New combinations in Jacobaea - Biodiversity Heritage Library
New combinations in Jacobaea - Biodiversity Heritage Library
New combinations in Jacobaea - Biodiversity Heritage Library
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Compositae newsletter.<br />
Columbus, Ohio :Dept. of Botany, Ohio State University,1975http://www.biodiversitylibrary.org/bibliography/12561<br />
no.44 2006: http://www.biodiversitylibrary.org/item/48846<br />
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Sponsored by: The LuEsther T Mertz <strong>Library</strong>, the <strong>New</strong> York Botanical Garden<br />
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[Beg<strong>in</strong> Page: Cover]<br />
7<br />
NEWSLETTER<br />
Number 44 20 February 2006<br />
Scientific Editor: Bertil Nordenstam<br />
Technical Editor: Gunnel Wirenius Nohl<strong>in</strong><br />
Published and distributed by The Swedish Museum of Natural History,<br />
Department of Phanerogamic Botany,<br />
PO. Box 50007, SE- 1 04 05 Stockholm, Sweden j<br />
ISSN 0284-8422<br />
APR 1 C 2005<br />
Contents<br />
Pelser, p. B., Veldkamp, J.-F. «& R. van der Meijden: <strong>New</strong> <strong>comb<strong>in</strong>ations</strong><br />
<strong>in</strong> <strong>Jacobaea</strong> Mill. (Asteraceae - Senecioneae) 1
Nordenstam, B.: Additions to the genus <strong>Jacobaea</strong> Mill.<br />
(Compositae-Senecioneae) 12<br />
LuNDiN, R.: Nordenstamia Lund<strong>in</strong> (Compositae-Senecioneae), a new<br />
genus from the Andes of South America 14<br />
X<br />
Nordenstam, B . : <strong>New</strong> <strong>comb<strong>in</strong>ations</strong> <strong>in</strong> Nordenstamia (Compositae- ^<br />
Senecioneae) from Argent<strong>in</strong>a, Bolivia, Peru and Ecuador 19<br />
Nordenstam, B.: Canariothanmus B. Nord., a new genus of the<br />
Compositae-Senecioneae, endemic to the Canary Islands 24<br />
Nordenstam, B., Kallersjo, M. & P. Eldenas: Nephrotheca, a new<br />
monotypic genus of the Compositae-Calenduleae from the southwestern<br />
Cape Prov<strong>in</strong>ce 32<br />
Nordenstam, B. : Generic revisions <strong>in</strong> the tribe Calenduleae (Compositae) 38<br />
Nordenstam, B.: <strong>New</strong> genera and <strong>comb<strong>in</strong>ations</strong> <strong>in</strong> the Senecioneae of<br />
the Greater Antilles 50
Tyler, T.: Lectotypification of names of Hieracium sect. Hieracium<br />
and sect. Vulgata based on material from southern Sweden 74<br />
<strong>New</strong> taxa and <strong>comb<strong>in</strong>ations</strong> published <strong>in</strong> this issue 94<br />
[Beg<strong>in</strong> Page: Text]<br />
[Beg<strong>in</strong> Page: Page 1]<br />
Comp. <strong>New</strong>sl. 44, 2006<br />
<strong>New</strong> <strong>comb<strong>in</strong>ations</strong> <strong>in</strong> <strong>Jacobaea</strong> Mill.<br />
(Asteraceae - Senecioneae)<br />
PlETER B. PeLSER' ^, J.-F. VeLDKAMP^, & RUUD VAN DER MeIJDEN^<br />
"Miami University, Department of Botany, 316 Pearson Hall,<br />
Oxford, Ohio 45056, USA<br />
pekerpb@muohio.edu<br />
^' Nationaal Herbarium Nederland, Universiteit Leiden branch,
P.O. Box 9514, 2300 RA Leiden, The Netherlands<br />
veldkampranhn.leidenuniv.nl<br />
This paper is dedicated to Berth Nordenstam on the occasion of his 70th birthday,<br />
Februarx' 20, 2006<br />
Abstract<br />
<strong>New</strong> <strong>in</strong>sights <strong>in</strong> the evolutionary relationships between the species of tribe<br />
Senecioneae show that at least 27 species currently assigned to Senecio sect.<br />
<strong>Jacobaea</strong> should be segregated from Senecio L. and are better placed <strong>in</strong> the genus<br />
<strong>Jacobaea</strong> Mill. In this paper, these species are listed and 19 new <strong>comb<strong>in</strong>ations</strong> <strong>in</strong><br />
<strong>Jacobaea</strong> are published.<br />
Introduction<br />
The results of recent molecular systematic studies on tribe Senecioneae show that the<br />
species of Senecio sect. <strong>Jacobaea</strong> (Mill.) Dumort. form a well-supported clade<br />
(Pelser et al. 2002, 2003, 2004, unpubl.) that is only distantly related to other species<br />
usually attributed to Senecio, <strong>in</strong>clud<strong>in</strong>g Senecio vulgaris L., the type species of this<br />
genus (Knox & Palmer 1 995, Kadereit & Jeffrey 1996, Ba<strong>in</strong> & Golden 2000, Pelser<br />
et al. 2002). These results suggest that sect. <strong>Jacobaea</strong> is more closely related to<br />
