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ISSN 1468-8158 

Tropical Bryology Research Reports 

No. 3 http://www.oshea.demon.co.uk/tbr.htm 2002 

Checklist and distribution of the liverworts and 

hornworts of sub-Saharan Africa, including the 

East African Islands 

(edition 1, February 2002) 

Martin J. Wigginton 

36, Big Green, Warmington, Peterborough, PE8 6TU, U.K. 

February 2002

Tropical Bryology Research Reports is published irregularly, and each issue will be 

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Editors: Brian J. O'Shea and Martin J. Wigginton 

Copyright © 2002, Tropical Bryology Research 

Date of publication: 27 February 2002 

Printed in United Kingdom 

All rights reserved. No part of this journal may be reproduced by film, microfilm or other means 

without written permission from the publisher

Tropical Bryology Research Reports (2002) 3: 1-88 

Checklist and distribution of the liverworts and hornworts of sub- 

Saharan Africa, including the East African Islands 

(edition 1, February 2002) 

Martin J. Wigginton 

36, Big Green, Warmington, Peterborough PE8 6TU, U.K. M.J.Wigginton@btinternet.com 

Abstract. The distribution by country or other geopolitical area is provided for each of the 1076 liverwort and 

hornwort taxa currently accepted in sub-Saharan Africa (i.e. Index Muscorum regions Afr 2, Afr 3 and Afr 4, 

excluding the part of Afr 4 to the south of South Africa). All country records are supported by at least one 

literature reference, and the number of taxa recorded for each country is given. Synonyms with their owning taxa 

are intercalated alphabetically in the list. 

Introduction 

Catalogues of the Hepaticae and Anthocerotae have been published for the East African Islands 

(Grolle, 1995 (1)) and for the rest of sub-Saharan Africa, including Atlantic Ocean islands and Socotra 

(Wigginton & Grolle 1996 (2)). A similar checklist of the bryophytes of North Africa was published 

by Ros et al. 1999 (3). These catalogues and checklist sought to bring together all published records of 

liverworts for these regions, and provided for each accepted taxon at least one literature reference for 

each of the countries in which it had been recorded. Together they summarised current knowledge of 

the distribution of liverworts throughout the continent. 

Since the publication of the East African Islands (1) and sub-Saharan African (2) catalogues, a 

considerable bulk of new data on the liverworts of these regions has been published, and this has 

provided the stimulus to produce a new checklist. Clearly, this checklist is not a substitute for (1) and 

(2): those publications contain much additional information and should be consulted for details of 

synonymy and literature sources for country records. Grolle 1995 (1) in addition provides essential 

notes on selected genera and on many excluded taxa. 

The scope of this checklist is all of the Index Muscorum regions Afr 2, Afr 3 and Afr 4, except for the 

part of Afr 4 which lies to the south of the African mainland (south of latitude 35°S). The latter area 

encompasses cool-temperate and subantarctic islands from St. Paul and Amsterdam islands in the 

southern Indian Ocean to the Tristan da Cunha group in the southern Atlantic, all of whose floristic 

affinities are antipodal rather than African. Western Sahara is also excluded as its floristic affinities are 

North African. There are, however, no bryophytes recorded from that geographical region (Ros et al. 

1999). 

Catalogues (1) and (2) have provided the main basis for the present checklist, and most country 

records are referred to those publications, which should be consulted for details of primary literature. 

In the few years since the catalogues appeared, many new publications have provided additional 

country records, and new taxa and synonymy, the latter being listed in Appendices 1 and 2. Also 

incorporated in this checklist are: 

a) the correction of errors, including some authorities of names, which remained undetected prior to 

the publication of (2), and the inclusion of some country records from overlooked literature,

2 M.J. Wigginton 

b) the reassignment of taxa which were placed in (2) in the genera Aneura, Aspiromitus, 

Anthoceros, Fimbriaria, Hygrolejeunea, Madotheca, Schisma and Strepsilejeunea, 

c) the reassignment of some records to Eritrea or Ethiopia following the re-checking of the 

geographical locations of places cited in (121, 502). 

In sub-Saharan Africa, including the islands, 1076 taxa (1029 species and 47 additional taxa of lower 

rank) are currently accepted in 140 genera and 46 families. One family (Wiesnerellaceae), 9 genera 

(Allisoniella, Bryopteris, Capillolejeunea, Geocalyx, Metalejeunea, Otolejeunea, Symbiezidium, 

Wiesnerella, Xylolejeunea) and 207 species or lower rank taxa have been recorded in Africa only from 

the East African Islands. 

The list of currently accepted taxa excludes those which have been placed in synonymy or have been 

published as 'Dubia. Excludenda' or 'Dubia. Indeterminate', but does include many taxa which have 

been insufficiently studied (or perhaps not re-examined since first described - which may be in the 

19th century), and some of doubtful occurrence in Africa. It seems likely that some or many of the 

latter will eventually be placed in synonymy or rejected for other reasons. Taxa whose validity, status 

or occurrence may be uncertain are not differentiated in this checklist, and reference should be made to 

(1) and (2) for further information. Two taxa, whose taxonomic position and validity is yet to be 

clarified, remain listed under the generic synonym Fimbriaria, though they are probably species of 

Asterella. 

Data for the checklist are held on the TAXA system (541), and the main taxon distribution list was 

generated directly from TAXA. The format of this checklist follows that established by O'Shea (171, 

540) for the Checklist of the mosses of sub-Saharan Africa, and subsequently adopted by Ros et al. (3) 

for the Bryophyte checklist of Northern Africa. 

The checklist is not a taxonomic document, and no new taxonomy is published here: it is merely a 

summation of published records. In its compilation, no judgements have been made by the author on 

the validity of taxa, or on the acceptability of regional records. Despite checking and re-checking, 

some errors doubtless remain undetected in this document, and literature overlooked, and the author 

would be very grateful to know of any of these. 

This checklist is published both as a printed version and an electronic version (via 

http://www.oshea.demon.co.uk/tbr.htm) so that it is made as widely available as possible. It is 

intended that further editions will be published in the same way at intervals of a few years. 

Acknowledgments 

I am indebted to Brian O'Shea for a careful reading of the text and detecting transcription and other 

errors. Many thanks also to Riclef Grolle who kindly read some of the final draft and pointed out some 

overlooked records and synonymy. However, any errors which may still remain in this document are 

entirely the responsibility of the author.

Checklist of the liverworts and hornworts of sub-Saharan Africa 3 

Map1. The countries of Index Muscorum regions Afr 2, Afr 3 and Afr 4 covered in this checklist. 

AGA Agalega Islands 

ALD Aldabra 

AMI Amirante Islands 

ANG Angola 

ANN Annobón Island 

ASC Ascension Island 

BEN Benin 

BIO Bioko (Fernando Po) 

BKN Burkina Faso 

BOT Botswana 

BUR Burundi 

CAB Cabinda 

CAF Central African Republic 

CGS Chagos Archipelago 

CHA Chad 

CMN Cameroun 

COM Comoro Islands 

CON Congo-Brazzaville 

CVI Cape Verde 

DIE Diego Garcia 

DJI Djibouti 

ERI Eritrea 

ETH Ethiopia 

GAB Gabon 

GAM Gambia 

GHA Ghana 

GNB Guinea Bissau 

GUI Guinea 

IVO Ivory Coast 

KEN Kenya 

LBR Liberia 

LES Lesotho 

MAU Mauritius 

MDG Madagascar 

MLI Mali 

MLW Malawi 

MOZ Mozambique 

MTN Mauritania 

NAM Namibia 

NGA Nigeria 

NGR Niger 

PRI Principé 

REU Réunion 

RIM Rio Muni 

ROD Rodrigues 

RWA Rwanda 

SAF South Africa 

SAO São Tomé 

SEN Senegal 

SEY Seychelles 

SIE Sierra Leone 

SOC Socotra 

SOM Somalia 

STH St. Helena 

SUD Sudan 

SWZ Swaziland 

TAN Tanzania 

TOG Togo 

UGA Uganda 

ZAI Democratic Republic of 

Congo (former Zaire) 

ZAM Zambia 

ZIM Zimbabwe


Synopsis of families and genera 

Families are arranged alphabetically within each Class and Order. The placement of genera closely 

follows Grolle 1983a (421), except for the Lejeuneaceae which follows Gradstein 1994 (122). Taxa are 

assigned to 140 genera in 46 families (excluding the generic synonym, Fimbriaria, placed in round 

brackets in this synopsis). The numbers in brackets against each genus indicate the number of taxa 

currently accepted within the genus (but this number includes doubtful taxa which have not (yet) been 

synonymised or rejected, and taxa identified only to the genus). The numbers in a small font refer to the 

African Index Muscorum region(s) in which the genus has been recorded. Since the publication of Grolle 

1995 (1) and Wigginton & Grolle 1996 (2), the genera Apomarsupella, Aureolejeunea, Kymatocalyx, 

Mnioloma, Paracromastigum, Sphenolobopsis and Xylolejeunea have been added to the sub-Saharan list, 

Hyalolepidozia has been removed, and Stenorrhipis placed in synonymy. 

ANTHOCEROTOPSIDA 

Anthocerotaceae 

Anthoceros (21) 

Megaceros (1) 2 

Phaeoceros (15) 

2 3 4N 

2 3 4N 

Dendrocerotaceae 

2 3 

Dendroceros (5) 

Notothyladaceae 

Notothylas (5) 2 

MARCHANTIOPSIDA 

CALOBRYALES 

Haplomitriaceae 

2 4N 

Haplomitrium (2) 

JUNGERMANNIALES 

Acrobolbaceae 

Acrobolbus (1) 4N 

Lethocolea (1) 

Marsupidium (1) 

Tylimanthus (3) 

2 3 4N 

2 3 4N 

2 3 4N 

Adelanthaceae 

2 3 4N 

Adelanthus (2) 

Antheliaceae 

Anthelia (1) 2 

Arnelliaceae 

Gongylanthus (4) 

Balantiopsaceae 

2 3 4N 

Isotachis (1) 

2 3 4N 

Calypogeiaceae 

Calypogeia (9) 

Mnioloma (1) 

2 3 4N 

2 3 4N 

Cephaloziaceae 

Alobiellopsis (2) 

Cephalozia (3) 

2 3 

Iwatsukia (1) 

Odontoschisma (1) 

2 3 4N 

2 3 4N 

Cephaloziellaceae 

Allisoniella (1) 3 

Cephalojonesia (1) 2 

Cephaloziella (17) 

Cylindrocolea (6) 

2 3 

Kymatocalyx (3) 

Chonecoleaceae 

Chonecolea (1) 2 

Frullaniaceae 

2 3 4N 

Frullania (54) 

Geocalycaceae 

Clasmatocolea (2) 

Conoscyphus (1) 

Geocalyx (1) 3 

Heteroscyphus (5) 

Leptoscyphus (4) 

Lophocolea (18) 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 4N 

Gymnomitriaceae 

Apomarsupella (1) 2 

Gymnomitrion (2) 

Marsupella (3) 

Stephaniella (1) 4N 

2 4N 

2 3 4N 

Herbertaceae 

2 3 4N 

Herbertus (12) 

Jungermanniaceae 

Anastrophyllum (5) 

Andrewsianthus (3) 

Chandonanthus (4) 

2 3 4N 

Cryptochila (1) 

2 3 

Gottschelia (1) 

Gymnocoleopsis (2) 

Jamesoniella (3) 

Jungermannia (11) 

2 4N 

Lophozia (3) 

2 3 

Nardia (1) 

Notoscyphus (1) 

Sphenolobopsis (1) 2 

2 3 

Syzygiella (4) 

Tetralophozia (1) 2 

2 4N 

Tritomaria (2) 

Lejeuneaceae 

Ptychanthoideae 

Ptychantheae 

2 3 4N 

2 3 

2 3 4N 

2 4N 

2 3 4N 

2 3 4N 

2 3 

2 3 4N 

Acrolejeunea (5) 

Archilejeunea (11) 

Bryopteris (1) 3 

Caudalejeunea (10) 

2 3 4N 

Frullanoides (1) 

Lopholejeunea (22) 

2 3 

Marchesinia (5) 

2 3 

Mastigolejeunea (6) 

2 3 4N 

Ptychanthus (2) 

Schiffneriolejeunea (11) 

2 3 4N 

Thysananthus (2) 

Lejeuneoideae 

Brachiolejeuneae 

Acanthocoleus (3) 

Brachiolejeunea (2) 

Odontolejeunea (1) 

2 3 

Stictolejeunea (1) 

Symbiezidium (1) 3 

Lejeuneae 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 4N 

2 3 

2 3 4N

Aureolejeunea (1) 2 

Capillolejeunea (1) 3 

Ceratolejeunea (10) 

Cheilolejeunea (29) 

Cladolejeunea (1) 2 

Drepanolejeunea (17) 

Evansiolejeunea (1) 2 

2 3 

Haplolejeunea (2) 

2 3 

Harpalejeunea (2) 

2 3 4N 

Lejeunea (57) 

2 3 

Lepidolejeunea (3) 

2 3 

Leptolejeunea (5) 

Leucolejeunea (3) 

Metalejeunea (1) 3 

Microlejeunea (13) 

Otolejeunea (2) 3 

2 3 

Prionolejeunea (2) 

2 3 

Pycnolejeunea (1) 

2 3 4N 

Taxilejeunea (6) 

Xylolejeunea (2) 3 

2 3 

2 3 4N 

2 3 4N 

Cololejeuneae 

Aphanolejeunea (10) 

Cololejeunea (107) 

2 3 4N 

Colura (22) 

Diplasiolejeunea (29) 

Lepicoleaceae 

Lepicolea (1) 4N 

Lepidoziaceae 

Acromastigum (1) 4N 

2 3 4N 

2 3 4N 


2 3 4N 

2 3 4N 

2 3 4N 

2 3 4N 

Arachniopsis (2) 

2 3 4N 

Bazzania (17) 

2 3 4N 

Kurzia (5) 

2 3 4N 

Lepidozia (14) 

Paracromastigum (2) 4N 

Psiloclada (1) 4N 

2 3 4N 

Telaranea (2) 

Mastigophoraceae 

2 3 

Mastigophora (1) 

Plagiochilaceae 

2 3 4N 

Plagiochila (69) 

Pleuroziaceae 

2 3 

Pleurozia (1) 

Porellaceae 

2 3 4N 

Porella (13) 

Pseudolepicoleaceae 

Blepharostoma (1) 2 

Radulaceae 

2 3 4N 

Radula (23) 

Scapaniaceae 

Diplophyllum (1) 2 

2 4N 

Scapania (4) 

Schistochilaceae 

Gottschea (3) 

Schistochila (4) 

2 3 4N 

METZGERIALES 

Allisoniaceae 

Calycularia (1) 2 

Aneuraceae 

Aneura (4) 

Riccardia (21) 

2 3 4N 

2 3 4N 

2 3 4N 

Fossombroniaceae 

2 3 4N 

Fossombronia (34) 

Metzgeriaceae 

2 3 4N 

Metzgeria (33) 

Pallaviciniaceae 

2 3 4N 

Jensenia (1) 

2 3 4N 

Pallavicinia (2) 

Symphyogyna (3) 

Symphyogynopsis (1) 

2 3 4N 

2 3 

SPHAEROCARPALES 

Monocarpaceae 

Monocarpus (1) 4N 

Checklist 

Riellaceae 

Riella (5) 4N 

Sphaerocarpaceae 

Sphaerocarpos (1) 4N 

MARCHANTIALES 

Aytoniaceae 

2 3 4N 

Asterella (14) 

Cryptomitrium (1) 4N 

(Fimbriaria) (2) 2 

2 4N 

Mannia (2) 

Plagiochasma (6) 

Reboulia (1) 2 

Cleveaceae 

Athalamia (2) 

2 4N 

2 3 4N 

Exormothecaceae 

2 3 4N 

Exormotheca (3) 

Lunulariaceae 

2 4N 

Lunularia (1) 

Marchantiaceae 

Dumortiera (2) 

Marchantia (8) 

Oxymitraceae 

Oxymitra (1) 4N 

Ricciaceae 

Riccia (86) 

Ricciocarpos (1) 

2 3 4N 

Targioniaceae 

Cyathodium (3) 

Targionia (3) 

2 3 4N 

2 3 4N 

2 4N 

2 3 4N 

2 3 4N 

Wiesnerellaceae 

Wiesnerella (1) 3 

The checklist is in alphabetic order of taxon name, with current names shown in bold face italic font. 

Names now placed in synonymy are indented and shown in a smaller font, together with the owning 

taxon. For synonymies, an = sign is used throughout, and no distinction made between nomenclatural and 

taxonomic synonyms. A source reference is given only for synonymy published since Grolle 1995 (1) and 

Wigginton & Grolle 1996 (2), which documents should be consulted for sources of other synonymy. 

Names published in recent literature as 'Dubia. Excludenda' and 'Dubia. Indeterminate' are also shown in


the smaller font. Explanatory notes are appended to selected taxa, especially where there has been an 

amendment to the distribution given in (1) and (2). Authorities of names follow Brummitt & Powell 1992 

(166), supplemented, including the suppression of spaces between initials and surname. 

The countries in which each current taxon has been recorded are denoted by 3-character abbreviations, 

which follow Hollis & Brummitt 1992 (539), with few exceptions. The name of the country and the Index 

Muscorum region to which it is assigned is shown in Appendix 3. The countries are given in alphabetic 

order of the abbreviation and not of the country name, which means that some names are slightly out of 

sequence - for example, Mauritius (MAU) being listed before Madagascar (MDG). However, the 

alphabetic arrangement of abbreviations makes it easier to 'read' the checklist. Literature references are 

given in brackets after each country abbreviation, and are translated in the References and Literature 

sections following this alphabetic list. 

Acanthocoleus aberrans var. laevis Gradst. ANG(2), CMN(2), KEN(2), MDG(1), NGA(2), SAF(2), 

TAN(2), UGA(2), ZAI(2), ZIM(2). A. aberrans var. laevis occurs in Africa and tropical America, whilst A. 

aberrans var. aberrans is restricted to Central America and the West Indies (Gradstein 1994 (122)). 

Acanthocoleus chrysophyllus (Lehm.) Kruijt CMN(2), IVO(53), KEN(2), LES(75), NGA(2), RWA(2), 

SAF(2,75), TAN(2), UGA(2), ZAM(2), ZIM(2). 

Acanthocoleus madagascariensis (Steph.) Kruijt BUR(2), COM(1), GUI(2), MAU(1), MDG(1,4), 

REU(1), RWA(2), SAF(2), SIE(2), TAN(2), ZIM(2). 

Acolea africana Steph. 1917 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996 (153). 

Acrobolbus bilobus (Mitt.) Steph. 1902 = Andrewsianthus bilobus (Mitt.) Grolle 1963. 

Acrobolbus campylodontus (Hook.f. et Taylor) Steph. 1902 = Adelanthus decipiens (Hook.) Mitt. 1864. 

Acrobolbus excisus (Mitt.) Schiffn. 1893 = A. ochrophyllus (Hook.f. et Taylor) R.M.Schust. 1961. 

Acrobolbus madagascariensis Steph. 1902 = Kymatocalyx madagascariensis (Steph.) Gradst. et Váòa 1999 (74). 

Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust. SAF(2). 

Acrolejeunea abnormis (Gottsche) Pearson 1892 = Phragmicoma abnormis Gottsche 1882 [Dubia et 

Excludenda] 

Acrolejeunea africana Steph., nom. inval. TAN(2). 

Acrolejeunea angustispica Steph. 1912 = A. emergens var. confertissima (Steph.) Gradst. 1975. 

Acrolejeunea aulacophora (Mont.) Steph. COM(1), MAU(1), MDG(1), MOZ(2), REU(1), TAN(2). 

Listed in error from Uganda in (2). 

Acrolejeunea borgenii Steph. 1892 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Acrolejeunea confertissima Steph. 1892 = A. emergens var. confertissima (Steph.) Gradst. 1975. 

Acrolejeunea convexa Steph. 1911 = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960. 

Acrolejeunea emergens var. emergens (Mitt.) Steph. ALD(34,35), ANG(2), BEN(11), BUR(2), CAF(2), 

COM(1), CON(2), CVI(2,37), GHA(2), GUI(2), IVO(2,53), KEN(2), MAU(1,4), MDG(1), 

MLW(2,563), NGA(2), REU(1), RIM(10), ROD(1), RWA(2), SEN(2), SEY(1,4), SIE(2), TAN(2), 

UGA(2,42,50,96), ZAI(2), ZAM(2). 

Acrolejeunea emergens var. confertissima (Steph.) Gradst. CMN(2), NGA(2), RIM(10), SEN(2), SIE(2). 

Acrolejeunea ferruginea Steph. 1895 = Schiffneriolejeunea ferruginea (Steph.) Gradst. 1974. 

Acrolejeunea fulva (Gottsche) Pearson 1892 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Acrolejeunea fuscescens Gola 1907 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 1907. 

Acrolejeunea inflexa (Gottsche) Pearson 1892 = Phragmicoma inflexa Gottsche 1882 [Dubia et Excludenda]. 

Acrolejeunea molleri (Steph.) Paris 1906 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Acrolejeunea occulta Steph. 1888 = Schiffneriolejeunea occulta (Steph.) Gradst. 1974. 

Acrolejeunea orchidae Pearson 1922 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Acrolejeunea pappeana (Nees) Schiffn. 1894 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Acrolejeunea parviloba Steph. 1890 = Schiffneriolejeunea parviloba (Steph.) Gradst. 1974. 

Acrolejeunea pusilla Steph. 1908, nom. nud. [non (Steph.) Grolle et Gradst. 1974] = A. emergens (Mitt.) Steph. 

1895. 

Acrolejeunea pycnoclada (Taylor) Schiffn. GHA(2), MDG(1), NGA(2), REU(1), SEY(1), ZAI(2). 

Acrolejeunea renauldii Steph. 1888, nom. inval. = A. aulacophora (Mont.) Steph. 1895.


Acrolejeunea roccatii Gola 1907 = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960. 

Acromastigum exile (Lindenb.) A.Evans SAF(2). 

Adelanthus decipiens (Hook.) Mitt. BIO(2,33), CMN(2), COM(4), ETH(2), MAU(1), MLW(2,109), 

REU(1), RWA(142), SAF(2,75), STH(2), TAN(2), UGA(42,50), ZAI(2). 

Adelanthus lindenbergianus (Lehm.) Mitt. KEN(2), LES(75), MAU(1), MDG(1), MLW(109), REU(1), 

RWA(2), SAF(2,75), TAN(2), UGA(2), ZAI(2). 

Adelanthus sphalerus (Hook.f. et Taylor) Steph. 1908 = A. lindenbergianus (Lehm.) Mitt. 1864. 

Adelanthus unciformis (Hook.f. et Taylor) Spruce 1876 = A. lindenbergianus (Lehm.) Mitt. 1864. 

Aitonia eximia Schiffn. in Steph. 1895 = Plagiochasma eximium (Schiffn. in Steph.) Steph. 1898. 

Aitonia fischeriana Steph. 1895 = Plagiochasma appendiculatum Lehm. et Lindenb. 1832. 

Aitonia microcephala Steph. 1895 = Plagiochasma microcephalum (Steph.) Steph. 1898. 

Alicularia vermicularis (Lehm.) Gottsche et al. 1844 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Allisoniella nigra (Rodway) R.M.Schust. REU(1). 

Alobiella borbonica (Steph.) Steph. 1908 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Alobiella chevalieri Steph. 1908 = Cylindrocolea chevalieri (Steph.) R.M.Schust. 1969. 

Alobiella heteromorpha (Lehm.) Steph. 1908 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969. 

Alobiella pillansii (Sim) S.W.Arnell 1963 = Alobiellopsis pillansii (Sim) R.M.Schust. 1969. 

Alobiellopsis heteromorpha (Lehm.) R.M.Schust. LES(75), MAS(1), MAU(1), MLW(114), SAF(2). 

Alobiellopsis pillansii (Sim) R.M.Schust. SAF(2). 

Anastrophyllum auritum (Lehm.) Steph. BIO(36), CMN(2), ETH(2), KEN(2,71,665), LES(75), 

MLW(114), RWA(2), SAF(2,75), TAN(2), UGA(2,42,96), ZAI(2). 

Anastrophyllum bessonii Steph. 1893 = Gottschelia schizopleura (Spruce) Grolle 1968. 

Anastrophyllum calcaratum Steph. 1911 = A. auritum (Lehm.) Steph. 1901. 

Anastrophyllum crebrifolium (Hook.f. et Taylor) Steph. REU(45). 

Anastrophyllum gambaragarae Gola 1907 = A. auritum (Lehm.) Steph. 1901. 

Anastrophyllum grossitextum Steph. 1911 = A. auritum (Lehm.) Steph. 1901. 

Anastrophyllum hintzeanum Steph. 1917 = A. auritum (Lehm.) Steph. 1901. 

Anastrophyllum leucocephalum (Taylor) Steph. 1893 = A. crebrifolium (Hook.f. et Taylor) Steph. 1893 (172, 

420). 

Anastrophyllum minutum (Schreb.) R.M.Schust. CMN(2), KEN(2), SAF(2,75), TAN(2), UGA(2), 

ZAI(667). 

Anastrophyllum obtusatum (Hook.f. et Taylor) Steph. 1901 = A. subcomplicatum (Lehm. et Lindenb.) Steph. 

1893. 

Anastrophyllum piligerum (Nees) Steph. ASC(2), MAU(1), MDG(1,4), MLW(114), REU(1), RWA(2), 

SEY(1,4), TAN(2), ZIM(2). 

Anastrophyllum schizopleurum (Spruce) Steph. 1893 = Gottschelia schizopleura (Spruce) Grolle 1968. 

Anastrophyllum subcomplicatum (Lehm. et Lindenb.) Steph. STH(2). 

Andrewsianthus aberrans (Nees et Mont.) Grolle MAU(1), MDG(4), REU(1,4). 

Andrewsianthus bilobus (Mitt.) Grolle BIO(2,33,36), CMN(2), ETH(2), RWA(2), TAN(2), UGA(2), 

ZAI(2). Listed in error from South Africa in (2). The specimen is Gymnocoleopsis capensis (163,164). 

Andrewsianthus jamesonii (Mont.) Váòa KEN(2), RWA(2), TAN(2), UGA(2), ZAI(2). 

Andrewsianthus kilimanjaricus (S.W.Arnell) Grolle et Váòa 1980 = A. jamesonii (Mont.) Váòa 1983. 

Aneura amazonica Spruce 1885 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Aneura angusticosta Steph. 1913 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Aneura caespitans Steph. 1891 = Riccardia sp. 

Aneura comosa Steph. 1890 = Riccardia sp. 

Aneura compacta Steph. 1893 = Riccardia compacta (Steph.) S.W.Arnell 1952. 

Aneura congoana Steph. 1917 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Aneura congoensis Steph. 1917 = Riccardia longispica (Steph.) Pearson 1893. 

Aneura corbieri Steph. 1917 = Riccardia sp. 

Aneura dismieri Steph. 1917 = Riccardia sp. 

Aneura erosa Steph. 1891 = Riccardia erosa (Steph.) E.W.Jones 1956. 

Aneura erronea Steph. TAN(2). 

Aneura exigua Steph. 1917 = Riccardia longispica (Steph.) Pearson 1893. 

Aneura fastigiata (Lehm.) Gottsche et al. 1846 = Riccardia fastigiata (Lehm.) Trevis. 1877.


Aneura grosselimbata Steph. 1917 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Aneura holstii Steph. 1917 = Riccardia holstii (Steph.) E.W.Jones 1956. 

Aneura inconspicua Steph. 1893 = Riccardia erosa (Steph.) E.W.Jones 1956. 

Aneura keniae Gola 1914 = Riccardia sp. 

Aneura latissima Spruce MAU(1), PRI(128), REU(1). 

Aneura laurentiana Steph. 1917 = A. pinguis (L.) Dumort. 1831 (652). 

Aneura lepervanchei Steph. 1899 = Riccardia sp. 

Aneura limbata Steph. 1891 = Riccardia limbata (Steph.) E.W.Jones 1956. 

Aneura longispica Steph. 1890 = Riccardia longispica (Steph.) Pearson 1893. 

Aneura nudiflora Steph. 1890 = Riccardia nudiflora (Steph.) Grolle 1995. 

Aneura palmata (Hedw.) Dumort. 1831 = Riccardia palmata (Hedw.) Carruth. 1865 [Dubia et Excludenda]. 

Aneura papulosa Steph. 1917, hom. illeg. [non Steph. 1893] = Riccardia longispica (Steph.) Pearson 1893. 

Aneura piliflora Steph. 1917 = Riccardia longispica (Steph.) Pearson 1893. 

Aneura pinguis (L.) Dumort. MAS(1), SAF(652), SEY(1), TAN(2). 

Aneura pinguis (L.) Dumort. sens. lat. ANG(2), CMN(2), GHA(2), NGA(2), SAO(2), SIE(2), ZAI(2). 

Aneura pinnatifida (F.Weber) Nees 1838 = Riccardia pinnatifida (F.Weber) Carruth. 1865 [Dubia et 

Excludenda]. 

Aneura pseudopinguis (Herzog) Pócs IVO(2), LES(75), MLW(114), RIM(10), SAF(2), SEY(4), 

TAN(2), UGA(42,50), ZAI(2), ZIM(2). 

Aneura ramosissima Steph. 1891 = Riccardia ramosissima (Steph.) Grolle 1995. 

Aneura reticulata Steph. 1891 = Riccardia limbata (Steph.) E.W.Jones 1956. 

Aneura saccatiflora Steph. 1890 = Riccardia saccatiflora (Steph.) S.W.Arnell 1952. 

Aneura stephanii Besch. ex Steph. 1899 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Aneura submarginata S.W.Arnell 1952 = A. pinguis (L.) Dumort. 1831 (652). 

Aneura tenera Steph. 1917 = Riccardia erosa (Steph.) E.W.Jones 1956. 

Aneura travisiana Pearson 1921 = Riccardia amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Anomalolejeunea decemplicata Steph. 1913 = Cheilolejeunea pluriplicata (Pearson) R.M.Schust. 1980. 

Anomalolejeunea pluriplicata (Pearson) Schiffn. 1893 = Cheilolejeunea pluriplicata (Pearson) R.M.Schust. 

1980. 

Anomalolejeunea pluriplicata var. tabularis S.W.Arnell 1953 = Cheilolejeunea pluriplicata (Pearson) 

R.M.Schust. 1980. 

Anthelia hirtella (F.Weber) Steph. 1891 = Chandonanthus hirtellus (F.Weber) Mitt. 1873. 

Anthelia juratzkana (Limpr.) Trevis. TAN(2). 

Anthoceros aethyopicus Gola 1914 = Phaeoceros sp. 

Anthoceros brunnthaleri Steph. 1913 = Phaeoceros sp. 

Anthoceros buettneri Steph. GHA(2), TOG(2). Listed in error from Sierra Leone in (2). 

Anthoceros caucasicus Steph. SIE(2), ZAI(2). 

Anthoceros chevalieri Steph. ZAI(2). 

Anthoceros communis Steph. 1897 = Phaeoceros carolinianus (Michx.) Prosk. 1951. 

Anthoceros cristatus Steph. ASC(2). 

Anthoceros decurvus Mitt. 1860 = Notothylas decurva (Mitt.) Steph. 1901. 

Anthoceros dichotomus Raddi 1818 = Phaeoceros bulbiculosus (Brot.) Prosk. 1954. 

Anthoceros dilatatus Steph. BIO(2,33). 

Anthoceros dimorphus Sim SAF(2), ZIM(2). 

Anthoceros ecklonii Steph. 1916 = Phaeoceros carolinianus (Michx.) Prosk. 1951. 

Anthoceros flagellaris Mitt. 1873 = Megaceros flagellaris (Mitt.) Steph. 1916. 

Anthoceros floribundus Steph. ASC(2). 

Anthoceros fuciformis Mont. ANN(2,33), REU(1), ROD(1), TAN(2). This species was placed under 

Folioceros in (1) and under Anthoceros in (2). Its placement remains uncertain because of the lack of a consensus 

on the status of Folioceros. The latter was treated as independent genus by Hässel de Menendez (1988), Hyvönen 

& Piippo (1993) and Piippo (1993), but treated as a subgenus of Anthoceros by Hasegawa (1994; 1995). 

Anthoceros fulvisporus Steph. 1916 = Phaeoceros fulvisporus (Steph.) J.Haseg. 1993. 

Anthoceros fusiformis Austin MAU(1). 

Anthoceros gasangorii Gola KEN(2). 

Anthoceros incurvus Steph. CMN(2). 

Anthoceros javanicus Nees 1830 = Dendroceros javanicus (Nees) Gottsche et al. 1846.


Anthoceros jollyanus Steph. 1916 = Phaeoceros jollyanus (Steph.) S.W.Arnell 1956. 

Anthoceros laevis L. 1753 = Phaeoceros laevis (L.) Prosk. 1951 [Dubia et Excludenda]. 

Anthoceros mandonii Steph. 1916 = A. caucasicus Steph. 1914. 

Anthoceros mildbraedii Steph. 1923 = Phaeoceros sp. 

Anthoceros minutus Mitt. 1877 = Phaeoceros minutus (Mitt.) S.W.Arnell 1963. 

Anthoceros myriandroecius Steph. KEN(2,665), RWA(2), TAN(2), UGA(96,115). 

Anthoceros natalensis Steph. LES(75), MLW(2), SAF(2). 

Anthoceros parvifrons Steph. 1912 = Phaeoceros sp. 

Anthoceros pinnatus Steph. CMN(2), SAO(2), TAN(2). 

Anthoceros pseudocostus Steph. 1916 = Phaeoceros sp. 

Anthoceros punctatus L. CVI(2,37), RWA(2), ZAI(2). 

Anthoceros rossoi Gola 1914 = Phaeoceros sp. 

Anthoceros sambesianus Steph. LES(75), RHO(2), RWA(2), UGA(115). 

Anthoceros schroederi Steph. TAN(2). 

Anthoceros stuhlmannii Steph. 1916 = Folioceros fuciformis (Mont.) Bharadw. 1975. 

Anthoceros tenuissimus Steph. 1893 = Phaeoceros sp. 

Anthoceros tigrinus Gola 1914 = Phaeoceros sp. 

Anthoceros usambarensis Steph. 1916, probably = Phaeoceros carolinianus (Michx.) Prosk. 1951. 

Anthoceros volkensii Steph. TAN(2). 

Anthoceros sp. (Aspiromitus dismieri Steph.) MDG(1). 

Anthoceros sp. (Aspiromitus expansus Steph.) CMN(2). Probably a species of Anthoceros, as Stephani 1916 

(140) described it as having "frons cavernosa ... sporae fuscae". 

Anthoceros sp. (Aspiromitus hildebrandtii Steph.) MDG(1). 

Aphanolejeunea capensis (S.W.Arnell) S.W.Arnell BUR(2), MLW(2,563), RWA(2), SAF(2), TAN(2), 

UGA(50), ZAI(2), ZIM(2). 

Aphanolejeunea corniculata Tixier in Pócs 1984, nom. inval. [Dubia et Excludenda]. 

Aphanolejeunea diaphana (A.Evans) R.M.Schust. 1980 = A. truncatifolia Horik. 1934 (117). 

Aphanolejeunea exigua A.Evans 1911 = A. microscopica var. exigua (A.Evans) Pócs et Lücking 1996 (117). 

Aphanolejeunea exigua var. africana Pócs 1984 = A. microscopica var. africana (Pócs) Pócs et Lücking 1996 

(117). 

Aphanolejeunea fadenii (Pócs) Pócs KEN(2), RWA(2), TAN(2), UGA(115). 

Aphanolejeunea jovetastiana Pócs CMN(2), GHA(2), IVO(2), RIM(10). 

Aphanolejeunea lisowskii Pócs RIM(10), UGA(115), ZAI(2). 

Aphanolejeunea mamillata sens. auct. [non (Ångstr.) Steph. 1916] = A. capensis (S.W.Arnell) S.W.Arnell 1957. 

Aphanolejeunea microscopica var. africana (Pócs) Pócs et Lücking KEN(2), MLW(563), RWA(2), 

TAN(2), UGA(42,96), ZAI(2). 

Aphanolejeunea microscopica var. exigua (A.Evans) Pócs et Lücking COM(1), IVO(2), KEN(2), 

MDG(1), MLW(563), RWA(2), TAN(2), UGA(42), ZAI(2). 

Aphanolejeunea microscopica var. microscopica (Taylor) A.Evans COM(4), MLW(563), TAN(2), 

UGA(42). 

Aphanolejeunea minima Tixier CMN(2). 

Aphanolejeunea moramangae (Tixier) Pócs COM(1), MDG(1), RIM(10), TAN(2), UGA(42,115), 

ZAI(2). 

Aphanolejeunea runssorensis Steph. 1916 = Cololejeunea runssorensis (Steph.) Pócs 1975. 

Aphanolejeunea subdiaphana (Ast) Pócs 1984 = A. truncatifolia Horik. 1934 (117). 

Aphanolejeunea truncatifolia Horik. BIO(72), MLW(563), UGA(115). 

Aphanolejeunea zenkeri Steph. 1916 = Cololejeunea zenkeri (Steph.) E.W.Jones 1954. 

Apomarsupella africana (Steph. ex Bonner) R.M.Schust. KEN(2), RWA(2), TAN(2), UGA(2), ZAI(2). 

Arachniopsis coactilis Spruce 1882 = A. diacantha (Mont.) M.Howe 1902. 

Arachniopsis diacantha (Mont.) M.Howe ANN(2,33), BIO(36), COM(4), GHA(2), IVO(2), 

KEN(71,665), MAU(1), MDG(1,4), MLW(2,92), MOZ(2), REU(1), RIM(10), RWA(2), SAF(2), 

SEY(1,4,13), SIE(2), TAN(2), UGA(42,50), ZAI(2), ZAM(2), ZIM(2). 

Arachniopsis diplopoda Pócs COM(4), MDG(1,4), REU(4). The record from D.R. Congo (ZAI), formerly 

considered to be doubtful, is now discounted. This species is now regarded as endemic to the East African Islands


(Pócs 1995 (4)). 

Arachniopsis madagascariensis Steph. 1922 = A. diacantha (Mont.) M.Howe 1902. 

Arachniopsis succulenta (Sim) S.W.Arnell 1955 = Paracromastigum succulentum (Sim) J.J.Engel et 

G.L.S.Merr. 2001 (556). 

Archilejeunea abbreviata (Mitt.) Vanden Berghen ANG(2), CAF(2), CMN(2), GAB(2), GHA(2), 

GUI(2), MLW(563), NGA(2), RIM(10), SIE(2), TAN(2), UGA(50). 

Archilejeunea africana Steph. BIO(2,33), CAF(2), CMN(2), CON(2), GAB(2), GHA(2), IVO(53), 

NGA(2), PRI(48), RIM(10), SIE(2), TAN(2), UGA(42,50,96), ZAI(2). 

Archilejeunea alata Steph. COM(1), MAU(1), MDG(1). 

Archilejeunea apiculata Pearson 1931 = Marchesinia excavata (Mitt.) Schiffn. 1893. 

Archilejeunea autoica Vanden Berghen CAF(2), CMN(2), GAB(2), GHA(2), IVO(2,53), NGA(2), 

RIM(10), SIE(2), UGA(42,115). 

Archilejeunea brevilobula Steph. MDG(1). 

Archilejeunea elobulata Steph. CAF(2), IVO(2,53), SAF(2), TAN(2), ZAI(2). 

Archi-lejeunea erronea Steph. 1888 = Leucolejeunea clypeata (Schwein) A.Evans 1907 [Dubia et Excludenda]. 

Grolle 1995 (1) placed the South African Archilejeunea erronea in synonymy with Leucolejeunea 

rotundistipula, but Gradstein & Geissler 1997 (55) re-examined the holotype, and found it inseparable from 

Leucolejeunea clypeata, a species endemic to eastern North America. Because of this extreme disjunction, 

they concluded that specimens had been confused by Stephani, and that there had been a labelling error. 

Archilejeunea jonesii Vanden Berghen CAF(2), IVO(53), NGA(2), RIM(10). 

Archilejeunea linguifolia Steph. CAF(2), CMN(2), GAB(2), IVO(2), NGA(2), RIM(10), SIE(2), 

TOG(2), ZAI(2). 

Archilejeunea mariana (Gottsche) Steph. 1897 = Cheilolejeunea mariana (Gottsche) B.Thiers et Gradst. 1989 

[Dubia et Excludenda]. 

Archilejeunea mauritiana Lindenb. ex Steph. MAS(1), MAU(1), TAN(2). 

Archilejeunea nobilis (Gottsche) Steph. MDG(1). 

Archilejeunea pappeana (Nees) Sim 1926 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Archilejeunea parvistipula Steph. 1924 = A. africana Steph. 1911. 

Archilejeunea planiuscula (Mitt.) Steph. SEY(1). 

Archilejeunea pobeguinii Steph. 1911 = A. abbreviata (Mitt.) Vanden Berghen 1951. 

Archilejeunea rotalis (Hook.f. et Taylor) Steph. 1911 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et 

Geissler 1997 (55). 

Archilejeunea rotundistipula (Lindenb. ex Lehm.) Sim 1926 = Leucolejeunea rotundistipula (Lindenb. ex 

Lehm.) A.Evans 1907. 

Archilejeunea staudtiana Steph. 1911 = A. africana Steph. 1911. 

Archilejeunea subinermis Steph. 1911, nom. nud. = A. africana Steph. 1911. 

Archilejeunea taxiantha Herzog, in sched. = A. africana Steph. 1911. 

Archilejeunea xanthocarpa (Lehm. et Lindenb.) Steph. 1895 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) 

A.Evans 1907. 

Aspiromitus dilatatus (Steph.) Steph. 1916 = Anthoceros dilatatus Steph. 1887. 

Aspiromitus dismieri Steph. 1916 = Anthoceros sp. 

Aspiromitus expansus Steph. 1916 = Anthoceros sp. 

Aspiromitus fuciformis (Mont.) Steph. 1916 = Folioceros fuciformis (Mont.) Bharadw. 1975. 

Aspiromitus hildebrandtii Steph. 1916 = Anthoceros sp. 

Aspiromitus incurvus (Steph.) Steph. 1916 = Anthoceros incurvus Steph. 1893. 

Aspiromitus maranguensis Steph. 1916 = Phaeoceros maranguensis (Steph.) Bapna 1967. 

Aspiromitus pinnatus (Steph.) Steph. 1916 = Anthoceros pinnatus Steph. 1887. 

Aspiromitus punctatus (L.) Schljakov 1976 = Anthoceros punctatus L. 1753. 

Aspiromitus schroederi Steph. 1916 = Anthoceros schroederi Steph. 1912. 

Asterella abyssinica (Gottsche) Grolle BUR(2), CMN(2), ETH(2), MLW(85), RWA(2), SAF(85), 

SIE(2), TAN(2), ZAI(2), ZIM(2). 

Asterella angolensis (Steph.) S.W.Arnell 1963 = A. wilmsii (Steph.) S.W.Arnell 1963. 

Asterella bachmannii (Steph.) S.W.Arnell BOT(2), LES(2,75), MLW(2,29), SAF(2,75), SWZ(2), 

ZAI(2), ZIM(2). 

Asterella boryana (Mont.) H.A.Mill. 1981 = A. syngenesica (Bory) Grolle 1974.


Asterella coronata (Steph.) H.A.Mill. MDG(1). 

Asterella dissoluta (Steph.) Grolle TAN(2). 

Asterella khasyana (Griff.) Pandé et al. BUR(93), UGA(93). 

Asterella linearis (Steph.) M.Howe KEN(2,71), TAN(2). 

Asterella marginata (Nees) S.W.Arnell SAF(2). 

Asterella muscicola (Steph.) S.W.Arnell BOT(2), LES(2,75), MLW(2), SAF(2), ZIM(2). 

Asterella pappii (Gola) Grolle ERI(121), ETH(2), SOC(2). 

Asterella syngenesica (Bory) Grolle MAU(1), REU(1). 

Asterella wilmsii (Steph.) S.W.Arnell ANG(2), LES(2,75), MDG(1), MLW(2,29), MOZ(2), SAF(2,75), 

SWZ(2), ZIM(2). 

Asterella sp. (Fimbriaria incrassata Steph.). ETH(2). Provisionally placed in synonymy with Asterella 

abyssinica by Long & Grolle, in litt. 

Asterella sp. (Fimbriaria kamerunensis Steph.). CMN(2). Provisionally placed in synonymy with Asterella 

abyssinica by Long & Grolle, in litt. 

Asterella sp. (Fimbriaria preussii Schiffn. ex Steph.). ETH(2). Provisionally placed in synonymy with 

Asterella bachmannii by Long & Grolle, in litt. 

Asterella sp. (Fimbriaria volkensii Steph.). KEN(2), TAN(2), UGA(2). Provisionally placed in synonymy 

with Asterella dissoluta by Long & Grolle, in litt. 

Athalamia pulcherrima (Steph.) S.Hatt. ETH(2). 

Athalamia spathysii (Lindenb.) S.Hatt. NAM(2), TAN(2). 

Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler STH(2,49,55). 

Aytonia abyssinica (Gola) Cufod. 1952 = Plagiochasma rupestre var. rupestre (J.R.Forst. et G.Forst.) Steph. 

1898. 

Aytonia appendiculata (Lehm. et Lindenb.) Kuntze 1891 = Plagiochasma appendiculatum Lehm. et Lindenb. 

1832. 

Aytonia appendiculata var. erythraeae (Gola) Cufod. 1952 = Plagiochasma appendiculatum Lehm. et Lindenb. 

1832. 

Aytonia beccariana (Steph.) Cufod. 1952 = Plagiochasma beccarianum Steph. 1898. 

Aytonia eximia Schiffn. ex Steph. 1895 = Plagiochasma eximium (Schiffn. ex Steph.) Steph. 1898. 

Aytonia microcephala Steph. 1895 = Plagiochasma microcephalum (Steph.) Steph. 1898. 

Aytonia rupestris J.R.Forst. et G.Forst. 1775 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Aytonia rupestris var. albociliata (Lindenb. et Nees) Cufod. 1952 = Plagiochasma rupestre (J.R.Forst. et 

G.Forst.) Steph. 1898. 

Aytonia schimperi (Steph.) Cufod. 1952 = Plagiochasma eximium (Schiffn.) Steph. 1898. 

Bazzania adnexa (Lehm. et Lindenb.) Trevis. 1877 [Dubia et Excludenda]. 

Bazzania approximata Onr. SEY(1,4). 

Bazzania borbonica (Steph.) Steph. 1891 = B. praerupta (Reinw. et al.) Trevis. 1877. 

Bazzania boschiana (Steph.) S.W.Arnell 1965, nom. inval. = B. decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Bazzania comorensis Steph. COM(1). 

Bazzania convexa (Lindenb.) Trevis. 1877 = B. nitida (F.Weber) Grolle 1960. 

Bazzania cucullata Onr. SEY(1,4). 

Bazzania curvidens Steph. MDG(1). 

Bazzania deciduifolia Onr. MDG(1). 

Bazzania decrescens (Lehm. et Lindenb.) Trevis. BIO(36), COM(1,4), CON(2), GAB(2), KEN(2), 

MAS(1), MAU(1), MDG(1), MLW(2,92), MOZ(2), REU(1), RWA(2), SAF(2), SEY(1,4), TAN(2), 

UGA(2), ZAI(2), ZAM(2), ZIM(2). 

Bazzania decrescens Mitt. in Melliss 1875, nom. inval. = B. decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Bazzania decrescens ssp. molleri (Steph.) E.W.Jones ANN(2,33), BIO(72), CMN(2), CON(2), GAB(2), 

GHA(2), IVO(51), MLW(92), MOZ(2), RIM(10), SAO(2), SIE(2), ZAI(2), ZIM(2). 

Bazzania decrescens ssp. pumila (Mitt.) Pócs COM(4), MDG(4), MLW(92), RWA(2), TAN(2), 

UGA(50), ZAI(2). 

Bazzania fusca Steph. 1893 = B. decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Bazzania integra (Nees et Mont.) Steph. 1894 = B. decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Bazzania lacouturei (Steph.) S.W.Arnell 1965, nom. inval. = Bazzania sp.


Bazzania laxifolia (Steph.) Demaret 1942 = B. roccatii Gola 1907. 

Bazzania longa (Nees) Trevis. 1877 [Dubia et Excludenda]. 

Bazzania madagassa (Steph.) S.W.Arnell 1965, nom. inval. = Bazzania sp. 

Bazzania mascarena (Steph.) Herzog COM(1), MDG(1), REU(1). 

Bazzania molleri Steph. 1891 = B. decrescens ssp. molleri (Steph.) E.W.Jones 1975. 

Bazzania nitida (F.Weber) Grolle BIO(36), BUR(2), CMN(2), COM(1), MAU(1,4), MDG(1,4), 

MLW(2,92), REU(1), RWA(2), SAF(2), SEY(1,4), TAN(2), UGA(50), ZAI(2). 

Bazzania papillosa S.W.Arnell MAU(1). 

Bazzania pellucida Sim 1926 = Acromastigum exile (Lindenb.) A.Evans 1934. 

Bazzania perrotana (Steph.) E.W.Jones MDG(1). 

Bazzania perrotana (Steph.) S.W.Arnell 1965, nom. inval. = B. mascarena (Steph.) Herzog 1947. 

Bazzania praerupta (Reinw. et al.) Trevis. BIO(36), MAS(1), MAU(1), MDG(1,4), MLW(2,92), 

REU(1), SAF(2), TAN(2). 

Bazzania pulvinata Steph. 1891 = B. decrescens ssp. pumila Pócs 1994. 

Bazzania pumila Mitt. 1886 = B. decrescens ssp. pumila (Mitt.) Pócs 1994. 

Bazzania radicans Sim 1926 = B. decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Bazzania reflexa (Gottsche) Steph. MDG(1). 

Bazzania robertii (Steph.) S.W.Arnell 1965, nom. inval. = B. decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Bazzania roccatii Gola CMN(2), MLW(92), RWA(2), TAN(2), UGA(2,42), ZAI(2). 

Bazzania schismoidea (Steph.) Steph. 1891 = B. praerupta (Reinw. et al.) Trevis. 1877. 

Bazzania seychellarum Gottsche ex Steph. 1893 = B. decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Bazzania sp. (Mastigobryum lacouturei Steph. = Bazzania lacouturei (Steph.) S.W.Arnell, nom. inval.) 

MDG(1). 

Bazzania sp. (Mastigobryum madagassum Steph. = Bazzania madagassa (Steph.) S.W.Arnell, nom. 

inval.) MDG(1). 

Bazzania stephanii (J.B.Jack) Steph. 1891 = B. nitida (F.Weber) Grolle 1960. 

Bazzania stolonifera (Sw.) Trevis. 1877 [Dubia et Excludenda]. 

Bellincinia abyssinica (Nees) Kuntze 1891 = Porella abyssinica (Nees) Trevis. 1877. 

Blasia javanica Sande Lac. 1856 = Notothylas javanica (Sande Lac.) Gottsche 1858. 

Blepharostoma cavallii Gola 1907 = Tetralophozia cavallii (Gola) Váòa 1993. 

Blepharostoma trichophyllum (L.) Dumort. KEN(2), RWA(2), TAN(2), UGA(2,96). 

Blytia byssophora (Lehm. et Lindenb.) Nees 1846 = Pallavicinia lyellii (Hook.) Carruth. 1865. 

Brachiolejeunea africana Steph. 1912 = Mastigolejeunea florea (Mitt.) Steph. 1912. 

Brachiolejeunea assimilis Steph. 1911 = Mastigolejeunea nigra (Steph.) Steph. 1895. 

Brachiolejeunea bidens Steph. 1912 = Mastigolejeunea auriculata (Wils.) Schiffn. 1893. 

Brachiolejeunea camerunensis E.W.Jones et Vanden Berghen 1951 = Frullanoides tristis (Steph.) van Slageren 

1985. 

