Preliminary Checklist of Philippine Pteridophytes - Co's Digital Flora ...
Preliminary Checklist of Philippine Pteridophytes - Co's Digital Flora ...
Preliminary Checklist of Philippine Pteridophytes - Co's Digital Flora ...
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<strong>Preliminary</strong> <strong>Checklist</strong> <strong>of</strong><br />
<strong>Philippine</strong> <strong>Pteridophytes</strong><br />
Leonardo L. Co<br />
Herbarium, Institute <strong>of</strong> Biology<br />
University <strong>of</strong> the <strong>Philippine</strong>s<br />
Diliman, Quezon City<br />
Family 1. OPHIOGLOSSACEAE<br />
(3 genera, 9 species, 1 variety, 2 formae + 2 interforma hybrids)<br />
1. Botrychium Swartz<br />
Botrychium daucifolium Wall. ex Hook. & Grev., Icon. Fil. (1829) t. 161; --Copel., FFP 1<br />
(1958) 20.<br />
Botrychium lanuginosum Wall. ex Hook. & Grev., Icon. Fil. (1828) t. 79; --Copel., FFP 1<br />
(1958) 19.<br />
.<br />
1. Helminthostachys Kaulfuss<br />
Helminthostachys zeylanica (L.) Hook., Genera (1840) t. 47; --Copel., FFP 1 (1958) 20; --<br />
Holttum, Rev. Fl. Mal. 2 (1954) 42, Fig. 2.<br />
Sri Lanka and India to Hainan, Taiwan, across Malesia to New Caledonia.<br />
Throughout the <strong>Philippine</strong>s. Terrestrial in open brushy places near sea level on damp<br />
ground.<br />
Revision:<br />
Wieffering,<br />
Ophioglossum costatum R Br.<br />
1. Ophioglossum Linnaeus
Ophioglossum gramineum Willd.<br />
var. gramineum<br />
Ophioglossum intermedium Hook., Icon. Pl. 10 (1854) t. 995; --Copel., FFP 1 (1958) 18;<br />
Ophioglossum nudicaule L.f., Suppl. (1781) 443; --Copel., FFP 1 (1958) 18;<br />
var. nudicaule<br />
var. macrorrhizum (Kunze) Clausen<br />
Ophioglossum pendulum L., Sp. Pl. ed. 2 (1763) 1518; --Copel., FFP 1 (1958) 19; --Holttum,<br />
Rev. Fl. Mal. 2 (1954) 40.<br />
forma pendulum<br />
Old World tropics from Madagascar through tropical Asia to Polynesia.<br />
Throughout the <strong>Philippine</strong>s. Epiphytic in everwet rainforests.<br />
forma pendulum x forma nutans Alderw.<br />
forma pendulum x forma falcatum (C Presl) Wief.<br />
Ophioglossum reticulatum L., Sp. Pl. (1753) 1063; --Copel., FFP 1 (1958) 17;<br />
forma reticulatum<br />
Ophioglossum petiolatum Hook., Exot. Fl. (1823) pl. 56; --Copel., FFP 1 (1958) 17;<br />
Ophioglossum nudicaule L. f. var. grandifolium Clausen, Mem. Torr. Bot. Club 19 (1938) 151; --Copel.,<br />
FFP 1 (1958) 18;<br />
? Ophioglossum ramosii Copel., PJS 56 (1935) 97, pl. 1, Fig. 1, 2; FFP 1 (1958) 18.<br />
Pantropic. Probably throughout the <strong>Philippine</strong>s.Terrestrial in humid mostly shaded<br />
sites. Only present during the wet season, 0-1600m.<br />
forma dilatatum (Miq.) Wief.<br />
forma complicatum (Miq.) Wief.<br />
Family 2. PSILOTACEAE (2 genera, 4 species)<br />
1. Psilotum Linnaeus<br />
Psilotum complanatum Sw., Syn. Fil. (1806) 188, 414, t. 4, Fig. 5.<br />
Psilotum nudum (L.) Griseb., Prodr. Aethéogam. (1805) 112.<br />
Psilotum triquetrum Sw., Schrad. J. Bot. 1800 (2) 1801<br />
2. Tmesipteris Bernhardi<br />
Tmesipteris lanceolata Dang, Le Botaniste 2 me Serie (1891)<br />
Tmesipteris tannensis auctt. non (Spreng.) Bernh.<br />
Family 3. EQUISETACEAE (1 genus, 1 species)<br />
Revision:<br />
Hauke, RI. 1963. A Taxonomic Monograph <strong>of</strong> the genus Equisetum subgenus Hippochaete.<br />
Beih. Nova Hedwigia 8: 1-123.
Laferrière, JE. 1998. Equisetaceae. <strong>Flora</strong> Malesiana series 2, 3: 287-288<br />
1. Equisetum Linnaeus<br />
Equisetum ramosissimum Desf.<br />
ssp. debile (Roxb. ex Vauch.) Hauke, Amer. Fern J. 52 (1962) 33.<br />
Family 4. MARATTIACEAE (4 genera, 5 species)<br />
1. Angiopteris H<strong>of</strong>fmann<br />
Angiopteris evecta Sw., Syn. Fil. (1806) 395; --Copel., FFP 1 (1958) 22, in obs., pro syn. A.<br />
palmiformis<br />
Angiopteris palmiformis (Cav.) C Chr., Dansk Bot. Arki 3, 9 (1937) 30, t. 3, Fig. 8, 9; --Copel., FFP 1<br />
(1958) 22.<br />
Angiopteris elmeriana Copel. in Elmer, LPB 5 (1913) 1679; --Copel., FFP 1 (1958) 23.<br />
Angiopteris uncinata Christ, PJS 2 c (1907) Bot. 186, non Vriese (1853); --Copel., FFP 1 (1958) 24.<br />
Angiopteris caudata Vriese, Monog. (1853) 20; --Copel., FFP 1 (1958) 24.<br />
Angiopteris cartilagidens Christ, Bull. Boiss. 6 (1898) 207, t. 4, Fig. 8; --Copel., FFP 1 (1958) 24.<br />
Angiopteris pruinosa Kunze, Bot. Zeit. 4 (1846) 417; --Copel., FFP 1 (1958) 25.<br />
Angiopteris angustifolia C Presl var. prunifera Hieron., Hedwigia 61 (1919) 257; --Copel., FFP 1 (1958)<br />
25.<br />
2. Christensenia Maxon<br />
Christensenia aesculifolia (Blume) Maxon, Proc. Biol. Soc. Wash. 18 (1905) 240; --<br />
Copel., FFP 1 (1958) 27.<br />
India (Assam) to Malesia.<br />
Macroglossum alidae Copel.<br />
3. Macroglossum Copeland<br />
4. Marattia Swartz<br />
Marattia pellucida C Presl, Suppl. Tent. (1845) 10; --Copel., FFP 1 (1958) 26.<br />
Marattia ternatea Christ, PJS 2 c (1907) Bot. 185; --Copel., FFP 1 (1958) 26.<br />
C LUZON southwards to MINDANAO: Mt. Matutum.<br />
Marattia sylvatica Blume, Enum. (1828) 256; --Copel., FFP 1 (1958) 27.<br />
LUZON: Cordillera Central. Common in mossy montane forests to ca. 2400 m.<br />
Family 5. OSMUNDACEAE (1 genus, 2-3 species)<br />
1. Osmunda Linnaeus<br />
Osmunda bromeliifolia (C Presl) Copel., PJS 2 c (1907) Bot. 147; --Copel., FFP 1 (1958) 30.<br />
Osmunda banksiifolia (C Presl) Kuhn, Ann. Lugd.-Bat. 4 (1869) 299; --Copel., FFP 1<br />
(1958) 30.<br />
Osmunda herbacea Copel., PJS 40 (1929) 291; FFP 1 (1958) 31;
Family 6. HYMENOPHYLLACEAE (5 genera, 60 species)<br />
Revision:<br />
Ebihara, A, J-Y Dubuisson, K Iwatsuki, S Hennequin, & M Ito. 2006: A Taxonomic Revision <strong>of</strong><br />
the Hymenophyllaceae. Blumea 51: 221-280.<br />
Conspectus to the genera:<br />
Subfamily I. Cardiomanoideae [petioles >7 cm; lamina usually <strong>of</strong> 4 cell-layers] –<br />
Only genus: Cardiomanes C Presl, not represented in the <strong>Philippine</strong>s.<br />
Subfamily II. Hymenophylloideae [petioles smaller; lamina usually 1 cell thick<br />
(except veins): formerly 2 polymorphous genera recognized – Hymenophyllum and<br />
Trichomanes but their limits still controversial. Following generic list is partly based<br />
on K Iwatsuki, 1984 and 1985].<br />
Abrodictyum C Presl =Cephalomanes<br />
Callistopteris C Presl =Cephalomanes<br />
1. Cephalomanes C Presl<br />
Cephalomanes apiifolia (C Presl) K Iwats.<br />
Trichomanes apiifolium C Presl, Hymen. (1843) 108, 136; --Copel., PJS 51 (1933) 227, pl. 42, Fig. 1;<br />
Callistopteris apiifolia (C Presl) Copel., PJS 67 (1938) 65; FFP 1 (1958) 75<br />
Cephalomanes atrovirens C Presl, Hymen. (1843) 17, pl. 5; --Copel., FFP 1 (1958) 77<br />
Trichomanes atrovirens (C Presl) Kunze, Bot. Zeit. 6 (1847) 371; --Copel., PJS 51 (1933) 251, pl. 52,<br />
Fig. 3; pl. 55, Fig. 2;<br />
Cephalomanes crassum (Copel.) MG Price<br />
Trichomanes crassum Copel.<br />
Cephalomanes cumingii (C Presl) K Iwats.<br />
Abrodictyum cumingii C Presl<br />
Trichomanes cumingii (C Presl) C Chr.<br />
Cephalomanes grande (Copel.) K Iwats.<br />
Trichomanes grande Copel., PJS 6 c (1911) 70; PJS 51 (1933) 224, pl. 40, Fig. 1-4;<br />
Nesopteris grandis (Copel.) Copel., PJS 67 (1938) 66; FFP 1 (1958) 76<br />
Cephalomanes javanicum (Blume) Bosch<br />
var. asplenioides (C Presl) K Iwats.<br />
Cephalomanes oblongifolium C Presl, Epim. Bot. (1849) 19, pl. 10; --Copel., FFP 1 (1923) 78<br />
Trichomanes asplenioides C Presl<br />
Trichomanes laciniatum Roxb.<br />
Cephalomanes meifolium (Bory ex Willd.) K Iwats.<br />
Trichomanes meifolium Bory ex Willd.<br />
Macroglena meifolia (Bory ex Willd.) Copel.<br />
Trichomanes gemmatum J Sm. ex Baker<br />
Cephalomanes obscurum (Blume) K Iwats.<br />
var. obscurum<br />
Trichomanes obscurum Blume<br />
Selenodesmium obscurum (Blume) Copel.<br />
Trichomanes rigidum Sw.
Cephalomanes thysanostomum (Makino) K Iwats.<br />
Trichomanes thysanostomum Makino, Bot. mag. Tokyo 12 (1898) 193;<br />
Nesopteris thysanostoma (Makino) Copel., PJS 67 (1938) 66;<br />
Cephalomanes<br />
Trichomanes extravagans Copel.<br />
Selenodesmium extravagans (Copel.) Copel.<br />
Cephalomanes setaceum (Bosch.) K Iwats.<br />
Trichomanes setaceum Bosch<br />
Macroglena setacea (Bosch) Copel.<br />
2. Crepidomanes (C Presl) C Presl<br />
Crepidomanes auriculatum (Blume) K Iwats.<br />
Trichomanes auriculatum Blume<br />
Vandenboschia auriculata (Blume) Copel.<br />
Crepidomanes brevipes (C Presl) Copel., PJS 67 (1938) 60; FFP 1 (1958) 71<br />
Trichomanes brevipes (C Presl) Baker, Syn. Fil. (1867) 84; --Copel., PJS 51 (1933) 182, pl. 20;<br />
Crepidomanes bipunctatum (Poir.) Copel., PJS 67 (1938) 59; FFP 1 (1958) 72<br />
Trichomanes bipunctatum Poir. in Lam., Encycl. 8 (1808) 69; --Copel., PJS 51 (1933) 177, pl. 18, Fig. 3,<br />
1-4;<br />
Crepidomanes christii (Copel.) Copel., PJS 67 (1938) 60; FFP 1 (1958) 73<br />
Trichomanes christii Copel., PJS 1 (1906) Suppl. 251; PJS 51 (1933) 185, pl. 21;<br />
Crepidomanes digitatum (Sw.) K Iwats.<br />
Trichomanes digitatum Sw., Syn. Fil. (1806) 370; --Copel., PJS 51 (1933) 159, pl. 7, Fig. 3, 4;<br />
Microtrichomanes digitatum (Sw.) Copel., PJS 67 (1938) 36; FFP 1 (1958) 55<br />
Crepidomanes humile (Forst.) Bosch<br />
Trichomanes humile Forst., Prodr. (1786) 12; --Copel., PJS 51 (1933) 164, pl. 12;<br />
Crepidopteris humilis (Forst.) Copel., PJS 67 (1938) 58; FFP 1 (1958) 70<br />
Trichomanes gracillimum Copel., PJS 51 (1933) 168, pl. 13;<br />
Crepidopteris gracillima (Copel.) Copel., PJS 67 (1938) 58; FFP 1 (1958) 70<br />
Crepidomanes johnstonense (Bailey) K Iwats.<br />
Trichomanes johnstonense Bailey<br />
Vandenboschia johnstonensis (Bailey) Copel.<br />
Crepidomanes minutum (Blume) K Iwats.<br />
Trichomanes minutum Blume, Enum. Pl. Jav. (1828) 223<br />
Gonocormus minutus (Blume) Bosch, Hymen. Javan. (1861) 7, pl. 3; --Copel., FFP 1 (1958) 68<br />
Crepidomanes pallidum (Blume) K Iwats.<br />
Pleuromanes pallidum (Blume) C. Presl, Epim. Bot. (1849) 258; --Copel., FFP 1 (1958) 67<br />
Trichomanes pallidum Blume, Enum. Pl. Jav. (1828) 226; --Copel., PJS 51 (1933) 141;<br />
Crepidomanes rupicolum (Raciborski) Copel., PJS 67 (1938) 59; FFP 1 (1958) 73<br />
Trichomanes rupicolum Raciborski, Pterid. Buitenz. (1898) 24<br />
Crepidomanes acutum (C. Presl) K Iwats.<br />
Trichomanes acutum C Presl, Epim. Bot. (1849) 258<br />
Crepidomanes minutum (Blume) K Iwats.<br />
Trichomanes minutum Blume forma alagensis (C Chr.) Copel.<br />
Trichomanes alagensis Christ, PJS 3 c (1908) Bot. 270; --Copel., PJS 51 (1933) 152, pl. 6, Fig. 4-7;<br />
Gonocormus alagensis (Christ) Copel., PJS 67 (1938) 57
Crepidomanes<br />
Trichomanes latifrons Bosch<br />
Vandenboschia latifrons (Bosch) Copel.<br />
Crepidomanes maximum (Blume) K Iwats.<br />
Trichomanes maximum Blume<br />
Vandenboschia maxima (Blume) Copel.<br />
Crepidomanes<br />
Microtrichomanes zamboanganum Copel., PJS 81 (1952) 2; FFP 1 (1958) 56<br />
Trichomanes zamboanganum (Copel.) C Morton<br />
Unplaced taxa:<br />
Trichomanes idoneum C Morton<br />
Trichomanes geminatum auct. non Baker<br />
Trichomanes bifidum C Presl non Willd.<br />
Trichomanes obtusum (Copel.) C Morton<br />
Crepidopteris Copeland =Crepidomanes<br />
Gonocormus Bosch =Crepidomanes<br />
3. Hymenophyllum Smith<br />
Hymenophyllum acanthoides (Bosch) Rosenst., Bull. Jard. Bot. Buitenz. 2, 2 (1911) 25;<br />
--Copel., PJS 64 (1937) 45, pl. 17;<br />
Meringium acanthoides (Bosch) Copel., PJS 67 (1938) 42; FFP 1 (1958) 63<br />
Hymenophyllum angulosum Christ, PJS 3 c (1908) Bot. 269; --Copel., PJS 64 (1937) 33, pl.<br />
13;<br />
Mecodium angulosum (Christ) Copel., PJS 64 (1937) 109, pl. 50; FFP 1 (1958) 49<br />
Hymenophyllum badium Hook. & Grev., Icon. Fil. (1828) pl. 76; --Copel., PJS 64 (1937)<br />
144, pl. 76;<br />
Mecodium badium (Hook. & Grev.) Copel., PJS 67 (1938) 23; FFP 1 (1958) 53<br />
Mecodium crispatum (Wall.) Copel., PJS 67 (1938) 23; FFP 1 (1958) 54<br />
Hymenophyllum crispatum Wall. in Hook. & Grev., Icon. Fil. (1828) pl. 72<br />
Hymenophyllum bartlettii (Copel.) C Morton<br />
Meringium bartlettii Copel., PJS 73 (1940) 464; FFP 1 (1958) 62<br />
Hymenophyllum bicolanum Copel., PJS 64 (1937) 31, pl. 10<br />
Meringium bicolanum (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 60<br />
Hymenophyllum blandum Raciborski, Pterid. Buitenz. (1898) 20; --Copel., PJS 64 (1937) 50,<br />
pl. 18;<br />
Meringium blandum (Raciborski) Copel., PJS 67 (1938) 43; FFP 1 (1958) 63<br />
Hymenophyllum bontocense Copel., PJS 64 (1937) 33, pl. 12;<br />
Meringium bontocense (Copel.) Copel., PJS 67 (1938) 41; FFP 1 (1958) 60<br />
Hymenophyllum campanulatum Christ, PJS 2 c (1907) Bot.155; --Copel., PJS 64 (1937) 32,<br />
pl. 11;<br />
Meringium campanulatum (Christ) Copel., PJS 67 (1938) 40; FFP 1 (1958) 60<br />
Hymenophyllum crispatum Wall. in Hook. & Grev., Icon. Fil. (1828) pl. 72; --Copel., PJS 64
(1937) 148, pl. 77;<br />
Mecodium crispatum (Wall.) Copel., PJS 67 (1938) 23;<br />
Hymenophyllum denticulatum Sw., Schrad. J. (1801) 100; --Copel., PJS 64 (1937) 41, pl. 15;<br />
Meringium denticulatum (Sw.) Copel., PJS 67 (1938) 42; FFP 1 (1958) 62<br />
Hymenophyllum edanoi (Copel.) C Morton<br />
Mecodium edanoi Copel., PJS 81 (1952) 2; FFP 1 (1958) 50<br />
Hymenophyllum emarginatum Sw., Schrad. J. 2 <strong>of</strong> 1800 (1801) 101; --Copel., PJS 64 (1937)<br />
118, pl. 58;<br />
Mecodium emarginatum (Sw.) Copel., PJS 67 (1938) 21; FFP 1 (1958) 52<br />
Hymenophyllum fimbriatum J Sm., J. Bot. 3 (1841) 418, nomen; --Hook., Sp. Fil. (1844) 102,<br />
pl. 36C; --Copel., PJS 64 (1937) 122, pl. 60;<br />
Mecodium fimbriatum (J Sm.) Copel., PJS 67 (1938) 21; FFP 1 (1958) 52<br />
Hymenophyllum holochilum (Bosch) C Chr., Index Fil. (1905) 226, 362; --Copel., PJS 64<br />
(1937) 34, pl. 14;<br />
Meringium holochilum (Bosch) Copel., PJS 67 (1938) 41; FFP 1 (1958) 62<br />
Hymenophyllum merrillii Christ, PJS 2 c (1907) Bot. 154; --Copel., PJS 64 (1937) 33, pl. 13;<br />
Meringium merrillii (Christ) Copel., PJS 67 (1938) 41; FFP 1 (1958) 61<br />
Hymenophyllum imbricatum Blume, Enum. Pl. Jav (1828) 220; --Copel., PJS 64 (1937) 137,<br />
pl. 70, 71;<br />
Mecodium imbricatum (Blume) Copel., PJS 67 (1938) 22; FFP 1 (1958) 53<br />
Hymenophyllum johorense Holttum, Gard. Bull. Sing. 4 (1929) 408, with fig.; --Copel., PJS 64<br />
(1937) 51, pl. 19;<br />
Meringium johorense (Holttum) Copel., FFP 1 (1958) 64<br />
Hymenophyllum reductum Copel., PJS 64 (1937) 53, pl. 20;<br />
Meringium reductum (Copel.) Copel., PJS 67 (1938) 43; FFP 1 (1958) 64<br />
Hymenophyllum klabatense Christ, Verh. Nat. Ges. Basel 11 (1894) 4; --Copel., PJS 64<br />
(1937) 30;<br />
Meringium klabatense (Christ) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59<br />
Hymenophyllum nitiduloides Copel., PJS 64 (1937) 112, pl. 52;<br />
