Bryological Monograph An annotated checklist of the mosses of ...

MOSSES OF EUROPE AND MACARONESIA 201Table 3.Non-native mosses in Europe.SpeciesAchrophyllum dentatumAtrichum crispumBryum apiculatumBryum valparaisenseCalomnion complanatumCalyptrochaeta apiculataCampylopus introflexusHennediella macrophyllaHennediella stanfordensisHypopterygium tamarisciLeptophascum leptophyllumLeptotheca gaudichaudiiMyuroclada maximowicziiOrthodontium lineareSematophyllum adnatumSplachnobryum obtusumSyntrichia bogotensisTortula amplexaTortula bogosicaTortula bolanderiThuidiopsis sparsaLocationGarden, EnglandBritain and Ireland, widespread (male only)TenerifeCanary Islands, PortugalGarden, IrelandBritain, rareC. and W. Europe, widespreadBritain, widespread and spreadingBritain and Ireland, widespread and spreadingPortugalWidespread in S. and W. Europe, apparently spreadingGarden, IrelandGarden, southern RussiaC. and W. Europe, widespreadLombardy and Piedmont, several localitiesHot springs, Hungary and Azores; elsewhere in greenhousesMadeiraBritain, confined to one small areaAzores and CanariesS. Europe (rare), MacaronesiaMadeira, in a former park areacombinations are given in the second edition of his flora(Smith, 2004).Heterocladium flaccidum (Schimp.) A.J.E.Sm., stat. etcomb. nov. Basionym: Heterocladium heteropterum var.flaccidum Schimp. in Bruch, Schimp. & W.Gümbel,Bryologia Europaea 5: 154 (fasc. 49–51). 1852.Hypnum cupressiforme var. heseleri (Ando & Higuchi)M.O.Hill, stat. et comb. nov. Basionym: Hypnum heseleriAndo & Higuchi, Journal of the Hattori BotanicalLaboratory 75: 98. f. 1–2. 1994.Pelekium atlanticum (Hedenäs) Hedenäs, comb. nov.Basionym: Thuidium atlanticum Hedenäs, Journal ofBryology 16: 387. f. 1–2. 1991.Pohlia flexuosa var. pseudomuyldermansii (Arts,Nordhorn-Richter & A.J.E.Sm.) A.J.E.Sm., comb. nov.Basionym: Pohlia muyldermansii var. pseudomuyldermansiiArts, Nordhorn-Richter & A.J.E.Sm. Journal of Bryology14: 642. f. 3. 1987.Weissia6mittenii (Bruch & Schimp.) Mitt. emend.A.J.E.Sm., stat. nov. Basionym: Weissia mittenii (Bruch &Schimp.) Mitt., Annals and Magazine of Natural History,ser. 2, 8: 317. 1851.Planta originis hybridae. Parens feminea Weissia multicapsularis.Parens mas forte W. rostellata.Composition of the moss floraThe moss flora comprises 1292 species in 278 genera, 71families and 22 orders. In addition, we recognize 46subspecies and 118 varieties. Of the total 1292 species, 53are confined in Europe to the Atlantic islands ofMacaronesia (Table 2). In addition, Bryoxiphium norvegicum,Didymodon brachyphyllus and Polytrichastrumsphaerothecium are confined to Iceland, and Bryumminiatum to the Faeroe islands.In the east and south-east, several species are known onlyfrom Russia, mainly from the Urals and the Caucasus. InAsia just outside Europe, Crumia latifolia, Tortula caucasicaand Trichostomum connivens are known from theCaucasus. An additional eight species are listed byKürschner & Erdağ (2005) from Anatolia.Twenty-onespeciesofmossarethoughttohavebeenintroduced to Europe by human agency (Table 3), aremarkably small number. Mosses must have beenintroduced from the Americas, especially to Macaronesia,before moss recording began; but invasions before1900 would not have been noticed. No non-native speciesis known to have arrived before 1800, though Corley &Frahm (1982) have speculated that Campylopus pyriformisis a late 18th century introduction from the SouthernHemisphere. Indeed, the only alien known to have reachedEurope before 1900 is the North American Atrichumcrispum, which has spread slowly and is probably a singleclone. There are several other species that may beintroduced. For example Bryum gemmiferum, now widespreadin central and west Europe, appears still to bespreading, and lacks pre-1940 records (HermanStieperaere, pers. comm.).TAXONOMIC HIERARCHYThe main source for the hierarchy is Goffinet & Buck(2004). A few genera have been moved between families totake account of suggestions from Ignatov and Hedenäs orwhere there is evidence from newer molecular phylogenies.The following classes and orders occur in Europe. Ordersare included in the list of taxa, below.

202 M. O. HILL ET AL.Sphagnopsida (Engl.) OchyraSphagnales Limpr.Andreaeopsida (Limpr.) Rothm.Andreaeales Limpr.Oedipodiopsida (Schimp.) Goffinet & W.R.BuckOedipodiales (Schimp.) Goffinet & W.R.BuckPolytrichopsida Ochyra, Żarnowiec & Bednarek-OchyraPolytrichales M.Fleisch.Tetraphidopsida (M.Fleisch.) Goffinet & W.R.BuckTetraphidales M.Fleisch.Bryopsida (Limpr.) Rothm.Buxbaumiales M.Fleisch.Diphysciales M.Fleisch.Timmiales (M.Fleisch.) OchyraEncalyptales DixonFunariales M.Fleisch.Bryoxiphiales H.A.Crum & L.E.AndersonGrimmiales M.Fleisch.Archidiales Limpr.Dicranales H.Philib. ex M.Fleisch.Pottiales M.Fleisch.Splachnales (M.Fleisch.) OchyraOrthotrichales DixonHedwigiales OchyraBryales Limpr.Rhizogoniales (M.Fleisch.) Goffinet & W.R.BuckHookeriales M.Fleisch.Hypnales (M.Fleisch.) W.R.Buck & VittLIST OF TAXASphagnales Limpr.Sphagnaceae Dumort.1 Sphagnum L. [1]section Sphagnum1 affine Renauld & Cardot (S. affine var. flagellare(Schlieph. ex Röll) L.Söderstr. & Hedenäs, S. imbricatumsubsp. affine (Renauld & Cardot) Flatberg)2 austinii Sull. (S. imbricatum subsp. austinii (Sull.)Flatberg)3 centrale C.E.O.Jensen (S. palustre var. centrale(C.E.O.Jensen) A.Eddy)4 magellanicum Brid.5 palustre L.6 papillosum Lindb.section Rigida (Lindb.) Limpr.7 compactum Lam. & DC.8 strictum Sull.section Cuspidata Lindb.9 angustifolium (C.E.O.Jensen ex Russow)C.E.O.Jensen (S. recurvum var. tenue H.Klinggr.)10 annulatum H.Lindb. ex Warnst.11 balticum (Russow) C.E.O.Jensen12 cuspidatum Ehrh. ex Hoffm. (S. viride Flatberg) [2]13 fallax (H.Klinggr.) H.Klinggr. (S. brevifolium (Lindb.ex Braithw.) Röll, S. fallax subsp. isoviitae (Flatberg)M.O.Hill, S. isoviitae Flatberg, S. recurvum var.brevifolium (Lindb. ex Braithw.) Warnst.) [3]14 flexuosum Dozy & Molk.15 jensenii H.Lindb.16 lenense H.Lindb. ex L.I.Savicz [4]17 lindbergii Schimp.18 majus (Russow) C.E.O.Jensensubsp. majussubsp. norvegicum Flatberg19 obtusum Warnst.20 pulchrum (Lindb. ex Braithw.) Warnst.21 riparium Ångstr.22 tenellum (Brid.) Pers. ex Brid.23 troendelagicum Flatbergsection Subsecunda (Lindb.) Schimp.24 auriculatum Schimp. (S. denticulatum Brid. [5], S.subsecundum var. rufescens (Nees ex Hornsch.)Huebener)25 contortum Schultz (S. subsecundum var. contortum(Schultz) Huebener)26 inundatum Russow (S. denticulatum var. inundatum(Russow) Kartt., S. subsecundum subsp. inundatum(Russow) Meyl., S. subsecundum var. inundatum(Russow) C.E.O.Jensen)27 platyphyllum (Lindb. ex Braithw.) Warnst.(S. subsecundum var. platyphyllum (Lindb. exBraithw.) Cardot)28 pylaesii Brid.29 subsecundum Neessection Squarrosa (Russow) Schimp.30 squarrosum Crome31 teres (Schimp.) Ångstr.32 tundrae Flatberg [6]section Polyclada (C.E.O.Jensen) Horrell33 wulfianum Girg. [7]section Insulosa Isov.34 aongstroemii C.Hartm. [8]section Acutifolia Wilson [9, 10]35 angermanicum Melin36 arcticum Flatberg & Frisvoll37 capillifolium (Ehrh.) Hedw. (S. nemoreum Scop., S.tenerum auct. eur. non Sull. & Lesq. ex Sull.) [11]38 fimbriatum Wilsonsubsp. concinnum (Berggr.) Flatberg & Frisvollsubsp. fimbriatum39 fuscum (Schimp.) H.Klinggr.40 girgensohnii Russow41 molle Sull.42 olafii Flatberg [12]43 quinquefarium (Braithw.) Warnst.44 rubellum Wilson (S. andersonianum R.E.Andrus [14],S. capillifolium subsp. rubellum (Wilson) M.O.Hill, S.capillifolium var. tenellum (Schimp.) H.A.Crum)45 rubiginosum Flatberg [13]46 russowii Warnst.47 skyense Flatberg48 subfulvum Sjors

MOSSES OF EUROPE AND MACARONESIA 203subsp. purpureum Flatbergsubsp. subfulvum49 subnitens Russow & Warnst.subsp. ferrugineum Flatberg (S. subnitens var.ferrugineum (Flatberg) M.O.Hill)subsp. subnitens50 warnstorfii RussowAndreaeales Limpr.Andreaeaceae Dumort.2 Andreaea Hedw. [15]section Chasmocalyx Lindb. ex Braithw.1 nivalis Hook.section Nerviae Cardot ex G.Roth2 blyttii Schimp.3 crassinervia Bruch4 frigida Huebener5 heinemannii Hampe & Müll.Hal. (A. planinerviaLindb. ex G.Roth)subsp. crassifolia (Luisier) Sérgio (A. crassifoliaLuisier) [16]subsp. heinemannii6 megistospora B.M.Murray7 rothii F.Weber & D.Mohrsubsp. falcata (Schimp.) Lindb.subsp. rothiisection Andreaea8 alpestris (Thed.) Schimp. (A. rupestris var. alpestris(Thed.) Sharp)9 alpina Hedw.10 mutabilis Hook.f. & Wilson11 obovata Thed. (A. hartmanii Thed.)12 rupestris Hedw.var. papillosa (Lindb.) Podp.var. rupestris13 sinuosa B.M.MurrayOedipodiales (Schimp.) Goffinet & W.R.BuckOedipodiaceae Schimp.3 Oedipodium Schwägr.1 griffithianum (Dicks.) Schwägr.Polytrichales M.Fleisch.Polytrichaceae Schwägr.4 Alophosia Cardot1 azorica (Renauld & Cardot) Cardot5 Atrichum P.Beauv., nom. cons.1 angustatum (Brid.) Bruch & Schimp. (A. angustatumvar. rhystophyllum (Müll.Hal.) P.W.Richards &E.C.Wallace) [17]2 crispum (James) Sull.3 flavisetum Mitt. (A. haussknechtii Jur. & Milde, A.undulatum var. gracilisetum Besch.) [18]4 tenellum (Röhl.) Bruch & Schimp.5 undulatum (Hedw.) P.Beauv.6 Oligotrichum DC., nom. cons.1 hercynicum (Hedw.) Lam. & DC.7 Pogonatum P.Beauv. [19]1 aloides (Hedw.) P.Beauv. [20]2 dentatum (Menzies ex Brid.) Brid. (P. capillare(Michx.) Brid.)3 nanum (Hedw.) P.Beauv. [20]4 neesii (Müll.Hal.) Dozy [21]5 urnigerum (Hedw.) P.Beauv. (P. urnigerum var.subintegrifolium (Arnell & C.E.O.Jensen) H.Möller)8 Polytrichastrum G.L.Sm. [22]1 alpinum (Hedw.) G.L.Sm. (Pogonatum alpinum(Hedw.) Rohl., Polytrichastrum alpinum var.septentrionale (Sw. ex Brid.) G.L.Sm., P. alpinum var.fragile (Bryhn) D.G.Long, P. norwegicum (Hedw.)Schljakov, Polytrichum alpinum Hedw., P. alpinumvar. arcticum (Sw. ex Brid.) Wahlenb.) [23]2 formosum (Hedw.) G.L.Sm. (Polytrichumformosum Hedw.)3 longisetum (Sw. ex Brid.) G.L.Sm. (P. longisetum var.anomalum (Milde) Ignatov & G.L.Merr., Polytrichumlongisetum Sw. ex Brid.)4 pallidisetum (Funck) G.L.Sm. (Polytrichum formosumvar. decipiens (Limpr.) Loeske, Polytrichumpallidisetum Funck)5 sexangulare (Brid.) G.L.Sm. (Polytrichum sexangulareHedw.)6 sphaerothecium (Besch.) J.-P.Frahm (P. sexangularevar. vulcanicum (C.E.O.Jensen) G.L.Merr.,Polytrichum sphaerothecium (Besch.) Müll.Hal.) [24]9 Polytrichum Hedw. [25]1 commune Hedw. (P. commune var. humile Sw., P.commune var. perigoniale (Michx.) Hampe, P.perigoniale Michx.)2 hyperboreum R.Br.3 jensenii I.Hagen (P. commune var. jensenii (I.Hagen)Mönk.)4 juniperinum Hedw.5 piliferum Hedw. (P. piliferum var. hoppei (Hornsch.)Haller)6 strictum Menzies ex Brid. (P. affine Funck, P. alpestreHoppe)7 swartzii Hartm. (P. commune var. swartzii (Hartm.)Nyholm)8 uliginosum (Wallr.) Schriebl (P. commune var.uliginosum Wallr.)10 Psilopilum Brid.1 cavifolium (Wilson) I.Hagen2 laevigatum (Wahlenb.) Lindb. (P. laevigatum var.aloma I.Hagen)Tetraphidales M.Fleisch.Tetraphidaceae Schimp.11 Tetraphis Hedw.1 pellucida Hedw.12 Tetrodontium Schwägr.1 brownianum (Dicks.) Schwägr.2 ovatum (Funck) Schwägr.3 repandum (Funck) Schwägr.Buxbaumiales M.Fleisch.Buxbaumiaceae Schimp.13 Buxbaumia Hedw.

204 M. O. HILL ET AL.1 aphylla Hedw.2 viridis (Moug. ex Lam. & DC.) Brid. ex Moug. & Nestl.Diphysciales M.Fleisch.Diphysciaceae M.Fleisch.14 Diphyscium D.Mohr1 foliosum (Hedw.) D.MohrTimmiales (M.Fleisch.) OchyraTimmiaceae Schimp.15 Timmia Hedw., nom. cons.section Timmiaurea Brassard1 austriaca Hedw.section Timmia2 bavarica Hessl.3 megapolitana Hedw.section Norvegica Brassard4 comata Lindb. & Arnell5 norvegica J.E.Zetterst.6 sibirica Lindb. & ArnellEncalyptales DixonEncalyptaceae Schimp.16 Bryobrittonia Williams [26]1 longipes (Mitt.) D.G.Horton (Encalypta longipes Mitt.)17 Encalypta Hedw. [27] [28]section Streptotheca (Kindb.) Broth.1 procera Bruch2 streptocarpa Hedw.section Pyromitrium Wallr. ex Hampe3 alpina Sm.4 mutica I.Hagen (E. vulgaris var. mutica Brid.)section Rhabdotheca Müll.Hal.5 intermedia Jur. [29]6 obovatifolia Nyholm [30]7 rhaptocarpa Schwägr.var. leptodon Lindb. (E. rhaptocarpa var. trachymitria(Ripart) Wijk & Margad., E. trachymitria Ripart) [31]var. rhaptocarpa8 spathulata Müll.Hal. (E. rhaptocarpa var. spathulata(Müll.Hal.) Husn.)9 vulgaris Hedw.section Megasporae D.G.Horton10 longicolla Bruch [32]section Encalypta11 affinis R.Hedw.subsp. affinissubsp. macounii (Austin) D.G.Horton (E. macouniiAustin) [33]12 brevicolla (Bruch & Schimp.) Ångstr. [32]13 brevipes Schljakov14 ciliata Hedw.15 microstoma Bals.-Criv. & De Not.Funariales M.Fleisch.Funariaceae Schwägr. [34] [35]18 Entosthodon Schwägr. [36]Subgenus Entosthodon1 attenuatus (Dicks.) Bryhn (E. templetonii (Sm.)Schwägr., Funaria attenuata (Dicks.) Lindb.)2 durieui Mont. (E. durieui var. mustaphae Trab.,E. mustaphae Trab., E. pallescens Jur., E. pallescensvar. mitratus (Casares-Gil) Wijk & Margad.,E. physcomitroides Casares-Gil & Beltrán, Funariapallescens (Jur.) Lindb.) [37]3 hungaricus (Boros) Loeske (E. maroccanus (Meyl.)Hébr. & Lo Giudice, Funaria hungarica Boros,Physcomitrium maroccanum Meyl.) [38]4 krausei Besch. (Funaria krausei (Besch.) Geh. &Herzog) [39]5 obtusus (Hedw.) Lindb. (E. ericetorum (De Not.)Müll.Hal., Funaria obtusa (Hedw.) Lindb.)Subgenus Plagiodus (Mitt.) Fife6 convexus (Spruce) Brugués (Funaria convexa Spruce,Funaria pustulosa Zodda)7 muhlenbergii (Turner) Fife (Funaria calcareaWahlenb., Funaria dentata Crome, Funaria hibernicaHook., Funaria mediterranea Lindb., Funariamuhlenbergii Turner)8 pulchellus (H.Philib.) Brugués (Funaria pulchellaH.Philib.)9 schimperi Brugués (Funaria algieriensis Lindb.,Funaria durieui Schimp.) [40]Subgenus Murcia Fife10 fascicularis (Hedw.) Müll.Hal. (Funaria fascicularis(Hedw.) Lindb.)11 mouretii (Corb.) Jelenc (Funaria mouretii Corb.) [41]19 Funaria Hedw. [36] [39] [42]1 aequidens Lindb. ex Broth.2 arctica (Berggr.) Kindb.3 hygrometrica Hedw.4 microstoma Bruch ex Schimp.20 Funariella Sérgio1 curviseta (Schwägr.) Sérgio (Entosthodon curvisetus(Schwägr.) Müll.Hal., Funaria curviseta (Schwägr.)Milde)21 Goniomitrium Hook.f. & Wilson [43]1 seroi Casas [44]22 Physcomitrella Bruch & Schimp.1 patens (Hedw.) Bruch & Schimp. (Aphanorrhegmapatens (Hedw.) Lindb., Physcomitrium patens (Hedw.)Mitt.)23 Physcomitrium (Brid.) Brid.1 arenicola Laz.2 eurystomum Sendtn.subsp. acuminatum (Bruch & Schimp.) Giacom.(P. acuminatum Bruch & Schimp.) [45]subsp. eurystomum3 pyriforme (Hedw.) Bruch & Schimp.4 sphaericum (C.F.Ludw. ex Schkuhr) Brid.24 Pyramidula Brid.1 tetragona (Brid.) Brid. (P. algeriensis Chud. & Douin,P. tetragona var. zoddae Bott.) [46]Disceliaceae Schimp.25 Discelium Brid.1 nudum (Dicks.) Brid.Gigaspermaceae Lindb.

MOSSES OF EUROPE AND MACARONESIA 20526 Gigaspermum Lindb.1 mouretii Corb.27 Oedipodiella Dixon1 australis (Wager & Dixon) Dixon (O. australis var.catalaunica P.de la Varde) [47]Bryoxiphiales H.A.Crum & L.E.AndersonBryoxiphiaceae Besch., nom. cons.28 Bryoxiphium Mitt., nom. cons.1 madeirense Á.Löve & D.Löve [48]2 norvegicum (Brid.) Mitt.Grimmiales M.Fleisch.Grimmiaceae Arn.29 Coscinodon Spreng.1 cribrosus (Hedw.) Spruce (C. cribrosus var. brevipilaM.Fleisch. & Warnst.)2 humilis Milde [49]30 Grimmia Hedw.1 alpestris (F.Weber & D.Mohr) Schleich.(Orthogrimmia alpestris (F.Weber & D.Mohr) Ochyra&Żarnowiec) [50]2 anodon Bruch & Schimp. (G. limprichtii Kern) [51]3 anomala Hampe ex Schimp. (Dryptodon anomalus(Hampe ex Schimp.) Loeske)4 arenaria Hampe (G. donniana var. curvula Spruce,Orthogrimmia arenaria (Hampe) Ochyra &Żarnowiec) [52]5 atrata Miel. ex Hornsch. (Streptocolea atrata (Miel. exHornsch.) Ochyra & Żarnowiec)6 caespiticia (Brid.) Jur. (G. pyrenaica Kern,Orthogrimmia caespiticia (Brid.) Ochyra & Żarnowiec)7 capillata De Not. [53]8 crinita Brid.9 curviseta Bouman [54]10 decipiens (Schultz) Lindb. (Dryptodon decipiens(Schultz) Loeske)11 dissimulata E.Maier [55]12 donniana Sm. (G. sudetica Spreng. ex Schkuhr,Orthogrimmia donniana (Sm.) Ochyra & Żarnowiec)13 elatior Bruch ex Bals.-Criv. & De Not. (Dryptodonincurvus (Hornsch.) Brid. [56], G. elatior var.pseudofunalis Limpr.)14 elongata Kaulf. (Dryptodon elongatus (Kaulf.) Hartm.)15 funalis (Schwägr.) Bruch & Schimp. (Dryptodonfunalis (Schwägr.) Brid., G. funalis var. calvescens(Kindb.) H.Möller) [57]16 fuscolutea Hook. (G. apiculata Hornsch., G. holleriMolendo)17 hartmanii Schimp. (Dryptodon hartmanii (Schimp.)Limpr.)18 incurva Schwägr. (Dryptodon contortus (Wahlenb.)Brid.)19 laevigata (Brid.) Brid. (Guembelia laevigata (Brid.)Ochyra & Żarnowiec)20 lisae De Not. (G. retracta Stirt.) [58]21 longirostris Hook. (G. affinis Hornsch., Guembelialongirostris (Hook.) Ochyra & Żarnowiec)22 mollis Bruch & Schimp. (Hydrogrimmia mollis (Bruch& Schimp.) Loeske)23 montana Bruch & Schimp. (G. pilosissima Herzog,Orthogrimmia montana (Bruch & Schimp.) Ochyra&Żarnowiec)24 muehlenbeckii Schimp. (Dryptodon muehlenbeckii(Schimp.) Loeske) [59]25 nutans Bruch (G. meteorae C.C.Towns.)26 orbicularis Bruch ex Wilson (Dryptodon orbicularis(Bruch ex Wilson) Ochyra & Żarnowiec)27 ovalis (Hedw.) Lindb. (Guembelia ovalis (Hedw.)Müll.Hal.)28 plagiopodia Hedw. (G. plagiopodia var. arvernica(H.Philib.) Boulay)29 poecilostoma Cardot & Sebille, nom. cons. prop.(G. tergestina var. poecilostoma (Cardot & Sebille)Loeske, G. tergestina var. tergestinoides Culm.) [59]30 pulvinata (Hedw.) Sm. (Dryptodon pulvinatus (Hedw.)Brid., G. pulvinata var. africana (Hedw.) Hook.f.,G. pulvinata var. obtusa (Brid.) Huebener) [60]31 ramondii (Lam. & DC.) Margad. (Dryptodon patens(Hedw.) Brid., G. curvata (Brid.) De Sloover)32 reflexidens Müll.Hal. (G. grisea Cardot [62],G. sessitana De Not., G. subsulcata Limpr. [63],Orthogrimmia sessitana (De Not.) Ochyra&Żarnowiec) [61]33 teretinervis Limpr.34 tergestina Tomm. ex Bruch & Schimp. (G. crassifoliaLindb. ex Broth. [64], Guembelia tergestina (Tomm. exBruch & Schimp.) Buyss.)35 torquata Drumm. (Dryptodon torquatus (Drumm.)Brid.)36 trichophylla Grev. (Dryptodon trichophyllus (Grev.)Brid., G. austrofunalis auct. eur. non Müll.Hal. [65],G. britannica A.J.E.Sm. [66], G. meridionalis (Müll.Hal.)E.Maier [67], G. trichophylla var. stirtonii (Schimp.)H.Möller, G. trichophylla var. subsquarrosa (Wilson)A.J.E.Sm., G. trichophylla var. tenuis (Wahlenb.) Wijk& Margad.)37 triformis Carestia & De Not. (Orthogrimmia triformis(Carestia & De Not.) Ochyra & Żarnowiec) [68]38 ungeri Jur. [69]39 unicolor Hook.31 Indusiella Broth. & Müll.Hal.1 thianschanica Broth. & Müll.Hal. [70]32 Jaffueliobryum Thér.1 latifolium Thér. [71]33 Racomitrium Brid. [72]Subgenus Racomitrium1 lanuginosum (Hedw.) Brid.Subgenus Niphotrichum Bednarek-Ochyra(Niphotrichum (Bednarek-Ochyra) Bednarek-Ochyra& Ochyra)2 canescens (Hedw.) Brid. (Niphotrichum canescens(Hedw.) Bednarek-Ochyra & Ochyra)subsp. canescens

206 M. O. HILL ET AL.subsp. latifolium (C.E.O.Jensen) Frisvoll(Niphotrichum canescens subsp. latifolium(C.E.O.Jensen) Bednarek-Ochyra & Ochyra)3 elongatum Ehrh. ex Frisvoll (Niphotrichum elongatum(Frisvoll) Bednarek-Ochyra & Ochyra, R. canescensvar. intermedium Venturi & Bott.)4 ericoides (Brid.) Brid. (Niphotrichum ericoides (Brid.)Bednarek-Ochyra & Ochyra, R. canescens var.ericoides (Brid.) Hampe)5 panschii (Müll.Hal.) Kindb. (Niphotrichum panschii(Müll.Hal.) Bednarek-Ochyra & Ochyra)Subgenus Cataracta Vilh. (Codriophorus P.Beauv.)6 aciculare (Hedw.) Brid. (Codriophorus acicularis(Hedw.) P.Beauv.)7 aquaticum (Brid. ex Schrad.) Brid. (Codriophorusaquaticus (Brid. ex Schrad.) Bednarek-Ochyra& Ochyra)8 fasciculare (Hedw.) Brid. (Codriophorus fascicularis(Hedw.) Bednarek-Ochyra & Ochyra)9 hespericum Sérgio, J.Muñoz & Ochyra (Codriophorushespericus (Sérgio, J.Muñoz & Ochyra) Bednarek-Ochyra & Ochyra) [73]Subgenus Ellipticodryptodon (Vilh.) Bednarek-Ochyra& Ochyra (Bucklandiella Roiv.)10 affine (F.Weber & D.Mohr) Lindb. (Bucklandiellaaffinis (F.Weber & D.Mohr) Bednarek-Ochyra& Ochyra, R. heterostichum var. affine (F.Weber& D.Mohr) Lesq.)11 ellipticum (Turner) Bruch & Schimp. (Bucklandiellaelliptica (Turner) Bednarek-Ochyra & Ochyra)12 heterostichum (Hedw.) Brid. (Bucklandiellaheterosticha (Hedw.) Bednarek-Ochyra & Ochyra)13 himalayanum (Mitt.) A.Jaeger (Bucklandiellahimalayana (Mitt.) Bednarek-Ochyra & Ochyra)14 lamprocarpum (Müll.Hal.) A.Jaeger (Bucklandiellalamprocarpa (Müll.Hal.) Bednarek-Ochyra & Ochyra)15 lusitanicum Ochyra & Sérgio (Bucklandiella lusitanica(Ochyra & Sérgio) Bednarek-Ochyra & Ochyra) [74]16 macounii Kindb. (Bucklandiella macounii (Kindb.)Bednarek-Ochyra & Ochyra)subsp. alpinum (E.Lawton) Frisvoll (Bucklandiellamacounii subsp. alpina (E.Lawton) Bednarek-Ochyra& Ochyra)subsp. macounii17 microcarpon (Hedw.) Brid. (Bucklandiella microcarpa(Hedw.) Bednarek-Ochyra & Ochyra)18 obtusum (Brid.) Brid. (Bucklandiella obtusa (Brid.)Bednarek-Ochyra & Ochyra)19 sudeticum (Funck) Bruch & Schimp. (Bucklandiellasudetica (Funck) Bednarek-Ochyra & Ochyra,R. heterostichum var. sudeticum (Funck) E.Bauer)34 Schistidium Bruch & Schimp., nom. cons. [75] [76]1 agassizii Sull. & Lesq. [77]2 apocarpum (Hedw.) Bruch & Schimp.3 atrofuscum (Schimp.) Limpr. (S. apocarpum var.atrofuscum (Schimp.) C.E.O.Jensen)4 boreale Poelt5 brunnescens Limpr.subsp. brunnescens (S. apocarpum var. brunnescens(Limpr.) Loeske)subsp. griseum (Nees & Hornsch.) H.H.Blom6 bryhnii I.Hagen7 confertum (Funck) Bruch & Schimp. (S. apocarpumvar. confertum (Funck) H.Möller)8 confusum H.H.Blom9 crassipilum H.H.Blom10 crenatum H.H.Blom11 dupretii (Thér.) W.A.Weber12 elegantulum H.H.Blomsubsp. elegantulumsubsp. wilsonii H.H.Blom13 flaccidum (De Not.) Ochyra (S. pulvinatum var.flaccidum (De Not.) De Not.)14 flexipile (Lindb. ex Broth.) G.Roth15 frigidum H.H.Blomvar. frigidumvar. havaasii H.H.Blom16 frisvollianum H.H.Blom17 grande Poelt18 grandirete H.H.Blom19 helveticum (Schkuhr) Deguchi (S. singarense (Schiffn.)Laz.) [78]20 holmenianum Steere & Brassard21 lancifolium (Kindb.) H.H.Blom22 maritimum (Sm. ex R.Scott) Bruch & Schimp. [79]subsp. maritimumsubsp. piliferum (I.Hagen) B.Bremer (S. maritimumvar. piliferum (I.Hagen) Loeske)23 occidentale (E.Lawton) S.P.Churchill [80]24 papillosum Culm. (S. apocarpum subsp. papillosum(Culm.) Poelt)25 platyphyllum (Mitt.) H.Perss. (S. alpicola auct. non(Hedw.) Limpr., S. rivulare subsp. latifolium(J.E.Zetterst.) B.Bremer, S. rivulare var. latifolium(J.E.Zetterst.) H.A.Crum & L.E.Anderson) [77]subsp. abrupticostatum (Bryhn) H.H.Blomsubsp. platyphyllum26 poeltii H.H.Blom27 pruinosum (Wilson ex Schimp.) G.Roth28 pulchrum H.H.Blom29 recurvum H.H.Blom30 rivulare (Brid.) Podp.31 robustum (Nees & Hornsch.) H.H.Blom32 scandicum H.H.Blom33 sinensiapocarpum (Müll.Hal.) Ochyra [81]34 sordidum I.Hagen35 spinosum H.H.Blom & Lüth [82]36 strictum (Turner) Loeske ex Martensson37 subjulaceum H.H.Blom38 submuticum H.H.Blomsubsp. arcticum H.H.Blomsubsp. submuticum39 tenerum (J.E.Zetterst.) Nyholm40 trichodon (Brid.) Poelt