Brachiolejeunea camerunensis var. angolensis S.W.Arnell 1962 = Frullanoides tristis (Steph.) van Slageren 

1985. 

Brachiolejeunea confertifolia Steph. 1912 = Frullanoides tristis (Steph.) van Slageren 1985. 

Brachiolejeunea crenata Sim 1926 = Frullanoides tristis (Steph.) van Slageren 1985. 

Brachiolejeunea hildebrandtii Steph. 1912 = Mastigolejeunea auriculata (Wils.) Schiffn. 1893. 

Brachiolejeunea hildebrandtii var. pluriplicata Gola 1914 = Frullanoides tristis (Steph.) van Slageren 1985. 

Brachiolejeunea laxifolia (Taylor) Schiffn. SAO(2). 

Brachiolejeunea nathalaensis Sim 1926 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988. 

Brachiolejeunea nigra (Steph.) Steph. 1912 = Mastigolejeunea nigra Steph. 1895. 

Brachiolejeunea parviflora Steph. 1912 = Frullanoides tristis (Steph.) van Slageren 1985. 

Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst. MDG(1), SAF(2), ZIM(2). The types of 

Dicranolejeunea lillieana, from Kenya, placed by Kruijt & Gradstein 1986 (133) under Brachiolejeunea 

phyllorhiza, are species of Acanthocoleus (Gradstein 1994 (122)). 

Brachiolejeunea rhodesica Vanden Berghen 1951 = Mastigolejeunea rhodesica (Vanden Berghen) E.W.Jones 

1957. 

Brachiolejeunea thomeensis Steph. 1912 = B. laxifolia (Taylor) Schiffn. 1893. 

Brachiolejeunea tristis (Steph.) Steph. 1912 = Frullanoides tristis (Steph.) van Slageren 1985. 

Brachiolejeunea usambarensis Steph. 1912 = Mastigolejeunea nigra Steph. 1895. 

Bryopteris filicina (Sw.) Nees 1845 [Dubia et Excludenda].


Bryopteris filicina var. fruticosa Gottsche et al. 1845, nom. inval. = Dendrolejeunea fruticosa (Lindenb. et 

Gottsche) Lacout. 1908 [Dubia et Excludenda]. 

Bryopteris fruticosa Lindenb. et Gottsche 1847 = Dendrolejeunea fruticosa (Lindenb. et Gottsche) Lacout. 1908 


Bryopteris gaudichaudii Gottsche MDG(1,4), REU(1). 

Bryopteris trinitensis (Lehm. et Lindenb.) Gottsche et al. 1845 = B. filicina (Sw.) Nees 1845 [Dubia et 

Excludenda]. 

Calobryum blumii Nees 1846 = Haplomitrium blumii (Nees) R.M.Schust. 1963. 

Calycularia crispula Mitt. ETH(2), KEN(71), MLW(2,29), TAN(2), ZAM(2). 

Calycularia golae Gerola 1947 = C. crispula Mitt. 1861. 

Calypogeia afrocaerulea E.W.Jones BIO(36), BUR(2), CMN(2), KEN(2), REU(4), RWA(2), SAO(2), 

SEY(4), TAN(2), UGA(42,50), ZAI(2). 

Calypogeia annabonensis Steph. ANN(2,33), GUI(2), KEN(2), SEY(1,4), TAN(2). 

Calypogeia arguta Nees et Mont. BUR(2), MAU(1,4), MDG(1), MLW(2), REU(1,4), SAF(2), SWZ(2), 

TAN(2), ZAI(2). 

Calypogeia bidentula (F.Weber) Nees LES(75), MDG(1), REU(1), RWA(2), SAF(2,75), ZIM(2). 

Calypogeia bidentula var. grandiretis S.W.Arnell 1963 = C. bidentula (F.Weber) Nees 1845. 

Calypogeia capensis S.W.Arnell 1953 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Calypogeia ericetorum Raddi 1818 = Gongylanthus ericetorum (Raddi) Nees 1836. 

Calypogeia fissa (L.) Raddi ANG(2), ANN(2,33), CMN(2), COM(4), KEN(71), REU(4), RWA(2), 

SAF(2), SAO(2), SIE(2), SWZ(2), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Calypogeia fusca (Lehm.) Steph. 1908 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Calypogeia latifolia Steph. 1908 = C. arguta Nees et Mont. 1838. 

Calypogeia longifolia Steph. BUR(2), LES(2), MDG(1), SAF(2), SIE(2), SWZ(2), ZIM(2). 

Calypogeia mascarenensis Bischl. COM(4), MAU(1), MDG(1), REU(1,4). 

Calypogeia microstipula (Steph.) Steph. COM(1), MDG(1), REU(1,4), SAF(2). 

Calypogeia muelleriana (Schiffn.) Müll.Frib. ETH(2). 

Calypogeia repandifolia Steph. 1924 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118) 

Campylolejeunea peculiaris (Herzog) Amakawa 1960 = Cololejeunea inflectens (Mitt.) Benedix 1953. 

Capillolejeunea mascarena S.W.Arnell COM(4), MAU(1). 

Cardiolejeunea dipterocarpa (E.W.Jones) R.M.Schust. 1980 = Lejeunea dipterocarpa E.W.Jones 1972. 

Cardiolejeunea papilionacea (Steph.) R.M.Schust. 1980 = Lejeunea papilionacea Steph. 1892. 

Carpolepidum dichotomum P.Beauv. 1805 = Plagiochila dichotoma (P.Beauv.) Nees et Mont. 1836. 

Carrpos sphaerocarpus (D.J.Carr) Prosk. 1961 = Monocarpus sphaerocarpus D.J.Carr 1956. 

Caudalejeunea acutifolia Gerola ETH(2). 

Caudalejeunea africana (Steph.) Steph. BIO(2,33), ETH(2), GHA(2), IVO(2), KEN(2), NGA(2), 

SAF(2), TAN(2), ZAI(2). 

Caudalejeunea dusenii Steph. CMN(2), GHA(2), NGA(2), TAN(2), UGA(96). 

Caudalejeunea fruticosa (Lindenb. et Gottsche) Steph. 1912 = Dendrolejeunea fruticosa (Lindenb. et Gottsche) 

Lacout. 1908 [Dubia et Excludenda]. 

Caudalejeunea grolleana Gradst. MDG(1). 

Caudalejeunea hanningtonii (Mitt.) Schiffn. BEN(22), CMN(2), GHA(2), GNB(2), GUI(2), IVO(2,53), 

KEN(2), MDG(1), MLW(563), NGA(2), PRI(48), RIM(10), SAF(2), SAO(48), SIE(2), TAN(2), 

UGA(50), ZAI(2), ZIM(2). 

Caudalejeunea katangensis Vanden Berghen ZAI(2). 

Caudalejeunea lehmanniana (Gottsche) A.Evans BEN(22), CMN(2), GNB(2), NGA(2), SIE(2). 

Caudalejeunea lessonii Steph. 1895 = C. recurvistipula (Gottsche) Schiffn. 1893. 

Caudalejeunea lewallei Vanden Berghen BUR(2), ETH(2), MDG(1), MLW(563), RWA(2), TAN(2), 

UGA(50,96), ZAI(2), ZIM(2). 

Caudalejeunea madagassa Steph. 1912 = C. recurvistipula (Gottsche) Schiffn. 1893. 

Caudalejeunea recurvistipula (Gottsche) Schiffn. COM(1), MDG(1). 

Caudalejeunea reniloba (Gottsche) Steph. 1912 [Dubia et Excludenda]. 

Caudalejeunea stephanii Spruce ex Steph. 1912 = C. reniloba (Gottsche) Steph. 1912 [Dubia et Excludenda]. 

Caudalejeunea tricariniata E.W.Jones 1953 = C. lehmanniana (Gottsche) A.Evans 1907.


Caudalejeunea yangambiensis (Vanden Berghen) E.W.Jones CMN(2), CON(2), GHA(23), RWA(2), 

UGA(2,42), ZAI(2). 

Caudalejeunea zenkeri Steph. 1912 = C. hanningtonii (Mitt.) Schiffn. 1893. 

Cephalojonesia incuba Grolle et Vanden Berghen KEN(2), NGA(2), ZAI(2). 

Cephalozia "atro-virens" Sim 1926, nom. inval. - a typographical error for C. atroviridis Sim 1926. 

Cephalozia africana Váòa RWA(2), ZAI(2). 

Cephalozia atro-viridis Sim 1926 = Cylindrocolea atroviridis (Sim) Váòa 1979. 

Cephalozia bicuspidata (L.) Dumort. KEN(2), MLW(2,114), REU(1), RWA(2), SAF(2), TAN(2), 

UGA(2,42), ZAI(2). 

Cephalozia borbonica (Steph.) Besch. 1895 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Cephalozia caffra J.B.Jack ex Bonner 1963, nom. nud. = C. bicuspidata (L.) Dumort. 1835. 

Cephalozia capensis Sim 1926 = Cephaloziella capensis (Sim) S.W.Arnell 1952. 

Cephalozia cavifolia S.W.Arnell 1956 = C. bicuspidata (L.) Dumort. 1835. 

Cephalozia connivens ssp. fissa Váòa BIO(2,33), CMN(2), CON(2), GAB(2), GHA(2), MDG(1,4), 

MLW(114), NGA(2), PRI(2), RIM(10), SAF(2), SAO(2), TAN(2), TOG(2), UGA(2,96), ZAI(2), 

ZAM(2), ZIM(2). 

Cephalozia connivens var. flagellifera Pearson 1886 = C. connivens ssp. fissa Steph. 1891. 

Cephalozia crassicaulis Steph. 1908 = C. connivens ssp. fissa Steph. 1891. 

Cephalozia denudata var. africana Pearson 1887 = Odontoschisma africanum (Pearson) Sim 1926. 

Cephalozia grossitexta Steph. 1908 = C. connivens ssp. fissa Steph. 1891. 

Cephalozia heteromorpha (Lehm.) Trevis. 1877 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969. 

Cephalozia jishibae Steph. 1924 = Iwatsukia jishibae (Steph.) N.Kitag. 1965. 

Cephalozia kiaeri (Austin) Pearson 1887 = Cephaloziella kiaeri (Austin) Douin 1920. 

Cephalozia ligulata (Steph.) Besch. 1895 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Cephalozia lycopodioides Sim 1926 = Cephaloziella lycopodioides (Sim) S.W.Arnell 1952. 

Cephalozia minutissima Kiaer et Pearson 1893 = Cephaloziella kiaeri (Austin) Douin 1920. 

Cephalozia natalensis Sim 1926 = Cephaloziella natalensis (Sim) S.W.Arnell 1952. 

Cephalozia nematodes Gottsche ex Austin 1879 = Telaranea nematodes (Gottsche ex Austin) M.Howe 1902. 

Cephalozia pillansii Sim 1926 = Alobiellopsis pillansii (Sim) R.M.Schust. 1969. 

Cephalozia radicans Sim 1926 = Cephaloziella kiaeri (Austin) Douin 1920. 

Cephalozia robusta Steph. 1908 = C. bicuspidata (L.) Dumort. 1835. 

Cephalozia tenuissima (Lehm. et Lindenb.) Trevis. 1877 = Cephaloziella tenuissima (Lehm. et Lindenb.) Steph. 

1893. 

Cephalozia vaginans (Steph.) Steph. 1924 = Cephaloziella vaginans Steph. 1911. 

Cephalozia vallis-gratiae Gottsche ex Steph. 1908 = C. bicuspidata (L.) Dumort. 1835. 

Cephalozia villaumei Steph. 1924 = Cephaloziella sp. 

Cephalozia vulcanicola Steph. 1911 = C. bicuspidata (L.) Dumort. 1835. 

Cephalozia welwitschii Steph. 1908 = Cephaloziella welwitschii (Steph.) Douin 1920. 

Cephaloziella abyssinica Gola ETH(2). 

Cephaloziella anthelioides S.W.Arnell SAF(2). 

Cephaloziella "atro-virens" (Sim) S.W.Arnell 1955, nom. inval. = Cylindrocolea atroviridis (Sim) Váòa 1979. 

Cephaloziella atroviridis (Sim) E.W.Jones 1958 = Cylindrocolea atroviridis (Sim) Váòa 1979. 

Cephaloziella capensis (Sim) S.W.Arnell SAF(2). 

Cephaloziella chevalieri (Steph.) Grolle 1964 = Cylindrocolea chevalieri (Steph.) R.M.Schust. 1969. 

Cephaloziella garsidei S.W.Arnell SAF(2), UGA(50). 

Cephaloziella gittinsii E.W.Jones 1960 = Cylindrocolea gittinsii (E.W.Jones) R.M.Schust. 1972. 

Cephaloziella kiaeri (Austin) Douin CMN(2), LES(17), MDG(1), MLW(2,114), MOZ(2), REU(1), 

SAF(2,75), SEY(1), TAN(2), ZAI(2). 

Cephaloziella leonensis S.W.Arnell 1952 = C. capensis (Sim) S.W.Arnell 1952. 

Cephaloziella lycopodioides (Sim) S.W.Arnell SAF(2). 

Cephaloziella natalensis (Sim) S.W.Arnell SAF(2). 

Cephaloziella nigerica E.W.Jones 1960 = Cylindrocolea nigerica (E.W.Jones) R.M.Schust. 1972. 

Cephaloziella pentagona Schiffn. ex Douin 1920 = C. kiaeri (Austin) Douin 1920. 

Cephaloziella pillansii (Sim) S.W.Arnell 1955 = Alobiellopsis pillansii (Sim) R.M.Schust. 1969. 

Cephaloziella radicans Sim 1926 = C. kiaeri (Austin) Douin 1920. 

Cephaloziella retusa E.W.Jones 1960 = Cylindrocolea chevalieri (E.W.Jones) R.M.Schust. 1969.


Cephaloziella schelpei S.W.Arnell SAF(2). 

Cephaloziella tabularis S.W.Arnell SAF(2). 

Cephaloziella tenuissima (Lehm. et Lindenb.) Steph. LES(75), SAF(2,75). 

Cephaloziella transvaalensis S.W.Arnell MLW(114), SAF(2), TAN(659), UGA(50), ZIM(143). 

Cephaloziella triplicata S.W.Arnell SAF(2). 

Cephaloziella ugandica E.W.Jones 1960 = Cylindrocolea ugandica (E.W.Jones) R.M.Schust. 1972. 

Cephaloziella umtaliensis S.W.Arnell MLW(114), MOZ(2), ZIM(2). 

Cephaloziella vaginans Steph. LES(75), MLW(114), REU(45), RWA(2), SIE(2), TAN(2), ZAI(2). Listed 

in error from Cameroun in (2). 

Cephaloziella vaginans var. camerunensis S.W.Arnell CMN(2). 

Cephaloziella welwitschii (Steph.) Douin ANG(2). 

Cephaloziella sp. (Cephalozia villaumei Steph.) MDG(1). 

Ceratolejeunea aberrans Steph. 1923 = Cladolejeunea aberrans (Steph.) Zwickel 1933. 

Ceratolejeunea belangeriana (Gottsche) Steph. COM(1), MAU(1), MDG(1), REU(1), SEY(1). Listed in 

error from Tanzania in (2). 

Ceratolejeunea beninensis E.W.Jones et Vanden Berghen BEN(2), CMN(2), GHA(2), IVO(53), 

NGA(2). 

Ceratolejeunea boschiana Steph. 1913 = C. belangeriana (Gottsche) Steph. 1913. 

Ceratolejeunea calabariensis Steph. ANN(2,33), BIO(2,33), CMN(2), COM(4), CON(2), GAB(2), 

GHA(2), IVO(2,53), MDG(4), MLW(563), NGA(2), PRI(2), RIM(10), SEY(4,108), SIE(2), TAN(2), 

UGA(42,96), ZAI(2). 

Ceratolejeunea cornuta (Lindenb.) Steph. 1895 [Dubia et Excludenda]. 

Ceratolejeunea cornutissima Steph. 1892 = C. diversicornua Steph. 1891 

Ceratolejeunea diversicornua Steph. CMN(2), GHA(2), ZAI(2). 

Ceratolejeunea floribunda Steph. SAO(2). 

Ceratolejeunea jungneri Steph. 1895 = C. calabariensis Steph. 1895 (27). 

Ceratolejeunea mascarena (Steph.) Steph. 1913 = C. belangeriana (Gottsche) Steph. 1913. 

Ceratolejeunea mauritiana (Steph.) Steph. 1892 = C. belangeriana (Gottsche) Steph. 1913. 

Ceratolejeunea papuliflora Steph. MDG(1). 

Ceratolejeunea renauldii Steph. 1913 = Lejeunea alata Gottsche 1845. 

Ceratolejeunea saxbyi Pearson 1921 = C. calabariensis Steph. 1895. 

Ceratolejeunea stictophylla Herzog ex Vanden Berghen NGA(2). 

Ceratolejeunea umbonata Steph. BIO(2,33), CMN(2). 

Ceratolejeunea usambarensis Steph. 1913 = C. jungneri Steph. 1913. 

Ceratolejeunea variabilis (Lindenb.) Schiffn. MAU(1), MDG(1), REU(1). 

Ceratolejeunea zenkeri Steph. BIO(2,33), CMN(2), GAB(2), GHA(2), IVO(53), NGA(2), SIE(2), 

TAN(2), ZAI(2). 

Chandonanthus birmensis Steph. MDG(1), MLW(114). 

Chandonanthus cavallii (Gola) S.W.Arnell 1956 = Tetralophozia cavallii (Gola) Váòa 1993. 

Chandonanthus giganteus Steph. 1911 = C. hirtellus ssp. giganteus (Steph.) Vanden Berghen 1965. 

Chandonanthus hirtellus (F.Weber) Mitt. BIO(36), CMN(2), COM(1), ETH(2), KEN(2,71), MAU(1), 

MDG(1), MLW(2,114), REU(1,4), RWA(2), SAF(2), SAO(2), SEY(1), SIE(2), TAN(2), 

UGA(2,42,50,96), ZAI(2), ZIM(2). 

Chandonanthus hirtellus ssp. giganteus (Steph.) Vanden Berghen ETH(2), KEN(2), RWA(2), TAN(2), 

UGA(2,96), ZAI(2). 

Chandonanthus perloi Gola KEN(2). 

Chandonanthus quadrifidus Steph. 1911 = Tetralophozia cavallii (Gola) Váòa 1993. 

Chandonanthus setiformis sensu Vanden Berghen 1965 [non (Ehrh.) Lindb.] = Tetralophozia cavallii (Gola) 

Váòa 1993. 

Cheilolejeunea africana (Steph.) E.W.Jones 1954 = C. surrepens (Mitt.) E.W.Jones 1976. 

Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle ASC(2), STH(2). 

Cheilolejeunea brachytoma (Gottsche) R.M.Schust. 1980 = C. trifaria (Reinw. et al.) Mizut. 1964. 

Cheilolejeunea brevifissa (Gottsche) R.M.Schust. 1963 = C. krakakammae (Lindenb.) R.M.Schust. 1963. 

Cheilolejeunea camerunensis S.W.Arnell CMN(2), NGA(2).


Cheilolejeunea compressa (Herzog) Grolle COM(1). 

Cheilolejeunea convexa (S.W.Arnell) S.W.Arnell SAF(2). 

Cheilolejeunea cordigera (Steph.) Grolle MAU(1), MDG(1). 

Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones BIO(2,33), CMN(2), ETH(2), KEN(2), 

MDG(1), MLW(563), REU(1), TAN(2), UGA(2). 

Cheilolejeunea crenulata Steph. in Paris 1902, nom. inval. = C. serpentina (Mitt.) Mizut. 1963. 

Cheilolejeunea cucullata Steph. 1914 = Haplolejeunea cucullata (Steph.) Grolle 1979. 

Cheilolejeunea decurrens (Steph.) X.-L.He 1995 = C. surrepens (Mitt.) E.W.Jones 1976. 

Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. CAF(2), CMN(2), COM(1), GUI(2), MDG(1), 

MLW(2), REU(1), SIE(2), TAN(2), ZAI(2). 

Cheilolejeunea diversifolia Augier CMN(2). 

Cheilolejeunea ecarinata Vanden Berghen REU(1,4). 

Cheilolejeunea exigua Steph. 1914 = Lejeunea caespitosa Lindenb. 1845. 

Cheilolejeunea exinnovata E.W.Jones GHA(2), SIE(2), ZIM(2). 

Cheilolejeunea inflata Steph. 1914 = C. intertexta (Lindenb.) Steph. 1897. 

Cheilolejeunea intertexta (Lindenb.) Steph. BEN(22), CMN(2), GHA(2), IVO(2), MDG(1), NGA(2), 

REU(1), SAF(2), SIE(2), TAN(2), UGA(42,50). 

Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. BUR(2), COM(1), ETH(2), KEN(2), MDG(1), 

MLW(563), REU(1), RWA(2), SAF(2,75), TAN(2), UGA(2,42), ZAI(2), ZIM(2). 

Cheilolejeunea kurzii Steph. 1890 = C. intertexta (Lindenb.) Steph. 1897. 

Cheilolejeunea latiflora Steph. CAF(2). 

Cheilolejeunea latistipula Steph. 1913 = Lejeunea eckloniana Lindenb. 1845. 

Cheilolejeunea laurentii Steph. ZAI(2). 

Cheilolejeunea longiflora (Taylor) R.M.Schust. SAO(2). 

Cheilolejeunea madagassa Steph. 1914 = C. serpentina (Mitt.) Mizut. 1963. 

Cheilolejeunea mariana (Gottsche) B.Thiers et Gradst. 1989 [Dubia et Excludenda]. 

Cheilolejeunea montagnei (Gottsche) R.M.Schust. COM(1), KEN(2), MAU(1), MDG(1), REU(1,4), 

RWA(2), SAO(2), TAN(2), ZAI(2). 

Cheilolejeunea newtonii Steph. ex Schiffn. CAF(2), CON(2), ETH(2), GHA(2), GUI(2), KEN(2), 

NGA(48), PRI(2), SIE(2). 

Cheilolejeunea obtusistipula Steph. 1914 = Lejeunea obtusistipula (Steph.) E.W.Jones 1989. 

Cheilolejeunea omphalogastria Pócs RWA(2). 

Cheilolejeunea pluriplicata (Pearson) R.M.Schust. ANG(2), ETH(2), KEN(2,71), MDG(1), MLW(563), 

MOZ(2), SAF(2), TAN(2), UGA(2,96), ZAI(2). 

Cheilolejeunea pocsii E.W.Jones KEN(21), MDG(1), MLW(563), RWA(2), SAF(2), TAN(2), 

UGA(115). 

Cheilolejeunea principensis Steph. ex Paris 1888, nom. inval. = C. serpentina (Mitt.) Mizut. 1963. 

Cheilolejeunea principensis Steph. ex Paris 1906 = C. serpentina (Mitt.) Mizut. 1963. 

Cheilolejeunea robillardii (Steph.) R.M.Schust. 1980 = C. trifaria (Reinw. et al.) Mizut. 1964. 

Cheilolejeunea rufescens (Lindenb.) Grolle SAF(2). 

Cheilolejeunea ruwenzorensis (S.W.Arnell) R.M.Schust. UGA(2), ZAI(2). 

Cheilolejeunea serpentina (Mitt.) Mizut. ANN(2,33), BIO(2,33), CMN(2), COM(1,4), GHA(2), GUI(2), 

IVO(2), KEN(48), MAU(1,108), MDG(1,4), NGA(2), PRI(2), REU(1), RIM(10), SEY(1,4,13,108), 

SIE(2), TAN(2), ZAI(2). 

Cheilolejeunea silvestris (Gottsche) E.W.Jones 1974 = C. surrepens (Mitt.) E.W.Jones 1976. 

Cheilolejeunea silvestris var. involuta (Steph.) E.W.Jones 1974 = C. surrepens (Mitt.) E.W.Jones 1976. 

Cheilolejeunea surrepens (Mitt.) E.W.Jones ANN(2,33), CMN(2), COM(1,4), GHA(2), IVO(2,53), 

KEN(2), MAU(1), MDG(1), MLW(563), NGA(2), REU(1), ROD(1), SEY(1,4), SIE(2), TAN(2), 

UGA(2,42,96), ZAI(2). 

Cheilolejeunea tisserantii Ast et Vanden Berghen 1951 = C. decursiva (Sande Lac.) R.M.Schust. 1963. 

Cheilolejeunea trapezia (Nees) Kachroo et R.M.Schust. ZAI(2). 

Cheilolejeunea trifaria (Reinw. et al.) Mizut. BIO(33), CAF(2), CMN(2), GAB(2), GHA(2), MAU(1), 

MDG(1), REU(1), RWA(2), SAO(2), SEY(1,4,108), SIE(2), TAN(2), TOG(48), UGA(2,96), ZAI(2),


ZIM(2). 

Cheilolejeunea usambarana (Steph.) Grolle CMN(2), CON(2), GUI(2), MLW(563), REU(1), SIE(2), 

TAN(2). 

Cheilolejeunea ventricosa (Schiffn.) X.-L.He MAU(108). 

Cheilolejeunea sp. (Strepsilejeunea vatovae Gerola) ETH(2). 

Chiloscyphus ankefinensis Gottsche ex Steph. 1907 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Chiloscyphus argutus (Reinw. et al.) Nees 1845 = Heteroscyphus argutus (Reinw. et al.) Schiffn. 1910 [Dubia et 

Excludenda]. 

Chiloscyphus armatistipulus Steph. 1893 = Heteroscyphus dubius (Gottsche) Schiffn. 1910. 

Chiloscyphus coadunatus (Sw.) J.J.Engel et R.M.Schust. 1984 = Lophocolea bidentata (L.) Dumort. 1835. 

Chiloscyphus concretus (Mont.) J.J.Engel et R.M.Schust. 1985 = Lophocolea concreta Mont. 1845. 

Chiloscyphus congoanus (Steph.) J.J.Engel et R.M.Schust. 1985 = Lophocolea martiana Nees 1845. 

Chiloscyphus connatus (Sw.) J.J.Engel et R.M.Schust. 1984 = Lophocolea connata (Sw.) Mont. et Nees 1839 


Chiloscyphus cuspidatus (Nees) J.J.Engel et R.M.Schust. 1985 = Lophocolea bidentata (L.) Dumort. 1835. 

Chiloscyphus decurrens Nees 1845 = Heteroscyphus splendens (Lehm. et Lindenb.) Grolle 1984. 

Chiloscyphus difformis (Nees) J.J.Engel et R.M.Schust. 1985 = Lophocolea difformis Nees 1845. 

Chiloscyphus dubius Gottsche 1882 = Heteroscyphus dubius (Gottsche) Schiffn. 1910. 

Chiloscyphus expansus (Lehm.) Nees 1845 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Chiloscyphus fasciculatus (Nees) Gottsche et al. 1845 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Chiloscyphus fragrans (Moris et De Not.) J.J.Engel et R.M.Schust. 1985 = Lophocolea fragrans (Moris et De 

Not.) Gottsche et al. 1845. 

Chiloscyphus grandistipus Steph. 1890 = Heteroscyphus grandistipus (Steph.) Schiffn. 1910. 

Chiloscyphus hamatistipulus Steph. 1907 = Heteroscyphus dubius (Gottsche) Schiffn. 1910. 

Chiloscyphus lepervanchei (Steph.) J.J.Engel et R.M.Schust. 1984 = Lophocolea lepervanchei Steph. 1907. 

Chiloscyphus lindenbergianus ("lindenbergii") Nees 1845 = Lophocolea semiteres (Lehm.) Mitt. 1877. 

Chiloscyphus loangensis Steph. 1893 = Heteroscyphus dubius (Gottsche) Schiffn. 1910. 

Chiloscyphus lucidus (Spreng. ex Lehm.) J.J.Engel et R.M.Schust. 1985 [non Mitt. 1876] = Lophocolea lucida 

(Spreng. ex Lehm.) Mont. 1839. 

Chiloscyphus madagascariensis (Gottsche) J.J.Engel et R.M.Schust. 1984 = Lophocolea madagascariensis 

Gottsche 1882. 

Chiloscyphus martianus (Nees) J.J.Engel et R.M.Schust. 1985 = Lophocolea martiana Nees 1845. 

Chiloscyphus mascarenensis S.W.Arnell 1965 = Heteroscyphus splendens (Lehm. et Lindenb.) Grolle 1984. 

Chiloscyphus minor (Nees) J.J.Engel et R.M.Schust. 1984 = Lophocolea minor Nees 1836. 

Chiloscyphus mollis (Nees) J.J.Engel et R.M.Schust. 1984 = Lophocolea mollis (Nees) Gottsche et al. 1845. 

Chiloscyphus muhavurensis S.W.Arnell 1956 = Lophocolea muhavurensis (S.W.Arnell) S.W.Arnell ex Pócs 

1979. 

Chiloscyphus muricatus (Lehm.) J.J.Engel et R.M.Schust. 1985 = Lophocolea muricata (Lehm.) Nees 1845. 

Chiloscyphus oblongifolius Mitt. 1860, hom. illeg. [non (Hook.f. et Taylor) Gottsche et al. 1847] = 

Heteroscyphus dubius (Gottsche) Schiffn. 1910. 

Chiloscyphus occidentalis (Steph.) J.J.Engel et R.M.Schust. 1985 = Heteroscyphus dubius (Gottsche) Schiffn. 

1910. 

Chiloscyphus rabenhorstii Steph. 1906 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Chiloscyphus regularis Steph. 1893 [non Steph. 1906] = Lophocolea onraedtii Grolle 1984. 

Chiloscyphus renauldii Steph. 1907 = Heteroscyphus cf. dubius (Gottsche) Schiffn. 1910. 

Chiloscyphus semiteres (Lehm.) Lindenb. 1845 = Lophocolea semiteres (Lehm.) Mitt. 1877. 

Chiloscyphus silvestris (Gottsche) J.J.Engel et R.M.Schust. 1984 = Lophocolea silvestris Gottsche 1882. 

Chiloscyphus spectabilis Steph. 1891 = Heteroscyphus spectabilis (Steph.) Schiffn. 1910. 

Chiloscyphus thomeensis Steph. 1893, perhaps = Heteroscyphus spectabilis (Steph.) Schiffn. 1910. 

Chiloscyphus trapezioides Sande Lac. 1855 = Conoscyphus trapezioides (Sande Lac.) Schiffn. 1898. 

Chonecolea ruwenzorensis E.W.Jones UGA(2,50,96). 

Ciliolejeunea capensis S.W.Arnell 1953 = Lejeunea villaumei (Steph.) Grolle 1977. 

Cincinnulus bidentulus (F.Weber) Steph. 1895 = Calypogeia bidentula (F.Weber) Nees 1845. 

Cladolejeunea aberrans (Steph.) Zwickel TAN(2). 

Clasmatocolea fasciculata (Nees) Grolle SAF(2,97). 

Clasmatocolea vermicularis (Lehm.) Grolle BUR(2,97), CMN(2), ETH(2), LES(2,75), MDG(97),


MLW(2,29), NGA(2), REU(1,97), RWA(2), SAF(2,97), TAN(2,97), UGA(115), ZAI(2,97), ZIM(2,97). 

Clevea pulcherrima Steph. 1895 = Athalamia pulcherrima (Steph.) S.Hatt. 1954. 

Clevea spathysii (Lindenb.) Müll.Frib. 1940 = Athalamia spathysii (Lindenb.) S.Hatt. 1954. 

Cololejeunea adhaesiva (Mitt.) R.M.Schust. TAN(2). 

Cololejeunea adnata Tixier MDG(1). 

Cololejeunea africana (Steph.) R.M.Schust. CMN(2), KEN(2), SAO(2), SIE(2), TAN(2). 

Cololejeunea amaniensis Pócs TAN(2). 

Cololejeunea andapania Tixier MDG(1). 

Cololejeunea androphylla E.W.Jones CMN(2), CON(2), GHA(2), IVO(2), NGA(2), RIM(10), SEY(1), 

SIE(2). 

Cololejeunea androphylla var. madecassa Tixier MDG(1). 

Cololejeunea ankaiana Tixier MDG(1). 

Cololejeunea apiculata (E.W.Jones) R.M.Schust. CMN(2), TAN(2), UGA(50). 

Cololejeunea appressa (A.Evans) Benedix COM(1), IVO(2), KEN(2), MDG(1), MLW(563), REU(1), 

SEY(1), SIE(2), TAN(2), UGA(2), ZAI(2). 

Cololejeunea augieri Tixier RWA(2). 

Cololejeunea auriculata (E.W.Jones) R.M.Schust. CMN(2), KEN(2), MDG(1), NGA(2), SEY(1), 

UGA(42). 

Cololejeunea autoica (Steph.) Grolle COM(1). 

Cololejeunea bandamiae Tixier COM(1). 

Cololejeunea bebourensis Tixier COM(1), REU(1). 

Cololejeunea bichiana Tixier 1974 = C. platyneura (Spruce) A.Evans 1902. 

Cololejeunea bidentula (Steph.) E.W.Jones KEN(2), MAU(1), MDG(1), REU(1), TAN(2), UGA(42). 

Cololejeunea bolombensis (Steph.) Vanden Berghen 1972 = C. tonkinensis Steph. 1895. 

Cololejeunea borbonica Tixier MDG(1). 

Cololejeunea borhidiana Pócs TAN(2). 

Cololejeunea bosseriana Tixier REU(1). 

Cololejeunea calcarata E.W.Jones GHA(2). 

Cololejeunea camusii Tixier MDG(1). 

Cololejeunea capensis S.W.Arnell 1953 = Aphanolejeunea capensis (S.W.Arnell) S.W.Arnell 1957. 

Cololejeunea capuronii Tixier MDG(1). 

Cololejeunea cardiocarpa (Mont.) A.Evans ASC(2), BUR(2), CMN(2), KEN(2,665), MDG(1), 

MLW(563), SAF(2), SIE(2), SWZ(2), TAN(2), UGA(2), ZAI(2), ZIM(2). 

Cololejeunea cardiocarpioides Tixier RWA(2). 

Cololejeunea ceatocarpa (Ångstr.) Steph. REU(1). 

Cololejeunea cornuta E.W.Jones GHA(2), NGA(2), RIM(10), SIE(2). 

Cololejeunea crenatiflora (Steph.) Vanden Berghen 1972 = C. obliqua (Nees et Mont.) Schiffn. 1897. 

Cololejeunea crenatiflora Steph. 1891, nom. inval. = C. obliqua (Nees et Mont.) Schiffn. 1897. 

Cololejeunea crenatiflora var. orientalis Tixier 1985 = C. obliqua (Nees et Mont.) Schiffn. 1897. 

Cololejeunea cristata (Steph.) R.M.Schust. KEN(2), MAU(1,108), MDG(1), ZAI(2). 

Cololejeunea cuneata (Lehm. et Lindenb.) Herzog COM(1), MAU(1), MDG(1,4), REU(1), ROD(1), 

TAN(2). 

Cololejeunea cuneifolia Steph. CMN(2), GHA(2), GUI(2), IVO(2), NGA(2), RIM(10), SIE(2), TAN(2), 

UGA(2,42,50), ZAI(2). 

Cololejeunea cuspidata Steph. ex Tixier SEY(4). 

Cololejeunea decemplicata (Steph.) Tixier MDG(1), REU(1). 

Cololejeunea dentata (E.W.Jones) R.M.Schust. CMN(2), GHA(2), KEN(2), SIE(2), TAN(2), UGA(50), 

ZAI(2). 

Cololejeunea deroinii Tixier MDG(1). 

Cololejeunea deslooveri Vanden Berghen ZAI(2). 

Cololejeunea diplasiolejeuneoides Tixier MDG(1). 

Cololejeunea disciflora Tixier MDG(1). 

Cololejeunea dissita E.W.Jones 1954 = C. minutissima ssp. myriocarpa (Nees et Mont.) R.M.Schust. 1955.


Cololejeunea distalopapillata (E.W.Jones) R.M.Schust. COM(1), KEN(2), MDG(1), MLW(563), 

RWA(2), TAN(2), UGA(42,50), ZAI(2), ZIM(2). 

Cololejeunea dolichodonta Tixier MDG(1). 

Cololejeunea duvignaudii E.W.Jones COM(1), GAB(2), KEN(2), MDG(1), MLW(563), REU(1), 

RIM(10), RWA(2), TAN(2), UGA(115), ZAI(2), ZIM(2). 

Cololejeunea duvignaudii var. papillata Tixier MDG(1), REU(1). 

Cololejeunea elegans Steph. CMN(2), COM(1), GHA(2), KEN(2), MDG(1), MLW(563), RIM(10), 

TAN(2). 

Cololejeunea erectifolia Steph. 1895 = C. cuneata (Lehm. et Lindenb.) Herzog 1947. 

Cololejeunea fadenii Pócs 1975 = Aphanolejeunea fadenii (Pócs) Pócs 1984. 

Cololejeunea falcata var. madecassa Tixier MDG(1). 

Cololejeunea filicaulis Steph. 1892 = C. hildebrandii (Austin) Steph. 1897. 

Cololejeunea fischeri Tixier RWA(2), ZAI(2). 

Cololejeunea floccosa (Lehm. et Lindenb.) Schiffn. IVO(2), MDG(1), SIE(2). 

Cololejeunea floccosa var. trivittata Tixier MDG(1). 

Cololejeunea frahmii Tixier RWA(2). 

Cololejeunea furcilobulata (Berrie et E.W.Jones) R.M.Schust. MDG(1), NGA(2), SEY(4), TAN(2). 

Cololejeunea grolleana Pócs TAN(2). 

Cololejeunea grossidens (Steph.) E.W.Jones 1953 = C. malanjae Steph. 1902. 

Cololejeunea grossidens Steph. 1900, nom. nud. = C. malanjae Steph. 1902. 

Cololejeunea guehoi Tixier MAU(1). 

Cololejeunea harrisii Pócs BUR(2), TAN(2). 

Cololejeunea harrisii sensu Vanden Berghen 1977 = C. magna (Tixier) Infante et Heras 1999 (39). 

Cololejeunea harrisii var. magna Tixier 1995 = C. magna (Tixier) Infante et Heras 1999 (39). 

Cololejeunea haskarliana (Lehm. et Lindenb.) Schiffn. REU(1), SEY(1). 

Cololejeunea heterolobula Tixier RWA(2), ZAI(2). 

Cololejeunea hildebrandii (Austin) Steph. CMN(2), COM(1), ETH(2), KEN(2), MDG(1), REU(1), 

RWA(2), TAN(2), ZAI(2). 

Cololejeunea himalayensis (Pandé et Misra) R.M.Schust. 1963 = C. latilobula (Herzog) Tixier 1985. 

Cololejeunea hyalinomarginata (Nees ex Mont.) Grolle 1981 = C. marginata (Lehm. et Lindenb.) Schiffn. 1898. 

Cololejeunea inflectens (Mitt.) Benedix COM(1), SEY(1,4,108). 

Cololejeunea iradieri Infante et Heras RIM(39). 

Cololejeunea jonesii Pócs TAN(2). 

Cololejeunea kahuziensis Tixier ZAI(2). 

Cololejeunea lanceolata E.W.Jones CMN(2), GHA(2), NGA(2), SIE(2). 

Cololejeunea latilobula (Herzog) Tixier BUR(2), COM(4), MLW(2,563), NGA(2), RIM(10), SEY(1), 

TAN(2), UGA(2,42), ZAI(2), ZAM(2). 

Cololejeunea leloutrei (E.W.Jones) R.M.Schust. CMN(2), COM(1), GAB(2), GHA(25), KEN(2), 

MAU(1), MDG(1), RIM(10), SAO(2), SEY(1,4,108), SIE(25), TAN(2,108), ZAI(2). Some current 

opinion favours the view that all African mainland records are probably of C. leloutrei, whilst C. marginata is 

endemic to the East African islands. Pending a clarification of this difficult complex, records from Ghana and 

Sierra Leone, uncertainly assigned to C. marginata in (2), have been re-assigned in this checklist to C. leloutrei. 

The record from Tanzania is also transferred. 

Cololejeunea leloutrei var. microlobulata Tixier MDG(1). 

Cololejeunea leloutrei var. ulugurica Pócs ex Tixier CMN(2), COM(1), KEN(2), MDG(1), TAN(2). 

Cololejeunea lemuriana Tixier MDG(1), TAN(108,129). 

Cololejeunea leonardii Vanden Berghen 1971 = C. runssorensis (Steph.) Pócs 1975. 

Cololejeunea lobulilineata Tixier ZAI(2). 

Cololejeunea magillii Pócs COM(1). 

Cololejeunea magna (Tixier) Infante et Heras RIM(39), RWA(2). 

Cololejeunea malanjae Steph. BUR(2), KEN(2), MLW(2,563), RWA(2), TAN(2), UGA(115), ZAI(2), 

ZIM(2).


Cololejeunea marginata (Lehm. et Lindenb.) Schiffn. COM(1), MAU(1,4,108), MDG(1), REU(1), 

SEY(4,108). See note under C. leloutrei. 

Cololejeunea mariana Tixier 1979 = C. obliqua (Nees et Mont.) Schiffn. 1897. 

Cololejeunea minutissima (Sm.) Schiffn. BEN(22), CVI(2,37), IVO(53), MDG(1), MLW(2,563), 

REU(1), ROD(1), RWA(2), SAF(2,75), TAN(2), UGA(42,50,115), ZAI(2), ZIM(2). 

Cololejeunea minutissima ssp. myriocarpa (Nees et Mont.) R.M.Schust. GHA(2), LES(75), 

MLW(2,563), NGA(2), SAF(2), TAN(2), UGA(42,115). 

Cololejeunea minutissima ssp. utriculifera Vanden Berghen BUR(2), RWA(2), TAN(2). 

Cololejeunea minutissima ssp. myriantha (Herzog) R.M.Schust. 1955 = C. minutissima ssp. myriocarpa (Nees et 

Mont.) R.M.Schust. 1955. 

Cololejeunea mocambiquensis S.W.Arnell MDG(1), MLW(563), MOZ(2), SAF(2), TAN(2), UGA(2). 

Cololejeunea moramangae Tixier 1979 = Aphanolejeunea moramangae (Tixier) Pócs 1984. 

Cololejeunea myriantha (Herzog) S.W.Arnell 1953 = C. minutissima ssp. myriocarpa (Nees et Mont.) 


Cololejeunea myriocarpa (Nees et Mont.) C.Massal. 1906 = C. minutissima ssp. myriocarpa (Nees et Mont.) 


Cololejeunea nigerica (E.W.Jones) R.M.Schust. CMN(2), GHA(2), IVO(2), NGA(2), SIE(2), 

TOG(2,101), UGA(42,115), ZAI(2). 

Cololejeunea nymanii (Steph.) Benedix 1953 = C. obliqua (Nees et Mont.) Schiffn. 1897. 

Cololejeunea obliqua (Nees et Mont.) Schiffn. BUR(2), CMN(2), COM(1), GHA(2), GUI(2), IVO(2), 

MAU(1), MDG(1), NGA(2), REU(1), RIM(10), SAO(2), SEY(1,4,108), SIE(2), TAN(2), 

UGA(2,42,115), ZAI(2), ZAM(2). 

Cololejeunea obtusifolia (E.W.Jones) Tixier BEN(22), BUR(2), CMN(2), ETH(2), GHA(2), GNB(2), 

GUI(2), IVO(2), KEN(2), MDG(1), MLW(563), NGA(2), RIM(10), RWA(2), SIE(2), TAN(2), 

UGA(42,50), ZAI(2). 

Cololejeunea occidentalis (E.W.Jones) Vanden Berghen KEN(2), MLW(563), NGA(2), SAF(2), 

TAN(2), ZIM(2). 

Cololejeunea oleana Sim SAF(2). 

Cololejeunea onraedtii Tixier COM(1), SEY(1). 

Cololejeunea parva Vanden Berghen RWA(2), UGA(115). 

Cololejeunea peculiaris (Herzog) Benedix 1953 = C. inflectens (Mitt.) Benedix 1953. 

Cololejeunea peponiformis Mizut. COM(1), REU(1), SEY(1). 

Cololejeunea peraffinis var. elegans Benedix GUI(2,106), NGA(2,104,106), SIE(2,25). The plants from 

Guinea, Nigeria and Sierra Leone, were published by E.W. Jones in (25, 104, 106) as C. peraffinis var. elegans. 

However in a later (unpublished) manuscript, he acknowledged that they showed conflicting characters, 

approaching C. peraffinis var. elegans in some, and matching C. usambarica (= C. platyneura) in others. At that 

later time he placed the collections under C. usambarica pending further investigations. (see also Pócs, in Pócs & 

Streimann 1999 (103)). 

Cololejeunea plagiochiliana Tixier MDG(1). 

Cololejeunea planissima (Mitt.) Abeyw. TAN(702). 

Cololejeunea platyneura (Spruce) A.Evans MDG(1), MLW(563), RWA(2), SIE(2), TAN(2), ZAI(2). 

Cololejeunea praeruptorum Tixier MDG(1). 

Cololejeunea pseudo-obliqua Tixier RWA(2), ZAI(2). 

Cololejeunea pseudopusilla Tixier ZAI(2). 

Cololejeunea pteroporum Tixier MDG(1). 

Cololejeunea punctata (E.W.Jones) R.M.Schust. 1963, nom. illeg. = C. vidaliana Tixier 1973. 

Cololejeunea punctata (Gottsche) Pearson 1892 [Dubia et Excludenda]. 

Cololejeunea pusilla Steph. CMN(2), GAB(2), GHA(2), GUI(2), NGA(2), RIM(10), RWA(2), 

TAN(108), TOG(101). 

Cololejeunea pusilla var. obtusifolia E.W.Jones 1954 = C. obtusifolia (E.W.Jones) Tixier 1995. 

Cololejeunea raduliloba Steph. COM(1), MDG(1), SEY(1). 

Cololejeunea runssorensis (Steph.) Pócs BUR(2), KEN(2), MLW(563), RWA(2), UGA(2,115), ZAI(2). 

Cololejeunea saltuum Tixier MDG(1). 

Cololejeunea sambiroana Tixier MDG(1).


Cololejeunea scabriflora Gottsche ex Steph. 1895 = C. obliqua (Nees et Mont.) Schiffn. 1897. 

Cololejeunea scabrifolia Schiffn. 1900, nom. inval. = C. obliqua (Nees et Mont.) Schiffn. 1897. 

Cololejeunea selangorensis Tixier MDG(1). 

Cololejeunea sphaerocarpa Tixier RWA(2), ZAI(2). 

Cololejeunea stenophylla Herzog 1947 = C. hildebrandii (Austin) Steph. 1897. 

Cololejeunea subciliata (Mitt.) R.M.Schust. 1963 = C. cuneata (Lehm. et Lindenb.) Herzog 1947. 

Cololejeunea subdiaphana Ast 1947 = Aphanolejeunea truncatifolia Horik. 1934 (117). 

Cololejeunea subinflata Tixier MDG(1). 

Cololejeunea takamakae Tixier REU(1). 

Cololejeunea tamatavensis Tixier MDG(1). 

Cololejeunea tanneri Pócs MLW(563), TAN(2). 

Cololejeunea tanzaniae Pócs COM(1), MAU(108), MDG(1), MLW(563), TAN(2). 

Cololejeunea tenella Benedix RWA(2). 

Cololejeunea tenuiparietata Tixier RWA(2), ZAI(2). 

Cololejeunea tonkinensis Steph. BUR(2), CMN(2), COM(1), GUI(2), IVO(2), KEN(2), MDG(1), 

MLW(563), NGA(2), RIM(10), SIE(2), TAN(2), UGA(2,42), ZAI(2), ZIM(2). C. tonkinensis and C. 

bolombensis are treated as synonyms of C. lanciloba Steph. in Zhu & So, 2000 (702). 

Cololejeunea tribracteata Tixier ZAI(2). 

Cololejeunea usambarica E.W.Jones 1954 = C. platyneura (Spruce) A.Evans 1902. 

Cololejeunea vidaliana Tixier KEN(2), MAU(1), MDG(1), SEY(1,4,108), TAN(2). 

Cololejeunea xavieri (Lacout. ex Steph.) E.W.Jones COM(1), KEN(2). Made a synonym of C. elegans in 

Tixier 1979 (152), but considered probably distinct by others, and on that basis is kept separate here. 

Cololejeunea zenkeri (Steph.) E.W.Jones BUR(2), CMN(2), MLW(563), PRI(48), RIM(10), RWA(2), 

SEY(4,108), SIE(2), TAN(2), UGA(50), ZAI(2). 

Colura alata (Gottsche) Trevis. 1877 = Lejeunea alata Gottsche 1845. 

Colura benoistii Ast COM(1), MDG(1). 

Colura berghenii Ast RWA(2), TAN(2). 

Colura bicornis Ast MDG(1,4). 

Colura calyptrifolia (Hook.) Dumort. BUR(2), MLW(110), RWA(2), SAF(2,75), TAN(2), UGA(42,50), 

ZAI(2), ZIM(2). 

Colura calyptrifolia ssp. tenuicornis (A.Evans) Vanden Berghen 1972 = C. tenuicornis (A.Evans) Steph. 1916. 

Colura cylindrica Herzog TAN(2). 

Colura digitalis (Mitt.) Steph. ANN(2,33), BUR(2), CAF(2), CMN(2), COM(1,4), CON(2), GAB(2), 

GHA(2), GUI(2), IVO(2), MAU(1,108), MDG(4), MLW(110), NGA(2), PRI(2), REU(1), RIM(10), 

RWA(2), SAO(2), SEY(1,108), SIE(2), TAN(2), UGA(2,42), ZAI(2). 

Colura digitalis var. mucronata Pócs SEY(4). 

Colura dusenii (Steph.) Steph. CMN(2), GHA(2), KEN(2), NGA(2), SIE(2), TAN(2). 

Colura hattoriana Pócs COM(1), MLW(110). 

Colura hedbergiana Pócs TAN(2), UGA(42). 

Colura heimii Ast COM(1), MAU(1,108), MDG(1), REU(1). 

Colura humbertii Ast MAU(1), MDG(1), REU(1). 

Colura inornata Ast MDG(1,4). 

Colura kilimanjarica Pócs et Ast TAN(2), UGA(42). 

Colura mauritiana Pócs MAU(6,108). 

Colura obesa Ast MDG(1), REU(1). 

Colura obtusa (Steph.) Steph. 1916 = C. digitalis (Mitt.) Steph. 1916. 

Colura obvoluta Ast MAU(108), MDG(1), SEY(4,108). 

Colura pluridentata Ast SEY(1). 

Colura pungens Herzog 1947 = C. tenuicornis (A.Evans) Steph. 1916. 

Colura saroltae Pócs RWA(2), TAN(2). 

Colura superba (Mont.) Steph. 1916 [Dubia et Excludenda]. 

Colura tenuicornis (A.Evans) Steph. ASC(2), BUR(2), CMN(2), COM(1), GAB(2), KEN(2), MAU(1), 

MDG(1), MLW(110), REU(1), RIM(10), RWA(2), SAF(2), SAO(2), SEY(1,4), SIE(2), TAN(2),


UGA(42,50), ZAI(2). 

Colura usambarica E.W.Jones MLW(2,110), TAN(2). 

Colura zoophaga E.Fischer KEN(52). 

Colurolejeunea dusenii Steph. 1892 = Colura dusenii (Steph.) Steph. 1916. 

Colurolejeunea obtusa Steph. 1891 = Colura digitalis (Mitt.) Steph. 1916. 

Colurolejeunea tenuicornis A.Evans 1900 = Colura tenuicornis (A.Evans) Steph. 1916. 

Conoscyphus inflexifolius Mitt. 1873 = C. trapezioides (Sande Lac.) Schiffn. 1898. 

Conoscyphus trapezioides (Sande Lac.) Schiffn. MDG(1,4), MLW(29), REU(1,4), SAO(2), SEY(1,4), 

SIE(2), TAN(2). Erroneously listed from Cameroun in (2). 

Crossotolejeunea kilimandjarica S.W.Arnell 1960 = Taxilejeunea pulchriflora Pearson 1924. 