Mecodium nitiduloides (Copel.) Copel., PJS 64 (1937) 112, pl. 52; FFP 1 (1958) 50<br />
Hymenophyllum pachydermicum Cesati, Atti Accad. Napoli 8 (1876) 8;<br />
Meringium pachydermicum (Cesati) Copel., PJS 67 (1938) 41; FFP 1 (1958) 57<br />
Hymenophyllum paniculiflorum C Presl, Hymen. (1843) 147; --Copel., PJS 64 (1937) 110,<br />
pl. 51;<br />
Mecodium paniculiflorum (C Presl) Copel., PJS 67 (1938) 19; FFP 1 (1958) 50<br />
Hymenophyllum polyanthos (Sw.) Sw., Schrad. J. ii <strong>of</strong> 1800 (1801) 102; --Copel., PJS 64<br />
(1937) 97, pl. 46, 47;<br />
Mecodium polyanthos (Sw.) Copel., PJS 67 (1938) 19; FFP 1 (1958) 49<br />
Hymenophyllum productum Kunze, Bot. Zeit. 6 (1848) 305; --Copel., PJS 64 (1937) 118, pl.<br />
54;<br />
Mecodium productum (Kunze) Copel., PJS 67 (1938) 20; FFP 1 (1958) 51<br />
Hymenophyllum ramosii Copel., PJS 64 (1937) 34, pl. 9, Fig. 4-6;<br />
Meringium ramosii (Copel.) Copel., PJS 67 (1938) 31; FFP 1 (1958) 61<br />
Hymenophyllum reinwardtii Bosch., Pl. Jungh. 1 (1856) 567; --Copel., PJS 64 (1937) 115, pl.<br />
55;<br />
Mecodium reinwardtii (Bosch.) Copel., PJS 67 (1938) 20; FFP 1 (1958) 51
Hymenophyllum serrulatum (C Presl) C Chr.<br />
Meringium meyenianum C Presl, Hymen. (1843) 116, pl. 8 B; --Copel., FFP 1 (1958) 58.<br />
Hymenophyllum meyenianum (C Presl) Copel., PJS 64 (1937) 25, pl. 8; FFP 1 (1958) 58<br />
Hymenophyllum edentulum (Bosch) C Chr., Index Fil. (1905) 360;<br />
Meringium edentulum (Bosch) Copel., PJS 67 (1938) 41; FFP 1 (1958) 58<br />
Hymenophyllum bakeri Copel., Sarawak Mus. J. 2 (1917) 309; --Copel., PJS 64 (1937) 29;<br />
Meringium bakeri (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59<br />
Hymenophyllum pulchrum Copel., PJS 64 (1937) 22;<br />
Meringium pulchrum (Copel.) Copel., PJS 67 (1938) 42; FFP 1 (1958) 57<br />
Hymenophyllum thuidium Harrington, J. Linn. Soc. Bot. 16 (1877) 25; --Copel., PJS 64<br />
(1937) 116, pl. 56;<br />
Mecodium thuidium (Harrington) Copel., PJS 67 (1938) 20; FFP 1 (1958) 52<br />
Hymenophyllum vittatum Copel., PJS 64 (1937) 31, pl. 9, Fig. 1-3; FFP 1 (1958) 59<br />
Meringium vittatum (Copel.) Copel., PJS 67 (1938) 40; FFP 1 (1958) 59<br />
Macroglena (C Presl) Copeland =Cephalomanes<br />
Mecodium C Presl =Hymenophyllum subg. Mecodium Copeland sect. Mecodium<br />
Meringium C Presl =Hymenophyllum<br />
Microgonium C Presl =Trichomanes L. emend. K Iwatsuki (1984).<br />
Microtrichomanes (Mettenius) Copeland =Crepidomanes<br />
Nesopteris Copeland =Cephalomanes<br />
Pleuromanes Blume =Crepidomanes<br />
Selenodesmium (Prantl) Copeland =Cephalomanes<br />
4. Sphaerocionium C Presl<br />
Sphaerocionium pilosissimum (C Chr.) Copel., PJS 67 (1938) 33; FFP 1 (1958) 54<br />
Hymenophyllum pilosissimum C Chr., Gard. Bull. 7 (1934) 213;<br />
5. Trichomanes Linnaeus<br />
Trichomanes mindorense Christ, PJS 3 c (1908) Bot. 270; --Copel., PJS 51 (1933) 209, pl. 34,<br />
Fig. 1, 2;<br />
Microgonium mindorense (Christ) Copel., PJS 67 (1938) 61; PJS 51 (1933) 164, pl. 12;<br />
Trichomanes craspedoneurum Copel., PJS 7 c (1912) Bot. 58; PJS 51 (1933) 208, pl. 33, Fig. 5-7;<br />
Microgonium craspedoneurum (Copel.) Copel., PJS 67 (1938) 61; FFP 1 (1958) 74.<br />
Trichomanes motleyi Bosch, Ned. Kruid. Arch. 5 (1861) 145; --Copel., PJS 51 (1933) 201, pl.<br />
xx, Fig. 1-4;
Microgonium motleyi Bosch, Hymen. Javan. (1861) 5, pl. 1; --Copel., FFP 1 (1958) 75<br />
Trichomanes beccarianum Cesati, Atti Accad. Napoli, Part 8 7 (1876) 8, pl. 1, Fig. 2; --Copel., PJS 51<br />
(1933) 200, pl. 29;<br />
Microgonium beccarianum (Cesati) Copel., PJS 67 (1938) 63; FFP 1 (1958) 74<br />
Vandenboschia = Crepidomanes<br />
Family 7. GLEICHENIACEAE (2 genera, 16 species, 4 varieties)<br />
Conspectus to the genera:<br />
Subfamily I. Gleichenioideae [frond at least once-forked]<br />
Dicranopteris Bernh.<br />
Gleichenia Sm.<br />
Subfamily II. Stromatopteridoideae [frond once-pinnate]: Monotypic, not<br />
represented in the <strong>Philippine</strong>s.<br />
1. Dicranopteris Bernhardi<br />
Dicranopteris curranii Copel., PJS 81 (1952) 4; --Copel., FFP 1 (1958) 46;<br />
Sumatra, Peninsular Malaysia, W Java, Lesser Sunda Isls (Flores), Borneo,<br />
Sulawesi, <strong>Philippine</strong>s. LUZON (Laguna), NEGROS, MINDANAO<br />
Dicranopteris linearis (Burm.) Underw., Bull. Torrey Bot. Club 34 (1907) 250; --Copel.,<br />
FFP 1 (1958) 45;<br />
var. linearis<br />
var. altissima Holttum<br />
var. latiloba Holttum<br />
var. rigida (Blume) Holttum<br />
Dicranopteris crassifolia (C Presl) Ching, Sunyatsenia 5 (1940) 275; --Copel., FFP 1 (1958) 46;<br />
var. subspeciosa Holttum<br />
Diplopterygium (Diels) Nakai =Gleichenia<br />
2. Gleichenia Smith<br />
Gleichenia dicarpa R Br., Prodr. Fl. Nov. Holl. (1810) 161; --Copel., FFP 1 (1958) 38.<br />
Australia, New Caledonia, Malesia: New Guinea and <strong>Philippine</strong>s. MINDANAO<br />
(Agusan del Norte, Mt Urdaneta; Davao, Mt Apo).<br />
Gleichenia elmeri Copel. in Elmer, LPB 3 (1910) 799;<br />
Hicriopteris elmeri (Copel.) Copel., FFP 1 (1958) 43<br />
Gleichenia hirta Blume, Enum. Pl. Jav. (1828) 250; --Copel., FFP 1 (1958) 40.<br />
Sticherus hirtus (Blume) Ching, Sunyatsenia 5 (1940) 203 (err. ‘hirsutus’)<br />
Moluccas and the <strong>Philippine</strong>s. LUZON, MINDORO, MINDANAO.
Gleichenia hispida Mett. ex Kuhn, Verh. K.K. Zool.-Bot. Ges. Wien 25 (1875) 600;<br />
Sticherus hispidus (Mett. ex Kuhn) Copel., Gen. Fil. (1947) 27; FFP 1 (1958) 41;<br />
Sticherus koordersii (Christ) Copel., FFP 1 (1958) 40.<br />
Sticherus pinnatus Copel.<br />
Sumatra, Java, Bali, N Sulawesi, Ternate and <strong>Philippine</strong>s.<br />
LUZON: Benguet, Albay (Mt Mayon), NEGROS.<br />
Gleichenia laevissima Christ, Bull. Ac. Inst. Geogr. Bot. 3, 11 (1902) 268;<br />
Hicriopteris laevissima (Christ) Ching, Sunyatesenia 5 (1940) 280; --Copel., FFP 1 (1958) 44<br />
Gleichenia loheri Christ, Bull. Herb. Boiss. 2, 6 (1906) 1009;<br />
Sticherus loheri (Christ) Copel., FFP 1 (1958) 42<br />
Sticherus perpaleaceus Copel., PJS 81 (1952) 3; FFP 1 (1958) 42.<br />
var. loheri<br />
LUZON: Benguet (Mt Pulog), Rizal, Quezon (Mt Banahaw).<br />
var. major Holttum, Reinwardtia 4 (1959) 272;<br />
NEGROS: Negros Occidental (Mt Kanlaon).<br />
Gleichenia longissima Blume, Enum. Pl. Jav. (1828) 250;<br />
Hicriopteris glauca (Thunb.) Ching, Sunyatsenia 5 (1940) 279; --Copel., FFP 1 (1958) 43<br />
S China to Indochina through Malesia to Melanesia (Fiji, Tahiti). In Malesia not<br />
collected in the Lesser Sunda Isls E <strong>of</strong> Flores, Sulawesi, the Moluccas and New<br />
Guinea.<br />
Throughout the <strong>Philippine</strong>s, usually at 800-1500m alt. in exposed places.<br />
Gleichenia microphylla R Br.<br />
Gleichenia paleacea (Copel.) Holttum, Reinwardtia 4 (1957) 265;<br />
Hicriopteris paleacea Copel., PJS 81 (1952) 3; FFP 1 (1958) 44<br />
Gleichenia peltophora Copel., PJS 40 (1929) 292, t. 1; --Copel., FFP 1 (1958) 39.<br />
New Guinea, S C Sulawesi, Borneo and <strong>Philippine</strong>s.<br />
MINDANAO: South Cotabato (Mt Matutum), alt. 1600m.<br />
Gleichenia sordida Copel. in Elmer, LPB 3 (1910) 798;<br />
Hicriopteris sordida (Copel.) Copel., FFP 1 (1958) 44<br />
Gleichenia truncata (Willd.) Sprengel, Syst. Veg. ed. 16, 4 (1827) 25;<br />
var. truncata<br />
Sticherus laevigatus (Willd.) C Presl, Tent. (1836) 52; --Copel., FFP 1 (1958) 41;<br />
Throughout Malesia, apparently most common in the west.<br />
Gleichenia vulcanica Blume, Enum. Pl. Jav. (1828) 251;<br />
Sumatra, Peninsular Malaysia, W java, Borneo, <strong>Philippine</strong>s, Sulawesi and New<br />
Guinea.<br />
MINDORO: Mindoro Oriental (Mt Halcon), PALAWAN (Mt Pulgar and Mt<br />
Mantalingahan, Co 1773), MINDANAO<br />
Of doubtful occurrence in the <strong>Philippine</strong>s:<br />
Gleichenia milnei Baker, Syn. Fil. (1874) 449;<br />
Sulawesi, Moluccas, New Guinea, Admiralty Isls, Solomons, New Hebrides.<br />
?LUZON<br />
Sticherus C Presl = Gleichenia
Family 8. DIPTERIDACEAE (including Cheiropleuriaceae)<br />
(1 genus, 2 species)<br />
1. Cheiropleuria C Presl<br />
Revision:<br />
Laferrière, JE. 1998. Cheiropleuriaceae. <strong>Flora</strong> Malesiana series 2, 3: 285-286.<br />
Cheiropleuria bicuspis (Blume) C Presl<br />
Dipteris conjugata Reinw.<br />
Dipteris lobbiana (Blume) Moore<br />
2. Dipteris Reinw.<br />
Family 9. MATONIACEAE (1 genus, 1 species)<br />
Revision:<br />
Kato, M. 1998. Matoniaceae. <strong>Flora</strong> Malesiana series 2, 3: 289-294<br />
Matonia foxworthyi Copel.<br />
Revision:<br />
1. Matonia R Brown<br />
Family 10. LYGODIACEAE<br />
1. Lygodium Swartz<br />
Lygodium auriculatum (Willd.) Alston, Reinwardtia 5 (1959) 16;<br />
Lygodium semihastatum (Cav.) Desv., Prodr. (1827) 203; --Copel., FFP 1 (1958) 34.<br />
Indochina, Micronesia, Malesia: E Borneo, <strong>Philippine</strong>s.<br />
LUZON, POLILLO, MINDORO, SAMAR, MINDANAO. Terrestrial, usually in thickets<br />
and disturbed forests.<br />
Lygodium circinnatum (Burm.) Sw., Syn. Fil. (1806) 153; --Copel., FFP 1 (1958) 33.<br />
Lygodium basilanicum Christ, PJS 2 c (1907) Bot. 179; --Copel., FFP 1 (1958) 34.<br />
Sri Lanka and India to S China, across Malesia to the Solomons.<br />
Throughout the <strong>Philippine</strong>s. Terrestrial, common in brush and open sites and<br />
disturbed lowland forests.<br />
Lygodium flexuosum (L.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 35.<br />
Sri Lanka, India, Himalayas to S China, Hongkong, Taiwan, Ryukyus, across<br />
Malesia, N Queensland, Melanesia. Terrestrial, in brush and open forests at low<br />
altitudes.<br />
Lygodium japonicum (Thunb.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 36.<br />
Lygodium mearnsii Copel., PJS 3 c (1908) Bot. 37; --Copel., FFP 1 (1958) 35.<br />
Sri Lanka, Himalayas (Kashmir eastwards) to E China (Zejiang), Korea, Japan and<br />
southwards to Thailand and S China; in Malesia: Banka, C & E Java, Sulawesi,
<strong>Philippine</strong>s, Moluccas, Lesser Sunda Isls and New Guinea. Naturalized in Florida<br />
and Texas. Terrestrial in open forests and throroughly cleared sites at low to medium<br />
altitudes.<br />
Lygodium longifolium (Willd.) Sw. in Schrader’s J. Bot. 1801, pt. 2 (1803) 305;<br />
S India, Hainan. In Malesia: Sumatra, Peninsular Malaysia, Riouw and Lingga Isls.,<br />
Borneo, <strong>Philippine</strong>s.<br />
LUZON (doubtful), PALAWAN.<br />
Lygodium merrillii Copel., PJS 2 c (1907) Bot. 146, t. 4; FFP 1 (1958) 37.<br />
Lygodium matthewii Copel., PJS 3 c (1908) Bot. 36; --Copel., FFP 1 (1958) 37.<br />
SW China (Guizhou), N Vietnam, Sumatra, Borneo (Sarawak), <strong>Philippine</strong>s.<br />
LUZON: Isabela; S Luzon, MINDORO, LEYTE.<br />
Lygodium microphyllum (Cav.) R Br., Prodr. Fl. Nov. Holl. (1810) 162;<br />
Lygodium scandens (L.) Sw., Schrader’s J. (1801) 106; --Copel., FFP 1 (1958) 36.<br />
Tropical Africa, SE Asia, N to Bangladesh and Hongkong and the Ryukyus, across<br />
Malesia to the Solomons, New Caledonia, N and E Australia S to New South Wales.<br />
LUZON to MINDANAO (along the Pacific side), MINDORO, CULION, BASILAN.<br />
Terrestrial in moe or less open sites.<br />
Lygodium salicifolium Sw.<br />
Lygodium versteegii Christ, Nova Guinea 8 (1909) 161; --Copel., FFP 1 (1958) 37.<br />
New Guinea and <strong>Philippine</strong>s.<br />
LUZON: Laguna (Mt Makiling), Quezon (Guinayangan), Camarines (Bicol National<br />
Park), MINDANAO: Agusan del Norte (Mt Hilong-hilong).<br />
Family 11. ANEMIACEAE (no <strong>Philippine</strong> representatives)<br />
Family 12. SCHIZAEACEAE (2 genera, 13 species)<br />
Revision:<br />
Holttum, RE. 1959. Schizaeaceae. <strong>Flora</strong> Malesiana series 2, 1: 37-61.<br />
1. Schizaea Smith<br />
Revision:<br />
Barcelona, JF, BF Hernaez & MG Price. 1996. xxxxxxxxx Asia Life Sciences 5 (1996) 27-34<br />
Schizaea dichotoma (L.) Sm., Mém. Acad. Turin 5 (1793) 422; --Copel., FFP 1 (1958) 32.<br />
Schizaea digitata (L.) Sw., Syn. Fil. (1806) 150, 380, t. 4, Fig. 1; --Copel., FFP 1 (1958) 32.<br />
Actinostachys digitata (L.) Wall. ex Brackenr.<br />
Schizaea inopinata Selling, Svensk. Bot. Tidskr. 40 (1946) 274, Fig. 1-7; --Copel., FFP 1 (1958)<br />
32.<br />
Actinostachys inopinata (Selling) Reed<br />
Schizaea malaccana Baker<br />
Family 13. MARSILEACEAE<br />
1. Marsilea Linnaeus
Marsilea crenata C Presl<br />
Salvinia<br />
Family 14. SALVINIACEAE (including Azollaceae)<br />
1. Salvinia Seguier<br />
1. Azolla Lamarck<br />
Revision:<br />
Saunders, RMK. 1998. Azollaceae. <strong>Flora</strong> Malesiana series 2, 3: 277-284<br />
Azolla pinnata R Br.<br />
Azolla pinnata R Br. var. imbricata (Roxb. ex Griff.) Bonap.<br />
Azolla imbricata (Roxb. ex Griff.) Nakai<br />
Family 15. THYRSOPTERIDACEAE<br />
(No <strong>Philippine</strong> representatives)<br />
Family 16. LOXOMATACEAE<br />
(No <strong>Philippine</strong> representatives)<br />
Family 17. CULCITACEAE<br />
(No <strong>Philippine</strong> representatives)<br />
Family 18. PLAGIOGYRIACEAE (1 genus, 7 species)<br />
Revision:<br />
Zhang XC & HP Nooteboom. 1998. Plagiogyriaceae. <strong>Flora</strong> Malesiana series 2, 3: 295-316<br />
Plagiogyria adnata (Blume) Bedd.<br />
Plagiogyria egenolfioides (Baker) Copel.<br />
var. latipinna (Copel.) XC Zhang & Noot.<br />
Plagiogyria tuberculata Copel.<br />
Plagiogyria euphlebia (Kunze) Mett.<br />
Plagiogyria christii Copel.<br />
Plagiogyria falcata Copel.<br />
Plagiogyria glauca (Blume) Mett.<br />
Plagiogyria nana Copel.<br />
Plagiogyria glauca var. nana (Copel.) C Chr.<br />
Plagiogyria pycnophylla (Kunze) Mett.<br />
Plagiogyria stenoptera (Hance) Diels<br />
1. Plagiogyria (Kunze) Mettenius
19. Family CIBOTIACEAE<br />
1. Cibotium Kaulfuss<br />
Cibotium barometz (L.) J Sm.<br />
Assam to S China, W Malesia.<br />
LUZON: Mountain Province (near Bontoc). Known in the <strong>Philippine</strong>s by a single<br />
collection, CAHUP 16635 Pancho.<br />
Cibotium cumingii Kunze<br />
Cyathea acuminata Copel.<br />
Cyathea apoensis Copel.<br />
Cyathea lobata Copel.<br />
Family 20. CYATHEACEAE (1 genus, 40 species)<br />
Cyathea atropurpurea Copel.<br />
Gymnosphaera atropurpurea (Copel.) Copel.<br />
Cyathea biliranensis Copel.<br />
Cyathea binuangensis Copel.<br />
Cyathea callosa Christ<br />
Cyathea foxworthyi Copel.<br />
Cyathea camiguinensis Copel.<br />
Cyathea caudata (J Sm.) Copel.<br />
Cyathea arborescens Copel.<br />
Cyathea brevipes Copel.<br />
Cyathea dupaxensis Copel.<br />
Cyathea christii Copel.<br />
Cyathea cinerea Copel.<br />
1. Cyathea Smith<br />
Cyathea contaminans (Wall.) Copel., PJS 4 c (1909) Bot. 60;<br />
Cyathea clementis Copel.<br />
New Guinea to India. Throughout the <strong>Philippine</strong>s. Usually common in open places<br />
and light forests from near sea-level to 1300m.<br />
Cyathea curranii Copel.<br />
Cyathea edanoi Copel.<br />
Cyathea elmeri (Copel.) Copel.<br />
Cyathea argyrolepis Copel.