Packera and several other ma<strong>in</strong>ly <strong>New</strong> World genera <strong>in</strong> Senecioneae. These f<strong>in</strong>d<strong>in</strong>gs<br />
are conf<strong>in</strong>ned <strong>in</strong> prelim<strong>in</strong>ary phylogenetic analyses of nuclear and plastid DNA<br />
sequence data that are currently carried out as part of ongo<strong>in</strong>g phylogenetic studies<br />
of the tribe focus<strong>in</strong>g on the <strong>in</strong>tergeneric relationships between its estimated 1 50
genera and the delimitation of Senecio (Pelser et al. unpubl.). We therefore propose to<br />
transfer the 27 species of sect. <strong>Jacobaea</strong> that have been identified us<strong>in</strong>g DNA<br />
sequence data from Senecio to <strong>Jacobaea</strong> Mill. Several new <strong>comb<strong>in</strong>ations</strong> to<br />
accommodate them <strong>in</strong> <strong>Jacobaea</strong> have recently been made elsewhere (Wiebe 2000,<br />
Pelser & van der Meijden 2005). The results of our phylogenetic analyses (Pelser et<br />
[Beg<strong>in</strong> Page: Page 2]<br />
Conip. <strong>New</strong>sl. 44, 2006<br />
al. 2002, unpubl.), however, show that three species that were transferred by Wiebe<br />
(2000; S. kirghisicus DC, S. nemorensis L., and S. sarracenicus L.) are only distantly<br />
related to those oi <strong>Jacobaea</strong>, and at present are better reta<strong>in</strong>ed <strong>in</strong> Senecio.<br />
In this paper, we list all species and a number of subspecific taxa for which analyses of<br />
DNA sequence data <strong>in</strong>dicate that they should be considered members of <strong>Jacobaea</strong><br />
and make new <strong>comb<strong>in</strong>ations</strong> where necessary. On basis of morphological similarities<br />
to these species, we believe that it is likely that other taxa currently assigned to<br />
Senecio are also better placed <strong>in</strong> <strong>Jacobaea</strong>. <strong>New</strong> <strong>comb<strong>in</strong>ations</strong> for these taxa will be<br />
made elsewhere (Nordenstam 2006).<br />
When nearly f<strong>in</strong>ished with this report we accidentally discovered a publication by<br />
Gandoger (1883) where 1 9 new species of presumably <strong>Jacobaea</strong> were described and<br />
at the time of writ<strong>in</strong>g were not listed <strong>in</strong> IPNI. The types probably are <strong>in</strong> LY, but we have<br />
not <strong>in</strong>spected them and <strong>in</strong>vite a curious reader to sort this problem out. An isotype of<br />
S. batavicus Gand. (K. Ankersmit, 'Bassimi'= ? Bussum) was expected <strong>in</strong> L, where<br />
KoK Ankersmit's herbarium is deposited, but could not be found.
<strong>Jacobaea</strong> Mill., Gard. Diet. Abr. Ed. 4 ( 1 754) [667] . - Lectotype: <strong>Jacobaea</strong> vulgaris<br />
Gaertn., designated here.<br />
1. <strong>Jacobaea</strong> abrotanifolia (L.) Moench<br />
<strong>Jacobaea</strong> abrotanifolia (L.) Moench, Methodus (1794) 587. - Senecio<br />
abrotanifolius L., Sp. PI. (1753) S69. -Herbichia abrotanifolia (L.) Zaw. <strong>in</strong> Guill.,<br />
Arch. Bot. 1 ( 1 832) 45 1 , nom. superfl.- Senecio herbichia Steud., Nomencl. Bot., ed.<br />
2,1(1 840) 750; 2 ( 1 84 1 ) 557, 56 1 (sub Herbichia carpatica Zaw.).<br />
Senecio carpathicus Herbich, Addit. Fl. Galic. (1831) 43 (n.v.). - Senecio<br />
abrotanifolius subsp. carpathicus (Herbich) Nyman, Consp. Fl. Eur. (1879) 356. -<br />
Senecio abrotanifolius van carpathicus (Herbich) Hayek, Repert. Spec. Nov. Regni<br />
Veg. Beih. 30, 2 ( 1 93 1 ) 679 Ccarpaticus').<br />
Senecio monocephalus Schur, Verb. Siebenb. Ver. Naturw. 1 (1 850) 107.<br />
Senecio tiroliensis Kerner ex Dalla Torre <strong>in</strong> Sonklar et al., Anleit. Wiss. Beob.<br />
Alpenreisen 2 (ante Sep 1 882) 247; <strong>in</strong> Hart<strong>in</strong>ger, Atlas Alpenfl. (1 882) 1 33 ('i. sched.');<br />
Europ. Alpenpfl. ( 1 890) 325. - Senecio abrotanifolius van tiroliensis (Dalla Torre)<br />
Kerner ex Hegi, Illust. Fl. Mit.-Eun 6, 2 (1 914) 269. - Senecio abrotanifolius subsp.<br />
tiroliensis (Dalla Torre) Gams, Jahrb. Vere<strong>in</strong>s Schutze Alpenpfl. Alpentiere 1 ( 1 938)<br />
22.<br />
2. <strong>Jacobaea</strong> adonidifolia (Loisel.) Pelser & Veldk., comb. nov.<br />
Senecio adonidifolius Loisel., Fl. Gall. 2 (Apr 1807) 566, nom. nov. pro Senecio<br />
abrotanifolius auct., non h.- Senecio artemisiifolius Pers., Syn. 2 (Sep 1 807) 435, id..