Cryptochila grandiflora (Lindenb. et Gottsche) Grolle MDG(1), MLW(114), REU(1,4), SAF(2). 

Cryptomitrium oreades Perold LES(2,75). 

Cuspidatula contracta (Reinw. et al.) Steph. 1901 = Jamesoniella contracta (Reinw. et al.) N.Kitag. 1981. 

Cyathodium africanum Mitt. 1886 = C. cavernarum Kunze 1834 (119). 

Cyathodium aureonitens (Griff.) Mitt., non Schiffn. CMN(2). 

Cyathodium cavernarum Kunze ANN(2,33), BIO(2,33,36), CAF(2), CMN(2), COM(1), CON(2), 

CVI(2,20,37), GHA(2), GUI(2), IVO(2), MLW(2), NAM(2), NGA(2), SAO(2), SIE(2), TAN(2), 

TOG(2), UGA(42,50,96), ZAI(2), ZIM(2). Jones 1952 (650) concluded that C. cavernarum and the Asian C. 

smaragdinum Schiffn. are conspecific, but Srivastava & Dixit 1996 (119) retained them as distinct, mainly on 

thallus size and spore morphology. They reported the latter species from Africa, citing, on p. 158, a specimen from 

Tanzania (Jones 679). However, on p. 188, they also cite Jones 679 under C. cavernosum. Clarification is 

required. 

Cyathodium foetidissimum Schiffn. CMN(2), UGA(96). The Uganda plants are sterile, and Lye & Pócs (96) 

considered their identity is uncertain. 

Cyathodium griffithii Schiffn. 1938, nom. illeg. = C. aureonitens (Griff.) Mitt. 1886. 

Cyclolejeunea annobonensis S.W.Arnell 1956 = Prionolejeunea grata (Gottsche) Schiffn. 1893. 

Cylindrocolea atroviridis (Sim) Váòa CAF(2), CMN(2), GHA(2), IVO(51), LES(75), MDG(1), MOZ(2), 

NGA(2), RWA(2), SAF(2), SIE(2), UGA(50), ZAI(2), ZIM(2). 

Cylindrocolea chevalieri (Steph.) R.M.Schust. CON(2), GAB(2), GHA(2), NGA(2). 

Cylindrocolea gittinsii (E.W.Jones) R.M.Schust. CMN(2), KEN(21), RWA(2), UGA(2). 

Cylindrocolea madagascariensis (Steph.) R.M.Schust. MDG(1). 

Cylindrocolea nigerica (E.W.Jones) R.M.Schust. DJI(2), NGA(2). 

Cylindrocolea ugandica (E.W.Jones) R.M.Schust. KEN(21), SIE(2), UGA(2,42). 

Cystolejeunea lineata (Lindenb. et Lehm.) A.Evans 1906 [Dubia et Excludenda]. 

Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche et al. STH(2). 

Dendroceros africanus Steph. CMN(2), TAN(2). 

Dendroceros borbonicus Steph. MAU(1), REU(1), SEY(1,108). 

Dendroceros crispatus Nees SAO(95). 

Dendroceros javanicus (Nees) Gottsche et al. TAN(2). 

Dendroceros lacerus Nees 1846 = Folioceros fuciformis (Mont.) Bharadw. 1975. 

Dendrolejeunea fruticosa (Lindenb. et Gottsche) Lacout. [Dubia et Excludenda]. 

Dicranolejeunea africana Steph. 1912 = Caudalejeunea sp. [non C. africana (Steph.) Schiffn. 1893]. 

Dicranolejeunea annobonensis S.W.Arnell 1956 = Marchesinia sp. 

Dicranolejeunea atra (Mitt.) Vanden Berghen 1953 = Acanthocoleus aberrans var. laevis Gradst. 1994. 

Dicranolejeunea bovonei Gola [Dubia et Excludenda] (542). 

Dicranolejeunea chrysophylla (Lehm.) Grolle 1966 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988. 

Dicranolejeunea lillieana Pearson 1924 = Acanthocoleus sp. 

Dicranolejeunea madagascariensis Steph. 1912 = Acanthocoleus madagascariensis (Steph.) Kruijt 1988. 

Dicranolejeunea madagascariensis var. obtusifolia E.W.Jones 1970 = Acanthocoleus chrysophyllus (Lehm.) 

Kruijt 1988. 

Dicranolejeunea nathalensis (Sim) S.W.Arnell 1961 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988. 

Dicranolejeunea nathalensis sensu S.W.Arnell 1963, hom. illeg. [non (Sim) S.W.Arnell 1961] = 

Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst. 1986. 

Dicranolejeunea phyllorhiza (Nees) Schiffn. 1893 = Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst. 1986. 

Dicranolejeunea usambarensis Steph. 1912 = Acanthocoleus aberrans var. laevis Gradst. 1994.


Diplasiolejeunea albifolia (Taylor) E.W.Jones KEN(2), NGA(2), TAN(2). 

Diplasiolejeunea albifolia ssp. orientalis Pócs, ined. TAN(129). 

Diplasiolejeunea aulae E.W.Jones GHA(2), MDG(1), RWA(2), UGA(50). 

Diplasiolejeunea auriculata Tixier MDG(1). 

Diplasiolejeunea bifidula Steph. 1916 = D. cornuta Steph. 1916. 

Diplasiolejeunea brachyclada A.Evans 1912 = D. cavifolia Steph. 1895. 

Diplasiolejeunea cavifolia Steph. COM(1), GHA(2), KEN(2), MAU(1,108), MDG(1), NGA(2), 

RIM(10), RWA(2), SAF(2), SAO(2), TAN(2), UGA(42), ZAI(2). 

Diplasiolejeunea cobrensis Gottsche ex Steph. GHA(2), MDG(1), SIE(2). 

Diplasiolejeunea cobrensis subsp. antsirananae Pócs MDG(709). 

Diplasiolejeunea cogoensis Infante, Heras et Pócs RIM(38). 

Diplasiolejeunea comorensis Tixier COM(1), MAU(108), MDG(1), ZAI(2). 

Diplasiolejeunea cornuta Steph. COM(1), GHA(2), KEN(2), MAU(1), MDG(1,4), MLW(563), 

REU(1,4), RWA(2), SEY(4), TAN(2), UGA(42,115), ZAI(2), ZIM(2). 

Diplasiolejeunea cyangugensis Tixier RWA(2). 

Diplasiolejeunea deslooveri Vanden Berghen RWA(2), TAN(2). 

Diplasiolejeunea drepanolejeunoides Tixier 1977 = D. magnistipulata Tixier 1975. 

Diplasiolejeunea ensifera Tixier MDG(1). 

Diplasiolejeunea gradsteinii Tixier ZAI(2). 

Diplasiolejeunea hamata Tixier MDG(1). 

Diplasiolejeunea harpaphylla Steph. 1916 = D. cobrensis Gottsche ex Steph. 1916. 

Diplasiolejeunea incurvata Ast et Tixier 1962 = D. cobrensis Gottsche ex Steph. 1916. 

Diplasiolejeunea insignis Tixier MDG(1). 

Diplasiolejeunea integerrima Tixier 1979 [Dubia et Excludenda]. 

Diplasiolejeunea jonesii Tixier MDG(1). 

Diplasiolejeunea kraussiana (Lindenb.) Steph. BUR(2), MDG(1), MLW(563), RWA(2), SAF(2), 

TAN(2), UGA(50), ZAI(2), ZIM(2). 

Diplasiolejeunea lemuriana Tixier MDG(1). 

Diplasiolejeunea magnistipulata Tixier CMN(2), MDG(1). 

Diplasiolejeunea palustrium Tixier MDG(1). 

Diplasiolejeunea pellucida (Meissn. ex Spreng.) Schiffn. 1893 [Dubia et Excludenda]. This name has 

sometimes been applied erroneously to African specimens of D. albifolia (Taylor) E.W.Jones. 

Diplasiolejeunea phyllarthonii Tixier MDG(1). 

Diplasiolejeunea ramicola Tixier MDG(1). 

Diplasiolejeunea runssorensis Steph. BUR(2), KEN(2), MDG(1), RWA(2), TAN(2), UGA(50), ZAI(2). 

Diplasiolejeunea runssorensis var. australis E.W.Jones 1974 = D. villaumei Steph. 1916. 

Diplasiolejeunea subcornuta Tixier MDG(1), MLW(563), SEY(4). 

Diplasiolejeunea symoensii Vanden Berghen BUR(2), COM(1), KEN(2), MDG(1), MLW(563), 

RWA(2), TAN(2), UGA(42,50), ZAI(2). 

Diplasiolejeunea tubulata Grolle, nom. herb. = D. zakiae Tixier 1977. 

Diplasiolejeunea utriculata Steph. MDG(1), TAN(545). 

Diplasiolejeunea vandenberghenii Grolle 1960 = D. cavifolia Steph. 1895. 

Diplasiolejeunea villaumei Steph. MDG(1), MLW(563), TAN(2). 

Diplasiolejeunea zakiae Tixier KEN(2), MDG(26), TAN(2). 

Diplophyllaea biapiculata (Hook.f. et Taylor) Trevis. 1877 = Anastrophyllum piligerum (Nees) Steph. 1893. 

Diplophylleia imbricata (Wils.) Trevis. 1877 = Anastrophyllum piligerum (Nees) Steph. 1893. 

Diplophylleia incumbens (Lehm. et Lindenb.) Trevis. 1877 = Anastrophyllum piligerum (Nees) Steph. 1893. 

Diplophylleia piligera (Nees) Trevis. 1877 = Anastrophyllum piligerum (Nees) Steph. 1893. 

Diplophyllum africanum S.W.Arnell KEN(2,71), RWA(2), TAN(2). 

Drepanolejeunea africana Steph. 1913, possibly = D. cultrella (Mitt.) Steph. 1913. 

Drepanolejeunea angustifolia (Mitt.) Grolle 1965 [Dubia et Excludenda]. 

Drepanolejeunea ankasica E.W.Jones GHA(2). 

Drepanolejeunea cambouena Steph. COM(1), MAU(1), MDG(1), SEY(1,108), TAN(2). 

Drepanolejeunea capensis Steph. 1913 = D. hamatifolia (Hook.) Schiffn. 1893.


Drepanolejeunea capulata (Taylor) Steph. NGA(2). 

Drepanolejeunea clavicornis Steph. 1913 = D. physaefolia (Gottsche) Steph. 1913. 

Drepanolejeunea claviformis Steph. 1896 = D. physaefolia (Gottsche) Steph. 1913. 

Drepanolejeunea cristata Steph. 1891 = D. cultrella (Mitt.) Steph. 1913. 

Drepanolejeunea cultrella (Mitt.) Steph. BIO(72), BUR(2), CMN(2), COM(1), GUI(2), IVO(2), 

KEN(2), MLW(563), RIM(10), RWA(2), SAO(2), SIE(2), TAN(2), UGA(115), ZAI(2). 

Drepanolejeunea deslooveri Vanden Berghen 1977 = D. hamatifolia (Hook.) Schiffn. 1893. 

Drepanolejeunea filicuspis Steph. 1913 = Harpalejeunea filicuspis (Steph.) Mizut. 1973. 

Drepanolejeunea friesii Vanden Berghen 1951 = D. physaefolia (Gottsche) Steph. 1913. 

Drepanolejeunea geisslerae Pócs MDG(709). 

Drepanolejeunea gomphiae Steph. 1913 = Leptolejeunea maculata (Mitt.) Schiffn. 1898. 

Drepanolejeunea hamatifolia (Hook.) Schiffn. MOZ(2), RWA(2), SAF(2,75), ZAI(2), ZIM(2). 

Drepanolejeunea helenae Pócs REU(6). 

Drepanolejeunea intorta (Besch. et Spruce) Steph. 1913 = D. physaefolia (Gottsche) Steph. 1913. 

Drepanolejeunea madagascariensis (Steph.) Grolle COM(1), KEN(2), MAU(1), MDG(1), REU(1), 

SEY(1,4,108), TAN(2). 

Drepanolejeunea mauritiana Tixier MAU(1). 

Drepanolejeunea molleri Steph. 1913 = D. cultrella (Mitt.) Steph. 1913. 

Drepanolejeunea papillosa S.W.Arnell SAF(2). 

Drepanolejeunea physaefolia (Gottsche) Steph. BIO(2,33), CMN(2), COM(1,4), GHA(2), GUI(2), 

IVO(2), KEN(2,71), MAU(1), MDG(1,4), MLW(2,563), MOZ(2), PRI(2), REU(1), RIM(10), RWA(2), 

SAF(2), SAO(2), SEY(1), TAN(2), UGA(2), ZAI(2), ZIM(2). D. physaefolia (Gottsche) Steph. was treated 

as a synonym of D. vesiculosa by Tixier (130), and for that reason records were placed under the latter name in (2). 

However, Grolle 1995 (1) considered that synonymy had yet to be proven and listed East African Island 

collections under D. physaefolia. In this checklist, all records are placed under D. physaefolia. 

Drepanolejeunea pocsii Grolle COM(1), KEN(2), MLW(563), TAN(2), ZIM(2). 

Drepanolejeunea riddleana Steph. 1914 = Harpalejeunea filicuspis (Steph.) Mizut. 1973. 

Drepanolejeunea ruandensis Vanden Berghen RWA(2), ZAI(2). 

Drepanolejeunea securifolia (Gottsche) Steph. 1913 = D. physaefolia (Gottsche) Steph. 1913. 

Drepanolejeunea symoensii Vanden Berghen et Grolle BUR(2), RWA(2), UGA(115), ZAI(2). 

Drepanolejeunea symoensii var. minor Tixier RWA(2). 

Drepanolejeunea taxiophthalma Herzog 1947 = D. trematodes (Nees) Bischl. 1968. 

Drepanolejeunea tenuis (Nees) Schiffn. 1898 = D. angustifolia (Mitt.) Grolle 1965 [Dubia et Excludenda]. 

Drepanolejeunea ternatensis (Gottsche) Schiffn. SEY(1,4,108). 

Drepanolejeunea thwaitesiana (Mitt.) Steph. 1913 [Dubia et Excludenda]. 

Drepanolejeunea trematodes (Nees) Bischl. COM(1), KEN(2), MAU(1,108), MDG(1), REU(1), 

SEY(108), TAN(2). 

Drepanolejeunea vesiculosa (Mitt.) Steph. - see note under D. physaefolia (Gottsche) Steph. 

Drepanolejeunea vesiculosa ssp. affinis Schiffn. ex Herzog 1939, nom. inval. = D. vesiculosa (Mitt.) Steph. 1913 


Dumortiera denudata Mitt. 1861 = Wiesnerella javanica Schiffn. 1896 (18). 

Dumortiera hirsuta (Sw.) Nees ANG(2), BIO(2,33,36), BUR(2), CMN(2), COM(1), ETH(2), 

KEN(2,71,665), MAU(1), MLW(2), MOZ(2), REU(1), RWA(2), SAF(2), SAO(2), SIE(2), SWZ(2), 

TAN(2), UGA(42,50,115), ZAI(2), ZIM(2). 

Dumortiera hirsuta var. nepalensis (Taylor) R.M.Schust. ZAI(2). 

Dumortiera trichocephala (Hook.) Nees 1838 = D. hirsuta var. nepalensis (Taylor) R.M.Schust. 1992. 

Eulejeunea abyssinica Gola 1914 = Lejeunea abyssinica (Gola) Cufod. 1952. 

Eulejeunea breutelii Steph. 1896 = Lejeunea caespitosa Lindenb. 1845. 

Eulejeunea caespitosa (Lindenb.) Steph. 1890 = Lejeunea caespitosa Lindenb. 1845. 

Eulejeunea camerunensis Steph. 1923 = Lejeunea camerunensis (Steph.) E.W.Jones 1972. 

Eulejeunea compressistipula Pearson 1922, perhaps = Lejeunea confusa E.W.Jones 1972. 

Eulejeunea corbieri Steph. 1912, perhaps = Lejeunea ibadana A.J.Harr. et E.W.Jones 1982. 

Eulejeunea cucullata (Reinw. et al.) Schiffn. 1898 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Eulejeunea cuculliloba Steph. 1891, nom. nud. = Lejeunea caespitosa Lindenb. 1845. 

Eulejeunea ecarinata Steph. Nov 1890 = Lejeunea ecarinata (Steph.) Steph. Dec 1890.


Eulejeunea ecklonii Lindenb. ex Steph. 1890, nom. inval. = Lejeunea eckloniana Lindenb. 1845. 

Eulejeunea elobulata Sim 1926 = Lejeunea elobulata (Sim) S.W.Arnell 1963. 

Eulejeunea expansa Steph. 1912 = Lejeunea tabularis (Spreng.) Gottsche et al. 1845. 

Eulejeunea flava (Sw.) Steph. Mar-Apr 1888 = Lejeunea flava (Sw.) Nees 1838. 

Eulejeunea flavovirens (Ångstr.) Steph. 1913 = Lejeunea flavovirens Ångstr. 1873. 

Eulejeunea hepaticola Steph. 1888 = Lejeunea hepaticola (Steph.) Steph. 1915. 

Eulejeunea isomorpha (Gottsche) Steph. 1901 = Lejeunea tabularis (Spreng.) Gottsche et al. 1845. 

Eulejeunea jungneri Steph. 1896, perhaps = Taxilejeunea conformis (Nees et Mont.) Steph. 1914. 

Eulejeunea lamacerina Steph. 1896 = Lejeunea lamacerina (Steph.) Schiffn. 1902. 

Eulejeunea microclada Pearson 1931, perhaps = Lejeunea camerunensis (Steph.) E.W.Jones 1972. 

Eulejeunea obtusata (Gottsche) Pearson 1892, nom. inval. = Lejeunea obtusata Gottsche 1882. 

Eulejeunea rodriguezii Steph. Mar-Apr 1888 = Lejeunea rodriguezii (Steph.) Steph. 1891. 

Eulejeunea saccatiloba Steph. 1891 = Lejeunea caespitosa Lindenb. 1845. 

Eulejeunea setacea Steph. 1912 = Lejeunea setacea (Steph.) Steph. 1914. 

Eulejeunea squarrosa Steph. 1895 = Lejeunea squarrosa (Steph.) Steph. 1915. 

Eulejeunea tabularis (Spreng.) Steph. 1890 = Lejeunea tabularis (Spreng.) Gottsche et al. 1845. 

Eulejeunea vulgariformis (Gottsche) Pearson 1892, nom. inval. = Lejeunea vulgariformis Gottsche 1882. 

Eulejeunea wilmsii Steph. 1892 = Lejeunea eckloniana Lindenb. 1845. 

Euosmolejeunea africana (Steph.) R.M.Schust. et Kachroo 1968 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 

1976. 

Euosmolejeunea brachytoma (Gottsche) Steph. 1914 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964. 

Euosmolejeunea contigua (Nees) Steph. 1890 = Pycnolejeunea contigua (Nees) Grolle 1979. 

Euosmolejeunea grandistipula (Steph.) Steph. 1914 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964. 

Euosmolejeunea inflata (Steph.) R.M.Schust. et Kachroo 1968 = Cheilolejeunea intertexta (Lindenb.) Steph. 

1897. 

Euosmolejeunea longiflora (Taylor) Steph. 1914 = Cheilolejeunea longiflora (Taylor) R.M.Schust. 1980. 

Euosmolejeunea montagnei (Gottsche) Steph. 1895 = Cheilolejeunea montagnei (Gottsche) R.M.Schust. 1963. 

Euosmolejeunea principensis (Steph.) R.M.Schust. et Kachroo 1968 = Cheilolejeunea serpentina (Mitt.) Mizut. 

1963. 

Euosmolejeunea robillardii Steph. 1914 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964. 

Euosmolejeunea ruwenzorensis S.W.Arnell 1956 = Cheilolejeunea ruwenzorensis (S.W.Arnell) R.M.Schust. 

1963. 

Euosmolejeunea serpentina (Mitt.) Steph. 1914 = Cheilolejeunea serpentina (Mitt.) Mizut. 1963. 

Euosmolejeunea thomeensis Steph. 1914 = Cheilolejeunea montagnei (Gottsche) R.M.Schust. 1963. 

Euosmolejeunea trifaria (Reinw. et al.) Steph. 1888 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964. 

Evansiolejeunea renistipula (Steph.) Vanden Berghen 1948 = E. roccatii (Gola) Vanden Berghen 1960. 

Evansiolejeunea roccatii (Gola) Vanden Berghen RWA(2), UGA(2,96), ZAI(2). 

Exormotheca africana Steph. 1924 = E. pustulosa Mitt. 1870. 

Exormotheca bulbigena Bornefeld, O.H.Volk et R.Wolf NAM(19). 

Exormotheca holstii Steph. NAM(2,19), SAF(2,19), TAN(2), ZIM(2,9). 

Exormotheca megastomata Marquand 1930 = E. holstii Steph. 1899. 

Exormotheca pustulosa Mitt. ANG(2), BOT(2), CHA(2), COM(1), CVI(2,37), ETH(2), KEN(2), 

LES(2,75), NAM(2), REU(1), SAF(2), STH(2), TAN(2), ZIM(2,9). 

Exormotheca youngii S.W.Arnell 1954 = E. holstii Steph. 1899. 

Fimbraria gondarensis Gola 1914 = Asterella pappii (Gola) Grolle 1976. 

Fimbraria marginata Nees 1820 = Asterella marginata (Nees) S.W.Arnell 1963. 

Fimbriaria abyssinica Gottsche 1846 = Asterella abyssinica (Gottsche ) Grolle 1972. 

Fimbriaria angolensis Steph. 1899 = Asterella angolensis (Steph.) S.W.Arnell 1963. 

Fimbriaria bachmannii Steph. 1894 = Asterella bachmannii (Steph.) S.W.Arnell 1963. 

Fimbriaria boryana Mont. 1838 = Asterella syngenesica (Bory) Grolle 1974. 

Fimbriaria coronata Steph. 1917 = Asterella coronata (Steph.) H.A.Mill. 1981. 

Fimbriaria dissoluta Steph. 1895 = Asterella dissoluta (Steph.) Grolle 1989. 

Fimbriaria gigantea Steph. TAN(2). 

Fimbriaria incrassata Steph. 1899 = Asterella sp. 

Fimbriaria kamerunensis Steph. 1917 = Asterella sp. 

Fimbriaria linearis Steph. 1895 = Asterella linearis (Steph.) M.Howe 1898. 

Fimbriaria muscicola Steph. 1892 = Asterella muscicola (Steph.) S.W.Arnell 1963.


Fimbriaria pappii Gola 1914 = Asterella pappii (Gola) Grolle 1976. 

Fimbriaria pirottae Gola ETH(2). 

Fimbriaria preussii Schiffn. in Steph. 1895 = Asterella sp. 

Fimbriaria pusilla Mitt. 1888 = Asterella pappii (Gola) Grolle 1976. 

Fimbriaria volkensii Steph. 1899 = Asterella sp. 

Fimbriaria wilmsii Steph. 1892 = Asterella wilmsii (Steph.) S.W.Arnell 1963. 

Folioceros fuciformis (Mont.) Bharadw. 1975 = Anthoceros fuciformis Mont. 1843 

Folioceros incurvus (Steph.) Bharadw. 1972 = Anthoceros incurvus Steph. 1893. 

Fossombronia angulifolia Perold LES(86,91). 

Fossombronia angulosa (Dicks.) Raddi CVI(2,37). 

Fossombronia caespitiformis De Not. ex Rabenh. ETH(2). 

Fossombronia capensis S.W.Arnell SAF(2,91). 

Fossombronia capensis var. spiralis Perold SAF(87,91). 

Fossombronia cederbergensis Perold SAF(84,91). 

Fossombronia crispa Nees SAF(2,83,91), ZIM(2). Kürschner, 2001 (558) considered that "probably F. crispa 

is erroneously reported from Socotra (Al-Gifri et al, 1995 (692)) and refers to R. caespitiformis". 

Fossombronia cristata Lindb. 1873 = F. wondraczekii (Corda) Dumort. ex Lindb. 1873. 

Fossombronia densilamellata S.W.Arnell SAF(2,79,91). 

Fossombronia elsieae Perold SAF(88,91). 

Fossombronia foveolata Lindb. REU(45), SAF(2). 

Fossombronia gemmifera Perold LES(86), SAF(91). 

Fossombronia glenii Perold LES(86), SAF(91). 

Fossombronia grandis Steph. KEN(2), TAN(2), UGA(96). 

Fossombronia hamato-hirta Steph. 1894 = F. wondraczeckii (Corda) Dumort. 1835. 

Fossombronia husnotii Corb. CMN(2), ETH(2), SIE(2), TAN(2), ZAI(2), ZAM(2). 

Fossombronia hyalorhiza Perold SAF(90,91). 

Fossombronia indica Steph. CMN(2), GHA(2), NGA(2), SIE(2), TOG(2). 

Fossombronia intestinalis Taylor SAF(2). 

Fossombronia leucoxantha (Lehm.) Gottsche et al. SAF(2,91). 

Fossombronia marindae Perold SAF(90,91). 

Fossombronia montaguensis S.W.Arnell LES(75), SAF(2,91). 

Fossombronia monticola Perold SAF(90,91). 

Fossombronia nyikaensis Perold MLW(664). 

Fossombronia occidento-africana S.W.Arnell 1952 = F. indica Steph. 1917. 

Fossombronia porphyrorhiza (Nees) Prosk. [Dubia et Excludenda]. According to Jones (679), this species was 

once recorded, as Noteroclada porphyrorhiza, from Mt. Kilimanjaro, Tanzania. However, as no published 

details have been located, the record is treated as dubious, pending further information coming to light. 

Fossombronia pulvinata Steph. RWA(2). 

Fossombronia pusilla (L.) Nees CVI(2,37), LES(75), MLW(2), REU(1), RWA(2), ZAI(2). Though 

reported from southern Africa, Perold 1999e (91) considered the presence there of F. pusilla is yet to be 

confirmed. The specimen reported from Lesotho in Hodgetts et al. 1999 (75) should be re-examined. 

Fossombronia renateae Perold SAF(89,91). 

Fossombronia rwandaensis Perold RWA(85). 

Fossombronia spinifolia Steph. SAF(2,91). 

Fossombronia spinosa Perold SAF(88,91). 

Fossombronia stephanii Schiffn. ex Steph. REU(1). 

Fossombronia straussiana Perold MLW(79,91), SAF(79,91). 

Fossombronia swaziensis Perold SAF(86,91), SWZ(86,91). 

Fossombronia tumida Mitt. SAF(2,91). 

Fossombronia wondraczekii (Corda) Dumort. ex Lindb. ASC(2), MAU(1). 

Fossombronia zeyheri Steph. 1900 = F. crispa Nees 1846 (82, 83). 

Fossombronia zuurbergensis Perold SAF(663). 

Frullania abyssinica Steph. 1910, perhaps = F. trinervis (Lehm.) Drège 1843. 

Frullania acutiloba Gerola ETH(2).


Frullania affinis Nees et Mont. 1843 [Dubia et Excludenda]. 

Frullania africana Steph. 1891 = F. riojaneirensis (Raddi) Ångstr. 1876. 

Frullania albertii Steph. BUR(2), KEN(2). 

Frullania anderssonii Ångstr. COM(1), MAU(1,43), MDG(1), REU(1,4), SAF(2), SEY(1). 

Frullania angolensis Steph. 1910 = F. obscurifolia Mitt. 1879. 

Frullania angulata Mitt. ANG(2), ANN(2,33), BIO(2,33,36,72), BUR(2), CMN(2), COM(1), ETH(2), 

KEN(2,665), MAU(1), MDG(1), MLW(2,114), MOZ(2), NGA(2), REU(1,4), RWA(2), SAO(2), 

TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Frullania angulata var. goetzeana (Steph.) Demaret et Vanden Berghen 1950 = F. angulata var. angulata Mitt. 

1863. 

Frullania angulata var. laciniata Demaret et Vanden Berghen MDG(1), MLW(2). 

Frullania angulata fo. serratoides Vanden Berghen ZAI(2). 

Frullania angustifolia Steph. 1910 = F. trinervis (Lehm.) Drège 1843. 

Frullania apertilobula Gerola ETH(2). 

Frullania apicalis Mitt. ANN(2), BIO(2,33,36), BUR(2), COM(1), ETH(2), GHA(2), GUI(2), IVO(2), 

KEN(2,71), LBR(2), MAU(1,4), MDG(1,4), MLW(114), MOZ(2), NGA(2), REU(1,4), RIM(10), 

ROD(1), RWA(2), SAO(2), SEY(4,108), SIE(2), TAN(2), UGA(2), ZAI(2), ZIM(2). 

Frullania apicalis var. camerunensis Vanden Berghen CMN(2). 

Frullania apiculata (Reinw. et al.) Nees CMN(2), COM(1), GAB(2), GHA(2), GUI(2), MAU(1), 

MDG(1,4), REU(1,4), RIM(10), RWA(2), SEY(1,4), ZAI(2). 

Frullania apiculata Dumort., nom. inval. = F. apiculata (Reinw. et al.) Nees in Gottsche et al. 1845. 

Frullania arecae (Spreng.) Gottsche BIO(2,33), BUR(2), CMN(2), ETH(2), GHA(2), KEN(2,71,665), 

MDG(1), MLW(2), MOZ(2), NGA(2), REU(1), RWA(2), SAF(2,75), SAO(2), SIE(2), TAN(2), 

UGA(2,42,96), ZAI(2), ZIM(2). 

Frullania armatistipula Steph. 1910 = F. ericoides (Nees) Mont. 1839. 

Frullania bangiensis Steph. 1910 = F. squarrosa Dumort. 1835. 

Frullania bessonii Steph. 1893 = F. loricata Pearson 1891. 

Frullania bilabiata Steph. 1911 = F. loricata Pearson 1891. 

Frullania boivinii Steph. 1911 = F. serrata Gottsche 1845. 

Frullania borbonica Lindenb. MAU(1), MDG(1), REU(1). 

Frullania borgenii Pearson 1891 = F. obscurifolia Mitt. 1879. 

Frullania brasiliensis Raddi 1822 [Dubia et Excludenda]. 

Frullania brunnea (Spreng.) Gottsche et al. MLW(2), SAF(2). 

Frullania brunnthaleri Steph. 1911 = F. apicalis var. apicalis Mitt. 1879. 

Frullania bullata Steph. MAU(1), MDG(1), MOZ(2), TAN(2). 

Frullania bursicula Steph. 1911 = F. socotrana Mitt. 1888. 

Frullania bystroemii S.W.Arnell 1961 = F. spongiosa Steph. 1894. 

Frullania caffraria Steph. ANG(2), BIO(2,33), CMN(2), DJI(2), ERI(2), ETH(2), KEN(2,21,71,665), 

MAU(1), MDG(1), MLW(2,114), NGA(2), REU(1), RWA(2), SAF(2), TAN(2), UGA(2,50,96), 

ZAI(2), ZAM(2), ZIM(2). 

Frullania caffraria fo. anomala Vanden Berghen ZAI(2). 

Frullania cambouena Steph. 1891 = F. loricata Pearson 1891. 

Frullania camerunensis Steph. 1911 = F. apiculata (Reinw. et al.) Nees 1845. 

Frullania capensis Gottsche MAU(1), MDG(1), MLW(114), MOZ(2), REU(1,4), SAF(2), TAN(2), 

ZIM(2). 

Frullania capensis sensu Sim 1926 [non Gottsche 1845] = F. obscurifolia Mitt. 1879. 

Frullania cardotii Steph. 1910 = F. obscurifolia Mitt. 1879. 

Frullania cavallii Gola UGA(2). 

Frullania chiovendae Steph. ETH(2). 

Frullania clavellifera Steph. 1911 = F. lindenbergii Lehm. 1844. 

Frullania colonica Steph. 1910 = F. spongiosa Steph. 1894. 

Frullania conjugata Steph. 1911 = F. serrata var. serrata Gottsche 1845. 

Frullania cordata Mitt. 1863 = F. serrata var. serrata Gottsche 1845. 

Frullania cordifolia Steph. 1911 = F. angulata var. angulata Mitt. 1863.


Frullania creberrima Steph. 1911 = F. loricata Pearson 1891. 

Frullania crenuliflora Pearson 1924, perhaps = F. trinervis (Lehm.) Drège 1843. 

Frullania crististipula Steph. 1894 = F. arecae (Spreng.) Gottsche 1863. 

Frullania cuculliloba Steph. 1911 = F. obscurifolia Mitt. 1879. 

Frullania cyparioides (Schwägr.) Nees 1838 [Dubia et Excludenda]. 

Frullania dalleana Gerola 1947, perhaps = F. socotrana Mitt. 1888. 

Frullania densepinnata Steph. 1911 = F. apicalis Mitt. 1878. 

Frullania dentilobula Steph. 1894 = F. arecae (Spreng.) Gottsche 1863. 

Frullania depressa Mitt. BUR(2), CMN(2), COM(4), ETH(2), KEN(2), LES(75), MDG(1), MLW(114), 

NGA(2), RWA(2), SAF(2,75), SIE(2), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Frullania dilatata (L.) Dumort. CVI(37,124). 

Frullania diptera (Lehm.) Drège ANG(2), ANN(2,33), BEN(22), GHA(2), KEN(2,665), MAU(1), 

MDG(1), MLW(114), NGA(2), REU(1,4), SAF(2), SAO(2), TAN(2), TOG(101), UGA(42,50), ZAI(2). 

Frullania dupuisana Steph. ex Bonner 1965, nom. nud. = F. ericoides (Nees) Mont. 1839. 

Frullania dusenii Schiffn. in sched. [non Steph. 1905] = F. spongiosa Steph. 1894. 

Frullania ecklonii (Spreng.) Gottsche et al. CMN(2), KEN(2), MLW(2), SAF(2), SAO(2), TAN(2), 

UGA(2,96), ZAI(2). 

Frullania elliotii Steph. 1894 = F. angulata Mitt. 1863. 

Frullania emergens Mitt. 1863 = F. ecklonii (Spreng.) Gottsche et al. 1845. 

Frullania epiphylla Vanden Berghen 1976, hom. illeg. [non S.Hatt. 1974] = F. vandenberghenii Pócs 1979. 

Frullania eplicata Steph. GHA(2), MLW(114), MOZ(2), SAF(2), TAN(2), ZAI(2), ZAM(2), ZIM(2). 

Frullania ericoides (Nees) Mont. ALD(34), ANG(2), ANN(2,33), ASC(2), BEN(22), BIO(2,33), 

BUR(2), CAF(2), CMN(2), COM(1), CON(2), CVI(2,37), ERI(121), ETH(2), GAB(2), GHA(2), 

GUI(2), IVO(51,53), KEN(2), MAU(1,43), MDG(1), MLW(2,114), MOZ(2), NGA(2), PRI(2), REU(1), 

ROD(1), RWA(2), SAF(2,75), SEY(1,4), SIE(2), SOC(2), TAN(2), TOG(2,101), UGA(2,96), ZAI(2), 

ZAM(2), ZIM(2). 

Frullania erosolimbata Steph. 1924 = F. caffraria Steph. 1894. 

Frullania erronea Steph. 1911 = F. depressa Mitt. 1863. 

Frullania exigua Steph. 1910 = F. obscurifolia Mitt. 1879. 

Frullania filiformis Steph. 1910 = F. loricata Pearson 1891. 

Frullania fusca Steph. 1894 = F. lindenbergii Lehm. 1844. 

Frullania gabonensis Vanden Berghen CMN(2), GAB(2), MDG(1,4). 

Frullania garsidei S.W.Arnell 1953 = F. variegata Steph. 1894. 

Frullania goetzeana Steph. 1901 = F. angulata var. angulata Mitt. 1863. 

Frullania gracilis (Reinw. et al.) Gottsche et al. 1845 [Dubia et Excludenda]. 

Frullania grossiclava Steph. MDG(1,4), MLW(114), MOZ(2), TAN(2), ZAI(2), ZIM(2). 

Frullania hamatirostra Gerola 1947, perhaps = F. ericoides (Nees) Mont. 1839. 

Frullania hedbergii Vanden Berghen 1953 = F. albertii Steph. 1916. 

Frullania hians (Lehm. et Lindenb.) Mont. et Nees 1839 = F. arecae (Spreng.) Gottsche 1863. 

Frullania hildebrandtii Steph. 1910 = F. trinervis (Lehm.) Drège 1843. 

Frullania hispida Steph. 1911 = F. angulata Mitt 1863. 

Frullania hoehneliana Steph. 1894 = F. caffraria Steph. 1894. 

Frullania holstii Steph. 1894 = F. angulata var. angulata Mitt. 1863. 

Frullania humbertii Vanden Berghen MDG(1), REU(1). 

Frullania imerinensis Steph. MDG(1), RWA(2), SEY(4). 

Frullania inflatiloba Steph. 1910 = F. diptera (Lehm. et Lindenb.) Drège 1843. 

Frullania insularis Beauverd 1924 = F. apicalis Mitt. 1878. 

Frullania johannensis Steph. 1911 = F. serrata var. pertenuis (Nees) Gottsche et al. 1845. 

Frullania junghuhniana Gottsche 1845 [Dubia et Excludenda]. 

Frullania jungneri Steph. 1911 = F. angulata var. angulata Mitt. 1863. 

Frullania kilimanjarica Vanden Berghen 1953 = F. apicalis var. apicalis Mitt. 1879. 

Frullania kizarensis Steph. 1911 = F. angulata var. angulata Mitt. 1863. 

Frullania lacerata Steph. 1911 = F. serrata Gottsche 1845. 

Frullania laceriloba Steph. 1911 = F. apicalis var. apicalis Mitt. 1879. 

Frullania laceriloba var. acutifolia S.W.Arnell 1956 = F. apicalis var. apicalis Mitt. 1879. 

Frullania laxepinnata Steph. 1895, nom. nud. [non Steph 1911] = F. angulata var. angulata Mitt. 1963.


Frullania lepervanchii Steph. 1911 = F. lindenbergii Lehm. 1844. 

Frullania letestui Vanden Berghen CAF(2), CMN(2), GAB(2), ZAI(2). 

Frullania levicalyx Steph. 1912 = F. trinervis (Lehm.) Drège 1843. 

Frullania lindenbergii Lehm. KEN(2), MAU(1), MDG(1,4), MLW(114), MOZ(2), REU(1,4), SAF(2), 

SEY(4,108), TAN(2), UGA(50), ZIM(2). 

Frullania lobangensis Steph. 1910 = F. caffraria Steph. 1894. 

Frullania longirostris Steph. 1894 = F. ecklonii (Spreng.) Gottsche et al. 1845. 

Frullania longistipula Steph. MDG(1). 

Frullania loricata Pearson MDG(1), MLW(2,114), TAN(2), UGA(2). 

Frullania loricata var. laxa Pearson MDG(1). 

Frullania macroloba Steph. 1910 = F. anderssonii Ångstr. 1873. 

Frullania madagascariensis Gottsche 1882, nom. dub. [Dubia et Excludenda]. 

Frullania mauritiana Austin 1870, nom. dub. [Dubia et Excludenda]. 

Frullania mildbraedii Steph. 1911 = F. serrata var. serrata Gottsche 1845. 

Frullania molleri Steph. 1894, perhaps = F. caffraria Steph. 1894 (147). 

Frullania multiramosa Steph. 1894 = F. capensis Gottsche 1845. 

Frullania mundiana Lindenb. et Gottsche 1847 = F. arecae (Spreng.) Gottsche 1863. 

Frullania natalensis Sim 1926 = F. depressa Mitt. 1863. 

Frullania nervosa Mont. 1840 = F. tamarisci (L.) Dumort. 1835. 

Frullania newtonii Steph. 1911, perhaps = F. angulata var. angulata Mitt. 1863. 

Frullania nigricans Steph. 1911 = F. apiculata (Reinw. et al.) Nees 1845. 

Frullania nodulosa (Reinw. et al.) Gottsche et al. CMN(2), COM(1), GHA(2), IVO(2), MDG(1,4), 

NGA(2), SEY(1), SIE(2), TAN(2), ZAI(2). 

Frullania nutans Steph. 1901, perhaps = F. caffraria Steph. 1894. 

Frullania obconica Steph. 1910 = F. trinervis (Lehm.) Drège 1843. 

Frullania obscura Steph. 1910, hom. illeg. [non (Lehm. et Lindenb.) Nees 1845] = F. anderssonii Ångstr. 1873. 

Frullania obscurifolia Mitt. ANG(2), ANN(2,33), BUR(2), CAF(2), CMN(2), COM(4), CON(2), DJI(2), 

ETH(2), GHA(2), KEN(2,21,71,665), LES(75), MAU(1), MDG(1,4), MLW(2,114), MOZ(2), NGA(2), 

ROD(1), RWA(2), SAF(2,75), SEY(1,4), SIE(2), SWZ(2), TAN(2), UGA(2,42), ZAI(2), ZAM(2), 

ZIM(2). 

Frullania onraedtii Vanden Berghen MAU(1). 

Frullania ovalifolia Kiaer et Pearson 1891 = F. anderssonii Ångstr. 1873. 

Frullania ovifolia Kiaer et Pearson ex Steph. 1910 = F. anderssonii Ångstr. 1873. 

Frullania parisii Steph. 1908 = F. trinervis (Lehm.) Drège 1843. 

Frullania perrotana Steph. 1911 = F. apiculata (Reinw. et al.) Nees 1845. 

Frullania picta Steph. 1894 [Dubia et Excludenda]. 

Frullania platyflora Pearson 1922 = F. eplicata Steph. 1911. 

Frullania purpurea Steph. COM(4), MAU(1), MDG(1,4), NGA(2), RIM(10), SAO(2), SEY(4), SIE(2). 

Frullania purpureopicta Steph. 1910 = F. socotrana Mitt. 1888. 

Frullania rehmannii Steph. 1911 = F. capensis Gottsche 1845. 

Frullania renauldii Steph. 17 Jul 1911 [non 28 Feb 1911] = F. apicalis Mitt. 1878. 

Frullania renauldii Steph. 28 Feb 1911 [nec 17 Jul 1911] = F. variegata Steph. 1894. 

Frullania renistipula Steph. 1910 = F. anderssonii Ångstr. 1873. 

Frullania repandistipula Sande Lac. COM(1), MDG(1), REU(1,4), SEY(1,4,108), TAN(2). 

Frullania rigida Steph. MDG(1). 

Frullania riojaneirensis (Raddi) Ångstr. CMN(2), GAB(2), GHA(2), GUI(2), NGA(2), PRI(2), SAO(2), 

TAN(2), TOG(101), ZAI(2). 

Frullania riparia Hampe ex Lehm. 1838 - recorded in error for F. ericoides in Cape Verde Islands. 

Frullania robillardii Steph. 1893 = F. anderssonii Ångstr. 1873. 

Frullania robusta Kiaer 1891, nom. inval. [non (Spruce) Steph. 1911] = F. loricata Pearson 1891. 

Frullania rodriguezii Steph. 1894 = F. apicalis Mitt. 1878. 

Frullania rupestris Steph. 1894 = F. schimperi Nees 1845. 

Frullania rutenbergii Gottsche 1882 = F. apicalis Mitt. 1879. 

Frullania sandei Kiaer ex Steph. 1894 = F. caffraria Steph. 1894. 

Frullania schimperi Nees BUR(2), CMN(2), ETH(2), KEN(2,71,665), MDG(1), RWA(2), TAN(2),


UGA(50). 

Frullania schimperi var. laciniata Vanden Berghen MDG(1). 

Frullania semienana Gola ETH(2). 

Frullania serrata Gottsche CMN(2), COM(1), ETH(2), KEN(2), MAU(1), MDG(1,4), MLW(2,114), 

MOZ(2), REU(1,4), RWA(2), SAF(2), SAO(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Frullania serrata var. pertenuis (Nees) Gottsche et al. COM(1), MDG(1,4), RWA(2), TAN(2), ZAI(2). 

Frullania silvestris Sim 1926 = F. depressa Mitt. 1863. 

Frullania socotrana Mitt. CMN(2), CVI(2,37), KEN(2), LES(75), MDG(1), MOZ(2), SAF(2), SOC(2), 

TAN(2), UGA(2,50,96), ZAI(2). 

Frullania sphaeroflora Steph. 1911 = F. bullata Steph. 1910. 

Frullania spongiosa Steph. BEN(22), CMN(2), CVI(2,37), GHA(2), IVO(2,53), MDG(4), NGA(2), 

RIM(10), SAF(2), SEY(4), TAN(2), TOG(101), UGA(50), ZAI(2), ZIM(2). 

Frullania squarrosa Gottsche et al. 1845 = F. ericoides (Nees) Mont. 1839. 

Frullania staudtiana Steph. 1911, probably = F. apiculata (Reinw. et al.) Nees 1845. 

Frullania stephanii Schiffn. 1893, perhaps = F. caffraria Steph. 1894. 

Frullania stipatiflora Steph. 1911 = F. apicalis Mitt. 1878. 

Frullania stricta (F.Weber) Lindenb. 1845 = F. serrata Gottsche 1845. 

Frullania stylata Gerola 1947, perhaps = F. trinervis (Lehm.) Drège 1843. 

Frullania subatrata Steph. 1911 = F. angulata var. angulata Mitt. 1863. 

Frullania subplana Gottsche ex Steph. 1910 = F. bullata Steph. 1910. 

Frullania tamarisci (L.) Dumort. CVI(2,20,37). 

Frullania tatanarivensis Steph. 1894 = F. ericoides (Nees) Mont. 1839. 

Frullania thomeensis Steph. 1910, probably = F. arecae (Spreng.) Gottsche 1863. 

Frullania tixieriana Vanden Berghen 1976 = F. repandistipula Sande Lac. 1855. 

Frullania traversiana Steph. 1910 = F. depressa Mitt. 1863. 

Frullania trinervis (Lehm.) Drège ANG(2), BIO(2,33), CAF(2), CON(2), ETH(2), GHA(2), GUI(2), 

KEN(2), LES(2,75), MAU(4), MDG(1), MLW(2,114), MOZ(2), NGA(2), SAF(75), SEY(4), SIE(2), 

SOC(94), TAN(2), TOG(101), UGA(2,42), ZAI(2), ZAM(2), ZIM(2). 

Frullania truncatiloba sensu Vanden Berghen 1961 [non Steph. 1910] = F. depressa Mitt. 1863. 

Frullania truncatiloba Steph. 1910 [non sensu Vanden Berghen 1961] = F. riojaneirensis (Raddi) Ångstr. 1876. 

Frullania ubangiensis Steph. 1910 = F. trinervis (Lehm.) Drège 1843. 

Frullania usagara Mitt. 1886 = F. ericoides (Nees) Mont. 1839. 

Frullania usambarana Schiffn. ex Steph. MDG(1,4), REU(4), TAN(2). 

Frullania usambarana var. reducta Vanden Berghen MDG(1), REU(1). 

Frullania usambarensis Steph. 1910 = F. spongiosa Steph. 1894. 

Frullania vandenberghenii Pócs MDG(1,4), UGA(50). 

Frullania vanderystii Steph. 1910 = F. obscurifolia Mitt. 1879. 

Frullania varia Gottsche 1882 = F. angulata Mitt. 1863. 

Frullania variegata Steph. BUR(2), CMN(2), MAU(1), MDG(1,4), MLW(114), REU(1), RWA(2), 

SAF(2), TAN(2). 

Frullania welwitschii Steph. 1901 = F. ericoides (Nees) Mont. 1839. 

Frullanoides riojaneirensis Raddi 1822 = Frullania riojaneirensis (Raddi) Ångstr. 1876. 

Frullanoides tristis (Steph.) van Slageren ANG(2), BUR(2), CAF(2), CMN(2), ETH(2), IVO(2), 

KEN(2), MDG(1), NGA(2), RWA(2), SAF(2,75), SIE(2), TAN(2), UGA(50), ZAI(2). 

Gamochaetium concretum (Mont.) Trevis. 1879 = Lophocolea concreta Mont. 1845. 

Geocalyx orientalis Besch. et Spruce REU(1). 

Gongylanthus ericetorum (Raddi) Nees BUR(2), CMN(2), ETH(2), KEN(2), LES(75), MDG(1), 

MLW(2,29), REU(1), SAF(2,75), SIE(2), TAN(2), ZAI(2), ZAM(2), ZIM(2). Listed in error from 

Rwanda in (2). 

Gongylanthus renifolius (Mitt.) Steph. RWA(2), SAF(2), TAN(2). Listed in error from Burundi in (2). 

Gongylanthus richardsii E.W.Jones CMN(2), SIE(2), TAN(2), UGA(42). 

Gongylanthus scariosus (Lehm.) Steph. CMN(2), ETH(2), MLW(29), REU(1), SAF(2). 

Gottschea alata (Lehm.) Nees 1844 = Schistochila alata (Lehm.) Schiffn. 1893. 

Gottschea borbonica (Steph.) Bonner 1966 = G. neesii Mont. 1843.


Gottschea engleriana (Steph.) Grolle et Zijlstra COM(1,4), MLW(2,109), RWA(2), TAN(2). 

Gottschea neesii Mont. COM(4), MAS(1), MAU(1,4), MDG(1,4), REU(1,4). 

Gottschea schwaegricheniana Nees et Mont. 1843, nom. inval. = G. neesii Mont. 1843. 

Gottschea sphagnoides (Schwägr.) Lindb. MAU(1), MLW(2,109), MOZ(2), REU(1,4), SAF(2), ZIM(2). 

Gottschea thouarsii (Hook.) Nees 1844 = G. sphagnoides (Schwägr.) Lindb. 1876. 

Gottschelia schizopleura (Spruce) Grolle MDG(1,4), MLW(2,114), REU(1,4), TAN(2). 

Grimaldia abyssinica Gola 1914 = Plagiochasma eximium (Schiffn. in Steph.) Steph. 1898. 

Grimaldia angustifolia (Necker) Schiffn. 1893 = Mannia androgyna (L. emend. Lindb.) A.Evans 1938. 

Grimaldia capensis Steph. 1898 = Mannia capensis (Steph.) S.W.Arnell 1963. 

Grimaldia delessertii Mont. 1846, nom. inval. = Marchantia globosa Brid. ex F.Weber 1815. 

Grimaldia dichotoma Raddi 1818 = Mannia androgyna (L. emend. Lindb.) A.Evans 1938. 

Gymnanthe anisodonta (Hook.f. et Taylor) Gottsche 1864 = Tylimanthus anisodontus (Hook.f. et Taylor) Mitt. 

1875. 

Gymnanthe belangeriana (Lehm. et Lindenb.) Steph. 1894 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 

1873. 

Gymnanthe biloba Mitt. 1863 = Andrewsianthus bilobus (Mitt.) Grolle 1963. 

Gymnanthe decipiens (Hook.) Mitt. 1863 = Adelanthus decipiens (Hook.) Mitt. 1864. 

Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust. SAF(2). 

Gymnocoleopsis multiflorus (Steph.) R.M.Schust. TAN(2), ZAI(2). 

Gymnomitrion africanum (Steph.) Horik. 1943 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996 

(153). 

Gymnomitrion andinum (Herzog) Herzog SAF(2,111). 

Gymnomitrion belangerianum (Lehm. et Lindenb.) Gottsche 1844 = Notoscyphus lutescens (Lehm. et Lindenb.) 

Mitt. 1873. 

Gymnomitrion elgonense S.W.Arnell 1956 = G. laceratum (Steph.) Horik. 1943. 

Gymnomitrion elgonense var. squarrosum S.W.Arnell 1956 = G. laceratum (Steph.) Horik. 1943. 

Gymnomitrion laceratum (Steph.) Horik. KEN(2), RWA(2), SAF(2,75), TAN(2), ZAI(2). 

Gymnomitrion scariosum (Lehm.) Nees 1844 = Gongylanthus scariosus (Lehm.) Steph. 1906. 

Haplolejeunea cucullata (Steph.) Grolle CMN(2). 

Haplolejeunea sticta Grolle MDG(1), REU(4). 

Haplomitrium andinum (Spruce) R.M.Schust. 1963 = H. blumii (Nees) R.M.Schust. 1963. 