Cyathea dimorphotricha Copel.<br />
Cyathea fenicis Copel.<br />
Cyathea ferruginea Christ<br />
Cyathea ferrugineoides Copel.<br />
Cyathea fuliginosa (Christ) Copel.<br />
Cyathea bicolor Copel.<br />
Cyathea dura Copel.<br />
Cyathea lanaensis Christ<br />
Cyathea lepidigera Copel.<br />
Cyathea mindanaensis (Christ) Copel.<br />
Cyathea squamicostata Copel.<br />
Cyathea sulitii Copel.<br />
Cyathea warihon Copel.<br />
Cyathea halconensis Christ<br />
Cyathea mearnsii Copel.<br />
Cyathea melanophlebia Copel.<br />
Cyathea heterochlamydea Copel.<br />
Cyathea merrillii Copel.<br />
Cyathea integra J Sm. ex Hook.<br />
Cyathea arguta Copel.<br />
Cyathea breviloba Copel.<br />
Cyathea bulusanensis Copel.<br />
Cyathea latebrosa (Wall.) Copel., PJS 4 c (1909) Bot. 52;<br />
Cyathea latipinnula Copel.<br />
Cyathea lepifera (J Sm. ex Hook.) Copel.<br />
Cyathea calocoma (Christ) Copel.<br />
Cyathea pteridioides Copel.<br />
Cyathea loheri Christ, Bull. Herb. Boiss. 2, 6 (1906) 1007;<br />
Cyathea campbellii Copel.<br />
Cyathea fructuosa Copel.<br />
Cyathea indusiosa Copel.<br />
Cyathea matthewii (Christ) Domin<br />
Cyathea mitrata Copel.<br />
Cyathea lurida (Blume) Copel.<br />
Gymnosphaera melanorhachis (Copel.) Copel.<br />
Cyathea masapilidensis Copel.<br />
Cyathea microchlamys Holttum<br />
Cyathea negrosiana Christ<br />
Cyathea obliqua Copel.<br />
Gymnosphaera obliqua (Copel.) Copel.<br />
Cyathea philippinensis Baker, Ann. Bot. 5 (1891) 186; --Copel., FFP 2 (1960) 205; --Holttum,<br />
Fl. Males. ser. 2, 1 (1963) 149. –Type: From a cultivated plant originating from the <strong>Philippine</strong>s,<br />
ex Hort. Veitch., Feb. 1878), cult. Kew (K).<br />
Cyathea bicolana Copel. in Elmer, LPB 9 (1920) 3108; --Copel., FFP 2 (1960) 204. –Type: Elmer 16528<br />
(L*, U*, US*, isotypes). Luzon: Sorsogon prov., Mt Bulusan, July 1916.<br />
Cyathea bontocensis Copel., PJS 46 (1931) 209; FFP 2 (1960) 205. –Type: Vanoverbergh 813. Luzon:
Mountain Province.<br />
Cyathea heteroloba Copel., PJS 38 (1929) 134; FFP 2 (1960) 204. –Type: BS 45633 Ramos & Edaño<br />
(NY*, isotype). Luzon: Quirino prov., Mt Alzapan, altitude 1500m, May-June 1925.<br />
Cyathea ramosiana Alderw., Bull. Jard. Bot. Buitenz. iii, 2 (1920) 137; --Copel., FFP 2 (1960) 206.<br />
–Type: BS 23411 Ramos (L*, US*, isotypes). Luzon: Sorsogon prov., July-Aug. 1915.<br />
LUZON: Mountain Province, Quirino (Mt Alzapan), Aurora (Mt Camatis), Quezon (Mt<br />
Banahaw), Camarines Sur (Mt Isarog, apparently common), Sorsogon (Mt Bulusan),<br />
MINDORO: Mindoro Oriental (Mt Halcon). Montane forests, alt. 800-1500m.<br />
Cyathea polypoda Baker in Stapf, Trans. Linn. Soc. Lond. Bot. 4 (1894) 250;<br />
Gymnosphaera dinagatensis (Copel.) Copel.<br />
Gymnosphaera glabra sensu Copel. Fern Fl. Philip. 2 (1960) 235, non Blume<br />
Cyathea robinsonii Copel.<br />
Cyathea pseudoalbizzia Copel.<br />
Cyathea rufopannosa Christ<br />
Cyathea setulosa Copel.<br />
Cyathea sibuyanensis Copel.<br />
Cyathea squamulata (Blume) Copel., PJS 4 c (1909) Bot. 37;<br />
Gymnosphaera squamulata Blume<br />
Cyathea suluensis Baker<br />
Cyathea sessilipinnula Copel.<br />
Cyathea trichophora Copel., PJS 6 c (1911) Bot. 363;<br />
Gymnosphaera trichophora (Copel.) Copel.<br />
Cyathea tripinnata Copel., PJS 1 (1906) Suppl. 251;<br />
Cyathea densisora Alderw.<br />
Cyathea leucostegia Copel.<br />
Cyathea leytensis Copel.<br />
Cyathea zamboangana Copel.<br />
Cyathea urdanetensis Copel.<br />
Calochlaena javanica<br />
Culcita javanica (Blume) Maxon<br />
Culcita copelandii (Christ) Maxon<br />
Culcita straminea (Labill.) Maxon<br />
Dicksonia mollis Holttum<br />
Dicksonia blumei (Kunze) T Moore<br />
Gymnosphaera Blume =Cyathea<br />
Family 21. DICKSONIACEAE<br />
1. Calochlaena<br />
2. Dicksonia L’Heritier
Family 22. METAXYACEAE (No <strong>Philippine</strong> representaives)<br />
Family 23 LINDSAEACEAE<br />
3 genera, 37 species, 1 ssp., 5 varieties, 1 possible natural hybrid.<br />
Lindsaea Dryander –29 spp., 4 vars.<br />
Sphenomeris Maxon – 3 spp., 1 ssp.<br />
Tapeinidium (C Presl) C Chr. –5 spp. 1 var., 1 poss. hybrid<br />
Revision:<br />
Kramer, KU. 1971. Lindsaea Group. <strong>Flora</strong> Malesiana series 2, 1: 177-254<br />
Lindsaea adiantoides J Sm.<br />
Lindsaea tropidorhachis Alderw.<br />
Lindsaea apoensis Copel.<br />
Lindsaea havicei Copel.<br />
Isoloma J Smith =Lindsaea<br />
1. Lindsaea Dryander<br />
3. Lindsaea bouillodii Christ, Not. Syst. 1 (1909) 59;<br />
Thailand and N Vietnam to Malesia: Peninsular Malaysia, Sumatra, W Java, Borneo,<br />
Banka and Natuna Is. New to the <strong>Philippine</strong>s: PALAWAN: Mt. Mantalingahan, alt.<br />
1200m, Co 1793 (A, PNH, PUH).<br />
Lindsaea capillacea Christ<br />
Lindsaea cultrata (Willd.) Sw., Syn. Fil. (1806) 119;<br />
Lindsaea decomposita sensu Copel. (1960) non Willd., pro parte.<br />
Lindsaea divergens Hook. & Grev.<br />
Isoloma divergens (Hook. & Grev.) J Sm.<br />
Lindsaea doryphora Kramer<br />
Lindsaea parasitica sensu Copel. (1960) non Wall.<br />
Lindsaea ensifolia Sw.<br />
ssp. ensifolia<br />
Lindsaea fissa Copel.<br />
Lindsaea hymenophylloides sensu Copel. (1960), non Blume<br />
Lindsaea gueriniana (Gaudich.) Desv., Mem. Soc. Linn. Paris 6 (1827) 312;<br />
Isoloma guerinianum (Gaudich.) Fée<br />
Isoloma fuligineum (Copel.) Copel.<br />
Lindsaea heterophylla Dryander<br />
Lindsaea javanensis Blume<br />
Lindsaea lobata Poir. in Lam., Encycl. Suppl. 3 (1813) 448;<br />
Lindsaea longifolia Copel.<br />
Lindsaea lucida Blume, Enum. Pl. Jav. (1828) 216;
Lindsaea securifolia C Presl ex Goldm.<br />
Lindsaea gracilis Blume in Copel. (1960), excl. syn.<br />
ssp. lucida<br />
Lindsaea merrillii Copel.<br />
ssp. merrillii<br />
Lindsaea oblanceolata Alderw., Bull. Jard. Bot. Buitenz. 2, 23 (1916) 15;<br />
Lindsaea obtusa J Sm. ex Hook., Sp. Fil. 1 (1846) 224;<br />
Lindsaea decomposita sensu Copel. (1960) non Willd., excl. syn.<br />
Lindsaea davallioides sensu Copel. (1960) non Blume<br />
Lindsaea odorata Roxb., Calc. J. Nat. Hist. 4 (1844) 511;<br />
Lindsaea cultrata sensu Copel. (1960), pro parte<br />
Lindsaea cultrata (Willd.) Sw. var. varia Copel.<br />
? Lindsaea cultrata var. minor Hook.<br />
Lindsaea loheriana Christ<br />
Lindsaea orbiculata (Lam.) Mett. ex Kuhn<br />
var. orbiculata<br />
var. conmixta (Tagawa) Kramer<br />
Lindsaea ovata J Sm.<br />
Isoloma ovatum (J Sm.) C Presl<br />
Lindsaea pellaeformis Christ<br />
Luzon: Isabela prov., Divilacan, Aubarede peninsula, 50 m alt., L. Co 3580,<br />
distributed as L. gueriniana (A, PNH, PUH, CAHUP, L, US, K, MICH). In here<br />
sympatrically occurring with L. gueriniana, which according to Kramer (1971), might<br />
just be an extreme form <strong>of</strong> that variable species. New to the <strong>Philippine</strong>s. Previously<br />
known only from Sulawesi.<br />
Lindsaea philippinensis Kramer<br />
Isoloma angustum (Copel.) Tard., comb. superfl.; non L. angusta Copel. (1952).<br />
According to Kramer (1971), doubtfully distinct from L. jamesonioides Baker <strong>of</strong><br />
Sabah.<br />
Lindsaea polyctena Kramer<br />
Lindsaea pulchella (J Sm.) Mett. ex Kuhn<br />
var. pulchella<br />
Lindsaea adiantifolia (Hook.) Copel.<br />
Lindsaea adiantoides sensu Copel. (1960), pro parte.<br />
var. blanda (Mett. ex Kuhn) Kramer<br />
Lindsaea cyathicola Copel.<br />
Lindasea ramosii Copel.<br />
Very doubtfully distinct from L. longifolia.<br />
Lindsaea repens (Bory) Thwaites<br />
var. sessilis (Copel.) Kramer, Blumea 15 (1968) 568;<br />
Lindsaea longa Copel.<br />
Lindsaea cultripinna Copel.<br />
var. pectinata (Blume) Mett. ex Kuhn in Miq., Ann. Mus. Bot. Lugd.-Bat. 4 (1869) 277;<br />
Lindsaea pectinata Blume
Lindsaea macraeana (Hook. & Arn.) Copel., at least in part.<br />
?Lindsaea roxasii Copel.<br />
Lindsaea angusta Copel.<br />
Aberrant form from Mindanao called L. angusta Copel. may be distinguished as<br />
forma angusta (Copel.) Kramer, Blumea 15 (1968) 568.<br />
Lindsaea rigida J Sm. ex Hook., Sp. Fil. 1 (1846) 217, t. 63A;<br />
Lindsaea monosora Copel.<br />
Lindsaea tetragona Kramer<br />
var. tetragona<br />
Lindsaea tenuifolia sensu Copel. (1960) non Blume, excl. syn.<br />
Doubtful species:<br />
Lindsaea decrescens Copel.<br />
“As stated by Copeland, this is intermediate between L. repens (‘L. macraena,<br />
L. longa’) and L. fissa. The isotype in MICH from Copeland’s herbarium is rather<br />
scrappy. I doubt whether it represents a distinct species” –Kramer, Fl. Males. Ser. ii 1<br />
(1971) 253.<br />
Lindsaea longa Copel.<br />
Probably only an usually deeply incised form <strong>of</strong> L. repens (Bory) Thwaites var.<br />
sessilis (Copel.) Kramer.<br />
2. Sphenomeris Maxon<br />
Sphenomeris biflora (Kaulf.) Tagawa<br />
Sphenomeris chusana sensu Copel. (1960), pro parte<br />
Sphenomeris chinensis (L.) Maxon<br />
Sphenomeris chusana (L.) Copel.<br />
ssp. chinensis<br />
Tropical to subtropical Asia and Oceania.<br />
Throughout the <strong>Philippine</strong>s.<br />
Terrestrial or epilithic in exposed sites, from low altitudes to 2100m.<br />
ssp. divaricata (Christ) Kramer<br />
Sphenomeris retusa (Cav.) Maxon<br />
Tapeinidium acuminatum Kramer<br />
Tapeinidium calomelanos Kramer<br />
Tapeinidium gracile (Blume) Alderw.<br />
Tapeinidium luzonicum (Hook.) Kramer<br />
var. luzonicum<br />
var. leptophyllum Kramer<br />
Tapeinidium pinnatum (Cav.) C Chr.<br />
3. Tapeinidium (C Presl) C Christensen
Tapeinidium biserratum (Blume) Alderw.<br />
Probably a natural hybrid. According to Kramer (Fl. Males. Ser. ii 1 (1971): “The type<br />
<strong>of</strong> this ‘species’ is intermediate between T. luzonicum and T. pinnatum; there are<br />
25-30 intermediates… Their status is not clear. They occur throughout the common<br />
range <strong>of</strong> the two species”.<br />
Orthiopteris campylura (Kunze) Copel.<br />
Family 24. SACCOLOMATACEAE<br />
Orthiopteris Copeland =Saccoloma<br />
1. Saccoloma<br />
Family 25. DENNSTAEDTIACEAE (including Hypolepidaceae, Monachosoraceae,<br />
and Pteridiaceae)<br />
(7 genera, xx species, x subspecies, x varieties)<br />
Dennstaedtia articulata Copel.<br />
1. Dennstaedtia Bernhardi<br />
Dennstaedtia dennstaedtioides (Copel.) Copel.<br />
Dennstaedtia elmeri Copel.<br />
Dennstaedtia fusca Copel.<br />
Dennstaedtia glabrata (Cesati) C Chr.<br />
Dennstaedtia hooveri Christ<br />
Dennstaedtia macgregori Copel.<br />
Dennstaedtia merrillii Copel.<br />
Dennstaedtia philippinensis Copel.<br />
Dennstaedtia resinifera (Blume) Mett. ex Kuhn<br />
Dennstaedtia cuneata (J Sm. ex Hook.) T Moore<br />
Dennstaedtia rufidula C Chr. in C Chr. & Holltum, Gard. Bull. Sing. 7 (1934) 226, t. 51<br />
Dennstaedtia scabra (Wall.) T Moore<br />
Dennstaedtia scandens (Blume) T Moore<br />
Dennstaedtia smithii (Hook.) T Moore<br />
Dennstaedtia williamsii Copel.
2. Histiopteris (Agardh) J Smith<br />
Histiopteris incisa (Thunb.) J Sm., Hist. Fil. (1875) 295;<br />
Histiopteris montana Copel.<br />
Revision:<br />
Blumea 32 (1987) 227-276<br />
1. Hypolepis Bernhardi<br />
Hypolepis alpina (Blume) Hook., Sp. Fil. 2 (1852) 63;<br />
Hypolepis bamleriana Rosenst., Feddes Repert. 10 (1912) 325;<br />
Hypolepis glandulifera Brownsey & Chinnock, J. Adelaide Bot. Gard. 10 (1987) 16;<br />
Hypolepis malesiana Brownsey<br />
Hypolepis pallida (Blume) Hook.?, Sp. Fil. 2 (1852) 64;<br />
Doubtfully occurring in the <strong>Philippine</strong>s (Negros: Amlan, MG Price 2479)<br />
Hypolepis polypodioides (Blume) Hook.<br />
Hypolepis punctata (Thunb.) Mett. ex Kuhn<br />
Doubtfully occurring in the <strong>Philippine</strong>s (Luzon: Ilocos Norte, Mt. Bubungkayo,<br />
Iwatsuki et al. P375 (L).<br />
Hypolepis tenuifolia (Forst. f.) Bernh. ex C Presl<br />
Doubtfully occurring in the <strong>Philippine</strong>s:<br />
Hypolepis punctata (Thunb.) Mett.<br />
4. Microlepia C Presl<br />
Microlepia herbacea Ching & C Chr. apud Tard. ex C Chr.<br />
Microlepia trapeziformis (Roxb.) Kuhn<br />
Microlepia platyphylla (D Don) J Sm.<br />
Microlepia protracta Copel.<br />
Microlepia speluncae (L.) T Moore<br />
Microlepia sablanensis Christ<br />
?Microlepia manilensis C Presl<br />
Microlepia strigosa (Thunb.) C Presl<br />
Microlepia todayensis Christ<br />
Microlepia trichotricha J Sm.<br />
5. Monachosorum Kunze
Monachosorum henryi Christ<br />
Luzon: Cordillera Central Mtn. Range, Mt. Data, alt. 2220 m, Co 4086 (L, PNH,<br />
PUH). New to the <strong>Philippine</strong>s. Previously known from Taiwan and Mainland China<br />
(Guangdong, Guangxi, Guizhou and NE & W Yunnan).<br />
Monachosorum subdigitatum (Blume) Kuhn, Chaetopt. (1882) 345;<br />
Paesia radula (Baker) C Chr.<br />
Paesia luzonica Christ<br />
Paesia elmeri Copel.<br />
6. Paesia St. Hilaire<br />
7. Pteridium Gleditsch ex Scopoli<br />
Revision:<br />
Tryon. 1941. Rhodora 43: 1-70.<br />
Pteridium aquilinum (L.) Kuhn<br />
var. wightianum (J Agardh) Tryon<br />
Pteridium caudatum (L.) Maxon<br />
Pteridium aquilinum (L.) Kuhn var. yarrabense Domin<br />
Family 26. PTERIDACEAE (including Acrostichaceae, Adiantaceae, Antrophyaceae,<br />
Ceratopteridaceae, Cheilanthaceae, Hemionitidaceae, Parkeriaceae,<br />
Sinopteridaceae, Taenitidaceae, Vittariaceae)<br />
(x genera, xx species)<br />
Subfamily I. PLATYZOMATOIDEAE – not represented in the <strong>Philippine</strong>s<br />
Subfamily 2. PARKERIOIDEAE (1 genus, 1 species in the <strong>Philippine</strong>s)<br />
Ceratopteris thalictroides (L.) Brongn.<br />
Ceratopteris siliquosa (L.) Copel.<br />
1. Ceratopteris Brongniart<br />
PTERIDACEAE Subfamily 3. ADIANTOIDEAE (x genus, xx species)<br />
1. Adiantum capillus-veneris L.<br />
Adiantum caudatum L.<br />
Adiantum cupreum Copel.<br />
Adiantum edgeworthii Hook.<br />
Adiantum flabellulatum L.<br />
1. Adiantum Linnaeus
Adiantum hispidulum Sw.<br />
Adiantum malesianum Ghatak<br />
Adiantum mindanaoense Copel.<br />
Adiantum philippense L.<br />
Adiantum scabripes Copel.<br />
Adiantum setulosum J Sm.<br />
Adiantum diaphanum sensu Copel. non Blume, Enum. Pl. Jav. (1828) 215;<br />
Adiantum soboliferum Wall.<br />
Adiantum stenochlamys Baker<br />
Adiantum opacum Copel., PJS 1 (1906) Suppl. 255, t. 3.<br />
2. Antrophyum Kaulfuss<br />
Antrophyum immersum (Bory ex Willd.) Mett.<br />
Antrophyum nanum Fée<br />
Antrophyum latifolium Blume<br />
Antrophyum ledermannii Hieron.<br />
Antrophyum plantagineum (Cav.) Kaulf.<br />
Antrophyum reticulatum (Forst.) Kaulf.<br />
Antrophyum alatum Brackenr..<br />
Antrophyum callifolium Blume<br />
Antrophyum sessilifolium (Cav.) Sprengel, syn. nov.<br />
Antrophyum clementis Christ<br />
Antrophyum semicostatum Blume<br />
Antrophyum williamsii Benedict<br />
3. Monogramma Comm. ex Schkuhr.<br />
Monogramma capillaris Copel., PJS 6c (1911) Bot. 147<br />
Monogramma dareicarpa Hook.<br />
Monogramma<br />
Vaginularia paradoxa (Fée) Mett.<br />
Monogramma trichoidea J Sm. ex Hook.<br />
Vaginularia trichoidea (J Sm. ) Fée<br />
Vaginularia Fée = Monogramma
Vittaria alternans Copel.<br />
Vittaria anguste-elongata Hayata<br />
Vittaria stenophylla Copel.<br />
Vittaria elongata Sw.<br />
Vittaria zosterifolia Willd.<br />
Vittaria ensiformis Sw.<br />
Vittaria hecistophylla Copel.<br />
Vittaria pachystemma Christ<br />
Vittaria scolopendrina (Bory) Thwaites<br />
Vittaria subcoriacea Christ<br />
Vittaria taeniophylla Copel..<br />
4. Vittaria Smith<br />
PTERIDACEAE Subfamily 4. CONIOGRAMMOIDEAE, subfam. nov.<br />
1. Coniogramme Fée<br />
Coniogramme fraxinea (D Don) Diels in Engl. & Prantl, Nat. Pflanzenfam. 1 (1899) 262;<br />
Coniogramme macrophylla (Blume) Hieron., Hedwigia 57 (1916) 291;<br />
Coniogramme subcordata Copel.<br />
PTERIDACEAE Subfamily 5. CHEILANTHOIDEAE<br />
(5 genera, 14 species in the <strong>Philippine</strong>s)<br />
Aleuritopteris farinosa (Forssk.) Fée<br />
Cheilanthes agetae (Saiki) Kuo<br />
Cheilanthes farinosa (Forssk.) Kaulf.<br />
1. Aleuritopteris<br />
Aleuritopteris alb<strong>of</strong>usca (Baker) Pic.-Serm.<br />
Cheilanthes anceps Blanf.<br />
Cheilanthes alb<strong>of</strong>usca Baker<br />
Aleuritopteris rufa (D Don) Copel.<br />
Cheilanthes dubia Hope<br />
Cheilanthes rufa D Don<br />
Cheilanthes belangeri (Bory) C Chr.<br />
Cheilanthes javensis (Willd.) T Moore<br />
Cheilanthes densa (J Sm.) Fée<br />
2. Cheilanthes Swartz
Cheilanthes dilimanensis PM Zamora<br />
Cheilanthes mysurensis Wall. ex Hook.<br />
Cheilanthes tenuifolia (NL Burm.) Sw., Syn. Fil. (1806) 129, 332;<br />
3. Doryopteris J Sm.<br />
Doryopteris concolor (Langsdorff & Fischer) Kuhn<br />
Cheilanthes concolor (Langsdorff & Fischer) R & A Tryon<br />
Doryopteris cuspidata Copel.<br />
Doryopteris ludens (Wall.) J Sm.<br />
Gymnopteris vestita (Wall.) Underw.<br />
4 Gymnopteris Bernhardi<br />
Hemionitis arifolia (Burm. f.) T Moore =Parahemionitis<br />
5. Parahemionitis Panigrahi<br />
Revision:<br />
Panigrahi, G.1993. American Fern Journal 83: 90-92<br />
Parahemionitis arifolia (Burm. f.) Panigr.<br />
Hemionitis arifolia (Burm. f.) T Moore<br />
PTERIDACEAE Subfamily 6. Pteridoideae (incl. Taenitoideae)<br />
(x genera, xx species)<br />
Acrostichum aureum L.<br />
Acrostichum speciosum Willd.<br />
Onychium japonicum (Thunb.) Kunze<br />
Onychium siliculosum (Desv.) C Chr.<br />
1. Acrostichum Linnaeus<br />
Copelandiopteris Stone = Pteris<br />
2. Onychium Kaulfuss<br />
3. Pityrogramma Link
Pityrogramma calomelanos (L.) Link, Handb. Gewachs. 3 (1833) 20;<br />
Pteris armata C Presl<br />
Pteris biaurita L.<br />
Pteris pacifica Hieron.<br />
Pteris cumingii Hieron<br />
Pteris blumeana J Agardh<br />
Pteris brevis Copel.<br />
Pteris calocarpa (Copel.) MG Price<br />
Schizostegeopsis calocarpa Copel.<br />
Copelandiopteris calocarpa (Copel.) Stone<br />
4. Pteris Linnaeus<br />
Pteris cretica L.<br />
Pteris stenophylla sensu Copel., FFP 1 (1958) 129, non Hook. & Grev.<br />
Pteris dataensis Copel.<br />
Pteris distans J Sm. ex Hook.<br />
Possibly a hybrid or a bizarre freak.<br />
Pteris edanoi Copel., PJS 81 (1952) 8, `edanyoi’<br />
Pteris elmeri Christ<br />
Pteris endoneura MG Price<br />
Copelandiopteris endoneura (MG Price) Stone<br />
Pteris ensiformis Burm.<br />
var. ensiformis<br />
Pteris mutilata L.<br />
var. victoriae Hort.<br />
Pteris fauriei Hieron.,Hedwigia 55 (1914) 345; --Hatusima, Mem. Fac. Kagoshima Univ. 5 (1966)<br />
20.<br />
Pteris rigidula Copel., FFP 1 (1958) 137.<br />
Pteris glaucovirens Goldm.<br />
Pteris heteromorpha Fée<br />
Pteris loheri Copel.<br />
Pteris longipinnula Wall. ex J Agardh<br />
Pteris luzonensis Hieron.<br />
Pteris macgregorii Copel.