MOSSES OF EUROPE AND MACARONESIA 207var. nutans H.H.Blomvar. trichodon41 umbrosum (J.E.Zetterst.) H.H.Blom42 venetum H.H.BlomPtychomitriaceae Schimp.35 Campylostelium Bruch & Schimp.1 pitardii (Corb.) E.Maier (Grimmia pitardii Corb.) [83]2 saxicola (F.Weber & D.Mohr) Bruch & Schimp.3 strictum Solms36 Ptychomitrium Fürnr., nom. cons.1 incurvum (Schwägr.) Spruce2 nigrescens (Kunze) Wijk & Margad.3 polyphyllum (Dicks. ex Sw.) Bruch & Schimp.Seligeriaceae Schimp.37 Blindia Bruch & Schimp.1 acuta (Hedw.) Bruch & Schimp.2 caespiticia (F.Weber & D.Mohr) Müll.Hal.38 Brachydontium Fürnr.1 trichodes (F.Weber) Milde (Seligeria transylvanicaPlam. [84])39 Seligeria Bruch & Schimp. [85]Subgenus Seligeria1 acutifolia Lindb.2 brevifolia (Lindb.) Lindb.3 galinae Mogensen & I.Goldberg [86]4 pusilla (Hedw.) Bruch & Schimp.Subgenus Cyrtoseligeria Vitt5 campylopoda Kindb.6 diversifolia Lindb.7 recurvata (Hedw.) Bruch & Schimp.Subgenus Anodus (Bruch & Schimp.) Boulay8 calcarea (Hedw.) Bruch & Schimp.9 calycina Mitt. ex Lindb. (S. paucifolia auct. non(With.) Carruth.)10 donniana (Sm.) Müll.Hal.Subgenus Blindiadelphus Lindb.11 polaris Berggr.12 subimmersa Lindb.Subgenus Megalosporia Vitt13 austriaca T.Schauer14 carniolica (Breidl. & Beck) Nyholm (Trochobryumcarniolicum Breidl. & Beck)15 irrigata (H.K.G.Paul) Ochyra & Gos (S. trifaria var.irrigata H.K.G.Paul) [87]16 oelandica C.E.O.Jensen & Medelius17 patula (Lindb.) I.Hagen (S. alpestris T.Schauer,S. patula var. alpestris (T.Schauer) Gos & Ochyra,S. tristichoides var. patula (Lindb.) Broth.)18 trifaria (Brid.) Lindb.19 tristichoides Kindb.Archidiales Limpr.Archidiaceae Schimp.40 Archidium Brid.1 alternifolium (Hedw.) Mitt.Dicranales H.Philib. ex M.Fleisch.Fissidentaceae Schimp.41 Fissidens Hedw.Subgenus Pachyfissidens (Müll.Hal.) Kindb.section Pachyfissidens1 adianthoides Hedw.2 azoricus (P.de la Varde) Bizot3 dubius P.Beauv. (F. cristatus Wilson ex Mitt.,F. decipiens De Not.) [88]4 grandifrons Brid.5 luisieri P.de la Varde [89]6 osmundoides Hedw.7 polyphyllus Wilson ex Bruch & Schimp. [90]8 serrulatus Brid. [91]9 taxifolius Hedw.subsp. pallidicaulis (Mitt.) Mönk. (F. pallidicaulisMitt., F. taxifolius var. pallidicaulis (Mitt.) Corb.)subsp. taxifoliussection Amblyothallia (Müll.Hal.) Pursell &Brugg.-Nann.10 asplenioides Hedw.Subgenus Octodiceras (Brid.) Broth.11 fontanus (Bach.Pyl.) Steud. (Octodiceras fontanum(Bach.Pyl.) Lindb.)Subgenus Fissidens [92]12 arcticus Bryhn [93]13 arnoldii R.Ruthe14 bryoides Hedw. [94]var. bryoidesvar. caespitans Schimp. (F. bryoides var. curnovii(Mitt.) J.J.Amann, F. curnovii Mitt.) [95]15 coacervatus Brugg.-Nann.16 crassipes Wilson ex Bruch & Schimp.subsp. crassipes (F. crassipes var. rufipes Schimp.)subsp. warnstorfii (M.Fleisch.) Brugg.-Nann.(F. crassipes var. philibertii Besch., F. mouretii Corb.)17 crispus Mont. (F. herzogii R.Ruthe ex Herzog,F. limbatus Sull., F. minutulus Sull.) [96]18 curvatus Hornsch. (F. algarvicus Solms)19 gracilifolius Brugg.-Nann. & Nyholm (F. viridulus var.tenuifolius (Boulay) A.J.E.Sm.)20 gymnandrus Büse [97]21 jansenii Sérgio & Pursell [98]22 microstictus Dixon & Luisier [99]23 monguillonii Thér.24 nobreganus Dixon & Luisier [100]25 ovatifolius R.Ruthe (F. perssonii P.de la Varde) [101]26 pusillus (Wilson) Milde (F. exiguus auct. eur. partim[103], F. viridulus var. pusillus Wilson) [102]27 rivularis (Spruce) Schimp.28 rufulus Bruch & Schimp.29 sublimbatus Grout [104]30 sublineaefolius (P.de la Varde) Brugg.-Nann.31 viridulus (Sw. ex anon.) Wahlenb. [105] [106]var. incurvus (Starke ex Röhl.) Waldh. (F. incurvusStarke ex Röhl., F. incurvus var. tamarindifolius(Turner) Braithw.) [107]var. viridulus (F. bambergeri Milde [108], F. bryoidesvar. inconstans (Schimp.) R.Ruthe, F. bryoides var.viridulus (Sw. ex anon.) Broth., F. exiguus auct. eur.

208 M. O. HILL ET AL.partim [92] [103], F. haraldii (Lindb.) Limpr., F.limbatus var. bambergeri (Schimp. ex Milde) Düll, F.marginatulus Meln. [109], F. viridulus var. bambergeri(Schimp. ex Milde) Waldh., F. viridulus var.haraldii (Lindb.) C.E.O.Jensen, F. viridulus var.intralimbatulus (R.Ruthe) Düll) [92]Subgenus Aloma Kindb.32 celticus Paton33 exilis Hedw.34 serratus Müll.Hal. (F. papillosus Sande Lac.) [110]Ditrichaceae Limpr., nom. cons.42 Ceratodon Brid.1 conicus (Hampe) Lindb. (C. purpureus var. conicus(Hampe) Husn.)2 purpureus (Hedw.) Brid. [111]subsp. purpureus (C. purpureus var. brevifolius Milde,C. purpureus var. flavisetus Limpr., C. purpureus var.pallidisetus Luisier, C. purpureus var. xanthopus Sull.)[112]subsp. stenocarpus (Bruch. & Schimp. ex Müll.Hal.)Dixon (C. corsicus Bruch & Schimp, C. stenocarpusBruch & Schimp. ex Müll.Hal.) [112]43 Cheilothela Broth.1 chloropus (Brid.) Broth.44 Cleistocarpidium Ochyra & Bednarek-Ochyra1 palustre (Bruch & Schimp.) Ochyra & Bednarek-Ochyra (Pleuridium palustre (Bruch & Schimp.) Bruch& Schimp., Sporledera palustris (Bruch & Schimp.)Hampe)45 Distichium Bruch & Schimp., nom. cons.1 capillaceum (Hedw.) Bruch & Schimp. (D. capillaceumvar. compactum (Huebener) Dalla Torre & Sarnth.)2 hagenii Ryan ex H.Philib.3 inclinatum (Hedw.) Bruch & Schimp.46 Ditrichum Timm ex Hampe, nom. cons.1 cornubicum Paton2 flexicaule (Schwägr.) Hampe3 gracile (Mitt.) Kuntze (D. crispatissimum (Müll.Hal.)Paris, D. flexicaule var. longifolium (J.E.Zetterst.)I.Hagen, D. flexicaule var. sterile (De Not.) Limpr.,D. giganteum R.S.Williams) [113]4 heteromallum (Hedw.) E.Britton (D. homomallum(Hedw.) Hampe)5 lineare (Sw.) Lindb. (D. vaginans (Sull.) Hampe)6 pallidum (Hedw.) Hampe7 plumbicola Crundw.8 punctulatum Mitt. (Campylopus brevipilus var.marginatulus (Geh.) Paris, Campylopus madeirensisLuisier, Campylopus marginatulus Geh.) [114]9 pusillum (Hedw.) Hampe (D. tortile (Schrad.) Brockm.)10 subulatum Hampe11 zonatum (Brid.) Kindb. (D. heteromallum var. zonatum(Brid.) Podp., D. zonatum var. scabrifolium Dixon)47 Pleuridium Rabenh., nom. cons. [115]1 acuminatum Lindb.2 subulatum (Hedw.) Rabenh. (P. alternifolium auct.non (Dicks. ex Hedw.) Rabenh.)48 Pseudephemerum (Lindb.) I.Hagen1 nitidum (Hedw.) Loeske (Pleuridium nitidum (Hedw.)Rabenh., P. axillare (Dicks.) I.Hagen)49 Rhamphidium Mitt.1 purpuratum Mitt. [116]50 Saelania Lindb.1 glaucescens (Hedw.) Broth.51 Trichodon Schimp.1 cylindricus (Hedw.) Schimp. (Ditrichum cylindricum(Hedw.) Grout, Ditrichum cylindricum var. oblongum(Lindb.) C.E.O.Jensen)Bruchiaceae Schimp.52 Bruchia Schwägr.1 flexuosa (Schwägr.) Müll.Hal. (B. trobasiana De Not.)2 vogesiaca Nestl. ex Schwägr.53 Trematodon Michx.1 ambiguus (Hedw.) Hornsch.2 brevicollis Hornsch. [32]3 laetevirens Hakelier & J.-P.Frahm4 longicollis Michx. [32]5 perssoniorum P.Allorge & Thér. ex V.Allorge [117]Rhabdoweisiaceae Limpr.54 Amphidium Schimp., nom. cons.1 lapponicum (Hedw.) Schimp.2 mougeotii (Schimp.) Schimp.3 tortuosum (Hornsch.) Cufod. (A. curvipes (Müll.Hal.)Broth.) [118]55 Arctoa Bruch & Schimp.1 anderssonii Wich. (A. fulvella var. anderssonii (Wich.) Grout)2 fulvella (Dicks.) Bruch & Schimp.3 hyperborea (Gunnerus ex Dicks.) Bruch & Schimp.,nom. cons.56 Cnestrum I.Hagen1 alpestre (Wahlenb. ex Huebener) Nyholm exMogensen (Cynodontium alpestre (Wahlenb. exHuebener) Milde)2 glaucescens (Lindb. & Arnell) Holmen ex Mogensen& Steere3 schisti (F.Weber & D.Mohr) I.Hagen, nom. cons.57 Cynodontium Bruch & Schimp., nom. cons.1 asperifolium (Lindb. ex Arnell) Paris [119]2 bruntonii (Sm.) Bruch & Schimp. (Oreoweisia bruntonii(Sm.) Milde)3 fallax Limpr.4 gracilescens (F.Weber & D.Mohr) Schimp.5 jenneri (Schimp.) Stirt.6 polycarpon (Hedw.) Schimp. [120]7 strumiferum (Hedw.) Lindb. (C. polycarpon var.strumiferum (Hedw.) Schimp.)8 suecicum (Arnell & C.E.O.Jensen) I.Hagen9 tenellum (Schimp.) Limpr. (C. torquescens Limpr.)58 Dichodontium Schimp.1 flavescens (Dicks.) Lindb. (D. pellucidum var.flavescens (Dicks.) Moore)2 palustre (Dicks.) M.Stech (Anisothecium palustre(Dicks.) I.Hagen, Dicranella palustris (Dicks.)Crundw., Diobelonella palustris (Dicks.) Ochyra) [121]

MOSSES OF EUROPE AND MACARONESIA 2093 pellucidum (Hedw.) Schimp. (D. pellucidum var.fagimontanum Brid., D. pellucidum var.propaguliferum (Correns) Casares-Gil)59 Dicranoweisia Milde [122]1 cirrata (Hedw.) Lindb.2 compacta (Schleich. ex Schwägr.) Schimp. (D. crispulavar. compacta (Schwägr.) Lindb., Hymenolomacompactum (Schwägr.) Ochyra)3 crispula (Hedw.) Milde (Hymenoloma crispulum(Hedw.) Ochyra)60 Glyphomitrium Brid.1 daviesii (Dicks.) Brid.61 Kiaeria I.Hagen1 blyttii (Bruch & Schimp.) Broth.2 falcata (Hedw.) I.Hagen3 glacialis (Berggr.) I.Hagen4 riparia (H.Lindb.) M.F.V.Corley (Dicranella riparia(H.Lindb.) Martensson & Nyholm, K. starkei var.riparia I.Hagen, Oncophorus riparius H.Lindb.)5 starkei (F.Weber & D.Mohr) I.Hagen62 Oncophorus (Brid.) Brid.1 elongatus (I.Hagen) Hedenäs (O. wahlenbergii var.elongatus I.Hagen) [123]2 virens (Hedw.) Brid.3 wahlenbergii Brid.var. compactus (Bruch & Schimp.) Braithw.(O. compactus (Bruch & Schimp.) Kindb.)var. wahlenbergii63 Oreas Brid.1 martiana (Hoppe & Hornsch.) Brid.64 Oreoweisia (Bruch & Schimp.) De Not.1 torquescens (Hornsch. ex Brid.) Wijk & Margad.(O. serrulata (Funck) De Not.)65 Rhabdoweisia Bruch & Schimp.1 crenulata (Mitt.) H.Jameson2 crispata (Dicks.) Lindb. (R. denticulata (Brid.) Bruch& Schimp., R. kusenevae Broth. [124])3 fugax (Hedw.) Bruch & Schimp. (R. striata (Schrad.)Lindb.)Schistostegaceae Schimp. [125]66 Schistostega D.Mohr1 pennata (Hedw.) F.Weber & D.MohrDicranaceae Schimp.67 Aongstroemia Bruch & Schimp., nom. cons.1 longipes (Sommerf.) Bruch & Schimp.68 Dicranella (Müll.Hal.) Schimp., nom. cons.1 campylophylla (Taylor) A.Jaeger [126]2 cerviculata (Hedw.) Schimp.3 crispa (Hedw.) Schimp. (Anisothecium vaginale(Dicks.) Loeske)4 grevilleana (Brid.) Schimp. (Anisothecium grevilleanum(Brid.) Lindb.)5 heteromalla (Hedw.) Schimp.6 howei Renauld & Cardot7 humilis R.Ruthe (Anisothecium humile (R.Ruthe)Lindb., Anisothecium rigidulum (Hedw.)C.E.O.Jensen)8 rufescens (Dicks.) Schimp. (Anisothecium rufescens(Dicks.) Lindb.)9 schreberiana (Hedw.) Dixon (Anisotheciumschreberianum (Hedw.) Dixon)10 staphylina H.Whitehouse (Anisothecium staphylinum(H.Whitehouse) Sipman, Rubers & Riemann)11 subulata (Hedw.) Schimp. (D. curvata (Hedw.)Schimp., D. secunda Lindb.)12 varia (Hedw.) Schimp. (Anisothecium rubrum Lindb.,Anisothecium varium (Hedw.) Mitt.)69 Dicranum Hedw. [127]section Dicranum1 bonjeanii De Not. (D. palustre Bruch & Schimp.,D. undulatum Turner, hom. illeg.)2 crassifolium Sérgio, Ochyra & Séneca [128]3 leioneuron Kindb.4 majus Sm.5 polysetum Sw. ex anon. (D. rugosum Brid., D. undulatumEhrh. ex F.Weber & D.Mohr, hom. illeg.) [106]6 scoparium Hedw.7 transsylvanicum Lüth [129]section Spuria Bruch & Schimp.8 acutifolium (Lindb. & Arnell) C.E.O.Jensen (D. bergerivar. acutifolium Lindb. & Arnell, D. muehlenbeckii var.acutifolium (Lindb. & Arnell) Nyholm)9 brevifolium (Lindb.) Lindb. (D. muehlenbeckii var.brevifolium Lindb., D. muehlenbeckii var. cirrhatum(Schimp.) Lindb.)10 dispersum Engelmark [130]11 drummondii Müll.Hal. (D. elatum Lindb., D. robustumBruch & Schimp.)12 spurium Hedw.13 undulatum Schrad. ex Brid. (D. affine Funck, D.bergeri Blandow) [131]section Fuscescentiformia (Kindb.) Ochyra14 flexicaule Brid. (D. congestum Brid., D. fuscescens var.congestum (Brid.) Kindb., D. fuscescens var. flexicaule(Brid.) Wilson)15 fuscescens Sm.section Convolutifolia (Kindb.) Ochyra16 angustum Lindb. [132]17 muehlenbeckii Bruch & Schimp.18 spadiceum J.E.Zetterst. (D. muehlenbeckii var. neglectum(De Not.) Pfeff., D. muehlenbeckii var. spadiceum(J.E.Zetterst.) Podp., D. neglectum Jur. ex De Not.)section Elongata I.Hagen19 elongatum Schleich. ex Schwägr. (D. elongatum var.sendtneri (Limpr.) Mönk., D. sendtneri Limpr. [133])20 fragilifolium Lindb.21 groenlandicum Brid. (D. elongatum subsp.groenlandicum (Brid.) Mönk.)22 laevidens R.S.Williams [132]section Crassinervia G.Roth23 canariense Hampe ex Müll.Hal. (D. scottianum subsp.canariense (Hampe) Corb.) [134]24 fulvum Hook. (Orthodicranum fulvum (Hook.)G.Roth ex Casares-Gil)

210 M. O. HILL ET AL.25 scottianum Turner ex R.Scott (Orthodicranum scottianum(Turner ex R.Scott) G.Roth ex Casares-Gil) [79]26 viride (Sull. & Lesq.) Lindb.section Montana Hartm.27 flagellare Hedw. (Orthodicranum flagellare (Hedw.)Loeske)28 montanum Hedw. (Orthodicranum montanum (Hedw.)Loeske)29 tauricum Sapjegin (D. strictum Schleich. ex D.Mohr,hom. illeg., Orthodicranum tauricum (Sapjegin)Smirnova)70 Paraleucobryum (Limpr.) Loeske1 enerve (Thed.) Loeske2 longifolium (Hedw.) Loeske3 sauteri (Bruch & Schimp.) Loeske (P. longifolium var.sauteri (Bruch & Schimp.) C.E.O.Jensen)Leucobryaceae Schimp.71 Atractylocarpus Mitt., nom. cons.1 alpinus (Schimp. ex Milde) Lindb. (Metzlerella alpina(Schimp. ex Milde) I.Hagen, Metzleria alpina Schimp.ex Milde)72 Campylopus Brid.1 atrovirens De Not. (C. atrovirens var. adustus (DeNot.) Husn., C. atrovirens var. falcatus Braithw., C.atrovirens var. gracilis Dixon)2 brevipilus Bruch & Schimp.3 cygneus (Hedw.) Brid. (C. setaceus Cardot)4 flaccidus Renauld & Cardot5 flexuosus (Hedw.) Brid.6 fragilis (Brid.) Bruch & Schimp.7 gracilis (Mitt.) A.Jaeger (C. schwarzii Schimp.)8 incrassatus Müll.Hal. (C. eximius Reichardt) [135]9 introflexus (Hedw.) Brid.10 oerstedianus (Müll.Hal.) Mitt. (C. mildei Limpr.)11 pilifer Brid.12 pyriformis (Schultz) Brid. (C. pyriformis var. azoricus(Mitt.) M.F.V.Corley)13 schimperi Milde (C. subulatus var. schimperi (Milde)Husn.)14 setifolius Wilson15 shawii Wilson (C. carreiroanus Cardot)16 subulatus Schimp. ex Milde73 Dicranodontium Bruch & Schimp.1 asperulum (Mitt.) Broth.2 denudatum (Brid.) E.Britton (D. denudatum var.alpinum (Schimp.) I.Hagen)3 subporodictyon Broth. (Campylopus subporodictyon(Broth.) B.H.Allen & Ireland, Dicranumsubporodictyon (Broth.) C.Gao & T.Cao) [136]4 uncinatum (Harv.) A.Jaeger (D. circinatum (Wilson)Schimp.)74 Leucobryum Hampe1 albidum (P.Beauv.) Lindb. [137]2 glaucum (Hedw.) Ångstr.3 juniperoideum (Brid.) Müll.Hal. (L. albidum auct. eur.non (P.Beauv.) Lindb., L. minus Hampe)75 Microcampylopus (Müll.Hal.) M.Fleisch.1 laevigatus (Thér.) Giese & J.-P.FrahmCalymperaceae Kindb. [138]76 Calymperes Sw. ex F.Weber1 erosum Müll.Hal. (C. sommieri Bott.)Pottiales M.Fleisch. [139]Pottiaceae Schimp., nom. cons. [140]Timmielloideae R.H.Zander77 Timmiella (De Not.) Limpr.1 anomala (Bruch & Schimp.) Limpr.2 barbuloides (Brid.) Mönk.3 flexiseta (Bruch) Limpr.Merceyoideae Broth.78 Scopelophila (Mitt.) Lindb.1 cataractae (Mitt.) Broth.2 ligulata (Spruce) SpruceTrichostomoideae (Limpr.) Broth.79 Anoectangium Schwägr., nom. cons.1 aestivum (Hedw.) Mitt. (A. angustifolium Mitt.)2 handelii Schiffn. [141]80 Aschisma Lindb.1 carniolicum (F.Weber & D.Mohr) Lindb.2 cuynetii (Bizot & R.B.Pierrot) J.Guerra & M.J.Cano(Phascum cuynetii Bizot & R.B.Pierrot)81 Ephemerum Hampe, nom. cons.1 cohaerens (Hedw.) Hampe2 hibernicum Holyoak & V.S.Bryan [142]3 minutissimum Lindb. (E. serratum var.angustifolium (Bruch & Schimp.) Bruch & Schimp.,E. serratum var.minutissimum (Lindb.) Grout) [143]4 recurvifolium (Dicks.) Boulay5 serratum (Hedw.) Hampe (E. serratum var. rutheanum(Schimp.) Jur.) [143]6 sessile (Bruch) Müll.Hal.7 spinulosum Bruch & Schimp. ex Schimp. [144]8 stellatum H.Philib.82 Eucladium Bruch & Schimp.1 verticillatum (With.) Bruch & Schimp. [145]var. angustifolium Lindb.var. verticillatum83 Gymnostomum Nees & Hornsch., nom. cons. [146]1 aeruginosum Sm.var. aeruginosumvar. obscurum J.Guerra [147]2 boreale Nyholm & Hedenäs3 calcareum Nees & Hornsch.4 lanceolatum M.J.Cano, Ros & J.Guerra [146]5 viridulum Brid.84 Gyroweisia Schimp., nom. cons.1 reflexa (Brid.) Schimp.2 tenuis (Hedw.) Schimp. (G. tenuis var. badia Limpr.)85 Hymenostylium Brid.1 recurvirostrum (Hedw.) Dixon (Gymnostomumrecurvirostrum Hedw.) [148]var. insigne (Dixon) E.B.Bartram (H. insigne (Dixon)Podp.)var. recurvirostrum

MOSSES OF EUROPE AND MACARONESIA 21186 Hyophila Brid., nom. cons.1 involuta (Hook.) A.Jaeger87 Leptobarbula Schimp.1 berica (De Not.) Schimp.88 Micromitrium Austin1 tenerum (Bruch & Schimp.) Crosby89 Molendoa Lindb.1 hornschuchiana (Hook.) Lindb. ex Limpr.(Anoectangium hornschuchianum (Hook.) Funck exHornsch.)2 schliephackei (Schlieph.) R.H.Zander (Anoectangiumschliephackei (Schlieph.) Paris)3 sendtneriana (Bruch & Schimp.) Limpr.(Anoectangium sendtnerianum Bruch & Schimp.)4 taeniatifolia Herzog (Anoectangium taeniatifolium(Herzog) M.O.Hill)5 tenuinervis Limpr. (Anoectangium tenuinerve (Limpr.)Paris)6 warburgii (Crundw. & M.O.Hill) R.H.Zander(Anoectangium warburgii Crundw. & M.O.Hill)90 Oxystegus (Broth.) Hilp.1 hibernicus (Mitt.) Hilp. (Trichostomum hibernicum(Mitt.) Dixon)2 tenuirostris (Hook. & Taylor) A.J.E.Sm.(Trichostomum tenuirostre (Hook. & Taylor) Lindb.)91 Pleurochaete Lindb. [149]1 squarrosa (Brid.) Lindb.92 Pottiopsis Blockeel & A.J.E.Sm.1 caespitosa (Brid.) Blockeel & A.J.E.Sm. (Pottiacaespitosa (Brid.) Müll.Hal., Trichostomumcaespitosum (Brid.) Jur.)93 Splachnobryum Müll.Hal.1 obtusum (Brid.) Müll.Hal. (S. delicatulum Broth.) [150]94 Tortella (Müll.Hal.) Limpr., nom. cons. [151]1 alpicola Dixon [152]2 bambergeri (Schimp.) Broth. (T. tortuosa var.bambergeri (Schimp.) Düll) [153]3 flavovirens (Bruch) Broth. (T. flavovirens var. minorLindb.)var. flavovirens (T. flavovirens var. viridiflava (DeNot.) Casares-Gil)var. glareicola (T.A.Chr.) Crundw. & Nyholm (T.glareicola T.A.Chr.)var. papillosissima Sérgio & Casas4 fragilis (Hook. & Wilson) Limpr.5 humilis (Hedw.) Jenn.6 inclinata (R.Hedw.) Limpr.var. densa (Lorentz & Molendo) Limpr. (T. densa(Lorentz & Molendo) Crundw. & Nyholm) [154]var. inclinata7 inflexa (Bruch) Broth.8 limbata (Schiffn.) Geh. & Herzog [155]9 limosella (Stirt.) P.W.Richards & E.C.Wallace [156]10 nitida (Lindb.) Broth. (T. cirrifolia (Mitt.) Broth.)[157]11 rigens Alberts.12 tortuosa (Hedw.) Limpr.var. fragilifolia (Jur.) Limpr.var. tortuosa95 Trichostomum Bruch, nom. cons. [158]1 arcticum Kaal.2 brachydontium Bruch (Hyophila treleasei Cardot [159])3 crispulum Bruch4 triumphans De Not. (T. pallidisetum H.Müll., Weissiatriumphans (De Not.) M.O.Hill)96 Weissia Hedw. (Astomum Hampe) [160] [161]1 brachycarpa (Nees & Hornsch.) Jur. (Hymenostomummicrostomum (Hedw.) R.Br. ex Nees & Hornsch.)2 condensa (Voit) Lindb.var. armata (Thér. & Trab.) M.J.Cano, Ros &J.Guerra (W. papillosissima Laz.)var. condensa3 controversa Hedw.var. controversavar. crispata (Nees & Hornsch.) Nyholm (W. fallaxSehlm.)var. densifolia (Bruch & Schimp.) Wilson4 levieri (Limpr.) Kindb. (Astomum levieri Limpr.)5 longifolia Mitt. (Astomum crispum (Hedw.) Hampe)6 multicapsularis (Sm.) Mitt. (Astomum multicapsulare(Sm.) Bruch & Schimp.)7 perssonii Kindb.8 rostellata (Brid.) Lindb.9 rutilans (Hedw.) Lindb.10 squarrosa (Nees & Hornsch.) Müll.Hal.(Hymenostomum squarrosum Nees & Hornsch.)11 sterilis W.E.Nicholson12 tyrrhena M.Fleisch.13 wimmeriana (Sendtn.) Bruch & Schimp.(W. controversa var. wimmeriana (Sendtn.) Blockeel &A.J.E.Sm., W. wimmeriana subsp. pallescens (Schimp.ex Besch.) Giacom.) [162]Pottioideae (Limpr.) Broth.97 Acaulon Müll.Hal.1 casasianum Brugués & H.A.Crum2 dertosense Casas, Sérgio, Cros & Brugués3 fontiquerianum Casas & Sérgio4 mediterraneum Limpr. (A. muticum var. mediterraneum(Limpr.) Sérgio) [163]5 muticum (Hedw.) Müll.Hal.6 piligerum (De Not.) Limpr.7 triquetrum (Spruce) Müll.Hal.98 Aloina Kindb., nom. cons.1 aloides (Koch ex Schultz) Kindb.2 ambigua (Bruch & Schimp.) Limpr. (A. aloides var.ambigua (Bruch & Schimp.) E.J.Craig)3 bifrons (De Not.) Delgad.4 brevirostris (Hook. & Grev.) Kindb.5 humilis M.T.Gallego, M.J.Cano & Ros [164]6 obliquifolia (Müll.Hal.) Broth. (A. rigida var.mucronulata (Bruch & Schimp.) Limpr., A. rigida var.obliquifolia (Müll.Hal.) Delgad.) [165]7 rigida (Hedw.) Limpr.99 Barbula Hedw., nom. cons. [139] [166]