Haplomitrium blumii (Nees) R.M.Schust. RWA(2), ZAI(2). 

Haplomitrium gibbsiae (Steph.) R.M.Schust. LES(75), UGA(115). 

Harpalejeunea delessertii (Nees et Mont.) Steph. 1913 = Lepidolejeunea delessertii (Nees et Mont.) Grolle 

1984. 

Harpalejeunea filicuspis (Steph.) Mizut. COM(1), SEY(1). 

Harpalejeunea fischeri Tixier RWA(2), UGA(42). 

Harpalejeunea punctata (Herzog) S.W.Arnell 1965 [Dubia et Excludenda]. 

Harpalejeunea riddleana (Steph.) Mizut. 1973 = H. filicuspis (Steph.) Mizut. 1973. 

Harpalejeunea solitaria (Gottsche) Steph. 1913 [Dubia et Excludenda]. 

Herbertus capensis (Steph.) Sim COM(1), ETH(2), MDG(1), MLW(2), SAF(2,75), UGA(50,96). 

Herbertus capillaris Steph. 1892 [Dubia et Excludenda]. 

Herbertus dicranus (Taylor ex Gottsche et al.) Trevis. KEN(2,71), MAU(1), MDG(1), REU(1), 

RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Herbertus doggeltianus (Steph.) Demaret UGA(50), ZAI(2). 

Herbertus grossevittatus (Steph.) S.W.Arnell ex Grolle MAU(1), MDG(1), SEY(1,4). 

Herbertus juniperinus Trevis. 1877 = H. juniperoideus (Sw.) Grolle 1961 [Dubia et Excludenda]. 

Herbertus juniperoideus (Sw.) Grolle 1961 [Dubia et Excludenda]. 

Herbertus leioclada (Taylor) Kuntze 1891 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838. 

Herbertus lobatus (Steph.) Demaret UGA(50), ZAI(2). 

Herbertus mascarenicus (Steph.) S.W.Arnell ex Váòa in Váòa et al. MAS(1), REU(1). 

Herbertus mollis (Steph.) Dusén CMN(2), SAO(2). 

Herbertus ochroleuca (Spreng.) Schiffn. 1898 = Lepicolea ochroleuca (Spreng.) Spruce 1885. 

Herbertus seriatus (Steph.) S.W.Arnell 1965, nom. inval. = H. grossevittatus (Steph.) S.W.Arnell ex Grolle 

1995.


Herbertus stuhlmannii (Steph.) Demaret UGA(2), ZAI(2). 

Herbertus subdentatus (Steph.) Fulford ETH(2), MDG(4), TAN(2), ZAI(2). 

Herbertus sp. (Schisma balboi Gola) KEN(2). 

Herbertus sp. (Schisma kilimandjarense Steph.) TAN(2). 

Herbertus sp. (Schisma mascarenicum var. kikujense Gola) KEN(2). 

Herpetium integrum Nees et Mont. 1843 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Heteroscyphus argutus (Reinw. et al.) Schiffn. 1910 [Dubia et Excludenda]. 

Heteroscyphus decurrens (Nees) Schiffn. 1910 = H. splendens (Lehm. et Lindenb.) Grolle 1984. 

Heteroscyphus dubius (Gottsche) Schiffn. CAF(2), CMN(2), GHA(2,165), IVO(2), MAU(1), MDG(1), 

MLW(2,29), NGA(2), PRI(2), REU(1), RIM(10), ROD(1), SAF(2), SEY(1,13), SIE(2), TAN(2), 

TOG(101), UGA(50,115), ZAI(2). The record from British Togoland published in Jones 1953b (165) was 

erroneously listed in (2) under Togo. The site is in Ghana. 

Heteroscyphus fasciculatus (Nees) Schiffn. 1910 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Heteroscyphus grandistipus (Steph.) Schiffn. REU(1). 

Heteroscyphus grandistipus ssp. henkelii (Sim) Grolle ZIM(2). 

Heteroscyphus hamatistipulus (Steph.) Schiffn. 1910 = H. dubius (Gottsche) Schiffn. 1910. 

Heteroscyphus loangensis (Steph.) Schiffn. 1910 = H. dubius (Gottsche) Schiffn. 1910. 

Heteroscyphus lucidus (Spreng. ex Lehm.) Schiffn. 1910 = Lophocolea lucida (Spreng. ex Lehm.) Mont. 1839. 

Heteroscyphus rabenhorstii (Steph.) Schiffn. 1910 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Heteroscyphus spectabilis (Steph.) Schiffn. CMN(2), SAO(2). 

Heteroscyphus splendens (Lehm. et Lindenb.) Grolle MAU(1), MDG(4), REU(1), SEY(1), TAN(2). 

Homalolejeunea acutifolia (Taylor) Steph. 1890 = Marchesinia brachiata (Sw.) Schiffn. 1893. 

Homalolejeunea excavata (Mitt.) Steph. 1912 = Marchesinia excavata (Mitt.) Schiffn. 1893. 

Homalolejeunea henriquesii Steph. 1888 = Marchesinia excavata (Mitt.) Schiffn. 1893. 

Hyalolepidozia bicuspidata (C.Massal.) Grolle 1964 - see Paracromastigum tristanianum. 

Hygrolejeunea acuta (Mitt.) Vanden Berghen 1961 = Lejeunea acuta Mitt. 1863. 

Hygrolejeunea alata (Gottsche) Steph. 1914 = Lejeunea alata Gottsche 1845. 

Hygrolejeunea breuteliana Steph. 1914 = Lejeunea eckloniana Lindenb. 1845. 

Hygrolejeunea cordigera Steph. 1896 = Cheilolejeunea cordigera (Steph.) Grolle 1977. 

Hygrolejeunea dismieri Steph. 1913 = Acanthocoleus madagascariensis (Steph.) Kruijt 1988. 

Hygrolejeunea grossecristata Steph. 1896 = Lejeunea grossecristata (Steph.) E.W.Jones 1968. 

Hygrolejeunea leucosis (Besch. et Spruce) Steph. 1914 = Lejeunea leucosis Besch. et Spruce 1890. 

Hygrolejeunea lyratiflora (Steph.) Steph. 1914 = Lejeunea lyratiflora Steph. 1892. 

Hygrolejeunea papilionacea (Steph.) Steph. 1914 = Lejeunea papilionacea Steph. 1892. 

Hygrolejeunea parva Steph. 1914 = Lejeunea eckloniana Lindenb. 1845. 

Hygrolejeunea patellirostris Steph. 1896 = Lejeunea sp. 

Hygrolejeunea pterota (Hook.f. et Taylor) Steph. 1914 = Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle 

1988. 

Hygrolejeunea pulcherrima Steph. 1914 = Lejeunea acuta Mitt. 1863. 

Hygrolejeunea staudtiana Steph. 1914 = Lejeunea sp. 

Hygrolembidium heteromorphum (Lehm.) Grolle 1971 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 

1969. 

Hypenantron abyssinicum (Gottsche) Kuntze 1891 = Asterella abyssinica (Gottsche) Grolle 1972. 

Hypenantron gondarense (Gola) Cufod. 1952 = Asterella pappii (Gola) Grolle 1976. 

Hypenantron incrassatum (Steph.) Cufod. 1952 = Fimbriaria incrassata Steph. 1899. 

Hypenantron pappii (Gola) Cufod. 1952 = Asterella pappii (Gola) Grolle 1976. 

Hypenantron pirottae (Gola) Cufod. 1952 = Fimbriaria pirottae Gola 1914. 

Inflatolejeunea capensis S.W.Arnell 1953 = Lejeunea capensis Gottsche 1845. 

Inflatolejeunea capensis S.W.Arnell 1961 = Lejeunea capensis Gottsche 1845. 

Inflatolejeunea ecarinata (Steph.) S.W.Arnell 1961 = Lejeunea ecarinata (Steph.) Steph. 1890. 

Inflatolejeunea eplicata S.W.Arnell 1961 = Lejeunea ramosissima Steph. 1886. 

Isopaches decolorans (Limpr.) Buch 1932 = Lophozia decolorans (Limpr.) Steph. 1902. 

Isotachis aspera Steph. 1911 = I. aubertii (Schwägr.) Mitt. 1886. 

Isotachis aspera var. transvaalensis S.W.Arnell 1963, nom. inval. = I. aubertii (Schwägr.) Mitt. 1886. 

Isotachis aubertii (Schwägr.) Mitt. ASC(2), BUR(2), CMN(2), KEN(2), MAU(1), MDG(1), MLW(114), 

REU(1,4), ROD(1), RWA(2), SAF(2), TAN(2), UGA(2), ZAI(2), ZIM(2).


Isotachis capensis Herzog 1952 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Isotachis conistipula Steph. 1911 = I. aubertii (Schwägr.) Mitt. 1886. 

Isotachis gordonii Steph. 1895 = I. aubertii (Schwägr.) Mitt. 1886. 

Isotachis perfoliata Steph. 1886 = Heteroscyphus spectabilis (Steph.) Schiffn. 1910. 

Isotachis renistipula Steph. 1911 = I. aubertii (Schwägr.) Mitt. 1886. 

Isotachis rutenbergii Gottsche 1882 = I. aubertii (Schwägr.) Mitt. 1886. 

Isotachis serrulata (Sw.) Gottsche 1864 [Dubia et Excludenda]. 

Isotachis uncinata (F.Weber) Gottsche 1864 = I. aubertii (Schwägr.) Mitt. 1886. 

Iwatsukia jishibae (Steph.) N.Kitag. BIO(36), COM(4), MAU(1), MDG(4), MLW(114), REU(1,4), 

SEY(1,4). 

Jamesoniella aspera Steph. 1917 = Gongylanthus ericetorum (Raddi) Nees 1836. 

Jamesoniella colorata (Lehm.) Schiffn. SAF(2,75). 

Jamesoniella contracta (Reinw. et al.) N.Kitag. COM(4), MAU(1), MDG(4), REU(4), SEY(1,4), 

TAN(2). 

Jamesoniella grandiflora (Lindenb. et Gottsche) Steph. 1901 = Cryptochila grandiflora (Lindenb. et Gottsche) 

Grolle 1971. 

Jamesoniella oenops (Lindenb. et Gottsche) Steph. 1901 = J. colorata (Lehm.) Schiffn. 1893. 

Jamesoniella purpurascens Steph. MAU(1), MDG(1,4), MLW(2,114), REU(1), SAF(2), TAN(2), 

ZAI(2), ZAM(2). 

Jamesoniella rehmannii (Steph.) Steph 1901 = J. purpurascens Steph. 1891. 

Jamesoniella seychellarum Steph. 1917 = J. contracta (Reinw. et al.) N.Kitag. 1981. 

Jensenia spinosa (Lindenb. et Gottsche) Grolle KEN(2,71,665), MAU(1), MLW(2,29), REU(1), 

RWA(2), SAF(2), STH(2), TAN(2), UGA(50), ZAI(2). 

Jubula cambouena Steph. 1924 = J. tonkinensis Steph. 1908 [Dubia et Excludenda]. 

Jubula hutchinsiae ssp. javanica (Steph.) Verd. 1928 [Dubia et Excludenda]. 

Jubula javanica Steph. 1911 = J. hutchinsiae ssp. javanica (Steph.) Verd. 1928 [Dubia et Excludenda]. 

Jubula tonkinensis Steph. 1908 [Dubia et Excludenda]. 

Jungermannia aberrans Nees et Mont. 1843 = Andrewsianthus aberrans (Nees et Mont.) Grolle 1963. 

Jungermannia abyssinica Nees ETH(2), KEN(2), TAN(2), UGA(2). 

Jungermannia acutiloba Hook.f. et Taylor 1845 = Marchesinia brachiata (Sw.) Schiffn. 1893. 

Jungermannia adnexa Lehm. et Lindenb. 1832 = Bazzania adnexa (Lehm. et Lindenb.) Trevis. 1877 [Dubia et 

Excludenda]. 

Jungermannia alata Lehm. 1829 = Schistochila alata (Lehm.) Schiffn.1893. 

Jungermannia anisodonta Hook.f. et Taylor 1845 = Tylimanthus anisodontus (Hook.f. et Taylor) Mitt. 1875. 

Jungermannia apiculata Reinw. et al. 1824 = Frullania apiculata (Reinw. et al.) Nees 1845. 

Jungermannia arboris-vitae With. 1776 = Porella arboris-vitae (With.) Grolle 1969. 

Jungermannia arecae Spreng. 1821 = Frullania arecae (Spreng.) Gottsche 1863. 

Jungermannia ascensionis Hook.f. et Taylor 1845 = Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle 1988. 

Jungermannia atrata var. β stricta F.Weber 1815 = F. serrata Gottsche 1845. 

Jungermannia aubertii Schwägr. 1814 = Isotachis aubertii (Schwägr.) Mitt. 1886. 

Jungermannia auriculata Wils. 1841 = Mastigolejeunea auriculata (Wils.) Schiffn. 1893. 

Jungermannia aurita Lehm. 1829 = Anastrophyllum auritum (Lehm.) Steph. 1901. 

Jungermannia austroafricana S.W.Arnell 1958 = J. mildbraedii Steph. 1911. 

Jungermannia balfourii Váòa MAU(1). 

Jungermannia belangeriana Lehm. et Lindenb. 1833 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Jungermannia bergiana Lehm. et Lindenb. 1832 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Jungermannia biapiculata Hook.f. et Taylor 1845 = Anastrophyllum piligerum (Nees) Steph. 1893. 

Jungermannia bicuspidata L. 1753 = Cephalozia bicuspidata (L.) Dumort. 1835. 

Jungermannia bidentata L. 1753 = Lophocolea bidentata (L.) Dumort. 1835. 

Jungermannia bidentula F.Weber 1815 = Calypogeia bidentula (F.Weber) Nees 1845. 

Jungermannia borgenii Gottsche ex Pearson ANG(2), BOT(2), CMN(2), CON(2), KEN(2), MAU(1), 

MDG(1), MLW(2,114), REU(1), RWA(2), SAF(2), SAO(2), TAN(2), UGA(2,96), ZAI(2), ZAM(2), 

ZIM(2). 

Jungermannia boryana F.Weber 1815 = Radula boryana (F.Weber) Mont. 1842. 

Jungermannia brasiliensis Nees 1830, hom. illeg. [non Raddi 1822] = Symphyogyna brasiliensis Nees et Mont. 

1836.


Jungermannia brunnea Spreng. 1827 = Frullania brunnea (Spreng. ) Gottsche et al. 1845. 

Jungermannia byssophora Lehm. et Lindenb. 1832 = Pallavicinia lyellii (Hook.) Carruth. 1865. 

Jungermannia calyptrifolia Hook. 1816 = Colura calyptrifolia (Hook.) Dumort. 1835. 

Jungermannia campylodonta Hook.f. et Taylor 1845 = Adelanthus decipiens (Hook.) Mitt. 1864. 

Jungermannia capillaris Sw. 1788 = Kurzia capillaris (Sw.) Grolle 1964. 

Jungermannia chrysophylla Lehm. 1835 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988. 

Jungermannia colorata Lehm. 1829 = Jamesoniella colorata (Lehm.) Schiffn. 1893. 

Jungermannia concreta Gottsche 1863 = Syzygiella concreta (Gottsche) Spruce 1876. 

Jungermannia congesta Lehm. 1829 = Lethocolea congesta (Lehm.) S.W.Arnell 1955. 

Jungermannia contigua Nees 1833 = Pycnolejeunea contigua (Nees) Grolle 1979. 

Jungermannia contracta Reinw. et al. 1824 = Jamesoniella contracta (Reinw. et al.) N.Kitag. 1981. 

Jungermannia convexa Thunb. 1800, hom. illeg. [non Scop. 1772] = Bazzania nitida (F.Weber) Grolle 1960. 

Jungermannia convoluta Hook.f. et Taylor 1845, hom. illeg. [non Huebener 1834] = Symphyogyna brasiliensis 

Nees et Mont. 1836. 

Jungermannia cordistipula var. pertenuis Nees 1830 = Frullania serrata var. pertenuis (Nees) Gottsche 1845. 

Jungermannia cornuta Lindenb. 1829 = Ceratolejeunea cornuta (Lindenb.) Steph. 1895 [Dubia et Excludenda]. 

Jungermannia crispa Spreng. in sched. Hb. Zeyher = Fossombronia crispa Nees 1846 (83). 

Jungermannia cucullata Reinw. et al. 1824 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Jungermannia cucullata var. maior Nees 1830 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Jungermannia cuneata Lehm. et Lindenb. 1832 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947. 

Jungermannia cupressina Sw. 1788 = Lepidozia cupressina (Sw.) Lindenb. 1845. 

Jungermannia cuspiduligera Nees 1833 = Scapania cuspiduligera (Nees) Müll.Frib. 1915. 

Jungermannia decipiens Hook. 1813 = Adelanthus decipiens (Hook.) Mitt. 1864. 

Jungermannia decolorans Limpr. 1880 = Lophozia decolorans (Limpr.) Steph. 1902. 

Jungermannia decrescens Lehm. et Lindenb. 1832 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Jungermannia decurrens Reinw. et al. 1824, hom. illeg. [non Desv. 1818] = Heteroscyphus splendens (Lehm. & 

Lindenb.) Grolle in Grolle & Piippo 1984. 

Jungermannia diacantha Mont. 1856 = Arachniopsis diacantha (Mont.) M.Howe 1902. 

Jungermannia dichotoma var. γ Nees 1831 = Plagiochila paucidentata Mont. et Gottsche 1856. 

Jungermannia dichotoma var. α Nees 1831 = Plagiochila pectinata Willd. ex Lindenb. 1839. 

Jungermannia diclados Brid. ex F.Weber 1815 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838. 

Jungermannia diptera Lehm. 1835 = Frullania diptera (Lehm.) Drège 1843. 

Jungermannia dusenii (Steph.) Steph. CMN(2), SAO(2). 

Jungermannia ecklonii Spreng. 1827 = Frullania ecklonii (Spreng.) Gottsche et al. 1845. 

Jungermannia elliptica Lehm. et Lindenb. 1833 = Leptolejeunea elliptica (Lehm. et Lindenb.) Schiffn. 1893. 

Jungermannia elongata Lehm. 1835 = Frullania trinervis (Lehm.) Drège 1843. 

Jungermannia emarginata Ehrh. 1784 = Marsupella emarginata (Ehrh.) Dumort. 1835. 

Jungermannia ericoides Nees 1833 = Frullania ericoides (Nees) Mont. 1839. 

Jungermannia exigua Taylor 1844 = Plagiochila exigua (Taylor) Taylor 1846. 

Jungermannia expansa Lehm. 1829 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Jungermannia exsecta Schmidel 1797 = Tritomaria exsecta (Schmidel) Loeske 1909. 

Jungermannia fasciculata Nees 1820 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Jungermannia fastigiata Lehm. 1829 = Riccardia fastigiata (Lehm.) Trevis. 1877. 

Jungermannia filicina Sw. 1788 = Bryopteris filicina (Sw.) Nees 1845 [Dubia et Excludenda]. 

Jungermannia filiformis Sw. 1788 = Omphalanthus filiformis (Sw.) Nees 1845 [Dubia et Excludenda]. 

Jungermannia flava Sw. 1788 = Lejeunea flava (Sw.) Nees 1838. 

Jungermannia floccosa Lehm. et Lindenb. 1833 = Cololejeunea floccosa (Lehm. et Lindenb.) Schiffn. 1898. 

Jungermannia fragrans Moris et De Not. 1839 = Lophocolea fragrans (Moris et De Not.) Gottsche et al. 1845. 

Jungermannia fulvifolia Hook.f. et Taylor 1845 = Radula fulvifolia (Hook.f. et Taylor) Gottsche et al. 1845. 

Jungermannia furcata L. 1753 = Metzgeria furcata (L.) Dumort. 1835. 

Jungermannia fusca Lehm. 1829 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Jungermannia geminiflora Nees 1833 = Omphalanthus filiformis (Sw.) Nees 1845 [Dubia et Excludenda]. 

Jungermannia geminifolia Mitt. 1863 = Syzygiella geminifolia (Mitt.) Steph. 1902. 

Jungermannia gigantea F.Weber 1815 = Pleurozia gigantea (F.Weber) Lindb. 1874. 

Jungermannia grandiflora Lindenb. et Gottsche 1847 = Cryptochila grandiflora (Lindenb. et Gottsche) Grolle 

1971. 

Jungermannia heteromorpha Lehm. 1829 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969.


Jungermannia hirtella F.Weber 1815 = Chandonanthus hirtellus (F.Weber) Mitt. 1873. 

Jungermannia humistrata Hook.f. et Taylor 1845 = Lophocolea humistrata (Hook.f. et Taylor) Gottsche et al. 

1847. 

Jungermannia incerta Gottsche 1882 [Dubia et Excludenda]. 

Jungermannia jamesonii Mont. 1856 = Andrewsianthus jamesonii (Mont.) Váòa 1983. 

Jungermannia javanica Sw. 1781 = Plagiochila javanica (Sw.) Dumort. 1835 [Dubia et Excludenda]. 

Jungermannia juratzkana Limpr. 1876 = Anthelia juratzkana (Limpr.) Trevis. 1877. 

Jungermannia kiaeri Austin 1875 = Cephaloziella kiaeri (Austin) Douin 1920. 

Jungermannia laevifolia Hook.f. et Taylor 1844 = Lepidozia laevifolia (Hook.f. et Taylor) Gottsche et al. 1845. 

Jungermannia leioclada Hook.f. et Taylor 1845 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838. 

Jungermannia leucoxantha Lehm. 1829 = Fossombronia leucoxantha (Lehm.) Gottsche et al. 1845. 

Jungermannia limbata Hook.f. et Taylor 1845 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Jungermannia lindenbergiana Lehm. 1829 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864. 

Jungermannia lineata Lehm. et Lindenb. 1832 = Cystolejeunea lineata (Lehm. et Lindenb.) A.Evans 1906 


Jungermannia longa Nees 1831 = Bazzania longa (Nees) Trevis. 1877 [Dubia et Excludenda]. 

Jungermannia lucida Spreng. ex Lehm. 1833 = Lophocolea lucida (Spreng. ex Lehm.) Mont. 1839. 

Jungermannia lunulata F.Weber 1815 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893. 

Jungermannia lutescens Lehm. et Lindenb. 1832 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Jungermannia lyellii Hook. 1816 = Pallavicinia lyellii (Hook.) Carruth. 1865. 

Jungermannia macrocalyx Mont. 1843 = Syzygiella macrocalyx (Mont.) Spruce 1876 [Dubia et Excludenda]. 

Jungermannia marginalis Hook.f. et Taylor 1845 = Harpalejeunea marginalis (Hook.f. et Taylor) Steph. 1913. 

Jungermannia marginata Lehm. et Lindenb. 1833 = Cololejeunea marginata (Lehm. et Lindenb.) Schiffn. 1898. 

Jungermannia mauritiana Nees 1831 = Plagiochila repanda (Schwägr.) Lindenb. 1840. 

Jungermannia microphylla var. β Hook. 1818 = Lepidozia africana Steph. 1922. 

Jungermannia microscopica Taylor 1836 = Aphanolejeunea microscopica (Taylor) A.Evans 1911. 

Jungermannia microscypha Hook.f. et Taylor 1845 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 

1907. 

Jungermannia mildbraedii Steph. RWA(2), SAF(2), TAN(2), UGA(2). 

Jungermannia minuta Schreb. ex Cranz 1770 = Anastrophyllum minutum (Schreb.) R.M.Schust. 1949. 

Jungermannia multifida L. 1753 = Riccardia multifida (L.) Gray 1821. 

Jungermannia multifida var. pinnatifida F.Weber 1815 = Riccardia pinnatifida (F.Weber) Carruth. 1865 [Dubia 

et Excludenda]. 

Jungermannia muricata Lehm. 1829 = Lophocolea muricata (Lehm.) Nees 1845. 

Jungermannia nemoides Hook.f. et Taylor 1845 = Kurzia nemoides (Hook.f. et Taylor) Grolle 1964. 

Jungermannia nitida F.Weber 1815 = Bazzania nitida (F.Weber) Grolle 1960. 

Jungermannia nodulosa Reinw. et al. 1824 = Frullania nodulosa (Reinw. et al.) Nees 1845. 

Jungermannia obtusata Hook.f. et Taylor 1845 = Anastrophyllum subcomplicatum (Lehm. et Lindenb.) Steph. 

1893. 

Jungermannia ochroleuca Spreng. 1827 = Lepicolea ochroleuca (Spreng.) Spruce 1884. 

Jungermannia onraedtii Váòa MDG(1), REU(1). 

Jungermannia palmata Hedw. 1784 = Riccardia palmata (Hedw.) Carruth. 1865 [Dubia et Excludenda]. 

Jungermannia pectinata Willd. ex F.Weber 1815, nom. inval. = Plagiochila pectinata Willd. ex Lindenb. 1839. 

Jungermannia pellucida Meissn. ex Spreng. 1827 = Diplasiolejeunea pellucida (Meissn. ex Spreng.) Schiffn. 

1893 [Dubia et Excludenda]. 

Jungermannia perloi (Gola) Váòa KEN(2), REU(1). 

Jungermannia phyllorhiza Nees 1833 = Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst. 1986. 

Jungermannia piligera Nees 1824 = Anastrophyllum piligerum (Nees) Steph. 1893. 

Jungermannia pinguis L. 1753 = Aneura pinguis (L.) Dumort. 1831. 

Jungermannia pocsii Váòa KEN(2), SAF(2), TAN(2). 

Jungermannia podophylla Thunb. 1800 = Symphyogyna podophylla (Thunb.) Mont. et Nees 1846. 

Jungermannia polycarpa Nees 1833 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Jungermannia praerupta Reinw. et al. 1824 = Bazzania praerupta (Reinw. et al.) Trevis. 1877. 

Jungermannia procumbens Hook.f. et Taylor 1845 = Symphyogyna brasiliensis Nees et Mont. 1836. 

Jungermannia pterota Hook.f. et Taylor 1845 = Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle 1988. 

Jungermannia pumila With. LES(75), SAF(75), TAN(2). 

Jungermannia purpurascens (Steph.) Besch. 1895 = Jamesoniella purpurascens Steph. 1891.


Jungermannia pusilla L. 1753 = Fossombronia pusilla (L.) Nees 1838. 

Jungermannia pusilla Lehm. = Fossombronia crispa Nees 1846. 

Jungermannia refracta Brid. ex Gottsche 1857, nom. inval. = Plagiochila paucidentata Mont. et Gottsche 1856. 

Jungermannia rehmannii Steph. 1892 = Jamesoniella purpurascens (Steph.) Steph. 1901. 

Jungermannia renauldii Steph. MDG(1), REU(1), UGA(2,96). 

Jungermannia repanda Schwägr. 1814 = Plagiochila repanda (Schwägr.) Lindenb. 1840. 

Jungermannia reptans L. 1753 = Lepidozia reptans (L.) Dumort. 1835. 

Jungermannia rotalis Hook.f. et Taylor 1845 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 

1997 (55). 

Jungermannia rotundistipula Lindenb. ex Lehm. 1829 = Leucolejeunea rotundistipula (Lindenb. ex Lehm.) 

A.Evans 1907. 

Jungermannia sarmentosa Lehm. et Lindenb. 1835 = Plagiochila sarmentosa (Lehm. et Lindenb.) Lindenb. 

1844. 

Jungermannia scariosa Lehm. 1829 = Gongylanthus scariosus (Lehm.) Steph. 1906. 

Jungermannia schizopleura Spruce 1885 = Gottschelia schizopleura (Spruce) Grolle 1968. 

Jungermannia semiteres Lehm. 1829 = Lophocolea semiteres (Lehm.) Mitt. 1877. 

Jungermannia serrulata Sw. 1788 = Isotachis serrulata (Sw.) Gottsche 1864 [Dubia et Excludenda]. 

Jungermannia simplex Sw. 1788 = Plagiochila simplex (Sw.) Lindenb. 1840 [Dubia et Excludenda]. 

Jungermannia spathulistipa Reinw. et al. 1824 = Thysananthus spathulistipus (Reinw. et al.) Lindenb. 1845. 

Jungermannia sphaerocarpa Hook. CMN(2), LES(75), MDG(1), MLW(2), REU(1), RWA(2), SAF(75), 

TAN(2), UGA(2,42), ZAI(2), ZIM(2). 

Jungermannia sphaerophora Lehm. et Lindenb. 1833 = Lopholejeunea sphaerophora (Lehm. et Lindenb.) 

Steph. 1912. 

Jungermannia sphagnoides Schwägr. 1814 = Gottschea sphagnoides (Schwägr.) Lindb. 1870. 

Jungermannia sphagnoides sensu Richard ex Hook. 1818, hom. illeg. [non Schwägr. 1814] = Pleurozia gigantea 

(F.Weber) Lindb. 1874. 

Jungermannia splendens Lehm. et Lindenb. 1832 = Heteroscyphus splendens (Lehm. et Lindenb.) Grolle 1984. 

Jungermannia squarrosa Reinw. et al. 1824, hom. illeg. [non Hook 1818] = Frullania ericoides (Nees) Mont. 

1839. 

Jungermannia stolonifera (Steph.) Steph. 1901, hom. illeg. [non Sw. 1788] = J. borgenii Gottsche ex Pearson 

1893. 

Jungermannia stolonifera Sieber ex Nees 1831, hom. illeg. [non Sw. 1788] = Bazzania longa (Nees) Trevis. 

1877 [Dubia et Excludenda]. 

Jungermannia stolonifera Sw. 1788 = Bazzania stolonifera (Sw.) Trevis. 1877 [Dubia et Excludenda]. 

Jungermannia striata Lehm. et Lindenb. 1832 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838. 

Jungermannia subaequifolia Nees et Mont. 1844 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838. 

Jungermannia subcomplicata Lehm. et Lindenb. 1838 = Anastrophyllum subcomplicatum (Lehm. et Lindenb.) 

Steph. 1901. 

Jungermannia subfusca Nees 1830 = Lopholejeunea subfusca (Nees) Schiffn. 1897. 

Jungermannia subtilissima (Schiffn.) Grolle. A record from the Cape Verde Islands (125) is based on a 

misidentification (37). 

Jungermannia superba Nees ex Spreng. 1827 = Plagiochila superba (Nees ex Spreng.) Mont. et Nees 1839 


Jungermannia tabularis Spreng. 1827 = Lejeunea tabularis (Spreng.) Gottsche et al. 1845. 

Jungermannia tamarisci L. 1753 = Frullania tamarisci (L.) Dumort. 1835. 

Jungermannia tenuis Reinw. et al. 1824, hom. illeg. [non Ehrh. 1789] = Drepanolejeunea angustifolia (Mitt.) 

Grolle 1965. 

Jungermannia tenuissima Lehm. et Lindenb. 1845 = Cephaloziella tenuissima (Lehm. et Lindenb.) Steph. 1893. 

Jungermannia thouarsii Hook. 1818 = Gottschea sphagnoides (Schwägr.) Lindb. 1870. 

Jungermannia tortuosa Lehm. et Lindenb. 1832 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893. 

Jungermannia trichophylla L. 1753 = Blepharostoma trichophyllum (L.) Dumort. 1835. 

Jungermannia trifaria Reinw. et al. 1824 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964. 

Jungermannia trinervis Lehm. 1835 = Frullania trinervis (Lehm.) Drège 1843. 

Jungermannia trinitensis Lehm. et Lindenb. 1833 = Bryopteris filicina (Sw.) Nees 1845 [Dubia et Excludenda]. 

Jungermannia tuberculata Lehm. 1835 = Frullania ericoides (Nees) Mont. 1839. 

Jungermannia ulicina Taylor 1841 = Microlejeunea ulicina (Taylor) A.Evans 1902. 

Jungermannia unciformis Hook.f. et Taylor 1844 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864.


Jungermannia uncinata F.Weber 1815 = Isotachis aubertii (Schwägr.) Mitt. 1886. 

Jungermannia vermicularis Lehm. 1829 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Jungermannia verrucosa (Steph.) Steph. 1901 = J. dusenii (Steph.) Steph. 1901. 

Jungermannia violacea Ach. 1805 = Metzgeria violacea (Ach.) Dumort. 1835. 

Jungermannia vitrea Nees 1830 = Leptolejeunea vitrea (Nees) Schiffn. 1893. 

Jungermannia volkensii Steph. 1917 = J. borgenii Gottsche ex Pearson 1893. 

Jungermannia xanthocarpa Lehm. et Lindenb. 1833 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans 

1907. 

Kantius bidentulus (F.Weber) Trevis. 1877 [Kantia auct.] = Calypogeia bidentula (F.Weber) Nees 1845. 

Kantius heteromorphus (Lehm.) Steph. 1888 [Kantia auct.] = Alobiellopsis heteromorpha (Lehm.) R.M.Schust 

1969. 

Kantius microstipulus Steph. 1895 [Kantia auct.] = Calypogeia microstipula (Steph.) Steph. 1908. 

Kantius muelleriana Schiffn. 1900 [Kantia auct.] = Calypogeia muelleriana (Schiffn.) Müll.Frib. 1908. 

Kurzia capillaris ssp. capillaris (Sw.) Grolle MDG(4), MLW(2,92), RWA(2), SAF(2), SEY(4), TAN(2). 

Kurzia capillaris ssp. capillaris var. verrucosa (Steph.) Pócs TAN(2). 

Kurzia capillaris ssp. stephanii (Renauld ex Steph.) Pócs MAU(1), MDG(1,4), MLW(2), REU(1,4), 

TAN(2). 

Kurzia irregularis (Steph.) Grolle RWA(2), SAF(2), TAN(2), UGA(2), ZAI(2). 

Kurzia nemoides (Hook.f. et Taylor) Grolle STH(2,49). 

Kurzia stephanii (Renauld ex Steph.) Grolle 1964 = Kurzia capillaris ssp. stephanii (Renauld ex Steph.) Pócs 

1984. 

Kurzia tabularis (Steph.) Grolle 1964 = K. capillaris (Sw.) Grolle 1964. 

Kymatocalyx africanus Váòa et Wigginton MLW(74), TAN(74). 

Kymatocalyx dominicensis (Spruce) Váòa MDG(74). 

Kymatocalyx madagascariensis (Steph.) Gradst. et Váòa COM(4,74), MAU(1,4,74), MDG(1,4,74), 

REU(1,4,74), TAN(2). 

Leioscyphus borbonicus Steph. 1892 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963. 

Leioscyphus gottscheanus (Lindenb. ex Lehm.) Steph. 1906 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Leioscyphus infuscatus Mitt. 1886 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963. 

Leioscyphus iversenii Pearson 1887 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Leioscyphus motleyi Mitt. 1886 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963. 

Leioscyphus repens Mitt. 1863, hom. illeg. [non Mitten 1854] = Leptoscyphus infuscatus (Mitt.) E.W.Jones 

1963. 

Lejeunea abbreviata Mitt. 1863 = Archilejeunea abbreviata (Mitt.) Vanden Berghen 1951. 

Lejeunea abnormis (Gottsche) Steph. 1891 = Phragmicoma abnormis Gottsche 1882 [Dubia et Excludenda]. 

Lejeunea abortiva Mitt. 1879 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898. 

Lejeunea abyssinica (Gola) Cufod. ETH(2). 

Lejeunea acuta Mitt. ANN(2,33), BIO(2,33,72), BUR(2), CMN(2), COM(1), KEN(2), RIM(10), 

RWA(2), SAO(2), TAN(2), UGA(50), ZAI(667). 

Lejeunea acutiloba (Hook.f. et Taylor) Gottsche et al. 1845 = Marchesinia brachiata (Sw.) Schiffn. 1893. 

Lejeunea adhaesiva Mitt. 1886 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963. 

Lejeunea aethiopica E.W.Jones ETH(2). 

Lejeunea africana (Steph.) Steph. 1891 = Microlejeunea africana Steph. 1888. 

Lejeunea alata (Steph.) Steph. 1894, hom. illeg. [non Gottsche 1845] = Archilejeunea alata Steph. 1893. 

Lejeunea alata Gottsche COM(1), MAS(1), MAU(1), MDG(1), REU(1), SEY(1), TAN(2). 

Lejeunea albicans Nees 1845 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Lejeunea albifolia Taylor 1845 = Diplasiolejeunea albifolia (Taylor) E.W.Jones 1954. 

Lejeunea alata var. patriciae Pócs MDG(709). 

Lejeunea aloba Sande Lac. ZAI(2). 

Lejeunea alternifolia Mitt. 1873 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Lejeunea amaniensis E.W.Jones CMN(2), MDG(1), RWA(2), TAN(2). 

Lejeunea amentulifera Steph. UGA/ZAI(2). 

Lejeunea amplectens (Steph.) Steph. 1890 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Lejeunea angulifolia Mitt. ROD(1). 

Lejeunea angustifolia Mitt. 1861 = Drepanolejeunea angustifolia (Mitt.) Grolle 1965 [Dubia et Excludenda].


Lejeunea arnelliana R.M.Schust. 1965 = L. villaumei (Steph.) Grolle 1977. 

Lejeunea ascensionis (Hook.f. et Taylor) Gottsche et al. 1845 = Cheilolejeunea ascensionis (Hook.f. et Taylor) 

Grolle 1988. 

Lejeunea asperifolia Steph. ZAI(2). 

Lejeunea astroidea Mitt. 1860 = Leptolejeunea astroidea (Mitt.) Steph. 1913. 

Lejeunea atra Mitt. 1886 = Acanthocoleus aberrans var. laevis Gradst. 1994. 

Lejeunea autoica R.M.Schust. BIO(2,33), CMN(2), GHA(2), TAN(2). 

Lejeunea balfourii Mitt. 1879 = Stictolejeunea balfourii (Mitt.) E.W.Jones 1976. 

Lejeunea belangeri Gottsche ex Ångstr. 1874, nom. inval. = Ceratolejeunea belangeriana (Gottsche) Steph. 

1913. 

Lejeunea belangeriana Gottsche 1845 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913. 

Lejeunea borbonica Hampe et Gottsche 1895, nom. inval. = Lopholejeunea borbonica Steph. 1896. 

Lejeunea borgenii Steph. 1890 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Lejeunea brachytoma Gottsche 1882 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964. 

Lejeunea brenanii E.W.Jones CMN(2), GHA(2), RIM(10), SIE(2). 

Lejeunea breutelii (Steph.) Steph. 1901, hom. illeg. [non Gottsche 1845] = L. caespitosa Lindenb. 1845. 

Lejeunea brevifissa Gottsche 1882 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963. 

Lejeunea brittoniae (A.Evans) Grolle 1995 = L. phyllobola Nees et Mont. 1842 (168). 

Lejeunea byssoides Gottsche 1882 = Microlejeunea byssoides (Gottsche) Pearson 1892 [Dubia et Excludenda]. 

Lejeunea caespitosa Lindenb. ANG(2), ANN(33), BIO(2,33), BUR(2), CAF(2), CMN(2), COM(1), 

CON(2), CVI(2,37), GHA(2), IVO(2,53), KEN(2,71), MAU(1), MLW(563), NGA(2), PRI(2), REU(1), 

RIM(10), RWA(2), SAF(2), SAO(2), SEY(1,4,108), SIE(2), TAN(2), UGA(2,42,96), ZAI(2), ZIM(2). 

Lejeunea calyptrifolia (Hook.) Spreng. 1827, hom. illeg. [non Dumort. 1822] = Colura calyptrifolia (Hook.) 

Dumort. 1835. 

Lejeunea calyptrifolia Dumort. 1822, nom. nud. = Colura calyptrifolia (Hook.) Dumort. 1835. 

Lejeunea camerunensis (Steph.) E.W.Jones ANG(2), ANN(2,33), CAF(2), CMN(2), GHA(2), GUI(2), 

IVO(2), KEN(2), MAU(1), MDG(1), MLW(2,563), MOZ(2), NGA(2), REU(1), SEY(1,4), SIE(2), 

TAN(2), UGA(96), ZIM(2). 

Lejeunea cancellata Nees et Mont. 1842 [Dubia et Excludenda]. 

Lejeunea cantabrigiensis E.W.Jones CMN(2). 

Lejeunea capensis (S.W.Arnell) R.M.Schust. 1963, hom. illeg. [non Gottsche 1845] = L. villaumei (Steph.) 

Grolle 1977. 

Lejeunea capensis Gottsche KEN(2), LES(75), MDG(1), MLW(2), RWA(2), SAF(2), TAN(2), ZAI(2). 

Lejeunea capulata Taylor 1846 = Drepanolejeunea capulata (Taylor) Steph. 1913. 

Lejeunea cardiantha Gottsche 1882 = Prionolejeunea grata (Gottsche) Schiffn. 1893. 

Lejeunea cardiocarpa Mont. 1842 = Cololejeunea cardiocarpa (Mont.) A.Evans 1902. 

Lejeunea cardiophylla Gottsche 1882 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914. 

Lejeunea cavifolia (Ehrh.) Lindb. SOC(2). 

Lejeunea cavifolia Steph. 1886, hom. illeg. [non (Ehrh.) Lindb. 1871] = Diplasiolejeunea cavifolia Steph. 1895. 

Lejeunea chrysophylla (Lehm.) Gottsche et al. 1845 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988. 

Lejeunea compressistipula (Pearson) Demaret 1942, perhaps = L. confusa E.W.Jones 1972. 

Lejeunea confertissima (Steph.) Steph. 1892 = Acrolejeunea emergens var. confertissima (Steph.) Gradst. 1975. 

Lejeunea conformis Nees et Mont. 1843 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914. 

Lejeunea confusa E.W.Jones CAF(2), CMN(2), COM(1), GAB(2), GHA(2), KEN(2), MLW(563), 

NGA(2), RIM(10), RWA(2), SEY(1), SIE(2), TAN(2), UGA(50,96), ZAI(2). 

Lejeunea contigua (Nees) Mont. 1840 = Pycnolejeunea contigua (Nees) Grolle 1979. 

Lejeunea convexa S.W.Arnell 1953 = Cheilolejeunea convexa (S.W.Arnell) S.W.Arnell 1955. 

Lejeunea corbieri Steph. 1915, perhaps = L. ibadana A.J.Harr. et E.W.Jones 1982. 

Lejeunea cornuta (Lindenb.) Nees 1838 = Ceratolejeunea cornuta (Lindenb.) Steph. 1895 [Dubia et 

Excludenda]. 

Lejeunea cornutissima Steph. 1892 = Ceratolejeunea diversicornua (Steph.) Steph. 1913. 

Lejeunea crenatiflora (Steph.) Steph. 1891 ("crenatifolia") = Cololejeunea obliqua (Nees et Mont.) Schiffn. 

1897. 

Lejeunea cristata (Steph.) Steph. 1891 = Drepanolejeunea cultrella (Mitt.) Steph. 1913. 

Lejeunea cucullata (Reinw. et al.) Nees 1838 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Lejeunea cucullata var. maior Nees 1845 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995.


Lejeunea cucullata var. parasita Gottsche et al. 1843 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Lejeunea cultrella Mitt. 1863 = Drepanolejeunea cultrella (Mitt.) Steph. 1913. 

Lejeunea cuneata (Lehm. et Lindenb.) Gottsche et al. 1845 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 

1947. 

Lejeunea cuneifolia (Steph.) Steph. Nov-Dec 1892 = Cololejeunea cuneifolia Steph. Jul-Aug 1892. 

Lejeunea cupulirostris Augier CMN(2). 

Lejeunea cyathearum E.W.Jones COM(1,4), MLW(2), RWA(2), TAN(2), ZAI(2). 

Lejeunea decursiva Sande Lac. 1855 = Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. 1963. 

Lejeunea delessertii Nees et Mont. 1843 = Lepidolejeunea delessertii (Nees et Mont.) Grolle 1984. 

Lejeunea deplanata Mitt. 1886, hom. illeg. [non Nees 1845] = Prionolejeunea grata (Gottsche) Schiffn. 1893. 

Lejeunea digitalis Mitt. 1886 = Colura digitalis (Mitt.) Steph. 1916. 

Lejeunea dipterocarpa E.W.Jones CMN(2), NGA(2). 

Lejeunea dispar (Ast) R.M.Schust. 1962 = Microlejeunea dispar Ast 1959. 

Lejeunea diversicornua (Steph.) Steph. 1891 = Ceratolejeunea diversicornua (Steph.) Steph. 1913. 

Lejeunea duncaniae (Sim) S.W.Arnell SAF(2). 

Lejeunea dusenii Steph. 1891 = L. ramosissima Steph. 1886. 

Lejeunea dusenii Steph. 1892, hom. illeg. = Colura dusenii (Steph.) Steph. 1916. 

Lejeunea ecarinata (Steph.) Steph. MDG(1), REU(1). 

Lejeunea eckloniana Lindenb. ANG(2), CMN(2), CON(2), CVI(2,37), DJI(2), GHA(2), GUI(2), 

KEN(2,21,71), LES(75), MAU(1), MDG(1), MLW(2,563), NGA(2,127), REU(1), ROD(1), RWA(2), 

SAF(2,75), SIE(2), TAN(2), TOG(2,101), UGA(2,96), ZAI(2), ZAM(2), ZIM(2). 

Lejeunea ecklonii Lindenb. ex Steph. 1890, nom. inval. = L. eckloniana Lindenb. 1845. 

Lejeunea elegans (Steph.) Steph. Nov-Dec 1891, hom. illeg. [non Gottsche 1845] = Cololejeunea elegans Steph. 

1891. 

Lejeunea elliptica (Lehm. et Lindenb.) Mont. 1840 = Leptolejeunea elliptica (Lehm. et Lindenb.) Schiffn. 1893. 

Lejeunea elobulata (Sim) S.W.Arnell, nom. inval. SAF(2). 

Lejeunea emergens (Mitt.) Steph. 1894 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Lejeunea epiphylla Mitt. 1861 = Leptolejeunea epiphylla (Mitt.) Steph. 1913. 

Lejeunea epiphyta (Steph.) Steph. 1891, hom. illeg. [non Gottsche 1845] = Taxilejeunea conformis (Nees et 

Mont.) Steph. 1914. 

Lejeunea eplicata Steph. 1915 = L. ramosissima Steph. 1887. 

Lejeunea erectifolia Spruce 1884 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Lejeunea eulopha Taylor 1846 = Lopholejeunea eulopha (Taylor) Schiffn. 1893. 

Lejeunea excavata Mitt. 1860 = Marchesinia excavata (Mitt.) Schiffn. 1893. 

Lejeunea expansa (Steph.) Steph. 1915 = L. tabularis (Spreng.) Gottsche et al. 1845. 

Lejeunea filicaulis (Steph.) Steph. Nov-Dec 1892 = Cololejeunea hildebrandii (Austin) Steph. 1897. 

Lejeunea filiformis (Sw.) Nees ex Mont. 1839 = Omphalanthus filiformis (Sw.) Nees 1845 [Dubia et 

Excludenda]. 

Lejeunea fimbriata Gottsche 1880 = Lopholejeunea eulopha (Taylor) Schiffn. 1893. 

Lejeunea flava (Sw.) Nees CAF(2), COM(1), CON(2), CVI(37), GHA(2), KEN(71), MDG(1), RWA(2), 

SEY(1,4,108), SIE(2), TAN(2), TOG(101), UGA(2,96), ZAI(2). 

Lejeunea flava agg. BIO(72), ETH(2), KEN(2), SAO(2), ZAI(2), ZAM(2). 

Lejeunea flava ssp. tabularis (Spreng.) S.W.Arnell 1963 = L. tabularis (Spreng.) Gottsche et al. 1845. 

Lejeunea flava var. convexiuscula Pearson 1886 = L. tabularis (Spreng.) Gottsche et al. 1845. 

Lejeunea flavovirens Ångstr. COM(1), MAU(1), MDG(1), REU(1), RWA(2), SIE(2), TAN(2), 

UGA(2,50), ZAI(2). 

Lejeunea fulva (Gottsche) Steph. 1890, hom. illeg. [non Spruce 1884] = Schiffneriolejeunea pappeana (Nees) 

Gradst. 1974. 

Lejeunea furva Mitt. 1878 = L. eckloniana Lindenb. 1845. 

Lejeunea fusca (Lehm.) Drège 1843 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Lejeunea galliotii Steph. 1915, probably = L. eckloniana Lindenb. 1845. 

Lejeunea gaudichaudii (Gottsche) Steph. 1894 [non Lindenb. 1845] = Bryopteris gaudichaudii Gottsche 1857. 

Lejeunea gracillima Mitt. 1861 = Microlejeunea ulicina (Taylor) A.Evans 1902. 

Lejeunea grandistipula Steph. 1886 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964. 

Lejeunea grata Gottsche 1882 = Prionolejeunea grata (Gottsche) Schiffn. 1893. 

Lejeunea grossecristata (Steph.) E.W.Jones ANN(2,33), BIO(2,33), CMN(2), GHA(2), IVO(2), KEN(2),


NGA(2), RWA(2), SAO(2), SIE(2), TAN(2), UGA(2,96), ZAI(2). 

Lejeunea hamatifolia (Hook.) Spreng. 1893 = Drepanolejeunea hamatifolia (Hook.) Schiffn. 1895. 

Lejeunea hanningtonii Mitt. 1886 = Caudalejeunea hanningtonii (Mitt.) Schiffn. 1893. 

Lejeunea haskarliana (Gottsche) Spruce 1884, comb. illeg. = Schiffneriolejeunea tumida (Nees) Gradst. 1974. 

Lejeunea haskarliana Lehm. & Lindenb. in Lehmann 1844 = Cololejeunea haskarliana (Lehm. & Lindenb.) 

Schiffn. 1898. 

Lejeunea helenae Pearson ANG(2), ETH(2), KEN(2), MLW(2), RWA(2), SAF(2,75), TAN(2), 

UGA(2,96), ZAI(2), ZIM(2). 

Lejeunea hepaticola (Steph.) Steph. KEN(2), MLW(563), TAN(2), UGA(115), ZAI(2). 

Lejeunea hildebrandii Austin 1876 = Cololejeunea hildebrandii (Austin) Steph. 1897. 

Lejeunea holtii Spruce ANN(2), UGA(2). 

Lejeunea hyalinomarginata Nees ex Mont. 1842 = Cololejeunea marginata (Lehm. et Lindenb.) Schiffn. 1898. 

Lejeunea ibadana A.J.Harr. et E.W.Jones CAF(2), GHA(2), IVO(2), NGA(2), RIM(10), SIE(2), 

UGA(2). 

Lejeunea inflexa (Gottsche) Steph. 1891, hom. illeg. [non Lehm. et Lindenb. 1838] = Phragmicoma inflexa 

Gottsche 1882 [Dubia et Excludenda]. 

Lejeunea intermedia Lindenb. 1845 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898. 

Lejeunea intertexta Lindenb. 1845 = Cheilolejeunea intertexta (Lindenb.) Steph. 1897. 

Lejeunea intorta Besch. et Spruce 1890 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913. 

Lejeunea isomorpha Gottsche 1882 = L. tabularis (Spreng.) Gottsche et al. 1845. However, Jones 1968 (106) 

pointed out that the names L. isomorpha and L. isophylla have been confused. Vanden Berghen 1972 (158) 

considered that plants from southern Africa named as L. isomorpha might often represent L. isophylla. 

Lejeunea isophylla E.W.Jones BUR(2), COM(1), IVO(2), KEN(2), MDG(1), MLW(2,563), MOZ(2), 

RWA(2), SAF(2), SEY(1), TAN(2), UGA(50,96), ZAI(2), ZIM(2). 

Lejeunea julacea Steph. MDG(1). 

Lejeunea jungneri (Steph.) Steph. 1901 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914. 

Lejeunea kamerunensis (Steph.) Vanden Berghen 1972 = Microlejeunea kamerunensis Steph. 1915. 

Lejeunea kilimandjarensis Steph. TAN(2). 

Lejeunea kindbergii Steph. 1892 = Prionolejeunea grata (Gottsche) Schiffn. 1893. 

Lejeunea krakakammae Lindenb. 1845 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963. 

Lejeunea kraussiana Lindenb. 1845 = Diplasiolejeunea kraussiana (Lindenb.) Steph. 1895. 

