a catalog of the cecidomyiidae - Agricultural Research Service - US ...

UPDATE FOR 

A CATALOG OF THE 

CECIDOMYIIDAE (DIPTERA) 

OF THE WORLD 

By 

Raymond J. Gagné 

Systematic Entomology Laboratory, Agricultural Research Service, U.S. Department of Agriculture 

U. S. National Museum NHB 168, P.O. Box 37012, Washington, DC 20013-7012, USA (raymond.gagne@ars.usda.gov). 

F

PREFACE TO THIS 2010 UPDATE 

This update to A Catalog of the Cecidomyiidae (Diptera) of the World (Gagné 2004b) will allow scientists 

and others to track nomenclatural changes and the additional 680 species and 185 genera that have been described since 

the cut-off date (sometime in 2003) for Gagné (2004b). The present totals are 6131 species and 783 genera, . This 

update lists the taxonomic papers on the group known to me as of March 21, 2010. The original introduction that 

follows is changed in minor ways to remain current, but the main points and modus operandi remain the same. 

Some changes in presentation of the catalog are made here. All genera, species, and literature cited new to this 

update are flagged by an asterisk. All fossil species, including those cited in the original catalog, are now preceded by a 

dagger to designate them more clearly. Some commentary following particular genera has been made, added to, or 

changed because of new developments or personal realizations. New family group names that have been proposed 

since Gagné (2004b) are discussed under Appendix 1, Classification of the Cecidomyiidae. The commentary in that 

section is largely rewritten to reflect changes that have occurred in the family since Gagné (2004b). The original 

Appendix 2 of Gagné (2004b) listing new type species, species, names, synonyms, and combinations has been omitted. 

No new formal taxonomic changes are made here. Some 20 species are now seen to be misplaced in their 

present genera or nomina dubia. They remain as were in Gagné (2004b), but are clearly marked, mostly as “per M. 

Jaschhof (in litt.).” Future authors are welcome to formalize these suggested changes. 

Since 2003, I have found or been made aware of some errors and a few missed papers in the original catalog. 

These are remedied in this present update and are listed for the record in an Errata section that follows "Literature 

Cited." 

Mathias Jaschhof of the Senckenberg Deutsches Entomologishes Institut, Müncheberg, Germany has carefully 

reviewed the sections on Catotrichinae, Lestremiinae, and Micromyinae, and has helpfully agreed to keep track of 

changes to these groups, the wood midges, from now on. Netta Dorchin and Keith M. Harris kindly read and 

commented on Appendix 1 for this update, as did Junichi Yukawa, who also helped with a complete list of literature, 

changes and additions by him and colleagues so that I could double-check those sources and results. I am grateful also 

for the general reviews of the manuscript and suggestions made by Alexander S. Konstantinov and Allen L. Norrbom, 

both of the Systematic Entomology Laboratory (SEL). I thank Mark Metz of the SEL for his technical advice and 

expertise, and Alma Solis, Research Leader of SEL, for making this update available on the Laboratory's website. 

The correct citation for this update is: Gagné, R.J. 2010. Update for a catalog of the Cecidomyiidae (Diptera) 

of the world. Digital version 1. 

The file can be downloaded from: 

http://www.ars.usda.gov/SP2UserFiles/Place/12754100/Gagne_2010_World_Catalog_Cecidomyiidae.pdf 

Cover illustration: The tumor stem gall midge, Asphondylia atriplicis, on saltbush, Atriplex canescens. Adult hardening 

before flight (photographed by Bradford A. Hawkins, University of California, Riverside).

INTRODUCTION 

Cecidomyiidae are a large family of nematocerous Diptera best known as plant gall makers. Included in this 

family are some of the most destructive pests of grains, fruits, and vegetables as well as important predators of aphids, 

scale insects and mites. Larvae show a great range of feeding habits, including fungivory, herbivory, and predation on 

various arthropods. The family has an ancient origin, with a fossil known from the Jurassic Period, but the largest 

subfamily, the Cecidomyiinae, expanded greatly on flowering plants during the Cretaceous Period. That subfamily now 

comprises the largest gall-making group among insects and mites and its great diversity parallels that of acalyptrate 

Diptera. Of the 6,131 known species in the family, a number that includes 107 fossil species, approximately four-fifths 

are associated with flowering plants or plant-feeding arthropods. Nevertheless, comparatively little is known of this 

group of insects, and species new to science are continually being found. To illustrate, The Plant-Feeding Gall Midges 

of North America (Gagné 1989b) listed the 900 examples of cecidomyiid plant associations reported for North America 

up to 1987. Since that book appeared, more than 279 additional examples have been discovered (mostly unpublished) 

of phytophagous species on the continent, and this without any great concerted effort. The number of species of all 

Cecidomyiidae still unknown in the world, especially in the tropics, is inestimable. 

The only other world catalog of Cecidomyiidae, Kieffer 1913i, appeared a little less than 100 years ago. 

Several regional catalogs were published in the interim: Foote 1965 for the Nearctic Region, Gagné 1973a for the 

Oriental, Harris 1980 for the Afrotropical, Skuhravá 1986 for the Palearctic, Gagné 1989c for the Australasian and 

Oceanian, and Gagné 1968d, 1994a for the Neotropical Region. These catalogs, now all much out of date, were very 

useful and most welcome when they appeared, especially those that included host data. Having only regional coverage, 

they lacked the breadth needed to convey the full geographical extent of the various genera and higher categories and 

the number of species they contain. Further, the separate catalogs appeared at different times so were inconsistent in 

their treatment of genera and higher groups. 

In preparing this catalog I examined anew all the literature cited except for the approximately 25 items I could 

not find and that are clearly marked in the catalog as not seen. All published generic and specific names of 

Cecidomyiidae, recent and fossil, are listed, together with their original references of author, date, and page. For genera, 

the catalog includes synonyms, emendations, misspellings, type species and kind of fixation, general distribution, 

number of species, suprageneric classification, and occasional notes and helpful references. Information provided for 

species includes range, hosts (for the subfamily Cecidomyiinae only), original spelling and original genus, synonyms, 

subsequent misspellings and emendations, type locality, kind of type, stage(s) of type(s), type deposition, and 

occasional notes. 

Organization and Format 

The arrangement of taxa in this catalog is first by subfamily, in order of presumed age, then by genera in 

alphabetical order. Supertribal and tribal dispositions for genera, when pertinent and if known, are noted in the generic 

entries and summarized in Appendix 1. At the end of each subfamily section are lists of species unplaced to genus and 

dubious names, if any. References to family group names are not given. Most subfamily and tribal names referred to 

were catalogued in Sabrosky 1999 so are not treated here except that new ones proposed since that paper are followed 

by their proper citation in Appendix 1. 

Appendix 1 lists the genera in context of their known classification and recommends useful publications for 

the study and identification of gall midges. Appendices 2 and 3 list and explain general abbreviations and four-letter 

abbreviations for type collections, respectively. Appendix 4 presents some notes on a few cecidomyiid collections. 

Generic entries.--Currently used generic names are centered, capitalized, and in boldface. Listed in 

chronological order below the heading are that name and any synonyms, misspellings, and emendations, each with 

author, date, and page. These citations are followed by the name of the type species (the original combination given 

only if not the current name), the kind of type designation, and, if the type species name is a junior synonym, the 

present senior synonym. New names or taxonomic decisions made here are in boldface following the entry. A separate 

paragraph following the generic entry gives geographic and temporal (for fossils) distribution of the genus, the number 

of included species, and tribal affiliation, if applicable. For Cecidomyiinae, host ranges are usually summarized for 

genera of two or more species. Useful references, reasons for any new taxonomic decisions, and additional notes are 

given as warranted, but general taxonomic references from Appendix 1 are not repeated. 

Species entries.--Valid specific names appear in boldface on the left margin, followed by distribution and, for 

2

3 

Cecidomyiinae, host or hosts if known. Indented below that line in chronological order is the original spelling and 

combination of that name, followed by any synonyms, misspellings, and emendations, with author, date, and page or 

plate for each name. Following each species are: the type locality or localities; kind of type; stage of type; and 

depository. Any new taxonomic decisions are signalled in boldface following the pertinent entries. Explanatory notes 

and further references are occasionally provided. 

Misspellings.--Alternative spellings accompanying the original description are cited, as well as those in 

subsequent papers that might tend to mislead because they are in headings, lists, or keys. Textual typographical errors 

in papers subsequent to the original that can be understood as such in context were not cited. Only the first example 

known to me of a misspelled name is indicated, so the term "of authors" is not used. 

Misidentifications.--These are only rarely noted, as in the case of economically important species, e.g. the 

North American Asphondylia websteri that was at one time cited as A. miki, a European species. 

Distribution.--Ranges given are from various sources, including revisions, previous catalogs, and faunal lists. I 

cannot vouch for all localities cited in the literature. Most historical locality records based on galls alone, if they do not 

stretch a range unduly, have been presumed correctly identified, although ultimate certainty depends on identification 

of the cecidomyiid inside the gall. 

The distribution of a species is outlined by country, generally from west to east and north to south. State, 

province or territory is specified for some of the larger countries, viz., Argentina, Brazil, Canada, China, India, Mexico, 

and USA. Russia is divided into four parts: Europe, west of the Urals; Western Siberia, the Urals east to the Yenisey 

River; Eastern Siberia, the Yenisey River east to Magadan Province; and the Far East, Magadan Province eastward. 

This generally follows the scheme used earlier in the Catalogue of Palaearctic Diptera (cf. Soós & Papp 1986) for the 

division of the USSR. Larger island distribution is given for Indonesia and Japan. Abbreviations are used for the names 

of only two countries, UK for United Kingdom and USA for United States of America. Fossil distribution is given as 

epoch first, followed by geographic locality. 

Far-flung states or parts of certain countries, e.g., Hawaii (as Hawaiian Islands here), Trinidad, and Tobago, 

are treated for convenience as if they were separate countries. Washington, DC, is given as "District of Columbia" so 

that it will not be confused with the State of Washington. No differentiation was made between Czech Republic and 

Slovakia for older Czechoslovakian records or among the present states formerly part of Yugoslavia, except for typelocalities 

and more recent records. 

Distribution is drawn here with a broad brush. A listing of "France, Germany" could mean that the species 

extends from the Baltic to the Mediterranean or only that it has been found on opposite banks of the Rhine. Most gall 

midges are probably not very restricted within a biome and most are probably more widely distributed than presently 

known. When a species has a broad range, e.g., one that has been recorded from the United Kingdom, Sweden, 

Germany, Latvia, European Russia, Hungary, Spain, Italy, and Greece, its distribution is summarized here as 

"widespread Europe." This avoids taking space for long lists of countries for common species. In the same way, I felt 

confident in giving a designation of "widespread eastern Nearctic" for species known from, e.g., Ontario, Canada and 

Florida, USA and many states between when the host occurs throughout the entire region. 

Hosts.--The type host or hosts for the senior name is given first, separated by a semicolon from any additional 

original or subsequent hosts. If the host became known only after the original description, the entry reads "Orig. unk.;" 

followed by the host. Three or more additional original or subsequent hosts of the same genus as the type host are not 

tabulated but summarized by "spp." to obviate long lists for oligophagous species. Older names for hosts are corrected 

to current names, but some recent name changes are not reflected here. Authors of host names are unnecessary in the 

context of this catalog and are not supplied, but the family for plant hosts, order for fungal hosts, and family and order 

for animal hosts are provided because they are valuable to anyone looking for cecidomyiids that occur on certain hosts 

or host groups. Many European gall makers have, subsequent to their original descriptions, been identified and 

recorded based on galls. These records are included uncritically here, but some records of gall associates, including 

predators and inquilines, that I suspect were not based on reared adults to critically determine them, are left out. 

Type localities.--Place names are as much as possible corrected to current and consistent use. For instance, 

where J.-J. Kieffer used or implied only Lorraine, I have used Moselle, the current department name for that part of 

Lorraine where he lived all of his life. For another example, Kieffer variously used "La Grange," "Lagrange," "Château 

de La Grange," "château de la Grange," and "forêt de La Grange" for a domicile and its surrounds near Thionville in 

Moselle where he spent two months one summer tutoring a student (Fleur 1929) and collected a large number of 

species that he subsequently described. All mentions of this type locality are given in this catalog as "château de La 

Grange, nr Thionville." I have not added the department name for uniquely named towns in France as the French Code

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Postal makes this superfluous. Russian localities for the most part are transliterated using the American National 

Standards Institute scheme, but spellings of higher category names, i.e., cities and provinces or territories, as in Alma- 

Ata and Kamchatka, follow common usage of current English language atlases. 

Kind of type.--This information is given as one- or two-letter designations: HT, LT, NT, ST, and T, for, 

respectively, holotype, lectotype, neotype, syntypes, and type (when the kind of type could not be determined). If the 

type is a lectotype or neotype, LT or NT are followed by parentheses enclosing the citation for the designation, as well 

as, for neotypes, the new type locality. 

Stage or sex of type.--This is rendered as: A for adult of unknown sex; ♂ for adult male; ♀ for adult female; P 

for pupa; L for larva; E for egg; and G for gall. In the case of syntypes of multiple life stages or sexes, the symbols are 

run together separated only by a comma or commas. For species described prior to 1931, galls, being considered the 

work of a species, can be considered type specimens. I list the gall as a type only by default, in absence of the insect: if 

someone originally based a species on a larva and its gall, I list only the larva as a type. 

Type depository.--Four-letter abbreviations used here for type depositories are mainly from Arnett et al. 1993, 

but others were newly coined as needed. No abbreviations were used for certain collections of some authors who 

described fewer than three species, in which case depository information is written out in full. The abbreviations and 

their equivalents are listed in Appendix 3. A question mark is used when the depository was left unstated and could not 

otherwise be determined. A question mark follows the word "lost" when it is possible a type might be found 

somewhere. Notes on types in some collections are given in Appendix 4. 

Unplaced species of Cecidomyiidae.--This category at the end of the catalog combines species of all 

subfamilies that are currently unplaced to genus. Most of these were originally combined in catchall genera, e.g., 

Cecidomyia, that have since been more narrowly defined. Many Cecidomyiinae are considered unplaced for the first 

time here because they obviously do not belong in the genera to which they were assigned originally or subsequently. 

Most may be properly placed some day because they have or may have extant types or were described with illustrations 

that may suffice for identification. Many Cecidomyiinae were described from known hosts and plant galls that can be 

found again to obtain additional specimens. Numerous unplaced species, particularly from the Oriental Region, appear 

to represent new genera. 

Nomina dubia.--This category, at the end of the catalog as well, includes genera and species that were too 

poorly described to be recognized now and whose type species are unknown or whose type specimens are lost, 

presumed lost, or reduced to fragments. If hosts are known, the cecidomyiids are only collaterally involved with galls or 

damage. These names could be applied to new species that generally fit an old description. Dubious species names are 

not included in counts of species in the catalog. 

Some Taxonomic Considerations 

Certain taxonomic matters, some peculiar to gall midges, should be explained or mentioned here. These 

concern names based on galls only, polynomial names once used in naming gall midges, and forgotten names or 

nomina oblita. 

The names of some cecidomyiid species described long ago are based only on galls. For example, Osten 

Sacken (1875) described a leaf gall on Urtica as Cecidomyia urnicola (a species now tentatively placed in Dasineura). 

The description was adequate for the time because the gall could easily be recognized and lead directly to the gall 

maker. According to present rules of nomenclature, such a name, as long as it was proposed before 1931, is available 

because it is based on the work of an animal (International Code of Zoological Nomenclature 1999). One might counter 

that a gall is the work of the plant, but the gall is at least a manifestation of the interaction between two organisms. Even 

before 1931, though, not everyone agreed that such a practice was proper or that such a name was available (Löw 

1877). Löw (1877) and Rübsaamen (1899b) even treated a name based on the larval stage as unavailable because that 

was not the "perfect form" of the insect. They each renamed some species that had been based only on galls or galls and 

larvae. Nonetheless, the earlier names have precedence. 

Some specific names were coined as two separate words as a device to categorize the host and kind of gall in 

one scientific name. For example, Walsh (1864b) coined the name Cecidomyia salicis batatas for the gall midge that 

forms a tuber-shaped enlargement on willow twigs. That name is unavailable now because the rules require that a 

specific name be a single word based on the name of a single entity. A name for that gall midge became available when 

Osten Sacken (1878) listed the two words as one, salicisbatatas. Cecidomyia bigeloviae-strobiloides Townsend was 

available when originally published because the words were united by a hyphen. Because present rules require that

5 

names be spelled without a hyphen, that name is now properly written as bigeloviaestrobiloides. The name proposed as 

Cecidomyia pini inopis Osten Sacken 1862 is considered valid with the original author and date, now as piniinopis, 

because, although the name was actually two separate words, it referred to a single entity, i.e., the plant Pinus inops, 

now Pinus virginiana. 

The fourth edition of the International Code of Zoological Nomenclature 1999 added a new provision (Article 

23.9) for upsetting precedence of names when certain senior names have not been used as valid names after 1899. This 

unfortunate provision neither fosters nor rewards thorough scholarship and implies that an occasional new name change 

is too great a hardship for the scientific community to accommodate. Forgotten names are not so numerous and are 

certainly finite. In my opinion it is better to leave a principle whole than devise a set of rules to undermine, or, as the 

Code expresses it, "moderate" its application. 

It did not take long for the new rule to be invoked in Cecidomyiidae, although not all its provisions were 

followed. For example, Bland (2000) designated Cecidomyia rosarum Hardy 1850 as senior synonym of the earlier 

Cecidomyia rosae Bremi 1847 (now in Dasineura), considering C. rosae a forgotten name or nomen oblitum. Unnoted 

by Bland was that Hardy (1854f) himself synonymized his rosarum under rosae Bremi. Also, the name rosae Bremi 

was used in Gagné 1972b, so it was not a forgotten name. Bland did not meet the conditions in the Code for designating 

rosae a nomen oblitum. It is the responsibility of the designator to give evidence that the younger name, the name to be 

preserved, has been cited in the literature as a valid name in at least 25 works published by at least 10 authors in the 

immediately preceding 50 years and encompassing a span of not less than 10 years. These strictures appear to be 

written so that the new rule would apply only to species of considerable significance. Inasmuch as these conditions are 

not met in this case, I am replacing the older name as senior synonym. Rule 23.10 of the Code states that if an action is 

erroneous, the matter is to be referred to the International Commission on Zoological Nomenclature. The onus in such a 

case as this, then, is unfairly placed on the challenger and not the proposer. 

During the preparation of this catalog several names from the 1700s and 1800s surfaced that had been unused 

since originally proposed. Most of them have seniority over other names for the same species and are accordingly used 

as senior names here because the tests of the rules for designating them nomina oblita do not apply. 

SUBFAMILY CATOTRICHINAE 

CATOTRICHA 

Catotricha Edwards 1938e: 102. Type species, Catocha americana Felt (orig. des.). 

Holarctic; 5 spp. Ref.: Jaschhof 2001a (rev.). 

americana. USA (New Hampshire). 

Catocha americana Felt 1908b: 309. USA: New Hampshire, Franconia; HT; ♂; USNM. 

marinae. Russia (Western Siberia). 

Catotricha marinae Mamaev 1985: 24. Russia: Tuva, Ishtii-Khem; HT; ♀; HYNM. 

nipponensis. Japan (Honshu). 

Catocha nipponensis Alexander 1924: 83. Japan: Honshu, Osaka Pref., Mt Minomo, Settsu-no-kuni; HT; ♂; USNM. 

Catotricha antennata Alexander 1959: 39. Japan: Honshu, Akigami, On-take, Shinano-Hida; HT; ♂; USNM. 

subobsoleta. USA (Washington, California). 

Catocha subobsoleta Alexander 1924: 82. USA: Washington, Mt Rainier, Longmire Springs; HT; ♂; USNM. 

subterranea. Russia (Far East). 

Catotricha subterranea Mamaev 1985: 25. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♀; HYNM. 

*MESOTRICHOCA 

Mesotrichoca Jaschhof & Jaschhof 2008: 58. Type species, Catotricha mesozoica Kovalev (orig. des.). 

Sediment fossil, Upper Jurassic/Lower Cretaceous of Siberia (Russia); 1 sp.

†mesozoica. Upper Jurassic/Lower Cretaceous - Russia (Siberia). 

Catotricha mesozoica Kovalev 1990: 175. Russia: Siberia, Upper Jurassic/Lower Cretaceous; HT; ♂; ?. Ref.: 

Jaschhof & Jaschhof 2008. 

*TRICHOTOCA 

Trichotoca Jaschhof & Jaschhof 2008: 54. Type species, edentula Jaschhof & Jaschhof (orig. des.). 

Australasian; 2 spp. 

*edentula. Australia (Tasmania). 

Trichotoca edentula Jaschhof & Jaschhof 2008: 56. Australia: Tasmania, Warra Long Term Ecological Research Site, 

Mt. Weld; HT; ♂; AMSY. 

fraterna. Australia (Australian Capitol Terr., New South Wales, Victoria, Tasmania). 

Catotricha fraterna Jaschhof 2001a: 643. Australia: Australian Capital Terr., Black Mtn; HT; ♂; ANIC. Ref.: 

Jaschhof & Jaschhof 2008. 

SUBFAMILY LESTREMIINAE 

ALLARETE 

Allarete Pritchard 1951: 250. Type species, Lestremia vernalis Felt (orig. des.). 

Allaretina Mamaev 1994: 28, as subg. of Allarete. Type species, Allarete nigra Mamaev (orig. des.). 

Holarctic, Afrotropical, Oriental; 11 spp.. Ref: Jaschhof & Jaschhof 2009a (rev.). 

africana. Tanzania. 

Anaretella africana Enderlein 1911c: 193. Tanzania: S L. Victoria, Myembe; HT; ♀; STZM. 

barberi. USA (California, Arizona, New Mexico, Minnesota), Germany, Italy, Croatia, Greece. 

Lestremia barberi Felt 1908b: 310. USA: New Mexico, Las Vegas; HT; ♀; USNM. 

bharatica. India (Uttar Pradesh). 

Allarete bharatica Grover & Bakhshi 1978: 6. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

*bhokarensis. India (Maharashtra). 

Anaretella bhokarensis Deshpande, Shaikh & Sharma 2002b: 41. India: Maharashtra, Nanded District, Bhokar; HT; 

♂; DESH. 

bicornuta. USA (Arizona). 

Allarete bicornuta Jaschhof 1997b: 526. USA: Arizona, Dragoon Mtns, Cochise Stronghold; HT; ♂; USNM. 

*deepica. India (Maharashtra). 

Anaretella deepica Deshpande, Shaikh & Sharma 2002b: 37. India: Maharashtra, Nanded District, Bhokar; HT; ♂; 

DESH. 

*indica. India (Maharashtra). 

Anaretella indica Deshpande, Shaikh & Sharma 2002b: 39. India: Maharashtra, Nanded District, Bhokar; HT; ♂; 

DESH. 

nigra. Finland, Austria, Kirghizstan, Russia (Far East). 

Allarete nigra Mamaev 1994: 28. Russia: Kamchatka, Kozyrevsk; HT; ♂; HYNM. 

Allarete kirghizica Mamaev & Zaitzev 1998a: 3. Kirghizstan: Sary-Chelek Res.; HT; ♂; ZMMU. 

spatuliformis. India (West Bengal). 

Allarete spatuliformis Grover 1979a: 3. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

*spinosa. India (Maharashtra). 

Anaretella spinosa Deshpande, Shaikh & Sharma 2002b: 35. India: Maharashtra, Nanded District, Bhokar; HT; ♂; 

DESH. 

6

vernalis. USA (Minnesota, Kansas, Texas), Germany, Uzbekistan, Turkmenistan. 

Lestremia vernalis Felt 1908b: 311. USA: Kansas, Wichita; HT; ♂; NYSM. 

Allarete distincta Mamaev 1963c: 71. Uzbekistan: Kitab; HT; ♂; UASS (actually in ZMMU). 

Allarete turkmenica Mamaev & Zaitzev 1998a: 2. Turkmenistan: Badkhyz Res.; HT; ♂; ZMMU. 

ALLARETELLA 

Allaretella Meyer & Spungis 1994: 436. Type species, germanica Meyer & Spungis (orig. des.). 

Western Palearctic; 1 sp. 

germanica. Germany, Austria. 

Allaretella germanica Meyer & Spungis 1994: 436. Germany: sea dikes nr Meldorf; HT; ♂; LASS. 

ANARETE 

Anarete Haliday 1833: 156. Type species, candidata Haliday (mon.). 

Pseudanarete Kieffer 1906d: 342. Type species, Anarete crassipalpis Kieffer (mon.) = lacteipennis Kieffer. 

Microcerata Felt 1908b: 309. Type species, Micromya corni Felt (orig. des.) 

Limnopneumella Enderlein 1911c: 195. Type species, Anarete stettinensis Enderlein (orig. des.). 

Holarctic, Neotropical, Oriental, Oceanian; 33 spp. Refs: Kim 1967 (New World spp.), Berest 2000 (Palearctic spp.), 

Jaschhof & Jaschhof 2009a (redescr.). 

albipennis. Germany. 

Anarete albipennis Loew 1845b: 395. Germany: Kassel-Nordshausen; ST; ♂,♀; ZMHB?. 

allahabadensis. India (Uttar Pradesh). 

Anarete allahabadensis Grover 1964a: 116. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

anepsia. USA (Oregon, California). 

Anarete anepsia Pritchard 1951: 264. USA: California, Orinda; HT; ♂; USNM. 

basmatensis. India (Maharashtra). 

Anarete basmatensis Bhalerao, Shaikh and Deshpande 2009: 227. India: Maharashtra, Basmathnagar; HT; ♀; 

DZNM. 

bharati. India (Maharashtra). 

Anarete bharati Bhalerao, Shaikh and Deshpande 2009: 226. India: Maharashtra, Nanded, Botanical Garden, Science 

College Campus; HT; ♂; DZNM. 

buscki. USA (Arizona, Minnesota), Cuba, Puerto Rico, Galapagos Is. 

Microcerata buscki Felt 1915a: 198. Cuba: Baracoa; HT; ♂; USNM. 

candidata. UK, Sweden, Germany, Russia (Europe). 

Anarete candidata Haliday 1833: 156. UK: Northern Ireland, Holywood; T; A; NMID. 

corni. USA (widespread), UK, Germany, Latvia, Russia (Europe). 

Micromya corni Felt 1907a: 6. USA: New York, Albany; HT; ♂; NYSM. 

Microcerata perplexa Felt 1908b: 310. USA: New York, Albany; HT; ♀; NYSM. 

Microcerata borealis Felt 1913c: 147. USA: New York, Speculator; HT; ♂; NYSM. 

Anarete angustata Edwards 1929a: 16. UK: England, Herts, Letchworth; HT; ♂; BMNH. 

coronata. Russia (Europe). 

Anarete coronata Mamaev 1964a: 778. Russia: Moscow Prov., Bykovo; HT; ♂; HYNM. 

diervillae. USA (New York). 

Micromyia diervillae Felt 1907a: 6. USA: New York, Karner; HT; ♂; NYSM. 

edwardsi. USA (Minnesota). 

Anarete edwardsi Pritchard 1951: 259. USA: Minnesota, Anoka; HT; ♂; USNM. 

felti. USA (Oregon, Minnesota, Florida). 

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8 

Anarete felti Pritchard 1951: 262. USA: Minnesota, Wayzata; HT; ♂; USNM. 

flavitarsis. Russia (Near East). 

Anarete flavitarsis Mamaev 1964a: 778. Russia: Dagestan, Arani; HT; ♂; HYNM. 

iridis. USA (North Dakota, Minnesota, Illinois, Arkansas), China. 

Microcerata iridis Cockerell 1914a: 460. USA: Colorado, ca 3 mi E Boulder; ST; ♂; USNM. 

jagdyevi. Turkmenistan. 

Anarete jagdyevi Mamaev 1986: 66. Turkmenistan: Kushka, Badkhyz Res.; HT; ♂; HYNM. 

johnsoni. USA (widespread), UK, Germany, Russia (Europe), Japan, Hawaiian Is. 

Microcerata johnsoni Felt 1908b: 310. USA: Pennsylvania, Philadelphia; HT; ♂; NYSM. 

Microcerata cockerelli Felt 1908b: 310. USA: New Mexico, Mesilla; ST; ♂; USNM. 

Microcerata spinosa Felt 1913c: 145. USA: Texas, Plano; LT (Pritchard 1951: 260); ♂; NYSM. 

Microcerata aldrichii Felt 1915e: 226. USA: Indiana, Lafayette; LT (Pritchard 1951: 260); ♂; NYSM. 

Anarete heracleana Edwards 1938d: 29. UK: England, Herts, Baldock; ST; ♂,♀; BMNH. 

lacteipennis. UK, Finland, Germany, Austria, Russia (Europe), USA (Minnesota). 

Anarete lacteipennis Kieffer 1906d: 345. Austria: Styria; T; ♂; lost?. 

Anarete crassipalpis Kieffer 1906d: 346. Austria: Styria; T; ♀; lost?. 

longipalpi. India (Maharashtra). 

Anarete longipalpi Bhalerao, Shaikh and Deshpande 2009: 228. India: Maharashtra, Nanded, Science College 

Campus; HT; ♀; DZNM. 

mamajevi. Ukraine. 

Anarete mamajevi Berest 1987: 81. Ukraine: Odessa Prov., Datchnoe; HT; ♂; UASK. 

manii. India (Uttar Pradesh). 

Anarete manii Rao 1953: 307. India: Uttar Pradesh, Agra; HT; ♀; ZSCI. 

medicaginis. Uzbekistan. 

Anarete medicaginis Mamaev 1963c: 72. Uzbekistan: Bukhara Prov., Mubarek; HT; ♂; MAMC. 

pallida. Samoa. 

Anarete pallida Edwards 1928: 39. Samoa: Tutuila; HT; ♀; BMNH. 

perplexia. Ukraine. 

Anarete perplexa Berest 2000: 40, jun. sec. hom. perplexa Felt. Ukraine: Chernigov Prov., Nezhinskii; HT; ♂; 

UASK. 

Anarete perplexia Gagné 2004b: 11, new name for Anarete perplexa Berest. 

pilipennis. Slovenia. 

Anarete pilipennis Strobl 1910: 225. Slovenia: Save Valley, nr Brezice; HT; ♀; NMBA. 

pritchardi. USA (Minnesota, Illinois). 

Anarete pritchardi Kim 1967: 528. USA: Minnesota, Waseca; HT; ♂; USNM. 

rubra. France, USA (Oregon, California, Minnesota). 

Anarete rubra Kieffer 1906d: 343. France: vic. Bitche; ST; ♂,♀; lost. 

serena. Ukraine. 

Anarete serena Berest 2000: 39. Ukraine: Dnepropetrovsk; HT; ♂; UASK. 

sitapurensis. India (Uttar Pradesh). 

Anarete sitapurensis Grover 1970c: 148. India: Uttar Pradesh, Sitapur; HT; ♂; PGCA. 

stettinensis. Poland. 

Anarete stettinensis Enderlein 1911a: 132. Poland: Radu� nr Stepnica, nr Szczecin; HT; ♀; ZMPA. 

taimyrensis. Russia (Eastern Siberia). 

Anarete taimyrensis Mamaev 1990b: 141. Russia: Taimyr, Tareya; HT; ♂; HYNM. 

*taishanensis. China (Shandong). 

Anarete taishanensis Mo & Xiao 2004: 163. China: Shandong, Taishan; HT; ♂; SAUT. 

triarthra. UK, Germany. 

Anarete triarthra Edwards 1938d: 31. UK: England, Herts, Letchworth; HT; ♀; BMNH. 

zhengi. China (Inner Mongolia Aut. Reg.). 

Anarete zhengi Bu & Li 2001: 26. China: Inner Mongolia Aut. Reg., Guyang, Xingshunxi; HT; ♂; NKUM.

ANARETELLA 

Anaretella Enderlein 1911c: 193. Type species, Lestremia defecta Winnertz (orig. des.). 

Neptunimyia Felt 1912l: 237. Type species, tridens Felt (orig. des.) = defecta Winnertz. 

Plocimas Enderlein 1936: 59. Type species, Plocimas hirsutus Enderlein (mon.) = defecta Winnertz. 

Holarctic, Oriental, Oceanian; 6 spp. Ref.: Jaschhof 1998b (rev.), Jaschhof & Jaschhof 2009a (redescr.). 

acutissima. Russia (Far East). 

Anaretella acutissima Mamaev 1998d: 1. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

defecta. Widespread Holarctic, Taiwan. 

Lestremia defecta Winnertz 1870b: 33. Germany; ST; ♂,♀; lost. 

Campylomyza acerifoliae Felt 1907a: 5. USA: New York, Albany; HT; ♂; NYSM. 

Catocha spiraeina Felt 1907a: 6. USA: New York, Albany; HT; ♂; NYSM. 

Lestremia pini Felt 1907a: 7. USA: New York, Albany; HT; ♂; NYSM. 

Campylomyza acerifolia Felt 1907c: 101, unjust. emend. of acerifoliae. 

Neptunimyia tridens Felt 1912l: 237. USA: New York, Nassau; LT (Pritchard 1951: 249); ♀; NYSM. 

Neptunimyia bromleyi Barnes 1928g: 173. UK: England, Suffolk, Wickhambrook; ST; ♂,♀; BMNH. 

Plocimas hirsutus Enderlein 1936: 59. Germany: Krefeld; ST; ♂; lost?. 

Lestremia strobli Edwards 1938d: 27. UK: Wales, Brecon, Crickhowell; HT; ♂; BMNH. 

Anaretella cincta Mamaev 1964a: 777. Russia: Moscow Prov.; HT; ♂; HYNM. 

Anaretella elegantula Mamaev 1964a: 777. Russia: Krasnodarskiï Terr., Novo-Prokhladnoye; HT; ♂; HYNM. 

Anaretella magnicornis Mamaev 1964a: 777. Russia: Voronezh Prov., Tellerman; HT; ♂; HYNM. 

*Anaretella bicincta Mamaev 1969a: 371, error for cincta. 

glacialis. Russia (Western Siberia). 

Anaretella glacialis Mamaev & Økland 1996: 3. Russia: Yamal Peninsula, Khadyta; HT; ♂; ZMMU. 

iola. USA (Washington), Fennoscandia, Netherlands, Latvia, Germany, Austria, Russia (Western Siberia). 

Anaretella iola Pritchard 1951: 250. USA: Washington, American R.; HT; ♂; USNM. 

Anaretella borealis Mamaev 1985: 28. Russia: Yamal Peninsula, Khadyta; HT; ♂; HYNM. 

Anaretella corniculata Mamaev 1985: 28. Russia: Yamal Peninsula, Khadyta; HT; ♂; HYNM. 

nitida. Samoa. 

Lestremia nitida Edwards 1928: 39. Samoa: Upolu, Malololelei; HT; ♀; BMNH. 

supermagna. Russia (Europe). 

Anaretella supermagna Mamaev & Økland 1996: 3. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ZMMU. 

CONARETE 

Conarete Pritchard 1951: 255. Type species, crebra Pritchard (orig. des.). 

Holarctic, Neotropical, Afrotropical, Oriental; 12 spp. Refs: Pritchard 1951 (Nearctic spp.), Gagné 1995d (Old 

World, review). 

*brevipalpa. China (Hainan). 

Conarete brevipalpa Li & Bu 2002b: 222. China: Hainan, Mt. Jianfeng; HT; ♂; NKUM. 

calcuttaensis. India (West Bengal), South Africa. 

Anarete calcuttaensis Nayar 1949: 79. India: West Bengal, Calcutta; HT; ♂; ZSCI. 

crebra. USA (Minnesota). 

Conarete crebra Pritchard 1951: 256. USA: Minnesota, Brauerville; HT; ♂; USNM. 

deepica. India. 

Conarete deepica Deshpande 1983: 124. India; T; A; DESH. 

eluta. USA (Minnesota), Dominica. 

Conarete eluta Pritchard 1951: 257. USA: Minnesota, Houston Co.; HT; ♂; USNM. 

9

eschata. USA (Minnesota). 

Conarete eschata Pritchard 1951: 257. USA: Minnesota, Houston Co.; HT; ♂; USNM. 

indica. India (Maharashtra). 

Microcerata indica Mani 1934b: 382. India: Maharashtra, Satara Distr., Yenna Valley, Medha; ST; ♂; ZSCI. 

indorensis. India (Madhya Pradesh), China (Shaanxi). 

Conarete indorensis Grover 1970c: 141. India: Madhya Pradesh, Indore; HT; ♂; PGCA. 

mihijamensis. India (Bihar), China (Hainan). 

Conarete mihijamensis Grover 1964a: 114. India: Bihar, Mihijam; HT; ♂; PGCA. 

orientalis. India (Uttar Pradesh). 

Conarete orientalis Rao 1956a: 67. India: Uttar Pradesh, Agra; HT; ♂; ZSCI. 

*sicyoidea. China (Hainan). 

Conarete sicyoidea Li & Bu 2002b: 222. China: Hainan, Mt. Jianfeng; HT; ♂; NKUM. 

texana. USA (Minnesota, Texas). 

Microcerata texana Felt 1913c: 147. USA: Texas, Plano; HT; ♂; NYSM. 

*EOMASTIX 

Eomastix Jaschhof 2009a: 66. Type species, Gongromastix incerta Jaschhof (orig. des.). 

Western Palearctic; 1 sp. 

incerta. Norway, Sweden. 

Gongromastix incerta Jaschhof 2002: 428. Sweden: Södermanland, Tyresta Nature Res.; HT; ♀; NHRS. Ref.: 

Jaschhof & Jaschhof 2009a (redescr.). 

GONGROMASTIX 

Gongromastix Enderlein 1936: 60. Type species, andorrana Enderlein (mon.) = angustipennis. 

Pararete Pritchard 1951: 253, preocc. Ijima 1927. Type species, Lestremia elongata Felt (orig. des.). 

*Pritchardaea Ozdikmen 2009: 202, new name for Pararete Pritchard. 

Holarctic, Oriental; 7 spp. Ref.: Jaschhof & Jaschhof 2009a (redescr.). 

angustipennis. USA (Washington, California, Minnesota), Spain, Andorra, Italy, Austria, Croatia, Ukraine, Russia 

(Western Siberia). 

Lestremia angustipennis Strobl 1902: 511. Croatia: Lesina; HT; ♂; NMBA. 

Gongromastix andorrana Enderlein 1936: 60. Andorra: Lago Engolastes; HT; ♂; ZMHB. 

Gongromastix epista Pritchard 1951: 253. USA: Washington, American R.; HT; ♂; USNM. 

Gongromastix schalis Pritchard 1951: 252. USA: Minnesota, Anoka; HT; ♂; USNM. 

Gongromastix major Mamaev 1985: 27. Russia: Tuva, Ishtii-Khem; HT; ♂; HYNM. 

Gongromastix minor (also as major), Mamaev 1985: 27. Russia: Tuva, Ishtii-Khem; HT; ♂; HYNM. 

bharatae. India (Maharashtra). 

Gongromastix bharatae Sharma & Rao 1979c: 29. India: Maharashtra, Aurangabad, Khadkeshwar; HT; ♂; RCMU. 

elongata. USA (California). 

Lestremia elongata Felt 1908b: 310. USA: California, Argus Mts; HT; ♀; USNM. 

ignigena. Sweden. 

Gongromastix ignigena Jaschhof 2002: 430. Sweden: Södermanland, Tyresta Nature Res.; HT; ♂; NHRS. 

indica. India (Uttar Pradesh). 

Gongromastix indica Grover 1964a: 111. India: Uttar Pradesh, Allahabad; HT; ♀; PGCA. 

indicata. India (Maharashtra). 

Pararete indica Rao & Adwant 1975: 179, jun. sec. hom. indica Grover. India: Maharashtra, Ganesh Nagar, Nanded; 

HT; ♀; RCMU. 

10

11 

Gongromastix indicata Gagné 2004b: 21, new name for Pararete indica Rao & Adwant. 

orientalis. India (Maharashtra). 

Pararete orientalis Rao & Adwant 1975: 177. India: Maharashtra, Aurangabad, Marathwada Univ.; HT; ♀; RCMU. 

*INSULESTREMIA 

Insulestremia Jaschhof 2004c: 101. Type species, sinclairi Jaschhof (orig. des). 

Neotropical; 1 sp. 

*sinclairi. Galapagos Is. 

Insulestremia sinclairi Jaschhof 2004c: 103. Galapagos Is.: San Cristóbal, 3 km E Baquerizo; HT; ♂; CNCI. 

LESTREMIA 

Lestremia Macquart 1826: 173. Type species, cinerea Macquart (mon.). 

Cecidogona Loew 1844: 324, as subg. of Lestremia. Type species, Lestremia carnea Loew (mon.) = cinerea 

Macquart. 

Lestremya Rondani 1856: 213, missp. of Lestremia. 

Lestremina Shinji 1944: 175, missp. of Lestremia. 

Lestremyia Shinji 1944: 209, missp. of Lestremia. 

Cosmopolitan and Eocene amber; 18 spp. Refs.: Jaschhof 1998b (rev.), Jaschoff & Jaschhof 2003: 104 (New 

Zealand), Jaschhof & Jaschhof 2009a (redescr.). 


Lestremia allahabadensis Grover 1970c: 128. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

calcuttaensis. India (West Bengal). 

Lestremia calcuttaensis Mani 1937: 281. India: West Bengal, Calcutta, Portland Pk; HT; ♀; ZSCI. 

ceylanica. Sri Lanka. 

Lestremia ceylanica Kieffer 1912a: 29. Sri Lanka: Peradeniya; HT; ♀; NZSI?. 

cinerea. Widespread Holarctic, Chile, Hawaiian Is, New Zealand. Ref.: Jaschhof & Jaschhof 2009a (notes). 

Lestremia cinerea Macquart 1826: 173. France; T; A; MNHN?. 

Lestremia fusca Meigen 1830: 309. Germany; T; ♀; lost. 

Lestremia albipennis Waltl 1837: 280. Germany: Bavaria; T; A; lost. 

Lestremia albipennis Meigen 1838: 50. Germany: Bavaria; T; A; lost. 

Lestremia carnea Loew 1844: 324. Poland: Posnan; T; ♀; ZMHB?. 

Catocha sylvestris Felt 1907a: 5. USA: North Carolina, Davidson's R.; HT; ♀; NYSM. 

Lestremia dyari Felt 1908b: 311. Canada: British Columbia, Kaslo; HT; ♂; USNM. 

Lestremia franconiae Felt 1908b: 311. USA: New Hampshire, Franconia; HT; ♂; USNM. 

Lestremia kansensis Felt 1908b: 311. USA: Kansas, Douglas Co.; HT; ♀; NYSM. 

Lestremia declinata Kieffer 1910e: 232. Austria: Admont; ST; ♂,♀; lost?. 

Zygoneura fenestrata Malloch 1914: 233. USA: Illinois, Urbana; LT (Pritchard 1951: 246); ♂; INHS. 

Lestremia floridana Felt 1915e: 226. USA: Florida, Jacksonville; HT; ♀; NYSM. 

Lestremia garretti Felt 1926f: 265. Canada: British Columbia, Wilson Creek, Michel; HT; ♀; NYSM. 

Lestremia vittata Vimmer 1930: 28. Israel: Tel Aviv; HT; ♂; lost. 

Catocha iwatensis Monzen 1936: 45. Japan: Honshu, Morioka; ST; ♂,♀; KUEC. 

Lestremia intercalaria Mamaev 1998d: 1. Turkmenistan: Kopet-Dag, Ipai-Kala; HT; ♂; ZMMU. 

Lestremia albidipennis Gagné 2004b: 22, new name for Lestremia albipennis Meigen, jun. prim. hom. albipennis 

Waltl. 

Lestremia intercalaris Gagné 2004b: 22, missp. of intercalaria Mamaev. 

clivicola. Hawaiian Is.

12 

Lestremia clivicola Hardy 1960: 249. Hawaiian Is: Hawaii, Keanakolu; HT; ♂; BPBM. 

* † deploegi. Lower Eocene - amber, northern France. 

Lestremia deploegi Nel & Prokop 2006: 41. Amber fossil, northern France: Oise, Creil region, Chevrière; HT; ♂; 

NMHN. 

* † eocenica. Lower Eocene - amber, northern France. 

Lestremia eocenica Nel & Prokop 2006: 38. Amber fossil, northern France: Oise, Creil region, Chevrière; HT; ♂; 

NMHN. 

indica. India (West Bengal). 

Lestremia indica Kieffer 1909c: 129. India: West Bengal, Calcutta; HT; ♀; lost. 

leucophaea. Widespread Holarctic, Hawaiian Is, New Zealand. Ref.: Jaschhof & Jaschhof 2009a (notes). 

Sciara leucophaea Meigen 1818: 288. Germany; T; ♀; lost. 

Catocha sambuci Felt 1907a: 5. USA: New York, Albany; HT; ♀; NYSM. 

Lestremia setosa Felt 1908b: 311. USA: New York, Albany; HT; ♂; NYSM. 

Lestremia occidentalis Felt 1926f: 265. Canada: British Columbia, Cranbrook; HT; ♀; NYSM. 

Lestremia inexpectata Mamaev 1986: 65. Turkmenistan: Kopetdag, Ipaï-Kala; HT; ♂; HYNM. 

nigra. Chile. 

Lestremia nigra Blanchard 1852: 349. Chile: Carelmapu & Chiloe; ST; A; lost?. 

novaezealandiae. New Zealand. 

Lestremia novaezealandiae (as novae-zealandiae) Marshall 1896: 247. New Zealand; NT (Jaschhof & Jaschhof 

2003; South I., Nelson, Kahurangi Natl Pk, Takaka R. valley); ♂; NZAC. Ref.: Jaschoff & Jaschhof 2003: 

104. 

Trichosia remota Hutton 1902: 192. New Zealand: Christchurch; HT; ♀; CMNZ. Ref.: Jaschoff & Jaschhof 2003: 

104. 


Lestremia orientalis Grover 1964a: 108. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

palikuensis. Hawaiian Is. 

Lestremia palikuensis Hardy 1960: 252. Hawaiian Is: Maui, Haleakala Crater; HT; ♂; BPBM. 

parvostylia. Morocco, Estonia, Latvia. 

Lestremia parvostylia Jaschhof 1994: 123. Morocco: Abeino, 15 km N Goulimine; HT; ♂; DEIC. 

† 

pinites. Upper Eocene - Baltic amber. 

Lestremia pinites Meunier 1904: 42. Baltic amber; ST; ♂,♀; lost. 

sanctijohanni. India (Uttar Pradesh). 

Lestremia sanctijohanni (as sancti-johanni) Rao 1951: 109. India: Uttar Pradesh, Agra; HT; ♀; ZSCI. 

Lestremia sanctijohani (as sancti-johani) Rao & Adwant 1970: 113, missp. of sanctijohanni. 

solidaginis. USA (Minnesota, Wisconsin, New York), Sweden. Ref.: Jaschhof & Jaschhof 2009a (notes). 

Catocha solidaginis Felt 1907a: 6. USA: New York, Newport; HT; ♂; NYSM. 

ugandae. Uganda. 

Lestremia ugandae Barnes 1936: 279. Uganda: Bukumbi; ST; ♂; BMNH. 

MANGOGROSTIX 

Mangogrostix Mamaev 1985: 27. Type species, Gongromastix orientalis Grover (orig. des.). 

Oriental; 1 sp. 


Gongromastix orientalis Grover 1964a: 112. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

NEOLESTREMIA 

Neolestremia Mani 1934b: 380. Type species, boerhaaviae Mani (orig. des.).

Oriental; 3 spp. 

boerhaaviae. India (Tamil Nadu). 

Neolestremia boerhaaviae Mani 1934b: 380. India: Tamil Nadu, Madras, Madras Medical College Grounds; HT; ♀ 

(HT given as ♂ but only ♀ descr.); MANC. 

longipalpia. India (Maharashtra). 

Neolestremia longipalpia Deshpande, Shaikh & Sharma 2002a: 817. India: Maharashtra, Nanded District, Bhokar; 

HT; ♂; DESH. 


Neolestremia orientalis Sharma & Rao 1979a: 36. India: Maharashtra, Aurangabad; HT; ♂; RCMU. 

WASMANNIELLA 

Wasmanniella Kieffer 1898f: 49. Type species, aptera Kieffer (mon.). 

Holarctic, Oriental; 3 spp. Refs.: Jaschhof 1998b, Jaschhof & Jaschhof 2009a (revs.). 

aptera. France. 

Wasmanniella aptera Kieffer 1898f: 49. France: Moselle; T; ♀; lost. 

clauda. USA (Minnesota), Russia (Far East)?, Japan (Honshu, Kyushu)?. 

Wasmanniella clauda Pritchard 1951: 254. USA: Minnesota, John Latch St. Pk; HT; ♂; USNM. 


Wasmaniella indica Grover 1970c: 136. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

SUBFAMILY MICROMYINAE 

ACOENONIA 

Acoenonia Pritchard 1947a: 14. Type species, perissa Pritchard (orig. des.). 

Holarctic; 4 spp.; Acoenoniini. Ref.: Jaschhof 1998b (rev.), Jaschhof & Jaschhof 2009a (redescr.). 

europaea. Sweden, Finland, Latvia, Germany, Ukraine, Russia (Europe). 

Acoenonia europaea Mamaev 1964d: 132. Russia: nr Ribinsk; HT; ♂; ZMMU. 

Acoenonia parvolobata Mamaev & Berest 1992: 69. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; HYNM. 

nana. Germany, Canada (Northwest Terr.), USA (Oregon). 

Acoenonia nana Meyer & Spungis 1994: 434. Germany: SW Kiel; HT; ♂; LASS. 

perissa. USA (Minnesota), Germany. 

Acoenonia perissa Pritchard 1947a: 15. USA: Minnesota, Anoka, along Coon Creek; HT; ♂; UMSP. 

*sidorenkoi. Russia (Far East). 

Acoenonia sidorenkoi Fedotova 2004i: 569. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

AMEDIA 

Amedia Jaschhof 1997b: 523. Type species, floridana Jaschhof (orig. des.). 

Nearctic; 1 sp.; Strobliellini. 

floridana. USA (Florida). 

Amedia floridana Jaschhof 1997b: 524. USA: Florida, Gainesville; HT; ♂; USNM. 

13

*AMEDIELLA 

Amediella Jaschhof 2003b: 106. Type species, involuta Jaschhof (orig. des.). 

Australasian; 1 sp.; Strobliellini. 

*involuta. New Zealand. 

Amediella involuta Jaschhof 2003b: 106. New Zealand: North I., Auckland, Kawau I.; HT; ♂; NZAC. 

ANODONTOCERAS 

Anodontoceras Yukawa 1967b: 194. Type species, saigusai Yukawa (orig. des.). 

Palearctic, Afrotropical, Oriental; 3 spp.; Micromyini. Ref.: Jaschhof 1998b (rev.). 

*harrisi. South Africa. 

Anodontoceras harrisi Jaschhof & Jaschhof 2009b: 118. South Africa: KwaZulu-Natal, Royal Natal Natl Pk, Gudu 

For.; HT; ♂; DEIC. 

saigusai. Russia (Far East), Japan (widespread), Malaysia. 

Anodontoceras saigusai Yukawa 1967b: 196. Japan: Ku, Fukuoka Pref., Mt Inunaki; HT; ♂; KUEC. Ref.: Jaschhof 

& Jaschhof 2009b: 118 (descr. ♀). 

yukawai. Japan (Kyushu, Okinawa), Malaysia. 

Anodontoceras yukawai Jaschhof 1998c: 322. Malaysia: Selangor, Ulu Gombak; HT; ♂; NHRS. 

*ANSIFERA 

Ansifera Jaschhof 2009c: 513. Type species, japonica Jaschhof (orig. des.). 

Palearctic, Oriental, 5 spp.; Campylomyzini. 

*asetosa. West Malaysia. 

Ansifera asetosa Jaschhof 2009c: 522. West Malaysia: Selangor, Ulu Gombak, nr Univ. of Malaya Field Studies 

Centre; HT; ♂; NHRS. 

*gombakensis. West Malaysia. 

Ansifera gombakensis Jaschhof 2009c: 520. West Malaysia: Selangor, Ulu Gombak, nr Univ. of Malaya Field Studies 


*japonica. Japan (Hokkaido). 

Ansifera japonica Jaschhof 2009c: 516. Japan: Hokkaido, Sapporo, Toyohira; HT; ♂; DEIC. 

*longipalpus. West Malaysia. 

Ansifera longipalpus Jaschhof 2009c: 523. West Malaysia: Pahang, Genting Highlands; HT; ♂; NHRS. 

*malayensis. West Malaysia. 

Ansifera malayensis Jaschhof 2009c: 518. West Malaysia: Selangor, Ulu Gombak, nr Univ. of Malaya Field Studies 


APRIONUS 

Apriona Kieffer 1894e: clxxvi, preocc. Chevrolat 1852. Type species, Aprionus spiniger Kieffer (subs. des. Kieffer 

1895: 95). 

Aprionus Kieffer 1894i: 205, new name for Apriona Kieffer. 

Apriocryptus Mamaev & Berest 1992: 67, as subg. of Aprionus. Type species, Aprionus carpathicus Mamaev & 

Berest (orig. des.). 

Apionus Berest 1992: 33, missp. of Aprionus. 

14

Crenuatus Berest 1992: 33, as subg. of Aprionus. Type species, Aprionus confusus Mamaev (orig. des.). 

Operirus Berest 1992: 33, as subg. of Aprionus. Type species, Aprionus cardiophorus Mamaev (orig. des.). 

Subapionus Mamaev 1998: 4, nomen nudum. 

Porianus Mamaev 1998: 5, nomen nudum. 

Holarctic, Oriental, New Zealand; 120 spp.; Aprionini. Refs.: Jaschhof 1998b (rev.), Jaschhof & Jaschhof 2009a 

(rev.). 

aberrantis. Russia (Europe). 

Aprionus aberrantis Mamaev 1998a: 1. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

abiskoensis. Sweden. 

Aprionus abiskoensis Jaschhof 1996b: 342. Sweden: Torne Lappmark, Abisko; HT; ♂; DEIC. 

accipitris. Sweden, Finland, Estonia, Latvia, Germany. 

Aprionus accipitris Jaschhof 1997c: 114. Germany: Potthagen, 6 km S Greifswald; HT; ♂; DEIC. 

aciculatus. Russia (Far East). 

Aprionus aciculatus Mamaev 1997b: 4. Russia: Khasan Region, Zanadvornaya; HT; ♂; ZMMU. 

acutus. UK, Denmark, Sweden, Finland, Russia (Europe), Latvia, Germany. 

Aprionus acutus Edwards 1938h: 233. UK: England, Herts, Hitch Wood; HT; ♂; BMNH. 

*adventitius. Sweden, Finland, Russia (Europe). 

Aprionus adventitius Jaschhof 2009a: 275. Finland: Ostrobottnia kajanensis, Hiidenportti Natl Pk, Urpovaara; HT; ♂; 

NHRS. 

angeloides. Sweden. 

Aprionus angeloides Jaschhof 1997c: 115. Sweden: Pite Lappmark, Arjeplog, Laisvik, Vuordnats; HT; ♂; NHRS. 

angulatus. Sweden, Russia (Europe). 

Aprionus angulatus Mamaev 1963a: 447. Russia: Moscow Prov., Glubokoye L.; HT; ♂; ZMMU. 

arcticus. Fennoscandia, Germany, Russia (Western Siberia). 

Aprionus arcticus Mamaev 2001b: 1. Russia: Northern Urals, Denezhkin Kamen'; ♂; ZMMU. 

*aquilonius. Finland. 

Aprionus aquilonius Jaschhof 2009a: 277. Finland: Tavastia borealis, Saarijärvi, W Saaripuro; HT; ♂; NHRS. 

asemus. Canada (Ontario), USA (Minnesota), Sweden. 

Aprionus asemus Pritchard 1947b: 72. USA: Minnesota, Bayport; HT; ♂; UMSP. 

aviarius. Fennoscandia, Estonia, Latvia, Germany, Ukraine, Russia (Far East). 

Aprionus aviarius Mamaev & Berest 1990a: 23. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; ZMMU. Ref.: 

Jaschhof & Jaschhof 2009a: 229 (redescr.). 

Aprionus ungulatus Jaschhof 1997c: 121. Germany: NSG Galenbecker See, 11 km SE Friedland; HT; ♂; DEIC. 

Aprionus bicruris Mamaev 1998a: 1. Russia: Kuril Is, Kunashir; HT; ♂; ZMMU. 

barbatus. Russia (Europe, Far East). 

Aprionus barbatus Mamaev 1963a: 450. Russia: Vologda Prov., Kadnikovskaya; HT; ♂; ZMMU. 

*berestae. Russia (Far East). 

Aprionus berestae Fedotova 2004i: 569. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

betulae. Fennoscandia, Estonia, Latvia, Germany, Russia (widespread). 

Aprionus betulae Jaschhof 1996b: 343. Norway: Finnmark, Varanger-Halbinsel, Vadsø; HT; ♂; DEIC. 

bicorniger. Russia (Far East). 

Aprionus bicorniger Mamaev 1998a: 1. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

bidentatus. Sweden, Finland, Netherlands, France, Germany, Latvia. 

Apriona bidentatus Kieffer 1894e: clxxvi. France: Bitche; NT (Jaschhof 1998b: 342; Germany: NSG Galenbecker 

See, 11 km SE Friedland); ♂; BMNH. 

bifidus. Northern Europe (widespread), Germany, Russia (widespread), USA (Minnesota, Maine). 

Aprionus bifidus Mamaev 1963a: 450. Russia: Yaroslavl Prov., Volkovo; HT; ♂; ZMMU. 

bifurcatus. China (Inner Mongolia Aut. Reg.). 

Aprionus bifurcatus Bu & Zheng 1996: 224. China: Inner Mongolia Aut. Reg., Ergun Right Banner; HT; ♂; NKUM. 

binarius. Russia (Far East). 

15

16 

Aprionus bifurcatus Mamaev 1997b: 3, jun. prim. hom. bifurcatus Bu & Zheng. Russia: Primorskiï Terr., upper 

reaches Bolshoi Shufen R.; HT; ♂; ZMMU. 

Aprionus binarius Mamaev 1998d: 9, new name for Aprionus bifurcatus Mamaev. 

bispinosus. Northern Europe (widespread), Germany, Ukraine, Russia (widespread). 

Aprionus bispinosus Edwards 1938h: 233. UK: England, Herts, Hitch Wood; HT; ♂; BMNH. 

blanditus. Russia (Western Siberia). 

Aprionus blanditus Mamaev 1998a: 2. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

bostrichus. Denmark, Germany, Ukraine. 

Aprionus bostrichus Berest 1997: 16. Ukraine: Volyn Prov., Kivertsovskiï Reg., Zverevo; HT; ♂; UASK. Ref.: 


Aprionus cariflavidus Jaschhof 1998b: 347. Germany: Elisenhain, 5 km E Greifswald; HT; ♂; DEIC. 

brachypterus. Northern Europe (widespread), Germany, Ukraine, Russia (widespread). 

Aprionus brachypterus Edwards 1938h: 234. UK, England, Cheshire, Longdendale; HT; ♀; BMNH. 

*brevitegminis. Sweden, Finland. 

Aprionus brevitegminis Jaschhof 2009a: 223. Finland: Lapponia kemensis, W part, Kittilä, Pallas-Yllästunturi Natl 

Pk, SE Lehtirova; HT; ♂; NHRS. 

*bullerenis. New Zealand. 

Aprionus bullerenis Jaschhof 2004a: 434. New Zealand: South I., Buller, Nelson Lakes Natl Pk, L. Rotoroa; HT; ♂; 

NZAC. 

cardiophorus. Denmark, Fennoscandia, Estonia, Latvia, Germany, Russia (Europe). 

Aprionus cardiophorus Mamaev 1963a: 448. Russia; Yaroslavl Prov., Rybinskoye Reservoir; HT; ♂; ZMMU. 

carinatus. Northern Europe, Germany, Czech Republic, Ukraine, Russia (Europe). 

Aprionus carinatus Jaschhof 1996b: 345. Sweden: Torne Lappmark, Abisko; HT; ♂; DEIC. 

carpathicus. Ukraine. 

Aprionus carpathicus Mamaev & Berest 1992: 67. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; ZMMU. 

caucasicus. Russia (Near East), Germany. 

Aprionus caucasicus Mamaev & Jaschhof 1997: 458. Russia: Krasnodarskiï Terr., Krasnaya Polyana, Medvezi 

Vorota; HT; ♂; ZMMU. 

complicatus. Fennoscandia, Latvia, Germany, Ukraine. 

Aprionus rostratus Mamaev & Berest 1990a: 20, jun. prim. hom. rostratus Yukawa. Ukraine: Zakarpatskaya Prov., 

Rakhov; HT; ♂; HYNM. 

Aprionus complicatus Mamaev & Berest 1996: 11, new name for Aprionus rostratus Mamaev & Berest. 

confusus. Fennoscandia, Estonia, Latvia, Germany, Ukraine, Russia (widespread). 

Aprionus confusus Mamaev 1969a: 360. Russia: Yaroslav Prov., Volkovo; HT; ♂; HYNM. 

congenericus. Russia (Far East). 

Aprionus lobatus Mamaev 1997b: 3, jun. prim. hom. lobatus Berest. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; 

HT; ♂; ZMMU. 

Aprionus congenericus Mamaev 1998a: 9, new name for Aprionus lobatus Mamaev. 

corniculatus. Sweden, Finland, Estonia, Latvia, Germany, Russia (widespread), Japan (Hokkaido). 

Aprionus corniculatus Mamaev 1963a: 448. Russia: Tula Prov., Tula For.; HT; ♂; ZMMU. 

cornutus. Fennoscandia, Estonia, Latvia, Germany, Ukraine, Russia (Europe). 

Aprionus cornutus Berest 1986: 77. Ukraine: Volyn Prov., vic. Ratno; HT; ♂; UASK. 

dalarnensis. Fennoscandia. 

Aprionus dalarnensis Mamaev 1998a: 2. Sweden: Dalarna; HT; ♂; ZMMU. 

delectabilis. Russia (Europe, Far East). 

Aprionus delectabilis Mamaev 1998a: 2. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

demonstrativus. Norway, Sweden, Germany, Russia (Europe). 

Aprionus demonstrativus Mamaev 1998a: 3. Russia: Vologda Prov., Kadnikovskaya; HT; ♂; ZMMU. 

denticulus. Russia (Europe), Germany, Ukraine, Greece. 

Aprionus denticulus Berest 1986: 77. Ukraine: Zhitomir Prov., Korosten Reg., Usomir; HT; ♂; UASK. 

dentifer. Fennoscandia, Estonia, Latvia, Lithuania, Germany, Russia (Europe), Ukraine. 

Aprionus dentifer Mamaev 1965a: 266. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM.

17 

dispar. Sweden, Finland, Russia (Europe), Germany, Ukraine. 

Aprionus dispar Mamaev 1963a: 446. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

dissectus. Sweden, Finland, Russia (Europe), Ukraine. 

Aprionus dissectus Mamaev & Berest 1990a: 23. Ukraine: Zakarpatskaya Prov., Kvasy, Mt Mincul'; HT; ♂; ZMMU. 

duplicatus. Fennoscandia, Germany. 

Aprionus duplicatus Mamaev 1998a: 3. Sweden: Garpenberg; HT; ♂; ZMMU. 

ellipticus. Russia (Far East). 

Aprionus ellipticus Mamaev 1997b: 4. Russia: Primorskiï Terr., upper reaches Bolshoi Shufan R.; HT; ♂; ZMMU. 

ensiferus. Fennoscandia, Latvia, Germany. 

Aprionus ensiferus Jaschhof 1996b: 346. Sweden: Torne Lappmark, Abisko; HT; ♂; DEIC. 

*fennicus. Sweden, Finland, Russia (Europe). 

Aprionus fennicus Jaschhof 2009a: 254. Finland: Karelia borealis, Lieksa, Patvinsuo Natl Pk, Autiovaara; HT; ♂; 

NHRS. 

ferulae. Turkmenistan. 

Aprionus ferulae Mamaev 1998a: 3. Turkmenistan: Badkhyzskiï Res.; HT; ♂; ZMMU. 

flavidus. UK, Sweden, Finland, France, Germany, Latvia, Lithuania, Ukraine, Russia (widespread), Uzbekistan. 

Campylomyza flavida Winnertz 1870b: 13. Germany; NT (Jaschhof 1998b: 352; UK: England, Bucks, Ivinghoe); ♂; 

BMNH. 

Apriona digitata Kieffer 1894e: clxxvi. France: Bitche; ST; ♂,♀,imm.; lost. 

Aprionus aequatus Mamaev 1963c: 74. Uzbekistan: Samarkand, Takhtakaracha Pass; HT; ♂; HYNM. 

flavoscuta. USA (Washington, New York). 

Campylomyza flavoscuta Felt 1907a: 1. USA: New York, Albany; HT; ♂; NYSM. 

*foliosus. Fennoscandia. 

Aprionus foliosus Jaschhof 2009a: 246. Finland: Ostrobottnia borealis, S part, Pudasjärvi, Syöte Natl Pk, Sotivaara; 

HT; ♂; NHRS. 

giganteus. Ukraine. 

Aprionus giganteus Berest 1991a: 104. Ukraine: Kiyev Prov., Belaya Tserkov'; HT; ♂; UASK. 

*gladiator. Finland, Russia (Europe). 

Aprionus gladiator Jaschhof 2009a: 258. Russia: Karelia, Kondopoga, Kivach Strict Nature Res.; HT; ♂; NHRS. 

halteratus. Northern Europe (widespread), Netherlands, Germany, Russia (Europe). 

Campylomyza halterata Zetterstedt 1852: 4351. Sweden: Östergötland, Winnerstad; HT; ♀; MZLU. 

Campylomyza albicauda Winnertz 1870b: 22. Germany; ST; ♂,♀; lost. 

Campylomyza flaviventris Winnertz 1870b: 21. Germany; ST; ♀; lost. 

hamulatus. China (Sichuan). 

Aprionus hamulatus Bu & Zheng 1996: 226. China: Sichuan, Wenchuan Co.; HT; ♂; NKUM. 

*heothinos. Finland, Germany, Russia (Western Siberia). 

Aprionus heothinos Jaschhof 2009a: 279. Finland: Karelia borealis, Lieksa, Jongunjoki Nature Res., 3 km S Telio; 


*hintelmannorum. Sweden, Finland, Russia (Europe). 

Aprionus hintelmannorum Jaschhof 2009a: 265. Sweden: Dalarna, Rättvik, Gåsberget Nature Res., 10 km N Dalfors; 


*hybridus. Sweden, Russia (Europe). 

Aprionus hybridus Jaschhof 2009a: 240. Sweden: Lule Lappmark, Vettasjärvi, E Puolva Nature Res., Annavaara; 


* † improvisus. Rovno amber (Ukraine). 

Aprionus improvisus Fedotova & Perkovsky 2007a: 87. Ukrainian amber: Rovno; HT; ♀; UASK. Misplaced, per M. 

Jaschhof (in litt.). 

*inaccessibilis. Russia (Far East). 

Aprionus inaccessibilis Fedotova 2004i: 572. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

indictus. Russia (Far East). 

Aprionus indictus Mamaev & Jaschhof 1997: 456. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU.

18 

indicus. India (Uttar Pradesh). 

Aprionus indicus Jaiswal 1988c: 142. India: Uttar Pradesh, Himalayas, Garhwal Hills, Kotdwara; HT; ♂; CSIA. 

inquisitor. Norway, Sweden, Estonia, Latvia, Germany, Switzerland, Russia (Europe). 

Aprionus inquisitor Mamaev 1963a: 447. Russia: Yaroslavl Prov., Volkovo; HT; ♂; HYNM. 

insignis. Northern Europe (widespread) Germany, Ukraine, Russia (widespread), Canada (Ontario). 

Aprionus insignis Mamaev 1963a: 450. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

interruptus. Japan (Kyushu). 

Aprionus interruptus Yukawa 1967b: 192. Japan: Kyushu, Fukuoka City; HT; ♂; KUEC. 

*karsios. Sweden, Russia (Europe). 

Aprionus karsios Jaschhof 2009a: 280. Sweden: Dalarna, Rättvik, 5 km NE Rättvik; HT; ♂; NHRS. 

*kivachensis. Sweden, Russia (Europe). 

Aprionus kivachensis Jaschhof 2009a: 266. Russia: Karelia, Kondopoga, Kivach Strict Nature Res.; HT; ♂; NHRS. 

laevis. Sweden, Finland, Russia (Europe), Latvia, Germany, Austria. 

Aprionus laevis Mohrig 1967: 453. Germany: Wrangelsburger For., nr Greifswald; HT; ♀; DEIC. 

lapponicus. Fennoscandia, Estonia, Latvia, Russia (Western Siberia). 

Aprionus lapponicus Jaschhof & Mamaev 1997: 452. Norway: Finnmark, Pasvik-Tal, Vaggatem; HT; ♂; DEIC. Ref.: 


Aprionus gracilipennis Mamaev 2001b: 2. Russia: Northern Urals, Denezhkin Kamen'; ♂; ZMMU. 

laricis. Sweden, Finland, Russia (Western Siberia). 

Aprionus laricis Mamaev & Jaschhof 1997: 460. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

latens. Ukraine. 

Aprionus latens Mamaev & Berest 1990a: 21. Ukraine: Zakarpatskaya, Kvasy, Mt Mincul'; HT; ♂; HYNM. 

*latitegminis. Sweden. 

Aprionus latitegminis Jaschhof 2009a: 225. Sweden: Gästrikland, Hyttön, 20 km SE Gävle; HT; ♂; DEIC. 

lobatus. Ukraine. 

Aprionus lobatus Berest 1986: 75. Ukraine: Volyn Prov., Kivercovskiy Reg., Zverevo; HT; ♂; UASK. 

longicollis. Fennoscandia, Latvia, Lithuania, Germany, Russia (Europe, Western Siberia). 

Aprionus longicollis Mamaev 1963a: 448. Russia: Yaroslavl Prov.; HT; ♂; HYNM. 

longipennis. USA (New York). 

Campylomyza longipennis Felt 1908b: 314. USA: New York, Albany; HT; ♀; NYSM. 

longisetus. Germany. 

Aprionus longisetus Mohrig 1967: 456. Germany: Potthagen, S Greifswald; HT; ♀; SMNS. 

longitegminis. Sweden, Finland, Russia (Europe), Germany, Austria, Japan. 

Aprionus longitegminis Yukawa 1967c: 63. Japan: Kyushu, Fukuoka Pref., Mt Hiko; HT; ♂; KUIC. 

marginatus. Fennoscandia, Russia (Europe). 

Aprionus marginatus Mamaev 1963a: 450. Russia: Moscow Prov., Il'icha For.; HT; ♂; ZMMU. 

miki. Northern Europe (widespread), France, Germany, Russia (widespread). 

Aprionus miki Kieffer 1895l: 96. France: Bitche; LT (Jaschhof 1995: 84); ♂; NYSM. Ref.: Jaschhof & Jaschhof 

2009a: 248 (redescr.). 

Aprionus pullatus Mamaev 1998a: 4. Sweden: Skäralid; HT; ♂; ZMMU. 

Aprionus trunculus Mamaev 1998a: 6. Russia: Krasnodarskiï Terr., Novo-Prokhladnoe; HT; ♂; ZMMU. 

miniusculus. Russia (Far East). 

Aprionus miniusculus Mamaev 1998a: 4. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

miserandus. Russia (Western Siberia). 

Aprionus miserandus Mamaev 2001b: 3. Russia: Northern Urals, Denezhkin Kamen'; ♀; ZMMU. 

monticola. USA (Minnesota, New York, Maine). 

Campylomyza monticola Felt 1920a: 281. USA: New York, Keene Valley; HT; ♂; NYSM. 

multispinosus. Japan (Hokkaido, Kyushu). 

Aprionus multispinosus Yukawa 1971: 40. Japan: Kyushu, Kagoshima Pref., Iriki-Pass; HT; ♂; KUIC. 

*mycophiloides. New Zealand. 

Aprionus mycophiloides Jaschhof 2004a: 436. New Zealand: South I., Buller, Nelson Lakes Natl Pk, L. Rotoiti; HT; 

♂; NZAC.

19 

myrmecophilus. France. 

myrmecophilus Kieffer 1913a: 235 (Aprionus). France: Bitche; T; ♀; lost. A nomen dubium, per M. Jaschhof (in 

litt.). 

*oligodactylus. Sweden, Finland. 

Aprionus oligodactylus Jaschhof 2009a: 252. Finland: Satakunta, Ikaalinen, Seitseminen Natl Pk, Kuru, N Pitkajärvi; 


onychophorus. Ukraine. 

Aprionus onychophorus Berest 1991a: 102. Ukraine: Sumy Prov., Bishkin'; HT; ♂; UASK. 

piceae. Fennoscandia, Germany. 

Aprionus piceae Jaschhof 1997c: 116. Germany: Karbow, 15 km SE Greifswald; HT; ♂; DEIC. 

pigmentalis. Norway, Sweden, Russia (Eastern Siberia). 

Aprionus pigmentalis Mamaev 1998a: 4. Russia: Altaï, L. Teletskoe; HT; ♂; ZMMU. 

pinicorticis. USA (New Jersey). 

Campylomyza pinicorticis Felt 1908b: 315. USA: New Jersey, Riverton; HT; ♀; NYSM. 

*plicatus. Russia (Far East). 

Campylomyza plicata Fedotova 2004i: 574. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

pommeranicus. Germany, Ukraine. 

Aprionus pommeranicus Jaschhof & Mamaev 1997: 461. Germany: Wampener For., 4 km N Greifswald; HT; ♂; 

DEIC. 

*praecipuus. Fennoscandia, Russia (Near East). 

Aprionus praecipuus Jaschhof 2009a: 285. Finland: Satakunta, Ikaalinen, Seitseminen Natl Pk, Kuru, N Pitkajärvi; 


pratincolus. Norway, Estonia, Latvia, Germany. 

Aprionus pratincolus Jaschhof & Meyer 1995: 218. Germany: NSG Galenbecker See & Friedländer Grosse Wiese, 

11 km SE Friedland; HT; ♂; DEIC. 

pseudispar. Sweden, Germany. 

Aprionus pseudispar Jaschhof 1997c: 118. Germany: Steffenshäger For., W Greifswald; HT; ♂; DEIC. 

pyxidiifer. Fennoscandia. 

Aprionus pyxidiifer Mamaev 1998a: 5. Sweden: Knutby; HT; ♂; ZMMU. 

*reduncus. Finland, Russia (Europe). 

Aprionus reduncus Jaschhof 2009a: 268. Finland: Karelia borealis, Lieksa, Patvinsuo Natl Pk, Autiovaara; HT; ♂; 

NHRS. 

*remotus. New Zealand. 

Aprionus remotus Jaschhof 2004a: 437. New Zealand: Auckland I., Laurie Harbor, Terror Cove; HT; ♂; NZAC. 

rostratus. Japan (Kyushu). 

Aprionus rostratus Yukawa 1971: 37. Japan: Kyushu, Fukuoka Pref., Mt Inunaki; HT; ♂; KUIC. 

separatus. Sweden, Finland, Latvia, Germany, Russia (Far East). 

Aprionus separatus Mamaev & Jaschhof 1997: 457. Russia: Primorskiï Terr., Ussuriyskiï Distr., Suputinskiï Res.; 


*sievertorum. Fennoscandia. 

Aprionus sievertorum Jaschhof 2009a: 260. Sweden: Lule Lappmark, Vettasjärvi, E Puolva Nature Res., Annavaara; 


similis. Sweden, Finland, Estonia, Latvia, France, Germany, Ukraine, Russia (Europe, Far East). 

Aprionus similis Mamaev 1963a: 447. Russia: Moscow Prov., Levinskaya; HT; ♂; HYNM. 

smirnovi. Russia (Europe). 

Aprionus smirnovi Mamaev 1961a: 614. Russia: Krasnodarskiï Terr., Novo-Prokhladnoye; HT; ♂; ZMMU. 

spiniferus. Fennoscandia, Germany, Ukraine. 

Aprionus spiniferus Mamaev & Berest 1990a: 19. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; ZMMU. 

spiniger. Widespread Europe and Far East, USA (Maine). 

Apriona spinigera Kieffer 1894e: clxxvi. France: Bitche; NT (Jaschhof 1998b: 401; UK: England, Hants., New 

Forest); ♂; BMNH. Ref.: Jaschhof & Jaschhof 2009a: 282 (redescr.).

20 

Aprionus subanalis Yukawa 1967: 194. Japan: Kyushu, Kurume City, Mt Kôra; HT; ♂; KUEC. 

Aprionus bisectus Mamaev 1998d: 2. Russia: Moscow Prov., Lugovaya; HT; ♂; ZMMU. 

*stiktos. Fennoscandia. 

Aprionus stiktos Jaschhof 2009a: 271. Norway: inner southern Nordland, Saltdal, Lønsdalen, Semska, 10 km S 

Lønsdal; HT; ♂; NHRS. 

stylatus. Russia (Far East). 

Aprionus stylatus Mamaev & Jaschhof 1997: 453. Russia: Primorskiï Terr., Ussuriyskiï Distr., Suputinskiï Res.; HT; 

♂; ZMMU. 

stylifer. Fennoscandia, Gernmany, Russia (Europe). 

Aprionus stylifer Mamaev 1998d: 3. Russia: Moscow Prov., Lugovaya; HT; ♂; ZMMU. Jaschhof 2009a: 242 

(redescr.). 

Aprionus internuntius Jaschhof 2003a: 176. Germany: Saxony, Dresden; HT; ♂; SMTD. 

styloideus. Northern Europe (widespread), Germany, Hungary, Ukraine, Russia (widespread). 

Aprionus styloideus Mamaev & Berest 1990a: 21. Ukraine: Zakarpatskaya Prov., Kvasy, Mt Mincul'; HT; ♂; 

ZMMU. Jaschhof & Jaschhof 2009a: 243 (redescr.). 

Aprionus paludosus Jaschhof & Mamaev 1997: 454. Norway: Finnmark, Kirkenes; HT; ♂; DEIC. 

Aprionus bicuspidatus Mamaev 1998a: 2. Ukraine: Rakhov; HT; ♂; ZMMU. 

subacutus. Denmark, Norway, Sweden, Lithuania, Germany, Russia (Europe, Far East). 

Aprionus subacutus Jaschhof 1997c: 119. Germany: Karbow, 15 km SE Greifswald; HT; ♂; DEIC. 

svecicus. Fennoscandia. 

Aprionus svecicus Jaschhof 1996b: 347. Sweden: Torne Lappmark, Abisko; HT; ♂; DEIC. 

*taigaensis. Sweden, Finland, Russia (Europe). 

Aprionus taigaensis Jaschhof 2009a: 249. Finland: Karelia borealis, Patvinsuo Natl Pk, Autiovaara; HT; ♂; NHRS. 

terrestris. Germany, Russia (Europe). 

Aprionus terrestris Mamaev 1963a: 447. Russia: Tula Prov., Tula For.; HT; ♂; HYNM. 

tiliamcorticis. Sweden, Finland, Russia (Europe, Far East), Latvia, Germany. 

Aprionus tiliamcorticis Mamaev 1963a: 448. Russia: Tula Prov., Tula For.; HT; ♂; HYNM. 

transitivus. Sweden, Russia (Far East). 

Aprionus transitivus Mamaev 1998a: 5. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

transmutatus. Russia (Far East). 

Aprionus transmutatus Mamaev 1998a: 5. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

umbrellus. Ukraine. 

Aprionus umbrellus Mamaev & Berest 1990a: 18. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

*victoriae. Sweden, Finland. 

Aprionus victoriae Jaschhof 2009a: 261. Finland: Karelia borealis, Lieksa, Koivusuo Strict Nature Res.; HT; ♂; 

NHRS. 

* † vlaskini. Rovno amber (Ukraine). 

Aprionus vlaskini Fedotova & Perkovsky 2007a: 89. Ukrainian amber: Rovno; HT; ♀; UASK. Misplaced, per M. 


wildeni. Spain. 

Aprionus wildeni Jaschhof 1997c: 122. Spain: Andalusia, Jimena de la Frontera; HT; ♂; DEIC. 

BRYOMYIA 

Bryomyia Kieffer 1895i: 61. Type species, bergrothi Kieffer (subs. des. Kieffer 1895k: 78). 

Tomonomyia Berest 1992: 31, as subg. of Bryomyia. Type species, Bryomyia gibbosa Felt (orig. des.). 

Holarctic, Oriental; 8 spp.; Bryomyiini. Ref.: Jaschhof 1998b (rev.), Jaschhof & Jaschhof 2009a (redescr.). 

amurensis. Russia (Far East), Japan (Hokkaido). 

Bryomyia amurensis Mamaev & Økland 1998: 148. Russia: Bychiha, Khekhzir Res.; HT; ♂; ZMMU. 

apsectra. Widespread Palearctic.

21 

Bryomyia apsectra Edwards 1938g: 210. UK: England, Herts, Hitch Wood; HT; ♂; BMNH. Ref.: Jaschhof & 

Jaschhof 2009a: 183 (restored from syn'y, redescr.). 

Bryomyia floccosa Fedotova 2004i: 585. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

bergrothi. Widespread Europe, Russia (Western Siberia, Near East), China. 

Bryomyia bergrothi Kieffer 1895k: 78. France: Bitche; LT (Jaschhof 1995: 87); ♂; NYSM. Jaschhof & Jaschhof 

2009a: 183 (redescr.). 

Bryomyia uralensis Mamaev 2001b: 5. Russia: Northern Urals, Denezhkin Kamen'; ♂; ZMMU. 

gibbosa. Widespread Holarctic. 

Campylomyza gibbosa Felt 1907a: 3. USA: New York, Lake Clear; HT; ♂; NYSM. 

Campylomyza cerasi Felt 1907a: 4. USA: New York, Nassau; HT; ♂; NYSM. 

Neptunimyia flavida Felt 1920a: 279. USA: New York, Keene Valley; HT; ♀; NYSM. 

Bryomyia trifida Edwards 1938g: 209. UK: England, N. Devon, Clovelly; HT; ♂; BMNH. 

Bryomyia incisa Mamaev 1963a: 441. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

Bryomyia longipennis Mamaev 1963a: 441. Russia: Yaroslavl Prov.; HT; ♂; ZMMU. 

Bryomyia montana Mamaev 2001b: 4. Russia: Northern Urals, Denezhkin Kamen'; ♂; ZMMU. 

helmuti. Germany. 

Bryomyia helmuti Jaschhof 1998a: 267. Germany: Karbow, 15 km SE Greifswald; HT; ♂; DEIC. 

multispinata. Ukraine. 

Bryomyia multispinata Berest 1988b: 151. Ukraine: Kiyev Prov., Feofaniya; HT; ♂; UASK. 

* † necessaria. Rovno amber (Ukraine). 

Bryomyia necessaria Fedotova & Perkovsky 2007a: 94. Ukrainian amber: Rovno; HT; ♂; UASK. Misplaced, per M. 


producta. Widespread Holarctic. 

Campylomyza producta Felt 1908b: 315. USA: New York, Nassau; HT; ♂; NYSM. 

CALOSPATHA 

Calospatha Kieffer 1913i: 303. Type species, Calospatha fagicola Kieffer (orig. des.). 

Western Palearctic; 1 sp., a nomen dubium, per M. Jaschhof (in litt.). 

fagicola. France. 

fagicola Kieffer 1913i: 303 (Calospatha). France: Moselle; T; LT; lost. 

CAMPYLOMYZA 

Campylomyza Meigen 1818: 101. Type species, flavipes Meigen (subs. des. Westwood 1840: 126). 

Campolomyia Siebke 1877: 206, missp. of Campylomyza. 

Prionota Kieffer 1894e: clxxvi, preocc. Wulp 1885. Type species, pini Kieffer (mon.) = bicolor Meigen. 

Prionella Kieffer 1894i: 205, poss. as new name for Prionota Kieffer but not so stated, preocc. Robineau-Desvoidy 

1830. Type species, Prionota pini Kieffer (subs. des. Pritchard 1947a: 17) = bicolor Meigen. 

Prionellus Kieffer 1895l: 91, new name for Prionella Kieffer. 

Amblyspatha Kieffer 1913f: 52. Type species, ormerodi Kieffer (subs. des. Edwards 1938: 49). 

Cylophora Kieffer 1913f: 55. Type species, fasciata Kieffer (mon.) = bicolor Meigen. 

Urosema Kieffer 1913i: 293. Type species, Prionellus villosus Kieffer (mon.). 

Holarctic, Neotropical, Oriental, Australasian; 39 spp.; Campylomyzini. Ref.: Jaschhof 1998b (rev.), Jaschhof & 

Jaschhof 2009a (redescr.). 

*abbreviata. Fennoscandia. 

Campylomyza abbreviata Jaschhof 2009a: 110. Sweden: Uppland, Östhammar, Andersby Nature Res., SW

22 

Österbybruk; HT; ♂; NHRS. 

abjecta. Russia (Far East). 

Campylomyza abjecta Mamaev 1998a: 6. Russia: Primorskiï Terr., nr Spasska; HT; ♂; ZMMU. 

aborigena. Russia (Far East). 

Campylomyza aborigena Mamaev 1998a: 6. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

aemula. UK, Fennoscandia, Germany, Russia (Europe). 

Campylomyza aemula Mamaev 1998a: 6. Russia: Voronezh Prov., Tellerman For.; HT; ♂; ZMMU. 

*alnea. Sweden, Germany. 

Campylomyza alnea Jaschhof 2009a: 111. Sweden: Uppland, Knivsta, Rickebasta; HT; ♂; NHRS. 

alpina. UK, Fennoscandia, China, Japan (Hokkaido). 

Campylomyza alpina Siebke 1863: 71 (publ'n not seen). Norway: Oppland, Dovrefjell; HT; ♂; MZLU. 

Cordylomyia pinetorum Edwards 1938g: 202. UK: England, Devon (Sidbury), Herts (Hitch Wood and Sharrard's 

Wood), Bucks (Ivinghoe Common), Lancashire (Low Wood & Manchester), and Scotland (Cromarty); ST; 

♂,♀; BMNH. 

*arcuata. Fennoscandia, Germany. 

Campylomyza arcuata Jaschhof 2009a: 118. Finland: Karelia borealis, Lieksa, Koivusuo Strict Nature Res., S Pirhu; 


armata. Fennoscandia, Estonia, Latvia, Netherlands, Germany, Russia (Europe). 

Campylomyza armata Mamaev 1963a: 452. Russia: Voronezh Prov., Tellerman For.; HT; ♂; ZMMU. 

bicolor. UK, Sweden, Netherlands, France, Germany, Latvia. 

Campylomyza bicolor Meigen 1818: 102. Germany; T; ♀; NMHN. 

Prionota pini Kieffer 1894e: clxxvi. France: Bitche; ST; ♂,♀; lost. 

Cylophora fasciata Kieffer 1913f: 55. France: Moulins; T; ♀; lost. 

*borealis. Norway, Sweden. 

Campylomyza borealis Jaschhof 2009a: 104. Norway, northern Finnmark, Varanger peninsula, mouth of Vesterelva 

W Kroknes NW Vardø; HT; ♂; DEIC. 

cavitata. Sweden, Finland, Germany, Russia (Europe). 

Campylomyza cavitata Mamaev 1998a: 7. Russia: Moscow Prov., Lugovaya; HT; ♂; ZMMU. 

*cingulata. Fennoscandia, Germany. 

Campylomyza cingulata Jaschhof 2009a: 119. Finland: Tavastia borealis, Saarijärvi, W Saaripuro; HT; ♂; NHRS. 

cornuta. Germany, Lithuania. 

Campylomyza cornuta Jaschhof 1998a: 260. Germany: Klepelshagen, 6 km N Strasburg; HT; ♂; DEIC. 

coronoidea. Sweden, Latvia. 

Campylomyza coronoidea Jaschhof 1998a: 261. Sweden: Dalarna, W Evertsberg; HT; ♂; DEIC. 

† 

crassitarsis. Upper Eocene - Baltic amber. 

Campylomyza crassitarsis Meunier 1904: 40. Baltic amber; ST; ♂,♀; DGOT?. 

cruciata. Russia (Far East). 

Campylomyza cruciata Mamaev 1998a: 7. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

dilatata. Widespread Holarctic. 

Campylomyza dilatata Felt 1907c: 149. USA: Massachusetts, Worcester; LT (Pritchard 1947a: 27); ♂; NYSM. 

Campylomyza boulderi Felt 1908b: 314. USA: Colorado, Boulder; HT; ♀; NYSM. 

Campylomyza silvana Felt 1908b: 314. Canada: British Columbia, Kokanee Mtn; HT; ♀; NYSM. 

Campylomyza simulator Felt 1908b: 314. Canada: British Columbia, Kaslo; HT; ♀; NYSM. 

Campylomyza tuckeri Felt 1908b: 316. USA: Texas, Plano; HT; ♂; NYSM. 

Prionellus monilis Felt 1913c: 175. USA: Texas, Plano; HT; ♀; NYSM. 

Prionellus boulderensis Felt 1913c: 177, unjust. emend. of boulderi. 

Campylomyza lobata Edwards 1938f: 181. UK: England, Herts, Breachwood Green; HT; ♂; BMNH. 

*falcifera. Fennoscandia. 

Campylomyza falcifera Jaschhof 2009a: 105. Sweden: Uppland, Sollentuna, Järvafältet; HT; ♂; NHRS. 

flavipes. Widespread Holarctic, New Zealand. 

Campylomyza flavipes Meigen 1818: 102. Germany; T; A; MNHN. Ref.: Jaschoff & Jaschhof 2003: 108. 

Campylomyza aceris Meigen 1818: 102. Germany; ST; ♂,♀; MNHN.

23 

Campylomyza fuscipes Meigen 1830: 272. Germany; HT; ♂; MNHN. 

Campylomyza pallipes Zetterstedt 1850: 3672. Sweden: Lund; LT (Jaschhof 1998b: 154); ♂; MZLU. 

Campylomyza aequalis Winnertz 1870b: 12. Germany; T; ♂; lost. 

Campylomyza flavicoxa Winnertz 1870b: 15. Germany; HT; ♂; lost. 

Campylomyza pumila Winnertz 1870b: 11. Germany; HT; ♂; lost. 

Campylomyza analis Winnertz 1870b: 11. Germany; T; ♂; lost. 

Campylomyza munda Winnertz 1870b: 18. Germany; T; ♀; lost. 

Campylomyza vivida Winnertz 1870b: 14. Germany; ST; ♂; lost. 

Prionellus minor Kieffer 1895l: 92. France: vic. Guinkirchen; ST; ♂; lost. 

Prionellus strobli Kieffer 1901c: 39. Austria: Styria; ST; ♀; lost. 

Campylomyza balsamicola Felt 1907a: 3. USA: New York, Lake Clear; HT; ♂; NYSM. 

Campylomyza graminea Felt 1907a: 2. USA: New York, Karner; HT; ♂; NYSM. 

Campylomyza karnerensis Felt 1907a: 5. USA: New York, Karner; HT; ♂; NYSM. 

Campylomyza leguminicola Felt 1907a: 2. USA: New York, Albany; HT; ♂; NYSM. 

Campylomyza pomiflorae Felt 1907a: 3. USA: New York, Karner; LT (Pritchard 1947a: 22); ♂; NYSM. 

Campylomyza tsugae Felt 1907a: 5. USA: New York, Lake Clear; HT; ♂; NYSM. 

Campylomyza defectiva Felt 1908b: 314. USA: New York, Newport; HT; ♀; NYSM. 

Campylomyza hesperia Felt 1908b: 315. USA: New York, Newport; LT (Pritchard 1947a: 22); ♂; NYSM. 

Campylomyza pomifolia Felt 1908b: 315. USA: New York, Nassau; LT (Pritchard 1947a: 22); ♂; NYSM. 

Prionellus montana Felt 1913c: 182. USA: Colorado, Boulder; HT; ♂; NYSM. 

Prionellus graminae Felt 1913g: 379, missp. of graminea. 

Monardia lateralis Felt 1914d: 109. USA: Massachusetts, Boston; HT; ♂; NYSM. 

Prionellus eremi Felt 1919b: 219. USA: Colorado, Boulder Co., Brainard L.; HT; ♂; NYSM. 

Monardia seaci Barnes 1927a: 167. UK: England, Kent, Wye; HT; ♂; BMNH. 

Campylomyza edwardsi Nijveldt 1959: 78. Netherlands: Wageningen; HT; ♂; ZMAN. 

Campylomyza moehni Kleesattel 1979: 75. Germany: Baden-Württemberg, Dauborn/Taunus; LT (Jaschhof 1998b: 

155); ♀; SMNS. 

furva. UK, Sweden, Finland, Latvia, Germany, Austria. 

Campylomyza furva Edwards 1938f: 179. UK: Herts, Breachwood Green; HT; ♂; BMNH. 

fusca. Widespread Holarctic. 

Campylomyza fusca Winnertz 1870b: 12. Germany; T; ♂; lost. 

Campylomyza barlowi Felt 1908b: 316. USA: Rhode Island, Kingston; HT; ♂; NYSM. 

Campylomyza latipennis Felt 1908b: 314. USA: New York, Karner; LT (Pritchard 1947a: 25); ♂; NYSM. 

Campylomyza obscura Winnertz 1870b: 19. Germany; HT; ♀; lost. 

Campylomyza populi Felt 1907a: 2. USA: New York, Albany; HT; ♂; NYSM. 

hedysari. Italy.. 

hedysari Kieffer 1913f: 53 (Amblyspatha). Italy: Vallombrosa; T; LT; lost. A nomen dubium, per M. Jaschhof (in 

litt.). 

*hybrida. Sweden, Russia (Europe). 

Campylomyza hybrida Jaschhof 2009a: 96. Sweden: Uppland, Uppsala, Fiby Nature Res.; HT; ♂; NHRS. 

*inornata. Fennoscandia, Germany. 

Campylomyza inornata Jaschhof 2009a: 114. Finland: Karelia ladogensis, Parikkala, Lake Siikalahti, W Kaukola; 


*insolita. Finland, Germany. 

Campylomyza insolita Jaschhof 2009a: 100. Germany: Mecklenburg-Vorpommern, Buddenhagen, 8km SW Wolgast; 

HT; ♂; DEIC. 

*mohrigi. Sweden, Germany, Switzerland, Italy, Morocco, Turkey, Uzbekistan, Turkmenistan, Russia (Western 

Siberia). 

Campylomyza mohrigi Jaschhof 2009a: 107. Germany: Mecklenburg-Vorpommern, Klepelshagen, 6 km N Strasburg; 


† 

monilifera. Upper Eocene - Baltic amber. 

Campylomyza monilifera Loew 1850b: 32. Baltic amber; T; A; lost?.

24 

Campylomyza globulifera Keilbach 1982: 346, error for monilifera. 

Campylomyza moniliformis Meunier 1899a: 161, error for monilifera. 

mucoris.France. 

mucoris Kieffer 1913f: 53 (Amblyspatha). France: Bitche; T; LT; lost. A nomen dubium, per M. Jaschhof (in litt.). 

nigroliminata. Russia (Europe). 

Campylomyza nigroliminata Mamaev 1998a: 7. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 


Campylomyza orientalis Grover 1970c: 162. India: Uttar Pradesh, Allahabad, Naini; HT; ♂; PGCA. 

ormerodi. UK, Fennoscandia, Germany. 

Amblyspatha ormerodi Kieffer 1913f: 52. UK, Scotland, vic. Edinburgh; NT (Jaschhof 1998b: 165; UK: England, 

Herts, Radwell); ♂; BMNH. 

*paenebicolor. Norway, Sweden, Germany. 

Campylomyza paenebicolor Jaschhof 2009a: 98. Germany: Mecklenburg-Vorpommern, Greifswald; HT; ♂; DEIC. 

*pubescens. Sweden, Germany. 

Campylomyza pubescens Jaschhof 2009a: 116. Sweden: Uppland, Uppsala, Fiby Nature Res.; HT; ♂; NHRS. 

serrata. Widespread northern and western Europe. 

Campylomyza serrata Jaschhof 1998a: 263. Germany: Ladebow, 5 km E Greifswald; HT; ♂; DEIC. 

spatulata. Sweden, Finland. 

Campylomyza spatulata Mamaev 1998a: 8. Sweden: Dalarna; HT; ♂; ZMMU. 

spinata. Russia (Far East). 

Campylomyza spinata Jaschhof 1998a: 264. Russia: Chabarovskiï Terr., Boltsovo, 20 km N Bikin; HT; ♂; DEIC. 

*stegetfore. Fennoscandia. 

Campylomyza stegetfore Jaschhof 2009a: 94. Sweden: Lule Lappmark, Vettasjärvi, Haapavaara/Ammavaara; HT; ♂; 

NHRS. 

tridentata. Sweden, Germany, Latvia, Estonia, Russia (Europe). 

Campylomyza tridentata Jaschhof 1998a: 266. Sweden: Dalarna, W Evertsberg; HT; ♂; DEIC. 

villosa. Romania. 

Prionellus villosus Kieffer 1901c: 40. Romania; T; ♀; lost. Misplaced, per Jaschhof (in litt.). 

CATARETE 

Catarete Edwards 1929a: 11. Type species, Lestremia brevinervis Zetterstedt (orig. des.). 

Western Palearctic; 2 spp.; Catochini. Ref.: Jaschhof & Jaschhof 2009a (rev.). 

brevinervis. Sweden. 

Lestremia brevinervis Zetterstedt 1851: 3770. Sweden: Åsele Lappmark, Vilhelmina; HT; ♀?; MZLU. 

*minuta. Sweden, Finland. 

Catarete minuta Jaschhof 2009a: 76. Sweden: Västerbotten, Skellefteå, Stensträsk, Björnhultet State For. Res.; HT; 

♂; NHRS. 

CATOCHA 

Catocha Haliday 1833: 156. Type species, latipes Haliday (mon.). 

Mimosciara Rondani 1840: 25. Type species, molobrina Rondani (subs. des. Coquillett 1910: 570) = latipes Haliday. 

Furcinerva Rondani 1846: 369. Type species, Mimosciara molobrina Rondani (subs. des. Coquillett 1910: 570) = 

latipes Haliday. 

Macrostyla Winnertz 1846: 20. Type species, latipes Winnertz (mon.) = latipes Haliday. 

Molobraea Loew 1850a: 12. Type species, Mimosciara molobrina Rondani (subs. des. Pritchard 1948: 667) = latipes 

Haliday. 

Yposatoea Rondani 1856: 198. Type species, Mimosciara molobrina Rondani (subs. des. Coquillett 1910: 621) = 

latipes Haliday.

Eucatocha Edwards 1938e: 106. Type species, Catocha barberi Felt (orig. des.). 

Anocha Pritchard 1948a: 669. Type species, Neocatocha spinosa Felt (orig. des.). 

Holarctic, Oriental; 9 spp.; Catochini. Refs.: Jaschhof 1998b (rev.), Jaschhof & Jaschhof 2009a (redescr.). 

*angulata. Denmark, Sweden. 

Catocha anglata Jaschhof 2009a: 79. Sweden: Uppland, Uppsala, Fiby Nature Res.; HT; ♂; NHRS. 

barberi. Canada (Yukon Terr.), USA (Wisconsin, Minnesota). 

Catocha barberi Felt 1913c: 131. USA: Wisconsin, Vilas Co., Crab L.; LT (Pritchard 1948a: 668); ♂; USNM. Felt 

1913c wrote that "two midges were taken" and he purportedly described a female. Both specimens in the 

USNM are males and the antenna described would fit that of a male, so Felt's "female" at the beginning of his 

description was evidently in error. 

betsyae. USA (California). 

Eucatocha betsyae Pritchard 1960c: 195. USA: California, Tuolumne Co., Strawberry; HT; ♀; CASC. 

celesteana. USA (Wyoming). 

Anocha celesteana Pritchard 1960c: 197. USA: Wyoming, Cheyenne; HT; ♀; CASC. 

*incisa. Finland. 

Catocha incisa Jaschhof 2009a: 79. Finland: Tavastia borealis, Saarijärvi, Pyhä-Häkki Natl Pk; HT; ♂; NHRS. 

indica. Pakistan. 

Catocha indica Mani 1934b: 378. Pakistan: North-West Frontier Prov., Chitral, Lutkoh Valley; ST; ♀; ZSCI. 

latipes. Widespread Holarctic, Taiwan. Ref.: Jaschhof & Jaschhof 2009a (redescr.). 

Catocha latipes Haliday 1833: 156. UK: Northern Ireland, Down Co., Holywood; T; A; NMID. 

Mimosciara lestremina Rondani 1840: 26. Italy: vic. Parma; T; A; MZUF. 

Mimosciara molobrina Rondani 1840: 25. Italy: vic. Parma; T; A; MZUF. 

Macrostyla latipes Winnertz 1846: 20. Germany; HT; ♀; lost. 

Catocha crassitarsis Wulp 1874: 113. Netherlands: Rotterdam; HT; ♀; ZMAN. 

Catocha muscicola Kieffer 1898f: 52, nomen nudum. 

Catocha muscicola Kieffer 1901b: 453. France: Moselle; T; ♀; lost. 

Catocha slossonae Felt 1908b: 309. USA: New Hampshire, Franconia; HT; ♂; USNM. 

Catocha kiefferi Strobl 1910: 231. Austria: Zirbitzkogel und Seitenstetten; ST; ♂; lost. 

Neocatocha sylvana Felt 1920a: 280. USA: New York, Keene Valley; HT; ♀; NYSM. 

spinosa. USA (Minnesota, Maine), Czech Republic. 

Neocatocha spinosa Felt 1913c: 152. USA; HT; ♀; USNM. 

Neocatocha nylanderi Felt 1926c: 207. USA: Maine, Woodland; HT; ♀; NYSM. 

*subalpina. Norway, Sweden. 

Catocha subalpina Jaschhof 2009a: 84. Sweden: Pite Lappmark, Arjeplog, Kungsleden SW Jäkkvik; HT; ♂; NHRS. 

CORINTHOMYIA 

Corinthomyia Felt 1911c: 35. Type species, Campylomyza hirsuta Felt (orig. des.) = brevicornis Felt. 

Holarctic, Neotropical; 1 sp.; Campylomyzini. Ref.: Jaschhof 1998b (redescr.). 

brevicornis. Canada (British Columbia, Newfoundland), USA (widespread), Chile, UK, Norway, Sweden, Austria, 

Russia (Far East). 

Campylomyza brevicornis Felt 1907a: 1. USA: New York, Nassau; HT; ♀; NYSM. 

Campylomyza bryanti Felt 1908b: 313. Canada: Newfoundland, Little River; HT; ♀; USNM. 

Campylomyza currei Felt 1908b: 315. Canada: British Columbia, Kaslo; HT; ♂; USNM. 

Campylomyza hirsuta Felt 1908b: 315. USA: New York, Albany; HT; ♂; NYSM. 

Campylomyza luna Felt 1908b: 313. USA: New York, Westfield; HT; ♀; NYSM. 

Corinthomyia gracilis Felt 1912f: 102. USA: Pennsylvania, Hazelton; HT; ♂; NYSM. 

Corinthomyia cincinna Felt 1913c: 200, also p. 201 as gracilis. USA: New York, Albany; HT; ♂; NYSM. 

25

Corinthomyia tumida Felt 1913c: 197. USA: New York, Albany; HT; ♀; NYSM. 

Corinthomyia curreyi Foote 1965: 249, missp. of currei. 

CRETOCATOCHA 

Cretocatocha Gagné 1977a: 58. Type species, mcalpinei Gagné (orig. des.). 

Canadian Cretaceous amber; 1 sp.; Catochini. 

†mcalpinei. Upper Cretaceous amber - Canada (Alberta). 

Cretocatocha mcalpinei Gagné 1977a: 58. Upper Cretaceous amber, Canada: Alberta, nr Medicine Hat; HT; ♂; 

CNCI. 

CRETOCORDYLOMYIA 

Cretocordylomyia Gagné 1977a: 60. Type species, quadriseries Gagné (orig. des.). 

Canadian Cretaceous amber; 1 sp.; Campylomyzini. 

†quadriseries. Upper Cretaceous amber - Canada (Manitoba). 

Cretocordylomyia quadriseries Gagné 1977a: 60. Upper Cretaceous amber, Canada: Manitoba, Cedar L.; HT; ♂; 

MCZ. 

ELENIELLA 

Eleniella Berest 2001: 72. Type species, kyseluci Berest (orig. des.). 

Palearctic; 1 sp.; Strobliellini. 

kyseluci. Ukraine. 

Eleniella kyseluci Berest 2001: 72. Ukraine: Ivalo-Frankovskaya Prov., Yaremcha, Mikulichin, Carpathian Natl 

Nature Pk; HT; ♂; SIZK. 

ELTXO 

Eltxo Arillo & Nel 2000: 285. Type species, cretaceus Arillo & Nel (orig. des.). 

Spanish amber, Lower Cretaceous; 1 sp.; Strobliellini. 

† cretaceus. Lower Cretaceous amber - Spain. 

Eltxo cretaceus Arillo & Nel 2000: 285. Lower Cretaceous amber, Spain: Alava, nr Peñacerrada; HT; ♂; MCNA. 

EXCRESCENTIA 

Excrescentia Mamaev & Berest 1991b: 75. Type species, mutuata Mamaev & Berest (orig. des.). 

Western Palearctic; 1 sp.; Campylomyzini. 

mutuata. Norway, Sweden, Finland, Latvia, Germany, Ukraine, Russia (Europe). 

Excrescentia mutuata Mamaev & Berest 1991b: 75. Ukraine: Zakarpatskaya Prov., Rakhov Region, Mt Mincul'; HT; 

♂; HYNM. 

26

FORBESOMYIA 

Forbesomyia Malloch 1914: 234. Type species, atra Malloch (orig. des.). 

Nearctic; 1 sp.; Catochini. Ref.: Jaschhof & Jaschhof 2009a (review). 

atra. Canada (British Columbia), USA (Washington, Montana, Illinois). 

Forbesomyia atra Malloch 1914: 235. USA: Illinois, Urbana; HT; ♀; SIUC. 

GAGNEA 

Gagnea Jaschhof 2001b: 38. Type species, tsutaensis Jaschhof (orig. des.). 

Eastern Palearctic; 1 sp.; Peromyiini. 

tsutaensis. Japan (Hokkaido, Honshu). 

Gagnea tsutaensis Jaschhof 2001b: 38. Japan: Honshu, Aomori Pref., Tsuta hot spring; HT; ♂; KUEC. 

GROVERIELLA 

Groveriella Mamaev 1978: 1. Type species, carpathica Mamaev (orig. des.). 

Western Palearctic; 2 spp.; Strobliellini. Ref.: Jaschhof & Jaschhof 2009a (redescr.). 

baltica. Sweden, Estonia, Latvia. 

Groveriella baltica Spungis & Jaschhof 2000: 309. Estonia: Puurmani; HT; ♂; FBUL. 

carpathica. Ukraine. 

Groveriella carpathica Mamaev 1978: 2. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; ZMMU. 

HETEROGENELLA 

Heterogenella Mamaev 1963a: 440. Type species, hybrida Mamaev (orig. des.). 

Cervuatina (also as Gervuatina) Berest 1993: 6. Type species, Bryomyia cambrica Edwards (orig. des.). 

Sidorenkomyia Fedotova 2004i: 585. Type species, aurita Fedotova (orig. des.). 

Holarctic; 13 spp.; Bryomyiini. Ref.: Jaschhof 1998b (rev.), Jaschhof & Jaschhof 2009a (redescr.). 

*aurita. Russia (Far East). 

Sidorenkomyia aurita Fedotova 2004i: 585. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

bigibbata. Ukraine. 

Heterogenella bigibbata Mamaev & Berest 1991a: 29. Ukraine: Zakarpatskaya Prov., Kvasy, Mt Mincul'; HT; ♂; 

ZMMU. 

Heterogenella monteviva Mamaev & Berest 1991a: 29. Ukraine: Zakarpatskaya Prov., Kvasy, Mt Mincul'; HT; ♂; 

ZMMU. 

californica. USA (California). 

Heterogenella californica Jaschhof 1997b: 532. USA: California, Inverness; HT; ♂; USNM. 

cambrica. Widespread Europe. 

Bryomyia cambrica Edwards 1938g: 210. UK, England, Brecon, Llangattock; HT; ♂; BMNH. Ref.: Jaschhof & 

Jaschhof 2009a (redescr.). 

Heterogenella miscella Mamaev 1998d: 2. Italy: Siena; HT; ♂; ZMMU. 

27

28 

*Heterogenella biradiata Mamaev & Berest 2005: 83. Ukraine: Zakarpatskaya Prov., Rakhiv; HT; ♂; ZMMU. 

dolini. Ukraine, China (Sichuan). 

Heterogenella dolini Berest 1989a: 84. Ukraine: Crimea, Alma R. valley; HT; ♂; UASK. 

finitima. Sweden, Finland, Estonia, Latvia, Germany, Russia (Far East). 

Heterogenella finitima Mamaev 1998d: 2. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

Heterogenella multifurcata Spungis & Jaschhof 2000: 309. Estonia: Valga; HT; ♂; FBUL. 

hybrida. Fennoscandia, Baltic countries, Germany, Russia (Europe), China (Inner Mongolian Autonomous Region). 

Heterogenella hybrida Mamaev 1963a: 440. Russia: Vologda Prov., Kaldikovskaya; HT; ♂; HYNM. 

linearis. Sweden, Germany, Ukraine, Russia (Far East), China (Yunnan), Japan (Honshu). 

Heterogenella linearis Yukawa 1971: 51. Japan: Honshu, Tottori Pref., Mt Daisen; HT; ♂; KUEC. Ref.: Jaschhof & 

Jaschhof 2009a: 190 (redescr.). 

Heterogenella carinata Mamaev & Berest 1991a: 27. Ukraine: Zakarpatskaya Prov., Kvasy, Mt Mincul'; HT; ♂; 

ZMMU. 

Neurolyga punctata Fedotova 2004i: 576. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

Campylomyza subsicca Fedotova 2004i: 574. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

Neurolyga vasilii Fedotova 2004i: 629. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

mamajevi. Japan (Kyushu). 

Heterogenella mamajevi Yukawa 1967a: 22. Japan: Kyushu, Fukuoka Pref., Mt Inunaki; HT; ♂; KUIC. 

*minuta. Russia (Europe), Germany. 

Heterogenella minuta Jaschhof 2009a: 191. Russia: Karelia, Kondopoga, Kivach Strict Nature Res.; HT; ♂; NHRS. 

puberula. China (Guangdong). 

Bryomyia puberula Li & Bu 2001b: 98. China: Guangdong, Fengkai; HT; ♂; NKUM. 

transgressoris. Norway, Estonia, Latvia, Germany. 

Heterogenella transgressoris Jaschhof 1998a: 269. Germany: NSG Galenbecker See, 11 km SE Friedland; HT; ♂; 

DEIC. 

transversa. China (Yunnan). 

Heterogenella transversa Li & Bu 2002a: 356. China: Yunnan, Zhongdian, Hutiaoxia; HT; ♂; NKUM. 

MICROMYA 

Micromya Rondani 1840: 23. Type species, lucorum Rondani (mon.). 

Micromyia Anonymous 1844: 451, missp. of Micromya. 

Ceratomyia Felt 1911c: 33. Type species, johannseni Felt (orig. des.) = lucorum Rondani. 

Crespiniella Kieffer 1924: 88. Type species, sahariensis Kieffer (mon.) = lucorum Rondani. 

Holarctic, Neotropical, Oriental; 11 spp.; Micromyini. Ref.: Jaschhof 1998b (rev.). 

brevisegmenta. China (Shandong). 

Micromyia brevisegmenta Mo 1990: 57. China: Shandong, Taian; HT; ♂; SAUT. 

championii. India (Uttar Pradesh). 

Micromyia championii Grover 1962e: 441. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

fusongensis. China (Jilin). 

Micromyia fusongensis Mo 1990: 58. China: Jilin, Fusong, Donggang; HT; ♂; SAUT. 

gurbaxii. India (Uttar Pradesh). 

Micromyia gurbaxii Grover 1970c: 154. India: Uttar Pradesh, Allahabad; HT; ♀; PGCA. 

indica. India (Delhi). 

Micromyia indica Mani 1937: 281. India: Delhi; ST; ♂; ZSCI. 

kyushuensis. Japan (Honshu, Kyushu). 

Micromya kyushuensis Yukawa 1967b: 191. Japan: Kyushu, Kagoshima Pref., Ôdomari; HT; ♂; KUEC. 

longicauda. China (Shandong).

29 

Micromya longicauda Mo 2000a: 209. China: Shandong, Dongying; HT; ♂; SAUT. 

longispina. China (Shandong). 

Micromyia longispina Mo 1990: 59. China: Shandong, Taian; HT; ♂; SAUT. 

lucorum. Widespread Europe, Algeria, Uzbekistan, Russia (Far East), USA (Minnesota), Mexico, Costa Rica, 

Dominica. 

Micromya lucorum Rondani 1840: 23. Italy; NT (Jaschhof 1998b: 242; UK: Oxford); ♂; BMNH. 

Campylomyza globifera Haliday 1856: 62. UK: Northern Ireland, Holywood; ST; ♂,♀; NMID. 

Ceratomyia johannseni Felt 1911c: 33. Mexico: Ocotlán; HT; ♂; NYSM. 

Crespiniella sahariensis Kieffer 1924: 88. Algeria: Oasis de Biskra; ST; ♂,♀; lost. 

Micromya mana Pritchard 1947b: 63. USA: Minnesota, St. Paul; HT; ♂; UMSP. 

Micromyia taurica Mamaev 1963a: 445. Ukraine: Crimea, Anapa; HT; ♂; HYNM. 

orientalis. India (Bihar). 

Micromyia orientalis Grover 1962e: 439. India: Bihar, Mihijam; HT; ♂; PGCA. 

transispina. China (Jilin). 

Micromyia transispina Mo 1990: 59. China: Jilin, Fusong, Donggang; HT; ♂; SAUT. 

MICROPTEROMYIA 

Micropteromyia Mamaev 1960b: 952. Type species, ghilarovi Mamaev (orig. des.). 

Palearctic; 1 sp.; Campylomyzini. Ref.: Jaschhof 1998b (rev.). 

ghilarovi. Russia (Europe). 

Micropteromyia ghilarovi Mamaev 1960b: 954. Russia: Tula Prov., Tula For.; LT (Mamaev 1963a: 453); ♂; IEME. 

MONARDIA 

Monardia Kieffer 1895i: 61. Type species, stirpium Kieffer (subs. des. Kieffer 1895m: 111). 

Pezomyia Kieffer 1913b: 92. Type species, Monardia vanderwulpi Meijere (orig. des.) = stirpium Kieffer. 

Mycopriona Mamaev 1965a: 266. Type species, Monardia caucasica Mamaev (orig. des.). 

Vulcanardia Mamaev 1993: 60. Type species, kamtshatica Mamaev (mon). 

Furcapriona Mamaev & Zaitzev 1996a: 6. Type species, Mycopriona nigrita Mamaev (orig. des.). 

Procnomyia Mamaev & Zaitzev 1996a: 4. Type species, Mycopriona incerta Mamaev (orig. des.) = lignivora Felt. 

Mycoporina Mamaev 1999: 241, missp. of Mycopriona. 

Holarctic, Afrotropical, Oceanian, and Baltic amber; 53 spp. in four subgenera and one group of unplaced species; 

Micromyini. Refs.: Jaschhof 1998b, Jaschhof & Jaschhof 2009a (rev.). 

SUBGENUS ANTENNARDIA 

Antennardia Mamaev 1993: 48. Type species, Antennardia gallicola Mamaev (orig. des.). 

antennata. UK, Germany, Japan, USA (Minnesota). 

Campylomyza antennata Winnertz 1870b: 23. Germany; T; ♀; lost. 

gallicola. Russia (Far East). 

Antennardia gallicola Mamaev 1993: 48. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

*saxonica. Germany. 

Monardia saxonica Jaschhof 2003a: 178. Germany: Saxony, Meissen Distr., 2 km NW Moritzburg; HT; ♂; SMTD. 

SUBGENUS MONARDIA 

abnormis. Denmark, Sweden, Finland, Latvia, Germany, Russia (Europe).

30 

Monardia abnormis Mamaev 1963a: 443. Russia: Yaroslavl Prov., Volkovo; HT; ♂; ZMMU. 

*armata. Sweden, Finland, Germany. 

Monardia armata Jaschhof 2003a: 180. Germany: Saxony, Meissen Distr., Friedewald, 2 km NW Moritzburg; HT; 

♂; SMTD. 

canadensis. Canada (British Columbia), USA (Idaho). 

Monardia canadensis Felt 1926f: 267. Canada: British Columbia, Michel, Wilson Creek; HT; ♀; NYSM. 

caucasica. Russia (Near East). 

Monardia caucasica Mamaev 1963a: 442. Russia: Krasnodarskiï Terr., Dakhavskiï For.; HT; ♂; ZMMU. 

crassicornis. Uzbekistan. 

Monardia crassicornis Mamaev 1963c: 73. Uzbekistan: Samarkand, Takhtakaracha Pass; HT; ♂; ZMMU. 

*dividua. New Zealand. 

Monardia dividua Jaschhof 2004a: 424. New Zealand: North I., Coromandel, Opoutere, Wharekawa Picnic Area; 

HT; ♂; NZAC. 

kamtshatica. Russia (Far East). 

Vulcanardia kamtshatica Mamaev 1993: 60. Russia: Kamchatka, Kozyrevsk; HT; ♂; HYNM. 

kollari. Germany, Austria. 

Campylomyza kollari Winnertz 1870b: 23. Austria; LT (Jaschhof 1998b: 265); ♀; NHMW. 

lignivora. USA (North Carolina), Norway, Finland, Russia (Europe, Far East). 

Campylomyza lignivora Felt 1907a: 4. USA: North Carolina, Davidson's R.; LT (Pritchard 1947: 56); ♂; USNM. 

Mycopriona incerta Mamaev 1993: 50. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. 

Mycopriona lateralis Mamaev 1993: 50. Russia: Moscow Prov., Malinki; HT; ♂; HYNM. 

magna. UK. 

Monardia magna Edwards 1938h: 238. UK: England, Herts, Welwyn, Sherrard's Wood; HT; ♀; BMNH. 

*malaisei. Sweden. 

Monardia malaisei Jaschhof 2009a: 199. Sweden: Uppland, Uppsala, Fiby Nature Res., Trefalddighets källa; HT; ♂; 

NHRS. 

misella. Russia (Europe). 

Mycopriona misella Mamaev 1993: 51. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ZMMU. 

monilicornis. UK, Norway, Sweden, Latvia, Germany, Russia (Europe). 

Cecidomyza monilicornis Zetterstedt 1838: 823. Norway: Lappland, Dovrefjell Mtn; HT; ♀; MZLU. 

multiarticulata. USA (New Hampshire, Maine). 

Monardia multiarticulata Felt 1914d: 109. USA: New Hampshire, Franconia; HT; ♀; NYSM. 

nigrita. Russia (Far East). 

Mycopriona nigrita Mamaev 1993: 51. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

obsoleta. UK, Sweden, Finland, Latvia, Germany, Italy, Russia (Europe, Far East). 

Monardia obsoleta Edwards 1938h: 240. UK: England, Herts, Hitch Wood; HT; ♀; BMNH. 

pediculata. Sweden, Finland, Russia (Europe). 

Mycopriona pediculata Mamaev 1993: 51. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

recondita. Hawaiian Is. 

Monardia recondita Hardy 1960: 252. Hawaiian Is: Oahu, Ewa; HT; ♂; BPBM. 

recta. Sweden, Russia (Europe). 

Mycopriona recta Mamaev 1993: 51. Russia: Voronezh Prov., Tellerman; HT; ♂; ZMMU. 

separata. Russia (Europe). 

Mycopriona separata Mamaev 1993: 52. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

stirpium. UK, Sweden, Netherlands, France, Germany, Ukraine, Russia (Europe, Far East), Uzbekistan, Japan, New 

Zealand. 

Monardia stirpium Kieffer 1895m: 111. France: vic. Bitche; ST; ♂,♀,P,L; NYSM. Ref.: Jaschhof & Jaschhof 2009a: 

202 (redescr.). 

Monardia vanderwulpi (as van der wulpi) Meijere 1899: 140. Netherlands: Zuid-Holland, Zwammerdam; LT 

(Jaschhof 1998b: 280); ♂; ZMAN. 

Monardia longicauda Mamaev 1963a: 444. Uzbekistan: Samarkand, Takhtakaracha Pass; HT; ♂; ZMMU. 

† 

submonilifera. Upper Eocene - Baltic amber.

31 

Monardia submonilifera Meunier 1904: 39. Baltic amber; ST; ♂,♀; lost. 

ulmaria. UK, Ukraine. 

Monardia ulmaria Edwards 1938h: 238. UK: England, Bucks, Chesham; HT; ♀; BMNH. 

yasumatsui. Finland, Germany, Russia (Near East), Japan (Kyushu). 

Monardia yasumatsui Yukawa 1967b: 199. Japan: Kyushu, Fukuoka Pref., Mt Inunaki; HT; ♂; KUIC. 

Mycopriona holoptica Mamaev 1993: 50. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

Mycopriona ornata Mamaev 1993: 51. Russia: Krasnodarskiï Terr., Krasnaya Polyana, Medvezh'i Vorota; HT; ♂; 

ZMMU. 

Mycopriona acuta Mamaev & Zaitzev 1996a: 3. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

SUBGENUS TRICHOPTEROMYIA 

Trichopteromyia Williston 1896: 255. Type species, modesta Williston (mon.). 

Projoannisa Kieffer 1912h: 2. Type species, Joannisia latipennis Kieffer (orig. des.) = modesta (Williston). 

absurda. Russia (Far East). 

Trichopteromyia absurda Mamaev 1993: 57. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. 

denticauda. USA (Ohio, Virginia). 

Trichopteromyia denticauda Gagné 1985a: 131. USA: Ohio, Canal Fulton; HT; ♂; USNM. 

*dissecta. Russia (Far East). 

Polyardis dissecta Fedotova 2004i: 569. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

gracilis. Russia (Europe). 

Trichopteromyia gracilis Mamaev 1993: 58. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

magnifica. Fennoscandia, Germany, Russia (Europe, Far East). 

Trichopteromyia magnifica Mamaev 1963a: 445. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

Trichopteromyia difficilis Mamaev 1993: 57. Russia: Primorskiï Terr.; HT; ♂; ZMMU. 

manii. India (Kerala). 

Trichopteromyia manii Nayar 1945: 69. India: Kerala, Trivandrum; HT; ♀; ETKI. 

modesta. USA (Minnesota, Maryland, New Hampshire), Mexico, Costa Rica, Dominica, St. Vincent, UK, Norway, 

Sweden, Latvia, Germany, Ukraine, Russia (Europe, Far East), Japan, Seychelles Is. 

Trichopteromyia modesta Williston 1896: 255. St. Vincent; ST; ♀; BMNH. 

Joannisia latipennis Kieffer 1911: 328. Seychelles Is: Mahé, Trois Frères; ST; ♀; BMNH. 

Monardia rugosa Felt 1914d: 110. USA: New Hampshire, Hanover; HT; ♀; NYSM. 

Trichopteromyia japonica Yukawa 1967b: 197. Japan: Kyushu, Fukuoka Pref., Mt Inunaki; HT; ♂; KUEC. 

Trichopteromyia bicolorata Mamaev 1993: 57. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

Trichopteromyia carpatica Mamaev 1993: 57. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; ZMMU. 

Trichopteromyia fungicola Mamaev 1993: 58. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

Trichopteromyia kovalevi Mamaev 1993: 57. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

Trichopteromyia pallida Mamaev 1993: 58. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

*relicta. Finland. 

Monardia relicta Jaschhof 2009a: 207. Finland: Tavastia borealis, Saarijärvi, Pyhä-Häkki Natl Pk; HT; ♂; NHRS. 

SUBGENUS XYLOPRIONA 

Xylopriona Kieffer 1913g: 291. Type species, Campylomyza pulchricornis Kieffer (orig. des.) = atra Meigen. 

Tetraxyphus Kieffer 1913g: 290. Type species, Campylomyza melanopterus Kieffer (mon.) = atra Meigen. 

adentis. Sweden, Finland, Germany. 

Xylopriona adentis Jaschhof 1998a: 271. Germany: Thüringen, Wittersroda, 15 km W Kahla; HT; ♂; DEIC. 

* † aristata. Rovno amber (Ukraine). 

Xylopriona aristata Fedotova & Perkovsky 2007a: 86. Ukrainian amber: Rovno; HT; ♂; UASK. Misplaced, per M. 

Jaschhof (in litt.).

32 

articulosa. USA (Virginia, New Hampshire). 

Campylomyza articulosa Felt 1908b: 315. USA: New Hampshire, White Mtns; HT; ♀; USNM. 

atra. Widespread Palearctic, USA (California). 

Cecidomyia atra Meigen 1804: 40. Germany; HT; ♂; MNHM. Ref.: Jaschhof & Jaschhof 2009a: 210 (redescr.). 

Campylomyza valida Winnertz 1870b: 20. Germany; ST; ♂,♀; lost. 

Campylomyza melanoptera Kieffer 1904c: 405. France: vic. Bitche; ST; ♂,♀; lost. 

Campylomyza pulchricornis Kieffer 1904c: 404. France: vic. Bitche; T; ♀; lost. 

Xylopriona querceti Edwards 1938g: 205. UK: England, Beds., Whipsnade; HT; ♀; BMNH. 

Monardia longicornis Mamaev 1963a: 444. Russia: Yaroslavl Prov., Volkovo; HT; ♂; ZMMU. 

furcifera. Sweden, Latvia, Lithuania, Germany, Hungary, Ukraine, Russia (Europe, Far East). 

Monardia furcifera Mamaev 1963a: 444. Russia: Yaroslavl Prov., Volkovo; HT; ♂; ZMMU. 


Xylopriona indica Sharma & Rao 1979d: 299. India: Maharashtra, Aurangabad; HT; ♂; ELMU. 

monotheca. UK, Fennoscandia, Germany, Ukraine, Russia (Europe), Japan, Canada (British Columbia), USA 

(California). 

Monardia monotheca Edwards 1938h: 241. UK: England, Herts, Hitch Wood; HT; ♀; BMNH. 

*ornata. Russia (Far East). 

Xylopriona ornata Fedotova 2004i: 582. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

radiella. Sweden, Finland, Russia (Europe). 

Xylopriona radiella Mamaev 1993: 60. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

sejuncta. Russia (Western Siberia). 

Xylopriona sejuncta Mamaev 1993: 60. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

toxicodendri. USA (widespread), Mexico, Europe (widespread), Morocco, Russia (Europe, Western Siberia), 

Uzbekistan, China, Japan. Ref.: Jaschhof & Jaschhof 2009a: 212 (redescr.). 

Campylomyza toxicodendri Felt 1907a: 1. USA: New York, Albany; HT; ♀; NYSM. 

Campylomyza gilletti Felt 1908b: 314. USA: Pennsylvania, Highspire; HT; ♀; NYSM. 

Monardia toxicodendron Felt 1913c: 186, missp. of toxicodendri. 

Monardia alexanderi Felt 1913c: 187. USA: New York, Sport I., Sacandaga R.; HT; ♀; NYSM. 

Monardia modesta Felt 1913f: 142. USA: Connecticut, New Haven; HT; ♀; NYSM. 

Monardia illinoiensis Felt 1935b: 47. USA: Illinois, Urbana; ST; ♂,♀; NYSM & INHS. 

Monardia nigricans Edwards 1938h: 242. UK: England, Herts, Breachwood Green; HT; ♂; BMNH. 

Xylopriona plicans Mamaev 1993: 59. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

*truncata. Sweden. 

Monardia truncata Jaschhof 2009a: 213. Sweden: Uppland, Lövstabruk; HT; ♂; NHRS. 

unguifera. Finland, Ukraine. 

Xylopriona unguifera Berest & Mamaev 1997: 87. Ukraine: Zhitomir Prov., Korostenskiy, Ushomir; HT; ♂; UASK. 

vividula. Russia (Far East). 

Xylopriona vividula Mamaev 1993: 60. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

MONARDIA Incertae Sedis 

*fumea. New Zealand. 

Monardia fumea Jaschhof 2004a: 426. New Zealand: South I., Buller, Ahaura, Granville S.F.; HT; ♂; NZAC. 

*furcillata. New Zealand. 

Monardia furcillata Jaschhof 2004a: 427. New Zealand: South I., Buller, Ahaura, Granville S.F.; HT; ♂; NZAC. 

* † impellucida. Rovno amber (Ukraine). 

Monardia impellucida Fedotova & Perkovsky 2007a: 84. Ukrainian amber: Rovno; HT; ♂; UASK. 

*modica. New Zealand. 

Monardia modica Jaschhof 2004a: 428. New Zealand: South I., Westland, "South Westland"; HT; ♂; NZAC. 

MYCOPHILA

Mycophila Felt 1911c: 33. Type species, fungicola Felt (orig. des.). 

Holarctic, Oriental, Australasian, Oceanian; 7 spp.; Aprionini. Ref.: Jaschhof 1998b (rev.). 

echinoidea. China (Sichuan). 

Mycophila echinoidea Bu & Mo 1996: 113. China: Sichuan, Wenchuan, Wolong; HT; ♂; NKUM. 

fungicola. USA (California, Indiana, Ohio, Pennsylvania), UK, Netherlands, Germany, Russia (Far East), China, 

Hawaiian Is., Australasia. Ref.: Jaschhof & Jaschhof 2004a: 439. 

Mycophila fungicola Felt 1911c: 33. USA: California, San Rafael; LT (Pritchard 1947b: 65); ♂; USNM. 

Mycophila barnesi Edwards 1938i: 254. UK: England, Herts, Letchworth; HT; ♂; BMNH. 

indica. India (Kerala). 

Mycophila indica Nayar 1949: 81. India: Kerala, Trivandrum; ST; ♀; ZSCI. 

lampra. USA (Minnesota, Wisconsin). 

Mycophila lampra Pritchard 1947b: 65. USA: Minnesota, St. Paul, L. Johanna; HT; ♂; USNM. 

longispina. China (Shandong). 

Mycophila longispina Bu & Mo 1996: 112. China: Shandong, Yantai, Mt Kunyu; HT; ♂; SAUT. 

nikoleii. Germany, Russia (Europe). 

Mycophila nikoleii Möhn 1960a: 4. Germany: Göttingen, Gartetal; HT; ♂; SMNS. 

speyeri. UK, Netherlands, Denmark, Germany, Italy, China, Republic of Korea, Taiwan, Japan, USA (Michigan, 

Pennsylvania, Virginia, Maryland). Ref.: Lee & Kim 2003 (biol., illus.). 

Pezomyia speyeri Barnes 1926: 90. UK: England, Herts, Cheshunt; LT (Jaschhof 1998b: 424); ♂; BMNH. 

NEOCATOCHA 

Neocatocha Felt 1912l: 236. Type species, marilandica Felt (orig. des.). 

Konisomyia Felt 1914f: 118. Type species, fusca Felt (orig. des.) = marilandica Felt. 

Holarctic; 1 sp.; Catochini. Ref.: Jaschhof 1998b (redescr.), Jaschhof & Jaschhof 2009a (distr.). 

marilandica. Canada (Ontario, Newfoundland), USA (New York, Maryland), Sweden, Russia (Western Siberia), 

Japan (Kyushu). 

Neocatocha marilandica Felt 1912l: 236. USA: Maryland, Montgomery Co., Plummers I.; HT; ♀; NYSM. 

Konisomyia fusca Felt 1914f: 118. USA: New York, Albany; HT; ♀; NYSM. 

Konisomyia borealis Felt 1920a: 279. USA: New York, Keene Valley; HT; ♀; NYSM. 

Neocatocha asiatica Mamaev 1979: 186. Russia: Tuva, Ishtii-Khem; HT; ♀; HYNM. 

NEUROLYGA 

Neurolyga Rondani 1840: 24. Type species, fenestralis Rondani (subs. des. Rondani 1856: 199). 

Cordylomyia Felt 1911c: 35. Type species, coprophila Felt (orig. des.) = fenestralis Rondani. 

Prosaprionus Kieffer 1913f: 54. Type species, cellularis Kieffer (mon.), a nomen dubium, per M. Jaschhof (in litt.). 

Microcordylomyia Mamaev & Mohrig 1975: 126. Type species, degenerans Mamaev & Mohrig (orig. des.). 

Holarctic and Eocene amber; 29 spp.; Campylomyzini. Refs.: Jaschhof 1998b (rev.), Jaschhof & Jaschhof 2009a 

(redescr.). 

*acuminata. Fennoscandia, Russia (Western Siberia). 

Neurolyga acuminata Jaschhof 2009a: 126. Russia: Karelia, Kondopoga, Kivach Strict Nature Res.; HT; ♂; NHRS. 

*angulosa. Sweden, Kazakhstan. 

Neurolyga angulosa Jaschhof 2009a: 127. Sweden: Uppland, Knivsta, Rickebasta; HT; ♂; NHRS. 

barsovi. Ukraine. 

Cordylomyia barsovi Berest 1991a: 106. Ukraine: Dnepropetrovsk; HT; ♂; UASK. 

33

34 

bifida. UK, Sweden, Latvia, Germany, Ukraine, Russia (Europe, Far East). 

Cordylomyia bifida Edwards 1938g: 201. UK: England, Herts, Letchworth & Knebworth Wood; ST; ♂,♀; BMNH. 

Cordylomyia coronata Mamaev & Rozhnova 1983: 64. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

bilobata. Fennoscandia, Estonia, Latvia, Germany, Ukraine. 

Cordylomyia bilobata Mamaev & Rozhnova 1982: 20. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; ZMMU. Ref.: 


Cordylomyia subarctica Mamaev 1990b: 142. Russia: Yamal Peninsula, Khadyta; HT; ♂; ZMMU. 

* † cauta. Rovno amber (Ukraine). 

Cordylomyia cauta Fedotova & Perkovsky 2007a: 92. Ukrainian amber: Rovno: HT; ♀; UASK. Misplaced, per M. 


cellularis. France. 

cellularis Kieffer 1913f: 54 (Prosaprionus). France: Bitche; T; ♀; lost. A nomen dubium, per M. Jaschhof (in litt.). 

collaris. Russia (Europe). 

Cordylomyia collaris Mamaev 1963a: 453. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ZMMU. 

*constricta. Sweden. 

Neurolyga constricta Jaschhof 2009a: 129. Sweden: Södermanland, Tyresta For.; HT; ♂; NHRS. 

degenerans. Germany. 

Microcordylomyia degenerans Mamaev & Mohrig 1975: 126. Germany: Thüringen, Kyffhäuser, nr Bad 

Frankenhausen; HT; ♀; DEIC. 

denningi. USA (Minnesota). 

Cordylomyia denningi Pritchard 1947a: 37. USA: Minnesota, Anoka Co., Fridley; HT; ♂; UMSP. 

excavata. Fennoscandia, Germany, Austria, Hungary, Russia (Europe, Far East), Japan (Kyushu). 

Cordylomyia excavata Yukawa 1967b: 189. Japan: Kyushu, Kurume, Mt Kôra; HT; ♂; KUEC. Ref.: Jaschhof & 


Cordylomyia obsoleta Mamaev & Rozhnova 1982: 23. Russia: Moscow Prov., Il'icha For.; HT; ♂; ZMMU. 

Cordylomyia terricola Mamaev & Rozhnova 1983: 65. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

fenestralis. UK, Norway, Sweden, Latvia, Germany, Italy, Russia (Europe), Canada (British Columbia), USA 

(widespread). 

Neurolyga fenestralis Rondani 1840: 24. Italy; LT (Jaschhof 1998b: 194); ♀; MZUF. 

Campylomyza fuscinervis Winnertz 1870b: 19. Germany; ST; ♂,♀; lost. 

Campylomyza rudis Winnertz 1870b: 16. Germany; HT; ♀; lost. 

Campylomyza texana Felt 1908b: 316. USA: Texas, Plano; HT; ♂; NYSM. 

Cordylomyia coprophila Felt 1911c: 35. USA: District of Columbia; LT (Pritchard 1947a: 32); ♂; NYSM. 

Cordylomyia americana Felt 1913c: 199. USA: Colorado, Boulder; HT; ♀; NYSM. 

Cordylomyia coloradensis Felt 1913c: 199. USA: Colorado, Boulder; LT (Pritchard 1947a: 32); ♂; NYSM. 

Monardia foliata Felt 1916e: 195. USA: Colorado, Boulder; HT; ♂; NYSM. 

Cordylomyia fulva Felt 1926f: 266. Canada: British Columbia, Cranbrook; HT; ♀; NYSM. 

Cordylomyia praelonga Felt 1926f: 266. Canada: British Columbia, Cranbrook; LT (Pritchard 1947a: 34); ♂; NYSM. 

Cordylomyia scutellata Felt 1926f: 267. Canada: British Columbia, Cranbrook; HT; ♀; NYSM. 

Cordylomyia xylophila Edwards 1938g: 200. UK, England, Herts, Welwyn, Sherrard's Wood; LT (Jaschhof 1998b: 

194); ♂; BMNH. 

hastagera. Ukraine, Russia (Western Siberia, Far East). 

Cordylomyia hastagera Mamaev & Rozhnova 1982: 23. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; HYNM. 

hyperborea. Russia (Western Siberia, Far East). 

Cordylomyia hyperborea Mamaev 1990b: 141. Russia: Yamal Peninsula, Khadyta; HT; ♂; HYNM. 

*interrupta. Norway, Finland, Russia (Europe). 

Neurolyga interrupta Jaschhof 2009a: 132. Russia: Karelia, Kondopoga, Kivach Strict Nature Res.; HT; ♂; NHRS. 

*londalensis. Fennoscandia. 

Neurolyga lonsdalensis Jaschhof 2009a: 133. Norway: inner southern Nordland, Saltdal, Lønsdalen, Semska, 10 km 

S Lønsdal; HT; ♂; NHRS. 

longipes. USA (Oregon). 

Neurolyga longipes Jaschhof 1997b: 528. USA: Oregon, Willamette Natl For.; HT; ♂; USNM.

35 

* † magnifica. Lower Eocene - amber, northern France. 

Neurolyga magnifica Nel & Prokop 2006: 43. Amber fossil, northern France: Oise, Creil region, Chevrière; HT; ♀; 

NMHN. 

ovata. Sweden, Russia (Europe), Canada (Northwest Terr.). 

Neurolyga ovata Jaschhof 1996b: 349. Sweden: Torne Lappmark, Abisko; HT; ♂; DEIC. 

*paludosa. Fennoscandia. 

Neurolyga paludosa Jaschhof 2009a: 135. Sweden: Västerbotten, Lövänger, Kallviken HT; ♂; NHRS. 

pritchardi. USA (California). 

Neurolyga pritchardi Jaschhof 1997b: 530. USA: California, Inverness; HT; ♂; USNM. 

semicircula. China (Sichuan), Sweden. 

Cordylomyia semicircula Bu 1996: 480. China: Sichuan, Wenchuan (Wolong); HT; ♂; NKUM. 

spinifera. Japan (Honshu). 

Cordylomyia spinifera Yukawa 1971: 34. Japan: Honshu, Kyoto Pref., Serio; HT; ♂; KUIC. 

subbifida. Russia (Europe). 

Cordylomyia subbifida Mamaev 1963a: 453. Russia: Yaroslavl Prov., Rybinskoye Reservoir; HT; ♂; ZMMU. 

sylvestris. USA (Washington, California, North Dakota, Minnesota, North Carolina), Sweden, Finland, Latvia, 

Lithuania, Ukraine, Russia (Europe). 

Campylomyza sylvestris Felt 1907a: 1. USA: North Carolina, Davidson's R.; HT; ♀; NYSM. 

Cordylomyia commutata Mamaev & Rozhnova 1982: 21. Russia: Moscow Prov., Danki; HT; ♂; HYNM. 

truncata. USA (Pennsylvania), UK, Norway, Sweden, Estonia, Latvia, Austria, Uzbekistan. Jaschhof & Jaschhof 

2009a: 138 (redescr.). 

Campylomyza truncata Felt 1912f: 102. USA: Pennsylvania, Hazelton; HT; ♂; NYSM. 

Cordylomyia hammi Edwards 1938g: 201. UK: England, Herts (Letchworth), Bucks (Ivinghoe Common), Oxon 

(Univ. Museum); ST; ♂; BMNH. 

Cordylomyia montana Mamaev 1963c: 72. Uzbekistan: Samarkand, Takhtakaracha Pass; HT; ♂; HYNM. 

venusta. Russia (Europe, Far East). 

Cordylomyia venusta Mamaev & Rozhnova 1983: 66. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

verna. Finland, Latvia, Germany, Bulgaria, Russia (Europe, Far East), USA (Connecticut). 

Cordylomyia verna Mamaev 1963a: 453. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

*PROACOENONIA 

Proacoenonia Nel & Prokop 2006: 47. Type species, olgae Nel & Prokop (orig. des.). 

Eocene amber from northern France; 1 sp.; Acoenoniini. 

* † olgae. Lower Eocene - amber, northern France. 

Proacoenonia olgae Nel & Prokop 2006: 47. Amber fossil, northern France: Oise, Creil region, Chevrière; HT; ♂; 

NMHN. 

PEROMYIA 

Peromyia Kieffer 1894e: clxxv. Type species, leveillei Kieffer (mon.). 

Joannisia Kieffer 1894e: clxxv, preocc. Monterosato 1984. Type species, aurantiaca Kieffer (subs. des. Coquillett 

1910: 556). 

Joanisia Enderlein 1911c: 196, missp. of Joannisia. 

Joanissia Felt 1911c: 32, missp. of Joannisia. 

Johannisia Enderlein 1936: 62, missp. of Joannisia. 

Camptoza Enderlein 1936: 62. Type species, Joannisia kiefferiana Enderlein = fungicola Kieffer (subs. des. Pritchard 

1947: 74). 

Cosmopolitan and Baltic amber; 152 spp.; Peromyiini. Refs: Jaschhof 1998b (rev.), 2001b (Japanese spp.), Jaschhof

& Jaschhof 2004b (New Zealand spp.), Jaschhof & Jaschhof 2009a (rev.). 

*abdita. Finland. 

Peromyia abdita Jaschhof 2009a: 143. Finland: Ostrobottnia borealis, S part, Pudasjärvi, Syöte Natl Pk, Sotivaara; 


aberrans. Russia (Europe). 

Peromyia aberrans Mamaev 1963a: 438. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

abnormis. Sweden, Finland, Germany, Ukraine. 

Peromyia abnormis Mamaev & Berest 1990b: 18. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; ZMMU. 

acutula. Japan (Honshu). 

Peromyia acutula Jaschhof 2001b: 81. Japan: Honshu, Yamanashi Pref., Oyama Town, Mt Fuji; HT; ♂; KUEC. 

albicornis. Fennoscandia, Germany, ?Japan. 

Campylomyza albicornis Meigen 1830: 271. Germany; HT; ♂; MNHN. 

aleemkhani. Japan (widespread). 

Peromyia aleemkhani Jaschhof 2001b: 62. Japan: Ryukyu Is, Amami Oshima, Kawauchi Village, Kamiya For.; HT; 

♂; KUEC. 

almensis. Ukraine. 

Peromyia almensis Berest 1989: 85. Ukraine: Crimea, Alma R. valley; HT; ♂; UASK. 

ampla. Japan (Honshu). 

Peromyia ampla Jaschhof 2001b: 104. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa Research 

For.; HT; ♂; KUEC. 

anatina. Sweden, Finland, Germany, Ukraine, Japan (Hokkaido, Honshu). 

Peromyia anatina Mamaev & Berest 1990b: 20. Ukraine: Zakarpatskaya Prov., Kvasy, Mt Mincul'; HT; ♂; ZMMU. 

Ref.: Jaschhof & Jaschhof 2009a: 157 (redescr.). 

Peromyia subanatina Mamaev & Zaitzev 1997a: 3. Sweden: Bladåker, Bennebol; HT; ♂; ZMMU. 

angellifera. ennoscandia, Germany. 

Peromyia angellifera Jaschhof 1997a: 34. Germany: NSG Galenbecker See, 11 km SE Friedland; HT; ♂; DEIC. 

angulata. Japan (Honshu). 

Peromyia angulata Jaschhof 2001b: 142. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa Research 


anisotoma. Sweden, Finland, Russia (Far East). 

Peromyia anisotoma Mamaev 1994: 30. Russia: Kamchatka, Kozyrevsk; HT; ♂; ZMMU. 

anocellata. Japan (Honshu, Shikoku). 

Peromyia anocellata Jaschhof 2001b: 105. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa For.; 

HT; ♂; KUEC. 

apposita. Sweden, Finland, Russia (Europe), Latvia, Germany, Japan (widespread). 

Peromyia apposita Jaschhof 1997a: 49. Germany: Steffenshäger For., W Greifswald; HT; ♂; DEIC. 

aurantiaca. France, Germany. 

Joannisia aurantiaca Kieffer 1894e: clxxv. France: Bitche; LT (Jaschhof 1995: 89); ♂; NYSM. 

austrina. Japan (Kyushu). 

Peromyia austrina Jaschhof 2001b: 120. Japan: Kyushu, Miyazaki Pref., Aya Town, Aya Res. For.; HT; ♂; KUEC. 

autumnalis. Japan (Kyushu). 

Peromyia autumnalis Jaschhof 2001b: 63. Japan: Kyushu, Miyazaki Pref., Aya Town, Aya Res. For.; HT; ♂; KUEC. 

avia. Japan (Honshu, Kyushu). 

Peromyia avia Jaschhof 2001b: 136. Japan: Honshu, Tokyo City, Imperial Palace Gardens; HT; ♂; KUEC. 

ayaensis. Japan (Kyushu). 

Peromyia ayaensis Jaschhof 2001b: 143. Japan: Kyushu, Miyazaki Pref., Aya Town, Aya Res. For.; HT; ♂; KUEC. 

*bavarica. Germany. 

Peromyia bavarica Jaschhof 2009b: 148. Germany: Bavaria, Berchtesgaden Nat'l Pk, Schapbach-Alm, Schapbach; 


bengalensis. India (West Bengal). 

Peromyia bengalensis Kieffer 1905a: 158. India: West Bengal, vic. Kurseong; T; ♂; lost. 

36

37 

*bertviklundi. Sweden. 

Peromyia bertviklundi Jaschhof 2009a: 168. Sweden: Norrbotten, Niemisel, Blåkölen For. Res.; HT; ♂; NHRS. 

bicolor. Widespread northern Europe, Germany, Czech Republic. 

Joannisia bicolor Edwards 1938i: 261. UK: England, Herts, Letchworth; HT; ♀; BMNH. 

bidentata. Sweden, Latvia, Lithuania, Hungary, Ukraine, Japan (Hokkaido, Honshu). 

Peromyia bidentata Berest 1988b: 152. Ukraine: Zhitomir Prov., Korostensk, Pugachevka; HT; ♂; UASK. 

bihamata. Russia (Far East), Japan (Honshu). 

Peromyia bihamata Mamaev & Zaitzev 1997a: 2. Russia: Amur Prov., Zeja; HT; ♂; ZMMU. 

borealis. USA (Minnesota, New York), UK, Sweden, Latvia, Germany, Kirghizstan. Jaschhof & Jaschhof 2009a: 149 

(redescr.). 

Joannisia borealis Felt 1920a: 280. USA: New York, Keene Valley; HT; ♂; NYSM. Jaschhof & Jaschhof 2009a: 


Joannisia roralis Edwards 1938i: 260. UK: England, Herts, Harpenden; HT; ♂; BMNH. 

Peromyia palposa Mamaev & Zaitzev 1997a: 3. Kirghizstan: Sary-Chelek Res.; HT; ♂; ZMMU. 

boreojaponica. Japan (Hokkaido, Honshu). 

Peromyia boreojaponica Jaschhof 2001b: 65. Japan: Hk, Soya Sub-Pref., Rishiri I., Rishiri Town; HT; ♂; KUEC. 

boreophila. Finland, Russia (Europe), Japan (Hokkaido, Honshu). 

Peromyia boreophila Jaschhof 2001b: 107. Japan: Hk, Soya Sub-Pref., Rishiri I., Rishiri Town; HT; ♂; KUEC. 

capitata. Japan (Honshu). 

Peromyia capitata Jaschhof 2001b: 137. Japan: Honshu, Nagano Pref., Azumi Village, Norikurakogen; HT; ♂; 

KUEC. 

caricis. Widespread Palearctic, USA (California). 

Joannisia caricis Kieffer 1901c: 36. France: Bitche; NT (Jaschhof 1998b: 449; UK: England, Herts, Letchworth); ♂; 

BMNH. Ref: Jaschhof & Jaschhof 2009a: 150. 

Peromyia trifurcata Mamaev 1998a: 8. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

carinata. Japan (widespread, incl. Bonin Is), New Zealand (South I.). 

Peromyia carinata Jaschhof 2001b: 92. Japan: Honshu, Kukizaki Town, Matsunosato; HT; ♂; KUEC. 

cassa. Japan (Honshu). 

Peromyia cassa Jaschhof 2001b: 109. Japan: Honshu, Iwate Pref., Ashiro Town, Appi Highland, Mt Appidake; HT; 

♂; KUEC. 

*clandestina. New Zealand. 

Peromyia clandestina Jaschhof 2004b: 90. New Zealand: North I., Waikato, Waitomo Caves 12 km SW Otorohanga; 


centrosa. Japan (Kyushu, Ryukyu Is). 

Peromyia centrosa Jaschhof 2001b: 94. Japan: Kyushu, Miyazaki Pref., Aya Town, Aya Res. For.; HT; ♂; KUEC. 

*clancula. Russia (Europe). 

Peromyia clancula Jaschhof 2009a: 177. Russia: Karelia, Kondopoga, Kivach Strict Nature Res.; HT; ♂; NHRS. 

composita. Sweden, Latvia, Germany. 

Peromyia composita Jaschhof 1997a: 51. Germany: Mecklenburg-Vorpommern, Altwarp, 14 km E �ckermünde; 


concitata. Sweden, Finland, Germany, Russia (Europe), Ukraine. 

Peromyia concitata Mamaev & Berest 1994: 77. Ukraine: Zakarpatskaya Prov., Kvasy, Mt Mincul'; HT; ♂; ZMMU. 

Ref.: Jaschhof & Jaschhof 2009a: 162 (redescr.). 

Peromyia decurvata Mamaev 1998a: 8. Russia: Moscow Prov., Il'icha For.; HT; ♂; ZMMU. 

Peromyia miniuscula Mamaev 1998a: 8. Sweden, Dalarna; HT; ♂; ZMMU. 

cornuta. Widespread northern Europe, Germany, Ukraine, Russia (Europe, Far East), Japan (widespread). Ref.: 


Joannisia cornuta Edwards 1938i: 260. UK: England, Herts, Letchworth; HT; ♀; BMNH. 

Peromyia sphaerica Berest 1991a: 104. Ukraine: Cherkassy Prov., Uman; HT; ♂; UASK. 

*culta. New Zealand. 

Peromyia culta Jaschhof 2004b: 100. New Zealand: South I., Buller, Nelson Lakes Natl Pk, St. Arnaud; HT; ♂; 

NZAC.

38 

curta. Finland, Latvia, Lithuania, Germany, Ukraine, Russia (Europe), Japan (widespread). 

Peromyia curta Jaschhof 1997a: 53. Germany: Gützkow; HT; ♂; DEIC. Ref.: Jaschhof & Jaschhof 2009a: 157 

(redescr.). 

Peromyia baccha Mamaev 1998d: 3. Russia: Moscow Prov., Lugovaya; HT; ♂; ZMMU. 

*denotata. Sweden. 

Peromyia denotata Jaschhof 2009a: 160. Sweden: Uppland, Östhammar, Andersby Nature Res., SW Österbybruk; 


*derupta. New Zealand. 

Peromyia derupta Jaschhof 2004b: 96. New Zealand: South I., Buller, Nelson Lakes Natl Pk, L. Rotoiti; HT; ♂; 

NZAC. 

diadema. Widespread northern Europe, Germany, Russia (Europe). 

Peromyia diadema Mamaev 1963a: 438. Russia: Vologda Prov., Kadnikovskaya; HT; ♂; ZMMU. 

*didhami. New Zealand. 

Peromyia didhami Jaschhof 2004b: 108. New Zealand: Stewart I., Christmas Village Hut; HT; ♂; NZAC. 

directa. Ukraine. 

Peromyia directa Mamaev & Berest 1990b: 24. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

discreta. Sweden. 

Peromyia discreta Jaschhof 1997a: 47. Sweden: Pite Lappmark, Arjeplog, Laisvik, Vuordnats; HT; ♂; NHRS. Ref.: 


Peromyia longicostalis Mamaev & Zaitzev 1997a: 4. Sweden, Overgren, Biskops-Arnö; HT; ♂; ZMMU. 

*dissona. New Zealand. 

Peromyia dissona Jaschhof 2004b: 90. New Zealand: North I., Waikato, Waitomo Caves 12 km SW Otorohanga; 


*doci. New Zealand. 

Peromyia doci Jaschhof 2004b: 83. New Zealand: South I., Nelson Lakes Natl Pk, L. Rotoroa; HT; ♂; NZAC. 

edwardsi. UK, Sweden, Finland, Latvia, Germany, Ukraine. 

Peromyia edwardsi Berest 1994: 26. Ukraine: Zakarpatskaya Prov., Tyachev Region; HT; ♂; LASS. 

emarginata. Ukraine. 

Peromyia emarginata Berest 1988a: 74. Ukraine: Crimea, Simferopol Region, Krasnolesye; HT; ♂; UASK. 

extensa. Ukraine. 

Peromyia extensa Berest 1991a: 106. Ukraine: Kiyev Prov., Belaja Cerkov'; HT; ♂; UASK. 

fagiphila. Widespread northern Europe, Germany, Austria, Russia (Western Siberia, Far East), Japan (widespread). 

Peromyia fagiphila Jaschhof 1997a: 36. Germany: Elisenhain, 5 km E Greifswald; HT; ♂; DEIC. 

*fibyensis. Sweden. 

Peromyia fibyensis Jaschhof 2009a: 163. Sweden: Uppland, Uppsala, Fiby Nature Res.; HT; ♂; NHRS. 

fujiensis. Japan (Honshu). 

Peromyia fujiensis Jaschhof 2001b: 133. Japan: Honshu, Yamanashi Pref., Oyama Town, Mt Fuji; HT; ♂; KUEC. 

fungicola. Widespread Palearctic, Canada (British Columbia), USA (California). 

Joannisia fungicola Kieffer 1898f: 48, nomen nudum. 

Joannisia fungicola Kieffer 1901c: 37. France: vic. Bitche; NT (Jaschhof 1998b: 474; UK: England, Herts, Welwyn, 

Sherrard's Wood); ♂; BMNH. 

Joannisia kiefferiana Enderlein 1911b: 573. Poland: Wollin, Warnow-See; HT; ♀; ZMPA. 

Peromyia brevispina Yukawa 1967b: 188. Japan: Kyushu, Fukuoka Pref., Mt Hiko; HT; ♂; KUEC. 

Peromyia subapicalis Mamaev & Berest 1990b: 21. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; HYNM. 

*galapagensis. Galapagos Is. 

Peromyia galapagensis Jaschhof 2004c: 1043. Galapagos Is.: Santa Cruz, Los Gemelos, 31 km N Santa Rosa; HT; ♂; 

CNCI. 

gemella. Japan (widespread). 

Peromyia gemella Jaschhof 2001b: 84. Japan: Ryukyu Is, Amami Oshima, Kawauchi Village, Kamiya For.; HT; ♂; 

KUEC. 

gotohi. Japan (Honshu, Kyushu, Ryukyu Is). 

Peromyia gotohi Jaschhof 2001b: 79. Japan: Honshu, Iwate Pref., Tamayama Village, Koma; HT; ♂; KUEC.

39 

gryphiswaldensis. Germany, Japan (Honshu). 

Peromyia gryphiswaldensis Jaschhof 1997a: 43. Germany: Greifswald; HT; ♂; DEIC. 

horridula. Denmark, Germany, Japan (Honshu). 

Peromyia horridula Jaschhof 1997a: 41. Germany: Greifswald; HT; ♂; DEIC. 

ibarakiensis. Japan (Honshu). 

Peromyia ibarakiensis Jaschhof 2001b: 111. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa 

Research For.; HT; ♂; KUEC. 

imperatoria. Japan (Honshu, Ryukyu Is). 

Peromyia imperatoria Jaschhof 2001b: 51. Japan: Honshu, Tokyo City, Imperial Palace Gardens; HT; ♂; KUEC. 

indica. India (Madhya Pradesh). 

Peromyia indica Grover 1970c: 158. India: Madhya Pradesh, Indore; HT; ♂; PGCA. 

*insueta. New Zealand. 

Peromyia insueta Jaschhof 2004b: 106. New Zealand: South I., Buller, Ahaura, Granville S.F.; HT; ♂; NZAC. 

*intecta. New Zealand. 

Peromyia intecta Jaschhof 2004b: 113. New Zealand: South I., Buller, Nelson Lakes Natl Pk, L. Rotoroa; HT; ♂; 

NZAC. 

intermedia. Sweden, Finland, Russia (Europe), France, Germany, Latvia, Japan (Honshu, Kyushu). 

Joannisia intermedia Kieffer 1895i: 62. France: Bitche; NT (Jaschhof 1998b: 458; Germany: Thüringen, Waldeck SE 

Bürgel); ♂; BMNH. 

*intonsa. New Zealand. 

Peromyia intonsa Jaschhof 2004b: 91. New Zealand: South I., Buller, Ahaura, Granville S.F.; HT; ♂; NZAC. 

*katieae. New Zealand. 

Peromyia katieae Jaschhof 2004b: 98. New Zealand: South I., Westland, Franz Josef Village, Canavans Knob; HT; 

♂; NZAC. 

*latebrosa. New Zealand. 

Peromyia latebrosa Jaschhof 2004b: 103. New Zealand: Stewart I., Smoky Beach; HT; ♂; NZAC. 

leveillei. France. 

Peromyia leveillei Kieffer 1894e: clxxv. France: Bitche; LT (Jaschhof 1998b: 501); ♀; ZMHB. 

lobata. Japan (Honshu, Kyushu). 

Peromyia lobata Yukawa 1971: 29. Japan: Kyushu, Fukuoka Pref., Mt Tachibana; HT; ♂; KUIC. 

lobuscorum. Japan (Hokkaido, Honshu). 

Peromyia lobuscorum Jaschhof 2001b: 128. Japan: Honshu, Aomori Pref., Towadako Town, Tsuta hot spring; HT; 

♂; KUEC. 

maetoi. Japan (Hokkaido, Honshu). 

Peromyia maetoi Jaschhof 2001b: 58. Japan: Honshu, Ibaraki Pref., Abukuma Highlands, Kitaibaraki City, 

Sadanami, Ogawa For.; HT; ♂; KUEC. 

mediterranea. Italy. 

Peromyia mediterranea Mamaev 1998a: 8. Italy: Siena; HT; ♂; ZMMU. 

*memoranda. New Zealand. 

Peromyia memoranda Jaschhof 2004b: 102. New Zealand: South I., Nelson Lakes Natl Pk, L. Rotoroa; HT; ♂; 

NZAC. 

*menzeli. Sweden, Germany. 

Peromyia menzeli Jaschhof 2009a: 145. Sweden: Västerbotten, 5 km SE Lövänger, Kallviken; HT; ♂; NHRS. 

mica. Italy. 

Peromyia mica Mamaev & Zaitzev 1998b: 127. Italy: Pontignano; HT; ♂; ZMMU. 

minutissima. Germany, Latvia, Ukraine, Russia (Europe). 

Peromyia minutissima Mamaev 1963a: 438. Russia: Tula Prov., Tula For.; HT; ♂; ZMMU. 

mitrata. Sweden, Finland, Latvia, Lithuania, Germany. 

Peromyia mitrata Jaschhof 1997a: 55. Germany: Elisenhain, 5 km E Greifswald; HT; ♂; DEIC. 

miyazakiensis. Japan (Honshu, Kyushu). 

Peromyia miyazakiensis Jaschhof 2001b: 97. Japan: Kyushu, Miyazaki Pref., Hinokage Town; HT; ♂; KUEC. 

modesta. USA (New York), widespread Palearctic.

40 

Campylomyza modesta Felt 1907a: 3. USA: New York, Lake Clear; HT; ♂; NYSM. 

† 

monilifera. Upper Eocene - Baltic amber. 

Joannisia monilifera Meunier 1904: 41. Baltic amber; ST; ♂,♀; lost. 

monilis. UK, Fennoscandia, Latvia, Germany, Hungary, Ukraine, Russia (Europe), USA (Minnesota). 

Peromyia monilis Mamaev 1965a: 266. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

Peromyia alni Kleesattel 1979: 156. Germany: Baden-Württemberg, Täferrot, nr Schwäbisch-Gmünd; LT (Jaschhof 

1998b: 453); ♂; SMNS. 

montivaga. Japan (Honshu). 

Peromyia montivaga Jaschhof 2001b: 113. Japan: Honshu, Akita Pref., Fujisato Town, Shirakami Mts; HT; ♂; 

KUEC. 

*mountalbertiensis. New Zealand. 

Peromyia mountalbertiensis Jaschhof 2004b: 84. New Zealand: North I., Northland, Waipoua For., along S.H. 12; 


*multifurcata. New Zealand. 

Peromyia multifurcata Jaschhof 2004b: 119. New Zealand: South I., Buller, Nelson Lakes Natl Pk, St. Arnaud; HT; 

♂; NZAC. 

muscorum. Widespread Palearctic, Japan (Bonin Is), New Zealand. 

Joannisia muscorum Kieffer 1895i: 62. France: vic. Bitche; NT (Jaschhof 1998b: 476; UK: England, Herts, 

Letchworth); ♂; BMNH. 

Peromyia penisulae Mamaev 1998a: 9. Russia: Kamchatka, Kozyrevsk; HT; ♂; ZMMU. 

Peromyia peninsulae Mamaev & Zaitzev 1999: 254, missp. of penisulae. 

nemorum. UK, Sweden, Estonia, Latvia, Germany, Japan (Honshu). 

Joannisia nemorum Edwards 1938i: 263. UK: England, Herts, Welwyn, Sherrard's Wood; LT (Jaschhof 1998b: 464); 

♂; BMNH. 

neomexicana. USA (New Mexico), Germany, Ukraine, Russia (Europe, Far East), Japan (widespread). 

Joannisia neomexicana Felt 1913c: 160. USA: New Mexico, Pecos; HT; ♂; NYSM. 

Peromyia carpathica Mamaev & Berest 1990b: 20. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; HYNM. 

*novaezealandiae. New Zealand. 

Peromyia novaezealandiae Jaschhof 2004b: 87. New Zealand: North I., Northland, Parataiko Ra., 45 km NW 

Dargaville; HT; ♂; NZAC. 

obesa. Japan (Hokkaido). 

Peromyia obesa Jaschhof 2001b: 121. Japan: Hokkaido, Iburi Sub-Pref., Tomakomai City, Hokkaido Univ. 

Experimental For.; HT; ♂; KUEC. 

*obunca. New Zealand. 

Peromyia obunca Jaschhof 2004b: 115. New Zealand: South I., Buller, Nelson Lakes Natl Pk, St. Arnaud; HT; ♂; 

NZAC. 

ogawaensis. Japan (Honshu, Kyushu). 

Peromyia ogawaensis Jaschhof 2001b: 115. Japan: Honshu, Ibaraki Pref., Abukuma Highlands, Kitaibaraki City, 


okochii. Japan (Hokkaido, Honshu). 

Peromyia okochii Jaschhof 2001b: 68. Japan: Honshu, Ibaraki Pref., Abukuma Highlands, Kitaibaraki City, 



Peromyia orientalis (also as indica) Grover 1964a: 123. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

ornata. Japan (widespread). 

Peromyia ornata Jaschhof 2001b: 122. Japan: Hokkaido, Iburi Sub-Pref., Tomakomai City, Hokkaido Univ. 

Experimental For.; HT; ♂; KUEC. 

ovalis. UK, Fennoscandia, Latvia, Germany, Russia (Europe, Far East), USA (Minnesota). 

Joannisia ovalis Edwards 1938i: 258. UK: England, Herts, Letchworth; HT; ♂; BMNH. 

paliformia. Russia (Far East). 

Peromyia paliformia Berest 1994: 25. Russia: Primorskiï Terr., Chasanskiï R., Zanadvorovka; HT; ♂; UASK. 

palustris. Widespread western Palearctic, Russia (Far East), Japan, USA (New York), New Zealand (widespread).

41 

Joannisia palustris Kieffer 1895i: 62. France: Bitche; NT (Jaschhof 1998b: 478; UK: England, London); ♂; BMNH. 

Peromyia prominens Yukawa 1967b: 186. Japan: Kyushu, Fukuoka-shi; HT; ♂; KUIC. 

penicillata. Germany. 

Peromyia penicillata Jaschhof 1997a: 58. Germany: Karbow, 15 km SE Greifswald; HT; ♂; DEIC. 

*perardua. New Zealand. 

Peromyia perardua Jaschhof 2004b: 89. New Zealand: North I., Northland, Dargaville; HT; ♂; NZAC. 

perpusilla. Denmark, Fennoscandia, Lithuania, Germany, Russia (Near East), Japan (Honshu). 

Campylomyza perpusilla Winnertz 1870b: 15. Germany; NT (Jaschhof 1998b: 487; Germany: Gehren, 9 km N 

Strasburg); ♂; BMNH. 

*pertrita. New Zealand. 

Peromyia pertrita Jaschhof 2004b: 93. New Zealand: North I., Gisborne, Kaimai Ra., Whakamarama 10 km SW 

Tauranga; HT; ♂; NZAC. 

photophila. USA (widespread eastern), UK, Fennoscandia, Latvia, Germany, Italy, Russia (Europe, Far East), Japan 

(widespread). 

Campylomyza photophila Felt 1907a: 3. USA: New York, Albany; LT (Pritchard 1947b: 77); ♂; NYSM. 

Campylomyza carolinae Felt 1907a: 4. USA: North Carolina, Davidson's R.; LT (Pritchard 1947b: 77); ♂; NYSM. 

Joannisia flavoscuta Felt 1908b: 313. USA: New York, Nassau; HT; ♂; NYSM. 

Joannisia flavopedalis Felt 1908b: 313. USA: New York, Newport; HT; ♀; NYSM. 

Joannisia pennsylvanica Felt 1911m: 476. USA: Pennsylvania, Reading; LT (Pritchard 1947b: 77); ♂; NYSM. 

Joannisia nodosa Edwards 1938i: 263. UK: England, Yorkshire, Mulgrave Woods, nr Whitby; HT; ♂; BMNH. 

pilosa. Japan (Honshu, Kyushu). 

Peromyia pilosa Jaschhof 2001b: 139. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa For.; HT; ♂; 

KUEC. 

*plena. New Zealand. 

Peromyia plena Jaschhof 2004b: 97. New Zealand: North I., Northland, Waipoua For., 45 km NW Dargaville; HT; 

♂; NZAC. 

podolica. Ukraine. 

Peromyia podolica Berest 1991b: 557. Ukraine: Vinnitsa Prov., Kalinovka, Uladovo; HT; ♂; UASK. 

*praeclara. New Zealand. 

Peromyia praeclara Jaschhof 2004b: 118. New Zealand: North I., Waikato, Waitomo Caves 12 km SW Otorohanga; 


pumila. Fennoscandia, Japan (widespread). 

Peromyia pumila Jaschhof 2001b: 70. Japan: Ryukyu Is, Okinawa Pref., Iriomote I., Komi Village, Maera R. valley; 


puncta. Japan (Honshu, Ryukyu Is). 

Peromyia puncta Jaschhof 2001b: 54. Japan: Honshu, Akita Pref., Fujisato Town, Shirakami Mts, Mt Futatsumori; 


ramosa. Widespread northern Europe, Germany, Poland, Ukraine, Russia (Europe, Siberia, Far East), Japan 

(widespread). 

Joannisia ramosa Edwards 1938i: 258. UK: England, Herts, Letchworth; HT; ♂; BMNH. 

Joannisia trimera Edwards 1938i: 258. UK: England, Herts, Letchworth; HT; ♂; BMNH. 

*ramosoides. Finland. 

Peromyia ramosoides Jaschhof 2009a: 155. Finland: Satakunta, Ikaalinen, Seitseminen Natl Pk, Kuru, N Pitkajärvi; 


*rara. New Zealand. 

Peromyia rara Jaschhof 2004b: 100. New Zealand: South I., Westland, 5 km SW Arawhata; HT; ♂; NZAC. 

revelata. Ukraine, Japan (widespread). 

Peromyia revelata Mamaev & Berest 1990b: 23. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

rhombica. Japan (widespread). 

Peromyia rhombica Jaschhof 2001b: 72. Japan: Honshu, Shizuoka Pref., Fujinomiya City, Mt Fuji; HT; ♂; KUEC. 

*rotoitiensis. New Zealand. 

Peromyia rotoitiensis Jaschhof 2004b: 104. New Zealand: South I., Buller, Nelson Lakes Natl Pk, L. Rotoiti; HT; ♂;

42 

NZAC. 

ryukyuensis. Japan (Ryukyu Is). 

Peromyia ryukyuensis Jaschhof 2001b: 124. Japan: Ryukyu Is, Okinawa Pref., Yambaru Peninsula, Yona Village; 


sacculiformia. Ukraine. 

Peromyia sacculiformia Mamaev & Berest 1990b: 21. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; ZMMU. 

sanguinea. Fennoscandia, France, Germany, Japan (Honshu, Kyushu). 

Joannisia sanguinea Kieffer 1894e: clxxv. France: Bitche; LT (Jaschhof 1998b: 498); ♀; ZMHB. 

*scirrhosa. Sweden, Finland, Russia (Europe). 

Peromyia scirrhosa Jaschhof 2009a: 170. Russia: Karelia, Kondopoga, Kivach Strict Nature Res.; HT; ♂; NHRS. 

scutellata. Fennoscandia, Latvia, Germany, Austria, Russia (Europe, Western Siberia), Japan (Honshu). Jaschhof & 


Peromyia scutellata Mamaev 1990b: 142. Russia: Yamal Peninsula, Khadyta; HT; ♂; ZMMU. 

Peromyia viklundi Jaschhof 1997a: 38. Sweden: Pite Lappmark, Arjeplog, Laisvik, Vuordnats; HT; ♂; NHRS. 

Peromyia bipuncta Jaschhof 2001b: 50. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa Research 


semota. Japan (Hokkaido, Honshu). 

Peromyia semota Jaschhof 2001b: 74. Japan: Honshu, Aomori Pref., Towadako Town, Tsuta hot spring; HT; ♂; 

KUEC. 

*semotoides. Sweden, Finland. 

Peromyia semotoides Jaschhof 2009a: 152. Finland: Karelia ladogensis, Parikkala, Lake Siikalahti, W Kaukola; HT; 

♂; NHRS. 

*sera. New Zealand. 

Peromyia sera Jaschhof 2004b: 85. New Zealand: South I., Nelson Lakes Natl Pk, L. Rotoroa; HT; ♂; NZAC. 

*serrata. New Zealand. 

Peromyia serrata Jaschhof 2004b: 81. New Zealand: South I., Buller, Nelson Lakes Natl Pk, L. Rotoroa; HT; ♂; 

NZAC. 

*setosa. New Zealand. 

Peromyia setosa Jaschhof 2004b: 95. New Zealand: North I., Wanganui, Palmerston North, "in reserve"; HT; ♂; 

NZAC. 

seychellensis. Seychelles Is. 

Joannisia seychellensis Kieffer 1911: 329. Seychelles Is: Mahé, Cascade Estate; ST; ♂,♀; BMNH. 

simpla. Japan (Honshu). 

Peromyia simpla Jaschhof 2001b: 86. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa For.; HT; ♂; 

KUEC. 

*sinuosa. New Zealand. 

Peromyia sinuosa Jaschhof 2004b: 81. New Zealand: South I., North Canterbury, Arthur's Pass Natl Pk, Bealey Spur, 

Waimakariri Track; HT; ♂; NZAC. 

sphenoides. Japan (Honshu). 

Peromyia sphenoides Jaschhof 2001b: 59. Japan: Honshu, Ibaraki Pref., Abukuma Highlands, Kitaibaraki City, 


*spinigera. New Zealand. 

Peromyia spinigera Jaschhof 2004b: 79. New Zealand: North I., Northland, Dargaville; HT; ♂; NZAC. 

spinosa. Sweden, Japan (widespread). 

Peromyia spinosa Jaschhof 2001b: 75. Japan: Honshu, Ibaraki Pref., Abukuma Highlands, Kitaibaraki City, 


*squamigera. New Zealand. 

Peromyia squamigera Jaschhof 2004b: 112. New Zealand: South I., Waikato, Westland, 5 km SW Arawhata; HT; ♂; 

NZAC. 

*subbicolor. Sweden, Finland, Russia (Europe). 

Peromyia subbicolor Jaschhof 2009a: 147. Finland: Lapponia kemensis, W part, Kittilä, Pallas-Yllästunturi Natl Pk, 

Lake Pyhäjärvi; HT; ♂; NHRS.

43 

subborealis. Fennoscandia, Latvia, Germany, Japan (Hokkaido). 

Peromyia subborealis Jaschhof 1997a: 55. Germany: Steffenshäger For., W Greifswald; HT; ♂; DEIC. 

subcurta. Japan (Honshu, Kyushu). 

Peromyia subcurta Jaschhof 2001b: 86. Japan: Honshu, Kyoto Pref., Kyoto City, Kuta Res. For.; HT; ♂; KUEC. 

suberis. Spain, Germany. 

Peromyia suberis Jaschhof 1997a: 45. Spain: Andalusia, Jimena de la Frontera; HT; ♂; DEIC. 

syltefjordensis. Fennoscandia. 

Peromyia syltefjordensis Jaschhof 1996b: 350. Norway: Finnmark, Varanger-Halbinsel, Ytre Syltefjord, 35 km SE 

Batsfjord; HT; ♂; DEIC. 

*tecta. New Zealand. 

Peromyia tecta Jaschhof 2004b: 110. New Zealand: South I., Otago Lakes, Fiordland Natl Pk, Eglinton R. valley, 

Deer Flat; HT; ♂; NZAC. 

tenella. Japan (Honshu). 

Peromyia tenella Jaschhof 2001b: 89. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa For.; HT; ♂; 

KUEC. 

trifida. Japan (Honshu). 

Peromyia trifida Jaschhof 2001b: 102. Japan: Honshu, Iwate Pref., Tamayama Village, Koma; HT; ♂; KUEC. 

tripuncta. Japan (Honshu, Ryukyu Is). 

Peromyia tripuncta Jaschhof 2001b: 56. Japan: Honshu, Tokyo City, Imperial Palace Garden; HT; ♂; KUEC. 

truncata. Spain, Sweden, Finland, Germany, Russia (Europe, Far East), Japan (widespread). 

Peromyia truncata Yukawa 1967c: 65. Japan: Kyushu, Fukuoka Pref., Mt Hiko; HT; ♂; KUEC. Ref.: Jaschhof & 


Peromyia membranacea Fedotova 2004i: 576. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

Peromyia squarrosa Fedotova 2004i: 579. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

tschirnhausi. Sweden. 

Peromyia tschirnhausi Jaschhof 1996b: 352. Sweden: Torne Lappmark, Abisko; HT; ♂; DEIC. 

tsukubasanensis. Japan (Honshu). 

Peromyia tsukubasanensis Jaschhof 2001b: 116. Japan: Honshu, Ibaraki Pref., Mt Tsukuba; HT; ♂; KUEC. 

*tumida. New Zealand. 

Peromyia tumida Jaschhof 2004b: 94. New Zealand: South I., Nelson Lakes Natl Pk, L. Rotoroa; HT; ♂; NZAC. 

tundrae. Sweden, Finland, Germany, Russia (Europe), Japan (Hokkaido, Honshu). 

Peromyia tundrae Jaschhof 1996b: 353. Sweden: Torne Lappmark, Abisko; HT; ♂; DEIC. 

upupoides. Sweden, Finland, Russia (Europe), Latvia, Germany. 

Peromyia upupoides Jaschhof 1997a: 61. Germany: Gützkow; HT; ♂; DEIC. 

valens. Japan (Hokkaido, Honshu). 

Peromyia valens Jaschhof 2001b: 145. Japan: Honshu, Aomori Pref., Towadako Town, Tsuta hot spring; HT; ♂; 

KUEC. 

vernalis. Japan (Honshu). 

Peromyia vernalis Jaschhof 2001b: 118. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa Research 


yezoensis. Japan (Hokkaido, Honshu). 

Peromyia yezoensis Jaschhof 2001b: 140. Japan: Hokkaido, Kamikawa Sub-Pref., Hagashikawa Town, Ishikari Mts, 

Tenninkyo hot spring; HT; ♂; KUEC. 

* † zherikhini. Rovno amber (Ukraine). 

Peromyia zherikhini Fedotova 2004a: 48. Ukrainian amber: Rovno; HT; ♂; UASK. Ref.: Jaschhof 2007a: 105. 

Misplaced, per M. Jaschhof (in litt.). 

POLYARDIS 

Polyardis Pritchard 1947b: 47. Type species, Campylomyza carpini Felt (orig. des.) = silvalis Rondani.

44 

Campyloneura Lengersdorf 1939: 1, preocc. Fieber 1860. Type species, recondita Lengersdorf (orig. des. as n. g., n. 

sp.). 

Holarctic and New Zealand; 10 spp.; Micromyini. Ref.: Jaschhof 1998b, Jaschhof & Jaschhof 2009a (rev.). 

adela. USA (Minnesota), widespread Europe, Russia (Western Siberia). 

Polyardis adela Pritchard 1947b: 49. USA: Minnesota, Afton; HT; ♂; UMSP. Ref.: Jaschhof & Jaschhof 2009a: 216 

(redescr.). 

Monardia edwardsi Kleesattel 1979: 312. UK: England, Oxford, Ditchley; HT; ♂; BMNH. 

Polyardis anomala Mamaev 1993: 53. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

Polyardis delicata Mamaev 1993: 54. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; ZMMU. 

Polyardis explicata Mamaev 1993: 54. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ZMMU. 

Polyardis pontignanorum Mamaev & Zaitzev 1998b: 127. Italy: Pontignano; HT; ♂; ZMMU. 

bispinosa. Widespread Europe, Russia (Western Siberia, Far East). 

Monardia bispinosa Mamaev 1963a: 443. Russia: Moscow Prov., Abramtsevo; HT; ♂; HYNM. 

crebra. USA (Minnesota). 

Xylopriona crebra Pritchard 1947b: 45. USA: Minnesota, Afton; HT; ♂; UMSP. 

*illustris. New Zealand. 

Polyardis illustris Jaschhof 2004a: 431. New Zealand: South I., Buller, Nelson Lakes Natl Pk, St. Arnaud; HT; ♂; 

NZAC. 

micromyoides. Germany. 

Polyardis micromyoides Jaschhof 1998a: 272. Germany: Greifswald; HT; ♂; DEIC. 

occulta. Canada (Ontario). 

Polyardis occulta Jaschhof 1997b: 534. Canada: Ontario, 18 km E Gananogue; HT; ♂; USNM. 

recondita. Romania. 

Campyloneura recondita Lengersdorf 1939: 1. Romania: Carpathia, Muntii Faget, 90 km NW Cluj; HT; ♂; ZFMK. 

silvalis. Widespread Holarctic. 

Neurolyga silvalis Rondani 1840: 24. Italy: vic. Parma; LT (Jaschhof 1998b: 323); ♂; MZUF. Ref.: Jaschhof & 


Campylomyza carpini Felt 1907a: 4. USA: New York, Albany; HT; ♂; NYSM. 

Campylomyza kasloensis Felt 1908b: 314. Canada: British Columbia, Kaslo; HT; ♀; NYSM. 

Campylomyza versicolor Felt 1908b: 314. USA: New York, Albany; HT; ♀; NYSM. 

Polyardis aporia Pritchard 1947b: 51. USA: Minnesota, Bemidji; HT; ♂; UMSP. 

Monardia elongata Mamaev 1963c: 73. Uzbekistan: Samarkand, Takhtakaracha Pass; HT; ♂; HYNM. 

Aprionus micropterus Mamaev & Mohrig 1975: 128. Denmark: Hestehaven; HT; ♀; DEIC. 

Polyardis carbonaria Mamaev 1993: 54. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

Polyardis brevis Mamaev 1993: 54. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

Polyardis detriticola Mamaev 1993: 54. Russia: Primorskiï Terr., Chapigou; HT; ♂; ZMMU. 

Polyardis fallax Mamaev 1993: 55. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

Polyardis glabra Mamaev 1993: 55. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

*triangula. New Zealand. 

Polyardis triangula Jaschhof 2004a: 432. New Zealand: South I., Mid Canterbury, Banks Peninsula, Prices Valley; 


vitinea. USA (New York). 

Campylomyza vitinea Felt 1907a: 2. USA: New York, Albany; HT; ♂; NYSM. 

*PSEUDOMONARDIA 

Pseudomonardia Jaschhof 2003b: 115. Type species, hutchesoni Jaschhof (orig. des.). 

Australasian; 11 spp.; Pteridomyiini.

45 

*australis. New Zealand. 

Pseudomonardia australis Jaschhof 2003b: 121. New Zealand: South I., Mid Canterbury, Cass; HT; ♂; NZAC. 

*communis. New Zealand. 

Pseudomonardia communis Jaschhof 2003b: 125. New Zealand: North I., Waikato, Waitomo Caves 12 mi SW 

Otorohanga; HT; ♂; NZAC. 

*elongata. New Zealand. 

Pseudomonardia elongata Jaschhof 2003b: 129. New Zealand: South I., Buller, Ahaura, Granville St. For.; HT; ♂; 

NZAC. 

*glacialis. New Zealand. 

Pseudomonardia glacialis Jaschhof 2003b: 130. New Zealand: South I., Westland, Westland Natl Pk, Fox Glacier 

Village; HT; ♂; NZAC. 

*hutchesoni. New Zealand. 

Pseudomonardia hutchesoni Jaschhof 2003b: 116. New Zealand: South I., Buller, Ahaura, Granville St. For.; HT; ♂; 

NZAC. 

*invisitata. New Zealand. 

Pseudomonardia invisitata Jaschhof 2003b: 119. New Zealand: North I., Taupo, Tongariro Natl Pk; HT; ♂; NZAC. 

*neurolygoides. New Zealand. 

Pseudomonardia neurolygoides Jaschhof 2003b: 118. New Zealand: South I., Nelson Lakes Natl Pk, L. Rotoroa; HT; 

♂; NZAC. 

*pallida. New Zealand. 

Pseudomonardia pallida Jaschhof 2003b: 124. New Zealand: South I., Buller, Nelson Lakes Natl Pk, St. Arnaud; 


*parva. New Zealand. 

Pseudomonardia parva Jaschhof 2003b: 128. New Zealand: South I., Nelson Lakes Natl Pk, L. Rotoroa; HT; ♂; 

NZAC. 

*parvolobata. New Zealand. 

Pseudomonardia parvolobata Jaschhof 2003b: 122. New Zealand: South I., Westland, Westland Natl Pk, Canavan's 

Knob; HT; ♂; NZAC. 

*vicina. New Zealand. 

Pseudomonardia vicina Jaschhof 2003b: 123. New Zealand: North I., Taupo, Pureora For., Waipapa Res.; HT; ♂; 

NZAC. 

PSEUDOPEROMYIA 

Pseudoperomyia Jaschhof & Hippa 1999a: 148. Type species, longicornis Jaschhof & Hippa (orig. des.). 

Oriental, Eastern Palearctic; 29 spp.; Micromyini. 

acutistyla. Malaysia. 

Pseudoperomyia acutistyla Jaschhof & Hippa 1999a: 150. Malaysia: Pahang, Cameron Highlands, Gunung Jasar; 


bidentata. Malaysia. 

Pseudoperomyia bidentata Jaschhof & Hippa 1999a: 152. Malaysia: Selangor, Genting Highlands, Gunung Ulu Kali; 


bispinata. Malaysia. 

Pseudoperomyia bispinata Jaschhof & Hippa 1999b: 347. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 

Studies Centre; HT; ♂; NHRS. 

bryomyoides. Malaysia, Japan (Kyushu). 

Pseudoperomyia bryomyoides Jaschhof & Hippa 1999b: 342. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya 

Field Studies Centre; HT; ♂; NHRS. 

composita. Japan (Honshu). 

Pseudoperomyia composita Jaschhof 2000: 23. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa

46 


fagiphila. Japan (Honshu). 

Pseudoperomyia fagiphila Jaschhof 2000: 25. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa 


furcillata. Japan (Ryukyu Is). 

Pseudoperomyia furcillata Jaschhof 2000: 27. Japan: Ryukyu Is, Okinawa, Yambaru Peninsula, Yona; HT; ♂; 

KUEC. 

gemina. Malaysia. 

Pseudoperomyia gemina Jaschhof & Hippa 1999b: 344. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


hondoensis. Japan (Honshu). 

Pseudoperomyia hondoensis Jaschhof 2000: 28. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa 


humilis. Malaysia, China (Yunnan), Japan (Ryukyu Is). 

Pseudoperomyia humilis Jaschhof & Hippa 1999a: 154. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


intermedia. Malaysia. 

Pseudoperomyia intermedia Jaschhof & Hippa 1999a: 157. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


japonica. Japan (Honshu). 

Pseudoperomyia japonica Jaschhof 2000: 30. Japan: Honshu, Ibaraki Pref., Kitaibaraki City, Sadanami, Ogawa 


labellata. Malaysia. 

Pseudoperomyia labellata Jaschhof & Hippa 1999b: 353. Malaysia: Selangor, Genting Highlands, Gunung Ulu Kali; 


longicornis. Malaysia. 

Pseudoperomyia longicornis Jaschhof & Hippa 1999a: 158. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


longidentata. Malaysia. 

Pseudoperomyia longidentata Jaschhof & Hippa 1999b: 355. Malaysia: Pahang, Bukit Fraser; HT; ♂; NHRS. 

macrostyla. Malaysia. 

Pseudoperomyia macrostyla Jaschhof & Hippa 1999a: 160. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


obtecta. Malaysia. 

Pseudoperomyia obtecta Jaschhof & Hippa 1999b: 349. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


obtusidentata. Malaysia. 

Pseudoperomyia obtusidentata Jaschhof & Hippa 1999b: 350. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya 


oculibunda. Malaysia. 

Pseudoperomyia oculibunda Jaschhof & Hippa 1999b: 356. Malaysia: Selangor, Genting Highlands, Gunung Ulu 

Kali; HT; ♂; NHRS. 

orophila. Malaysia. 

Pseudoperomyia orophila Jaschhof & Hippa 1999b: 361. Malaysia: Selangor, Genting Highlands, Gunung Ulu Kali; 


parvolobata. Malaysia. 

Pseudoperomyia parvolobata Jaschhof & Hippa 1999a: 155. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya 


platystyla. Malaysia. 

Pseudoperomyia platystyla Jaschhof & Hippa 1999a: 162. Malaysia: Selangor, Gunung Ulu Kali; HT; ♂; NHRS. 

polyardioides. Malaysia. 

Pseudoperomyia polyardioides Jaschhof & Hippa 1999b: 359. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya

47 


psittacephala. China (Fujian). 

Pseudoperomyia psittacephala Li & Bu 2001a: 81. China: Fujian, Wuyishan; HT; ♂; NKUM. 

pyramidata. Malaysia, Japan (Hokkaido, Shikoku). 

Pseudoperomyia pyramidata Jaschhof & Hippa 1999b: 362. Malaysia: Selangor, Genting Highlands, Gunung Ulu 

Kali; HT; ♂; NHRS. 

trispinata. Malaysia. 

Pseudoperomyia trispinata Jaschhof & Hippa 1999b: 345. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


variabilis. Malaysia. 

Pseudoperomyia variabilis Jaschhof & Hippa 1999b: 352. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


velata. Japan (Ryukyu Is). 

Pseudoperomyia velata Jaschhof 2000: 32. Japan: Ryukyu Is, Okinawa, Yambaru Peninsula, Kunigami; HT; ♂; 

KUEC. 

ventricosa. Malaysia. 

Pseudoperomyia ventricosa Jaschhof & Hippa 1999b: 358. Malaysia: Selangor, Ulu Gombak, nr Univ. Malaya Field 


*PTERIDOMYIA 

Pteridomyia Jaschhof 2003b: 111. Type species, Cordylomyia gressitti Yukawa (orig. des.). 

Australasian; 2 spp.; Pteridomyiini. 

*bilobata. New Zealand. 

Pteridomyia bilobata Jaschhof 2003b: 114. New Zealand: North I., Taupo, Pureora For., Waipapa Res.; HT; ♂; 

NZAC. 

gressitti. New Zealand. 

Cordylomyia gressitti Yukawa 1964: 300. New Zealand: Campbell I., Tucker Cove; HT; ♂; NZAC. Ref.: Jaschhof & 

Jaschhof 2003: 112. 

SKUHRAVIANA 

Skuhraviana Mamaev 1963a: 439. Type species, triangulifera Mamaev (orig. des.). 

Holarctic; 1 spp.; Bryomyiini. Ref.: Jaschhof 1998b (rev.); Jaschhof & Jaschhof 2009 (redescr.). 

triangulifera. Fennoscandia, Estonia, Lavia, Germany, Ukraine, Russia (Near East, Far East), USA (Virginia). 

Skuhraviana triangulifera Mamaev 1963a: 440. Russia: Krasnodarskiï Terr., Novo-Prokhladnoye; HT; ♂; HYNM. 

Skuhraviana camelina Mamaev & Berest 1991a: 31. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; HYNM. 

Skuhraviana media Mamaev 1994: 29. Russia: Kamchatka Peninsula, Kozyrevsk; HT; ♂; ZMMU. 

STENOSPATHA 

Stenospatha Kieffer 1913f: 55. Type species, eriophori Kieffer (orig. des. as n. g., n. sp.). 


eriophori. France. 

eriophori Kieffer 1913f: 55 (Stenospatha). France: Haspelscheidt bog, nr Bitche; T; LT; lost.

STROBLIELLA 

Strobliella Kieffer 1898f: 51. Type species, intermedia Kieffer (mon.). 

Holarctic; 2 spp.; Strobliellini. 

brachycornis. Latvia, Germany. 

Strobliella brachycornis Spungis & Jaschhof 2000: 313. Latvia: Jumprava; HT; ♀; FBUL. 

intermedia. UK, Austria, USA (Alaska). 

Strobliella intermedia Kieffer 1898f: 51. Austria: Admont; HT; ♂; lost. 

TEKOMYIA 

Tekomyia Möhn 1960a: 5. Type species, populi Möhn (orig. des.). 

Western Palearctic; 1 sp.; Aprionini. Ref.: Jaschhof 1998b (redescr.). 

populi. Germany. 

Tekomyia populi Möhn 1960a: 6. Germany: Niedersachsen, DGOT, Gartetal; HT; ♂; SMNS. 

TRITOZYGA 

Tritozyga Loew 1862: 179. Type species, sackeni Felt (subs. des. Felt 1911: 32). 

Holarctic; 2 spp.; Catochini. Ref.: Jaschhof 1998b (redescr.). 

sackeni. Canada (British Columbia), USA (New York, District of Columbia). 

Tritozyga sackeni Felt 1911c: 32. USA: District of Columbia; HT; ♀; MCZ. 

Tritozyga fenestra Felt 1914f: 117. USA: New York, Albany; HT; ♀; NYSM. 

Tritozyga borealis Felt 1920a: 278. USA: New York, Keene Valley; LT (Pritchard 1948a: 665); ♀; NYSM. 

Neocatocha cranbrooki Felt 1926f: 266. Canada: British Columbia, Cranbrook; HT; ♀; NYSM. 

tyrestaensis. Sweden. 

Tritozyga tyrestaensis Jaschhof 2002: 437. Sweden: Södermanland, Tyresta Nature Res.; HT; ♂; NHRS. 

YUKAWAMYIA 

Yukawamyia Mamaev & Zaitzev 1996c: 1. Type species, gratia Mamaev & Zaitzev (orig. des.). 

Eastern Palearctic; 1 sp.; Aprionini. 

gratia. Russia (Far East). 

Yukawamyia gratia Mamaev & Zaitzev 1996c: 3. Russia: Kuril Is, Kunashir I., Mendeleevo; HT; ♂; ZMMU. 

Mycromyinae Unplaced to Tribe 

*BERESTELLA 

Berestella Fedotova & Perkovsky 2007a: 82. Type species, insuperabilis Fedotova & Perkovsky (orig. des.). 

Rovno amber (Ukraine); 1 sp. 

* † insuperabilis. Rovno amber (Ukraine). 

48

49 

Berestella insuperabilis Fedotova & Perkovsky 2007a: 82. Ukrainian amber: Rovno, Fedorovka site; HT; ♀; UASK. 

IPOMYIA 

Ipomyia Colless 1965: 145. Type species, bornemisszai Colless (orig. des. as n. g., n. sp.). 

Australasian; 1 sp. 

bornemisszai. Australia (Western Australia). 

Ipomyia bornemisszai Colless 1965: 145. Australia: Western Australia, Perth, King's Pk; HT; ♀; ANIC. 

PSADARIA 

Psadaria Enderlein 1940: 668. Type species, pallida Enderlein (orig. des.). 


pallida. Juan Fernandez Is. 

Psadaria pallida Enderlein 1940: 669. Juan Fernandez Is: Masafuera; ST; ♂,♀; ZMHB?. 

TERMITOMASTUS 

Termitomastus Silvestri 1901: 1. Type species, leptoproctus Silvestri (mon.). 


leptoproctus. Brazil (Minas Gerais), Argentina (Misiones). 

Termitomastus leptoproctus Silvestri 1901: 1. Argentina: Misiones, S. Ana, & Brazil: MG, Cuiabá; ST; ♀; ?. 

TRICAMPYLOMYZA 

Tricampylomyza Kieffer 1919: 198. Type species, parvula Kieffer (orig. des.). 

Afrotropical; 1 sp., a nomen dubium, per M. Jaschhof (in litt.). 

parvula. Cameroon. 

parvula Kieffer 1919: 198 (Tricampylomyza). Cameroon: Kaibi, on Kaibi R.; T; ♀; lost. 

TRICHELOSPATHA 

Trichelospatha Kieffer 1913i: 302. Type species, Campylomyza conigera Kieffer (mon.). 


conigera. France. 

conigera Kieffer 1913i: 302 (Trichelospatha). France: Moselle; T; LT; lost. 

TRICOLPODIA 

Tricolpodia Kieffer 1913g: 55. Type species, Colpodia anomala of Kieffer 1900: 464 (larva only). 

Western Palearctic; 1 sp., a nomen dubium, per M. Jaschhof (in litt.).

anomala. France. 

anomala Kieffer 1898f: 43 (Colpodia). France: Moselle; T; A; lost. 

*VICEMYIA 

Vicemyia Fedotova & Perkovsky 2007a: 90. Type species, immediata Fedotova & Perkovsky (orig. des.). 

Rovno amber (Ukraine); 1 sp. 

* † immediata. Rovno amber (Ukraine). 

Vicemyia immediata Fedotova & Perkovsky 2007a: 90. Ukrainian amber: Rovno; HT; ♂; UASK. 

SUBFAMILY PORRICONDYLINAE 

*ADSUMYIA 

Adsumyia Fedotova & Perkovsky 2008b: 60. Type species, integra Fedotova & Perkovsky (orig. des.). 

Rovno amber (Ukraine; late Eocene); 1 sp.; Dicerurini 

* † integra. Late Eocene - Ukrainian amber. 

Adsumyia integra Fedotova & Perkovsky 2008b: 61. Rovno amber, Ukraine; HT; ♀; UASK. 

*ALOCOLPODIA 

Alocolpodia Fedotova & Sidorenko 2007c: 78. Type species, apta Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Porricondylini. 

*apta. Russia (Far East). 

Alocolpodia apta Fedotova & Sidorenko 2007c: 80. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; 

♂; ZMAS. 

*ANCOREPIDOSIS 

Ancorepidosis Fedotova & Sidorenko 2008a: 236. Type species, electa Fedotova & Sidorenko (orig. des.). 


*electa. Russia (Far East). 

Ancorepidosis electa Fedotova & Sidorenko 2008a: 236. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 

HT; ♂; ZMAS. 

*ANISEPIDOSIS 

Anisepidosis Fedotova & Sidorenko 2005e: 99. Type species, nulla Fedotova & Sidorenko (orig. des.). 

Palearctic; 3 spp.; Porricondylini. 

*modica. Russia (Far East). 

Anisepidosis modica Fedotova & Sidorenko 2007b: 42. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 

50

51 


*nulla. Russia (Far East). 

Anisepidosis nulla Fedotova & Sidorenko 2005e: 101. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

*plicata. Russia (Far East). 

Anisepidosis plicata Fedotova & Sidorenko 2005e: 102. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


ARCTEPIDOSIS 

Arctepidosis Mamaev 1990a: 15. Type species, jamalensis Mamaev (orig. des.). 


jamalensis. Russia (Western Siberia). 

Arctepidosis jamalensis Mamaev 1990a: 15. Russia: Yamal Peninsula, Khadyta; HT; ♂; ZMMU. 

paneliusi. Finland. 

Arctepidosis paneliusi Mamaev & Zaitzev 1998e: 85. Finland: Grankulla; HT; ♂; ZMMU. 

*ARMATEPIDOSIS 

Armatepidosis Fedotova & Sidorenko 2007c: 72. Type species, cuspidis Fedotova & Sidorenko (orig. des.). 


*cuspidis. Russia (Far East). 

Armatepidosis cuspidis Fedotova & Sidorenko 2007c: 74. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


decussata. Germany, Japan (Honshu). 

Porricondyla decussata Yukawa 1971: 67. Japan: Honshu, Wakayama-City, Ichibanchô; HT; ♂; KUIC. 

ASYCOLA 

Asycola Spungis 1991: 197. Type species, connata Spungis (orig. des.). 

Holarctic; 1 sp.; Asynaptini. 

connata. Latvia. 

Asycola connata Spungis 1991: 197 Latvia: Saulkalne; HT; ♂; ZMMU. 

ASYNAPTA 

Asynapta Loew 1850a: 21, 39, as subg. of Cecidomyia. Type species, Cecidomyia longicollis Loew (subs. des. 

Karsch 1877: 14). 

Asinapta Rondani 1860: 287, missp. of Asynapta 

Rubsaamenia Kieffer 1894b: 83. Type species, flava Kieffer (mon.) = pectoralis Winnertz. 

Ruebsaamenia (as Rübsaamenia) Kieffer 1894c: 333, unjust. emend. of Rubsaamenia. 

Camptomyiella (also as Camptomuiella and Camptomuella) Marikovskij 1958a: 139. Type species, abieticola 

Marikovskij (orig. des.) = strobi Kieffer. 

Tjanshaniomyia Marikovskij 1958b: 1848. Type species, brachycera Marikovskij (orig. des.). 

Holarctic, Neotropical, Afrotropical, Oriental; 46 spp.; Asynaptini. Refs: Panelius 1965 (rev., Eur. spp.), Parnell 1971

(review, Nearctic spp.), Spungis 1988 (key, Eur. spp.). 

artocarpi. India (Kerala). 

Rubsaamenia artocarpi Nayar 1949: 83. India: Kerala, Trivandrum; ST; ♀; ZSCI. 

aurangabadensis. India (Maharashtra). 

Asynapta aurangabadensis Sharma 1987b: 170. India: Maharashtra, Aurangabad, Khadkeshwar; HT; ♂; INPC. 

baltica. Latvia, Russia (Far East). 

Asynapta baltica Spungis 1988: 108. Latvia: Bulduri; HT; ♂; LASS. 

borealis. USA (New York). 

Asynapta borealis Felt 1920a: 285. USA: New York, Keene Valley; HT; ♀; NYSM. 

brachycera. Kazakhstan, Russia (Eastern Siberia). 

Tjanshaniomyia brachycera Marikovskij 1958b: 1848. Kazakhstan: Alma-Ata; T; ♂; MARC. 

breviata. Latvia, Russia. 

Asynapta breviata Spungis 1988: 110. Latvia: Darzini; HT; ♂; LASS. 

cactophila. USA (Arizona). 

Ruebsaamenia cactophila Samuelson 1964: 5. USA: Arizona, Pima Co., Santa Rita Mts, Florida Canyon; HT; ♂; 

USNM. 

canadensis. Canada (Quebec), USA (New York). 

Asynapta canadensis Felt 1908b: 421. Canada: Quebec; HT; ♂; MCZC. 

Asynapta dolens Felt 1920a: 285. USA: New York, Keene Valley; HT; ♂; NYSM. 

caudata. USA (New York). 

Asynapta caudata Felt 1908b: 421. USA: New York, Albany; HT; ♂; NYSM. 

cerasi. USA (New York). 

Asynapta cerasi Felt 1907a: 51. USA: New York, Albany; HT; ♂; NYSM. 

cerealis. Germany. 

Tipula cerealis Sauter 1817: 47 (publ'n not seen). Germany: vic. Baden & in Wurtemburg; ST; A,L; lost?. 

citrinae. Puerto Rico. 

Asynapta citrinae Felt 1932c: 117. Puerto Rico: Isabela; ST; ♂,♀,L; USNM. 

communis. Somalia. 

Asynapta communis Mamaev & Zaitzev 1997c: 8. Somalia: Hargeisa; HT; ♂; ZMMU. 

dolichocera. Kazakhstan, Kirghizstan. 

Tjanshaniomyia dolichocera Marikovskij 1958b: 1849. Kazakhstan: Alma-Ata, & Kirghizstan: Frunze; ST; ♂; 

MARC. 

flavida. USA (New York). 

Asynapta flavida Felt 1908b: 421. USA: New York, Nassau; HT; ♂; NYSM. 

frosti. USA (Massachusetts). 

Asynapta frosti Felt 1913f: 143. USA: Massachusetts, Framingham; HT; ♀; NYSM. 

fungivora. India (Uttar Pradesh). 

Asynapta fungivora Grover 1969a: 145. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

furcifer. Italy, Cyprus. 

Asynapta furcifer Barnes 1932a: 51. Italy: Sicily, Patti, & Cyprus; ST; ♂,♀; BMNH. 

gossypii. Barbados. 

Porricondyla gossypii Coquillett 1905: 200. Barbados; ST; ♂,♀,L; USNM. 

*groverae. India (Madhya Pradesh), China (Shandong). 

Dicerura indica Grover 1971: 22, jun. sec. hom. indica Grover. India: Madhya Pradesh, Indore; HT; ♂; PGCA. 

Asynapta groverae Jiang & Bu 2004c: 786, new name for Dicerura indica Grover. 

himalayensis. India (Uttar Pradesh). 

Asynapta himalayensis Jaiswal 1989c: 112. India: Uttar Pradesh, Himalayas, Garhwal Hills, Kotdwara; HT; ♂; CSIA. 

hopkinsi. Canada (British Columbia), USA (Washington, Oregon, California, Montana, Florida). 

Asynapta hopkinsi Felt 1935c: 48. USA: Florida, Lake City; LT (Parnell 1971: 295); ♂; USNM. 

Rubsaamenia keeni Foote 1956: 48. USA: California, Butte Co; HT; ♂; USNM. 

ilicis. Algeria. 

52

53 

Rubsaamenia ilicis Kieffer 1919: 196. Algeria: Djurdjura, forest of Dra-Inguel, Haïzer; T; ♂; lost?. 


Rubsaamenia indica Grover 1964b: 203. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

inflata. Latvia. 

Asynapta inflata Spungis 1988: 111. Latvia: Salaspils; HT; ♂; LASS. 

lacunosa. Somalia. 

Asynapta lacunosa Mamaev & Zaitzev 1997c: 8. Somalia: Hargeisa; HT; ♂; ZMMU. 

laminata. Somalia. 

Asynapta laminata Mamaev & Zaitzev 1997c: 10. Somalia: Hargeisa H; ♂; ZMMU. 

longicollis. Poland. 

Cecidomyia longicollis Loew 1850a: 39. Poland; ST; ♀; ZMHB?. 

Cecidomyia longicauda Loew 1850a: 39. Poland; ST; ♀; ZMHB?. 

magdalini. UK, Finland, Latvia. 

Asynapta magdalini Panelius 1965: 81. Finland: Munksnäs; HT; ♂; MZHF. 

mangiferae. Barbados. 

Asynapta mangiferae Felt 1909a: 299. Barbados; ST; ♂,♀; USNM. 

marilandica. USA (New York, Maryland). 

Asynapta marilandica Felt 1916f: 412. USA: Maryland, Montgomery Co., Plummers I.; HT; ♂; USNM. 

Camptomyia dentata Felt 1920a: 286. USA: New York, Keene Valley; HT; ♂; NYSM. 

Holoneurus inflatus Felt 1920a: 287. USA: New York, Keene Valley; HT; ♂; NYSM. 

mira. Somalia. 

Asynapta mira Mamaev & Zaitzev 1997c: 8. Somalia: Hargeisa; HT; ♂; ZMMU. 

*northi. Seychelles. 

Asinapta northi Spungis 2006a: 45. Seychelles, North Island; HT; ♂; BMNH. 


Rubsaamenia orientalis Grover 1964b: 205. India: Uttar Pradesh, Allahabad; HT; ♀; PGCA. 

pectoralis. UK, Sweden, Finland, Denmark, Netherlands, France, Germany, Latvia, Czechoslovakia, Austria. 

Cecidomyia pectoralis Winnertz 1853: 303. Germany; ST; ♂,♀; MNHN. 

Cecidomyza winnertzii Zetterstedt 1855: 4880. Sweden: Lund; ST; ♂,♀; MZLU. 

Rubsaamenia flava Kieffer 1894b: 83. France: Bitche; ST; LT; lost?. 

Asynapta longipennis Löw 1874a: 148. Austria: Vienna, Wienerwald; ST; ♀; NHMW. 

Ruebsaamenia ruficornis Kieffer 1894c: 334. France: Bitche; ST; ♀; lost?. 

Asynapta macrura Kieffer 1913f: 47. France: Bitche; T; ♀; lost?. 

phragmitis. UK, Sweden, Netherlands, Latvia, Czechoslovakia, Austria, Kazakhstan. 

Cecidomyia phragmitis Giraud 1863: 1260. Austria; T; ♀; lost?. 

Ruebsaamenia harundinea Marikovskij 1961: 44. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, vic. middle & 

lower reaches of Ili R.; ST; ♂,♀; ZMAS. 

populnea. UK, Netherlands, Germany, Finland, Austria. 

Asynapta populnea Panelius 1965: 82. Finland: Helsingfors; HT; ♂; MZHF. 

rufomaculata. Finland. 

Asynapta rufomaculata Panelius 1965: 83. Finland: Tvärminne; HT; ♂; MZHF. 

saliciperda. USA (Illinois, Pennsylvania). 

Asynapta saliciperda Felt 1908b: 421. USA: Illinois, Evanston; LT (Parnell 1971: 295); ♂; NYSM. 

Asynapta americana Felt 1912f: 103. USA: Pennsylvania, Hazelton; HT; ♂; NYSM. 

simplex. Somalia. 

Asynapta simplex Mamaev & Zaitzev 1997c: 8. Somalia: Hargeisa; HT; ♂; ZMMU. 

strobi. Widespread Europe. 

Camptomyia strobi Kieffer 1920a: 20. Germany: vic. Munich; ST; ♂,♀; lost?. 

Camptomuiella abieticola Marikovskij 1958a: 140. Russia: Karelia; ST; ♂,♀; ?. 

Camptomuella larici Marikovskij 1958a: 141. Russia: Orel Prov.; ST; ♂,♀; ?. 

Asynapta laricis Skrzypczy�ska 1977: 185. Poland: Krasiczyn For. Inspectorate, nr Lublin; HT; ♂; SKRP. 

strobilophila. USA (Oregon, California).

Holoneurus strobilophilus Foote 1956: 49. USA: California, Butte Co.; HT; ♂; USNM. 

thuraui. Germany, Latvia, Czech Republic, Hungary, Russia (Europe). 

Asynapta thuraui Rübsaamen 1893: 165. Germany: vic. Berlin; ST; ♂,♀,L; ZMHB. 

Asynapta thurani Kieffer 1894c: 340, missp. of thuraui. 

umbra. USA (New York). 

Asynapta umbra Felt 1914c: 128. USA: New York, Johnstown; HT; ♂; NYSM. 

vasiljevae. Ukraine. 

Asynapta vasiljevae Spungis 1988: 108. Ukraine: Crimea, Yalta; ST; ♂,♀; LASS. 

viscicola. Italy. 

Asynapta viscicola Skuhravá 2007b: 196. Italy: South Tyrol, Tschötsch; HT; ♂,♀; LASS. 

BASICONDYLA 

Basicondyla Parnell 1971: 327. Type species, Porricondyla fultonensis Felt (orig. des.). 

Nearctic; 1 sp.; Porricondylini. 

fultonensis. USA (New York). 

Porricondyla fultonensis Felt 1919b: 222. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

BRITTENIA 

Brittenia Edwards 1941: 2. Type species, fraxinicola Edwards (orig. des.) 

Palearctic; 1 sp.; Heteropezini. Ref.: Wyatt 1967 (redescr.). 

fraxinicola. UK. 

Brittenia fraxinicola Edwards 1941: 3. England: Mulgrave Wood nr Whitby; HT; ♀; BMNH. 

BRYOCRYPTA 

Bryocrypta Kieffer 1896g: 8. Type species, dubia Kieffer (orig. des.). 

Palearctic, Oriental, and Baltic and Ukrainian amber; 16 spp.; Bryocryptini. Ref.: Panelius 1965 (rev., Eur. spp.). 

angustata. Russia (Europe). 

Bryocrypta angustata Mamaev 1966: 230. Russia: Rostov Prov., Nizhnyaya Kundryucheskhaya; HT; ♂; MAMC. 

† asiatica. Turkmenistan. 

Bryocrypta asiatica Mamaev 1966: 231. Turkmenistan: Bakhyz Res.; HT; ♂; ZMMU. 

† capitosa. Upper Eocene - Baltic amber. 

Bryocrypta capitosa Meunier 1904: 26. Baltic amber; ST; ♂,♀; DGOT?. 

* † catenulata. Late Eocene - Ukrainian amber. 

Bryocrypta catenulata Fedotova 2005e: 42. Rovno amber, Ukraine; HT; ♀; UASK. 

* † concinna. Late Eocene - Ukrainian amber. 

Bryocrypta concinna Fedotova 2005e: 43. Rovno amber, Ukraine, Klesoc excavation; HT; ♀; UASK. 

dubia. France. 

Bryocrypta dubia Kieffer 1896g: 8. France: vic. Bitche; ST; ♂,♀,L,egg; lost? 

† elegantula. Upper Eocene - Baltic amber. 

Bryocrypta elegantula Meunier 1904: 27. Baltic amber; HT; ♀; DGOT?. 

† fagioides. Upper Eocene - Baltic amber. 

Bryocrypta fagioides Meunier 1904: 28. Baltic amber; HT; ♂; DGOT?. 

54

† girafa. Upper Eocene - Baltic amber. 

Bryocrypta girafa Meunier 1904: 25. Baltic amber; ST; ♂,♀; DGOT?. 

indubitata. Latvia, Poland, Russia (Europe). 

Bryocrypta indubitata Mamaev 1964c: 903. Russia: Moscow Prov., Bykovo; HT; ♂; ZMMU. 

*vlaqueata. Late Eocene - Ukrainian amber. 

Bryocrypta laqueata Fedotova 2005e: 44. Rovno amber, Ukraine; HT; ♀; UASK. 

lobata. Ukraine. 

Bryocrypta lobata Mamaev 1966: 230. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

maculata. Germany, Yugoslavia, Russia (Europe). 

Bryocrypta maculata Mamaev 1965b: 1667. Russia: Moscow Prov., Zavety Ilyicha; HT; ♂; ZMMU. 

triangulobata. India (Uttar Pradesh). 

Bryocrypta triangulobata Grover 1973: 70. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

vesiculosa. France. 

Bryocrypta vesiculosa Kieffer 1913i: 263. France: Moselle; T; LT; lost?. 

† vetusta. Upper Eocene - Baltic amber. 

Bryocrypta vetusta Meunier 1904: 27. Baltic amber; T; ♂; DGOT?. 

BULBEPIDOSIS 

Bulbepidosis Mamaev 1990a: 14. Type species, tarsalis Mamaev (orig. des.). 

Palearctic; 1 sp.; Dicerurini. 

tarsalis. Ukraine. 

Bulbepidosis tarsalis Mamaev 1990a: 14. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; HYNM. 

CAMPTOMYIA 

Camptomyia Kieffer 1894b: 86. Type species, erythromma Kieffer (mon.). 

Prodirhiza Kieffer 1913i: 269. Type species, Dirhiza multiarticulata Felt (orig. des.). 

Cecidophila Rübsaamen 1916b: 551. Type species, artemisiae Rübsaamen (mon.) = corticalis Loew. 

Neocamptomyia Mamaev 1961f: 1683, as subg. of Camptomyia. Type species, Camptomyia abnormis Mamaev (orig. 

des.). 

Paracamptomyia Mamaev 1961f: 1685, as subg. of Camptomyia. Type species, Camptomyia salicicola Mamaev 

(orig. des.) = antennata Felt. 

Procamptomyia Mamaev 1961f: 1682, as subg. of Camptomyia. Type species, Camptomyia spinifera Mamaev (orig. 

des.). 

Xylocamptomyia Mamaev 1961f: 1689, as subg. of Camptomyia. Type species, Camptomyia heterobia Mamaev 

(orig. des.). 

Holarctic, Neotropical, Oriental, and Baltic and Ukrainian amber; 73 spp.; Asynaptini. Refs: Panelius 1965 (rev., Eur. 

spp.), Parnell 1971 (review, Nearctic spp.), Spungis 1988 (key, Eur. spp.). 

abnormis. Denmark, Finland, Latvia, Poland, Russia (Europe). 

Camptomyia abnormis Mamaev 1961f: 1683. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

accepta. Russia (Far East). 

Camptomyia accepta Mamaev & Zaitzev 1998c: 5. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

addenda. Latvia. 

Camptomyia addenda Spungis 1989: 67. Latvia: Salaspils; HT; ♂; ZMMU. 

aestiva. USA (New York, Pennsylvania). 

Camptomyia aestiva Felt 1912f: 104. USA: Pennsylvania, Hazelton; HT; ♀; NYSM. 

albidula. Russia (Far East). 

55

56 

Camptomyia albidula Mamaev 1975d: 84. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

alnicola. Algeria. 

Camptomyia alnicola Kieffer 1919: 195. Algeria: Kabylia, Yakouren; ST; ♂,♀; lost?. 

antennata. USA (New York), Finland, Latvia, Poland, Russia (Europe). 

Camptomyia antennata Felt 1920a: 286. USA: New York, Keene Valley; HT; ♂; NYSM. 

Camptomyia populicola Mamaev 1961f: 1686. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

Camptomyia salicicola Mamaev 1961f: 1685. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

artocarpi. India (Kerala). 

Camptomyia artocarpi Nayar 1949: 83. India: Kerala, Trivandrum; ST; ♂,♀; ZSCI. 

*attenuata. Russia (Far East). 

Camptomyia attenuata Fedotova 2004i: 589. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

* † aufuga. Late Eocene - Ukrainian amber. 

Camptomyia aufuga Fedotova 2004d: 77. Rovno amber, Ukraine; HT; ♀; UASK. 

aurora. Finland. 

Cecidomyia aurora Mannerheim 1823: 182. Finland; ST; ♂,♀,P,L; lost. 

binotata. France. 

Camptomyia binotata Kieffer 1894c: 324. France: Bitche; LT (Panelius 1965: 94); ♂; NYSM. 

Camptomyia similis Kieffer 1894c: 349, pl. IV, fig. 26. France; T; LT; lost?. 

breviradicis. Japan (Kyushu). 

Camptomyia breviradicis Yukawa 1968a: 146. Japan: Kyushu, Fukuoka Pref., Mt Inunaki; HT; ♂; KUEC. 

calcarata. Russia (Europe), Latvia. 

Camptomyia calcarata Mamaev 1964c: 896. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

canadensis. Canada, USA (New York). 

Dirhiza canadensis Felt 1908b: 420. Canada; HT; ♀; USNM. 

Dirhiza photophila Felt 1908b: 420. USA: New York, Nassau; HT; ♀; NYSM. 

cognata. Russia (Far East). 

Camptomyia cognata Mamaev 1975a: 39. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. 

concolor. France. 

Camptomyia concolor Kieffer 1894c: 331. France: Bitche; ST; ♀,L; lost?. 

conformis. Russia (Far East). 

Camptomyia conformis Mamaev & Zaitzev 1998c: 7. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; MAMC. 

*cornuta. Russia (Far East). 

Camptomyia cornuta Fedotova & Sidorenko 2005e: 123. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 

Kamenushka; HT; ♂; ZMAS. 

corticalis. Widespread northern Europe. 

Cecidomyia corticalis Loew 1851: 377. Poland: Wroclaw; ST; ♂,♀; ZMHB?. 

Epidosis lutescens Kieffer 1888a: 202. France: vic. Bitche; ST; ♂,♀,L; lost?. 

Camptomyia innotata Kieffer 1894c: 329. France: Bitche; ST; ♂,♀,L; lost?. 

Cecidophila artemisiae Rübsaamen 1916b: 552. Germany; ST; ♀,L; lost. 

*crucicula. Russia (Far East). 

Camptomyia crucicula Fedotova & Sidorenko 2005e: 120. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


denticuligera. Russia (Far East). 

Camptomyia denticuligera Mamaev & Zaitzev 1998c: 2. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

derivata. Russia (Far East). 

Camptomyia derivata Mamaev & Zaitzev 1998c: 8. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

drymophila. Russia (Far East). 

Camptomyia drymophila Mamaev & Zaitzev 1998c: 8. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMMU. 

emarginata. UK. 

Camptomyia emarginata Panelius 1965: 100. UK: England; Headington Hill; HT; ♂; MZHF.

57 

erythromma. France. 

Epidosis erythromma Kieffer 1888a: 200. France: vic. Bitche; ST; ♂,♀,L; lost?. 

fenestralis. Switzerland. 

Cecidomyia fenestralis Bremi 1847a: 55. Switzerland: Zurich; ST; A; ETHZ?. 

flavocinerea. Finland, Latvia, Russia (Europe). 

Camptomyia flavocinerea Panelius 1965: 96. Finland: Tvärminne; HT; ♂; MZHF. 

fulva. Denmark, Latvia, Russia (Europe). 

Camptomyia fulva Mamaev 1961f: 1688. Russia: Tula Prov., Krapiven For.; HT; ♂; ZMMU. 

Camptomyia juglandis Panelius 1965: 91. Denmark: Bagsvaerd; HT; ♂; MZHF. 

furcellata. Russia (Far East). 

Camptomyia furcellata Mamaev & Zaitzev 1998c: 3. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

gigantea. Latvia. 

Camptomyia gigantea Spungis 1989: 63. Latvia: Valmiera; HT; ♂; ZMMU. 

helveola. Germany. 

Epidosis helveolus Rübsaamen 1890b: 18. Germany: vic. Siegen; T; ♀; ZMHB. 

heterobia. Latvia, Russia (Europe). 

Camptomyia heterobia Mamaev 1961f: 1689. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

hibisci. India (Tamil Nadu) 

Camptomyia hibisci Felt 1926i: 241. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

incognita. Russia (Far East). 

Camptomyia incognita Mamaev & Zaitzev 1998c: 10. Russia: Kuril Is, Kunashir I., Mendeleevo; HT; ♂; ZMMU. 

mamaevi. Russia (Far East). 

Camptomyia mamaevi Spungis 1989: 65. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

maritima. Russia (Far East). 

Camptomyia maritima Mamaev 1972b: 113. Russia: Primorskiï Terr., Ussuriyskiï Distr., Suputinskiï Res.; HT; ♂; 

HYNM. 

maxima. Finland, Latvia, Russia (Europe). 

Camptomyia maxima Mamaev 1961f: 1686. Russia: Tula Prov., Krapiven For.; HT; ♂; HYNM. 

minima. Latvia. 

Camptomyia minima Spungis 1989: 68. Latvia: Valmiera; HT; ♂; ZMMU. 

montana. USA (New York, New Hampshire). 

Dirhiza montana Felt 1908b: 420. USA: New Hampshire, White Mtns; HT; ♀; USNM. 

Camptomyia tsugae Felt 1913d: 214. USA: New York, New York City; LT (Parnell 1971: 300); ♂; NYSM. 

morindae. India (Tamil Nadu). 

Camptomyia morindae Felt 1926i: 242. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

mucronata. Russia (Far East). 

Camptomyia mucronata Mamaev & Zaitzev 1998c: 7. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

multiarticulata. USA (New York, New Jersey). 

Dirhiza multiarticulata Felt 1908b: 420. USA: New York, Palisades; HT; ♀; NYSM. 

multinoda. USA (Illinois, New York), UK, Latvia, Poland, Austria, Russia (Europe, Far East), Uzbekistan. 

Porricondyla multinoda Felt 1908b: 419. USA: Illinois, Lake Forest; HT; ♂; NYSM. 

Camptomyia pectinata Felt 1920a: 287. USA: New York, Keene Valley; HT; ♂; NYSM. 

Camptomyia tiliarum Mamaev 1961f: 1686. Russia: Tula Prov., Krapiven For.; HT; ♂; ZMMU. 

multinodica. Philippines. 

Rubsaamenia multinoda Felt 1915a: 198, jun. sec. hom. multinoda Felt. Philippines: Mindanao, Fort Pikit; HT; ♀; 


Camptomyia multinodica Gagné 2004b: 41, new name for Rubsaamenia multinoda Felt. 

nigricornis. France. 

Camptomyia nigricornis Kieffer 1894c: 331. France: Bitche; ST; ♂,♀,,P,L; lost?. 

nodicornis. Germany, Russia (Europe). 

Cecidomyia nodicornis Winnertz 1853: 300. Germany; ST; ♂,♀; lost. 

Cecidomyia nemoralis Winnertz 1853: 302. Germany; ST; ♀; lost.

58 

pallida. France. 

Camptomyia pallida Kieffer 1894c: 326. France: Bitche; ST; ♀,L; lost?. 

panelii. USA (New York). 

Camptomyia montana Felt 1915j: 180, jun. sec. hom. Dirhiza montana. USA: New York, Wells; HT; ♂; NYSM. 

Camptomyia panelii Parnell 1971: 300, new name for Camptomyia montana Felt. 

parrishi. Guyana. 

Porricondyla parrishi Felt 1915h: 152. Guyana: Bartica; HT; ♂; NYSM. 

piceae. Denmark, Finland, Latvia, Ukraine, Russia (Eastern Siberia, Far East). 

Camptomyia piceae Panelius 1965: 91. Denmark: Bagsvaerd; HT; ♂; MZHF. 

pinicola. Finland, Latvia, Russia (Europe). 

Camptomyia pinicola Mamaev 1961f: 1687. Russia: Moscow Prov., Levinskaya; HT; ♂; HYNM. 

Camptomyia similis Mamaev 1961f: 1688, jun. prim. hom. similis Kieffer. Russia: Dagestan, Tlyarata; HT; ♂; 

MAMC. 

Camptomyia dubia Mamaev 1966: 236, new name for Camptomyia similis Mamaev. 

piptopori. Finland, Latvia. 

Camptomyia piptopori Panelius 1965: 98. Finland: Koli; HT; ♂; MZHF. 

populi. France. 

Cecidomyia populi Dufour 1841: 261. France; ST; ♂,♀,P,L; lost. 

pseudotsugae. Canada (British Columbia), USA (Washington, Oregon). 

Camptomyia pseudotsugae Hedlin & Johnson 1968: 532. USA: Washington, Centralia; HT; ♂; CNCI. 

recta. France. 

Camptomyia recta Kieffer 1896g: 27. France: Moselle; ST; ♂,P; lost?. 

regia. Latvia. 

Camptomyia regia Spungis 1989: 69. Latvia: Valmiera; HT; ♂; ZMMU. 

ribiformis. India (Uttar Pradesh). 

Camptomyia ribiformis Jaiswal 1991a: 76. India: Uttar Pradesh, Himalayas, Garhwal Hills, Kotdwara; HT; ♂; CSIA. 

ricini. India (Tamil Nadu). 

Camptomyia ricini Felt 1921j: 23. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

sajanorum. Russia (Western Siberia). 

Camptomyia sajanorum Mamaev & Zaitzev 1998c: 5. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

shibuyai. Japan (Kyushu). 

Camptomyia shibuyai Yukawa 1968a: 144. Japan: Kyushu, Fukuoka Pref., Mt Inunaki; HT; ♂; KUEC. 

sickliformis. India (Uttar Pradesh). 

Camptomyia sickliformis Jaiswal 1988b: 120. India: Uttar Pradesh, Himalayas, Garhwal Hills, Kotdwara; HT; ♂; 

CSIA. 

† 

sinuosa. Upper Eocene - Baltic amber. 

Camptomyia sinuosa Meunier 1904: 36. Baltic amber; T; ♀; lost. 

spinifera. Finland, Latvia, Poland, Russia (Europe), Japan. 

Camptomyia spinifera Mamaev 1961f: 1682. Russia: Tula Prov., Krapiven For.; HT; ♂; HYNM. 

stylosa. Russia (Far East). 

Camptomyia stylosa Mamaev 1994: 30. Russia: Kamchatka, Kozyrevsk; HT; ♂; HYNM. 

subepidermalis. Russia (Far East). 

Camptomyia subepidermalis Mamaev & Zaitzev 1998c: 11. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMMU. 

szadziewskii. Latvia. 

Camptomyia szadziewskii Spungis 1989: 70. Latvia: Valmiera; HT; ♂; ZMMU. 

triangularis. India (Uttar Pradesh). 

Camptomyia triangularis Jaiswal 1991a: 78. India: Uttar Pradesh, Himalayas, Garhwal Hills, Kotdwara; HT; ♂; 

CSIA. 

*triplicata. Russia (Far East). 

Camptomyia triplicata Fedotova 2004i: 589. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS.

ulmicola. Latvia, Poland, Russia (Europe). 

Camptomyia ulmicola Mamaev 1961f: 1684. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

unisaetosa. Latvia. 

Camptomyia unisaetosa Spungis 1989: 64. Latvia: Valmiera; HT; ♂; ZMMU. 

valvata. Germany. 

Cecidomyia valvata Winnertz 1853: 302. Germany; ST; ♀; lost. 

webii. India (Uttar Pradesh). 

Camptomyia webii Kashyap 1989b: 99. India: Uttar Pradesh, Allahabad, Ashok Nagar; HT; ♂; PGCA. 

CASSIDOIDES 

Cassidoides Mamaev 1960a: 1521. Type species, pini Mamaev (orig. des. as n. g., n. sp.). 

Palearctic; 8 spp.; Holoneurini. Ref.: Mamaev 2001c (review). 

bifurcatus. Russia (Far East). 

Cassidoides bifurcatus Mamaev 2001c: 5. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; ♂; ZMMU. 

caucasicus. Russia (Europe). 

Cassidoides caucasicus Mamaev 2001c: 5. Russia: Krasnodarskiï Terr., Krasnaya Polyana; ♂; ZMMU. 

corticalis. Russia (Europe). 

Holoneurus corticalis Mamaev 1964c: 908. Russia: Tula Prov., Krapivenskiï For.; HT; ♂; ZMMU. 

fulviventris. Russia (Europe). 

Holoneurus fulviventris Mamaev 1964c: 908. Russia: Yaroslavl Prov., Volkovo-Rassokhino; HT; ♂; ZMMU. 

omnivagus. Ukraine. 

Cassidoides omnivagus Mamaev 2001c: 6. Ukraine: Zakarpatskaya Prov., Rakhov, Mincul'; ♂; ZMMU. 

pini. UK, Finland, Russia (Europe). 

Cassidoides pini Mamaev 1960a: 1523. Russia: Moscow Prov., Levinskaja; ST; ♂,♀; HYNM. 

raptor. Russia (Europe). 

Holoneurus elongatus Mamaev 1964c: 908, jun. prim. hom. elongatus Felt. Russia: Yaroslavl Prov., 

Volkovo-Rassokhino; HT; ♂; ZMMU. 

Holoneurus raptor Mamaev 1966: 236, new name for Holoneurus elongatus Mamaev. 

*volgensis. Russia (Europe). 

Cassidoides volgensis Fedotova & Sidorenko 2007d: 254. Russia: Middle Volga, Ust-Kinelskii, 30 km SW Samara; 


CEDROCRYPTA 

Cedrocrypta Kieffer 1919: 194. Type species, montana Kieffer (mon.) 

Palearctic, Oriental; 2 spp.; Porricondylini. Ref.: Panelius 1965 (redescr.). 

bifidata. India (West Bengal). 

Cedrocrypta bifidata Grover 1979b: 8. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

montana. Algeria, Germany. 

Cedrocrypta montana Kieffer 1919: 194. Algeria: Djurdjura, Crète du Haïzer; ST; ♂,♀; lost?. 

CLASPETTOMYIA 

Claspettomyia Grover 1964b: 198. Type species, octoclaspettii Grover (mon.). 

Pachylabis Panelius 1965: 54. Type species, Cecidomyza niveitarsis Zetterstedt (orig. des.). 

Claspittomyia Deshpande 1982: 41, missp. of Claspettomyia. 

59

60 

Holarctic, Oriental; 30 spp.; Porricondylini. Refs: Panelius 1965 (rev., Eur. spp.), Parnell 1971 (review, Nearctic 

spp.), Jiang & Bu 2004a (review, Chinese spp.). 

bidentata. USA (New York). 

Porricondyla bidentata Felt 1920a: 283. USA: New York, Keene Valley; HT; ♂; NYSM. 

carpatica. Ukraine. 

Claspettomyia carpatica Mamaev 1998c: 1. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

chrysanthemi. UK, Sweden, Germany, Latvia, Lithuania, Poland, Ukraine, Yugoslavia. 

Pachylabis chrysanthemi Panelius 1965: 56. Sweden: Lund; HT; ♂; MZHF. 

corniculata. Russia (Far East). 

Claspettomyia corniculata Mamaev 1998c: 2. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

dentata. India (Maharashtra). 

Claspettomyia dentata Kashyap 1988d: 126. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

formosa. Switzerland, China (Yunnan, Jilin). 

Cecidomyia formosa Bremi 1847a: 47. Switzerland: Hottingerberg For.; HT; ♂; ETHZ. 

hamata. USA (Massachusetts, North Carolina), UK, Netherlands, Germany, Finland, Latvia, Poland, Ukraine, Croatia. 

Porricondyla hamata Felt 1907a: 49. USA: North Carolina, Davidson's R.; HT; ♂; NYSM. 

Porricondyla johnsoni Felt 1920a: 282. USA: Massachusetts, Brookline; HT; ♂; NYSM. 

Dicroneurus montanus Mamaev 1965a: 267. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; HYNM. 


Claspettomyia indica Rao & Sharma 1978: 115. India: Maharashtra, Aurangabad; HT; ♂; RCMU. 

kirghizica. Kirghizstan. 

Claspettomyia kirghizica Mamaev 1998c: 2. Kirghizstan: Sary-Chelek Res.; HT; ♂; ZMMU. 

*longicornis. China (Shaanxi). 

Claspettomyia longicornis Jiang & Bu 2004a: 353. China: Shaanxi, Fengxian, Qinling Stn; HT; ♂; NKUM. 

montiviva. Kirghizstan. 

Claspettomyia montiviva Mamaev 1998c: 2. Kirghizstan: Sary-Chelek Res.; HT; ♂; ZMMU. 

nipponensis. China (Sichuan,, Gansu, Hubei), Japan (Kyushu). 

Claspettomyia nipponensis Yukawa 1971: 74. Japan: Kyushu, Fukuoka City, Ropponmatsu; HT; ♂; KUIC. 

niveitarsis. Norway, Finland, Poland, Latvia, Russia, China (Guizhou). 

Cecidomyza niveitarsis Zetterstedt 1850: 3688. Norway: Verdal, Garnes; HT; ♂; MZLU. 

octoclaspettii. India (Uttar Pradesh). 

Claspettomyia octoclaspettii Grover 1964b: 198. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

oculata. Turkmenistan, Kirghizstan. 

Claspettomyia oculata Mamaev 1986: 66. Turkmenistan: Kopetdag, Ipaï-Kala; HT; ♂; HYNM. 

orientalis. Taiwan. 

Claspettomyia orientalis Yukawa 1968b: 252. Taiwan: Alishan, Chiayi Hsien; HT; ♂; KUEC. 

paneliusi. Ukraine. 

Claspettomyia paneliusi Mamaev 1998c: 2. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

perlongitegminis. Japan (Honshu). 

Claspettomyia perlongitegminis Yukawa 1971: 79. Japan: Honshu, Gunma Pref., Tone, Marunuma; HT; ♂; KUIC. 

pini. USA (New York, Florida). 

Porricondyla pini Felt 1907a: 48. USA: New York, Albany; HT; ♂; NYSM. 

Porricondyla pennulae Felt 1921c: 142. USA: Florida, St. Augustine; HT; ♂; NYSM. 

quadriclaspettii. India (Uttar Pradesh). 

Claspettomyia quadriclaspettii Jaiswal 1991b: 103. India: Uttar Pradesh, Himalayas, Garhwal Hills, Kotdwara; HT; 

♂; CSIA. 

*retrorsa. China (Yunnan). 

Claspettomyia retrorsa Jiang & Bu 2004a: 353. China: Yunnan, Pingbian, Mt Dawei; HT; ♂; NKUM. 

rossica. Russia (Europe). 

Claspettomyia rossica Mamaev 1998c: 3. Russia: Kursk, Kursk Res.; HT; ♂; ZMMU. 

serrata. China (Fujian, Shaanxi, Gansu), Japan (Kyushu, Shikoku).

Claspettomyia serrata Yukawa 1971: 77. Japan: Kyushu, Fukuoka Pref., Mt Inunaki; HT; ♂; KUIC. 

serrulata. India (Maharashtra). 

Claspettomyia serrulata Kashyap 1988d: 128. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

spinii. India (Uttar Pradesh). 

Didactylomyia spinii Grover 1964b: 201. India: Uttar Pradesh, Allahabad, Tagore Town; HT; ♂; PGCA. 

spinosa. India (Maharashtra). 

Claspettomyia spinosa Kashyap 1988d: 128. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

tenuiforceps. Japan (Honshu). 

Claspettomyia tenuiforceps Yukawa 1971: 76. Japan: Honshu, Morioka City, Kuriyagawa; HT; ♂; KUIC. 

toelgi. Austria. 

Epidosis toelgi (as tölgi) Kieffer 1913a: 234. Austria; T; ♂; lost?. 

ussuriensis. Russia (Far East). 

Claspettomyia ussuriensis Mamaev 1998c: 3. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

zatsepini. Somalia. 

Claspettomyia zatsepini Mamaev 1998c: 3. Somalia: Hargeisa; HT; ♂; ZMMU. 

CLINOPHAENA 

Clinophaena Kieffer 1913i: 268. Type species, Holoneurus mahensis Kieffer (mon.). 

Clinophena Mani 1946: 196, missp. of Clinophaena. 

Afrotropical; 1 sp.; unplaced to tribe. 

mahensis. Seychelles Is. 

Holoneurus mahensis Kieffer 1911: 328. Seychelles Is: Mahé, Cascade Estate; HT; ♂; BMNH. 

CLINORHYTIS 

Clinorhytis Kieffer 1896g: 30. Type species, Clinorhiza flavitarsis Kieffer (orig. des.). 

Palearctic; 1 sp.; Winnertziini. Ref.: Spungis 1992 (redescr.). 

flavitarsis. France, Latvia, Ukraine, Russia (Far East). 

Clinorhiza flavitarsis Kieffer 1895n: cccxix. France: vic. Bitche; NT (Spungis 1992: 10, Latvia: Saulkalne); ♂; 

LASS. 

*COCCOPSILIS 

Coccopsis Meijere 1901: 1, preocc. Reichenbach 1850. Type species, marginata Meijere (orig. des. as n. g., n. sp.). 

*Coccopsilis Harris 2004b: 40, new name for Coccopsis Meijere. 

Holarctic; 5 spp.; Holoneurini. Refs: Panelius 1965 (redescr.), Mamaev 2001c (review). 

*alexandrae. Russia (Far East). 

Coccopsis alexandrae Fedotova & Sidorenko 2007d: 244. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya 

Bay; HT; ♂; ZMAS. 

formosa. Russia (Far East). 

Cassidoides formosus Mamaev 2001c: 7. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; ♂; ZMMU. 

*indigena. Russia (Far East). 

Coccopsis indigena Fedotova & Sidorenko 2007d: 247. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


marginata. UK, Netherlands, Germany, Finland, Russia (Europe), Japan, USA (New Jersey). 

61

Coccopsis marginata Meijere 1901: 1. Netherlands: Amsterdam; LT (Yukawa & Nijveldt 1975: 106); ♂; ZMAN. 

Holoneurus obscurus Mamaev 1964c: 908. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

positiva. Russia (Far East). 

Coccopsis positiva Mamaev 1998d 5. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

*COGNITEPIDOSIS 

Cognitepidosis Fedotova & Sidorenko 2005e: 103. Type species, lepida Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Holoneurini (Fedotova & Sidorenko 2007d). 

*lepida. Russia (Far East). 

Cognitepidosis lepida Fedotova & Sidorenko 2005e: 104. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

COLOMYIA 

Colomyia Kieffer 1892e: 259. Type species, clavata Kieffer (mon.). 

Holarctic, Oriental; 5 spp.; Asynaptini. Ref.: Panelius 1965 (redescr.) 

appendiculata. UK, France. 

Colomyia appendiculata Kieffer 1901a: 36. France: Bitche; T; ♀; lost?. 

caricis. Germany. 

Colomyia caricis Rübsaamen 1899c: 566. Germany; T; ♂; ZMHB. 

caudata. Latvia, Ukraine. 

Colomyia caudata Spungis 1991: 200. Latvia: Skulte; HT; ♂; ZMMU. 

clavata. France, Germany, Latvia, Poland, Russia (Europe). 

Colomyia clavata Kieffer 1892e: 259. France: Bitche; ST; ♂,♀,P,L,egg; lost?. 

Colomyia minor Kieffer 1892e: 261, as var. of clavata Kieffer. France: Bitche; ST; ♂,♀,L; lost?. 

Colomyia picta Kieffer 1892e: 262, as var. of clavata Kieffer. France: Bitche; ST; ♀; lost?. 

corticii. Indonesia (Java). 

Colomyia corticii Edwards 1923: 540. Indonesia: Java, Bogor; ST; ♂,♀; BMNH. 

COLPODIA 

Colpodia Winnertz 1853: 185. Type species, Cecidomyia angustipennis Winnertz (mon.). 

Copolodia Shinji 1944: 164, missp. of Colpodia. 

Palearctic, Oriental, and Baltic amber; 6 spp.; Porricondylini. Ref.: Panelius 1965 (redescr.). 

angustipennis. Netherlands, Germany. 

Cecidomyia angustipennis Winnertz 1853: 293. Germany; ST; ♀; lost. 

† brevicornis. Upper Eocene - Baltic amber. 

Colpodia brevicornis Meunier 1904: 32. Baltic amber; T; ♀; DGOT?. 

† curvinervis. Upper Eocene - Baltic amber. 

Colpodia curvinervis Meunier 1904: 31. Baltic amber; ST; ♂,♀; DGOT?. 

dhakae. India (Uttar Pradesh). 

Colpodia dhakae Mani 1935: 430. India: Uttar Pradesh, Dehra Dun, Lachiwala; HT; ♀; ZSCI. 

fletcheri. India (Bihar). 

Colpodia fletcheri Felt 1916g: 400. India: Bihar, Pusa; HT; ♂; NYSM. 

62

† xylophaga. Upper Eocene - Baltic amber. 

Colpodia xylophaga Meunier 1904: 30. Baltic amber; ST; ♂,♀; DGOT?. 

CRETOHAPLUSIA 

Cretohaplusia Arillo & Nel 2000: 287. Type species, ortunoi Arillo & Nel (orig. des.). 

Spanish amber, Lower Cretaceous; 1 sp.; Diallactini. 

†ortunoi. Lower Cretaceous amber - Spain. 