212 M. O. HILL ET AL.1 bicolor (Bruch & Schimp.) Lindb.2 bolleana (Müll.Hal.) Broth. (B. ehrenbergii (Lorentz)M.Fleisch.)3 convoluta Hedw. (Streblotrichum convolutum (Hedw.)P.Beauv.)var. convolutavar. sardoa Schimp. (B. commutata Jur., B. convolutavar. commutata (Jur.) Husn., B. convoluta var.uliginosa (Limpr.) Limpr., B. sardoa (Schimp.)J.-P.Frahm, nom. inval.) [167]4 crocea (Brid.) F.Weber & D.Mohr5 enderesii Garov.6 indica (Hook.) Spreng.7 unguiculata Hedw.100 Bryoerythrophyllum P.C.Chen1 alpigenum (Venturi) P.C.Chen (B. recurvirostrum var.dentatum (Schimp.) H.A.Crum, Steere &L.E.Anderson)2 caledonicum D.G.Long3 campylocarpum (Müll.Hal.) H.A.Crum (B. lusitanicum(Cardot & Dixon) M.O.Hill, B. machadoanum (Sérgio)M.O.Hill, Hyophila lusitanica Cardot & Dixon,Hyophila machadoana Sérgio) [168]4 ferruginascens (Stirt.) Giacom.5 inaequalifolium (Taylor) R.H.Zander6 recurvirostrum (Hedw.) P.C.Chen7 rubrum (Jur. ex Geh.) P.C.Chen101 Cinclidotus P.Beauv., nom. cons. [169]1 aquaticus (Hedw.) Bruch & Schimp.2 confertus Lüth [170]3 danubicus Schiffn. & Baumgartner4 fontinaloides (Hedw.) P.Beauv.5 pachylomoides Bizot6 riparius (Host ex Brid.) Arn.7 vivesii Ederra [171]102 Crossidium Jur., nom. cons.1 aberrans Holz. & E.B.Bartram2 crassinerve (De Not.) Jur.3 davidai Catches. [172]4 geheebii (Broth.) Broth. [173]5 laevipilum Thér. & Trab. [174]6 laxefilamentosum W.Frey & Kürschner [175]7 squamiferum (Viv.) Jur.var. pottioideum (De Not.) Mönk.var. squamiferum103 Dialytrichia (Schimp.) Limpr.1 fragilifolia (Bizot & J.Roux) F.Lara (D. mucronatavar. fragilifolia Bizot & J.Roux) [176]2 mucronata (Brid.) Broth. (Cinclidotus mucronatus(Brid.) Guim.)104 Didymodon Hedw. [177] [178] [179]1 acutus (Brid.) K.Saito (Barbula acuta (Brid.) Brid.)2 asperifolius (Mitt.) H.A.Crum, Steere & L.E.Anderson(Barbula asperifolia Mitt., Barbula kneuckeri Loeske &Osterwald, D. ferrugineus var. kneuckeri (Loeske &Osterwald) Düll, D. rufus Lorentz) [180]3 australasiae (Hook. & Grev.) R.H.Zander (D. aaronis(Lorentz) J.Guerra, D. incrassatus (Lindb.) Broth.,Trichostomopsis aaronis (Lorentz) S.Agnew &C.C.Towns., Trichostomopsis australasiae (Hook. &Grev.) H.Rob.) [181]4 bistratosus Hébr. & R.B.Pierrot [182]5 brachyphyllus (Sull.) R.H.Zander [183]6 cordatus Jur. (Barbula cordata (Jur.) Loeske)7 erosus J.A.Jiménez & J.Guerra [184]8 fallax (Hedw.) R.H.Zander (Barbula adriaticaBaumgartner, Barbula fallax Hedw.) [180]9 ferrugineus (Schimp. ex Besch.) M.O.Hill (Barbulareflexa (Brid.) Brid.)10 giganteus (Funck) Jur. (Barbula gigantea Funck,Geheebia gigantea (Funck) Boulay) [180]11 glaucus Ryan (Barbula rigidula subsp. verbana(W.E.Nicholson & Dixon) Podp., Barbula rigidulavar. glauca (Ryan) J.J.Amann)12 icmadophilus (Schimp. ex Müll.Hal.) K.Saito(Barbula acuta var. icmadophila (Schimp. exMüll.Hal.) H.A.Crum)13 insulanus (De Not.) M.O.Hill (Barbula cylindrica(Taylor) Schimp., Barbula vinealis var. cylindrica(Taylor) Boulay, D. vinealis var. flaccidus (Bruch &Schimp.) R.H.Zander)14 johansenii (R.S.Williams) H.A.Crum (Barbulajohansenii R.S.Williams)15 lamyanus (Schimp.) Thér. [185]16 luridus Hornsch. (Barbula lurida Hornsch., D. trifariusauct. non (Hedw.) Röhl.)17 maximus (Syed & Crundw.) M.O.Hill (Barbula reflexavar. robusta Braithw.) [180]18 nicholsonii Culm.19 rigidulus Hedw. (Barbula rigidula (Hedw.) Milde, D.mamillosus (Crundw.) M.O.Hill) [186]20 sicculus M.J.Cano, Ros, García-Zamora & J.Guerra[187]21 sinuosus (Mitt.) Delogne (Barbula sinuosa (Mitt.)Grav.)22 spadiceus (Mitt.) Limpr. (Barbula insidiosa Jur. &Milde, Barbula spadicea (Mitt.) Braithw., D.barbuloides Lib. ex Marchal, D. zetterstedtii Schimp.,Limneria viridula Stirt.) [180]23 subandreaeoides (Kindb.) R.H.Zander (D. rigidulussubsp. andreaeoides (Limpr.) Wijk & Margad.) [188]24 tomaculosus (Blockeel) M.F.V.Corley25 tophaceus (Brid.) Lisa (Barbula tophacea (Brid.) Mitt.,D. bosniacus Głow., D. spadiceus var. siluricus Velen.)[180]26 umbrosus (Müll.Hal.) R.H.Zander (D. australasiaevar. umbrosus (Müll.Hal.) R.H.Zander, D. trivialis(Müll.Hal.) J.Guerra, Trichostomopsis trivialis(Müll.Hal.) H.Rob., Trichostomopsis umbrosa(Müll.Hal.) H.Rob.) [189]27 vinealis (Brid.) R.H.Zander (Barbula vinealis Brid.)105 Hennediella Paris

MOSSES OF EUROPE AND MACARONESIA 2131 heimii (Hedw.) R.H.Zander (Desmatodon heimii(Hedw.) Mitt., Pottia heimii (Hedw.) Hampe)var. arctica (Lindb.) R.H.Zander (Desmatodon heimiivar. arcticum (Lindb.) H.A.Crum)var. heimii2 macrophylla (R.Br.bis) Paris [190]3 stanfordensis (Steere) Blockeel106 Hilpertia R.H.Zander1 velenovskyi (Schiffn.) R.H.Zander107 Leptodontium (Müll.Hal.) Lindb.1 flexifolium (Dicks.) Hampe2 gemmascens (Mitt.) Braithw.3 styriacum (Jur.) Limpr.108 Leptophascum (Müll.Hal.) J.Guerra & M.J.Cano1 leptophyllum (Müll.Hal.) J.Guerra & M.J.Cano(Chenia leptophylla (Müll.Hal.) R.H.Zander, Cheniarhizophylla (Sakurai) R.H.Zander, Phascumleptophyllum Müll.Hal., Tortula rhizophylla(Sakurai) Z.Iwats. & K.Saito)109 Microbryum Schimp. [191]1 curvicollum (Hedw.) R.H.Zander (M. piptocarpum(Durieu & Mont.) J.Guerra & M.J.Cano, Phascumcurvicollum Hedw., Phascum piptocarpum Durieu &Mont.) [32] [19]2 davallianum (Sm.) R.H.Zander (Pottia commutataLimpr., Pottia conica (Schleich. ex Schwägr.) Fürnr.ex Paris, Pottia davalliana (Sm.) C.E.O.Jensen, Pottiastarckeana subsp. conica (Schleich. ex Schwägr.)D.F.Chamb., Pottia starckeana subsp. minutula(Schwägr.) D.F.Chamb., Pottia starckeana var.minutula (Schwägr.) Corb.) [193]3 floerkeanum (F.Weber & D.Mohr) Schimp. (Phascumfloerkeanum F.Weber & D.Mohr)4 fosbergii (E.B.Bartram) Ros, O.Werner & Rams (M.starckeanum var. fosbergii (E.B.Bartram) R.H.Zander,Pottia 6andalusica Ros & R.Oliva) [194]5 longipes (J.Guerra, J.J.Martínez & Ros) R.H.Zander(Phascum longipes J.Guerra, J.J.Martínez & Ros)6 rectum (With.) R.H.Zander (Pottia recta (With.) Mitt.)7 starckeanum (Hedw.) R.H.Zander (Pottia muticaVenturi, Pottia starckeana (Hedw.) Müll.Hal., Pottiastarckeana var. brachyodus (Bruch & Schimp.)Müll.Hal.) [193] [195]110 Paraleptodontium D.G.Long1 recurvifolium (Taylor) D.G.Long (Trichostomumrecurvifolium (Taylor) R.H.Zander)111 Phascum Hedw.1 cuspidatum Hedw. (Tortula acaulon (With.)R.H.Zander, Tortula atherodes R.H.Zander)var. cuspidatum (P. cuspidatum var. curvisetum(Dicks.) Nees & Hornsch., P. cuspidatum var.mitraeforme Limpr., Tortula atherodes var. curviseta(Dicks.) R.H.Zander)var. papillosum (Lindb.) G.Roth (P. cuspidatumsubsp. papillosum (Lindb.) J.Guerra & Ros, Tortulaacaulon var. papillosa (Lindb.) R.H.Zander, Tortulaatherodes var. papillosa R.H.Zander)var. piliferum (Hedw.) Hook. & Taylor (Tortulaacaulon var. pilifera (Hedw.) R.H.Zander Tortulaatherodes var. pilifera (Hedw.) R.H.Zander)var. retortifolium J.Guerra & Rosvar. schreberianum (Dicks.) Brid. (Tortula acaulon var.schreberiana (Dicks.) R.H.Zander, Tortula atherodesvar. schreberiana (Dicks.) R.H.Zander)2 vlassovii Laz. (Microbryum vlassovii (Laz.)R.H.Zander)112 Protobryum J.Guerra & M.J.Cano1 bryoides (Dicks.) J.Guerra & M.J.Cano (Pottiabryoides (Dicks.) Mitt., Tortula protobryoidesR.H.Zander)113 Pseudocrossidium R.S.Williams1 hornschuchianum (Schultz) R.H.Zander (Barbulahornschuchiana Schultz)2 obtusulum (Lindb.) H.A.Crum & L.E.Anderson(P. revolutum var. obtusulum (Lindb.) B.C.Tan,R.H.Zander & Taylor,Terry) [196]3 replicatum (Taylor) R.H.Zander [197]4 revolutum (Brid.) R.H.Zander (Barbula revoluta Brid.)114 Pterygoneurum Jur., nom. cons.1 compactum M.J.Cano, J.Guerra & Ros [198]2 crossidioides W.Frey, Herrnst. & Kürschner [199]3 kozlovii Laz.4 lamellatum (Lindb.) Jur.5 ovatum (Hedw.) Dixon (P. ovatum var. incanum Jur.)6 papillosum Oesau [200]7 sampaianum (Guim.) Guim.8 squamosum Segarra & Kürschner [201]9 subsessile (Brid.) Jur.115 Stegonia Venturi1 latifolia (Schwägr.) Venturi ex Broth. (S. latifolia var.latifolia, S. latifolia var. pilifera (Brid.) Broth.)116 Syntrichia Brid. [202]1 bogotensis (Hampe) R.H.Zander [203]2 calcicola J.J.Amann (S. densa (Velen.) J.-P.Frahm,Tortula calcicolens W.A.Kramer, Tortula densa (Velen.)J.-P.Frahm, Tortula ruralis var. calcicola (J.J.Amann)Barkman, Tortula ruralis var. densa Velen.) [204]3 caninervis Mitt. (Tortula caninervis (Mitt.) Broth.) [205]var. abranchesii (Luisier) R.H.Zander (S. abranchesii(Luisier) Ochyra, Tortula abranchesii Luisier) [206]var. astrakhanica Ignatov, Ignatova & Suragina [207]var. caninervisvar. gypsophila (J.J.Amann ex G.Roth) Ochyra (S.caninervis var. spuria (J.J.Amann) R.H.Zander, S.ruralis var. gypsophila (J.J.Amann ex G.Roth)J.J.Amann, S. ruralis var. spuria (J.J.Amann) Podp.,Tortula caninervis subsp. spuria (J.J.Amann)W.A.Kramer, Tortula caninervis subsp. spuria var.gypsophila (J.J.Amann ex G.Roth) W.A.Kramer,Tortula ruralis var. gypsophila J.J.Amann ex G.Roth,Tortula spuria J.J.Amann)4 echinata (Schiffn.) Herrnst. & Ben-Sasson (S. princepssubsp. echinata (Schiffn.) Podp., S. princeps var.

214 M. O. HILL ET AL.echinata (Schiffn.) R.H.Zander, Tortula echinataSchiffn.) [202]5 fragilis (Taylor) Ochyra (Barbula alpina var. inermisMilde, S. mutica Giacom., Tortula fragilis Taylor)6 glabra J.-P.Frahm & M.T.Gallego [208]7 handelii (Schiffn.) S.Agnew & Vondr. (S. montanasubsp. handelii (Schiffn.) Podp., Tortula handeliiSchiffn., Tortula intermedia subsp. handelii (Schiffn.)Wijk & Margad.)8 laevipila Brid. (S. laevipila var. laevipilaeformis (DeNot.) J.J.Amann, S. pagorum (Milde) J.J.Amann,Tortula laevipila (Brid.) Schwägr., Tortula laevipilavar. meridionalis (Schimp.) Wijk & Margad., Tortulalaevipila var. notarisii Barkman, Tortula laevipila var.wachteri Barkman, Tortula laevipilaeformis De Not.,Tortula pagorum (Milde) De Not., Tortulasaccardoana De Not.) [209]9 latifolia (Bruch ex Hartm.) Huebener (Tortula latifoliaBruch ex Hartm.)10 minor (Bizot) M.T.Gallego, J.Guerra, M.J.Cano, Ros&Sánchez-Moya [210]11 montana Nees (S. intermedia Brid., Tortula crinita (DeNot.) De Not., Tortula intermedia (Brid.) Berk., hom.illeg. [211])var. calva (Durieu & Sagot ex Bruch & Schimp.)J.J.Amann (Tortula crinita var. calva (Durieu & Sagotex Bruch & Schimp.) Nebel & Heinrichs, Tortularuralis var. calva (Durieu & Sagot ex Bruch &Schimp.) C.Hartm.)var. montana12 norvegica F.Weber (Tortula norvegica (F.Weber)Lindb.)13 papillosa (Wilson) Jur. (Tortula papillosa var.meridionalis Warnst. [212], Tortula papillosaWilson)14 papillosissima (Copp.) Loeske (S. ruralis var. hirsuta(Venturi) Podp., Tortula hirsuta (Venturi) Laz.,Tortula papillosissima (Copp.) Broth., Tortula ruralissubsp. hirsuta (Venturi) W.A.Kramer, Tortula ruralisvar. hirsuta (Venturi) Paris) [202]15 princeps (De Not.) Mitt. (Tortula princeps De Not.)[213]16 rigescens (Broth. & Geh.) Ochyra (Tortula rigescensBroth. & Geh.) [202]17 ruralis (Hedw.) F.Weber & D.Mohr (Tortula ruralis(Hedw.) P.Gaertn., B.Mey. & Scherb.)var. ruraliformis (Besch.) Delogne (S. ruraliformis(Besch.) Cardot, S. ruralis var. arenicola J.J.Amann,Tortula ruraliformis (Besch.) Ingham, Tortula ruralisvar. ruraliformis (Besch.) De Wild.) [214]var. ruralis (S. ruralis var. glacialis J.J.Amann)18 sinensis (Müll.Hal.) Ochyra (Tortula sinensis(Müll.Hal.) Broth.)19 subpapillosissima (Bizot & R.B.Pierrot exW.A.Kramer) M.T.Gallego & J.Guerra (Tortulapapillosissima var. submamillosa (W.A.Kramer)Heinrichs & Caspari, Tortula ruraliformis var.subpapillosissima Bizot & R.B.Pierrot exW.A.Kramer, Tortula ruralis var. submamillosaW.A.Kramer) [215]20 virescens (De Not.) Ochyra (Tortula virescens (DeNot.) De Not.)117 Tortula Hedw., nom. cons. (Desmatodon Brid., PottiaEhrh. ex Fürnr.) [216] [217]1 amplexa (Lesq.) Steere (Syntrichia amplexa (Lesq.)R.H.Zander)2 ampliretis Crundw. & D.G.Long [218]3 atrovirens (Sm.) Lindb.4 bogosica (Müll.Hal.) R.H.Zander (Desmatodonbogosicus Müll.Hal.)5 bolanderi (Lesq. & James) M.Howe (Syntrichiabolanderi (Lesq. & James) R.H.Zander) [219]6 brevissima Schiffn. [221]7 canescens Mont.8 cernua (Huebener) Lindb. (Desmatodon cernuus(Huebener) Bruch & Schimp.)9 cuneifolia (Dicks.) Turner10 freibergii Dixon & Loeske11 guepinii (Bruch & Schimp.) Broth. (Desmatodonguepinii Bruch & Schimp.)12 hoppeana (Schultz) Ochyra (Desmatodon latifolius(Hedw.) Brid., T. eucalyptrata Lindb., T. euryphyllaR.H.Zander) [222]13 inermis (Brid.) Mont. (Syntrichia inermis (Brid.)Bruch) [219]14 israelis Bizot & F.Bilewsky (T. baetica (Casas &R.Oliva) J.Guerra & Ros, T. muralis var. baeticaCasas & R.Oliva) [220]15 lanceola R.H.Zander (Pottia lanceolata (Hedw.)Müll.Hal.)16 laureri (Schultz) Lindb. (Desmatodon laureri (Schultz)Bruch & Schimp.)17 leucostoma (R.Br.) Hook. & Grev. (Desmatodonleucostoma (R.Br.) Berggr.)18 lingulata Lindb.19 marginata (Bruch & Schimp.) Spruce (Desmatodonmeridionalis Luisier) [223]20 modica R.H.Zander (Pottia intermedia (Turner)Fürnr.)21 mucronifolia Schwägr.22 muralis Hedw.23 obtusifolia (Schwägr.) Mathieu24 pallida (Lindb.) R.H.Zander (Pottia pallida Lindb.,P. cuneifolia Solms ex Schimp., T. zoddaeR.H.Zander) [224]25 randii (Kenn.) R.H.Zander (Desmatodon oxneri Laz.,Desmatodon randii (Kenn.) Laz.)26 revolvens (Schimp.) G.Roth27 rhodonia R.H.Zander (Desmatodon wilczekii Meyl.)[225]28 schimperi M.J.Cano, O.Werner & J.Guerra(T. angustata Lindb., hom. illeg., T. subulata var.angustata (Schimp.) Limpr.) [226]

MOSSES OF EUROPE AND MACARONESIA 21529 solmsii (Schimp.) Limpr. (T. marginata subsp. limbata(Lindb.) Podp.) [227]30 subulata Hedw. (T. subulata var. graeffii Warnst.,T. subulata var. subinermis (Bruch & Schimp.) Wilson)[226]31 systylia (Schimp.) Lindb. (Desmatodon systyliusSchimp.)32 truncata (Hedw.) Mitt. (Pottia truncata (Hedw.) Bruch& Schimp.)33 ucrainica (Laz.) R.H.Zander (Desmatodon ucrainicusLaz.)34 vahliana (Schultz) Mont.35 viridifolia (Mitt.) Blockeel & A.J.E.Sm. (Pottia crinitaBruch & Schimp.)36 wilsonii (Hook.) R.H.Zander (Pottia wilsonii (Hook.)Bruch & Schimp.)118 Triquetrella Müll.Hal.1 arapilensis LuisierSplachnales (M.Fleisch.) OchyraSplachnaceae Grev. & Arn.119 Aplodon R.Br.1 wormskioldii (Hornem.) R.Br. [32]120 Splachnum Hedw.1 ampullaceum Hedw.2 luteum Hedw.3 melanocaulon (Wahlenb.) Schwägr.4 pensylvanicum (Brid.) Grout ex H.A.Crum5 rubrum Hedw.6 sphaericum Hedw.7 vasculosum Hedw.121 Tayloria Hook.1 acuminata Hornsch.2 froelichiana (Hedw.) Mitt. ex Broth.3 hornschuchii (Grev. & Arn.) Broth.4 lingulata (Dicks.) Lindb.5 rudolphiana (Garov.) Bruch & Schimp.6 serrata (Hedw.) Bruch & Schimp.7 splachnoides (Schleich. ex Schwägr.) Hook.8 tenuis (Dicks.) Schimp. (T. serrata var. tenuis (Dicks.)Bruch & Schimp.)122 Tetraplodon Bruch & Schimp.1 angustatus (Hedw.) Bruch & Schimp.2 blyttii Frisvoll3 mnioides (Hedw.) Bruch & Schimp.4 pallidus I.Hagen5 paradoxus (R.Br.) I.Hagen6 urceolatus (Hedw.) Bruch & Schimp. [228]123 Voitia Hornsch.1 hyperborea Grev. & Arn.2 nivalis Hornsch.Meesiaceae Schimp.124 Amblyodon P.Beauv., nom. cons.1 dealbatus (Hedw.) P.Beauv.125 Leptobryum (Bruch & Schimp.) Wilson1 pyriforme (Hedw.) Wilson126 Meesia Hedw., nom. cons.1 hexasticha (Funck) Bruch2 longiseta Hedw.3 triquetra (L. ex Jolycl.) Ångstr.4 uliginosa Hedw.127 Paludella Brid.1 squarrosa (Hedw.) Brid.Orthotrichales DixonOrthotrichaceae Arn.128 Orthotrichum Hedw.Subgenus Orthotrichum [229]1 anomalum Hedw.2 cupulatum Hoffm. ex Brid.var. bistratosum Schiffn.var. cupulatumvar. fuscum (Venturi) Boulay (O. limprichtii I.Hagen)[230]var. riparium Huebener (O. cupulatum var. nudum(Dicks.) Braithw.)3 pellucidum Lindb.4 urnigerum MyrinSubgenus Pulchella (Schimp.) Vitt5 alpestre Bruch & Schimp.6 casasianum F.Lara, Garilleti & Mazimpaka [231]7 consimile Mitt.8 crenulatum Mitt. [231] [232]9 diaphanum Schrad. ex Brid.10 handiense F.Lara, Garilleti & Mazimpaka [233]11 hispanicum F.Lara, Garilleti & Mazimpaka [234]12 macrocephalum F.Lara, Garilleti & Mazimpaka [235]13 microcarpum De Not.14 pallens Bruch ex Brid. (O. paradoxum Gronvall)15 patens Bruch ex Brid.16 philibertii Venturi17 pulchellum Brunt.18 pumilum Sw. ex anon. [106]19 rivulare Turner20 rogeri Brid.21 scanicum Gronvall (O. lewinskyae F.Lara, Garilleti &Mazimpaka) [236]22 schimperi Hammar [237]23 sprucei Mont.24 stellatum Brid.25 stramineum Hornsch. ex Brid. (O. rogeri var. defluens(Venturi) Venturi) [238]26 tenellum Bruch ex Brid. (O. australe Jur.) [239]27 vittii F.Lara, Garilleti & Mazimpaka [240]Subgenus Callistoma (Z.Iwats. & Sharp) Lewinsky28 callistomum Fisch.-Oost. ex Bruch & Schimp.Subgenus Orthophyllum Delogne29 gymnostomum Bruch ex Brid.30 obtusifolium Brid.Subgenus Phaneroporum Delogne31 laevigatum J.E.Zetterst. (O. arcangelianum Massari)[241]32 rupestre Schleich. ex Schwägr.33 shawii Wilson [242]Subgenus Gymnoporus (Braithw.) Limpr.34 acuminatum H.Philib.

216 M. O. HILL ET AL.35 affine Schrad. ex Brid. (O. fastigiatum Bruch ex Brid.)36 ibericum F.Lara & Mazimpaka [243]37 lyellii Hook. & Taylor38 pylaisii Brid.39 sordidum Sull. & Lesq. (O. caucasicum Venturi) [244]40 speciosum Nees (O. elegans auct. eur. non Schwägr.)[245]var. brevisetum F.Lara, Garilleti & Mazimpaka [246]var. speciosum41 striatum Hedw.42 tortidontium F.Lara, Garilleti & Mazimpaka [247]43 vladikavkanum Venturi [248]129 Ulota D.Mohr1 bruchii Hornsch. ex Brid. (U. crispa var. norvegica(Gronvall) A.J.E.Sm. & M.O.Hill)2 calvescens Wilson3 coarctata (P.Beauv.) Hammar4 crispa (Hedw.) Brid. (U. intermedia Schimp.)5 curvifolia (Wahlenb.) Lilj.6 drummondii (Hook. & Grev.) Brid.7 hutchinsiae (Sm.) Hammar8 macrospora E.Bauer & Warnst. [249]9 phyllantha Brid.10 rehmannii Jur.130 Zygodon Hook. & Taylor1 conoideus (Dicks.) Hook. & Taylorvar. conoideusvar. lingulatus S.R.Edwards2 dentatus (Limpr.) Kartt. (Z. viridissimus var. dentatusLimpr.)3 forsteri (Dicks.) Mitt. (Codonoblepharon forsteri(Dicks.) Goffinet, Zygodon madeirensis Dixon &Luisier) [250]4 gracilis Wilson5 rupestris Schimp. ex Lorentz (Z. baumgartneri Malta,Z. viridissimus var. rupestris C.Hartm., Z. vulgarisNyholm)6 sibiricus Ignatov, Ignatova, Z.Iwats. & B.C.Tan [251]7 stirtonii Schimp. ex Stirt. [252]8 viridissimus (Dicks.) Brid.Hedwigiales OchyraHedwigiaceae Schimp.131 Braunia Bruch & Schimp.1 alopecura (Brid.) Limpr.132 Hedwigia P.Beauv., nom. cons.1 ciliata (Hedw.) P.Beauv. [253]var. ciliatavar. leucophaea Bruch & Schimp. [254]2 integrifolia P.Beauv. (Hedwigidium integrifolium(P.Beauv.) Dixon)3 stellata Hedenäs [255]Bryales Limpr.Catoscopiaceae Boulay ex Broth.133 Catoscopium Brid.1 nigritum (Hedw.) Brid.Bartramiaceae Schwägr.134 Anacolia Schimp., nom. cons.1 laevisphaera (Taylor) Flowers [256]2 menziesii (Turner) Paris [257]3 webbii (Mont.) Schimp.135 Bartramia Hedw., nom. cons.section Bartramia1 halleriana Hedw.2 pomiformis Hedw. [258]section Pyridium Müll.Hal. [259]3 breviseta Lindb. (B. ithyphylla var. breviseta (Lindb.)Kindb.) [260]4 ithyphylla Brid. [261]5 subulata Bruch & Schimp.section Strictidium Müll.Hal.6 stricta Brid.136 Breutelia (Bruch & Schimp.) Schimp.1 azorica (Mitt.) Cardot2 chrysocoma (Hedw.) Lindb.137 Conostomum Sw. ex F.Weber & D.Mohr1 tetragonum (Hedw.) Lindb.138 Philonotis Brid.section Bartramidula (Bruch & Schimp.) Mitt.1 cernua (Wilson) D.G.Griffin & W.R.Bucksection Philonotula (Schimp.) A.Jaeger2 rigida Brid.section Homomorphae (Kindb.) Ochyra3 arnellii Husn. (P. capillaris auct. non Lindb.)4 hastata (Duby) Wijk & Margad.5 marchica (Hedw.) Brid. (P. marchica var. laxa(Limpr.) Loeske & Warnst. [262])6 uncinata (Schwägr.) Brid. [263]section Philonotis7 caespitosa Jur.8 calcarea (Bruch & Schimp.) Schimp.9 fontana (Hedw.) Brid.10 seriata Mitt.11 tomentella Molendo (P. fontana var. pumila (Turner)Brid.)139 Plagiopus Brid.1 oederianus (Sw.) H.A.Crum & L.E.Anderson(P. oederi (Brid.) Limpr.)var. alpinus (Schwägr.) Ochyravar. oederianusBryaceae Schwägr. [264]140 Anomobryum Schimp.1 concinnatum (Spruce) Lindb. (A. filiforme subsp.concinnatum (Spruce) J.J.Amann, A. julaceum var.concinnatum (Spruce) J.E.Zetterst., A. leptostomoidesSchimp., Bryum concinnatum Spruce, Bryum filiformevar. concinnatum (Spruce) Boulay, Bryum julaceumvar. concinnatum (Spruce) Wilson) [265]2 julaceum (Schrad. ex P.Gaertn. et al.) Schimp. (A.filiforme (Dicks.) Husn., hom. illeg., A. filiforme var.juliforme (Solms) Husn., A. juliforme Solms, Bryumfiliforme Dicks., Bryum julaceum Schrad. ex P.Gaertn.et al., Bryum juliforme (Solms) Schimp., Bryumleptostomum Schimp.)3 lusitanicum (I.Hagen ex Luisier) Thér. [266]