[Beg<strong>in</strong> Page: Page 3]<br />
Comp. <strong>New</strong>sl. 44, 2006 3<br />
nom. superfl. Cartemisiaefolius').<br />
3. <strong>Jacobaea</strong> alp<strong>in</strong>a (L.) Moench<br />
<strong>Jacobaea</strong> alp<strong>in</strong>a (L.) Moench, Methodus (1794) 587. - Solidago alp<strong>in</strong>a L., Sp. PI.<br />
( 1 753) 880; Hubs., F1. Angl. Ed. 2 ( 1 778) 870 (not new). - C<strong>in</strong>eraria alp<strong>in</strong>a (L.) L., Sp.<br />
PI., ed. 2, 2 ( 1 763) 1 243, excl. van; All., F1. Pedem. 1 ( 1 785) 203, t. 38, f. 2 (not new);<br />
WiLLD., Sp. PI. Ed. 4, 3, 3 ( 1 803) 2084, isonym. - Senecio alp<strong>in</strong>us (L.) Scop., F1. Cam.,<br />
ed. 2,2(1 772) 1 64; L. f. Suppl. ( 1 782) 371, isonym. - C<strong>in</strong>eraria cordifolia Gouan, 111.<br />
Observ. Bot. ( 1 773) 69; Jacq., F1. Austr. 2 ( 1 774) 47, 1. 1 76 (not new); L. f, Suppl. ( 1 782)<br />
375, (not new); nom. superfl. -Senecio cordifolius (Gouan) Clairv., Man. Herb. (1811)<br />
24 1 , nom. superfl. (n.v.); L<strong>in</strong>k, Handb. ( 1 829) 748, isonym; non L. f. ( 1 782). - Senecio<br />
cordatus Koch, Flora 17(1 834) 6 1 4, nom. superfl. -<strong>Jacobaea</strong> cordata Opiz, Seznam<br />
(1852) 54, nom. superfl.<br />
Senecio baldensis Poir., Encycl. Meth. 7 (1 806) 1 09.<br />
Note - This synonymy of J. alp<strong>in</strong>a is probably <strong>in</strong>complete; the <strong>in</strong>fra-specific names<br />
have been omitted. Some names have been <strong>in</strong>cluded, although not new, because they<br />
are often cited as if they were, or were encountered <strong>in</strong> the Index Kewensis.<br />
4. <strong>Jacobaea</strong> ambigua (Biv.) Pelser & Veldk., comb. nov.
C<strong>in</strong>eraria ambigua Biv., Stirp. Rar. Sicilia 3(1815)5- Senecio ambiguus (Biv.) DC,<br />
Prodr. 6 ( 1 838) 356, excl. syn. C<strong>in</strong>eraria bicolor Willd. ( 1 803). - C<strong>in</strong>eraria bicolor<br />
subsp. ambigua (Biv.) Nyman, Consp. Fl. Eur. ( 1 879) 350. - Senecio gibbosus subsp.<br />
ambiguus (Biv.) Arcanc, Comp. Fl. Ital. ( 1 882) 349.<br />
C<strong>in</strong>eraria gibbosa Guss., Cat. PI. Hort. Reg. Boccadifalco (1821) 71. - Senecio<br />
gibbosus (Guss.) DC, Prodr. 6 ( 1 838) IfSS.- Senecio ambiguus subsp. gibbosus (Guss.)<br />
Chater, Bot. J. L<strong>in</strong>n. Soc. 68 ( 1 974) 274.<br />
Senecio tcngeteus Boiss. & Heldr., Diagn. PI. Orient. 1, 6 ( 1 846, ' 1 845') 95. - C<strong>in</strong>eraria<br />
nebrodensis subsp. tay^getea (Boiss. & Heldr.) Nyman, Consp. Fl. Eur (1879) 350. -<br />
Senecio ambiguus subsp. taygeteus (Boiss. & Heldr.) Greuter, Willdenowia 33 (2003)<br />
247.<br />
5. <strong>Jacobaea</strong> analoga (DC.) Veldk., comb. nov.<br />
Senecio analogus DC, Prodr. 6 ( 1 838) 366. Selected over the next by Nordenstam [Fl.<br />
h-an. 164(1989)81].<br />
Senecio spectabilis Wall, ex DC, Prodr. 6 ( 1 838) 366. - Senecio chrysanthemoides<br />
var. spectabilis (Wall, ex DC.) Hook, f., Fl. Brit. India 3(1881)339.<br />
Senecio chrysanthemoides DC, Prodr 6 ( 1 838) 365, non Schrank, Baier. Fl. 2 ( 1 789)<br />
386.