Cretohaplusia ortunoi Arillo & Nel 2000: 287. Lower Cretaceous amber, Spain: Alava, nr Peñacerrada; HT; ♂; 

MCNA. 

CRETOMIASTOR 

Cretomiastor Gagné 1977a: 62. Type species, ferejunctus Gagné (orig. des.). 

Cretaceous Canadian amber; 1 sp.; Heteropezini. 

†ferejunctus. Upper Cretaceous - Canadian amber (Alberta). 

Cretomiastor ferejunctus Gagné 1977a: 62. Cretaceous amber - Canada: Alberta, nr Medicine Hat; HT; ♀; CNCI. 

CRETOWINNERTZIA 

Cretowinnertzia Gagné 1977a: 60. Type species, angustala Gagné (orig. des.). 

Canadian amber; 1 sp.; Winnertziini. 

† angustula. Upper Cretaceous - Canadian amber (Manitoba). 

Cretowinnertzia angustula Gagné 1977a: 60. Cretaceous amber - Canada: Manitoba, Cedar Lake; HT; ♀; CNCI. 

*CRUSTEPIDOSIS 

Crustepidosis Fedotova & Sidorenko 2008b: 374. Type species, delicata Fedotova & Sidorenko (orig. des). 


*delicata. Russia (Far East). 

Crustepidosis delicata Fedotova & Sidorenko 2008b: 374. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


CRYPTONEURUS 

Cryptoneurus Mamaev 1964c: 909. Holoneurus muscicolus Kieffer (orig. des.). 

Palearctic; 3 spp.; Holoneurini. Ref.: Mamaev 2001c (review). 

muscicola. France, Romania, Russia (Europe). 

Holoneurus muscicola Kieffer 1896g: 16. France: Bitche; ST; ♂,♀,P,L; lost. Mamaev (1964c: 909) invalidly 

designated a neotype; no characters were given to distinguish this species from congeners. 

nigridens. Russia (Europe). 

Cryptoneurus nigridens Mamaev 1964c: 910. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

63

tridentatus. France. 

Holoneurus tridentatus Kieffer 1896g: 15. France: Bitche; ST; ♂,♀,L; lost?. 

CRYPTOXYLOMYIA 

Cryptoxylomyia Mamaev 1996: 65. Rhipidoxylomyia excavata Mamaev (orig. des.). 

Palearctic; 2 spp.; Winnertziini. Ref.: Fedotova & Sidorenko 2007c: 100 (rev.). 

excavata. Russia (Europe), Algeria. 

Rhipidoxylomyia excavata Mamaev 1964c: 900. Russia: Vologda Prov., Kadnikovskaya; HT; ♂; ZMMU. 

perfecta. Russia (Europe). 

Rhipidoxylomyia perfecta Mamaev 1998d: 4. Russia: Krasnodarskiï Terr., Ubinskaya; HT; ♂; ZMMU. 

DALLAIELLA 

Dallaiella Mamaev 1997a: 70. Type species, petrosa Mamaev (orig. des.). 

Palearctic; 1 sp.; Holoneurini. Ref.: Mamaev 2001c (review). 

petrosa. Italy. 

Dallaiella petrosa Mamaev 1997a: 70. Italy: Siena, Monte Amiata; HT; ♂; ZMMU. 

DENDREPIDOSIS 

Dendrepidosis Mamaev 1990a: 21. Type species, trispinula Mamaev (orig. des.). 


bifidula. Russia (Europe). 

Dendrepidosis bifidula Mamaev & Zaitzev 1998e: 86. Russia: Yaroslavl Prov., Volkovo; HT; ♂; ZMMU. 

longipennis. Latvia. 

Dendrepidosis longipennis Spungis 1981: 50. Latvia: Darzini; HT; ♂; ZMMU. 

*obstipa. Russia (Far East). 

Dendrepedosis obstipa Fedotova & Sidorenko 2008a: 231. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 

America; HT; ♂; ZMAS. 

trispinula. Russia (Europe). 

Dendrepidosis trispinula Mamaev 1990a: 21. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

DENTEPIDOSIS 

Dentepidosis Mamaev 1990a: 18. Type species, ussuriensis Mamaev (orig. des.). 



Dentepidosis ussuriensis Mamaev 1990a: 19. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

DESERTEPIDOSIS 

Desertepidosis Mamaev & Soyunov 1989: 65. Type species, Stackelbergiella phrygamophila Mamaev (orig. des.). 

64


pallida. Turkmenistan. 

Desertepidosis pallida Mamaev & Sojunov 1989: 66. Turkmenistan: Sarykamysh; HT; ♂; ZMMU. 

phryganophila. Turkmenistan. 

Stackelbergiella phrygamophila (also as phryganophila) Mamaev 1986: 66. Turkmenistan: Kushka, Badkhyz Res.; 

HT; ♂; HYNM. 

DIALLACTIA 

Diallactes Kieffer 1894c: 347, preocc. Reichenbach 1850. Type species, croceus Kieffer (orig. des.). 

Diallactia Gagné 2004b: 45, new name for Diallactes Kieffer. 

Holarctic; 7 spp.; Diallactini. Ref.: Panelius 1965 (redescr.). 

croceus. France, Latvia. 

Diallactes croceus Kieffer 1894c: 347. France; ST; A; lost?. 

fulvus. Ukraine. 

Diallactes fulvus Mamaev 1966: 221. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; ZMMU. 

gracilis. USA (Maryland). 

Diallactes gracilis Plakidas 1999: 125. USA: Maryland, Murray Hill Rd., 7 mi. S Columbia; HT; ♂; USNM. 

incomparabilis. Russia (Europe). 

Diallactes incomparabilis Mamaev 1998d: 3. Russia: Moscow Prov., Lugovaya; HT; ♂; ZMMU. 

obscuripes. Finland, Latvia. 

Diallactes obscuripes Spungis 1985: 51. Latvia: Saulkalne; HT; ♂; LASS. 

ocellaris. Russia (Europe). 

Diallactes ocellaris Mamaev 1964c: 901. Russia: Krasnodarskiï Terr., Novo-Prokhladnoye; HT; ♂; ZMMU. 

subcroceus. Ukraine. 

Diallactes subcroceus Mamaev 1968a: 613. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; HYNM. 

DICERURA 

Dicerura Kieffer 1898f: 57. Type species, scirpicola Kieffer (mon.). 

Iridomyza Rübsaamen 1899a: 67. Type species, kaltenbachii Rübsaamen (mon.) = iridis Kaltenbach. 

Hormosomyia Felt 1919b: 220. Type species, oregonensis Felt (orig. des.). 

Ulmomyia Mamaev 1960a: 1521. Type species, rossica Mamaev (orig. des. as n. g., n. sp.). 

Neosynepidosis Parnell 1971: 313. Type species, Winnertzia furcata Felt (orig. des.). 

Holarctic, Oriental; 28 spp.; Dicerurini. Ref.: Panelius 1965 (redescr.). 

adunca. Canada (Ontario). 

Dicerura adunca Borkent 1990: 574. Canada: Ontario, Wentworth Co., Ancaster Spring Valley Conservation Area; 

HT; ♂; CNCI. 

barbata. Ukraine. 

Dicerura barbata Mamaev 1966: 226. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; HYNM. 

carpiensis. Canada (Yukon Terr., Ontario), USA (New Jersey). 

Dicerura carpiensis Borkent 1990: 575. Canada: Ontario, E Carp; HT; ♂; CNCI. 

complicata. Latvia. 

Dicerura complicata Spungis 1987: 27. Latvia: Saulkalne; HT; ♂; LASS. 

cooperi. Canada (Newfoundland). 

Dicerura cooperi Borkent 1990: 576. Canada: Newfoundland, 3 km N Picadilly; HT; ♂; CNCI. 

curva. Canada (Ontario, Nova Scotia). 

65

66 

Dicerura curva Borkent 1990: 576. Canada: Ontario, Springwater Conservation Area, nr Aylmer; HT; ♂; CNCI. 

*cymbiformis. Russia (Far East). 

Dicerura cymbiformis Fedotova & Sidorenko 2008b: 363. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


dentata. Latvia. 

Dicerura dentata Spungis 1979: 84. Latvia: Moritsala Res.; HT; ♂; IEME. 

*divaricata. Russia (Far East). 

Dicerura divaricata Fedotova 2004i: 591. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

elongata. USA (Arizona). 

Dicerura elongata Borkent 1990: 579. USA: Arizona, Cochise Co., Portal; HT; ♂; CNCI. 

foliicola. Russia (Far East). 

Dicerura foliicola Mamaev 1968a: 614. Russia: Primorskiï Terr., Ussuriyskiï Distr., Suputinskiï Res.; HT; ♂; 

ZMMU. 

formosa. Russia (Western Siberia). 

Dicerura formosa Mamaev 1998d: 4. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

fungicola. Russia (Europe). 

Ulmomyia fungicola Mamaev 1964c: 904. Russia: Moscow Prov., Glubokoye L.; HT; ♂; ZMMU. 

furcata. USA (New York). 

Winnertzia furcata Felt 1907a: 52. USA: New York, Nassau; HT; ♂; NYSM. 

furculata. Finland, Latvia, Ukraine. 

Dicerura furculata Mamaev 1968a: 614. Ukraine: Zakarpatskaya Prov., Rakhov; HT; ♂; HYNM. 

*infundibularis. Russia (Far East). 

Dicerura infundibularis Fedotova & Sidorenko 2005e: 106. Russia: Primorskiï Terr., Vladivostok, Sputnik; HT; ♂; 

ZMAS. 

iridis. UK, Germany, Latvia, Lithuania, Czech Republic, Russia (Europe). 

Cecidomyia iridis Kaltenbach 1873: 717. Germany; ST; LT; NHMW?. 

Iridomyza kaltenbachii Rübsaamen 1899a: 67. Germany; ST; ♂,♀,P,L; ZMHB. 

Dicerura kaltenbachi Kieffer 1913i: 268, missp. of kaltenbachii. 

loba. Canada (Ontario). 

Dicerura loba Borkent 1990: 580. Canada: Ontario, 3 mi. E Carp; HT; ♂; CNCI. 

mixta. Finland, Latvia. 

Dicerura mixta Spungis 1987: 24. Latvia: Kauguri; HT; ♂; LASS. 

oregonensis. USA (Oregon). 

Hormosomyia oregonensis Felt 1919b: 220. USA: Oregon, Forest Grove; HT; ♂; NYSM. 

padi. Russia (Far East). 

Dicerura padi Mamaev 1975c: 60. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

rossica. Russia (Europe). 

Ulmomyia rossica Mamaev 1960a: 1521. Russia: Voronezh Prov., Tellerman For.; ST; ♂,♀; HYNM. 

scirpicola. France. 

Dicerura scirpicola Kieffer 1898f: 57. France: Moselle; ST; ♂,♀,L; lost?. 

Dicerura scirpi Kieffer 1899a: 165, error for scirpicola Kieffer. 

separata. Latvia. 

Dicerura separata Spungis 1987: 26. Latvia: Darzini; HT; ♂; LASS. 

stipator. Russia (Far East). 

Dicerura stipator Mamaev 1972b: 113. Russia: Primorskiï Terr., Ussuriyskiï Distr., Suputinskiï Res.; HT; ♂; 

ZMMU. 

triangularis. Latvia, Ukraine. 

Dicerura triangularis Mamaev 1966: 227. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; MAMC. 

unidentata. Latvia. 

Dicerura unidentata Spungis 1987: 20. Latvia: Kauguri; HT; ♂; LASS. 

xylophila. Latvia, Ukraine.

Dicerura xylophila Mamaev 1966: 227. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; ZMMU. 

DIRHIZA 

Dirhiza Loew 1850a: 38, as subg. of Cecidomyia. Type species, Cecidomyia lateritia Loew (mon.). 

Dirhyza Rondani 1860: 288, 293, missp. of Dirhiza. 

Diorhiza Neave 1939b: 101, missp. of Dirhiza. 

Holarctic; 2 spp.; Dirhizini. Ref.: Panelius 1965 (redescr.). 

lateritia. UK, Latvia, Poland, Russia (Europe). 

Cecidomyia lateritia Loew 1950: 38. Poland; ST; ♀; ZMHB?. 

papillata. USA (New Hampshire). 

Porricondyla papillata Felt 1914d: 111. USA: New Hampshire, Mt Washington; HT; ♀; NYSM. 

*DIVELLEPIDOSIS 

Divellepidosis Fedotova & Sidorenko 2007b: 17 (also as Dollerepidosis, p. 36). Type species, kuznetzovi Fedotova & 

Sidorenko (orig. des.). 

*Faucistyla Fedotova & Sidorenko 2007b: 22, as subg. of Divellepidosis, Type species, Divellepidosis fecunda 

Fedotova & Sidorenko (orig. des.). 

*Epitoxepidosis Fedotova & Sidorenko 2007b: 29, as subg. of Divellepidosis, Type species, Divellepidosis lassa 


*Soligenistyla Fedotova & Sidorenko 2007b: 32, as subg. of Divellepidosis, Type species, Divellepidosis honorata 


Palearctic; 16 spp.; Porricondylini. The subgenera of this genus described in Fedotova & Sidorenko 2007b are listed 

above but not employed here. 

*epipeda. Russia (Far East). 

Divellepidosis epipeda Fedotova & Sidorenko 2007b: 35. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


*fecunda. Russia (Far East). 

Divellepidosis fecunda Fedotova & Sidorenko 2007b: 22. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


*fermita. Russia (Far East). 

Stomatocolpodia fermita Fedotova & Sidorenko 2005e: 93 (also as fermitas, p. 93). Russia: Primorskiï Terr., 30 km 

SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

*honorata. Russia (Far East). 

Divellepidosis honorata Fedotova & Sidorenko 2007b: 32. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 


*kuznetzovi. Russia (Far East). 

Divellepidosis kuznetzovi Fedotova & Sidorenko 2007b: 18. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya 

Pad'; HT; ♂; ZMAS. 

*lassa. Russia (Far East). 

Divellepidosis lassa Fedotova & Sidorenko 2007b: 29. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*latebrosa. Russia (Far East). 

Divellepidosis latebrosa Fedotova & Sidorenko 2007b: 26. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


lutescens. Finland, Latvia. 

Porricondyla lutescens Spungis 1981: 53. Latvia: Lautere; HT; ♂; LASS. 

67

68 

*nonsectaria. Russia (Far East). 

Divellepidosis nonsectaria Fedotova & Sidorenko 2007b: 26 (also as consectaria, pp. 15, 28). Russia: Primorskiï 

Terr., Lazovskiï Res., Cordon America; HT; ♂; ZMAS. 

quadridens. Latvia. 

Porricondyla quadridens Spungis 1981: 49. Latvia: Moritsala; HT; ♂; LASS. 

rotundata. Japan (Kyushu). 

Porricondyla rotundata Yukawa 1971: 73. Japan: Kyushu, Fukuoka Pref., Mt Wakasugi; HT; ♂; KUIC. 

separata. Japan (Kyushu). 

Porricondyla separata Yukawa 1971: 72. Japan: Kyushu, Fukuoka City, Ropponmatsu; HT; ♂; KUIC. 

*spumida. Russia (Far East). 

Divellepidosis spumida Fedotova & Sidorenko 2007b: 20. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*superba. Russia (Far East). 

Divellepidosis superba Fedotova & Sidorenko 2007b: 24. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


*tenuisecta. Russia (Far East). 

Porricondyla tenuisecta Fedotova & Sidorenko 2005e: 90. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*verticalis. Russia (Far East). 

Stomatocolpodia verticalis Fedotova 2004i: 592. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*DOLEREPIDOSIS 

Dolerepidosis Fedotova & Sidorenko 2005e: 104. Type species, sola Fedotova & Sidorenko (orig. des.). 


*sola. Russia (Far East). 

Dolerepidosis lepida Fedotova & Sidorenko 2005e: 104. Russia: Primorskiï Terr., 30 km Vladivostok; HT; ♀; 

ZMAS. 

*ELECTROXYLOMYIA 

Electroxylomyia Nel & Prokop 2006: 49. Type species, eocenica Nel & Prokop (orig. des.). 

Eocene fossil from northern France; 1 sp.; Heteropezini. 

* † mirabilis. Latvia, Poland. 

Electroxylomyia eocenica Nel & Prokop 2006: 49. Amber fossil, northern France: Oise, Creil region, Chevrière; HT; 

♂; NMHN. 

EPICOLA 

Epicola Spungis 1991: 201. Type species, mirabilis Spungis (orig. des.). 


mirabilis. Latvia, Poland. 

Epicola mirabilis Spungis 1991: 201. Latvia: Sigulda; HT; ♂; ZMMU. 

*ERAEPIDOSIS

Eraepidosis Fedotova & Sidorenko 2007d: 267. Type species, errora Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Winnertziini. 

*errora. Russia (Far East). 

Eraepidosis errora Fedotova & Sidorenko 2007d: 268. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 

HT; ♀; ZMAS. 

*EUDOKIMYIA 

Eudokimyia Fedotova & Sidorenko 2005e: 123. Type species, lobata Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Asynaptini. 

*lobata. Russia (Far East). 

Eudokimyia lobata Fedotova & Sidorenko 2005e: 125. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


FELTOMYINA 

Feltomyia Alexander 1936: 12, preocc. Kieffer 1913. Type species, polymera Alexander (orig. des.). 

Feltomyina Alexander 1937: 60, new name for Feltomyia Alexander. 

Neotropical; 1 sp.; Asynaptini. 

polymera. Panama. 

Feltomyia polymera Alexander 1936: 14. Panama: Chiriqui, Potrerillos; HT; ♀; USNM. 

FEROVISENDA 

Ferovisenda Mamaev 1972b: 114. Type species, pigrum Mamaev (orig. des.). 

Eastern Palearctic; 1 sp.; Diallactini. 

pigrum. Russia (Far East). 

Ferovisenda pigrum Mamaev 1972b: 114. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. 

FRIRENIA 

Frirenia Kieffer 1894i: 201. Type species, tenella Kieffer (mon.). 

Palearctic and Baltic amber; 2 spp.; Heteropezini. Ref.: Panelius 1965 (redescr.). 

†eocenica. Upper Eocene - Baltic amber. 

Frirenia eocenica Meunier 1904: 43. Baltic amber; ST; ♂; lost. 

tenella. France. 

Frirenia tenella Kieffer 1894c: 206. France: Bitche; ST; ♂,♀,P,L,egg; ZMHB. 

*FERTEPIDOSIS 

Fertepidosis Fedotova & Sidorenko 2007d: 268. Type species, ferocis Fedotova & Sidorenko (orig. des.). 

69


*ferocis. Russia (Far East). 

Fertepidosis ferocis Fedotova & Sidorenko 2007d: 270. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 

HT; ♀ (erroneously given as ♂ but only ♀ described); ZMAS. 

FURCEPIDOSIS 

Furcepidosis Mamaev 1990a: 20. Type species, gamma Mamaev (orig. des.). 


gamma. Russia (Far East). 

Furcepidosis gamma Mamaev 1990a: 20. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

*GLOMEPIDOSIS 

Glomepidosis Fedotova & Sidorenko 2008a: 234. Type species, solida Fedotova & Sidorenko (orig. des.). 


*solida. Russia (Far East). 

Glomepidosis solida Fedotova & Sidorenko 2008a: 234. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*GRATOMYIA 

Gratomyia Fedotova & Perkovsky 2008b: 61. Type species, inexigentis Fedotova & Perkovsky (orig. des.). 

Rovno amber (Ukraine; late Eocene); 1 sp.; Holoneurini. 

* † inexigentis. Late Eocene - Ukrainian amber. 

Gratomyia inexigentis Fedotova & Perkovsky 2008b: 63. Rovno amber, Ukraine; HT; ♀; UASK. 

GRISEPIDOSIS 

Grisepidosis Mamaev 1968a: 612. Type species, bilobata Mamaev (orig. des.). 

Palearctic; 2 spp.; Solntseviini. 

bilobata. Russia (Far East). 

Grisepidosis bilobata Mamaev 1968a: 612. Russia: Primorskiï Terr., Ussuriyskiï Distr., Suputinskiï Res.; HT; ♂; 

HYNM. 

transmarina. Russia (Far East). 

Grisepidosis transmarina Mamaev & Zaitzev 1998e: 87. Russia: Kuril Is, Kunashir I.; HT; ♂; ZMMU. 

GYNAPTEROMYIA 

Gynapteromyia Mamaev 1965a: 266. Type species, carpatica Mamaev (orig. des.). 

Palearctic; 1 sp.; Diallactini. 

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carpatica. Ukraine. 

Gynapteromyia carpatica Mamaev 1965a: 266. Ukraine: Zakarpatskaya Prov., Kvasy; HT; ♂; ZMMU. 

HAPLUSIA 

Haplusia Karsch 1877: 16. Type species, plumipes Karsch (orig. des.). 

Chastomera Skuse 1888: 112. Type species, bella Skuse (mon.). 

Palaeocolpodia Meunier 1902: 101, nomen nudum. 

Palaeocolpodia Meunier 1904: 29. Type species, eocenica Meunier (mon.). 

Johnsonomyia Felt 1908b: 417. Type species, rubra Felt (orig. des.). 

Paleocolpodia Felt 1911c: 40, missp. of Palaeocolpodia. 

Cosmopolitan and Baltic amber; 19 spp.; Diallactini. Ref.: Gagné 1978d (review). 

alexanderi. Cameroon. 

Johnsonomyia alexanderi Felt 1921d: 96. Cameroon: Efulan; HT; ♀; NYSM. 

bella. Australia (New South Wales). 

Cecidomyia bella Skuse 1888: 112. Australia: New South Wales, Gosford; HT; ♀; ANIC. 

braziliensis. Brazil (Pará). 

Johnsonomyia braziliensis Felt 1915h: 153. Brazil: Pará, Igarape Açú; HT; ♀; NYSM. 

brevipalpis. Russia. 

Chastomera brevipalpis Mamaev 1964c: 903. Russia: Yaroslavl Prov., Volkovo-Rassokhino; HT; ♀; ZMMU. 

cincta. Guatemala. 

Johnsonomyia cincta Felt 1912f: 103. Guatemala: Polochic R.; HT; ♀; USNM. 

† eocenica. Upper Eocene - Baltic amber. 

Palaeocolpodia eocenica Meunier 1904: 29. Baltic amber; ST; ♂,♀; lost. 

funebris. USA (Pennsylvania). 

Haplusia funebris Plakidas 2007: 81. USA: Pennsylvania, Allegheny Co., Aspinwall; HT; ♂; USNM. 

fusca. USA (New York). 

Johnsonomyia fusca Felt 1908b: 417. USA: New York, Albany; HT; ♂; NYSM. 

heteroptera. Latvia. 

Chastomera heteroptera Mamaev & Spungis 1980: 149. Latvia: Brotzeni; HT; ♂; MAMC. 

hondrui. Romania. 

Chastomera hondrui Mamaev 1964c: 902. Romania: nr Bucharest; HT; ♂; ZMMU. 

indica India (Uttar Pradesh). 

Chastomera indica Grover 1971: 4. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

longipalpis. Russia (Europe). 

Chastomera longipalpis Mamaev 1964c: 902. Russia: Voronezh Prov., Volkovo-Rassokhino; HT; ♀; ZMMU. 

obscuripes. Russia (Far East). 

Johnsonomyia obscuripes Mamaev 1968a: 612. Russia: Primorskiï Terr., Ussuriyskiï Distr., Suputinskiï Res.; HT; ♂; 

ZMMU. 

pallida. Russia (Far East). 

Johnsonomyia pallida Mamaev 1966: 220. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. 

palpata. Poland, Russia (Europe). 

Johnsonomyia palpata Mamaev 1966: 219. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; IEME. 

plumipes. Brazil (Bahia). 

Haplusia plumipes Karsch 1877: 16. Brazil: Bahia; HT; ♀; ZMHB. 

rubra. USA (Vermont, New Hampshire, Maryland). 

Johnsonomyia rubra Felt 1908b: 417. USA: Vermont, Montpelier; HT; ♂; NYSM. 

spiculosa. Malaysia. 

Chastomera spiculosa Barnes 1927e: 271. Malaysia: Pahang, Cameron Highlands; HT; ♂; BMNH. 

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72 

*stricta. Russia (Far East). 

Haplusia stricta Fedotova & Sidorenko 2005e: 120. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

HARPOMYIA 

Harpomyia Felt 1916g: 401. Type species, indica Felt (orig. des.). 

Oriental; 1 sp.; unplaced to tribe. 

indica. India (Bihar). 

Harpomyia indica Felt 1916g: 401. India: Bihar, Pusa; ST; ♂,♀,P,L,egg; NYSM. 

HENRIA 

Henria Wyatt 1959: 175. Type species, psalliotae Wyatt (orig. des.). 

Barnesina Pritchard 1960a: 314. Type species, hylecoites Pritchard (orig. des.). 

Holarctic; 3 spp.; Heteropezini. Ref.: Panelius 1965 (redescr.). 

hylecoites. USA (California). 

Barnesina hylecoites Pritchard 1960a: 315. USA: California, Berkeley; HT; ♂; USNM. 

psalliotae. UK. 

Henria psalliotae Wyatt 1959: 175. UK: England, Cornwall, Padstow; HT; ♀; BMNH. 

quercivora. USA (Minnesota, New York). 

Leptosyna querci Felt 1911l: 467, nomen nudum. 

Leptosyna quercus Felt 1911n: 546, jun. prim. hom. quercus Kieffer 1904. USA: New York, Nassau; HT; ♂; NYSM. 

Leptosyna quercivora Felt 1912a: 123, new name for Leptosyna quercus Felt. 

Leptosyna quercicola Kieffer 1913a: 235, new name for Leptosyna quercus Felt. 

Leptosyna americana Felt 1913c: 216. USA: New York, Nassau; HT; ♂; NYSM. 

HETEROPEZA 

Heteropeza Winnertz 1846: 13. Type species, pygmaea Winnertz (mon.). 

Oligarces Meinert 1865: 237. Type species, paradoxus Meinert (mon.) = pygmaea Winnertz. 

Oligarices Shinji 1944: 210, missp. of Oligarces. 

Holarctic and Baltic amber; 3 spp.; Heteropezini. Ref.: Panelius 1965 (redescr.). 

† pulchella. Upper Eocene - Baltic amber. 

Heteropeza pulchella Meunier 1904: 43. Baltic amber; T; ♀; lost. 

pygmaea. Widespread Europe, Israel, USA (California, New York, Pennsylvania, New Jersey, Maryland). 

Heteropeza pygmaea Winnertz 1846: 14. Germany; ST; ♂,♀; lost. 

Oligarces paradoxus Meinert 1865: 238. Denmark; ST; ♀,L; ZMUC. 

Oligarces noveboracensis Felt 1907d: 5. USA: New York, Albany; HT; ♀; NYSM. 

ulmi. USA (New York). 

Oligarces ulmi Felt 1911m: 477. USA: New York, Nassau; ST; ♂; NYSM. 

HETEROPEZINA 

Heteropezina Pritchard 1960a: 311. Type species, cathistes Pritchard (orig. des.).

Holarctic, Oriental, Ukrianian amber; 3 spp.; Heteropezini. 

bharatika. India (Uttar Pradesh). 

Heteropezina bharatika Kashyap & Grover 1986: 30. India: Uttar Pradesh, Allahabad, Ashok Nagar; HT; ♂; PGCA. 

cathistes. USA (Florida), Latvia, Russia (Europe). 

Heteropezina cathistes Pritchard 1960a: 311. USA: Florida, Tampa; HT; ♀; USNM. 

* † marikovskii. Late Eocene - Ukrainian amber. 

Heteropeza (Heteropezina) marikovskii Fedotova & Perkovsky 2008c: 410. Rovno amber, Ukraine, Klesov; HT; ♂; 

UASK. 

HETEROPEZULA 

Heteropezula Wyatt 1967: 85. Type species, tenuis Wyatt (orig. des.). 

Palearctic; 1 sp.; Heteropezini. 

tenuis. UK. 

Heteropezula tenuis Wyatt 1967: 85. UK: England, Sussex, Slindon; HT; ♀; BMNH. 

HILVERSIDEA 

Hilversidia Mamaev 1966: 227. Type species, autumnalis Mamaev (orig. des.). 


autumnalis. Latvia, Russia (Europe). 

Hilversidea autumnalis Mamaev 1966: 228. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

HOLONEURUS 

Holoneura Kieffer 1894c: 316, preocc. Tetens 1891. Type species, cincta Kieffer (orig. des.). 

Holoneurus Kieffer 1895g: 115, new name for Holoneura Kieffer. 

Palearctic, Neotropical, Oriental, New Zealand; 13 spp.; Holoneurini. Refs: Panelius 1965 (rev., Eur. spp.), Parnell 

1971 (review, Nearctic spp.), Mamaev 2001c (review). 

altifilus. USA (New York). 

Porricondyla altifila Felt 1907a: 51. USA: New York, Karner; HT; ♀; NYSM. 

ciliatus. France, Italy. 

Holoneurus ciliatus Kieffer 1896g: 15. France: Bitche; ST; ♂,♀; lost?. 

cinctus. France, Romania. 

Holoneura cincta Kieffer 1894c: 316. France: Moselle; ST; ♂,♀; lost?. 

defectus. Poland. 

Cecidomyia defecta Loew 1850a: 38. Poland; ST; ♂,♀; ZMHB?. 

Colomyia winnertzi Kieffer 1894c: 317. Germany; ST; ♂,♀; lost?. 

fulvus. France, Russia (Europe). 

Holoneurus fulvus Kieffer 1896g: 18. France: Bitche; ST; ♂,♀,P,L; lost?. 

humilis. USA (New York). 

Johnsonomyia humilis Felt 1908b: 417. USA: New York, Albany; HT; ♀; NYSM. 

occidentalis. Panama. 

Holoneurus occidentalis Felt 1911k: 190. Panama: Canal Zone, Paraíso; ST; ♂,♀,L; USNM. 

olschwangi. Russia (Western Siberia). 

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74 

Holoneurus olschwangi Mamaev 1990b: 144. Russia: Yamal Peninsula, Khadyta; HT; ♂; ZMMU. 

paneliusi. UK, Netherlands, Germany, Finland, Russia (Europe, Far East), Japan (Kyushu). 

Holoneurus paneliusi Yukawa 1971: 64. Japan: Kyushu, Fukuoka Pref., Mt Inunaki; HT; ♂; KUIC. 

pectinatus. India (Uttar Pradesh). 

Holoneurus pectinatus Jaiswal 1989a: 30. India: Uttar Pradesh, Garhwal Hills, Kotdwara; HT; ♂; CSIA. 

*recusatus. Russia (Far East). 

Holoneurus recusatus Fedotova & Sidorenko 2005e: 106. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


setiger. France. 

Holoneurus setiger Kieffer 1896g: 17. France: Bitche; ST; ♂,♀,L; lost?. 

tarsalis. USA (New York). 

Holoneurus tarsalis Felt 1915j: 192. USA: New York, North Creek; HT; ♂; NYSM. 

*INCREMENTISTYLA 

Incrementistyla Fedotova & Sidorenko 2007c: 76. Type species, militis Fedotova & Sidorenko (orig. des.). 


*militis. Russia (Far East). 

Incrementistyla militis Fedotova & Sidorenko 2007c: 76. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


ISOCOLPODIA 

Isocolpodia Parnell 1971: 318. Type species, Porricondyla graminis Felt (orig. des.). 

Nearctic; 3 spp.; Porricondylini. 

alta. USA (New York). 

Colpodia alta Felt 1908b: 416. USA: New York, Karner; HT; ♀; NYSM. 

diervillae. USA (New York). 

Porricondyla diervillae Felt 1907a: 49. USA: New York, Karner; HT; ♂; NYSM. 

graminis. USA (New York, Massachusetts, District of Columbia). 

Porricondyla graminis Felt 1907a: 50. USA: New York, Albany; HT; ♀; NYSM. 

Porricondyla trifolii Felt 1907a: 49. USA: New York, Albany; HT; ♂; NYSM. 

Colpodia pratensis Felt 1908b: 416. USA: New York, Nassau; HT; ♀; NYSM. 

Colpodia sanguinea Felt 1908b: 416. USA: New York, Syracuse; HT; ♀; NYSM. 

Colpodia terrena Felt 1908b: 416. USA: New York, Albany; HT; ♀; NYSM. 

Colpodia cornuta Felt 1913f: 135, nomen nudum. 

Colpodia cornuta Felt 1915j: 153. USA: Massachusetts, Boston, Arnold Arboretum; HT; ♂; NYSM. 

JAMALEPIDOSIS 

Jamalepidosis Mamaev 1990b: 143. Type species, longipes Mamaev (orig. des.). 

Palearctic; 3 spp.; unplaced to tribe. 

abludentis. Russia (Far East). 

Jamalepidosis abludentis Mamaev 1998d: 4. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

alienigena. Russia (Far East). 

Jamalepidosis alienigena Mamaev & Zaitzev 1997b: 211. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU.

longipes. Russia (Western Siberia). 

Jamalepidosis longipes Mamaev 1990b: 143. Russia: Yamal Peninsula, Khadyta; HT; ♂; ZMMU. 

KRONOMYIA 

Kronomyia Felt 1911m: 476. Type species, populi Felt (orig. des.). 

Trichoxylomyia Mamaev 1964c: 897. Type species, ovalis Mamaev (orig. des.). 

Klonomyia Shinji 1944: 211, missp. of Kronomyia. 

Holarctic; 6 spp.; Winnertziini. Ref.: Spungis 1992 (redescr.). 

*conspissata. Russia (Far East). 

Kronomyia conspissata Fedotova & Sidorenko 2005e: 112. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*incisa. Russia (Far East). 

Kronomyia incisa Fedotova & Sidorenko 2005e: 109. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


melanura. Russia (Far East). 

Kronomyia melanura Mamaev 1966: 224. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. 

ovalis. Latvia, Ukraine, Russia (Europe). 

Trichoxylomyia ovalis Mamaev 1964c: 897. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

Trichoxylomyia variegata Mamaev 1965a: 148. Ukraine: Zakarpatskaya Prov., Khust; HT; LT; HYNM. 

pilosa. Russia (Europe). 

Trichoxylomyia pilosa Mamaev 1964c: 898. Russia: Voronezh Prov., Tellerman For.; HT; ♀; ZMMU. 

populi. USA (New York, Maryland). 

Kronomyia populi Felt 1911m: 476. USA: New York, Nassau; HT; ♀; NYSM. 

LAMELLEPIDOSIS 

Lamellepidosis Mamaev 1990a: 16. Type species, spungisi Mamaev (orig. des.). 


spungisi. Ukraine. 

Lamellepidosis spungisi Mamaev 1990a: 16. Ukraine: Crimea; HT; ♂; ZMMU. 

*LARIMYIA 

Larimyia Fedotova & Sidorenko 2007c: 88. Type species, larvalis Fedotova & Sidorenko (orig. des.). 


*larvalis. Russia (Far East). 

Larimyia larvalis Fedotova & Sidorenko 2007c: 90. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; HT; 

♂; ZMAS. 

*LEDOMEPIDOSIS 

Ledomepidosis Fedotova & Sidorenko 2007c: 70. Type species, nigella Fedotova & Sidorenko (orig. des.). 


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76 

*nigella. Russia (Far East). 

Ledomepidosis nigella Fedotova & Sidorenko 2007c: 72. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya 


LEPTOSYNA 

Leptosyna Kieffer 1894i: 201. Type species, acutipennis Kieffer (mon.). 

Palearctic; 4 spp.; Heteropezini. Ref.: Wyatt 1967 (redescr.). 

acutipennis. France. 

Leptosyna acutipennis Kieffer 1894i: 209. France: Bitche; ST; E,♂,♀; ZMHB. 

edwardsi. Russia (Europe). 

Leptosyna edwardsi Mamaev 1964b: 91. Russia: Moscow Prov., Danki; HT; ♂; IEME. 

nervosa. UK, Germany. 

Heteropeza nervosa Winnertz 1852: 50. Germany; T; ♀; lost. 

Leptosyna setipennis Edwards 1919: 104. UK: England, Letchworth; ST; ♂,♀; BMNH. 

quercus. France. 

Leptosyna quercus Kieffer 1904c: 407. France: vic. Bitche; ST; ♂,♀,L; lost?. 

*LOBOPEDOSIS 

Lobopedosis Fedotova & Sidorenko 2005e: 115. Type species, maritima Fedotova & Sidorenko (orig. des.). 


*maritima. Russia (Far East). 

Lobopedosis maritima Fedotova & Sidorenko 2005e: 118. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


LOBOWINNERTZIA 

Lobowinnertzia Mamaev 1963b: 565, as subg. of Winnertzia. Type species, Winnertzia globifera Mamaev (orig. 

des.). 

Palearctic; 8 spp.; Winnertziini. Ref.: Fedotova & Sidorenko 2007c: 90 (revision). 

*aperta. Russia (Far East). 

Lobowinnertzia aperta Fedotova & Sidorenko 2007c: 92. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


globifera. Germany, Finland, Estonia, Latvia, Poland, Ukraine, Russia (Europe), Turkmenistan. 

Winnertzia globifera Mamaev 1963b: 566. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

Winnertzia xylostei Mamaev 1963b: 566. Russia: Tula Prov., Krapiven For.; HT; ♂; ZMMU. 

Winnertzia argentata Mamaev 1964c: 897. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ZMMU. 

*nota. Russia (Far East). 

Lobowinnertzia nota Fedotova & Sidorenko 2007c: 93. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


*parma. Russia (Far East). 

Lobowinnertzia parma Fedotova & Sidorenko 2007c: 95. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


prolongata. Turkmenistan. 

Winnertzia prolongata Mamaev 2001d: 3. Turkmenistan: Kushka, Badkhyz Res.; HT; ♂; ZMMU.

tamariciphila. Uzbekistan. 

Winnertzia tamariciphila Mamaev 1963b: 567. Uzbekistan: Bukhara Prov., Shafrikan; HT; ♂; HYNM. 

tridens. UK, Estonia, Latvia, Lithuania, Poland, Ukraine, Moldova, Russia (Europe). 

Winnertzia tridens Panelius 1965: 118. UK: England, Herts, Letchworth; HT; ♂; MZHF. 

tumida. UK, Germany, Latvia. 

Winnertzia tumida Panelius 1965: 121. UK: England, Herts, Letchworth; HT; ♂; BMNH. 

*LOCUSEPIDOSIS 

Locusepidosis Fedotova & Sidorenko 2007c: 86. Type species, summa Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Bryocryptini. 

*summa. Russia (Far East). 

Locusepidosis summa Fedotova & Sidorenko 2007c: 88. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


*MELICEPIDOSIS 

Melicepidosis Fedotova & Sidorenko 2007d: 256. Type species, perplexa Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Holoneurini. 

*perplexa. Russia (Far East). 

Melicepidosis perplexa Fedotova & Sidorenko 2007d: 256 (also as perplexas, p. 257). Primorskiï Terr., Lazovskiï 

Res., Koreiskaya Pad'; HT; ♂; ZMAS. 

MEUNIERIA 

Meunieria Kieffer 1904c: 408. Type species, succini Felt (subs. des. Felt 1911: 36). 

Baltic amber; 1 sp.; unplaced to tribe. 

†succini. Upper Eocene - Baltic amber. 

Meunieria succini Felt 1911c: 36. Baltic amber; T; A; lost. 

MIASTOR 

Miastor Meinert 1864: 42. Type species, metraloas Meinert (mon.). 

Peromiastor Kieffer 1913g: 54. Type species, dryobius Kieffer (orig. des. as n. g., n. sp.) = metraloas Meinert. 

Miaster Shinji 1944: 210, missp. of Miastor. 

Promiaster Shinji 1944: 210. 60, missp. of Peromiastor. 

Holarctic and Ukraine amber; 3 spp.; Heteropezini. Ref.: Wyatt 1967 (redescr.). 

castaneae. UK. 

Miastor castaneae Wyatt 1967: 76. UK: England, Sussex, Littlehampton; HT; ♂; BMNH. 

metraloas. Widespread Palearctic, USA (Minnesota, Indiana. New York, Connecticut, New Jersey, Maryland). 

Miastor metraloas Meinert 1864: 42. Denmark: Fredericksdal; ST; ♂,♀,P,L; ZMUC. 

Miastor hospes Winnertz 1870a: 6. Germany; T; A; lost. 

Miastor subterraneus Karsch 1887: 199. Germany: Saxony, Burgk; ST; ♀; ZMHB?. 

Miastor hastatus Kieffer 1898f: 54, nomen nudum. 

77

Miastor americana Felt 1907d: 5. USA: New York, Highland; HT; ♀; NYSM. 

Peromiastor dryobius Kieffer 1913g: 54. France: Bitche; T; A; lost?. 

Miastor hastatus Kieffer 1913g: 53. France: Bitche; ST; ♂,♀; lost?. 

Miastor similis Kieffer 1913g: 54. France; T; A; lost?. 

MISOCOSMUS 

Misocosmus Kieffer 1913g: 55. Type species, Epidosis ceylanica Kieffer (orig. des.). 

Mysocosmus Felt 1918c: 301, missp. of Misocosmus. 

Oriental; 1 sp.; unplaced to tribe. 

ceylanicus. Sri Lanka. 

Epidosis ceylanica Kieffer 1912a: 29. Sri Lanka: Peradeniya; HT; ♀; NZSI?. 

MONEPIDOSIS 

Monepidosis Mamaev 1966: 235. Type species, pectinata Mamaev (orig. des.). Ref.: Spungis 2006b (rev.). 

Holarctic; 8 spp.; Porricondylini. 

*acuta. Japan. 

Monepidosis acuta Spungis 2006b: 27. Japan: Hyogo, Sanda, Fukada-Koen; HT; ♂; HYNM. 

carolinae. USA (North Carolina), northern Europe, Japan (Kyushu). 

Porricondyla carolinae Felt 1907a: 49 USA: North Carolina, Davidson's R.; HT; ♂; NYSM. 

Monepidosis furcata Mamaev 1966: 236. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

Monepidosis pectinata Mamaev 1966: 235. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; IEME. 

Monepidosis bulgarica Mamaev & Dimitrova 1992: 174. Bulgaria: Vitosha Mtn, nr Sofia; HT; ♂; BIZS. 

*digitata. Japan. 

Monepidosis digitata Spungis 2006b: 29. Japan: Hyogo, Sanda, Fukada-Koen; HT; ♂; HYNM. 

duplicis. Latvia, Ukraine, Russia (Far East). 

Monepidosis duplicis Mamaev 1998d: 5. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; MAMC. 

*laccata. Russia (Far East). 

Monepidosis laccata Fedotova & Sidorenko 2008a: 228. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya 


*munita. Russia (Far East). 

Monepidosis munita Fedotova & Sidorenko 2005e: 94. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


*scepteri. Latvia. 

Monepidosis scepteri Spungis 2006b: 25. Latvia: Bramberge; HT; ♂; FBUL. 

*spatulata. Latvia. 

Monepidosis spatulata Spungis 2006b: 27. Latvia: Bramberge; HT; ♂; FBUL. 

MONOCOLPODIA 

Monocolpodia Mamaev 1990a: 17. Type species, Neocolpodia spiniformis Mamaev (orig. des.). 


spiniformis. Russia (Europe). 

Colpodia spiniformis Mamaev 1964c: 904. Russia: Yaroslavl Prov., Volkovo-Rassokhino; HT; ♂; IEME. 

78

MONODICRANA 

Monodicrana Loew 1850b: 32. Type species, terminalis Loew (mon.). 

Monodicrania Schiner 1868: 5, missp. of Monodicrana. 

Monodricana Bigot 1854: 464, missp. of Monodicrana. 

Baltic amber; 1 sp.; Heteropezini. 

Hterminalis. Upper Eocene - Baltic amber. 

Monodicrana terminalis Loew 1850b: 32. Fossil; Baltic amber; T; A; lost?. 

*NEBULEPIDOSIS 

Nebulepidosis Fedotova & Sidorenko 2008b: 376. Type species, urceolata Fedotova & Sidorenko (orig. des). 

Palearctic; 2 spp.; Dicerurini. 

*urceolata. Russia (Far East). 

Nebulepidosis urceolata Fedotova & Sidorenko 2008b: 377. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 


*velutina. Russia (Far East). 

Nebulepidosis velutina Fedotova & Sidorenko 2008b: 380. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


NEOCOLPODIA 

Neocolpodia Mamaev 1964c: 904, as subg. of Colpodia. Type species, Colpodia paradoxa Mamaev (orig. des.). 


paradoxa. Netherlands, Germany, Russia (Europe). 

Colpodia paradoxa Mamaev 1964c: 904. Russia: Moscow Prov., Abramtsevo; HT; ♂; HYNM. 

pinea. USA (North Carolina). 

Porricondyla pinea Felt 1907a: 48. USA: North Carolina, Davidson's R.; HT; ♂; NYSM. 

NEOSTENOPTERA 

Stenoptera Meunier 1901: 201, preocc. Duponchel 1838. Type species, kiefferi Meunier (mon.). 

Neostenoptera Meunier 1902: 102, new name for Stenoptera Meunier. 

Afrotropical and African copal; 2 spp.; Heteropezini. 

†congoensis. Congo. 

Neostenoptera congoensis Gagné 1979: 347. Congo: Mayombe, Stn Dimonika; HT; ♀; MNHN. 

† kiefferi. African copal. 

Stenoptera kiefferi Meunier 1901: 201. African copal; HT; ♂; lost. 

NEOTETRANEUROMYIA 

Neotetraneuromyia Spungis 1987: 36. Type species, lenticularis Spungis (orig. des). 


79

lenticularis. Latvia. 

Neotetraneuromyia lenticularis Spungis 1987: 37. Latvia: Yumprava; HT; ♂; LASS. 

moldaviensis. Moldova. 

Neotetraneuromyia moldaviensis Spungis 1987: 37. Moldova: Durleshty; HT; ♂; LASS. 

NEUREPIDOSIS 

Neurepidosis Spungis 1987: 39. Type species, gracilis Spungis (orig. des). 

*Lanepidosis Fedotova & Sidorenko 2008b: 367, as subgenus of Neurepidosis. Type species, Neurepidosis conchata 

Fedotova & Sidorenko (orig. des). 


*conchata. Russia (Far East). 

Neurepidosis (Lanepidosis) conchata Fedotova & Sidorenko 2008b: 367. Russia: Primorskiï Terr., Lazovskiï Res.; 


gracilis. Latvia. 

Neurepidosis gracilis Spungis 1987: 39. Latvia: Salaspils; HT; ♂; LASS. 

minuta. Latvia. 

Neurepidosis minutus Spungis 1987: 40. Latvia: Darzini; HT; ♂; LASS. 

*paradisica. Russia (Far East). 

Neurepidosis (Lanepidosis) paradisica Fedotova & Sidorenko 2008b: 370. Russia: Primorskiï Terr., Lazovskiï Res., 

Koreiskaya Pad'; HT; ♂; ZMAS. 

solinasi. Italy. 

Neurepidosis solinasi Mamaev & Zaitzev 1998b: 129. Italy: Pontignano; HT; ♂; ZMMU. 

*suffusa. Russia (Far East). 

Neurepidosis (Lanepidosis) suffusa Fedotova & Sidorenko 2008b: 369. Russia: Primorskiï Terr., Lazovskiï Res., 

Cordon America; HT; ♂; ZMAS. 

NIKANDRIA 

Nikandria Mamaev 1964b: 93. Type species, brevitarsis Mamaev (orig. des.). 

Holarctic; 1 sp.; Heteropezini. Ref.: Panelius 1965 (redescr.). 

brevitarsis. Russia (Europe). 

Nikandria brevitarsis Mamaev 1964b: 93. Russia: Voronezh Prov., Tellerman For.; HT; ♂; IEME. 

*OBRYZEPIDOSIS 

Obryzepidosis Fedotova & Sidorenko 2007b: 38.Type species, declivis Fedotova & Sidorenko (orig. des.). 


*declivis. Russia (Far East). 

Obryzepidosis declivis Fedotova & Sidorenko 2007b: 39. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


PALAEOSPANIOCERA 

Palaeospaniocera Meunier 1901: 192. Type species, Palaeospaniocera paleospaniocera Evenhuis (subs. des. 

80

Evenhuis 1994: 186). 

African copal; 1 sp.; unplaced to tribe. 

†paleospaniocera. African copal. 

Palaeospaniocera paleospaniocera Evenhuis 1994: 186. African copal; T; A; lost. 

PALLIDEPIDOSIS 

Pallidepidosis Mamaev & Zaitzev 1996b: 75. Type species, pilosa Mamaev & Zaitzev (orig. des.). 

Palearctic; 3 spp.; unplaced to tribe. 

*adunca. Russia (Far East). 

Pallidepidosis adunca Fedotova & Sidorenko 2005e: 96. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 



Pallidepidosis pallida Mamaev & Zaitzev 1996b: 76. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

pilosa. Russia (Europe). 

Pallidepidosis pilosa Mamaev & Zaitzev 1996b: 76. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

PARACOLPODIA 

Paracolpodia Parnell 1971: 321. Type species, Colpodia capitata Felt (orig. des.). 


capitata. USA (New York). 

Colpodia capitata Felt 1914c: 125. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

furcata. Russia (Europe). 

Paracolpodia furcata Mamaev 2001b: 7. Russia: Northern Urals, Denezhkin Kamen'; ♂; ZMMU. 

lunularis. Russia (Far East). 

Paracolpodia lunularis Mamaev & Zaitzev 1998e: 89. Russia: Amur Prov., Sega; HT; ♂; ZMMU. 

PARASYNAPTA 

Parasynapta Panelius 1965: 84. Type species, intermedia Panelius (orig. des.). 

Holarctic; 2 spp.; Asynaptini. 

canadensis. Canada (Quebec). 

Porricondyla canadensis Felt 1908b: 418. Canada: Quebec; HT; ♂; MCZ?. 

intermedia. Finland. 

Parasynapta intermedia Panelius 1965: 84. Finland: Källvik, Ekenäs Lk; HT; ♂; MZHF. 

PARATETRANEUROMYIA 

Paratetraneuromyia Spungis 1987: 29. Type species, Asynapta nobilis Felt (orig. des.). 

Holarctic; 2 spp.; Dicerurini. 

nobilis. USA (New York, Maine), UK, Latvia. 

81

Asynapta nobilis Felt 1913f: 142. USA: Maine, Bridgeton; HT; ♀; NYSM. 

Asynapta apicalis Felt 1914c: 127. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

Asynapta mediana Felt 1914c: 128. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

Clinorhytis ramosa Panelius 1965: 28. UK: England: Gloucester, Symond's Yat; HT; ♂; BMNH. 

vernalis. Finland, Latvia. 

Paratetraneuromyia vernalis Spungis 1987: 31. Latvia: Kauguri; HT; ♂; LASS. 

PAREPIDOSIS 

Parepidosis Kieffer 1913a: 234. Type species, Cecidomyia venusta Winnertz sensu Kieffer (orig. des.) = venustior 

Gagné. 

Parpidosise Mani 1946: 198, missp. of Parepidosis. 

Holarctic, Oriental; 14 spp.; Porricondylini. Ref.: Panelius 1965 (rev., Eur. spp.). 

arcuata. Latvia, Poland, Russia (Europe), Uzbekistan. 

Parepidosis arcuata Mamaev 1964c: 912. Russia: Vologda Prov., Kadnikovskaya; HT; ♂; ZMMU. 

argentifera. Netherlands, Germany. 

Porricondyla argentifera Meijere 1906: 24. Netherlands: Hilversum; LT (Yukawa & Nijveldt 1975: 104); ♂; ZMAN. 

longinodis. UK, Finland. 

Parepidosis longinodis Panelius 1965: 53. England: Herts, Harpenden; HT; ♂; MZHF. 

mamaevi. Romania. 

Parepidosis mamaevi Neacsu 1968: 899. Romania: Turnu Severin, Oltenia; HT; ♂; NEAC. 

militaris. Russia (Far East). 

Parepidosis militaris Mamaev 1966: 235. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

peculiaris. Russia (Europe). 

Parepidosis peculiaris Mamaev 1966: 235. Russia: Yaroslavl Prov., Volkovo; HT; ♂; MAMC. 

porrecta USA (New York). 

Colpodia porrecta Felt 1914c: 126. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♀; NYSM. 

rectangularis. India (Jammu and Kashmir). 

Parepidosis rectangularis Kashyap 1993: 113. India: Jammu and Kashmir, Srinagar; HT; ♂; PGCA. 

*strigillata. Russia (Far East). 

Parepidosis strigilata Fedotova 2004i: 594. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

sylvestris. USA (New York). 

Colpodia sylvestris Felt 1914c: 126. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♀; NYSM. 

trilobata. India (Maharashtra). 

Parepidosis trilobata Sharma & Rao 1979b: 125. India: Maharashtra, Aurangabad; HT; ♂; RCMU. 

ulmicorticis. Russia (Europe). 

Parepidosis ulmicorticis Mamaev 1964c: 911. Russia: Voronezh Prov., Tellerman For.; HT; ♂; ZMMU. 

ventralis. Japan (Kyushu). 

Parepidosis ventralis Yukawa 1971: 80. Japan: Kyushu, Kagoshima City, Iso; HT; ♂; KUIC. 

venustior. Widespread Europe. 

Epidosis venusta Winnertz of Kieffer 1901a: pl. 21, fig. 6, pl. 25, fig. 2. 

Parepidosis venustior Gagné 2004b: 53. France; L of Kieffer 1901b: pl. 25, fig. 2; lost. Ref.: Penttinen & Spungis 

2007 (tax'c notes). 

PARWINNERTZIA 

Parwinnertzia Felt 1920a: 281. Type species, notmani Felt (orig. des.). 

Parawinnertzia Curran 1934: 107, missp. of Parwinnertzia. 

82

Holarctic; 2 spp.; Winnertziini. 

italiana. Italy. 

Parwinnertzia italiana Mamaev & Zaitzev 1998e: 89. Italy: Siena; HT; ♂; ZMMU. 

notmani. USA (New York). 

Parwinnertzia notmani Felt 1920a: 281. USA: New York, Keene Valley; HT; ♀; NYSM. 

PORRICONDYLA 

Porricondyla Rondani 1840: 13, as subg. of Cecidomyia. Type species, Cecidomyia albitarsis Meigen (mon.) = 

nigripennis Meigen. 

Epidosis Loew 1850a: 21, 38. Type species, Cecidomyia fragilis Loew (subs. des. Brunetti 1920: 17). 

*Dicraneura Kieffer 1894k: unnumbered p. 2. Type species, Dicraneura albimana Kieffer (mon.). 

Dicroneurus Kieffer 1895g: 122. Type species, Epidosis lineata Kieffer (mon.). 

Phaenepidosis Kieffer 1913i: 273. Type species, Epidosis abbreviata Kieffer (orig. des.). 

Synaptella Kieffer 1913g: 55. Type species, sulfurea Kieffer (orig. des.). 

Synarthrella Kieffer 1913g: 55. Type species, flavida Kieffer (orig. des.). 

Poricondylia Shinji 1942a: 15, missp. of Porricondyla. 

Vesperomyia Marikovskij 1958b: 1846. Type species, montana Marikovskij (orig. des.). 

Cosmopolitan and Baltic amber; 95 spp.; Porricondylini. Refs: Panelius 1965 (rev., Eur. spp.), Parnell 1971 (review, 

Nearctic spp.). 

abbreviata. Kenya. 

Epidosis abbreviata Kieffer 1913i: 114. Kenya: Kijabé RR Stn, in forest of Kikuyu escarpment; HT; ♀; MNHN. 

acuta. Latvia. 

Porricondyla acuta Spungis 1981: 51. Latvia: Vidrizhi; HT; ♂; LASS. 

albiceps. UK. 

Cecidomyia albiceps Walker 1856a: 99. UK: England; ST; ♂; BMNH. 

*albimana. France. 

Dicraneura albimana Kieffer 1894k: unnumbered p. 2. Unstated, presumably France: Bitche; HT; egg; lost? 

*albipalpis. Germany. 

Cecidomyia albipalpis Meigen 1838: 22. Germany; T; ♀; MNHN. 

americana. USA (New York). 

Colpodia americana Felt 1914c: 124. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

amurensis. Russia (Far East). 

Porricondyla amurensis Mamaev & Zaitzev 1996d: 209. Russia: Amur Prov., nr Zeja; HT; ♂; ZMMU. 

anemotis. Kenya. 

Epidosis anemotis Kieffer 1913i: 114. Kenya: Kijabé RR Stn, in forest of Kikuyu escarpment; HT; ♀; MNHN. 

antennata. USA (New York). 

Porricondyla antennata Felt 1915j: 169. USA: New York, Johnstown; HT; ♂; NYSM. 

armata. Russia (Europe). 

Porricondyla armata Spungis 1981: 53. Russia: Voronezh Prov., Ramon'; HT; ♂; LASS. 

asiatica. Uzbekistan. 

Porricondyla asiatica Mamaev 1963c: 76. Uzbekistan: Samarkand, Takhtakaracha Pass; HT; ♂; HYNM. 

aurantiaca. Finland, Germany, Poland, Austria, Russia (Europe), Japan (Honshu, Kyushu). 

Porricondyla aurantiaca Panelius 1965: 52. Finland: Grankulla; HT; ♂; MZHF. 

auriculata. Western Samoa. 

Phaenepidosis auriculata Barnes 1928e: 107. Western Samoa: Upolu: Malololelei; HT; ♀; BMNH. 

barberi. USA (Arizona, New York, Maine). 

Porricondyla barberi Felt 1908b: 418. USA: Arizona, Williams; HT; ♂; USNM. 

Porricondyla tumidosa Felt 1920a: 284. USA: New York, Keene Valley; HT; ♂; NYSM. 

83

84 

Porricondyla sylvestris Felt 1926c: 207. USA: Maine, Bar Harbor; HT; ♂; NYSM. 

bicincta. France. 

Dicroneurus bicinctus Kieffer 1913f: 46. France: Bitche; T; ♀; lost?. 

bifurcata. Uzbekistan. 

Porricondyla bifurcata Mamaev 1963c: 74. Uzbekistan: Takhtakaracha Pass, nr Samarkand; HT; ♂; MAMC. 


Porricondyla borealis Felt 1907a: 51. USA: New York, Lake Clear; HT; ♀; NYSM. 

carolina. USA (North Carolina). 

Porricondyla carolina Felt 1908b: 418. USA: North Carolina, Davidson's R.; HT; ♀; NYSM. 

caucasicola Russia (Europe). 

Porricondyla caucasicola Mamaev & Zaitzev 1996d: 211. Russia: Krasnodarskiï Terr., Sober-Oash Mtn nr 

Ubinskaya; HT; ♂; ZMMU. 

citrina. France. 

Epidosis citrina Kieffer 1894c: 321. France: Bitche; ST; ♂,L; lost?. 

colpodioides. Germany, Russia (Europe), Uzbekistan. 

Porricondyla colpodioides Mamaev 1963c: 75. Uzbekistan: Takhtakaracha Pass, nr Samarkand; HT; ♂; HYNM. 

consobrina. Canada (Ontario). 

Porricondyla consobrina Felt 1919b: 221. Canada: Ontario, Gull L., nr Muskoka; HT; ♂; NYSM. 

dietzii. USA (Pennsylvania). 

Porricondyla dietzii Felt 1912f: 105. USA: Pennsylvania, Hazelton; HT; ♀; NYSM. 

dilatata. USA (New York), UK, Latvia, Lithuania, Poland, Russia (Europe). 

Porricondyla dilatata Felt 1908b: 418. USA: New York, Highland; HT; ♂; NYSM. 

distinguenda. Sweden, Germany, Latvia, Poland, Russia (Europe), Uzbekistan. 

Porricondyla distinguenda Mamaev 1963c: 75. Uzbekistan: Kitab; HT; ♂; HYNM. 

† 

elegantula. Upper Eocene - Baltic amber. 

Dicroneurus elegantulus Meunier 1904: 33. Baltic amber: T; ♀; lost. 

elongata. Canada (British Columbia). 

Holoneurus elongatus Felt 1908b: 420. Canada: British Columbia, Kaslo; HT; ♂; USNM. 

ermakovi. Russia (Europe). 

Porricondyla ermakovi Mamaev 2001b: 5. Russia: Northern Urals, Denezhkin Kamen'; ♂; ZMMU. 

errabunda. Russia (Europe). 

Porricondyla errabunda Mamaev 2001b: 6. Russia: Northern Urals, Denezhkin Kamen'; ♂; ZMMU. 

flava. Germany. 

Cecidomyia flava Meigen 1818: 99. Germany; T; ♂; MNHN. 

flavescens. Austria. 

Epidosis flavescens Löw 1874a: 147. Austria: Vienna, Wienerwald; ST; ♀; NHMW. 

flavida. France. 

Synarthrella flavida Kieffer 1913g: 55. France: Bitche; T; A; lost?. 

fragilis. Poland. 

Cecidomyia fragilis Loew 1850a: 38. Poland; ST; ♂,♀; ZMHB?. 

fulvescens. UK, Finland. 

Porricondyla fulvescens Panelius 1965: 45. Finland: Tvärminne; HT; ♂; MZHF. 

fuscostriata. Finland, Latvia. 

Porricondyla fuscostriata Panelius 1965: 50. Finland: Tvärminne; HT; ♂; MZHF. 

† 

gibbosa. Upper Eocene - Baltic amber. 

Epidosis gibbosa Meunier 1904: 33. Baltic amber; ST; ♂,♀; lost. 

globosa. Finland, Latvia. 

Porricondyla globosa Spungis 1981: 49. Latvia: Darzini; HT; ♂; LASS. 

gracilis. Germany. 

Cecidomyia gracilis Winnertz 1853: 298. Germany: S; ♂,♀; lost. 

hamata. USA (New York, New Hampshire). 

Dirhiza hamata Felt 1907a: 48. USA: New York, Albany; HT; ♀; NYSM.

85 

Porricondyla novaeangliae (as novae-angliae) Felt 1914d: 110. USA: New Hampshire, Franconia; HT; ♀; NYSM. 

hamifera. Uzbekistan. 

Porricondyla hamifera Mamaev 1963c: 76. Uzbekistan: Takhtakaracha Pass, nr Samarkand; HT; ♂; MAMC. 

hyalinata. Russia (Western Siberia). 

Porricondyla hyalinata Mamaev 1990b: 144. Russia: Yamal Peninsula, Khadyta; HT; ♂; ZMMU. 

hypoxantha. Netherlands, Germany, Finland, Ukraine, Russia (Europe). 

Porricondyla hypoxantha Panelius 1965: 49. Finland: Tvärminne; HT; ♂; MZHF. 

indica. India. 

Epidosis indica Kieffer 1913c: 200. India: Himalayas; T; ♂; lost?. 

juvenalis. USA (New York). 

Porricondyla juvenalis Felt 1912l: 239. USA: New York, Adirondack Mts, Elm L.; HT; ♂; NYSM. 

Colpodia ovata Felt 1914c: 125. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

lamellata. Japan (Kyushu). 

Porricondyla lamellata Yukawa 1971: 71. Japan: Kyushu, Kumamoto Pref., Mt Ichifusa; HT; ♂; KUIC. 

lata. Lithuania, Poland, Russia (Europe). 

Porricondyla lata Mamaev 1965a: 267. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; MAMC. 

leacheana. UK, Austria, Russia (Europe). 

Cecidomyia leacheana Stephens 1829a: 53 (publ'n not seen); also in Stephens 1929b: 240. Nomen nudum. 

Cecidomyia leacheana Walker 1856a: 109. UK: England, nr London & Gloucestershire, Forest of Dean; ST; ♂,♀; 

BMNH. 

lineata. France. 

Epidosis lineata Kieffer 1894c: 320. France: Bitche; ST; ♀; lost?. 

lobata. France. 

Synaptella lobata Kieffer 1913f: 45. France: Bitche; ST; ♂,♀; lost?. 

lobifera. Russia (Far East). 

Porricondyla lobifera Mamaev & Zaitzev 1996d: 211. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

longipes. Poland. 

Cecidomyia longipes Loew 1850a: 38. Poland; ST; ♀; ZMHB?. 

Hmagnifica. Upper Eocene - Baltic amber. 

Epidosis magnifica Meunier 1904: 32. Baltic amber; T; ♂; lost. 

microcera. France. 

Epidosis microcera Kieffer 1901a: 35. France: Bitche; T; ♀; lost?. 

minor. Canada (British Columbia). 

Porricondyla minor Felt 1926f: 268. Canada: British Columbia, Cranbrook; HT; ♀; NYSM. 

Hminuta. Upper Eocene - Baltic amber. 

Epidosis minuta Meunier 1904: 34. Baltic amber; T; ♀; lost. 

minutina. India (Uttar Pradesh). 

Porricondyla minuta Chandra 1993b: 2, jun. sec. hom. minuta Meunier. India: Uttar Pradesh, Allahabad; HT; ♂; 

CSIA. 

Porricondyla minutina Gagné 2004b: 55, new name for Porricondyla minuta Chandra. 

modesta. Latvia. 

Porricondyla modesta Spungis 1981: 50. Latvia: Koknese; HT; ♂; LASS. 

montana. Kazakhstan, Kirghizstan. 

Vesperomyia montana Marikovskij 1958b: 1847. Kazakhstan: Alma-Ata, & Kirghizstan: Frunze; ST; ♂; MARC. 

monticola. Kenya. 

Epidosis monticola Kieffer 1913h: 113. Kenya: Naivasha, on L. Naivasha; HT; ♀; MNHN. 

neglecta. Finland, Latvia, Czechoslovakia, Russia (Europe). 

Porricondyla neglecta Mamaev 1965a: 267. Russia: Moscow Prov., Il'icha For.; HT; ♂; HYNM. 

nervosa. Germany. 

Cecidomyia nervosa Meigen 1838: 21. Germany: Bavaria; T; ♂; MNHN. 

nigripennis. Widespread Holarctic. 

Cecidomyia nigripennis Meigen 1830: 269. Germany; T; ♀; MNHN.

86 

Cecidomyia albitarsis Meigen 1830: 268. Germany; T; ♀; lost. 

Cecidomyia analis Winnertz 1853: 300. Germany; ST; ♂; lost. 

Cecidomyia dorsalis Winnertz 1853: 297. Germany; ST; ♀; lost. 

Porricondyla wellsi Felt 1915e: 227. USA: New York, Wells; HT; ♂; NYSM. 

Colpodia colei Felt 1919b: 223. USA: Oregon, Forest Grove; HT; ♂; NYSM. 

*nimia. Russia (Far East). 

Porricondyla nimia Fedotova & Sidorenko 2007c: 83. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


nocturna. Kazakhstan. 

Synaptella nocturna Marikovskij 1958b: 1842. Kazakhstan: Alma-Ata; T; ♂; MARC. 

Hnoduliformis. Upper Eocene - Baltic amber. 

Epidosis noduliformis Meunier 1904: 35. Baltic amber; T; ♀; lost. 

pallescens. Finland. 

Porricondyla pallescens Panelius 1965: 51. Finland: Ekhamn; HT; ♂; MZHF. 

pallida. Germany. 

Cecidomyia pallida Meigen 1830: 270. Germany; T; ♂; MNHN. 

pallipes. Germany. 

Cecidomyia pallipes Winnertz 1853: 295. Germany; ST; ♀; lost. 

pergrata. Turkmenistan. 

Porricondyla pergrata Mamaev 1986: 67. Turkmenistan: Kopetdag, Ipaï-Kala; HT; ♂; HYNM. 

petiolata. Turkmenistan. 

Porricondyla petiolata Mamaev 1986: 67. Turkmenistan: Kushka, Badkhyz Res.; HT; ♂; ZMMU. 

petrophila. Turkmenistan. 

Porricondyla petrophila Mamaev 1986: 67. Turkmenistan: Kopetdag, Ipai-Kala; HT; ♂; HYNM. 

peyerimhoffi. Algeria. 

Synaptella peyerimhoffi Kieffer 1919: 193. Algeria: For. Bouïra-Sahary, betw. Aïn Oussera & Djelfa; T; ♂; lost?. 

photophila. Germany, Latvia. 

Porricondyla photophila Spungis 1981: 51. Latvia: Salaspils; HT; ♂; LASS. 

porrecta. USA (North Carolina). 

Porricondyla porrecta Felt 1912f: 105. USA: North Carolina, Davidson's R.; HT; ♀; NYSM. 

prayagensis. India (Uttar Pradesh). 

Porricondyla prayagensis Grover & Bakhshi 1978: 11. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

pubescens. UK, Finland, Latvia, Macedonia, Ukraine, Russia (Europe). 

Cecidomyia pubescens Walker 1856a: 104. UK: England; ST; ♂; BMNH. 

quercina. USA (New York). 

Porricondyla quercina Felt 1907a: 50. USA: New York, Karner; HT; ♀; NYSM. 

Porricondyla karnerensis Felt 1908b: 418. USA: New York, Karner; HT; ♀; NYSM. 

Porricondyla dorsata Felt 1912l: 238. USA: New York, Adirondack Mts, Dug Mtn; HT; ♀; NYSM. 

ramadei. France. 

Porricondyla ramadei Thomas 1980: 226. France: Hautes-Pyrénées, la Neste de Couplan; HT; ♂; ?. 

*recondita. USA (Pennsylvania). 

Porricondyla recondita Plakidas 2005: 652. USA: Pennsylvania, Allegheny Co., 15 km NE Pittsburgh; HT; ♂; 


rostellata. Finland, Latvia, Lithuania, Poland. 

Porricondyla rostellata Panelius 1965: 49. Finland: Tvärminne; HT; ♂; MZHF. 

rufescens. UK, Netherlands, Finland, Latvia, Lithuania, Yugoslavia, Turkmenistan, Russia (Far East). 

Porricondyla rufescens Panelius 1965: 45. Finland: Tvärminne; HT; ♂; MZHF. 

rufocinerea. Finland. 

Porricondyla rufocinerea Panelius 1965: 47. Finland: Helsingfors; HT; ♂; MZHF. 

sericata. Poland. 

Cecidomyia sericata Loew 1850a: 38. Poland; ST; ♀; ZMHB?. 

setosa. USA (New York).

87 

Porricondyla setosa Felt 1914c: 129. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♀; NYSM. 

spinigera. USA (New York). 

Porricondyla spinigera Felt 1920a: 284. USA: New York, Keene Valley; HT; ♂; NYSM. 

srinagarensis. India (Jammu and Kashmir). 

Porricondyla srinagarensis Kashyap 1993: 116. India: Jammu and Kashmir, Srinagar; HT; ♂; PGCA. 

sulfurea. France. 

Synaptella sulfurea Kieffer 1913g: 55. France: Bitche; T; ♂; lost?. 

sylvestris. USA (New Jersey). 

Porricondyla sylvestris Felt 1907a: 50. USA: New York, Lake Clear; HT; ♀; NYSM. 

tetraschistica. Russia (Europe). 

Porricondyla tetraschistica Mamaev 1998d: 5. Russia: Krasnodarskiï Terr., Ubinskaya; HT; ♂; ZMMU. 

Htitana. Upper Eocene - Baltic amber. 

Epidosis titana Meunier 1904: 35. Baltic amber; T; ♀; lost. 

tuckeri. USA (Kansas, New York). 

Porricondyla tuckeri Felt 1908b: 418. USA: Kansas, Lawrence; HT; ♀; NYSM. 

Porricondyla caudata Felt 1908b: 418. USA: New York, Nassau; HT; ♀; NYSM. 

unidentata. USA (New York, Pennsylvania). 

Porricondyla flava Felt 1907a: 50, jun. sec. hom. Cecidomyia flava Meigen. USA: New York, Lake Clear; HT; ♂; 

NYSM. 

Porricondyla unidentata Parnell 1971: 333, new name for Porricondyla flava Felt 1907. 

ussuriorum. Russia (Far East). 

Porricondyla ussuriorum Mamaev & Zaitzev 1996d: 211. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; MAMC. 

venusta. Germany. 

Cecidomyia venusta Winnertz 1853: 299. Germany; ST; ♂; lost. 

vernalis. USA (Pennsylvania). 

Porricondyla vernalis Felt 1912f: 104. USA: Pennsylvania, Hazelton; HT; ♀; NYSM. 

varians. USA (New York). 

Porricondyla varians Parnell 1971: 331. USA: New York, Keene Valley; HT; ♂; NYSM. 

PROSEPIDOSIS 

Prosepidosis Kieffer 1913a: 233. Type species, pectinata Kieffer (orig. des.). 

Western Holarctic; 1 sp.; Porricondylini. Ref.: Panelius 1965 (redescr.). 

pectinata. France. 

Prosepidosis pectinata Kieffer 1913a: 234. France: Bitche; ST; ♂,♀; lost?. 

PSEUDEPIDOSIS 

Pseudepidosis Mamaev 1966: 233. Type species, lunaris Mamaev (orig. des.). 


armilla. Russia (Far East). 

Pseudepidosis armilla Mamaev 1994: 31. Russia, Kamchatka, Kozyrevsk; HT; ♂; HYNM. 

bifida. Latvia. 

Pseudepidosis bifida Spungis 1981: 46. Latvia: Brotseny; HT; ♂; LASS. 

*dives. Russia (Far East). 

Dendrepedosis dives Fedotova & Sidorenko 2008a: 232. Russia: Primorskiï Terr., Lazovskiï Res., Proselochhnaya 


lunaris. Russia (Europe).

Pseudepidosis lunaris Mamaev 1966: 233. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

trifida. Finland, Poland, Russia (Europe). 

Pseudepidosis trifida Mamaev 1966: 233. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

zonata. Russia (Eastern Siberia). 

Pseudepidosis zonata Mamaev 1990b: 145. Russia: Taimyr, Tareya; HT; ♂; ZMMU. 

PSEUDOCAMPTOMYIA 

Pseudocamptomyia Parnell 1971: 304. Type species, Asynapta phosphila Felt (orig. des.). 

Asycampta Mamaev & Zaitzev 1997c: 11, as subg. of Pseudocamptomyia. Type species, Pseudocamptomyia palpata 

Mamaev & Zaitzev. 

Nearctic, Afrotropical; 3 spp.; Asynaptini. 

africana. Somalia. 

Pseudocamptomyia africana Mamaev & Zaitzev 1997c: 11. Somalia: Hargeisa; HT; ♂; ZMMU. 

palpata. Somalia. 

Pseudocamptomyia palpata Mamaev & Zaitzev 1997c: 11. Somalia: Hargeisa; HT; ♂; ZMMU. 

phosphila. USA (New York). 

Asynapta phosphila Felt 1907a: 52. USA: New York, Nassau; HT; ♂; NYSM. 

Asynapta photophila Felt 1907c: 148, unjust. emend. of phosphila. 

Holoneurus multinodus Felt 1908b: 420. USA: New York, Nassau; HT; ♂; NYSM. 

Holoneurus multineurus Kieffer 1913i: 266, missp. of multinodus. 

Pseudocamptomyia photophyla Mamaev & Zaitzev 1997c: 13, missp. of photophila. 

PTEREPIDOSIS 

Pterepidosis Mamaev 1990a: 14. Type species, apicalis Mamaev (orig. des.). 


apicalis. Russia (Europe). 

Pterepidosis apicalis Mamaev 1990a: 14. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

varimezovi. Bulgaria. 

Pterepidosis varimezovi Mamaev & Dimitrova 1998: 317. Bulgaria: Vitosha Mtn, nr Sofia; HT; ♂; BIZS. 

*PUTEPIDOSIS 

Putepidosis Fedotova & Sidorenko 2007d: 258. Type species, inevitabilis Fedotova & Sidorenko (orig. des.). 


*inevitabilis. Russia (Far East). 

Putepidosis inevitabilis Fedotova & Sidorenko 2007d: 259. Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; ♂; 

ZMAS. 

*PYXICOCCOPSIS 

Pyxicoccopsis Fedotova & Sidorenko 2007d: 249. Type species, dulcis Fedotova & Sidorenko (orig. des.). 


88

89 

*dulcis. Russia (Far East). 

Pyxicoccopsis dulcis Fedotova & Sidorenko 2007d: 250 (also as dulicis, pp. 251, 252). Russia: Primorskiï Terr., 

Lazovskiï Res., Koreiskaya Pad'; HT; ♂; ZMAS. 

RABINDRODIPLOSIS 

Rabindrodiplosis Grover 1964b: 193. Type species, champakii Grover (mon). 

Oriental; 2 spp.; unplaced to tribe. 

champakii. India (Uttar Pradesh). 

Rabindrodiplosis champakii Grover 1964b: 193. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 


Rabindrodiplosis orientalis Sharma & Rao 1980b: 290. India: Maharashtra; HT; ♂; ELMU. 

*RECESSEPIDOSIS 

Recessepidosis Fedotova & Sidorenko 2008a: 235. Type species, umbonis Fedotova & Sidorenko (orig. des.). 


*umbonis. Russia (Far East). 

Glomepidosis umbonis Fedotova & Sidorenko 2008a: 235. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


RHIPIDOXYLOMYIA 

Rhipidoxylomyia Mamaev 1964c: 898. Type species, rubella Mamaev (orig. des.). 

Palearctic, Oriental, and Eocene amber; 14 spp.; Winnertziini. Ref.: Spungis 1992 (key, Eur. spp.), Jiang & Bu 2004b 

(review, key, Chinese spp.). 

*aequabilis. China (Yunnan). 

Rhipidoxylomyia aequabilis Jiang & Bu 2004b: 358. China: Yunnan, Simao, Caiyanghe Res.; HT; ♂; NKUM. 

brevicornis. Russia (Europe). 

Rhipidoxylomyia brevicornis Mamaev 1964c: 900. Russia: Tula Prov., Krapivenskiï For.; HT; ♂; HYNM. 

concava. Japan (Kyushu). 

Trichoxylomyia concava Yukawa 1967c: 66. Japan: Kyushu, Fukuoka Pref., Mt Hiko; HT; ♂; KUEC. Removed from 

Kronomyia by Jiang & Bu 2004b. 

*elongata. China (Guizhou). 

Rhipidoxylomyia elongata Jiang & Bu 2004b: 359. China: Guizhou, Mt Fanjing, Huguosi; HT; ♂; NKUM. 


Rhipidoxylomyia indica Grover 1971: 12. India: Uttar Pradesh, Allahabad; HT; ♀; PGCA. 

*longilobata. China (Jiangxi). 

Rhipidoxylomyia longilobata Jiang & Bu 2004b: 359. China: Jiangxi, Mt Jinggang, Xiaoxidong; HT; ♂; NKUM. 

minor. Russia (Europe). 

Rhipidoxylomyia minor Mamaev 1964c: 900. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

*nata. Russia (Far East). 

Rhipidoxilomyia nata Fedotova & Sidorenko 2007c: 98. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*orbiculata. China (Henan). 

Rhipidoxylomyia orbiculata Jiang & Bu 2004b: 360. China: Henan, Luanchuan, Longyuwan, Jijiaojian; HT; ♂;

90 

NKUM. 

*ramifera. Russia (Far East). 

Rhipidoxylomyia incisa Fedotova & Sidorenko 2005e: 113. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


rubella. Russia (Europe). 

Rhipidoxylomyia rubella Mamaev 1964c: 899. Russia: Tula Prov., Krapivenskiï For.; HT; ♂; HYNM. 

*simplicis. China (Yunnan). 

Rhipidoxylomyia simplicis Jiang & Bu 2004b: 362. China: Yunnan, Simao, Caiyanghe Res.; HT; ♂; NKUM. 

* † vaga. Late Eocene - Ukrainian amber. 

Rhipidoxylomyia vaga Fedotova & Perkovsky 2008a: 30. Rovno amber, Ukraine, Durovitsa; HT; ♀; UASK. 

*yunnanensis. China (Yunnan). 

Rhipidoxylomyia yunnanensis Jiang & Bu 2004b: 363. China: Yunnan, Pingbian, Mt Dawei; HT; ♂; NKUM. 

*SALPISTEPIDOSIS 

Salpistepidosis Fedotova & Sidorenko 2007d: 272. Type species, silidis Fedotova & Sidorenko (orig. des.). 


*silidis. Russia (Far East). 

Salpistepidosis silidis Fedotova & Sidorenko 2007d: 272. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


SCHISTONEURUS 

Schistoneurus Mamaev 1964c: 910. Type species, impressus Mamaev (orig. des.). 

Schistoneura Skuhravá 1997: 110, missp. Schistoneurus. 

Holarctic; 3 spp.; Holoneurini. Ref.: Mamaev 2001c (review). 

impressus. Russia (Europe). 

Schistoneurus impressus Mamaev 1964c: 910. Russia: Moscow Prov., Kratovo; HT; ♂; ZMMU. 

irregularis. Sweden, Finland, Estonia, Russia (Europe). 

Schistoneurus irregularis Mamaev 1964c: 910. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ZMMU. 

pectinatus. USA (New York). 

Bryocrypta pectinata Felt 1907d: 23. USA: New York, Albany; HT; ♂; NYSM. 

SCLEREPIDOSIS 

Sclerepidosis Mamaev 1990a: 18. Type species, bulbosa Mamaev (orig. des.). 


bulbosa. Russia (Far East). 

Sclerepidosis bulbosa Mamaev 1990a: 18. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

SENSEPIDOSIS 

Sensepidosis Mamaev 1990a: 19. Type species, gukasiani Mamaev (orig. des.). 

Palearctic; 1 sp.; Porricondylini.

gukasiani. Russia (Eastern Siberia). 

Sensepidosis gukasiani Mamaev 1990a: 20. Russia: Tuva, Ishtii-Khem; HT; ♂; HYNM. 

*SEYCHELLEPIDOSIS 

Seychellepidosis Spungis 2007: 63. Type species, spinosa Spungis (orig. des.). 

Afrotropical; 1 sp.; Porricondylini. 

spinosa. Seychelles Is 

Seychellepidosis spinosa Spungis 2007: 63. Seychelles Is: Aldabra; HT; ♂; BMNH. 

*SHEVCHENKIA 

Shevchenkia Fedotova & Sidorenko 2005e: 115. Type species, pulla Fedotova & Sidorenko (orig. des.). 


*pulla. Russia (Far East). 

Shevchenkia pulla Fedotova & Sidorenko 2005e: 115. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


SOLNTSEVIA 

Solntsevia Mamaev 1965a: 266. Type species, nigripes Mamaev (orig. des.). 

Holarctic; 3 spp.; Solntseviini. 

maculata. USA (New York). 

Colpodia maculata Felt 1908b: 416. USA: New York, Albany; HT; ♀; NYSM. 

nigripecta. Russia (Far East). 

Solntsevia nigripecta Mamaev & Zaitzev 1998e: 87. Russia: Kuril Is, Kunashir I.; HT; ♂; ZMMU. 

nigripes. Finland, Estonia, Latvia, Russia (Europe). 

Solntsevia nigripes Mamaev 1965a: 266. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

SPUNGISOMYIA 

Spungisomyia Mamaev & Zaitzev 1996d: 209. Type species, Porricondyla media Spungis (orig. des.). 

Palearctic; 1 sp.; unplaced to tribe. 

media. Finland, Latvia. 

Porricondyla media Spungis 1981: 54. Latvia: Brotseny; HT; ♂; LASS. 

STACKELBERGIELLA 

Stackelbergiella Marikovskij 1958b: 1845. Type species, paradoxa Marikovskij (orig. des.) = hordei. 


hordei. UK, Germany, Latvia, Kazakhstan, USA (New Jersey). 

Colomyia hordei Barnes 1927a: 167. UK, England, Kent, Wye; ST; ♂,♀; BMNH. 

91

Stackelbergiella paradoxa Marikovskij 1958b: 1845. Kazakhstan: Alma-Ata; T; ♂; MARC. 

STOMATOCOLPODIA 

Stomatocolpodia Mamaev 1990a: 17. Type species, iridis Mamaev (orig. des.). 

*Avicolpodia Fedotova & Sidorenko 2008a: 237, as subgenus of Stomatocolpodia. Type species, Stomatocolpodia 

gracilis Fedotova & Sidorenko (orig. des.). 


*cristiformis. Russia (Far East). 

Stomatocolpodia cristiformis Fedotova 2004i: 593. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

*gracilis. Russia (Far East). 

Stomatocolpodia gracilis Fedotova & Sidorenko 2005e: 91. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


iridis. Lithuania, Russia (Europe). 

Stomatocolpodia iridis Mamaev 1990a: 18. Lithuania: Junvintas Res.; HT; ♂; HYNM. 

sinuosa. Russia (Far East). 

Stomatocolpodia sinuosa Mamaev & Zaitzev 1998e: 87. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMMU. 

unidentatus. Kazakhstan. 

Holoneurus unidentatus Marikovskij 1958b: 1844. Kazakhstan: Alma-Ata; T; ♂; MARC. 

SYLVENOMYIA 

Sylvenomyia Mamaev & Zaitzev 1998d: 211. Type species, sueciae Mamaev & Zaitzev (orig. des.) = spinigera 

(Spungis). 

Western Palearctic; 2 spp.; unplaced to tribe. Ref.: Penttinen & Jaschhof 2009 (rev.). 

*fennica. Finland. 

Sylvenomyia fennica Penttinen & Jaschhof 2009: 52. Finland: Ok, Kuhmo, Kieverrysjärvet; HT; ♂; MZHF. 

spinigera. Sweden, Finland, Latvia, Russia (European). 

Chastomera spinigera Spungis 1985: 46. Latvia: Koknese; HT; ♂; LASS. 

Sylvenomyia sueciae Mamaev & Zaitzev 1998d: 212. Sweden: Skane, Klippan, Skaralid; HT; ♂; ZMMU. 

SYNEPIDOSIS 

Synepidosis Mamaev 1964c: 905. Type species, longiventris Mamaev (orig. des.). 


longiventris. Russia (Europe). 

Synepidosis longiventris Mamaev 1964c: 906. Russia: Voronezh Prov., Tellerman For.; HT; ♂; ZMMU. 

voluptaria. Russia (Eastern Siberia). 

Synepidosis voluptaria Mamaev & Zaitzev 1998e: 87. Russia: Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

*TENEPIDOSIS 

Tenepidosis Fedotova & Sidorenko 2005e: 125. Type species, simplificata Fedotova & Sidorenko (orig. des.). 

92


*simplificata. Russia (Far East). 

Tenepidosis simplificata Fedotova & Sidorenko 2005e: 125. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*TERSEPIDOSIS 

Tersepidosis Fedotova & Sidorenko 2007d: 236. Type species, vigilis Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Solntseviini 

*vigilis. Russia (Far East). 

Tersepidosis vigilis Fedotova & Sidorenko 2007d: 236. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


TETRANEUROMYIA 

Tetraneuromyia Mamaev 1964c: 906. Type species, bulbifera Mamaev (orig. des.). 


bulbifera. Latvia, Russia (Europe). 

Tetraneuromyia bulbifera Mamaev 1964c: 906. Russia: Voronezh Prov., Tellerman For.; HT; ♂; IEME. 

hirticornis. Denmark, Latvia, Poland, Austria, Russia (Europe). 

Cecidomyza hirticornis Zetterstedt 1850: 3680. Denmark: Copenhagen; LT (Panelius 1965: 26); ♂; ZMUC. 

lamellata. Latvia. 

Tetraneuromyia lamellata Spungis 1987: 35. Latvia: Darzini; HT; ♂; LASS. 

multipartita. Russia (Europe). 

Tetraneuromyia multipartita Mamaev 1998d: 3. Russia: Moscow Prov., Lugovaya; HT; ♂; ZMMU. 

similis. Latvia, Russia (Europe). 

Tetraneuromyia similis Mamaev 1964c: 907. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

*THAUMACECIDOMYIA 

Thaumacecidomyia Yang 1998: 10. Type species, sinica Yang (orig. des.). 

Palearctic; 1 sp.; Diallactini. 

*sinica. China (Zhejiang). 

Thaumacecidomyia sinica Yang 1998: 10. China: Zhejiang, Anjico, Longwangshan; HT; ♂; BAUC. 

TRICHEPIDOSIS 

Trichepidosis Mamaev 1990a: 15. Type species, petiolata Mamaev (orig. des.). 


petiolata. Russia (Europe). 

Trichepidosis petiolatus Mamaev 1990a: 16. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ZMMU. 

*TUTKOWSKIA 

93

Tutkowskia Fedotova & Perkovsky 2008c: 417. Type species, ukrainica Fedotova & Perkovsky (orig. des.) 

Rovno amber (Ukraine); 1 sp.; Heteropezini. 

* † Hukrainica. Late Eocene - Ukrainian amber. 

Tutkowskia ukrainica Fedotova & Perkovsky 2008c: 418. Rovno amber, Ukraine; HT; ♂; UASK. 

UBINOMYIA 

Ubinomyia Mamaev 1990a: 13. 10 OD. Type species, cardinalis Mamaev (orig. des.). 


cardinalis. Russia (Europe). 

Ubinomyia cardinalis Mamaev 1990a: 13. Russia: Krasnodarskiï Terr., Ubinskaya; HT; ♂; ZMMU. 

*USTINEPIDOSIS 

Ustinepidosis Fedotova & Sidorenko 2005e: 99 9also as Ustnieipidosis, p. 100). Type species, korkishkoi Fedotova & 

Sidorenko (orig. des.). 


*korkishkoi. Russia (Far East). 

Ustinepidosis korkishkoi Fedotova & Sidorenko 2005e: 99. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

*VASILIOLA 

Vasiliola Fedotova 2004i: 585. Type species, evanida Fedotova (orig. des.). 

Palearctic; 1 sp.; Winnertziini?. Ref.: Jaschhof 2009: 86 (placement). 

*evanida. Russia (Far East). 

Vasiliola evanida Fedotova 2004i: 586. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

*VELAFACERA 

Velafacera Fedotova & Perkovsky 2008a: 28. Type species, vectabilis Fedotova & Perkovsky (orig. des.) 

Rovno amber (Ukraine); 1 sp.; Dicerurini. 

* † vectabilis. Late Eocene - Ukrainian amber. 

Velafacera vectabilis Fedotova & Perkovsky 2008a: 28. Rovno amber, Ukraine, Durovitsa; HT; ♀; UASK. 

*VENTOSAGLORIA 

Ventosagloria Fedotova & Perkovsky 2008c: 412. Type species, gratshevi Fedotova & Perkovsky (orig. des.) 

Rovno amber (Ukraine); 3 spp.; Heteropezini. 

94

* † contradictoria. Late Eocene - Ukrainian amber. 

Ventosagloria contradictoria Fedotova & Perkovsky 2008c: 415. Ukraine amber: Dubrovitsa; HT; ♂; UASK. 

* † fassa. Late Eocene - Ukrainian amber. 

Ventosagloria fassa Fedotova & Perkovsky 2008c: 416. Ukraine amber: Klesov; HT; ♂; UASK. 

* † gratshevi. Late Eocene - Ukrainian amber. 

Ventosagloria gratshevi Fedotova & Perkovsky 2008c: 414. Ukraine amber: Dubrovitsa; HT; ♂; UASK. 

*VOLNOCOCCOPSIS 

Volnococcopsis Fedotova & Perkovsky 2008a: 35. Type species, korniushini Fedotova & Perkovsky (orig. des.) 

Rovno amber (Ukraine); 1 sp.; Holoneurini. 

* † korniushini. Late Eocene - Ukrainian amber. 

Volnococcopsis korniushini Fedotova & Perkovsky 2008a: 35. Rovno amber, Ukraine, Durovitsa; HT; ♀; UASK. 

*VULNEPIDOSIS 

Vulnepidosis Fedotova & Sidorenko 2007d: 239. Type species, reburra Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Solntseviini. 

*reburra. Russia (Far East). 

Vulnepidosis reburra Fedotova & Sidorenko 2007d: 239. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


WINNERTZIA 

Winnertzia Rondani 1860: 290. Type species, Cecidomyia lugubris Winnertz (orig. des.). 

Clinorhiza Kieffer 1894c: 340. Type species, citrina Kieffer (orig. des.) = anomala Kieffer. 

Clinorrhyza Kieffer 1895h: 121, missp. of Clinorhiza. 

Prowinnertzia Kieffer 1911: 327. Type species, mahensis Kieffer (mon.). 

Winnertziola Kieffer 1913a: 235. Type species, Winnertzia rubra Kieffer (orig. des.). 

Winnertiziola Shinji 1944: 213, missp. of Winnertziola. 

Xylowinnertzia Mamaev 1963b: 564, as subg. of Winnertzia. Type species, Winnertzia rubricola Mamaev (orig. des.). 

Mycowinnertzia Mamaev 1963b: 567, as subg. of Winnertzia. Type species, Winnertzia nigra Mamaev (orig. des.). 

Holarctic, Oriental, Afrotropical, also Baltic, Ukrainian, and Burmese amber; 113 spp.; Winnertziini. Refs: Panelius 

1965 (rev., Eur. spp.), Parnell 1971 (review, Nearctic spp.), Spungis 1992 (key, Eur. spp.). 

†affinis. Upper Eocene - Baltic amber. 

Winnertzia affinis Meunier 1904: 38. Baltic amber; T; ♀; DGOT?. 

alba. Russia (Far East). 

Winnertzia alba Mamaev 2002b: 11. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

amoena. Russia (Far East). 

Winnertzia amoena Mamaev 1998f: 41. Russia: Amur Prov., Khingansky Res.; HT; ♂; ZMMU. 

ampelophila, USA (New York, Maryland). 

Porricondyla ampelophila Felt 1907a: 48. USA: New York, Albany; HT; ♂; NYSM. 

anomala. France. 

Clinorhiza citrina Kieffer 1894c: 340, jun. sec. hom. Winnertzia citrina Kieffer. France: Bitche; ST; A; lost?. 

Winnertzia anomala Kieffer 1896g: 34, new name for Clinorhiza citrina Kieffer. 

arizoniensis. USA (Arizona). 

95

96 

Winnertzia arizoniensis Felt 1908b: 421. USA: Arizona, Williams; HT; ♀; NYSM. 

Winnertzia arizonensis Kieffer 1913i: 282, missp. of arizoniensis. 

asiatica. Uzbekistan. 

Winnertzia asiatica Mamaev 1963b: 570. Uzbekistan: Kyzyl Kum, Kuldzhuktau Mts; HT; ♂; ZMMU. 

assimilata. Russia (Europe, Far East). 

Winnertzia assimilata Mamaev 1972b: 114. Russia: Primorskiï Terr., Ussuriyskiï Distr., Suputinskiï Res.; HT; ♂; 

HYNM. 

* † bellata. Late Eocene - Ukrainian amber. 

Winnertzia bellata Fedotova 2005e: 46. Rovno amber, Ukraine; HT; ♀; UASK. 

betulicola. Russia (Europe, Far East). 

Winnertzia betulicola Mamaev 1963b: 568. Russia: Tula Prov., Tula For.; HT; ♂; ZMMU. 

Winnertzia befulicola Mamaev 1998f: 46, missp. of betulicola. 

blanda. Russia (Europe). 

Winnertzia blanda Mamaev 2002b: 7. Russia: Krasnodarskiï Terr., Krasnaya Polyana, Medvezh'i Vorota; HT; ♂; 

ZMMU. 

brachypalpa. Ukraine, Russia (Far East). 

Winnertzia brachypalpa Mamaev 1975d: 85. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

brevis. Italy. 

Winnertzia brevis Mamaev 2002c: 1. Italy: Siena, Monte Amiata; HT; ♂; ZMMU. 

bulbifera. Germany, Latvia, Russia (Europe). 

Winnertzia bulbifera Mamaev 1963b: 571. Russia: Moscow Prov., Kropotovo; HT; ♂; ZMMU. 

† 

burmitica. Eocene - Burmese amber. 

Winnertziola burmitica Cockerell 1917: 42. Burmese amber, Burma; HT; ♀; BMNH. 

carinata. Russia (Far East). 

Winnertzia carinata Mamaev 2002c: 2. Russia: Khabarovsk Terr., Khekhtsyr Res.; HT; ♂; ZMMU. 

carpini. USA (New York). 

Winnertzia carpini Felt 1907a: 52. USA: New York, Albany; HT; ♂; NYSM. 

carpinicola. France. 

Winnertzia carpinicola Kieffer 1913f: 47. France: Bitche; ST; ♂,♀,P,L; lost?. 

cauta. Russia (Europe). 

Winnertzia cauta Mamaev 2002c: 2. Russia: Stavropol; HT; ♂; ZMMU. 

centralis. Russia (Europe). 

Winnertzia centralis Mamaev 2001e: 1. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

citrina. France. 

Winnertzia citrina Kieffer 1888b: 243. France: vic. Bitche; ST; ♂,♀,L; lost?. 

colubrifera. Russia (Far East). 

Winnertzia colubrifera Mamaev 1975b: 54. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

conorum. Germany, Hungary. 

Winnertzia conorum Kieffer 1920a: 21. Germany: vic. Munich; ST; ♂,♀; lost?. 

cornigera. Russia (Europe). 

Winnertzia cornigera Mamaev 2002c: 3. Russia: Moscow Prov., Glubokoe L.; HT; ♂; ZMMU. 

corticis. France. 

Winnertzia corticis Kieffer 1913f: 52. France: Bitche; ST; ♀,L; lost?. 

cumulata. Kirghizstan 

Winnertzia cumulata Mamaev 2002b: 6. Kirghizstan: Sary-Chelek; HT; ♂; ZMMU. 

curvata. UK, Germany, Estonia, Latvia, Russia (Europe). 

Winnertzia curvata Panelius 1965: 119. UK: England, Harpenden, Rothhamsted Lodge; HT; ♂; BMNH. 

† 

cylindrica. Upper Eocene - Baltic amber. 

Winnertzia cylindrica Meunier 1904: 37. Baltic amber; T; ♀; DGOT?. 

deserticola. Uzbekistan. 

Winnertzia deserticola Mamaev 1963b: 569. Uzbekistan: Kyzyl Kum, Kuldzhukhtau Mts; HT; ♂; HYNM. 

detrita. Russia (Far East).

97 

Winnertzia detrita Mamaev 1975d: 85. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

discreta. Turkmenistan. 

Winnertzia discreta Mohrig & Mamaev 1970: 334. Turkmenistan: Badkhyz Res., 80 km NW Kushka; HT; ♀; 

ZMMU. 

discretella. Latvia, Russia (Europe). 

Winnertzia discretella Spungis 1992: 19. Latvia: Broc ni; HT; ♂; LASS. 

dispar. Russia (Europe). 

Winnertzia dispar Mamaev 2002c: 4. Russia: Voronezh Prov., Tellerman; HT; ♂; ZMMU. 

divergens. Italy. 

Winnertzia divergens Mamaev 2002c: 4. Italy: Siena, Monte Amiata; HT; ♂; ZMMU. 

diversicornis. Russia (Far East). 

Winnertzia diversicornis Mamaev 1998f: 44. Russia: Khabarovsk Prov., Bychikha; HT; ♂; ZMMU. 

equestris. Germany, Russia (Europe). 

Winnertzia equestris Mamaev 1963b: 572. Russia: Dagestan, Tlyarata; HT; ♂; HYNM. 

feralis. Ukraine. 

Winnertzia feralis Mamaev 2002b: 8. Ukraine: Zakarpatskaya Prov., Rakhov; H; ♂; ZMMU. 

finitima. Russia (Far East). 

Winnertzia finitima Mamaev 2002b: 8. Russia: Khabarovsk Terr., Khekhtsyr Res.; HT; ♂; ZMMU. 

flaveola. Russia (Far East). 

Winnertzia flaveola Mamaev 2002a: 6. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

fumosa. Russia (Far East). 

Winnertzia fumosa Mamaev 2002a: 8. Russia: Primorskiï Terr., Ussuriyskiï (Suputinskiï) Res.; HT; ♂; ZMMU. 

fungicola. USA (New Jersey). 

Winnertzia fungicola Felt 1921b: 118. USA: New Jersey, Plainfield; LT (Parnell 1971: 288); ♂; NYSM. 

fungorum. Russia (Europe). 

Winnertzia fungorum Mamaev 2002c: 5. Russia: Krasnodarskiï Terr., Kudepsta; HT; ♂; ZMMU. 

fusca. France, Latvia, Russia (Far East). 

Winnertzia fusca Kieffer 1901b: 464. France: Bitche; NT (Spungis 1992: 27, Latvia: Saulkalne); ♂; LASS. 

graduata. Finland, Estonia, Latvia, Ukraine. 

Winnertzia graduata Spungis 1992: 28. Latvia: Darzini; HT; ♂; LASS. 

griseipennis. Austria. 

Asynapta griseipennis Löw 1874a: 148. Austria: Vienna, Wienerwald; ST; ♀; NHMW. 

hikosanensis. Japan (Kyushu). 

Winnertzia hikosanensis Yukawa 1967c: 68. Japan: Kyushu, Fukuoka Pref., Mt Hiko; HT; ♂; KUEC. 

hudsonici. USA (New York). 

Winnertzia hudsonici Felt 1908b: 422. USA: New York, Albany; T; ♀; NYSM. 

Winnertzia karnerensis Felt 1908b: 422. USA: New York, Karner; HT; ♀; NYSM. 


Winnertzia indica Grover 1971: 8. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

invisibilis. Italy. 

Winnertzia invisibilis Mamaev 2001e: 2. Italy: Pontignano; HT; ♂; ZMMU. 

iridis. Russia (Europe). 

Winnertzia iridis Mamaev 2002b: 4. Russia: Kaliningrad Prov., Ribachiï; HT; ♂; ZMMU. 

* † isotoma. Late Eocene - Ukrainian amber. 

Winnertzia isotoma Fedotova 2005e: 46. Rovno amber, Ukraine; HT; ♂; UASK. 

kamchatica. Russia (Far East). 

Winnertzia kamchatica Mamaev 2002c: 5. Russia: Kamchatka, Kozyrevsk; HT; ♂; ZMMU. 

* † kapustini. Late Eocene - Ukrainian amber. 

Winnertzia kampustini Fedotova & Perkovsky 2008a: 74. Rovno amber, Ukraine, Durovitsa; HT; ♀; UASK. 

kovalevi. Russia (Far East). 

Winnertzia kovalevi Mamaev 2002c: 6. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

kushkensis. Turkmenistan.

98 

Winnertzia kushkensis Mamaev 2001d: 2. Turkmenistan: Kushka, Badkhyz Res.; HT; ♂; ZMMU. 

levicollis. France. 

Winnertzia levicollis Kieffer 1901b: 460. France: Bitche; T; LT; lost?. 

longiptera. Sweden. 

Winnertzia longiptera Mamaev 2002c: 7. Sweden: Uppsala; HT; ♂; ZMMU. 

lugubris. Germany, Czechoslovakia, Austria. 

Winnertzia lugubris Winnertz 1853: 305. Germany; ST; ♂,♀; lost. 

maacki. Russia (Far East). 

Winnertzia maacki Mamaev 1975b: 53. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 


Prowinnertzia mahensis Kieffer 1911: 327. Seychelles Is: Mahé, Cascade Estate; HT; ♀; BMNH. 

maxima. Russia (Europe). 

Winnertzia maxima Mamaev 1963b: 568. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM (Yagi & Sawada 

2000) - ZMMU (Krivosheina & Ozerov 2006). 

monarthra. Russia (Western Siberia). 

Winnertzia monarthra Mamaev 1990b: 143. Russia: Yamal Peninsula, Khadyta; HT; ♂; ZMMU. 

mycetaula. Russia (Europe). 

Winnertzia mycetaula Mamaev 2002a: 5. Russia: Moscow Prov., Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

nigra. Latvia, Ukraine, Russia (Europe, Far East). 

Winnertzia nigra Mamaev 1963b: 567. Russia: Voronezh Prov., Tellerman For.; HT; ♂; HYNM. 

nigricolor. Russia (Europe). 

Winnertzia nigricolor Mamaev 2002a: 5. Russia: Moscow Prov., Danki; HT; ♂; ZMMU. 

nigripennis. France, Germany, Finland, Latvia, Russia (Europe). 

Clinorhiza nigripennis Kieffer 1894c: 340, nomen nudum. 

*Clinorhiza nigripennis Kieffer 1894k: unnumbered p. 1. Unstated, presumably France: Bitche; HT; egg (illustrated); 

lost? 

Winnertzia nigripennis Kieffer 1896g: 36. France: Bitche; ST; ♂,♀,L,P,egg; lost?. 

Winnertzia pinicola Kieffer 1913f: 49. France: Bitche; ST; ♂,♀,L; lost?. 

obscura. France. 

Winnertzia obscura Kieffer 1894c: 345. France: Bitche; ST; ♂,♀,L,egg; lost?. 

obscurella. Russia (Far East). 

Winnertzia obscurella (also as obcurella) Mamaev 2002a: 7. Russia: Primorskiï Terr., Ussuriyskiï (Suputinskiï) Res.; 


obscuricornis. Russia (Europe). 

Winnertzia obscuricornis Mamaev 2001e: 2. Russia: Voronezh Prov., Tellerman For.; HT; ♂; ZMMU. 

orientalis. India (Madhya Pradesh). 

Winnertzia orientalis Grover 1971: 9. India: Madhya Pradesh, Indore; HT; ♀; PGCA. 

padicola. Latvia, USA (Maryland). 

Winnertzia padicola Spungis 1992: 32. Latvia: Kauguri; HT; ♂; LASS. 

pallidicauda. Russia (Europe). 

Winnertzia pallidicauda Mamaev 2002c: 7. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

pallidula. Russia (Europe). 

Winnertzia pallidula Mamaev 2002c: 8. Russia: Krasnodarskiï Terr., Ubinskaya; HT; ♂; ZMMU. 

palpina. Russia (Far East). 

Winnertzia palpina Mamaev 1998f: 44. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

palustris. USA (New York). 

Winnertzia palustris Felt 1915j: 133. USA: New York, Canada L., Caroga; HT; ♀; NYSM. 

pectinulata. Russia (Far East). 

Winnertzia pectinulata Mamaev 1975d: 86. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

pellucida. Russia (Far East). 

Winnertzia pellucida Mamaev 2002c: 8. Russia: Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

peramoena. Russia (Far East).

99 

Winnertzia peramoena Mamaev 1998f: 43. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

pictipes. France. 

Clinorhiza pictipes Kieffer 1894c: 340, nomen nudum. 

Winnertzia pictipes Kieffer 1896g: 37. France: Bitche; ST; ♂,P,L,egg; lost?. 

pinicorticis. USA (Virginia). 

Winnertzia pinicorticis Felt 1907d: 23. USA: Virginia, Strom; T; ♂; USNM. 

plagiata. Russia (Far East). 

Winnertzia plagiata Mamaev 1975a: 39. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

*plastica. Russia (Far East). 

Winnertzia plastica Fedotova & Sidorenko 2005e: 109. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


populicola. Latvia, Poland. 

Winnertzia populicola Spungis 1992: 32. Latvia: Saulkalne; HT; ♂; LASS. 

pravdini. Estonia, Latvia, Ukraine, Russia (Europe). 

Winnertzia pravdini Mamaeva & Mamaev 1971: 9. Russia: Krasnodarskiï Terr., Adlerskiï Region, Prekhoko Mts; 


propria. Turkmenistan. 

Winnertzia propria Mamaev 2001d: 3. Turkmenistan: Kushka, Badkhyz Res.; HT; ♂; ZMMU. 

proxima. France. 

Winnertzia proxima Kieffer 1894c: 344. France: Bitche; ST; ♂,♀,L; lost?. 

pustulata. Latvia, Ukraine. 

Winnertzia pustulata Spungis 1992: 24. Latvia: Jumprava; HT; ♂; LASS. 

quercicola. France. 

Winnertzia quercicola Kieffer 1913f: 50. France: Bitche; ST; ♂,♀,L; lost?. 

† 

radiata. Upper Eocene - Baltic amber. 

Winnertzia radiata Meunier 1904: 36. Baltic amber; ST; ♂,♀; DGOT?. 

rara. Russia (Europe). 

Winnertzia rara Mamaev 2002b: 10. Russia: Krasnodarskiï Terr., Novo-Prokhladnoye; HT; ♂; ZMMU. 

* † recusata. Late Eocene - Ukrainian amber. 

Winnertzia recusata Fedotova & Perkovsky 2008b: 64. Rovno amber, Ukraine; HT; ♂; UASK. 

reducta. Turkmenistan. 

Winnertzia reducta Mamaev 2001d: 4. Turkmenistan: Kushka, Badkhyz Res.; HT; ♂; ZMMU. 

regia. Russia (Europe). 

Winnertzia regia Mamaev 2002c: 9. Russia: Moscow Prov., Tuva, Ishtii-Khem; HT; ♂; ZMMU. 

rotundata. Latvia, Estonia, Russia (Far East). 

Winnertzia rotundata Spungis 1992: 17. Latvia: Jumprava; HT; ♂; LASS. 

rubida. USA (New York). 

Winnertzia rubida Felt 1908b: 422. USA: New York, Albany; HT; ♂; NYSM. 

rubra. France. 

Winnertzia rubra Kieffer 1894c: 343. France: Bitche; ST; ♂,♀,L,egg; lost?. 

rubricola. Russia (Europe), Azerbaijan. 

Winnertzia rubricola Mamaev 1963b: 564. Russia: Tula Prov., Krapiven For.; HT; ♂; HYNM. 

salicis. Germany. 

Cecidomyia salicis Bouché 1834: 27, jun. prim. hom. salicis Schrank 1803. Germany; ST; ♂,♀,P,L; ZMHB?. 

Cecidomyia boucheana Loew 1850a: 24, new name for Cecidomyia salicis Bouché. 

semideserta. Turkmenistan. 

Winnertzia semideserta Mamaev 2001d: 5. Turkmenistan: Kushka, Badkhyz Res.; HT; ♂; ZMMU. 

† 

separata. Upper Eocene - Baltic amber. 

Winnertzia separata Meunier 1904: 38. Baltic amber; T; ♀; DGOT?. 

sequentis. Sweden. 

Winnertzia sequentis Mamaev 2001e: 3. Sweden: Dalarna; HT; ♂; ZMMU. 

solidaginis. USA (New York, Pennsylvania), Finland, Estonia, Latvia, Ukraine, Russia (Europe, Far East), Japan.

Winnertzia solidaginis Felt 1907a: 53. USA: New York, Nassau; HT; ♂; NYSM. 

Winnertzia calciequina Felt 1907c: 161. USA: New York, Albany; LT (Parnell 1971: 290) ♂; NYSM. 

Winnertzia pectinata Felt 1911l: 457, nomen nudum. 

Winnertzia pectinata Felt 1911m: 478. USA: New York, Nassau; LT (Parnell 1971: 290); ♂; NYSM. 

Winnertzia aceris Felt 1913d: 213. USA: New York, Nassau; LT (Parnell 1971: 290); ♂; NYSM. 

Winnertzia clypeata Mamaev 1975b: 54. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

speciosa. Russia (Far East). 

Winnertzia speciosa Mamaev 2002a: 9. Russia: Primorskiï Terr., Ussuriyskiï (Suputinskiï) Res.; HT; ♂; ZMMU. 

striaticollis. France. 

Winnertzia straticollis Kieffer 1901b: 460. France; T; LT; lost?. 

subglobifera. Russia (Far East). 

Winnertzia subglobifera Mamaev 1998f: 46. Russia: Kuril I., Kunashir; HT; ♂; ZMMU. 

sublongiptera. Italy. 

Winnertzia sublongiptera Mamaev 2002c: 9. Italy: Siena, Monte Amiata; HT; ♂; ZMMU. 

superna. Russia (Far East). 

Winnertzia superna Mamaev 2002a: 9. Russia: Primorskiï Terr., Ussuriyskiï (Suputinskiï) Res.; HT; ♂; ZMMU. 

tarsata. Russia (Europe). 

Winnertzia tarsata Mamaev 2002c: 10. Russia: Moscow Prov., Glubokoe L.; ST; ♂, ♀; ZMMU. 

tenella. UK. 

Cecidomyia tenella Walker 1856a: 129. UK: England; ST; ♂,♀; lost. 

tergata. Russia (Far East). 

Winnertzia tergata Mamaev 2002c: 10. Russia: Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

triangulifera. Italy. 

Winnertzia triangulifera Mamaev 2001e: 4. Italy: Pontignano; HT; ♂; ZMMU. 

ussurica. Russia (Far East). 

Winnertzia ussurica Mamaev 1975d: 84. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; ZMMU. 

vexans. France. 

Winnertziola vexans Kieffer 1913f: 47. France: Bitche; T; ♀; lost?. 

WYATTELLA 

Wyattella Mamaev 1966: 221. Type species, ussuriensis Mamaev (orig. des.). 

Palearctic, Oriental; 3 spp.; Diallactini. 

fungicola. Russia (Far East). 

Wyattella fungicola Mamaev 1976: 1. Russia: Khabarovsk Terr., Khekhtsyr Res., nr Khabarovsk; HT; ♂; HYNM. 

lobata. China (Hong Kong). 

Wyattella lobata Yukawa 1968b: 251. China: Hong Kong; HT; ♂; KUEC. 


Wyattella ussuriensis Mamaev 1966: 221. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

*YUKAWAEPIDOSIS 

Yukawaepidosis Fedotova & Sidorenko 2007d: 265. Type species, aliculata Fedotova & Sidorenko (orig. des.). 


aliculatus. New Zealand (Campbell I.). 

Holoneurus aliculatus Yukawa 1964: 302. New Zealand: Campbell I., Beeman Hill; HT; ♂; NZAC. 

100

*ZAITZEVIOLA 

Zaitzeviola Fedotova & Sidorenko 2007c: 82. Type species, Neocolpodia dubitabila Mamaev & Zaitzev (orig. des.). 


dubitabila. Russia (Europe, Far East). 

Neocolpodia dubitabila Mamaev & Zaitzev 1998e: 88. Russia: Kamchatka, Kozyrevsk; HT; ♂; ZMMU. 

Zaitzeviola dubitabilis Fedotova 2007c: 82, missp. dubitabila. 

gracilipennis. Japan (Kyushu), Russia (Far East). 

Porricondyla gracilipennis Yukawa 1971: 68. Japan: Kyushu, Fukuoka-City, Ropponmatsu; HT; ♂; KUIC. 

ZATSEPINOMYIA 

Zatsepinomyia Mamaev & Zaitzev 1997c: 5. Type species, inopina Mamaev & Zaitzev (orig. des.). 

Afrotropical; 2 spp.; unplaced to tribe. 

inopina. Somalia. 

Zatsepinomyia inopina Mamaev & Zaitzev 1997c: 6. Somalia: Hargeisa; HT; ♂; ZMMU. 

intercalaris. Somalia. 

Zatsepinomyia intercalaris Mamaev & Zaitzev 1997c: 6. Somalia: Hargeisa; HT; ♂; ZMMU. 

*ZEPHYREPIDOSIS 

Zephyrepidosis Fedotova & Sidorenko 2008a: 238. Type species, pilifera Fedotova & Sidorenko (orig. des.). 


*pilifera. Russia (Far East). 

Dendrepidosis pilifera Fedotova & Sidorenko 2005e: 96. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


SUBFAMILY CECIDOMYIINAE 

ACACIDIPLOSIS 

Acacidiplosis Gagné 1993c: 84. Type species, spinosa Gagné (orig. des.). 

Afrotropical; 13 species, all forming bud galls on acacias; Cecidomyiidi, Cecidomyiini. 

ananas. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis ananas Gagné 1993c: 88. Kenya: Voi; HT; ♂; USNM. 

cespitosa. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis cespitosa Gagné 1993c: 92. Kenya: Yatta; HT; ♂; USNM. 

conica. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis conica Gagné 1993c: 92. Kenya: Yatta; HT; ♂; USNM. 

crispa. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis crispa Gagné 1993c: 94. Kenya: Kima, nr Sultan Hamid; HT; ♂; USNM. 

echinata. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis echinata Gagné 1993c: 94. Kenya: Makindu; HT; ♂; USNM. 

erupta. Kenya. Acacia tortilis (Fabaceae). 

101

102 

Acacidiplosis erupta Gagné 1993c: 95. Kenya: Tsavo East Natl Pk; HT; ♂; USNM. 

hamata. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis hamata Gagné 1993c: 95. Kenya: L. Baringo, Baringo Town; HT; P; USNM. 

imbricata. Kenya. Acacia nilotica (Fabaceae). 

Acacidiplosis imbricata Gagné 1993c: 96. Kenya: Rt. A109, Kapiti Plains, Malili Ranch, ca. 70 km SE Nairobi; HT; 

LT; USNM. 

ramosa. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis ramosa Gagné 1993c: 96. Kenya: southern base Ngong Hills; HT; P; USNM. 

rugosa. Kenya. Acacia drepanolobium (Fabaceae). 

Acacidiplosis rugosa Gagné 1993c: 96. Kenya: southern base Ngong Hills; HT; ♂; USNM. 

spinosa. Kenya. Acacia nilotica (Fabaceae). 

Acacidiplosis spinosa Gagné 1993c: 97. Kenya: Rt. A109, pipeline trees, Kilugu Ranch, ca. 80 km SE Nairobi; HT; 

♂; USNM. 

undulata. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis undulata Gagné 1993c: 98. Kenya: Yatta; HT; LT (orig. as ♂ in error); USNM. 

verticillata. Kenya. Acacia tortilis (Fabaceae). 

Acacidiplosis verticillata Gagné 1993c: 98. Kenya: 54 km E Athi R. Township; HT; ♂; USNM. 

*ACANTHACIDIPLOSIS 

Acanthacidiplosis Fedotova 2003e: 59. Type species, erinacea Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. This genus should be compared to the cosmopolitan 

Dichodiplosis. Fedotova 2003e erected for this one genus the tribe Acanthacidiplosini but it is not used here (see 

Appendix). 

*erinacea. Russia (Far East). 

Acanthacidiplosis erinaceus Fedotova 2003e: 60. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

ACERICECIS 

Acericecis Gagné 1983e: 707. Type species, Cecidomyia ocellaris Osten Sacken (orig. des.). 

Harrisomyia Skuhravá & Skuhravý 1986: 259. Type species, Perrisia vitrina Kieffer (orig. des.). 

Holarctic and Miocene; 4 spp., all from leaf galls on Acer; Lasiopteridi, unplaced to tribe. 

campestre. United Kingdom, Italy. Acer campestre (Aceraceae). 

Acericecis campestre Harris 2004c: 41. United Kingdom: Surrey, East Horsley, Sheepleas; HT; larva; BMNH. 

† chaneyi. Miocene - USA (Oregon). Acer osmonti (Aceraceae). 

Cecidomyia chaneyi Cockerell 1927: 64. Miocene, John Day Formation - USA: Oregon, 11 km E Post, locality 3930; 

T; G; ?. 

ocellaris. Widespread Nearctic. Acer rubrum; Acer spp. (Aceraceae). 

Cecidomyia ocellaris Osten Sacken 1862: 199. USA: District of Columbia; LT (Gagné 1983e: 709); G; MCZ. 

Sciara ocellata Beutenmüller 1904: 119, missp. of ocellaris. 

Mayetiola virginiana Felt 1908b: 369. USA: New York, Albany; HT; ♀; NYSM. 

vitrina. Widespread Europe. Acer pseudoplatanus (Aceraceae). 

Perrisia vitrina Kieffer 1909b: 1. France: Bitche; NT (Skuhravá & Skuhravý 1986: 260, Czech Republic: Závisín, nr 

Mariánské Lázn); ♂; SKCR. 

*ACEROVESICULOMYIA

Acerovesiculomyia Fedotova 2003c: 14. Type species, marikovskii Fedotova (orig. des.). 

Palearctic; 1 sp., from leaf galls on Acer; Cecidomyiidi, Ceciodmyiini. 

*marikovskii. Russia (Far East). Acer mono (Salicaceae). 

Acerovesiculomyia marikovskii Fedotova 2003c: 15. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


*ACINACISTYLA 

Acinacistyla Fedotova & Sidorenko 2006b: 11. Type species, papposa Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Brachineuridi. 

*papposa. Russia (Far East). 

Acinacistyla papposa Fedotova & Sidorenko 2006b: 12. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


ACODIPLOSIS 

Acodiplosis Kieffer 1895f: cxciv. Type species, Cecidomyia inulae Loew (orig. des.). 

Arthrocerastis Rubsaamen 1895b: 186. Type species, Cecidomyia inulae Loew (orig. des.). 

Neurodiplosis Kieffer 1913b: 90, as subg. of Acodiplosis. Type species, Acodiplosis pulicariae Kieffer (orig. des. as 

n. subg., n. sp.). 

Western Palearctic; 2 spp.; Cecidomyiidi, unplaced to tribe. The two species were placed by Kieffer in two separate 

subgenera that do not necessarily form a natural group. The female flagellomeres have a definite constriction in A. 

inulae but not in A. pulicariae, and the male of the latter is unknown. Ref.: Rübsaamen 1926 (illus.). 

inulae. Widespread Europe. Inula britannica; I. ensifolia (Asteraceae). 

Cecidomyia inulae Loew 1847: 299. Poland; ST; ♂,♀,L,P; ZMHB?. 

pulicariae. Portugal, Algeria. Pulicaria odora (Asteraceae). 

Acodiplosis pulicariae Kieffer 1913b: 90. Algeria: Sahel, vic. Bouzaréa, nr Algiers; ST; ♀,P,L,egg; lost?. 

ACTILASIOPTERA 

Actilasioptera Gagné 1999a: 23, invalid, described without type species designation. 

Actilasioptera Gagné 1999c: 697. Type species, tumidifolium Gagné (orig. des.). 

Oriental, Australasian; 6 spp., all from leaf galls on Avicennia; Lasiopteridi, Lasiopterini. 

coronata. Australia (Queensland). Avicennia marina (Avicenniaceae). 

Actilasioptera coronata Gagné 1999a: 32. Australia: Queensland, Sandgate; HT; ♂; ANIC. 

falcaria. Indonesia (Java). Avicennia officinalis (Avicenniaceae). 

Stefaniella falcaria Felt 1921h: 141. Indonesia: Java, Semarang; ST; ♂,♀; NYSM. 

pustulata. Australia (Queensland). Avicennia marina (Avicenniaceae). 

Actilasioptera pustulata Gagné 1999a: 32. Australia: Queensland, Sandgate; HT; ♂; ANIC. 

subfolium. Australia (Queensland). Avicennia marina (Avicenniaceae). 

Actilasioptera subfolium Gagné 1999a: 28. Australia: Queensland, Brighton; HT; ♂; ANIC. 

tuberculata. Australia (Queensland). Avicennia marina (Avicenniaceae). 

Actilasioptera tuberculata Gagné 1999a: 33. Australia: Queensland, Wellington Point; HT; ♂; ANIC. 

tumidifolium. Australia (Queensland). Avicennia marina (Avicenniaceae). 

103

Actilasioptera tumidifolium Gagné 1999a: 25. Australia: Queensland, Sandgate; HT; ♂; ANIC. 

*ACULEATODIPLOSIS 

Aculeatodiplosis Fedotova & Sidorenko 2005a: 214. Type species, fasciata Fedotova & Sidorenko (orig. des.). 

Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. 

*fasciata. Russia (Far East). 

Aculeatodiplosis fasciata Fedotova & Sidorenko 2005a: 216. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. 


*ACUMYIA 

Acumyia Harris 2008: 22. Type species, acericola Harris (orig. des.). 

Palearctic; 1 sp.; Lasiopteridi, Dasineurini. 

*acericola. UK (England). Acer campestre; A. platanoides (Aceraceae) 

Acumyia acericola Harris 2008: 23. UK: England, Essex, Hainault Forest/Lambourne Common; HT; ♂; BMNH. 

ADIPLOSIS 

Adiplosis Felt 1908b: 405. Type species, Cecidomyia toxicodendri Felt (orig. des.). 

Nearctic; 1 sp.; Cecidomyiidi, Lestodiplosini. This genus is similar to many species now under Trisopsis, so the 

characters used to separate these genera need to be re-evaluated. Adiplosis is senior to Trisopsis. 

toxicodendri. USA (New York, Connecticut). 

Cecidomyia toxicodendri Felt 1907a: 40. USA: New York, Nassau; HT; ♂; NYSM. 

*ADMAREVERSA 

Admareversa Fedotova & Sidorenko 2009d: 244. Type species, illudia Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Clinodiplosini. This genus is close to Clinodiplosis if the two must be 

separated. 

*illudia. Russia (Far East). 

Admareversa illudia Fedotova & Sidorenko 2009d: 244. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


AFRODIPLOSIS 

Afrodiplosis Felt 1926h: 44. Type species, tarchonanthi Felt (mon.). 

Afrotropical; 1 sp.; Cecidomyiidi, unplaced to tribe. 

tarchonanthi. South Africa. Tarchonanthus camphoratus (Asteraceae). 

Afrodiplosis tarchonanthi Felt 1926h: 44. South Africa: East London; ST; ♂,♀; NYSM. 

*AFROLASIOPTERA 

104

Afrolasioptera Dorchin 2007: 234. Type species, tumida Dorchin (orig. des.). 

Afrotropical; 1 sp.; Lasiopteridi, Lasiopterini. 

*tumida. South Africa. Elytropappus rhinicerotis (Asteraceae). 

Afrolasioptera tumida Dorchin 2007: 237. South Africa: Northern Cape, Kamiesberge, Studers Pass; HT; ♀; SANC. 

*ALATOSTYLA 

Alatostyla Fedotova & Sidorenko 2006b: 2. Type species, velutina Fedotova & Sidorenko (orig. des.). 


*velutina. Russia (Far East). 

Alatostyla velutina Fedotova & Sidorenko 2006b: 3. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; 

♂; ZMAS. 

ALEXOMYIA 

Alexomyia Felt 1921c: 141. Type species, ciliata Felt (orig. des.). 

Neotropical; 1 sp.; Cecidomyiidi, Anadiplosini. 

ciliata. Brazil (Pará). 

Alexomyia ciliata Felt 1921c: 141. Brazil: Pará, Prata; HT; ♀; NYSM. 

ALLIOMYIA 

Allomyia Fedotova 1991d: 21, preocc. Banks 1916. Type species, saurica Fedotova (orig. des.). 

Alliomyia Gagné 2004b: 66, new name for Allomyia Fedotova. 


saurica. Kazakhstan. Allium lineare (Liliaceae). 

Allomyia saurica Fedotova 1991d: 21. Kazakhstan: Saur Mtn Range, Zaïsan, nr Zhanaturmys; HT; ♀; ZMAS. 

ALLOBREMIA 

Allobremia Barnes 1928e: 103. Type species, upolui Barnes (orig. des.). 

Oceanic and Oriental; 2 spp.; Cecidomyiidi, Aphidoletini. The two included species are not necessarily closely 

related. 

pincerifera. India (Uttar Pradesh). 

Allobremia pincerifera Grover & Bakhshi 1978: 83. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

upolui. Samoa, Hawaiian Is. 

Allobremia upolui Barnes 1928e: 104. Samoa: Upolu, Malololelei; HT; ♂; BMNH. 

ALLODIPLOSIS 

Allodiplosis Kieffer & Jörgensen 1910: 389. Type species, crassa Kieffer & Jörgensen. 

105

Neotropical; 1 sp.; Cecidomyiidi, Lopesiini. 

crassa. Argentina (Mendoza). Geoffraea decorticans (Fabaceae). 

Allodiplosis crassa Kieffer & Jörgensen 1910: 389. Argentina: Mendoza; ST; ♂,♀,L,egg; lost. 

ALYCAULUS 

Alycaulus Rübsaamen 1916a: 476. Type species, mikaniae Rübsaamen (mon.). 

Neotropical; 3 spp., all from Mikania; Lasiopteridi, Alycaulini. 

globulus. Brazil (Rio de Janeiro). Mikania glomerata (Asteraceae). 

Alycaulus globulus Gagné 2001: 113. Brazil: Rio de Janeiro, Poço das Antas; HT; ♂; QBUM. 

mikaniae. Brazil (Amazonas). Mikania sp. (Asteraceae). 

Alycaulus mikaniae Rübsaamen 1916a: 476. Brazil: Amazonas, Auristella on Rio Acre; ST; ♂,♀,P,L; ZMHB. 

trilobatus. El Salvador, Colombia. Mikania micrantha; M. cordifolia (Asteraceae). 

Alycaulus trilobatus Möhn 1964b: 583. El Salvador: Cuscatlán, NE Cojutepeque; HT; LT; SMNS. 

AMELIOMYIA 

Ameliomyia Möhn 1960c: 237. Type species, zexmeniae Möhn (orig. des.). 

Neotropical; 1 sp.; Cecidomyiidi, Asphondyliini, Schizomyiina. 

zexmeniae. El Salvador. Zexmenia frutescens (Asteraceae). 

Ameliomyia zexmeniae Möhn 1960c: 237. El Salvador: Santa Ana, S Candelaria; HT; ♀; SMNS. 

AMERHAPHA 

Amerhapha Rübsaamen 1914: 99. Type species, gracilis Rübsaamen (orig. des.). 

Amerapha Rübsaamen & Hedicke 1938a: 285, missp. of Amerhapha. 

Western Palearctic; 1 sp.; Lasiopteridi, unplaced to tribe. 

gracilis. UK, Germany, Czechoslovakia. Daucus carota and possibly other Apiaceae, from galls of Kiefferia 

pericarpiicola (Cecidomyiidae). 

Amerhapha gracilis Rübsaamen 1914: 100. Germany; T; ♂; ZMHB. 

AMETRODIPLOSIS 

Ametrodiplosis Rübsaamen 1910d: 289. Type species, Clinodiplosis thalictricola Rübsaamen (mon.). 

Loewodiplosis (as Löwodiplosis) Kieffer 1912h: 1. Type species, Diplosis auripes Loew (orig. des.). 

Cyrtodiplosis Kieffer 1912h: 1. Type species, Clinodiplosis crassinerva Kieffer (orig. des). 

Nouryodiplosis Tavares 1930: 156. Type species, duclosii Tavares (mon.). 

Ameterodiplosis Grover 1970b: 66, missp. of Ametrodiplosis. 

Palearctic; 20 spp.; Cecidomyiidi, Clinodiplosini. Refs: Möhn 1955a (larva), Harris 1966 (illus.). 

acutissima. Korea, Japan (Honshu to Kyushu). Quercus acutissima (Fagaceae). 

Loewodiplosis acutissima Monzen 1937: 181. Japan: Ishikawa, Shizuoka, Aomori Prefectures; ST; ♂,♀; KUEC. 

auripes. UK, Germany, Poland, Hungary, Yugoslavia. Galium mollugo (Rubiaceae). 

106

107 

Diplosis auripes Löw 1888: 233. Germany: Stuttgart; ST; ♂,♀,P,L; NHMW. 

*campanulae. Russia (Far East). Campanula cephalotes (Campanulaceae). 

Ametrodiplosis campanulae Fedotova 2003c: 2. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

crassinerva. UK, Netherlands, France. Stachys silvatica (Lamiaceae). 

Clinodiplosis crassinerva Kieffer 1901d: 172. France: Moselle, betw. Stürzelbronn & Erbsenthal; T; ♂; lost?. 

duclosii. France, Germany. Stellaria graminea (Caryophyllaceae). 

Nouryodiplosis duclosii Tavares 1930: 157. France: Seine Maritime, St. Saëns; ST; ♂,♀,P,L; TAVC. 

dulichii. USA (Massachusetts). Dulichium arundinaceum (Cyperaceae). 

Thecodiplosis dulichii Felt 1912l: 241. USA: Massachusetts, Magnolia; ST; ♂,♀,L; NYSM. 

*lysimachiae. Russia (Far East). Lysimachia davurica (Primulaceae). 

Ametrodiplosis lysimachiae Fedotova 2003c: 4. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*maackiae. Russia (Far East). Maackia amurensis (Fabaceae). 

Ametrodiplosis maackiae Fedotova 2002c: 30. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

mamajevi. Russia (Far East). Abies holophylla (Pinaceae). 

Ametrodiplosis mamajevi Kovalev 1972: 423. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

medialis. Russia (Europe). Crataegus oxyacantha (Rosaceae). 

Ametrodiplosis medialis Mamaev 1961e: 384. Russia: Krasnodarskiï Terr., Kudepsta; HT; ♂; HYNM. 

nivea. Portugal, Spain. Prunella vulgaris (Lamiaceae). 

Ametrodiplosis nivea Tavares 1916b: 97. Spain: Gallicia, Tuy, & Santiago; ST; ♂,♀,L; TAVC. 

pareirae. India (Uttar Pradesh). Cissampelos pareira (Menispermaceae). 

Ameterodiplosis pareirae Grover 1970b: 66. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

populicola. Russia (Far East). Populus maximoviczii (Salicaceae). 

Ametrodiplosis populicola Kovalev 1972: 425. Russia: Primorskiï Terr., Suputinskiy Res.; HT; ♂; ZMAS. 

*primorskii. Russia (Far East). Vicia sorbifolia (Fabaceae) 

Ametrodiplosis primorskii Fedotova & Sidorenko 2004a: 11. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*sorbariaflora. Russia (Far East). Sorbaria sorbifolia (Rosaceae) 

Ametrodiplosis sorbariaflora Fedotova & Sidorenko 2004a: 10. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*spiraeaflorae. Russia (Far East). Spiraea salicifolia (Rosaceae) 

Ametrodiplosis spiraeaflorae Fedotova 2004e: 10. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

thalictricola. Widespread Europe. Thalictrum flavum (Ranunculaceae). 

Clinodiplosis thalictricola Rübsaamen 1895c: 257. Germany; ST; ♂,L; ZMHB. 

urticae. Yugoslavia, Russia (Far East). Urtica angustifolia (Urticaceae). 

Ametrodiplosis urticae Kovalev 1972: 426. Russia: Primorskiï Terr., Suputinskiy Res.; HT; ♂; ZMAS. 

*veronicastrum. Russia (Far East). Veronicastrum sibiricum (Scrophulariaceae). 

Ametrodiplosis veronicastrum Fedotova 2003c: 6. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

viciae. Latvia, Russia (Europe). Vicia cracca (Fabaceae). 

Ametrodiplosis viciae Mamaev 1961e: 386. Russia: Moscow Prov., Sofrino; HT; ♂; HYNM. 

AMPELOSUCTA 

Ampelosucta De Stefani 1912: 7 (publ'n not seen). Type species, illata De Stefani (mon.). 

Italy; 1 sp.; Cecidomyiidi, unplaced to tribe. De Stefani 1912, which I could not locate, was first cited in Kieffer 

1913i; all subsequent citations, including Neave 1939a, may be secondarily from Kieffer's citation.

illata. Italy. Vitis vinifera (Vitaceae). 

Ampelosucta illata De Stefani 1912: 7 (publ'n not seen). Italy: Sicily; ST; ♂,♀,P,L; lost. 

ANABREMIA 

Anabremia Kieffer 1912h: 2. Type species, Macrodiplosis bellevoyei Kieffer (orig. des.). 

Western Palearctic; 6 spp., all inquilines or possibly inquilines in galls of Dasineura spp. on Fabaceae; Cecidomyiidi, 

unplaced to tribe. Ref.: Solinas 1965b (illus.). 

bellevoyei. UK, France, Germany, Poland, Czechoslovakia. Lathyrus pratensis (Fabaceae). 

Macrodiplosis bellevoyei Kieffer 1896j: 384. France: château de La Grange, nr Thionville; T; A; lost?. 

inquilina. Italy. Medicago sativa (Fabaceae) in galls of Jaapiella medicaginis. 

Anabremia inquilina Solinas 1965b: 253. Italy; ST; ♂,♀,P,L; Solinas Coll. 

massalongoi. Italy. Vicia villosa, V. dasycarpa (Fabaceae). 

Clinodiplosis massalongoi Kieffer 1909b: 34. Italy; ST; LT; lost?. 

medicaginis. Czechoslovakia, Croatia. Medicago orbiculare, M. hispida, M. falcata (Fabaceae). 

Anabremia medicaginis Rübsaamen 1917: 65. Croatia: nr Split; ST; ♀,L; ZMHB. 

trotteri. Italy. Vicia lutea (Fabaceae). 

Clinodiplosis trotteri Kieffer 1909b: 32. Italy: Verona; T; LT; lost?. Italy. 

viciae. UK, France, Poland, Czechoslovakia, Romania, Russia (Western Siberia). Vicia lutea (Fabaceae). 

Anabremia viciae Kieffer 1913i: 169. France: château de La Grange, nr Thionville; T; LT; lost?. 

ANADIPLOSIS 

Anadiplosis Tavares 1916a: 41. Type species, pulchra Tavares (orig. des.). 

Neotropical; 4 spp., on Fabaceae; Cecidomyiidi, Anadiplosini. 

caetetensis. Brazil (Bahia). Undet. Fabaceae. 

Anadiplosis caetetensis Tavares 1920e: 70. Brazil: Bahia, Caeteté; ST; P; TAVC. 

procera. Brazil (Bahia). Undet. sp. of mimosoid Fabaceae. 

Anadiplosis procera Tavares 1920e: 66. Brazil: Bahia, Salvador, Itaparica; ST; ♂,♀,P; TAVC. 

pulchra. Brazil (Rio de Janeiro). Machaerium sp. (Fabaceae). 

Anadiplosis pulchra Tavares 1916a: 42. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♂,♀,egg; TAVC. 

venusta. Brazil (Rio de Janeiro). Machaerium sp. (Fabaceae). 

Anadiplosis venusta Tavares 1916a: 46. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♂,♀,L; TAVC. 

ANASPHODIPLOSIS 

Anasphondylia Blanchard 1939: 173, preocc. Tavares 1920. Type species, aspidospermae Blanchard (orig. des.). 

Anasphodiplosis Gagné 1994a: 172, new name for Anasphondylia Blanchard. 

Neotropical; 1 sp.; Cecidomyiidi, unplaced to tribe. 

aspidospermae. Argentina (Santa Fé). Aspidosperma quebracho-blanco (Apocynaceae). 

Anasphondylia aspidospermae Blanchard 1939: 173. Argentina: Santa Fé, San Cristóbal; ST; ♀; DPBA. 

ANASPHONDYLIA 

Anasphondylia Tavares 1920a: 38. Type species, myrtacea Tavares (orig. des.). 

108


myrtacea. Brazil (Rio de Janeiro). Undet. sp. of Myrtaceae. 

Anasphondylia myrtacea Tavares 1920f: 125, pl. III. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♀,P; TAVC. 

ANDIRODIPLOSIS 

Andirodiplosis Tavares 1920d: 93. Type species, bahiensis Tavares (mon.). 


bahiensis. Brazil (Bahia). Andira sp. (Fabaceae). 

Andirodiplosis bahiensis Tavares 1920d: 94. Brazil: Bahia, Salvador; ST; ♂,♀,P; TAVC. 

ANDRODIPLOSIS 

Androdiplosis Felt 1915g: 182. Type species, coccidivora Felt (orig. des.). 

Oriental; 1 sp.; Cecidomyiidi, Lestodiplosini. Distinct only in the android female antennae, the one included species 

could belong to Lestodiplosis or Dentifibula. Ref.: Harris 1968 (rev.). 

coccidivora. Sri Lanka. Aspidiotus sp. or Chrysomphalus sp. (Hemiptera: Diaspididae). 

Androdiplosis coccidivora Felt 1915g: 183. Sri Lanka: Peradeniya, Botanic Gardens; HT; ♀; lost. 

ANDROSACEMYIA 

Androsacemyia Fedotova 1992h: 18. Type species, alatavica Fedotova (orig. des.). 


alatavica. Kazakhstan. Androsace lehmanniana (Primulaceae). 

Androsacemyia alatavica Fedotova 1992h: 19. Kazakhstan: Kungeï Alatau, Karabulak canyon, 12 km SW Malyï 

Zhalanash; HT; ♀; ZMAS. 

ANGEIOMYIA 

Angeiomyia Kieffer & Herbst 1909: 124. Type species, spinulosa Kieffer & Herbst (orig. des. as n. g., n. sp.). 

Neotropical; 1 sp.; Lasiopteridi, unplaced to tribe. 

spinulosa. Chile. Hydrangea scandens (Hydrangeaceae). 

Angeiomyia spinulosa Kieffer & Herbst 1909: 124. Chile: Concepción; T; LT; lost. 

*ANISODIPLOSIS 

Anisodiplosis Maia 2005c: 28. Type species, waltheriae Maia (orig. des.). 


†waltheriae. Brazil. Waltheria indica (Sterculiaceae). 

Anisodiplosis waltheriae Maia 2005c: 29. Brazil: Minas Gerais, Aimorés; HT; ♂; MNRJ. Ref.: Almeida et al. 2006 

(ecol., biol.). 

109

ANISOSTEPHUS 

Anisostephus Rübsaamen 1917: 87. Type species, Diplosis betulina Kieffer (orig. des.). 

Palearctic; 2 spp., from leaf galls of Betula; Cecidomyiidi, Cecidomyiini. Ref.: Möhn 1955a (larva). 

betulinus. Widespread Europe; Russia (Far East). Betula pubescens (Betulaceae). 

Diplosis betulina Kieffer 1889a: 153. France: Bitche & Karlingen; ST; ♀,L; lost?. 

hydrophilus. Russia (Far East). Betula mandshurica (Betulaceae). 

Anisostephus hydrophilus, as ssp. of betulinus Kieffer, Kovalev 1972: 422. Russia: Primorskiï Terr., Tudagou R. 

valley, S Vinogradovka; HT; ♀; ZMAS. 

ANTHODIPLOSIS 

Anthodiplosis Kieffer 1912h: 1. Type species, Clinodiplosis rudimentalis Kieffer (orig. des.). 

Western Palearctic; 1 sp.; Cecidomyiidi, Clinodiplosini. Ref.: Stelter 1989a (redescr.). 

rudimentalis. France, Czechoslovakia, Romania, Yugoslavia. Artemisia vulgaris (Asteraceae). 

Clinodiplosis rudimentalis Kieffer 1901d: 172. France: vic. Guinkirchen; T; ♂; lost?. 

ANTICHIRIDIUM 

Antichira Rübsaamen 1910d: 285, preocc. Agassiz & Loew 1846. Type species, striata Rübsaamen (orig. des. as n. 

g., n. sp.). 

Antichiridium Rübsaamen 1911e: 168, new name for Antichira Rübsaamen. 

Trichodiplosis Kieffer 1913a: 229. Type species, Clinodiplosis caricis Kieffer (orig. des.). 

Palearctic; 2 spp.; Cecidomyiidi, unplaced to tribe. 

caricis. UK, France, Romania. Carex stellulata (Cyperaceae). 

Clinodiplosis caricis Kieffer 1898f: 61. France: Moselle; T; A; lost?. 

striatum. UK, Germany, Czechoslovakia. Molinia caerulea (Poaceae). 

Antichira striata Rübsaamen 1910d: 285. Germany; T; A; ZMHB. 

*ANULIDIPLOSIS 

Anulidiplosis Fedotova 2004h: 175. Type species, trilicis Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, tribe Aphidoletini. This genus is close to Bremia if the two must be separated. 

*trilicis. Russia (Far East). 

Anulidiplosis trilicis Fedotova 2004h: 175. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

APAGODIPLOSIS 

Apagodiplosis Gagné 1973c: 862. Type species, Oligotrophus papyriferae Gagné (orig. des.). 

Nearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. 

110

papyriferae. Canada (Quebec), USA (Washington, Oregon, Michigan). Betula papyrifera (Betulaceae). 

Oligotrophus papyriferae Gagné 1967a: 132. USA: Michigan, Sanilac Co.; HT; ♂; USNM. 

APHIDOLETES 

Aphidoletes Kieffer 1904c: 385. Type species, Bremia abietis Kieffer (subs. des. Felt 1911: 53). 

Cryptobremia Kieffer 1912h: 1. Type species, aegyptiaca Kieffer (orig. des.) = Aphidoletes aphidimyza Rondani. 

Phaenobremia Kieffer 1912h: 1. Type species, Bremia urticariae Kieffer (orig. des.). 

Isobremia Kieffer 1913a: 231. Type species, Bremia sonchi Kieffer (orig. des.) = Aphidoletes aphidimyza Rondani. 

Phenobremia Del Guercio 1918a: 182, missp. of Phaenobremia. 

Rondaniella Del Guercio 1918a: 187, preocc. Johannsen 1909. Type species, bezzii Del Guercio (orig. des. as n. g., n. 

sp.) = Aphidoletes aphidimyza Rondani. 

Guerciobremia Barnes 1927f: 221, new name for Rondaniella Del Guercio. 

Phaerobremia Shinji 1944: 236, missp. of Phaenobremia. 

Aphdoletes Mani 1946: 216, missp. of Aphidoletes. 

Cosmopolitan; 4 spp., their larvae all predators of aphidoids; Cecidomyiidi, Aphidoletini. Refs: Harris 1966 (illus.), 

1973 (taxonomy, list of prey), Solinas 1968 (anat., larva). 

abietis. UK, France, Germany, Switzerland, Latvia, Poland, Czechoslovakia, Russia (Europe), Canada (British 

Columbia, Alberta, Ontario, Newfoundland), USA (Maine, Virginia, North Carolina). Adelges abietis; A. 

piceae, A. nusslini (Hemiptera: Adelgidae). 

Bremia abietis Kieffer 1896j: 382. France: Bitche; ST; A; lost?. See note under Aphidoletes thompsoni. 

aphidimyza. Widespread Europe, Turkey, Israel, Egypt, Russia (Far East), Japan (Hokkaido to Kyushu), Hawaiian Is, 

widespread Nearctic, Brazil, Chile, New Zealand. Aphis sp.; many genera of aphids (Hemiptera: Aphididae), 

cf. Harris 1973. 

Cecidomya aphidimyza Rondani 1847: 443. Italy; ST; ♂,♀,L; MZUF. 

Cecidomyia cerasi Loew 1850a: 34. Poland; ST; ♂,♀; ZMHB?. 

Cecidomyia napi Kaltenbach 1858: 145. Germany; ST; LT; NHMW?. 

Diplosis aphidisuga Rübsaamen 1891a: 8. Germany; ST; ♂,♀,L; ZMHB. 

Diplosis aphidivora Rübsaamen 1891a: 14. Germany; ST; ♂,♀,P,L; ZMHB. 

Bremia sonchi Kieffer 1896j: 382. France: château de La Grange, nr Thionville; ST; A; lost?. 

Diplosis cucumeris Lintner 1897: 166. USA: Massachusetts, Lowell; ST; ♂,♀; NYSM. 

Diplosis rosivora Coquillett 1900a: 46. USA: District of Columbia; LT (Gagné 1971c: 178); ♂; USNM. 

Bremia impatientis Kieffer 1901d: 169. France: Bitche; ST; ♂,♀; lost?. 

Bremia hamamelidis Felt 1907a: 29. USA: New York, Albany; HT; ♂; NYSM. 

Aphidoletes carnifex Kieffer 1908: 200. Russia: Astrakhan; ST; ♂,♀,L; lost?. 

Aphidoletes basalis Felt 1908b: 397. USA: New York, Albany; HT; ♂; NYSM. 

Aphidoletes borealis Felt 1908b: 397. USA: New York, Albany; LT (Gagné 1971c: 178); ♂; NYSM. 

Aphidoletes flavida Felt 1908b: 397. USA: New York, Albany; HT; ♂; NYSM. 

Aphidoletes fulva Felt 1908b: 397. USA: New York, Albany; HT; ♂; NYSM. 

Aphidoletes marginata Felt 1908b: 397. USA: New York, Nassau; HT; ♂; NYSM. 

Aphidoletes marina Felt 1908b: 397. USA: New York, New York, Staten I.; HT; ♂; NYSM. 

Aphidoletes meridionalis Felt 1908b: 397. USA: District of Columbia; HT; ♂; USNM. 

Cryptobremia aegyptiaca Kieffer 1912h: 1. Egypt; T; A; lost?. 

Phaenobremia cardui Kieffer 1913g: 50. France: Bitche; ST; ♂,P,L; lost?. 

Phaenobremia vacunae Kieffer 1913g: 51. France: Bitche; T; LT; lost?. 

Rondaniella bezzii Del Guercio 1918a: 187. Italy; ST; ♂,♀,P,L; lost. 

Rondaniella cucullata Del Guercio 1918a: 218. Italy; T; LT; lost. 

Rondaniella macrosiphoniellae Del Guercio 1918a: 213. Italy; T; LT; lost. 

Rondaniella macrosiphonis Del Guercio 1918a: 210. Italy; T; LT; lost. 

Rondaniella ornata Del Guercio 1918a: 215. Italy; T; LT; lost. 

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112 

Rondaniella phorodontis Del Guercio 1918a: 208. Italy; T; LT; lost. 

Rondaniella toxopterae Del Guercio 1918a: 203. Italy: Florence; T; LT; lost. 

Phaenobremia kiefferiana Tölg 1921: 33. Austria: vic. Vienna; ST; ♂,♀; lost. 

Isobremia kiefferi Voukassovitch 1925: 357. Serbia and Montenegro: Belgrade; ST; P,L,egg; ?. 

Phaenobremia helichrysis Barnes 1927d: 176. Ireland: Co. Dublin; ST; ♂,♀; BMNH. 

Phaenobremia aphidiphora Nijveldt 1953a: 193, missp. of aphidivora. 

Tribremia aphidophaga Marikovskij 1956: 186. Georgia: Batumi; ST; ♂,♀; ZMAS. 

Phaenobremia doutti Pritchard 1961b: 100. USA: CA, Albany; HT; ♂; USNM. 

Aphidoletes aegypta Skuhravá 1986: 224, missp. of aegyptiaca. 

Aphidoletes aphidosuga Skuhravá 1886: 224, missp. of aphidisuga. 

thompsoni. UK, Germany, Switzerland, Czechoslovakia; introduced into Canada & USA, established?. Adelges piceae 

(Hemiptera: Adelgidae). 

Aphidoletes thompsoni Möhn 1954: 462. Germany: Laimbach, nr Munich; HT; ♂; SMNS. This species was 

reportedly introduced into Canada in the 1950s and from there into the USA (Dowden 1962), but could have 

been confused in Europe and in North America with the widespread A. abietis, a species perhaps native to 

North America. Aphidoletes thompsoni and A. abietis are similar, possibly identical, species presently being 

evaluated by K.M. Harris (in litt.). Ref.: Harris & Bennett 2003. 

urticaria. Widespread Europe, USA (Washington, Colorado, New York, Massachusetts, Virginia). Aphis urticata; 

many genera of aphids (Hemiptera: Aphididae), cf. Harris 1973. 

Bremia urticaria Kieffer 1895p: 9. France; NT (Harris 1973: 310; Netherlands: Amsterdam); ♂; BMNH. 

Bremia urticariae Kieffer 1901a: 333, missp. of urticaria. 

Aphidoletes recurvata Felt 1908b: 397. USA: Massachusetts, Brookline; HT; ♂; NYSM. 

Phaenobremia macrorosae Barnes 1927d: 177. UK: Scotland, St. Andrew's; ST; ♂,♀; BMNH. 

APIOMYIA 

Apiomyia Kieffer 1913a: 220. Type species, Hormomyia bergenstammi Wachtl (orig. des.). 

Palearctic; 1 sp.; Lasiopteridi, unplaced to tribe. 

bergenstammi. Widespread Europe, Middle East. Pyrus salicifolia; P. communis (Rosaceae). 

Hormomyia bergenstammi Wachtl 1882: 289. Greece: Corfu; ST; ♂,♀; NHMW?. 

APLECUS 

Aplecus Kieffer 1913h: 105. Type species, validus Kieffer (orig. des.). 

Aplecs Shinji 1944: 250, missp. of Aplecus. 


validus. Tanzania. 

Aplecus validus Kieffer 1913h: 105. Tanzania: "Neu-Moschi", lower slopes Mt. Kilimanjaro; HT; ♀; MNHN. 

APLONYX 

Aplonyx De Stefani 1908b: 174. Type species, chenopodii De Stefani (mon). 

Palearctic; 1 sp.; Lasiopteridi, Lasiopterini. The single species was only superficially described and apparently has not 

been rediscovered since. 

chenopodii. Italy. Atriplex patula, originally, but subsequently as Chenopodium album (ref.: Skuhravá, M., V. 

Skuhravý and B. Massa, 2007) (Chenopodiaceae).

Aplonyx chenopodii De Stefani 1908b: 174. Italy: Sicily, Palermo; ST; ♂,♀,P,L,egg; lost. 

APOASPHONDYLIA 

Diceromyia Felt 1918c: 283, preocc. Theobald 1911. Type species, vernoniae Felt (orig. des.). 

Apoasphondylia Gagné 1973a: 493, new name for Diceromyia Felt. 

Oriental; 2 spp.; Cecidomyiidi, Asphondyliini, Asphondyliina. 

orientalis. Indonesia (Sumatra). Phoebe declinata (Lauraceae). 

Diceromyia orientalis Felt 1927d: 388. Indonesia: Sumatra, nr Sibolangit; HT; ♂; NYSM. 

vernoniae. Philippines. Vernonia lancifolia (Asteraceae). 

Diceromyia vernoniae Felt 1918c: 284. Philippines: Luzon, Laguna Prov., Mt Makiling; ST; ♂; NYSM. 

APODIPLOSIS 

Apodiplosis Tavares 1922a: 37. Type species, praecox Tavares (orig. des.). 


praecox. Brazil (Rio de Janeiro). Psychotria sp. (Rubiaceae). 

Apodiplosis praecox Tavares 1922a: 35. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♀,P; TAVC. 

APOSCHIZOMYIA 

Aposchizomyia Gagné 1993c: 100. Type species, acuta Gagné (orig. des.). 

Afrotropical; 6 spp., from bud and leaf galls of Acacia; Cecidomyiidi, Asphondyliini, Schizomyiina. 

acuta. Kenya. Acacia nilotica (Fabaceae). 

Aposchizomyia acuta Gagné 1993c: 104. Kenya: Rt. A109, Kapiti Plains, Malili Ranch, ca. 70 km SE Nairobi; HT; 


brevis. Kenya. Acacia drepanolobium (Fabaceae). 

Aposchizomyia brevis Gagné 1993c: 104. Kenya: southern base Ngong Hills; HT; P; USNM. 

crenata. Kenya. Acacia drepanolobium (Fabaceae). 

Aposchizomyia crenata Gagné 1993c: 104. Kenya: Rt. A109, Kapiti Plains, Malili Ranch, ca. 70 km SE Nairobi; HT; 

P; USNM. 

longa. Kenya. Acacia drepanolobium (Fabaceae). 

Aposchizomyia longa Gagné 1993c: 106. Kenya: Rt. A109, Kapiti Plains, Malili Ranch, ca. 70 km SE Nairobi; HT; 

P; USNM. 

striata. Kenya. Acacia tortilis (Fabaceae). 

Aposchizomyia striata Gagné 1993c: 106. Kenya: Yatta; HT; P; USNM. 

turnouri. Kenya. Acacia drepanolobium (Fabaceae). 

Aposchizomyia turnouri Gagné 1993c: 107. Kenya: Rt. A109, Kapiti Plains, Malili Ranch, ca. 70 km SE Nairobi; HT; 

♀; USNM. 

*AEQUIDIPLOSIS 

Aequidiplosis Fedotova 2004h: 167. Type species, cornuta Fedotova (orig. des.). 

Aquidiplosis Fedotova 2006a: 154, misspelling of Aequidiplosis. 

Eastern Palearctic; 3 spp.; Cecidomyiidi, Mycodiplosini. This genus is close to Mycodiplosis if the two must be 

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separated. 

*ampla. Russia (Far East). 

Aquidiplosis ampla Fedotova 2006a: 154. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

*cornuta. Russia (Far East). 

Aequidiplosis cornuta Fedotova 2004h: 168. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*rara. Russia (Far East). 

Aequidiplosis rara Fedotova 2004h: 169. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

ARCIVENA 

Arcivena Gagné 1984c: 129. Type species, kielmeyerae Gagné (orig. des.). 

Neotropical; 1 sp.; Lasiopteridi, Dasineurini. 

kielmeyerae. Brazil (São Paulo). Kielmeyera rosea (Clusiaceae). 

Arcivena kielmeyerae (also as keilmeyerae) Gagné 1984c: 130. Brazil: São Paulo, Mogi-Guaçu; HT; ♂; MZSP. 

ARNOLDIOLA 

Arnoldia Kieffer 1895b: 7, preocc. Mayer 1887. Type species, Cecidomyia quercus Binnie (subs. des. Coquillett 

1910: 510). 

Arnoldiola Strand 1928: 48, new name for Arnoldia Kieffer. 

Arnoldiana Anonymous 1929: 322, missp. of Arnoldiola. 

Holarctic; 15 spp., most associated with Quercus and Castanea; Lasiopteridi, Dasineurini. The European species on 

Rosa, Pentaphylloides, and Sambucus possibly do not belong here. Ref.: Gagné 2009a (rev. Nearctic spp.). 

atra. USA (Texas). Quercus virginiana (Fagaceae). 

Arnoldiola atra Gagné 1999d: 160. USA: Texas, Houston; HT; ♂; USNM. 

azaleae. USA (New York). 

Oligotrophus azaleae Felt 1907a: 26. USA: New York, Albany; HT; ♂; NYSM. Ref.: Gagné 2009a: 108 (redescr.). 

Phytophaga caudata Felt 1915j: 199. USA: New York, Albany; ST; ♂; NYSM. 

baldratii. Italy. Quercus ostryaefolia (Fagaceae). 

Arnoldia baldratii Kieffer 1909b: 23. Italy; T; G; lost?. 

brevicornis. USA (New York). 

Oligotrophus brevicornis Felt 1907a: 25. USA: New York, Nassau; HT; ♂; NYSM. Ref.: Gagné 2009: 109 

(redescr.). 

Oligotrophus tiliacei Felt 1907a: 25. USA: New York, Westfield; HT; ♂; NYSM. 

Oligotrophus tiliaceus Felt 1907c: 121, unjust. emend. of tiliacei. 

castaneae. USA (Massachusetts). Castanea dentata (Fagaceae). 

Rhopalomyia castaneae Felt 1909b: 291. USA: Massachusetts, Stowe; HT; ♀; NYSM. Ref.: Gagné 2009: 109 

(redescr.). 

Janetiella ligni Felt 1915j: 220. USA: New York, Nassau; HT; ♀; NYSM. 

dryophila. France, Germany. Quercus cerris (Fagaceae). 

Arnoldia dryophila Kieffer 1909b: 21. France; ST; LT; lost?. 

gemmae. UK, Sweden, Germany, Czechoslovakia. Quercus sp.; Q. petraea. (Fagaceae). 

Cecidomyia gemmae Giraud 1868: cxii. France; ST; G; MNHN. 

Cecidomyia gemmae Rübsaamen 1892b: 401, jun. prim. hom. of gemmae Giraud. Germany: vic. Berlin; ST; ♀; 

ZMHB. As new synonym in Gagné 2004b: 71, to avoid coining a new name for gemmae Rübsaamen. 

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115 

libera. Widespread Europe. Quercus petraea, Q. robur (Fagaceae). 

Perrisia libera Kieffer 1909b: 21. France; T; G; lost?. 

margaritae. Poland. 

Arnoldiola margaritae Szadziewski 1975: 365. Poland: Toru� Distr., Las Piwnicki; HT; ♂; SZAC. 

marikovskii. Kazakhstan. Rosa nanothamnus; R. platyacantha (Rosaceae). 

Arnoldiola marikovskii Fedotova 1987d: 33. Kazakhstan: Tarbagataï Mtn Range, 59 km NW Urdzhara; HT; ♂; 

ZKAS. 

pentaphylloidis. Kazakhstan. Pentaphylloides fruticosa (Rosaceae). 

Arnoldiola pentaphylloidis Fedotova 1990b: 101. Kazakhstan: southern Altaï, northeastern shore L. Markakol', 10 km 

N Urunkhaïka; HT; ♂; ZKAS. 

quercus. Widespread Europe. Quercus sp.; Q. petraea, Q. robur (Fagaceae). 

Cecidomyia quercus Binnie 1877: 179. UK: Scotland, Cambuslang; ST; ♂,♀,L; ?. This name is a jun. prim. hom. of 

Cecidomyia quercus Loew, now in Macrolabis. According to Article 23.9.5 of the International Code of 

Zoological Nomenclature 1999, when two primary homonyms are both in use in separate genera, the case is to 

be referred to the Commission for a ruling under the plenary power and that in the meanwhile prevailing usage 

of both names is to be maintained. 

Arnoldia quercicola Kieffer 1909b: 21. France; ST; LT; lost?. 

Arnoldia quercina Tavares 1920c: 53. Spain: La Guardia (Pontevedra); ST; ♀; TAVC. 

sambuci. UK, France, Germany, Italy, Russia (Europe). Sambucus nigra (Caprifoliaceae). 

Arnoldia sambuci Kieffer 1901c: 21. France: Bitche; ST; ♂,♀; lost?. 

trotteri. Italy. Quercus cerris (Fagaceae). 

Arnoldia trotteri Kieffer 1909b: 22. Italy; T; LT; lost?. 

tympanifex. Italy, Greece. Quercus ilex, Q. suber (Fagaceae). 

Arnoldia tympanifex Kieffer 1909b: 22. Italy: Sicily; T; G; lost?. 

ARRABIADAEAMYIA 

Arrabiadaeamyia Maia 2001a: 2. Type species, serrata Maia (orig. des.). 


serrata. Brazil (Rio de Janeiro). Arrabidaea conjugata (Bignoniaceae). 

Arrabiadaeamyia serrata Maia 2001a: 2. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

ARTEMISIOMYIA 

Artemisiomyia Kovalev 1972: 412. Type species, Asteromyia gorovoji Kovalev (orig. des.). 

Eastern Palearctic; 1 sp.; Lasiopteridi, Lasiopterini. 

gorovoji. Russia (Far East). Artemisia sp. (Asteraceae). 

Asteromyia gorovoji Kovalev 1967: 98. Russia: Primorskiï Terr., Khasan; HT; ♂; KOVC. 

ARTHROCNODAX 

Arthrocnodax Rübsaamen 1895a: 189. Type species, vitis Rübsaamen (subs. des. Coquillett 1910: 510). 

Feltodiplosis Kieffer 1913i: 154. Type species, Endaphis americana (subs. des. Gagné 2004b: 72), one of the three of 

the originally included spp. 

Feltdiplosis Del Guercio 1918a: 182, missp. of Feltodiplosis. 

Asyomyia Marikovskij 1953a: 132. Type species, humilis Marikovskij (orig. des.). 

Cosmopolitan; 49 spp.; Cecidomyiidi, Lestodiplosini. Almost all of these species have been reared in association with

or are known predators of Eriophyidae, but three have been associated only with other Cecidomyiidae. 

abdominalis. Peru. Undet. sp. of Acarina. 

Endaphis abdominalis Felt 1911e: 128. Peru: Piura; ST; ♂,♀; USNM. 

alatavicus. Kazakhstan. Acarina on Artemisia absinthium (Asteraceae). 

Arthrocnodax alatavicus Fedotova 2001h: 1485. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak Canyon, 20 km SE Alma- 

Ata; HT; ♂; ZMAS. 

alyssiiphilus. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax alyssiiphilus Fedotova 1994a: 128, nomen nudum. 

Arthrocnodax alyssiiphilus Fedotova 1994b: 80. Kazakhstan: 28 km W Urdzhara; HT; ♂; ZMAS. 

americanus. USA (Arizona). Eriophyes fraxiniflora (Acarina: Eriophyidae). 

Endaphis americana Felt 1911e: 129. USA: Arizona, Prescott; ST; ♀; NYSM. 

annulatus. USA (New York). 

Rhabdophaga annulata Felt 1907a: 17. USA: New York, Albany; HT; ♀; NYSM. 

asiaticus. Kazakhstan. Undet. sp. of Acarina. 

Arthrocnodax asiaticus Fedotova 1998a: 29. Kazakhstan: Azutau Mtn Range, 20 km SE Alekseevka; HT; ♂; ZMAS. 

berteroaphilus. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax berteroaphilus Fedotova 1994b: 79. Kazakhstan: Tarbagataï Mtn Range, 30 km E Tarbagataï; HT; ♂; 

ZMAS. 

bromiphilus. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax bromiphilus Fedotova 1997a: 45. Kazakhstan: Dzhungarskiï Alatau, 13 km E Sarkand; HT; ♂; ZMAS. 

campanulae. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax campanulae Fedotova 1995a: 14. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 18 km SE 

Novonikolaevka; HT; ♂; ZMAS. 

chondrillaphilus. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax chondrillaphilus Fedotova 1998a: 33. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 20 km NW 


clematitis. Algeria. Phyllocoptes populi (Acarina: Eriophyidae). 

Arthrocnodax clematitis Marchal 1897a: 23. Algeria: Blidah, Vallon des Singes; ST; ♂,♀; MNHN?. 

Arthrocnodax clematidis Skuhravá 1989: 226, missp. of clematitis. 

coprae. Philippines. 

Arthrocnodax coprae Felt 1919a: 291. Philippines: Luzon, Laguna Prov., Los Baños; ST; ♂,♀; NYSM. 

coryligallarum. Widespread Europe. Phytoptus avellanae (Acarina: Eriophyidae). 

Diplosis coryligallarum Targioni-Tozzetti 1887: 422. Italy: Palermo; ST; ♂,♀,L; ?. 

erianeus. Switzerland. Eriophyes sp. (Acarina: Eriophyidae). 

Cecidomyia erianea Bremi 1847a: 30. Switzerland: vic. Zurich; ST; LT; ETHZ?. 

fagi. France. 

Arthrocnodax fagi Kieffer 1901b: 459. France; T; LT; lost?. 

fragariae. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax fragariae Fedotova 1997a: 44. Kazakhstan: Dzhungarskiï Alatau, nr Lepsinsk; HT; ♂; ZMAS. 

fraxinellus. Ireland, UK. Aceria fraxinivora (Acarina: Eriophyidae). 

Diplosis fraxinella Meade 1888: 77. UK: England; ST; ♂,♀; ?. Ref.: Ashe & O'Connor 2005 (ecol., distr.). 

galiobiae. Kazakhstan. Aceria galiobiae (Acarina: Eriophyidae). 

Arthrocnodax galiobiae Fedotova 1997c: 331. Kazakhstan, northern spurs Dzhungarskiï Alatau, Dzhaman-Kotyr-Tas 

Ridge, nr Topolevka; HT; ♂; ZMAS. 

gemmarum. France. Eriophyes sp. (Acarina: Eriophyidae). 

Arthrocnodax gemmarum Kieffer 1895p: 9. France; ST; ♂,♀; lost?. 

geranii. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax geranii Fedotova 1995a: 13. Kazakhstan: southern Altaï, Markakol'skiï Res., L. Markakol', Topolevka 

peak, nr Urunkhaïka; HT; ♂; ZMAS. 

haloxylonis. Kazakhstan. Aegyptobia traegardhy (Acarina: Tenuipalpidae). 

Arthrocnodax holoxylonis Fedotova 1993e: 89. Kazakhstan: eastern Mangyshlak basin, 65 km SE Novyï-Uzen', 30 

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117 

km SE Senek; HT; ♂; ZMAS. 

hieraciis. Kazakhstan. Undet. sp. of Acarina. 

Arthrocnodax hieraciis Fedotova 1998a: 31. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Ul'ken-Kaïndy 

canyon, 20 km S Novonikolaevka; HT; ♂; ZMAS. 

humilis. Kazakhstan. 

Asyomyia humilis Marikovskij 1953a: 132. Kazakhstan: Alma-Ata; ST; ♂,♀; ZKAS & ZMAS. 

incanus. Germany. Dasineura populeti (Diptera: Cecidomyiidae). 

Diplosis incana Rübsaamen 1890b: 20. Germany: vic. Siegen; ST; ♀,L; ZMHB. 

jaapi. Denmark, Germany. Contarinia tanaceti (Diptera: Cecidomyiidae). 

Arthrocnodax jaapi Rübsaamen 1921: 42. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

lepidiis. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax lepidiis Fedotova 1994b: 81. Kazakhstan: Dzhungarskiï Alatau, 15 km S Pokatilovka; HT; ♂; ZMAS. 

limonii. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax limonii Fedotova 1995a: 11. Kazakhstan: Dzhusan-Dala Desert, 40 km SE Aksuek; HT; ♂; ZMAS. 

lycii. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax lycii Fedotova 1997a: 42. Kazakhstan: right shore Syr-Darya R., 7 km S Terenozek; HT; ♂; ZMAS. 

mali. Germany. Phyllocoptes schlechtendali (Acarina: Eriophyidae). 

Arthrocnodax mali Kieffer 1926: 103. Germany; ST; ♂,♀,P,L; lost?. 

medicaginis. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax medicaginis Fedotova 1995a: 24. Kazakhstan: 101 km SW Ust'-Kamenogorsk, nr Georgievka; HT; ♂; 

ZMAS. 

meridionalis. USA (Florida), St. Vincent, Trinidad, Peru (paralectotypes of Lestodiplosis peruviana). Eriophyes sp. 

(Acarina: Eriophyidae). 

Arthrocnodax meridionalis Felt 1912b: 176. St. Vincent; ST; ♂,♀,L; NYSM. 

minutus. Germany. 

Cecidomyia minuta Winnertz 1853: 250. Germany; T; ♂; lost. 

nepetae. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax nepetae Fedotova 1995a: 22. Kazakhstan: southern Kazakhstan, 12 km S Zhantas, upper Shabykty R.; 



Arthrocnodax orientalis Grover & Bakhshi 1978: 53. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; PGCA. 

origani. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax origani Fedotova 1995a: 17. Kazakhstan: Dzhungarskiï Alatau, Kaïkan Mtn Range, 7 km NE 

Dzerzhinsk; HT; ♂; ZMAS. 

paeoniae. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax paeoniae Fedotova 1997a: 39. Kazakhstan: Dzhungarskiï Alatau, Konyrtau Mtn Range, 10 km W 

Sarkand; HT; ♂; ZMAS. 

peregrinus. Germany. Acarus sp. (Acarina: Acaridae). 

Cecidomyia peregrina Winnertz 1853: 252. Germany; ST; ♂,L; lost. 

reaumuriae. Kazakhstan. Dasineura flosculorum (Diptera: Cecidomyiidae). 

Lestodiplosis reaumuriae Fedotova 1983c: 27. Kazakhstan: Dzhungarian Alatau, mid course Ili R., nr Poyushchei 

Mts; HT; ♂; ZMAS. 

rhoinus. USA (New York). 

Arthrocnodax rhoina Felt 1908b: 404. USA: New York, Albany; HT; ♂; NYSM. 

Arthrocnodax sambucifolia Felt 1908b: 404. USA: New York, Albany; HT; ♂; NYSM. 

rodiolae. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax rodiolae Fedotova 1995a: 24. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Ul'ken-Kaïndy 

pass; HT; ♂; ZMAS. 

sajramicus. Kazakhstan. Undet. sp. of Acarina. 

Arthrocnodax sajramicus Fedotova 1998a: 28. Kazakhstan: Saïramskiï Mtn Range, upper reaches Saïramsu R., 10 

km SW Kaskasu; HT; ♂; ZMAS. 

saliciphilus. Kazakhstan. Undet. sp. of Eriophyidae (Acarina).

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Arthrocnodax saliciphilus Fedotova 1995a: 16. Kazakhstan: Saur Mtn Range, 13 km S Zaïsan, nr Zhanaturmys; HT; 

♂; ZMAS. 

salviae. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax salviae Fedotova 1995a: 19. Kazakhstan: Zailiïskiï Alatau, nr Kok-Pek; HT; ♂; ZMAS. 

*sophorae. Kazakhstan. 

Arthrocnodax sophorae Fedotova 1992o: 125. Kazakhstan: vic. Bakh-Bakhty; HT; ♂; ZMAS. 

spiraeae. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax spiraeae Fedotova 1997a: 41. Kazakhstan: foothills Saur Mtn Range, along Kenderlyk R., 20 km SE 

Zaïsan, nr Przheval'sk; HT; ♂; ZMAS. 

tanaceti. France, Germany. 

Arthrocnodax tanaceti Kieffer 1925: 35. France; HT; LT; lost?. 

thymiphilus. Kazakhstan. Undet. sp. of Eriophyidae (Acarina). 

Arthrocnodax thymiphilus Fedotova 1995a: 21. Kazakhstan: Terskeï Alatau, upper Karkara R., 24 km S Kegen'; HT; 

♂; ZMAS. 

vitis. Germany. Eriophyes vitis (Acarina: Eriophyidae). 

Arthrocnodax vitis Rübsaamen 1895a: 189. Germany: Rhembsohl; ST; ♂,♀,P,L; ZMHB. 

wissmanni. Germany. Phyllocoptes schlechtendali (Acarina: Eriophyidae). 

Arthrocnodax wissmanni Kieffer 1924: 89. Germany: Geisenheim; ST; ♂,♀; lost?. 

ASCHISTONYX 

Aschistonyx Rübsaamen 1917: 66. Type species, carpinicolus Rübsaamen (mon.). 


carpinicolus. Netherlands, Germany, Poland, Czechoslovakia, Austria, Ukraine, Romania, Yugoslavia, Greece. 

Carpinus betulus (Betulaceae). 

Aschistonyx carpinicolus Rübsaamen 1917: 66. Germany; ST; ♂,♀,L; ZMHB. 

ASPHONDYLIA 

Asphondylia Loew 1850a: 21, 37, as subg. of Cecidomyia. Type species, sarothamni Loew (subs. des. Karsch 1877: 

15). 

Phyllophaga Rondani 1856: 199. Type species, Cecidomyia fusca Meigen (orig. des.). 

Asphondilia Rondani 1860: 290, missp. of Asphondylia. 

Cylindrocera Lioy 1864: 503. Type species, Asphondylia sarothamni Loew (subs. des. Gagné 2004b). 

Monasphondylia Kieffer 1913g: 48. Type species, Asphondylia phlomidis Trotter (orig. des.). 

Eumarchalia (also as Eumarschalia) Del Guercio 1914: 231, as subg. of Schizomyia. Type species, Schizomyia 

gennadii Marchal (orig. des.). 

Gisonobasis Rübsaamen 1916a: 432. Type species, tournefortiae Rübsaamen (subs. des. Rübsaamen 1916c: 12). 

Ischnonyx Rübsaamen 1916c: 5. Type species, Cecidomyia verbasci Vallot (subs. des. Möhn 1955a: 147). 

Gisomobasis Shinji 1944: 227, missp. of Gisonobasis. 

Eoasphondylia Möhn 1960c: 214, as subg. of Asphondylia. Type species, Asphondylia convolvuli Möhn (orig. des.). 

Ussurasphondylia Kovalev 1964: 438, as subg. of Asphondylia. Type species, Asphondylia aceris Kovalev. 

Halimodendromyia Marikovskij 1965: 75. Type species, heptopotamica Marikovskij (orig. des.). 

Chenasphondylia Mamaev 1972a: 898, as subg. of Asphondylia. Type species, Asphondylia aelleniae Mamaev (orig. 

des.). 

Aspondilia Fedotova & Kovalev 2001: 547, missp. of Asphondylia. 

Cosmopolitan; 289 spp.; Cecidomyiidi, Asphondyliini, Asphondyliina. Asphondylia species mostly form galls on 

flowers or flower buds and prevent fruiting in a great many families of plants, but some form complex galls on other 

plant parts. All are associated with symbiotic fungi. Some species are of economic importance, e.g., A. websteri (cf.

119 

Gagné & Wuensche 1986) and A. gennadii (cf. Gagné and Orphanides 1992) and include those that affect buds and 

have alternate or multiple hosts. Some species alternate generations between two parts of a single plant species, e.g., A. 

rudbeckiaeconspicua, that forms a winter crown gall at the base of the plant and a summer flower bud gall (Plakidas 

1988). Many species groups within this genus are the result of radiation on particular plant genera, e.g. Atriplex 

(Hawkins et al. 1986) and Larrea (Gagné and Waring 1990), and are closely host specific. Refs: Möhn 1961a (rev., 

Neotr. spp.); Gagné 2004b (syn. note.). 

abutilon. USA (Texas). Abutilon incanum (Malvaceae). 

Asphondylia abutilon Felt 1935a: 1. USA: Texas, Austin; ST; ♂,♀; NYSM. 

aceris. Russia (Far East). Acer mono (Aceraceae). 

Asphondylia aceris Kovalev 1964: 438. Russia: Primorskiï Terr., Okeanskaya Station, nr Vladivostok; HT; ♂; 

KOVC. 

achatocarpi. El Salvador. Achatocarpus nigricans (Phytolaccaceae). 

Asphondylia achatocarpi Möhn 1959: 304. El Salvador: La Libertad, San Diego, E La Libertad; HT; ♀; SMNS. 

acinata. Israel, Eritrea. Suaeda monoica (Chenopodiaceae). 

Asphondylia acinata Dorchin 2001: 565. Israel: Ne'ot HaKikkar; HT; ♂; TAUI. 

adenocarpi. Portugal. Adenocarpus intermedius (Fabaceae). 

Asphondylia adenocarpi Tavares 1902b: 31. Portugal: Ponte do Lima; ST; ♀,P; TAVC. 

adenostoma. USA (California). Adenostoma fasciculatum (Asteraceae). 

Asphondylia adenostoma Felt 1916e: 177. USA: California, Los Angeles, Laurel Canyon; ST; ♀,P; NYSM. 

aelleniae. Turkmenistan, Uzbekistan. Aellenia subaphylla (Chenopodiaceae). 

Asphondylia aelleniae Mamaev 1972a: 898. Uzbekistan: Bukhara Prov., Artesianskiy; HT; ♂; ZMAS. 

africana. South Africa. Cissus cuneifolia (Vitaceae). 

Asphondylia africana Felt 1926h: 45. South Africa: East London; ST; ♂,♀; NYSM. 

ajallai. El Salvador. Vernonia canescens; V. patens, Vernonia sp. (Asteraceae). 

Asphondylia ajallai Möhn 1959: 366. El Salvador: La Libertad, Los Chorros; HT; ♀; SMNS. 

albiventris. Kenya. 

Asphondylia albiventris Kieffer 1913h: 93. Kenya: Likoni, nr Mombasa; HT; ♀; MNHN. 

altani. Nicaragua. 

Asphondylia altani Felt 1915a: 206. Nicaragua: San Marcos; HT; ♀; USNM. 

amaranthi. USA (Texas, Florida), Mexico (Tamaulipas). Amaranthus blitoides, A. "spinosus" (Amaranthaceae). 

Asphondylia amaranthi Felt 1935a: 1. USA: Texas, Austin; ST; ♂,♀; NYSM. 

ambrosiae. USA (California, Texas, Florida). Ambrosia sp.; Ambrosia spp. (Asteraceae). 

Asphondylia ambrosiae Gagné 1975b: 51. USA: Florida, Miami; HT; P; USNM. 

anaceliae. El Salvador. Bidens squarrosa; B. refracta (Asteraceae). 

Asphondylia anaceliae Möhn 1959: 355. El Salvador: La Libertad, Cumbre, Hacienda Chilata; HT; ♂; SMNS. 

anatolica. Turkey. Astragalus lagurus (Fabaceae). 

Asphondylia anatolica Skuhravá & Çam 1998: 218. Turkey: Ta�liçiftlik, nr Tokat; HT; ♂; SKCR. 

andirae. El Salvador. Andira inermis (Fabaceae). 

Asphondylia andirae Möhn 1959: 314. El Salvador: La Libertad, San Diego, E La Libertad; HT; LT; SMNS. 

annonae. El Salvador. Annona cherimola (Annonaceae). 

Asphondylia annonae Möhn 1959: 309. El Salvador: Santa Ana, N El Roblarón; HT; ♀; SMNS. 

anotisae. India (Tamil Nadu). Hedyotis leschanaultiana (Rubiaceae). 

Asphondylia anotisae Mani 1986: 16. India: Tamil Nadu, Madras; HT; ♀; ZSCI. 

antennariae. USA (Wisconsin, Maine). Antennaria plantaginifolia (Asteraceae). 

Asynapta antennariae Wheeler 1889: 212. USA: Wisconsin, Milwaukee; ST; ♂,♀,P,L; USNM. 

anthocercidis. Australia (Western Australia). Anthocercis littorea (Solanaceae). 

Asphondylia anthocercidis Kolesik 1997b: 157. Australia: Western Australia, Hillarys; HT; ♂; SAMA. 

apicata. USA (California, Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia apicata Gagné 1990c: 658. USA: Arizona, Black Canyon City dog track; HT; LT; USNM. 

ardisiae. El Salvador. Ardisia paschalis (Myrsinaceae). 

Asphondylia ardisiae Möhn 1959: 333. El Salvador: La Libertad, N El Cimarrón; HT; ♀; SMNS.

120 

artemisiae. USA (Arizona). Artemisia sp. (Asteraceae). 

Asphondylia artemisiae Felt 1908b: 377. USA: Arizona, Fort Grant; ST; ♀; USNM. 

atriplicicola. USA (California, Arizona, New Mexico). Atriplex canescens (Chenopodiaceae). 

Diplosis atriplicicola Cockerell 1898: 326. USA: New Mexico, Mesilla Park; LT (Gagné 1986d: 66); ♂; USNM. 

atriplicis. USA (California, Arizona, New Mexico). Atriplex canescens (Chenopodiaceae). 

Cecidomyia atriplicis Townsend 1893a: 1021. USA: New Mexico, nr Las Cruces; T; G; lost. 

Cecidomyia atriplicis Cockerell 1895: 766. USA: New Mexico, Las Cruces, College Farm; ST; ♀,P; lost. 

Asphondylia atriplics Gagné 1986d: 65, missp. of atriplicis. 

attenuatata. West Indies. Clerodendrum aculeatum (Verbenaceae). 

Asphondylia attenuatata Felt 1909a: 300. West Indies; ST; ♂; USNM. 

aucubae. Japan (Hokkaido to Kyushu). Aucuba japonica (Cornaceae). 

Asphondylia aucubae Yukawa & Ohsaki 1988: 368. Japan: Kagoshima; HT; ♂; KUIC. 

auripila. USA (California, Arizona, New Mexico, Texas), Mexico (SIN). Larrea tridentata (Zygophyllaceae). 

Asphondylia auripila Felt 1907d: 14. USA: Arizona, Tucson; LT (Gagné 1990c: 660); ♂; USNM. 

autumnalis. USA (Missouri, North Carolina). Helenium autumnale (Asteraceae). 

Asphondylia autumnalis Beutenmüller 1907a: 386. USA: North Carolina, Buncombe Co., Valley of the Black Mts; 

ST; ♂,♀; AMNH. 

ayeniae. El Salvador. Ayenia pusilla (Sterculiaceae). 

Asphondylia ayeniae Möhn 1960c: 216. El Salvador: Santa Ana, rd to Metapán, NE Santa Ana; HT; ♂; SMNS. 

azaleae. USA (New York, Maryland, Virginia). Azalea sp. (Ericaceae). 

Asphondylia azaleae Felt 1907d: 14. USA: New York, Albany; ST; ♂,♀; NYSM. 

baca. Korean Peninsula, Japan (Hokkaido to Kyushu). Ampelopsis heterophylla; Vitis labrusca and Cayratia japonica 

(Vitaceae), Weigela coraeensis and W. hortensis (Caprifoliaceae). 

Asphondylia baca Monzen 1937: 189. Japan: Honshu, Aomori Pref., Towada; NT (Yukawa & Ohsaki 1988: 374; 

Japan: Honshu, Asama-onsen, Matsumoto City); ♂; KUEC. Ref.: Uechi et al. 2004 (host alternation). 

Contarinia ampelopsivora Shinji 1938f: 321. Japan: Honshu; ST; ♂,♀; lost. 

Asphondylia vaca Shinji 1944: 135, missp. of baca. 

bacchariola. USA (Arizona, Texas). Baccharis pteronioides (Asteraceae). 

Asphondylia bacchariola Gagné 1995c: 768. USA: Arizona, Santa Cruz Co., Appleton-Whittel Research Ranch, 4 mi 

S Elgin; HT; LT; USNM. 

baccharis. Chile. Baccharis sp. (Asteraceae). 

Asphondylia baccharis Kieffer & Herbst 1905: 65. Chile: vic. San Vicente; T; G; lost. 

bahiensis. Brazil (Bahia). Undet. sp. of Rubiaceae. 

Asphondylia bahiensis Tavares 1917b: 156. Brazil: Bahia, betw. Bahia & R. de Vermelho; ST; ♂,♀,P; TAVC. 

barbata. USA (California, Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia barbata Gagné 1990c: 660. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

baroni. USA (Arizona). 

Asphondylia baroni Felt 1908b: 377. USA: Arizona; ST; ♀; NYSM. 

baudysi. Widespread Europe. Coronilla varia (Fabaceae). 

Asphondylia baudysi Vimmer 1937: 107. Czech Republic: Blansko nr Brno; T; ♂; ?. 

baumanni. Togo. Vitis sp. (Vitaceae). 

Asphondylia baumanni Rübsaamen 1899e: 274. Togo: Misahöhe; ST; ♀,P; ZMHB. 

bea. USA (California). Senecio douglassi (Asteraceae). 

Asphondylia bea Felt 1925b: 19. USA: California, Claremont, Pomona College; ST; ♂,♀; NYSM. 

beguni. India (Andhra Pradesh). Solanum melongena (Solanaceae). 

Asphondylia beguni Mani 1953a: 119. India: Andhra Pradesh, Hyderabad; HT; ♀; ZSCI. 

betheli. Widespread southwestern Nearctic. Opuntia sp. (Cactaceae). 

Asphondylia betheli Cockerell 1907: 324. USA: Colorado, Boulder; ST; ♂,P; USNM. 

Asphondylia arizonensis Felt 1907d: 13. USA: Arizona, Fort Grant; ST; ♂,♀; USNM. 

Asphondylia opuntiae Felt 1908b: 377. USA: Texas, Sinton; ST; ♂,♀; NYSM. 

Asphondylia optuniae Pritchard 1953a: 135, missp. of opuntiae. 

bidens. USA (Florida). Bidens pilosa (Asteraceae).

121 

Asphondylia bidens Johannsen 1945: 9. USA: Florida, Englewood; HT; ♂; CUIC. 

bigeloviaebrassicoides. USA (New Mexico). Chrysothamnus graveolens (Asteraceae). 

Cecidomyia bigeloviaebrassicoides (as bigeloviae-brassicoides) Townsend 1893b: 491. USA: New Mexico, a few 

mi. E Gallo Spring; ST; G; lost. 

blechi. Colombia. Blechum pyramidatum (Acanthaceae). 

Asphondylia blechi Wünsch 1979: 31. Colombia: Magdalena Prov., R. Buritaca; HT; ♂; SMNS. 

boerhaaviae. Mexico, El Salvador, Colombia, Jamaica. Boerhaavia erecta; Boerhaavia spp. (Nyctaginaceae). 

Asphondylia boerhaaviae Möhn 1959: 302. El Salvador: San Vicente, nr R. San Felipe; HT; ♂; SMNS. 

borreriae. Brazil (Rio de Janeiro). Borreria sp.; B. verticillata (Rubiaceae). 

Asphondylia borreriae Rübsaamen 1905a: 74. Brazil: Rio de Janeiro, Harpoador; T; P; ZMHB. 

borrichiae. USA (South Carolina, Florida); Mexico. Borrichia frutescens (Asteraceae). 

Asphondylia borrichiae Rossi & Strong 1990: 732. USA: Florida, Wakulla Co., St. Mark's Wildlife Refuge; HT; ♂; 


borzi. UK, France, Portugal, Italy. Rhamnus alaternus (Rhamnaceae). 

Cecidomyia borzi De Stefani 1898a: 104. Italy: Sicily, Palermo; ST; ♀,P,L; lost. 

brevicauda. USA (Arizona). 

Asphondylia brevicauda Felt 1907d: 14. USA: Arizona, Fort Yuma; HT; ♀; USNM. 

brevipalpis. Kenya. 

Asphondylia brevipalpis Kieffer 1913h: 92. Kenya: Likoni, nr Mombasa; HT; ♂; MNHN. 

buddleia. USA (Texas); El Salvador, France (immigr., teste Béguinot 1999). Buddleia racemosa, B. americana 

(Loganiaceae). 

Asphondylia buddleia Felt 1935a: 2. USA: Texas, Austin; ST; ♂,♀; NYSM. 

bullata. USA (Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia bullata Gagné 1990c: 663. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

bursaria. Indonesia (Java). Symplocos fasciculata (Symplocaceae). 

Asphondylia bursaria Felt 1927d: 387. Indonesia: Java, Tjibodas, Mt Gede; ST; ♂,♀,P; NYSM. 

cabezasae. El Salvador. Crotalaria mucronata; Gliricidia sepium, Desmodium nicaraguense (Fabaceae). 

Asphondylia cabezasae Möhn 1959: 312. El Salvador: La Libertad, Los Chorros; HT; ♂; SMNS. 

calaminthae. Italy, Yugoslavia. Calamintha alpina (Lamiaceae). 

Asphondylia calaminthae Kieffer 1909b: 3. Italy; T; G; lost?. 

caleae. El Salvador. Calea integrifolia (Asteraceae). 

Asphondylia caleae Möhn 1959: 357. El Salvador: San Salvador, Picacho; HT; ♂; SMNS. 

callicarpae. Java, Philippines. Callicarpa erioclona; C. longifolia (Verbenaceae). 

Asphondylia callicarpae Felt 1918c: 285. Philippines: Luzon, Laguna Prov., Mt Makiling; ST; ♂,♀; NYSM. 

calycotomae. Italy, Greece, Algeria, Tunisia. Calycotome intermedia (Fabaceae). 

Asphondylia calycotomae Kieffer 1912e: 107. Algeria: St-Denis-du-Sig; ST; ♂,P; lost?. 

camarae. El Salvador, Colombia. Lantana camara (Verbenaceae). 

Asphondylia camarae Möhn 1959: 340. El Salvador: Cuscatlán, 27 km E San Martin; HT; ♂; SMNS. 

canastrae. Brazil (Minas Gerais). Hyptis sp. (Lamiaceae). 

Asphondylia canastrae Urso-Guimarães & Amorim 2002: 562. Brazil: Minas Gerais, Delfinópolis, Estância Carmen 

Silvia; HT; ♂; MZSP. 

canavaliae. Colombia. Canavalia maritima (Fabaceae). 

Asphondylia canavaliae Wünsch 1979: 33. Colombia: Magdalena Prov., Cañaverales; HT; ♀; SMNS. 

capensis. South Africa. "Pabella sp." (?). 

Asphondylia capensis Kieffer 1914: 517. South Africa: Cape Prov.; ST; ♀,P; lost?. 

capparis. Circum-Mediterranean. Capparis spinosa (Capparidaceae). 

Asphondylia capparis Rübsaamen 1894a: 363. Italy, Catania; ST; ♂,♀,P; ZMHB. 

caprariae. Colombia. Capraria biflora (Scrophulariaceae). 

Asphondylia caprariae Wünsch 1979: 53. Colombia: Magdalena Prov., Bahia Nenguange; HT; ♂; SMNS. 

caudicis. USA (California). Atriplex canescens (Chenopodiaceae). 

Asphondylia caudicis Gagné 1986d: 66. USA: California, Indio; HT; P; USNM. 

ceanothi. USA (Oregon, California). Ceanothus sp.; C. cuneatus (Rhamnaceae).

122 

Asphondylia ceanothi Felt 1908b: 377. USA: California, Oakland; ST; ♀; NYSM. 

celsiae. Southern Europe. Celsia orientalis (Scrophulariaceae). 

Asphondylia celsiae Kieffer 1909b: 6. Southern Europe; T; G; lost?. 

cestri. El Salvador. Cestrum nocturnum (Solanaceae). 

Asphondylia cestri Möhn 1959: 344. El Salvador: La Libertad, Los Chorros; HT; ♂; SMNS. 

*chapae. Russia. Acinos arvensis (Lamiaceae). 

Asphondylia chapae Fedotova 2008: 135. Russia: Central Povolzhye, Zhigulevsk Nature Res., nr Bakhilova Polyana; 


chrysothamni. USA (Utah). Chrysothamnus graveolens (Asteraceae). 

Asphondylia chrysothamni Felt 1916e: 178. USA: Utah, nr Murray; ST; ♂,♀,P; NYSM. 

cissi. El Salvador. Cissus sicyoides (Vitaceae). 

Asphondylia cissi Möhn 1959: 324. El Salvador: La Paz, ca. 8 km before Herradura; HT; ♂; SMNS. 

clavata. USA (California, Arizona), Mexico (Baja California). Larrea tridentata (Zygophyllaceae). 

Asphondylia clavata Gagné 1990c: 663. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

clematidis. USA (Texas), El Salvador. Clematis drummondi; C. grossa, C. dioica (Rosaceae). 

Asphondylia clematidis Felt 1935a: 3. USA: Texas, Austin; ST; ♂,♀; NYSM. 

*clinopodiiflorae. Russia (Europe). Clinopodium vulgare (Lamiaceae). 

Asphondylia clinopodiiflorae Fedotova 2003b: 981. Russia: Samara Oblast, Zigulevskiy Res., nr Vakhilova Polyana; 


communis. Brazil (Rio de Janeiro). Ximenia americana (Olacaceae). 

Asphondylia communis Maia & Couri 1992: 659. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; 

QBUM. 

conglomerata. Spain, Italy, Greece, Israel. Atriplex halimus (Chenopodiaceae). 

Asphondylia conglomerata De Stefani 1900: 16. Italy: Sicily; ST; ♂,♀; lost. 

conspecta. Togo. Commelina communis (Commelinaceae). 

Asphondylia conspecta Kieffer 1913i: 93. Togo: vic. Misahöhe; T; P; ZMHB. 

convoluli. El Salvador. Ipomoea arborescens (Convolvulaceae). 

Asphondylia convoluli Möhn 1960c: 214. El Salvador: La Libertad, NE Sitio del Nino, W lava fields; HT; ♂; SMNS. 

corbulae. El Salvador, Guadeloupe, Trinidad. Eupatorium sp.; Chromolaena odorata, Fleischmannia microstemon 

(Asteraceae). 

Asphondylia corbulae Möhn 1960c: 197. El Salvador: Ahuachapán, NW Las Chinamas; HT; ♂; SMNS. 

cordiae. El Salvador, Brazil. Cordia dentata; C. alba (Boraginaceae). 

Asphondylia cordiae Möhn 1959: 338. El Salvador: La Libertad, W Ateos; HT; ♂; SMNS. 

coronillae. France, Italy, Yugoslavia, Hungary, Greece. Coronilla minima; C. emeroides, C. emerus (Fabaceae). 

Cecidomyia coronillae Vallot 1829: 112. France; ST; G, not L (the larva Vallot described was red and made a 

cocoon, neither of which fits Asphondylia larvae); lost?. 

Asphondylia jaapi Rübsaamen 1916b: 562. Serbia and Montenegro: Hercegnovi; ST; A,P,L; ZMHB. 

crassinervis. Kenya. 

Asphondylia crassinervis Kieffer 1913h: 92. Kenya: Shimoni; HT; ♀; MNHN. 

curatellae. El Salvador. Curatella americana (Dilleniaceae). 

Asphondylia curatellae Möhn 1959: 330. El Salvador: Chalatenango, SE La Palma; HT; ♂; SMNS. 

cyamopsii. India (Gujarat). Cyamopsis tetragonoloba (Fabaceae). 

Asphondylia cyamopsii Chari 1971: 892, with reference to Patel & Chari 1970: 90. India: Gujarat, Anad; ST; ♀,P,L; 

?. 

cytisi. Widespread Europe, Russia (Western Siberia); immigr., not estab.: USA (New York). Cytisus austriacus; 

Cytisus sp. (Fabaceae). 

Asphondylia cytisi Frauenfeld 1873: 186. Austria: Neusiedlersee; ST; ♀,P; ?. 

deserticola. Turkmenistan, Uzbekistan. Salsola rigida (Chenopodiaceae). 

Asphondylia deserticola Mamaev 1972a: 898. Uzbekistan: Bukhara Prov., Kul'dzhuktau; HT; ♂; MAMC. 

diervillae. USA (New York, Massachusetts). Diervilla lonicera (Caprifoliaceae). 

Asphondylia diervillae Felt 1907c: 165. USA: New York, Albany; T; ♀; NYSM. This species was erroneously 

recorded from Japan (Shinji 1938l); the subject of that record has been shown to be A. baca (Uechi et al. 2002,

123 

Uechi et al. 2004). 

Cecidomyia inaequalis Stebbins 1910: 48. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

Asphondylia dievervillae Shinji 1944: 103, missp. of diervillae. 

digerae. India (Delhi). Digera arvensis (Amaranthaceae). 

Asphondylia digerae Mani 1944: 155. India: Delhi; HT; ♂; INPC. 

digitata. USA (California, Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia digitata Gagné 1990c: 664. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

diplaci. USA (California). Diplacus longiflorus (Scrophulariaceae). 

Asphondylia diplaci Felt 1912h: 151. USA: California, Whittier, Puente Hills; ST; ♂,P; NYSM. 

discalis. USA (Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia discalis Gagné 1990c: 665. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

dodonaeae. Australia (South Australia). Dodonaea viscosa (Sapindaceae). 

Asphondylia dodonaeae Kolesik 1995c: 171. Australia: South Australia, Morialta Conservation Pk, 13 km NE 

Adelaide; HT; ♂; SAMA. 

dondiae. USA (California). Suaeda multiflora (Chenopodiaceae). 

Asphondylia dondiae Felt 1918b: 381. USA: California, Point Fermine; ST; ♂,P,L; NYSM. 

dorycnii. Widespread Europe. Dorycnium suffruticosum (Fabaceae). 

Cecidomyia dorycnii Müller 1870: 76. France: Menton; ST; A,P,L,egg; ?. 

Asphondylia dorycnii Löw 1880: 37. Austria: on the Mauberge nr Mödling and vic. Vienna; ST; ♂,♀,L; NHMW. 

duplicornis. Colombia. Melanthera aspera (Asteraceae). 

Asphondylia duplicornis Wünsch 1979: 51. Colombia: Magdalena Prov., Cañaverales; HT; ♂; SMNS. 

echii. Germany, Poland, Czechoslovakia, Austria, Italy, Hungary, Romania, Yugoslavia. Echium vulgare 

(Boraginaceae). 

Cecidomyia echii Loew 1850a: 38. Poland: T; A; ZMHB?. 

enceliae. USA (California). Encelia californica (Asteraceae). 

Asphondylia enceliae Felt 1912h: 152. USA: California, Whittier, Puente Hills; ST; ♀,P; NYSM. 

ericiformis. Australia (South Australia). Holosarcia indica subsp. leiostachya (Chenopodiaceae). 

Asphondylia ericiformis Kolesik 1997a: 63. Australia: South Australia, Lyndhurst; HT; ♂; SAMA. 

ervi. Widespread Europe, Russia (Western Siberia), Japan. Vicia hirsuta (Fabaceae). 

Asphondylia ervi Rübsaamen 1895c: 260. Germany: vic. St. Goar; T; A; ZMHB. 

erythroxylis. El Salvador. Erythroxylis mexicanum (Erythroxylaceae). 

Asphondylia erythroxylis Möhn 1959: 315. El Salvador: La Libertad, NE Sitio del Nino, W lava fields; HT; ♀; 

SMNS. 

eupatorii. USA (New York, Maryland). Ageratina altissima (Asteraceae). 

Asphondylia eupatorii Felt 1911l: 460. USA: New York, Medina; T; G; NYSM. 

evae. Colombia. Chamizoa sp. (Amaranthaceae). 

Asphondylia evae Wünsch 1979: 39. Colombia: Magdalena Prov., Bahia Gairaca; HT; ♂; SMNS. 

fabalis. USA (Arizona). Larrea tridentata. (Zygophyllaceae). 

Asphondylia fabalis Gagné 1990c: 665. USA: Arizona, Black Canyon City dog track; HT; LT; USNM. 

floccosa. USA (California, Arizona). Atriplex polycarpa (Chenopodiaceae). 

Asphondylia floccosa Gagné 1986d: 69. USA: California, Riverside; HT; P; USNM. 

Asphondylia flocossa Dixon, Craig & Hughes 1998: 592. missp. of floccosa. 

florea. USA (Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia florea Gagné 1990c: 666. USA: Arizona, Grand Canyon Natl Pk, Parashant Canyon, R. mile 198; HT; 


florida. USA (Florida). 

Asphondylia florida Felt 1908b: 376. USA: Florida, Coconut Grove; ST; ♂,P; USNM. 

*floriformis. Australia (Victoria). Sarcocornia quinqueflora (Chenopodiaceae). 

Asphondylia floriformis Veenstra-Quah & Kolesik 2007: 200. Australia: Victoria, Yaringa Marine Natl Pk, Tyabb; 

HT; ♂; SAMA. 

foliata. USA (California). Atriplex polycarpa (Chenopodiaceae). 

Asphondylia foliata Gagné 1986d: 70. USA: California, nr Thermal; HT; LT; USNM.

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foliosa. USA (Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia foliosa Gagné 1990c: 666. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

*fructicola. Brazil (Pará). Solanum sp. (Solanaceae). 

Asphondylia fructicola Maia 2009c: 167. Brazil: Pará, Porto Trompetas; HT; ♂; QBUM. 

fulvopedalis. USA (New York, Vermont). 

Asphondylia fulvopedalis Felt 1907a: 22. USA: New York, Westfield; HT; ♂; NYSM. 

fusca. Belgium. 

Cecidomyia fusca Meigen 1830: 270. Belgium: vic. Liège; T; ♀; NMHN. 

garryae. USA (Oregon, California). Garrya sp.; G. buxifolia, G. fremontii (Cornaceae). 

Asphondylia garryae Felt 1912k: 757. USA: California; T; G; NYSM?. 

Asphondylia garryae Felt 1934c: 34. USA: California, Lake Co, 1 mi SW Mt Konocti; ST; ♂,♀; USNM. This name 

is not necessarily synonymous with the earlier garryae as the gall descriptions do not exactly agree; they are 

kept as synonyms to avoid coining a possibly unnecessary new name. 

gemmae. USA (California). Atriplex canescens (Chenopodiaceae). 

Asphondylia gemmae Gagné 1986d: 67. USA: California, nr Valle Vista; HT; P; USNM. 

genistae. Widespread Europe. Genista sp.; G. germanica, G. sagittalis (Fabaceae). 

Cecidomyia genistae Loew 1850a: 38. Poland; ST; ♂,♀; ZMHB?. 

Asphondylia bitensis Kieffer 1888c: 264. France: Bitche; ST; ♂,♀,P,L; lost?. 

Asphondylia moraviae Vimmer 1928a: 53. Czech Republic: Ba kovce nad B�ezinou, nr Tišnov; T; ♂; ?. 

gennadii. Italy, Malta, Greece, Cyprus, Turkey. Ceratonia siliqua (Fabaceae); Capparis spinosa (Capparidaceae), 

Capsicum annuum (Solanaceae), Urginea maritima (Liliaceae). 

Schizomyia gennadii Marchal 1904: 272. Cyprus; ST; ♀,P,L; MNHN?. This may be a synonym of capparis 

Rübsaamen (Gagné & Orphanides 1992). 

Schyzomya gennadiosi Del Guercio 1918a: 126, unjust. emend. of gennadii. 

Cecidomyia ceratoniae Bevan 1918: 11. Cyprus; T; G; lost. 

Asphondylia capsici Barnes 1932b: 478. Cyprus; ST; ♂,♀; BMNH. 

glomeratae. Brazil (Rio de Janeiro). Mikania glomerata (Asteraceae). 

Asphondylia glomeratae Gagné 2001: 116. Brazil: Rio de Janeiro, Rio de Janeiro; HT; ♂; QBUM. 

*gochnatiae. Brazil (Minas Gerais). Gochnatia polymorpha (Asteraceae). 

Asphondylia gochnatiae Maia 2008: 54. Brazil: Minas Gerais, Luz; HT; ♂; MNRJ. 

godmaniae. El Salvador. Godmania aesculifolia (Bignoniaceae). 

Asphondylia godmaniae Möhn 1959: 346. El Salvador: W Chalatenango; HT; LT; SMNS. 

grewiae. Philippines. Grewia stylocarpa (Tiliaceae). 

Asphondylia grewiae Felt 1919a: 289. Philippines: Luzon, Laguna Prov., Mt Makiling; HT; ♂; NYSM. 

guareae. El Salvador. Guarea sp. (Meliaceae). 

Asphondylia guareae Möhn 1959: 316. El Salvador: Usulután, for. of San Marcos Lempa; HT; ♂; SMNS. 

helianthiglobulus. USA (widespread eastern). Helianthus sp.; Helianthus spp. (Asteraceae). 

Asphondylia helianthi globulus Osten Sacken 1869: 301, invalid, a polynomial. 

Asphondylia helianthiglobulus (as helianthi-globulus) Osten Sacken 1878: 5. USA: Illinois, Rock Island; T; G; 


Asphondylia globulus Beutenmüller 1907a: 387, error for helianthiglobulus. 

helianthiflorae. Canada (Quebec), USA (New York). Helianthus strumosus (Asteraceae). 

Asphondylia helianthiflorae Felt 1908b: 376. USA: New York, Highland; ST; ♂,♀; NYSM. 

helicteris. El Salvador. Helicteres mexicana (Sterculiaceae). 

Asphondylia helicteris Möhn 1959: 327. El Salvador: La Paz, NW Olocuilta; HT; ♀; SMNS. 

heptopotamica. Kazakhstan. Halimodendron halodendron (Fabaceae). 

Halimodendromyia heptopotamica Marikovskij 1965: 75. Kazakhstan: mid course Ili R., Karachingil', & mid course 

Charyn R., Septogai; ST; ♂,♀; MARC. 

herculesi. El Salvador. Vernonia canescens (Asteraceae). 

Asphondylia herculesi Möhn 1959: 363. El Salvador: Chalatenango, SE La Palma; HT; ♂; SMNS. 

hieronymi. Argentina. Baccharis salicifolia (Asteraceae) 

Lasioptera hieronymi Weyenbergh 1875: 164 (publ'n not seen). Argentina; T; ?; NHMW?.

125 

hilli. Australia (Northern Territory). 

Asphondylia hilli Edwards 1916: 498. Australia: Northern Territory, Darwin; ST; ♀,P; BMNH?. 

hornigi. Poland, Czechoslovakia, Austria, Hungary, Romania, Bulgaria, Yugoslavia, Russia (Europe). Origanum 

vulgare (Lamiaceae). 

Asphondylia hornigi Wachtl 1880: 531. Austria: vic. Vienna; ST; ♂,♀,P; NHMW?. 

hydrangeae. USA (Missouri, Maryland, Virginia). Hydrangea arborescens (Hydrangeaceae). 

Asphondylia hydrangeae Felt 1907d: 15. USA: Virginia; ST; ♂; USNM. 

hyptis. El Salvador. Hyptis verticillata; H. suaveolens (Lamiaceae). 

Asphondylia hyptis Möhn 1960c: 206. El Salvador: San Salvador; HT; P; SMNS. 

ilicicola. USA (Pennsylvania, Connecticut, Maryland, Virginia, North Carolina). Ilex sp.; Ilex spp. (Aquifoliacae). 

Asphondylia ilicicola Foote 1953: 197. USA: Virginia, Williamsburg; HT; ♂; USNM. 

ilicoides. USA (New York). Nemopanthus mucronatus (Aquifoliaceae). 

Asphondylia ilicoides Felt 1907d: 15. USA: New York, Old Forge; ST; ♂,♀; NYSM. 


Asphondylia indica Rao 1957: 1. India: Uttar Pradesh, Pradesh, Agra; HT; ♀; SJCA. 

indigoferae. El Salvador. Indigofera suffruticosa (Fabaceae). 

Asphondylia indigoferae Möhn 1960c: 204. El Salvador: San Salvador; HT; ♂; SMNS. 

inflata. Australia (South Australia). Halosarcia pergranulata (Chenopodiaceae). 

Asphondylia inflata Kolesik 1997a: 61. Australia: South Australia, Port Adelaide; HT; ♂; SAMA. 

integrifoliae. USA (California, Utah). Rhus integrifolia; R. trilobata (Anacardiaceae). 

Asphondylia integrifoliae Felt 1908b: 376. USA: California, Los Angeles Co.; ST; ♀,P; USNM. 

ipomoeae. India (Tamil Nadu). Ipomoea staphylina (Convolvulaceae). 

Asphondylia ipomoeae Felt 1926i: 243. India: Tamil Nadu, Coimbatore; ST; ♀,P; NYSM. 

Asphondylia ipomaeae Rao 1955: 243, missp. of ipomoeae. 

*itoi. Japan (Kyushu, Okinawa). Distylium racemosum (Hamamelidaceae). 

Asphondylia itoi Uechi & Yukawa 2004: 31. Japan: Okinawa Pref., Okinawa I., Kunigami Village, Mt. Katsuudake; 


ixora. Indonesia (Java). Ixora timoriensis (Rubiaceae). 

Asphondylia ixora Felt 1927d: 388. Indonesia: Java, Djakarta, Klein Kombius I.; ST; ♀; NYSM. 

jatrophae. El Salvador. Jatropha curcas (Euphorbiaceae). 

Asphondylia jatrophae Möhn 1959: 319. El Salvador: La Paz, ca. 8 km before Herradura; HT; ♀; SMNS. 

johnsoni. USA (Pennsylvania). Solidago sp. (Asteraceae). 

Asphondylia johnsoni Felt 1908b: 377. USA: Pennsylvania, Lansdowne; ST; ♂,P; NYSM. 

kopetdagica. Turkmenistan. Scrophularia turcomanica (Scrophulariaceae). 

Asphondylia kopetdagica Fedotova 1993p: 58. Turkmenistan: Kopetdag Res., nr Parkhaï, 7 km NE Garrygala; HT; 

♀; ZMAS. 

lacinariae. USA (Kansas, Texas). Lacinaria punctata; L. pycnostacha (Asteraceae). 

Asphondylia lacinariae Felt 1935a: 3. USA: Texas, Austin; ST; ♂,♀; NYSM. 

lantanae. India (Tamil Nadu). Lantana camara, L. indica (Verbenaceae). 

Asphondylia lantanae Felt 1920b: 2. India: Tamil Nadu, Nilgiri Hills, Kallar; ST; ♂,♀; NYSM. 

lathyri. Widespread Europe, Russia (Western Siberia). Lathyrus pratensis (Fabaceae). 

Asphondylia lathyri Rübsaamen 1914: 110. Germany; T; ♂; ZMHB. 

leeae. Indonesia (Java). Leea sambucina (Vitaceae). 

Asphondylia leeae Felt 1921h: 148. Indonesia: Java, Bogor; ST; ♂,♀,P; NYSM. 

ligustrinae. Russia (Far East). Ligustrina amurensis (Oleaceae). 

Asphondylia ligustrinae Kovalev 1964: 440. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♀; KOVC. 

lippiae. El Salvador. Lippia cardiostegia (Verbenaceae). 

Asphondylia lippiae Möhn 1959: 342. El Salvador: San Salvador, E San Marcos; HT; ♂; SMNS. 

litseae. Indonesia (Java). Litsea sp. (Lauraceae). 

Asphondylia litseae Felt 1921h: 149. Indonesia: Java, Ungaran Mts; ST; ♀,P; NYSM. 

Asphondylia listeae Rao 1955: 243, missp. of litseae. 

lonchocarpi. El Salvador. Lonchocarpus minimiflorus (Fabaceae).

126 

Asphondylia lonchocarpi Möhn 1960c: 205. El Salvador: Sonsonate, S Los Cóbanos; HT; ♀; SMNS. 

lopezae. Colombia. Amaranthus dubius, A. spinosus, Iresine angustifolia (type host not specified) (Amaranthaceae). 

Asphondylia lopezae Wünsch 1979: 43. Colombia: Magdalena Prov., Santa Marta; HT; ♂; SMNS. 

lupini. Italy. Lupinus albus (Fabaceae). 

Asphondylia lupini Silvestri 1908: 3. Italy: Campania, nr Nola; ST; ♂,♀,P,L; ?. 

lupulinae. UK, France, Germany, Czechoslovakia, Ukraine. Medicago lupulina (Fabaceae). 

Asphondylia lupulinae Kieffer 1909b: 14. France; T; G; lost?. 

*lycopidiflorae. Russia (Europe). Lycopus europaeus (Lamiaceae). 

Asphondylia lycopidiflorae Fedotova 2003b: 981. Russia: Samara Oblast, 30 km SE Samara, nr Ust'Kinelsk; HT; ♂; 

ZMAS. 

malvavisci. El Salvador. Malvaviscus arboreus (Malvaceae). 

Asphondylia malvavisci Möhn 1959: 325. El Salvador: San Salvador; HT; ♂; SMNS. 

maricensis. Brazil (Rio de Janeiro). Struthanthus maricensis (Loranthaceae). 

Asphondylia maricensis Maia & Couri 1992: 658. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♀; 

QBUM. 

massalongoi. Czechoslovakia, Hungary, Romania, Italy, Yugoslavia, Greece. Ajuga chamaepitys (Lamiaceae). 

Asphondylia massalongoi Rübsaamen 1893: 163. Italy; ST; ♂,♀; ZMHB. 

*mcneilli. Australia (South Australia). Scleroderma diacantha (Chenopodiaceae). 

Asphondylia mcneilli Kolesik & Veenstra-Quah 2008: 218. Australia: South Australia, Eyre Peninsula, Minnipa; HT; 

♂; SAMA. 

melanopus. Widespread Europe. Lotus corniculatus (Fabaceae). 

Asphondylia melanopus Kieffer 1890a: 31. France: vic. Bitche; T; ♀; lost?. 

melantherae. El Salvador. Melanthera nivea (Asteraceae). 

Asphondylia melantherae Möhn 1959: 360. El Salvador: San Miguel, N El Delirio; HT; ♂; SMNS. 

menaschei. India (West Bengal). 

Asphondylia menaschei Mani 1937: 283. India: West Bengal, Calcutta, Portland Park; HT; ♂; ZSCI. 

menthae. Widespread Europe. Mentha pulegium, M. macrostachya; M. rotundifolia (Lamiaceae). 

Asphondylia menthae Kieffer 1902a: 365. France, Italy; T; G; lost?. 

Gisonobasis ignorata Rübsaamen 1916c: 9. Portugal; T; P; TAVC?. 

mentzeliae. USA (New Mexico). Mentzelia multiflora (Loasaceae). 

Asphondylia mentzeliae Cockerell 1900b: 302. USA: New Mexico, Raton; ST; ♂,P; USNM. 

mesembrianthemi. South Africa. Mesembryanthemum sp. (Aizoaceae). 

Asphondylia mesembrianthemi Schiner 1868: 9. South Africa: Cape of Good Hope; HT; ♀; NHMW. 

*microcapillata. Brazil (Minas Gerais). Bauhinea brevipes (Fabaceae). 

Asphondylia microcapillata Maia 2005d: 28. Brazil: Minas Gerais, Três Marias; HT; ♂; MNRJ. 

miki. Widespread Europe, Russia (Western Siberia). Medicago sativa (Fabaceae). 

Asphondylia miki Wachtl 1880: 535. Austria: vic. Vienna; ST; ♂,♀,P; NHMW?. 

mimosae. USA (Texas). Mimosa sp. (Fabaceae). 

Asphondylia mimosae Felt 1934a: 77. USA: Texas, Brownsville; ST; ♀,P; USNM. 

mitroshinae. Kazakhstan. Salsola richteri (Chenopodiaceae). 

Asphondylia mitroshinae Fedotova 1991f: 10. Kazakhstan: northeastern Kyzylkumy, nr Karaktau Mtn, 64 km NW 

Bairkum; HT; ♂; ZMAS. 

moehni. Brazil (Rio Grande do Sul, Rio de Janeiro). Mikania guaco; M. glomerata (Asteraceae). 

Asphondylia tavaresi Möhn 1973: 4, jun. sec. hom. tavaresi Rübsaamen. Brazil: Rio Grande do Sul, São Leopoldo; 

HT; ♂; Tavares. 

Asphondylia moehni Skuhravá 1989: 202, new name for Asphondylia tavaresi Möhn. 

monacha. Widespread eastern Nearctic. Solidago canadensis (Asteraceae). 

Asphondylia monacha Osten Sacken 1869: 299. USA: New York, nr Brooklyn; ST; ♂,♀,P; USNM. 

morindae. India (Tamil Nadu). Morinda tinctoria (Rubiaceae). 

Asphondylia morindae Mani 1934a: 134. India: Tamil Nadu, Thanjavur; HT; ♂; ZSCI. 

morivorella. Japan (Hokkaido, Honshu). Morus alba (Moraceae). 

Diplosis morivorella Naito 1919: 29. Japan; ST; ♂,♀,P,L; lost?.

127 

Diplosis moricola Matsumura 1931: 403. Japan: Hokkaido, Sapporo; HT; ♀; Matsumura Coll. 

murrayae. India (Uttar Pradesh). Murraya koenigii (Rutaceae). 

Asphondylia murrayae Grover & Jaiswal 1993: 56. India: Uttar Pradesh, Kotdwara, Garhwal; HT; ♂; PGCA. 

napiformis. Kenya. Acacia mellifera (Fabaceae). 

Asphondylia napiformis Gagné 1993c: 107. Kenya: Rt. A109, Kapiti Plains, Malili Ranch, ca. 70 km SE Nairobi; HT; 


neomexicana. USA (California, Arizona, New Mexico). Atriplex canescens (Chenopodiaceae). 

Cecidomyia neomexicana Cockerell 1896: 204. USA: New Mexico, Organ; LT (Gagné 1986d: 68); ♂; USNM. 

*nepetaflorae. Russia (Europe). Nepeta pannonica; N. nuda, N. ukrainica (Lamiaceae). 

Asphondylia nepetaflorae Fedotova 2003b: 973. Russia: Samara Oblast, 30 km SE Samara, nr Ust'Kinelsk; HT; ♂; 

ZMAS. 

nodula. USA (California). Atriplex polycarpa (Chenopodiaceae). 

Asphondylia nodula Gagné 1986d: 70. USA: California, Riverside Co., Torres Martinez Indian Res.; HT; P; USNM. 

obscura. Australia (New South Wales). Solanum physalifolium var. nitidibaccatum; S. chenopodioides (Solanaceae). 

Asphondylia obscura Kolesik 2000a: 34. Australia: New South Wales, Mt. Tomah; HT; ♂; ANIC. 

ocimi. El Salvador. Ocimum micranthum (Lamiaceae). 

Asphondylia ocimi Möhn 1959: 343. El Salvador: Morazán, SW Jocoro; HT; ♂; SMNS. 

ononidis. Widespread Europe, Libya. Ononis spinosa; Ononis spp. (Fabaceae). 

Asphondylia ononidis Löw 1873: 139. Austria: nr Pressbaum, Vienna Woods; ST; ♂,♀,P,L; NHMW. 

Asphondylia frauenfeldi Kieffer 1909b: 15. Austria; T; G; NHMW?. 

osbeckiae. India (Assam). Osbeckia sp. (Malastomataceae). 

Asphondylia osbeckiae Senior-White 1922: 107. India: Assam, Shillong; HT; ♀; ZSCI?. 

palaciosi. El Salvador. Crotalaria longirostrata (Fabaceae). 

Asphondylia palaciosi Möhn 1959: 310. El Salvador: Santa Ana, NW Ciudad Arce; HT; ♂; SMNS. 

parasiticola. El Salvador. Struthanthus marginatus (Loranthaceae). 

Asphondylia parasiticola Möhn 1960c: 202. El Salvador: nr Ahuachapán; HT; ♀; SMNS. 

parva. Brazil (Bahia). Undet. sp. of Rubiaceae. 

Asphondylia parva Tavares 1917b: 159. Brazil: Bahia, Roça da Madre de Deus; ST; ♂,♀,P; TAVC. 

pattersoni. St. Vincent. Citharexylum quadrangulare (Verbenaceae). 

Asphondylia pattersoni Felt 1911j: 301. St. Vincent: S; ♂,♀; NYSM. 

paucidentata. Australia (Queensland, New South Wales). Solanum aviculare; S. linearifolium (Solanaceae). 

Asphondylia paucidentata Kolesik 2000a: 34. Australia: Queensland, Bunya Mtns; HT; ♂; ANIC. 

peploniae. Brazil (Rio de Janeiro). Peplonia asteria (Asclepiadaceae). 

Asphondylia peploniae Maia 2001a: 4. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

phlomidis. Malta, Greece. Phlomis fruticosa; P. tuberosa (Lamiaceae). 

Asphondylia phlomidis Trotter 1901: 345. Greece: vic. Volo in Thessaly, on Corinth Acropolis, & Corfu; T; ♂; lost?. 

Asphondylia phlomidis Kieffer 1909b: 15. Greece: vic. Volo in Thessaly, Corinth Acropolis, & Corfu; T; G; lost?. 

photiniae. USA (California). Photinia arbutifolia (Rosaceae). 

Asphondylia photiniae Pritchard 1952a: 16. USA: California, Palo Alto; HT; ♂; USNM. 

phyllanthi. Burma. Phyllanthus emblica (Euphorbiaceae). 

Asphondylia phyllanthi Felt 1920b: 2. Burma: Maymyo; ST; ♂,♀; NYSM. 

pictipennis. Kenya. 

Asphondylia pictipennis Kieffer 1913h: 91. Kenya: Shimoni, 75 km from Monbasa; HT; ♀; MNHN. 

pila. USA (Arizona), Mexico (Baja California). Larrea tridentata (Zygophyllaceae). 

Asphondylia pilosa Gagné 1990c: 667, jun. prim. hom. pilosa Kieffer. USA: Arizona, Saguaro Natl Mon., East, 20 

mi. E Tucson; HT; LT; USNM. 

Asphondylia pila Gagné 2004b: 82, new name for pilosa Gagné. 

pilosa. Widespread Europe; immigr.: USA (Washington, Oregon). Cytisus scoparius (Fabaceae). 

Asphondylia pilosa Kieffer 1898f: 59. France: Moselle; T; LT; lost?. Considered a synonym of A. sarothamni Loew 

in Skuhravá 1986, the two species have now been shown to be distinct in Harris 2002. 

Ischnonyx pilosa Rübsaamen 1916c: 10. Germany; ST; ♀,P,L; ZMHB. 

*plaksinae. Russia. Ajuga plaksinae (Lamiaceae).

128 

Asphondylia plaksinae Fedotova 2008: 138. Russia: Central Povolzhye, Zhigulevsk Nature Res., nr Bakhilova 

Polyana; HT; ♂; ZMAS. 

pongamiae. India (Tamil Nadu). Pongamia glabra (Fabaceae). 

Asphondylia pongamiae Felt 1921j: 24. India: Tamil Nadu, Coimbatore; ST; ♂,♀,P; NYSM. 

portulacae. USA (Florida), El Salvador, Colombia, West Indies, Bolivia, Argentina. Portulaca oleracea 

(Portulacaceae). 

Asphondylia portulacae Möhn 1959: 308. El Salvador: La Libertad, San Diego, E La Libertad; HT; P; SMNS. 

prosopidis. USA (New Mexico, Texas). Prosopis glandulosa (Fabaceae). 

Asphondylia prosopidis Cockerell 1898: 329. USA: New Mexico, Mesilla Park; ST; ♂,P; USNM. 

pruniperda. Widespread Europe. Prunus domestica; P. spinosa (Rosaceae). 

Asphondylia pruniperda Rondani 1867: 37. Italy; ST; ♂,♀; MZUF. 

Asphondylia prunorum Wachtl 1880: 205. Austria: vic. Vienna; ST; ♂,♀,L; NHMW?. 

Asphondylia prunorum Kieffer 1889a: 149. France: Moselle, forest nr Courcelles sur Nied & vic. Guinkirchen; ST; 

♂,P,L; lost?. 

psychotriae. El Salvador. Psychotria pubescens (Rubiaceae). 

Asphondylia psychotriae Möhn 1959: 349. El Salvador, San Salvador; HT; ♂; SMNS. 

pterosparti. Portugal, Spain, Greece. Genista tridentata (Fabaceae). 

Asphondylia pterosparti Tavares 1901: 69. Portugal: S. Fiel, Sobral do Campo; ST; ♂; TAVC. 

punica. Greece, Algeria, Tunisia, Libya. Atriplex halimus (Chenopodiaceae). 

Asphondylia punica Marchal 1897a: 20. Tunisia: betw. Sousse & Kairouan; ST; A,P,L; MNHN?. 

randiae. El Salvador. Randia aculeata (Rubiaceae). 

Asphondylia randiae Möhn 1959: 350. El Salvador: La Libertad, S. Ciudad Arce; HT; ♂; SMNS. 

ratibidae. USA (Texas). Ratibida columnaris (Asteraceae). 

Asphondylia ratibidae Felt 1935a: 5. USA: Texas, Austin & Brownwood; ST; ♂,♀; NYSM. 

recondita. USA (New York, Maryland, North Carolina). Aster patens; Aster spp. (Asteraceae). 

Asphondylia recondita Osten Sacken 1875: 202. USA: New York, Long Island, Lloyd's Neck; T; P; lost. 

Asphondylia patens Beutenmüller 1907a: 386. USA: North Carolina, Buncombe Co., Valley of the Black Mts; ST; ♀; 

AMNH. 

resinosa. USA (Arizona), Mexico (Jalisco). Larrea tridentata (Zygophyllaceae). 

Asphondylia resinosa Gagné 1990c: 667. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; 


ricini. India (Andhra Pradesh), South Africa. Ricinus communis (Euphorbiaceae). 

Asphondylia ricini Mani 1947: 439. India: Andhra Pradesh, Hyderabad; HT; ♀; ZSCI. 

riveae. India (Tamil Nadu). Rivea hypocrateriformis (Convolvulaceae). 

Asphondylia riveae Mani 1934b: 411. India: Tamil Nadu, Thanjavur; HT; ♂; ZSCI. 

rochae. Brazil (Ceará). Jussiaea sp. (Onagraceae). 

Asphondylia rochae Tavares 1918c: 68. Brazil: Ceará, nr Fortaleza; ST; ♂,♀,P; TAVC. 

rondeletiae. El Salvador. Rondeletia strigosa (Rubiaceae). 

Asphondylia rondeletiae Möhn 1959: 352. El Salvador: San Salvador, forest of Picacho; HT; ♂; SMNS. 

rosetta. USA (California, Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia rosetta Gagné 1990c: 668. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

rosmarini. France, Italy, Slovenia, Yugoslavia, Tunisia. Rosmarinus officinalis (Lamiaceae). 

Asphondylia rosmarini Kieffer 1896e: 217. France: Marseille; ST; A,P,L; lost?. 

rubescens. South Africa. Tarchonanthus sp. prob. camphoratus (Asteraceae). 

Asphondylia rubescens Schiner 1868: 8. South Africa: nr Constantia, Cape of Good Hope; HT; ♀; NHMW?. 

rudbeckiaeconspicua. USA (Arizona, widespread eastern). Rudbeckia laciniata (Asteraceae). 

Asphondylia rudbeckiae conspicua Osten Sacken 1870: 51, invalid, a polynomial. 

Asphondylia rudbeckiaeconspicua Osten Sacken 1878: 5. USA: Pennsylvania; T; G; USNM. 

Asphondylia conspicua Beutenmüller 1907a: 387, error for rudbeckiaeconspicua. 

ruebsaameni. Hungary. Ferula heuffelii (Apiaceae). 

Asphondylia ruebsaameni (as rübsaameni) Kertész 1898: 245. Hungary; ST; ♂,P; ?. 

ruelliae. El Salvador. Ruellia albicaulis (Acanthaceae).

129 

Asphondylia ruelliae Möhn 1959: 347. El Salvador: Santa Ana, bank of L. Coatepeque; HT; ♂; SMNS. 

rutae. Italy. Ruta graveolens (Rutaceae). 

Asphondylia rutae Kieffer 1909b: 26. Italy; T; G; lost?. 

salictaria. USA (Indiana). Salix sp. (Salicaceae). 

Asphondylia salictaria Felt 1907d: 16. USA: Indiana, Pleasantville; HT; ♀; USNM. 

salsolae. Namibia. Salsola aphylla (Chenopodiaceae). 

Asphondylia salsolae Rübsaamen 1908c: 253. Namibia: Gaiaub Desert; ST; P,L; ZMHB. 

salsolarum. Tunisia. Salsola sp. (Chenopodiaceae). 

Asphondylia salsolae Kieffer 1912e: 77, jun. prim. hom. salsolae Rübsaamen. Tunisia: Gabès; ST; ♀,P; lost?. 

Asphondylia salsolarum Kieffer 1913g: 48, new name for Asphondylia salsolae Kieffer. 

salvadorensis. El Salvador. Hymenostephium cordatum (Asteraceae). 

Asphondylia salvadorensis Möhn 1959: 358. El Salvador: San Salvador; HT; ♂; SMNS. 

*salviaflorae. Russia (Europe). Salvia stepposa; Salvia spp. (Lamiaceae). 

Asphondylia salviaflorae Fedotova 2003b: 976. Europe: Samara Oblast, 30 km SE Samara, nr Ust'Kinelsk; HT; ♂; 

ZMAS. 

sambuci. USA (District of Columbia). Sambucus sp. (Caprifoliaceae). 

Asphondylia sambuci Felt 1908b: 377. USA: District of Columbia; HT; ♀; USNM. 

sanctipetri. Brazil (São Paulo). Didymopanax morototoni (Araliaceae). 

Asphondylia sanctipetri Urso-Guimarães & Amorim 2002: 562. Brazil: São Paulo, Ribeirão Preto, Fazenda 

Palmeiras; HT; ♂; MZSP. 

*sarcocorniae. Australia (Victoria). Sarcocornia quinqueflora (Chenopodiaceae). 

Asphondylia sarcocorniae Veenstra-Quah & Kolesik 2007: 203. Australia: Victoria, French Island Natl Pk, Gartside; 


sarothamni. Widespread Europe; Algeria. Cytisus scoparius (Fabaceae). 

Cecidomyia sarothamni Loew 1850a: 38. Poland; ST; ♂,♀; ZMHB?. 

Asphondylia mayeri Liebel 1889a: 266. France: Moselle, Karlingen & Bitche; ST; ♂,♀,P,L; lost?. 

scopariae. El Salvador. Scoparia dulcis (Scrophulariaceae). 

Asphondylia scopariae Möhn 1959: 345. El Salvador: Chalatenango, E Santa Barbara; HT; ♂; SMNS. 

scrophulariae. Widespread Europe. Scrophularia canina (Scrophulariaceae). 

Cecidomyia scrophulariae Schiner 1856: 220. Austria; ST; ♂,♀,P; NHMW. 

Asphondylia scrophulariae Tavares 1907a: 50, jun. sec. hom. scrophulariae Schiner. Portugal: Algarve, Portimão, & 

vic. Tavira; ST; ♂,♀,P; TAVC. 

Asphondylia scrophularina Tavares 1919a: 20, new name for Asphondylia scrophulariae Tavares. 

sennae. Brazil (Rio de Janeiro). Senna bicapsularis (Fabaceae). 

Asphondylia sennae Maia & Couri 1992: 656. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♀; QBUM. 

serjaniae. El Salvador. Serjania goniocarpa (Sapindaceae). 

Asphondylia serjaniae Möhn 1959: 322. El Salvador: San Salvador; HT; ♂; SMNS. 

serpylli. Widespread Europe. Thymus serpyllum (Lamiaceae). 

Asphondylia serpylli Kieffer 1898f: 20. France: Moselle, & Austria; T; G; lost?. 

Asphondylia thymi Kieffer 1898f: 59. France: Moselle; T; ♀; lost?. 

Asphondylia proxima Kieffer 1909b: 30. France; T; LT; lost?. 

*serrata. Brazil (Minas Gerais). Vanilosmopsis erythropappa (Asteraceae). 

Asphondylia serrata Maia 2004a: 71. Brazil: Minas Gerais, Tiradentes, Serra de São José; HT; ♂; MNRJ. 

sesami. India (widespread), Nigeria, Sudan, Uganda. Sesamum indicum; S. orientale (Pedaliaceae). 

Asphondylia sesami Felt 1916a: 31, India: Tamil Nadu, Coimbatore; ST; ♂,♀,P; NYSM. 

Asphondylia sesamiae Rao 1956a: 70. missp. of sesami. 

setosa. Kenya. 

Asphondylia setosa Kieffer 1913h: 92. Kenya: Likoni, nr Mombasa; HT; ♀; MNHN. 

shepherdiae. USA (Colorado). Shepherdia canadensis (Eleagnaceae). 

Asphondylia shepherdiae Felt 1916f: 414. USA: Colorado, Lake Eldora; ST; ♂,♀,P; NYSM. 

siccae. Jamaica. Phyllanthus distichus (Euphorbiaceae). 

Asphondylia siccae Felt 1908b: 376. Jamaica; ST; ♂,♀; NYSM.

130 

sidae. Colombia. Sida acuta; S. rhombifolia (Malvaceae). 

Asphondylia sidae Wünsch 1979: 36. Colombia: Magdalena Prov., Cañaverales; HT; ♂; SMNS. 

silicula. USA (Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia silicula Gagné 1990c: 668. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

smilacinae. USA (Maryland, Virginia). Smilacina racemosa (Liliaceae). 

Asphondylia smilacinae Felt 1907d: 17. USA: District of Columbia; T; ♂; USNM. 

solani. Mozambique. Solanum sp. (Solanaceae). 

Asphondylia solani Tavares 1908: 170. Mozambique; ST; ♂,♀,P; TAVC. 

solidaginis. Widespread eastern Nearctic. Solidago canadensis (Asteraceae). 

Asphondylia solidaginis Beutenmüller 1907b: 305. USA: New York, New York, Bronx Pk & Staten I., New Jersey, 

Fort Lee, & North Carolina, Buncombe Co., Valley of the Black Mts; ST; ♂,♀; AMNH. 

Cecidomyia bifolia Stebbins 1910: 49. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

*sophorae. Kazakhstan. Vexibia alopecuroides (Fabaceae) 

Asphondylia sophorae Fedotova 1988c: 45. Kazakhstan: western Kazakhstan, E of Darinskoe; HT; ♂; ZMAS. 

sphaera. Japan (Honshu to Okinawa). Ligustrum ibota; Ligustrum spp. (Oleaceae), Rhus succedanea, R. sylvestris 

(Anacardiaceae). 

Asphondylia sphaera Monzen 1937: 189. Japan; NT (Yukawa & Miyamoto 1979: 104; Saitama Pref., Yorii); ♂; 

KUEC. Ref.: Uechi & Yukawa 2006 (alternative hosts, biol.). 

*stachidiflorae. Russia (Europe). Stachis recta (Lamiaceae). 

Asphondylia stachidiflorae Fedotova 2003b: 979. Russia: Samara Oblast, 30 km SE Samara, nr Ust'Kinelsk; HT; ♂; 

ZMAS. 

stachytarpheta. Trinidad. Stachytarpheta cayennensis (Verbenaceae). 

Asphondylia stachytarpheta Barnes 1932b: 482. Trinidad; ST; ♂,♀; BMNH. 

stefanii. France, Italy, Malta. Diplotaxis tenuifolia (Brassicaceae). 

Asphondylia stefanii Kieffer 1898f: 59. Italy: Palermo; T; ♂; lost?. 

strobilanthi. Indonesia (Java). Strobilanthus cernuus (Acanthaceae). 

Asphondylia strobilanthi Felt 1921h: 150. Indonesia: Java, Tjibodas; ST; ♂,P,L; NYSM. 

struthanthi. Brazil (Ceará). Struthanthus sp. (Loranthaceae). 

Asphondylia struthanthi Rübsaamen 1916a: 432. Brazil: Ceará, Serra do Baturité; LT (Möhn 1973: 9); ♂; ZMHB. 

sturtiana. Australia (Queensland). Solanum sturtianum (Solanaceae). 

Asphondylia sturtiana Kolesik 2000a: 32. Australia: Queensland, Gregory Hwy, 52 km S Charters Towers; HT; ♂; 

ANIC. 

sulphurea. Brazil (Rio Grande do Sul). Poss. Smilax sp. (Smilacaceae). 

Asphondylia sulphurea Tavares 1909: 14. Brazil: Rio Grande do Sul, vic. São Leopoldo; HT; ♂; TAVC. 

swaedae. Southern Europe. Suaeda vermiculata (Chenopodiaceae). 

Asphondylia swaedae Kieffer 1909b: 30. Southern Europe; T; G; lost?. 

Asphondylia suaedae Skuhravá 1986: 213, missp. of swaedae. 

swaedicola. Argentina (Mendoza, San Juan). Suaeda divaricata (Chenopodiaceae). 

Asphondylia swaedicola Kieffer & Jörgensen 1910: 435. Argentina: Mendoza, La Paz, Potrerillos, & San Ignacio, & 

San Juan, nr Cancete; ST; ♂,♀,P; lost. 

tabernaemontanae. El Salvador. Tabernaemontana amygdalifolia (Apocynaceae). 

Asphondylia tabernaemontanae Möhn 1959: 333. El Salvador: Sonsonate, N Los Cóbanos; HT; ♂; SMNS. 

talini. EL Salvador. Talinum paniculatum (Portulacaceae). 

Asphondylia talini Möhn 1959: 305. El Salvador: La Libertad, Cumbre, Hacienda Chilata; HT; ♂; SMNS. 

tavaresi. Portugal. Lavandula stoechas (Lamiaceae). 

Ischnonyx tavaresi Rübsaamen 1916c: 7. Portugal: T; ♂; TAVC?. 

tectonae. India (Tamil Nadu). Tectona grandis (Verbenaceae). 

Asphondylia tectonae Mani 1974: 61. India: Tamil Nadu, Madras, Top Slip; HT; ♂; FRDD. 

tephrosiae. (Delhi, Tamil Nadu). Tephrosia candida (Fabaceae). 

Asphondylia tephrosiae Mani 1944: 152. India: Delhi; HT; ♂; INPC. 

thalictri. USA (New York, New Hampshire, Massachusetts). Thalictrum sp. (Ranunculaceae). 

Asphondylia thalictri Felt 1911l: 473. USA: Massachusetts, Magnolia; T; G; NYSM.

131 

thevetiae. El Salvador. Thevetia plumeriaefolia (Apocynaceae). 

Asphondylia thevetiae Möhn 1959: 337. El Salvador: San Vicente, km 54, SE Santo Domingo; HT; ♂; SMNS. 

tithoniae. El Salvador. Tithonia rotundifolia (Asteraceae). 

Asphondylia tithoniae Möhn 1960c: 200. El Salvador: La Libertad, E Sitio del Nino; HT; ♂; SMNS. 

*tonsura. Australia (South Australia). Enchylaena tomentosa (Chenopodiaceae). 

Asphondylia tonsura Kolesik & Veenstra-Quah 2008: 218. Australia: South Australia, Eyre Peninsula, Minnipa; HT; 

♂; SAMA. 

tournefortiae. El Salvador, Brazil (Amazonas). Tournefortia angustiflora; T. volubilis (Boraginaceae). 

Asphondylia tournefortiae Rübsaamen 1916a: 433. Brazil: Amazonas, Auristella & S. Francisco on R. Acre; ST; 

♀,P,L; ZMHB. 

trabuti. Algeria, Italy. Solanum tuberosum (Solanaceae). 

Asphondylia trabuti Marchal 1896: 97. Algeria; ST; ♀,P; MNHN?. 

trichiliae. El Salvador. Trichilia sp. (Meliaceae). 

Asphondylia trichiliae Möhn 1959: 318. El Salvador: La Libertad, NE Sitio del Nino, W lava fields; HT; ♂; SMNS. 

trichocecidarum. India (Tamil Nadu). Acacia leucophloea (Fabaceae). 

Asphondylia trichocecidarum Mani 1934b: 413. India: Tamil Nadu, Thanjavur; HT; ♂; ZSCI. 

trixidis. El Salvador. Trixis radialis (Asteraceae). 

Asphondylia trixidis Möhn 1959: 362. El Salvador: La Libertad, NE Sitio del Nino, W lava fields; HT; ♂; SMNS. 

ulei. Brazil (Rio de Janeiro). Mikania sp. (Asteraceae). 

Asphondylia ulei Rübsaamen 1908a: 172. Brazil: Rio de Janeiro, Palmeiras; LT (Möhn 1973: 7); LT; ZMHB. 

ulicis. UK, Portugal. Ulex europaeus (Fabaceae). 

Asphondylia ulicis Trail 1873: 172. UK: Scotland, Aberdeen; T; G; ?. 

utriculae. India (Tamil Nadu). Dichrostachys cineria (Fabaceae). 

Asphondylia utriculae Mani 1934b: 410. India: Tamil Nadu, Thanjavur; HT; ♀; ZSCI. 

vavilovi. Turkmenistan. Salsola gemmascens (Chenopodiaceae). 

Asphondylia vavilovi Fedotova 1990d: 62. Turkmenistan: southern Ustyurt, Kaplankyrskiï Res., 300 km W Tashauz; 


verbasci. Widespread Europe, Algeria. Verbascum sp.; Verbascum spp. (Scrophulariaceae). 