Pteris melanocaulon Fée<br />
Pteris melanorhachis Copel.<br />
Pteris mertensioides Willd.<br />
Pteris gardneri (Fée) Hook.<br />
Pteris excelsa Gaudich.<br />
Pteris micracantha Copel.<br />
Pteris mucronulata Copel.<br />
Pteris opaca (C Presl) J Sm. ex Fée<br />
Pteris taenitis Copel.<br />
Pteris oppositipinnata Fée<br />
Pteris asperula J Sm.<br />
Pteris caesia Copel.<br />
Pteris pachysora (Copel.) MG Price<br />
Schizostegeopsis pachysora Copel.<br />
Copelandiopteris pachysora (Copel.) Stone<br />
Pteris pedicellata Copel.<br />
Pteris pellucida C Presl<br />
Pteris philippinensis Fée<br />
Pteris longipes D Don, Prod. Fl. Nepal. (1825) 15; --Copel., FFP 1 (1958) 133<br />
Pteris longipes D Don var. philippinensis (Fée) Christ,<br />
Pteris purpureorhachis Copel.<br />
Pteris ramosii Copel.<br />
Possibly a variant <strong>of</strong> P. philippinensis.<br />
Pteris rangiferina C Presl<br />
Pteris ryukyuensis Tagawa, Acta Phytotax. Geobot. 4 (1935) 204; --Hatusima, Mem. Fac.<br />
Kagoshima Univ. 5 (1966) 20.<br />
?Pteris cretica sensu Copel., FFP 1 (1958) 129, quoad pl. ex Batanes Islands, non L.<br />
Pteris schlechteri Brause<br />
Pteris semipinnata L.<br />
Pteris spinescens C Presl<br />
Pteris setuloso-costulata Hayata<br />
Pteris squamipes Copel.<br />
Pteris tripartita Sw.<br />
Pteris microdictyon (Fée) Hook.<br />
Pteris vittata L.<br />
Pteris wallichiana J Agardh<br />
Confirmed to be in Luzon: Ilocos Norte prov., Mt. Burnay (Iwatsuki & Price, 1975).
Pteris whitfordii Copel.<br />
Schizoloma Gaudich. = Taenitis<br />
5. Syngramma J Smith<br />
Revision:<br />
Holttum, RE. 1974. Kew Bulletin 30: 327-343 + pl.<br />
Syngramma alismifolia (C Presl) J Sm., Lond. J. Bot. 4 (1845) 168, t. 7-8, Fig. B.<br />
Syngramma vittiformis J Sm.<br />
Syngramma wallichii (Hook.) Bedd.<br />
6. Taenitis Willdenow<br />
Taenitis blechnoides (Willd.) Sw., Syn. Fil. 24 (1806) 220;<br />
Taenitis luzonica (Alderw.) Holttum<br />
Syngramma luzonica Alderw.<br />
Taenitis cordata (Gaudich.) Holttum<br />
Schizoloma cordatum Gaudich.<br />
Family 27. ASPLENIACEAE (1 genus, 52 species)<br />
1. Asplenium Linnaeus<br />
Asplenium affine Sw., Schrad. J. Bot. 1800 (2) (1801) 56<br />
Asplenium spathulinum J Sm. ex Hook.<br />
Asplenium gracilifolium Copel.<br />
Asplenium amboinense Willd., Sp. Pl. 5 (1810) 303;<br />
Asplenium apoense Copel.<br />
Asplenium caudatum G Forst., Prodr. (1786) 80;<br />
?Asplenium chamissoanum C Presl<br />
Asplenium cheilosorum Kunze ex Mett., Aspl., 133, n. 104 (1859) t. 5, Fig. 12-13;<br />
Asplenium colubrinum Christ<br />
Asplenium crinicaule Hance<br />
Asplenium cuneatum Lam.<br />
Asplenium cymbifolium Christ, Bull. Boiss. 2, 6 (1906) 999;<br />
Asplenium elmeri Christ, PJS 2 c (1907) Bot. 164;
Asplenium epiphyticum Copel.<br />
Asplenium excisum C Presl<br />
Asplenium obscurum Blume<br />
Asplenium exiguum Bedd.<br />
Asplenium griseum Copel.<br />
Asplenium haenkei Copel.<br />
Asplenium insiticium Brackenr.<br />
Asplenium lepturus J Sm. ex C Presl<br />
Asplenium contiguum Kaulf.<br />
Asplenium longissimum Blume<br />
Asplenium acutiusculum Blume<br />
?Asplenium anisodontum C Presl<br />
Asplenium mantalingahanum PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32<br />
(1980) 46, t. 3.<br />
Asplenium militare Copel.<br />
Asplenium nidus L., Sp. Pl. (1753) 1079;<br />
var. nidus<br />
var. curtisorum (C Chr.) Holttum, Gard. Bull. Sing. 27 (1974) 148;<br />
Asplenium normale D Don, Prodr. Fl. Nepal. (1825) 7;<br />
Asplenium <strong>of</strong>eliae Salgado, Amer. Fern J. 93(4) (2003) 193. –Type: Merrill Phil. Plts. 700. (US,<br />
holo; iso: PRC). Luzon: Benguet prov., May 1911.<br />
Asplenium pellucidum Lam., Encycl. 2 (1786) 305;<br />
Asplenium persicifolium J Sm. ex Mett., Aspl. 97 (1859) no. 32<br />
Asplenium tripinnatifidum Copel.<br />
Asplenium phyllitidis D Don<br />
ssp. malesicum Holttum, Gard. Bull. Sing. 27 (1974) 153;<br />
Asplenium nidus L. var. phyllitidis (D. Don) Alderw., Handb. Malayan Ferns Suppl. (1917) 282;<br />
–Probably incl. A. colubrinum<br />
Asplenium polyodon G Forst., Prodr. (1786) 80;<br />
Asplenium falcatum Lam.<br />
Asplenium robustum Blume<br />
Asplenium laserpitiifolium Lam.<br />
Asplenium scandens J Sm. ex Mett.<br />
Asplenium schizocarpum Copel.<br />
Asplenium scolopendrioides J Sm. ex Hook., Icon. Pl. (1854) t. 930
Asplenium semivarians Viane & Reichst., Pteridol. New Millennium (2003) 78 (-81; fig. 3).<br />
78 (-81; fig. 3) 2003<br />
Tamil Nadu (India, Indian Subcontinent, Asia-Tropical);Yunnan (China South-Central,<br />
China, Asia-Temperate);West-Central Tropical Africa (Africa);Manipur (Assam, Indian<br />
Subcontinent, Asia-Tropical);<strong>Philippine</strong>s (Malesia, Asia-Tropical);Sri Lanka (Indian<br />
Subcontinent, Asia-Tropical);Uganda (East Tropical Africa, Africa);Zimbabwe (South<br />
Tropical Africa, Africa);<br />
Type Information<br />
Collector(s): Fraser-Jenkins 9063<br />
Locality: Shevaroy Hills, NW <strong>of</strong> Yercaud, NE <strong>of</strong> Salem, c. 1550m<br />
Collection Date: 1978-12-13<br />
Type Location: holotype GENT<br />
isotype Herb. R.Viane<br />
32. Asplenium setisectum Blume, Enum. Pl. Jav. (1828) 187;<br />
33. Asplenium stantoni Copel.<br />
34. Asplenium steerei Harrington<br />
35. Asplenium subnormale Copel. in Perkins, Fragm. Fl. Philip. (1905) 183, t. 4, Fig. B.<br />
36. Asplenium subresectum Copel.<br />
37. Asplenium tenerum G Forst., Prodr. (1786) 80;<br />
38. Asplenium tenuifolium Don<br />
39. Asplenium thunbergii Kunze, Linnaea 10 (1836) 517;<br />
Asplenium decorum Kunze<br />
49. Asplenium unilaterale Lam., Encycl. 2 (1786) 305;<br />
Asplenium filipes Copel., PJS 3 c (1908) Bot. 304;<br />
50. Asplenium vittiforme Cav., Descr. (1802) 255;<br />
51. Asplenium vulcanicum Blume, Enum. Pl. Jav. (1828) 176;<br />
Possibly conspecific with A. salignum Blume<br />
52. Asplenium yoshinagae Makino<br />
var. planicaule (Wall. ex Mett.) C Morton, Contr. US Nat’l. Herb. 38 (1973) 227;<br />
Asplenium planicaule Wall. ex Mett., Abhandl. Senckenb. Naturf. Gessell. 3 (1859) 201, non Lowe<br />
(1858).<br />
Family 28. THELYPTERIDACEAE (19 genera, 123 taxa: 118 species (1 possibly<br />
new) +4 varieties, + 1 possibly new forma).<br />
Revision: R.E. Holttum. Fl. Males. Ser. ii 1 (1981) 331-599<br />
1. Ampelopteris Kunze
1. Ampelopteris prolifera (Retz.) Copel.<br />
Tropical and subtropical regions <strong>of</strong> the Old World. In the <strong>Philippine</strong>s recorded from<br />
LUZON: Abra, Rizal, Laguna, MINDANAO: Davao del Norte (Maco, Lake Leonard),<br />
Agusan del Sur (Agusan Marsh). Terrestrial, <strong>of</strong>ten forming mats or tangles in<br />
swampy sites, lake margins or river embankments. Low altitudes.<br />
2. Amphineuron Holttum<br />
1. Amphineuron attenuatum (Kuntze) Holttum<br />
Cyclosorus alatellus sensu Copel. (1960), excl. syn.<br />
Lastrea erubescens sensu Copel. (1960)<br />
2. Amphineuron immersum (Blume) Holttum<br />
Lastrea immersa (Blume) T Moore<br />
Lastrea verrucosa C Presl<br />
3. Amphineuron opulentum (Kaulf.) Holttum<br />
Cyclosorus extensus (Blume) Ching<br />
4. Amphineuron terminans (Hook.) Holttum<br />
Cyclosorus interruptus sensu Copel. (1960), non (Willd.) H Ito<br />
Sri Lanka to Polynesia. Throughout the <strong>Philippine</strong>s in the more seasonal regions.<br />
Terrestrial, common in thickets on hillsides and in valleys at low and medium<br />
altitudes.<br />
1. Chingia christii (Copel.) Holttum<br />
Cyclosorus christii (Copel.) Copel.<br />
2. Chingia clavipilosa Holttum<br />
var. clavipilosa<br />
3. Chingia ferox (Blume) Holttum<br />
Cyclosorus ferox (Blume) Ching<br />
4. Chingia paucipaleata Holttum<br />
5. Chingia pricei Holttum<br />
6. Chingia urens Holttum<br />
1. Christella acuminata (Houtt.) Lév.<br />
Cyclosorus acuminatus (Houtt.) Ching<br />
2. Christella arida (D Don) Holttum<br />
Cyclosorus aridus (D Don) Ching<br />
3. Chingia Holttum<br />
4. Christella Léville emend. Holttum<br />
3. Christella dentata (Forssk.) Brownsey & Jermy<br />
Cyclosorus dentatus sensu Copel., 1960, pro parte<br />
Cyclosorus mindanaensis (Christ) Copel.<br />
Pantropic. Throughout the <strong>Philippine</strong>s. Terrestrial in exposed sites, light forests,<br />
along stream embankments, etc. Common at low altitudes.
4. Christella hispidula (Decne.) Holttum<br />
Cyclosorus falcatulus (Christ) Copel.<br />
5. Christella parasitica (L.) Lév.<br />
Cyclosorus dentatus sensu Copel. 1960, pro parte<br />
Cyclosorus benguetensis Copel.<br />
Cyclosorus procurrens (Mett.) Copel.<br />
SE Asia (confined here?, or more widely distributed?). Throughout the <strong>Philippine</strong>s.<br />
Terrestrial, open sites and degraded habitats generally, along stream embankments.<br />
Altitude?<br />
6. Christella subdentata Holttum<br />
7. Christella subpubescens (Blume) Holttum<br />
Cyclosorus subpubescens (Blume) Ching<br />
Cyclosorus latipinna (Hook. & Baker) Tard.<br />
1. Coryphopteris borealis Holttum<br />
2. Coryphopteris dura (Copel.) Holttum<br />
Lastrea dura (Copel.) Copel.<br />
5. Coryphopteris Holttum<br />
3. Coryphopteris pubirhachis (Baker) Holttum<br />
var. philippinensis Holttum<br />
Lastrea viscosa sensu Copel. (1960) non (Baker) Bedd., pro specim. philipp.<br />
4. Coryphopteris squamipes (Copel.) Holttum<br />
Lastrea squamipes (Copel.) Copel.<br />
Excluded:<br />
Coryphopteris viscosa (J Sm.) Holttum<br />
6. Cyclogramma Tagawa<br />
1. Cyclogramma auriculata (J Sm.) Ching<br />
Thelypteris simulans Ching<br />
7. Cyclosorus Link<br />
1. Cyclosorus interruptus (Willd.) H Ito<br />
Cyclosorus gongylodes auctt., doubtfully based on Aspidium gongylodes Schkuhr<br />
Haplodictyum C Presl =Pronephrium, Sphaerostephanos<br />
8. Macrothelypteris Ching<br />
1. Macrothelypteris polypodioides (Hook.) Holttum<br />
Lastrea leucolepis C Presl<br />
2. Macrothelypteris torresiana (Gaudich.) Ching<br />
Lastrea torresiana (Gaudich.) T Moore
9. Mesophlebion Holttum<br />
1. Mesophlebion crassifolium (Blume) Holttum<br />
Lastrea crassifolia (Blume) Moore<br />
2. Mesophlebion rufescens Holttum<br />
10. Metathelypteris Ching<br />
1. Metathelypteris dayi (Bedd.) Holttum<br />
Thelypteris singalanensis sensu Holttum, non (Baker) Ching<br />
2. Metathelypteris gracilescens (Blume) Ching<br />
Lastrea gracilescens (Blume) Moore<br />
Lastrea calva (Copel.) Copel.<br />
3. Metathelypteris uraiensis (Rosenst.) Ching<br />
11. Nannothelypteris Holttum<br />
1. Nannothelypteris aoristisora (Harrington) Holttum<br />
Cyclosorus aoristisorus (Harrington) Copel.<br />
2. Nannothelypteris camarinensis Holttum<br />
3. Nannothelypteris inaequilobata Holttum<br />
Lastrea nervosa sensu Copel. (1960), pro parte<br />
4. Nannothelypteris nervosa (Fée) Holttum<br />
Lastrea nervosa (Fée) Copel., pro parte<br />
5. Nannothelypteris philippina (C Presl) Holttum<br />
Lastrea confusa (Copel.) Copel.<br />
12. Parathelypteris (H Ito) Ching emend. Holttum<br />
1. Parathelypteris beddomei (Baker) Ching<br />
var. beddomei<br />
Lastrea beddomei (Baker) Bedd.<br />
var. eugracilis (Copel.) Holttum<br />
Lastrea eugracilis Copel.<br />
2. Parathelypteris glanduligera (Kunze) Ching<br />
3. Parathelypteris grammitoides (Christ) Holttum<br />
Lastrea grammitoides (Christ) Copel.<br />
13. Plesioneuron (Holttum) Holttum<br />
1. Plesioneuron savaiense (Baker) Holttum<br />
Lastrea quadriaurita (Christ) Copel.
14. Pneumatopteris Nakai<br />
1. Pneumatopteris costata (Brackenr.) Holttum<br />
Lastrea cavitensis Copel.<br />
2. Pneumatopteris glabra (Copel.) Holttum<br />
Cyclosorus glaber Copel.<br />
Cyclosorus viridis Copel.<br />
3. Pneumatopteris laevis (Mett.) Holttum<br />
Cyclosorus nitidulus sensu Copel. (1960), pro parte<br />
Cyclosorus microlonchus sensu Copel. (1960), pro parte<br />
4. Pneumatopteris ligulata (C Presl) Holttum<br />
Lastrea ligulata C Presl<br />
Lastrea foxii (Christ) Copel.<br />
5. Pneumatopteris lithophila Holttum<br />
6. Pneumatopteris michaelis Holttum<br />
7. Pneumatopteris microloncha (Christ) Holttum<br />
Cyclosorus microlonchus (Christ) Copel.<br />
8. Pneumatopteris nitidula (C Presl) Holttum<br />
Cyclosorus nitidulus (C Presl) Copel. (1960), p.p<br />
9. Pneumatopteris truncata (Poir.) Holttum<br />
Cyclosorus truncatus (Poir.) Farw.<br />
1. Pronephrium amphitrichum Holttum<br />
15. Pronephrium C Presl<br />
2. Pronephrium asperum (C Presl) Holttum<br />
Cyclosorus philippinarum (Fée) Copel.<br />
3. Pronephrium bakeri (Harrington) Holttum<br />
Haplodictyum bakeri (Harrington) Ching<br />
Cyclosorus bakeri (Harrington) Copel.<br />
4. Pronephrium balabacensis PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32<br />
(1980) 45, t. 2.<br />
Possibly synonymous with P. merrilliii. The exindusiate condition <strong>of</strong> the sori<br />
mentioned in the original description may actually be due to small, fugaceous<br />
indusia.<br />
5. Pronephrium bulusanicum (Holttum) Holttum<br />
Haplodictyum bulusanicum Holttum<br />
6. Pronephrium clemensiae (Copel.) Holttum<br />
var. clemensiae<br />
Cyclosorus clemensiae (Copel.) Ching<br />
var. degenerum (Christ) Holttum<br />
Cyclosorus degener (Christ) Copel.<br />
7. Pronephrium cuspidatum (Blume) Holttum<br />
Cyclosorus cuspidatus (Blume) Copel.