Lejeunea kurzii (Steph.) Steph. 1890 = Cheilolejeunea intertexta (Lindenb.) Steph. 1897. 

Lejeunea lamacerina (Steph.) Schiffn. CVI(2,37). 

Lejeunea lehmanniana Gottsche 1845 = Caudalejeunea lehmanniana (Gottsche) A.Evans 1907. 

Lejeunea letabaensis S.W.Arnell 1954 = L. rhodesiae (Sim) R.M.Schust. 1962. 

Lejeunea leucosis Besch. et Spruce MDG(1). 

Lejeunea lineata (Lehm. et Lindenb.) Nees 1838 = Cystolejeunea lineata (Lehm. et Lindenb.) A.Evans 1906 


Lejeunea lomana E.W.Jones SIE(2), UGA(115). 

Lejeunea longirostris (Steph.) E.W.Jones 1968, hom. illeg. [non Spruce 1884] = L. grossecristata (Steph.) 

E.W.Jones 1968. 

Lejeunea lunulata (F.Weber) Nees 1845 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893. 

Lejeunea lyratiflora Steph. BUR(2), CMN(2), RWA(2), TAN(2), UGA(50). 

Lejeunea maculata Mitt. 1861 = Leptolejeunea maculata (Mitt.) Schiffn. 1898. 

Lejeunea madagascariensis Gottsche 1882, hom. illeg. [non Nees et Mont. 1836] = Acanthocoleus 

madagascariensis (Steph.) Kruijt 1988. 

Lejeunea madagascariensis Nees et Mont. 1836 = Porella madagascariensis (Nees et Mont.) Trevis. 1877. 

Lejeunea marginata (Lehm. et Lindenb.) Gottsche et al. 1845 = Cololejeunea marginata (Lehm. et Lindenb.) 


Lejeunea mariana Gottsche 1845 = Cheilolejeunea mariana (Gottsche) B.Thiers et Gradst. 1989 [Dubia et 

Excludenda]. 

Lejeunea mascarena Steph. 1890 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913. 

Lejeunea mauritiana Steph. 1890 = Ceratolejeunea belangeriana (Gottsche) Steph. 1913. 

Lejeunea microloba Gola 1920 perhaps = L. eckloniana Lindenb. 1845 (158). 

Lejeunea microlobulata S.W.Arnell 1953 = L. eckloniana Lindenb. 1845. 

Lejeunea microscopica (Taylor) Gottsche et al. 1845 = Aphanolejeunea microscopica (Taylor) A.Evans 1911.


Lejeunea microscypha (Hook.f. et Taylor) Gottsche et al. 1845 = Leucolejeunea xanthocarpa (Lehm. et 

Lindenb.) A.Evans 1907. 

Lejeunea minutissima (Sm.) Spreng. 1827, hom. illeg. [non Dumort. 1822] = Cololejeunea minutissima (Sm.) 


Lejeunea minutissima Dumort. 1822, nom. inval. = Cololejeunea minutissima (Sm.) Schiffn. 1893. 

Lejeunea mitracalyx (Eifrig) Mizut. 1970 = L. alata Gottsche 1845. 

Lejeunea monoica (Bischl.) R.M.Schust. 1980 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Lejeunea montagnei Gottsche 1843 = Cheilolejeunea montagnei (Gottsche) R.M.Schust. 1963. 

Lejeunea multilacera (Steph.) Steph. Dec 1890 = Lopholejeunea multilacera Steph. Nov 1890. 

Lejeunea myriocarpa Nees et Mont. 1842 = Cololejeunea minutissima ssp. myriocarpa (Nees et Mont.) 


Lejeunea nigra Steph. 1891 = Mastigolejeunea nigra (Steph.) Steph. 1895. 

Lejeunea nigricans Lindenb. 1845 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898. 

Lejeunea nobilis Gottsche 1882 = Archilejeunea nobilis (Gottsche) Steph. 1911. 

Lejeunea obliqua Nees et Mont. 1843 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897. 

Lejeunea oblongistipula Gottsche 1882 = Microlejeunea oblongistipula (Gottsche) Steph. 1915. 

Lejeunea obtusa (Steph.) Steph. 1891 = Colura digitalis (Mitt.) Steph. 1916. 

Lejeunea obtusata Gottsche GHA(2), MAU(1), MDG(1), SIE(2), TAN(2), UGA(2,96). 

Lejeunea obtusistipula (Steph.) E.W.Jones GUI(2), SIE(2). 

Lejeunea occulta Steph. 1888 = Schiffneriolejeunea occulta (Steph.) Gradst. 1974. 

Lejeunea ocellifera (S.W.Arnell) R.M.Schust. 1955, nom. inval. = Microlejeunea africana Steph. 1888. 

Lejeunea okomuensis E.W.Jones CMN(2), IVO(2), NGA(2), UGA(42). 

Lejeunea pallidissima Gola ZAI(2). 

Lejeunea papilionacea Steph. CMN(2,670), COM(1,670), GHA(2,670), GUI(670), MDG(1,670), 

REU(670), RIM(10), SIE(2,670). 

Lejeunea pappeana (Nees) Mitt. 1863 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Lejeunea papulosa Gottsche 1882 = Microlejeunea papulosa (Gottsche) Pearson 1892. 

Lejeunea parallela Schiffn. 1890 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Lejeunea parisii Steph. 1915 = L. caespitosa Lindenb. 1845. 

Lejeunea parviloba (Steph.) Steph. 1890, hom. illeg. [non Ångstr. 1877] = Schiffneriolejeunea parviloba 

(Steph.) Gradst. 1974. 

Lejeunea pellucida (Meissn. ex Spreng.) Gottsche et al. 1845 = Diplasiolejeunea pellucida (Meissn. ex Spreng.) 

Schiffn. 1893 [Dubia et Excludenda]. 

Lejeunea perforata Schiffn. 1886 = Drepanolejeunea trematodes (Nees) Bischl. 1968. 

Lejeunea phyllobola Nees et Mont. COM(1), KEN(2), MDG(4), SAF(2), SEY(4), SIE(2), TAN(2), 

ZAI(2). 

Lejeunea physaefolia Gottsche 1882 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913. 

Lejeunea planiuscula Mitt. 1861 = Archilejeunea planiuscula (Mitt.) Steph. 1911. 

Lejeunea platyneura Spruce 1884 = Cololejeunea platyneura (Spruce) A.Evans 1902. 

Lejeunea pluriplicata Pearson 1887 = Cheilolejeunea pluriplicata (Pearson) R.M.Schust. 1980. 

Lejeunea polyandra Gola 1920 = L. eckloniana Lindenb. 1845. 

Lejeunea pterota (Hook.f. et Taylor) Gottsche et al. 1845 = Cheilolejeunea ascensionis (Hook.f. et Taylor) 

Grolle 1988. 

Lejeunea pulcherrima Steph. 1886 = L. acuta Mitt. 1863. 

Lejeunea punctata Gottsche 1882 = Cololejeunea punctata (Gottsche) Pearson 1892 [Dubia et Excludenda]. 

Lejeunea pycnoclada (Taylor) Mitt. 1861 = Acrolejeunea pycnoclada (Taylor) Schiffn. 1893. 

Lejeunea quintasii Steph. 1891 = Leptolejeunea epiphylla (Mitt.) Steph. 1913. 

Lejeunea ramosissima Steph. ANN(2,33), BIO(2,33), CMN(2), CON(2), GUI(2), PRI(2), REU(1), 

RWA(2), SAO(2), SEY(1), SIE(2), TAN(2), TOG(2), UGA(96), ZAI(2). 

Lejeunea renauldii Renauld et Cardot 1891, hom. illeg. = Acrolejeunea aulacophora (Mont.) Steph. 1895. 

Lejeunea renauldii Steph. Dec 1890 = L. alata Gottsche 1845. 

Lejeunea repanda Schiffn 1886 = Leptolejeunea maculata (Mitt.) Schiffn. 1898. 

Lejeunea rhodesiae (Sim) R.M.Schust. ANG(2), CAF(2), CON(2), KEN(21), LES(75), MLW(2), 

RWA(2), SAF(2), TAN(2), UGA(2,50), ZAI(2), ZAM(2), ZIM(2). 

Lejeunea rodriguezii (Steph.) Steph. MAU(1), MDG(1), REU(1).


Lejeunea rotalis (Hook.f. et Taylor) Gottsche et al. 1845 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et 


Lejeunea rotundistipula (Lindenb. ex Lehm.) Mont. 1842 = Leucolejeunea rotundistipula (Lindenb. ex Lehm.) 

A.Evans 1907. 

Lejeunea rufescens Lindenb. 1845 = Cheilolejeunea rufescens (Lindenb.) Grolle 1982. 

Lejeunea rutenbergiana Gottsche 1882 = Prionolejeunea grata (Gottsche) Schiffn. 1893. 

Lejeunea rutenbergii Gottsche ex Steph. 1891, nom. inval. = Prionolejeunea grata (Gottsche) Schiffn. 1893. 

Lejeunea saccatiloba (Steph.) Steph. Nov-Dec 1891 = L. caespitosa Lindenb. 1845. Jones 1972 (157) stated that 

most plants Stephani named as L. saccatiloba are L. confusa, but that the type is L. caespitosa. 

Lejeunea sagraeana (Mont.) Gottsche et al. 1845 = Lopholejeunea subfusca (Nees) Schiffn. 1897. 

Lejeunea scabriflora Gottsche 1882, nom. inval. = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897. 

Lejeunea securifolia Gottsche 1882 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913. 

Lejeunea serpentina Mitt. 1861 = Cheilolejeunea serpentina (Mitt.) Mizut. 1963. 

Lejeunea serrula Mitt. 1886 = Prionolejeunea grata (Gottsche) Schiffn. 1893. 

Lejeunea setacea (Steph.) Steph. BEN(22), BUR(2), CAF(2), CMN(2), COM(1), CON(2), GHA(2), 

IVO(2), KEN(2), NGA(2), PRI(2), SAF(2), SAO(2), SIE(2), ZAI(2), ZAM(2). 

Lejeunea sieberiana Gottsche 1845 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893. 

Lejeunea sikorae (Steph.) Steph. 1894 = Taxilejeunea sikorae Steph. 1893. 

Lejeunea silvestris Gottsche 1882 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976. 

Lejeunea sinuata Mitt. 1886 = Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph. 1913. 

Lejeunea solitaria Gottsche 1882 = Harpalejeunea solitaria (Gottsche) Steph. 1913 [Dubia et Excludenda]. 

Lejeunea spathulistipa (Reinw. et al.) Mitt. 1861 = Thysananthus spathulistipus (Reinw. et al.) Lindenb. 1845. 

Lejeunea sphaerophora (Lehm. et Lindenb.) Gottsche et al. 1845 = Lopholejeunea sphaerophora (Lehm. et 

Lindenb.) Steph. 1912. 

Lejeunea squarrosa (Steph.) Steph. TAN(2). 

Lejeunea stephanii Spruce ex Steph. 1892, nom. inval. = Caudalejeunea reniloba (Gottsche) Steph. 1912 [Dubia 

et Excludenda]. 

Lejeunea striata (Lehm. et Lindenb.) Steph. 1890 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838. 

Lejeunea subciliata Mitt. 1878 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947. 

Lejeunea subfusca (Nees) Nees et Mont. 1836 = Lopholejeunea subfusca (Nees) Schiffn. 1897. 

Lejeunea subinermis (Steph.) Steph. 1901 = Lopholejeunea nigricans (Lindenb.) Schiffn. 1898. 

Lejeunea subintegra (Hook.f. et Taylor) Gottsche et al. 1845 = Clasmatocolea vermicularis (Lehm.) Grolle 

1960. 

Lejeunea surrepens Mitt. 1878 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976. 

Lejeunea tabularis (Spreng.) Gottsche et al. BIO(2,33), BUR(2), CAF(2), CMN(2), COM(1), CON(2), 

GUI(2), IVO(2), KEN(2), LES(75), MAU(108), MDG(1), MLW(2,563), MOZ(2), NGA(2), PRI(2), 

RWA(2), SAF(2,75), TAN(2), UGA(2,42,96), ZAI(2). 

Lejeunea tenuis (Nees) Gottsche et al. 1845 [non (Ehrh.) Gottsche et al. 1845] = Drepanolejeunea angustifolia 

(Mitt.) Grolle 1965 [Dubia et Excludenda]. 

Lejeunea thomeensis Steph. 1886 = Leptolejeunea maculata (Mitt.) Schiffn. 1898. 

Lejeunea thwaitesiana Mitt. 1861 = Drepanolejeunea thwaitesiana (Mitt.) Steph. 1913 [Dubia et Excludenda]. 

Lejeunea tortuosa Lehm. et Lindenb. 1845 = Odontolejeunea lunulata (F.Weber) Schiffn. 1893. 

Lejeunea trematodes Nees 1836 = Drepanolejeunea trematodes (Nees) Bischl. 1968. 

Lejeunea trichomanoides Mont. et Nees 1845 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914. 

Lejeunea trifaria (Reinw. et al.) Gottsche et al. 1845 = Cheilolejeunea trifaria (Reinw. et al.) Mizut. 1964. 

Lejeunea trinitensis Lindenb. COM(4). 

Lejeunea triquetra Mitt. 1863 = Thysananthus spathulistipus (Reinw. et al.) Gottsche et al. 1845. 

Lejeunea tristis Steph. 1890 = Frullanoides tristis (Steph.) van Slageren 1985. 

Lejeunea truncatiloba Steph. 1892 = Leptolejeunea maculata (Mitt.) Schiffn. 1898. 

Lejeunea tuberculiflora E.W.Jones ex Pócs 1979, nom. illeg. = L. camerunensis (Steph.) E.W.Jones 1972. 

Lejeunea turgida Steph. 1892, hom. illeg. [non Mitt. 1861] = Mastigolejeunea turgida Steph. 1912. 

Lejeunea ulicina (Taylor) Gottsche et al. 1945 = Microlejeunea ulicina (Taylor) A.Evans. 1902. 

Lejeunea ulicina ssp. africana (Steph.) Vanden Berghen 1972 = Microlejeunea africana Steph. 1888. 

Lejeunea ulicina ssp. ocellifera (S.W.Arnell) R.M.Schust. 1962 = Microlejeunea africana Steph. 1888. 

Lejeunea unciloba Lindenb. 1845 = Leucolejeunea unciloba (Lindenb.) A.Evans 1907. 

Lejeunea variabilis Lindenb. 1845 = Ceratolejeunea variabilis (Lindenb.) Schiffn. 1893.


Lejeunea villaumei (Steph.) Grolle KEN(2), MDG(1), MLW(563), REU(1), RWA(2), SAF(2), SEY(1), 

TAN(2). 

Lejeunea vogelii Taylor 1846 = L. caespitosa Lindenb. 1845. 

Lejeunea vulgariformis Gottsche MDG(1). 

Lejeunea wilmsii (Steph.) Steph. 1915 = L. eckloniana Lindenb. 1845. 

Lejeunea xanthocarpa (Lehm. et Lindenb.) Gottsche et al. 1845 = Leucolejeunea xanthocarpa (Lehm. et 


Lejeunea sp. (Hygrolejeunea patellirostris Steph.). Treated as synonymous with Lejeunea acuta Mitt. in Tixier 

1995 (130). However, E.W.Jones (in litt.) previously commented that although a specimen under the name 

Hygrolejeunea patellirostris in Hb. Stephani (G) is L. acuta, it is not the type, "nor does it agree well with the 

description of the type". 

Lejeunea sp. (Hygrolejeunea staudtiana Steph.) BIO(2), CMN(2), UGA(2). 

Lejeunea sp. (Prionolejeunea corbisieri Pearson) ZAI(2). 

Lembidium borbonicum Steph. 1893 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Lembidium heteromorphum (Lehm.) Steph. 1891 = Alobiellopsis heteromorpha (Lehm.) R.M.Schust. 1969. 

Leperoma ochroleuca (Spreng.) Mitt. 1877 = Lepicola ochroleuca (Spreng.) Spruce 1885. 

Lepicolea ochroleuca (Spreng.) Spruce SAF(2,107). 

Lepidolejeunea bidentula (Steph.) R.M.Schust. MDG(1), SEY(1,4,108). 

Lepidolejeunea delessertii (Nees et Mont.) Grolle MAU(1), REU(1,4). 

Lepidolejeunea serrulata (Steph.) Grolle CMN(2), GHA(2), RIM(10). 

Lepidozia abyssinica S.W.Arnell 1960 = L. stuhlmannii var. stuhlmannii Steph. 1895 (92). 

Lepidozia africana Steph. MAU(1), MDG(1), REU(1,4), TAN(2). 

Lepidozia bicuspidata C.Massal. 1885 = Hyalolepidozia bicuspidata (C.Massal.) S.W.Arnell ex Grolle 1964. 

Lepidozia capillaris (Sw.) Lindenb. 1845 = Kurzia capillaris (Sw.) Grolle 1964. 

Lepidozia carnosa Steph. 1911 = L. stuhlmannii var. carnosa (Steph.) Pócs (96). 

Lepidozia cupressina (Sw.) Lindenb. KEN(2), LES(75), RWA(2), SAF(2), TAN(2), UGA(2,96), ZAI(2). 

Lepidozia cupressina ssp. natalensis (Steph.) Pócs SAF(2), TAN(2). 

Lepidozia cupressina ssp. quinquefida (Steph.) Pócs TAN(2), ZAI(2). 

Lepidozia hexiloba Pearson GHA(2). 

Lepidozia hyalina Steph. 1911 = L. pearsonii Spruce 1891. 

Lepidozia irregularis Steph. 1911 = Kurzia irregularis (Steph.) Grolle 1982. 

Lepidozia lacerata Steph. 1909 = L. stuhlmannii Steph. 1895. 

Lepidozia laevifolia (Hook.f. et Taylor) Gottsche et al. STH(2). 

Lepidozia laurentii Steph. 1909 = L. succida Mitt. 1860. 

Lepidozia longitexta Steph. 1909 = L. succida Mitt. 1860. 

Lepidozia microphylla var. ß (Hook.) Lindenb. 1845 [quoad pl. Madagascar] = L. africana Steph. 1922. 

Lepidozia muenchiana Steph. 1909 = L. cupressina (Sw.) Lindenb. 1845. 

Lepidozia natalensis Steph. 1909 = L. cupressina ssp. natalensis (Steph.) Pócs 1984. 

Lepidozia nematodes (Gottsche ex Austin) Spruce 1885 = Telaranea nematodes (Gottsche ex Austin) M.Howe 

1902. 

Lepidozia nemoides (Hook.f. et Taylor) Lindenb. et Gottsche 1846 = Kurzia nemoides (Hook.f. et Taylor) Grolle 

1964. 

Lepidozia pearsonii Spruce MLW(2), SAF(2), UGA(2), ZAI(2). 

Lepidozia pearsonii var. lacerata (Steph.) Pócs 1984 = L. stuhlmannii Steph. 1895. 

Lepidozia pearsonii var. pulvinata (Steph.) Pócs 1984 = L. stuhlmannii ssp. pulvinata (Steph.) Pócs 1994. 

Lepidozia procumbens Mitt. 1875 = L. laevifolia (Hook.f. et Taylor) Gottsche et al. 1845. 

Lepidozia pulvinata Steph. 1911 = L. stuhlmannii ssp. pulvinata (Steph.) Pócs 1994. 

Lepidozia quinquefida Steph. 1911 = L. cupressina ssp. quinquefida (Steph.) Pócs 1984. 

Lepidozia quintasii Steph. 1922 = Telaranea nematodes (Gottsche ex Austin) M.Howe 1902. 

Lepidozia redacta Steph. 1911 = Telaranea nematodes (Gottsche ex Austin) M.Howe 1902. 

Lepidozia reducta Steph. 1922, nom. illeg. = Telaranea nematodes (Gottsche ex Austin) M.Howe 1902. 

Lepidozia reptans (L.) Dumort. ZAI(2). 

Lepidozia spinosa S.W.Arnell 1954 = Psiloclada clandestina Mitt. 1855. 

Lepidozia stephanii Renauld ex Steph. 1890 = Kurzia capillaris ssp. stephanii (Renauld) Pócs 1984. 

Lepidozia stuhlmannii Steph. MLW(92), UGA(96).


Lepidozia stuhlmannii var. abyssinica (S.W.Arnell) Pócs 1994 = L. stuhlmannii var. stuhlmannii Steph. 1895 

(92). 

Lepidozia stuhlmannii ssp. pulvinata (Steph.) Pócs KEN(2), RWA(2), UGA(2), ZAI(2). 

Lepidozia stuhlmannii ssp. stuhlmannii Steph. BIO(36), CMN(2), ETH(2), KEN(2), MLW(2), RWA(2), 

SAF(2), TAN(2), UGA(2,42,50), ZAI(2). 

Lepidozia stuhlmannii var. carnosa (Steph.) Pócs KEN(7), UGA(7,96), ZAI(274). 

Lepidozia succida Mitt. ANG(2), BIO(2,33,36), CAF(2), CMN(2), CON(2), GAB(2), GHA(2), GUI(2), 

IVO(2), MDG(1,4), NGA(2), RIM(10), SAO(2), SIE(2), UGA(2,42,96), ZAI(2). 

Lepidozia succida var. ubangiensis (Steph.) Vanden Berghen 1983 = L. ubangiensis Steph. 1909. 

Lepidozia succulenta Sim 1926 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 2001 (556). 

Lepidozia tabularis Steph. 1909 = Kurzia capillaris (Sw.) Grolle 1964. 

Lepidozia trifida Steph. 1911 = Telaranea trifida (Steph.) R.M.Schust. 1966. 

Lepidozia truncatella Nees 1845 = L. cupressina (Sw.) Lindenb. 1845. 

Lepidozia ubangiensis Steph. CAF(2), CON(2), GAB(2), ZAI(2). 

Lepidozia ulothrix (Schwägr.) Lindenb. MAU(1). 

Lepidozia verrucosa Steph. 1885 = Kurzia capillaris var. verrucosa (Steph.) Pócs 1984. 

Lepidozia wallisiana Steph. 1909 = L. cupressina (Sw.) Lindenb. 1845. 

Leptocolea adhaesiva (Mitt.) E.W.Jones 1953 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963. 

Leptocolea africana (Steph.) E.W.Jones 1953 = Cololejeunea africana (Steph.) R.M.Schust. 1963. 

Leptocolea apiculata E.W.Jones 1954 = Cololejeunea apiculata (E.W.Jones) R.M.Schust. 1963. 

Leptocolea appressa A.Evans 1912 = Cololejeunea appressa (A.Evans) Benedix 1953. 

Leptocolea auriculata E.W.Jones 1953 = Cololejeunea auriculata (E.W.Jones) R.M.Schust. 1963. 

Leptocolea autoica (Steph.) E.W.Jones 1953, nom. inval. = Cololejeunea autoica (Steph.) Grolle 1995. 

Leptocolea bolombensis (Steph.) E.W.Jones 1953 = Cololejeunea tonkinensis Steph. 1895. 

Leptocolea cardiocarpa (Mont.) A.Evans 1911 = Cololejeunea cardiocarpa (Mont.) A.Evans 1902. 

Leptocolea crenatiflora Steph. 1915 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897. 

Leptocolea crenatifolia Steph. ex S.W.Arnell 1965, nom. inval. = Cololejeunea obliqua (Nees et Mont.) Schiffn. 

1897. 

Leptocolea cristata (Steph.) E.W.Jones 1953 = Cololejeunea cristata (Steph.) R.M.Schust. 1963. 

Leptocolea cristata var. lanciloba S.W.Arnell 1963 = Cololejeunea cardiocarpa (Mont.) A.Evans 1902. 

Leptocolea cristata var. occidentalis E.W.Jones 1953 = Cololejeunea occidentalis (E.W.Jones) Vanden Berghen 

1978 (654). 

Leptocolea cuneifolia (Steph.) A.Evans 1911 = Cololejeunea cuneifolia Steph. 1892. 

Leptocolea dentata E.W.Jones 1953 = Cololejeunea dentata (E.W.Jones) R.M.Schust. 1963. 

Leptocolea distalopapillata E.W.Jones 1957 = Cololejeunea distalopapillata (E.W.Jones) R.M.Schust. 1963. 

Leptocolea furcilobulata Berrie et E.W.Jones 1954 = Cololejeunea furcilobulata (Berrie et E.W.Jones) 


Leptocolea himalayensis Pandé et Misra 1943 = Cololejeunea latilobula (Herzog) Tixier 1985. 

Leptocolea latilobula Herzog 1930 = Cololejeunea latilobula (Herzog) Tixier 1985. 

Leptocolea leloutrei E.W.Jones 1953 = Cololejeunea leloutrei (E.W.Jones) R.M.Schust. 1963. 

Leptocolea mascarena Steph. ex S.W.Arnell 1965, nom. inval. = Leptolejeunea maculata (Mitt.) Schiffn. 1898. 

Leptocolea montagnei (Gottsche) Steph. 1916 = Cheilolejeunea montagnei (Gottsche) Schiffn. 1898. 

Leptocolea nigerica E.W.Jones 1953 = Cololejeunea nigerica (E.W.Jones) R.M.Schust. 1963. 

Leptocolea nymanii Steph. 1916 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897. 

Leptocolea perforata Schiffn. ex S.W.Arnell 1965, nom. inval. = Drepanolejeunea trematodes (Nees) Bischl. 

1968. 

Leptocolea perforata sensu S.W.Arnell 1965 [non Schiffn. ex S.W.Arnell 1965, nom. inval.] = Drepanolejeunea 

madagascariensis (Steph.) Grolle 1974. 

Leptocolea punctata E.W.Jones 1957 = Cololejeunea vidaliana Tixier 1973. 

Leptocolea quintasii var mascarenica S.W.Arnell 1965, nom. inval. = Leptolejeunea epiphylla (Mitt.) Steph. 

1913. 

Leptocolea scabrifolia var. africana Herzog 1947, nom. inval. = Cololejeunea obliqua (Nees et Mont.) Schiffn. 

1897. 

Leptocolea subciliata (Mitt.) E.W.Jones 1953 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947. 

Leptocolea tonkinensis (Steph.) Steph. 1916 = Cololejeunea tonkinensis Steph. 1895. 

Leptolejeunea adhaesiva (Mitt.) Steph. 1895 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963.


Leptolejeunea astroidea (Mitt.) Steph. BUR(2), CMN(2), CON(2), GHA(2), GNB(2), GUI(2), IVO(2), 

NGA(2), RIM(10), SIE(2), UGA(42,50,115), ZAI(2), ZIM(2). 

Leptolejeunea convexistipa Bischl. 1969 = L. maculata (Mitt.) Schiffn. 1898. 

Leptolejeunea degelii S.W.Arnell 1962 = Diplasiolejeunea symoensii Vanden Berghen 1960. 

Leptolejeunea elliptica (Lehm. et Lindenb.) Schiffn. COM(1). 

Leptolejeunea epiphylla (Mitt.) Steph. CMN(2), COM(1), CON(2), GHA(2), GUI(2), IVO(2), KEN(2), 

MAU(1,4), NGA(2), RIM(10), SAO(2), SEY(4,108), SIE(2), TAN(2), UGA(42), ZAI(2). 

Leptolejeunea maculata (Mitt.) Schiffn. CMN(2), COM(1,4), GAB(2), GHA(2), GUI(2), IVO(2), 

KEN(2), MAU(1), NGA(2), RIM(10), SAO(2), SEY(1,4,108), SIE(2), TAN(2), TOG(2), 

UGA(42,50,96), ZAI(2). 

Leptolejeunea madagascariensis Steph. 1913 = Drepanolejeunea madagascariensis (Steph.) Grolle 1974. 

Leptolejeunea mascarena Steph. 1913 = L. maculata (Mitt.) Schiffn. 1898. 

Leptolejeunea papuliflora Steph. 1913 = L. epiphylla (Mitt.) Steph. 1913. 

Leptolejeunea perforata (Schiffn.) Schiffn. 1897 = Drepanolejeunea trematodes (Nees) Bischl. 1968. 

Leptolejeunea quintasii (Steph.) Steph. 1913 = L. epiphylla (Mitt.) Steph. 1913. 

Leptolejeunea quintasii var. mascarenica S.W.Arnell 1965, nom. inval. = L. epiphylla (Mitt.) Steph. 1913. 

Leptolejeunea radiata (Mitt.) Steph. 1896 = L. maculata (Mitt.) Schiffn. 1898. 

Leptolejeunea repanda (Schiffn.) Steph. 1913 = L. maculata (Mitt.) Schiffn. 1898. 

Leptolejeunea symoensii Vanden Berghen 1961 = Drepanolejeunea symoensii (Vanden Berghen) Grolle 1976. 

Leptolejeunea thomeensis (Steph.) Steph. 1913 = L. maculata (Mitt.) Schiffn. 1898. 

Leptolejeunea trematodes (Nees) Steph. 1913 = Drepanolejeunea trematodes (Nees) Bischl. 1968. 

Leptolejeunea triocellata Steph. 1913 = Drepanolejeunea trematodes (Nees) Bischl. 1968. 

Leptolejeunea truncatiflora Steph. 1911 = L. astroidea (Mitt.) Steph. 1913. 

Leptolejeunea truncatiloba (Steph.) Steph. 1896 = L. maculata (Mitt.) Schiffn. 1898. 

Leptolejeunea vitrea (Nees) Schiffn. MAS(1). 

Leptoscyphus bewsii Sim 1926 = Lophocolea bewsii (Sim) Grolle 1959. 

Leptoscyphus diversifolius (Gottsche) Grolle SAF(2). 

Leptoscyphus expansus (Lehm.) Grolle LES(75), MDG(1), MLW(29), RWA(2), SAF(2), ZAI(2), 

ZIM(2). 

Leptoscyphus gottscheanus (Lindenb. ex Lehm.) Gottsche 1858 = L. expansus (Lehm.) Grolle 1963. 

Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust. KEN(2), RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Leptoscyphus henkelii Sim 1926 = Heteroscyphus grandistipus ssp. henkelii (Sim) Grolle 1963. 

Leptoscyphus infuscatus (Mitt.) E.W.Jones BIO(2,33,36), CMN(2), ETH(2), MLW(2,29), REU(1,4), 

RWA(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Leptoscyphus iversenii (Pearson) Sim 1926 = L. expansus (Lehm.) Grolle 1963. 

Leptoscyphus leightonii Sim 1926 = Lophocolea martiana Nees 1845. 

Leptoscyphus stephensii Sim 1926 = Lophocolea bidentata (L.) Dumort. 1835. 

Lethocolea congesta (Lehm.) S.W.Arnell COM(1,4), KEN(2), LES(75), REU(1,4), RWA(2), SAF(2), 

TAN(2), UGA(2), ZAI(2). 

Leucolejeunea capensis S.W.Arnell 1953 = L. xanthocarpa (Lehm. et Lindenb.) A.Evans 1907. 

Leucolejeunea knysnana S.W.Arnell 1953 = L. unciloba (Lindenb.) A.Evans 1907. 

Leucolejeunea rotalis (Hook.f. et Taylor) S.W.Arnell 1961 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. 

et Geissler 1997 (55). 

Leucolejeunea rotundistipula (Lindenb. ex Lehm.) Steph. LES(75), MAU(1), MLW(563), REU(1), 

SAF(2,75), ZIM(2). 

Leucolejeunea unciloba (Lindenb.) A.Evans SAF(2), TAN(2). 

Leucolejeunea xanthocarpa (Lehm. et Lindenb.) A.Evans BIO(2,33), COM(4), ETH(2), KEN(2), 

MDG(4), MLW(563), MOZ(2), REU(1,4), RWA(2), SAF(2), STH(2), TAN(2), UGA(2,42), ZAI(2), 

ZIM(2). 

Lindigina renifolia Mitt. 1877 = Gongylanthus renifolius (Mitt.) Steph. 1906. 

Lindigina scariosa (Lehm.) Mitt. 1877 = Gongylanthus scariosus (Lehm.) Steph. 1906. 

Lophocolea amanica Steph. 1922 = L. martiana Nees 1845. 

Lophocolea armatistipula Steph. 1911 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963. 

Lophocolea ascensionis Steph. ASC(2). 

Lophocolea bewsii (Sim) Grolle SAF(2).


Lophocolea bidentata (L.) Dumort. BIO(2,33,36), BUR(2), CMN(2), CVI(2,37), ETH(2), KEN(2,21), 

MAU(1), MDG(1), MLW(2,29), REU(1), RWA(2), SAF(2), STH(49), TAN(2), UGA(2,42,96), ZAI(2). 

Lophocolea bidentata sensu Sim 1926 = L. difformis Nees 1845. 

Lophocolea bidentata var. δ capensis Gottsche et al. 1845 = L. difformis Nees 1845. 

Lophocolea borbonica Steph. 1890 = L. difformis Nees 1845. 

Lophocolea cagnii Gola 1907 = L. difformis Nees 1845. 

Lophocolea cambouena Steph. 1907 = L. concreta Mont. 1845. 

Lophocolea concreta Mont. ANN(2,33), BUR(2), GHA(2), KEN(2), LES(75), MAU(1), MDG(1), 

MLW(2,29), NGA(2), REU(1), ROD(1), RWA(2), SAF(2,75), SIE(2), SWZ(2), TAN(2), TOG(2), 

UGA(42,50), ZAI(2), ZAM(2), ZIM(2). 

Lophocolea congoana Steph. 1905 = L. martiana Nees 1845. 

Lophocolea connata (Sw.) Mont. et Nees 1839 [Dubia et Excludenda]. 

Lophocolea cuspidata (Nees) Limpr. 1876 = L. bidentata (L.) Dumort. 1835 (684). 

Lophocolea devexa Mitt. 1863 = Conoscyphus trapezioides (Sande Lac.) Schiffn. 1898. 

Lophocolea difformis Nees BIO(2,33,36), CAF(2), CMN(2), ETH(2), GHA(2), IVO(51), KEN(2), 

LES(75), MAU(4), MLW(2,29), NGA(2), REU(1), RWA(2), SAF(2,75), SAO(2), SIE(2), TAN(2), 

TOG(2), UGA(2,96), ZAI(2), ZIM(2). 

Lophocolea diversifolia Gottsche 1845 = Leptoscyphus diversifolius (Gottsche) Grolle 1963. 

Lophocolea diversifolia sensu Sim 1926 [non Gottsche 1845] = L. difformis Nees 1845. 

Lophocolea dusenii Steph. 1907 = L. martiana Nees 1845 or L. lucida (Spreng. ex Lehm.) Mont. 1839. 

Lophocolea esterhuysenii S.W.Arnell ex E.W.Jones 1953 = L. bewsii (Sim) Grolle 1959. 

Lophocolea expansa (Lehm.) Drège 1843 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Lophocolea fasciculata (Nees) S.W.Arnell et Grolle 1959 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Lophocolea fragrans (Moris et De Not.) Gottsche et al. COM(1), MLW(29), REU(1), SAF(2), TAN(2), 

ZAI(2). 

Lophocolea frappieri Steph. 1907 = L. fragrans (Moris et De Not.) Gottsche et al. 1845. 

Lophocolea hians Steph. 1901, perhaps = L. lucida (Spreng. ex Lehm.) Mont. 1839. 

Lophocolea humistrata (Hook.f. et Taylor) Gottsche et al. STH(2). 

Lophocolea inflata Steph. 1911 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963. 

Lophocolea integrifolia Steph. 1892 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Lophocolea lepervanchei Steph. REU(1). 

Lophocolea lillieana Steph. KEN(2). 

Lophocolea longifolia Steph. 1890 = L. difformis Nees 1845. 

Lophocolea longispica Steph. 1892 = L. bidentata (L.) Dumort. 1835. 

Lophocolea lucida (Spreng. ex Lehm.) Mont. CMN(2), COM(1), MDG(1), MLW(2,29), REU(1), 

ROD(1), SAF(2), SEY(1), TAN(2), UGA(96), ZAI(2), ZAM(2). 

Lophocolea macleana Steph. 1907 = L. difformis Nees 1845. 

Lophocolea madagascariensis Gottsche MDG(1). 

Lophocolea magnistipula S.W.Arnell 1954 = L. semiteres (Lehm.) Mitt. 1877. 

Lophocolea martiana Nees ANG(2), BIO(36,72), BUR(2), CAF(2), CMN(2), CON(2), GAB(2), 

GHA(2), IVO(2,53), MAU(1), MDG(1), MLW(29), NGA(2), PRI(2), REU(1), RIM(10), RWA(2), 

SAF(2), SEY(1,4), SIE(2), SWZ(2), TAN(2), TOG(2), UGA(2,96), ZAI(2), ZIM(2). 

Lophocolea minor Nees REU(1). 

Lophocolea moelleri Steph. 1886 = L. difformis Nees 1845. 

Lophocolea mollis (Nees) Gottsche et al. The distribution of this taxon is given only as "îles austro-africaines 

de l'Indien Océan". It is close to L. difformis and may be conspecific (fide Grolle 1995 (1)). 

Lophocolea muhavurensis (S.W.Arnell) S.W.Arnell ex Pócs KEN(2), TAN(2), UGA(2). 

Lophocolea muricata (Lehm.) Nees BIO(33,36,72), BUR(2), CMN(2), COM(1), ETH(2), MDG(1), 

MLW(29), MOZ(2), REU(1), RWA(2), SAF(2), TAN(2), UGA(2,42,50,96), ZAI(2), ZIM(2). 

Lophocolea newtonii Steph. 1907 = L. martiana Nees 1845. 

Lophocolea obscura Steph. 1907 = L. lucida (Spreng. ex Lehm.) Mont. 1839. 

Lophocolea occidentalis Steph. 1922 = Heteroscyphus dubius (Gottsche) Schiffn. 1910. 

Lophocolea onraedtii Grolle MDG(1), REU(1). 

Lophocolea opposita Mitt. 1879 = L. lucida (Lehm.) Mont. 1839. 

Lophocolea ovistipula Herzog 1952 = Clasmatocolea vermicularis (Lehm.) Grolle 1960.


Lophocolea rehmannii Steph. 1892 = L. concreta Mont. 1845. 

Lophocolea rubescens Steph. 1890 = L. lucida (Spreng. ex Lehm.) Mont. 1839. 

Lophocolea scott-elliottii Steph. 1907 = L. bidentata (L.) Dumort. 1835. 

Lophocolea semiteres (Lehm.) Mitt. SAF(2). 

Lophocolea setacea sensu Sim 1926 [non Steph. 1892] = L. difformis Nees 1845. 

Lophocolea setacea Steph. 1892 = L. bidentata (L.) Dumort. 1835. 

Lophocolea silvestris Gottsche 1882 [Dubia et Excludenda]. 

Lophocolea spiniflora Steph. 1907 = L. muricata (Lehm.) Nees 1845. 

Lophocolea staudtiana Steph. 1907 = L. martiana Nees 1845. 

Lophocolea subintegra (Hook.f. et Taylor) Grolle 1959 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Lophocolea subrotunda Mitt. 1879 = L. concreta Mont. 1845. 

Lophocolea subulistipa Herzog 1952 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Lophocolea sylvestris Gottsche ex Steph. 1907, nom. inval. = L. silvestris Gottsche 1882 [Dubia et Excludenda]. 

Lophocolea vermicularis Herzog 1952 = Clasmatocolea fasciculata (Nees) Grolle 1960. 

Lopholejeunea abbreviata (Mitt.) Steph. 1912 = Archilejeunea abbreviata (Mitt.) Vanden Berghen 1951. 

Lopholejeunea abortiva (Mitt.) Steph. 1912 = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea abortiva var. alata Vanden Berghen 1984 = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea abortiva var. fragilis (Steph.) Vanden Berghen 1984 = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea angustiflora Steph. 1912 = L. eulopha (Taylor) Schiffn. 1893. 

Lopholejeunea atra (Mitt.) Steph. 1895 = Acanthocoleus aberrans var. laevis Gradst. 1994. 

Lopholejeunea boivinii Steph. 1912 = L. subfusca (Nees) Schiffn. 1897. 

Lopholejeunea borbonica Steph. COM(1), MDG(1), REU(1), TAN(2). 

Lopholejeunea camerunensis Steph. 1911, nom. nud = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea erugata B.Thiers MDG(1). 

Lopholejeunea eulopha (Taylor) Schiffn. COM(1), MAU(1), MDG(1), REU(1), RWA(2), SEY(1), 

TAN(2). 

Lopholejeunea fimbriata (Gottsche) Schiffn. 1893 = L. eulopha (Taylor) Schiffn. 1893. 

Lopholejeunea fragilis Steph. 1912 = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea gabonensis Vanden Berghen GAB(2). 

Lopholejeunea grandicrista Steph. MDG(1), REU(1). 

Lopholejeunea jonesii Vanden Berghen BIO(2,33), CMN(2), IVO(2), NGA(2), SIE(2). 

Lopholejeunea kilimanjarica E.W.Jones 1957 = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea laceriloba Steph. 1923 = L. eulopha (Taylor) Schiffn. 1893. 

Lopholejeunea laciniata E.W.Jones MLW(563), TAN(2), ZIM(2). 

Lopholejeunea leioptera Gyarmati MDG(708). 

Lopholejeunea lepidoscypha Kiaer et Pearson MDG(1). 

Lopholejeunea minima Vanden Berghen REU(1). 

Lopholejeunea moenkemeyeri Steph. 1912 = L. subfusca (Nees) Schiffn. 1897. 

Lopholejeunea multilacera Steph. MAU(1), MDG(1), REU(1). 

Lopholejeunea nigricans (Lindenb.) Schiffn. BIO(2,33), BUR(2), CAF(2), CMN(2), CON(2), ETH(2), 

GAB(2), GHA(2), GUI(2), IVO(2,53), MAU(1), MDG(1), MLW(2,563), MOZ(2), NGA(2), REU(1), 

RIM(10), ROD(1), RWA(2), SAF(2,75), SEY(1), SIE(2), TAN(2), TOG(2), ZAI(2), ZAM(2), ZIM(2). 

Lopholejeunea obtusilacera Herzog CMN(2), IVO(2), NGA(2). 

Lopholejeunea onraedtii Vanden Berghen MDG(1). 

Lopholejeunea palustris (Sim) S.W.Arnell 1953 = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea paramultilacera Vanden Berghen REU(1). 

Lopholejeunea quinquecarinata Vanden Berghen MDG(1), REU(1), TAN(2). 

Lopholejeunea revoluta E.W.Jones UGA(2,50). 

Lopholejeunea sagraeana (Mont.) Schiffn. 1893 = L. subfusca (Nees) Schiffn. 1897. 

Lopholejeunea sinuata (Mitt.) Steph.1912 = L. sphaerophora (Lehm. et Lindenb.) Steph. 1912. 

Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph. MAU(1). 

Lopholejeunea subfusca (Nees) Schiffn. ANG(2), ANN(2,33), BIO(2,33), CAF(2), CMN(2), COM(1), 

GAB(2), GHA(2), GUI(2), IVO(2,53), KEN(2), MAU(1), MDG(1), NGA(2), REU(1,2), RIM(10), 

RWA(2), SAF(2), SEY(1), SIE(2), TAN(2), TOG(2,101), UGA(96,115), ZAI(2). 

Lopholejeunea subfusca var. elongata Vanden Berghen SEY(1).


Lopholejeunea subinermis Steph. 1895, probably = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea tixieriana Vanden Berghen MDG(1). 

Lopholejeunea utriculata Steph. MDG(1). 

Lopholejeunea vermoesenii Pearson 1922, probably = L. nigricans (Lindenb.) Schiffn. 1898. 

Lopholejeunea zenkeri Steph. 1912 = L. nigricans (Lindenb.) Schiffn. 1898. 

Lophozia aberrans (Nees et Mont.) Steph. 1901 = Andrewsianthus aberrans (Nees et Mont.) Grolle 1963. 

Lophozia argentina (Steph.) R.M.Schust. LES(75), SAF(2). 

Lophozia capensis S.W.Arnell 1953 = Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust. 1995. 

Lophozia decolorans (Limpr.) Steph. CMN(2), TAN(2), ZAI(2). 

Lophozia hedbergii S.W.Arnell LES(75), UGA(2). 

Lophozia incisa (Schrad.) Dumort. 1835. The plant named as L. incisa in Váòa, Pócs & De Sloover 1979 (156) 

is Andrewsianthus jamesonii (Mont.) Váòa 1983. 

Lophozia kilimanjarica S.W.Arnell 1956 = A. jamesonii (Mont.) Váòa 1983. 

Lophozia kinangopii Gola 1914 = Isotachis aubertii (Schwägr.) Mitt. 1886. 

Lophozia madagascariensis Steph. 1917 = Cylindrocolea madagascariensis (Steph.) R.M.Schust. 1969. 

Lophozia montaguensis S.W.Arnell 1954 = Lophozia argentina (Steph.) R.M.Schust. 1966. 

Lophozia multiflora Steph. 1917 = Gymnocoleopsis multiflora (Steph.) R.M.Schust. 1978. 

Lophozia ruwenzorensis S.W.Arnell 1956 = Andrewsianthus bilobus (Mitt.) Grolle 1963. 

Lunularia cruciata (L.) Dumort. ex Lindb. CMN(48), CVI(2), ERI(2), ETH(2,113), KEN(2,71,665), 

MLW(2,114), NAM(2), RWA(2), SAF(2), TAN(2), UGA(2,42,96), ZAI(2), ZIM(2). 

Lunularia vulgaris Raddi 1818, nom. illeg. = L. cruciata (L.) Dumort. ex Lindb. 1822. 

Madotheca abyssinica Nees 1845 = Porella abyssinica (Nees) Trevis. 1877. 

Madotheca capensis Gottsche 1845 = Porella capensis (Gottsche) Mitt. 1886. 

Madotheca cucullistipula (Steph.) Paris 1903 = Porella cucullistipula Steph. 1893. 

Madotheca hoehneliana Steph. 1895 = Porella abyssinica var. hoehnelii (Steph.) Pócs 1994. 

Madotheca katangae Gola 1920 = Porella abyssinica (Nees) Trevis. 1877. 

Madotheca keniana Steph. 1924 = Porella abyssinica var. hoehnelii (Steph.) Pócs 1994. 

Madotheca lobata Steph. 1910 = Porella montantii (Steph.) E.W.Jones 1963. 

Madotheca madagascariensis (Nees et Mont.) Gottsche et al. 1845 = Porella madagascariensis (Nees et Mont.) 

Trevis. 1877. 

Madotheca matongae Herzog 1952 = Porella vallis-gratiae (Gottsche ex Steph.) S.W.Arnell 1963. 

Madotheca mattirolii Gola 1914 = Porella mattirolii (Gola) S.W.Arnell 1960. 

Madotheca montantii Steph. 1910 = Porella montantii (Steph.) E.W.Jones 1963. 

Madotheca nilgheriensis Mont. 1842 = Porella nilgheriensis (Mont.) Trevis. 1877 [Dubia et Excludenda]. 

Madotheca ovifolia Steph. 1910 = Porella triquetra (Steph.) E.W.Jones 1963. 

Madotheca prolixa Gottsche ex Steph. 1910 = Porella prolixa (Gottsche ex Steph.) E.W.Jones 1963. 

Madotheca schroederi Steph. 1924 = Porella abyssinica var. hoehnelii (Steph.) Pócs 1994. 

Madotheca steudneri Gottsche 1867 = Porella abyssinica var. hoehnelii (Steph.) Pócs 1994. 

Madotheca subdentata Mitt. 1863 = Porella subdentata (Mitt.) E.W.Jones 1963. 

Madotheca thomeensis Steph. 1910 = Porella subdentata (Mitt.) E.W.Jones 1963. 

Madotheca triquetra Steph. 1895 = Porella triquetra (Steph.) E.W.Jones 1963. 

Madotheca vallis-gratiae Gottsche ex Steph. 1910 = Porella vallis-gratiae (Gottsche ex Steph.) S.W.Arnell 

1955. 

Madotheca vastifolia Steph. 1910 = Porella triquetra (Steph.) E.W.Jones 1963. 

Mannia androgyna (L. emend. Lindb.) A.Evans CHA(2,3), CVI(2,37), ERI(2), ETH(2), NAM(2). 

Mannia capensis (Steph.) S.W.Arnell NAM(2), SAF(2), TAN(2), ZAM(2), ZIM(2). 

Mannia dichotoma S.W.Arnell 1961, nom. illeg. = M. androgyna (L.) A.Evans 1938. 

Marchantia androgyna L. 1753 = Mannia androgyna (L. emend. Lindb.) A.Evans 1938. 

Marchantia angustifolia Necker 1771 = Mannia androgyna (L. emend. Lindb.) A.Evans 1938. 

Marchantia balboa Gola 1914 [Dubia. Indeterminate] (132). 

Marchantia balboa var. acutisquamata Gerola 1947 [Dubia. Indeterminate] (132). 

Marchantia berteroana Lehm. et Lindenb. ASC(2), SAF(2), STH(2), ZIM(2). 

Marchantia cagnii Gola 1907 [Dubia. Indeterminate] (132). 

Marchantia cengiana Gerola 1947 [Dubia. Indeterminate] (132). 

Marchantia chevalieri Steph. ex Bonner 1953 = M. debilis Goebel 1915. 

Marchantia contracta Bisch. ex C.Krauss 1846 = M. berteroana Lehm. et Lindenb. 1834.


Marchantia crassa A.Richard 1834 = M. berteroana Lehm. et Lindenb. 1834. 

Marchantia cruciata L. 1753 = Lunularia cruciata (L.) Dumort. ex Lindb. 1822. 

Marchantia debilis Goebel ANG(2), BUR(2), CHA(2), CMN(2), COM(1), CON(2), GAB(2), GHA(2), 

GUI(2), IVO(2), LES(2,75), MDG(1), MLW(2), NGA(2), RWA(2), SAF(2,75), SIE(2), SWZ(2), 

TAN(2), TOG(2), UGA(2,42), ZAI(2), ZAM(2), ZIM(2). 

Marchantia flavescens Steph. ex Bonner 1953 = M. pappeana Lehm. 1857. 

Marchantia globosa Brid. ex F.Weber COM(1), MAU(1), REU(1). 

Marchantia hemisphaerica L. 1753 = Reboulia hemisphaerica (L.) Raddi 1818. 

Marchantia hirsuta Sw. 1788 = Dumortiera hirsuta (Sw.) Nees 1824. 

Marchantia keniae Gola 1914 [Dubia. Indeterminate] (132). 

Marchantia marginata (Nees) Lehm. 1823 [non Steph. ex Bonner 1953] = Asterella marginata (Nees) 

S.W.Arnell 1963. 

Marchantia paleacea Bertol. CVI(2,37), ETH(2), REU(1), SAF(2). 

Marchantia pappeana Lehm. ANG(2), BIO(2,33,36), BUR(2), CMN(2), COM(1), CON(2), 

CVI(2,20,37), ETH(2), GHA(2), GUI(2), IVO(2), KEN(2,665), LES(2,75), MDG(1), MLW(2,114), 

RWA(2), SAF(2,75), SAO(2), SIE(2), SWZ(2), TAN(2), TOG(2), UGA(2,96), ZAI(2), ZAM(2), 

ZIM(2). 

Marchantia papyracea Gola 1907 [Dubia. Indeterminate] (132). 

Marchantia parviloba Steph. 1895 = M. pappeana Lehm. 1857. 

Marchantia planiloba Steph. 1886 = M. pappeana Lehm. 1857. 

Marchantia polymorpha L. BUR(2), ETH(2), KEN(2), REU(1), RWA(2), TAN(2), UGA(2), ZAI(2). 

Reported in Frahm, Lindlar et al. 1996 (37) as occurring in the Cape Verde Islands, but the specimen they cite 

(Chevalier H23) is identified in Bischler-Causse 1993 (132) as M. paleacea. 

Marchantia polymorpha ssp. montivagans Bischl. et Boisselier ETH(2), UGA(96). 

Marchantia polymorpha ssp. ruderalis Bischl. et Boisselier SAF(2). 