Cecidomyia verbasci Vallot 1827: 92. France; ST; A,P; ?. 

Asphondylia dufouri Kieffer 1902b: 546. France; T; LT; lost?. 

verbenae. USA (Texas). Verbena hallei (Verbenaceae). 

Asphondylia verbenae Felt 1935a: 6. USA: Texas, Austin; ST; ♂,♀; NYSM. 

vernoniae. USA (Virginia, Florida). Vernonia noveboracensis (Asteraceae). 

Asphondylia vernoniae Felt 1908b: 377. USA: Virginia, S; ♂,♀; USNM. 

*vesicaria. Australia (South Australia). Atriplex vesicaria (Chenopodiaceae). 

Asphondylia vesicaria Kolesik & Veenstra-Quah 2008: 214. Australia: South Australia, Eyre Peninsula, Ceduna; HT; 

♂; SAMA. 

villosa. USA (Arizona). Larrea tridentata (Zygophyllaceae). 

Asphondylia villosa Gagné 1990c: 669. USA: Arizona, Saguaro Natl Mon., East, 20 mi. E Tucson; HT; LT; USNM. 

vincenti. El Salvador, St. Vincent. Jussiaea linifolia, J. suffruticosa; J. angustifolia (Onagraceae). 

Asphondylia vincenti Felt 1911d: 109. St. Vincent; ST; ♂,♀,P,L; NYSM. 

vitea. Philippines. Cissus trifolia (Vitaceae). 

Asphondylia vitea Felt 1918c: 284. Philippines: Luzon, Manila; ST; ♂,♀; NYSM. 

viticola. Indonesia (Java). Vitis trifolia (Vitaceae). 

Asphondylia viticola Kieffer & Docters van Leeuwen-Reijnvaan 1910: 124. Indonesia: Java, Salatiga; ST; ♂,♀,P; 

lost?. 

waltheriae. El Salvador. Waltheria americana (Sterculiaceae). 

Asphondylia waltheriae Möhn 1959: 328. El Salvador: Santa Ana, E Tablón del Coco; HT; ♂; SMNS. 

websteri. USA (California, Arizona, Oklahoma, Texas), Mexico, Honduras, Dominican Republic, Guadeloupe. 

Medicago sativa; Cyamopsis tetragonoloba, Mimosa sp., Parkinsonia spp. (Fabaceae), & Simmondsia 

chinensis (Simmondsiaceae). 

Asphondylia websteri Felt 1917d: 562. USA: Arizona, Tempe; LT (Gagné & Wuensche 1986: 248); ♀; USNM.

132 

Misident as miki Wachtl in Webster 1912. 

xanthii. USA (Texas). Xanthium speciosum (Asteraceae). 

Asphondylia xanthii Felt 1936b: 231. USA: Texas, Navasota; ST; ♂,♀; USNM. 

ximeniae. El Salvador. Ximenia americana (Olacaceae). 

Asphondylia ximeniae Möhn 1959: 301. El Salvador: Dept. La Libertad, NE Sitio del Nino, W lava fields; HT; ♂; 

SMNS. 

xylosmatis. El Salvador. Xylosma flexuosum (Flacourtiaceae). 

Asphondylia xylosmatis Möhn 1959: 331. El Salvador: Santa Ana, SE Candelaria, Hacienda Cortez; HT; ♂; SMNS. 

yukawai. Colombia. Arenaria sp. (Caryophyllaceae), Melochia spp. (Sterculiaceae). 

Asphondylia yukawai Wünsch 1979: 47. Colombia: Magdalena Prov., Bahia Nenguange; HT; ♂; SMNS. 

yushimai. Japan (Honshu, Shikoku, Kyushu), China, Indonesia. Glycine max (Fabaceae); Osmanthus heterophyllus 

(Oleaceae), Prunus zippeliana (Rosaceae). 

Asphondylia yushimai Yukawa & Uechi 2003a: 77, nomen nudum. 

Asphondylia yushimai Yukawa & Uechi 2003b: 265. Japan: Kyushu, Kagoshima City, Kamifukumoto; HT; ♂; 

KUEC. Ref.: Uechi, Yukawa & Usuba. 2005 (host alternation). 

zacatechichi. El Salvador. Calea zacatechichi (Asteraceae). 

Asphondylia zacatechichi Möhn 1960c: 199. El Salvador: San Vicente, SE Santo Domingo, km 54; HT; LT; SMNS. 

zexmeniae. El Salvador. Zexmenia frutescens (Asteraceae). 

Asphondylia zexmeniae Möhn 1960c: 201. El Salvador: San Salvador; HT; P; SMNS. 

ziziphorae. Kazakhstan. Ziziphora clinopodioides (Lamiaceae). 

Asphondylia ziziphorae Fedotova 1985b: 31. Kazakhstan: Talasskiï Alatau, Tyul'kubas valley; HT; ♂; ZKAS. 

ASPHOTROPHIA 

Asphotrophia Barnes 1928a: 23. Type species, fici Barnes (orig. des.). 

Anjeerodiplosis Mani 1938: 333. Type species, peshawarensis Mani (orig. des.). 

Udumbaria Grover 1963: 37. Type species, nainiensis Grover (mon.). 

Oriental; 3 spp., all from Ficus; Cecidomyiidi, unplaced to tribe. 

fici. Philippines. Ficus mora (Moraceae). 

Asphotrophia fici Barnes 1928a: 23. Philippines: Luzon, Laguna Prov., Los Baños; HT; ♀; BMNH. 

nainensis. India (Uttar Pradesh). Ficus carica (Moraceae). 

Udumbaria nainensis Grover 1963: 38. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

peshawarensis. Pakistan. Ficus carica (Moraceae). 

Anjeerodiplosis peshawarensis Mani 1938: 333. Pakistan: Peshawar; HT; ♀; INPC. 

ASPHOXENOMYIA 

Asphoxenomyia Felt 1927c: 382. Type species, smilacis Felt (orig. des.). 

Oriental; 1 sp.; Cecidomyiidi, Asphondyliini, Schizomyiina. 

smilacis. Indonesia (Java). Smilax modesta (Smilacaceae). 

Asphoxenomyia smilacis Felt 1927c: 382. Indonesia: Java, Tjibodas, Mt Gede; ST; ♂,♀; NYSM. 

ASTERALOBIA 

Asteralobia Kovalev 1964: 419. Type species, doellingeriae Kovalev (orig. des.). 

Euasteralobia Kovalev 1964: 430, as subg. of Asteralobia. Type species, Asteralobia calathidiphaga Kovalev 

(mon.).

133 

Eastern Palearctic; 12 spp. on various plant families; Cecidomyiidi, Asphondyliini, Schizomyiina. This genus could 

possibly be subsumed under Schizomyia, q.v. for remarks. Ref.: Tokuda, Tabuchi, et al. 2004 (ecol., biol. of spp. on 

Ilex). 

asteris. Russia (Far East). Aster tataricus; A. scaber (Asteraceae). 

Asteralobia asteris Kovalev 1964: 425. Russia: Primorskiï Terr., Lyanchikhe R., nr Vladivostok; HT; ♂; KOVC. 

Ref.: Tokuda et al. 2003 (descr. larva). 

calathidiphaga. Russia (Far East). Cacalia hastata; Senecio sp. (Asteraceae). 

Asteralobia calathidiphaga Kovalev 1964: 430. Russia: Primorskiï Terr., Shufan R.; HT; ♂; KOVC. 

*clematidis. Russia (Far East). Clematis brevicaudata (Rosaceae). 

Asteralobia clematidis Fedotova 2002c: 4. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

doellingeriae. Russia (Far East), Japan (Honshu). Aster scaber (Asteraceae). 

Asteralobia doellingeriae Kovalev 1964: 422. Russia: Primorskiï Terr., Okeanskaya Station, Bogataya Crest; HT; ♂; 

KOVC. Ref.: Tokuda et al. 2003 (descr. larva; distr.). 

Asteralobia doellingerae Fedotova & Kovalev 2001: 527, missp. of doellingeriae. 

humuli. Japan (Honshu, Kyushu), Republic of Korea. Humulus japonicus; Humulus lupulus (Cannabaceae). 

Asphondilia humuli Shinji 1939h: 224. Japan; NT (Yukawa 1983: 100; Japan: Honshu, Kagoshima City, Toso); ♀; 

KUIC. 

kovalevi. Russia (Far East). Patrinia scabiosifolia; P. rupestris (Valerianaceae). 

Asteralobia patriniae Kovalev 1964: 428, jun. sec. hom. patriniae Shinji. Russia: Primorskiï Terr., Chapigou R.; HT; 

♂; KOVC. 

Asteralobia kovalevi Skuhravá 1986: 215, new name for Asteralobia patriniae Kovalev. 

patriniae. Japan (Hokkaido to Shikoku). Patrinia villosa (Valerianaceae). 

Schizomyia patriniae Shinji 1938i: 372. Japan; NT (Yukawa 1983: 106; Japan: Ishikawa Pref., Kanazawa City, 

Iôzan); ♂; KUIC. 

Asphondylia partriniae Shinji 1944: 376, missp. of patriniae. 

Parasphondylia patrici Shinji 1944: 44, missp. of patriniae. 

sasakii. Japan (Hokkaido to Kyushu). Ilex crenata; I. maximowicziana (Aquifoliaceae). 

Parasphondylia sasakii Monzen 1937: 191. Japan: Iwate, Tokyo, Toyama, Fukui, Wakayama, Yamaguchi, Fukuoka 

Prefs; ST; ♂,♀; KUEC. Ref.: Tokuda, Tabuchi et al. 2004 (redescr., host range). 

Shizomyia ilexicola Shinji 1938k: 239. Japan: Honshu; ST; ♂,♀; lost. 

solidaginis. Russia (Far East). Solidago pacifica (Asteraceae). 

Asteralobia solidaginis, as subsp. of asteris, Kovalev 1964: 427. Russia: Primorskiï Terr., Bogataya crest, nr 

Vladivostok; HT; ♂; KOVC. 

Asteralobia asterisolidaginis Skuhravá 1986: 215, unnec. new name for Asteralobia solidaginis Kovalev. 

soyogo. Japan (Honshu, Kyushu, Nansei Is.). Ilex pedunculosa; I. chinensis, I. integra, I. leucoclada (Aquifoliaceae). 

Schizomyia soyogo Kikuti 1939: 239. Japan: Honshu, Kyoto, Higasiyama, Kwazan; HT; ♂; Kikuti. Ref.: Tokuda, 

Tabuchi et al. 2004 (redescr., host range). 

*spiraeae. Russia (Far East). Spiraea salicifolia (Rosaceae). 

Asteralobia spiraeae Fedotova 2002c: 6. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

*veronicastrum. Russia (Far East). Veronicastrum sibiricum (Scrophulariaceae). 

Asteralobia veronicastrum Fedotova 2002c: 8. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

ASTEROMYIA 

Asteromyia Felt 1910d: 348. Type species, Cecidomyia carbonifera Felt (orig. des.) = euthamiae Gagné. 

Nearctic; 9 spp., all in leaf or stem galls of Astereae (Asteraceae); Lasiopteridi, Alycaulini. Ref.: Gagné 1968 (rev.).

134 

carbonifera. Widespread Nearctic. Solidago sp.; known from almost all Solidago spp. (Asteraceae). 

Cecidomyia carbonifera Osten Sacken 1862: 195. USA: District of Columbia; ST; G; USNM. 

Choristoneura abnormis Felt 1907a: 14. USA: New York, Albany; HT; ♀; NYSM. 

Choristoneura albomaculata Felt 1907a: 14. USA: New York, Albany; HT; ♀; NYSM. 

Choristoneura convoluta Felt 1907a: 14. USA: New York, Albany; HT; ♀; NYSM. 

Lasioptera rubra Felt 1907a: 7. USA: New York, Albany; HT; ♂; NYSM. 

Lasioptera tuberculata Felt 1907a: 7. USA: New York, Albany; HT; ♂; NYSM. 

Choristoneura flavolunata Felt 1907c: 154. USA: New York, Albany; HT; ♀; NYSM. 

Lasioptera rosea Felt 1907c: 152. USA: New York, Albany; HT; ♀; NYSM. 

Baldratia flavoanulata Felt 1908b: 329. USA: New York, West Nyack; HT; ♀; NYSM. 

Baldratia flavoscuta Felt 1908b: 328. USA: New York, Albany; HT; ♀; NYSM. 

Baldratia socialis Felt 1908b: 328. USA: New York, Albany; HT; ♀; NYSM. 

Baldratia squarrosae Felt 1908b: 329. USA: New York, Rhinecliff; HT; ♀; NYSM. 

chrysothamni. Widespread western Nearctic. Chrysothamnus sp.; Chrysothamnus spp., Haplopappus spp. 

(Asteraceae). 

Asteromyia chrysothamni Felt 1918a: 214. USA: Colorado, nr Boulder; LT (Gagné 1968b: 28); ♀; NYSM. 

clarkei. Widespread eastern Nearctic. Aster sp.; Aster spp. (Asteraceae). 

Lasioptera clarkei Felt 1909b: 287. USA: Massachusetts, Magnolia; LT (Gagné 1968b: 25); ♂; NYSM. 

Asteromyia nigrina Felt 1911m: 481. USA: Massachusetts, Magnolia; LT (Gagné 1968b: 25); ♀; NYSM. 

Asteromyia sylvestris Felt 1915e: 228. USA: New York, Mount Kisco; LT (Gagné 1968b: 25); ♀; NYSM. 

euthamiae. Widespread eastern Nearctic. Euthamia graminifolia; Euthamia spp. (Asteraceae). 

Lasioptera carbonifera Felt 1906b: 116, jun. sec. hom. carbonifera Osten Sacken. USA: New York, Albany & 

Kenwood; ST; ♂,♀,P,L; NYSM. 

Asteromyia euthamiae Gagné 1968b: 28. USA: Maryland, Montgomery Co., Cabin John Pk; HT; ♀; USNM. 

grindeliae. USA (California, Minnesota, Texas, Maryland). Grindelia robusta; Grindelia spp. (Asteraceae). 

Asteromyia grindeliae Felt 1912h: 149. USA: California, nr Santa Barbara; LT (Gagné 1968b: 27); LT; NYSM. 

gutierreziae. Western Nearctic. Gutierrezia sarothrae; G. microcephala, Baccharis angustifolia, Gymnosperma 

glutinosum, Isocoma palmeri (Asteraceae). 

Asteromyia gutierreziae Felt 1916e: 179. USA: Utah, nr Salt Lake City; LT (Gagné 1968b: 30); ♀; NYSM. 

laeviana. Widespread Nearctic. Aster laevis; many Aster spp. (Asteraceae). 

Choristoneura laeviana Felt 1907a: 12. USA: New York, Albany; HT; ♂; NYSM. 

Choristoneura basalis Felt 1907a: 13. USA: New York, Albany; HT; ♂; NYSM. 

Choristoneura paniculata Felt 1907a: 13. USA: New York, Albany; HT; ♂; NYSM. 

Lasioptera asterifoliae Beutenmüller 1907a: 395. USA: North Carolina, Buncombe Co., Valley of the Black Mts; LT 

(Gagné 1968b: 22); ♂; AMNH. 

Choristoneura helena Felt 1907d: 7. USA: New York, Highland; HT; ♀; NYSM. 

Baldratia divaricata Felt 1908b: 330. USA: New York, Albany; LT (Gagné 1968b: 22); ♀; NYSM. 

Baldratia flavomaculata Felt 1908b: 329. USA: New York, Albany; HT; ♀; NYSM. 

Baldratia fuscoanulata Felt 1908b: 329. USA: New York, Highland; LT (Gagné 1968b: 23); ♀; NYSM. 

Baldratia pustulata Felt 1908b: 328. USA: New York, Albany; HT; ♂; NYSM. 

Baldratia dumosae Felt 1909b: 286. USA: Massachusetts, Annisquam; HT; ♀; NYSM. 

Baldratia nitida Felt 1909b: 286. USA: New York, Albany; HT; ♀; NYSM. 

Baldratia vesiculosa Felt 1909b: 286. USA: Massachusetts, Magnolia; LT (Gagné 1968b: 23); ♀; NYSM. 

Baldratia waldorfi Felt 1909b: 286. USA: New York, Albany; LT (Gagné 1968b: 23); ♀; NYSM. 

Asteromyia asterifolia Felt 1910d: 349, missp. of asterifoliae. 

Asteromyia marginata Felt 1910d: 349. USA: New York, Albany; HT; ♀; NYSM. 

Asteromyia reducta Felt 1911m: 481. USA: Massachusetts, Magnolia; LT (Gagné 1968b: 23); ♀; NYSM. 

Asteromyia fuscoannulata Felt 1925c: 190, unjust. emend. of fuscoanulata. 

modesta. Widespread eastern Nearctic, Brazil (Minas Gerais), Argentina (Buenos Aires). Conyza canadensis; Erigeron 

strigosus (Asteraceae). 

Choristoneura modesta Felt 1907c: 163. USA: New York, Albany; LT (Gagné 1968b: 27); ♀; NYSM. 

tumifica. USA (Minnesota, New York, Massachusetts, West Virginia, Maryland, North Carolina). Solidago rugosa; S.

135 

canadensis (Asteraceae). 

Lasioptera tumifica Beutenmüller 1907a: 394. USA: New York, New York, Staten I.; LT (Gagné 1969d: 1354); ♀; 

AMNH. 

ASTICTONEURA 

Astictoneura Gagné 1969d: 1357. Type species, Lasioptera muhlenbergiae Marten (orig. des.). 

Nearctic; 1 sp.; Lasiopteridi, Alycaulini. 

muhlenbergiae. USA (New Mexico, South Dakota, Iowa, Illinois, Connecticut). Muhlenbergia sp. (Poaceae). 

Lasioptera muhlenbergiae Marten 1893: 155. USA: Illinois, Champaign; ST; ♂,♀,P,L; ?. 

Lasioptera carbonitens Cockerell 1902: 183. USA: New Mexico, Las Valles; LT (Gagné 1969d: 1357); ♀; USNM. 

ASTRAGALOMYIA 

Astragalomyia Fedotova 1993j: 75. Type species, buzaczensis Fedotova (orig. des.). 


buzaczensis. Kazakhstan. Astragalus longiflorus (Fabaceae). 

Astragalomyia buzaczensis Fedotova 1993j: 76. Kazakhstan: Buzachi Peninsula, 12 km SW Shebir; HT; ♂; ZMAS. 

ASTRODIPLOSIS 

Astrodiplosis Felt 1913d: 217. Type species, speciosa Felt (orig. des.). 


speciosa. Guatemala. Cissus sp. (Vitaceae). 

Astrodiplosis speciosa Felt 1913d: 218. Guatemala: Puerto Barrios; ST; ♂,♀,P,L; NYSM. 

ATHIDIPLOSIS 

Athidiplosis Gagné 1993c: 108. Type species, walteri Gagné (orig. des.). 

Afrotropical; 2 spp.; Cecidomyiidi, unplaced to tribe. 

bullata. Kenya. Acacia mellifera (Fabaceae). 

Athidiplosis bullata Gagné 1993c: 109. Kenya: Kima, nr Sultan Hamid; HT; ♂; USNM. 

walteri. Kenya. Acacia mellifera (Fabaceae). 

Athidiplosis walteri Gagné 1993c: 109. Kenya: Yatta; HT; ♂; USNM. 

ATOPODIPLOSIS 

Atopodiplosis Gagné 1997a: 119. Type species, wirthi Gagné (orig. des.). 


wirthi. Dominica. 

Atopodiplosis wirthi Gagné 1997a: 122. Dominica: Clarke Hall; HT; ♀; USNM.

ATOLASIOPTERA 

Atolasioptera Möhn 1975: 8. Type species, calida Möhn (orig. des.). 

Neotropical; 1 sp.; Lasiopteridi, Alycaulini. 

calida. El Salvador. Achatocarpus nigricans (Phytolaccaceae). 

Atolasioptera calida Möhn 1975: 8. El Salvador: E La Libertad, San Diego; HT; ♀; SMNS. 

ATRAPHAXIOLA 

Atraphaxiola Fedotova 1986b: 629. Type species, bogutensis Fedotova (orig. des.). 

Palearctic; 1 sp.; Lasiopteridi, Lasiopterini. 

bogutensis. Kazakhstan. Atraphaxis virgata (Polygonaceae). 

Atraphaxiola bogutensis Fedotova 1986b: 631. Kazakhstan: northern Tyan'-Shan', Kyrtogaï canyon, 30 km N 

Chundzha; HT; ♀; ZMAS. 

ATRICHOSEMA 

Atrichosema Kieffer 1904b: 347. Type species, aceris Kieffer (mon.). 


aceris. UK, France, Germany, Czechoslovakia, Yugoslavia. Acer campestre (Aceraceae). 

Atrichosema aceris Kieffer 1904b: 348. France; ST; ♀,P,L; lost?. 

*AURITADIPLOSIS 

Auritadiplosis Fedotova 2004f: 110. Type species, rhombea Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Clinodiplosini. 

*rhombea. Russia (Far East). 

Auritadiplosis rhombeus Fedotova 2004f: 110. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

AUSTRODIPLOSIS 

Austrodiplosis Brèthes 1914: 154. Type species, argentina Brèthes (orig. des.). 


argentina. Argentina (Buenos Aires). 

Austrodiplosis argentina Brèthes 1914: 154. Argentina: Buenos Aires, General Urquiza; HT; ♀; MACN. 

AUSTROLAUTHIA 

Austrolauthia Brèthes 1914: 153. Type species, spegazzinii Brèthes (orig. des.). 

136

Neotropical; 1 sp.; Lasiopteridi, Dasineurini. 

spegazzinii. Argentina (Buenos Aires). 

Austrolauthia spegazzinii Brèthes 1914: 153. Argentina: Buenos Aires, La Plata; ST; ♂,♀; MACN. 

Austrolauthia spegazzini Gagné 1968d: 88, missp. of spegazzinii. 

AUSTROLOPESIA 

Austrolopesia Kolesik 1999a: 32. Type species, melaleucae Kolesik (orig. des.). 

Australasian; 1 sp.; Cecidomyiidi, Lopesiini. 

melaleucae. Australia (South Australia). Melaleuca halmaturorum (Myrtaceae). 

Austrolopesia melaleucae Kolesik 1999a: 33. Australia: South Australia, Coorong Natl Pk, Loop Road; HT; ♂; 

SAMA. 

AUTODIPLOSIS 

Eudiplosis Tavares 1916a: 51, preocc. Kieffer 1894. Type species, parva Tavares (orig. des.). 

Autodiplosis Tavares 1920: 106, new name for Eudiplosis Tavares. 

Neotropical; 1 sp.; Cecidomyiidi, Clinodiplosini. 

parva. Brazil (Bahia). Undet. sp. of Faboideae (Fabaceae). 

Eudiplosis parva Tavares 1916a: 51. Brazil: Bahia, Salvador; ST; ♂,♀,L; TAVC. 

BACCHAROMYIA 

Baccharomyia Tavares 1917b: 129. Type species, ramosina Tavares (orig. des.). 

Neotropical; 4 spp., all on Baccharis; Lasiopteridi, Alycaulini. Ref.: Gagné 1994a (review). 

cordobensis. Argentina (Córdoba). Baccharis coridifolia (Asteraceae). 

Lasioptera cordobensis Kieffer & Jörgensen 1910: 363. Argentina: Córdoba, nr Arias & La Carlota; T; LT; lost. 

interrupta. Argentina (Mendoza). Baccharis subulata (Asteraceae). 

Lasioptera interrupta Kieffer & Jörgensen 1910: 375. Argentina: Mendoza, nr Chacras de Coria; ST; ♂,♀,P,L; lost. 

ornaticornis. Argentina (Mendoza). Baccharis salicifolia (Asteraceae). 

Lasioptera ornaticornis Kieffer & Jörgensen 1910: 368. Argentina: Mendoza, nr Chacras de Coria; ST; ♂,♀,P,egg; 

lost. 

ramosina. Brazil (Rio de Janeiro). Baccharis trimera (Asteraceae). 

Baccharomyia ramosina Tavares 1917b: 130. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♀,P,L; TAVC. 

BAEODIPLOSIS 

Baeodiplosis Kieffer 1913h: 111. Type species, tropicalis Kieffer (orig. des.). 


tropicalis. Tanzania. 

Baeodiplosis tropicalis Kieffer 1913h: 112. Tanzania: Tavéta; ST; ♀,egg; MNHN. 

BALDRATIA 

137

Baldratia Kieffer 1897c: 7. Type species, salicorniae Kieffer (orig. des.). 

Baldratiella Kieffer 1913g: 45. Type species, Baldratia hyalina Kieffer (orig. des.) = salicorniae Kieffer. 

Baldolatiola Shinji 1944: 217, missp. of Baldratiella. 

Baldolatia Shinji 1944: 217, missp. of Baldratia. 

Arafavilla Mamaev & Mirumian 1990b: 139. Type species, terteriani Mamaev & Mirumian (orig. des.). 

Palearctic; 38 spp., all on Chenopodiaceae; Lasiopteridi, Lasiopterini. Refs: Dorchin 2001 (descr.), Gagné 2004b 

(syn. note.). 

aegyptiaca. Egypt. Anabasis setifera (Chenopodiaceae). 

Baldratia aegyptiaca Möhn 1969: 184. Egypt: Wadi Feiran; HT; LT; SMNS. 

aelleni. Iran. Suaeda microphylla (Chenopodiaceae). 

Baldratia aelleni Möhn 1969: 172. Iran: Central Elbrus, Simin-Dascht; HT; LT; SMNS. 

algeriensis. Algeria. Salsola longifolia (Chenopodiaceae). 

Baldratia algeriensis Möhn 1969: 194. Algeria: Oran; HT; LT; SMNS. 

anabasidiramea. Kazakhstan. Anabasis cretacea (Chenopodiaceae). 

Baldratia anabasidiramea Fedotova 1991e: 51. Kazakhstan: eastern Mangyshlak basin, 50 km SE Novyï Uzen', nr 

Senek; HT; ♀; ZMAS. 

anabasis. Iran, Kazakhstan. Anabasis aphylla; A. brachiata, A. salsa (Chenopodiaceae). 

Baldratia anabasis Möhn 1969: 179. Iran: Kerman Prov., Kuh-Hasar; HT; LT; SMNS. 

araliensis. Kazakhstan. Salsola orientalis (Chenopodiaceae). 

Baldratia araliensis Fedotova 1992b: 45. Kazakhstan: NE Aral Sea, 5 km E Akesle; HT; ♀; ZMAS. 

arbustifolia. Kazakhstan. Salsola arbuscula (Chenopodiaceae). 

Baldratia arbustifolia Fedotova 1985a: 47. Kazakhstan: left bank of middle course of Ili R., 20 km NW Barandysu; 

HT; ♀; ZKAS. 

balchanensis. Turkmenistan. Salsola aucheri (Chenopodiaceae). 

Baldratia balchanensis Fedotova 1992d: 32. Turkmenistan: Bolshie Balkhany, nr Nebit-Dag; HT; ♀; ZMAS. 

caspica. Kazakhstan. Salsola lanata (Chenopodiaceae). 

Baldratia caspica Möhn 1969: 198. Kazakhstan: Transcaspian Region, Kisil Arwat; HT; LT; SMNS. 

chiwensis. Turkmenistan. Salsola chiwensis (Chenopodiaceae). 

Baldratia chiwensis Fedotova 1994d: 42. Turkmenistan: southern Ustyurt, Burchliburun escarpment, 300 km W 

Tashauza; HT; ♂; ZMAS. 

climacopterifolia. Kazakhstan. Climacoptera obtusifolia; C. affinis, C. crassa (Chenopodiaceae). 

Baldratia climacopterifolia Fedotova 1991e: 52. Kazakhstan: S L. Balkhash region, Tesik R. valley, 50 km E 

Akseuk; HT; ♀; ZMAS. 

dendroides. Turkmenistan. Salsola dendroides (Chenopodiaceae). 

Baldratia dendroides Fedotova 1992d: 32. Turkmenistan: Kugitang Mtn Range, nr Charshanga; HT; ♀; ZMAS. 

desertorum. Egypt. Anabasis setifera (Chenopodiaceae). 

Baldratia desertorum Möhn 1969: 190. Egypt: Feu Schel Wrab; HT; LT; SMNS. 

grandis. Middle East. Anabasis aphylla (Chenopodiaceae). 

Baldratia grandis Möhn 1969: 190. "Palestine, Mashetta"; HT; LT; SMNS. 

halimocnemiis. Kazakhstan. Halimocnemis sclerosperma (Chenopodiaceae). 

Baldratia halimocnemiis Fedotova 1992b: 43. Kazakhstan: NE Aral Sea, 14 km SE Saksaulskaya RR stn; HT; ♀; 

ZMAS. 

halogetonis. Egypt. Halogeton alopecuroides (Chenopodiaceae). 

Baldratia halogetonis Möhn 1969: 197. Egypt: Guari/Sinai, Ham't; HT; LT; SMNS. 

halophila. China (Xinjiang-Uygur Aut. Reg.). Anabasis phyllophora (Chenopodiaceae). 

Baldratia halophila Möhn 1969: 188. China: Xinjiang-Uygur Aut. Reg., Dzhungaria; HT; LT; SMNS. 

haloxyli. Jordan, Tunisia. Haloxylon articulatum (Chenopodiaceae). 

Baldratia haloxyli Möhn 1969: 178. Jordan: Esh-Shara, Wadi Musa; HT; LT; SMNS. 

jaxartica. Kazakhstan. Salicornia europaea (Chenopodiaceae). 

138

139 

Baldratia jaxartica Fedotova 1992b: 46. Kazakhstan: NE Aral Sea, 60 km SE Aralsk, nr Kamyshlybash RR stn; HT; 

♀; ZMAS. 

jordaniensis. Jordan. Salsola longifolia (Chenopodiaceae). 

Baldratia jordaniensis Möhn 1969: 196. Jordan: Jordan Valley; HT; LT; SMNS. 

kaplankyrica. Turkmenistan. Salsola richteri (Chenopodiaceae). 

Baldratia kaplankyrica Fedotova 1992k: 70. Turkmenistan: southern Ustyurt, Kaplankyrskiï Res., Burchliburun 

escarpment, 300 km W Tashauza; HT; ♀; ZMAS. 

kermanensis. Iran, Kazakhstan. Anabasis setifera (Chenopodiaceae). 

Baldratia kermanensis Möhn 1969: 183. Iran: Kerman; HT; LT; SMNS. 

kozlovi. Kazakhstan, Turkmenistan. Haloxylon ammodendron, H. aphyllum (Chenopodiaceae). 

Baldratia koslovi Marikovskij 1955b: 104, nomen nudum. 

Baldratia kozlovi Marikovskij 1955d: 311. Kazakhstan: Kzyl-Orda; ST; ♂,♀; ?. 

marikovskiji. Turkmenistan. Salsola crassa (Chenopodiaceae). 

Baldratia marikovskiji Möhn 1969: 192. Turkmenistan: Skobelewka, nr Askhabad; HT; LT; SMNS. 

nitrariacarpa. Kazakhstan. Salsola nitraria (Chenopodiaceae). 

Baldratia nitrariacarpa Fedotova 1991e: 53. Kazakhstan: Ustyurt plateau, Karakumy Desert, 38 km NE Sarykamys, 

Kyzyltan; HT; ♀; ZMAS. 

occulta. Israel. Suaeda monoica (Chenopodiaceae). 

Baldratia occulta Dorchin 2001: 568. Israel: Newe Zohar; HT; ♀; TAUI. 

orientalis. Jordan, Arabia. Anabasis ramosissima (Chenopodiaceae). 

Baldratia orientalis Möhn 1969: 182. Jordan: Wuste Ziza; HT; LT; SMNS. 

przewalskii. Widespread central Asia. Haloxylon ammodendron, H. aphyllum (Chenopodiaceae). 

Baldratia przewalskii Marikovskij 1955b: 104, nomen nudum. 

Baldratia przewalskii Marikovskij 1955d: 310. Kazakhstan: Ili R.; ST; ♂,♀; ?. 

salicorniae. UK, circum-Mediterranean, Eritrea. Salicornia fructicosa (Chenopodiaceae). 

Baldratia salicorniae Kieffer 1897c: 7. Italy; L (Dorchin & Freidberg 2008: 4); ♀; NYSM. Ref.: Dorchin & 

Freidberg 2008 (redescr.; biol.). 

Baldratia hyalina Kieffer 1912e: 74. Tunisia: Oasis de Nefta; ST; ♀,P; lost?. 

salsolae. Turkmenistan. Salsola aurantiaca (Chenopodiaceae). 

Baldratia salsolae Möhn 1969: 193. Turkmenistan: Krasnowodsk; HT; LT; SMNS. 

salsoliramea. Kazakhstan. Salsola paulsenii (Chenopodiaceae). 

Baldratia salsoliramea Fedotova 1991e: 48. Kazakhstan: lower reaches Ili R., 75 km N Bakanas; HT; ♀; ZMAS. 

seidlitziae. Iraq. Seidlitzia rosmarinus (Chenopodiaceae). 

Baldratia seidlitziae Möhn 1969: 199. Iraq: Karbala; HT; LT; SMNS. 

similis. Iran, Syria, Russia (Europe). Anabasis aphylla; Anabasis spp. (Chenopodiaceae). 

Baldratia similis Möhn 1969: 186. Iran: Rayin, SE Kerman; HT; LT; SMNS. 

suaedae. Canary Is, Algeria, Israel. Suaeda vera (Chenopodiaceae). 

Baldratia suaedae Möhn 1969: 171. Israel: Jaffa; HT; LT; SMNS. 

suaedifolia. Kazakhstan. Suaeda acuminata (Chenopodiaceae). 

Baldratia suaedifolia Fedotova 1991e: 51. Kazakhstan; Karatau Mtn Range, Berkutinka R. valley, nr Zhanatas; HT; 

♀; ZMAS. 

terteriani. Armenia. Suaeda altissima (Chenopodiaceae). 

Arafavilla terteriani Mamaev & Mirumian 1990b: 139. Armenia: Masisskii Region; HT; ♂; HYNM. 

tragani. Egypt. Traganum nudatum (Chenopodiaceae). 

Baldratia tragani Möhn 1969: 200. Egypt: Nouébe, El Hamd; HT; LT; SMNS. 

tubulata. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Baldratia tubulata Mamaev 1972a: 888. Turkmenistan: Mary, Akibay; HT; ♀; ZMAS. 

BAYERIOLA 

Bayeria Rübsaamen 1914: 97, preocc. Fritsch in Fritsch and Bayer 1905. Type species, erysimi Rübsaamen (mon.). 

Bayeriola Gagné 1991: 791, new name for Bayeria Rübsaamen.

Western Palearctic; 4 spp.; Lasiopteridi, Dasineurini. 

buhri. Germany. Gypsophila fastigata (Caryophyllaceae). 

Bayeria buhri Möhn 1958: 6. Germany: Kyffhäuser, Ochsenburg; HT; ♂; SMNS. 

erysimi. Germany, Poland, Czechoslovakia, Romania. Erysimum virgatum (Brassicaceae). 

Bayeria erysimi Rübsaamen 1914: 97. Czech Republic: Brno; ST; ♂,♀,L; ZMHB. 

salicariae. Widespread Europe. Lythrum salicaria (Lythraceae). 

Cecidomyia salicariae Kieffer 1888e: 96. France: vic. Guinkirchen; ST; ♀,L; lost?. 

thymicola. Widespread Europe, Morocco, Tunisia. Thymus serpyllum, T. chamaedrys (Lamiaceae). 

Cecidomyia thymicola Kieffer 1888e: 102. Germany: Ohrdruf; ST; ♂,♀,,L; lost?. 

BERBERISOMYIA 

Berberisomyia Kovalev 1972: 413. Type species, Diarthronomyia sobolevi Kovalev (orig. des.). 

Eastern Palearctic; 1 sp.; Lasiopteridi, unplaced to tribe. 

sobolevi. Russia (Far East). Berberis amurensis (Berberidaceae). 

Diarthronomyia sobolevi Kovalev 1967: 103. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; KOVC. 

*BICORNIDIPLOSIS 

Bicornidiplosis Fedotova 2003e: 43. Type species, accrescentis Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. 

*accrescentis. Russia (Far East). 

Bicornidiplosis accrescentis Fedotova 2003e: 44. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

BLAESODIPLOSIS 

Blaesodiplosis Gagné 1973c: 864. Type species, Hormomyia crataegifolia Felt (orig. des.). 

Nearctic; 3 spp., all on Rosaceae; Cecidomyiidi, unplaced to tribe. 

canadensis. USA (New York, Maine, Massachusetts). Amelanchier canadensis (Rosaceae). 

Hormomyia canadensis Felt 1908b: 388. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

crataegifolia. Canada (Ontario), USA (Michigan, New York, Massachusetts, Virginia). Crataegus sp. (Rosaceae). 

Hormomyia crataegifolia Felt 1907c: 160. USA: New York, Nassau; ST; ♂,♀; NYSM. 

venae. USA (Oregon, New York, Massachusetts, Virginia). Crataegus sp.; C. punctata (Rosaceae). 

Cecidomyia venae Stebbins 1910: 39 10 vic. Springfield -- USA: Massachusetts, vic. Springfield; T; G; NHSM. 

Lobopteromyia venae Felt 1914f: 120. USA: New York, Nassau; ST; ♀,L,egg; NYSM. 

BLASTODIPLOSIS 

Blastodiplosis Kieffer 1912c: 8. Type species, Clinodiplosis artemisiae Kieffer (orig. des.). 

Palearctic, Oriental; 2 spp.; Cecidomyiidi, unplaced to tribe. 

artemisiae. UK, Denmark, France, Germany, Czechoslovakia, Romania, Greece. Artemisia vulgaris (Asteraceae). 

140

Clinodiplosis artemisiae Kieffer 1901c: 34. France: Moselle; ST; ♂,L; lost?. 

thalictrina. Spain. Thalictrum flavum (Ranunculaceae). 

Blastodiplosis thalictrina Tavares 1916b: 100. Spain: Gallicia, Tuda; ST; ♂,♀; TAVC. 

BLASTOMYIA 

Blastomyia Kieffer 1913i: 54. Type species, Oligotrophus origani Tavares (mon.). 


origani. Widespread Europe, Morocco, Algeria. Origanum virens (Lamiaceae). 

Oligotrophus origani Tavares 1901: 66. Portugal: Setubal; ST; ♂,♀,L; TAVC. 

BRACHINEURA 

Brachineura Rondani 1840: 16. Type species, fuscogrisea Rondani (mon.). 

Brachyneura Anonymous 1844: 450, missp. of Brachineura. 

Brachynevra Agassiz 1847: 51, unjust. emend. of Brachineura. 

Spaniocera Winnertz 1853: 190. Type species, squamigera Winnertz (mon.). 

Acroectasis (as Acroëctasis) Rübsaamen 1910a: 131. Type species, maura Rübsaamen (mon.). 

Brachyneurella Kieffer 1913i: 112. Type species, Brachyneura squamata Kieffer (orig. des.). 

Ramineura Fedotova & Sidorenko 2006d: 84, as subgenus of Brachineura. Type species, Brachinura ussurica 

Fedotova (orig. des.). 

Cornistyla Fedotova & Sidorenko 2006d: 87, as subgenus of Brachineura. Type species, Brachineura nijveldti 

Fedotova (orig. des.). 

Cosmopolitan; 30 spp.; Brachineuridi. Larvae are associated with fungi and decaying vegetable matter. Fedotova and 

Sidorenko erected two new subgenera for Brachineura that are not separately used here. Ref.: Gagné & Solinas 1996 

(descr.). 

aberrata. Russia (Europe). 

Brachyneura aberrata Mamaev 1967: 876. Russia: Moscow Prov., Danki; HT; ♂; HYNM. 

alia. Russia (Far East). 

Brachineura alia Mamaev 1994: 31. Russia: Kamchatka, Kozyrevsk; HT; ♂; ZMMU. 


Brachyneura americana Felt 1907d: 5. USA: New York, Albany; HT; ♀; NYSM. 

antennata. Ukraine. 

Brachyneura antennata Mamaev 1967: 875. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

apicata. Russia (Europe). 

Brachineura apicata Mamaev 1967: 876. Russia: Moscow Prov., Pavlovskaja Sloboda; HT; ♂; ZMMU. 

australis. Australia (New South Wales). 

Spaniocera australis Skuse 1890: 385. Australia: New South Wales, Hogan's Brush, Navara Creek, nr Gosford; HT; 

♂; ANIC. 

*enigmatosa. Russia (Far East). 

Brachineura enigmatosa Fedotova & Sidorenko 2006d: 78. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya 


eupatorii. USA (New York). 

Brachineura eupatorii Felt 1908b: 317. USA: New York, Poughkeepsie; HT; ♀; NYSM. 

*eximia. Russia (Far East). 

Brachineura eximia Fedotova & Sidorenko 2006d: 80. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*flabellata. Russia (Far East). 

141

142 

Brachineura flabellata Fedotova & Sidorenko 2006d: 87. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


fungicola. Russia (Europe). 

Brachyneura fungicola Mamaev 1967: 874. Russia: Yaroslavl Prov., Volkovo; HT; ♂; HYNM. 

fuscogrisea. Italy. 

Brachineura fuscogrisea Rondani 1840: 17. Italy: vic. Parma; LT (Gagné & Solinas 1996: 73); ♂; MZUF. 

*impleta. Russia (Far East). 

Brachineura impleta Fedotova & Sidorenko 2006d: 82. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


*inconspicua. Russia (Far East). 

Brachineura inconspicua Fedotova 2004i: 598. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*infirma. Russia (Far East). 

Brachineura infirma Fedotova & Sidorenko 2006d: 88. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


maura. Germany. 

Acroectasis maura Rübsaamen 1910a: 131. Germany: Remagen; T; ♂; ZMHB. 

minima. France. 

Brachyneura minima Kieffer 1904c: 379. France: vic. Bitche; T; ♂; lost?. 

modesta. Russia (Europe). 

Brachyneura modesta Mamaev 1967: 875. Russia: Yaroslav Prov., Volkovo; HT; ♂; HYNM. 

*musiva. Russia (Far East). 

Brachineura musiva Fedotova & Sidorenko 2006d: 85. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*nijveldti. Russia (Far East). 

Brachineura nijveldti Fedotova 2004i: 601. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

quercina. UK, Latvia, Russia (Europe). 

Brachyneura quercina Edwards 1937: 151. UK: England, Herts, Welwyn, Sherrard's Wood; HT; ♂; BMNH. 

scotica. UK, Ukraine. 

Brachyneura scotica Edwards 1937: 151. UK: Scotland, Morayshire, Logie; HT; ♂; BMNH. 

squamata. France. 

Brachyneura squamata Kieffer 1904c: 377. France: vic. Bitche; ST; ♂,♀; lost?. 

squamigera. UK, Netherlands, France, Germany. 

Spaniocera squamigera Winnertz 1853: 306. Germany; ST; ♂,♀; MNHN?. 

stygia. Germany. 

Lasioptera stygia Meigen 1818: 91. Germany; T; A; lost. 

stylosa. Sweden. 

Brachineura stylosa Mamaev 1998d: 5. Sweden: Dalarna; HT; ♂; ZMMU. 

*subulata. Russia (Far East). 

Brachineura subulata Fedotova & Sidorenko 2006d: 85. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


1*transversa. Russia (Far East). 

Brachineura transversa Fedotova & Sidorenko 2006d: 84. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*ussurica. Russia (Far East). 

Brachineura ussurica Fedotova 2004i: 601. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

vitis. USA (New York). 

Brachineura vitis Felt 1908b: 317. USA: New York, West Nyack; HT; ♂; NYSM. 

BRACHYDIPLOSIS

Brachydiplosis Rübsaamen 1910d: 286. Type species, caricum Rübsaamen (orig. des. as n. g., n. sp.). 


caricum. Germany, Czechoslovakia. Carex sp. (Cyperaceae). 

Brachydiplosis caricum Rübsaamen 1910d: 286. Germany: vic. Berlin; ST; ♂,♀; ZMHB. 

BRACHYLASIOPTERA 

Brachylasioptera Möhn 1964b: 576. Type species, rotunda Möhn (orig. des.). 

Neotropical; 2 spp.; Lasiopteridi, Alycaulini. 

gilibertiae. El Salvador. Gilibertia arborea (Araliaceae). 

Brachylasioptera gilibertiae Möhn 1964b: 579. El Salvador: La Libertad, nr Los Chorros; HT; LT; SMNS. 

rotunda. El Salvador. Pseudocalymma macrocarpa (Bignoniaceae). 

Brachylasioptera rotunda Möhn 1964b: 576. El Salvador: La Libertad, nr San Diego, E La Libertad; HT; ♂; SMNS. 

BRACHYNEURINA 

Brachyneurina Mamaev 1967: 876. Type species, xylophila Mamaev (orig. des.). 

Palearctic; 5 spp.; Brachineuridi. 

angulata. Russia (Europe). 

Brachyneurina angulata Mamaev 1967: 876. Russia: Moscow Prov., Pavlovskaja Sloboda; HT; ♂; HYNM. 

peniophorae. UK. Peniophora cinerea (Aphylloporales). 

Brachyneurina peniophorae Harris 1979a: 25. UK: England, Warwickshire, Knavenhill Wood; HT; ♂; BMNH. 

pleiomorpha. Russia (Far East). 

Brachyneurina pleiomorpha Mamaev 1998d: 6. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

*Hpyxidiiformis. Late Eocene - Ukrainian amber. 

Brachineurina pyxidiiformis Fedotova 2005e: 49. Ukraine amber: Rovno; HT; ♂; UASK. 

xylophila. Poland, Russia (Europe). 

Brachyneurina xylophila Mamaev 1967: 876. Russia: Yaroslavl Prov., Volkovo; HT; ♂; HYNM. 

BRASSOMYIA 

Brassomyia (also as Brassomya & Brassomyiia) Mitroshina & Fedotova 1993: 67. Type species, beknazarovae 

Mitroshina & Fedotova (orig. des.). 

Palearctic; 2 spp., on Brassicaceae; Lasiopteridi, Dasineurini. 

beknazarovae. Turkmenistan. Leptalleum filifolium (Brassicaceae). 

Brassomyia beknazarovae Mitroshina & Fedotova 1993: 68. Turkmenistan: southern Istyurt, Kaplankyrskiï Res., 300 

km W Tashauz; HT; ♂; ZMAS. 

strigosellae. Kazakhstan. Strigosella turkestanica; Strigosella spp. (Brassicaceae). 

Brassomyia strigosellae Fedotova 1994a: 125. Kazakhstan: northeastern Kyzylkum Desert, Karatau Mtn Range, 30 

km W Bairkum; HT; ♂; ZMAS. 

BRAUERIELLA 

143

Braueriella Kieffer 1896b: 4. Type species, Diplosis phillyreae Löw (orig. des.). 

Braucriella De Stefani 1898b: 3, missp. of Braueriella. 

Western Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. 

phillyreae. Portugal, Italy, Yugoslavia, Greece, Algeria, Tunisia. Phillyrea media (Oleaceae). 

Diplosis phillyreae Löw 1877: 13. Italy: Miramare nr Trieste; ST; ♂,♀,L; NHMW. 

BREMIA 

Bremia Rondani 1860: 289, as subg. of Diplosis. Type species, Cecidomyia decorata Loew (orig. des.). 

Cosmopolitan; 19 spp.; Cecidomyiidi, Aphidoletini. The host is known for only one species, an egg predator of a 

species of Odonata. Ref: Harris 1981a (biology, larva). 

agilis. Cameroon. 

Homobremia agilis Felt 1920c: 495. Cameroon: Lonji, nr Ulou R.; ST; ♂,♀; NYSM. 


Toxomyia americana Felt 1914c: 130. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

bifurcata. France. 

Bremia bifurcata Kieffer 1904c: 396. France: vic. Bitche; T; ♂; lost?. 


Bremia borealis Felt 1914c: 130. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

caricis. USA (New York). 

Mycodiplosis caricis Felt 1907a: 32. USA: New York, Nassau; HT; ♂; NYSM. 

ciliata. UK, France. 

Bremia ciliata Kieffer 1904c: 392. France: vic. Bitche; T; ♂; lost?. 

cilipes. Germany. 

Cecidomyia cilipes Winnertz 1853: 280. Germany; ST; ♂,♀; MNHN?. 

decorata. UK, Germany, Poland, Russia (Europe). 

Cecidomyia decorata Loew 1850a: 34. Poland; ST; ♂,♀; ZMHB?. 

Cecidomyia decorata Winnertz 1853: 281. Germany; ST; ♂,♀; lost. 

filicis. USA (New York). 

Bremia filicis Felt 1907a: 29. USA: New York, Karner; HT; ♂; NYSM. 

legrandi. Gabon. Malgassophlebia aequatoris (Odonata: Libellulidae). 

Bremia legrandi Harris 1981a: 29. Gabon: Makokou, rd to Libreville; HT; ♂; MNHN. 

longicornis. France. 

Bremia longicornis Kieffer 1904c: 401. France: vic. Bitche; T; ♂; lost?. 

longipes. UK, France. 

Dicrodiplosis longipes Kieffer 1901c: 29. France: Bitche; ST; ♂,♀; lost?. 

macrofilum. Philippines. 

Bremia macrofilum Felt 1919a: 290. Philippines: Mindanao, Lanao Distr., Kolambugan; ST; ♂,♀; NYSM. 

mirifica. Colombia. 

Bremia mirifica Gagné 1994a: 176. Colombia: Valle; HT; ♂; USNM. 

montana. USA (New York). 

Bremia montana Felt 1914c: 131. USA: New York, Adirondack Mts, Mountain L.; HT; ♂; NYSM. 

obconica. India (West Bengal). 

Bremia obconica Grover 1979c: 89. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

podophyllae. USA (New York). 

Bremia podophyllae Felt 1907a: 29. USA: New York, Albany; HT; ♂; NYSM. 

Bremia podophylli Felt 1907c: 126, unjust. emend. of podophyllae. 

sylvestris. USA (New York). 

144

Bremia sylvestris Felt 1920a: 289. USA: New York, Keene Valley; HT; ♂; NYSM. 

tristis. USA (New York), UK. 

Bremia tristis Felt 1914c: 131. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

BREMIOLA 

Bremiola Rübsaamen 1916b: 491. Type species, Cecidomyia onobrychidis Bremi (orig. des.). 

Intranerviola Fedotova 1986a: 33, as subg. of Bremiola. Type species, Bremiola decipiens Fedotova (orig. des.). 

Radicaliola Fedotova 1993j: 78, as subg. of Bremiola. Type species, Bremiola rosulae Fedotova (orig. des.). 

Palearctic; 15 spp., 13 of them on Fabaceae, 2 on Chenopodiaceae; Lasiopteridi, Dasineurini. 

alpina. Kazakhstan. Oxytropis platysema; O. microsphaera (Fabaceae). 

Bremiola alpina Fedotova 1990h: 108. Kazakhstan: Dzhungarskiï Alatau, 10 km SW Kabal; HT; ♂; ZMAS. 

astragalicola. Kazakhstan. Astragalus alpinus; A. balchaschensis, A. paucijugus (Fabaceae). 

Bremiola astragalicola Fedotova 1986a: 37. Kazakhstan: Zailiïskiï Alatau, Turgen' canyon, 40 km S Turgen'; HT; ♂; 

ZMAS. 

calophacae. Kazakhstan. Calophaca tianschanica; C. turkestanica (Fabaceae). 

Bremiola calophacae Fedotova 1986a: 33. Kazakhstan: Karzhantau Mtn Range, upper course Dongustau R., 15 km 

SE Lenger; HT; ♂; ZMAS. 

caraganae. Kazakhstan. Caragana aurantiaca; Caragana spp. (Fabaceae). 

Bremiola caraganae Fedotova 1986a: 40. Kazakhstan: Narynkol'skaya Valley, nr Tekes, 20 km NW Narynkol; HT; 

♂; ZMAS. 

caraganicola. Kazakhstan. Caragana arborescens (Fabaceae). 

Bremiola caraganicola Fedotova 1990h: 114. Kazakhstan: Altaï, Ivanovskiï Mts, 32 km NE Leninogorsk, nr 

Poperechnoe; HT; ♂; ZMAS. 

decipiens. Kazakhstan. Halimodendron halodendron (Fabaceae). 

Bremiola decipiens Fedotova 1986a: 39. Kazakhstan: northeast Karatau Mtn Range, nr Bilikul', 5 km NW 

Karabastau; HT; ♂; ZMAS. 

deserta. Kazakhstan. Astragalus paucijugus; Astragalus spp. (Fabaceae). 

Bremiola deserta Fedotova 1990h: 110. Kazakhstan: mid Ili R., vic. Bakanas; HT; ♂; ZMAS. 

halimodendronis. Kazakhstan. Halimodendron halodendron (Chenopodiaceae). 

Bremiola halimodendronis Fedotova 1986a: 34. Kazakhstan: Northeast Karatau Mtn Range, nr. Bilikul', 5 km NW 

Karabastau; HT; ♂; ZMAS 

hedysarii. Kazakhstan. Hedysarum cephalotes; Hedysarum spp. (Fabaceae). 

Bremiola hedysarii Fedotova 1986a: 36. Kazakhstan: western Talasskiï Alatau, Ul'ken-Kaindy canyon, 30 km SE 

Bannovka; HT; ♂; ZMAS. 

karatavica. Kazakhstan. Astragalus skorniakovii (Fabaceae). 

Bremiola karatavica Fedotova 1994h: 32. Kazakhstan: Karatau Mtn Range, 6 km NE Ashchisaï; HT; ♂; ZMAS. 

onobrychidis. Widespread Europe, Kazakhstan. Onobrychis sativa; Onobrychis spp. (Fabaceae). 

Cecidomyia onobrychidis Bremi 1847a: 53. Switzerland: Zurich and Aargau Cantons; ST; ♂,♀,L; ETHZ?. 

oxytropicola. Kazakhstan. Oxytropis puberula; O. glabra, O. microsphaera (Fabaceae). 

Rhopalomyia oxytropicola Fedotova 1984e: 1031. Kazakhstan: southern shore L. Balkhash, Karatal R., vic. 

Golovnoe; HT; ♂; ZKAS. 

rosulae. Kazakhstan. Astragalus mucidus (Fabaceae). 

Bremiola rosulae Fedotova 1993j: 78. Kazakhstan: Moyynkum Desert, 17 km NE Sarykoby; HT; ♂; ZMAS. 

sphaerophysae. Kazakhstan. Sphaerophysa salsula (Fabaceae). 

Bremiola sphaerophysae Fedotova 1990h: 118. Kazakhstan: lower Karatal R., nr Kopberlik; HT; ♂; ZMAS. 

tarbagataica. Kazakhstan. Calophaca soongorica (Fabaceae). 

Bremiola tarbagataica Fedotova 1990h: 112. Kazakhstan: Tarbagataï Mtn Range, 40 km NW Urdzhara, nr 

Blagodatnoye; HT; ♂; ZMAS 

145

BREPHOMETRA 

Pero Meinert 1870: 463, preocc. Herrich-Schaeffer 1856. Type species, fasciata Meinert (mon.). 

Brephometra Strand 1910: 155, new name for Pero Meinert. 

Meinertomyia Felt 1911c: 37, new name for Pero Meinert. 

Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. Ref.: Edwards 1941, placement. 

fasciata. Denmark. Carpinus betulus (Betulaceae). 

Pero fasciata Meinert 1870: 465. Sweden: Westrabygaard, Skane; ST; ♂,♀,L; ZMUC. Ref.: Skuhravá 2005: 82 

(rediscovery of type); Skuhravá et al. 2006: 68 (type. loc.). 

BRETHESIA 

Brethesia Maia 2009a: 38 Type species, retorta Maia (orig. des.). 

Neotropical; 1 sp., a leaf gall maker on Myrcia (Myrtaceae); Cecidomyiidi, Asphondyliini, Schizomyiina. 

*retorta. Brazil (Minas Gerais). Myrcia retorta (Myrtaceae). 

Brethesia retorta Maia 2009a: 39. Brazil: Minas Gerais: Três Marias; HT; ♂; MNRJ. 

BRUGGMANNIA 

Bruggmannia Tavares 1906b: 81. Type species, braziliensis Tavares (mon.). 

Feltomyia Kieffer 1913i: 100. Type species, Uleella pisonifolia Felt (orig. des.). 

Brugmannia Maia 2004c: 761, missp. of Bruggmannia. 

Neotropical; 19 spp. from complex leaf galls, mostly on Nyctaginaceae, but one each on Myrsinaceae and Rubiaceae; 

Cecidomyiidi, Asphondyliini, Schizomyiina. Ref.: Möhn 1962 (illus.). 

*acaudata. Brazil (Rio de Janeiro). Guapira opposita (Nyctaginaceae). 

Bruggmannia acaudata Maia 2004c: 761. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

braziliensis. Brazil (Rio Grande do Sul). Myrsine (Myrsinaceae). 

Bruggmannia braziliensis Tavares 1906b: 82. Brazil: Rio Grande do Sul, São Leopoldo; LT (Möhn 1962: 217); ♂; 

SMNS. 

depressa. Brazil (Amazonas, Rio de Janeiro, Santa Catarina). Neea sp. (Nyctaginaceae). 

Uleella depressa Kieffer 1913i: 101. Brazil: Amazonas, Belém & Juruá Miri; Rio de Janeiro, Floresta da Tijuca; & 

Santa Catarina, Minas & Teresopolis; T; LT; ZMHB?. 

elongata. Brazil (Rio de Janeiro). Guapira opposita (Nyctaginaceae). 

Bruggmannia elongata Maia & Couri 1993: 210. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; 

QBUM. 

globulifex. Brazil (Rio de Janeiro). Neea sp. (Nyctaginaceae). 

Uleella globulifex Kieffer 1913i: 101. Brazil: Rio de Janeiro, Serra dos Orgãos; T; LT; ZMHB?. 

lignicola. Brazil (Rio de Janeiro). Neea sp. (Nyctaginaceae). 

Uleella lignicola Kieffer 1913i: 101. Brazil: Rio de Janeiro, Serra do Macahé; T; LT; ZMHB?. 

longicauda. Brazil (Rio de Janeiro). Neea sp. (Nyctaginaceae). 

Uleella longicauda Kieffer 1913i: 101. Brazil: Rio de Janeiro, R. de Janeiro; T; LT; ZMHB?. 

longiseta. Brazil (Amazonas). Neea sp. (Nyctaginaceae). 

Uleella longiseta Kieffer 1913i: 101. Brazil: Amazonas, Marari, Juruá; T; LT; ZMHB?. 

micrura. Brazil (Santa Catarina). Neea sp. (Nyctaginaceae). 

Uleella micrura Kieffer 1913i: 101. Brazil: Santa Catarina; T; LT; ZMHB?. 

monteiroi. Brazil (Rio de Janeiro). Guapira opposita (Nyctaginaceae). 

146

147 

Bruggmannia monteiroi Maia & Couri 1993: 214. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♀; 

QBUM. 

neeana. Brazil (Rio de Janeiro). Neea sp. (Nyctaginaceae). 

Uleella neeana Kieffer 1913i: 101. Brazil: Rio de Janeiro; T; LT; ZMHB?. 

pisoniae. Cuba. Pisonia sp. (Nyctaginaceae). 

Cecidomyia pisoniae Cook 1909: 144. Cuba; T; G; lost. 

pisonifolia. St. Vincent. Pisonia nigricans (Nyctaginaceae). 

Uleella pisonifolia Felt 1912i: 353. St. Vincent; ST; ♂,♀,P,L; NYSM. 

pisonioides. St. Vincent. Pisonia nigricans (Nyctaginaceae). 

Bruggmanniella pisoniae Felt 1912b: 174, jun. sec. hom. pisoniae Cook. St. Vincent; ST; ♂,♀,P,L; NYSM. 

Bruggmannia pisonioides Gagné 1994a: 137, new name for Bruggmanniella pisoniae Felt. 

psychotriae. El Salvador. Psychotria carthaginensis (Nyctaginaceae). 

Bruggmannia psychotriae Möhn 1960d: 352. El Salvador: La Libertad, San Diego, E La Libertad; HT; ♂; SMNS. 

pustulans. El Salvador. Pisonia macranthocarpa (Nyctaginaceae). 

Bruggmannia pustulans Möhn 1960d: 348. El Salvador: La Libertad, SE Ciudad Arce; HT; ♂; SMNS. 

randiae. El Salvador. Randia spinosa (Rubiaceae). 

Bruggmannia randiae Möhn 1960d: 354. El Salvador: San Salvador; HT; ♂; SMNS. 

robusta. Brazil (Rio de Janeiro). Guapira opposita (Nyctaginaceae). 

Bruggmannia robusta Maia & Couri 1993: 212. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

ruebsaameni. Brazil (Santa Catarina). Neea sp. (Nyctaginaceae). 

Uleella ruebsaameni (as rübsaameni) Kieffer 1913i: 101. Brazil: Santa Catarina, Pedras Grandes; T; LT; ZMHB?. 

BRUGGMANNIELLA 

Bruggmanniella Tavares 1909: 19. Type species, braziliensis Tavares (mon.). 

Hemibruggmanniella Möhn 1961a: 6. Type species, Bruggmanniella oblita Tavares (orig. des.). 

Brugmanniella Felt 1911l: 465, missp. of Bruggmanniella. 

Burggmanniella Del Guercio 1918a: 126, missp. of Bruggmanniella. 

Southern Nearctic, Neotropical, Palearctic, Oriental; 10 spp.; Cecidomyiidi, Asphondyliini, Asphondyliina. Yukawa 

(1971) pointed out the similarity of this genus to the Asian and Oceanian Pseudasphondylia. Refs: Möhn 1962 (illus.); 

Gagné et al. 2004 (review, key); Tokuda & Yukawa 2006 (rev.). 

*actinodaphnes. Japan (Honshu to Kyushu). Actinodaphne lancifolia (Lauraceae). 

Bruggmanniella actinodaphnes Tokuda & Yukawa 2006: 630. Japan: Kyushu, Kagoshima, Shiroyama; HT; ♂; 

KUEC. 

braziliensis. Brazil (Rio Grande do Sul). Sorocea illicifolia (Moraceae). 

Bruggmanniella braziliensis Tavares 1909: 20. Brazil: Rio Grande do Sul, vic. São Leopoldo; LT (Möhn 1963b: 12); 

♀; SMNS. 

bumeliae. USA (New Mexico, Texas, Louisiana, South Carolina, Florida). Bumelia lanuginosa (Sapotaceae). 

Asphondylia bumeliae Felt 1907d: 15. USA: Texas, Nuecestown; ST; ♂,♀; USNM. 

byrsonimae. Brazil (Rio de Janeiro). Byrsonima seriacea (Malpighiaceae). 

Asphondylia byrsonimae Maia & Couri 1992: 657. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; 

QBUM. 

*cinnamomi. Japan (Nansei Is.). Cinnamomum japonicum (Lauraceae). 

Bruggmanniella cinnamomi Tokuda & Yukawa 2006: 633. Japan: Okinawa, Nakagusuku; HT; ♂; KUEC. 

*duguetiae. Brazil (São Paulo). Duguetia furfuraceae (Annonaceae). 

Bruggmanniella duguetiae Urso-Guimarães & Amorim 2005: 430. Brazil: São Paulo, São Carlos; HT; ♀; MZSP. 

*ingae. Brazil (São Paulo). Inga edulis (Fabaceae). 

Bruggmanniella ingae Urso-Guimarães & Amorim 2005: 431. Brazil: São Paulo, betw. São José do Rio Preto & 

Tapiratiba; HT; ♂; MZSP. 

maytenuse. Brazil (Rio de Janeiro). Maytenus obtusifolia (Celastraceae).

148 

Asphondylia maytenuse Maia & Couri 1992: 655. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♀; 

QBUM. Ref.: Maia 1999a (note). 

oblita. Brazil (Rio de Janeiro). Schinus sp. (Anacardiaceae). 

Bruggmanniella oblita Tavares 1920a: 34. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♀,P; TAVC. 

perseae. Costa Rica, Colombia. Persea americana (Lauraceae). 

Bruggmanniella perseae Gagné 2004d: 549. Colombia: Caldas, Villamaria; HT; Cenicafé Insect Collection, 

Chinchiná, Caldas, Colobia. 

BUHRIELLA 

Buhriella Stelter 1960: 111. Type species, rubicola Stelter (orig. des.). 


rubicola. Sweden, Germany, Czechoslovakia, Austria, Greece. Rubus idaeus (Rosaceae). 

Buhriella rubicola Stelter 1960: 112. Germany: Kr. Rostock, Gross Lüsewitz, Kattawall; HT; ♂; DEIC. 

BUHROMYIELLA 

Buhromyiella Holz 1970: 98. Type species, giganteosaetosa Holz (orig. des.). 

Western Palearctic; 1 sp.; Cecidomyiidi, Mycodiplosini. 

giganteosaetosa. Germany. Erysiphe sp., Sphaerotheca sp. (Erysiphales). 

Buhromyiella giganteosaetosa Holz 1970: 99. Germany: Thüringen, Mühlhäusen; HT; ♂; SMNS. 

BUNGOMYIA 

Bungomyia Nayar 1949: 85. Type species, brachyrhyncha Nayar (orig. des.). 

Oriental; 1 sp.; Cecidomyiidi, unplaced to tribe. 

brachyrhyncha. India (West Bengal). 

Bungomyia brachyrhyncha Nayar 1949: 86. India: West Bengal, Calcutta; HT; ♂; ZSCI. 

BURSERAMYIA 

Burseramyia Möhn 1960c: 232. Type species, burserae Möhn (orig. des.). 


burserae. El Salvador. Bursera simaruba (Burseraceae). 

Burseramyia burserae Möhn 1960c: 233. El Salvador: La Libertad, NE Sitio del Nino, W lava fields; HT; ♂; SMNS. 

CACOPLECUS 

Cacoplecus Kieffer 1913h: 112. Type species, brevitarsis Kieffer (orig. des.). 


brevitarsis. Kenya. 

Cacoplecus brevitarsis Kieffer 1913h: 113. Kenya; HT; ♀; MNHN.

heterotarsus. Kenya. 

Cacoplecus heterotarsus, as var. of brevitarsis, Kieffer 1913h: 113. Kenya; HT; ♀; MNHN. 

CALAMOMYIA 

Calamomyia Gagné 1969d: 1356. Type species, Lasioptera echinochloa Felt (orig. des.). 

Nearctic; 19 spp., the 7 reared spp. all from culms of Poaceae; Lasiopteridi, Alycaulini. 

agrostis. USA. Eragrostis minor (Poaceae). 

Neolasioptera agrostis Felt 1908b: 331 (as n. sp. also in Felt 1911m: 483). USA; ST; ♂,♀; USNM. 

Neolasioptera agrostidis Kieffer 1913i: 22, unjustified new name for Neolasioptera agrostis Felt. 

alterniflorae. USA (Maine, North Carolina). Spartina alterniflora (Poaceae). 

Calamomyia alterniflorae Gagné 1981a: 132. USA: North Carolina, Carteret Co.; HT; ♂; USNM. 

canadensis. USA (New York). 

Lasioptera canadensis Felt 1907a: 9. USA: New York, Albany; HT; ♂; NYSM. 

consobrina. USA (New York). 

Lasioptera consobrina Felt 1907a: 8. USA: New York, Albany; HT; ♂; NYSM. 

danthoniae. USA (New York). Danthonia sp. (Poaceae). 

Lasioptera danthoniae Felt 1909b: 287. USA: New York, Ithaca; ST; ♂; NYSM. 

echinochloa. USA (South Dakota). Echinochloa crusgalli (Poaceae). 

Lasioptera echinochloa Felt 1916e: 181. USA: South Dakota, Elk Point; ST; ♂,♀,P,L; USNM. 

flavipes. USA (New York). 

Lasioptera flavipes Felt 1908b: 325. USA: New York, Albany; HT; ♂; NYSM. 

hecate. USA (New York). 

Lasioptera hecate Felt 1908b: 326. USA: New York, Albany; HT; ♀; NYSM. 

inustorum. USA (South Dakota, Oklahoma). Panicum virgatum (Poaceae). 

Lasioptera inustorum Felt 1916e: 182. USA: South Dakota, Elk Point; ST; ♂,♀,P,L; USNM. 

juvenalis. USA (New York). 

Lasioptera juvenalis Felt 1908b: 327. USA: New York, Newport; HT; ♀; NYSM. 

liriodendri. USA (New York). 

Choristoneura liriodendri Felt 1907a: 13. USA: New York, Albany; HT; ♂; NYSM. 

nassauensis. USA (New York). 

Lasioptera nassauensis Felt 1908b: 324. USA: New York, Nassau; HT; ♀; NYSM. 

panici. USA (New York). Dichanthelium latifolium (Poaceae). 

Lasioptera panici Felt 1908b: 326. USA: New York, Albany; HT; ♀; NYSM. 

phragmites. USA (Michigan, Ohio). Phragmites australis (Poaceae). 

Asteromyia phragmites Felt 1936a: 9. USA: Michigan, Maple Bay, Douglas L., & Ohio, Oak Harbor; LT (Gagné 

1968b: 31, Ohio: Oak Harbor); ♀; USNM. 

sexmaculata. USA (New York). 

Neolasioptera sexmaculata Felt 1908b: 331. USA: New York, Nassau; ST; ♂; NYSM. 

squamosa. USA (Missouri). (Poaceae). 

Neolasioptera squamosa Felt 1908b: 333 (as n. sp. also in Felt 1911m: 483). USA: Missouri, Cadet; ST; ♂; USNM. 

Neolasioptera subsquamosa Kieffer 1913i: 23, unjustified new name for Neolasioptera squamosa Felt of 1911m: 

483. 

Neolasioptera squamosella Kieffer 1913i: 23, new name for nonexistent Neolasioptera squamosa Felt "1909: 483." 

tenuitas. USA (New York). 

Neolasioptera tenuitas Felt 1908b: 331. USA: New York, Albany; HT; ♀; NYSM. 

tiliaginea. USA (New York). 

Neolasioptera tiliaginea Felt 1908b: 332. USA: New York, Nassau; HT; ♂; NYSM. 

tripunctata. USA (New York). 

Neolasioptera tripunctata Felt 1908b: 331. USA: New York, Nassau; HT; ♂; NYSM. 

149

CALLITRIDIPLOSIS 

Callitridiplosis Kolesik 2000c: 248. Type species, jana Kolesik (orig. des.). 

Australasian; 1 sp.; Cecidomyiidi, unplaced to tribe. 

jana. Australia (South Australia). Callitris gracilis (Cupressaceae). 

Callitridiplosis jana Kolesik 2000c: 249. Australia: South Australia, Sandy Creek Conservation Pk; HT; ♂; SAMA. 

CALODIPLOSIS 

Calodiplosis Tavares 1908: 165. Type species, parinarii Tavares (mon.). 


parinarii. Mozambique. Parinarium mobola (Rosaceae). 

Calodiplosis parinarii Tavares 1908: 165. Mozambique; ST; ♂,♀,P; TAVC. 

*CALYPTRADIPLOSIS 

Calyptradiplosis Fedotova 2004f: 102. Type species, turriformis (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. This genus does not differ from Resseliella. 

*turriformis. Russia (Far East). 

Calyptradiplosis turriformis Fedotova 2004f: 102. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

CAMPTODIPLOSIS 

Camptodiplosis Kieffer 1913a: 232. Type species, Mycodiplosis boleti Kieffer (orig. des.). 

Western Palearctic; 2 spp., fungivores; Cecidomyiidi, Mycodiplosini. Ref.: Holz 1970 (rev.). 

auriculariae. England. Auricularia auricula (Tremellales: Auriculariaceae). 

Camptodiplosis auriculariae Buxton & Barnes 1953: 197. UK: England, Bucks, Gerrard's Cross; HT; ♀; BMNH. 

boleti. UK, France, Germany. Boletus confluens; Albatrellus sp., Grifola sp., Poria sp. (Agaricales: Boletaceae, 

Polyporaceae?). 

Camptodiplosis boleti Kieffer 1901c: 28. France: vic. Bitche; ST; ♀; lost?. 

Mycodiplosis poriae Rübsaamen 1912b: 99. Germany: along Rhine, Ahr, & Mosel Rivers; ST; ♂,♀,L; ZMHB. 

CAMPTONEUROMYIA 

Camptoneuromyia Felt 1908b: 334. Type species, Dasineura virginica Felt, (subs. des. Coquillett 1910: 518). 

Mostly Neotropical, also Nearctic, but see Verbasciola; 15 spp., mostly inquilines in galls of other Cecidomyinae; 

Lasiopteridi, Camptoneuromyiini. 

adhesa. USA (New York). Solidago canadensis (Asteraceae). 

Dasyneura adhesa Felt 1907d: 10. USA: New York, West Nyack; ST; ♂,♀; NYSM. 

boerhaaviae. El Salvador, Colombia. Boerhaavia erecta; B. caribaea (Nyctaginaceae). 

150

151 

Camptoneuromyia boerhaaviae Möhn 1975: 22. El Salvador: San Vicente, km 81 nr Rio San Felipe; HT; ♂; SMNS. 

burserae. El Salvador. Bursera simaruba (Burseraceae). 

Camptoneuromyia burserae Möhn 1975: 16. El Salvador: La Libertad, NE Sitio del Niño, W lava fields; HT; ♂; 

SMNS. 

calliandrae. El Salvador. Calliandra houstoniana (Fabaceae). 

Camptoneuromyia calliandrae Möhn 1975: 26. El Salvador: Chalatenango, SE La Palma; HT; ♀; SMNS. 

crotalariae. El Salvador. Crotalaria longirostrata (Fabaceae). 

Camptoneuromyia crotalariae Möhn 1975: 24. El Salvador: San Salvador; HT; ♀; SMNS. 

exigua. El Salvador. Boerhaavia erecta (Nyctaginaceae). 

Camptoneuromyia exigua Möhn 1975: 23. El Salvador: San Vicente, km 81 nr Rio San Felipe; HT; ♀; SMNS. 

flavescens. USA (New York, New Jersey). Solidago graminifolia (Asteraceae). 

Lasioptera flavescens Felt 1908b: 327. USA: New York, West Nyack; HT; ♀; NYSM. 

fulva. USA (New York). 

Camptoneuromyia fulva Felt 1908b: 334. USA: New York, Albany; HT; ♀; NYSM. 

hamamelidis. USA (New York). 

Dasineura hamamelidis Felt 1907a: 20. USA: New York, Albany; HT; ♂; NYSM. 

meridionalis. St. Vincent, Trinidad. Ipomoea sp. (Convolvulaceae). 

Camptoneuromyia meridionalis Felt 1910c: 269. St. Vincent; ST; ♀; NYSM. 

rubifolia. USA (Massachusetts, Connecticut). Rubus sp. (Rosaceae). 

Camptoneuromyia rubifolia Felt 1908b: 334. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

siliqua. El Salvador. Jussiaea suffruticosa (Onagraceae). 

Camptoneuromyia siliqua Möhn 1975: 20. El Salvador: Sonsonate, rd to Los Cóbanos, km 5; HT; ♀; SMNS. 

virginica. USA (New York). 

Dasineura virginica Felt 1907a: 19. USA: New York, Albany; HT; ♂; NYSM. 

waltheriae. El Salvador. Waltheria americana (Sterculiaceae). 

Camptoneuromyia waltheriae Möhn 1975: 14. El Salvador: Santa Ana, E Tablón del Coco; HT; ♀; SMNS. 

xylosmatis. El Salvador. Xylosma flexuosum (Flacourtiaceae). 

Camptoneuromyia xylosmatis Möhn 1975: 18. El Salvador: Santa Ana, SE Candelaria, Hacienda Cortez; HT; ♂; 

SMNS. 

CAREOPALPIS 

Careopalpis Marikovskij 1955b: 104, nomen nudum. 

Careopalpis Marikovskij 1961: 45. Type species, anabasidis Marikovskij (orig. des.). 

Palearctic; 15 spp., all on Chenopodiaceae; Lasiopteridi, Lasiopterini. 

*akko. Israel. Suaeda splendens (Chenopodiaceae). 

Careopalpis akko Dorchin & Freidberg 2008: 10. Israel: Akko; HT; ♀; TAUI. 

anabasidis. Russia (Europe), Kazakhstan. Anabasis aphylla (Chenopodiaceae). 

Careopalpis anabasidis Marikovskij 1961: 46. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, middle & lower 

reaches Ili R.; ST; ♂,♀; ZKAS. 

balkhashensis. Kazakhstan, China (Xinjiang-Uygur Aut. Reg.). Kalidium foliatum (Chenopodiaceae). 

Careopalpis balkhashensis Marikovskij 1965: 77. Kazakhstan: NW shore L. Balkhash; ST; ♂,♀; MARC. 

contermina. Kazakhstan. Salsola arbusculaeformis (Chenopodiaceae). 

Stefaniola contermina Fedotova 1984d: 296. Kazakhstan: north shore middle course Ili R., Malye Kalkany Mtn; HT; 

♀; ZKAS. 

halocnemicola. Kazakhstan. Halocnemum strobilaceum (Chenopodiaceae). 

Careopalpis halocnemicola Fedotova 1990c: 29. Kazakhstan: mid Ili R., 20 km NW Barandysu; HT; ♀; ZMAS. 

halocnemii. Kazakhstan. Halocnemum strobilaceum (Chenopodiaceae). 

Careopalpis halocnemii Fedotova 1985e: 1196. Kazakhstan: middle course of Ili R., north shore, 20 km NW 

Barandysu; HT; ♀; ZKAS.

152 

halostachysis. Kazakhstan. Halostachys belangeriana (Chenopodiaceae). 

Careopalpis halostachysis Fedotova 1985e: 1195. Kazakhstan: middle course of Ili R., north shore, 20 km NW 

Barandysu; HT; ♀; ZKAS. 

harenosa. Armenia, Azerbaijan, Iran, Turkmenistan, Uzbekistan, Russia (Western Siberia). Kalidium caspicum; 

Halocnemum strobilaceum (Chenopodiaceae). 

Stefaniola harenosa Möhn 1971: 241. Azerbaijan: Baku Prov., Saljany; HT; LT; SMNS. 

kalidiis. Kazakhstan. Kalidium capsicum (Chenopodiaceae). 

Careopalpis kalidiis Fedotova 1992b: 48. Kazakhstan: 247 km NE Alma-Ata, Kurchaksholak R. valley; HT; ♂; 

ZMAS. 

kenderlensis. Kazakhstan. Suaeda microphylla (Chenopodiaceae). 

Careopalpis kenderlensis Fedotova 1998b: 16. Kazakhstan: eastern Mangyshlak basin, 60 km E Akkukuk, Kenderli 

spring; HT; ♀; ZMAS. 

roborovskiy. Kazakhstan, Turkmenistan. Anabasis salsa (Chenopodiaceae). 

Careopalpis roborovskiy Marikovskij 1965: 78. Kazakhstan: SW shore L. Balkhash; ST; ♂,♀; MARC. 

Stefaniola roborovskyi Skuhravá 1986: 137, unjust. emend. of roborovskiy. 

salicorniaramaea. Kazakhstan. Salicornia europaea (Chenopodiaceae). 

Careopalpis salicicorniaramaea Fedotova 1990c: 28. Kazakhstan: Ustyurt plateau, 14 km E Opornoï Stn; HT; ♀; 

ZMAS. 

suaedae. Armenia, Kazakhstan. Suaeda physophora (Chenopodiaceae). 

Stefaniola suaedae Fedotova 1983c: 22. Kazakhstan: Dzhungarskiï Alatau, left bank Ili R.; HT; ♀; ZMAS. 

suaedicola. Armenia, Kazakhstan. Suaeda physophora (Chenopodiaceae). 

Careopalpis suaedicola Fedotova 1985e: 1197. Kazakhstan: middle course of Ili R., north shore, 20 km NW 

Barandysu; HT; ♀; ZKAS. 

suaediphila. Turkmenistan. Suaeda acuminata; S. physophora, S. crassifolia (Chenopodiaceae). 

Careopalpis suaediphila Fedotova 1992d: 34. Turkmenistan: Kugitang Mtn Range, nr Charshanga; HT; ♀; ZMAS. 

*CARINATIDIPLOSIS 

Carinatidiplosis Fedotova 2004h: 166. Type species, scapiformis Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced here to tribe, but may belong to Mycodiplosini. 

*scapiformis. Russia (Far East). 

Carinatidiplosis scapiformis Fedotova 2004h: 166. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

CARTODIPLOSIS 

Cartodiplosis Gagné 1973c: 865. Type species, Cecidomyia nyssaecola Beutenmüller (orig. des.). 

Nearctic; 1 sp.; Cecidomyiidi, Lestodiplosini. 

nyssaecola. USA (New York, Maryland). Phylloxera nyssae (Hemiptera: Phylloxeridae). 

Cecidomyia nyssaecola Beutenmüller 1907a: 387. USA: New York, vic. New York; ST; ♂,♀; AMNH. 

*CARYADIPLOSIS 

Caryadiplosis Gagné 2008c: 88. Type species, venicola Gagné (orig. des.). 

Nearctic; 2 spp., from simple leaf galls on Carya (Juglandaceae); Cecidomyiidi, Cecidomyiini. 

*biconvexa. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae).

Caryadiplosis biconvexa Gagné 2008c: 89. USA: Washington, DC, National Arboretum; HT; LT; USNM. 

*venicola. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryadiplosis venicola Gagné 2008c: 89. USA: Maryland, Ridgely; HT; LT; USNM. 

CARYOMYIA 

Caryomyia Felt 1909b: 292. Type species, Cecidomyia tubicola Osten Sacken (orig. des.). 

Nearctic; 56 spp., all from complex leaf galls of Carya (Juglandaceae); Cecidomyiidi, Cecidomyiini. Ref.: Gagné 

2008c (rev.). 

*aggregata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia aggregata Gagné 2008c: 21. USA: Maryland, Beltsville; HT; LT; USNM. 

*albipilosa. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia albipilosa Gagné 2008c: 22. USA: Maryland, Beltsville, Entomology Road; HT; LT; USNM. 

*ansericollum. Eastern Nearctic. Carya cordiformis (Juglandaceae). 

Caryomyia caryaecola Felt 1921a: 114, sec. hom. Diplosis caryae Osten Sacken. USA: New York, Hamburg; L, 

desig. Gagné 2008c: 24; LT; NYSM. 

Caryomyia ansericollum Gagné 2008c: 23, new name for Caryomyia caryaecola Felt. 

antennata. Eastern Nearctic. Carya sp.; Carya spp. (Juglandaceae). 

Caryomyia antennata Felt 1909b: 292. USA: New York, vic. Albany; L, desig. Gagné 2008c: 25; ♂; NYSM. 

*arcuata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia arcuata Gagné 2008c: 25. USA: Missouri, St Clair; HT; LT; USNM. 

*asteris. Midwestern USA. Carya ovata; Carya spp. (Juglandaceae). 

Caryomyia asteris Gagné 2008c: 26. USA: Missouri, St Clair; HT; LT; USNM. 

*biretta. Eastern Nearctic. Carya ovata; Carya spp. (Juglandaceae). 

Caryomyia biretta Gagné 2008c: 27. USA: Massachusetts, Petersham; HT; LT; USNM. 

*caminata. Eastern Nearctic. Carya ovata; Carya spp. (Juglandaceae). 

Caryomyia caminata Gagné 2008c: 27. USA: Indiana, Spencer, I 64 rest stop, 2 mi E U.S. 231; HT; LT; USNM. 

caryae. Eastern Nearctic, NE Mexico. Carya sp.; Carya spp. (Juglandaceae). 

Diplosis caryae Osten Sacken 1862: 191. USA: District of Columbia; L, desig. Gagné 2008: 29; LT; USNM. 

Hormomyia arcuaria Felt 1908b: 388. USA: Massachusetts, Worcester; HT; ♂; NYSM. 

caryaecola. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Cecidomyia caryaecola Osten Sacken 1862: 192. USA: District of Columbia; ST; G; USNM. 

*cilidolium. Eastern Nearctic. Carya cordiformis. (Juglandaceae). 

Caryomyia cilidolium Gagné 2008c: 32. USA: Maryland, Ellicott City; HT; LT; USNM. 

*conoidea. Eastern Nearctic. Carya cordiformis. (Juglandaceae). 

Caryomyia conoidea Gagné 2008c: 34. USA: Maryland, Montgomery Co., Wheaton Pk; HT; LT; USNM. 

*cucurbitata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia cucurbitata Gagné 2008c: 35. USA: Maryland, Lusby; HT; LT; USNM. 

cynipsea. Eastern Nearctic. Carya glabra; Carya spp. (Juglandaceae). 

Cecidomyia cynipsea Osten Sacken 1862: 193. USA: Maryland, Wheaton, Wheaton Pk; N, desig. Gagné 2008c: 36; 


*deflexipili. Eastern Nearctic. Carya ovata; Carya spp. (Juglandaceae). 

Caryomyia deflexipili Gagné 2008c: 37. USA: Connecticut, Amston; HT; LT; USNM. 

*echinata. Central and southcentral Nearctic. Carya texana; Carya spp. (Juglandaceae). 

Caryomyia echinata Gagné 2008c: 38. USA: Missouri, Rolla; HT; LT; USNM. 

*eumaris. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia eumaris Gagné 2008c: 38. USA: Maryland, Beltsville, BARC, Entomology Road; HT; LT; USNM. 

*flaticrustum. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia flaticrustum Gagné 2008c: 40. USA: Maryland, Beltsville, BARC; HT; LT; USNM. 

*glauciglobus. Eastern Nearctic. Carya ovata; Carya spp. (Juglandaceae). 

153

154 

Caryomyia glauciglobus Gagné 2008c: 42. USA: Connecticut, Amston; HT; LT; USNM. 

*glebosa. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia glebosa Gagné 2008c: 42. USA: Missouri, Wildwood; HT; LT; USNM. 

*guttata. USA: Florida. Carya floridana (Juglandaceae). 

Caryomyia guttata Gagné 2008c: 43. USA: Florida, St. Lucie; HT; LT; USNM. 

*hirtidolium. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia hirtidolium Gagné 2008c: 43. USA: Maryland, Lusby; HT; LT; USNM. 

*hirtiglobus. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia hirtiglobus Gagné 2008c: 43. USA: Maryland, Silver Spring; HT; LT; USNM. 

holotricha. Eastern Nearctic. Carya sp.; Carya spp. (Juglandaceae). 

Cecidomyia holotricha Osten Sacken 1862: 193. USA: District of Columbia; L, desig. Gagné 2008c: 46; G; USNM. 

Dirhiza caryae Felt 1907a: 47. USA: New York, Albany; HT; ♀; NYSM. 

inanis. Eastern Nearctic. Carya sp.; Carya spp. (Juglandaceae). 

Caryomyia inanis Felt 1909b: 292. USA: New York, Nassau; L, desig. Gagné 2008c: 48; ♀; NYSM. 

*inclinata. Eastern Nearctic, NE Mexico. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia inclinata Gagné 2008c: 49. USA: West Virginia, 6 mi NW Hedgesville; HT; LT; USNM. 

*inflata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia inflata Gagné 2008c: 50. USA: West Virginia, 6 mi NW Hedgesville; HT; LT; USNM. 

*lenta. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia lenta Gagné 2008c: 51. USA: West Virginia, 6 mi NW Hedgesville; HT; LT; USNM. 

*levicrustum. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia levicrustum Gagné 2008c: 52. USA: Maryland, Beltsville, BARC, Entomology Road; HT; LT; USNM. 

*leviglobus. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia leviglobus Gagné 2008c: 54. USA: Maryland, Ridgely; HT; LT; USNM. 

*marginata. Eastern Nearctic. Carya ovata; Carya spp. (Juglandaceae). 

Caryomyia marginata Gagné 2008c: 55. USA: Massachusetts, Leominster; HT; LT; USNM. 

*melicrustum. Eastern Nearctic. Carya glabra; Carya spp. (Juglandaceae). 

Cecidomyia melicrustum Gagné: 56. USA: Maryland, Wheaton Pk; HT; LT; USNM. 

*ovalis. USA: Maryland. Carya glabra; Carya spp. (Juglandaceae). 

Cecidomyia ovalis Gagné: 58. USA: Maryland, Kemp Mill; HT; LT; USNM. 

persicoides. Eastern Nearctic. Carya sp.; Carya spp. (Juglandaceae). 

Cecidomyia persicoides Osten Sacken 1862: 193. USA: District of Columbia; ST; G; USNM. 

*procumbens. Eastern Nearctic. Carya ovata; Carya spp. (Juglandaceae). 

Caryomyia procumbens Gagné 2008c: 60. USA: West Virginia, Morgan Co.; HT; LT; USNM. 

*purpurea. Eastern Nearctic. Carya glabra; Carya spp. (Juglandaceae). 

Caryomyia purpurea Gagné 2008c: 61. USA: Maryland, Beltsville; HT; LT; USNM. 

*recurvata. Eastern Nearctic. Carya glabra; Carya spp. (Juglandaceae). 

Caryomyia recurvata Gagné 2008c: 62. USA: Washington, DC, National Arboretum; HT; LT; USNM. 

sanguinolenta. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Cecidomyia sanguinolenta Osten Sacken 1862: 192. USA: Maryland, Silver Spring, Walter Reed Medical Center 

Annex; N, desig. Gagné 2008c: 64; LT; USNM. 

*shmoo. Eastern Nearctic. Carya glabra; Carya spp. (Juglandaceae). 

Cecidomyia shmoo Gagné: 65. USA: Maryland, Wheaton; HT; LT; USNM. 

*spherica. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia spherica Gagné 2008c: 67. USA: Maryland, Beltsville Agicultural Research Center; HT; LT; USNM. 

*spiniglobus. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia spiniglobus Gagné 2008c: 68. USA: Maryland, Caroline Co., Ridgely; HT; LT; USNM. 

*spinulosa. Eastern Nearctic. Carya cordiformis (Juglandaceae). 

Caryomyia spinulosa Gagné 2008c: 69. USA: Pennsylvania, Pavia; HT; LT; USNM. 

*stellata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia stellata Gagné 2008c: 70. USA: Tennessee, Jackson; HT; LT; USNM. 

*striolacrustum. Eastern Nearctic. Carya glabra; Carya spp. (Juglandaceae).

155 

Caryomyia striolacrustum Gagné 2008c: 70. USA: Washington, DC, National Arboretum; HT; LT; USNM. 

*striolata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia striolata Gagné 2008c: 72. USA: Maryland, Beltsville, BARC-East, Entomology Road; HT; LT; USNM. 

*subulata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia lenta Gagné 2008c: 74. USA: West Virginia, Morgan Co., 6 mi NW Hedgesville; HT; LT; USNM. 

*supina. Eastern Nearctic. Carya glabra; Carya spp. (Juglandaceae). 

Cecidomyia supina Gagné: 74. USA: Maryland, Wheaton; HT; LT; USNM. 

thompsoni. Eastern Nearctic. Carya sp.; Carya spp. (Juglandaceae). 

Hormomyia thompsoni Felt 1908b: 388. USA: Massachusetts, Worcester; L, desig. Gagné 2008c: 75; G; NYSM. 

Caryomyia consobrina Felt 1909b: 292. USA: New York, Nassau; L, desig. Gagné 2008c: 76; ♂; NYSM. 

Caryomyia similis Felt 1909b: 292. USA: New York, Nassau; L, desig. Gagné 2008: 76; LT; NYSM. 

*tuberculata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia tuberculata Gagné 2008c: 77. USA: Maryland, vic. Ridgely; HT; LT; USNM. 

*tuberidolium. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia tuberidolium Gagné 2008c: 79. USA: Maryland, Gaithersburg; HT; LT; USNM. 

tubicola. Eastern Nearctic. Carya sp.; Carya spp. (Juglandaceae). 

Cecidomyia tubicola Osten Sacken 1862: 192. USA: District of Columbia; ST; L,G; USNM. 

*tumida. USA: Illinois, Pennsylvania. Carya ovata; C. tomentosa (Juglandaceae). 

Caryomyia tumida Gagné 2008c: 82. USA: Pennsylvania, Schellsburg; HT; LT; USNM. 

*turbanella. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia turbanella Gagné 2008c: 83. USA: Maryland, Beltsville, BARC, Entomology Road; HT; LT; USNM. 

*turbinata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia turbinata Gagné 2008c: 84. USA: Maryland, Beltsville, BARC, Entomology Road; HT; LT; USNM. 

*urnula. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia urnula Gagné 2008c: 85. USA: Maryland, Beltsville; HT; LT; USNM. 

*viscidolium. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Caryomyia viscidolium Gagné 2008c: 86. USA: Maryland, Silver Spring, Brunett Ave.; HT; LT; USNM. 

CECIDOMYIA 

Itonida Meigen 1800: 19. Type species, Tipula pini De Geer (subs. des. Coquillett 1910: 556). Suppressed by ICZN 

1963: 339. 

Cecidomyia Meigen 1803: 261. Type species, Tipula pini De Geer (mon.). 

Cedomyia Vallot 1827: 95, missp. of Cecidomyia. 

Cecydomyja Bouché 1833: 123, missp. of Cecidomyia. 

Phytophaga Rondani 1840: 13 (also as Fitophaga, p. 14). Type species, Tipula pini De Geer (subs. des. Gagné 

2004b: 100). 

Cecidomia Passerini 1850: 475, missp. of Cecidomyia. 

Diplosis Loew 1850a: 20, 32, as subg. of Cecidomyia. Type species, Tipula pini De Geer (subs. des. Rondani 1860: 

289). 

Cecydomyia Schiner 1854: 175, missp. of Cecidomyia. 

Cecidomya Rondani 1864: 189, missp. of Cecidomyia. 

Cryptodiplosis Kieffer 1895f: cxciv. Type species, Tipula pini De Geer (orig. des.). 

Retinodiplosis Kieffer 1912h: 1. Type species, Diplosis resinicola Osten Sacken (orig. des.). 

Stelechodiplosis Möhn 1955b: 141. Type species, magna Möhn (orig. des.). 

Holarctic, Neotropical, Oriental; 19 spp., all associated with resin of conifers; Cecidomyiidi, Cecidomyiini. Refs: 

Gagné 1978f (rev.); Nijveldt 1987 (Palearctic spp.); Gagné 2004b (syn. notes). 

bisetosa. USA (Florida). Pinus elliottii (Pinaceae). 

Cecidomyia bisetosa Gagné 1978e: 194. USA: Florida, Nassau Co.; HT; ♂; USNM. 

brevispatula. USA (Maryland, Florida). Pinus elliottii; Pinus spp. (Pinaceae).

156 

Cecidomyia brevispatula Gagné 1978f: 4. USA: Florida, Glades Co., Lykes Fire Tower; HT; LT; USNM. 

candidipes. Nearctic (widespread northeastern). Pinus strobus (Pinaceae). 

Cecidomyia albitarsis Felt 1918b: 383, jun. prim. hom. albitarsis Meigen 1830. USA: New York, Round L.; LT 

(Gagné 1978f: 5); ♂; NYSM. 

Cecidomyia candidipes Foote 1965: 287, new name for Cecidomyia albitarsis Felt. 

fortunactus. USA (Maryland, Florida). Pinus virginiana; P. elliottii (Pinaceae). 

Cecidomyia fortunactus Gagné 1978f: 5. USA: Maryland, Montgomery Co., Wheaton Pk.; HT; LT; USNM. 

harrisi. UK, Netherlands. Pinus sylvestris (Pinaceae). 

Cecidomyia harrisi Nijveldt 1987: 218. Netherlands: Ede; HT; LT; ZMAN. 

japonica. Japan (Kyushu). Pinus thunbergii (Pinaceae). 

Cecidomyia japonica Nijveldt 1987: 218. Japan: Kyushu, Fukuoka Pref., Koga; HT; ♂; KUIC. 

lamellata. USA (Florida). Taxodium distichum (Taxodiaceae). 

Cecidomyia lamellata Gagné 2008d: 431. USA: Florida, Ft. Pierce; HT; ♂; USNM. 

magna. UK, Germany, Latvia, Ukraine, Russia (Europe). Picea sp. (Pinaceae). 

Stelechodiplosis magna Möhn 1955b: 141. Germany: Hebertshausen nr Munich; HT; ♂; SMNS. 

mesasiatica. Kirghizstan. Picea schrenkiana (Pinaceae). 

Stelechodiplosis mesasiatica Mamaev 1971: 504. Kirghizstan: Sary-Chelek Res.; HT; ♂; HYNM. 

phagwariae. Pakistan. Pinus roxburghii (Pinaceae). 

Cecidomyia phagwariae Nijveldt 1987: 220. Pakistan; HT; ♂; BMNH. 

pini. Widespread northern & central Europe. Pinus maritima; P. sylvestris, Picea sp. (Pinaceae). 

Tipula pini De Geer 1776: 417. Sweden; ST; ♂,♀,P,L,egg; lost. 

Cecidomyza laterella Zetterstedt 1838: 823. Sweden: Kengis & Haparanda; ST; ♀,P; MZLU?. 

Cecidomyia pinimaritimae (as pini maritimae) Dufour 1838: 294. Southwestern France; ST; ♀; ?. 

Cecidomyia pilosa Bremi 1847a: 61. Switzerland: vic. Zurich; T; A; ETHZ. 

piniinopis. Widespread Nearctic. Pinus virginiana; Pinus spp. (Pinaceae). 

Cecidomyia piniinopis (as pini inopis) Osten Sacken 1862: 196. USA: District of Columbia; ST; LT; MCZ. 

Cecidomyia inopis Felt 1912g: 368, error for piniinopis. 

Cecidomyia banksianae Vockeroth 1960: 73. Canada: Manitoba, Stead; HT; ♂; CNCI. 

pumila. Russia (Far East). 

Cecidomyia pumila Mamaev & Efremova 1994: 32. Russia: Kamchatka, Petropavlovsk-Kamshatski; HT; ♂; MAMC. 

reburrata. USA (Maryland, Louisiana, Florida), Cuba. Pinus taeda; P. caribaea (Pinaceae). 

Cecidomyia reburrata Gagné 1978f: 6. USA: Maryland, Charles Co., Cedarville St. For.; HT; ♂; USNM. 

resinicola. Widespread Nearctic. Pinus virginiana; Pinus spp. (Pinaceae). 

Diplosis resinicola Osten Sacken 1871: 345. USA: New York, Tarrytown; LT (Gagné 1978f: 7); ♂; MCZ. 

Retinodiplosis palustris Felt 1915f: 408. USA: AL, Talladega; LT (Gagné 1978f: 7); ♂; NYSM. 

Cecidomyia reeksi Vockeroth 1960: 70. Canada: Manitoba, Stead; HT; ♂; CNCI. 

resinicoloides. Widespread Nearctic. Pinus radiata; Pinus spp. (Pinaceae). 

Cecidomyia resinicoloides Williams 1909: 2. USA: California, Stanford Univ. Arboretum; LT (Gagné 1978f: 8); ♂; 

NYSM. 

Cecidomyia accola Vockeroth 1960: 76. Canada: Ontario, Espanola; HT; ♂; CNCI. 

sarae. UK, Netherlands. Pinus sylvestris (Pinaceae). 

Cecidomyia sarae Nijveldt 1987: 218. Netherlands: Ede; HT; LT; ZMAN. 

tortilis. USA (Maryland, Florida). Pinus virginiana; P. clausa (Pinaceae). 

Cecidomyia tortilis Gagné 1978f: 8. USA: Maryland, Montgomery Co., Wheaton Pk.; HT; ♂; USNM. 

yunnanensis. China (Yunnan). Pinus yunnanensis (Pinaceae). 

Cecidomyia yunnanensis Wu & Zou 1989: 315. China: Yunnan, Mi-le Co.; HT; ♂; ?. 

CELTICECIS 

Celticecis Gagné 1983a: 436. Type species, Mayetiola celtiphyllia Felt (orig. des.). 

Holarctic; 11 spp., all from Celtis; Lasiopteridi, unplaced to tribe. Ref.: Moser 1965 (biology).

capsularis. USA (Texas). Celtis sp. (Ulmaceae). 

Cecidomyiaceltis capsularis Patton 1897: 248. USA: Texas; ST; G; USNM. 

celtiphyllia. USA (Iowa, Texas). Celtis sp. (Ulmaceae). 

Mayetiola celtiphyllia Felt 1908b: 371. USA: Iowa, Keokuk; LT (Gagné 1983a: 437); ♂; USNM. 

japonica. Japan (Honshu, Kyushu). Celtis sinensis var. japonica; C. jessoensis (Ulmaceae). 

Celticecis japonica Yukawa & Tsuda 1987: 124. Japan: Kyushu, Korimoto, Kagoshima City; HT; ♂; KUIC. 

oviformis. USA (Texas). Celtis sp. (Ulmaceae). 

Cecidomyiaceltis oviformis Patton 1897: 248. USA: Texas; ST; G; USNM. 

painteri. USA (Texas). Celtis mississippiensis, C. reticulata (Ulmaceae). 

Phytophaga painteri Felt 1935a: 7. USA: Texas, Austin & San Antonio; ST; ♂,♀; NYSM. 

pubescens. USA (Texas). Celtis sp. (Ulmaceae). 

Cecidomyiaceltis pubescens Patton 1897: 248. USA: Texas; ST; G; USNM. 

semenrumicis. USA (Texas). Celtis sp. (Ulmaceae). 

Cecidomyiaceltis semenrumicis Patton 1897: 248. USA: Texas; ST; G; USNM. 

spiniformis. USA (Texas). Celtis sp. (Ulmaceae). 

Cecidomyiaceltis spiniformis Patton 1897: 248. USA: Texas; ST; G; USNM. 

texana. USA (Texas). Celtis reticulata (Ulmaceae). 

Phytophaga texana Felt 1935a: 8. USA: Texas, Austin; HT; ♂; NYSM. 

unguicula. USA (Ohio). Celtis occidentalis (Ulmaceae). 

Cecidomyia unguicula Beutenmüller 1907a: 388. USA: Ohio, Cincinnati; ST; ♂,♀; AMNH. 

wellsi. USA (Ohio). Celtis occidentalis (Ulmaceae). 

Phytophaga wellsi Wells 1916: 273. USA: Ohio, vic. Columbus; ST; G; OHUC. 

Phytophaga wellsi Felt 1916e: 190. USA: Ohio, vic. Columbus; ST; ♂,♀,L; NYSM. 

CENTRODIPLOSIS 

Centrodiplosis Kieffer & Jörgensen 1910: 405. Type species, crassipes Kieffer & Jörgensen (mon.). 

Neotropical; 1 sp.; Cecidomyiidi, Centrodiplosini. 

crassipes. Argentina (Mendoza). Lycium chilense (Solanaceae). 

Centrodiplosis crassipes Kieffer & Jörgensen 1910: 406. Argentina: Mendoza, Chacras de Coria; ST; ♂,♀,P,egg; 

lost. 

CHAETODIPLOSIS 

Chaetodiplosis Kieffer 1913h: 103. Type species, tropicus Kieffer (orig. des.). 


gymnastica. Nigeria. 

Chaetodiplosis gymnastica Collin 1922: 507. Nigeria: Moor Plantation, nr Ibadan; ST; ♂,♀; BMNH. 

tropica. Tanzania. 

Chaetodiplosis tropicus Kieffer 1913h: 103. Tanzania: Tavéta; HT; ♀; MNHN. 

CHAMAEDIPLOSIS 

Chamaediplosis Gagné 1990a: 146. Type species, nootkatensis Gagné (orig. des.). 

Nearctic; 2 spp.; Cecidomyiidi, unplaced to tribe. 

157

158 

nootkatensis. Canada (British Columbia). Cupressus nootkatensis (Cupressaceae). 

Chamaediplosis nootkatensis Gagné & Duncan 1990: 151. Canada: British Columbia, Saanichton; HT; ♂; CNCI. 

rugosa. USA (California); immigr.: Hawaiian Is. Cupressus arizonica (Cupressaceae). 

Contarinia rugosa Gagné 1986a: 127. USA: Hawaii, Maui, Olinda; HT; ♂; USNM. 

CHANCHUDIPLOSIS 

Chanchudiplosis Grover & Bakhshi 1978: 115. Type species, striata Grover & Bakhshi (orig. des.). 

Oriental; 1 sp.; Cecidomyiidi, Lestodiplosini. 

striata. India (Uttar Pradesh). 

Chanchudiplosis striata Grover & Bakhshi 1978: 116. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

CHAULIODONTOMYIA 

Chauliodontomyia Gagné 1969b: 108. Type species, parianae Gagné (orig. des.). 

Nearctic; 2 spp., known from adults associated with Pariana flowers; Cecidomyiidi, Clinodiplosini. 

egregia. Venezuela. Pariana stenolemma (Poaceae). 

Chauliodontomyia egregia Gagné 1969b: 111. Venezuela: nr Maracay, Rancho Grande; HT; ♂; USNM. 

parianae. Venezuela, Brazil. Pariana stenolemma (Poaceae). 

Chauliodontomyia parianae Gagné 1969b: 110. Venezuela: nr Maracay, Rancho Grande; HT; ♂; USNM. 

CHELOBREMIA 

Chelobremia Kieffer 1912h: 1. Type species, Clinodiplosis sublevis Kieffer (orig. des.). 

Western Palearctic; 1 sp.; Cecidomyiidae, Clinodiplosini. Ref.: Möhn 1955a (larva). 

sublevis. France. Sonchus oleraceus; S. arvensis (Asteraceae). 

Clinodiplosis sublevis Kieffer 1909b: 29. France: vic. chateau de La Grange, nr Thionville; ST; LT; lost?. 

Chelobremia insignis Kieffer 1912b: 138. France: Moselle; T; ♂; lost?. 

CHILOPHAGA 

Chilophaga Gagné 1969d: 1358. Type species, Lasioptera colorati Felt (orig. des.). 

Nearctic; 4 spp., from Poaceae; Lasiopteridi, Alycaulini. 

coloradensis. USA (Colorado). 

Neolasioptera coloradensis Felt 1918a: 194. USA: Colorado, Boulder; HT; ♀; NYSM. 

colorati. USA (South Dakota, Iowa, Connecticut). Muhlenbergia sp. (Poaceae). 

Lasioptera colorati Felt 1918a: 129. USA: South Dakota, Elk Point; ST; ♂,♀,L; USNM. 

gyrantis. USA (Florida). Aristida gyrans (Poaceae). 

Chilophaga gyrantis Gagné 1971d: 336. USA: Florida, Miami; HT; ♂; USNM. 

tripsaci. USA (Texas). Tripsacum dactyloides (Poaceae). 

Lasioptera tripsaci Felt 1910a: 10. USA: Texas, Plano; ST; ♀,L; USNM. 

CHRYBANEURA

Chrybaneura Gagné 1968a: 33. Type species, harrisoni Gagné (orig. des.). 

Neotropical; 1 sp.; Brachineuridi. 

harrisoni. Panama. Associated with spider egg shells. 

Chrybaneura harrisoni Gagné 1968a: 34. Panama: Changuinola; HT; ♂; USNM. Ref.: Gagné & Etienne 2009: 242 

(descr. ♀, pupa, larva). 

CHRYSODIPLOSIS 

Chrysodiplosis Kieffer 1911: 318. Type species, pulchricornis Kieffer (subs. des. Kieffer 1913h: 145). 

Afrotropical, Oriental; 2 spp.; Cecidomyiidi, unplaced to tribe. 

pulchricornis. Seychelles Is. 

Chrysodiplosis pulchricornis Kieffer 1911: 318. Seychelles Is: Mahé, Cascade Estate; HT; ♂; BMNH. 

squamatipes. Sri Lanka. 

Chrysodiplosis squamatipes Kieffer 1912a: 28. Sri Lanka: Peradeniya; HT; ♂; NZSI?. 

*CINGOLA 

Cingola Fedotova & Sidorenko 2006d: 96. Type species, certa Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Brachineuridi. This genus is close to Brachineura if the two must be separated. 

*certa. Russia (Far East). 

Cingola certa Fedotova & Sidorenko 2006d: 96. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; ♂; 

ZMAS. 

CLEITODIPLOSIS 

Cleitodiplosis Tavares 1921e: 108. Type species, Necrophlebia graminis Tavares (mon.). 


graminis. Brazil (Bahia, Rio de Janeiro). Paspalum conjugatum (Poaceae). 

Necrophlebia graminis Tavares 1916a: 37. Brazil: Bahia & Rio de Janeiro; ST; ♂,♀; TAVC. 

CLINODIPLOSIS 

Clinodiplosis Kieffer 1894d: 120. Type species, Diplosis cilicrus Kieffer (mon). 

Chinodiplosis Rübsaamen 1906c: 226, missp. of Clinodiplosis. 

Calamodiplosis Rübsaamen 1910d: 289. Type species, Clinodiplosis coriscii Kieffer (mon.) = cilicrus Kieffer. 

Ciinodiplosis Rübsaamen 1911a: 14, missp. of Clinodiplosis. 

Alethediplosis Tavares 1916b: 89. Type species, pulchricornis Tavares (orig. des.) = cilicrus Kieffer. 

Bainidiplosis Mani 1935: 427. Type species, championi Mani (orig. des.). 

Carpodiplosis Kjellander 1945: 38. Type species, papaveris Kjellander (orig. des.) = cilicrus Kieffer. 

Cosmopolitan; 102 spp.; Cecidomyiidi, Clinodiplosini. Most, particularly the European species, are mycophagous, 

but many New World species are phytophagous, including some Neotropical species that cause complex plant galls. A 

few are at least facultatively predaceous (personal observation). Ref.: Skuhravá 1973 (review Palearc.). 

159

160 

*achyrophoridis. Russia. Achyrophorus maculatus (Asteraceae). 

Clinodiplosis achyrophoridis Fedotova 2008: 141. Russia: Central Povolzhye, Zhigulevsk Nature Res., nr Bakhilova 


aestiva. USA (New York). 

Clinodiplosis aestiva Felt 1908b: 402. USA: New York, Nassau; HT; ♂; NYSM. 

agraria. USA (New York). 

Cecidomyia agraria Felt 1908b: 413. USA: New York, Nassau; ST; ♂; NYSM. 


Feltiella allahabadensis Grover 1965a: 23. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

americana. Panama. Ficus sp. (Moraceae). 

Hyperdiplosis americana Felt 1911k: 192. Panama: Canal Zone, Paraíso; ST; ♂,♀,L; USNM. 

*alternantherae. Paraguay, Brazil, Uruguay, Argentina. Alternanthera philoxeroides; A. aquatica (Amaranthaceae). 

Clinodiplosis alternantherae Gagné 2004c: 306. Argentina: Buenos Aires, 4 km from Dolores; HT; ♂; MLPA. 

apicis. USA (New York). 

Cecidomyia apicalis Felt 1908b: 413, jun. prim. hom. apicalis Walker 1856. USA: New York, Newport; ST; ♂; 

NYSM. 

Cecidomyia apicis Kieffer 1913i: 213, new name for Cecidomyia apicalis Felt. 

apocyni. USA (New York). Apocynum sp. (Apocynaceae). 

Cecidomyia apocyni Felt 1908b: 414. USA: New York, Nassau; HT; ♂; NYSM. 

araneosa. USA (Maryland, Virginia). 

Clinodiplosis araneosa Felt 1912h: 154. USA: District of Columbia; ST; ♂,♀; USNM. 

artemisiarum. India (West Bengal). Artemisia sp. (Asteraceae). 

Clinodiplosis artemisiarum Kieffer 1909a: 156. India: West Bengal; T; LT; lost?. 

bahiensis. Brazil (Bahia). Undet. sp. of Asteraceae. 

Eudiplosis bahiensis Tavares 1917b: 145. Brazil: Bahia, Salvador, Roça da Madre de Deus; ST; ♀,P,L; TAVC. 

baraldajensis. Kazakhstan. Rumex pamiricus (Polygonaceae). 

Clinodiplosis baraldajensis Fedotova 1997g: 881. Kazakhstan: western Karatau, Baraldai Mtn Range, upper course 

Baraldai R., nr Mt. Chulak-Kurgan; HT; ♂; ZMAS. 

botularia. UK, France, Denmark, Germany, Poland, Czechoslovakia. Fraxinus excelsior (Oleaceae). 

Cecidomyia botularia Winnertz 1853: 266. Germany; ST; ♂; lost. 

canadensis. USA (Massachusetts). Amelanchier sp., Prunus virginiana (Rosaceae). 

Itonida canadensis Felt 1911l: 453, 466; as n. sp. also in Felt 1911n: 558. USA: Massachusetts, Magnolia; T; G; 

NYSM. 

capsici. Guadeloupe, Costa Rica, French Guiana. Capsicum frutescens; C. annuum (Solanaceae). 

Clinodiplosis capsici Gagné 2000: 833. Guadeloupe: Ravine Chaude; HT; ♂; USNM. 

captiva. USA (New York). 

Mycodiplosis captiva Felt 1908b: 401. USA: New York, Nassau; HT; ♂; NYSM. 

carolina. USA (North Carolina). 

Mycodiplosis carolina Felt 1911l: 463, nomen nudum. 

Mycodiplosis carolina Felt 1911n: 549. USA: North Carolina, Kiwani; HT; ♂; USNM. 

cattleyae. Mexico, Ecuador, Brazil, Jamaica; immigr: Hawaiian Is, USA, western Europe with commerce (not 

established). Cattleya sp. (Orchidaceae). 

Cecidomyia cattleyae Molliard 1903: 165. France (intercepted on Neotropical plant material; ST; ♂,♀; ?. 

Clinodiplosis cattleyae Felt 1908b: 412. USA: MA (intercepted on Neotropical plant material; ST; ♂,♀; USNM. 

cearensis. Brazil (Ceará). Undet. sp. of Asteraceae. 

Eudiplosis cearensis Tavares 1917b: 153. Brazil: Ceará, Fortaleza; ST; ♂,♀; TAVC. 

cellularis. India (West Bengal). 

Clinodiplosis cellularis Kieffer 1909a: 156. India: West Bengal, Balason Valley, vic. Kurseong; T; LT; lost?. 

championi. India (Uttar Pradesh). Quercus dilatata (Fagaceae). 

Bainidiplosis championi Mani 1935: 428. India: Uttar Pradesh, Mundali, Chakrata; ST; ♂,♀; ZSCI. 

chlorophorae. Brazil (Rio de Janeiro). Chlorophora tinctoria (Moraceae). 

Clinodiplosis chlorophorae Rübsaamen 1905a: 82. Brazil: Rio de Janeiro, Fabrica; ST; LT; ZMHB.

161 

cilicrus. Widespread Europe; Russia (Western Siberia). Centaurea jacea, Scabiosa sp., Cirsium lanceolatum, Carlina 

vulgaris (Asteraceae); many other genera of various plant families (Skuhravá 1973). 

Diplosis cilicrus Kieffer 1889a: 152. France: Bitche; ST; ♂,♀,L; lost?. 

Diplosis pallescens Kieffer 1890f: 196. France: Metz; ST; ♂,♀; lost?. 

Diplosis rosiperda Rübsaamen 1892c: 54. Germany: Weidenau; ST; ♂,♀,L; ZMHB. 

Clinodiplosis coriscii Kieffer 1896c: 96. France: Moselle; T; LT; lost?. 

Clinodiplosis vitis Lüstner 1900: 81. Germany: Weinberg; ST; ♂,♀,P,L,egg; ?. 

Clinodiplosis impatientis Kieffer 1901d: 173. France: Bitche; T; ♂; lost?. 

Clinodiplosis sarothamni Kieffer 1902c: 117. France: vic. Bitche, & Alsace; ST; ♀,L; lost?. 

Clinodiplosis urticae Kieffer 1902c: 119. France: Bitche; ST; ♀,L; lost?. 

Clinodiplosis dahliae Kieffer 1904d: 92. Italy: Vallombrosa; ST; ♂,♀,P,L; lost?. 

Clinodiplosis acinorum Rübsaamen 1906c: 231. Germany; ST; ♂,♀,L; ZMHB. 

Clinodiplosis betonicae Kieffer 1909b: 3. France: vic. Bitche; ST; LT; lost?. 

Clinodiplosis biorrhizae Kieffer 1909b: 20. France: Thionville & Bitche; ST; ♂,♀,L; lost?. 

Clinodiplosis breviseta Kieffer 1909b: 21. France: château de La Grange, nr Thionville; ST; LT; lost?. 

Clinodiplosis cirsii Kieffer 1909b: 6. Italy; T; G; lost?. 

Clinodiplosis crassistigma Kieffer 1909b: 27. France: vic. Bitche; ST; LT; lost?. 

Clinodiplosis spinulosa Kieffer 1909b: 10. France: vic. Bitche; ST; LT; lost?. 

Clinodiplosis strobi Kieffer 1909b: 17. France: vic. Bitche; ST; ♂,♀,L; lost?. 

Clinodiplosis tetrahit Kieffer 1909b: 9. France: vic. Thionville; ST; LT; lost?. 

Clinodiplosis schlechtendali Rübsaamen 1911a: 16. Germany: Bad Godesberg; ST; ♂,♀,P,L; ZMHB. 

Clinodiplosis gallicola Rübsaamen 1911b: 55. Germany; ST; ♀,L; ZMHB. 

Clinodiplosis rhynchitou Rübsaamen 1911b: 52. Germany: several sites on Mosel R.; ST; ♂,♀,L; ZMHB. 

Clinodiplosis mali Kieffer 1913i: 238, nomen nudum. 

Alethediplosis pulchricornis Tavares 1916b: 89. Spain: Gallicia, Vianna del Bollo, Humoso, &Carballino; ST; ♂,♀; 

TAVC. 

Clinodiplosis cerricola Rübsaamen 1917: 72. Austria: Vienna; ST; ♂,♀,L; ZMHB. 

Clinodiplosis lathyri Rübsaamen 1917: 73. Germany: Metternich; ST; ♂,♀,L; ZMHB. 

Clinodiplosis scorzonerae Rübsaamen 1917: 74. Germany: Remagen; ST; ♂,♀,L; ZMHB. 

Clinodiplosis sorbicola Rübsaamen 1917: 74. Germany: nr Reinbeck; ST; ♂,L; ZMHB. 

Clinodiplosis piceae Kieffer 1920a: 18. Germany: vic. Munich; ST; ♂,♀; lost?. 

Clinodiplosis kiefferiana Tölg 1921: 34. Austria: vic. Vienna; ST; ♂,♀; ?. 

Clinodiplosis hedickeana Fahringer 1922: 80, unnec. new name for Clinodiplosis kiefferiana Tölg. 

Clinodiplosis pisicola Barnes 1928h: 184. UK: England, Aberystwyth & Harpenden; ST; ♂,♀; BMNH. 

Clinodiplosis hydrangeae Barnes 1944a: 129. UK: Cornwall, Bodmin; ST; ♂,♀; BMNH. 

Clinodiplosis papaveris Kjellander 1945: 38. Sweden: Åkarp, Landkrona, Svalöv; HT; ♂; MZLU. 

Clinodiplosis hyperici Barnes 1952: 702. UK: England, Surrey, Box Hill; ST; ♂,♀; BMNH. 

Clinodiplosis leguminicola Milne 1960: 96. UK: 11 counties in England & Wales; ST; ♂,♀; BMNH. 

Clinodiplosis fagopyri Mamaeva 1964a: 209. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ?. 

cingulata. Germany. 

Cecidomyia cingulata Winnertz 1853: 263. Germany; ST; ♀; lost. 

coffeae. Dominica, St. Vincent. Coffea liberica (Rubiaceae). 

Hyperdiplosis coffeae Felt 1911j: 305. St. Vincent; ST; ♂,♀; NYSM. 

*conica. Brazil (Rio de Janeiro). Sebastiana glandulosa (Euphorbiaceae). 

Clinodiplosis conica Oliveira and Maia 2008: 355. Brazil: Rio de Janeiro, Maricá; HT; ♂; QBUM. 

contariniperda. Italy. Pyrus communis (Rosaceae). 

Moreschiella contariniperda Del Guercio 1918a: 254. Italy; ST; ♂,♀; lost. The original description indicates that this 

species is a Clinodiplosis. 

contracta. USA (New York). 

Mycodiplosis contracta Felt 1908b: 401. USA: New York, Albany; HT; ♂; NYSM. 

corticis. USA (District of Columbia). 

Parallelodiplosis corticis Felt 1915f: 407. USA: District of Columbia; HT; ♂; USNM.

162 

corylicola. Japan (Honshu), Republic of Korea. Corylus heterophylla (Betulaceae). 

Clinodiplosis corylicola Shinji 1944: 386. Japan: Honshu; T; A; lost. 

Schizoneura corylicole Shinji 1938l: 272, missp. of corylicola. 

corylifolia. USA (New York). Corylus sp. (Betulaceae). 

Mycodiplosis corylifolia Felt 1907d: 20. USA: New York, West Nyack; ST; ♂,♀; NYSM. 

*costai. Brazil (Rio de Janeiro). Paullinia weinmanniaefolia (Sapindaceae). 

Clinodiplosis costai Maia 2005b: 676. Brazil: Rio de Janeiro, APA de Maricá; HT; ♂; MNRJ. 

cyanococci. USA (New York). 

Mycodiplosis cyanococci Felt 1907a: 32. USA: New York, Karner; HT; ♂; NYSM. 

denotata. Costa Rica. Theobroma cacao (Sterculiaceae). 

Clinodiplosis denotata Gagné 1984c: 127. Costa Rica: Limón Prov., nr Siquirres, Finca La Lola; HT; ♂; USNM. 

diodiae. Brazil (Rio de Janeiro). Diodia gymnocephala (Rubiaceae). 

Clinodiplosis diodiae Maia 2001a: 6. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

eupatorii. St. Vincent, Trinidad, Costa Rica, Brazil (Pará). Chromolaena odorata; Eupatorium spp. (Asteraceae). 

Hyperdiplosis eupatorii Felt 1911d: 110. St. Vincent; ST; ♂,♀,P; NYSM. 

examinis. USA (New York). 

Clinodiplosis examinis Felt 1913e: 306. USA: New York, Nassau; ST; ♂,♀; NYSM. 

fibulata. USA (New York). 

Mycodiplosis fibulata Felt 1908b: 401. USA: New York, Newport; HT; ♂; NYSM. 

*floricola. Brazil (Rio de Janeiro). Heteropteris nitida (Malpigiaceae). 

Clinodiplosis floricola Novo-Guedes and Maia 2008: 360. Brazil: Rio de Janeiro, Maricá; HT; ♂; QBUM. 

fulva. USA (Massachusetts). Quercus sp. (Fagaceae). 

Dicrodiplosis fulva Felt 1913f: 140 (as n. sp. also in Felt 1918e: 153). USA: Massachusetts, Boston, Arnold 

Arboretum; T; G; NYSM. 

gillettei. USA (Colorado). Pinus scopulorum (Pinaceae). 

Dicrodiplosis gillettei Felt 1911l: 465. USA: Colorado, Fort Collins; T; G; USNM. 

graminis. USA (New York). 

Cecidomyia cerealis Fitch 1845b: 263, jun. prim. hom. cerealis Sauter 1817. USA: New York, Salem; ST; ♂,♀; 


Cecidomyia graminis Fitch 1861: 832, new name for Cecidomyia cerealis Fitch.. 

holotricha. USA (New York). 

Mycodiplosis holotricha Felt 1908b: 401. USA: New York, Albany; HT; ♂; NYSM. 

iheringi. Brazil (Santa Catarina). Aegiphila arborescens (Verbenaceae). 

Autodiplosis iheringi Tavares 1925a: 45. Brazil: Santa Catarina, Joinville; ST; ♀,P,L; TAVC. 

ilicicola. Italy. Quercus ilex (Fagaceae). 

Moreschiella ilicicola Del Guercio 1918a: 180. Italy; T; LT; lost. The original description indicates that this species is 

a Clinodiplosis. 


Charidiplosis indica Rao 1953: 312. India: Uttar Pradesh, Agra; HT; ♂; ZSCI. 

indorensis. India (Madhya Pradesh). 

Clinodiplosis indorensis Grover 1979c: 32. India: Madhya Pradesh, Indore; HT; ♂; PGCA. 

infirma. USA (New York). 

Cecidomyia infirma Felt 1908b: 413. USA: New York, Albany; HT; ♂; NYSM. 

*infrequens. Russia (Far East). 

Clinodiplosis infrequens Fedotova 2003e: 59. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

insularum. Seychelles Is. 

Clinodiplosis insularum Kieffer 1911: 326. Seychelles Is: Mahé, Cascade Estate; ST; ♂,♀; BMNH. 

intermedia. USA (New York). 

Mycodiplosis intermedia Felt 1920a: 290. USA: New York, Keene Valley; HT; ♂; NYSM. 

invocata. Germany. Fraxinus excelsior (Oleaceae). 

Cecidomyia invocata Winnertz 1853: 269. Germany; ST; ♂,♀,L; MNHN?.

163 

kumayanagii. Japan (Honshu). Berchemia racemosa (Rhamnaceae). 

Orthodiplosis kumayanagii Shinji 1938m: 340. Japan: Honshu; T; ♀; lost. 

Clinodiplosis kumayanagi & kumayanigi Shinji 1944: 389, missp. of kumayanagii. 

lappa. USA (Massachusetts). Spiraea salicifolia (Rosaceae). 

Cecidomyia lappa Stebbins 1910: 35. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

Parallelodiplosis clarkei Felt 1911l: 472. USA: Massachusetts, Magnolia; T; G; NYSM. 

latibulorum. Germany. 

Cecidomyia latibulorum Winnertz 1853: 247. Germany; ST; ♂,♀; lost. 

lenis. USA (New York). 

Mycodiplosis lenis Felt 1920a: 290. USA: New York, Keene Valley; HT; ♂; NYSM. 

longicornis. USA (New York). 

Dicrodiplosis longicornis Felt 1918e: 156. USA: New York, Nassau; HT; ♀; NYSM. 

longistemmis. India (Uttar Pradesh). 

Clinodiplosis longistemmis Grover & Bakhshi 1978: 22. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

marcetiae. Brazil (Rio de Janeiro). Marcetia sp. (Melastomataceae). 

Eudiplosis marcetiae Tavares 1917a: 22. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♂,♀,P; TAVC. 

meibomiifoliae. USA (New Jersey). Desmodium canadense (Fabaceae). 

Cecidomyia meibomiifoliae Beutenmüller 1907b: 306. USA: New Jersey, Carlstadt; ST; ♂,♀; AMNH. 

melissae. Brazil (Rio de Janeiro). Melissa officinalis (Lamiaceae). 

Clinodiplosis melissae Maia 1994c: 695. Brazil: Rio de Janeiro, Maricá; HT; ♂; QBUM. 

microphysae. Kazakhstan. Microphysa elongata (Rubiaceae). 

Clinodiplosis microphysae Fedotova 1997c: 334. Kazakhstan: delta Ili R., 15 km NW Aral-Tyube; HT; ♂; ZMAS. 

modesta. USA (New York). 

Mycodiplosis modesta Felt 1908b: 402. USA: New York, Nassau; HT; ♂; NYSM. 

monticola. Tanzania. 

Clinodiplosis monticola Kieffer 1913h: 111. Tanzania: Kilema; HT; ♂; MNHN. 

mutabilis. Germany. 

Cecidomyia mutabilis Winnertz 1853: 248. Germany; ST; ♀; lost. 

nidorum. UK, Austria. 

Clinodiplosis nidorum Kieffer 1913a: 231. Austria; T; ♀; lost?. 

nodifex. India (West Bengal). 

Clinodiplosis nodifex Kieffer 1909a: 155. India: West Bengal, Kurseong; T; LT; lost?. 

nympha. Tanzania. 

Coprodiplosis nympha Kieffer 1913h: 96. Tanzania; Taveta; HT; ♀; MNHN. 

obscura. USA (New York). 

Mycodiplosis obscura Felt 1908b: 402. USA: New York, Albany; HT; ♂; NYSM. 

oleracei. Germany, Yugoslavia. Cirsium oleraceum (Asteraceae). 

Clinodiplosis oleracei Rübsaamen 1917: 75. Germany: Prignitz, Triglitz; ST; ♀,L; ZMHB. 

orientalis. India (Andhra Pradesh). 

Charidiplosis orientalis Rao 1956b: 19. India: Andhra Pradesh, Guntur; HT; ♂; ZSCI. 

paucifili. USA (New York). 

Cecidomyia paucifili Felt 1908b: 413. USA: New York, Albany; HT; ♂; NYSM. 

Itonida paucifila Felt 1921: 182, missp. of paucifili. 

*phalacrolomae. Russia (Far East). Phalacroloma strigosa (Asteraceae) 

Clinodiplosis palacrolomae Fedotova 2004e: 16. Russia: Primorskiï Terr., 23 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

phlox. USA (Ohio, Maine). Phlox sp. (Polemoniaceae). 

Hyperdiplosis phlox Greene 1941: 547. USA: Ohio, Chagrin Falls; ST; ♂,♀,L; USNM. 

picturata. Guyana. 

Lestodiplosis picturata Felt 1915h: 156. Guyana: Bartica; HT; ♀; NYSM. 

populi. USA (Massachusetts, New Jersey). Populus sp. (Salicaceae). 

Dicrodiplosis populi Felt 1908b: 394. USA: New Jersey, ♂; NYSM.

164 

pratensis. USA (New York). 

Clinodiplosis pratensis Felt 1908b: 412. USA: New York, Albany; HT; ♂; NYSM. 

profusa. Brazil (Rio de Janeiro). Eugenia uniflora (Myrtaceae). 

Clinodiplosis profusa Maia 2001a: 8. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

pulchra. Brazil (Bahia). Lantana sp. (Verbenaceae). 

Eudiplosis pulchra Tavares 1917b: pl. X, figs. 3-7; pl. XI, fig. 6. Brazil: Bahia, Salvador, Roça da Madre de Deus; 

ST; ♂,♀,L; TAVC. 

pyricola. Germany, Italy. Pyrus communis (Rosaceae). 

Cecidomyia pyricola Nördlinger 1855: 527. Germany; T; A; ?. Skuhravá 1986: 232 places this in questionable 

synonymy under cilicrus; if the two are synonyms, pyricola is senior. 

rhododendri. USA (New York, North Carolina). Rhododendron sp. (Ericaceae). 

Giardomyia rhododendri Felt 1939a: 42. USA: New York, Huntington Station; ST; ♂,♀; USNM. 

robusta. USA (New York). 

Mycodiplosis robusta Felt 1908b: 401. USA: New York, Albany; HT; ♂; NYSM. 

rosaefoliae. Korean Peninsula, Japan (Honshu to Kyushu). Rosa multiflora (Rosaceae). 

Schizeneura rosaefoliae Shinji 1939d: 66. Japan: Honshu, Morioka; ST; ♂,♀; lost. 

rubiae. Brazil (Rio de Janeiro). Rubia sp. (Rubiaceae). 

Eudiplosis rubiae Tavares 1918c: 70. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♂,♀; TAVC. 

rubida. USA (New York). 

Dicrodiplosis rubida Felt 1918e: 158. USA: New York, Albany; HT; ♀; NYSM. 

rubisolita. USA (New York, Connecticut). 

Clinodiplosis rubisolita Felt 1908b: 412. USA: New York, Albany; HT; ♂; NYSM. 

sanguinia. USA (New York). 

Cecidomyia sanguinia Felt 1908b: 413. USA: New York, Nassau; HT; ♂; NYSM. 

santhali. India (Bihar). 

Charidiplosis santhali Grover 1961b: 337. India: Bihar, Mahijam, Santhal Pargana; HT; ♂; PGCA. 

setariae. USA (New York, Florida). Setaria sp. (Poaceae). 

Cecidomyia setariae Felt 1907d: 22. USA: New York, Albany; HT; ♂; NYSM. 

*skuhravae. Russia (Far East). Vicea cracca (Fabaceae) 

Clinodiplosis skuhravae Fedotova 2004e: 17. Russia: Primorskiï Terr., 23 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

socialis. Germany. Rubus sp. (Rosaceae). 

Cecidomyia socialis Winnertz 1853: 251. Germany; ST; ♀; lost. 

spiraeaflorae. USA (New York). Spiraea salicifolia (Rosaceae). 

Cecidomyia spiraeaflorae Felt 1907d: 23. USA: New York, Albany; HT; ♂; NYSM. 

spiraeina. USA (Massachusetts). Spiraea salicifolia (Rosaceae). 

Itonida spiraeina Felt 1911l: 472. USA: Massachusetts, Magnolia; T; G; NYSM. 

styracifoliae. Japan (Honshu). Styrax obassia (Styracaceae). 

Clinodiplosis styracifoliae Shinji 1944: 390. Japan: Honshu; T; A; lost. 

tenuitas. USA (New York). 

Mycodiplosis tenuitas Felt 1908b: 401. USA: New York, Poughkeepsie; HT; ♂; NYSM. 

terrestris. USA (New York). 

Cecidomyia terrestris Felt 1908b: 413. USA: New York, Poughkeepsie; HT; ♂; NYSM. 

tsugae. USA (New York). 

Mycodiplosis tsugae Felt 1907a: 34. USA: New York, Lake Clear; HT; ♂; NYSM. 

uliginosa. USA (New York). 

Itonida uliginosa Felt 1914c: 133. USA: New York, Adirondack Mts, Woodworth's L.; ST; ♂,♀; NYSM. 

variabilis. USA (New York). 

Mycodiplosis variabilis Felt 1908b: 402. USA: New York, Nassau; HT; ♂; NYSM. 

venitalis. USA (New York). Crataegus sp. (Rosaceae). 

Dicrodiplosis venitalis Felt 1914f: 121. USA: New York, Nassau; HT; ♀; NYSM. 

verbenae. USA (New York, New Jersey). Verbena urticifolia (Verbenaceae).

165 

Cecidomyia verbenae Beutenmüller 1907b: 306. USA: New York, New York, Bronx Pk, & NJ, Fort Lee; ST; ♂,♀; 

AMNH. 

Cecidomyia urtifolia Felt 1908b: 414. USA: New York, Karner; HT; ♂; NYSM. 

*vitidis. Russia (Far East). Vitis amurensis (Vitaceae) 

Clinodiplosis vitidis Fedotova 2004e: 16. Russia: Primorskiï Terr., 23 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

CLUSIAMYIA 

Clusiamyia Maia 1997: 829. Type species, nitida Maia (orig. des.). 

Neotropical; 2 spp.; Cecidomyiidi, unplaced to tribe. 

granulosa. Brazil (Rio de Janeiro). Clusia hilariana (Clusiaceae). 

Clinodiplosis granulosa Maia 2001a: 10. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

nitida. Brazil (Rio de Janeiro). Clusia lanceolata (Clusiaceae). 

Clusiamyia nitida Maia 1997: 831. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

COCCIDOMYIA 

Coccidomyia Felt 1911c: 45. Type species, pennsylvanica Felt (orig. des.). 

Neotropical; 1 sp., associated with a coccid scale, its role not exactly known; Brachineuridi. Ref.: Gagné 1994a 

(review). 

pennsylvanica. USA (Pennsylvania). Lecanium sp. (Hemiptera: Coccidae). 

Coccidomyia pennsylvanica Felt 1911c: 45. USA: Pennsylvania, Highspire; ST; ♂,♀; USNM. 

COCCOMYZA 

Coccomyza Del Guercio 1918a: 264. Type species, brittini Del Guercio (orig. des. as n. g., n. sp.). 


brittini. New Zealand. Undet sp. of Eriococcidae (Hemiptera). 

Coccomyza brittini Del Guercio 1918a: 264. New Zealand: Waipara; ST; ♂,♀,P,L; lost. 

COELODIPLOSIS 

Coelodiplosis Kieffer 1913e: 43. Type species, magnipennis Kieffer (orig. des.). 


magnipennis. Taiwan. 

Coelodiplosis magnipennis Kieffer 1913e: 43. Taiwan: Taihorin; ST; ♂; lost?. 

COLLULA 

Collinia Kieffer 1913a: 232, preocc. Cépede 1910. Type species, acaciae Kieffer (orig. des.). 

Collula Strand 1928: 49, new name for Collinia Kieffer. 

Palearctic; 2 spp.; on Acacia; Cecidomyiidi, unplaced to tribe. Ref.: Gagné 1993c (review).

acaciae. Egypt. Acacia nilotica (Fabaceae). 

Perrisia acaciae Kieffer 1909b: 1. Egypt: Cairo; ST; G; NHMW?. 

kiefferi. Egypt. Acacia nilotica (Fabaceae). 

Collinia acaciae Kieffer 1913a: 233, jun. sec. hom. acaciae Kieffer 1909. 

Collula kiefferi Gagné 1993c: 112, new name for Collinia acaciae Kieffer 1912. 

*COMPOSITOLA 

Compositola Fedotova & Sidorenko 2006b: 18. Type species, competiva Fedotova & Sidorenko (orig. des.). 


*competiva. Russia (Far East). 

Compositola competiva Fedotova & Sidorenko 2006b: 18. Russia: Primorskiï Terr., Lazovskiï Res.; HT; ♂; ZMAS. 

COMPSODIPLOSIS 

Compsodiplosis Tavares 1909: 14. Type species, luteoalbida Tavares (mon.). 


luteoalbida. Brazil (Rio Grande do Sul). ?Smilax sp. (Smilacaceae). 

Compsodiplosis luteoalbida (as luteo-albida) Tavares 1909: 14. Brazil: Rio Grande do Sul, vic. São Leopoldo; ST; 

♂,♀,P; TAVC. 

CONIOPHORA 

Coniophora Nijveldt 1959: 81. Type species, graminicola Nijveldt (orig. des.). 

Palearctic; 5 spp.; Lasiopteridi, Dasineurini. 

acutissima. Russia (Far East). 

Coniophora acutissima Mamaev 1998d: 7. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♀; ZMMU. 

autumnalis. UK, Netherlands, Russia (Europe). Ulmus sp. (Ulmaceae). 

Procystiphora autumnalis Mamaev 1961d: 227. Russia: Voronezh Prov., Tellerman For.; ST; ♂,♀; MAMC. 

graminicola. UK, Netherlands, Russia (Europe). Phalaris arundinacea (Poaceae). 

Coniophora graminicola Nijveldt 1959: 81. Netherlands: nr Kapelle; HT; ♂; ZMAN. 

nijveldti. Bulgaria. 

Coniophora nijveldti Dimitrova 1992: 79. Bulgaria: Vitosha Mtn, by Selskata R.; HT; ♂; ZISB. 

semimarina. Russia (Europe). 

Coniophora semimarina Mamaev & Przhiboro 2001: 235. Russia: Pskov Prov., Sebezh Distr., shore of L. Anniskoe, 

nr Anniskoe; HT; ♂; ZMMU. 

CONODIPLOSIS 

Conodiplosis Kieffer 1913h: 102. Type species, dolichopsis Kieffer (orig. des.). 


dolichopsis. Kenya. 

Conodiplosis dolichopsis Kieffer 1913h: 102. Kenya: Likoni, nr Mombasa; HT; ♀; MNHN. 

166

CONTARINIA 

Contarinia Rondani 1860: 289, as subg. of Cecidomyia. Type species, Tipula loti De Geer (orig. des.). 

Eudiplosis Kieffer 1894a: xxviii. Type species, Contarinia sorbi Kieffer (subs. des., Gagné 2004b: 108). Kieffer 

1895a: cclxxx designated Tipula loti as type species, but it was neither of the two originally included species. 

Stictodiplosis Kieffer 1894a: xxviii. Type species, Contarinia picridis Kieffer (subs. des., Gagné 2004b: 108). Kieffer 

1895a: cclxxx designated Contarinia nubilipennis as type species, but it was neither of the two originally 

included species. 

Contariuia Rübsaamen 1906b: 194, missp. of Contarinia. 

Syndiplosis Rübsaamen 1910d: 284. Type species, winnertzi Rübsaamen (orig. des. as n. g., n. sp.) = petioli Kieffer. 

Atylodiplosis Rübsaamen 1910e: 342. Type species, Diplosis acetosellae Rübsaamen (subs. des. Rübsaamen & 

Hedicke 1926: 261). 

Doxodiplosis Kieffer 1912h: 1. Type species, picridis Kieffer (orig. des.). 

Dryodiplosis Kieffer 1912h: 1. Type species, Contarinia subulifex Kieffer (orig. des.). 

Navasodiplosis Tavares 1920c: 65. Type species, camphorosmae Tavares (orig. des.). 

Sissudiplosis Mani 1943: 44. Type species, chatterjeei Mani (orig. des.). 

Bothriochloamyia Rao & Sharma 1977b: 237. Type species, orientalis Rao & Sharma (orig. des.). 

Contarinomyia Fedotova 1991c: 51. Type species, Contarinia reaumuriae Fedotova (orig. des.). 

*Achillinia Fedotova 1992o: 115, as subgenus Contarinia. Type species, Contarinia achilleae Fedotova (orig. des.). 

Cosmopolitan; 311 spp.; Cecidomyiidi, Cecidomyiini. Most of these species live gregariously in flower heads or leaf 

roll galls. Some, such as C. maculipennis, are pests and evidently have a wide host range. Species placed here that were 

reared from Australian grasses may belong to Stenodiplosis; they need to be examined for the pertinent characters 

(Gagné 1994a). Ref.: Gagné 2004b (syn. note.). 

acerplicans. Widespread Europe. Acer pseudoplatanus; A. monspessulanum (Aceraceae). 

Diplosis acerplicans Kieffer 1889b: 171. France: vic. Bitche; ST; ♂,♀,L; lost?. 

acetosellae. UK, Netherlands, France, Germany, Czechoslovakia, Romania, Russia (Europe); immigr.: USA (Illinois, 

Georgia, Florida). Rumex acetosella (Polygonaceae). 

Diplosis acetosellae Rübsaamen 1891d: 128. Germany: betw. Siegen & Buschgotthardshütten; ST; ♂,♀,P,L; ZMHB. 

Contarinia acetosae Kieffer 1901c: 31. France: Bitche; ST; ♂,♀; lost?. 

Contarinia rumicis Kieffer (1896c: 99) of Skuhravá 1986: 235, but Kieffer 1896c referred his species to rumicis 

Loew. 

*achilleae. Kazakhstan. Achillea millefolii (Asteraceae). 

Contarinia achilleae Fedotova 1992o: 116. Kazakhstan: Tarbagataï Mtns, 30 km E Tarbagataï; HT; ♂; ZMAS. 

aconitifloris. Germany, Czechoslovakia, Switzerland, Austria, Yugoslavia. Aconitum lycoctonum (Ranunculaceae). 

Contarinia aconitifloris Stelter 1962a: 332. Germany: vic. Mühlhausen in Thuringen; HT; ♂; DEIC. 

acrocecis. Germany, Poland, Czechoslovakia, Yugoslavia. Galium mollugo (Rubiaceae). 

Contarinia acrocecis Stelter 1962b: 750. Germany: Kr. Rostock, Gross Lüsewitz; HT; ♂; DEIC. 

acuta. USA (Mississippi, Florida). Pinus elliottii; P. taeda (Pinaceae). 

Contarinia acuta Gagné 1984a: 222. USA: Florida, Gainesville; HT; ♂; USNM. 

aequalis. Widespread Europe. Senecio saracenicus; S. nemoralis (Asteraceae). 

Contarinia aequalis Kieffer 1898f: 61. Germany: Kreuznach; ST; ♂,♀; lost?. 

agrimoniae. USA (New York, Virginia). Agrimonia eupatoria (Rosaceae). 

Contarinia agrimoniae Felt 1907d: 21. USA: New York, Bath; ST; ♂,♀; NYSM. 

ajaguzensis. Kazakhstan. Tragopogon scoparius (Asteraceae). 

Contarinia ajaguzensis Fedotova 1995e: 23. Kazakhstan: Tarbagataï Mtn Range, 46 km E Ayaguz, upper Ayaguz R.; 


albotarsa. USA (New York). 

Cecidomyia albotarsa Felt 1907a: 36. USA: New York, Albany; HT; ♂; NYSM. 

Cecidomyia albotarsus Foote 1965: 289, unjust. emend. of albotarsa. 

167

168 

*allii. Kazakhstan. Allium trachyscordum (Liliaceae). 

Contarinia allii Fedotova 1992o: 120. Kazakhstan: Bet-Pak-Dala desert, 58 km N Furnanovka; HT; ♂; ZMAS. 

alloteropsidis. Australia (Queensland). Alloteropsis semialata (Poaceae). 

Contarinia alloteropsidis Harris 1979b: 179. Australia: Queensland, Cooktown; HT; ♀; QMBA. 

amenti. Italy. Quercus pubescens (Fagaceae). 

Contarinia amenti Kieffer 1909b: 23. Italy; ST; LT; lost?. 

ampelophila. USA (New York). 

Contarinia ampelophila Felt 1907a: 35. USA: New York, Albany; HT; ♂; NYSM. 

angreni. Kazakhstan. Astragalus angreni (Fabaceae). 

Contarinia angreni Fedotova 1993k: 91. Kazakhstan: foothills Talasskiï Alatau, Aksu R. valley, nr Belyye Vody; 


anthobia. Widespread Europe. Crataegus oxyacantha (Rosaceae). 

Diplosis anthobia Löw 1877: 16. Austria: vic. Vienna betw. Weidling & Kirling; ST; ♂,♀,L; NHMW. 

anthonoma. France. Sarothamnus scoparius (Fabaceae). 

Diplosis anthonoma Kieffer 1890d: 135. France: vic. Bitche; ST; ♀,L; lost?. 

anthophthora. Germany, Czechoslovakia, Austria, Yugoslavia. Verbascum orientale (Scrophulariaceae). 

Diplosis anthophthora Löw 1880: 36. Austria: Hundskogel nr Mödling; ST; ♂,♀,L; NHMW. 

aprilina. France. 

Contarinia aprilina Kieffer 1901c: 30. France: vic. Bitche; ST; ♂,♀; lost?. 

arrhenatheri. UK, France, Poland, Czechoslovakia. Arrhenatherum elatius (Poaceae). 

Contarinia arrhenatheri Kieffer 1901c: 33. France: Bitche; T; ♀; lost?. 

artemisiae. Germany, Latvia, Poland, Czechoslovakia. Artemisia vulgaris (Asteraceae). 

Contarinia artemisiae Rübsaamen 1917: 92. Germany; ST; ♂,♀,L; ZMHB. 

asclepiadis. Netherlands, Germany, Czechoslovakia, Austria. Asclepias vincetoxicum (Asclepiadaceae). 

Cecidomyia asclepiadis Giraud 1863: 1303. Austria; ST; ♂,♀,L; ?. 

asperulae. UK, France, Hungary. Asperula tinctoria (Rubiaceae). 

Contarinia asperulae Kieffer 1909b: 2. France; ST; LT; lost?. 

astragalicarpa. Kazakhstan. Astragalus unifoliatus (Fabaceae). 

Contarinia astragalicarpa Fedotova 1991c: 50. Kazakhstan: Buzachi Peninsula, 12 km SW Shebir; HT; ♂; ZMAS. 

atraphaxiflorae. Kazakhstan. Atraphaxis laetevirens (Polygonaceae). 

Contarinia atraphaxiflorae Fedotova 1988a: 62. Kazakhstan: Zailiïskiï Alatau, Alma-Ata Res., Pravyï Talgar canyon; 

HT; ♂; ZKAS. 

avenae. France, Russia (Europe). Avena pubescens (Poaceae). 

Contarinia avenae Kieffer 1901c: 32. France: Bitche; ST; ♂,♀,L; lost?. 

baeri. Widespread Europe; immigr.: Canada (New Brunswick), USA (Michigan, Pennsylvania). Pinus sylvestris 

(Pinaceae). 

Cecidomyia baeri Prell 1931: 51. Germany: Dresden; T; LT; ?. 

baggendorfi. UK, Germany, Czechoslovakia. Angelica sylvestris (Apiaceae). 

Contarinia baggendorfi Stelter 1982a: 75. Germany: Kr. Grimmen, Kirch Baggendorf; HT; ♂; Stelter. 

bajankolica. Kazakhstan. Galium septentrionale (Rubiaceae). 

Contarinia bajankolica Fedotova 1997c: 330. Kazakhstan: Terskei Alatau, upper course Bayankol R., 28 km SW 

Narynkol; HT; ♂; ZMAS. 

ballotae. UK, France, Italy, Germany, Poland, Czechoslovakia, Hungary, Yugoslavia, Greece. Ballota nigra 

(Lamiaceae). 

Contarinia ballotae Kieffer 1898f: 60. France: Aisne, Coeuvres; ST; ♂,♀; lost?. 

balsamifera. USA (New York). 

Contarinia balsamifera Felt 1907a: 34. USA: New York, Lake Clear; HT; ♂; NYSM. 

barbichei. Widespread Europe. Lotus corniculatus; L. uliginosus (Fabaceae). 

Diplosis barbichi Kieffer 1890a: 29. France: Metz, Bitche, Sierck, & Mengen; ST; ♂,♀,L; lost?. 

Contarinia barbichei Kieffer 1898a: 25, justified emendation of barbichi. 

*berulae. Kazakhstan. Berula erecta (Apicaeae). 

Contarinia berulae Fedotova 1992o: 123. Kazakhstan: Aigyrkumy desert, nr Ardynka; HT; ♂; ZMAS.

169 

beskokotovi. Kazakhstan. Onobrychis echidna (Fabaceae). 

Contarinia beskokotovi Fedotova 1996e: 48. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Kaska-Bulak pass, 

30 km SE Novonikolaevki; HT; ♂; ZMAS. 

bitensis. France. 

Stenodiplosis bitensis Kieffer 1909b: 35. France: Bitche; HT; ♀; lost?. 

bivalviae. India (Maharashtra). Acacia catechu (Fabaceae). 

Cincticornia bivalviae Rao 1952b: 35. India: Maharashtra, Thana; HT; ♀; ZSCI. 

bothriochloae. Australia (Queensland). Bothriochloa bladhii (Poaceae). 

Contarinia bothriochloae Harris 1979b: 175. Australia: Queensland, W Cooktown; HT; ♀; QMBA. 

brevipalpus. Australia (Queensland). Eragostis brownii (Poaceae). 

Contarinia brevipalpus Harris 1979b: 179. Australia: Queensland, nr Kennedy; HT; ♀; QMBA. 

brizae. Sweden, France, Czechoslovakia. Briza media (Poaceae). 

Contarinia brizae Kieffer 1896c: 99. France: Bitche; ST; ♂,L; lost?. 

*bulliformis. USA (Louisiana). Carya illinoiensis (Juglandaceae). 

Contarinia bulliformis Gagné 2008c: 91. USA: Louisiana, Caddo Parish, Greensleeves Orchard; HT; LT; USNM. 

bursariae. Australia (South Australia). Bursaria spinosa (Pittosporaceae). 

Contarinia bursariae Kolesik 1995d: 177. Australia: South Australia, Morialta Conservation Pk, 13 km NE Adelaide; 


calophacae. Kazakhstan. Calophaca soongorica (Fabaceae). 

Contarinia calophacae Fedotova 1989e: 36. Kazakhstan: Tarbagataï Mtn Range, 25 km S Ayaguz; HT; ♂; ZKAS. 

campanulae. UK, Sweden, France, Germany, Poland, Czechoslovakia, Romania, Yugoslavia. Campanula 

rapunculoides (Campanulaceae). 

Eudiplosis campanulae Kieffer 1895p: 9. France; T; A; lost?. 

camphorosmae. Spain. Camphorosma monspeliaca (Chenopodiaceae). 

Navasodiplosis camphorosmae Tavares 1920c: 66. Spain: Saragossa; ST; ♀,P,L; TAVC. 

canadensis. Canada (Ontario). Poss. Fraxinus sp. (Oleaceae). 

Contarinia canadensis Felt 1908b: 394. Canada: Ontario, Toronto; ST; ♂,♀; USNM. 

capparis. Kazakhstan. Capparis spinosa (Capparidaceae). 

Contarinia capparis Fedotova 1983b: 43. Kazakhstan: right bank Ili R., Kapchagaïskoe reservoir, nr "Singing 

Mountain"; HT; ♂; ZMAS. 

caraganae. Kazakhstan. Caragana frutex (Fabaceae). 

Desertomyia caraganae Fedotova 1990g: 57. Kazakhstan: Tarbagataï Mtn Range, Ashisu R. floodplain, 30 km N 

Ayaguz Mtn; HT; ♂; ZKAS. Placed here because it resembles in every respect a Contarinia except that it has 

tricircumfilar male flagellomeres, as do other Contarinia spp. (cf. C. catalpae and C. tecomae). 

caraganicola. Kirghizstan, Russia (Western Siberia). Caragana laucophela (Fabaceae). 

Contarinia caraganicola Marikovskij 1955e: 24. Kirghizstan: Kapkak valley, western end Issyk-Kul' L.; ST; ♂,♀; ?. 

cardariae. Kazakhstan. Cardaria draba; C. pubescens (Brassicaceae). 

Contarinia cardariae Fedotova 1994b: 82. Kazakhstan: Buzachi Peninsula, 25 km E Shebir, Mastek; HT; ♂; ZMAS. 

carolinae. Kenya. Acacia senegal (Fabaceae). 

Contarinia carolinae Gagné 1993c: 112. Kenya: Kima, nr Sultan Hamid; HT; ♂; USNM. 

carpini. Widespread Europe; Pliocene/Germany. Carpinus betulus (Betulaceae). 

Contarinia carpini Kieffer 1897c: 15. France; ST; ♀; lost?. 

catalpae. USA (Indiana, Maryland, District of Columbia). Catalpa bignonioides; C. speciosa (Bignoniaceae). 

Diplosis catalpae Comstock 1881: 266. USA: District of Columbia; ST; ♂,♀,L; USNM. 

caudata. India (Tamil Nadu). Apluda varia, Andropogon schoenanthus (Poaceae). 

Contarinia caudata Felt 1920b: 4. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

cerasiphila. USA (Colorado). Prunus sp. (Rosaceae). 

Cecidomyia cerasiphila Felt 1911l: 466, nomen nudum. 

Cecidomyia cerasiphila Felt 1911n: 554. USA: Colorado, Glen Ayrie; HT; ♀; USNM. 

cerasiserotinae. USA (Nebraska, New York, Maryland, Virginia). Prunus serotina (Rosaceae). 

Cecidomyia cerasiserotinae (as cerasi serotinae) Osten Sacken 1871: 346. USA: New York, Tarrytown; T; LT; lost. 

cerriperda. Czech Republic. Quercus cerris (Fagaceae).

170 

Contarinia cerriperda Skuhravá 1991: 407. Czech Republic: Mly�any; HT; ♂; SKCR. 

chrysanthemi. UK, Denmark, France, Czechoslovakia, Russia (Europe). Chrysanthemum leucanthemum (Asteraceae). 

Eudiplosis chrysanthemi Kieffer 1895p: 9. France; T; A; lost?. 

citri. Mauritius, Italy, Cyprus?, Turkey. Citrus sp. (Rutaceae). 

Contarinia citri Barnes 1944b: 212. Mauritius: Beau Bassin; ST; ♂,♀; BMNH. 

citrina. USA (Illinois, New York, Massachusetts, Maryland). Tilia americana (Tiliaceae). 

Cecidomyia tiliae citrinae Osten Sacken 1870: 53, invalid, a polynomial. 

Cecidomyia citrina Osten Sacken 1878: 6. USA: New York; T; G; USNM. 

Cecidomyia citricola Felt 1915k: 58, error for citrina Osten Sacken. 

clarkei. USA (New York, Massachusetts). Spiraea sp. (Rosaceae). 

Hormomyia clarkei Felt 1908b: 388. USA: Massachusetts, Magnolia; HT; ♀; NYSM. 

Trishormomyia clarkeae Sabrosky 1967: 119, unjust. emend. of clarkei. 

clematidis. USA (New York, Massachusetts). Clematis sp. (Rosaceae). 

Contarinia clematidis Felt 1908b: 393. USA: New York, Newport; HT; ♂; NYSM. 

cockerelli. USA (Colorado). Pinus edulis (Pinaceae). 

Thecodiplosis cockerelli Felt 1918b: 381. USA: Colorado, Oxford; HT; ♀; NYSM. 

coffeae. Uganda. Coffea arabica (Rubiaceae). 

Contarinia coffeae Harris 1970: 671. Uganda: Toro, Kicheche; HT; ♂; BMNH. 

coloradensis. USA (Colorado). Pinus scopulorum (Pinaceae). 

Contarinia coloradensis Felt 1912l: 240. USA: Colorado, Denver; ST; ♀,P,L; NYSM. 

constricta. Canada (British Columbia), USA (Washington, Oregon, California). Pseudotsuga menziesii (Pinaceae). 

Contarinia constricta Condrashoff 1961: 126. Canada: British Columbia, Oyama; HT; ♂; CNCI. 

convallaria. Germany. Convallaria majalis (Liliaceae). 

Contarinia convallaria Rübsaamen 1925: 47. Germany; T; LT; ZMHB. 

convolvulicola. Kazakhstan. Convolvulus tragacantoides; C. fruticosus (Convolvulaceae). 

Contarinia convolvulicola Fedotova 1991c: 41. Kazakhstan: Zailiïskiï Alatau, SW end Boguty Mts nr Kop-Pek, 160 

km NE Alma-Ata; HT; ♂; ZMAS. 

convolvuliflora. Kazakhstan. Convolvulus fruticosus (Convolvulaceae). 

Contarinia convolvuliflora Fedotova 1991c: 43. Kazakhstan: Karakumy Desert, Kulzhan Valley, 22 km SE Opornaya 

Stn; HT; ♂; ZMAS. 

coronillae. Slovenia, Yugoslavia. Coronilla emerus, C. emeroides (Fabaceae). 

Contarinia coronillae Jane�i 1978: 14. Slovenia: vic. Portoro�; ST; ♂,♀,L; ?. 

coryli. Widespread Europe, China. Corylus sp. (Betulaceae). 

Cecidomyia coryli Kaltenbach 1859: 284. Germany; ST; LT; NHMW?. 

Diplosis corylina Löw 1878: 396. Austria: vic. Vienna; ST; ♂,♀,L; NHMW. 

cybelae. UK, Sweden, France, Latvia, Poland, Czechoslovakia. Corylus avellana (Betulaceae). 

Contarinia coryli Kieffer 1909b: 6, jun. sec. hom. coryli Kaltenbach 1859. France; T; LT; lost?. 

Contarinia cybelae Gagné 1972b: 322, new name for Contarinia coryli Kieffer. 

cotini. Czechoslovakia to Serbia, E to Ukraine. Rhus cotinus (Anacardiaceae). 

Contarinia cotini Kieffer 1901d: 170. Hungary: Budapest; ST; ♂,♀; lost?. 

craccae. Widespread Europe, Russia (Western Siberia). Vicia cracca; Vicia spp. (Fabaceae). 

Cecidomyia craccae Loew 1850a: 22. Poland; ST; LT; ZMHB?. 

Contarinia craccae Kieffer 1897c: 15. France; ST; ♂,♀; lost?. 

crispans. UK, France, Germany, Czechoslovakia, Romania. Valeriana officinalis (Valerianaceae). 

Contarinia crispans Kieffer 1909b: 31. France; ST; LT; lost?. 

cucubali. UK, Italy. Silene vulgaris (Caryophyllaceae). 

Contarinia cucubali Kieffer 1909b: 28. Italy; ST; LT; lost?. 

*cucumata. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). 

Contarinia cucumata Gagné 2008c: 92. USA: Maryland, Clarksburg; HT; LT; USNM. 

cuniculator. Canada (British Columbia), USA (Washington, Oregon, California). Pseudotsuga menziesii (Pinaceae). 

Contarinia cuniculator Condrashoff 1961: 128. Canada: British Columbia, Oyama; HT; ♂; CNCI. 

czalikovae. Kazakhstan. Medicago tianschanica (Fabaceae).

171 

Contarinia czalikovae Fedotova 1996e: 46. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Baldabrek Mtn 

Range, Chooldak area, 57 km NE Georgievka; HT; ♂; ZMAS. 

dactylidis. UK, Poland, Czechoslovakia, Russia (Europe), Greece. Dactylis glomerata (Poaceae). 

Cecidomyia dactylidis Loew 1851: 383. Poland; T; ♀; ZMHB?. 

dalbergiae. India (Uttar Pradesh). Dalbergia sp. (Fabaceae). 

Contarinia dalbergiae Mani 1943: 43. India: Uttar Pradesh, Dehra Dun; HT; ♂; INPC. 

Sissudiplosis chatterjeei Mani 1943: 44. India: Uttar Pradesh, Dehra Dun; HT; ♂; INPC. 

desertophila. Kazakhstan. Astragalus alopecias; Astragalus spp. (Fabaceae). 

Contarinia desertophila Fedotova 1993k: 98. Kazakhstan: middle course Ili R., nr Kapchagay; HT; ♂; ZMAS. 

desertorum. Armenia, Kazakhstan. Alhagi pseudalhagi (Fabaceae). 

Contarinia desertorum Marikovskij 1955b: 105, nomen nudum. 

Contarinia desertorum Marikovskij 1961: 48. Kazakhstan: Kzyl-Orda Region, loess desert of Daryalyk; ST; ♂,♀; 

ZMAS. 

dichanthii. Australia (Northern Territory, Queensland). Dichanthium annulatum var. fecundum; Dichanthium spp. 

(Poaceae). 

Contarinia dichanthii Harris 1979b: 176. Australia: Queensland, Bullock Creek; HT; ♀; QMBA. 

digitata. Germany, Poland. 

Cecidomyia digitata Loew 1850a: 39. Poland; ST; ♀; ZMHB?. 

Cecidomyia digitata Winnertz 1853: 272. Germany; ST; ♂,♀; lost. 

dipsacearum. Sweden, Netherlands, Germany, Latvia, Russia (Europe), Yugoslavia. Knautia arvensis, Succisa 

pratensis (Dipsacaceae). 

Contarinia dipsacearum Rübsaamen 1921: 45. Germany: Prignitz, Triglitz, & Eibsee nr Oberstdorf; ST; ♂,♀,L; 

ZMHB. 

divaricata. USA (New York). 

Contarinia divaricata Felt 1908b: 392. USA: New York, Albany; HT; ♂; NYSM. 

echii. France, Czechoslovakia, Romania. Echium vulgare (Boraginaceae). 

Eudiplosis echii Kieffer 1895p: 9. France; T; A; lost?. 

elaeagniflorae. Kazakhstan. Elaeagnus oxycarpa (Elaeagnaceae). 

Contarinia elaeagniflorae Fedotova 1988a: 62. Kazakhstan: left bank middle course Karatal R., nr Ush-Tobe; HT; ♂; 

ZKAS. 

enceliae. USA (California). Encelia californica (Asteraceae). 

Rhopalomyia enceliae Felt 1916e: 183. USA: California; ST; ♀; NYSM. 

eragrostidis. Indonesia (Java). Eragrostis unioloides (Poaceae). 

Cecidomyia eragrostidis Felt 1927b: 381. Indonesia: Java; ST; ♂,♀; NYSM. 

erigeronis. France. Erigeron acer (Asteraceae). 

Contarinia erigeronis Kieffer 1909b: 7. France; ST; LT; lost?. 

erucastris. Kazakhstan. Erucastrum armoracoides (Brassicaceae). 

Contarinia erucastris Fedotova 1994b: 83. Kazakhstan: Ili R. delta, Topar R. floodplain, nr Zhil-Turanga; HT; ♂; 

ZMAS. 

excavationis. USA (New York). 

Cecidomyia excavationis Felt 1907a: 42. USA: New York, Albany; HT; ♂; NYSM. 

Cecidomyia excavata Felt 1907c: 139, error for excavationis Felt. 

fagi. Widespread Europe. Fagus sylvatica (Fagaceae). 

Contarinia fagi Rübsaamen 1921: 44. Germany: Partenkirchen; ST; ♂,♀,L; ZMHB. Ref.: Goffau & Nijveldt 2005 

(morph., biol.). 

festucae. UK, Sweden, Germany, Poland, Czechoslovakia, Bulgaria. Festuca pratensis, F. rubra (Poaceae). 

Contarinia festucae Jones 1940: 539. UK: Wales, Aberystwyth; ST; ♂,♀; BMNH. 

fimbristylidis. Australia (Northern Territory). Fimbristylis bisumbellata (Cyperaceae). 

Contarinia fimbristylidis Harris 1979b: 180. Australia: Northern Territory, Swim Creek; HT; ♀; QMBA. 

fitchii. USA (New York). 

Contarinia fitchii Felt 1912e: 288. USA: New York; LT (Gagné 1966a: 1155); ♂; NYSM. 

flavolinea. USA (New York).

172 

Contarinia flavolinea Felt 1908b: 392. USA: New York, Albany; HT; ♂; NYSM. 

floricola. UK, Sweden, Germany, Poland, Czechoslovakia. Poa sp.; P. pratensis, P. trivialis (Poaceae). 

Phytophaga floricola Oettingen 1927: 660. Poland: Szczecin; ST; ♂,♀,L; BMNH. 

Contarinia poae Barnes 1946b: 100, with ref. to Tomaszewski 1931. Germany: Randowbruch; ST; ♂,♀,L; BMNH. 

floriperda. Widespread Europe. Sorbus aucuparia (Rosaceae). 

Contarinia floriperda Rübsaamen 1917: 89. Germany: Prignitz, Triglitz; ST; ♂,♀; ZMHB. 

florum. Germany, Poland, Czechoslovakia, Hungary. Asparagus officinalis, Convallaria majalis, Polygonatum 

multiflorum (Liliaceae). 

Contarinia florum Rübsaamen 1917: 90. Poland: Zielona Gora, & Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

forskalei. Egypt. Astragalus forskalei (Fabaceae). 

Contarinia forskalei Debski 1918: 21. Egypt: Wadi Abou-Zouleiqa, nr Hélouan; ST; LT; ?. 

fraxini. USA (New York). Fraxinus sp. (Oleaceae). 

Phytophaga fraxini Felt 1915j: 206. USA: New York, Poughkeepsie; ST; ♂,♀,L; NYSM. 

*galatellae. Kazakhstan. Galatella punctata (Asteraceae). 

Contarinia galatellae Fedotova 2003a: 831. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE Alma-Ata; 


galeobdolontis. France, Germany. Lamium galeobdolon (Lamiaceae). 

Contarinia galeobdolontis Kieffer 1909b: 12. France; ST; LT; lost?. 

galii. France, Greece. Galium lucidum (Rubiaceae). 

Contarinia galii Kieffer 1909b: 9. France; ST; LT; lost?. 

gei. Widespread Europe. Geum rivale; G. urbanum (Rosaceae). 

Contarinia gei Kieffer 1909b: 9. Germany; T; G; ZMHB?. 

Contarinia geicola Rübsaamen 1917: 93. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

gemmae. Brazil (Rio de Janeiro). Calophyllum brasiliense (Clusiaceae). 

Contarinia gemmae Maia 2003: 45. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

glycyrrhizae. Kazakhstan. Glycyrrhiza uralensis; G. glabra (Fabaceae). 

Contarinia glycyrrhizae Fedotova 1983b: 42. Kazakhstan: south bank L. Balkhash, Karatal R., nr Golovnoe; HT; ♂; 

ZMAS. 

goebeliae. Kazakhstan. Goebelia alopecuroides (Fabaceae). 

Contarinia goebeliae Fedotova 1987a: 39. Kazakhstan: right bank Lepsy R., 8 km NW Saratovki; HT; ♂; ZKAS. 

gossypii. Colombia, U.S. Virgin Is, Antigua, Montserrat, Barbados. Gossypium sp. (Malvaceae). 

Contarinia gossypii Felt 1908a: 210. Antigua; ST; ♂,♀; USNM. 

halimodendronis. Kazakhstan. Halimodendron halodendron (Fabaceae). 

Contarinia halimodendronis Fedotova 1991c: 46. Kazakhstan: Karatal R. valley, 60 km NW Ush-Tobe; HT; ♂; 

ZMAS. 

hedysari. Kazakhstan. Hedysarum cephalotes (Fabaceae). 

Contarinia hedysari Fedotova 1993f: 680. Kazakhstan: Talasskiï Alatau, Ul'Ken Kaïndy gorge, 30 km SE Vannovka; 


hedysarocarpi. Kazakhstan. Hedysarum semenovii (Fabaceae). 

Contarinia hedysarocarpi Fedotova 1993f: 677. Kazakhstan: NE spurs Dzhungarskiï Alatau, Kungeï Mts, 78 km SE 

Uch-Aral, 8 km SE Koktum; HT; ♂; ZKAS. 

helianthemi. UK, Germany, Poland, Czechoslovakia, Austria, Yugoslavia, Tunisia. Helianthemum vulgare 

(Asteraceae). 

Cecidomyia helianthemi Hardy 1850: 187. UK: Scotland, vic. Penmanshiel; ST; ♂,L; lost. 

heptapotamica. Kazakhstan. Astragalus polyceras; Astragalus spp. (Fabaceae). 

Contarinia heptapotamica Fedotova 1991c: 48. Kazakhstan: SE L. Balkhash, Karatau Mtn Range, Arkharly Mtn 

ridge, 64 km NE Saratovka; HT; ♂; ZMAS. 

heraclei. UK, Germany, Latvia, Poland, Czechoslovakia. Heracleum sphondylium (Apiaceae). 

Diplosis heraclei Rübsaamen 1889b: 373. Germany: Berlin, vic. Bruch'sches Walzwerk; ST; ♂,♀,L; ZMHB. 

hongoi. Kenya. Acacia drepanolobium (Fabaceae). 

Contarinia hongoi Gagné 1993c: 114. Kenya: southern base Ngong Hills; HT; ♂; USNM. 

hudsonici. USA (New York).

173 

Contarinia hudsonici Felt 1908b: 393. USA: New York, Poughkeepsie; HT; ♂; NYSM. 

humuli. UK, Austria, Bulgaria. Humulus lupulus (Cannabaceae). 

Diplosis humuli Theobald 1909: 152. UK: England, many localities in the Weald, Kent, Hampshire, Surrey, & 

Worcestershire; T; LT; ?. 

Contarinia humuli Tölg 1921: 33. Austria: vic. Vienna; ST; ♂,♀; ?. 

hydrangeae. Japan (Honshu). Hydrangea paniculata (Hydrangeaceae). 

Contarinia hydranthi Shinji 1939c: 583, nomen nudum. 

Contarinia hydrangeae Shinji 1939f: 130. Japan: Honshu, Morioka; ST; ♂,♀; lost. 

hyperici. UK. Hypericum perforatum (Hypericaceae). 

Contarinia hyperici Barnes 1952: 700. UK: England, Surrey, Box Hill; ST; ♂,♀; BMNH. 

hypochoeridis. Widespread northern Europe. Hypochaeris radicata (Asteraceae). 

Diplosis hypochoeridis Rübsaamen 1891b: 1. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

ilicis. France, Portugal, Italy, Greece, Algeria. Quercus ilex (Fagaceae). 

Contarinia ilicis Kieffer 1898f: 61. France & Algeria; T; ♀; lost?. 

inouyei. Japan (Hokkaido to Kyushu). Cryptomeria japonica (Taxodiaceae). 

Contarinia inouyei Mani 1954b: 112. Japan: Kyushu, Kagoshima; HT; ♂; ZSCI. 

inquilina. Germany, Czechoslovakia. Pimpinella saxifraga (Apiaceae). 

Contarinia inquilina Rübsaamen 1917: 94. Germany; ST; ♂,♀,L; ZMHB. 

intrans. Australia (Northern Territory). Sorghum intrans (Poaceae). 

Contarinia intrans Harris 1979b: 173. Australia: Northern Territory, Cullen R.; HT; ♀; QMBA. 

inulicola. Germany, Czechoslovakia, Romania. Inula britannica (Asteraceae). 

Contarinia inulicola Stelter 1965: 98. Germany: Bad Sülze in Mecklenberg; HT; ♂; DEIC. 

ishkovi. Kazakhstan. Polygonum alpinum; P. hissaricum, P. coriarium (Polygonaceae). 

Contarinia ishkovi Fedotova 1997g: 879. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Baldabrek Mtn 

Range, 57 km NE Georgievka; HT; ♂; ZMAS. 

istriana. Croatia, Yugoslavia. Coronilla emerus, C. emeroides (Fabaceae). 

Contarinia istriana Jane�i 1980: 49. Croatia: Istria, Buje; ST; ♂,♀,L; ?. 

jaapi. UK, Germany, Poland, Czechoslovakia, Yugoslavia. Lathyrus pratensis (Fabaceae). 

Contarinia jaapi Rübsaamen 1914: 110. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

jacobaeae. Widespread Europe. Senecio jacobaea; S. vulgaris (Asteraceae). 

Cecidomyia jacobaeae Loew 1850a: 33. Poland; ST; ♂; ZMHB?. 

Diplosis senecionis Rübsaamen 1891c: 43. Germany: Giersberg nr Siegen; ST; ♂,L; ZMHB. 

johnsoni. USA (New York). Vitis sp. (Vitaceae). 

Contarinia johnsoni Slingerland & Johnson 1904: 72, nomen nudum. 

Contarinia johnsoni Felt 1909c: 15. USA: New York, Chatauqua Co.; ST; ♂,♀,L; NYSM. 

juniperina. Canada (Ontario), USA (Nebraska, Kansas, Missouri, Ohio, Pennsylvania). Juniperus sabinoides 

(Cupressaceae). 

Contarinia juniperina Felt 1939b: 159. USA: Missouri, Columbia; ST; ♂,♀; USNM. 

juniperiramea. Kazakhstan. Juniperus turkestanica (Cupressaceae). 

Contarinia juniperiramea Fedotova 1985d: 1105. Kazakhstan: Western Tyan'-Shan', Ugamskiï Mtn Range, nr 

Pervomaïskiï; HT; ♀; ZKAS. 

kanervoi. UK, Sweden, Finland. Phleum pratense (Poaceae). 

Contarinia kanervoi Barnes 1958a: 60. Finland: Nivala; ST; ♂,♀; BMNH. 

karataliensis. Kazakhstan. Sisymbrium loeselii (Brassicaceae). 

Contarinia karataliensis Fedotova 1994b: 83. Kazakhstan: middle part Karatal R., 42 km SW Taldy-Kurgan, nr 

Golovnoye; HT; ♂; ZMAS. 

karatavica. Kazakhstan. Silene guntensis (Caryophyllaceae). 

Contarinia karatavica Fedotova 1994e: 46. Kazakhstan: Karatau Mtn Range, nr Karatau; HT; ♂; ZMAS. 

kiefferi. Germany, Poland. Sisymbrium sophia (Brassicaceae). 

Diplosis kiefferi Schlechtendahl 1891: 52. Germany: Zwickau; T; LT; ?. 

*kochiae. Kazakhstan. Kochia prostrata (Chenopodiaceae). 

Contarinia kochiae Fedotova 1988b: 40. Kazakhstan: Tarbagataï Mtn Range, 60 SW Ayaguz; HT; ♂; ZKAS.

174 

kurenzovi. Russia (Far East). Phellodendron amurense (Rutaceae). 

Contarinia kurenzovi Kovalev 1972: 419. Russia: Primorskiï Terr., Suputinskiy Res.; HT; ♂; ZMAS. 

lamii. Portugal, Spain. Lamium maculatum (Lamiaceae). 

Contarinia lamii Kieffer 1909b: 13. Europe: S; ♂,L; lost?. 

lamiicola. Germany, Czechoslovakia, Hungary. Lamium maculatum (Lamiaceae). 

Contarinia lamiicola Rübsaamen 1916b: 495. Germany; ST; ♂,♀; ZMHB. 

lathyri. UK, France, Hungary, Yugoslavia, Russia (Western Siberia). Lathyrus pratensis (Fabaceae). 

Contarinia lathyri Kieffer 1909b: 13. France; ST; LT; lost?. 

lentis. France, Czechoslovakia, Hungary. Lens culinaris (Fabaceae). 

Contarinia lentis Aczél 1944: 166 (publ'n not seen). Hungary: Alsózsolca (Komitat Borsod); ?; ?; ?. 

lepidii. Germany. Lepidium draba (Brassicaceae). 

Contarinia lepidii Kieffer 1909b: 14. Germany; T; G; ZMHB. 

lespedezifolia. Russia (Far East). Lespedeza bicolor (Fabaceae). 

Contarinia lespedezifolia Kovalev 1972: 418. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

liliacea. Sweden. Lilium candidum (Liliaceae). 

Contarinia liliacea Wahlgren 1957: 203. Sweden; ST; A,L; ?. 

lilii. Austria. Lilium martagon (Liliaceae). 

Contarinia lilii Kieffer 1909b: 14. Austria; T; G; NHMW?. 

*limonii. Kazakhstan. Limonium gmelinii (Plumbaginaceae). 

Contarinia limonii Fedotova 1992o: 118. Kazakhstan: 6 km E Buran; HT; ♂; ZMAS. 

*lini. Serbia. Linum austriacum (Linaceae). 

Contarinia lini Simova & Skuhravá 2007; 144. Serbia: Novi Beograd; HT; ♂; Natural History Museum, Belgrade. 

lolii. UK. Lolium pratense (Poaceae). 

Contarinia lolii Metcalfe 1933: 336. UK: England, Harpenden; ST; ♂,♀; BMNH. 

lonicerae. Austria, Italy, Yugoslavia. Lonicera coerulea (Caprifoliaceae). 

Contarinia lonicerae Kieffer 1909b: 14. Austria: S. Tyrol, Bad Ratzes; T; G; NHMW?. 

loti. Widespread Europe. Lotus corniculatus (Fabaceae). 

Tipula loti De Geer 1776: 420 (publ'n not seen). Sweden; ?; ?; ?. 

luteola. Portugal, Italy, Yugoslavia. Quercus coccifera (Fagaceae). 

Contarinia luteola Tavares 1902b: 34. Portugal: Perto do Sobral do Campo e Soalheira, Louza e Oledo, & Villa 

Velha de Rodão; ST; ♂,♀,L; TAVC. 

lycii. Egypt. Lycium arabicum (Solanaceae). 

Contarinia lycii Debski 1918: 33. Egypt: Wadi Rished, Wadi Abu-Zouleiqa; ST; LT; ?. 

lyciicola. Kazakhstan. Lycium dasystemum (Solanaceae). 

Contarinia lycii Fedotova 1983b: 44, jun. prim. hom. lycii Debski. Kazakhstan: Ili R. delta, nr Karoï; HT; ♂; ZMAS. 

Contarinia lyciicola Fedotova 1985b: 35, new name for Contarinia lycii Fedotova. 

lycopersici. Belize, West Indies (Guadeloupe to Trinidad), Guyana. Lycopersicum esculentum (Solanaceae). 

Contarinia lycopersici Felt 1911j: 303. St. Vincent; ST; ♂,♀,L; NYSM. 

lysimachiae. Sweden, Denmark, Germany, Latvia, Poland, Czechoslovakia, Yugoslavia. Lysimachia vulgaris 

(Primulaceae). 

Diplosis lysimachiae Rübsaamen 1893: 162. Germany: vic. Berlin; T; ♀; ZMHB. 

maculipennis. Thailand, Vietnam, Taiwan; immigr.: Japan, Hawaiian Is, USA (not estab., California, Florida), 

Guadeloupe. Hibiscus sp. (Malvaceae); Brassica chinensis (Brassicaceae), Dendrobium spp. (Orchidaceae), 

Jasminum spp. (Oleaceae), Lycopersicon spp., Solanum spp. (Solanaceae), Momordica charantia 

(Cucurbitaceae). 

Contarinia maculipennis Felt 1933b: 247. Hawaiian Is: Honolulu; ST; ♂,♀; USNM. Refs: Gagné 1995b, Tokuda et 

al. 2002. 

maculosa. USA (New York). 

Contarinia maculosa Felt 1908b: 393. USA: New York, Albany; HT; ♂; NYSM. 

majanthemi. Germany. Maianthemum bifolium (Liliaceae). 

Contarinia majanthemi Rübsaamen 1925: 47. Germany; T; LT; ZMHB. 

mali. Japan (Honshu). Pyrus malus (Rosaceae).

175 

Contarinia mali Barnes 1939d: 126. Japan: Honshu, Nagano Pref.; ST; ♂,♀; BMNH. 

marchali. UK, France, Germany, Poland, Czechoslovakia, Hungary, Yugoslavia, Ukraine, Russia (Europe), Algeria. 

Fraxinus excelsior (Oleaceae). 

Contarinia marchali Kieffer 1896f: 236. France: Bitche; ST; A,L; lost?. 

martagonis. Germany, Czechoslovakia, Austria. Lilium martagon (Liliaceae). 

Contarinia martagonis Kieffer 1909b: 14. Austria; ST; LT; lost?. 

matusintome. Korean Peninsula, Japan (Honshu to Kyushu). Pinus densiflora; P. thunbergii (Pinaceae). 

Contarinia matusintome Haraguti & Monzen 1955b: 44. Japan: Honshu, Morioka; ST; ♂,♀; Monzen. 

Contarinia tonensis Uchida & Inouye 1956: 34. Japan: Honshu; HT; ♂; HFES. 

medicaginis. Widespread Europe, Armenia, Russia (Western Siberia). Medicago sativa (Fabaceae). 

Contarinia medicaginis Kieffer 1895q: 150. France: Moselle; ST; ♂,L; lost?. 

melanocera. UK, France, Germany, Poland, Czechoslovakia, Hungary, Romania, Yugoslavia. Genista tinctoria 

(Fabaceae). 

Contarinia melanocera Kieffer 1904a: 76. France: Alençon & Metz; ST; ♂,♀,P,L,egg; lost?. 

melissitis. Kazakhstan. Melissitus popovi (Fabaceae). 

Contarinia melissitis Fedotova 1992g: 53. Kazakhstan: Talasskiï Alatau, 10 km S Kaskasu; HT; ♂; ZMAS. 

merceri. Widespread Europe. Alopecurus pratensis (Poaceae). 

Contarinia merceri Barnes 1930a: 352. UK (many localities throughout) & Finland; ST; ♂,♀,L; BMNH. 

meristotropea. Kazakhstan. Meristotropis triphylla (Fabaceae). 

Contarinia meristotropeus Fedotova 1993f: 679. Kazakhstan: Karatau Mtn Range, foothills Kuyuk Mts, 10 km S L. 

Bilikul', Bilikul'; HT; ♂; ZKAS. 

minima. Italy. Quercus ilex (Fagaceae). 

Contarinia minima Kieffer 1909b: 22. Italy: Sicily; T; LT; lost?. 

molluginis. UK, Denmark, Germany, Lithuania, Czechoslovakia, Italy, Yugoslavia. Galium mollugo (Rubiaceae). 

Diplosis molluginis Rübsaamen 1889a: 51. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

montana. Kazakhstan. Astragalus alpinus; A. lasiosemius, A. tibetanus (Fabaceae). 

Contarinia montana Fedotova 1993k: 94. Kazakhstan: NE spurs Kungeï Alatau, 20 km NE Saryzhas; HT; ♂; ZMAS. 

morindae. India (Uttar Pradesh). Morinda tinctoriae (Rubiaceae). 

Stictodiplosis morindae Grover 1966b: 25. India: Uttar Pradesh, Allahabad; ST; ♂,♀; PGCA. 

moringae. India (Tamil Nadu). Moringa sp. (Moringaceae). 

Stictodiplosis moringae Mani 1936: 195. India: Tamil Nadu, Coimbatore; ST; ♂,♀; ZSCI. 

mucidus. Kazakhstan. Astragalus mucidus; A. flexus (Fabaceae). 

Contarinia mucidus Fedotova 1993k: 92. Kazakhstan: Moyynkum Desert, 17 km NE Sarykoby, 100 km NE Tatty; 


nasturtii. Widespread Europe, Turkey; immigr.: Canada (Ontario, Quebec), USA (New York). Nasturtium palustre; 

many other genera and species of Brassicaceae, incl. Brassica spp. (Barnes 1946a). 

Diplosis nasturtii Kieffer 1888c: 263. France: Bitche; ST; ♂,♀,L; lost?. 

Diplosis ruderalis Kieffer 1890e: 198. France: vic. Bitche; ST; ♂,♀,L; lost?. 

Contarinia torquens Meijere 1906: 18. Netherlands: North Holland, Schagen; ST; ♂,♀,P,L; ZMAN. 

Contarinia isatidis Rübsaamen 1910f: 421. Germany: Erpel nr Linz & nr Casbach; ST; ♀,L; ZMHB. 

Contarinia perniciosa Rübsaamen 1914: 112. Germany: vic. Coblenz; ST; ♂,♀; ZMHB. 

Contarinia tudensis Tavares 1916b: 109. Spain: Gallicia, Tuy; ST; ♀; TAVC. 

Contarinia gallaica Tavares 1916c: 192. Spain: Tuy & Pontevedra; ST; ♂,♀; TAVC. 

Contarinia pontevedrensis Tavares 1916c: 188. Spain: S. Teresa, vic. Pontevedra; ST; ♂,♀; TAVC. 

Contarinia geisenheyneri Rübsaamen 1917: 92. Germany; ST; ♂,♀,L; ZMHB. 

negundinis. Widespread eastern Nearctic. Acer negundo (Aceraceae). 

Cecidomyia negundinis Gillette 1890: 392. USA, Iowa, Ames, Iowa State University; ST; ♀,egg; ISUI. 

Contarinia negundifolia Felt 1908b: 394. USA: Virginia; ST; ♂,♀; USNM. 

nicolayi. UK, Netherlands, Germany, Poland, Czechoslovakia, Hungary, Romania. Heracleum spondylium (Apiaceae). 

Eudiplosis nicolayi Rübsaamen 1895b: 183. Germany; ST; ♂,♀; ZMHB. 

nitensis. Kazakhstan. Polygonum nitens (Polygonaceae). 

Contarinia nitensis Fedotova 1997g: 876. Kazakhstan: southern Altaï, Markakol'skiï Res., L. Markakol', nr

176 

Urunkhaïka, 60 km N Alekseyevka; HT; ♂; ZMAS. 

nitrariae. Kazakhstan. Nitraria sibirica (Zygophyllaceae). 

Contarinia nitrariae Fedotova 1988a: 64. Kazakhstan: Dzhungarskiï Alatau, nr Panfilova, 10 km S Koktal; HT; ♂; 

ZKAS. 

nitrariagemmae. Kazakhstan. Nitraria sibirica (Zygophyllaceae). 

Contarinia nitrariagemmae Fedotova 1988a: 65. Kazakhstan: right bank Ili R., 63 km NW Bakanas, 4 km NW 

Akkol'; HT; ♂; ZKAS. 

nubilipennis. France. 

Diplosis nubilipennis Kieffer 1889a: 150. France: Bitche; T; ♀; lost?. 

obesa. USA (New York). 

Contarinia obesa Felt 1918e: 110. USA: New York, Albany; HT; ♂; NYSM. 

okadai. Japan (Honshu to Kyushu). Citrus unshiu (Rutaceae). 

Diplosis okadai Miyoshi 1926: 342. Japan; NT (Harris & Yukawa 1980: 279; Japan: Yamaguchi Pref., Ooshima, 

Agenosho); ♂; KUIC. 

onobrychidis. UK, France, Germany, Czechoslovakia, Hungary, Ukraine, Romania, Bulgaria, Armenia, Russia 

(Western Siberia). Onobrychis sativa (Fabaceae). 

Contarinia onobrychidis Kieffer 1895q: 150. France: Moselle; ST; ♂,♀,L; lost?. 

ononidis. UK, France, Czechoslovakia. Ononis repens (Fabaceae). 

Contarinia ononidis Kieffer 1899b: 104. France: Seine Maritime, vic. Les Petites Dalles; ST; ♂,♀,P,L; lost?. 

opuntiae. USA (Texas), Mexico. Opuntia sp. (Cactaceae). 

Cecidomyia opuntiae Felt 1910a: 10. USA: New York, New York, Bronx Botanical Garden; ST; ♂,♀,L; NYSM. 

oregonensis. Canada (British Columbia), USA (Washington, Oregon). Pseudotsuga menziesii (Pinaceae). 

Contarinia oregonensis Foote 1956: 54. USA: Oregon; HT; ♂; USNM. 

orientalis. India (Maharashtra). Bothriochloa pertusa (Poaceae). 

Bothriochloamyia orientalis Rao & Sharma 1977b: 237. India: Maharashtra, Aurangabad; HT; ♂; RCMU. 

ovipositosclera. Kazakhstan. Astragalus cognatus; Astragalus spp. (Fabaceae). 

Contarinia ovipositosclera Fedotova 1993k: 94. Kazakhstan: left bank middle course Ili R., 40 km NE Chilik, 20 km 

NE Barandysu; HT; ♂; ZMAS. 

oxytropeocarpi. Kazakhstan. Oxytropis pseudofrigida (Fabaceae). 

Contarinia oxytropeocarpi Fedotova 1993f: 675. Kazakhstan: foothills Saur Mtn Range, 20 km S L. Zaïsan, nr 

Zhanaturmys; HT; ♂; ZKAS. 

oxytropiflora. Kazakhstan. Oxytropis puberula; O. glabra (Fabaceae). 

Contarinia oxytropiflora Fedotova 1984e: 1033. Kazakhstan: Zailiïskiï Alatau, along Charyn R.; HT; ♂; ZKAS. 

partheniicola. USA (California, Arizona, New Mexico, Texas, Florida). Parthenium incanum; Ambrosia spp. 

(Asteraceae). 

Diplosis partheniicola Cockerell 1900a: 201. USA: New Mexico, Mesilla Valley, foot Picacho Mtn; LT (Gagné 

1975b: 53); ♂; NYSM. 

passlowi. Australia (Queensland). Dicanthium annulatum var. fecundum (Poaceae). 

Contarinia passlowi Harris 1979b: 178. Australia: Queensland, Townsville; HT; ♀; QMBA. 

pastinacae. UK, Germany, Czechoslovakia, Romania, Yugoslavia. Pastinaca sativa (Apiaceae). 

Diplosis pastinacae Rübsaamen 1892b: 395. Germany: vic. Berlin; ST; ♂,♀,L; ZMHB. 

Contarinia pastinaceae Skuhravá 1986: 243, missp. of pastinacae. 

pentaphylloidifolia. Kazakhstan. Pentaphylloides fruticosa (Rosaceae). 

Contarinia pentaphylloidifolia Fedotova 1990b: 107. Kazakhstan: southern Altaï, northeastern shore L. Markakol', 10 

km S Urunkhaïka; HT; ♂; ZKAS. 

perfoliata. USA (New York). Eupatorium perfoliatum (Asteraceae). 

Contarinia perfoliata Felt 1908b: 391. USA: New York, Albany; HT; ♂; NYSM. 

peritomatis. USA (New Mexico). Cleome serrulata (Capparidaceae). 

Cecidomyia peritomatis Cockerell 1913: 280. USA: New Mexico, Santa Fe; ST; LT; USNM. 

perplicata. Kazakhstan. Galium verum (Rubiaceae). 

Contarinia perplicata Fedotova 1997c: 326. Kazakhstan: Tarbagataï Mtn Range, 45 km W Ayaguz Mtn, Ayaguz 

valley; HT; ♂; ZMAS.

177 

petioli. Widespread Europe, Kazakhstan, Russia (Europe, Far East). Populus tremula (Salicaceae). 

Harmandia petioli Kieffer 1898f: 61. France: Moselle; T; ♀; lost?. 

Syndiplosis winnertzi Rübsaamen 1910d: 284. Germany; T; ♀; ZMHB. 

phellodendrobia. Russia (Far East). Phellodendron amurense (Rutaceae). 

Contarinia phellodendrobia Kovalev 1972: 421. Russia: Primorskiï Terr., Petra Velikoe, Popova I.; HT; ♂; ZMAS. 

picridis. Netherlands, France, Germany, Romania. Picris hieracioides (Asteraceae). 

Doxodiplosis picridis Kieffer 1894j: 9, fig. 8 (publ'n not seen); further descr. in Kieffer 1913a: 229. France: Moselle; 

ST; egg; lost?. 

pilosellae. Widespread Europe. Hieracium pilosella (Asteraceae). 

Contarinia pilosellae Kieffer 1896c: 100. France: Bitche; ST; ♂,♀; lost?. 

pimpinellae. Portugal. Pimpinella villosa (Apiaceae). 

Contarinia pimpinellae Tavares 1902b: 32. Portugal: Castello Branco, Louza, & Setubal; ST; ♂,♀,P; TAVC. 

piri. Spain. Crataegus monogyna (Rosaceae). 

Contarinia piri Tavares 1922c: 144. Spain: Rosal, Pontevedra; ST; ♂,♀,P,L; TAVC. 

pisi. Widespread Europe, Russia (Western Siberia). Pisum sativum; P. arvense (Fabaceae). 

Cecidomyia pisi Loew 1850: 30. Poland; ST; G; ZMHB?. 

Cecidomyia pisi Winnertz 1854: 325. Germany; ST; ♂,♀,L; MNHN?. 

Contarinia pisicola Meijere 1911: 185. Netherlands: Wageningen, Ulrum, & vic. Amsterdam; ST; ♂,♀,P,L; ZMAN. 

plicata. Kenya. Acacia drepanolobium (Fabaceae). 

Contarinia plicata Gagné 1993c: 116. Kenya: Rt. A109, Kapiti Plains, Malili Ranch, ca. 70 km SE Nairobi; HT; LT; 


plumosi. Australia (Queensland). Sorghum plumosum (Poaceae). 

Contarinia plumosi Harris 1979b: 172. Australia: Queensland, Mt Molloy; HT; ♀; QMBA. 

polygonati. UK, Sweden, Germany, Poland, Russia (Europe). Polygonatum multiflorum (Liliaceae). 

Contarinia polygonati Rübsaamen 1921: 43. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

populi. Widespread Europe. Populus tremula (Salicaceae). 

Syndiplosis populi Rübsaamen 1917: 81. Germany; ST; ♂,♀; ZMHB. 

pratula. Kazakhstan. Astragalus chlorodontus (Fabaceae). 

Contarinia pratula Fedotova 1993k: 96. Kazakhstan: Tarbagataï Mtn Range, 40 km NW Urdzhara, nr Blagodatnoye; 


pravdini. Turkmenistan, Kirghizstan. Lycium ruthenicum (Solanaceae). 

Contarinia pravdini Becknazarova & Mamaeva 1981: 82. Turkmenistan: Mary, Akibay; HT; ♂; HYNM. 

prolixa. Venezuela, Bolivia. 

Contarinia prolixa Gagné & Byers 1985: 736. Venezuela: Zulia; HT; ♀; USNM. 

prosopidis. India (Delhi). Prosopis spicigera (Fabaceae). 

Lobopteromyia prosopidis Mani 1938: 334. India: New Delhi; HT; ♀; INPC. 

pruniflorum. France, Czechoslovakia. Prunus spinosa, P. mahaleb (Rosaceae). 

Contarinia pruniflorum Coutin & Rambier 1955: 106. France: Normandy & Gard; HT; ♂; INRV. 

psammophila. Turkmenistan, Kazakhstan. Ammodendron argenteum (Chenopodiaceae). 

Contarinia psammophila Marikovskij 1975: 53. Kazakhstan: along middle course Ili R.; ST; ♂,♀,L; MARC. 

pseudotsugae. Canada (British Columbia), USA (Washington, Oregon, California, Montana; immmigr.: 

Pennsylvania). Pseudotsuga menziesii (Pinaceae). 

Contarinia pseudotsugae Condrashoff 1961: 124. Canada: British Columbia, Oyama; HT; ♂; CNCI. 

pulcherrima. India (West Bengal). Symplocos theaeformis (Styracaceae). 

Contarinia pulcherrima Kieffer 1909c: 130. India: West Bengal, Kurseong; ST; ♂,♀,L,P,egg; lost?. 

pulchripes. UK, Denmark, Netherlands, France, Czechoslovakia. Sarothamnus scoparius (Fabaceae). 

Contarinia pulchripes Kieffer 1890d: 133. France: vic. Bitche; ST; ♂,♀,L; lost?. 

pyrivora. Widespread Europe; immigr.: USA (New York, Massachusetts, Connecticut, Pennsylvania, Maryland). 

Pyrus communis (Rosaceae). 

Cecidomyia pyrivora Riley 1886: 287. USA: Connecticut, Meriden; ST; ♂,♀,P,L; USNM. 

quercicola. Czechoslovakia, Austria, Hungary, Ukraine, Romania, Yugoslavia, Greece. Quercus cerris (Fagaceae). 

Diplosis quercicola Rübsaamen 1899d: 599. Austria: Vienna; ST; ♀,L; ZMHB.

178 

quercina. Widespread Europe. Quercus robur (Fagaceae). 

Diplosis quercina Rübsaamen 1890b: 21. Germany: vic. Siegen; ST; ♂,♀,L; ZMHB. 

Diplosis dryophila Kieffer 1890e: 197. France: vic. Bitche; ST; ♂,♀,L; lost?. 

Contarinia quaesita Tavares 1916b: 106. Portugal: Castello Novo, Spain: Tuy, Pontevedra, & Santiago; ST; ♂,♀,L; 

TAVC. 

quinquenotata. Widespread Europe; immigr: Canada (BC); USA (Washington, Oregon); New Zealand. Hemerocallis 

fulva (Liliaceae). 

Diplosis quinquenotata Löw 1888: 235. Austria: Vienna & Seebenstein; ST; ♂,♀,L; NHMW. 

racemi. USA (Massachusetts, Maryland). Prunus virginiana (Rosaceae). 

Cecidomyia racemi Stebbins 1910: 39. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

ramachandrani. India (Tamil Nadu). Acacia ferruginea (Fabaceae). 

Lobopteromyia ramachandrani Mani 1953b: 262. India: Tamil Nadu, Coimbatore, Walayar For.; HT; ♂; ZSCI. 

ramicola. Germany. Tilia sp. (Tiliaceae). 

Cecidomyia ramicola Rudow 1875b: 239. Germany; ST; ♀,L; ?. 

reaumuriae. Kazakhstan. Reaumuria fruticosa (Tamaricaceae). 

Contarinia reaumuriae Fedotova 1991c: 51. Kazakhstan: eastern Mangyshlak basin, 18 km NE Senek, Besokty well; 


rhamni. UK, Norway, Germany, Poland, Czechoslovakia, Yugoslavia, Russia (Europe). Rhamnus frangula 

(Rhamnaceae). 

Contarinia rhamni Rübsaamen 1892a: 392. Germany: vic. Siegen; T; LT; ZMHB. 

rhodendorfi. Kazakhstan. Caragana arborescens (Fabaceae). 

Contarinia rhodendorfi Fedotova 1991c: 44. Kazakhstan: northern range Tarbagataï Mtn Range, 101 km E Ayaguz; 


ribis. Netherlands, France, Germany, Poland, Czechoslovakia. Ribes grossularia (Grossulariaceae). 

Contarinia ribis Kieffer 1909b: 25. France: Moselle, quarries betw. Sierck & Montenach; ST; LT; lost?. 

Contarinia ribis Meijere 1911: 181. Netherlands: Lent; ST; ♂,♀,P,L; ZMAN. 

roperi. Australia (Northern Territory). Sorghum plumosum (Poaceae). 

Contarinia roperi Harris 1979b: 171. Australia: Northern Territory, Roper R. Rd; HT; ♂; QMBA. 

rosaecarpae. Kazakhstan. Rosa beggeriana (Rosaceae). 

Contarinia rosaecarpae (also as rasaecarpae) Fedotova 1992a: 86. Kazakhstan: Karzhantau Mtn Range, floodplain 

Saïramsu R., 10 km SW Kaska-Su; HT; ♂; ZMAS. 

rubicola. UK, Sweden, Denmark, Netherlands, France, Germany, Czechoslovakia, Russia (Europe). Rubus sp.; R. 

caesius (Rosaceae). 

Contarinia rubicola Kieffer 1909b: 26. France; ST; LT; lost?. 

Contarinia rubicola Rübsaamen 1910f: 424. Germany: vic. Lieser an der Mosel; ST; ♂,♀,L; ZMHB. 

*rubsaameni. France. 

*Stictodiplosis rubsaameni Kieffer 1894k: unnumbered p. 1. Unstated, presumably France: Bitche; HT; egg 

(illustrated); lost? 

rumicina. Spain. Rumex acetosella (Polygonaceae). 

Atylodiplosis rumicina Tavares 1919b: 75. Spain: Gallicia, Pontevedra; ST; ♂,♀,P,L; TAVC. 

rumicis. UK, Portugal, France, Germany, Poland, Czechoslovakia, Austria, Romania, Yugoslavia; immigr.: USA (New 

York). Rumex sp.; R. crispus (Polygonaceae). 

Cecidomyia rumicis Loew 1850a: 34. Poland; ST; ♂,♀; ZMHB?. 

salatica. Kazakhstan. Lactuca tatarica (Asteraceae). 

Contarinia salaticola Fedotova 1996b: 203. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE Alma-Ata; 


salicola. Japan (Honshu). Salix reinii (Salicaceae). 

Contarinia salicola Shinji 1939f: 133. Japan: Honshu, Morioka; ST; ♂,♀,P,L; lost. 

sambuci. Widespread Europe. Sambucus nigra (Caprifoliaceae). 

Cecidomyia sambuci Kaltenbach 1873: 785. Germany; ST; LT; NHMW?. 

Diplosis lonicerearum Löw 1877: 17. Austria: Lower Austria; ST; ♂,♀,L; NHMW. 

sambucifoliae. USA (New York).

179 

Contarinia sambucifoliae Felt 1907a: 35. USA: New York, Lake Clear; HT; ♂; NYSM. 

Contarinia sambucifolia Felt 1907c: 131, unjust. emend. of sambucifoliae. 

saussureaflora. Kazakhstan. Saussurea elegans (Asteraceae). 

Contarinia saussureaflora Fedotova 1996b: 205. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE Alma- 

Ata; HT; ♂; ZMAS. 

scabiosae. UK, Sweden, France, Germany. Scabiosa columbaria (Dipsacaceae). 

Contarinia scabiosae Kieffer 1898f: 60. France: Aisne, vic. Coeuvres; ST; A; lost?. 

schlechtendaliana. UK, France, Italy, Germany, Poland, Russia (Europe). Sonchus arvensis (Asteraceae). 

Diplosis schlechtendaliana Rübsaamen 1893: 162. Germany: Sinzig; ST; ♂,♀,L; ZMHB. 

Contarinia sonchi Kieffer 1896a: 14. France: Moselle; T; A,L; lost?. 

schulzi. USA (North Dakota, Minnesota, Kansas, Texas). Helianthus annuus; Helianthus spp. (Asteraceae). 

Contarinia schulzi Gagné 1972c: 279. USA: North Dakota, Cass Co., Fargo; HT; ♂; USNM. 

scirpi. Australia (Northern Territory). Scirpus lacustris (Cyperaceae). 

Contarinia scirpi Harris 1979b: 180. Australia: Northern Territory, Elsey Creek; HT; ♀; QMBA. 

scoparii. Widespread Europe. Cytisus scoparius (Fabaceae). 

Diplosis scoparii Rübsaamen 1889a: 48. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

scrophulariae. Widespread Europe. Scrophularia nodosa (Scrophulariaceae). 

Contarinia scrophulariae Kieffer 1896c: 100. France: Bitche; ST; ♂,♀; lost?. 

scutati. Netherlands, Germany, Czechoslovakia, Romania. Rumex scutatus (Polygonaceae). 

Contarinia scutati Rübsaamen 1910f: 418. Germany: St. Goar; ST; ♂,♀,L; ZMHB. 

sehimae. Australia (Queensland). Sehima nervosum (Poaceae). 

Contarinia sehimae Harris 1979b: 173. Australia: Queensland, Normanton; HT; ♀; QMBA. 

selevini. Kazakhstan. Clematis soongarica (Rosaceae). 

Contarinia selevini Fedotova 1984f: 643. Kazakhstan: southeastern spur Zailiïskiï Alatau, Syugaty Mts, Kok-Pek 

pass; HT; ♂; ?. 

sennicola. Australia (New South Wales). Senna artemisioides (Fabaceae). 

Contarinia sennicola Kolesik 2000b: 122. Australia: New South Wales, Scotia Sanctuary; HT; ♂; SAMA. 

sesami. India (Madhya Pradesh). Sesamum orientale (Pedaliaceae). 

Contarinia sesami Grover & Bakhshi 1978: 94. India: Madhya Pradesh, Raipur; HT; ♂; PGCA. 

setigera. USA (Massachusetts). 

Diplosis setigera Lintner 1897: 168. USA: Massachusetts, Lowell; ST; ♂,♀; NYSM. 

shelahovi. Russia (Western Siberia). Trifolium sp.; Trifolium spp. (Fabaceae). 

Contarinia shelahovi Kolomoets 1986: 164. Russia: Sverdlovsk Prov., Beloyarskiy; HT; ♂; UASK. 

shevtshenkoi. Kazakhstan. Astragalus ammodendron; A. unifoliatus (Fabaceae). 

Contarinia shevtshenkoi Fedotova 1991c: 47. Kazakhstan: Mangyshlak Peninsula, nr Kuybyshevo, 80 km NS 

Shevchenko; HT; ♂; ZMAS. 

silenei. Spain. Lychnis dioica (Caryophyllaceae). 

Contarinia silenei Tavares 1916b: 112. Spain: Gallicia; ST; ♂,♀,L; TAVC. Skuhravá et al. 2006 consider this species 

synonymous with Contarinia steini. 

silvestris. France, Germany, Czechoslovakia, Hungary. Lathyrus sylvestris (Fabaceae). 

Contarinia silvestris Kieffer 1897a: 15. France; T; A; lost?. 

solani. Widespread Europe. Solanum dulcamara (Solanaceae). 

Diplosis solani Rübsaamen 1892b: 396. Germany: vic. Berlin; ST; ♂,♀,L; ZMHB. 

soongarica. Kazakhstan. Barbarea vulgaris (Brassicaceae). 

Contarinia soongarica Fedotova 1994b: 86. Kazakhstan: Dzhungarskiï Alatau, Altyn Emel' pass, 5 km E 

Golubinovka; HT; ♂; ZMAS. 

*sorbariaflora. Russia (Far East). Sorbaria sorbifolia (Rosaceae). 

Contarinia sorbariaflora Fedotova 2004e: 20. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

sorbi. UK, Sweden, Netherlands, France, Germany, Poland, Czechoslovakia, Yugoslavia, Ukraine. Sorbus aucuparia 

(Rosaceae). 

*Eudiplosis sorbi Kieffer 1894k: unnumbered p. 2. Unstated, presumably France: Bitche; HT; egg; lost?

180 

Contarinia sorbi Kieffer 1896c: 99. France: Bitche; ST; ♀,L; lost?. Redescription. 


Contarinia sphaerophysae Fedotova 1984f: 644. Kazakhstan: north shore L. Balkash, nr Zhiltyn Peninsula; HT; ♂; ?. 

spiraeaphaga. Kazakhstan. Spiraea hypericifolia (Rosaceae). 

Contarinia spiraeaphaga Fedotova 1988a: 60. Kazakhstan: Dzhungarskiï Alatau, Altyn-Emel' Pass; HT; ♂; ZKAS. 

spiraeina. USA (Maine, Massachusetts). Spiraea salicifolia (Rosaceae). 

Contarinia spiraeina Felt 1911l: 473. USA: Massachusetts, Magnolia; T; G; NYSM. 

squamulicola. USA (Massachusetts, Connecticut). Corylus americana (Betulaceae). 

Cecidomyia squamulicola Stebbins 1910: 16. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

stackelbergi, Russia (Far East). Sorbaria sorbifolia (Rosaceae). 

Contarinia stackelbergi Kovalev 1972: 418. Russia: Primorskiï Terr., Mt Oblachnaya; HT; ♂; ZMAS. 

steini. UK, Sweden, Denmark, Netherlands, France, Germany, Poland, Czechoslovakia, Hungary, Romania, 

Yugoslavia, Armenia. Lychnis dioica (Caryophyllaceae). 

Cecidomyia steini Karsch 1881: 227. Germany: vic. Berlin; ST; ♂,♀,L; ZMHB?. Skuhravá et al. 2006 consider this 

species synonymous with Contarinia silenei. 

subulifex. France, Czechoslovakia, Austria, Hungary, Romania, Yugoslavia. Quercus cerris (Fagaceae). 

Contarinia subulifex Kieffer 1897a: 15. Hungary: Budapest; T; ♂; lost?. 

symphyti. France. Symphytum officinale (Boraginaceae). 

Contarinia symphyti Kieffer 1909b: 30. France; T; G; lost?. 

tanaceti. Denmark, Germany, Czechoslovakia, Russia (Europe). Tanacetum vulgare (Asteraceae). 

Contarinia tanaceti Rübsaamen 1921: 47. Germany; ST; ♂,♀,L; ZMHB. 

tecomae. USA (New York). Campsis radicans (Bignoniaceae). 

Bremia tecomae Felt 1906b: 127. USA: New York, Albany; ST; ♂,♀,P,L; NYSM. 

tephrosiae. India (Delhi). Tephrosia candida (Fabaceae). 

Stictodiplosis tephrosiae Mani 1944: 157. India: Delhi; HT; ♀; INPC. 

texana. USA (Arizona, Oklahoma, Texas), Mexico. Orig. unk.; Cyamopsis tetragonoloba, Parkinsonia aculeata, P. 

macrophyllum (Fabaceae). 

Itonida texana Felt 1921a: 204. USA: Texas, Plano; HT; ♂; NYSM. 

thalactri. Canada (Ontario), USA (New York, Maryland). Fraxinus sp. (Oleaceae). 

Oligotrophus thalactri Felt 1907a: 27. USA: New York, Albany; HT; ♂; NYSM. 

Mayetiola thalictri Felt 1908b: 369, missp. of thalactri. 

thermopsidis. Kazakhstan. Thermopsis turkestanica (Fabaceae). 

Contarinia thermopsidis Fedotova 1993f: 678. Kazakhstan: Narynkol Valley, Tekes, 20 km NW Narynkol; HT; ♂; 

ZKAS. 

thlaspeos. Sweden, Germany. Thlaspi arvense (Brassicaceae). 

Contarinia thlaspeos Rübsaamen 1910f: 419. Germany: Laacher See; ST; ♂,♀,L; ZMHB. 

Contarinia major, as var. thlaspeos, Rübsaamen 1910f: 420. Germany: Laacher See; T; A; ZMHB. 

tianschanica. Kazakhstan. Rumex tianschanicus (Polygonaceae). 

Contarinia tianschanica Fedotova 1997g: 874. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Baldabrek Mtn 

Range, 57 km NE Georgievka; HT; ♂; ZMAS. 

tiliarum. Widespread Europe, Russia (Far East), Japan (Hokkaido, Honshu). Tilia grandifolia, T. parvifolia; T. 

japonica, T. maximowicziana (Tiliaceae). 

Diplosis tiliarum Kieffer 1890f: 193. France: Metz, Bitche, Bensdorf, & Rodalben; ST; ♂,♀; lost?. 

tragopogonis. UK, Sweden, France, Germany, Poland, Czechoslovakia, Romania, Yugoslavia. Tragopogon officinalis; 

T. dubius (Asteraceae). 

Contarinia tragopogonis Kieffer 1909b: 30. France; T; G; lost?. 

Contarinia tragopogonis Barnes 1927g: 217. UK: England, Kent; ST; ♂,♀; BMNH. 

tremulae. UK, Sweden, France, Poland, Czechoslovakia, Yugoslavia. Populus tremula (Salicaceae). 

Contarinia tremulae Kieffer 1909b: 18. France; ST; LT; lost?. 

trifolii. USA (New York). 

Contarinia trifolii Felt 1907a: 35. USA: New York, Albany; HT; ♂; NYSM. 

tritici. Widespread Europe, Israel. Triticum vulgare (Poaceae).

181 

Tipula tritici Kirby 1798: 232. UK: England; ST; A,P,L; ?. 

Contarinia venturii (also as bayeri) Vimmer 1936a: 28. Italy: Bologna; T; ♂; ?. 

trizni. Kazakhstan. Astragalus ugamicus (Fabaceae). 

Contarinia trizni Fedotova 1996e: 49. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Baldabrek Mtn Range, 

Chooldak area, 57 km NE Georgievka; HT; ♂; ZMAS. 

trotteri. France, Denmark, Yugoslavia. Carpinus betulus (Betulaceae). 

Contarinia trotteri Kieffer 1909b: 6. France; ST; LT; lost?. 

truncata. USA (New York). 

Contarinia truncata Felt 1908b: 393. USA: New York, Albany; ST; ♂; NYSM. 

turkmenica. Uzbekistan, Kirghizstan. Calligonum sp. (Polygonaceae). 

Contarinia turkmenica Becknazarova & Mamaeva 1981: 82. Uzbekistan: Bukhara Prov., Mt Kul'dzhuktau; HT; ♂; 

HYNM. 

ubiquita. Brazil (Rio de Janeiro). Mikania glomerata (Asteraceae). 

Contarinia ubiquita Gagné 2001: 120. Brazil: Rio de Janeiro, Poço das Antas Biol. Res.; HT; LT; QBUM. 

umbellatarum. UK, Germany. Pimpinella saxifraga (Apiaceae). 

Contarinia umbellatarum Rübsaamen 1910f: 422. Germany; ST; ♀,L; ZMHB. 

utechini. Kazakhstan. Astragalus ugamicus (Fabaceae). 

Contarinia utechini Fedotova 1996e: 51. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Baldabrek Mtn Range, 


valerianae. UK, Netherlands, Germany, Czechoslovakia. Valeriana officinalis (Valerianaceae). 

Diplosis valerianae Rübsaamen 1890c: 231. Germany: vic. Siegen; ST; ♂,♀,L; ZMHB. 

variabilis. Germany, Serbia. Rumex scutatus; R. crispus (Polygonaceae). 

Contarinia variabilis Rübsaamen 1917: 96. Germany: Werlau nr St. Goar; ST; ♂,♀,L; ZMHB. 

vera. Kazakhstan. Galium verum; G. pamiro-alaicum (Rubiaceae). 

Contarinia vera Fedotova 1997c: 328. Kazakhstan: southern Altaï, Markakol'skiï Res., L. Markakol', 60 km N 

Alekseyevka; HT; ♂; ZMAS. 

verna. UK. Lamium luteum (Lamiaceae). 

Cecidomyia verna Curtis 1827: item 178. UK: England; ST; ♂,♀,L; ?. Curtis's orig. illus. shows this to belong in 

Contarinia. 

vernalis. USA (New York). 

Oligotrophus vernalis Felt 1908b: 368. USA: New York, Karner; HT; ♀; NYSM. 

*veronicastum. Russia (Far East). on Veronicastrum sibiricum (Scrophulariaceae) 

Contarinia veronicastrum Fedotova 2003c: 22. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

verrucicola. USA (widespread eastern). Tilia americana (Tiliaceae). 

Cecidomyia verrucicola Osten Sacken 1875: 201. USA: New York, Niagara Falls & West Point, & Massachusetts, 

Cambridge; ST; G; USNM. 

viatica. USA (New York). 

Contarinia viatica Felt 1908b: 393. USA: New York, Albany; HT; ♂; NYSM. 

viburnorum. Widespread Europe. Viburnum lantana, V. opulus (Caprifoliaceae). 

Contarinia viburni Kieffer 1913a: 230, jun. prim. hom. viburni Felt. France: Bitche; ST; ♂,♀,L,egg; lost?. 

Contarinia viburnorum Kieffer 1913a: 230, new name for Contarinia viburni Kieffer. 

viciocarpi. Kazakhstan. Vicia tetrasperma (Fabaceae). 

Contarinia viciocarpi Fedotova 1993f: 682. Kazakhstan: NE spurs Dzhungarskiï Alatau, Kungeï Mts., 78 km SE 

Uch-Aral, 8 km SE Koktum; HT; ♂; ZKAS. 

vincetoxici. Sweden, Germany, Czechoslovakia. Vincetoxicum officinale (Asclepiadaceae). 

Contarinia vincetoxici Kieffer 1909b: 34. Germany: Kreuznach; T; LT; lost?. 

virginianiae. Widespread Nearctic. Prunus sp. (Rosaceae). 

Cecidomyia virginianiae Felt 1906b: 130. USA: New York, Keene Valley; HT; LT; NYSM. 

viridiflava. USA (New York). 

Contarinia viridiflava Felt 1908b: 392. USA: New York, Albany; HT; ♂; NYSM. 

virosa. Kazakhstan. Hieracium virosum (Asteraceae).

182 

Contarinia virosa Fedotova 1996b: 205. Kazakhstan: central Altaï, Kaksuïskiï Mts, Mt. Lymin Belok, 60 km NE 

Leninogorsk; HT; ♂; ZMAS. 

viticola. France, Italy, Germany, Czechoslovakia. Vitis vinifera (Vitaceae). 

Contaruia viticola Rübsaamen 1906b: 194. Germany; ST; ♂,P,L; ZMHB. 

washingtonensis. Canada (British Columbia), USA (Washington, Oregon). Pseudotsuga menziesii (Pinaceae). 

Contarinia washingtonensis Johnson 1963: 94. USA: Washington, nr Centralia; HT; ♂; USNM. 

zauschneriae. USA (California). Epilobium canum (Onagraceae). 

Thecodiplosis zauschneriae Felt 1912h: 152. USA: California, Whittier, Puente Hills; ST; ♂,♀,L; NYSM. 

ziziphorae. Kazakhstan. Zizyphora clinopodioides (Lamiaceae). 

Contarinia ziziphorae Fedotova 1998b: 13. Kazakhstan: southern Altaï, Markakol'skii Res., L. Markakol', nr 

Urunkhaïka; HT; ♂; ZMAS. 

zygophylli. Egypt. Zygophyllum album (Zygophyllaceae). 

Contarinia zygophylli Debski 1918: 23. Egypt: Hélouan, nr Walda; ST; LT; ?. 

zygophylliflorae. Armenia, Turkmenistan, Kazakhstan. Zygophyllum oxianum; Zygophyllum spp. (Zygophyllaceae). 

Contarinia zygophylliflorae Fedotova 1990c: 25. Kazakhstan: right bank Syr-Darya R., 30 km SW Turkestan, nr 

Baltakol'; HT; ♂; ZMAS. 

CONTODIPLOSIS 

Contodiplosis Gagné 1994a: 180. Type species, Compsodiplosis friburgensis Tavares (orig. des.). 

Neotropical; 3 spp., all on Styrax; Cecidomyiidi, unplaced to tribe. 

friburgensis. Brazil (Rio de Janeiro). Styrax sp. (Styracaceae). 

Compsodiplosis friburgensis Tavares 1915b: 150. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♂,♀; TAVC. 

humilis. Brazil (Rio de Janeiro). Styrax sp. (Styracaceae). 

Compsodiplosis humilis Tavares 1915b: 154. Brazil: Rio de Janeiro, Nova Friburgo; T; ♂; TAVC. 

tristis. Brazil (Rio de Janeiro). Styrax sp. (Styracaceae). 

Compsodiplosis tristis Tavares 1915b: 156. Brazil: Rio de Janeiro, Nova Friburgo; T; ♂; TAVC. 

COPINOLASIOPTERA 

Copinolasioptera Möhn 1975: 39. Type species, salvadorensis Möhn (orig. des.). 

Neotropical; 1 sp.; Lasiopteridi, Camptoneuromyiini. 

salvadorensis. El Salvador. Hymenaea courbaril (Fabaceae). 

Copinolasioptera salvadorensis Möhn 1975: 40. El Salvador: San Salvador; HT; ♂; SMNS. 

COQUILLETTOMYIA 

Coquillettomyia Felt 1908b: 398. Type species, Mycodiplosis lobata Felt (orig. des.). 

Almatamyia Marikovskij 1953a: 135. Type species, mirifica Marikovskij (orig. des.). 

Strobilodiplosis Möhn 1955b: 127. Type species, uvae Möhn (orig. des.). 

Pelodiplosis Möhn 1955b: 132. Type species, umida Möhn (orig. des.). 

Picrodiplosis Möhn 1955b: 137. Type species, caricis Möhn (orig. des.). 

Serratomyia Grover 1965: 25. Type species, mediospina Grover (mon.). 

Ancylodiplosis Gagné 1973c: 862. Type species, Coquillettomyia bryanti Felt (orig. des.). 

Calliperodiplosis Grover 1979c: 85. Type species, indica Felt (orig. des.). 

Halteromyia Grover 1979c: 122. Type species, quadrihaltera Grover (orig. des.). 

Coquillettomya Fedotova & Sidorenko 2006a: 134, missp. of Coquillettomyia. 

*Nescitodiplosis Fedotova & Sidorenko 2009c: 73, as subgenus of Coquillettomyia. Type species, Coquillettomyia

183 

bidens Mamaev (orig. des.). 

*Obviodiplosis Fedotova & Sidorenko 2009c: 73, as subgenus of Coquillettomyia. Type species, Karshomyia 

townsendi Felt (orig. des.). 

Holarctic, Neotropical, Oriental; 42 spp.; Cecidomyiidi, Mycodiplosini. Known larvae are associated with fungi and 

decaying vegetable matter. Adults of C. obliqua were taken in numbers from cacao flowers. Fedotova and Sidorenko 

2009c resurrect Pelodiplosis, Picrodiplosis, and Strobilodiplosis from synonymy under Coquillettomyia to subgeneric 

status and describe two new subgenera, Obviodiplosis and Nescitodiplosis. These subgenera are not separated here 

because they do not appear distinctive or otherwise serve the purpose of classification. 

*aprica. Russia (Far East). 

Coquillettomyia martia Fedotova & Sidorenko 2009c: 66. Russia: Primorskiï Terr., Vladivostok, Sputnik; HT; ♂; 

ZMAS. 

bidens. Russia (Far East). 

Coquillettomyia bidens Mamaev 1973a: 49. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. 

bidenticulata. China (Heilongjiang). 

Coquillettomyia bidenticulata Bu & Zheng 1994: 353. China: Heilongjiang, Shangzhi; HT; ♂; NKUM. 

bryanti. USA (New York, Massachusetts). 

Coquillettomyia bryanti Felt 1913f: 144. USA: Massachusetts, North Adams; ST; ♂,♀; NYSM. 

Lobodiplosis cincta Felt 1918e: 165. USA: New York, Newport; HT; ♀; NYSM. 

bulbiformis. China (Yunnan). 

Coquillettomyia bulbiformis Bu & Zheng 1994: 353. China: Yunnan, Wuding; HT; ♂; NKUM. 

*combinata. Russia (Far East). 

Coquillettomyia combinata Fedotova & Sidorenko 2005c: 229. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


dentata. USA (New York, Massachusetts), UK, Germany, Poland, Latvia, Lithuania, Ukraine, Russia (Europe, Far 

East), Kazakhstan, Kirghizstan. Carex spp. (Cyperaceae). Harris 2004a: 36 (synonymy). 

Coquillettomyia dentata Felt 1908b: 398. USA: New York, Newport; HT; ♂; NYSM. 

Picrodiplosis caricis Möhn 1955b: 137. Germany: Taunus, Dauborn; HT; ♂; SMNS. 

Almatamyia pegeli Marikovskij 1960: 183. Kazakhstan: vic. Alma-Ata, Kirghiztan: Frunze; ST; ♂; MARC. 

divergens. Russia (Europe). 

Coquillettomyia divergens Mamaev 1973a: 51. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; HYNM. 

elongata. Russia (Europe), China (Heilongjiang, Sichuan). 

Coquillettomyia elongata Bu & Zheng 1994: 354. China: Heilongjiang, Harbin; HT; ♂; NKUM. 

*exercita. Russia (Far East). 

Coquillettomyia exercita Fedotova & Sidorenko 2009c: 71. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 


extensa. Germany, Russia (Europe). 

Coquillettomyia extensa Mamaev 1973a: 50. Russia: Moscow Prov., Pavlovskaja Sloboda; HT; ♂; HYNM. 

fungivora. India (Kerala). Undet. spp. of Uredinales. 

Octodiplosis fungivora Nayar 1949: 89. India: Kerala, Trivandrum, Venganur; ST; ♂,♀; ZSCI. 

*hyppocrepica. Russia (Far East). 

Coquillettomyia hyppocrepica Fedotova & Sidorenko 2006a: 136. Russia: Primorskiï Terr., Vladivostok; HT; ♂; 

ZMAS. 

indiana. India (West Bengal). 

Calliperodiplosis indica Grover 1979c: 86, jun. sec. hom. indica Rao & Saksena. India: West Bengal, Darjeeling; HT; 

♂; PGCA. 

Coquillettomyia indiana Gagné 2004b: 122, new name for Coquillettomyia indica Grover. 


Dactylodiplosis indica Rao & Saksena 1959: 133. India: Maharashtra, Nagpur; HT; ♀; RCNU. 

inflata. India (Uttar Pradesh). 

Coquillettomyia inflata Kashyap 1986: 86. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA.

184 

*insita. Russia (Far East). 

Coquillettomyia insita Fedotova & Sidorenko 2009c: 61. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


insolita. Kazakhstan, Uzbekistan. 

Almatamyia insolita Marikovskij 1960: 184. Kazakhstan: vic. Alma-Ata; HT; ♂; MARC. 

kurilensis. Russia (Far East). 

Coquillettomyia kurilensis Mamaev 1998d: 7. Russia: Kuril Is, Kunashir; HT; ♂; ZMMU. 

labiata. India (Uttar Pradesh). 

Coquillettomyia labiata Grover 1979c: 82. India: Uttar Pradesh, India; HT; ♂; PGCA. 

lobata. USA (New York), UK, Netherlands, Germany, Lithuania, Russia (Europe, Far East). 

Mycodiplosis lobata Felt 1907a: 31. USA: New York, Lake Clear; HT; ♂; NYSM. 

longicurvatus. India (Jammu and Kashmir). 

Coquillettomyia longicurvatus Kashyap 1986: 88. India: Jammu and Kashmir, Srinagar; HT; ♂; PGCA. 

longipalpi. India (Uttar Pradesh). 

Microplecus longipalpi Rao 1953: 317. India: Uttar Pradesh, Agra; HT; ♂; ZSCI. 

lunata. India (Uttar Pradesh). 

Coquillettomyia lunata Grover 1979c: 82. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

*martia. Russia (Far East). 

Coquillettomyia martia Fedotova & Sidorenko 2009c: 64. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


mediospina. India (Uttar Pradesh). 

Serratomyia mediospina Grover 1965a: 26. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

mirifica. Germany, Latvia, Russia (Europe, Far East), Kazakhstan. 

Almatamyia mirificus Marikovskij 1953a: 135. Kazakhstan: Alma-Ata; ST; ♂,♀; ZKAS & ZMAS. 

nigricornis. UK, Russia (Europe). 

Coquillettomyia nigricornis Mamaev 1973a: 50. Russia: Kursk Prov.; HT; ♂; ZMMU. 

obliqua. Costa Rica. 

Coquillettomyia obliqua Gagné 1984c: 125. Costa Rica: Limón Prov., nr Siquirres, Finca La Lola; HT; ♂; USNM. 

photophila. India (Tripura). 

Coquillettomyia photophila Chandra 1993c: 11. India: Tripura, Agartala; HT; ♂; CSIA. 

*primorskii. Russia (Far East). 

Coquillettomyia primorskii Fedotova & Sidorenko 2005c: 229. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


quadrihaltera. India (West Bengal). 

Halteromyia quadrihaltera Grover 1979c: 123. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

quadrispina. India (Uttar Pradesh, Madhya Pradesh, West Bengal). 

Octodiplosis quadrispina Grover 1979c: 99. India: Madhya Pradesh, Indore; HT; ♂; PGCA. 

regionalis. Russia (Europe). 

Coquillettomyia regionalis Mamaev 1998d: 8. Russia: Moscow Prov., Lugovaya; HT; ♂; ZMMU. 

serratospina. India (Uttar Pradesh). 

Coquillettomyia serratospina Grover 1979c: 83. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

spinosa. Russia (Far East). 

Coquillettomyia spinosa Mamaev 1973a: 50. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. Ref.: 

Fedotova & Sidorenko 2009c: 58 (redescr.). 

texana. USA (Texas). 

Coquillettomyia texana Felt 1908b: 398. USA: Texas, HT; ♂; NYSM. 

townsendi. Peru. 

Karschomyia townsendi Felt 1912h: 155. Peru: Cajamarca, Jaén; ST; ♂,♀; NYSM. 

triradiata. India (Maharashtra). 

Coquillettomyia triradiata Chandra 1993c: 12. India: Maharashtra, Parbhani; HT; ♂; CSIA. 

truncata. China (Sichuan). 

Coquillettomyia truncata Bu & Zheng 1994: 355. China: Sichuan, Miyaluo; HT; ♂; NKUM.

umida. Germany, Latvia, Russia (Europe). 

Pelodiplosis umida Möhn 1955b: 133. Germany: Taunus, Dauborn; HT; ♂; SMNS. 

uvae. Germany. 

Strobilodiplosis uvae Möhn 1955b: 128. Germany: Hebertshausen, nr Munich; HT; ♂; SMNS. 

CORDIAMYIA 

Cordiamyia Maia 1996c: 579. Type species, globosa Maia (orig. des.). 


globosa. Brazil (Rio de Janeiro). Cordia verbenacea (Boraginaceae). 

Cordiamyia globosa Maia 1996c: 581. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

CORDYLODIPLOSIS 

Cordylodiplosis Gagné 1973c: 868. Type species, molliterga Gagné (orig. des.). 


molliterga. USA (Florida). 

Cordylodiplosis molliterga Gagné 1973c: 868. USA: Florida, Miami; HT; ♂; USNM. 

COTONEASTEROMYIA 

Cotoneasteromyia Fedotova 1991d: 19. Type species, Dasineura cotoneasteris Fedotova (orig. des.). 


cotoneasteris. Kazakhstan. Cotoneaster oligantha (Rosaceae). 

Dasineura cotoneasteris Fedotova 1990f: 74. Kazakhstan: Tarbagataï Mtn Range, upper Karabuga R., 80 km S 

Aksuat; HT; ♂; ZMAS. 

*COURIDIPLOSIS 

Couridiplosis Maia 2004a: 71. Type species, Couridiplosis vena Maia (mon.). 


*vena. Brazil (Minas Gerais). Croton floribundus (Euphorbiaceae). 

Couridiplosis vena Maia 2004a: 72. Brazil: Minas Gerais, Tiradentes, Serra de São José; HT; ♂; MNRJ. 

CRANEIOBIA 

Craneiobia Kieffer 1913g: 55. Type species, Cecidomyia corni Giraud (orig. des.). 

Holarctic; 2 spp.; Lasiopteridi, unplaced to tribe. 

corni. Widespread Europe. Cornus sanguinea (Cornaceae). 

Cecidomyia corni Giraud 1863: 1301. France; ST; ♂,♀; MNHN. 

tuba. USA (Illinois, New York, Massachusetts). Cornus amomum (Cornaceae). 

Cecidomyia tuba Stebbins 1910: 46. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

185

CTENODACTYLOMYIA 

Ctenodactylomyia Felt 1915a: 199. Type species, watsoni Felt (orig. des.). 


coccolobae. Cuba. Coccoloba uvifera (Polygonaceae). 

Cecidomyia coccolobae Cook 1909: 145. Cuba; T; G; lost. 

watsoni. USA (Florida). Coccoloba diversifolia; C. uvifera (Polygonaceae). 

Ctenodactylomyia watsoni Felt 1915a: 199. USA: Florida, Palm Beach; ST; ♂,♀,P,L; USNM. 

CTENODIPLOSIS 

Ctenodiplosis Kieffer 1913h: 109. Type species, elongatus Kieffer (orig. des.). 


elongata. Kenya. 

Ctenodiplosis elongatus Kieffer 1913h: 110. Kenya: Ramisi; ST; ♀,egg; MNHN. 

subrotunda. Tanzania. 

Ctenodiplosis subrotundus Kieffer 1913h: 110. Tanzania: Tavéta; HT; ♀; MNHN. 

CYSTIPHORA 

Cystiphora Kieffer 1892e: 212. Type species, Cecidomyia hieracii Loew (subs. des. Rübsaamen 1910: 337) = 

sanguinea Bremi. 

Laubertia Rübsaamen 1914: 94. Type species, schmidti Rübsaamen (mon.). 

Palearctic and introduced into Nearctic and Australasian Regions for biological control; 7 spp., all from leaf galls of 

western Palearctic Cichorieae (Asteraceae); Lasiopteridi, unplaced to tribe. Refs: Gagné 1975d (descr.), Harris 1975a 

(descr.), Peschken et al. 1993 (descr.). 

canadensis. Canada (Ontario), USA (Maine). Prenanthes altissima or P. alba; P. serpentaria (Asteraceae). 

Cystiphora canadensis Felt 1913h: 417. Canada: Ontario, Toronto; ST; ♀; NYSM. 

leontodontis. UK, France, Germany, Switzerland, Poland, Czechoslovakia, Austria. Leontodon hastilis; Taraxacum 

officinale (Asteraceae). 

Cecidomyia leontodontis Bremi 1847a: 19. Switzerland: vic. Zurich; ST; LT; lost. 

Cystiphora leontodontis Kieffer 1909b: 14. Austria; T; G; NHMW. 

sanguinea. Widespread Europe. Hieracium murorum; H. pilosella, H. praealtum (Asteraceae). 

Cecidomyia sanguinea Bremi 1847a: 19. Switzerland: vic. Zurich; ST; LT; lost. 

Cecidomyia hieracii Löw 1874a: 145. Austria: Vienna, Wienerwald; ST; ♀,P,L; NHMW. 

Cystiphora pilosellae Kieffer 1892f: 213. France: Bitche; ST; ♂,L; lost?. 

schmidti. Widespread Europe and Middle East; introd.: USA (Oregon, Idaho, California), Australia (New South 

Wales, Victoria). Chondrilla juncea (Asteraceae). 

Laubertia schmidti Rübsaamen 1914: 94. Poland: Zielona Gora; T; ♀; ZMHB. 

scorzonerae. France, Germany, Latvia, Hungary. Scorzonera humilis (Asteraceae). 

Cystiphora scorzonerae Kieffer 1909b: 27. France; T; G; lost?. 

sonchi. Widespread Europe; introd.: Canada (Alberta, Saskatchewan, Manitoba, Nova Scotia). Sonchus oleraceus; S. 

arvensis (Asteraceae). 

Cecidomyia sonchi Vallot 1827: 94. France; ST; LT; lost. 

Cecidomyia sonchi Bremi 1847a: 19. Switzerland: vic. Zurich; ST; LT; lost. 

186

Cecidomyia sonchi Löw 1875: 18. Austria: vic. Klosterneuberg and Weidling, nr Vienna; ST; ♂,♀,L; NHMW. 

taraxaci. Widespread Europe, Pakistan; introd.: Canada (Saskatchewan). Taraxacum officinale (Asteraceae). 

Cecidomyia taraxaci Kieffer 1888e: 98. France: Moselle; ST; ♂,♀,L; lost?. 

CYSTODIPLOSIS 

Cystodiplosis Kieffer & Jörgensen 1910: 395. Type species, longipennis Kieffer & Jörgensen (mon.). 


longipennis. Argentina (Mendoza). Grabowskia obtusa (Solanaceae). 

Cystodiplosis longipennis Kieffer & Jörgensen 1910: 395. Argentina: Mendoza, Chacras de Coria; ST; ♂,♀,P,L,egg; 

lost. 

*DACTYLASIOPTERA 

Dactylasioptera Kolesik & Veenstra-Quah 2008: 220. Type species, adentata Kolesik & Veenstra-Quah (orig. des.). 

Australasian; 2 spp. from leaf galls on Scleroderma (Chenopodiaceae); Lasiopteridi, Lasiopterini. 

*adentata. Australia (South Australia). Scleroderma diacantha (Chenopodiaceae). 

Dactylasioptera adentata Kolesik & Veenstra-Quah 2008: 221. Australia: South Australia, Eyre Peninsula, Minnipa; 


*dentata. Australia (South Australia). Scleroderma diacantha (Chenopodiaceae). 

Dactylasioptera dentata Kolesik & Veenstra-Quah 2008: 222. Australia: South Australia, Eyre Peninsula, Minnipa; 


DACTYLODIPLOSIS 

Dactylodiplosis Rübsaamen 1916a: 452. Type species, heisteriae Rübsaamen (mon.). 


*heptaphylli. Brazil (Rio de Janeiro). Protium heptaphyllum (Burseraceae). 

Dactylodiplosis heptaphylli Maia 2004b: 485. Brazil: Rio de Janeiro, Parque Nacional da Restinga de Jurubatiba, 

Lagoa de Carepebus; HT; ♂; MNRJ. 

heisteriae. Brazil (Amazonas). Heisteria cyanocarpa (Olacaceae). 

Dactylodiplosis heisteriae Rübsaamen 1916a: 454. Brazil: Amazonas, betw. Auristella & S. Francisco on R. Acre; 

ST; ♂,♀,P,L; ZMHB. 

icicaribae. Brazil (Rio de Janeiro). Protium icicariba (Burseraceae). 

Dactylodiplosis icicaribae Maia 2002b: 172. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

*DAEDALIDIPLOSIS 

Daedalidiplosis Fedotova 2004h: 176. Type species, galeidis Fedotova (orig. des.). 

Dacdalidiplosis Fedotova 2004i: 616, missp. of Daedalidiplosis. 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Aphidoletini. This genus is close to Bremia if the two must be separated. 

*galeidis. Russia (Far East). 

Daedalidiplosis galeidis Fedotova 2004h: 178. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

187

DAPHNEPHILA 

Daphnephila Kieffer 1905a: 144. Type species, haasi Kieffer (subs. des. Felt 1911c: 48). 

Eastern Palearctic and Oriental; 9 spp., all from galls of Lauraceae; Cecidomyiidi, Asphondyliini, Asphondyliina. 

Ref.: Tokuda, Yang & Yukawa 2008 (redescr.). 

glandifex. India (West Bengal). Machilus gamblei (Lauraceae). 

Daphnephila glandifex Kieffer 1905a: 148. India: West Bengal, vic. Kurseong; ST; ♀,P,L; lost?. 

haasi. India (West Bengal). Machilus gamblei (Lauraceae). 

Daphnephila haasi Kieffer 1905a: 144. India: West Bengal, vic. Kurseong; ST; ♂,♀,P,L,egg; lost?. 

Daphnephila hassi Mani 1953b: 258, missp. of haasi. 

linderae. India (West Bengal). Lindera pulcherrima (Lauraceae). 

Daphnephila linderae Kieffer 1905a: 149. India: West Bengal, vic. Kurseong; ST; ♂,P,L; lost?. 

machilicola. Japan (Honshu to Okinawa), Republic of Korea. Machilus thunbergii (Lauraceae). 

Daphnephila machilicola Yukawa 1974: 297. Japan: Kyushu, Kagoshima Pref., Mt Inao; HT; ♂; KUEC. 

*ornithocephala. Taiwan. Machilus thunbergii (Lauraceae). 

Daphnephila ornithocephala Tokuda, Yang & Yukawa 2008: 539. Taiwan: Taipei, Yangmingshan Nat'l Pk, 

Chungchengshan; HT; ♂; NMNS. 

*stenocalia. Taiwan. Machilus thunbergii (Lauraceae). 

Daphnephila stenocalia Tokuda, Yang & Yukawa 2008: 539. Taiwan: Taipei, Yangmingshan Nat'l Pk, 


*sueyenae. Taiwan. Machilus thunbergii (Lauraceae). 

Daphnephila sueyenae Tokuda, Yang & Yukawa 2008: 536. Taiwan: Taipei, Mt. Tatun; HT; ♂; NMNS. 

*taiwanensis. Taiwan. Machilus thunbergii (Lauraceae). 

Daphnephila taiwanensis Tokuda, Yang & Yukawa 2008: 540. Taiwan: Taipei, Yangmingshan Nat'l Pk, 


*truncicola. Taiwan. Machilus thunbergii (Lauraceae). 

Daphnephila truncicola Tokuda, Yang & Yukawa 2008: 540. Taiwan: Taipei, Yangmingshan Nat'l Pk, Erhtzupin; 

HT; ♂; NMNS. 

DASINEURA 

Dasineura Rondani 1840: 17. Type species, Tipula sisymbrii Schrank, subs. des. ICZN 1998: 246, Opinion 1911, 

setting aside previous des. of obscura Rondani by Rondani 1856: 200. 

Dasyneura Anonymous 1844: 450, missp. of Dasineura. 

Perrisia Rondani 1846: 371. Type species, Cecidomyia urticae Perris. 

Dasynevra Agassiz 1847: 117, unjust. emend. of Dasineura. 

Dasyneura Agassiz & Loew 1846: 11, poss. as emend. of Dasineura but preocc. Saunders 1842. 

Neocerata Coquillett 1900a: 47. Type species, rhodophaga Coquillett (mon.). 