8. Pronephrium granulosum (C Presl) Holttum<br />
Cyclosorus granulosus (C Presl) Copel.<br />
Cyclosorus subdimorphus Copel.<br />
9. Pronephrium gymnopteridifrons (Hayata) Holttum<br />
Cyclosorus pustulosus Copel.<br />
10. Pronephrium heterophyllum (C Presl) Holttum<br />
Haplodictyum heterophyllum C Presl<br />
11. Pronephrium hosei (Baker) Holttum<br />
Cyclosorus hosei (Baker) Copel.<br />
12. Pronephrium lineatum (Blume) Presl<br />
13. Pronephrium melanophlebium (Copel.) Holttum<br />
forma melanophlebium<br />
Cyclosorus melanophlebius (Copel.) Copel.<br />
forma proliferum Co, ined.<br />
14. Pronephrium menisciicarpon (Blume) Holttum<br />
Cyclosorus latifolius (C Presl) Copel.<br />
15. Pronephrium merrillii (Christ) Holttum<br />
Cyclosorus merrillii (Christ) Copel.<br />
16. Pronephrium nitidum Holttum<br />
Cyclosorus urophyllus sensu Copel. (1960), excl. loc. Malacca<br />
17. Pronephrium ramosii (Christ) Holttum<br />
var. ramosii<br />
Cyclosorus cuspidatus sensu Copel. (1960), excl. syn. Meniscium cuspidatum Blume<br />
18. Pronephrium rhombeum (Christ) Holttum<br />
Cyclosorus rhombeus (Christ) Copel.<br />
Cyclosorus lanceolus (Christ) Copel.<br />
19. Pronephrium rubidum (J Sm. ex Hook.) Holttum<br />
Cyclosorus rubidus (Hook.) Copel.<br />
20. Pronephrium samarense (Copel.) Holttum<br />
Cyclosorus samarensis Copel.<br />
21. Pronephrium solsonicum Holttum<br />
22. Pronephrium trachyphyllum Holttum<br />
23. Pronephrium triphyllum (Sw.) Holttum<br />
Cyclosorus triphyllus (Sw.) Tard.<br />
24. Pronephrium x xiphioides (Christ) Holttum<br />
Cyclosorus xiphioides (Christ) Copel.<br />
Cyclosorus edanyoi Copel.<br />
Cyclosorus rhombeus sensu Copel. (1960), pro parte<br />
16. Pseudocyclosorus Ching<br />
1. Pseudocyclosorus tylodes (Kunze) Moore
Lastrea tylodes (Kunze) Moore; ‘xylodes’ in Copeland’s Fern <strong>Flora</strong>.<br />
17. Pseudophegopteris Ching<br />
1. Pseudophegopteris aurita (Hook.) Ching<br />
2. Pseudophegopteris paludosa (Blume) Ching<br />
Lastrea pyrrhorhachis sensu Copel., Fern Fl. Philipp. (1960) 330, pro parte<br />
18. Sphaerostephanos J Sm.<br />
Revision:<br />
Holltum, RE. 1975. Studies in the family Thelypteridaceae. IX. The genus Sphaerostephanos<br />
in the <strong>Philippine</strong>s. Kalikasan 4: 47-68.<br />
1. Sphaerostephanos alcasidii Holttum<br />
2. Sphaerostephanos acrostichoides (Desv.) Holttum<br />
Sphaerostephanos smithianus (C Presl) Holttum<br />
Cyclosorus boholensis Copel.<br />
Cyclosorus productus sensu Copel. (1960), pro parte<br />
3. Sphaerostephanos angustifolius (C Presl) Holttum<br />
Cyclosorus angustifolius (C Presl) Copel.<br />
4. Sphaerostephanos appendiculatus (Blume) Holttum<br />
5. Sphaerostephanos cartilagidens PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp.<br />
32 (1980) 43, t. 1.<br />
6. Sphaerostephanos convergens Holttum<br />
7. Sphaerostephanos dichrotrichoides (Alderw.) Holttum<br />
Lastrea dichrotrichoides (Alderw.) Copel.<br />
8. Sphaerostephanos diminutus (Copel.) MG Price<br />
Cyclosorus diminutus (Copel.) Copel.<br />
Pronephrium diminutum (Copel.) Holttum<br />
Sphaerostephanos lucbanii Holttum<br />
9. Sphaerostephanos diversilobus (C Presl) Holttum<br />
Cyclosorus diversilobus (C Presl) Ching<br />
10. Sphaerostephanos ellipticus (Rosenst.) Holttum<br />
var. ellipticus<br />
Cyclosorus ellipticus (Rosenst.) Copel.<br />
var. glabrior Holttum<br />
11. Sphaerostephanos erectus (Copel.) Holttum<br />
Cyclosorus erectus Copel.<br />
12. Sphaerostephanos foxworthyi Holttum<br />
13. Sphaerostephanos grandescens Holttum<br />
14. Sphaerostephanos hernaezii Holttum<br />
15. Sphaerostephanos heterocarpus (Blume) Holttum<br />
Eight variants <strong>of</strong> this polymorphic species were enumerated (but no formal names
given) by Holttum in the his FM revision. Two are definitely occurring in the<br />
<strong>Philippine</strong>s, while another 2 resembles those ascribed to Borneo and New Guinea<br />
forms. Thus:<br />
-Mindanao form<br />
-Luzon form [as Cyclosorus bordenii (Christ) Ching in ‘Fern <strong>Flora</strong>’].<br />
From Palawan is one collection resembling the Borneo form, while another one from<br />
San Ramon, Zamboanga is nearer to the New Guinea form.<br />
16. Sphaerostephanos hirsutus (Kunze ex Mett.) Holttum<br />
var. hirsutus<br />
Cyclosorus adenophorus (C Chr.) Ching<br />
17. Sphaerostephanos irayensis (Copel.) Holttum<br />
Plesioneuron irayense (Copel.) Holttum<br />
Cyclosorus irayensis Copel.<br />
Sphaerostephanos fenixii Holttum<br />
18. Sphaerostephanos lastreoides (C Presl) Holttum<br />
Cyclosorus duplosetosus Copel.<br />
19. Sphaerostephanos latebrosus (Kunze ex Mett.) Holttum<br />
Sphaerostephanos todayensis (Christ) Holttum<br />
Cyclosorus todayensis (Christ) Copel.<br />
Cyclosorus sagittifolioides Copel.<br />
Cyclosorus halconensis Copel.<br />
Cyclosorus polypterus Copel.<br />
20. Sphaerostephanos lobatus (Copel.) Holttum<br />
Cyclosorus lobatus Copel.<br />
Cyclosorus bordenii sensu Copel. (1960), pro parte<br />
21. Sphaerostephanos loherianus (Christ) Holttum<br />
Cyclosorus loherianus (Christ) Copel.<br />
22. Sphaerostephanos magnus (Copel.) Holttum<br />
Cyclosorus magnus Copel.<br />
23. Sphaerostephanos major (Copel.) Holttum<br />
Haplodictyum majus Copel.<br />
Cyclosorus dimorphus Copel.<br />
24. Sphaerostephanos norrisii (Rosenst.) Holttum<br />
Sphaerostephanos elmerorum (Copel.) Holttum<br />
Cyclosorus elmerorum Copel.<br />
Cyclosorus ellipticus sensu Copel. (1960), pro parte<br />
25. Sphaerostephanos perglanduliferus (Alderw.) Holttum<br />
26. Sphaerostephanos polisianus Holttum<br />
27. Sphaerostephanos polycarpus (Blume) Copel.<br />
Sphaerostephanos asplenioides J Sm.<br />
28. Sphaerostephanos porphyricola (Copel.) Holttum<br />
Cyclosorus pophyricola (Copel.) Ching<br />
29. Sphaerostephanos productus (Kaulf.) Holttum<br />
Cyclosorus productus (Kaulf.) Ching<br />
Cyclosorus serratus Copel.<br />
Cyclosorus weberi Copel.<br />
Cyclosorus rigidus Copel.
Cyclosorus leucadenius Copel.<br />
Sphaerostephanos punctatus Holttum<br />
Cyclosorus megaphyllus sensu Copel. (1960)<br />
Cyclosorus nitidulus sensu Copel. (1960), pro parte<br />
30. Sphaerostephanos santomasii Holttum<br />
31. Sphaerostephanos sessilipinna (Copel.) Holttum<br />
Cyclosorus sessilipinna (Copel.) Ching<br />
Cyclosorus austrophilippinus (Copel.) Copel.<br />
32. Sphaerostephanos simplicifolius (J Sm. ex Hook.) Holttum<br />
Cyclosorus simplicifolius (J Sm.) Copel.<br />
33. Sphaerostephanos spenceri (Christ) Holttum<br />
Cyclosorus spenceri (Christ) Copel.<br />
34. Sphaerostephanos stenodontus (Copel.) Holttum<br />
Cyclosorus stenodontus Copel.<br />
35. Sphaerostephanos subcordatus Holttum<br />
36. Sphaerostephanos tephrophyllus (Copel.) Holttum<br />
Cyclosorus tephrophyllus (Copel.)<br />
37. Sphaerostephanos unitus (L.) Holttum<br />
var. unitus<br />
Cyclosorus unitus (L.) Ching<br />
Sri Lanka, S India, across Malesia. Throughout the <strong>Philippine</strong>s. Terrestrial. Degraded<br />
habitats generally, along waterways, etc. at low to medium altitudes. Very common.<br />
var. mucronatus (Christ) Holttum<br />
38. Sphaerostephanos urdanetensis (Copel.) Holttum<br />
Cyclosorus urdanetensis Copel.<br />
Cyclosorus matutumensis Copel.<br />
39. Sphaerostephanos williamsii (Copel.) Holttum<br />
Lastrea williamsii Copel.<br />
Status?:<br />
Sphaerostephanos mindorensis Holltum, Kalikasan 4 (1975) 54. Type: MG Price 690 (K,<br />
holotype; isotype in PNH). Mindoro: Mt Halcon.<br />
MINDORO: Mt Halcon, in forest, 750m. Known from two collections from the type<br />
locality. Not accounted for in Holltum’s <strong>Flora</strong> Malesiana revision.<br />
19. Stegnogramma Blume<br />
1. Stegnogramma calcicola Co, sp. nov., ined.<br />
BOHOL.<br />
2. Stegnogramma dissitifolia Holttum<br />
3. Stegnogramma gymnocarpa (Copel.) K Iwats.<br />
ssp. gymnocarpa<br />
Lastrea gymnocarpa Copel.
1. Athyrium anisopterum Christ<br />
2. Athyrium aristulatum Copel.<br />
3. Athyrium brevipinnulum Copel.<br />
Athyrium meansianum (Copel.) Alderw.<br />
Athyrium philippinense Christ apud Copel.<br />
Family 29. WOODSIACEAE (incl. Athyriaceae)<br />
(6 genera, xx species)<br />
Anisocampium Presl =Athyrium<br />
1. Athyrium Roth<br />
4. Athyrium cumingianum (C Presl) Ching<br />
Anisocampium cumingianum C Presl<br />
5. Athyrium drepanopteron (Kunze) A. Braun<br />
6. Athyrium oreopteris Copel.<br />
7. Athyrium puncticaule (Blume) Moore<br />
8. Athyrium ramosii Copel.<br />
9. Athyrium stramineum Copel.<br />
Callipteris Bory =Diplazium<br />
2. Cornopteris Nakai<br />
1. Cornopteris banahaoensis (C Chr.) K Iwats. & MG Price<br />
Athyrium nudum Copel.<br />
2. Cornopteris<br />
Athyrium elmeri Copel.<br />
3. Cornopteris<br />
Athyrium gymnocarpum Copel.<br />
Currania Copeland =Gymnocarpium<br />
3. Cystopteris Bernhardi<br />
1. Cystopteris tenuisecta (Blume) Mett., Ann. Lugd. Bat. 1 (1864) 241<br />
Acystopteris tenuisecta (Blume) Tagawa<br />
Ctenitis tenerifrons (Christ) Copel.<br />
4. Deparia Hooker & Greville<br />
1. Deparia boryana (Willd.) Kato, Bot. Mag. Tokyo 90 (1977) 36.
Dryoathyrium boryanum (Willd.) Ching<br />
Ctenitis boryana (Willd.) Copel.<br />
2. Deparia confluens (Kunze) Kato<br />
3. Deparia formosana (Rosenst.) Sano, Act. Phytotax. Geobot. 51 (2000) 17.<br />
Diplaziopsis formosana (Rosenst.) MG Price, Contrib. Univ. Mich. Herb. 17 (1990) 269; -- Diplazium<br />
formosanum Rosenst., Hedw. 56 (1915) 337<br />
3. Deparia japonica (Thunb.) Kato, Bot. Mag. Tokyo 90 (1977) 37.<br />
Lunathyrium japonicum (Thunb.) Kurata<br />
Athyrium japonicum (Thunb.) Ching<br />
Athyrium grammitoides (C Presl) Milde<br />
4. Deparia subfluviatilis (Hayata) Kato<br />
`<br />
1. Diplazium allantoideum MG Price<br />
2. Diplazium atratum Christ<br />
Athyrium atratum (Christ) Copel.<br />
3. Diplazium bantamense Blume<br />
Athyrium bantamense (Blume) Milde<br />
4. Diplazium bolsteri Copel.<br />
Athyrium bolsteri (Copel.) Copel.<br />
5. Diplazium brachysoroides Copel.<br />
Athyrium brachysoroides (Copel.) Copel.<br />
Diplaziopsis C Christensen =Diplazium<br />
5. Diplazium Swartz<br />
6. Diplazium calliphyllum (Copel.) MG Price, Contrib. Univ. Mich. Herb. 17 (1990) 270.<br />
Athyrium calliphyllum Copel.<br />
7. Diplazium cordifolium Blume<br />
Athyrium cordifolium (Blume) Copel.<br />
Athyrium parvum Copel.<br />
8. Diplazium costulisorum (Copel.) C Chr.<br />
Athyrium costulisorum Copel.<br />
9. Diplazium cultratum C Presl<br />
Athyrium cultratum (C Presl) Milde<br />
10. Diplazium cumingii (C Presl) C Chr.<br />
Athyrium cumingii (C Presl) Milde<br />
11. Diplazium davaoense Copel.<br />
Athyrium davaoense (Copel.) Copel.<br />
12. Diplazium deltoideum (C Presl) C Presl<br />
Athyrium deltoideum (C Presl) Milde<br />
13. Diplazium dilatatum Blume<br />
Athyrium dilatatum (Blume) Milde
14. Diplazium doederleinii (Luerssen) Makino<br />
Athyrium irayense Copel.<br />
15. Diplazium dolichosorum Copel.<br />
Athyrium dolichosorum (Copel.) Copel.<br />
16. Diplazium egenolfioides MG Price, Brit. Fern Gaz. 10 (1973) 258, t. 1<br />
LUZON: Mt. Makiling (several collections from this type locality); Nueva Vizcaya<br />
prov., Sta. Fe, mtns N <strong>of</strong> Imugan, c. 1200m, L.L.Co 1640 (MICH, PNH, PUH).<br />
17. Diplazium esculentum (Retz.) Sw.<br />
Athyrium esculentum (Retz.) Copel.<br />
Asplenium manilense Sprengel<br />
18. Diplazium fraxinifolium C Presl<br />
Athyrium fraxinifolium (C Presl) Milde<br />
19. Diplazium fructuosum Copel.<br />
Athyrium fructuosum (Copel.) Copel.<br />
20. Diplazium fuliginosum (Hook.) MG Price, Brit. Fern Gaz. 10 (1973) 260.<br />
Asplenium fuliginosum Hook.<br />
Diplazium longissimum (Copel.) C Chr.<br />
Athyrium longissimum Copel.<br />
21. Diplazium geophilum (Copel.) Alderw.<br />
Athyrium geophilum Copel.<br />
22. Diplazium griffithii T Moore<br />
Athyrium platyphyllum Copel., 1908, non Diplazium platyphyllum Christ, 1906<br />
Diplazium mearnsii Hieron.<br />
Athyrium platyphyllum Copel.<br />
23. Diplazium irigense (Copel.) MG Price, Contrib. Univ. Mich. Herb. 17 (1990) 273<br />
Athyrium irigense Copel.<br />
24. Diplazium javanicum (Blume) Makino, Phan. Pter. Jap. Ic. I11. 1(1901) pl. 46<br />
Diplaziopsis javanica (Blume) C Chr.<br />
25. Diplazium lanceum (Thunb.) C Presl<br />
Diplazium subsinuatum (Wall. ex Hook. & Grev.) Tagawa<br />
Athyrium dubium (Don) Copel.<br />
26. Diplazium lomariaceum (Christ) MG Price, Gard. Bull. Sing. 36 (1983) 27<br />
Diplazium altum (Copel.) C Chr.<br />
Athyrium altum Copel.<br />
Diplazium merrillii Copel.<br />
Athyrium merrillii (Copel.) Copel.<br />
27. Diplazium macrosorum (Copel.) MG Price, Contrib. Herb. Univ. Mich. 17 (1990) 273<br />
Athyrium macrosorum Copel.<br />
28. Diplazium magnificum (Copel.) MG Price, 1982<br />
Athyrium magnificum Copel.<br />
29. Diplazium melanopodium Fée<br />
Athyrium melanopodium (Fée) Copel.<br />
30. Diplazium mollifrons (C Chr.) MG Price
31. Diplazium muricatum (Mett.) Alderw.<br />
Athyrium muricatum (Mett.) Copel.<br />
32. Diplazium oligosorum Copel.<br />
Athyrium oligosorum (Copel.) Copel.<br />
33. Diplazium pallidum (Blume) Moore<br />
Athyrium pallidum Blume<br />
34. Diplazium pariens (Copel.) C Chr.<br />
Athyrium pariens Copel.<br />
35. Diplazium petiolare C Presl<br />
Athyrium petiolare (C Presl) Milde<br />
36. Diplazium pinatubicum MG Price, Contrib. Herb. Univ. Michigan 17 (1990) 273<br />
37. Diplazium polypodioides Blume<br />
Athyrium blumei (Bergsmann) Copel.<br />
Diplazium ophiodontum (Copel.) C Chr.<br />
Athyrium ophiodontum Copel.<br />
Diplazium asperum Blume<br />
Athyrium asperum (Blume) Milde<br />
38. Diplazium proliferum (Lam.) Thouars<br />
Callipteris prolifera (Lam.) Bory<br />
Diplazium accedens Blume<br />
39. Diplazium propinquum (Copel.) Alderw.<br />
Athyrium propinquum Copel.<br />
40. Diplazium pseudocyatheifolium Rosenst.<br />
Athyrium pseudocyatheifolium (Rosenst.) Copel.<br />
Athyrium ebenirachis Copel.<br />
41. Diplazium sibuyanense (Copel.) Alderw.<br />
Athyrium sibuyanense Copel.<br />
42. Diplazium sorzogonense (C Presl) C Presl<br />
Athyrium sorzogonense (C Presl) Milde<br />
43. Diplazium symmetricum (Copel.) MG Price, Gard. Bull. Sing. 36 (1983) 26.<br />
Athyrium symmetricum Copel.<br />
SAMAR: Mt. Cabayanan, Edano PNH 15148, lectotype (MICH).<br />
44. Diplazium tenuifolium (Copel.) Lellinger, Am. Fern J. 74 (1954) 57<br />
Athyrium tenuifolium Copel.<br />
Formally published new combination?<br />
45. Diplazium tomentosum Blume<br />
Athyrium crenato-serratum (Blume) Milde<br />
46. Diplazium vestitum C Presl<br />
Athyrium vestitum (C Presl) Milde<br />
47. Diplazium whitfordii Copel.<br />
Athyrium whitfordii (Copel.) Copel.<br />
48. Diplazium williamsii Copel.<br />
Athyrium williamsii (Copel.) Copel.
Athyrium banahaoense Copel.<br />
Diplazium banahaoense (Copel.) C Chr.<br />
49. Diplazium woodwardioides (C Presl) Holttum<br />
Athyrium sylvaticum (Blume) Milde<br />
50. Diplazium xiphophyllum (Baker) Copel.<br />
Athyrium xiphophyllum (Baker) Copel.<br />
Diplazium accedens Blume<br />
Polillo, PNBG 184 Sotalbo (PUH). New record for the <strong>Philippine</strong>s?<br />
Most specimens in D. proliferum; the rest in D. vestitum & D. dolichosorum<br />
Dryoathyrium Ching =Deparia<br />
6. Gymnocarpium Newman<br />
1. Gymnocarpium oyamense (Baker) Ching<br />
Currania gracilipes Copel.<br />
Conspectus <strong>of</strong> the genera:<br />
Lunathyrium Koidzumi =Deparia<br />
Family 30. BLECHNACEAE<br />
Subfamily I. Blechnoideae [scales not peltate, petiole vascular bundles U-shaped in<br />
cross section]:<br />
Blechnum L.<br />
Brainea J SM.<br />
Woodwardia SM.<br />
Subfamily II. Stenochlaenoideae [scales peltate, petiole vascular bundles in 2<br />
circles]:<br />
Stenochlaena J SM.<br />
1. Blechnum Linnaeus<br />
1. Blechnum egregium Copel. in Perkins, Fragm. Fl. Philip. (1905) 187;<br />
2. Blechnum finlaysonianum Hook. & Grev., Icon. Fil., t. (1831) 225;<br />
3. Blechnum fraseri (A. Cunningham) Luerss., <strong>Flora</strong> (1876) 292;<br />
4. Blechnum orientale L., Sp. Pl. (1753) 1077;<br />
5. Blechnum patersonii (R Br.) Mett., Fil. Lips. (1856) 64, t. 4, Fig. 4-10;<br />
6. Blechnum vestitum (Blume) Kuhn, Ann. Lugd.-Bat. 4 (1869) 284;<br />
Blechnum capense (L.) Schlechtr.<br />
7. Blechnum vulcanicum (Blume) Kuhn, Ann. Lugd.-Bat. 4 (1869) 284;
1. Brainea insignis (Hook.) J Sm.<br />
2. Brainea J Smith<br />
1. Stenochlaena J Smith<br />
1. Stenochlaena areolaris (Harrington) Copel.<br />
2. Stenochlaena cumingii Holttum<br />
Stenochlaena laurifolia C Presl<br />
3. Stenochlaena milnei Underw.<br />
Stenochlaena juglandifolia C Presl<br />
4. Stenochlaena palustris (NL Burm.) Bedd., Ferns Brit. Ind. Suppl. (1876) 26;<br />
1. Woodwardia orientalis Sw.<br />
var. prolifera (Hook. & Arn.) Hook.<br />
4. Woodwardia Smith<br />
2. Woodwardia unigemmata (Makino) Nakai<br />
Family 31. ONOCLEACEAE (No <strong>Philippine</strong> representatives)<br />
Family 32. DRYOPTERIDACEAE (including “Aspidiaceae”, Bolbitidaceae,<br />
Elaphoglossaceae, Hypodematiaceae, Peranemataceae)<br />
(xx genera, xxx species)<br />
Revisions:<br />
Holttum, RE. 1978. <strong>Flora</strong> Malesiana series 2, 1: 255-330 (Bolbitis treatment by E.<br />
Hennipman)<br />
Tribe 2. Dryopterideae –7 genera, 44 species in the <strong>Philippine</strong>s.<br />
Acrophorus C Presl -1<br />
Arachniodes Blume – 5<br />
Cyrtomium C Presl -2<br />
Didymochlaena Desv. -1<br />
Dryopteris Adans. -14<br />
Dryopsis Holttum & Edwards -1<br />
Hypodematium Kunze<br />
Lastreopsis Ching -1<br />
Leucostegia<br />
Lomagramma<br />
Nothoperanema (Tagawa) Ching -1<br />
Peranema D Don -2<br />
Polystichum Roth -11<br />
1. Acrophorus C Presl
1. Acrophorus nodosus C Presl<br />
Acrophorus stipellatus (Wall.) T Moore<br />
1. Arachniodes amabilis (Blume) Tindale<br />
Rumohra amabilis (Blume) Ching<br />
2. Arachniodes aristata (Forster) Tindale<br />
Rumohra aristata (Forst.) Ching<br />
Acystopteris Nakai =Cystopteris<br />
2. Arachniodes Blume<br />
3. Arachniodes coniifolia (Wall. ex T Moore) Ching<br />
Rumohra wallichii Ching<br />
4. Arachniodes hasseltii (Blume) Ching<br />
Rumohra hasseltii (Blume) Ching<br />
5. Arachniodes tripinnata (Goldm.) Sledge<br />
Rumohra carvifolia (Kunze) Ching<br />
Arthrobotrya J Smith =Teratophyllum<br />
3. Bolbitis Schott<br />
Revision:<br />
E. Hennipman, Leid. Bot. Ser. 2 (1977) 123-<br />
1. Bolbitis angustipinna (Hayata) H Ito<br />
2. Bolbitis appendiculata (Willd.) K Iwats.<br />
ssp. appendiculata<br />
Egenolfia appendiculata (Willd.) J Sm.<br />
3. Bolbitis heteroclita (C Presl) Ching<br />
Bolbitis cuspidata (C Presl) Ching<br />
Bolbitis pseudoscalpturata (Copel.) Ching<br />
Bolbitis foxworthyi (Copel.) Ching<br />
Bolbitis tenuissima (Copel.) Copel.<br />
Bolbitis membranacea (Copel.) C Chr.<br />
Bolbitis edanyoi Copel.<br />
Edanyoa difformis Copel.<br />
4. Bolbitis quoyana (Gaudich.) Ching<br />
Bolbitis enormis (Copel.) C Chr.<br />
Stenosemia dimorpha Copel.<br />
5. Bolbitis repanda (Blume) Schott<br />
6. Bolbitis rhizophylla (Kaulf.) Hennipman<br />
Egenolfia intermedia (J Sm.) Fée<br />
Egenolfia rhizophylla (Kaulf.) Fée<br />
Egenolfia fluviatilis Copel.<br />
7. Bolbitis scalpturata (Fée) Ching
8. Bolbitis sinuata (C Presl) Hennipman<br />
Bolbitis diversifolia (Blume) Schott<br />
Bolbitis subsimplex (Fée) Ching<br />
Bolbitis hydrophylla (Copel.) Ching<br />
Hybrids:<br />
H. 1. Bolbitis x arguta (Fée) Ching<br />
Hennipman doubts whether this fern warrants a separate treatment as<br />
a hybrid; it may be close to Edanyoa difformis (=B. heteroclita).<br />
H. 2. Bolbitis heteroclita x rhizophylla Hennipman, Leid. Bot. Ser. 2 (1977) 291, Fig. 831o<br />
H. 3. Bolbitis x sinuosa (Fée) Copel., two nothomorphs distinguished:<br />
nm. sinuosa<br />
nm. foxii (Copel.) Hennipman<br />
Bolbitis x foxii Copel.<br />
Dubious species:<br />
Leptochilus stolonifer Christ, Bull. Herb. Boiss. Ii, 6 (1906) 1004.<br />
Luzon: Rizal prov., Angilog (type locality). Possibly synonymous with B. heteroclita<br />
but the venation pattern as given by Christ is deviating. Type not traced (Hennipman,<br />
Fl. Males. Ser ii 1 (1978) 330).<br />
Excluded:<br />
Bolbitis presliana (Fée) Ching<br />
Included in Copeland’s treatment in Fern Fl. Philipp. 2 (1960) 260. This taxon<br />
does not occur in the <strong>Philippine</strong>s nor anywhere in Malesia.<br />
4. Cyrtomium C Presl<br />
1. Cyrtomium cf. caryotideum (Wall.) Presl<br />
Phanerophlebia caryotidea (Wall. ex Hook. & Grev.) Copel.<br />
<strong>Philippine</strong> record based on a sterile specimen by Mearns from Mt. Sto. Tomas<br />
[Copeland, Fern Fl. Philipp. 2 (1960) 248].<br />
2. Cyrtomium cf. falcatum (L.f.) C Presl<br />
Phanerophlebia falcata (L. f.) Copel.<br />
<strong>Philippine</strong> record based on a Loher collection from Mt. Data [Copeland, Fern Fl.<br />
Philipp. 2 (1960) 248] at the turn <strong>of</strong> the 1900s.<br />
Diacalpe Blume =Peranema<br />
5. Didymochlaena Desvaux<br />
1. Didymochlaena truncatula (Sw.) J Sm.<br />
6. Dryopsis Holttum & PJ Edwards<br />
1. Dryopsis manipurensis (Bedd.) Holttum & PJ Edwards, Kew Bull. 41 (1986) 200.<br />
Ctenitis copelandii (Christ) Copel.