MOSSES OF EUROPE AND MACARONESIA 217141 Brachymenium Schwägr.1 commutatum (Müll.Hal.) A.Jaeger [267]2 notarisii (Mitt.) A.J.Shaw (Haplodontium notarisii(Mitt.) Broth.) [268]3 paradoxum (Herzog) A.J.Shaw (Mielichhoferiaparadoxa Herzog) [269]4 philonotula Broth. [270]142 Bryum Hedw. (Imbribryum N.Pedersen) [271] [272][273] [274] [275]1 algovicum Sendtn. ex Müll.Hal. [276]var. algovicum (B. algovicum var. compactum(Hornsch.) Düll, B. compactum (Hornsch.) Kindb.)var. rutheanum (Warnst.) Crundw.2 alpinum Huds. ex With. (Imbribryum alpinum (Huds.ex With.) N.Pedersen)3 apiculatum Schwägr.(B. cruegeri Hampe, B. nitens Hook.,Pohlia apiculata (Schwägr.) H.A.Crum & L.E.Anderson,Pohlia cruegeri (Hampe) A.L.Andrews) [277]4 archangelicum Bruch & Schimp. (B. amblyodonMüll.Hal., B. curvatum Kaurin & Arnell, B.imbricatum auct. non? (Schwägr.) Bruch & Schimp.,B. inclinatum (Brid.) Turton, hom. illeg. non (Hedw.)Dicks., B. stenotrichum Müll.Hal.) [278]5 arcticum (R.Br.) Bruch & Schimp. (B. bryoides (R.Br.)Wijk & Margad., B. lawersianum H.Philib., B.purpurascens (R.Br.) Bruch & Schimp.) [278]6 argenteum Hedw. (Anomobryum lanatum (P.Beauv.)J.R.Spence & H.P.Ramsay, B. argenteum subsp.veronense (De Not.) J.J.Amann, B. argenteum var.lanatum (P.Beauv.) Hampe [280], B. argenteum var.veronense (De Not.) Molendo, B. lanatum (P.Beauv.)Brid., B. veronense De Not.) [279]7 blindii Bruch & Schimp. [281]8 bornholmense Wink. & R.Ruthe [282]9 caespiticium Hedw. (B. badium (Brid.) Schimp., B.comense Schimp.) [283]10 calophyllum R.Br. (B. acutiforme Limpr., B.axel-blyttii Kaurin ex H.Philib.) [278]11 canariense Brid. (B. provinciale H.Philib.) [284]12 capillare Hedw. (B. platyloma Schwägr.,B. rufifolium(Dixon) Demaret & R.Wilczek, B. validicostatumCardot & Dixon, Rosulabryum capillare (Hedw.)J.R.Spence) [278]13 caucasicum (Schimp. ex Broth.) C.J.Cox & Hedd.(Mielichhoferia caucasia Schimp. ex Broth.,Mielichhoferia himalayana Mitt. non Bryumhimalayanum (Mitt.) Müll.Hal.) [285]14 cellulare Hook. (Brachymenium cellulare (Hook.) A.Jaeger,B. splachnoides (Harv.) Müll.Hal. non Dicks.) [269]15 creberrimum Taylor (B. affine F.W.Schultz nonJ.F.Gmel. ex Broth., B. cuspidatum (Bruch & Schimp.)Schimp., B. lisae De Not.)16 cryophilum Martensson (B. cyclophyllum var.obtusifolium Lindb., B. obtusifolium Lindb.,hom. illeg.)17 cyclophyllum (Schwägr.) Bruch & Schimp.(B. tortifolium Funck ex Brid.)18 demaretianum Arts [286]19 dichotomum Hedw. (B. balticum Nyholm & Hedenäs,B. barnesii J.B.Wood ex Schimp., B. bicolor Dicks.,B. dunense A.J.E.Sm. & H.Whitehouse, B. excurrensLindb., B. versicolor A.Braun ex Bruch & Schimp.) [287]20 dixonii Cardot ex W.E.Nicholson (B. bicolor subsp.dixonii (Cardot ex W.E.Nicholson) Podp.)21 donianum Grev. (B. obovatum Mitt., B. pachylomaCardot)22 dyffrynense Holyoak [287]23 elegans Nees (B. capillare var. elegans (Nees) Husn., B.capillare var. ferchelii (Funck ex Brid.) Bruch &Schimp., B. chibinense Schljakov, B. stirtonii Schimp.,Rosulabryum elegans (Nees) Ochyra) [278]24 funckii Schwägr. (B. funkii auct. non Schwägr.) [278]25 gemmiferum R.Wilczek & Demaret26 gemmilucens R.Wilczek & Demaret27 gemmiparum De Not. (B. alpinum subsp. gemmiparum(De Not.) Kindb.)28 intermedium (Brid.) Blandow (B. nitidulum Lindb.)[278]29 klinggraeffii Schimp.30 knowltonii Barnes (B. lacustre (F.Weber & D.Mohr)Blandow non Brid.)31 kunzei Hornsch. (B. caespiticium subsp. kunzei(Hornsch.) Podp., B. caespiticium var. imbricatumBruch & Schimp., B. caespiticium var. kunzei(Hornsch.) Braithw., nom. illeg.) [278] [283] [288]32 longisetum Blandow ex Schwägr. (B. inclinatum subsp.longisetum (Blandow ex Schwägr.) Podp.)33 marratii Hook.f. & Wilson34 mildeanum Jur. (B. alpinum var. mildeanum (Jur.) Podp.)35 miniatum Lesq. [289]36 minii Podp. [290]37 moravicum Podp. (B. flaccidum auct. non Brid., B.laevifilum Syed, B. subelegans auct. non Kindb.,Rosulabryum laevifilum (Syed) Ochyra) [278]38 muehlenbeckii Bruch & Schimp. (B. alpinum var.brevifolium Myrin, Imbribryum muehlenbeckii (Bruch& Schimp.) N.Pedersen) [291]39 neodamense Itzigs. (B. cavifolium Wilson, nom. nud. insynon., B. crispulum Hampe ex Müll.Hal., B.neodamense var. ovatum (Lange & C.E.O.Jensen)Lindb. & Arnell, B. pseudotriquetrum subsp.neodamense (Itzigs.) J.J.Amann, B. pseudotriquetrumvar. cavifolium Schimp., B. subneodamense Kindb.)[292]40 oblongum Lindb. (B. blindii subsp. oblongum (Lindb.)Kindb., B. blindii var. oblongum (Lindb.) Mönk.) [281]41 pallens Sw. ex anon. (B. aeneum Blytt ex Bruch &Schimp. [293], B. oeneum Blytt ex Bruch & Schimp.emend. Wijk et al., B. rutilans auct. eur. non Brid., B.subelegans Kindb.) [106] [278]42 pallescens Schleich. ex Schwägr. (B. lonchocaulonMüll.Hal. [294], B. obconicum Hornsch. ex Bruch &Schimp.)

218 M. O. HILL ET AL.43 pseudotriquetrum (Hedw.) P.Gaertn. et al. (B.castaneum I.Hagen, B. ventricosum Dicks., nom. illeg.)var. bimum (Schreb.) Lilj. (B. bimum (Schreb.) Turner,B. pseudotriquetrum subsp. bimum (Schreb.) Hartm.)[295]var. pseudotriquetrum44 radiculosum Brid.45 riparium I.Hagen46 rubens Mitt.47 ruderale Crundw. & Nyholm48 salinum I.Hagen ex Limpr. [296]49 sauteri Bruch & Schimp.50 schleicheri DC. (B. schleicheri var. latifolium(Schwägr.) Schimp., B. turbinatum subsp. schleicheri(Schwägr.) Kindb., B. turbinatum var. latifolium(Schwägr.) Bruch & Schimp.) [297]51 subapiculatum Hampe (B. erythrocarpum auct. partim,B. microerythrocarpum Müll.Hal. & Kindb. exMacoun)52 tenuisetum Limpr.53 torquescens Bruch & Schimp. (B. capillare subsp.icodense (H.Winter) Podp., B. capillare subsp.torquescens (Bruch & Schimp.) Kindb., B. capillarevar. torquescens (Bruch & Schimp.) Husn., B. icodenseH.Winter, B. obconicum auct. non Hornsch. ex Bruch& Schimp., Rosulabryum torquescens (Bruch &Schimp.) J.R.Spence) [278] [298]54 turbinatum (Hedw.) Turner (B. syriacum Lorentz)55 uliginosum (Brid.) Bruch & Schimp. (B. cernuum(Hedw.) Bruch & Schimp.)56 valparaisense Thér. (B. pyriferum Crundw. &H.Whitehouse) [299]57 violaceum Crundw. & Nyholm58 warneum (Röhl.) Brid. (B. mamillatum Lindb., B.oelandicum H.Philib.) [278]59 weigelii Spreng. (B. duvalii Voit)60 wrightii Sull. & Lesq. (B. globosum Lindb., B.mamillatum var. globosum (Lindb.) Berggr.)143 Plagiobryum Lindb.1 demissum (Hook.) Lindb. (Bryum demissum Hook.)2 zieri (Hedw.) Lindb. (Bryum zieri Hedw.) [300]144 Rhodobryum (Schimp.) Limpr., nom. cons.1 ontariense (Kindb.) Kindb. (Bryum ontariense Kindb.,R. spathulatum auct. non (Hornsch.) Pócs) [301]2 roseum (Hedw.) Limpr. (Bryum roseum (Hedw.)P.Gaertn. et al.)Mielichhoferiaceae Schimp. (Mniaceae subfam.Pohlioideae auct., nom. nud.) [302]145 Epipterygium Lindb.1 tozeri (Grev.) Lindb.146 Mielichhoferia Nees & Hornsch.1 elongata (Hoppe & Hornsch. ex Hook.) Hornsch.(M. mielichhoferiana var. elongata (Hoppe & Hornsch.ex Hook.) Wijk & Margad.) [303]2 mielichhoferiana (Funck) Loeske (M. mielichhoferi(Hook.) Wijk & Margad., nom. inval.)147 Pohlia Hedw. [304] [305]section Pohlia1 bolanderi (Lesq.) Broth. [306]2 cruda (Hedw.) Lindb.3 crudoides (Sull. & Lesq.) Broth.4 elongata Hedw.var. acuminata (Hornsch.) Huebener (P. acuminataHornsch., P. ambigua (Limpr.) Broth.) [288] [307]var. elongatavar. greenii (Brid.) A.J.Shaw (P. elongata var.polymorpha (Hornsch.) Nyholm [288]5 longicolla (Hedw.) Lindb. [32]6 nutans (Hedw.) Lindb. [308]subsp. nutanssubsp. schimperi (Müll.Hal.) Nyholm (P. nutans var.purpurascens Latzel, P. schimperi (Müll.Hal.)A.L.Andrews) [308]7 obtusifolia (Vill. ex Brid.) L.F.Koch8 saprophila (Müll.Hal.) Broth. [309]9 sphagnicola (Bruch & Schimp.) Broth.section Cacodon Lindb. ex Broth.10 andalusica (Höhn.) Broth.11 andrewsii A.J.Shaw12 annotina (Hedw.) Lindb. (P. grandiflora H.Lindb.)13 bulbifera (Warnst.) Warnst.14 camptotrachela (Renauld & Cardot) Broth.15 drummondii (Müll.Hal.) A.L.Andrews16 erecta Lindb. (P. defecta (Sanio) A.L.Andrews)17 filum (Schimp.) Martensson (P. schleicheri H.A.Crum)18 flexuosa Hook. (P. muyldermansii R.Wilczek &Demaret) [310]var. flexuosavar. pseudomuyldermansii (Arts, Nordhorn-Richter &A.J.E.Sm.) A.J.E.Sm. (P. muyldermansii var.pseudomuyldermansii Arts, Nordhorn-Richter &A.J.E.Sm.) [311]19 lescuriana (Sull.) Ochi (Mniobryum pulchellum(Hedw.) Loeske)20 ludwigii (Spreng. ex Schwägr.) Broth.21 lutescens (Limpr.) H.Lindb.22 proligera (Kindb.) Lindb. ex Broth.23 scotica Crundw.24 tundrae A.J.Shaw [312]section Apalodictyon (Müll.Hal.) Ochyra (sectionMniobryum Nyholm, nom. inval.)25 atropurpurea (Wahlenb.) H.Lindb.26 melanodon (Brid.) A.J.Shaw (Mniobryum delicatulum(Hedw.) Dixon)27 vexans (Limpr.) H.Lindb.28 wahlenbergii (F.Weber & D.Mohr) A.L.Andrews(Mniobryum wahlenbergii (F.Weber & D.Mohr) Jenn.)var. calcarea (Warnst.) E.F.Warb.var. glacialis (Brid.) E.F.Warb.var. wahlenbergii148 Schizymenium Harv.1 pontevedrense (Luisier) Sérgio, Casas, Cros & BruguésMniaceae Schwägr. [302]149 Mnium Hedw., nom. cons.

MOSSES OF EUROPE AND MACARONESIA 219section Mnium1 hornum Hedw.section Spinosa (Kindb.) T.J.Kop.2 spinosum (Voit) Schwägr.3 spinulosum Bruch & Schimp.4 thomsonii Schimp.section Laevinervia P.C.Chen ex X.J.Li & M.Zang5 lycopodioides Schwägr. (M. ambiguum H.Müll.) [313]6 marginatum (Dicks.) P.Beauv.var. dioicum (H.Müll.) Crundw. (M. marginatum var.riparium (Mitt.) Husn.)var. marginatumsection Stellariformia (Kindb.) T.J.Kop.7 blyttii Bruch & Schimp.8 heterophyllum (Hook.) Schwägr.9 stellare Hedw.150 Trachycystis T.J.Kop.1 ussuriensis (Maack & Regel) T.J.Kop. (T. immarginata(Broth.) Laz.) [314]Cinclidiaceae Kindb. (Mniaceae tribus CinclidieaeT.J.Kop.) [302]151 Cinclidium Sw.1 arcticum (Bruch & Schimp.) Schimp.2 latifolium Lindb.3 stygium Sw.4 subrotundum Lindb.152 Cyrtomnium Holmen1 hymenophylloides (Huebener) T.J.Kop.2 hymenophyllum (Bruch & Schimp.) Holmen153 Rhizomnium (Broth.) T.J.Kop.1 andrewsianum (Steere) T.J.Kop.2 gracile T.J.Kop.3 magnifolium (Horik.) T.J.Kop.4 pseudopunctatum (Bruch & Schimp.) T.J.Kop.5 punctatum (Hedw.) T.J.Kop.Plagiomniaceae T.J.Kop. (Mniaceae tribus PlagiomnieaeT.J.Kop.) [302]154 Plagiomnium T.J.Kop.section Plagiomnium1 cuspidatum (Hedw.) T.J.Kop.2 drummondii (Bruch & Schimp.) T.J.Kop.section Rosulata (Kindb.) T.J.Kop.3 affine (Blandow ex Funck) T.J.Kop.4 curvatulum (Lindb.) Schljakov (P. medium subsp.curvatulum (Lindb.) T.J.Kop.) [315]5 elatum (Bruch & Schimp.) T.J.Kop.6 ellipticum (Brid.) T.J.Kop.7 medium (Bruch & Schimp.) T.J.Kop.section Undulata (Kindb.) T.J.Kop.8 confertidens (Lindb. & Arnell) T.J.Kop.9 undulatum (Hedw.) T.J.Kop.var. madeirense T.J.Kop. & Sérgio [316]var. undulatumsection Rostrata (Kindb.) T.J.Kop.10 rostratum (Schrad.) T.J.Kop.155 Pseudobryum (Kindb.) T.J.Kop.1 cinclidioides (Huebener) T.J.Kop.Aulacomniaceae Schimp.156 Aulacomnium Schwägr., nom. cons.1 androgynum (Hedw.) Schwägr.2 palustre (Hedw.) Schwägr.3 turgidum (Wahlenb.) Schwägr.Orthodontiaceae (Broth.) Goffinet157 Orthodontium Schwägr.1 gracile (Wilson) Schwägr. ex Bruch & Schimp.2 lineare Schwägr. (O. australe Hook.f. & Wilson [317])3 pellucens (Hook.) Bruch & Schimp.Rhizogoniales (M.Fleisch.) Goffinet & W.R.BuckRhizogoniaceae Broth.158 Leptotheca Schwägr.1 gaudichaudii Schwägr. [318]Calomniaceae Kindb.159 Calomnion Hook.f. & Wilson1 complanatum (Hook.f. & Wilson) Lindb. [319]Hookeriales M.Fleisch.Hypopterygiaceae Mitt.160 Hypopterygium Brid.1 tamarisci (Sw.) Brid. ex Müll.Hal. (H. muelleriHampe) [320]Daltoniaceae Schimp.161 Achrophyllum Vitt & Crosby1 dentatum (Hook.f. & Wilson) Vitt & Crosby [321]162 Calyptrochaeta Desv.1 apiculata (Hook.f. & Wilson) Vitt (Eriopus apiculatus(Hook.f. & Wilson) Mitt.)163 Daltonia Hook. & Taylor, nom. cons.1 splachnoides (Sm.) Hook. & Taylor2 stenophylla Mitt. [322]164 Distichophyllum Dozy & Molk.1 carinatum Dixon & W.E.NicholsonHookeriaceae Schimp.165 Hookeria Sm., nom. cons.1 lucens (Hedw.) Sm.Leucomiaceae Broth.166 Tetrastichium (Mitt.) Cardot1 fontanum (Mitt.) Cardot2 virens (Cardot) S.P.Churchill (Lepidopilum virensCardot)Pilotrichaceae Kindb.167 Cyclodictyon Mitt.1 laetevirens (Hook. & Taylor) Mitt. [323]Hypnales (M.Fleisch.) W.R.Buck & VittFontinalaceae Schimp.168 Dichelyma Myrin1 capillaceum (L. ex Dicks.) Myrin2 falcatum (Hedw.) Myrin169 Fontinalis Hedw.1 antipyretica Hedw. [324]subsp. antipyreticasubsp. bryhnii (Limpr.) Podp. (F. bryhnii Limpr. exI.Hagen) [325]subsp. gracilis (Lindb.) Kindb. (F. antipyretica var.gracilis (Lindb.) Schimp.)

220 M. O. HILL ET AL.subsp. kindbergii (Renauld & Cardot) Cardot (F.kindbergii Renauld & Cardot)2 dalecarlica Schimp.3 dichelymoides Lindb. [326]4 hypnoides C.Hartm.var. duriaei (Schimp.) Kindb.var. hypnoides5 squamosa Hedw.var. curnowii Cardotvar. dixonii (Cardot) A.J.E.Sm.var. squamosaClimaciaceae Kindb.170 Climacium F.Weber & D.Mohr1 dendroides (Hedw.) F.Weber & D.MohrAmblystegiaceae Kindb. [327]171 Amblystegium Schimp. [327]1 confervoides (Brid.) Schimp. (Platydictya confervoides(Brid.) H.A.Crum, Serpoleskea confervoides (Brid.)Loeske)2 radicale (P.Beauv.) Schimp. (A. saxatile Schimp.,Campylium radicale (P.Beauv.) Grout)3 serpens (Hedw.) Schimp. (A. juratzkanum Schimp.)4 subtile (Hedw.) Schimp. (Platydictya subtilis (Hedw.)H.A.Crum, Serpoleskea subtilis (Hedw.) Loeske)172 Anacamptodon Brid.1 splachnoides (Froel. ex Brid.) Brid.173 Campyliadelphus (Kindb.) R.S.Chopra1 chrysophyllus (Brid.) R.S.Chopra (Campyliumchrysophyllum (Brid.) Lange)2 elodes (Lindb.) Kanda (Campylium elodes (Lindb.)Kindb.)174 Campylium (Sull.) Mitt.1 laxifolium Engelmark & Hedenäs [328]2 longicuspis (Lindb. & Arnell) Hedenäs3 protensum (Brid.) Kindb. (C. stellatum subsp.protensum (Brid.) C.E.O.Jensen, C. stellatum var.protensum (Brid.) Bryhn)4 stellatum (Hedw.) Lange & C.E.O.Jensen(Campyliadelphus stellatus (Hedw.) Kanda)175 Conardia H.Rob. [327]1 compacta (Drumm. ex Müll.Hal.) H.Rob.(Amblystegium compactum (Drumm. ex Müll.Hal.)Austin, Rhynchostegiella compacta (Drumm. exMüll.Hal.) Loeske)176 Cratoneuron (Sull.) Spruce1 curvicaule (Jur.) G.Roth (Callialaria curvicaulis (Jur.)Ochyra, Cratoneuron filicinum var. curvicaule (Jur.)Mönk.)2 filicinum (Hedw.) Spruce177 Drepanocladus (Müll.Hal.) G.Roth, nom. cons.1 aduncus (Hedw.) Warnst. (D. polycarpos (Blandow exVoit) Warnst., D. simplicissimus Warnst., D. stagnatusŻarnowiec)2 arcticus (R.S.Williams) Hedenäs (Campylium arcticum(R.S.Williams) Broth.)3 longifolius (Mitt.) Paris (D. capillifolius (Warnst.)Warnst.)4 polygamus (Schimp.) Hedenäs (Campyliadelphuspolygamus (Schimp.) Kanda, Campylium polygamum(Schimp.) Lange & C.E.O.Jensen)5 sendtneri (Schimp. ex H.Müll.) Warnst.6 sordidus (Müll.Hal.) Hedenäs (D. tenuinervis T.J.Kop.)178 Hygroamblystegium Loeske, nom. cons. [327] [329]1 fluviatile (Hedw.) Loeske (Amblystegium fluviatile(Hedw.) Schimp.)2 humile (P.Beauv.) Vanderp., Goffinet & Hedenäs(Amblystegium humile (P.Beauv.) Crundw.,Leptodictyum humile (P.Beauv.) Ochyra,Leptodictyum kochii (Schimp.) Warnst.)3 tenax (Hedw.) Jenn. (Amblystegium tenax (Hedw.)C.E.O.Jensen)4 varium (Hedw.) Mönk. (Amblystegium varium (Hedw.)Lindb., Orthotheciella varia (Hedw.) Ochyra) [330]179 Hygrohypnum Lindb. [327] [331]1 alpestre (Hedw.) Loeske (Ochyraea alpestris (Hedw.)Ignatov & Ignatova)2 alpinum (Lindb.) Loeske3 cochlearifolium (Venturi) Broth. (Ochyraeacochlearifolia (Venturi) Ignatov & Ignatova)4 duriusculum (De Not.) D.W.Jamieson (Hygrohypnelladuriuscula (Turner ex Wilson) Ignatov & Ignatova, H.dilatatum (Wilson) Loeske)5 eugyrium (Schimp.) Broth. (Pseudohygrohypnumeugyrium (Schimp.) Kanda)6 luridum (Hedw.) Jenn. (Pictus scoticus C.C.Towns.)[332]7 molle (Hedw.) Loeske (H. molle var. schimperianum(Lorentz) Loeske)8 montanum (Lindb.) Broth. (Ochyraea montana(Lindb.) Ignatov & Ignatova)9 norvegicum (Schimp.) J.J.Amann (Ochyraea norvegica(Schimp.) Ignatov & Ignatova)10 ochraceum (Turner ex Wilson) Loeske (Hygrohypnellaochracea (Turner ex Wilson) Ignatov & Ignatova)11 polare (Lindb.) Loeske (Hygrohypnella polaris(Lindb.) Ignatov & Ignatova)12 smithii (Sw.) Broth. (Ochyraea smithii (Sw.) Ignatov& Ignatova)13 styriacum (Limpr.) Broth.14 subeugyrium (Renauld & Cardot) Broth.(Pseudohygrohypnum subeugyrium (Renauld &Cardot) Ignatov & Ignatova)180 Leptodictyum (Schimp.) Warnst. [333]1 riparium (Hedw.) Warnst. (Amblystegium riparium(Hedw.) Schimp.)181 Ochyraea Váňa1 tatrensis Váňa [331]182 Palustriella Ochyra1 commutata (Hedw.) Ochyra (Cratoneuroncommutatum (Hedw.) G.Roth, P. commutata var.fluctuans (Schimp.) Ochyra, P. commutata var.virescens (Schimp.) Ochyra)2 decipiens (De Not.) Ochyra (Cratoneuron decipiens(De Not.) Loeske)

MOSSES OF EUROPE AND MACARONESIA 2213 falcata (Brid.) Hedenäs (Cratoneuron falcatum (Brid.)G.Roth, P. commutata var. falcata (Brid.) Ochyra,P. commutata var. sulcata (Lindb.) Ochyra)4 pluristratosa M.Stech & J.-P.Frahm [334]183 Pseudocalliergon (Limpr.) Loeske1 angustifolium Hedenäs [335]2 brevifolium (Lindb.) Hedenäs (Drepanocladusbrevifolius (Lindb.) Warnst.) [336]3 lycopodioides (Brid.) Hedenäs (Drepanocladuslycopodioides (Brid.) Warnst.)4 trifarium (F.Weber & D.Mohr) Loeske (Calliergontrifarium (F.Weber & D.Mohr) Kindb.)5 turgescens (T.Jensen) Loeske (Scorpidium turgescens(T.Jensen) Loeske)184 Sanionia Loeske [327]1 georgicouncinata (Müll.Hal.) Ochyra & Hedenäs(S. nivalis Hedenäs)2 orthothecioides (Lindb.) Loeske3 uncinata (Hedw.) Loeske (Drepanocladus uncinatus(Hedw.) Warnst.)185 Tomentypnum Loeske [327]1 nitens (Hedw.) LoeskeCalliergonaceae (Kanda) Vanderp., Hedenäs, C.J.Cox &A.J.Shaw [327]186 Calliergon (Sull.) Kindb. [337]1 cordifolium (Hedw.) Kindb.2 giganteum (Schimp.) Kindb.3 megalophyllum Mikut.4 richardsonii (Mitt.) Kindb. (C. obtusifolium Karczm.)[338]187 Hamatocaulis Hedenäs1 lapponicus (Norrl.) Hedenäs (Drepanocladuslapponicus (Norrl.) Smirnova)2 vernicosus (Mitt.) Hedenäs (Drepanocladus vernicosus(Mitt.) Warnst.)188 Loeskypnum H.K.G.Paul1 badium (Hartm.) H.K.G.Paul (Drepanocladus badius(Hartm.) G.Roth)189 Scorpidium (Schimp.) Limpr.1 cossonii (Schimp.) Hedenäs (Drepanocladus cossonii(Schimp.) Loeske, Drepanocladus intermedius(Lindb.) Warnst., Limprichtia cossonii (Schimp.)L.E.Anderson, H.A.Crum & W.R.Buck, Limprichtiaintermedia (Lindb.) Loeske)2 revolvens (Sw. ex anon.) Rubers (Drepanocladusrevolvens (Sw. ex anon.) Warnst., Limprichtiarevolvens (Sw. ex anon.) Loeske) [106]3 scorpioides (Hedw.) Limpr.190 Straminergon Hedenäs1 stramineum (Dicks. ex Brid.) Hedenäs (Calliergonstramineum (Dicks. ex Brid.) Kindb.)191 Warnstorfia Loeske1 exannulata (Schimp.) Loeske (Drepanocladusexannulatus (Schimp.) Warnst.)2 fluitans (Hedw.) Loeske (Drepanocladus fluitans(Hedw.) Warnst., Drepanocladus h-schulzei (Limpr.)Loeske, W. fluitans var. falcata (Sanio exC.E.O.Jensen) H.A.Crum & L.E.Anderson, W.h-schulzei (Limpr.) Loeske)3 procera (Renauld & Arnell) Tuom. (Drepanocladusprocerus (Renauld & Arnell) Warnst.) [339]4 pseudostraminea (Müll.Hal.) Tuom. & T.J.Kop.(Drepanocladus pseudostramineus (Müll.Hal.) G.Roth)5 sarmentosa (Wahlenb.) Hedenäs (Calliergonsarmentosum (Wahlenb.) Kindb.)6 trichophylla (Warnst.) Tuom. & T.J.Kop.(Drepanocladus trichophyllus (Warnst.) Podp.)7 tundrae (Arnell) Loeske (Drepanocladus tundrae(Arnell) Loeske)Leskeaceae Schimp. [340]192 Haplocladium (Müll.Hal.) Müll.Hal.1 angustifolium (Hampe & Müll.Hal.) Broth.(Bryohaplocladium angustifolium (Hampe &Müll.Hal.) R.Watan. & Z.Iwats., Thuidiumangustifolium (Hampe & Müll.Hal.) A.Jaeger)2 microphyllum (Hedw.) Broth. (Bryohaplocladiummicrophyllum (Hedw.) R.Watan. & Z.Iwats.)3 virginianum (Brid.) Broth. (Bryohaplocladiumvirginianum (Brid.) R.Watan. & Z.Iwats., Thuidiumvirginianum (Brid.) Schimp.)193 Lescuraea Schimp.1 mutabilis (Brid.) Lindb. ex I.Hagen2 saxicola (Schimp.) Molendo3 secunda Arnell [341]194 Leskea Hedw.1 polycarpa Hedw.195 Lindbergia Kindb.1 brachyptera (Mitt.) Kindb. [342]196 Pseudoleskea Schimp. [343] [344]1 artariae Thér. (Pseudoleskeopsis artariae (Thér.) Thér.)2 incurvata (Hedw.) Loeske (Lescuraea incurvata(Hedw.) E.Lawton)3 patens (Lindb.) Kindb. (Lescuraea patens Lindb.)4 radicosa (Mitt.) Macoun & Kindb. (Lescuraearadicosa (Mitt.) Mönk.)5 saviana (De Not.) Latzel (Lescuraea saviana (De Not.)E.Lawton)197 Pseudoleskeella Kindb. [345]1 catenulata (Brid. ex Schrad.) Kindb.2 nervosa (Brid.) Nyholm (Leskeella incrassata (Lindb.ex Broth.) Broth. [346], Leskeella nervosa (Brid.)Loeske)3 papillosa (Lindb.) Kindb.4 rupestris (Berggr.) Hedenäs & L.Söderstr. (P. nervosavar. rupestris (Berggr.) Nyholm, P. sibirica (Arnell)P.S.Wilson & D.H.Norris) [347]5 tectorum (Funck ex Brid.) Kindb. ex Broth.198 Ptychodium Schimp.1 plicatum (Schleich. ex F.Weber & D.Mohr) Schimp.(Lescuraea plicata (Schleich. ex F.Weber & D.Mohr)Broth., Pseudoleskea plicata (Schleich. ex F.Weber &D.Mohr) Kindb.)Thuidiaceae Schimp. [348]199 Abietinella Müll.Hal.