Marchantia sellae Gola 1907 [Dubia. Indeterminate] (132). 

Marchantia spathysii Lindenb. 1829 = Athalamia spathysii (Lindenb.) S.Hatt. 1954. 

Marchantia stephanii Vanden Berghen 1954 = M. pappeana Lehm. 1857. 

Marchantia syngenesica Bory 1804 = Asterella syngenesica (Bory) Grolle 1974. 

Marchantia tabularis Nees 1838 = M. berteroana Lehm. et Lindenb. 1834. 

Marchantia tusii Gola 1914 [Dubia. Indeterminate] (132). 

Marchantia umbellata Steph. 1895, hom. illeg. [non Scopoli 1772] = M. pappeana Lehm. 1857. 

Marchantia viridula Lehm. et Lindenb. 1857 = M. globosa Brid. ex F.Weber 1815. 

Marchantia wilmsii Steph. 1892 = M. pappeana Lehm. 1857. 

Marchantia winkleri Steph. ex Bonner 1953 = M. pappeana Lehm. 1857. 

Marchesinia acutiloba (Hook.f. et Taylor) Trevis. 1877 = M. brachiata (Sw.) Schiffn. 1893. 

Marchesinia brachiata (Sw.) Schiffn. STH(2). 

Marchesinia chrysophylla (Lehm.) Steph. 1912 = Acanthocoleus chrysophyllus (Lehm.) Kruijt 1988. 

Marchesinia deslooveri Vanden Berghen RWA(2), TAN(2). Geissler 1997 (126) named this taxon 

Marchesinia cf. acutiloba. 

Marchesinia excavata (Mitt.) Schiffn. ANN(2,33), BIO(2,33,72), CMN(2), GAB(2), GHA(2), GUI(2), 

KEN(2), MLW(2), NGA(2), RIM(10), SAO(2), SIE(2), SOC(558), TAN(2), TOG(2), ZAI(2), ZAM(2). 

Marchesinia mackaii (Hook.) Gray CVI(124). The collection from Socotra reported in Kürschner 2000 (94) 

was later redetermined as M. excavata (Kürschner 2001 (558)). 

Marchesinia madagassa Steph. BUR(2), CMN(2), COM(4), KEN(2,665), MDG(1), MLW(2,563), 

REU(1), SIE(2), TAN(2), ZAI(2), ZIM(2). The (sterile) plant from Mt Kasigau, Kenya reported in Bizot & 

Pócs 1974 (136), is probably a Lopholejeunea (fide Geissler 1997 (126)). 

Marchesinia microscypha (Hook.f. et Taylor) Trevis. 1877 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) 

A.Evans 1907. 

Marchesinia moelleriana Pearson 1924 = M. madagassa Steph. 1912. 

Marchesinia obtusifolia Vanden Berghen 1984 = M. madagassa Steph. 1912. 

Marchesinia pappeana (Nees) Kuntze 1891 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Marsupella africana Steph. ex Bonner 1953 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996 

(153). 

Marsupella aurita (Lehm.) Sim 1926 = Anastrophyllum auritum (Lehm.) Steph. 1901.


Marsupella boeckii var. ruwenzorensis S.W.Arnell 1957 = Nardia arnelliana Grolle 1964. 

Marsupella capensis S.W.Arnell 1957 = Gymnomitrion andinum (Herzog) Herzog 1934 (111). 

Marsupella emarginata (Ehrh.) Dumort. RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Marsupella hedbergii S.W.Arnell 1956 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996 (153). 

Marsupella integra N.Kitag. 1967 = M. subintegra S.W.Arnell 1956. 

Marsupella sparsifolia (Lindb.) Dumort. SAF(2). 

Marsupella subintegra S.W.Arnell MLW(29), REU(1), RWA(2), UGA(2). 

Marsupella subquadrata Steph., in sched. et icones ined. = Apomarsupella africana (Steph. ex Bonner) 

R.M.Schust. 1996 (153). 

Marsupidium brevifolium Steph. 1908 = Adelanthus decipiens (Hook.) Mitt. 1864. 

Marsupidium excisum Mitt. 1877 = Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust. 1961. 

Marsupidium limbatum (Steph.) Grolle MAU(1), MLW(114), REU(1), SAF(2), TAN(2). 

Mastigobryum borbonicum Steph. 1885 = Bazzania praerupta (Reinw. et al.) Trevis. 1877. 

Mastigobryum boschianum Steph. 1924 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Mastigobryum comorense (Steph.) Steph. 1908 = Bazzania comorense Steph. 1891 [B. decrescens complex]. 

Mastigobryum convexum Lindenb. 1845 = Bazzania nitida (F.Weber) Grolle 1960. 

Mastigobryum curvidens (Steph.) Steph. 1908 = Bazzania curvidens Steph. 1891 [B. decrescens complex]. 

Mastigobryum decrescens (Lehm. et Lindenb.) Gottsche et al. 1845 = Bazzania decrescens (Lehm. et Lindenb.) 

Trevis. 1877. 

Mastigobryum diclados (in Stephani 1909 (161)) - a typographical error for Mastigophora diclados. 

Mastigobryum exile Lindenb. 1845 = Acromastigum exile (Lindenb.) A.Evans 1934. 

Mastigobryum fuscum (Steph.) Besch. 1895 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Mastigobryum integrum (Nees et Mont.) Gottsche et al. 1845 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 

1877. 

Mastigobryum lacouturei Steph. 1924 = Bazzania sp. [B. decrescens complex]. 

Mastigobryum laxifolium Steph. 1911 = Bazzania roccatii Gola 1907. 

Mastigobryum madagassum Steph. 1924 = Bazzania sp. [B. decrescens complex]. 

Mastigobryum mascarenum Steph. 1886 = Bazzania mascarenum (Steph.) Herzog 1947 [B. decrescens 

complex]. 

Mastigobryum molleri Steph. 1886 = Bazzania decrescens ssp. molleri (Steph.) E.W.Jones 1975. 

Mastigobryum novae-hollandiae Nees 1845 = Bazzania adnexa (Lehm. et Lindenb.) Trevis. 1877 [Dubia et 

Excludenda]. 

Mastigobryum pellucidum Steph. 1908 = Bazzania decrescens ssp. molleri (Steph.) E.W.Jones 1975. 

Mastigobryum perrotanum Steph. 1908 [non Steph. 1924] = Bazzania mascarena (Steph.) Herzog 1947 [B. 

decrescens complex]. 

Mastigobryum perrotanum Steph. 1924, hom. illeg. [non Steph. 1908] = Bazzania perrotana E.W.Jones 1975. 

Mastigobryum pulvinatum Steph. 1891 = Bazzania decrescens ssp. pumila (Mitt.) Pócs 1994. 

Mastigobryum pumilum (Mitt.) Steph. 1908 = Bazzania decrescens ssp. pumila (Mitt.) Pócs 1994. 

Mastigobryum reflexum Gottsche 1882 = Bazzania reflexa (Gottsche) Steph. 1891. 

Mastigobryum robertii Steph. 1908 = Bazzania decrescens (Lehm. et Lindenb.) Trevis. 1877. 

Mastigobryum schismoideum Steph. 1888 = Bazzania praerupta (Reinw. et al.) Trevis. 1877. 

Mastigobryum seychellarum (Gottsche ex Steph.) Steph. 1908 = Bazzania decrescens (Lehm. et Lindenb.) 

Trevis. 1877. 

Mastigobryum staudtianum Steph. 1908 = Bazzania decrescens ssp. molleri (Steph.) E.W.Jones 1975. 

Mastigobryum stephanii J.B.Jack ex Steph. 1886 = Bazzania nitida (F.Weber) Grolle 1960. 

Mastigobryum stoloniferum (Sw.) Lindenb. 1845 = Bazzania stolonifera (Sw.) Trevis. 1877 [Dubia et 

Excludenda]. 

Mastigolejeunea africana Steph. ex Paris 1911, nom. inval. = M. florea (Mitt.) Steph. 1912. 

Mastigolejeunea auriculata (Wils.) Schiffn. ANN(2,33), BEN(22), CAF(2), CMN(2), CON(2), GAB(2), 

GHA(2), GUI(2), IVO(2,53), MAU(4), MDG(1), MOZ(2), NGA(2,127), REU(1), RIM(10), ROD(1), 

SAO(2), SEN(2), SIE(2), SUD(2), TAN(2), TOG(2,101), UGA(96), ZAI(2). 

Mastigolejeunea buettneri Steph. 1888 = M. auriculata (Wils.) Schiffn. 1893. 

Mastigolejeunea carinata (Mitt.) Steph. 1912 = M. auriculata (Wils.) Schiffn. 1893. 

Mastigolejeunea corbieri Steph. 1912 = M. florea (Mitt.) Steph. 1912. 

Mastigolejeunea crispula Steph. 1912 = M. auriculata (Wils.) Schiffn. 1893. 

Mastigolejeunea dusenii Steph. 1912 = M. auriculata (Wils.) Schiffn. 1893.


Mastigolejeunea florea (Mitt.) Paris BEN(22), CAF(2), CMN(2), CON(2), GAB(2), GHA(2), GUI(2), 

IVO(2), NGA(2), PRI(48), RIM(10), SIE(2), TAN(2), TOG(2,101), UGA(115), ZAI(2). 

Mastigolejeunea fusco-virens Steph. 1912 = M. auriculata (Wils.) Schiffn. 1893. 

Mastigolejeunea gilletana Steph. 1912 = M. auriculata (Wils.) Schiffn. 1893. 

Mastigolejeunea humilis (Gottsche) Schiffn. 1893 = M. auriculata (Wils.) Schiffn. 1893. 

Mastigolejeunea jungneri Steph. CMN(2). 

Mastigolejeunea lacomblezii Pearson 1922 = M. nigra (Steph.) Steph. 1895. 

Mastigolejeunea microscypha (Hook.f. et Taylor) Steph. 1912 = Leucolejeunea xanthocarpa (Lehm. et 


Mastigolejeunea nigra Steph. BEN(22), CMN(2), GAB(2), GHA(2), GUI(2), IVO(2), MLW(2), 

NGA(2), SAO(2), SIE(2), TAN(2), UGA(96), ZAI(2). 

Mastigolejeunea rhodesica (Vanden Berghen) E.W.Jones CAF(2), KEN(2), MDG(1), MLW(2), REU(1), 

TAN(2), ZAI(2), ZAM(2). 

Mastigolejeunea robusta Steph. 1912 = Schiffneriolejeunea ferruginea (Steph.) Gradst. 1974. 

Mastigolejeunea sudanensis Herzog 1948 = M. auriculata (Wils.) Schiffn. 1893. 

Mastigolejeunea tarkwana Pearson 1931 = Thysananthus spathulistipus (Reinw. et al.) Lindenb. 1845. 

Mastigolejeunea trigona Steph. 1895 = M. auriculata (Wils.) Schiffn. 1893. 

Mastigolejeunea turgida Steph. BIO(2,33), CMN(2), GHA(2), NGA(2), PRI(2), SAO(2). 

Mastigophora africana S.W.Arnell 1957 = M. diclados (Brid. ex F.Weber) Nees 1838. 

Mastigophora africana var. annobonensis S.W.Arnell 1956, nom. inval. = M. diclados (Brid. ex F.Weber) Nees 

1838. 

Mastigophora diclados (Brid. ex F.Weber) Nees ANN(2,33), ASC(2), BIO(2,33), CMN(2), COM(1,4), 

ETH(2), GAB(2), MAS(1), MAU(1,4), MDG(1), MLW(114), REU(1), RIM(10), RWA(2), SAO(2), 

SEY(1,4,13), TAN(2), UGA(50). 

Mastigophora leioclada (Hook.f. et Taylor) Mitt. 1875 = M. diclados (Brid. ex F.Weber) Nees 1838. 

Mastigophora madagassa Steph. 1909 = Chandonanthus hirtellus (F.Weber) Mitt. 1873. 

Mastigophora mascarenica (Mitt.) Steph. 1891 = M. diclados (Brid. ex F.Weber) Nees 1838. 

Megaceros flagellaris (Mitt.) Steph. TAN(2). 

Megaceros lacerus (Nees) Steph. 1916 = Folioceros fuciformis (Mont.) Bharadw. 1975. 

Metacalypogeia fusca (Lehm.) N.Kitag. 1988 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Metalejeunea cucullata (Reinw. et al.) Grolle COM(1), MDG(1), SEY(1,4). 

Metzgeria africana Steph. TAN(543). To be clarified. M. africana was made a synonym of M. quadrifaria 

Steph. by Vanden Berghen (167), but supposedly differentiating characters were later described by Kuwahara 

(543). 

Metzgeria africana Steph. 1917 = M. quadrifaria Steph. 1899. 

Metzgeria agnewii Kuwah. ETH(2), KEN(2,21,71), LES(75), RWA(2), SAF(75), TAN(2), 

UGA(42,50,96,115). 

Metzgeria albinea Spruce SEY(1). 

Metzgeria allanii Steph. 1917 = M. australis Steph. 1889. Vanden Berghen 1948 (211) treated M. allanii as a 

synonym of M. camerunensis, whilst Kuwahara 1986 (155) treated M. allanii and M. camerunensis as 

synonyms of M. leptoneura. 

Metzgeria allionii Steph. MLW(2). 

Metzgeria attenuata Steph. BUR(2), RWA(2), UGA(42), ZAI(2). 

Metzgeria australis Steph. ANN(2,33), BIO(36), CMN(2) COM(1), ETH(2), GHA(2), IVO(2), KEN(2), 

MDG(1), MLW(2), MOZ(2), NGA(2), RIM(10), RWA(2), SAF(2), SAO(2), SEY(4), SIE(2), TAN(2), 

UGA(96), ZAI(2). 

Metzgeria borneensis Kuwah. MDG(1). 

Metzgeria brunnthaleri Steph. 1913 = M. nudifrons Steph. 1892. 

Metzgeria bystroemii S.W.Arnell BIO(2,33). 

Metzgeria camerunensis Steph. 1899 = M. australis Steph. (649). 

Metzgeria capensis S.W.Arnell 1953 = M. decipiens (C.Massal.) Schiffn. 1890. 

Metzgeria conjugata Lindb. MAU(1), MDG(1), MOZ(2), REU(1), SAF(2). 

Metzgeria consanguinea Schiffn. COM(1), KEN(2,71), LES(75), RWA(2), TAN(2), UGA(50), ZAI(2). 

Metzgeria convexa Vanden Berghen ETH(2), KEN(2), TAN(2), UGA(2).


Metzgeria corralensis Steph. SAF(2). 

Metzgeria decipiens (C.Massal.) Schiffn. COM(1), MAU(1), MDG(1), SAF(2), UGA(2), ZAI(2), 

ZIM(2). 

Metzgeria elliotii Steph. BUR(2), ETH(2), KEN(2), RWA(2), SAF(2), TAN(2), UGA(2), ZAI(2). 

Metzgeria epiphylla A.Evans UGA(2). 

Metzgeria furcata (L.) Dumort. CMN(2), ETH(2), MDG(1), MLW(2), REU(1), UGA(2), ZIM(2). There 

are no recent records of M. furcata from Africa, all the country records listed here having been made between 

1864 and 1922. Their status should be re-examined. 

Metzgeria furcata ssp. crassipilis Lindb. 1878 = M. crassipilis (Lindb.) A.Evans 1909. 

Metzgeria furcata var. decipiens C.Massal. 1885 = M. decipiens (C.Massal.) Schiffn. 1890. 

Metzgeria hamata auct. [non Lindb. 1878] = M. leptoneura Spruce 1885. 

Metzgeria hamata Lindb. 1878 [non M. hamata auct.] = M. procera Mitt. 1855 [Dubia et Excludenda]. 

Metzgeria hedbergii Vanden Berghen KEN(2), UGA(2,42,96). 

Metzgeria latifrons Steph. 1911 = M. elliotii Steph. 1899. 

Metzgeria leptoneura Spruce MAU(1), MDG(1), MLW(2), REU(1), RWA(2), SAO(2), SEY(4), 

TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Metzgeria limbato-setosa Steph. ETH(2), KEN(2), MOZ(2), RWA(2), SAF(2), TAN(2), UGA(2,42,96), 

ZAI(2), ZIM(2). 

Metzgeria madagassa Steph. 1899 = M. australis Steph. 1889. 

Metzgeria muscicola Steph. CMN(2), KEN(71), RWA(2), ZAI(2). 

Metzgeria nicomariei Veltman & Potgieter SAF(533). 

Metzgeria nudifrons Steph. CMN(2), LES(75), SAF(2,75), TAN(2), UGA(2), ZAI(2), ZIM(2). 

Metzgeria papulosa Steph. 1899 = M. leptoneura Spruce 1885 (424). 

Metzgeria perrotana Steph. 1899 = M. decipiens (C.Massal.) Schiffn. 1890. 

Metzgeria procera Mitt. 1855 [Dubia et Excludenda]. 

Metzgeria propagulifera Vanden Berghen MDG(1), TAN(2), ZAI(2). Although Kuwahara 1981 (154) 

placed M. propagulifera in synonymy with M. crassipilis (Lindb.) A.Evans, he seems to have abandoned that 

synonymy in 1986 (155). At any rate, in the later publication the distribution of M. crassipilis is shown to be 

confined to the New World, and M. propagulifera is not mentioned. Because of the uncertainty, M. propagulifera 

is kept as an independent species in this list, and the record from Madagascar, listed under M. crassipilis in (1) but 

originally named as M. propagulifera, is also included here. 

Metzgeria quadrifaria Steph. BUR(2), ETH(2), KEN(2), LES(75), SOM(2), TAN(2), ZAI(2), ZAM(2). 

Metzgeria recurva Steph. 1886 = M. leptoneura Spruce 1885. 

Metzgeria renauldii Steph. 1899 = M. leptoneura Spruce 1885. 

Metzgeria rufula Spruce CMN(2), UGA(2). 

Metzgeria ruwenzorensis Gola UGA(2). 

Metzgeria saxbyi Pearson BIO(2,33), CMN(2), GHA(2), SAF(2), SWZ(2), UGA(2,96). 

Metzgeria tabularis Steph. SAF(2), TAN(2). 

Metzgeria thomeensis Steph. 1891 = M. australis Steph. 1889 (1,624). 

Metzgeria tusii Gola KEN(2). 

Metzgeria vandenberghenii Kuwah. UGA(96), ZAI(2). 

Metzgeria violacea (Ach.) Dumort. BUR(2), KEN(2), REU(1), SAF(2), TAN(2), ZAI(2). 

Metzgeria violascens (Ach.) Dumort. ex S.W.Arnell 1965, nom. inval. = M. violacea (Ach.) Dumort. 1835. 

Metzgeria vivipara A.Evans REU(1). 

Metzgeria warnstorfii Steph. CMN(2), TAN(2), ZAI(2). 

Microlejeunea africana Steph. ANN(2,33), BEN(2), BIO(2,33,72), BUR(2), CAF(2), CMN(2), COM(1), 

CON(2), ETH(2), GHA(2), GNB(2), GUI(2), IVO(2,53), KEN(2,665), LES(75), MAU(108), MDG(1), 

MLW(2,563), MOZ(2), NGA(2), PRI(2), RIM(10), RWA(2), SAF(2,75), SAO(2), SEY(1,108), SIE(2), 

SOC(94), TAN(2), TOG(101), UGA(2,42), ZAI(2), ZIM(2). 

Microlejeunea albicans (Nees) Steph. 1890 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea alternifolia (Mitt.) Steph. 1915 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea ankasica E.W.Jones GHA(2), SIE(2). 

Microlejeunea brunnea Pearson 1922 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea byssoides (Gottsche) Pearson 1892 [Dubia et Excludenda].


Microlejeunea cochlearifolia Steph. 1888, perhaps = M. kamerunensis Steph. 1915. 

Microlejeunea colensoana Steph. 1916 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea cucullata (Reinw. et al.) Steph. 1915 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea cumingiana Steph. 1896 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea dispar Ast MAU(1), REU(1). 

Microlejeunea erectifolia (Spruce) Steph. 1889 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea fissistipula Steph. MDG(1). 

Microlejeunea furva (Mitt.) Steph. 1915 = Lejeunea eckloniana Lindenb. 1845. 

Microlejeunea gracillima (Mitt.) Steph. 1895 = M. ulicina (Taylor) A.Evans 1902. 

Microlejeunea gracillima sensu Sim 1926 [? non (Mitt.) Steph. 1895] = M. africana Steph. 1888. 

Microlejeunea helenae (Pearson) Sim 1926 = Lejeunea helenae Pearson 1886. 

Microlejeunea heterophylla Steph. 1915 = Lejeunea caespitosa Lindenb. 1845. 

Microlejeunea inflata Steph. COM(1), MDG(1). 

Microlejeunea jungneri Steph. 1915 = Lejeunea confusa E.W.Jones 1972. 

Microlejeunea kamerunensis Steph. BUR(2), CMN(2), IVO(2,53), KEN(2), MLW(563), MOZ(2), 

NGA(2), PRI(2), RIM(10), RWA(2), SAO(2), SIE(2), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Microlejeunea magnilobula Gola UGA(2). 

Microlejeunea minutistipula Steph. TAN(2). 

Microlejeunea monoica Bischl. 1963 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea oblongistipula (Gottsche) Pearson MDG(1), SEY(4,108). 

Microlejeunea ocellifera S.W.Arnell 1953 = M. africana Steph. 1888. 

Microlejeunea ovistipula Steph. ZAI(2). 

Microlejeunea papulosa (Gottsche) Pearson MDG(1). 

Microlejeunea parallela (Schiffn.) Steph. 1915 = Metalejeunea cucullata (Reinw. et al.) Grolle 1995. 

Microlejeunea saccula Steph. 1915 = M. africana Steph. 1888. 

Microlejeunea ulicina (Taylor) A.Evans CVI(37). 

Microlejeunea usambarensis Steph. TAN(2). 

Microlepidozia stephanii (Renauld ex Steph.) S.W.Arnell 1965, nom. inval. = Kurzia capillaris ssp. stephanii 

(Renauld) Pócs 1984. 

Microlepidozia succulenta (Sim) S.W.Arnell 1962 = Paracromastigum succulentum (Sim) J.J.Engel et 

G.L.S.Merr. 2001 (556). 

Microlepidozia tabularis (Steph.) S.W.Arnell 1962 = Kurzia capillaris (Sw.) Grolle 1964. 

Mnioloma fuscum (Lehm.) R.M.Schust. BIO(36), CMN(2), ETH(2), LES(2), MDG(1), REU(1,4), 

RWA(2), SAF(2), SEY(1,4), STH(2), SWZ(2), TAN(2), UGA(2), ZAI(2). 

Monocarpus sphaerocarpus D.J.Carr. SAF(2,76). 

Monoclea adglutinata Hook.f. et Taylor 1845 = Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche et al. 

1846. 

Monoclea gottschei Lindb. - erroneously recorded from Ethiopia. 

Mylia diversifolia (Gottsche) E.W.Jones ex S.W.Arnell 1955 = Leptoscyphus diversifolius (Gottsche) Grolle 

1963. 

Mylia expansa (Lehm.) S.W.Arnell 1955 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Mylia hedbergii S.W.Arnell 1956 = Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust. 1959. 

Mylia infuscata (Mitt.) S.W.Arnell 1956 = Leptoscyphus infuscatus (Mitt.) E.W.Jones 1963. 

Mylia iversenii (Pearson) S.W.Arnell 1955 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Nardia abyssinica (Nees) Cufod. 1952 = Jungermannia abyssinica Nees 1844. 

Nardia arnelliana Grolle REU(1), UGA(2). 

Nardia aurita (Lehm.) Trevis. 1877 = Anastrophyllum auritum (Lehm.) Steph. 1901. 

Nardia dusenii Steph. 1891 = Jungermannia dusenii (Steph.) Steph. 1901. 

Nardia exserta Steph. ex Besch. 1895, hom. inval. [non A.Evans 1891] = Jungermannia perloi (Gola) Váòa 

1973. 

Nardia jackii Steph. 1892 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1983. 

Nardia stolonifera Steph. 1892 = Jungermannia borgenii Gottsche ex Pearson 1893. 

Nardia verrucosa Steph. 1892 = Jungermannia dusenii (Steph.) Steph. 1901. 

Nesolejeunea intercalaris Herzog 1947 = Lejeunea acuta Mitt. 1864. 

Noteroclada porphyrorhiza (Nees) Mitt. 1855 = Fossombronia porphyrorhiza (Nees) Prosk. 1955. 

Notoscyphus africanus (Pearson) S.W.Arnell 1955 = Odontoschisma africanum (Pearson) Sim 1926.


Notoscyphus belangerianus (Lehm. et Lindenb.) Mitt. 1877 = N. lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Notoscyphus congoanus Steph. 1917 = N. lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Notoscyphus jackii (Steph.) Steph. 1901 = N. lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. BUR(2), CMN(2), COM(1,4), MAU(1), MDG(1,4), 

MLW(114), REU(1), RWA(2), SAF(2), ZAI(2). 

Notoscyphus natalensis Sim 1926 = N. lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Notoscyphus variifolius Mitt. 1877 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Notoscyphus vermicularis (Lehm.) Steph. 1901 = Clasmatocolea vermicularis (Lehm.) Grolle 1960. 

Notothylas angolensis Steph. 1917 = N. decurva (Mitt.) Steph. 1901. 

Notothylas decurva (Mitt.) Steph. ANG(2), CMN(2), GHA(2), NGA(2), SIE(2). Following Schuster 1992 

(141), N. decurva was treated in (2) as a synonym of N. orbicularis. However, Jones 1985 (104) described some 

observed differences between the two taxa, and concluded "on present evidence" that N. decurva should be 

maintained. It is unclear whether the two taxa have been compared in sufficient detail, and for that reason Jones 

(1985) is followed in this checklist, pending any further evidence presented to support synonymy with N. 

orbicularis. 

Notothylas flabellata Steph. ANG(2), CMN(2), TAN(2). 

Notothylas indica Kashyap GHA(2). 

Notothylas javanica (Sande Lac.) Gottsche IVO(2). 

Notothylas minuta (Mitt.) Steph. 1916 = Phaeoceros minutus (Mitt.) S.W.Arnell 1963. 

Notothylas sp. SIE(25). Jones (in litt.) considered the Sierra Leone plant may represent a new taxon, but that its 

status must remain uncertain until more copious material is available. 

Odonotoschisma ligulatum Steph. 1890 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Odontolejeunea hanningtonii (Mitt.) Steph. 1895 = Caudalejeunea hanningtonii (Mitt.) Schiffn. 1909. 

Odontolejeunea lunulata (F.Weber) Schiffn. BIO(2,33), BUR(2), CMN(2), COM(1), CON(2), ETH(2), 

GUI(2), IVO(2), KEN(2), MAU(1), MDG(1,4), MOZ(2), NGA(2), REU(2), RIM(10), RWA(2), 

SAO(2), SIE(2), TAN(2), UGA(42,50), ZAI(2). 

Odontolejeunea mauritiana Steph. 1912 = O. lunulata (F.Weber) Schiffn. 1893. 

Odontolejeunea sieberiana (Gottsche) Schiffn. 1893 = O. lunulata (F.Weber) Schiffn. 1893. 

Odontolejeunea sieberiana var. africana Steph. 1892, nom. inval. = O. lunulata (F.Weber) Schiffn. 1893. 

Odontolejeunea thomeensis Steph. 1912 = O. lunulata (F.Weber) Schiffn. 1893. 

Odontolejeunea tortuosa (Lehm. et Lindenb.) Steph. 1912 = O. lunulata (F.Weber) Schiffn. 1893. 

Odontoschisma africanum (Pearson) Sim MAU(1), MLW(114), REU(45), SAF(2), TAN(2). 

Odontoschisma denudatum var. africanum (Pearson) S.W.Arnell 1957, nom. inval. = O. africanum (Pearson) 

Sim 1926. 

Odontoschisma ligulatum Steph. 1890 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Odontoschisma variabile Sim 1926, hom. illeg. [non (Lindenb. et Gottsche) Trevis. 1877] = Clasmatocolea 

vermicularis (Lehm.) Grolle 1960. 

Omphalanthus filiformis (Sw.) Nees 1845 [Dubia et Excludenda]. 

Omphalanthus geminiflorus (Nees) Gottsche et al. 1845 = O. filiformis (Sw.) Nees 1845 [Dubia et Excludenda]. 

Omphalanthus renistipulus Steph. 1911, hom. illeg. [non Lindenb. 1845] = Evansiolejeunea roccatii (Gola) 

Vanden Berghen 1960. 

Omphalanthus roccatii (Gola) R.M.Schust. 1963 = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960. 

Omphalolejeunea filiformis (Sw.) Lacout. 1908 = Omphalanthus filiformis (Sw.) Nees 1845 [Dubia et 

Excludenda]. 

Otigoniolejeunea villaumei Steph. 1914 = Lejeunea villaumei (Steph.) Grolle 1977. 

Otiona aitonia (Lindenb. et Nees) Mitt. 1877 = Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Otolejeunea moniliata Grolle MDG(1). 

Otolejeunea rabenorii Tixier MDG(1). 

Oxymitra cristata Garside ex Perold NAM(2), SAF(2). 

Pallavicinia attenuata Steph. 1893 = P. lyellii (Hook.) Carruth. 1865. 

Pallavicinia byssophora (Lehm. et Lindenb.) Trevis. 1877 = P. lyellii (Hook.) Carruth. 1865. 

Pallavicinia camisassai Gola KEN(120). 

Pallavicinia capensis S.W.Arnell 1954 = Symphyogyna brasiliensis Nees et Mont. 1836. 

Pallavicinia lyellii (Hook.) Carruth. BUR(2), LES(75), MLW(2,29), REU(1), RWA(2), SAF(2,75), 

SAO(2), TAN(2), UGA(50,96), ZAI(2), ZIM(2). 

Pallavicinia pilifera Steph. 1891 = P. lyellii (Hook.) Carruth. 1865.


Pallavicinia procumbens (Hook.f. et Taylor) Mitt. 1875 = Symphyogyna brasiliensis Nees et Mont. 1836. 

Pallavicinia rubescens Steph. 1917 = Symphyogyna brasiliensis Nees et Mont. 1836. 

Pallavicinia serrata (Mitt.) Grolle 1984 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986. 

Pallavicinia spinosa (Lindenb. et Gottsche) Grolle 1979 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986. 

Pallavicinia stephanii J.B.Jack 1892 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986. 

Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. SAF(2,556). 

Paracromastigum tristanianum (R.M.Schust.) R.M.Schust. SAF(2,504). Following the redefinition of 

Hyalolepidozia, South African records of H. bicuspidata (C.Massal.) Grolle have been reassigned by Schuster 

(504) to P. tristanianum. 

Paraschistochila engleriana (Steph.) R.M.Schust. 1971 = Gottschea engleriana (Steph.) Grolle et Zijlstra 1984. 

Paraschistochila neesii (Mont.) R.M.Schust 1971 = Gottschea neesii Mont. 1843. 

Pedinolejeunea himalayensis (Pandé et Misra) Chen et Wu 1964 = Cololejeunea latilobula (Herzog) Tixier 

1985. 

Phaeoceros bolusii (Sim) S.W.Arnell SAF(2), ZIM(2). 

Phaeoceros bulbiculosus (Brot.) Prosk. CMN(2), ERI(2). 

Phaeoceros carolinianus (Michx.) Prosk. ANG(2), CMN(2), CVI(2,37), ETH(2), KEN(2), LES(75), 

MDG(1), MLW(2,114), NAM(2), NGA(2), REU(1), RWA(2), SAF(2,75), SIE(2), STH(2,49), TAN(2), 

UGA(2,96), ZAI(2), ZIM(2). 

Phaeoceros fulvisporus (Steph.) J.Haseg. RWA(2), SAF(2), TAN(2). 

Phaeoceros jollyanus (Steph.) S.W.Arnell BIO(2,33). 

Phaeoceros laevis (L.) Prosk. 1951 [Dubia et Excludenda]. 

Phaeoceros laevis ssp. carolinianus (Michx.) Prosk. 1954 = P. carolinianus (Michx.) Prosk. 1951 (683). 

Phaeoceros maranguensis (Steph.) Bapna TAN(2). 

Phaeoceros minutus (Mitt.) S.W.Arnell SAF(2), ZIM(2). 

Phaeoceros sp. (Anthoceros aethyopicus Gola) ETH(2). Probably a species of Phaeoceros as Gola 1914b 

(121) described it as having "frons haud cavernosa". 

Phaeoceros sp. (Anthoceros brunnthaleri Steph.) TAN(2). Probably a species of Phaeoceros as Stephani 

1916 (140) described it as having "frons solida ... sporae pallidae". 

Phaeoceros sp. (Anthoceros mildbraedii Steph.) CAM(2). Probably a species of Phaeoceros, as Stephani 

1923 (139) described it as having "frons solida ... sporae pallidae". E.W. Jones (in litt. ca. 1990) considered it is 

probably synonymous with Phaeoceros carolinianus (Michx.) Prosk. 

Phaeoceros sp. (Anthoceros parvifrons Steph.) TAN(2). Probably a species of Phaeoceros as Stephani 1912 

(138) described it as having "frons solida ... sporae pallidae". 

Phaeoceros sp. (Anthoceros pseudocostus Steph.) MDG(1). The spores are described as "pallidae" (Stephani 

1916 (140)), hence probably a Phaeoceros species. 

Phaeoceros sp. (Anthoceros rossoi Gola) KEN(2). Probably a species of Phaeoceros as it is described as 

"frons haud cavernosa ... sporae flavescentes" (Gola 1914a (120)). 

Phaeoceros sp. (Anthoceros tenuissimus Steph.) TAN(2). West Africa (Lobango)(2). Probably a species of 

Phaeoceros as the spores are described as "flavidae" (Stephani 1923 (139)). 

Phaeoceros sp. (Anthoceros tigrinus Gola) ETH(2). Gola 1914b (121) described it as having "frons haud 

cavernosa ... sporae flavae", so it is probably a species of Phaeoceros. 

Phragmicoma abnormis Gottsche 1882 [Dubia et Excludenda]. 

Phragmicoma acutiloba (Hook.f. et Taylor) Gottsche 1847 = Marchesinia brachiata (Sw.) Schiffn. 1893. 

Phragmicoma amplectens Steph. 1886 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Phragmicoma aulacophora Mont. 1843 = Acrolejeunea aulacophora (Mont.) Steph. 1895. 

Phragmicoma carinata Mitt. 1879 = Mastigolejeunea auriculata (Wils.) Schiffn. 1893. 

Phragmicoma emergens Mitt. 1879 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Phragmicoma excavata (Mitt.) Steph. 1886 = Marchesinia excavata (Mitt.) Schiffn. 1893. 

Phragmicoma florea Mitt. 1886 = Mastigolejeunea florea (Mitt.) Paris 1906. 

Phragmicoma fulva Gottsche 1882 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Phragmicoma inflexa Gottsche 1882 [Dubia et Excludenda]. 

Phragmicoma microscypha (Hook.f. et Taylor) Mitt. 1875 = Leucolejeunea xanthocarpa (Lehm. et Lindenb.) 

A.Evans 1907. 

Phragmicoma molleri Steph. 1886 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Phragmicoma palisoti Taylor ex Mitt. 1886, nom. nud. = Odontolejeunea lunulata (F.Weber) Schiffn. 1893.


Phragmicoma pappeana Nees 1845 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Phragmicoma reniloba Gottsche 1845 = Caudalejeunea reniloba (Gottsche) Steph. 1912 [Dubia et Excludenda]. 

Phragmicoma rotalis (Hook.f. et Taylor) Mitt. 1875 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et 


Phragmicoma sagraeana Mont. 1842 = Lopholejeunea subfusca (Nees) Schiffn. 1897. 

Phragmicoma sphaerophora (Lehm. et Lindenb.) Schiffn. 1886 = Lopholejeunea sphaerophora (Lehm. et 

Lindenb.) Steph. 1912. 

Phragmilejeunea molleri (Steph.) R.M.Schust. 1961 = Schiffneriolejeunea polycarpa (Steph.) Gradst. 1974. 

Phragmilejeunea occulta (Steph.) Bischl. 1965, nom. inval. = Schiffneriolejeunea occulta (Steph.) Gradst. 1974. 

Phragmilejeunea pappeana (Nees) R.M.Schust. 1954 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Physiotium sphagnoides Richard ex Nees 1838, nom. illeg. = Pleurozia gigantea (F.Weber) Lindb. 1874. 

Physocolea adhaesiva (Mitt.) Steph. 1916 = Cololejeunea adhaesiva (Mitt.) R.M.Schust. 1963. 

Physocolea africana Steph. 1916 = Cololejeunea africana (Steph.) R.M.Schust. 1963. 

Physocolea autoica Steph. 1916 = Cololejeunea autoica (Steph.) Grolle 1995. 

Physocolea bidentula Steph. 1916 = Cololejeunea bidentula (Steph.) E.W.Jones 1954. 

Physocolea bolombensis Steph. 1916 = Cololejeunea tonkinensis Steph. 1895. 

Physocolea cristata Steph. 1916 = Cololejeunea cristata (Steph.) R.M.Schust. 1963. 

Physocolea cuneata (Lehm. et Lindenb.) Steph. 1916 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947. 

Physocolea decemplicata Steph. 1916 = Cololejeunea decemplicata (Steph.) Tixier 1979. 

Physocolea elegans (Steph.) Steph. 1916 = Cololejeunea elegans Steph. 1891. 

Physocolea erectifolia (Steph.) Steph. 1916 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947. 

Physocolea filicaulis (Steph.) Steph. 1916 = Cololejeunea hildebrandii (Austin) Steph. 1897. 

Physocolea flegelii Steph. 1916 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897. 

Physocolea grossidens (Steph.) Steph. 1916 = Cololejeunea malanjae Steph. 1902. 

Physocolea malanjae Steph. 1916 = Cololejeunea malanjae Steph. 1902. 

Physocolea myriantha Herzog 1952 = Cololejeunea minutissima ssp. myriocarpa (Nees et Mont.) R.M.Schust. 

1955. 

Physocolea peculiaris Herzog 1931 = Cololejeunea inflectens (Mitt.) Benedix 1953. 

Physocolea punctata (Gottsche) Steph. 1916 = Cololejeunea punctata (Gottsche) Pearson 1892 [Dubia et 

Excludenda]. 

Physocolea pusilla (Steph.) Steph. 1916 = Cololejeunea pusilla Steph. 1895. 

Physocolea scabrifolia Schiffn. ex Steph. 1916 = Cololejeunea obliqua (Nees et Mont.) Schiffn. 1897. 

Physocolea subciliata (Mitt.) Steph. 1916 = Cololejeunea cuneata (Lehm. et Lindenb.) Herzog 1947. 

Physocolea villaumei Steph. 1916 = Cololejeunea hildebrandii (Austin) Steph. 1897. 

Physocolea vittata Steph. 1916, perhaps = Cololejeunea platyneura (Spruce) A.Evans 1902. 

Physocolea xavieri Lacout. ex Steph. 1916 = Cololejeunea xavieri (Lacout. ex Steph.) E.W.Jones 1954. 

Plagiochasma abyssinicum Gola 1914 = P. rupestre var. rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Plagiochasma aitonia Lindenb. et Nees 1836 = Plagiochasma rupestre var. rupestre (J.R.Forst. et G.Forst.) 

Steph. 1898. 

Plagiochasma appendiculatum Lehm. et Lindenb. ERI(2), KEN(2), SAF(2), SOC(2), TAN(2), ZIM(2). 

Plagiochasma appendiculatum var. erythraeum Gola 1914 = P. appendiculatum Lehm. et Lindenb. 1832. 

Plagiochasma beccarianum Steph. ERI(2), MLW(29), NAM(2), SAF(2), SOC(2), TAN(2), UGA(115), 

ZAM(2). 

Plagiochasma capense Sim 1926 = P. rupestre var. rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Plagiochasma dinteri Steph. 1901 = P. microcephalum (Steph.) Steph. 1898. 

Plagiochasma dschallanum Steph. 1898 = P. rupestre var. rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Plagiochasma eximium (Schiffn. in Steph.) Steph. CMN(2), CVI(2,37), DJI(2), ERI(2), ETH(2,113), 

GUI(2), KEN(2), REU(1), SAF(2,75), SIE(2), SOC(2), TAN(2), UGA(2,96), ZAI(2). The collection from 

Malawi (Crundwell 369), has been re-determined by T. Pócs as P. beccarianum. 

Plagiochasma fischerianum (Steph.) Steph. 1898 ("P. fischeri") = P. appendiculatum Lehm. et Lindenb. 1832. 

Plagiochasma limbatum (Hook.f. et Taylor) Nees 1846 = P. rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Plagiochasma microcephalum (Steph.) Steph. ANG(2), ETH(2,113), MDG(1), NAM(2), SAF(2), 

TAN(2), UGA(2). 

Plagiochasma muricatum Steph. 1901 = P. rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph. CHA(3), CVI(37), LES(75), MAU(1), REU(1), 

SAF(75), UGA(42).


Plagiochasma rupestre var. rupestre (J.R.Forst. et G.Forst.) Steph. ANG(2), ASC(2), CHA(2), 

CVI(2,534), DJI(2), ERI(2), ETH(2,113), KEN(2), LES(2), NAM(2), SAF(2), STH(2), SUD(2), 

TAN(2), UGA(2), ZAI(2), ZIM(2). 

Plagiochasma rupestre var. volkii Bischl. LES(2,75), NAM(2), SAF(2), ZIM(2). 

Plagiochasma schimperi Steph. 1898 = P. eximium (Schiffn.) Steph. 1898. 

Plagiochasma tenue Steph. 1898 = P. rupestre var. rupestre (J.R.Forst. et G.Forst.) Steph. 1898. 

Plagiochila abyssinica Mitt. 1886 = P. squamulosa var. squamulosa Mitt. 1863. 

Plagiochila africana Steph. BIO(2,33), CAF(2), CMN(2), GHA(2), GUI(48), IVO(2,53), NGA(2), 

RIM(10), SIE(2). 

Plagiochila aloysii-sabaudiae Gola 1907 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864. 

Plagiochila amphigastriata Herzog 1938 = P. pinniflora Steph. 1891. 

Plagiochila amplifolia Steph. 1901, perhaps = P. kiaeri Gottsche 1882. 

Plagiochila angolensis Steph. ex Paris, nom. inval. ANG(2). 

Plagiochila angusta Lindenb. MDG(1), REU(1). 

Plagiochila angustifolia Steph. 1913 = P. strictifolia Steph. 1891. 

Plagiochila angustissima Steph. 1902 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila angustitexta Steph. MLW(114), TAN(2). 

Plagiochila anjouana Gottsche ex Dugas 1928 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila ankefinensis Steph. 1904 = P. kiaeri Gottsche 1882. 

Plagiochila armata Steph. 1891 = P. barteri var. barteri Mitt. 1886. 

Plagiochila attenuata Steph. 1911 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864. 

Plagiochila bamingensis Steph. CAF(2). 

Plagiochila barteri Mitt. BIO(2,33,36), CMN(2), COM(1), ETH(2), GAB(2), KEN(2,71), MAU(1), 

MDG(1), MLW(2,114), PRI(2), REU(1), RWA(2), SAO(2), SIE(2), TAN(2), TOG(2), UGA(96), 

ZAI(2), ZIM(2). 

Plagiochila barteri ssp. colorans (Steph.) Pócs ETH(2), KEN(2,71), RWA(2), TAN(2), UGA(2), ZAI(2). 

Plagiochila barteri var. valida (Steph.) Vanden Berghen ETH(2), MLW(2), REU(1). 

Plagiochila batangensis Steph. 1917 = P. praemorsa Steph. 1886. 

Plagiochila berthieui Steph. 1893 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila bidentula Steph. ANN(2,33), CMN(2). 

Plagiochila blanda Steph. 1911 = P. integerrima Steph. 1886. 

Plagiochila boivinii Steph. MDG(1), REU(1). 

Plagiochila bomanensis Steph. 1891 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila borbonica Mont. et Gottsche 1856 = P. pectinata Willd. ex Lindenb. 1839. 

Plagiochila borgenii Gottsche 1882, nom. inval. = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila boryana Gottsche ex Steph. MAU(1), MDG(1), REU(1,4), TAN(2), UGA(42). 

Plagiochila bovonei Gola 1920 = P. fusifera Taylor 1846. 

Plagiochila breviramea Steph. 1911 = P. squamulosa Mitt. 1863. 

Plagiochila brevisedens Steph. 1918 = P. kiaeri Gottsche 1882. 

Plagiochila brunneola Steph. ETH(2), SAO(2). 

Plagiochila brunnthaleri Steph. 1913 = P. effusa Steph. 1895. 

Plagiochila bueensis Steph. 1904 = P. moenkemeyeri Steph. 1904. 

Plagiochila buettneriana Steph. 1895 = P. strictifolia Steph. 1891. 

Plagiochila bursata (Desv.) Lindenb. 1840 [Dubia et Excludenda]. 

Plagiochila cacuminis Steph. 1918 = P. praemorsa Steph. 1887. 

Plagiochila calva (Nees) Lindenb. 1839 = P. javanica (Sw.) Dumort. 1836. 

Plagiochila cambouena Steph. 1904, nom. inval. = P. cambuena Steph. 1890. 

Plagiochila cambuena Steph. IVO(53), MDG(1), MLW(114), TAN(2). 

Plagiochila camerunensis Steph. 1902 = P. africana Steph. 1902. 

Plagiochila candelabra Steph. 1904 = P. pinniflora Steph. 1891. 

Plagiochila capensis Steph. 1904 = P. kiaeri var. capensis (Steph.). 

Plagiochila carduifolia Steph. 1904 = P. africana Steph. 1902. 

Plagiochila chenagonii Steph. 1892 = P. angusta Lindenb. 1843. 

Plagiochila chiovendae Gola ETH(2). 

Plagiochila circumvoluta Gerola ETH(2).


Plagiochila clavaeflora Steph. 1891 = P. integerrima Steph. 1886. 

Plagiochila collicalyx Steph. 1904 = P. flabellata Steph. 1886. 

Plagiochila colorans Steph. 1911 = Plagiochila barteri ssp. colorans (Steph.) Pócs 1993. 

Plagiochila comorensis Steph. 1888 = P. pectinata Willd. ex Lindenb. 1839. 

Plagiochila comorensis fo. johannensis Herzog 1947, nom. inval. = P. terebrans Nees et Mont. ex Lindenb. 

1840. 

Plagiochila congoana Steph. 1918 = P. neckeroidea Mitt. 1860. 

Plagiochila corniculata auct. [non Dumort. 1835] = P. exigua (Taylor) Taylor 1846. 

Plagiochila corymbulosa Pearson 1886 = P. heterostipa Steph. 1892. 

Plagiochila crispi-crista Steph. 1904 = P. squamulosa var. squamulosa Mitt. 1863. 

Plagiochila crispulo-caudata Gottsche 1882 = P. squamulosa var. crispulo-caudata (Gottsche) Vanden Berghen 

1981. 

Plagiochila cristato-dentata Steph. 1904 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864. 

Plagiochila crollii Steph. 1904 = P. incerta Gottsche 1857. 

Plagiochila cumminsiana Steph. 1918 = P. praemorsa Steph. 1886. 

Plagiochila curvatifolia Steph. 1886 = P. neckeroidea Mitt. 1860. 

Plagiochila decurrens Steph. 1901 = P. moenkemeyeri Steph. 1904. 

Plagiochila dichotoma (P.Beauv.) Nees et Mont. NGA(2). 

Plagiochila dichotoma var. elongata Lindenb. ex Gottsche et al. 1844 = P. paucidentata Mont. et Gottsche 

1856. 

Plagiochila dichotoma var. laxa Gottsche et al. 1844 = P. integerrima Steph. 1886. 

Plagiochila dicrana Mitt. 1886 = P. drepanophylla Sande Lac. 1857. 

Plagiochila divergens Steph. 1891 = P. kiaeri Gottsche 1882. 

Plagiochila drepanophylla Sande Lac. MDG(1), REU(1), TAN(2). 

Plagiochila dschaggana Steph. 1891 = P. squamulosa var. sinuosa (Mitt.) Vanden Berghen 1981. 

Plagiochila effusa Steph. ANG(2), BIO(33,72), KEN(2), MLW(114), RIM(10), TAN(2), UGA(2). 

Plagiochila elgonensis Pearson 1924 = P. integerrima Steph. 1886. 

Plagiochila elongata Gottsche ex Geissler et Bischl. 1989, nom. inval. = P. paucidentata Mont. et Gottsche 

1856. 

Plagiochila emarginata Mont. et Gottsche 1856 = Adelanthus decipiens (Hook.) Mitt. 1864. 

Plagiochila engleriana Steph. 1895 = P. lastii Mitt. 1886. 

Plagiochila ericicola Steph. KEN(2), RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Plagiochila erythraeae Herzog ERI(2). 

Plagiochila esterhuysenii S.W.Arnell 1954, probably = P. squamulosa var. crispulo-caudata (Gottsche) Vanden 

Berghen 1981. 

Plagiochila evansii Steph. 1904 = P. kiaeri Gottsche 1882. 

Plagiochila exigua (Taylor) Taylor ETH(2), KEN(2), LES(75), MLW(114), REU(1), SAF(2,75), 

TAN(2), UGA(42), ZAI(2,667). 

Plagiochila expallescens Steph. 1904 = P. kiaeri Gottsche 1882. 

Plagiochila filicicola Steph. UGA(2). 

Plagiochila filiformis Gottsche ex Dugas 1928 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila fissicalyx Steph. 1902 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila flabellata Steph. CMN(2), REU(1), SAO(2), SEY(1), TAN(2). 

Plagiochila flagellaris Steph. ex Renauld 1909, nom. inval. = P. integerrima Steph. 1886. 

Plagiochila fragilifolia Pearson 1922 = P. praemorsa Steph. 1886. 

Plagiochila frapieri Dugas 1928 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila frappieri Gottsche ex Besch. 1895, nom. inval. = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila frondescens (Nees) Lindenb. ETH(2). 

Plagiochila furcata Steph. 1892 = P. angusta Lindenb. 1843. 

Plagiochila fuscovirens Steph. 1918, probably = P. squamulosa var. crispulo-caudata (Gottsche) Vanden 

Berghen 1981. 

Plagiochila fusifera Taylor CAF(2), CMN(2), ETH(2), GAB(2), GHA(2), GUI(2), IVO(2,53), KEN(2), 

MLW(2,114), NGA(2), RWA(2), SEN(2), SIE(2), SOM(2), TAN(2), TOG(2,101), UGA(2), ZAI(2), 

ZAM(2). 

Plagiochila aff. fusifera Taylor SEY(4). 

Plagiochila gaffatensis Gottsche ETH(2).


Plagiochila gasangorii Gola 1914 = P. lastii Mitt. 1886. 

Plagiochila gentiliana Steph. 1902 = P. neckeroidea Mitt. 1860. 

Plagiochila gibbiflora Steph. SAO(2). 

Plagiochila gottscheana Lindenb. ex Lehm. 1844 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Plagiochila granditexta Steph. MDG(1). 

Plagiochila grossealata Steph. 1918, perhaps = P. moenkemeyeri Steph. 1904. 

Plagiochila haumannii Herzog KEN(21), MOZ(2), SAF(2), UGA(2), ZAI(2). 

Plagiochila hedbergii S.W.Arnell 1956 = P. ericicola Steph. 1904. 

Plagiochila henriquesii Steph. 1904, perhaps = P. heterostipa Steph. 1892 or P. kiaeri Gottsche 1882. 

Plagiochila heterostipa Steph. ANG(2), BUR(2), GAB(2), IVO(53), KEN(2), MLW(2,114), RWA(2), 

SAF(2), SWZ(2), UGA(50,96), ZAI(2), ZIM(2). 

Plagiochila heudelotiana Steph. 1904 = P. fusifera Taylor 1846. 

Plagiochila heudelotiana Steph. ex Paris 1902, nom. inval. = P. fusifera Taylor 1846. 