Ctenitis manipurensis (Bedd.) Ching<br />
1. Dryopteris cacaina Tagawa<br />
2. Dryopteris cochleata (Don) C Chr.<br />
3. Dryopteris chrysocoma C Chr.<br />
4. Dryopteris formosana (Christ) C Chr.<br />
5. Dryopteris hirtipes (Blume) Kuntze<br />
6. Dryopteris permagna MG Price<br />
7. Dryopteris polita Rosenst.<br />
7. Dryopteris Adanson<br />
8. Dryopteris pulvinulifera (Bedd.) Kuntze<br />
9. Dryopteris purpurascens (Blume) Christ<br />
10. Dryopteris sparsa (Don) Kuntze<br />
11. Dryopteris subarborea (Baker) C Chr.<br />
12. Dryopteris uropinna MG Price<br />
Dryopteris erythrosora sensu Copel. (1960), non (Eaton) Kuntze<br />
13. Dryopteris varia (L.) Kuntze<br />
14. Dryopteris wallichiana (Sprengel) Alston & Bonner<br />
Dryopteris paleacea (Sw.) C Chr., nom. illeg.<br />
Edanyoa Copeland = Bolbitis heteroclita<br />
Egenolfia Schott = Bolbitis<br />
8. Elaphoglossum Schott ex J Smith<br />
1. Elaphoglossum angulatum (Blume) T Moore<br />
2. Elaphoglossum apoense Holttum<br />
3. Elaphoglossum blumeanum (Fée) J Sm.<br />
var. blumeanum<br />
Elaphoglossum copelandii Christ<br />
var. philippinense Christ ex Holttum<br />
4. Elaphoglossum calanasanicum Holttum
5. Elaphoglossum callifolium (Blume) T Moore<br />
6. Elaphoglossum luzonicum Copel.<br />
Elaphoglossum elmeri Copel.<br />
7. Elaphoglossum melanostictum (Blume) T Moore<br />
Elaphoglossum basilanicum Copel.<br />
8. Elaphoglossum negrosensis Holttum<br />
Elaphoglossum angulatum sensu Copel. (1960), pro parte, non (Blume) T Moore<br />
9. Elaphoglossum ophioglossoides (Goldm.) Holttum<br />
Elaphoglossum merrillii Christ<br />
Hemigramma Christ =Tectaria<br />
9. Hypodematium Kunze<br />
1. Hypodematium crenatum (Forssk.) Kuhn<br />
1. Lastreopsis tenera (R Br.) Tindale<br />
Ctenitis recedens (J Sm.) Copel.<br />
10. Lastreopsis Ching<br />
11. Leucostegia C Presl<br />
1. Leucostegia immersa (Wall.) C Presl, Tent. (1836) 95, t. 4, Fig. 11;<br />
2. Leucostegia pallida Mett.<br />
Lomagramma brooksii Copel.<br />
Lomagramma copelandii Holttum<br />
Lomagramma merrillii Holttum<br />
Lomagramma pteroides J Sm.<br />
Lomagramma subcoriacea Copel.<br />
Lomagramma pedicellata Copel.<br />
Lomagramma cordata Copel.<br />
12. Lomagramma J Smith<br />
13. Nothoperanema (Tagawa) Ching<br />
Nothoperanema hendersonii (Bedd.) Ching<br />
Ctenitis mearnsii Copel.<br />
As Ct. mearnsii:<br />
LUZON: Benguet (Pauai; Mt Pulag). Terrestrial in mossy forest, altitude 2100-2750m.
Peranema aspidioides (Blume) Mett.<br />
Diacalpe aspidioides Blume<br />
14. Peranema D Don<br />
Peranema cyatheoides D Don<br />
var. luzonicum (Copel.) Ching & SH Wu, Acta Phytotax. Sin. 21 (1983) 373.<br />
Peranema luzonica Copel.<br />
Phanerophlebia C Presl =Cyrtomium<br />
15. Polystichum Roth<br />
Polystichum biaristatum (Blume) T Moore<br />
Polystichum aquilinum Copel.<br />
Polystichum copelandii Christ<br />
Polystichum deltodon (Baker) Diels<br />
Polystichum doianum Tagawa<br />
Polystichum pulogense Copel.<br />
Polystichum elmeri Copel.<br />
Polystichum fuscum Copel.<br />
Polystichum horizontale C Presl<br />
Polystichum nepalense (Sprengel) C Chr.<br />
Polystichum nudum Copel.<br />
Polystichum obliquum (D Don) T Moore<br />
Polystichum obtusum J Sm. ex C Presl<br />
Polystichum edanoi Copel.<br />
Rumohra sensu Copel. (1960) non Raddi =Arachniodes<br />
16. Teratophyllum Mettenius<br />
Teratophyllum aculeatum (Blume) Mett. ex Kuhn<br />
var. aculeatum<br />
Teratophyllum arthropteroides (Christ) Holttum<br />
Teratophyllum articulatum (J Sm. ex Fée) Mett. ex Kuhn<br />
Arthrobotrya articulata J Sm.<br />
Teratophyllum koordersii Holttum<br />
Teratophyllum leptocarpum (Fée) Holttum
Teratophyllum luzonicum Holttum<br />
Family 33. LOMARIOPSIDACEAE (including Nephrolepidaceae)<br />
Cyclopeltis crenata (Fee) C Chr.<br />
Cyclopeltis zamboangana Copel.<br />
Cyclopeltis presliana (J.Sm.) Berkeley<br />
Cyclopeltis cumingiana (Fee) C Morton<br />
Lomariopsis kingii (Copel.) Holttum<br />
Lomariopsis lineata (C Presl) Holttum<br />
Lomariopsis smithii Fée<br />
Lomariopsis papyracea Copel.<br />
1. Cyclopeltis J Smith<br />
2. Lomariopsis Fée<br />
Lomariopsis subtrifoliata (Copel.) Holttum<br />
Doubtfully occurring in the <strong>Philippine</strong>s:<br />
Lomariopsis spectabilis (Kunze) Mett.<br />
<strong>Philippine</strong> specimens formerly so-named to this species all appear to have narrower<br />
fertile pinnae and hence believed by Holttum to be L. kingii. (Fl. Males. ser. 2, 1<br />
(1978) 262.<br />
Nephrolepis acutangula C Presl<br />
Nephrolepis pubescens Copel.<br />
Nephrolepis auriculata (L.) Trimen<br />
Nephrolepis biserrata (Sw.) Schott<br />
Nephrolepis clementis Christ<br />
Probably synonymous with N. radicans.<br />
Nephrolepis cordifolia (L.) Presl<br />
Nephrolepis glabra Copel.<br />
Nephrolepis exaltata (L.) Schott<br />
Nephrolepis falcata (Cav.) C Chr.<br />
Nephrolepis hirsutula (G Forst.) C Presl<br />
3. Nephrolepis Schott<br />
Nephrolepis multiflora (Roxb.) Jarrett ex C Morton<br />
Nephrolepis radicans (Burm.) Kuhn
Family 34. TECTARIACEAE<br />
Revision:<br />
Holttum, RE. 1991. Tectaria Group. <strong>Flora</strong> Malesiana series 2, 2: 1-132. (Tectaria Group =<br />
Tribe Tectarieae)<br />
Aenigmopteris Holttum -2<br />
Arthropteris J Sm. ex JD Hook. -4<br />
Ctenitis (C Chr.) C Chr. – 14<br />
Heterogonium C Presl - 6<br />
Pleocnemia C Presl -6<br />
Psomiocarpa C Presl -1<br />
Pteridrys C Chr. & Ching -2<br />
Tectaria Cav. –32 spp., 3 hybrids<br />
Tectaridium Copel. -1<br />
1. Aenigmopteris Holttum<br />
Aenigmopteris dubia (Copel.) Holttum<br />
Ctenitis dubia (Copel.) Copel.<br />
LUZON: Quezon (Mt Banahaw de Lucban), NEGROS: Negros Oriental (Cuernos<br />
Mtns), LEYTE, CAMIGUIN. Terrestrial, altitude 850-1200m.<br />
Aenigmopteris mindanaensis Holttum<br />
Arthropteris beckleri (Hook.) Mett.<br />
Arthropteris obliterata (R Br.) J Sm.<br />
Arthropteris palisotii (Desv.) Alston<br />
Arthropteris glabra Copel.<br />
Arthropteris repens (Brackenr.) C Chr.<br />
Arthropteris wollastonii (Ridley) Holttum<br />
Arthropteris orientalis (J. Gmel.) C Chr.<br />
Ctenitis aciculata (Baker) Ching<br />
2. Arthropteris J Smith ex JDHook.<br />
3. Ctenitis (C Christensen) C Christensen<br />
Ctenitis atrorubens Holttum<br />
Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching<br />
Ctenitis boholensis Holttum<br />
Ctenitis bulusanica Holttum<br />
Ctenitis decurrentipinnata (Ching) Ching<br />
Ctenitis erythradenia Holttum<br />
Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching<br />
Ctenitis humilis Holttum
Ctenitis paleolata Copel.<br />
LUZON: Benguet (Pauai), altitude c. 2400m. Known only from the type collection.<br />
Ctenitis pallens (Brackenr.) MG Price<br />
Ctenitis rizalensis (Christ) Copel.<br />
As C. rizalensis: LUZON: Rizal, Laguna (Mt Makiling), SAMAR, MINDANAO:<br />
Zamboanga (San Ramon), Davao (Mt Apo). Terrestrial under moist forest shade,<br />
altitude 400-1200m.<br />
Ctenitis propinqua (J Sm. ex C Presl) Copel.<br />
LUZON: Pampanga (Mt Arayat), Laguna (Mt Makiling), Quezon (Mt Banahaw),<br />
Batangas (Mt Makulod).<br />
Ctenitis setosa (C Presl) Holttum<br />
Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching<br />
Ctenitis silvatica Holttum<br />
Ctenitis vilis sensu Copel. (1960), pro parte, non (Kunze) Ching<br />
Ctenitis subglandulosa (Hance) Ching<br />
Ctenitis rhodolepis (Clarke) Ching<br />
Ctenitis subobscura (Christ) Holttum<br />
Excluded:<br />
Ctenitis subconnexa (Christ) Holttum var. subconnexa and var. alstonii Holttum<br />
A mislocalization to the <strong>Philippine</strong>s by Holttum, Fl. Males. ser. 2, 2 (1991) 28. The<br />
locality Batjan is in Halmahera (<strong>of</strong> the Moluccas).<br />
Heterogonium aspidioides C Presl<br />
Heterogonium lobulatum Holttum<br />
4. Heterogonium C Presl<br />
Heterogonium pinnatum (Copel.) Holttum<br />
Stenosemia pinnata Copel.<br />
Heterogonium pr<strong>of</strong>ereoides (Christ) Copel.<br />
Heterogonium aspidioides sensu Holttum, Kalikasan 4 (1975) 224, pro parte<br />
Heterogonium sagenioides (Mett.) Holttum<br />
Ctenitis sagenioides (Mett.) Copel.<br />
Ctenitis obscura (Hook.) Copel.<br />
Heterogonium obscurum (Hook.) Holttum<br />
Heterogonium wenzelii (Copel.) Holttum<br />
Ctenitis wenzelii Copel.<br />
LEYTE.<br />
Pleocnemia brongniartii (Bory) Holttum<br />
Arcypteris brongniartii (Bory) Holttum<br />
Tectaria irregularis sensu Copel. (1960), pro parte<br />
5. Pleocnemia C Presl
Pleocnemia conjugata (Blume) C Presl<br />
Pleocnemia cumingiana C Presl<br />
Tectaria leuzeana sensu Copel. (1960), pro parte<br />
Pleocnemia irregularis (C Presl) Holttum<br />
Arcypteris irregularis (C Presl) Ching<br />
Tectaria irregularis sensu Copel. (1960), pro parte<br />
Pleocnemia macrodonta (Fee) Holttum<br />
Arcypteris macrodonta (C Presl) Holttum<br />
Tectaria irregularis sensu Copel. (1960), pro parte<br />
Pleocnemia presliana Holttum<br />
LUZON: Cagayan, Aurora, Laguna, PALAWAN (Mt Gantung; Imolnod), LEYTE<br />
(Palo), MINDANAO: Davao (Mt Apo, Todaya), Surigao del Norte. Terrestrial, usually<br />
in partly shaded area, near sea-level to at least 400m.<br />
6. Psomiocarpa C Presl<br />
Psomiocarpa apiifolia C Presl<br />
LUZON: Cagayan, Isabela, Bataan, Rizal, Laguna (Mt Makiling), Quezon, POLILLO,<br />
MASBATE, LEYTE, SAMAR, MINDANAO: Zamboanga (San Ramon). Terrestrial in<br />
forest, from near sea-level to c. 450m altitudes.<br />
7. Pteridrys C Christensen & Ching<br />
Pteridrys microthecia (Fee) C Chr. & Ching<br />
Pteridrys syrmatica (Willd.) C Chr. & Ching<br />
Tectaria adenophora Copel.<br />
SIBUYAN.<br />
Tectaria angulata (Willd.) Copel.<br />
Tectaria trifolia (Alderw.) C Chr.<br />
Stenosemia C Presl =Tectaria, Heterogonium<br />
8. Tectaria Cavanilles<br />
Tectaria athyriosora MG Price<br />
BOHOL, SAMAR, MINDANAO: Surigao del Norte.<br />
Tectaria aurita (Sw.) S. Chandra<br />
Stenosemia aurita (Sw.) Presl<br />
Tectaria barberi (Hook.) Copel.<br />
Tectaria beccariana (Cesati) C Chr.<br />
Tectaria calcarea (C Presl) Copel.