[Beg<strong>in</strong> Page: Page 4]<br />
Comp. <strong>New</strong>sl. 44, 2006<br />
Senecio sisymbhiformis DC, Prodr. 6 ( 1 838) 366. - Senecio civysanthemoides van<br />
sisymbriiformis (DC.) Hook, f., Fl. Brit. India 3 ( 1 88 1 ) 339. -Sewec/o laetm Edgew. var.<br />
sisymhriiformis (DC.) Aswai. <strong>in</strong> Aswai. & Mhhrotra, Fl. Lahaul-Spiti ( 1 994) 384, nom.<br />
<strong>in</strong>val., contra priorit.<br />
C<strong>in</strong>eraria decipiens Spreng. ex DC, Prodr. 6 (1 838) 366, cited as a synonym by the<br />
Index kewensis, is an <strong>in</strong>valid name, published as a synonym of S. spectabilis.<br />
Senecio laetiis Edgew., Trans. L<strong>in</strong>n. Soc. Bot. 20 ( 1 846) 74.<br />
6. <strong>Jacobaea</strong> aquatica (Hill) P. Gaertn., B. Mey. & Scherb.<br />
<strong>Jacobaea</strong> aquatica (Hill) P. Gaertn., B. Mey. & Scherb., Oekon. Fl. Wetterau 3 ( 1 802)<br />
2 1 0. - Senecio aquaticus Hill, Veg. Syst. 2 ( 1 76 1 ) 1 20; Huds., Fl. Angl. ( 1 762 ) 3 1 7,<br />
\sonym. - Senecio jacobaea var. aquaticus (Hill) Fiori & Beg., Fl. Ital. 3 (1903) 214,<br />
nom. superfl.<br />
Senecio praealtus Bertol., Opusc. Sci. 3 (1 819) 1 83 (n.v.).<br />
6b. <strong>Jacobaea</strong> aquatica var. erratica (Bertol.) Pelser & Meijden<br />
<strong>Jacobaea</strong> aquatica var. erratica (Bertol.) Pelser & Meijden, <strong>in</strong> Heuicels, Fl. Nederl.,<br />
ed. 23 (2005) 677.
Senecio erraticus Bertol., Rar. Ital. PI. Dec. 3 (1810) 62. - Senecio aquaticus var.<br />
erraticus (Bertol.) Coss. <strong>in</strong> Germ. & Wedd., Introd. Fl. Anal. Paris ( 1 842) 77. - Senecio<br />
jacobaea var. erraticus (Bertol.) Neilr., FI. Wien ( 1 846) 25 1 ; Paoli & Beg., FI. Ital. 3<br />
(1903) 214 (<strong>in</strong>cl. subvar. erraticus); Icon. Fl. Ital. (1903) 412, isonym. - Senecio<br />
aquaticus subsp. erraticus (Bertol.) Tourlet, Cat. PI. Vase. Indre-et-Loire (1908) 273;<br />
V.A. Matthews, Fl. Turkey 5 ( 1 950) 1 50, isonym. - Senecio jacobaea subsp. erraticus<br />
(Bertol.) Sudre, Fl. Toulous. ( 1907) 1 15 (n.v.); Rouy, Consp. Fl. France (1927) 143,<br />
isonym.<br />
Senecio 'barbareae foliis ' Krocr., Fl. Siles. 2, 2 (1790) 421, nom. <strong>in</strong>val. - Senecio<br />
aquaticus [van] a barbaraeifolius Wimm. & Grab., FI. Siles. 3 ( 1 829) 151, nom. <strong>in</strong>val. -<br />
Senecio jacobaea var. barbaraeifolius [Krock.] Fiori <strong>in</strong> Fiori & Paol., Fl. Ital. 3 ( 1 903)<br />
214; Icon. Fl. Ital. (1903) 412 {' barbareaefolius"). - Senecio erraticus subsp.<br />
barbaraeifolius [Wimm. & Grab.] Beger <strong>in</strong> Hegi, 111. Fl. Mitt.-Eur. 6, 2 (1928) 778<br />
{' barbaraeaefolius'). - Senecio aquaticus subsp. barbaraeifolius [Wimm. & Grab.]<br />
A. Pedersen, Bot. Tidsskr. 57 (I96I) 173 {"barbaraeifolius'); Walters, Bot. J. L<strong>in</strong>n.<br />
Soc. 71(1 976, ' 1 975 ' ) 273 ( 'barbareifolius '), isonym.<br />
Senecio pratens is Klett & Right., Fl. Leipzig (1830) 692 (n.v.), cited sub Senecio<br />
barbareaifolius Krock. ex Rchb., Fl. Germ. Excurs. (1832) 244 (' barbareaefolius'),<br />
nom. superfl.<br />
[Beg<strong>in</strong> Page: Page 5]<br />
Comp. <strong>New</strong>sl. 44, 2006
7. <strong>Jacobaea</strong> arnautomm (Velen.) Pelser, comb. nov.<br />
Senecio arnautorum Velen., Sitzungsber. Konigl. Bohm. Ges. Wiss., Math.-<br />
Naturwiss. CI. 1 888 (1 889) 45 - C<strong>in</strong>eraria arnautorum (Velen.) Nyman, Consp. Fl.<br />
Eur., Suppl. 2 ( 1 889) 1 63 . - Senecio alpimis subsp. arnautorum (Velen.) Hayek, Rep-<br />