Gioliella Del Guercio 1918a: 56. Type species, Dasyneura lathierei Del Guercio (mon.). 

Dasynewca Shinji 1938m: 335, missp. of Dasineura. 

Dassyneura Shinji 1944: 41, missp. of Dasineura. 

Gioriella Shinji 1944: 211, missp. of Gioliella. 

Cosmopolitan; 466 spp.; Lasiopteridi, Dasineurini. This broadly defined polyphyletic genus is a large dumping 

ground to which are assigned mostly flower-dwelling or leaf-rolling Lasiopteridi sharing certain primitive 

characteristics, but also many species that form galls, e.g., cedri Coutin and rhois Coquillett, some of which may not 

belong here. Refs: Sylvén 1975 (review of larvae), Sylvén & Tastás-Duque 1993 (inquiry). 

aberrata. USA (New York). 

188

189 

Dasyneura aberrata Felt 1908b: 346. USA: New York, Albany; HT; ♂; NYSM. 

abiesemia. USA (Oregon, California). Abies magnifica; A. concolor, A. grandis (Pinaceae). 

Dasyneura abiesemia Foote 1956: 52. USA: Oregon, Palmerlee's Ranch; HT; ♂; USNM. 

abietiperda. Widespread Europe. Picea abies (Pinaceae). 

Cecidomyia abietiperda Henschel 1880: 371. Austria; T; A; ?. 

Cecidomyia piceae Henschel 1881: 505. Austria: Vienna; ST; ♂,♀,egg; ?. 

Cecidomyia piceae Hartig 1893: 6. Germany: Ebersberg; ST; LT; ?. 

acaciaelongifoliae. Australia (Victoria, New South Wales, ACT, Queensland). Acacia longifolia, Acacia spp. 

(Fabaceae). Ref.: Kolesik 2005: 469 (redescr.) 

Cecidomyia acaciaelongifoliae (as acaciae-longifoliae) Skuse 1890: 374. Australia: New South Wales, Sydney; ST; 

♂,♀; ANIC. 

acerifoliae. USA (New York). 

Rhabdophaga acerifoliae Felt 1907a: 16. USA: New York, Albany; HT; ♂; NYSM. 

Dasyneura acerifolia Felt 1907c: 112, unjust. emend. of acerifoliae. 

aceris. Widespread eastern Nearctic. Acer saccharinum (Aceraceae). 

Cecidomyia aceris Shimer 1868: 281. USA: Illinois, Mount Carroll; ST; ♀,L; ?. 

acerobia. Russia (Far East). Acer mono (Aceraceae). 

Dasyneura acerobia Kovalev 1967: 114. Russia: Primorskiï Terr., Okeanskaya, nr Vladivostok; HT; ♂; KOVC. 

acrophila. Widespread Europe, Algeria. Fraxinus excelsior (Oleaceae). 

Cecidomyia acrophila Winnertz 1853: 233. Germany; ST; ♂,♀; MNHN?. 

acuminata. Germany, Czechoslovakia, Romania, Yugoslavia. Campanula rapunculoides; C. bononiensis 

(Campanulaceae). 

Dasyneura acuminata Rübsaamen 1916b: 511. Germany: vic. Jena; ST; ♂,L; ZMHB. 

adenophorae. Russia (Far East). Adenophora sp. (Campanulaceae). 

Dasyneura adenophorae Kovalev 1967: 116. Russia: Primorskiï Terr., Bogataya Griva, nr Vladivostok; HT; ♂; 

KOVC. 

affinis. Widespread Europe, Algeria, Turkey. Viola silvestris; V. odorata (Violaceae). 

Cecidomyia affinis Kieffer 1886: 330. France: vic. Bitche; ST; ♀,L; lost?. 

airae. France. Aira flexuosa (Poaceae). 

Perrisia airae Kieffer 1897b: 300. France: Moselle; ST; ♂,♀; lost?. 

alatavica. Kazakhstan. Lonicera kerelini (Caprifoliaceae). 

Dasineura alatavica Fedotova 1990f: 72. Kazakhstan: Terskeï Alatau, upper Bayankol R., 28 km SW Narynkol; HT; 

♂; ZMAS. 

albohirta. USA (New York). 

Dasyneura albohirta Felt 1908b: 346. USA: New York, Albany; HT; ♀; NYSM. 

albovittata. Widespread Nearctic. Salix spp. (Salicaceae). 

Cecidomyia albovittata Walsh 1864b: 621. USA: Illinois, nr Rock Island; ST; ♂,♀,P,L; USNM. 

alliicola. Kazakhstan. Allium galanthum (Liliaceae). 

Dasineura alliicola Fedotova 1993l: 81. Kazakhstan: Dzhungarskiï Alatau, Kungeï Alatau, 10 km SE Koktum; HT; 

♂; ZMAS. 

alopecuri. Widespread northern Europe; immigr: Canada (Ontario, New Brunswick), New Zealand (immigr.). 

Alopecurus pratensis; Agropyron repens (Poaceae). 

Oligotrophus alopecuri Reuter 1895: 3. Finland: Kirchspiel Pargas, nr Åbo; ST; ♂,♀,P,L; ?. 

Dasyneura agropyronis Barnes 1927f: 214. UK: Scotland, Aberdeenshire; HT; ♀; BMNH. 

alpestris. Widespread Europe. Arabis alpina, A. hirsuta; A. albida (Brassicaceae). 

Perrisia alpestris Kieffer 1909b: 2. Austria; T; G; NHMW?. 

Dasyneura schneideri Rübsaamen 1917: 48. Switzerland: Zurich; ST; ♂,♀,L; ZMHB. 

Dasyneura arabis Barnes 1927b: 54. UK: Surrey, Byfleet; ST; ♂,♀; ZMAN. 

altajensis. Kazakhstan. Galium boreale (Rubiaceae). 

Dasineura altajensis Fedotova 1997b: 43. Kazakhstan: Altaï, Narymskiï Mtn Range, 15 km S Bol'shenarymskoe; HT; 

♂; ZMAS. 

alyssi. Czechoslovakia, Hungary, Yugoslavia. Alyssum calycinum (Brassicaceae).

190 

Perrisia alyssi Kieffer 1901d: 168. Hungary: Budapest; T; ♀; lost?. 

amaramanjarae. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Dasyneura amaramanjarae Grover 1964c: 204. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

americana. USA (Massachusetts, North Carolina). Galium sp. (Rubiaceae). 

Dasyneura galii Felt 1908b: 348, jun. sec. hom. Cecidomyia galii Loew. USA: Massachusetts, Magnolia; ST; ♂,♀; 

NYSM. 

Dasyneura americana Felt 1913f: 136, new name for Dasyneura galii Felt. 

Perrisia galiorum Kieffer 1913i: 77, new name for Dasyneura galii Felt. 

ammodendronae. Kazakhstan. Astragalus ammodendron (Fabaceae). 

Dasineura ammodendronae Fedotova 1993j: 87. Kazakhstan: Bol'shiye Baruski Desert, 128 km NW Kulandy; HT; 

♂; ZMAS. 

andrieuxi. Portugal. Halimium libanotis (Cistaceae). 

Perrisia andrieuxi Tavares 1902a: 99. Portugal: vic. Setubal; ST; ♂,♀,L.; TAVC. 

anemone. USA (New York). Anemone canadense (Ranunculaceae). 

Dasyneura anemone Felt 1907d: 11. USA: New York, Kinderhook & Nassau; ST; ♂,♀; NYSM. 

angelicae. Widespread Europe. Angelica silvestris (Apiaceae). 

Dasyneura angelicae Rübsaamen 1916b: 512. Germany: Triglitz in Prignitz, Alt Rahlstedt nr Hamburg, & vic. 

Braunfels; ST; ♂,L; ZMHB. 

anglica. UK. Vaccinium vitis-idaea (Ericaceae). 

Perrisia anglica Kieffer 1909b: 31. UK: Scotland; ST; LT; lost?. 


Dasyneura antennata Felt 1908b: 347. USA: New York, Albany; HT; ♀; NYSM. 

aparines. Widespread Europe, Algeria. Galium aparine (Rubiaceae). 

Cecidomyia aparines Kieffer 1889d: 208. France: Bitche; ST; ♀,L; lost?. 

aparinicola. Kazakhstan. Galium aparine (Rubiaceae). 

Dasineura aparinicola Fedotova 1997b: 45. Kazakhstan: Trans-Ili Alatau, 32 km SE Chilik, 10 km NW Kok-Pek; 


apicata. USA (New York, Pennsylvania). Fraxinus sp. (Oleaceae). 

Dasyneura apicatus Felt 1908b: 345. USA: New York, Highland; ST; ♀; NYSM. 

armoraciae. Germany, Poland, Czechoslovakia, Romania, Yugoslavia. Armoracia rusticana (Brassicaceae). 

Dasyneura armoraciae (also as armoracia) Vimmer 1936b: 134. Czech Republic: Hrušov nr Opava; T; ♂; ?. 

aromaticae. USA (New York, Massachusetts). Mentha sp. (Lamiaceae). 

Dasyneura aromaticae Felt 1909b: 289. USA: Massachusetts, Barre; ST; ♂; NYSM. 

artemisiae. UK, Germany, Latvia. Artemisia campestris (Asteraceae). 

Dasyneura artemisiae Rübsaamen 1916b: 516. Germany: vic. Bergedorf nr Hamburg; ST; ♂,♀,L; ZMHB. 

asparagi. Portugal, Spain, Malta. Asparagus aphyllus (Liliaceae). 

Perrisia asparagi Tavares 1902b: 25. Portugal: Setubal; ST; ♂,♀,L; TAVC. 

asparagiiflora. Kazakhstan. Asparagus persicus (Liliaceae). 

Dasineura asparagiiflora Fedotova 1993i: 49. Kazakhstan: Charyn R., 21 km N Chundzha; HT; ♂; ZMAS. 

asperulae. Widespread Europe. Asperula tinctoria; A. cinanchica, A. ciliata (Rubiaceae). 

Cecidomyia asperulae Löw 1875: 13. Austria: vic. Klosterneuberg and Weidling, nr Vienna; ST; ♂,♀,L; NHMW. 

Perrisia asperularum Kieffer 1909b: 2. France; T; LT; lost?. 

asteriae. Japan (Honshu), Republic of Korea. Aster scaber (Asteraceae). 

Fulvomyia asteriae Shinji 1938h: 1066, nomen nudum. 

Mikiola asteriae Shinji 1942b: 2. Japan: Honshu, Tokyo; T; ♂; lost. 

astragalorum. France, Poland, Czechoslovakia, Romania. Astragalus arenarius, A. glyciphyllos (Fabaceae). 

Perrisia astragalorum Kieffer 1909b: 2. France; T; G; lost?. 

attenuata. USA (New York). 

Dasyneura attenuata Felt 1908b: 350. USA: New York, Albany; HT; ♂; NYSM. 

aucupariae. France. Sorbus aucuparia (Rosaceae). 

Perrisia aucupariae Kieffer 1909b: 29. France; T; G; lost?. 

augusta. USA (New York).

191 

Dasyneura augusta Felt 1908b: 348. USA: New York, Albany; HT; ♀; NYSM. 

auricomi. UK, France, Germany. Ranunculus auricomus (Ranunculaceae). 

Perrisia auricomi Kieffer 1909b: 25. Austria: nr Ohrdruf; T; G; NHMW? 

aurihirta. USA (New York). 

Dasyneura aurihirta Felt 1908b: 342. USA: New York, Albany; HT; ♀; NYSM. 

auritae. Widespread Europe. Salix aurita; S. cinerea (Salicaceae). 

Dasyneura auritae Rübsaamen 1916b: 506. Germany: nr Alt-Rahlstedt nr Hamburg; ST; ♀; ZMHB. 

axillaris. Widespread Europe. Trifolium medium (Fabaceae). 

Dasyneura axillaris Kieffer 1896c: 87. France: château de La Grange, nr Thionville; ST; ♀,L; lost?. 

azutavica. Kazakhstan. Erigeron acris (Asteraceae). 

Dasineura azutavica Fedotova 1996a: 538. Kazakhstan: southern Altaï, 45 km SW Katon-Karagaï, nr Ornek; HT; ♂; 

ZMAS. 

balsamicola. Canada (Nova Scotia, New Brunswick), USA (California?, Wisconsin, Michigan, New York, North 

Carolina). Abies balsamea; A. fraseri, A. concolor? (Pinaceae). 

Cecidomyia balsamicola Lintner 1888a: 60. USA: New York, Hamilton Co., Sageville; ST; LT; lost. 

balsamifera. USA (New York). 

Mayetiola balsamifera Felt 1908b: 369. USA: New York, Lake Clear; HT; ♀; NYSM. 

banksiae. Australia (South Australia, Western Australia). Banksia cocciniea (Proteaceae). 

Dasineura banksiae Kolesik 2007a: 41. Australia: Western Australia, Redmond West, nr Albany; HT; ♂; SAMA. 

bayeri. Lithuania, Poland, Czechoslovakia, Hungary, Romania. Sisymbrium leoselii (Brassicaceae). 

Dasyneura bayeri Rübsaamen 1914: 104. Czech Republic: Brno; ST; ♂,♀; ZMHB. 

beccariella. Eritrea. Momordica pteromorfa (Cucurbitaceae). 

Perrisia beccariella Del Guercio 1918a: 153. Eritrea; T; LT; lost. 

berberidis. Widespread Europe. Berberis vulgaris (Berberidaceae). 

Perrisia berberidis Kieffer 1909b: 3. Austria; T; G; NHMW?. 

berestae. Kazakhstan. Geranium albiflorum; G. collinum (Geraniaceae). 

Dasineura berestae Fedotova 1993o: 19. Kazakhstan: Dzhungarskiï Alatau, Kargaly R. valley, 10 km W Aral-Tyube; 


berteroae. Germany. Barbarea vulgaris (Brassicaceae). 

Dasineura berteroae Stelter 1976: 89. Germany: Warin; HT; ♂; SMTD. 

berti. Norway. Astragalus alpinus (Fabaceae). 

Dasineura berti Sylvén 1993: 294. Norway: Kongsvold; HT; ♀; NHRS. 

betuleti. France. Betula alba (Betulaceae). 

Cecidomyia betuleti Kieffer 1886: 326. France: vic. Bitche; ST; ♀,L; lost?. 

bidentata. USA (New York). 

Dasyneura bidentata Felt 1907a: 18. USA: New York, Albany; HT; ♂; NYSM. 

bistortae. Widespread Europe, Kazakhstan. Polygonum bistortum; P. alpinum (Polygonaceae). 

Perrisia bistortae Kieffer 1909b: 18. Austria; T; G; NHMW?. 

Dasyneura polygoni Rübsaamen 1921: 48. France: Haut-Rhin, Hohneck, Longemer; ST; ♀,L; ZMHB. 


Dasyneura borealis Felt 1907a: 21. USA: New York, Lake Clear; HT; ♀; NYSM. 

bragancae. Portugal. Thalictrum glaucum (Rhamnaceae). 

Perrisia bragancae Tavares 1904b: 298. Portugal: Matta do Fundão; ST; ♂,♀,P; TAVC. 

braziliensis. Brazil (Bahia). Protium heptaphyllum (Burseraceae). 

Perrisia braziliensis Tavares 1922a: 12. Brazil: Bahia, vic. Bahia; ST; ♀,L; TAVC. 

broteri. Portugal, Spain. Erica ciliaris (Ericaceae). 

Perrisia broteri Tavares 1901: 58. Portugal: betw. Runa & Turcifal, & Perto de la Cruz; ST; ♂,♀,P,L; TAVC. 

brunellae. Spain, Italy. Prunella vulgaris (Lamiaceae). 

Perrisia brunellae Kieffer 1909b: 3. Italy; ST; LT; lost?. 

Dasineura brunnelae Skuhravá 1986: 147, missp. of brunellae. 

bugaevi. Kazakhstan. Artemisia absinthium (Asteraceae). 

Dasineura bugaevi Fedotova 2001h: 1483. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak Canyon, 20 km SE Alma-Ata;

192 


bupleuri. Germany, Czechoslovakia, Austria, Hungary. Bupleurum falcatum (Apiaceae). 

Cecidomyia bupleuri Wachtl 1883: 478. Austria: vic. Vienna; ST; ♂,♀,L; NHMW?. 

bursakovi. Kazakhstan. Saussurea elegans (Asteraceae). 

Dasineura bursakovi Fedotova 1996b: 195. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE Alma-Ata; 


bursifex. France. Cytisus biflorus (Fabaceae). 

Perrisia bursifex Kieffer 1909b: 7. France; T; G; lost?. 

californica. USA (California). Salix sp. (Salicaceae). 

Dasyneura californica Felt 1908b: 347. USA: California, Alameda; HT; ♀; USNM. 

callitridis. Australia (South Australia). Callitris gracilis (Cupressaceae). 

Dasineura callitridis Kolesik 2000c: 252. Australia: South Australia, Sandy Creek Conservation Pk; HT; ♂; SAMA. 

calophacifoliae. Kazakhstan. Calophaca soongorica (Fabaceae). 

Dasineura calophacifoliae Fedotova 1989e: 34. Kazakhstan: Tarbagataï Mtn Range, 40 km NW Urdzhar, nr 

Blagodatnoye; HT; ♂; ZKAS. 

campanulae. UK, Germany, Czechoslovakia. Campanula rotundifolia (Campanulaceae). 

Dasyneura campanulae Rübsaamen 1914: 102. Germany: St. Goar; ST; ♂,♀; ZMHB. 

campanularum. France. Campanula glomerata (Campanulaceae). 

Perrisia campanularum Kieffer 1909b: 4. France; ST; LT; lost?. 

capsulae. Widespread Europe, Algeria, Tunisia. Euphorbia cyparissias (Euphorbiaceae). 

Dasyneura capsulae Kieffer 1901d: 167. France: Bitche; ST; ♂,♀; lost?. 

Perrisia cornifex Kieffer 1909b: 7. Algeria and Spain; T; G; lost?. 

carbonaria. USA (New York, Pennsylvania). Euthamia graminifolia (Asteraceae). 

Dasyneura carbonaria Felt 1907a: 21. USA: New York, Albany; HT; ♀; NYSM. Ref.: Dorchin et al. 2007 (tax'y, 

biol.) 

Dasyneura flavicornis Felt 1908b: 345. USA: New York, Albany; ST; ♀; NYSM. 

cardaminicola. UK, Germany. Cardamine amara (Brassicaceae). 

Dasyneura cardaminicola Rübsaamen 1916b: 512. Germany: Stützerbach and Gehlberg in the Thuringian For.; ST; 

♂,♀,L; ZMHB. 

cardaminis. Widespread Europe. Cardamine pratense; C. flexuosa (Brassicaceae). 

Cecidomyia cardaminis Winnertz 1853: 225. Germany; ST; ♂,♀,L; MNHN. 

cardariae. Kazakhstan. Cardaria draba (Brassicaceae). 

Dasineura cardariae Fedotova 1994a: 121. Kazakhstan: Syr Darya R. valley, 16 km N Bairkum; HT; ♂; ZMAS. 

caricicola. USA (New York). 

Dasyneura caricis Felt 1907a: 19, jun. prim. hom. caricis Kieffer. USA: New York, Albany; HT; ♂; NYSM. 

Perrisia caricicola Kieffer 1913i: 74, new name for Dasineura caricis Felt. 

caricis. France. Carex spp. (Cyperaceae). 

Dasyneura caricis Kieffer 1901c: 20. France: Bitche; T; ♀; lost?. 

Perrisia caricina Kieffer 1913i: 74, unnec. new name for Dasyneura caricis Kieffer. 

carpesii. France. Carpesium cernuum (Asteraceae). 

Perrisia carpesii Kieffer 1909b: 6. France; T; G; lost?. 

cecconiana. Italy, Czechoslovakia. Campanula trachelium (Campanulaceae). 

Perrisia cecconiana Kieffer 1909b: 4. Italy; ST; LT; lost?. 

cedri. Lebanon. Cedrus libani (Pinaceae). 

Dasineura cedri Coutin 2001a: 487. Lebanon: Tannourine; HT; ♂; MNHN. 

centaureae. Czechoslovakia, Austria. Centaurea montana, C. sp. (Asteraceae). 

Perrisia centaureae Kieffer 1909b: 6 (bis). France, Austria: Bernese Alps; T; G; NHMW?. 

cerastii. UK, Portugal, France, Czechoslovakia, Yugoslavia. Cerastium viscosum; Cerastium sp. (Caryophyllaceae). 

Cecidomyia cerastii Binnie 1877: 181. UK: Scotland, Possil Distr.; ST; ♂,L; ?. 

cerastiiflora. Kazakhstan. Cerastium ceratoides (Caryophyllaceae). 

Dasineura cerastiiflora Fedotova 1993i: 51. Kazakhstan: Tarbagataï Mtn Range, 80 km S Aksuat; HT; ♂; ZMAS. 

cercocarpi. (USA (Oregon, California, Colorado). Cercocarpus betuloides (Rosaceae).

193 

Dasyneura cercocarpi Felt 1913d: 215. USA: Colorado, Golden; ST; ♂,♀,L; NYSM. 

Dasineura cerocarpi Foote 1965: 259, missp. of cercocarpi. 

chilensis. Chile. Baccharis rosmarinifolia (Asteraceae). 

Perrisia chilensis Kieffer & Herbst 1909: 121. Chile: Concepción; ST; ♂,♀,P; lost. 

chrysanthemi. UK, Denmark. Chrysanthemum leucanthemum (Asteraceae). 

Dasyneura chrysanthemi Heath 1962: 8. UK: England, Cornwall, Devon, Derbyshire, & Jersey; ST; ♂,♀; BMNH. 

chrysophyllae. Eritrea. Olea chrysophylla (Oleaceae). 

Perrisia chrysophyllae Del Guercio 1918a: 49. Eritrea; T; LT; lost. 

cirsioni. USA (New York). 

Dasyneura cirsioni Felt 1908b: 346. USA: New York, Albany; HT; ♀; NYSM. 

citri. India (Uttar Pradesh, Maharashtra). Citrus sp. (Rutaceae). 

Pectinodiplosis citri Rao & Grover 1960: 745. India: Maharashtra, Nagpur; HT; ♂; RCMU. 

Dasyneura citri Grover & Prasad 1966b: 25. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

citrigemmia. China (Guangdong). Citrus sp. (Rutaceae). 

Dasineura citrigemmia (also as citrigemmina) Yang & Tang 1991: 74. China: Guangdong, Guangzhou City, Shipai; 

HT; ♂; BAUC. 

citrigemina. China (Guangdong). Citrus sp. (Rutaceae). 

Dasineura citrigemmia Yang & Tang 1991: 76. China: Guangdong, Guangzhou City, Shipai; HT; ♂; SCUC. 

clematidis. USA (New York). Clematis sp. (Rosaceae). 

Dasyneura clematidis Felt 1908b: 344. USA: New York, Highland; ST; ♂,♀; NYSM. 

clematidina. Austria, Armenia. Clematis viticella; C. vitalba (Rosaceae). 

Perrisia clematidis Kieffer 1909b: 6, jun. sec. hom. clematidis Felt. Austria; T; G; NHMW?. 

Perrisia clematidina Kieffer 1913i: 75, new name for Perrisia clematidis Kieffer. 

clematifolia. Kazakhstan. Clematis orientalis (Rosaceae). 

Dasineura clematifolia Fedotova 1985f: 35. Kazakhstan: north bank Ili R., nr Malaïsary; HT; ♂; ZKAS. 

coffeae. Congo. Coffea sp. (Rubiaceae). 

Dasyneura coffeae Barnes 1939f: 325. Democratic Republic of Congo: Rutshuru; ST; ♂,♀; BMNH. 

collinsoniae. USA (New York, Maryland). Collinsonia canadensis (Lamiaceae). 

Cecidomyia collinsoniae Beutenmüller 1908: 73. USA: New York, Garrison; HT; LT; AMNH. 

columnae. Austria, Hungary, Romania. Ononis columnae (Fabaceae). 

Perrisia columnae Kieffer 1909b: 15. Austria; ST; LT; NHMW?. 

communis. Widespread eastern Nearctic. Acer sp.; Acer spp. (Aceraceae). 

Dasyneura communis Felt 1911m: 478. USA: New York, Albany; ST; ♂,♀; NYSM. 

comosae. Germany, Czechoslovakia, Italy. Hippocrepis comosa (Fabaceae). 

Dasyneura comosae Rübsaamen 1916b: 515. Germany: Hausberg nr Jena; ST; ♂,♀,L; ZMHB. 


Dasyneura consobrina Felt 1907a: 21. USA: New York, Albany; HT; ♀; NYSM. 

copacabanensis. Brazil (Rio de Janeiro). Eugenia copacabanensis (Myrtaceae). 

Dasineura copacabanensis Maia 1993: 717. Brazil: Rio de Janeiro, Arraial do Cabo; HT; ♂; QBUM. 

copiosa. Kazakhstan. Aster altaicus (Asteraceae). 

Dasineura copiosa Fedotova 1996a: 544. Kazakhstan: Tarbagataï Mtn Range, Kel'dymurat R. valley; HT; ♂; ZMAS. 

corniculata. Germany, Czechoslovakia. Lamium album (Lamiaceae). 

Perrisia corniculata Kieffer 1909b: 12. Germany; T; G; ZMHB?. 

corollae. Guadeloupe, Trinidad. Chromolaena odorata; Eupatorium sp. (Asteraceae). 

Dasineura corollae Gagné 1977b: 118. Trinidad: Curepe; HT; ♂; USNM. 

coronillae. Portugal. Coronilla glauca (Fabaceae). 

Perrisia coronillae Tavares 1901: 56. Portugal: Puchaleiros Valley; ST; ♂,♀,L; TAVC. 

corticis. USA (Massachusetts). Salix sp. (Salicaceae). 

Dasyneura corticis Felt 1909b: 289. USA: Massachusetts, Boston, Arnold Arboretum; ST; ♂,♀; NYSM. 

corylina. UK, Netherlands, Sweden, Germany. Corylus avellana (Betulaceae). 

Dasyneura coryli Rübsaamen 1912e: 288, jun. prim. hom. coryli Felt. Germany; ST; ♂,♀,L; ZMHB. 

Perrisia corylina Kieffer 1913i: 75, new name for Dasyneura coryli Rübsaamen.

194 

cotini. Slovenia. Cotinus coggygria (Anacardiaceae). 

Dasyneura cotini Jane�i 1978: 9. Slovenia: betw. Koper & Portoro�; ST; ♂,♀,L; ?. 

couepiae. Brazil (Rio de Janeiro). Couepia ovalifolia (Chrysobalanaceae). 

Dasineura couepiae Maia 2001a: 13. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

crataegi. Widespread Europe. Crataegus oxyacantha; C. coccinea, C. monogyna (Rosaceae). 

Cecidomyia crataegi Winnertz 1853: 228. Germany; ST; ♂,♀,L; lost. 

crataegibedeguar. Canada (Ontario), USA (Illinois, Massachusetts, Virginia). Crataegus crus-galli; Crataegus spp. 

(Rosaceae). 

Cecidomyia crataegi bedeguar Walsh 1869: 79, a polynomial, invalid. 

Cecidomyia crataegibedeguar (as crataegi-bedeguar) Osten Sacken 1878: 6. USA: Illinois, Rock Island; ST; G; 


cyanococci. USA (Massachusetts, New Jersey, Georgia). Vaccinium sp. (Ericaceae). 

Dasyneura cyanococci Felt 1907d: 11. USA: Massachusetts, Stowe; HT; ♀; NYSM. 

cytisi. France, Romania, Ukraine, Russia (Western Siberia). Cytisus biflorus (Fabaceae). 

Perrisia cytisi Kieffer 1909b: 7. France; ST; LT; lost?. 

dactylidis. UK, Czechoslovakia, Russia (Europe). Dactylis glomerata (Poaceae). 

Dasyneura dactylidis Metcalfe 1933: 329. UK: Herts, Rothhamsted & Harpenden; ST; ♂,♀; BMNH. 

daphnephila. Austria, Romania, Yugoslavia. Daphne striata (Thymelaeaceae). 

Perrisia daphnephila Kieffer 1909b: 7. Austria: Graubünden; T; G; NHMW?. 

daphnes. Widespread Europe. Daphne cneorum (Thymelaeaceae). 

Perrisia daphnes Kieffer 1901c: 18. France: vic. Main-du-Prince, nr Bitche; T; ♂; lost?. 

dentatae. Canada (Ontario), USA (Ohio, Massachusetts). Castanea dentata (Fagaceae). 

Eriophyes dentatae Stebbins 1910: 16. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

denticulata. USA (New York). 

Dasyneura denticulata Felt 1907a: 21. USA: New York, Lake Clear; HT; ♀; NYSM. 

dianthi France. Dianthus carthusianorum (Caryophyllaceae). 

Perrisia dianthi Kieffer 1909b: 7. France; T; G; lost?. 

dielsi. Australia (Western Australia, Victoria), South Africa (introd.). Acacia cyclops, Acacia spp. (Fabaceae). Ref.: 

Kolesik 2005: 471 (redescr.) 

Dasyneura dielsi Rübsaamen 1916a: 478. Australia: Western Australia, nr Karocketta, & Victoria; ST; ♂,♀; ZMHB. 

dioicae. UK, Netherlands, Germany, Czechoslovakia, Hungary, Romania. Urtica dioica (Urticaceae). 

Dichelomyia dioicae Rübsaamen 1895b: 179. Germany: Tegeler See; ST; A,L; ZMHB. 

dombrovskae. Kazakhstan. Lonicera tatarica (Caprifoliaceae). 

Dasineura dombrovskae Fedotova 1985f: 34. Kazakhstan: Kirgizskiï Alatau, S Merke; HT; ♂; ZKAS. 

dryophila. UK, Germany, Russia (Europe). Quercus robur (Fagaceae). 

Dasyneura dryophila Rübsaamen 1917: 47. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

dzhanokmenae. Kazakhstan. Descurainia sophia (Brassicaceae). 

Dasineura dzhanokmenae Fedotova 1994a: 119. Kazakhstan: Karakumy Desert, Kul'zhan, 22 km SW Opornaya Stn; 


elatostemmae. Indonesia (Java). Elatostema sp. (Urticaceae). 

Dasyneura elatostemmae Felt 1921h: 140. Indonesia: Java, Ungaran Mts; ST; ♂,♀; NYSM. 

elegans. Portugal. Erica umbellata; E. australis (Ericaceae). 

Perrisia elegans Tavares 1907a: 51. Portugal: Gerez, nr Braga; ST; ♂,♀; TAVC. 

engstfeldi. Widespread Europe. Spiraea ulmaria (Rosaceae). 

Cecidomyia engstfeldi Rübsaamen 1889b: 375. Germany: vic. Siegen; ST; ♂,L; ZMHB. Ref.: Harris 2010a (gall). 

Perrisia rostrupiana Kieffer 1909b: 29. France; ST; LT; lost?. 

epilobii. Widespread Europe. Epilobium angustifolium (Onagraceae). 

Cecidomyia epilobii Löw 1889a: 201. Germany: vic. Aachen, & Austria: vic. Vienna; ST; ♂,♀,L; NHMW. 

ericaescopariae. Portugal, France, Italy, Yugoslavia, Algeria, Tunisia. Erica scoparia (Ericaceae). 

Cecidomyia ericaescopariae (as ericae scopariae) Dufour 1837: 87. France: Bordeaux; ST; ♂,♀; ?. 

erigerontis. Germany. Erigeron acris (Asteraceae). 

Dasyneura erigerontis Rübsaamen 1912e: 286. Germany: Rhine Prov.; ST; ♂,♀,L; ZMHB.

195 

erodiicola. Portugal, Malta, Tunisia. Erodium moschatum (Geraniaceae). 

Dasineura erodiicola Sylvén 1993: 296. Portugal: Algarve, Albufeira; HT; ♀; NHRS. 

erysimicola. Kazakhstan. Erysimum diffusum (Brassicaceae). 

Dasineura erysimicola Fedotova 1994a: 114. Kazakhstan: Karatau Mtn Range, 8 km N Kentau; HT; ♂; ZMAS. 

eugeniae. USA (Florida), Puerto Rico. Eugenia buxifolia (Myrtaceae). 

Dasyneura eugeniae Felt 1912f: 106. USA: Florida, Key West; ST; ♂,♀; USNM. 

euphorbiarum. France. Euphorbia cyparissias (Euphorbiaceae). 

Perrisia euphorbiarum Kieffer 1909b: 7. France; T; G; lost?. 

excavans. UK, Germany, Czechoslovakia, Switzerland, Austria, Italy. Lonicera xylosteum (Caprifoliaceae). 

Perrisia excavans Kieffer 1909b: 14. Switzerland: nr Neuchatel & Rhonethal; T; G; NHMW. 

ezomatsue. Japan (Hokkaido, Honshu). Picea jezoensis (Pinaceae). 

Dasyneura ezomatsue Uchida & Inouye 1954: 73. Japan: Hokkaido; HT; ♂; HFES. 

fairmairei. UK, France, Czechoslovakia, Romania, Russia (Western Siberia). Lathyrus sylvestris (Fabaceae). 

Perrisia fairmairei Kieffer 1897b: 301. France: Moselle; ST; ♂,L; lost?. 

fastidiosa. Netherlands, Denmark, Norway. Betula pubescens (Betulaceae). 

Dasineura fastidiosa Roskam 1979: 299. Netherlands: Meijendel; HT; ♂; RMNH. 

fedtshenkoi. Kazakhstan. Clematis soongarica (Rosaceae). 

Dasyneura fedtshenkoi Fedotova 1984f: 640. Kazakhstan: Zailiïskiï Alatau, Syugaty Mts, Kok-Pek pass; HT; ♂; 

unstated. 

festucae. UK, Germany, Poland, Czechoslovakia. Festuca rubra (Poaceae). 

Dasyneura festucae Barnes 1939e: 172. UK: England, Oxfordshire; ST; ♂,♀,L; BMNH. 

filicae. USA (New York). 

Dasyneura filicae Felt 1907a: 19. USA: New York, Albany; HT; ♂; NYSM. 

Dasyneura filicis Felt 1907c: 115, unjust. emend. of filicae. 

filipendulae. UK, Sweden, France, Czechoslovakia, Yugoslavia. Filipendula vulgaris; F. hexapetala (Rosaceae). 

Perrisia filipendulae Kieffer 1909b: 29. France; T; G; lost?. 

*fistulosa. Australia (New South Wales, Victoria). Acacia mearnsii, A. irrorata (Fabaceae). 

Dasineura fistulosa Kolesik 2005: 468. Australia: Victoria: Bundoora; HT; ♂; SAMA. 

flavescens. USA (New York). 

Dasyneura flavescens Felt 1908b: 344. USA: New York, Albany; HT; ♀; NYSM. 

flavoabdominalis. USA (New York). 

Dasyneura flavoabdominalis Felt 1908b: 350. USA: New York, Albany; HT; ♀; NYSM. 

flavoscuta. USA (New York). 

Dasyneura flavoscuta Felt 1908b: 350. USA: New York, Albany; HT; ♀; NYSM. 

floralis. Ukraine. Onobrychis sp. (Fabaceae). 

Dasyneura floralis Marikovskij 1956: 184. Ukraine: Kiyev Prov., Belaya Tserkov'; ST; ♂,♀; ZMAS. 

florida. USA (Florida). 

Dasyneura florida Felt 1908b: 346. USA: Florida, Key West; HT; ♀; USNM. 

foliumcrispans. Germany, Czechoslovakia. Symphytum officinale (Boraginaceae). 

Dichelomyia foliumcrispans Rübsaamen 1895c: 259. Germany; ST; ♂,♀,L; ZMHB. 

folliculi. USA (Pennsylvania to Massachusetts). Solidago "canadensis"; S. gigantea, S, rugosa (Asteraceae). 

Dasyneura folliculi Felt 1908b: 348. USA: New Jersey, Jamesburgh; HT; ♀; NYSM. Ref.: Dorchin et al. 2007 (tax'y, 

biol.) 

Dasyneura radifolii Felt 1909b: 289. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

frangulae. UK, Sweden, Germany. Rhamnus frangula (Rhamnaceae). 

Dasyneura frangulae Rübsaamen 1917: 51. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

fraxinea. Widespread Europe. Fraxinus excelsior (Oleaceae). 

Dasyneura fraxinea Kieffer 1907c: 523. Germany: Annaberg; ST; ♂,♀,P,L; lost?. 

fraxini. Widespread Europe, Algeria; Pliocene fossil/Germany. Fraxinus excelsior (Oleaceae). 

Cecidomyia fraxini Bremi 1847a: 18. Switzerland: vic. Zurich; ST; LT; ETHZ?. 

Cecidomyia fraxinicola Hardy 1854e: 141. UK: Scotland, vic. Penmanshiel; T; LT; lost. 

Perrisia fraxini Kieffer 1897b: 301. France: Moselle; ST; ♂,♀; lost?.

196 

fraxinifolia. USA (New York). Fraxinus sp. (Oleaceae). 

Dasyneura fraxinifolia Felt 1907d: 12. USA: New York, Bath; ST; ♂; NYSM. 

fructicola. France. Myosotis palustris (Boraginaceae). 

Perrisia fructicola Kieffer 1909b: 15. France: Moulins; T; LT; lost?. 

fructum. UK, Germany, Poland, Czechoslovakia. Cerastium triviale (Caryophyllaceae). 

Dichelomyia fructum Rübsaamen 1895c: 258. Germany: Berlin, "Tegeler Weg"; ST; ♂,♀,L; ZMHB. 

Perrisia fructuum Kieffer 1913i: 76, unjust. emend. for fructum Rübsaamen. 

fulva. USA (New York, Pennsylvania). 

Dasyneura fulva Felt 1908b: 349. USA: New York, Nassau; HT; ♀; NYSM. 

fulvicola. Japan (Honshu). Artemisia sp. (Asteraceae). 

Dasynewca fulvicola Shinji 1938m: 335. Japan: Honshu; T; ♀; lost. 

*furcata. Australia (Victoria). Acacia melanoxylon (Fabaceae). 

Dasineura furcata Kolesik 2005: 471. Australia: Victoria: Braeside; HT; ♂; SAMA. 

fusca. UK, Germany. Crataegus sp.; Crataegus oxyacantha (Rosaceae). 

Dasyneura fusca Rübsaamen 1914: 103. Germany; ST; ♂,♀,L; ZMHB. 

galeopsis. France. Galeopsis tetrahit (Lamiaceae). 

Dasyneura galeopsis Kieffer 1897a: 19. France: château de La Grange, nr Thionville; T; LT; lost?. 

galiicola. Widespread Europe. Galium mollugo (Rubiaceae). 

Cecidomyia galiicola Löw 1880: 33. Austria: Wartenstein Castle nr Gloggnitz; ST; ♂,♀,L; NHMW. 

gallica. UK, France. Ulex europaeus (Fabaceae). 

Perrisia gallica Kieffer 1909b: 31. France; T; G; lost?. 

gardoquiae. Chile. Gardoquia gilliesii (Lamiaceae). 

Dasyneura gardoquiae Kieffer & Herbst 1905: 65. Chile: vic. Concepción; T; LT; lost. 

geisenheyneri. Germany, Yugoslavia. Hippocrepis comosa (Fabaceae). 

Perrisia geisenheyneri Kieffer 1904a: 67. Germany: vic. Kreuznach; ST; ♂,♀,P,L; lost?. 

gemmae. USA (Utah). Salix sp. (Salicaceae). 

Dasyneura gemmae Felt 1909b: 288. USA: Utah, Logan; ST; ♂,♀; NYSM. 

genistarum. Czechoslovakia, Austria, Yugoslavia. Genista pilosa (Fabaceae). 

Perrisia genistarum Kieffer 1909b: 9. Austria; T; LT; NHMW?. 

gentianae. UK, Netherlands, Romania. Gentiana campestris (Gentianaceae). 

Perrisia gentianae Kieffer 1909b: 9. UK: Scotland; T; G; lost?. 

gentneri. UK, Sweden, Finland, Netherlands, Germany, Czechoslovakia, Russia (Europe, Western Siberia); poss. 

immigr.: USA (Washington, Oregon, New York). Trifolium repens; T. hybridum, T. pratense. (Fabaceae). 

Dasyneura gentneri Pritchard 1953b: 131. USA: Oregon, Talent; HT; ♂; USNM. 

geranii. UK, Sweden, France, Germany, Czechoslovakia, Hungary. Geranium cicutarium (Geraniaceae). 

Perrisia geranii Kieffer 1907b: 44. France: Seine et Marne, Vulaines sur Seine, près d'Avon; ST; ♀,P,L,egg; lost?. 

gibsoni. UK; poss. immigr.: Canada (British Columbia, Ontario), USA (Indiana, Massachusetts). Cirsium arvense 

(Asteraceae). This species is possibly synonymous with Jaapiella cirsiicola (Rübsaamen) 

Dasyneura gibsoni Felt 1911m: 479. Canada: Ontario, Ottawa; ST; ♂,♀; NYSM. 

gigantea. Brazil (Paraná, Santa Catarina). Psidium cattleianum (Myrtaceae). 

Dasyneura gigantea Angelo & Maia 1999: 193. Brazil: Paraná, Piraquara; HT; ♂; MNRJ. 

giraudi. Austria. Astragalus austriacus (Fabaceae). 

Cecidomyia giraudi Frauenfeld 1863: 1233. Austria: Berndorf; ST; ♂,♀; ?. 

glandis. USA (Missouri). Quercus sp. (Fagaceae). 

Dasyneura glandis Felt 1908b: 342. USA: Missouri, Kirkwood; ST; ♀,P; USNM. 

*glauca. Australia (South Australia). Acacia pendula (Fabaceae). 

Dasineura glauca Kolesik 2005: 468. Australia: South Australia: Adelaide, Goodwood; HT; ♂; SAMA. 

glechomae. Widespread Europe; immigr.: USA (Maryland). Glechoma hederacea (Lamiaceae). 

Cecidomyia glechomae Kieffer 1889e: 263. France: Bitche; ST; ♂,♀,L; lost?. 

gleditchiae. USA (widespread eastern); immigr.: USA (California, Idaho), widespread Europe. Gleditsia triacanthos 

(Fabaceae). 

Cecidomyia gleditchiae Osten Sacken 1866: 219. USA: Rhode Island, Newport; ST; ♂,♀,L; USNM.

197 

Cecidomyia gleditschiae Beutenmüller 1892: 266, missp. of gleditchiae. 

globosa. Brazil (Rio de Janeiro). Eugenia rotundifolia (Myrtaceae). 

Dasineura globosa Maia 1996b: 1002. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

*glomerata. Australia (New South Wales, Victoria). Acacia mearnsii, Acacia spp. (Fabaceae). 

Dasineura glomerata Kolesik 2005: 466. Australia: Victoria: Keilor; HT; ♂; SAMA. 

glyciphyli. UK, France, Germany, Poland, Czechoslovakia, Austria, Hungary, Romania, Yugoslavia. Astragalus cicer 

(Fabaceae). 

Dasyneura glyciphylli Rübsaamen 1912e: 286. Germany: vic. Laacher See; ST; ♂,♀,L; ZMHB. 

glycyrrhizicola. Kazakhstan. Glycyrrhiza uralensis (Fabaceae). 

Dasineura glycyrrhizicola Fedotova 1985c: 37. Kazakhstan: Karzhantau Mtn Range, Badanskoe canyon, 30 km S 

Lenger; HT; ♂; ZKAS. 

gmelinii. Kazakhstan. Lathyrus gmelinii (Fabaceae). 

Dasineura gmelinii Fedotova 1992g: 47. Kazakhstan: northeastern Kungeï Alatau, 20 km SE Saryzhas; HT; ♂; 

ZMAS. 

gossypii. India (Bihar). Gossypium sp. (Malvaceae). 

Dasyneura gossypii Felt 1916a: 29. India: Bihar, Pusa; ST; ♀; NYSM. 

gracilicornis. Chile. Unspecified shrub. 

Perrisia gracilicornis Kieffer & Herbst 1905: 64. Chile: vic. Concepción; T; G; lost. 

graminis. USA (New York). Agrostis tenuis (Poaceae). 

Dasyneura graminis Felt 1908b: 342. USA: New York, Albany; ST; ♀; NYSM. 

grasseti. Algeria. Erica arborea (Ericaceae). 

Dasyneura grasseti Barnes 1935c: 140. Algeria: Reghaïa; ST; ♂,♀; BMNH. 

halimii. Portugal, Spain. Halimium occidentale, H. heterophyllum (Cistaceae). 

Perrisia halimii Tavares 1902c: 183. Portugal: Serra da Estrella & Arredores de Setubal; ST; ♂,♀; TAVC. 

halimodendronifolia. Kazakhstan. Halimodendron halodendron (Fabaceae). 

Dasineura halimodendronifolia Fedotova 1990f: 76. Kazakhstan: northeastern Karatau Mtn Range, nr Bilikul' R., 5 

km NW Karabastau; HT; ♂; ZMAS. 

harrisoni. UK, Netherlands, Germany, Czechoslovakia, Romania, Yugoslavia. Filipendula ulmaria; F. hexapetala 

(Rosaceae). 

Dasyneura jaapiana Rübsaamen 1917: 50, jun. prim. hom. jaapiana Rübsaamen 1914. Germany: Prignitz, Triglitz; 

ST; ♂,♀,L; ZMHB. 

Perrisia harrisoni Bagnall 1922: 184, new name for Dasyneura jaapiana Rübsaamen 1917. Ref.: Harris 2010a (gall). 

hebefolia. New Zealand. Hebe salicifolia (Scrophulariaceae). 

Dasyneura hebefolia Lamb 1951: 210. New Zealand: North I., Mayor I.; HT; ♂; BMNH. 

helenae. Sweden. Fraxinus excelsior (Oleaceae). 

Dasineura helenae Sylvén 1993: 295. Sweden: Öland, Kalkstad; HT; ♀; NHRS. 

helianthemiflorae. Kazakhstan. Helianthemum soongoricum (Asteraceae). 

Dasineura helianthemiflorae Fedotova 1992a: 84. Kazakhstan: floodplain Timirlik R., 25 km SE Chundzha; HT; ♂; 

ZMAS. 

herminii. Portugal, Spain. Halimium occidentale (Cistaceae). 

Perrisia herminii Tavares 1902c: 182. Portugal: Estrella; ST; ♂,♀,L.; TAVC. 

hildebrandi. USA (California). Arctostaphylos sp. (Ericaceae). 

Rhabdophaga hildebrandi Felt 1928b: 58. USA: California, 7 mi SE Kelseyville; ST; ♂,♀,L; NYSM. 

hirticornis. USA (New York). 

Rhabdophaga hirticornis Felt 1909b: 290. USA: New York, S; ♂,♀; NYSM. 

hisarenis. India (Haryana). Brassica campestris, B. rapa, B. juncea (Brassicaceae). 

Dasineura hisarenis Sharma & Singh 1991: 431. India: Haryana, Hisar; HT; ♂; NZSI. 

holosteae. UK, France. Stellaria holostea (Caryophyllaceae). 

Perrisia holosteae Kieffer 1909b: 29. France; T; LT; lost?. 

hybanthi. Australia (South Australia). Hybanthus floribundus (Violaceae). 

Dasineura hybanthi Kolesik & Skuhravá 1997: 240. Australia: South Australia, Morialta Conservation Pk, 13 km NE 

Adelaide; HT; ♂; SAMA.

198 

hygrophila. Widespread Europe. Galium palustre (Rubiaceae). 

Cecidomyia hygrophila Mik 1883: 209. Austria: nr Hammern in Mühlviertel; ST; ♂,♀,P,L; NHMW. 

hyperici. Widespread Europe. Hypericum perforatum (Hypericaceae). 

Cecidomyia hyperici Bremi 1847a: 53. Switzerland: on the Hottingerberge & in Burghölzi; ST; ♂,♀,L; ETHZ?. 

hyssopi. Austria. Hyssopus sp. (Lamiaceae). 

Perrisia hyssopi Kieffer 1909b: 10. Austria; T; G; NHMW?. 

ilicis. Portugal. Quercus ilicis (Fagaceae). 

Perrisia ilicis Tavares 1919b: 61. Portugal: Sobral do Campo, Beira Baixa; ST; ♂,P; TAVC. 

incola. Kazakhstan. Spiraea hypericifolia (Rosaceae). 

Dasineura incola Fedotova 1990f: 77. Kazakhstan: Tarbagataï Mtn Range, 30 km N Ayagguza; HT; ♂; ZMAS. 

inflata. Germany. Carex otrubae (Cyperaceae). 

Dasineura inflata Stelter 1986: 197. Germany: Kr. Grimmen, nr Kirch Baggendorf; HT; ♂; SMTD. 

ingeris. Sweden. Salix viminalis (Salicaceae). 

Dasineura ingeris Sylvén & Lövgren 1995: 61. Sweden: Scania, Svalöv; HT; ♀; NHRS. 

interbracta. Netherlands, Norway, Sweden. Betula pubescens; B. pendula (Betulaceae). 

Dasineura interbracta Roskam 1979: 292. Netherlands: Meijendel; HT; ♂; RMNH. 

inventa. Kazakhstan. Aster limonifolius (Asteraceae). 

Dasineura inventa Fedotova 1996a: 547. Kazakhstan: Kirgizskiï Alatau, 18 km S Merke, upper course Merke R.; HT; 

♂; ZMAS. 

irregularis. Widespread Europe. Acer pseudoplatanus; A. campestre (Aceraceae). 

Cecidomyia irregularis Bremi 1847a: 30. Switzerland: Burghölzchen, nr Zurich; ST; LT; ETHZ?. 

Cecidomyia acercrispans Kieffer 1888c: 266. France: Moselle; ST; ♂,♀,L; lost?. 

ivashenkoae. Kazakhstan. Erigeron cabulicus (Asteraceae). 

Dasineura ivashenkoae Fedotova 1996a: 542. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Ul'ken-Kaïndy 

pass, 20 km SE Novonikolaevka; HT; ♂; ZMAS. 

jaapi. Austria. Veronica fruticans (Scrophulariaceae). 

Dasyneura jaapi Rübsaamen 1914: 101. Austria: Tirol; ST; ♂,L; ZMHB. 

karnerensis. USA (New York). 

Dasyneura karnerensis Felt 1908b: 341. USA: New York, Karner; HT; ♀; NYSM. 

kellneri. Widespread Europe, Russia (Western Siberia). Larix decidua (Pinaceae). 

Cecidomyia kellneri Henschel 1875: 183. Austria: Oberermark; T; LT; NHMW?. 

Cecidomyia laricis Löw 1878: 393. Austria: Steiermark & Semmering; ST; ♂,♀,L; NHMW. 

kiefferi. UK, France, Ukraine. Hedera helix (Araliaceae). 

Dasyneura kiefferi Marchal 1896: 99. France: Hauts de Seine, Fontenay-aux-Roses; ST; A,L; MNHN?. 

kiefferiana. Widespread Europe. Epilobium angustifolium (Onagraceae). 

Cecidomyia kiefferiana Rübsaamen 1891b: 1. Germany: Siegen; ST; ♂,♀,P,L; ZMHB. 

kleini. Germany. Sisymbrium officinale (Brassicaceae). 

Cecidomyia kleini Rübsaamen 1892b: 398. Germany; ST; ♀,L; ZMHB. 

koesterbecki. Denmark, Germany. Carex acutiformis (Cyperaceae). 

Dasineura kösterbecki Stelter 1986: 191. Germany: Kr. Rostock, betw. Kösterbeck & Fresendorf; HT; ♂; SMTD. 

kungeica. Kazakhstan. Geranium collinum (Geraniaceae). 

Dasineura kungeica Fedotova 1993o: 20. Kazakhstan: Dzhungarskiï Alatau, Kungeï Alatau, 78 km SE Uch-Aral, 8 

km SE Koktumy; HT; ♂; ZMAS. 

lamii. UK, France, Poland, Czechoslovakia, Austria, Romania. Lamium maculatum (Lamiaceae). 

Perrisia lamii Kieffer 1909b: 13. Austria; T; G; NHMW?. 

lamiicola. UK, France, Czechoslovakia, Austria. Lamium maculatum (Lamiaceae). 

Cecidomyia lamiicola Mik 1888: 35. Austria: Salzburg; ST; ♂,♀,L; NHMW. 

lappulae. Kazakhstan. Lappula microcarpa (Boraginaceae). 

Dasineura lappulae Fedotova 1993l: 80. Kazakhstan; Dzhungarskiï Alatau, Altyn-Emel' Mts, Koayandytau, nr 

Lugovoye; HT; ♂; ZMAS. 

lathierei. Italy, Middle East. Olea europaea (Oleaceae). 

Dasyneura lathierei Del Guercio 1910: 292. Italy; ST; ♀,L; lost.

199 

lathyri. UK, France, Italy, Czechoslovakia, Yugoslavia, Russia (Europe). Lathyrus pratensis (Fabaceae). 

Perrisia lathyri Kieffer 1909b: 13. France; ST; LT; lost?. 

lathyricola. Widespread Europe, Russia (Western Siberia). Lathyrus pratensis; L. latifolius (Fabaceae). 

Cecidomyia lathyricola Rübsaamen 1890b: 26. Germany: vic. Siegen; ST; ♂,♀,L; ZMHB. 

lathyrina. Germany. Lathyrus pratensis (Fabaceae). 

Cecidomyia lathyrina Rübsaamen 1890b: 28. Germany: vic. Siegen; ST; ♂,♀,L; ZMHB. 

leguminicola. Widespread Europe; poss. immigr.: widespread Nearctic. Trifolium pratense; T. medium (Fabaceae). 

Cecidomyia trifolii Lintner 1879a: 44, jun. prim. hom. trifolii Löw. USA: New York, Elmira; ST; LT; NYSM. 

Cecidomyia leguminicola Lintner 1879b: 121, new name for Cecidomyia trifolii Lintner. 

Cecidomyia flosculorum Kieffer 1890e: 200. France: betw. Bitche & Sturzelbronn; ST; ♂,♀,L; lost?. 

lenkiewicziae. Egypt. Zilla spinosa (Brassicaceae). 

Perrisia lenkiewicziae Debski 1918: 18. Egypt: Wadi-Hof & Hélouan, Wadi Tuck-Key; T; G; ?. 

lepidii. USA (District of Columbia). Lepidium sp. (Brassicaceae). 

Dasyneura lepidii Felt 1908b: 346. USA: District of Columbia; HT; ♀; USNM. 

lepidiis. Kazakhstan. Lepidium perfoliatum (Brassicaceae). 

Dasineura lepidiis Fedotova 1992n: 106. Kazakhstan: Dzhungarskiï Alatau, 255 km NW Alma-Ata; HT; ♂; ZMAS. 

Dasineura lepidiiphaga Fedotova 1994a: 124, unjust. new name for Dasineura lepidiis Fedotova. 

*thalictrifoliae. Kazakhstan. Thalictrum collinum (Ranunculaceae). 

Dasineura thalictrifoliae Fedotova 1992n: 104. Kazakhstan: Aigyr Kum Desert, 50 km NE Zaisan; HT; ♂; ZMAS. 

ligulariae. Kazakhstan. Ligularia naryensis; L. thyrsoidea (Asteraceae). 

Dasineura ligulariae Fedotova 1993l: 76. Kazakhstan: Dzhungarskiï Alatau, 45 km NW Panfilov, Borakhudzir R. 

valley; HT; ♂; ZMAS. 

lini. India (Bihar). Linum usitatissimum (Linaceae). 

Dasyneura lini Barnes 1936: 273. India: Bihar, Pusa; ST; ♂,♀; BMNH. 

linosyridis. Germany, Romania. Aster linosyris (Asteraceae). 

Dasyneura linosyridis Möhn 1958: 3. Germany: Kyffhäuser, Ochsenburg; HT; ♂; SMNS. 

lithospermi. Widespread Europe. Lithospermum officinalis (Boraginaceae). 

Cecidomyia lithospermi Loew 1850a: 36. Poland; ST; ♂,♀; ZMHB?. 

loewiana. UK, Netherlands, Germany, Poland, Czechoslovakia. Vicia cracca (Fabaceae). 

Dasyneura loewiana Rübsaamen 1917: 45. Germany: Kreuznach, Hamburg, & Rhine Prov.; ST; ♂,♀,L; ZMHB. 

loniceraegemmae. Kazakhstan. Lonicera tatarica (Caprifoliaceae). 

Dasineura loniceraegemmae Fedotova 1992a: 83. Kazakhstan: Aïgyrkumy desert, 50 km NE Zaïsana, 4 km S 

Kabyrgatal; HT; ♂; ZMAS. 

lotharingiae. Widespread Europe. Cerastium glomeratum, C. triviale, C. arvense (Caryophyllaceae). 

Cecidomyia lotharingiae Kieffer 1888e: 107. France: Moselle, Bitche & Hohlwegen; ST; ♂,♀,P,L; lost?. 

lupini. USA (California). Lupinus spp. (Fabaceae). 

Dasyneura lupini Felt 1916f: 413. USA: California, San Francisco, Buena Vista Pk; ST; ♂,♀; NYSM. 

lupinorum. USA (California). Lupinus arboreus; Lupinus spp. (Fabaceae). 

Dasineura lupinorum Gagné 1993b: 542. USA: California, Bodega Bay, Bodega Bay Marine Res.; HT; ♂; USNM. 

lupulinae. Widespread Europe. Medicago lupulina; M. sativa (Fabaceae). 

Cecidomyia lupulinae Kieffer 1891c: 233. France: Guinkirchen & château de La Grange, nr Thionville; ST; G; lost?. 

lysimachiae. USA (Indiana, New York, Massachusetts, New Jersey). Lysimachia quadrifolia (Primulaceae). 

Cecidomyia lysimachiae Beutenmüller 1907b: 305. USA: New York, New York, Bronx Pk, & New Jersey, Fort Lee; 

ST; ♂,♀; AMNH. 

mali. Widespread Europe, Russia (WS, Far East); immigr.: USA (Massachusetts, New Hampshire), Canada (New 

Brunswick), Argentina, New Zealand. Pyrus malus (Rosaceae). 

*Dichelomyia mali Kieffer 1894k: unnumbered p. 2. Unstated, presumably France: Bitche; HT; egg; lost? 

Perrisia mali Kieffer 1904b: 345. France: Bitche; ST; ♂,♀,L; lost?. Redescription. 

marginalis. Brazil (Rio de Janeiro). Eugenia umbelliflora & E. rotundifolia (Myrtaceae). 

Dasineura marginalis Maia 2005a: 349. Brazil: Rio de Janeiro, Macaé; HT; ♂; MNRJ. 

marginata. USA (New York). 

Rhabdophaga marginata Felt 1908b: 352. USA: New York, Albany; HT; ♀; NYSM.

200 

mariae. Spain. Astragalus lusitanicus (Fabaceae). 

Dasineura mariae Sylvén 1993: 293. Spain: Sierra Blanca, Ojén; HT; ♀; NHRS. 

maritima. USA (Massachusetts). Lathyrus maritimus (Fabaceae). 

Dasyneura maritima Felt 1909b: 288. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

markakolica. Kazakhstan. Scutellaria supina (Lamiaceae). 

Dasineura markakolica Fedotova 1996d: 12. Kazakhstan: southern Altaï, Markakol'skiï Res., nr L. Markakol', nr 


medicaginis. Widespread Europe, Russia (Western Siberia). Medicago sativa (Fabaceae). 

Cecidomyia medicaginis Bremi 1847a: 18. Switzerland: vic. Zurich; ST; LT; ETHZ?. 

Cecidomyia ignorata Wachtl 1884: 163. Austria: vic. Vienna; ST; ♂,♀; NHMW?. 

medullaris. France. Salix aurita (Salicaceae). 

Cecidomyia medullaris Kieffer 1892d: 253. France: Bitche; ST; ♂,♀,P,L,egg; lost?. 

meibomiae. USA (New York, North Carolina). Desmodium rigida, D. laevigata, D. pauciflora (Fabaceae). 

Cecidomyia meibomiae Beutenmüller 1907a: 390. USA: New York, New York, Staten I., & North Carolina, 

Buncombe Co., Valley of the Black Mts; ST; LT; AMNH. 

meliloti. USA (New York). 

Dasyneura meliloti Felt 1907a: 20. USA: New York, Albany; HT; ♂; NYSM. 

miki. UK, Austria. Centaurea scabiosa (Asteraceae). 

Perrisia miki Kieffer 1909b: 6. Austria: T; G; NHMW?. 

mimosae. Egypt. Mimosa nilotica (Fabaceae). 

Perrisia mimosae Kieffer 1909b: 14. Egypt: Cairo; T; G; NHMW?. 

minardii. Italy. Asparagus spp. (Liliaceae). 

Perrisia minardii De Stefani 1913: 19 (publ'n not seen). Italy: Sicily; ?; ?; lost. 

minoterminalis. Germany. Salix aurita (Salicaceae). 

Dasyneura minoterminalis Stelter 1969b: 137. Germany: Kr. Rostock, betw. Teschendorf & Gross Lüsewitz; HT; ♂; 

SMTD. 

minungula. Germany. Carex paniculata (Cyperaceae). 

Dasineura minungula Stelter 1986: 193. Germany: Kr. Rostock, betw. Kösterbeck & Fresendorf; HT; ♂; SMTD. 

miranda. Kazakhstan. Erigeron pseudoseravshanicus (Asteraceae). 

Dasineura miranda Fedotova 1996a: 540. Kazakhstan: Kungeï Alatau, Karabulak canyon, 12 km SW Malyï 

Zhalanash; HT; ♂; ZMAS. 

modesta. USA (New York). 

Dasyneura modesta Felt 1908b: 345. USA: New York, Albany; HT; ♀; NYSM. 

mojynkumensis. Kazakhstan. Sisymbrium polymorphum (Brassicaceae). 

Dasineura mojynkumensis Fedotova 1994a: 115. Kazakhstan: Moïynkumy Desert, 17 km NE Sarykoba; HT; ♂; 

ZMAS. 

multianulata. USA (New York). 

Dasyneura multianulata Felt 1908b: 346. USA: New York, Nassau; HT; ♀; NYSM. 

Dasyneura multiannulata Felt 1915j: 156, unjust. emend. of multianulata. 

myosotidis. UK, Sweden, Denmark, Netherlands, France, Germany, Poland, Czechoslovakia. Myosotis palustris 

(Boraginaceae). 

Perrisia myosotidis Kieffer 1902c: 115. France: Althorn, nr Bitche; ST; ♂,♀,L; lost?. 

Perrisia palustris Kieffer 1909b: 15. France; ST; LT; lost?. 

myrciariae. Brazil (Rio de Janeiro). Myrciaria floribunda (Myrtaceae). 

Dasineura myrciariae Maia 1996b: 1003. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

myrtylli. Germany. Vaccinium myrtillus (Ericaceae). 

Dasyneura myrtylli Rübsaamen 1916b: 518. Germany; ST; ♀; ZMHB. 

napi. Widespread Europe. Brassica napi; B. oleracea. (Brassicaceae). 

Cecidomyia napi Loew 1850a: 30. Poland; ST; G; ZMHB?. Loew's description of the gall fits exactly that of 

brassicae Winnertz. 

Cecidomyia brassicae Winnertz 1853: 231. Germany; ST; ♂,♀,L; MNHN?. 

nasturtii. Yugoslavia. Nasturtium sp. (Brassicaceae).

201 

Dasyneura nasturtii Rübsaamen 1916b: 517. Yugoslavia: Montenegro, Hercegnovi; ST; ♂,♀,L; ZMHB. 

nervicola. Germany. Hieracium auricula, H. pilosella (Asteraceae). 

Perrisia nervicola Kieffer 1909b: 9. Germany; T; G; ZMHB?. 

nipponica. Japan (Honshu). Larix decidua (Pinaceae). 

Dasyneura nipponica Inouye 1966: 20. Japan: Honshu, Yamanashi Pref.; HT; ♂; HFES. 

nodosa. USA (New York). 

Oligotrophus nodosus Felt 1907a: 27. USA: New York, Albany; HT; ♂; NYSM. 

nummulariifoliae. Kazakhstan. Lonicera nummulariifolia (Caprifoliaceae). 

Dasineura nummulariifoliae Fedotova 1996f: 50. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 12 km SE 

Novonikolaevki; HT; ♂; ZMAS. 

odoratae. Germany. Viola odorata (Violaceae). 

Dasineura odoratae Stelter 1982c: 261. Germany: Schmalkalden; HT; ♂; SMTD. 

*oldfieldii. Australia (Western Australia). Acacia oldfieldii, A. neurophylla (Fabaceae). 

Dasineura oldfieldii Kolesik 2005: 472. Australia: Western Australia: Kalbarri; HT; ♂; SAMA. 

oleae. Mediterranean Region. Olea europaea (Oleaceae). 

Corethra oleae Angelini 1831: 313. Italy: Verona Prov.; ST; A, L; lost. 

Cecidomyia oleae Löw 1885a: 109. Croatia: Rijeka; ST; ♂,♀,L; NHMW. 

oleiperda. Eritrea. Olea chrysophylla (Oleaceae). 

Rhabdophaga oleiperda Del Guercio 1918a: 34. Eritrea; ST; A,P,L; lost. Removed from Rabdophaga because larvae 

and biology (i.e., not on Salicaceae) not typical of that genus. 

*oshanesii. Australia (Queensland, New South Wales). Acacia oshanesii (Fabaceae). 

Dasineura oshanesii Kolesik 2005: 464. Australia: Queensland, Beerwah; HT; ♂; SAMA. 

oxyacanthae. Widespread Europe. Crataegus oxyacantha; C. monogyna (Rosaceae). 

Dasyneura oxyacanthae Rübsaamen 1914: 103. Germany: nr Metternich; ST; ♂,L; ZMHB. 

Perrisia oyensis Tavares 1922c: 140. Spain: Gallicia, Oya; ST; ♂,♀,P,L; TAVC. 

oxycoccana. Widespread eastern Nearctic; immigr.: Latvia, Italy. Vaccinium sp.; Vaccinium spp. (Ericaceae). 

Cecidomyia vaccinii Smith 1890: 31, jun. prim. hom. vaccinii Osten Sacken. USA: New Jersey, Jamesburg; ST; ♂; 


Cecidomyia oxycoccana Johnson 1899: 80, new name for Cecidomyia vaccinii Smith. 

oxytropifolia. Kazakhstan. Oxytropis glabra, O. puberula (Fabaceae). 

Dasineura oxytropifolia Fedotova 1984e: 1029. Kazakhstan: Zailiïskiï Alatau, along Charyn R.; HT; ♂; ZKAS. 

oxytropigemma. Kazakhstan. Oxytropis glabra (Fabaceae). 

Dasineura oxytropigemma Fedotova 1984e: 1027. Kazakhstan: Zailiïskiï Alatau, along Charyn R.; HT; ♂; ZKAS. 

paeoniae. Kazakhstan. Paeonia hybrida (Paeoniaceae). 

Dasineura paeoniae Fedotova 1989d: 25. Kazakhstan: Altaï, Ivanovskiï Mts, 32 km NE Leninogorsk, nr 

Poperechnoe; HT; ♂; ZKAS. 

pallasi. Kazakhstan. Aster alpinus (Asteraceae). 

Dasineura pallasi Fedotova 1996a: 546. Kazakhstan: Kungeï Alatau, Kegen' pass, 40 km SW Chundzha; HT; ♂; 

ZMAS. 

panteli. UK, Spain. Quercus petraea, Q. robur (Fagaceae). 

Perrisia panteli Kieffer 1909b: 21. Spain; T; G; lost?. 

papaveris. Widespread Europe, Israel. Papaver rhoeas, P. dubium (Papaveraceae). 

Cecidomyia papaveris Winnertz 1853: 229. Germany: Aachen; ST; ♂,♀,L; MNHN?. 

papivora. India (Uttar Pradesh). Carica papaya (Caricaceae). 

Dasineura papivora Grover & Prasad 1966c: 121. India: Uttar Pradesh, Allahabad; HT; ♀; PGCA. 

parthenocissi. Canada (Ontario), USA (widespread northeastern). Parthenocissus quinquefolia (Vitaceae). 

Cecidomyia parthenocissi Stebbins 1910: 44. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

pedalis. USA (New York). 

Dasyneura pedalis Felt 1908b: 350. USA: New York, Newport; HT; ♂; NYSM. 

*pediculariflorae. Kazakhstan. Pedicularis achilleifolia (Scrophulariaceae). 

Dasineura pediculariflorae Fedotova 2005b: 897. Kazakhstan: Tarbagatai Mts., 40 km S Tarbagatai Village; HT; ♂; 

ZMAS.

202 

peinei. UK, Germany, Russia (Europe). Sanguisorba officinalis (Rosaceae). 

Cecidomyia peinei Rübsaamen 1890a: 27. Germany: Weidenau, vic. Münkershütten; ST; ♂,♀,L; ZMHB. 

pellex. USA (widespread eastern). Fraxinus americana (Oleaceae). 

Cecidomyia pellex Osten Sacken 1862: 199. USA: District of Columbia; T; LT; lost. 

pentaphylloidiflora. Kazakhstan. Pentaphylloides parviflora (Rosaceae). 

Dasineura pentaphylloidiflora Fedotova 1990b: 105. Kazakhstan: Tarbagataï Mtn Range, Maïlybaï valley, 78 km SE 

Koktum; HT; ♂; ZKAS. 

pergandei. USA (Colorado). Prunus sp. (Rosaceae). 

Dasyneura pergandei Felt 1911m: 480. USA: Colorado, Glenn Ayrie; ST; ♂,♀; USNM. 

periclymeni. Widespread Europe. Lonicera periclymenum (Caprifoliaceae). 

Cecidomyia periclymeni Rübsaamen 1889a: 54. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

perigonialis. Kazakhstan. Erigeron seravschanicus (Asteraceae). 

Dasineura perigonialis Fedotova 1996a: 543. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Ul'ken-Kaïndy 

pass, 20 km SE Novonikolaevka; HT; ♂; ZMAS. 

peyerimhoffi. Algeria. Coronilla valentina (Fabaceae). 

Perrisia peyerimhoffi Kieffer 1919: 191. Algeria: Oued Djer; Zaouïa des Mouzaïa; ST; ♂,♀,L; lost?. 

phlomicola. Russia (Far East). Phlomis maximoviczii (Lamiaceae). 

Dasyneura phlomicola Kovalev 1967: 115. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; KOVC. 

phyteumatis. France, Germany, Poland, Czechoslovakia, Switzerland, Austria, Italy. Phyteuma orbiculare, P. 

spicatum (Campanulaceae). 

Cecidomyia phyteumatis Löw 1885b: 487. Austria, Germany, Switzerland; ST; ♀,L; NHMW. 

*pilifera. Australia (Western Australia, Victoria, ACT). Acacia baileyana, A. dealbata, A. decurrens (Fabaceae). 

Dasineura pilifera Kolesik 2005: 462. Australia: Victoria: Lilydale; HT; ♂; SAMA. 

pini. USA (New York). 

Oligotrophus pini Felt 1907a: 28. USA: New York, Albany; HT; ♂; NYSM. 

piperitae. USA (New York). Mentha gentilis, M. piperita (Lamiaceae). 

Dasyneura piperitae Felt 1908b: 342. USA: New York, Nassau; ST; ♀; NYSM. 

pirolae. France. Pyrola minor (Ericaceae). 

Perrisia pirolae Kieffer 1909b: 17. France; T; G; lost?. 

plectranthi. Russia (Far East). Plectranthus excisus (Lamiaceae). 

Dasyneura plectranthi Kovalev 1967: 113. Russia: Primorskiï Terr., Shtokovskoe Region; HT; ♂; KOVC. 

plicata. USA (New York, Massachusetts). Salix sp. (Salicaceae). 

Rhabdophaga plicata Felt 1908b: 352. USA; HT; ♀; USNM. 

plicatrix. Widespread Europe, Algeria; immigr.: Canada (British Columbia). Rubus caesius; Rubus spp. (Rosaceae). 

Ref.: Sinclair et al. 2009.zzz 

Cecidomyia plicatrix Loew 1850a: 36. Poland; ST; ♂,♀; ZMHB?. Ref.: Sinclair et al. 2009 (North American record, 

redescr.). 

poae. Sweden, Germany, Poland. Poa pratensis (Poaceae). 

Dasyneura poae Mühle 1957: 547. Germany: Steinach nr Straubing; ST; ♂,♀,L; ?. 

polygalae. UK, Austria, Italy. Polygala alpestris (Polygalaceae). 

Perrisia polygalae Kieffer 1909b: 18. Austria; T; LT; NHMW?. 

populeti. Widespread Europe. Populus tremula (Salicaceae). 

Cecidomyia populeti Rübsaamen 1889a: 57. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

populicola. Ukraine. Populus nigra (Salicaceae). 

Dasyneura populicola Kozarzhevskaya 1957: 1502 (attributed to P.I. Marikovskij but such description as is present is 

by Kozarzhevskaya). Ukraine: Voroshilovgrad Distr., Derkul' For. Stn; ST; ♂,♀; ?. 

populnea. France, Czechoslovakia, Austria, Hungary. Populus alba (Salicaceae). 

Perrisia populnea Kieffer 1909b: 18. Austria; T; G; NHMW?. 

potentillae. UK, Germany, Latvia, Lithuania, Poland, Czechoslovakia, Hungary, Romania, Yugoslavia. Potentilla 

argentea (Rosaceae). 

Cecidomyia potentillae Wachtl 1885: 193. Austria: vic. Znaim (Mähren), & Czech Republic: Znojmo; ST; ♂,♀,L; 

NHMW?.

203 

*potentillaeflora. Kazakhstan. Potentilla asiatica (Rosaceae). 

Dasineura potentillaeflora Fedotova 2005b: 899. Kazakhstan: Tarbagatai Mts., 40 km S Tarbagatai Village; HT; ♂; 

ZMAS. 

praecox. USA (Maryland, District of Columbia). Rhododendron periclymenoides (Ericaceae). 

Dasineura praecox Gagné 1996a: 228. USA: Maryland, Beltsville Agricultural Research Center; HT; ♂; USNM. 

pratensis. UK, France, Germany, Romania. Lathyrus pratensis (Fabaceae). 

Perrisia pratensis Kieffer 1909b: 13. Germany; T; G; lost?. 

praticola. Widespread Europe. Lychnis floscuculi (Caryophyllaceae). 

Cecidomyia praticola Kieffer 1892f: 216. France: Bitche; ST; ♂,♀,P,L,egg; lost?. 

procera. Germany, Romania. Linosyris vulgaris (Asteraceae). 

Dasyneura procera Rübsaamen 1914: 104. Germany: Rheinfels ruins nr St. Goar; ST; ♀,L; ZMHB. 

proxima. Eritrea. Olea chrysophylla (Oleaceae). 

Perrisia proxima Del Guercio 1918a: 51. Eritrea; T; LT; lost. 

prunicola. Czechoslovakia, Austria, Romania, Yugoslavia. Prunus spinosa; P. domestica (Rosaceae). 

Cecidomyia prunicola Löw 1889b: 538. Austria: vic. Vienna; ST; ♂,♀,L; NHMW. 

pseudacaciae. USA (New York, New Jersey). Robinia pseudoacacia (Fabaceae). 

Cecidomyia pseudacaciae Fitch 1859: 833. USA: New York, Salem; ST; ♀,L; lost. 

pseudococcus. UK, Sweden, Netherlands, France, Germany, Switzerland, Czechoslovakia, Romania. Salix caprea; S. 

aurita, S. cinerea (Salicaceae). 

Cecidomyia pseudococcus Thomas 1890: 301. Germany: nr Ohrdruf; ST; LT; ?. 

psoraleae. India (Maharashtra). Psoralea corylifolia (Fabaceae). 

Dasineura psoraleae Sharma 1987c: 186. India: Maharashtra, Aurangabad, Himayat Bagh; HT; ♂; INPC. 

pteridicola. Widespread Europe. Pteridium aquilinum (Polypodiaceae). 

Perrisia pteridicola Kieffer 1901c: 19. France: vic. Bitche; T; ♂; lost?. 

pteridis. Widespread Europe, Japan. Pteridium aquilina (Polypodiaceae). 

Cecidomyia pteridis Müller 1871b: 100. UK: England, vic. S. Norwood; ST; LT; lost. This name has been 

overlooked since its first use but the gall described is the same as that described for C. filicina. 

Cecidomyia filicina Kieffer 1889c: 191. France: Bitche & St Avold; ST; ♂,♀,L,egg; lost?. 

pudibunda. Nearctic (widespread eastern). Carpinus caroliniana (Betulaceae). 

Cecidomyia pudibunda Osten Sacken 1862: 202. USA: District of Columbia; T; LT; lost. 

pulsatillae. Sweden, Denmark, France, Germany, Latvia, Czechoslovakia, Romania. Pulsatilla vernalis, P. vulgaris 

(Ranunculaceae). 

Dichelomyia pulsatillae Kieffer 1894h: 297. France: Moselle; ST; LT; lost?. 

purpurea. USA (New York, Massachusetts, Virginia). Eupatoriadelphus sp. (Asteraceae). 

Dasyneura purpurea Felt 1908b: 349. USA: New York, New York, Staten I.; HT; ♀; NYSM. 

pustulans. Widespread Europe. Spiraea ulmaria (Rosaceae). 

Cecidomyia pustulans Rübsaamen 1889b: 378. Germany: vic. Sieghütte in Siegen region; ST; ♂,♀,L; ZMHB. Ref.: 

Harris 2010a (gall). 

*pyrethridis. Russia. Pyrethrum corymbosum (Asteraceae). 

Dasineura pyrethridis Fedotova 2008: 133. Russia: Central Povolzhye, Zhigulevsk Nature Res., nr Bakhilova 


pyri. Widespread Europe; immigr: Nearctic (eastern), New Zealand. Pyrus communis (Rosaceae). 

Cecidomyia pyri Bouché 1847: 144. Germany; ST; ♂,♀,P,L; ZMHB?. 


Dasyneura quercina Felt 1907a: 20. USA: New York, Albany; HT; ♂; NYSM. 

ranunculi. Widespread Europe, Russia (Far East). Ranunculus bulbosus (Ranunculaceae). 

Cecidomyia ranunculi Bremi 1847a: 54. Switzerland: Zurich; ST; ♂,♀,L; ETHZ?. 

rapunculi. Germany. Campanula rapunculi (Campanulaceae). 

Perrisia rapunculi Kieffer 1906d: 347. Germany; ST; ♂,P,L; lost?. 

rhododendri. UK, France, Yugoslavia, Romania. Rhododendron ferrugineum (Ericaceae). 

Perrisia rhododendri Kieffer 1909b: 25. France; T; G; lost?. 

rhodophaga. Widespread Nearctic. Rosa sp. (Rosaceae).

204 

Neocerata rhodophaga Coquillett 1900a: 47. USA: District of Columbia; ST; ♂,♀; USNM. 

rhois. USA (New York, Maryland, Virginia). Rhus toxicodendron (Anacardiaceae). 

Cecidomyia rhois Coquillett 1895a: 348. USA: New York, Lebanon Springs; ST; ♂,♀; USNM. 

ribis. Sweden, Finland, Germany, Latvia, Poland, Bulgaria, Russia (Europe). Ribes nigrum; R. alpinum 

(Saxifragaceae). 

Dasyneura ribis Barnes 1940a: 85. Finland; ST; ♂,♀; BMNH. 

rileyana. USA (Iowa). Acer sp. (Aceraceae). 

Rhabdophaga rileyana Felt 1909b: 289. USA: Iowa, Ames; HT; ♀; USNM. 

rosae. Widespread Europe; Kazakhstan, Russia (Far East). Rosa canina; Rosa spp. (Rosaceae). 

Cecidomyia rosae Bremi 1847a: 27. Switzerland: vic. Zurich; ST; LT; ETHZ?. Bland (2000: 10) designated this 

species as a nomen oblitum and junior synonym of rosarum Hardy under Article 23.9, ICZN 1999, but Hardy 

(1854f) had synonymized his own name under rosae. Bland neglected to show that the conditions of Article 

29.9.2 were met. In addition, the name rosae Bremi was used in Gagné 1972b, so it was not a forgotten name. 

Cecidomyia rosarum Hardy 1850: 186. UK: Scotland, vic. Penmanshiel; T; ♀; lost. 

rosmarini. Portugal, Spain, France. Rosmarinus officinalis (Lamiaceae). 

Dasyneura rosmarini Tavares 1902b: 27. Portugal: S. Fiel, Setubal; ST; ♂,♀; TAVC. 

rossi. UK, Germany, Czechoslovakia, Ukraine, Russia (Western Siberia). Astragalus danicus (Fabaceae). 

Dasyneura rossi Rübsaamen 1914: 106. Germany: Gerolzhofen; ST; ♂,♀; ZMHB. 

rostratae. Germany. Sisymbrium loeselii (Brassicaceae). 

Dasineura rostratae Stelter 1992e: 201. Germany: Berlin-Baumsschulenweg, vic. railway stn.; HT; ♂; SMTD. 

rozhkovi. Russia (Western Siberia), Kazakhstan. Larix sp. (Pinaceae). 

Dasineura rozhkovi Mamaev & Nikolskij 1983: 109. Russia: Krasnoyarsk, Akademgorodok; HT; ♂; ZMMU. 

rubella. Widespread Europe. Acer campestre (Aceraceae). 

Dasyneura rubella, as var. of acercrispans, Kieffer 1896h: 37. France; T; ♀; lost?. 

rubiflorae. USA (Virginia). Rubus sp. (Rosaceae). 

Dasyneura rubiflorae Felt 1908b: 343. USA: Virginia, S; ♂,♀; USNM. 

*rubiformis. Australia (Western Australia), South Africa (introd.). Acacia mearnsii (Fabaceae). 

Dasineura rubiformis Kolesik 2005: 460. Australia: Western Australia, 5 km E Albany; HT; ♂; SAMA. 

ruebsaameni. UK, Netherlands, Germany, Czechoslovakia, Austria, Italy, Yugoslavia, Romania; Pliocene 

fossil/Germany. Carpinus betulus (Betulaceae). 

Perrisia ruebsaameni (as rübsaameni) Kieffer 1909b: 6. Germany; T; G; ZMHB?. 

rufescens. Portugal, Italy, Yugoslavia, Bulgaria, Greece, Turkey, Algeria. Phyllirea variabilis (Rhamnaceae). 

Perrisia rufescens De Stefani 1898b: 9. Italy: Sicily, nr Palermo; ST; ♂,♀,P,L; lost. 

rufipedalis. USA (New York). 

Dasyneura rufipedalis Felt 1908b: 349. USA: New York, Karner; HT; ♀; NYSM. 

rumicicola. Germany. Rumex acetosella (Polygonaceae). 

Dasyneura rumicicola Rübsaamen 1914: 108. Germany: betw. Oberwesel & Engewohl; ST; ♀,L; ZMHB. 

rutifolia. Kazakhstan. Artemisia rutifolia (Asteraceae). 

Dasineura rutifolia Fedotova 2001h: 1481. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 18 km SE 


salicifolia. USA (New York, Massachusetts). Salix sp. (Salicaceae). 

Dasyneura salicifolia Felt 1907d: 12. USA: New York, Albany; HT; ♂; NYSM. 

salicifoliae. Widespread Nearctic. Spiraea salicifolia (Rosaceae). 

Cecidomyia salicifoliae Osten Sacken 1866: 220. Canada: Quebec; ST; G; USNM. 

Rhabdophaga salicifolia Felt 1907d: 12. USA: New York, Albany; ST; ♂,♀; NYSM. 

salviae. Widespread Europe. Salvia pratensis (Lamiaceae). 

Perrisia salviae Kieffer 1909b: 27. France; ST; LT; lost?. 

sampaina. UK, Portugal, France, Italy, Germany, Czechoslovakia, Hungary, Algeria. Linum usitatissimum; Linum spp. 

(Linaceae). 

Perrisia sampaina Tavares 1902b: 24. Portugal: Porto; ST; ♀,P,egg; TAVC. 

sanguisorbae. UK, Germany, Lithuania, Poland, Czechoslovakia, Hungary, Russia (Europe). Sanguisorba officinalis 

(Rosaceae).

205 

Cecidomyia sanguisorbae Rübsaamen 1890a: 26. Germany: Weidenau, vic. Münkershütten; ST; ♂,♀,L; ZMHB. 

sassafras. Canada (Ontario). Sassafras albidum (Lauraceae). 

Dasyneura sassafras Felt 1916a: 29. Canada: Ontario, Gordon; HT; ♀; NYSM. 

saurica. Kazakhstan. Campanula wolgensis (Campanulaceae). 

Dasineura saurica Fedotova 1993l: 82. Kazakhstan: Saur Mtn Range, 20 km S Zaïsan, nr Churchutsu; HT; ♂; 

ZMAS. 

saussureae. Sweden, Kazakhstan. Saussurea latifolia; S. alpina (Asteraceae). 

Dasineura saussureae Fedotova 1996b: 192. Kazakhstan: Altaï, Ivanovskiï Mts, 32 km NE Leninogorsk, nr 

Poperechnoe; HT; ♂; ZMAS. 

saxifragae. UK, Sweden, France, Germany, Czechoslovakia. Saxifraga granulata (Saxifragaceae). 

Cecidomyia saxifragae Kieffer 1891d: 210. France: Bitche; ST; G; lost?. 

schulzei. Germany, Poland, Czechoslovakia, Romania, Yugoslavia. Euphorbia palustris; E. helioscopia, E. seguierana 

(Euphorbiaceae). 

Dasyneura schulzei Rübsaamen 1917: 55. Germany: Havelberg; LT (Gagné 1990b: 342); ♂; ZMHB. 

scirpi. France. Scirpus sylvaticus (Cyperaceae). 

Dasyneura scirpi Kieffer 1898f: 58. France: Moselle; ST; ♂,♀; lost?. 

scorpii. France. Genista scorpius (Fabaceae). 

Perrisia scorpii Kieffer 1909b: 9. France; T; G; lost?. 

Perrisia rosifex Kieffer 1909b: 9. France; T; G; lost?. 

scorzonerifloris. Kazakhstan. Scorzonera parviflora (Asteraceae). 

Dasineura scorzonerifloris Fedotova 1996d: 8. Kazakhstan: Tarbagataï Mtn Range, 10 km S Ayaguz Mtn; HT; ♂; 

ZMAS. 

scutata. USA (New York). 

Dasyneura scutata Felt 1908b: 346. USA: New York, Albany; HT; ♀; NYSM. 

sedicola. Russia (Far East). Sedum aizoon (Crassulaceae). 

Dasyneura sedicola Kovalev 1967: 112. Russia: Primorskiï Terr., Popova I., Petra Velikoe Bay, nr Vladivostok; HT; 

♂; KOVC. 

semenivora. Canada (Ontario), USA (Massachusetts, New Jersey). Viola spp. (Violaceae). 

Cecidomyia semenivora Beutenmüller 1907a: 390. USA: New Jersey, Plainfield; T; LT; AMNH. 

senebrierae. Egypt. Senebiera nilotica (Brassicaceae). 

Perrisia senebrierae Kieffer 1909b: 28. Egypt: on route to Pyramids of Gizeh; T; G; NHMW?. 

seneciocola. Kazakhstan. Senecio praticola (Asteraceae). 

Dasineura seneciocola Fedotova 1998a: 34. Kazakhstan: southern Altaï, Markakol'skiï Res., L. Markakol', nr 


senecioflora. Kazakhstan. Senecio nemorensis (Asteraceae). 

Dasineura senecioflora Fedotova 1993l: 78. Kazakhstan: Altaï, Ivanovskiï Mts, 32 km NE Leninogorsk; HT; ♂; 

ZMAS. 

senecionis. Germany. Senecio sp. (Asteraceae). 

Dasyneura senecionis Rübsaamen 1925: 68. Germany; T; P; ZMHB. 

serotina. Widespread Europe. Hypericum humifusum (Hypericaceae). 

Cecidomyia serotina Winnertz 1853: 316. Germany; ST; ♂,♀,L; MNHN?. 

serrulatae. Canada (Ontario), USA (Massachusetts, North Carolina). Alnus serrulata (Betulaceae). 

Cecidomyia serrulatae Osten Sacken 1862: 198. USA: District of Columbia; ST; ♂,♀,L; MCZC. 

sesami. India (Uttar Pradesh). Sesamum orientale (Pedaliaceae). 

Dasineura sesami Grover & Prasad 1966c: 119. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

Dasyneura gingellii Grover & Prasad 1966d: 357, nomen nudum. 

severzovi. Kazakhstan. Polygonatum severzovi (Liliaceae). 

Dasineura severzovi Fedotova 1996d: 10. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 12 km SE 


shinjii. Japan (Honshu). Vicia pseudoorobus (Fabaceae). 

Dasyneura vicicola Shinji 1939e: 99, jun. sec. hom. Perrisia vicicola Tavares. Japan: Honshu, Morioka; T; A; lost. 

Dasineura shinjii Skuhravá 1986: 164, new name for Dasyneura vicicola Shinji.

206 

sibirica. Russia (Western Siberia). Caragana arborescens (Fabaceae). 

Dasyneura sibirica Marikovskij 1962: 21. Russia: vic. Tomsk, Novosibirsk, & Klyuchevskaya; ST; ♂,♀; MARC. 

silvestris. UK, France, Poland, Czechoslovakia, Romania, Bulgaria, Russia (Western Siberia). Lathyrus sylvestris 

(Fabaceae). 

Perrisia silvestris Kieffer 1909b: 13. France; ST; LT; lost?. 

silvicola. UK, Germany. Stellaria holostea (Caryophyllaceae). 

Perrisia silvicola Kieffer 1909b: 30. Germany: Kreuznach; ST; LT; lost?. 

similis. Widespread Europe. Veronica scutellata; V. anagallis-aquatica (Scrophulariaceae). 

Cecidomyia similis Löw 1888: 232. France: vic. Bitche; ST; ♂,♀,L; NHMW. 

simillima. USA (New York). 

Dasyneura similis Felt 1908b: 346, jun. sec. hom. similis Löw. USA: New York, Albany; HT; ♂; NYSM. 

Perrisia simillima Kieffer 1913i: 81, new name for Dasyneura similis Felt. 

sisymbrii. Widespread Europe, Kazakhstan. Rorippa palustris; Barbarea vulgaris, Sisymbrium zannonii 

(Brassicaceae). 

Tipula sisymbrii Schrank 1803: 83. Germany: Bavaria; ST; A,L; lost?. 

Cecidomyia barbareae Curtis 1845: 400. UK: England; ST; ♀,L; ?. 

Cecidomyia sophiae Loew 1850a: 29. Poland; ST; G; ZMHB?. 

Perrisia fiorii Cecconi 1904: 184. Italy: Vallombrosa, Monte Secchieta; ST; ♂,♀,P,L; ?. 

Perrisia zannonii Kieffer 1909b: 28. Italy; ST; LT; lost?. 

Dasineura zannonnii Skuhravá 1986: 164, missp. of zannonii. 

smilacifolia. USA (Massachusetts, Maryland). Smilax sp. (Smilacaceae). 

Dasyneura smilacifolia Felt 1911m: 480. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

smilacinae. Canada (Ontario), USA (New York). Smilacina racemosa (Liliaceae). 

Dasyneura smilacinae Bishop 1911: 346. USA: New York, Ithaca; ST; ♂,♀; ?. 

socialis. UK, France, Poland. Erigeron acris (Asteraceae). 

Perrisia socialis Kieffer 1909b: 7. France; ST; LT; lost?. 

sodalis. Widespread Europe. Prunus spinosa; Prunus spp. (Rosaceae). 

Cecidomyia sodalis Löw 1877: 7. Austria: vic. Klosterneuberg, nr Vienna; ST; ♂,♀,L; NHMW. 

solonaica. Kazakhstan. Lonicera altaica (Caprifoliaceae). 

Dasineura solonaica Fedotova 1996f: 52. Kazakhstan: southern Altaï, Solonoï R. valley, nr Urunkhaïka, 60 km N 


soongarica. Kazakhstan. Barbarea vulgaris (Brassicaceae). 

Dasineura soongarica Fedotova 1994a: 115. Kazakhstan: Dzhungarskiï Alatau, Altyn-Emel' pass, 5 km S 

Golubinovka; HT; ♂; ZMAS. 

spadicea. Widespread Europe, Russia (Western Siberia). Vicia cracca (Fabaceae). 

Dasyneura spadicea Rübsaamen 1917: 45. Germany: Kreuznach, Hamburg, & Rhine Prov.; ST; ♂,♀,L; ZMHB. 

spiraeae. UK, Sweden, France. Spiraea ulmaria (Rosaceae). 

Perrisia spiraeae Loiselle 1912: 26. France: Lisieux; ST; ♂,♀; ?. 

Dasineura spireae Skuhravá 1986: 165, missp. of spiraeae. 

spiraeina. USA (New York). 

Dasyneura spiraeina Felt 1908b: 341. USA: New York, Karner; HT; ♀; NYSM. 

squamosa. UK, Spain. Quercus robur, Q. lusitanica (Fagaceae). 

Perrisia squamosa Tavares 1919b: 63. Spain: Gallicia, Carballino; ST; ♂,♀; TAVC. 

stanleyae. USA (Colorado). Stanleya glauca (Brassicaceae). 

Perrisia stanleyae Cockerell 1914b: 241. USA: Colorado, 4 mi. N Boulder; ST; ♂,L; USNM. 

stellariae. Denmark, France, Germany, Poland, Czechoslovakia, Romania. Stellaria holostea (Caryophyllaceae). 

Dasyneura stellariae Rübsaamen 1916b: 514. Germany: vic. Sattenfeld nr Oldesloe; ST; ♂,♀,L; ZMHB. 

stelteri. Germany. Barbarea vulgaris (Brassicaceae). 

Dasineura barbareae Stelter 1992e: 199, jun. sec. hom. of barbareae Curtis. Germany, Kr. Rostock, Gross Lüsewitz; 

HT; ♂; SMTD. 

Dasineura stelteri Gagné 2004b: 140, new name for Dasineura barbareae Stelter. 

*storozhenkoi. Russia (Far East). on Vicia cracca (Fabaceae).

207 

Dasineura storozhenkoi Fedotova 2003c: 24. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

strumosa. Widespread Europe. Galeobdolon luteum (Lamiaceae). 

Cecidomyia strumosa Bremi 1847a: 25. Switzerland: nr Hottingen; ST; LT; ETHZ?. 

Cecidomyia galeobdolontis Winnertz 1853: 238. Germany; ST; ♀,L; lost. 

subinermis. Chile. Baccharis rosmarinifolia (Asteraceae). 

Perrisia subinermis Kieffer & Herbst 1911: 696. Chile: Valparaíso; ST; ♂,♀,P; lost. 

subterranea, France. Silene inflata (Caryophyllaceae). 

Perrisia subterranea Kieffer 1909b: 28. France; T; G; lost?. 

*sulcata. Australia (Western Australia?, New South Wales). Acacia saligna, Acacia spp. (Fabaceae). 

Dasineura sulcata Kolesik 2005: 465. Australia: New South Wales: Wollongong; HT; ♂; SAMA. 

swainei. Canada (Alberta to Newfoundland, New Brunswick). Picea sp. (Pinaceae). 

Rhabdophaga swainei Felt 1914h: 77. Canada: Ontario, Ottawa; ST; ♂,♀; NYSM. 

symphyti. UK, Netherlands, Germany, Poland, Czechoslovakia, Hungary, Romania, Yugoslavia. Sisymbrium officinale 

(Brassicaceae). 

Cecidomyia symphyti Rübsaamen 1892b: 399. Germany; ST; ♂,♀,L; ZMHB. 

syreniae. Kazakhstan. Syrenia siliculosa, S. sessiliflora (Brassicaceae). 

Dasineura syreniae Fedotova 1994a: 117. Kazakhstan: Ustyurt plateau, Sam Desert, Akshi, 12 km E Sam; HT; ♂; 

ZMAS. 

szepligetii. Hungary. Campanula rapunculoides (Campanulaceae). 

Perrisia szepligetii Kieffer 1909b: 4. Hungary; T; LT; lost?. 

tamaricicarpa. Kazakhstan. Tamarix ewersmannii (Tamaricaceae). 

Dasyneura tamaricicarpa Fedotova 1983a: 39. Kazakhstan: Ili R. delta, nr Karoi; HT; ♂; ZKAS. 

tamaricicola. Kazakhstan. Tamarix ewersmannii (Tamaricaceae). 

Dasyneura tamaricicola Fedotova 1983a: 37. Kazakhstan: Ili R. delta, nr Karoi; HT; ♂; ZKAS. 

tamariciflora. Kazakhstan. Tamarix litvinovi; T. hispida (Tamaricaceae). 

Dasyneura tamaricifolia Fedotova 1983a: 36. Southern Kazakhstan: 20 km W Abai; HT; ♂; ZMAS. 

tamaricina. Libya, Egypt. Tamarix africana (Tamaricaceae). 

Perrisia tamaricina Kieffer 1909b: 30. Egypt: Oasis Tarfa; T; G; NHMW?. 

tanaitica. Kazakhstan. Onobrychis tanaitica (Fabaceae). 

Dasineura tanaitica Fedotova 1992g: 50. Kazakhstan: Kungeï Alatau, nr Karabulak; HT; ♂; ZMAS. 

tavaresi. Brazil (Rio de Janeiro). Neomitranthes obscura (Myrtaceae). 

Dasineura tavaresi Maia 1996b: 1006. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♀; QBUM. 

*tavolga. Russia (Far East). Spiraea salicifolia (Rosaceae). 

Dasineura tavolga Fedotova 2002c: 2. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

tetensi. Widespread Europe. Ribes nigrum (Saxifragaceae). 

Cecidomyia tetensi Rübsaamen 1892b: 400. Germany: Berlin; ST; ♂,♀,L; ZMHB. 

Perrisia ribicola Kieffer 1909b: 25. Germany: nr Ohrdruf; T; G; NHMW?. 

tetragynae. Egypt. Tamarix tetragyna (Tamaricaceae). 

Dasyneura tetragynae Debski 1918: 28. Egypt: Hélouan; ST; LT; ?. 

tetrahit. UK, Netherlands, France, Czechoslovakia. Galeopsis tetrahit (Lamiaceae). 

Perrisia tetrahit Kieffer 1909b: 8. France; ST; LT; lost?. 

tetrastigma. Indonesia (Java). Tetrastigma pergamaceum (Vitaceae). 

Dasyneura tetrastigma Felt 1927d: 385. Indonesia: Java, Tjibodas, Mt Gede; ST; ♀; NYSM. 

teucrii. UK, Portugal, Spain, France, Germany, Czechoslovakia, Yugoslavia. Teucrium lusitanicum, T. scorodonia; T. 

chamaedrys (Lamiaceae). 

Perrisia teucrii Tavares 1903: 178. Portugal: Matta do Fundão e Rodão, Nine, Gerez, vic. Braga, & Serra da Estrella; 

ST; ♂,♀,P; TAVC. 

Perrisia teucriicola Kieffer 1909b: 30. France; ST; LT; lost?. 

*thalictrifoliae. Kazakhstan. Thalictrum collinum (Ranunculaceae). 

Dasineura thalictrifoliae Fedotova 1992n: 104. Kazakhstan: Aigyr Kum Desert, 50 km NE Zaisan; HT; ♂; ZMAS. 

*theobromae. Peru. Theobroma bicolor (Sterculiaceae).

208 

Dasineura theobromae Maia 2006b: 126. Peru: Loreto, Iquitos; HT; ♂; MNRJ. 

thlaspicarpae. Kazakhstan. Thlaspi arvense (Brassicaceae). 

Dasineura thlaspicarpae Fedotova 1990c: 23. Kazakhstan: Tarbagataï Mtn Range, 30 km SW Ayaguza, Baskan R. 

floodplain; HT; ♂; ZMAS. 

thomasi. France, Germany, Switzerland, Italy, Poland, Romania. Campanula pusilla, C. scheuchzeri (Campanulaceae). 

Cecidomyia thomasi Kieffer 1909b: 4. Austria: Tyrol; ST; LT; NHMW?. 

Dasyneura thomasi Rübsaamen 1912f: 354. Switzerland: Bernese Alps, nr Wengen; ST; ♂,♀; ZMHB. 

thomasiana. Widespread Europe. Tilia parvifolia, T. grandifolia (Tiliaceae). 

Cecidomyia thomasiana Kieffer 1888e: 95. France: Falkenstein ruins nr Bitche; ST; ♂,♀,L; lost?. 

Perrisia tiliae Kieffer 1909b: 30, jun. sec. hom. Cecidomyia tiliae Schrank. Italy; ST; LT; lost?. 

Perrisia tiliarum Kieffer 1913i: 82, new name for Perrisia tiliae Kieffer. 

tiliae. Widespread Europe, Russia (Far East). Tilia sp.; T. cordata, T. platyphyllos (Tiliaceae). 

Tipula tiliae Schrank 1803: 87. Germany: Bavaria; ST; LT; lost?. 

Cecidomyia tiliamvolvens Rübsaamen 1889a: 55. Germany: Siegen; ST; ♀,L; ZMHB. 

*tjanshanica. Kazakhstan. Galatella punctata (Asteraceae). 

Dasineura tjanshanica Fedotova 2003a: 830. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE Alma-Ata; 


torontoensis. Canada (Ontario). Maianthemum canadense (Liliaceae). 

Dasyneura torontoensis Felt 1915f: 405. Canada: Ontario, Toronto; ST; ♀,L; NYSM. 

tortilis. Widespread Europe. Alnus incana; A. glutinosa (Betulaceae). 

Cecidomyia tortilis Bremi 1847a: 29. Switzerland: vic. Zurich; ST; LT; ETHZ?. 

Cecidomyia alni Löw 1877: 2. Austria: Vienna; ST; ♀,L; NHMW. 

tortrix. Widespread Europe. Prunus spinosa (Rosaceae). 

Cecidomyia tortrix Löw 1877: 6. Austria: vic. Klosterneuberg nr Vienna; ST; ♂,♀,L; NHMW. 

toweri. USA (Massachusetts). Hypericum mutilum (Hypericaceae). 

Dasyneura toweri Felt 1909b: 289. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

tragopogonicola. Kazakhstan. Tragopogon scoparius (Asteraceae). 

Dasineura tragopogonicola Fedotova 1995e: 18. Kazakhstan: Tarbagataï Mtn Range, 46 km E Ayaguz, upper 

Ayaguz R.; HT; ♂; ZMAS. 

traili. UK, Sweden, Netherlands, Czechoslovakia, Romania, Yugoslavia. Ranunculus acris, R. repens 


Perrisia traili Kieffer 1909b: 25. UK: Scotland; T; G; lost?. 

trifolii. Widespread Europe, Algeria, Russia (Western Siberia); immigr.: USA (Nebraska, Maryland, Massachusetts). 

Trifolium pratense (Fabaceae). 

Cecidomyia trifolii Löw 1874a: 143. Austria: Vienna; ST; ♂,♀,P,L; NHMW. 

tripolii. Romania. Aster tripolium (Asteraceae). 

Dasyneura tripolii Neacsu 1968: 901. Romania: Dobroudja, Mamaia; HT; ♂; NEAC. 

triseti. Austria. Trisetum flavescens (Poaceae). 

Dasyneura triseti Barnes 1939e: 174. Austria: Tyrol, Leutasch; ST; ♂,♀; BMNH. 

trotteri. UK, Portugal, Spain. Cytisus albus, Sarothamnus scoparius (Fabaceae). 

Perrisia trotteri Tavares 1902c: 184. Portugal: Guarda, Estrella, Marváo, & Castro Laboreiro; ST; ♂,♀,L; TAVC. 

tubicoloides. UK, Netherlands, France, Germany, Poland, Greece. Sarothamnus scoparius (Fabaceae). 

Cecidomyia tubicola Kieffer 1889c: 188, jun. prim. hom. tubicola Osten Sacken. France: vic. Bitche; ST; ♂,♀,L; 

lost?. 

Dasineura tubicoloides Gagné 2004b: 142, new name for Cecidomyia tubicola Kieffer. 

tubularis. Czechoslovakia, Austria, Hungary, Romania, Yugoslavia, Greece. Quercus cerris (Fagaceae). 

Perrisia tubularis Kieffer 1909b: 22. Austria; T; G; NHMW?. 

tumidosae. Canada (Ontario), USA (Michigan, New York, Maine, Massachusetts, Connecticut, Maryland); immigr.: 

USA (California). Fraxinus sp. (Oleaceae). 

Dasyneura tumidosae Felt 1908b: 346. USA: New York, Bath & Highland; ST; ♂,♀; NYSM. 

turgenica. Kazakhstan. Astragalus alpinus (Fabaceae). 

Dasineura turgenica Fedotova 1993j: 86. Kazakhstan: Zailiïskiï Alatau, Turgen' gorge, 40 km S Turgen'; HT; ♂;

209 

ZMAS. 

turionum. France, Italy, Yugoslavia, Greece, Algeria. Asparagus officinalis (Liliaceae). 

Perrisia turionum Kieffer & Trotter 1904a: 7. Italy: Avellino Prov., Valle dell'Ofanto nr Lioni; ST; ♂,♀,L; lost?. 

tympani. Widespread Europe. Acer campestre, Acer spp. (Aceraceae). 

Perrisia tympani Kieffer 1909b: 1. France; T; G; lost?. 

ulicis. UK, France. Ulex europaeus (Fabaceae). 

Perrisia ulicis Kieffer 1909b: 31. France; T; G; lost?. 

ulmaria. Widespread Europe, Russia (Far East). Spiraea ulmaria (Rosaceae). 

Cecidomyia ulmaria Bremi 1847a: 52. Switzerland: vic. Zurich; ST; ♂,♀,L; ETHZ?. Ref.: Harris 2010a (gall). 

ulmea. USA (Illinois, New York, Massachusetts, Maryland). Ulmus sp. (Ulmaceae). 

Dasyneura ulmea Felt 1908b: 349. USA: vic. District of Columbia; ST; ♀; USNM. 

ulmicola. UK, Sweden, France, Germany. Ulmus minor (Ulmaceae). 

Perrisia ulmicola Kieffer 1909b: 31. France; T; LT; lost?. 

umbrosa. Austria. Cardamine flexuosa (Brassicaceae). 

Perrisia umbrosa Kieffer 1909b: 4. Austria: Thüringen; T; G; NHMW?. 

urnicola. USA (New York). Urtica dioica (Urticaceae). 

Cecidomyia urnicola Osten Sacken 1875: 202. USA: New York, Lake George & Trenton Falls; ST; G; lost. 

urticae. Widespread Europe, Russia (Far East). Urtica dioica (Urticaceae). 

Cecidomyia urticae Perris 1840: 403. France: Landes, vic. Mont de Marsan; ST; ♂,♀,L; ?. 

vagans. Italy, Greece. Erica vagans (Ericaceae). 

Perrisia vagans Kieffer 1909b: 7. Italy; T; G; lost?. 

valerianae. Georgia. Valeriana alliariaefolia (Valerianaceae). 

Perrisia valerianae Kieffer 1909b: 31. Georgia: nr Kisslowodsk; T; G; ZMHB?. 

vallisumbrosae. UK, Italy, Yugoslavia, Romania. Sarothamnus scoparius (Fabaceae). 

Perrisia vallisumbrosae Kieffer 1904d: 91. Italy: Vallombrosa; ST; ♂,♀,P,L; lost?. 

verbasci. France, Austria, Romania, Yugoslavia. Verbascum lynchitis (Scrophulariaceae). 

Perrisia verbasci Kieffer 1909b: 31. France; ST; LT; lost?. 

vernalis. USA (New York). 

Dasyneura vernalis Felt 1908b: 341. USA: New York, Nassau; HT; ♀; NYSM. 

verrucosa. Eritrea. Olea chrysophylla (Oleaceae). 

Perrisia verrucosa Del Guercio 1918a: 53. Eritrea; T; LT; lost. 

viciae. Widespread Europe, Russia (Western Siberia). Vicia sepium (Fabaceae). 

Cecidomyia viciae Kieffer 1888e: 105. France: Moselle; ST; ♂,♀,L; lost?. 

vicicola. Portugal. Vicia angustifolia bobartii (Fabaceae). 

Perrisia vicicola Tavares 1905b: 260. Portugal; ST; ♂,♀; TAVC. 

vincae. Italy, Hungary, Romania, Yugoslavia. Vinca major (Apocynaceae). 

Perrisia vincae Kieffer & Trotter 1904b: 64. Italy: vic. Avellino; ST; ♂,P,L; lost?. 

violae. Widespread Europe. Viola tricolor (Violaceae). 

Cecidomyia violae Löw 1880: 34. Austria: nr Payerbach & nr Baden; ST; ♂,♀,L; NHMW. 

violahirtae. Denmark, Germany, Hungary. Viola hirta (Violaceae). 

Dasyneura violaehirtae Buhr 1965: 1357, nomen nudum. 

Dasineura violahirtae Stelter 1982c: 264. Germany: Kyffhäuser, Barbarossahöhle; HT; ♂; SMTD. 

Dasineura violaehirtae Skuhravá 1986: 168, missp. of violahirtae. 

virgaeaureae. Widespread Europe. Solidago virgaurea (Asteraceae). 

Cecidomyia virgaeaureae (as virgae aureae) Liebel 1889b: 283. France: Boulay; ST; ♀,L; ?. 

vitis. USA (New York). Vitis sp. (Vitaceae). 

Dasyneura vitis Felt 1908b: 341. USA: New York, West Nyack; HT; ♀; NYSM. 

vitisidaea. UK, Czechoslovakia, Italy, Austria. Vaccinium vitis-idaea (Ericaceae). 

Perrisia vitisidaea (as vitis idaea) Kieffer 1909b: 31. Austria; ST; LT; NHMW?. 

volantis, USA (New York). 

Dasyneura acerifoliae Felt 1907a: 18, jun. sec. hom. Rhabdophaga acerifoliae Felt. USA: New York, Albany; HT; 

♂; NYSM.

210 

Dasyneura acerifolia Felt 1907c: 115, unjust. emend. of acerifoliae. 

Dasineura volantis Gagné 2004b: 143, new name for Dasyneura acerifoliae Felt. 

vulgatiformiae. Sweden. Hieracium sp., vulgatiformia group (Asteraceae). 

Dasineura vulgatiformiae Sylvén 1998b: 248. Sweden: Öland, Böda Crown For., Fiskåsen; HT; ♂; NHRS. 

wahlenbergiae. Australia (South Australia). Wahlenbergia stricta (Campanulaceae). 

Dasineura wahlenbergiae Kolesik 1998c: 147. Australia: South Australia, Morialta Conservation Pk, 13 km NE 


weigelaefolia. Russia (Far East). Weigela praecox (Caprifoliaceae). 

Dasyneura weigelaefolia Kovalev 1967: 114. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♀; KOVC. 

wistariae. Japan (Honshu, Kyushu). Wisteria floribunda (Fabaceae). 

Dasyneura wistariae Mani 1954c: 9. Japan: Kyushu, Fukuoka; HT; ♂; ZSCI. 

Dasineura wisteriae Yukawa 1971: 105, missp. of wistariae. 

xylostei. Germany, Czechoslovakia. Lonicera xylosteum (Caprifoliaceae). 

Perrisia xylostei Kieffer 1909b: 14. Germany; T; G; ZMHB?. 

zillae. Egypt. Zilla myagoides (Brassicaceae). 

Perrisia zillae Kieffer 1909b: 34. Egypt: both shores of Red Sea; T; G; NHMW?. 

zimmermanni. Portugal, Spain, Yugoslavia. Erica arborea (Ericaceae). 

Perrisia zimmermanni Tavares 1901: 60. Portugal: Perto da Covilhan & Gardunha; ST; ♂,♀,P; TAVC. 

DASYNEURIOLA 

Dasyneuriola Marikovskij 1955b: 104, nomen nudum. 

Dasyneuriola Marikovskij 1961: 42. Type species, suaedae Marikovskij (orig. des.). 

Palearctic; 5 spp.; Lasiopteridi, Dasineurini. This genus was separated from Dasineura by characters that appear 

superficial and do not exclude some Dasineura spp. With the exception of D. tamariciphila from Tamarix, which does 

not appear to have a particular affinity to the remaining four species, all are from Suaeda (Chenopodiaceae) and may 

form a natural group, possibly with the addition of species now in Dasineura that are also found on Chenopodiaceae. 

salicorniae. Armenia, Kazakhstan. Suaeda altissima; Suaeda spp. (Chenopodiaceae) (Host orig. given as Salicornia 

europaea but changed to Suaeda in Fedotova 1995d: 706). 

Janetiella salicorniae (also as salicornidae) Fedotova 1984d: 297. Kazakhstan: north shore L. Balkash, Baïgabyl; 


suaedae. Kazakhstan, Turkmenistan. Suaeda microphylla (Chenopodiaceae). 

Dasyneuriola suaeda Marikovskij 1955b: 104. Nomen nudum. 

Dasyneuriola suaedae Marikovskij 1961: 43. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, vic. middle & lower 

reaches Ili R.; ST; ♂,♀; ZMAS. 

suaedigemmae. Kazakhstan. Suaeda acuminata (Chenopodiaceae). 

Dasyneuriola suaedigemmae Fedotova 1995d: 707. Kazakhstan: Dzhungarskiï Alatau, 3 km W Konurolen, nr 

Alatyn-Emel' pass; HT; ♂; ZMAS. 

suaediramea. Kazakhstan. Suaeda physophora (Chenopodiaceae). 

Dasyneuriola suaediramea Fedotova 1995d: 708. Kazakhstan: upper course Ili R., left bank, 3 km above 

Poyushchaya Gora, 45 km NE Chilik; HT; ♂; ZMAS. 

tamariciphila. Kazakhstan. Tamarix sp. (Tamaricaceae). 

Dasyneuriola tamariciphila Marikovskij 1961: 43. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, vic. middle & 

lower reaches Ili R.; HT; ♂; ZMAS. 

DELODIPLOSIS 

Delodiplosis Tavares 1908: 157. Type species, copaibae Tavares (mon.). 

Afrotropical; 1 sp.; Cecidomyiidi, unplaced to tribe.

copaibae. Mozambique. Copaiba mopane (Fabaceae). 

Delodiplosis copaibae Tavares 1908: 158. Mozambique; ST; ♂,♀; TAVC. 

DENTIFIBULA 

Dentifibula Felt 1908b: 385, 389. Type species, Cecidomyia viburni Felt (orig. des.). 

Muirodiplosis Grover 1965b: 111. Type species, spinosa Grover (orig. des.). 

Holarctic, Oriental; 8 spp., the 2 with known hosts feed on scale insects; Cecidomyiidi, Lestodiplosini. 

*hastata. Russia (Far East). 

Dentifibula hastata Fedotova & Sidorenko 2005a: 214. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. 


*magna. China (Zhejiang). 

Dentifibula magna Mo & Liu 2003: 51. China: Zhejiang, Anji Co.; HT; ♂; SAUT. 

*marikovskajae. Russia (Far East). 

Dentifibula marikovskajae Fedotova & Sidorenko 2004b: 98. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


nigritarsis. China (Shandong). 

Dentifibula nigritarsis Mo 1992: 293. China: Shandong, Taian; HT; ♂; SAUT. 

obtusilobae. Sri Lanka. Pinnaspis aspidistrae, Hemiberlesia lataniae (Hemiptera: Diaspididae). 

Dentifibula obtusilobae Felt 1915g: 176. Sri Lanka: Peradeniya, Botanic Gardens; ST; ♂,♀; NYSM. 

Dentifibula ceylanica Felt 1915g: 175. Sri Lanka: Peradeniya, Botanic Gardens; HT; ♂; NYSM. 

spinosa. India (Uttar Pradesh). 

Muirodiplosis spinosa Grover 1965b: 112. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

turkmenorum. Turkmenistan. 

Dentifibula turkmenorum Mamaev 1986: 65. Turkmenistan: Kopetdag, Ipaï-Kala; HT; ♂; HYNM. 

viburni. USA (Illinois, New York, Maryland, Florida), Italy. Orig. unkn.; Diaspidiotus uvae, Pseudaulacapis spp., 

Targiona vitis (Hemiptera: Diaspididae). 

Contarinia viburni Felt 1907a: 36. USA: New York, Albany; HT; ♂; NYSM. 

Contarinia caryae Felt 1907a: 36. USA: New York, Albany; HT; ♂; NYSM. 

Dentifibula cocci Felt 1908b: 389. USA: Illinois, HT; ♂; USNM. 

*DEXTERMANA 

Dextermana Fedotova & Sidorenko 2009d: 253. Type species, Tuguridiplosis cordata Fedotova & Sidorenko (orig. 

des.). 

Eastern Palearctic; 1 spp.; Cecidomyiidi, Clinodiplosini. This genus is close to Clinodiplosis if the two must be 

separated. 

*cordata. Russia (Far East). 

Tuguridiplosis cordata Fedotova 2005c: 219. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. Kamenushka; HT; 

♂; ZMAS. 

DIADIPLOSIS 

Diadiplosis Felt 1911c: 54. Type species, cocci Felt (orig. des.). 

Coccodiplosis Meijere 1917: 238. Type species, pseudococci Meijere (mon.) = smithi Felt. 

Cleodiplosis Felt 1922d: 1. Type species, aleyrodici Felt (orig. des.). 

Ghesquierinia Barnes 1939c: 327. Type species, megalamellae Barnes (orig. des.). 

211

212 

Kalodiplosis Felt 1915e: 229. Type species, Dicrodiplosis multifila Felt (orig. des.). 

Schizobremia Felt 1926d: 183. Type species, formosana Felt (orig. des.). 

Olesicoccus Borgmeier 1931: 186. Type species, costalimai Borgmeier (orig. des.) = coccidivora Felt. 

Cieodiplosis Shinji 1944: 232, missp. of Cleodiplosis. 

Nipponodiplosis Harris 1968: 458. Type species, Diadiplosis hirticornis Felt (orig. des.). 

Golanudiplosis Grover & Prasad 1968: 213. Type species, japonica Grover & Prasad (mon.). 

Phagodiplosis Blanchard 1938: 345. Type species, haywardi Blanchard (orig. des.) = coccidivora Felt. 

Vincentodiplosis Harris 1968: 473. Type species, Lobodiplosis coccidarum Felt (orig. des.) = coccidarum Cockerell. 

Cosmopolitan, chiefly tropical; 28 spp.; Cecidomyiidi, unplaced to tribe. Larvae of these species are all predaceous on 

whiteflies and scale insects, chiefly Pseudococcidae. Three genera, Diadiplosis, Megommata, and Triommata, share 

similar straplike adult abdominal sclerites and stout tarsal claws that are bent at the basal third. Eyes of Megommata are 

greatly enlarged on the ventral half and usually mostly divided laterally; in one species, M. pulvinariae, they are 

completely divided. That is a character shared with some Diadiplosis and both species of Triommata. Additionally, 

Megommata has gynecoid male flagellomeres. Streptodiplosis, known from one Indian species, may be a relative of the 

genera above, but it must first be rediscovered, the type being lost. Tricontarinia may also be close to this group of 

genera if not synonymous with one of them. Refs: Harris 1968 (rev., illus.), Gagné 1994a (review). 

aleyrodici. Panama, Ecuador. Aleurycus chagnetios (Hemiptera: Aleyrodidae). 

Cleodiplosis aleyrodici Felt 1922d: 1. Panama: Panama City; ST; ♂,♀; USNM. 

ananasae. Hawaiian Is. Predator possibly of Dysmicoccus brevipes (Hemiptera: Pseudococcidae). 

Coccodiplosis ananasae Hardy 1960: 283. Hawaiian Is: Oahu, Honolulu; HT; ♂; BPBM. 

citri. South Africa. Planococcus citri (Hemiptera: Pseudococcidae). 

Coccodiplosis citri Barnes 1935d: 526. South Africa: Pretoria; ST; ♂,♀; BMNH. 

cocci. Cuba, St. Vincent. Parasaissetia nigra (Hemiptera: Coccidae). 

Diadiplosis cocci Felt 1911c: 55. St. Vincent; ST; ♂,♀,P,L; NYSM. 

coccidarum. USA (Florida), West Indies (Cuba to Trinidad), Colombia, Guyana, Peru. Orig. from a box containing 

various Hemiptera: Aleurodes sp. (Aleyrodidae); Aonidiella aurantii & Dactylopius sp. (Diaspididae); and 

Ferrisia sp., Dysmicoccus spp., Nipaecoccus sp., Paracoccus marginatus, Phenacoccus spp., Pseudococcus 

spp., & Saccharicoccus spp. (Pseudococcidae). 

Diplosis coccidarum Cockerell 1892: 181. Jamaica; HT; ♀; USNM. 

Lobodiplosis coccidarum Felt 1911i: 195. St. Vincent; ST; ♂,♀; NYSM. 

Karschomyia cocci Felt 1913e: 304. Puerto Rico: Patillas; ST; ♂,♀; lost. 

coccidivora. USA (Florida), Bermuda, Jamaica, Guadeloupe, Panama, Guyana, Brazil, Argentina. Pulvinaria urbicola 

(Hemiptera: Coccidae); Alichtensia sp., Coccus sp., Pulvinaria spp., & Saissetia spp. (Hemiptera: Coccidae), 

& Eriococcus sp. (Hemiptera: Eriococcidae). Ref.: Parnell 1966 (biol., anat.). 

Mycodiplosis coccidivora Felt 1911l: 475, nomen nudum. 

Mycodiplosis coccidivora Felt 1911n: 549. Jamaica: Kingston; ST; ♂,♀; USNM. 

Olesicoccus costalimai (as costa-limai) Borgmeier 1931: 186. Brazil: São Paulo; ST; ♂,♀; IBSP. 

Phagodiplosis haywardi Blanchard 1938: 345. Argentina: Entre Rios, Concordia; ST; ♂,♀; DPBA. 

coffeae. Ghana, Nigeria, Democratic Republic of Congo, Uganda, Tanzania. Pseudococcus sp; Planococcus sp. 

(Hemiptera: Pseudococcidae). 

Schizobremia coffeae Barnes 1935d: 528. Uganda: Kampala; ST; ♂,♀; BMNH. 

donaldi. Nigeria. Planococcus citri (Hemiptera: Coccoidea). 

Ghesquierinia donaldi Harris 1968: 446. Nigeria: Ibadan; HT; ♂; BMNH. 

duni. New Britain, Java. Planococcus sp.; Planococcus lilacinus (Hemiptera: Pseudococcidae). 

Nipponodiplosis duni Harris 1968: 460. Papua New Guinea: New Britain, Keravat; HT; ♂; BMNH. 

floridana. USA (Florida), Cuba, Paraguay. 

Kalodiplosis floridana Felt 1915e: 230. USA: Florida, Miami; ST; ♂,♀; USNM. 

formosana. Taiwan. Nipaecoccus filamentosus (Hemiptera: Pseudococcidae). 

Schizobremia formosana Felt 1926d: 183. Taiwan: Taihoku; ST; ♂,♀; NYSM. Taiwan. 

generosi. Sri Lanka. Howardia biclavis and or Pseudaulacaspis sp. nr. pentagona (Hemiptera: Diaspididae).

213 

Dyodiplosis generosi Felt 1915g: 183. Sri Lanka: Peradeniya, Botanic Gardens; ST; ♀; NYSM. 

hirticornis. India, Japan (Honshu, Shikoku), Yemen. Pseudococcus vapor; Planococcus spp. (Hemiptera: 

Pseudococcidae). 

Diadiplosis hirticornis Felt 1915g: 179. Japan; ST; ♂,♀; NYSM. 

Diadiplosis horticornis Fedotova & Kovalev 2001: 578, missp. of hirticornis. 

indica. India (Karnataka). Undet. sp. of Pseudococcidae (Hemiptera). 

Kamptodiplosis indica Rao & Grover 1959: 300. India: Karnataka, Bangalore; HT; ♂; RCMU. 

japonica. Japan (Honshu, Kyushu). Pseudococcus comstocki; Crisococcus matsumotoi (Hemiptera: Pseudococcidae). 

Golanudiplosis japonicus Grover & Prasad 1968: 216. Japan: Honshu, Aomori Pref., Kuroishi; HT; ♂; PGCA. 

jujubae. Mauritius. Pseudococcus sp. (Hemiptera: Pseudococcidae). 

Schizobremia jujubae Barnes 1935d: 529. Mauritius: Rose Hill; ST; ♂,♀; BMNH. 

koebelei. Australia, New Zealand, Hawaiian Is, USA (California). Dactylopius sp. (Hemiptera: Dactylopiidae) & 

Megaspidiotus fimbriatus (as Diaspis fimbriatus) (Hemiptera: Diaspididae); Nipaecoccus vastator, 

Pseudococcus calceolariae, Pseudococcus longispinus (Hemiptera: Pseudococcidae). 

Diplosis koebelei Koebele 1893: 38. Australia; Northern Territory (Gagné & Stein 1982: 66; USA: California, 

Riverside); ♂; USNM. 

Silvestrina koebelei Felt 1932b: 167. USA: California, Riverside; LT (Gagné & Stein 1982: 66); ♂; USNM. 

megalamellae. Ghana, Nigeria, Democratic Republic of Congo, Kenya, Tanzania. Host orig. unkn.; Planococcus 

lilacinus, Planococcus citri (Hemiptera: Pseudococcidae). 

Ghesquierinia megalamellae Barnes 1939f: 327. Democratic Republic of Congo; ST; ♂,♀; BMNH. 

menoni. India (Delhi). Pseudococcus sp. (Hemiptera: Pseudococcidae). 

Dactylodiplosis menoni Mani 1944: 160. India: Delhi; HT; ♂; INPC. This species is newly placed in this genus 

because of its reportedly strongly angled, toothed claws and its predation on a Pseudococcus. 

Dactylodiplosis memoni Rao 1955: 276, missp. of menoni. 

multifila. West Indies (Dominican Republic to Trinidad), Brazil, Fiji. Scale insect; Ferrisia sp., Planococcus citri 

(Hemiptera: Pseudococcidae), & Icerya montserratensis (Hemiptera: Margarodidae) 

Dichrodiplosis multifila Felt 1907d: 19. Puerto Rico; HT; ♂; USNM. 

Dicrodiplosis coccidarum Felt 1911l: 475, nomen nudum. 

Dicrodiplosis coccidarum Felt 1911n: 548. Puerto Rico: Mayaguez; HT; ♀; USNM. 

Diadiplosis buscki Felt 1914f: 122. Puerto Rico; ST; ♂,♀; USNM. 

pseudococci. Mexico, Guatemala, Honduras, Jamaica, Guyana, Brazil, Hawaiian Is. Pseudococcus bromeliae; 

Dysmicoccus brevipes, D. neobrevipes (Hemiptera: Pseudococcidae). 

Diadiplosis pseudococci Felt 1921l: 225. Guyana: Kartabo; ST; ♀,P,L; NYSM. 

Lobodiplosis pseudococci Felt 1933a: 87. Hawaiian Is: Oahu, Konia; ST; ♂,♀; USNM. 

pulvinariae. USA (Florida), West Indies from Jamaica & Dominican Republic to Trinidad, Venezuela, Guyana. 

Protopulvinaria pyriformis; Philephedra tuberculosa (Hemiptera: Coccidae). 

Mycodiplosis pulvinariae Felt 1912b: 175. St. Vincent; ST; ♂,♀,P,L; NYSM. 

Mycodiplosis moznettei Felt 1922a: 46. USA: Florida, Miami; ST; ♂,♀; USNM. 

smithi. Indonesia (Java), Philippines, Papua New Guinea (New Britain). Planococcus citri, P. lilacinus (Hemiptera: 

Pseudococcidae). 

Diadiplosis smithi Felt 1915g: 178. Philippines: Luzon, Manila; ST; ♂,♀; NYSM. 

Coccodiplosis pseudococci Meijere 1917: 239. Indonesia: Java, Salatiga; ST; ♂,♀,L; ZMAN. 

theobromae. Ghana. Delottococcus sp. nr proteae (Hemiptera: Pseudococcidae). 

Coccodiplosis theobromae Harris 1968: 431. Ghana: Tafo; HT; ♂; BMNH. 

tortuosa. Nigeria. Pseudococcus hargreavesi (Hemiptera: Pseudococcidae). 

Ghesquierinia tortuosa Harris 1968: 448. Nigeria: Agodi; HT; ♂; BMNH. 

trinetris. India. 

Diadiplosis trinetris Grover & Bakhshi 1978: 64. India; HT; ♂; PGCA. 

unca. Nigeria. Pseudococcus njalensis; Planococcus citri, Planococcus kenyae (Hemiptera: Pseudococcidae). 

Ghesquierinia unca Harris 1968: 448. Nigeria: Agunmu; HT; ♂; BMNH. 

vaupedis. Colombia, Guadeloupe. Undet. sp. of coccoid; Planococcus sp. (Hemiptera: Pseudococcidae). 

Ghesquierinia vaupedis Harris 1968: 450. Colombia: Vaupes; HT; ♂; BMNH.

DIALERIA 

Dialeria Tavares 1918c: 78. Type species, styracis Tavares (orig. des.). 


styracis. Brazil (Bahia). Styrax sp. (Styracaceae). 

Dialeria styracis Tavares 1918c: 78. Brazil: Bahia, Caeteté; HT; ♀; TAVC. 

DICHAETIA 

Dichaetia Mamaev 1965a: 267. Type species, pusilla Mamaev (orig. des.). 


pusilla. Russia (Europe). 

Dichaetia pusilla Mamaev 1965a: 267. Russia: Tula Prov., Tulskie zaseki; HT; ♂; MAMC. 

DICHODIPLOSIS 

Dichodiplosis Rübsaamen 1910d: 288. Type species, langeni Rübsaamens (orig. des. as n. g., n. sp.). 

Charidiplosis Tavares 1918c: 80. Type species, concinna Tavares (orig. des.). 

Aphodiplosis Gagné 1973c: 864. Type species, Clinodiplosis triangularis Felt (orig. des.). 

Holarctic, Neotropical; 3 spp.; Cecidomyiidi, unplaced to tribe. Dichodiplosis langeni and D. triangularis have been 

reared from mushrooms (Bruns 1982) and from various substrates where the larvae are presumably mycophagous. 

Stenohypodiplosis Fedotova, q.v., appears to be an objective synonym of Dichodiplosis. Ref.: Gagné 2004b (syn. 

notes). 

clavijica. Ecuador. Adults repeatedly assoc. with flowers of Clavija weberbaueri (Theophrastaceae). 

Charidiplosis clavijica Gagné 1997b: 111. Ecuador: Jatun Sacha; HT; ♂; MECN. 

langeni. UK, Germany, Czechoslovakia. 

Dichodiplosis langeni Rübsaamen 1910d: 288. Germany: Remagen; T; ♂; ZMHB. 

triangularis. USA (widespread), Costa Rica, Brazil (Bahia). 

Clinodiplosis triangularis Felt 1908b: 411. USA: New York, Nassau; ST; ♂; NYSM. 

Charidiplosis concinna Tavares 1918c: 81. Brazil: Bahia, Salvador; ST; ♂,♀; TAVC. 

DICRODIPLOSIS 

Dicrodiplosis Kieffer 1895f: cxciv. Type species, fasciata Kieffer (orig. des.). 

Dichrodiplosis Felt 1907d: 19, missp. of Dicrodiplosis. 

Xiphodiplosis Felt 1915g: 180. Type species, fulva Felt (orig. des.). 

Peridiplosis Felt 1918e: 160. Type species, Dichrodiplosis quercina Felt (orig. des., misidentified as Cecidomyia 

quercina Felt; cf. Gagné 1986c: 605). 

Xiphodiplosia Rao 1955: 272, missp. of Xiphodiplosis. 

Tjanshaniella Marikovskij 1953a: 133. Type species, pectinata Marikovskij (orig. des.). 

Cosmopolitan; 13 spp.; Cecidomyiidi, unplaced to tribe. All species with known hosts are predators of scale insects. 

Tjanshaniella, synonymized under Lestodiplosis in Gagné 1973c, more properly fits Dicrodiplosis because its tarsal 

claws are toothed. Ref: Harris 1968 (revision). 

214

215 

antennata. USA (Maine). Phenacoccus acericola (Hemiptera: Pseudococcidae). 

Dicrodiplosis antennata Felt 1912l: 243. USA: Maine, Orono; ST; ♀,L; NYSM. 

bifurcata. India (West Bengal). 

Dicrodiplosis bifurcata Grover 1979c: 113. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

californica. USA (California). Pseudococcus sp. (Hemiptera: Pseudococcidae). 

Dicrodiplosis californica Felt 1912l: 244. USA: California, Riverside; ST; ♂,♀; USNM. 

fasciata. France. 

Dicrodiplosis fasciata Kieffer 1895f: cxciv. France: Moselle; T; ♀; lost?. 

fulva. Sri Lanka, Tanzania. Parasaissetia nigra (Hemiptera: Coccidae). 

Xiphodiplosis fulva Felt 1915g: 180. Sri Lanka: Peradeniya, Botanic Gardens; ST; ♂,L; NYSM. 

guatemalensis. Guatemala, Hawaiian Is. Undet. sp. of Pseudococcidae (Hemiptera). 

Dicrodiplosis guatemalensis Felt 1938: 43. Guatemala; ST; ♂,♀,L; USNM. 

insolens. USA (New York). 

Dicrodiplosis insolens Felt 1920a: 288. USA: New York, Keene Valley; HT; ♀; NYSM. 

kimberleyensis. Ghana, South Africa. Pseudococcus sp. (Hemiptera: Pseudococcidae). 

Dicrodiplosis kimberleyensis Harris 1968: 437. South Africa: Kimberley; HT; ♂; BMNH. 

manihoti. Senegal, Democratic Republic of Congo, Middle East. Phenacoccus manihoti; Ferrisia virgata, 

Planococcus citri (Hemiptera: Pseudococcidae). 

Dicrodiplosis manihoti Harris 1981b: 339. Democratic Republic of Congo: Kombé; HT; ♂; MNHN. 

*marikovskii. Russia (Far East). 

Dicrodiplosis marikovskii Fedotova 2006a: 161. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♀; ZMAS. 

pectinata. Kazakhstan. 

Tjanshaniella pectinata Marikovskij 1953a: 134. Kazakhstan: Zailiïskiï Alatau; ST; ♂,♀; ZKAS & ZMAS. 

pseudococci. Mediterranean Region, Middle East, Nigeria. Planococcus citri (Hemiptera: Pseudococcidae). 

Acaroletes pseudococci Felt 1914a: 148. Italy: Sicily; ST; ♂,♀; NYSM. 

quercina. USA (Georgia). 

Dichrodiplosis quercina Felt 1907d: 19. USA: Georgia, Augusta; HT; ♂; USNM. 

DIDACTYLOMYIA 

Didactylomyia Felt 1911c: 39. Type species, Colpodia longimana Felt (orig. des.). 

Hallomyia Kieffer 1912a: 25. Type species, iris Kieffer (mon.). 

Ditactylomyia Shinji 1944: 213, missp. of Didactylomyia. 

Cosmopolitan; 3 spp.; Stomatosematidi. 

*arcana. Russia (Far East). 

Didactylomyia arcana Fedotova & Sidorenko 2003: 3. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. 


conspicua. India (Uttar Pradesh). 

Didactylomyia conspicua Grover & Bakhshi 1978: 16. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

longimana. Cosmopolitan. Ref.: Sivinsky & Stowe 1981 (biol.). 

Colpodia longimana Felt 1908b: 416. USA: Massachusetts, Auburndale; HT; ♂; NYSM. 

Hallomyia iris Kieffer 1912a: 25. Sri Lanka: Peradeniya; ST; ♀; NZSI?. The orig. illustrations indicate that this sp. 

belongs to Didactylomyia, and, without further information and actual type specimens, there is no reason to 

keep it a distinct species. 

Didactylomyia capitata Felt 1913f: 135, nomen nudum. 

Didactylomyia capitata Felt 1913j: 174. USA: Massachusetts, North Adams, Mt Greylock; HT; ♂; NYSM. 

Didactylomyia flava Felt 1915j: 145. USA: New York, Albany; HT; ♀; NYSM. 

Didactylomyia maculata Felt 1915j: 144. USA: New York, Poughkeepsie; HT; ♀; NYSM. 

DIDYMOMYIA

Didymomyia Rübsaamen 1912d: 216. Type species, Hormomyia reaumuriana Löw (subs. des. Rübsaamen 1916b: 

493) = Cecidomyia tiliacea Bremi. 

Palearctic, recent and Pliocene; 2 spp., but D. accidentalis is probably misplaced here; Lasiopteridi, unplaced to tribe. 

* † accidentalis. Late Eocene - Ukrainian amber. 

Didymomyia accidentalis Fedotova 2005e: 52. Ukraine amber: Rovno; HT; ♀; UASK. 

tiliacea. Widespread Europe, Russia (Far East); Pliocene/Germany. Tilia europaea (Tiliaceae). 

Cecidomyia tiliacea Bremi 1847a: 13. Switzerland: vic. Zurich; ST; LT; ETHZ?. 

Cecidomyia frauenfeldi Kaltenbach 1872: 79, jun. prim. hom. frauenfeldi Schiner 1868. Germany: Aachen; ST; LT; 

NHMW?. 

Hormomyia reaumuriana Löw 1878: 387. Austria: Schönbrunn & Vienna Woods nr Mödling; ST; ♂,♀,L; NHMW. 

*DILATODIPLOSIS 

Dilatodiplosis Fedotova & Sidorenko 2006c: 130. Type species, exerta Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 2 spp.; Cecidomyiidi, Lestodiplosini. This genus is close to Lestodiplosis if the two must be 

separated. 

*exerta. Russia (Far East). 

Dilatodiplosis exerta Fedotova & Sidorenko 2006c: 130. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

*saxosa. Russia (Far East). 

Dilatidiplosis saxosa Fedotova & Sidorenko 2006c: 132. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

*DIMOCARPOMYIA 

Dimocarpomyia Tokuda & Yukawa 2008: 302. Type species, folicola Tokuda & Yukawa (orig. des.). 

Oriental; 1 sp., Asphondyliidi, Asphondyliina. 

*folicola. Thailand. Dimocarpus longan (Sapindaceae). 

Dimocarpomyia folicola Tokuda & Yukawa 2008: 303. Thailand: Changmai, San Sai Distr., Faek Mai, Tambol Mae; 


DIODAULUS 

Diodaulus Rübsaamen 1917: 87. Type species, Cecidomyia linariae Winnertz (orig. des.). 


linariae. Widespread Europe. Linaria vulgaris (Scrophulariaceae). 

Cecidomyia linariae Winnertz 1853: 260. Germany; ST; ♂,♀; lost. 

traili. Widespread Europe. Pimpinella saxifraga (Apiaceae). 

Diplosis traili Kieffer 1889e: 262. France: vic. Bitche; ST; ♂,♀,L; lost?. 

DIPLECUS 

Diplecus Kieffer 1913g: 55. Type species, Coprodiplosis inconspicuus Kieffer (orig. des.). 

216


inconspicuus. Sri Lanka. 

Coprodiplosis inconspicuus Kieffer 1912a: 29. Sri Lanka: Peradeniya; HT; ♀; NZSI?. 

DIPLODONTOMYIA 

Diplodontomyia Rao 1951: 120. Type species, orientalis Rao (orig. des.). 

Oriental; 2 spp.; Cecidomyiidi, unplaced to tribe. 

deepica. India (Uttar Pradesh). 

Diplodontomyia deepica Rao 1953: 310. India: Uttar Pradesh, Agra, St. John's College; HT; ♂; ZSCI. 


Diplodontomyia orientalis Rao 1951: 121. India: Uttar Pradesh, Agra, Hewitt Park; HT; ♂; ZSCI. 

DIPLOSIOLA 

Diplosiola Solinas 1965c: 55. Type species, bursaria Solinas (orig. des. as n. g., n. sp.). 


bursaria. Italy. Medicago sativa (Fabaceae). 

Diplosiola bursaria Solinas 1965c: 58. Italy: northwestern Emilia-Romagna; ST; ♂,♀,P,L; Solinas Coll. 

*DISSIMILIDIPLOSIS 

Dissimilidiplosis Fedotova 2005c: 224. Type species, strumosa Fedotova (orig. des.). 


*strumosa. Russia (Far East). 

Dissimilidiplosis strumosa Fedotova 2005c: 225. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. Kamenushka; 


DOMOLASIOPTERA 

Domolasioptera Möhn 1975: 31. Type species, adversaria Möhn (orig. des.). 

Neotropical; 7 spp., all inquilines in galls formed by other Cecidomyiinae; Lasiopteridi, Camptoneuromyiini. 

acuario. Colombia. Mikania cordifolia (Asteraceae). 

Domolasioptera acuario Wünsch 1979: 115. Colombia: Magdalena, Rio Buritaca; HT; ♂; SMNS. 

adversaria. El Salvador. Xylosma flexuosum (Flacourtiaceae). 

Domolasioptera adversaria Möhn 1975: 32. El Salvador: Santa Ana, SE Candelaria, Hacienda Cortez; HT; ♂; 

SMNS. 

ayeniae. El Salvador. Ayenia pusilla (Sterculiaceae). 

Domolasioptera ayeniae Möhn 1975: 35. El Salvador: NE Santa Ana, rd to Metapán; HT; LT; SMNS. 

baca. El Salvador. Indigofera suffruticosa (Fabaceae). 

Domolasioptera baca Möhn 1975: 38. El Salvador: San Salvador; HT; LT; SMNS. 

curatellae. El Salvador. Curatella americana (Dilleniaceae). 

217

218 

Domolasioptera curatellae Möhn 1975: 37. El Salvador: Chalatenango, SE La Palma; HT; LT; SMNS. 

securidacae. El Salvador. Securidaca sylvestris (Polygalaceae). 

Domolasioptera securidacae Möhn 1975: 33. El Salvador: San Salvador; HT; ♂; SMNS. 

thevetiae. El Salvador. Thevetia plumeriaefolia (Apocynaceae). 

Domolasioptera thevetiae Möhn 1975: 34. El Salvador: San Vicente, SE Santo Domingo, km 54; HT; LT; SMNS. 

DRISINA 

Drisina Giard 1893: cccxliii. Type species, glutinosa Giard (orig. des.). 


glutinosa. Widespread Europe. Acer pseudoplatanus (Aceraceae). 

Drisina glutinosa Giard 1893: cccxlii. France; NT (Skuhravá & Skuhravy 1986: 2; Czech Republic: Závissín, nr 

Mariánske Lázne); ♂; SKCR. 

Massalongia aceris Rübsaamen 1921: 40. Germany: Partenkirchen; ST; ♀,L; ZMHB. 

DRYOMYIA 

Dryomyia Kieffer 1898f: 17. Type species, Cecidomyia circinans Giraud (subs. des. Sharp 1901: 296). 

Palearctic; 4 spp., all from leaf galls on Quercus; Lasiopteridi, Dasineurini. 

circinans. Italy, Czechoslovakia, Austria, Hungary, Romania, Balkan Peninsula, Turkey, Japan?. Quercus cerris 

(Fagaceae). 

Cecidomyia circinans Giraud 1861: 475. Austria; ST; ♂,♀,P,L; MNHN. 

Dryomyia circinnans Yukawa 1971: 102, missp. of circinans. 

cocciferae. Portugal, France, Algeria, Tunisia. Quercus coccifera (Fagaceae). 

Cecidomyia cocciferae Marchal 1897a: 19. Algeria: Bouzarea, nr Algiers; ST; ♂,♀,P,L; MNHN?. 

dubia. Spain. 

Dryomyia dubia Tavares 1916b: 121. Spain: Gallicia, Tuda; HT; ♀; TAVC. 

lichtensteinii. France, Poland, Italy, Yugoslavia, Morocco, Algeria. Quercus ilex (Fagaceae). 

Cecidomyia lichtensteinii Löw 1878: 392. France: Montpellier; ST; ♂,♀,L; NHMW. 

Dryomyia lichtensteini Skuhravá 1986: 171, missp. of lichtensteinii 

ECHINELLA 

Echinella Mamaev 1965a: 267, preoccupied Swainson 1840 (Mollusca); needs to be renamed. Type species, 

albiventris Mamaev (orig. des.). 


albiventris. Russia (Europe). 

Echinella albiventris Mamaev 1965a: 267. Russia: Yaroslav Prov., Volkovo; HT; ♂; HYNM. 

polypori. Latvia, Russia (Europe). Fomes marginatus (Polyporales). 

Echinella polypori Mamaev 1965a: 267. Russia: Yaroslav Prov., Volkovo; HT; ♂; HYNM. 

EDESTOCHILUS 

Edestochilus Gagné 1969d: 1358. Type species, Lasioptera allioides Pritchard (orig. des.). 

Nearctic; 1 sp.; Lasiopteridi, Alycaulini.

allioides. USA (California). Paspalum distichum (Poaceae). 

Lasioptera allioides Pritchard 1961a: 56. USA: California, Fresno; HT; ♀; USNM. 

EDESTOSPERMA 

Edestosperma Gagné 1969d: 1356. Type species, aristidae Gagné (orig. des.). 


aristidae. USA (Idaho). Aristida longiseta (Poaceae). 

Edestosperma aristidae Gagné 1969d: 1357. USA: Idaho, Idaho Co., Salmon River Canyon; HT; ♀; USNM. 

*EFFERATODIPLOSIS 

Efferatodiplosis Fedotova 2006a: 160. Type species, ornata Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Cecidomyiini. 

*ornata. Russia (Far East). 

Efferatodiplosis ornata Fedotova 2006a: 161. Russia: Primorskiï Terr., Vladivostok; HT; ♂; ZMAS. 

*EFFUSOMYIA 

Effusomyia Fedotova 2004i: 596. Type species, pusilla Fedotova (orig. des.). 

Palearctic; 1 sp.; Brachineuridi. 

*pusilla. Russia (Far East). 

Effusomyia pusilla Fedotova 2004i: 596. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

*EMARGINATODIPLOSIS 

Emarginatodiplosis Fedotova 2003e: 51. Type species, libera Fedotova (orig. des.). 


separated. 

*libera. Russia (Far East). 

Emarginatodiplosis liberum Fedotova 2003e: 51. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

ENALLODIPLOSIS 

Enallodiplosis Gagné 1994a: 185. Type species, discordis Gagné (orig. des.). 

Neotropical; 1 sp., possibly predaceous; Cecidomyiidi, unplaced to tribe. 

discordis. Chile, Argentina. Prosopis tamarugo (Fabaceae). 

Enallodiplosis discordis Gagné 1994a: 187. Chile: Canchones; HT; ♂; USNM. 

ENDAPHIS 

219

Endaphis Kieffer 1896j: 383. Type species, perfida Kieffer (mon.). 

Pseudendaphis Barnes 1954a: 772. Type species, maculans Barnes (mon.). 

Occuloxenium Mamaev 1973b: 51. Type species, compitale Mamaev (mon.). 

Holarctic, Oriental; 6 spp., internal parasitoids of aphids; Cecidomyiidi, Lestodiplosini. 

aphidimyza. India (Maharashtra). Dactynotus sonchi (Hemiptera: Aphididae). 

Endaphis aphidimyza Shivpuje & Raodeo 1985: 61. India: Maharashtra, Parbhani; HT; ♂; Shivpuje Coll., 

Maharashtra Univ., Parbhani. 

compitalis. Uzbekistan. Aphis pomi (Hemiptera: Aphididae). 

Occuloxenium compitale Mamaev 1973b: 53. Uzbekistan: Namangan Prov.; HT; ♂; HYNM. 

fugitiva. Hawaiian Is. Pentalonia nigronervosa (Hemiptera: Aphididae). 

Endaphis fugitiva Gagné & Muratori 2009: 187. Hawaiian Islands: Kauai, Kauai Agricultural Res. Stn; HT; ♂; 


gregaria. Canada (British Columbia). Euceraphis gillettei (Hemiptera: Aphididae). 

Endaphis gregaria Gagné 1981c: 222. Canada: British Columbia, Burnaby; HT; ♂; USNM. 

maculans. USA (Florida), Guadeloupe, Costa Rica, Trinidad, Hawaiian Is. Toxoptera aurantii; Aphis gossypii, A. 

spiraecola (Hemiptera: Aphididae). 

Pseudendaphis maculans Barnes 1954a: 772. Trinidad; ST; ♂,♀; BMNH. 

perfida. UK, France, Russia (Europe). Undet. aphid; Drepanosiphum platanoides (Hemiptera: Aphididae). 

Endaphis perfida Kieffer 1896j: 383. France: château de La Grange, nr Thionville; T; ♀; lost?. 

ENDOPSYLLA 

Endopsylla Meijere 1907: 172. Type species, agilis Meijere (mon.). 

Palearctic; 2 spp., internal parasitoids of Hemiptera; Cecidomyiidi, Lestodiplosini. Ref.: Harris 1966 (illus.). 

agilis. UK, Netherlands, Czechoslovakia. Psylla foersteri (Hemiptera: Psyllidae). 

Endopsylla agilis Meijere 1907: 173. Netherlands: Kortenhoef; ST; ♂,♀,L; ZMAN. 

endogena. Portugal. Tingis piri (Hemiptera: Tingidae). 

Cecidomyia endogena Kieffer 1907a: 129. Portugal: St. Fiel; ST; ♂,P; lost?. 

EOCINCTICORNIA 

Eocincticornia Felt 1915a: 206. Type species, australasiae Felt (orig. des.). 

Australasian; 2 spp., the one reared species from Eucalyptus; Cecidomyiidi, Asphondyliini, Schizomyiina. 

australasiae. Australia (Western Australia). 

Eocincticornia australasiae Felt 1915a: 207. Australia: Western Australia, Perth; ST; ♂,♀; USNM. 

malarskii. Australia (South Australia). Eucalyptus fasciculosa (Myrtaceae). 

Eocincticornia malarskii Kolesik 1995a: 148. Australia: South Australia, Morialta Conservation Pk, 13 km NE 


EOHORMOMYIA 

Eohormomyia Felt 1915a: 208. Type species, howardi Felt (orig. des.). 


220

howardi. Mozambique. 

Eohormomyia howardi Felt 1915a: 208. Mozambique: Maputo, along Umbeluzi R.; HT; ♀; USNM. 

EPHEDROMYIA 

Ephedromyia Marikovskij 1953a: 130. Type species, debilopalpis Marikovskij (orig. des.). 

Palearctic; 6 spp., all from Ephedra; Lasiopteridi, unplaced to tribe. 

debilopalpis. Russia (Europe), Kazakhstan, Turkmenistan. Ephedra sp. (Gnetaceae). 

Ephedromyia debilopalpis Marikovskij 1953a: 130. Kazakhstan: Dzhungarskiï Alatau, right bank mid Ili R., Chulak; 

ST; ♂,♀; ZKAS & ZMAS. 

issykkulensis. Kirghizstan. Ephedra sp. (Gnetaceae). 

Ephedromyia issykkulensis Marikovskij 1955e: 22. Kirghizstan: north shore Issyk-Kul' L.; ST; ♂,♀; ?. 

kazenasi. Kazakhstan. Ephedra intermedia (Gnetaceae). 

Ephedromyia kazenasi Fedotova 1993d: 90. Kazakhstan: NE spurs Zailiïskiï Alatau, 32 km SE Chilik, 10 km NW 

Kok-Pek; HT; ♂; ZMAS. 

kopetdagica. Turkmenistan. Ephedra equisetina (Gnetaceae). 

Ephedromyia kopetdagica Fedotova 1993d: 92. Turkmenistan: Kopetdag, Ioldere ravine, 18 km S Kara-Kala; HT; ♂; 

ZMAS. 

kugitangica. Turkmenistan. Ephedra intermedia (Gnetaceae). 

Ephedromyia kugitangica Fedotova 1993d: 93. Turkmenistan: Kugitang Mtn Range, nr summit Ayri-Baba, 70 km 

NE Charshanga; HT; ♂; ZMAS. 

strobilaceae. Turkmenistan. Ephedra strobilacea (Gnetaceae). 

Ephedromyia strobilaceae Fedotova 1993d: 90. Turkmenistan: eastern Karakumy Desert, 70 km SW Chardzhou, 

Repetek Stn; HT; ♂; ZMAS. 

EPICALAMUS 

Epicalamus Sylvén 1998a: 100. Type species, phalaridis Sylvén (orig. des.). 

Western Palearctic; 1 sp.; Lasiopteridi, Poomyini. The general similarity of the adult abdomen of both sexes and of 

the larvae (except for the lack of a puparium) with Mayetiola indicates a relationship between the two genera. 

phalaridis. Sweden. Phalaris arundinacea (Poaceae). 

Epicalamus phalaridis Sylvén 1998a: 100. Sweden: Norrbotten, Vojakkala; HT; ♀; NHRS. 

EPIDIPLOSIS 

Epidiplosis Felt 1908b: 406. Type species, sayi Felt (orig. des.). 

Gersonomyia Nijveldt 1965: 42. Type species, filifera Nijveldt (orig. des.). 

Epdiplosis Shinji 1944: 244, missp. of Epidiplosis. 

Holarctic, Oriental; 13 spp., one a predator of a coccoid (Hemiptera), the remainder caught in flight; Cecidomyiidi, 

Lestodiplosini. Fedotova and Sidorenko 2009a erected for this one genus the new subtribe Epidiplosina and placed it 

within the tribe Karshomyini, but the suite of characters of the type species of both generic synonyms of this genus 

definitely fit Lestodiplosini instead. Ref.: Li & Bu 2006 (review, key to spp.). 

*badenkoi. Russia (Far East). 

Epidiplosis badenkoi Fedotova & Sidorenko 2009a: 15. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


*bicornuta. China (Shaanxi). 

221

222 

Epidiplosis bicornuta Li & Bu 2006: 81. China: Shaanxi, Ningshan, Huoditang; HT; ♂; NKUM. 

*campanulata. China (Shaanxi). 

Epidiplosis campanulata Li & Bu 2006: 82. China: Shaanxi, Ningshan, Xunyangba; HT; ♂; NKUM. 

filifera. UK, Israel, Russia (Far East). Ceroplastes floridensis (Hemiptera: Coccidae); Lepidosaphes beckii (Hemiptera; 

Diaspididae). 

Gersonomyia filifera Nijveldt 1965: 42. Israel: nr Kfar Hess; HT; ♂; ZMAN. 

*furcata. China (Zhejiang). 

Epidiplosis furcata Mo & Zheng 2004: 563. China: Zhejiang, West Tianmushan; HT; ♂; SAUT. 

indica. India (Assam), China (Guangxi, Yunnan). 

Epidiplosis indica Grover & Chandra 1989: 158. India: Assam, Shillong; HT; ♂; PGCA. 

*lari. China (Guizhou). 

Epidiplosis lari Li & Bu 2006: 84. China: Guizhou, Libo; HT; ♂; NKUM. 

*mica. Russia (Far East). 

Opinadiplosis mica Fedotova 2004g: 27. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

reflexa. Russia (Europe), China (Heilongjiang). 

Epidiplosis reflexa Mamaev 1969a: 416. Russia: Moscow Prov., Danki; HT; ♂; IEME. 

sayi. USA (New York). 

Epidiplosis sayi Felt 1908b: 406. USA: New York, Nassau; HT; ♂; NYSM. 

simulifilifera. China (Shandong). 

Epidiplosis simulifilifera Mo 2000b: 184. China: Shandong, Taian; HT; ♂; SAUT. 

triangularis. China (Hubei, Shaanxi). 

Epidiplosis triangularis Mo & Liu 2000: 122. China: Hubei; HT; ♂; SAUT. 

*tuberoforcipata. China (Fujian, Yunnan, Guangxi). 

Epidiplosis tuberoforcipata Li & Bu 2006: 85. China: Fujian, Wuyishan, Tongmu, Qiliqiao; HT; ♂; NKUM. 

EPIHORMOMYIA 

Epihormomyia Felt 1915h: 155. Type species, auripes Felt (mon.). 


auripes. Guyana. 

Epihormomyia auripes Felt 1915h: 156. Guyana: Mallali, Demerara R.; HT; ♀; NYSM. 

miconiae. Brazil (Rio de Janeiro). Miconia cinnamomifolia (Melastomataceae). 

Epihormomyia miconiae Maia 2001a: 14. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

EPILASIOPTERA 

Epilasioptera Möhn 1964b: 573. Type species, iresinis Möhn (orig. des.). 


iresinis. El Salvador. Iresine calea (Amaranthaceae). 

Epilasioptera iresinis Möhn 1964b: 573. El Salvador: La Libertad, Cumbre, Hacienda Chilata; HT; ♂; SMNS. 

EPIMYIA 

Epimyia Felt 1911c: 38. Type species, carolina Felt (orig. des.). 

Nearctic, Palearctic; 2 spp.; Brachineuridi. Refs: Bruns 1982 (hosts), Gagné 1994a (review).

223 

*calcarata. Russia (Far East). 

Epimyia calcarata Fedotova & Sidorenko 2006d: 92. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad; HT; 

♂; ZMAS. 

carolina. USA (Minnesota, Maine, North Carolina). Orig. unk.; Boletus, Leccinium, Paxillus, Suillus, & Strobilocyces 

(Boletales). 

Epimyia carolina Felt 1911c: 38. USA: North Carolina; HT; ♂; NYSM. 

EPIMYIELLA 

Epimyiella Mamaev 1967: 879. Type species, palustris Mamaev (orig. des.). 

Palearctic; 1 sp.; Brachineuridi. 

palustris. Latvia, Russia (Europe). 

Epimyiella palustris Mamaev 1967: 880. Russia: Moscow Prov., Glubokoye L.; HT; ♂; HYNM. 

ETSUHOA 

Etsuhoa Inouye 1959: 10. Type species, okayamana Inouye (orig. des.). 

Etsuchoa Marikovskij 1969: 96, missp. of Etsuhoa. 

Palearctic; 10 spp., all from Juniperus; Cecidomyiidi, unplaced to tribe. 

kopetdagica. Turkmenistan. Juniperus turcomanica (Cupressaceae). 

Etsuhoa kopetdagica Fedotova 1993b: 20. Turkmenistan: western Kopetdag, 18 km S Mala-Kara; HT; ♂; ZMAS. 

kugitangica. Turkmenistan. Juniperus seravschanica (Cupressaceae). 

Etsuhoa kugitangica Fedotova 1993b: 17. Turkmenistan: Kugitang Mtn Range, on Aïri-Baba, 70 km NE Charshanga; 


okayamana. Japan (Honshu). Juniperus rigida (Cupressaceae). 

Etsuhoa okayamana Inouye 1959: 4. Japan: Honshu, Okayama Pref.; HT; ♂; HFES. 

renifolia. Russia (Europe), Kazakhstan. Juniperus pseudosabina (Cupressaceae). 

Etsuhoa renifolia Fedotova 1985b: 32. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak, vic. Alma-Ata; HT; ♂; ZKAS. 

sabinae. Spain or France, Austria, Kazakhstan. Juniperus communis or J. phoenica; J. sabina (Cupressaceae). 

Oligotrophus sabinae Kieffer 1898a: 17. Spain or France, & Austria; T; G; lost?. 

severtzovi. Kirghizstan. Juniperus sp.; J. turkestanica (Cupressaceae). 

Etsuchoa severtzovi Marikovskij 1969: 96. Kirghizstan: Talasskiï Alatau; ST; ♂; MARC. 

tarbagataica. Kazakhstan. Juniperus pseudosabina (Pinaceae). 

Etsuhoa tarbagataica Fedotova 1990g: 65. Kazakhstan: Tarbagatai Mtn Range, upper Karabuga R., 80 km S Aksuat; 


thuriferae. Spain. Juniperus thurifera (Cupressaceae). 

Etsuhoa thuriferae Skuhravá 1996: 135. Spain: Zaragoza Prov., Monegros; HT; ♂; SKCR. 

tjanshanica. Kirghizstan. Juniperus sp.; J. semoglobosa (Cupressaceae). 

Etsuhoa tjanshanica Marikovskij 1969: 96. Kirghizstan: Talasskiï Alatau; ST; ♂,♀; MARC. 

Etsuhoa tianshanica Fedotova 1990g: 68, missp. of tjanshanica. 

yakhontovi. Uzbekistan. Juniperus seravschanica (Cupressaceae). 

Etsuhoa yakhontovi Mamaev 1969b: ? (publ'n not seen). Uzbekistan: ?; ?; ?; MAMC. 

EUCALYPTODIPLOSIS 

Eucalyptodiplosis Kolesik 2002: 24. Type species, germinis Kolesik (orig. des.). 

Australasian; 3 spp., on Eucalyptus; Cecidomyiidi, unplaced to tribe.

*chionochloae. New Zealand. Chionochloa pallens; Chionochloa spp. 

Eucalytodiplosis chionochloae Kolesik 2007b: 109. New Zealand: [South Island], Mt. Hutt; HT; ♂; NZAC. 

germinis. Australia (South Australia). Eucalyptus cosmophylla; E. camaldulensis (Myrtaceae). 

Eucalytodiplosis germinis Kolesik 2002: 25. Australia: South Australia, nr Bridgewater; HT; ♂; SAMA. 

mcintyrei. Australia (New South Wales). Eucalyptus grandis (Myrtaceae). 

Eucalytodiplosis germinis Kolesik 2002: 25. Australia: New South Wales, nr Kyogle; HT; ♂; SAMA. 

EUGENIAMYIA 

Eugeniamyia Maia, Mendonça & Romanowski 1997: 1087. Type species, dispar Maia, Mendonça & Romanowski 

(orig. des. as n. g., n. sp.). 


dispar. Brazil (Rio Grande do Sul). Eugenia uniflora (Myrtaceae). 

Eugeniamyia dispar Maia, Mendonça, & Romanowski 1997: 1089. Brazil: Rio Grande do Sul, Porto Alegre; HT; ♂; 

QBUM. 

EUMEROSEMA 

Eumerosema Kieffer 1913e: 42. Type species, dispar Kieffer (orig. des.). 


dispar. Taiwan. 

Eumerosema dispar Kieffer 1913e: 42. Taiwan: Tainan; ST; ♂,♀; lost?. 

EUPHORBOMYIA 

Euphorbomyia Fedotova 1994i: 456. Type species, Cecidomyia loewii Mik (orig. des.). 

Palearctic; 4 spp., all on Euphorbia; Lasiopteridi, Dasineurini. 

jaxarticae. Kazakhstan. Euphorbia jaxartica; E. microcarpa (Euphorbiaceae). 

Euphorbomyia jaxarticae Fedotova 1994i: 458. Kazakhstan: Talasskiï Alatau, Zhalpaksaï gorge, 10 km S Kaskasu; 


latifoliae. Kazakhstan. Euphorbia latifolia (Euphorbiaceae). 

Euphorbomyia latifoliae Fedotova 1994i: 459. Kazakhstan: SE L. Balkhash area, Karatau Mtn Range, Arkarly Mtn; 


loewii. France, Germany, Czechoslovakia, Austria, Hungary, Romania, Bulgaria, Ukraine, Kazakhstan. Euphorbia 

seguierana (Euphorbiaceae). 

Cecidomyia loewii Mik 1882: 265. Austria: nr Oberweiden; ST; ♂,♀; NHMW. Fedotova (1994i: 457) invalidly fixed 

a neotype for Cecidomyia loewii Mik when designating it as type species of Euphorbomyia: that Mik's types 

might exist was not discussed. 

turkestanica. Kazakhstan. Euphorbia jaxartica (Euphorbiaceae). 

Euphorbomyia turkestanica Fedotova 1994i: 461. Kazakhstan: right bank Syr-Darya R., 30 km SW Turkestan, nr 

Baltakol'; HT; ♂; ZMAS. 

*EXIGUIDIPLOSIS 

Exiguidiplosis Fedotova 2004h: 175. Type species, flaccida Fedotova (orig. des.). 

224


*flaccida. Russia (Far East). 

Exiguidiplosis flaccida Fedotova 2004h: 176. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

EXOPSYLLOMYIA 

Exopsyllomyia Mamaev 1972a: 886. Type species, symbia Mamaev (orig. des.). 

Palearctic; 3 spp., all on Chenopodiaceae; Lasiopteridi, Lasiopterini. 

karakumensis. Turkmenistan. Salsola rigida (Chenopodiaceae). 

Stephaniella karakumensis Marikovskij 1957: 942. Turkmenistan: Taschauz Region, Kuna Urgench; ST; ♂,♀; 

MARC. 

symbia. Turkmenistan, Mongolia. Salsola sp. (Chenopodiaceae). 

Exopsyllomyia symbia Mamaev 1972a: 886. Turkmenistan: Mary; HT; ♀; HYNM. 

sympegmae. Kazakhstan. Sympegma regeli (Chenopodiaceae). 

Exopsyllomyia sympegmae Fedotova 1986b: 629. Kazakhstan: northern Tyan'-Shan', Kyrtogaï canyon, 40 km NW 

Kegen; HT; ♀; ZMAS. 

FABOMYIA 

Fabomyia Fedotova 1991d: 23. Type species, Dasineura sphaerophysae Fedotova (orig. des.). 

Palearctic; 8 spp., all on Fabaceae; Lasiopteridi, Dasineurini. 

arbuscula. Kazakhstan. Astragalus arbuscula (Fabaceae). 

Fabomyia arbuscula Fedotova 1993j: 81. Kazakhstan: 118 km NE Alma-Ata, 18 km NE Sary-Ozek; HT; ♂; ZMAS. 

biczurini. Kazakhstan. Astragalus neolipskianus; A. falcigerus (Fabaceae). 

Fabomyia biczurini Fedotova 1993j: 82. Kazakhstan: 118 km NE Alma-Ata, 18 km NE Sary-Ozek; HT; ♂; ZMAS. 

elenae. Kazakhstan. Medicago tianschanica (Fabaceae). 

Fabomyia elenae Fedotova 1996e: 44. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Baldabrek Mtn Range, 


karatavica. Kazakhstan. Astragalus pseudomacropterus (Fabaceae). 

Fabomyia karatavica Fedotova 1992g: 51. Kazakhstan: Karatau Mtn Range, 20 km SW Baïzhansaï, nr Leont'evkï; 


medicaginis. Widespread Europe, Russia (Western Siberia), Kazakhstan. Medicago sativa; M. falcata (Fabaceae). 

Dasyneura medicaginis Rübsaamen 1912e: 287. Germany; ST; ♀,L; ZMHB. 

melissitis. Kazakhstan. Melissitus popovi (Fabaceae). 

Fabomyia melissitis Fedotova 1992g: 51. Kazakhstan: Talasskiï Alatau, Zhalpaksaï gorge, 10 km S Kaskasu; HT; ♂; 

ZMAS. 

paucijugus. Kazakhstan. Astragalus paucijugus (Fabaceae). 

Fabomyia paucijugus Fedotova 1993j: 84. Kazakhstan: southeastern Kyzylkum Desert, 20 km NW Koksu, 68 km 

NW Chardara; HT; ♂; ZMAS. 


Dasyneura sphaerophysae Fedotova 1983c: 26. Kazakhstan: Ili R. delta, Apistan canal, Zhydeli; HT; ♂; ZMAS. 

*viciafoliae. Russia (Far East). Vicia voroshilovi (Fabaceae). 

Fabomyia viciafoliae Fedotova 2004e: 21. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

225

*FALCIFORMIDIPLOSIS 

Falciformidiplosis Fedotova 2003e: 62. Type species, stylis Fedotova (orig. des.). 

Eastern Palearctic; 2 spp.; Cecidomyiidi, Aphidoletini. This genus is close to Bremia if the two must be separated. 

*laqueata. Russia (Far East). 

Falciformidiplosis laqueata Fedotova 2003e: 65. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

*stylis. Russia (Far East). 

Falciformidiplosis stylis Fedotova 2003e: 63. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*FARISTODIPLOSIS 

Faristodiplosis Fedotova 2005d: 232 (also as Fareastodiplosis, p.227). Type species, glochidiata Fedotova (orig. 

des.). 

Palearctic; 13 spp.; Cecidomyiidi, Mycodiplosini. This genus does not appear to comprise a monophyletic 

assemblage. 

*aculeata. Russia (Far East). 

Faristodiplosis aculeata Fedotova 2005d: 232. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*aliformis. Russia (Far East). 

Faristodiplosis aliformis Fedotova & Sidorenko 2005c: 237. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*assueata. Russia (Far East). 

Faristodiplosis asssueta Fedotova & Sidorenko 2009c: 83. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 


*collaris. Russia (Far East). 

Faristodiplosis collaris Fedotova 2005d: 233. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*clara. Russia (Far East). 

Coquillettomya (sic!) clara Fedotova & Sidorenko 2006a: 134. Russia: Primorskiï Terr., vic. Vladivostok, Sputnik; 


*finitima. Russia (Far East). 

Faristodiplosis finitima Fedotova & Sidorenko 2009c: 86. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


*furcata. Russia (Far East). 

Faristodiplosis furcata Fedotova 2005d: 234. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*glochidiata. Russia (Far East). 

Faristodiplosis glochidiata Fedotova 2005d: 232. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

*rara. Russia (Far East). 

Coquillettomya (sic!) rara Fedotova & Sidorenko 2006a: 134. Russia: Primorskiï Terr., Vladivostok, Sputnik; HT; ♂; 

ZMAS. 

*scalaris. Russia (Far East). 

Faristodiplosis scalaris Fedotova & Sidorenko 2005c: 235. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*spathulata. Russia (Far East). 

226

227 

Faristodiplosis spathulata Fedotova & Sidorenko 2005c: 236. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*stratiosa. Russia (Far East). 

Faristodiplosis stratiosa Fedotova & Sidorenko 2005c: 235. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*urceolata. Russia (Far East). 

Faristodiplosis urceolata Fedotova 2005d: 237. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

FARQUHARSONIA 

Farquharsonia Collin 1921: 504. Type species, rostrata Collin (mon.). 

Megauchomyia Felt 1928a: 123. Type species, africana Felt (orig. des.) = rostrata Collin. 

Afrotropical; 2 spp.; Aphidoletini. 

manicata. Cameroon. 

Megauchomyia manicata Edwards 1929b: 68. Cameroon: Mo Tyan; HT; ♀; BMNH. 

rostrata. Ghana, Liberia, Nigeria. Adults assoc. with Crematogaster sp. (Hymenoptera: Formicidae) and hovering over 

undet. Coccoidea (Hemiptera). 

Farquharsonia rostrata Collin 1922: 505. Nigeria: Agege, 16 mi N Lagos, & Moor Plantation, nr Ibadan; ST; ♂,♀ ; 

BMNH. 

Megauchomyia africana Felt 1928a: 123. Liberia: Bakratown; ST; ♂,♀; NYSM. 

FELTIELLA 

Feltiella Rübsaamen 1910d: 285. Type species, tetranychi Rübsaamen (orig. des. as n. g., n. sp.) = acarisuga Vallot. 

Therodiplosis Kieffer 1912h: 2. Type species, persicae Kieffer (orig. des.) = acarisuga Vallot. 

Acaroletes Kieffer 1913a: 229. Type species, Arthrocnodax tetranychi Kieffer (orig. des.). 

Acroletes Mani 1946: 215, missp. of Acaroletes. 

Cosmopolitan; 10 spp.; Cecidomyiidi, Lestodiplosini. Larvae of this cosmopolitan genus prey almost exclusively on 

red spider mites. Refs: Gagné 1995a (rev.), 2004b (syn. note). 

acarisuga. Europe (widespread), North Africa, Israel, Japan, India, Sri Lanka, Taiwan, Nearctic (widespread), 

Australia (ACT), New Zealand. Tetranychus sp.; many tetranychids (Acarina: Tetranychidae). 

Cedomyia acarisuga Vallot 1827: 95. France: Dijon; NT (Gagné 1995a: 23; Switzerland: la Tour de Peilz); ♂; 

BMNH. 

Mycodiplosis minuta Felt 1907a: 31. USA: New York, Nassau; HT; ♂; NYSM. 

Feltiella tetranychi Rübsaamen 1910d: 285. Germany: vic. Berlin; ST; ♂,♀,L; ZMHB. 

Therodiplosis persicae Kieffer 1912h: 2. France; ST; ♂,♀; lost. 

Feltiella davisi Felt 1915f: 406. USA: Indiana, Lafayette; ST; ♂,♀; NYSM. 

Feltiella americana Felt 1916a: 33. USA: New York, Geneva; ST; ♂,♀; NYSM. 

Feltiella ithacae Felt 1926b: 141. USA: New York, Ithaca; LT (Gagné 1995a: 23); ♂; NYSM. 

Therodiplosis beglarovi Mamaev 1965a: 267. Russia: Leningrad Prov.; HT; ♂; MAMC. 

Feltiella quadrata Kashyap 1988b: 60. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

Arthrocnodax rutherfordi Felt 1915g: 180. Sri Lanka: Peradeniya, Botanic Gardens; LT (Gagné 1995a: 23); ♂; 

NYSM. 

acarivora. Indonesia (Java), Australia (Northern Territory). Tetranychus exsiccator (Acarina: Tetranychidae). 

Diplosis acarivora Zehntner 1901: 203. Java; ST; ♂,♀; lost?. 

curtistylus. Brazil (Pernambuco). Tetranychus evansi (Acarina: Tetranychidae). 

Feltiella curtistylus Gagné 1984c: 128. Brazil: Pernambuco, Petrolina; HT; ♂; MZSP.

228 

insularis. USA (Illinois, New Jersey, Florida), Jamaica, Puerto Rico, Trinidad, Colombia, Argentina. Tetranychus sp.; 

many tetranychids (Acarina). 

Mycodiplosis insularis Felt 1913e: 305. Puerto Rico: Rio Piedras; LT (Gagné 1995a: 26); ♂; NYSM. 

Arthrocnodax carolina Felt 1913i: 488. USA: South Carolina, Batesburg; LT (Gagné 1995a: 26); ♂; NYSM. 

Arthrocnodax constricta Felt 1914i: 481. Puerto Rico: Rio Piedras; LT (Gagné 1995a: 26); ♂; NYSM. 

Feltiella venatoria Felt 1917b: 195. USA: Illinois, Chicago; LT (Gagné 1995a: 26); ♂; NYSM. 

kanchanjungaensis. India (West Bengal). 

Triommata kanchanjungaensis Grover 1979c: 59. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

ligulata. Cape Verde Is. Tetranychus sp. (Acarina: Tetranychidae). 

Feltiella ligulata Gagné 1995a: 26. Cape Verde Is: Ribeira de Mangue; HT; ♂; BMNH. 

occidentalis. USA (California), Japan (Honshu). Tetranychus sp.; many tetranychids (Acarina). 

Cecidomyia acarivora Felt 1907b: 242, jun. sec. hom. acarivora Zehntner. USA: California, Chula Vista; LT (Gagné 

1995a: 27); ♂; NYSM. 

Arthrocnodax occidentalis Felt 1912j: 402. USA: California, Whittier; LT (Gagné 1995a: 27); ♂; NYSM. 

pini. North America (widespread); Bahamas, Nicaragua, Australia (New South Wales). Tetranychus sp.; many 

tetranychids (Acarina). 

Mycodiplosis pini Felt 1907a: 31. USA: New York, Albany; HT; ♂; NYSM. 

Mycodiplosis acerifoliae Felt 1907a: 31. USA: New York, Albany; HT; ♂; NYSM. 

Rhabdophaga borealis Felt 1907a: 17. USA: New York, Lake Clear; HT; ♀; NYSM. 

Feltiella acerifolia Felt 1907c: 127, unjust. emend. of acerifoliae. 

Mycodiplosis spinosa Felt 1911l: 455, nomen nudum. 

Mycodiplosis spinosa Felt 1911n: 550. USA: California, Los Angeles; LT (Gagné 1995a: 27); ♂; USNM. 

Mycodiplosis macgregori Felt 1915b: 149. USA: South Carolina, Batesburg; HT; ♂; NYSM. 

reducta. USA (New York). 

Mycodiplosis reducta Felt 1908b: 400. USA: New York, Karner; HT; ♂; NYSM. 

tetranychi. Germany, Czechoslovakia, Ukraine, Bulgaria. Tetranychus telarius (Tetranychidae; Acarina). This species 

is possibly synonymous with F. acarisuga Vallot. 

Arthrocnodax tetranychi Kieffer 1908: 201. "Southern Russia"; ST; ♂,L; lost. 

Acaroletes tetranychorum Kieffer 1913i: 201, unnec. new name for Arthrocnodax tetranychi Kieffer. 

FICIOMYIA 

Ficiomyia Felt 1922b: 5. Type species, perarticulata Felt (orig. des.). 

Neotropical; 1 sp.; Lasiopteridi, unplaced to tribe. Ref.: Roskam & Nadel 1990 (redescr.). 

perarticulata. USA (Florida), Guadeloupe. Ficus citrifolia (Moraceae). 

Ficiomyia perarticulata Felt 1922b: 5. USA: Florida, Miami; LT (Roskam & Nadel 1990: 785); ♂; USNM. 

Ficiomyia birdi Felt 1934b: 132. USA: Florida, Dade Co., Royal Palm St. Pk; LT (Roskam & Nadel 1990: 785); ♂; 


*FILIDIPLOSIS 

Filidiplosis Fedotova 2003e: 49. Type species, mollis Fedotova (orig. des.). 

Eastern Palearctic; 4 spp.; Cecidomyiidi, unplaced to tribe. Fedotova and Sidorenko 2009a propose the new tribe 

Filidiplosini to include this genus and four others, viz., Exiguidiplosis, Sicituradastra, Flexipidiplosis, and Rotadiplosis. 

The abdominal terga described for these genera are so different, from entire to laterally dissected, from strongly 

pigmented to weakly so, and the tarsal claws as drawn are of such a variety, that the few superficial similarities outlined 

for the genitalia of these genera do not make a convincing argument that they comprise a monophyletic assemblage. 

*admixta. Russia (Far East).

229 

Gybbosidiplosis admixta Fedotova 2006a: 143. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*laesa. Russia (Far East). 

Gybbosidiplosis laesus Fedotova 2003e: 46. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*mollis. Russia (Far East). 

Filidiplosis mollis Fedotova 2003e: 49. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

*vasta. Russia (Far East). 

Filidiplosis vasta Fedotova & Sidorenko 2009a: 28. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya Bay; 


*FIMBRIATODIPLOSIS 

Fimbriatodiplosis Fedotova 2004g: 20. Type species, calcea Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Karshomyiini. This genus is close to Karshomyia if the two must be 

separated. 

*calcea. Russia (Far East). 

Fimbriatodiplosis calceus Fedotova 2004g: 21. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*FISSURATIDIPLOSIS 

Fissuratidiplosis Fedotova 2004h: 170. Type species, stylaris Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Mycodiplosini. This genus does not essentially differ from Mycodiplosis. 

*stylaris. Russia (Far East). 

Fissuratidiplosis stylaris Fedotova 2004h: 170. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*FLEXIPIDIPLOSIS 

Flexipidiplosis Fedotova & Sidorenko 2009a: 32. Type species, flustra Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe here. Fedotova and Sidorenko 2009a place this genus in 

their tribe Filidiplosini (see Appendix). 

*flustra. Russia (Far East). 

Flexipidiplosis flustra Fedotova & Sidorenko 2009a: 32 (erroneously as Rotadiplosis flustra, p. 38). Russia: 

Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; ♂; ZMAS. 

*FOVEOLADIPLOSIS 

Foveoladiplosis Fedotova 2004g: 16. Type species, capitulifera Fedotova (orig. des.). 


separated. Fedotova and Sidorenko 2009a propose the new subtribe Foveoladiplosina within the tribe Karshomyini to 

include this genus and Prorsodiplosis. 

*capitulifera. Russia (Far East).

230 

Foveoladiplosis capitulifera Fedotova 2004g: 16. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

FRAUENFELDIELLA 

Frauenfeldiella Rübsaamen 1905b: 122. Type species, coussapoae Rübsaamen (orig. des. as n. g., n. sp.). 


coussapoae. Brazil (Amazonas, Rio de Janeiro). Coussapoa sp. (Moraceae). 

Frauenfeldiella coussapoae Rübsaamen 1905b: 122. Brazil: Amazonas, Juruà Miry, & RJ, Gavea; ST; ♀,L; ZMHB. 

*GALATELLOMYIA 

Galatellomyia Fedotova 2003a: 825. Type species, asiatica Fedotova (orig. des.). 


*asiatica. Kazakhstan. Galatella punctata; G. chromopappus (Asteraceae). 

Galatellomyia asiatica Fedotova 2003a: 825. Kazakhstan: northwestern Dzhungarskiï Alatau, Lepsy R., nr 

Petropavlovka-Cherkasskaya; HT; ♂; ZMAS. 

*GALEIDIPLOSIS 

Galeidiplosis Fedotova 2005d: 230. Type species, quadricoxita Fedotova (orig. des.). 

Palearctic; 2 spp.; Cecidomyiidi, Mycodiplosini. This genus does not essentially differ from Coquillettomyia. 

*pluridens. Russia (Far East). 

Galeidiplosis pluridens Fedotova & Sidorenko 2005e: 231. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


*quadricoxita. Russia (Far East). 

Galeidiplosis quadricoxita Fedotova 2005d: 230. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

*GARUGADIPLOSIS 

Garugadiplosis Deshpande, Shaikh & Sharma 2002c: 111. Type species, brevipalpis Deshpande, Shaikh & Sharma 

(orig. des.). 

Oriental; 1 sp.; Cecidomyiidi, Cecidomyiini. 

*brevipalpis. India (Maharashtra). Garuga pinnata (Burseraceae). 

Garugadiplosis pinnata Deshpande, Shaikh & Sharma 2002c: 112. India: Maharashtra, Nanded Distr., Bhokar; HT; 

♂; Deshpande Collection. 

GEOCRYPTA 

Geocrypta Kieffer 1913i: 68. Type species, Cecidomyia braueri Handlirsch (mon.). 

Palearctic; 6 spp.; Lasiopteridi, Dasineurini. Ref.: Fedotova 1997b (rev.).

231 

braueri. Widespread Europe. Hypericum perforatum; H. pulchrum (Hypericaceae). 

Cecidomyia braueri Handlirsch 1884: 135. Czech Republic: Vranov nr Znojmo; ST; ♂,♀; ?. 

Geocrypta hypericina Tavares 1919a: 41. Spain: Pontevedra; ST; ♂,♀,L; TAVC. 

campanulae. UK, Sweden, Denmark, Germany, Poland, Czechoslovakia, Austria, Romania, Yugoslavia. Campanula 

rotundifolia (Campanulaceae). 

Cecidomyia campanulae Müller 1871a: viii. UK: Scotland, Aberdeen; ST; LT; lost. This name has not been used 

since 1871 but the description of the gall is similar to that formed by C. trachelii. 

Cecidomyia trachelii Wachtl 1885: 195. Austria: nr Aggsbach on Donau R.; ST; ♀,L; NHMW?. 

galii. Widespread Europe, Kazakhstan, Russia (Far East). Galium uliginosum, G. mollugo; G. verum (Rubiaceae). 

Cecidomyia galii Loew 1850a: 37. Poland; ST; ♂,♀; ZMHB?. 

Cecidomyia molluginis Loew 1850a: 30. Poland; ST; G; ZMHB?. 

Cecidomyia galii Winnertz 1853: 235. Germany; ST; ♂,♀,L; lost. 

heterophylli. Germany, Yugoslavia, Armenia. Lathyrus heterophyllos, L. platyphyllos (Fabaceae). 

Dasyneura heterophylli Rübsaamen 1914: 105. Germany: Munich, & Yugoslavia: Dalmatia; ST; ♂; ZMHB. 

rostriformis. UK, Kazakhstan. Galium verum; Galium spp. (Rubiaceae). 

Geocrypta rostriformis Fedotova 1997b: 51. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Baldabrek Mtn 

Range, Chooldak area, 57 km NE Georgievka, 7 km SE Baldabrek; HT; ♂; ZMAS. 

spongiosa. Kazakhstan. Galium verum (Rubiaceae). 

Geocrypta spongiosa Fedotova 1997b: 49. Kazakhstan: Trans-Ili Alatau, Oizhailyau area, Turgen' canyon, 36 km S 

Turgen'; HT; ♂; ZMAS. 

*GEOMYIA 

Geomyia Skuhravá 2006: 108. Type species, alpina Skuhravá (orig. des.). 


*alpina. Switzerland. Geum reptans (Rosaceae). 

Geomyia alpina Skuhravá 2006: 109. Switzerland: 15 km S Davos; HT; ♀; SKCR. 

GEPHYRAULUS 

Gephyraulus Rübsaamen 1916b: 490. Type species, Cecidomyia raphanistri Kieffer (orig. des.). 

Paragephyraulus Solinas 1982: 318. Type species, diplotaxis Solinas (orig. des.). 

Lepidiomyia Fedotova 1992j: 669. Type species, sylveni Fedotova (orig. des.). 

Spirorhynchomyia Fedotova 1992j: 670. Type species, solinasi Fedotova (orig. des.). 

Lepidomyia Fedotova 1992j (in 1993 translation only): 13, missp. of Lepidiomyia. 

Palearctic and Neotropical (introd.); 11 spp., nine on Brassicaceae, two on mango (Anacardiaceae); Lasiopteridi, 

Dasineurini. Refs: Sylvén & Solinas 1989 (review); Sylvén & Tastás-Duque 1993 (review); Gagné & Etienne 2006: 

931 (addition of spp. on mango). Gagné 2004b (syn. notes). 

diplotaxis. Italy, Malta. Diplotaxis muralis; D. erucoides. (Brassicaceae). 

Paragephyraulus diplotaxis Solinas 1982: 320. Italy: Apulia, Bari; ST; ♂,♀,P,L; Solinas Coll. 

erysimi. Kazakhstan. Erysimum marschallianum (Brassicaceae). 

Gephyraulus erysimi Fedotova 1992j: 665. Kazakhstan: 45 km NW Panfilov, floodplain Borakhudzir River; HT; ♂; 

ZMAS. 

indica. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Procystiphora indica Grover & Prasad 1966a: 4. India: Uttar Pradesh, Lucknow; HT; ♂; PGCA. Ref.: Gagné & 

Etienne 2006: 936 (n. comb.). 

isatidis. Kazakhstan. Isatis violascens (Brassicaceae). 

Gephyraulus isatidis Fedotova 1992j: 665. Kazakhstan: southern L. Balkhash region, 40 km NW Kolshengel'; HT; ♂;

232 

ZMAS. 

*mangiferae. India (Tamil Nadu); immigr.: Hawaii, Puerto Rico, Guadeloupe. Mangifera indica (Anacardiaceae). 

Dasyneura mangiferae Felt 1927a: 1. India: Tamil Nadu, Coimbatore; ST; ♀; NYSM. Ref.: Gagné & Etienne 2006: 

933 (n. comb., redescr., immigr. to New World). 

moricandiae. Tunisia. Moricandia arvensis (Brassicaceae). 

Gephyraulus moricandiae Sylvén & Solinas 1989: 23. Tunisia: Sousse; HT; ♀; NHRS. 

raphanistri. Widespread Europe. Raphanistrum silvestre (Brassicaceae). 

Cecidomyia raphanistri Kieffer 1886: 324. France: Metz, Boulay, Sarraguemines, Bitche; ST; ♂,♀,L; lost?. 

sisymbrii. Kazakhstan. Sisymbrium loeselii (Brassicaceae). 

Gephyraulus sisymbrii Fedotova 1992j: 667. Kazakhstan: southern L. Balkhash region, 40 km NW Kolshengel'; HT; 

♂; ZMAS. 

solinasi. Kazakhstan. Spirorhynchus sabulosus (Brassicaceae). 

Spirorhynchomyia solinasi Fedotova 1992j: 671. Kazakhstan: Moïynkumy Desert, 30 km SW Ulan-Bel'; HT; ♂; 

ZMAS. 

strigosellacola. Kazakhstan. Strigosella circinnata; S. turkestanica (Brassicaceae). 

Lepidiomyia strigosellacola Fedotova 1994a: 127. Kazakhstan: Kyzylkum Desert, left bank Syr-Darya R., 30 km S 

Yanykurgan, 24 km SW Kyrkense; HT; ♂; ZMAS. 

sylveni. Kazakhstan. Lepidium sibiricum (Brassicaceae). 

Lepidiomyia sylveni Fedotova 1992j: 669. Kazakhstan: Ili R. delta, Ketmen'-Kaldy branch of Zhideli R., Bayguti; HT; 

♂; ZMAS. 

GERALDESIA 

Geraldesia Tavares 1917b: 134. Type species, eupatorii Tavares (orig. des.). 

Neotropical; 3 spp.; Lasiopteridi, Alycaulini. 

cumbrensis. El Salvador. Baccharis trinervis (Asteraceae). 

Geraldesia cumbrensis Möhn 1975: 5. El Salvador: Santa Ana, Hacienda San José, nr San José; HT; ♂; SMNS. 

eupatorii. Brazil. Eupatorium sp. (Asteraceae). 

Geraldesia eupatorii Tavares 1917b: 134. Brazil: Rio de Janeiro; ST; ♀,P; TAVC. 

polygonarum. Colombia. Triplaris/Symmeria/Ruprechtia sp. (Polygonaceae). 

Geraldesia polygonarum Wünsch 1979: 111. Colombia: Magdalena, Cerro San Fernando; HT; ♂; SMNS. 

GERANIOMYIA 

Geraniomyia Fedotova 1991d: 24. Type species, Lathyromyza geraniicola Fedotova (orig. des.). 


geraniicola. Kazakhstan. Geranium collinum (Geraniaceae). 

Lathyromyza geraniicola Fedotova 1984c: 47. Kazakhstan: Zailiïskiï Alatau, Sredniï Talgar, 40 km SE Alma-Ata; 


GEROMYIA 

Geromyia Coutin & Harris 1969: 259. Type species, Itonida penniseti Felt (orig. des.). 

Afrotropical, Oriental, Eastern Palearctic; 3 spp.; Cecidomyiidi, unplaced to tribe. 

akolaensis. India (Maharashtra). 

Geromyia akolaensis Grover & Bakhshi 1978: 122. India: Maharashtra, Akola; HT; ♂; PGCA.

233 

nawai. Japan (Honshu to Kyushu). Arundinaria simonii (Poaceae). 

Clinodiplosis nawai Monzen 1937: 182. Japan: Honshu, Aici Pref.; ST; ♂,♀; KUEC. 

penniseti. Senegal, Ghana, Nigeria, Sudan, Uganda, Madagascar, India (Tamil Nadu). Pennisetum cenchroides; P. 

typhoideum (Poaceae). 

Itonida penniseti Felt 1920b: 9. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

Cecidomyia penniseti Felt 1920b: 11. India: Tamil Nadu, Coimbatore; HT; ♀; NYSM. 

Itonida seminis Felt 1921j: 26. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

Cecidomyia pennisetoides Barnes 1956: 188, new name for Cecidomyia penniseti Felt. 

GIARDOMYIA 

Giardomyia Felt 1908b: 405. Type species, Cecidomyia photophila Felt (orig. des.). 

Holarctic, Oriental, Oceanian; 12 spp.; Cecidomyiidi, unplaced to tribe. 

britannica. UK, Netherlands, Czechoslovakia, Russia (Europe). Trifolium repens, T. pratense (Fabaceae). 

Giardomyia britannica Milne 1960: 99. UK: Kent, Shropshire, Montgomeryshire, Cardiganshire; ST; ♂,♀; BMNH. 

emarginata. USA (New York). 

Cecidomyia emarginata Felt 1907a: 38. USA: New York, Albany; HT; ♂; NYSM. 

fragariae. USA (New York). 

Cecidomyia fragariae Felt 1907a: 37. USA: New York, Albany; HT; ♂; NYSM. 

furvescens. Hawaiian Is. 

Giardomyia furvescens Hardy 1960: 287. Hawaiian Is: Oahu, Ewa; HT; ♂; HDOA. 

hudsonica. USA (New York). 

Giardomyia hudsonica Felt 1908b: 406. USA: New York, Poughkeepsie; HT; ♂; NYSM. 

Giardomyia montana Felt 1908b: 406. USA: New York, Newport & New York, Staten I.; ST; ♂; NYSM. 

Cecidomyia ruricola Felt 1908b: 413. USA: New York, Nassau; HT; ♂; NYSM. 


Giardomyia indica Grover & Bakhshi 1978: 131. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; PGCA. 

menthae. USA (New York). Mentha piperita (Lamiaceae). 

Giardomyia menthae Felt 1908b: 405. USA: New York, Karner; HT; ♂; NYSM. 

pallidithorax. Hawaiian Is. 

Giardomyia pallidithorax Hardy 1960: 288. Hawaiian Is: Oahu, Ewa; HT; ♂; HDOA. 


Cecidomyia phosphila Felt 1907a: 37. USA: New York, Newport; HT; ♂; NYSM. 

Cecidomyia photophila Felt 1907c: 134, unjust. emend. of phosphila. 

Giardomyia emarginata Felt 1908b: 405. USA: New York, Newport; HT; ♂; NYSM. 

Clinodiplosis extensa Felt 1908b: 412. USA: New York, Albany; HT; ♂; NYSM. 

Giardomyia noveboracensis Felt 1908b: 405. USA: New York, Nassau; HT; ♂; NYSM. 

piperitae. USA (New York). 

Cecidomyia piperitae Felt 1907d: 22. USA: New York, Nassau; HT; ♀; NYSM. 

sesami. India (Madhya Pradesh). Sesamum orientale (Pedaliaceae). 

Giardomyia sesami Grover & Bakhshi 1978: 127. India: Madhya Pradesh, Raipur; HT; ♂; PGCA. 

tripurensis. India (Tripura). 

Giardomyia tripurensis Chandra 1989: 106. India: Tripura, Agartala; HT; ♂; CSIA. 

*GIGANTODIPLOSIS 

Gigantodiplosis Fedotova 2004g: 23. Type species, pilosa Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe here. Fedotova and Sidorenko 2009a proposed the new tribe 

Gigantodiplosini for this genus alone and include the tribe in their supertribe Coquillettomyiidi. Neither the tribe or

supertribe is used here (see Appendix). 

*pilosa. Russia (Far East). 

Gigantodiplosis pilosa Fedotova 2004g: 23. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

GIRAUDIELLA 

Giraudiella Rübsaamen 1916b: 489. Type species, Cecidomyia inclusa Frauenfeld (orig. des.). 

Palearctic; 2 spp.; Lasiopteridi, Dasineura. Ref.: Skuhravá & Skuhravý 1981 (bionomics, descr.). 

inclusa. Widespread Europe, Iraq, China; immigr.: USA (eastern). Phragmites australis (Poaceae). 

Cecidomyia inclusa Frauenfeld 1862: 1175. Austria; ST; ♂,♀; ?. 

Giraudiella incurvans Nijveldt 1953b: 282. Netherlands: Guelderland, nr Winterswijk; HT; ♂; ZMAN. 

scirpicola. Russia (Europe). Scirpus silvaticus (Cyperaceae). 

Giraudiella scirpicola Mamaeva 1964a: 211. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; ?. 

GLADIODIPLOSIS 

Gladiodiplosis Kolesik 2000c: 250. Type species, Cecidomyia frenelae Skuse (orig. des.). 


frenelae. Australia (New South Wales). Callitris endlicheri (Cupressaceae). 

Cecidomyia frenelae Skuse 1890: 377. Australia: New South Wales, Wagga Wagga Distr.; HT; ♂; ANIC. 

GLENODIPLOSIS 

Glenodiplosis Gagné 1973c: 870. Type species, callipus Gagné (orig. des.). 


callipus. USA (Virginia). 

Glenodiplosis callipus Gagné 1973c: 870. USA: Virginia, Falls Church, Holmes Run; HT; ♂; USNM. 

*GLIASPILOTA 

Gliaspilota Gagné 2008c: 93. Type species, glutinosa Gagné (orig. des.). 

Nearctic; 1 sp., from sticky leaf spot on Carya (Juglandaceae); Lasiopteridi, unplaced to tribe. 

glutinosa. Eastern Nearctic. Carya tomentosa; Carya spp. (Juglandaceae). Ref.: Gagné 2008c: 95 (redescr.). 

Cecidomyia glutinosa Osten Sacken 1862: 193. USA: District of Columbia; ST; L, G; USNM. 

*GLOMERULIDIPLOSIS 

Glomerulidiplosis Fedotova 2003e: 53. Type species, crenata Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Clinodiplosini. This genus is close to Clinodiplosis, if the two must be 

separated. Fedotova & Sidorenko (2009d: 254) erect the subtribe Glomerulidiplosina for this monotypic genus alone, 

but that subdivision is not used here (see Appendix under Clinodiplosini). 

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*crenata. Russia (Far East). 

Glomerulidiplosis crenatus Fedotova 2003e: 54. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

GNESIODIPLOSIS 

Gnesiodiplosis Tavares 1917b: 162. Type species, itaparicae Tavares (orig. des.). 

Gneciodiplosis Shinji 1944: 252, missp. of Gnesiodiplosis. 


itaparicae. Brazil (Bahia). Undet. sp. of Rubiaceae. 

Gnesiodiplosis itaparicae Tavares 1917b: 162. Brazil: Bahia, Salvador; ST; ♂,♀; TAVC. 

GOBIDIPLOSIS 

Gobidiplosis Mamaev 1972a: 896. Type species, kandybinae Mamaev (orig. des.). 

Cobidiplosis Fedotova 1992i: 45, missp. of Gobidiplosis. 

Ilidiplosis Fedotova 1992i: 45. Type species, mamaevi Fedotova (orig. des.). 

Palearctic; 2 spp.; Cecidomyiidi, Cecidomyiini. This genus is very similar to Contarinia except for the pronounced 

swelling at the base of the ovipositor. Both species make similar galls on Reaumuria soongarica. Ref.: Gagné 2004b 

(syn. note). 

kandybinae. Mongolian People's Republic. Reaumuria soongarica (Tamaricaceae). 

Gobidiplosis kandybinae Mamaev 1972a: 897. Mongolian People's Republic: South Gobi Admin. Distr., Bulgana; 


mamaevi. Kazakhstan. Reaumuria soongarica (Tamaricaceae). 

Ilidiplosis mamaevi Fedotova 1992i: 46. Kazakhstan: north shore middle course Ili R., 40 km NE Chilik; HT; ♂; 

ZMAS. 

GONGRODIPLOSIS 

Gongrodiplosis Gagné 1973c: 870. Type species, Mayetiola latipes Felt (orig. des.). 


latipes. USA (New York). 

Mayetiola latipes Felt 1908b: 370. USA: New York, Albany; HT; ♂; NYSM. 

*GUAREAMYIA 

Guareamyia Maia 2007a: 450. Type species, purpura Maia (orig. des.). 

Neotropical; 1 sp.; Lasiopteridi, unplaced to tribe. See notes under Guarephila about possible placement of this 

genus. 

*purpura. Brazil (São Paulo). Guarea macrophylla (Meliaceae). 

Guareamyia purpura Maia 2007a: 450. Brazil: São Paulo, Bertioga, Itaguaré; HT; ♂; QBUM. 

GUAREPHILA 

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Guarephila Tavares 1909: 18. Type species, albida Tavares (mon.). 

Neotropical; 1 sp.; Lasiopteridi, unplaced to tribe. Maia 2007a described two new genera, Guareamyia and 

Sphaeramyia, of one species each that show distinct similarities with Guarephila and are separated from it by 

seemingly superficial characters. The three species form spherical leaf galls on Guarea. 

albida. Brazil (Rio Grande do Sul). Guarea sp. poss. trichilioides (Meliaceae). 

Guarephila albida Tavares 1909: 18. Brazil: Rio Grande do Sul, vic. São Leopoldo; ST; ♂,♀; TAVC. 

GUIGNONIA 

Guignonia Kieffer 1913a: 219. Type species, potentillae Kieffer (orig. des.). 

Guigonia Shinji 1944: 226, missp. of Guignonia. 


potentillae. France. Potentilla verna (Rosaceae). 

Guignonia potentillae Kieffer 1913a: 219. France; T; ♂; lost?. 

Guignonia potentillarum Kieffer 1913i: 42, unjust. emend. of potentillae. 

*GYBBOSIDIPLOSIS 

Gybbosidiplosis Fedotova 2003e: 44. Type species, matutina Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Mycodiplosini. 

*matutina. Russia (Far East). 

Gybbosidiplosis matutinus Fedotova 2003e: 46. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

GYNANDROBREMIA 

Gynandrobremia Mamaev 1965c: 679. Type species, viciae Mamaev (orig. des.). 

Palearctic; 5 spp.; Cecidomyiidi, unplaced to tribe. This genus should be compared to Resseliella. 

*maackiaefloris. Russia (Far East). Maackia amurensis (Fabaceae). 

Gynandrobremia maackiaefloris Fedotova 2002c: 12. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; 


*populicola. Russia (Far East). Populus koreana (Salicaceae). 

Gynandrobremia populicola Fedotova 2002c: 8. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

salicicola. Latvia, Russia (Europe). 

Gynandrobremia salicicola Mamaev 1965c: 680. Russia: Yaroslavl Prov., Volkovo; HT; ♂; HYNM. 

*salicifoliae. Russia (Far East). Salix nipponica (Salicaceae). 

Gynandrobremia salicifoliae Fedotova 2003c: 8. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

viciae. Russia (Europe). 

Gynandrobremia viciae Mamaev 1965c: 680. Russia: Yaroslavl Prov., Volkovo; HT; ♂; HYNM. 

GYNODIPLOSIS 

236

Gynodiplosis Felt 1927c: 383. Type species, humata Felt (orig. des.). 


humata. Indonesia (Java). Davallia alpina (Davalliaceae). 

Gynodiplosis humata Felt 1927c: 383. Indonesia: Java, Tjibodas, Mt Gede; ST; ♂,♀; NYSM. 

GYPSOPHYLLOMYIA 

Gypsophyllomyia (also as Gypsophillomyia) Fedotova 1991d: 19. Type species, Wachtliella gypsophilae Fedotova 

(orig. des.). 


gypsophilae. Kazakhstan. Gypsophila trichotoma (Caryophyllaceae). 

Wachtliella gypsophilae Fedotova 1983c: 25. Kazakhstan: upper Ili R., Ketmen'-Kaldy canal, 15 km NW Koktal; HT; 

♂; ZMAS. 

Gypsophyllomyia gypsophylae Fedotova 1991d: 20, missp. of gypsophilae. 

HADROBREMIA 

Hadrobremia Kieffer 1912h: 1. Type species, Clinodiplosis longiventris Kieffer (orig. des.). 

Hadrobremyia Fedotova & Kovalev 2001: 423, 598, missp. of Hadrobremia. 

Palearctic; 1 sp.; Cecidomyiidi, Clinodiplosini. 

longiventris. UK, France, Germany, Russia (Western Siberia). Trifolium medium (Fabaceae). 

Clinodiplosis longiventris Kieffer 1901a: pl. 21, fig. 8 (as n. sp. also in Kieffer 1909b: 34). France: château de La 

Grange, nr. Thionville; T; ♀; lost?. 

Clinodiplosis trifolii Kieffer 1909b: 31. France: château de La Grange, nr. Thionville; ST; LT; lost?. 

HALODIPLOSIS 

Halodiplosis Kieffer 1912e: 78. Type species, salsolae Kieffer (orig. des. as n. g., n. sp.). 

Onodiplosis Felt 1916e: 175. Type species, sarcobati Felt (orig. des.). 

Asidiplosis Marikovskij 1953a: 136, nomen nudum. 

Asiodiplosis Marikovskij 1955a: 336, invalid, described without type species designation. 

Tyloceramyia Marikovskij 1955b: 105, nomen nudum. 

Haloxylonophaga Marikovskij 1955b: 105, nomen nudum. 

Asiodiplosis Marikovskij 1955d: 303. Type species, noxia Marikovskij (orig. des.). 

Haloxylophaga Marikovskij 1955d: 298. Type species, inornata Marikovskij (orig. des.). 

Tyloceramyia Marikovskij 1956: 192. Type species, steinbergi Marikovskij (orig. des.). 

Monarthropselaphus Marikovskij 1957b: 939. Type species, hebes Marikovskij (orig. des.). 

Halocnemomyia Marikovskij 1965: 80. Type species, schnitnikovi Marikovskij (orig. des.). 

Desertomyia Marikovskij 1975: 54. Type species, dissimmetrica Marikovskij (mon.). 

Micrasiodiplosis Mamaeva 1980: 45. Type species, constricta Mamaeva (orig. des.). 

Bojalodiplosis Fedotova 1985a: 48. Type species, arbocarpis Fedotova (orig. des.). 

Gemmidiplosis Fedotova 1992c: 77, as subg. of Halodiplosis. Type species, Halodiplosis aurantiaca Fedotova (orig. 

des.). 

Holarctic, mainly Palearctic; 99 spp., all forming bud galls on Chenopodiaceae and Fabaceae; Cecidomyiidi, 

237

238 

Cecidomyiini. Halodiplosis appears to be a monophyletic group, comprising species that have either two or three 

circumfila on the male flagellomeres. Ref.: Gagné 2004b (syn. notes). 

aestivas. Kazakhstan. Salsola laricifolia (Chenopodiaceae). 

Asidiplosis aestivas Marikovskij 1953a: 138. Kazakhstan: Chulak Mts, middle course Ili R.; ST; ♂,♀; ZKAS & 

ZMAS. 

Asiodiplosis aestivus Marikovskij 1955b: 104, missp. of aestivas. 

anabasidicola. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis anabasidicola Fedotova 1989f: 826. Kazakhstan: N shore L. Balkhash, 16 km SE of Burylbaytal; HT; 

♀; ZKAS. 

anabasidigemmae. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis anabasidigemmae Fedotova 1989f: 824. Kazakhstan: Karatal R. valley, 70 km NW Ush-Tobe; HT; ♂; 

ZKAS. 

anabasidis. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis anabasidis Fedotova 1989f: 828 (as n. sp. also in Fedotova 1990a: 228). Kazakhstan: north shore L. 

Balkhash, 16 km SE Burylbaytal; HT; ♂; ZMAS. 

anabasidia. Kazakhstan. Anabasis aphylla (Chenopodiaceae). 

Bojalodiplosis anabasidis Fedotova 1989f: 828 (as n. sp. also in Fedotova 1990a: 231), new jun. sec. hom. 

Halodiplosis anabasidis Fedotova. Kazakhstan: 20 km NW Zhalanash; HT; ♂; ZMAS. 

Halodiplosis anabasidia Gagné 2004b: 158, new name for Bojalodiplosis anabasidis Fedotova. 

aphylla. Kazakhstan. Anabasis aphylla (Chenopodiaceae). 

Halodiplosis aphylla Fedotova 1992c: 73. Kazakhstan: Mangyshlak Mtn, 10 km NE Kuybyshevo, nr Shair; HT; ♂; 

ZMAS. 

araratica. Armenia. Salsola ericoides (Chenopodiaceae). 

Halodiplosis araratica Mirumian 1991a: 45. Armenia: Ararat Reg., nr Urtsadzor; HT; ♂; ASAY. 

arbocarpis. Kazakhstan. Salsola arbusculaeformis (Chenopodiaceae). 

Bojalodiplosis arbocarpis Fedotova 1985a: 48. Kazakhstan: Karatal R. valley, 21 km NW Ush-Tobe; HT; ♂; ZKAS. 

aristiformis. Kazakhstan. Salsola arbuscula (Chenopodiaceae). 

Halodiplosis aristiformis Fedotova 1993m: 21. Kazakhstan: northeastern Kyzlkumy, 30 km S Yanykurgana, 24 km 

SW Kyrkense; HT; ♂; ZMAS. 

arthrophytumis. Kazakhstan. Arthrophytum iliense (Chenopodiaceae). 

Halodiplosis arthrophytumis Fedotova 1992e: 46. Kazakhstan, north end Kuluktau Mts, Timirlik R. floodplain, 45 

km SE Chundzha; HT; ♂; ZMAS. 

arthrophytumidis. Kazakhstan. Arthrophytum longibracteatum (Chenopodiaceae). 

Bojalodiplosis arthrophytumis Fedotova 1992e: 49. Kazakhstan: Sengir-Kum Desert, Kyzylodyr landmark, 60 km 

NE Senek; HT; ♀; ZMAS. 

Halodiplosis arthrophytumidis Gagné 2004b: 158, new name for Bojalodiplosis arthrophytumis Fedotova. 

aurantiaca. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis aurantiacus Fedotova 1992c: 78. Kazakhstan: Usturt Plateau, 155 km NE Senek; HT; ♂; ZMAS. 

bacilliformis. Kazakhstan. Salsola richteri (Chenopodiaceae). 

Halodiplosis bacilliformis (also as baciliformis) Fedotova 1991g: 36. Kazakhstan: northeastern Kyzylkum Desert, nr 

Karaktau Mtn, 64 km NW Bairkum; HT; ♂; ?. 

balchashensis. Kazakhstan. Anabasis eriopoda (Chenopodiaceae). 

Halodiplosis balchashensis Fedotova 1993e: 83. Kazakhstan: SW shore L. Balkhash, 8 km S Chiganak; HT; ♂; 

ZMAS. 

bassiae. Turkmenistan. Bassia hyssopifolia (Chenopodiaceae). 

Halodiplosis bassiae Fedotova 1993h: 51. Turkmenistan: northern Kopetdag, Sumbar R., nr Kara-Kala; HT; ♂; 

ZMAS. 

biennis. Kazakhstan, Turkmenistan. Haloxylon ammodendron, H. aphyllum (Chenopodiaceae). 

Haloxylophaga biennis Marikovskij 1955d: 302. Kazakhstan: Ili and Syr-Darya Rivers; ST; ♂,♀; ?. 

botryoidea. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Halodiplosis botryoideus Fedotova 1993e: 87. Turkmenistan: eastern Karakumy Desert, Repetek Nature Res., 70 km

239 

SW Chardzhou; HT; ♂; ZMAS. 

buzaczensis. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis buzaczensis Fedotova 1993c: 106. Kazakhstan: Buzachi Peninsula, 75 km SW Shebir, Tomb of Aksyn; 


caspica. Kazakhstan. Salicornia europaea (Chenopodiaceae). 

Bojalodiplosis caspicus Fedotova 1990c: 26. Kazakhstan: northwestern Ustyurt, Kumin L., floodplain Karasaï R., 90 

km NW Sam; HT; ♂; ZMAS. 

caspicana. Kazakhstan. Anabasis eriopoda; A. hispidula (Chenopodiaceae). 

Halodiplosis caspicus Fedotova 1993c: 110, jun. sec. hom. caspica Fedotova. Kazakhstan: Buzachi Peninsula, 

Suyursha, 12 km SE Shebir; HT; ♂; ZMAS. 

Halodiplosis caspicana Gagné 2004b: 159, new name for Halodiplosis caspica Fedotova. 

ceratoidis. Kazakhstan. Ceratoides papposa (Chenopodiaceae). 

Halodiplosis ceratoidis Fedotova 1994f: 31. Kazakhstan: middle course Ili R., 20 km NE Chilik, nr Buryndysu ; HT; 

M ; ZMAS. 

climacopterae. Kazakhstan. Climacoptera lanata (Chenopodiaceae). 

Halodiplosis climacopterae Fedotova 1993e: 81. Kazakhstan, middle course Ili R., 20 km NW Bakanas; HT; ♂; 

ZMAS. 

conmixta. Turkmenistan. Salsola richteri (Chenopodiaceae). 

Halodiplosis conmixta Mamaev & Pak 1990: 58. Turkmenistan: Tashauz Prov., Chiryshlya; HT; ♂; HYNM. 

consociata. Kazakhstan. Haloxylon ammodendron, H. aphyllum (Chenopodiaceae). 

Haloxylonophaga consociata Marikovskij 1955b: 105, nomen nudum. 

Haloxylonophaga consociata Marikovskij 1955d: 301. Kazakhstan: Ili and Syr-Darya Rivers; ST; ♂; ?. 

constricta. Turkmenistan, Kazakhstan. Anabasis aphylla (Chenopodiaceae). 

Micrasiodiplosis constricta Mamaeva 1980: 45. Kazakhstan: Chimkent Reg., Yermolovka; HT; ♂; HYNM. 

czuensis. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis czuensis Fedotova 1993c: 108. Kazakhstan: Chu R. valley, 10 km SE Ulan-Bel'; HT; ♂; ZMAS. 

densipila. Turkmenistan. Salsola rigida (Chenopodiaceae). 

Tyloceramyia densipila Marikovskij 1957: 938. Turkmenistan: Tashauz Prov., Kuna Urgench; ST; ♂,♀; MARC. 

deserta. Kazakhstan, Turkmenistan. Salsola arbuscula (Chenopodiaceae). 

Asiodiplosis deserta Marikovskij 1955a: 341. Kazakhstan: Kzyl-Orda, lower Syr-Darya R., and Sorgali Desert, 

southwestern Turkmenistan; ST; ♂,♀; ZMAS. 

dissimmetrica. Kazakhstan. Halimodendron halodendron (Fabaceae). 

Desertomyia dissimmetrica Marikovskij 1975: 54. Kazakhstan: lower course Ili R, nr Bakanas; ST; ♂,♀; MARC. 

dzhanokmenae. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis dzhanokmenae Fedotova 1993c: 114. Kazakhstan: middle course Ili R., 20 km NW Bakanas; HT; ♂; 

ZMAS. 

eremobia. Turkmenistan. Anabasis sp. (Chenopodiaceae). 

Halodiplosis eremobia Mamaev & Pak 1989: 32. Turkmenistan: Tashauz Prov., Shakhsenem; HT; ♂; HYNM (Yagi 

& Sawada 2000) - ZMMU (Krivosheina & Ozerov 2006). 

fedtschenkovi. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Haloxylophaga fedtschenkovi Marikovskij 1965: 82. Kazakhstan: southwestern shore L. Balkhash; ST; ♂,♀; MARC. 

festinans. Kazakhstan. Haloxylon aphyllum, H. ammodendron (Chenopodiaceae). 

Asiodiplosis festinans Marikovskij 1955b: 104, nomen nudum. 

Asiodiplosis festinans Marikovskij 1955d: 307. Kazakhstan: Ili R.; ST; ♂,♀; ?. 

filipjevi. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halocnemomyia filipjevi Fedotova 1993g: 18. Kazakhstan: 84 km NW Kyzl-Orda, 57 km NW Karaozek; HT; ♂; 

ZMAS. 

floripara. Turkmenistan, Uzbekistan. Haloxylon persicum (Chenopodiaceae). 

Haloxylophaga floripara Mamaev 1972a: 895. Uzbekistan: Bukhara Prov., Kul'dzhuktau; HT; ♂; ZMAS. 

foliosae. Kazakhstan. Salsola foliosa (Chenopodiaceae). 

Halocnemomyia foliosae Fedotova 1993g: 15. Kazakhstan: 18 km NW Zaïsana, nr Karabulak; HT; ♂; ZMAS. 

germinantis. Kazakhstan. Salsola richteri (Chenopodiaceae).

240 

Halodiplosis germinantis Fedotova 1991g: 38. Kazakhstan: northeastern Kyzylkum Desert, 30 km S Yanykurgana; 

HT; ♂; ?. 

halimocnemii. Kazakhstan. Halimocnemis sclerosperma (Chenopodiaceae). 

Halocnemomyia halomocnemii Fedotova 1993g: 17. Kazakhstan: Zaembinskiï basin, 45 km SE Opornaya Stn; HT; 

♂; ZMAS. 

halogetonis. Kazakhstan. Halogeton glomeratus (Chenopodiaceae). 

Bojalodiplosis halogetonis Fedotova 1992f: 49. Kazakhstan: south of L. Zaisan, nr Karasu; HT; ♂; ZMAS. 

hammadae. Turkmenistan. Hammada leptoclada (Chenopodiaceae). 

Halodiplosis hammadae Fedotova 1993n: 13. Turkmenistan: Kugitang Mtn Range, 55 km NE Charshanga; HT; ♂; 

ZMAS. 

hebes. Turkmenistan. Salsola rigida (Chenopodiaceae). 

Monarthropselaphus hebes Marikovskij 1957: 940. Turkmenistan: Tashauz Prov., Kuna Urgench; ST; ♂,♀; MARC. 

hirsuta. Uzbekistan. Salsola gemmascens (Chenopodiaceae). 

Asiodiplosis hirsuta Mamaev 1972a: 896. Uzbekistan: Bukhara Prov., Kul'dzhuktau; HT; ♂; ZMAS. 

hirta. Kazakhstan. Anabasis aphylla (Chenopodiaceae). 

Asiodiplosis hirta Marikovskij 1955a: 338. Kazakhstan: Kzyl-Orda Prov., L. Telekul', & Alma-Ata Prov., Bela- 

Boguty Mts; ST; ♂,♀; ZMAS. 

hodukini. Kazakhstan. Kalidium caspicum (Chenopodiaceae). 

Asiodiplosis hodukini Marikovskij 1965: 83. Kazakhstan: lower Charyn R.; ST; ♂,♀; MARC. 

iliensis. Kazakhstan. Anabasis aphylla (Chenopodiaceae). 

Asiodiplosis iliensis Marikovskij 1955a: 339. Kazakhstan: Sary-Chagan, nr Ili R.; ST; ♂,♀; ZMAS. 

indurentis. Kazakhstan. Salsola arbuscula (Chenopodiaceae). 

Halodiplosis indurentis Fedotova 1993m: 23. Kazakhstan: right shore middle course Ili R., Malye Kalkany, nr 

Poyushcheï Mtn; HT; ♀; ZMAS. 

iljiniae. Kazakhstan. Iljinia regelii (Chenopodiaceae). 

Halocnemomyia iljiniae Fedotova 1993g: 14. Kazakhstan: mouth L. Alakol, 40 km E. Koktum; HT; ♂; ZMAS. 

infestans. Kazakhstan. Haloxylon aphyllum (Chenopodiaceae). 

Haloxylonophaga infestans Marikovskij 1955b: 105, nomen nudum. 

Haloxylophaga infestans Marikovskij 1955d: 300. Kazakhstan: Kyzl-Orda Prov.; ST; ♂,♀; ?. 

inflorescentiae. Kazakhstan. Sympegma regelii (Chenopodiaceae). 

Halodiplosis inflorescentiae Fedotova 1990e: 45. Kazakhstan: 19 km NW Chunzha, 16 km N Aktaï; HT; ♂; ZMAS. 

inornata. Kazakhstan. Haloxylon ammodendron, H. aphyllum (Chenopodiaceae). 

Haloxylonophaga inornata Marikovskij 1955b: 105, nomen nudum. 

Haloxylophaga inornata Marikovskij 1955d: 299. Kazakhstan: Ili R.; ST; ♂,♀; ?. 

insularis. Kazakhstan. Salsola arbuscula (Chenopodiaceae). 

Halodiplosis insularis Fedotova 1993m: 19. Kazakhstan: northern Pre-Aral region, 30 km NW Saksaul'skoï, 98 km 

NW Aral'ska; HT; ♂; ZMAS. 

invisibilis. Kazakhstan. Anabasis truncata (Chenopodiaceae). 

Halodiplosis invisibilis Fedotova 1989f: 823. Kazakhstan: Tarbagatai Mts., upper Ayaguz R., 18 km E Ayaguz; HT; 

♂; ZKAS. 

latentis. Turkmenistan. Salsola sp. (Chenopodiaceae). 

Halodiplosis latentis Mamaev & Pak 1989: 33. Turkmenistan: Tashauz Prov., Kankakyr; HT; ♂; HYNM. 

ludmilae. Turkmenistan. Salsola chiwensis (Chenopodiaceae). 

Halodiplosis ludmilae Fedotova 1994d: 46. Turkmenistan: southern Ustyurt, Burchliburun escarpment, 300 km W 

Tashauza; HT; ♀; ZMAS. 

magna. Turkmenistan, Uzbekistan. Haloxylon aphyllum (Chenopodiaceae). 

Haloxylophaga magna Mamaev 1972a: 894. Turkmenistan: Mary, Akibay; HT; ♂; ZMAS. 

meridiana. Kazakhstan. Haloxylon aphyllum (Chenopodiaceae). 

Asiodiplosis meridianus Marikovskij 1955b: 104, nomen nudum. 

Asiodiplosis meridianus Marikovskij 1956: 193. Kazakhstan: vic. Kyzl-Orda; ST; ♂,♀; ZMAS. 

morbosa. Turkmenistan, Kazakhstan. Anabasis sp. (Chenopodiaceae). 

Halodiplosis morbosa Mamaev & Pak 1989: 33. Turkmenistan: Tashauz Prov., Shakhsenem; HT; ♂; HYNM.

241 

mutabilis. Kazakhstan. Anabasis eriopoda (Chenopodiaceae). 

Asiodiplosis mutabilis Marikovskij 1955a: 344. Kazakhstan: Boguta Mts, left bank middle course Ili R.; ST; ♂,♀; 

ZMAS. 

nanophytonigemmae. Kazakhstan. Nanophyton erinaceum; Arthrophytum balchaschense, A. longibracteatum 

(Chenopodiaceae). 

Halodiplosis nanophytonigemmae Fedotova 1990a: 229. Kazakhstan: upper reaches Charyn R., Sartogay landmark, 8 

km NW Chundza; HT; ♂; ZMAS. 

nanophytonis. Kazakhstan. Nanophyton erinaceum (Chenopodiaceae). 

Halodiplosis nanophytonis Fedotova 1990a: 228. Kazakhstan: Dzhungarskiï Alatau, 70 km SE Uch-Aral; HT; ♂; 

ZMAS. 

noxia. Kazakhstan, Turkmenistan. Haloxylon aphyllum, H. ammodendron (Chenopodiaceae). 

Asidiplosis noxia Marikovskij 1953a: 138, nomen nudum. 

Asidiplosis noxia Marikovskij 1955b: 104, nomen nudum. 

Asidiplosis noxia Marikovskij 1955c: 127, nomen nudum. 

Asiodiplosis noxia Marikovskij 1955d: 304. Kazakhstan: Ili and Syr-Darya Rivers; ST; ♂,♀; ?. 

orientalis. Kazakhstan. Salsola orientalis (Chenopodiaceae). 

Halodiplosis orientalis Fedotova 1994f: 34. Kazakhstan: lower course Ili R., 20 km NW Bakanas; HT; M ; ZMAS. 

oxyglumis. Kazakhstan. Haloxylon aphyllum (Chenopodiaceae). 

Halodiplosis oxyglumis Fedotova 1993e: 85. Kazakhstan, middle course Ili R., left bank, opposite "Singing Mtn"; 


palpata. Kazakhstan. Anabasis aphylla (Chenopodiaceae). 

Asiodiplosis palpata Marikovskij 1955a: 340. Kazakhstan: Yanykurgan to Dzhusaly, Syr-Darya R., Boguty Mts, left 

bank middle course Ili R.; ST; ♂,♀; ZMAS. 

panderiae. Kazakhstan. Panderia turkestanica (Chenopodiaceae). 

Halodiplosis panderiae Fedotova 1992f: 49. Kazakhstan: lower course Ili R., 75 km N Bakanas; HT; ♂; ZMAS. 

petrosimoniae. Kazakhstan. Petrosimonia monandra (Chenopodiaceae). 

Halocnemomyia petrosimoniae Fedotova 1993g: 20. Kazakhstan: Saur Mtn Range, 18 km NW Zaïsan, 6 km NW 

Karabulak; HT; ♂; ZMAS. 

petrosimoniana. Kazakhstan. Petrosimonia triandra; Petrosimonia spp. (Chenopodiaceae). 

Halodiplosis petrosimoniae Fedotova 1994f: 33, new secondary homonym of Halocnemomyia petrosimoniae 

Fedotova. Kazakhstan: eastern shore L. Balkhash, nr Pybach'e; HT; ♂; ZMAS. 

Halodiplosis petrosimoniana Gagné 2004b: 161, new name for Halodiplosis petrosimoniae Fedotova. 

primoveris. Kazakhstan. Salsola laricifolia (Chenopodiaceae). 

Asidiplosis primoveris Marikovskij 1953a: 136. Kazakhstan: Chulak Mts, middle course Ili R.; ST; ♂,♀; ZKAS & 

ZMAS. 

propria. Kazakhstan, Uzbekistan. Salsola rigida (Chenopodiaceae). 

Asiodiplosis propria Marikovskij 1955a: 336. Kazakhstan: Karatau Mtn Range; ST; ♂,♀; ZMAS. 

repetekensis. Turkmenistan. Salsola richteri (Chenopodiaceae). 

Halodiplosis repetekensis Kaplin 1990: 15. Turkmenistan: Chardzhou Prov., Repetek; HT; ♂; ZMAS. 

rhaphidophytonis. Kazakhstan. Rhaphidophyton regelii (Chenopodiaceae). 

Halodiplosis rhaphidophytonis (also as rhaphidophitonis) Fedotova 1989d: 27. Kazakhstan: Karatau Mtn Range, 

upper Koktal R., 27 km NW Karatau; HT; ♂; ZKAS. 

rosaria. Mongolian People's Republic. Haloxylon sp. (Chenopodiaceae). 

Asiodiplosis rosaria Mamaev 1972a: 895. Mongolian People's Republic, South Gobi Admin. Distr.; HT; ♂; ZMAS. 

salsae. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Asiodiplosis salsae Marikovskij 1955a: 345. Kazakhstan: Dzhambul Region, Karatau Mtn Range; ST; ♂; ZMAS. 

Asiodiplosis salsa Marikovskij 1955b: 104, missp. of salsae. 

salsola. Kazakhstan, Uzbekistan. Anabasis salsa (Chenopodiaceae). 

Asiodiplosis salsola Marikovskij 1955a: 337. Kazakhstan: Dzhambul Region, Karatau Mtn Range; ST; ♂; ZMAS. 

salsolae. Tunisia. Salsola tetragona (Chenopodiaceae). 

Halodiplosis salsolae Kieffer 1912e: 78. Tunisia: Gabès; ST; ♂,P; lost?. 

salsolicola. Turkmenistan, Uzbekistan. Salsola richteri (Chenopodiaceae).

242 

Haloxylonophaga salsoliicola Marikovskij 1955b: 105, nomen nudum. 

Haloxylophaga salsolicola Marikovskij 1956: 194. Turkmenistan (southeastern); ST; ♂,♀; ZMAS. 

salsuginea. Uzbekistan. Nanophyton erinaceum (Chenopodiaceae). 

Asiodiplosis salsuginea Mamaev 1972a: 896. Uzbekistan: Bukhara Prov., Kul'dzhuktau; HT; ♂; ZMAS. 

sarcobati. USA (Idaho, Utah). Sarcobatus vermiculatus (Chenopodiaceae). 

Onodiplosis sarcobati Felt 1916e: 176. USA: Utah; ST; ♀,P; NYSM. 

savojskella. Kirghizstan. Suaeda physophora (Chenopodiaceae). 

Asiodiplosis savojskella Marikovskij 1955e: 25. Kirghizstan: Kapkak valley, western end Issyk-Kul' L.; ST; ♂,♀; ?. 

saxauli. Turkmenistan. Haloxylon aphyllum; H. persicum (Chenopodiaceae). 

Halodiplosis saxauli (also as caxauli) Kaplin 1991: 63. Turkmenistan: Chardzhou Prov., Repetek; HT; ♂; ZMAS. 

schnitnikovi. Kazakhstan. Halocnemum strobilaceum (Chenopodiaceae). 

Halocnemomyia schnitnikovi Marikovskij 1965: 80. Kazakhstan: upper reaches Ili R., nr Karatau Mtn Range; ST; 

♂,♀; MARC. 

slyvkini. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis slyvkini Fedotova 1992c: 71. Kazakhstan: 18 km NW Zaysan L., 6 km NW Karabulak; HT; ♂; ZMAS. 

sperandus. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis sperandus Fedotova 1989f: 822. Kazakhstan: left bank middle course Ili R., 20 km NE of Chilik, nr 

Barandysu; HT; ♂; ZKAS. 

sphaerobia. Kazakhstan, Turkmenistan. Salsola arbuscula (Chenopodiaceae). 

Asiodiplosis sphaerobia Marikovskij 1955a: 341. Kazakhstan: Kzyl-Orda Prov., Telekul' L., and middle course Ili R.; 

ST; ♂,♀; ZMAS. 

Asiodiplosis spherobia Marikovskij 1955b: 105, missp. of sphaerobia. 

stackelbergi. Kazakhstan, Turkmenistan. Haloxylon aphyllum, H. ammodendron (Chenopodiaceae). 

Asiodiplosis stackelbergi Marikovskij 1955b: 105, nomen nudum. 

Asiodiplosis stackelbergi Marikovskij 1955d: 306. Kazakhstan: middle course Ili R.; ST; ♂,♀; ?. 

steinbergi. Kazakhstan. Salsola sp. (Chenopodiaceae). 

Tyloceramyia steinbergi Marikovskij 1955b: 105, nomen nudum. 

Tyloceramyia steinbergi Marikovskij 1956: 192. Kazakhstan: Guryev Prov., Saraichik; ST; ♂,♀; ZMAS. 

strobiliformis. Kazakhstan. Salsola richteri (Chenopodiaceae). 

Halodiplosis strobiliformis Fedotova 1991g: 35. Kazakhstan: northeastern Kyzylkum Desert, 30 km S Yanykurgana, 

24 km SW Kyrkense; HT; ♂; ?. 

sympegmae. Kazakhstan. Sympegma regelii (Chenopodiaceae). 

Halodiplosis sympegmae Fedotova 1990e: 45. Kazakhstan: 19 km SW Chundzha, 16 km N Aksaï; HT; ♂; ZMAS. 

syrdarjensis. Kazakhstan, Turkmenistan. Salsola arbuscula (Chenopodiaceae). 

Asiodiplosis syrdarjensis Marikovskij 1955a: 342. Kazakhstan: Kzyl-Orda Prov., Telekul' L., and Dzhusaly in 

western Karatau Mtn Range; ST; ♂,♀; ZMAS. 

truncata. Kazakhstan. Anabasis truncata (Chenopodiaceae). 

Halodiplosis truncata Fedotova 1992c: 75. Kazakhstan: eastern L. Zaysan Reg., 6 km E Ardynka, 8 km NE Buran; 


truncatula. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Haloxylophaga truncatula Mamaev 1972a: 895. Turkmenistan: central Karakumy Desert, Dzhar; HT; ♂; ZMAS. 

tsherkesi. Turkmenistan. Salsola richteri (Chenopodiaceae). 

Halodiplosis tsherkesi Kaplin 1990: 17. Turkmenistan: Chardzhou Prov., Repetek; HT; ♂; ZMAS. 

ulkulkalkani. Kazakhstan, Turkmenistan, Uzbekistan. Haloxylon aphyllum, H. ammodendron (Chenopodiaceae). 

Asiodiplosis ulkunkalkani Marikovskij 1955b: 105, nomen nudum. 

Asiodiplosis ulkulkalkani Marikovskij 1955d: 305. Kazakhstan: Ulkulkalkan Mtn, right bank, middle course Ili R.; 

ST; ♂,♀; ?. 

unica. Kazakhstan. Arthrophytum iliense (Chenopodiaceae). 

Halodiplosis unicus Fedotova 1992e: 46.: upper reaches Charyn R., Sartogay landmark, 8 km NW Chundza; HT; ♂; 

ZMAS. 

*urceolatus. Kazakhstan. Kochia prostrata (Chenopodiaceae). 

Halodiplosis urceolatus Fedotova 1988b: 41. Kazakhstan: 55 km N Alma-Ata; HT; ♂; ZKAS.

243 

uzenensis. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Halodiplosis uzenensis Fedotova 1993c: 112. Kazakhstan: eastern Mangyshlak basin, 150 km NE Senek, Kyzyloy; 


vernalis. Kazakhstan, Turkmenistan. Haloxylon ammodendron (Chenopodiaceae). 

Asiodiplosis vernalis Marikovskij 1955b: 105, nomen nudum. 

Asiodiplosis vernalis Marikovskij 1955d: 308. Kazakhstan: Charyn, upper course Ili R.; ST; ♂,♀; ?. 

vicina. Kazakhstan, Uzbekistan. Kochia prostrata (Chenopodiaceae). 

Asiodiplosis vicina Marikovskij 1955a: 343. Kazakhstan: Dzhambul Prov., Karatau Mtn Range; ST; ♂,♀; ZMAS. 

zajsanicus. Kazakhstan. Haloxylon ammodendron (Chenopodiaceae). 

Halodiplosis zajsanicus Fedotova 1993c: 114. Kazakhstan: northern L. Zaysan Region, 6 km N Amanat, Karabyurek 

Mtn; HT; ♂; ZMAS. 

HAPLODIPLOSIS 

Haplodiplosis Rübsaamen 1910d: 288. Type species, Diplosis equestris Wagner (mon.) = marginata Roser. 


marginata. Widespread Europe. Unknown orig. host; Triticum, Hordeum, Avena, Secale, and other genera of Poaceae 

(Barnes 1956). 

Cecidomyia marginata Roser 1840: 50. Germany: T; A; lost. 

Diplosis equestris Wagner 1871: 414. Germany: Fulda; ST; ♂,♀,P,L; lost. 

Haplodiplosis incerta, as form of equestris, Rübsaamen 1926: 140. Germany: T; ♂; ZMHB. 

HAPLOPALPUS 

Haplopalpus Rübsaamen 1916a: 471. Type species, serjaneae Rübsaamen (mon.). 

Hablopalpus Rübsaamen 1916a: 474, missp. of Haplopalpus. 


serjaneae. Brazil (Amazonas). Serjania sp. (Sapindaceae). 

Haplopalpus serjaneae Rübsaamen 1916a: 473. Brazil: Amazonas, nr Auristella on Rio Acre; ST; ♀,P; ZMHB. 

HARMANDIOLA 

Harmandia Kieffer 1896b: 5, preocc. Rochebrune 1882. Type species, Cecidomyia tremulae Winnertz (orig. des.). 

Harmandiola Skuhravá 1997: 166, new name for Harmandia Kieffer. 

Holarctic; 14 spp., on Populus except two on Castanea and one on Carya; Cecidomyiidi, unplaced to tribe. 

amisae. USA (Georgia). Castanea pumila (Fagaceae). 

Harmandia amisae Gagné 1992a: 388. USA: Georgia, Peach Co., Byron; HT; ♂; USNM. 

castaneae. USA (Massachusetts). Castanea dentata (Fagaceae). 

Cecidomyia castaneae Stebbins 1910: 17. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

cavernosa. Widespread Europe, Kazakhstan. Populus tremula (Salicaceae). 

Diplosis cavernosa Rübsaamen 1899d: 602. Germany; T; A; ZMHB. 

globuli. Widespread Europe, Kazakhstan. Populus tremula (Salicaceae). 

Diplosis globuli Rübsaamen 1889a: 49. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

helena. USA (Massachusetts). Populus tremuloides (Salicaceae). 

Dicrodiplosis helena Felt 1912l: 245. USA: Massachusetts, Magnolia; HT; ♂; NYSM. 

hudsoni. USA (New York, New Hampshire).

244 

Cecidomyia hudsoni Felt 1907a: 39. USA: New York, Poughkeepsie; HT; ♂; NYSM. 

Itonida reflexa Felt 1913f: 146. USA: NH, Hampton; HT; ♂; NYSM. 

nucicola. Eastern Nearctic. Carya ovata; C. tomentosa (Juglandaceae). Ref.: Gagné 2008c: 95 (redescr.). 

Cecidomyia caryae nucicola Osten Sacken 1870: 53, invalid, a polynomial. 

Cecidomyia nucicola Osten Sacken 1878: 6. USA: New York, nr Rhinebeck; ST; L, G; USNM. 

Cecidomyia caryaenucicola Jarvis 1909: 84. 

polymorpha. Switzerland. Populus tremula (Salicaceae). 

Cecidomyia polymorpha Bremi 1847a: 16. Switzerland: vic. Zurich; ST; LT; lost. 

populea. Germany. Populus tremula (Salicaceae). 

Tipula populea Schrank 1803: 87. Germany: Bavaria; T; G; ?. Placed here because it forms a pea-shaped gall on 

poplar leaves as do some other Harmandia spp. 

populi. UK, Sweden, Germany, Poland, Czechoslovakia, Hungary, Romania, Yugoslavia, Kazakhstan. Populus 

tremula (Salicaceae). 

Harmandia populi Rübsaamen 1917: 82. Germany; ST; ♂,♀,L; ZMHB. 

pustulans. UK, Sweden, France, Germany, Czechoslovakia, Yugoslavia. Populus tremula (Salicaceae). 

Harmandia pustulans Kieffer 1909b: 18. France; T; G; lost?. 

reginae. Canada (Saskatchewan). 

Itonida reginae Felt 1921a: 196. Canada: Saskatchewan, Regina; HT; ♂; NYSM. 

stebbinsae. USA (Massachusetts, Maryland). Populus grandidentata (Salicaceae). 

Cecidomyia irregularis Stebbins 1910: 9, jun. prim. hom. irregularis Bremi. USA: Massachusetts, vic. Springfield; T; 

G; NHSM. 

Cecidomyia stebbinsae Gagné 1972b: 324, new name for Cecidomyia irregularis Stebbins. 

tremulae. Widespread Europe, Kazakhstan. Populus tremula (Salicaceae). 

Cecidomyia tremulae Winnertz 1853: 273. Germany; ST; ♂,♀,L; MNHN?. 

Diplosis loewii Rübsaamen 1892c: 49. Germany: vic. Weidenau; ST; ♂,♀,L; ZMHB. 

HARTIGIOLA 

Hartigiola Rübsaamen 1912c: 161. Type species, Cecidomyia annulipes Hartig (orig. des.). 

Phegobia Kieffer 1913i: 61. Type species, Cecidomyia tornatella Bremi (mon.) = annulipes Hartig. 

Palearctic; 2 spp.; Lasiopteridi, Dasineurini. 

annulipes. Widespread Europe, Turkey; Pliocene/Germany (Straus 1977). Fagus sylvatica (Fagaceae). 

Cecidomyia annulipes Hartig 1839: 641. Germany; ST; ♂,♀; ?. 

Cecidomyia tornatella Bremi 1847a: 13. Switzerland: Montallein nr Chur; T; G; ETHZ. 

Cecidomyia piligera Loew 1850a: 32. Poland; ST; ♀; ZMHB?. 

Oligotrophus fagineus Kieffer 1909b: 7. Austria?; T; G; NHMW?. 

Hartigiola fossilus Evenhuis 1994: 182, error and missp. of "fossilis" Straus 1977: 50, 70, which was not a new 

scientific name but a record of a fossil gall identified as being that of either Mikiola fagi or Hartigiola 

annulipes. 

faggalli. Japan (Honshu to Kyushu). Fagus crenata (Fagaceae). 

Oligotrophus faggalli Monzen 1955a: 47. Japan: Honshu, Aomori, Iwate, Tochigi, Wakayama, & Tottori Prefs; NT 

(Sato & Yukawa 2004: 20, Japan: Iwate Pref., Morioka City, Kuriyagawa); ♂; KUEC. 

HASEGAWAIA 

Hasegawaia Monzen 1937: 192. Type species, sasacola Monzen (mon). 


sasacola. Japan (Hokkaido, Honshu). Sasa ramosa; Sasa spp. (Poaceae).

Hasegawaia sasacola Monzen 1937: 192. Japan: Honshu, Iwate, & Gifu PRefs; ST; ♂,♀,L; KUEC. 

Rhabdophaga sasae Shinji 1938a: 232. Japan: Honshu; ST; ♂,♀,L; lost. 

HELIODIPLOSIS 

Heliodiplosis Felt 1918c: 291. Type species, spatholobi Felt (orig. des.). 

Spatholobomyia Rao 1952b: 41, unnec. new name for Heliodiplosis Felt. 


spatholobi. Philippines. Spatholobus gyrocarpus (Fabaceae). 

Heliodiplosis spatholobi Felt 1918c: 291. Philippines: Luzon, Laguna Prov., Mt Makiling; HT; ♀; NYSM. 

HEMIASPHONDYLIA 

Hemiasphondylia Möhn 1960c: 229. Type species, mimosae Möhn (orig. des.). 

Neotropical; 2 spp., from bud galls of Mimosoideae (Fabaceae); Cecidomyiidi, Asphondyliini, Asphondyliina. 

enterolobii. Costa Rica. Enterolobium cyclocarpum (Fabaceae). 

Asphondylia enterolobii Gagné 1978c: 515. Costa Rica: Guanacaste Prov., Santa Rosa Natl Pk; HT; P; USNM. 

mimosae. El Salvador, Colombia. Mimosa albida; M. leiocarpa, Prosopis juliflora (Fabaceae). 

Hemiasphondylia mimosae Möhn 1960c: 230. El Salvador: La Libertad, N El Cimarrón; HT; ♂; SMNS. 

Asphondylia mimosicola Gagné 1978c: 514, new name for then sec. hom. Asphondylia mimosae Möhn. 

HERBOMYIA 

Herbomyia Möhn 1955b: 147. Type species, robusta Möhn (orig. des.). 


robusta. Germany. Carex spp. (Cyperaceae). 