222 M. O. HILL ET AL.1 abietina (Hedw.) M.Fleisch. (Thuidium abietinum(Hedw.) Schimp.)var. abietinavar. hystricosa (Mitt.) Sakurai (A. hystricosa (Mitt.)Broth., Thuidium abietinum subsp. hystricosum(Mitt.) Kindb., Thuidium abietinum var. hystricosum(Mitt.) Loeske & Lande) [32]200 Helodium Warnst., nom. cons.1 blandowii (F.Weber & D.Mohr) Warnst. (Thuidiumblandowii (F.Weber & D.Mohr) Schimp.)201 Pelekium Mitt., nom. cons. [348]1 atlanticum (Hedenäs) Hedenäs (Cyrto-hypnumatlanticum (Hedenäs) Hedenäs & Sérgio, Thuidiumatlanticum Hedenäs) [349]2 minutulum (Hedw.) Touw (Cyrto-hypnum minutulum(Hedw.) W.R.Buck & H.A.Crum, Microthuidiumminutulum (Hedw.) Warnst., Thuidium minutulum(Hedw.) Schimp.)202 Thuidiopsis (Broth.) M.Fleisch.1 sparsa (Hook.f. & Wilson) Broth. (Cyrto-hypnummontei Hedenäs) [350]203 Thuidium Schimp.1 assimile (Mitt.) A.Jaeger (T. delicatulum var. radicans(Kindb.) H.A.Crum, Steere & L.E.Anderson, T.philibertii Limpr.) [351]2 delicatulum (Hedw.) Schimp. (T. erectum Duby)3 recognitum (Hedw.) Lindb.4 tamariscinum (Hedw.) Schimp.Brachytheciaceae Schimp. [352]Eurhynchioideae Milde (Rhynchostegioideae Ignatov &Huttunen) [352]204 Pseudoscleropodium (Limpr.) M.Fleisch.1 purum (Hedw.) M.Fleisch. (Scleropodium purum(Hedw.) Limpr.)205 Scorpiurium Schimp.1 circinatum (Bruch) M.Fleisch. & Loeske (Thamniumcossyrense var. melitense Bott.) [353]2 deflexifolium (Solms) M.Fleisch. & Loeske3 sendtneri (Schimp.) M.Fleisch. (Thamnium cossyrenseBott. var. cossyrense, Thamnobryum cossyrense (Bott.)A.J.E.Sm.) [353]206 Palamocladium Müll.Hal.1 euchloron (Müll.Hal.) Wijk & Margad.207 Plasteurhynchium M.Fleisch. ex Broth.1 meridionale (Schimp.) M.Fleisch. (Eurhynchiummeridionale (Schimp.) De Not.)2 striatulum (Spruce) M.Fleisch. (Eurhynchiumstriatulum (Spruce) Schimp., Isothecium striatulum(Spruce) Kindb.)208 Eurhynchium Schimp.1 angustirete (Broth.) T.J.Kop. (E. striatum subsp.zetterstedtii (P.Størmer) Podp., E. striatum var.pachycladum G.Roth)2 striatum (Hedw.) Schimp.209 Platyhypnidium M.Fleisch.1 grolleanum Ochyra & Bednarek-Ochyra [354]2 lusitanicum (Schimp.) Ochyra & Bednarek-Ochyra(P. alopecuroides (Brid.) A.J.E.Sm., Rhynchostegiumalopecuroides (Brid.) A.J.E.Sm., Rhynchostegiumlusitanicum (Schimp.) A.J.E.Sm., hom. illeg.)3 mutatum Ochyra & Vanderp. [355]4 riparioides (Hedw.) Dixon (Rhynchostegium riparioides(Hedw.) Cardot)5 torrenticola (Ochyra, C.Schmidt & Bültmann) Ochyra& Bednarek-Ochyra (Gradsteinia torrenticola Ochyra,C.Schmidt & Bültmann) [356]210 Rhynchostegium Schimp.1 arcticum (I.Hagen) Ignatov & Huttunen (R. muralevar. arcticum I.Hagen) [357]2 confertum (Dicks.) Schimp. (R. surrectum (Mitt.)A.Jaeger) [358]3 megapolitanum (Blandow ex F.Weber & D.Mohr)Schimp. (Brachythecium cardotii H.Winter) [359]4 murale (Hedw.) Schimp.5 rotundifolium (Scop. ex Brid.) Schimp.6 strongylense (Bott.) W.R.Buck & Privitera (Barbellastrongylensis Bott.) [360]Helicodontioideae M.Fleisch. (RhynchostegielloideaeIgnatov & Huttunen) [352] [361]211 Nobregaea Hedenäs1 latinervis Hedenäs [362]212 Helicodontium Schwägr.1 capillare (Hedw.) A.Jaeger (H. italicum (Schimp.)M.Fleisch.) [363]213 Rhynchostegiella (Schimp.) Limpr., nom. cons.1 bourgaeana (Mitt.) Broth. [364]2 curviseta (Brid.) Limpr. (R. curviseta var. curviseta, R.curviseta var. laeviseta (W.E.Nicholson & Dixon)Podp., R. letourneuxii (Besch.) Broth.)3 durieui (Mont.) P.Allorge & Perss.4 litorea (De Not.) Limpr. (R. tenella var. litorea (DeNot.) Mönk.) [365]5 macilenta (Renauld & Cardot) Cardot [366]6 tenella (Dicks.) Limpr.var. meridionalis (Boulay) Zoddavar. tenella7 teneriffae (Mont.) Dirkse & Bouman (R. jacquinii(Garov.) Limpr., R. teesdalei (Schimp.) Limpr.) [367]8 tenuicaulis (Spruce) Kartt. (Cirriphyllum germanicum(Grebe) Loeske & M.Fleisch., Cirriphyllum tenuicaule(Spruce) Wijk & Margad.) [368]9 trichophylla Dirkse & Bouman [369]214 Cirriphyllum Grout1 crassinervium (Taylor) Loeske & M.Fleisch.(Eurhynchium crassinervium (Taylor) Schimp.)2 piliferum (Hedw.) Grout215 Oxyrrhynchium (Schimp.) Warnst.1 hians (Hedw.) Loeske (Eurhynchium hians (Hedw.)Sande Lac., O. swartzii (Turner) Warnst.)2 pumilum (Wilson) Loeske (Eurhynchium pumilum(Wilson) Schimp., Rhynchostegiella pallidirostris(Brid.) Loeske, nom. illeg., Rhynchostegiella pumila(Wilson) E.F.Warb.)

MOSSES OF EUROPE AND MACARONESIA 2233 schleicheri (R.Hedw.) Röll (Eurhynchium schleicheri(R.Hedw.) Milde)4 speciosum (Brid.) Warnst. (Eurhynchium speciosum(Brid.) Jur.)Brachythecioideae Engler216 Kindbergia Ochyra1 praelonga (Hedw.) Ochyra (Eurhynchium praelongum(Hedw.) Schimp., Eurhynchium praelongum var.stokesii (Turner) Dixon, Eurhynchium stokesii(Turner) Schimp., Oxyrrhynchium praelongum (Hedw.)Warnst.)217 Bryhnia Kaurin1 scabrida (Lindb.) Kaurin (B. novae-angliae auct. eur.non (Sull. & Lesq.) Grout) [370]218 Myuroclada Besch.1 maximowiczii (G.G.Borshch.) Steere & W.B.Schofield[371]219 Sciuro-hypnum Hampe1 flotowianum (Sendtn.) Ignatov & Huttunen(Cirriphyllum reichenbachianum (Huebener) Wijk &Margad., Cirriphyllum velutinoides (Schimp.) Loeske& M.Fleisch., nom. illeg., Eurhynchium flotowianum(Sendtn.) Kartt.)2 glaciale (Schimp.) Ignatov & Huttunen(Brachythecium dovrense (Limpr.) Amman,Brachythecium glaciale Schimp., Brachytheciumglaciale var. dovrense Limpr.) [372]3 latifolium (Kindb.) Ignatov & Huttunen(Brachythecium latifolium Kindb.)4 oedipodium (Mitt.) Ignatov & Huttunen(Brachythecium curtum (Lindb.) Limpr.,Brachythecium oedipodium (Mitt.) A.Jaeger,Brachythecium starkei var. curtum (Lindb.) Warnst.)5 ornellanum (Molendo) Ignatov & Huttunen(Brachythecium ornellanum (Molendo) Venturi &Bott., Scleropodium apiculigerum (Lindb. & Arnell)J.-P.Frahm, Scleropodium ornellanum (Molendo)Lorentz)6 plumosum (Hedw.) Ignatov & Huttunen, nom. cons.(Brachythecium plumosum (Hedw.) Schimp.)7 populeum (Hedw.) Ignatov & Huttunen(Brachythecium populeum (Hedw.) Schimp.)8 reflexum (Starke) Ignatov & Huttunen (Brachytheciumreflexum (Starke) Schimp.)9 starkei (Brid.) Ignatov & Huttunen (Brachytheciumstarkei (Brid.) Schimp., Brachythecium starkei var.tromsoeense (Kaurin & Arnell) Nyholm)220 Brachythecium Schimp. [373]1 albicans (Hedw.) Schimp.2 campestre (Müll.Hal.) Schimp. (B. ryanii Kaurin) [374]3 capillaceum (F.Weber & D.Mohr) Giacom. (B.rotaeanum De Not. [375], B. salebrosum subsp.rotaeanum (De Not.) Amman, B. salebrosum var.capillaceum (F.Weber & D.Mohr) Lorentz)4 cirrosum (Schwägr.) Schimp. (Cirriphyllum cirrosum(Schwägr.) Grout)5 coruscum I.Hagen (B. groenlandicum (C.E.O.Jensen)Schljakov) [376]6 erythrorrhizon Schimp.subsp. asiaticum Ignatov [377]subsp. erythrorrhizonvar. erythrorrhizonvar. thedenii (Schimp.) Lindb. [378]7 geheebii Milde (Homalothecium geheebii (Milde) Wigh)8 glareosum (Bruch ex Spruce) Schimp.9 laetum (Brid.) Schimp. (B. oxycladon auct. non (Brid.)A.Jaeger) [379]10 mildeanum (Schimp.) Schimp.var. mildeanumvar. udum (I.Hagen) Mönk. (B. udum I.Hagen)11 percurrens Hedenäs [380]12 rivulare Schimp.13 rutabulum (Hedw.) Schimp.var. atlanticum Hedenäs [381]var. rutabulum14 salebrosum (Hoffm. ex F.Weber & D.Mohr) Schimp.,nom. cons.15 tommasinii (Sendtn. ex Boulay) Ignatov & Huttunen(Cirriphyllum tenuinerve (Lindb.) Wijk & Margad.,Cirriphyllum tommasinii (Sendtn. ex Boulay) Grout)16 turgidum (Hartm.) Kindb.Homalothecioideae Ignatov & Huttunen221 Scleropodium Bruch & Schimp.1 cespitans (Wilson ex Müll.Hal.) L.F.Koch(Brachythecium appleyardiae McAdam & A.J.E.Sm.)[382]2 touretii (Brid.) L.F.Koch [32]222 Eurhynchiastrum Ignatov & Huttunen1 pulchellum (Hedw.) Ignatov & Huttunen(Eurhynchium pulchellum (Hedw.) Jenn.)var. diversifolium (Schimp.) Ochyra & Żarnowiec(Eurhynchium pulchellum var. diversifolium (Schimp.)C.E.O.Jensen)var. praecox (Hedw.) Ochyra & Żarnowiec(Eurhynchium pulchellum var. praecox (Hedw.) Dixon)var. pulchellum223 Brachytheciastrum Ignatov & Huttunen1 collinum (Schleich. ex Müll.Hal.) Ignatov & Huttunen(Brachythecium collinum (Schleich. ex Müll.Hal.) Schimp.)2 dieckii (Röll) Ignatov & Huttunen (Brachytheciumdieckii Röll, Brachythecium salteri Cardot & Dixon)[383]3 fendleri (Sull.) Vanderp. et al. (Brachythecium fendleri(Sull.) A.Jaeger) [384]4 olympicum (Jur.) Vanderp. et al. (Brachytheciumolympicum Jur.) [384]5 trachypodium (Brid.) Ignatov & Huttunen (Brachytheciumtrachypodium (Brid.) Schimp., Brachytheciumtrachypodium var. payotianum (Boulay) Bott.)6 vanekii (Šmarda) Ochyra & Żarnowiec (Brachytheciumvanekii Šmarda)7 velutinum (Hedw.) Ignatov & Huttunen(Brachythecium velutinum (Hedw.) Schimp.)

224 M. O. HILL ET AL.var. salicinum (Schimp.) Ochyra & Żarnowiec(Brachythecium velutinum var. salicinum (Schimp.)Mönk.)var. vagans (Milde) Ochyra & Żarnowiec(Brachythecium velutinum var. vagans (Milde) Warnst.)var. velutinum224 Homalothecium Schimp.1 aureum (Spruce) H.Rob.2 lutescens (Hedw.) H.Rob. (Camptothecium lutescens(Hedw.) Schimp.)var. fallax H.Philib. ex Schimp.var. lutescens3 philippeanum (Spruce) Schimp.4 sericeum (Hedw.) Schimp.Myriniaceae Schimp.225 Myrinia Schimp., nom. cons.1 pulvinata (Wahlenb.) Schimp.Fabroniaceae Schimp.226 Fabronia Raddi1 ciliaris (Brid.) Brid.2 pusilla RaddiHypnaceae Schimp. [340][385]227 Andoa Ochyra1 berthelotiana (Mont.) Ochyra228 Breidleria Loeske1 pratensis (W.D.J.Koch ex Spruce) Loeske (Hypnumpratense W.D.J.Koch ex Spruce) [386]229 Callicladium H.A.Crum1 haldanianum (Grev.) H.A.Crum230 Calliergonella Loeske1 cuspidata (Hedw.) Loeske2 lindbergii (Mitt.) Hedenäs (Hypnum lindbergii Mitt.)[386]231 Campylophyllum (Schimp.) M.Fleisch. [387]1 calcareum (Crundw. & Nyholm) Hedenäs(Campylidium calcareum (Crundw. & Nyholm)Ochyra, Campylium calcareum Crundw. & Nyholm)2 halleri (Hedw.) M.Fleisch. (Campylium halleri (Hedw.)Lindb.)3 sommerfeltii (Myrin) Hedenäs (Campylidiumsommerfeltii (Myrin) Ochyra, Campylium hispidulumauct. eur. non (Brid.) Mitt. [388], Campyliumsommerfeltii (Myrin) Lange, Campylophyllumhispidulum auct. eur. non (Brid.) Hedenäs [388])232 Ctenidium (Schimp.) Mitt.1 molluscum (Hedw.) Mitt. [389]233 Homomallium (Schimp.) Loeske1 incurvatum (Schrad. ex Brid.) Loeske234 Hyocomium Bruch & Schimp.1 armoricum (Brid.) Wijk & Margad.235 Hypnum Hedw., nom. cons.1 andoi A.J.E.Sm. (H. cupressiforme var. mammillatumBrid., H. mammillatum (Brid.) Loeske, nom. inval.)2 bambergeri Schimp. (Stereodon bambergeri (Schimp.)Lindb.)3 callichroum Brid. (Stereodon callichrous (Brid.)Braithw.)4 cupressiforme Hedw. [390]var. cupressiformevar. filiforme Brid.var. heseleri (Ando & Higuchi) M.O.Hill (H. heseleriAndo & Higuchi) [391]var. lacunosum Brid. (H. cupressiforme subsp.lacunosum (Brid.) Bertsch, H. lacunosum (Brid.)Hoffm. ex Brid.)var. resupinatum (Taylor) Schimp. (H. cupressiformesubsp. resupinatum (Taylor) Hartm., H. resupinatumTaylor)var. subjulaceum Molendo5 fertile Sendtn. (Stereodon fertilis (Sendtn.) Lindb.)6 hamulosum Schimp. (Stereodon hamulosus (Schimp.)Lindb.) [392]7 holmenii Ando (Stereodon holmenii (Ando) Ignatov& Ignatova) [393]8 imponens Hedw.9 jutlandicum Holmen & E.Warncke (H. cupressiformevar. ericetorum Schimp.)10 pallescens (Hedw.) P.Beauv. (Stereodon pallescens(Hedw.) Mitt.)11 plicatulum (Lindb.) A.Jaeger (Stereodon plicatulusLindb.)12 procerrimum Molendo (Ctenidium procerrimum(Molendo) Lindb.)13 recurvatum (Lindb. & Arnell) Kindb. (Drepaniumrecurvatum (Lindb. & Arnell) G.Roth)14 revolutum (Mitt.) Lindb. (Stereodon revolutus Mitt.)var. dolomiticum (Milde) Mönk. (H. dolomiticumMilde)var. revolutum15 sauteri Schimp.16 subimponens Lesq. [394]17 uncinulatum Jur.18 vaucheri Lesq. (Stereodon vaucheri (Lesq.) Lindb. exBroth.)236 Ptilium De Not.1 crista-castrensis (Hedw.) De Not.237 Pylaisia Schimp., nom. cons.1 polyantha (Hedw.) Schimp. (Pylaisiella polyantha(Hedw.) Grout)2 selwynii Kindb. (Pylaisiella selwynii (Kindb.)H.A.Crum, Steere & L.E.Anderson)238 Taxiphyllum M.Fleisch.1 densifolium (Lindb. ex Broth.) Reimers2 wissgrillii (Garov.) Wijk & Margad.239 Vesicularia (Müll.Hal.) Müll.Hal.1 reimersiana Bizot & P.de la Varde [395]Pterigynandraceae Schimp.240 Habrodon Schimp.1 perpusillus (De Not.) Lindb.241 Heterocladium Schimp.1 dimorphum (Brid.) Schimp.2 flaccidum (Schimp.) A.J.E.Sm. (H. heteropterum var.flaccidum Schimp.) [396]3 heteropterum (Brid.) Schimp.

MOSSES OF EUROPE AND MACARONESIA 2254 wulfsbergii I.Hagen (H. heteropterum subsp.wulfsbergii (I.Hagen) C.E.O.Jensen & Perss.) [397]242 Iwatsukiella W.R.Buck & H.A.Crum1 leucotricha (Mitt.) W.R.Buck & H.A.Crum [398]243 Pterigynandrum Hedw.1 filiforme Hedw.var. filiformevar. majus (De Not.) De Not.Hylocomiaceae (Broth.) M.Fleisch.244 Hylocomiastrum Broth.1 pyrenaicum (Spruce) M.Fleisch. (Hylocomiumpyrenaicum (Spruce) Lindb.)2 umbratum (Hedw.) M.Fleisch. (Hylocomium umbratum(Hedw.) Schimp.)245 Hylocomium Schimp., nom. cons.1 splendens (Hedw.) Schimp. [399]246 Loeskeobryum Broth.1 brevirostre (Brid.) M.Fleisch. (Hylocomium brevirostre(Brid.) Schimp.)247 Pleurozium Mitt., nom. cons.1 schreberi (Willd. ex Brid.) Mitt.248 Rhytidiadelphus (Limpr.) Warnst.1 loreus (Hedw.) Warnst.2 squarrosus (Hedw.) Warnst. (Rhytidiastrumsquarrosum (Hedw.) Ignatov & Ignatova)3 subpinnatus (Lindb.) T.J.Kop. (R. squarrosus var.calvescens (Kindb.) Warnst., Rhytidiastrumsubpinnatum (Lindb.) Ignatov & Ignatova)4 triquetrus (Hedw.) Warnst.Rhytidiaceae Broth.249 Rhytidium (Sull.) Kindb.1 rugosum (Hedw.) Kindb.Plagiotheciaceae (Broth.) M.Fleisch.250 Herzogiella Broth. [400]1 seligeri (Brid.) Z.Iwats.2 striatella (Brid.) Z.Iwats.3 turfacea (Lindb.) Z.Iwats.251 Isopterygiopsis Z.Iwats.1 alpicola (Lindb. & Arnell) Hedenäs (Isopterygiumalpicola (Lindb. & Arnell) Nyholm)2 muelleriana (Schimp.) Z.Iwats. (Isopterygiummuellerianum (Schimp.) A.Jaeger)3 pulchella (Hedw.) Z.Iwats. (Isopterygium pulchellum(Hedw.) A.Jaeger)252 Myurella Schimp.1 julacea (Schwägr.) Schimp. [401]2 sibirica (Müll.Hal.) Reimers3 tenerrima (Brid.) Lindb.253 Orthothecium Schimp., nom. cons.1 chryseon (Schwägr.) Schimp. [402]2 intricatum (Hartm.) Schimp.3 lapponicum (Schimp.) C.Hartm.4 rufescens (Dicks. ex Brid.) Schimp.5 strictum Lorentz254 Plagiothecium Schimp.1 berggrenianum Frisvoll2 cavifolium (Brid.) Z.Iwats. (P. roeseanum Schimp.)3 curvifolium Schlieph. ex Limpr. (P. laetum var.curvifolium (Schlieph. ex Limpr.) Mastracci &M.Sauer, P. laetum var. secundum (Lindb.) Frisvollet al.) [403]4 denticulatum (Hedw.) Schimp.var. denticulatumvar. obtusifolium (Turner) Moorevar. undulatum R.Ruthe ex Geh. (P. denticulatum var.ruthei (Limpr.) Riehm., P. ruthei Limpr.) [404]5 laetum Schimp.6 latebricola Schimp.7 neckeroideum Schimp. (P. neckeroideum subsp.noricum (Molendo ex Limpr.) J.J.Amann, P. noricumMolendo ex Limpr.) [405]8 nemorale (Mitt.) A.Jaeger9 piliferum (Sw.) Schimp.10 platyphyllum Mönk.11 succulentum (Wilson) Lindb. [406]12 svalbardense Frisvoll [407]13 undulatum (Hedw.) Schimp. (Buckiella undulata(Hedw.) Ireland)255 Platydictya Berk.1 jungermannioides (Brid.) H.A.Crum (Amblystegiumjungermannioides (Brid.) A.J.E.Sm.)256 Pseudotaxiphyllum Z.Iwats.1 elegans (Brid.) Z.Iwats. (Isopterygium elegans (Brid.)Lindb.)2 laetevirens (Dixon & Luisier ex F.Koppe & Düll)Hedenäs [408]Entodontaceae Kindb., nom. cons.257 Entodon Müll.Hal.1 challengeri (Paris) Cardot (E. compressus Müll.Hal.ex Cardot non (Hedw.) Müll.Hal.) [409]2 cladorrhizans (Hedw.) Müll.Hal.3 concinnus (De Not.) Paris4 schleicheri (Schimp.) Demet.Pylaisiadelphaceae Goffinet & W.R.Buck258 Brotherella Loeske ex M.Fleisch.1 lorentziana (Molendo ex Lorentz) Loeske ex M.Fleisch.259 Heterophyllium (Schimp.) Kindb.1 affine (Hook.) M.Fleisch.260 Isopterygium Mitt.1 tenerum (Sw.) Mitt. (I. bottinii (Breidl.) Kindb.,Sematophyllum bottinii (Breidl.) Podp.) [410]261 Platygyrium Schimp., nom. cons.1 repens (Brid.) Schimp.Sematophyllaceae Broth., nom. cons.262 Hageniella Broth.1 micans (Mitt.) B.C.Tan & Y.Jia (Hygrohypnum micans(Mitt.) Broth., Sematophyllum micans (Mitt.)Braithw.) [411]263 Sematophyllum Mitt.1 adnatum (Michx.) E.Britton [412]2 demissum (Wilson) Mitt.3 substrumulosum (Hampe) E.BrittonCryphaeaceae Schimp.264 Cryphaea D.Mohr

226 M. O. HILL ET AL.1 heteromalla (Hedw.) D.Mohr265 Dendrocryphaea Paris & Schimp. ex Broth.1 lamyana (Mont.) P.Rao (Cryphaea lamyana (Mont.)Müll.Hal.) [413]Leucodontaceae Schimp.266 Antitrichia Brid.1 californica Sull.2 curtipendula (Hedw.) Brid.267 Leucodon Schwägr.1 canariensis (Brid.) Schwägr.2 flagellaris Lindb. ex Broth. [414]3 immersus Lindb.4 pendulus Lindb. [415]5 sciuroides (Hedw.) Schwägr.var. morensis (Schwägr.) De Not.var. sciuroides6 treleasei (Cardot) Paris [416]268 Pterogonium Sw.1 gracile (Hedw.) Sm.Neckeraceae Schimp.269 Homalia Brid., nom. cons.1 lusitanica Schimp. (H. lusitanica var. subrecta (Mitt.)Düll-Herm., H. subrecta (Mitt.) A.Jaeger, Neckerasubrecta Mitt.)2 trichomanoides (Hedw.) Brid. [417]3 webbiana (Mont.) Schimp. (Neckera webbiana (Mont.)Düll)270 Neckera Hedw., nom. cons.1 besseri (Lobarz.) Jur. (Homalia besseri Lobarz.)2 cephalonica Jur. & Unger (N. pennata subsp.cephalonica (Jur. & Unger) Giacom.)3 complanata (Hedw.) Huebener (Homalia complanata(Hedw.) De Not., N. complanata var. longifoliaSchimp., N. complanata var. secunda Grav., N.complanata var. tenella Schimp.)4 crispa Hedw. (N. crispa var. falcata Müll.Hal.,N. pseudopennata (Warnst.) Schlieph. ex Żmuda)5 intermedia Brid. (N. elegans Jur., N. elegans var.laevifolia Schiffn., N. intermedia var. laevifolia(Schiffn.) Renauld & Cardot, N. laeviuscula Geh.)6 menziesii Drumm. (Metaneckera menziesii (Drumm.)Steere) [418]7 oligocarpa Bruch (N. pennata subsp. oligocarpa(Bruch) Giacom., N. pennata var. tenera Müll.Hal.)8 pennata Hedw.9 pumila Hedw. (N. fontinaloides Lindb.,N. fontinaloides var. philippeana (Schimp.) Guim.,N. philippeana Schimp., N. pumila var. philippeana(Schimp.) Milde, N. pumila var. pilifera Jur.)271 Thamnobryum Nieuwl.1 alopecurum (Hedw.) Gangulee (T. alopecurum var.gracillimum Bott., T. mediterraneum (Bott.) G.Roth)2 angustifolium (Holt) Nieuwl.3 cataractarum N.G.Hodgetts & Blockeel [419]4 fernandesii Sérgio (Crassiphyllum fernandesii (Sérgio)Ochyra) [420]5 maderense (Kindb.) Hedenäs (T. alopecurum var.maderense (Kindb.) M.Stech, Ros & O.Werner) [421]6 neckeroides (Hook.) E.Lawton [422]7 rudolphianum Mastracci [423]Echinodiaceae Broth.272 Echinodium Jur.1 prolixum (Mitt.) Broth. (Scleromnium knyi Jur.)2 renauldii (Cardot) Broth.3 setigerum (Mitt.) Jur. (E. setigerum var. integrifoliumLuisier) [424]4 spinosum (Mitt.) Jur. (E. madeirense Jur.)Leptodontaceae Schimp.273 Cryptoleptodon Renauld & Cardot1 longisetus (Mont.) Enroth (Leptodon longisetus Mont.,Neckera longipedunculata Müll.Hal.) [425]274 Leptodon D.Mohr, nom. cons.1 smithii (Hedw.) F.Weber & D.MohrLembophyllaceae Broth.275 Isothecium Brid.1 algarvicum W.E.Nicholson & Dixon (Forsstroemiacanariensis (Renauld & Cardot) Enroth, I. atlanticumHedenäs, nom. illeg., I. canariense H.Winter,Thamnium canariense Renauld & Cardot,Thamnobryum canariense (Renauld & Cardot)D.G.Long) [426]2 alopecuroides (Lam. ex Dubois) Isov. (I. alopecuroidesvar. robustum (Schimp.) Düll, I. circinans Saut.,I. myurum Brid., I. viviparum Lindb.)3 holtii Kindb. (Eurhynchium myosuroides var. rivulare(Limpr.) Paris, I. myosuroides var. rivulare Limpr.)[427]4 myosuroides Brid.subsp. brevinerve Lindb. (I. myurum var. piliferumC.E.O.Jensen, I. myurum var. tenuinerve (Kindb.)Limpr.)subsp. myosuroides (I. myosuroides var. cavernarumMolendo, I. myosuroides var. debile Braithw., I.myosuroides var. falcatum Jaap ex G.Roth, I.myosuroides var. filescens (Renauld) Warnst., I.myosuroides var. filiforme Jaap ex G.Roth,I. myosuroides var. integrifolium Papp)var. brachythecioides (Dixon) Braithw. [427]Myuriaceae M.Fleisch.276 Myurium Schimp.1 hochstetteri (Schimp.) Kindb. (M. hebridarum Schimp.)Anomodontaceae Kindb.277 Anomodon Hook. & Taylor1 attenuatus (Hedw.) Huebener2 longifolius (Schleich. ex Brid.) Hartm.3 rostratus (Hedw.) Schimp.4 rugelii (Müll.Hal.) Keissl.5 tristis (Ces.) Sull. & Lesq. (Haplohymenium triste(Ces.) Kindb.)6 viticulosus (Hedw.) Hook. & Taylor278 Claopodium (Lesq. & James) Renauld & Cardot1 whippleanum (Sull.) Renauld & Cardot(C. whippleanum var. cavernicola Luisier)