ert. Spec. Nov. Regni Veg. Beih. 30, 2 ( 1 93 1 ) 680.<br />
Senecio erubescens Pan^ic, Elem. FI. Bulg. (1 883) 4 1 , non W.T. Aiton (181 3).- Senecio<br />
pancicii Degen, Magyar Bot. Lapok 1 ( 1 902) 92.<br />
8. <strong>Jacobaea</strong> auricula (Bourg.) Pelser, comb. nov.<br />
Senecio auricula Bourg. <strong>in</strong> Coss., Notes PI. Crit. ( 1 852) 1 69.<br />
9. <strong>Jacobaea</strong> boissieri (DC.) Pelser, comb. nov.<br />
Senecio boissieri DC, Prodr. 7 ( 1 838) 300.<br />
10. <strong>Jacobaea</strong> cannabifolia (Less.) E. Wiebe<br />
<strong>Jacobaea</strong> cannabifolia (Less.) E. Wiebe, Turczan<strong>in</strong>owia 3 (2000) 62. - Senecio<br />
cannabifolius Less., L<strong>in</strong>naea 6 (1 83 1 ) 242.<br />
n. <strong>Jacobaea</strong> carniolica (Willd.) Schrank<br />
<strong>Jacobaea</strong> carniolica (Willd.) Schrank, Denkschr. Acad. Munch. 1813 (1814) 316<br />
(n.v.). -Senecio carniolicus Willd. Sp. PI., ed. 4, 3, 3 ( 1 803) 1 993. - ? Senecio <strong>in</strong>canus<br />
var. carniolicus (Willd.) Heathcote, Fl . Engad<strong>in</strong>e ( 1 89 1 ) 1 2, 1. 1 2 1 , comb, <strong>in</strong>val. (s<strong>in</strong>e<br />
basion.) - Senecio <strong>in</strong>canus subsp. carniolicus (Willd.) Braun-Blanq., Neue<br />
Denkschr. Schweiz. Naturf. Ges. 48 ( 1 9 1 3) 300.
1 1 b. <strong>Jacobaea</strong> carniolica subsp. <strong>in</strong>subrica (Chenevard) Pelser, comb. nov.<br />
Senecio carniolicus van <strong>in</strong>subricus Chenevard, Bull. Herb. Boiss. II, 6 (1906) 368. -<br />
Senecio carniolicus subsp. <strong>in</strong>subricus (Chenevard) Braun-Blanq., Neue Denkschr.<br />
Schweiz. Naturf. Ges. 48 ( 1 9 1 3) 300.<br />
12. <strong>Jacobaea</strong> delph<strong>in</strong>iifoUa (Vahl) Pelser & Veldk., comb. nov.<br />
Senecio delph<strong>in</strong>iifolius Vahl, Symb. Bot. 2 ( 1 79 1 ) 9 1 , t. 45 {' delph<strong>in</strong>ifolius').<br />
Senecio cr<strong>in</strong>itus Bertol., Opusc. Sci. 1 (1817) 234 (n.v.).<br />
<strong>Jacobaea</strong> anthorifolia J. Presl & C. Presl, Delic. Prag. 1 ( 1 832) 92 {' anthoraefolia').<br />
13. <strong>Jacobaea</strong> erucifolia (L.) P. Gaertn., B. Mev. & Scherb.<br />
<strong>Jacobaea</strong> erucifolia (L.) P. Gaertn., B. Mey. & Scherb., Oekon. Fl. Wetterau 3, 1 ( 1 80 1 )<br />
208; M. BiEB. ex C. A. Mey., Verz. Pfl. Casp. Meer. ( 1 83 1 ) 8 1 ("erucaefolia'), isonym. -<br />
Senecio erucifoliiis L., Sp. PI. (1753) 869 (descr.), Addenda (2) {tpxihoi).- <strong>Jacobaea</strong><br />
rubella Moench, Methodus ( 1 794) 586, nom. superfl.<br />
[Beg<strong>in</strong> Page: Page 6]<br />
Comp. <strong>New</strong>sl. 44, 2006<br />
Senecio tenuifolius Jacq., F1. Austriac. 3 (1775) 42, t. 278. - Senecio emcifolius L.<br />
subsp. tenuifolius (Jacq.) Schubl. & G. Martens, FI. Wurtemberg (1834) 541. -
Senecio emcifolius L. var. tenuifolius Cariot, Etude Fl., ed. 5,2(1 872) 344.<br />
<strong>Jacobaea</strong> suaveolens G\UB., Fl. Lituan. 1 ( 1782) 201, nom. <strong>in</strong>val. (n.v.)<br />
<strong>Jacobaea</strong> tenuifolia J. Presl & C. Presl, Fl. Cech. (1819)175 (no ref. to Jacqu<strong>in</strong> ! ).<br />
Senecio delph<strong>in</strong>iifolius Rchb., Iconogr. Bot. PI. Crit. 4 (1826) 29 {'delph<strong>in</strong>ifolius'),<br />
nom. <strong>in</strong>val., non Vahl ( 1 79 1 ).<br />
Senecio emcifolius L. subvar. subglabm Cariot, Etude FL, ed. 5, 2 (1872) 344. -<br />
Senecio amassensis Gand., Fl. Lyon. ( 1 875) 123.<br />