POLILLO, in wet forest, altitude 50m, BOHOL, LEYTE.<br />
Tectaria christii Copel.<br />
LUZON: Mountain Province, Benguet, MINDORO: Mindoro Oriental (Mt Halcon).<br />
Usually on limestone, altitude c. 1500m.<br />
Tectaria crenata Cav.<br />
Tectaria grandifolia sensu Copel. (1960), pro parte<br />
Tectaria decurrens (C Presl) Copel.<br />
Tectaria peralata Copel.<br />
Tectaria devexa (Kunze ex Mett.) Copel.<br />
var. devexa<br />
Tectaria dissecta (G Forst.) Lellinger<br />
Ctenitis dissecta (G Forst.) Copel.<br />
Tectaria dolichosora Copel.<br />
LUZON: Cagayan, Quezon (Infanta, Tignoan). Steep slopes near summits.<br />
Tectaria griffithii (Baker) C Chr.<br />
var. griffithii<br />
Tectaria malayensis (Christ) Copel.<br />
Tectaria hilocarpa (Fée) MG Price<br />
Hemigramma latifolia (Meyen ex Goldm.) Copel., excl. Hemionitis zollingeri and other latter names<br />
based on this.<br />
Tectaria laxa (Copel.) MG Price<br />
Ctenitis laxa Copel.<br />
LUZON: Ilocos Norte (Mt Quebrada; Mt Siminublan). Altitude 80-1100m.<br />
Tectaria lobbii (Hook.) Copel.<br />
var. lobbii<br />
Tectaria manilensis (C Presl) Holttum<br />
var. manilensis<br />
Lastreopsis manilensis (C Presl) MG Price<br />
Ctenitis manilensis (C Presl) Holttum<br />
Ctenitis loheri (Christ) Copel.<br />
Tectaria melanocaula (Blume) Copel.<br />
Tectaria ‘melanocaulis’ in Copel. (1960).<br />
Tectaria menyanthidis (C Presl) Copel.<br />
Tectaria mesodon (Copel.) MG Price<br />
Ctenitis mesodon (Copel.) Copel.<br />
Dryopteris mesodon Copel.<br />
LUZON: Ilocos Norte, MINDANAO: Agusan del Norte (Mt Hilong-hilong), altitude<br />
120m.<br />
Tectaria polymorpha (Wall. ex Hook.) Copel.<br />
Tectaria ramosii (Copel.) Holttum<br />
Ctenitis ramosii Copel.<br />
As C. ramosii: BOHOL (Dimiao), SAMAR (Catubig River). Possibly near sea-level to
c. 300m.<br />
Probable hybrids:<br />
Tectaria ramosii x Psomiocarpa apiifolia C Presl<br />
Tectaria ramosii x Tectaria aurita (Sw.) S. Chandra<br />
Tectaria repanda (Willd.) Holttum<br />
Tectaria crenata sensu Copel. (1960), pro parte<br />
Probable hybrid:<br />
Tectaria repanda x Tectaria hilocarpa (Fée) MG Price<br />
Tectaria siifolia (Willd.) Copel.<br />
Tectaria stalactica MG Price<br />
Tectaria sulitii Copel.<br />
Tectaria loheri Copel.<br />
Tectaria grandifolia sensu Copel. (1960), pro parte<br />
According to Holttum, Fl. Males. ser. 2, 2 (1991) 91, there may be intermediates<br />
between this species and T. crenata.<br />
As T. loheri: LUZON: Rizal (Montalban; Sumag).<br />
Tectaria suluensis Holttum<br />
Tectaria tabonensis MG Price<br />
Tectaria taccifolia (Fée) MG Price<br />
Hemigramma taccifolia (Fée) Copel.<br />
Tectaria trichotoma (Fée) Tagawa<br />
Tectaria balabacensis (Christ) MG Price<br />
Ctenitis balabacensis (Christ) Copel.<br />
As Ct. balabacensis: LUZON: Quezon (Guinayangan), PALAWAN, BALABAC,<br />
SIBUYAN. Petrophytic along rivers and streams in forests, altitude 100-200m in<br />
Palawan.<br />
Tectaria trifida (Fée) MG Price<br />
Tectaria irrigua (J Sm. ex Presl) Copel.<br />
Tectaridium macleanii Copel.<br />
Tectaridium primitivum Copel.<br />
1. Oleandra benguetensis Copel.<br />
2. Oleandra cumingii J Sm.<br />
Oleandra scandens Copel.<br />
3. Oleandra herrei Copel.<br />
9. Tectaridium Copeland<br />
Family 35. OLEANDRACEAE<br />
1. Oleandra Cavanilles
4. Oleandra maquilingensis Copel.<br />
Oleandra colubrina sensu Copel., Fern Fl. Philipp. 1 (1958) 181, pro parte<br />
5. Oleandra nitida (Copel.) Copel.<br />
6. Oleandra pistillaris (Sw.) C Chr.<br />
Oleandra neriiformis Cav.<br />
Oleandra colubrina (Blanco) Copel.<br />
7. Oleandra sibbaldii Grev.<br />
Family 36. DAVALLIACEAE (2 genera, 15 species, 2 varieties)<br />
1. Davallia angustata Hook. & Grev.<br />
Humata angustata (Wall.) J Sm.<br />
Humata microsora Alderw.<br />
Araiostegia Copeland =Davallia<br />
1. Davallia J Smith<br />
2. Davallia brevipes Copel. ; --Hovenkamp & de Joncheere, Blumea 33 (1988) 408.<br />
<strong>Philippine</strong>s, Sulawesi, New Guinea.<br />
3. Davallia denticulata (NL Burm.) Mett., ex Kuhn, Fil. Deck., (1867) 27;<br />
var. denticulata<br />
var. elata (G Forst.) Kuhn<br />
4. Davallia divaricata Blume<br />
var. divaricata<br />
5. Davallia embolostegia Copel., PJS 1 (1906) Suppl. 147, t. 3;<br />
6. Davallia falcinella (J Sm.) C Presl<br />
Trogostolon falcinellus (C Presl) Copel.<br />
7. Davallia heterophylla Sm.<br />
Humata heterophylla (J Sm.) Desv., Prodr. (1825) 323<br />
Humata ophioglossa Cav.<br />
8. Davallia hymenophylloides (Blume) Kuhn<br />
Araiostegia hymenophylloides (Blume) Copel., PJS 34 (1927) 241;<br />
9. Davallia pectinata Sm.<br />
Humata pectinata (J Sm.) Desv.<br />
10. Davallia repens (L.f.) Kuhn<br />
Humata intermedia C Chr.<br />
Humata obtusata Alderw.<br />
Humata pusilloides Copel.<br />
Humata repens (L.f.) Diels in Engl. & Prant, Nat. Pflanzenfam. 1 4 (1899) 209;<br />
Humata trifoliata Cav.<br />
Humata vestita (Blume) T Moore, Index (1857) 92;<br />
11. Davallia sessilifolia Blume
12. Davallia solida (Forst.) Sw., Schrad. J. Bot. 1800 (1801) 87;<br />
Davallia elmeri Copel.<br />
Davallia robinsonii Copel.<br />
Malesia to Polynesia. Throughout the <strong>Philippine</strong>s. Epiphytic or epilithic, sea-level to c.<br />
600m.<br />
13. Davallia trichomanoides Blume<br />
var. trichomanoides<br />
var. lorrainii (Hance) Holttum<br />
Davallia lorrainii Hance, Ann. Sci. Nat. 5, 5 (1866) 254;<br />
14. Davallia wagneriana Copel.<br />
1. Davallodes hirsutum (C Presl) Copel.<br />
Davallodes grammatosorum Copel.<br />
Davallodes gymnocarpum Copel.<br />
Davallodes laxum Copel.<br />
Davallodes congestum Copel.<br />
2. Davallodes (Copeland) Copeland<br />
Humata Cavanilles =Davallia<br />
Trogostolon Copeland =Davallia falcinella<br />
Family 37. POLYPODIACEAE (including Drynariaceae, Grammitidaceae,<br />
Loxogrammaceae, Platyceriaceae)<br />
(23 genera, 166 species, 2 varieties; in addition, 1 possible new record in<br />
Loxogramme)<br />
Revision:<br />
Hovenkamp, PH, MTM Bosman, E Hennipman, HP Nooteboom, G. Rodl-Linder & MC Roos.<br />
1998. Polypodiaceae. <strong>Flora</strong> Malesiana series 2, 3: 1-234.<br />
1. Acrosorus Copeland<br />
1. Acrosorus friderici-et-pauli (Christ) Copel.,<br />
Acrosorus exaltatus Copel.<br />
Acrosorus merrillii Copel.<br />
Acrosorus symmetricus Copel.<br />
2. Acrosorus nudicarpus PM Zamora & Co, Nat. Appl. Sc. Bull. Univ. Philipp. 32 (1980) 47,<br />
t. 2<br />
Xiphopteris nudicarpa (PM Zamora & Co) Parris in Parris et al., Pl. Mt Kinabalu 1 (1992) 77.<br />
Grammitis palasaba MG Price, Contr. Univ. Mich. Herb. 16 (1987) 196.<br />
Acrosorus streptophyllus (Baker) Copel.<br />
Recorded by Price (1972); some material so identified belongs to A. friderici-et-pauli.<br />
Occurrence in the <strong>Philippine</strong>s needs confirmation.<br />
Acrosorus triangularis (Scort. ex Bedd.) Copel.
2. Aglaomorpha Schott<br />
1. Aglaomorpha acuminata (Willd.) Hovenkamp<br />
Photinopteris acuminata (Willd.) C Morton<br />
Photinopteris speciosa C Presl<br />
2. Aglaomorpha cornucopia (Copel.) Roos<br />
Thayeria cornucopia Copel..<br />
3. Aglaomorpha heraclea (Kunze) Copel.<br />
Drynariopsis heraclea (Kunze) Ching<br />
4. Aglaomorpha meyeniana (Hook.) Schott<br />
5. Aglaomorpha pilosa (Hook. & Bauer) Copel.<br />
6. Aglaomorpha splendens (Hook. & Bauer) Copel.<br />
1. Arthromeris proteus (Copel.) Tagawa<br />
1. Belvisia mucronata (Fée) Copel.<br />
var. mucronata<br />
Anapausia C Presl =Leptochilus<br />
3. Arthromeris (T Moore) J Sm.<br />
4. Belvisia Mirbel<br />
2. Belvisia platyrhynchos (Kunze) Copel.<br />
3. Belvisia spicata (L.f.) Copel.<br />
Belvisia revoluta (Blume) Copel.<br />
4. Belvisia validinervis (Kunze) Copel.<br />
var. validinervis<br />
1. Calymmodon clavifer (Hook.) T Moore<br />
5. Calymmodon C Presl<br />
2. Calymmodon cucullatus (Nees & Blume) C Presl<br />
3. Calymmodon gracilis (Fée) Copel.<br />
4. Calymmodon muscoides (Copel.) Copel.<br />
5. Calymmodon ordinatus Copel.<br />
Chrysogrammitis Parris
Chrysogrammitis glandulosa (J Sm.) Parris<br />
Ctenopteris merrittii (Copel.) Tagawa<br />
Chrysogrammitis musgraviana (Baker) Parris<br />
Xiphopteris apoensis (Copel.) Copel.<br />
Ctenopteris Blume = To Chrysogrammitis, Ctenopterella, Dasygrammitis,<br />
Prosaptia, Themelium, Tomophyllum<br />
1. Christopteris sagitta (Christ) Copel.<br />
Christopteris copelandii Christ<br />
Christiopteris Copeland =Christopteris<br />
7. Christopteris Copeland<br />
[probably assignable to Tribe Microsoreae]<br />
Colysis Presl =Leptochilus<br />
Crypsinus Presl =Selliguea<br />
Ctenopterella Parris<br />
Ctenopterella blechnoides (Grev.) Parris, Gard. Bull. Sing. 58 (2007) 235.<br />
Ctenopteris blechnoides (Grev.) Wagner & Grether<br />
Ctenopterella denticulata (Blume) Parris<br />
Ctenopteris solida Kunze in Zoll., Syst. Verz. (1854) 37<br />
Ctenopterella pediculata (Baker) Parris, Gard. Bull. Sing. 58 (2007) 237.<br />
Ctenopteris pediculata (Baker) Copel.<br />
Dasygrammitis Parris<br />
Dasygrammitis fuscata (Blume) Parris, Gard. Bull. Sing. 58 (2007) 239.<br />
Ctenopteris fuscata (Blume) Kunze<br />
Dasygrammitis malaccana (Baker) Parris<br />
Ctenopteris matutumensis Copel., Fern Fl. Philip. 3 (1960) 528, anglice, non rite publice<br />
Ctenopteris pachycaulis (Copel.) Copel.<br />
Dasygrammitis mollicoma (Nees & Blume) Parris, Gard. Bull. Sing. 58 (2007) 239.<br />
Ctenopteris mollicoma (Nees & Blume) Kunze<br />
Dendroglossa C Presl =Leptochilus<br />
Diblemma J Smith =Microsorum samarense
1. Drynaria descensa Copel.<br />
Drymoglossum C Presl =Pyrrosia<br />
8. Drynaria (Bory) J Smith<br />
2. Drynaria quercifolia (L.) J Sm.<br />
India and S China, across Malesia to tropical Australia and Fiji. Throughout the<br />
<strong>Philippine</strong>s. A very common epiphyte or lithophyte in seasonal areas, in open forests,<br />
about settlements, etc., usually at low altitudes.<br />
3. Drynaria rigidula (Sw.) Bedd.<br />
4. Drynaria sparsisora (Desv.) T Moore<br />
Drynariopsis (Copeland) Ching =Aglaomorpha<br />
9. Goniophlebium (Blume) C Presl<br />
1. Goniophlebium benguetense (Copel.) Copel.<br />
Schellolepis benguetensis (Copel.) Pic.-Serm.<br />
2. Goniophlenium coadunatum Barcelona & MG Price<br />
3. Goniophlebium mengtzeense (Christ) Rödl-Linder<br />
Goniophlebium argutum auctt. philipp. non J Sm.<br />
4. Goniophlebium percussum (Cav.) Wagner & Grether<br />
Schellolepis percussa Pic.-Serm.<br />
5. Goniophlebium persicifolium (Desv.) Bedd.<br />
Schellolepis persicifolia (Desv.) Pic.-Serm.<br />
6. Goniophlebium pseudoconnatum (Copel.) Copel.<br />
7. Goniophlebium subauriculatum (Blume) C Presl<br />
Goniophlebium tomentellum (C Chr.) Copel.<br />
Schellolepis tomentella (C Chr.) Pic.-Serm.<br />
Goniophlebium integrum Copel.<br />
8. Goniophlebium terrestre Copel.<br />
Schellolepis terrestris (Copel.) MG Price<br />
Grammitis auctt. Males., non Swartz =Oreogrammitis<br />
10. Lecanopteris Reinwardt<br />
1. Lecanopteris curtisii Baker, J. Bot. (1881) Icon. Plant, pl. 1607; --H Gay, Gard. Bull. Sing. 45<br />
(1993) 301.<br />
Lecanopteris deparioides (Cesati) Baker<br />
Lecanopteris pumila Copel., Polyp. Philipp. (1905) 134; Fern Fl. Philip. 3 (1960) 499, non Blume (1829)<br />
Lecanopteris philippinensis Alderw., Bull. Jard. Bot. Buitenz. Ser. 2, no. 1 (1911) 8.<br />
Sumatra, Peninsular Malaysia, <strong>Philippine</strong>s to the Moluccas. MINDANAO: Lanao del
Sur, Davao (Mt Apo), Davao Oriental (Mt Hamiguitan). Altitude 800-1200m.<br />
2. Lecanopteris lomarioides (Mett.) Copel.<br />
LUZON: Nueva Ecija.<br />
3. Lecanopteris luzonensis Hennipman, Blumea 32<br />
Lecanopteris carnosa auctt. philipp. non (Reinw.) Blume<br />
LUZON: Pangasinan, Pampanga, Bataan (Mt Mariveles), Laguna, Quezon,<br />
Camarines Norte.<br />
4. Lecanopteris sinuosa (Wall. ex Hook.) Copel.<br />
11. Lemmaphyllum C Presl<br />
1. Lemmaphyllum accedens (Blume) Donk<br />
Weatherbya accedens (Blume) Copel.<br />
2. Lemmaphyllum carnosum (Hook.) C Presl<br />
var. carnosum<br />
Lemmaphyllum spathulatum (C Presl) C Presl<br />
1. Lepisorus longifolius (Blume) Holttum<br />
Pleopeltis longifolia (Blume) Bedd.<br />
Paragramma longifolia (Bl) T Moore<br />
12. Lepisorus (J Smith) Ching<br />
2. Lepisorus thunbergianus (Kaulf.) Ching<br />
Pleopeltis thunbergiana Kaulf.<br />
1. Leptochilus axillaris (Cav.) Kaulf.<br />
2. Leptochilus decurrens Blume<br />
Anapausia decurrens (Blume) C Presl<br />
3. Leptochilus ellipticus (Thunb.) Noot.<br />
Colysis elliptica (Thunb.) Ching<br />
Colysis pothifolia (D Don) C Presl<br />
13. Leptochilus Kaulfuss<br />
4. Leptochilus macrophyllus (Blume) Noot.<br />
var. macrophyllus<br />
Colysis macrophylla (Blume) C Presl<br />
Colysis bolsteri (Copel.) Alderw.<br />
var. fluviatilis (Lauterb.) Noot.<br />
var. pedunculatus (Hook. & Grev.) Noot.<br />
Colysis membranacea (Blume) C Presl<br />
5. Leptochilus minor Fée<br />
Dendoglossa minor (Fée) Copel.<br />
Colysis minor (Fée) MG Price<br />
6. Leptochilus
Colysis hemionitidea (Wall.) C Presl<br />
For some reason, this is not accounted for in the FM revision, although quite a<br />
common species in the <strong>Philippine</strong>s.<br />
14. Loxogramme (Blume) Presl<br />
1. Loxogramme antrophyoides (Baker) C Chr.<br />
2. Loxogramme avenia (Blume) C Presl<br />
Loxogramme malayana Copel.<br />
3. Loxogramme conferta Copel.<br />
4. Loxogramme dimorpha Copel.<br />
5. Loxogramme forbesii Copel.<br />
6. Loxogramme parallela Copel.<br />
7. Loxogramme scolopendrioides (Gaudich.) C Morton<br />
Loxogramme scolopendrina (Bory) C Presl<br />
8. Loxogramme subselliguea (Baker) Alston<br />
Loxogramme major Copel.<br />
Possible new record to the <strong>Philippine</strong>s:<br />
9. Loxogramme chinensis<br />
Micropolypodium Hayata<br />
Micropolypodium pulogense (Copel.) AR Sm.<br />
Xiphopteris pulogensis (Copel.) Copel.<br />
15. Microsorum Link<br />
1. Microsorum commutatum (Blume) Copel.<br />
Phymatosorus commutatus (Blume) Pic.-Serm.<br />
Microsorum phanerophlebium (Copel.) Copel.<br />
2. Microsorum congregatifolium (Alderw.) Holttum<br />
Microsorum mindanaense (Christ) Copel.<br />
3. Microsorum ensatum (Thunb.) H Ito<br />
Neocheiropteris ensata (Thunb.) Ching<br />
4. Microsorum heterocarpum (Blume) Ching<br />
5. Microsorum heterolobum (C Chr.) Copel.<br />
Probably a hybrid.<br />
6. Microsorum insigne (Blume) Copel.<br />
Microsorum decurrens (Brackenr.) Copel.<br />
Microsorum palmatum auct. non (Blume) Fee<br />
7. Microsorum longissimum Fée<br />
Microsorum sablanianum (Christ) Copel.
8. Microsorum membranifolium (R Br.) Ching<br />
Microsorum plukenetii (C Presl) MG Price<br />
Microsorum alternifolium auct.: Copel., Gen. Fil. (1947), pro parte<br />
Microsorum longipes Copel., nom. nud.<br />
Microsorum nigrescens (Blume) Copel.<br />
Phymatosorus nigrescens (Blume) Pic.-Serm.<br />
?Microsorum schneideri (Christ) Copel.<br />
9. Microsorum monstruosum (Copel.) Copel.<br />
10. Microsorum pentaphyllum (Baker) Copel.<br />
Microsorum curranii (Copel.) Copel.<br />
11. Microsorum pteropus (Blume) Copel.<br />
12. Microsorum punctatum (L.) Copel.<br />
Microsorum musifolium (Blume) Copel.<br />
Microsorum validum (Copel.) Ching<br />
13. Microsorum rubidum (J Sm.) Copel.<br />
14. Microsorum samarense (J Sm.) Bosman<br />
Diblemma samarense J Sm.<br />
Microsorum tenuilore (J Sm. ex Mett.) Copel.<br />
15. Microsorum sarawakense (Baker) Ching<br />
Microsorum rizalense Copel.<br />
16. Microsorum scolopendria (Burm. f.) Copel.<br />
Phymatosorus scolopendria (Burm. f.) Pic.-Serm.<br />
Tropical Africa to Polynesia and Australia. Throughout the <strong>Philippine</strong>s. Epiphytic or<br />
lithophytic in exposed sites and disturbed forests at low elevations, even on coastal<br />
bluffs subjected to salt spray. Common.<br />
17. Microsorum zippelii (Blume) Ching<br />
Note: Included in Copeland’s Fern <strong>Flora</strong> but not ascribed to the <strong>Philippine</strong>s in the FM<br />
revision is:<br />
Microsorum linguiforme (Mett.) Copel.<br />
What is Microsorum trifidum?<br />
Neocheiropteris Christ =Microsorum<br />
16. Oreogrammitis Copeland<br />
Oreogrammitis adspersa (Blume) Parris, Gard. Bull. Sing. 58 (2007) 255.<br />
Grammitis adspersa (Blume) Blume<br />
Oreogrammitis caespitosa (Blume) Parris, Gard. Bull. Sing. 58 (2007) 257.<br />
Grammitis caespitosa Blume<br />
Oreogrammitis congener (Blume) Parris, Gard. Bull. Sing. 58 (2007) 258.<br />
Grammitis congener Blume<br />
Oreogrammitis dolichosora (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 258.<br />
Grammitis dolichosora (Copel.) Copel.
Oreogrammitis fasciata (Blume) Parris, Gard. Bull. Sing. 58 (2007) 259.<br />
Grammitis fasciata Blume<br />
Oreogrammitis fenicis (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 259.<br />
Grammitis fenicis Copel.<br />
Oreogrammitis knutsfordiana (Baker) Parris, Gard. Bull. Sing. 58 (2007) 262.<br />
Grammitis knutsfordiana (Baker) Copel.<br />
Oreogrammitis locellata (Baker) Parris, Gard. Bull. Sing. 58 (2007) 262.<br />
Grammitis locellata (Baker) Copel.<br />
Grammitis stenocrypta Copel.<br />
Oreogrammitis marivelesensis (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 263.<br />
Grammitis marivelesensis Copel.<br />
Oreogrammitis merrillii (Baker) Parris, Gard. Bull. Sing. 58 (2007) 263.<br />
Grammitis merrillii (Copel.) Copel.<br />
Oreogrammitis oncobasis (MG Price) Parris, Gard. Bull. Sing. 58 (2007) 264.<br />
Grammitis oncobasis MG Price,<br />
Oreogrammitis padangensis (Baker) Parris, Gard. Bull. Sing. 58 (2007) 264.<br />
Grammitis padangensis (Baker) Copel.<br />
Oreogrammitis pilosiuscula (Blume) Parris<br />
Oreogrammitis reinwardtii (Blume) Parris, Gard. Bull. Sing. 58 (2007) 266.<br />
Grammitis reinwardtii Blume<br />
Oreogrammitis scabristipes (Baker) Parris, Gard. Bull. Sing. 58 (2007) 267.<br />
Grammitis scabristipes (Baker) Copel.<br />
Grammitis bulbotricha (Copel.) Copel.<br />
Grammitis limapes Copel.<br />
Oreogrammitis sumatrana (Baker) Parris, Gard. Bull. Sing. 58 (2007) 269.<br />
Grammitis sumatrana (Baker) Copel.<br />
Oreogrammitis torricellana (Brause) Parris, Gard. Bull. Sing. 58 (2007) 270.<br />
Grammitis torricellana (Brause) Parris<br />
Grammitis microtricha Copel.<br />
Oreogrammitis aff. insularis<br />
Oreogrammitis aff. knutsfordiana<br />
Photinopteris J Smith =Aglaomorpha<br />
Phymatodes C Presl =Microsorum<br />
Phymatopsis J Smith =Selliguea<br />
Phymatopteris Pichi-Sermolli =Selliguea
Phymatosorus Pichi-Sermolli =Microsorum<br />
17. Platycerium Desvaux<br />
1. Platycerium coronarium (König ex Müller) Desv.<br />
Tenasserim, Thailand and Malesia. In the <strong>Philippine</strong>s reported from LUZON:<br />
Cagayan to MINDANAO: Davao. High canopy epiphyte in lowland primary<br />
dipterocarp forests. Also cultivated as an ornamental.<br />
2. Platycerium grande (Fée) Kunze<br />
MINDANAO: Zamboanga, Lanao, Davao (Mt Apo). Also reported by Copeland (1960)<br />
from S LUZON (Mt Cristobal), possibly in error. Majestic crown epiphyte in forest,<br />
lowland to 1000m. Also cultivated as an ornamental.<br />
1. Podosorus angustatus Holttum<br />
Prosaptia alata (Blume) Christ<br />
Prosaptia pinnatifida C Presl<br />
Prosaptia toppingii Copel.<br />
Prosaptia ancestralis Copel.<br />
Pleopeltis Humboldt & Bonpland =Lepisorus<br />
18. Podosorus Holttum<br />
19. Prosaptia C Presl<br />
Prosaptia barathrophylla (Baker) MG Price<br />
Prosaptia celebica (Blume) Tagawa & K Iwats., Acta Phytotax. Geobot. 24 (1969) 61.<br />
Ctenopteris celebica (Blume) Copel.<br />
Ctenopteris pseudoarticulata (Copel.) Tagawa<br />
Prosaptia contigua (Forst.) C Presl<br />
Prosaptia cryptocarpa Copel.<br />
Prosaptia davalliacea (F Muell. & Baker) Copel.<br />
Prosaptia linearis Copel.<br />
Prosaptia khasyana (Hook.) C Chr. & Tardieu, Notul. Syst. (Paris) 8 (1939) 180.<br />
Ctenopteris khasyana (Hook.) Holttum<br />
Prosaptia micropora (Copel.) Parris<br />
Ctenopteris microcarpa Copel. Fern Fl. Philip. 3 (1960) 531, anglice, non rite publice<br />
Prosaptia multicaudata (Copel.) Parris, Kew Bull. 50 (1995) 166.<br />
Ctenopteris multicaudata (Copel.) Copel.