Herbomyia robusta Möhn 1955b: 147. Germany: Dauborn; HT; ♂; SMNS. 

HETERASPHONDYLIA 

Heterasphondylia Möhn 1960c: 227. Type species, salvadorensis Möhn (orig. des.). 

Neotropical; 1 sp.; Cecidomyiidi, Asphondyliini, Asphondyliina. 

salvadorensis. El Salvador. Piper sp. (Piperaceae). 

Heterasphondylia salvadorensis Möhn 1960c: 228. El Salvador: San Salvador; HT; ♂; SMNS. 

HETEROBREMIA 

Heterobremia Felt 1920c: 494. Type species, furcata Felt (orig. des.). 

Afrotropical; 1 sp.; Cecidomyiidi, Aphidoletini. 

furcata. Cameroon. 

Heterobremia furcata Felt 1920c: 494. Cameroon: Lonji, nr Ulou R.; HT; ♂; NYSM. 

245

HETEROCONTARINIA 

Heterocontarinia Hardy 1960: 271. Type species, spinosa Hardy (orig. des.). 

Oceanian; 1 sp.; Cecidomyiidi, Lestodiplosini. 

spinosa. Hawaiian Is. 

Heterocontarinia spinosa Hardy 1960: 272. Hawaiian Is: Oahu, Palolo; HT; ♂; HDOA. 

*HETEROSTYLIDIPLOSIS 

Heterostylidiplosis Fedotova 2004f: 104. Type species, serpentina (orig. des.). 

Eastern Palearctic; 4 spp.; Cecidomyiidi, Karshomyiini. This genus, for which Fedotova (2004f) erected the tribe 

Heterostylidiplosini, appears to be close to Karshomyia. 

*capituliformis. Russia (Far East). 

Heterostylidiplosis capituliformis Fedotova 2004i: 610. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


fungicola. Russia (Far East). 

Karschomyia fungicola Mamaev & Krivosheina 1998: 78. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMMU. 

hemisphaerica. Russia (Far East), Canada (Ontario). 

Karshomyia hemisphaerica Kovalev & Mamaev 1966: 231. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

HYNM. 

Karschomyia insolita Gagné 1973d: 351. Canada: Ontario, N. Burgess Township, Black L.; HT; ♂; USNM. 

*serpentina. Russia (Far East). 

Heterostylidiplosis serpentinus Fedotova 2004f: 105. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


HOLOBREMIA 

Holobremia Kieffer 1912h: 1. Type species, Bremia fallacicornis Kieffer (orig. des.). 

Holobremyia Shinji 1944: 232, missp. of Holobremia. 


cylindrica. India (Jammu and Kashmir). 

Holobremia cylindricus Kashyap 1987: 108. India: Jammu and Kashmir, Srinagar; HT; ♂; PGCA. 

fallacicornis. France, Russia (Europe). 

Holobremia fallacicornis Kieffer 1904c: 391. France: vic. Bitche; T; ♂; lost?. 

lignicola. France, Russia (Europe). 

Holobremia lignicola Kieffer 1913f: 45. France: Bitche; T; ♂; lost?. 

HOLODIPLOSIS 

Holodiplosis Kieffer 1913h: 103. Type species, alticola Kieffer (orig. des.). 


alticola. Kenya. 

Holodiplosis alticola Kieffer 1913h: 104. Kenya: along Burgerett R. betw. Narémuru and West Kenya For. Stn; HT; 

246

♂; MNHN. 

imicola. Kenya. 

Holodiplosis imicola Kieffer 1913h: 104. Kenya: Tiwi, S Mombasa; HT; ♂; MNHN. 

HOMOBREMIA 

Homobremia Kieffer 1912h: 1. Type species, Bremia emarginata Kieffer (orig. des.). 

Palearctic; 1 sp.; Cecidomyiidi, Aphidoletini. 

emarginata. France. 

Bremia emarginata Kieffer 1904c: 399. France: vic. Bitche; ST; ♂,♀; lost?. France. 

HORIDIPLOSIS 

Horidiplosis Felt 1920b: 10. Type species, fici Felt (orig. des.). 

Hordiplosis Shinji 1944: 252, missp. of Horidiplosis. 

Oriental; 7 spp., all on Ficus; Cecidomyiidi, unplaced to tribe. 

caricae. India (Uttar Pradesh). Ficus carica (Moraceae). 

Xylodiplosis fici Grover 1967c: 134, jun. sec. hom. fici Felt. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

Horidiplosis caricae Gagné 2004b: 165, new name for Xylodiplosis fici Grover. 

fici. India (Bihar). Ficus infectoria (Moraceae). 

Horidiplosis fici Felt 1920b: 10. India: Bihar, Pusa; ST; ♂,♀; NYSM. 

ficicola. India (Uttar Pradesh). Ficus glomerata (Moraceae). 

Dyodiplosis fici Rao 1951: 123, jun. sec. hom. fici Felt. India: Uttar Pradesh, Agra, Hewitt Park; HT; ♂; ZSCI. 

Horidiplosis ficicola Gagné 1973a: 502, new name for Dyodiplosis fici Rao. 

*ficifolii. China, Taiwan; immigr: Netherlands, Denmark, Italy, USA (Florida). Ficus benjamina; F. microcarpa 

(Moraceae). 

Horidiplosis ficifolii Harris 2003a: 304. Taiwan (through Honselersdijk, Netherlands): HT; ♂; ZMAN. Ref.: Steck, 

G.J. and S. Krueger 1908 (biol., distr.). 

glomerataii. India (Uttar Pradesh). Ficus glomerata (Moraceae). 

Dyodiplosis glomerataii Grover 1962b: 276. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

mathuri. India (Uttar Pradesh). Ficus glomerata (Moraceae). 

Horidiplosis mathuri Mani 1935: 440. India: Uttar Pradesh, Dehra Dun, New Forest; HT; ♀; ZSCI. 

orientalis. India (Uttar Pradesh). Ficus religiosa (Moraceae). 

Isodiplosis orientalis Grover 1967b: 48. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

HOUARDIELLA 

Houardiella Kieffer 1912e: 69. Type species, salicorniae Kieffer (orig. des. as n. g., n. sp.). 

Kalidiomyia Fedotova 1984a: 1. Type species, distincta Fedotova (orig. des.). 

Palearctic; 3 spp., on Chenopodiaceae; Cecidomyiidi, Asphondyliini, Asphondyliina. Ref.: Gagné 2004b (syn. note). 

*gracilis. Israel. Arthrocnemum macrostachyum (Chenopodiaceae). 

Houardiella gracilis Dorchin & Freidberg 2008: 13. Israel: Akko; HT; ♂; TAUI. 

distincta. Kazakhstan. Kalidium schrenkianum (Chenopodiaceae). 

Kalidiomyia distincta Fedotova 1984a: 3. Kazakhstan: left bank upper course Ili R., 20 km NW Barandysu; HT; ♂; 

ZKAS. 

salicorniae. Italy, Tunisia, Libya. Salicicornia fruticosa (Chenopodiaceae). 

247

Houardiella salicorniae Kieffer 1912e: 69. Tunisia: Tunis, Oasis de Nefta; ST; ♂,P; lost?. 

HOUARDODIPLOSIS 

Houardodiplosis Tavares 1925a: 13. Type species, rochae Tavares (orig. des.). 

Neotropical; 1 sp.; Cecidomyiidi, Clinodiplosini. Ref.: Maia 2002a (larva). 

rochae. Brazil (Ceará). Combretum leprosum (Combretaceae). 

Houardodiplosis rochae Tavares 1925a: 14. Brazil: Ceará, vic. Fortaleza; ST; ♂,♀,P; TAVC. 

HURADIPLOSIS 

Huradiplosis Nijveldt 1968: 61. Type species, surinamensis Nijveldt (orig. des.). 


surinamensis. Surinam. Hura crepitans (Euphorbiaceae). 

Huradiplosis surinamensis Nijveldt 1968: 62. Surinam: Paramaribo; HT; ♂; ZMAN. 

HYBOLASIOPTERA 

Hybolasioptera Rübsaamen 1916b: 560. Type species, Cecidomyia cerealis of Lindeman not Fitch (mon.) = fasciata 

Kieffer. 

Holarctic; 2 spp., on Poaceae; Lasiopteridi, Lasiopterini. 

elymi. USA (Kansas). Elymus virginicus (Poaceae). 

Hybolasioptera elymi Gagné 1969d: 1352. USA: Kansas, Geary Co.; HT; ♂; USNM. 

fasciata. Widespread Europe. Type host unknown; Secale cereale & spp. of Agropyron, Agrostis, Avenella, 

Calamagrostis, Holcus, Triticum (Poaceae). 

Lasioptera fasciata Kieffer 1904b: 342, as var. of cerealis of Lindeman (misident.) not Fitch. France: vic. Bitche; ST; 

♂,♀; lost?. 

HYGRODIPLOSIS 

Hygrodiplosis Kieffer 1912h: 1. Type species, Diplosis vaccinii Kieffer (orig. des.). 


vaccinii. UK, Sweden, Finland, Netherlands, France, Czechoslovakia, Italy, Austria, Romania, Italy. Vaccinium 

uliginosum (Ericaceae). 

Diplosis vaccinii (as caccinii) Kieffer 1897c: 14. France: Bitche; T; ♀; lost?. 

HYPERDIPLOSIS 

Hyperdiplosis Felt 1908b: 405. Type species, Cecidomyia lobata Felt (orig. des.). 

Holarctic, Oriental; 11 spp.; Cecidomyiidi, Clinodiplosini. This genus appears to be an artificial collection of species. 


Cecidomyia americana Felt 1908b: 413. USA: New York, Poughkeepsie; HT; ♂; NYSM. 

248

249 

banksi. Philippines. Cissus adnata (Vitaceae). 

Hyperdiplosis banksi Felt 1918c: 294. Philippines: Luzon, Laguna Prov., Los Baños Falls, nr Los Baños; HT; ♀; 

NYSM. 

bharatensis. India (Uttar Pradesh). 

Hyperdiplosis bharatensis Grover & Bakhshi 1978: 139. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

bryanti. USA (Massachusetts, Connecticut), UK, Germany. 

Hyperdiplosis bryanti Felt 1913f: 146. USA: Massachusetts, North Adams; HT; ♂; NYSM. 

emarginata. India (Tripura). 

Hyperdiplosis emarginata Chandra 1993f: 190. India: Tripura, Agartala; HT; ♂; CSIA. 

lobata. USA (New York), UK, Germany. 

Cecidomyia lobata Felt 1907a: 39. USA: New York, Karner; HT; ♂; NYSM. 

oryzae. India (Andhra Pradesh). 

Hyperdiplosis oryzae Grover & Bakhshi 1978: 135. India: Andhra Pradesh, Hyderabad; HT; ♂; PGCA. 

prayagensis. India (Uttar Pradesh). 

Clinodiplosis prayagensis Chandra 1992: 100. India: Uttar Pradesh, Allahabad, Prayag; HT; ♂; CSIA. 

recurvata. USA (New York, District of Columbia). 

Cecidomyia recurvata Felt 1907a: 38. USA: New York, Poughkeepsie; HT; ♂; NYSM. 

Hyperdiplosis fungicola Felt 1911l: 461, nomen nudum. 

Hyperdiplosis fungicola Felt 1911n: 552. USA: District of Columbia; HT; ♂; USNM. 

relicta. Philippines. 

Hyperdiplosis relicta Felt 1918c: 295. Philippines: Luzon, Laguna Prov., Los Baños; HT; ♀; NYSM. 

tritici. USA. Triticum aestivum (Poaceae). 

Itonida tritici Felt 1912e: 289. USA; ST; ♂,♀,L; USNM. 

Cecidomyia triticicola Kieffer 1913i: 220, unnec. new name for Itonida tritici Felt (Cecidomyia tritici Kirby in 

Contarinia when new name proposed). 

Cecidomyia triticoides Barnes 1956: 28, unnec. new name for Itonida tritici Felt. 

HYPODIPLOSIS 

Hypodiplosis Kieffer 1913i: 187. Type species, Clinodiplosis paederiae Kieffer and J. & W. Docters van 

Leeuwen-Reujnvaan (mon.). 


paederiae. Indonesia (Java). Paederia foetida (Rubiaceae). 

Clinodiplosis paederiae Kieffer & Docters van Leeuwen-Reijnvaan 1910: 129. Indonesia: Java, Salatiga; ST; ♂,♀,P; 

lost. 

*HYPOPRODIPLOSIS 

Hypoprodiplosis Fedotova 2004e: 18. Type species, universalis Fedotova (orig. des.). 


*universalis. Russia (Far East). Vicia cracca (Fabaceae). 

Hypoprodiplosis universalis Fedotova 2004e: 18. Russia: Primorskiï Terr., 23 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

IATROPHOBIA 

Iatrophobia Rübsaamen 1916a: 469. Type species, Clinodiplosis brasiliensis Rübsaamen (orig. des.). 

Latrophobia Shinji 1944: 244, missp. of Iatrophobia.

Iatrodiplosis Mani 1946: 222, missp. of Iatrophobia. 

Jatrophobia Barnes 1946b: 32, unjust. emend. of Iatrophobia. 


brasiliensis. Costa Rica, Guadeloupe, St. Vincent, Trinidad, Tobago, Guyana, Surinam, Brazil (Amazonas, Rio de 

Janeiro, Santa Catarina). Manihot utilissima (Euphorbiaceae). 

Clinodiplosis brasiliensis Rübsaamen 1908a: 156. Brazil: Amazonas, Fortaleza & Jurua Miri, Rio de Janeiro, Maua & 

Palmerias, & Santa Catarina, Tubarão; ST; P,L; ZMHB. 

Cecidomyia manihot Felt 1910c: 268. St. Vincent; ST; ♂,♀; USNM. 

*ILLICIOMYIA 

Illiciomyia Tokuda 2004: 4. Type species, yukawai Tokuda (orig. des.). 

Palearctic; 1 sp.; Cecidomyiidi, Asphondyliini, Asphondyliina. 

*yukawai. Japan (Honshu to Kyushu & islands). Illicium anisatum (Illiciaceae). 

Illiciomyia yukawai Tokuda 2004: 1-11. Japan: Kyushu, Fukuoka, Sasaguri, Mt. Wakasugi; HT; ♂; SMNS. 

*INCLINATIDIPLOSIS 

Inclinatidiplosis Fedotova 2004h: 173. Type species, coronalis Fedotova (orig. des.). 

Eastern Palearctic; 2 spp.; Cecidomyiidi, Clinodiplosini. This genus should be compared with Bicornidiplosis. 

Fedotova and Sidorenko (2009) base a new tribe on this genus (see Appendix). 

*coronalis. Russia (Far East). 

Inclinatidiplosis coronalis Fedotova 2004h: 173. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*crystallina. Russia (Far East). 

Inclinatidiplosis crystallina Fedotova & Sidorenko 2009d: 227. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 


INCOLASIOPTERA 

Incolasioptera Möhn 1975: 64. Type species, siccida Möhn (orig. des.). 


siccida. El Salvador. Celtis iguanea (Ulmaceae). 

Incolasioptera siccida Möhn 1975: 65. El Salvador: nr La Libertad; HT; LT; SMNS. 

INULOMYIA 

Inulomyia Stelter 1965: 91. Type species, Cecidomyia subterranea Frauenfeld (orig. des.). 


subterranea. Germany, Austria, Hungary, Italy. Inula ensifolia (Asteraceae). 

Cecidomyia subterranea Frauenfeld 1861: 172. Austria: L (Stelter 1965: 96); ♀; NHMW. 

250

ISOGYNANDROMYIA 

Isogynandromyia Spungis 1981: 43. Type species, terricola Spungis (orig. des.). 

Palearctic; 1 sp.; Lasiopteridi, Ledomyiini. 

terricola. Latvia. 

Isogynandromyia terricola Spungis 1981: 44. Latvia: Brotseny; HT; ♂; LASS. 

ISOLASIOPTERA 

Isolasioptera Möhn 1964b: 584. Type species, eupatoriensis Möhn (orig. des.). 

Neotropical; 2 spp., on Eupatorium; Lasiopteridi, Lasiopterini. 

eupatoriensis. El Salvador. Eupatorium morifolium; Eupatorium sp. (Asteraceae). 

Isolasioptera eupatoriensis Möhn 1964b: 585. El Salvador: Sonsonate, on rd betw. Armenia & Izalco; HT; ♂; 

SMNS. 

palmae. El Salvador. Eupatorium collinum (Asteraceae). 

Isolasioptera palmae Möhn 1975: 78. El Salvador: Santa Ana, nr San José, Hacienda San José; HT; ♂; SMNS. 

*ISTODIOMYIA 

Istodiomyia Fedotova 2008: 119. Type species, alexandrae Fedotova (orig. des.). 


*alexandrae. Russia. Polygala hybrida (Polygalaceae). 

Istodiomyia alexandrae Fedotova 2008: 120. Russia: Central Povolzhye, Zhigulevsk Nature Res., nr Zol'noy; HT; ♂; 

ZMAS. 

ITEOMYIA 

Iteomyia Kieffer 1913g: 55. Type species, Cecidomyia capreae Winnertz (orig. des.). 

Holarctic; 5 spp., from Salix leaf galls; Lasiopteridi, unplaced to tribe. 

capreae. Widespread Europe, Turkey, China, Japan (Hokkaido, Honshu). Salix caprea; S. aurita, S. oleifolia 

(Salicaceae). 

Cecidomyia capreae Winnertz 1853: 291. Germany; ST; ♂,♀,L; lost. 

Cecidomyia salicisfolii (as salicis-folii) Hardy 1854c: 80. UK: Scotland, vic. Penmanshiel; T; LT; lost. 

major. UK, France. Salix caprea (Salicaceae). 

Oligotrophus major, as var. of capreae, Kieffer 1898f: 22. France: Moselle; T; A; lost?. Synonymized under capreae 

in Nijveldt & Yukawa 1982: 33, this species is resurrected in Redfern & Askew 1992: 73. 

peyerimhofi. Algeria. Salix pedicellata (Salicaceae). 

Oligotrophus peyerimhofi Kieffer 1909b: 27. Algeria: Kabylia; T; LT; lost?. 

Iteomyia peyerimhoffi Houard 1912: 19, missp. of peyerimhofi. 

salicifolia. USA (New York). Salix sp. (Salicaceae). 

Oligotrophus salicifolius Felt 1910d: 354. USA: New York, Karner; LT (Gagné 1967a: 131); ♂; NYSM. 

salicisverruca. USA (Washington, Oregon, Illinois, New York, Massachusetts). Salix humilis (Salicaceae). 

Cecidomyia salicis verruca Walsh 1864b: 606, invalid, a polynomial. 

Cecidomyia salicisverruca (as salicis-verruca) Osten Sacken 1878: 7. USA: Illinois, nr Rock Island; ST; LT; USNM. 

251

Cecidomyia verruca Felt 1906a: 745, error for salicisverruca Osten Sacken. 

IZENIOLA 

Izeniola Fedotova 1985a: 46. Type species, Stefaniola potanini Fedotova (orig. des.). 

Palearctic; 3 spp., all on Chenopodiaceae; Lasiopteridi, Lasiopterini. Ref.: Dorchin 2001 (descr.). 

bassiae. Turkmenistan. Bassia hyssopifolia (Chenopodiaceae). 

Izeniola bassiae Fedotova 1993h: 48. Turkmenistan: Kugitang Mtn Range, 25 km SW Kugitang, 55 km NE 

Charshanga; HT; ♀; ZMAS. 

obesula. Israel, Eritrea. Suaeda monoica (Chenopodiaceae). 

Izeniola obesula Dorchin 2001: 571. Israel: Newe Zohar; HT; ♀; TAUI. 

potanini. Kazakhstan. Kochia prostrata (Chenopodiaceae). 

Stefaniola potanini Fedotova 1982b: 43. Kazakhstan: Kerbulak, right bank Ili R.; HT; ♀; ?. 

Stefaniola deserta Fedotova 1982b: 44. Kazakhstan: Kerbulak, right bank Ili R.; HT; ♀; ?. 

JAAPIELLA 

Jaapiella Rübsaamen 1916b: 490. Type species, catariae Rübsaamen (orig. des.). 

Palearctic; 79 spp.; Lasiopteridi, Dasineurini. No distinction between this genus and Dasineura has been made for the 

North American fauna. The difference between the two genera cited for European species is a posteromedial 

protuberance on the gonocoxite, but this is not present in all species presently assigned to this genus (Sylvén & Tastás- 

Duque 1993) and occurs on some species assigned to Dasineura. Compare also Neomikiella & Wachtliella. 

* † acinaciformis. Late Eocene - Ukrainian amber. 

Jaapiella acinaciformis Fedotova 2005e: 49. Ukraine amber: Rovno; HT; ♀; UASK. 

acroptiloniflorae. Kazakhstan. Acroptilon repens (Asteraceae). 

Jaapiella acroptiloniflorae Fedotova 1987a: 36. Kazakhstan: Talasskiï Alatau, Daubaba, 14 km S Kal'ty-Mashat; HT; 

♂; ZMAS. 

adpressae. Kazakhstan. Centaurea adpressa (Asteraceae). 

Jaapiella adpressae Fedotova 1995e: 20. Kazakhstan: Kirghiz Mtn Range, Merke gorge, 20 km S Merke; HT; ♂; 

ZMAS. 

alaris. Kazakhstan. Sonchus arvensis (Asteraceae). 

Jaapiella alaris Fedotova 1985f: 38. Kazakhstan: left bank middle course Ili R., 20 km NW Barandysu; HT; ♂; 

ZKAS. 

alpina. UK, Sweden, Poland, Czechoslovakia, Switzerland, Austria, Romania, Kazakhstan. Silene acaulus; Silene spp. 

(Caryophyllaceae). 

Cecidomyia alpina Löw 1885a: 110. Austria: several localities in Lower Austria, & UK: Scotland, nr Glasgow; ST; 

♂,♀,L; NHMW. 

antennariae. Kazakhstan. Antennaria dioica (Asteraceae). 

Jaapiella antennariae Fedotova 1993i: 46. Kazakhstan: Sarymsakty Mtn Range, 45 km SW Katon-Karagaya, nr 

Ornek; HT; ♂; ZMAS. 

azutavica. Kazakhstan. Rumex acetosa (Polygonaceae). 

Jaapiella azutavica Fedotova 1997f: 450. Kazakhstan: southern Altaï, Markakol'skii Res., L. Markakol', nr 


bajankolica. Kazakhstan. Galium septentrionale (Rubiaceae). 

Jaapiella bajankolica Fedotova 1997b: 46. Kazakhstan: Terskey Alatau, upper Bayankol R., 28 km SW Narynkol; 


blavatskae. Kazakhstan. Spiraea media (Rosaceae). 

Jaapiella blavatskae Fedotova 1996f: 54. Kazakhstan: southern Altaï, nr L. Markakol', nr Urunkhaïka, 60 km N 

252

253 


bryoniae. Widespread Europe, Algeria. Bryonia alba (Cucurbitaceae). 

Cecidomyia bryoniae Bouché 1847: 144. Germany; ST; ♂,♀,P,L; ZMHB?. 

buhri. Germany. Peucedanum oreoselinum (Apiaceae). 

Jaapiella buhri Stelter 1975b: 240. Germany: Bocksberg Nature Res., Kösterbeck nr Rostock; HT; ♂; SMTD. 

calophaciflorae. Kazakhstan. Calophaca soongorica (Fabaceae). 

Jaapiella calophaciflorae Fedotova 1989e: 36. Kazakhstan: Tarbagataï Mtn Range, 25 km S Ayaguz; HT; ♂; ZKAS. 

catariae. Germany. Nepeta cataria (Lamiaceae). 

Jaapiella catariae Rübsaamen 1916b: 502. Germany: Prignitz, Triglitz; L (Sylvén 1993: 291); ♂; ZMHB. 

*chelidonii. Russia. Chelidonium majus (Papaveraceae). 

Jaapiella chelidonii Fedotova 2008: 125. Russia: Central Povolzhye, Zhigulevsk Nature Res., nr Bakhilova Polyana; 


*chondrillae. Turkey. Chondrilla juncea (Asteraceae). 

Jaapiella chondrillae Skurravá 2004: 236. Turkey: nr Selcuk, ca 100 mi S Izmir; HT; ♂; SKCR. 

cirsiicola. Widespread Europe, Kazakhstan. Cirsium acaule; Cirsium spp. (Asteraceae). 

Jaapiella cirsiicola Rübsaamen 1916b: 503. Germany: Prignitz, Triglitz; L (Sylvén 1998b: 257); ♂; ZMHB. 

clethrophila. Widespread Europe. Alnus glutinosa (Betulaceae). 

Jaapiella clethrophila Rübsaamen 1917: 41. Germany: Ilmenau; ST; ♂,♀,L; ZMHB. 

compositarum. UK, Netherlands, France, Germany, Romania. Hypochoeris glabra, Hieracium pilosella, Hieracium 

murorum (Asteraceae). 

Cecidomyia compositarum Kieffer 1888d: 310. France: Bitche; ST; ♂,♀,L; lost?. 

cousiniae. Kazakhstan. Cousinia bonvalotii (Asteraceae). 

Jaapiella cousiniae Fedotova 1998a: 35. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Ul'ken-Kaïndy pass, 20 

km SE Novonikolaevka; HT; ♀; ZMAS. 

crinita. UK, Germany. Senecio vulgaris (Asteraceae). 

Cecidomyia crinita Rübsaamen 1891c: 45. Germany: vic. Siegen; ST; ♂,L; ZMHB. 

cucubali. Widespread Europe. Cucubalus baccifer (Caryophyllaceae). 

Perrisia cucubali Kieffer 1909b: 6. France, Italy; T; G; lost?. 

Perrisia cucubalina Tavares 1919b: 55. Spain: Tuy; ST; ♂,♀,L; TAVC. 

Jaapiella cucubali Rübsaamen 1921: 49. Poland: Scinawa; ST; ♂,L; ZMHB. 

dittrichi. UK, France, Germany, Poland, Czechoslovakia. Silaus pratensis (Apiaceae). 

Dichelomyia dittrichi Rübsaamen 1895b: 178. Poland: Wroclaw; ST; ♂,♀,L; ZMHB. 

dracocephalicola. Kazakhstan. Dracocephalum nutans (Lamiaceae). 

Jaapiella dracocephalicola Fedotova 1998b: 10. Kazakhstan: southern Altaï, Markakol'skii Res., L. Markakol', nr 


*dracocephalis. Kazakhstan. Dracocephalum integrifolium (Lamiaceae). 

Jaapiella dracocephalis Fedotova 2005b: 903. Kazakhstan: Zailiiskii Alatau, Toraigyr Mts., 8 km from Kok-Pek 

Village; HT; ♂; ZMAS. 

floriperda. Widespread Europe, Kazakhstan. Silene inflata; S. vulgaris, S. wolgensis (Caryophyllaceae). 

Cecidomyia floriperda Löw 1888: 231. Austria: nr Kaltenleutgeben; ST; ♂,♀,L; NHMW. 

Cecidomyia bergrothiana Mik 1889: 236. Austria; ST; ♂,♀; NHMW. 

*galatellagemmae. Kazakhstan. Galatella punctata (Asteraceae). 

Jaapiella galatellagemmae Fedotova 2003a: 828. Kazakhstan: northwestern Dzhungarskiï Alatau, Lepsy R., nr 

Petropavlovka-Cherkasskaya; HT; ♂; ZMAS. 

genistamtorquens. Widespread Europe. Genista pilosa (Fabaceae). 

Cecidomyia genistamtorquens Kieffer 1888d: 311. France: Bitche; ST; ♂,♀,L; lost?. 

genisticola. Widespread Europe, Russia (Western Siberia). Genista tinctoria (Fabaceae). 

Cecidomyia genisticola Löw 1877: 4. Austria: vic. Klosterneuberg nr Vienna; ST; ♂,♀,L; NHMW. 

geraniifolia. Kazakhstan. Geranium collinum (Geraniaceae). 

Jaapiella geraniifolia Fedotova 1993o: 23. Kazakhstan: Kungeï Alatau, 12 km SW Malyï Zhalanash; HT; ♂; ZMAS. 

glycyrrhizae. Kazakhstan. Glycyrrhiza uralensis; G. glabra (Fabaceae). 

Jaapiella glycyrrhizae Fedotova 1983b: 44. Kazakhstan: along Karatal R., nr Golovnoe; HT; ♂; ZKAS.

254 

goebeliae. Kazakhstan. Goebelia alopecuroides (Fabaceae). 

Jaapiella goebeliae Fedotova 1987a: 37. Kazakhstan: right bank mid-Ili R., Kalkan-Mataï valley, Kalinino; HT; ♂; 

ZMAS. 

halimodendronis. Kazakhstan. Halimodendron halodendron (Fabaceae). 

Jaapiella halimodendronis Fedotova 1990f: 80. Kazakhstan: Karatal R. valley, 60 km NW Ush-Tobe; HT; ♂; 

ZMAS. 

hedickei. Widespread Europe. Pimpinella saxifraga (Apiaceae). 

Jaapiella hedickei Rübsaamen 1921: 50. Poland: nr Szczecin; ST; ♂,♀,L; ZMHB. 

hypochoeridis. Germany, Sweden. Hypochoeris radicata (Asteraceae). 

Jaapiella hypochoeridis Sylvén 1998b: 252. Germany: Husum; HT; ♀; NHRS. 

inflatae. UK, Germany, Latvia. Silene inflata (Caryophyllaceae). 

Dasyneura inflatae Rübsaamen 1914: 108. Germany: nr Freiberg; ST; ♂,♀,L; ZMHB. 

inulicola. Denmark, Sweden, Kazakhstan. Inula salicina (Asteraceae). 

Jaapiella inulicola Fedotova 1993l: 84. Kazakhstan: Ili River valley, 20 km NW Bakanas; HT; ♂; ZMAS. 

ivannikovi. Kazakhstan. Acroptilon picris (Asteraceae). 

Jaapiella ivannikovi Fedotova 1985f: 37. Kazakhstan: Western Tyan'-Shan', Karzhantau Mtn Range, Kyryk-Kyz, nr 

Turbat; HT; ♂; ZKAS. 

jaapiana. Widespread Europe. Medicago lupulina (Fabaceae). 

Dasyneura jaapiana Rübsaamen 1914: 107. Germany: Prignitz, Triglitz; ST; ♂,♀; ZMHB. 

jurineae. Kazakhstan. Jurinea multiflora (Asteraceae). 

Jaapiella jurineae Fedotova 1993l: 85. Kazakhstan: Tarbagataï Mtn Range, 101 km E Ayaguz; HT; ♂; ZMAS. 

karkarensis. Kazakhstan. Rumex pamiricus (Polygonaceae). 

Jaapiella karkarensis Fedotova 1997f: 453. Kazakhstan: Terskey Alatau, upper reaches Karkara R., 45 km S Keghen; 


knautiae. UK, Sweden, Germany, Latvia, Czechoslovakia, Romania. Knautia arvensis (Dipsacaceae). 

Jaapiella knautiae Rübsaamen 1917: 40. Germany: Jena; ST; ♂,♀,L; ZMHB. 

konyrtauensis. Kazakhstan. Serratula kirghisorum (Asteraceae). 

Jaapiella konyrtauensis Fedotova 1993l: 88. Kazakhstan: Konyrtau Mts, 10 km SW Sarybien; HT; ♂; ZMAS. 

kovalevi. Kazakhstan. Berberis oblonga; B. heteropoda (Berberidaceae). 

Jaapiella kovalevi Fedotova 1990f: 86. Kazakhstan: 40 km SW Chundzha, floodplain Timirlik R.; HT; ♂; ZMAS. 

lactucicola. Kazakhstan. Lactuca tatarica (Asteraceae). 

Jaapiella lactucicola Fedotova 1996b: 200. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE Alma-Ata; 


loniceriflora. Kazakhstan. Lonicera tatarica (Caprifoliaceae). 

Jaapiella loniceriflora Fedotova 1990f: 83. Kazakhstan: Tarbagataï Mtn Range, 40 km NW Urdzhara, vic. 

Blagodatnoye; HT; ♂; ZMAS. 

loticola. Widespread Europe; Russia (Western Siberia). Lotus corniculatus; L. uliginosus (Fabaceae). 

Cecidomyia loticola Rübsaamen 1889a: 52. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

Perrisia loti Kieffer 1909b: 14. Italy; ST; LT; lost?. 

microphysae. Kazakhstan. Microphysa elongata (Rubiaceae). 

Jaapiella microphysae Fedotova 1997c: 333. Kazakhstan: delta Ili R., 15 km NW Aral-Tyube; HT; ♀; ZMAS. 

moraviae. Czech Republic. Lychnis viscaria (Caryophyllaceae). 

Cecidomyia moraviae Wachtl 1883: 478. Czech Republic: Znojmo; ST; ♂,L; NHMW?. 

narymica. Kazakhstan. Cirsium serratuloides (Asteraceae). 

Jaapiella narymica Fedotova 1996b: 199. Kazakhstan: Narymskiï Mtn Range, 15 km S Bol'shenarymskoe; HT; ♂; 

ZMAS. 

narynensis. Kazakhstan. Serratula coronata (Asteraceae). 

Jaapiella narynensis Fedotova 1993l: 87. Kazakhstan: Narym Mts, 15 km S Bol'shenarymskiy; HT; ♂; ZMAS. 

*oxytropicarpa. Kazakhstan. Oxytropis pseudofrigida (Fabaceae). 

Jaapiella oxytropicarpa Fedotova 2005b: 905. Kazakhstan: Saur Mt. Range, 20 km S Zaisan, nr. Zhanaturmys 

Village; HT; ♂; ZMAS. 

parvula. UK, Netherlands, France, Portugal, Libya. Bryonia dioica (Cucurbitaceae).

255 

Cecidomyia parvula Liebel 1889b: 283. France: Bitche; ST; ♂,♀,L; ?. 

pentaphylloidis. Kazakhstan. Pentaphylloides phyllocalyx (Rosaceae). 

Jaapiella pentaphylloidis Fedotova 1990f: 88. Kazakhstan: western Tyan'-Shan', Karzhantau Mtn Range, Ular Mtn, 

Saryaïgyr canyon, 15 km SE Kaskasu; HT; ♂; ZMAS. 

picridis. Germany. Picris hieracioides (Asteraceae). 

Dasyneura picridis Rübsaamen 1912e: 287. Germany: Rhine Prov.; ST; ♀,L; ZMHB. 

pilosellae. Sweden. Hieracium pilosella (Asteraceae). 

Jaapiella pilosellae Sylvén 1998b: 253. Sweden: Uppland, Danderyd; HT; ♀; NHRS. 

plantaginicola. Kazakhstan. Plantago maxima (Plantaginaceae). 

Jaapiella plantaginicola Fedotova 1996d: 14. Kazakhstan: Tarbagataï Mtn Range, 70 km E Ayaguz; HT; ♂; ZMAS. 

reichardiae. Malta, Croatia, Greece. Reichardia picroides (Asteraceae). 

Jaapiella reichardiae Sylvén 1998b: 255. Greece: Delphi; HT; ♀; NHRS. 

rubiae. Kazakhstan. Rubia tinctorum (Rubiaceae). 

Jaapiella rubiae Fedotova 1987a: 38. Kazakhstan: upper Charyn R., Sartogaï, nr Chundzha; HT; ♂; ZKAS. 

rubicundula. UK, Germany, Poland, Czechoslovakia, Yugoslavia, Russia (Eur). Rumex acetosella (Chenopodiaceae). 

Cecidomyia rubicundula Rübsaamen 1891d: 131. Germany: betw. Siegen & Buschgotthardshütten; ST; ♂,♀,L; 

ZMHB. 

ruthenicae. Kazakhstan. Centaurea ruthenica (Asteraceae). 

Jaapiella ruthenicae Fedotova 1995e: 21. Kazakhstan: Tarbagataï Mtn Range, 7 km N Blagodarnos, upper 

Kel'dymurat R.; HT; ♂; ZMAS. 

sanguisorbacola. Kazakhstan. Sanguisorba alpina (Rosaceae). 

Jaapiella alpina Fedotova 1993i: 48, jun. sec. hom. alpina Löw. Kazakhstan: Sarymsakty Mtn Range, 45 km SW 

Katon-Karagaya, nr Ornek; HT; ♂; ZMAS. 

Jaapiella sanguisorbacola Fedotova 1996d: 13, new name for Jaapiella alpina Fedotova. 

sarothamni. UK, Germany. Sarothamnus scoparius (Fabaceae). 

Jaapiella sarothamni Rübsaamen 1917: 39. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

scabiosae. UK, France, Germany, Poland, Czechoslovakia, Austria. Scabiosa columbaria (Dipsacaceae). 

Cecidomyia scabiosae Kieffer 1888e: 97. France: Lorraine; ST; ♂,♀,L; lost?. 

schmidti. Widespread Europe. Plantago lanceolata (Plantaginaceae). 

Dasyneura schmidti Rübsaamen 1912e: 284. Germany: nr Werlau, vic. St. Goar; ST; ♂,♀,L; ZMHB. 

schrenkiana. Kazakhstan. Lonicera microphylla (Caprifoliaceae). 

Jaapiella schrenkiana Fedotova 1990f: 85. Kazakhstan: Dzhungarskiï Alatau, Altyn-Emel' Pass, vic. Konurolen; HT; 

♂; ZMAS. 

sileneicola. Kazakhstan. Silene odoratissima; S. gebleriana (Caryophyllaceae). 

Jaapiella sileneicola Fedotova 1994e: 45. Kazakhstan: Barsuki desert, 76 km NW Aral'ska; HT; ♂; ZMAS. 

sphaerophysaflorae. Kazakhstan. Sphaerophysa salsula (Fabaceae). 

Jaapiella sphaerophysaflorae Fedotova 1992g: 44. Kazakhstan: Karatal R., nr Kopbirlik; HT; ♂; ZMAS. 

spungisi. Kazakhstan. Geranium collinum (Geraniaceae). 

Jaapiella spungisi Fedotova 1993o: 22. Kazakhstan: Zailiïskiï Alatau, 32 km S Turgen'; HT; ♂; ZMAS. 

tarbagataica. Kazakhstan. Caragana frutex (Fabaceae). 

Jaapiella tarbagataica Fedotova 1990f: 79. Kazakhstan: Tarbagataï Mtn Range, Ashchysu R. floodplain, 30 km N 

Ayaguza; HT; ♂; ZMAS. 

tekesensis. Kazakhstan. Lonicera alberti (Caprifoliaceae). 

Jaapiella tekesensis Fedotova 1990f: 82. Kazakhstan: floodplain Tekes R., vic. Tekes, 20 km NW Narynkol; HT; ♂; 

ZMAS. 

thalictri. UK, Sweden, Netherlands, Germany, Poland, Czechoslovakia, Russia (Far East). Thalictrum flavum 


Dichelomyia thalictri Rübsaamen 1895a: 181. Germany: Berlin; ST; ♂,♀,L; ZMHB. 

thyrsiflora. Kazakhstan. Rumex thyrsiflorus (Polygonaceae). 

Jaapiella thyrsiflora Fedotova 1997f: 451. Kazakhstan: Listvyaga Mtn Range, nr Rakhmanovskie, 30 mi E Berel'; 


vacciniorum. Widespread Europe. Vaccinium myrtillus (Ericaceae).

256 

Dichelomyia vaccinii Rübsaamen 1895c: 258, jun. sec. hom. Cecidomyia vaccinii Smith. Germany; ST; ♂,♀,L; 

ZMHB. 

Perrisia vacciniorum Kieffer 1913i: 83, new name for Dichelomyia vaccinii Rübsaamen. 

veronicae. Widespread Europe. Veronica chamaedrys, V. spicata (Scrophulariaceae). 

Cecidomyia veronicae Vallot 1827: 93. France; ST; A; lost?. 

Cecidomyia veronicae Bremi 1847a: 49. Switzerland: Zurich; ST; A,L; ETHZ. 

Cecidomyia chamaedrys Inchbald 1860b: 196. UK: England, nr Huddersfield; ST; A; ?. 

Perrisia spicatae Kieffer 1909b: 32. Germany; T; G; lost?. 

viciafoliae. Kazakhstan. Vicia tetrasperma (Fabaceae). 

Jaapiella viciafoliae Fedotova 1992f: 45. Kazakhstan: Saur Mtn Range, 20 km S Zaïsan, nr Zhanaturmys; HT; ♂; 

ZMAS. 

viscariae. France, Czechoslovakia. Lychnis viscaria (Caryophyllaceae). 

Cecidomyia viscariae Kieffer 1886: 332. France: vic. Bitche; ST; ♀,L; lost?. 

*volgensis. Russia. Filipendula vulgaris (Rosaceae). 

Jaapiella volgensis Fedotova 2008: 123. Russia: Central Povolzhye, 30 km SE of Samara R., nr Ust-Kinelskii; HT; 

♂; ZMAS. 

volvens. UK, Germany, Poland, Czechoslovakia, Yugoslavia, Ukraine, Russia (Europe, Western Siberia). Lathyrus 

pratensis (Fabaceae). 

Jaapiella volvens Rübsaamen 1917: Germany: Prignitz, Triglitz, & Steinau; ST; ♂,♀,L; ZMHB. 

zygophylliflora. Kazakhstan. Zygophyllum oxianum (Zygophyllaceae). 

Jaapiella zygophylliflora Fedotova 1985f: 40. Kazakhstan: right bank Syr Darya R., 30 km SW Turkestan, nr 

Baltakol'; HT; ♂; ZKAS. 

JANETIA 

Janetia Kieffer 1896f: 236. Type species, Lasioptera cerris Kollar (subs. des. Skuhravá 1986: 177). 

Kiefferiola Tavares 1920b: 43. Type species, Arnoldia panteli Kieffer (orig. des.). 

Palearctic; 7 spp., from leaf galls on Quercus; Lasiopteridi, Dasineurini. 

cerris. Widespread Europe, Japan?. Quercus cerris (Fagaceae). 

Lasioptera cerris Kollar 1850: 349. Austria; ST; ♂,♀; ? 

Cecidomyia cerrifoliae Passerini 1850: 475. Italy: vic. Florence; ST; G; lost. 

homocera. Italy, Czechoslovakia, Austria, Hungary, Romania, Bulgaria, Yugoslavia, Greece. Quercus cerris 

(Fagaceae). 

Cecidomyia homocera Löw 1877: 8. Austria: vic. Vienna; ST; ♂,♀,L; NHMW. 

nervicola. Czechoslovakia, Austria, Hungary, Romania, Bulgaria, Yugoslavia. Quercus cerris (Fagaceae). 

Arnoldia nervicola Kieffer 1909b: 21. Austria; T; LT; lost?. 

panteli Spain. Quercus lusitanica var. faginea (Fagaceae). 

Arnoldia panteli Kieffer 1909b: 23. Spain; T; LT; lost?. 

plicans. Austria, Greece. Quercus cerris (Fagaceae). 

Arnoldia plicans Kieffer 1909b: 21. Austria; ST; LT; lost?. 

pustularis. Czechoslovakia, Austria, Hungary, Yugoslavia, Greece. Quercus cerris (Fagaceae). 

Arnoldia pustularis Kieffer 1909b: 22. Austria; T; LT; NHMW?. 

szepligetii. Italy, Czechoslovakia, Hungary, Romania, Bulgaria, Yugoslavia, Greece, Turkey. Quercus cerris 

(Fagaceae). 

Janetia szepligetii Kieffer 1896f: 236. Hungary; ST; A,L; lost?. 

JANETIELLA 

Janetiella Kieffer 1898f: 23. Type species, Cecidomyia thymi Kieffer (subs. des. Coquillett 1910: 556). 

Ceratomyia Fedotova 1991d: 19, preocc. Felt 1911. Type species, Janetiella gemmicola Marikovsky.

Holarctic; 24 spp; Lasiopteridi, Dasineurini. The Nearctic species are a diverse assemblage placed here only because 

they lack basal teeth on the tarsal claws. Many otherwise share a resemblance to Dasineura spp. Ceratomyia Fedotova 

was placed as a new synonym in Gagné (2004b) because its uniqueness was not apparent from the description and the 

action precluded having to rename the preoccupied Ceratomyia. 

acerifolii. USA (New York). 

Oligotrophus acerifolii Felt 1907a: 27. USA: New York, Albany; HT; ♂; NYSM. 

Oligotrophus acerifolius Felt 1907c: 124, unjust. emend. of acerifolii. 


Janetiella americana Felt 1908b: 372. USA: New York, Albany; HT; ♂; NYSM. 

asplenifolia. USA (Massachusetts). 

Oligotrophus asplenifolia Felt 1907c: 159. USA: Massachusetts, Worcester; ST; ♂,♀; NYSM. 

breviaria. USA (New York). 

Janetiella breviaria Felt 1908b: 372. USA: New York, Albany; HT; ♀; NYSM. 

coloradensis. USA (Colorado, Arizona). Pinus sp. (Pinaceae). 

Janetiella coloradensis Felt 1912h: 148. USA: Colorado, Denver; ST; ♀,L; NYSM. 

euphorbiae. Italy, Greece. Euphorbia characias (Euphorbiaceae). 

Janetiella euphorbiae De Stefani 1908a: 108. Italy: Sicily, Mt Renda nr Monreale; ST; ♂,♀,L; lost. 

fallax. Germany, Hungary, Bulgaria, Albania. Alyssum arenarium (Brassicaceae). 

Janetiella fallax Kieffer 1904a: 69. Germany: vic. Kreuznach; ST; ♂,♀,P,L; lost?. 

foliicola. Ukraine. Onobrychis sp. (Fabaceae). 

Janetiella foliicola Marikovskij 1956: 185. Ukraine: Kiyev Prov., Belaya Tserkov'; ST; ♂,♀; ZMAS. 

fortiana. Greece. Erysimum graecum (Brassicaceae). 

Janetiella fortiana Trotter 1901: 346. Greece: Acropolis of Corinth; ST; ♂,L; ?. 

*frankumi. UK. Rosa pimpinellifolia (Rosaceae). 

Janetiella frankumi Harris 2003b: 52. UK: Wales, Llanbedr, Shell I.; HT; ♂; BMNH. 

gemmicola. Kazakhstan. Eurotia ceratoides (Chenopodiaceae). 

Janetiella gemmicola Marikovskij 1961: 47. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, middle & lower 


genistae. France. Genista corsica (Fabaceae). 

Janetiella genistae Kieffer 1909b: 9. France; ST; LT; lost?. Skuhravá 1986: 242 places this species as jun. syn. of 

Contarinia melanocera Kieffer. 

glechomae. France. Glechoma hederacea (Lamiaceae). 

Janetiella glechomae Tavares 1930: 153. France: Seine Maritime, Bois Guilbert; ST; ♂,♀; TAVC. 

goiranica. Italy. Carpesium cernuum (Asteraceae). 

Janetiella goiranica Kieffer & Trotter 1905: 54. Italy: Treviso, Volpago For.; ST; ♀,L; lost?. 

infrafoli. Japan (Honshu to Kyushu). Fagus crenata (Fagaceae). 

Janetiella infrafoli Monzen 1955a: 47. Japan: Honshu, Aomori, Iwate, Tochigi, Wakayama, & Tottori Prefs; ST; 

♂,♀; KUEC. 

inquilina. USA (New York). Solidago sp. (Asteraceae). 

Oligotrophus inquilinus Felt 1908b: 368. USA: New York, Albany; HT; ♂; NYSM. 

kimurai. Japan (Honshu). Pinus parviflora (Pinaceae). 

Janetiella kimurai Inouye 1964: 22. Japan: Honshu, Miyagi Pref.; HT; ♂; HFES. 

lemeei. Widespread Europe; Turkey. Ulmus spp. (Ulmaceae). 

Oligotrophus lemeei Kieffer 1904a: 71. France: Aisne, Ardennes, Orne, & Seine-Inférieure; ST; ♂,♀,P,L; lost?. 

maculata. Portugal, Spain. Cytisus albus (Fabaceae). 

Janetiella maculata Tavares 1901: 65. Portugal: S. Fiel; ST; ♂,L; TAVC. 

sanguinea. USA (New York). 

Janetiella sanguinea Felt 1908b: 372. USA: New York, Nassau; HT; ♀; NYSM. 

siskiyou. USA (Washington, Oregon); introd.: UK, Netherlands, France, Denmark, Poland, Czechoslovakia, Cyprus. 

257

258 

Chamaecyparis lawsoniana (Cupressaceae). 

Janetiella siskiyou Felt 1917b: 194. USA: Oregon, 1 mi N Siskiyou Natl For. along south fork Coquille R.; ST; ♀,L; 


Craneiobia lawsonianae Meijere 1935: 129. Netherlands: Wageningen; ST; ♂,♀,P,L; ZMAN. 

thymi. Widespread Europe. Thymus serpyllum, T. chamaedrys (Lamiaceae). 

Cecidomyia thymi Kieffer 1888e: 100. France: Bitche; ST; ♂,♀,L; lost?. Ref.: Gagné 2009b: 400 (redescr.). 

tuberculi. UK, France, Germany, Poland, Portugal. Sarothamnus scoparius (Fabaceae). 

Cecidomyia tuberculi Rübsaamen 1889b: 381. Germany; ST; ♀,L; ZMHB. 

ulmii. Canada (Ontario), USA (New York, Maryland). Ulmus americana (Ulmaceae). 

Cecidomyia ulmii Beutenmüller 1907a: 387. USA: New York, New York; ST; ♂,♀; AMNH. 

JORGENSENIA 

Jorgensenia Kieffer 1913i: 148. Type species, Centrodiplosis falcigera Kieffer & Jörgensen (mon.). 


falcigera. Argentina (Mendoza). Lycium gracile (Solanaceae). 

Centrodiplosis falcigera Kieffer & Jörgensen 1910: 411. Argentina: Mendoza; T; ♀; lost. 

*JORGENSENIELLA 

Jorgenseniella Maia 2005a: 350. Type species, Jorgenseniella eugeniae Maia (orig. des.). 

Neotropical; 1 sp.; Cecidomyiidi, unplaced Lasiopteridi. 

*eugeniae. Brazil (Rio de Janeiro). Eugenia umbelliflora & E. rotundifolia (Myrtaceae). 

Jorgenseniella eugeniae Maia 2005a: 351. Brazil: Rio de Janeiro, Arraial do Cabo; HT; ♂; MNRJ. 

KALTENBACHIOLA 

Kaltenbachiella Rübsaamen 1916b: 490, preocc. Schouteden 1906. Type species, Cecidomyia strobi Winnertz (orig. 

des.). 

Kaltenbachiola Hedicke 1938b: 297, new name for Kaltenbachiella Rübsaamen. 

Holarctic; 3 spp., from Picea cones; Lasiopteridi, Dasineurini. 

canadensis. Canada (Saskatchewan, Ontario). Picea glauca (Pinaceae). 

Dasyneura canadensis Felt 1907c: 157. Canada: Ontario, vic. Ottawa; ST; ♂,♀; NYSM. 

rachiphaga. Canada (Ontario, Quebec). Picea glauca (Pinaceae). 

Dasyneura rachiphaga Tripp 1955: 259. Canada: Ontario, Kemptville; HT; ♂; CNC. 

strobi. Widespread Europe. Picea abies (Pinaceae). 

Cecidomyia strobi Winnertz 1853: 234. Germany; ST; ♂,♀; lost. 

KAMPTODIPLOSIS 

Kamptodiplosis Felt 1918c: 290. Type species, reducta Felt (orig. des.). 


ramanathi. India (Uttar Pradesh). undet. bamboo (Poaceae). 

Kamptodiplosis ramanathi Mani 1963: 13. India: Uttar Pradesh, Dehra Dun, New Forest; HT; ♂; ZSCI.

259 

reducta. Philippines. Siphonodon celastrineus (Celastraceae). 

Kamptodiplosis reducta Felt 1918c: 290. Philippines: Luzon, Laguna Prov., Los Baños, Balong Bulo Hill; L (Gagné 

1969e: 566); ♂; NYSM. 

KARSHOMYIA 

Karshomyia Felt 1908b: 398. Type species, Mycodiplosis viburni Felt (orig. des.). 

Lobodiplosis Felt 1908b: 397. Type species, Mycodiplosis acerina Felt (orig. des.). 

Metadiplosis Felt 1908b: 406. Type species, spinosa Felt (orig. des.). 

Karschomyia Felt 1911c: 54, unjust. emend. of Karshomyia. 

Plesiobremia Kieffer 1912h: 1. Type species, Bremia ramosa Kieffer (orig. des.). 

Tristephanus Kieffer 1913h: 106. Type species, atricauda Kieffer (orig. des.). 

Tristephanes Shinji 1944: 244, missp. of Tristephanus. 

Kartschomyia Mani 1946: 215, missp. of Karshomyia. 

Hiastatus Marikovskij 1956: 188. Type species, concinnus Marikovskij (orig. des.). 

Tristephana Marikovskij 1960: 185, missp. of Tristephanus. 

Apamargamyia (also as Apramargamyia) Sharma & Rao 1978: 287. Type species, orientalis Sharma & Rao (orig. 

des.). 

Holarctic, Neotropical, Afrotropical, Oriental; 49 spp.; Cecidomyiidi, Karshomyiini. Several species have been reared 

from mushrooms (Bruns 1982), others were taken in association with decaying plant matter. 

abnormis. Poland, Russia (Europe). 

Hiastatus abnormis Mamaev 1961c: 449. Russia: Moscow Prov., Glubokoye L.; HT; ♂; HYNM. 

acerina. USA (New York). 

Mycodiplosis acerina Felt 1907a: 33. USA: New York, Albany; HT; ♂; NYSM. 

acietata. Russia (Far East). 

Karschomyia acietata Mamaev & Krivosheina 1998: 78. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMMU. 

*alata. Russia (Far East). 

Karshomyia alata Fedotova 2004g: 8. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

apicalis. India (Uttar Pradesh). 

Karschomyia apicalis Grover & Bakhshi 1978: 145. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; PGCA. 

apiculata. Russia (Far East). Ulmus sp. (Ulmaceae). 

Karschomyia apiculata Mamaev & Krivosheina 1998: 77. Russia: Primorskiï Terr., Suputinskiy Res.; HT; ♂; 

ZMMU. 

arcuata. India (Uttar Pradesh). 

Therodiplosis arcuata Grover & Bakhshi 1978: 47. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; PGCA. 

*asyndeta. Russia (Far East). 

Karshomyia asyndeta Fedotova & Sidorenko 2009a: 8. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


atricauda. Kenya. 

Tristephanus atricauda Kieffer 1913h: 107. Kenya: Kyambu Distr., 25 km from Nairobi, & Bura, on road to 

Kilimanjaro, west of Voi; ST; ♂,♀; MNHN. 


Lobodiplosis borealis Felt 1920a: 289. USA: New York, Keene Valley; HT; ♂; NYSM. 

caulicola. USA (New Hampshire), UK, Netherlands, Germany, Lithuania, Ukraine, Russia (Europe). Papaver 

nudicaule (Papaveraceae). 

Diplosis caulicola Coquillett 1895b: 401. USA: New Hampshire, Isles of Shoals; ST; ♂,♀; USNM. 

Hiastatus concinnus Marikovskij 1956: 189. Russia: Leningrad Prov.; ST; ♂,♀; ZMAS. 

ciliata. Latvia, Lithuania, Russia (Europe). 

Karschomyia ciliata Mamaev & Krivosheina 1998: 78. Russia: Vologda Prov., Kadnikovskaya; HT; ♂; ZMMU.

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curiosa. USA (Maryland, North Carolina). Orig. unk.; Paxillus involutus (Boletales). 

Karschomyia curiosa Gagné 1973d: 348. USA: Maryland, Montgomery Co., Wheaton Pk; HT; ♂; USNM. 

curvidentata. Latvia, Russia (Europe). 

Hiastatus curvidentatus Mamaev 1961c: 448. Russia: Moscow Prov., Pavlovskaja Sloboda; HT; ♂; HYNM. 

ectopia. USA (Virginia). Orig. unk.; Strobilocyces sp. (Boletales). 

Karschomyia ectopia Gagné 1973d: 348. USA: Virginia, Holmes Run; HT; ♂; USNM. 

figurata. Latvia, Russia (Western Siberia). 

Karschomyia figurata Mamaev & Krivosheina 1998: 78. Russia: Gorno-Altayskaya Aut. Reg., Teletskoye L., 

Artybash; HT; ♂; ZMMU. 

garhwalensis. India (Uttar Pradesh). 

Karschomyia garhwalensis Jaiswal 1988a: 103. India: Uttar Pradesh, Garhwal Hills, Kotdwara; HT; ♂; CSIA. 

insolens. USA (New York). 

Hyperdiplosis insolens Felt 1920a: 291. USA: New York, Keene Valley; HT; ♂; NYSM. 

inusitata. USA (Louisiana, Maryland, South Carolina). Orig. unk.; Boletus sp., Suillus sp. (Boletales). 

Karschomyia inusitata Gagné 1973d: 351. USA: Louisiana, Elizabeth; HT; ♂; USNM. 

longilobis. India (Uttar Pradesh). 

Karschomyia longilobis Jaiswal 1988a: 101. India: Uttar Pradesh, Garhwal Hills, Kotdwara; HT; ♂; CSIA. 

*mamaevi. Russia (Far East). 

Karshomyia mamaevi Fedotova & Sidorenko 2004c: 6. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


marikovskii. Germany, Lithuania, Russia (Europe). 

Hiastatus marikovskii Mamaev 1961c: 447. Russia: Moscow Prov., Bykovo; HT; ♂; HYNM. 

*melioria. Russia (Far East). 

Karshomyia melioria Fedotova & Sidorenko 2004c: 138. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

mira. USA (Maryland). 

Karschomyia mira Gagné 1973d: 351. USA: Maryland, Silver Spring; HT; ♂; USNM. 

*obstructa. Russia (Far East). 

Karshomyia obstructa Fedotova 2004g: 8. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

oklandi. Norway. 

Karschomyia oklandi Mamaev & Krivosheina 1998: 79. Norway: Roelingen, Tappenberg; HT; ♂; NFRI. 

orientalia. India (Maharashtra). Achyranthes aspera (Amaranthaceae). 

Apamargamyia orientalis Sharma & Rao 1978: 287, jun. sec. hom. orientalis Grover. India: Maharashtra, 

Aurangabad Univ; HT; ♂; RCMU. The female associated with this species apparently belongs to Contarinia. 

Karshomyia orientalia Gagné 2004b: 174, new name for Apamargamyia orientalis Sharma & Rao. 


Macrodiplosis orientalis Grover 1965a: 37. India: Uttar Pradesh, Allahabad, Tagore Town; HT; ♂; PGCA. 

peculiaris. India (Tripura). 

Karschomyia peculiaris Chandra 1997: 98. India: Tripura, Agartala; HT; ♂; CSIA. 

perissa. USA (South Carolina). 

Karschomyia perissa Gagné 1973d: 351. USA: South Carolina, Clemson; HT; ♂; USNM. 

praecipua. USA (California). 

Karschomyia praecipua Gagné 1973d: 351. USA: California, Sonoma Co.; HT; ♂; USNM. 

producta. Latvia, Ukraine, Russia (Europe, Far East). 

Karschomyia producta Mamaev & Krivosheina 1998: 79. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; 

ZMMU. 

*pusilla. Russia (Far East). 

Karshomyia pusilla Fedotova & Sidorenko 2004c: 141. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


quadrii. India (Uttar Pradesh). 

Karschomyia quadrii Jaiswal 1988a: 99. India: Uttar Pradesh, Garhwal Hills, Kotdwara; HT; ♂; CSIA.

261 

quercina. USA (New York, Massachusetts). 

Mycodiplosis quercina Felt 1907a: 33. USA: New York, Albany; HT; ♂; NYSM. 

ramosa. UK, France, Latvia, Lithuania, Russia (Europe, Far East). 

Bremia ramosa Kieffer 1904c: 394. France: vic. Bitche; ST; ♂,♀; lost?. 

Karschomyia elegans Mamaev 1961d: 229. Russia: Moscow Prov. & Voronezh Prov.; ST; ♂; MAMC. 

setosa. Russia (Europe, Far East). 

Karschomyia setosa Mamaev & Krivosheina 1998: 80. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; 

ZMMU. 

speciosa. USA (Massachusetts). 

Lobodiplosis speciosa Felt 1913f: 143. USA: Massachusetts, North Adams; HT; ♂; NYSM. 

spinosa. USA (Minnesota, New York). 

Metadiplosis spinosa Felt 1908b: 406. USA: New York, Albany; HT; ♂; NYSM. 

spinulifera. Russia (Europe, Far East). 

Karschomyia spinulifera Mamaev & Krivosheina 1998: 81. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMMU. 

spinulosa. Antigua. 

Lobodiplosis spinosa Felt 1909a: 301, jun. sec. hom. spinosa Felt 1908. Antigua: St Johns; L (Gagné 1994a: 190); ♂; 


Karschomyia spinulosa Gagné 1994a: 190, new name for Lobodiplosis spinosa Felt. 

spungisi. Latvia, Lithuania, Russia (Europe). 

Karschomyia spungisi Mamaev & Krivosheina 1998: 77. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; 

ZMMU. 

*striatella. Russia (Far East). 

Karshomyia striatella Fedotova & Sidorenko 2004c: 4. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 


terminalis. India (Uttar Pradesh). 

Lobodiplosis terminalis Grover & Bakhshi 1978: 151. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; PGCA. 

triangularis. USA (Massachusetts). 

Lobodiplosis triangularis Felt 1918e: 163. USA: Massachusetts, Mt Greylock; HT; ♂; NYSM. 

twiarcus. India (Uttar Pradesh). 

Karshomyia twiarcus Grover & Kashyap 1991: 88. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

viburni. Canada (British Columbia), USA (California, New York, Maine, Maryland, South Carolina), Russia (Europe, 

Far East). 

Mycodiplosis viburni Felt 1907a: 34. USA: New York, Albany; HT; ♂; NYSM. 

Karschomyia aceri Mamaev 1960a: 1523. Russia: Voronezh Prov., Tellerman; HT; ♂; HYNM. 

xylogena. Russia (Far East). 

Karschomyia xylogena Mamaev & Krivosheina 1998: 81. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMMU. 

xylophila. Russia (Europe). 

Hiastatus xylophilus Mamaev 1961c: 446. Russia: Caucasus Mts, Kudepsta; HT; ♂; HYNM. 

KIEFFERIA 

Kiefferia Mik 1895: 96. Type species, Asphondylia pimpinellae Loew (orig. des.) = Kiefferia pericarpiicola Bremi. 

Palearctic; 1 sp.; Cecidomyiidi, Asphondyliini, Schizomyiina. 

pericarpiicola. Widespread Europe; Russia (Far East). Daucus carota; many other genera of Apiaceae (Barnes 1946a). 

Cecidomyia pericarpiicola (also as dauci) Bremi 1847a: 21. Switzerland: Zurich; ST; LT; ETHZ?. 

Cecidomyia pimpinellae Loew 1850a: 30. Poland; T; G; ZMHB?. 

Cecidomyia tysselini Loew 1950: 22. Poland; ST; G; ZMHB?. 

Asphondylia pimpinellae Löw 1874b: 326. Austria: Pfalzauthale, nr Vienna; ST; ♂,♀,L; NHMW.

Asphondylia umbellatarum Löw 1877: 31, new name for Asphondylia pimpinellae Löw. 

KIMADIPLOSIS 

Kimadiplosis Gagné 1993c: 118. Type species, diversa Gagné (orig. des.). 


diversa. Kenya. Acacia tortilis (Fabaceae). 

Kimadiplosis diversa Gagné 1993c: 118. Kenya: Kima, nr Sultan Hamid; HT; LT; USNM. 

*KITELLA 

Kitella Deshpande, Shaikh & Sharma. 2005: 80. Type species, orientalis Deshpande, Shaikh & Sharma (orig. des.). 


*orientalis. India (Maharashtra). 

Kitella orientalis Deshpande, Shaikh & Sharma 2005: 80. India: Maharashtra, Nanded Distr., Hadgaon; HT; ♂; 

Deshpande collection, Nanded. 

KOCHIOMYIA 

Turkmenomyia Marikovskij 1957b: 936, preocc. Paramonov 1938. Type species, lanugiviva Marikovskij (orig. des.). 

Kochiomyia Mamaev 1969a: 395, new name for Turkmenomyia Marikovskij. 

Desertovelum Mamaev 1972a: 897. Type species, stackelbergi Mamaev (orig. des.). 

Southcentral Palearctic; 7 spp., all from Chenopodiaceae; Cecidomyiidi, Asphondyliini, Schizomyiina. The 

distinctions between this genus and Pseudokochiomyia with its 5 spp., all also from Chenopodiaceae, seem trivial and a 

restudy may show that the two genera grade into one another. 

heptapotamica. Kazakhstan. Salsola paulsenii (Chenopodiaceae). 

Desertovelum heptapotamicum Fedotova 1982a: 44. Kazakhstan: Sary-Taukumy desert; HT; ♂; ZMAS. 

horaninoviae. Turkmenistan. Horaninovia ulicina (Chenopodiaceae). 

Kochiomyia horaninoviae Fedotova 1993p: 59. Turkmenistan: eastern Karakumy Desert, Sundukli Desert, nr Kyzyl- 

Burun L., 80 km SE Chardzheva; HT; ♂; ZMAS. 

kaplankyrum. Turkmenistan. Salsola orientalis (Chenopodiaceae). 

Kochiomyia kaplankyrum Mamaev 1988: 61. Turkmenistan: Tashauz Prov, Kankakyr; HT; ♂; ZMMU. 

kochiae. Hungary, Ukraine, Russia (Europe). Kochia prostrata (Chenopodiaceae). 

Rhopalomyia koehliae Kieffer 1909b: 12. Ukraine: Crimea; T; G; ZMHB?. 

Misospatha kochiae Kieffer 1913i: 46, justified emendation of koehliae. 

lanugiviva. Turkmenistan. Salsola rigida (Chenopodiaceae). 

Turkmenomyia lanugiviva Marikovskij 1957: 936. Turkmenistan: Amu-Darja R. delta; ST; ♂,♀; MARC. 

londesiae. Turkmenistan. Londesia eriantha (Chenopodiaceae). 

Kochiomyia londesiae Mamaev 1972a: 898. Turkmenistan: Mary, Akibay; HT; ♂; ZMAS. 

stackelbergi. Turkmenistan. Salsola pellucida (Chenopodiaceae). 

Desertovelum stackelbergi Mamaev 1972a: 897. Turkmenistan: Mary, Akibay; HT; ♂; ZMAS. 

*KOVALEVIOLA 

Kovaleviola Fedotova & Perkovsky 2008b: 65. Type species, injusta Fedotova & Perkovsky (orig. des.). 

262

263 

Rovno amber (Ukraine; late Eocene); 1 sp.; originally placed with the Brachineuridi but its placement below the level 

of subfamily Cecidomyiinae is not at all certain from its description. 

* † injusta. Rovno amber (Ukraine). 

Kovaleviola injusta Fedotova & Perkovsky 2008b: 66. Ukraine amber: Rovno; HT; ♀; UASK. 

KRONODIPLOSIS 

Kronodiplosis Felt 1918c: 289. Type species, uichancoi Felt (orig. des.). 


uichancoi. Philippines. Barringtonia luzonensis (Lecythidaceae). 

Kronodiplosis uichancoi Felt 1918c: 289. Philippines: Luzon, Laguna Prov., Los Baños; HT; ♂; NYSM. 

LABIATOMYIA 

Labiatomyia Fedotova 1987b: 32. Type species, Rhopalomyia hyssopi Fedotova (orig. des.). 

Palearctic; 6 spp., on Lamiaceae; Lasiopteridi, Dasineurini. 

dracocephalii. Kazakhstan. Dracocephalum bipinnatum (Lamiaceae). 

Labiatomyia dracocephalii Fedotova 1987b: 34. Kazakhstan: Kungeï Alatau, Kegenskiï pass, nr Kegen'; HT; ♂; ?. 

hyssopi. Kazakhstan. Hyssopus macranthus (Lamiaceae). 

Rhopalomyia hyssopi Fedotova 1983b: 45. Kazakhstan: Mt Bektay-Ata, 70 km N L. Balkhash; HT; ♂; ZMAS. 

marikovskii. Kazakhstan. Dracocephalum integrifolium (Lamiaceae). 

Labiatomyia marikovskii Fedotova 1987b: 35. Kazakhstan: Tarbagataï Mtn Range, 5 km NE Novotroiskoe, 55 km 

NW Urdzhar; HT; ♂; ZMAS. 

markakolica. Kazakhstan. Dracocephalum nutans (Lamiaceae). 

Labiatomyia markakolica Fedotova 1998b: 12. Kazakhstan: southern Altaï, Markakol'skii Res., L. Markakol', nr 


*nepetaeflorae. Kazakhstan. Nepeta mariae (Lamiaceae). 

Jaapiella nepetaeflorae Fedotova 2005b: 907. Kazakhstan: Aksu-DZhabagly Nature Res., 20 km E Novonikolaevka 

Village, Ulkun-Kaindy Canyon; HT; ♂; ZMAS. 

ziziphorae. Kazakhstan. Ziziphora clinopoidioides (Lamiaceae). 

Labiatomyia ziziphorae Fedotova 1987b: 32. Kazakhstan: Karatau Mtn Range, 12 km SW Leont'evkï; HT; ♂; ?. 

LAMPRODIPLOSIS 

Lamprodiplosis Kieffer 1913a: 229. Type species, Lestodiplosis rhopalothrix Kieffer (orig. des.). 

Lamplodiplosis Shinji 1944: 244, missp. of Lamprodiplosis. 

Western Palearctic; 1 sp.; Cecidomyiidi, Lestodiplosini. 

rhopalothrix. France. Rumex acetosella (Polygonaceae). 

Lestodiplosis rhopalothrix Kieffer 1904a: 77. France: vic. Bitche; T; ♂; lost?. 

*LANIDIPLOSIS 

Lanidiplosis Fedotova 2004f: 113. Type species, lata Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Aphidoletini. This genus does not essentially differ from Bremia.

*lata. Russia (Far East). 

Lanidiplosis latus Fedotova 2004f: 113. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; ZMAS. 

LASIODIPLOSIS 

Lepidodiplosis Kieffer 1912h: 2, preocc. Kieffer 1911. Type species, Chrysodiplosis homotomus Kieffer (orig. des.). 

Lasiodiplosis Kieffer 1913a: 232, new name for Lepidodiplosis Kieffer. 


homotomus. Seychelles Is. 

Chrysodiplosis homotomus Kieffer 1911: 319. Seychelles Is: Mahé, Cascade Estate; HT; ♂; BMNH. 

LASIOPTERA 

Lasioptera Meigen 1818: 88. Type species, Lasioptera picta Meigen (subs. des. Karsch 1877: 14; validated ICZN 

1970: 75) = rubi Schrank. 

Diomyza Meigen 1818: 88. Type species, Lasioptera picta Meigen (subs. des. Coquillett 1910: 533) = rubi Schrank. 

Westwood 1840: 126 designated Diomyza fuliginosa Stephens as type sp., which was but a nomen nudum. 

Dyomiza Bigot 1854: 462, missp. of Diomyza. 

Genista Bigot 1854: 462, nomen nudum. 

Prolasioptera Kieffer 1913g: 47. Type species, Lasioptera niveocincta Kieffer (mon) = francoisi Kieffer. 

Thomasiella Rübsaamen 1916b: 559. Type species, Cecidomyia eryngii Vallot (subs. des. Möhn 1968a: 69). 

Monilipennella Del Guercio 1918a: 151. Type species, trapezospatha Del Guercio (mon). 

Gastinella Del Guercio 1918a: 59. Type species, Lasioptera kiefferiana Del Guercio (mon.) = berlesiana Paoli. 

Lasioptesa Shinji 1939b: 567, missp. of Lasioptera. 

Lasioptere Shinji 1944: 56, missp. of Lasioptera. 

Lacioptera Shinji 1944: 138, missp. of Lasioptera. 

Lasipotera Shinji 1944: 148, missp. of Lasioptera. 

Neoprotoplonyx Rao 1952b: 33. Type species, indica Rao (orig. des.). 

Bimba Grover 1962d: 355. Type species, toombii Grover (orig. des.). 

Old World, Australia, and North America; 128 spp.; Lasiopteridi, Lasiopterini. Lasioptera is largely Old World with 

only a few North American species, which supports the assumption that the migration into the New World is a recent 

phenomenon in the history of the genus. While most species appear to be gall makers, particularly in stems, some 

species, e.g., L. cerasiphera, L. tibialis, and L. yadokariae, are successor species that live in galls vacated by the gall 

makers (Yukawa & Haitsuka 1994). At least one species, L. berlesiana, is a facultative predator (Solinas 1967). Refs.: 

Tastás-Duque & Sylvén 1989 (anatomy), Gagné 2004b (syn. notes.). 

abhamata. USA (New York). 

Lasioptera abhamata Felt 1907a: 11. USA: New York, Karner; HT; ♀; NYSM. 

achyranthesae. India (Maharashtra). Achyranthes aspera (Amaranthaceae). 

Lasioptera achyranthesae Sharma 1988b: 376. India: Maharashtra, Aurangabad, University Campus; HT; ♂; NZSI. 

achyranthii. Japan (Honshu to Kyushu), Republic of Korea. Achyranthes bidentata, A. japonica, A. longifolia 

(Amaranthaceae). 

Lasioptera achyranthii Shinji 1939b: 565. Japan: Honshu; ST; ♂,♀; lost. 

Lasioptera inokozuchii Shinji 1944: 328. Japan: Honshu; T; ♂; lost. 

Lasioptera achranthi Fedotova & Kovalev 2001: 510, missp. of achyranthii. 

aeschynanthusperottetti. India (Kerala). Aeschynanthus perottetti (Gesneriaceae). 

Prolasioptera aeschynanthusperottetti (as aeschynanthus-perottetti) Mani 1944: 151. India: Kerala, Western Ghats, 

Travancore, Pampadampari Ranges; HT; ♀; INPC. 

264

265 

annandalei. India (Orissa). 

Prolasioptera annandalei Mani 1934b: 395. India: Orissa, Barkuda I., Chilka L.; HT; ♀; ZSCI. 

anonae. Mozambique. Anona senegalensis (Anonaceae). 

Lasioptera anonae Tavares 1908: 141. Mozambique; ST; ♂,♀; TAVC. 

argentata. Poland. 

Lasioptera argentata Loew 1850a: 39. Poland; T; A; ZMHB?. 

arizonensis. USA (Arizona). Senecio arizonensis (Asteraceae). 

Lasioptera arizonensis Felt 1908b: 325 (as n. sp. also in Felt 1911l: 470 & Felt 1911m: 482). USA: Arizona, Oracle; 

ST; ♂; USNM. 

Lasioptera arizonae Kieffer 1913i: 31, unjust. new name for Lasioptera arizonensis Felt. 

artemisiae. Russia (Europe). Artemisia vulgaris; A. japonica, A. rubripes (Asteraceae). Ref.: Fedotova & Sidorenko 

2005: 459 (redescr.). 

Lasioptera artemisiae Dombrovskaya 1940b: 195. Russia: Penza Prov., Sosnovy Bor For. Res., nr Penza; ST; 

♂,♀,P,L; ZMAS. 

artemisifoliae. Japan (Honshu to Kyushu), Russia (Far East). Artemisia japonica (Asteraceae). 

Lasioptera artemisifoliae Shinji 1939a: 383. Japan: Honshu; T; ♀; lost. 

Lasioptera artemisifolia Mamaeva & Kritskaya 1980: 147, missp. of artemisifoliae. 

arundinis. Widespread Europe. Phragmites australis (Poaceae). 

Lasioptera arundinis Schiner 1854: 175. Austria: Vienna, Prater; ST; ♂,♀; NHMW. 

astericola. Japan (Honshu). Aster scaber (Asteraceae). 

Lasioptera astericola Shinji 1939g: 162. Japan: Honshu, Morioka; ST; A; lost. 

asterspinosae. USA (Texas). Aster spinosus (Asteraceae). 

Lasioptera asterspinosae White 1950: 67. USA: Texas, vic. Austin; ST; ♂,♀,P,L; USNM. 

asystasiae. India (Kerala). Asystasia violacea (Acanthaceae). 

Lasioptera asystasiae Nayar 1945: 71. India: Kerala, Cardamom Hills, Pampadampara; HT; ♂; ETKI. 

aurata. Australia (New South Wales). 

Lasioptera aurata Skuse 1888: 129. Australia: New South Wales, Middle Harbour; HT; ♂; ANIC. 

auricincta. Germany, Poland. Festuca pratensis (Poaceae). 

Lasioptera auricincta Loew 1850a: 39. Poland; ST; A; ZMHB?. 

Lasioptera auricincta Winnertz 1853: 308. Germany; ST; ♂,♀,L; MNHN?. 

azamii. Japan (Honshu). Saussurea affinis (Asteraceae). 

Lasioptera azamii Shinji 1939h: 227. Japan: Honshu; T; ♀; lost. 

Lasioptera azami Shinji 1944: 320, missp. of azamii. 

basiflava. USA (New York). 

Lasioptera basiflava Felt 1908b: 324. USA: New York, Albany; HT; ♀; NYSM. 

berberina. Austria. Berberis vulgaris (Berberidaceae). 

Tipula berberina Schrank 1781: 435. Austria; ST; A,P; lost?. 

berlesiana. Circum-Mediterranean. Olea europaea (Oleaceae); Pistacia lentiscus (Anacardiaceae). Ref.: Sasso & 

Viggiani 2001: 55 (biol., new host). 

Lasioptera berlesiana Paoli 1907: 45. Italy: Tuscany, Maremma; ST; ♀,L; ?. 

Lasioptera brevicornis Melis 1925: 41 (publ'n not seen); also as n. sp. in Melis 1928: 41. Italy: Pisa, San Vicenzo; T; 

A; ?. 

Lasioptera carpophila Del Guercio 1918a: 62. Italy: Abruzzi, Teramo; T; LT; lost. 

Lasioptera kiefferiana Del Guercio 1910: 288. Italy; T; ♀; lost. 

bothriochloae. India (Maharashtra). Bothriochloa pertusa (Poaceae). 

Lasioptera bothriochloae Rao & Sharma 1977a: 148. India: Maharashtra, Aurangabad, Marathwada Univ. campus; 

HT; ♂; RCMU. 

bryoniae. India (Tamil Nadu). Bryonia sp. (Cucurbitaceae). 

Lasioptera bryoniae Schiner 1868: 5. India: Tamil Nadu, Madras; ST; ♂,♀; NHMW?. 

buhri. Germany. Crepis biennis; also spp. of Hieracium, Lapsana, Mycelis, Picris, Tragopogon (Asteraceae), 

Campanula (Campanulaceae), & Melilotus (Fabaceae). 

Lasioptera buhri Möhn 1968b: 125. Germany: Mühlhausen; HT; LT; SMNS.

266 

calamagrostidis. UK, Netherlands, France, Germany, Latvia, Poland, Czechoslovakia, Hungary, Ukraine. 

Calamagrostis epigeios; also other grasses (Poaceae). 

Lasioptera calamagrostidis Rübsaamen 1893: 164. Germany: vic. Berlin; T; ♂; ZMHB. 

Lasioptera graminicola Kieffer 1898f: 58. France: Moselle; T; A; lost?. 

callicarpae. Japan (Honshu, Kyushu). Callicarpa japonica (Verbenaceae). 

Calopedila callicarpae Shinji 1938i: 371. Japan: Honshu, Iwate Pref., Morioka; ST; ♂,♀; lost. 

camelliae. Japan (Honshu to Okinawa). Camellia japonica (Theaceae). 

Lasioptera camelliae Ohno & Yukawa 1984: 429. Japan: Kyushu, Kagoshima City, Shiroyama; HT; ♂; KUIC. 

carbonaria. South Africa. ?Passerina sp. (Thymelaeaceae). 

Lasioptera carbonaria Schiner 1868: 6. South Africa: Cape of Good Hope, vic. Constantia; HT; ♀; NHMW. 

carophila. Widespread Europe, North Africa. Carum carvi; various genera and spp. of Apiaceae (Möhn 1968: 88 ff., 

as L. argentata). 

Lasioptera carophila Löw 1874a: 149. Austria: Wienerwald; ST; ♀,P,L; NHMW. 

Lasioptera thapsiae Kieffer 1898f: 58. Algeria; T; ♀; lost?. 

caryae. USA (New York). 

Lasioptera caryae Felt 1907a: 10. USA: New York, Albany; HT; ♂; NYSM. 

centerensis. USA (New York). Impatiens aurea (Balsaminaceae). 

Lasioptera centerensis Felt 1918a: 170. USA: New York, Karner; HT; LT; NYSM. 

cephalandrae. India (widespread). Coccinia indica (Cucurbitaceae). 

Neolasioptera cephalandrae Mani 1934b: 397. India: Tamil Nadu, Thanjavur; HT; ♀; ZSCI. 

cerasiphera. Germany. Cerasus vulgaris (Rosaceae). 

Lasioptera cerasiphera Stelter 1990: 73. Germany: vic. Kleinmachnow nr Berlin; HT; ♂; SMTD. 

chichindae. India (Uttar Pradesh). Trichosanthes cucumerina (Cucurbitaceae). 

Bimba chichindae Grover 1966a: 249. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

cimicifugae. Russia (Far East). Cimicifuga simplex (Ranunculaceae). 

Lasioptera cimicifugae Kovalev 1967: 92. Russia: Primorskiï Terr., Suputinskiy Res.; HT; ♀; KOVC. 

cinerea. USA (New York). 

Lasioptera cinerea Felt 1907a: 8. USA: New York, Albany; HT; ♂; NYSM. 

clinopodii. Russia (Far East). Clinopodium chinense (Lamiaceae). 

Lasioptera clinopodii Kovalev 1967: 92. Russia: Primorskiï Terr., Popova I., in Petra Velikoe Bay, nr Vladivostok; 

HT; ♀; KOVC. 

collinsonifolia. USA (New York, Maryland). Collinsonia canadensis (Lamiaceae). 

Cecidomyia collinsonifolia Beutenmüller 1908: 74. USA: New York, Garrison; HT; LT; AMNH. 

corni. USA (New York, Massachusetts). Cornus paniculata (Cornaceae). 

Lasioptera corni Felt 1907a: 11. USA: New York, Albany; HT; ♀; NYSM. 

corusca. Australia (New South Wales). 

Lasioptera corusca Skuse 1888: 131. Australia, New South Wales, Elizabeth Bay; ST; ♂; ANIC. 

crataevae. India (Tamil Nadu). Crateva religiosa (Capparidaceae). 

Neolasioptera crataevae Mani 1934b: 399. India: Tamil Nadu, Thanjavur, Grand Anicut Canal Bank; HT; ♀; ZSCI. 

cubitalis. Kenya. 

Lasioptera cubitalis Kieffer 1913h: 88. Kenya: Tiwi, 20 km S Mombasa; HT; ♀; MNHN. 

cylindrigallae. USA (New York, Massachusetts). Solidago sp. (Asteraceae). 

Lasioptera cylindrigallae Felt 1907c: 150. USA: New York, New York, Staten I.; ST; ♂,♀; NYSM. 

dioscoreae. Russia (Far East). Dioscorea polystachya (Dioscoreaceae). 

Lasioptera dioscoreae Kovalev 1967: 94. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♀; KOVC. 

Lasioptera dioscorae Skuhravá 1986: 127, missp. of dioscoreae. 

*dombrovskajae. Russia (Far East). Artemisia rubripes (Asteraceae). 

Lasioptera dombrovskajae Fedotova & Kovalev 2005: 457. Russia: Primorskiï Terr., Sedanka nr Vladivostok; HT; 

♀; ZMAS. 

donacis. France, Italy, Malta, Greece, Bulgaria, Egypt. Arundo donax (Poaceae). 

Lasioptera donacis Coutin & Faivre-Amiot 1981: 196, nomen nudum. 

Lasioptera donacis Coutin 2001b: 105. France: Montpellier; HT; ♀; MNHN.

267 

ephedrae. USA (California, Arizona, New Mexico). Ephedra trifurca (Gnetaceae). 

Lasioptera ephedrae Cockerell 1898: 327. USA: New Mexico, Mesilla Park & Paraje; ST; ♀,L; USNM. 

ephedricola. USA (California, Arizona, New Mexico). Ephedra trifurca; E. californica, E. nevadensis (Gnetaceae). 

Lasioptera ephedricola Cockerell 1902: 184. USA: New Mexico, Mesilla Park; ST; ♂,L; USNM. 

eriochloa. India (Tamil Nadu). Eriochloa polystachya (Poaceae). 

Lasioptera eriochloa Felt 1926i: 244. India: Tamil Nadu, Coimbatore; ST; ♀; NYSM. 

eryngii. Widespread Europe, Algeria. Eryngium campestre; Eryngium spp. (Apiaceae). 

Cecidomyia eryngii Vallot 1829: 114. France; ST; A,L; ?. 

Lasioptera eryngii Dufour 1846b: 124. France: Landes, St. Sever; ST; ♀,P,L; ?. 

Lasioptera eryngii Giraud 1861: 483. Europe; ST; ♂,♀,P,L; MNHN. 

excavata. USA (New York). Crataegus sp. (Rosaceae). 

Lasioptera excavata Felt 1907d: 6. USA: New York, vic. Albany: T; ♀; NYSM. 

falcata. India, Philippines. Undet. sp. of Cucurbitaceae. 

Lasioptera falcata Felt 1919a: 288. Philippines: Luzon, Laguna Prov., Los Baños; HT; ♀; NYSM. 

flavoventris. USA (New York). 

Neolasioptera flavoventris Felt 1908b: 333. USA: New York, Karner; ST; ♂; NYSM. 

fluitans. India (Tamil Nadu). Panicum fluitans (Poaceae). 

Lasioptera fluitans Felt 1917a: 73. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

francoisi. UK, France, Germany, Romania. Achillea millefolium (Asteraceae). 

Perrisia francoisi (also as fancoisi) Kieffer 1902a: 244. France; T; G; lost?. 

Lasioptera niveocincta Kieffer 1904b: 341. France: Bitche; T; ♂; lost?. 

fructuaria. USA (Maine). Vaccinium pennsylvanicum, V. canadense (Ericaceae). 

Lasioptera fructuaria Felt 1916b: 266. USA: Maine, Orono, Cherryfield, Auburn, & Searsport; ST; ♂,♀,P,L,E; 

NYSM. 

*garugai. India (Maharashtra). Garuga pinnata (Burseraceae). 

Lasioptera graugai Bhalerao, Shaikh and Deshpande 2007b: 287. India: Maharashtra, Nanded District, Bhokar; HT; 

♀; Dept. Zoology, N.E.S. Science College, Nanded, Maharashtra. Ref.: Bhalerao, Shaikh and Deshpande 

2007a. 

gibaushi. Japan (Honshu). Aster scaber (Asteraceae). 

Lasioptesa gibaushi Shinji 1939b: 567. Japan: Honshu; T; ♀; lost. 

hamata. USA (New York). ?Desmodium sp. (Fabaceae). 

Lasioptera hamata Felt 1907a: 11. USA: New York, Nassau; HT; ♀; NYSM. 

Choristoneura hamata Felt 1907c: 155. USA: New York, Albany; ST; ♂,♀; NYSM. 

helvipes. Australia (New South Wales). 

Lasioptera helvipes Skuse 1888: 132. Australia: New South Wales, Sydney; ST; ♂; ANIC. 

howardi. USA (California). Quercus sp. (Fagaceae). 

Lasioptera howardi Felt 1921g: 94. USA: California, Contra Costa Co., Mt Diablo, Pine Canyon; HT; ♀; USNM. 

humulicaulis. USA (New York). Humulus sp. (Cannabaceae). 

Lasioptera humulicaulis Felt 1907c: 151. USA: New York, Rye; ST; ♂,♀; NYSM. 

hungarica. Widespread Europe; immigr: USA (eastern). Phragmites australis (Poaceae). 

Lasioptera hungarica Möhn 1968b: 136. Hungary: Bátorliget; HT; LT; SMNS. 

Thomasiella massa Erdös 1957: 171 (publ'n not seen). A nomen nudum (Möhn 1968b: 138). 

Lasioptera erdosi Möhn 1968b: 138. Hungary: Gárdony; HT; LT; SMNS. 

hygrophila. Russia (Far East). Senecio cannabifolius (Asteraceae). 

Lasioptera hygrophila Kovalev 1967: 93. Russia: Primorskiï Terr., Suputinskiy Res.; HT; ♀; KOVC. 


Lasioptera indica Rao 1952a: 50. India: Uttar Pradesh, Agra, Hewett Park; HT; ♀; ZSCI. 

Neoprotoplonyx indica Rao 1952b: 33, jun. sec. hom. indica Rao 1952a. India: Uttar Pradesh, Agra, St. John's 

College; HT; ♀; ZSCI. This species is tentatively synonymized here to avoid coining a possibly unnecessary 

new name for it. 

javanica. Indonesia (Java). Melothria perpusilla (Cucurbitaceae). 

Lasioptera javanica Kieffer & Docters van Leeuwen-Reijnvaan 1910: 123. Indonesia: Java, Salatiga & elsewhere;

268 

ST; ♂,♀,L; lost?. 

kadaladi. India (Kerala). 

Neoprotaplonyx kadaladi Rao 1956a: 70. India: Kerala, Kadaladi; HT; ♀; ZSCI. 

kallstroemia. USA (Texas), Mexico, Guatemala, Nicaragua. Kallstroemia intermedia (Zygophyllaceae); Solanum spp. 

(Solanaceae). 

Lasioptera kallstroemia Felt 1935a: 6. USA: Texas, Austin; ST; ♀; NYSM. 

koreana. Russia (Far East). Sambucus coreana (Caprifoliaceae). 

Lasioptera koreana Kovalev 1967: 89. Russia: Primorskiï Terr., Khasan, Golubinnyi Rocks; HT; ♀; KOVC. 

kosarzewskella. Czechoslovakia, Ukraine. Rhamnus catharticus (Rhamnaceae). 

Lasioptera kosarzewskella Kozarzhevskaya 1957: 1502 (attributed to Marikovskij but such description as appears is 

by Kozarzhevskaya). Ukraine: Voroshilovgrad Prov., Derkul' For. Stn; ST; ♂,♀; ?. 

Lasioptera kasarzewskella Marikovskij 1958a: 142. Ukraine: Voroshilovgrad Prov., Derkul' For. Stn; ST; ♂,♀; ?. 

lactucae. USA (New York, New Hampshire, Massachusetts, Maryland). Lactuca sp. (Asteraceae). 

Lasioptera lactucae Felt 1907c: 151. USA: Massachusetts, Worcester; ST; ♂,♀; NYSM. 

lespedezae. Japan (Honshu to Kyushu), Republic of Korea, Russia (Far East). Lespedeza bicolor; L. cyrtobotrya 

(Fabaceae). 

Lasioptera lespedezae Shinji 1939c: 583. Nomen nudum. 

Lasioptera lespedezae Shinji 1939e: 100. Japan: Honshu, Morioka; T; A; lost. 

lignicola. South Africa. Muraltia sp. (Polygalaceae). 

Lasioptera lignicola Schiner 1868: 6. South Africa: Cape of Good Hope; T; A; NHMW?. 

longipes. France. 

Lasioptera longipes Kieffer 1904b: 340. France: Bitche; ST; ♂,♀; lost?. 

longispatha. India (West Bengal). Camellia drupifera (Theaceae). 

Lasioptera longispatha Kieffer 1909a: 157. India: West Bengal, Kurseong; T; LT; lost?. 

lonicericola. Russia (Far East). Lonicera sp. (Caprifoliaceae). 

Lasioptera lonicericola Kovalev 1967: 95. Russia: Primorskiï Terr., Khualaza, southern Sikhote-Alin' Mts; HT; ♀; 

KOVC. 

loyolai. India (Tamil Nadu). Panicum sp. (Poaceae). 

Lasioptera loyolai Mani 1986: 13. India: Tamil Nadu, Madras; HT; ♀; ZSCI. 

mangiflorae. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Meunieriella mangiflorae Grover 1968a: 18. India: Uttar Pradesh, Lucknow; HT; ♂; PGCA. 

manilensis. Java, Philippines. Leea manillensis, L. sambucina (Vitaceae). 

Lasioptera manilensis Felt 1918c: 288. Philippines: Luzon, Laguna Prov., Los Baños & Mt Makiling; ST; ♂,♀; 

NYSM. 

melampyri. Germany. Melampyrum arvense (Scrophulariaceae). 

Lasioptera melampyri Möhn 1968b: 130. Germany: Mühlhausen, Eigenrieden; HT; LT; SMNS. 

miscella. Australia (New South Wales). 

Lasioptera miscella Skuse 1888: 131. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC. 

moliniae. Germany. Molinia caerulea (Poaceae). 

Lasioptera moliniae Möhn 1968b: 135. Germany: Plötzensee, nr Berlin; HT; LT; SMNS. 

nenuae. India (Uttar Pradesh). Luffa acutangula (Cucurbitaceae). 

Bimba nenuae Grover 1966a: 251. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

nigrocincta. Algeria. Quercus sp. (Fagaceae). 

Lasioptera nigrocincta Kieffer 1904b: 339. Algeria: Philippeville; T; ♂; lost?. 

nodosae. Australia (New South Wales). Melaleuca nodosa (Myrtaceae). 

Lasioptera nodosae Skuse 1888: 130. Australia: New South Wales, Homebush; HT; ♀; ANIC. 


Lasioptera orientalis Rao 1960a: 183. India: Uttar Pradesh, Agra; HT; ♀; ZSCI. 

paederiae. Japan (Honshu, Kyushu). Paederia chinensis; P. scandens (Rubiaceae). 

Dasyneura paederiae Shinji 1944: 349. Japan: Honshu; T; A; lost. 

paniculi. Philippines. Panicum carinatum (Poaceae). 

Lasioptera paniculi Felt 1920d: 232. Philippines: Luzon, Laguna Prov., Los Baños; ST; ♂,♀; NYSM.

269 

populnea. Widespread Europe. Populus alba, P. canescens (Salicaceae). 

Lasioptera populnea Wachtl 1883: 477. Austria: vic. Vienna; ST; ♂,♀,L; NHMW?. 

psederae. USA (Ohio). Parthenocissus quinquefolia (Vitaceae). 

Lasioptera psederae Felt 1934a: 77. USA: Ohio, Columbus; ST; ♀,L; NYSM. 

puerariae. Japan (Honshu), Korea. Pueraria thunbergiana (Fabaceae). 

Calopedila puerariae Shinji 1938k: 241. Japan: Honshu; ST; ♂,♀; lost. 

Lasioptera puerarire Shinji 1944: 332, missp. of puerariae. 

querciflorae. USA (Oregon, Arizona, New York, Maryland). Quercus sp. (Fagaceae). 

Lasioptera querciflorae Felt 1908b: 325. USA: Arizona, Fort Grant; HT; ♀; USNM. 

Lasioptera quercifloriae Pritchard 1953a: 137, missp. of querciflorae. 

querciperda. USA (New York, Massachusetts, Rhode Island, Virginia). Quercus sp. (Fagaceae). 

Lasioptera querciperda Felt 1908b: 324. USA: Virginia; ST; ♀; USNM. 

quercirami. USA (Utah). Quercus sp. (Fagaceae). 

Lasioptera quercirami Felt 1926g: 80. USA: UT, Logan; HT; ♂; NYSM. 

rubi. Widespread Europe, Russia (Far East), Japan (Honshu to Kyushu). Rubus spp. (Rosaceae). 

Tipula rubi Schrank 1803: 87. France; T; LT; lost?. 

Cecidomyia albipennis Meigen 1804: 40. Germany; T; A; MNHN. 

Lasioptera fusca Meigen 1818: 89. Austria; T; ♀; lost. 

Lasioptera juniperi Meigen 1818: 89. Austria; T; ♀; lost. 

Lasioptera picta Meigen 1818: 89. Germany; T; ♀; lost. 

Diomyza fuliginosa Stephens 1829a: 55 (publ'n not seen); also in Stephens 1929b: 240. Nomen nudum. 

Lasioptera argyrosticta Meigen 1830: 266. Germany; T; ♀; MNHN. 

Lasioptera pulchra Meigen 1830: 267. Germany: T; ♀; lost. 

Lasioptera annulata Costa 1844: 83. Italy: Marana di Castiglione, nr Foggia; HT; ♀; ?. 

Lasioptera rubi Heeger 1850: 39. Nomen nudum. 

Lasioptera rubi Heeger 1851: 203. Austria; ST; ♀,P,L; ?. 

Lasioptera ichigo Shinji 1939b: 561. Japan: Honshu; ST; ♂,♀; lost. 

Neolasioptera rubicola Monzen 1955a: 39. Japan: Iwate Pref., Shimohei; ST; ♂,♀; KUEC. 

ruebsaameni. Germany. Sonchus sp. (Asteraceae). 

Lasioptera ruebsaameni Möhn 1968b: 147. Germany: Rhineland; HT; LT; SMNS. 

rufa. France. 

Lasioptera rufa Kieffer 1904b: 338. France: Bitche; T; ♂; lost?. 

salviae. South Africa. Salvia sp. (Lamiaceae). 

Lasioptera salviae Schiner 1868: 7. South Africa: Cape of Good Hope; HT; ♀; NHMW?. 

serotina. USA (New York). 

Lasioptera serotina Felt 1908b: 326. USA: New York, Albany; HT; ♀; NYSM. 

solani. USA (Maryland, Virginia, South Carolina). Solanum carolinense (Solanaceae). 

Choristoneura solani Felt 1907c: 164. USA: District of Columbia; T; ♀; NYSM. 

solidaginis. USA (widespread eastern). Solidago canadensis (Asteraceae). 

Lasioptera solidaginis Osten Sacken 1863: 370. USA; ST; ♂,♀; MCZC. 

Lasioptera dorsimaculata Felt 1908b: 325. USA: New York, Karner; HT; ♀; NYSM. 

soongarica. Kazakhstan. Artemisia proceraeformis (Asteraceae). 

Lasioptera soongarica Fedotova 1991a: 27. Kazakhstan: Dzungarskiï Alatau, 30 km SE Uch-Aral; HT; ♀; ZMAS. 

sorghivora. Nigeria. Sorghum vulgare (Poaceae). 

Thomasiella sorghivora Harris 1960: 661. Nigeria: Zaria; HT; ♂; BMNH. 

spiraeafolia. Canada (British Columbia), USA (Washington, New York, Maine, Massachusetts, Maryland). Spiraea 

salicifolia; Spiraea spp. (Rosaceae). 

Lasioptera spiraeafolia Felt 1909b: 287. USA: Massachusetts, Magnolia; HT; ♂; NYSM. 

stelteri. Germany. Heracleum sphondylium (Apiaceae). 

Lasioptera stelteri Möhn 1968b: 146. Germany: Taunus, Dauborn; HT; LT; SMNS. 

tarbagataica. Kazakhstan. Artemisia dracunculus (Asteraceae). 

Lasioptera tarbagataica Fedotova 1991a: 25. Kazakhstan: Tarbagataï Mtn Range, 7 km N Plagodatnogo,

270 

Kel'dymurat R. floodplain; HT; ♀; ZMAS. 

taroiae. India (Uttar Pradesh). Luffa cylindrica (Cucurbitaceae). 

Bimba taroiae Grover 1966a: 250. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

terminaliae. Mozambique. Terminalia sericea (Combretaceae). 

Lasioptera terminaliae Tavares 1908: 153. Mozambique; ST; ♂,♀,P; TAVC. 

textor. India (West Bengal). Polygonum molle (Polygonaceae). 

Lasioptera textor Kieffer 1905a: 155. India: West Bengal, vic. Kurseong; ST; A,P,L; lost?. 

thuringica. Germany. Pimpinella major (Apiaceae). 

Lasioptera thuringica Möhn 1968b: 147. Germany: Thüringen, Mühlhausen; HT; LT; SMNS. 

tibialis. USA (California). Keckiella antirrhinoides (Scrophulariaceae); Atriplex canescens (Chenopodiaceae). 

Lasioptera tibialis Felt 1914f: 119. USA: California, nr La Jolla; L (Gagné & Hawkins 1983: 383); ♀; NYSM. 

tiliarum. Germany, Czechoslovakia, Russia (Europe). Tilia sp. (Tiliaceae). 

Lasioptera tiliarum Mamaeva 1964a: 208. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♀; HYNM. 

tomentosae. India (Uttar Pradesh). Setaria tomentosa (Poaceae). 

Thomasiella tomentosae Grover 1967d: 152. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

toombii. India (Uttar Pradesh). Coccinia indica (Cucurbitaceae). 

Bimba toombii Grover 1962d: 355. India: Uttar Pradesh, Allahabad; ST; ♂,♀,P,L,E; PGCA. 

trapezospatha. Eritrea. Asclepias sp. (Asclepiadaceae). 

Monilipennella trapezospatha Del Guercio 1918a: 147. Eritrea; ST; A,P,L; lost. 

trilobata. India (West Bengal). Schima walchii (Theaceae) or Echinocarpus dasycarpus (Elaeocarpaceae). 

Lasioptera trilobata Kieffer 1909a: 157. India: West Bengal, Kurseong; T; LT; lost?. 

tuberosa. Tanzania. 

Lasioptera tuberosa Kieffer 1913h: 88. Tanzania: Neu-Moschi, nr Rau R., lower slopes of Kilimanjaro; HT; ♀; 

MNHN. 

turcica. Turkey. Echinophora anatolica (Apiaceae). 

Lasioptera turcica Möhn 1968b: 113. Turkey: Paphlagonia, Wilajet Kastambuli; HT; LT; SMNS. 

ukogi. Japan (Honshu). Acanthopanax sp. (Apiaceae). 

Lasioptera ukogi Shinji 1940: 34. Japan: Honshu; ST; ♂; lost. 

Lasioptera ukogii Shinji 1944: 25, missp. of ukogi. 

umbelliferarum. Italy, Russia (Europe), Georgia, Kurdistan, Iran, Palestine. Seseli sp.; Hippomarathrum spp. 

(Apiaceae). 

Lasioptera umbelliferarum Kieffer 1909b: 28. Russia: Dagestan, Makhachkala; ST; LT; lost?. 

uncinata. Australia (QLD). Melaleuca dealbata; Melaleuca spp. (Myrtaceae). 

Lasioptera uncinata Gagné 1997d: 330. Australia: QLD, 3 km N Cardwell; HT; ♀; ANIC. 

ussurica. Russia (Far East). Plectrantus incisus (Lamiaceae). 

Lasioptera ussurica Mamaev & Kritskaya 1980: 410. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; HYNM. 

variipalpis. India (West Bengal). 

Neolasioptera variipalpis Mani 1937: 282. India: West Bengal, Calcutta, Portland Park; ST; ♀; ZSCI. 

vastatrix. Australia (New South Wales). Undet. sp. of Poaceae. 

Lasioptera vastatrix Skuse 1888: 128. Australia: New South Wales, Parkes Distr.; ST; ♂,♀,P,L; ANIC. 

viburni. USA (New York). 

Lasioptera viburni Felt 1907a: 8. USA: New York, Albany; HT; ♂; NYSM. 

vitis. USA (New York, Massachusetts, Maryland). Vitis sp.; Vitis spp. (Vitaceae). 

Lasioptera vitis Osten Sacken 1862: 201. USA: DC; ST; A,L; lost. 

wildi. Australia (QLD). 

Lasioptera wildi Skuse 1890: 387. Australia: QLD, Brisbane, Botanical Gardens; ST; ♂,♀; ANIC. 

yadokariae. Japan (Kyushu). Neolitsea sericea (Lauraceae); Machilus japonica (Lauraceae), Euonymus japonicus 

(Celastraceae). 

Lasioptera yadokariae Yukawa & Haitsuka 1994: 712. Japan: Kyushu, Kagoshima City, Ishiki; HT; ♂; KUIC. 

ziziae. USA (Illinois). Zizia sp. (Asteraceae). 

Lasioptera ziziae Felt 1908b: 327. USA: Illinois, Evanston; ST; ♂,♀; NYSM. 

Lasioptera zigiae Kieffer 1913i: 33, missp. of ziziae.

LATHYROMYZA 

Lathyromyza Rübsaamen 1916b: 491. Type species, Cecidomyia schlechtendali Kieffer (orig. des.). 

Palearctic; 4 spp., 3 on Vicia or Lathyrus; Lasiopteridi, Dasineurini. 

abruptis. Kazakhstan. Vicia cracca; V. tenuifolia (Fabaceae). 

Lathyromyza abruptis Fedotova 1991d: 24. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, nr Alma-Ata; HT; ♂; 

ZMAS. 

florum. Denmark, Germany, Czechoslovakia. Lathyrus sylvestris; L. tuberosus (Fabaceae). 

Lathyromyza florum Rübsaamen 1916b: 521. Germany; ST; ♂,♀,L; ZMHB. 

*recedens. Russia (Far East). 

Ledomyia recedens Fedotova & Sidorenko 2005d: 10. Russia: Primorskiï Terr., Vladivostok; HT; ♀; ZMAS. 

schlechtendali. Widespread Europe, Russia (Western Siberia), Kazakhstan. Lathyrus linifolia; Vicia cracca 

(Fabaceae). 

Cecidomyia schlechtendali Kieffer 1886: 328. France: betw. Bitche & Reyersweiler; ST; ♂,♀,L; lost?. 

LAUTHIA 

Lauthia Kieffer 1912h: 2. Type species, Ledomyia divisa Kieffer (orig. des.). 

Cryptolauthia Kieffer 1912h: 2. Type species, Ledomyia connata Kieffer (orig. des.). 

Microperrisia Kieffer 1912h: 2. Type species, Perrisia brachypsectra Kieffer (orig. des.). 

Phaenolauthia Kieffer 1912h: 2. Type species, Ledomyia obscuripennis Kieffer (orig. des.). 

Properrisia Kieffer 1912h: 2. Type species, Ledomyia aurofulgens Kieffer (orig. des.). 

Sphaerolauthia Kieffer 1912h: 2. Type species, Ledomyia monilicornis Kieffer (orig. des.). 

Nanolauthia Kieffer 1913i: 110. Type species, Ledomyia mahensis Kieffer (mon.). 

Cryptolanthia Kieffer 1919: 192, missp. of Cryptolauthia. 

Phaenolanthia Kieffer 1919: 192, missp. of Phaenolauthia. 

Cosmopolitan; 30 spp.; Lasiopteridi, Ledomyiini. Most species were caught in flight but at least two, L. acariphaga 

and L. acerina, were associated with eriophyid galls and L. cardui was reared from old galls of a tephritid fly. Others 

were reared from freshly cut or partly decayed wood. Some of these species were originally in Ledomyia, which is now 

limited to its strict sense. Lauthia shows an amazing variety in male and female genitalia and some of the synonymous 

genera will doubtless resurface to separate use someday. Ref.: Gagné 2004b (syn. note). 

acariphaga. Kazakhstan. Eriophyidae galls on Acroptilon picris (Asteraceae). 

Lasiopteryx acariphaga Marikovskij 1968: 110. Kazakhstan: vic. Alma-Ata & Sarkand; ST; ♂,♀; MARC. 

acerina. France, Switzerland, Austria. Acer pseudoplatanus (Aceraceae). 

Cecidomyia acerina Giraud 1863: 1304. France: vic. La Grande Chartreuse Monastery, & Switzerland: slope of 

Salève mountain, nr Geneva; ST; ♂,♀; MNHN. 

ampelophila. USA (New York). 

Dasyneura ampelophila Felt 1908b: 343. USA: New York, Albany; HT; ♂; NYSM. 

arizonensis. USA (Arizona). 

Lasiopteryx arizonensis Felt 1915d: 195. USA: Arizona, Prescott; ST; ♂,♀; NYSM. 

aurofulgens. Seychelles Is. 

Ledomyia aurofulgens Kieffer 1911: 316. Seychelles Is: Mahé, Cascade Estate; ST; ♂,♀; BMNH. 

brachypsectra. France. 

Perrisia brachypsectra Kieffer 1904c: 379. France: vic. Bitche; T; ♂; lost?. 

cardui. UK, France. Cirsium arvense (Asteraceae). 

Ledomyia cardui Kieffer 1904b: 346. France: vic. Paris; ST; ♂,♀; lost?. 

carpini. USA (New York). 

271

272 

Asphondylia carpini Felt 1907a: 23. USA: New York, Albany; HT; ♀; NYSM. 

connata. France. 

Ledomyia connata Kieffer 1904c: 370. France: vic. Bitche; ST; ♀; lost?. 

coryli. USA (New York). Corylus sp. (Betulaceae). 

Dasyneura coryli Felt 1907d: 11. USA: New York, West Nyack; ST; ♂,♀; NYSM. 

crispata. USA (Massachusetts). 

Lasiopteryx crispata Felt 1914d: 111. USA: Massachusetts, Magnolia; ST; ♀,L; NYSM. 

divisa. France. 

Lauthia divisa Kieffer 1904c: 372. France: vic. Bitche; ST; ♂,♀; NYSM?. 

ilicis. Algeria. Ilex aquifolium (Aquifoliaceae). 

Phaenolauthia ilicis Kieffer 1919: 193. Algeria: Dra-Inguel, Haïzer - (Djurdjura); T; ♂; lost?. 

maculosa. USA (New York). 

Dasyneura maculosa Felt 1908b: 341. USA: New York, Nassau; HT; ♀; NYSM. 


Ledomyia mahensis Kieffer 1911: 317. Seychelles Is: Mahé, Cascade Estate; HT; ♂; BMNH. 

manihot. St. Vincent. Manihot utilissima (Euphorbiaceae). 

Lasiopteryx manihot Felt 1912c: 144. St. Vincent; ST; ♂,♀; NYSM. 

minuta. Sweden. 

Cecidomyza minuta Zetterstedt 1838: 824. Sweden: Lycksele; HT; ♂; MZLU?. 

monilicornis. Seychelles Is. 

Ledomyia monilicornis Kieffer 1911: 318. Seychelles Is: Mahé, Cascade Estate; HT; ♂; BMNH. 

obscuripennis. France, Russia (Europe). 

Ledomyia obscuripennis Kieffer 1904c: 371. France: vic. Bitche; T; ♀; lost?. 

parma. USA (New York). 

Janetiella parma Felt 1914c: 129. USA: New York, Adirondack Mts, Woodworth's L.; HT; ♂; NYSM. 

primigenia. Russia (Far East). 

Ledomyia primigenia Mamaev 1994: 31. Russia: Kamchatka, Kozyrevsk; HT; ♂; ZMMU. 

pudorosa. USA (New York). 

Dasyneura pudorosa Felt 1908b: 344. USA: New York, Nassau; HT; ♂; NYSM. 

schwarzi. Panama. 

Lasiopteryx schwarzi Felt 1911k: 191. Panama: Canal Zone, Paraíso; ST; ♂,♀; USNM. 

setosa. USA (New York). 

Dasyneura setosa Felt 1907a: 19. USA: New York, Nassau; HT; ♂; NYSM. 

simulator. USA (New York). 

Dasyneura simulator Felt 1908b: 344. USA: New York, Nassau; HT; ♂; NYSM. 

spilota. Ukraine, Russia (Europe, ES). 

Lauthia spilota Mamaev 1967: 880. Russia: Yaroslavl Prov., Volkovo; HT; ♂; HYNM. 

spinigerella. Ukraine, Russia (Europe). 

Lauthia spinigerella Mamaev 1967: 880. Russia: Yaroslavl Prov., Volkovo; HT; ♂; HYNM. 

unguicula. USA (New York). 

Dasyneura unguicula Felt 1908b: 344. USA: New York, Nassau; HT; ♂; NYSM. 

volucris. Russia (Far East). 

Ledomyia volucris Mamaev 1972b: 113. Russia: Maritime Terr., Ussuriyskiï Distr., Suputinskiy Res.; HT; ♂; 

HYNM. 

yuccae. USA (Arizona). Yucca sp. (Liliaceae). 

Dasyneura yuccae Felt 1908b: 343. USA: Arizona; HT; ♂; USNM. 

*LAXADIPLOSIS 

Laxadiplosis Fedotova & Sidorenko 2009b: 15. Type species, latebra Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Mycodiplosini, and evidently close to Coquillettomyia.

*latebra. Russia (Far East). 

Laxadiplosis latebra Fedotova & Sidorenko 2009b: 16. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


LEDOMYIA 

Lepidomyia Kieffer 1894i: 201, preocc. Loew 1864. Type species, lugens Kieffer (mon.). 

Ledomyia Kieffer 1895o: cccxx, new name for Lepidomyia Kieffer. 

Neuromyia Felt 1911c: 44. Type species, Arnoldia minor Felt (orig. des.). 

Cosmopolitan and in Eocene amber (undescr.); 26 spp.; Lasiopteridi, Ledomyiini. Most of the included species have 

been reared from fresh-cut logs. Ledomyia collarata was reared from galls on a fungus (Larew et al. 1987); its adults 

are unknown and the species is only tentatively placed in this genus. Refs: Gagné 1985a (rev.), Rock & Jackson 1985 

(biol.). 

*adunca. Russia (Far East). 

Ledomyia adunca Fedotova & Sidorenko 2007a: 5. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya Bay; HT; 

♂; ZMAS. 

aestiva. Russia (Europe). 

Lasiopteryx aestiva Mamaev 1967: 882. Russia: Moscow Prov., Danki; HT; ♂; HYNM. 

*alterna. Russia (Far East). 

Ledomyia alterna Fedotova & Sidorenko 2007a: 14. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; 

♂; ZMAS. 

alternata. Russia (Europe). 

Lasiopteryx alternata Mamaev 1967: 882. Russia: Moscow Prov., Abramtsevo; HT; ♂; ZMMU. 

brevicauda. USA (New York). 

Dasineura brevicauda Felt 1908b: 343. USA: New York, Nassau; ST; ♀; NYSM. 

*caespitosa. Russia (Far East). 

Ledomyia caespitosa Fedotova & Sidorenko 2007a: 8. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; 

♂; ZMAS. 

collarata. Venezuela. Xylaria enterogena (Xylariaceae; Ascomycetes). 

Ledomyia collarata Gagné 1987a: 502. Venezuela: Amazonian Federal Terr., Cerro de la Neblina; HT; LT; IZAV. 

*conspecta. Russia (Far East). 

Ledomyia conspecta Fedotova & Sidorenko 2007a: 13. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


* † dextra. Rovno amber (Ukraine). 

Ledomyia dextra Fedotova & Perkovsky 2008a: 37. Rovno amber, Ukraine, Durovitsa (late Eocene); HT; ♀; UASK. 

emilyae. USA (Ohio, Massachusetts, Maryland). 

Ledomyia emilyae Gagné 1985a: 129. USA: Ohio, Akron; HT; ♀; USNM. 

*eminens. Russia (Far East). 

Ledomyia eminens Fedotova & Sidorenko 2005d: 8. Russia: Primorskiï Terr., Vladivostok; HT; ♀; ZMAS. 

flavotibialis. USA (New York). 

Dasyneura flavotibialis Felt 1907c: 157. USA: New York, Nassau; ST; ♂,♀; NYSM. 

lepida. Russia (Europe). 

Lasiopteryx lepida Mamaev 1967: 882. Russia: Moscow Prov., Danki; HT; ♂; HYNM. 

*lignosa. Russia (Far East). 

Ledomyia lignosa Fedotova & Sidorenko 2007a: 6. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskay Pad'; HT; ♂; 

ZMAS. 

lineata. France. 

Ledomyia lineata Kieffer 1904c: 369. France: vic. Bitche; T; ♀; lost?. 

lugens. UK, France. 

273

274 

Lepidomyia lugens Kieffer 1894: 211. France: Bitche; HT; ♀; NYSM?. 

minor. USA (New York). 

Arnoldia minor Felt 1907d: 9. USA: New York, Nassau; HT; ♂; NYSM. 

mira. USA (Ohio). 

Ledomyia mira Gagné 1985a: 130. USA: Ohio, Canal Fulton; HT; ♀; USNM. 

*necessaria. Russia (Far East). 

Ledomyia necessaria Fedotova & Sidorenko 2007a: 15. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*obturata. Russia (Far East). 

Ledomyia obturata Fedotova & Sidorenko 2007a: 17. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; HT; 

♂; ZMAS. 

parva. USA (Illinois, Ohio). 

Ledomyia parva Gagné 1985a: 131. USA: Ohio, Canal Fulton; HT; ♀; USNM. 


Dasyneura phosphila Felt 1907a: 18. USA: New York, Nassau; HT; ♂; NYSM. 

Dasyneura photophila Felt 1907c: 114, unjust. emend. of phosphila. 

ramicola. Algeria. 

Cryptolanthia ramicola Kieffer 1919: 192. Algeria: forest of Farghen, marshes of Mazafran; ST; ♂,♀; lost?. 

*rustica. Russia (Far East). 

Ledomyia rustica Fedotova & Sidorenko 2007a: 10. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskay Pad'; HT; ♂; 

ZMAS. 

styloptera. Seychelles Is. 

Ledomyia styloptera Kieffer 1911: 317. Seychelles Is: Mahé, Cascade Estate; ST; ♀; BMNH. 

*vaga. Russia (Far East). 

Ledomyia vaga Fedotova & Sidorenko 2007a: 10. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; HT; ♂; 

ZMAS. 

LEDOMYIELLA 

Ledomyiella Meunier 1904: 44. Ledomyiella succini Meunier (subs. des. Felt 1911: 38). 

Lidomyiella Mani 1946: 194, missp. of Ledomyiella. 

Baltic amber; 5 spp.; Lasiopteridi, Ledomyiini. 

† crassipes. Upper Eocene; Baltic amber. 

Ledomyiella crassipes Meunier 1904: 47. Baltic amber; ST; ♀; lost. 

† eocenica. Upper Eocene; Baltic amber. 

Ledomyiella eocenica Meunier 1904: 43. Baltic amber; HT; ♀; lost. 

† pygmaea. Upper Eocene; Baltic amber. 

Ledomyiella pygmaea Meunier 1904: 46. Baltic amber; ST; ♂,♀; lost. 

† rotundata. Upper Eocene; Baltic amber. 

Ledomyiella rotundata Meunier 1904: 45. Baltic amber; T; ♀; lost. 

† succini. Upper Eocene; Baltic amber. 

Ledomyiella succini (also as succinea) Meunier 1904: 45. Baltic amber; ST; ♂,♀; lost. 

LEPIDOBREMIA 

Lepidobremia Kieffer 1912h: 1. Type species, Bremia mahensis Kieffer (orig. des.). 


mahensis. Seychelles Is.

Bremia mahensis Kieffer 1911: 326. Seychelles Is: Mahé, I. Anonyme; HT; ♂; lost?. 

LEPIDODIPLOSIS 

Lepidodiplosis Kieffer 1911: 319. Type species, mahensis Kieffer (sub. des. Kieffer 1913i: 227). 


calopus. Seychelles Is. 

Lepidodiplosis calopus Kieffer 1911: 323. Seychelles Is: Mahé, Cascade Estate; HT; ♀; BMNH?. 


Lepidodiplosis mahensis Kieffer 1911: 320. Seychelles Is: Mahé, I. Anonyme; ST; ♂,♀; BMNH. 

seychellensis. Seychelles Is. 

Lepidodiplosis seychellensis Kieffer 1911: 320. Seychelles Is: Mahé, Cascade Estate; HT; ♀; BMNH. 

LESTODIPLOSIS 

Leptodiplosis Kieffer 1894a: xxviii. Type species, Lestodiplosis septemguttata Kieffer (subs. des. Kieffer 1895a: 

cclxxx). Replaced by Lestodiplosis as valid original spelling (ICZN 1958: 291). 

Lestodiplosis Kieffer 1894b: 84. Type species, Cecidomyia septemguttata Kieffer (subs. des. Kieffer 1895a: cclxxx). 

Coprodiplosis Kieffer 1894b: 84. Type species, Cecidomyia polypori Loew (subs. des. Kieffer 1895a: cclxxx). 

Hemidiplosis Kieffer 1894j: 9 (publ'n not seen). Type species, Lestodiplosis rosea Kieffer (mon.). 

Adelgimyza Del Guercio 1918a: 231. Type species, strobilobii Del Guercio (orig. des. as n.g, n.sp.). 

Cecidomyella Del Guercio 1918a: 247. Type species, aulacaspidis Del Guercio (orig. des. as n. g., n. sp.). 

Moreschiella Del Guerico 1918a: 157. Type species, moricola Del Guerico (mon.) = moricolata Gagné. 

Theatodiplosis Tavares 1922c: 148. Type species, quercina Tavares (orig. des.). 

Cecidomyela Felt 1925c: 163, missp. of Cecidomyella. 

Cecidomyiella Mani 1934: 428, missp. of Cecidomyella. 

Chiliodiplosis Möhn 1955d: 415. Type species, vasta Möhn (orig. des.). 

Phonodiplosis Möhn 1955d: 420. Type species, casta Möhn (orig. des.). 

Coplodiplosis Shinji 1944: 245, missp. of Coprodiplosis. 

Cosmopolitan and copal fossil; 182 spp.; Cecidomyiidi, Lestodiplosini. Larvae are predators of other arthropods. This 

genus contains many apparent synonyms that only a proper revision will root out. When the arthropod host is unknown, 

the niche or plant on which larvae were found is given. Refs: Baylac 1985, 1986, 1987 (reviews, illus), Cole & Harris 

2002 (illus.). 

acanthoidis. UK. Dasineura sp. (Diptera: Cecidomyiidae). 

Lestodiplosis acanthoidis Barnes 1928b: 69, fig.1C. UK: Northern Ireland, Cos. Armagh & Tyrone, & England, 

Kent; HT; ♂; BMNH. 

acerina. USA (New York). 

Cecidomyia acerina Felt 1907a: 40. USA: New York, Lake Clear; HT; ♂; NYSM. 

achilleae. UK. Macrolabis achilleae (Diptera: Cecidomyiidae). 

Lestodiplosis achilleae Barnes 1928b: 71, fig. 2F. UK: England, Kent; HT; ♂; BMNH. 

aestiva. Spain. Ametrodiplosis thalictricola (Diptera: Cecidomyiidae). 

Ametrodiplosis aestiva, as var. of thalictricola, Tavares 1916b: 93. Spain: Gallicia, Tuda; ST; ♀; TAVC. 

affinis. UK. Dasineura affinis (Diptera: Cecidomyiidae). 

Lestodiplosis affinis Barnes 1928f: 146. UK: Kent; HT; ♀; BMNH. 

alternans. France, Czechoslovakia. Winnertzia sp. (Diptera: Cecidomyiidae). 

Lestodiplosis alternans Kieffer 1898f: 62. France: Moselle; T; ♂; lost?. 

alvei. UK. Acarina. 

Lestodiplosis alvei Barnes 1934: 594. UK: Sussex, Horsham, Selehurst; ST; ♂,♀; BMNH. 

275

276 

anthemidis. Poland. 

Cecidomyia anthemidis Loew 1850a: 32. Poland; ST; ♂,♀; ZMHB?. 

aonidiellae. South Africa, Italy. Aonidiella aurantii (Hemiptera: Diaspididae). 

Lestodiplosis aonidiellae Harris 1968: 452. South Africa: Letaba; HT; ♂; BMNH. Ref.: Siscaro et al. 1999 (biol.). 

apocyniflorae. USA (New York). In flower buds Apocynum androsaemifolium (Apocynaceae). 

Lestodiplosis apocyniflorae Felt 1908b: 409. USA: New York, Nassau; HT; ♂; NYSM. 

aprimiki. UK. Aprionus miki (Diptera: Cecidomyiidae). 

Lestodiplosis aprimiki Barnes 1928d: 143. UK: Kent; HT; ♂; BMNH. 

arcuata. Germany. On rotten wood. 

Cecidomyia arcuata Winnertz 1853: 256. Germany; ST; ♂,♀; lost. 

asclepiae. USA (New York). Rolled leaves of Asclepias cornuta (Asclepidaceae). 

Lestodiplosis asclepiae Felt 1908b: 409. USA: NY: Albany & Bath; ST; ♂; NYSM. 

asphodeli. Italy. From inflorescence Asphodelus sp. (Liliaceae). 

Lestodiplosis asphodeli Barnes 1935b: 73. Italy: Sardinia, Cagliari; ST; ♂,♀; BMNH. 

asteris. USA (New York). 

Cecidomyia asteris Felt 1907a: 45. USA: New York, Albany; HT; ♂; NYSM. 

aulacaspidis. Italy. Aulacaspis rosae (Hemiptera: Diaspididae). 

Cecidomyella aulacaspidis Del Guercio 1918a: 247. Italy; ST; ♂,♀,L; lost. 

auriculata. India (Uttar Pradesh, West Bengal). 

Lestodiplosis auriculata Grover 1979c: 50. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

basalis. USA (New York). 

Lestodiplosis basalis Felt 1908b: 408. USA: New York, Albany; HT; ♂; NYSM. 

braziliensis. Brazil (Rio de Janeiro). Anadiplosis pulchra (Diptera: Cecidomyiidae). 

Coprodiplosis braziliensis Tavares 1920f: 113. Brazil: Rio de Janeiro, Nova Friburgo; HT; ♀; TAVC. 

brevilobata. India (Maharashtra). 

Lestodiplosis brevilobata Sharma 1988a: 155. India: Maharashtra, Aurangabad; HT; ♂; INPC. 

*cacaliae. Russia (Far East). on Cacalia hastata (Asteraceae). 

Lestodiplosis cacaliae Fedotova 2003c: 18. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

caliptera. USA (New York). 

Cecidomyia caliptera Fitch 1845b: 262. USA: New York, Salem; ST; ♂,♀; USNM. 

callida. UK, Germany. Dasineura papaveris (Diptera: Cecidomyiidae). 

Cecidomyia callida Winnertz 1853: 255. Germany; ST; ♂,♀; lost. 

callipus. Guadeloupe, Trinidad. Apion brunneonigrum (Coleoptera: Curculionidae), Recurvaria sp. (Lepidoptera: 

Gelechiidae), Adaina bipunctata (Lepidoptera: Pterophoridae), Melanagromyza minima (Diptera: 

Agromyzidae), and Dasineura corollae (Diptera: Cecidomyiidae). 

Lestodiplosis callipus Gagné 1977b: 120. Trinidad: Simla; HT; ♂; USNM. 

carolinae. USA (North Carolina). Rosette bud gall of Solidago canadensis (Asteraceae). 

Cecidomyia carolinae Felt 1907a: 43. USA: North Carolina, Asheville; HT; ♂; NYSM. 

casta. Germany. Parepidosis sp. (Diptera: Cecidomyiidae). 

Phonodiplosis casta Möhn 1955d: 421. Germany: Taunus, Dauborn; HT; ♂; SMNS. 

centralis. UK, Netherlands, Germany. 

Cecidomyia centralis Winnertz 1853: 277. Germany; ST; ♀; lost. 

cerasi. USA (New York). In twig swellings Prunus serotina (Rosaceae). 

Lestodiplosis cerasi Felt 1908b: 407. USA: New York, West Nyack; HT; ♂; NYSM. 


Lestodiplosis ceylanicus Kieffer 1912a: 28. Sri Lanka: Peradeniya; HT; ♀; NZSI?. 

chrysanthemi. UK, France. Contarinia chrysanthemi (Diptera: Cecidomyiidae). 

Lestodiplosis chrysanthemi Kieffer 1913a: 226. France: Bitche; T; ♀; lost?. 

ciliatipennis. Seychelles Is. 

Lestodiplosis ciliatipennis Kieffer 1911: 325. Seychelles Is: Mahé, Cascade Estate; HT; ♀; BMNH?. 

cincta. USA (New York).

277 

Lestodiplosis cincta Felt 1908b: 408. USA: New York, Albany; HT; ♂; NYSM. 

cincticornis. Tanzania. 

Lestodiplosis cincticornis Kieffer 1913h: 99. Tanzania: Tavéta; HT; ♀; MNHN. 

cinctipes. USA (Massachusetts). 

Parallelodiplosis cinctipes Felt 1914d: 113. USA: Massachusetts, Framingham; HT; ♂; NYSM. 

cirsii. UK, Netherlands. Macrolabis cirsii (Diptera: Cecidomyiidae). 

Lestodiplosis cirsii Barnes 1928f: 146. UK: Kent; HT; ♀; BMNH. 

*cirsiiflorae. Russia (Far East). Cirsium schantarense (Asteraceae) 

Lestodiplosis cirsiiflorae Fedotova 2004e: 14. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

clematiflorae. USA (New York). Closed flowers of Clematis virginiana (Rosaceae). 

Lestodiplosis clematiflorae Felt 1908b: 409. USA: New York, Karner; HT; ♂; NYSM. 

coni. Germany, Latvia, Hungary, Russia, (Europe, ES). From cones Picea excelsa (Pinaceae). 

Coprodiplosis coni Kieffer 1920a: 16. Germany: vic. Munich; ST; ♂,♀,L; lost?. 

conyzae. Italy. Neomikiella beckiana (Diptera: Cecidomyiidae). 

Lestodiplosis conyzae Kieffer 1909b: 12. Italy: Vallombrosa; ST; ♂,♀; lost?. 

corticalis. France. Asynapta sp. (Diptera: Cecidomyiidae). 

Lestodiplosis corticalis Kieffer 1898f: 63. France: Moselle; T; ♀; lost?. 

cossae. USA (Illinois). Phylloxera spp. (Hemiptera: Phylloxeridae). 

Cecidomyia cossae Shimer 1869: 395. USA: Illinois, vic. Mt. Carroll; T; A; lost?. 

crataegifolia. USA (New York). Dasineura crataegibedeguar (Diptera: Cecidomyiidae). 

Lestodiplosis crataegifolia Felt 1908b: 408. USA: New York, Bath; ST; ♂,♀; NYSM. 

cruenta. France. Winnertzia sp. (Diptera: Cecidomyiidae). 

Lestodiplosis cruenta Kieffer 1898f: 62. France: Moselle; ST; ♂,♀; lost?. 

cryphali. France, Germany. Cryphalus fagi (Coleoptera: Scolytidae). 

Coprodiplosis cryphali Kieffer 1894ab: 1; as n. sp. also in Kieffer 1894d: 119 and 1901c: 34. France: vic. Bitche; ST; 

LT; lost?. 

czalikovae. Kazakhstan. Acarina on Artemisia dracunculus (Asteraceae). 

Lestodiplosis czalikovae Fedotova 2001h: 1487. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 18 km SE 


dactylopii. Italy. Dactylopius sp. (Hemiptera: Dactylopiidae). 

Adelgimyza dactylopii Del Guercio 1918a: 237. Italy; ST; ♂,♀; lost. 

diaspidis. Italy. Diaspis pentagona (Hemiptera: Diaspididae). 

Arthrocnodax diaspidis Kieffer 1910b: 131. Italy: Acerra, nr Naples; T; ♀; lost?. An original drawing of a female 

flagellomere of this species shows the long neck more typical of Lestodiplosis; that this species feeds on a 

diaspidid instead of Acarina, the usual diet of Arthrocnodax spp., is a further indication. 

erecta. India (Kerala). 

Moreschiella erecta Nayar 1949: 88. India: Kerala, Trivandrum; HT; ♂; ZSCI. 

eupatorii. USA (New York). Schizomyia eupatoriflorae (Diptera: Cecidomyiidae). 

Cecidomyia eupatorii Felt 1907a: 44. USA: New York, L. Champlain, Glen L., South Bay; HT; ♂; NYSM. 

fascipennis. UK, Netherlands, Germany. In rotten wood. 

Cecidomyia fascipennis Winnertz 1853: 253. Germany; ST; ♂,♀; lost. 

fenestra. USA (New York). 

Arthrocnodax fenestra Felt 1908b: 404. USA: New York, Nassau; HT; ♂; NYSM. 


Cecidomyia filicis Felt 1907a: 40. USA: New York, Karner; HT; ♂; NYSM. 

*fimbripetalis. Russia (Far East). on Fimbripetalum radians (Caryophyllaceae). 

Lestodiplosis fimbripetalis Fedotova 2003c: 22. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

fimicola. Hawaiian Is. In rotting organic matter. 

Lestodiplosis fimicola Hardy 1960: 291. Hawaiian Is: Oahu, Honolulu, Univ. Hawaii campus; HT; ♂; BPBM. 

flaveolata. Germany.

278 

Cecidomyia flaveolata Winnertz 1853: 265. Germany; ST; ♂; lost. 

flavomarginata. USA (New York). 

Cecidomyia flavomarginata Felt 1907a: 42. USA: New York, Albany; HT; ♂; NYSM. 

florida. Canada (Ontario), USA (Florida). Leaf rolls on Crataegus sp. (Rosaceae). 

Lestodiplosis florida Felt 1908b: 409. USA: Florida, Ft. George; ST; ♂,♀; USNM. 

floridana. USA (Florida). Various insect larvae in flower heads Bidens pilosa (Asteraceae). 

Lestodiplosis floridana Johannsen 1945: 8. USA: Florida, Englewood; HT; ♂; CUIC. 

fratricida. France. Mycodiplosis tremulae (Diptera: Cecidomyiidae). 

Lestodiplosis fratricida Kieffer 1895p: 9, nomen nudum. 

Lestodiplosis fratricida Kieffer 1898f: 63. France: Moselle; T; ♂; lost?. 

fraxinifolia. USA (New Jersey). Rolled leaves of Fraxinus sp. (Oleaceae). 

Lestodiplosis fraxinifolia Felt 1908b: 408. USA: New Jersey, Newfoundland; HT; ♂; NYSM. 

frirenii. UK, France, Czechoslovakia. Contarinia ramicola (Diptera: Cecidomyiidae). 

Cecidomyia frirenii Kieffer 1888e: 109. France: Metz; ST; ♀; lost?. 

Lestodiplosis friereni Kieffer 1895f: cxciii, missp. of frirenii. 

Lestodiplosis frireni Kieffer 1913i: 197, missp. of frirenii. 

gagnei. Colombia. Elaeidobius subvittatus (Coleoptera: Curculionidae). 

Lestodiplosis gagnei Baylac 1987b: 125. Colombia: San-Alberto; HT; ♂; MNHN. 

gammae. UK. Dasineura or Rabdophaga sp. (Diptera: Cecidomyiidae). 

Lestodiplosis gammae Barnes 1928b: 72. UK: England, Kent; HT; ♂; BMNH. 

giardi. UK, France. Contarinia jacobeae (Diptera: Cecidomyiidae). 

Lestodiplosis giardi Kieffer 1896a: 14. France: Moselle; T; A; lost?. 

globosa. USA (New Jersey). Subglobular leaf gall of Populus tremuloides (Salicaceae). 

Lestodiplosis globosa Felt 1908b: 409. USA: New Jersey, Newfoundland; HT; ♂; NYSM. 

gracilis. Netherlands. Giraudiella inclusa (Diptera: Cecidomyiidae). 

Lestodiplosis gracilis Nijveldt 1953b: 284. Netherlands: nr Winterswijk; HT; ♂; ZMAN. 

grassator. Nearctic (widespread eastern). Daktuosphaira vitifoliae (Hemiptera: Phylloxeridae); various insects, incl. 

larvae in flower heads of Chromolaena odorata (Asteraceae) (cf. to list under Lestodiplosis callipus). 

Diplosis grassator Fyles 1883: 237. Canada: Quebec, Cowansville; ST; A,P,L,E; CNCI?. 

guttata. Poland. 

Cecidomyia guttata Loew 1850a: 32. Poland; ST; ♂,♀; ZMHB?. 

heterobiae. UK, Netherlands. Rabdophaga heterobia (Diptera: Cecidomyiidae). 

Lestodiplosis heterobiae Barnes 1928b: 70. UK: England, Kent, Wye, King's Wood; HT; ♂; BMNH. 

heterofila. India (Uttar Pradesh). 

Hadrobremia heterofila Grover 1965a: 35. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

hicoriae. USA (New York). 

Cecidomyia hicoriae Felt 1907a: 41. USA: New York, Nassau; HT; ♂; NYSM. 

hieracii. UK. Contarinia sp. (Diptera: Cecidomyiidae). 

Lestodiplosis hieracii Barnes 1928d: 144. UK: Kent; HT; ♀; BMNH. 

holstei. Germany, Poland, Czechoslovakia. Cones of Picea sp. (Pinaceae). 

Lestodiplosis holstei Kieffer 1920a: 17. Germany: vic. Munich; ST; ♂,L; lost?. 

hopkinsi. USA (California). In burrows of Scolytidae (Coleoptera). 

Cecidomyia hopkinsi Felt 1911l: 465, nomen nudum. 

Cecidomyia hopkinsi Felt 1911n: 554. USA: California, San Francisco; HT; ♀; USNM. 

hordei. UK. Stackelbergiella hordei (Diptera: Cecidomyiidae). 

Lestodiplosis hordei Barnes 1928f: 145. UK: England, Kent; HT; ♀; BMNH. 

hyperici. Spain. Geocrypta braueri (Diptera: Cecidomyiidae). 

Coprodiplosis hyperici Tavares 1919b: 82. Spain: Pontevedra; ST; ♂,L; TAVC. 

inclusae. France. Giraudiella inclusa (Diptera: Cecidomyiidae). 

Lestodiplosis inclusae Kieffer 1909b: 16. France: Coeuvres; T; ♀; lost?. 

inermis. France. Cecidomyiidae (Diptera) in rotten wood. 

Lestodiplosis inermis Kieffer 1913a: 227. France: Bitche; T; ♂; lost?.

279 

irenae. USA (New York). 

Cecidomyia fraxini Felt 1907a: 42, jun. prim. hom. fraxini Bremi 1847a. USA: New York, Albany; HT; ♂; NYSM. 

Arthrocnodax irenae Gagné 1972b: 325, new name for Cecidomyia fraxini Felt. 

iridipennis. USA (New York). 

Lestodiplosis iridipennis Johnson 1929: 216. USA: New York, Oneonta; HT; ♂; MCZI. 

irregularifila. India (Maharashtra). 

Lestodiplosis irregularifila Kashyap 1989c: 120. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

jacobeae. UK. Contarinia jacobeae (Diptera: Cecidomyiidae). 

Lestodiplosis jacobeae Barnes 1928b: 69, fig. 1B. UK: England, Kent; HT; ♂; BMNH. 

juniperina. USA (New York). 

Cecidomyia juniperina Felt 1907a: 44. USA: New York, Nassau; HT; ♂; NYSM. 

† 

kiefferis. Quaternary - Madagascar copal. 

Lestodiplosis kiefferis Meunier 1905: 90. Madagascar, a copal fossil or sub-fossil; HT; ♂; lost?. 

lacciferi. Malaysia, Indonesia (Java). Kerria sp. (Hemiptera: Kerriidae). 

Dentifibula lacciferi Barnes 1935d: 525. Malaya: Bukit Badong; ST; ♂,♀; BMNH. 

*lactucae. Russia (Far East). Lactuca sibirica (Asteraceae) 

Lestodiplosis lactucae Fedotova 2004e: 13. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

lanceolatae. UK. Dasineura sp. (Diptera: Cecidomyiidae). 

Lestodiplosis lanceolatae Barnes 1928f: 145. UK: England, Kent; HT; ♀; BMNH. 

laticaulis. USA (Florida). Diaspis echinocacti (Hemiptera: Diaspididae). 

Lestodiplosis laticaulis Gagné 1993a: 342. USA: Florida, Monroe Co., Long Key St. Pk; HT; ♂; USNM. 

lineata. France. Cecidomyiidae (Diptera) in rotten wood. 

Lestodiplosis lineata Kieffer 1898f: 62. France: Moselle; T; ♀; lost?. 

liviae. UK, Poland. Livia juncorum (Hemiptera: Psyllidae). 

Lestodiplosis liviae Rübsaamen 1901: 94. Poland: vic. Sp dziny woj. Olsztyn; ST; ♂,♀; ZMHB. 

longifilis. France, Czechoslovakia, Russia (Europe). Mycodiplosis boleti (Diptera: Cecidomyiidae) and Sciaridae 

(Diptera) in a woody fungus. 

Coprodiplosis longifilis Kieffer 1901d: 174. France: Bitche; ST; ♂,L; lost?. 

*lonicericola. Russia (Far East). on Lonicera maackii (Caprifoliaceae). 

Lestodiplosis lonicericola Fedotova 2003c: 20. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

lumiensis. Tanzania. 

Coprodiplosis lumiensis Kieffer 1913h: 97. Tanzania: Tavéta; HT; ♀; MNHN. 

lunata. India (Uttar Pradesh). 

Lestodiplosis lunata Grover & Bakhshi 1978: 42. India: Uttar Pradesh, Allahabad, HT; ♂; PGCA. 

luzonensis. Philippines. Parashorea malaanonan (Dipterocarpaceae). 

Tricontarinia luzonensis Felt 1918c: 293. Philippines: Luzon, Laguna Prov., Mt Makiling; ST; ♂,♀; NYSM. 

*maackiae. Russia (Far East). Gynandrobremia maackiaefloris and Ametrodiplosis maackiae (Diptera: 

Cecidomyiidae). 

Lestodiplosis maackiae Fedotova 2002c: 24. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

macrorosae. UK. Aphidoletes urticariae (Diptera: Cecidomyiidae). 

Lestodiplosis macrorosae Barnes 1928b: 72. UK: Scotland, St. Andrews; HT; ♂; BMNH. 

maculata. Germany. 

Cecidomyia maculata Winnertz 1853: 258. Germany; ST; ♀; lost. 

maculipennis. USA (Ohio). Clinodiplosis phlox (Diptera: Cecidomyiidae). 

Lestodiplosis maculipennis Greene 1941: 550. USA: Ohio, Chagrin Falls; ST; ♂,♀,L; USNM. 

*maricaensis. Brazil (Rio de Janeiro). 

Lestodiplosis maricaensis Santos & Maia 2009: 486. Brazil: Rio de Janeiro, Maricá (Restinga da Barra de Maricá); 

HT; ♂; MNRJ. 

marini. Spain. Contarinia scoparii (Diptera: Cecidomyiidae).

280 

Coprodiplosis marini Tavares 1919b: 83. Spain: Gallicia, Tuda; HT; ♂; TAVC. 

massalongoi. Italy. Dasineura salviae (Diptera: Cecidomyiidae). 

Lestodiplosis massalongoi Rübsaamen 1895b: 185. Italy; T; ♀; ZMHB. 

miastoris. France. Miastor sp. (Diptera: Cecidomyiidae). 

Lestodiplosis miastoris Kieffer 1913a: 228. France: Bitche; T; ♀; lost?. 

miki. Ireland, UK. Dasineura miki (Diptera: Cecidomyiidae). 

Lestodiplosis miki Barnes 1928b: 69, fig. 1A. UK: Caernarvonshire, & Ireland: Co. Dublin; HT; ♂; BMNH. 

mirabilis. France. 

Coprodiplosis mirabilis Kieffer 1895h: 120. France; T; A; lost?. 

morchellae. Germany. Insects in Morchella sp. (Pezizales). 

Lestodiplosis morchellae Rübsaamen 1911g: 350. Germany: Berlin; ST; ♂,♀; ZMHB. 

moricolata. Italy. Morus alba (Moraceae). 

Moreschiella moricola Del Guercio 1918a: 157. Italy: Abruzzi, Notaresco; ST; ♂,♀,L,E; lost. The original 

description indicates that the adults of this species are a Lestodiplosis and the larva is a Clinodiplosis. 

Moreschiella mori Del Guercio 1918a: 158, missp. of moricola. 

Lestodiplosis moricolata Gagné 2004b: 189, new name for Moreschiella moricola Del Guercio. 

moricola. South Africa. Diaspis pentagona (Hemiptera: Diaspididae). 

Arthrocnodax moricola Kieffer 1910b: 129. South Africa: Cape Town; T; ♀; lost?. An original drawing of a female 

flagellomere of this species shows the long neck more typical of Lestodiplosis; that this species feeds on a 

diaspidid instead of Acarina, the usual diet of Arthrocnodax spp., is a further indication. 

muricatae. UK. Wachtliella caricis (Diptera: Cecidomyiidae). 

Lestodiplosis muricatae Barnes 1928f: 147. UK: Kent; HT; ♀; BMNH. 

nana. Kazakhstan, Kirghizstan. 

Coprodiplosis nana Marikovskij 1960: 186. Kazakhstan: vic. Alma-Ata, & Kirghiztan: Frunze; ST; ♂; MARC. 

necans. Germany. Andricus fecundatrix (Hymenoptera: Cynipidae). 

Diplosis necans Rübsaamen 1892a: 384. Germany: vic. Berlin; ST; ♂,♀,L; ZMHB. 

novangliae. USA (Massachusetts). Epinotia "nanana" (Lepidoptera: Tortricidae). 

Lestodiplosis novangliae Felt 1933c: 114. USA: Massachusetts, Melrose Highlands; ST; ♂,♀,L; USNM. 

obtusilobata. Hawaiian Is. 

Lestodiplosis obtusilobata Hardy 1960: 292. Hawaiian Is: Oahu, Waialua; HT; ♂; HDOA. 

oomeni. Indonesia (Java). Calacarus carinatus (Acarina: Eriophyidae). 

Lestodiplosis oomeni Harris 1982: 20. Indonesia: Java, Bandung, Gambung; HT; ♂; BMNH. 

pallidicornis. Netherlands, France, Finland, Latvia, Lithuania, Russia (Europe). Dasineura leguminicola (Diptera: 


Lestodiplosis pallidicornis Kieffer 1898f: 63. France: Moselle; T; ♀; lost?. 

parricida. Germany. Clinodiplosis cilicrus (Diptera: Cecidomyiidae). 

Lestodiplosis parricida Rübsaamen 1906c: 233. Germany; ST; ♀,L; ZMHB. Germany. 

pavonia. Poland. 

Cecidomyia pavonia Loew 1850a: 33. Poland; ST; ♀; ZMHB?. 

perrisi. France. Tomicus ramulorum (Coleoptera: Scolytidae). 

Cecidomyia pictipennis Perris 1870: 170, jun. prim. hom. pictipennis Meigen. France: Landes; ST; ♂,♀,P,L; MNHN. 

Lestodiplosis perrisi Gagné 2004b: 189, new name for Cecidomyia pictipennis Perris 

peruviana. Peru, Guadeloupe. Pinnaspis strachani; Diaspis boisduvallii (Hemiptera: Diaspididae). 

Lestodiplosis peruviana Felt 1911a: 10. Peru: Piura; L (Gagné & Etienne 2009: 245); ♂; USNM. Ref.: Gagné & 

Etienne 2009: 245 (redescr.). 

phlomicoliphaga. Russia (Far East). Dasineura phlomicola (Diptera: Cecidomyiidae). 

Lestodiplosis phlomicoliphaga Kovalev 1972: 427. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

pini. UK, Netherlands. Bremia sp. (Diptera: Cecidomyiidae) & Pineus pini (Hemiptera: Aphididae). 

Lestodiplosis pini Barnes 1928b: 71, fig. 2G. UK: Cheshire, Delamere Forest; HT; ♂; BMNH. 

pisi. UK. Contarinia pisi (Diptera: Cecidomyiidae). 

Lestodiplosis pisi Barnes 1928c: 75, fig. 3A. UK: Kent; HT; ♂; BMNH. 

platanifolia. USA (New York). Prodiplosis platani (Diptera: Cecidomyiidae).

281 

Lestodiplosis platanifolia Felt 1908b: 410. USA: New York, New York; HT; ♂; NYSM. 

plicatricis. UK. Dasineura plicatrix (Diptera: Cecidomyiidae). 

Lestodiplosis plicatricis Barnes 1928d: 144. UK: Kent; HT; ♀; BMNH. 

polypori. UK, Germany, Latvia, Poland, Czechoslovakia, Russia (Europe). Insects in Polyporus spp. (Aphylloprales). 

Cecidomyia polypori Loew 1850a: 33. Poland; ST; ♂,♀; ZMHB?. 

Cecidomyia polypori Winnertz 1853: 249. Germany; ST; ♂,♀; lost. 

populifolia. USA (New York). Prodiplosis morrisi (Diptera: Cecidomyiidae). 

Lestodiplosis populifolia Felt 1908b: 408. USA: New York, Albany; HT; ♂; NYSM. 

*prunellaflorae. Russia (Far East). Prunella vulgaris (Lamiaceae) 

Lestodiplosis prunellaflorae Fedotova 2004e: 13. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

pugionis. USA (New York). Miastor metraloas (Diptera: Cecidomyiidae). 

Itonida pugionis Felt 1911l: 457, nomen nudum. 

Itonida pugionis Felt 1911n: 557. USA: New York, Nassau; ST; ♂,♀; NYSM. 

pulchella. Germany. In rotten wood. 

Cecidomyia pulchella Winnertz 1853: 254. Germany; ST; ♂,♀; lost. 

pyri. UK. Dasineura pyri (Diptera: Cecidomyiidae). 

Lestodiplosis pyri Barnes 1928c: 75, fig. 3B. UK: Devonshire, Essex; HT; ♂; BMNH. 

quercina. Portugal, Spain. Quercus robur (Fagaceae). 

Theatodiplosis quercina Tavares 1922c: 149. Spain: Gallicia, & Portugal; ST; ♂,♀; TAVC. 

quercus. Spain. Dasineura squamosa (Diptera: Cecidomyiidae). 

Coprodiplosis quercus Tavares 1919b: 80. Spain: Pontevedra; ST; ♂,♀; TAVC. 

*rabdosiae. Russia (Far East). Rabdosia excisa (Lamiaceae). 

Lestodiplosis rabdosiae Fedotova 2002c: 22. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

raphani. UK, Denmark, Netherlands, Germany, Czechoslovakia. Acarina in stored seed. 

Lestodiplosis raphani Barnes 1929: 119. Denmark: Lyngby; ST; ♂,♀; BMNH. 

roburella. Italy. On Quercus robur (Fagaceae). 

Moreschiella roburella Del Guercio 1918a: 171. Italy; ST; ♂,♀,L; lost. The original description indicates that the 

male and female of this species are a Lestodiplosis and the larva is a Clinodiplosis. 

rosarum. UK, Netherlands. Wachtliella rosae (Diptera: Cecidomyiidae). 

Lestodiplosis rosarum Barnes 1928b: 71, fig. 2E. UK: Kent; HT; ♂; BMNH. 

rosea. France. Peromyia aurantiaca (Diptera: Cecidomyiidae). 

Hemidiplosis rosea Kieffer 1894k: unnumbered p. 2. Unstated, presumably France: Bitche; HT; egg; lost? 

Lestodiplosis rosea Kieffer 1898f: 63. France: Moselle; T; ♀; lost?. Redescription. 

*rubripes. Russia (Far East). Artemisia rubripes (Asteraceae). 

Lestodiplosis rubripes Fedotova 2005a: 464. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♂; ZMAS. 

rudbeckiae. USA (North Carolina). In flowerheads of Rudbeckia hirta (Asteraceae). 

Cecidomyia rudbeckiae Beutenmüller 1907a: 388. USA: North Carolina, Buncombe Co., Valley of the Black Mts; 

ST; ♂,♀,L; AMNH. 

rufa. USA (New York). 

Arthrocnodax rufus Felt 1908b: 403. USA: New York, Albany; HT; ♂; NYSM. 

rugosae. USA (New York). Solidago sp. (Asteraceae). 

Cecidomyia rugosae Felt 1907a: 45. USA: New York, Albany; HT; ♂; NYSM. 

Cecidomyia rugosa Felt 1907c: 141, unjust. emend. of rugosae. 

rumicis. USA (New York). Poss. Contarinia rumicis (Diptera: Cecidomyiidae). 

Lestodiplosis rumicis Felt 1908b: 410. USA: New York, Newport; ST; ♂,♀; NYSM. 

satiata. USA (New York, PA, North Carolina). 

Lestodiplosis satiata Felt 1920a: 292. USA: New York, Keene Valley; HT; ♂; NYSM. 

scottianus. Seychelles Is. 

Lestodiplosis scottianus Kieffer 1911: 324. Seychelles Is: Mahé, Cascade Estate; HT; ♀; BMNH. 

scrophulariae. USA (New York). Distorted flower buds of Scrophularia marilandica (Scrophulariaceae).

282 

Cecidomyia scrophulariae Felt 1907d: 22. USA: New York, West Nyack; HT; ♂; NYSM. USA (New York). 

septemguttata. France. Cryphalus fagi (Coleoptera: Scolytidae). 

Lestodiplosis septemguttata Kieffer 1894k: unnumbered p. 2. Unstated, presumably France: Bitche; HT; egg; lost? 

Lestodiplosis septemguttata Kieffer 1898f: 62. France; ST; ♂,♀; lost?. Redescription. 

septemmaculata. USA (Illinois). Rabdophaga strobiloides (Diptera: Cecidomyiidae). 

Cecidomyia septemmaculata (as septem-maculata) Walsh 1864b: 630. USA: Illinois, nr Rock Island; ST; ♂,♀; 


solani. UK. Contarinia solani (Diptera: Cecidomyiidae). 

Lestodiplosis solani Barnes 1928d: 144. UK: England, Kent; HT; ♀; BMNH. 

solidaginis. USA (New York). Rhopalomyia albipennis (Diptera: Cecidomyiidae). 

Lestodiplosis solidaginis Felt 1908b: 409. USA: New York, Albany; HT; ♂; NYSM. 

*spicigera. Russia (Far East). 

Lestodiplosis spicigera Fedotova & Sidorenko 2006c: 111. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

spiraeafolia. USA (New York). Spiraea sp. (Rosaceae). 

Lestodiplosis spiraeafolia Felt 1908b: 410. USA: New York, Albany; HT; ♂; NYSM. 

strobilobii. Italy. Adelges laricis (Adelgidae; Hemiptera). 

Adelgimyza strobilobii Del Guercio 1918a: 231. Italy; ST; ♂,♀; lost. 

sylvestris. USA (North Carolina). 

Cecidomyia sylvestris Felt 1907a: 47. USA: North Carolina, Davidson's R.; HT; ♂; NYSM. 

syringopais. Indonesia (Sumatra), Australia (QLD). Phyllocnistis nepenthae (Lepidoptera: Phyllocnistidae). 

Coprodiplosis syringopais Hering 1931: 59. Indonesia: Sumatra, vic. L. Toba; ST; ♂,♀,P,L; ZMHB?. 

tarsonemi. Netherlands, Germany. Tarsonemus sp. (Acarina: Tarsonemidae). 

Lestodiplosis tarsonemi Rübsaamen 1895b: 184. Germany; ST; ♀,L; ZMHB. 

taxiconis. USA (Oregon). In cones of Pseudotsuga menziesii (Pinaceae). 

Lestodiplosis taxiconis Foote 1956: 56. USDA: Oregon, Colestin; HT; ♂; USNM. 

tetrachaeta. France. 

Coprodiplosis tetrachaeta Kieffer 1913a: 228. France: Bitche; T; ♂; lost?. 

tibialis. UK, Germany. Salix alba (Salicaceae). 

Cecidomyia tibialis Winnertz 1853: 253. Germany; ST; ♂; lost. 

tjanshanica. Kazakhstan, Kirghiztan. 

Plagiodiplosis tjanshanica Marikovskij 1960: 188. Kazakhstan: steppe and alpine meadows of Zailiïskiï Alatau, and 

Kirghiztan: Frunze; ST; ♂; MARC. 

tragopogonis. UK. Contarinia tragopogonis (Diptera: Cecidomyiidae). 

Lestodiplosis tragopogonis Barnes 1928f: 146. UK: England, Kent; HT; ♀; BMNH. 

traili. UK. Dasineura traili (Diptera: Cecidomyiidae). 

Lestodiplosis traili Barnes 1928b: 69, fig. 1D. UK: England, Kent; HT; ♂; BMNH. 

triangularis. USA (New York). Leaf gall Solidago sp. (Asteraceae). 

Cecidomyia triangularis Felt 1907a: 42. USA: New York, Lebanon; HT; ♂; NYSM. 

triangulata. India (Uttar Pradesh). 

Lestodiplosis triangularis Grover & Bakhshi 1978: 38, jun. sec. hom. triangularis Felt. India: Uttar Pradesh, 

Allahabad, Mumfordganj; HT; ♂; PGCA. 

Lestodiplosis triangulata Gagné 2004b: 191, new name for Lestodiplosis triangularis Grover & Bakhshi. 

trifaria. Japan (Kyushu). Resseliella odai (Diptera: Cecidomyiidae). 

Lestodiplosis trifaria Yukawa & Sanui 1978: 245. Japan: Kyushu, Miyazaki Pref., Ebino-City, Suenaga; HT; ♂; 

KUIC. 

trifolii. UK, France, Czechoslovakia. Dasineura trifolii (Diptera: Cecidomyiidae). 

Lestodiplosis trifolii Barnes 1928f: 146. UK: England, Kent; HT; ♀; BMNH. 

tripidiperda. Italy. Liothrips oleae (Thysanoptera: Phlaeothripidae). 

Adelgimyza tripidiperda Del Guercio 1918a: 240. Italy; T; ♀; lost. 

trivittata. Germany. On rotten wood. 

Lestodiplosis trivittata Rübsaamen 1921: 41. Germany: Kiel; ST; ♂,♀,L; ZMHB.

283 

tsugae. USA (New York). 

Cecidomyia tsugae Felt 1907a: 43. USA: New York, Lake Clear; HT; ♂; NYSM. 

urticae. UK, Netherlands. Dasineura urticae (Diptera: Cecidomyiidae). 

Lestodiplosis urticae Nijveldt 1951: 313. Netherlands: Amsterdam; HT; ♂; ZMAN. 

variegata. Netherlands, France, Germany. 

Cecidomyia variegata Macquart 1826: 171. France; T; A; MNHN?. 

vasta. Germany. Polyxenus lagurus (Diplopoda, Pselaphognatha). 

Chiliodiplosis vasta Möhn 1955d: 416. Germany: Mainz-Gonzenheim; HT; ♂; SMNS. 

venusta. USA (Missouri). Psilocorsis quercicella (Lepidoptera: Oecophoridae). 

Lestodiplosis venusta Gagné 1999b: 333. USA: Missouri, Troy Co., Cuivre R. St. Pk; HT; ♂; USNM. 

*verbasci. Russia (Europe). Verbascum orientale (Scrophulariaceae). 

Lestodiplosis verbasci Fedotova 2004b: 821. Russia: 30 km SE Samara, nr Ust-Kinel'skii; HT; ♂; ZMAS. 

verbenifolia. USA (New York). Rolled leaves of Verbena urticifolia (Verbenaceae). 

Lestodiplosis verbenifolia Felt 1908b: 408. USA: New York, Karner; HT; ♂; NYSM. 

*veronicae. Russia (Far East). Veronica longifolia (Scrophulariaceae) 

Lestodiplosis veronicae Fedotova 2004e: 14. Russia: Primorskiï Terr., 23 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

viburni. UK. Contarinia viburnorum (Diptera: Cecidomyiidae). 

Lestodiplosis viburni Barnes 1928b: 71, fig. 2H. UK: England, Kent; HT; ♂; BMNH. 

*viciaflorae. Russia (Far East). Vicia cracca (Fabaceae) 

Lestodiplosis viciaflorae Fedotova 2004e: 12. Russia: Primorskiï Terr., 23 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

vorax. UK, Germany. 

Diplosis vorax Rübsaamen 1892a: 384. Germany; ST; ♂,♀,L; ZMHB. 

winnertziae. France. Hartigiola annulipes (Diptera: Cecidomyiidae). 

Lestodiplosis winnertziae Kieffer 1909b: 8. France: Bitche; T; ♀; lost?. 

woeldickii. Italy. Acarina on feathers of stuffed birds. 

Cecidomya woeldickii Contarini 1839: 130. Italy; ST; ♂,♀; lost. 

xylodiplosuga. Germany. Xylodiplosis sp. (Diptera: Cecidomyiidae). 

Lestodiplosis xylodiplosuga Skuhravá 2001: 59. Germany: Rottenburg am Neckar; HT; ♂; SKCR. 

yuccae. USA. Pods of Yucca angustifolia (Liliaceae). 

Lestodiplosis yuccae Felt 1908b: 408. USA; HT; ♂; USNM. 

*LIANODIPLOSIS 

Lianodiplosis Fedotova & Sidorenko 2006a: 143. Type species, taeniata Fedotova & Sidorenko (orig. des.). 


*taeniata. Russia (Far East). 

Lianodiplosis taeniata Fedotova & Sidorenko 2006a: 144. Russia: Primorskiï Terr., Vladivostok; HT; ♂; ZMAS. 

LIEBELIOLA 

Liebeliola Kieffer & Jörgensen 1910: 428. Type species, prosopidis Kieffer & Jörgensen (mon.). 

Liebliola Felt 1918c: 320, missp. of Liebeliola 


prosopidis. Argentina (Mendoza, San Juan). Prosopis strombulifera (Fabaceae). 

Liebeliola prosopidis Kieffer & Jörgensen 1910: 428. Argentina: Mendoza, San Juan; ST; ♀,E; lost.

*LIGULODIPLOSIS 

Ligulodiplosis Fedotova 2006a: 150. Type species, fimbriata Fedotova (orig. des.). 


*fimbriata. Russia (Far East). 

Ligulodiplosis fimbriata Fedotova 2006a: 150. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

LIODIPLOSIS 

Liodiplosis Gagné 2001: 122. Type species, cylindrica Gagné (orig. des.). 

Neotropical; 3 spp., all from Mikania; Cecidomyiidi, Clinodiplosini. 

cylindrica. Brazil (Rio de Janeiro). Mikania glomerata (Asteraceae). 

Liodiplosis cylindrica Gagné 2001: 124. Brazil: Rio de Janeiro, Silva Jardim, Poço das Antas Biol. Res.; HT; ♂; 

QBUM. 

conica. Brazil (Rio de Janeiro). Mikania glomerata (Asteraceae). 

Liodiplosis conica Gagné 2001: 126. Brazil: Rio de Janeiro, Parque Nacional da Tijuca; HT; ♂; QBUM. 

spherica. Brazil (Rio de Janeiro). Mikania glomerata (Asteraceae). 

Liodiplosis spherica Gagné 2001: 127. Brazil: Rio de Janeiro, Silva Jardim, Poço das Antas Biol. Res.; HT; ♂; 

QBUM. 

LITCHIOMYIA 

Litchiomyia Yang 1999: 129. Type species, chinensis Yang & Luo (orig. des.). 

Oriental; 1 sp.; Lasiopteridi, unplaced to tribe. 

chinensis. China (Guangdong). Litchi sinensis (Sapindaceae). 

Litchiomyia chinensis Yang & Luo 1999: 129. China: Guangdong, Guangzhou; HT; ♂; SCUC. 

LOBOLASIOPTERA 

Lobolasioptera Möhn 1964b: 580. Type species, media Möhn (orig. des.). 


media. El Salvador. Pseudocalymma macrocarpa (Bignoniaceae). 

Lobolasioptera media Möhn 1964b: 581. El Salvador: La Libertad, nr San Diego; HT; ♂; SMNS. 

LOBOPTEROMYIA 

Lobopteromyia Felt 1908b: 385, 389. Type species, Contarinia filicis Felt (orig. des.). 

Nearctic; 1 sp.; Cecidomyiidi, Cecidomyiini. 


Contarinia filicis Felt 1907a: 35. USA: New York, Karner; HT; ♂; NYSM. 

Contarinia consobrina Felt 1907c: 161. USA: New York, Karner; HT; ♂; NYSM. 

284

Contarinia tiliae Felt 1907c: 161. USA: New York, Karner; HT; ♂; NYSM. 

Lobopteromyia abdominalis Felt 1908b: 390. USA: New York, Albany; HT; ♂; NYSM. 

Lobopteromyia apicalis Felt 1908b: 390. USA: New York, Albany; ST; ♂; NYSM. 

Lobopteromyia caricis Felt 1908b: 390. USA: New York, Karner; HT; ♂; NYSM. 

Lobopteromyia foetedi Felt 1908b: 390. USA: New York, Karner; HT; ♂; NYSM. 

Lobopteromyia symplocarpi Felt 1908b: 390. USA: New York, Karner; ST; ♂,♀; NYSM. 

LOEWIOLA 

Loewiola Kieffer 1896b: 5. Type species, Diplosis centaureae Loew (orig. des.). 

Löwiola Kieffer 1913i: 206, unjust. emend. of Loewiola 

Palearctic; 3 spp., on Asteraceae; Cecidomyiidi, unplaced to tribe. 

acroptilon. Kazakhstan. Acroptilon picris (Asteraceae). 

Loewiola acroptilon Marikovskij 1968: 118. Kazakhstan: Zailiïskiï Alatau; ST; ♂,♀,L; MARC. 

centaureae. Widespread Europe. Centaurea scabiosa; C. jacea (Asteraceae). 

Diplosis centaureae Löw 1875: 20. Austria: vic. Klosterneuberg & Weidling, nr Vienna; ST; ♂,♀,L; NHMW. 

serratulae. UK, France, Czechoslovakia, Yugoslavia. Serratula tinctoria (Asteraceae). 

Loewiola serratulae Kieffer 1905b: 201. France: Seine et Marne, Vulaines sur Seine; ST; ♂,P,L; lost?. 

LOPESIA 

Lopesia Rübsaamen 1908b: 29. Type species, brasiliensis Rübsaamen (mon.). Tavares 1908 used "Lopesia n. g." but 

noted that Rübsaamen had used the name earlier for the same genus and had priority. 

Lopesla Shinji 1944: 214, missp. of Lopesia. 

Nearctic, Neotropical, Afrotropical, Australasia; 18 spp.; Cecidomyiidi, Lopesiini. 

armata. Kenya. Acacia tortilis (Fabaceae). 

Lopesia armata Gagné 1993c: 120. Kenya: Kima, nr Sultan Hamid; HT; LT; USNM. 

*bilobata. Brazil (Minas Gerais). Guapira sp. (Nyctaginaceae). 

Lopesia bilobata Maia 2004a: 73. Brazil: Minas Gerais, Tiradentes, Serra de São José; HT; ♂; MNRJ. 

brasiliensis. Brazil (Rio de Janeiro, Santa Catarina). Ossaea sp. (Melastomataceae). 

Lopesia brasiliensis Rübsaamen 1908b: 30. Brazil: Rio de Janeiro, Fabrica, Santa Catarina, Tubarão; ST; ♂,♀,L; 

ZMHB. Ref.: Maia, V.C. 2007c (descr. pupa). 

caulinaris. Brazil (Rio de Janeiro). Calophyllum brasiliense (Clusiaceae). 

Lopesia caulinaris Maia 2003: 32. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

conspicua. Brazil (Rio de Janeiro). Calophyllum brasiliense (Clusiaceae). 

Lopesia caulinaris Maia 2003: 34. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

elliptica. Brazil (Rio de Janeiro). Calophyllum brasiliense (Clusiaceae). 

Lopesia elliptica Maia 2003: 37. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

grandis. Brazil (Rio de Janeiro). Dalbergiamyia ecastophylla (Fabaceae). 

Lopesia grandis Maia 2001a: 17. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

licaniae. USA (Florida). Licania michauxii (Chrysobalanaceae). 

Lopesia licaniae Gagné 1996b: 429. USA: Florida, St. Lucie Co., Indrio; HT; ♂; FSCA. 

linearis. Brazil (Rio de Janeiro). Calophyllum brasiliense (Clusiaceae). 

Lopesia linearis Maia 2003: 41. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

marginalis. Brazil (Rio de Janeiro). Couepia ovalifolia (Chrysobalanaceae). 

Lopesia marginalis Maia 2001a: 18. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

niloticae. Kenya. Acacia nilotica (Fabaceae). 

Lopesia niloticae Gagné 1993c: 121. Kenya: Rt. A109, Kapiti Plains, Malili Ranch, ca. 70 km SE Nairobi; HT; ♂; 

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286 


parinarii. Mozambique. Parinarium mobola (Rosaceae). 

Lopesia parinarii Tavares 1908: 163. Mozambique; ST; ♂,P; TAVC. 

*quadrata. Australia (South Australia). Melaleuca lanceolata (Myrtaceae). 

Lopesia quadrata Kolesik 1999b: 116. Australia: South Australia, Coorong Natl Pk; ♂; SAMA. 

*similis. Brazil (Rio de Janeiro). Protium heptaphyllum (Burseraceae). 

Lopesia similis Maia 2004b: 486. Brazil: Rio de Janeiro, Parque Nacional da Restinga de Jurubatiba, Lagoa de 

Carepebus; HT; ♂; MNRJ. 

simplex. Brazil (Rio de Janeiro). Protium icicariba (Burseraceae). 

Lopesia simplex Maia 2002b: 175. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

singularis. Brazil (Rio de Janeiro). Pouteria venosa (Sapotaceae). 

Lopesia singularis Maia 2001a: 20. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

*spinosa. Brazil (Minas Gerais). Croton floribundus (Euphorbiaceae). 

Lopesia spinosa Maia 2004a: 75. Brazil: Minas Gerais, Tiradentes, Serra de São José; HT; ♂; MNRJ. 

*tibouchinae. Brazil (Minas Gerais). Tibouchina candolleana (Melastomataceae). 

Lopesia tibouchinae Maia 2004a: 77. Brazil: Minas Gerais, Tiradentes, Serra de São José; HT; ♂; MNRJ. 

LOPESIELLA 

Lopesiella Tavares 1908: 145. Type species, combreti Tavares (mon). 

Lopeziella Felt 1911c: 41, missp. of Lopesiella. 

Afrotropical; 1 sp.; Cecidomyiidi, Lopesiini. 

combreti. Mozambique. Combretum galpinii (Combretaceae). 

Lopesiella combreti Tavares 1908: 148. Mozambique; ST; ♂,♀,P; TAVC. 

LOPHODIPLOSIS 

Lophodiplosis Gagné 1997d: 313. Type species, indentata Gagné (orig. des.). 

Australasian; 5 spp., on Melaleuca; Cecidomyiidi, unplaced to tribe. 

bidentata. Australia (Queensland). Melaleuca quinquenervia (Myrtaceae). 

Lophodiplosis bidentata Gagné 1997d: 322. Australia: Queensland, 9.4 mi. E Tully; HT; ♂; ANIC. 

cornuata. Australia (Queensland). Melaleuca viridiflora; M. nervosa (Myrtaceae). 

Lophodiplosis cornuata Gagné 1997d: 325. Australia: Queensland, Magnetic I.; HT; ♂; ANIC. 

denticulata. Australia (Queensland). Melaleuca quinquenervia; M. viridiflora (Myrtaceae). 

Lophodiplosis denticulata Gagné 1997d: 321. Australia: Queensland, Tully Heads Rd, 15.3 mi. ESE Tully; HT; ♂; 

ANIC. 

indentata. Australia (Queensland). Melaleuca quinquenervia; Melaleuca spp. (Myrtaceae). 

Lophodiplosis indentata Gagné 1997d: 317. Australia: Queensland, Tully Heads Rd, 15.3 mi. ESE Tully; HT; ♂; 

ANIC. 

trifida. Australia (Queensland). Melaleuca quinquenervia; M. dealbata (Myrtaceae). 

Lophodiplosis trifida Gagné 1997d: 327. Australia: Queensland, Tully Heads Rd., 15.3 mi. ESE Tully; HT; ♂; ANIC. 

Refs: Purcell et al. 2007 (biol), Gagné et al. 2009 (descr. larva). 

LUZONOMYIA 

Luzonomyia Felt 1918c: 282. Type species, symphoremae Felt (orig. des.). 

Oriental; 1 sp.; Cecidomyiidi, Asphondyliini, Schizomyiina.

symphoremae. Philippines. Symphorema luzonicum (Verbenaceae). 

Luzonomyia symphoremae Felt 1918c: 282. Philippines: Luzon, Laguna Prov., Mt Makiling; L (Gagné 1969e: 568); 

♂; NYSM. 

LYCIOMYIA 

Lyciomyia Kieffer & Jörgensen 1910: 412. Type species, gracilis Kieffer & Jörgensen (mon.). 


gracilis. Argentina (Mendoza). Lycium gracile (Solanaceae). 

Lyciomyia gracilis Kieffer & Jörgensen 1910: 412. Argentina: Mendoza; ST; ♂,♀; lost. 

LYGOCECIS 

Lygocecis Gagné 1975d: 7. Type species, Sackenomyia packardi Felt (orig. des.). 

Holarctic; 4 spp., all on Salix stems; Lasiopteridi, Dasineurini. 

caulentis. Kazakhstan. Salix caprea (Salicaceae). 

Lygocecis caulentis Fedotova 1985b: 29. Kazakhstan: Zailiïskiï Alatau, valley of Maloï Almatinki R.; HT; ♂; ZKAS. 

packardi. USA (Texas, Illinois, Massachusetts). Salix sp. (Salicaceae). 

Sackenomyia packardi Felt 1909b: 290. USA: Massachusetts, Canton; ST; ♂,♀; NYSM. 

porterae. USA (New Mexico). Salix sp. (Salicaceae). 

Rhabdophaga porterae Cockerell 1904: 155. USA: New Mexico, nr Las Vegas; ST; ♀,P; USNM. 

yanagi. Japan (Honshu, Kyushu). Salix sp.; S. babylonica, S. hondoensis (Salicaceae). 

Zygiobia yanagi Shinji 1938c: 178. Japan: Honshu; T; A; lost. 

Helicomyia chidai Monzen 1955b: 36. Japan: Iwate, Tokyo, & Wakayama Prefs; ST; ♂; KUEC. 

MACHAERIOBIA 

Machaeriobia Rübsaamen 1916a: 448. Type species, brasiliensis Rübsaamen (mon.). 

Ruebsaamenodiplosis, as Rübsaamenodiplosis, Tavares 1920e: 61. Type species, Uleela machaerii Kieffer (orig. 

des.). 


machaerii. Brazil (Santa Catarina). Machaerium sp. (Fabaceae). 

Uleella machaerii Kieffer 1913i: 101. Brazil: Santa Catarina, Tubarão; T; LT; ZMHB?. 

Machaeriobia brasiliensis Rübsaamen 1916a: 448. Brazil: Santa Catarina, Tubarão; ST; ♀,P,L; ZMHB. 

MACRODIPLOSIS 

Macrodiplosis Kieffer 1895f: cxciv. Type species, Diplosis dryobia Loew (orig. des.) = pustularis Bremi. 

Holarctic; 14 spp., the reared species all from Quercus; Cecidomyiidi, Cecidomyiini. 


Cecidomyia antennata Felt 1908b: 414. USA: New York, Albany; HT; ♂; NYSM. 

claytoniae. USA (New York). 

Cecidomyia claytoniae Felt 1907a: 36. USA: New York, Albany; HT; ♂; NYSM. 

287

288 

electra. USA (New York). 

Mayetiola electra Felt 1908b: 369. USA: New York, Albany; HT; ♀; NYSM. 

erubescens. USA (widespread eastern). Quercus sp.; Q. coccinea (Fagaceae). 

Cecidomyia erubescens Osten Sacken 1862: 200. USA: District of Columbia; T; G; USNM. 

Cecidomyia foliora Russell & Hooker 1908: 350. USA: Massachusetts; ST; ♂,♀,L,E; USNM, NYSM, UMEC. 

flavoscuta. USA (New York). 

Cecidomyia flavoscuta Felt 1907a: 41. USA: New York, Albany; HT; ♂; NYSM. 

flexa. Russia (Far East). Quercus mongolica (Fagaceae). 

Macrodiplosis flexa Kovalev 1972: 414. Russia: Sikhote-Alin Mts, Pidan Range, Mt Chandolaz; HT; ♂; ZMAS. 

inflexa. Switzerland. Quercus sp. (Fagaceae). 

Cecidomyia inflexa Bremi 1847b: 10. Switzerland: vic. of Zurich; T; G; lost?. 

majalis. Nearctic (widespread eastern). Quercus palustris (Fagaceae). 

Cecidomyia quercus majulis Osten Sacken 1870: 53, invalid, a polynomial. 

Cecidomyia quercus majalis Osten Sacken 1871: 347, invalid, a polynomial. 

Cecidomyia majalis Osten Sacken 1878: 6. USA: New York; T; G; lost. 

niveipila. Nearctic (widespread eastern). Quercus sp. (Fagaceae). 

Cecidomyia niveipila Osten Sacken 1862: 199. USA: DC; T; LT; lost. 

pustularis. Widespread Europe, Kazakhstan. Quercus sp.; Q. robur, Q. petraea (Fagaceae). 

Cecidomyia pustularis Bremi 1847b: 10. Switzerland: vic. of Zurich; T; G; lost?. 

Cecidomyia reaperiens Bach 1859: 252. Austria; T; G; ?. Cf. Löw 1877: 14. 

Diplosis dryobia Löw 1877: 14. Austria: Vienna; ST; ♂,♀,L; NHMW. 

putrida. USA (New York). 

Itonida putrida Felt 1912l: 246. USA: New York, Albany; HT; ♂; NYSM. 

qoruca. USA (Illinois). Quercus tinctoria (Fagaceae). 

Cecidomyia qoruca (as q-oruca) Felt 1925c: 61. USA: Illinois; T; G; USNM. 

Macrodiplosis quercusoruca Gagné 1973c: 874, unjust. emend. of qoruca. 

quercus. Poland. Quercus robur (Fagaceae). 

Cecidomyia quercus Loew 1850a: 22. Poland; ST; LT; ZMHB?. This name has not been used since originally used 

except for a reference in Löw 1877. 

roboris. Widespread Europe, Turkey, Kazakhstan. Quercus robur; Quercus spp. (Fagaceae). 

Cecidomyia roboris Hardy 1854d: 108. UK: Scotland, vic. Penmanshiel; ST; ♀,L; lost. Bland (2000: 10) designated 

roboris as a nomen oblitum and the junior synonym of volvens Kieffer citing Article 23.9, International Code 

of Zoological Nomenclature 1999, but neglected to show that the conditions of Article 23.9.2 were met. 

Macrodiplosis volvens Kieffer 1895p: 9. France: Moselle; ST; ♂,L; lost?. 

MACROLABIS 

Macrolabis Kieffer 1892e: 214. Type species, Cecidomyia pilosellae Binnie (subs. des. Felt 1911: 44). 

Schueziella Möhn 1961c: 1. Type species, Dasyneura fagicola Barnes (orig. des.). 

Holarctic; 60 spp.; Lasiopteridi, Dasineurini. All except one species are Palearctic, the remaining one eastern 

Nearctic. Most species are inquilines in galls of other cecidomyiids. Males have a strikingly enlarged gonocoxite. Refs: 

Rübsaamen & Hedicke 1938b (descr.), Sylvén & Tastás-Duque 1993 (descr.), Fedotova 2000a (host list), Dorchin et al. 

2006 (first N. Amer. record). 

achilleae. UK, Germany, Russia (Europe). Achillea millefolium (Asteraceae). 

Macrolabis achilleae Rübsaamen 1893: 164. Germany; ST; ♂; ZMHB. 

aconiti. Kazakhstan. Aconitum leucostomum (Ranunculaceae). 

Macrolabis aconiti Fedotova 2000a: 465. Kazakhstan: southern Altaï, L. Markakol', nr Urunkhaïka, 60 km N 


*aedopodiicola. Kazakhstan. Aegopodium alpestre (Apiaceae). 

Macrolabis aedopodiicola Fedotova 2004c: 730. Kazakhstan: Zailiskii Alatau, Issykskoe Canyon; HT; ♂; ZKAS.

289 

alatauensis. Kazakhstan. Rosa beggeriana (Rosaceae). 

Macrolabis alatauensis Fedotova 2000a: 469. Kazakhstan: Ketmen Mtn Range, Kyrgyz Sai canyon, nr Podgornoe; 


alnicola. Germany, Czechoslovakia. Alnus incana (Betulaceae). 

Macrolabis alnicola Rübsaamen 1914: 101. Germany: Remagen & Jungfernheide nr Berlin; ST; ♂,L; ZMHB. 

*altaicus. Kazakhstan. Achillea asiatica (Asteraceae). 

Macrolabis ptarmicae Fedotova 2004c: 728. Kazakhstan: Azutau Range, 20 km NE Alekseeva; HT; ♂; ZKAS. 

*americana. USA (Pennsylvania). Solidago gigantea, S. rugosa (Asteraceae). 

Macrolabis americana Dorchin 2006: 657. USA: Pennsylvania, Lairdsville; HT; ♂; USNM. 

aquilegiae. Netherlands, France, Czechoslovakia, Portugal. Aquilegia vulgaris (Ranunculaceae). 

Perrisia aquilegiae Kieffer 1909b: 2. France: Nancy; ST; LT; lost?. 

bedeguariformis. Germany. Populus tremula (Salicaceae). 

Cecidomyia bedeguariformis Rudow 1875a: 87. Germany; T; A; ?. 

Cecidomyia bedeguariformans Rudow 1875b: 250, missp. of bedeguariformis. 

brunellae. UK, Portugal, Spain, Russia (Europe). Prunella vulgaris (Lamiaceae). 

Macrolabis brunellae Tavares 1907b: 111. Portugal: Gerez; ST; ♂,♀,P,L; TAVC. Skuhravá et al. 2006: 119 

considers this species synonymous with M. ruebsaameni. 

buhri. Germany. Inula britannica (Asteraceae). 

Macrolabis buhri Stelter 1956: 117. Germany: Kr. Rostock, Gross Lüsewitz; ST; ♂,♀; DEIC. 

calophacae. Kazakhstan. Calophaca soongorica (Fabaceae). 

Macrolabis calophacae Fedotova 1989e: 33. Kazakhstan: Tarbagataï Mtn Range, 40 km NW Urdzhar, nr 


*chamaecytisicola. Russia. Chamaecytisus ruthenicus (Fabaceae). 

Macrolabis chamaecytisicola Fedotova 2004c: 731. Russia: Middle Volga area, Zhiguli Nature Res., nr Bakhilova 

Polyana; HT; ♂; ZKAS. 

cirsii. UK, Germany, Czechoslovakia. Cirsium arvense, C. lanceolatum (Asteraceae). 

Cecidomyia cirsii Rübsaamen 1890c: 236. Germany: vic. Siegen; ST; ♂,♀,L; ZMHB. 

*cotoneasteris. Kazakhstan. Cotoneaster oligantha (Rosaceae). 

Macrolabis cotoneasteris Fedotova 2004c: 720. Kazakhstan: Cental Tarbagatai, upper course of Karabuga R.,80 km 

S Aksaut; HT; ♂; ZKAS. 

delphinii. Kazakhstan. Delphinium iliense; D. elatum (Ranunculaceae). 

Macrolabis delphinii Fedotova 1987d: 35. Kazakhstan: Zailiïskiï Alatau, Turgen' canyon, 19 km S Turgen', 80 km SE 

Alma-Ata; HT; ♂; ZKAS. 

dulcamarae. Germany. Solanum dulcamara (Solanaceae). 

Cecidomyia dulcamarae Rübsaamen 1892b: 397. Germany: vic. Berlin; ST; ♂,L; ZMHB. 

euphorbiae. Kazakhstan. Euphorbia soongarica (Euphorbiaceae). 

Macrolabis euphorbiae Fedotova 1983c: 24. Kazakhstan: Dzhungarskiï Alatau, left bank Ili R.; HT; ♂; ZMAS. 

fagicola. Sweden, Germany, Czechoslovakia. Fagus sylvatica (Fagaceae). 

Dasyneura fagicola Barnes 1939a: 42. Germany: Schleswig-Holstein; ST; ♂,♀,L; BMNH. 

floricola. Germany, Poland. Tilia parvifolia; T. platyphyllos, T. cordata (Tiliaceae). 

Cecidomyia floricola Rudow 1875b: 240. Germany; ST; ♂,♀,L; ?. 

heraclei. Widespread Europe. Heracleum sphondylium, Chaerophyllum aromaticum (Apiaceae). 

Cecidomyia heraclei Kaltenbach 1862: 34. Germany; ST; LT; NHMW?. 

Cecidomyia corrugans Löw 1877: 11. Austria: Vienna; ST; ♂,♀,L; NHMW. 

Macrolabis tamujana Tavares 1922c: 128. Spain: Ribeira do Tamuje (Pontevedra), & Portugal: Entre-os-Rios 

(Douro); ST; ♂,♀,P,L; TAVC. 

hieracii. Widespread Europe. Hieracium boreale, H. murorum; H. umbellatum (Asteraceae). 

Macrolabis hieracii Rübsaamen 1917: 36. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

*hieraciflorae. Kazakhstan. Hieracium virosum (Asteraceae). 

Macrolabis hieraciflorae Fedotova 2004c: 724. Kazakhstan: Central Altai, Koksiuskii Range, Lyamin Belok Mt., 60 

km NE Leninogorsk; HT; ♂; ZKAS. 

hippocrepidis. UK, Spain, France, Germany, Switzerland, Austria, Italy. Hippocrepis comosa (Fabaceae).

290 

Macrolabis hippocrepidis Kieffer 1898f: 59. France, Switzerland, Germany, Austria; ST; ♂,♀; lost?. 

holosteae. UK, Germany, Poland, Czechoslovakia. Stellaria holostea (Caryophyllaceae). 

Macrolabis holosteae Rübsaamen 1917: 37. Germany: Reinbeck; ST; ♂,♀,L; ZMHB. 

incognita. Germany. Peucedanum oreoselinum (Apiaceae). 

Macrolabis incognita Stelter 1975b: 242. Germany: Bocksberg Nature Res., Kösterbeck nr Rostock; HT; ♂; SMTD. 

incolens. UK, Sweden, Netherlands, Germany, Czechoslovakia, Russia (Europe). Veronica chamaedrys 

(Scrophulariaceae). 

Macrolabis incolens Rübsaamen 1895b: 177. Germany; ST; ♂,♀; ZMHB. 

ivashczenkoae. Kazakhstan. Rosa kokanica (Rosaceae). 

Macrolabis ivashczenkoae Fedotova 2000a: 471. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Kshi-Kaïndy 

canyon, 12 km SE Novonikolaevka; HT; ♂; ZMAS. 

jaapi. UK, Netherlands, Denmark, Germany, Czechoslovakia. Galium aparine (Rubiaceae). 

Macrolabis jaapi Rübsaamen 1916b: 495. Germany: Jena; ST; ♂,♀; ZMHB. 

lamii. Widespread Europe. Lamium album (Lamiaceae). 

Macrolabis lamii Rübsaamen 1916b: 494. Germany; ST; ♂; ZMHB. 

laserpitii. Sweden, Germany, Czechoslovakia. Laserpitium latifolium; Chenopodium fischeri. Cnidium dubium, 

Pastinaca silvestris, Pimpinella saxifraga, Silaum silaus (Apiaceae). 

Macrolabis laserpitii Rübsaamen 1917: 38. Germany: Thüringen, Martinroda; ST; ♂,♀,L; ZMHB. 

leontopodii. Kazakhstan. Leontopodium fedtschenfoanum (Asteraceae). 

Macrolabis leontopodii Fedotova 1987d: 36. Kazakhstan: Terskeï Alatau, upper Bayankol R., 28 km SW Narynkol; 


linii. Kazakhstan. Linum heterosepalum; L. violascens (Linaceae). 

Macrolabis linii Fedotova 1987d: 34. Kazakhstan: Kungeï Alatau, 20 km SE Saryzhas; HT; ♂; ZKAS. 

lonicerae. UK, Netherlands, Germany, Poland, Czechoslovakia, Serbia. Lonicera periclymenum; L. caprifolium 

(Caprifoliaceae). 

Macrolabis lonicerae Rübsaamen 1912f: 355. Germany: vic. Remagen; ST; ♂,♀,L; ZMHB. 

lutea. Germany, Greece. Euphorbia cyparissias (Euphorbiaceae). 

Macrolabis lutea Rübsaamen 1914: 100. Germany; ST; ♂,♀,L; ZMHB. 

*mali. Italy. Malus domestica (Rosaceae). 

Macrolabis mali Anfora 2005: 96. Italy: 15 km N Trento, S. Michele all'Adige; HT; ♂; TNSM. 

markakolicus. Kazakhstan. Polygonum alpinum (Polygonaceae). 

Macrolabis markakolicus Fedotova 1997f: 448. Kazakhstan: southern Altaï, Markakol'skii Res., L. Markakol', nr 


marteli. UK, France. Hypericum perforatum (Hypericaceae). 

Macrolabis marteli Kieffer 1892f: 215. France: Elbeuf; ST; ♂,♀,L; lost?. 

orobi. Finland, Germany, Czechoslovakia, Austria, Hungary, Romania, Yugoslavia. Lathyrus vernus (Fabaceae). 

Cecidomyia orobi Löw 1877: 10. Austria: vic. Vienna; ST; ♂,♀,L; NHMW. 

*oxytropicola. Kazakhstan. Oxytropis puberula (Fabaceae). 

Macrolabis oxytropicola Fedotova 1992n: 102. Kazakhstan: Aigyr Kum Desert, 50 km NE Zaisan; HT; ♂; ZMAS. 

pavida. UK, Sweden, Netherlands, Germany, Czechoslovakia, Yugoslavia. Fraxinus excelsior (Oleaceae). 

Cecidomyia pavida Winnertz 1853: 245. Germany; ST; ♂,♀,L; MNHN?. 

*pedicularidis. Kazakhstan. Pedicularis achilleifolia (Scrophulariaceae). 

Macrolabis pedicularidis Fedotova 2004c: 722. Kazakhstan: Tarbagatai Range, upper coursof Ayaguz R., 40 km S 

Tarbagatai village; HT; ♂; ZKAS. 

pilosellae. Widespread Europe. Hieracium pilosella; H. echioides (Asteraceae). 

Cecidomyia pilosellae Binnie 1877: 179. UK: Scotland: Kilsyth Glen; Milngavie; ST; ♂,L; ?. 

podagrariae. Sweden, Germany, Poland, Czechoslovakia, Austria, Yugoslavia. Aegopodium podagraria (Apiaceae). 

Cecidomyia podagrariae Loew 1850a: 29. Poland; ST; G; ZMHB?. 

Cecidomyia aegopodii Loew 1850a: 22. Poland; ST; LT; ZMHB?. 

Macrolabis podagrariae Stelter 1962c: 85. Germany: Kr. Rostock, Gross Lüsewitz; HT; ♂; orig. DEIC, now SMTD. 

polemonii. Kazakhstan. Polemonium coeruleum (Polemoniaceae). 

Macrolabis polemonii Fedotova 2000a: 467. Kazakhstan: southern Altaï, L. Markakol', nr Urunkhaïka, 60 km N

291 


*ptarmicae. Kazakhstan. Ptarmica impatiens (Asteraceae). 

Macrolabis ptarmicae Fedotova 2004c: 727. Kazakhstan: Central Altai, Koksiuskii Range, Lyamin Belok Mt., 60 km 

NE Leninogorsk; HT; ♂; ZKAS. 

quercicola. Germany. Quercus robur (Fagaceae). 

Schueziella quercicola Stelter 1994a: 400. Germany: Kr. Rostock, betw. Gross Lüsewitz & Teschendorf; HT; ♂; 

DEIC?. 

rhodophila. Widespread Europe. Rosa sp.; R. cinnamomea (Rosaceae). 

Cecidomyia rhodophila Hardy 1850: 186. UK: Scotland, vic. Penmanshiel; T; ♀; lost. 

Macrolabis luceti Kieffer 1899b: 103. France: Seine Maritime, Les Petites-Dalles, & Bitche; ST; A,L; lost?. 

rubiae. Russia (Far East). Rubia sp. (Rubiaceae). 

Macrolabis rubiae Fedotova & Kovalev 2001: 453. Russia: Primorskiï Terr., Kedrovaya Pad'; HT; ♂; ZMAS. 

ruebsaameni. Sweden, Netherlands, Germany, Poland, Czechoslovakia, Hungary, Romania, Yugoslavia. Prunella 

grandiflora (Lamiaceae). 

Macrolabis brunellae Rübsaamen 1921: 50, jun. prim. hom. brunellae Tavares. Germany: aus dem Oytal nr 

Oberstdorf; ST; ♂,♀,L; ZMHB. 

Macrolabis ruebsaameni Hedicke 1938b: 322, new name for Macrolabis brunellae Rübsaamen. Skuhravá et al. 

2006: 119 considers this species synonymous with M. brunellae Tavares. 

saliceti. Germany, Czechoslovakia. Salix fragilis (Salicaceae). 

Cecidomyia saliceti Loew 1850a: 37. Germany; ST; ♂,♀; ZMHB?. 

Cecidomyia saliceti Winnertz 1853: 244. Germany; ST; ♂,♀,L; MNHN?. 

scrophulariae. Portugal, Spain. Scrophularia scorodonia (Scrophulariaceae). 

Macrolabis scrophulariae Tavares 1906a: 77. Portugal: Matta do Fundão; ST; ♂,♀; TAVC. 

*solidaginis. Kazakhstan. Solidago virgaurea (Asteraceae). 

Macrolabis solidaginis Fedotova 2004c: 724. Kazakhstan: Central Altai, Koksiuskii Range, Lyamin Belok Mt., 60 

km NE Lninogorsk; HT; ♂; ZKAS. 

sorbariae. Russia (Far East). Sorbaria sorbifolia (Rosaceae). 

Macrolabis sorbariae Fedotova & Kovalev 2001: 453. Russia: Primorskiï Terr., Arsen'yevka R. valley; HT; ♂; 

ZMAS. 

stellariae. UK, Sweden, Netherlands, France, Germany, Poland, Czechoslovakia, Romania, Yugoslavia, Russia 

(Europe). Stellaria media (Caryophyllaceae). 

Cecidomyia stellariae Liebel 1889b: 282. France: Bitche; ST; ♂,♀,L; lost. 

*swertiae. Kazakhstan. Swertia obtusa (Gentianaceae). 

Macrolabis swertiae Fedotova 2004c: 720. Kazakhstan: Sarymsakty Range, 45 km SW Katon-Karatai, nr Ornek; HT; 

♂; ZKAS. 

taraxaci. Kazakhstan. Taraxacum officinale (Asteraceae). 

Macrolabis taraxaci Fedotova 1987d: 36. Kazakhstan: Tarbagataï Mtn Range, 40 km NW Urdzhara, nr 


thalictri. Kazakhstan. Thalictrum collinum; T. minus (Ranunculaceae). 

Macrolabis thalictri Fedotova 2000a: 463. Kazakhstan: southern Altaï, L. Markakol', nr Urunkhaïka, 60 km N 


viciae. Russia (Far East). Vicia unijuga (Fabaceae). 

Macrolabis viciae Fedotova & Kovalev 2001: 453. Russia: Primorskiï Terr., Kedrovaya Pad'; HT; ♂; ZMAS. 

vicicola. Germany. Vicia cracca (Fabaceae). 

Macrolabis vicicolus Stelter 1992a: 179. Germany: Kr. Rostock, Freienholz, NNW Sanitz; HT; ♂; SMTD. 

MACROPORPA 

Macroporpa Rübsaamen 1916a: 437. Type species, peruviana Rübsaamen (subs. des. Gagné 1968d: 17). 

Neotropical; 2 spp.; Cecidomyiidi, Asphondyliini, Schizomyiina. Ref.: Möhn 1962 (illus.).

292 

peruviana. Brazil (Amazonas). Undet. sp. of Malpighiaceae. 

Macroporpa peruviana Rübsaamen 1916a: 437. Brazil: Amazonas, Auristella on R. Acre; L (Möhn 1962: 228); ♂; 

ZMHB. 

ulei. Brazil (Amazonas). Undet. sp. of Lauraceae. 

Macroporpa ulei Rübsaamen 1916a: 441. Brazil: Amazonas, S. Francisco on R. Acre; L (Möhn 1962: 231); ♂; 

ZMHB. 

*MAGADIPLOSIS 

Magadiplosis Fedotova & Sidorenko 2009b: 24. Type species, mera Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe here, but Fedotova and Sidorenko 2009b erect for this 

monotypic genus the subtribe Magadiplosina within the tribe Heterodiplosini (see Appendix). 

*mera. Russia (Far East). 

Magadiplosis mera Fedotova & Sidorenko 2009b: 24. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; 


MAMAEVIA 

Mamaevia Skuhravá 1967: 103. Type species, vysineki Skuhravá (orig. des.). 

Palearctic; 2 spp.; Cecidomyiidi, unplaced to tribe here, but Fedotova and Sidorenko 2009b place it in their subtribe 

Marikovskidiplosina within their tribe Quadridiplosini (See Appendix). 

hamifera. Kazakhstan. 

Tristephana hamifera Marikovskij 1960: 185. Kazakhstan: vic. Alma-Ata; HT; ♂; MARC. 

vysineki. UK, Germany, Latvia, Poland, Czech Republic. 

Mamaevia vysineki Skuhravá 1967: 103. Czech Republic: Bohemia, Vyšínek near Slaný; HT; ♂; SKCR. 

MANILKARAMYIA 

Manilkaramyia Maia 2001a: 22. Type species, notabilis Maia (orig. des.). 


notabilis. Brazil (Rio de Janeiro). Manilkara subsericea (Sapotaceae). 

Manilkaramyia notabilis Maia 2001a: 23. Brazil: Rio de Janeiro, Carapebus; HT; ♂; MNRJ. 

*MARIKOVSKIDIPLOSIS 

Marikovskidiplosis Fedotova & Sidorenko 2009b: 12. Type species, bullata Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe here, but Fedotova and Sidorenko 2009b erect the subtribe 

Marikovskidiplosina for this monotypic genus and place it within their tribe Quadridiplosini (see Appendix). 

*bullata. Russia (Far East). 

Marikovskidiplosis bullata Fedotova & Sidorenko 2009b: 13. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 


MARILASIOPTERA

Marilasioptera Möhn 1975: 62. Type species, tripartita Möhn (orig. des.). 


tripartita. El Salvador. Coccoloba sp. (Polygonaceae), in gall of undet. Cecidomyiidae. 

Marilasioptera tripartita Möhn 1975: 63. El Salvador: nr La Libertad; HT; ♀; SMNS. 

MASAKIMYIA 

Masakimyia Yukawa & Sunose 1976: 160. Type species, pustulae Yukawa & Sunose (orig. des.). 

Massakimaya Fedotova & Kovalev 2001: 407, 429, missp. of Masakimaya. 

Eastern Palearctic; 1 sp.; Lasiopteridi, unplaced to tribe. 

pustulae. Japan (Hokkaido to Kyushu), Republic of Korea. Euonymus japonicus; E. fortunei (Celastraceae). 

Masakimyia pustulae Yukawa & Sunose 1976: 160. Japan: Kagoshima City, Kôrimoto; HT; ♂; KUEC. 

MASSALONGIA 

Massalongia Kieffer 1897c: 12. Type species, Hormomyia rubra Kieffer (orig. des.). 

Palearctic; 4 spp., all on Betula; Cecidomyiidi, unplaced to tribe. 

altaica. Kazakhstan. Betula rotundifolia (Betulaceae). 

Massalongia altaica Fedotova 1990g: 61. Kazakhstan: central Altaï, Koksuïskiï Mtn Range, Lymin Belok Mt., 60 km 

NE of Leninogorsk; HT; ♂; ZKAS. 

bachmaieri. Germany, Russia (Europe). Betula nana (Betulaceae). 

Massalongia bachmaieri Möhn 1958: 1. Germany: Bavaria, nr Bernreid; HT; ♂; SMNS. 

betulifolia. UK. Betula pendula; B. pubescens (Betulaceae). 

Massalongia betulifolia Harris 1974: 137. UK: England, Cheshire, Lindow Common; HT; ♂; BMNH. 

rubra. Widespread Europe, Kazakhstan. Betula sp.; B. alba, B. pubescens (Betulaceae). 

Hormomyia rubra Kieffer 1890e: 199. France: vic. Bitche; T; G; lost?. 

Oligotrophus ruber Kieffer 1895c: lxxi. France: vic. Bitche; T; G; lost?. 

MAYETIOLA 

Mayetia Kieffer 1896b: 5, preocc. Mulsant & Rey 1875. Type species, Cecidomyia destructor Say (orig. des.). 

Mayetiola Kieffer 1896c: 89, new name for Mayetia Kieffer. 

Poomyia Rübsaamen 1910e: 337. Type species, Cecidomyia destructor Say (orig. des.). 

Chortomyia Kieffer 1913i: 63. Type species, Cecidomyia poae Bosc (orig. des.). 

Caulomyia Rübsaamen 1916b: 558. Type species, Oligotrophus radicificus Rübsaamen (orig. des.). 

Pemphigocecis Rübsaamen 1916b: 493. Type species, Oligotrophus ventricolus Rübsaamen (orig. des.). 

Palearctic except one Atlantic coast Nearctic species and M. destructor, an immigrant in North America and New 

Zealand; 29 spp., all on grasses; Lasiopteridi, Poomyini. Refs: Ertel 1975 (rev., larvae), Gagné and Hatchett 1989 

(descr. larva), Gagné et al. 1991 (descr.). 

agrostidis. Germany. Agrostis tenuis; A. stolonifera (Poaceae). 

Mayetiola agrostidis Ertel 1975: 18. Germany: Taunus, Dauborn; HT; LT; SMNS. 

Mayetiola agrostidis Coutin 2000: 353, jun. prim. hom. agrostidis Ertel. France: Chambourcy; ♂; MNHN. Ertel 1975 

was not cited in Coutin 2000 so, because Coutin's name may be a synonym of Ertel's, it is placed here to avoid 

coining a possibly unnecessary replacement name for the homonym. 

293

294 

agrostivora. Germany. Agrostis stolonifera (Poaceae). 

Mayetiola agrostivora Meyer 1985: 455. Germany: Schleswig-Holstein, 32 km NW Husum; HT; ♂; BMNH. 

alopecuri. Germany. Alopecurus pratensis (Poaceae). 

Mayetiola alopecuri (also as alopecura) Ertel 1975: 11. Germany: Leipzig; HT; LT; SMNS. 

ammophila. USA (New Jersey). Ammophila breviligulata (Poaceae). 

Mayetiola ammophila Gagné 1975e: 86. USA: New Jersey, Ocean Co., Island Beach St. Pk; HT; ♂; USNM. 

avenae. Widespread Europe; Algeria. Avena sativa; A. fatua (Poaceae). 

Cecidomyia avenae Marchal 1895a: 1. France; ST; A,L; MNHN?. 

baudysi. Czech Republic. Agropyron repens (Poaceae). 

Mayetiola baudysi Ertel 1975: 15. Czech Republic: Bohemia, Tauwald; HT; LT; SMNS. 

bifida. Germany. Calamagrostis lanceolata (Poaceae). 

Mayetiola bifida Kieffer 1909b: 3. Germany; T; LT; ZMHB?. 

bimaculata. UK, Netherlands, France, Germany, Latvia. Calamagrostis lanceolata; C. canescens, C. epigeios 

(Poaceae). 

Oligotrophus bimaculatus Rübsaamen 1895b: 181. Germany; T; ♀; ZMHB. 

Mayetiola calamagrostidis Kieffer 1909b: 3. Germany; T; G; ZMHB. 

Mayetiola spinulosa Kieffer 1909b: 3. Germany; T; LT; ZMHB. 

bromi. Austria, Romania. Bromus secalinus, B. tectorum (Poaceae). 

Cecidomyia bromi Hammerschmidt 1834: 719. Austria; ST; LT; ?. 

Phytophaga bromi Neacsu 1967: 185. Romania: Dobroudja, Agigea; ST; ♂,♀; NEAC. 

bromicola. Italy, Czechoslovakia, Hungary. Bromus sterilis, B. inermis, B. ramosus (Poaceae). 

Mayetiola bromicola Roberti 1953: 139. Italy: Campania, Portici; ST; ♂,♀,L; ?. 

buhri. Germany. Melica uniflora (Poaceae). 

Mayetiola buhri Ertel 1975: 8. Germany: Mühlhausen, Stadtwald; HT; LT; SMNS. 

clavata. France. 

Oligotrophus clavatus Kieffer 1901c: 23. France: Bitche; T; ♂; lost?. 

culacera. Germany. Calamagrostis canescens (Poaceae). 

Mayetiola culacera Stelter 1992b: 184. Germany: Kr. Rostock, Gross Lüsewitz, vic. Lüsewitzer Kruges; HT; ♂; 

SMTD. 

dactylidis. UK, France, Germany, Czechoslovakia, Ukraine, Yugoslavia. Dactylis glomerata (Poaceae). 

Mayetiola dactylidis Kieffer 1896e: 217. France: château de La Grange, nr Thionville; ST; A,P,L; lost?. 

destructor. Palearctic (widespread Europe, western Asia, Morocco to Tunisia); immigr.: Nearctic (wherever wheat is 

grown), New Zealand. Triticum aestivum; Triticum spp. Also occasional hosts: Hordeum vulgare, Secale 

cereale, Elymus sp., Bromus sp. (Poaceae). 

Cecidomyia destructor Say 1817: 45. USA: eastern; ST; F,L,E; lost. 

Phytophaga cerealis Rondani 1843: 159. Italy: vic. Parma; ST; ♂,♀,L; lost. 

Cecidomya frumentaria Rondani 1864: 189. Italy; ST; ♂,♀; MZUF. 

Cecidomyia culmicola Morris 1850: 194. USA: PA, Germantown; T; P; lost. 

Cecidomyia secalina Loew 1859: 7. Poland: Posen & Silesia; T; ♀; lost?. 

Rhabdophaga occidentalis Felt 1908b: 353. USA: California, Santa Clara Co.; HT; ♂; NYSM. 

Rhabdophaga pratensis Felt 1908b: 353. USA: New York, Albany; HT; ♂; NYSM. 

Rhabdophaga elymi Felt 1909b: 289. USA: California, Alameda; HT; ♀; USNM. 

Mayetiola orientalis Bollow 1955: 284. Turkey: Nihat Shevkey (eastern Turkey); HT; ♂; DEIC. 

Mayetiola secalis Bollow 1955: 273. Germany: Kasendorf; ST; ♂,♀; DEIC. 

festucae. Belgium. Festuca rubra (Poaceae). 

Mayetiola festucae Ertel 1975: 13. Belgium: St. Idesbald; HT; LT; SMNS. 

graminis. Widespread Europe. Poa trivialis; P. alpina, P. cenisia (Poaceae). 

Cynips graminis Fourcroy 1785: 385. France: Paris; T; G; lost. Fourcroy's description of the gall matches that of M. 

poae exactly. 

Cecidomyia poae Bosc d'Antic 1817: 133. France: Plessis-Piquet; ST; ♂,♀,L; lost. 

Cecidomyia graminicola Winnertz 1853: 292. Germany; ST; ♂,♀,L; lost. 

Cecidomyia graminis Brischke 1869: 1. Poland: vic. Gdansk; ST; ♂,♀,L; ?.

295 

hellwigi. UK, France, Netherlands, Denmark, Germany, Poland, Czechoslovakia, Hungary, Romania. Brachypodium 

sylvaticum; B. pinnatum (Poaceae). 

Poomyia hellwigi Rübsaamen 1912d: 217. Germany: Remagen; ST; ♂,♀,L; ZMHB. 

Mayetiola brachypodii Barnes 1927f: 212. UK: England, Kent, Wye; ST; ♂,♀; BMNH. 

hierochloae. Ukraine. Hierochloe repens (Poaceae). 

Cecidomyia hierochloae (as heirochloae) Lindeman 1888: 50. Ukraine: Poltava; ST; ♂,♀,L; ?. 

holci. UK, Netherlands, France, Germany, Czechoslovakia. Holcus mollis, H. lanatus (Poaceae). 

Mayetiola holci Kieffer 1896i: 333. France: château de La Grange, nr Thionville; ST; ♂,♀,L; lost?. 

hordei. Widespread Europe; Morocco, Algeria. Hordeum vulgare (Poaceae). 

Mayetiola hordei Kieffer 1909b: 9. Poland: Halicz; T; G; lost?. 

Phytophaga mimeuri Mesnil 1934: 246. Morocco & Algeria; ST; ♂,♀,L; ?. 

joannisi. UK, Finland, France, Germany. Poa nemoralis (Poaceae). 

Mayetiola joannisi (as joannissi) Kieffer 1896e: 216. France: château de La Grange, nr Thionville; ST; A,P,L; lost?. 

lanceolatae. UK, Germany, Latvia, Poland, Czechoslovakia. Calamagrostis lanceolata; Calamagrostis spp. (Poaceae). 

Oligotrophus lanceolatae Rübsaamen 1895b: 181. Germany; ST; ♂,♀; ZMHB. 

moliniae. UK, Germany, Latvia, Czechoslovakia. Molinia caerulea; M. arundinacea (Poaceae). 

Oligotrophus moliniae Rübsaamen 1895b: 180. Germany; T; ♀; ZMHB. 

phalaris. UK, Germany, Czechoslovakia. Phalaris arundinacea (Poaceae). 

Mayetiola phalaris Barnes 1927i: 375. Poland: Pomerania; ST; ♂,♀; BMNH. 

puccinelliae. Germany. Puccinellia maritima (Poaceae). 

Mayetiola puccinelliae Meyer 1984: 40. Germany: Schleswig-Holstein, Meldorf Inlet; HT; ♂; ZMKG. 

radicifica. UK, Netherlands, Germany, Poland, Romania. Poa nemoralis (Poaceae). 

Oligotrophus radicificus Rübsaamen 1895b: 179. Germany: Plötzensee & Berlin; ST; ♂,♀; ZMHB. 

Caulomyia radifica Barnes 1946b: 99, missp. of radicificus. 

schoberi. Denmark, Netherlands, Germany. Poa pratensis (Poaceae). 

Mayetiola schoberi Barnes 1958b: 690. Germany: Ratzeburg nr Lübeck; ST; ♂,♀; BMNH. 

ventricola. UK, Netherlands, Germany, Latvia, Czechoslovakia. Molinia coerulea; M. arundinacea (Poaceae). 

Oligotrophus ventricolus Rübsaamen 1899c: 566. Germany; T; A; ZMHB. 

MAYTENIELLA 

Mayteniella Maia 2001a: 24. Type species, distincta Maia (orig. des.). 


distincta. Brazil (Rio de Janeiro). Maytenus obtusifolia var. obovata (Celastraceae). 

Mayteniella distincta Maia 2001a: 25. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

MEGAULUS 

Megaulus Rübsaamen 1916a: 461. Type species, sterculiae Rübsaamen (mon.). 


sterculiae. Brazil (Amazonas). Sterculia sp. (Sterculiaceae). 

Megaulus sterculiae Rübsaamen 1916a: 463. Brazil: Amazonas, nr S. Francisco & Cobija on R. Acre; ST; ♀,P,L; 

ZMHB. 

MEGOMMATA 

Megommata Barnes 1939f: 333. Type species, seychelli Barnes (orig. des.).

296 

Afrotropical and Oriental; 6 spp., all predaceous as larvae on Coccidae or Pseudococcidae; Cecidomyiidi, unplaced to 

tribe. This genus is related to Diadiplosis, q.v. for comments. Ref.: Harris 1968 (rev.). 

donaldi. Ghana. Pseudococcus hargreavesi (Hemiptera: Pseudococcidae). 

Coccomyza donaldi Pritchard 1952b: 477. Ghana: Wiawso; HT; ♂; USNM. 

leefmansi. Indonesia (Java). Pulvinaria polygonata (Hemiptera: Coccidae). 

Coccomyza leefmansi Nijveldt 1954: 92. Indonesia: Java, Bogor; HT; ♂; ZMAN. 

psidii. Democratic Republic of Congo. Pulvinaria sp. (Hemiptera: Coccidae). 

Megommata psidii Barnes 1939f: 335. Democratic Republic of Congo: Rutshuru; ST; ♀; MRAC, BMNH. 

pulvinariae. Philippines. Pulvinaria sp. (Hemiptera: Coccidae). 

Microperrisia pulvinariae Felt 1915g: 174. Philippines: Luzon, Manila; ST; ♂,♀; NYSM. 

raoi. Malaysia. Megapulvinaria maxima (Hemiptera: Coccidae). 

Megommata raoi Harris 1968: 456. Malaysia: Selangor, Rubber Research Institute; HT; ♂; BMNH. 

seychelli. Seychelles Is. Pulvinaria sp. (Hemiptera: Coccidae). 

Megommata seychelli Barnes 1939f: 334. Seychelles Is: Derris I.; ST; ♂,♀; BMNH. 

MESODIPLOSIS 

Mesodiplosis Kolesik 2000c: 246. Type species, callitridis Kolesik (orig. des.). 


callitridis. Australia (South Australia). Callitris gracilis (Cupressaceae). 

Mesodiplosis callitridis Kolesik 2000c: 246. Australia: South Australia, Sandy Creek Conservation Pk; HT; ♂; 

SAMA. 

METASPHONDYLIA 

Metasphondylia Tavares 1918a: 41. Type species, squamosa Tavares (orig. des.). 

Neotropical; 1 sp.; Cecidomyiidi, Asphondyliini, Schizomyiina. The single known species might well fit into 

Schizomyia, q.v. for remarks. 

squamosa. Brazil (Bahia). Undet. sp. of Malvaceae. 

Metasphondylia squamosa Tavares 1918a: 42. Brazil: Bahia, Salvador; ST; ♂,♀,P,L; TAVC. 

METERDIPLOSIS 

Meterdiplosis Barnes 1927e: 272. Type species, nigrolineata Barnes (mon.). 


fici. Philippines. Ficus pruniformis, F. longipedunculata (Moraceae). 

Meterdiplosis fici Barnes 1928a: 22. Philippines: Luzon, Laguna Prov., Los Baños; ST; ♂,♀; BMNH. 

nigrolineata. Malaya. 

Meterdiplosis nigrolineata Barnes 1927e: 272. Malaya: Kuala Lumpur, Selangor; HT; ♀; BMNH. 

MEUNIERIELLA 

Meunieria Rübsaamen 1905b: 137, preocc. Kieffer 1904). Type species, dalechampiae Rübsaamen (orig. des.). 

Meunieriella Kieffer 1909b: 35, new name for Meunieria Rübsaamen. 

Dolicholabis Tavares 1918c: 72. Type species, lantanae Tavares (orig. des.).

Neotropical, Nearctic; 21 spp.; Lasiopteridi, Alycaulini. The genus is overwhelmingly Neotropical. Most species are 

known to be inquilinous in galls of other Cecidomyiidi but at least two, M. aquilonia and M. avicenniae, are gall 

makers. Ref.: Möhn 1975 (rev.). 

acaciae. El Salvador. Acacia riparioides (Fabaceae). 

Meunieriella acaciae Möhn 1975: 52. El Salvador: Ahuachapan, SW Turin, rd to Ahuachapan; HT; LT; SMNS. 

acalyphae. El Salvador. Acalypha unibracteata (Euphorbiaceae). 

Meunieriella acalyphae Möhn 1975: 45. El Salvador: San Salvador; HT; ♂; SMNS. 

aquilonia. USA (Pennsylvania). Gleditsia triacanthos (Fabaceae). 

Meunieriella aquilonia Gagné 1984b: 546. USA: Pennsylvania, Dauphin Co., Middle Paxton Township, 0.4 mi. N 

Clark Creek; HT; ♀; USNM. 

armeniae. El Salvador. Lonchocarpus sp. (Fabaceae). 

Meunieriella armeniae Möhn 1975: 50. El Salvador: La Libertad, W Ateos; HT; ♂; SMNS. 

avicenniae. Cuba, Guadeloupe. Avicennia germinans (Verbenaceae). 

Cecidomyia avicenniae Cook 1909: 145. Cuba; T; G; ?. 

cordiae. El Salvador. Cordia alliodora (Boraginaceae). 

Meunieriella cordiae Möhn 1975: 43. El Salvador: Santa Ana, SE Candelaria; HT; ♂; SMNS. 

dalechampiae. Brazil (Rio de Janeiro). Dalechampia filicifolia (Euphorbiaceae). 

Meunieria dalechampiae Rübsaamen 1905b: 137. Brazil: Rio de Janeiro, Palmeira; ST; ♂,♀,P; ZMHB. 

datxeli. Colombia. Triplaris/Symmeria/Ruprechtia sp. (Polygonaceae). 

Meunieriella datxeli Wünsch 1979: 123. Colombia: Magdalena, Bahia Nenguange; HT; ♂; SMNS. 

eugeniae. El Salvador. Eugenia sp. (Myrtaceae). 

Meunieriella eugeniae Möhn 1975: 51. El Salvador: La Libertad, N El Cimarrón; HT; LT; SMNS. 

gairae. Colombia. Cordia subtruncata (Boraginaceae). 

Meunieriella gairae Wünsch 1979: 127. Colombia: Magdalena, Bella-Vista; HT; ♂; SMNS. 

graciliforceps. Argentina (Mendoza). Prosopis strombulifera (Fabaceae). 

Lasioptera graciliforceps Kieffer & Jörgensen 1910: 429. Argentina: Mendoza; ST; ♂,♀,P; lost. 

ingae. El Salvador. Inga leptoloba (Fabaceae). 

Meunieriella ingae Möhn 1975: 59. El Salvador: La Libertad, N El Cimarrón; HT; ♂; SMNS. 

insignis. Brazil (Bahia). Protium heptaphyllum (Burseraceae). 

Dolicholabis insignis Tavares 1922a: 13. Brazil: Bahia, vic. Salvador; ST; ♂,♀; TAVC. 

lantanae. Brazil (Rio de Janeiro). Lantana sp. (Verbenaceae). 

Dolicholabis lantanae Tavares 1918c: 73. Brazil: Rio de Janeiro, Nova Friburgo; HT; ♂; TAVC. 

lonchocarpi. El Salvador. Lonchocarpus sp. (Fabaceae). 

Meunieriella lonchocarpae Möhn 1975: 49. El Salvador: La Libertad, Los Chorros; HT; ♀; SMNS. 

lucida. El Salvador. Ficus ovalis (Moraceae). 

Meunieriella lucida Möhn 1975: 54. El Salvador: La Libertad, NE Sitio del Niño, W lava fields; HT; ♂; SMNS. 

machaerii. El Salvador. Machaerium biovulatum (Fabaceae). 

Meunieriella machaerii Möhn 1975: 58. El Salvador: San Salvador; HT; ♀; SMNS. 

magdalenae. Colombia. Coccoloba candolleana (Polygonaceae). 

Meunieriella magdalenae Wünsch 1979: 119. Colombia: Magdalena, Bahia Nenguange; HT; ♂; SMNS. 

meridiana. El Salvador. Undet. sp. of Vernonieae (Asteraceae). 

Meunieriella meridiana Möhn 1975: 60. El Salvador: La Libertad, N El Cimarrón; HT; ♂; SMNS. 

pisoniae. El Salvador. Pisonia macranthocarpa (Nyctaginaceae). 

Meunieriella pisoniae Möhn 1975: 47. El Salvador: Ahuachapan, NW Las Chinamas; HT; LT; SMNS. 

randiae. El Salvador. Randia spinosa (Rubiaceae). 

Meunieriella randiae Möhn 1975: 56. El Salvador: San Salvador; HT; LT; SMNS. El Salvador. 

MICRODIPLOSIS 

Microdiplosis Tavares 1908: 155. Type species, zambezensis Tavares (mon.). 

297


zambezensis. Mozambique. Royena sp. (Ebenaceae). 

Microdiplosis zambezensis Tavares 1908: 155. Mozambique; ST; ♂,♀; TAVC. 

MICROLASIOPTERA 

Microlasioptera Skuhravá & Skuhravý 1981: 95. Lasioptera flexuosa Winnertz (orig. des.). 

Western Palearctic; 1 sp.; Lasiopteridi, Lasiopterini. 

flexuosa. Czechoslovakia, Austria, Hungary. Phragmites australis (Poaceae). 

Lasioptera flexuosa Winnertz 1853: 308. Germany; N (Skuhravá & Skuhravý 1981: 99, Czech Republic: nr 

B�eclav); ♂; SKCR. 

Lasioptera flexuosella Möhn 1968b: 139. Hungary: Gárdony; HT; LT; SMNS. 

MICROPLECUS 

Microplecus Kieffer 1913h: 107. Type species, Microplecus brevipalpis Kieffer (orig. des.). 


brevipalpis. Tanzania. 

Microplecus brevipalpis Kieffer 1913h: 108. Tanzania: Tavéta; HT; ♂; MNHN. 

MIKANIADIPLOSIS 

Mikaniadiplosis Gagné 2001: 128. Type species, annulipes Gagné (orig. des.). 


annulipes. Brazil (Rio de Janeiro). Mikania glomerata (Asteraceae). 

Mikaniadiplosis annulipes Gagné 2001: 129. Brazil: Rio de Janeiro, Rio de Janeiro; HT; M; QBUM. 

MIKIOLA 

Mikiola Kieffer 1896b: 5. Type species, Cecidomyia fagi Hartig (orig. des.). 

Palearctic; 5 spp., on Fagus leaf galls; Lasiopteridi, unplaced to tribe. 

*bicornis. Japan (Hokkaido to Kyushu). Fagus crenata (Fagaceae). 

Mikiola bicornis Sato & Yukawa 2009: 155. Japan: Kyoto Pref., Miyama Town, Asiu; HT; ♂; KUEC. 

cristata. France. Fagus sylvatica (Fagaceae). 

Mikiola cristata Kieffer 1898f: 60. France: Moselle; T; ♀; lost?. 

fagi. Widespread Europe; also Pliocene/Pas-de-la-Mougudo, Cantal, France. Fagus sylvatica; F. pliocenica (Fagaceae). 

Cecidomyia fagi Hartig 1839: 641. Germany; T; ♀; ?. 

*glandaria. Japan (Honshu and Kyushu). Fagus crenata (Fagaceae). 

Mikiola glandaria Sato & Yukawa 2009: 158. Japan: Hiroshima Pref., Higashihiroshima City, Mt Taganosu; HT; ♂; 

KUEC. 

orientalis. Turkey. Fagus sp. (Fagaceae). 

Mikiola orientalis Kieffer 1909a: 158. Turkey; T; LT; lost?. 

298

MIKOMYA 

Mikomya Kieffer 1912h: 2. Type species, Oligotrophus coryli Kieffer (orig. des.). 

Mikomyia Kieffer 1913i: 51, missp. of Mikomya. 


coryli. Widespread Europe. Corylus avellana (Betulaceae). 

Oligotrophus coryli Kieffer 1901c: 22. France: Bitche; T; ♂; lost?. 

Oligotrophus tympanifex Kieffer 1909b: 6. France; T; G; lost?. 

*MIRABILIDIPLOSIS 

Mirabilidiplosis Fedotova 2003e: 60. Type species, alutacea Fedotova (orig. des.). 

Eastern Palearctic; 4 spp.; Cecidomyiidi, Aphidoletini. This genus is close to Bremia if the two must be separated. 

*alutacea. Russia (Far East). 

Mirabilidiplosis alutaceus Fedotova 2003e: 62. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*spinosa. Russia (Far East). 

Mirabilidiplosis spinosa Fedotova 2004h: 184. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*undata. Russia (Far East). 

Mirabilidiplosis undata Fedotova 2004h: 181. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*villosa. Russia (Far East). 

Mirabilidiplosis villosa Fedotova 2004h: 181. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

MITODIPLOSIS 

Mitodiplosis Kieffer 1914: 516. Type species, graminis Kieffer (orig. des.). 


graminis. South Africa. Ehrhartia villosa (Poaceae). 

Mitodiplosis graminis Kieffer 1914: 516. South Africa: nr Cape Town; ST; ♀,P; lost?. 

MOEHNIELLA 

Moehniella Wünsch 1979: 66. Type species, fernandi Wünsch (orig. des.). 


fernandi. Colombia. Mimosa leiocarpa (Fabaceae). 

Moehniella fernandi Wünsch 1979: 67. Colombia, Magdalena Prov., Cerro San Fernando; HT; ♀; SMNS. 

MOHWADIPLOSIS 

Mohwadiplosis Rao 1957: 5. Type species, orientalis Rao (orig. des.). 

299


orientalis. India (Uttar Pradesh). Poss. predaceous on a sp. of Aleurodidae (Hemiptera). 

Mohwadiplosis orientalis Rao 1957: 5. India: Uttar Pradesh, Agra; HT; ♂; SJCA. 

MONARTHROPALPUS 

Monarthropalpus Rübsaamen 1892d: 381. Type species, Tipula flava Schrank (mon.). 

Palearctic, immigr. Nearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. 

flavus. Widespread Europe, Turkey; immigr.: USA (California, widespread eastern). Buxus sempervirens (Buxaceae). 

Tipula flava Schrank 1776: 55. Germany; T; A; lost?. 

Diplosis schineri Frauenfeld 1873: 184. Italy: Miramare; ST; ♀,P,L; ?. 

Cecidomyia buxi Laboulbène 1873: 321. France; L (Baylac 1987a: 264); ♂; MNHN. 

MONOBREMIA 

Monobremia Kieffer 1912h: 1. Type species, Bremia subterranea Kieffer (orig. des.). 

Palearctic, Oriental; 4 spp., the only reared species a predator of aphids; Cecidomyiidi, Aphidoletini. 

lobata. India (Uttar Pradesh). 

Monobremia lobata Grover & Bakhshi 1978: 88. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

longicornis. Russia (Far East). 

Monobremia longicornis Kovalev & Mamaev 1966: 229. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

HYNM. 

rishikeshensis. India (Uttar Pradesh). 

Monobremia rishikeshensis (also as rishikeshnsis) Grover 1979c: 70. India: Uttar Pradesh, rd to Rishikesh; HT; ♂; 

PGCA. 

subterranea. UK, Netherlands, France, Latvia, Romania, Russia (Europe). Metanipponaphis vandergrooti; Aphis spp., 

Dysaphis plantaginea, Nerudea roseus, Symydobius oblongus (Hemiptera: Aphididae). 

Bremia subterranea Kieffer 1888f: 27, nomen nudum. 

Monobremia subterranea Kieffer 1913g: 51. France: Bitche; ST; ♂,L; lost?. 

Phaenobremia rosea Barnes 1927d: 175. UK: England, Kent, Wye; HT; ♀; BMNH. 

MONODIPLOSIS 

Monodiplosis Rübsaamen 1910d: 289. Type species, Diplosis liebeli Kieffer (mon.). 


liebeli. Widespread Europe. Quercus sp.; Q. robur (Fagaceae). 

Diplosis liebeli Kieffer 1889b: 174. France: Moselle; ST; ♀,L; lost?. 

Schizomyia sociabilis Rübsaamen 1889a: 43. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

*MONTOSIDIPLOSIS 

Montosidiplosis Fedotova 2005c: 224. Type species, Montosidiplosis crenata Fedotova (orig. des.). 

Palearctic; 1 sp.; Cecidomyiidi, Lestodiplosini. 

300

*crenata. Russia (Far East). 

Montosidiplosis crenata Fedotova 2005c: 224. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. Kamenushka; HT; 

♂; ZMAS. 

MYCETODIPLOSIS 

Mycetodiplosis Kieffer 1912h: 1. Type species, Clinodiplosis boleti Kieffer (orig. des.). 


boleti. France. Polyporus confluens (Aphyllophorales). 

Clinodiplosis boleti Kieffer 1909b: 19. France: vic. Bitche; ST; ♂,♀,L; lost?. 

MYCOCECIS 

Mycocecis Edwards 1922: 105. Type species, ovalis Edwards (mon.). 


ovalis. UK, France, Germany, Denmark, Sweden, Russia (Europe). Hypoxylon rubiginosum (orig. as Hypochnus sp.) 

(Xylariales). 

Mycocecis ovalis Edwards 1922: 106. UK: England, North Sussex, Verdly; ST; ♂,♀; BMNH. Ref.: Spooner 2010 

(host, distr.). 

MYCODIPLOSIS 

Mycodiplosis Rübsaamen 1895b: 186. Type species, Cecidomyia coniophaga Winnertz (orig. des.). 

Isodiplosis Rübsaamen 1910d: 287. Type species, involuta Rübsaamen (orig. des. as n. g., n. sp.) = inimica Fitch. 

Toxomyia Felt 1911j: 302. Type species, fungicola Felt (orig. des.). 

Trilobiella Del Guercio 1918a: 244. Type species, siphae Del Guercio (orig. des. as n. g., n. sp.). 

Mycodiplossi Barnes 1927h: 220, missp. of Mycodiplosis. 

Kittada Agarwal 1956: 129. Type species, coimbatorensis Agarwal (orig. des. as n. g., n. sp.). 

Kittada Rao 1960a: 112, preocc. Agarwal 1956. Type species, pucciniae Rao (orig. des.). 

Cosmopolitan; 49 spp., mycophagous, especially on rusts and mildews; Cecidomyiidi, Mycodiplosini. Ref. Holz 1970 

(rev. Palearctic spp.). 

asiatica. Kazakhstan, Kirghiztan. 

Feltiella asiatica Marikovskij 1960: 187. Kazakhstan: vic. Alma-Ata, Kirghiztan: Frunze; T; ♂; MARC. 

australis. Argentina (Buenos Aires). 

Neobaeomyza australis Brèthes 1914: 152. Argentina: Buenos Aires, General Urquiza; HT; ♀; MACN. 

beliae. India (Maharashtra). A rust (Uredinales). 

Mycodiplosis beliae Chandra 1993d: 88. India: Maharashtra, Parbhani; HT; ♂; CSIA. 

buhri. Germany. Hyalopsora polypodii (Uredinales). 

Mycodiplosis buhri Holz 1970: 45. Germany: Haarz, Mägdesprung; HT; ♂; SMNS. 

ceomatis. Germany, Austria. Caeoma miniatum (Uredinales). 

Cecidomyia ceomatis Winnertz 1853: 278. Germany; ST; ♂,♀,L; lost. 

coimbatorensis. India (Tamil Nadu). Puccinia kuehnii (Uredinales). 

Kittada coimbatorensis Agarwal 1956: 129. India: Tamil Nadu, Coimbatore; HT; ♂; INPC. 

Kittada pucciniae Rao 1960a: 112. India: Tamil Nadu, Coimbatore; HT; ♂; ZSCI. 

Kittada saccharii Grover 1960: 1. India; HT; ♀; PGCA. 

301

302 

coniophaga. UK, Netherlands, France, Germany, Austria, Egypt, USA (New York, New Hampshire, New Jersey), 

Mexico (Texcoco). Phragmidium sp.; many other rusts (Uredinales). 

Cecidomyia coniophaga Winnertz 1853: 267. Germany: vic. Krefeld; ST; ♂,♀,L; MNHN?. 

Cecidomyia cerasifolia Felt 1907d: 21. USA: New Jersey, Newfoundland; HT; ♂; NYSM. 

Mycodiplosis cucurbitae Felt 1911l: 459, nomen nudum. 

Itonida cucurbitae Felt 1911l: 459, nomen nudum. 

Mycodiplosis cucurbitae Felt 1911n: 550. USA; ST; ♀,L; USNM. 

Itonida cucurbitae Felt 1911n: 555. USA; ST; ♂,♀,L; USNM. 

Mycodiplosis impatientis Felt 1908b: 401. USA: New York, West Nyack; HT; ♂; NYSM. 

Mycodiplosis populifolia Felt 1908b: 400. USA: New York, Albany; ST; ♂,♀; NYSM. 

Mycodiplosis reaumuri Kieffer 1901d: 173. France: Bitche; ST; ♂,♀; lost?. 

cylindrica. Argentina (Entre Ríos). Puccinia coronata avenae (Uredinales). 

Mycodiplosis cylindrica Gagné 1987b: 4. Argentina: Entre Ríos, Paraná; HT; ♂; MLPA. 

emarginata. USA (New York). 

Mycodiplosis emarginata Felt 1907a: 33. USA: New York, Nassau; HT; ♂; NYSM. 

erysiphes. Netherlands, Germany. Erysiphe lamprocarpa; Microsphaera sp., Sphaerotheca sp. (Erysiphales). 

Diplosis erysiphes Rübsaamen 1889c: 379. Germany: vic. Siegen; ST; ♂,♀,L; ZMHB. 

explicata. USA (New York). 

Cecidomyia explicata Felt 1908b: 413. USA: New York, Albany; HT; ♂; NYSM. 

*filipendulae. Russia (Far East). Filipendula palmata (Rosaceae). 

Mycodiplosis filipendulae Fedotova 2002c: 26. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*fraxinicola. Russia (Far East). Fraxinus mandshurica (Oleaceae). 

Mycodiplosis fraxinicola Fedotova 2002c: 28. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

fungiperda. USA (District of Columbia). In undet. fungus. 

Mycodiplosis fungiperda Felt 1915f: 407. USA: District of Columbia; ST; ♂,♀; USNM. 

gloeopeniophorae. Germany. Peniophora aurantiaca; P. cinerea (Aphyllophorales). 

Isodiplosis gloeopeniophorae Rübsaamen 1925: 44. Germany: Prignitz, Triglitz; HT; LT; ZMHB. 

glycyrrhizae. Kazakhstan. Glycyrrhiza uralensis (Fabaceae). 

Mycodiplosis glycyrrhizae Fedotova 1985c: 38. Kazakhstan: Zhety-Zhol Mts, Kara-Konyz canyon, 16 km SW 

Masanchi; ♂; ZMAS. 

gymnosporangii. France, Greece. Gymnosporangium clavariaeforme (Uredinales). 

Mycodiplosis gymnosporangii Kieffer 1904a: 73. France: vic. Paris & Alpes-Maritimes; ST; ♂,♀,P,L; lost?. 

hemileiae. Tanzania. Hemileia vastatrix (Uredinales). 

Mycodiplosis hemileiae Barnes 1939f: 330. Tanzania: Bukoba, Kaboia; ST; ♂,♀; BMNH. 

heterosaetosa. Germany. Erysiphe spp., Sphaerotheca spp. (Erysiphales). 

Mycodiplosis heterosaetosa Holz 1970: 79. Germany: Thüringen, Kyffhäuser, Kelbra; HT; LT; SMNS. 

indica. India (Tamil Nadu). A rust (Uredinales). 

Mycodiplosis indica Felt 1920b: 5. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

inimica. UK, Netherlands, Germany, Czechoslovakia, Russia (Europe), USA (New York). On wheat; Plasmopara 

viticola (Peronosporales). 

Cecidomyia inimica Fitch 1861: 830. USA: New York, Salem; ST; ♂,♀,L; USNM. 

Mycodiplosis plasmoparae Rübsaamen 1906c: 234. Germany: Mosel, Lieser; ST; ♀,L; ZMHB. 

Cecidomyia angulata Felt 1907a: 39. USA: New York, Albany; HT; ♂; NYSM. 

Cecidomyia urticae Felt 1907a: 40. USA: New York, Albany; HT; ♂; NYSM. 

Mayetiola socialis Felt 1908b: 369. USA: New York, Albany; HT; ♀; NYSM. 

Isodiplosis involuta Rübsaamen 1910d: 287. Germany: Mosel, Oberbillig; T; ♂; ZMHB. 

Itonida aphidivora Felt 1912l: 245. USA: New York, Nassau; ST; ♂,♀; NYSM. 

Isodiplosis deutera Milne 1960: 100. UK: England, Lincolnshire and Hertfordshire; ST; ♂,♀; BMNH. 

isosaetosa. Germany. Erysiphe spp., Sphaerotheca spp. (Erysiphales). 

Mycodiplosis isosaetosa Holz 1970: 73. Germany: Wittenberg, Lorch; HT; ♂; SMNS.

303 

kittabhakshi. India (Uttar Pradesh). 

Mycodiplosis kittabhakshi Grover 1970a: 1. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

kraussei. Poland, Czechoslovakia. 

Itonida kraussei Wolff 1910: 410. Poland: Bydgosz; ST; ♂,♀,P,L; ?. 

ligulata. Costa Rica. Adult assoc. with flowers of Theobroma cacao (Sterculiaceae). 

Mycodiplosis ligulata Gagné 1984c: 124. Costa Rica: Limón Prov., nr Siquirres, Finca La Lola; HT; ♂; USNM. 

*ligustrinaphila. Russia (Far East). Ligustrina amurensis (Oleaceae) 

Mycodiplosis ligustrinaphila Fedotova 2004e: 15. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

limbata. Germany. Melampsora salicina (Uredinales). 

Cecidomyia limbata Winnertz 1853: 266. Germany; ST; ♀; lost. 

longiterminalis. India (Uttar Pradesh). 

Mycodiplosis longiterminalis Grover & Bakhshi 1978: 101. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; 

PGCA. 

*loniceracarpae. Russia (Far East). Lonicera maackii (Caprifoliaceae). 

Mycodiplosis loniceracarpae Fedotova 2003c: 9. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

melampsorae. Denmark, Germany, Czechoslovakia, Austria, Romania. Melampsora salicina (Uredinales); several 

genera of rusts (Uredinales). 

Diplosis melampsorae Rübsaamen 1889a: 46. Germany: Siegen; ST; ♀,L; ZMHB. 

Mycodiplosis jaapi Rübsaamen 1925: 44. Germany: Siegen; HT; LT; ZMHB. 

nitrariae. Kazakhstan. A rust (Uredinales). 

Mycodiplosis nitrariae Fedotova 1993e: 90. Kazakhstan: SW shore L. Balkhash, 8 km S Chiganak; HT; ♂; ZMAS. 

oidii. UK. Oidium sp.; Erysiphe sp., Microsphaera sp., Sphaerotheca sp. (Erysiphales). 

Cecidomyia oidii Hardy 1854f: 170. UK: Scotland, vic. Penmanshiel; T; LT; lost. 

padi. Russia (Europe). Ex galls of Eriophyes padi (Acarina: Eriophyidae). 

Mycodiplosis padi Mamaeva 1964a: 210. Russia: Pavlovskaja Sloboda; HT; ♂; MAMC. 

pritchardi. USA (California). Puccinia evadens (Uredinales). 

Clinodiplosis pucciniae Pritchard 1948b: 29, jun. sec. hom. pucciniae Rübsaamen. USA: California, Palo Alto; HT; 


Mycodiplosis pritchardi Gagné 2004b: 204, new name for Clinodiplosis pucciniae Pritchard. 

*pteridiis. Russia (Far East). Pteridium aquilinum (Polypodiaceae). 

Mycodiplosis pteridiis Fedotova 2003c: 11. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

pucciniacola. Hawaiian Is. Puccinia sp. (Uredinales). 

Nanodiplosis pucciniacola Hardy 1960: 298. Hawaiian Is: Oahu, Kailua; HT; ♂; BPBM. 

pucciniae. UK, Germany, Latvia, Russia (Europe). Puccinia compositarum; Melampsora spp., Puccinia spp. 

(Uredinales). 

Diplosis pucciniae Rübsaamen 1889c: 382. Germany; ST; ♀,L; ZMHB. 

pulsatillae. France. Pulsatilla vulgaris (Ranunculaceae). 

Diplosis pulsatillae Kieffer 1888c: 262. France: Bitche; ST; ♂,♀,L; lost?. 

rotundata. USA (New York). 

Mycodiplosis rotundata Felt 1908b: 401. USA: New York, Newport; ST; ♂; NYSM. 

rubida. Jamaica, St. Vincent, Brazil, Hawaiian Is. Uromyces pisi; Puccinia sp. (Uredinales). Ref.: Parnell 1969 (biol., 

anat.). 

Toxomyia rubida Felt 1911i: 194. St. Vincent; ST; ♂,♀; NYSM. 

Toxomyia fungicola Felt 1911j: 302. St. Vincent; ST; ♂,♀,L; NYSM. This species was treated as senior synonym of 

rubida in Parnell 1969: 121 but the description of rubida appeared earlier. 

saundersi. UK, Czechoslovakia. A rust (Uredinales). 

Mycodiplossi saundersi Barnes 1927h: 220. UK: vic. Cambridge; ST; ♂,♀; BMNH. 

sesami. India (Maharashtra). On buds Sesamum indicum (Pedaliaceae). 

Clinodiplosis sesami Rao & Sharma 1981: 26. India: Maharashtra, Aurangabad; HT; ♂; RCMU.

304 

simulacri. Sri Lanka. 

Mycodiplosis simulacri Felt 1915g: 177. Sri Lanka: Peradeniya, Botanic Gardens; HT; ♂; NYSM. 

siphae. Italy. Host given as Sipha maydis (Hemiptera: Aphididae), but M. siphae is presumably a mycophage. 

Trilobiella siphae Del Guercio 1918a: 244. Italy; T; LT; lost. 

sphaerothecae. UK, Netherlands, Germany. Sphaerotheca humuli; Erysiphe sp., Microsphaera sp., Sphaerotheca sp. 

(Erysiphales. 

Diplosis sphaerothecae Rübsaamen 1889c: 381. Germany: vic. Siegen; ST; ♂,♀,L; ZMHB. 

tianschanica. Kazakhstan. Uromyces polygoni (Uredinales). 

Mycodiplosis tianschanicus Fedotova 1997g: 883. Kazakhstan: Zailiïskiï Alatau, 20 km S Alma-Ata; HT; ♂; ZMAS. 

tremulae. UK, France. Melampsora tremulae (Uredinales). 