Senecio chabertii Gand. ex Cariot, Etude FL, ed. 5,2(1 872) 344; Gand., Fl. Lyon. ( 1 875)<br />
Mli^chaberti').<br />
Senecio emcifolius L. var. autumnalis Gand. ex Cariot, Etude FL, ed. 5, 2 ( 1 872) 344. -<br />
Senecio autumnalis (Gand. ex Cariot) Gand., Fl. Lyon. ( 1 875) 123.<br />
13b. <strong>Jacobaea</strong> erucifolia subsp. argunensis (Turcz.) Veldk., comb. nov.<br />
Senecio argunensis Turcz., Bull. Soc. Nat. Mosc. 20, 2 (1847) 18 (n.v.).- Senecio<br />
emcifolius L. subsp. argunensis (Turcz.) E. Wiebe, Fl. Sibir. 13 (1998, '1997') 166.<br />
Senecio jacobaea L. var. grandifloms Turcz. <strong>in</strong> DC, Prodr. 6 ( 1 838) 350.<br />
Senecio praealtus var. dahuricus DC, Prodr. 6 ( 1 838) 35 1 .<br />
Senecio bl<strong>in</strong>ii H. Lev., Fedde, Repert. 8 ( 1 9 1 0) 1 38.
14. <strong>Jacobaea</strong> gallerandiana (Coss. & Durieu) Pelser, comb. nov.<br />
Senecio gallerandianus Coss. & Durieu, Bull. Soc. Bot. France 2 (1 855) 365.<br />
15. <strong>Jacobaea</strong> gigantea (Desf.) Pelser, comb. nov.<br />
Senecio giganteus Desf., FL Atlant. 2 ( 1 800) 273, t. 234.<br />
16. <strong>Jacobaea</strong> gnaphalioides (Sieber ex Spreng.) Veldk., comb. nov.<br />
Conyza gnaphalioides Sieber, Reise Kreta2 (1823) 322, t. 10, f. 14, {'gnaphalodes'),<br />
non KuNTH (1818).- Senecio gnaphalioides Sieber (Reise Kreta 1 ( 1 823) 352, nom.<br />
<strong>in</strong>val., 'gnaphalodes') ex Spreng., Syst. Veg. 4 ( 1 826) 554. - Conyza sieberi Steud.,<br />
Nomencl. Bot., ed. 2, 1 (1 840) 414. - C<strong>in</strong>eraria gnaphalioides Nyman, Consp. Fl. Eur.<br />
(1879) 351 ("gnaphalodes'). - Lectotype: Sieber s.n. (L), 'Senecio gnaphalodes<br />
Sieber. Stia', designated here.<br />
Senecio sariacus Beauverd, Bull. Soc. Bot. Geneve II, 1 2 ( 1 92 1 ) 1 5.<br />
[Beg<strong>in</strong> Page: Page 7]<br />
Comp. <strong>New</strong>sl. 44, 2006 7<br />
17. <strong>Jacobaea</strong> <strong>in</strong>cana (L.) Veldk., comb. nov.<br />
Senecio <strong>in</strong>camis L., Sp. PI. (1753) 869.<br />
Note -<strong>Jacobaea</strong> <strong>in</strong>cana Raf. (Med. Repos. <strong>New</strong> York 5 ( 1 808) 354) is a nomen nudum,
C<strong>in</strong>eraria Candida C Presl <strong>in</strong> J. Presl & C Presl, Delic. Prag. (1822) 95.- Senecio<br />
candidus (C Presl) DC, Prodr. 6 ( 1 838) 355.<br />
Senecio c<strong>in</strong>eraria DC. var. plattii Meikle, Kew Bull. 1954 (1954) 142. - Senecio<br />
bicolor wax. plattii (Meikle) Matthews, Notes Roy. Bot. Gard. Ed<strong>in</strong>burgh 33 (1975)<br />
434.<br />
Note - Othonna maritima L. and Othonna c<strong>in</strong>eraria L. were published<br />
simultaneously. We have not discovered who made the first choice between the two,<br />
but the first epithet is the one generally encountered, while for the epithet c<strong>in</strong>eraria <strong>in</strong><br />
the present context one is always referred to De Candolle ( 1 838), a nomen novum for<br />
[Beg<strong>in</strong> Page: Page 8]<br />
8 Comp. <strong>New</strong>sl. 44, 2006<br />
Othonna c<strong>in</strong>eraria necessitated <strong>in</strong> Senecio because of the older S. marilima L. f.<br />
(1781).<br />
20. <strong>Jacobaea</strong> maroccana (P.H. Davis) Pelser, comb. nov.<br />
Senecio maroccanus P.H. Davis, Notes Roy. Bot. Gard. Ed<strong>in</strong>burgh 38 (1980)445.<br />
21. <strong>Jacobaea</strong> m<strong>in</strong>uta (Cav.) Pelser & Veldk., comb. nov.<br />
C<strong>in</strong>eraria m<strong>in</strong>uta Can ., Icon. 1 (1791)21, t. 33, f. 3.-5e«ec/ow/«M/us(CAV.)DC.,Prod.