Prosaptia negrosensis (Copel.) MG Price, Contr. Univ. Michigan Herb. 17 (1990) 276.<br />
Ctenopteris negrosensis (Copel.) Copel.<br />
Ctenopteris spongiosa (Copel.) Copel.<br />
Prosaptia nutans (Blume) Mett.<br />
Ctenopteris nutans (Blume) J Sm.<br />
Prosaptia obliquata (Blume) C Chr. & Tardieu, Notul. Syst. (Paris) 8 (1939) 180.<br />
Ctenopteris obliquata (Blume) Copel.<br />
Prosaptia pectinata T Moore<br />
Prosaptia leysii (Baker) Ching<br />
Prosaptia urceolaris (Hayata) Copel.<br />
Prosaptia venulosa (Blume) MG Price, Contr. Univ. Michigan Herb. 16 (1987) 197.<br />
Ctenopteris venulosa (Blume) Kunze<br />
Prosaptia aff. cavisora<br />
Pteropsis Desvaux = Pyrrosia<br />
20. Pyrrosia Mirbel<br />
1. Pyrrosia lanceolata (L.) Farw.<br />
Pyrrosia adnascens (Sw.) Ching<br />
Pyrrosia varia (Kaulf.) Farwell<br />
India and S China, across Malesia to Polynesia. Throughout the <strong>Philippine</strong>s. In<br />
exposed sites, open forests, etc., Epiphytic or lithophytic, usually at low altitude.<br />
2. Pyrrosia longifolia (Burm.f.) C Morton<br />
Pyrrosia acrostichoides (G Forst.) Ching<br />
3. Pyrrosia nummularifolia (Sw.) Ching<br />
4. Pyrrosia piloselloides (L.) MG Price<br />
Drymoglossum piloselloides (L.) Presl<br />
Pteropsis piloselloides (L.) Desv.<br />
Pteropsis heterophyllum auct. non (L.) Trimen<br />
India to New Guinea. Throughout the <strong>Philippine</strong>s. Epiphytic on tree trunks and<br />
branches or epilithic on outcrops and boulders along rivers and streams at low<br />
altitudes. Common.<br />
5. Pyrrosia porosa (C Presl) Hovenkamp<br />
var. porosa<br />
Pyrrosia mollis auct. non (Kunze) Ching<br />
6. Pyrrosia rasamalae (Raciborski) Shing<br />
Pyrrosia lanuginosa (Giesenh.) Copel.<br />
This species has <strong>of</strong>ten been named P. floccigera, properly a synonym <strong>of</strong> P.<br />
albicans, a species not known to occur in the <strong>Philippine</strong>s.
7. Pyrrosia samarensis (C Presl) Ching<br />
Pyrrosia philippinensis Copel., nom. superfl.<br />
8. Pyrrosia sphaerotricha (Mett.) Ching<br />
9. Pyrrosia splendens (C Presl) Ching<br />
Radiogrammitis Parris<br />
Radiogrammitis alepidota (MG Price) Parris, Gard. Bull. Sing. 58 (2007) 241.<br />
Grammitis alepidota MG Price, Philip. Agriculturist 57 (1973) 34, Fig. 1;<br />
Taiwan (BJ Wang 17447-B, HAST) and <strong>Philippine</strong>s. LUZON (Ilocos Norte, Mt<br />
Burnay; Cagayan, Mt Cetaceo; Quezon, Mt Banahaw, type locality), PALAWAN (Mt<br />
Mantalingahan).<br />
Radiogrammitis jagoriana (Mett. ex Kuhn) Parris, Gard. Bull. Sing. 58 (2007) 242.<br />
Grammitis jagoriana (Mett.) Copel.<br />
Radiogrammitis setigera (Blume) Parris<br />
Grammitis intromissa (Christ) Parris<br />
Grammitis fasciculata Blume, nom. illegit. pro Polypodium setigerum Blume<br />
Radiogrammitis subpinnatifida (Blume) Parris<br />
Xiphopteris subpinnatifida (Blume) Copel.<br />
Schellolepis Pic.-Serm. =Goniophlebium<br />
21. Scleroglossum Alderwerelt<br />
Scleroglossum pusillum (Blume) Alderw.<br />
Scleroglossum minus (Fée) C Chr.<br />
Scleroglossum sulcatum (Kuhn) Alderw.<br />
Scleroglossum intermedium (Copel.) C Chr.<br />
Scleroglossum pusillum Copel., non (Blume) Alderw.<br />
22. Selliguea Bory<br />
1. Selliguea albidosquamata (Blume) Parris<br />
Crypsinus albidosquamatus (Blume) Copel.<br />
2. Selliguea caudiformis (Blume) Carruth.<br />
Selliguea calophlebia (Copel.) Copel.<br />
3. Selliguea elmeri (Copel.) Ching<br />
Selliguea feei Bory var. elmeri (Copel.) Tagawa<br />
4. Selliguea enervis (Cav.) Ching<br />
Crypsinus enervis (Cav.) Copel.<br />
5. Selliguea feei Bory<br />
Selliguea feei Bory var. vulcanica (Blume) Tagawa<br />
6. Selliguea glauca (J Sm. ex T Moore) Hovenkamp<br />
Crypsinus glaucus (J Sm. ex T Moore) Copel.
7. Selliguea heterocarpa (Blume) Blume<br />
8. Selliguea lagunensis (Christ) Hovenkamp<br />
Crypsinus lagunensis (Christ) Copel.<br />
Phymatopteris lagunensis (Christ) Pic.-Serm.<br />
9. Selliguea lateritia (Baker) Hovenkamp<br />
10. Selliguea oodes (Kunze) Hovenkamp<br />
Crypsinus oodes (Kunze) Copel.<br />
11. Selliguea pyrolifolia (Goldm.) Hovenkamp<br />
Crypsinus pyrolifolius (Goldm.) Copel.<br />
12. Selliguea stenophylla (Blume) Parris<br />
Crypsinus stenophyllus (Blume) Copel.<br />
13. Selliguea taeniata (Sw.) Parris<br />
Crypsinus taeniatus (Sw.) Copel., including var. palmatus C Chr.<br />
Phymatopteris taeniata (Sw.) Pic.-Serm.<br />
Crypsinus ramosii Copel.<br />
14. Selliguea triloba (Houtt.) MG Price<br />
Crypsinus trilobus (Houtt.) Copel.<br />
Phymatopteris triloba (Houtt.) Pic.-Serm.<br />
Selliguea matutumensis Copel.<br />
15. Selliguea whitfordii (Copel.) Hovenkamp<br />
Crypsinus whitfordii (Copel.) Copel.<br />
Crypsinus subdimorphus Copel.<br />
Doubtful occurrence in the <strong>Philippine</strong>s:<br />
Crypsinus hastatus (Thunb.) Copel.<br />
Phymatopteris hastata (Thunb.) Pic.-Serm.<br />
India, China, Japan. In the <strong>Philippine</strong>s known only from LUZON: Benguet. Epiphytic<br />
or lithophytic, alt. c. 2400m.<br />
According to Hovenkamp et.al. [Fl. Males. ser. 2, 3 (1998) 227], simple<br />
fronded plants from the <strong>Philippine</strong>s have been identified as Phymatopteris hastata by<br />
Zamora & Co [Guide to Philipp. <strong>Flora</strong> & Fauna 2 (1986) 29], and that other than<br />
through this reference, P. hastata is not known from the <strong>Philippine</strong>s.<br />
This is incorrect as Copeland (Fern Fl. Philipp. 2 (1960) 505) reported this species to<br />
be common in Benguet at 2400 m.<br />
Themelium (T Moore) Parris<br />
Themelium cf. blechnifrons (Hayata) Parris<br />
Ctenopteris curtisii (Baker) Copel.<br />
Themelium halconense (Baker) Parris, Kew Bull. 59 (2004) 225.<br />
Ctenopteris halconensis (Copel.) Copel.<br />
Themelium loherianum (Christ) Parris, Kew Bull. 59 (2004) 225.<br />
Grammitis loheriana (Christ) Copel.<br />
Themelium taxodioides (Baker) Parris, Kew Bull. 52 (1997) 739.<br />
Ctenopteris taxodioides (Baker) Copel.
Themelium tenuisectum (Blume) Parris, Kew Bull. 52 (1997) 740.<br />
Ctenopteris tenuisecta (Blume) J Sm.<br />
Themelium yoderi (Copel.) Parris, Kew Bull. 52 (1997) 739.<br />
Ctenopteris yoderi (Copel.) Copel.<br />
Themelium aff. tenuisectum sp. 1<br />
Themelium aff. tenuisectum sp. 2<br />
1. Thylacopteris papillosa (Blume) J Sm.<br />
23. Thylacopteris Kunze ex J Smith<br />
Tomophyllum (E Fournier) Parris<br />
Tomophyllum macrum (Copel.) Parris, Gard. Bull. Sing. 58 (2007) 247.<br />
Ctenopteris macra (Copel.) Copel.<br />
Tomophyllum millefolium (Blume) Parris, Gard. Bull. Sing. 58 (2007) 247.<br />
Ctenopteris millefolia (Blume) Copel.<br />
Tomophyllum minutum (Blume) Parris, Gard. Bull. Sing. 58 (2007) 247.<br />
Ctenopteris minuta (Blume) Holttum<br />
Possibly conspecific with C. macra. Blume’s name has priority.<br />
Tomophyllum repandulum (Mett.) Parris<br />
Tomophyllum subsecundodissectum (Zoll.) Parris, Gard. Bull. Sing. 58 (2007) 249.<br />
Ctenopteris subsecundodissecta (Zoll.) Copel.<br />
Weatherbya Copel. =Lemmaphyllum<br />
Xiphopteris Kaulfuss<br />
Family ISOETACEAE (1 genus, 1 species)<br />
Revision:<br />
Alston, AHG. 1959. Isoetaceae. <strong>Flora</strong> Malesiana series 2, 1: 63-64.<br />
1. Isoetes Linnaeus<br />
1. Isoetes philippinensis Merr. & Perry, Amer. Fern J. 30 (1940) 19, fig.; --Alston, Fl. Males.<br />
ser. 2, 1 (1959) 64.<br />
MINDANAO (Lanao, near Momungan, vicinity <strong>of</strong> Olangu). Bottom <strong>of</strong> a stream, 400-<br />
500m.
Family LYCOPODIACEAE (7 genera, 32 spp. + 1 var.)<br />
● Lycopodium barbatum Kaulf. -- Enum. Filic. 18 (-19). 1824 [Apr-May 1824]<br />
Diphasium Rothmaler =Diphasiastrum<br />
1. Diphasiastrum Holub<br />
Diphasiastrum multispicatum (J Wilce) Holub, Preslia 47 (1975) 108.<br />
Lycopodium multispicatum J Wilce, Nova Hedwigia 3 (1961) 103<br />
Lycopodium complanatum non L.<br />
Diphasiastrum sitchense (C Presl) Holub, Preslia 47 (1975) 108.<br />
ssp. nikoense (Franch. & Savigny) A Löve & D Löve, Taxon 26 (1977) 324.<br />
Diphasiastrum nikoense (Franch. & Savigny) Holub, Preslia 47 (1975) 108.<br />
Lycopodium sitchense Rupr. var. nikoense (Franch. & Sav.) Takeda<br />
Diphasiastrum<br />
Diphasium scariosum (G Forst.) Rothm., Repert. Spec.Nov. Reg. Veg. (1944) 54 .<br />
Lycopodium scariosum G Forst., Fl. Ins. Austr. (1786) 87<br />
Diphasiastrum wightianum (Wall. ex Grev. & Hook.) Holub, Preslia 47 (1975) 108.<br />
Lycopodium wightianum Wall. ex Grev. & Hook., Bot. Misc. 2 (1831) 379.<br />
Diphasium wightianum (Wall. ex Grev. & Hook.) Pic.-Serm., Webbia 24 (1970) 720.<br />
2. Huperzia Bernhardi<br />
Huperzia banayanica (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 71.<br />
Urostachys banayanicus Herter,<br />
Lycopodium banayanicum (Herter) Copel.<br />
Huperzia carinata (Desv.) Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) xxx<br />
Phlegamariurus carinatum (Desv.) Ching,<br />
Lycopodium carinatum Desv.<br />
Huperzia delbrueckii (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 72.<br />
Urostachys delbrueckii Herter,<br />
Huperzia edanoi (Copel.) Holub, Folia Geobot. Phytotax. 20 (1985) 72.<br />
Lycopodium edanoi Copel. Type: BS 77930 Edaño<br />
Huperzia elmeri (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 72.<br />
Urostachys elmeri Herter<br />
Huperzia fordii (Baker) Holub, Folia Geobot. Phytotax. 20 (1985) 72.<br />
Lycopodium fordii Baker, Fern Allies (1887) 17; --Hatusima, Mem. Fac. Agr. Kagoshima Univ. 5 (1966)<br />
18.<br />
Huperzia gigantea (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 73.<br />
Urostachys verticillatus L.f. var. giganteus Herter<br />
Huperzia magnusiana (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 74.<br />
Lycopodium magnusianum Herter Type: MS Clemens s.n., April 1907.<br />
Huperzia merrillii (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 74.<br />
Lycopodium merrillii (Herter) Copel.<br />
Urostachys merrillii Herter, Type: Merrill 965
Huperzia minima (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 74.<br />
Urostachys minimus Herter<br />
Huperzia nummulariifolia (Blume) Jermy in T Chambers, Jermy & Crabbe, Brit. Fern Gaz. 10<br />
(1971) 176.<br />
Phlegmariurus nummularifolius (Blume) Ching, Acta Bot. Yunnanica 3 (1981) 298<br />
Lycopodium nummularifolium Blume,<br />
Huperzia phlegmaria (L) Rothm., Feddes Repert. 54 (1944) 62.<br />
Lycopodium phlegmaria L., Sp. Pl. 2 (1753) 1101.<br />
Urostachys phlegmaria (L.) Herter, Bot. Arch. 3 (1923) 17; PJS 22 (1923) 66.<br />
Huperzia phyllantha (Hook. & Arn.) Holub, Folia Geobot. Phytotax. 20 (1985) 75.<br />
Lycopodium phyllanthum Hook. & Arn., Bot. Veechey Voy. (1832) 102.<br />
Huperzia pinifolia Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) 247<br />
Lycopodium pinifolium Blume<br />
Urostachys pinifolius (Blume) Herter, Bot. Arch. 3 (1923) 65.<br />
Huperzia piscium (Herter) Holub, Folia Geobot. Phytotax. 26 (1991) 93.<br />
Lycopodium piscium (Herter) Tagawa & K Iwatsuki, Acta Phytotax. Geobot. 22<br />
(1967) 103.<br />
Urostachys piscium Herter, Ind. Lycopod. (1949) 75.<br />
Huperzia salvinioides (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 76.<br />
Urostachys salvinioides Herter<br />
Lycopodium salvinioides (Herter) Tagawa<br />
Phlegmariurus salvinioides (Herter) Ching,<br />
Huperzia selago (L.) Bernh. ex Schrank ex Mart., Hortus Reg. Monac. (1829) 3.<br />
var. miniata (Spring) Co, stat. nov.<br />
Lycopodium selago L. var. miniatum (Spring) BC Tan & Tolentino<br />
Lycopodium miniatum Spring,<br />
Huperzia serrata (Thunb. ex Murray) Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) 248.<br />
Lycopodium serratum Thunb. ex Murray, Syst. Veg. ed.14 (1784) 944.<br />
Huperzia squarrosa (G Forst.) Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) 247<br />
Lycopodium squarrosum G Forst., Fl. Ins. Aust. (1786) 479.<br />
Huperzia toppingii (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 77.<br />
Urostachys toppingii Herter<br />
Huperzia verticillata (L.f.) Trevis., Atti Soc. Ital. Sci. Nat. 17 (1874) 248.<br />
Lycopodium verticillatum L.f., Suppl., pl. 448.<br />
Lycopodium verticillatum var. maximum Herter<br />
Huperzia whitfordii (Herter) Holub, Folia Geobot. Phytotax. 20 (1985) 78.<br />
Urostachys whitfordii Herter,<br />
3. Lateristachys Holub<br />
Lateristachys halconensis (Copel.) Holub, Folia Geobot. Phytotax., 18 (1983) 441.<br />
Lycopodium halconense Copel.<br />
4. Lycopodiastrum Holub ex RD Dixit<br />
Lycopodiastrum casuarinoides (Spring) Holub ex RD Dixit, J. Bombay Nat. Hist.Soc. 77<br />
(1980 publ. 1981<br />
Lycopodium casuarinoides Spring, Bull. Acad. Roy. Sci. Belg. 8 (1841) 521;
5. Lycopodiella Holub<br />
Lycopodiella cernua (L.) Pic.-Serm., Webbia 23 (1968) 166.<br />
Lycopodium cernuum L., Sp. Pl. 2 (1753) 1103; Herter, PJS 22 (1923) 69.<br />
var. cernua<br />
var. ingens (BC Tan & Tolentino) stat. nov.<br />
6. Lycopodium Linnaeus<br />
Lycopodium clavatum L., Sp. Pl. (1753) 1101;<br />
Phlegmariurus (Herter) Holub =Huperzia sect. ---<br />
7. Pseudodiphasium Holub<br />
Pseudodiphasium volubile (G Forst.) Holub,<br />
Lycopodium volubile G Forst., Fl. Ins. Austr. (1786) 86.<br />
Don’t know in what segregate genus to put this:<br />
Lycopodium subsentaceum C Chr.<br />
Selaginella aenea Warb.<br />
Selaginella alligans Hieron.<br />
Selaginella apoensis Hieron.<br />
Selaginella aristata Spring<br />
Family SELAGINELLACEAE<br />
1. Selaginella Palisot de Beauvois<br />
Selaginella atimonanensis BC Tan & Jermy, Fern Gaz. 12 (1981) 170, Fig. 1-6. --Type: BC<br />
Tan 77-2005 (CAHUP,holo; iso: BM, KYO, L, US).<br />
LUZON (Quezon, Atimonan, Quezon National Park). Forest on limestone, 250-430m.<br />
Known only from the type locality.<br />
Selaginella auriculata Spring<br />
Selaginella boninensis Baker<br />
Selaginella ciliaris (Retz.) Spring, Bulletin de L'Academie Royale des Sciences et Belles-<br />
Lettres de Bruxelles 10 (1) 1843<br />
Selaginella cumingiana Spring<br />
Selaginella cupressina (Willd.) Spring<br />
Selaginella flabelloides Warb.
Selaginella delicatula (Desv.) Alston, J. Bot. 70 (1932) 282.<br />
Selaginella elmeri Hieron.<br />
Selaginella engleri Hieron.<br />
Selaginella eschscholzii Hieron.<br />
Selaginella fenixii Hieron.<br />
Selaginella flagellifera Bull., Cat. (Bull.) 225 (1886) 9.<br />
Selaginella biformis A. Braun, Forsch. Gaz. 4, Bot. 6 (1889) 17.<br />
Selaginella gastrophylla Warb.<br />
Selaginella heterostachys Baker<br />
Selaginella intermedia (Blume) Spring, Bull. Acad. Brux. 10 (1843) 144;<br />
Selaginella ascendens Alderw.<br />
Selaginella intertexta Spring<br />
Selaginella involvens (Sw.) Spring, Bull. Acad. Brux. 10 (1843) 136;<br />
Selaginella peltata C Presl<br />
Selaginella polyura Warb.<br />
Selaginella jagori Warb.<br />
Selaginella halconensis Hieron.<br />
Selaginella lacerata Warb.<br />
Selaginella latifrons Warb.<br />
Selaginella agusanensis Hieron.<br />
Selaginella copelandii Hieron.<br />
Selaginella llanosii Hieron.<br />
Selaginella longiaristata Hieron.<br />
Selaginella springiana Alderw.<br />
Selaginella luzonensis Hieron.<br />
Selaginella magnifica Hieron.<br />
Selaginella moellendorfii Hieron.<br />
Selaginella myosuroides (Kaulf.) Spring<br />
Lycopodium myosuroides Kaulf. -- Enum. Filic. 19. 1824 [Apr-May 1824]<br />
Selaginella neei Hieron.<br />
Selaginella negrosensis Hieron.<br />
Selaginella nummularia Warb.<br />
Selaginella opaca Warb.
Selaginella ornata (Hook. & Grev.) Spring, Bull. Acad. Brux. 10 (1843) 232;<br />
Selaginella philippina Spring<br />
Selaginella presliana Spring<br />
Selaginella pricei BC Tan & Jermy, Fern Gaz. 12 (1981) 170, Fig. 7-10. --Type: BC Tan 75-<br />
414 (CAHUP, holo; iso: BM, KYO, L, US).<br />
SAMAR (Western Samar, trail from Concord to Bagacay, Hinabangan, type locality;<br />
Gandara and Matuginao). Primary forest on limestone, along trail and cliffs, c. 200m.<br />
Selaginella procera Alston<br />
Selaginella ramosii Hieron.<br />
Selaginella remotifolia Spring<br />
Selaginella repanda (Desv.) Spring<br />
Selaginella tamariscina (P Beauv.) Spring<br />
Selaginella usterii Hieron.<br />
Cultivated/Introduced<br />
Selaginella plana (Desv.) Hieron.<br />
Selaginella uncinata (Desv.) Spring<br />
Occurring as an escape in Luzon. Native to China.<br />
Selaginella willdenovii (Desv.) Baker<br />
Burma and Indochina to Peninsular Malaysia and Sumatra. Pantropic in cultivation.<br />
Introduced to the <strong>Philippine</strong>s.