Mycodiplosis tremulae Kieffer 1895p: 9. France: Moselle; T; A; lost?. 

triticina. Egypt, Kenya. Puccinia triticina (Uredinales). 

Hyperdiplosis triticina Barnes 1936: 275. Kenya; ST; ♂,♀; BMNH. 

tussilaginis. France. Aecidium tussilaginis (Uredinales). 

Mycodiplosis tussilaginis (as sussilaginis) Kieffer 1895p: 9. France: Moselle; T; A; lost?. This species was originally 

indicated as coming from tussilaginis so the original spelling of sussilaginis was an inadvertent error. 

MYRCIAMYIA 

Myrciamyia Maia 1996a: 567. Type species, maricaensis Maia (orig. des.). 

Neotropical; 1 sp.; Cecidomyiidi; Lopesiini. 

maricaensis. Brazil (Rio de Janeiro). Myrcia ovata (Myrtaceae). 

Myrciamyia maricaensis Maia 1996a: 569. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

MYRCIARIAMYIA 

Myrciariamyia Maia 1995: 635. Type species, bivalva Maia (orig. des. as n. g., n. sp.). 

Neotropical; 3 spp.; Lasiopteridi, unplaced to tribe. 

*admirabilis. Brazil (Minas Gerais). Erythroxylum suberosum (Erythroxylaceae). 

Myrciariamyia admirabilis Maia 2007b: 42. Brazil: Minas Gerais, Serra de São José, Tiradentes; HT; ♂; QBUM. 

bivalva. Brazil (Rio de Janeiro). Myrciaria floribunda (Myrtaceae). 

Myrciariamyia bivalva Maia 1995: 637. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

*fernandesi. Brazil (Minas Gerais). Myrciaria tenella (Myrtaceae). 

Myrciaramyia fernandesi Maia 2004a: 79. Brazil: Minas Gerais, Tiradentes, Serra de São José; HT; ♂; MNRJ. 

MYRICOMYIA 

Myricomyia Kieffer 1901b: 470. Type species, Diplosis mediterranea Loew (mon.). 


mediterranea. UK, circum-Mediterranean. Erica arborea, E. carnea, E. mediterranea, E. scoparia (Ericaceae). Ref.: 

Skuhravá & Skuhravý 2003: 18 (illus.). 

Diplosis mediterranea Löw 1885b: 485. Spain, France, Italy, Croatia; ST; ♂,♀,P,L; NHMW. 

Myricomyia longipalpis Kieffer 1913a: 232. Portugal; T; ♀; lost?. 

NANODIPLOSIS

Nanodiplosis Kieffer 1913g: 55. Type species, Lestodiplosis squamosus Kieffer (orig. des.). 


squamosus. Seychelles Is. 

Lestodiplosis squamosus Kieffer 1911: 323. Seychelles Is: Mahé, Cascade Estate; HT; ♂; BMNH?. 

NEOBAEZOMYIA 

Neobaeomyza Brèthes 1914: 151. Type species, maculata Brèthes (orig. des.). 


maculata. Argentina (Buenos Aires). 

Neobaeomyza maculata Brèthes 1914: 151. Argentina: Buenos Aires, General Urquiza; HT; ♀; MACN. 

NEODASYNEURIOLA 

Neodasyneuriola Marikovskij 1958b: 1850. Type species, tuberculifera Marikovskij (orig. des.). 


tuberculifera. Kazakhstan. 

Neodasyneuriola tuberculifera Marikovskij 1958b: 1850. Kazakhstan: southern slope Kungeï Alatau; HT; ♂; MARC. 

NEOISODIPLOSIS 

Neoisodiplosis Holz 1970: 116. Type species, Isodiplosis corticii Rübsaamen (orig. des.). 

Western Palearctic; 2 spp., fungivorous; Cecidomyiidi, Mycodiplosini. 

corticii. Germany. Corticium lacteum; Peniophora aurantiaca, P. cinerea (Aphylloporales). 

Isodiplosis corticii Rübsaamen 1925: 44. Germany: Prignitz, Triglitz; HT; LT; ZMHB. 

longisaetosa. Germany. Peniophora cinerea (Aphylloporales). 

Neoisodiplosis longisaetosa Holz 1970: 123. Germany: Prignitz, Triglitz; HT; LT; SMNS. 

NEOLASIOPTERA 

Neolasioptera Felt 1908b: 330. Type species, Lasioptera vitinea Felt (subs. des. Coquillett 1910: 575). 

Luisieria Tavares 1922a: 43. Type species, fariae Tavares (orig. des.). 

Dilasioptera Möhn 1964b: 570, as subg. of Neolasioptera. Type species, Neolasioptera serrata Möhn (orig. des.). 

Neurolasioptera Brèthes 1922: 138. Type species, baezi Brèthes (mon.). 

Physalidicola Brèthes 1917a: 240. Type species, argentata Brèthes (orig. des.). 

Nearctic, Neotropical; 134 spp.; Lasiopteridi, Alycaulini. These form mainly stem galls, but also petiole, leaf vein, or 

flower galls; most live on Asteraceae, but many other plant families are hosts also. Host specificity usually seems close, 

even to cases of two distinct species on the same host (Plakidas 1994), although there are examples of species, e.g., N. 

hibisci, with hosts in different families. Ref.: Gagné 1994a (review). 

aeschynomensis. Argentina (Buenos Aires). Aeschynomene montevidensis (Fabaceae). 

Lasioptera aeschynomensis Brèthes 1918: 312. Argentina: Buenos Aires, Buenos Aires; ST; ♀; MACN. 

albipes. USA (New Jersey). 

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306 

Neolasioptera albipes Felt 1918a: 190. USA: New Jersey, Riverton; HT; ♀; NYSM. 

albitarsis. USA (New York). Aster cordifolius (Asteraceae). 

Choristoneura albitarsis Felt 1907c: 153. USA: New York, Nassau; ST; ♂,♀; NYSM. 

albolineata. USA (New York). 

Neolasioptera albolineata Felt 1908b: 332. USA: New York, Albany; HT; ♀; NYSM. 

allioniae. USA (Colorado, Texas). Allionia sp.; A. nyctaginae (Nyctaginaceae). 

Lasioptera allioniae Felt 1911m: 482. USA: Colorado, Boulder; ST; ♂,♀; NYSM. 

amaranthi. El Salvador. Amaranthus spinosus (Amaranthaceae). 

Neolasioptera amaranthi Möhn 1964a: 47. El Salvador: San Salvador; HT; ♂; SMNS. 

ambrosiae. Widespread Nearctic. Ambrosia trifida; Ambrosia spp. (Asteraceae). 

Neolasioptera ambrosiae Felt 1909b: 288. USA: New York, Ithaca; ST; ♂,♀; NYSM. 

angelicae. USA (New Jersey). Angelica villosa (Apiaceae). 

Cecidomyia angelicae Beutenmüller 1908: 74. USA: New Jersey, Middlesex Co.; T; LT; AMNH. 

aphelandrae. El Salvador. Aphelandra deppeana (Acanthaceae). 

Neolasioptera aphelandrae Möhn 1964a: 90. El Salvador: La Libertad, N El Cimarrón; HT; LT; SMNS. 

apocyni. USA (New Jersey). Apocynum androsaemifolium (Apocynaceae). 

Lasioptera apocyni Felt 1921e: 149. USA: New Jersey, Trenton; HT; ♀; NYSM. 

argentata. Argentina (Buenos Aires). Physalis viscosa (Solanaceae). 

Physalidicola argentata Brèthes 1917a: 240. Argentina: Buenos Aires, vic. Buenos Aires; ST; A; MACN. 

argentisquamae. USA (New York). Solidago sp. (Asteraceae). 

Lasioptera argentisquamae Felt 1908b: 324. USA: New York, West Nyack; HT; ♀; NYSM. 

asclepiae. USA (New York). Asclepias incarnata (Asclepiadaceae). 

Neolasioptera asclepiae Felt 1908b: 332. USA: New York, Albany; HT; ♀; NYSM. 

baccharicola. USA (Virginia). Baccharis halimifolia (Asteraceae). 

Neolasioptera baccharicola Gagné 1971a: 156. USA: Virginia, Virginia Beach; HT; ♂; USNM. 

baezi. Argentina (Entre Ríos). Teucrium inflatum (Lamiaceae). 

Neurolasioptera baezi Brèthes 1922: 138. Argentina: Entre Ríos, Paraná; ST; ♂,♀; MACN. 

boehmeriae. Canada (Ontario), USA (New York, New Jersey, Maryland). Boehmeria cylindrica (Urticaceae). 

Cecidomyia boehmeriae Beutenmüller 1908: 74. USA: New York, Shushank, & New Jersey, Fort Lee; T; LT; 

AMNH. 

borreriae. El Salvador. Borreria verticillata (Rubiaceae). 

Neolasioptera borreriae Möhn 1964a: 93. El Salvador: San Salvador, E San Marcos; HT; ♀; SMNS. 

brevis. USA (Missouri, Michigan, Pennsylvania). Gleditsia triacanthos (Fabaceae). 

Neolasioptera brevis Gagné 1984b: 544. USA: Pennsylvania, Dauphin Co., Harrisburg, E. Harrisburg Cemetery; HT; 

♀; USNM. 

brickelliae. El Salvador. Brickellia pacayensis (Asteraceae). 

Neolasioptera brickelliae Möhn 1964b: 560. El Salvador: Chalatenango, ca 15 km from la Palma; HT; ♀; SMNS. 

caleae. El Salvador. Calea integrifolia (Asteraceae). 

Neolasioptera caleae Möhn 1964b: 556. El Salvador: La Libertad, nr Los Chorros; HT; ♂; SMNS. 

camarae. El Salvador. Lantana camara (Verbenaceae). 

Neolasioptera camarae Möhn 1964b: 565. El Salvador: San Salvador; HT; ♂; SMNS. 

capsici. El Salvador. Capsicum baccatum (Solanaceae). 

Neolasioptera capsici Möhn 1964a: 78. El Salvador: La Libertad, nr San Diego, E La Libertad; HT; ♀; SMNS. 

cassiae. USA (Arizona). Cassia nictitans (Fabaceae). 

Lasioptera cassiae Felt 1909b: 287. USA: Arizona, Fort Huachuca; ST; ♂,♀; USNM. 

caulicola. USA (New York). Lonicera sp. (Caprifoliaceae). 

Lasioptera caulicola Felt 1907c: 162. USA: New York, vic. Albany; HT; ♀; NYSM. 

celastri. USA (New York). 

Neolasioptera celastri Felt 1908b: 330. USA: New York, Albany; HT; ♀; NYSM. 

celtis. El Salvador. Celtis iguanea (Ulmaceae). 

Neolasioptera celtis Möhn 1964b: 553. El Salvador: E La Libertad, nr San Diego; HT; ♂; SMNS. 

cerei. Brazil (Rio de Janeiro). Selenicereus setaceus (Cactaceae).

307 

Lasioptera cerei Rübsaamen 1905a: 81. Brazil: Rio de Janeiro, Cabo Frio; ST; F,P,L; ZMHB. Ref.: Maia 1999b 

(redescr.). 

cestri. El Salvador. Cestrum aurantiacum; C. lanatum (Solanaceae). 

Neolasioptera cestri Möhn 1964a: 80. El Salvador: Santa Ana, nr San Jose, Hacienda San Jose; HT; ♂; SMNS. 

cimmaronensis. El Salvador. Undet. sp. of Vernonieae (Asteraceae). 

Neolasioptera cimmaronensis Möhn 1975: 76. El Salvador: La Libertad, N El Cimarrón; HT; ♀; SMNS. 

cinerea. USA (New York). 

Choristoneura cinerea Felt 1907a: 15. USA: New York, Albany; HT; ♀; NYSM. 

cissampeli. El Salvador. Cissampelos pareira (Menispermaceae). 

Neolasioptera cissampeli Möhn 1964a: 55. El Salvador: San Salvador; HT; ♀; SMNS. 

clematicola. El Salvador. Clematis dioica (Ranunculaceae). 

Neolasioptera clematicola Möhn 1964a: 52. El Salvador: Santa Ana, NW Ciudad Arce; HT; ♂; SMNS. 

clematidis. USA (New York, Massachusetts). Clematis virginiana (Ranunculaceae). 

Choristoneura clematidis Felt 1907d: 6. USA: New York, Newport; HT; ♀; NYSM. 

combreti. El Salvador. Combretum farinosum (Combretaceae). 

Neolasioptera combreti Möhn 1964a: 68. El Salvador: La Libertad, N El Cimarrón; HT; ♂; SMNS. 

compositarum. El Salvador. Tridax procumbens; Hymenostephium cordatum (Asteraceae). 

Neolasioptera compositarum Möhn 1964b: 563. El Salvador: San Salvador; HT; ♂; SMNS. 

convolvuli. USA (Nebraska, New York, Massachusetts, Maryland). Convolvulus sepium (Convolvulaceae). 

Lasioptera convolvuli Felt 1907c: 149. USA: New York, Bath; ST; ♂,♀; NYSM. 

cordiae. El Salvador. Cordia cana (Boraginaceae). 

Neolasioptera cordiae Möhn 1964a: 71. El Salvador: La Libertad, SE Ciudad Arce; HT; ♀; SMNS. 

cornicola. USA (New York, Massachusetts, Rhode Island, Maryland). Cornus stolonifera (Cornaceae). 

Lasioptera cornicola Beutenmüller 1907a: 394. USA: New York, New York, Bronx Pk & Staten I.; ST; ♂,♀,L; 

AMNH. 

crotalariae. USA (Massachusetts). Crotalaria sagittalis (Fabaceae). 

Cecidomyia crotalariae Stebbins 1910: 40. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

cruttwellae. Colombia, Trinidad. Chromolaena odorata (Asteraceae). 

Neolasioptera cruttwellae Gagné 1977b: 115. Trinidad: Simla; HT; ♀; USNM. 

Neolasioptera odorati Wünsch 1979: 100. Colombia: Magdalena, Cañaverales; HT; ♂; SMNS. 

cupheae. Brazil (Rio de Janeiro). Cuphea carthagenensis (Lythraceae). 

Neolasioptera cupheae Gagné 1998: 522. Brazil: Rio de Janeiro, Silva Jardim; HT; ♂; QBUM. 

cusani. Colombia. Undet. sp. of Amaranthaceae. 

Neolasioptera cusani Wünsch 1979: 104. Colombia: Magdalena, Donama; HT; LT; SMNS. 

dentata. El Salvador. Eupatorium sp. (Asteraceae). 

Neolasioptera dentata Möhn 1964b: 567. El Salvador: Santa Ana, nr San Jose; HT; LT; SMNS. 

desmodii. USA (New York, New Jersey, Maryland). Desmodium cuspidatum or D. canadense; Desmodium spp. 

(Fabaceae). 

Lasioptera desmodii Felt 1907a: 10. USA: New York, Albany; ST; ♂,♀; NYSM. 

diclipterae. Colombia. Dicliptera assurgens (Acanthaceae). 

Neolasioptera diclipterae Wünsch 1979: 78. Colombia: Magdalena, Rio Manzanares; HT; ♂; SMNS. 

diplaci. USA (California). Mimulus longiflorus (Scrophulariaceae). 

Lasioptera diplaci Felt 1912h: 151. USA: California, Whittier, Puente Hills; ST; F,L; NYSM. 

donamae. Colombia. Iresine angustifolia (Amaranthaceae). 

Neolasioptera donamae Wünsch 1979: 97. Colombia: Magdalena, Donama; HT; LT; SMNS. 

eregeroni. Widespread eastern Nearctic. Conyza canadensis (Asteraceae). 

Diplosis eregeroni Brodie 1894a: 13. Canada: Ontario, Toronto; ST; A,L; USNM. 

Diplosis erigerontis Aldrich 1905: 162, error for eregeroni. 

Cecidomyia erigeroni Felt 1925c: 54, missp. of eregeroni. 

erigerontis. Widespread Nearctic, El Salvador. Conyza canadensis (Asteraceae). 

Choristoneura erigerontis Felt 1907c: 163. USA: New York, Albany; ST; ♂,♀; NYSM. 

Lasioptera podagrae Beutenmüller 1913: 414. USA: New York, New York; New Jersey, Fort Lee; ST; ♂,♀;

308 

AMNH. 

erythroxyli. El Salvador. Erythroxylon mexicanum (Erythroxylaceae). 

Neolasioptera erythroxyli Möhn 1964b: 62. El Salvador: Sonsonate, N Los Cóbanos; HT; ♀; SMNS. 

eugeniae. Brazil (Rio de Janeiro). Eugenia uniflora (Myrtaceae). 

Neolasioptera eugeniae Maia 1993: 718. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

eupatoriflorae. USA (New York, Massachusetts, Maryland). Eupatorium perfoliatum; E. hyssopifolium (Asteraceae). 

Lasioptera eupatoriflorae Felt 1907d: 6. USA: New York, Karner; ST; ♂,♀; NYSM. 

eupatorii. USA (New York). Ageratina altissima (Asteraceae). 

Choristoneura eupatorii Felt 1907c: 154. USA: New York, New York, Staten I.; ST; ♂,♀; NYSM. 

exeupatorii. Cuba. Eupatorium villosum (Asteraceae). 

Cecidomyia eupatorii Cook 1909: 144, jun. sec. hom. eupatorii Felt. Cuba; T; G; ?. 

Neolasioptera exeupatorii Gagné. 1994a: 73, new name for Cecidomyia eupatorii Cook. 

exigua. El Salvador. Solanum umbellatum (Solanaceae). 

Neolasioptera exigua Möhn 1964a: 82. El Salvador: La Libertad, N El Cimarrón; HT; ♀; SMNS. 

fariae. Brazil (Bahia). Ex galls of unkn. plant. 

Luisieria fariae Tavares 1922a: 44. Brazil: Bahia, vic. Salvador; ST; ♂,♀,P,L; TAVC. 

farinosa. Widespread eastern Nearctic. Rubus sp. (Rosaceae). 

Cecidomyia farinosa Osten Sacken 1862: 204. USA: DC; T; LT; lost. 

flavomaculata. USA (New York). 

Neolasioptera flavomaculata Felt 1908b: 332. USA: New York, Albany; HT; ♀; NYSM. 

fontagrensis. USA (New York, Massachusetts). Celastrus scandens (Celastraceae). 

Cecidomyia celastri Stebbins 1910: 41, jun. sec. hom. celastri Felt. USA: Massachusetts, vic. Springfield; T; G; 

NHSM. 

Neolasioptera fontagrensis Gagné 1989b: 173, new name for Cecidomyia celastri Stebbins. 

fraxinifolia. USA (New York). Fraxinus sp. (Oleaceae). 

Lasioptera fraxinifolia Felt 1908b: 327. USA: New York, New York; HT; ♂; NYSM. 

frugivora. Trinidad. Chromolaena odorata (Asteraceae). 

Neolasioptera frugivora Gagné 1977b: 116. Trinidad: Simla; HT; ♀; USNM. 

galeopsidis. USA (Massachusetts). Galeopsis tetrahit (Lamiaceae). 

Lasioptera galeopsidis Felt 1909b: 287. USA: Massachusetts, Boston; ST; ♂,♀; NYSM. The host is an alien sp., so, 

assuming the original plant identification was correct, N. galeopsidis shifted to it from a native host. 

grandis. El Salvador. Pseudocalymma macrocarpa (Bignoniaceae). 

Neolasioptera grandis Möhn 1964a: 87. El Salvador: San Vicente, km 81 nr Rio San Felipe; HT; LT; SMNS. 

hamamelidis. USA (New York). 

Choristoneura hamamelidis Felt 1907a: 15. USA: New York, Albany; HT; ♀; NYSM. 

helianthi. Widespread eastern Nearctic. Helianthus strumosus; Helianthus spp. (Asteraceae). 

Choristoneura helianthi Felt 1907d: 7. USA: New York, Highland; HT; ♀; NYSM. 

Lasioptera murtfeldtiana Felt 1909b: 288. USA: Missouri, Kirkwood; ST; ♂; USNM. 

heliocarpi. El Salvador. Heliocarpus donnellsmithii (Tiliaceae). 

Neolasioptera heliocarpi Möhn 1964a: 66. El Salvador: La Libertad, E San Juan Higenio; HT; LT; SMNS. 

heterothalami. Argentina (Mendoza, San Juan). Baccharis artemisioides, B. spartioides (Asteraceae). 

Lasioptera heterothalami Kieffer & Jörgensen 1910: 399. Argentina: Mendoza, La Paz, & San Juan, Cancete; ST; 

♂,♀,P,L,E; lost. 

hibisci. USA (New York, North Carolina, Louisiana), Honduras. Hibiscus moscheutos (Malvaceae); Cucumis melo 

(Cucurbitaceae). 

Choristoneura hibisci Felt 1907c: 155. USA: New York, New York, Staten I.; ST; ♂,♀; NYSM. 

hirsuta. USA (Arizona). 

Neolasioptera hirsuta Felt 1908b: 331. USA: Arizona, Fort Huachuca; HT; ♀; USNM. 

hyptis. El Salvador. Hyptis suaveolens (Lamiaceae). 

Neolasioptera hyptis Möhn 1964a: 74. El Salvador: Santa Ana, NW Ciudad Arce; HT; ♂; SMNS. 

impatientifolia. Widespread eastern Nearctic. Impatiens capensis; I. pallida (Balsaminaceae). 

Lasioptera impatientifolia Felt 1907a: 9. USA: New York, Albany; ST; ♂,♀; NYSM. Shinji 1938g recorded this

309 

species from Japan (as both Lasioptera and Cecidomyia impatientis O.S.), but the indication in Shinji 1938n is 

that the Japanese species is a Lasioptera and not a Neolasioptera. 

Cecidomyia fulva Beutenmüller 1908: 75. USA: New York, Shushank, & New Jersey, Fort Lee; T; LT; AMNH. 

incisa. USA (Pennsylvania, Maryland, Virginia). Verbesina alternifolia (Asteraceae). 

Neolasioptera incisa Plakidas 1994: 214. USA: Pennsylvania, Allegheny Co., 7 km E Ingomar; HT; ♀; USNM. 

indigoferae. El Salvador. Indigofera suffruticosa (Fabaceae). 

Neolasioptera indigoferae Möhn 1964a: 60. El Salvador: San Salvador; HT; ♂; SMNS. 

ingae. El Salvador. Inga spuria; I. leptoloba (Fabaceae). 

Neolasioptera ingae Möhn 1964a: 57. El Salvador: La Libertad, Cumbre, SE San Julian, Hacienda San Julian; HT; 

♀; SMNS. 

iresinis. El Salvador. Iresine celosia (Amaranthaceae). 

Neolasioptera iresinis Möhn 1964a: 50. El Salvador: San Salvador; HT; ♂; SMNS. 

lantanae. Brazil (Bahia). Lantana sp. (Verbenaceae). 

Luisieria lantanae Tavares 1922a: 48. Brazil: Bahia, vic. Salvador; T; LT; TAVC. 

lapalmae. El Salvador. Calea zacatechichi (Asteraceae). 

Neolasioptera lapalmae Möhn 1964a: 94. El Salvador: Chalatenango, SE La Palma; HT; ♂; SMNS. 

lathami. USA (Arizona & widespread eastern). Baccharis halimifolia; Baccharis spp. (Asteraceae). 

Neolasioptera lathami Gagné 1971a: 154. USA: South Carolina, Yemassee; HT; ♂; USNM. 

linderae. USA (New York, Virginia). Lindera benzoin (Lauraceae). 

Lasioptera linderae Beutenmüller 1907a: 398. USA: New York, New York, Bronx; T; LT; AMNH. 

lupini. USA (Washington, California). Lupinus sp.; L. arboreus (Fabaceae). 

Lasioptera lupini Felt 1908b: 326. USA: California; HT; ♀; USNM. 

lycopi. USA (New York, Massachusetts, Maryland). Lycopus communis; L. uniflorus (Lamiaceae). 

Lasioptera lycopi Felt 1907c: 152. USA: New York, vic. Albany; HT; ♀; NYSM. 

major. USA (Colorado). 

Neolasioptera major Felt 1918a: 175. USA: Colorado, Colorado Springs; HT; ♀; NYSM. 

malvavisci. El Salvador. Malvaviscus arboreus (Malvaceae). 

Neolasioptera malvavisci Möhn 1975: 72. El Salvador: La Libertad, N El Cimarrón; HT; LT; SMNS. 

martelli. Mexico (Distrito Federal). Agave sp. (Agavaceae). 

Neolasioptera martelli Nijveldt 1967b: 125. Mexico: DF, Chapingo; HT; ♂; ZMAN. 

melantherae. El Salvador. Melanthera nivea (Asteraceae). 

Neolasioptera melantherae Möhn 1964a: 97. El Salvador: Sonsonate, NW Salcoatitlán; HT; LT; SMNS. 

menthae. USA (Illinois). Mentha sp. (Lamiaceae). 

Neolasioptera menthae Felt 1909b: 288. USA: Illinois, Chicago; ST; ♂,♀; USNM. 

merremiae. El Salvador. Merremia umbellata (Convolvulaceae). 

Neolasioptera merremiae Möhn 1964a: 70. El Salvador: La Libertad, nr . San Diego, E La Libertad; HT; LT; SMNS. 

mimuli. USA (California). Mimulus "glutinosus" (Scrophulariaceae). 

Neolasioptera mimuli Felt 1908b: 332. USA: California, Alameda; HT; ♀; USNM. 

mincae. Colombia. Melanthera aspera (Asteraceae). 

Neolasioptera mincae Wünsch 1979: 95. Colombia: Magdalena, Minca; HT; LT; SMNS. 

mitchellae. USA (District of Columbia). Lycopus virginicus (Lamiaceae). 

Lasioptera mitchellae Felt 1908b: 325. USA: District of Columbia; ST; ♂,♀; USNM. 

monardi. Canada (Ontario). Monarda fistulosa (Lamiaceae). 

Diplosis monardi Brodie 1894d: 109. Canada: Ontario, Toronto; ST; A,L; USNM. 

monticola. Chile. Gymnophyton polycephalum (Apiaceae). 

Lasioptera monticola Kieffer & Herbst 1909: 124. Chile: Santiago, Pennalolen; ST; ♂,♀,P,L; lost. 

neofusca. USA (New York). 

Lasioptera neofusca Felt 1908b: 327. USA: New York, Albany; HT; ♀; NYSM. 

nodulosa. Widespread eastern Nearctic. Rubus "villosus"; Rubus spp. (Rosaceae). 

Lasioptera nodulosa Beutenmüller 1907a: 397. USA: Ohio, New York, New York, & New Jersey; ST; F,L; AMNH. 

odontonemae. El Salvador. Odontonema stricta (Acanthaceae). 

Neolasioptera odontonemae Möhn 1964a: 91. El Salvador: nr Sonsonate, betw. Armenia & Izalco; HT; ♂; SMNS.

310 

olivae. Colombia. Calea caracasana (Asteraceae). 

Neolasioptera olivae Wünsch 1979: 92. Colombia: Magdalena, Donama; HT; ♀; SMNS. 

palmeri. USA (New Mexico). Amaranthus palmeri (Amaranthaceae). 

Cecidomyia palmeri Felt 1912k: 757. USA: New Mexico, Mesilla Park; HT; LT; USNM. 


Lasioptera palustris Felt 1907c: 162. USA: New York, West Nyack; ST; ♂,♀; NYSM. 

parvula. El Salvador. Calea integrifolia (Asteraceae). 

Neolasioptera parvula Möhn 1964b: 558. El Salvador: La Libertad, Volcán Boquerón; HT; ♂; SMNS. 

perfoliata. USA (northeastern). Eupatorium perfoliatum; E. hyssopifolium (Asteraceae). 

Choristoneura perfoliata Felt 1907c: 156. USA: Massachusetts, Worcester; ST; ♂,♀; NYSM. 

phaseoli. El Salvador. Phaseolus sp. (Fabaceae). 

Neolasioptera phaseoli Möhn 1975: 71. El Salvador: Santa Ana, nr San José, Hacienda San José; HT; ♀; SMNS. 

pierrei. USA (New York, Massachusetts). Sambucus canadensis (Caprifoliaceae). 

Cecidomyia sambuci Felt 1906b: 131, jun. prim. hom. sambuci Kaltenbach. USA: New York, Nassau; HT; LT; 

NYSM. 

Neolasioptera pierrei Gagné 1972b: 323, new name for Cecidomyia sambuci Felt. 

piriqueta. Puerto Rico. Piriqueta ovata (Turneraceae). 

Lasioptera piriqueta Felt 1917b: 193. Puerto Rico: Mayaguez; ST; ♂,♀,P; NYSM. 

portulacae. Circumcaribbean. Portulaca oleracea (Portulacaceae). 

Cecidomyia portulacae Cook 1906: 251. Cuba; T; G; ?. 

Cecidomyia portulacae Cook 1909: 145. Cuba; T; G; ?. 

Lasioptera portulacae Felt 1911f: 84. St. Vincent; ST; ♂,♀,P,L; USNM. 

potentillaecaulis. USA (Massachusetts). Potentilla canadensis (Rosaceae). 

Cecidomyia potentillaecaulis Stebbins 1910: 37. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

pseudocalymmae. El Salvador. Pseudocalymma macrocarpa (Bignoniaceae). 

Neolasioptera pseudocalymmae Möhn 1964a: 84. El Salvador: La Libertad, nr San Diego, E La Libertad; HT; ♂; 

SMNS. 

punicei. Canada (Ontario). Aster puniceus (Asteraceae). 

Diplosis punicei Brodie 1909a: 159. Canada: Ontario, Toronto; ST; A,L; USNM. 


Lasioptera quercina Felt 1907a: 12. USA: New York, Albany; HT; ♀; NYSM. 

*ramicola. Brazil (Rio Grande do Sul). Physalis angulata (Solanaceae). 

Neolasioptera ramicola Maia 2009b: 163. Brazil: Rio Grande do Sul, Bento Gonçalves; HT; ♂; QBUM. 

ramuscula. USA (widespread eastern). Aster dumosus, A. patens (Asteraceae). 

Cecidomyia ramuscula Beutenmüller 1907a: 392. USA: North Carolina, Buncombe Co., Valley of the Black Mts; 

ST; LT; AMNH. 

rostrata. USA (New Mexico, Texas, Maryland, Florida), Dominican Republic, Argentina. Baccharis glomeruliflora; 

Baccharis spp. (Asteraceae). 

Neolasioptera rostrata Gagné 1989a: 170. USA: Florida, Miami, Dodge I.; HT; ♂; USNM. 

rudbeckiae. USA (New York, Pennsylvania, Virginia). Rudbeckia laciniata (Asteraceae). 

Lasioptera rudbeckiae Felt 1908b: 324. USA: Pennsylvania, Highspire; HT; ♀; NYSM. 

salvadorensis. El Salvador. Pithecoctenium echinatum (Bignoniaceae). 

Neolasioptera salvadorensis Möhn 1964a: 88. El Salvador: San Miguel, S El Delirio; HT; ♂; SMNS. 

salviae. El Salvador. Salvia occidentalis (Lamiaceae). 

Neolasioptera salviae Möhn 1964a: 76. El Salvador: San Salvador; HT; ♀; SMNS. 

samariae. Colombia. Adenocalymma dugandi (Bignoniaceae). 

Neolasioptera samariae Wünsch 1979: 89. Colombia: Magdalena, Santa Marta; HT; ♂; SMNS. 

senecionis. El Salvador. Senecio kermesinus (Asteraceae). 

Neolasioptera senecionis Möhn 1964a: 98. El Salvador: La Libertad, N El Cimarrón; HT; ♂; SMNS. 

serjaniae. El Salvador. Serjania racemosa (Sapindaceae). 

Neolasioptera serjaniae Möhn 1964a: 64. El Salvador: La Libertad, NW Quezaltepeque, La Toma forest; HT; ♂; 

SMNS.

311 

serrata. El Salvador. Calea zacatechichi (Asteraceae). 

Neolasioptera serrata Möhn 1964b: 571. El Salvador: Chalatenango, SE La Palma; HT; ♀; SMNS. 

sidae. El Salvador. Heliocarpus donnellsmithii (Tiliaceae). 

Neolasioptera sidae Möhn 1964a: 67. El Salvador: Chalatenango, nr Citalá on bank Lempa R.; HT; LT; SMNS. 

spinulae. USA (Arizona). 

Lasioptera spinulae Felt 1908b: 325. USA: Arizona, Huachuca; ST; ♂,♀; USNM. 

tertia. USA (New Mexico). Undet. sp. of Asteraceae. 

Lasioptera tertia Cockerell 1898: 328. USA: New Mexico, Paraje; HT; ♀; USNM. 

thurstoni. Nearctic (widespread). Helianthus divaricatus; Helianthus spp. (Asteraceae). 

Diplosis thurstoni Brodie 1894c: 73. Canada: Ontario, Toronto; ST; ♂,♀,L; USNM. 

Lasioptera weldi Felt 1908b: 326. USA: Illinois, Evanston; ST; ♂,♀; NYSM. 

Neolasioptera trimera Felt 1911m: 484. USA: Arizona, Fort Grant; HT; ♀; USNM. 

tournefortiae. El Salvador. Tounefortia volubilis (Boraginaceae). 

Neolasioptera tournefortiae Möhn 1964a: 73. El Salvador: La Libertad, NE Sitio del Nino; HT; LT; SMNS. 

triadenii. USA (New York, Massachusetts, New Jersey). Triadenium virginicum (Hypericaceae). 

Cecidomyia triadenii Beutenmüller 1908: 74. USA: New Jersey, Middlesex Co.; T; LT; AMNH. 

Lasioptera virginica Felt 1909b: 287 (as n. sp. also in Felt 1911: 462). USA: Massachusetts, Magnolia; HT; ♀; 

NYSM. 

tribulae. Colombia. Tribulus cistoides (Zygophyllaceae). 

Neolasioptera tribulae Wünsch 1979: 81. Colombia: Magdalena, Santa Marta; HT; ♂; SMNS. 

tridentifera. Argentina (Mendoza, San Juan). Heliotropium curassavicum (Boraginaceae). 

Lasioptera tridentifera Kieffer & Jörgensen 1910: 398. Argentina: Mendoza, La Paz, & San Juan, Cancete; ST; 

F,P,L; lost. 

urvilleae. Brazil (RS). Urvillea uniloba (Sapindaceae). 

Lasioptera urvilleae Tavares 1909: 25. Brazil: RS, vic. São Leopoldo; ST; ♂,♀; TAVC. 

verbenae. USA (California). Verbena prostrata (Verbenaceae). 

Lasioptera verbenae Felt 1912h: 150. USA: California, Whittier, Puente Hills; ST; ♂,♀,L; NYSM. 

verbesinae. El Salvador. Verbesina turbacensis (Asteraceae). 

Neolasioptera verbesinae Möhn 1964a: 100. El Salvador: Chalatenango, nr La Palma; HT; ♂; SMNS. 

vernoniae. USA (Nebraska, Indiana, New York, North Carolina). Vernonia noveboracensis; V. baldwini (Asteraceae). 

Cecidomyia vernoniae Beutenmüller 1907a: 389. USA: New York, New York, Staten I., & North Carolina: 


Lasioptera vernoniflorae Felt 1908b: 324. USA: District of Columbia; ST; ♂,♀; USNM. 

vernoniensis. El Salvador. Vernonia sp. (Asteraceae). 

Neolasioptera vernoniensis Möhn 1964a: 103. El Salvador: La Libertad, SE Ciudad Arce; HT; LT; SMNS. 

viburnicola. USA (New York, Connecticut, Maryland, Virginia). Viburnum dentatum (Caprifoliaceae). 

Lasioptera viburnicola Beutenmüller 1907a: 398. USA: New York, New York, Bronx; ST; ♂,♀,L; AMNH. 

vitinea. USA (widespread eastern). Vitis sp. (Vitaceae). 

Lasioptera vitinea Felt 1907c: 153. USA: New York, New York, Staten I.; HT; ♀; NYSM. 

Lasioptera riparia Felt 1909b: 287. USA: New York, Westfield; HT; ♂; NYSM. 

Baldratia petiolicola Felt 1908b: 328. USA: Rhode Island; HT; ♀; USNM. 

willistoni. USA (California, Arizona, New Mexico). Atriplex canescens; A. polycarpa (Chenopodiaceae). 

Lasioptera willistoni Cockerell 1898: 327. USA: New Mexico, Mesilla Park; L (Gagné & Hawkins 1983: 380); LT; 


NEOMIKIELLA 

Mikiella Rübsaamen 1916b: 489, preocc. Meunier 1893. Type species, Cecidomyia beckiana Mik (orig. des.). 

Neomikiella Hedicke 1938a: 291, new name for Mikiella Rübsaamen. 

Palearctic; 3 spp.; Lasiopteridi, Dasineurini. Sylvén & Tastás-Duque (1993) indicate that this genus is differentiated 

from Dasineura on only a flimsy difference of the male genitalia.

eckiana. Widespread Europe. Inula conyza (Asteraceae). 

Cecidomyia beckiana Mik 1885: 140. Austria: Lower Austria; ST; ♂,♀,P,L; NHMW. 

kolomoetzae. Kazakhstan. Inula salicina (Asteraceae). 

Neomikiella kolomoetzae Fedotova 1992h: 16. Kazakhstan: Aïgyrkumy desert, 50 km NE Zaïsana, 4 km N 

Kabyrgatal; HT; ♂; ZMAS. 

lychnidis. Widespread Europe. Lychnis dioica (Caryophyllaceae). 

Cecidomyia lychnidis Vallot 1827: 94. France; ST; G; lost. 

Cecidomyia lychnidis Heyden 1861: 97. Germany; ST; A,P,L; ?. 

NEOMITRANTHELLA 

Neomitranthella Maia 1996a: 571. Type species, robusta Maia (orig. des.). 


robusta. Brazil (Rio de Janeiro). Neomitranthes obscura (Myrtaceae). 

Neomitranthella robusta Maia 1996a: 571. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

NEOMYCODIPLOSIS 

Neomycodiplosis Holz 1970: 112. Type species, ruebsaameni Holz (orig. des.). 

Western Palearctic; 1 sp.; Cecidomyiidi, Mycodiplosini. 

ruebsaameni. Germany. Peniophora pubera (Aphylloporales). 

Neomycodiplosis ruebsaameni Holz 1970: 113. Germany: Prignitz, Triglitz; HT; LT; SMNS. 

*NODALISTYLA 

Nodalistyla Fedotova & Sidorenko 2006d: 91. Type species, undata Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Brachineuridi. This genus is close to Brachineura if the two must be separated. 

*undata. Russia (Far East). 

Nodalistyla undata Fedotova & Sidorenko 2006d: 91. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; 

♂; ZMAS. 

*NORMANBYOMYIA 

Normanbyomyia Kolesik 2006: 40. Type species, fructivora Kolesik (orig. des.). 

Australasia; 1 sp.; Lasiopteridi, unplaced to tribe. 

fructivora. Australia (Queensland). Normanbya normanbyi (Arecaceae) 

Normanbyomyia fructivora Kolesik 2006: 42. Australia: Queensland, Cape Tribulation; HT; ♂; SAMA. 

NOVOCALMONIA 

Calmonia Tavares 1917b: 173, preocc. Clarke 1913. Type species, urostigmatis Tavares (orig. des.). 

*Novocalmonia Ozdikmen 2009: 201, new name for Calmonia Tavares. 

Paracalmonia Gagné & Etienne 2009: 338, error for Novocalmonia Ozdikmen. 

312

Neotropical; 3 spp., from Ficus; Lasiopteridi, unplaced to tribe. Ref.: Gagné & Etienne 2009: 240 (redescr.). 

fici. Brazil (Bahia). Ficus sp. (Moraceae). 

Asteromyia urostigmatis Tavares 1917b: 169, jun. sec. hom. Calmonia urostigmatis Tavares. Brazil: Bahia, Itaparica 

and S. Antonio da Barra; ST; ♀,P,L; TAVC. 

Calmonia fici Gagné 1994a: 88, new name for Asteromyia urostigmatis Tavares. 

*paucula. Guadeloupe. Ficus citrifolia (Moraceae). 

Paracalmonia paucula Gagné & Etienne 2009: 340. Guadeloupe: Petit-Canal; HT; ♂; USNM. 

urostigmatis. Brazil (Rio de Janeiro). Ficus sp. (Moraceae). 

Calmonia urostigmatis Tavares 1917b: 174. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♂,♀,P,L; TAVC. 

OBOLODIPLOSIS 

Obolodiplosis Felt 1908b: 410. Type species, Cecidomyia orbiculata Felt (orig. des.) = robiniae Haldeman. 

Nearctic, immigr. eastern and western Palearctic; 1 sp.; Cecidomyiidi, Lopesiini. 

robiniae. USA (New York, Maine, Pennsylvania, Maryland, Virginia); immigr.: widespread Europe, China, Republic 

of Korea, Japan. Robinia pseudoacacia (Fabaceae). Refs.: Kodoi et al. 2003 & Woo, et al. 2003 (rec. Republic 

of Korea & Japan, illus.); Uechi et al. 2005 (biol., descr.); Skuhravá et al. 2007 (distr. in Europe); Yang et al. 

2006 (rec. China). 

Cecidomyia robiniae Haldeman 1847: 193. USA: Pennsylvania; ST; ♂,♀,P,L; ?. 

Cecidomyia orbiculata Felt 1907a: 37. USA: New York, Albany; HT; ♂; NYSM. 

OCTODIPLOSIS 

Octodiplosis Giard 1894: cxli. Type species, Diplosis glyceriae Rübsaamen (orig. des.). 


glyceriae. UK, Germany. Glyceria maxima (Poaceae). 

Diplosis glyceriae Rübsaamen 1895a: 184. Germany: Plötzensee nr Berlin & Tegeler See; ST; ♂,♀,P,L; ZMHB. 

OCHNEPHILA 

Ochnephila Kieffer 1914: 514. Type species, socialis Kieffer (orig. des.). 

Afrotropical; 1 sp.; Lasiopteridi, unplaced to tribe. 

socialis. South Africa. Ochna arborea (Ochnaceae). 

Ochnephila socialis Kieffer 1914: 515. South Africa: vic. Cape Town; ST; F,P; lost?. 

ODINADIPLOSIS 

Odinadiplosis Mani 1935: 435. Type species, odinae Mani (orig. des.). 


amygdali. Malta, Greece, Turkey, Lebanon, Afghanistan. Prunus amygdalus (Rosaceae). 

Cecidomyia amygdali Anagnostopoulos 1929: 173. Greece: Attica; ST; ♂,♀,P,L,E; ?. 

odinae. India (Tamil Nadu, West Bengal). Odina wodier (Anacardiaceae). 

313

314 

Odinadiplosis odinae Mani 1935: 436. India: Tamil Nadu, Thanjavur, & West Bengal, Calcutta; ST; ♂,♀; ZSCI. 

ODONTODIPLOSIS 

Odontodiplosis Felt 1908b: 404. Type species, Cecidomyia karnerensis Felt. 

Ischnodiplosis Kieffer 1913g: 55. Type species, Lestodiplosis longiforceps Kieffer (orig. des.). 

Holarctic, Oriental; 14 spp., presumably predaceous; Cecidomyiidi, Lestodiplosini. 


Odontodiplosis americana Felt 1908b: 404. USA: New York, Albany; ST; ♂,♀; NYSM. 

bulbiformis. Ukraine. 

Odontodiplosis bulbiformis Mamaev 1998d: 9. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 

*cristiformis. Russia (Far East). 

Odontodiplosis cristiformis Fedotova & Sidorenko 2006c: 113. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; 

♂; ZMAS. 

fingeriformis. India (Maharashtra). 

Odontodiplosis fingeriformis Kashyap 1988a: 2. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

jonesi. India (Kerala). 

Odontodiplosis jonesi Nayar 1949: 86. India: Kerala, Trivandrum; HT; ♂; ZSCI. 


Cecidomyia karnerensis Felt 1907a: 45. USA: New York, Karner; HT; ♂; NYSM. 

longiforceps. France, Poland. 

Lestodiplosis longiforceps Kieffer 1904c: 403. France: vic. Bitche; ST; ♂,♀; lost?. Ref.: Szadziewski 1990 (redescr.). 

longiuscula. Ukraine. 

Odontodiplosis longiuscula Mamaev 1998d: 8. Ukraine: Zakarpatskaya Prov., Khust; HT; ♂; ZMMU. 


Odontodiplosis montana Felt 1908b: 404. USA: New York, Newport; HT; ♂; NYSM. 

muirshikha. India (Uttar Pradesh). 

Odontodiplosis muirshikha Grover & Bakhshi 1978: 78. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; 

PGCA. 

producta. Russia (Europe). 

Odontodiplosis producta Mamaev 1998d: 8. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; ZMMU. 

punctipennis. India (Uttar Pradesh). 

Odontodiplosis punctipennis Grover & Bakhshi 1978: 74. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; 

PGCA. 

raoi. India (Maharashtra). 

Odontodiplosis raoi Sharma 1986: 165. India: Maharashtra, Aurangabad; HT; ♂; INPC. 

travancorica. India (Kerala). 

Trisopsis travancoricus Nayar 1949: 87. India: Kerala, Trivandrum; ST; ♂,♀; ZSCI. 

*OGDODIPLOSIS 

Ogdodiplosis Fedotova 2004g: 14. Type species, utkini Fedotova (orig. des.). 


separated. 

*utkini. Russia (Far East). 

Ogdodiplosis utkini Fedotova 2004g: 15. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; KOVC. 

OKRIOMYIA

Okriomyia Kolesik 1998a: 46. Type species, schwarzi Kolesik (orig. des.). 

Australasian; 2 spp., from Eucalyptus; Cecidomyiidi, Asphondyliini, Schizomyiina. 

flabellidentata. Australia (South Australia). Eucalyptus cosmophylla (Myrtaceae). 

Okriomyia flabellidentata Kolesik 1998a: 48. Australia: South Australia, Cleland Conservation Pk, 10 km SE 


schwarzi. Australia (South Australia). Eucalyptus gracilis (Myrtaceae). 

Okriomyia schwarzi Kolesik 1998a: 48. Australia: South Australia, Nadda; HT; ♂; SAMA. 

OLIGOTROPHUS 

Oligotrophus Latreille 1805: 288. Type species, Tipula juniperina Linnaeus (mon.). 

Arceuthomyia Kieffer 1913g: 49. Type species, Rhopalomyia valerii Tavares (orig. des.). 

Schmidtiella Rübsaamen 1914: 89. Type species, gemmarum Rübsaamen (mon.). 

Allomyia Felt 1918b: 380, preocc. Banks 1916. Type species, juniperi Felt (orig. des.). 

Alassomyia Felt 1925: 147, new name for Allomyia Felt. 

Oligotrephus Shinji 1944: 225, missp. of Oligotrophus. 

Allasomyia Nayar 1945: 71, missp. of Alassomyia. 

Allassomyia Grover 1981: 74, missp. of Alassomyia. 

Holarctic; 10 spp., on Juniperus buds; Lasiopteridi, Oligotrophini. Ref.: Harris et al. 2006. 

betheli. Nearctic (widespread). Juniperus utahensis; Juniperus spp. (Cupressaceae). 

Oligotrophus betheli Felt 1912h: 148. USA: Colorado, McCoy; L (Gagné 1967a: 131); ♀; NYSM. 

Phytophaga sabinae Felt 1935a: 7. USA: Texas, Austin; ST; ♀; NYSM. 

Oligotrophus apicis Appleby & Neiswander 1965: 167. USA: Ohio, Wayne Co., Wooster; HT; ♂; USNM. 

gemmarum. UK, Netherlands, Germany, Poland, Czechoslovakia. Juniperus communis (Cupressaceae). 

Schmidtiella gemmarum Rübsaamen 1914: 89. Poland: Zielona Góra; ST; ♂,♀; ZMHB. Ref.: Harris et al. 2006. 

juniperi. USA (Colorado). Juniperus sp. (Cupressaceae). 

Allomyia juniperi Felt 1918b: 380. USA: Colorado, Oxford; HT; ♀; NYSM. 

juniperinus. Widespread Europe. Juniperus communis (Cupressaceae). 

Tipula juniperina Linnaeus 1758: 588. Sweden; T; A; LSUK. Ref.: Harris et al. 2006. 

nezu. Russia (Far East), Japan (Honshu). Juniperus rigida (Cupressaceae). 

Oligotrophus nezu Kikuti 1940: 35. Japan: Honshu, Kwazan, Higasiyama, Kyoto; HT; ♂; Kikuti. Ref.: Harris et al. 

2006. 

Arceuthomyia nakaharai Inouye 1959: 5. Japan: Honshu, Okayama Pref., Saidaiji City; HT; ♂; HFES. 

panteli. Widespread Europe, Algeria. Juniperus sp.; J. communis, J. sabina (Cupressaceae). 

Oligotrophus panteli Kieffer 1898a: 18. France; ST; A,L; lost?. Ref.: Harris et al. 2006. 

pattersoni. USA (Texas). Juniperus mexicana (Cupressaceae). 

Oligotrophus pattersoni White 1950: 72. USA: Texas, vic. Austin; ST; ♂,♀; USNM. 

schmidti. UK, Germany, Poland, Czechoslovakia, Yugoslavia. Juniperus communis (Cupressaceae). 

Oligotrophus schmidti Rübsaamen 1914: 91. Poland: Zielona Gora; ST; ♂,♀; ZMHB. 

uetsukii. Japan (Honshu). Juniperus rigida (Cupressaceae). 

Rhopalomyia uetsukii Inouye 1959: 6. Japan: Honshu, Okayama Pref., Okayama City; HT; ♂; HFES. The elongated 

empodia place this species with other juniper-infesting Lasiopteridi rather than with Rhopalomyia. 

valerii. UK, Portugal, Spain, France, Greece, Libya. Juniperus oxycedrus (Cupressaceae). 

Rhopalomyia valerii Tavares 1904b: 299. Portugal: Portas do Rodão, vic. Setubal; ST; ♂,♀,P; TAVC. 

Oligotrophus oxycedri (also as oxcedri) Rübsaamen 1916b: 555. Southern Europe; T; ♀; ZMHB. 

OLIGOXENOMYIA 

315

Oligoxenomyia Felt 1927d: 386. Type species, radicis Felt (orig. des.). 


radicis. Indonesia (Java). Freycinetia valida (Pandanaceae). 

Oligoxenomyia radicis Felt 1927d: 386. Indonesia: Java, Bolang, nr Bogor; ST; ♂,♀,P; NYSM. 

*OLLAEDIPLOSIS 

Ollaediplosis Fedotova 2004h: 179. Type species, ramosa Fedotova (orig. des.). 

Olladiplosis Fedotova 2006a: 153, misspelling. 

Eastern Palearctic; 1 sp.; Cecidomyiidi, tribe Aphidoletini. This genus is close to Bremia if the two must be separated. 

*ramosa. Russia (Far East). 

Ollaediplosis ramosa Fedotova 2004h: 179. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

OLPODIPLOSIS 

Olpodiplosis Gagné 1973c: 875. Type species, Diplosis helianthi Brodie (orig. des.). 


helianthi. Canada (Ontario), USA (Iowa, Missouri, Texas). Helianthus decapetalus, H. divaricatus; Helianthus spp. 

(Asteraceae). 

Diplosis helianthi Brodie 1894b: 44. Canada: Ontario, Toronto; ST; ♂,♀; USNM. 

*OPINATODIPLOSIS 

Opinatodiplosis Fedotova 2004g: 25. Type species, turriformis Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Karshomyiini. 


Opinatodiplosis turriformis Fedotova 2004g: 26. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

ORIBREMIA 

Oribremia Kieffer 1913c: 199. Type species, multifida Kieffer (orig. des.). 


multifida. India (Uttar Pradesh). 

Oribremia multifida Kieffer 1913c: 199. India: Uttar Pradesh, nr Bhowali, Kumaun; T; ♂; lost?. 

ORSEOLIA 

Orseolia Kieffer & Massalongo 1902: 56. Type species, cynodontis Kieffer & Massalongo (mon.). 

Courteia Kieffer & Docters van Leeuwen-Reijnvann 1910: 131. Type species, graminis Kieffer & Docters van 

316

317 

Leeuven-Reijnvann (orig. des. as n. g., n. sp.). 

Orseoliella Kieffer 1913a: 231. Type species, Orseolia javanica Kieffer & Docters van Leeuwen-Reijnvaan (orig. 

des.). 

Oreseolia Shinji 1944: 250, missp. of Orseolia. 

Palearctic, Afrotropical, Oriental; 28 spp., all on Poaceae; Cecidomyiidi, unplaced to tribe. Refs.: Gagné 1985b (rev.), 

Williams et al. 2002 (biology, ecology, and management of one species). 

andropogonis. India (Andhra Pradesh, Orissa). Dichanthium annulatum (Poaceae). 

Dyodiplosis andropogonis Felt 1917c: 371. India: Andhra Pradesh; L (Gagné 1985b: 136); ♂; NYSM. 

Dyodiplosis andropoginis Felt 1921f: 18, missp. of andropogonis. 

Dyodiplosis plumosa Felt 1921f: 21. India: Andhra Pradesh, Samalkot; L (Gagné 1985b: 136); ♂; NYSM. 

apludae. India (Tamil Nadu). Apluda varia (Poaceae). 

Orseoliella apludae Felt 1920b: 8. India: Tamil Nadu, Coimbatore; L (Gagné 1985b: 138); ♂; NYSM. 

bonzii. Nigeria. Paspalum scrobiculatum; P. orbiculare (Poaceae). 

Orseolia bonzii Harris 2000: 442. Nigeria: Ikwo LGA, nr Abakaliki; HT; ♂; BMNH. 

caulicola. Indonesia (Java). Sacciolepis indica (Poaceae). 

Parallelodiplosis javanica Felt 1921k: 91, jun. sec. hom. Orseolia javanica Kieffer & Docters van 

Leeuwen-Reijnvaan. Indonesia: Java, Bogor; L (Gagné 1985b: 139); ♂; NYSM. 

Orseolia caulicola Gagné 1973a: 506, new name for Parallelodiplosis javanica Felt. 


Clinodiplosis ceylanicus Kieffer 1912a: 26. Sri Lanka: Peradeniya; ST; ♀; NZSI?. 

Orseolia ceylonica Gagné 1973a: 506, missp. of ceylanica. 

cornea. India (Tamil Nadu). Ischaemum indicum (Poaceae). 

Pseudhormomyia cornea Felt 1917a: 76. India: Tamil Nadu, Coimbatore; L (Gagné 1985b: 140); ♂; NYSM. 

cynodontis. France, Italy, Hungary, Romania, Morocco, Algeria. Cynodon dactylon (Poaceae). 

Orseolia cynodontis Kieffer & Massalongo 1902: 57. Italy: Provinces of Verona, Tuscany, & Emilia; ST; ♂,♀,P,L,E; 

lost?. 

difficilis. Widespread India. Cynodon dactylon (Poaceae). 

Tristephanus cynodontis Grover 1968b: 166, jun. prim. hom. cynodontis Kieffer & Massalongo. India: Uttar Pradesh, 

Allahabad; HT; ♂; PGCA. 

Orseolia difficilis Gagné 1973a: 507, new name for Tristephanus cynodontis Grover. 

eragrostisae. Widespread India. Eragrostis tenella (Poaceae). 

Cecidomyia eragrostisae Mani 1936: 196. India: Kerala, Cannanore; L (Gagné 1985b: 141); ♂; ZSCI. 

fluvialis. India (Tamil Nadu). Paspalidium geminatum; Paspalidium spp. (Poaceae). 

Pseudhormomyia fluvialis Felt 1917a: 74. India: Tamil Nadu, Coimbatore; L (Gagné 1985b: 142); ♂; NYSM. 

graminicola. Indonesia (Java). Cynodon dactylon (Poaceae). 

Clinodiplosis graminicola Kieffer & Docters van Leeuwen-Reijnvaan 1910: 127. Indonesia: Java, Salatiga; ST; ♂,♀; 

lost?. 

graminis. Indonesia (Java). Ottochloa nodosa (Poaceae). 

Courteia graminis Kieffer & Docters van Leeuwen-Reijnvaan 1910: 131. Indonesia: Java, Salatiga; HT; ♀; lost?. 

indica. India (Mysore, Tamil Nadu). Cymbopogon schoenanthus (Poaceae). 

Dyodiplosis indica Felt 1921f: 19. India: Mysore, Bellary Distr., Yemmiganur; L (Gagné 1973a: 507); ♂; NYSM. 

ischaemi. India. Ischaemum pilosum (Poaceae). 

Oligotrophus ischaemi Kieffer 1910c: 71. Southern India; ST; P,L; lost?. 

javanica. Indonesia (Java). Imperata cylindrica (Poaceae). 

Orseolia javanica Kieffer & Docters van Leeuwen-Reijnvaan 1910: 125. Indonesia: Java, Salatiga; ST; ♂,♀; lost?. 

lourdusamyi. India (Tamil Nadu). Panicum psilopodium (Poaceae). 

Orseoliella lourdusamyi Mani 1986: 20. India: Tamil Nadu, Madras; HT; ♀; ZSCI. 

miscanthi. Japan (Honshu to Okinawa, Hachijo Is). Miscanthus sinensis (Poaceae). 

Contarinia miscanthi Shinji 1938d: 315. Japan: Honshu, Morioka; ST; ♂,♀; lost. 

Asphondylia sasae Shinji 1944: 106. Japan; ST; A,L; lost.

318 

mnesitheae. India (Orissa). Mnesithea laevis (Poaceae). 

Orseolia mnesitheae Gagné 1985b: 146. India: Orissa, Cuttack; HT; ♂; NZSI. 

monticola. India (Mysore). Chrysopogon fulvus (Poaceae). 

Dyodiplosis monticola Felt 1921f: 20. India: Mysore, Bellary Distr., Muruvani; L (Gagné 1985b: 147); ♂; NYSM. 

nwanzii. Nigeria. Eragrostis atrovirens (Poaceae). 

Orseolia nwanzii Harris & Nwilene 2007: 261. Nigeria: Ebonyi State, Ikwo, Abakaliki; HT; ♂; BMNH. 

orientalis. Indonesia (Java). Oplismenus compositus (Poaceae). 

Orseoliella orientalis Felt 1921k: 90. Indonesia: Java, Bogor; L (Gagné 1985b: 148); ♂; NYSM. 

oryzae. Widespread India, southern China, Thailand, Indonesia. Oryza sativa; O. perrenis (Poaceae). 

Cecidomyia oryzae Riley 1881: 149, nomen nudum. 

Cecidomyia oryzae Wood-Mason 1889: 103. India: "Kurruckpore;" S; ♀,P; NZSI. 

oryzivora. Senegal, Ivory Coast, Burkina Faso, Nigeria, Sudan. Oryza glaberrima (Poaceae). 

Orseolia oryzivora Harris & Gagné 1982: 469. Senegal: Djibelor; HT; ♂; BMNH. 

paspali. Widespread India, Indonesia (Java). Paspalum scrobiculatum; P. conjugatum (Poaceae). 

Parallelodiplosis paspali Felt 1921k: 92. Indonesia: Java, Bogor; L (Gagné 1985b: 150); ♂; NYSM. 

Itonida paspalumi Mani 1936: 196. India: Kerala, Cannanore; L (Gagné 1985b: 150); ♂; ZSCI. 

polliniae. India (Kerala). Eulalia trispicata (Poaceae). 

Lopesiella polliniae Felt 1927a: 2. India: Kerala, Taliparamba; L (Gagné 1985b: 151); ♀; NYSM. 

sallae. India (Andhra Pradesh). Pseudoraphis spinascens (Poaceae). 

Orseoliella graminis Felt 1921f: 22, jun. sec. hom. graminis Kieffer & J. Docters van Leeuwen-Reijnvaan. India: 

Andhra Pradesh, Achanta; L (Gagné 1985b: 151); ♀; NYSM. 

Orseolia sallae Gagné 1973a: 508, new name for Orseoliella graminis Felt. 

similis. India (Tamil Nadu). Ischaemum indicum (Poaceae). 

Orseoliella ischaemi Mani 1947: 445, jun. sec. hom. ischaemi Kieffer. India: Tamil Nadu, Coimbatore; HT; ♂; ZSCI. 

Orseolia similis Gagné 1973a: 508, new name for Orseoliella ischaemi Mani. India: Tamil Nadu. 

visvanathi. India (Uttar Pradesh). 

Macrodiplosis visvanathi Rao 1953: 314. India: Uttar Pradesh, Dehra Dun; HT; ♂; ZSCI. 

ORTHODIPLOSIS 

Orthodiplosis Kieffer 1913h: 109. Type species, macrurus Kieffer (orig. des.). 


macrurus. Kenya. 

Orthodiplosis macrura Kieffer 1913h: Kenya: Tiwi; HT; ♀; MNHN. 

OURADIPLOSIS 

Ouradiplosis Felt 1915h: 154. Type species, aurata Felt (orig. des.). 


aurata. Brazil (Pará). 

Ouradiplosis aurata Felt 1915h: 154. Brazil: Pará, Igarape-Açú; HT; ♀; CUIC. 

*OXYCEPHALOMYIA 

Oxycephalomyia Tokuda & Yukawa 2004: 54. Type species, Asphondylia styraci Shinji (orig. des.). 

Palearctic; 1 sp.; Cecidomyiidi, Asphondyliini, Schizomyiina.

319 

styraci. Japan (Honshu, Okinawa). Styrax japonica (Styracaceae). 

Asphondylia styraci Shinji 1944: 376 Japan; NT (Yukawa 1983:103; Japan: Saitama Pref., Urawa City, Funayama); 

♂; KUIC. Ref.: Tokuda et al. 2004 (illus., biol.). 

OZIRHINCUS 

Ozirhincus Rondani 1840: 16. Type species, longicollis Rondani (mon.). 

Oxyrhynchus Anonymous 1844: 450, missp. of Ozirhincus. 

Oxyrhynchus Loew 1845a: 11, new name for Ozirhincus Rondani. 

Ozirhyncus Rondani 1846: 372, missp. of Ozirhincus. 

Clinorhyncha Loew 1850a: 21, new name for Ozirhincus Rondani. 

Oxyrynchus Loew 1850a: 21, missp. of Oxyrhynchus. 

Ozirhynchus Bigot 1854: 461, missp. of Ozirhincus. 

Clinorrhyncha Loew 1856: 79, missp. of Clinorhyncha. 

Oxyrhyncus Rondani 1856: 200, missp. of Ozirhincus. 

Oxyrrhynchus Walker 1856a: xxii, missp. of Ozirhincus. 

Oxyrhincus Rondani 1860: 171, missp. of Ozirhincus. 

Ozyrhyncus Rondani 1860: 171, missp. of Ozirhincus. 

Ozyrhinchus Rondani 1860: 291, missp. of Ozirhincus. 

Acorhynchus Rondani 1860: 291, unjust. new name for Ozirhincus Rondani. 

Acyrhynchus Rondani 1860: 288, missp. of Acorhynchus. 

Ceinorhynca Rondani 1860: 291, missp. of Clinorhyncha. 

Chinorhyncha Rondani 1860: 291, missp. of Clinorhyncha. 

Clinorhynca Rondani 1860: 291, missp. of Clinorhyncha. 

Oxyrincus Lioy 1864: 501. 60., missp. of Ozirhincus. 

Oziorhincus Schiner 1868: 3, missp. of Ozirhincus. 

Oziorhynchus Walker 1874: 14, missp. of Ozirhincus. 

Achorhychus Scudder 1882: 4, missp. of Acorhynchus. 

Acorrhynchus Scudder 1882: 5, missp. of Acorhynchus. 

Oxirhynchus Scudder 1882: 240, missp. of Ozirhincus. 

Oziorhinchus Scudder 1882: 241, missp. of Ozirhincus. 

Chinorrhynca Scudder 1882: 70, missp. of Clinorhyncha. 

Clinorrhynca Scudder 1882: 77, missp. of Clinorhyncha. 

Clinorhynca Scudder 1884: 64, missp. of Clinorhyncha. 

Ozyrrhynchus Kieffer 1898f: 4, missp. of Ozirhincus. 

Ozirhinchus Kieffer 1901b: 438, missp. of Ozirhincus. 

Oziorhyncus Kertész 1902: 335, missp. of Ozirhincus. 

Ozirchyncus Kertész 1903: 180, missp. of Ozirhincus. 

Clinoryncha Rübsaamen 1916b: 561, missp. of Clinorhyncha. 

Palearctic, immigr. Nearctic; 5 spp., all in achenes of Anthemideae (Asteraceae); Lasiopteridi, Lasiopterini. Refs: 

Verrall 1889 (discussion of earlier misspellings of Ozirhincus and Clinorhyncha), Möhn 1966, 1968a (larvae and host 

records). 

anthemidis. Germany & Circummediterranean. Anthemis arvensis, A. tinctoria; Anthemis spp. (Asteraceae). 

Clinoryncha anthemidis Rübsaamen 1916b: 561. Germany: Rhineland; T; A; ZMHB. 

Ozirhincus anatolicus, as subsp. of anthemidis, Möhn 1968a: 62. Turkey: East Anatolia, Amasya; HT; LT; SMNS. 

Ozirhincus dalmaticus Möhn 1968a: 66. Croatia: Dalmatia, Clissa; HT; LT; SMNS. 

Ozirhincus ibericus, as subsp. of anthemidis, Möhn 1968a: 64. Spain: Malaga Prov., Ronda; HT; LT; SMNS. 

Ozirhincus kabylensis Möhn 1968a: 65. Algeria: Kabylei, Kap Aôkas; HT; LT; SMNS. 

Ozirhincus mallorcensis, as subsp. of anthemidis, Möhn 1968a: 63. Spain: Mallorca, Palma; HT; LT; SMNS. 

longicollis. Widespread western Palearctic. Orig. unkn.; Chrysanthemum spp. & several other genera of Anthemideae

320 

(Asteraceae). 

Ozirhincus longicollis Rondani 1840: 16. Italy; T; A; lost. 

Lasioptera chrysanthemi Loew 1850a: 39. Poland; T; A; ZMHB?. 

Clinorhyncha crassipes Winnertz 1853: 309. Germany; ST; ♂,♀,L; lost. 

Clinorrhyncha leucanthemi Kieffer 1898f: 4. France: Moselle; T; G; lost?. 

Ozirhincus planus, as subsp. of longicollis, Möhn 1966: 40. Croatia: Istria, Canfanaro; HT; LT; SMNS. 

millefolii. Widespread Europe, Turkey, Russia (Far East); immigr: widespread Nearctic. Achillea millefolium; Achillea 

spp., Santolina spp. (Asteraceae). 

Clinorrhyncha millefolii Wachtl 1884: 161. Austria: vic. Vienna; ST; ♂,♀,L; NHMW?. 

Clinorhyncha filicis Felt 1907a: 12. USA: New York, Nassau; HT; ♀; NYSM. 

Clinorhyncha karnerensis Felt 1908b: 333. USA: New York, Karner; HT; ♀; NYSM. 

Ozirhincus amurensis, as subsp. of millefolii, Möhn 1968a: 52. Russia: Zejskaja Pristán am Zejafluss, Amur/UddSR; 

HT; LT; SMNS. 

Ozirhincus lusitanicus, as subsp. of millefolii, Möhn 1968a: 50. Portugal: Serra da Boa Viageur; HT; LT; SMNS. 

Ozirhincus minor, as subsp. of millefolii Wachtl, Möhn 1968a: 51. Slovakia: Nová Ba�a; HT; LT; SMNS. 

tanaceti. Widespread Europe. Tanacetum vulgare; Anthemis tuberculata, Chrysanthemum spp. (Asteraceae). 

Clinorrhyncha tanaceti Kieffer 1889d: 209. France: Bitche; ST; ♂,♀,L; lost?. 

Ozirhincus hispanicus Möhn 1968a: 57. Spain: Sierra Nevada, Cartijo de la Vibora; HT; LT; SMNS. 

Ozirhincus hungaricus Möhn 1968a: 55. Hungary: Budapest; HT; LT; SMNS. 

Ozirhincus parvus Möhn 1968a: 56. Italy: Etruscan Apennines, Monte Cuccioli; HT; LT; SMNS. 

trichatus. Syria. Achillea fragrantissima (Asteraceae). 

Ozirhincus trichatus Möhn 1968a: 67. Syria: nr Damascus; HT; LT; SMNS. 

PACHYDIPLOSIS 

Pachydiplosis Kieffer 1913h: 108. Type species, apricans Kieffer (orig. des.). 


apricans. Kenya. 

Pachydiplosis apricans Kieffer 1913h: 109. Kenya: Shimoni; HT; ♀; MNHN. 

*PALMATODIPLOSIS 

Palmatodiplosis Fedotova 2004g: 29. Type species, fasciculifera Fedotova (orig. des.). 

Eastern Palearctic; 3 spp.; Cecidomyiidi, Karshomyiini. This genus is close to Karshomyia if the two must be 

separated. 

*fasciculifera. Russia (Far East). 

Palmatodiplosis fasciculifera Fedotova 2004g: 29. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

*galeriformis. Russia (Far East). 

Karshomyia galeriformis Fedotova 2004g: 10. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

Palmatodiplosis galeriformes Fedotova & Sidorenko 2009a: 25, missp. of galeriformis. 

*variana. Russia (Far East). 

Palmatodiplosis variana Fedotova & Sidorenko 2009a: 23. Russia: Primorskiï Terr., Lazovskiï Res., Koeiskaya Pad'; 


PARADIPLOSIS

Paradiplosis Felt 1908b: 410. Type species, Cecidomyia obesa Felt (orig. des.). 

Agevillea Hubault 1945: 23. Type species, abietis Hubault (orig. des. as n. g., n. sp.) = abietispectinatae Tubeuf. 

Polydiplosis Perny 1994: 11, missp. of Paradiplosis. 

Holarctic; 4 spp., on Abies and Picea; Cecidomyiidi, Cecidomyiini. Ref.: Osgood & Gagné 1978 (illus., biol.). 

abietispectinatae. Central and southeastern Europe. Abies alba (Pinaceae). 

Diplosis abietispectinatae (as abietis pectinatae) Tubeuf 1930: pl. II B. Germany; ST; LT; ?. 

Agevillea abietis Hubault 1945: 23. France: Ageville; ST; ♂,♀,L,egg; ?. 

manii. Japan (Hokkaido to Kyushu). Abies mayriana; A. firma (Pinaceae). 

Aschistonyx abietis Mani 1954b: 115, jun. sec. hom. abietis Hubault. Japan: Hokkaido; HT; ♂; ZSCI. Ref.: Tokuda & 

Yukawa 2003: 59 (illus.). 

Agevillea manii Inouye 1959: 1, new name for Aschistonyx abietis Mani 1954. 

obesa. USA (New York). Picea glauca (Pinaceae). 

Cecidomyia obesa Felt 1907a: 38. USA: New York, Lake Clear; HT; ♂; NYSM. 

tumifex. Canada (Quebec, Newfoundland, New Brunswick), USA (Wisconsin, Maine, North Carolina). Abies 

balsamea (Pinaceae). 

Paradiplosis tumifex Gagné 1978a: 88. USA: Maine, Penobscot Co.; HT; ♂; USNM. 

PARALLELODIPLOSIS 

Parallelodiplosis Rübsaamen 1910d: 287. Type species, Diplosis galliperda Löw (mon.). 

Parellalodiplosis Mani 1946: 224, missp. of Parallelodiplosis. 

Holarctic, Oriental, Oceanian; 22 spp.; Cecidomyiidi, Clinodiplosini. Several species are associated with gall makers 

of various insect orders. Ref.: Gagné 2008c (descr., biol. of 1 sp.). 

abdominalis. USA (New York). 

Oligotrophus caryae Felt 1907a: 28, jun. sec. hom. Cecidomyia caryae Felt. USA: New York, Albany; HT; ♀; 

NYSM. 

Itonida abdominalis Felt 1921a: 207, new name for Oligotrophus caryae Felt. 

acernea. USA (Minnesota, New York, North Carolina). Orig. unk.; Phylloxera spp. (Hemiptera: Phylloxeridae) 

Cecidomyia acernea Felt 1907a: 46. USA: New York, Nassau; HT; ♂; NYSM. 

aprilis. USA (New York). 

Itonida aprilis Felt 1912l: 247. USA: New York, Albany; ST; ♂; NYSM. 

autumnalis. India (Uttar Pradesh). 

Parallelodiplosis autumnalis Grover & Bakhshi 1978: 32. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; 

PGCA. 

bimaculata. Hawaiian Is. 

Parallelodiplosis bimaculata Hardy 1960: 300. Hawaiian Is: Oahu, Ewa; HT; ♂; HDOA. 

bupleuri. Germany, Czechoslovakia. Bupleurum falcatum (Apiaceae). 

Clinodiplosis bupleuri Rübsaamen 1895b: 182. Germany: Sinzig; ST; ♂,♀,L; ZMHB. 

carpinicola. USA (New York). 

Cecidomyia carpini Felt 1907a: 38, jun. prim. hom. carpini Löw 1874. USA: New York, Albany; HT; ♂; NYSM. 

Cecidomyia carpinicola Kieffer 1913i: 214, new name for Cecidomyia carpini Felt. 

caryae. USA (Minnesota, New York, Maryland). Orig. unk; Caryomyia spp. (Diptera: Cecidomyiidae). Ref.: Gagné 

2008c: 97 (redescr., biol.). 

Cecidomyia caryae Felt 1907a: 45. USA: New York, Albany; HT; ♂; NYSM. 

florida. USA (Florida). Macrodiplosis sp. (Diptera: Cecidomyiidae). 

Clinodiplosis florida Felt 1908b: 411. USA: Florida, Georgiana; HT; ♂; USNM. 

galliperda. Widespread Europe. Neuroterus lenticularis (Hymenoptera: Cynipidae). 

321

322 

Diplosis galliperda Löw 1889a: 202. France: Bitche; ST; ♂,♀,L; NHMW. 

hartmaniae. USA (New York). 

Itonida hartmaniae Felt 1921a: 201. USA: New York, Albany; HT; ♂; NYSM. 

indorensis. India (Madhya Pradesh). 

Parallelodiplosis indorensis Grover 1979c: 37. India: Madhya Pradesh, Indore; HT; ♂; PGCA. 


Clinodiplosis montana Felt 1908b: 412. USA: New York, Newport; HT; ♂; NYSM. 

nixoni. USA (New York). 

Cecidomyia nixoni Felt 1908b: 414. USA: New York, Albany; HT; ♂; NYSM. 

ramuli. USA (New York). Cornus racemosa (Cornaceae). 

Cecidomyia ramuli Felt 1907c: 164. USA: New York, Albany; HT; ♀; NYSM. 

rotunda. India (Maharashtra). 

Parallelodiplosis rotunda Chandra 1993e: 182. India: Maharashtra, Parbhani; HT; ♂; CSIA. 

rubrascuta. USA (New York). 

Cecidomyia rubrascuta Felt 1907a: 46. USA: New York, Albany; HT; ♂; NYSM. 

Cecidomyia rubroscuta Felt 1907c: 142, missp. of rubrascuta. 

sarae. USA (New York). 

Cecidomyia coryli Felt 1907a: 46, jun. prim. hom. coryli Kaltenbach 1859. USA: New York, Albany; HT; ♂; NYSM. 

Parallelodiplosis sarae Gagné 1972b: 322, new name for Cecidomyia coryli Felt. 

spirae. USA (New York, Massachusetts). Spiraea salicifolia (Rosaceae). 

Clinodiplosis spirae Felt 1909b: 293. USA: Massachusetts, Magnolia; HT; ♂; NYSM. 

subtruncata. USA (Minnesota, New York, Massachusetts, Maryland). Orig. unk.; C. florida, C. stolonifera 

(Cornaceae). 

Cecidomyia subtruncata Felt 1907a: 44. USA: New York, Albany; HT; ♂; NYSM. 

tolhurstae. USA (New York). 

Cecidomyia tolhurstae Felt 1908b: 414. USA: New York, Albany; HT; ♂; NYSM. 

triangularis. India (Maharashtra). 

Parallelodiplosis triangularis Grover & Kashyap 1991: 92. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

*PARAMETASPHONDYLIA 

Parametasphondylia Maia & Santos 2007: 137. Type species, piperis Maia & Santos (orig. des.). 


*piperis. Brazil (Minas Gerais). Piper sp. (Piperaceae). 

Parametasphondylia piperis Maia & Santos 2007: 138. Brazil: Minas Gerais, Tiradentes; HT; ♂; QBUM. 

PARASCHIZOMYIA 

Paraschizomyia Möhn 1961a: 4. Type species, Cecydomyia buboniae Frauenfeld (orig. des.). 

North African; 1 sp.; Cecidomyiidi, Asphondyliini, Schizomyiina. 

buboniae. Widespread North Africa. Deverra tortuosa; D. scoparia (Apiaceae). 

Cecydomyia buboniae Frauenfeld 1859: 325. Egypt: Cairo to Suez; T; A; NHMW. 

Schizomyia deverrae Kieffer 1912f: 141. Tunisia: Matmata, Oued Chahal, 50 km S Sfax, & Metlaoui; ST; ♀,P; lost?. 

PARAZALEPIDOTA 

Parazalepidota Maia 2001a: 26. Type species, clusiae Maia (orig. des.).

323 

Neotropical; 1 sp.; Cecidomyiidi, Asphondyliini, Asphondyliina. Parazalepidota differs from Zalepidota on 

seemingly trivial characters, which can be evaluated as more species are discovered. 

clusiae. Brazil (Rio de Janeiro). Clusia fluminensis (Clusiaceae). 

Parazalepidota clusiae Maia 2001a: 27. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

*PARKIAMYIA 

Parkiamyia Maia 2006a: 2. Type species, paraensis Maia (orig. des.). 

Neotropical; 1 sp.; Cecidomyiidi, Cecidomyiini. This genus appears to differ from Contarinia only in the elongate 

aedeagus. It should be compared with Contarinia species from other Mimosoideae. 

*paraensis. Brazil (Pará). Parkia pendula (Fabaceae). 

Parkiamyia paraensis Maia 2006a: 3. Brazil: Pará: Porto Trombetas; HT; ♂; MNRJ. Ref.: Fernandes et al. 2007: 471 

ff (biol.). 

PAULLINIAMYIA 

Paulliniamyia Maia 2001a: 29. Type species, ampla Maia (orig. des.). 


ampla. Brazil (Rio de Janeiro). Paullinia weinmanniaefolia (Sapindaceae). 

Paulliniamyia ampla Maia 2001a: 30. Brazil: Rio de Janeiro, Arrial do Cabo; HT; ♂; MNRJ. 

PECTINODIPLOSIS 

Pectinodiplosis Felt 1918e: 132. Type species, Contarinia erratica Felt (orig. des.). 

Nearctic; 1 sp.; Cecidomyiidi, Lestodiplosini. Ref.: Gagné & Etienne 2009: 248 (redescr.). 

erratica. USA (District of Columbia); Guadeloupe (West Indies). Hemerocampa leucostigma (Lepidoptera: 

Lymantriidae); Parasaissetia nigra (Hemiptera: Coccidae); Crypticerya genistae (Hemiptera: 

Monophlebidae). 

Contarinia erratica Felt 1908b: 391. USA: District of Columbia; HT; ♂; USNM. 

*PEDETENTIA 

Pedetentia Fedotova & Sidorenko 2009d: 248. Type species, polita Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 3 spp; Cecidomyiidi, Clinodiplosini. This genus is close to Clinodiplosis if the two must be 

separated. Fedotova & Sidorenko (2009d: 250) erect the subtribe Pedetentiina for this genus and nest it within the tribe 

Inclinatidiplosini but neither subdivision is used here (see Appendix under Clinodiplosini). 

*escifera. Russia (Far East). 

Pedetentia escifera Fedotova & Sidorenko 2009d: 250. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya Bay; 


*maritima. Russia (Far East). 

Pedetentia maritima Fedotova & Sidorenko 2009d: 250. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya 


*polita. Russia (Far East).

324 

Pedetentia polita Fedotova & Sidorenko 2009d: 248. Russia: Primorskiï Terr., Lazovskiï Res., Cordon America; HT; 

♂; ZMAS. 

PERASPHONDYLIA 

Perasphondylia Möhn 1960c: 218. Type species, reticulata Möhn (orig. des.). 

Neotropical; 2 spp., on Eupatorieae (Asteraceae); Cecidomyiidi, Asphondyliini, Asphondyliina. 

mikaniae. Brazil (Rio de Janeiro). Mikania glomerata (Asteraceae). 

Perasphondylia mikaniae Gagné 2001: 132. Brazil: Rio de Janeiro, Silva Jardim, Poço das Antas Biol. Res.; HT; M; 

QBUM. 

reticulata. Mexico, El Salvador, Trinidad, Bolivia, Brazil. Eupatorium sp.; Chromolaena odorata (Asteraceae). 

Perasphondylia reticulata Möhn 1960c: 219. El Salvador: W Chalatenango; HT; ♂; SMNS. 

PERODIPLOSIS 

Perodiplosis Kieffer 1913h: 100. Type species, coronata Kieffer (orig. des.). 


coronatus. Kenya. 

Perodiplosis coronatus Kieffer 1913h: 100. Kenya: Ramisi R.; HT; ♀; MNHN. 

PERNETTYELLA 

Pernettyella Kieffer & Herbst 1909: 126. Type species, longicornis Kieffer & Herbst (orig des. as n. g., n. sp.). 

Parnettyella Neave 1940: 614, missp. of Pernettyella. 

Fernettyella Shinji 1944: 219, missp. of Pernettyella. 


longicornis. Chile. Pernettya furens (Ericaceae). 

Pernettyella longicornis Kieffer & Herbst 1909: 126. Chile: Concepción; ST; ♂,♀,P,L; lost. 

PHEGOMYIA 

Phegomyia Kieffer 1913g: 49. Type species, Oligotrophus fagicola Kieffer (orig. des.). 

Palearctic; 2 spp., on Fagus; Lasiopteridi, unplaced to tribe. 

fagicola. Widespread Europe. Fagus sylvatica (Fagaceae). 

Oligotrophus fagicola Kieffer 1901c: 23. France: Bitche; ST; ♂,♀; lost?. 

tokunagai. Japan (Honshu). Fagus crenata (Fagaceae). 

Phegomyia tokunagai Sasakawa & Koyama 1953: 4. Japan: Honshu, Ashiu, Kyoto Univ. Experimental For.; ST; 

♂,♀; ELSU. 

PHYLLODIPLOSIS 

Phyllodiplosis Kieffer 1912h: 1. Type species, Contarinia cocciferae Tavares (orig. des.). 

Western Palearctic; 1 sp.; Cecidomyiidi, Cecidomyiini. This genus shoud be compared with Prodiplosis (cf. Gagné

1986b). 

cocciferae. UK, Portugal, Spain, Italy, Malta, Yugoslavia, Greece, Morocco. Quercus ilex (Fagaceae). 

Contarinia cocciferae Tavares 1901: 72. Portugal: Arrabida; Sobral do Campo; ST; ♀,P,egg; TAVC. 

PHYSEMOCECIS 

Physemocecis Rübsaamen 1914: 88. Type species, ulmi Rübsaamen (mon) = ulmi Kieffer. 

Palearctic; 2 spp.; Lasiopteridi, unplaced to tribe. 

hartigi. Widespread Europe, Turkey. Tilia grandifolia; T. cordata (Tiliaceae). 

Hormomyia hartigi Liebel 1892: 285. France: vic. Bitche; ST; ♂,♀,L,egg; lost. 

ulmi. Widespread Europe. Ulmus campestris; Ulmus spp. (Ulmaceae). 

Oligotrophus ulmi Kieffer 1909b: 31. France; T; LT; lost?. 

Physemocecis ulmi Rübsaamen 1914: 88. Germany; ST; F,L; ZMHB. 

PILODIPLOSIS 

Pilodiplosis Gagné 1973c: 876. Type species, Cecidomyia helianthibulla Walsh (orig. des.). 


helianthibulla. Widespread eastern Nearctic. Helianthus sp.; Helianthus spp. (Asteraceae). 

Cecidomyia helianthibulla (as helianthi-bulla) Walsh 1866: 228. USA: Illinois, nr Rock Island; T; G; USNM. 

Hormomyia bulla Felt 1914g: 286. USA: Illinois, Evanston; ST; ♂,♀; NYSM. 

PINYONIA 

Pinyonia Gagné 1970a: 153. Type species, edulicola Gagné (orig. des.). 

Nearctic; 1 sp.; Cecidomyiidi, Cecidomyiini. 

edulicola. USA (Colorado, New Mexico). Pinus edulis (Pinaceae). 

Pinyonia edulicola Gagné 1970a: 154. USA: New Mexico, Colfax Co., Raton; HT; ♂; USNM. 

PIPALDIPLOSIS 

Pipaldiplosis Mani 1943: 46. Type species, pipaldiplosis Mani (orig. des.). 


pipaldiplosis. India (Delhi). Ficus religiosa (Moraceae). 

Pipaldiplosis pipaldiplosis Mani 1943: 46. India: Delhi; HT; ♂; INPC. India. 

PIRANEA 

Piranea Jane�i 1990: 29. Type species, spartii Jane�i (orig. des. as n. g., n. sp.). 

Palearctic; 1 sp.; Cecidomyiidi, Clinodiplosini. The single species appears to belong to Clinodiplosis, but one cannot 

be certain from the original description alone. 

325

spartii. Slovenia. Spartium junceum (Fabaceae). 

Piranea spartii Jane�i 1980: 49. Slovenia: Piran & vic.; ST; ♂,♀,L; ?. 

PISONIAMYIA 

Pisoniamyia Möhn 1960d: 338. Type species, armeniae Möhn (orig. des.). 

Neotropical; 3 spp., on Pisonia; Cecidomyiidi, Asphondyliini, Schizomyiina. Ref.: Möhn 1962 (illus.). 

armeniae. El Salvador. Pisonia macranthocarpa (Nyctaginaceae). 

Pisoniamyia armeniae Möhn 1960d: 339. El Salvador: La Libertad, W Ateos; ♂; SMNS. 

mexicana. Mexico (Tamaulipas). Pisonia aculeata (Nyctaginaceae). 

Brugmanniella mexicana Felt 1911l: 465. Mexico: Tamaulipas, Tampico; T; G; lost. 

similis. El Salvador. Pisonia macranthocarpa (Nyctaginaceae). 

Pisoniamyia similis Möhn 1960d: 342. El Salvador: La Libertad, W Ateos; ♂; SMNS. 

PISPHONDYLIA 

Pisphondylia Möhn 1960d: 344. Type species, salvadorensis Möhn (orig. des.). 

Neotropical; 2 spp., on Nyctaginaceae; Cecidomyiidi, Asphondyliini, Schizomyiina. 

brasiliensis. Brazil (Rio de Janeiro). Guapira opposita (Nyctaginaceae). 

Pisphondylia brasiliensis Couri & Maia 1992: 730. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; 

QBUM. 

salvadorensis. El Salvador. Pisonia macranthocarpa (Nyctaginaceae). 

Pisphondylia salvadorensis Möhn 1960d: 345. El Salvador: La Libertad, W Ateos; HT; ♂; SMNS. 

PITYDIPLOSIS 

Pitydiplosis Gagné 1973c: 877. Type species, Mycodiplosis packardi Felt (orig. des.). 


packardi. USA (New York). Pinus strobus (Pinaceae). 

Mycodiplosis packardi Felt 1918b: 382. USA: N.Y., Round L.; HT; ♂; NYSM. 

PLACOCHELA 

Placochela Rübsaamen 1916c: 4. Type species, Epidosis nigripes Löw (orig. des.). 

Palearctic; 2 spp.; Cecidomyiidi, Asphondyliini, Schizomyiina. This genus might easily be subsumed under 

Schizomyia, q.v. for remarks. Ref.: Tokuda et al. 2005 (morph. and molecular studies). 

araliae. Russia (Far East). Aralia mandshurica (Araliaceae). 

Placochela araliae Mamaev & Kritskaya 1981: 404. Russia: Primorskiï Terr., Ussuriyskiï Res.; HT; ♂; HYNM. 

nigripes. Widespread Europe. Lonicera xylosteum, Sambucus nigra, S. ebulus, Viburnum lantana (Caprifoliaceae); 

Ligustrum vulgare (Oleaceae). Ref.: Tokuda et al. 2005: 421. 

Epidosis nigripes Löw 1877: 19. Austria; ST; ♂,♀,L; NHMW. 

Schizomyia propinqua Rübsaamen 1889a: 45. Germany: Siegen; ST; ♂,♀,L; ZMHB. 

Placochela ligustri Rübsaamen 1899d: 598. Germany: Linz; T; ♀; ZMHB. 

326

PLANETELLA 

Planetes Walker 1837: 178, preocc. Macleay 1825. Type species, extrema Walker (orig. des.). 

Planetella Westwood 1840: 126, new name for Planetes Walker. 

Hormomyia Loew 1850a: 31. Type species, crassipes Loew (subs. des. Coquillett 1910: 553) = Planetes extremus 

Walker. 

Angelinia Rondani 1860: 290. Type species, gibbosa Rondani (mon.). 

Pseudhormomyia Kieffer 1898f: 57. Type species, granifex Kieffer (mon.). 

Dichrona Rübsaamen 1899b: 542. Type species, gallarum Rübsaamen (orig. des. as n. g., n. sp.). 

Homomyia Rübsaamen 1899d: 603, missp. of Hormomyia. 

Amaurosiphon Rübsaamen 1910d: 287. Type species, caricis Rübsaamen (subs. mon. Rübsaamen 1911h: 391). 

Dyodiplosis Rübsaamen 1910d: 287. Type species, Hormomyia arenariae Rübsaamen (mon.). 

Pseudohormomyia Rübsaamen 1911h: 390, missp. of Pseudhormomyia. 

Dishormomya Kieffer 1912h: 2. Type species, Hormomyia cornifex Kieffer (orig. des.). 

Taphodiplosis Kieffer 1912h: 2. Type species, Pseudhormomyia subterranea Kieffer & Trotter (orig. des.). 

Trishormomya Kieffer 1912h: 2. Type species, Hormomyia strobli Kieffer (orig. des.). 

Dishormomyia Kieffer 1913i: 138, missp. of Dishormomya. 

Proshormomyia Kieffer 1913i: 136. Type species, Hormomyia winnertzi Kieffer (mon.). 

Trishormomyia Kieffer 1913i: 139, missp. of Trishormomya. 

Jaapiola Rübsaamen 1914: 95. Type species, tarda Rübsaamen (mon.). 

Diplolaboncus Rübsaamen 1917: 59. Type species, Hormomyia tumorifica Rübsaamen (orig. des.). 

Paurosphondylus Rübsaamen 1917: 59. Type species, Hormomyia rosenhaueri Rübsaamen (orig. des.). 

Pseudohormyia Shinji 1944: 248, missp. of Pseudhormomyia. 

Disharmomya Mani 1946: 227, missp. of Dishormomya. 

Holarctic; 51 spp., all known hosts belong to Carex or Cyperus (Cyperaceae); Cecidomyiidi, Hormomyiini. Ref.: 

Edwards 1938b (rev.). 

alexanderi. USA (New York). 

Hormomyia alexanderi Felt 1921a: 220. USA: New York, Johnstown; HT; ♀; NYSM. 


Hormomyia americana Felt 1907a: 28. USA: New York, Nassau; HT; ♂; NYSM. 

arenariae. Widespread Europe. Carex arenaria (Cyperaceae). 

Homomyia arenariae Rübsaamen 1899d: 602. Germany; ST; ♂,♀; ZMHB. 

atlantica. USA (New Jersey). 

Hormomyia atlantica Felt 1908b: 387. USA: New Jersey, Clementon; HT; ♀; NYSM. 

bambusae. Japan (Kyushu). Bambusa sp. (Poaceae). 

Trishormomyia bambusae Felt 1932a: 124. Japan: Kyushu, Oita Pref., Yasaka, Mura, Hayami-gun; ST; ♂; USNM. 

billoti. France. Carex davalliana (Cyperaceae). 

Hormomyia billoti Kieffer 1909b: 5. France: Strasbourg; T; LT; lost?. 

bremii. Switzerland. 

Hormomyia bremii Kieffer 1898f: 25, based on fasciata Meigen of Bremi 1847a. Switzerland; T; A; ETHZ?. 

brunnea. Switzerland. 

Hormomyia brunnea Rübsaamen 1892d: 396. Switzerland: Graubünden, Bergûn; T; ♂; ZMHB. 

caricis. UK, France, Germany, Czechoslovakia. Carex spp.; C. glauca, C. stellulata (Cyperaceae). 

Amaurosiphon caricis Rübsaamen 1911h: 391. Germany; ST; ♂,♀; ZMHB. 

Amaurosiphon baudysi Vimmer 1925: 112 (publ'n not seen). Czech Republic: i�kovec nr Ji ín; T; ?; ?. 

caudata. USA (Tennessee). Cyperus sp. (Cyperaceae). 

Hormomyia caudata Felt 1916e: 176. USA: Tennessee, Clarksville; ST; ♀,P; USNM. 

cincta. USA (New Hampshire). 

Hormomyia cincta Felt 1921a: 216. USA: New Hampshire, Hampton; HT; ♀; NYSM. 

coloradensis. USA (Colorado). 

327

328 

Hormomyia coloradensis Cockerell 1908c: 421. USA: Colorado, Boulder; HT; ♂; USNM. 


Hormomyia consobrina Felt 1907d: 18. USA: New York, Albany; HT; ♂; NYSM. 

cornifex. UK, France, Czechoslovakia, Romania. Carex pallescens, C. stricta (Cyperaceae). 

Hormomyia cornifex Kieffer 1898f: 60. France: Moselle; ST; ♂,♀; lost?. 

cornuta. Switzerland. 

Cecidomyia cornuta Bremi 1847a: 48. Switzerland; ST; ♀; ETHZ?. 

cucullata. Germany. 

Cecidomyia cucullata Meigen 1818: 96. Germany; T; ♂; MNHN. 

davisi. USA (New York). 

Dyodiplosis davisi Felt 1921a: 208. USA: New York, New York, Staten I.; HT; ♂; NYSM. 

dilatata. USA (New York). 

Trishormomyia dilatata Felt 1921a: 226. USA: New York, Newport; HT; ♂; NYSM. 

extrema. UK, Netherlands, Poland. 

Planetes extremus Walker 1837: 179. UK: England, nr London; ST; ♀; BMNH. 

Cecidomyia crassipes Loew 1850a: 31. Poland; ST; ♀; lost?. 

Cecidomyia funesta Walker 1856a: 120. UK: England; ST; ♂; BMNH. 

fenestra. USA (New York). 

Hormomyia fenestra Felt 1915e: 231. USA: New York, Fulton Co., Woodworth's L.; ST; ♂,♀; NYSM. 

fischeri. UK, Denmark, Germany, Poland, Czechoslovakia, Austria, Yugoslavia, Romania. Carex pilosa; C. rostrata 

(Cyperaceae). 

Hormomyia fischeri Frauenfeld 1867: 781. Austria: vic. Vienna; ST; ♂,♀,P,L; ?. 

Hormomyia hieronymi Kieffer 1909b: 5. Germany; T; LT; lost?. 

frireni. UK, France, Czechoslovakia, Yugoslavia. Carex pallescens, C. stricta (Cyperaceae). 

Hormomyia frireni Kieffer 1909b: 5. France: vic. Bitche; T; LT; lost?. 

fulva. USA (Massachusetts). 

Hormomyia fulva Felt 1926c: 208. USA: Massachusetts, Sherborn; HT; ♀; NYSM. 

gallarum. UK, Netherlands, Germany, Latvia, Lithuania, Poland, Czechoslovakia, Yugoslavia. Carex stricta, C. 

gracilis, C. goodenoughii (Cyperaceae). 

Dichrona gallarum Rübsaamen 1899b: 542. Germany; ST; ♂,♀,L; ZMHB. 

gibba. Sweden. 

Cecidomyza gibba Zetterstedt 1850: 3694. Sweden; T; ♀; MZLU?. 

gibbosa. Italy. 

Angelinia gibbosa Rondani 1860: 290. Italy: Parma; HT; ♂; MZUF. 

grandis. UK, Finland, Germany, Poland, Czechoslovakia, Austria, Russia (Europe). 

Cecidomyia grandis Meigen 1804: 39. Germany; T; A; lost. 

Cecidomyia fasciata Meigen 1818: 94. Germany; T; ♀; MNHN. 

Cecidomyia grossa Bremi 1847a: 46. Switzerland: vic. Zurich; ST; A,L; ETHZ?. 

Cecidomyia ampla Walker 1856a: 120. England; ST; ♂,♀; BMNH. 

Hormomyia dubitata Rübsaamen 1892d: 393. Poland; ST; ♀; ZMHB. 

granifex. UK, France, Germany, Poland, Czechoslovakia, Hungary, Romania, Yugoslavia. Carex stellulata, C. 

pallescens, C. stricta (Cyperaceae). 

Pseudhormomyia granifex Kieffer 1898f: 58. France: Moselle; ST; ♂,♀; lost?. 

incisa. USA (New York). 

Trishormomyia incisa Felt 1921a: 226. USA: New York, Nassau; HT; ♂; NYSM. 

johnsoni. USA (Massachusetts). 

Hormomyia johnsoni Felt 1907d: 18. USA: Massachusetts, Auburndale; HT; ♂; NYSM. 

kneuckeri. France. Carex stellulata (Cyperaceae). 

Hormomyia kneuckeri Kieffer 1909b: 6. France; T; G; lost?. 

lambertoni. France. 

Hormomyia lambertoni Kieffer 1901c: 25. France: Bitche; T; ♂; lost?. 

maxima. USA (New York).

329 

Hormomyia maxima Felt 1921a: 216. USA: New York, Johnstown; HT; ♀; NYSM. 

modesta. USA (Massachusetts). 

Hormomyia modesta Felt 1913f: 145. USA: Massachusetts, Auburndale; HT; ♀; NYSM. 

montana. USA (Colorado). 

Hormomyia montana Felt 1921a: 217. USA: Colorado; HT; ♀; USNM. 

needhami. USA (Illinois). 

Hormomyia needhami Felt 1907c: 160. USA: Illinois, Lake Forest; HT; ♂; NYSM. 


Hormomyia palustris Felt 1907d: 19. USA: New York, Ithaca; ST; ♂; NYSM. 

producta. UK, Netherlands, Germany. 

Cecidomyia producta Meigen 1830: 267. Germany; T; ♂; MNHN. 

proteana. USA (Massachusetts). 

Hormomyia proteana Felt 1914d: 113. USA: Massachusetts, Auburndale; HT; ♂; NYSM. 

pudica. USA (New Hampshire). 

Hormomyia pudica Felt 1913f: 146. USA: New Hampshire, Hampton; HT; ♀; NYSM. 

rosenhaueri. UK, Germany, Czechoslovakia. Carex acuta (Cyperaceae). 

Hormomyia rosenhaueri Rübsaamen 1892d: 394. Germany; ST; ♀,P,L; ZMHB. 

saturni. USA (New York). 

Hormomyia saturni Felt 1914c: 133. USA: New York, Adirondack Mts; HT; ♂; NYSM. 

shawi. USA (New Hampshire). 

Hormomyia shawi Felt 1913f: 145. USA: New Hampshire, Hampton; HT; ♂; NYSM. 

strobli. Austria. 

Hormomyia strobli Kieffer 1901c: 26. Austria: Styria; T; ♂; NMBA?. 

subaptera. India (Tamil Nadu). 

Hormomyia subaptera Felt 1926i: 244. India: Tamil Nadu, Nilgiri Hills, Ootacamund, Snowdon Peak; HT; ♀; 

NYSM. 

subterranea. UK, Czechoslovakia, Italy. Carex divulsa (Cyperaceae). 

Pseudhormomyia subterranea Kieffer & Trotter 1904b: 65. Italy: vic. Avellino; ST; ♀,P,L; lost?. 

tarda. Germany. Carex vesicaria (Cyperaceae). 

Jaapiola tarda Rübsaamen 1914: 95. Germany: Prignitz, Triglitz; ST; ♂,♀,L; ZMHB. 

tuberifica. UK, Germany, Czechoslovakia. Carex stricta (Cyperaceae). 

Hormomyia tuberifica Rübsaamen 1899b: 546, fig.7. Germany; ST; ♀; ZMHB. 

tumorifica. Germany, Czechoslovakia, Italy. Carex pseudocyperus (Cyperaceae). 

Hormomyia tumorifica Rübsaamen 1899d: 603. Germany; ST; ♂,♀; ZMHB. 

westermanni. Germany. 

Cecidomyia westermanni Meigen 1830: 269. Germany: nr Kiel; T; ♂; lost. 

winnertzi. UK, Germany. 

Hormomyia winnertzi Kieffer 1898f: 26. Germany; ST; ♂,♀; lost?. 

PLANODIPLOSIS 

Planodiplosis Kieffer 1911: 323. Type species, insularis Kieffer (mon.). 


insularis. Seychelles Is. 

Planodiplosis insularis Kieffer 1911: 323. Seychelles Is: Mahé, Cascade Estate; HT; ♀; BMNH. 

PLATYDIPLOSIS 

Platydiplosis Gagné 1973c: 877. Type species, nigricauda Gagné (orig. des.).


nigricauda. USA (Maryland). 

Platydiplosis nigricauda Gagné 1973c: 878. USA: Maryland, Montgomery Co., Seneca St. Pk; HT; ♂; USNM. 

PLECOPHORUS 

Plecophorus Kieffer 1913h: 104. Type species, fuscocinctus Kieffer (orig. des.). 


fuscocinctus. Tanzania. 

Plecophorus fuscocinctus Kieffer 1913h: 105. Tanzania: Neu-Moschi, lower elevations Mt Kilimanjaro; HT; ♀; 

MNHN. 

PLECTRODIPLOSIS 

Plectrodiplosis Gagné 1973c: 878. Type species, fascipennis Gagné (orig. des.). 


fascipennis. USA (Maryland). 

Plectrodiplosis fascipennis Gagné 1973c: 879. USA: Maryland, Montgomery Co., Plummer's I.; HT; ♂; USNM. 

PLEMELIELLA 

Plemeliella Seitner 1908: 185. Type species, abietina Seitner (mon.). 

Palearctic; 1 sp.; Cecidomyiidi, Cecidomyiini. 

abietina. Sweden, Finland, Germany, Poland, Czechoslovakia, Austria, Slovenia, Hungary, Ukraine, Russia (Europe). 

Picea abies; P. pubescens, P. excelsa, P. pungens (Pinaceae). 

Plemeliella abietina Seitner 1908: 185. Slovenia: Idria, & Carpathian Mts; ST; ♂,♀,P,L; Seitner Coll. 

PLESIODIPLOSIS 

Plesiodiplosis Kieffer 1912h: 1. Type species, Clinodiplosis chilensis Kieffer & Herbst (orig. des.). 


chilensis. Chile. 

Clinodiplosis chilensis Kieffer & Herbst 1906: 231. Chile: vic. Concepción; ST; ♂,♀; lost. 

PLESIOLAUTHIA 

Plesiolauthia Mamaev 1967: 880. Type species, crassicornis Mamaev (orig. des.). 

Palearctic; 1 sp.; Lasiopteridi, Ledomyiini. 

crassicornis. Russia (Europe). 

Plesiolauthia crassicornis Mamaev 1967: 881. Russia: Voronezh Prov., Tellerman For.; HT; ♂; IEME. 

330

PLUTODIPLOSIS 

Plutodiplosis Kieffer 1912h: 27. Type species, magnifica Kieffer (mon.). 


heterofila. India (Uttar Pradesh). 

Plutodiplosis heterofila Grover & Bakhshi 1978: 160. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; PGCA. 

magnifica. Sri Lanka. 

Plutodiplosis magnificus Kieffer 1912a: 27. Sri Lanka: Peradeniya; HT; ♂; NZSI?. 

POLYGONOMYIA 

Polygonomyia Fedotova 1991d: 23. Type species, Dasineura atraphaxiflorae Fedotova (orig. des.). 

Palearctic; 2 spp., on Polygonaceae; Lasiopteridi, Dasineurini. 

alabastris. Kazakhstan. Polygonum soongoricum; P. lapathifolium, P. undulatum (Polygonaceae). 

Polygonomyia alabastris Fedotova 1991d: 23. Kazakhstan: Dzhungarskiï Alatau, floodplain Lepsy R., nr Pokrovka- 

Cherkasskaya; HT; ♂; ZMAS. 

atraphaxiflorae. Kazakhstan. Atraphaxis frutescens; Atraphaxis spp. (Polygonaceae). 

Dasyneura atraphaxiflorae Fedotova 1984f: 642. Kazakhstan: northern Balkash area, Katanemel' Mtn; HT; ♂; ?. 

POLYSTEPHA 

Polystepha Kieffer 1897c: 11. Type species, quercus Kieffer (orig. des.). 

Cincticornia Felt 1908b: 379. Type species, Asphondylia transversa Felt (orig. des.). 

Holarctic and south into Central America; 24 spp., probably all leaf gall makers on Quercus; Cecidomyiidi, 

Asphondyliini, Schizomyiina. 

aceris. USA (New York). 

Oligotrophus aceris Felt 1907a: 26. USA: New York, Albany; HT; ♂; NYSM. 

americana. USA (New York). Quercus sp. (Fagaceae). 

Cincticornia americana Felt 1908b: 380. USA: New York, Albany; ST; ♂,♀; NYSM. 

canadensis. Canada (Ontario). Quercus sp. (Fagaceae). 

Cincticornia canadensis Felt 1908b: 380. Canada: Ontario, Toronto; HT; ♀; USNM. 

caryae. USA (Massachusetts, Maine). Quercus sp. (Fagaceae). 

Cincticornia caryae Felt 1908b: 380. USA: Massachusetts, Worcester; HT; ♂; NYSM. 

connecta. USA (Connecticut). 

Cincticornia connecta Felt 1908b: 381. USA: Connecticut, Westville; HT; ♂; NYSM. 

cornifolia. USA (New York). 

Oligotrophus cornifolius Felt 1907a: 28. USA: New York, Albany; HT; ♀; NYSM. 

globosa. USA (Massachusetts, Connecticut). Quercus sp. (Fagaceae). 

Cincticornia globosa Felt 1909b: 291. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

lapalmae. El Salvador. Quercus hondurensis (Fagaceae). 

Polystepha lapalmae Möhn 1960d: 335. El Salvador: Chalatenango, rd to La Palma, ca. 15 km before La Palma; HT; 

LT; SMNS. 

malpighii. UK, Sweden, France, Germany, Czechoslovakia, Yugoslavia, Greece. Quercus petraea, Q. robur 

(Fagaceae). 

Perrisia malpighii Kieffer 1909b: 21. France; T; G; lost?. 

multifila. USA (New York). 

331

332 

Asphondylia multifila Felt 1907a: 22. USA: New York, Albany; HT; ♂; NYSM. 

pilulae. USA (Illinois, New York, Maine, Connecticut, Maryland). Quercus spp. (Fagaceae). 

Cecidomyia quercus pilulae Walsh 1864a: 481, invalid, a polynomial. 

Cecidomyia pilulae Beutenmüller 1892: 269. USA: New York, vic. New York; ST; G; AMNH. 

podagrae. USA (Massachusetts). Quercus velutina (Fagaceae). 

Cincticornia podagrae Felt 1909b: 291. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

pustulata. USA (Massachusetts). Quercus velutina (Fagaceae). 

Cincticornia pustulata Felt 1909b: 291. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

pustuloides. USA (widespread eastern). Quercus spp. (Fagaceae). 

Cecidomyia pustuloides Beutenmüller 1907a: 390. USA: New Jersey, Lakehurst, & North Carolina, Buncombe Co., 

Valley of the Black Mts; ST; LT; AMNH. 

quercifolia. USA (District of Columbia). Quercus sp. (Fagaceae). 

Cincticornia quercifolia Felt 1908b: 380. USA: District of Columbia; ST; ♂,♀; USNM. 

quercus. Spain, France, Latvia, Lithuania, Czechoslovakia, Russia (Europe). Quercus sp.; Q. robur (Fagaceae). 

Polystepha quercus Kieffer 1897c: 11. France: Bitche; T; A; lost?. 

Polystepha titilivii (as titi-livii) Tavares 1924a: 38. Spain: Gallicia, vic. La Guardia; ST; ♂,♀; TAVC. 

rhoina. USA (New York). 

Oligotrophus rhoina Felt 1907a: 26. USA: New York, Albany; HT; ♂; NYSM. 

rossica. Russia (Europe). Quercus sp. (Fagaceae). 

Polystepha rossica Mamaeva 1981: 405. Russia: Moscow Prov., Sofrino; HT; ♂; IEME. 

salvadorensis. El Salvador. Quercus eugeniaefolia (Fagaceae). 

Polystepha salvadorensis Möhn 1960d: 333. El Salvador: Chalatenango, rd to La Palma, ca. 15 km before La Palma; 

HT; ♂; SMNS. 

serrata. USA (Massachusetts). Quercus sp. (Fagaceae). 

Cincticornia serrata Felt 1908b: 380. USA: Massachusetts, nr Boston; ST; ♂; NYSM. 

simpla. USA (New York, Massachusetts). Quercus velutina (Fagaceae). 

Cincticornia simpla Felt 1909b: 291. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

sobrina. USA (Massachusetts). Quercus velutina (Fagaceae). 

Asphondylia sobrina Felt 1907c: 158. USA: Massachusetts, Worcester; ST; ♂,♀; NYSM. 

symmetrica. USA (widespread eastern). Quercus sp.; Quercus spp. (Fagaceae). 

Cecidomyia symmetrica Osten Sacken 1862: 200. USA: District of Columbia; T; LT; lost. 

transversa. USA (New York). 

Asphondylia transversa Felt 1907a: 22. USA: New York, Albany; HT; ♂; NYSM. 

*PORIDIPLOSIS 

Poridiplosis Fedotova 2006a: 144. Type species, semiaperta Fedotova (orig. des.). 


*semiaperta. Russia (Far East). 

Poridiplosis semiaperta Fedotova 2006a: 147. Russia: Primorskiï Terr., Vladivostok; HT; ♂; ZMAS. 

POTENTILLOMYIA 

Potentillomyia Fedotova 1990f: 89. Type species, kasenasae Fedotova (orig. des.). 


kazenasae. Kazakhstan. Potentilla bifurca (Rosaceae). 

Potentillomyia kazenasae Fedotova 1990f: 89. Kazakhstan: Karatal R. valley, 60 km NW Ush-Tobe; HT; ♂; ZMAS.

PRIMAVERA 

Primavera Gagné 2002a: 40. Type species, Rhabdophaga porrecta Felt (orig. des.). 

Eastern Nearctic; 1 sp.; Lasiopteridi, unplaced to tribe. Ref.: Gagné 2002a, redescr. 

porrecta. USA (Minnesota, Wisconsin, Iowa, New York, West Virginia, Maryland). Ulmus americana (Ulmaceae). 

Rhabdophaga porrecta Felt 1915d: 101. USA: New York, Gloversville; HT; ♂; NYSM. 

PRIMOFAVILLA 

Primofavilla Mamaev 1972a: 887. Type species, initialis Mamaev (orig. des.). 

Palearctic; 3 spp., on Chenopodiaceae; Lasiopteridi, Lasiopterini. 

cystiphorae. Kazakhstan. Atriplex tatarica (Chenopodiaceae). 

Primofavilla cystiphorae Fedotova 1991b: 47. Kazakhstan: Berkutinka R. valley nr Zhanatas; HT; ♀; ZMAS. 

initialis. Armenia, Kazakhstan. Atriplex salina (Chenopodiaceae). 

Primofavilla initialis Mamaev 1972a: 888. Kazakhstan: Bugun'; HT; ♀; HYNM. 

kaplini. Kazakhstan. Atriplex nana (Chenopodiaceae). 

Primofavilla kaplini Fedotova 1991b: 45. Kazakhstan: western Ustyurt, Kartyuleïskaya lowland, Sarshanaï valley, 34 

km SE Akzhigit; HT; ♀; ZMAS. 

PROASPHONDYLIA 

Proasphondylia Felt 1915a: 203. Type species, brasiliensis Felt (orig. des.). 

Neotropical; 3 spp., the 2 reared species both from Nyctaginaceae; Cecidomyiidi, Asphondyliini, Schizomyiina. 

brasiliensis. Brazil (Pernambuco). 

Proasphondylia brasiliensis Felt 1915a: 203. Brazil: Pernambuco, Bonito; ST; ♂,♀,P; USNM. 

formosa. Brazil (Rio de Janeiro). Guapira opposita (Nyctaginaceae). 

Proasphondylia formosa Maia 1994a: 217. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

guapirae. Brazil (Rio de Janeiro). Guapira opposita (Nyctaginaceae). 

Proasphondylia guapirae Maia 1994a: 216. Brazil: Rio de Janeiro, Arraial do Cabo; HT; ♂; QBUM. 

PROBRUGGMANNIELLA 

Probruggmanniella Möhn 1961a: 1. Type species, Schizomyia phillyreae Tavares (orig. des.). 

Southwestern Palearctic; 1 sp.; Cecidomyiidi, Asphondyliini, Asphondyliina. 

phillyreae. Portugal, France, Italy, Yugoslavia, Greece, Turkey, Algeria. Phillyrea angustifolia (Oleaceae). 

Schizomyia phillyreae Tavares 1907a: 52. Portugal: Vergão & Cardigos; ST; ♂,♀,P; TAVC. 

PROCONTARINIA 

Procontarinia Kieffer & Cecconi 1906: 135. Type species, matteiana Kieffer & Cecconi (orig. des. as n. g, n. sp.). 

Erosomyia Felt 1911c: 49. Type species, mangiferae Felt (orig. des.). 

Indodiplosis Felt 1916g: 403. Type species, mangiferae Felt (orig. des.). 

Mangodiplosis Tavares 1918a: 47. Type species, mangiferae Tavares (orig. des.) = mangiferae Felt. 

Amradiplosis Mani 1947: 441. Type species, echinogalliperda Mani (orig. des.). 

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Amraemyia Rao 1950: 33. Type species, viridigallicola Rao (orig. des.). 

Oriental, immigr. Nearctic; 15 spp., all associated with mango; Cecidomyiidi, Cecidomyiini. Refs: Gagné & Medina 

2004 (review, synonymy), Harris 1966 (illus.), Harris & Schreiner 1992 (illus., biol.). 

allahabadensis. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Amraemyia allahabadensis Grover 1962c: 312. India: Uttar Pradesh, Allahabad, Tagore Town; HT; ♂; PGCA. 

amraeomyia. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Amraemyia amraeomyia Rao 1950: 37. India: Uttar Pradesh, Kanpur; HT; ♂; ZSCI. 

Amraemyia amraemyia Grover 1962c: 312, missp. amraeomyia Rao. 

biharana. India (Bihar). Mangifera indica (Anacardiaceae). 

Indodiplosis mangiferae Felt 1916g: 403, jun. sec. hom. of Erosomyia mangiferae Felt. India: Bihar, Pusa; ST; 

♂,♀,P; NYSM. 

Procontarinia biharana Gagné 2004a: 24, new name for Indodiplosis mangiferae Felt. 

brunneigallicola. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Amraemyia brunneigallicola Rao 1950: 39. India: Uttar Pradesh, Kanpur; HT; ♂; ZSCI. 

echinogalliperda. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Amradiplosis echinogalliperda Mani 1947: 443. India: Uttar Pradesh, Dehra Dun; HT; ♀; ZSCI. 

*frugivora. Philippines (Luzon). Mangifera indica (Anacardiaceae). 

Procontarinia frugivora Gagné 2004a: 20. Philippines: Luzon, Bulacan Prov., Alagao; HT; ♂; University of 

Philippines Museum of Natural History, Los Baños, Laguna, Philippines. 

keshopurensis. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Amraemyia keshopurensis Rao 1952a: 52. India: Uttar Pradesh, Keshopur; HT; ♂; ZSCI. 

mangicola. China (Guangxi Zhuang Aut. Reg.); immigr.: Japan (Okinawa), Guam. Mangifera indica (Anacardiaceae). 

Erosomyia mangicola Shi 1980: 131. China: Guangxi Zhuang Aut. Reg., Nanning; HT; ♂; IEAS. Ref.: Uechi et al. 

2002. 

Procontarinia schreineri Harris 1992: 42. Guam: Barrigada; HT; ♂; BMNH. 

mangiferae. India, Réunion; immigr.: Iran, Guadeloupe, St. Lucia, St. Vincent, Trinidad, Brazil (Bahia). Mangifera 

indica (Anacardiaceae). Ref.: Pezhman & Askari 2004 (biol.). 

Erosomyia mangiferae Felt 1911c: 49. St. Vincent; ST; ♂,♀,P,L; USNM. 

Mangodiplosis mangiferae Tavares 1918a: 48. Brazil, Bahia, Salvador, Roça da Madre de Deus; ST; ♂,♀,P,L; 

TAVC. 

Rhabdophaga mangiferae Mani 1938: 331, jun. sec. hom. mangiferae Felt. India: Kerala, Trichur; HT; ♀; INPC. 

Erosomyia indica Grover & Prasad 1966a: 7. India: Tamil Nadu, Coimbatore; ST; ♂,♀; PGCA. 

mangifoliae. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Indodiplosis mangifoliae Grover 1965b: 115. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

matteiana. Kenya, South Africa, Mauritius, Réunion, India, Java. Mangifera indica (Anacardiaceae). 

Procontarinia matteiana Kieffer & Cecconi 1906: 135. Italy: Sicily, Palermo, Botanic Gardens; ST; ♂,♀,L; lost?. 

*pustulata. Australia (QLD), Papua New Guinea). Mangifera indica (Anacardiaceae). 

Procontarinia pustulata Kolesic 2009: 312. Australia: Queensland, Torres Strait, Badu I.; HT; ♂; SAMA. 

*robusta. China (Fujian). Mangifera indica (Anacardiaceae). 

Procontarinia robusta Li, Bu & Zhang 2003: 148. China: Fujian, Xiamen, Xinglin District; HT; ♂; NKUM. 

tenuispatha. India (West Bengal). Mangifera indica (Anacardiaceae). 

Oligotrophus tenuispatha Kieffer 1909a: 150. India: West Bengal, nr Ranchi; T; LT; lost?. A neotype designated by 

Nayar 1945: 72 is invalid because there was no indication of how it differed from related species. 

viridigallicola. India (Uttar Pradesh). Mangifera indica (Anacardiaceae). 

Amraemyia viridigallicola Rao 1950: 34. India: Uttar Pradesh, Kanpur; HT; ♂; ZSCI. 

PROCYSTIPHORA 

Procystiphora Felt 1915d: 212. Type species, coloradensis Felt (orig. des.). 

334

335 

Holarctic; 4 spp., on Cyperaceae, Juncaceae, and Poaceae; Lasiopteridi, unplaced to tribe. Ref.: Sato et al. 2009 

(review). 

coloradensis. USA (Colorado). ?Carex (Cyperaceae). 

Procystiphora coloradensis Felt 1915d: 212. USA: Colorado, Long's Peak Inn; ST; ♂,♀; NYSM. 

gerardi. Germany. Juncus gerardi (Juncaceae). 

Procystiphora gerardi Meyer 1984: 35. Germany: Schleswig-Holstein, Hauke-Haien-Koog; HT; ♂; ZMKG. 

junci. USA (Illinois). Juncus sp. (Juncaceae). 

Procystiphora junci Felt 1922c: 166. USA: Illinois, Centralia; ST; ♂,♀; NYSM. 

*uedai. Japan (Honshu). Sasa nipponica (Poaceae). 

Procystiphora uedai Sato & Yukawa 2009: 43. Japan: Honshu, Nara Pref., Kamikitayama Village, Mt Ôdaigahara; 


PRODIPLOSIS 

Prodiplosis Felt 1908b: 403. Type species, Cecidomyia floricola Felt (orig. des.). 

Geisenheyneria Rübsaamen 1910d: 289. Type species, rhenana Rübsaamen (orig. des. as n. g., n. sp.). 

Holarctic, Neotropical; 12 spp.; Cecidomyiidi, Cecidomyiini. This is a morphologically and biologically diverse 

genus that includes polyphagous, economically important species. Refs: Gagné 1986b (revision). 

citrulli. USA (Arizona). Citrullus sp., Cucumis sp.; Cucurbita spp. (Cucurbitaceae). 

Itonida citrulli Felt 1935e: 79. USA: Arizona, Tucson; LT (Gagné 1973c: 879); ♂; USNM. 

falcata. USA (California). Eriogonum latifolium (Polygonaceae); Ericameria ericoides (Asteraceae). 

Prodiplosis falcata Gagné 1986b: 244. USA: California, Montana; HT; ♂; USNM. 

floricola. USA (New York), Brazil. Spiraea salicifolia (Rosaceae); Clematis sp. (Ranunculaceae), Caryocar 

brasiliense (Caryocaraceae). 

Cecidomyia floricola Felt 1907d: 21. USA: New York, Albany; LT (Gagné 1986b: 242); ♂; NYSM. 

longifila. USA (Virginia, Florida), Colombia, Ecuador, Peru. Gossypium sp. (Malvaceae); Lycopersicon esculentum & 

Solanum tuberosum (Solanaceae), Citrus aurantifolia (Rutaceae), Medicago sativa & Phaseolus sp. 

(Fabaceae), Chenopodium ambrosioides (Chenopodiaceae), & Ricinus communis (Euphorbiaceae). 

Prodiplosis longifila Gagné 1986b: 240. USA: Florida, Monroe Co., Marco I.; HT; ♂; USNM. 

morrisi. Widespread eastern Nearctic. Populus deltoides; P. tremuloides (Salicaceae). 

Prodiplosis morrisi Gagné 1966a: 1155. USA: MS, Stoneville; HT; ♂; USNM. 

myricae. USA (New Jersey). Myrica cerifera (Myricaceae). 

Cecidomyia myricae Beutenmüller 1907b: 306. USA: New Jersey, Carlstadt; LT (Gagné 1986b: 237); ♂; AMNH. 

orientalis. Kazakhstan. Erigeron pseudoseravschanicus (Asteraceae). 

Geisenheyneria orientalis Fedotova 1995c: 54. Kazakhstan, foothills Saur Mtn Range, 20 km S Zaïsan, nr 

Zhanturmys; HT; ♂; ZMAS. 

platani. USA (New York, Maryland). Platanus occidentalis (Platanaceae). 

Prodiplosis platani Gagné 1986b: 237. USA: Maryland, Beltsville; HT; ♂; USNM. 

rhenana. Germany, Hungary. Erigeron acer (Asteraceae). 

Geisenheyneria rhenana Rübsaamen 1910d: 289. Germany; T; ♂; ZMHB. 

saurica. Kazakhstan. Erigeron acris (Asteraceae). 

Geisenheyneria saurica Fedotova 1995c: 56. Kazakhstan, southern Altaï, northeastern shore L. Markakol', 10 km N 


vaccinii. USA (Pennsylvania, New Jersey). Vaccinium sp. (Ericaceae). 

Contarinia vaccinii Felt 1926a: 85. USA: New Jersey, Whitesbog; LT (Gagné 1986b: 240); ♂; NYSM. 

violicola. USA (Minnesota, Michigan, New York, Connecticut, New Jersey, Virginia), Sweden, Netherlands. Viola 

spp. (Violaceae). 

Diplosis violicola Coquillett 1900b: 50. USA: District of Columbia; LT (Gagné 1986b: 244); ♂; USNM.

PROLAUTHIA 

Prolauthia Rübsaamen 1916b: 505. Type species, Cecidomyia circumdata Winnertz (mon.). 

Prolanthia Neave 1940: 913, missp. of Prolauthia. 

Western Palearctic; 1 sp.; Lasiopteridi, Ledomyiini. 

circumdata. UK, Germany. Crataegus oxyacantha (Rosaceae). 

Cecidomyia circumdata Winnertz 1853: 226. Germany; ST; ♂,♀,L; lost. 

PROMIKIOLA 

Promikiola Kieffer & Herbst 1911: 700. Type species, rubra Kieffer & Herbst (orig. des. as n. g., n. sp.). 


rubra. Chile. Colliguaja odorifera (Euphorbiaceae). 

Promikiola rubra Kieffer & Herbst 1911: 700. Chile: Valparaíso; ST; ♂,♀,P; lost. 

*PRORSODIPLOSIS 

Prorsodiplosis Fedotova & Sidorenko 2009a: 11. Type species, credula Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 2 spp.; Cecidomyiidi, Karshomyiini. This genus is close to Karshomyia if the two must be 

separated. 

*credula. Russia (Far East). 

Prorsodiplosis credula Fedotova & Sidorenko 2009a: 11. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


dubia. Russia (Far East). 

Holoneurus dubius Kovalev & Mamaev 1966: 232. Russia: Primorskiï Terr., Chapigou; HT; ♂; ZMMU. Ref.: 

Fedotova & Sidorenko 2009a: 14 (n. comb.). 

PROTAPLONYX 

Protaplonyx Felt 1916c: 202. Type species, hagani Felt (orig. des.). 


sarcobati. USA (Idaho, Utah, Colorado). Sarcobatus vermiculatus (Chenopodiaceae). 

Aplonyx sarcobati Felt 1914e: 93. USA: Colorado, Canyon City; LT (Gagné 1969d: 1355); ♀; NYSM. 

Protaplonyx hagani Felt 1916c: 202. USA: Utah, Wellington, nr Price; LT (Gagné 1969d: 1355); ♀; NYSM. 

PROTERODIPLOSIS 

Proterodiplosis Wyatt 1963: 103. Type species, radicis Wyatt (orig. des.). 


radicis. New Zealand. Metrosideros robusta (Myrtaceae). 

Proterodiplosis radicis Wyatt 1963: 103. New Zealand: Whakarewarewa St. For., Rotorua area; HT; ♂; FRNZ. 

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PRUTHIDIPLOSIS 

Pruthidiplosis Mani 1934b: 389. Type species, mimusopsicola Mani (orig. des.). 


mimusopsicola. India (Tamil Nadu). Mimusops hexandra (Sapotaceae). 

Pruthidiplosis mimusopsicola Mani 1934b: 390. India: Tamil Nadu, Thanjavur; ST; ♂; ZSCI. 

PSECTROSEMA 

Psectrosema Kieffer 1904b: 343. Type species, Cecidomyia tamaricis De Stefani (orig. des.). 

Amblardiella Kieffer 1912g: 169. Type species, Oligotrophus tamaricum Kieffer (mon.). 

Isosandalum Kieffer 1913i: 40. Type species, Psectrosema provinciale Kieffer (mon.). 

Mamaeviana Fedotova 1992m: 91, as subg. of Psectrosema. Type species, Psectrosema diversicorne Mamaev & 

Becknazharova (mon.). 

Harrisiana Fedotova 1992m: 91 (as n. g. also in Fedotova 1993a: 64). Type species, mamaevi Fedotova (mon.). 

Marikovskiana Fedotova 1992m: 91 (as n. g. also in Fedotova 1993a: 66). Type species, Isosandalum dentipes 

Marikovskij (orig. des.). 

Debskiana Mamaev & Becknazarova 1983: 69, as subg. of Psectrosema. Type species, Psectrosema turkmenicum 

Mamaev & Becknazarova (orig. des.). 

Ptectrosema Shinji 1944: 225, missp. of Psectrosema. 

Palearctic; 26 spp., all on Tamarix; Lasiopteridi, unplaced to tribe. Ref.: Gagné et al. 1996 (rev.). 

acuticorne. France. Tamarix gallica (Tamaricaceae). 

Psectrosema acuticorne Gagné 1996c: 63. France: Aigues Mortes, 30 km SE Montpellier; HT; P; USNM. 

album. France. Tamarix sp. (Tamaricaceae). 

Psectrosema album Gagné 1996c: 62. France: 15 km SE Montpellier; HT; P; USNM. 

alfierii. Egypt. Tamarix arborea (Tamaricaceae). 

Psectrosema alfierii Debski 1922: 36. Egypt: Cairo; ST; ♂,♀,P,L; USNM. 

barbatum. Kazakhstan, Uzbekistan. Tamarix ramosissima (Tamaricaceae). 

Isosandalum barbatum Marikovskij 1961: 37. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, vic. middle & lower 


becknazarovae. Kazakhstan. Tamarix hispida (Tamaricaceae). 

Psectrosema becknazarovae Fedotova 1992m: 96. Kazakhstan: Achi-Say R. valley, 40 km SW Abay, 30 km NE 

Tashkent; HT; ♂; ZMAS. 

debskii. Egypt. Tamarix nilotica (Tamaricaceae). 

Cecidomyia debskii Kieffer 1912g: 171. Egypt: Hélouan, nr Cairo; ST; L,P,F,egg; lost?. 

dentipes. Armenia, Kazakhstan. Tamarix ramosissima (Tamaricaceae). 

Isosandalum dentipes Marikovskij 1961: 39. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, vic. middle & lower 


diversicorne. Turkmenistan. Tamarix sp. (Tamaricaceae). 

Psectrosema diversicornis Mamaev & Becknazharova 1983: 68. Turkmenistan: Mary, Akibay; HT; ♀; HYNM. 

grummgrzhimajloi. Kazakhstan. Tamarix ramosissima (Tamaricaceae). 

Isosandalum grummgrzhimajloi (as Grumm-Grzhimajloi) Fedotova 1983a: 35. Kazakhstan: southwestern banks L. 

Balkhash, 10 km S Mirnyï; HT; ♂; ZKAS. 

iliense. Kazakhstan. Tamarix gracilis, T. ramosissima, T. leptostachys (Tamaricaceae). 

Isosandalum iliense Marikovskij 1961: 40. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, vic. middle & lower 


indicum. Pakistan. Tamarix indica (Tamaricaceae). 

Psectrosema indicum Harris 1983: 451. Pakistan: Muzaffargarh; HT; ♂; BMNH. 

337

338 

mamaevi. Turkmenistan. Tamarix laxa; T. kotschyi, T. ramosissima (Tamaricaceae). 

Harrisiana mamaevi Fedotova 1992m: 92 (as n. sp. also in Fedotova 1993a: 65). Turkmenistan: southern Ustyurt, 

Kaplankyrskiï Res., 300 km W Tashauz; HT; ♂; ZMAS. 

manii. Pakistan (Multan). Tamarix dioica (Tamaricaceae). 

Misospatha tamaricis Mani 1935: 432, jun. sec. hom. tamaricis Tavares. Pakistan: Multan; ST; ♂,♀; ZSCI. 

Psectrosema manii Harris 1983: 451, new name for Misospatha tamaricis Mani. 

mitjaevi. Kazakhstan. Tamarix ramosissima (Tamaricaceae). 

Isosandalum mitjaevi Marikovskij 1961: 38. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, vic. middle & lower 

reaches Ili R.; ST; ♂; ZMAS. 

nigrum. France. Tamarix parviflora; T. gallica, T. ramosissima (Tamaricaceae). 

Psectrosema nigrum Gagné 1996c: 59. France: 15 km SE Montpellier; HT; P; USNM. 

noxium. Kazakhstan. Tamarix gracilis, T. ramosissima, T. leptostachys (Tamaricaceae). 

Isosandalum noxium Marikovskij 1961: 41. Kazakhstan: Alma-Ata & Taldy-Kurgan Regions, vic. middle & lower 


parvum. Pakistan. Tamarix dioica; T. indica (Tamaricaceae). 

Psectrosema parvum Harris 1983: 454. Pakistan: Lahore; HT; ♂; BMNH. 

provinciale. France, Portugal. Tamarix gallica (Tamaricaceae). 

Psectrosema provinciale Kieffer 1912g: 171. France: Marseille; ST; ♂,♀,P,L; lost?. 

reticulatum. Pakistan. Tamarix hispida; T. androssowii, T. arceuthoides (Tamaricaceae). 

Psectrosema reticulatum Harris 1983: 452. Pakistan: Jungle Thana; HT; ♂; BMNH. 

squamosum. Turkmenistan. Tamarix sp. (Tamaricaceae). 

Isosandalum squamosum Marikovskij 1955e: 21. Turkmenistan: northeastern Karakumy Desert; ST; ♂,♀; ?. 

tamaricinum. Egypt. 

Perrisia tamaricina Kieffer 1909b: 30. Egypt: Wadi Tarfa; ST; G; lost?. 

tamaricis. UK, Portugal, Italy, Tunisia. Tamarix gallica, T. tetrandra (Tamaricaceae). 

Rhopalomyia tamaricis De Stefani 1902: 114. Italy: Sicily, Catania & Palermo; ST; ♂,♀,P,L; lost. 

tamaricum. Algeria, Tunisia, Libya, Egypt. Tamarix africana (Tamaricaceae). 

Oligotrophus tamaricum Kieffer 1912e: 126. Algeria: St. Denis-du-Sig, nr Oran; ST; P; MNHN?. A neotype series, 

not a neotype, for this species was invalidly designated by Grover 1972: 6. When a neotype is designated for 

this species, it should come from a place closer to the original locality than India and also from the original 

host. 

turkmenicum. Turkmenistan. Tamarix sp. (Tamaricaceae). 

Psectrosema turkmenica Mamaev & Becknazharova 1983: 69. Turkmenistan: Mary, Akibay; HT; ♀; HYNM. 

unicorne. Pakistan. Tamarix dioica (Tamaricaceae). 

Psectrosema unicorne Harris 1983: 454. Pakistan: Lahore; HT; ♂; BMNH. 

xinjiangense. China (Xinjiang Uygur Aut. Reg.). Tamarix sp. (Tamaricaceae). 

Psectrosema xinjiangense Bu & Zheng 1991: 63. China: Xinjiang Uygur Aut. Reg., Ta-cheng; HT; ♂; NKUM. 

PSEUDASPHONDYLIA 

Pseudasphondylia Monzen 1955b: 34. Type species, rokuharensis Monzen (orig. des.). 

Philadelphella Kovalev 1964: 440. Type species, philadelphi Kovalev (orig. des.). 

Eastern Palearctic, Oriental, and Oceanian; 10 spp., on as many families of plants; Cecidomyiidi, Asphondyliini, 

Asphondyliina. Ref.: Tokuda & Yukawa 2005 (Japanese species). 

campanulata. India (Himachal Pradesh). Sabia campanulata (Sabiaceae). 

Acroectasis campanulata Mani 1954a: 17. India: Himachal Pradesh, Kalatop Res. For., nr Dalhousie; HT; ♀; ZSCI. 

diospyri. China (Zhejiang). Diospyros kaki (Ebenaceae). 

Pseudasphondylia diospyri Mo & Xu 1999: 36. China: Zhejiang, Lanxi; HT; ♂; SAUT. 

*elaeocarpi. Japan (Honshu to Okinawa). Elaeocarpus sylvestris (Elaeocarpaceae). 

Pseudasphondylia elaeocarpi Tokuda & Yukawa 2005: 265. Japan: Kyushu, Kagoshima, Shiroyama; HT; ♂; KUEC.

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*kiritanii. Japan (Honshu). Cornus controversa (Cornaceae). 

Pseudasphondylia kiritanii Tokuda & Yukawa 2005: 261. Japan: Honshu, Shizuoka, Ito, Futo; HT; ♂; KUEC. 

matatabi. Japan (Honshu, Kyushu). Actinidia polygama (Actinidiaceae). 

Asphondylia matatabi Yuasa & Kumazawa 1938: 204. Japan: Honshu, Mt. Takao, nr Tokyo; ST; ♂,♀; Natl Inst. for 

Agro-Environmental Sciences, Tsukuba, Japan. 

neolitseae. Japan (Shikoku, Kyushu), Republic of Korea. Neolitsea sericea (Lauraceae). 

Pseudasphondylia neolitseae Yukawa 1974: 294. Japan: Kyushu, Kagoshima, Shiroyama; HT; ♂; KUEC. 

philadelphi. Russia (Far East). Philadelphus tenuifolius (Hydrangeaceae). 

Philadelphella philadelphi Kovalev 1964: 441. Russia: Primorskiï Terr., Okeanskaya Station, nr Vladivostok; HT; ♂; 

KOVC. 

rauwolfiae. New Caledonia. Rauwolfia schumanniana (Apocynaceae). 

Pseudasphondylia rauwolfiae Coutin 1980: 502. New Caledonia: R. des Pirogues, for. de Faux Bon Secours; HT; ♂; 

INRV. 

rokuharensis. Japan (Honshu, Kyushu). Viburnum dilatatum (Caprifoliaceae). 

Pseudasphondylia rokuharensis Monzen 1955b: 41. Japan: Honshu, Iwate, Fukushima, Wakayama, Oita Prefs; ST; 

♂,♀; KUEC. Refs.: Tokuda & Yukawa 2002 (anat.), Tokuda et al. 2007 (biol.). 

Pseudasphondylia rukuharaensis Yukawa 1971: 124, missp. of rokuharensis. 

*zanthoxyli. China (Yunnan). Zanthoxylum bungeanum (Rutaceae). 

Pseudasphondylia zanthoxyli Mo, Bu & Li. 2007: 974. China: Yunnan, Yongshan Co; HT; ♂; Saut. 

PSEUDOKOCHIOMYIA 

Pseudokochiomyia Fedotova 1984a: 6. Type species, Polystepha vicina Marikovskij (orig. des.). 

Kazakhstan; 5 spp., all on Chenopodiaceae; Cecidomyiidi, Asphondyliini, Schizomyiina. The distinctions between 

this genus and Kochiomyia seem trivial and a restudy may show that the two genera grade into one another. 

camphorosmae. Armenia, Kazakhstan. Camphorosma monspeliaca (Chenopodiaceae). 

Kochiomyia camphorosmae Fedotova 1984d: 299. Kazakhstan: north shore L. Balkash, Baïgabyl; HT; ♂; ZKAS. 

mesasiatica. Armenia, Kazakhstan. Kochia prostrata; K. laniflora (Chenopodiaceae). 

Kochiomyia mesasiatica Fedotova 1982b: 44. Kazakhstan: Kerbulak, right bank Ili R.; HT; ♂; ? 

panderiae. Kazakhstan. Panderia turkestanica (Chenopodiaceae). 

Pseudokochiomyia panderiae Fedotova 1992f: 46. Kazakhstan: Moïynkum Desert, 22 km E Furmanovka; HT; ♂; 

ZMAS. 

taranovii. Kazakhstan. Kochia prostrata (Chenopodiaceae). 

Pseudokochiomyia taranovii Fedotova 1992f: 44. Kazakhstan: Karakumy Desert, 38 km NE Sarykamys; HT; ♂; 

ZMAS. 

vicina. Armenia, Kazakhstan. Kochia prostrata; K. iranica (Chenopodiaceae). 

Kochiomyia vicina Marikovskij 1955b: 105, nomen nudum. 

Polystepha vicina Marikovskij 1961: 50. Kazakhstan: Dzhambul Region, loess desert nr Karatau Mtn Range; ST; 

♂,♀; ZMAS. 

PSEUDOMIKIOLA 

Pseudomikiola Brèthes 1917b: 411. Type species, lippiae Brèthes (orig. des.). 


lippiae. Argentina (Entre Rios). Lippia sp. (Verbenaceae). 

Pseudomikiola lippiae Brèthes 1917b: 413. Argentina: Entre Rios, Concepción del Uruguay; ST; ♂,♀; MACN. 

PUMILOMYIA

Pumilomyia De Stefani 1919: 72. Type species, protrahenda De Stefani (mon.). 

Pulmilomyia De Stefani 1929: 262, missp. of Pumilomyia. 

Southeastern Palearctic; 1 sp.; Cecidomyiidi, Asphondyliini, Asphondyliina. 

protrahenda. Italy. Artemisia arborescens (Asteraceae). 

Pumilomyia protrahenda De Stefani 1919: 72. Italy: Sicily, Palermo; ST; ♂,♀,P,L,egg; lost. 

Pulmilomyia artemisiae De Stefani 1929: 262. Italy: Sicily, Palermo; ST; A,P,L,egg; lost. 

PUNARNAVOMYIA 

Punarnavomyia Mani 1953a: 122. Type species, Cecidomyia boerhaaviaefoliae Mani (orig. des.). 


boerhaaviaefoliae. India (Rajasthan). Boerhavia sp. (Nyctaginaceae). 

Punarnavomyia boerhaaviaefoliae Mani 1953a: 124. India: Rajahstan, Bharatpur, Bund Bareta; HT; ♀; INPC. 

PUTONIELLA 

Putoniella Kieffer 1896b: 4. Type species, Diplosis marsupialis Loew (orig. des.) = pruni Kaltenbach. 

Holarctic; 2 spp., on Prunus; Cecidomyiidi, unplaced to tribe. 

gracilis. USA (Georgia). Prunus angustata; P. persica (Rosaceae). 

Putoniella gracilis Gagné 1992a: 89. USA: Georgia, Byron; HT; ♂; USNM. 

pruni. Widespread Europe. Prunus sp.; P. domestica, P. spinosa (Rosaceae). 

Cecidomyia pruni Kaltenbach 1872: 175. Germany; ST; LT; NHMW?. 

Diplosis marsupialis Löw 1889b: 536. Austria; ST; ♂,♀,L; NHMW. 

*QUADRIDIPLOSIS 

Quadridiplosis Fedotova 2003e: 65. Type species, Karschomyia sidorenkoi Fedotova (orig. des.). 

Eastern Palearctic; 4 spp.; Cecidomyiidi, unplaced to tribe. This genis is close to Coquillettomyia if the two must be 

separated. Fedotova 2003e erected the tribe Quadriplosini for this genus but it is not used here (see Appendix). 

*astrica. Russia (Far East). 

Quadridiplosis astrica Fedotova & Sidorenko 2009b: 8. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*flexa. Russia (Far East). 

Quadridiplosis flexa Fedotova 2003e: 66. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*lucis. Russia (Far East). 

Quadridiplosis lucis Fedotova 2003e: 66. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*sidorenkoi. Russia (Far East). Cirsium schantarense (Asteraceae). 

Karschomyia sidorenkoi Fedotova 2003c: 16. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. Fedotova 2003e: 65 (n. comb.). 

RABDOPHAGA 

340

Rabdophaga Westwood 1847: 588. Type species, Cecidomyia viminalis Westwood (mon). 

Rhabdophaga Kieffer 1901b: 444, unjust. emend. of Rabdophaga. 

Bertieria Kieffer 1896d: 189. Type species, Bertieria gemmicola Kieffer (orig. des.). 

Dichelomyia Rübsaamen 1892d: 346. Type species, Tipula salicis Schrank (subs. des. Gagné 2004b: 229). 

Rübsaamen originally listed a number of species in this genus, but his generic designation appears to describe 

salicis most closely and the types of T. salicis possibly still exist. 

Helicomyia Rübsaamen 1916b: 489. Type species, Lasioptera saliciperda Dufour (orig. des.). 

Phabdophaga Shinji 1944: 172, missp. of Rabdophaga. 

Epourenia Rapp 1946: 128. Type species, viva Rapp (orig. des.). 

Bremiolina Mamaev & Mirumian 1990a: 92. Type species, gemmicola Mamaev & Mirumian = gemmicolata Gagné. 

Holarctic; 82 spp., all on Salix except two on Populus and one on Cephalanthus; Lasiopteridi, Dasineurini. Although 

sometimes considered close to Dasineura or synonymous with it (Skuhravá 1986), Rabdophaga is separated from 

Dasineura by at least one important character of the female postabdomen, which is that the eighth tergite is no longer 

than the seventh and never more than superficially divided, even when the ovipositor is elongated. This is in contrast 

with the situation in Dasineura in which the eighth tergite is typically much longer than the seventh and definitely 

divided into two discrete sclerites. Some species listed below may properly belong to Dasineura. Others have been 

alternately in or out of synonymy and it is difficult at present to judge which authority to follow without better 

knowledge of this large genus. For example, R. degeerii was placed in synonymy under R. salicis in Skuhravá 1986 but 

was treated as a separate species in Stelter 1988b. In another case, R. cineararum was treated as a synonym of R. 

rosaria in Stelter 1993 but as a separate species in Robbins 1996. I have generally followed Stelter here because of his 

many years of experience with willow galls. Nevertheless, the identity of many species appears to be based more on 

tradition than on type specimens. An originally poorly described species, its types lost and no neotype yet designated, 

cannot be identified with certainty. There is also host specificity to consider. While R. rigidae is evidently fairly wideranging 

in host preference (Smith et al. 1983) as well as distribution, other species, most recently R. repenticornua 

Bland (2001), are assumed unique strictly on the basis of host difference. Refs: Harris 2006, Nijveldt 1971, 1979, 1985 

(studies, Palearctic species), Nijveldt and Yukawa 1982 (review Japanese spp.), Smith et al. 1983 (host range), Stelter 

1977, 1980, 1982b-c, 1989 b-d (studies, Palearctic spp.). 

absobrina. USA (New York). 

Rhabdophaga absobrina Felt 1907a: 16. USA: New York, Albany; HT; ♂; NYSM. 

albipennis. Widespread northern Europe. Salix alba (Salicaceae). 

Cecidomyia albipennis Loew 1850a: 35. Poland; ST; ♂; ZMHB?. 

Cecidomyia albipennis Winnertz 1853: 216, jun. prim. hom. albipennis Loew. Germany; ST; ♂,♀; MNHN?. This 

homonym is placed here for now to avoid coining a possibly unnecessary new name. 

americana. USA (Illinois, Connecticut). Salix sp. (Salicaceae). 

Mayetiola americana Felt 1908b: 370. USA: Illinois, Maywood; ST; ♂,♀; NYSM. 

caudicis. USA (Illinois). Salix sp. (Salicaceae). 

Rhabdophaga caulicola Felt 1909b: 290, jun. sec. hom. caulicola Felt 1908. USA: Illinois, Evanston; ST; ♂,♀; 

NYSM. 

Rabdophaga caudicis Gagné 1989b: 274, new name for Rhabdophaga caulicola Felt 1909. 

caulicola. USA (Illinois). Salix sp. (Salicaceae). 

Mayetiola caulicola Felt 1908b: 370. USA: Illinois, Evanston; ST; ♂,♀; NYSM. 

cephalanthi. USA (New York). Cephalanthus occidentalis (Rubiaceae). 

Rhabdophaga cephalanthi Felt 1908b: 355. USA: New York, Nyack; ST; ♂,♀; USNM. 

clavifex. Widespread Europe, Japan (Honshu), Republic of Korea. Salix aurita, S. caprea, S. cinerea; Salix spp. 

(Salicaceae). 

Cecidomyia clavifex Kieffer 1891c: 238. France: Bitche; ST; G; lost?. 

Rhabdophaga repenticola Stelter 1964: 78. Germany: Kr. Rostock, vic. Gross Lüsewitz; HT; ♀; DEIC. 


Rhabdophaga consobrina Felt 1907a: 17. USA: New York, Albany; HT; ♂; NYSM. 

341

342 

cornuta. USA (Illinois). Salix spp. (Salicaceae). 

Cecidomyia cornuta Walsh 1864b: 625. USA: Illinois, nr Rock Island; ST; ♂,P; USNM. 

degeerii. Widespread Europe. Salix purpurea; S. daphnoides (Salicaceae). 

Cecidomyia degeerii (also as degerii) Bremi 1847a: 17, 45. Switzerland: Canton Zurich, nr Dällikon, & Sweden (De 

Geer illustration); ST; LT; ETHZ?. 

Rhabdophaga ramicola Rübsaamen 1916b: 534. Germany: vic. Remagen; ST; ♂,♀,P,L; ZMHB. 

Rabdophaga degeri Simova-Toši et al. 2000: 77, missp. degeerii Bremi. 

deletrix. Netherlands, Germany, Latvia, Poland. Salix alba (Salicaceae). 

Helicomyia deletrix Rübsaamen 1921: 36. Germany; ST; ♂,♀,L; ZMHB. 

dubiosa. Widespread Europe. Salix aurita, S. cinerea (Salicaceae). 

Cecidomyia dubia Kieffer 1891d: 208, jun. prim. hom. dubia Heyden 1862. France: Moselle; ST; LT; lost?. 

Rhabdophaga dubiosa Kieffer 1913i: 70, new name for Cecidomyia dubia Kieffer. 

essigi. USA (California). Salix sp. (Salicaceae). 

Rhabdophaga essigi Felt 1926g: 79. USA: California, Alviso; ST; ♂,♀; NYSM. 

exsiccans. UK, Netherlands, Germany, Latvia. Salix repens (Salicaceae). 

Rhabdophaga exsiccans Rübsaamen 1916b: 533. Germany: Prignitz, Triglitz; NT (Stelter 1982a: 22; Germany: Kr. 

Rostock, Gross Lüsewitz); ♂; DEIC. 

frater. USA (Colorado). Salix sp. (Salicaceae). 

Cecidomyia frater Cockerell 1890b: 280. USA: Colorado, West Cliff; HT; ♂; USNM. 

Cecidomyia frata Felt 1912k: 757, missp. of frater. 

gemmae. USA (New York). Salix sp. (Salicaceae). 

Rhabdophaga gemmae Felt 1908b: 354. USA: New York, Albany; HT; ♂; NYSM. 

gemmicola. Widespread Europe. Salix aurita; S. cinerea (Salicaceae). 

Bertieria gemmicola Kieffer 1896d: 189. France; NT (Stelter 1977: 251; Germany: Kr. Rostock, Gross Lüsewitz); ♂; 

DEIC. 

Rhabdophaga gemmarum Rübsaamen 1916b: 540. Germany: Prignitz, Triglitz; ST; ♂,L; ZMHB. 

gemmicolata. Armenia. Salix triandra (Salicaceae). 

Bremiolina gemmicola Mamaev & Mirumian 1990a: 92, jun. sec. hom. gemmicola Kieffer. Armenia: Echmiadzin 

region, vic. Aratashen; HT; ♂; HYNM. 

Rabdophaga gemmicolata Gagné 2004b: 231, new name for Bremiolina gemmicola Mamaev & Mirumian. 

giraudiana. UK, Denmark, France, Germany, Austria, Romania, Ukraine. Populus tremula, P. alba (Salicaceae). 

Cecidomyia giraudiana Kieffer 1898e: 159. Austria?; T; G; NHMW?. 

globosa. USA (New York, Connecticut). Salix sp. (Salicaceae). 

Rhabdophaga globosa Felt 1908b: 354. USA: New York, Albany; HT; ♂; NYSM. 

heterobia. Widespread Europe, Japan. Salix triandra, Salix spp. (Salicaceae). 

Cecidomyia heterobia Loew 1850a: 35. Poland; ST; ♂,♀; ZMHB?. 

Rhabdophaga amenticola Kieffer 1909b: 27. Austria: nr St. Gertrud in Suldenthal; T; G; NHMW?. 

insignis. Germany, Poland, Italy. Salix purpurea (Salicaceae). 

Rhabdophaga insignis Kieffer 1906b: 70. Italy; ST; F,P; lost?. 

iteobia. Widespread Europe. Salix caprea (Salicaceae). 

Cecidomyia iteobia Kieffer 1890e: 201. France: vic. Bitche & Niederhof; ST; ♂,♀,L; lost?. 

iteophila. Poland. Salix alba (Salicaceae). 

Cecidomyia iteophila Loew 1850a: 35. Poland; ST; ♂,♀; ZMHB?. 

jaapi. Widespread Europe. Salix repens (Salicaceae). 

Rhabdophaga jaapi Rübsaamen 1916b: 526. Germany: Hamburg; ST; ♂,♀,L; ZMHB. 

Dasineura repentis Skuhravá 1986: 160, new name for Rhabdophaga jaapi Rübsaamen while jaapi combined with 

Dasineura. 

justini. UK, Netherlands. Salix purpurea (Salicaceae). 

Rhabdophaga justini Barnes 1935a: 98. UK: England, Lancashire; ST; ♂,♀,P,L; BMNH. 

karschi. UK, France, Germany. Salix aurita, S. cinerea (Salicaceae). 

Cecidomyia karschi Kieffer 1891c: 239. France: Bitche; NT (Stelter 1980: 141; Germany: Kr. Rostock, betw. Gross 

Lüsewitz & Sanitz); ♂; DEIC.

343 

Rhabdophaga oculiperda Rübsaamen 1921: 38. Germany; ST; F,L; ZMHB. 

latebrosa. USA (New York). Salix sp. (Salicaceae). 

Rhabdophaga latebrosa Felt 1909b: 290. USA: New York, Albany; HT; ♂; NYSM. 

latipennis. USA (Illinois). Salix sp. (Salicaceae). 

Rhabdophaga latipennis Felt 1908b: 353. USA: Illinois, Lake Forest; ST; ♂,♀; NYSM. 

lattkei. Germany. Salix pentandra (Salicaceae). 

Rabdophaga lattkei Stelter 1994b: 193. Germany: Kr. Grimmen, Kirch Baggendorf; HT; ♂; SMTD. 

lindhardti. Germany. Salix purpurea (Salicaceae). 

Rabdophaga lindhardti Stelter 1989b: 192. Germany: Kr. Königswusterhausen, Töpchin; HT; ♂; SMTD. 

nervorum. UK, France. Salix aurita (Salicaceae). 

Dichelomyia nervorum Kieffer 1895d: clxxiv. France; ST; A,P; lost?. 

nielsenii. UK, Denmark, Germany, Poland, Czechoslovakia. Salix sp. (Salicaceae). 

Rhabdophaga nielsenii Kieffer 1906a: 1. Denmark: Copenhagen; ST; ♂,♀,P,L; lost?. 

Dasineura nielseni Skuhravá 1986: 157, missp. of nielsenii. 

normaniana. Canada (Manitoba). Salix sp. (Salicaceae). 

Rhabdophaga normaniana Felt 1908b: 354. Canada: Manitoba, Aweme; ST; ♂,♀; NYSM. 

occidua. USA (California). Salix sp. (Salicaceae). 

Phytophaga occidentalis Felt 1926g: 79, jun. prim. hom. occidentalis Felt 1908. USA: California, Berkeley; ST; ♂,♀; 

NYSM. 

Rabdophaga occidua Gagné 1989b: 274, new name for Phytophaga occidentalis Felt 1926g. 

palliumparens. Germany. Salix aurita, S. cinerea (Salicaceae). 

Rhabdophaga palliumparens Stelter 1977: 251. Germany: Kr. Rostock, Gross Lüsewitz; HT; ♂; SMTD. 

perocculta. USA (Colorado). Salix sp. (Salicaceae). 

Cecidomyia perocculta Cockerell 1904: 156. USA: Colorado, Colorado Springs; ST; ♂,P; USNM. 

persimilis. USA (New York). Salix sp. (Salicaceae). 

Rhabdophaga persimilis Felt 1908b: 351. USA: New York, Albany; HT; ♀; NYSM. 

pierreana. Widespread Europe. Salix aurita (Salicaceae). 

Perrisia pierreana Kieffer 1909b: 26. France: Moulins; T; G; lost?. 

pierrei. Widespread Europe. Salix sp. (Salicaceae). 

Bertieria pierrei Kieffer 1896e: 218, referring to earlier misidentification of saliciperda Dufour (Kieffer 1892d: 249). 

France: Bitche; ST; ♂,♀,P,L,egg; lost?. 

podagrae, USA (New York). Salix sp. (Salicaceae). 

Rhabdophaga podagrae Felt 1908b: 355. USA: New York, East Schodack; ST; ♂,♀; NYSM. 

populi. USA (New York). Populus sp. (Salicaceae). 

Rhabdophaga populi Felt 1907a: 16. USA: New York, Albany; ST; ♂,♀; NYSM. 

pulvini. Widespread Europe. Salix aurita, S. cinerea; Salix spp. (Salicaceae). 

Cecidomyia pulvini Kieffer 1891d: 209. France: Moselle; NT (Stelter 1977: 247, Germany: Kr. Rostock, Gross 

Lüsewitz); ♂; DEIC. 

Bertieria superna Kieffer 1896d: 189, nomen nudum. 

Bertieria superna Kieffer 1897a: 24. France: Bitche; T; LT; lost?. 

purpureaperda. UK, Denmark. Salix purpurea (Salicaceae). 

Rhabdophaga purpureaperda Barnes 1935a: 95. UK: England, Lancashire; ST; ♂,♀,P,L; BMNH. 

racemi. Canada (Manitoba). Salix sp. (Salicaceae). 

Rhabdophaga racemi Felt 1908b: 352. Canada: Manitoba, Aweme; ST; ♂,♀; NYSM. 

ramuscula. USA (New York). Salix sp. (Salicaceae). 

Rhabdophaga ramuscula Felt 1908b: 351. USA: New York, New York, Staten I.; ST; ♂,♀; NYSM. 

repenticornua. UK. Salix repens (Salicaceae). 

Rabdophaga repenticornua Bland 2001: 195. UK: Scotland, Perthshire, Rannoch, nr Lochan an Diàm; HT; ♀; 

RSME. 

repentiperda. Germany. Salix repens ssp. argentea (Salicaceae). 

Rhabdophaga repentiperda Stelter 1982b: 23. Germany: Kr. Ribnitz-Damgarten, Halbinsel Rerik, nr Neuhaus (S 

Wustrow); HT; ♂; SMTD.

344 

rigidae. Widespread Nearctic. Salix humilis, Salix spp. (Salicaceae). 

Cecidomyia salicis Fitch 1845a: 263, jun. prim. hom. salicis Schrank 1803. USA: New York, Salem; ST; ♂,♀; 

MCZC. 

Cecidomyia rigidae Osten Sacken 1862: 189, new name for Cecidomyia salicis Fitch. . Ref.: Sato & Yukawa 2006 

(redescr., compared with R. salicivora). 

Cecidomyia salicis siliqua Walsh 1864b: 591, invalid, a polynomial. 

Cecidomyia siliqua Osten Sacken 1878: 4. USA: Illinois, nr Rock Island; ST; F,P,L; USNM. 

rosacea. Canada (Manitoba). Salix sp. (Salicaceae). 

Rhabdophaga rosacea Felt 1908b: 354. Canada: Manitoba, Aweme; HT; ♂; NYSM. 

rosaeformis. Russia (Far East); Japan (Honshu). Salix nipponica, Salix triandra (type host unspec.) (Salicaceae). 

Rhabdophaga rosaeformis Kovalev 1967: 109. Russia: Primorskiï Terr., Khasan; HT; ♂; KOVC. 

rosaria. Widespread Europe. Salix alba; Salix spp. (Salicaceae). 

Cecidomyia rosaria Loew 1850a: 35. Poland; ST; ♂,♀; ZMHB?. Ref.: Harris 2006: 34 (restored from synonymy in 

Gagné 2004 under R. strobilina (Bremi)). 

rosariella. Widespread Europe. Salix aurita, S. cinerea (Salicaceae). 

Bertieria rosariella Kieffer 1896d: 189, nomen nudum. 

Bertieria rosariella Kieffer 1897a: 23. France: Bitche; T; G; lost?. Ref.: Harris 2009: 27 (restored from synonymy). 

roskami. UK, Germany. Salix viminalis (Salicaceae). 

Rabdophaga roskami Stelter 1989c: 91. Germany: Wendenkönig nr Burg/Spreewald; HT; ♂; SMTD. 

salicifoliae. Japan (Honshu). Salix sp. (Salicaceae). 

Rhabdophaga salicifoliae Shinji 1942b: 227. Japan: Honshu, Tokyo; T; A; lost. 

saliciperda. Widespread Europe, Turkey, Japan. Salix sp.; Salix spp. (Salicaceae). 

Lasioptera saliciperda Dufour 1841: 262. France; ST; ♂,♀,P,L; ?. 

Cecidomyia terebrans Loew 1851: 373. Germany; ST; P; ZMHB?. 

salicivora. Eastern Palearctic: Russia (Far East), Korea, Japan (Honshu, Kyushu). Salix babylonica, S. jessoensis, and 

Salix spp. 

Rhabdophaga salicivora Shinji 1938b: 116. Japan: Honshu; NT (Sato & Yukawa 2006: 429); ♂; KUEC. Ref.: Sato 

& Yukawa 2006 (redescr., removed from synonymy with R. rigidae). 

salicis. Widespread Europe, Algeria, Turkey, China, Korea, Japan (Hokkaido to Kyushu), prob. Nearctic; Salix spp. 

(Salicaceae). 

Tipula salicis Schrank 1803: 69. Germany: Bavaria; ST; A,L; lost?. 

Cecidomyia salicina Bouché 1834: 27. Germany; ♂,♀; ZMHB?. 

Cecidomyia gallarumsalicis (as gallarum-salicis) Hardy 1850: 185. UK: Scotland, vic. Penmanshiel; T; ♂; lost. 

Cecidomyia argyrosticta Macquart 1854: 21 (publ'n not seen). France; T; ?; MNHN?. 

Dichelomyia noduli Rübsaamen 1895b: 177. Germany; ST; F,L; ZMHB. 

Rhabdophaga saliyonai (as sali-yonai) Shinji 1938c: 180. Japan: Honshu; T; A; lost. 

salicisbatatas. Widespread Nearctic. Salix humilis; Salix spp. (Salicaceae). 

Cecidomyia salicis batatas Walsh 1864b: 601, invalid, a polynomial. 

Cecidomyia salicisbatatas (as salicis-batatas) Osten Sacken 1878: 4. USA: Illinois, nr Rock Island; ST; ♂,♀,P,L; 


Cecidomyia batatas Beutenmüller 1892: 268, error for salicisbatatas. 

salicisbrassicoides. Widespread Nearctic. Salix longifolia (Salicaceae). 

Cecidomyia salicis brassicoides Walsh 1864b: 577, invalid, a polynomial. 

Cecidomyia salicisbrassicoides Packard 1869: 377. USA: Illinois, nr Rock Island; T; G; USNM. 

Cecidomyia brassicoides Beutenmüller 1892: 268, error for salicisbrassicoides. 

saliciscornu. USA (Illinois). Salix humilis (Salicaceae). 

Cecidomyia salicis cornu Walsh 1864b: 590, invalid, a polynomial. 

Cecidomyia saliciscornu (as salicis-cornu) Osten Sacken 1878: 4. USA: Illinois, nr Rock Island; ST; LT; USNM. 

Cecidomyia cornu Brodie 1909b: 252, error for saliciscornu. 

saliciscoryloides. Widespread Nearctic. Salix discolor (Salicaceae). 

Cecidomyia salicis coryloides Walsh 1864b: 588, invalid, a polynomial. 

Cecidomyia saliciscoryloides (as salicis-coryloides) Osten Sacken 1878: 7. USA: Illinois, nr Rock Island; ST; LT;

345 


Rhabdophaga coryloides Felt 1906a: 745, error for saliciscoryloides. 

salicisgnaphaloides. USA (Illinois). Salix candida, S. humilis (Salicaceae). 

Cecidomyia salicis gnaphaloides Walsh 1864b: 583, invalid, a polynomial. 

Cecidomyia salicisgnaphaloides (as salicis-gnaphaloides) Osten Sacken 1878: 4. USA: Illinois, nr Rock Island; ST; 

F,P,L; USNM. 

Rhabdophaga gnaphaloides Felt 1906a: 745, error for salicisgnaphaloides. 

Rhabdophaga gnaphalioides Felt 1940: 57, missp. and error for salicisgnaphaloides. 

salicisnodulus. USA (widespread eastern). Salix sp; Salix spp. (Salicaceae). 

Cecidomyia salicis nodulus Walsh 1864b: 599, invalid, a polynomial. 

Cecidomyia salicisnodulus (as salicis-nodulus) Osten Sacken 1878: 7. USA: Illinois, nr Rock Island; ST; LT; USNM. 

Rhabdophaga nodulus Felt 1906a: 745, error for salicisnodulus. 

salicisrhodoides USA (Illinois). Salix humilis (Salicaceae). 

Cecidomyia salicis rhodoides Walsh 1864b: 586, invalid, a polynomial. 

Cecidomyia salicisrhodoides (as salicis-rhodoides) Osten Sacken 1878: 4. USA: Illinois, Rock Island Co.; ST; 

♂,♀,P,L; USNM. 

Rhabdophaga rhodoides Felt 1906a: 745, error for salicisrhodoides. 

salicisstrobiliscus. USA (Illinois). Salix discolor, S. rostrata (Salicaceae). 

Cecidomyia salicis strobiliscus Walsh 1864b: 582, invalid, a polynomial. 

Cecidomyia salicisstrobiliscus (as salicis-strobiliscus) Osten Sacken 1878: 4. USA: Illinois, Winnebago Co.; T; G; 


Rhabdophaga strobiliscus Felt 1906a: 745, error for salicisstrobiliscus. 

salicistriticoides. USA (Illinois, New York). Salix cordata, S. humilis (Salicaceae). 

Cecidomyia salicis triticoides Walsh 1864b: 598, invalid, a polynomial. 

Cecidomyia salicis hordeoides Walsh 1864b: 599, invalid, a polynomial. 

Cecidomyia salicistriticoides (as salicis-triticoides) Osten Sacken 1878: 7. USA: Illinois, nr Rock Island; T; G; 


Cecidomyia salicishordoides (as salicis-hordoides) Osten Sacken 1878: 7. USA: Illinois, nr Rock Island; T; G; 


Cecidomyia triticoides Felt 1906a: 745, error for salicistriticoides. 

Cecidomyia hordeoides Felt 1906a: 745, error for salicishordoides. 

salignea. India (Uttar Pradesh). Salix elegans (Salicaceae). 

Oligotrophus saligneus Anonymous 1903: 115, pl. 16. India: Uttar Pradesh, Tehri Garhwal; ST; ♂,♀,P; NZSI. 

Coquillett (1903: 1) formally described this species, but the plate, published earlier, was made by someone 

else. 

schicki. Denmark, Germany. Salix repens ssp. rosmarinifolia (Salicaceae). 

Rhabdophaga schicki Stelter 1982b: 26. Germany: Kr. Rostock, Gross Lüsewitz; HT; ♂; SMTD. 

schreiteri. dENMARK, Germany. Salix repens ssp. rosmarinifolia (Salicaceae). 

Rhabdophaga schreiteri Stelter 1982d: 77. Germany: Mecklenburg, Kr. Bützow, Warnow R. valley nr Baumgarten; 

HT; ♂; SMTD. 

schwangarti. Germany. Salix sp. (Salicaceae). 

Rhabdophaga schwangarti Rübsaamen 1916b: 536. Germany: Tharandt, Neustadt a.d. Haardt; ST; ♂,♀,L; ZMHB. 

sodalitatis. USA (New York). Salix sp. (Salicaceae). 

Rhabdophaga sodalitatis Felt 1908b: 351. USA: New York, Albany; HT; ♀; NYSM. 

strobilina. Widespread Europe, Turkey, Syria, Kazakhstan, Russia (Far East), Korea, China, Japan. Salix purpurea; 

Salix spp. (Salicaceae). 

Cecidomyia strobilina Bremi 1847a: 22. Switzerland: Sihl, nr Zurich; NT (Stelter 1982d: 76, Germany: Kr. 

Königswusterhausen, W Töpchin); ♂; SMTD. 

Cecidomyia cinerearum Hardy 1850: 185. UK: Scotland, vic. Penmanshiel; T; ♂; lost. 

Cecidomyia capreae Hardy 1854b: 45. UK: Scotland, vic. Penmanshiel; ST; ♂,♀,L; lost. 

strobiloides. Widespread Nearctic. Salix cordata; Salix spp. (Salicaceae). 

Cecidomyia salicis strobiloides Walsh 1864b: 580, invalid, a polynomial.

Cecidomyia strobiloides Osten Sacken 1862: 203. USA: Illinois; T; LT; lost. 

Cecidomyia salicisstrobiloides (as salicis-strobiloides) Osten Sacken 1878: 4, unjust. emend. of strobiloides. 

terminalis. Widespread Europe, Japan?. Salix fragilis; Salix spp. (Salicaceae). 

Cecidomyia terminalis Loew 1850a: 35. Poland; ST; ♂,♀; ZMHB?. 

Cecidomyia saligna Hardy 1850: 185. UK: Scotland, vic. Penmanshiel; ST; ♂,♀; lost. 

timberlakei. USA (Utah). Salix fendleriana (Salicaceae). 

Phytophaga timberlakei Felt 1916e: 191. USA: Utah, nr Salt Lake City; ST; ♂,♀,P; NYSM. 

triandraperda. UK, Netherlands, Latvia. Salix triandra (Salicaceae). 

Rhabdophaga triandraperda Barnes 1935a: 92. UK: England, Suffolk, Leicester; ST; ♂,♀,P,L; BMNH. 

tumidosae. Canada (Ontario), USA (Illinois). Salix sp. (Salicaceae). 

Mayetiola tumidosae Felt 1908b: 370. USA: Illinois, Evanston; ST; ♀; NYSM. 

vigemmae. Germany. Salix viminalis (Salicaceae). 

Rabdophaga vigemmae Stelter 1989d: 271. Germany: Plau; HT; ♂; SMTD. 

viminalis. UK, Netherlands, Sweden, Germany. Salix viminalis (Salicaceae). 

Cecidomyia viminalis Westwood 1847: 588. UK: England; ST; ♂,♀,P,L; ?. 

Rhabdophaga perforans Kieffer 1906b: 71. France: Moulins & Bitche; ST; ♀,P; lost?. 

viva. USA (Illinois). 

Epourenia viva Rapp 1946: 128. USA: Illinois, Champaign Co., Mahomet; HT; ♀; lost. 

walshii. USA (Illinois). Salix sp. (Salicaceae). 

Mayetiola walshii Felt 1908b: 371. USA: Illinois, L. Michigan beach; ST; ♂,♀; NYSM. 

RADULELLA 

Radulella Del Guercio 1918a: 46. Type species, aureocephala Del Guercio (orig. des. as n.g., n.sp.). 


aureocephala. Eritrea. Olea chrysophylla (Oleaceae). 

Radulella aureocephala Del Guercio 1918a: 46. Eritrea; T; LT; lost. 

RAODIPLOSIS 

Raodiplosis Felt 1920b: 6. Type species, orientalis Felt (orig. des.). 


orientalis. Burma. 

Raodiplosis orientalis Felt 1920b: 7. Burma: Thaton; ST; ♂,♀; NYSM. 

RESSELIELLA 

Resseliella Seitner 1906: 174. Type species, piceae Seitner (mon.). 

Geodiplosis Kieffer 1909b: 24. Type species, ranunculi Kieffer (orig. des. as n. g., n. sp.). 

Thomasia Rübsaamen 1910d: 288, preocc. Poche 1908. Type species, Diplosis oculiperda Rübsaamen (mon.). 

Profeltiella Kieffer 1912h: 2. Type species, Lestodiplosis ranunculi Kieffer (orig. des.). 

Thomasiniana Strand 1917: 82, new name for Thomasia Rübsaamen. 

Gillotiella Kieffer 1924: 87. Type species, carnea Kieffer (mon.). 

Thomasodiplosis Barnes 1927g: 220, error for Thomasiniana. 

Wichmaniella Möhn 1955c: 97. Type species, crassa Möhn (orig. des.). 

Cosmopolitan; 53 spp., with very diverse plant hosts; Cecidomyiidi, unplaced to tribe. Many species live under bark 

or in flower heads, but some form simple galls. 

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aurata. USA (New York). 

Mycodiplosis aurata Felt 1908b: 402. USA: New York, Albany; HT; ♂; NYSM. 

betulicola. Widespread Europe. Betula alba, B. pubescens (Betulaceae). 

Diplosis betulicola Kieffer 1889a: 155. France; ST; ♂,♀,L; lost?. 

californica. Canada (British Columbia), USA (Oregon, California). Symphoricarpos sp.; S. alba (Caprifoliaceae). 

Thomasia californica Felt 1914c: 132. USA: California, Lake Tahoe; HT; ♀; NYSM. 

carnea. France. 

Gillotiella carnea Kieffer 1924: 87. France: Bitche; HT; ♀; lost?. 

cincta. USA (New York). 

Mycodiplosis cincta Felt 1918e: 203. USA: New York, Jamestown; HT; ♀; NYSM. 

cinctella. USA (Illinois). Quercus sp. (Fagaceae). 

Itonida cincta Felt 1911l: 467, nomen nudum. 

Itonida cincta Felt 1911n: 558, jun. sec. hom. Cecidomyia cincta Felt. USA: Illinois; HT; ♀; USNM. 

Cecidomyia cinctella Kieffer 1913i: 214, new name for Itonida cincta Felt. 

clavula. Canada (Ontario), USA (Indiana, New York, Massachusetts, Maryland, Virginia). Cornus florida (Cornaceae). 

Cecidomyia clavula Beutenmüller 1892: 269. USA: New York, New York, Staten I.; HT; LT; AMNH. 

Mycodiplosis alternata Felt 1907a: 30. USA: New York, Albany; HT; ♂; NYSM. 

conicola. Canada (British Columbia), USA (Washington to California); introd.: Denmark. Abies magnifica; A. 

concolor; Picea sitchensi (in Denmark, Skuhravá et al. 2006) (Pinaceae). 

Mycodiplosis conicola Foote 1956: 54. USA: California, Butte Co; HT; ♂; USNM. 

coryli. USA (New York). 

Mycodiplosis coryli Felt 1907a: 32. USA: New York, Albany; HT; ♂; NYSM. 

coryloides. Canada (British Columbia), USA (Washington, Oregon, California, Montana). Abies concolor; A. 

magnifica, Pseudotsuga menziesii (Pinaceae). 

Mycodiplosis coryloides Foote 1956: 55. USA: California, Butte Co.; HT; ♂; USNM. 

crassa. Germany. Abies alba (Pinaceae). 

Wichmanniella crassa Möhn 1955c: 97. Germany: vic. Dachau, Schönbrunn For., Röhrmoos; HT; ♂; SMNS. 

crataegi. UK, Netherlands, Germany, Russia (Europe), Yugoslavia. Crataegus spp. (Rosaceae). 

Thomasiniana crataegi Barnes 1939c: 343. UK: Staffs., Shropshire, Warwickshire, & Herts; ST; ♂,♀; BMNH. 

dizygomyzae. UK, Netherlands, Czechoslovakia. Salix spp. (Salicaceae). 

Profeltiella dizygomyzae Barnes 1933: 514. UK: Rothamsted, Lancashire, Herts, & Suffolk; ST; ♂,♀,L; BMNH. 

emblicae. India (Uttar Pradesh). Emblica officinalis (Euphorbiaceae). 

Resseliella emblicae Chandra 1991a: 52. India: Uttar Pradesh, Lucknow; HT; ♂; CSIA. 

fruticosi. UK. Rubus fruticosus (Rosaceae). 

Thomasiniana fruticosi Pitcher 1955: 35. UK: Surrey, Wisley; HT; ♂; BMNH. 

galegae. Romania. Galega officinalis (Fabaceae). 

Golanudiplosis galegae Neacsu 1971: 95. Romania: Braïla; T; ♂; NEAC. 

hudsoni. USA (New York). 

Mycodiplosis hudsoni Felt 1907a: 33. USA: New York, Albany; HT; ♂; NYSM. 

ingrica. Poland, Russia (Europe). Picea abies (Pinaceae). 

Thomasiniana ingrica Mamaev 1971: 504. Russia: Leningrad Prov.; HT; ♂; HYNM. 

lavandulae. France. Lavandula officinalis (Lamiaceae). 

Thomasiniana lavandulae Barnes 1953: 125. France: Allemagne-en-Provence; ST; ♂,♀; BMNH. 

liriodendri. USA (widespread eastern). Liriodendron tulipifera (Magnoliaceae). 

Cecidomyia liriodendri Osten Sacken 1862: 202. USA: District of Columbia; T; LT; lost. 

maccus. USA (District of Columbia). 

Diplosis maccus Loew 1862: 187. USA: District of Columbia; ST; ♂,♀; MCZC. 

marginemtorquens. Widespread Europe, Kazakhstan, Japan?. Salix viminalis (Salicaeae). 

Cecidomyia marginemtorquens Bremi 1847a: 28. Switzerland: vic. Zurich; NT (Stelter 1989c: 91; Germany: Kr. 

Rostock, Trebel R. valley betw. Langsdorf & Tribsees); ♂; SMTD. 

meridionalis. Russia (Europe). 

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348 

Profeltiella meridionalis Mamaev 1965a: 267. Russia: Krasnodar Terr., Kudepsta; HT; ♂; HYNM. 

oculiperda. UK, Sweden, Denmark, Netherlands, France, Germany, Latvia, Poland, Czechoslovakia, Ukraine, Russia 

(Europe). Rosa sp. (Rosaceae). 

Diplosis oculiperda Rübsaamen 1893: 161. Germany: Sinzig; ST; ♀,L; ZMHB. 

odai. Japan (Honshu to Kyushu). Cryptomeria japonica (Taxodiaceae). 

Thomasiniana odai Inouye 1955: 8. Japan: Kyushu, Miyazaki Pref.; HT; ♂; HFES. 

oleisuga. Morocco, Italy, Malta, Yugoslavia, Greece, Syria, Israel. Olea europaea (Oleaceae). 

Diplosis oleisuga Targioni-Tozzetti 1887: 428. Italy; ST; ♀,L; lost. 

orientalis. Philippines. Siphonodon celastrineus (Celastraceae). 

Profeltiella orientalis Felt 1918c: 292. Philippines: Luzon, Laguna Prov., Balong Bulo Hill, nr Los Baños; HT; ♂; 

NYSM. 

perplexa. USA (New York). 

Mycodiplosis perplexa Felt 1908b: 402. USA: New York, Albany; HT; ♂; NYSM. 

piceae. Widespread Europe. Abies alba; A. nordmanniana, A. pectinatax (Pinaceae). 

Resseliella piceae Seitner 1906: 174. Slovenia: Idria; ST; ♂,♀,P,L; NHMW. 

pinifoliae. Canada (Ontario), USA (Connecticut). Pinus strobus (Pinaceae). 

Itonida pinifoliae Felt 1936a: 7. USA: Connecticut, Stamford; ST; ♂,♀,L; USNM. 

*poecilantha. Russia (Far East). 

Resseliella poecilantha Fedotova & Sidorenko 2004b: 107. Russia: Primorskiï Terr., 30 km SE Ussuriysk, 


proteae. South Africa. Protea repens (Proteaceae). 

Resseliella proteae Gagné 1983d: 589. South Africa: Cape Prov., Caledon Distr., Tygerhoek; HT; ♂; UNSA. 

quadrifasciata. Japan (Honshu), Korea. Morus alba (Moraceae). 

Diplosis quadrifasciata (as 4-fasciata) Niwa 1910: 1 (publ'n not seen). Japan: Honshu; ST; ♂,♀,L; ?. 

Diplosis fasciata Yokoyama 1929: 388 (publ'n not seen), error for quadrifasciata Niwa. 

quercivora. Russia (Europe). 

Profeltiella quercivora Mamaev 1965a: 267. Russia: Krasnodar Terr., Maykop; HT; ♂; HYNM. 

radicis. USA (Alabama). Xanthium sp. (Asteraceae). 

Mycodiplosis radicis Felt 1936b: 232. USA: Alabama, Mobile; HT; ♂; USNM. 

ranunculi. UK, France. Ranunculus acris (Ranunculaceae). 

Geodiplosis ranunculi Kieffer 1909b: 24. France: château de La Grange, nr Thionville; ST; ♂,♀,L; lost?. 

Lestodiplosis ranunculi Kieffer 1909b: 25, jun. sec. hom. Geodiplosis ranunculi Kieffer. France: château de La 

Grange, nr Thionville; ST; ♀,L; lost?. Considered a new synonym in Gagné 2004b: 236 in preference to 

renaming the homonym. 

*resinicola. Japan (Kyushu). Cryptomeria japonica (Taxodiaceae). 

Resseliella resinicola Sanui & Yukawa 1985: 29. Japan: Kyushu, Miyazaki Pref.; HT; ♂; KUIC. 

resinophaga. Russia (Europe). Abies alba (Pinaceae). 

Thomasiniana resinophaga Mamaev 1971: 504. Russia: Krasnodarskiï Terr., Krasnaya Polyana; HT; ♂; HYNM. 

ribis. Finland, Latvia, Poland, Ukraine, Bulgaria, Russia (Europe). Ribes nigrum (Grossulariaceae). 

Thomasiniana ribis Marikovskij 1956: 190. Russia: Leningrad Prov., Siverskaya; ST; ♂,♀; ZMAS. 

*salicicola. Russia (Far East). Salix gracilistyla (Salicaceae). 

Resseliella salicicola Fedotova 2003c: 12. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

salvadorae. India (Uttar Pradesh, Madhya Pradesh). Salvadora persica (Salvadoraceae). 

Thomasiniana salvadorae Rao 1951: 117. India: Uttar Pradesh, Agra, Castle Grant Garden; HT; ♂; ZSCI. 

sibirica. Kazakhstan, Russia (Eastern Siberia). Larix sp.; L. sibirica (Pinaceae). 

Thomasiniana sibirica Mamaev 1971: 504. Russia: Nizhneudinsk; HT; ♂; HYNM. 

silvana. USA (New York). 

Mycodiplosis silvana Felt 1908b: 402. USA: New York, Albany; HT; ♂; NYSM. 

skuhravyorum. Netherlands, Poland, Czechoslovakia, Austria. Larix decidua, L. polonica (Pinaceae). 

Resseliella skuhravyorum Skrzypczy�ska 1975: 147. Poland: Odrow �ek, Skar�ysko For. Inspectorate (OZLP 

Radom); HT; ♂; Skrzypczy�ska Coll.

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soya. Japan (Hokkaido, Honshu), Korea. Glycine max (Fabaceae). 

Contarinia soya Monzen 1936: 51. Japan: Honshu, Iwate Pref., Shimohei; ST; ♂,♀; KUEC. 

syringogenea. France. Heracleum sphondylium (Apiaceae). 

Clinodiplosis syringogenea Hering 1943: 211. France: Caen; ST; ♂,♀; ZMHB?. 

*tenera. Russia (Far East). 

Resseliella tenera Fedotova 2006a: 158. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

tenuis. UK, Poland. 

Cecidomyia tenuis Loew 1850a: 33. Poland; ST; ♀; lost?. 

theobaldi. Widespread Europe. Rubus sp.; R. idaeus (Rosaceae). 

Thomasiniana theobaldi Barnes 1927g: 219. UK: England, Kent; ST; ♂,♀; BMNH. 

trianguliceps. Egypt. Acacia arabica var. nilotica (Fabaceae). 

Thomasia trianguliceps Debski 1918: 19. Egypt: Marg; ST; LT; ?. 

tulipiferae. Nearctic (widespread eastern). Liriodendron tulipifera (Magnoliaceae). 

Cecidomyia tulipiferae Osten Sacken 1862: 202. USA: District of Columbia; T; LT; lost. 

vespicoloris. UK. Arabis albida (Brassicaceae). 

Profeltiella vespicoloris Barnes 1933: 516. UK: England, Rothamsted; HT; ♀; BMNH. UK. 

*yagoi. Japan (Honshu), Republic of Korea. Pyrus pyrifolia (Rosaceae). 

Resseliella yagoi Yukawa & Sato 2009: 659. Japan: Honshu, Fukushima Pref., Minamisouma City, Kashima; HT; ♂; 

KUEC. 

RHIZOMYIA 

Rhizomyia Kieffer 1898f: 56. Type species, perplexa Kieffer (mon.). 

Coccomorpha Rübsaamen 1899b: 534. Type species, circumspinosa Rübsaamen (mon.). 

Holarctic, Oriental; 30 spp.; Brachineuridi. 

absobrina. USA (New York). 

Arnoldia absobrina Felt 1907d: 8. USA: New York, Albany; ST; ♂,♀; NYSM. 

*applanata. Russia (Far East). 

Rhizomyia applanata Fedotova & Sidorenko 2005d: 6. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

*arsenjevi. Russia (Far East). 

Brachineurina arsenjevi Fedotova 2004j: 603. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. N. comb., Fedotova & Sidorenko 2005d: 8. 

binaria. Russia (Far East). 

Rhizomyia binaria Mamaev 1998d: 7. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

cincta. USA (New York). 

Rhizomyia cincta Felt 1915d: 204. USA: New York, Albany; HT; ♂; NYSM. 

circumspinosa. Germany. Carex sp. (Cyperaceae). 

Coccomorpha circumspinosa Rübsaamen 1899b: 534. Germany; ST; ♂,L; ZMHB. 

clavellata. Italy. 

Rhizomyia clavellata Mamaev 1998d: 6. Italy: Siena; HT; ♂; ZMMU. 

detrita. UK, Russia (Europe). 

Rhizomyia detrita Mamaev 1967: 879. Russia: Vologda Prov., Kadnikovskaya; HT; ♂; HYNM (Yagi & Sawada 

2000) - ZMMU (Krivosheina & Ozerov 2006). 

elongata. Italy. 

Rhizomyia elongata Mamaev & Zaitzev 2002: 5. Italy: Siena, Pontignano; HT; ♂; ZMMU. 

fasciata. Netherlands, France, Russia (Europe). 

Rhizomyia fasciata Kieffer 1904c: 374. France: vic. Bitche; ST; ♂,♀; lost. Mamaev 1967: 878 invalidly designated a 

neotype: no characters were given to separate this species from congeners. 

fraxinifolia. USA (New Jersey). Fraxinus sp. (Oleaceae).

350 

Arnoldia fraxinifolia Felt 1907d: 8. USA: New Jersey, Newfoundland; HT; ♂; NYSM. 

*furcata. Russia (Far East). 

Rhizomyia furcata Fedotova & Sidorenko 2005d: 5. Russia: Primorskiï Terr., Vladivostok; HT; ♂; ZMAS. 

hirta. USA (New York). 

Rhizomyia hirta Felt 1911m: 478. USA: New York, Bath; HT; ♀; NYSM. 

hispida. USA (New York). 

Arnoldia hispida Felt 1907d: 9. USA: New York, Albany; HT; ♂; NYSM. 

improbabilis. Russia (Eastern Siberia). 

Rhizomyia improbabilis Mamaev 1998d: 6. Russia: Khabarovskiï Terr., Khekhtsirskiï Res.; HT; ♂; ZMMU. 


Rhizomyia indica Grover 1972: 21. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

*jaapi. Germany. 

Coccomorpha jaapi Rübsaamen 1925: 42 (Fig. 34). Germany; T; LT; ZMHB. 

*operculata. Russia (Far East). 

Rhizomyia operculata Fedotova & Sidorenko 2006b: 7. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


ornata. Russia (Europe). 

Rhizomyia ornata Mamaev 1967: 878. Russia: Moscow Prov., Il'icha For.; HT; ♂; HYNM. 

perplexa. France, Russia (Europe). Carex glauca (Cyperaceae). 

Rhizomyia perplexa Kieffer 1898f: 57. France: Moselle; ST; A,L; lost?. Mamaev 1967: 878 invalidly designated a 

neotype: no characters were given to separate this species from others. 

*propensa. Russia (Far East). 

Rhizomyia propensa Fedotova & Sidorenko 2006b: 7. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya Bay; 


rossica. Russia (Far East). 

Rhizomyia rossica Mamaev & Zaitzev 2002: 4. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

selecta. UK, Russia (Europe). 

Rhizomyia selecta Mamaev 1967: 879. Russia: Moscow Prov., Pavlova Sloboda; HT; ♂; ZMMU. 

silvicola. France, Russia (Europe). 

Rhizomyia silvicola Kieffer 1904c: 375. France: vic. Bitche; ST; ♂,♀; lost?. Mamaev 1967: 878 invalidly designated 

a neotype: no characters were given to separate this species from others. 

Rhizomyia sylvicola Mamaev 1967: 878, missp. of silvicola. 

tiliacea. USA (New York). 

Dasyneura tiliacea Felt 1907a: 18. USA: New York, Albany; HT; ♂; NYSM. 

Dasyneura cerasi Felt 1907c: 114. USA: New York, Albany; HT; ♂; NYSM. 


Rhizomyia turriformis Fedotova & Sidorenko 2005d: 2. Russia: Primorskiï Terr., Vladivostok; HT; ♂; ZMAS. 

*umbra. Russia (Far East). 

Rhizomyia umbra Fedotova & Sidorenko 2006b: 5. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; ♂; 

ZMAS. 

ungulata. USA (New York). 

Arnoldia ungulata Felt 1907d: 9. USA: New York, Albany; HT; ♂; NYSM. 

vitis. USA (New York). Vitis sp. (Vitaceae). 

Arnoldia vitis Felt 1907d: 9. USA: New York, Albany; ST; ♂,♀; NYSM. 

vittata. Russia (Europe). 

Rhizomyia vittata Mamaev 1967: 879. Russia: Moscow Prov., Pavlovskaja Sloboda; HT; ♂; ZMMU. 

RHOASPHONDYLIA 

Rhoasphondylia Möhn 1960c: 224. Type species, Oxasphondylia friburgensis Tavares (orig. des.). 

Neotropical; 3 spp., on Asteraceae; Cecidomyiidi, Asphondyliini, Asphondyliina.

crassipalpis. Argentina (Mendoza, San Juan). Baccharis salicifolia (Asteraceae). 

Asphondylia crassipalpis Kieffer & Jörgensen 1910: 365. Argentina: Mendoza, nr Pedregal & N Chacras de Coria, & 

San Juan, nr Cancete; ST; ♂,♀,P,L; lost. 

friburgensis. El Salvador, Brazil (Rio de Janeiro). Baccharis schultzi, B. dracunculifolia; B. trinervis (Asteraceae). 

Oxasphondylia friburgensis Tavares 1917b: 126. Brazil: Rio de Janeiro, Nova Friburgo; ST; ♀,P; TAVC. 

sanpedri. Colombia. Chaetospira funcki (Asteraceae). 

Rhoasphondylia sanpedri Wünsch 1979: 63. Colombia: Magdalena Prov., San Pedro de la Sierra; HT; ♂; SMNS. 

RHOPALOMYIA 

Rhopalomyia Rübsaamen 1892d: 370. Type species, Oligotrophus tanaceticola Karsch (subs. des. Kieffer 1896: 89). 

Diarthronomyia Felt 1908b: 339. Type species, artemisiae Felt (orig. des.) = pomum Gagné. 

Diathronomyia Coquillett 1910: 532, missp. of Diarthronomyia. 

Calopedila Kieffer 1913g: 49. Type species, Rhopalomyia herbsti Kieffer (orig. des.). 

Misospatha Kieffer 1913h: 48. Type species, Rhopalomyia globifex Kieffer & Jörgensen (orig. des.). 

Panteliola Kieffer 1913g: 49. Type species, Rhopalomyia haasi Kieffer (orig. des.). 

Boucheella Rübsaamen 1914: 93. Type species, Cecidomyia artemisiae Bouché (orig. des.). 

Dichelonyx Rubsaamen 1914: 94. Type species, Cecidomyia foliorum Loew (orig. des.). 

Dictyomyia Tavares 1919a: 25. Type species, navasina Tavares (orig. des.). 

Navasia Tavares 1919a: 34, preocc. Kirby 1914. Type species, santolinae Tavares (orig. des.). 

Navasiella Tavares 1919b: 93, new name for Navasia Tavares. 

Eudictyomyia Tavares 1920c: 55. Type species, Rhopalomyia navasi Tavares (orig. des.). 

Rhopalomya Shinji 1938h: 1063, missp. of Rhopalomyia. 

Misospatna Shinji 1939c: 588, missp. of Misospatha. 

Phopalomyia Shinji 1939c: 588, missp. of Rhopalomyia. 

Ropalomyia Shinji 1939c: 588, missp. of Rhopalomyia. 

Calopedia Shinji 1944: 323, missp. of Calopedila. 

Mesospatha Shinji 1944: 191, missp. of Misospatha. 

Mesospathia Shinji 1944: 29, missp. of Misospatha. 

Mesospathi Shinji 1944: 42, missp. of Misospatha. 

Rhapalomyia Shinji 1944: 162, missp. of Rhopalomyia. 

Artemisiobia Kovalev 1967: 102. Type species, globosa Kovalev (orig. des.). 

Calopedilla Gagné 1994: 88, missp. of Calopedila. 

Dracunculomyia Fedotova 1999c: 834. Type species, kashkarovi Fedotova (orig. des.). 

Arenaromyia Fedotova 1999a: 588. Type species, caspica Fedtova (orig. des.). 

Absinthomyia (also as Absinhomyia), as subg. of Dracunculomyia, Fedotova 1999c: 835. Type species, 

Dracunculomyia bergi Fedotova (orig. des.). 

Pupascleromyia (also as Pupascleromtyia) Fedotova 1999c: 844, nomen nudum. 

Seriphidomyia Fedotova 1999c: 841, nomen nudum. 

Polynomyia Fedotova 1999c: 846, nomen nudum. 

Seriphidomyia Fedotova 2000d: 1421. Type species, butakovi Fedotova (orig. des.). 

Polynomyia, as subg. of Seriphidomyia, Fedotova 2001a: 59. Type species, Seriphidomyia tarbagataica Fedotova, 

orig. des. 

Yukawyx, as subg of Dichelonyx, Fedotova 2001d: 954. Type species, Dichelonyx ustjurtensis Fedotova (orig. des.). 

Pupascleromyia Fedotova 2001e: 1084. Type species, iliensis (orig. des.). 

Cosmopolitan; 278 spp.; Lasiopteridi, Rhopalomyiini. Most species in this genus form galls, most commonly 

complex galls, on Asteraceae, chiefly of the tribe Anthemideae. Rhopalomyia spp. from other host families that remain 

here are: R. audibertiae and R. salviae on Salvia spp. (Lamiaceae), R. goodeniae on Goodenia lunata (Goodeniaceae), 

R. lawrenciae on Lawrencia squamata (Malvaceae), and R. salsolae and R. salsolata on Salsola. 

Dictyomyia and Artemisiobia were synonymized in Gagné (2004b) because they bore no distinctive characters to 

351

352 

differentiate them clearly from other Rhopalomyia spp. Fedotova (1999a-f, 2000a, 2000b-d, 2001a-h, 2004e, 2005a-b), 

in her overview of the gall midge fauna from Artemisia in Kazakhstan and Turkmenistan, unnecessarily resurrected 

previously synonymized generic names and proposed new genera and subgenera that were considered synonyms in 

Gagné (2004b): Dracunculomyia, Arenaromyia, Absinthomyia, Seriphidomyia, and Polynomyia. Fedotova's divisions 

of Rhopalomyia are not convincing because they were based on local representatives of a genus that is widely 

distributed in the Holarctic Region and on seemingly superficial adult characters and adaptive pupal characters. 

Fedotova species described since 2004 in Dracunculomyia and Seriphidomyia are placed here under Rhopalomyia. 

Jones, Barr, & Gagné 1983 showed the great diversity in 28 Rhopalomyia spp. on Artemisia tridentata in southern 

Idaho and that pupal head development appeared to be correlated more with the relative hardness and position of the 

gall on the plant than on presumed consanguinity. Even if these 28 species could be divided cleanly into Fedotova's 

scheme, one would still have to be convinced that the genera were natural ones and that the various character states 

defining them occurred only once. The seemingly adaptive nature of the distinguishing characters argue against this 

(e.g., see Russo 2007). 

abdominalis. Japan (Honshu). Artemisia princeps (Asteraceae). 

Rhopalomyia abdominalis Shinji 1938m: 339. Japan: Honshu; T; ♀; lost. 

abnormis. USA (New York). 

Rhopalomyia abnormis Felt 1908b: 365. USA: New York, Huguenot Pk.; HT; ♂; NYSM. 

abrotani. UK. Artemisia abrotanum (Asteraceae). 

Hormomyia abrotani Trail 1886: 250. UK: Scotland, Old Aberdeen; ST; ♂,♀,L; ?. 

achillearum. Bulgaria. Achillea pseudopectinata (Asteraceae). 

Rhopalomyia achilleae Trotter 1901: 346, jun. sec. hom. achilleae Inchbald. Romania: nr Plovdiv; ST; ♂,♀,P,L; ?. 

Rhopalomyia achillearum Kieffer 1913i: 45, new name for Rhopalomyia achilleae Trotter. 

ajgurcumensis. Kazakhstan. Artemisia arenaria (Asteraceae). 

Boucheella ajgurcumensis (also as ajgyrcumensis & ajgyrkumensis) Fedotova 1999a: 581. Kazakhstan: Aïgyrkumy 

desert, 6 km N Rozhkovo; HT; ♂; ZMAS. 

alatavica. Kazakhstan. Artemisia santolinifolia (Asteraceae). 

Dichelonyx alatavicus Fedotova 2001d: 959. Kazakhstan: Zailiïskiï Alatau, 20 km SE Alma-Ata; HT; ♂; ZMAS. 

albicerata. Kazakhstan. Artemisia albicerata (Asteraceae). 

Dracunculomyia albicerata Fedotova 1999e: 1184. Kazakhstan: Sary-Taukumy desert, 23 km NE Aïdarly; HT; ♂; 

ZMAS.; HT; ♂; ZMAS. 

almataica. Kazakhstan. Artemisia santolinifolia (Asteraceae). 

Dracunculomyia almataica Fedotova 1999f: 1311. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE 

Alma-Ata; HT; ♂; ZMAS. 

altaica. Kazakhstan. Artemisia scoparia (Asteraceae). 

Dracunculomyia altaica Fedotova 1999d: 973. Kazakhstan: Altaï, Listvyaga Mtn Range, 17 km E Katon-Karagaï; 


alticola. USA (Colorado). Artemisia canadensis (Asteraceae). 

Cecidomyia alticola Cockerell 1890b: 281. USA: Colorado, West Cliff; ST; ♂,♀,P,L; USNM. Recorded from Japan 

(Miyoshi 1929) but only on basis of similarity of galls. 

ambrosiae. USA (Texas, Florida). Ambrosia artemisiifolia; A. psilostachya (Asteraceae). 

Rhopalomyia ambrosiae Gagné 1975b: 54. USA: Florida, Hialeah; HT; ♂; USNM. 

ambrosinae. Spain. Ambrosia sp. (Asteraceae). 

Navasia santolinae Tavares 1919a: 34, jun. sec. hom. santolinae Tavares 1902. Spain: Saragossa; ST; ♀,P; TAVC. 

This species has been treated as a junior synonym of santolinae Tavares 1902 (Skuhravá 1986), possibly 

because the host was originally given as Santolina chamaecyparissus. The name of the host was subsequently 

corrected in Tavares 1919b: 97 to a species of Ambrosia, so ambrosinae is treated as a distinct species here. 

Ref.: Skuhravá et al. 2007 (tax'c notes). 

Rhopalomyia ambrosinae Gagné 2004b: 239, new name for santolinae Tavares 1919. 

ampullaria. USA (Washington, Idaho, Missouri, Utah). Artemisia tridentata (Asteraceae). 

Rhopalomyia ampullaria Felt 1916e: USA: Utah, nr Salt Lake City; LT (Gagné 1983b: 33); ♀; NYSM. 

annua. Kazakhstan. Artemisia annua (Asteraceae).

353 

Dracunculomyia annua Fedotova 1999f: 1317. Kazakhstan: Dzhungarskiï Alatau, Kalmak-Emel' Mtn, nr Sayak; HT; 

♀; ZMAS. 

antennariae. USA (Wisconsin, New York, Maine). Antennaria plantaginifolia (Asteraceae). 

Cecidomyia antennariae Wheeler 1889: 209. USA: Wisconsin, Milwaukee; ST; ♂,♀,P,L,egg; ?. 

anthoides. USA (Washington, Idaho, Nevada, Utah). Artemisia tridentata (Asteraceae). 

Rhopalomyia anthoides Gagné 1983b: 45. USA: Idaho, Elmore Co., 3 mi. N Mountain Home; HT; ♂; USNM. 

anthophila. Canada (Ontario), USA (Missouri, New York, Massachusetts, North Carolina). Solidago altissima 

(Asteraceae) 

Cecidomyia anthophila Osten Sacken 1869: 302. USA: New York, nr Brooklyn; ST; ♂,L; MCZC. Ref.: Dorchin et 

al. 2009 (redescr.). 

apicata. USA (New York). 

Rhopalomyia apicata Felt 1908b: 364. USA: New York, Nassau; HT; ♂; NYSM. 

aralensis. Kazakhstan. Artemisia tomentella (Asteraceae). 

Dichelonyx aralensis Fedotova 2001d: 963. Kazakhstan: northern Aral area, Malye Barsuki Desert, 76 km N Aralsk; 


aralocaspica. Kazakhstan. Tanacetum achilleifolium (Asteraceae). 

Rhopalomyia aralocaspica Fedotova 1995b: 24. Kazakhstan: Mangyschlak basin, 75 km W Opornoï Stn; HT; ♂; 

ZMAS. 

arcuata. USA (New York). 

Rhopalomyia arcuata Felt 1907c: 158. USA: New York, Albany; HT; ♂; NYSM. 

ardynka. Kazakhstan. Artemisia marschalliana (Asteraceae). 

Dracunculomyia ardynka Fedotova 1999c: 836. Kazakhstan: Aïgyrkumy desert, nr Ardynka; HT; ♂; ZMAS. 

arenariaflora. Kazakhstan. Artemisia arenaria (Asteraceae). 

Dracunculomyia arenariaflora Fedotova 1999c: 848. Kazakhstan: Mangyschlak basin, 38 km NW Sarykamys; HT; 

♀; ZMAS. 

arenariagemma. Kazakhstan. Artemisia arenaria (Asteraceae). 

Dracunculomyia arenariagemma Fedotova 1999c: 846. Kazakhstan: Mangyschlak basin, 38 km NW Sarykamys; 


artemisiae. Widespread Europe, Japan (Honshu). Artemisia campestris; Artemisia spp. (Asteraceae). 

Cecidomyia artemisiae Bouché 1834: 27. Germany; ST; ♂,♀,P,L; ZMHB?. 

astericaulis. USA (Massachusetts). Aster sp. (Asteraceae). 

Rhopalomyia astericaulis Felt 1907c: 159. USA: Massachusetts, Worcester; HT; ♂; NYSM. 

astericola. Hungary, Kazakhstan. Aster alpinus (Asteraceae). 

Perrisia astericola Kieffer 1909b: 2. Hungary; T; G; lost?. 

asteriflorae. USA (New York). Aster paniculatus (Asteraceae). 

Rhopalomyia asteriflorae Felt 1907d: 17. USA: New York, Albany; HT; ♀; NYSM. 

audibertiae. USA (California), Mexico (Baja California). Salvia apiana (Lamiaceae). 

Rhopalomyia audibertiae Felt 1907d: 18. USA: California; HT; ♂; NYSM. 

azutavica. Kazakhstan. Artemisia sericea (Asteraceae). 

Seriphidomyia azutavica Fedotova 2001b: 695. Kazakhstan: southern Altaï, Azutau Mts, 30 km NE Alekseyevka; 


baccarum. Widespread Europe, Japan. Artemisia vulgaris (Asteraceae). 

Cecidomyia baccarum Wachtl 1883: 477. Austria: vic. Vienna; ST; ♂,♀,P,L; NHMW?. 

baccharis. USA (California). Baccharis pilularis (Asteraceae). 

Rhopalomyia baccharis Felt 1908b: 364. USA: California; ST; ♂,♀; NYSM. Ref.: Russo 2007 (gall, biol., pupa). 

badhyzi. Turkmenistan. Artemisia badhyzi (Asteraceae). 

Pupascleromyia badhyzi Fedotova 2001e: 1088. Turkmenistan: Badkhyz, Badkhyz Res., Akarcheshme Station; HT; 

♂; ZMAS. 

bajansulu. Kazakhstan. Artemisia sublessingiana (Asteraceae). 

Seriphidomyia bajansulu Fedotova 2001a: 66. Kazakhstan: Tarbagataï Mtn Range, 75 km S Ayagyz, nr Tansyk; HT; 

♀; ZMAS. 

balchanica. Turkmenistan. Artemisia kemrudica (Asteraceae).

354 

Dracunculomyia balchanica Fedotova 1999c: 842. Turkmenistan: Bol'shie Balchany Mtn Range, nr Nebit-Dar; HT; 

♂; ZMAS. 

barsukensis. Kazakhstan. Artemisia albicerata (Asteraceae). 

Dichelonyx barsukensis Fedotova 2001d: 962. Kazakhstan: Bol'shie Barsuki Desert, 128 km NW Kulandy, 100 km 

SW Chelkar; HT; ♀; ZMAS. 

baudysi. Czech Republic, Serbia. Artemisia pontica (Asteraceae). 

Rhopalomyia baudysi Vimmer 1928b: 54. Czech Republic: Hustope e; T; ♂; ?. 

bedeguaris. Argentina (Mendoza). Lycium chilense (Solanaceae). 

Panteliola bedeguaris Kieffer & Jörgensen 1910: 403. Argentina: Mendoza, Cordillera de Mendoza; ST; ♂,♀,P; lost. 

bekoviczi. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Seriphidomyia bekoviczi Fedotova 2001a: 58. Kazakhstan: Chu-Iliïskie Mts, nr Khantau; HT; ♂; ZMAS. 

bergi. Kazakhstan. Artemisia sublessingiana (Asteraceae). 

Dracunculomyia bergi Fedotova 1999c: 838. Kazakhstan: Dzhungarskiï Alatau, 30 km SE Uch-Aral; HT; ♂; ZMAS. 

betheliana. USA (Colorado). Artemisia frigida (Asteraceae). 

Rhopalomyia betheliana Cockerell 1909: 150. USA: Colorado, vic. Denver; LT (Gagné 1983c: 69); ♂; USNM. 

bigeloviae. USA (Colorado). Chrysothamnus sp. (Asteraceae). 

Cecidomyia bigeloviae Cockerell 1889: 324. USA: Colorado; T; G; USNM. 

bigeloviaestrobiloides. USA (New Mexico). Chrysothamnus graveolens (Asteraceae). 

Cecidomyia bigeloviaestrobiloides (as bigeloviae-strobiloides) Townsend 1894: 176. USA: New Mexico, nr Gallo 

Spring; ST; G; lost. 

bigelovioides. USA (California). Chrysothamnus sp. (Asteraceae). 

Rhopalomyia bigelovioides Felt 1908b: 366. USA: California, Los Angeles; ST; ♂,♀; USNM. 

botryosa. Kazakhstan. Artemisia scoparia (Asteraceae). 

Rhopalomyia botryosa Fedotova 1984b: 34. Kazakhstan: Chu-Iliïskie Mts, vic. Khantau; HT; ♂; ZKAS. 

*brassiformis. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia brassiformis Fedotova 2005a: 457. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♀; ZMAS 

brevibulla. USA (Idaho, Missouri, Utah). Artemisia tridentata (Asteraceae). 

Rhopalomyia brevibulla Gagné 1983b: 29. USA: Idaho, Elmore Co., 3 mi. N Mountain Home; HT; ♂; USNM. 

buchtulensis. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Pupascleromyia buchtulensis Fedotova 1999c: 844, nomen nudum. 

Pupascleromyia buchtulensis Fedotova 2001e: 1086. Kazakhstan: middle course Syr Darya R., Bukhtulen Station, 5 

km N Khodzhatogaï; HT; ♂; ZMAS. 

bulbula. USA (Massachusetts). Solidago juncea (Asteraceae). 

Rhopalomyia bulbula Felt 1908b: 365. USA: Massachusetts, Worcester; ST; ♂,♀; NYSM. Ref.: Dorchin et al. 2009 

(redescr.). 

butakovi. Kazakhstan. Artemisia lercheana (Asteraceae). 

Seriphidomyia butakovi Fedotova 2000d: 1422. Kazakhstan: Bolshie Barsuki Desert, 7 km NE Chelkar; HT; ♂; 

ZMAS. 

californica. USA (Oregon, California); immigr.: Australia (Queensland). Baccharis pilularis; B. halimifolia 

(Asteraceae). 

Rhopalomyia californica Felt 1908b: 364. USA: California, Alameda Co.; ST; ♂,♀; USNM. Refs.: Rosenheim et al. 

2007 (biol.); Russo 2007 (gall, pupa). 

calvipomum. USA (Idaho, Nevada). Artemisia tridentata (Asteraceae). 

Rhopalomyia calvipomum Gagné 1983b: 23. USA: Idaho, Twin Falls Co., Twin Falls; HT; ♂; USNM. 

campestris. Denmark, Germany, Poland, Ukraine. Artemisia campestris (Asteraceae). 

Misospatha campestris Rübsaamen 1916b: 550. Germany: Rhine Prov.; ST; ♂,♀; ZMHB. 

capitata. USA (New York, Pennsylvania, Maryland, West Virginia). Solidago gigantea; S. canadensis, S. 

leavenworthii (Asteraceae). 

Rhopalomyia capitata Felt 1908b: 363. USA: New York, West Nyack; ST; ♂,♀; NYSM. Ref.: Dorchin et al. 2009 

(redescr.). 

caspica. Kazakhstan. Artemisia arenaria (Asteraceae). 

Arenaromyia caspica Fedotova 1999a: 589. Kazakhstan: Karakumy Desert, Kul'zhan, 22 km SW Opornaya Stn; HT;

355 

♂; ZMAS. 

caterva. Japan (Honshu). Artemisia japonica; A. vulgaris var. parviflora (Asteraceae). 

Rhopalomyia caterva Monzen 1937: 186. Japan: Iwate Pref., Morioka & Koiwai; ST; ♂; KUEC. 

Rhopalomyia tubiartemisiae (also as atubiartemisiae) Shinji 1938e: 318. Japan: Honshu; T; ♂; lost. 

*cephaliformis. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia cephaliformis Fedotova 2005a: 438. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♀; ZMAS. 

chionophylla. Kazakhstan. Artemisia marschalliana (Asteraceae). 

Dracunculomyia chionophylla Fedotova 1999c: 836, nomen nudum. 

Dracunculomyia chionophylla Fedotova 1999d: 975. Kazakhstan: eastern shore L. Zaysan, 17 km W 

Bol'shenarymsk; HT; ♂; ZMAS. 

chrysanthemi. Origin unknown, presumably from Asia with chrysanthemum trade; elsewhere probably an immigrant: 

widespread Nearctic, widespread northern Europe, Chile, Colombia, Argentina, Hawaiian Is, New Zealand. 

Chrysanthemum spp. (Asteraceae). 

Diarthronomyia chrysanthemi Ahlberg 1939: 276. Europe; LT (Gagné 1983c: 68; USA: MI, East Lansing); ♂; 

NYSM. 

chrysanthemum. Japan (Honshu to Kyushu). Chrysanthemum japonicum (Asteraceae). 

Rhopalomyia chrysanthemum Monzen 1937: 183. Japan: Nagano & Fukuoka Prefs; ST; ♂,♀; KUEC. 

chrysopsidis. USA (Minnesota, Maryland). Chrysopsis mariana (Asteraceae). 

Cecidomyia chrysopsidis Loew 1862: 203. USA: District of Columbia; T; G; MCZC. 

chrysothamni. USA (Utah). Chrysothamnus graveolens (Asteraceae). 

Rhopalomyia chrysothamni Felt 1916e: 187. USA: Utah, nr Murray; ST; ♂,♀,P; NYSM. 

cinerarius. Japan (Honshu to Okinawa), Russia (Far East). Artemisia princeps (Asteraceae). 

Rhopalomyia cinerarius Monzen 1937: 187. Japan: Iwate, Yamagata, Akita, Kanagawa, Wakayama, & Okayama 

Prefs; ST; ♂,♀; KUEC. 

clarkei. Canada (Ontario), USA (New York to North Carolina, New Hampshire, Massachusetts). Solidago rugosa; S. 

altissima (Asteraceae). 

Rhopalomyia clarkei Felt 1907d: 18. USA: New Hampshire, Tamworth; HT; ♀; NYSM. Ref.: Dorchin et al. 2009 

(redescr.). 

clinata. USA (California). Artemisia "californica" (Asteraceae). 

Diarthronomyia californica Felt 1912d: 752, jun. sec. hom. californica Felt 1908b. USA: California, Claremont; LT 

(Gagné 1983c: 69); ♂; USNM. 

Rhopalomyia clinata Gagné 1975c: 483, new name for Diarthronomyia californica Felt 1912d. 

cockerelli. USA (Colorado). 

Rhopalomyia cockerelli Felt 1915j: 237. USA: Colorado, Boulder; ST; ♂; USNM. 

coloradella. USA (Colorado). Artemisia sp. (Asteraceae). 

Diplosis coloradella Cockerell 1904: 155. USA: Colorado, Colorado Springs, nr Monument Creek; LT (Gagné 

1983c: 69); ♂; USNM. 

conica. USA (Washington, Idaho, Nevada). Artemisia tridentata; Artemisia spp. (Asteraceae). 

Rhopalomyia conica Gagné 1983b: 50. USA: Idaho, Twin Falls Co., 5 mi. E Twin Falls; HT; ♂; USNM. 

costacoxita. Kazakhstan. Artemisia diffusa (Asteraceae). 

Seriphidomyia costacoxita Fedotova 2000d: 1427. Kazakhstan: Ul'ken Karatau plateau, 10 mi SW Zhanatas, nr 

Turkestan; HT; ♂; ZMAS. 

cramboides. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia cramboides Gagné 1983b: 45. USA: Idaho, Cassia Co., 4 mi. W Almo; HT; ♂; USNM. 

crassulina. USA (Colorado). Aster crassulus (Asteraceae). 

Cecidomyia crassulina Cockerell 1908b: 89. USA: Colorado, Boulder; ST; ♂; USNM. 

cristaegalli. UK, Netherlands, Germany, Czechoslovakia. Rhinanthus minor (Scrophulariaceae). 

Dasyneura cristae galli Karsch 1877: 31, invalid, a polynomial. 

Rhopalomyia cristaegalli (as cristae-galli) Kieffer 1898f: 21. Germany: vic. Finkenkrug nr Berlin; ST; ♂,♀; ZMHB?. 

cruziana. USA (California). Solidago sp. (Asteraceae). 

Rhopalomyia cruziana Felt 1908b: 366. USA: California, Santa Cruz Mts; ST; ♂,♀; USNM. Ref.: Dorchin et al. 

2009 (redescr.). Ref.: Dorchin et al. 2009 (redescr.).

356 

culmata. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia culmata Gagné 1983b: 36. USA: Idaho, Elmore Co., 3 mi. N Mountain Home; HT; ♂; USNM. 

czarschanga. Turkmenistan. Artemisia badhyzi (Asteraceae). 

Seriphidomyia czarchanga Fedotova 1999c: 841, nomen nudum. 

Seriphidomyia czarschanga Fedotova 2000d: 1424. Turkmenistan: Kugitang Mtn Range, nr Charshanga; HT; ♂; 

ZMAS. 

disciformans. Russia (Far East). Artemisia gmelinii (Asteraceae). 

Rhopalomyia disciformans Kovalev 1967: 100. Russia: Primorskiï Terr., Khasan; HT; ♂; KOVC. 

dracunculiflora. Kazakhstan. Artemisia dracunculus (Asteraceae). 

Dracunculomyia dracunculiflora Fedotova 1999e: 1193. Kazakhstan: Saur Mtn Range, 13 km S Zaïsan, nr 

Zhanaturmys; HT; ♂; ZMAS. 

dracunculis. Kazakhstan. Artemisia dracunculis (Asteraceae). 

Arenaromyia dracunculus Fedotova 1999a: 593. Kazakhstan: Terskeï Alatau, floodplain Tekes R., 20 km N 

Narynkol; HT; ♂; ZMAS. 

efremovi. Kazakhstan. Artemisia dracunculus (Asteraceae). 

Dracunculomyia efremovi Fedotova 1999e: 1189. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Kshi-Kaïndy 

canyon, 12 km NW Novonikolaevki; HT; ♂; ZMAS. 

ehstragoni. Kazakhstan. Artemisia dracunculus (Asteraceae). 

Dracunculomyia ehstragoni Fedotova 1999e: 1191. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE 


*eudokiae. Russia (Far East). Artemisia scoparia (Asteraceae). 

Arenaromyia eudokiae Fedotova 2004e: 20. Russia: Primorskiï Terr., 23 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

ericameriae. USA (California). Ericameria palmeri (Asteraceae). 

Rhopalomyia ericameriae Felt 1916e: 190. USA: California, Puente Hills nr Whittier; ST; ♂,♀,P; NYSM. 

erigerontis. USA (California). Erigeron "fragilis" (Asteraceae). 

Rhopalomyia erigerontis Felt 1916e: 189. USA: California, Whittier; ST; ♂,P; NYSM. 

erii. USA (California). Artemisia californica (Asteraceae). 

Coccidomyia erii Felt 1912h: 147. USA: California, Ventura Co., Carpenteria; ST; ♂,♀; NYSM. 

ermeni. Kazakhstan. Artemisia absinthium (Asteraceae). 

Seriphidomyia ermeni Fedotova 2001b: 697. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE Alma-Ata; 


fedczenkoae. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Seriphidomyia fedczenkoae Fedotova 2000d: 1429. Kazakhstan: Ili R. delta, nr Karoi; HT; ♂; ZMAS. 

flavipalpis. Czech Republic. Artemisia vulgaris (Asteraceae). 

Rhopalomyia flavipalpis Vimmer 1928b: 59. Czech Republic: Lipnik nad Be vou; ST; ♂,♀; ?. 

floccosa. USA (California). Artemisia californica (Asteraceae). 

Diarthronomyia floccosa Felt 1916e: 195. USA: California, Marin Co., betw. Mill Valley & Muir Woods; ST; ♀; 

NYSM. 

florella. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia florella Gagné 1983b: 49. USA: Idaho, Elmore Co., 3 mi. N. Mountain Home; HT; ♂; USNM. 

florum. UK, France, Germany, Latvia, Czechoslovakia, Hungary, Japan. Artemisia vulgaris (Asteraceae). 

Cecidomyia florum Kieffer 1890c: 37. France: Bitche; ST; ♂,♀; lost?. 

foliorum. Widespread Europe, Japan (Hokkaido to Okinawa). Artemisia vulgaris; A. montana, A. princeps 

(Asteraceae). 

Cecidomyia foliorum Loew 1850a: 36. Poland; ST; ♂; ZMHB?. Ref.: Ganaha et al. 2007 (redescr., biol.). 

frigida. Kazakhstan. Artemisia frigida (Asteraceae). 

Navasiella frigida Fedotova 1999b: 691. Kazakhstan: Saur Mtn Range, 15 km S Zaïsan, nr Zhanaturmys; HT; ♂; 

ZMAS. 

fusiformae. USA (New York, Connecticut, New Jersey). Solidago graminifolia; E. tenuifolia (Asteraceae). 

Rhopalomyia fusiformae Felt 1907a: 24. USA: New York, Albany; HT; ♂; NYSM. Ref.: Dorchin et al. 2009 

(redescr.).

357 

Rhopalomyia fusiformis Felt 1907c: 120, unjust. emend. of fusiformae. 

gelleri. Kazakhstan. Artemisia scoparia (Asteraceae). 

Dracunculomyia gelleri Fedotova 1999d: 969. Kazakhstan: eastern Mangyshlak basin, 125 km E Senek; HT; ♂; 

ZMAS. 

gemmaria. USA (Massachusetts). Aster ericoides (Asteraceae). 

Cecidomyia gemmaria Stebbins 1910: 53. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

gerasimovi. Kazakhstan. Artemisia rutifolia (Asteraceae). 

Dracunculomyia gerasimovi Fedotova 1999f: 1306. Kazakhstan: Dzhungarskiï Alatau, Altyn Emel' pass, nr Baschi; 


*gina. USA (Pennsylvania). Solidago juncea (Asteraceae). 

Rhopalomyia gina Dorchin 2009: 13. USA: Pennsylvania, Mauses Creek; ♂; USNM. 

giraldii. China (Shanxi), Russia (Far East), Japan (Honshu to Kyushu). Artemisia sp.; Artemisia princeps (Asteraceae). 

Rhopalomyia giraldii Kieffer & Trotter 1900: 233. China: Shanxi, Huo-Tria-Zaez; ST; ♂,♀,P; lost. 

Rhopalomyia neoartemisiae Shinji 1938e: 317. Japan: Honshu; ST; ♂,♀; lost. 

Rhopalomyia gossypii Monzen 1955a: 44. Japan: Honshu, Aomori, Iwate, Shizuoka, Wakayama, Okayama, & 

Fukuoka Prefs; ST; ♂,♀; KUEC. 

globifex. Argentina (Mendoza, San Juan). Baccharis salicifolia (Asteraceae). 

Rhopalomyia globifex Kieffer & Jörgensen 1910: 364. Argentina: Mendoza, nr Chacras de Coria, Pedregal, 

Potrerillos, & San Ignacio, & San Juan, nr Cancete; ST; ♂,♀,P,L; lost. 

globosa. Russia (Far East). Artemisia rubripes (Asteraceae). 

Artemisiobia globosa Kovalev 1967: 102. Russia: Primorskiï Terr., Suputinskiy Res.; HT; ♂; KOVC. 

glutinosa. USA (Utah?). Chrysothamnus graveolens (Asteraceae). 

Rhopalomyia glutinosa Felt 1916e: 188. USA: Utah?; ST; ♂,P; NYSM. 

gnaphalodis. USA (Colorado). Artemisia gnaphalodes (Asteraceae). 

Rhopalomyia gnaphalodis Felt 1911m: 484. USA: Colorado, Boulder; LT (Gagné 1983c: 70); ♀; NYSM. 

goodeniae. Australia (South Australia). Goodenia lunata (Goodeniaceae). 

Rhopalomyia goodeniae Kolesik 1996b: 155. Australia: South Australia, William Creek; HT; ♂; SAMA. 

gossypina. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia gossypina Gagné 1983b: 37. USA: Idaho, Minidoka Co., 2 mi. E Acequia; HT; ♂; USNM. 

grindeliae. USA (California). Grindelia cuneifolia (Asteraceae). 

Rhopalomyia grindeliae Felt 1916e: 186. USA: California, nr Millbrae; ST; ♂,♀; NYSM. 

gutierreziae. USA (New Mexico). Gutierrezia sarothrae (Asteraceae). 

Asphondylia gutierreziae Cockerell 1901: 23. USA: New Mexico, Las Vegas; ST; ♂,P; USNM. 

*guttata. USA (Connecticut). Solidago bicolor (Asteraceae). 

Rhopalomyia guttata Dorchin 2009: 15. USA: Connecticut, Meriden; ♂; USNM. 

gynaptera. Mongolia. Brachanthemum gobicum (Asteraceae). 

Diarthronomyia gynaptera Mamaev 1972a: 894. Mongolian People's Republic: South Gobi Admin. Distr.; HT; ♀; 

ZMAS. 

haasi. India (West Bengal). Artemisia sp. (Asteraceae). 

Rhopalomyia haasi Kieffer 1905a: 151. India: West Bengal, vic. Kurseong; ST; ♂,♀,P,L,egg; lost?. 

heptapotamica. Kazakhstan. Artemisia marshalliana (Asteraceae). 

Dracunculomyia heptapotamica Fedotova 1999d: 967. Kazakhstan: Dzhungarskiï Alatau, Malaïsary pass, 168 km 

NW Alma-Ata; HT; ♂; ZMAS. 

herbsti. Chile. Baccharis rosmarinifolia (Asteraceae). 

Rhopalomyia herbsti Kieffer 1903: 226. Chile: vic. Concepción; ST; ♂,♀,P,L; lost. 

heteropalpis. Kazakhstan. Artemisia cina (Asteraceae). 

Diarthronomyia heteropalpis Marikovskij 1964: 94 (publ'n not seen). Kazakhstan: Alma-Ata; ?; ?; ?. 

hirtibulla. USA (Idaho, Missouri, Utah). Artemisia tridentata (Asteraceae). 

Rhopalomyia hirtibulla Gagné 1983b: 28. USA: Idaho, Elmore Co., 3 mi. N Mountain Home; HT; ♂; USNM. 

hirticaulis. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia hirticaulis Gagné 1983b: 63. USA: Idaho, Lincoln Co., 11.5 mi. W Kimama; HT; P; USNM. 

hirtipes. Widespread eastern Nearctic. Solidago juncea (Asteraceae).

358 

Cecidomyia hirtipes Osten Sacken 1862: 195. USA: New York, Eldridge Wilderness; N (desig. Dorchin et al. 2009: 

16); ♂; USNM. Ref.: Dorchin et al. 2009 (redescr.). 

hirtipomum. USA (Washington, Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia hirtipomum Gagné 1983b: 21. USA: Washington, Adams Co., 1.2 mi. W Washtucna; HT; ♂; USNM. 

hispanica. Spain. Artemisia herba-alba (Asteraceae). 

Rhopalomyia hispanica Tavares 1904a: 295. Spain: Huesca, Serra de Guara; ST; ♂,♀,P; TAVC. 

hypogaea. UK, France, Germany, Czechoslovakia, Austria, Italy, Russia (Europe). Chrysanthemum atratum 

(Asteraceae). 

Cecidomyia hypogaea Löw 1885b: 488. Austria: Raxalpe; ST; M,P,L; NHMW. 

Rhopalomyia hypogaea of Felt 1915c: 267, misident. of chrysanthemi Ahlberg. 

iliensis. Kazakhstan. Artemisia santolina (Asteraceae). 

Pupascleromyia iliensis Fedotova 1999c: 844, nomen nudum. 

Pupascleromyia iliensis Fedotova 2001e: 1084. Kazakhstan: left bank middle course Ili R., 20 km SE Chilik, nr 

Nurly; HT; ♂; ZMAS. 

iliica. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Navasiella iliica Fedotova 1999b: 686. Kazakhstan: middle course Ili R., 20 km NW Bakanas; HT; ♂; ZMAS. 

incognita. Kazakhstan. Tanacetum achilleifolium (Asteraceae). 

Rhopalomyia incognita Fedotova 1995b: 27. Kazakhstan: Mangyschlak, 75 km W Opornoï Stn; HT; ♂; ZMAS. 

inquisitor. USA (New York). Solidago gigantea (Asteraceae). 

Rhopalomyia inquisitor Felt 1908b: 364. USA: New York, West Nyack; ST; ♂,♀; NYSM. Ref.: Dorchin et al. 2009 

(redescr.). 

integrifoliae. Russia (Far East). Artemisia princeps (Asteraceae). 

Navasiella integrifoliae Fedotova & Kovalev 2001: 437. Russia: Primorskiï Terr., Golubinnyi, nr Tumyntszyan R.; 


iwatensis. Japan (Honshu, Kyushu). Artemisia princeps (Asteraceae). 

Rhopalomyia iwatensis Shinji 1938m: 338. Japan: Honshu, Iwate Pref.; T; ♀; lost. 

japonica. Japan (Honshu). Artemisia japonica, A. princeps (Asteraceae). 

Rhopalomyia japonica Monzen 1937: 185. Japan: Honshu, Iwate Pref., Morioka; ST; ♂,♀; KUEC. 

jaxartica. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Dracunculomyia jaxartica Fedotova 1999c: 844. Kazakhstan: mid course Syr-Darya R., Bukhtulen Stn, 5 km W 

Khodzhatogaï; HT; ♂; ZMAS. 

juncea. Kazakhstan. Artemisia juncea (Asteraceae). 

Seriphidomyia juncea Fedotova 2000d: 1430. Kazakhstan: Karatau Mtn Range, 6 km NE Achchisai; HT; ♂; ZMAS. 

junceaflora. Kazakhstan. Artemisia juncea (Asteraceae). 

Dracunculomyia junceaflora Fedotova 1999c: 838. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 12 km SE 


Dracunculomyia junceafora Fedotova 2000c: 1317, missp. of junceaflora. 

kalkani. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Seriphidomyia kalkani Fedotova 2000d: 1432. Kazakhstan: right bank Ili R., Kalkan-Matai Valley, nr Kalinino; HT; 

♂; ZMAS. 

kalmaki. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Dracunculomyia kalmaki Fedotova 2000c: 1313. Kazakhstan: north shore L. Balkhash, Kalmak-Emel' Mtn, nr Sayak; 


karatavica. Kazakhstan. Artemisia karatavica (Asteraceae). 

Navasiella karatavica Fedotova 1999b: 690. Kazakhstan: Karatau Mtn Range, Kuyuk pass, 22 km SW Dzhambul; 


karmyshevae. Kazakhstan. Artemisia lehmanniana (Asteraceae). 

Dracunculomyia karmyshevae Fedotova 1999c: 840. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Kshi- 

Kaïndy pass; HT; ♂; ZMAS. 

kashkarovi. Kazakhstan. Artemisia arenaria (Asteraceae). 

Dracunculomyia kashkarovi Fedotova 1999c: 835. Kazakhstan: Buzachi Peninsula, 12 km SE Shebir; HT; ♂; ZMAS. 

Dracunculomyia kashcarovi Fedotova 2000c: 1314, missp. of kashkarovi.

359 

kemrudica. Turkmenistan. Artemisia kemrudica (Asteraceae). 

Navasiella kemrudica Fedotova 1999b: 682. Turkmenistan: Bolshie Bolkany Mtn Range, nr Nebit Dag; HT; ♂; 

ZMAS. 

kiefferi. Italy, Yugoslavia. Artemisia camphorata (Asteraceae). 

Rhopalomyia kiefferi Trotter 1900: 285. Italy: Modena & Treviso; ST; ♂,♀,P,L,egg; ?. 

koktalica. Kazakhstan. Artemisia pontica (Asteraceae). 

Dracunculomyia koktalica Fedotova 1999c: 846. Kazakhstan: north bank upper Ili R., 10 km S Koktal; HT; ♀; 

ZMAS. 

korovini. Kazakhstan. Artemisia dracunculus (Asteraceae). 

Dracunculomyia korovini Fedotova 1999e: 1198. Kazakhstan: Saur Mtn Range, 20 km S Zaïsan, nr Churchutsu; HT; 

♀; ZMAS. 

*kovalevi. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia kovalevi Fedotova 2005a: 451. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♂; ZMAS. 

kozlovi. Kazakhstan. Artemisia absinthium (Asteraceae). 

Boucheella kozlovi Fedotova 1999a: 584. Kazakhstan: Azutau Mtn Range, 2.5 km N Uspenka; HT; ♂; ZMAS. 

kugitangica. Turkmenistan. Artemisia badhyzi (Asteraceae). 

Dracunculomyia kugitangica Fedotova 1999c: 841. Turkmenistan: Kugitang Mtn Range, nr Charshanga; HT; ♂; 

ZMAS. 

Dracunculomyia ugitangica Fedotova 2000c: 1321, missp. of kugitangica. 

kunini. Kazakhstan. Artemisia albicerata (Asteraceae). 

Dracunculomyia kunini Fedotova 1999f: 1318. Kazakhstan: Bol'shie Barsuki desert, 100 km SW Chelkar; HT; ♂; 

ZMAS. 

*kuznetzovi. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia kuznetzovi Fedotova 2005a: 438. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♂; ZMAS. 

lateriflori. USA (New York, Massachusetts). Aster sp. (Asteraceae). 

Rhopalomyia lateriflori Felt 1908b: 364. USA: New York, Lake George; ST; ♂,♀; NYSM. 

Rhopalomyia thomi Felt 1915k: 50. USA: Massachusetts; ST; G; Clark Univ., Worcester MA, USA. 

Rhopalomyia lateriflorae Foote 1965: 266, unjust. emend. of lateriflori. 

lawrenciae. Australia (South Australia). Lawrencia squamata (Malvaceae). 

Rhopalomyia lawrenciae Kolesik 1998b: 140. Australia: South Australia, Hindmarsh I.; HT; ♂; SAMA. 

*leleji. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia leleji Fedotova 2005a: 443. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♂; ZMAS. 

*leucanthemidis. Russia. Leucanthemum vulgaris (Asteraceae). 

Rhopalomyia leucanthemidis Fedotova 2008: 128. Russia: Central Povolzhye, Zhigulevsk Nature Res., nr Bakhilova 


levczenkoi. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Seriphidomyia levczenkoi Fedotova 2000d: 1431. Kazakhstan: mid course Syr Darya R., Bukhtulen Stn, 5 km N 

Khodzhatogaï; HT; ♂; ZMAS. 

lignea. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia lignea Gagné 1983b: 41. USA: Idaho, Custer Co., 8 mi. NW Mackay; HT; ♂; USNM. 

lignitubus. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia lignitubus Gagné 1983b: 44. USA: Idaho, Cassia Co., 6 mi. W Burley; HT; ♂; USNM. 

lobata. USA (Illinois, New York, Pennsylvania, Maine). Euthamia graminifolia (Asteraceae). 

Rhopalomyia lobata Felt 1908b: 366. USA: New York, West Nyack; HT; ♀; NYSM. Ref.: Dorchin et al. 2009 

(redescr.). 

Rhopalomyia lanceolata Felt 1908b: 367. USA: Illinois, Lake Forest; ST; ♂,♀; NYSM. 

lobulifera. Russia (Eastern Siberia). Artemisia thilesii (Asteraceae). 

Rhopalomyia lobulifera Mamaev 1990b: 145. Russia: Taimyr, Tareya; HT; ♂; HYNM. 

*longicauda. China (Hebei, Hubei), Republic of Korea. Chrysanthemum morifolium; C. indicum (Asteraceae). 

Rhopalomyia longicauda Sato, Ganaha & Yukawa 2009: 64. China: Hebei Prov., Anguo; HT; ♂; BAUC. 

longitubifex. Japan (Hokkaido to Kyushu), Republic of Korea. Artemisia montana, A. princeps (Asteraceae). 

Misospatha longitubifix (also as longitubifex) Shinji 1939a: 382. Japan: Honshu; ST; ♂,♀; lost. Ref.: Ganaha et al.

360 

2007: 157 (tax'y, syst., biol.) 

Rhopalomyia yomogi Shinji 1938e: 317. Japan: Honshu; ST; ♂,♀; lost. 

Misospatha artemisiae Shinji 1939f: 132, jun. sec. hom. artemisiae Bouché. Japan: Honshu, Morioka; T; ?; lost. 

Panteliola ampulla Monzen 1955a: 45. Japan: Honshu, Aomori, Iwate, Fukushima, Wakayama, Okayama Prefs; ST; 

♀; KUEC. 

Misospatha ampuila Yukawa 1971: 97, missp. of ampulla. 

Rhopalomyia shinjii Gagné 1975c: 483, new name for Misospatha artemisiae Shinji. 

Rhopalomyia longitubifx Yukawa & Masuda 1996: 306, missp, of longitubifex. 

luetkemuelleri. Czechoslovakia, Austria, Italy, Hungary, Yugoslavia, Japan. Artemisia spicata (Asteraceae). 

Rhopalomyia luetkemuelleri (as lütkemülleri) Thomas 1893: 306. Austria: Tyrol; ST; ♂,♀,P; ?. 

magnusi. Germany. Artemisia campestris, A. vulgaris (Asteraceae). 

Rhopalomyia magnusi Rübsaamen 1893: 162. Germany; ST; A,L; ZMHB. 

major. USA (New York). 

Rhopalomyia major Felt 1907a: 25. USA: New York, New York, Staten I.; HT; ♂; NYSM. 

malajsaryensis. Kazakhstan. Artemisia marshalliana (Asteraceae). 

Dracunculomyia malajsaryensis (also as malaisaryensis) Fedotova 1999d: 972. Kazakhstan: Dzhungarskiï Alatau, 

Malaïsary pass, 168 km NW Alma-Ata; HT; ♂; ZMAS. 

mammilla. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia mammilla Gagné 1983b: 52. USA: Idaho, Bingham Co., 15 mi. NW Blackfoot; HT; ♂; USNM. 

mangyshlakensis. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Navasiella mangyshlakensis Fedotova 1999b: 683. Kazakhstan: eastern Mangyshlak basin, 127 km NE Senek; HT; 

♂; ZMAS. 

marschalliana. Kazakhstan. Artemisia marschalliana (Asteraceae). 

Arenaromyia marschalliana Fedotova 1999a: 590. Kazakhstan: right bank Ili R., Malaïsary pass, 168 km NW 


medusa. USA (Idaho, Missouri, Nevada). Artemisia tridentata; Artemisia spp. (Asteraceae). 

Rhopalomyia medusa Gagné 1983b: 59. USA: Idaho, Custer Co., 8 mi. NW Mackay; HT; ♂; USNM. 

medusirrasa. USA (Idaho, Nevada). Artemisia tridentata (Asteraceae). 

Rhopalomyia medusirrasa Gagné 1983b: 62. USA: Idaho, Twin Falls Co., 22.5 mi. S Hansen; HT; ♂; USNM. 

micranthae. Kazakhstan. Achillea micrantha (Asteraceae). 

Rhopalomyia micranthae (also as microcantha) Fedotova 1995b: 25. Kazakhstan: Karakumy Desert, 65 km NW 

Novokazalinska, 45 km SE Aral'ska; HT; ♂; ZMAS. 

millefolii. Widespread Europe, Egypt, India. Achillea millefolium; A. ptarmica (Asteraceae). 

Cecidomyia millefolii Loew 1850a: 37. Poland; ST; ♀; ZMHB?. 

Cecidomyia achilleae Inchbald 1860a: 195. UK: England, nr Huddersfield; T; G; ?. 

mojinkumensis. Kazakhstan. Artemisia marshalliana (Asteraceae). 

Dracunculomyia mojinkumensis Fedotova 1999d: 970. Kazakhstan: Moïynkumy desert, 17 km NE Sarykoby; HT; ♂; 

ZMAS. 

mongolica. Mongolia. Ajania achilleoides (Asteraceae). 

Rhopalomyia mongolica Mamaev 1972a: 894. Mongolian People's Republic: South Gobi Admin. Distr.; HT; ♂; 

ZMAS. 

monogyna. Kazakhstan. Artemisia monogyna (Asteraceae). 

Dichelonyx monogynus Fedotova 2001d: 958. Kazakhstan: western Ustyurt, Koshkuduk well, 162 km NE Senek; 


monogynasphaera. Kazakhstan. Artemisia monogyna (Asteraceae). 

Navasiella monogynasphaera Fedotova 1999b: 685. Kazakhstan: Karakum Desert, Kul'zhan, 22 km NE Opornaya 


muravjevi. Kazakhstan. Artemisia santolinifolia (Asteraceae). 

Pupascleromyia muravjevi Fedotova 2001e: 1090. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak Canyon, 20 km SE 


mushketovi. Kazakhstan. Artemisia sericea (Asteraceae). 

Seriphidomyia mushketovi Fedotova 2001b: 693. Kazakhstan: Saur Mtn Range, 20 km S Zaïsan, nr Churchutsu; HT;

361 

♂; ZMAS. 

navasi. Spain, Romania, Algeria to Egypt. Artemisia herba-alba (Asteraceae). 

Rhopalomyia navasi Tavares 1904a: 296. Spain: Huesca, Serra de Guara; ST; ♂,♀,P; TAVC. 

navasina. Spain. Santolina chamaecyparissus (Asteraceae). 

Dictyomyia navasina Tavares 1919a: 26. Spain: Salvador, nr Saragossa; ST; ♂,♀,P; TAVC. 

Dictyomyia navasiana Skuhravá 1986: 170, missp. of navasina. 

novitzkii. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Pupascleromyia novitzkii Fedotova 2001e: 1095. Kazakhstan: Buzachi Peninsula, 75 km NW Shebir; HT; ♀; ZMAS. 

nucula. USA (Washington, Idaho, Montana, Nevada, Utah). Artemisia tridentata; also A. nova (Asteraceae). 

Rhopalomyia nucula Gagné 1983b: 57. USA: Idaho, Twin Falls Co., 5 mi. E Twin Falls; HT; ♂; USNM. 

obovata. USA (Washington, Idaho, Nevada). Artemisia tridentata; Artemisia spp. (Asteraceae). 

Rhopalomyia obovata Gagné 1983b: 54. USA: Idaho, Twin Falls Co., 5 mi. E Twin Falls; HT; ♂; USNM. 

occidentalis. USA (California, Utah). Artemisia heterophylla (Asteraceae). 

Diarthronomyia occidentalis Felt 1916e: 194. USA: California, San Diego Co., Sweet Water Dam; LT (Gagné 

1983c: 70); ♂; NYSM. 

olgae. Kazakhstan. Artemisia dracunculus (Asteraceae). 

Dracunculomyia olgae Fedotova 1999e: 1193. Kazakhstan: Saur Mtn Range, 20 km NW Zaïsan; HT; ♂; ZMAS. 

palearum. UK, France, Denmark, Germany, Poland, Ukraine. Achillea ptarmica (Asteraceae). 

Hormomyia palearum Kieffer 1890b: 28. France: Guinkirchen, Gesselingen, and Bitche; ST; ♂,♀; lost?. 


Rhopalomyia palustris Felt 1908b: 365. USA: New York, Nassau; HT; ♀; NYSM. 

paratubifex. Russia (Far East). Artemisia desertorum (Asteraceae). 

Rhopalomyia paratubifex Kovalev 1967: 99. Russia: Vladivostok; HT; ♂; KOVC. 

pavlovi. Kazakhstan. Artemisia marschalliana (Asteraceae). 

Boucheella pavlovi Fedotova 1999a: 583. Kazakhstan: Altaï, southern spurs Listvyaga Mts, 17 km E Katon-Karagaï; 


pedicellata. USA (Minnesota, New York, Pennsylvania, Maine, Massachusetts, Maryland, Virginia). Euthamia 

graminifolia (Asteraceae). 