6(1838)346.<br />
C<strong>in</strong>eraria heterophylla Ortega, Nov. PI. Descr. Dec. ( 1 798) 7 1 , 1. 1 0, f. 2.<br />
22. <strong>Jacobaea</strong> othonnae (M. Bieb.) Spreng. ex C. A. Mev.<br />
<strong>Jacobaea</strong> othonnae (M. Bieb.) Spreng. ex C.A. Mey., Verz. Pfl. Casp. Meer. ( 1 83 1 ) 8 1 .<br />
- Senecio othonnae M. Bieb., F1. Taur.-Cauc. 2 (1808) 308. - Iranecio othonnae<br />
(M. Bieb.) B. Nord., F1. Iran. 1 64 ( 1 989) 58.<br />
C<strong>in</strong>eraria anomala Sm., F1. Graec. Prodr 2 ( 1 8 1 3) 1 79; Fl. Graec. 9 (1 839) 56, t. 872.<br />
Cacalia p<strong>in</strong>nata Willd., Enum. Hort Berol. (1 809) 850. - Senecio p<strong>in</strong>natus (Willd.)<br />
ScH.Bip., Flora 28 (1845) 499.<br />
Senecio rariflonts K. Koch, L<strong>in</strong>naea 17(1 843) 47. - Senecio othonnae var. rariflora<br />
(K. Koch) K. Koch, L<strong>in</strong>naea 25 ( 1 85 1 ) 366.<br />
23. <strong>Jacobaea</strong> paludosa (L.) P. Gaertn., B. Mev. & Scherb.<br />
<strong>Jacobaea</strong> paludosa (L.) P. Gaertn., B. Mey. & Scherb., Oekon. Fl. Wetterau 3 (1 802)<br />
2 1 1 .-Senecio pahdosiis L., Sp. PI. (1 753) 870. - C<strong>in</strong>eraria serratifolia Gray, Nat. Arr.<br />
Brit. PI. 2 ( 1 82 1 ) 469, nom. superfl. - Dona pahidosa (L.) Fourr., Ann. Soc. L<strong>in</strong>n. Lyon<br />
n.s. 16(1868)404.<br />
C<strong>in</strong>eraria aurea L., Sp. PI., ed. 2 (1 763) 1244. -Senecio auratus DC, Prodr. 6 (1 838)<br />
348, non 5'. aureus L. (1 753).
Senecio tataricus Less., L<strong>in</strong>naea 9 ( 1 834) 1 92. - <strong>Jacobaea</strong> tatarica (Less.) E. Weebe,<br />
Turczan<strong>in</strong>owia 3 (2000) 62.<br />
Solidago serratifolia Gilib., Fl. Lituan. 1 (1 782) 202, nom. <strong>in</strong>val. (n.v.).<br />
Senecio riparius Wallr., Sched. Crit. ( 1 822) 482.<br />
Senecio bohemicus Tausch, Syll. PI. Nov. 2 (1828) 253. -<strong>Jacobaea</strong> kosteleckii Opiz,<br />
Seznam (1852) 54, nom. superfl. - Senecio paludosus subsp. bohemicus (Tausch)<br />
Celak., Prodr. Fl. Bohmen (1871) 242.<br />
Senecio Sadler i Sadler, Fl. Comit. Pest., ed. 2 (1840)403 (n.v.).<br />
[Beg<strong>in</strong> Page: Page 9]<br />
Comp. Nevvsl. 44, 2006 9<br />
Senecio paludosus subsp. tomentosus Celak., Prodr. Fl. Bohmen (1871)242.<br />
24. <strong>Jacobaea</strong>persoonii (De Not.) Pelser, comb. nov.<br />
Senecio persoonii De Not., Repert. Fl. Ligust. (1844) 229 (n.v.). - Senecio <strong>in</strong>canus<br />
suhs^. persoonii (De Not.) Nyman, Consp. Fl. Eur. ( 1 879) 356.<br />
Senecio <strong>in</strong>canus var. italicus Pers., Syn. PI. 2 (1807) 435 - Senecio <strong>in</strong>canus subsp.<br />
italicus (Pers.) Pers. ex Arcang., Comp. Fl. Ital. (1882) 349.