MOSSES OF EUROPE AND MACARONESIA 227ANNOTATIONS1. Ignatov & Afonina (1992) report Sphagnum imbricatumHornsch. ex Russow (section Sphagnum), S.orientale L.I.Savicz (section Subsecunda) and S.perfoliatum L.I.Savicz (section Subsecunda) fromarctic European Russia. The record of S. imbricatumwould, if correct, be likely to refer to S. steereiR.E.Andrus, which is seemingly the only taxon of theS. imbricatum complex collected in arctic areas ofNorth America and Eurasia (Flatberg, 1984), andwhich is listed by Dierssen (2001). However, Afonina(pers. comm. to Flatberg) tells us that there are noreliable herbarium specimens of European material ofany of these species. They are therefore omitted.2. Sphagnum viride is closely related to S. cuspidatum(Flatberg, 1988), and isoenzymatic results indicatethat gene flow exists between the two morphs(Hanssen, Såstad & Flatberg, 2000). More geneticdata are required to evaluate their taxonomic status.3. Sphagnum brevifolium and S. isoviitae belong to asmall group of closely related taxa in the S. recurvumcomplex with yellow spores (Flatberg, 1992, 1992[1993]). In typical morphological appearance they arewell separated from S. fallax, but seem not clearlysegregated genetically in sympatric populations(Såstad, Stenøien & Flatberg, 1999). Their speciesstatus can therefore be questioned, and more geneticdata are required before a decisive taxonomic conclusioncan be drawn.4. The name Sphagnum lenense was published as anomen nudum in 1915, and was not validated until1936 (Afonina and Egorova, pers. comm. to Flatberg).5. Sphagnum auriculatum is retained here as the name forthe taxon that has, following a paper by Dirkse &Isoviita (1986), been called S. denticulatum in recentlists. Sphagnum denticulatum is an aquatic form withabnormally large, nearly isophyllous leaves. Althoughmost such plants belong to S. auriculatum, similarforms of S. inundatum are found not uncommonlygrowing under aquatic conditions in mire pools alongthe western coast of Norway. The type specimen of S.denticulatum is therefore unsatisfactory.6. Sphagnum tundrae was described from Svalbard byFlatberg (1994).7. Sphagnum wulfianum is either nested within sectionAcutifolia (Shaw, 2000b) or is sister to sectionAcutifolia, lying between it and section Squarrosa(Shaw, Cox & Boles, 2005). It is left here in its ownsection Polyclada.8. Sphagnum aongstroemii may be nested within sectionAcutifolia (Shaw, 2000b, Shaw et al., 2005). It is lefthere in its own section Insulosa.9. Sphagnum nitidulum Warnst. was described fromTerceira in the Azores, but type material haspresumably been destroyed and its identity is obscure.10. Sphagnum subtile (Russow) Warnst. (S. capillifoliumvar. subtile (Russow) Kartt., S. rubellum var. subtile(Russow) Amann) is a disputed taxon whose distinctnesscan be questioned. Natcheva & Cronberg (2002)consider European herbarium material studied tobelong to S. rubellum. Shaw et al. (2005) claim thatNorth American plants named S. subtile cannot beseparated genetically from S. capillifolium. The nameis based on European material, but original materialhas not been traced. It is premature to disregard theexistence of a European taxon underlying this name.11. Sphagnum tenerum Sull. & Lesq. ex Sull. (S. capillifoliumvar. tenerum (Sull. & Lesq. ex Sull.) H.A.Crum)is an eastern North American species, which accordingto Shaw et al. (2005) is highly differentiated geneticallyfrom S. capillifolium. European plants labelled S.tenerum have been examined by Flatberg, and belongto hemi-isophyllous forms of S. capillifolium.12. Sphagnum olafii was described from Svalbard byFlatberg (1993a).13. Sphagnum rubiginosum was described from Norway byFlatberg (1993b).14. Sphagnum andersonianum has been reported fromEurope, but Shaw et al. (2005) claim that examinedmaterial from Europe and North America cannot bedistinguished by genetic markers from S. rubellum.15. Sections of Andreaea follow Murray (1988).16. Sérgio (2004) recognized Andreaea heinemannii subsp.crassifolia, stating that it does not intergrade morphologicallywith subsp. heinemannii and that they arefound in one mixed population. The taxon may thusmerit species status.17. Smith (2004) treated the East Asian Atrichum rhystophyllum(Müll.Hal.) Paris as a synonym of A.angustatum, following Richards & Wallace (1950),who treated it as a variety of A. angustatum. Therelationships of North American, European and EastAsian representatives of this complex and theiraffinity with A. undulatum are not clear and are inneed of thorough revision, as stated by Lou &Koponen (1986).18. A critical, global revision is required to clarify thestatus of Atrichum flavisetum and its relationship to A.undulatum.19. Pogonatum inflexum (Lindb.) Sande Lac. is listed forTurkey by Kürschner & Erdağ (2005). Pogonatuminflexum is an exclusively East Asian species confinedto China, Japan, Korea and the Russian Far East.Several records from other parts of mainland Asiahave been confirmed as misidentified, and are mostlyP. neesii (Hyvönen, 1989). We have not studied anyTurkish material but are inclined to assume that theserecords also represent P. neesii, a widespread Asianspecies with a disjunct occurrence in the Caucasusarea (see below).20. According to Schratz (1928), Pogonatum aloides var.minimum (Crome) Molendo and P. nanum var. long-

228 M. O. HILL ET AL.isetum Hampe ex Bruch & Schimp. are hybridsbetween P. aloides and P. nanum.21. Pogonatum neesii was reported for European Russiafor the first time by Akatova & Ignatova (2000).22. According to the latest phylogenetic analyses(Hyvönen et al., 2004, Koskinen & Hyvönen, 2004),neither Polytrichastrum nor the traditional Polytrichumsensu lato is monophyletic. Furthermore, severalPolytrichastrum species have been shown to beallopolyploids (Derda & Wyatt, 2000; Van der Velde& Bijlsma, 2001). However, as Polytrichum sensustricto (Smith, 1971) may be monophyletic, and as theparaphyly of Polytrichastrum is due to some taxabeing more closely related to some Pogonatum species,we continue to accept Polytrichastrum until such timeas more comprehensive phylogenetic analyses havebeen completed.23. No infraspecific taxa of Polytrichastrum alpinum arerecognized here, not even var. fragile, which Long(1985) recognized, although with reservations.Results obtained by Yli-Rekola (1980) andSchriebl (1991) in culture experiments support thiswide circumscription of the species, althoughneither study included var. fragile. Although thistaxon is easily identified by its caducous leaves thatare constricted at the line of dehiscence, thistendency may be a result of phenotypic plasticityassociated with certain habitats or environmentalstimuli. A similar feature has been observed byLong (1988) at the population level in Pogonatumurnigerum (which recent phylogenetic analyses suggestis closely related to Polytrichastrum alpinum)and by both Holmen (1960) and Long (1985) forPogonatum dentatum. As the other distinguishingcharacters of var. fragile are well within the rangeof variation for the species, it would be inconsistentto treat this feature as taxonomically significant inone taxon only.24. Polytrichastrum sphaerothecium was treated by Corleyet al. (1981) as a synonym of P. sexangulare and byMerrill (1992) as a variety of P. sexangulare.25. Recent genetic (Van der Velde & Bijlsma, 2000) andculture experiments (Schriebl, 1991) based onEuropean material support the recognition ofseveral closely related species formerly treated asvarieties of Polytrichum commune. Differencesbetween some of these are small, and studies ofNorth American material have not supported all ofthem (Derda & Wyatt, 1990). A global studyincluding examination of type material is requiredto stabilize the taxonomy. We follow Long (1985)and Bijlsma et al. (2000) in reducing P. communevar. humile and var. perigoniale to synonymy underP. commune. As noted by Long (1985), perichaetialleaf length varies significantly in P. commune andminor differences in form hardly warrant taxonomicrecognition.26. The generic status of Bryobrittonia is questionable.Recent European checklists followed Horton (1983) intreating it as a genus, whereas Nyholm (1998) treatedit as a member of Encalypta section Streptotheca. Thequestion is still unresolved, so we follow previouschecklists by treating Bryobrittonia as a genus.27. Encalypta sections follow Horton (1983).28. Encalypta serbica Katić was described from Serbiain 1906, but has not been refound. Horton (1983)was unable to trace a type specimen, but thought fromthe description that it was probably a form of E.ciliata.29. The occurrence of Encalypta intermedia in Europe hashitherto not been well documented. It was describedfrom Harput and Berit Dagt in Turkey (D.G. Horton,pers. comm.), and is listed for Turkey by Kürschner &Erdağ (2005). It was added to the European list byCorley & Crundwell (1991) because Horton (1983)stated that it occurred in Europe. Horton (pers.comm.) has confirmed two specimens (B) collectedby J. Bornmüller in 1907 from Gröden in the SouthTyrol and determined by him as E. commutata.30. Encalypta obovatifolia was described from Gotlandand Öland by Nyholm (1995 [1996]).31. The status of Encalypta rhaptocarpa var. leptodon isdisputed. Nyholm (1998) and Mogensen (2001) accordit species rank as E. trachymitria, while Ignatov &Ignatova (2003) treat it as a form of E. rhaptocarpaand Kučera & Váňa (2003) treat it as doubtful.32. The original spelling of names terminating with -collaor -collum is retained, in accordance with article 60.1of the International Code (Greuter et al., 2000). Theargument of Crundwell (1970) in favour of the -collisending is not sustainable. Trematodon brevicollis andT. longicollis are, however, correct. We have alsochecked the original spelling of Hypnum(Scleropodium) touretii Brid., Splachnum (Aplodon)wormskioldii Hornem. and Thuidium hystricosum Mitt.(Abietinella abietina var. hystricosa).33. Encalypta affinis subsp. macounii is commonly treatedas a species, but Horton (1983; and pers. comm. toHill) points out that it differs from subsp. affinis onlyin the muticous rather than hair-pointed leaves.34. Entosthodon subpallescens Laz. (Funaria subpallescens(Laz.) Smirnova) has been included in European listsbut is Asian.35. The hybrid taxa Funaria6hybrida R.Ruthe ex Limpr.(Entosthodon fascicularis6Funaria hygrometrica) andPhyscomitrella6hampei Limpr. (usually interpreted asPhyscomitrella patens6Physcomitrium eurystomum)are omitted from the main list.36. The genera Funaria and Entosthodon are distinguishedaccording to the revision by Fife (1985).37. For synonyms of Entosthodon durieui see Brugués(1998).38. For synonymy of Entosthodon hungaricus see Canoet al. (1999).

MOSSES OF EUROPE AND MACARONESIA 22939. The poorly known Macaronesian endemic Funariafritzei is treated by Losada-Lima, Dirkse &Rodríguez-Núñez (2001) as a synonym ofEntosthodon krausei. It needs further elucidation.40. Entosthodon schimperi Brugués is a new name forthe moss described by Schimper as Funaria durieui.The epithet durieui is not available in Entosthodon.The species has been found in Portugal, Spain andthe Canary Islands (Brugués, Dirkse & Sérgio,2001).41. Entosthodon mouretii was reported from Spain byBrugués et al. (1999).42. Funaria maireana Copp., from Greece, is too poorlyknown to be included in the checklist. F. anomala Jur.and Funaria handelii Schiffn. (Entosthodon handelii(Schiffn.) Laz.) are listed for Turkey by Kürschner &Erdağ (2005).43. Goffinet & Buck (2004) placed Goniomitrium inPottiales on the basis of a molecular study byGoffinet & Cox (2000). However, the followingmorphological features favour Funariales: laminalcells smooth, large and thin-walled; stomata withsimple guard cells; calyptra mitriform.44. Fife & Seppelt (2001) expressed doubts about thedistinctness of Goniomitrium seroi from the AustralianG. acuminatum Hook. & Wilson. A fuller study ofAustralian and Spanish material is required beforethese two species can be treated as synonyms.45. Physcomitrium eurystomum subsp. acuminatum istreated as a species in the Polish checklist (Ochyraet al., 2003) but as a synonym of subsp. eurystomum inthe alpine countries from which it was described.46. For synonymy of Pyramidula tetragona see Brugués,Casas & Belmonte (1998).47. For synonymy of Oedipodiella australis see Brugués(2003).48. Bryoxiphium madeirense was described from Madeiraby Löve & Löve (1953) but ignored by Corley et al.(1981) and Corley & Crundwell (1991). It is listed byEggers (1982), Düll (1992) and Dierssen (2001).49. Coscinodon humilis was described from Europe, butlargely ignored by European authors although recognizedin Japan; European plants lack capsules(Greven, 1995).50. For typification of Grimmia alpestris, see Muñoz(1997).51. According to Maier (2002a), Grimmia limprichtii is asynonym of G. tergestina. However, Greven (2000)treats it as distinct.52. Grimmia donniana var. curvula is homotypic with G.arenaria.53. Grimmia capillata, originally described from Sardinia,was generally treated as a variety of G. crinita duringthe 20th century. Both Greven (1995) and Muñoz &Pando (2000) recognize it at species rank.54. Grimmia curviseta was described from Tenerife byBouman (1991).55. Grimmia dissimulata was described from Cyprus byMaier (2002b), who also listed localities from Turkey,Syria and southern and western Europe.56. Dryptodon incurvus is homotypic with Grimmia elatior.57. Grimmia funalis var. calvescens is homotypic with G.funalis f. epilifera J.E.Zetterst., a heterotypic synonymof G. funalis.58. Although the name Grimmia retracta remains to betypified, it is now treated as a synonym of G. lisae bySmith (2004).59. Grimmia muehlenbeckii and G. poecilostoma wereincluded by Corley et al. (1981) in G. trichophyllaand G. tergestina, respectively. Both Greven (1995)and Muñoz & Pando (2000) recognize them as species.60. Grimmia pulvinata var. obtusa is homotypic with G.pulvinata var. africana.61. The identity of Grimmia reflexidens, which wasdescribed from Chile, is controversial; Maier (2002a)and Ochyra (2004b) treat it as a species of Coscinodon.62. Grimmia grisea has been reported from Iceland byGreven (1998). This taxon was originally describedfrom South Georgia (Cardot, 1906) after materialscollected by Skottsberg. There are no originalmaterials at PC, and all the available syntypes at S(Skottsberg original herbarium) and H-BR are conspecificwith G. reflexidens.63. Grimmia subsulcata was considered a synonym of G.reflexidens by Muñoz (1998), although he laterrealized (Muñoz & Pando, 2000) that this typificationwas in error (the name is homotypic with G. alpestrisvar. microstoma Bruch & Schimp.). No type of thelatter name could be found in BM (originalSchimper’s herbarium) or any of the many herbariato which materials were requested. Pending furtherdevelopments the name is tentatively maintainedunder G. reflexidens.64. The status of Grimmia crassifolia is controversial.There are no capsules in the holotype, and thus it isimpossible to decide if it belongs to G. poecilostoma orG. tergestina. Muñoz (1999) considered it a synonymof G. poecilostoma, but in a more recent paper it wasplaced under G. tergestina (Ignatova & Muñoz, 2005).65. Grimmia austrofunalis is an Australasian and SouthAmerican taxon that does not grow in Europe. It ischaracterized by large gemmae (100–200 mm) formedon short stalks arising at dorsal costa base, whoseliberation does not cause any damage to the leaves.Other characters, such as very large size for the genus(up to 8 cm) and shiny leaves, are more variable(Muñoz, 1999).66. The name Grimmia britannica, now treated by Smith(2004) as a form of G. trichophylla, is a new name forG. robusta Braithw., a later homonym for G. robustaNees & Hornsch. (? Schistidium robustum (Nees &Hornsch.) H.H.Blom) and thus illegitimate. WhenBraithwaite (1872) published his new species, hementioned four syntypes from different Scotland

230 M. O. HILL ET AL.localities, collected by Fergusson, Moore, Stirton andHunt, which are, at least Fergusson’s, a mixture ofseveral species. Not all possible syntypes have beenlocated, and the name remains to be typified.67. Grimmia meridionalis, originally described as G.trichophylla var. meridionalis Müll.Hal. and raised tospecies rank by Maier (2002b), is listed here as asynonym of G. trichophylla, in accordance with theopinion of Muñoz (1999).68. Grimmia triformis was generally treated as a synonymor variety of G. donniana until revised by Muñoz(1998).69. Grimmia ungeri was originally described from Cyprus.Its presence in Europe is disputed by Greven (1995)but asserted by Muñoz & Pando (2000).70. Indusiella thianschanica was reported by Thériot(1918) from Dagestan, Russian Caucasus.71. Jaffueliobryum latifolium has been found in theRussian Caucasus (Kharzinov et al., 2005).72. Bednarek-Ochyra & Ochyra in Ochyra et al. (2003)divide Racomitrium into four genera, namelyRacomitrium Brid., Niphotrichum (Bednarek-Ochyra)Bednarek-Ochyra & Ochyra, Codriophorus P.Beauv.and Bucklandiella Roiv. In the interests of nomenclaturalstability, these subdivisions are recognized hereat the rank of subgenus.73. Racomitrium hespericum was described from theIberian Peninsula by Sérgio, Muñoz & Ochyra (1995).74. Racomitrium lusitanicum was described from Portugalby Ochyra & Sérgio (1992).75. Schistidium andreaeopsis (Müll.Hal.) Laz., knownfrom Beringia, is probably not European.Schistidium marginale, S. pratense, S. scabrum and S.subflaccidium are mapped for Austria on the website‘Mooskartierung in Österreich’ http://www.sbg.ac.at/bot/, on the basis of specimens determined by H.H.Blom (pers. comm.). These names are nomina nudaand cannot be included in the checklist although themapped taxa are thought to be good species.76. Schistidium apocarpum s.l. has been comprehensivelyrevised in northern Europe by Blom (1996). Hesubsequently wrote a substantial flora account ofSchistidium (Blom in Nyholm, 1998) and hasdescribed additional species from elsewhere. Ourtreatment of Schistidium follows his flora accountexcept where otherwise indicated.77. The name Schistidium alpicola (Hedw.) Limpr., nom.rejic., was for long applied to the plant now called S.platyphyllum. It is a synonym of Schistidium agassizii(Blom in Nyholm, 1998) and is now rejected. Blom(pers. comm. to Hill) points out that the authorcitation S. platyphyllum (Mitt.) Kindb., often used inNorth America, is incorrect because Kindberg did notrecognize Schistidium as a genus.78. According to Ochyra et al. (2003), Schistidiumhelveticum (Schkuhr) Deguchi is a synonym of S.singarense and has priority. Blom (1996) was unable tostudy the lectotype of S. helveticum, which was notfound at LAU. He has subsequently written (pers.comm.) ‘All evidence (including the protologue)points towards the taxonomic identity of S. helveticumand S. singarense, further strengthened by the articleby Ochyra et al. (2003)’. Given that there is apparentlyno Schkuhr material at HAL, he concludes that if weaccept the lectotypification by Deguchi, then we mayuse the name S. helveticum. Therefore the name S.helveticum is recommended here, even though a newlectotype to replace the lost type specimen chosen byDeguchi has not been designated.79. For protologues of Grimmia maritima and Dicranumscottianum, see Nelson (1997).80. Schistidium occidentale, a species previously knownonly from the mountains of southwestern NorthAmerica, was found in the Sierra Nevada, southernSpain (Casas, Blom & Cros, 2001).81. Schistidium sinensiapocarpum is mapped for twolocalities in Austria on the website ‘Mooskartierungin Österreich’ http://www.sbg.ac.at/bot/, on the basisof specimens determined by H.H. Blom (pers. comm.).82. Schistidium spinosum was described from the BlackForest, with localities elsewhere in Germany andFrance (Blom & Lüth, 2002).83. Campylostelium pitardii was transferred from Grimmiaby Maier (1998).84. Seligeria transylvanica was synonymized withBrachydontium trichodes by Ochyra (2004c).85. Subgenera of Seligeria are those of Vitt (1976).86. Seligeria galinae was described from the Urals byMogensen & Goldberg (2002).87. Seligeria irrigata was raised to species rank by Ochyra& Gos (1992).88. Fissidens dubius var. mucronatus (Breidl. ex Limpr.)Kartt., Hedenäs & L.Söderstr. is included here in var.dubius.89. For information on Fissidens luisieri refer to Sérgio,Iwatsuki & Ederra (1997).90. Like Sérgio & Carvalho (2003) we include pendingfurther investigation Fissidens polyphyllus var. newtoniiHusn. in F. polyphyllus var. polyphyllus.91. Sérgio et al. (1997) provide a useful discussion ofFissidens serrulatus.92. Limbidia can be greatly reduced in many limbatespecies. Most taxa that are primarily or onlycharacterized by their limbidia restricted to thevaginant lamina, e.g. Fissidens kosaninii Latzel, F.exiguus Sull. and F. viridulus var. intralimbatus(R.Ruthe) Düll, are probably just poorly limbateforms of limbate species.93. Pursell (2005) provides new insight on the characterizationof Fissidens arcticus.94. The European concept of Fissidens bryoides andrelated species diverges considerably from that ofNorth American authors, e.g. Crum & Anderson(1981), Pursell (2005). European floras emphasize

MOSSES OF EUROPE AND MACARONESIA 231differences and recognize several species. NorthAmerican authors stress the presence of intergradingforms and recognize only one species, F. bryoidesHedw. On both continents many specimens are easilyidentified as F. bryoides sensu stricto (with numerousaxillary antheridial buds), F. viridulus (most antheridiaterminal on dwarf male or longer plants) or F. incurvus(incurved capsules). Also on both continents sexualcharacters are more or less correlated to othercharacters (extension of the limbidium, shape of theleaf apex, sporophyte, etc.). Intergrading specimensare also known from both continents and are probablymost common in southern and northern regions.Whether they are more common in North Americais unknown. In the Netherlands most specimens canbe easily identified. Variation in sexuality in Europeanmaterial of this complex has led to the description ofmany varieties and species. In this checklist, we adopta conservative (from a European point of view)perspective, but omit F. exiguus Sull. and F. kosaniniiLatzel, and reduce F. curnovii and F. incurvus tovarietal rank.95. Cortini Pedrotti (2001) and Smith (2004) treatFissidens bryoides var. caespitans at species level. It ishere included as variety because, though extremeforms are clearly distinct from F. bryoides var.bryoides by red rhizoids, a thicker limbidium, inclinedcapsules and a cushion-like growth form, manycollections cannot be identified with certainty.96. In a recent provisional publication Pursell (2005)recognizes Fissidens minutulus as a distinct species,closely related to, yet distinct from, F. crispus(including F. limbatus). We are not convinced thatthe two taxa are distinct in Europe and they aretreated as synonyms here.97. Fissidens gymnandrus is treated by North Americanauthors and Ignatov & Afonina (1992) as a synonymof F. bryoides. However it differs from F. bryoidessensu stricto in being polyoicous, having oblong leaveswith a wide apex, and by its preference for woodysubstrates in regularly flooded localities.98. Fissidens jansenii was described by Sérgio & Pursell(2001). The resemblance of this species to Fissidensrufulus was overlooked by its authors. Both specieshave small cells and limbidia that reach the insertionof the dorsal laminae. F. jansenii has bistratose leaves;the leaves of F. rufulus are irregularly bistratose orunistratose.99. Fissidens microstictus is a relatively unknown speciescharacterized by stems with numerous wide, ellipticalto ovate leaves, broad, acute leaf tips and dorsallaminae that are narrowed below and typically reachthe insertion. All leaves of a stem can be elimbate orlimbate, or limbate and elimbate leaves may occur onthe same stem, limbidia are found on the vaginantlaminae and sometimes also in the middle of thedorsal laminae. It is closely related to F. arnoldii fromwhich it differs in the broadly acute not acuminate leaftips.100. Fissidens nobreganus resembles F. sublineaefoliuswhich also has linear leaves and smooth cells. Itdiffers from this species in having bud-shaped axillaryperigonia and perigonia terminal on short axillarybranches, by smaller, almost isodiametric laminal cellsand limbidia that are restricted to the upper leaves ofperichaetial plants. Fissidens sublineaefolius has terminalperigonia, larger, hexagonal laminal cells andwell-developed limbidia on all laminae of all leaves.101. Fissidens perssonii was synonymized with F. ovatifoliusby Dirkse, Bouman & Losada-Lima (1993).102. Fissidens pusillus is treated by North Americanauthors as an expression of F. bryoides.103. European specimens identified as Fissidens exiguus arepoorly limbate forms of either F. viridulus or of F.pusillus. Fissidens exiguus is treated by NorthAmerican authors as a poorly limbate expression ofF. bryoides.104. Fissidens sublimbatus was reported from the CanaryIslands by Ros et al. (2001).105. Fissidens viridulus is treated by North Americanauthors as an expression of F. bryoides.106. Bryum pallens, Dicranum polysetum, D. viridulum(Fissidens), Hypnum revolvens (Scorpidium) andOrthotrichum pumilum are Swartz names from beforeSpecies Muscorum that were validated in a bookreview (Anon., 1801); thus ‘ex anon.’.107. Fissidens viridulus var. incurvus is linked to var.viridulus by transitional forms. North Americanauthors treat both taxa as expressions of F. bryoides.108. Re-examination of the isotype of Fissidens bambergeriin U showed that it is a poorly limbate form ofF. viridulus. It differs from F. crispus (F. limbatus)with which it has been confused by larger, hardlybulging, less than twice as deep as wide, laminalcells.109. Fissidens marginatulus is not accepted by Ignatov &Ignatova (2003) and seems hardly distinct from F.viridulus.110. Fissidens serratus was reported as F. papillosus, newfor the Canary Islands, by Dirkse, Bruggeman-Nannenga & Bouman (1991). Bruggeman-Nannenga& Pursell (1995) treated F. papillosus as a synonym ofF. serratus.111. Ceratodon antarcticus Cardot and C. heterophyllusKindb. have been reported from arctic and alpineparts of Europe. According to Ochyra (1998), C.antarcticus is merely a developmental phase of C.purpureus with very large mid-leaf cells and plane leafmargins. European records of C. heterophyllus arebased on plants that lack capsules and thereforeaccording to the account of Burley & Pritchard (1990)are not reliable. Nevertheless, the occurrence in coldparts of Europe of Ceratodon plants with rounded,entire leaf apices is remarkable. Cultivation experi-

232 M. O. HILL ET AL.ments and additional genetic data are needed toelucidate their status.112. The assignment of synonyms to the subspecies ofCeratodon purpureus follows Burley & Pritchard(1990).113. Ditrichum gracile was synonymized with D. crispatissimumby Allen (1994).114. Ditrichum punctulatum is a subantarctic species,known in Europe only from Madeira and theAzores. Records of D. difficile (Duby) M.Fleischprobably belong to this species.115. Pleuridium serrulatum Luisier & Dixon, treated asdoubtful by Corley et al. (1981) but included in recentSpanish lists, is represented in Luisier’s herbarium(INA) by non-fruiting plants. It therefore remainsdoubtful.116. Rhamphidium purpuratum, with peristome resemblingSaelania, is excluded from Pottiaceae by Zander(1993). Its leaves resemble Dicranella and its tubersresemble those of Trichodon (Arts, 1989).117. Trematodon perssoniorum, endemic to S. Miguel,Azores, is closely related to T. longicollis. It wasdescribed by Allorge (1951) on the basis of suggestionsmade by her husband (died 1944) and Thériot (died1947). She makes it absolutely clear that the descriptionis newly written by her for the paper. Inaccordance with Article 60.11 of the InternationalCode, the epithet perssonorum is corrected here toperssoniorum (noting that the form perssonii is used for‘of Persson’ in several bryophyte epithets).118. Amphidium tortuosum is a tropical montane species, inEurope only in Tenerife and Madeira, where it hadbeen reported as A. curvipes. Synonymy followsFrahm et al. (2000).119. Cynodontium asperifolium is similar to C. fallax, withrather slight differences. It is known from the UralMountains (Ignatov & Ignatova, 2003).120. The basionym of Cynodontium polycarpon isFissidens polycarpos, which appears incorrectly in theindex of Species Muscorum (Hedwig, 1801) as F.polycarpus.121. Although Dichodontium palustre has been variouslyplaced in Dicranella, Diobelon or Diobelonella, amolecular study by Stech (1999) supports its affiliationwith Dichodontium; which seems reasonable in view ofits leaf shape.122. The systematic position of Dicranoweisia is controversial.Ochyra (1993) argued that its peristomeplaces it in Seligeriaceae. He also (in Ochyraet al., 2003) divided the European species intoDicranoweisia, which occurs in the NorthernHemisphere and Africa, and Hymenoloma, whosedistribution is mainly in the Southern Hemisphere.We recognize the merits of his argument but preferto a adopt a conservative position until furtherwork has been done.123. Oncophorus elongatus, from Norway and Sweden, waselevated to the rank of species by Hedenäs (2005).124. Rhabdoweisia kusenevae Broth. was included byCorley & Crundwell (1991) but treated by Ignatov &Afonina (1992) as a synonym of R. crispata.125. The placement of Schistostega in Dicranales iscontroversial, but we follow Goffinet & Buck (2004)in this.126. Dicranella campylophylla, not listed by Corleyet al. (1981), was reported for Madeira by Eggers(1982).127. Sections of Dicranum follow those of Nyholm (1987),with some nomenclatural changes due to Ochyra et al.(2003).128. Dicranum crassifolium was described from Portugal(Sérgio, Ochyra & Séneca, 1995).129. Dicranum transsylvanicum was described fromRomania by Lüth (2002).130. Dicranum dispersum was described from Germany byEngelmark (1999).131. Dicranum undulatum Schrad. ex Brid. 1801 haspriority over D. bergeri Blandow 1809. Dicranumundulatum Ehrh. ex F.Weber & D.Mohr 1803 is anillegitimate homonym. Although many recentEuropean authors have avoided the name, it is currentin North America and should also be used in Europe(Ochyra et al., 2003).132. Dicranum angustum and D. laevidens are treated asdistinct species, following Hedenäs & Bisang (2004).133. An isotype of Dicranum sendtneri at S consists of amixture of D. fuscescens and D. elongatum. FollowingHedenäs & Bisang (2004), D. sendtneri is notrecognized here as a distinct species.134. Dicranum canariense, often treated as a subspecies ofD. scottianum, is known from the Canary Islands andSpain.135. Campylopus incrassatus was reported under the nameC. eximius from Madeira by Koppe & Düll (1986),having been previously known under other namesincluding C. dixonii Luisier. The synonymy of C.eximius and C. incrassatus is due to Frahm (1987).136. Dicranodontium subporodictyon is only knownas sterile. Therefore its generic position is stilldoubtful.137. Leucobryum albidum is a North American species, inEurope only in the Azores. For differences between itand L. juniperoideum see Frahm (2005b). The distinctionbetween L. albidum and L. juniperoideum iscontroversial; according to Vanderpoorten, Boles &Shaw (2003) they should be treated as synonyms.138. Octoblepharum albidum Hedw. was reported fromTenerife by Düll (1992) on the basis of a 19th centuryherbarium specimen. It is omitted from a recent list forthe Canary Islands (Losada-Lima et al., 2001). If therecord is genuine, the species had probably beenintroduced from the tropics.