Rhopalomyia pedicellata Felt 1908b: 365. USA: New York, Albany; ST; ♂,♀; NYSM. Ref.: Dorchin et al. 2009 

(redescr.). 

Cecidomyia euthamiae Stebbins 1910: 53. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

petrovi. Kazakhstan. Artemisia gurganica (Asteraceae). 

Seriphidomyia petrovi Fedotova 2000d: 1427. Kazakhstan: northeastern Ustyurt, Kalenkely Mts, 70 km SE Opornaya 


pevtzovi. Kazakhstan. Artemisia halophila (Asteraceae). 

Navasiella pevtzovi Fedotova 1999b: 692. Kazakhstan: Dzhungarskiï Alatau, floodplain Tentek R., nr Uch-Aral; HT; 

♀; ZMAS. 

pilosa. Canada (Northwest Territories). Antennaria sp. (Asteraceae). 

Rhopalomyia pilosa Felt 1908b: 366. Canada: Northwest Territories, Medicine Head; ST; ♂,♀; NYSM. 

pini. USA (New York). 

Rhopalomyia pini Felt 1907a: 24. USA: New York, Albany; HT; ♂; NYSM. 

plumula. Kazakhstan. Erigeron acris (Asteraceae). 

Rhopalomyia plumula Fedotova 1995c: 45. Kazakhstan: southern Altaï, Azutau Mts, 30 km NE Alekseyevka; HT; ♂; 

ZMAS. 

polyni. Kazakhstan. Artemisia dracunculus (Asteraceae). 

Dracunculomyia polyni Fedotova 1999e: 1196. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Kshi-Kaïndy 

canyon, 12 km NW Novonikolaevki; HT; ♀; ZMAS. 

polynovi. Kazakhstan. Artemisia sericea (Asteraceae). 

Seriphidomyia polynovi Fedotova 2001b: 696. Kazakhstan: southern Altaï, Azutau Mts, 20 km NE Alekseyevka; HT; 

♂; ZMAS. 

pomum. USA (Washington, Idaho, Montana, California, Nevada, Utah, Colorado, Nebraska). Artemisia tridentata 

(Asteraceae).

362 

Diarthronomyia artemisiae Felt 1908b: 339, jun. sec. hom. artemisiae Bouché. USA: Colorado, Fort Garland; LT 

(Gagné 1983b: 14); ♂; USNM. 

Rhopalomyia pomum Gagné 1975c: 483, new name for Diarthronomyia artemisiae Felt. 

potanini. Kazakhstan. Artemisia gracilescens (Asteraceae). 

Navasiella potanini Fedotova 1999b: 691. Kazakhstan: 45 km W Ayaguz, Ayaguz R. valley; HT; ♀; ZMAS. 

producticeps. Algeria, Libya. Artemisia herba-alba (Asteraceae). 

Rhopalomyia producticeps Kieffer 1912f: 180. Algeria: mts S Saint-Denis-du-Sig; ST; ♀,P,L; lost?. 

przhevalskii. Kazakhstan. Artemisia pontica (Asteraceae). 

Seriphidomyia przhevalskii Fedotova 1999c: 846, nomen nudum. 

Seriphidomyia przhevalskii Fedotova 2001a: 61. Kazakhstan: right bank upper course Ili R., 10 km S Koktal; HT; ♂; 

ZMAS. 

pseudofoliorum. Czech Republic. Artemisia campestris (Asteraceae). 

Rhopalomyia pseudofoliorum Vimmer 1924: 60. Czech Republic: nr Mikulov; T; ♂; ?. 

*ptarmicacola. Kazakhstan. Ptarmica impatiens (Asteraceae). 

Rhopalomyia ptarmicacola Fedotova 2005b: 909. Kazakhstan: Central Altai, Kok siiskii Mt. Rang, Lyamin Belok 

Mt., 60 km NE Leninogorsk; HT; ♂; ZMAS. 

ptarmicae. Widespread Europe. Achillea ptarmica (Asteraceae). 

Cecidomyia ptarmicae Vallot 1849: 98. France; ST; ♀,L; ?. 

Cecidomyia floricola Winnertz 1853: 289. Germany; ST; ♂,♀,L; MNHN?. 

*pusilla. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia pusilla Fedotova 2005a: 438. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♂; ZMAS. 

quinqueloba. Kazakhstan. Artemisia quinqueloba (Asteraceae). 

Arenaromyia quinqueloba Fedotova 1999a: 592. Kazakhstan: eastern shore Aral Sea, 20 km NE Kazalinsk; HT; ♂; 

ZMAS. 

racemicola. USA (North Carolina, Florida). Solidago altissima; S. fistulosa? (Asteraceae). 

Rhopalomyia racemicola Felt 1907a: 24. USA: North Carolina, Asheville; ST; ♂,♀; NYSM. Ref.: Dorchin et al. 


rerichi. Kazakhstan. Artemisia austriaca (Asteraceae). 

Seriphidomyia rerichi Fedotova 2001b: 700. Kazakhstan: Tarbagataï Mtn Range, nr Novo-Andreevka, 18 km N 

Urdzhara; HT; ♂; ZMAS. 

rozhkovi. Kazakhstan. Artemisia porrecta (Asteraceae). 

Seriphidomyia rozhkovi Fedotova 2001a: 52. Kazakhstan: Aïgyrkumy desert, 6 km N Rozhkovo; HT; ♂; ZMAS. 

*rubripes. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia rubripes Fedotova 2005a: 452. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♂; ZMAS. 

*rubripigemmae. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia rubripigemmae Fedotova 2005a: 447. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♂; ZMAS. 

ruebsaameni. Austria, Italy, Kazakhstan. Erigeron uniflorus; E. tianschanicus, E. pseudoseravschanicus (Asteraceae). 

Rhopalomyia ruebsaameni (as rübsaameni) Thomas 1893: 301. Austria: betw. Arosa & the Brüggerhorn; ST; 

♂,♀,P,egg; NHMW?. 

Perrisia alpicola Kieffer 1909b: 7. Austria; T; G; NHMW?. 

Perrisia erigeronis Kieffer 1909b: 7. Austria: Tyrol, nr St. Gertrud; T; G; NHMW?. 

rugosa. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia rugosa Gagné 1983b: 27. USA: Idaho, Twin Falls Co., 23 mi. S Hansen; HT; ♀; USNM. 

rutifolia. Kazakhstan. Artemisia rutifolia (Asteraceae). 

Seriphidomyia rutifolia Fedotova 2001b: 699. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 12 km NW 

Novonikolaevki; HT; ♂; ZMAS. 

saissanica. Kazakhstan. Artemisia austriaca (Asteraceae). 

Dracunculomyia saissanica Fedotova 1999f: 1309. Kazakhstan: Saur Mtn Range, Zaïsan valley, 6 km NW 

Karabulak; HT; ♀; ZMAS. 

salsolae. Spain. Salsola vermiculata (Chenopodiaceae). 

Misospatha salsolae Tavares 1924a: 44. Spain: Saragossa; HT; ♀; TAVC. 

salsolata. Spain. Salsola vermiculata (Chenopodiaceae).

363 

Dictyomyia salsolae Tavares 1924a: 42, new sec. hom. salsolae Tavares 1924. Spain: Saragossa; HT; ♀; TAVC. 

Rhopalomyia salsolata Gagné 2004b: 247, new name for Dictyomyia salsolae Tavares. 

salviae. USA (California). Salvia californica, S. nivea (Lamiaceae). 

Rhopalomyia salviae Felt 1916e: 184. USA: California, Puente Hills nr Whittier; ST; ♂,♀,P; NYSM. 

santolina. Kazakhstan. Artemisia arenaria (Asteraceae). 

Dracunculomyia santolina Fedotova 1999c: 847. Kazakhstan: left bank mid Ili R., 20 km NE Chilik; HT; ♂; ZMAS. 

santolinae. Portugal, Spain. Santolina rosmarinifolia (Asteraceae). 

Rhopalomyia santolinae Tavares 1902b: 28. Portugal; ST; ♂,♀,P; TAVC. 

santolinifolia. Kazakhstan. Artemisia santolinifolia (Asteraceae). 

Dracunculomyia santolinifolia Fedotova 1999f: 1315. Kazakhstan: Zailiïskiï Alatau, Bel'-Bulak canyon, 20 km SE 


saurica. Kazakhstan. Artemisia frigida (Asteraceae). 

Dracunculomyia saurica Fedotova 1999f: 1308. Kazakhstan: Saur Mtn Range, 10 km S Zaïsan, nr Zhanaturmys; HT; 

♂; ZMAS. 

schrenkiana. Kazakhstan. Artemisia schrenkiana (Asteraceae). 

Navasiella schrenkiana Fedotova 1999b: 689. Kazakhstan: Buzachi Peninsula, 14 km NE Shebir, Zhumazhe area; 


scoparia. Kazakhstan, Turkmenistan. Artemisia scoparia (Asteraceae). 

Dracunculomyia scoparia Fedotova 1999d: 965. Kazakhstan: eastern Mangyshlak basin, 87 km NE Senek; HT; ♂; 

ZMAS. 

scopariaflora. Kazakhstan. Artemisia scoparia (Asteraceae). 

Dracunculomyia scopariaflora Fedotova 1999d: 976. Kazakhstan: Altaï, Listvyaga Mts, 17 km E Katon-Karagaï; 


senekensis. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Dichelonyx senekensis Fedotova 2001d: 955. Kazakhstan: Ustyurt plateau, 125 km NE Senek; HT; ♂; ZMAS. 

serotina. Kazakhstan. Artemisia serotina (Asteraceae). 

Seriphidomyia serotina Fedotova 2001a: 56. Kazakhstan: Dzhungarskiï Alatau, upper Yrgaïty R., 40 km E Koktumy; 


setubalensis. Portugal, Spain, Italy, Algeria. Santolina rosmarinifolia var. vulgaris (Asteraceae). 

Rhopalomyia setubalensis Tavares 1902a: 98. Portugal: vic. Setubal; ST; ♂,♀,P,L; TAVC. 

shevczenkoi. Kazakhstan. Artemisia dracunculus (Asteraceae). 

Boucheella shevczenkoi Fedotova 1999a: 585. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., 12 km NW 


shlagintvejti. Kazakhstan. Artemisia sericea (Asteraceae). 

Pupascleromyia shlagintvejti Fedotova 2001e: 1091. Kazakhstan: Azutau Mtn Range, 20 km NE Alekseevka; HT; ♂; 

ZMAS. 

simulans. Czech Republic. Artemisia campestris (Asteraceae). 

Rhopalomyia simulans Vimmer 1924: 61. Czech Republic: Suchá hora nr Brno; T; ♀; ?. 

simultanea. Kazakhstan. Artemisia scoparia (Asteraceae). 

Rhopalomyia simultanea Fedotova 1984b: 35. Kazakhstan: Chu-Iliïskie Mts, vic. Khantau; HT; ♂; ZKAS. 

solidaginis. Widespread eastern Nearctic. Solidago altissima; S. canadensis, S. rugosa (Asteraceae). 

Cecidomyia solidaginis Loew 1862: 194. USA: New York, West Dryden; N (desig. Dorchin et al. 2009: 30); ♂; 

USNM. Ref.: Dorchin et al. 2009 (redescr.). 

Rhopalomyia albipennis Felt 1908b: 364. USA: New York, Albany; ST; ♂,♀; NYSM. 

Rhopalomyia carolina Felt 1908b: 363. USA: North Carolina, Asheville; HT; ♀; NYSM. 

soongarica. Kazakhstan. Artemisia soongarica (Asteraceae). 

Dracunculomyia soongarica Fedotova 1999e: 1186. Kazakhstan: Sary-Taukumy desert, 23 km NE Aïdarly; HT; ♂; 

ZMAS. 

spongiosa. Kazakhstan. Erigeron pseudoseravschanicus (Asteraceae). 

Rhopalomyia spongiosa Fedotova 1995c: 49. Kazakhstan: Dzhungarskiï Alatau, 6 km S Pokatilovka; HT; ♂; ZMAS. 

strobiligemma. USA (New York, Massachusetts). Aster ericoides (Asteraceae). 

Cecidomyia strobiligemma Stebbins 1910: 53. USA: Massachusetts, vic. Springfield; T; G; NHSM.

364 

struma. Japan (Hokkaido to Kyushu), Korea, Russia (Far East). Artemisia montana; A. princeps, Artemisia spp. 

(Asteraceae). 

Rhopalomyia struma Monzen 1937: 188. Japan: Honshu: Aomori, Iwate, Tokyo, Toyama, Wakayama, Okayama 

Prefs; ST; ♂,♀; KUEC. 

subhumilis. USA (Utah). Artemisia ludoviciana (Asteraceae). 

Rhopalomyia subhumilis Gagné 1978b: 426. USA: Utah, Sanpete Co., 12 mi. E Ephraim, Philadelphia Flat; HT; ♂; 


sublessaingiana. Kazakhstan. Artemisia sublessingiana (Asteraceae). 

Dracunculomyia sublessingiana Fedotova 1999c: 838, nomen nudum. 

Seriphidomyia sublessingiana Fedotova 2001a: 54. Kazakhstan: 4 km S Ush-Tobe, nr Dal'niï; HT; ♂; ZMAS. 

sulcata. USA (Arizona, New Mexico, Texas), Mexico (Durango, Nuevo León). Baccharis salicina; Baccharis spp. 

(Asteraceae). 

Rhopalomyia sulcata Gagné 1995c: 773. USA: New Mexico, Eddy Co., Loving, on Pecos R.; HT; ♂; USNM. 

syngenesiae. UK, France, Germany, Poland, Czechoslovakia, Hungary, Romania, Ukraine. Chrysothamnus inodorum, 

Anthemis arvensis (Asteraceae). 

Cecidomyia syngenesiae Loew 1850a: 39. Poland; ST; ♂,♀; ZMHB?. 

talassica. Kazakhstan. Artemisia santolinifolia (Asteraceae). 

Dracunculomyia talassica Fedotova 1999f: 1314. Kazakhstan: Talasskiï Alatau, Aksu-Dzhabagly Res., Ul'ken- 

Kaïndy canyon, 18 km SE Novonikolaevka; HT; ♂; ZMAS. 

tanaceticola. Widespread Europe, Kazakhstan. Tanacetum vulgare (Asteraceae). 

Oligotrophus tanaceticolus Karsch 1879: 28. Germany: Westphalia; ST; G; NHMW?. 

tarbagataica. Kazakhstan. Artemisia santolinifolia (Asteraceae). 

Seriphidomyia tarbagataica Fedotova 2001a: 59. Kazakhstan: Tarbagataï Mtn Range, upper Karabuga R., 80 km S 

Aksuat; HT; ♂; ZMAS. 

tavaresi. Spain. Artemisia herba-alba (Asteraceae). 

Eudictyomyia artemisiae Tavares 1920c: 59, jun. sec. hom. artemisiae Bouché. 

Rhopalomyia tavaresi Gagné 1975c: 483, new name for Eudictyomyia artemisiae Tavares. 

tenteki. Kazakhstan. Artemisia halophila (Asteraceae). 

Seriphidomyia tenteki Fedotova 2000d: 1428. Kazakhstan: Tarbagataï Mtn Range, Tentek R. floodplain, nr Uch-Aral; 


terraealbae. Kazakhstan. Artemisia terraealbae (Asteraceae). 

Dichelonyx terraealbae Fedotova 2001d: 961. Kazakhstan: right bank middle course Ili R., Kerbulak, nr Kapchagaï; 


terskeica. Kazakhstan. Aster alpinus (Asteraceae). 

Rhopalomyia terskeica Fedotova 1995c: 52. Kazakhstan: Terskeï Alatau, Shybyndysai gorge, 49 km S Kegen'; HT; 

♂; ZMAS. 

tetradymia. USA (California). Tetradymia comosa (Asteraceae). 

Phytophaga tetradymia Felt 1925a: 15. USA: California, Claremont; ST; ♂,♀; NYSM. 

thompsoni. USA (New York, Massachusetts). Solidago altissima (Asteraceae). 

Rhopalomyia thompsoni Felt 1907c: 159. USA: Massachusetts, Worcester; ST; ♂,♀; NYSM. Ref.: Dorchin et al. 


tianschanensis. Kazakhstan. Artemisia tianschanica (Asteraceae). 

Navasiella tianschanica Fedotova 1999b: 687, jun. sec. hom. Rhopalomyia tianschanica Fedotova 1995. Kazakhstan: 

W Tien Shan, floodplain Aksu R., nr Belye Vody; HT; ♂; ZMAS. 

Rhopalomyia tianschanensis Gagné 2004b: 249, new name for Navasiella tianschanica Fedotova 1999b. 

tianschanica. Kazakhstan. Erigeron tianschanicus (Asteraceae). 

Rhopalomyia tianschanica Fedotova 1995c: 48. Kazakhstan: Dzhungarskiï Alatau 25 km S Arasan, Biendy-Koksu 

gorge; HT; ♂; ZMAS. 

tjulkubasica. Kazakhstan. Lepidolapha kuratavica (Asteraceae). 

Dracunculomyia tjulkubasica Fedotova 1999c: 834, nomen nudum. 

Dracunculomyia tjulkubasica Fedotova 1999f: 1313. Kazakhstan: Talasskiï Alatau, Tyulkubas canyon, nr Tyulkubas; 

HT; ♂; ZMAS.

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tomentella. Kazakhstan. Artemisia tomentella (Asteraceae). 

Dracunclomyia tomentella Fedotova 1999e: 1186. Kazakhstan: Arshakumy desert, 20 km NE Aral'ska, nr Shizhaga; 


tournefortiana. Kazakhstan. Artemisia tournefortiana (Asteraceae). 

Seriphidomyia tournefortiana Fedotova 1999c: 846, nomen nudum. 

Seriphidomyia tournefortiana Fedotova 2001a: 61. Kazakhstan: Aïgyrkumy desert, 6 km E Buran; HT; ♂; ZMAS. 

tridentatae. USA (California). Artemisia tridentata (Asteraceae). 

Rhopalomyia tridentatae Rübsaamen 1893: 163. USA: California; LT (Gagné 1983c: 71); ♂; ZMHB. 

tripleurospermi. Denmark, Czech Republic, Slovakia, Austria, Hungary, Greece, Russia (Europe); introd.: Canada 

(Alberta). Tripleurospermum perforatum (Asteraceae). 

Rhopalomyia tripleurospermi Skuhravá 2000: 427. Austria: Eisenstadt; HT; ♂; SKCR. 

truncata. USA (Massachusetts). 

Hormomyia truncata Felt 1907c: 160. USA: Massachusetts, nr Boston; HT; ♂; NYSM. 

Rhopalomyia uniformis Felt 1915j: 244, unjust. new name for Hormomyia truncata Felt 1907c. 

truncula. USA (California). 

Rhopalomyia truncata Felt 1908b: 365, jun. sec. hom. truncata Felt 1907c. USA: California, Los Angeles; ST; ♂; 


Rhopalomyia truncula Foote 1965: 267, new name for Rhopalomyia truncata Felt 1908b. 

tubifex. Widespread Europe, Algeria, Tunisia, Libya, Japan. Artemisia campestris (Asteraceae). 

Cecidomyia tubifex Bouché 1847: 144. Germany; ST; ♀,P,; ZMHB?. 

tubulus. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia tubulus Gagné 1983b: 25. USA: Idaho, Gooding Co., 9 mi. W Bliss; HT; ♂; USNM. 

tumidibulla. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia tumidibulla Gagné 1983b: 31. USA: Idaho, Elmore Co., 3 mi. N Mountain Home; HT; ♂; USNM. 

tumidicaulis. USA (Idaho). Artemisia tridentata (Asteraceae). 

Rhopalomyia tumidicaulis Gagné 1983b: 65. USA: Idaho, Canyon Co., 15 mi. S Nampa; HT; ♂; USNM. 

uniloculata. Kazakhstan. Artemisia scoparia (Asteraceae). 

Rhopalomyia uniloculata Fedotova 1984b: 36. Kazakhstan: Zailiïskiï Alatau, Syugaty Mtn; HT; ♂; ZKAS. 

*ussurica. Russia (Far East). Artemisia scoparia (Asteraceae). 

Dracunculomyia ussurica Fedotova 2004e: 21. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*ussurica. Russia (Far East). Artemisia rubripes (Asteraceae). 

Seriphidomyia ussurica Fedotova 2005a: 455. Russia: Primorskiï Terr., 23 km SE Ussuriysk; HT; ♂; ZMAS. If this 

species remains in Rhopalomyia, it will need to be renamed because is is junior to Dracunculomyia ussurica. 

ustjurtensis. Kazakhstan. Artemisia arenaria (Asteraceae). 

Dichelonyx ustjurtensis Fedotova 1999c: 846, nomen nudum. 

Dichelonyx ustjurtensis Fedotova 2001d: 958. Kazakhstan: western Ustyurt, Caspian Karakum Desert, 38 km NW 

Sarakamys; HT; ♂; ZMAS. 

utahensis. USA (Utah). Chrysothamnus graveolens (Asteraceae). 

Rhopalomyia utahensis Felt 1916e: 186. USA: Utah, nr Murray; ST; ♂,♀; NYSM. 

valichanovi. Kazakhstan. Artemisia austriaca (Asteraceae). 

Seriphidomyia valichanovi Fedotova 2001a: 64. Kazakhstan: Dzhungarskiï Alatau, Altyn-Emel' pass; HT; ♂; ZMAS. 

vamberi. Kazakhstan. Artemisia arenaria (Asteraceae). 

Pupascleromyia vamberi (also as vamberii) Fedotova 2001e: 1093. Kazakhstan: Buzachi Peninsula, 10 km NE 

Shebir; HT; ♂; ZMAS. 

verticillata. Kazakhstan. Artemisia schrenkiana (Asteraceae). 

Seriphidomyia verticillata Fedotova 2000d: 1434. Kazakhstan: Altaï, 70 km SW Georgievka, 175 km SW 

Ust'Kamenogorsk; HT; ♂; ZMAS. 

weldi. USA (Illinois). Aster macrophyllus (Asteraceae). 

Rhopalomyia weldi Felt 1921b: 116. USA: Illinois, Glencoe; ST; ♂,♀,P,L; NYSM. 

yomogicola. Korea, Japan (Hokkaido to Okinawa). Artemisia sp.; A. japonica, A. montana, A. princeps, Artemisia spp. 

(Asteraceae). Ref.: Nohara et al. 2007 (distr., biol., redescr.).

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Asynapta yomogicola Matsumura 1931: 403. Japan: Hokkaido, Sapporo; ST; ♀; Matsumura. 

yrgajtensis. Kazakhstan. Artemisia tournefortiana (Asteraceae). 

Dracunculomyia yrgajtensis Fedotova 1999c: 844. Kazakhstan: Dzhungarskiï Alatau, upper Yrgaïty R., 40 km E 

Koktumy; HT; ♂; ZMAS. 

Dracunculomyia yrgaitensis Fedotova 2000c: 1321, missp. of yrgajtensis. 

RIVERAELLA 

Riveraella Kieffer & Herbst 1911: 698. Type species, colliguayae Kieffer & Herbst (orig. des. as n. g., n. sp.). 


colliguayae. Chile. Colliguaja odorifera (Euphorbiaceae). 

Riveraella colliguayae Kieffer & Herbst 1911: 698. Chile: Valparaíso; T; ♀,P,L; lost. 

ROCHADIPLOSIS 

Rochadiplosis Tavares 1917a: 33. Type species, tibouchinae Tavares (orig. des.). 

Roachadiplosis Felt 1931: 449, missp. of Rochadiplosis. 

Roachodiplosis Mani 1946: 219, missp. of Rochadiplosis. 


tibouchinae. Brazil (Rio de Janeiro). Tibouchina sp. (Melastomataceae). 

Rochadiplosis tibouchinae Tavares 1917a: 34. Brazil: Rio de Janeiro, Petropolis, Tijuca, & Nova Friburgo; ST; 

♂,♀,P,L; TAVC. 

RONDANIOLA 

Rondaniella Rübsaamen 1916b: 493, preocc. Johannsen 1909. Type species, Cecidomyia bursaria Bremi (orig. des.). 

Rondaniola Hedicke 1938a: 290, new name for Rondaniella Rübsaamen. 

Western Palearctic; 1 sp.; Lasiopteridi, Dasineurini. 

bursaria. Widespread Europe. Glechoma hederacea (Lamiaceae). 

Cecidomyia bursaria Bremi 1847a: 52. Switzerland: vic. Zurich; ST; ♂,♀,L; ETHZ?. 

ROSOMYIA 

Rosomyia Fedotova 1987b: 36. Type species, spiraeae Fedotova (orig. des.). 

Palearctic; 3 spp., all on Rosaceae; Lasiopteridi, unplaced to tribe. 

oshanini. Kazakhstan. Spiraea media (Rosaceae). 

Rosomyia oshanini Fedotova 1996f: 55. Kazakhstan: southern Altaï, nr L. Markakol', nr Urunkhaïka, 60 km N 


pentaphylloidigemma. Kazakhstan. Pentaphylloides fruticosa (Rosaceae). 

Rosomyia pentaphylloidigemma Fedotova 1990b: 103. Kazakhstan: southern Altaï, northeastern shore L. Markakol', 

Kurchum Mtn, 10 km S Urunkhaïka; HT; ♂; ZMAS. 

spiraeae. Kazakhstan. Spiraea hypericifolia (Rosaceae). 

Rosomyia spiraeae Fedotova 1987b: 37. Kazakhstan: Dzhungarskiï Alatau, Altyn-Emel' pass, nr Konurolen; HT; ♂; 

ZMAS.

*ROTADIPLOSIS 

Rotadiplosis Fedotova & Sidorenko 2009a: 33. Type species, eximia Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe here. Fedotova and Sidorenko 2009a placed this genus in 

their tribe Filidiplosini but that classification is not used here (see Appendix). 

*eximia. Russia (Far East). 

Rotadiplosis eximia Fedotova & Sidorenko 2009a: 39 (erroneously as Flexidiplosis eximia on pp. 40-41). Russia: 

Primorskiï Terr., Lazovskiï Res., Proselochnaya Bay; HT; ♂; ZMAS. 

*RUIDADIPLOSIS 

Ruidadiplosis Fedotova & Sidorenko 2009b: 20. Type species, fluida Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe here, but Fedotova and Sidorenko 2009b place this single 

monotypic genus in their subtribe Ruidadiplosina but that classification is not used here (see Appendix). 

*fluida. Russia (Far East). 

Ruidadiplosis fluida Fedotova & Sidorenko 2009b: 22. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


SACKENOMYIA 

Sackenomyia Felt 1908b: 361. Type species, Rhopalomyia acerifoliae (as Oligotrophus acerifolius) Felt (orig. des.). 

Phlyctidobia Kieffer 1913a: 220. Type species, Oligotrophus solmsi Kieffer (orig. des.) = reaumurii Bremi. 

Holarctic; 5 spp., at least three on Viburnum, one on Ribes; Lasiopteridi, unplaced to tribe. 

acerifoliae. USA (New York). 

Rhopalomyia acerifoliae Felt 1907a: 25. USA: New York, Albany; HT; ♀; NYSM. 

Rhopalomyia acerifolia Felt 1907c: 121, unjust. emend. of acerifoliae. 

commota. USA (northeastern). Viburnum sp.; Viburnum spp.(Caprifoliaceae). 

Cystiphora viburnifolia Felt 1911m: 480, jun. sec. hom. Sackenomyia viburnifolia Felt 1909b. USA: Massachusetts, 

Magnolia; HT; ♀; NYSM. 

Sackenomyia commota Gagné 1975d: 12. new name for Cystiphora viburnifolia Felt 1911m. 

reaumurii. Widespread Europe. Viburnum lantana (Caprifoliaceae). 

Cecidomyia reaumurii Bremi 1847a: 19. Switzerland: vic. Zurich; ST; LT; ETHZ?. 

Oligotrophus solmsi Kieffer 1906c: 179. France: Moselle; ST; ♀,P,L,egg; lost?. 

ribesifolia. Kazakhstan. Ribes heterotrichum (Grossulariaceae). 

Sackenomyia ribesifolia Fedotova 1987c: 1108. Kazakhstan: Zailiïskiï Alatau, Topaïyr Mts, 150 km E Alma-Ata, 8 

km S Kok-Pek; HT; ♂; ZKAS. 

viburnifolia. USA (northeastern). Viburnum dentatum (Caprifoliaceae). 

Sackenomyia viburnifolia Felt 1909b: 290. USA: Massachusetts, Magnolia; ST; ♂,♀; NYSM. 

*SAGITTIDIPLOSIS 

Sagittidiplosis Fedotova 2004h: 163. Type species, tenua Fedotova (orig. des.). 

Eastern Palearctic; 2 spp.; Cecidomyiidi, Mycodiplosini. This genus does not differ from Mycodiplosis. 

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368 

*impreta. Russia (Far East). 

Sagittidiplosis impreta Fedotova 2004h: 165. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*tenua. Russia (Far East). 

Sagittidiplosis tenua Fedotova 2004h: 164. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

SALVALASIOPTERA 

Salvalasioptera Möhn 1975: 28. Type species, merremiae Möhn (orig. des.). 


merremiae. El Salvador. Merremia umbellata (Convolvulaceae). 

Salvalasioptera merremiae Möhn 1975: 29. El Salvador: Sonsonate, W Armenia; HT; ♂; SMNS. 

*SAMARADIPLOSIS 

Samaradiplosis Fedotova 2005c: 221. Type species, devexa Fedotova (orig. des.). 

Palearctic; 1 sp.; Cecidomyiidi, Clinodiplosini. 

*devexa. Russia (Far East). 

Samaradiplosis devexa Fedotova 2005c: 223. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. Kamenushka; HT; 

♂; ZMAS. 

SCHEUERIA 

Scheueria Kieffer & Herbst 1909: 120. Type species, longicornis Kieffer & Herbst (orig. des. as n. g., n. sp.). 

Scheuria Mani 1946: 202, missp. of Scheueria. 

Neotropical; 2 spp., on Baccharis; Lasiopteridi, unplaced to tribe. 

agglomerata. Chile. Baccharis eupatorioides (Asteraceae). 

Scheueria agglomerata Kieffer 1913i: 39. Chile: Concepción; T; G; lost. 

longicornis. Chile. Baccharis eupatorioides (Asteraceae). 

Scheueria longicornis Kieffer & Herbst 1909: 120. Chile: Concepción; ST; ♂,♀,P; lost. 

SCHISMATODIPLOSIS 

Schismatodiplosis Rübsaamen 1916a: 467. Type species, Clinodiplosis lantanae Rübsaamen (mon.). 


lantanae. Mexico (Veracruz, Tabasco, Quintana Roo), Guadeloupe, Trinidad, Brazil (Rio de Janeiro, Santa Catarina). 

Lantana sp. (Verbenaceae). 

Clinodiplosis lantanae Rübsaamen 1908a: 151. Brazil: Rio de Janeiro, Insel Cabo Frio & Palmeiras: Santa Catarina, 

Tubar�o; ST; ♂,♀,L; ZMHB. 

SCHIZANDROBIA 

Schizandrobia Kovalev 1964: 432. Type species, Tetrasphondylia arsenjevi Marikovskij & Kovalev (orig. des.).

Palearctic; 2 spp. on Schisandra; Cecidomyiidi, Asphondyliini, Schizomyiina. This genus could easily be subsumed 

under Schizomyia, q.v. for remarks. 

accidentalis. Russia (Far East). Schisandra chinensis (Schisandraceae). 

Schizandrobia accidentalis Kovalev 1964: 436. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; KOVC. 

arsenjevi. Russia (Far East). Schisandra chinensis (Schisandraceae). 

Tetrasphondylia arsenjevi Marikovskij & Kovalev 1962: 133. Russia: Primorskiï Terr.; ST; ♂,♀; MARC. 

SCHIZODIPLOSIS 

Schizodiplosis Kieffer 1913a: 233. Type species, Clinodiplosis floralis Kieffer & Herbst (orig. des.). 

Scizodiplosis Shinji 1944: 236, missp. of Schizodiplosis. 


floralis. Chile. Adults assoc. with flowers of an undet. Myrtaceae. 

Schizodiplosis floralis Kieffer & Herbst 1906: 233. Chile: vic. Concepción; T; ♀; lost. 

SCHIZOMYIA 

Schizomyia Kieffer 1889c: 183. Type species, galiorum Kieffer (mon.). 

Parasphondylia Kieffer 1913h: 93. Type species, variicornis Kieffer (orig. des.). 

Schyzomya, Schizomya Del Guercio 1818a: 126, missp. of Schizomyia. 

Shizomyia Shinji 1938k: 239, missp. of Schizomyia. 

Cosmopolitan; 53 spp., from a great diversity of plant hosts; Cecidomyiidi, Asphondyliini, Schizomyiina. Schizomyia 

serves as a catchall genus for species with needlelike ovipositors, four-segmented palpi, and larvae with all four pairs of 

terminal papillae present. The included species are such a broad representation within this definition that the species 

now in Asteralobia, Metasphondylia, Placochela, and Scizandrobia could easily fit under this umbrella. Möhn (1955a) 

distinguished larvae of Placochela from those of Schizomyia by the disposition of the papillae of the eighth abdominal 

segment. This distinction works with a small regional representation, but gradations between the two extremes can be 

seen in Schizomyia species known to me. Differences among these genera, including important and unnoted aspects of 

the ovipositor, need to be tabulated for all the species. Three Indian species formerly placed in Oxasphondylia were 

tentatively placed here by Gagné (2004b); they do not belong in Oxasphondylia. Ref.: Gagné and Menjivar 2008 

(review Neotrop. spp.). 

acaciae. India (Tamil Nadu). Acacia leucophloea (Fabaceae). 

Schizomyia acaciae Mani 1934b: 406. India: Tamil Nadu, Thanjavur; HT; ♀; ZSCI. 

acalyphae. Philippines. Acalypha stipulacea (Euphorbiaceae). 

Schizomyia acalyphae Felt 1918c: 286. Philippines: Luzon, Laguna Prov., Los Baños; ST; ♀; NYSM. 

Schizomyia acalypjae Rao 1955: 240, missp. of acalyphae. 

altifila. USA (New York). 

Asphondylia altifila Felt 1907a: 23. USA: New York, Lake Clear; HT; ♂; NYSM. 

assamensis. India (Assam), Indonesia (Java). Rubus assamensis (Rosaceae). 

Schizomyia assamensis Felt 1920b: 3. India: Assam, Shillong; ST; ♂,♀; NYSM. 

caryaecola. USA (New York). 

Schizomyia caryaecola Felt 1908b: 378. USA: New York; HT; ♀; NYSM. 

cheriani. India (Tamil Nadu). Ipomoea sepiaria (Convolvulaceae). 

Schizomyia cheriani Mani 1936: 194. India: Tamil Nadu, Madras; ST; ♀; ZSCI. 

cissusaeflorae. India (Tamil Nadu). Cissus vitigenea (Vitaceae). 

Schizomyia cissusaeflorae Mani 1986: 19. India: Tamil Nadu, Madras; HT; ♀; ZSCI. 

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370 

clerodendri. India (Tamil Nadu). Clerodendron enerme (Verbenaceae). 

Schizomyia clerodendri Mani 1986: 18. India: Tamil Nadu, Madras; HT; ♀; ZSCI. 

cocculi. India (Tamil Nadu). Cocculus hirsutus (Menispermaceae). 

Schizomyia cocculi Mani 1954b: 109. India: Tamil Nadu, Coimbatore, Marudamalai Hills; HT; ♂; ZSCI. 

cryptostegiae. Madagascar. Cryptostegia grandiflora; Pentapetia androsaemifolia, Leptadenia madagascariensis 

(Asclepiadaceae). 

Schizomyia cryptostegiae Gagné 1997c: 15. Madagascar: Tulear, Miary; HT; ♂; BMNH. 

diplodisci. Philippines. Diplodiscus paniculatus (Tiliaceae). 

Schizomyia diplodisci Felt 1918c: 287. Philippines: Luzon, Laguna Prov., Mt Makiling; ST; ♂,♀; NYSM. 

dosua. India (Himachal Pradesh). Indigofera dosua (Fabaceae). 

Oxasphondylia dosua Mani 1954a: 22. India: Himachal Pradesh, Kangra Distr., Dharamsala; HT; ♂; ZSCI. 

echinata. India (Himachal Pradesh). Indigofera gerardiana (Fabaceae). 

Oxasphondylia echinata Mani 1954a: 20. India: Himachal Pradesh, Kangra Distr., 10 mi. from Dharamsala; HT; ♂; 

ZSCI. 

ericae. South Africa. Erica sp. (Ericaceae). 

Schizomyia ericae Rübsaamen 1916a: 444. South Africa; T; ♀; ZMHB. 

eupatoriflorae. Canada (Ontario), USA (New York, West Virginia, Maryland, North Carolina). Ageratina altissima 

(Asteraceae). 

Cecidomyia eupatoriflorae Beutenmüller 1907a: 391. USA: New York, New York, Staten I., & North Carolina, 


floricola. India (Himachal Pradesh). Indigofera gerardiana (Fabaceae). 

Oxasphondylia floricola Mani 1954a: 25. India: Himachal Pradesh, Dalhousie; HT; ♀; ZSCI. 

galiorum. Widespread Europe, Algeria, Kazakhstan. Galium mollugo; Galium spp. (Rubiaceae). 

Schizomyia galiorum Kieffer 1889c: 184. France: Metz & Bitche; ST; ♂,♀,L; lost?. 

impatientis. USA (New York, Maine, Massachusetts, Maryland, Virginia). Impatiens capensis; I. pallida 

(Balsaminaceae). 

Cecidomyia impatientis Osten Sacken 1862: 204. USA: District of Columbia; T; LT; lost. 

incerta. India (West Bengal). Eurya japonica (Theaceae). 

Schizomyia incerta Kieffer 1909a: 153. India: West Bengal, Kurseong; T; LT; lost?. 


Schizomyia indica Kieffer 1909a: 154. India: West Bengal, Kurseong; T; LT; lost?. 

ipomoeae. Guadeloupe, St. Vincent. Ipomoea sp. (Convolvulaceae). 

Schizomyia ipomoeae Felt 1910b: 160. St. Vincent; ST; ♂,♀,P; NYSM. 

laporteae. Indonesia (Java). Laportea stimulans (Urticaceae). 

Schizomyia laporteae Felt 1921h: 143. Indonesia: Java, Ungaran Mts; ST; ♀,P; NYSM. 

*loroco. Guatemala to Costa Rica. Fernaldia pandurata (Apocynaceae). 

Schizomyia loroco Gagné 2008b: 286. El Salvador: Nueva Conceción, Chalatenango; HT; ♂; USNM. 

macarangae. India (Kerala). Macaranga indica (Euphorbiaceae). 

Schizomyia macarangae Nayar 1953: 131. India: Kerala, Trivandrum; HT; ♂; ZSCI. 

macrofila. USA (California). Amsinckia lycopsoides (Boraginaceae). 

Asphondylia macrofila Felt 1907d: 16. USA: California, Los Angeles & Alameda; ST; ♂,♀; USNM. 

maeruae. India (Tamil Nadu). Maerua arenaria (Capparidaceae). 

Schizomyia maeruae Felt 1926i: 243. India: Tamil Nadu, Coimbatore; HT; ♀; NYSM. 

manihoti. Colombia, Brazil (Ceará). Manihot utilissima (Euphorbiaceae). 

Schizomyia manihoti Tavares 1925a: 22. Brazil: Ceará; ST; ♂,♀,P,L; TAVC. 

*maricaensis. Brazil (Rio de Janeiro). Tetrapterys phlomoides (Malpighiaceae). 

Schizomyia maricaensis Sousa & Maia 2007: 348. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; 

QBUM. 

*microcapillata. Brazil (Minas Gerais). Bauhinea brevipes (Fabaceae). 

Schizomyia microcapillata Maia 2005d: 33. Brazil: Minas Gerais, Três Marias; HT; ♂; QBUM. 

nodosa. Indonesia (Java). Moschosma polystachum (Lamiaceae). 

Schizomyia nodosa Felt 1921h: 144. Indonesia: Java, Semarang; ST; ♂,♀,P,L; NYSM.

371 

orientalis. India (Uttar Pradesh). Acacia leucophloea (Fabaceae). 

Schizomyia orientalis Grover 1967a: 261. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

petiolicola. USA (New York, Massachusetts). Vitis bicolor (Vitaceae). 

Schizomyia petiolicola Felt 1908b: 379. USA: New York, Westfield; ST; ♂,♀; NYSM. 

psoraleae. South Africa. Psoralea pinnata (Fabaceae). 

Schizomyia psoraleae Rübsaamen 1910b: 32. South Africa: Transvaal, Lydenburg Distr., nr Flussufer; T; LT; 

ZMHB. 

racemicola. USA (widespread eastern). Solidago sp.; Solidago spp., principally nemoralis (Asteraceae). 

Cecidomyia racemicola Osten Sacken 1862: 196. USA: District of Columbia; T; LT; lost. 

rivinae. USA (Florida). Rivina humilis (Phytolaccaceae). 

Schizomyia rivinae Felt 1908b: 379. USA: Florida, Georgiana; HT; ♀; NYSM. 

rubi. USA (New York). 

Schizomyia rubi Felt 1907a: 23. USA: New York, Karner; HT; ♀; NYSM. 

*samaralukensis. Russia (Europe). Tripleurospermum perforatum (Asteraceae). 

Schizomyia samaralukensis Fedotova 2007: 466. Russia: Middle Volga River Area, Zhiguli Nature Res., nr 

Bakhilova Polyana, 15 km SE Zol'noe; HT; ♀; ZMAS. 

*santosi. Brazil (Rio de Janeiro). Jacquemontia holosericea (Convolvulaceae). 

Schizomyia santosi Maia & Araújo 2009: 357. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

scheppigi. South Africa. Stoebe cinerea (Asteraceae). 

Schizomyia scheppigi Rübsaamen 1910b: 36. South Africa; ST; P,L; ZMHB. 

serjaniae. El Salvador. Serjania goniocarpa (Sapindaceae). 

Schizomyia serjaniae Möhn 1960c: 235. El Salvador: San Salvador; HT; ♀; SMNS. 

sesami. Tanzania. Sesamum angustifolium (Pedaliaceae). 

Schizomyia sesami Barnes 1939b: 456. Tanzania: Nyakato, Bukoba; ST; ♂,♀; BMNH. 

speciosa. USA (New Hampshire). 

Schizomyia speciosa Felt 1914d: 112. USA: New Hampshire, Franconia; HT; ♀; NYSM. 

*spherica. Brazil (Rio de Janeiro). Sebastiana glandulosa (Euphorbiaceae). 

Schizomyia spherica Maia & Oliveira 2007: 97. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

stachytarphetae. Guadeloupe, Trinidad. Stachytarpheta cayennensis; S. jamaicensis (Verbenaceae). 

Schizomyia stachytarphetae Barnes 1932b: 483. Trinidad; ST; ♂,♀; BMNH. 

tami. Italy. Tamus communis (Dioscoreaceae). 

Schizomyia tami Kieffer 1902b: 523. Italy; T; G; lost?. 

tuiuiu. Brazil (Mato Grosso). Bauhinea rufa (Fabaceae). 

Schizomyia tuiuiu Urso-Guimarães & Amorim 2002: 568. Brazil: Mato Grosso, Cuiabá, Universidade Federal de 

Mato Grosso campus; HT; ♀; MZSP. 

umbellicola. USA (widespread eastern). Sambucus racemosa (Caprifoliaceae). 

Cecidomyia sambuci umbellicola Osten Sacken 1870: 52, invalid, a polynomial. 

Cecidomyia umbellicola Osten Sacken 1878: 7. USA: New York & New Jersey; T; G; lost. 

Cecidomyia sambuciumbellicola (as sambuci-umbellicola) Beutenmüller 1892: 269, emend. of sambuci umbellicola. 

variicornis. Kenya. 

Parasphondylia variicornis Kieffer 1913h: 93. Kenya: Shimoni; HT; ♀; MNHN. 

viburni. USA (New York). Viburnum sp. (Caprifoliaceae). 

Schizomyia viburni Felt 1908b: 378. USA: New York, Old Forge; HT; ♀; NYSM. 

villebrunneae. Indonesia (Java). Villebrunnea rubescens (Urticaceae). 

Schizomyia villebrunneae Felt 1921h: 146. Indonesia: Java, Ungaran Mts; ST; ♂,♀,P,L; NYSM. 

viticola. Nearctic (widespread eastern). Vitis sp.; Vitis spp. (Vitaceae). 

Cecidomyia viticola Osten Sacken 1862: 202. USA: District of Columbia; T; LT; lost. 

Cecidomyia vitis lituus Walsh & Riley 1870: 28, invalid, a polynomial. 

Cecidomyia vitislituus Osten Sacken 1878: 7. USA: New York & Pennsylvania; T; G; USNM. 

Cecidomyia lituus Felt 1911l: 458, error for vitislituus. 

Dasineura vitis Shinji 1944: 179, error for viticola, recorded from Japan only on basis of similarity of galls. 

vitiscoryloides. Nearctic (widespread eastern). Vitis cordifolia; Vitis spp. (Vitaceae).

372 

Cecidomyia vitis coryloides Walsh & Riley 1869: 106, invalid, a polynomial. 

Cecidomyia vitiscoryloides Packard 1869: 377. USA: Illinois, Rock Island; T; G; USNM. 

Cecidomyia coryloides Felt 1906a: 745, error for vitiscoryloides. 

vitispomum. USA (widespread eastern). Vitis cordifolia; Vitis spp. (Vitaceae). 

Cecidomyia vitis pomum Walsh & Riley 1869: 106, invalid, a polynomial. 

Cecidomyia vitispomum (as vitis-pomum) Osten Sacken 1878: 7. USA: Illinois, Alton, & Pennsylvania; T; LT; 


Schizomyia pomum Felt 1908b: 379, error for vitispomum. 

SCIASPHONDYLIA 

Sciasphondylia Möhn 1960c: 221. Type species, salviae Möhn (orig. des.). 


salviae. El Salvador. Salvia sp. (Lamiaceae). 

Sciasphondylia salviae Möhn 1960c: 222. El Salvador: Chalatenango, 15 km. from La Palma; HT; ♀; SMNS. 

SCOPODIPLOSIS 

Scopodiplosis Felt 1915a: 209. Type species, speciosa Felt (orig. des.). 


speciosa. Paraguay. 

Scopodiplosis speciosa Felt 1915a: 210. Paraguay: San Bernardino; HT; ♀; USNM. 

SCORZONEROMYIA 

Scorzoneromyia Fedotova 1991d: 20. Type species, Dasineura scorsonerae Fedotova (orig. des.). 


scorsonerae. Kazakhstan. Scorzonera pusilla (Asteraceae). 

Dasyneura scorsonerae Fedotova 1982a: 45. Kazakhstan: Sary-Taukumy Desert; HT; ♂; ZMAS. 

SEMUDOBIA 

Semudobia Kieffer 1913g: 55. Type species, Cecidomyia betulae Winnertz (orig. des.). 

Holarctic; 7 spp., all from Betula; Lasiopteridi, unplaced to tribe. Ref.: Roskam 1977 (rev.). 

betulae. Widespread Palearctic; immigr.: Canada (Ontario), USA (Pennsylvania). Betula pendula; Betula spp. 

(Betulaceae). 

Cecidomyia betulae Winnertz 1853: 234. Germany; NT (Roskam 1977: 168; Netherlands: Renkum); ♂; RMNH. 

brevipalpis. Widespread northeastern Nearctic. Betula papyrifera; B. populifolia (Betulaceae). 

Semudobia brevipalpis Roskam 1977: 169. Canada: Quebec, along Trans-Canada Highway; HT; ♂; RMNH. 

markakolica. Kazakhstan. Betula pendula (Betulaceae). 

Semudobia markakolica Fedotova 1996c: 38. Kazakhstan: southern Altaï, nr L. Markakol', nr Urunkhaïka, 60 km N 


rotundifoliae. Kazakhstan. Betula rotundifolia (Betulaceae). 

Semudobia rotundifoliae Fedotova 1996c: 36. Kazakhstan: central Altaï, Koksuïskiï Mtn Range, Mt. Lymin Belok,

373 

60 km NE Leninogorsk; HT; ♂; ZMAS. 

skuhravae. Widespread Holarctic. Betula pendula; Betula spp. (Betulaceae). 

Semudobia skuhravae Roskam 1977: 174. Netherlands: Renkum; HT; ♂; RMNH. 

steenisi.Widespread northwestern Nearctic. Betula occidentalis (Betulaceae). 

Semudobia steenisi Roskam 1977: 170. USA: Wyoming, Fremont Co., Wind R., 10 mi N Dubois; HT; ♂; RMNH. 

tarda. Widespread Palearctic; immigr.: Canada (Quebec), USA (Idaho). Betula pendula, Betula spp. (Betulaceae). 

Semudobia tarda Roskam 1977: 172. Netherlands: Renkum; HT; ♂; RMNH. 

SEQUOIOMYIA 

Sequoiomyia Möhn 1960b: 515. Type species, kraeuseli Möhn (orig. des.). 

Holarctic; 2 spp., one a Pliocene fossil, from Taxodiaceae; Cecidomyiidi, Cecidomyiini. 

† kraeuseli. Pliocene (coal) - Germany. Sequoia langsdorfii (Taxodiaceae). 

Sequoiomyia kraeuseli Möhn 1960b: 515. Pliocene/Germany: in coal, Düren; HT; LT; SMNS. 

taxodii. Widespread southeastern Nearctic. Taxodium distichum (Taxodiaceae). 

Retinodiplosis taxodii Felt 1916f: 415. USA: Missouri, Charleston; LT (Gagné 1968e: 271); ♂; NYSM. 

*SETODIPLOSIS 

Setodiplosis Fedotova 2006a: 153. Type species, unifaria Fedotova (orig. des.). 


*unifaria. Russia (Far East). 

Setodiplosis unifaria Fedotova 2006a: 153. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

*SHARMADIPLOSIS 

Sharmadiplosis Fedotova & Sidorenko 2009b: 11. Type species, Odontodiplosis orientalis Sharma & Rao (orig. 

des.). 

Oriental; 1 sp.; Cecidomyiidi, Mycodiplosini. Fedotova and Sidorenko 2009a placed this genus in their subtribe 

Quadridiplosina but that classification is not used here (see Appendix). 


Odontodiplosis orientalis Sharma & Rao 1980a: 299. India: Maharashtra, Aurangabad; HT; ♂; ELMU. 

*SICITURADASTRA 

Sicituradastra Fedotova & Sidorenko 2009a: 28. Type species, proboscidea Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe here. The gender of the name was unstated but was 

presumably meant to be feminine because the type species as designated had the feminine form. Fedotova and 

Sidorenko 2009a placed this genus in their tribe Filidiplosini but that classification is not used here (see Appendix). 

*proboscidea. Russia (Far East). 

Siciturasastra proboscidea Fedotova & Sidorenko 2009a: 30. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya 


*SIGNASICCA

Signasicca Fedotova & Sidorenko 2009d: 246. Type species, subita Fedotova & Sidorenko (orig. des.). 


*subita. Russia (Far East). 

Signasicca subita Fedotova & Sidorenko 2009d: 246 (as T[uguridiplosis] subita, p. 229). Russia: Primorskiï Terr., 

Lazovskiï Res., Koreiskaya Pad'; HT; ♂; ZMAS. 

SILVESTRIOLA 

Silvestrina Kieffer 1912d: 172, preocc. Prewer 1904. Type species, Arthrocnodax silvestrii Kieffer (orig. des.) = 

cincta Felt. 

Silvestria Shinji 1944: 379, missp. of Silvestrina. 

Silvectria Shinji 1944: 87, missp. of Silvestrina. 

Silvestriola Skuhravá 1997: 170, new name for Silvestrina Kieffer. 

Cosmopolitan; 14 spp.; Cecidomyiidi, unplaced to tribe. The species are all surely predators but some species are 

known only from having been reared from particular plant hosts. 

artemisiae. Japan (Honshu). Artemisia princeps (Asteraceae). 

Sitodiplosis artemisiae Shinji 1939h: 228. Japan: Honshu; ST; ♂,♀; lost. 

asphodeli. Italy, Greece. Asphodelus sp. (Liliaceae). 

Silvestrina asphodeli Barnes 1935b: 72. Italy: Sardinia, Cagliari; ST; ♂,♀; BMNH. 

beardsleyi. Hawaiian Is. 

Microdiplosis beardsleyi Hardy 1960: 296. Hawaiian Is: Oahu, Ewa; HT; ♂; HDOA. 

chinagliana. Italy. Diaspis sp. (Hemiptera: Diaspididae). 

Silvestrina chinagliana Del Guercio 1918a: 259. Italy; ST; ♂,♀; lost. 

cincta. Cosmopolitan. With mites assoc. with bees (Acarina); Pseudaulacaspis pentagona (Hemiptera: Diaspididae). 

Cecidomyia cincta Felt 1907a: 47. USA: New York, Albany; HT; ♂; NYSM. 

Cecidomyia macrofila Felt 1907: 21. USA: New Mexico, Las Vegas; HT; ♂; NYSM. 

Arthrocnodax apiphila Felt 1907d: 20. USA: California; HT; ♂; NYSM. 

Arthrocnodax silvestrii Kieffer 1910b: 133. South Africa: Cape Town; T; ♀; lost?. 

Silvestrina cecconiana, as var. of silvestrii, Kieffer 1920b: 296. Italy; ST; ♂,♀; lost?. 

euphorbiae. Japan (Hokkaido to Kyushu). Hydrangea paniculata (Hydrangeaceae). 

Contarinia euphorbiae Shinji 1939g: 163. Japan: Honshu, Morioka; T; A; lost. 

Silvestria euphobiae Shinji 1944: 381, missp. of euphorbiae. 

farinicola. UK, Netherlands, Italy. Dermatophagoides farinae (Acarina: Pyroglyphidae). 

Arthrocnodax farinicola Barnes 1929: 121. Italy: Naples; ST; ♂,♀; BMNH. 

ficorum. Algeria. On dried figs (Moraceae). 

Silvestrina ficorum Barnes 1932b: 476. Algeria: Algiers, Bougie; ST; ♂,♀; BMNH. 

lobata. India (Uttar Pradesh). 

Silvestrina lobata Grover 1965a: 30. India: Uttar Pradesh, Allahabad, Tagore Town; HT; ♂; PGCA. 

*mescheryakovi. Russia (Far East). Pseudaulacaspis cockerelli (Homoptera: Diaspididae) 

Silvestrina mescheryakovi Fedotova 2004f: 100. Russia: Primorskiï Terr., nr Vladivostok; HT; ♂; ZMAS. 

minima. Germany. 

Diplosis minima Rübsaamen 1891c: 50. Germany: Giersberg nr Siegen; ST; ♂; ZMHB. 

quercifoliae. Japan (Honshu). Quercus grandulifera (Fagaceae). 

Silvestrina quercifoliae Shinji 1944: 383. Japan: Honshu; T; A; lost. 

Diplosis mitsukurii Shinji 1944: 125, nomen nudum. 

Silvestrina quercicola Shinji 1944: 126, missp. of quercifoliae. 

spatulata. India (Uttar Pradesh). 

374

Silvestrina spatulata Grover & Bakhshi 1978: 58. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

tyrophagi. Ukraine. Tyrophagus noxius (Acarina: Acaridae). 

Silvestrina tyrophagi Dombrovskaja 1940a: 87. Ukraine: Glukhov; ST; ♂,♀,L; ZMAS. 

SITODIPLOSIS 

Sitodiplosis Kieffer 1913g: 49. Type species, Cecidomyia mosellana Géhin (orig. des.). 

Palearctic, immigr. Nearctic; 4 spp., all associated with grasses; Cecidomyiidi, Clinodiplosini. Refs.: Harris 1966 

(review), Gagné & Doane 1999 (larvae). 

cambriensis. UK. Poa trivialis (Poaceae). 

Sitodiplosis cambriensis Jones 1940: 542. UK: Wales, Cardiganshire, & England, Montgomeryshire; ST; ♂,♀; 

BMNH. 

dactylidis. UK. Dactylis glomerata (Poaceae). 

Sitodiplosis dactylidis Barnes 1940b: 112. UK: Northern Ireland, Co. Armagh & Co. Fermanagh, & England, 

Shropshire & Leicestershire; ST; ♂,♀; BMNH. 

mosellana. Widespread Palearctic; immigr.: widespread Nearctic. Triticum aestivum; T. turgidum, T. vulgare, 

(Poaceae). 

Cecidomyia mosellana Géhin 1857: 21. France: Moselle; ST; LT; lost. 

Diplosis aurantiaca Wagner 1866: 82. Germany: Fulda; ST; ♂,♀,L; lost. 

phalaridis. UK. Phalaris arundinacea (Poaceae). 

Sitodiplosis phalaridis Abbass 1986: 65. UK: England, Manchester, Didsbury, Fletcher Moss; HT; ♂; BMNH. 

SKUSEMYIA 

Skusemyia Kolesik 1995b: 41. Type species, allocasuarinae Kolesik (orig. des.). 


allocasuarinae. Australia (South Australia). Allocasuarina verticillata (Casuarinaceae). 

Skusemyia allocasuarinae Kolesik 1995b: 42. Australia: South Australia, Black Hill Conservation Pk, 13 km NE 


SMILASIOPTERA 

Smilasioptera Möhn 1975: 10. Type species, candelariae Möhn (orig. des.). 


candelariae. El Salvador. Smilax mexicana (Smilacaceae). 

Smilasioptera candelariae Möhn 1975: 11. El Salvador: Santa Ana, SE Candelaria, Hacienda Cortez; HT; ♂; SMNS. 

SOPHOROMYIA 

Sophoromyia Fedotova 1984c: 47. Type species, goebeliae Fedotova (orig. des.). 

Palearctic; 2 spp., from Fabaceae; Lasiopteridi, unplaced to tribe. 

armenica. Armenia. Glycyrrhiza glabra (Fabaceae). 

Sophoromyia armenica Mamaev & Mirumian 1989: 45. Armenia: Abovyanskiy Region, Dzhrvezhskoe; HT; ♂; 

HYNM. 

375

376 

goebeliae. Kazakhstan. Goebelia pachycarpa (Fabaceae). 

Sophoromyia goebeliae Fedotova 1984c: 48. Kazakhstan: Karatau Mtn Range, Abaï R. valley, 15 km NE 

Babaïkurgan; HT; ♂; ZKAS. 

SPARTIOMYIA 

Spartiomyia Kieffer 1913i: 57. Type species, Janetiella martinsi Tavares (mon.). 

Sportiomyia Felt 1931: 435, missp. of Spartiomyia. 

Spaltiomyia Shinji 1944: 219, missp. of Spartiomyia. 


martinsi. Portugal. Echinospartum ibericum (Fabaceae). 

Janetiella martinsi Tavares 1901: 63. Portugal: Gardunha; ST; ♂,♀,P,L; TAVC. 

*SPHAERAMYIA 

Sphaeramyia Maia 2007a: 452. Type species, flava Maia (orig. des.). 

Neotropical; 1 sp.; Lasiopteridi, unplaced to tribe. See notes under Guarephila about possible placement of this 

genus. 

*flava. Brazil (São Paulo). Guarea macrophylla (Meliaceae). 

Sphaeramyia flava Maia 2007a: 453. Brazil: São Paulo, Bertioga, Fazenda Pinto; HT; ♂; QBUM. 

SPHAERODIPLOSIS 

Sphaerodiplosis Rübsaamen 1916a: 461. Type species, dubia Rübsaamen (mon.). 

Neotropical; 1 sp.; Cecidomyiidi, Cecidomyiini. 

dubia. Brazil. 

Sphaerodiplosis dubia Rübsaamen 1916a: 461. Brazil; T; ♂; ZMHB. 

*SPICULATIDIPLOSIS 

Spiculatidiplosis Fedotova 2003e: 54. Type species, longistylis Fedotova (orig. des.). 

Eastern Palearctic; 4 spp.; Cecidomyiidi, Clinodiplosini. This genus is close to Clinodiplosis if the two must be 

separated. 

*fastigiata. Russia (Far East). 

Spiculatidiplosis fastigiatus Fedotova 2003e: 56. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*longistylus. Russia (Far East). 

Spiculatidiplosis longistylis Fedotova 2003e: 56. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*mira. Russia (Far East). 

Spiculatidiplosis mira Fedotova 2004h: 172. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*superposita. Russia (Far East).

377 

Spiculatidiplosis superposita Fedotova & Sidorenko 2006c: 136. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; 

♂; ZMAS. 

SPILODIPLOSIS 

Spilodiplosis Kovalev & Mamaev 1966: 228. Type species, occidentalis Kovalev & Mamaev (orig. des.). 

Palearctic; 1 sp.; Cecidomyiidi, Aphidoletini. 

occidentalis. Russia (Far East). 

Spilodiplosis occidentalis Kovalev & Mamaev 1966: 229. Russia: Primorskiï Terr., Suputinskiy Res.; HT; ♂; 

HYNM. 

SPIRAEANTHOMYIA 

Spiraeanthomyia Fedotova 1989d: 23. Type species, karatavica Fedotova (orig. des.). 

Palearctic; 2 spp., from Rosaceae; Lasiopteridi, Dasineurini. The distinction between this genus and Dasineura (s.l.) 

is not apparent from the original description. 

hulthemiae. Kazakhstan. Hulthemia persica (Rosaceae). 

Spiraeanthomyia hulthemiae Fedotova 1992a: 81. Kazakhstan: 75 km NW Kyzl-Orda; HT; ♂; ZMAS. 

karatavica. Kazakhstan. Spiraeanthus schrenkianus (Rosaceae). 

Spiraeanthomyia karatavica Fedotova 1989d: 24. Kazakhstan: northwestern Karatau Mtn Range, Abaï valley, 15 km 

NE Babaïkurgan; HT; ♂; ZKAS. 

SPIROMYIA 

Spiromyia Fedotova 1991d: 20. Type species, Dasineura cystiphorae Fedotova (orig. des.). 


cystiphorae. Kazakhstan. Spiraea hypericifolia (Rosaceae). 

Dasineura cystiphorae Fedotova 1985f: 32. Kazakhstan: Kirgizskiï Alatau, S Merke; HT; ♂; ZKAS. 

*SPIRODIPLOSIS 

Spirodiplosis Fedotova 2006a: 163. Type species, implexa Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Lestodiplosini. This genus is close to Lestodiplosis if the two must be 

separated. 

*implexa. Russia (Far East). 

Spirodiplosis implexa Fedotova 2006a: 164. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

*SPUNGISIOLA 

Spungisiola Fedotova & Perkovsky 2008b: 67. Type species, insuperabilis Fedotova & Perkovsky (orig. des.). 

Rovno amber (Ukraine; late Eocene); 1 sp.; originally placed with the Brachineuridi, q.v., but its placement below the 

level of subfamily Cecidomyiinae is not at all certain from its description.

* † insuperabilis. Rovno amber (Ukraine). 

Spungisiola insuperabilis Fedotova & Perkovsky 2008b: 67. Ukraine amber: Rovno; HT; ♀; UASK. 

SPURGIA 

Spurgia Gagné 1990b: 335. Type species, Cecidomyia capitigena Bremi (orig. des.) = euphorbiae Vallot. 

Palearctic, two spp. introduced into Nearctic; 8 spp., all on Euphorbia; Lasiopteridi, Dasineurini. 

alatavica. Kazakhstan. Euphorbia alatavica (Euphorbiaceae). 

Spurgia alatavica (also as altaica) Fedotova 1994i: 449. Kazakhstan: Tarbagataï Mtn Range, 40 km NW Urdzhara, 

nr Blagodatnoye; HT; ♂; ZMAS. 

esulae. Italy; introd.: USA (Montana, North Dakota). Euphorbia esula (Euphorbiaceae). 

Spurgia esulae Gagné 1990b: 340. Italy: Pisa, San Rossore; HT; ♂; USNM. 

euphorbiae. Widespread Europe; immigr. with introd. stock of S. esulae: Canada, USA (North Dakota). Euphorbia 

cyparissias (Euphorbiaceae). 

Cecidomyia euphorbiae Vallot 1827: 94. France; ST; G; lost. Vallot's description of the gall matches that of this 

species exactly. 

Cecidomyia capitigena Bremi 1847a: 50. Switzerland: vic. Zurich; LT (Gagné 1990b: 337); ♀; ETHZ. 

Cecidomyia euphorbiae Loew 1850a: 36. Poland; ST; ♂,♀; ZMHB?. 

Cecidomyia subpatula Bremi 1847a: 50. Switzerland: Zurich; LT (Gagné 1990b: 337); ♂; ETHZ. 

gagnei. Kazakhstan. Euphorbia soongarica; E. virgata (Euphorbiaceae). 

Spurgia gagnei Fedotova 1994i: 453. Kazakhstan: border Dzhambul Prov. & Kirghizia, Karabalt R. valley, Stepnoye; 


lamprocarpae. Kazakhstan. Euphorbia lamprocarpa (Euphorbiaceae). 

Spurgia lamprocarpae Fedotova 1994i: 452. Kazakhstan: Tabargatay Mtn Range, Tentek R. valley, nr Uch-Aral; HT; 

♂; ZMAS. 

macrorrizae. Kazakhstan. Euphorbia macrorrhiza (Euphorbiaceae). 

Spurgia macrorrizae Fedotova 1996c: 34. Kazakhstan: southern Altaï, nr L. Markakol', nr Urunkhaïka, 60 km N 


pachyrizae. Kazakhstan. Euphorbia pachyrhiza (Euphorbiaceae). 

Spurgia pachyrizae Fedotova 1994i: 450. Kazakhstan: Terskey Alatau, upper Bayankol R., 28 km SW Narynkol; HT; 

♂; ZMAS. 

seguierianae. Kazakhstan. Euphorbia seguierianae (Euphorbiaceae). 

Spurgia sequierianae Fedotova 1994i: 451. Kazakhstan: Buzachi Peninsula, 10 km NE Shebir; HT; ♂; ZMAS. 

*STABILIOLA 

Stabiliola Fedotova & Sidorenko 2006b: 14. Type species, serrata Fedotova & Sidorenko (orig. des.). 


*serrata. Russia (Far East). 

Stabiliola serrata Fedotova & Sidorenko 2006b: 14. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; 

♂; ZMAS. 

STEFANIELLA 

Stefaniella Kieffer 1898f: 55. Type species, atriplicis Kieffer (subs. des. Felt 1911: 42). 

Steffaniela Shinji 1939c: 588, missp. of Stefaniella. 

Palearctic, 9 spp., eight on Atriplex, one on Chenopodium (Chenopodiaceae); Lasiopteridi, Lasiopterini. 

378

atriplicis. Italy, Czechoslovakia, Jugoslavia, Greece, Russia (Europe), Algeria. Atriplex halimus; A. patula 


Stefaniella atriplicis Kieffer 1898f: 56. Algeria; ST; ♂,♀; lost?. 

brevipalpis. UK, circum-Mediterranean. Atriplex portulacoides; A. patula (Chenopodiaceae). 

Stefaniella brevipalpis Kieffer 1898f: 56. Israel: Akko; N (Dorchin & Freidberg 2008: 15); ♀; TAUI. Ref.: Dorchin 

& Freidberg 2008 (redescr., biol.). 

cecconii. Italy, Czechoslovakia, Romania, Yugoslavia. Atriplex patula (Chenopodiaceae). 

Stefaniella cecconii Kieffer 1909b: 2. Italy; T; LT; lost?. 

gigantea. Turkmenistan. Chenopodium vulvaria (Chenopodiaceae). 

Stefaniella gigantea Fedotova 1992d: 29. Turkmenistan; Kugitang Mtn Range, nr Charshanga; HT; ♀; ZMAS. 

hilversidae. Armenia, Kazakhstan. Atriplex salina; A. tatarica (Chenopodiaceae). 

Stefaniella hilversidae Mamaev 1972a: 887. Kazakhstan: Bugun'; HT; ♀; ZMAS. 

kazenasae. Kazakhstan. Atriplex verrucifera (Chenopodiaceae). 

Stefaniella kazenasae Fedotova 1991b: 42. Kazakhstan: Alakul lowland, 16 km N Uch-Aral, 11 km S Rybach'e; HT; 

♀; ZMAS. 

pusilla. Kazakhstan. Atriplex micrantha (Chenopodiaceae). 

Stefaniella pusilla Fedotova 1994h: 29. Kazakhstan: Ili R. estuary, 14 km N Karoï; HT; ♀; ZMAS. 

trinacriae. Spain, Italy, Malta, Greece, Algeria, Israel. Atriplex halimus (Chenopodiaceae). 

Stefaniella trinacriae De Stefani 1900: 8. Italy: Sicily; ST; ♂,♀; lost. Ref.: Dorchin & Freidberg 2008: 16 (redescr.). 

ustjurtensis. Kazakhstan. Atriplex aucherii; A. nitens (Chenopodiaceae). 

Stefaniella ustjurtensis Fedotova 1991b: 43. Kazakhstan: northwestern Ustyurt, 90 km SW Sam-1, Karasai R. valley, 

Kumin L.; HT; ♀; ZMAS. 

STEFANIOLA 

Stefaniola Kieffer 1913g: 45. Type species, Stefaniella salsolae Tavares (orig. des.). 

Baldratiola Kieffer 1913g: 45. Type species, Baldratia houardi Kieffer (orig. des.). 

Dibaldratia Kieffer 1913g: 46. Type species, bilobata Kieffer (orig. des.). 

Salsolomyia Tavares 1919a: 17. Type species, parva Tavares (orig. des.). 

Haloxylonomyia Marikovskij 1953b: 331. Type species, deformans Marikovskij (orig. des.). 

Haloxylonobia Marikovskij 1955c: 126, missp. of Haloxylonomyia. 

Haloxylomyia Marikovskij 1955d: 300, missp. of Haloxylonomyia. 

Aridofavilla Mamaev 1972a: 888. Type species, bicuspidata Mamaev (orig. des.). 

Palearctic; 109 spp., all on Chenopodiaceae, most associated with simple leaf galls but some in complex stem and bud 

galls; Lasiopteridi, Lasiopterini. Many additional species are known but undescribed (Dorchin, in litt.). Refs: Fedotova 

1989c (key to spp.), Dorchin 2001 (redescr.). Gagné 2004b (syn. note). 

accola. Kazakhstan, Turkmenistan. Haloxylon ammodendrum, H. aphyllum (Chenopodiaceae). 

Haloxylonomyia accola Marikovskij 1953b: 340. Kazakhstan: Ili & Syr-Darya R. valleys; ST; ♂,♀; ZMAS. 

adaptata. Kazakhstan. Arthrophytum lehmannianum (Chenopodiaceae). 

Stefaniola adaptata Fedotova 1992e: 41. Kazakhstan: Sengir-Kum Desert, 45 km NW Senek; HT; ♀; ZMAS. 

adstricta. Iraq. Salsola spissa (Chenopodiaceae). 

Stefaniola adstricta Möhn 1971: 231. Iraq: Baghdad; HT; LT; SMNS. 

aelleniafoliae. Kazakhstan. Aellenia glauca (Chenopodiaceae). 

Stefaniola aelleniafoliae Fedotova 1989a: 71. Kazakhstan: western Ustyurt, Ustyurtskiï Res., 50 SE Akkuduk; HT; 

♀; ZKAS. 

africana. Eritrea. Suaeda fruticosa; S. baccata (Chenopodiaceae). 

Stefaniola africana Möhn 1971: 247. Eritrea: Daukalia, Assab; HT; LT; SMNS. 

anabasidigemmae. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Stefaniola anabasidigemmae Fedotova 1989a: 63. Kazakhstan: Buzachi Peninsula, 60 km NW Staryï Kalamkas; HT; 

379

380 

♀; ZKAS. 

angusta. Russia (Eastern Siberia). Salsola collina (Chenopodiaceae). 

Stefaniola angusta Möhn 1971: 230. Russia: Eastern Siberia, Nerchinsk; HT; LT; SMNS. 

arbuscula. Kazakhstan, Turkmenistan. Salsola arbuscula (Chenopodiaceae). 

Stefaniola arbuscula Möhn 1971: 230. Turkmenistan; HT; LT; SMNS. 

arthrophytumicola. Kazakhstan. Arthrophytum iliense; A. lehmannianum (Chenopodiaceae). 

Stefaniola arthrophytumicola Fedotova 1989a: 63. Kazakhstan: upper Charyn R., nr Chundzha; HT; ♀; ZKAS. 

arthrophytumilaesa. Kazakhstan. Arthrophytum lehmannianum; A. longibracteatum (Chenopodiaceae). 

Stefaniola arthrophytumilaesa Fedotova 1992e: 43. Kazakhstan: Sengir-Kum Desert, 45 km NW Senek; HT; ♀; 

ZMAS. 

asiatica. Turkmenistan, Uzbekistan. Salsola rigida; S. orientalis (Chenopodiaceae). 

Dibaldratia asiatica Marikovskij 1957: 941. Turkmenistan: Mangyr-Tai, old delta of Amu-Darya; ST; ♀; MARC. 

aurata. Kazakhstan, Turkmenistan, Uzbekistan. Haloxylon ammodendrum, H. aphyllum; H. persicum 


Haloxylonomyia aurata Marikovskij 1953b: 337. Kazakhstan: Ili R. valley; ST; ♂,♀; ZKAS & ZMAS. 

badhyzica. Turkmenistan. Salsola richteri (Chenopodiaceae). 

Stefaniola badhyzica Fedotova 1992l: 69. Turkmenistan: Badkhyzskiï Res., 100 km NW Gushgi; HT; ♀; ZMAS. 

bamianica. Afghanistan. Salsola bamianica (Chenopodiaceae). 

Stefaniola bamianica Möhn 1971: 232. Afghanistan: Bamian; HT; LT; SMNS. 

bassiagemmae. Turkmenistan. Bassia hyssopifolia (Chenopodiaceae). 

Stefaniola bassiagemmae Fedotova 1993h: 50. Turkmenistan: Kugitang Mtn Range, 25 km SW Kugitang, 55 km NE 

Charshanga; HT; ♀; ZMAS. 

begimbetovae. Kazakhstan. Salsola nitraria (Chenopodiaceae). 

Stefaniola begimbetovae Fedotova 1993e: 79. Kazakhstan: 10 km SE Ulan-Bel', Chu R. floodplain; HT; ♀; ZMAS. 

bicuspidata. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Aridofavilla bicuspidata Mamaev 1972a: 889. Turkmenistan: Mary, Akibay; HT; ♀; ZMAS. 

bilobata. Spain, Morocco, Algeria. Salsola vermiculata (Chenopodiaceae). 

Dibaldratia bilobata Kieffer 1913g: 46. Spain; ST; ♂,♀; lost?. 

brevis. Tunisia, Israel. Traganum nudatum (Chenopodiaceae). 

Stefaniola brevis Möhn 1971: 245. Tunisia: Oasis Nefta; HT; LT; SMNS. 

camphorosmae. Kazakhstan. Camphorosma monspeliacum (Chenopodiaceae). 

Stefaniola camphorosmae (also as camphorasmae) Fedotova 1992b: 47. Kazakhstan: middle part Ili R., 20 km NW 

Bakanas; HT; ♂; ZMAS. 

caspica. Kazakhstan. Arthrophytum longibracteatum; A. lehmannianum (Chenopodiaceae). 

Stefaniola caspica Fedotova 1989b: 86. Kazakhstan: eastern Mangyschlak basin, 18 km E Senek; HT; ♀; ZKAS. 

caulicola. Israel. Anabasis setifera (Chenopodiaceae). 

Stefaniola caulicola Möhn 1971: 237. Israel: Engedi; HT; LT; SMNS. 

chiwensicola. Kazakhstan. Salsola chiwensis (Chenopodiaceae). 

Stefaniola chiwensicola Fedotova 1998b: 14. Kazakhstan: eastern Mangyshlak basin, Ustyurtskiï Res., 140 km E 

Novyï-Uzen', 45 km S Akkuduk; HT; ♀; ZMAS. 

clavifaciens. Kazakhstan. Haloxylon ammodendrum, H. aphyllum (Chenopodiaceae). 

Haloxylonomyia clavifaciens, as subspecies of deformans, Marikovskij 1953b: 334. Kazakhstan: Ili & Syr-Darya R. 

valleys; ST; ♂,♀; ZKAS & ZMAS. 

climacopterae. Kazakhstan. Climacoptera crassa (Chenopodiaceae). 

Stefaniola climacopterae Fedotova 1989b: 85. Kazakhstan: Karakumy Desert, 22 km SW Opornaya Stn; HT; ♀; 

ZKAS. 

completa. Turkmenistan. Salsola richteri (Chenopodiaceae). 

Stefaniola completa Fedotova 1992k: 72. Turkmenistan: Badkhyzskiï Res., 100 km NW Kushki; HT; ♀; ZMAS. 

congregata. Russia (Western Siberia), Kazakhstan, Turkmenistan, Uzbekistan. Haloxylon ammodendrum, H. aphyllum 


Haloxylonomyia congregata, as subspecies of deformans, Marikovskij 1953b: 334. Kazakhstan: Ili R. valley; ST; 

♂,♀; ZKAS & ZMAS.

381 

*crispa. Israel. Arthrocnemum macrostachyum (Chenopodiaceae). 

Stefaniola crispa Dorchin & Freidberg 2008: 20. Israel: Akko; HT; ♀; TAUI. 

davletshinae. Turkmenistan. Salsola rigida; S. orientalis (Chenopodiaceae). 

Careopalpis davletshinae Marikovskij 1957: 943. Turkmenistan: Mangyr-Tai, old delta of Amu-Darya R.; T; ♀; 

MARC. 

defoliata. Israel, Egypt. Suaeda monoica (Chenopodiaceae). 

Stefaniola defoliata Dorchin 2001: 574. Israel: Ne'ot HaKikkar; HT; ♀; TAUI. 

deformans. Kazakhstan, Turkmenistan, Uzbekistan, Afghanistan, Iran. Haloxylon ammodendrum, H. aphyllum; 

Haloxylon spp. (Chenopodiaceae). 

Haloxylonomyia deformans Marikovskij 1953b: 333. Kazakhstan: Ili & Syr-Darya R. valleys; ST; ♂,♀; ZKAS & 

ZMAS. 

Haloxylomyia solitaria, as ssp. of deformans, Marikovskij 1955c: 124, nomen nudum. 

diversa. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola diversa Mamaev 1972a: 891. Turkmenistan: Mary, Akibay; HT; ♀; ZMAS. 

domina. Kazakhstan, Turkmenistan. Haloxylon ammodendrum, H. aphyllum (Chenopodiaceae). 

Haloxylonomyia domina Marikovskij 1953b: 338. Kazakhstan: Ili R. valley; ST; ♂,♀; ZKAS & ZMAS. 

dubia. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola dubia Mamaev 1972a: 892. Turkmenistan: Mary, Akibay; HT; ♀; ZMAS. 

dupla. Libya. Haloxylon salicornium (Chenopodiaceae). 

Stefaniola dupla Möhn 1971: 217. Libya: Uadi Tininai; HT; LT; SMNS. 

elatiorae. Kazakhstan. Anabasis elatior (Chenopodiaceae). 

Stefaniola elatiorae Fedotova 1993e: 77. Kazakhstan: Aïgyrkumy desert, 6 km E Buran, nr Ardynka; HT; ♀; ZMAS. 

excelsa. Armenia, Iran, Kazakhstan, Turkmenistan, Uzbekistan. Aellenia glauca; A. subaphylla (Chenopodiaceae). 

Stefaniola excelsa Möhn 1971: 235. Armenia: Maman; HT; LT; SMNS. 

fistula. Israel, Iraq. Seidlitzia rosmarinus (Chenopodiaceae). 

Stefaniola fistula Möhn 1971: 247. Israel: Wadi Zmvivah, nr Dead Sea; HT; LT; SMNS. 

foliata. Central Asia. Aellenia auricula (Chenopodiaceae). 

Stefaniola foliata Möhn 1971: 233. Central Asia; HT; LT; SMNS. 

folicola. Israel. Anabasis setifera (Chenopodiaceae). 

Stefaniola folicola Möhn 1971: 238. Israel: M. Baghik, nr Dead Sea; HT; LT; SMNS. 

foliosae. Kazakhstan. Salsola foliosa (Chenopodiaceae). 

Stefaniola foliosae Fedotova 1992f: 47. Kazakhstan: Ustyurt, 45 km SE Opornaya Stn; HT; ♀; ZMAS. 

fragosa. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola fragosa Mamaev 1972a: 890. Turkmenistan: Mary, Akibay; HT; ♀; ZMAS. 

fraudulenta. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola fraudulenta Mamaev 1972a: 892. Turkmenistan: Mary, Akibay; HT; ♀; ZMAS. 

fructua. Kazakhstan, Turkmenistan, Azerbaijan, Iran. Aellenia glauca; A. cinerascens (Chenopodiaceae). 

Stefaniola fructua Möhn 1971: 234. Iran: Hableh-Rud valley, central Elburs; HT; LT; SMNS. 

furtiva. Kazakhstan, Turkmenistan, Iran. Haloxylon ammodendrum, H. aphyllum (Chenopodiaceae). 

Haloxylonomyia furtiva Marikovskij 1953b: 338. Kazakhstan: Ili R. valley; ST; ♂,♀; ZKAS & ZMAS. 

gigas. Kazakhstan, Turkmenistan, Uzbekistan, Iran, Mongolia. Haloxylon ammodendrum, H. aphyllum 


Haloxylonomyia gigas Marikovskij 1953b: Kazakhstan: Ili & Syr-Darya R. valleys; ST; ♂,♀; ZKAS & ZMAS. 

globosa. Turkmenistan, Uzbekistan. Haloxylon ammodendron (Chenopodiaceae). 

Stefaniola globosa Möhn 1971: 226. Turkmenistan: Bala-Ischem; HT; LT; SMNS. 

gloma. Spain, Algeria, Egypt, Israel, Syria, Iraq. Salsola vermiculata; Salsola spp. (Chenopodiaceae). 

Stefaniola gloma Möhn 1971: 207. Algeria: Biskra; HT; LT; SMNS. 

halimocnemiis. Kazakhstan. Halimocnemis sclerocarpa (Chenopodiaceae). 

Stefaniola halimocnemiis Fedotova 1992b: 48. Kazakhstan: NE Priaral'e , 14 km SE Saksaulnaya RR Stn; HT; ♀; 

ZMAS. 

halocharii. Kazakhstan. Halocharis hispida (Chenopodiaceae). 

Stefaniola halocharii Fedotova 1994h: 30. Kazakhstan: Dzhusan-Dala desert, 40 km SE Aksuek; HT; ♀; ZMAS.

382 

halogetonicola. Kazakhstan. Halogeton glomeratus (Chenopodiaceae). 

Stefaniola halogetonicola Fedotova 1992f: 48. Kazakhstan: Saur Mtn Range, S L. Zaïsan, nr Karasu; HT; ♂; ZMAS. 

halothamnis. Turkmenistan. Halothamnus hispidulus (Chenopodiaceae). 

Stefaniola halothamnis Fedotova 1992d: 38. Turkmenistan: Bolshie Balkhany Mts, nr NebitDag; HT; ♀; ZMAS. 

hammadae. Turkmenistan. Hammada leptoclada (Chenopodiaceae). 

Stefaniola hammadae Fedotova 1993n: 11. Turkmenistan: Kugitang Mtn Range, 55 km NE Charshanga; HT; ♀; 

ZMAS. 

heterobia. Kazakhstan, Turkmenistan. Haloxylon ammodendrum, H. aphyllum (Chenopodiaceae). 

Haloxylonomyia heterobia Marikovskij 1953b: 339. Kazakhstan: Ili & Syr-Darya R. valleys; ST; ♂,♀; ZKAS & 

ZMAS. 

houardi. Tunisia. Bassia muricata (Chenopodiaceae). 

Baldratia houardi Kieffer 1912e: 66. Tunisia: Gafsa; ST; ♀,P,egg; lost? 

iliensis. Kazakhstan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola iliensis Fedotova 1989a: 65. Kazakhstan: mid Ili R., nr Bakanas; HT; ♀; ZKAS. 

inobservabilis. Kazakhstan, Turkmenistan, Uzbekistan. Haloxylon ammodendrum, H. aphyllum (Chenopodiaceae). 

Haloxylonomyia inobservabilis, as subspecies of gigas, Marikovskij 1953b: 336. Kazakhstan: Ili & Syr-Darya R. 

valleys; ST; ♂,♀; ZKAS & ZMAS. 

insignata. Kazakhstan, Turkmenistan, Uzbekistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola insignata Mamaev 1972a: 892. Turkmenistan: central Karakumy Desert, Dzhar; HT; ♀; ZMAS. 

insignis. Kazakhstan, Turkmenistan, Uzbekistan. Haloxylon ammodendrum, H. aphyllum (Chenopodiaceae). 

Haloxylonomyia insignis, as subspecies of gigas, Marikovskij 1953b: 336. Kazakhstan: Ili & Syr-Darya R. valleys; 

ST; ♂,♀; ZMAS. 

junia. Turkmenistan. Salsola richteri (Chenopodiaceae). 

Stefaniola junia Fedotova 1992l: 68. Turkmenistan: Sundukli desert, Kyzyl-Burun L., 80 km SE Chardzheva; HT; ♀; 

ZMAS. 

kaplini. Turkmenistan. Salsola richteri (Chenopodiaceae). 

Stefaniola kaplini Fedotova 1991f: 14. Turkmenistan: eastern Karakumy Desert, 70 km SW Chardzhou; HT; ♀; 

ZMAS. 

kovalevi. Kazakhstan. Climacoptera brachiata; C. affinis (Chenopodiaceae). 

Stefaniola kovalevi Fedotova 1989b: 84. Kazakhstan: Karakumy Desert, 22 km SW Opornaya Stn; HT; ♀; ZKAS. 

kugitangica. Turkmenistan. Hammada leptoclada (Chenopodiaceae). 

Stefaniola kugitangica Fedotova 1993n: 13. Turkmenistan: Kugitang Mtn Range, 55 km NE Charshanga; HT; ♀; 

ZMAS. 

kyzylkumensis. Kazakhstan. Salsola richteri (Chenopodiaceae). 

Stefaniola kyzylkumensis Fedotova 1991f: 12. Kazakhstan: northeastern Kyzylkum Desert, 24 km SW Kyrkense; HT; 

♀; ZMAS. 

lepidosa. Turkmenistan, Uzbekistan. Salsola rigida (Chenopodiaceae). 

Stefaniola lepidosa Mamaev 1972a: 892. Uzbekistan: Bukhara Prov., Kul'dzhuktau; HT; ♀; ZMAS. 

libanensis. Lebanon. Salsola canescens (Chenopodiaceae). 

Stefaniola libanensis Möhn 1971: 231. Lebanon; HT; LT; SMNS. 

lignea. Afghanistan. Haloxylon multiflorum (Chenopodiaceae). 

Stefaniola lignea Möhn 1971: 216. Afghanistan; HT; LT; SMNS. 

mangyshlakensis. Kazakhstan. Arthrophytum lehmannianum; A. longibracteatum (Chenopodiaceae). 

Stefaniola mangyshlakensis Fedotova 1989a: 61. Kazakhstan: eastern Mangyshlak basin, 60 km NE Senek; HT; ♀; 

ZKAS. 

Stefaniola mangyshlakensus Fedotova 1989c: 89, missp. of mangyshlakensis. 

mediterranea. Italy, Algeria. Salicornia europaea; Halocnemum strobilaceum (Chenopodiaceae). 

Stefaniola mediterranea Möhn 1971: 243. Italy: Venice; HT; LT; SMNS. 

miscella. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola miscella Mamaev 1972a: 892. Turkmenistan: Mary, Akibay; HT; ♀; HYNM. 

moehni. Kazakhstan. Salsola arbuscula (Chenopodiaceae). 

Stefaniola moehni (as möhni) Fedotova 1992d: 36. Kazakhstan: 5 km SE Karabas well, 22 km NW Novyy Uzen';

383 


nanophytonicola. Kazakhstan. Nanophyton erinaceum (Chenopodiaceae). 

Stefaniola nanophytonicola Fedotova 1989a: 68. Kazakhstan: central Ustyurt, 71 km SE Akzhigit; HT; ♀; ZKAS. 

nitrariae. Kazakhstan. Salsola nitraria (Chenopodiaceae). 

Stefaniola nitrariae Fedotova 1989b: 82. Kazakhstan: western Ustyurt, 38 km NE Sarykamys; HT; ♀; ZKAS. 

nodosa. Iraq. Halocnemum strobilaceum (Chenopodiaceae). 

Stefaniola nodosa Möhn 1971: 243. Iraq: Hor Hammah, vic. Djelibah; HT; LT; SMNS. 

nucula. Algeria. Nucularia perrinii (Chenopodiaceae). 

Stefaniola nucula Möhn 1971: 246. Algeria: Oued Inçoki; HT; LT; SMNS. 

ochracea. Kazakhstan, Turkmenistan. Salsola richteri (Chenopodiaceae). 

Stefaniola ochracea Fedotova 1992l: 69. Kazakhstan: northeastern Kyzylkumy, nr Karaktau, 64 km NW Bairkum; 


opulenta. Morocco, Tunisia, "Palestine". Anabasis aphylla; A. haussknechtii (Chenopodiaceae). 

Stefaniola opulenta Möhn 1971: 240. "Palestine": Mashetta; HT; LT; SMNS. 

orientalis. Kazakhstan. Salsola orientalis (Chenopodiaceae). 

Stefaniola orientalis Fedotova 1989b: 79. Kazakhstan: Ustyurt, 70 km NW Beïneu; HT; ♀; ZKAS. 

orta. Uzbekistan, Afghanistan. Haloxylon ammodendron (Chenopodiaceae). 

Stefaniola orta Möhn 1971: 225. Afghanistan; HT; LT; SMNS. 

pakae. Turkmenistan. Salsola dendroides (Chenopodiaceae). 

Stefaniola pakae Fedotova 1992d: 38. Turkmenistan: foothills Kugitang Mtn Range, nr Charshanga; HT; ♀; ZMAS. 

parva. Spain. Salsola vermiculata (Chenopodiaceae). 

Salsolomyia parva Tavares 1919a: 18. Spain: Saragossa; T; ♀; TAVC. 

petrosimoniae. Kazakhstan. Petrosimonia sibirica, P. triandra (Chenopodiaceae). 

Stefaniola petrosimoniae Fedotova 1989b: 82. Kazakhstan: Karatal R. valley, 60 km NW Ush-Tobe; HT; ♀; ZKAS. 

polita. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola polita Mamaev 1972a: 890. Turkmenistan: Mary, Akibay; HT; ♀; HYNM. 

praestans. Kazakhstan. Salsola richteri (Chenopodiaceae). 

Stefaniola praestans Fedotova 1991f: 11. Kazakhstan: northeastern Kyzylkum, 81 km SW Kzyl-Orda; HT; ♀; 

ZMAS. 

procera. Azerbaijan, Kazakhstan. Anabasis aphylla (Chenopodiaceae). 

Stefaniola procera Möhn 1971: 219. Azerbaijan: nr Kur R.; HT; LT; SMNS. 

psammophila. Turkmenistan. Haloxylon persicum (Chenopodiaceae). 

Stefaniola psammophila Mamaev 1972a: 891. Turkmenistan: central Karakumy Desert; HT; ♀; ZMAS. 

punjabensis. Pakistan/India. Salsola foetida (Chenopodiaceae). 

Stefaniola punjabensis Möhn 1971: 208. "Punjab"; HT; LT; SMNS. 

pusilla. Kazakhstan, Turkmenistan, Uzbekistan, Mongolia. Haloxylon ammodendrum, H. aphyllum (Chenopodiaceae). 

Haloxylonomyia pusilla Marikovskij 1953b: 337. Kazakhstan: mid Ili R.; ST; ♂,♀; ZKAS & ZMAS. 

ramicola. Turkmenistan, Uzbekistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola ramicola Mamaev 1972a: 889. Turkmenistan: Mary, Akibay; HT; ♀; ZMAS. 

ramosa. Iraq. Haloxylon persicum (Chenopodiaceae). 

Stefaniola ramosa Möhn 1971: 225. Iraq: Ur/Al Busaiya; HT; LT; SMNS. 

rotunda. Kazakhstan, China (Xinjiang-Uygur Aut. Reg.). Arthrophytum lehmannianum; Arthrophytum spp. 


Stefaniola rotunda Möhn 1971: 227. Kazakhstan; HT; LT; SMNS. 

rufa. Israel, ?Eritrea. Suaeda monoica (Chenopodiaceae). 

Stefaniola rufa Dorchin 2001: 576. Israel: Newe Zohar; HT; ♀; TAUI. 

salsolae. Spain, Portugal. Salsola vermiculata, S. microphylla (Chenopodiaceae). 

Stefaniella salsolae Tavares 1904a: 293. Spain: Huesca, Sierra de Guara; ST; ♂,♀,egg; TAVC. 

salsolicarpae. Kazakhstan. Salsola paulsenii (Chenopodiaceae). 

Stefaniola salsolicarpae Fedotova 1989a: 70. Kazakhstan: Karakumy Desert, 38 km SW Sarykamys; HT; ♀; ZKAS. 

shubenkinae. Turkmenistan. Salsola chiwensis (Chenopodiaceae). 

Stefaniola shubenkinae (also as schubenkinae) Fedotova 1994d: 44. Turkmenistan: southern Ustyurt, Burchliburun

384 

escarpment, 300 km W Tashauza; HT; ♀; ZMAS. 

siliqua. Israel, Eritrea. Suaeda monoica (Chenopodiaceae). 

Stefaniola siliqua Dorchin 2001: 579. Israel: Newe Zohar; HT; ♀; TAUI. 

similata. Turkmenistan, Uzbekistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola similata Mamaev 1972a: 890. Turkmenistan: Mary, Akibay; HT; ♀; ZMAS. 

spinosa. Iran, Iraq. Aellenia glauca; A. subaphylla (Chenopodiaceae). 

Stefaniola spinosa Möhn 1971: 236. Iran: Sharad-Bustan; HT; LT; SMNS. 

subterminalis. Kazakhstan, Turkmenistan. Salsola richteri (Chenopodiaceae). 

Stefaniola subterminalis Mamaev & Pak 1989: 35. Turkmenistan: Sarykamysh; HT; ♀; HYNM. 

transversa. Afghanistan. Haloxylon multiflorum (Chenopodiaceae). 

Stefaniola transversa Möhn 1971: 229. Afghanistan; HT; LT; SMNS. 

tripla. Algeria. Haloxylon schmittianum (Chenopodiaceae). 

Stefaniola tripla Möhn 1971: 218. Algeria: Biskra; HT; LT; SMNS. 

tscharynensis. Kazakhstan. Anabasis salsa (Chenopodiaceae). 

Stefaniola tscharynensis Fedotova 1989a: 66. Kazakhstan: mid Ili R., 20 km NW Bakanas; HT; ♀; ZKAS. 

tubifera. Turkmenistan. Haloxylon aphyllum (Chenopodiaceae). 

Stefaniola tubifera Mamaev 1975e: 73. Turkmenistan: Mary, Akibay; HT; ♀; MAMC. 

turcistana. Central Asia. Haloxylon persicum (Chenopodiaceae). 

Stefaniola turcistana Möhn 1971: 224. "Turkistan"; HT; LT; SMNS. 

unita. Algeria, Tunisia, Egypt. Haloxylon schweinfurthii; H. schmittianum (Chenopodiaceae). 

Stefaniola unita Möhn 1971: 217. Egypt: nr Suez; HT; LT; SMNS. 

ustjurtensis. Kazakhstan. Anabasis eriopoda (Chenopodiaceae). 

Stefaniola ustjurtensis Fedotova 1989a: 66. Kazakhstan: western Ustyurt, Ustyurtskiï Res., 60 km NW Akkuduk; HT; 

♀; ZKAS. 

vastita. Spain, Algeria, Egypt, Israel, Jordan. Anabasis articulata; A. prostrata (Chenopodiaceae). 

Stefaniola vastita Möhn 1971: 238. Algeria: Laghouat; HT; LT; SMNS. 

ventriosa. Tunisia, Morocco, Egypt, Israel. Traganum nudatum (Chenopodiaceae). 

Stefaniola ventriosa Möhn 1971: 245. Tunisia: Gabès; HT; LT; SMNS. 

vexillata. Kazakhstan, Turkmenistan, Uzbekistan. Salsola richteri (Chenopodiaceae). 

Stefaniola vexillata Mamaev & Pak 1989: 34. Uzbekistan: Shafirkan; HT; ♀; HYNM. 

Stefaniola vexibia Fedotova 2000b: 646, missp. of vexillata. 

STENODIPLOSIS 

Stenodiplosis Reuter 1895: 9. Type species, geniculati Reuter (subs. des. Kieffer 1896: 92). 

Tinconaria Gagné 1967b: 155. Type species, albescentis Gagné (orig. des.). 

Allocontarinia Solinas 1986: 23. Type species, Diplosis sorghicola Coquillett (orig. des.). 

Cosmopolitan; 10 spp., all from inflorescences of grasses; Cecidomyiidi, Cecidomyiini. This genus may include also 

some species now under Contarinia that are associated with grasses. Refs: Harris 1976 (illus.), Solinas 1986 (illus.). 

albescentis. USA (Texas, South Carolina), Mexico. Triodia flava (Poaceae). 

Tinconaria albescentis Gagné 1967b: 155. USA: Texas, Brownsville; HT; ♀; USNM. 

bromicola. Germany, Poland, Ukraine, Russia (Europe); immigr.: Canada (Saskatchewan, Alberta), USA (Oklahoma, 

Missouri, Wisconsin, Maryland). Bromus inermis (Poaceae). 

Stenodiplosis bromicola Marikovskij & Agafonova 1961: 272. Russia: Kursk Prov.; ST; ♂,♀; Leningrad. 

dactylidis. New Zealand (presumed immigr.). Dactylis glomerata (Poaceae). 

Stenodiplosis dactylidis, as var. of geniculati, Barnes 1940b: 114. New Zealand: Auckland (Mt. Albert), Wellington, 

& Canterbury; ST; ♂,♀; BMNH. 

gambae. Nigeria. Andropogon gayanus (Poaceae). 

Contarinia gambae Harris 1964: 242. Nigeria: Zaria; HT; ♂; BMNH. 

geniculati. UK, Sweden, Finland, Poland, Bulgaria; immigr.: Canada (Ontario, New Brunswick), USA (South Dakota),

385 

New Zealand. Alopecurus geniculatus; A. arundinaceus, A. pratensis (Poaceae). 

Stenodiplosis geniculati Reuter 1895: 10. Finland: Kirchspiel Pargas, nr Åbo; ST; ♂,♀,P,L; ?. 

panici. Yugoslavia, Ukraine, Russia (Europe), Kazakhstan, Uzbekistan. Panicum miliaceum, P. crus-galli (Poaceae). 

Stenodiplosis panici Plotnikov 1926: 236 (attributed to B.B. Rohdendorf but such description as is present is by 

Plotnikov). Uzbekistan: Tashkent, & Kazakhstan: Chimkent Prov.; ST; ♂,♀,P; ?. 

sorghi. Senegal, Nigeria, Uganda. Sorghum arundinaceum; S. guineense, S. verticilliflorum (Poaceae). 

Stenodiplosis sorghi Harris 1964: 244. Nigeria: Ilugun; HT; ♂; BMNH. 

sorghicola. Africa, India (Tamil Nadu), Philippines (Luzon); immigr.: USA (southern), Mexico, Dominican Republic, 

Argentina (Buenos Aires) . Originated in Africa, now found throughout the world betw. latitudes 40 deg. N & 

S, wherever sorghum is grown (Harris 1970). Sorghum bicolor; Sorghum spp. (Poaceae). 

Contarinia sorghicola Coquillett 1899: 82. USA: Texas, College Station; ST; ♂,♀; USNM. 

Contarinia andropoginis Felt 1921j: 25. India: Tamil Nadu, Coimbatore; ST; ♂,♀; NYSM. 

Contarinia saltata Felt 1919a: 289. Philippines: Luzon, Laguna Prov., Los Baños; LT (Gagné 1969e: 562); ♂; 

NYSM. 

Contarinia palposa Blanchard 1958: 423. Argentina: Buenos Aires; ST; ♂,♀; DPBA. 

Contarinia andropogonis Harris 1964: 238, missp. of andropoginis. 

stenotaphri. Mauritius. Stenotaphrum dimidiatum (Poaceae). 

Stenodiplosis stenotaphri Barnes 1944b: 211. Mauritius: Reduit; ST; ♂,♀; BMNH. 

wattsi. USA (South Dakota, Nebraska, New Mexico, Florida). Schizachyrium scoparium; A. gerardii, A. hallii, 

Sorghastrum nutans (Poaceae). 

Contarinia wattsi Gagné 1966b: 319. USA: New Mexico, Los Lunas; HT; ♂; USNM. 

Contarinia halliicola Gagné 1966b: 319. USA: New Mexico, Los Lunas; HT; ♂; USNM. 

*STENOHYPODIPLOSIS 

Stenohypodiplosis Fedotova 2005c: 220. Type species, sejuncta Fedotova (orig. des.). 

Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. The single known species described for this genus does not appear 

to differ from Dichodiplosis langeni Rübsaamen. 

*sejuncta. Russia (Far East). 

Stenohypodiplosis sejuncta Fedotova 2005c: 221. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. Kamenushka; 


STEPHODIPLOSIS 

Stephodiplosis Tavares 1908: 140. Type species, lanneae Tavares (mon.). 

Afrotropical, Australasian; 2 spp.; Cecidomyiidi, unplaced to tribe. 

lanneae. Mozambique. Lannea fulva (Anacardiaceae). 

Stephodiplosis lanneae Tavares 1908: 140. Mozambique; ST; ♂,♀,P; TAVC. 

nothofagi. New Zealand. Nothofagus cliffortioides (Fagaceae). 

Stephodiplosis nothofagi Barnes 1936: 276. New Zealand: Cass; ST; ♂,♀; BMNH. 

STEPHOMYIA 

Stephomyia Tavares 1916a: 54. Type species, eugeniae Tavares (orig. des.). 

Neotropical; 7 spp., all from leaf galls on Myrtaceae; Cecidomyiidi, Asphondyliini, Schizomyiina. 

clavata. Brazil (Bahia). An unknown Myrtaceae.

386 

Oxasphondylia clavata Tavares 1920f: 125, pl. III. Brazil: Bahia, Roça da Madre de Deus, vic. Salvador; T; ♂; 

TAVC. 

epeugeniae. Brazil (Rio de Janeiro). Eugenia sp. (Myrtaceae). 

Stephomyia eugeniae Tavares 1916a: 54, jun. sec. hom. eugeniae Felt. Brazil: Rio de Janeiro, Botanical Garden; LT 

(Möhn 1962: 234); ♂; TAVC. 

Stephomyia epeugeniae Gagné 1994a: 144, new name for Stephomyia eugeniae Tavares. 

espiralis. Brazil (Rio de Janeiro). Eugenia capacabanensis (Myrtaceae). 

Stephomyia espiralis Maia 1994b: 526. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♀; QBUM. 

eugeniae. USA (Florida). Eugenia buxifolia (Myrtaceae). 

Cystodiplosis eugeniae Felt 1913a: 175. USA: Florida, Key West; ST; ♂,♀,P; USNM. 

mina. Brazil (Rio de Janeiro). Neomitranthes obscura (Myrtaceae). 

Stephomyia mina Maia 1994b: 528. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

rotundifoliorum. Brazil (Rio de Janeiro). Eugenia rotundifolia (Myrtaceae). 

Stephomyia rotundifoliorum Maia 1994b: 523. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♂; QBUM. 

tetralobae. Brazil (Rio de Janeiro). Eugenia capacabanensis (Myrtaceae). 

Stephomyia tetralobae Maia 1994b: 525. Brazil: Rio de Janeiro, Restinga de Barra de Maricá; HT; ♀; QBUM. 

STERRHAULUS 

Sterrhaulus Rübsaamen 1916b: 490. Type species, Cecidomyia corneola Rübsaamen (orig. des.). 


corneolus. Germany, Russia (Europe). Carex sp. (Cyperaceae). 

Cecidomyia corneola Rübsaamen 1899b: 545. Germany; ST; ♀,L; ZMHB. 

STICTOBREMIA 

Stictobremia Kieffer 1912h: 138. Type species, campylomyzae Kieffer (orig. des.). 


campylomyzae. France. Peromyia leveillei (Diptera: Cecidomyiidae). 

Stictobremia campylomyzae Kieffer 1912b: 138. France: Bitche; ST; ♂,♀; lost?. 

*STIGMODIPLOSIS 

Stigmodiplosis Fedotova & Sidorenko 2006c: 124. Type species, plicata Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 3 spp.; Cecidomyiidi, Lestodiplosini. This genus is close to Lestodiplosis if the two must be 

separated. 

*additionalis. Russia (Far East). 

Stigmodiplosis additionalis Fedotova & Sidorenko 2006c: 128. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; 

♂; ZMAS. 

*plicata. Russia (Far East). 

Stigmodiplosis plicata Fedotova & Sidorenko 2006c: 124. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

*uniseptata. Russia (Far East). 

Stigmodiplosis uniseptata Fedotova & Sidorenko 2006c: 127. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS.

STOMATOSEMA 

Stomatosema Kieffer 1904c: 380. Type species, nemorum Kieffer (mon). 

Jeannellomyia Kieffer 1913d: 40. Type species, gracilipes Kieffer (orig. des.). 

Baeomyza Kieffer 1913h: 89. Type species, monticola Kieffer (orig. des.). 

Vanchidiplosis Nayar 1949: 81. Type species, vanchii Nayar (orig. des.). 

Neohaplusia Rao 1951: 114. Type species, neohaplusia Rao (orig. des.) = vanchii Nayar. 

Haplusiella Rao 1952a: 49. Type species, pectiniclava Rao (orig. des.) = vanchii Nayar. 

Cosmopolitan; 20 spp.; Stomatosematidi. 

biclaspettii. India (Maharashtra). 

Stomatosema biclaspettii Kashyap 1989a: 22. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

brevicorne. Russia (Europe). 

Vanchidiplosis brevicornis Mamaev 1967: 878. Russia: Volodga Prov., Kadnikovskaya; HT; ♂; HYNM. 

*breviseta. Russia (Far East). 

Stomatosema breviseta Fedotova & Sidorenko 2006b: 22. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


ceylanicum. Sri Lanka. 

Didactylomyia ceylanica Felt 1915g: 173. Sri Lanka: Peradeniya; ST; ♂,♀; NYSM. 

*cuneata. Russia (Far East). 

Stomatosema cuneata Fedotova & Sidorenko 2006b: 24. Russia: Primorskiï Terr., Lazovskiï Res.; HT; ♂; ZMAS. 

dominicense. Dominica. 

Stomatosema dominicensis Gagné 1994a: 54. Dominica: 1.7 mi. E Pont Cassé; HT; ♂; USNM. 

gracilipes. Kenya. 

Jeannellomyia gracilipes Kieffer 1913d: 41. Kenya: Ramisi; HT; ♀; MNHN. 

kamali. India (Bihar), Germany. 

Vanchidiplosis kamali Grover 1961a: 172 (as n. sp. also in Grover 1964b: 191). India: Bihar, Mahijam; HT; ♀; 

PGCA. 

Vanchidiplosis kamalii Grover 1964b: 191, missp. of kamali. 

*medularis. Russia (Far East). 

Stomatosema medularis Fedotova & Sidorenko 2006b: 20, also as madularis, pp. 21-22. Russia: Primorskiï Terr., 

Lazovskiï Res., Proselochnaya Bay; HT; ♂; ZMAS. 

monticola. Tanzania. 

Baeomyza monticola Kieffer 1913h: 90. Tanzania: Tavéta; HT; ♀; MNHN. 

nemorum. France, Latvia, Lithuania, Poland, Russia (Europe). 

Stomatosema nemorum Kieffer 1904c: 380. France: vic. Bitche; ♀; lost?. A neotype designated by Mamaev (1967: 

877) is invalid because description inconsistent with original. 

obscurum. Lithuania, Latvia, Macedonia, Russia (Europe), USA (Minnesota, New Jersey), Mexico. 

Vanchidiplosis obscura Mamaev 1967: 877. Russia: Yaroslavl Prov., Volkovo; HT; ♂; HYNM. 

pedicellatum. Kenya. 

Baeomyza pedicellata Kieffer 1913h: 90. Kenya: Likoni, nr Mombasa; HT; ♀; MNHN. 

robustum. USA (New York). 

Didactylomyia robusta Felt 1920a: 282. USA: New York, Keene Valley; HT; ♂; NYSM. 

spinellosum. Italy. 

Stomatosema spinellosa Mamaev & Zaitzev 1998b: 130. Italy: Siena; HT; ♂; ZMMU. 

subsessile. Kenya. 

Baeomyza subsessilis Kieffer 1913h: 89. Kenya: Tiwi, S Mombasa; HT; ♀; MNHN. 

*taiga. Russia (Far East). 

Stomatosema taiga Fedotova 2003d: 9. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. Kamenushka; HT; ♂; 

ZMAS. 

trichodera. USA (Pennsylvania). 

387

388 

Stomatosema trichodera Gagné 1975a: 90. USA: Pennsylvania, York Co., Conewago Cr.; HT; ♂; USNM. 

vanchii. India (Uttar Pradesh, Kerala). 

Vanchidiplosis vanchii Nayar 1949: 82. India: Kerala, Trivandrum; ST; ♂,♀; ZSCI. 

Haplusia brevipalpia Rao 1951: 113. India: Uttar Pradesh, Agra, Hewett Park; HT; ♀; ZSCI. 

Neohaplusia agraensis Rao 1951: 116. India: Uttar Pradesh, Agra, Hewett Park; HT; ♀; ZSCI. 

Haplusia longipalpia Rao 1951: 112. India: Uttar Pradesh, Agra, Hewett Park; HT; ♀; ZSCI. 

Neohaplusia neohaplusia Rao 1951: 114. India: Uttar Pradesh, Agra, Hewett Park; HT; ♀; ZSCI. 

Haplusiella pectiniclava Rao 1952a: 49. India: Uttar Pradesh, Agra, St John's College; HT; ♂; ZSCI. 

Vanchidiplosis agraensis Rao 1952b: 31, jun. sec. hom. agraensis Rao 1951. India: Uttar Pradesh, Agra; HT; ♀; 

ZSCI. 

Haplusiella indica Rao 1953: 309. India: Uttar Pradesh, Agra; HT; ♂; ZSCI. 

Vanchidiplosis brevipalpi Rao 1960b: 189. India: Uttar Pradesh, Agra; HT; ♀; ZSCI. 

Haplusia elliptica Rao 1960b: 179. India: Uttar Pradesh, Agra, St John's College; HT; ♀; ZSCI. 

Neohaplusia inaequipalpis Rao 1960b: 181. India; HT; ♀; ZSCI. 

Vanchidiplosis longipalpi Rao 1960b: 191. India: Uttar Pradesh, Agra; HT; ♀; ZSCI. 

Vanchidiplosis similis Gagné 1973a: 516, new name for Vanchidiplosis agraensis Rao. 

*zaitzevi. Russia (Far East). 

Stomatosema zaitzevi Fedotova 2003d: 9. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. Kamenushka; HT; ♂; 

ZMAS. 

STREPTODIPLOSIS 

Streptodiplosis Felt 1916g: 405. Type species, indica Felt (orig. des.). 


indica. India. Poss. Lepidosaphes piperis (Hemiptera: Diaspididae). 

Streptodiplosis indica Felt 1916g: 405. India: South India, N. Wynaad; HT; ♂; lost. 

STROBLOPHILA 

Stroblophila Kieffer 1912h: 1. Type species, Clinodiplosis aberrans Kieffer (subs. des. Kieffer 1912: 9). 


aberrans. Austria. 

Clinodiplosis aberrans Kieffer 1901c: 34. Austria: Styria; ST; ♂,♀; lost?. 

STYRAXDIPLOSIS 

Styraxdiplosis Tavares 1915b: 147. Type species, caetitensis Tavares (orig. des.). 

Styracodiplosis Tavares 1918c: 77, unjust. emend. of Styraxdiplosis. 


caetitensis. Brazil (Bahia). Styrax sp. (Styracaceae). 

Styraxdiplosis caetitensis Tavares 1915b: 147. Brazil: Caetité, nr Salvador; ST; ♂; TAVC. 

Styraxdiplosis caetetensis Tavares 1918c: 75, missp. of caetitensis. 

cearensis. Brazil (Ceará). Croton hemiargyreus (Euphorbiaceae). 

Styraxdiplosis cearensis Tavares 1925a: 20. Brazil: Ceará; ST; ♀,P; TAVC. 

SYRENOMYIA

Syrenomyia Fedotova 1991d: 17. Type species, ramaea Fedotova (orig. des.). 


ramaea. Kazakhstan. Syrenia seliculosa (Brassicaceae). 

Syrenomyia ramaea Fedotova 1991d: 17. Kazakhstan: Buzachi Peninsula, 12 km SW Shebir; HT; ♂; ZMAS. 

TANAODIPLOSIS 

Tanaodiplosis Gagné 1973c: 880. Type species, Dicrodiplosis androgynes Felt (orig. des.). 

Nearctic; 1 sp., a hermaphrodite; Cecidomyiidi, Lestodiplosini. 

androgynes. USA (New York). 

Dicrodiplosis androgynes Felt 1908b: 394. USA: New York, Albany; HT; ♂/♀; NYSM. 

TAVARESOMYIA 

Tavaresomyia Möhn 1961a: 9. Type species, Schizomyia mimosae Tavares (orig. des.). 


mimosae. Brazil (Ceará). Mimosa caesalpinifolia (Fabaceae). 

Schizomyia mimosae Tavares 1925a: 28. Brazil: Ceará; ST; ♂,♀; TAVC. 

TAVOLGOMYIA 

Tavolgomyia Fedotova 1991d: 19. Type species, Wachtliella karelini Fedotova (orig. des.). 


karelini. Russia (Europe), Kazakhstan. Spiraea hypericifolia (Rosaceae). 

Wachtliella karelini Fedotova 1982a: 47. Kazakhstan: Kapchagaïskoe on Ili R.; HT; ♂; ZMAS. 

TAXODIOMYIA 

Taxodiomyia Gagné 1968e: 271. Type species, Cecidomyia cupressianananssa Osten Sacken (orig. des.). 

Nearctic; 3 spp., from Taxodium; Cecidomyiidi, Cecidomyiini. 

cupressiananassa. Southeastern Nearctic, Idaho (Miocene). Taxodium distichum (Taxodiaceae). 

Cecidomyia cupressi ananassa Riley 1870: 244, invalid, a polynomial. 

Cecidomyia cupressiananassa (as cupressi-ananassa) Osten Sacken 1878: 3. USA: Tennessee, Savannah; LT (Gagné 

1968e: 273); ♂; USNM. 

Cecidomyia ananassae Riley & Howard 1891: 125, error for cupressiananassa. 

Cecidomyia ananassi Felt 1906a: 755, error for cupressiananassa. 

Thecodiplosis ananassa Felt 1918e: 125, error for cupressiananassa. 

Thecodiplosis clarkiensis Lewis, Heikes, & Lewis 1990: 1, nomen nudum. Based on galls found nr Clarkia, Idaho, in 

Miocene sediments, Latah Formation. 

cupressi. Southeastern Nearctic. Taxodium distichum (Taxodiaceae). 

Merulius cupressi Schweinitz 1822: 92. USA: Georgia; T; G; ?. 

389

Itonida anthici Felt 1913b: USA: Alabama, Barachias; LT (Gagné 1968e: 273); ♂; USNM. 

taxodii. Southeastern Nearctic. Taxodium distichum (Taxodiaceae). 

Itonida taxodii Felt 1911l: 473. USA: Florida; LT (Gagné 1968e:273); ♀; USNM. 

TAXOMYIA 

Taxomyia Rubsaamen 1912d: 216. Type species, Cecidomyia taxi Inchbald (mon.). 

Palearctic; 1 sp.; Lasiopteridi, unplaced to tribe. Ref.: Redfern 1975 (biol., anatomy). 

taxi. Widespread Europe. Taxus baccata (Taxaceae). 

Cecidomyia taxi Inchbald 1861: 76. UK; England, Yorkshire; ST; ♂,♀; ?. 

TESSARADIPLOSIS 

Tessaradiplosis Baylac 1988: 31. Type species, Cecidomyia entomophila Perris (orig. des.). 


entomophila. UK, France. Acarus sp. (Acarina: Acaridae). 

Cecidomyia entomophila Perris 1855: 274. France: Landes, vic. Monts de Marsan; LT (Baylac 1888: 34); ♂; MNHN. 

TETRADIPLOSIS 

Tetradiplosis Kieffer & Jörgensen 1910: 421. Type species, sexdentata Kieffer & Jörgensen (mon.). 


sexdentata. Argentina (Mendoza). Prosopis alpataco, P. campestris (Fabaceae). 

Tetradiplosis sexdentatus Kieffer & Jörgensen 1910: 421. Argentina: Mendoza; ST; ♀,P,L; lost. 

TETRASPHONDYLIA 

Tetrasphondylia Kieffer 1913g: 48. Type species, Polystepha terminaliae Tavares (orig. des.). 

Afrotropical; 1 sp.; Cecidomyiidi, Asphondyliini, Asphondyliina. 

terminaliae. Mozambique. Terminalia sericea (Combretaceae). 

Polystepha terminaliae Tavares 1908: 152. Mozambique: Lourenço Marques, Nkonono; ST; ♂,♀; TAVC. 

THAUMADIPLOSIS 

Thaumadiplosis Gagné 1973c: 881. Type species, magnicauda Gagné (orig. des.). 


magnicauda. USA (Maryland). 

Thaumadiplosis magnicauda Gagné 1973c: 881. USA: Maryland, Bethesda; HT; ♂; USNM. 

THECODIPLOSIS 

Thecodiplosis Kieffer 1895f: cxciv. Type species, Cecidomyia brachyntera Schwägrichen (orig. des.). 

390

Thecordiplosis Shinji 1944: 232, missp. of Thecodiplosis. 

Holarctic; 6 spp., on Pinus; Cecidomyiidi, Cecidomyiini. 

brachyntera. Widespread Europe. Pinus sylvestris (Pinaceae). 

Cecidomyia brachyntera Schwägrichen 1835: 163. Germany: Leipzig; ST; ♂,♀,P,L; ?. 

Thecodiplosis brachyptera Skuhravá 1986: 288, missp. of brachyntera. 

brachynteroides. USA (Maryland to Virginia). Pinus virginiana (Pinaceae). 

Cecidomyia brachynteroides Osten Sacken 1862: 198. USA: District of Columbia; T; LT; lost. 

Cecidomyia brachypteroides Felt 1915i: 75, missp. of brachynteroides. 

japonensis. China, Korea, Japan (Hokkaido to Okinawa). Pinus densiflora, P. thunbergii (Pinaceae). 

Thecodiplosis japonensis Uchida & Inouye 1955: 44. Japan: Kyushu; HT; ♂; HFES. 

Thecodiplosis pinicola Kim 1955: 231. Korea: Seoul, Zun-Ra-Nam-Do, Pusan; ST; ♂,♀,P,L,egg; ?. 

piniradiatae. USA (California). Pinus radiata; Pinus spp. (Pinaceae). 

Diplosis piniradiatae (as pini-radiatae) Snow & Mills 1900: 491. USA: California, Palo Alto, Stanford University; 

ST; ♂,♀,P,L,egg; CASC. 

piniresinosae. Canada (Ontario), USA (Minnesota, Wisconsin, New York). Pinus resinosa (Pinaceae). 

Thecodiplosis piniresinosae Kearby 1963: 414. USA: Wisconsin, Adams Co.; HT; ♂; IRCW. 

pinirigidae. USA (Maine). Pinus rigida (Pinaceae). 

Diplosis pinirigidae (as pini-rigidae) Packard 1878: 527. USA: Maine, Brunswick; ST; ♀,L; USNM. 

THORODIPLOSIS 

Thorodiplosis Felt 1921k: 89. Type species, impatientis Felt (orig. des.). 


impatientis. Indonesia (Java). Impatiens platypetala (Balsaminaceae). 

Thorodiplosis impatientis Felt 1921k: 89. Indonesia: Java, Tjibodas, Mount Gede; ST; ♂,♀; NYSM. 

THRIPSOBREMIA 

Thripsobremia Barnes 1930b: 331. Type species, liothripis Barnes (orig. des.). 

Nearctic, Neotropical; 3 spp., predators of thrips and psyllids; Cecidomyiidi, Lestodiplosini. 

liothripis. Trinidad. Liothrips urichi (Thysanoptera: Phlaeothripidae). 

Thripsobremia liothripis Barnes 1930b: 331. Trinidad; ST; ♂,♀; BMNH. 

quercifolia. USA (Virginia). 

Contarinia quercifolia Felt 1908b: 391. USA: Virginia; HT; ♂; USNM. 

thripivora. USA (Florida). Gynaikothrips ficorum (Thysanoptera: Phlaeothripidae); Trioza magnoliae (Hemiptera: 

Psyllidae). 

Thripsobremia thripivora Gagné 1993a: 346. USA: Florida, Palm Beach Co., West Palm Beach; HT; ♂; USNM. 

THURAUIA 

Thurauia Rübsaamen 1899a: 58. Type species, aquatica Rübsaamen (subs. des. Felt 1911: 50). 

Holarctic; 2 spp., from Carex; Cecidomyiidi, unplaced to tribe. 

aquatica. Netherlands, Germany, Czechoslovakia, Russia (Europe), Jugoslavia. Carex paradoxa (Cyperaceae). 

Thurauia aquatica Rübsaamen 1899a: 59. Germany: Berlin, vic. Paulsborn in Grünewald; ST; ♂,♀,P,L; ZMHB. 

391

uliginosa. Germany. Carex sp. (Cyperaceae). 

Thurauia uliginosa Rübsaamen 1899a: 66. Germany: Berlin, vic. Paulsborn in Grünewald; ST; LT; ZMHB. 

*TINGIDOLETES 

Tingidoletes Gagné 2008a: 43. Type species, praelonga Felt (orig. des.). 

Nearctic; 1 sp.; Cecidomyiidi, Lestodiplosini. 

*praelonga. USA (Florida). Pseudacysta perseae (Heteroptera: Tingidae) 

Tingidoletes praelonga Gagné 2008a: 47. USA: Florida, Homestead; HT; ♂; USNM. 

TOKIWADIPLOSIS 

Tokiwadiplosis Simbolon & Yukawa 1992: 610. Type species, matecola Simbolon & Yukawa (orig. des.). 


matecola. Japan (Kyushu). Lithocarpus edulis (Fagaceae). 

Tokiwadiplosis matecola Simbolon & Yukawa 1992: 612. Japan: Kyushu, Kagoshima City, Terayama; HT; ♂; 

KUIC. 

*TOLLEREADASTRA 

Tollereadastra Fedotova & Sidorenko 2009a: 17. Type species, verecunda Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 2 spp.; Cecidomyiidi, unplaced to tribe here. The gender of the name was unstated but was 

presumably meant to be feminine because the type species was given the feminine form. Fedotova and Sidorenko 

2009a propose the new tribe Tollereadastrini to include this genus, Palmatodiplosis, and Fimbriatodiplosis, but the 

three genera do not appear to form a natural group. The tribal name is not used here (see Appendix). 

*torta. Russia (Far East). 

Tollereadastra torta Fedotova & Sidorenko 2009a: 21. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya pad'; 


*verecunda. Russia (Far East). 

Tollereadastra verecunda Fedotova & Sidorenko 2009a: 18. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya 

pad'; HT; ♂; ZMAS. 

*TONSIDIPLOSIS 

Tonsidiplosis Fedotova & Sidorenko 2005: 226. Type species, incurva Fedotova & Sidorenko (orig. des.). 

Palearctic; 4 spp.; Cecidomyiidi, Lestodiplosini. This genus does not essentially differ from Lestodiplosis. 

*amplissima. Russia (Far East). 

Tonsidiplosis amplissima Fedotova & Sidorenko 2006c: 122. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 

*incurva. Russia (Far East). 

Tonsidiplosis incurva Fedotova & Sidorenko 2005: 227. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. 


*rostriformis. Russia (Far East). 

Tonsidiplosis rostriformis Fedotova & Sidorenko 2005: 227. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. 

392

393 


*tuberculata. Russia (Far East). 

Tonsidiplosis tuberculata Fedotova & Sidorenko 2005: 228. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. 


*TRANSVERSALIDIPLOSIS 

Transversalidiplosis Fedotova 2003e: 69. Type species, siderenkoi Fedotova (orig. des.). 


separated. 

*ollae. Russia (Far East). 

Transversalidiplosis ollae Fedotova 2003e: 69. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

TRIBREMIA 

Tribremia Kieffer 1912h: 1. Type species, Bremia brevitarsis Kieffer (orig. des.). 


brevitarsis. France. 

Bremia brevitarsis Kieffer 1904c: 398. France: vic. Bitche; T; ♂; lost?. 

TRICHOLABA 

Tricholaba Rübsaamen 1917: 60. Type species, trifolii Rübsaamen (subs. des. Rübsaamen & Hedicke 1938: 170). 

Western Palearctic; 5 spp., inquilines or possibly inquilines in galls of Dasineura spp. on Fabaceae; Cecidomyiidi, 

unplaced to tribe. 

lespedezicola. Russia (Far East). Lespedeza bicolor (Fabaceae). 

Tricholaba lespedezicola Kovalev 1972: 416. Russia: southern Sikhote-Alin Mts, Pidan Range, Mt Khualaza; HT; ♂; 

ZMAS. 

similis. UK, Germany, Russia (Europe, Western Siberia). Vicia sp. (Fabaceae) in galls of Dasineura viciae (Diptera: 


Tricholaba similis Rübsaamen 1917: 64. Germany; ST; ♀,L; ZMHB. 

trifolii. UK, Sweden, Netherlands, Germany, Latvia, Lithuania, Poland, Czechoslovakia, Austria, Romania, Russia 

(Europe, Western Siberia). Trifolium sp. in galls of Dasineura trifolii (Diptera: Cecidomyiidae); T. pratense, 

T. repens (Fabaceae). 

Tricholaba trifolii Rübsaamen 1917: 62. Germany; ST; ♂,♀,L; ZMHB. 

Tricholaba barnesi Milne 1960: 85. UK: 14 counties in England & Wales; ST; ♂,♀; BMNH. 

viciarum. Germany, Russia. Vicia cracca (Fabaceae). 

Tricholaba viciarum Stelter 1963: 109. Germany: Kr. Rostock, Gross Lüsewitz; HT; ♂; orig. as DEIC, now in 

SMTD. 

viciobia. Germany. Vicia sylvatica (Fabaceae). 

Tricholaba viciobia Buhr 1965: 1382, nomen nudum. 

Tricholaba viciobia Stelter 1988: 98. Germany: Mühlhausen, Stadtwald; HT; ♂; SMTD. 

TRICHOPERRISIA

Trichoperrisia Kieffer 1913c: 199. Type species, pipericola Kieffer (orig. des.). 

Oriental; 1 sp.; Lasiopteridi, unplaced to tribe. 

pipericola. Sri Lanka. Piper nigrum (Piperaceae). 

Trichoperrisia pipericola Kieffer 1913c: 199. Sri Lanka: Peradeniya; ST; ♂; lost?. 

TRICONTARINIA 

Tricontarinia Kieffer 1910a: 71. Type species, ciliatipennis Kieffer (mon.). 

Eastern Palearctic; 2 spp., predators of Diaspididae; Cecidomyiidi, unplaced to tribe. This genus appears to grade into 

Diadiplosis. 

ciliatipennis. Japan (Honshu). Pseudaulacaspis pentagona (Hemiptera: Diaspididae). 

Tricontarinia ciliatipennis Kieffer 1910a: 71. Japan: Honshu, Tokyo; ST; ♂,♀ (Kieffer 1910b: 129 limited sp. to the 

♂); lost?. 

japanica. Japan (Honshu). Pseudaulacaspis pentagona (Hemiptera: Diaspididae). 

Tricontarinia japanica (as "n. nov." but actually new sp.) Kieffer 1910b: 129. Japan: Honshu, Tokyo; T; ♀; lost?. 

Tricontarinia japonica Skuhravá 1986: 289, missp. of japanica. 

TRIGONODIPLOSIS 

Trigonodiplosis Rübsaamen 1917: 68. Type species, fraxini Rübsaamen (mon.). 


fraxini. Yugoslavia. Fraxinus ornis (Oleaceae). 

Trigonodiplosis fraxini Rübsaamen 1917: 68. Serbia and Montenegro: nr Hercegnovi; ST; ♂,♀,L; ZMHB. 

TRIGONOMYIA 

Trigonomyia Kolesik 1996a: 62. Type species, ananas Kolesik (orig. des.). 

Australasian; 3 spp., all from Olearia; Lasiopteridi, unplaced to tribe. 

ananas. Australia (South Australia). Olearia ramulosa (Asteraceae). 

Trigonomyia ananas Kolesik 1996a: 62. Australia: South Australia, Black Hill Conservation Pk, 13 km NE Adelaide; 


cristata. Australia (South Australia). Olearia axilaris (Asteraceae). 

Trigonomyia cristata Kolesik 1996a: 66. Australia: South Australia, Beachport; HT; ♂; SAMA. 

tulipa. Australia (South Australia). Olearia axilaris (Asteraceae). 

Trigonomyia tulipa Kolesik 1996a: 66. Australia: South Australia, Beachport; HT; ♂; SAMA. 

TRILOBIA 

Trilobia Del Guercio 1918a: 223. Type species, aphidisuga Del Guercio (orig. des. as n. g., n. sp.). 


aphidisuga. Austria, Italy. Aphis fabae; several other Aphididae (Hemiptera) (Harris 1973). 

Trilobia aphidisuga Del Guercio 1918a: 223. Italy; T; LT; lost. 

394

TRILOBOMYIA 

Trilobomyia Marikovskij 1956: 187. Type species, insolita Marikovskij (orig. des.). 


insolita. Russia (Europe). Padus racemosa (Rosaceae). 

Trilobomyia insolita Marikovskij 1956: 188. Russia: Leningrad Prov.; ST; ♂; ZMAS. 

TRILOBOPHORA 

Trilobophora Marikovskij 1953a: 128. Type species, nitrariae Marikovskij (orig. des.). 


nitrariae. Kazakhstan. Nitraria schoberi (Zygophyllaceae). 

Trilobophora nitrariae Marikovskij 1953a: 129. Kazakhstan: vic. Iliskii, mid Ili R.; ST; ♂,♀; ZKAS & ZMAS. 

TRIOMMATA 

Triommata Barnes 1931a: 205. Type species, coccotroctes Barnes (mon.). 

Afrotropical, Oriental; 2 spp., predators of Pseudococcidae; Cecidomyiidi, unplaced to tribe. This genus is related to 

Diadiplosis, q.v. for comments. Ref.: Harris 1968 (illus.). 

coccidivora. India (Bihar, Karnataka, Kerala), Ceylon. Pseudococcus sp.; P. corymbatus, Ferrisia virgata, 

Maconellicoccus hirsutus (Hemiptera: Pseudococcidae). 

Diadiplosis coccidivora Felt 1914b: 86. Sri Lanka: Peradeniya; ST; ♂,♀; NYSM. 

Diadiplosis indica Felt 1920b: 6. India: Bihar, Pusa; ST; ♂,♀; NYSM. 

Schizobremia malabarensis Felt 1927a: 1. India: Kerala, Taliparamba; ST; ♂,♀; NYSM. 

Schizobremia bangalorensis Rao 1952a: 54. India: Karnataka, Bangalore; HT; ♂; ZSCI. 

coccotroctes. Sierra Leone, Democratic Republic of Congo. Ferrisia virgata (Hemiptera: Pseudococcidae). 

Triommata coccotroctes Barnes 1931a: 206. Sierra Leone: Njala; ST; ♂,♀,L; BMNH. 

TRIOMMATOMYIA 

Triommatomyia Mamaev 1961b: 405. Type species, barnesi Mamaev (orig. des.). 

Triomkatomyia Fedotova & Kovalev 2001: 417, 571, missp. of Triommatomyia. 

Palearctic; 2 spp.; Cecidomyiidi, Lestodiplosini. 

barnesi. Russia (Europe). 

Triommatomyia barnesi Mamaev 1961b: 406. Russia: Krasnodar Terr., Novo-Prokhladnoye; HT; ♂; HYNM. 

papillata. Russia (Far East). 

Triommatomyia papillata Mamaev 1998d: 9. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMMU. 

TRISOPSIS 

Trisopsis Kieffer 1912d: 171. Type species, oleae Kieffer (orig. des.). 

Plagiodiplosis Kieffer 1913g: 55. Type species, Lestodiplosis nana Kieffer (orig. des.). 

395

396 

Cosmopolitan; 26 spp., presumably all predators although most were caught in flight; Cecidomyiidi, Lestodiplosini. 

This genus, containing diverse species, may not be monophyletic; some species may be referable to Adiplosis or 

Odontodiplosis. 

abdominalis. Netherlands, Poland, Yugoslavia, Russia (Europe, Western Siberia), Uzbekistan. 

Trisopsis abdominalis Mamaev 1961b: 407 Russia: Tula Prov., Tula For.; HT; ♂; HYNM. 

acicularis. Ukraine, Russia (Europe). 

Trisopsis acicularis Mamaev 1961b: 409. Russia: Moscow Prov., Glubokoye L.; HT; ♂; HYNM. 

alluaudi. Kenya. 

Trisopsis alluaudi Kieffer 1913h: 94. Kenya: Mt Kenya; HT; ♀; MNHN. 

*coroniformis. Russia (Far East). 

Trisopsis coroniformis Fedotova & Sidorenko 2006c: 119. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; 

ZMAS. 


Trisopsis deepica Grover 1962a: 143. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

globularis. Russia (Europe, Far East). 

Trisopsis globularis Mamaev 1961b: 410. Russia: Moscow Prov., Bykovo; HT; ♂; ZMMU. 

hibisci. USA (Louisiana). 

Trisopsis hibisci Felt 1935d: 76. USA: Louisiana, Tallulah; ST; ♂,♀; USNM. 

hyperici. UK, Spain. Geocrypta braueri Handlirsch (Diptera: Cecidomyiidae). 

Trisopsis hyperici Tavares 1919b: 85. Spain: Gallicia, Redondela (Pontevedra); T; ♀; TAVC. 

incisa. USA (Washington, New York, District of Columbia), Trinidad, Colombia, Hawaiian Is (T. oleae of Hardy), 

Greece, Israel. Orig. unkn; from laboratory colonies of Anopheles sp. (Culicidae), Perileucoptera coffeella 

(Lepidoptera: Lyonetiidae), Pedronia sp. (Hemiptera: Pseudococcidae), on shoots of Sorghum sp., on leaves 

of Cyrtandra sp. (Gesneriaceae), in fruiting bodies of fungi, and various insect larvae in flower heads of 

Chromolaena odorata (Asteraceae) (cf. to list under Lestodiplosis callipus). 

Cecidomyia incisus Felt 1907a: 43. USA: New York, Albany; HT; ♂; NYSM. 


Trisopsis indica Grover 1962a: 141. India: Maharashtra, Nagpur; HT; ♀; PGCA. 

karelini. Ukraine, Russia (Europe, Far East), Kazakhstan, Uzbekistan. 

Trisopsis karelini Marikovskij 1958b: 1851. Kazakhstan: Alma-Ata, Kirghizstan: Frunze; T; ♂; MARC. 

longitarsis. Russia (Europe). 

Trisopsis longitarsis Mamaev 1961b: 407. Russia: Krasnodar Terr., Ubinsk For.; HT; ♂; HYNM. 

macrolobis. Russia (Europe). 

Trisopsis macrolobis Mamaev 1961b: 411. Russia: Moscow Prov., Bykovo; HT; ♂; ZMMU. 

nana. Seychelles Is. 

Lestodiplosis nana Kieffer 1911: 325. Seychelles Is: Mahé, Cascade Estate; ST; ♂; BMNH. 

nandedensis. India (Maharashtra). 

Trisopsis nandedensis Deshpande 1985: 59. India: Maharashtra, Nanded, Itwara; HT; ♂; DESH. 

narendrii. India (Madhya Pradesh). 

Adiplosis narendrii Grover 1979c: 41. India: Madhya Pradesh, Indore; HT; ♂; PGCA. 

obscura. USA (New York). 

Arthrocnodax obscurus Felt 1908b: 404. USA: New York, Karner; HT; ♂; NYSM. 

oleae. South Africa, New Zealand, Colombia. 

Trisopsis oleae Kieffer 1912d: 171. South Africa: Wellington; ST; ♂,♀; lost?. 

photophila. India (Uttar Pradesh). 

Trisopsis photophila Grover & Bakhshi 1978: 68. India: Uttar Pradesh, Allahabad, Mumfordganj, HT; ♂; PGCA. 

punctiventris. Germany, Poland, Ukraine, Russia (Europe), Uzbekistan. 

Trisopsis punctiventris Mamaev 1961b: 410. Russia: Moscow Prov., Pavlovskaya Sloboda; HT; ♂; HYNM. 

quercina. USA (New York, Georgia). 

Cecidomyia quercina Felt 1907a: 41. USA: New York, Albany; HT; ♂; NYSM. 

spiniformis. India (Maharashtra).

397 

Adiplosis spiniformis Grover & Kashyap 1991: 86. India: Maharashtra, Parbhani; HT; ♂; PGCA. 

*torta. Russia (Far East). 

Trisopsis torta Fedotova & Sidorenko 2006c: 116. Russia: Primorskiï Terr., Kedrovaya Pad' Res.; HT; ♂; ZMAS. 

tyroglyphi. Russia (Europe), Israel. Dermatophagoides farinae (Acarina: Acaridae). 

Trisopsis tyroglyphi Barnes 1951b: 565. Russia: Leningrad; ST; ♂,♀; BMNH. 

vulgaris. Latvia, Ukraine, Russia (Europe). 

Trisopsis vulgaris Mamaev 1961b: 408. Russia: Moscow Prov., Glubokoye L.; HT; ♂; HYNM. 

walkeriana. Sri Lanka, Hawaii. Pseudococcus sp. (Hemiptera: Pseudococcidae) & Walkeriana sp. (Margarodidae). 

Arthrocnodax walkeriana Felt 1915g: 181. Sri Lanka: Peradeniya, Botanic Gardens; ST; ♂,♀; NYSM. The type 

series contains two species, both tentatively in Trisopsis, each with the eyes separated laterally. 

TRISSODIPLOSIS 

Trissodiplosis Kieffer 1913h: 101. Type species, longicornis Kieffer (orig. des.). 


longicornis. Kenya. 

Trissodiplosis longicornis Kieffer 1913h: 101. Kenya: Likoni, nr Mombasa; HT; ♀; MNHN. 

TROGODIPLOSIS 

Trogodiplosis Gagné 1985a: 121. Type species, flexuosa Gagné (orig. des.). 


flexuosa. USA (Ohio, Maryland). 

Trogodiplosis flexuosa Gagné 1985a: 122. USA: Ohio, Canal Fulton; HT; ♂; USNM. 

TROPIDIPLOSIS 

Tropidiplosis Gagné 1973c: 881. Type species, pectinata Gagné (orig. des.). 


pectinata. USA (Minnesota). 

Tropidiplosis pectinata Gagné 1973c: 882. USA: Minnesota, Detroit Lakes; HT; ♂; USNM. 

TROTTERIA 

Choristoneura Rübsaamen 1892d: 342, preocc. Lederer 1859. Type species, Cecidomyia obtusa Loew. 

Trotteria Kieffer 1902b: 561, new name for Choristoneura Rübsaamen. 

Holarctic, Neotropical, Afrotropical; 23 spp.; Lasiopteridi, Trotteriini. Most Trotteria species whose biology is known 

live in galls caused by various gall midges of the tribe Asphondyliini. At least T. obtusa (as T. scoparii Möhn) lives also 

in galls formed by another gall midge, a Contarinia. Möhn (1963a) noted a unique behavior in this genus, that the hind 

legs are carried outstretched and do not touch the ground when the fly is at rest. Fedotova (2007) described five species 

on superficial grounds and with poor, sketchy drawings, each from a different host plant, but it is not definitely known 

that a one species/one host situation is the reality in Trotteria. Refs.: Skuhravá 1989 (notes); Fedotova 2007 (notes, incl. 

a list of Trotteria spp., their plant hosts and gall-making associates). 

*asteralobiphila. Russia (Far East). Spiraea salicifolia (Rosaceae).

398 

Trotteria asteralobiphila Fedotova 2002c: 20. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*bochkarevi. Russia (Europe). Verbascum orientale (Scrophulariaceae). 

Trotteria bochkarevi Fedotova 2004b: 814. Russia: 30 km SE Samara, nr Ust-Kinel'skii; HT; ♂; ZMAS. 

caryae. USA (New York), Dominica. 

Choristoneura caryae Felt 1907a: 14. USA: New York, Albany; HT; ♂; NYSM. 

Trotteria argenti Felt 1908b: 335. USA: New York, Albany; HT; ♂; NYSM. 

Trotteria caudata Felt 1908b: 335. USA: New York, Albany; HT; ♂; NYSM. 

Trotteria metallica Felt 1908b: 335. USA: New York, Albany; HT; ♂; NYSM. 

Trotteria solidaginis Felt 1908b: 335. USA: New York, West Nyack; HT; ♀; NYSM. 

Trotteria squamosa Felt 1908b: 335. USA: New York, Albany; HT; ♂; NYSM. 

Trotteria subfuscata Felt 1908b: 335. USA: New York, Albany; HT; ♀; NYSM. 

Trotteria tarsata Felt 1908b: 335. USA: New York, Albany; HT; ♂; NYSM. 

*clematicola. Russia (Far East). Clematis brevicaudata (Rosaceae). 

Trotteria clematicola Fedotova 2002c: 18 Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*coronillacola. Russia (Europe). Coronilla varia (Fabaceae). 

Trotteria coronillacola Fedotova 2007: 457. Russia: Middle Volga River Area, Zhiguli Nature Res., nr Bakhilova 


*cytisiphila. Russia (Europe). Citisus ruthenicus (Fabaceae). 

Trotteria cytisiphila Fedotova 2007: 460. Russia: Middle Volga River Area, Zhiguli Nature Res., nr Bakhilova 

Polyana, 15 km SE Zol'noe; HT; ♂; ZMAS. 

galii. UK, Sweden, Netherlands, Denmark, Germany, Poland, Czechoslovakia. Galium sylvaticum, G. mollugo 

(Rubiaceae). 

Trotteria galii Rübsaamen 1912g: 376. Germany: Remagen; ST; ♀,L; ZMHB. 

*humulopsidis. Russia (Far East). Humulopsis scandens (Cannabaceae). 

Trotteria humulopsidis Fedotova 2002c: 14. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*jigulensis. Russia (Europe). Tripleurospermum perforatum (Asteraceae). 

Trotteria jigulensis Fedotova 2007: 464. Russia: Middle Volga River Area, Zhiguli Nature Res., nr Bakhilova 

Polyana, 15 km SE Zol'noe; HT; ♀; ZMAS. 

*kalininae. Russia (Europe). Stachys recta (Lamiaceae). 

Trotteria kalininae Fedotova 2007: 462. Russia: Middle Volga River Area, 30 km SE Samara, nr Ust-Kinel'skii; HT; 

♂; ZMAS. 


Trotteria karnerensis Felt 1908b: 335. USA: New York, Karner; HT; ♀; NYSM. 

lapalmae. El Salvador. Eupatorium sp. (Asteraceae). 

Trotteria lapalmae Möhn 1975: 68. El Salvador: Chalatenango, SE La Palma; HT; ♂; SMNS. 

ligustri. UK, Czechoslovakia. Ligustrum vulgare (Oleaceae). 

Trotteria ligustri Barnes 1954b: 282. UK: England, Norfolk, Drayton; HT; ♂; BMNH. 

lindneri. El Salvador. Indigofera suffruticosa (Fabaceae). 

Trotteria lindneri Möhn 1963a: 1. El Salvador: San Salvador; HT; ♂; SMNS. 

obtusa. Widespread Europe. Orig. unkn.; various genera of Fabaceae. 

Lasioptera obtusa Loew 1845b: 394. Germany: vic. Cassel; T; M; ZMHB?. 

Lasioptera sarothamni Kieffer 1890d: 136. France: vic. Bitche; ST; ♀,L; lost?. 

Trotteria coronillae Kieffer 1913g: 47. Italy: Vallombrosa; ST; LT; lost?. 

Trotteria dalmatica Rübsaamen 1916b: 561. Serbia and Montenegro: Hercegnovi; ST; ♂,♀,L; ZMHB. 

Trotteria lathyri Rübsaamen 1916b: 563. Germany: vic. Sinzig; ST; ♀,L; ZMHB. 

Trotteria genistae Möhn 1966: 16. Germany: Bubenreuth nr Erlangen; HT; LT; SMNS. 

Trotteria ononidis Möhn 1966: 19. Hungary: Makád; HT; LT; SMNS. 

Trotteria viciae Möhn 1966: 20. Czech Republic: D�evohostice; HT; LT; SMNS. 

Trotteria scoparii Möhn 1966: 14. Germany: Westerwald, Hundsangen; HT; LT; SMNS.

399 

*perkovskii. Russia (Europe). Nepeta pannonica (Lamiaceae). 

Trotteria perkovskii Fedotova 2007: 459. Russia: Middle Volga River Area, 30 km SE Samara, nr Kamennyi; HT; ♂; 

ZMAS. 

quadridentata. Brazil (Rio de Janeiro). Pouteria caimito var. laurifolia (Sapotaceae). 

Trotteria quadridentata Maia 2001b: 107. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

rivinae. Colombia. Rivina humilis (Phytolaccaceae). 

Trotteria rivinae Wünsch 1979: 143. Colombia: Magdalena, Bahia Concha; HT; ♀; SMNS. 

salvadorensis. El Salvador; Colombia. Boerhaavia erecta (Sterculiaceae); Ayenia pusilla (Nyctaginaceae). 

Trotteria salvadorensis Möhn 1963a: 3. El Salvador: San Vicente, km 81 nr Rio San Felipe; HT; ♀; SMNS. 

sesami. Tanzania. Sesamum angustifolium (Pedaliaceae). 

Trotteria sesami Barnes 1939b: 455. Tanzania: Nyakato, Bukoba; ST; ♂,♀; BMNH. 

umbelliferarum. UK, France, Germany, Czechoslovakia, Russia (Far East). Daucus carota; other genera of Apiaceae 

(Möhn 1966: 23). 

Trotteria umbelliferarum Kieffer 1902a: 305 (as n. sp. also in Kieffer 1913g: 47). France: Bitche; T; ♂; lost?. 

Trotteria inquilina Rübsaamen 1921: 51. Germany: Partenkirchen; ST; ♂,♀,L; ZMHB. 

Trotteria spinosa Kovalev 1967: 96. Russia: Primorskiï Terr., Chapigou, Shufan; HT; ♂; KOVC. Fedotova (2007) 

removes this species from synonymy. 

ussuriana. Russia (Far East). Cacalia hastata (Asteraceae). 

Trotteria ussuriana Kovalev 1967: 97. Russia: Primorskiï Terr., Chapigou, Shufan; HT; ♂; KOVC. 

*veronicastricola. Russia (Far East). Veronicastrum sibiricum (Scrophulariaceae). 

Trotteria veronicastricola Fedotova 2002c: 17. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*TUGURIDIPLOSIS 

Tuguridiplosis Fedotova 2005c: 216. Type species, phaseoliformis Fedotova (orig. des.). 

Palearctic; 10 spp.; Cecidomyiidi, Clinodiplosini. This genus is close to Clinodiplosis if the two must be separated. 

Compare the slight differences among species with the variation found for Clinodiplosis cilicrus in Skuhravá (1973). 

*abdita. Russia (Far East). 

Tuguridiplosis abdita Fedotova 2005c: 217. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. Kamenushka; HT; ♂; 

ZMAS. 

*aspectabilis. Russia (Far East). 

Tuguridiplosis aspectabilis Fedotova & Sidorenko 2009d: 233. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskya 


*brochata. Russia (Far East). 

Tuguridiplosis brochata Fedotova & Sidorenko 2009d: 236. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 


*flammosa. Russia (Far East). 

Tuguridiplosis flammosa Fedotova & Sidorenko 2009d: 230. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskya 


*pallida. Russia (Far East). 

Tuguridiplosis pallida Fedotova & Sidorenko 2009d: 239. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskya Pad'; 


*pergrata. Russia (Far East). 

Tuguridiplosis pergrata Fedotova & Sidorenko 2009d: 230. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya 


*phaseoliformis. Russia (Far East). 

Tuguridiplosis phaseoliformis Fedotova 2005c: 217. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, vic. 


*turris. Russia (Far East).

400 

Tuguridiplosis turris Fedotova & Sidorenko 2009d: 236. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskya Pad'; 


*vafra. Russia (Far East). 

Tuguridiplosis vafra Fedotova & Sidorenko 2009d: 219. Russia: Primorskiï Terr.: 30 km SE Ussuriysk, Kamenushka; 


*vespertina. Russia (Far East). 

Tuguridiplosis vespertina Fedotova & Sidorenko 2009d: 233. Russia: Primorskiï Terr., Lazovskiï Res., Cordon 


ULEELLA 

Uleella Rübsaamen 1908a: 120. Type species, dalbergiae Rübsaamen (orig. des.). 

Uleela Felt 1912i: 353, missp. of Uleella. 


dalbergiae. Brazil (Rio de Janeiro). Dalbergia sp. (Fabaceae). 

Uleella dalbergiae Rübsaamen 1908a: 121. Brazil: Rio de Janeiro, Jacarapaguá; ST; P,L; ZMHB. 

ULEIA 

Uleia Rübsaamen 1905a: 85. Type species, clusiae Rübsaamen (mon.). 

Ulea Houard 1933: 249, missp. of Uleia. 


clusiae. Brazil (Amazonas). Clusia sp. (Clusiaceae). 

Uleia clusiae Rübsaamen 1905a: 85. Brazil: Amazonas, Santa Clara & Bom Fim on Juruá R.; ST; ♂,♀,L; ZMHB. 

UNCINULELLA 

Uncinulella Del Guercio 1918a: 227. Type species, eriosomiperda Del Guercio (orig. des. as n. g. n. sp.). 

Western Palearctic; 1 sp.; Cecidomyiidi, Lestodiplosini. The larval head and terminal segments as originally 

illustrated resemble those of a Feltiella, but the aphid host would be unlikely for a Feltiella. 

eriosomiperda. Italy. Eriosoma lanigerum (Hemiptera: Aphididae). 

Uncinulella eriosomiperda Del Guercio 1918a: 227. Italy; T; LT; lost. 

*UNDADIPLOSIS 

Undadiplosis Fedotova 2004g: 17. Type species, tomentosa Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, Karshomyiini. Fedotova 2009 erected the subtribe Undadiplosini, in which 

were also included Opinatodiplosis and Laxadiplosis, but these three genera do not appear to form a natural group (see 

Appendix). 

*tomentosa. Russia (Far East). 

Undadiplosis tomentosa Fedotova 2004g: 20. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

*UNDONEURA

Undoneura Fedotova & Sidorenko 2006d: 94. Type species, fasciculata Fedotova & Sidorenko (orig. des.). 

Eastern Palearctic; 2 spp.; Brachineuridi. This genus is close to Brachineura if the two must be separated. 

*fasciculata. Russia (Far East). 

Undoneura fasciculata Fedotova & Sidorenko 2006d: 94. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; 


*phaseoliformis. Russia (Far East). 

Brachineura phaseoliformis Fedotova 2004i: 598. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

*USSURIDIPLOSIS 

Ussuriplosis Fedotova 2004g: 12. Type species, Lobodiplosis pallida Fedotova (orig. des.). 

Eastern Palearctic; 1 sp.; Cecidomyiidi, unplaced to tribe. This genus is close to Karshomyia if the two must be 

separated. 


Lobodiplosis pallida Kovalev & Mamaev 1966: 231. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; 

HT; ♂; KOVC. 

VACCINIDIPLOSIS 

Vaccinidiplosis Gagné 1994c: 601. Type species, Cecidomyia vaccinii Osten Sacken (orig. des.). 


vaccinii. USA (New York, Pennsylvania, Maine, Maryland, District of Columbia, Georgia). Vaccinium stamineum 

(Ericaceae). 

Cecidomyia vaccinii Osten Sacken 1862: 196. USA: District of Columbia; ST; LT; lost. 

Cecidomyia gaylussacii Felt 1925c: 55, unnec. new name for Cecidomyia vaccinii Osten Sacken. 

*VERBASCIOLA 

Verbasciola Fedotova 2004b: 811. Type species, mamaevae Fedotova (orig. des.). 

Palearctic; 3 species; Lasiopteridi, Camptoneuromyiini. Fedotova 2004b erected the tribe Verbasciolini for this genus. 

This is unnecessary inasmuch as Verbasciola and Camptoneuromyia can hardly be separated. 

*mamaevae. Russia (Europe). Verbascum orientale; V. nigrum (Scrophulariaceae). 

Verbasciola mamaevae Fedotova 2004b: 812. Russia: 30 km SE Samara, nr Ust-Kinel'skii; HT; ♂; ZMAS. 

petioli. Czechoslovakia, Russia (Europe). Populus tremula (Salicaceae). 

Camptoneuromyia petioli Mamaeva 1964a: 207. Russia: Moscow Prov., Pavlovskaja Sloboda; HT; ♀; IEME. 

*volgensis. Russia (Europe). Veronica chamaedris (Scrophulariaceae). 

Verbasciola volgensis Fedotova 2007: 469. Russia: Middle Volga River Area, Zhiguli Nature Res., nr Bakhilovo; 


*VERSUSFACERA 

Versusvacera Fedotova & Sidorenko 2009c: 74. Type species, crepera Fedotova & Sidorenko (orig. des.). 

401

Eastern Palearctic; 2 spp.; Cecidomyiidi, Mycodiplosini here, but Fedotova & Sidorenko 2009c place it in their tribe 

Coquillettomyiini (see Appendix). The genus is evidently close to Coquillettomyia. 

*crepera. Russia (Far East). 

Versusfacera crepera Fedotova & Sidorenko 2009c: 75. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya pad'; 


*dolosa. Russia (Far East). 

Versusfacera dolosa Fedotova & Sidorenko 2009c: 75. Russia: Primorskiï Terr., Lazovskiï Res., Proselochnaya Bay; 


*VITISIELLA 

Vitisiella Fedotova & Kovalev 2003c: 25. Type species, vesicula Fedotova (orig. des.). 

Palearctic; 3 spp.; Lasiopteridi, Dasineurini. Known species form leaf galls on grape. Ref.: Gagné 2009b: 404 

(redescr.). 

brevicauda. Canada (Ontario), USA (Missouri, New York, New Hampshire, Maryland, North Carolina). Vitis sp. 

(Vitaceae). 

[Cecidomyia] vitis tomatos Riley 1873: 117, invalid, a polynomial. 

Janetiella brevicauda Felt 1908b: 372. USA: unspecified; ST; ♂,♀; NYSM. Ref.: Gagné 2009b: 405 (redescr.). 

oenephila. Widespread central to southern Europe; Turkey, Lebanon; Russia (Far East). Vitis vinifera (Vitaceae). 

Cecidomyia oenephila Haimhoffen 1875: 809. Austria: Gersthof, nr Vienna; ST; ♀,L; ?. 

Janetiella oenophila Kieffer 1913i: 59, missp. of oenephila. 

*vesicula. Russia (Far East). on Vitis amurensis (Vitaceae). 

Vitisiella vesicula Fedotova & Kovalev. 2003c: 26. Russia: Primorskiï Terr., 30 mi SE Ussuriysk, Kamenushka; HT; 

♂; ZMAS. 

*VOLSATIOLA 

Volsatiola Fedotova & Sidorenko 2006b: 16. Type species, aequa Fedotova & Sidorenko (orig. des.). 


*aequa. Russia (Far East). 

Volsatiola aequa Fedotova & Sidorenko 2006b: 16. Russia: Primorskiï Terr., Lazovskiï Res., Koreiskaya Pad'; HT; 

♂; ZMAS. 

WACHTLIELLA 

Wachtliella Rübsaamen 1916b: 491. Type species, dalmatica Rübsaamen (orig. des.). 

Palearctic; 9 spp.; Lasiopteridi, Dasineurini. Sylvén & Tastás-Duque (1993) indicate that this genus is differentiated 

from Dasineura on slight differences. No distinction between these two genera has been made for the Nearctic fauna, 

for which Wachtliella has not been used. 

caricis. Widespread Europe; Algeria. Carex muricata; C. riparia (Cyperaceae). 

Cecidomyia caricis Loew 1850a: 30. Poland; T; G; ZMHB?. 

Cecidomyia riparia Winnertz 1853: 218. Germany: Aachen; ST; ♂,♀; lost. 

Cecidomyia muricatae Meade 1886: 153. UK: England, nr Harrogate; ST; ♂,♀; ?. 

*cytisicola. Russia. Cytisus ruthenicus (Fabaceae). 

402

403 

Wachtliella cytisicola Fedotova 2008: 131. Russia: Central Povolzhye, Zhigulevsk Nature Res., nr Bakhilova 


dalmatica. Czechoslovakia, Croatia, Ukraine, Russia (Western Siberia). Medicago falcata, M. prostrata (Fabaceae). 

Wachtliella dalmatica Rübsaamen 1916b: 544. Croatia: betw. Trogir & Split; ST; ♂,♀,L; ZMHB. 

ericina. Widespread Europe, Algeria, Tunisia. Erica carnea (Ericaceae). 

Cecidomyia ericina Löw 1885b: 484. Austria, Croatia, Italy; ST; ♂,♀,P,L; NHMW. 

*galatellaflorae. Kazakhstan. Galatella hauptii; G. trinervifolia (Asteraceae). 

Wachtliella galatellaflorae Fedotova 2003a: 829. Kazakhstan: Azutau Mtn Range, 25 km SE Alekseevka; HT; ♂; 

ZMAS. 

krumbholzi. Germany, Czechoslovakia. Rhamnus catharticus (Rhamnaceae). 

Wachtliella krumbholzi Stelter 1975a: 158. Germany: Kr. Rostock, Gross Lüsewitz; HT; ♂; SMTD. 

niebleri. Germany, Poland, Czechoslovakia, Italy, Hungary, Romania, Greece, Ukraine, Russia (Western Siberia). 

Cytisus nigricans (Fabaceae). 

Wachtliella niebleri Rübsaamen 1916b: 543. Germany: nr Amberg; ST; ♂,♀,L; ZMHB. 

persicariae. Widespread Europe, Kazakhstan. Polygonum amphibium; Polygonum spp. (Polygonaceae). 

Tipula persicariae Linnaeus 1767: 977. Sweden; T; A; LSUK?. 

stachydis. Widespread Europe. Stachys sylvatica; S. recta (Lamiaceae). 

Cecidomyia stachydis Bremi 1847a: 55. Switzerland: Burghölzli nr Zurich; ST; A,L; ETHZ? 

Perrisia granulata Kieffer 1909b: 29. Germany: Kreuznach, & Hungary: Budapest; ST; LT; lost?. 

WALSHOMYIA 

Walshomyia Felt 1908b: 359. Type species, juniperina Felt (orig. des.). 

Nearctic; 5 spp., all on Cupressaceae; Lasiopteridi, Oligotrophini. 

cupressi. USA (California). Cupressus pygmaea (Cupressaceae). 

Walshomyia cupressi Gagné 1969a: 19. USA: California, Mendocino Co., Ft. Bragg; HT; ♂; USNM. 

insignis. USA (Texas). Juniperus sp. 

Walshomyia insignis Felt 1921b: 117. USA: Texas, Austin; ST; ♀,P; NYSM. 

juniperina. USA (California). Juniperus californica (Cupressaceae). 

Walshomyia juniperina Felt 1908b: 360. USA: California, New Indria; ST; ♂,♀; NYSM. 

sabinae. (Utah, Colorado, Texas). Juniperus "sabinoides" (Cupressaceae). 

Rhopalomyia sabinae Patterson 1919: 346. USA: Texas, T; G; UTAT?. 

Rhopalomyia sabinae Felt 1921b: 115. USA: Colorado, Denver; ST; ♂,♀,P; NYSM. 

texana. USA (Texas). Juniperus "sabinoides" (Cupressaceae). 

Walshomyia texana Felt 1916a: 30. USA: Texas, Austin; ST; ♂,♀; NYSM. 

XENASPHONDYLIA 

Xenasphondylia Felt 1915a: 202. Type species, albipes Felt (orig. des.). 

Xenasphohdylia Shinji 1944: 227, missp. of Xenasphondylia. 

Neotropical; 1 sp.; Cecidomyiidi, unplaced to tribe. Ref.: Gagné 1994a (illus.). 

albipes. Dominican Republic. 

Xenasphondylia albipes Felt 1915a: 202. Dominican Republic: San Francisco Mts; ST; ♀; USNM. 

XENHORMOMYIA 

Xenhormomyia Felt 1920c: 491. Type species, africana Felt (orig. des.).


africana. South Africa. 

Xenhormomyia africana Felt 1920c: 492. South Africa: Pietermaritzburg; HT; ♂; UNSA. 

natalensis. South Africa. 

Xenhormomyia natalensis Felt 1920c: 493. South Africa: Pietermaritzburg; HT; ♀; UNSA. 

XENODIPLOSIS 

Allodiplosis Rübsaamen 1910d: 287, preocc. Kieffer & Jörgensen 1910. Type species, laeviusculi Rübsaamen (orig. 

des. as n. g., n. sp.). 

Xenodiplosis Felt 1911c: 61, new name for Allodiplosis Rübsaamen. 


laeviusculi. UK, Germany, Czechoslovakia. Quercus robur (Fagaceae). 

Allodiplosis laeviusculi Rübsaamen 1910d: 287. Germany: vic. Berlin; T; ♂; ZMHB. 

*lonicerae. Russia (Far East). Lonicera maackii (Caprifoliaceae). 

Xenodiplosis lonicerae Fedotova 2004e: 19. Russia: Primorskiï Terr., 30 km SE Ussuriysk, Kamenushka; HT; ♂; 

ZMAS. 

XEREPHEDROMYIA 

Xerephedromyia Mamaev 1972a: 893. Type species, bipartita Mamaev (orig. des.). 

Palearctic; 3 spp., all from Ephedra; Lasiopteridi, Dasineurini. 

bipartita. Uzbekistan. Ephedra strobilacea (Gnetaceae). 

Xerephedromyia bipartita Mamaev 1972a: 893. Uzbekistan: Bukhara Prov., Kul'dzhuktau; HT; ♂; HYNM. 

mitroshinae. Turkmenistan. Ephedra strobilacea; E. lomatolepis (Gnetaceae). 

Xerephedromyia mitroshinae Fedotova 1992a: 76. Turkmenistan: southern Karakumy Desert, 70 km SW Chardzhou, 

Repetek Stn; HT; ♂; ZMAS. 

ustjurtensis. Spain, Kazakhstan. Ephedra distachya (Gnetaceae). 

Xerephedromyia ustjurtensis Fedotova 1992a: 78. Kazakhstan: eastern Mangyschlak basin, 80 km NE Novyï-Uzen', 

45 km NE Senek; HT; ♂; ZMAS. 

XIPHOLASIOPTERA 

Xipholasioptera Gagné 1995c: 776. Type species, ensata Gagné (orig. des.). 


ensata. USA (Texas). Baccharis pteronioides (Asteraceae). 

Xipholasioptera ensata Gagné 1995c: 776. USA: Texas, Guadelupe Mts Natl Pk; HT; ♀; USNM. 

XYLODIPLOSIS 

Xylodiplosis Kieffer 1895a: cclxxx. Type species, Cecidomyia praecox Winnertz (orig. des.) = nigritarsis Zetterstedt. 

Holarctic, Oriental; 5 spp., including at least 3 spp. that live in xylem vessels of fresh-cut wood; Cecidomyiidi, 


404

aestivalis. France. 

Xylodiplosis aestivalis Kieffer 1904c: 382. France: vic. Bitche; ST; ♂,♀; lost?. 

kempi. India (Karnataka). 

Xylodiplosis kempi Mani 1934b: 439. India: Karnataka, North Kanara, Castle Rock; HT; ♀; ZSCI. 

kizilkumensis. Uzbekistan. 

Xylodiplosis kizilkumensis Mamaev 1998d: 8. Uzbekistan: Kyzyl Kum, Kuldzhuktau Mts; HT; ♂; ZMMU. 

longistylus. USA (Ohio, Maryland). 

Xylodiplosis longistylus Gagné 1985a: 119. USA: Ohio, Canal Fulton; HT; ♂; USNM. 

nigritarsis. UK, Denmark, Netherlands, France, Germany, Finland. 

Cecidomyza nigritarsis Zetterstedt 1850: 3683. Denmark; ST; ♀; MZLU?. 

Cecidomyia praecox Winnertz 1853: 276. Germany; ST; ♀; lost. 

XYLOPERRISIA 

Xyloperrisia Kieffer 1913g: 68. Type species, Perrisia azarae Kieffer (mon.). 

Neotropical; 2 spp.; Lasiopteridi, unplaced to tribe. 

azarae. Chile. Pernettya furens (Ericaceae). 

Perrisia azarae Kieffer 1903: 227. Chile: vic. Concepción; ST; ♂,P,L; lost. 

Perrisia azarai Kieffer & Herbst 1905: 64, unjust. emend. of azarae. 

mayteni. Argentina (Neuquén). Maytenus boaria (Celastraceae). 

Xyloperrisia mayteni Gagné 2002b: 39. Argentina: Neuquén, San Martín de los Andes; HT; ♂; USNM. 

YOUNGOMYIA 

Youngomyia Felt 1908b: 398. Type species, Dicrodiplosis podophyllae Felt (orig. des.). 

Delphodiplosis Felt 1915h: 155. Type species, cinctipes Felt (orig. des.). 

Nearctic, Neotropical, Oriental; 6 spp., inquilines, possibly predaceous, in galls of other cecidomyiines; Cecidomyiidi, 


cinctipes. Guyana. 

Delphodiplosis cinctipes Felt 1915h: 155. Guyana: Bartica; HT; ♀; NYSM. 

knabi. Costa Rica. 

Coquillettomyia knabi Felt 1912h: 154. Costa Rica: Port Limón; ST; ♂,♀; USNM. 

podophyllae. USA (New York, Maine, Pennsylvania, Virginia), Guatemala, Panama. Orig. unk.; inquiline in galls of 

various Cecidomyiinae (cf. Gagné 1989b). 

Dicrodiplosis podophyllae Felt 1907a: 30. USA: New York, Albany; HT; ♂; NYSM. 

Dicrodiplosis podophylli Felt 1907c: 126, unjust. emend. of podophyllae. 

Youngomyia rubida Felt 1908b: 399. USA: New York, Nassau; HT; ♂; NYSM. 

Youngomyia vernoniae Felt 1911l: 474, nomen nudum. 

Youngomyia vernoniae Felt 1911n: 552. USA: Virginia; ST; ♂,♀; USNM. 

Youngomyia pennsylvanica Felt 1912f: 106. USA: Pennsylvania, Hazelton; HT; ♂; NYSM. 

Youngomyia producta Felt 1918e: 150. USA: New York, Karner; HT; ♀; NYSM. 

pouteriae. Brazil (Rio de Janeiro). Pouteria caimito var. laurifolia (Sapotaceae). 

Youngomyia pouteriae Maia 2001b: 104. Brazil: Rio de Janeiro, Maricá; HT; ♂; MNRJ. 

quercina. USA (California). Quercus sp. (Fagaceae). 

Youngomyia quercina Felt 1911l: 467, nomen nudum. 

Youngomyia quercina Felt 1911n: 551. USA: California, Donner; HT; ♂; USNM. 

spinosa. India (Uttar Pradesh). 

405

Youngomyia spinosa Grover 1979c: 159. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

ZALEPIDOTA 

Zalepidota Rübsaamen 1908b: 38. Type species, piperis Rübsaamen (orig. des.). 

Ozobia Kieffer 1913i: 97. Type species, tavaresi Kieffer (orig. des.). 

Oxasphondylia Felt 1915a: 204. Type species, reticulata Felt (orig. des.). 

Neotropical; 4 spp.; Cecidomyiidi, Asphondyliini, Asphondyliina. Ref.: Möhn 1962 (illus.). 

ituensis. Brazil (São Paulo). Porophyllum sp. (Asteraceae). 

Oxasphondylia ituensis Tavares 1917b: 139. Brazil: São Paulo, Salto de Itu; ST; ♀,P; TAVC. 

piperis. Brazil (Rio de Janeiro). Piper sp. (Piperaceae). 

Zalepidota piperis Rübsaamen 1908a: 121. Brazil: Rio de Janeiro, Tijuca; LT (Möhn 1963b: 15); ♂; ZMHB. 

reticulata. Guatemala. 

Oxasphondylia reticulata Felt 1915a: 205. Guatemala: Trece Aguas, Alta V. Paz; HT; ♀; USNM. 

tavaresi. Brazil (Rio Grande do Sul). Piper sp. (Piperaceae). 

Ozobia tavaresi Kieffer 1913i: 98. Brazil: Rio Grande do Sul, vic. São Leopoldo; LT (Möhn 1963b: 20); P; TAVC. 

ZEUXIDIPLOSIS 

Zeuxidiplosis Kieffer 1904b: 349. Type species, Diplosis giardiana Kieffer (orig. des) = Diplosis giardi Kieffer. 

Palearctic, Afrotropical, introd. western Nearctic and Australasian; 1 sp.; Cecidomyiidi, Cecidomyiini. 

giardi. Widespread Europe; introd: USA (California), Australia, New Zealand; immigr.: South Africa. Hypericum 

perforatum (Hypericaceae). 

Diplosis giardi Kieffer 1896j: 383. France: château de La Grange, nr Thionville; ST; ♂,♀; lost?. 

Diplosis giardiana Kieffer 1897a: 20, unjust. new name for Diplosis giardi Kieffer. 

ZYGIOBIA 

Zygiobia Kieffer 1913h: 55. Type species, Cecidomyia carpini Loew (orig. des.). 

Zygobia Mani 1946: 207, missp. of Zygiobia. 

Western Palearctic; 2 spp., from Carpinus leaf galls; Lasiopteridi, Dasineurini. 

carpini. Widespread Europe. Carpinus betulus (Betulaceae). 

Cecidomyia carpini Löw 1874b: 322. Austria: Pfalzauthale, nr Vienna; ST; ♂,♀,P,L; NHMW. Stelter (1992c: 379) 

designated a neotype, which was invalid inasmuch as there was no indication that a search was made for the 

types, which may be in NHMW. 

ruebsaameni. Germany. Carpinus betulus (Betulaceae). 

Zygiobia ruebsaameni Stelter 1992c: 374. Germany: Kr. Rostock, Tessin; HT; ♂; DEIC. 

UNPLACED SPECIES OF CECIDOMYIIDAE 

Most of the species listed here may be referable to a known genus once available types are studied or the 

species are reared again. This list contains some junior primary homonyms that are not renamed because of the 

likelihood they will be considered synonymous with the senior homonym. 

Lestremiinae 

406

sydneyensis. Australia: (New South Wales). 

Lestremia sydneyensis Skuse 1890: 389. Australia: New South Wales, Sydney; HT; ♀; ANIC. 

Micromyinae 

* † admirandus. Rovno amber (Ukraine). 

Aprionus admirandus Fedotova 2004a: 46. Ukrainian amber: Rovno; HT; ♀; UASK. Ref.: Jaschhof 2007a: 105. 

aeratipennis. Australia (New South Wales). 

Campylomyza aeratipennis Skuse 1888: 135. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC. 

amplipennis. Australia (New South Wales). 

Campylomyza amplipennis Skuse 1888: 142. Australia: New South Wales, Middle Harbour; HT; ♂; ANIC. 

* † appropinquata. Rovno amber (Ukraine). 

Strobliella appropinquata Fedotova 2004d: 69. Ukrainian amber: Rovno; HT; ♀; UASK. Ref.: Jaschhof 2007a: 105. 

* † autonoma. Rovno amber (Ukraine). 

Peromyia autonoma Fedotova 2004a: 51. Ukrainian amber: Rovno; HT; ♂; UASK. Ref.: Jaschhof 2007a: 105. 

* † capitata. Rovno amber (Ukraine). 

Strobliella capitata Fedotova 2004d: 69. Ukrainian amber: Rovno; HT; ♀; UASK. Ref.: Jaschhof 2007a: 105. 

* † convoluta. Rovno amber (Ukraine). 

Micromyia convoluta Fedotova 2004a: 45. Ukrainian amber: Rovno; HT; ♀; UASK. Ref.: Jaschhof 2007a: 105. 

crocea. Australia (New South Wales). 

Campylomyza crocea Skuse 1888: 136. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC. 

* † declinata. Rovno amber (Ukraine). 

Neurolyga declinata Fedotova 2004d: 75. Ukrainian amber: Rovno; HT; ♂; UASK. Ref.: Jaschhof 2007a: 105. 

* † falciformis. Rovno amber (Ukraine). 

Campylomyza falciformis Fedotova 2004d: 72. Ukrainian amber: Rovno; HT; ♀; UASK. Ref.: Jaschhof 2007a: 105. 

grandiuscula. Australia (New South Wales). 

Campylomyza grandiuscula Skuse 1890: 388. Australia: New South Wales, Sydney; HT; ♂; ANIC. 

impexa. Australia (New South Wales). 

Campylomyza impexa Skuse 1888: 139. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

* † laesus. Rovno amber (Ukraine). 

Conarete laesus Fedotova 2004a: 53. Ukrainian amber: Rovno; HT; ♀; UASK. Ref.: Jaschhof 2007a: 105. 

* † miranda. Rovno amber (Ukraine). 

Peromyia miranda Fedotova 2004a: 50. Ukrainian amber: Rovno; HT; ♂; UASK. Ref.: Jaschhof 2007a: 105. 

pellax. Australia (New South Wales). 

Campylomyza pellax Skuse 1888: 140. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

perpallida. Australia (New South Wales). 

Campylomyza perpallida Skuse 1888: 134. Australia: New South Wales, Sydney; HT; ♀; ANIC. 

persimilis. Australia (New South Wales). 

Campylomyza persimilis Skuse 1888: 136. Australia: New South Wales, Sydney; HT; ♀; ANIC. 

sidneyensis. Australia (New South Wales). 

Campylomyza sidneyensis Schiner 1868: 9. Australia: New South Wales, Sydney; T; A; NHMW?. 

Campylomyza sydneyensis Skuse 1888: 141, missp. of sidneyensis. 

* † sparsa. Rovno amber (Ukraine). 

Heterogenella sparsa Fedotova 2004a: 46. Ukrainian amber: Rovno; HT; ♂; UASK. Ref.: Jaschhof 2007a: 105. 

* † superposita. Rovno amber (Ukraine). 

Campylomyza superposita Fedotova 2004d: 71. Ukrainian amber: Rovno; HT; ♀; UASK. Ref.: Jaschhof 2007a: 105. 

subtilis. Australia (New South Wales). 

Campylomyza subtilis Skuse 1888: 137. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

* † sukachevae. Rovno amber (Ukraine). 

Peromyia sukachevae Fedotova 2004a: 49. Ukrainian amber: Rovno; HT; ♂; UASK. Ref.: Jaschhof 2007a: 105. 

vicina. Australia (New South Wales). 

Campylomyza vicina Skuse 1888: 138. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

407

Porricondylinae 

* † bifurcata. Late Eocene - Ukrainian amber. 

Diadocidia bifurcata Fedotova 2004d: 76. Rovno amber, Ukraine; HT; ♀; UASK. 

distenta. Australia (New South Wales). 

Cecidomyia distenta Skuse 1888: 115. Australia: NSW, Elizabeth Bay; HT; ♀; ANIC. 

exigua. Australia (New South Wales). 

Cecidomyia exigua Skuse 1888: 122. Australia: New South Wales, Glenbrook, Blue Mts; HT; ♂ (orig. mistakenly as 

♀); ANIC. 

indubitata. Australia (New South Wales). 

Cecidomyia indubitata Skuse 1888: 114. Australia: New South Wales, Middle Harbour; HT; ♂; ANIC. 

leucopeza. Germany. 

Cecidomyia leucopeza Meigen 1830: 268. Germany; T; ♀; MNHN. 

longiptera. India (West Bengal). 

Clinophaena longiptera Nayar 1949: 82. India: West Bengal, Calcutta; ST; ♂,♀; ZSCI. 

magnifica. Australia (New South Wales). 

Cecidomyia magnifica Skuse 1888: 116. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

opipara. Australia (New South Wales). 

Cecidomyia opipara Skuse 1888: 118. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC. 

pallidina. Australia (New South Wales). 

Cecidomyia pallida Skuse 1888: 120, jun. prim. hom. pallida Meigen. Australia: New South Wales, Elizabeth Bay; 

ST; ♂,♀; ANIC. 

Porricondyla pallidina Gagné 1989c: 156, new name for Cecidomyia pallida Skuse. 

parkensis. India (Uttar Pradesh). 

Clinophena parkensis Grover 1964b: 195. India: Uttar Pradesh, Allahabad; HT; ♀; PGCA. 

perexilis. Australia (New South Wales). 

Cecidomyia perexilis Skuse 1888: 119. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

skusei. Australia (New South Wales). 

Cecidomyia gracilis Skuse 1888: 117, jun. prim. hom. gracilis Winnertz. Australia: New South Wales, Elizabeth 

Bay; HT; ♂; ANIC. 

Porricondyla skusei Kieffer 1913i: 273, new name for Cecidomyia gracilis Skuse. 

Porricondyla exilis Gagné 1989c: 156, unnec. new name for Cecidomyia gracilis Skuse. 

† spectabilis. fossil - Baltic amber. 

Cecidomyia spectabilis Meunier 1899a: 161, with reference to Meunier 1899b, pl. I, fig. 1. Baltic amber; T; ?; 

Königsburg collection. The type has recently been found, so this species should be placed here instead of with 

nomina dubia, per M. Jaschhof (in litt.). 

temeritatis. USA (New York). 

Colpodia temeritatis Felt 1908b: 416. USA: New York, vic. Albany; HT; ♀; NYSM. 

Cecidomyiinae 

acanthopanaci. Japan (Honshu). Eleutherococcus sp. (Araliaceae). 

Poricondylia acanthopanaci Shinji 1942a: 161. Japan: Honshu; T; ♀; lost. 

Poricondylia acanthopanici Shinji 1944: 316, missp.of acanthopanaci. 

actiosa. Australia (New South Wales). 

Cecidomyia actiosa Skuse 1888: 105. Australia: New South Wales, Peat's Ferry; ST; ♂,♀; ANIC. 

aequipalpis. India (Uttar Pradesh). 

Meinertomyia aequipalpis Mani 1935: 426. India: Uttar Pradesh, Dehra Dun, Jhajra; ST; ♀; ZSCI. 

aethiops. UK. 

Cecidomyia aethiops Walker 1856a: 118. UK: England; ST; ♂; BMNH. 

agraensis. India (Uttar Pradesh). 

408

409 

Feltiella agraensis Rao 1957: 8. India: Uttar Pradesh, Agra; HT; ♂; SJCA. 

agrostis. USA (New York). Agrostis lateriflora (Poaceae). 

Cecidomyia agrostis Osten Sacken 1862: 204. USA: New York; T; P; lost. 

albipes. UK. 

Cecidomyia albipes Walker 1856a: 100. UK: England; ST; ♂; BMNH. 

albonotata. UK. 

Cecidomyia albonotata Walker 1856a: 118. UK: England; ST; ♂; BMNH. 


Blastodiplosis allahabadensis Grover & Bakhshi 1978: 111. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

antidesme. Philippines. Antidesma moritzii (Euphorbiaceae). 

Ctenodactylomyia antidesme Felt 1919a: 287. Philippines: Luzon, Laguna Prov., Mt Makiling; HT; ♀; NYSM. 

apicalis. UK. 

Cecidomyia apicalis Walker 1856a: 107. UK: England; ST; ♀; BMNH. 

araneosa. Australia (New South Wales). 

Cecidomyia araneosa Skuse 1888: 82. Australia: New South Wales, Elizabeth Bay; HT; ♀ (orig. as ♂); ANIC. 

articulata. UK. 

Cecidomyia articulata Walker 1856a: 108. UK: England; ST; ♂; BMNH. 

artocarpi. India (Maharashtra). Artocarpus sp. (Moraceae). 

Cecidomyia artocarpi Felt 1921j: 27. India: Maharashtra, Godavari; ST; ♀; NYSM. 

astilbensis. Japan (Honshu). Astilbe thunbergii (Saxifragaceae). 

Schizoneura astilbensis Shinji 1939d: 68. Japan: Honshu, Morioka; ST; ♂,♀; lost. 

atricapilla. Italy. Aristolochia rotunda (Aristolochiaceae). 

Cecidomyia atricapilla Rondani 1869: 190. Italy; ST; ♀; MZUF. 

atriceps. UK. 

Cecidomyia atriceps Walker 1856a: 102. UK: England; ST; ♂; BMNH. 

aurantiaca. France. 

Cecidomyia aurantiaca Macquart 1826: 172. France; HT; A; MNHN?. 

baccata. Australia (New South Wales). 

Cecidomyia baccata Skuse 1888: 65. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

barnaii. India (Uttar Pradesh). Crateva religiosa (Capparidaceae). 

Aschistonyx barnaii (also as baranii) Grover 1965a: 33. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

basalis. UK. 

Cecidomyia basalis Walker 1856a: 88. UK: England; ST; ♀; BMNH. 

bassiaflorae. India (Tamil Nadu). Bassia latifolia (Chenopodiaceae). 

Mikiola bassiaflorae Mani 1986: 15. India: Tamil Nadu, Madras; HT; ♀; ZSCI. 


Orseoliella bengalensis Mani 1937: 285. India: West Bengal, Calcutta, Portland Park; HT; ♂; ZSCI. 


Stefaniola bengalensis Mani 1934b: 401. India: West Bengal, Port Canning; HT; ♀; ZSCI. 

bicolora. Switzerland. 

Cecidomyia bicolor Bremi 1847a: 54, jun. prim. hom. bicolor Meigen. Switzerland: vic. Zurich; ST; ♂,♀; ETHZ. 

Cecidomyia bicolora Kieffer 1913i: 214, new name for Cecidomyia bicolor Bremi. 

biconica. Cuba. 

Cecidomyia biconica (as bi-conica) Cook 1906: 251. Cuba: Prov. Santiago de Cuba; T; G; ?. 

bidenticulata. Malaysia. 

Cecidomyia bidenticulata Barnes 1927e: 273. Malaysia: Cameron Highlands, Pahang; HT; ♂; BMNH. 

bifurcata. Samoa Is. 

Liebeliola bifurcata Barnes 1928e: 106. Samoa Is: Upolu, Malololelei; HT; ♀; BMNH. 

bispina. India (Maharashtra). 

Octodiplosis bispina Sharma 1987a: 2. India: Maharashtra, Aurangabad, Khadkeshwar; HT; ♂; INPC. 

boerhaaviaefoliae. India (Uttar Pradesh). Boerhaavia repanda (Nyctaginaceae). 

Cecidomyia boerhaaviaefoliae Mani 1944: 161. India: Uttar Pradesh, Delhi; HT; ♀; INPC.

410 

bremii. Miocene - Switzerland. Populus sp. (Salicaceae). 

Cecidomyia bremii Heer 1876: 54. Miocene/Switzerland: Oeningen; ST; G; ?. 

brevicornis. UK. 

Cecidomyia brevicornis Walker 1856a: 99. UK: England; ST; ♂; BMNH. 

brevipalpis. India (Karnataka). 

Blastodiplosis brevipalpis Sharma 1999: 169 India: Karnataka, Nagarhole Natl Pk; HT; ♂; ZCSI. 

brevipennis. Australia (New South Wales). 

Cecidomyia brevipennis Skuse 1888: 71. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

brevis. UK. 

Cecidomyia brevis Walker 1856a: 93. UK: England; ST; ♂; BMNH. 

brideliae. Philippines. Bridelia stipularis (Euphorbiaceae). 

Toxomyia brideliae Felt 1920d: 233. Philippines: Luzon, Laguna Prov., Los Baños; ST; ♀; NYSM. 

calcuttaensis. India (West Bengal). 

Hormomyia calcuttaensis Mani 1937: 284. India: West Bengal, Calcutta, Portland Park; HT; ♀; ZSCI. 

californica. USA (California). Ribes sp. (Grossulariaceae). 

Mayetiola californica Felt 1908b: 370. USA: California, Alameda Co.; ST; ♂,♀; NYSM?. 

californica. USA (California). 

Rhabdophaga californica Felt 1908b: 353. USA: California, Santa Clara Co.; HT; ♂; NYSM. 

callicarpae. Japan (Honshu, Kyushu). Callicarpa japonica (Verbenaceae). 

Rhopalomyia callicarpae Shinji 1939b: 563. Japan: Honshu; ST; ♂,♀; lost. 

capensis. South Africa. Phylica ericoides (Rhamnaceae). 

Cecidomyia capensis Schiner 1868: 7. South Africa: Cape of Good Hope; HT; ♀; NHMW?. 

cardamomi. India (Kerala). Elettaria cardamomi (Zingiberaceae). 

Hallomyia cardamomi Nayar 1949: 84. India: Kerala, high ranges of Travancore, Pampadamparai Hills; ST; ♀; 

ZSCI. 

carnea. Germany. 

Cecidomyia carnea Meigen 1818: 98. Germany; T; ♂; MNHN. 

carpophaga. Canada (Ontario). Picea glauca (Pinaceae). 

Phytophaga carpophaga Tripp 1955: 261. Canada: Ontario, Dunrobin; HT; ♂; CNCI. 

cayamacensis. Cuba. Prob. a sp. of Fabaceae. 

Cecidomyia cayamacensis Cook 1906: 251. Cuba: Prov. Santa Clara, Cayamas; T; G; ?. 

cecropiae. Cuba. Cecropia obtusa (Moraceae). 

Cecidomyia cecropiae Cook 1909: 145. Cuba; T; G; ?. 

chilkaensis. India (Tamil Nadu). 

Thurauia chilkaensis Mani 1934b: 416. India: Tamil Nadu, Chilka L., Barkuda I.; HT; ♀; ZSCI. 

chinquapin. USA (North Carolina). Castanea pumila (Fagaceae). 

Cecidomyia chinquapin Beutenmüller 1907a: 389. USA: North Carolina, Buncombe Co., Valley of the Black Mts; 

HT; LT; AMNH. 

ciliata. UK. 

Cecidomyia ciliata Walker 1856a: 105. UK: England; ST; ♀; BMNH. 

cinctipes. UK. 

Cecidomyia cinctipes Walker 1856a: 90. UK: England; HT; ♂; BMNH. 

clausilia. Switzerland, Spain. 

Cecidomyia clausilia Bremi 1847a: 28. Switzerland: Canton Zurich, nr Dättlikon; ST; LT; ETHZ?. Gall on Salix alba 

(Salicaceae). Skuhravá et al. 2006: 135 point out that similar galls on Salix are forned by Eriophyidae 

(Acarina), but the larva originally described (only as yellow) by Bremi would still be the name bearer. This 

species would probably be an inquiline or predator, so is removed from Rabdophaga where it was placed in 

Gagné 2004. 

Cecidomyia inchbaldiana Mik 1886: 317, unjust. new name for Cecidomyia clausilia Bremi. 

clematidis. Japan (Honshu). Clematis apiifolia (Rosaceae). 

Mycetodiplosis clematidis (also as cleamatidis) Shinji 1944: 393. Japan: Honshu; T; ♀; lost. 

Silvestris clematii Shinji 1944: 146, error for Mycetodiplosis clematidis.

411 

concinna. UK. 

Cecidomyia elegans Walker 1856a: 99, jun. prim. hom. for elegans Walker. UK: England; ST; ♂; BMNH. 

Cecidomyia concinna Walker 1856a: 342, new name for Cecidomyia elegans Walker. 

concolor. UK. 

Cecidomyia concolor Walker 1856a: 98 (also as discalis p. 242, as unnec. new name for Cecidomyia concolor 

Walker). UK: England; ST; ♂; BMNH. 

condigna. Australia (New South Wales). 

Cecidomyia condigna Skuse 1888: 85. Australia: New South Wales, Elizabeth Bay; ST; ♂; ANIC. 

conformis. UK. 

Cecidomyia conformis Walker 1856a: 112. UK: England; ST; ♀; BMNH. 

coprosmae. New Zealand. Coprosma crassifolia (Rubiaceae). 

Kiefferia coprosmae Barnes & Lamb 1954: 814. New Zealand, Auckland, Bethell's Beach; HT; ♂; BMNH. 

coprosmae. New Zealand. Coprosma arborea (Rubiaceae). 

Oligotrophus coprosmae Barnes & Lamb 1954: 815. New Zealand, Auckland, Titirangi; HT; ♂; BMNH. 

costalis. UK. 

Cecidomyia costalis Walker 1856a: 83. UK: England; ST; ♂; BMNH. 

costata. Sri Lanka. 

Lowiola costata Felt 1915g: 182. Sri Lanka: Peradeniya, Botanic Gardens; ST; ♀; NYSM. 

crassicornis. UK. 

Cecidomyia crassicornis Walker 1856a: 119. UK: England; ST; ♂; BMNH. 

crataevae. India (Tamil Nadu). Crateva religiosa (Capparidaceae). 

Cecidomyiella crataevae Mani 1934b: 428. India: Tamil Nadu, Thanjavur; ST; ♂,♀; ZSCI. 

cylindriformis. India (Jammu and Kashmir). 

Dicrodiplosis cylindriformis Kashyap 1988c: 112. India: Jammu and Kashmir, Srinagar; HT; ♂; PGCA. 

danae. UK. 

Cecidomyia danae Haliday 1856: 131. UK: England; ST; ♀; NMID. 

dattai. India (West Bengal). Aegle marmelos (Rutaceae). 

Cecidomyia dattai Mani 1937: 285. India: West Bengal, Calcutta, nr Barrackpore; HT; ♀; ZSCI. 

decolorata. UK. 

Cecidomyia decolorata Walker 1856: 87. UK: England; ST; ♀; BMNH. 

decora. UK. 

Cecidomyia decora Walker 1856a: 97. UK: England; ST; ♂; BMNH. 


Bryocrypta deepica Rao 1960b: 184. India: Uttar Pradesh, Agra, Hewett Park; HT; ♀; ZSCI. 

deferenda. Malaysia (Sarawak). 

Cecidomyia deferenda Walker 1856b: 105. Malaysia: Sarawak; HT; ♀; BMNH. 

deformans. South Africa. Osteospermum moniliferum (Asteraceae). 

Cecidomyia deformans Schiner 1868: 8. South Africa: Cape of Good Hope, betw. Simonsbay & Cape Town; HT; ♂; 

NHMW?. 

dehradunensis. India (Uttar Pradesh). 

Liebeliola dehradunensis Rao 1960b: 186. India: Uttar Pradesh, Dehra Dun; HT; ♀; ZSCI. 

dibapha. Australia (New South Wales). 

Cecidomyia dibapha Skuse 1888: 98. Australia: New South Wales, Illawarra Distr., Coogee Bay; ST; ♀; ANIC. 

difficilis. Australia (New South Wales). 

Cecidomyia difficilis Skuse 1888: 63. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC. 

dioscoreae. Japan (Honshu). Dioscorea nipponica (Dioscoreaceae). 

Riveraella dioscoreae Shinji 1939e: 98. Japan: Honshu, Morioka; T; A; lost. 

discolor. UK. 

Cecidomyia discolor Walker 1856a: 87. UK: England; ST; ♂; BMNH. 

disjuncta. UK. 

Cecidomyia disjuncta Walker 1856a: 104. UK: England; ST; ♂; BMNH. 

diurni. India (Uttar Pradesh).

Liebeliola diurni Rao 1960b: 187. India: Uttar Pradesh, Agra, St. John's College; HT; ♀; ZSCI. 

dolosa. UK. 

Cecidomyia dolosa Walker 1856a: 114. UK: England; ST; ♀; BMNH. 

4dubia. Tanzania. 

Lamprodiplosis dubius Kieffer 1913h: 95. Tanzania; HT; ♀; MNHN. 

eppoi. Japan (Honshu). Juniperus chinensis var. globosa (Cupressaceae). 

Aschistonyx eppoi Inouye 1964: 26. Japan: Honshu, Okayama Pref.; HT; ♂; HFES. 

erronea. Australia (New South Wales). 

Cecidomyia erronea Skuse 1888: 94. Australia: New South Wales, Sydney; HT; ♀; ANIC. 

eucalypti. Australia (New South Wales). Eucalyptus haemastoma (Myrtaceae). 

Cecidomyia eucalypti Skuse 1890: 381. Australia: New South Wales, Botany Bay; HT; ♀; lost. 

eugeniae. Chile. Myrceugenia stenophylla (Myrtaceae). 

Oligotrophus eugeniae Kieffer & Herbst 1909: 125. Chile: Concepción; T; LT; lost. 

evanescens. UK. 

Cecidomyia evanescens Walker 1856a: 102. UK: England; ST; ♂; BMNH. 

expandens. UK. 

Cecidomyia expandens Walker 1856a: 101. UK: England; ST; ♂; BMNH. 

fici. Cuba. Ficus sp. (Moraceae). 

Cecidomyia fici Cook 1909: 145. Cuba; T; G; ?. 

filipes. UK. 

Cecidomyia filipes Walker 1856a: 103. UK: England; ST; ♀; BMNH. 

filipes. Seychelles Is. 

Lepidodiplosis filipes Kieffer 1911: 321. Seychelles Is: Silhouette I.; HT; ♀; BMNH. 

finalis. UK. 

Cecidomyia terminalis Walker 1856a: 100, jun. prim. hom. terminalis Loew. UK: England; ST; ♂; BMNH. 

Cecidomyia finalis Walker 1856a: 342, new name for Cecidomyia terminalis Walker. 

fixa. UK. 

Cecidomyia fixa Walker 1856a: 107. UK: England; ST; ♂; BMNH. 

fleothripetiperda. Italy. Liothrips oleae (Thysanoptera: Phlaeothripidae). 

Diplosis fleothripetiperda Del Guercio 1931: 133. Italy; ST; A,P,L; lost. 

flexa. UK. 

Cecidomyia arcuata Walker 1856a: 112, jun. prim. hom. arcuata Winnertz. UK: England; T; ♀; BMNH. 

Cecidomyia flexa Walker 1856a: 342, new name for Cecidomyia arcuata Walker. 

floridensis. USA (Florida). 

Phytophaga floridensis Felt 1921c: 143. USA: Florida, St. Augustine; HT; ♂; NYSM. 

frauenfeldi. Australia (New South Wales). Melaleuca sp. (Myrtaceae). 

Cecidomyia frauenfeldi Schiner 1868: 7. Australia: New South Wales, Sydney; ST; ♂,♀; NHMW?. 

frequens. Australia (New South Wales). 

Cecidomyia frequens Skuse 1888: 99. Australia: New South Wales, Elizabeth Bay; ST; ♂,♀; ANIC. 

furcata. Chile. 

Lasioptera furcata Philippi 1865: 631. Chile: Santiago; T; A; MNNC?. 

fuscescens. Chile. 

Cecidomyia fuscescens Philippi 1865: 628. Chile: Santiago; T; A; MNNC?. 

gagaimo. Japan (Honshu). Metaplexis japonica (Asclepiadaceae). 

Psectrosema gagaimo Monzen 1955b: 39. Japan: Iwate & Morioka Prefs; ST; ♂,♀,L; KUEC. 

Psectrosema gagaima Skuhravá 1986: 295, missp. of gagaimo. 

garcinia. Indonesia (Java). Garcinia dioica (Clusiaceae). 

Gnesiodiplosis garcinia Felt 1927d: 388. Indonesia: Java, Depok, nr Djakarta; ST; ♂,♀,P; NYSM. 

gemini. Switzerland. Hieracium pilosella (Asteraceae). 

Cecidomyia gemini Bremi 1847a: 17. Switzerland: Katzensee; ST; LT; lost. 

globata. India (Uttar Pradesh). 

Therodiplosis globata Grover 1965a: 28. India: Uttar Pradesh, Allahabad, Tagore Town; HT; ♂; PGCA. 

412

413 

gnaphalii. Germany. Gnaphalium leontopodium (Asteraceae). 

Oligotrophus gnaphalii Kieffer 1909b: 9. Germany; T; LT; ZMHB?. 

gollmeri. Venezuela. 

Cecidomyia gollmeri Karsch 1880: 297. Venezuela: Caracas. HT; ♂; ZMHB. 

gracilis. Tanzania. 

Coprodiplosis gracilis Kieffer 1913h: 97. Tanzania: Taveta; HT; ♀; MNHN. 

grossulariae. USA (Wyoming, New York). Ribes grossularia (Saxifragaceae). 

Cecidomyia grossulariae Fitch 1855: 880. USA: New York, Salem; ST; ♀,L; USNM. 

grossulariae. USA (Ohio). Ribes grossularia (Grossulariaceae). 

Rhopalomyia grossulariae Felt 1911h: 347. USA: Ohio, Camp Chase; ST; ♂,♀; NYSM. 

hageni. USA (Massachusetts). Aristolochia macrophylla (Aristolochiaceae). 

Cecidomyia hageni Aldrich 1905: 162. USA: Massachusetts; T; G; MCZC?. 

hirta. Kenya. 

Coprodiplosis hirtus Kieffer 1913h: 96. Kenya: Voi; HT; ♀; MNHN. 

hirta. Sri Lanka. Dactylopius sp. (Hemiptera: Dactylopiidae). 

Endaphis hirta Felt 1911g: 224. Sri Lanka: Tangalla; HT; ♂; USNM. 

ilexifoliae. Japan (Honshu). Ilex serrata (Aquifoliaceae), Symplocos chinensis (Symplocaceae). 

Rhopalomyia ilexifoliae (also as ilicifoliae & ilecifoliae) Shinji 1944: 358. Japan: Honshu; T; ♀; lost. 

inaequipalpis. India (Uttar Pradesh, Assam). 

Meinertomyia inaequipalpis Mani 1935: 426. India: Assam, Srimangal, Sylhet, & UP, Kotdwara, Lansdowne; ST; ♀; 

ZSCI. 

incompleta. UK. 

Cecidomyia incompleta Walker 1856a: 101. UK: England; ST; ♂; BMNH. 

indianus. India (West Bengal). Capparis viminea (Capparidaceae). 

Oligotrophus indianus Kieffer 1909a: 152. India: West Bengal, Kurseong; T; LT; lost?. 

Oligotrophus indicus Kieffer 1913i: 50, unnec. new name for Oligotrophus indianus Kieffer; no Oligotrophus 

indianus Felt exists. 


Clinophaena indica Mani 1934b: 385. India: West Bengal, Kurseong; HT; ♂; ZSCI. 


Johnsonomyia indica Rao 1960b: 177. India: Uttar Pradesh, Agra; HT; ♀; ZSCI. 


Toxomyia indica Grover 1979c: 153. India: West Bengal, Darjeeling; HT; ♂; PGCA. 


Triommata indica Grover 1965a: 21, jun. sec. hom. Diadiplosis indica Felt. India: Uttar Pradesh, Allahabad; HT; ♀; 

PGCA. 

Triommata cybele Gagné 1973a: 511, new name for Triommata indica Grover. 

innotata. UK. 

Cecidomyia innotata Walker 1856a: 82. UK: England; ST; A; BMNH. 

innotata. UK. 

Cecidomyia innotata Walker 1856a: 117. UK: England; ST; ♂; BMNH. 

invaria. UK. 

Cecidomyia invaria Walker 1856a: 102. England; ST; ♂; BMNH. 

itaparicana. Brazil (Bahia). Undet. plant. 

Compsodiplosis itaparicana Tavares 1922a: 41. Brazil: Bahia, Salvador, Itaparica; HT; ♂; TAVC. 

jamboolii. India (Uttar Pradesh). Eugenia jambolana (Myrtaceae). 

Frauenfeldiella jamboolii Grover 1969b: 156. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

japonicus. Japan (Honshu). Weigela hortensis (Caprifoliaceae). 

Oligotrophus japonicus Monzen 1955a: 46. Japan: Honshu, Aomori, Iwate, Tochigi, Niigata, Wakayama, & Tottori 

Prefs; ST; ♀; KUEC. 

jujubae. India (Uttar Pradesh). Zizyphus jujuba (Rhamnaceae). 

Phyllodiplosis jujubae Grover & Bakhshi 1978: 155. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA.

414 

jujubae. India (Tripura). Zizyphus jujuba (Rhamnaceae). 

Silvestrina jujubae Chandra 1988: 164. India: Tripura, Agartala; HT; ♂; CSIA. 

kaalae. Hawaiian Is. 

Mayetiola kaalae Hardy 1960: 276. Hawaiian Is: Hawaii, Oahu, Mt Kaala; HT; ♂; HDOA. 

labiatidis. India (Uttar Pradesh). Terammus labiatis (Fabaceae). 

Tricholaba labiatidis Grover & Bakhshi 1978: 105. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

latipennis. UK. 

Cecidomyia latipennis Walker 1856a: 99. UK: England; ST; ♂; BMNH. 

latiuscula. UK. 

Cecidomyia latiuscula Walker 1856a: 108. UK: England; ST; ♀; BMNH. 

leptospermi. Australia (New South Wales). Leptospermum sp. (Myrtaceae). 

Cecidomyia leptospermi Skuse 1888: 68. Australia: New South Wales, Sydney; HT; ♂; ANIC. 

limbitorquens. Switzerland. 

Cecidomyia limbitorquens Bremi 1847a: 48. Switzerland: vic. Zurich; ST; ♂; ETHZ. 

linearis. UK. 

Cecidomyia linearis Walker 1856a: 112. UK: England; ST; ♂,♀; BMNH. 

lippiae. Argentina (Mendoza). Lippia foliolosa (Verbenaceae). 

Rhopalomyia lippiae Kieffer & Jörgensen 1910: 401. Argentina: Mendoza, Cordillera de Mendoza; ST; ♂,♀,P,egg; 

lost. 

loewianus. UK, France, Austria. Carex arenaria (Cyperaceae). 

Oligotrophus loewianus (as löwianus) Kieffer 1909b: 4. Austria; T; LT; NHMW?. 

longicauda. Tanzania. 

Coprodiplosis longicauda Kieffer 1913h: 99. Tanzania: Taveta; HT; ♀; MNHN. 

longicornis. Sweden. 

Tipula longicornis Linnaeus 1758: 588. Sweden; T; A; LSUK?. 

longipennis. India (Karnataka). 

Blastodiplosis longipennis Sharma 1999: 167. India: Karnataka, Nagarhole Natl Pk; HT; ♂; ZCSI. 

lonicera. USA (California). Lonicera subspicata (Caprifoliaceae). 

Rhopalomyia lonicera Felt 1925a: 15. USA: California, Claremont; HT; ♂; NYSM?. 

lutea. Germany. 

Cecidomyia lutea Meigen 1804: 40. Germany; ST; ♂,♀; MNHN. 

lyciicola. Argentina (Mendoza). Lycium chilense; L. gracile (Solanaceae). 

Oligotrophus lyciicola Kieffer & Jörgensen 1910: 409, 414. Argentina: Mendoza, Cordillera de Mendoza; T; G; lost. 

lythri. Poland. Lythrum salicaria (Lythraceae). 

Cecidomyia lythri Loew 1950: 30. Poland; ST; G; ZMHB?. 

macleayi. Australia (New South Wales). 

Cecidomyia macleayi Skuse 1888: 97. Australia: New South Wales, Woronora; HT; ♀; ANIC. 

malabarensis. India (Kerala). Piper nigrum (Piperaceae) 

Cecidomyia malabarensis Felt 1927a: 3. India: Kerala, Taliparamba; HT; ♀; NYSM. 

mangiferae. India (West Bengal). Mangifera indica (Anacardiaceae). 

Oligotrophus mangiferae Kieffer 1909a: 150. India: West Bengal, nr Ranchi; HT; P; lost?. The damage attributed to 

this species is the same as that of Procontarinia mangiferae Felt, but Kieffer described his own species as 

having frontal armature, which Felt's species does not. Possibly Kieffer meant to write "cervical armature" 

instead, because no other known species from mango has frontal prominences of the type Kieffer described. 

manii. India (Uttar Pradesh). 

Meinertomyia manii Rao 1951: 110. India: Uttar Pradesh, Agra; HT; ♀; ZSCI. 

marginata. UK. 

Cecidomyia marginata Walker 1856a: 107. UK: England; ST; ♂; BMNH. 

maura. UK. 

Cecidomyia maura Walker 1856a: 102. UK: England; ST; ♂; BMNH. 

mazaiana. Cuba. 

Cecidomyia mazaiana Cook 1906: 252. Cuba: Prov. Santiago de Cuba; T; G; ?.

415 

minima. Austria. 

Asphondylia minima Strobl 1880: 42. Austria: nr Amstetten; HT; ♂; NMBA?. 

minima. Austria. 

Diplosis minima Strobl 1880: 63. Austria: Styria; ST; ♀; NMBA?. 

minuta. Japan (Honshu). Artemisia princeps (Asteraceae). 

Dicrodiplosis minuta Shinji 1939a: 384. Japan: Honshu; ST; ♂,♀; lost. 

minuta. India (Madhya Pradesh). 

Feltiella minuta Grover 1979c: 46. India: Madhya Pradesh, Indore; HT; ♂; PGCA. Ref.: Gagné 1995a. 

moesta. UK. 

Cecidomyia moesta (also as sobria, p. 342) Walker 1856a: 104. UK: England; ST; ♂; BMNH. 

mollipes. Australia (New South Wales). 

Cecidomyia mollipes Skuse 1888: 92. Australia: New South Wales, Sydney; ST; ♂,♀; ANIC. 

montivaga. Argentina. Senecio mendocinus (Asteraceae). 

Janetiella montivaga Kieffer & Jörgensen 1910: 432. Argentina: Mendoza, Cordillera de Mendoza; ST; ♂,♀,P; lost. 

mori. Korean Penunsula, Japan (Honshu to Kyushu). Morus alba (Moraceae). 

Diplosis mori Yokoyama 1929: 381 (publ'n not seen). Japan; ST; ♂,♀,P,L,egg; ?. 

Trishormomyia maculata Sasaki 1931: 10. Japan; ST; ♂,♀,L; ?. 

nectandrae. Paraguay. Nectandra megapotamica (Lauraceae). 

Oligotrophus nectandrae Kieffer 1910d: 442. Paraguay; ST; A,P; ZMHB?. 

nigricollis. Germany. 

Cecidomyia nigricollis Meigen 1818: 97. Germany; T; ♀; MNHN. 

nigrina. UK. 

Cecidomyia nigrina Walker 1856a: 128. UK: England; ST; ♀; BMNH. Ref.: Panelius 1965: 18. 

nitida. Netherlands. 

Epidosis nitida Wulp 1874: 113. Netherlands: Driebergen; HT; ♀; ZMAN. Ref.: Panelius 1965: 18. 

niveonigra. Thailand. 

Doxodiplosis niveonigra Barnes 1927e: 269. Thailand: Khao Luang, Nakhon Sri Thammarat; HT; ♀; BMNH. 

nocticolor. Seychelles Is. 

Lepidodiplosis nocticolor Kieffer 1911: 322. Seychelles Is: Mahé, Cascade Estate; HT; ♂; BMNH?. 

notabilis. UK. 

Cecidomyia notabilis Walker 1856a: 114. UK: England; ST; ♂; BMNH. 

nothofagi. Chile. Nothofagus obliqua (Fagaceae). 

Rhopalomyia nothofagi Gagné 1973e: 447. Chile: Valdivia; HT; ♂; USNM. 

nubeculosa. UK. 

Cecidomyia nubeculosa Walker 1856a: 96. UK: England; ST; ♂; BMNH. 

nubilipennis. Australia (New South Wales). 

Cecidomyia nubilipennis Skuse 1890: 375. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

oleariae. New Zealand. Olearia paniculata (Asteraceae). 

Cecidomyia oleariae Maskell 1888: 257. New Zealand; ST; ♂,♀,P,L,egg; NZAC?. 

oleiphila. Eritrea. Olea chrysophylla (Oleaceae). 

Hormomyia oleiphila Del Guercio 1918a: 42. Eritrea; ST; A,P,L; lost. 

omalanthi. Australia (New South Wales). Omalanthus populifolius (Euphorbiaceae). 

Cecidomyia omalanthi Skuse 1890: 383. Australia: New South Wales, Illawarra Distr., Mount Kembla; HT; ♀; 

ANIC. 

oreas. Australia (New South Wales). 

Cecidomyia oreas Skuse 1888: 107. Australia: New South Wales, Glenbrook; HT; ♂; ANIC. 

oreiplmisia drana. Argentina (Mendoza). Verbena seriphioides (Verbenaceae). 

Rhopalomyia oreiplana Kieffer & Jörgensen 1910: 441. Argentina: Mendoza, Cordillera de Mendoza, nr Blanco 

Encallado; ST; ♀,P; lost. 


Pseudohormomyia orientalis Rao 1956b: 23. India: Uttar Pradesh, Dehra Dun, New Forest area; HT; ♂; ZSCI. 

orientalis. Indonesia (Java). Lepidagathis javanica (Acanthaceae).

416 

Stefaniella orientalis Felt 1921h: 142. Indonesia: Java, Ungaran Mts; HT; ♂; 

ovalis. India (Uttar Pradesh). 

Meinertomyia ovalis Grover 1961c: 411. India: Uttar Pradesh, Allahabad, Tagore Town; HT; ♀; PGCA. 

oxyacanthae. Germany. Crataegus sp. (Rosaceae). 

Tipula oxyacanthae Schrank 1803: 87. Germany: Bavaria; T; LT; lost?. 

paederiae. Philippines. Paederia tomentosa (Rubiaceae). 

Itonida paederiae Felt 1919a: 293. Philippines: Luzon, Laguna Prov., Los Baños; ST; ♂,♀; NYSM. 

pallipes. Chile. 

Lasioptera pallipes Philippi 1865: 630. Chile: Santiago; T; A; MNNC?. 

palustris. Sweden. 

Tipula palustris Linnaeus 1758: 588. Sweden; T; A; LSUK?. 

pandani. Indonesia (Java). Pandanus sp. (Pandanaceae). 

Trishormomyia pandani Felt 1921i: 270. Indonesia: Java, Tjibodas; ST; ♂,♀; NYSM. 

parilis. Australia (New South Wales). Eucalyptus corymbosa (Myrtaceae). 

Cecidomyia parilis Skuse 1888: 87. Australia: New South Wales, Middle Harbour; HT; ♀; ANIC. 

parva. Canada (Ontario). 

Lasioptera parva Walker 1848: 29. Canada: Ontario, Hudson's Bay; ST; A; BMNH. 

percita. Australia (New South Wales). 

Cecidomyia percita Skuse 1888: 72. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC. 

philippinensis. Philippines. Poss. Spatholobus philippinensis (Fabaceae). 

Cecidomyia philippinensis Felt 1919a: 294. Philippines: Luzon, Laguna Prov., Mt Makiling; HT; ♀; NYSM. 

piceae. Canada (Manitoba to New Brunswick), USA (New York, Maine, Connecticut). Picea canadensis (Pinaceae). 

Phytophaga picaea Felt 1926e: 229. Canada: Manitoba, Aweme and unstated, & Quebec, Fort Coulonge; ST; ♂,♀; 

NYSM & CNCI. 

Phytophaga piceae Anonymous 1926: 240, emend. of Phytophaga piceae Felt. 

pictipennis. Germany. 

Cecidomyia pictipennis Meigen 1830: 270. Germany; T; ♂; MNHN. 

pictipes. St. Vincent. 

Diplosis pictipes Williston 1896: 253. St. Vincent; ST; ♂; BMNH?. Cecidomyiinae. 

pinnicornis. Sweden. 

Tipula pinnicornis Linnaeus 1758: 589. Sweden; T; A; LSUK?. 

Cecidomyia pennicornis Kieffer 1913i: 218, missp. of pinnicornis. 

plagiata. UK. 

Cecidomyia plagiata Walker 1856a: 96. UK: England; ST; ♂; BMNH. 

plena. UK. 

Cecidomyia plena Walker 1856a: 118. UK: England; ST; ♂; BMNH. 

plumbea. Australia (New South Wales). 

Cecidomyia plumbea Skuse 1888: 69. Australia: New South Wales, Elizabeth Bay; ST; ♂,♀; ANIC. 

polyalthiae. Indonesia (Java). Polyalthia subcordata (Annonaceae). 

Calopedila polyalthiae Felt 1927d: 385. Indonesia: Java, Tjibodas, Mt Gede; ST; ♂,♀,P; NYSM. 


Microdiplosis pongamiae Mani 1934b: 425. India: Tamil Nadu, Thanjavur; HT; ♂; ZSCI. 


Myricomyia pongamiae Mani 1934b: 420. India: Tamil Nadu, Thanjavur; HT; ♂; ZSCI. 

pontaniiformis. Tertiary/Florissant shale. Myrica drymeja (Myricaceae). 

Cecidomyia pontaniiformis Cockerell 1908a: 66. USA: Colorado, Florissant shale; T; G; ?. 

populi. Japan (Honshu?). Populus maximowiczii (Salicaceae). 

Mikiola populi Shinji 1938j: 210. Japan: Honshu?; T; A; lost. 

populicola. Japan (Honshu). Populus nigra (Salicaceae). 

Mikiola populicola Shinji 1938j: 211. Japan: Honshu; T; A; lost. 

pratorum. Germany. 

Cecidomyia pratorum Meigen 1838: 21, jun. prim. hom. pratorum Waltl. Germany; T; ♀; MNHN.

417 

pratorum. Germany. 

Cecidomyia pratorum Winnertz 1853: 237, jun. prim. hom. pratorum Waltl.. Germany; ST; ♀; lost. 

producta. St. Vincent. Stachytarpheta jamaicensis (Verbenaceae). 

Hyperdiplosis producta Felt 1912b: 177. St. Vincent; ST; ♂,♀; NYSM. 

prosopidis. Argentina (Mendoza, San Luis). Prosopis alpataco, P. campestris, P. flexuosa (Fabaceae). 

Rhopalomyia prosopidis Kieffer & Jörgensen 1910: 427. Argentina: Mendoza, nr Chacras de Coria & La Paz, & San 

Luis, nr Alto Pencoso; ST; P,L; lost. 

pygmaea. France. 

Cecidomyia pygmaea Macquart 1826: 173. France; HT; ♂; MNHN?. 

quadrata. India (Tripura). 

Blastodiplosis quadrata Chandra 1993a: 244. India: Tripura, Agartala; HT; ♂; CSIA. 

quadrilobatus. India (West Bengal). Maesa indica (Myrsinaceae). 

Oligotrophus quadrilobatus Kieffer 1909a: 151. India: West Bengal, Kurseong; T; LT; lost?. 

reniformis. USA (Massachusetts). Fragaria sp. (Rosaceae). 

Cecidomyia reniformis Stebbins 1910: 36. USA: Massachusetts, vic. Springfield; T; G; NHSM. 

repleta. UK. 

Cecidomyia repleta Walker 1856a: 102. UK: England; ST; ♂; BMNH. 

schefflerae. Philippines. Schefflera insularum (Araliaceae). 

Scheuria schefflerae Felt 1920d: 231. Philippines: Luzon, Laguna Prov., Mt Makiling; HT; ♀; NYSM. 

scotti. Seychelles Is. 

Clinodiplosis scotti Kieffer 1911: 326. Seychelles Is: Silhouette I.; HT; ♀; BMNH. 

scutellata. Germany. 

Cecidomyia scutellata Meigen 1830: 270. Germany; T; ♂; MNHN. 

semiopaca. UK. 

Cecidomyia semiopaca Walker 1856a: 101. UK: England; ST; ♂; BMNH. 

shawiae. New Zealand. Olearia paniculata (Asteraceae). 

Dryomyia shawiae Anderson 1935: 422. New Zealand; ST; ♂,♀; BMNH. 

silvestrii. USA (Oregon, California). Quercus sp.; Q. chrysolepis, Q. garryana, Q. vaccinifolia (Fagaceae). 

Diplosis silvestrii Trotter 1911: 108. USA: California, Yosemite; T; G; ?. 

sissua. India (Uttar Pradesh). Dalbergia sp. (Fabaceae). 

Xylodiplosis sisso Bakhshi & Grover 1976: 100, nomen nudum. 

Xylodiplosis sissua Grover 1979c: 157. India: Uttar Pradesh, Allahabad; HT; ♂; PGCA. 

sobria. UK. 

Cecidomyia sobria Walker 1856a: 342 (also as moesta, p. 108). UK: England; ST; ♂; BMNH. 

sociata. UK. 

Cecidomyia sociata Walker 1856a: 87. UK: England; ST; ♀; BMNH. 

solennis. UK. 

Cecidomyia solennis Walker 1856a: 104. UK: England; ST; ♂; BMNH. 

spatulata. India (West Bengal). 

Aphidoletes spatulatus Grover 1979c: 66. India: West Bengal, Darjeeling; HT; ♂; PGCA. 

spatulata. India (Uttar Pradesh). 

Prodiplosis spatulata Jaiswal 1989b: 92. India: Uttar Pradesh, Garhwal Hills, Kotdwara; HT; ♂; CSIA. 

spondiasi. Philippines. Spondias mombin (Anacardiaceae). 

Mycodiplosis spondiasi Felt 1920d: 234. Philippines: Luzon, Laguna Prov., Los Baños; ST; ♀; NYSM. 

spongivora. Canada (Ontario). 

Cecidomyia spongivora Walker 1848: 30. Canada: Ontario, Hudson Bay, St. Martin's Falls, Albany R.; ST; ♂,♀; 

BMNH. 

squamosipes. Seychelles Is. 

Lepidodiplosis squamosipes Kieffer 1911: 321. Seychelles Is: I. Anonyme; HT; ♀; BMNH. 

stercoraria. Germany. 

Diplosis stercoraria Rübsaamen 1891d: 123. Germany: vic. Siegen; ST; ♂,♀; ZMHB. 

subhashensis. India (Uttar Pradesh).

418 

Sitodiplosis subhashensis Grover & Bakhshi 1978: 27. India: Uttar Pradesh, Allahabad, Mumfordganj; HT; ♂; 

PGCA. 

szeplegetii. Czechoslovakia, Hungary, Ukraine. Acer tataricum (Aceraceae). 

Oligotrophus szeplegetii Kieffer 1902a: 239 (as n. sp. also in Kieffer 1909b: 2). Hungary: Budapest; T; LT; lost?. 

tamaricis. Egypt. Tamarix articulata (Tamaricaceae). 

Cecidomyia tamaricis Kollar 1858: 159. Egypt: nr Cairo; ST; ♂,♀; ?. 

tausaghyzi. Kazakhstan. Scorzonera tausaghyz (Asteraceae). 

Perrisia tausaghyzi (as tau-saghyzi) Pravdin 1935: ? (publ'n not seen; cf. Barnes 1949: 157). Kazakhstan: Karatau 

Mts; T; ?; lost?. 

tecta. UK. 

Cecidomyia tecta Walker 1856a: 88. UK: England; ST; ♀; BMNH. 

tendens. UK. 

Cecidomyia tendens Walker 1856a: 112. England; ST; ♂; BMNH. 

tenuicornis. UK. 

Cecidomyia tenuicornis Walker 1856a: 109. UK: England; ST; ♂; BMNH. 

testacea. UK. 

Cecidomyia testacea Walker 1856a: 91. UK: England; ST; ♂; BMNH. 

thalictri. Poland. Thalictrum sp. (Ranunculaceae). 

Cecidomyia thalictri Loew 1850a: 22. Poland; ST; LT; ZMHB?. Known from the same host as Jaapiella thalictri 

Rübsaamen, this species appears to form a different gall. 

thujae. Canada (British Columbia). Thuja plicata (Pinaceae). 

Phytophaga thujae Hedlin 1959: 719. Canada: British Columbia, Langford; HT; ♂; CNCI. 

tormentillae. Poland. Potentilla recta (Rosaceae). 

Cecidomyia tormentillae Loew 1950a: 30. Poland; ST; G; ZMHB?. 

torreana. Cuba. 

Cecidomyia torreana Cook 1906: 252. Cuba: Prov. Santiago de Cuba; T; G; ?. 

torreyi. USA (New Mexico). Lycium torreyi (Solanaceae). 

Cecidomyia torreyi Felt 1925c: 63. USA: New Mexico; HT; ♀; USNM. 

transmarina. Australia (New South Wales). Host: Callistemon sp. (Myrtaceae). 

Heteropeza transmarina Schiner 1868: 5. Australia: New South Wales, vic. Sydney; ST; ♀,P; NHMW?. 

tricyclae. Argentina (Mendoza). Tricycla spinosa (Nyctaginaceae). 

Rhopalomyia tricyclae Kieffer & Jörgensen 1910: 440. Argentina: Mendoza, vic. Blanco Encalado & San Ignacio; T; 

P; lost. 

trinodata. India (Uttar Pradesh). 

Odontodiplosis trinodata Rao 1957: 11. India: Uttar Pradesh, Agra, Hewett Park; HT; ♂; SJCA. 

tristis. Germany. 

Cecidomyia tristis Meigen 1838: 22, jun. prim. hom. tristis Waltl. Germany: Bavaria; T; ♂; MNHN. 

tritomus. Tanzania. 

Coprodiplosis tritomus Kieffer 1913h: 98. Tanzania: Taveta; HT; ♀; MNHN. 

tropicalis. Tanzania. 

Coprodiplosis tropicalis Kieffer 1913h: 98. Tanzania: Taveta; HT; ♀; MNHN. 

tsugae. Canada (Ontario), USA (New York). 

Oligotrophus tsugae Felt 1907a: 27. USA: New York, Lake Clear; HT; ♂; NYSM. 

umbra. South America. 

Cecidomyia umbra Walker 1856c: 421. South America: HT; A; BMNH. 

verbenae. Argentina (Mendoza). Verbena aspera (Verbenaceae). 

Rhopalomyia verbenae Kieffer & Jörgensen 1910: 441. Argentina: Mendoza, Cordillera Mendoza; T; P; lost. 

verbesinae. USA (North Carolina). Verbesina alterniflora (Asteraceae). 

Cecidomyia verbesinae Beutenmüller 1907a: 391. USA: North Carolina, Buncombe Co., Valley of the Black Mts; 

HT; LT; AMNH. 

violacea. Australia (New South Wales). 

Cecidomyia violacea Skuse 1888: 101. Australia: New South Wales, Sydney, Woronora, Middle Harbour, & Blue

Mts; ST; ♂,♀; ANIC. 

vitulans. Australia (New South Wales). 

Cecidomyia vitulans Skuse 1888: 63. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC. 

Cecidomyiidae 

BAEONOTUS 

Baeonotus Byers 1969: 366. Type species, microps Byers (orig. des.) 

Eastern Nearctic; 1 sp.; Baeonotini. Ref.: Jaschhof 2009 (review). 

microps. USA (Virginia). 

Baeonotus microps Byers 1969: 366. USA: Virginia, Giles Co., Mountain Lake Biol. Stn; HT; ♀; SEMC. 

NOMINA DUBIA 

These taxa cannot be identified. Types are lost, presumed lost, or reduced to fragments. Their hosts are either 

unknown or the cecidomyiids are only collaterally involved with galls or damage. In the family-level subdivisions 

below, any genera are listed first, followed by species. 

yasukunii Shinji 1944: 315, Lestremia. Japan; T; ♀; lost. 

Lestremiinae 

Micromyinae 

Lasiopteryx Stephens 1829a: 53, (publ'n not seen); also in Stephens 1929b: 239. Cecidomyia obfuscata Meigen (subs. 

des. Westwood 1840: 126). 

brachyptera Kieffer 1913b: 92, Pezomyia. Algeria: Algiers; T; ♀; lost. Jaschhof 1998b: 548. 

cinerascens Rübsaamen 1926: 8, Pezomyia. Unknown; HT; ♀; lost. Based on illustration of head. 

coronatus Kieffer 1895l: 93, Prionellus. France: vic. Bitche; T; ♀; lost. 

dimorphogyna Rübsaamen 1899b: 543, Campylomyza. Germany: S; ♂,♀,P; lost. 

hirta Marshall 1896: 244, Campylomyza. New Zealand: Lincoln; HT; A; lost. 

lepida Winnertz 1870b: 14, Campylomyza. Germany; T; ♀; lost. 

lincolniensis Marshall 1896: 243, Campylomyza. New Zealand: Lincoln; ST; ♂; lost. 

luculenta Meigen 1830: 271, Campylomyza. Germany; ST; ♀; lost. 

maculatus Kieffer 1901c: 41, Prionellus. France: vic. Bitche; T; ♀; lost. 

magna Marshall 1896: 245, Campylomyza. New Zealand: Lincoln; HT; ♀; lost. 

minuta Marshall 1896: 243, Campylomyza. New Zealand: Lincoln; HT; ♀; lost. 

mori Sasaki 1931: 4, Urosema. Japan; ST; ♂,♀,L; lost. 

muscicola Kieffer 1895l: 93, Prionellus. France: Guinkirchen; ST; ♂; lost. 

nitida Marshall 1896: 244, Campylomyza. New Zealand: Lincoln; HT; ♀; lost. 

obfuscata Meigen 1818: 90, Lasiopteryx. Germany; T; A; lost. 

ordinaria Marshall 1896: 247, Campylomyza. New Zealand: Lincoln; HT; ♂,♀; lost. 

perrisi Kieffer 1895m: 109, Aprionus. France; ST; ♂,P,L; lost. 

picea Winnertz 1870b: 20, Campylomyza. Germany; T; ♀; lost. 

praecox Kieffer 1894e: clxxvi, Prionota. France: Bitche; T; ♀; lost. 

robusta Marshall 1896: 246, Campylomyza. New Zealand: Lincoln; HT; ♂; lost. 

sejunctus Kieffer 1901c: 40, Prionellus. France: vic. Bitche; T; ♀; lost. 

squalida Winnertz 1870b: 16, Campylomyza. Germany; T; ♀; lost. 

squamata Marshall 1896: 245, Campylomyza. New Zealand: Lincoln; HT; ♀; lost. 

419

sylvicola Winnertz 1870b: 17, Campylomyza. Germany; T; ♀; lost. 

tenuis Marshall 1896: 242, Campylomyza. New Zealand: Lincoln; HT; A; lost. 

turmalis Rondani 1840: 25, Neurolyga. Italy; T; A; lost. 

vittata Winnertz 1870b: 17, Campylomyza. Germany; T; ♀; lost. 


agricolae Marshall 1896: 232, Miastor. New Zealand: Lincoln; T; A; lost. 

agricolae Marshall 1896: 240, Cecidomyia. New Zealand: Lincoln; T; ♀; lost 

albimana Winnertz 1853: 296, Cecidomyia. Germany; ST; ♀; lost. 

aurea Marshall 1896: 241, Cecidomyia. New Zealand: Mt Torlesse; HT; A; lost. 

difficilis Marshall 1896: 233, Miastor. New Zealand: Lincoln; HT; A; lost. 

Hincompleta Loew 1850b: 32, Cecidomyia. Upper Eocene - Baltic amber; T; A; lost?. 

magna Marshall 1896: 239, Cecidomyia. New Zealand; ST; ♂,♀; lost. 

mastersi Skuse 1888: 59, Miastor. Australia: New South Wales, Elizabeth Bay; HT; ♂; lost. 

Hminutissima Loew 1850b: 32, Cecidomyia. Upper Eocene - Baltic amber; T; A; lost?. 

ordinaria Marshall 1896: 241, Cecidomyia. New Zealand: Lincoln; T; ♂; lost. 

pallidula Wulp 1874: 113, Colpodia. Netherlands: Schevening; ST; ♀; lost?. 

procax Skuse 1888: 58, Miastor. Australia: New South Wales, Elizabeth Bay, nr Sydney; HT; ♀; lost. 

protogaea Heer 1856: 32, Cecidomyia. France: Aix en Provence, a Miocene fossil; ST; ♀; lost. 

Cecidomyiinae 

Gonioclema Skuse 1888: 60, as subg. of Cecidomyia. Type species, Cecidomyia pauxillula Skuse (mon.). The original 

description does not sufficiently characterize this genus and the type of pauxillula is damaged beyond 

recognition. 

Necrophlebia Skuse 1888: 111, as subg. of Cecidomyia. Type species, Cecidomyia volitans Skuse (mon.). The original 

description does not sufficiently characterize this genus and the type series of C. volitans is damaged beyond 

recognition. 

abbreviata Waltl 1837: 279, Cecidomyia. Germany: Bavaria; T; ♂; lost?. 

acarinivora Tölg 1921: 33, Feltiella. Austria: vic. Vienna; HT; ♂; lost. Cf. Gagné 1995: 28. 

acarivora Kieffer 1913i: 202, Feltiella, nomen nudum. 

acuticauda Kieffer & Herbst 1906: 227, Janetiella. Chile: vic. Concepción; T; ♀; lost. 

acuticauda Kieffer & Herbst 1905: 64, Janetiella, nomen nudum. 

adusta Skuse 1888: 82, Cecidomyia. Australia: New South Wales, Woronora. HT; ♀ (orig. given as ♂); ANIC 

(fragments). 

agilis Winnertz 1853: 265, Cecidomyia. Germany; ST; ♂; lost. 

albilabris Winnertz 1853: 217, Cecidomyia. Germany; ST; ♂,♀; lost. 

albulipennis Skuse 1890: 378, Cecidomyia. Australia: New South Wales, Sydney, Dunoon & Richmond R., & 

Queensland, Burpengary, nr Brisbane; ST; ♂,♀; ANIC (fragments). 

amoena Loew 1850a: 31, Cecidomyia. Poland; ST; ♀; lost?. 

amyotii Fitch 1861: 773, Cecidomyia. USA: New York, Salem; ST; ♀; lost. Cf. Gagné & Doane 1999:62. 

annulipes Meigen 1830: 268, Cecidomyia. Germany; T; ♀; lost. 

annulipes Walsh 1864b: 629, Cecidomyia. USA: Illinois, nr Rock Island, on Salix spp. (Salicaceae); ST; ♀; lost. 

ardens Skuse 1888: 75, Cecidomyia. Australia: New South Wales, Sydney; HT; ♀; ANIC (fragments). 

atra Roser 1840: 50, Cecidomyia. Germany; T; A; presumed lost. 

atricornis Walsh 1864b: 628, Cecidomyia. USA: Illinois, nr Rock Island, on Salix spp. (Salicaceae); HT; ♂; lost. 

atrocularis Walsh 1864b: 626, Cecidomyia. USA: Illinois, nr Rock Island, on Salix spp. (Salicaceae); ST; ♂,♀; lost. 

bellula Skuse 1888: 87, Cecidomyia. Australia: New South Wales, Peat's Ferry; HT; ♂; ANIC (fragments). 

bicolor Meigen 1818: 98, Cecidomyia. Germany; T; ♂; lost. 

bifida Brèthes 1914: 154, Trisopsis. Argentina: Buenos Aires, Buenos Aires: HT; ♂; lost. Cf. Gagné 1994a: 200. 

bipunctata Winnertz 1853: 288, Cecidomyia. Germany; ST; ♂; lost. 

bombycina Skuse 1888: 79, Cecidomyia. Australia: New South Wales, Sydney; HT; ♀; ANIC (fragments). 

420

421 

brachyptera Kieffer 1913b: 92, Pezomyia. Algeria: nr Algiers; T; ♀; lost. 

brevitata Gagné 1989c: 161, Cecidomyia, new name for Cecidomyia abbreviata Skuse. 

abbreviata Skuse 1888: 75, Cecidomyia, jun. prim. hom. abbreviata Waltl. Australia: New South Wales, Sydney; 

HT; ♀; ANIC (fragments). 

caeca Skuse 1888: 76, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♀ (orig. given as ♂); ANIC 

(fragments). 

certa Skuse 1888: 100, Cecidomyia. Australia: New South Wales, Woronora; HT; ♀; ANIC (fragments). 

cineraria Skuse 1888: 106, Cecidomyia. Australia: New South Wales, Woronora; HT; ♂; ANIC (fragments). 

conferta Skuse 1888: 118, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC (fragments). 

confinis Skuse 1888: 70, Cecidomyia. Australia: New South Wales, Sydney; HT; ♂; ANIC (fragments). 

conspecta Skuse 1888: 91, Cecidomyia. Australia: New South Wales, Richmond, Hawkesbury Distr.; HT; ♀; ANIC 

(fragments). 

contigua Skuse 1888: 103, Cecidomyia. Australia: New South Wales, Wheeny Creek, Hawkesbury Distr.; HT; ♂; lost. 

decemmaculata (as decem-maculata) Walsh 1864b: 631, Cecidomyia. USA: Illinois, nr Rock Island, on Salix spp. 

(Salicaceae); ST; ♂,♀; lost. 

discedens Skuse 1888: 93, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC (fragments). 

dubiella Gagné 1989c: 162, Cecidomyia, new name for Cecidomyia dubia Marshall. 

dubia Marshall 1896: 235, Cecidomyia, jun. prim. hom. dubia Kieffer. New Zealand: Lincoln; HT; ♀; lost. 

dumetorum Winnertz 1853: 264, Cecidomyia. Germany; ST; ♂; lost. 

elegans Winnertz 1853: 287, Cecidomyia. Germany; ST; ♂; lost. 

faceta Skuse 1888: 81, Cecidomyia. Australia: New South Wales, Sydney; HT; ♂; lost. 

flammula Skuse 1888: 124, Cecidomyia. Australia: New South Wales, Sydney; HT; ♀; ANIC (fragments). 

flavella Kieffer 1913i: 215, Cecidomyia, new name for Cecidomyia flava Marshall. 

flava Marshall 1896: 239, Cecidomyia, jun. prim. hom. flava Meigen. New Zealand: Wanganui; HT; ♂; lost. 

flavida Blanchard 1852: 350, Cecidomyia. Chile: Coquimbo, La Serena; T; A; lost. 

fragilina Gagné 1989c: 162, Cecidomyia, new name for Cecidomyia fragilis Marshall. 

fragilis Marshall 1896: 237, Cecidomyia, jun. prim. hom. fragilis Loew. New Zealand: Lincoln; ST; ♂; lost. 

frischii Bremi 1847a: 24, Cecidomyia. Switzerland; ST; A,P,L; lost (based on descr. in Frisch 1820). 

furva Skuse 1888: 90, Cecidomyia. Australia: New South Wales, Middle Harbour; HT; ♀; ANIC (fragments). 

fuscicollis Meigen 1818: 97, Cecidomyia. Germany; T; ♂; lost. This name has been attributed to Bouché 1834: 25, but 

Bouché's mention was only a misidentification of fuscicollis Meigen. 

fuscipennis Meigen 1818: 98, Cecidomyia. Germany; T; ♀; lost. 

geniculata Winnertz 1853: 251, Cecidomyia. Germany; ST; ♂; lost. 

gibberosa Skuse 1888: 123, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC (fragment). 

gibbula Skuse 1888: 67, Cecidomyia. Australia: New South Wales, Middle Harbour; HT; ♀; ANIC (fragments). 

gilva Skuse 1888: 93, Cecidomyia. Australia: New South Wales, Gosford; HT; ♂; ANIC (fragments). 

grandipennis Skuse 1890: 384, Cecidomyia. Australia: New South Wales, Mossman's Bay, nr Sydney; HT; ♂; ANIC 

(fragments). 

grisea Bremi 1847a: 49, Cecidomyia. Switzerland: Zurich; ST; A; lost. 

helmsi Skuse 1890: 379, Cecidomyia. Australia: New South Wales, Dunoon, Richmond R.; ST; ♂,♀; ANIC 

(fragments). 

hirta Marshall 1896: 237, Cecidomyia. New Zealand: Lincoln; ST; ♀; lost. 

humilis Skuse 1888: 95, Cecidomyia. Australia: New South Wales, Wheeny Creek, Hawkesbury Distr.; ST; ♂,♀; 

ANIC (fragments). 

impudica Winnertz 1853: 269, Cecidomyia. Germany; ST; ♂; lost. 

indotata Skuse 1888: 107, Cecidomyia. Australia: New South Wales, Middle Harbour; HT; ♂; ANIC (fragments). 

Hlargo Meunier 1901: 192, Cecidomyia. Holocene/African copal; T; A; lost?, based only on an antenna. 

lentipes Winnertz 1853: 291, Cecidomyia. Germany; ST; ♀; lost. 

loewi Skuse 1888: 108, Cecidomyia. Australia: New South Wales, Como; HT; ♂ (orig. given as ♀); ANIC (fragments). 

lucida Skuse 1888: 73 Cecidomyia. Australia: New South Wales, Woronora; HT; ♂; ANIC (fragments). 

luteofusca Rondani 1840: 17, Dasineura. Italy: vic. Parma; T; A; lost. 

lutulenta Skuse 1888: 110, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC (fragments). 

macella Skuse 1888: 121, Cecidomyia. Australia: New South Wales, Sydney; HT; ♀; lost.