25. <strong>Jacobaea</strong> subalp<strong>in</strong>a (W. D. J. Koch) Pelser i& Veldk., comb. nov.<br />
Senecio subalp<strong>in</strong>us W. D. J. Koch, Flora 17(1 834) 6 1 4, nom. nov. pro C<strong>in</strong>eraria alp<strong>in</strong>a<br />
auct. non L.f., e.g. Wahlenb., Fl. Carpat. Pr<strong>in</strong>c. (1815) 269. - C<strong>in</strong>eraria alp<strong>in</strong>a [var. B]<br />
alata L., Sp. PI., ed. 2 (1763) 1243. -Senecio alpimis (L.) Scop, subsp. subalp<strong>in</strong>us<br />
(W. D. J. Koch) Hayek <strong>in</strong> Repert. Spec. Nov. Regni Veg. Beih. 30, 2 ( 1 93 1 ) 680.<br />
C<strong>in</strong>eraria cordifolia var. auriculata Jacq., Fl. Austriac. 2 ( 1 774) 47, t. 177. -Senecio<br />
alp<strong>in</strong>us (L.) Scop, subsp. auriculatus (Jacq.) Rchb., Iconogr. Bot. PI. Crit. 2 ( 1 824) 23,<br />
1. 135. -Senecio cordatus subsp. auriculatus (Jacq.) Jacq. ex Arcang., Comp. Fl. Ital.<br />
(1882)348.<br />
26. <strong>Jacobaea</strong> imiflora (All.) Veldk., comb. nov.<br />
Solidago uniflora All., Melanges Philos. Math. Soc. Roy. Tur<strong>in</strong> 2 ( 1 773) 1 8. Senecio<br />
unijlorus (All.) All., Fl. Pedem. 1 ( 1 785) 200, t. 1 7, f. 3, non Retz. ( 1 783). -Senecio<br />
halleri Dandy, Taxon 19(1 970) 625 .<br />
Senecio chabertii Petitm., Monde PI. II, 6 (1904) 46 ('chaberti'), non Gand. ex Carjot,<br />
Etude Fl.,ed. 5, 2 (1872)344.<br />
27. <strong>Jacobaea</strong> vulgaris Gaertn.<br />
<strong>Jacobaea</strong> vulgaris Gaertn., Friict. Sem. PI. 2 ( 1 79 1 ) 445, t. 1 70, f. 1 . -Senecio jacobaea<br />
L.,Sp.Pl.(1753)870.<br />
Senecio jacobaea var. nudus Weston, Bot. Univ. 2-3 (1772) 641 -Senecio jacobaea<br />
subsp. nudus (Weston) Sojak, Cas. Cas. Nar. Muz., Odd. Pfi. 148 (1980) 77.
<strong>Jacobaea</strong> sicca G]LiB., Fl. Lithuan. 3 (1782) 149, nom. <strong>in</strong>val. (n.v.).<br />
Senecio foliosus DC, Prodr. 6 ( 1 838) 35 1 .<br />
Senecio Jlosculosits Jord., Cat. Gra<strong>in</strong>es Jard. Dijon 1 848 ( 1 848) 30, non S. jacobaea var.<br />
Jlosculosus Lam. & DC.<br />
Senecio jacobaeoides Willk., <strong>in</strong> Willk. & Lange, Prodr. Fl. Hispan. 2 (1 865) 119.<br />
[Beg<strong>in</strong> Page: Page 10]<br />
1 Comp. <strong>New</strong>sl. 44, 2006<br />
27b. <strong>Jacobaea</strong> vulgaris subsp. dunensis (Dumort.) Pelser & Meijden<br />
<strong>Jacobaea</strong> vulgaris subsp. dunensis (Dumort.) Pelser & Meijden, <strong>in</strong> Heukels, F1.<br />
Nederl.,ed.23(2005)677.<br />
Senecio dunensis Dumort., F1. Belg. ( 1 827) 66.- Senecio jacobaea L. subsp. dunensis<br />
(Dumort.) Kadereit & P.D. Sell, Watsonia 16(1 986) 23.<br />
Senecio jacobaea van nudus auct. non Weston: Meijden, Gorteria 8 (1976) 57, t. 1 .<br />
Senecio jacobaea \a.r.jIosculosus Lam. & DC. <strong>in</strong> Lam., F1. Fran9. ed. 3,4(1 805) 1 63. -<br />
<strong>Jacobaea</strong> Jlosculosa Fourr., Ann. See. L<strong>in</strong>n. Lyon, n.s. 16 (1868) 404, non Senecio<br />
jlosculosus Jord. ( 1 848).
Acknowledgements<br />
We would like to thank Bertil Nordenstam and Werner Greuter for valuable<br />
suggestions on earlier drafts of this paper.<br />
[Beg<strong>in</strong> Page: Page 11]<br />
Comp. <strong>New</strong>sl. 44, 2006 1 1<br />
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