Plagiochila hildebrandtii Steph. 1902 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila hochstetteri Nees ex Lindenb. 1843, perhaps = P. squamulosa Mitt. 1863. 

Plagiochila imerinensis Steph. 1904 = P. kiaeri Gottsche 1882. 

Plagiochila incerta Gottsche MAU(1), MDG(1), REU(1). 

Plagiochila injasutiensis S.W.Arnell SAF(2,75). 

Plagiochila insularia Mitt. STH(2). 

Plagiochila integerrima Steph. BIO(2,33,36,72), BUR(2), CAF(2), CMN(2), CON(2), GAB(2), GHA(2), 

IVO(2,53), KEN(2), MAU(1), MDG(1), MLW(114), NGA(2), REU(1), RIM(10), RWA(2), SAO(2), 

SEY(1), SIE(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Plagiochila javanica (Sw.) Nees et Mont. 1836 [Dubia et Excludenda]. 

Plagiochila johannensis Steph. 1902 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila jollyana Steph. 1902 = P. neckeroidea Mitt. 1860. 

Plagiochila jungneri Steph. 1904 = P. barteri Mitt. 1886. 

Plagiochila kamerunensis Steph. 1918 = P. strictifolia Steph. 1891. 

Plagiochila katangae Gola 1920 = P. squamulosa var. crispulo-caudata (Gottsche) Vanden Berghen 1981. 

Plagiochila kiaeri Gottsche ANG(2), BUR(2), KEN(2), MDG(1), MLW(2,114), RWA(2), SEY(4), 

TAN(2), UGA(2), ZAI(2). 

Plagiochila kiaeri var. capensis (Steph.) ANG(2), SAF(2,75), ZIM(2). 

Plagiochila kiaeri var. myriocarpa (Pearson) ETH(2), KEN(2,21), MLW(2,114), RWA(2), TAN(2), 

UGA(2,96), ZAI(2), ZAM(2). 

Plagiochila knysnana S.W.Arnell 1954 = Leptoscyphus expansus (Lehm.) Grolle 1963. 

Plagiochila laevifolia Gola KEN(2). 

Plagiochila lastii Mitt. ANG(2), ETH(2), KEN(2,71), MLW(2,114), RWA(2), SAF(2,75), TAN(2), 

UGA(2,96), ZAI(2), ZIM(2). 

Plagiochila laxifolia Gottsche 1882 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila lecontei Steph. Congo francais(2). 

Plagiochila ledieui Steph. Congo(2). 

Plagiochila lenormandii Gottsche 1882, nom. inval. = P. repanda (Schwägr.) Lindenb. 1840. 

Plagiochila ligulata Steph. 1904 = P. repanda (Schwägr.) Lindenb. 1840. 

Plagiochila liliputana Herzog 1947 = P. rodriguezii Steph. 1890. 

Plagiochila lindblomii Pearson 1924, probably = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila lindenbergiana (Lehm.) Gottsche et al. 1844 = Adelanthus lindenbergianus (Lehm.) Mitt. 1864. 

Plagiochila loloensis Steph. BIO(33), CMN(2), NGA(2), SAO(2). 

Plagiochila longicauda Steph. 1921 = P. pinniflora Steph. 1891. 

Plagiochila lophophora R.M.Schust. 1959 = P. exigua (Taylor) Taylor 1846. 

Plagiochila lunata S.W.Arnell LES(75), SAF(2). 

Plagiochila lurida Steph. 1911 = P. squamulosa Mitt. 1863. 

Plagiochila macrostachya Lindenb. 1840 [Dubia et Excludenda]. 

Plagiochila madagascariensis Gottsche 1882, nom. inval. [non Steph. 1904] = P. repanda (Schwägr.) Lindenb. 

1840. 

Plagiochila madagascariensis Steph. 1904 [nec Gottsche 1882] = P. cambuena Steph. 1890. 

Plagiochila maranguana Steph. 1895 = P. kiaeri Gottsche 1882.


Plagiochila mascarena Gottsche 1857 = P. paucidentata Mont. et Gottsche 1856. 

Plagiochila mauritiana (Nees) Lindenb. 1840 = P. repanda (Schwägr.) Lindenb. 1840. 

Plagiochila mediocris Dugas 1928 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila moenkemeyeri Steph. BIO(2,33), CMN(2), ETH(2), GHA(2), NGA(2), RIM(10), SAO(2), 

SIE(2), TAN(2), ZAI(2). 

Plagiochila molleri Steph. 1886, perhaps = P. flabellata Steph. 1886. 

Plagiochila moschiensis Steph. 1904, perhaps = P. strictifolia Steph. 1891 or P. moenkemeyeri Steph. 1904. 

Plagiochila multiflora Steph. TAN(2). 

Plagiochila multispica Steph. 1905 = P. moenkemeyeri Steph. 1904. 

Plagiochila myriocarpa Pearson 1924 = P. kiaeri var. myriocarpa (Pearson). 

Plagiochila natalensis Pearson 1886, perhaps = P. heterostipa Steph. 1892. 

Plagiochila naveauiana Pearson 1922 = P. integerrima Steph. 1886. 

Plagiochila neckeroidea Mitt. BIO(36), CMN(2), CON(2), GAB(2), GHA(2), IVO(2,53), LBR(2), 

NGA(2), RIM(10), SIE(2), UGA(42), ZAI(2). 

Plagiochila negrii Gola ETH(2). 

Plagiochila nemophila Gottsche 1882 = P. incerta Gottsche 1857. 

Plagiochila noditexta Steph. 1904 = P. repanda var. perrotana (Steph.) Vanden Berghen 1981. 

Plagiochila nudicaulis Steph. 1895 = P. integerrima Steph. 1886. 

Plagiochila nudiuscula Gottsche 1882 = P. repanda (Schwägr.) Lindenb. 1840. 

Plagiochila oblongotrigona Steph. 1921 = P. effusa Steph. 1895. 

Plagiochila ovatotrigona Steph. 1904 = P. moenkemeyeri Steph. 1904. 

Plagiochila pallida Gottsche 1882 = P. angusta Lindenb. 1843. 

Plagiochila pallidevirens Steph. 1921 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila paucidentata Mont. et Gottsche MAS(1), MAU(1), MDG(1), REU(1). 

Plagiochila pearsonii Steph. 1921, perhaps = P. kiaeri Gottsche 1882. 

Plagiochila pectinata Willd. ex Lindenb. BIO(72), CMN(2), COM(1,4), GHA(2), MAS(1), MAU(1), 

MDG(1), MLW(114), REU(1), RIM(10), SAO(2), SEY(1,4), SIE(2), UGA(50), ZAI(2). 

Plagiochila perflorens Gola 1920 = P. squamulosa var. crispulo-caudata (Gottsche) Vanden Berghen 1981. 

Plagiochila perloi Gola 1914 = P. squamulosa var. sinuosa (Mitt.) Vanden Berghen 1981. 

Plagiochila perrotana Steph. 1904 = P. repanda (Schwägr.) Lindenb. 1840. 

Plagiochila pinniflora Steph. CMN(2), CON(2), GAB(2), GHA(2), IVO(2), NGA(2), RIM(10), SIE(2), 

TOG(2), UGA(2,42), ZAI(2). 

Plagiochila pomai Pearson 1922 = P. neckeroidea Mitt. 1860. 

Plagiochila praemorsa Steph. ANN(2,33), BIO(2,33,36), CAF(2), CMN(2), CON(2), GAB(2), GHA(2), 

IVO(2,53), NGA(2), RIM(10), SAO(2), SIE(2), UGA(2), ZAI(2), ZAM(2). 

Plagiochila pseudoattenuata S.W.Arnell KEN(2), LES(75), MDG(1), MLW(2), RWA(2), SAF(2,75), 

TAN(2), UGA(2), ZAI(2). Includes the record from Madagascar listed under P. subalpina in (1). 

Plagiochila punctata Taylor BIO(2,33). 

Plagiochila quintasii Steph. 1904 = P. barteri Mitt. 1886. 

Plagiochila radicans Steph. 1902 = P. integerrima Steph. 1886. 

Plagiochila ragazzii Gola ERI(121), ETH(2). 

Plagiochila ramentacea Steph. 1921 = P. pinniflora Steph. 1891. 

Plagiochila refracta Brid. ex Gottsche 1857, nom. inval. = P. paucidentata Mont. et Gottsche 1856. 

Plagiochila renauldii Steph. MAU(1), REU(1). 

Plagiochila repanda (Schwägr.) Lindenb. KEN(2), MAU(1), MDG(1), MLW(2), REU(1). 

Plagiochila repanda var. perrotana (Steph.) Vanden Berghen COM(1), MDG(1). 

Plagiochila replicatula Steph. 1902 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila richardiana Steph. 1902 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila rivana Steph. 1921 = P. fusifera Taylor 1846 or P. moenkemeyeri Steph. 1904. 

Plagiochila rivularis Steph. 1921 = P. integerrima Steph. 1886. 

Plagiochila rodriguezii Steph. COM(1), MDG(1), REU(1). 

Plagiochila rotundifolia Steph. 1904, hom. illeg. [non Colenso 1888] = P. loloensis Steph. 1904. 

Plagiochila rubricaulis Steph. ZAI(2,159). 

Plagiochila rudolfii Pócs TAN(2). 

Plagiochila runssorensis Steph. 1895 = P. kiaeri Gottsche 1882.


Plagiochila rupicola Steph. 1895 = P. integerrima Steph. 1886. 

Plagiochila ruspoliana Steph. 1904 = P. squamulosa Mitt. 1863. 

Plagiochila rutenbergiana Gottsche 1882 = P. repanda (Schwägr.) Lindenb. 1840. 

Plagiochila rutenbergii Gottsche 1882, nom. inval. = P. repanda (Schwägr.) Lindenb. 1840. 

Plagiochila ruwenzorensis Steph. 1911 = Syzygiella geminifolia (Mitt.) Steph. 1902. 

Plagiochila ruwenzoris Steph. 1921 = P. barteri ssp. colorans (Steph.) Pócs 1993. 

Plagiochila salvadorica Steph. ANG(2), CAF(2), GHA(2), GUI(48), IVO(53), SIE(2), ZAI(2). 

Plagiochila sarmentosa (Lehm. et Lindenb.) Lindenb. SAF(2). 

Plagiochila schimperiana Steph. 1895 = P. squamulosa var. squamulosa Mitt. 1863. 

Plagiochila securifolia Nees in Lindenb. 1840 = Syzygiella securifolia (Nees in Lindenb.) Inoue 1979. 

Plagiochila sikorae Steph 1893 = P. barteri Mitt. 1886. 

Plagiochila simplex (Sw.) Lindenb. 1840 [Dubia et Excludenda]. 

Plagiochila sinuosa Mitt. 1886 = P. squamulosa var. sinuosa (Mitt.) Vanden Berghen 1981. 

Plagiochila socorana Gerola 1947, probably = P. squamulosa var. squamulosa Mitt. 1863. 

Plagiochila sparsa Steph. 1903 = P. kiaeri Gottsche 1882. 

Plagiochila sprengeri Steph. 1904 = P. lastii Mitt. 1886. 

Plagiochila squamulosa Mitt. BIO(2,33), BUR(2), CMN(2), DJI(2), ERI(2), ETH(2), KEN(2,21,71,665), 

MLW(114), RWA(2), SOM(2), TAN(2), UGA(2,96), ZAI(2), ZAM(2). 

Plagiochila squamulosa var. crispulo-caudata (Gottsche) Vanden Berghen CMN(2), ETH(2), 

KEN(2,21,71,665), LES(75), MDG(1), MLW(2,114), MOZ(2), REU(1), RWA(2), SAF(2,75), TAN(2), 

UGA(50), ZAI(2), ZAM(2), ZIM(2). 

Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen ANG(2), BUR(2), ETH(2), 

KEN(2,21,71,665), MAU(1), MDG(1), MLW(2,114), REU(1), RWA(2), TAN(2), UGA(2,96), ZAI(2), 

ZAM(2), ZIM(2). 

Plagiochila squarrosifolia Steph. 1921 = P. integerrima Steph. 1886. 

Plagiochila stahlii Steph. 1921 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila strictifolia Steph. ANG(2), ANN(2,33), BIO(2), CAF(2), CMN(2), GHA(2), IVO(2,53), 

MLW(2,114), NGA(2), PRI(2), SAO(2), SIE(2), TAN(2), TOG(2,101), UGA(2,96), ZAI(2), ZAM(2). 

Plagiochila stuhlmannii Steph. 1895 = P. kiaeri Gottsche 1882. 

Plagiochila subalpina Steph. 1891, hom. illeg. [non (Nees ex Lindenb.) Mont. et Nees 1838] = P. 

pseudoattenuata S.W.Arnell 1962. 

Plagiochila subcrispula Steph. 1921 = P. africana Steph. 1902. 

Plagiochila subintegerrima (Reinw. et al.) Mont. et Nees 1839 = Syzygiella subintegerrima (Reinw. et al.) 

Spruce 1895 [Dubia et Excludenda]. 

Plagiochila sublinearis Gottsche 1882 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila subrepanda Steph. 1921 = P. pinniflora Steph. 1891. 

Plagiochila subtenuis Steph. 1902 [Dubia et Excludenda]. 

Plagiochila superba (Nees ex Spreng.) Mont. et Nees 1839 [Dubia et Excludenda]. 

Plagiochila telekii Steph. 1891, probably = P. squamulosa var. crispulo-caudata (Gottsche) Vanden Berghen 

1981. 

Plagiochila tenax Steph. 1890 = P. pectinata Willd. ex Lindenb. 1839. 

Plagiochila terebrans Nees et Mont. ex Lindenb. ANG(2), ANN(2,33), BIO(36), CMN(2), COM(1,4), 

IVO(53), KEN(2), MAS(1), MDG(1), MLW(2,114), REU(1), RIM(10), RWA(2), SAF(2), SAO(2), 

TAN(2), UGA(2,96), ZAI(2). 

Plagiochila thomeensis Steph. 1886 = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila togoensis Steph. 1895 = P. strictifolia Steph. 1891. 

Plagiochila triangularis Steph. 1886 = P. barteri var. barteri Mitt. 1886. 

Plagiochila tricuspis Steph. ANN(2), BIO(33), SAO(2). 

Plagiochila tridenticulata auct. [non Dumort. 1835] = P. exigua (Taylor) Taylor 1846. 

Plagiochila ubangiensis Steph. 1921, perhaps = P. terebrans Nees et Mont. ex Lindenb. 1840. 

Plagiochila uhligii Steph. 1921 = P. barteri ssp. colorans (Steph.) Pócs 1993. 

Plagiochila undulata Dugas 1928, hom. illeg. [non (L.) Mont. et Nees 1838] = P. cambuena Steph. 1890. 

Plagiochila uondana Gola 1914, probably = P. squamulosa Mitt. 1863. 

Plagiochila valida Steph. 1904 = P. barteri var. valida (Steph.) Vanden Berghen 1981. 

Plagiochila vignolii Gola 1914 = P. squamulosa var. sinuosa (Mitt.) Vanden Berghen 1981. 

Plagiochila volkensii Steph. 1895 = P. lastii Mitt. 1886.


Plagiochila wilmsiana Steph. CMN(2), SAF(2). 

Plagiochila winteri Steph. 1902, perhaps = P. strictifolia Steph. 1891 or P. moenkemeyeri Steph. 1904. 

Pleuroschisma convexa Steph. 1895 = Bazzania nitida (F.Weber) Grolle 1960. 

Pleurozia gigantea (F.Weber) Lindb. MAU(1), REU(1,4), SAO(2), SEY(4), STH(2,49), TAN(2). 

Pleurozia gigantea var. major (J.B.Jack) A.Evans 1892 = P. gigantea (F.Weber) Lindb. 1874. 

Pleurozia sphagnoides Trevis. 1877 = P. gigantea (F.Weber) Lindb. 1874. 

Porella abyssinica (Nees) Trevis. ETH(2), KEN(2,71), RWA(2), TAN(2), ZAI(2). 

Porella abyssinica var. hoehnelii (Steph.) Pócs BUR(2), ETH(2), KEN(2,21,665), MDG(1), MLW(109), 

MOZ(2), SAF(2), TAN(2), UGA(2,96). 

Porella arboris-vitae (With.) Grolle CVI(2,37). 

Porella capensis (Gottsche) Mitt. ETH(2), KEN(2), MLW(109), SAF(2,75), TAN(2), UGA(96), ZIM(2). 

Porella cucullistipula Steph. COM(1), MAU(1,43), MDG(1). 

Porella hoehneliana Steph. ex S.W.Arnell 1957, nom. inval. = P. abyssinica var. hoehnelii (Steph.) Pócs 1994. 

Porella hoehnelii Steph. 1891 = P. abyssinica var. hoehnelii (Steph.) Pócs 1994. 

Porella keniana (Steph.) S.W.Arnell 1956, nom. inval. = P. abyssinica var. hoehnelii (Steph.) Pócs 1994. 

Porella laevigata (Schrad.) Pfeiffer 1855 = P. arboris-vitae (With.) Grolle 1969. 

Porella madagascariensis (Nees et Mont.) Trevis. MDG(1). 

Porella matongae (Herzog) S.W.Arnell 1963 = P. vallis-gratiae (Gottsche ex Steph.) S.W.Arnell 1955. 

Porella mattirolii (Gola) S.W.Arnell ETH(2). 

Porella montantii (Steph.) E.W.Jones MDG(1). 

Porella nilgheriensis (Mont.) Trevis. 1877 [Dubia et Excludenda]. 

Porella prolixa (Gottsche ex Steph.) E.W.Jones REU(1). 

Porella subdentata var. camerunensis E.W.Jones BIO(2,33), CMN(2), GHA(2). 

Porella subdentata var. subdentata (Mitt.) E.W.Jones ANG(2), CAF(2), CMN(2), ETH(2), GAB(2), 

GHA(2), GUI(2), IVO(51), KEN(2,665), MLW(109), NGA(2), PRI(2), RWA(2), SAO(2), SIE(2), 

TAN(2), TOG(2), UGA(2,96), ZAI(2), ZAM(2). 

Porella triquetra (Steph.) E.W.Jones ETH(2), MDG(1), MLW(2), REU(1), TAN(2), ZIM(2). 

Porella vallis-gratiae (Gottsche ex Steph.) S.W.Arnell MOZ(2), SAF(2,75). 

Potamolejeunea holtii (Spruce) Greig-Smith 1948 = Lejeunea holtii Spruce 1887. 

Prionolejeunea aberrans Steph. 1923 = Lejeunea papilionacea Steph. 1892. 

Prionolejeunea corbisieri Pearson 1922 = Lejeunea sp. 

Prionolejeunea deplanata Schiffn. 1893 = P. grata (Gottsche) Schiffn. 1893. 

Prionolejeunea exarmata Steph. 1913 = Lejeunea papilionacea Steph. 1892. 

Prionolejeunea fissistipula Steph. 1913 = Drepanolejeunea physaefolia (Gottsche) Steph. 1913. 

Prionolejeunea grata (Gottsche) Schiffn. ANN(2,33), BIO(72), CMN(2), COM(1,4), GAB(2), GHA(2), 

IVO(2), MAU(1), MDG(1), MLW(563), NGA(2), PRI(2), REU(1), RIM(10), RWA(2), SAO(2), 

SEY(1), TAN(2), UGA(50), ZAI(2), ZIM(2). 

Prionolejeunea hamidens Herzog 1947 = P. grata (Gottsche) Schiffn. 1893. 

Prionolejeunea kindbergii (Steph.) Schiffn. 1893 = P. grata (Gottsche) Schiffn. 1893. 

Prionolejeunea kindbergii Steph. 1892, nom. inval. = P. grata (Gottsche) Schiffn. 1893. 

Prionolejeunea madagascariensis Steph. 1923 = P. grata (Gottsche) Schiffn. 1893. 

Prionolejeunea principensis Vanden Berghen PRI(2). 

Prionolejeunea pyriflora Steph. 1923 = Cheilolejeunea cordigera (Steph.) Grolle 1977. 

Prionolejeunea rutenbergiana (Gottsche) Steph. 1913 = P. grata (Gottsche) Schiffn. 1893. 

Prionolejeunea rutenbergii Gottsche ex Pearson 1892, nom. inval. = P. grata (Gottsche) Schiffn. 1893. 

Prionolejeunea serrula (Mitt.) Steph. 1913 = P. grata (Gottsche) Schiffn. 1893. 

Prionolejeunea uncatifolia Pearson 1922, probably = P. grata (Gottsche) Schiffn. 1893. 

Pseudomarsupidium decipiens (Hook.) Grolle 1963 = Adelanthus decipiens (Hook.) Mitt. 1864. 

Pseudoneura multifida (L.) Gottsche 1863 = Riccardia multifida (L.) Gray 1821. 

Psiloclada clandestina Mitt. SAF(2). 

Ptychanthus africanus Steph. 1910 = P. striatus (Lehm. et Lindenb.) Nees 1838. 

Ptychanthus dioicus (Sim) S.W.Arnell 1953 = P. striatus (Lehm. et Lindenb.) Nees 1838. 

Ptychanthus integrifolius Steph. 1912 = P. striatus (Lehm. et Lindenb.) Nees 1838. 

Ptychanthus pallidus Steph. 1912 = P. striatus (Lehm. et Lindenb.) Nees 1838. 

Ptychanthus pycnocladus Taylor 1846 = Acrolejeunea pycnoclada (Taylor) Schiffn. 1893.


Ptychanthus squarrosus Mont. ex Lehm. MOZ(2). 

Ptychanthus striatus (Lehm. et Lindenb.) Nees BUR(2), CMN(2), ETH(2), GUI(48), IVO(51), KEN(2), 

MDG(1), MLW(2,563), MOZ(2), RWA(2), SAF(2,75), SIE(2), TAN(2), UGA(2,96), ZAI(2), ZAM(2), 

ZIM(2). 

Ptychocoleus abnormis (Gottsche) Steph. 1912 = Phragmicoma abnormis Gottsche 1882 [Dubia et Excludenda]. 

Ptychocoleus africanus Steph. 1912 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Ptychocoleus amplectens Steph. 1912 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Ptychocoleus borgenii Steph. 1912 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Ptychocoleus confertissimus (Steph.) Steph. 1912 = Acrolejeunea emergens var. confertissima (Steph.) Gradst. 

1975. 

Ptychocoleus convexus (Steph.) Steph. 1912 = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960. 

Ptychocoleus emergens (Mitt.) Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Ptychocoleus ferrugineus (Steph.) Steph. 1912 = Schiffneriolejeunea ferruginea (Steph.) Gradst. 1974. 

Ptychocoleus flagelliferus Steph. 1912 = Acrolejeunea emergens var. confertissima (Steph.) Gradst. 1975. 

Ptychocoleus floribundus Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Ptychocoleus fulvus (Gottsche) Steph. 1912 = Phragmicoma fulvus Gottsche 1882 [Dubia et Excludenda]. 

Ptychocoleus fuscescens (Gola) Steph. 1912, hom. illeg. [non (Hampe ex Lehm.) Trevis. 1877] = Leucolejeunea 

xanthocarpa (Lehm. et Lindenb.) A.Evans 1907. 

Ptychocoleus grossispicus Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Ptychocoleus henriquesii Steph. 1912 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Ptychocoleus inermis Steph. 1912 = Lopholejeunea erugata B.Thiers 1984. 

Ptychocoleus inflexus (Gottsche) Steph. 1912 = Phragmicoma inflexus Gottsche 1882 [Dubia et Excludenda]. 

Ptychocoleus madagascariensis Steph. 1912 = Schiffneriolejeunea madagascariensis (Steph.) Gradst. 1974. 

Ptychocoleus molleri (Steph.) Steph. 1912 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Ptychocoleus occultus (Steph.) Steph. 1912 = Schiffneriolejeunea occulta (Steph.) Gradst. 1974. 

Ptychocoleus orchidae (Pearson) Demaret 1942 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Ptychocoleus pappeanus (Nees) Steph. 1912 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Ptychocoleus parvilobus (Steph.) Steph. 1912 = Schiffneriolejeunea parviloba (Steph.) Gradst. 1974. 

Ptychocoleus polycarpus (Nees) Trevis. 1877 = Schiffneriolejeunea polycarpa (Nees) Gradst. 1974. 

Ptychocoleus pusillus Steph. 1912 = Acrolejeunea emergens var. confertissima (Steph.) Gradst. 1975. 

Ptychocoleus quintasii Steph. 1912 = Schiffneriolejeunea pappeana (Nees) Gradst. 1974. 

Ptychocoleus renauldii Steph. 1912 = Acrolejeunea aulacophora (Mont.) Steph. 1895. 

Ptychocoleus renauldii var. victoriae E.W.Jones 1954 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Ptychocoleus roccatii (Gola) Steph. 1912 = Evansiolejeunea roccatii (Gola) Vanden Berghen 1960. 

Ptychocoleus rupestris Herzog 1948 = Acrolejeunea emergens (Mitt.) Steph. 1895. 

Ptychocoleus vanderystii Steph. 1912 = Acrolejeunea emergens var. confertissima (Steph.) Gradst. 1975. 

Ptychocoleus yangambiensis Vanden Berghen 1951 = Caudalejeunea yangambiensis (Vanden Berghen) 

E.W.Jones 1957. 

Ptycholejeunea dioica Sim 1926 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838. 

Ptycholejeunea striata (Lehm. et Lindenb.) Steph. 1895 = Ptychanthus striatus (Lehm. et Lindenb.) Nees 1838. 

Pycnolejeunea africana Steph. 1923 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976. 

Pycnolejeunea angustiflora Steph. 1914 = Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones 1985. 

Pycnolejeunea bidentula Steph. in Jack et Steph. 1894 = Lepidolejeunea bidentula (Steph.) R.M.Schust. 1980. 

Pycnolejeunea contigua (Nees) Grolle COM(4), GHA(2,14), MDG(1,14), NGA(2,14), RIM(10), 

SIE(2,14), TAN(2,14). 

Pycnolejeunea decurrens Steph. 1914 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976. 

Pycnolejeunea delessertii (Nees et Mont.) Grolle 1974 = Lepidolejeunea delessertii (Nees et Mont.) Grolle 1984. 

Pycnolejeunea involuta Steph. 1914 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976. 

Pycnolejeunea multiflora (Steph.) Steph. 1914 = Lepidolejeunea bidentula (Steph.) R.M.Schust. 1980. 

Pycnolejeunea nicobarica Steph. 1896 = Lepidolejeunea bidentula (Steph.) R.M.Schust. 1980. 

Pycnolejeunea silvestris (Gottsche) Steph. 1914 = Cheilolejeunea surrepens (Mitt.) E.W.Jones 1976. 

Pycnolejeunea ventricosa Schiffn. 1898 = Cheilolejeunea ventricosa (Schiffn.) X.-L.He 1999 (14). 

Radula allamanoi Gola 1914 = R. voluta Taylor ex Gottsche et al. 1845. 

Radula angustata Steph. 1886 = R. appressa Mitt. 1879. 

Radula ankefinensis Gottsche ex Steph. BIO(36), BUR(2), CMN(2), ETH(2), KEN(661,665), MAU(1), 

MDG(1,661), MLW(2,114), REU(1,661), RWA(2), SAF(2,661), SWZ(2), TAN(2,661), UGA(2,96),


ZAI(2,661), ZIM(2). The collections from Ghana and Sierra Leone, reported as Radula ?holstiana, could not be 

named with certainty (25). Grolle (661) stated that past reports from Ethiopia require verification. 

Radula aphylla Mitt. ex Steph., nom. inval. TAN(2). 

Radula appressa Mitt. ANN(2,33), BIO(2,33,72), CAF(2), CMN(2), COM(1), GHA(2), GUI(2), 

IVO(53), KEN(71), MAU(1), MDG(1,4), MLW(114), MOZ(2), NGA(2), PRI(2), REU(1,112), 

RIM(10), ROD(1), SAO(2), SEY(1), SIE(2), SOC(94), TAN(2), UGA(96), ZAI(2). 

Radula autoica Steph. 1913 = R. ankefinensis Gottsche ex Steph. 1884. 

Radula bipinnata Mitt. 1863 = R. boryana (F.Weber) Mont. 1842. 

Radula boryana (F.Weber) Mont. ANN(2,33), BIO(2,33,36), CMN(2), COM(1,661), GHA(2), 

KEN(2,661,665), MAU(1), MDG(1,661), MLW(2,114), PRI(2), REU(1,112,661), RIM(10), 

RWA(2,661), SAF(2,661), SAO(2,661), TAN(2,661), TOG(2), UGA(2), ZAI(2,661), ZIM(2,661). The 

collection from Ethiopia (669) has been redetermined as R. fulvifolia (661). 

Radula caespitosa Steph. 1888 = R. madagascariensis Gottsche 1882. 

Radula capensis Steph. 1884 = R. lindenbergiana Gottsche ex C.Hartm. 1864. 

Radula carringtonii J.B.Jack REU(112). 

Radula commutata Gottsche ex J.B.Jack 1881 = R. lindenbergiana Gottsche ex C.Hartm. 1864. 

Radula comorensis Steph. COM(1), LES(75), MAU(1), MLW(114), REU(1,112), RWA(2), UGA(50), 

ZAI(2). 

Radula complanata (L.) Dumort. SAF(2). 

Radula corniculata auct. [non Dumort. 1831] = Plagiochila exigua (Taylor) Taylor 1846. 

Radula delessertii Steph. 1893 = R. voluta Taylor ex Gottsche et al. 1845 (661). 

Radula epiphylla Mitt. ex Steph. 1884 = R. flaccida Lindenb. et Gottsche 1847. 

Radula evelynae Yamada KEN(2,665), REU(1), TAN(2), ZAI(2). 

Radula exigua Steph. 1910 = R. madagascariensis Gottsche 1882 (661). 

Radula flaccida Lindenb. et Gottsche ANG(2), CMN(2), COM(1), GHA(2), GUI(2), IVO(2), NGA(2), 

PRI(145), RIM(10), RWA(2), SAO(2), SIE(2), TAN(2), UGA(2,42), ZAI(2). 

Radula foliicola Steph. 1910 = R. flaccida Lindenb. et Gottsche 1847. 

Radula fulvifolia (Hook.f. et Taylor) Gottsche et al. CAF(661), CMN(661), COM(661), ETH(2,661), 

GHA(661), KEN(2,661,665), MAU(661), MDG(661), PRI(661), REU(661,662), RIM(661), 

SAF(2,661,662), STH(2,49,661), TAN(2,661), UGA(2,96,661), ZAI(2,661), ZIM(661). 

Radula guineensis Steph. 1884 = R. appressa Mitt. 1879. 

Radula hallapensis Mont. ex Castle SAF(2). 

Radula hans-meyeri Steph. ex S.W.Arnell 1960, nom. inval. - an orthographic error for R. meyeri Steph. 

Radula heteroica Steph. 1910 = R. ankefinensis Gottsche ex Steph. 1884 (661). 

Radula holstiana Steph. 1895 = R. ankefinensis Gottsche ex Steph. 1884 (661). 

Radula javanica Gottsche 1845 [Dubia et Excludenda]. 

Radula lespagnolii Steph. 1910 = R. voluta Taylor ex Gottsche et al. 1845 (661). 

Radula lindbergiana Gottsche ex Lindb. 1881, nom. inval. - an orthographic variant of R. lindenbergiana 

Gottsche ex C.Hartm. 

Radula lindenbergiana Gottsche ex C.Hartm. CVI(2,37), LES(75), MLW(114), SAF(2,75), TAN(2), 

ZIM(2). 

Radula macroloba Steph. 1892 = R. voluta Taylor ex Gottsche et al. 1845 (661). 

Radula macroloba fo. gigantea S.W.Arnell 1957, nom. inval. = R. voluta Taylor ex Gottsche et al. 1845. 

Radula madagascariensis Gottsche COM(661), KEN(661,665), MAU(1,108,661), MDG(1,4,661), 

MLW(114), REU(1,661), TAN(2,661). 

Radula marojezica E.W.Jones MDG(1). 

Radula mascarena Steph. 1884 = R. appressa Mitt. 1878. 

Radula mauritiana Mitt. ex Steph. 1884 = R. complanata (L.) Dumort. 1831. 

Radula molleri Steph. 1910 = R. appressa Mitt. 1878. 

Radula meyeri Steph. 1888 = R. fulvifolia (Hook.f. et Taylor) Gottsche et al. 1845 (661). 

Radula multiramea Gola 1920 = R. voluta Taylor ex Gottsche et al. 1845. 

Radula multiramosa Steph. 1924 = R. boryana (F.Weber) Mont. 1842. 

Radula newtonii Steph. 1910 = R. stipatiflora Steph. 1910. 

Radula oblongiloba Steph. 1884 = R. appressa Mitt. 1878. 

Radula pallens (Sw.) Nees et Mont. 1839 [Dubia et Excludenda].


Radula perrottetii Gottsche ex Steph. 1884 [Dubia et Excludenda]. 

Radula physoloba Mont. 1843 - a record of this taxon from South Africa seems likely to represent R. 

lindenbergiana. 

Radula pirottae Gola 1914 = R. quadrata Gottsche 1845. 

Radula pseudoflaccida E.W.Jones TAN(2). 

Radula quadrata Gottsche CMN(2), COM(1), ERI(121), ETH(2), KEN(2,71,665), MAU(1), MDG(1), 

MLW(114), REU(1), RWA(2), SAF(2), TAN(2), UGA(96), ZAI(2). 

Radula recurvifolia Steph. 1888 = R. quadrata Gottsche 1845. 

Radula retroflexa Taylor REU(112). 

Radula sarogliae Gola 1914 = R. boryana (F.Weber) Mont. 1842. 

Radula silvestris Gottsche MDG(1). 

Radula socorana Gerola ETH(503). 

Radula spongiosa Steph. 1910 = R. quadrata Gottsche et al. 1845. 

Radula stenocalyx Mont. BUR(2), CMN(2), COM(1), GAB(2), GHA(2), GUI(2), KEN(2), MAU(1), 

MDG(1,4), MLW(114), REU(1), RIM(10), RWA(2), SAF(2), SAO(2), SEY(73), SIE(2), TAN(2), 

UGA(50), ZAI(2). 

Radula stipatiflora Steph. 1910 = R. voluta Taylor ex Gottsche et al. 1845 (661). 

Radula tabularis Steph. SAF(2,661). 

Radula tamariscina Mitt. 1863 = R. boryana (F.Weber) Mont. 1842 (661). 

Radula thomeensis Steph. 1910 = R. boryana (F.Weber) Mont. 1842. 

Radula tubiflora Steph. 1892 ("tubaeflora") = R. stenocalyx Mont. 1855. 

Radula vaginata Steph. 1910 = R. stenocalyx Mont. 1855. 

Radula voluta Taylor ex Gottsche et al. BIO(2,33,36), ETH(2,661), KEN(2,71,661,665), MAU(1), 

MDG(1,4,661), MLW(2,114), REU(1,112), RWA(2), SAF(2,75,661), TAN(2,661), UGA(2,96,661), 

ZAI(2,661). 

Radula wilmsiana Steph. ex Castle, nom. inval. SAF(2). 

Radula woodiana Steph. 1910 = R. boryana (F.Weber) Mont. 1842 (660) 

Radula xalapensis Nees et Mont. 1836 [Dubia et Excludenda]. 

Reboulia hemisphaerica (L.) Raddi CHA(2,3), CVI(2,37), ERI(2), ETH(2), MLW(2), SOC(2), TAN(2). 

Rectolejeunea aloba (Sande Lac.) Steph. 1914 = Lejeunea aloba Sande Lac. 1856. 

Rectolejeunea arnellii E.W.Jones 1974 = Lejeunea phyllobola Nees et Mont. 1842 (168). 

Rectolejeunea brittoniae A.Evans 1911 = Lejeunea phyllobola Nees et Mont. 1842 (168). 

Rectolejeunea pililoba (Spruce) R.M.Schust. 1980 = Lejeunea pililoba Spruce 1895. 

Rectolejeunea rhodesiae (Sim) S.W.Arnell 1953 = Lejeunea rhodesiae (Sim) R.M.Schust. 1962. 

Rectolejeunea santae-mariae Steph. 1914 = Cheilolejeunea intertexta (Lindenb.) Steph. 1897. 

Rectolejeunea setacea (Steph.) R.M.Schust. 1980 = Lejeunea setacea (Steph.) Steph. 1914. 

Riccardia amazonica (Spruce) Schiffn. ex Gradst. BIO(36), CMN(2), CON(2), GAB(2), GHA(2), 

IVO(51), KEN(2), LES(75), NGA(2), RIM(10), SAF(2), SIE(2), TAN(2), UGA(2,96), ZAI(2), ZAM(2), 

ZIM(2). 

Riccardia angusticosta (Steph.) Grolle 1973 = R. amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Riccardia caespitans (Steph.) E.W.Jones 1956, nom. inval. = Aneura caespitans Steph. 1891. 

Riccardia campanuliflora S.W.Arnell 1952 = R. amazonica (Spruce) Schiffn. ex Gradst. 1979. 

Riccardia capensis S.W.Arnell SAF(2). 

Riccardia comosa (Steph.) E.W.Jones 1956, nom. inval. = Aneura comosa Steph. 1890. 

Riccardia compacta (Steph.) S.W.Arnell SAF(2), TAN(2), UGA(2). 

Riccardia congoana (Steph.) Demaret 1942 = R. amazonica (Steph.) Schiffn. ex Gradst. 1979. 

Riccardia erosa (Steph.) E.W.Jones CMN(2), CON(2), ETH(2), SAO(2), TAN(2). 

Riccardia exigua (Steph.) S.W.Arnell 1957, nom. inval. = R. longispica (Steph.) Pearson 1893. 

Riccardia fastigiata (Lehm.) Trevis. CAF(2), CON(2), ETH(2), KEN(2,665), LES(75), SAF(2,75), 

SEY(4), TAN(2), UGA(96), ZAI(2), ZAM(2), ZIM(2). 

Riccardia holstii (Steph.) E.W.Jones CMN(2), TAN(2), UGA(115), ZAI(2), ZIM(2). 

Riccardia kilimandjarica S.W.Arnell 1959 = R. compacta (Steph.) S.W.Arnell 1952. 

Riccardia laurentiana (Steph.) Demaret 1942 = Aneura pinguis (L.) Dumort. 1831 sens. lat. 

Riccardia lepervanchei (Steph.) S.W.Arnell 1965, nom. inval. = Aneura lepervanchei Steph. 1899. 

Riccardia limbata (Steph.) E.W.Jones ANN(2,33), BIO(36), CMN(2), CON(2), GHA(2), KEN(2,665),


MAU(1), MLW(2), NGA(2), REU(1), RIM(10), RWA(48), SAF(2), SAO(2), SIE(2), SWZ(2), TAN(2), 

TOG(2), ZAI(2), ZAM(2). 

Riccardia longispica (Steph.) Pearson BIO(36), CMN(2), COM(1), MAU(1), MDG(1), REU(1), 

RIM(10), SEY(1), TAN(2), UGA(115), ZAI(2). 

Riccardia multifida (L.) Gray MDG(1), SAF(653). 

Riccardia nudiflora (Steph.) Grolle MAU(1), MDG(1), REU(1). 

Riccardia obtusa S.W.Arnell SAF(2). 

Riccardia palmata (Hedw.) Carruth. 1865 [Dubia et Excludenda]. 

Riccardia pinguis (L.) Gray 1821 = Aneura pinguis (L.) Dumort. 1831. 

Riccardia pinnatifida (F.Weber) Carruth. 1865 [Dubia et Excludenda]. 

Riccardia pseudopinguis Herzog 1942 = Aneura pseudopinguis (Herzog) Pócs 1983. 

Riccardia ramosissima (Steph.) E.W.Jones 1956, nom. inval. = R. ramosissima (Steph.) Grolle 1995. 

Riccardia ramosissima (Steph.) Grolle REU(1). 

Riccardia rhodesiae S.W.Arnell ZIM(2). 

Riccardia saccatiflora (Steph.) S.W.Arnell KEN(2), MAU(1), MDG(1), REU(1,653), RIM(10), TAN(2), 

UGA(96). Perold 2001c (653) showed that Arnell's records of R. saccatiflora from South Africa should be 

referred to R. multifida. 

Riccardia stephanii (Besch. ex Steph.) E.W.Jones 1956, hom. illeg. [non S.Hatt. 1944] = R. amazonica (Spruce) 

Schiffn. ex Gradst. 1979. 

Riccardia submarginata S.W.Arnell 1953 = Aneura pinguis (L.) Dumort. (652). 

Riccardia sp. (Aneura caespitans Steph.) CMN(2), KEN(2), MDG(1), MAU(1), REU(1), UGA(2). 

Riccardia sp. (Aneura comosa Steph.) MAU(1), REU(1). 

Riccardia sp. (Aneura corbieri Steph.) MDG(1). 

Riccardia sp. (Aneura dismieri Steph.) MDG(1). 

Riccardia sp. (Aneura keniae Gola) KEN(2). 

Riccardia sp. (Aneura lepervanchei Steph.) REU(1). 

Riccardia sp. GHA(25), ?MAU(25), ?RWA(25), ?ZAI(25). 

Riccia abnormis (Steph.) Steph. 1898 = R. moenkemeyeri Steph. 1887. 

Riccia aegyptiaca S.W.Arnell 1963 = R. congoana Steph. 1898. 

Riccia africana Sim 1926 = R. crozalsii Levier 1902. 

Riccia alatospora O.H.Volk et Perold SAF(2). 

Riccia albolimbata S.W.Arnell KEN(2), NAM(2), SAF(2), TAN(2). 

Riccia albomarginata Bisch. ex C.Krauss SAF(2). 

Riccia alboporosa Perold SAF(2). 

Riccia albornata O.H.Volk et Perold SAF(2). 

Riccia albosquamata S.W.Arnell 1957 = R. albolimbata S.W.Arnell 1957. 

Riccia albovestita O.H.Volk NAM(2), SAF(2). 

Riccia ampullacea Perold LES(2,75), SAF(2). 

Riccia angolensis Steph. ANG(2), BOT(2), KEN(2), NAM(2), SAF(2). 

Riccia argenteolimbata O.H.Volk et Perold BOT(2), KEN(2), NAM(2), SAF(2), TAN(2). 

Riccia atromarginata var. jovet-astiae Rauh et Buchloh MDG(1). 

Riccia atropurpurea Sim ANG(2), BEN(11), BOT(2), GHA(2), MLW(2), NAM(2), NGA(2), RWA(2), 

SAF(2), SWZ(2), TAN(2), UGA(2), ZAM(2), ZIM(2,9). 

Riccia beccarii Steph. ex Chiov., nom. inval. ERI(2). 

Riccia berriei E.W.Jones 1957 = R. congoana Steph. 1898. 

Riccia bicolorata Perold SAF(2). 

Riccia biokoensis Perold BIO(98). 

Riccia bullosa Link ex Lindenb. LES(2,75), SAF(2). 

Riccia campbelliana M.Howe 1899 = R. macrocarpa Levier 1894. 

Riccia canescens Steph. 1898 = R. crinita Taylor 1846 (555). 

Riccia capensis auct. [non Steph. 1913] = R. nigrella DC. 1815. 

Riccia capensis Steph. 1913 = R. limbata Bisch. ex C.Krauss 1846. 

Riccia cavernosa Hoffm. ANG(2), BOT(2), CHA(2), CVI(37), DJI(2), ERI(2), KEN(2), LES(75), 

MDG(1), MOZ(2), MTN(3), NAM(2), NGR(2,3), SAF(2), SUD(2), TAN(2), ZAI(2), ZAM(2), ZIM(2).


Riccia chevalieri Steph. 1917, probably = R. moenkemeyeri Steph. 1887. 

Riccia ciliata Hoffm. CVI(2,37). 

Riccia ciliata ssp. canescens (Steph.) R.M.Schust. 1992 = R. crinita Taylor 1846 (555). 

Riccia ciliata ssp. trichocarpa (M.Howe) R.M.Schust. 1992 = R. crinita Taylor 1846 (555). 

Riccia commutata J.B.Jack ex Levier 1894 = R. warnstorfii Limpr. ex Warnst. 1885. 

Riccia compacta Garside SAF(2). 

Riccia concava Bisch. ex C.Krauss SAF(2). 

Riccia congoana Steph. ANG(2), BEN(11), BOT(2), CHA(2), CON(2), GHA(2), KEN(2,665), MLW(2), 

NAM(2), NGA(2), RWA(2), SAF(547), SIE(2), SUD(2), SWZ(2), TAN(2), UGA(2,96), ZAI(2), 

ZAM(2), ZIM(2). 

Riccia convexa Steph. ex Har. 1913, nom. inval. [? non Steph. 1917] = R. frostii Austin 1875. 

Riccia coronata Sim This species is insufficiently known. The specimen cited by Sim cannot be traced, and 

another specimen, which had been identified as 'near R. coronata' is placed under R. alatospora (fide Perold 

1999g (8)). 

Riccia crinita Taylor BOT(2), KEN(2), LES(2), MDG(1), MLW(2), NAM(2), NGR(2), RWA(2), 

SAF(2), SOC(2), TAN(2), UGA(2), ZIM(2). 

Riccia crozalsii Levier SAF(2), TAN(2). 

Riccia crystallina L. emend. Raddi CHA(3), CVI(2,37), ETH(2), SAF(2), TAN(2), ZIM(2). It is uncertain 

whether some early records represent this species or R. cavernosa. 

Riccia crystallina var. angustior sensu Gillet et Ast 1957 = R. cavernosa Hoffm. 1796. 

Riccia cupulifera A.V.Duthie SAF(2). 

Riccia curtisii (James ex Austin) Austin CHA(2), SAF(2). 

Riccia dinteri Steph. ex S.W.Arnell 1956, nom. inval. = R. stricta (Lindenb.) Perold 1990. 

Riccia discolor Lehm. et Lindenb. BEN(11), IVO(2,60), KEN(2), MLW(2), NGA(2), NGR(2), TAN(2), 

UGA(2), ZAI(2), ZAM(2). 

Riccia duthieae O.H.Volk et Perold 1985 = R. albovestita O.H.Volk 1981. 

Riccia elongata Perold SAF(2). 

Riccia erubescens Perold ANG(2), MLW(2), MOZ(2), ZAM(2). 

Riccia erythraea (Gola) Cufod. 1952 = R. lanceolata Steph. 1888. 

Riccia erythraeae Steph. ex Chiov. 1919, nom. inval. = R. lanceolata Steph. 1888. 

Riccia esulcata Steph. ex Har. 1913, nom. inval. [non Steph. 1917] = R. frostii Austin 1875. 

Riccia fluitans sens. lat. IVO(2), SAO(2), TAN(2), TOG(2), ZAM(2). 

Riccia fluitans var. δ stricta Lindenb. 1837 = R. stricta (Lindenb.) Perold 1990. 

Riccia frostii Austin CHA(2,3), CVI(2), MTN(2,3), SUD(2). 

Riccia furfuracea Perold SAF(2). 

Riccia garsidei Sim SAF(2). 

Riccia glauca L. ETH(2). 

Riccia gougetiana Durieu et Mont. CVI(2,37). 

Riccia hantamensis Perold SAF(2). 

Riccia hariotii Steph. ex Har., nom. inval. SUD(2). 

Riccia helenae Ast REU(1). 

Riccia hirsuta O.H.Volk et Perold SAF(2). 

Riccia hortorum Bory ex Lindenb. MAU(1), REU(1). 

Riccia huebeneriana Lindenb. ETH(2), REU(45). 

Riccia intermedia E.W.Jones 1957 = R. discolor Lehm. et Lindenb. 1832. 

Riccia lamellosa Raddi CHA(2,3), ERI(2), TAN(2). 

Riccia lanceolata Steph. BEN(2,11), ERI(2), ETH(2), GHA(2), IVO(2,60), MLW(2,114), MOZ(2), 

NGA(2), NGR(2), RWA(2), TAN(2), UGA(2,96), ZAM(2). 

Riccia lanceolata var erythraea Gola 1914 = R. lanceolata var. lanceolata Steph. 1888. 

Riccia laxisquamata (Steph.) Steph. ETH(2). 

Riccia limbata Bisch. ex C.Krauss ETH(2), SAF(2). 

Riccia limbatoides O.H.Volk 1984, nom. prov. = R. congoana Steph. 1898. 

Riccia macrocarpa Levier SAF(2), SOC(2). 

Riccia mammifera O.H.Volk et Perold SAF(2).


Riccia membranacea Gottsche et Lindenb. ANG(2), CMN(2), GHA(2), KEN(2), NGA(2), SIE(2), 

SUD(2), TAN(2), ZAI(2). 

Riccia microciliata O.H.Volk et Perold ANG(2), BOT(2), MLW(2), MOZ(2), NAM(2), SAF(2), 

SWZ(2), TAN(2), ZIM(9). 

Riccia moenkemeyeri Steph. ANG(2), BEN(11), BIO(33), CAF(2), CMN(2), GAM(2), GHA(2), 

IVO(2,60), MLW(2), NGA(2), RWA(2), SAF(2), SIE(2), TAN(2), ZAI(2), ZAM(2), ZIM(2). 

Riccia montaguensis S.W.Arnell 1952 = R. bullosa Link ex Lindenb. 1829. 

Riccia montana Perold LES(2,75), SAF(2). 

Riccia namaquensis Perold SAF(2). 

Riccia natalensis Sim SAF(2), ZIM(9). 

Riccia natans L. 1759 = Ricciocarpos natans (L.) Corda 1829. 

Riccia negrii Gola 1914 = R. crystallina L. 1753 emend. Raddi 1818. 

Riccia nigerica E.W.Jones CMN(2), NGA(2). 

Riccia nigrella DC. CVI(37), LES(2,75), NAM(2), SAF(2). 

Riccia nigrosquamata E.W.Jones 1957 = R. congoana Steph. 1898. 

Riccia okahandjana S.W.Arnell ANG(2), BOT(2), LES(75), MLW(2), MOZ(2), NAM(2), RWA(2), 

SAF(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Riccia papillispora Steph. ERI(2). 

Riccia parvo-areolata O.H.Volk et Perold SAF(2). 

Riccia perssonii Sultan Khan BOT(2), CHA(2), MDG(655), MLI(2), NAM(2). 

Riccia plana Taylor 1846 = R. crystallina L. 1753 emend. Raddi 1818. 

Riccia pottsiana Sim SAF(2). 

Riccia pseudolimbata S.W.Arnell 1957 = R. angolensis Steph. 1898. 

Riccia pulveracea Perold SAF(2). 

Riccia purpurascens Lehm. SAF(2). 

Riccia radicosa Pearson ZAI(2). 

Riccia rautanenii (Steph.) Steph. 1898 = R. cavernosa Hoffm. 1796. 

Riccia rhodesiae S.W.Arnell 1952 = R. congoana Steph. 1898. 

Riccia rosea O.H.Volk et Perold ANG(2), BOT(2), KEN(2), NAM(2), SAF(2), TAN(2), UGA(2), 

ZAI(2), ZAM(2), ZIM(11). 

Riccia rubricollis Garside et A.V.Duthie ex Perold SAF(2). 

Riccia runssorensis Steph. ANG(2), BOT(2), NAM(2), SAF(2), TAN(2), UGA(2). 

Riccia saharensis Steph. ex Ast NGR(2,3). Erroneously listed from Mauritania in (2). 

Riccia sarcosa O.H.Volk et Perold 1986 = R. albovestita O.H.Volk 1981. 

Riccia schelpei O.H.Volk et Perold SAF(2). 

Riccia schroederi Steph. TAN(2). 

Riccia schweinfurthii Steph. SUD(2). 

Riccia sibayenii Perold SAF(651). 

Riccia simii Perold LES(75), SAF(2). 

Riccia somaliensis Perold SOM(2). 

Riccia sorocarpa Bisch. CHA(2,3), CVI(37), KEN(2), LES(2), SAF(2), TAN(2). 

Riccia sorocarpa var. heegii Schiffn. CVI(2). 