MOSSES OF EUROPE AND MACARONESIA 233139. Molecular phylogenies of Pottiaceae by Werner et al.(2004a, 2005b) place the European species in the threegroups recognized here: Merceyoideae (Scopelophila),Trichostomoideae (including Splachnobryum,Ephemeraceae, Pleuroweiseae sensu Corley et al.,1981, and some Barbula species, e.g. B. bolleana),and Pottioideae (including Pottieae, Cinclidotus andBarbuleae sensu Corley et al., 1981). Timmiella doesnot belong in Pottiaceae but is listed here until a betterplace can be found for it. Likewise, those Barbulaspecies that belong in Trichostomoideae have beenretained in Barbula until their correct generic positioncan be ascertained.140. Crumia latifolia (Kindb.) W.B.Schofield is knownfrom the Caucasus but outside Europe.141. Anoectangium handelii is known from the Crimea. It isdescribed by Zander & Weber (2005), correcting anearlier report by Zander (1977) who had treated it as asynonym of Molendoa sendtneriana.142. Ephemerum hibernicum was described from Ireland byHolyoak & Bryan (2005).143. Holyoak has examined much material of Ephemerumserratum and E. minutissimum, and finds that in bothspecies occasional plants have leaves with a weaknerve. He has not seen the type of E. serratum var.praecox, which could be a synonym of either of thesetaxa.144. Ephemerum spinulosum has been found in Ireland andthe Basque region of Spain (Holyoak, 2001; Infante &Heras, 2005).145. Author citation for Eucladium verticillatum follows thetypification by Ochyra & Zijlstra (2005).146. Gymnostomum mosis (Lorentz) Jur. & Milde wasreported from Spain by Martínez-Sánchez, Ros &Guerra (1991), but later Cano, Ros & Guerra (1994)described the new species Gymnostomum lanceolatumbased on this material.147. Gymnostomum aeruginosum var. obscurum wasdescribed by Guerra (2004).148. Spelling of species epithet in Hymenostylium recurvirostrumfollows Hedwig (1801) not Crosby et al.(1999).149. Pleurochaete malacophylla (Müll.Hal.) Broth. is listedfor Turkey by Kürschner & Erdağ (2005).150. Splachnobryum delicatulum is treated by Arts (2001) asa synonym of S. obtusum. Splachnobryum obtusum wasnot included in the main list of Corley et al. (1981),because these authors considered that its occurrenceswere not sufficiently natural. However, it occurs in theopen in Hungary and the Azores, and is accordinglylisted here.151. Tortella arctica (Arnold) Crundw. & Nyholm has beenreported from Svalbard, but the records refer toTrichostomum arcticum (Frisvoll & Elvebakk,1996).152. Tortella alpicola was reported new for Europe byOtnyukova et al. (2004).153. Tortella bambergeri was not recognized as a goodspecies by Corley et al. (1981), but is generallyaccepted by bryologists in central Europe and alsoby us.154. Tortella densa is treated as a variety T. inclinata var.densa, following Eckel (1998).155. Tortella limbata has been refound on the CanaryIslands. Dirkse et al. (1993) compared their collectionwith the holotype and noted the distinctive denticulateleaf border, composed of long narrow cells, runningnearly to the leaf apex.156. Tortella limosella is known only from the Scottishtype-locality, and has not been refound since itsdiscovery in 1906.157. Tortella cirrifolia was synonymized with T. nitida bySjögren (2001).158. Molecular studies by Werner et al. (2005b) show thatthe genus Trichostomum as listed here is heterogeneous,with T. brachydontium and T. crispulum nestedwithin Weissia and T. triumphans very close toPottiopsis caespitosa. Trichostomum connivens(Lindb. ex Broth.) Paris is recorded from theCaucasus but is not known from Europe.Trichostomum contortum (Kunze) Sérgio is probablya synonym of T. brachydontium, although Sérgio(1985) thought that it was a good species.159. For synonymy of Hyophila treleasei, see Sollman(1993).160. Weissia leptocarpa Schimp. ex Besch. is treated bySérgio & Carvalho (2003) as a doubtful species, but isaccepted for Turkey by Kürschner & Erdağ (2005), inspite of the fact that Frey & Kürschner (1991) hadtreated it as doubtful and illegitimate.161. Weissia mittenii (Bruch & Schimp.) Mitt. is omitted,because it is designated above as a hybridW.6mittenii (Bruch & Schimp.) Mitt. emend.A.J.E.Sm. Smith (pers. comm. to Hill) notes that W.mittenii has malformed capsules and abnormal spores.Following Werner et al. (2005b), Astomum is includedin Weissia.162. A molecular study by Werner et al. (2004b) supportsthe species status of Weissia wimmeriana.163. Acaulon mediterraneum is treated by many authors asa variety of A. muticum, to which it is undoubtedlyclose.164. Aloina humilis was described from the Canary Islandsby Gallego et al. (1998).165. Aloina obliquifolia is accorded species rank followingGallego et al. (1999).166. The Macaronesian Barbula semilimbata Dixon &Luisier was listed by Eggers (1982) but is a poorlyknown taxon, not recorded since it was described.167. Barbula convoluta var. sardoa is treated by some atspecies rank. Frahm & Ahmed (2004a) suggested thename B. sardoa. However, the epithet sardoa is correctat the varietal level but superfluous at the species level.

234 M. O. HILL ET AL.If species rank is used, the correct name is B.commutata (see Frahm & Ahmed, 2004b).168. Although Corley & Crundwell (1991) did not acceptthe synonymy of Bryoerythrophyllum campylocarpum,Sollman (1990), Zander (1993) and Sérgio & Carvalho(2003) all accept it, as we do here.169. Cinclidotus bistratosus Kürschner & Lüb.-Nestle andC. nyholmiae Çetin have been described fromAnatolia.170. Cinclidotus confertus Lüth was described from northwestGreece. Dr Alicia Ederra, who treatedCinclidotus for the Flora Briofítica Ibérica, has written(pers. comm. to Hill) ‘I had the opportunity to see theholotype of Cinclidotus confertus from STU. It is,without any doubt, a Cinclidotus; its habit is verysimilar to C. riparius, but, as noted by Lüth, theperistomes are different. So, I think they must beclosely related species.’ She recommends retaining it inthe list, although it has not been found elsewhere.171. Cinclidotus vivesii was described by Ederra in Ederra& Guerra (2005).172. Crossidium davidai was reported new for our areafrom the Canary Islands by Dirkse & Bouman (1995b)and Stern (1995).173. Crossidium geheebii was reported from Italy, new toEurope, by Privitera & Puglisi (2000a).174. Crossidium laevipilum was described in 1931 fromNorth Africa and subsequently found in Spain (Casas,Cros & Brugués, 1993; Guerra et al., 1993).175. Crossidium laxefilamentosum was reported new toEurope by Pócs et al. (2004).176. Dialytrichia fragilifolia was raised to species rank byLara (2005).177. A molecular study by Werner et al. (2005a) shows thatprevious sectional divisions of Didymodon cannot bemaintained, but does not yet provide enough detail todemarcate new sections.178. Didymodon reedii H.Rob. (D. tectorum (Müll.Hal.) K.Saito), listed by Corley & Crundwell (1991), waserroneously reported from Britain (Blockeel & Smith,1998).179. According to Sérgio & Jiménez Fernández (2003),Didymodon soaresii is a non-fruiting species ofTortula, of uncertain species.180. The synonymy of Didymodon asperifolius, D. fallax,D. giganteus, D. maximus, D. spadiceus and D.tophaceus follows Jiménez et al. (2005b).181. Didymodon australasiae synonymy follows Jiménezet al. (2005a).182. For description of Didymodon bistratosus, see Hébrard(1994).183. Didymodon brachyphyllus is reported from Iceland(Jóhannsson, 2003).184. Didymodon erosus was described by Jiménez et al.(2004).185. Didymodon lamyanus, from France, has been omittedfrom recent European lists. It was studied by Werneret al. (2005a). Its closest relatives appear to be D.insulanus, D. nicholsonii and D. vinealis.186. Didymodon mamillosus is treated by Jiménez (2004) asa synonym of D. rigidulus.187. Didymodon sicculus was described by Cano et al.(1996).188. For synonymy of Didymodon subandreaeoides seeKučera & Köckinger (2000).189. Didymodon umbrosus synonymy follows Jiménez et al.(2005a).190. According to Ochyra (1998), Hennediella antarctica(Ångstr.) Ochyra & Matteri is a synonym of H.macrophylla and has priority. However, there appearto be some differences, and we have not changed thename applied to European plants pending furtherstudy of variation in the genus.191. Our circumscription of the genus Microbryum followsZander (1993).192. Phascum piptocarpum is treated as a synonym ofMicrobryum curvicollum following Ros, Werner &Rams (2005).193. The synonymy of Microbryum davallianum and M.starckeanum follows Ros et al. (1996).194. Microbryum fosbergii was raised to species rank andsynonymized with Pottia6andalusica by Ros et al.(2005).195. The basionym of Microbryum starckeanum is Weissiastarckeana, named by Hedwig (1801) after JohanChristian Starke. Although Hedwig’s spelling is notapt, Article 60.1 of the International Code (Greuter etal., 2000) requires that it shall be retained.196. Pseudocrossidium obtusulum, originally described fromSweden, is widespread in North America, from whichP. revolutum is absent. Reasons for treating it as adistinct species are given by Eckel, Hoy & Elliot (1997)and Zander (2002).197. Pseudocrossidium replicatum was found in Italy, newfor the European moss flora (Privitera & Puglisi,2000b).198. Pterygoneurum compactum was described from Spainby Cano, Guerra & Ros (1994).199. Pterygoneurum crossidioides, originally described fromthe Dead Sea area of Israel, was found in Hungary(Pócs, 1999). However, Erzberger & Papp (2004)regard the record as somewhat doubtful.200. Pterygoneurum papillosum was described by Oesau(2003).201. Pterygoneurum squamosum was described from Spainby Segarra et al. (1998).202. Syntrichia treatment follows Gallego (2005).203. Syntrichia bogotensis (Hampe) R.H.Zander has beenfound in Madeira (Gallego, Cano & Sérgio, 2005).204. Some recent authors such as Vanderpoorten (2001)treat Syntrichia calcicola as a synonym of S. ruralis.205. According to a revision by Gallego et al. (2002a),Spanish material ascribed to Syntrichia pseudohandeliiis S. caninervis var. caninervis. These authors treat S.

MOSSES OF EUROPE AND MACARONESIA 235pseudohandelii as a synonym of the non-European S.caninervis var. pseudodesertorum (Vondr.)M.T.Gallego, which is found in western Asia, includingTurkey.206. For the status of Syntrichia caninervis var. abranchesiirefer to Gallego et al. (2002a).207. Syntrichia caninervis var. astrakhanica was describedby Ignatov, Ignatova & Suragina (2002).208. Syntrichia glabra was described by Frahm & Gallego(2001).209. Following Gallego, Cano & Guerra (2004), noinfraspecific taxa are recognized within Syntrichialaevipila, which is synonymized with S. pagorum,Tortula laevipilaeformis and T. saccardoana.210. Syntrichia minor was recognized at species rank byGallego et al. (2000).211. According to Ochyra (1994), Tortula intermedia DeNot. 1838 is a synonym of Tortella humilis, makingTortula intermedia (Brid.) Berk. 1863 an illegitimatehomonym.212. According to Gallego (2005), Tortula papillosa var.meridionalis Warnst. should not be recognized as adistinct variety.213. According to Gallego (2005), Syntrichia princeps var.parnassica (Schiffn.) Podp. should not be recognizedas a distinct variety.214. Author citation of Syntrichia ruralis var. ruraliformisfollows Gallego (2005).215. Syntrichia subpapillosissima was recognized at speciesrank by Gallego et al. (2002b). These authors treatedTortula ruralis var. submamillosa W.A.Kramer as asynonym.216. Tortula buyssonii (H.Philib.) Broth., from theAuvergne, France, is a little-known taxon that hasapparently not been found since the 19th century.217. Tortula caucasica Lindb. (Pottia caucasica (Lindb.)Paris) is known from Georgia but not from Europe.Tortula grandiretis Broth. is listed for Turkey byKürschner & Erdağ (2005).218. Tortula ampliretis, described from the Canary Islandsin a paper by Crundwell, During & Long (1978), wasinexplicably not mentioned by Corley et al. (1981).219. Tortula bolanderi and T. inermis should be in Tortulanot Syntrichia (Werner et al., 2003a).220. Tortula israelis, described from Spain as T. muralisvar. baetica, was raised to species rank by Guerra &Ros in Guerra, Ros & Carrión (1992) and subsequentlysynonymized with T. israelis by Cano, Guerra& Ros (1996).221. According to Cano (2004a), European plants attributedto Crossidium seriatum Crum & Steere should bereferred to Tortula brevissima.222. The synonymy and nomenclature of Tortula hoppeanafollow Ochyra (2004a).223. For the synonymy of Desmatodon meridionalis seeSérgio & Granzow de la Cerda (2001).224. According to Sérgio & Carvalho (2003), the typematerial of Pottia cuneifolia corresponds to P. pallida.Therefore, P. pallida and Tortula zoddae, which is anew name for P. cuneifolia in Tortula, are included inthe synonymy of T. pallida.225. Zander (1993) provided the new name Tortularhodonia for Desmatodon wilczekii, because the epithetwilczekii is not available in Tortula.226. Cano, Werner & Guerra (2005) raised Tortula subulatavar. angustata with bistratose border to species rank, onthe basis of T. angustata Lindb., and provided the newname T. schimperi, because the epithet angustata is notavailable at species rank in Tortula. The other varietiesof T. subulata were sunk in a single variable species.227. According to Cano (2004a, b), Tortula marginatasubsp. limbata is a synonym of T. solmsii.228. Tetraplodon urceolatus is a good species which hasbeen misunderstood in Scandinavia, where it has beenconfused with forms of T. mnioides.229. Orthotrichum urnaceum Müll.Hal. was signified forEurope by Lewinsky-Haapasaari (1995) but thelocality is in Armenia.230. Synonymy of Orthotrichum limprichtii followsLewinsky-Haapasaari in Nyholm (1998).231. Orthotrichum casasianum is reported only from northernSpain (Mazimpaka et al., 1999).232. Orthotrichum crenulatum was reported as O.flowersii Vitt from the French Alps (Boudier &Pierrot, 1992).233. Orthotrichum handiense is an endemic fromFuerteventura, Canary Islands (Lara et al., 1999a;Lara, Garilleti & Mazimpaka, 2003).234. Orthotrichum hispanicum was described from Spain byLara, Garilleti & Mazimpaka (2000).235. Orthotrichum macrocephalum was described fromSpain by Lara, Garilleti & Mazimpaka (1994).236. For synonymy of Orthotrichum lewinskyae see Draperet al. (2003).237. Orthotrichum schimperi is recognized as a species inaccordance with Cortini & Lara in Cortini Pedrotti(2001).238. For synonymy of Orthotrichum rogeri var. defluens,see Garilleti, Lara & Mazimpaka (2002).239. For synonymy of Orthotrichum australe, see Lewinsky(1993).240. Orthotrichum vittii was described from Spain by Laraet al. (1999b).241. For synonymy of Orthotrichum arcangelianum seeLara et al. (2002) and Lewinsky (1993).242. Orthotrichum shawii has been reinstated as a distinctspecies (Mazimpaka et al., 2000).243. Orthotrichum ibericum was described from Spain andPortugal by Lara & Mazimpaka (1993).244. For the synonymy of Orthotrichum caucasicum, seeLewinsky (1993).245. We have excluded Orthotrichum elegans Schwägr. as aEuropean species because Nordic material named as

236 M. O. HILL ET AL.O. speciosum var. elegans (Schwägr.) Warnst. isreferable to O. speciosum. For differences betweenthese two taxa see Vitt & Darigo (1997).246. Orthotrichum speciosum var. brevisetum is known fromSpain & Sicily; see Draper et al. (2003).247. Orthotrichum tortidontium was described fromMorocco and Spain by Lara, Garilleti & Mazimpaka(1996).248. Orthotrichum vladikavkanum was described from theRussian Caucasus by Venturi (1887). It was listed inPodpěra’s (1954) Conspectus, but the Caucasus wasassigned to Asia by Corley et al. (1981). For thepresent enumeration, the Russian part of the Causasusis now again included in Europe.249. Corley et al. (1981) included Ulota macrospora in U.rehmannii, but the two species are distinguishedhere, following Boudier & Pierrot (1996) and Sauer(1998).250. The inclusion of Zygodon forsteri in Codonoblepharonis controversial (Goffinet and Vitt, 1998; Matcham &O’Shea, 2005). In our opinion it is best placed inZygodon. Casas et al. (1985) considered Z. madeirensisto be close to Z. forsteri; it is treated here as asynonym following Düll (1992). Further revision isneeded to assess its identity.251. Zygodon sibiricus was reported from the Europeanside of the Urals by Ignatov & Ignatova (2003).252. Zygodon stirtonii is recognized at species rank followingSmith (2004).253. Hedwigia ciliata var. viridis Bruch & Schimp. is listedby Casas (1991), but Hedenäs (1994) was unable totrace the type specimen, so it is uncertain whether thevariety belongs to H. ciliata or H. stellata.254. Hedwigia ciliata var. leucophaea was illustrated andmapped by Hedenäs (1994).255. Hedwigia stellata was described from Sweden byHedenäs (1994).256. Anacolia laevisphaera is reported from Europe in anabstract by Quandt & Frahm (2004). It is a tropicalmontane species and German specimens from theMosel region had been incorrectly identified asBartramia stricta.257. Anacolia menziesii was found new to Europe in theprovinces of Almería and Granada in southern Spain(García-Zamora et al., 1998). Its distinctness from A.webbii was confirmed by molecular data (Werner etal., 2003b).258. Bartramia pomiformis var. elongata Turner and B.pomiformis var. heteromalla (Brid.) Müll.Hal. aretreated with great scepticism by Nyholm (1998), andvar. elongata is treated as a synonym of var.pomiformis by Smith (2004). These varieties areaccordingly excluded from the checklist.259. Nomenclature for Bartramia section Pyridium (sectionVaginella Müll.Hal., nom. illeg.) follows Ochyra et al.(2003).260. Bartramia breviseta was originally described fromFinnmark, but was treated as a synonym of B.ithyphylla in many European checklists. Fransén(2004) has demonstrated that it is distinct, differing,for example, in the lack of an inner peristome.261. Bartramia ithyphylla var. strigosa (Wahlenb.) Hartm.was mapped by Söderström (1996). However, Fransén(2004) treats it as a synonym of var. ithyphylla, and itis accordingly not recognized here.262. Philonotis marchica var. laxa, described fromSwitzerland and Germany as P. laxa Limpr., is listedby Casas (1991) but is no longer recognized in Germanand Swiss lists.263. Philonotis uncinata (Schwägr.) Brid. is listed for theAzores by Gabriel et al. (2005).264. Molecular data now leave no doubt that the generaAnomobryum, Brachymenium, Plagiobryum andRhodobryum cannot be maintained with the circumscriptionused by Corley et al. (1981) since most areingroups within Bryum sensu lato (e.g. Pedersen, Cox& Hedenäs, 2003; Pedersen & Hedenäs, 2005). Themolecular data also show that there have beenmultiple instances of convergent evolution of morphologicalcharacters in Bryaceae, so that classificationsbased exclusively on the morphologicalcharacters currently known are unsound. Hence, forexample, the genus Rosulabryum proposed by Spence(1996) is not adopted here because molecular dataimply it is polyphyletic (Pedersen et al., 2003). A starthas been made on subdividing Bryum into segregategenera (e.g. Imbribryum N.Pedersen) on the basisof molecular data from multiple loci used in combinationwith morphological data (Pedersen, 2005;Pedersen & Hedenäs, 2005), but their treatmentcannot yet be applied to European species as awhole because molecular data are not availablefor many species. The generic treatment of Corleyet al. (1981) is therefore used here in an interimarrangement.265. Loeske (1932 [1933]) long ago interpreted Anomobryumleptostomoides as fertile A. concinnatum, andnumerous fertile specimens examined by Holyoakfrom its type-locality at Mont Dore (France)consistently show the gametophytic characters ofconcinnatum. This is therefore the only Europeanlocality at which A. concinnatum has been foundwith capsules, and it is significant because sporophytesof A. leptostomoides differ from those of A.julaceum in the erect not inclined to cernuouscapsules and their greatly reduced peristome, asdescribed in detail by Shaw & Fife (1984).Furthermore, similar capsules have been describedfrom Michigan (USA), again from plants withgametophytes indistinguishable from those of A.concinnatum. Shaw & Fife (1984) were ‘inclined torecognize A. leptostomoides at the specific level’ buthesitated to do so because they thought that the

MOSSES OF EUROPE AND MACARONESIA 237usual non-fertile plants would be impossible to namewith certainty. However, European bryologists havelong recognized A. concinnatum without greatdifficulty from non-fertile material, so this form issurely best treated at species rank as advocated byLoeske.266. Sérgio, Cros & Brugués (1996) re-examined Anomobryumlusitanicum in detail, retaining it in the genusAnomobryum.267. Brachymenium commutatum is known in Europe onlyfrom the Sierra Nevada. It was first reported byThériot (1932) as B. commutatum var. hispanicumThér. and rediscovered in 1997 (Brugués et al., 2003).European plants are sterile and less robust than var.attenuatum Thér. & Trab., which was described fromthe Hoggar mountains, Algeria. Small forms are alsofound in the Hoggar (Thériot, 1932). Ochi (1972,1973) did not examine any of the type material of B.commutatum during his review of African Bryoideae.However, a specimen assigned to B. commutatum var.attenuatum from Sudan and material from Eritreawere reidentified by him as the widespread B. exileDozy & Molk. It appears very likely that B.commutatum should be placed as a synonym of B.exile.268. Brachymenium notarisii was placed in the genusHaplodontium for many years, within which it is thetype of subgenus Ateleobryum (Mitt.) Broth. (Wijk,Margadant & Florschütz, 1962). It was transferredfrom Haplodontium to Brachymenium by Shaw (1987),who commented that ‘it certainly does not belong inthe Mielichhoferioideae’.269. Podpěra (1954) identified Mielichhoferia paradoxawith ‘Bryum splachnoides C. M.’, i.e. B. cellulare.However, Shaw (1987) reidentified it as a distinctspecies known only from Macedonia that he placed inBrachymenium, an arrangement that was followed byCorley et al. (1991). The recent checklist of mosses ofthe former Yugoslavia (Pavletic, Martincic & Düll,1999) continues to treat Brachymenium paradoxumerroneously as Bryum cellulare.270. Brachymenium philonotula is known in our area onlyfrom Madeira (Rocha da Pena, 1928, leg. C. Barreto,S). According to Ochi (1972) it is perhaps conspecificwith B. exile Dozy & Molk.271. Bryum pamirense H.Philib. ex Broth. was reportednew for Europe from Komi Republic in northernRussia (Zheleznova & Shubina, 1998). However,Ignatov & Afonina (1992) marked it with ?, to denotea ‘particularly poorly known and putatively ill-definedspecies’. Bryum pamirense is apparently close to B.savicziae and B. uliginosum (Savicz-Ljubitzkaja &Smirnova, 1970), but differs from both of these inbeing synoicous rather than autoicous. Like B.savicziae, it has the cilia of the endostome welldeveloped and appendiculate, whereas B. uliginosumhas short cilia. It might be a form of B. uliginosum orpossibly a hybrid.272. Bryum savicziae Schljakov was described from theKola peninsula in Russia in 1951 but overlooked byIndex Muscorum (Wijk, Margadant & Florschütz,1959, 1969). It is known only from the type-locality. Itresembles B. uliginosum but has the cilia of theendostome well developed and appendiculate. Sporesizes of 19–31 mm given by Savicz-Ljubitzkaja &Smirnova (1970) seem rather variable and perhapsthis suggests a hybrid is involved. Ignatov & Afonina(1992) also reported B. ekstamii L.I.Savicz, B.labradorense H.Philib., B. umbratum I.Hagen and B.zemliae Arnell & Jaderh. from the north of EuropeanRussia. These little-known species require furtherstudy.273. Bryum vermigerum Arnell & C.E.O.Jensen wasdiscovered at Hardanger, Norway in 1915 and1916; it is otherwise known only by a recent reportfrom Iceland. Bryum vermigerum was treated as avalid species by Nyholm (1993) and its combinationof characters is undoubtedly different from that inany other European Bryum species. Nevertheless,Nyholm reported an extraordinarily wide range ofspore sizes (10–20 mm) and, although one capsuleexamined by Holyoak (at S) contained spores ofless variable size (10–14 mm), these included aproportion of small shrunken and presumablyabortive spores. The rarity of B. vermigerum, itspeculiar characters and the variable and sometimesabortive spores may suggest that an interspecifichybrid is involved.274. Bryum colombii Meyl. and B. mesodon J.J.Amann arelisted in the current Swiss list http://www.bryolich.ch/,but their identity is uncertain. Bryum murmanicumBroth. is on the Red List for the Murmansk Oblast ofRussia as Data Deficient; its identity is also uncertain.Bryum geheebii Müll.Hal. was treated as probably asynonym of B. alpinum by Corley et al. (1981), as aspecies by Düll (1985), but very doubtful by Düll(1992); its identity is uncertain. The identity of B. teresLindb. is also uncertain; it was treated by Düll (1985)as a synonym of B. nitidulum (here treated as asynonym of B. intermedium), but this synonymy wasquestioned by Nyholm (1993).275. Bryum gerwigii (Müll.Hal.) Limpr. is known from itstype-locality on limestone rocks above the Rheinfallesnear Schaffhausen in Switzerland and from a fewother reports from central Europe. As with othersubmerged Bryum, it is apparently so greatly modifiedby growth in water (and perhaps by low light levels)that it is difficult to judge which species was itsimmediate progenitor. This uncertainty providesinsufficient reason for treating it as a valid species.Bryum gemmiparum seems closest in view of the leafshape, occurrence of bulbils and the habitat.

238 M. O. HILL ET AL.276. Crundwell (1970) recognized the widespread synoicousforms of Bryum algovicum as var. rutheanum andthe less common autoicous ones as var. algovicum (ofwhich he treated var. compactum as a synonym).Unpublished studies by Holyoak suggest that autoicousplants appear to occur mainly if not entirely onhigh ground in the Alps, whereas synoicous plantsoccur throughout the lowlands, from central Europenorth to Svalbard. The autoicous plants are oftensmall and densely tufted, they sometimes have narrowcapsules, and the endostome processes have narrowperforations. They might merit treatment as a separatespecies rather than merely as a variety.277. Bryum apiculatum has a Pantropical range extendinginto temperate areas, with a single confirmed record inour region, from Tenerife (Ochi, 1972). The synonymyof B. apiculatum follows Ochi (1994).278. Taxonomic treatment of several Bryum species followsHolyoak (2004).279. Bryum veronense was treated as a subspecies of B.argenteum by Podpěra (1954) and Urmi (1987), as ‘aweak species’ by Düll (1992) who noted that verificationis needed for most specimens, and as a distinctspecies allied to B. argenteum by Nyholm (1993).Some of the material referred to B. veronense is amodified form of B. argenteum, including plants witha proportion of achlorophyllose leaf tips thatundoubtedly intergrade with that species, and otherwisetypical B. veronense may have the papilloserhizoids of B. argenteum. However, other materialplaced as B. veronense in herbaria differs in variousrespects and may comprise similarly reduced, ratherfeatureless, modified forms of other Bryum species,e.g. diminutive B. pallens.280. Longton (1981) studied intraspecific variation inmorphology and physiology of Bryum argenteum ona global scale by means of cultures grown on agarunder standardized laboratory conditions. He establishedthat var. lanatum is of doubtful taxonomic valuebecause the excurrent nerve was not maintained incultivation by clones originating in Canada andHawaii. Ochi (1994) also noted ‘gradations with thetypical variety’. Nevertheless, lanatum was treated as adistinct species by Frahm (2002) and Spence &Ramsay (2002), on the basis of its longer acuminateleaf point, excurrent nerve and differing habitatpreferences, but without any mention of Longton’sdetailed study. Unless better evidence for the geneticdistinctness of lanatum can be found it should bedismissed as an inconstant form of B. argenteum.281. Bryum oblongum and B. blindii are treated as separatespecies based on the detailed study by Shaw (1981).282. Bryum bornholmense was redescribed by Crundwell &Whitehouse (2001).283. Corley et al. (1981) suggested that Bryum caespiticiumis probably an aggregate species, within which B.comense and B. kunzei may be good segregates. Thelatter is treated as a valid species here, but the case formaintaining B. comense appears weak. Although Düll(1985) listed B. comense as a species, he later (Düll,1992) expressed doubt. Nyholm (1993) treated it as asynonym of B. caespiticium, as we do here.284. Demaret & Wilczek (1982) examined the types of B.canariense and B. provinciale and concluded that theyrepresented different species, but Corley et al. (1991)did not follow them because ‘their work was based toorigidly on the study of types alone’. Studies byHolyoak support the conclusions of Ochi (1972),who noted that ‘The form called B. provinciale issynoicous, but there are also dioicous plants whichhave no other clear-cut differences from such synoicousones. These two forms should be better includedin a single taxon’.285. Bryum caucasicum is known in Europe only from theholotype from the Caucasus (S). It was transferredfrom Mielichhoferia to Bryum by Cox & Hedderson(2003).286. Bryum demaretianum was described from Belgium byArts (1992).287. Treatment of the Bryum bicolor complex followsHolyoak (2003), who described B. dyffrynense as anew species from Britain.288. Bryum kunzei, Pohlia acuminata and P. polymorphawere described by Hornschuch (1819), not by Hoppe& Hornschuch as commonly stated.289. Bryum miniatum is a predominantly North Americanspecies known in Europe only from the FaeroeIslands.290. Bryum minii was mainly overlooked until the paper bySérgio et al. (1999).291. Bryum muehlenbeckii is probably best regarded as aform of B. alpinum that grows where intermittentlysubmerged in cold water, since a substantial proportionof European specimens intergrade with thatspecies (Holyoak, unpublished). Some specimens havethe characteristic leaves of both B. muehlenbeckii andB. alpinum on different parts of the same stem,apparently as a result of growth under wet or dryconditions, respectively.292. Bryum neodamense is almost certainly a recurrentform of B. pseudotriquetrum, since intermediate plantsoccur intermixed with most populations of neodamense,often in large numbers (Holyoak, in preparation).Occurrence of the neodamense morphotypeappears to be closely associated with intermittentflooding by hard water, most often on lake shores, butalso in basic fens and dune-slacks. Detailed morphologicalanalyses and molecular studies have beencarried out (Holyoak & Hedenäs, in preparation).293. Bryum aeneum was later emended to B. oeneum; seeWijk et al. (1959).294. Zolotov (2000) treats the polyoicous (partly synoicous)B. lonchocaulon as a species distinct from theautoicous B. pallescens. However, he examined few

MOSSES OF EUROPE AND MACARONESIA 239autoicous specimens and the differences he reportedfrom B. lonchocaulon (other than in sexuality) appearinconstant in material from Britain.295. Bryum bimum may merit species rank since it differsfrom B. pseudotriquetrum in being synoicous ratherthan autoicous, perhaps in having twice as manychromosomes (Smith, 2004), and in average dimensionsof the leaf lamina cells and thickness of their cellwalls (Demaret & Empain, 1985; Zolotov, 2000).However, they apparently do not differ much, if at all,in ranges, frequency or ecology, and preliminarymolecular data (Hedenäs, pers. comm. to Holyoak)suggest they are very close.296. Bryum salinum differs from B. archangelicum in only afew characters; a few intermediate specimens areknown.297. Bryum schleicheri author citation follows Geissler(1985).298. Wijk et al. (1959) cited the original description ofBryum torquescens as Bruch ex De Not., Syllab.Musc., no. 163, 1838. Syed (1973) found this referenceto be wrong and he discounted ‘Fl. Sard. Exs., 1828’,given by other authors because he suspected it is anomen nudum. The original description is thereforetaken as that by Bruch & Schimper (Bryol. Eur., 4 p.119, pl. 20, 1839 (fasc. 6–9, Mon., p. 49, pl. 20)) partlybecause this was the earliest description found by Syedand partly because they refer to it as ‘Diese neue Art’.299. Arts, Crundwell & Whitehouse (1995) showed thatBryum pyriferum described from Tenerife is conspecificwith B. valparaisense, which was originallydescribed from Chile but is now also known from N.America, Africa and Portugal.300. Hedwig (1801) used the spelling zierii twice, but thename Bryum zierii commemorates John Zier as waspointed out by Crum & Anderson (1981). More recentauthors who amend it to zieri are therefore correctingit according to Article 60.11 of the International Code(Greuter et al., 2000).301. Touw (1984) gave reasons for rejecting the nameRhodobryum spathulatum in favour of R. ontariense.302. The large family Mniaceae of Goffinet & Buck (2004)is here divided following Koponen (1988) into thethree families traditionally included in Mniaceae andthe remainder, for which the only available name isMielichhoferiaceae.303. Mielichhoferia elongata is often treated as a synonymor variety of M. mielichhoferiana, but according toShaw (2000a), it is in reality a cryptic species.304. Pohlia ramannii Warnst. is recorded from the territorythat was formerly in Finland and now in Russia(Söderström, 1998). It is not mentioned by Ignatov &Afonina (1992) and is treated here as doubtful.Webera luisieri Dixon and W. maderensis Dixon &Luisier, listed by Eggers (1982), are obscure taxawhose identity is uncertain.305. Sections of Pohlia are based on Shaw (1984), exceptthat we follow Smith (2004) in placing P. crudoides insection Pohlia and P. flexuosa in section Cacodon.306. Pohlia bolanderi was found in the Sierra Nevada, newfor Europe (Rams et al., 2004).307. Pohlia ambigua was described from Austria butremains a poorly known taxon. According to Shaw(1982) it is a form of P. elongata. It is autoicous and istherefore assigned to var. acuminata.308. Species limits in the Pohlia nutans group requirefurther elucidation. Köckinger, Kučera & Stebel(2005) studied P. schimperi and established that theonly really stable character separating it from P.nutans is its red coloration; they accordingly gave itsubspecies rank and synonymized it with P. nutansvar. purpurascens. Pohlia marchica Osterwald wasdescribed from near Berlin and subsequently collectedon Bornholm. It has not been found for many years,and is signified by Nyholm (1993) as very close to P.schimperi. Pohlia turonensis J.M.Couderc & Guédès,nom. inval. was described from France. It is a closerelative of P. nutans and has not subsequently beenvalidated.309. Pohlia saprophila, a mainly Asian species, is reportedfrom the Komi Republic, Russia (Zheleznova, 1994).It may prove to be a dioicous race of P. elongata, butneeds further study.310. For the synonymy of Pohlia flexuosa see Townsend(1995).311. Smith (2004) recognized two varieties of Pohliaflexuosa in Europe, namely var. flexuosa and var.pseudomuyldermansii nom. nud. (P. muyldermansiivar. pseudomuyldermansii Arts, Nordhorn-Richter &A.J.E.Sm.). Townsend (1995) had noted that there iscontinuous variation in bulbil morphology in Nepaland India, and argued that the varieties shouldtherefore be treated as synonyms. He hypothesizedthat there might be only two clones in Europe, withvar. muyldermansii possibly a recent introduction.However Smith (pers. comm. to Hill) is of the opinionthat in Europe there has been differentiation into twomorphologically and ecologically distinct groups ofplants. In view of the distribution of var. pseudomuyldermansii(Austria, British Isles and Switzerland)it seems unlikely that the populations concernedrepresent a single clone. The habitat of the varietiesappears to be markedly different in Europe. For thesereasons, Smith (above) makes the new combination P.flexuosa var. pseudomuyldermansii.312. Pohlia tundrae is reported from Central Europe (Düll,1991, 1992; Müller, 2004).313. Mnium lycopodioides is treated here as a full synonymof M. ambiguum following Koponen (1994).314. There are a number of records of Trachycystisussuriensis in the Russian Caucasus, e.g. Kharzivovet al. (2004).