422 

marshalli Gagné 1989c: 162, Cecidomyia, new name for Cecidomyia difficilis Marshall. 

difficilis Marshall 1896: 235, Cecidomyia, jun. prim. hom. difficilis Skuse. New Zealand: Lincoln; HT; ♂; lost. 

mastersi Skuse 1888: 127, Lasioptera. Australia: New South Wales, Woronora; HT; ♀; ANIC (fragments). 

maxima Roser 1840: 50, Cecidomyia. Germany; T; A; presumed lost. 

melana Marshall 1896: 236, Cecidomyia. New Zealand: Lincoln; HT; ♀; lost. 

miniscula Gagné 1989c: 162, Cecidomyia, new name for Cecidomyia minuta Marshall. 

minuta Marshall 1896: 236, Cecidomyia, jun. prim. hom. minuta Winnertz. New Zealand: Lincoln; T; ♀; lost. 

modesta Winnertz 1853: 262, Cecidomyia. Germany; ST; ♂; lost. 

montana Skuse 1888: 84, Cecidomyia. Australia: New South Wales, Glenbrook; HT; ♀; ANIC (fragments). 

nana Winnertz 1853: 252, Cecidomyia. Germany; ST; ♂,♀; lost. 

negotiosa Skuse 1888: 104, Cecidomyia. Australia: New South Wales, North Willoughby; ST; ♂,♀; ANIC 

(fragments). 

fallax Skuse 1888: 85, Cecidomyia. Australia: New South Wales, Woronora; HT; ♀; lost. 

nervosa Waltl 1837: 279, Cecidomyia. Germany: Bavaria; T; ♂; lost?. 

nigra Meigen 1804: 39, Cecidomyia. Germany; ST; ♂,♀; lost. 

nigricornis Schrank 1803: 86, Tipula. Germany: Ingolstadt; T; A; lost. 

nobilis Skuse 1888: 66, Cecidomyia. Australia: New South Wales, Gosford; HT; ♂; ANIC (fragments). 

obscura Meigen 1838: 22, Cecidomyia. Germany: Bavaria; T; ♀; lost. 

obscura Rondani 1840: 18, Dasineura. Italy: vic. Parma; T; A; lost. 

obsoleta Skuse 1888: 83, Cecidomyia. Australia: New South Wales, Glenbrook; HT; ♀; ANIC (fragments). 

ochracea Winnertz 1853: 262, Cecidomyia. Germany; ST; ♂; lost. 

orbitalis Walsh 1864b: 623, Cecidomyia. USA: Illinois, nr Rock Island, on Salix spp. (Salicaceae); ST; ♂,♀,P; lost. 

ornata Say 1824: 357, Cecidomyia. USA: Pennsylvania, Philadelphia; HT; A; lost. 

parietina Skuse 1888: 125, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC (fragments). 

paula Skuse 1888: 74, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♀; lost. 

pauxillula Skuse 1888: 61, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♀; ANIC (fragments). 

pratorum Waltl 1837: 280, Cecidomyia. Germany: Bavaria; T; ♂; lost? (same specimen(s) as pratorum Meigen?). 

prisca Skuse 1888: 125, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC (fragments). 

probata Skuse 1888: 77, Cecidomyia. Australia: New South Wales, Middle Harbour, nr Sydney; HT; ♀; ANIC 

(fragments). 

pusilla Meigen 1818: 91, Lasioptera. Germany; T; A; lost. 

quaesita Skuse 1888: 90, Cecidomyia. Australia: New South Wales, Woronora; HT; ♀; ANIC (fragments). 

regilla Skuse 1888: 64, Cecidomyia. Australia: New South Wales, Elizabeth Bay, HT; ♀; ANIC (fragments). 

resupinans Bach 1859: 252, Cecidomyia. Germany; T; G (unspecified host); lost. 

ribesii Meigen 1818: 98, Cecidomyia. Austria; T; ♂; lost. 

rubicunda Skuse 1888: 109, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC (fragments). 

rusticula Skuse 1888: 89, Cecidomyia. Australia: New South Wales, Como; HT; ♂; ANIC (fragments). 

saxatilis Skuse 1888: 102, Cecidomyia. Australia: New South Wales, Elizabeth Bay, Sydney; ST; ♂,♀; ANIC 

(fragments). 

scelesta Skuse 1888: 96, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC (fragments). 

scenica Skuse 1888: 72, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC (fragments). 

scoparia Marshall 1896: 238, Cecidomyia. New Zealand: Lincoln; ST; ♀; lost. 

scutellata Say 1823: 17, Campylomyza. USA: MO; T; A; lost. Jaschhof 1998b: 550. 

senilis Skuse 1888: 88, Cecidomyia. Australia: New South Wales, Coogee Bay, nr Sydney; HT; ♀; ANIC (fragments). 

signata Winnertz 1853: 246, Cecidomyia. Germany; ST; ♂,♀,L; lost. 

simplex Loew 1850a: 34, Cecidomyia. Poland; ST; ♀; lost?. 

strobilina Rudow 1875b: 279, Cecidomyia. Germany; ST; ♀,P; presumed lost. Panelius 1965: 75. 

sulfurea Skuse 1888: 86, Cecidomyia. Australia: New South Wales, Sydney; HT; ♀; ANIC (fragments). 

sylvatica Winnertz 1853: 249, Cecidomyia. Germany; ST; A; lost. 

tergata Fitch 1845b: 264, Cecidomyia. USA: New York, Salem; ST; A; lost. 

thoracica Fitch 1845b: 264, Cecidomyia. USA: New York, Salem; ST; A; lost. Gagné 1973c: 875 combined with 

Mycodiplosis a specimen from the Fitch collection labelled thoracica and considered that species the senior 

synonym of M. coniophaga Winnertz, but that specimen is not one of Fitch's original specimens (Barnes 1988:

423 

110). 

tristis Waltl 1837: 279, Cecidomyia. Germany: Bavaria; T; ♂; lost? (same specimen(s) as tristis Meigen?). 

unicolor Loew 1850a: 33, Cecidomyia. Poland; ST; ♀; lost?. 

varicolor Bremi 1847a: 55, Cecidomyia. Switzerland: vic. Zurich; ST; ♂,♀; lost. 

varipes Roser 1840: 50, Cecidomyia. Germany; T; A; presumed lost. 

vegrandis Skuse 1888: 78, Cecidomyia. Australia: New South Wales, Elizabeth Bay; ST; ♂,♀; ANIC (fragments). 

ventralis Say 1824: 357, Lasioptera. USA: Pennsylvania, Philadelphia; HT; ♀; lost. 

verbasci Meigen 1838: 23, Cecidomyia. Germany; T; A; lost. 

villosa Skuse 1888: 80, Cecidomyia. Australia: New South Wales, Elizabeth Bay; HT; ♂; ANIC (fragments). 

virgata Skuse 1890: 386, Lasioptera. Australia: New South Wales, Elizabeth Bay; ST; ♂,♀; ANIC (fragments). 

vittata Meigen 1838: 22, Cecidomyia. Germany; T; A; lost. 

volitans Skuse 1888: 111, Cecidomyia. Australia: New South Wales, Middle Harbour & Woronora; ST; ♀; ANIC 

(fragments). 

wanganuiensis Marshall 1896: 238, Cecidomyia. New Zealand: Wanganui; ST; ♀; lost. 

xanthopyga Winnertz 1853: 263, Cecidomyia. Germany; ST; ♂; lost. 

CECIDOMYIIDAE NOMINA NUDA AND INVALID SPECIES 


bujunensis Naora 1933: 216, Cecidomyia. 

adwantii Deshpande 1982: 41, Claspittomyia. 

agropyri Moiseev 1962: 118 (pub'n not seen), Comtarinia. 

anomala Keilbach 1982: 344, Cecidomyia. 

astericola Shinji 1944: 98, Dasyneura. 

*citrifrugis Jiang 1994: 245, Resseliella. 

coloradensis Felt 1940: 56, Rhabdophaga 

confinis Stephens 1829b: 239, Lasiopteryx. 

conjuncta Meunier 1899a: 161, Cecidomyia. 

crumenalis Vogler 1906: 331, (pub'n not seen), genus not given, fide Skuhravá 1986: 294. 

*datifolia Jiang 1994: 245, Dasineura. 

Diplobremia Felt 1931: 449. Invalid; not accompanied by fixation of type species. No included species. 

Diplodasyomyia Felt 1931: 451. Invalid; not accompanied by fixation of type species. No included species. 

Diplohindomyia Felt 1931: 455. Invalid; not accompanied by fixation of type species. No included species. 

elegantissima Stephens 1829b: 239, Lasiopteryx. 

eupatrii Shinji 1944: 147, Lasioptera. 

euphobiae Shinji 1944: 324, Lasioptera, based on gall only. 

fagi Shinji 1938j: 210, Catocha. 

filicornis Keilbach 1982: 344, Cecidomyia. 

*flabilis Fedotova & Sidorenko 2009d: 229, Tuguridiplosis 

flavicans Stephens 1829b: 240, Cecidomyia. 

galiicaulis Buhr 1964: 527, Dasyneura. Originally attributed by Buhr to Stelter. 

gigantea Stephens 1829b: 240, Diomyza. 

*Hammamyia Vaillant 1955: 42. 

*hymenocleae Felt 1940: 326, Rhopalomyia. 

inops Stephens 1829b: 239, Lasiopteryx. 

intergerina Yagi & Sawada 2000: 7, Cordylomyia. 

iridescens Stephens 1829b: 240, Cecidomyia. 

kirii Shinji 1938j: 210, Catocha. 

leacheana Stephens 1829b: 240, Cecidomyia. 

liquidambaris Givulesçu 1984: 125, Phylocecidium. 

liquidambrus Evenhuis 1994: 187, Phylocecidium, missp. of liquidambaris Givulesçu. 

maculipennis Stephens 1829b: 240, Cecidomyia.

424 

*morulae Jiang 1994: 246, Contarinia. 

*muscicola Vaillant 1955: 42, Hammamyia (q.v. above). 

muscorum Skuse 1890: 377, Cecidomyia. 

nawai Shinji 1944: 127, Diplosis. 

neglecta Buhr 1965: 1375, Geocrypta. 

oxyacanthae Stephens 1829b: 240, Cecidomyia. 

Pararhizomyia Mamaev 1998d: 6. 

*pentagona Jiang 1994: 246, Lestodiplosis. 

*photophila Jaiswal 2005: 406 (Winnertzia). India: Uttar Pradesh, Garhwal Hills, Kotdwara; HT; ♂; ?. Invalid: type 

deposition not given (ICZN 1999). 

pirolae Buhr 1965: 912, Dasyneura. 

pontiensis Villalta 1957: 64, Mikiola. 

recta Keilbach 1982: 344, Cecidomyia. 

rosea Stephens 1829b: 240, Cecidomyia. 

rubra Stephens 1829b: 240, Diomyza. 

sambucicola Yagi & Sawada 2000: 16, Lasioptera. 

smilacifoliae Shinji 1944: 87, Silvectrina. 

stachydis Buhr 1965: 1215, Asphondylia. 

styracophila Shinji 1944: 33, Rhopalomyia. 

tamaricia Marikovskij 1955b: 104, Dasyneuriola. 

tenella Loew 1850b: 32, Molobraea. 

viburni Shinji 1939c: 586, Dasyneura. 

*weni. Jiang in Wen et al. 1998: 80. 

yakovlevi Fedotova 2000b: 697, Etsuhoa. Attributed to Mamaev, B.M. but never published (B.M. Mamaev, in litt.). 

yasukunii Shinji 1944: 109, Lestremine. 

yukawai Fedotova & Sidorenko 2008a: 230, Monepidosis (also as yukawi in key, p. 230), based on Monepidosis 

pectinata Mamaev of Yukawa 1971: 81. Name unavailable; no holotype designated. 

*zathoyli Huang et al. 2005: 646, Asphondylia. 

TAXA NO LONGER REFERABLE TO CECIDOMYIIDAE 

Cecidomium Westwood 1854: 394. Type species, grandaevum Westwood (mon.). 

grandaevum Westwood 1854: 394 and pl. XV, fig. 21, Cecidomium. Lower Cretaceous/UK: England, Dorset, Middle 

Purbecks of Durdlestone Bay; T; A; ?. Possibly Chironomidae. 

Cecidomyia dubia Heyden 1862: 80. Germany: Brown coal/Salzhausen; T; G; ?. Gall on Juglans acuminata. Possibly 

Acarina. 

Cecidomyia muscosa Stebbins 1910: 35. USA: Massachusetts, vic. Springfield; T; G; NHSM. Gall on Rubus 

allegheniensis (Rosaceae). Cynipidae (Hymenoptera). 

Cecidomyia poculum Osten Sacken 1862: 201. USA: Washington, DC; ST; galls; USNM. Gall on Quercus alba 

(Fagaceae). Xystoteras poculum Cynipidae (Hymenoptera). 

Cecidomyiaceltis Patton 1897: 247, (as Cecidomyia-celtis). Type species, deserta Patton (orig. des., as n. g., n. sp.). 

Cecidomyiaceltis deserta Patton 1897: 247. USA: Connecticut, Orange, on Celtis occidentalis. Agromyzidae (Diptera). 

Euprojoannisia Brèthes 1914: 155. Type species, platensis Brèthes (orig. des.). 

Euprojoannisia platensis Brèthes 1914: 156. Argentina: Buenos Aires, General Urquiza; HT; ♀; MACN. 

Ceratopogonidae (Diptera). 

Limnopneuma Enderlein 1911c: 195. Type species, Sciara coracina Zetterstedt (orig. des.). 

Sciara coracina Zetterstedt 1851: 3739. Norway: vic. Oslo; HT; ♀; MZLU?. Refer to Diptera: Sciaridae (= 

Scatopsciara vitripennis (Meigen), Jaschhof 1998b: 552). 

Lasioptera recessa Scudder 1877: 745. Eocene?/USA: Colorado, unstated but prob. White R., Chagrin Valley; T; A; ?. 

Probably not Cecidomyiidae. 

Lithomyza Scudder 1877: 746. Type species, Lithomyza condita Scudder (mon.). 

Lithomyza condita Scudder 1877: 746. Eocene?/USA: Colorado, White R. Chagrin Valley; HT; A; ?. Probably not 

Cecidomyiidae.

Moehnia Pritchard 1960a: 308. Type species, erema Pritchard (orig. des.). 

Moehnia erema Pritchard 1960a: 309. USA: California, Berkeley; HT; ♀; USNM. Sciaridae (Diptera). 

Villigera Karsch 1877: 17. Type species, frauenfeldi Karsch (orig. des.). 

Villigera frauenfeldi Karsch 1877: 18. China: Hong Kong; T; ♂; ?. Margarodidae (Hemiptera). 

APPENDIX 1: CLASSIFICATION OF THE FAMILY CECIDOMYIIDAE 

In this update to Gagné (2004) I have added the number of species included in each genus. Generic names 

described since or otherwise not included in Gagné (2004b) are preceded by an asterisk. Commentary for family group 

names and genera has been changed or augmented. 

This section places the family groups and genera in some context but is still largely imperfect. The best known 

groups presently are the old "Lestremiinae," now distributed among three subfamilies, the Catotrichinae, the 

Lestremiinae, and the Micromyinae. Advances in these groups are the result of recent comprehensive studies done by 

Mathias Jaschhof and co-authors that place the groups in context and newly illuminate some of the fauna of various 

parts of the world, previously nearly completely unexplored. 

Advances in the Porricondylinae have been minor despite a great number of new genera and species published 

since 2003 by Fedotova and Sidorenko based on specimens collected in Primorskiï Territory of Far East Russia and by 

Fedotova with Perkovsky on specimens from Ukrainian amber (see Literature Cited). More specific criticism of those 

authors' work is given under various headings elsewhere in this appendix, but these papers generally describe species 

on the basis of a limited number of study specimens, usually the holotype only, either caught in flight traps or found in 

amber. Descriptions and illustrations of many species are based on evidently imperfect, at least imperfectly prepared or 

rendered, specimens. Illustrations of the more diagnostic parts of male genitalia where several separate structures are 

superimposed are not clear and the use of non-morphological terms such as "lobes" do not properly convey the 

relationships among the various parts. This is especially unhelpful in groups that require at least dorsoventral and lateral 

views to properly illustrate the male genitalia. Genera are mostly monotypic, many separated from other genera on 

superficial grounds, and the descriptions often give no indication that existing genera were considered. Further, there is 

no evidence that the papers were peer-reviewed, despite the stated policy of some of the journals in which the papers 

appeared. 

Work has continued apace in the Cecidomyiinae by many authors on the fauna of various parts of the world. 

A similar situation to that outlined under Porricondylinae occurs in Cecidomyiinae in which Fedotova alone or with 

Sidorenko, and Fedotova with or without Perkovsky, described a total of 81 genera in this subfamily since 2003, 

inclusively (see Literature Cited). All these genera and their species will require revision by anyone taking on the study 

of these groups whose attraction has been diminished by these authors' substandard work. Fedotova et al. propose many 

new family group names, but only one, the Karshomyiini, is employed here and then only because Fedotova and 

Sidorenko described many genera that, because they are evidently related to Karshomyia, make using the tribe a handy 

excuse to remove those genera from the long list of genera that otherwise would not be placed to tribe. 

For the Cecidomyiinae I have provided the main shared derived characters that differentiate the supertribes, 

tribes, and subtribes. Many genera separately listed here under unplaced Cecidomyiidi and Lasiopteridi still cannot be 

grouped into tribes. One reason for this situation is that many of the old genera were too poorly and incompletely 

described, but these should be rediscovered as particular faunas become better known. Another reason is that, for the 

tropical areas of the world, at least, we know so little of the fauna that many new taxa seemingly stand alone. As more 

comprehensive revisions are made of the fauna generally or of the species found on particular host groups, the affinities 

of some of the unplaced genera should become clear. 

Subfamily Catotrichinae 

This subfamily consists of only eight known species distributed among three genera. This group is the most 

generalized of the family. The one known fossil species, from the Upper Jurassic/Lower Cretaceous of Siberia, is the 

oldest known cecidomyiid fossil. Refs.: Jaschhof 2001a, Jaschhof and Jaschhof 2008. 

Catotricha 5 

*Mesotrichoca 1 

*Trichotoca 2 

425

Subfamily Lestremiinae 

The Lestremiinae comprise 12 genera and 98 species, one of them unplaced to genus. This group was only 

recently separated from the remainder of the old Lestremiinae that are now placed under the subfamily Micromyinae 

(Jaschhof and Jaschhof 2009a). The Holarctic Lestremiinae, excluding Anarete and Conarete, were revised in Jaschhof 

1998b and further elucidated in Jaschhof and Jaschhof 2009a. The classification of Lestremiinae in the list below 

follows the latter paper. 

Allarete 11 

Allaretella 1 

Anarete 33 

Anaretella 6 

Conarete 12 

*Eomastix 1 

Gongromastix 7 

*Insulestremia 1 

Lestremia 18 

Mangogrostix 1 

Neolestremia 3 

Wasmanniella 3 

Tribe Lestremiini 

Subfamily Micromyinae 

Micromyinae are another ancient but more diverse group of 47 genera (one with four subgenera) and 557 

species, 23 of them unplaced to genus. Ten of the genera are unplaced to tribe and five of those should be relegated to 

nomina dubia, per M. Jaschhof (in litt.). The subfamily appears to be monophyletic and the sister group to the 

Porricondylinae + Cecidomyiinae (Jaschhof 2001a, Jaschhof & Jaschhof 2009a). It includes the oldest described fossil 

of the Cecidomyiidae, Eltxo cretaceus Arillo & Nel (2000), from the Lower Cretaceous. Micromyines are 

mycophagous and feed in mushrooms and various forms of detritus including rotten logs. Mamaev and Krivosheina 

(1965) presented a well-illustrated overview of larvae and Mamaev 1968b contains general information on biology. 

The arrangement of tribes and genera listed below follows Jaschhof & Jaschhof 2009a, but other treatments of parts of 

the subfamily are in Jaschhof and Hippa 1999a,b on the large new genus Pseudoperomyia from Malaysia with notes on 

the phylogeny of Micromyidi, and Jaschhof 2000, 2001b on Pseudoperomyia and the Peromyiini of Japan. The 

Micromyini, especially outside the Holarctic Region, still provide a vast field for further taxonomic study. This is 

exemplified by the list of only six named species from the Neotropics in Gagné 1994a, although indications are that the 

fauna is much richer (Gagné & Jaschhof 2009), and by the genus Aprionus, currently known from 101 named species 

but with an additional 150 undescribed species from Japan and peninsular Malaysia (Jaschhof in litt.). Jaschhof and 

Jaschhof 2003 studied the fauna of New Zealand, noting the features of the group there and described a new tribe 

peculiar to New Zealand. 

Tribe Acoenoniini 

Acoenonia 4 

*Proacoenonia 1 

Tribe Aprionini 

Aprionus 119 

Mycophila 7 

Tekomyia 1 

426

Yukawamyia 1 

Tribe Bryomyiini 

Bryomyia 7 

Heterogenella 13 

Skuhraviana 1 

Tribe Campylomyzini 

Ansifera 5 

Campylomyza 38 

Corinthomyia 1 

Cretocordylomyia 1 

Excrescentia 1 

Micropteromyia 1 

Neurolyga 29 

Tribe Catochini 

Catarete 2 

Catocha 9 

Cretocatocha 1 

Forbesomyia 1 

Neocatocha 1 

Tritozyga 2 

Tribe Micromyini 

Anodontoceras 3 

Micromya 11 

Monardia 53 

Subg. Antennardia 3 

Subg. Monardia 24 

Subg. Trichopteromyia 8 

Subg. Xylopriona 14 

Unplaced to subg. 4 

Polyardis 10 

Pseudoperomyia 29 

Tribe Peromyiini 

Gagnea 1 

Peromyia 150 

Tribe Pteridomyiini. Ref.: Jaschhof 2003b: 109. 

*Pseudomonardia 11 

*Pteridomyia 2 

Tribe Strobliellini 

Amedia 1 

427

*Amediella 1 

Eleniella 1 

Eltxo 1 

Groveriella 2 

Strobliella 2 

Micromyinae unplaced to tribe 

*Berestella 1 

Calospatha 1 

Ipomyia 1 

Psadaria 1 

Stenospatha 1 

Termitomastus 1 

Tricampylomyza 1 

Trichelospatha 1 

Tricolpodia 1 

*Vicemyia 1 

Subfamily Porricondylinae 

This diverse subfamily of 131 genera and 731 species, including 14 unplaced species, is especially poorly 

known outside the Palearctic Region. As an example, only nine species are known from the Neotropical Region, all 

separately described between 1877 and 1936 (Gagné 1994a). I cite this fact only to indicate how little is known there; I 

remain willing to wait forever for a judicious and careful taxonomist to study them properly. 

Porricondylines are among the oldest known fossils of Cecidomyiidae. Arillo & Nel (2000) described a 

species from Lower Cretaceous Spanish amber and Gagné (1977a) named others from Upper Cretaceous Canadian 

amber. Many porricondyline fossils have been named from Baltic amber (Meunier 1904) and Ukrainian amber 

(Fedotova and Perkovsky 2004, 2005, 2007a, 2008a, and Perkovsky and Fedotova 2004, 2008a, 2008b). 

This subfamily is paraphyletic with regard to the subfamily Cecidomyiinae (Panelius 1965, Jaschhof & 

Jaschhof 2009a). Most species are associated with fungi growing in detritus and beneath tree bark. Some may be 

regularly associated with particular niches, e.g., the species of Asynapta and Camptomyia that live in conifer cones. The 

small but ancient tribe Heteropezini is particularly noteworthy for its paedogenetic habit. Its species exhibit remarkable 

modifications in anatomical structures and reduction in size (Wyatt 1967). 

The most recent revision of Porricondylinae is that by Panelius (1965) for the European fauna. A review by Parnell 

(1971) of the Nearctic species described by Felt placed most of the known Nearctic species into Panelius's 

classification. Wyatt (1967) revised the Heteropezini with attention to all stages. Möhn (1955a) and Mamaev and 

Krivosheina (1968) presented keys and illustrations to larvae of the subfamily. Plakidas (1999) published a key to 

larvae of selected genera for the Nearctic Region. Keys to genera of Nearctic adults can be found in Gagné 1981b and 

to Palearctic adults by Skuhravá 1997. Spungis (1985, 1987, 1991, 1992, 2006b) reviewed the European fauna of 

certain tribes. Mamaev (2001c) differentiated Holoneurini from Porricondylini; I use his classification with the caution 

that Holoneurus is still based on a species without a type specimen. 

Since 2004, Fedotova (2004j) and Fedotova and Sidorenko (2005e, 2007b, 2007c, 2007d, 2008a, 2008b) 

described 76 species of porricondylines from Primorskiï Territory, Far East Russia, and Fedotova (2004d, 2004e), 

Fedotova and Perkovsky (2008a, 2008b, 208c), and Perkovsky and Fedotova (2008) described 17 species of this 

subfamily from Ukrainian amber. The number of species from the Primorskiï Territory, added to the 54 species 

previously described from there, gives a taste of the great diversity that is to be found in a small circumscribed area. Of 

the 33 genera and 4 subgenera described by Fedotova and Fedotova with Sidorenko to classify the species that they 

described from that area, all except four are monotypic. The practice of describing a separate genus for nearly every 

second species found does not give the category of genus very much to do. Further, five genera based only on females 

are presumably synonyms of other genera based on males. Inasmuch as females of the supertribe are so poorly known 

and lack the striking characters found in male genitalia, these genera based only on females offer no help in clarifying 

428

429 

relationships. Fedotova and Perkovsky, separately and together, and Perkovsky and Fedotova described six new genera 

for their species from Ukrainian amber. Five of them are monotypic, and four of those five are characterized only from 

females. Considering that the fossils appear to be of such poor quality and the papers describing them of such little help 

in recognizing the species in the future, there seems little purpose in giving names even to the species. A critique of this 

work on Ukrainian amber was made by Jaschhof (2007) to which Fedotova and Perkovsky (2007b) responded. 

I note here that Yang (1998) proposed the new subfamily Thaumacecidomyiinae in which to include his new 

genus Thaumacecidomyia. The genus is listed here under the tribe Diallactini under which the superfluous subfamily 

name is subsumed. 

Tribe Asynaptini 

Asycola 1 

Asynapta 46 

Camptomyia 73 

Colomyia 5 

Epicola 1 

*Eudokimyia 1 

Feltomyina 1 

*Larimyia 1 

*Lobopedosis 1 

Parasynapta 2 

Pseudocamptomyia 3 

Stackelbergiella 1 

*Tenepidosis 1 

Tribe Bryocryptini 

Bryocrypta 16 

*Locusepidosis 1 

Tribe Diallactini 

Cretohaplusia 1 

Diallactia 7 

Ferovisenda 1 

Gynapteromyia 1 

Haplusia 19 

*Thaumacecidomyia 1 

Wyattella 3 

Tribe Dicerurini 

*Adsumyia 1 

Bulbepidosis 1 

*Crustepidosis 1 

Dicerura 28 

Hilversidia 1 

*Nebulepidosis 2 

Neotetraneuromyia 2 

Neurepidosis 6 

Paratetraneuromyia 2 

Pterepidosis 2 

*Salpistepidosis 1

Synepidosis 2 

Tetraneuromyia 5 

Ubinomyia 1 

*Velafacera 1 

Tribe Dirhizini 

Dirhiza 5 

Tribe Heteropezini 

Perkovsky and Fedotova (2008b) raised this tribe to superfamily level and placed it under the Cecidomyiinae 

(specifically the Lasiopterinae, raised in that paper and earlier in Fedotova 2000b to subfamily rank). These 

phylogenetically unacceptable proposals are not followed here. The reason given in support of that placement is the 

authors' assertion that the first and second flagellomeres are fused in Heteropezini. I surveyed a number of 

Heteropezini, including specimens reliably determined as Heteropeza pygmaea, the type species of Heteropeza, for 

this character and found the opposite, that those flagellomeres are not fused. The Heteropezini more properly appear to 

be an early offshoot from the base of Porricondylinae that never developed ring-shaped sensoria (Panelius 1965). 

Perkovsky and Fedotova (2008b) consider Heteropezina a subgenus of Heteropeza, but that suggestion is also not 

followed here. 

Brittenia 1 

Cretomiastor 1 

*Electroxylomyia 1 

Frirenia 2 

Henria 3 

Heteropeza 3 

Heteropezina 3 

Heteropezula 1 

Leptosyna 4 

Miastor 3 

Monodicrana 1 

Neostenoptera 2 

Nikandria 1 

*Tutkowskia 1 

*Ventosagloria 3 

Tribe Holoneurini 

Cassidoides 8 

Coccopsilis 5 

*Cognitepidosis 1 

Cryptoneurus 3 

Dallaiella 1 

*Gratomyia 1 

Holoneurus 13 

*Melicepidosis 1 

*Putepidosis 1 

*Pyxicoccopsis 1 

Schistoneurus 3 

*Volnococcopsis 1 

Tribe Porricondylini 

430

*Alocolpodia 1 

*Ancorepidosis 1 

*Anisepidosis 1 

Arctepidosis 2 

*Armatepidosis 2 

Basicondyla 1 

Cedrocrypta 2 

Claspettomyia 30 

Colpodia 6 

Dendrepidosis 4 

Dentepidosis 1 

Desertepidosis 2 

*Divellepidosis 16 

*Dolerepidosis 1 

Furcepidosis 1 

*Glomepidosis 1 

*Incrementistyla 1 

Isocolpodia 3 

Lamellepidosis 1 

*Ledomepidosis 1 

Monepidosis 8 

Monocolpodia 1 

Neocolpodia 2 

*Obryzepidosis 1 

Paracolpodia 3 

Parepidosis 14 

Porricondyla 95 

Prosepidosis 1 

Pseudepidosis 6 

*Recessepidosis 1 

Sclerepidosis 1 

Sensepidosis 1 

*Seychellepidosis 1 

Stomatocolpodia 5 

Trichepidosis 1 

*Ustinepidosis 1 

*Yukawaepidosis 1 

*Zaitzeviola 1 

*Zephyrepidosis 2 

Tribe Solntseviini 

Grisepidosis 2 

Solntsevia 3 

*Tersepidosis 1 

*Vulnepidosis 1 

Tribe Winnertziini 

Clinorhytis 1 

Cretowinnertzia 1 

Cryptoxylomyia 2 

431

*Eraepidosis 1 

*Fertepidosis 1 

Kronomyia 6 

Lobowinnertzia 8 

Parwinnertzia 2 

Rhipidoxylomyia 14 

*Shevchenkia 1 

*Vasiliola 1 

Winnertzia 113 

Unplaced to tribe 

Clinophaena 1 

Harpomyia 1 

Jamalepidosis 3 

Meunieria 1 

Misocosmus 1 

Palaeospaniocera 1 

Pallidepidosis 3 

Rabindrodiplosis 2 

Spungisomyia 1 

Sylvenomyia 2 

Zatsepinomyia 2 

Subfamily Cecidomyiinae 

Cecidomyiinae comprise by far the largest subfamily of gall midges, with 595 genera and 4763 known species. The 

number of species includes 270 that are unplaced to genus but not 117 nomina dubia. The subfamily contains all plant 

feeding and predaceous cecidomyiids, as well as groups that have retained or reacquired the ancient fungal diet. Fossil 

species of this subfamily are common in amber but not yet known earlier than the Cenozoic (Gagné 1973b). The 

subfamily is monophyletic, characterized by the following synapomorphies: the ventrally separated gonocoxites; the 

reduced number of flagellomeres from the primitive 14 found in most other Sciaroidea, including the subfamily 

Porricondylinae, to 13, 12, or fewer, or an augmented number of more than 12 that is usually irregular within a species; 

and the presence of only two dorsal papillae on the eighth larval abdominal segment. 

Some general and indispensable references for any systematic study of this subfamily are: Barnes 1946a,b, 

1949, 1951a,b, 1952, 1955, and Nijveldt 1969, a series of books on gall midges of economic importance; Fedotova & 

Kovalev 2001, keys to Far East Russian taxa with many illustrations; Gagné 1989b on Nearctic plant-feeding gall 

midges; Gagné 1994a on Neotropical gall midges; Kieffer 1913i, an outdated but indispensable world monograph; 

Mamaev and Krivosheina 1968 on larvae of Palearctic genera; Möhn 1955a on larvae of Palearctic genera; Skuhravá 

1997 on adults of Palearctic genera, with a large collection of illustrations from various works; and Yukawa 1971, a 

revision of Japanese gall midges. Comprehensive gall books treating all gall makers can also be helpful, such as, Buhr 

1964, 1965, a two-volume work covering the galls of central and northern Europe, Yukawa and Masuda 1996 on the 

galls of Japan, Redfern et al. 2002 on British plant galls, Russo 2006 on the galls of California and other western states, 

and Roskam 2009 on the galls of The Netherlands. 

Cecidomyiinae are divided here into four supertribes: Brachineuridi, Cecidomyiidi, Lasiopteridi, and 

Stomatosematidi (Gagné 1994a). These are treated separately below in alphabetical order with the reasons justifying 

their uniqueness. 

In the 2004 catalog, family group names were not cataloged because they had been treated in Sabrosky 1999. 

New ones proposed since then, however, are covered here under the pertinent supertribes. Most of them are not 

employed here because they are not based on comprehensive studies. 

Supertribe Brachineuridi 

432

433 

This cosmopolitan but so far species-poor supertribe of 82 species is characterized by the following 

synapomorphies: antennae with 10 flagellomeres; tarsal claws narrow, curved near midlength; male seventh and eighth 

abdominal tergites each reduced to a strongly sclerotized, linear band and corresponding sternites shorter and more 

weakly developed than preceding sternites; and short, sclerotized, free, glabrous gonocoxite lobes, each with one to 

three terminal setae (Gagné 1994a). In this supertribe the female postabdomen is not protrusible and its cerci are 

separate. Larvae of some species have been associated with insect and spider remains and adults have been reared from 

rotting wood. The supertribe has no tribal arrangement. Fedotova & Perkovsky (2008) placed two genera, Kovaleviola 

and Spungisiola, from Ukrainian amber with the Brachineuridi, but these, from what one can judge from the 

illustrations, could belong to the Lasiopteridi instead. 

*Acinacistyla 1 

*Alatostyla 1 

Brachineura 30 

Brachyneurina 5 

Chrybaneura 1 

*Cingola 1 

Coccidomyia 1 

*Compositola 1 

*Effusomyia 1 

Epimyia 2 

Epimyiella 1 

*Kovaleviola 1 

*Nodalistyla 1 

Rhizomyia 30 

*Spungisiola 1 

*Stabiliola 1 

*Undoneura 2 

Volsatiola 1 

Supertribe Cecidomyiidi 

This supertribe and the Lasiopteridi comprise the plant-feeding gall midges. In addition to plant feeders, the 

Cecidomyiidi, with 2395 species, include many fungivores as well as all the predaceous cecidomyiids. The predaceous 

habit appears to have evolved several times within the supertribe. Unique characteristics of Cecidomyiidi include the 

following: the presence on the adult head of a vertical process bearing two large, anteriorly pointed setae (sometimes 

secondarily lost); the development, which may be separately lost, of the male antennal flagellomeres into two distinct 

nodes, each node with one or two separate, many-looped circumfila; and the loss of setae on the ventral papillae of the 

larval eighth abdominal segment. Males of Asphondyliini and some other Cecidomyiidi have secondarily lost the 

antennal modification and have gynecoid male flagellomeres. Most genera of Cecidomyiidi can be grouped at least 

tentatively among 11 tribes, the Anadiplosini, Aphidoletini, Asphondyliini, Cecidomyiini, Centrodiplosini, 

Clinodiplosini, Hormomyiini, Karshomyiini, Lestodiplosini, Lopesiini, and Mycodiplosini. These tribes are listed in 

alphabetical order below. Many genera are not yet grouped into formal tribes and that extensive list is given at the end 

of the supertribe. Useful references for Cecidomyiidi in addition to those mentioned under Cecidomyiinae are: Gagné 

1973c (synopsis of Nearctic genera) and Harris 1966 (review of selected genera with illustrations). 

The following new tribal and subtribal names for Cecidomyiidi have recently been proposed: 

Acanthacidiplosini Fedotova 2003e: 59, based on Acanthacidiplosis Fedotova; Epidiplosina Fedotova and Sidorenko 

2009a: 14, based on Epidiplosis Felt; Filidiplosini Fedotova and Sidorenko 2009a: 26, based on Filidiplosis Fedotova; 

Foveoladiplosina Fedotova and Sidorenko 2009a: 10, based on Foveoladiplosis Fedotova; Gigantodiplosini Fedotova 

and Sidorenko 2009a: 44, based on Gigantodiplosis Fedotova; Glomerulidiplosina Fedotova and Sidorenko 2009d: 

254, based on Glomerulidiplosis Fedotova 2003; Heterostylidiplosini Fedotova 2004g: 104, based on 

Heterostylidiplosis Fedotova; Inclinatidiplosini Fedotova & Sidorenko 2009d: 223, based on Inclinatidiplosis Fedotova 

2004h; Karshomyiini Fedotova and Sidorenko 2009a: 6, based on Karshomyia Felt; Mamaeviina Fedotova 2009: 13, 

based on Mamaevia Skuhravá; Marikovskidiplosina Fedotova and Sidorenko 2009b: 12, based on Marikovskidiplosis

434 

Fedotova and Sidorenko; Magadiplosina Fedotova 2009: 24, based on Magadiplosis Fedotova and Sidorenko; 

Pedetentiina Fedotova and Sidorenko 2009d: 247, based on Pedetentia Fedotova and Sidorenko 2009; Quadridiplosini 

Fedotova 2003e: 65, based on Quadridiplosis Fedotova; Ruidadiplosina Fedotova 2009: 20, based on Ruidadiplosis 

Fedotova and Sidorenko; Tollereadastrini Fedotova and Sidorenko 2009a: 17, based on Tollereadastra Fedotova; and 

Undadiplosina Fedotova 2009: 15, based on Undadiplosis Fedotova and Sidorenko. Fedotova and Sidorenko (2009a: 5) 

also raise Coquillettomyiini Mamaev 1968 to the level of supertribe. Of those tribes listed, only the tribe Karshomyiini 

is used here, but not with all of the genera Fedotova and Sidorenko proposed to include in it. Their tribal level 

descriptions are based mainly on seemingly superficial and non-exclusive characters of the male genitalia and do not 

take into account other important characters. As an example, Fedotova and Sidorenko 2009a propose the new tribe 

Filidiplosini to include Filidiplosis, Exiguidiplosis, Sicituradastra, Flexipidiplosis, and Rotadiplosis. The abdominal 

terga described for these genera are so different, from entire to laterally dissected, from strongly pigmented to weakly 

so, and the tarsal claws as drawn are of such a variety, that the few superficial similarities outlined for the male genitalia 

of these genera do not make a convincing argument that the tribe comprises a monophyletic assemblage. 

Tribe Anadiplosini 

This small, little known Neotropical tribe of eight species of plant-feeders has a number of apomorphies, chief 

of which are: the placement of wing vein Rs beyond midlength of R1; the slightly curved base of M; the shortened 

tibiae relative to the tarsi; and pointed, ventroapical prolongations (spurs) on the first tarsomeres. Ref.: Gagné 1994a 

(descr.). 

Alexomyia 1 

Anadiplosis 4 

Machaeriobia 1 

Scopodiplosis 1 

Uleella 1 

Tribe Aphidoletini 

Of this cosmopolitan tribe of 54 species, at least the species of Aphidoletes are aphid predators, and the others, 

mostly known from adults taken from rotting wood or caught in flight, are known to be or are possibly also predaceous 

on other insects. Synapomorphies include the elongate and irregular circumfila of the male flagellomeres, the middle 

one bandlike in most genera, and the modification of the larval terminal segment into two lateral lobes on which the 

terminal papillae are situated. Refs: Harris 1966 (review), 1973 (review, illustrations), 1981a (Bremia, illustrations). At 

least the following genera belong here. 

Allobremia 2 

*Anulidiplosis 1 

Aphidoletes 4 

Bremia 19 

*Daedalidiplosis 1 

*Falciformidiplosis 2 

Farquharsonia 2 

Heterobremia 1 

Homobremia 1 

*Lanidiplosis 1 

*Mirabilidiplosis 14 

Monobremia 4 

*Ollaediplosis 1 

Spilodiplosis 1 

Tribe Asphondyliini

435 

The Asphondyliini, all gallmakers, form a well-circumscribed, monophyletic group of 505 species, sharing 

unique characters of the adult postabdomen. These are the distinctive female seventh sternite that is much longer than 

the preceding sternite; the strongly sclerotized, wide, and laterally notched female eighth tergite; the ventrally 

lengthened gonocoxites; and the dorsally instead of apically disposed, short and compact gonostyli. The tribe was for a 

time treated as a supertribe equivalent to the Cecidomyiidi, but Gagné (1994a) placed it in the supertribe Cecidomyiidi 

because the antennae have 12 flagellomeres that were probably primitively binodal in the male, as they are today in 

Bruggmannia. The tribe is divided here into two subtribes, the Asphondyliina and the Schizomyiina following Gagné 

1994a. Refs.: Möhn 1961b (rev.); Tokuda & Yukawa 2007. 

Subtribe Asphondyliina 

The greatest diversity so far known of this subtribe of 343 species lies in the Neotropics. Asphondyliina have a 

pointed, ventroapical prolongation on the first tarsomere of each leg, pseudocerci at the posterior end of the female 

eighth tergite, the basal half of the ovipositor with numerous, parallel ridges lined with setulae, the apical half aciculate 

except in Daphnephila, the male genitalia usually without mediobasal gonocoxite lobes and the denticles of the 

gonostyli all or mostly fused into one or two solid teeth, the pupal head with strongly modified, sawtooth-like antennal 

bases and frontal projections used in cutting through gall tissue before ecdysis, and the pupal third through eighth 

abdominal segments with strong dorsal, cuticular spines that offer purchase against the gall tissue when exiting the gall, 

and the larva with a reduced number of lateral papillae, setose ventral papillae, and reduction in size of the anal segment 

with the anus placed subdorsally. 

Apoasphondylia 2 

Asphondylia 289 

Bruggmanniella 10 

Daphnephila 9 

*Dimocarpomyia 1 

Hemiasphondylia 2 

Heterasphondylia 1 

Houardiella 3 

*Illiciomyia 1 

Parazalepidota 1 

Perasphondylia 2 

Probruggmanniella 1 

Pseudasphondylia 10 

Pumilomyia 1 

Rhoasphondylia 3 

Sciasphondylia 1 

Tavaresomyia 1 

Tetrasphondylia 1 

Zalepidota 4 

Subtribe Schizomyiina 

Except for the cosmopolitan Schizomyia this subtribe of 162 species shows its greatest diversity outside the 

temperate Holarctic Region. Schizomyiina are probably paraphyletic to the Asphondyliina. They have no apical spur on 

the first tarsomere of each leg, no large lobes (occasionally a very small one) at the posterior end of the female eighth 

tergite, a primitively soft ovipositor that in most genera is elongate and tapered, and unfused denticles on the gonostyli. 

The pupa may or may not have prominent antennal bases and usually has projections of the frons, and small cuticular 

spines cover only the anterior part of the third to eighth abdominal segments. The larva has six or fewer lateral papillae, 

and a ventral anus. 

Ameliomyia 1 

Anasphondylia 1

Aposchizomyia 6 

Asphoxenomyia 1 

Asteralobia 12 

Brethesia 1 

Bruggmannia 19 

Burseromyia 1 

Eocincticornia 2 

Kiefferia 1 

Kochiomyia 7 

Luzonomyia 1 

Macroporpa 2 

Metasphondylia 1 

Okriomyia 2 

*Oxycephalomyia 1 

*Parametasphondylia 1 

Paraschizomyia 1 

Pisoniamyia 3 

Pisphondylia 2 

Placochela 2 

Polystepha 24 

Proasphondylia 3 

Pseudokochiomyia 5 

Schizandrobia 2 

Schizomyia 53 

Stephomyia 7 

Tribe Cecidomyiini 

This large tribe of 583 known species is exclusively plant-feeding. The distinguishing character is the makeup 

of the terminal papillae of the larvae: one pair is corniform and recurved and the remaining three pairs have short setae. 

If this modification evolved only once, then Procontarinia should fit and is listed here. Most Cecidomyiini have simple, 

untoothed tarsal claws and all are curved beyond midlength. 

Acacidiplosis 13 

*Acerovesiculomyia 1 

Anisostephus 2 

*Caryadiplosis 2 

Caryomyia 56 

Cecidomyia 20 

Chamaediplosis 2 

Contarinia 312 

Efferatodiplosis 1 

Garugadiplosis 1 

Gobidiplosis 1 

Halodiplosis 99 

Lobopteromyia 1 

Macrodiplosis 14 

Paradiplosis 4 

*Parkiamyia 1 

Phyllodiplosis 1 

Pinyonia 1 

Plemeliella 1 

Procontarinia 15 

436

Prodiplosis 12 

Sequoiomyia 2 

Sphaerodiplosis 1 

Stenodiplosis 10 

Taxodiomyia 3 

Thecodiplosis 6 

Zeuxidiplosis 1 

Tribe Centrodiplosini 

This tribe, known only from three monotypic Neotropical genera feeding on Solanaceae, is unique for the 

strongly sclerotized, aciculate ovipositor with fused cerci. 

Centrodiplosis 1 

Cystodiplosis 1 

Jorgensenia 1 

Tribe Clinodiplosini 

Clinodiplosini are a cosmopolitan tribe of 193 species. Many are evidently mycophagous but, some, especially 

in the Western Hemisphere, are phytophagous and free-living or form galls. Some are predaceous on other gall midges. 

The main synapomorphies of the tribe are the obtuse mediobasal lobe of the male gonocoxite and the distinctive 

terminal segment of the larva, with typically three pairs of corniform papillae, one pair smaller than the other two, and 

one pair of setiform papillae. Fedotova and Sidorenko (2009) elevate this tribe to supertribal status, with the new tribe 

Inclinatidiplosini, itself divided into three new subtribes: Inclinatidiplosina, Pedentiina, and Glomerulidiplosina. These 

new divisions are superfluous, based as they ultimately are on species that could as well be housed in Clinodiplosis and 

may be synonymous with existing species of Clinodiplosis. Ref.: Skuhravá 1973 (review of Clinodiplosis). 

*Admareversa 1 

Ametrodiplosis 20 

Anthodiplosis 1 

*Auritadiplosis 1 

Autodiplosis 1 

Chauliodontomyia 2 

Chelobremia 1 

Clinodiplosis 102 

*Dextermana 1 

*Emarginatodiplosis 1 

*Glomerulidiplosis 1 

Hadrobremia 1 

Houardodiplosis 1 

Hyperdiplosis 4 

*Hypoprodiplosis 1 

Iatrophobia 1 

*Inclinatidiplosis 2 

Liodiplosis 3 

Parallelodiplosis 22 

*Pedetentia 3 

Piranea 1 

*Samaradiplosis 1 

Schismatodiplosis 1 

*Signasicca 1 

Sitodiplosis 4 

437

*Spiculatidiplosis 4 

*Transversalidiplosis 1 

*Tuguridiplosis 10 

Tribe Hormomyiini 

The 51 species in a single genus make up a very diverse and rich group of sedge-feeding gall midges. A large 

number of generic names has been coined for members of this tribe, and some of those names may be reemployed one 

day when the tribe undergoes a revision. Their most conspicuous apomorphy is the long thorax that projects anteriorly 

over the head. Some species in this group have an augmented number of flagellomeres, up to 25 in one species. 

Planetella 51 

Tribe Karshomyiini 

This tribe comprises 63 spp. Fedotova and Sidorenko (2009a: 6) proposed the new tribe Karshomyiini (also as 

Karshomyini) for Karshomyia and several other genera, but not all the genera placed there by Fedotova and Sidorenko 

(ibid.) are placed here. Those gathered here share some similarities of male genitalia but mainly the partially divided 

abdominal terga. Placing them here is an alternative to leaving them undistinguished in the large pool of unplaced 

genera of Cecidomyiidi. These genera should be reevaluated as a group. 

*Fimbriatodiplosis 1 

*Foveoladiplosis 1 

*Heterostylidiplosis 4 

Karshomyia 49 

*Ogdodiplosis 1 

*Opinatodiplosis 1 

*Palmatidiplosis 3 

*Prorsodiplosis 2 

*Undadiplosis 1 

Tribe Lestodiplosini 

This is a large tribe of 341 species. Larvae are obligatory predators of insects, mites, and other arthropods, 

mostly free-ranging, and have elongate antennae and various means to effect locomotion, e.g., pseudopods in 

Lestodiplosis and some other genera. A few species are endoparasitoids of Hemiptera. The species are among the 

smallest cecidomyiids. Males show a tendency for, alternatively, mediobasal gonocoxite lobes or a variously setose or 

lobed ventral surface of the hypoproct. Ref.: Baylac 1987c (descr. larvae). 

Adiplosis 1 

Androdiplosis 1 

Arthrocnodax 49 

Cartodiplosis 1 

Chanchudiplosis 1 

Dentifibula 8 

*Dilatodiplosis 2 

Diplecus 1 

Endaphis 6 

Endopsylla 2 

Epidiplosis 13 

Feltiella 10 

Heterocontarinia 1 

Lamprodiplosis 1 

438

Lestodiplosis 182 

Mohwadiplosis 1 

*Montosidiplosis 1 

Odontodiplosis 14 

Pectinodiplosis 1 

*Spirodiplosis 1 

Stictobremia 1 

*Stigmodiplosis 3 

Tanaodiplosis 1 

Tessaradiplosis 1 

Thripsobremia 3 

*Tingidoletes 1 

*Tonsidiplosis 4 

Trilobia 1 

Triommatomyia 2 

Trisopsis 26 

Uncinulella 1 

Tribe Lopesiini 

This tribe of 27 species, all plant feeders, appears mainly tropical in distribution. Its separation as a tribe is 

still uncertain due to the small known sample and its still indistinct morphological limits that include: general 

robustness; reductions in size and circumfila of male flagellomeres; extra teeth on the tarsal claws of some species; and 

short, sensory ventral setae on the female cerci. Known larvae have diminutive terminal papillae with two or more pairs 

bearing tiny corniform setae. Ref.: Gagné 1994a (descr.). In addition to the genera listed directly below, Anabremia and 

Tricholaba may possibly belong here. 

Allodiplosis 1 

Austrolopesia 1 

Cordiamyia 1 

Lopesia 18 

Lopesiella 1 

Myrciamyia 3 

Obolodiplosis 1 

Rochadiplosis 1 

Tribe Mycodiplosini 

This tribe contains 123 mycophagous or presumably mycophagous species. The one known apomorphy 

unifying this group is the makeup of the larval terminal papillae: two pairs are short-corniform and two setose (cf. 

Möhn 1955a, 1955b). Ref.: Holz 1970 (rev.). 

*Aequidiplosis 3 

Buhromyiella 1 

Camptodiplosis 2 

Coquillettomyia 42 

*Faristodiplosis 13 

*Fissuratidiplosis 1 

*Galeidiplosis 2 

*Gybbosidiplosis 1 

*Laxadiplosis 1 

Mycodiplosis 49 

Neoisodiplosis 2 

439

Neomycodiplosis 1 

*Sagittidiplosis 2 

*Sharmadiplosis 1 

*Versusfacera 2 

Unplaced Genera of Cecidomyiidi 

Some of the following 184 genera with a total of 444 species may be referable to available tribes. Others, as 

they become better known, will sort into natural groups. For example, Diadiplosis, Megommata, and Triommata are 

evidently closely related. 

*Acanthacidiplosis 1 

Acodiplosis 2 

*Aculeatodiplosis 1 

Afrodiplosis 1 

Ampelosucta 1 

Anabremia 6 

Anasphodiplosis 1 

Andirodiplosis 1 

*Anisodiplosis 1 

Antichiridium 2 

Apagodiplosis 1 

Aplecus 1 

Apodiplosis 1 

Arrabiadaeamyia 1 

Aschistonyx 1 

Asphotrophia 3 

Astrodiplosis 1 

Athidiplosis 2 

Atopodiplosis 1 

Atrichosema 1 

Austrodiplosis 1 

Baeodiplosis 1 

*Bicornidiplosis 1 

Blaesodiplosis 3 

Blastodiplosis 2 

Brachydiplosis 1 

Braueriella 1 

Brephometra 1 

Buhriella 1 

Bungomyia 1 

Cacoplecus 2 

Callitridiplosis 1 

Calodiplosis 1 

*Calyptradiplosis 1 

*Carinatidiplosis 1 

Chaetodiplosis 2 

Chrysodiplosis 2 

Cleitodiplosis 1 

Clusiamyia 2 

Coccomyza 1 

Coelodiplosis 1 

Collula 2 

440

Compsodiplosis 1 

Conodiplosis 1 

Contodiplosis 3 

Cordylodiplosis 1 

Ctenodactylomyia 2 

Ctenodiplosis 2 

Dactylodiplosis 3 

Delodiplosis 1 

Diadiplosis 28 

Dichaetia 1 

Dichodiplosis 3 

Dicrodiplosis 13 

Diodaulus 2 

Diplodontomyia 2 

Diplosiola 1 

*Dissimilidiplosis 1 

Drisina 1 

Echinella 2 

Enallodiplosis 1 

Eohormomyia 1 

Epihormomyia 2 

Etsuhoa 10 

Eucalyptodiplosis 3 

Eumerosema 1 

Exiguidiplosis 1 

*Filidiplosis 4 

*Flexidiplosis 1 

Frauenfeldiella 1 

Geromyia 3 

Giardomyia 12 

*Gigantodiplosis 1 

Gladiodiplosis 1 

Glenodiplosis 1 

Gnesiodiplosis 1 

Gongrodiplosis 1 

Gynandrobremia 5 

Gynodiplosis 1 

Haplodiplosis 1 

Harmandiola 14 

Heliodiplosis 1 

Holobremia 3 

Holodiplosis 2 

Horidiplosis 7 

Huradiplosis 1 

Hygrodiplosis 1 

Hypodiplosis 1 

Inulomyia 1 

Kamptodiplosis 2 

Kimadiplosis 1 

*Kitella 1 

Kronodiplosis 1 

*Lianodiplosis 1 

Lasiodiplosis 1 

441

Lepidobremia 1 

Lepidodiplosis 3 

Liebeliola 1 

*Ligulodiplosis 1 

Loewiola 3 

Lophodiplosis 5 

*Magadiplosis 1 

Mamaevia 2 

Manilkaramyia 1 

*Marikovskidiplosis 1 

Massalongia 4 

Megaulus 1 

Megommata 6 

Mesodiplosis 1 

Meterdiplosis 2 

Microdiplosis 1 

Microplecus 1 

Mikaniadiplosis 1 

Mitodiplosis 1 

Moehniella 1 

Monarthropalpus 1 

Monodiplosis 1 

Mycetodiplosis 1 

Mycocecis 1 

Myricomyia 1 

Nanodiplosis 1 

Neobaezomyia 1 

Octodiplosis 1 

Odinadiplosis 2 

Oligoxenomyia 1 

Olpodiplosis 1 

Oribremia 1 

Orseolia 28 

Orthodiplosis 1 

Ouradiplosis 1 

Pachydiplosis 1 

Perodiplosis 1 

Pilodiplosis 1 

Pipaldiplosis 1 

Pitydiplosis 1 

Platydiplosis 1 

Plecophorus 1 

Plectrodiplosis 1 

Plesiodiplosis 1 

Plutodiplosis 2 

*Poridiplosis 1 

Proterodiplosis 1 

Pruthidiplosis 1 

Punarnavomyia 1 

Putoniella 2 

*Quadridiplosis 4 

Raodiplosis 1 

Resseliella 53 

442

*Rotadiplosis 1 

*Ruidadiplosis 1 

Schizodiplosis 1 

*Setodiplosis 1 

*Sicituradastra 1 

Silvestriola 14 

Skusemyia 1 

*Stenohypodiplosis 1 

Stephodiplosis 2 

Streptodiplosis 1 

Stroblophila 1 

Styraxdiplosis 2 

Tetradiplosis 1 

Thaumadiplosis 1 

Thorodiplosis 1 

Thurauia 2 

Tokiwadiplosis 1 

*Tollereadastra 2 

Tribremia 1 

Tricholaba 5 

Tricontarinia 2 

Trigonodiplosis 1 

Trilobomyia 1 

Trilobophora 1 

Triommata 2 

Trissodiplosis 1 

Trogodiplosis 1 

Tropidiplosis 1 

*Ussuridiplosis 1 

Vaccinidiplosis 1 

Xenasphondylia 1 

Xenhormomyia 2 

Xenodiplosis 2 

Xylodiplosis 5 

Youngomyia 6 

Supertribe Lasiopteridi 

This supertribe of 1993 species is made up mostly of plant feeders, most of them gall makers. Some are 

inquilines, including all Trotteria spp. and Camptoneuromyiini, probably most Macrolabis spp., and a few Alycaulini, 

Dasineurini, and Lasiopterini. Inquilines often affect their particular gall maker host by crowding or other form of 

competition, but some are merely successors in the galls. Species of the tribe Ledomyiini are probably all 

mycophagous. Two synapomorphies that delimit the Lasiopteridi are the weaker aedeagus relative to that of the 

Cecidomyiidi and the pair of mediobasal gonocoxite lobes that laterally clasp and appear to lend support to the 

aedeagus. 

Since early in the twentieth century most Lasiopteridi have been grouped into two tribes, Dasineurini and 

Oligotrophini. These were separated according to whether the tarsal claws were toothed or not. This classification was 

most recently reflected in Skuhravá 1997, but teeth of tarsal claws have evidently been lost many times so that their 

absence is too superficial for exclusive use (Sylvén & Tástas-Duque 1993). 

From the mass of Lasiopteridi, several well-circumscribed, monophyletic tribes, namely Alycaulini, 

Camptoneuromyiini, Lasiopterini, Ledomyiini, and Trotteriini, are distinct and easily separated. A few others are 

narrowly defined here because the tribal names are available and applicable, e.g. Oligotrophini, Poomyini, and 

Rhopalomyiini. I have assigned the remaining genera to two large categories, a large tenuous tribe, the Dasineurini, and 

443

444 

one other large group of unplaced genera that do not appear to fit into any of the tribes or are too poorly known to 

place. The tribes are treated below in alphabetical order with the unplaced genera at the end. 

Perkovsky and Fedotova 2008b raise the Lasiopteridi to subfamily rank but that proposal is not followed here. 

Tribe Alycaulini 

This is a large group of 210 described species, all restricted to the Americas. Most species are primary plant 

feeders causing simple swellings on stems, tendrils, or petioles of various plants, or living in flowers or achenes of 

Asteraceae. Most appear to be associated with symbiotic fungi, at least some of the time, but some evidently are not. A 

few are inquilines in galls of other cecidomyiids. Except for the speciose and diverse Neolasioptera, remaining genera 

are small but distinct and are each restricted to a particular host group. 

Unique, derived characters of Alycaulini involve unique modifications of the ovipositor and male posterior 

abdominal tergites. Dorsal, longitudinal ridges on the basal half of the ovipositor are enlarged and interrupted to form 

rows of sclerotized and rounded bumps that may be used to abrade plant tissue when laying eggs. The posterior margin 

of the male seventh tergite and occasionally the eighth is sclerotized and setose. Reduction in these two tergites begins 

at the anterior border, whereas in Lasiopterini and Dasineurini, any reduction proceeds from the posterior border of the 

tergites. The cerci of at least one Alycaulus sp. are separate, indicating the apparently primitive state for the tribe. 

Characters that are presumably separately derived and shared with some other tribes of Lasiopteridi are the short 

antennae of both sexes, the short R5 that is also straight and almost parallel with C, and the robust, toothed claws. The 

Alycaulini are superficially quite similar to the Lasiopterini, but substantial differences in reductions of the posterior 

abdominal tergites show the Alycaulini to be unique among Lasiopteridi and the Lasiopterini to be more similar to the 

Dasineurini. Most alycauline larvae show adaptations for cutting a tunnel through wood before pupation, including the 

cylindrical body shape and strongly sclerotized spatula. The lateral papillae are reduced to four or fewer per side and 

terminal papillae may be reduced from eight to six, four, or two. The robust spatula and reduced lateral papillae occur 

also in other groups that live in woody stem galls, including the Lasiopterini and the Oligotrophini of this supertribe and 

the genus Bruggmanniella of the Cecidomyiidi. 

Alycaulus 3 

Asteromyia 9 

Astictoneura 1 

Atolasioptera 1 

Baccharomyia 4 

Brachylasioptera 2 

Calamomyia 19 

Chilophaga 4 

*Couridiplosis 1 

Edestochilus 1 

Edestosperma 1 

Epilasioptera 1 

Geraldesia 3 

Lobolasioptera 1 

Marilasioptera 1 

Meunieriella 21 

Neolasioptera 134 

Protaplonyx 1 

Smilasioptera 1 

Xipholasioptera 1 

Tribe Camptoneuromyiini 

Camptoneuromyiini are a small tribe of 29 species in the Americas and 3 in the Palearctic. Most genera and 

species are exclusively Neotropical. The Palearctic genus Verbasciola, for which the tribe Verbasciolini Fedotova 

(2004b) was erected, does not appear to differ from Camptoneuromyia. At least all Neotropical species are inquilines in

445 

galls formed by other Cecidomyiinae. The only character unique to this tribe is in the relative length of the terminal 

papillae, with one pair shorter than the remaining three. Other, non-exclusive distinguishing characters of the tribe are 

the short R5 wing vein, which is only one-third to one-half the length of the wing and usually bowed but sometimes 

nearly straight, the short antennae of both sexes that show only slight sexual dimorphism, and the larval clove-shaped 

spatula with six lateral papillae on each side. Refs: Möhn 1975 (descr.), Gagné 1994a (review). 

Camptoneuromyia 15 

Copinolasioptera 1 

Dialeria 1 

Domolasioptera 7 

Incolasioptera 1 

Salvalasioptera 1 

*Verbasciola 3 

Tribe Dasineurini 

This tribe is a large, heterogeneous assemblage of 870 primary plant feeders. Most cause simple or complex 

plant galls, but some live freely in flower heads, and some others are inquilines. I place here those Lasiopteridi that 

share most of the following characteristics (with some exceptions): male flagellomeres with definite necks; wing with 

R5 joining C at least slightly before the wing apex; a protrusible ovipositor; female eighth abdominal tergite as long or 

longer than seventh and tending to divide longitudinally; and male genitalia with mediobasal gonocoxite lobes that 

gradually taper from base to apex and are nearly as long as the aedeagus. None of these characters is unique to this 

tribe, especially with regard to Camptoneuromyiini and Lasiopterini, which may be offshoots of Dasineurini, but the 

genera placed here show some general resemblance to one another. Some genera that are unplaced to tribe may also fit 

here. 

Sylvén & Tástas-Duque (1993), in their review of characters differentiating females of Dasineura and 

relatives, show how little basis there is for separating some of the included genera, although many good adaptive 

characters of the terminalia occur that relate to behavior, particularly oviposition. Certain characters, e.g., length of 

ovipositor and the point where the R5 and C wing vein meet, which once were given considerable importance occur so 

many times in genera that can be shown to be unrelated that they cannot be used alone as criteria for classification. 

Sylvén (1975), in his study of relationships between habits and integumental structures of larvae, showed that similar 

larval characters are evidently correlated with larval habits, indicating that similar losses and new structures likely 

appeared separately many times. 

When one canvasses individual species of Dasineura, one can presently find as much interspecific variety as 

there is among some genera. Dasineura spp. that occur on Fabaceae, e.g., D. mariae on Astragalus, have the fused 

female cerci pointed apically. The same is true for the type species of Wachtliella on Medicago and for all eight species 

of Fabomyia spp., all on Fabaceae, incl. Medicago and Astragalus. An analogous situation exists with Dasineurini on 

Brassicaceae: species of Brassomyia, Syrenomyia, and Gephyraulus, one of Bayeriola, two of Janetiella, and 28 

species of Dasineura occur on that host family. In fact, the type species of five of those genera live on Brassicaceae. 

The Dasineurini on Fabaceae and those on Brassicaceae may possibly one day prove to be separate monophyletic 

groups. No groupings within Dasineurini can be satisfactory until the tribe is revised in a comprehensive way. Refs: 

Sylvén 1975, Sylvén & Tástas-Duque 1993. 

*Acumyia 1 

Alliomyia 1 

Androsacemyia 1 

Arcivena 1 

Arnoldiola 15 

Astragalomyia 1 

Austrolauthia 1 

Bayeriola 4 

Brassomyia 2 

Bremiola 15

Coniophora 5 

Cotoneasteromyia 1 

Dasineura 466 

Dasyneuriola 5 

Dryomyia 4 

Euphorbomyia 4 

Fabomyia 9 

*Galatellomyia 1 

Geocrypta 6 

*Geomyia 1 

Gephyraulus 11 

Geraniomyia 1 

Giraudiella 2 

Gypsophyllomyia 1 

Hartigiola 2 

*Istodiomyia 1 

Jaapiella 79 

Janetia 7 

Janetiella 24 

Kaltenbachiola 3 

Labiatomyia 6 

Lathyromyza 4 

Lygocecis 4 

Macrolabis 60 

Neomikiella 3 

Polygonomyia 2 

Potentillomyia 1 

Rabdophaga 82 

Rondaniola 1 

Scorzoneromyia 1 

Spiraeanthomyia 2 

Spiromyia 1 

Spurgia 8 

Syrenomyia 1 

Tavolgomyia 1 

Vitisiella 3 

Wachtliella 9 

Xerephedromyia 3 

Zygiobia 2 

Tribe Lasiopterini 

Lasiopterini are a large, chiefly Old World tribe. Of the 330 known species, only 37 are native to North 

America and none is known from South America. Most Lasiopterini form simple swellings on buds, stems, petioles, 

and leaf ribs of various plants, but some live in galls or galleries originally formed by other insects. Most are associated 

with a symbiotic fungus. 

The synapomorphic characters of the Lasiopterini lie in the ovipositor: the first is the presence on each side, 

just beyond the eighth tergite, of a group of enlarged and flattened setae; the second and third apomorphies are the 

hooked setae and the glabrous, sclerotized area on the fused cerci. The tribe superficially resembles Alycaulini in the 

short antennae that are similar in both sexes, the short R5, the thick covering of scales, including the white spot at the 

juncture of R5 and the otherwise dark costa, and generally similar larvae, particularly in the presence of no more than 

four thoracic lateral papillae on each side of the spatula. Males of Lasiopterini differ from those of Alycaulini in the 

manner of reduction in length of the male seventh tergite, which loses sclerotization along its posterior margin, unlike 

446

in Alycaulini in which reduction proceeds from the anterior margin. 

Actilasioptera 6 

*Afrolasioptera 1 

Aplonyx 1 

Artemisiomyia 1 

Atraphaxiola 1 

Baldratia 38 

Careopalpis 15 

*Dactylasioptera 2 

Exopsyllomyia 3 

Hybolasioptera 2 

Isolasioptera 2 

Izeniola 3 

Lasioptera 128 

Microlasioptera 1 

Ozirhincus 5 

Primofavilla 3 

Stefaniella 9 

Stefaniola 109 

Tribe Ledomyiini 

This cosmopolitan tribe of 64 species is extremely diverse and presents a small universe of genitalic variety. 

While several problematic genera are used for the Palearctic fauna, all of the New World fauna is included at the 

moment in Ledomyia and the catchall genus Lauthia. Lauthia (sensu lato) is commonly found in Mexican (Gagné 

1973) and Dominican amber. Some species are associated with fungal fruiting bodies or rotting plant tissue, some with 

insects or insect remains. Several Nearctic species have been reared from xylem vessels in logs. This tribe has one 

synapomorphy, the largely divided mediobasal gonocoxite lobes: one part is setulose, the other, usually longer and 

glabrous except for apical setae. The antenna has 8-10 or a regular number of 12 sexually dimorphic flagellomeres. The 

head of a few species has a dorsal occipital prominence, unlike the remainder of the Lasiopteridi. The tarsal claws are 

thin, toothed on the forelegs, and toothed or simple on the mid- and hindlegs. Wing venation is variable, with R5 much 

shorter to longer than the wing. The ovipositor is short to elongate, but the cerci are separate. Refs: Gagné 1985a 

(review), Rock and Jackson 1985 (biology). 

Isogynandromyia 1 

Lauthia 30 

Ledomyia 26 

Ledomyiella 5 

Plesiolauthia 1 

Prolauthia 1 

Tribe Oligotrophini 

I treat this tribe extremely narrowly, limiting it to two genera and 15 species associated with Cupressaceae, all 

with simple tarsal claws with very long empodia. The latter is a possibly synapomorphic character here but one shared 

also with almost all conifer-feeding gall midges of both Lasiopteridi and Cecidomyiidi. Separate female cerci as well as 

fused cerci are present in this tribe and even in the same genus, Oligotrophus. The presence of both states here and in 

other tribes indicates that the development of fused cerci is a superficial occurrence that has come about many times. 

Oligotrophus 10 

Walshomyia 5 

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Tribe Poomyini 

I tentatively place the monotypic Epicalamus here with the large genus Mayetiola. The latter, with 30 species, 

is distinctive for the presence of a puparium, which Epicalamus does not have, but the larvae of both genera live in 

stems of grasses, have a curved aedeagus attended by short, blunt mediobasal gonocoxite lobes, and the female eighth 

tergite is no longer than the seventh. This tribe is wholly western Palearctic except for one native, east coast Nearctic 

species of Mayetiola and one immigrant to North America and New Zealand, the Hessian fly. Ref.: Ertel 1975 (rev.). 

Epicalamus 1 

Mayetiola 29 

Tribe Rhopalomyiini 

As limited here, this tribe is made up of one large cosmopolitan genus of 278 species. Nearly all occur on 

Asteraceae and most form complex galls. Perhaps some genera among those in the unplaced category may be placed 

here as separate genera or as species of Rhopalomyia. Possible synapomorphies include: sexual dimorphism with male 

legs and wings longer than those of the female, although this character does not seem to be universal; the female eighth 

tergite, when elongated, is not longitudinally divided into separate tergites as is the case in Dasineurini; and the 

mediobasal gonocoxite lobes are short and blunt. Refs.: Fedotova 2001f,g,h (rev. Palearctic), Jones et al. 1983 (fauna 

on Artemisia, Nearctic), Dorchin et al. 2009 (fauna on Solidago, Nearctic). 

Rhopalomyia 278 

Tribe Trotteriini 

This tribe consists of one cosmopolitan genus of 23 species. All are inquilines in galls of other cecidomyiids, 

chiefly Asphondyliini. Adults are extremely modified so that the tribe's closest relative among Lasiopteridi is not 

apparent. Although in certain adult characters the genus superficially resembles Lasiopterini, larvae are more 

plesiomorphic and the ovipositor resembles more closely that of Dasineurini than of Lasiopterini. Synapomorphies 

include: the elongate postocciput that covers the dorsal part of the head so that the eyes and antennae are displaced 

ventrad; the elongate antennal scape and pronotum; and the greatly enlarged hind femora. Ref.: Möhn 1975. 

Trotteria 23 

Unplaced genera of Lasiopteridi 

The following 59 genera with their 144 species cannot be placed at this time into any of the named tribes of 

Lasiopteridi. Reasons for this include: the genera have anatomical specializations that mask relationships; the genera are 

simply too poorly known for placement; or, more likely, the tribal classification is inadequate to the task of grouping 

the genera. Skuhravá 1997 erected the tribe Mikiolini, not used here, for Mikiola, one of the genera listed below, based 

on non-exclusive characters. 

Acericecis 4 

Amerhapha 1 

Angeiomyia 1 

Apiomyia 1 

Berberisomyia 1 

Blastomyia 1 

Celticecis 11 

Craneiobia 2 

Cystiphora 7 

Didymomyia 2 

Ephedromyia 6 

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Eugeniamyia 1 

Ficiomyia 1 

*Gliaspilota 1 

*Guareamyia 1 

Guarephila 1 

Guignonia 1 

Haplopalpus 1 

Hasegawaia 1 

Herbomyia 1 

Iteomyia 5 

*Jorgenseniella 1 

Litchiomyia 1 

Lyciomyia 1 

Masakimyia 1 

Mayteniella 1 

Mikiola 5 

Mikomya 1 

Myrciariamyia 3 

Neodasyneuriola 1 

Neomitranthella 1 

*Normanbyomyia 1 

Novocalmonia 3 

Ochnephila 1 

Paulliniamyia 1 

Pernettyella 1 

Phegomyia 2 

Physemocecis 2 

Planodiplosis 1 

Primavera 1 

Procystiphora 4 

Promikiola 1 

Psectrosema 26 

Pseudomikiola 1 

Radulella 1 

Riveraella 1 

Rosomyia 3 

Sackenomyia 5 

Scheueria 2 

Semudobia 7 

Sophoromyia 2 

Spartiomyia 1 

*Sphaeramyia 1 

Sterrhaulus 1 

Taxomyia 1 

Trichoperrisia 1 

Trigonomyia 3 

Uleia 1 

Xyloperrisia 2 

Supertribe Stomatosematidi 

This small supertribe of 23 species consists of two cosmopolitan genera. Nothing is known of the larval 

biology but the adult of one of the Didactylomyia spp. is a kleptoparasite on spider webs. Distinguishing anatomical 

449

450 

characters are: the presence of 13 flagellomeres, the last with an elongate, narrow, apical process, and a strong Rs wing 

vein. A synapomorphy is the distinct mediobasal gonocoxite lobe, usually divided into two distinct parts approximately 

equal in length but different in structure. A primitive character found only in Didactylomyia but lost in Stomatosema 

and presumably separately lost in the remainder of Cecidomyiinae is the two-segmented female cercus. 

Didactylomyia 3 

Stomatosema 20 

APPENDIX 2: ABBREVIATIONS 

A - adult 

anat. - anatomy 

assoc. - associated 

Aut. Reg. - Autonomous Region 

betw. - between 

biol. - biology 

Biol. - Biological 

Co. - County 

Coll. - Collection 

des. - designation 

descr. - description 

Distr. - District 

E - east 

emend. - emendation 

Eur. - European 

for., For. - forest, Forest 

G - gall 

hom. - homonym 

HT - holotype 

I. - Island 

ICZN - International Commission on Zoological Nomenclature 

illus. - illustrations 

imm. - immature; used only for stage of type 

immigr. - immigrant 

introd. - introduced 

Is - Islands 

jun. - junior 

jun. prim. hom. - junior primary homonym 

jun. sec. hom. - junior secondary homonym 

km - kilometer, kilometers 

Kr. - Kreis, a German political district 

L - larva 

L. - Lake 

LT - lectotype 

mi - mile, miles 

missp. - misspelling 

mon. - monotypy 

Mt - Mount 

mtn, Mtn - mountain, Mountain 

mts, Mts - mountains, Mountains 

N - north 

Natl - National 

NE - northeast

n. g. - new genus 

nr - near 

n. sp. - new species 

NT - neotype 

NW - northwest 

occ. - occurrence 

orig. - originally 

orig. des. - original designation 

P - pupa 

Pk - Park 

poss. - possibly 

Pref. - Prefecture 

Prefs - Prefectures 

preocc. - preoccupied 

prob. - probably 

Prov. - Province 

Provs - Provinces 

publ'n - publication 

R. - Rio, River, or Rivière 

rd - road 

redescr. - redescription 

Ref. - Reference 

Refs - References 

Reg. - Region 

Res. - Reserve or Reservation 

rev. - revision 

RR - railroad 

S - south 

SE - southeast 

s.l. - sensu lato 

s.s. - sensu stricto 

sp. - species 

spp. - species (plural) 

St - Saint (in type localities only) 

ST - syntypes 

St. - State 

Stn - Station 

subg. - subgenus 

subs. des. - subsequent designation 

subsp. - subspecies 

SW - southwest 

syn. - synonym 

T - type 

Terr. - Territory 

undet. - undetermined 

Univ. - University 

unjust. emend. - unjustified emendation 

unk. - unknown 

unnec. - unnecessary 

var. - variety 

vic. - vicinity 

W - west 

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APPENDIX 3: ABBREVIATIONS FOR TYPE DEPOSITORIES 

Four-letter abbreviations for type depositories are mainly from Arnett 1993, but some are original here. No 

new abbreviations were coined for collections of authors who described fewer than three species. 

AMNH - USA: American Museum of Natural History, New York, New York 

ANIC - Australia: Australian National Collection of Insects, Canberra 

ASAY - Armenia: Zoological Institute, Erevan 

BAUC - China: Agricultural University, Beijing 

BIZS - Bulgaria: Institute of Zoology, Bulgarian Academy of Sciences, Sofia 

BMNH - United Kingdom: The Natural History Museum, London 

BPBM - USA: Bernice P. Bishop Museum, Honolulu, Hawaii 

CASC - USA: California Academy of Sciences, San Francisco 

CMNZ - New Zealand: Canterbury Museum, Christchurch 

CNCI - Canada: Canadian National Collection, Ottawa, Ontario 

CSIA - India: Cecidological Society of India, Allahabad 

CUIC - USA: Cornell University, Ithaca, New York 

DEIC - Germany: Senckenberg Deutsches Entomologisches Institut, Müncheberg (formerly Eberswalde) 

DESH - India: Deshpande Collection, Science College, Nanded, Maharashtra 

DGOT - Germany: Institut und Museum für Geologie und Paläontologie der Universität Göttingen 

DPBA - Argentina: INTA, Departamento de Patologia Vegetal, Castellar, Buenos Aires 

DZNM - India: Post Graduate Department of Zoology, Science College, Nanded, Maharashtra 

DZSM - Germany: Zoologisches Staatssammlung München 

ELMU - India: Entomological Laboratory, Maharashtra University, Parbhani 

ELSU - Japan: Entomological Laboratory, Saikyo University, Kyoto 

ETHZ - Switzerland: Entomologisches Institut, Eidgenössische Technische Hochschule, Zurich 

ETKI - India: Entomology Laboratory, Central Research Institute, Trivandrum, Kerala 

FBUL - Latvia: Faculty of Biology, University of Latvia, Riga 

FRDD - India: Forest Research Institute, Dehra Dun, Andhra Pradesh 

FRNZ - New Zealand: Forest Research Institute, Rotorua 

FSCA - USA: Florida Collection of Arthropods, Gainesville 

HDOA - USA: Hawaii Department of Agriculture, Honolulu, Hawaii 

HFES - Japan: Hokkaido Government Forestry Experiment Station, Toyohira, Sapporo 

HYNM - Japan: Museum of Nature and Human Activities, Hyogo 

IBSP - Brazil: Instituto Biologico, Secretaria da Agricultura, São Paulo 

IEME - Russia, A.N. Severtzov Institute for Evolution, Morphology, and Ecology of Animals, Moscow 

IGGB - Romania: Institute of Geology and Geophysics, Bucharest 

INHS - USA: Illinois Natural History Survey, Urbana 

INPC - India: Indian Agriculture Research Institute, New Delhi 

INPC - India: Hayara National PUSA Collection 

INRA - France: Institut National de Recherche Agronomique, Versailles 

IRCW - USA: University of Wisconsin, Madison 

ISUI - USA: Iowa State University, Ames 

IZAV - Venezuela: Instituto de Zoologia Agricola, Universidad Central de Venezuela, Maracay 

KOVC - Russia: O.V. Kovalev Collection 

KUEC - Japan: Kyushu University, Fukuoka 

KUIC - Japan: Kagoshima University, Kagoshima 

LASS - Latvia: Institute of Biology, Latvian Academy of Sciences, Salaspils 

LSUK - United Kingdom: Linnaean Society, London 

MACN - Argentina: Museo Argentino de Ciencias Naturales, Buenos Aires 

MAMC - Russia: B.M. Mamaev Collection, Moscow 

MANC - India: M.S. Mani Collection 

MARC - Kazakhstan: P.I. Marikovskij Collection, Alma-Ata 

452

MCNA - Spain: Museum of Natural Sciences of Alava, Vitoria 

MCZC - USA: Museum of Comparative Zoology, Cambridge, Massachusetts 

MECN - Ecuador: Museo Ecuatoriano de Ciencias Naturales, Quito 

MLPA - Argentina: Universidad National de La Plata 

MNHN - France: Muséum National d'Histoire Naturelle, Paris 

MNNC - Chile: Museo National de Historia Natural, Santiago 

MRAC - Belgium: Musée Royal de l'Afrique Centrale, Tervuren 

MZCR - Costa Rica: Museo de Zoologia, Universidad de Costa Rica, San Jose 

MZHF - Finland: Zoological Museum, Helsinki 

MZLU - Sweden: Lund University 

MZSP - Brazil: Museu de Zoologia da Universidade de São Paulo 

MZUF - Italy: Museo Zoologico "La Specola," Florence 

NEAC - Romania: Neacs� Collection, Bucharest, Romania 

NFRI - Norway: Forestry Research Institute, Ås 

NHMW - Austria: Naturhistorisches Museum, Vienna 

NHRS - Sweden: Swedish Museum of Natural History, Stockholm 

NHSM - USA: Springfield Natural History Museum, Massachusetts 

NKUM - China: Nankai University, Tianjin 

NMBA - Austria: Naturhistorisches Museum der Benediktiner-Abtei, Admont 

NMID - Ireland: National Museum, Dublin 

NMNS - Taiwan: National Museum of Natural Science, Taichung 

NYSM - USA: New York State Museum, Albany. 

NZAC - New Zealand: New Zealand Arthropod Collection, Auckland 

OSUC - USA: Ohio State University, Columbus 

PGCA - India: P. Grover Collection, Allahabad 

QBUM - Brazil: Museu National, Quinta de Boa Vista, São Cristóvão, Rio de Janeiro 

QMBA - Australia: Queensland Museum, Brisbane 

RCMU - India: S.N. Rao Collection, Marathwada University 

RCNU - India: S.N. Rao Collection, Nagpur University 

RMNH - Netherlands: Natural History Museum, Leiden 

RSME - United Kingdom: Royal Museums of Scotland, National Museum of Scotland, Edinburgh 

SAMA - Australia: South Australian Museum, Adelaide 

SANV - South Africa: South African National Collection of Insects, Pretoria 

SAUT - China: Shandong Agricultural University, Tainan 

SCUC - China: South China University Collection, Guangzhou, Guangdong Province 

SEMC - USA: University of Kansas, Lawrence 

SIUC - USA: Illinois Natural History Survey, Champaign, IL 

SJCA - St. John's College, Agra, Andhra Pradesh 

SKCR - Skuhravá Collection, National Museum, Prague, Czech Republic 

SKRP - Skrzypczy�ska Collection, Cracow 

SMNS - Germany: Staatliches Museum für Naturkunde, Stuttgart 

SMTD - Germany: Senckenberg Naturhistorisches Sammlungen, Museum für Tierkunde, Dresden 

STZM - Poland: Zoological Museum, Szezecin 

SZAC - Poland: Szadziewski Collection, Warsaw 

TAUI - Israel: Zoological Museum, Tel Aviv University 

TAVC - Tavares Collection, in SMNS or MNHN (Houard Collection) 

TNSM - Tridentino Natural Sciences Museum, Trento, Italy 

UASK - Ukraine: Schmalhausen Institute of Zoology, Kiev 

UASS - Uzbekistan: Zoological Institute, Uzbek Academy of Sciences, Samarkand 

UMEC - USA: University of Massachusetts, Amherst 

UMSP - USA: University of Minnesota, St. Paul 

UNSA - Republic of South Africa: Natal Museum, Pietermaritzburg 

453

USNM - USA: Washington, DC, Smithsonian Institution 

UTAT - USA: University of Texas, Austin 

ZFMK - Germany: Zoologisches Forschunginstitut und Museum Alexander Koenig, Bonn 

ZIMG - Germany: Zoologisches Institut und Museum, Ernst Moritz Arndt- Universität, Greifswald 

ZISB - Bulgaria: Institute of Zoology, Sofia 

ZKAS - Kazakhstan: Zoological Institute, Kazakh Academy of Sciences, Alma-Ata 

ZMAN - Netherlands: Zoological Museum, Amsterdam 

ZMAS - Russia: Zoological Institute, Russian Academy of Sciences, St. Petersburg 

ZMHB - Germany: Museum für Naturkunde, Humboldt-Universität, Berlin 

ZMKG - Germany: Zoologisches Museum, Kiel 

ZMMU - Russia: Zoological Museum, Moscow State University, Moscow 

ZMPA - Poland: Polish Academy of Science, Warsaw 

ZMUC - Denmark: Zoology Museum, Copenhagen University 

ZSCI - India: National Zoological Collection, Calcutta 

APPENDIX 4: NOTES ON SOME AUTHORS' COLLECTIONS AND TYPES 

The following notes should help interested persons locate several authors' types and collections or at least to 

learn what happened to them. Subject authors are listed in alphabetical order. 

Bremi, J.J. Most of this author's extant collection is in the Entomologisches Institut, Eidgenössische 

Technische Hochschule in Zurich. Box #66 of the entomological collection contains three rows of Bremi's pinned 

cecidomyiid adults (Gagné 1990b). A few of Bremi's pinned specimens are also in the Rondani collection in Florence 

(Gagné and Solinas 1996). Material of Bremi's species that were based only on galls and larvae appears to be lost. 

Felt, E.P. The complete slide collection consisting mostly of type material is on indefinite loan from the New 

York State Museum in Albany, New York, to the Systematic Entomology Laboratory of the U S. Department of 

Agriculture in Washington, DC. The type deposition is given here as the New York State Museum to reflect the true 

ownership. Specimens on pins and a collection of galls remain in the New York State Museum in Albany. 

Jaschhof, J. As of January, 2009, the Jaschhof collection of Cecidomyiidae, formerly deposited in the 

Zoological Institute and Museum, University of Greifswald, is housed in the Senckenberg Deutsches Entomologisches 

Institut, Müncheberg. 

Kieffer, J.-J. Most of Kieffer's types can be considered as certainly lost. No collection remains in Bitche 

where Kieffer lived (Gagné 1994a), but some collections elsewhere, not all yet surveyed, contain Kieffer material. 

Among his type specimens that still exist are those in The Natural History Museum in London and the Muséum 

National d'Histoire Naturelle in Paris of at least some of his Seychelles and East African species. Others may have been 

returned following their description to the National Zoological Collection in Calcutta. Still other Kieffer specimens, not 

necessarily types although some are labelled as such, are in the Felt Collection (Gagné 1994a) and could possibly serve 

for designation as lectotypes or neotypes. Galls of some species are present in the Houard Collection in the Muséum 

National d'Histoire Naturelle in Paris. In addition, Kieffer occasionally based species on published references to galls 

by Rübsaamen and Löw, among others. Such specimens might someday be found in the original authors' home 

museums. Not being certain which of Kieffer's types are definitely lost, I have indicated most of them as lost followed 

by a question mark. 

Loew, H. Horn & Kahle (1937) indicate that part of Loew's collection of European Cecidomyiidae is in Berlin 

and that part of his cecidomyiid collection is in Vienna, but I have not searched for them. Some of Loew's sciarid types 

have been found in the Museum of Natural History, Humboldt University, Berlin (Jaschhof in litt.), so there is a chance 

the types of some of his European gall midge species are also there. I have indicated the type deposition of Loew's 

species as "ZMHB?," except for the types of two North American species that reside in the Museum of Comparative 

Zoology, Cambridge, Massachusetts. 

Mamaev, B.M. Most of Mamaev's and co-authors' types are in the Zoological Museum of Moscow State 

University (Krivosheina & Ozerov 2006), others are in the Museum of Nature and Human Activities, Hyogo, Japan 

(Yagi & Sawada 2000). Both museums claim the holotype of several Mamaev holotypes. Others are still listed here as 

in the Mamaev collection or the A.N. Severtzov Institute for Evolution, Morphology, and Ecology of Animals, 

Moscow. Information with the original descriptions of those species indicated that the types were earlier intended for 

454

455 

other depositories, chiefly the A.N. Severtzov Institute for Evolution, Morphology, and Ecology of Animals in Moscow 

and the Zoological Institute, Russian Academy of Sciences in Leningrad, now St. Petersburg. 

Marshall, P. The types of Marshall's 33 species from New Zealand are presumed lost (Miller 1950). 

*Meade, R.H. Possible type specimens reside in the Zoology Department of the University of Leeds, UK. 

According to Adrian Pont (in litt.), these are in a drawer of specimens donated by the Yorkshire dipterist Cheetham 

comprising reared Cecidomyiidae from Meade. 

Meigen, J.W. Some of the Meigen types are in the Muséum National d'Histoire Naturelle in Paris (L. Matile, 

in litt.) and are so indicated. The remainder are marked as lost. 

Meunier, F. Meunier described many Baltic amber fossils from the Königsburg collection. These were 

probably destroyed during World War II. A few survive that had been transferred to the Institut und Museum für 

Geologie und Paläontologie der Universität Göttingen before the war and are listed only by genus in Ritzkowski 1990. 

Meunier species in those genera are listed here as questionably in Göttingen. The remainder are listed as lost. 

Möhn, E. This collection is in the Staatliches Museum für Naturkunde, Stuttgart. Most specimens are stored 

in a cold room in vials containing alcohol mixed with glycerin and arranged by collection number. The numbers refer to 

Möhn's notebooks, which contain full collection data and are also kept at the Museum. To find specimens of a 

particular species, one first looks up the species name in a book to find the numbers under which the species is stored. 

Numbered vials are located, the specimens retrieved, and temporarily mounted in glycerin under a cover slip on a glass 

slide. After they have been studied the specimens are replaced in the vials. Additionally, some specimens are still on 

temporary slide mounts (N. Dorchin, in litt.). These have suffered some deterioration due to the evaporation of the 

glycerin and movement of coveer slips. 

Rübsaamen, E.H. All or most of the Rübsaamen collection is in the Museum für Naturkunde, Humboldt- 

Universität, Berlin. To judge from a sample of his specimens that I saw on loan elsewhere (Gagné 1994a) and that has 

since been returned to Berlin, a part is in alcohol and another is mounted on slides. At least some of this latter portion is 

mounted on microscope slides in glycerin that has dried with time so that the cover slips are easily separated, sometimes 

accidentally so, from the glass slides. Also, at least some material is labelled only by collection number. There is a great 

need to curate this large, very important collection, and, of course, this task should be undertaken or at least overseen 

not by a disinterested person but by someone with considerable experience in the care and study of gall midges. 

Skuse, F.A.A. I have examined the remaining Skuse collection of cecidomyiids in the Australian National 

Collection of Insects in Canberra. Many of the types of Skuse's species are missing or in fragments and their names are 

placed with nomina dubia here. The types of the remaining Porricondylinae and Cecidomyiinae have since been loaned 

for study to Dr. P. Kolesik in Adelaide, South Australia, and Jaschhof has a paper in preparation on the Lestremiinae 

and Micromyinae. 

Spungis, V. Some of his types originally stated as deposited in the Institute of Biology, Latvian Academy of 

Sciences, Salaspils, were listed in Krivosheina and Ozerov (2006) as in the Zoological Museum of Moscow State 

University. The changes have been made in this revision of Gagné (2004). 

Stelter, H. Place of type deposition in some of Stelter's papers is given as the Deutsches Entomologisches 

Institut in Eberswalde, but a number of those types are actually in the Staatliches Museum für Tierkunde Dresden, as is 

most of the Stelter Collection (U. Kallweit, in litt.). These are reported here as being in Dresden, the actual place of 

residence. 

Tavares, J.S. Most of Tavares's collection is lost, but examples of galls of some species are in the Houard gall 

collection in the Muséum National d'Histoire Naturelle in Paris. Another small part of Tavares's collection was found 

by E. Möhn (pers. comm.) in an attic of the Instituto Nun'Alvres, Caldas de Saude, Portugal and are presently in the 

care of the Staatliches Museum für Naturkunde in Stuttgart. 

Walker, F. The Walker types are in The Natural History Museum in London, but most are in poor condition 

and unidentifiable (K.M. Harris in litt.). Many are listed here under "Unplaced Species" as they are at present 

unassigned to genus, but some will doubtless later be relegated to nomina dubia. 

Winnertz, J. Most of Winnertz's types were destroyed in Bonn during World War II. F.W. Edwards saw the 

Winnertz collection in the 1930s and published notes on some types. A few Winnertz specimens, types possibly among 

them, can be found in the Muséum National d'Histoire Naturelle in Paris. Pont (in litt., 7/9/03): with Palearctic Diptera 

in Oxford University Museum of Natural History, he found a number of dry mounted Cecidomyiidae in the Verrall- 

Collin collection. He quotes from a letter to Verrall by Osten Sacken that OS intends to give them to Verrall. And in 

2003, Pont found additional Winnertz material in Zetterstedt's "Diptera Exotica" collection in Lund, Sweden.

LITERATURE CITED 

References added since Gagné 2004 are marked with an asterisk. References are arranged chronologically for 

each author as well as I could determine. Parts of series of papers published in separate journal numbers are cited 

individually. 

Translated titles of papers that appeared in Slavic languages or Japanese are in brackets, even if originally 

accompanied by a title translated into English, as were many of the Russian papers. The original translated titles of 

some Russian and Japanese articles were sometimes not in proper English and with misspellings due to anonymous 

translators, so I offer in some instances a more correct title that I think reflects better on the contents and authors. Titles 

in other languages are as originally given, including misspellings, except that generic and specific names are always 

italicized even if not italicized in the original. Russian words in titles are transliterated according to the American 

National Standards Institute system. 

Journal titles are spelled out in full except for some of the few items that I did not see. Issue number of 

volumes is given only when pages are not numbered consecutively through the entire volume. Plate numbers are 

included only when in addition to article pagination. 

Abbass, A.K. 1986. A new species of grass midge (Dipt., Cecidomyiidae) infesting the inflorescence of Phalaris 

arundinacea L. in Britain. The Entomologist's Monthly Magazine 122: 65-71. 

Aczél, M. Not seen. 1944. Die Linsenblütengallmücke (Contarinia lentis n. sp.), ein neuer Linsenschädling. 

Növényegészségügyi Evy. 1941-1943: 127-171. 

Agarwal, R.A. 1956. A new genus of Indian Itonididae: Kittada coimbatorensis, gen. et sp. nov. Current Science 25: 

129-130. 

Agassiz, L. 1847. Nomenclatoris Zoologici. Soloduri [=Solothurn, Switzerland]. viii & 393 pp. 

Agassiz, L. and H. Loew. 1846. Nomina systematica generum dipterorum. [Pt. 4], [vi] 42 pp. In Agassiz, L. 

Nomenclator zoologicus. Fasc. 9/10: Titulum et praefationem operis, Mollusca, Lepidoptera, Strepsiptera, 

Diptera, Myriapoda, Thysanura, Thysanoptera, Suctoria, Epizoa et Arachnidas. Soloduri [=Solothurn, 

Switzerland]. 

Ahlberg, O. 1939. Diarthronomyia chrysanthemi nom. nov. (hypogaea Felt nec Löw). Entomologisk Tidskrift 60: 

274-278. 

Aldrich, J.M. 1905. A catalogue of North American Diptera. Smithsonian Institution Miscellaneous Collections 46 (2): 

1-680. 

Alexander, C.P. 1924. Undescribed species of nematocerous Diptera from North America and Japan. Insecutor Inscitiae 

Menstruus 12: 81-84. 

Alexander, C.P. 1936. The maximum number of antennal segments in the Order Diptera, with the description of a new 

genus of Cecidomyiidae. Bulletin of the Brooklyn Entomological Society 31: 12-14. 

Alexander, C.P. 1937. Change of name in Diptera. Bulletin of the Brooklyn Entomological Society 32: 60. 

Alexander, C.P. 1959. Undescribed species of nematocerous Diptera. Part VI. Bulletin of the Brooklyn Entomological 

Society 54: 37-43. 

*Almeida, F.V.M., J.C. Santos, F.A.O. Silveira and G.W. Fernandes. 2006. Distribution and frequency of galls induced 

by Anisodiplosis waltheriae Maia (Diptera: Cecidomyiidae) on the invasive plant Waltheria indica L. 

(Sterculiaceae). Neotropical Entomology 35: 435-439. 

Anagnostopoulos, P.T. 1929. Blastomanie de l'amandier. Revue de Pathologie Végétale et d'Entomologie Agricole 16: 

168-177. 

Anderson, J.A.T. 1935. The description, bionomics, morphology, and anatomy of a new Dryomyia (Cecidomyidae, 

Diptera). Proceedings of the Zoological Society of London 1935: 421-430. 

*Anfora, G. 2005. Pp. 96-98. In Anfora, G., N. Isidoro, De Cristofaro, A. and C. Ioriatti. Description of Macrolabis 

mali sp. nov. (Diptera Cecidomyiidae), a new inquiline gall midge species from galls of Dasineura mali on 

apple in Italy. Bulletin of Insectology 58: 95-99. 

*Anfora, G., N. Isidoro, De Cristofaro, A. and C. Ioriatti. 2005. Description of Macrolabis mali sp. nov. (Diptera 

Cecidomyiidae), a new inquiline gall midge species from galls of Dasineura mali on apple in Italy. Bulletin of 

Insectology 58: 95-99. 

*Angelini, B. 1831. Degli insetti nocivi all'ulivo nella provincia Veronese. Memoria I. Atti e Memoriae della 

456

457 

Accademia di agricoltura, scienze e lettere di Verona 12: 295-323. 

Angelo, A.C. and V.C. Maia. 1999. Dasineura gigantea sp. n. (Diptera, Cecidomyiidae) associada a Psidium 

cattleianum Sabine (Myrtaceae) no Brasil. Revista Brasileira de Zoologia 16: 191-195. 

Anonymous. 1844. Memorie per servire alla ditterologia italiana di C. Ròndani. Isis (Oken) 1844: 449-452. 

Anonymous. 1903. Notes on insect pests from the Entomological Section, Indian Museum. Indian Museum Notes 5: 

103-202, 212, pls. VII-XX. 

Anonymous. 1926. Two corrections. The Canadian Entomologist 58: 240. 

Anonymous. 1929. Insecta. Zoological Record 65: 322. 

Appleby, J.E. and R.B. Neiswander. 1965. Oligotrophus apicis sp. n., a midge injurious to junipers; with key to species 

of Oligotrophus found in the United States. The Ohio Journal of Science 65: 166-175. 

Arillo, A. and A. Nel. 2000. Two new fossil cecidomyiids flies from the Lower Cretaceous amber of Alava (Spain) 

(Diptera, Cecidomyiidae). Bulletin de la Société Entomologique de France 105: 285-288. 

Arnett, R.H., G.A. Samuelson, and G.M. Nishida. 1993. The Insect and Spider Collections of the World. 2nd ed. 

Sandhill Crane Press, Inc. Gainesville, Florida. 310 pp. 

*Ashe, P. and J.P. O'Connor. 2005. Re-discovery and the first Irish record of Arthrocnodax fraxinella (Meade) 

(Diptera: Cecidomyiidae) reared from the cauliflower ash gall. The Entomologist's Monthly Magazine 141: 

139-142. 

Bach, M. 1859. Ueber gewisse Veränderungen und Auswüchse an verschiedenen Pflanzentheilen, welche durch den 

Einfluss von Insekten bewirkt werden. Natur und Offenbarung 5: 250-263. 

Bagnall, R.S. 1922. A cecidomyid, Perrisia harrisoni, nom. nov. The Entomologist's Monthly Magazine 58: 183-184. 

Bakhshi, M. and P. Grover. 1976. On the study of cocoons of some gall-midges (Cecidomyiidae: Diptera). Cecidologia 

Indica 11: 83-115. 

Barnes, H.F. 1926. An undescribed mushroom-feeding gall-midge. The Entomologist's Monthly Magazine 62: 89-92. 

Barnes, H.F. 1927a. British gall-midges. I. The Entomologist's Monthly Magazine 63: 164-168. 

Barnes, H.F. 1927b. Pp. 54-55. In Barnes, H.F. and F.V. Theobald. A new gall midge attacking Arabis albida. Journal 

of the Southeastern Agricultural College Wye 24: 54-55. 

Barnes, H.F. 1927c. Untitled, continuation of "British gall-midges. I." The Entomologist's Monthly Magazine 63: 

169-172. 

Barnes, H.F. 1927d. Four British aphid-eating gall-midges. The Entomologist 60: 174-180. 

Barnes, H.F. 1927e. Some Cecidomyidae from the Federated Malay States. Journal of the Federated Malay States 

Museum 13: 269-274. 

Barnes, H.F. 1927f. British gall-midges. I. The Entomologist's Monthly Magazine 63: 211-216. 

Barnes, H.F. 1927g. Untitled, continuation of "British gall-midges. I." The Entomologist's Monthly Magazine 63: 

217-221. 

Barnes, H.F. 1927h. A new rust-eating British gall-midge. The Entomologist 60: 220-221. 

Barnes, H.F. 1927i. A new gall midge (Mayetiola phalaris sp. n.) reared from Phalaris arundinacea. Zeitschrift für 

Angewandte Entomologie 13: 375-377. 

Barnes, H.F. 1928a. Gall midges (Cecidomyidae) infesting wild figs in the Philippine Islands. Hawaiian Sugar Planters' 

Association Experiment Station Entomological Series Bulletin 19: 21-23. 

Barnes, H.F. 1928b. British gall-midges. II. Lestodiplosis Kieffer. The Entomologist's Monthly Magazine 64: 68-72. 

Barnes, H.F. 1928c. Untitled, continuation of "British gall-midges. II. Lestodiplosis Kieffer." The Entomologist's 

Monthly Magazine 64: 73-75. 

Barnes, H.F. 1928d. British gall-midges. II. Lestodiplosis Kieffer. The Entomologist's Monthly Magazine 64: 142-144. 

Barnes, H.F. 1928e. Cecidomyiinae (gall midges). Pp. 103-108. In F.W. Edwards. Part VI. Fasc. 2. Nematocera. Pp. 

23-108. In Insects of Samoa and other Samoan Terrestrial Arthropoda. British Museum. 

Barnes, H.F. 1928f. Untitled, continuation of "British gall-midges. II. Lestodiplosis Kieffer." The Entomologist's 


Barnes, H.F. 1928g. Neptunimyia Felt, a gall midge genus new to Great Britain. The Entomologist 61: 173-175. 

Barnes, H.F. 1928h. The British gall midges of peas. Bulletin of Entomological Research 19: 183-185. 

Barnes, H.F. 1929. Two gall midges (Cecidomyidae) found in stored products. Bulletin of Entomological Research 20: 

119-122. 

Barnes, H.F. 1930a. On the biology of the gall-midges (Cecidomyidae) attacking meadow foxtail grass (Alopecurus 

pratensis), including the description of one new species. Annals of Applied Biology 17: 339-366.

458 

Barnes, H.F. 1930b. A new thrips-eating gall midge, Thripsobremia liothripis, gen. et sp. n. (Cecidomyidae). Bulletin 

of Entomological Research 21: 331-332. 

Barnes, H.F. 1931a. A new predacious gall midge (Dipt., Cecidomyidae). Bulletin of Entomological Research 22: 

205-207. 

Barnes, H.F. 1931b. Description of Amblardiella tamaricum Kieffer. Bulletin de la Société d'Histoire Naturelle de 

l'Afrique du Nord 1931: 271-272. 

Barnes, H.F. 1932a. A new saprophytic gall midge, Asynapta furcifer sp. n., on olives. Bollettino del Laboratorio di 

Zoologia Generale e Agraria della R. Istituto Superiore d'Agricoltura in Portici 26: 51-53. 

Barnes, H.F. 1932b. Notes on Cecidomyidae. Annals and Magazine of Natural History (10) 9: 475-484. 

Barnes, H.F. 1933. A cambium miner of basket willows (Agromyzidae) and its inquiline gall-midge (Cecidomyidae). 

Annals of Applied Biology 20: 498-519, pls. XXIX-XXX. 

Barnes, H.F. 1934. Lestodiplosis alvei sp.n. (Diptera, Cecidomyidae). Parasitology 26: 594-595. 

Barnes, H.F. 1935a. On the gall midges injurious to the cultivation of willows. II. The so-called "shot hole" gall midges 

(Rhabdophaga spp.). Annals of Applied Biology 22: 86-105, pls. XI-XIV. 

Barnes, H.F. 1935b. Two gall midges on Asphodelus in Sardinia. Bollettino del Laboratorio di Zoologia Generale e 

Agraria della R. Istituto Superiore d'Agricoltura in Portici 28: 70-74. 

Barnes, H.F. 1935c. Two gall midges on Erica arborea, including the description of a new species. Bulletin de la 

Société d'Histoire Naturelle de l'Afrique du Nord 26: 139-141. 

Barnes, H.F. 1935d. Some new coccid-eating gall midges (Cecidomyidae). Bulletin of Entomological Research 26: 

525-530. 

Barnes, H.F. 1936. Notes on Cecidomyidae. II. Annals and Magazine of Natural History (10) 17: 272-279. 

Barnes, H.F. 1939a. A new gall midge attacking beech buds. Arbeiten über physiologische und angewandte 

Entomologie 6: 41-43. 

Barnes, H.F. 1939b. Two new gall midges from Sesamum angustifolium. Annals and Magazine of Natural History (11) 

3: 453-456. 

Barnes, H.F. 1939c. Some gall midge species and their host plant range. Annals of Applied Biology 26: 318-347, pl. 

XXII. 

Barnes, H.F. 1939d. Contarinia mali n. sp. the apple blossom midge, with notes on the more common gall midges 

attacking apple and pear. Kontyû 13: 126-132. 

Barnes, H.F. 1939e. Grass seed Dasyneura gall midges. Arbeiten über physiologische und angewandte Entomologie 6: 

171-175. 

Barnes, H.F. 1939f. Gall midges (Cecidomyidae) associated with coffee. Revue de Zoologie et de Botanique Africaines 

32: 324-336. 

Barnes, H.F. 1940a. Two new pests of apple and black currant. Bulletin of Entomological Research 31: 85-87, pl. III. 

Barnes, H.F. 1940b. The gall midges attacking the seed-heads of cocksfoot, Dactylis glomerata, L. Bulletin of 

Entomological Research 31: 111-119, pls. IV-V. 

Barnes, H.F. 1944a. Clinodiplosis hydrangeae sp. n. (Cecidomyidae) causing a leaf blister gall on Hydrangea 

hortensis. The Entomologist 67: 129-130, pl. IV. 

Barnes, H.F. 1944b. Two new gall midges from Mauritius. Bulletin of Entomological Research 35: 211-213. 

Barnes, H.F. 1946a. Gall Midges of Economic Importance. I: Gall Midges of Root and Vegetable Crops. Crosby 

Lockwood & Son, Ltd. 104 pp., 10 pls. 

Barnes, H.F. 1946b. Gall Midges of Economic Importance. II: Gall Midges of Fodder Crops. Crosby Lockwood & Son, 

Ltd. frontispiece, 160 pp., 3 pls. 

Barnes, H.F. 1949. Gall Midges of Economic Importance. VII: Miscellaneous Crops. Crosby Lockwood & Son, Ltd. 

229 pp., 14 pls. 

Barnes, H.F. 1951a. Gall Midges of Economic Importance. V: Gall Midges of Trees. Crosby Lockwood & Son, Ltd. 

270 pp., 8 pls. 

Barnes, H.F. 1951b. A new gall midge (Dipt., Cecidomyidae) predaceous on the flour mite, Tyroglyphus farinae 

(Deg.). Bulletin of Entomological Research 42: 565-566. 

Barnes, H.F. 1952. The gall midges of St. John's wort (Hypericum spp.), with descriptions of two new species. Bulletin 

of Entomological Research 42: 697-705. 

Barnes, H.F. 1953. Description of the new gall midge found by M. R. Pussard on lavender, together with notes on the 

damage caused by some other species (Dipt. Cecidomyidae). Bulletin de la Société Entomologique de France

459 

58: 125-158. 

Barnes, H.F. 1954a. Gall-midge larvae as endoparasites, including the description of a species parasitising aphids in 

Trinidad, B.W.I. Bulletin of Entomological Research 45: 769-775. 

Barnes, H.F. 1954b. A new species of Trotteria Kieffer (Cecidomyidae) reared from unopened flower-buds of privet 

(Ligustrum vulgare L.) Entomologist's Record 66: 281-283. 

Barnes, H.F. 1956. Gall Midges of Economic Importance. VII: Gall Midges of Cereal Crops. Crosby Lockwood & Son, 

Ltd. 261 pp., 16 pls. 

Barnes, H.F. 1958a. The gall midges (Dipt., Cecidomyidae) of Timothy grass inflorescences, with the description of 

one new species. Annales Entomologici Fennici 24: 59-68. 

Barnes, H.F. 1958b. A new stem-inhabiting gall midge of Poa pratensis (Dipt., Cecidomyiidae). Beiträge zur 


Barnes, H.F. and K.P. Lamb. 1954. Gall midges (Cecidomyidae) on Coprosma (Rubiaceae) in New Zealand. 

Transactions of the Royal Society of New Zealand 82: 813-816, pl. 32. 

Barnes, J.K. 1988. Asa Fitch and the emergence of American entomology. New York State Museum Bulletin 461: iviii, 

1-120. 

Baylac, M. 1985. Etude biométrique de quelques larves du genre Lestodiplosis Kieffer (Dipt. Cecidomyiidae). Annales 

de la Société Entomologique de France 21: 367-379. 

Baylac, M. 1986. Note sur le polymorphisme alaire dans le genre Lestodiplosis Kieffer (Dipt. Cecidomyiidae). Bulletin 

de la Société Entomologique de France 91: 41-45. 

Baylac, M. 1987a. Redescription de Monarthropalpus buxi (Laboulbène), désignation d'un lectotype (Dipt. 

Cecidomyiidae). Bulletin de la Société Entomologique de France 91: 259-269. 

Baylac, M. 1987b. Description d'un Lestodiplosis nouveau, prédateur d'Elaeidobius subvittatus (Col. Curculionidae), 

pollinisateur du palmier à huile Elaeis guineensis en Colombie (Dipt. Cecidomyiidae). Bulletin de la Société 

Entomologique de France 92: 125-132. 

Baylac, M. 1987c. Morphologie comparée de quelques larves du genre Lestodiplosis Kieffer (Diptera, Cecidomyiidae). 

Nouvelle Revue d'Entomologie 4: 195-209. 

Baylac, M. 1988. Description de Tessaradiplosis genre nouveau pour Cecidomyia entomophila Perris [Diptera, 

Cecidomyiidae]. Revue Française d'Entomologie 10: 31-35. 

Becknazarova, O.Kh. and E.B. Antonova. 1973. [Gall midges damaging plant reproductive structures in Turkmenia 

(Diptera, Cecidomyiidae)]. Izvestiya Akademiï Nauk Turkmenskoï SSR, Seriya Biologicheskikh Nauk 1973 

(6): 86-88. 

Becknazarova, O.Kh. and E.B. Antonova. 1977. [Gall midges (Diptera, Cecidomyiidae) damaging fruit of Aellenia 

subaphylla.] Izvestiya Akademiï Nauk Turkmenskoï SSR, Seriya Biologicheskikh Nauk 1977 (6): 41-46. 

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equivalent later publication are as follows: pp. 33-70 = 1918b; 120-130 = 1918c; 145-147 = 1918d; 151-154 = 

1918e; 157-166 = 1918f; 170-181 = 1918g; 182-186 = 1918h; 187-202 = 1918i; 203-207 = 1918j; 208-209 =

465 

1919a; 210-212 = 1919b; 213-214 = 1919c; 215-217 = 1919d; 218-222 = 1919e; 223-226 = 1919f; 227-230 = 

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Del Guercio, G. 1918f. Notizie preliminari sul nuovo moscerino del gelso (Moreschiella moricola n. gen., n. sp.). 

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Del Guercio, G. 1918g. I moscerini delle ghiande della querce e del leccio in Italia. Agricoltura Coloniale 12: 358-369. 

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Del Guercio, G. 1918i. Il moscerino dell'Aphis symphyti Schr. Agricoltura Coloniale 12: 375-390. 

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Del Guercio, G. 1919b. Il moscerino del Microsiphon del carciofo (Rondaniella microsiphonis sp. n.). Agricoltura 


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Del Guercio, G. 1919d. Il moscerino dell'afide del cotogno ornamentale (Rondaniella ornata sp. n.). Agricoltura 


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Del Guercio, G. 1919f. Il moscerino dell'afide del papavero (Trilobia aphidisuga n. g., n. sp.). Agricoltura Coloniale 

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Del Guercio, G. 1919g. Il moscerino dell'afide del lanigero (Uncinulella eriosomiperda n. g., n. sp.). Agricoltura 


Del Guercio, G. 1919h. Il moscerino degli Adelges dell'abeto (Adelgimyza strobilobii n. g., n. sp.). Agricoltura 


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(1991) 70 (3): 49-62. 

Fedotova, Z.A. 1991d. [Review of the gall midges of the subtribe Dasineurina (Diptera, Cecidomyiidae, Oligotrophini) 

in Kazakhstan.] Izvestiya Akademii Nauk Kazakhskoï SSR, Seriya Biologicheskaya 1991 (4): 16-25. 

Fedotova, Z.A. 1991e. [Gall midges of the genus Baldratia (Diptera, Cecidomyiidae) from Kazakhstan.] 


Fedotova, Z.A. 1991f. [Gall midges (Diptera, Cecidomyiidae) injurious to Salsola richteri in Karakum and Kyzylkum. 

Part 1.] Izvestiya Akademiï Nauk Turkmenskoï SSR, Seriya Biologicheskikh Nauk 1991 (5): 9-16. 

Fedotova, Z.A. 1991g. [Gall midges (Diptera, Cecidomyiidae) injurious to Salsola richteri in Karakum and Kyzylkum. 


Fedotova, Z.A. 1992a. [New species of gall midges (Diptera, Cecidomyiidae) from brushwood in Kazakhstan and 

Turkmenia.] Zoologicheskiï Zhurnal 71 (1): 75-87.

470 

Fedotova, Z.A. 1992b. [Gall midges of the subtribe Baldratiina (Diptera, Cecidomyiidae, Lasiopterini) in Kazakhstan.] 


Fedotova, Z.A. 1992c. [New species of gall midges of the genus Halodiplosis (Diptera, Cecidomyiidae) from Anabasis 

spp. in Kazakhstan.] Zoologicheskiï Zhurnal 71 (6): 71-79. English translation in Entomological Review 

(1993) 71 (9): 95-103. 

Fedotova, Z.A. 1992d. [New species of gall midges (Diptera, Cecidomyiidae) developing on Chenopodiaceae in 

Kazakhstan and Turkmenia.] Zoologicheskiï Zhurnal 71 (7): 29-40. English translation in Entomological 

Review (1993) 72 (1): 66-77. 

Fedotova, Z.A. 1992e. [Review of species of gall midges (Diptera, Cecidomyiidae) on saksaul (Arthrophytum spp.) 

from Kazakhstan.] Zoologicheskiï Zhurnal 71 (7): 41-52. English translation in Entomological Review (1993) 

72 (1): 78-88. 

Fedotova, Z.A. 1992f. [New species of gall midges (Diptera, Cecidomyiidae) on Chenopodiaceae in Kazakhstan.] 

Izvestiya Akademiya Nauk Respubliki Kazakhstan, Seriya Biologicheskaya 1992 (1): 44-52. 

Fedotova, Z.A. 1992g. [New species of gall midges (Diptera, Cecidomyiidae) damaging plants in Kazakhstan.] 

Izvestiya Akademiya Nauk Respubliki Kazakhstan, Seriya Biologicheskaya 1992 (3): 43-54. 

Fedotova, Z.A. 1992h. [New and litle known gall midge genera (Diptera, Cecidomyiidae) from Kazakhstan.] Vestnik 

Zoologii 1992 (1): 16-21. 

Fedotova, Z.A. 1992i. [A new genus and a new species of gall midges (Diptera, Cecidomyiidae) from Kazakhstan.] 


Fedotova, Z.A. 1992j. [New species and genera of gall midges (Diptera, Cecidomyiidae) living on Brassicaceae in 

Kazakhstan.] Entomologicheskoe Obozrenie 71: 664-673. English translation in Entomological Review 

(1993) 72 (5): 8-18. 

Fedotova, Z.A. 1992k. [New species of gall-midges (Diptera, Cecidomyiidae) from Salsola richteri in Turkmenistan. 

Part 1.] Izvestiya Akademiï Nauk Turkmenskoï SSR, Seriya Biologicheskikh Nauk 1992 (4) 70-74. 

Fedotova, Z.A. 1992l. [New species of gall-midges (Diptera, Cecidomyiidae) from Salsola richteri in Turkmenistan. 


Fedotova, Z.A. 1992m. [Review of gall midges (Diptera, Cecidomyiidae) developing in Tamarix spp. with descriptions 

of new taxa from Kazakhstan and Turkmenistan. Part 1.] Zoologicheskiï Zhurnal 71 (12): 87-98. English 


*Fedotova, Z.A. 1992n. [Phytophagous gall midges (Diptera, Cecidomyiidae) on certain weeds and poisonous plants of 

Kazakhstan.] Pp. 96-110. In [Biological Methods for the Control of Injurious Weeds of Kazakhstan]. Alma- 

Ata. 

*Fedotova, Z.A. 1992o. Pp. 116-128. In Maralbaev, T.A. and Z.A. Fedotova. [Review of phytophagous gallmakers on 

pasture weeds in Kazakhstan]. Pp. 111-129. In [Biological Methods for the Control of Injurious Weeds of 

Kazakhstan]. Alma-Ata. 

Fedotova, Z.A. 1993a. [Review of gall midges (Diptera, Cecidomyiidae) developing on Tamarix spp. with descriptions 

of new taxa from Kazakhstan and Turkmenistan. Part 2.] Zoologicheskiï Zhurnal 72 (1): 63-73. English 


Fedotova, Z.A. 1993b. [Two new juniper inhabiting gall midges species (Diptera, Cecidomyiidae) from Turkmenistan.] 

Vestnik Zoologii 1993(1): 17-21. 

Fedotova, Z.A. 1993c. [New species of gall midges of the genus Halodiplosis (Diptera, Cecidomyiidae) from 

Kazakhstan.] Zoologicheskiï Zhurnal 72 (3): 106-118. English translation in Entomological Review (1993) 72 

(7): 121-132. 

Fedotova, Z.A. 1993d. [Gall midges of the genus Ephedromyia (Diptera, Cecidomyiidae) in Kazakhstan and 

Turkmenia.] Zoologicheskiï Zhurnal 72 (4): 88-96. English translation in Entomological Review (1994) 72 

(8): 131-139. 

Fedotova, Z.A. 1993e. [New species of gall midges (Diptera, Cecidomyiidae) on Chenopodiaceae from Kazakhstan and 

Turkmenia.] Zoologicheskiï Zhurnal 72 (5): 77-92. English translation in Entomological Review (1995) 74 

(1): 79-93. 

Fedotova, Z.A. 1993f. [New species of gall midges of the genus Contarinia Rd. (Diptera, Cecidomyiidae) developing 

on legumes (Fabaceae) in Kazakhstan.] Entomologicheskoe Obozrenie 72 (3): 675-683. English translation in 

Entomological Review (1994) 73 (4): 64-72. 

Fedotova, Z.A. 1993g. [Review of the gall midges of the tribe Cecidomyiini Cecidomyiini (Diptera, Cecidomyiidae)

471 

from Kazakhstan.] Izvestiya Natsional'noï Akademii Nauk Respubliki Kazakhstan, Seriya Biologicheskaya 3 

(177): 13-23. 

Fedotova, Z.A. 1993h. [New species of gall midges (Diptera, Cecidomyiidae) inhabiting Chenopodiaceae in Turkmenia 

and Kazakhstan.] Bulletin of the Moscow Society of Naturalists, Biological Series 98 (6): 48-54. 

Fedotova, Z.A. 1993i. [New species of gall midges (Diptera, Cecidomyiidae) from flower galls in Kazakhstan.] Bulletin 

of the Moscow Society of Naturalists, Biological Series 98 (4): 46-53. 

Fedotova, Z.A. 1993j. [Review of the gall midges (Diptera, Cecidomyiidae) damaging Astragalus spp. in Kazakhstan. 

Part 1.] Zoologicheskiï Zhurnal 72 (10): 75-89. English translation in Entomological Review (1994) 73 (6): 

107-120. 

Fedotova, Z.A. 1993k. [Review of the gall midges (Diptera, Cecidomyiidae) damaging Astragalus spp. in Kazakhstan. 


121-136. 

Fedotova, Z.A. 1993l. [New species of flower gall midges (Diptera, Cecidomyiidae) from the mountains of 

Kazakhstan.] Zoologicheskiï Zhurnal 72 (11): 76-90. English translation in Entomological Review (1994) 73 

(3): 93-106. 

Fedotova, Z.A. 1993m. [Gall midges (Diptera, Cecidomyiidae) injurious to Salsola arbuscula Pall. in Kazakhstan.] 

Bulletin of the Moscow Society of Naturalists, Biological Series 98 (2): 19-26. 

Fedotova, Z.A. 1993n. [New species of gall midges (Diptera, Cecidomyiidae) developing on Hammada in Turkestan.] 


Fedotova, Z.A. 1993o. [New geranium-inhabiting gall midges species (Diptera, Cecidomyiidae) from Kazakhstan.] 


Fedotova, Z.A. 1993p. [Two new species of gall-midges of the tribe Asphondyliini (Diptera, Cecidomyiidae) from 

Turkmenistan.] Izvestiya Akademii Nauk Turkmenistana, Seriya Biologicheskikh Nauk 1993 (2): 57-60. 

Fedotova, Z.A. 1993q. [Classification and phylogeny of the gall midges of the supertribe Lasiopteridi (Diptera, 

Cecidomyiidae).] Zoologicheskiye Issledovaniya v Kazakhstane 1993: 27-56. 

Fedotova, Z.A. 1993r. [Gall midges of the Repetek Biosphere Reservation.] Problemy Osvoeniya Pustyn' 1993 (3): 49- 

53. English translation in Problems of Desert Development, 1994, pp. 10-16, Allerton Press, Inc., New York, 

NY. 

Fedotova, Z.A. 1994a. [Gall midges (Diptera, Cecidomyiidae) of Brassicaceae: review of the Palearctic fauna with 

description of new species from Kazakhstan. Part 1.] Zoologicheskiï Zhurnal 73(2): 112-130. English 


Fedotova, Z.A. 1994b. [Gall midges (Diptera, Cecidomyiidae) of Brassicaceae: review of the Palearctic fauna with 

description of new species from Kazakhstan. Part 2.] Zoologicheskiï Zhurnal 73(4): 79-90. English translation 

in Entomological Review (1995) 74 (1): 114-125. 

Fedotova, Z.A. 1994c. [Gall midges (Diptera, Cecidomyiidae) of Brassicaceae: review of the Palearctic fauna with 

description of new species from Kazakhstan. Part 3.] Zoologicheskiï Zhurnal 73(4): 91-102. English 


Fedotova, Z.A. 1994d. [New species of gall-midges (Diptera, Cecidomyiidae) on Salsola chiwensis from Kazakhstan 

and Turkmenistan.] Izvestiya Natsional'noï Akademii Nauk Respubliki Kazakhstan, Seriya Biologicheskaya 

1994(1): 42-47. 

Fedotova, Z.A. 1994e. [Gall midges (Diptera, Cecidomyiidae) developing on Silene spp. in Kazakhstan.] Izvestiya 

Natsional'noï Akademii Nauk Respubliki Kazakhstan, Seriya Biologicheskaya 1994(2): 42-48. 

Fedotova, Z.A. 1994f. [New species of gall midges of the genus Halodiplosis Kieff. (Diptera, Cecidomyiidae) from 

Kazakhstan.] Izvestiya Natsional'noï Akademii Nauk Respubliki Kazakhstan, Seriya Biologicheskaya 

1994(5): 31-36. 

Fedotova, Z.A. 1994g. Not seen. [New species of gall-midges (Diptera, Cecidomyiidae) developing on legumes in 

Turkmenia.] Izvestiya Akademiï Nauk Turkmenskoï SSR, Seriya Biologicheskikh Nauk 1994 (5): ??. Per 

Fedotova 2000b: 624, item #325. 

Fedotova, Z.A. 1994h. [Three new species of gall midges (Diptera, Cecidomyiidae) from Kazakhstan.] Vestnik 

Zoologii 1994 (4-5): 29-34. 

Fedotova, Z.A. 1994i. [Gall midges (Diptera, Cecidomyiidae) injurious to spurges (Euphorbia spp.) in Kazakhstan. 

Entomologicheskoe Obozrenie 73: 447-464. English translation in Entomological Review (1995) 74 (6): 

23-40.

472 

Fedotova, Z.A. 1995a. [New species of gall-midges of the genus Arthrocnodax Rübsaamen (Diptera, Cecidomyiidae) 

from Kazakhstan.] Selevinia 3 (3): 10-26. 

Fedotova, Z.A. 1995b. [A review of Palaearctic gall midges (Diptera, Cecidomyiidae) from Achillea and Tanacetum, 

with description of three new species from Kazakhstan.] Vestnik Zoologii 1995 (2-3): 22-29. 

Fedotova, Z.A. 1995c. [Review of gall midges (Diptera, Cecidomyiidae) damaging Aster and Erigeron in Kazakhstan. 


166-179. 

Fedotova, Z.A. 1995d. [Gall midges of the genus Dasyneuriola Marikovskij (Diptera, Cecidomyiidae) from 

Kazakhstan.] Entomologicheskoe Obozrenie 74: 706-710. English translation in Entomological Review 

(1996) 75 (9): 61-65. 

Fedotova, Z.A. 1995e. [New species of gall midges (Diptera, Cecidomyiidae) from Asteraceae in Kazakhstan.] Vestnik 

Zoologii 1995 (5-6): 18-24. 

Fedotova, Z.A. 1996a. [Review of gall midges (Diptera, Cecidomyiidae) damaging Aster and Erigeron in Kazakhstan. 

Part 2.] Zoologicheskiï Zhurnal 75 (14): 538-551. 

Fedotova, Z.A. 1996b. [New species of gall-midges (Diptera, Cecidomyiidae) from flower heads of Asteraceae in 

Kazakhstan.] Entomologicheskoe Obozrenie 75: 192-207. 

Fedotova, Z.A. 1996c. [New species of gall midges (Diptera, Cecidomyiidae) from southern Kazakhstan.] Izvestiya 

Natsional'noï Akademii Nauk Respubliki Kazakhstan, Seriya Biologicheskaya 1996 (1): 34-39. 

Fedotova, Z.A. 1996d. [New species of flower gall-midges (Diptera, Cecidomyiidae) from Kazakhstan.] Vestnik 

Zoologii 1996(4-5): 8-15. 

Fedotova, Z.A. 1996e. [New species of gall-midges (Diptera, Cecidomyiidae) developing on legumes in Kazakhstan.] 

Bulletin of the Moscow Society of Naturalists, Biological Series 101 (3): 42-53. 

Fedotova, Z.A. 1996f. [New and little known species of gall midges (Diptera, Cecidomyiidae) from brushwood in 

Kazakhstan.] Bulletin of the Moscow Society of Naturalists, Biological Series 101 (5): 50-60. 

Fedotova, Z.A. 1997a. [New species of gall midges of the genus Arthrocnodax Rübsaamen (Diptera, Cecidomyiidae) 

from Kazakhstan.] Bulletin of the Moscow Society of Naturalists, Biological Series 102 (2): 39-47. 

Fedotova, Z.A. 1997b. [Review of gall midges (Diptera, Cecidomyiidae) developing on Rubiaceae with descriptions of 

new species from Kazakhstan. Part 1.] Zoologicheskiï Zhurnal 76 (1): 43-54. English Translation in 


Fedotova, Z.A. 1997c. [Review of gall midges (Diptera, Cecidomyiidae) developing on Rubiaceae with descriptions of 

new species from Kazakhstan. Part 2.] Zoologicheskiï Zhurnal 76 (3): 326-340. English Translation in 


Fedotova, Z.A. 1997d. [Review of gall midges (Diptera, Cecidomyiidae) developing on Palaearctic Fabaceae with a 

conspectus of the Kazakhstan fauna. Part 1.] Zoologicheskiï Zhurnal 76 (4): 448-459. English Translation in 


Fedotova, Z.A. 1997e. [Review of gall midges (Diptera, Cecidomyiidae) developing on Fabaceae with a conspectus of 

the Kazakhstan fauna. Part 2.] Zoologicheskiï Zhurnal 76 (5): 565-574. English Translation in Entomological 

Review (1997) 77 (9): 1129-1139. 

Fedotova, Z.A. 1997f. Review of gall midges (Diptera, Cecidomyiidae) developing on Palaearctic Polygonaceae with 

descriptions of new species from Kazakhstan. Part. 1.] Entomologicheskoe Obozrenie 76 (2): 448-458. 

English translation in Entomological Review (1997) 76 (2): 232-240. 

Fedotova, Z.A. 1997g. [Review of gall midges (Diptera, Cecidomyiidae) developing on Palaearctic Polygonaceae with 

descriptions of new species from Kazakhstan. Part. 2.] Entomologicheskoe Obozrenie 76 (4): 874-891. 


Fedotova, Z.A. 1998a. [New species of gall-midges (Diptera, Cecidomyiidae) from Asteraceae in Kazakhstan.] Bulletin 

of the Moscow Society of Naturalists, Biological Series 103 (1): 28-37. 

Fedotova, Z.A. 1998b. [New species of gall-midges (Diptera, Cecidomyiidae) from Kazakhstan.] Bulletin of Moscow 

Society of Naturalists 103 (2): 10-18. 

Fedotova, Z.A. 1999a. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp., in 

Kazakhstan and Turkmenistan. 1. The genera Boucheella and Arenaromyia, new genus.] Zoologicheskiï 

Zhurnal 78 (5): 580-595. English translation in Entomological Review (1999) 79 (9): 1199-1214. 

Fedotova, Z.A. 1999b. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp., in 

Kazakhstan and Turkmenistan. 2. The genus Navasiella.] Zoologicheskiï Zhurnal 78 (6): 681-696. English

473 


Fedotova, Z.A. 1999c. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp., in 

Kazakhstan and Turkmenistan. 3. The genus Dracunculomyia.] Zoologicheskiï Zhurnal 78 (7): 834-848. 

Fedotova, Z.A. 1999d. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp., in 

Kazakhstan and Turkmenistan. 4. Species of the genus Dracunculomyia from Artemisia marschalliana and A. 

scoparia.] Zoologicheskiï Zhurnal 78 (8): 965-978. 

Fedotova, Z.A. 1999e. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp., in 

Kazakhstan and Turkmenistan. 5. The Dracunculomyia species complex from Artemisia dracunculus.] 


Fedotova, Z.A. 1999f. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp., in 

Kazakhstan and Turkmenistan. 6. New species of the genus Dracunculomyia.] Zoologicheskiï Zhurnal 78 

(11): 1306-1319. 

Fedotova, Z.A. 2000a. [Review of gall midges of the genus Macrolabis Kieffer (Diptera, Cecidomyiidae) with 

description of new species from Kazakhstan.] Entomologicheskoe Obozrenie 79 (2): 463-481. English 


Fedotova, Z.A. 2000b. [Plant-Feeding Gall Midges (Diptera, Cecidomyiidae) of the Deserts and Mountains of 

Kazakhstan: Morphology, Biology, Distribution, Phylogeny, and Systematics.] Samara. 803 pp. 

Fedotova, Z.A. 2000c. Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp., in 

Kazakhstan and Turkmenistan. 7. A key to species of the genus Dracunculomyia and analysis of the fauna.] 

Zoologicheskiï Zhurnal 79 (11): 1313-1324. English translation in Entomological Review (2000) 80 (8): 982- 

994. 

Fedotova, Z.A. 2000d. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp. in 

Kazakhstan and Turkmenistan. 8. The genus Seriphidomyia gen. n.] Zoologicheskiï Zhurnal 79 (12): 

1421-1434. English translation in Entomological Review (2000) 80 (8): 995-1008. 

Fedotova, Z.A. 2001a. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp. in 

Kazakhstan and Turkmenistan. 9. The genus Seriphidomyia gen. n.] Zoologicheskiï Zhurnal 80 (1): 52-66. 


Fedotova, Z.A. 2001b. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia spp. in 

Kazakhstan and Turkmenistan. 10. Genus Seriphidomyia, subgenus Polynomyia.] Zoologicheskiï Zhurnal 80 

(6): 693-702. English translation in Entomological Review (2001) 81 (4): 463-472. 

Fedotova, Z.A. 2001c. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia: taxa from 

Kazakhstan and Turkmenistan. 11. A key to species of Seriphidomyia.] Zoologicheskiï Zhurnal (2001) 80 (7): 


Fedotova, Z.A. 2001d. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia: taxa from 

Kazakhstan and Turkmenistan. 12. The genus Dichelonyx.] Zoologicheskiï Zhurnal (2001) 80 (8): 952-967. 


Fedotova, Z.A. 2001e. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia: taxa from 

Kazakhstan and Turkmenistan. 13. Pupascleromyia, n. gen.] Zoologicheskiï Zhurnal (2001) 80 (9): 1083- 

1098. English translation in Entomological Review (2001) 81 (5): 565-580. 

Fedotova, Z.A. 2001f. [Review of gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia, with 

description of new taxa from Kazakhstan and Turkmenistan. 14. Palaearctic fauna. Types of pupae and 

relationships between gall midge genera and Artemisia subgroups.] Zoologicheskiï Zhurnal (2001) 80 (10): 


Fedotova, Z.A. 2001g. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia: taxa from 

Kazakhstan and Turkmenistan. 15. The types of galls in relation to plant hosts.] Zoologicheskiï Zhurnal 

(2001) 80 (11): 1338-1354. English translation in Entomological Review (2001) 81 (7): 829-846. 

Fedotova, Z.A. 2001h. [Gall-midges (Diptera, Cecidomyiidae) developing on wormwoods, Artemisia: taxa from 

Kazakhstan and Turkmenistan. 16. Species of genera nonspecific to Artemisia and a list of Palearctic species 

and their food plants.] Zoologicheskiï Zhurnal (2001) 80 (12): 1481-1493. English translation in 


Fedotova, Z.A. 2002a. [Gall-midges (Diptera, Cecidomyiidae) developing on Rosaceae in the Palaearctic and a 

summary of the fauna from Kazakhstan. Report 1.] Zoologicheskiï Zhurnal 81 (1): 60-70. English translation 

in Entomological Review (2002) 81 (1): 443-453.

474 

Fedotova, Z.A. 2002b. [Gall midges (Diptera, Cecidomyiidae) developing on Rosaceae in the Palaearctic and a 

summary of the fauna from Kazakhstan. Report 2.] Zoologicheskiï Zhurnal 81 (2): 201-211. English 


*Fedotova, Z.A. 2002c. New species of gall midges (Diptera, Cecidomyiidae) from the Russian Far East. Far Eastern 

Entomologist 2002 (118): 1-35. 

*Fedotova, Z.A. 2003a. [New species of gall midges (Diptera, Cecidomyiidae) developing on Galatella from 

Kazakhstan.] Zoologicheskiï Zhurnal 82: (7): 825-833. English translation in Entomological Review (2003) 

83 (3): 368-376. 

*Fedotova, Z.A. 2003b. [A review of Palaearctic gall midges of the genus Asphondylia (Diptera, Cecidomyiidae) with 

descriptions of new species from the middle Volga River Region.] Zoologicheskiï Zhurnal 82: (8): 972-985. 


*Fedotova, Z.A. 2003c. New genera and species of gall midges (Diptera, Cecidomyiidae) from the Russian Far East. 

Far Eastern Entomologist 125: 1-30. 

*Fedotova, Z.A. 2003d. Pp. 5-11. In Fedotova, Z.A. and V.S. Sidorenko. New species of gall midges of the tribe 

Stomatosematini (Diptera, Cecidomyiidae) from the Russian Far East. Far Eastern Entomologist 128: 1-11. 

*Fedotova, Z.A. 2003e. New species from new genera of gall midges (Diptera, Cecidomyiidae) from the Russian Far 

East. International Journal of Dipterological Research 14: 43-72. 

*Fedotova, Z.A. 2004a. Pp. 45-54. In Perkovsky, E.E. and Z.A. Fedotova. 2004. [New species of gall midges (Diptera, 

Cecidomyiidae) from Rovno amber: Subfamily Lestremiinae, tribes Micromyiini and Peromyiini.] 

Paleontologicheskii Zhurnal 2004 (5): 44-54. English translation in Paleontological Journal 38: 396-406. 

*Fedotova, Z.A. 2004b. [A review of gall midges (Diptera, Cecidomyiidae) developing on mullein (Verbascum spp.) 

with description of new taxa from the middle Volga River Basin.] Zoologicheskiï Zhurnal 83: (7): 809-825. 

*Fedotova, Z.A. 2004c. [New species of the gall midge genus Macrolabis Kieffer (Diptera, Cecidomyiidae) from 

Kazakhstan and Russia.] Entomologicheskoe Obozrenie 83 (3): 718-733. English translation in Entomological 

Review 84: 704-716. 

*Fedotova, Z.A. 2004d. Pp. 69-77. In Fedotova, Z.A. and E.E. Perkovsky. 2004. [New gall midges (Diptera, 

Cecidomyiidae) from the Rovno amber: Subfamily Lestremiinae, Tribes Strobliellini and Campylomyzini; 

Subfamily Porricondylinae, Tribes Diadocidiini and Asynaptini.] Paleontologicheskii Zhurnal 2004: 69-78. 

English translation in Paleontological Journal 39: 538-547. 

*Fedotova, Z.A. 2004e. Pp. 12-55. In Fedotova, Z.A. and V.S. Sidorenko. 2004a. A new genus and species of gall 

midges from the Russian Far East (Diptera: Cecidomyiidae). International Journal of Dipterological Research 

15: 9-55. 

*Fedotova, Z.A. 2004f. Pp. 100-114. In Fedotova, Z.A. and V.S. Sidorenko. 2004b. New taxa of gall midges from the 

Russian Far East (Diptera, Cecidomyiidae). Entomofauna: Zeitschrift für Entomologie 25; 97-115. 

*Fedotova, Z.A. 2004g. Pp. 8-31. In Fedotova, Z.A. and V.S. Sidorenko. 2004c. New species of gall midges of the 

genus Karshomyia Felt, 1908 (Diptera, Cecidomyiidae) and related new genera from the Russian Far East. Far 

Eastern Entomologist 137: 1-32. 

*Fedotova, Z.A. 2004h. Pp. 165-185. In Fedotova, Z.A. and V.S. Sidorenko. 2004d. New genera and species of gall 

midges (Diptera, Cecidomyiidae) from the Russian Far East. International Journal of Dipterological Research 

15: 163-185. 

*Fedotova, Z.A. 2004i. [Supplement. 22. Fam. Cecidomyiidae - gall midges.] Pp. 565-629. In Ler, P.A., ed. 

Opredelitel' Nasekomych Dal'nego Vostoka Russia v Shesti Tomakh. VI. Dvukrylye i Blokhi. Part 3. 

Vladivostok Dal'nauka. 

*Fedotova, Z.A. 2005a. Pp. 438-468. In Fedotova, Z.A. and V.S. Sidorenko. 2005b. New species of gall midges 

(Diptera, Cecidomyiidae) of Artemisia rubripes Nakai (Asteraceae) [= Compositae] from the Russian Far 

East. Studia Dipterologica 11 (2004): 437-469. 

*Fedotova, Z.A. 2005b. [New gall-midge species (Diptera, Cecidomyiidae) from southeastern Kazakhstan.] 

Entomologicheskoe Obozrenie 84 (4): 897-910. English translation in Entomological Review 85 (8): 972-983. 

*Fedotova, Z.A. 2005c. Pp. 216-229. In Fedotova, Z.A. and V.S. Sidorenko. 2005a. New taxa of gall midges from the 

Russian Far East (Diptera: Cecidomyiidae). Zoosystematica Rossica 13: 213-231. 

*Fedotova, Z.A. 2005d. Pp. 230-238. In Fedotova, Z.A. and V.S. Sidorenko. 2005c. New taxa of the tribe 

Coquillettomyiini Mamaev, 1968 (Diptera: Cecidomyiidae) from the Russian Far East. Russian Entomological 

Journal 14: 227-238.

475 

*Fedotova, Z.A. 2005e. Pp. 42-53. In Fedotova, Z.A. and E.E. Perkovsky. 2005. [New gall midges (Diptera, 

Cecidomyiidae) from the Rovno amber: Subfamily Porricondylinae (Tribes Bryocryptini and Winnertziini) 

and subfamily Lasiopterinae (Tribes Brachineurini and Oligotrophini).] Paleontologicheskii Zhurnal 2005 (1): 

42-53. English translation in Paleontological Journal 39: 41-51. 

*Fedotova, Z.A. 2006a. Pp. 143-165. In Fedotova, Z.A. and V.S. Sidorenko. 2006a. New genera and species of gall 

midges from the Russian Far East (Diptera: Cecidomyiidae). Entomofauna: Zeitschrift für Entomologie 27: 

133-166. 

*Fedotova, Z.A. 2007. [Relationships between Trotteria and Verbasciola (Diptera, Cecidomyiidae) gall midges with a 

description of new species from the Middle Volga River Area]. Zoologicheskiï Zhurnal 86: 452-474. English 

translation in Entomological Review 87: 598-620. 

*Fedotova, Z.A. 2008. [New genus and species of the gall-midges (Diptera, Cecidomyiidae) from Zhigulevskyy Nature 

Reserve.] Isvestiya Samarskogo Nauchnogo Tsentra Rosikaoi Akademii Nauk 10: 119-145. 

*Fedotova, Z.A. 2009. Pp. 12-24. In Fedotova, Z.A. and V.S. Sidorenko. 2009b. New taxa of gall midges from the 

tribes Quadridiplosini and Heterodiplosini (Diptera, Cecidomyiidae) from the Russian Far East. Far Eastern 

Entomologist 196: 1-27. 

Fedotova, Z.A. and O.V. Kovalev. 2001. [22. Fam. Cecidomyiidae - gall midges.] Pp. 390-614. In Ler, P.A., ed. 

Opredelitel' Nasekomych Dal'nego Vostoka Russia v Shesti Tomakh. VI. Dvukrylye i Blokhi. Part 2. 


*Fedotova, Z.A. and O.V. Kovalev. 2003c. Pp. 25-29. In Fedotova, Z.A. New genera and species of gall midges 

(Diptera, Cecidomyiidae) from the Russian Far East. Far Eastern Entomologist 125: 1-30. 

*Fedotova, Z.A. and O.V. Kovalev. 2005. Pp. 457-459. In Fedotova, Z.A. and V.S. Sidorenko. 2005b. New species of 

gall midges (Diptera, Cecidomyiidae) of Artemisia rubripes Nakai (Asteraceae) [= Compositae] from the 

Russian Far East. Studia Dipterologica 11 (2004): 437-469. 

*Fedotova, Z.A. and E.E. Perkovsky. 2004. [New gall midges (Diptera, Cecidomyiidae) from the Rovno amber: 

Subfamily Lestremiinae, Tribes Strobliellini and Campylomyzini; Subfamily Porricondylinae, Tribes 

Diadocidiini and Asynaptini.] Paleontologicheskii Zhurnal 2004 (5): 69-78. English translation in 

Paleontological Journal 38 (5): 538-547. 

*Fedotova, Z.A. and E.E. Perkovsky. 2005. [New gall midges (Diptera, Cecidomyiidae) from the Rovno amber: 

Subfamily Porricondylinae (Tribes Bryocryptini and Winnertziini) and subfamily Lasiopterinae (Tribes 

Brachineurini and Oligotrophini).] Paleontologicheskii Zhurnal 2005 (1): 42-53. English translation in 

Paleontological Journal 39: 41-51. 

*Fedotova, Z.A. and E.E. Perkovsky. 2007a. [New gall midges of the Subfamily Lestremiinae (Diptera, 

Cecidomyiidae) from the Rovno amber.] Paleontologicheskii Zhurnal 2007 (4): 82-95. English translation in 


*Fedotova, Z.A. and E.E. Perkovsky. 2007b. Problems and prospects in the study of fossil gall midges. Reply to the 

criticism of M. Jaschhof. Paleontologicheskii Zhurnal 2007 (6): 101-107. English translation in 


*Fedotova, Z.A. and E.E. Perkovsky. 2008a. [New taxa of gall midges (Diptera, Cecidomyiidae) from Dubrovitsa 

(Rovno amber).] Vestnik Zoologii 42: 27-40. 

*Fedotova, Z.A. and E.E. Perkovsky. 2008b. Pp. 60-68. In Perkovsky, E.E. and Z.A. Fedotova. 2008a. [New gall 

midge taxa of the subfamilies Porricondylinae and Lasiopterinae (Diptera, Cecidomyiidae) from the Rovno 

amber. Paleontologicheskii Zhurnal 2008 (2): 60-69. English translation in Paleontological Journal 42: 166- 

175. 

*Fedotova, Z.A. and E.E. Perkovsky. 2008c. Pp. 410-423. In Perkovsky, E.E. and Z.A. Fedotova. 2008b. [On the 

systematics of the gall midge supertribe Heteropezidi (Diptera, Cecidomyiidae) from the Rovno amber: new 

taxa and combinations of the tribes Heteropezini and Miastorini]. Vestnik zoologii 42: 403-425. 

*Fedotova, Z.A. and V.S. Sidorenko. 2003. New species of gall midges of the tribe Stomatosematini (Diptera, 

Cecidomyiidae) from the Russian Far East. Far Eastern Entomologist 128: 1-11. 

*Fedotova, Z.A. and V.S. Sidorenko. 2004a. A new genus and species of gall midges from the Russian Far East 

(Diptera: Cecidomyiidae). International Journal of Dipterological Research 15: 9-55. 

*Fedotova, Z.A. and V.S. Sidorenko. 2004b. New taxa of gall midges from Russian Far East (Diptera: Cecidomyiidae). 

Entomofauna: Zeitschrift für Entomologie 25: 97-115. 

*Fedotova, Z.A. and V.S. Sidorenko. 2004c. New species of gall midges of the genus Karshomyia Felt, 1908 (Diptera,

476 

Cecidomyiidae) and related new genera from the Russian Far East. Far Eastern Entomologist 137: 1-32. 

*Fedotova, Z.A. and V.S. Sidorenko. 2004d. New genera and species of gall midges (Diptera, Cecidomyiidae) from the 

Russian Far East. International Journal of Dipterological Research 15: 163-185. 

*Fedotova, Z.A. and V.S. Sidorenko. 2005a. New taxa of gall midges from the Russian Far East (Diptera: 

Cecidomyiidae). Zoosystematica Rossica 13: 213-231. 

*Fedotova, Z.A. and V.S. Sidorenko. 2005b. New species of gall midges (Diptera, Cecidomyiidae) of Artemisia 

rubripes Nakai (Asteraceae) [= Compositae] from the Russian Far East. Studia Dipterologica 11: 437-469. 

*Fedotova, Z.A. and V.S. Sidorenko. 2005c. New taxa of the tribe Coquillettomyiini Mamaev, 1968 (Diptera: 

Cecidomyiidae) from the Russian Far East. Russian Entomological Journal 14: 227-238. 

*Fedotova, Z.A. and V.S. Sidorenko. 2005d. New species of gall midges of the supertribe Oligotrophidi (Diptera: 


*Fedotova, Z.A. and V.S. Sidorenko. 2005e. New species of gall midges of the subfamily Porricondylinae from the 

Russian Far East (Diptera: Cecidomyiidae). International Journal of Dipterological Research 16: 89-127. 

*Fedotova, Z.A. and V.S. Sidorenko. 2006a. New genera and species of gall midges from the Russian Far East 

(Diptera: Cecidomyiidae). Entomofauna: Zeitschrift für Entomologie 27: 133-166. 

*Fedotova, Z.A. and V.S. Sidorenko. 2006b. New gall midges of the tribes Brachineurini Edwards, 1937 and 

Stomatosematini Mamaev, 1968 (Diptera: Cecidomyiidae) from the Russian Far East. Far Eastern 


*Fedotova, Z.A. and V.S. Sidorenko. 2006c. [Gall midges (Diptera: Cecidomyioidea) from the Kedrovaya Pad' Nature 

Reserve. Flora and Fauna of Kedrovaya Pad' Nature Reserve.] Pp. 106-139. Russian Academy of Sciences, 

Far Eastern Branch, Institute of Biology and Soil Sciences. Dalnauka, Vladivostok. 

*Fedotova, Z.A. and V.S. Sidorenko. 2006d. New species of gall midges from the genus Brachineura Rondani, 1840 

and new related genera (Diptera: Cecidomyiidae, Brachineurini) from the Russian Far East. International 

Journal of Dipterological Research 17: 77-97. 

*Fedotova, Z.A. and V.S. Sidorenko. 2007a. Nine new species of genus Ledomyia Kieffer, 1895 (Diptera, 


*Fedotova, Z.A. and V.S. Sidorenko. 2007b. New taxa of gall midges from genera related to Parepidosis Kieffer, 1913 

and Porricondyla Rondani, 1840 (Diptera, Cecidomyiidae) from the Russian Far East. International Journal of 

Dipterological Research 18: 15-45. 

*Fedotova, Z.A. and V.S. Sidorenko. 2007c. New taxa of gall midges from tribes Porricondylini, Bryocryptini, 

Asynaptini and Winnertziini (Diptera, Cecidomyiidae, Porricondylinae) from the Russian Far East. 

International Journal of Dipterological Research 18: 69-103. 

*Fedotova, Z.A. and V.S. Sidorenko. 2007d. New taxa of gall midges (Diptera, Cecidomyiidae, Porricondylinae) with 

reduced wing venation and flagellomeres from Russia. International Journal of Dipterological Research 18: 

235-277. 

*Fedotova, Z.A. and V.S. Sidorenko. 2008a. New gall midges related to the genus Monepidosis Mamaev, 1966 

(Diptera, Cecidomyiidae) from the Russian Far East. Russian Entomological Journal 17: 227-238. 

*Fedotova, Z.A. and V.S. Sidorenko. 2008b. New genera and species of gall midges of the tribe Dicerurini from the 

Russian Far East (Diptera, Cecidomyiidae, Porricondylinae). Entomofauna: Zeitschrift für Entomologie 29 

(24): 361-382. 

*Fedotova, Z.A. and V.S. Sidorenko. 2009a. New taxa of gall midges of the supertribe Coquillettomyiidi, stat. nov. 

(Diptera, Cecidomyiidae) from the Russian Far East. International Journal of Dipterological Research 20: 3- 

46. 

*Fedotova, Z.A. and V.S. Sidorenko. 2009b. New taxa of gall midges from the tribes Quadridiplosini and 

Heterodiplosini (Diptera, Cecidomyiidae) from the Russian Far East. Far Eastern Entomologist 196: 1-27. 

*Fedotova, Z.A. and V.S. Sidorenko. 2009c. Gall midges of the tribe Coquillettomyiini s. str. (Diptera, Cecidomyiidae, 

Cecidomyiidi) of Palaearctic Region with descriptions of new taxa. International Journal of Dipterological 

Research 20:51-93. 

*Fedotova, Z.A. and V.S. Sidorenko. 2009d. New taxa of gall midges from the tribe Inclinatidiplosini of the supertribe 

Clinodiplosidi, stat. n. (Diptera, Cecidomyiidae) from the Russian Far East. International Journal of 

Dipterological Research 20: 221-255. 

Felt, E.P. 1906a. Insects affecting park and woodland trees. New York State Museum Memoir 8 (vol. 2): 333-877. 

Felt, E.P. 1906b. Studies in Cecidomyiidae. Pp. 116-134. In his 21st report of the State Entomologist on injurious and

477 

other insects of the State of New York, 1905. New York State Museum Bulletin 104: 49-186, 9 pls. 

Felt, E.P. 1907a. New species of Cecidomyiidae. New York State Education Department, Albany. 53 pp. 

Felt, E.P. 1907b. Cecidomyia acarivora n. sp. Entomological News 18: 242. 

Felt, E.P. 1907c. Appendix: New species of Cecidomyiidae. Pp. 97-165. In his 22d report of the State entomologist on 

injurious and other insects of the State of New York 1906. New York State Museum Bulletin 110: 39-186, 3 

pls. 

Felt, E.P. 1907d. New species of Cecidomyiidae II. New York State Education Department, Albany. 23 pp. 

Felt, E.P. 1908a. Contarinia gossypii n. sp. Entomological News 19: 210-211. 

Felt, E.P. 1908b. Appendix D. Pp. 286-422, 489-510, pls. 33-34. In his 23d report of the State Entomologist on 

injurious and other insects of the State of New York 1907. New York State Museum Bulletin 124: 5-541, 44 

pls. 

Felt, E.P. 1909a. New species of West Indian Cecidomyiidae. Entomological News 20: 299-302. 

Felt, E.P. 1909b. Additional rearings in Cecidomyiidae. Journal of Economic Entomology 2: 286-293. 

Felt, E.P. 1909c. Injurious insects. Pp. 13-40. In his 24th Report of the State Entomologist on injurious and other 

insects of the State of New York 1908. New York State Museum Bulletin 134: 1-206, 17 pls. 

Felt, E.P. 1910a. Two new Cecidomyiidae. Entomological News 21: 10-12. 

Felt, E.P. 1910b. Schizomyia ipomoeae. Entomological News 21: 160-161. 

Felt, E.P. 1910c. West Indian Cecidomyiidae. Entomological News 21: 268-270. 

Felt, E.P. 1910d. Gall midges of Aster, Carya, Quercus and Salix. Journal of Economic Entomology 3: 347-356. 

Felt, E.P. 1911a. A new Lestodiplosis. Entomological News 22: 10-11. 

Felt, E.P. 1911b. Appendix: Miastor americana Felt, an account of pedogenesis. 82-104, 147-174, pls. 22-35. In his 

26th Report of the State Entomologist on injurious and other insects of the State of New York 1910. New 

York State Museum Bulletin 147: 5-180, 35 pls. 

Felt, E.P. 1911c. A generic synopsis of the Itonidae. Journal of the New York Entomological Society 19: 31-62. 

Felt, E.P. 1911d. Two new gall midges (Dipt.). Entomological News 22: 109-111. 

Felt, E.P. 1911e. Endaphis Kieff. in the Americas (Dipt.). Entomological News 22: 128-129. 

Felt, E.P. 1911f. A new species of Lasioptera with observations on certain homologies. Psyche 18: 84-86. 

Felt, E.P. 1911g. Endaphis hirta n. sp. (Dipt.). Entomological News 22: 224. 

Felt, E.P. 1911h. Rhopalomyia grossulariae n. sp. Journal of Economic Entomology 4: 347. 

Felt, E.P. 1911i. Two new gall midges. The Canadian Entomologist 43: 194-196. 

Felt, E.P. 1911j. Four new gall midges (Dipt.). Entomological News 22: 301-305. 

Felt, E.P. 1911k. Three new gall midges (Dipt.). Journal of the New York Entomological Society 19: 190-193. 

Felt, E.P. 1911l. Hosts and galls of American gall midges. Journal of Economic Entomology 4: 451-475. 

Felt, E.P. 1911m. New species of gall midges. Journal of Economic Entomology 4: 476-484. 

Felt, E.P. 1911n. New species of gall midges. Journal of Economic Entomology 4: 546-559. 

Felt, E.P. 1912a. P. 123. In his 27th Report of the State Entomologist on injurious and other insects of the State of New 

York 1911. New York State Museum Bulletin 155: 5-198, 27 pls. 

Felt, E.P. 1912b. New West Indian gall midges (Dipt.). Entomological News 23: 173-177. 

Felt, E.P. 1912c. Lasiopteryx manihot, n. sp. (Diptera). The Canadian Entomologist 44: 144. 

Felt, E.P. 1912d. Diarthronomyia californica n. sp. (Diptera, Itonidae). Pomona College Journal of Entomology 4: 752. 

Felt, E.P. 1912e. Observations on the identity of the wheat midge. Journal of Economic Entomology 5: 286-289. 

Felt, E.P. 1912f. New Itonididae (Dipt.). Journal of the New York Entomological Society 20: 102-107. 

Felt, E.P. 1912g. Itonida inopis O.S. Journal of Economic Entomology 5: 368-369. 

Felt, E.P. 1912h. New gall midges or Itonidae (Dipt.). Journal of the New York Entomological Society 20: 146-156. 

Felt, E.P. 1912i. Observations on Uleela Rubs. (Dipt.). Entomological News 23: 353-354. 

Felt, E.P. 1912j. Arthrocnodax occidentalis n. sp. (Dipt.). Journal of Economic Entomology 5: 402. 

Felt, E.P. 1912k. The gall midge fauna of western North America. Pomona College Journal of Entomology 4: 753-757. 

Felt, E.P. 1912l. Studies in Itonididae. Journal of the New York Entomological Society 20: 236-248. 

Felt, E.P. 1913a. Cystodiplosis eugeniae n. sp. (Dipt.). Entomological News 24: 175-176. 

Felt, E.P. 1913b. Itonida anthici n. sp. (Dipt.). Journal of Economic Entomology 6: 278-279. 

Felt, E.P. 1913c. Appendix: A study of gall midges. Pp. 127-226, 247-254, pls. 11-14. In his 28th Report of the State 

Entomologist on injurious and other insects of the State of New York 1912. New York State Museum Bulletin 

165: 5-265, 14 pls.

478 

Felt, E.P. 1913d. Descriptions of gall midges. Journal of the New York Entomological Society 21: 213-219. 

Felt, E.P. 1913e. Three new gall midges (Diptera). The Canadian Entomologist 45: 304-308. 

Felt, E.P. 1913f. The gall midge fauna of New England. Psyche 20: 133-147. 

Felt, E.P. 1913g. Adaptation in the gall midges. The Canadian Entomologist 45: 371-379. 

Felt, E.P. 1913h. Two new Canadian gall midges. The Canadian Entomologist 45: 417-419. 

Felt, E.P. 1913i. Arthrocnodax carolina n. sp. Journal of Economic Entomology 6: 488-489. 

Felt, E.P. 1913j. Didactylomyia capitata n. sp. Psyche 20: 174. 

Felt, E.P. 1914a. Acaroletes pseudococci n. sp. Journal of Economic Entomology 6: 148-149. 

Felt, E.P. 1914b. Diadiplosis coccidivora, n. sp. The Entomologist 47: 86. 

Felt, E.P. 1914c. Descriptions of gall midges. Journal of the New York Entomological Society 22: 124-134. 

Felt, E.P. 1914d. Additions to the gall midge fauna of New England. Psyche 21: 109-114. 

Felt, E.P. 1914e. Aplonyx sarcobati n. sp. Pomona College Journal of Entomology 6: 93-94. 

Felt, E.P. 1914f. New gall midges (Itonididae). Insecutor Inscitiae Menstruus 2: 117-123. 

Felt, E.P. 1914g. Hormomyia bulla, n. sp. The Canadian Entomologist 46: 286-287. 

Felt, E.P. 1914h. Gall midges as forest insects. The Ottawa Naturalist 28: 76-79. 

Felt, E.P. 1914i. Arthrocnodax constricta n. sp. Journal of Economic Entomology 7: 481. 

Felt, E.P. 1915a. New genera and species of gall midges. Proceedings of the United States National Museum 48: 

195-211. 

Felt, E.P. 1915b. Mycodiplosis macgregori n. sp. Journal of Economic Entomology 8: 149. 

Felt, E.P. 1915c. A new pest, the chrysanthemum midge (Rhopalomyia hypogaea H. Lw.). Journal of Economic 

Entomology 8: 267. 

Felt, E.P. 1915d. Appendix: A study of gall midges II. Pp. 79-242, pls. 1-14. In his 29th Report of the State 


175: 5-257, 16 pls. 

Felt, E.P. 1915e. New North American gall midges. The Canadian Entomologist 47: 226-232. 

Felt, E.P. 1915f. New gall midges. Journal of Economic Entomology 8: 405-409. 

Felt, E.P. 1915g. New Asian gall midges. Journal of the New York Entomological Society 23: 173-184. 

Felt, E.P. 1915h. New South American gall midges. Psyche 22: 152-157. 

Felt, E.P. 1915i. The gall midges of the pine. Bulletin of the Brooklyn Entomological Society 10: 74-76. 

Felt, E.P. 1915j. Appendix: A study of gall midges III. Pp. 127-288, 295-326, pls. 4-19. In his 30th Report of the State 


180: 7-336, 19 pls. 

Felt, E.P. 1915k. Pp. 1-116 (as editor). In Thompson, M.T. An Illustrated Catalogue of American Insect Galls. E.P. 

Felt, ed. Nassau, New York. 116 pp. 

Felt, E.P. 1916a. New gall midges. The Canadian Entomologist 48: 29-34. 

Felt, E.P. 1916b. Pp. 268-269. In Woods, W.C. Blueberry insects in Maine. Maine Agricultural Experiment Station 

Bulletin (1915) Ser. 2, 244: 249-288. 

Felt, E.P. 1916c. Gall midges of certain Chenopodiaceae (Dip.). Entomological News 27: 201-203. 

Felt, E.P. 1916d. Appendix: A study of gall midges IV. Pp. 101-172, 199-203, pls. 14-18. In his 31st Report of the 

State Entomologist on injurious and other insects of the State of New York 1915. New York State Museum 

Bulletin 186: 7-215, 18 pls. 

Felt, E.P. 1916e. New western gall midges. Journal of the New York Entomological Society 24: 175-196. 

Felt, E.P. 1916f. New North American gall midges (Dipt.). Entomological News 27: 412-417. 

Felt, E.P. 1916g. New Indian gall midges. The Canadian Entomologist 48: 400-406. 

Felt, E.P. 1917a. New Indian gall midges (Diptera). Entomological News 28: 73-76. 

Felt, E.P. 1917b. New gall midges. Journal of the New York Entomological Society 25: 193-196. 

Felt, E.P. 1917c. Indian gall midges (Cecidomyiidae, Dipt.). Entomological News 28: 369-372. 

Felt, E.P. 1917d. Asphondylia websteri n. sp. Journal of Economic Entomology 10: 562. 

Felt, E.P. 1918a. Appendix: A study of gall midges V. Pp. 101-252, 257-268, pls. 3-8. In his 32nd Report of the State 


198: 7-276, 8 pls. 

Felt, E.P. 1918b. New gall midges (Dipt.). Journal of Economic Entomology 11: 380-384. 

Felt, E.P. 1918c. New Philippine gall midges, with a key to the Itonididae. The Philippine Journal of Science 13:

479 

281-325, pl. I. 

Felt, E.P. 1918d. Key to American insect galls. New York State Museum Bulletin 200: 5-310, 16 pls. 

Felt, E.P. 1918e. Appendix: A study of gall midges VI. Pp. 76-205, 213-230, pls. 4-12. In his 33rd Report of the State 


202: 7-240, 12 pls. 

Felt, E.P. 1919a. New Philippine gall midges. The Philippine Journal of Science 14: 287-294. 

Felt, E.P. 1919b. Five non-gall-making midges (Dip., Cecidomyidae). Entomological News 30: 219-223. 

Felt, E.P. 1920a. New gall midges or Itonididae from the Adirondacks. Journal of the New York Entomological Society 

27 (1919): 277-292. 

Felt, E.P. 1920b. New Indian gall midges. Memoirs of the Department of Agriculture in India 7: 1-11. 

Felt, E.P. 1920c. Four new African gall midges. Annals of the Natal Museum 4: 491-496. 

Felt, E.P. 1920d. New Philippine gall midges. The Philippine Journal of Science 17: 231-234. 

Felt, E.P. 1921a. Appendix: A study of gall midges, VII. Pp. 81-240, 255-280, pls. 8-20. In his 34th Report of the State 


231-232: 7-288, 20 pls. 

Felt, E.P. 1921b. New species of reared gall midges (Itonididae). Journal of the New York Entomological Society 29: 

115-118. 

Felt, E.P. 1921c. Three new sub-tropical gall midges (Itonididae, Dipt.). Entomological News 32: 141-143. 

Felt, E.P. 1921d. Observations of Johnsonomyia Felt with a description of a new species. The Canadian Entomologist 

53: 96. 

Felt, E.P. 1921e. P. 149 in Weiss, H.B. and E. West. Notes on the insects of spreading dogbane, Apocynum 

androsaemifolium L., with a description of a new dogbane midge. The Canadian Entomologist 53: 146-150. 

Felt, E.P. 1921f. Indian grass gall midges. Memoirs of the Department of Agriculture in India, Entomological Series 7: 

15-22. 

Felt, E.P. 1921g. The number of antennal segments in gall midges and a new species. Bulletin of the Brooklyn 

Entomological Society 16: 93-95. 

Felt, E.P. 1921h. Javanese gall midges. Treubia 1: 139-151. 

Felt, E.P. 1921i. A new Javanese gall midge (Trishormomyia pandani n. sp.). Treubia 1: 270-271. 

Felt, E.P. 1921j. New Indian gall midges (Itonididae). Memoirs of the Department of Agriculture in India, 

Entomological Series 7: 23-28. 

Felt, E.P. 1921k. New Javanese gall midges. Treubia 2: 89-92. 

Felt, E.P. 1921l. A new Diadiplosis. Zoologica (Scientific Contributions of the New York Zoological Society) 3: 

225-226. 

Felt, E.P. 1922a. Mycodiplosis moznettii n. sp. The Florida Entomologist 5: 46. 

Felt, E.P. 1922b. A new and remarkable fig midge. The Florida Entomologist 6: 5-6. 

Felt, E.P. 1922c. A new gall midge on rushes. Entomological News 33: 166-168. 

Felt, E.P. 1922d. A new cecidomyiid parasite of the white fly. Proceedings of the United States National Museum 61: 

1-2. 

Felt, E.P. 1925a. New gall midges. Journal of Entomology and Zoology 17: 15. 

Felt, E.P. 1925b. Pp. 19-20. In Needham, J.G. Observations on a flower gall of the chaparral. Journal of Entomology 

and Zoology 17: 17-20. 

Felt, E.P. 1925c. Key to gall midges (A resumé of studies I-VII, Itonididae). New York State Museum Bulletin 257: 

3-239, 8 pls. 

Felt, E.P. 1926a. P. 85. In Driggers, B.F. The blueberry tip worm (Contarinia vaccinii Felt.), a new species of midge 

attacking cultivated blueberries. Journal of the New York Entomological Society 34: 82-85. 

Felt, E.P. 1926b. A new predaceous midge on roses (Dipt.: Cecidomyiidae). Entomological News 37: 141. 

Felt, E.P. 1926c. New gall midges from New England. Occasional Papers of the Boston Society of Natural History 5: 

207-208. 

Felt, E.P. 1926d. A new Formosan gall-midge (Diptera, Itonididae [Cecidomyiidae]). The Entomologist's Monthly 

Magazine 62: 183-184. 

Felt, E.P. 1926e. A new spruce gall midge (Itonididae). The Canadian Entomologist 58: 229-230. 

Felt, E.P. 1926f. New non-gall making Itonididae (Diptera). The Canadian Entomologist 58: 265-268. 

Felt, E.P. 1926g. Three western gall midges. Journal of Entomology and Zoology 18: 79-81.

480 

Felt, E.P. 1926h. New African gall midges. Entomology Memoirs, Department of Agriculture, Union of South Africa 

5: 44-45. 

Felt, E.P. 1926i. New species of Indian gall midges (Itonididae). Memoirs of the Department of Agriculture in India 9: 

241-245. 

Felt, E.P. 1927a. Four new Indian gall midges. Memoirs of the Department of Agriculture in India 10: 1-4. 

Felt, E.P. 1927b. A new seed head midge. Treubia 9: 381. 

Felt, E.P. 1927c. Two remarkable East Indian gall midges. Treubia 9: 382-384. 

Felt, E.P. 1927d. New species of East Indian gall midges. Treubia 9: 385-389. 

Felt, E.P. 1928a. A new African gall midge. Journal of the New York Entomological Society 36: 123-124. 

Felt, E.P. 1928b. A new western gall midge. Journal of Entomology and Zoology 20: 58. 

Felt, E.P. 1931. Gall midges or gall gnats of the Orient (Itonididae or Cecidomyiidae). Lingnan Science Journal 7 

(1931): 413-474. The journal issue shows "1928" on the cover, but the inside cover has printed in small 

letters: "Issued Mar. 1931." The date on a separate of the article also shows "1928," reflecting the cover of the 

journal issue. There is no evidence that the separate is a preprint. 

Felt, E.P. 1932a. A new Japanese gall midge. Bulletin of the Brooklyn Entomological Society 27: 124. 

Felt, E.P. 1932b. A new predaceous gall midge for California. Pan-Pacific Entomologist 8: 167-168. 

Felt, E.P. 1932c. A new citrus cambium miner from Puerto Rico. Journal of the Department of Agriculture of Puerto 

Rico 16: 117-118. 

Felt, E.P. 1933a. A new enemy of the pineapple mealybug and a list of gall midge enemies of mealybugs. Journal of the 

New York Entomological Society 41: 87-89. 

Felt, E.P. 1933b. A hibiscus bud midge new to Hawaii. Proceedings of the Hawaiian Entomological Society 8: 

247-248. 

Felt, E.P. 1933c. A new Lestodiplosis. Psyche 40: 113-114. 

Felt, E.P. 1934a. New gall midges. Bulletin of the Brooklyn Entomological Society 29: 77-78. 

Felt, E.P. 1934b. A new gall midge on fig (Diptera: Itonididae). Entomological News 45: 131-133. 

Felt, E.P. 1934c. Two western species of Asphondylia. Journal of Entomology and Zoology 26: 34. 

Felt, E.P. 1935a. New species of gall midges from Texas. Journal of the Kansas Entomological Society 8: 1-8. 

Felt, E.P. 1935b. A new gall midge. Journal of the New York Entomological Society 43: 47. 

Felt, E.P. 1935c. A gall midge on pine cones. Journal of the New York Entomological Society 43: 48-49. 

Felt, E.P. 1935d. Trisopsis in the United States (Dipt., Itonididae or Cecidomyiidae). Entomological News 46: 75-77. 

Felt, E.P. 1935e. A new melon gall midge. Bulletin of the Brooklyn Entomological Society 30: 79-80. 

Felt, E.P. 1936a. New midges on pine and grass. Journal of the New York Entomological Society 44: 7-9. 

Felt, E.P. 1936b. Two new cockle burr midges (Diptera: Cecidomyiidae). Entomological News 47: 231-233. 

Felt, E.P. 1938. A new species of gall midge predacious on mealybugs. Proceedings of the Hawaiian Entomological 

Society 10: 43. 

Felt, E.P. 1939a. A new gall midge on rhododendron. Journal of the New York Entomological Society 47: 41-42. 

Felt, E.P. 1939b. A new juniper gall midge (Diptera: Cecidomyiidae). Entomological News 50: 159-160. 

Felt, E.P. 1940. Plant Galls and Gall Makers. Comstock Publishing Co., Ithaca, New York. viii & 364 pp. 

*Fernandes, G.W., J.C. Santos, F.M.C. Castro and A. Castilho. 2007. Impact of a gall midge Parkiamyia paraensis 

(Diptera, Cecidomyiidae) on the Amazonian plant Parkia pendula (Fabaceae). Revista Brasileira de 

Entomologia 51: 471-475. 

Fitch, A. 1845a. Insects injurious to vegetation. - No. 2. Insects of the genus Cecidomyia, including the Hessian and 

wheat-fly. American Quarterly Journal of Agriculture and Science 1: 255-269, pl. II. 

Fitch, A. 1845b. Insects injurious to vegetation. - No. 3. The wheat-fly. American Quarterly Journal of Agriculture and 

Science 2: 233-264, pl. 5. 

Fitch, A. 1845c. An essay upon the wheat-fly, and some species allied to it. Carroll and Cook, Albany. 32 pp., 1 pl. 

Fitch, A. 1855. Report. Transactions of the New York State Agricultural Society 14 (1854): 705-880. 

Fitch, A. 1859. Fifth report on the noxious and other insects of the State of New York. Transactions of the New York 

State Agricultural Society 18 (1858): 781-854. 

Fitch, A. 1861. Sixth report on the noxious and other insects of the State of New York. Transactions of the New York 

State Agricultural Society 20 (1860): 745-868. 

Fleur, E. 1929. Monsieur l'Abbé Kieffer, Professeur au Collège de Bitche (Moselle). Bulletin de la Société d'Histoire 

Naturelle de la Moselle 8: 7-29, 2 unnumbered pls.

481 

Foote, R.H. 1953. A new gall midge infesting holly (Diptera: Itonididae). Entomological News 64: 197-201. 

Foote, R.H. 1956. Gall midges associated with cones of western forest trees (Diptera: Itonididae). Journal of the 

Washington Academy of Sciences 46: 48-57. 

Foote, R.H. 1965. Family Cecidomyiidae (Itonididae). Pp 241-295. In A. Stone, C.W. Sabrosky, W.W. Wirth, R.H. 

Foote, and J.R. Coulson, eds. A catalog of the Diptera of America north of Mexico. United States Department 

of Agriculture Handbook 276: i-iv, 1-1696. [Reprinted in 1983 by Smithsonian Institution Press, Washington, 

D.C.] 

Foote, R.H. and C.A. Thomas. 1959. Mycophila fungicola Felt: a redescription and review of its biology (Diptera, 

Itonididae). Annals of the Entomological Society of America 52: 331-334. 

Fourcroy, A.F. de. 1785. Entomologia Parisiensis; sive Catalogus Insectorum quae in Agro Parisiensi Reperiuntur; 

Secundum Methodum Geoffraeanam in Sectiones, Genera et Species Distributus: Qui Addita Funt Nomina 

Trivialia & Fere Trecentae Novae Species. Paris. viii & 544 pp. 

Frauenfeld, G. 1859. Ueber exotische Pflanzenauswüchse, erzeugt von Insekten. Verhandlungen der kaiserlichköniglichen 

zoologisch-botanischen Gesellschaft in Wien 9: 319-332, pls. VI-VII. 

Frauenfeld, G. 1861. Beitrag zur Kenntniss der Insekten-Metamorphose aus dem Jahre 1860. Verhandlungen der 

kaiserlich-königlichen zoologisch-botanischen Gesellschaft in Wien 11: 163-174, pl. II. D. 

Frauenfeld, G. 1862. Beitrag zur Insektengeschichte aus dem Jahre 1861. Verhandlungen der kaiserlich-königlichen 

zoologisch-botanischen Gesellschaft in Wien 12: 1171-1178. 

Frauenfeld, G. 1863. Beitrag zur Metamorphosengeschicte aus dem Jahre 1862. Verhandlungen der kaiserlichköniglichen 


Frauenfeld, G. 1867. Zoologische Miscellen. XII. Verhandlungen der kaiserlich-königlichen zoologisch-botanischen 

Gesellschaft in Wien 17: 775-784. 

Frauenfeld, G. 1873. Zoologische Miscellen. Verhandlungen der kaiserlich-königlichen zoologisch-botanischen 

Gesellschaft in Wien 23: 183-192, pl. II.B. 

Frisch, J.L. 1820. XX. Von den kleinen schlänglichen Maden, in allerlen feuchten faulenden Sachen, und den Fliegen, 

so daraus werden. Pp. 37-38, pl. IV, fig. X. In Beschreibung von allerlen Insecten in Deutschland, nebst 

nützlichen Anmerkungen und nöthigen Abbildungen von diesem kriechenden und fliegenden inländichen 

Gewürme. Vierter Theil. G.G.Nicolai, Berlin. 

Fyles, T.W. 1883. Description of a dipterous parasite of Phylloxera vastatrix. The Canadian Entomologist 14: 237-239. 

Gagné, R.J. 1966a. A new species of Prodiplosis (Diptera: Cecidomyiidae). Annals of the Entomological Society of 

America 59: 1154-1157. 

Gagné, R.J. 1966b. The Nearctic species of Contarinia which infest grasses (Diptera: Cecidomyiidae). Proceedings of 

the Entomological Society of Washington 68: 318-320. 

Gagné, R.J. 1967a. The genus Oligotrophus Latreille (Diptera: Cecidomyiidae) in North America and a new species 

injurious to Betula papyrifera Marsh. Entomological News 78: 129-134. 

Gagné, R.J. 1967b. Tinconaria albescentis, a new North American genus and species of Cecidomyiidae found in seeds 

of Triodia albescens Vasey (Diptera). Proceedings of the Entomological Society of Washington 69: 155-157. 

Gagné, R.J. 1967c. Notes on the genus Thecodiplosis Kieffer in North America and some generic reassignments 

(Diptera: Cecidomyiidae). Proceedings of the Entomological Society of Washington 69: 338-339. 

Gagné, R.J. 1968a. Chrybaneura harrisoni, a new genus and species from Central America (Diptera: Cecidomyiidae). 


Gagné, R.J. 1968b. A taxonomic revision of the genus Asteromyia (Diptera: Cecidomyiidae). Miscellaneous 

Publications of the Entomological Society of America 6: 1-40. 

Gagné, R.J. 1968c. Authorship of Pachydiplosis oryzae, a rice pest of Asia and Africa (Diptera: Cecidomyiidae). 

Proceedings of the Entomological Society of Washington 70: 136. 

Gagné, R.J. 1968d. Family Cecidomyiidae. Fascicle 23: 1-62. In Papavero, N., ed. A Catalogue of the Diptera of the 

Americas South of the United States. Departamento de Zoologia, Secretaria da Agricultura, São Paulo. 

Gagné, R.J. 1968e. Revision of the gall midges of bald cypress (Diptera: Cecidomyiidae). Entomological News 79: 

269-274. 

Gagné, R.J. 1969a. A review of the genus Walshomyia including a new species reared from Cupressus galls in 

California (Diptera: Cecidomyiidae). The Pan-Pacific Entomologist 45: 16-19. 

Gagné, R.J. 1969b. A new genus and two new species of Cecidomyiidae associated with Pariana spp. (Gramineae) in 

South America (Diptera). Proceedings of the Entomological Society of Washington 71: 108-111.

482 

Gagné, R.J. 1969c. New synonyms of Asphondylia betheli Cockerell (Diptera: Cecidomyiidae). Entomological News 

80: 176. 

Gagné, R.J. 1969d. A tribal and generic revision of the Nearctic Lasiopteridi (Diptera: Cecidomyiidae). Annals of the 

Entomological Society of America 62: 1348-1364. 

Gagné, R.J. 1969e. A review of Felt's Philippine genera and species of Cecidomyiidae (Diptera). Pacific Insects 11: 

561-570. 

Gagné, R.J. 1970a. A new genus and new species of Cecidomyiidae on pinyon pine (Diptera). Entomological News 81: 

153-156. 

Gagné, R.J. 1970b. A family reassignment for Moehnia Pritchard (Diptera: Sciaridae). Journal of the Washington 

Academy of Sciences 60: 60. 

Gagné, R.J. 1971a. Two new species of North American Neolasioptera from Baccharis (Diptera: Cecidomyiidae - 

Compositae). Proceedings of the Entomological Society of Washington 73: 153-157. 

Gagné, R.J. 1971b. Taxonomic notes on two racemicolous gall-makers on Solidago spp. (Diptera: Cecidomyiidae - 

Compositae). Proceedings of the Entomological Society of Washington 73: 169. 

Gagné, R.J. 1971c. The genus Aphidoletes Kieffer (Diptera: Cecidomyiidae) in North America. Entomological News 

82: 177-181. 

Gagné, R.J. 1971d. Pp. 335-338. In Gagné, R.J. and C.E. Stegmaier, Jr. A new species of Chilophaga on Aristida 

(Gramineae) in Florida (Diptera: Cecidomyiidae). The Florida Entomologist 54: 335-338. 

Gagné, R.J. 1972a. Craneiobia lawsonianae De Meijere, new synonym of Janetiella siskiyou Felt (Cecidomyiidae: 

Diptera). Proceedings of the Entomological Society of Washington 74: 128. 

Gagné, R.J. 1972b. New synonymy and homonymy in Cecidomyiidae (Diptera). Proceedings of the Entomological 

Society of Washington 74: 321-326. 

Gagné, R.J. 1972c. A new species of Contarinia (Diptera: Cecidomyiidae) from Helianthus annuus L. (Compositae) in 

North America. Entomological News 83: 279-281. 

Gagné, R.J. 1973a. Family Cecidomyiidae. Pp. 480-517. In Delfinado, M.D. and D.E. Hardy. A Catalog of the Diptera 

of the Oriental Region. Volume I. Order Nematocera. The University Press of Hawaii, Honolulu. 

Gagné, R.J. 1973b. Cecidomyiidae from Mexican Tertiary amber (Diptera). Proceedings of the Entomological Society 

of Washington 75: 169-171. 

Gagné, R.J. 1973c. A generic synopsis of the Nearctic Cecidomyiidi (Diptera: Cecidomyiidae: Cecidomyiinae). Annals 

of the Entomological Society of America 66: 857-889. 

Gagné, R.J. 1973d. A review of Karschomyia Felt with descriptions of seven new Nearctic species (Diptera: 

Cecidomyiidae). Proceedings of the Entomological Society of Washington 75: 345-354. 

Gagné, R.J. 1973e. A new species of Cecidomyiidae (Diptera) from Nothofagus in Chile. Studia Entomologica 16: 

447-448. 

Gagné, R.J. 1975a. A revision of the Nearctic Stomatosematidi (Diptera: Cecidomyiidae: Cecidomyiinae). Annals of 

the Entomological Society of America 68: 86-90. 

Gagné, R.J. 1975b. The gall midges of ragweed, Ambrosia, with descriptions of two new species (Diptera: 


Gagné, R.J. 1975c. A redefinition of Diarthronomyia Felt as a subgenus of Rhopalomyia Rübsaamen (Diptera: 

Cecidomyiidae: Oligotrophidi). Annals of the Entomological Society of America 68: 482-484. 

Gagné, R.J. 1975d. A review of the Nearctic genera of Oligotrophidi with piercing ovipositors (Diptera: 

Cecidomyiidae). Entomological News 86: 5-12. 

Gagné, R.J. 1975e. A new species of Mayetiola (Diptera: Cecidomyiidae) on American beachgrass in New Jersey. 

Entomological News 86: 85-87. 

Gagné, R.J. 1976. New Nearctic records and taxonomic changes in the Cecidomyiidae (Diptera). Annals of the 


Gagné, R.J. 1977a. Cecidomyiidae (Diptera) from Canadian amber. Proceedings of the Entomological Society of 

Washington 79: 57-62. 

Gagné, R.J. 1977b. The Cecidomyiidae (Diptera) associated with Chromolaena odorata (L.) K. & R. (Compositae) in 

the Neotropical Region. Brenesia 12/13: 113-131. 

Gagné, R.J. 1978a. Pp. 85-88. In Osgood, E.A. and R.J. Gagné. Biology and taxonomy of two gall midges (Diptera: 

Cecidomyiidae) found in galls on balsam fir needles with description of a new species of Paradiplosis. Annals 

of the Entomological Society of America 71: 85-91.

483 

Gagné, R.J. 1978b. A new species of Rhopalomyia (Diptera: Cecidomyiidae) from Artemisia ludoviciana Nuttall 

(Compositae) in Utah. Great Basin Naturalist 37: 426-428. 

Gagné, R.J. 1978c. A new species of Asphondylia (Diptera: Cecidomyiidae) from Costa Rica with taxonomic notes on 

related species. Proceedings of the Entomological Society of Washington 80: 514-516. 

Gagné, R.J. 1978d. New synonymy and a review of Haplusia (Diptera: Cecidomyiidae). Proceedings of the 


Gagné, R.J. 1978e. A new species of Cecidomyia injurious to cones of slash pine in Florida. The Florida Entomologist 

61: 193-195. 

Gagné, R.J. 1978f. A systematic analysis of the pine pitch midges, Cecidomyia spp. (Diptera: Cecidomyiidae). U.S. 

Department of Agriculture Technical Bulletin 1575: 1-18. 

Gagné, R.J. 1979. A new species of Neostenoptera from the Congo (Diptera: Cecidomyiidae). Annales de la Société 

Entomologique de France (n.s.) 15: 345-347. 

Gagné, R.J. 1981a. A new species of North American Cecidomyiidae (Diptera) from Spartina alterniflora (Poaceae). 


Gagné, R.J. 1981b. Cecidomyiidae. Pp. 257-292. In McAlpine, J.F., B.V. Peterson, G.E. Shewell, H.J. Teskey, J.R. 

Vockeroth, and D.M. Wood (eds.). Manual of Nearctic Diptera. Vol. 1. Research Branch, Agriculture. Canada 

Monograph No. 27. 

Gagné, R.J. 1981c. A new species of Endaphis (Diptera: Cecidomyiidae) endoparasitic in aphids in British Columbia. 


Gagné, R.J. 1983a. Celticecis (Diptera: Cecidomyiidae), a new genus for gall makers on hackberries, Celtis spp. 

(Ulmaceae). Proceedings of the Entomological Society of Washington 85: 435-438. 

Gagné, R.J. 1983b. Pp. 13-66. In Jones, R.G., R.J. Gagné and W.F. Barr. Biology and taxonomy of the Rhopalomyia 

gall midges (Diptera: Cecidomyiidae) of Artemisia tridentata Nuttall (Compositae) in Idaho. Contributions if 

the American Entomological Institute 21: [2+] 1-79 [+11]. 

Gagné, R.J. 1983c. Appendix. Pp. 67-72. In Jones, R.G., R.J. Gagné and W.F. Barr. Biology and taxonomy of the 

Rhopalomyia gall midges (Diptera: Cecidomyiidae) of Artemisia tridentata Nuttall (Compositae) in Idaho. 

Contributions if the American Entomological Institute 21: [2+] 1-79 [+11]. 

Gagné, R.J. 1983d. A new species of Cecidomyiidae (Diptera) injurious to Protea flowers in South Africa. Annals of 

the Natal Museum 25: 587-589. 

Gagné, R.J. 1983e. Acericecis Gagné, a new genus for Cecidomyia ocellaris Osten Sacken (Diptera: Cecidomyiidae), 

the maple leaf ocellate gall maker in North America. Proceedings of the Entomological Society of Washington 

85: 704-709. 

Gagné, R.J. 1984a. Pp. 222-227. In Gagné, R.J. and G.M. Beavers. Contarinia spp. (Diptera: Cecidomyiidae) from 

shoots of slash pine (Pinus elliottii Engelm.) with the description of a new species injurious to needles. The 

Florida Entomologist 67: 221-228. 

Gagné, R.J. 1984b. Pp. 544-548. In Gagné, R.J. and K. Valley. Two new species of Cecidomyiidae (Diptera) from 

honeylocust, Gleditsia triacanthos L. (Fabaceae), in eastern United States. Proceedings of the Entomological 


Gagné, R.J. 1984c. Five new species of Neotropical Cecidomyiidae (Diptera) associated with cacao flowers, killing the 

buds of Clusiaceae, or preying on mites. Brenesia 22: 123-138. 

Gagné, R.J. 1985a. Descriptions of new Nearctic Cecidomyiidae (Diptera) that live in xylem vessels of fresh-cut wood, 

and a review of Ledomyia (s.str.). Proceedings of the Entomological Society of Washington 87: 116-134. 

Gagné, R.J. 1985b. A taxonomic revision of the Asian rice gall midge, Orseolia oryzae (Wood-Mason), and its 

relatives (Diptera: Cecidomyiidae). Entomography 3: 127-162. 

Gagné, R.J. 1986a. A new Nearctic species of Contarinia (Diptera: Cecidomyiidae) recently introduced into Hawaii on 

Cupressus (Cupressaceae). Proceedings of the Entomological Society of Washington 88: 127-130. 

Gagné, R.J. 1986b. Revision of Prodiplosis (Diptera: Cecidomyiidae) with descriptions of three new species. Annals of 


Gagné, R.J. 1986c. The correct type species of Peridiplosis (Diptera: Cecidomyiidae). Proceedings of the 


Gagné, R.J. 1986d. Pp. 65-71.In Hawkins, B.A., R.D. Goeden and R.J. Gagné. Ecology and taxonomy of the 

Asphondylia spp. (Diptera: Cecidomyiidae) forming galls on Atriplex spp. (Chenopodiaceae) in southern 

California. Entomography 4: 55-107.

484 

Gagné, R.J. 1987a. Pp 502-504. In Larew, H.G, R.J. Gagné, and A.Y. Rossman. Fungal gall caused by a new species of 

Ledomyia (Diptera: Cecidomyiidae) on Xylaria enterogena (Ascomycetes: Xylariaceae). Annals of the 


Gagné, R.J. 1987b. Pp. 1-12. In Gagné, R.J. and M.L.A. Rios de Saluso. Una nueva especie de Mycodiplosis (Diptera: 

Cecidomyiidae) associada a la roya amarilla de Avena sativa (Poaceae) en Argentina. INTA Estación 

Experimental Agropecuaria Paraná Entre Ríos - Republica Argentina, Serie Técnica No. 54: 1-12. 

Gagné, R.J. 1988. A polynomial Riley name in Cecidomyiidae (Diptera) and implications of such names for Cynipidae 

(Hymenoptera). Proceedings of the Entomological Society of Washington 90: 394-395. 

Gagné, R.J. 1989a. Pp. 170-173. In Gagné, R.J. and P.E. Boldt. A new species of Neolasioptera (Diptera: 

Cecidomyiidae) from Baccharis (Asteraceae) in southern United States and the Dominican Republic. 


Gagné, R.J. 1989b. The Plant-Feeding Gall Midges of North America. Cornell University Press. xi & 356 pp. 

Gagné, R.J. 1989c. Family Cecidomyiidae. Pp. 152-163. In Evenhuis, N.L., ed. Catalog of the Diptera of the 

Australasian and Oceanian Regions. Bishop Museum Special Publication 86. Bishop Museum Press and E.J. 

Brill. 

Gagné, R.J. 1990a. Pp. 146-151. In Gagné, R.J. and R.W. Duncan. A new species of Cecidomyiidae (Diptera) 

damaging shoot tips of yellow cypress, Chamaecyparis nootkatensis, and a new genus for two gall midges on 

Cupressaceae. Proceedings of the Entomological Society of Washington 92: 146-151. 

Gagné, R.J. 1990b. Gall midge complex (Diptera: Cecidomyiidae) in bud galls of Palearctic Euphorbia 

(Euphorbiaceae). Annals of the Entomological Society of America 83: 335-345. 

Gagné, R.J. 1990c. Pp. 652-670. In Gagné, R.J. and G.L. Waring. The Asphondylia (Cecidomyiidae: Diptera) of 

creosote bush (Larrea tridentata) in North America. Proceedings of the Entomological Society of Washington 

92: 649-671. 

Gagné, R.J. 1991. Bayeriola Gagné, new name for Bayeria Rübsaamen 1914 (Diptera: Cecidomyiidae), preoccupied by 

Fritsch 1905 (Pisces). Proceedings of the Entomological Society of Washington 93: 791. 

Gagné, R.J. 1992a. Pp. 87-90. In Gagné, R.J. and J.A. Payne. A new species of Putoniella (Diptera: Cecidomyiidae) 

damaging leaves of Prunus spp. (Rosaceae) in southeastern United States. Journal of Entomological Science 

27: 85-91. 

Gagné, R.J. 1992b. Pp. 385-390. In Gagné, R.J. and J.A. Payne. A new species of Harmandia (Diptera: 

Cecidomyiidae) damaging leaves of Allegheny chinkapin in eastern United States and a redescription of the 

genus. Journal of Entomological Science 27: 383-391. 

Gagné, R.J. 1993a. Pp. 342-347. In Gagné, R.J. and F.D. Bennett. Two new species of Lestodiplosini (Diptera: 

Cecidomyiidae) preying on Homoptera and Thysanoptera in southern Florida. The Florida Entomologist 76: 

341-348. 

Gagné, R.J. 1993b. Pp. 542-545. In Gagné, R.J. and D.R. Strong. A new species of Dasineura (Diptera: 

Cecidomyiidae) galling leaves of Lupinus spp. (Fabaceae) in California. Proceedings of the Entomological 


Gagné, R.J. 1993c. Pp. 84-123. In Gagné, R.J. and J. Marohasy. The gall midges (Diptera: Cecidomyiidae) of Acacia 

spp. (Mimosaceae) in Kenya. Insecta Mundi 7: 77-124. 

Gagné, R.J. 1994a. The Gall Midges of the Neotropical Region. Cornell University Press, Ithaca, New York. xv & 352 

pp. 

Gagné, R.J. 1994b. Pp. 524-529. In Tang, Y.Q., R.K. Yokomi and R.J. Gagné. Life history and description of Endaphis 

maculans (Diptera: Cecidomyiidae), an endoparasitoid of aphids in Florida and the Caribbean Basin. Annals 

of the Entomological Society of America 87: 523-531. 

Gagné, R.J. 1994c. Pp. 601-605. In Gagné, R.J. and J.A. Payne. Vaccinidiplosis, a new genus for Cecidomyia vaccinii 

Osten Sacken (Diptera: Cecidomyiidae). Proceedings of the Entomological Society of Washington 96: 

599-606. 

Gagné, R.J. 1995a. Revision of tetranychid (Acarina) mite predators of the genus Feltiella (Diptera: Cecidomyiidae). 


Gagné, R.J. 1995b. Contarinia maculipennis (Diptera: Cecidomyiidae), a polyphagous pest newly reported for the 

Americas. Bulletin of Entomological Research 85: 209-214. 

Gagné, R.J. 1995c. Pp. 768-777. In Gagné, R.J and P.E. Boldt. The gall midges (Diptera: Cecidomyiidae) of Baccharis 

spp. (Asteraceae) in the United States. Proceedings of the Entomological Society of Washington 97: 767-778.

485 

Gagné, R.J. 1995d. First record of Conarete (Diptera: Cecidomyiidae: Lestremiinae) in the Afrotropical Region, and 

notes on the loss of ocelli in the tribe Lestremiini. Annals of the Natal Museum 36: 141-145. 

Gagné, R.J. 1996a. A new species of Dasineura (Diptera: Cecidomyiidae) infesting pinxterbloom azalea flowers in 

Maryland and Washington, D.C. Proceedings of the Entomological Society of Washington 98: 228-232. 

Gagné, R.J. 1996b. Pp. 429-433. In Gagné, R.J. and K.L. Hibbard. A new species of gall midge (Diptera: 

Cecidomyiidae) from subterranean stem galls of Licania michauxii (Chrysobalanaceae) in Florida. The Florida 


Gagné, R.J. 1996c. Pp. 55-68. In Gagné, R.J., R. Sobhian and N. Isidoro. A review of the genus Psectrosema (Diptera: 

Cecidomyiidae), Old World pests of Tamarix (Tamaricaceae), and description of three new species. Israel 

Journal of Entomology 30: 53-69. 

Gagné, R.J. 1997a. A new genus and species of Cecidomyiidae (Diptera) from Dominica. Memoirs of the 


Gagné, R.J. 1997b. Pp. 111-113. In Gagné, R.J., C. Ott and S.S. Renner. A new species of gall midge (Diptera: 

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Gagné, R.J. 1997c. Pp. 15-18. In Gagné, R.J. and J. Marohasy. A new species of Schizomyia Kieffer (Diptera: 

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Gagné, R.J. 1997d. Pp. 313-333. In Gagné, R.J., J.K. Balciunas and D.W. Burrows. Six new species of gall midges 

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Gagné, R.J. 1998. Pp. 522-524. In Gagné, R.J., F.F.F.Ferraz and R.E.Monteiro. A new species of Neolasioptera Felt 

(Diptera: Cecidomyiidae) on Cuphea carthagenensis (Jacq.) Macbride (Lythraceae) in Brazil, with notes on its 

biology. Proceedings of the Entomological Society of Washington 100: 521-525. 

Gagné, R.J. 1999a. Pp. 23-34. In Gagné, R.J. and L.L. Law. Actilasioptera (Diptera: Cecidomyiidae), a new genus for 

Australasian and Asian gall midges of grey mangroves, Avicennia spp. (Avicenniaceae). In Csóka, G., W.J. 

Mattson, G.N. Stone and P.W. Price, eds. The Biology of Gall-Inducing Arthropods. U.S. Department of 

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Gagné, R.J. 1999b. Pp. 333-336. In Gagné, R.J. and J.T. Lill. A new Nearctic species of Lestodiplosis (Diptera: 

Cecidomyiidae) preying on an oak leaf tier, Psilocorsis quercicella (Lepidoptera: Oecophoridae). Proceedings 

of the Entomological Society of Washington 101: 332-336. 

Gagné, R.J. 1999c. A type species designation for Actilasioptera Gagné (Diptera: Cecidomyiidae). Proceedings of the 


Gagné, R.J. 1999d. Pp. 160-165. In Gagné, R.J. and E.G. Riley. A new gall midge (Diptera: Cecidomyiidae) pest of 

live oak in Texas. Southwestern Entomologist 24: 159-165. 

Gagné, R.J. 2000. Pp. 833-837. In Gagné, R.J., H. Blanco-Metzler and J. Etienne. A new Neotropical species of 

Clinodiplosis (Diptera: Cecidomyiidae), an important new pest of cultivated peppers. Proceedings of the 


Gagné, R.J. 2001. Pp. 113-133 in Gagné, R.J., R.A.M. Oda and R.E. Monteiro. The gall midges (Diptera: 

Cecidomyiidae) of Mikania glomerata (Asteraceae) in Southeastern Brazil. Proceedings of the Entomological 


Gagné, R.J. 2002a. Primavera (new genus) porrecta (Felt), the elm samara midge: description and biological notes 

(Diptera: Cecidomyiidae). Annales de la Société Entomologique de France (n.s.) 38: 39-44. 

Gagné, R.J. 2002b. Pp. 39-42. In Gentili, M.O., R.J. Gagné and P. Cueto. A new species of Xyloperrisia (Diptera: 

Cecidomyiidae), a stem gallmaker on Maytenus boaria (Celastraceae) from Argentina. Revista de la Sociedad 

Entomológica Argentina 61: 37-42. 

*Gagné, R.J. 2004a. Pp. 20-24. In Gagné, R.J. and C. de R. Medina. A new species of Procontarinia (Diptera: 

Cecidomyiidae), an important new pest of mango in the Philippines. Proceedings of the Entomological Society 


*Gagné, R.J. 2004b. A catalog of the Cecidomyiidae (Diptera) of the world. Memoirs of the Entomological Society of 

Washington No. 23, 408 pp. 

*Gagné, R.J. 2004c. Pp. 306-309. In Gagné, R.J., A. Sosa and H. Cordo. A new Neotropical species of Clinodiplosis 

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*Gagné, R.J. 2008a. Pp. 43-47. In Gagné, R.J., J.E. Peña and F.E. Acevedo. A new lestodiplosine (Diptera: 

Cecidomyiidae) preying on the avocado lace bug, Pseudacysta perseae (Heteroptera: Tingidae), in southern 

Florida. Florida Entomologist 91: 43-48. 

*Gagné, R.J. 2008b. Pp. 285-290. In Gagné, R.J. and R. Menjivar. A new species of Schizomyia (Diptera: 

Cecidomyiidae), a pest of Fernaldia pandurata (Apocynaceae) in Central America. Proceedings of the 


*Gagné, R.J. 2008c. The gall midges (Diptera: Cecidomyiidae) of hickories (Juglandaceae: Carya). Memoirs of the 

American Entomological Society 48: 1-147. 

*Gagné, R.J. 2008d. Pp. 431-435. In Gagné, R.J. and K.L. Hibbard. A new species of Cecidomyia (Diptera: 

Cecidomyiidae) feeding on resin of baldcypress. Florida Entomologist 91: 431-435. 

*Gagné, R.J. 2009a. The genus Arnoldiola (Diptera: Cecidomyiidae) in the Nearctic Region, with new synonymies and 

combinations. Proceedings of the Entomological Society of Washington 111: 106-110. 

*Gagné, R.J. 2009b. Taxonomy of Janetiella thymi (Kieffer) (Diptera: Cecidomyiidae) and of the species formerly in 

Janetiella that feed on Vitis (Vitaceae). Proceedings of the Entomological Society of Washington 111: 399- 

409. 

Gagné, R.J. and F.D. Bennett. 1993. Two new species of Lestodiplosini (Diptera: Cecidomyiidae) preying on 

Homoptera and Thysanoptera in southern Florida. The Florida Entomologist 76: 341-348. 

Gagné, R.J. and G.W. Byers. 1985. A remarkable new Neotropical species of Contarinia (Diptera: Cecidomyiidae). 

Journal of the Kansas Entomological Society 57: 736-738. 

Gagné, R.J. and J.F. Doane. 1999. The larval instars of the wheat midge, Sitodiplosis mosellana (Géhin) (Diptera: 


Gagné, R.J. and R.W. Duncan. 1990. A new species of Cecidomyiidae (Diptera) damaging shoot tips of yellow 

cypress, Chamaecyparis nootkatensis, and a new genus for two gall midges on Cupressaceae. Proceedings of 

the Entomological Society of Washington 92: 146-151. 

Gagné, R.J. and J. Etienne. 1996. Meunieriella avicenniae (Cook) (Diptera: Cecidomyiidae) the leaf gall maker of 

black mangrove in the American tropics. Proceedings of the Entomological Society of Washington 98: 

527-532. 

*Gagné, R.J. and J. Etienne. 2006. Gephyraulus mangiferae (Felt), n. comb. (Diptera: Cecidomyiidae): a mango pest 

from India newly recorded from the Western Hemisphere. Proceedings of the Entomological Society of 


*Gagné, R.J. and J. Etienne. 2009. Note on the Cecidomyiidae from Guadeloupe (West Indies) with description of a 

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82: 73-79. 

Gagné, R.J., J.H. Hatchett, S. Lhaloui, and M. El Bouhssini. 1991. Hessian fly and barley stem gall midge, two 

different species of Mayetiola (Diptera: Cecidomyiidae) in Morocco. Annals of the Entomological Society of 

America 84: 436-443. 

Gagné, R.J. and B.A. Hawkins. 1983. Biosystematics of the Lasiopterini (Diptera: Cecidomyiidae: Cecidomyiinae) 

associated with Atriplex spp. (Chenopodiaceae) in southern California. Annals of the Entomological Society 

of America 76: 379-383. 

Gagné, R.J. and K.L. Hibbard. A new species of gall midge (Diptera: Cecidomyiidae) from subterranean stem galls of 

Licania michauxii (Chrysobalanaceae) in Florida. The Florida Entomologist 79: 428-434. 

*Gagné, R.J. and K.L. Hibbard. A new species of Cecidomyia (Diptera: Cecidomyiidae) feeding on resin of 

baldcypress. Florida Entomologist 91: 431-435. 

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pest of mango in the Philippines. Proceedings of the Entomological Society of Washington 106: 19-25. 

*Gagné, R.J. and R. Menjivar. 2008. A new species of Schizomyia (Diptera: Cecidomyiidae), a pest of Fernaldia 

pandurata (Apocynaceae) in Central America. Proceedings of the Entomological Society of Washington 110: 

384-291. 

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western Palearctic Region. Proceedings of the Entomological Society of Washington 99: 756. 

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avocado lace bug, Pseudacysta perseae (Heteroptera: Tingidae), in southern Florida. Florida Entomologist 91: 

43-48. 

*Gagné, R.J., F. Posada and Z.M. Gil. 2004. A new species of Bruggmanniella (Diptera: Cecidomyiidae) aborting 

young fruit of avocado, Persea americana (Lauraceae), in Colombia and Costa Rica. Proceedings of the 


Gagné, R.J., R. Sobhian and N. Isidoro. 1996. A review of the genus Psectrosema (Diptera: Cecidomyiidae), Old 

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Lophodiplosis trifida, an Australian gall midge (Diptera: Cecidomyiidae) and biocontrol agent of paperbark in 

Florida. Florida Entomologist 92: 593-597. 

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*Ganaha, T., M. Nohara, S. Sato, N. Uechi, K. Yamagishi, S. Yamauchi and J. Yukawa. 2007. Polymorphism of 

axillary bud galls induced by Rhopalomyia longitubifex (Diptera: Cecidomyiidae) on Artemisia princeps and 

A. montana (Asteraceae) in Japan and Korea, with designation of new synonyms. Entomological Science 10: 

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Maia, V.C. 1994a. Considerações sobre Proasphondylia Felt (Diptera, Cecidomyiidae), com descrições de duas 

espécies novas associadas com Guapira opposita (Velloso) Reitz (Nyctaginaceae). Revista Brasileira de 

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Maia, V.C. 1994b. Considerações sobre Stephomyia Tavares (Diptera, Cecidomyiidae, Asphondyliidi), com descrição 

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Revista Brasileira de Zoologia 10: 521-530. 

Maia, V.C. 1994c. Uma nova espécie de Clinodiplosis Kieffer (Diptera, Cecidomyiidae), associada com Melissa 

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Maia, V.C. 1995. Myrciariamyia bivalva, gen. n. e sp. n. (Diptera, Cecidomyiidae, Oligotrophini), associado com 

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Maia, V.C. 1996a. Dois gêneros novos de Cecidomyiidae (Diptera) associados à Myrtaceae, na restinga de Barra de 

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Maia, V.C. 1996b. Três espécias novas de Dasineura Rondani (Diptera, Cecidomyiidae) associadas a Myrtaceae, na 

restinga de Barra de Maricá, Rio de Janeiro. Revista Brasileira de Zoologia 12: 1001-1008. 

Maia, V.C. 1996c. Cordiamyia globosa gen. n. e sp. n. (Diptera, Cecidomyiidae) associado com Cordia verbenacea 

DC. (Boraginaceae) no Brasil. Revista Brasileira de Zoologia 13: 579-583. 

Maia, V.C. 1997. Clusiamyia nitida gen. n. e sp. n. (Diptera, Cecidomyiidae, Cecidomyiidi) associada com Clusia 

lanceolata Camb. (Clusiaceae) no Brasil. Revista Brasileira de Zoologia 13: 829-832. 

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Asphondylia maytenuse Maia & Couri (Diptera, Cecidomyiidae). Revista Brasileira de Zoologia 16: 775-778. 

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Maia, V.C. 2001b. Two new species of gall midges (Diptera, Cecidomyiidae) associated with Pouteria caimito var. 

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Maia, V.C. 2002a. Description of the larva of Houardodiplosis rochae Tavares, 1925 (Diptera, Cecidomyiidae, 

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*Maia, V.C. 2004a. A new genus and six new species of gall midges (Diptera, Cecidomyiidae) from serra de São José 

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*Maia, V.C. 2004c. Description of a new species of Bruggmannia Tavares (Diptera, Cecidomyiidae) associated with 

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*Maia, V.C. 2005b. Clinodiplosis costai, a new galler species (Diptera, Cecidomyiidae) associated with Paullinia 

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*Maia, V.C. 2008. Pp. 54-57. In Maia, V.C., F.A.O.Silveira, L.A. Oliveira, and M.F. Xavier. Asphondylia gochnatiae, 

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*Maia, V.C. and G. W. Fernandes. 2005a. A new genus and species of gall midges Diptera: Cecidomyiidae) associated 

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*Maia, V.C. and G. W. Fernandes. 2005b. Two new species of Asphondyliini (Diptera: Cecidomyiidae) associated 

with Bauhinea brevipes (Fabaceae). Zootaxa 1091: 27-40. 

*Maia, V.C. and G. W. Fernandes. 2006. A new genus and species of gall midge (Diptera, Cecidomyiidae) associated 

with Parkia pendula (Fabaceae, Mimosoideae). Revista Brasileira de Entomologia 50: 1-5. 

*Maia, V.C. and G. W. Fernandes. 2007. Myrciariamyia admirabilis, a new species of gall midge (Diptera, 

Cecidomyiidae) associated with Erythroxylum suberosum (Erythroxylaceae). Zootaxa 1554: 41-48. 

*Maia, V.C., G. W. Fernandes and D. Negreiros. 2009. A new genus and species of gall midge (Diptera, 

Cecidomyiidae) associated with Myrcia retorta (Myrtaceae) Revista Brasileira de Entomologia 53: 38-40. 

Maia, V.C., M. de S. Mendonça, Jr. and H.P. Romanowski. 1997. Eugeniamyia dispar gen. n. e sp. n. (Diptera, 

Cecidomyiidae, Lasiopteridi) associated with Eugenia uniflora L. (Myrtaceae) in Brazil. Revista Brasileira de 

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*Maia, V.C. and U.P. de Oliveira. 2007. Uma nova espécie de Cecidomyiidae (Diptera) associada com Sebastiania 

glandulosa (Euphorbiaceae). Iheringia, Série Zoologica 97: 97-101. 

*Maia, V.C., J.C. Santos and G.W. Fernandes. 2009. Asphondylia fructicola, a new species of Cecidomyiidae (Diptera) 

associated with Solanum sp. (Solanaceae) from Brazil. Revista Brasileira de Entomologia 53: 166-170. 

*Maia, V.C., F.A.O.Silveira, L.A. Oliveira and M.F. Xavier. 2008. Asphondylia gochnatiae, a new species of gall 

midge (Diptera, Cecidomyiidae) associated with Gochnatia polymorpha (Asteraceae). Zootaxa 1740: 53-58. 

*Maia, V.C. and J. Vasquez. 2006. A new species of gall midge (Diptera: Cecidomyiidae) associated with Theobroma 

bicolor (Sterculiaceae) from Peru. Arquivos do Museu Nacional, Rio de Janeiro 64: 125-129. 

*Maia, V.C., M. Zart and M. Botton. 2009. Neolasioptera ramicola, a new species of Cecidomyiidae (Diptera) 

associated with Physalis angulata (Solanaceae). Revista Brasileira de Entomologia 53: 163-165. 

Malloch, J,R. 1914. Notes on North American Diptera, with descriptions of new species in the collection of the Illinois 

State Laboratory of Natural History. Bulletin of the Illinois State Laboratory of Natural History 10: 213-243, 

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Mamaev, B.M. 1960a. [Description of two new genera and one species of gall midge (Itonididae, Diptera) developing 

in rotten wood.] Zoologicheskiï Zhurnal 39: 1521-1524. 

Mamaev, B.M. 1960b. [Micropteromyia ghilarovi, gen. et. sp. nov. (Diptera, Itonididae) with reduced wings.] 

Entomologicheskoe Obozrenie 39: 951-955. Translated in Entomological Review (1961) 39 (4): 690-693. 

Mamaev, B.M. 1961a. [Description of the gall midge Aprionus smirnovi sp. n. (Itonididae, Diptera).] Zoologicheskiï 

Zhurnal 40: 614-615. 

Mamaev, B.M. 1961b. [New species of the genera Trisopsis Kieff. and Triommatomyia, gen. n. (Diptera, Itonididae).] 


Mamaev, B.M. 1961c. Neue Hiastatus-Arten (Diptera: Itonididae). Beiträge zur Entomologie 11: 446-450. 

Mamaev, B.M. 1961d. [New representatives of gall midge genera, known in the Nearctic Region, among the fauna of 

the European part of the USSR (Itonididae, Diptera).] Doklady Akademii Nauk 139: 227-229. 

Mamaev, B.M. 1961e. [New gall midges of the genus Ametrodiplosis Rübs. (Löwodiplosis Kieff.) (Diptera, 

Itonididae).] asopis eskoslovenské Spole nosti Entomologické 58: 384-388.

508 

Mamaev, B.M. 1961f. [Gall midges of the USSR. New species of the genus Camptomyia Kieffer (Itonididae, Diptera).] 

Zoologicheskiï Zhurnal 40: 1677-1690. 

Mamaev, B.M. 1963a. [Gall midges of the USSR. 2. The tribe Micromyini (Diptera, Itonididae).] Entomologicheskoe 

Obozrenie 42: 436-454. Translated in Entomological Review (1964) 42 (2): 239-247. 

Mamaev, B.M. 1963b. [Gall midges of the USSR. Part 3. New species of the genus Winnertzia Rondani, developing in 

the soil, in fungal mycelium and under decaying bark of coniferous trees (Itonididae, Diptera).] Zoologicheskiï 

Zhurnal 42: 562-573. 

Mamaev, B.M. 1963c. [Gall midges of the USSR. 5. New Middle Asian species of the tribes Lestremiini, Micromyini, 

Porricondylini (Itonididae, Diptera).] Uzbekskiï Biologicheskii Zhurnal 1963 (2): 70-77. 

Mamaev, B.M. 1964a. [Gall-midges of the USSR. 4. New species of the tribe Lestremiini Itonididae, Diptera).] 


Mamaev, B.M. 1964b. [On the systematic position of the Tribe Leptosynini Enderlein (Diptera, Itonididae).] asopis 

eskoslovenské Spole nosti Entomologické 61: 89-99. 

Mamaev, B.M. 1964c. [Gall midges of the USSR. 6. New species of the tribe Porricondylini (Diptera, Cecidomyiidae).] 


Mamaev, B.M. 1964d. Acoenonia europaea sp. n.: the first Palaearctic species of the tribe Acoenoniini (Diptera: 

Itonididae). Proceedings of the Royal Entomological Society of London 33: 131-132. 

Mamaev, B.M. 1965a. Pp. 148, 266-267. In Mamaev, B.M. and N.P. Krivosheina. [The Larvae of Gall Midges 

(Diptera, Cecidomyiidae).] Akademiya Nauk USSR. Moscow. Translated, 1993, Roskam, J.C. and A.A. 

Balkema, eds. Rotterdam. ix & 293 pp. 

Mamaev, B.M. 1965b. [The practical genus and species conception in gall midges (Diptera, Cecidomyiidae).] 


Mamaev, B.M. 1965c. [Replacement of secondary sexual characters and origin of new taxonomic groups of insects 

exemplified by gall midges (Cecidomyiidae, Diptera).] Zhurnal Obshchei Biologii 26: 677-684. 

Mamaev, B.M. 1966. [New and little known Palaearctic gall midges of the tribe Porricondylini (Diptera, 

Cecidomyiidae).] Acta Entomologica Bohemoslovaca 63: 213-239. 

Mamaev, B.M. 1967. [Gall-midges of the USSR. 7. New species of non-gall-making gall midges of the tribe 

Oligotrophini (Diptera, Cecidomyiidae).] Entomologicheskoe Obozrenie 46: 873-883. Translated in 


Mamaev, B.M. 1968a. [New Nematocera of the USSR (Diptera, Axymyiidae, Mycetobiidae, Sciaridae, 

Cecidomyiidae).] Entomologicheskoe Obozrenie 47: 605-616. Translated in Entomological Review (1969) 47 

(3): 371-377. 

Mamaev, B.M. 1968b. [Evolution of Gall Forming Insects: Gall Midges.] Akademia Naul, Leningrad. 235 pp. 

Translated 1975, K.M. Harris, ed. British Library, Boston Spa, United Kingdom. iii & 316 pp. 

Mamaev, B.M. 1969a. 26. [Family Cecidomyiidae (Itonididae).] Pp. 356-420. In Bei-Bienko, G.Ya., ed. Keys to the 

Insects of the European Part of the USSR. Vol. V. Diptera and Siphonaptera. Part I. Leningrad. Published in 

English, 1988, Amerind Publishing Co., Ltd., New Delhi. 

Mamaev, B.M. 1969b. Not seen. [Discovery of a gall midge of the Japanese genus Etsuhoa Inouye on Juniperus 

seravschanica in Uzbekistan.] Just. Sci. Techn. Inform., Moscow, No. 1619: 4 pp. 

Mamaev, B.M. 1971. [The use of resin as a habitat by insects, with a review of insects living in resin.] Zhurnal 

Obshcheï Biologii 32: 501-508. 

Mamaev, B.M. 1972a. [New gall midges (Diptera, Cecidomyiidae) of the desert zone.] Entomologicheskoe Obozrenie 

51: 886-899. Translated in Entomological Review (1974) 51 (4): 526-534. 

Mamaev, B.M. 1972b. [Review of species and ecological relations of insect decomposers of wood of Ulmus propinqua 

Loidz.] Rol' Nasekomykh V Leshykh Biogeotsenozakh Primor'ya. Trudy Biologo-pochvennogo Instituta, 

Novaya Seriya 7 (110): 106-120. 

Mamaev, B.M. 1973a. [Revision of the Palearctic species of the genus Coquillettomyia Felt (Diptera, Cecidomyiidae).] 


Mamaev, B.M. 1973b. [A new genus of endoparasitic gall midges, Occuloxenium compitale Mamaev gen. et sp. n., 

developing on aphids (Diptera, Cecidomyiidae).] Uzbekskiï Biologicheskiï Zhurnal 17 (6): 52-54. 

Mamaev, B.M. 1975a. [Xylophagous insects developing on Quercus mongolica in southern Primor'e.] [Entomological 

Investigations in the Far East.] Trudy Biologo-pochvennogo Instituta, Novaya Seriya 28: 35-42.

509 

Mamaev, B.M. 1975b. [Endemic and subendemic elements in xylophagous entomocomplexes developing on Padus 

maacki.] [Entomological Investigations in the Far East.] Trudy Biologo-pochvennogo Instituta, Novaya Seriya 

28: 50-57. 

Mamaev, B.M. 1975c. [Comparative nature of entomocomplexes developing in wood of Padus asiatica in southern 

Maritime Territory.] [Entomological Investigations in the Far East.] Trudy Biologo-pochvennogo Instituta, 

Novaya Seriya 28: 58-62. 

Mamaev, B.M. 1975d. [Some regularities of infestation of Betula costata and Betula manshurica by xylophagous 

insects.] [Entomological Investigations in the Far East.] Trudy Biologo-pochvennogo Instituta, Novaya Seriya 

28: 81-88. 

Mamaev, B.M. 1975e. [Gall insects of saxaul forestation.] Pp. 61-82. In Mamaev, B.M. and F.N. Pravdin, eds. [Insects 

as Components of Biogenesis of the Saxaul Forest.] Moscow. 

Mamaev, B.M. 1976. Pp. 1-4. In Mamaev, B.M. and T.A. Rozhnova. The description of gall-midge Wyattella fungicola 

Mamaev sp. n. (Cecidomyiidae: Diptera) with special reference to characters under scanning electron 

microscope. Cecidologia Indica 11: 1-24. 

Mamaev, B.M. 1978. New genus Groveriella Mamaev, gen. n. of the tribe Strobliellini (Diptera, Cecidomyiidae). 

Cecidologia Indica 12-13: 1-3. 

Mamaev, B.M. 1979. [New species Neocatocha asiatica sp. n. - first record of the genus Neocatocha Felt (Diptera, 

Cecidomyiidae) in the Palaearctic Region.] Trudy Vsesoyuznoge Entomologicheskogo Trudy Obshchestva 

61: 185-186. 

Mamaev, B.M. 1980. Pp. 150-151. In Mohrig, W., B.M. Mamaev, and V. Spungis. Information on wing-reduced 

Diptera in the ground-litter. Part X: Cecidomyiidae (Genus Chastomera and Micropteromyia) and Sciaridae 

(Genus Plastosciara). Zoologischer Jahrbücher für Systematik 107: 148-153. 

Mamaev, B.M. 1985. [New gall midge and sciarid species (Diptera, Cecidomyiidae, Sciaridae) from the USSR.] 


Mamaev, B.M. 1986. [New gall midges from Turkmenistan (Diptera, Cecidomyiidae).] Izvestiya Akademii Nauk 

Turkmenskoi SSR, Seriya Biologicheskikh Nauk 1986 (5): 65-68. 

Mamaev, B.M. 1988. [Results of studies on gall midges damaging Salsola orientalis in northern Turkmenistan.] Pp. 56- 

64. In [The Insect Complex of the Northern Turkmanistan Desert.] Trudy Nauchno-Issledovatel'skiï Institut, 

Nauchno-Tekhnicheskoï Informatsii i Tekhniko-Èkonomicheskikh Isseledovaniï, Gosplan Turkmenskoï SSR, 

Ashkabad. 

Mamaev, B.M. 1990a. [Description of new genera and species and key to genera of Palaearctic gall midges of the 

subfamily Porricondylinae (Diptera, Cecidomyiidae).] Acta Zoologica Bulgarica 40: 12-28. 

Mamaev, B.M. 1990b. [Gall midges (Diptera, Cecidomyiidae) of the subarctic region.] Pp. 141-146. In Novosti 

Sistematiki i Faunistiki. Kiev: Naukova Dumka. 

Mamaev, B.M. 1993. [Gall midges of the Xylopriona group (Diptera, Cecidomyiidae) and their ecological 

significance.] Pp. 46-61. In Inozemtsev, A.A. Anthropogenic Factors on Structure and Function of 

Ecosystems and Their Separate Components. Moscow Pedagological University. 

Mamaev, B.M. 1994. [A contribution to the gall midge fauna (Diptera, Cecidomyiidae) of Kamchatka with descriptions 

of new species.] Vestnik Zoologii 1994 (2): 28-32. 

Mamaev, B.M. 1995. [Cryptoxylomyia Mamaev, gen. n. (Diptera, Cecidomyiidae).] Vestnik Zoologii 1995 (5-6): 65. 

Mamaev, B.M. 1996. Preliminary materials to the revision of gall midges belonging to genera Aprionus Kieffer and 

Peromyia Kieffer (Diptera, Cecidomyiidae). All-Russian Institute of Continuous Education in Forestry, 

Pushkino, Moscow Region 5: 1-5. 

Mamaev, B.M. 1997a. New genus of gall-midges, Dallaiella Mamaev, gen. nov. (Diptera, Cecidomyiidae). 

Entomologica (Bari) 30: 69-72. 

Mamaev, B.M. 1997b. Four new aberrant species of gall midges of the genus Aprionus Kieffer (Diptera, 

Cecidomyiidae). All-Russian Institute of Continuous Education in Forestry, Pushkino, Moscow Region 6: 1-6. 

Mamaev, B.M. 1997c. Preliminary discussion on taxonomy of the tribe Holoneurini of log midges (Diptera, 

Cecidomyiidae, Porricondylinae). All-Russian Institute of Continuous Education in Forestry, Pushkino, 

Moscow Region 8: 1-5. 

Mamaev, B.M. 1998a. [New species of gall midges of the subfamily Lestremiinae (Diptera, Cecidomyiidae).] 

All-Russian Institute of Continuous Education in Forestry, Pushkino, Moscow Region 10: 1-10. 

Mamaev, B.M. 1998b. Natural system and species composition of gall midges belonging to subfamily Lestremiinae

510 

(Diptera, Cecidomyiidae). All-Russian Institute of Continuous Education in Forestry, Pushkino, Moscow 

Region 11: 1-6. 

Mamaev, B.M. 1998c. [New species of gall midges of the genus Claspettomyia Grover (Diptera, Cecidomyiidae).] 

All-Russian Institute of Continuous Education in Forestry, Pushkino, Moscow Region 12: 1-10. 

Mamaev, B.M. 1998d. [New species of gall midges of various taxa (Diptera, Cecidomyiidae).] All-Russian Institute of 

Continuous Education in Forestry, Pushkino, Moscow Region 13: 1-10. 

Mamaev, B.M. 1998e. [Review of the publication, Revision of the "Lestremiinae" (Diptera, Cecidomyiidae) of the 

Holarctic (by M. Jaschhof).] All-Russian Institute of Continuous Education in Forestry, Pushkino, Moscow 

Region 14: 1-7. 

Mamaev, B.M. 1998f. New species of the gall midge genus Winnertzia from Far East Russia, with remarks on 

synonymy (Diptera, Cecidomyiidae). Journal of the Ukrainian Entomological Society 4: 41-46. 

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Russian Far East. VI, Diptera and Siphonaptera. Pt. 1. Dal'nauka, Vladivostok. 

Mamaev, B.M. 2001a. [Criteria and indicators of species validity of gall midges (Diptera, Cecidomyiidae).] All-Russian 

Institute of Continuous Education in Forestry, Pushkino 15: 3-18. 

Mamaev, B.M. 2001b. [New species of gall midges and sciarids (Diptera: Cecidomyiidae, Sciaridae) in mountain 

tundra of the northern Urals.] All-Russian Institute of Continuous Education in Forestry, Pushkino 16: 1-12. 

Mamaev, B.M. 2001c. [The tribe Holoneurini in the Palearctic (Diptera, Cecidomyiidae).] All-Russian Institute of 

Continuous Education in Forestry, Pushkino 17: 1-11. 

Mamaev, B.M. 2001d. [New species of gall midges of the genus Winnertzia Rond. in arid regions (Diptera, 

Cecidomyiidae).] All-Russian Institute of Continuous Education in Forestry, Pushkino 18: 1-7. 

Mamaev, B.M. 2001e. [Gall midges of the genus Winnertzia Rond. from the group with unarmed gonostyle (Diptera, 


Mamaev, B.M. 2002a. [New species of gall midges of the genus Winnertzia Rond. developing in tissue of fruit bodies 

of wood fungi (Diptera, Cecidomyiidae).] All-Russian Institute of Continuous Education in Forestry, 

Pushkino 20: 1-12. 

Mamaev, B.M. 2002b. [New species of gall midges of the genus Winnertzia Rond. and retention of the species W. 

xylostei Mam. and W. argentata Mam. (Diptera, Cecidomyiidae).] All-Russian Institute of Continuous 

Education in Forestry, Pushkino 21: 1-13. 

Mamaev, B.M. 2002c. [The main group of new species of gall midges of the genus Winnertzia Rond. (Diptera, 


Mamaev, B.M. and O.Kh. Becknazharova. 1983. [New species of gall midges on buds of tamarisk.] Izvestiya Akademii 

Nauk Turkmenskoi SSR 1983 (4): 68-69. 

Mamaev, B.M. and Z.L. Berest. 1990a. [New species of lestremiine gall midges (Diptera, Cecidomyiidae) from 

Carpathian and Transcarpathian regions.] Vestnik Zoologii 1990 (4): 18-24. 

Mamaev, B.M. and Z.L. Berest. 1990b. [New species of Lestremiine gall midges (Diptera, Cecidomyiidae) from 

Carpathian and Transcarpathian regions. Communication 2. Genus Peromyia.] Vestnik Zoologii 1990 (5): 

18-25. 

Mamaev, B.M. and Z.L. Berest. 1991a. [New species of lestremiine gall midges of the genera Heterogenella and 

Skuhraviana (Diptera, Cecidomyiidae) from Carpathian and Transcarpathian regions. Vestnik Zoologii 1991 

(2): 27-33. 

Mamaev, B.M. and Z.L. Berest. 1991b. [A new genus of lestremiine gall midges (Diptera, Cecidomyiidae) from the 

Transcarpathian Region.] Vestnik Zoologii 1991 (5): 75-77. 

Mamaev, B.M. and Z.L. Berest. 1992. [A new subgenus and new Aprionus and Acoenonia species (Diptera: 

Cecidomyiidae) from the Carpathians and Transcarpathia.] Vestnik Zoologii 1992 (1): 67-70. 

Mamaev, B.M. and Z.L. Berest. 1994. A new species of the genus Peromyia (Diptera: Cecidomyiidae) from the 

Transcarpathian region. Vestnik Zoologii 1994 (1): 77-78. 

Mamaev, B.M. and Z.L. Berest. 1996. [Aprionus complicatus Mamaev et Berest, nom. n. pro A. rostratus Mamaev et 

Berest, 1990 (Vestn. Zool. 4: 20) non Yukava, 1971 (Mem. Fac. Agric. Kagoshima Univ. 8, 1: 37).] Vestnik 

Zoologii 1995 (4): 11. 

*Mamaev, B.M. and Z.L. Berest. 2005. A new species of the genus Heterogenella (Diptera, Cecidomyiidae, 

Lestremiinae) Zacarpathia (Ukraine). Vestnik Zoologii 39: 83-85. 

Mamaev, B.M. and B.D. Dimitrova. 1992. A new species of gall-midges from Bulgaria (Insecta, Diptera,

511 

Cecidomyiidae: Porricondylinae). Reichenbachia Staatliches Museum für Tierkunde Dresden 29: 173-175. 

Mamaev, B.M. and B.D. Dimitrova. 1998. A new species of gall midges of the genus Pterepidosis Mamaev from 

Bulgaria (Insecta: Diptera: Cecidomyiidae: Porricondylinae). Reichenbachia 32: 317-319. 

Mamaev, B.M. and Efremova. 1994. P. 32. In Mamaev, B.M. 1994. [A contribution to the gall midge fauna (Diptera, 

Cecidomyiidae) of Kamchatka with description of new species.] Vestnik Zoologii 1994 (2): 28-32. 

Mamaev, B.M. and M. Jaschhof. 1997. Neue paläarktische Arten aus der Gattung Aprionus Kieffer, 1894 (Diptera: 

Cecidomyiidae, Lestremiinae). Beiträge zur Entomologie 47: 451-463. 

Mamaev, B.M. and Krivosheina, N.P. 1965. [The Larvae of Gall Midges (Diptera, Cecidomyiidae).] Akademiya Nauk 

USSR. Moscow. 277 pp. Translated, 1993, Roskam, J.C. and A.A. Balkema, eds. Rotterdam. ix & 293 pp. 

Mamaev, B.M. and M.G. Krivosheina. 1998. [On] the taxonomy of the gall midges of the genus Karschomyia Felt, 

1908 (Diptera: Cecidomyiidae) with description of 11 new species. Russian Entomological Journal 6: 75-81. 

Mamaev, B.M. and I.G. Kritskaya. 1980. Pp. 146-147. In Mamaeva, Kh.P. and I.G. Kritskaya. [New and little-known 

species of the tribe Lasiopterini (Diptera, Cecidomyiidae) from the USSR.] Entomologicheskoe Obozrenie 59: 

410-414. Translated in Entomological Review (1982) 59 (2): 146-150. 

Mamaev, B.M. and I.G. Kritskaya. 1981. Pp. 404-405. In Kritskaya, I.G. and Kh. P. Mamaeva. New systematic data on 

gallmidges of the tribe Asphondyliini (Diptera, Cecidomyiidae) of the USSR. Entomologicheskoe Obozrenie 

60: 401-406. Translated in Entomological Review (1983) 60 (2): 129-134. 

Mamaev, B.M. and L.S. Mirumian. 1989. [A new gall midge, Sophoromyia armenica Mamaev et Mirumian sp. n. 

(Diptera, Cecidomyiidae), damaging licorice (Glycyrrhiza glabra L.).] Doklady Akademii Nauk Armyanskoï 

SSR 88: 43-45. 

Mamaev, B.M. and L.S. Mirumian. 1990a. [A new genus of gall midges, Bremiolina gen. n., and new species, 

Bremiolina gemmicola sp. n. (Diptera, Cecidomyiidae), on the willow Salix triandra L. in Armenia.] Doklady 

Akademii Nauk Armyanskoï SSR 90: 91-94. 

Mamaev, B.M. and L.S. Mirumian. 1990b. [Description of the gall midge Arafavilla terteriani gen. et. sp. n. (Diptera, 

Cecidomyiidae), inducing galls on Suaeda altissima (L.) Pall. in Armenia.] Doklady Akademii Nauk 

Armyanskoï SSR 90: 139-141. 

Mamaev, B.M. and W. Mohrig. 1975. Zur Kenntnis flügelreduzierter Dipteren der Bodenstreu. VI. Beitrag: Gattungen 

Microcordylomyia, Aprionus und Trisopsis (Cecidomyiidae). Zoologischer Anzeiger Jena 194: 125-132. 

Mamaev, B.M. and V.I. Nikolskij. 1983. [New species - gall midge Dasyneura rozhkovi Mamaev et Nikolskij, sp. n.- 

an important pest of the larch in Siberia (Diptera, Cecidomyiidae).] Izvestiya Sibirskogo Otdeleniya Akademii 

Nauk SSSR, Seriya Biologocheskikh Nauk 1983 (3): 108-112. 

Mamaev, B.M. and B. Økland. 1996. Description of two new species and key to the genus Anaretella Enderlein 


Region 4: 1-12. 

Mamaev, B.M. and B. Økland. 1998. The genus Bryomyia Kieff. (Diptera, Cecidomyiidae): Palaearctic species and 

Fennoscandian records. Entomologica Fennica 26: 147-152. 

Mamaev, B.M. and S.M. Pak. 1989. [New species of phytophagous gall midges from northwestern Turkmenistan 

(Diptera, Cecidomyiidae).] Izvestiya Akademii Nauk Turkmenskoi SSR, Seriya Biologicheskikh Nauk 1989 

(3): 31-35. 

Mamaev, B.M. and S.M. Pak. 1990. [A new species of gall midge, Halodiplosis commixta sp. n., developing in 

transformed galls of Halodiplosis salsolicola Mar. on Salsola richteri (Diptera, Cecidomyiidae).] Izvestiya 

Akademii Nauk Turkmenskoi SSR, Seriya Biologicheskikh Nauk 1990 (6): 57-59. 

Mamaev, B.M. and A.A. Przhiboro. 2001. A new species of the gall midge genus Coniophora from Pskov Province of 

Russia (Diptera: Cecidomyiidae). Zoosystematica Rossica 9: 235-236. 

Mamaev, B.M. and T.A. Rozhnova. 1982. [A revision of species of the genus Cordylomyia (Diptera, Cecidomyiidae) 

from the fauna of the USSR.] Vestnik Zoologii 1982 (6): 18-24. 

Mamaev, B.M. and T.A. Rozhnova. 1983. [Gall midges of the genus Cordylomyia Felt (Diptera, Cecidomyiidae) from 

the fauna of the Far East.] Pp. 64-68. In Sistematika i Ekolo-faunisticeskii Obzor Otdel'nkh Otryadov 

Nasekomykh Dal'nego Vostoka. Dal'nevostochnyi Nauchnyi Tsentr, Akademiya Nauk SSR, Sbornik 

Nauchnyi Trudov. Vladivostok. 

Mamaev, B.M. and O.S. Soyunov. 1989. [Description of a new genus of non-gallmaking gall midges, Desertepidosis 

Mam. et Sojun. gen. n., from Turkmenistan (Diptera, Cecidomyiidae).] Izvestiya Akademii Nauk 

Turkmenskoi SSR, Seriya Biologichesikh Nauk 1989 (4): 65-67.

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Mamaev, B.M. and W. Spungis. 1980. Pp. 148-150. In Mohrig, W., B.M. Mamaev, and W. Spungis. Information on 

wing-reduced Diptera in the ground-litter. Part X: Cecidomyiidae (Genus Chastomera and Micropteromyia) 

and Sciaridae (Genus Plastosciara). Zoologische Jahrbucher für Systematik 107: 148-153. 

Mamaev, B.M. and A.I. Zaitzev. 1996a. Description of new species and key to gall midges of the genus Mycopriona 

Mamaev (Diptera, Cecidomyiidae). All-Russian Institute of Continuous Education in Forestry, Pushkino, 

Moscow Region [1]: 1-16. 

Mamaev, B.M. and A.I. Zaitzev. 1996b. New genus of gall midges Pallidepidosis gen. n. of the subfamily 

Porricondylinae (Diptera, Cecidomyiidae). International Journal of Dipterological Research 7: 75-77. 

Mamaev, B.M. and A.I. Zaitzev. 1996c. New genus of gall midges Yukawamyia Mamaev et Zaitzev, gen. n. (Diptera, 


Mamaev, B.M. and A.I. Zaitzev. 1996d. Revision of Palaearctic species of the genus Porricondyla Rondani, 1840 s. str. 

(Diptera, Cecidomyiidae). International Journal of Dipterological Research 7: 203-215. 

Mamaev, B.M. and A.I. Zaitzev. 1997a. Four new aberrant species of gall midges of the genus Peromyia Kieffer 


Region 7: 1-9. 

Mamaev, B.M. and A.I. Zaitzev. 1997b. A new species of Jamalepidosis Mamaev (Diptera, Cecidomyiidae). Studia 


Mamaev, B.M. and A.I. Zaitzev. 1997c. New genus and species of free-developing gall midges of the subfamily 

Porricondylinae from Somalia (Diptera, Cecidomyiidae). Journal of the Ukrainian Entomological Society 3 

(2): 5-13. 

Mamaev, B.M. and A.I. Zaitzev. 1998a. A review of gall midges of the genus Allarete Pritchard (Diptera, 


Mamaev, B.M. and A.I. Zaitzev. 1998b. Records of freedeveloping gall midges in Italy, with the description of four 

new species (Diptera, Cecidomyiidae). Entomologica 31 (1997): 125-132. 

Mamaev, B.M. and A.I. Zaitzev. 1998c. Ten new species of gall midges of the genus Camptomyia Kieffer (Diptera, 

Cecidomyiidae) from East Palaearctic Region. Far Eastern Entomologist No. 62: 1-12. 

Mamaev, B.M. and A.I. Zaitzev. 1998d. Sylvenomyia gen. n. in Sweden and a key to the genera of the tribe 

Winnertziini (Diptera: Cecidomyiidae, Porricondylinae). Entomologica Fennica 9: 211-213. 

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LIST OF ERRATA IN GAGNE 2004b 

Minor corrections in Gagné (2004b), e.g., unitalicized scientific names, inconsistent punctuation, or missing 

diacritical marks were simply corrected in the new edition and are not listed here. Errata that would tend to mislead, 

such as misspellings and missing earlier papers are listed below. Many of these errors were kindly brought to my 

attention by others, in which cases their names appear following the entry: M. Jaschhof, K.M. Harris, F.C. Thompson, 

N. Dorchin, and O. Rohfritsch. Some others were brought to my attention by the research papers cited.

543 

p. 9. In first paragraph, change Pinus inopis to Pinus inops. 

p. 10. Under Catotricha mesozoica, change sex of type from ♀ to ♂. [Jaschhof] 

p. 12. Under Anaretella magnicornis in the list of synonyms of Anaretella defecta, add "bicincta Mamaev 1969a: 371, 

error for cincta." 

p. 15. Under Aprionus pyxidiifer, the type locality is Knutby, not Knidti. [Jaschhof] 

p. 14. Under Aprionus demonstrativa, the type locality is Kadnikovskaya, not Kalnikovskaya. [Jaschhof] 

p. 16. Bryomyia is Oriental as well as Holarctic. [Jaschhof] 

p. 18. Under Campylomyza flavipes, the type of Prionellus strobli is ♀, not ♂, and it is lost. [Jaschhof] 

p. 22. Lestremia intercalaris: change spelling to intercalaria. [Jaschhof] 

p. 23. Under Micromyia lucorum, change spelling from Crespinella to Crespiniella. [Thompson] 

p. 32. Peromyia spinosa: the original page number is 75, not 68. [Jaschhof] 

p. 32. Peromyia subanatina: the type locality is Bennebol, not Binnebol. [Jaschhof] 

p. 33. Polyardis monotheca was originally described in Monardia, not Polyardis. [Jaschhof] 

p. 35. Skuhraviana has a Holarctic distribution. [Jaschhof] 

p. 44. Coccopsis is preoccupied and has been renamed Coccopsilis by Harris (2004b). 

p. 53. Under Porricondyla, add the previously unseen Kieffer (1894k) names, Dicraneura Kieffer and Dicraneura 

albimana Kieffer. 

p. 61. Per Kieffer (ibid.), add the earlier reference to Winnertzia (as Clinorhiza) nigripennis Kieffer. 

p. 66. The genus Alliomyia is out of alphabetical order; it should follow Alexomyia. 

p. 71. The original combination for Arthrocnodax incanus was Diplosis, not Arthrocnodax. 

p. 98. Calmonia was preoccupied and since renamed Novocalmonia by Ozdikmen (2009). 

p. 98. Following "(orig. des.)" under Calmonia delete "= fici Gagné." 

p. 98. Following the entry for Calmonia urostigmatis, add ♂ to type series. 

pp. 101-102. The type depository for Celticecis celtiphyllia is USNM, not NYSM. 

p. 103. The host of Contarinia silene should be Lychnis dioica [Tavares 1919b, pointed out by Skuhravá et al. 2006: 

103]. 

p. 110. The name brevipalpis should end as "us." [Harris] 

p. 116. The type depository for Contarinia nucicola is USNM, and the type is not lost. 

p. 117. Contarinia pisi was described in 1850, not 1950. [Harris] 

p. 118. The name ramachandri is properly spelled ramachandrani, and its host is Acacia ferruginea, not A. suma. 

[Rohfritsch] 

p. 118. Contarinia (as Stictodiplosis) rubsaameni was found in Kieffer 1894k, a reference I had not previously seen. 

p. 119. Contarinia scutati is not authoritatively recorded in the UK. [Harris] 

p. 119. An earlier reference, Kieffer (1894k), was found for Contarinia (as Eudiplosis) sorbi Kieffer. 

p. 134. An earlier reference, Kieffer (1894k), was found for Dasineura (as Dichelomyia) mali Kieffer. 

p. 135. Dasineura marginemtorquens should be moved to Rabdophaga, as suggested by Harris, per use in Stelter 

1989c. 

p. 136. Dasineura oleae (Angelini) predates oleae Loew, which is a junior synonym and homonym, as brought to light 

in Skuhravá, Skuhravy, and Massa 2007. 

p. 136. Under Dasineura oxycoccana, the type information for Cecidomyia vaccinii was not given. 

p. 147. Didactylomyia spinii Grover should have been placed in Claspettomyia as per Grover 1981. 

p. 149. Echinella is preoccupied so needs to be renamed. [Thompson] 

p. 166. Under Hybolasioptera fasciata, "Lindemann" is properly "Lindeman". [Harris] 

p. 172. Under siskiyou, Chamaecyparis lawsonianae should be spelled lawsoniana, and Craneiobia lawsoniana should 

be spelled lawsonianae. 

p. 187. Lestodiplosis cryphali was described in Kieffer (1894ab), a newly discovered preprint of Kieffer (1894d). 

p. 189. Change "Capetown" to "Cape Town" under: Lestodiplosis moricola, this page; Mitodiplosis graminis, p. 202; 

Ochnephila socialis, p. 212; Silvestriola cincta, p. 255; and Cecidomyia deformans, p. 276. [Dorchin] 

p. 190. An earlier reference, Kieffer (1894k), was found for Lestodiplosis (as Eudiplosis) rosea Kieffer and to 

Lestodiplosis septemguttata Kieffer. 

p. 192. Under Lopesia conspicua, change caulinaris to conspicua. 

p. 197. Massalongia ruber should read rubra. Also, the two mentions of the name in the entry should be reversed so 

that Hormomyia rubra is first. [Harris]

544 

p. 212. The type depository for Neomycodiplosis ruebsaameni at the top left of page should read "SMNS." 

p. 230. The reference to Stelter (1993) in the paragraph on the top left was left out of Literature Cited. [Harris] 

p. 233. Under Rabdophaga strobilina, change Stelter (1982d) page from 760 to 76. 

p. 236. The entry for Resseliella resinicola is missing, as well as the paper by Sanui & Yukawa 1985 now added to 

Literature Cited. 

p. 238. In first column, line 8, "Fedtova" should be "Fedotova." 

p. 237. Rhizomyia jaapi (Rübsaamen) is missing. 

p. 239, I missed a synonym of Pumilomyia protrahenda De Stefani, viz., Pulmilomyia artemisiae De Stefani (1929: 

262), per Skuhravá, Skuhravý & Massa 2007. 

p. 241. The type depository for Rhopalomyia cruziana is USNM, not NYSM. 

p. 248. The main heading for R. sublessaingiana should be spelled sublessingiana. 

p. 286. Cecidomyia poculum Osten Sacken, completely left out of the catalog, should have appeared as a species no 

longer referable to Cecidomyiidae. 

p. 287. In the second column, 3rd line, remove quotation marks from Micromyiidi, and in 2nd paragraph, 3rd line, 

Mycromyidi should be spelled Micromyidi. [Jaschhof] 

p. 298. In the list of new names, Procontarinia biharana is listed twice. 

p. 300. Primavera porrecta does not need to be in the list of new combinations. 

p. 301. In the list of new combinations, Rhopalomyia monogyna is out of alphabetical order. 

p. 317. In the reference to Felt, E.P. 1915d, "30th" in "In his 30th Report ..." should be changed to "29th." 

p. 327. A preprint was newly discovered for the combination of Kieffer 1894b, d, f and g and paged as 1-11. 

p. 327. An offprint was newly discovered for Kieffer 1894j (here as Kieffer 1894k). 

p. 391. In the first column, lines 12 and 13, change "Karshomyia" to "Kiefferia." 

p. 396. Under rufomaculata Panelius, move rufa Felt higher up the column, under "rufa Dorchin".

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