Riccia spongosa S.W.Arnell 1952 = R. curtisii (James ex Austin) Austin 1879. 

Riccia stricta (Lindenb.) Perold ANG(2), BIO(36), BOT(2), BUR(2), CMN(2), ETH(2), GHA(2), 

IVO(2), KEN(2,21,665), LES(75), MLW(2), MOZ(2), NAM(2), NGA(2), NGR(2), RWA(2), SAF(2), 

SUD(2), TAN(2), TOG(2), UGA(2,96), ZAI(2), ZAM(2), ZIM(2,9). 

Riccia stricta A.V.Duthie ex S.W.Arnell 1963, nom. inval. = R. stricta (Lindenb.) Perold 1990. 

Riccia sudanensis Steph. ex Har., nom. inval. SUD(2). 

Riccia suntica Pettet 1967 = R. cavernosa Hoffm. 1796. 

Riccia symoensii Vanden Berghen ZAI(2). 

Riccia tellinii Gola 1914 = R. cavernosa Hoffm. 1796. 

Riccia terracianoi Gola 1914 = R. cavernosa Hoffm. 1796. 

Riccia tomentosa O.H.Volk et Perold SAF(2).


Riccia trabutiana Steph. SOC(2). 

Riccia trachyglossum Perold LES(2). 

Riccia triangularis Steph. CAF(2). 

Riccia trichocarpa M.Howe 1898 = R. crinita Taylor 1846 (555). 

Riccia undulata S.W.Arnell 1952 = R. moenkemeyeri Steph. 1886. 

Riccia villosa Steph. SAF(2). 

Riccia vitrea Perold SAF(2). 

Riccia volkii S.W.Arnell NAM(2), SAF(2), ZIM(2,9). 

Riccia vulcanicola E.Fischer RWA(2). 

Riccia warnstorfii Limpr. SAF(2). Twice collected from a garden in 1954, but not subsequently recorded from 

South Africa, and considered to have been a short-lived introduction (Perold 1999g (8)). 

Riccia welwitschii Steph. 1898 = R. membranacea Gottsche et Lindenb. 1846. 

Ricciella abnormis Steph. 1891 = Riccia moenkemeyeri Steph. 1887. 

Ricciella laxisquamata Steph. 1895 = Riccia laxisquamata (Steph.) Steph. 1898. 

Ricciella purpurascens (Lehm.) Lehm. et Lindenb. 1832 = Riccia purpurascens Lehm. 1829. 

Ricciella rautanenii Steph. 1895 = Riccia cavernosa Hoffm. 1796. 

Ricciella stricta (Lindenb.) Trevis. 1877 = Riccia stricta (Lindenb.) Perold 1990. 

Ricciella tellinii (Gola) Cufod. 1952 = Riccia cavernosa Hoffm. 1796. 

Ricciella terracianoi (Gola) Cufod. 1952 = Riccia cavernosa Hoffm. 1796. 

Ricciella welwitschii Steph. 1901 = Riccia membranacea Gottsche et Lindenb. 1846. 

Ricciocarpos natans (L.) Corda BOT(2), GHA(2), KEN(2,665), MOZ(2), NGA(2), RWA(2), SAF(2), 

TAN(2), UGA(96), ZAI(2), ZAM(2). 

Riella affinis M.Howe et Underw. SAF(2,78). 

Riella alatospora Wigglesworth SAF(2,78). 

Riella capensis Cavers SAF(2,78). 

Riella echinospora Wigglesworth NAM(2), SAF(2,78). 

Riella purpureospora Wigglesworth SAF(2,78). 

Sarcocyphos auritus (Lehm.) Nees 1844 = Anastrophyllum auritum (Lehm.) Steph. 1901. 

Sarcocyphos sparsifolius Lindb. 1868 = Marsupella sparsifolia (Lindb.) Dumort. 1874. 

Scapania calcicola (S.W.Arnell et J.Perss.) Ingham LES(75). 

Scapania cuspiduligera (Nees) Müll.Frib. LES(75), SAF(75), ZAI(2). 

Scapania esterhuyseniae S.W.Arnell TAN(2), UGA(2). 

Scapania hedbergii S.W.Arnell KEN(2). 

Schiffneriolejeunea altimontana Vanden Berghen RWA(2), TAN(2), UGA(50), ZAI(2). 

Schiffneriolejeunea ferruginea (Steph.) Gradst. CMN(2), NGA(2). 

Schiffneriolejeunea fragilis Gradst. et E.W.Jones CMN(2), CON(2), GAB(2), GHA(2), GUI(2), IVO(2), 

NGA(2), SIE(2), UGA(42), ZAI(2). 

Schiffneriolejeunea madagascariensis (Steph.) Gradst. MDG(1). 

Schiffneriolejeunea occulta (Steph.) Gradst. CON(2), GHA(2), NGA(2), RIM(10), SIE(2). 

Schiffneriolejeunea pappeana (Nees) Gradst. ANG(2), ANN(2,33), BIO(2,33), CAF(2), CMN(2), 

ETH(2), GUI(2), KEN(2), MDG(1), MLW(2,563), REU(1), RWA(2), SAF(2), SAO(2), TAN(2), 

UGA(2,96), ZAI(2), ZIM(2). 

Schiffneriolejeunea pappeana var. bidentata Gradst. et Vanden Berghen MAU(1), MDG(1), REU(1). 

Schiffneriolejeunea pappeana var. integra Gradst. et Vanden Berghen MDG(1). 

Schiffneriolejeunea parviloba (Steph.) Gradst. MAU(1), REU(1). 

Schiffneriolejeunea polycarpa (Nees) Gradst. ANG(2), BIO(2,33), CAF(2), CON(2), GAB(2), GHA(2), 

GUI(2), MDG(1), MLW(563), MOZ(2), NGA(2), SAF(2), SAO(2), SIE(2), TAN(2), UGA(42,50,96), 

ZAI(2), ZAM(2), ZIM(2). 

Schiffneriolejeunea tumida (Nees) Gradst. SEY(1). 

Schisma balboi Gola 1914 = Herbertus sp. 

Schisma capense Steph. 1909 = Herbertus capensis (Steph.) Sim 1926. 

Schisma capillare (Steph.) Steph. 1909 = Herbertus capillaris Steph. 1892 [Dubia et Excludenda]. 

Schisma dicranum (Taylor ex Gottsche et al.) Steph. 1895 = Herbertus dicranus (Taylor ex Gottsche et al.) 

Trevis. 1877.


Schisma doggeltianum Steph. 1909 = Herbertus doggeltianus (Steph.) Demaret 1942. 

Schisma dusenii Steph. 1909 = Herbertus mollis (Steph.) Dusén 1893. 

Schisma grossevittatum Steph. 1909 = Herbertus grossevittatus (Steph.) S.W.Arnell ex Grolle 1995. 

Schisma hildebrandtii Steph. 1909 = Herbertus capensis (Steph.) Sim 1926. 

Schisma kilimandjarense Steph. 1909 = Herbertus sp. 

Schisma lobatum Steph. 1911 = Herbertus lobatus (Steph.) Demaret 1942. 

Schisma mascarenicum Steph. 1909 = Herbertus capensis (Steph.) Sim 1926. 

Schisma mascarenicum var. kikujense Gola 1914 = Herbertus sp. 

Schisma seriatum Steph. 1909 = Herbertus grossevittatus (Steph.) S.W.Arnell ex Grolle 1995. 

Schisma stuhlmannii Steph. 1909 = Herbertus stuhlmannii (Steph.) Demaret 1942. 

Schisma subdentatum Steph. 1901 = Herbertus subdentatus (Steph.) Fulford 1962. 

Schistochila alata (Lehm.) Schiffn. MDG(1,4), SAF(2). 

Schistochila borbonica Steph. 1890 = Gottschea neesii Mont. 1843. 

Schistochila congoana Steph. CON(2). 

Schistochila engleriana Steph. 1909 = Gottschea engleriana (Steph.) Grolle et Zijlstra 1984. 

Schistochila limbata S.W.Arnell 1957 = Gottschea sphagnoides (Schwägr.) Lindb. 1870. 

Schistochila neesii (Mont.) Lindb. 1872 = Gottschea neesii Mont. 1843. 

Schistochila pauciserrata Kiaer et Pearson MDG(1). 

Schistochila piligera Steph. MDG(1,4). 

Schistochila sphagnoides (Schwägr.) Steph. 1909 = Gottschea sphagnoides (Schwägr.) Lindb. 1870. 

Schistochila thouarsii (Hook.) Dumort. ex Trevis. 1877 = Gottschea sphagnoides (Schwägr.) Lindb. 1870. 

Schistochilaster englerianus (Steph.) H.A.Mill. 1970 = Gottschea engleriana (Steph.) Grolle et Zijlstra 1984. 

Schistochilaster neesii (Mont.) H.A.Mill. 1970 = Gottschea neesii Mont. 1843. 

Schistochilaster piliger (Steph.) H.A.Mill. 1970 = Schistochila piliger Steph. 1890. 

Sendtnera capillaris (Steph.) Renauld 1898 = Herbertus capillaris Steph. 1892 [Dubia et Excludenda]. 

Sendtnera diclados (Brid. ex F.Weber) Endl. ex Gottsche et al. 1845 = Mastigophora diclados (Brid. ex 

F.Weber) Nees 1838. 

Sendtnera dicrana Taylor ex Gottsche et al. 1845 = Herbertus dicranus (Taylor ex Gottsche et al.) Trevis. 1877. 

Sendtnera juniperina Nees 1845 = Herbertus juniperoideus (Sw.) Grolle 1961 [Dubia et Excludenda]. 

Sendtnera mascarenica Mitt. 1860 = Mastigophora diclados (Brid. ex F.Weber) Nees 1838. 

Sendtnera mollis Steph. 1886 = Herbertus mollis (Steph.) Dusén 1893. 

Simia atroviridis (Sim) S.W.Arnell 1962, nom. inval. = Cylindrocolea atroviridis (Sim) Váòa 1979. 

Solenostoma abyssinicum (Nees) Steph. 1901 = Jungermannia abyssinica Nees 1844. 

Solenostoma atrovirens Steph. 1901 = Jungermannia perloi (Gola) Váòa 1973. 

Solenostoma austroafricanum (S.W.Arnell) S.W.Arnell 1963 = Jungermannia mildbraedii Steph. 1914. 

Solenostoma belangerianum Steph. 1917 = Notoscyphus lutescens (Lehm. et Lindenb.) Mitt. 1873. 

Solenostoma borgenii (Gottsche ex Pearson) Steph. 1901 = Jungermannia borgenii Gottsche ex Pearson 1893. 

Solenostoma lacouturei Steph. 1917 = Jungermannia borgenii Gottsche ex Pearson 1893. 

Solenostoma perloi Gola 1914 = Jungermannia perloi (Gola) Váòa 1973. 

Solenostoma sphaerocarpum (Hook.) Steph. 1901 = Jungermannia sphaerocarpa Hook. 1815. 

Solenostoma stoloniferum (Steph.) S.W.Arnell 1963 = Jungermannia borgenii Gottsche ex Pearson 1893. 

Sphaerocarpos stipitatus Bisch. ex Lindenb. SAF(2,77). 

Sphenolobopsis pearsonii (Spruce) R.M.Schust. KEN(102), MLW(102). 

Sphenolobus argentinus Steph. 1902 = Lophozia argentinus (Steph.) R.M.Schust. 1966. 

Sphenolobus crispifolius Steph. 1924 = Plagiochila squamulosa var. crispulo-caudata (Gottsche) Vanden 

Berghen 1981. 

Sphenolobus flagellaris (S.Hatt.) Grolle 1963 = Iwatsukia jishibae (Steph.) N.Kitag. 1965. 

Sphenolobus laceratus Steph. 1902 = Gymnomitrion laceratum (Steph.) Horik. 1943. 

Sphenolobus minutus (Schreb.) Berggr. 1898 = Anastrophyllum minutum (Schreb.) R.M.Schust. 1949. 

Sphenolobus nigrus Rodway 1916 = Allisoniella nigra (Rodway) R.M.Schust. 1972. 

Sphenolobus perigonialis (Hook f. et Taylor) Steph. 1902 = Andrewsianthus perigonialis (Hook.f. et Taylor) 

R.M.Schust. 1966. [Excludenda]. Reports of S. perigonialis in Africa are based on misidentifications. 

Sphenolobus savioi Gola 1914 = Anastrophyllum auritum (Lehm.) Steph. 1901. 

Spruceanthus marianus (Gottsche) Mizut. 1966 = Cheilolejeunea marianus (Gottsche) B.Thiers et Gradst. 1989 


Sprucella moenkemeyeri Steph. 1886 = Lepidozia succida Mitt. 1860. 

Sprucella succida (Mitt.) Steph. 1891 = Lepidozia succida Mitt. 1860.


Sprucella ubangiensis (Steph.) Fulford et J.Taylor 1959 = Lepidozia ubangiensis Steph. 1909. 

Stenorrhipis madagascariensis (Steph.) Grolle 1963 = Kymatocalyx madagascariensis (Steph.) Gradst. et Váòa 

1999 (74). 

Stephaniella paraphyllina J.B.Jack LES(75), SAF(2,75,100). 

Stictolejeunea africana Vanden Berghen 1950 = S. balfourii (Vanden Berghen) E.W.Jones 1976. 

Stictolejeunea balfourii (Mitt.) E.W.Jones CAF(2), CMN(2), GHA(2), IVO(2,53), MAU(1), NGA(2), 

REU(1), RIM(10), ROD(1), SIE(2). 

Stictolejeunea richardsii Herzog 1950 = S. balfourii (Mitt.) E.W.Jones 1976. 

Strepsilejeunea acuta (Mitt.) Steph. 1895 = Lejeunea acuta Mitt. 1863. 

Strepsilejeunea brevifissa (Gottsche) Steph. 1895 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963. 

Strepsilejeunea brevifissa sensu Vanden Berghen 1960, p.p. = Cheilolejeunea usambarana (Steph.) Grolle 1979. 

Strepsilejeunea cardiophylla (Gottsche) Steph. 1913 = Taxilejeunea conformis (Nees et Mont.) Steph. 1914. 

Strepsilejeunea compressa Herzog 1947 = Cheilolejeunea compressa (Herzog) Grolle 1995. 

Strepsilejeunea cordistipula Steph. 1913 = Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones 1985. 

Strepsilejeunea decemplicata Steph. in Levier 1901, nom. inval. = Cheilolejeunea pluriplicata (Pearson) 


Strepsilejeunea decemplicata Steph. in Renauld 1909, nom. inval. = Cheilolejeunea pluriplicata (Pearson) 


Strepsilejeunea georgensis S.W.Arnell 1953 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963. 

Strepsilejeunea inflata Steph. 1913 [non Cheilolejeunea inflata Steph. 1914] = Cheilolejeunea krakakammae 

(Lindenb.) R.M.Schust. 1963. 

Strepsilejeunea knysnana S.W.Arnell 1953 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963. 

Strepsilejeunea krakakammae (Lindenb.) Steph. 1913 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 

1963. 

Strepsilejeunea usambarana Steph. 1913 = Cheilolejeunea usambarana (Steph.) Grolle 1979. 

Strepsilejeunea vatovae Gerola 1947 = Cheilolejeunea sp. 

Stylolejeunea duncaniae Sim 1926 "duncanii" = Lejeunea duncaniae (Sim) S.W.Arnell 1961. 

Stylolejeunea rhodesiae Sim 1926 = Lejeunea rhodesiae (Sim) R.M.Schust. 1962. 

Stylolejeunea setacea (Steph.) S.W.Arnell 1953 = Lejeunea setacea (Steph.) Steph. 1914. 

Symbiezidium balfourii (Mitt.) Steph. 1912 = Stictolejeunea balfourii (Mitt.) E.W.Jones 1976. 

Symbiezidium madagascariense Steph. MDG(1,4), SEY(1). 

Symbiezidium rotale (Hook.f. et Taylor) Trevis. 1877 = Aureolejeunea rotalis (Hook f. et Taylor) Gradst. et 


Symphyogyna aloysii-sabaudiae Gola 1907, probably = S. volkensii Steph. 1900. 

Symphyogyna brasiliensis Nees et Mont. ANG(2), ASC(2), BUR(2), CMN(2), GHA(2), MAU(1), 

MDG(1), MLW(2,29), NGA(2), REU(1), RWA(2), SAF(2), SIE(2), STH(2,49), SWZ(2), TAN(2), 

UGA(2,96), ZAI(2), ZAM(2), ZIM(2). 

Symphyogyna convoluta Gottsche 1846 = S. brasiliensis Nees et Mont. 1836. 

Symphyogyna filicum Nadeaud 1873 = Symphyogynopsis gottscheana (Mont. et Nees) Grolle 1987. 

Symphyogyna gottscheana Mont. et Nees 1846 = Symphyogynopsis gottscheana (Mont. et Nees) Grolle 1987. 

Symphyogyna harveyana Taylor 1846 = S. podophylla (Thunb.) Mont. et Nees 1846. 

Symphyogyna lehmanniana Mont. et Nees 1846 = S. brasiliensis Nees et Mont. 1836. 

Symphyogyna podophylla (Thunb.) Mont. et Nees CMN(2), ETH(2), KEN(2,71), MAU(1), MLW(2,29), 

REU(1), RWA(2), SAF(2), TAN(2), UGA(2,96), ZAI(2), ZIM(2). 

Symphyogyna rhizobola (Schwägr.) Nees 1846 = S. podophylla (Thunb.) Mont. et Nees 1846. 

Symphyogyna rigida Steph. 1911 = S. podophylla (Thunb.) Mont. et Nees 1846. 

Symphyogyna sellae Gola 1907 = S. podophylla (Thunb.) Mont. et Nees 1846. 

Symphyogyna serrata Mitt. 1875 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986. 

Symphyogyna spinosa Lindenb. et Gottsche 1847 = Jensenia spinosa (Lindenb. et Gottsche) Grolle 1986. 

Symphyogyna tenuicosta - an orthographic error for S. tenuicostata Steph. (in Arnell 1956 (7)). 

Symphyogyna tenuicostata Steph. 1895 = S. brasiliensis Nees et Mont. 1836. 

Symphyogyna valida Steph. 1917 = S. brasiliensis Nees et Mont. 1836. 

Symphyogyna volkensii Steph. KEN(2), RWA(2), TAN(2), UGA(2,96), ZAI(2). 

Symphyogynopsis filicum (Nadeaud) Grolle 1986 = S. gottscheana (Mont. et Nees) Grolle 1987. 

Symphyogynopsis gottscheana (Mont. et Nees) Grolle BIO(36), CMN(146), COM(1), MAU(1), 

MDG(1), MLW(29), REU(1).


Symphyomitra africana Steph. 1917 = Lethocolea congesta (Lehm.) S.W.Arnell 1955. 

Symphyomitra glossophylla Spruce 1885 [Dubia et Excludenda]. 

Symphyomitra tabularis S.W.Arnell 1953 = Lethocolea congesta (Lehm.) S.W.Arnell 1955. 

Synhymenium aureonitens Griff. 1849 = Cyathodium aureonitens (Griff.) Mitt. 1886. 

Syzygiella concreta (Gottsche) Spruce KEN(2), MDG(1), RWA(2), TAN(2), ZAI(2). 

Syzygiella geminifolia (Mitt.) Steph. BIO(36), ETH(2), MDG(1,4), REU(1,4), RWA(2), SAO(2), 

TAN(2), ZAI(2). 

Syzygiella macrocalyx (Mont.) Spruce 1876 [Dubia et Excludenda]. 

Syzygiella ruwenzorensis Steph. 1917 = S. geminifolia (Mitt.) Steph. 1902. 

Syzygiella securifolia (Nees in Lindenb.) Inoue PRI(2). 

Syzygiella subintegerrima (Reinw. et al.) Spruce 1895 [Dubia et Excludenda]. 

Targionia capensis Huebener 1834 = T. hypophylla L. 1753. 

Syzygiella sp. SAF(416). 

Targionia elongata Bisch. ex Gottsche ETH(2). 

Targionia fiorii Gola ERI(121). 

Targionia hypophylla L. CHA(2,3), CMN(2), COM(4), CVI(37), DJI(2), ERI(2), ETH(2,113), GHA(2), 

KEN(2,665), LES(2,75), MDG(1), MLW(2,114), NAM(2), NGA(2), REU(1), SAF(2,170), TAN(2), 

UGA(42,96), ZAI(2), ZAM(2), ZIM(2). T. lorbeeriana was placed in synonymy with T. hypophylla by Garcia- 

Zamora et al. 1989 (557). The conspecificity of the two taxa was supported by Boisselier-Dubayle & Bischler 

1999 (625). 

Targionia hypophylla var. capensis (Huebener) C.Krauss 1846 = T. hypophylla L. 1753. 

Targionia lorbeeriana Müll.Frib. = T. hypophylla L. (557). 

Targionia orbicularis Schwein. 1821 = Notothylas orbicularis (Schwein.) Sull. 1846. 

Taxilejeunea acuta (Mitt.) Steph. 1914 = Lejeunea acuta Mitt. 1863. 

Taxilejeunea borbonica S.W.Arnell 1965 = Lejeunea eckloniana Lindenb. 1845. 

Taxilejeunea conformis (Mont. et Nees) Steph. ANG(2), BUR(2), CAF(2), CMN(2), COM(1), ETH(2), 

GHA(2), IVO(2), KEN(2,71), MAU(1), MDG(1), MLW(2,563), REU(1), RWA(2), SAF(2,75), SEY(1), 

SIE(2), TAN(2), TOG(2), UGA(2), ZAI(2). 

Taxilejeunea cuneistipula Steph. CMN(2). 

Taxilejeunea dusenii Steph. 1891 = Lejeunea ramosissima Steph. 1887. 

Taxilejeunea epiphyta Steph. 1914 = T. conformis (Nees et Mont.) Steph. 1914. 

Taxilejeunea furcicornuta Grolle MAU(1), MDG(1), REU(1,4). 

Taxilejeunea heterofolia Steph. 1914 = Lejeunea eckloniana Lindenb. 1845. Tixier 1995 (130) treated T. 

heterofolia as a distinct species and recorded it from Rwanda. Chuah-Petiot 1997 (71) recorded it from Kenya. 

Taxilejeunea krakakammae (Lindenb.) Sim 1926 = Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 1963. 

Taxilejeunea lindblomii Pearson 1924 = T. conformis (Nees et Mont.) Steph. 1914. 

Taxilejeunea longirostris Steph. 1914 = Lejeunea grossecristata (Steph.) E.W.Jones 1968. 

Taxilejeunea lyratiflora (Steph.) Tixier 1995 = Lejeunea lyratiflora Steph. 1892. 

Taxilejeunea mitracalyx Eifrig 1936 = Lejeunea alata Gottsche 1845. 

Taxilejeunea nicholsonii Pearson 1931 = Lejeunea grossecristata (Steph.) E.W.Jones 1968. 

Taxilejeunea pulchriflora Pearson CAF(2), CMN(2), GHA(2), GUI(2), IVO(2,53), KEN(2), LBR(2), 

RIM(10), RWA(2), SIE(2), TAN(2), TOG(2), UGA(115), ZAI(2). 

Taxilejeunea pusilla Pearson 1924, hom. illeg. [non Steph. 1916] = Lejeunea eckloniana Lindenb. 1845. 

Taxilejeunea ramosissima (Steph.) Steph. 1914 = Lejeunea ramosissima Steph. 1886. 

Taxilejeunea sikorae Steph. MDG(1). 

Taxilejeunea trichomanoides (Mont. et Nees) Steph. 1914 = T. conformis (Nees et Mont.) Steph. 1914. 

Taxilejeunea vallis-gratiae Steph. LES(75), SAF(2,75). 

Telaranea nematodes (Gottsche ex Austin) M.Howe BIO(72), BUR(2), COM(1), GAB(2), GHA(2), 

KEN(48), MDG(1), MLW(2,92), REU(4), RIM(10), RWA(2), SAF(2), SAO(2), SIE(2), TAN(2), 

TOG(2), UGA(42,50,96), ZAI(2), ZAM(2), ZIM(2). 

Telaranea sejuncta (Ångstr.) S.W.Arnell 1957 = Arachniopsis diacantha (Mont.) M.Howe 1902. 

Telaranea sejuncta sensu auct. [non (Ångstr.) S.W.Arnell 1957] = Arachniopsis diacantha (Mont.) M.Howe 

1902. 

Telaranea succulenta (Sim) Grolle 1964 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 2001 

(556).


Telaranea trifida (Steph.) R.M.Schust. CMN(2), RWA(2), UGA(42), ZAI(2). 

Tetralophozia cavallii (Gola) Váòa ETH(2), KEN(2), MLW(114), RWA(2), TAN(2), UGA(2,42,96), 

ZAI(2), ZIM(2). 

Thysananthus africanus (Sim) S.W.Arnell, hom. illeg. [non Steph. 1887] SAF(2). 

Thysananthus africanus Steph. 1887 = Caudalejeunea africana (Steph.) Steph. 1895. 

Thysananthus fruticosus (Lindenb. et Gottsche) Schiffn. 1893 = Dendrolejeunea fruticosa (Lindenb. et Gottsche) 

Lacout. 1908 [Dubia et Excludenda]. 

Thysananthus moenkemeyeri Steph. 1895, nom. nud. = Caudalejeunea africana (Steph.) Steph. 1895. 

Thysananthus monoicus Steph. 1912 = T. spathulistipus (Reinw. et al.) Lindenb. 1845. 

Thysananthus planus Sande Lac. 1855 = T. retusus (Reinw. et al.) B.Thiers et Gradst. 1989 [Dubia et 

Excludenda]. 

Thysananthus retusus (Reinw. et al.) B.Thiers et Gradst. 1989 [Dubia et Excludenda]. 

Thysananthus spathulistipus (Reinw. et al.) Lindenb. CMN(2), GAB(2), GHA(2), IVO(2,53), MAU(1), 

MDG(1), NGA(2), REU(1), RIM(10), SEY(1,4), SIE(2), ZAI(2). 

Thysananthus triquetrus (Mitt.) Steph. 1912 = T. spathulistipus (Reinw. et al.) Lindenb. 1845. 

Thysanolejeunea africana Sim 1926 = Thysananthus africanus (Sim) S.W.Arnell 1953, hom. illeg. [non Steph. 

1887]. 

Thysanolejeunea spathulistipa (Reinw. et al.) J.B.Jack et Steph. 1894 = Thysananthus spathulistipus (Reinw. et 

al.) Lindenb. 1845. 

Trachylejeunea grolleana Pócs 1999 = Xylolejeunea grolleana (Pócs) X.-L.He et Grolle 2001 (149). 

Trachylejeunea serrulata Steph. 1913 = Lepidolejeunea serrulata (Steph.) Grolle 1984. 

Tritomaria camerunensis S.W.Arnell CMN(2). 

Tritomaria exsecta (Schmidel) Loeske ETH(2), KEN(2), MLW(114), RWA(2), SAF(75), TAN(2), 

UGA(2,42). 

Tylimanthus aetiopicus S.W.Arnell 1961 = T. ruwenzorensis S.W.Arnell 1956. 

Tylimanthus africanus Pearson SAF(2). 

Tylimanthus anisodontus (Hook.f. et Taylor) Mitt. STH(2,49). 

Tylimanthus chenagonii Steph. 1905 = Plagiochila integerrima Steph. 1886. 

Tylimanthus congoensis Steph. 1922 = Plagiochila integerrima Steph. 1886. 

Tylimanthus lespagnolii Steph. 1905 = Plagiochila integerrima Steph. 1886. 

Tylimanthus limbatus Steph. 1911 = Marsupidium limbatum (Steph.) Grolle 1965. 

Tylimanthus madagascariensis Steph. 1901, nom. inval. = Kymatocalyx madagascariensis (Steph.) Gradst. et 

Váòa 1999 (74). 

Tylimanthus retusus Steph. 1922 = Plagiochila integerrima Steph. 1886. 

Tylimanthus ruwenzorensis S.W.Arnell ETH(2), KEN(2), MLW(114), REU(1,4), RWA(2), TAN(2), 

UGA(2,96). The collection of Tylimanthus from Ghana could not be named with certainty (25). 

Tylimanthus schweinfurthii Steph. 1905 = Plagiochila integerrima Steph. 1886. 

Tylimanthus wilmsii Steph. 1922 = Marsupidium limbatum (Steph.) Grolle 1965. 

Wiesnerella denudata (Mitt.) Steph. 1899 = W. javanica (Mitt.) Schiffn. 1896 (18). 

Wiesnerella javanica Schiffn. REU(1,18). 

Xylolejeunea grolleana (Pócs) X.-L.He et Grolle MDG(5,149). 

Xylolejeunea muricella X.-L.He et Grolle SEY(149) 

References 

The full literature reference can be found by translating the numerical reference shown in brackets after 

the country (or in additional notes on the taxon) to the author + year equivalent and cross-referencing with 

the Literature list below. References 3 onwards refer to literature which has provided additional data on 

the liverworts of sub-Saharan Africa since the publication of (1) and (2), including new country records 

and synonymy, or are the sources for notes and corrections. 

1 Grolle 1995 

2 Wigginton & Grolle 1996 

3 Ros et al. 1999 

4 Pócs 1995


5 Pócs 1999b 

6 Pócs 1997 

7 Arnell 1956 

8 Perold 1999g 

9 During & Moyo 1999 

10 Infante & Heras 1998 

11 Frahm & Porembski 1998 

13 Een & Kristoferson 1999 

14 He 1999 

17 Anon 1994 

18 Bischler 1998 

19 Bornefeld et al. 1996 

20 Cardoso 1915 

21 Chuah-Petiot 1996 

22 Dilg & Frahm 2000 

23 Wigginton 2000 

25 Jones & Harrington 1983 

26 Tixier 1977 

27 Vanden Berghen 1973 

29 Wigginton & Porley 2001 

33 Heras Pérez & Infante Sánchez 1996 

34 Seaward et al. 1996 

35 Gradstein 1975 

36 Müller 1996 

37 Frahm et al. 1996 

38 Infante et al. 1999 

39 Infante & Heras 1999 

42 Pócs & Lye 1999 

43 Een 2000 


48 Müller et al. 2000 


50 Porley et al. 1999 

51 Müller & Schaefer-Verwimp 1999 

52 Fischer 1999 

53 Gyarmati 2001 

55 Gradstein & Geissler 1997 

60 Frahm et al. 1996 


72 Infante et al. 1997 

73 O'Shea et al. 1997 

74 Gradstein & Váòa 1999 

75 Hodgetts et al. 1999 

76 Perold 1999f 

77 Perold 2000a 

78 Perold 2000b 


80 Perold 1997c 

81 Perold 1997d 

82 Perold 1997e 

83 Perold 1997f 









92 Hodgetts & Pócs 1999 

93 Long 2001 

94 Kürschner 2000 

95 Mitten 1863 

96 Lye & Pócs 1997 

97 Engel 1980 


100 Schmitt & Winkler 1968 

101 Tixier 1983 

102 Váòa 1998 

103 Pócs & Streimann 1999 

104 Jones 1985 

106 Jones 1968 

107 Scott 1960 

108 Pócs & Tóthmérész 1997 

109 Hodgetts 1997 

110 Porley 1997 

111 Váòa 1999 

112 Arts & Yamada 1998 

113 Wigginton 2001a 

114 O'Shea et al. 2001 

115 Wigginton et al. 2001 

117 Pócs & Lücking 1996 

118 Schuster 1995a 

119 Srivastava & Dixit 1996 

120 Gola 1914a 

121 Gola 1914b 

122 Gradstein 1994 

123 Grolle 1981 

124 Jovet-Ast 1946 

125 Muhle 1982 

126 Geissler 1997 

127 Oyesiku & Egunyomi 2001 

128 Stephani 1888 

129 Pócs 1999a 

130 Tixier 1995 

131 Hasegawa 1995 

132 Bischler-Causse 1993 

133 Kruijt & Gradstein 1986 

134 Jones 1957 

135 Tixier 1985 

136 Bizot & Pócs 1974 



139 Stephani 1923


141 Schuster 1992 

142 Fischer 1993 

143 Arnell 1957 

145 Castle 1939 

146 Grolle & Piippo 1986 


148 Long & Grolle 1992 

149 He & Grolle 2001 

152 Tixier 1979 


154 Kuwahara 1981 


156 Váòa et al. 1979 

157 Jones 1972 



160 Grolle & Long 2000 


163 Váòa 1982 

164 Schuster 1995b 

165 Jones 1953b 

166 Brummitt & Powell 1992 


168 Reiner-Drehwald 2000 

170 Arnell 1963 

171 O'Shea 1995 

172 Engel 1978 

416 Váòa 1985 

420 Engel & Braggins 1998 

421 Grolle 1983a 

424 Piippo 1991 

502 Cufodontis 1952 

503 Gerola 1947 


533 Veltman et al. 2000 

539 Hollis & Brummitt 1992 

540 O'Shea 1999 

541 O'Shea 1993 


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558 Kürschner 2001 

563 Wigginton 2001b 


625 Boisselier-Dubayle & Bischler 1999 


650 Jones 1952 




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655 Perold 1995 

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657 Piippo 1993 

658 Hasegawa 1994 

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660 Castle 1937 

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662 Müller et al. 1998 




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80 Perold, S.M. 1997c. Studies in the liverwort genus Fossombronia (Metzgeriales) from southern 

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81 Perold, S.M. 1997d. Studies in the liverwort genus Fossombronia (Metzgeriales) from southern 

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Appendix 1. Additional current names 

These current names have been added to the checklist of sub-Saharan liverworts since the publication of 

Grolle 1995 (1) and Wigginton & Grolle (2). 

1 sp. nov. / ssp. nov. / var. nov. 

2 new to Africa 

3 comb. nov. 

4 new synonymy 

5 noted in (2) as occurring in Africa, but without a published reference. 

6 overlooked in (2), or otherwise reassigned 

Acanthocoleus aberrans var. laevis Gradst. 1 

2, 4 

Anastrophyllum crebrifolium (Hook.f. et Taylor) Steph. 

Aphanolejeunea microscopica var. africana (Pócs) Pócs et Lücking 3 

Aphanolejeunea microscopica var. exigua (A.Evans) Pócs & Lücking 3 

2, 5 

Aphanolejeunea truncatifolia Horik. 

Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 3 

Asterella khasyana (Griff.) Pandé, K.P.Srivast. et Sultan Khan 2 

Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et Geissler 3 

Cheilolejeunea ventricosa (Schiffn.) X.-L.He 2 

Cololejeunea cuspidata Steph. ex Tixier 2 

Cololejeunea iradieri Infante et Heras 1 

Cololejeunea magna (Tixier) Infante & Heras 3 

Cololejeunea peraffinis var. elegans Benedix 6 

Cololejeunea planissima (Mitt.) Abeyw. 2 

Colura digitalis var. mucronata Pócs 1 

Colura mauritiana Pócs 1 

Colura zoophaga E.Fischer 1 

2, 4 

Cyathodium cavernarum Kunze 

Diplasiolejeunea albifolia ssp. orientalis Pócs ined. 1 

Diplasiolejeunea cobrensis subsp. antsirananae Pócs 1 

Diplasiolejeunea cogoensis Infante, Heras et Pócs 1 

Drepanolejeunea geisslerae Pócs 1 

Drepanolejeunea helenae Pócs 1 

Exormotheca bulbigena Bornefeld, O.H.Volk et R.Wolf 1 

Fossombronia angulifolia Perold 1


Fossombronia capensis var. spiralis Perold 1 

Fossombronia cederbergensis Perold 1 

Fossombronia elsieae Perold 1 

Fossombronia gemmifera Perold 1 

Fossombronia glenii Perold 1 

Fossombronia hyalorhiza Perold 1 

Fossombronia marindae Perold 1 

Fossombronia monticola Perold 1 

Fossombronia nyikaensis Perold 1 

Fossombronia renateae Perold 1 

Fossombronia rwandaensis Perold 1 

Fossombronia spinosa Perold 1 

Fossombronia straussiana Perold 1 

Fossombronia swaziensis Perold 1 

Fossombronia zuurbergensis Perold 1 

Gymnomitrion andinum (Herzog) Herzog 

Haplomitrium gibbsiae (Steph.) R.M.Schust. 2 

Kymatocalyx africanus Váòa et Wigginton 1 

Kymatocalyx dominicensis (Spruce) Váòa 2 

Kymatocalyx madagascariensis (Steph.) Gradst. et Váòa 3 

Lejeunea alata var. patriciae Pócs 1 

Lejeunea phyllobola Nees et Mont. 

Lejeunea pililoba Spruce 2 

Lepidozia stuhlmannii var. carnosa (Steph.) Pócs 3 

Lopholejeunea leioptera Gyarmati 1 

Marchesinia mackaii (Hook.) Gray 2 

Metzgeria africana Steph. 6 

Metzgeria nicomariei Veltman & Potgeiter 1 

Mnioloma fuscum (Lehm.) R.M.Schust. 3 

Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 3 

Paracromastigum tristanianum (R.M.Schust.) R.M.Schust. 6 

2, 4 

Radula ankefinensis Gottsche ex Steph. 

Radula carringtonii J.B.Jack 2 

Radula retroflexa Taylor 2 

Riccia biokoensis Perold 1 

Riccia sibayenii Perold 1 

Scapania calcicola (S.W.Arnell et J.Perss.) Ingham 2 

Sphenolobopsis pearsonii (Spruce) R.M.Schust. 2 

Wiesnerella javanica Schiffn. 

Xylolejeunea grolleana (Pócs) X.-L.He et Grolle 1 

Xylolejeunea muricella X.-L.He et Grolle 1 

2, 4 

2, 4 

2, 4 

Appendix 2. Additional synonymy 

The following list includes new synonymy published since the publication of Grolle 1995 (1) and 

Wigginton & Grolle 1996 (2), and earlier synonymy not included in (2). Transfers to other genera are also 

listed, e.g. Anthoceros species to Phaeoceros. Reference numbers to source documentation are shown in 

brackets. 

Acolea africana Steph. 1917 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996 (153).


Archilejeunea rotalis (Hook.f. et Taylor) Steph. 1911 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. 


Acrobolbus madagascariensis Steph. 1902 = Kymatocalyx madagascariensis (Steph.) Gradst. et Váòa 

1999 (74). 

Anastrophyllum leucocephalum (Taylor) Steph. 1893 = A. crebrifolium (Hook.f. et Taylor) Steph. 1893 

(172, 420). 

Aneura keniae Gola 1914 = Riccardia sp. 

Aneura laurentiana Steph. 1917 = A. pinguis (L.) Dumort. 1831 (652). 

Aneura submarginata S.W.Arnell 1952 = A. pinguis (L.) Dumort. 1831 (652). 

Anthoceros aethyopicus Gola 1914 = Phaeoceros sp. 

Anthoceros brunnthaleri Steph. 1913 = Phaeoceros sp. 

Anthoceros mildbraedii Steph. 1923 = Phaeoceros sp. 

Anthoceros parvifrons Steph. 1912 = Phaeoceros sp. 

Anthoceros rossoi Gola 1914 = Phaeoceros sp. 

Anthoceros tenuissimus Steph. 1893 = Phaeoceros sp. 

Anthoceros tigrinus Gola 1914 = Phaeoceros sp. 

Aphanolejeunea diaphana (A.Evans) R.M.Schust. 1980 = A. truncatifolia Horik. 1934 (117). 

Aphanolejeunea exigua var. africana Pócs 1984 = A. microscopica var. africana (Pócs) Pócs et Lücking 

1996 (117). 

Aphanolejeunea exigua A.Evans 1911 = A. microscopica var. exigua (A.Evans) Pócs & Lücking 1996 

(117). 

Aphanolejeunea subdiaphana (Ast) Pócs 1984 = A. truncatifolia Horik. 1934 (117). 

Arachniopsis succulenta (Sim) S.W.Arnell 1955 = Paracromastigum succulentum (Sim) J.J.Engel et 

G.L.S.Merr. 2001 (556). 

Aspiromitus expansus Steph. 1916 = Anthoceros sp. 

Calypogeia capensis S.W.Arnell 1953 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Calypogeia fusca (Lehm.) Steph. 1908 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Calypogeia repandifolia Steph. 1924 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118) 

Ceratolejeunea jungneri Steph. 1895 = C. calabariensis Steph. 1895 (27). 

Cololejeunea harrisii var. magna Tixier 1995 = C. magna (Tixier) Infante et Heras 1999 (39). 

Cololejeunea harrisii sensu Vanden Berghen 1977 = C. magna (Tixier) Infante et Heras 1999 (39) 

Cololejeunea subdiaphana Ast 1947 = Aphanolejeunea truncatifolia Horik. 1934 (117). 

Cyathodium africanum Mitt. 1886 = C. cavernarum Kunze 1834 (119). 

Dumortiera denudata Mitt. 1861 = Wiesnerella javanica Schiffn. 1896 (18). 

Fimbriaria incrassata Steph. 1899 = Asterella sp. (provisionally treated as A. abyssinica). 

Fimbriaria kamerunensis Steph. 1917 = Asterella sp. (provisionally treated as A. abyssinica). 

Fimbriaria preussii Schiffn. ex Steph. 1895 = Asterella sp. (provisionally treated as A. bachmannii). 

Fimbriaria volkensii Steph. 1899 = Asterella sp. (provisionally treated as A. dissoluta). 

Fossombronia zeyheri Steph. 1900 = F. crispa Nees 1846 (82,83). 

Gymnomitrion africanum (Steph.) Horik. 1943 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 

1996 (153). 

Hygrolejeunea patellirostris Steph. 1896 = Lejeunea sp. 

Hygrolejeunea staudtiana Steph. 1914 = Lejeunea sp. 

Jungermannia crispa Spreng. in sched. Hb. Zeyher = Fossombronia crispa Nees 1846 (83). 

Jungermannia fusca Lehm. 1829 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Jungermannia rotalis Hook.f. et Taylor 1845 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et 


Lejeunea brittoniae (A.Evans) Grolle 1995 = Lejeunea phyllobola Nees et Mont. 1842 (168). 

Lejeunea fusca (Lehm.) Drège 1843 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Lejeunea rotalis (Hook.f. et Taylor) Gottsche et al. 1845 = Aureolejeunea rotalis (Hook.f. et Taylor) 

Gradst. et Geissler 1997 (55). 

Lepidozia abyssinica S.W.Arnell 1960 = L. stuhlmannii var. stuhlmannii Steph. 1895 (92).


Lepidozia carnosa Steph. 1911 = L. stuhlmannii var. carnosa (Steph.) Pócs (96). 

Lepidozia stuhlmannii var. abyssinica (S.W.Arnell) Pócs 1994 = L. stuhlmannii var. stuhlmannii Steph. 

1895 (92). 

Lepidozia succulenta Sim 1926 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 2001 

(556). 

Leptocolea cristata var. occidentalis E.W.Jones 1953 = Cololejeunea occidentalis (E.W.Jones) Vanden 

Berghen 1978 (654). 

Leucolejeunea rotalis (Hook.f. et Taylor) S.W.Arnell 1961 = Aureolejeunea rotalis (Hook.f. et Taylor) 

Gradst. et Geissler 1997 (55). 

Madotheca mattirolii Gola 1914 = Porella mattirolii (Gola) S.W.Arnell 1960. 

Marsupella africana Steph. ex Bonner 1953 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 

1996 (153). 

Marsupella capensis S.W.Arnell 1957 = Gymnomitrion andinum (Herzog) Herzog 1934 (111). 

Marsupella hedbergii S.W.Arnell 1956 = Apomarsupella africana (Steph. ex Bonner) R.M.Schust. 1996 

(153). 

Marsupella subquadrata Steph., in sched. et icones ined. = Apomarsupella africana (Steph. ex Bonner) 

R.M.Schust. 1996 (153). 

Metacalypogeia fusca (Lehm.) N.Kitag. 1988 = Mnioloma fuscum (Lehm.) R.M.Schust. 1995 (118). 

Metzgeria camerunensis Steph. 1899 = M. australis Steph. (649). 

Metzgeria papulosa Steph. 1899 = M. leptoneura Spruce 1885 (424). 

Metzgeria thomeensis Steph. 1891 = M. australis Steph. 1889 (1,624). 

Microlepidozia succulenta (Sim) S.W.Arnell 1962 = Paracromastigum succulentum (Sim) J.J.Engel et 

G.L.S.Merr. 2001 (556). 

Phragmicoma rotalis (Hook.f. et Taylor) Mitt. 1875 = Aureolejeunea rotalis (Hook.f. et Taylor) Gradst. et 


Pycnolejeunea ventricosa Schiffn. 1898 = Cheilolejeunea ventricosa (Schiffn.) X.-L.He 1999 (14). 

Radula autoica Steph. 1913 = R. ankefinensis Gottsche ex Steph. 1884 (661). 

Radula delessertii Steph. 1893 = R. voluta Taylor ex Gottsche et al. 1845 (661). 

Radula exigua Steph. 1910 = R. madagascariensis Gottsche 1882 (661). 

Radula heteroica Steph. 1910 = R. ankefinensis Gottsche ex Steph. 1884 (661). 

Radula holstiana Steph. 1895 = R. ankefinensis Gottsche ex Steph. 1884 (661). 

Radula lespagnolii Steph. 1910 = R. voluta Taylor ex Gottsche et al. 1845 (661). 

Radula macroloba Steph. 1892 = R. voluta Taylor ex Gottsche et al. 1845 (661). 

Radula meyeri Steph. 1888 = R. fulvifolia (Hook.f. et Taylor) Gottsche et al. 1845 (661). 

Radula stipatiflora Steph. 1910 = R. voluta Taylor ex Gottsche et al. 1845 (661). 

Radula tamariscina Mitt. 1863 = R. boryana (F.Weber) Mont. 1842 (661). 

Radula woodiana Steph. 1910 = R. boryana (F.Weber) Mont. 1842 (660) 

Rectolejeunea brittoniae A.Evans 1911 = Lejeunea phyllobola Nees et Mont. 1842 (168). 

Riccardia submarginata S.W.Arnell 1953 = Aneura pinguis (L.) Dumort. (652). 

Riccia canescens Steph. 1898 = R. crinita Taylor 1846 (555). 

Riccia ciliata ssp. canescens (Steph.) R.M.Schust. 1992 = R. crinita Taylor 1846 (555). 

Riccia ciliata ssp. trichocarpa (M.Howe) R.M.Schust. 1992 = R. crinita Taylor 1846 (555). 

Riccia trichocarpa M.Howe 1898 = R. crinita Taylor 1846 (555). 

Schisma balboi Gola 1914 = Herbertus sp. 

Schisma kilimandjarense Steph. 1909 = Herbertus sp. 

Schisma mascarenicum var. kikujense Gola 1914 = Herbertus sp. 

Stenorrhipis madagascariensis (Steph.) Grolle 1963 = Kymatocalyx madagascariensis (Steph.) Gradst. et 

Váòa 1999 (74). 

Strepsilejeunea vatovae Gerola 1947 = Cheilolejeunea sp. 

Symbiezidium rotale (Hook.f. et Taylor) Trevis. 1877 = Aureolejeunea rotalis (Hook f. et Taylor) Gradst. 


Syzygiella sp. (416).


Targionia lorbeeriana Müll.Frib. = T. hypophylla L. (557). 

Telaranea succulenta (Sim) Grolle 1964 = Paracromastigum succulentum (Sim) J.J.Engel et G.L.S.Merr. 

2001 (556). 

Trachylejeunea grolleana Pócs 1999 = Xylolejeunea grolleana (Pócs) X.-L.He et Grolle 2001 (149). 

Tylimanthus madagascariensis Steph. 1901, nom. inval. = Kymatocalyx madagascariensis (Steph.) 

Gradst. et Váòa 1999 (74). 

Wiesnerella denudata (Mitt.) Steph. 1899 = W. javanica (Mitt.) Schiffn. 1896 (18). 

Appendix 3. Country abbreviations and numbers of taxa 

The list of geographical areas includes 65 countries, part-countries or islands. Country abbreviations 

follow Hollis & Brummitt 1992 (539), except for the four Gulf of Guinea islands and the Republic of 

South Africa, which are not provided with 3-letter abbreviations in the Hollis & Brummitt list. 

The abbreviations for Madagascar (MDG) and Cameroun (CMN) differ from those given in (1) and (2), 

and RIM (Rio Muni) is an additional abbreviation. RHO is adopted for former Rhodesia (Zambia + 

Zimbabwe) for one record where the location is unclear, and SEG for Senegambia (Senegal + Gambia). 

The 'Group/Index Muscorum zone' indicates the Index Muscorum zone together with the group name to 

which it belongs in the 'Full name' column, if relevant. The sub-Saharan part of part of Africa-4 is 

designated Africa-4N (= North) to differentiate it from the subantarctic part (not included in this 

checklist). 

Abbr. Full name No. of taxa Group/ Index Muscorum zone 

AGA Agalega Islands 0 Africa-3 

ALD Aldabra 2 Seychelles [Africa-3] 

AMI Amirante Islands 0 Seychelles [Africa-3] 

ANG Angola 61 Africa-2 

ANN Annobón Island 38 Equatorial Guinea [Africa-2] 

ASC Ascension Island 14 Africa-2 

BEN Benin 20 Africa-2 

BIO Bioko (Fernando Po) 93 Equatorial Guinea [Africa-2] 

BKN Burkina Faso 0 Africa-2 

BOT Botswana 17 Africa-4N 

BUR Burundi 81 Africa-2 

CAB Cabinda 0 Angola [Africa-2] 

CAF Central African Republic 56 Africa-2 

CGS Chagos Archipelago 0 Africa-3 

CHA Chad 15 Africa-2 

CMN Cameroun 229 Africa-2 

COM Comoro Islands 146 Africa-3 

CON Congo-Brazzaville 49 Africa-2 

CVI Cape Verde 39 Africa-2 

DIE Diego Garcia 0 Chagos Archipelago [Africa-3] 

DJI Djibouti 9 Africa-2 

EQG Equatorial Guinea 0 Africa-2 

ERI Eritrea 22 Africa-2 

ETH Ethiopia 131 Africa-2 

GAB Gabon 50 Africa-2 

GAM Gambia 1 Senegambia [Africa-2]


GHA Ghana 136 Africa-2 

GNB Guinea Bissau 5 Africa-2 

GUI Guinea 57 Africa-2 

IVO Ivory Coast 95 Africa-2 

KEN Kenya 227 Africa-2 

LBR Liberia 3 Africa-2 

LES Lesotho 82 Africa-4N 

MAS Mascarene Islands 13 Africa-3 

MAU Mauritius 161 Mascarenes [Africa-3] 

MDG Madagascar 346 Africa-3 

MLI Mali 1 Africa-2 

MLW Malawi 229 Africa-2 

MOZ Mozambique 55 Africa-2 

MTN Mauritania 2 Africa-2 

NAM Namibia 32 Africa-4N 

NGA Nigeria 134 Africa-2 

NGR Niger 6 Africa-2 

PRI Principé 32 Africa-2 

REU Réunion 237 Mascarenes [Africa-3] 

RHO former Rhodesia 1 Africa-2 

RIM Rio Muni 89 Equatorial Guinea [Africa-2] 

ROD Rodrigues 18 Mascarenes [Africa-3] 

RWA Rwanda 222 Africa-2 

SAF South Africa 320 Africa-4N 

SAO São Tomé 85 Africa-2 

SEG Senegambia 0 Africa-2 

SEN Senegal 4 Senegambia [Africa-2] 

SEY Seychelles 111 Africa-3 

SIE Sierra Leone 141 Africa-2 

SOC Socotra 16 Africa-2 

SOM Somalia 4 Africa-2 

STH St. Helena 22 Africa-2 

SUD Sudan 10 Africa-2 

SWZ Swaziland 22 Africa-4N 

TAN Tanzania 403 Africa-2 

TOG Togo 41 Africa-2 

UGA Uganda 291 Africa-2 

ZAI Democratic Republic of Congo 305 Africa-2 

(former Zaire) 

ZAM Zambia 58 Africa-2 

ZIM Zimbabwe 141 Africa-2

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