240 M. O. HILL ET AL.315. Plagiomnium curvatulum is treated as a (rather cryptic)species distinct from P. medium on the basis of agenetic study by Wyatt, Odrzykoski & Stoneburner(1993).316. Plagiomnium undulatum var. madeirense was describedfrom Madeira by Koponen & Sérgio (2001).317. According to Ochyra & Broughton (2004),Orthodontium australe is a synonym of O. lineare.318. Leptotheca gaudichaudii is a New Zealand species thathas been introduced to a garden in Ireland (Smith,2004). It is not at present known in the wild in Europe.319. Calomnion complanatum is an Australasian speciesthat has been introduced to a garden in Ireland(Smith, 2004). It is not at present known in the wild inEurope.320. According to Pfeiffer et al. (2000), Hypopterygiumtamarisci is a variable and widely distributed species,which includes H. muelleri, introduced to Portugalfrom the Southern Hemisphere. We follow Sérgio &Carvalho (2003) in using the name H. tamarisci.321. Achrophyllum dentatum is an austral species that hasbeen introduced to a garden in England (Smith, 2004).It is not at present known in the wild in Europe.322. Daltonia stenophylla is reported from Azores bySjögren (2001) and Frahm (2004). According toSjögren, Azores records attributed to D. splachnoidesare actually D. stenophylla; this had been suggested tohim by R. Schumacker and confirmed by H.A. Crum.323. The species epithet of Cyclodictyon laetevirens wasoriginally hyphenated as ‘laete-virens’. This is correctedhere to ‘laetevirens’ in accordance with Article60.9 of the International Code (Greuter et al., 2000).324. The taxonomy of Fontinalis antipyretica is poorlyunderstood and is further complicated by the startlingresults of recent phylogenetic studies using molecularmethods (Shaw & Allen, 2000; Shaw, 2001). Theseshow that European F. antipyretica is more closelyrelated to the European endemic F. squamosa than toAmerican F. antipyretica, which is morphologicallyindistinguishable from the European species. Thesubspecies retained here are recognized as distinct inseveral European countries, but they may be merelyecotypes that have evolved in different locations tosuit similar selective pressures. Many other infraspecifictaxa are not included in the main list. RecentEuropean checklists have included subsp. lachenaudii(Cardot) Podp., subsp. sparsifolia (Limpr.) Kindb.,var. cymbifolia W.E.Nicholson, and var. gigantea(Sull.) Sull.325. Fontinalis antipyretica subsp. bryhnii is mapped as aspecies, F. bryhnii, by Söderström (1996), but istreated with great scepticism by Nyholm (1960) andsignified as a possible hybrid by P. Martiny in Stewart(1995). In view of the taxonomic difficulties in the F.antipyretica aggregate, F. bryhnii is retained here onlyas a subspecies.326. Fontinalis dichelymoides is recognized as a species inrecent lists from Fennoscandia, e.g. Söderström(1996).327. Generic concepts in Amblystegiaceae andCalliergonaceae mostly follow Vanderpoorten et al.(2002). Hygroamblystegium is separated fromAmblystegium according to the results ofVanderpoorten et al. (2003) and Serpoleskea isretained in Amblystegium. Hygrohypnum is not monophyletic,but the genus is retained in its traditionalsense awaiting ongoing studies regarding the relationshipsof its species. The positions of Conardia,Sanionia and Tomentypnum could not be decided byVanderpoorten et al. (2002), and these genera areretained in the Amblystegiaceae until further studieshave been made.328. Campylium laxifolium was described from northernSweden by Engelmark & Hedenäs (1990 [1992]).329. Vanderpoorten (2004) has argued that the fourEuropean Hygroamblystegium species should be synonymizedas H. varium. Further work is needed beforemaking changes.330. Hygroamblystegium varium is an aptly named speciesin a difficult genus. There is evidence that theHygroamblystegium species recognized here are notmonophyletic (Vanderpoorten, Cox & Shaw, 2004).331. Hygrohypnum as circumscribed here is known tocomprise heterogeneous elements, probably frommore than one family. Ochyraea tatrensis would comewithin this circumscription, but a new combination isnot made, pending a full revision of the group.According to Frahm (2005a), O. tatrensis is a mutantof Hygrohypnum smithii, but they are superficiallyvery different and H. smithii was present at only one ofthe O. tatrensis sites.332. The type of Pictus scoticus has been examined byIgnatov and others (pers. comm.), who consider it tobe a form of Hygrohypnum luridum. This view isfollowed here.333. Leptodictyum kurdicum (Schiffn.) Broth. is listed forTurkey by Kürschner & Erdağ (2005).334. Palustriella pluristratosa was described from Valais,Switzerland (Stech & Frahm, 2001). However, in alater publication Frahm (2005a) signifies that P.pluristratosa may be a somatic mutation of P. falcata.335. Pseudocalliergon angustifolium was described fromSweden by Hedenäs (1990 [1992]).336. The high-arctic Pseudocalliergon brevifolium wastransferred to Pseudocalliergon and recognized as aspecies by Hedenäs (1990 [1992]).337. Calliergon orbiculare-cordatum (‘orbicularicordatum’)(Renauld & Cardot) Broth. was listed by Corley et al.(1981) but the specimens are C. richardsonii(Söderström, 1996).338. For observations on Calliergon richardsonii, seeHedenäs (1993b).

MOSSES OF EUROPE AND MACARONESIA 241339. Warnstorfia procera is recognized as a species followingHedenäs (1993b).340. The circumscription of Leskeaceae and Hypnaceaehas been challenged by Gardiner et al. (2005). Werecognize the merit of their proposals, which wouldnecessitate substantial nomenclatural change. Weprefer not to make the changes until a more generaloverview of families in Hypnales is available.341. Lescuraea secunda is reported from European Russiain the Urals (Ignatov & Ignatova, 2004).342. Lindbergia brachyptera, known from the RussianCaucasus, was included by Podpěra (1954) butexcluded by Corley et al. (1981) because theCaucasus was treated by them as part of Asia.343. Pseudoleskea was reinstated as a genus distinct fromLescuraea by Corley & Crundwell (1991), on thebasis of peristome differences emphasized by Crum& Anderson (1981). The peristome characters nowseem less important, and Ignatov & Ignatova (2004)and Söderström (1998) included Pseudoleskea inLescuraea. A conservative position is adoptedhere, pending more detailed analysis by molecularmethods.344. Pseudoleskea brachyclados (Schwägr.) Kindb., originallydescribed from Austria, is listed for Turkey byKürschner & Erdağ (2005). According to Podpěra(1954) it is a variety of P. radicosa (but if so it shouldhave priority at species rank). Other Europeanauthors have treated it as a variety or synonym of P.incurvata.345. We follow Wilson & Norris (1989) in includingLeskeella (Limpr.) Loeske 1903 in PseudoleskeellaKindb. 1897.346. Leskeella incrassata type specimen has been examinedby Ignatov; it is very close to Pseudoleskeella nervosaand, if recognized as a species, will prove to becommon in southern Europe. For the time being thetwo are treated as synonyms.347. Pseudoleskeella rupestris was lectotypified and synonymizedwith P. sibirica by Hedenäs & Söderström(1991).348. In Thuidiaceae, the generic concepts of Touw (2001a)are adopted here. In particular Pelekium Mitt. (1868)emend. Touw is treated as a synonym of Cyrtohypnum(Hampe) Hampe & Lorentz (1869).349. Pelekium atlanticum was described from Madeira asThuidium atlanticum (Hedenäs, 1991). The namePelekium atlanticum (Hedenäs) Touw appears inTouw (2001a), p. 191, but he failed to validate thename by a combination, commenting that P. atlanticumis doubtfully distinct from the American P.muricatulum (Hampe) Touw. On the other hand,Touw was not prepared to treat P. atlanticum as asynonym of P. muricatulum. Hedenäs makes thenecessary combination above.350. For the synonymy of Cyrto-hypnum montei withThuidiopsis sparsa, see Touw (2001a, p. 205). Touwnotes that T. sparsa is known only from a former parkarea in Madeira, and suggests that it is an alien,introduced with ornamental plants.351. Thuidium assimile has priority over T. philibertii, basedon the large-scale revision of Touw (2001b). As notedby Touw, T. assimile belongs to a complex of poorlydefined taxa including T. delicatulum. Observations byMrs C.T. van Dorp on European specimens supportthe recognition of two weakly distinguished species,rather than treating T. assimile as a variety of T.delicatulum.352. Subfamilial and generic concepts in Brachytheciaceaemostly follow Ignatov & Huttunen (2002).These authors overlooked the names HelicodontioideaeM.Fleisch. and Eurhynchioideae Milde, andsuperfluously described these subfamilies asRhynchostegielloideae and Rhynchostegioideae,respectively.353. For the taxonomic status of Thamnium cossyrenseBott. var. cossyrense and var. melitense Bott. seeMastracci (2001).354. Platyhypnidium grolleanum was described from CzechRepublic by Ochyra & Bednarek-Ochyra (1999).355. Platyhypnidium mutatum was described by Ochyra &Vanderpoorten (1999); molecular studies did not finddifferences between this species and P. riparioides(Stech & Frahm, 1999). Frahm (2005a) asserts that itis a somatic mutant of P. riparioides.356. Platyhypnidium torrenticola was described asGradsteinia torrenticola from the Canary Islands byOchyra, Schmidt & Bultmann (1998) and subsequentlytransferred to the genus Platyhypnidium.357. The status of Rhynchostegium arcticum was reevaluatedby Ignatov & Huttunen (2002).358. Rhynchostegium surrectum was synonymized with R.confertum by Hedenäs (1992b).359. Brachythecium cardotii was synonymized withRhynchostegium megapolitanum by Hedenäs (1992b).360. Barbella strongylensis was transferred to Rhynchostegiumby Buck & Privitera (1999).361. Clasmatodon parvulus (Hampe) Sull. is excluded.Heras, Infante & Buck (2006) have found that everySpanish specimen named as Clasmatodon wasPseudoleskeella tectorum. An old German record from1851 is poorly localized (Müller, 2004) and is unlikelyto be C. parvulus.362. Nobregaea latinervis is a morphologically very peculiarspecies known from a single locality in Madeira; it wasdescribed by Hedenäs (1992b).363. Synonymy of Helicodontium capillare follows CortiniPedrotti (2006 [2005]), who indicates that a specimenof the supposedly endemic H. italicum had beenidentified as H. capillare by W.R. Buck.364. Rhynchostegiella bourgaeana is treated by Dirkse &Bouman (1995a) as an endemic of the Canary Islands.On the other hand, Corley et al. (1991) and Hedenäs(1992b) regarded it as a synonym of R. tenella.

242 M. O. HILL ET AL.365. The status of Rhynchostegiella litorea and its differentiationfrom R. tenella are discussed by Dirkse &Bouman (1995a).366. Rhynchostegiella macilenta, omitted by Corley et al.(1981) and Corley & Crundwell (1991), is treated as afull species and illustrated by Dirkse & Bouman(1995a).367. Rhynchostegiella teesdalei and R. jacquinii weresynonymized with R. teneriffae by Dirkse & Bouman(1995a).368. According to Ignatov & Huttunen (2002),Rhynchostegiella tenuicaulis does not belong toBrachytheciaceae and is more likely a member ofAmblystegiaceae, but more studies are needed.369. Rhynchostegiella trichophylla was described from theCanary Islands by Dirkse & Bouman (1995a).370. Molecular data do not confirm the traditional view onthe identity of European Bryhnia scabrida and NorthAmerican B. novae-angliae, so the resurrection of theformer name was suggested by Ignatov & Huttunen(2002).371. Popov et al. (2000) showed that Myuroclada maximowicziiwas reported from Europe on the basis of amisidentification and excluded it from the Europeanflora, but later Virchenko & Babenko (2001) found itas an alien plant in the grounds of the UniversityBotanical Garden, Rostov-na-Donu, southern Russia.372. The status of Brachythecium glaciale var. dovrenseneeds further study. The taxon may deserve recognitionas a species.373. Brachythecium umbilicatum Jur. & Milde is listed forTurkey by Kürschner & Erdağ (2005).374. Brachythecium ryanii was synonymized with B.campestre by Hedenäs (1996).375. Brachythecium rotaeanum and B. capillaceum aretreated as synonyms, as in most recent Europeanchecklists. However, their identity needs confirmation.376. Brachythecium coruscum was recognized as a synonymof B. groenlandicum by Nyholm (1965) but the namewas omitted by Corley et al. (1981).377. Brachythecium erythrorrhizon subsp. asiaticum wasdescribed from the Altai by Ignatov (1998), and isknown from the Urals.378. A typographic error in Wijk et al. (1962) makes itseem that Hypnum thedenii (Schimp.) C.Hartm. (1854)(not Hartm. 1845 as indicated) was published beforeBrachythecium thedenii Schimp. (1853).379. Robinson & Ignatov (1997) showed thatBrachythecium laetum is the correct name for thespecies often called B. oxycladon. In eastern NorthAmerica this species is very common. It is reportedfrom various parts of Europe, but confirmation of theidentity of European plants and the American one isneeded.380. Brachythecium percurrens was described from Madeiraby Hedenäs (1992a). Unpublished molecular data byVanderpoorten et al. and Huttunen et al. suggest thatit belongs to subfamily Helicodontioideae, and may bea member of Oxyrrhynchium.381. Brachythecium rutabulum var. atlanticum wasdescribed from Madeira by Hedenäs (1992a).382. Brachythecium appleyardiae is a synonym ofScleropodium cespitans (Blockeel et al., 2005).383. Brachythecium salteri was synonymized with B. dieckiiby Hedenäs (1993a).384. The combinations Brachytheciastrum fendleri and B.olympicum were made by Vanderpoorten et al. (2005).385. There is still much uncertainty about the correctcircumscription of Hypnaceae and related families.The classification adopted here is provisional, and weare well aware that it will be changed in future.386. The placement of Hypnum lindbergii and H. pratensein Calliergonella and Breidleria, respectively, followsHedenäs (1990).387. The circumscription of Campylophyllum is that ofHedenäs (1997).388. Campylophyllum hispidulum (Brid.) Hedenäs is anAmerican species not found in Europe (Crundwell &Nyholm, 1962).389. Ctenidium molluscum is a very variable species, forwhich numerous infraspecific taxa have beendescribed. The following varieties are recognized inmore than one European country: var. condensatum(Schimp.) E.Britton, var. molluscum, and var. robustumBoulay. The genetic basis of this variationrequires further elucidation.390. Hypnum cupressiforme has such a complex patternof variation that the delimitation of infraspecifictaxa remains highly controversial. The varieties inthe main list are those recognized in current checklistsof at least four European countries. Other names,some of which may have priority over more popularones, are var. brevisetum Schimp., var. imbricatumBoulay, var. julaceum Brid., var. tectorum Brid.,var. tenue Hook. & Taylor, and var. tectorum (Brid.)J.-P.Frahm.391. Hypnum heseleri was described from Saarland inGermany and two localities in the Netherlands(Ando & Higuchi, 1994). van Zanten & Hofman(1994) grew spores from capsules of H. heseleri anddemonstrated that the progeny were about 50% H.heseleri and 50% H. cupressiforme. Allozyme-electrophoresisprovided further convincing evidence that H.heseleri is a locally derived mutant of H. cupressiforme.Accordingly, H. heseleri is treated as a varietyof H. cupressiforme in a new combination, madeabove.392. Hypnum aemulans Breidl. ex Limpr., originallydescribed from Austria, is treated as a goodspecies by Ando (1994) but is listed withoutcomment as a synonym of H. hamulosum byEuropean authors (Koperski et al., 2000; Ulvinen,Syrjänen & Anttila, 2002). European occurrences needfurther investigation.

MOSSES OF EUROPE AND MACARONESIA 243393. Hypnum holmenii was described from Canada,Greenland and Finland (Ando, 1994), and has beenfound in the Arkhangelsk Oblast of Russia (Ignatov &Ignatova, 2004).394. Hypnum subimponens is reported from EuropeanRussia by Afonina (2004).395. Vesicularia sphaerocarpa (A.Jaeger) Broth. wasreported from Malta by Reimers in 1934 on the basisof a 19th century collection by Schweinfurth. Thespecimen (B) was destroyed in the Second World War,but Reimers in Bizot & Potier de la Varde (1952) wasconfident that it belonged to the taxon described bythese authors as V. reimersiana. According to Corleyet al. (1981), V. sphaerocarpa is a synonym of V.galerulata (Duby) Broth. Düll (1985) listed theMaltese plant as V. reimersiana, but also asserted thatV. galerulata had been found in mainland Italy.Corley & Crundwell (1991) accordingly added V.reimersiana to the list, retaining V. galerulata.However, no species of Vesicularia is listed for Italyby Cortini Pedrotti (2006 [2005]). It seems likely thatV. galerulata has never been found in Europe.396. Smith (2004) treated Heterocladium flaccidum as aspecies, but provided only a nomen nudum. He hasmade the necessary new combination above for thischecklist, noting (pers. comm. to Hill) that consultationwith a number of British field bryologists revealedthat, with one exception, none had encounteredintermediates between it and H. heteropterum.397. Heterocladium wulfsbergii is discussed in detail byCrundwell & Smith (2000).398. Iwatsukiella leucotricha was listed for Europe byPodpěra (1954) under the name Habrodon leucotrichus(Mitt.) Perss., with a locality in the southern Urals. Itwas omitted without comment by Corley et al.(1981), but is indicated from the European side ofthe Urals by Dierssen (2001) and Ignatov & Ignatova(2004).399. Alpine and arctic forms of Hylocomium splendens havebeen variously treated. Var. alpinum Schlieph. exLimpr. is mapped by Söderström (1996), but isdoubted and treated as possibly a synonym of var.obtusifolium (Geh.) Paris by Koperski et al. (2000).Var. obtusifolium is commonly treated as a synonymof H. alaskanum (Lesq. & James) Austin, whosedistinguishing characters have been shown by Ross etal. (2001) to be mainly under environmental ratherthan genetic control. It is not clear whether any ofthese forms is sufficiently distinct genetically todeserve taxonomic recognition.400. Herzogiella adscendens (Lindb.) Z.Iwats. & W.B.Schofieldhas been reported from Svalbard (Corley &Crundwell, 1991; Düll, 1985), but the recordwas subsequently rejected (Frisvoll & Elvebakk,1996).401. Myurella julacea var. ciliata (Chal.) Ochyra &Bednarek-Ochyra (syn. M. julacea var. scabrifoliaLindb. ex Limpr.) is treated as distinct in someEuropean countries, but intergrades completely withvar. julacea. It is not recognized here.402. Orthothecium chryseon is incorrectly listed as O.chryseum by Crosby et al. (1999). It was described asHypnum chryseon by Schwägrichen (1804).403. Plagiothecium curvifolium is sometimes treated as avariety (in Europe) or synonym (in North America)(Ireland, 2003) of P. laetum, to which it is undoubtedlyvery close. As a variety, its correct name is P. laetumvar. secundum.404. Plagiothecium denticulatum var. undulatum is treatedin many European lists as a species, P. ruthei, but inNorth America merely as a synonym of P. denticulatum(Ireland, 2003). It is not clearly distinct from var.denticulatum, and we follow Ignatov & Ignatova(2004), Koperski et al. (2000) and Smith (2004) intreating it as a variety.405. Plagiothecium noricum was described from thePinzgau, Austria and was thought by Molendo(but not Limpricht) to intergrade with P. neckeroideum.Grims (1999) listed it but noted that Düll(1992) had thought it a weak species, possiblyonly a variety of P. neckeroideum. It is not recognizedhere.406. Plagiothecium succulentum is a difficult taxon,intergrading on the one hand with P. nemorale(Hemeric, 1989) and on the other with P. cavifolium(Nyholm, 1965). It is also very close to P. platyphyllum.It is retained here, because it cannot satisfactorilybe subordinated to any of these species withouttreating the whole group as a single, rather-broadspecies.407. Plagiothecium svalbardense was described by Frisvollin Frisvoll & Elvebakk (1996).408. Pseudotaxiphyllum laetevirens was raised to speciesrank by Hedenäs (1992a).409. Entodon challengeri occurs in European Russia(Ignatov & Ignatova, 2004) and is the correct namefor the plant formerly known as E. compressus(Iwatsuki & Tan, 2001).410. According to Ochyra & Ireland (2004), Isopterygiumtenerum is absent from Europe but present in sub-Saharan Africa. However, Schumacker in Stewart(1995), Cortini Pedrotti (2006 [2005]) and Gabrielet al. (2005) list it for Italy and the Azores.Isopterygium bottinii was excluded from Sematophyllumby Guerra & Gallego (2005), and is includedhere in I. tenerum.411. Hageniella micans, previously known from Europeand North America, was synonymized with H.pacifica Broth., which occurs in China and Taiwan(Tan & Jia, 1999).412. Sematophyllum adnatum was found in Italy, where it isthought to be an introduction from North America(Brusa in Blockeel et al., 2000).

244 M. O. HILL ET AL.413. Cryphaea lamyana was transferred to the otherwisetropical and Australasian genus Dendrocryphaea byRao (2001).414. Leucodon flagellaris author citation is based onsynonymy in Podpěra (1954).415. Leucodon pendulus, whose main distribution is ineastern Asia, has been found in the Ivanovo OblastNE of Moscow (Ignatov & Ignatova, 2004).416. Leucodon treleasei was treated as a synonym of L.canariensis by Corley et al. (1981) but as a distinctspecies by Hedenäs (1992b).417. Homalia trichomanoides author citation follows that inthe International Code (Greuter et al., 2000) AppendixIIIa, rather than Corley & Crundwell (1991).418. The characters emphasized by Steere (1941) donot justify generic status for Metaneckera. Thereare, especially in Asia, several species of Neckera thathave some or all of those chracters (J. Enroth, pers.obs.).419. Thamnobryum cataractarum was described fromEngland by Hodgetts & Blockeel (1992).420. Thamnobryum fernandesii was described fromMadeira by Sérgio (1981).421. Thamnobryum maderense, signified by Hedenäs(1992b) as a good species, was reduced to a varietyof T. alopecurum by Stech, Ros & Werner (2001) onthe basis of molecular data. We are not fullyconvinced by their argument and prefer to retain T.maderense as a species for the time being.422. Thamnobryum neckeroides was reported from CzechRepublic, Germany and Italy by Mastracci (2003).423. Thamnobryum rudolphianum was described from theAzores by Mastracci (2004).424. The Madeiran endemic Echinodium setigerum was notmentioned by Corley et al. (1981) but was recognizedas a good species by Churchill (1986).425. Cryptoleptodon longisetus was omitted by Corley et al.(1981) but listed as Leptodon longisetus by Düll (1985).It was transferred to Cryptoleptodon by Enroth (1992).426. Synonymy of Isothecium algarvicum follows Enroth &Hedenäs (1993).427. According to Smith (2004), Isothecium myosuroidesvar. brachythecioides and I. holtii, although distinct intheir typical expressions, are linked to I. myosuroidesby intermediate forms.ACKNOWLEDGEMENTSFinancial support for publication of this checklist camefrom the British Bryological Society (through its BequestFund), from English Nature and from the Centre forEcology and Hydrology (CEH). We are grateful to thosemembers of the Steering Committee who are not authors,namely Michele Aleffi, Bill Buck, Klaus Dierssen and JiříVáňa, for wise advice during the project. Francis Rowlandof CEH designed an attractive website. We thank RonPorley (Newbury, England) for encouraging us to realizethe ECCB’s proposal to make a new European checklist.Bernard Goffinet (Connecticut) supplied an electronic copyof the Goffinet & Buck (2004) taxonomic hierarchy. JohnTweddle (London) checked the author abbreviationsagainst IPNI recommendations. We made extensive use ofdata on websites, especially the W 3 MOST database ofMissouri Botanical Garden, and the author query facility ofthe International Plant Names Index. For help onparticular points, we thank Olga Afonina (St Petersburg,Sphagnum), Hans Blom (Bergen, Schistidium), AliciaEderra (Navarra, Cinclidotus), Gerard Dirkse (Nijmegen,Tortella limbata), T.V. Egorova (St Petersburg, Sphagnum),Len Ellis (London, Conardia), Tomas Hallingbäck(Uppsala, various taxa), Patxi Heras & Marta Infante(Vitoria-Gasteiz, Clasmatodon), Diana Horton (Iowa,Encalypta), Juan Antonio Jiménez (St Louis, Didymodon),Eva Maier (Geneva, Grimmia), Rayna Natcheva (Lund,Bulgarian mosses), Ron Pursell (Pennsylvania, Fissidens),Cecília Sérgio (Lisbon, Portuguese and Macaronesianmosses), Tony Smith (north Wales, British mosses), AlainVanderpoorten (Liège, Macaronesian mosses). Holyoak’sBryum research was supported by the HIGH LATRESOURCE under the EC-funded IHP programme,Plantlife International, and facilitated by Jeff Duckett(London). Finally, we thank two anonymous referees forhelpful and detailed comments, which have resulted inmany corrections and improvements.TAXONOMIC ADDITIONS and CHANGES: Heterocladiumflaccidum (Schimp.) A.J.E.Sm., stat. et comb. nov.(Heterocladium heteropterum var. flaccidum Schimp.);Hypnum cupressiforme var. heseleri (Ando & Higuchi)M.O.Hill, stat. et comb. nov. (Hypnum heseleri Ando &Higuchi); Pelekium atlanticum (Hedenäs) Hedenäs, comb.nov. (Thuidium atlanticum Hedenäs); Pohlia flexuosa var.pseudomuyldermansii (Arts, Nordhorn-Richter &A.J.E.Sm.) A.J.E.Sm., comb. nov. (Pohlia muyldermansiivar. pseudomuyldermansii Arts, Nordhorn-Richter &A.J.E.Sm.); Weissia6mittenii (Bruch & Schimp.) Mitt.emend. A.J.E.Sm., stat. nov. (Weissia mittenii (Bruch &Schimp.) Mitt.).REFERENCESAfonina OM. 2004. 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MOSSES OF EUROPE AND MACARONESIA 253planinervia, 2/5rothii, 2/7rupestris, 2/12sinuosa, 2/13Anisotheciumgrevilleanum, 68/4humile, 68/7palustre, 58/2rigidulum, 68/7rubrum, 68/12rufescens, 68/8schreberianum, 68/9staphylinum, 68/10vaginale, 68/3varium, 68/12Anoectangium, 79aestivum, 79/1angustifolium, 79/1handelii, 79/2, Ann141hornschuchianum, 89/1schliephackei, 89/2sendtnerianum, 89/3taeniatifolium, 89/4tenuinerve, 89/5warburgii, 89/6Anomobryum, 140concinnatum, 140/1, Ann265filiforme, 140/2julaceum, 140/2, Ann265juliforme, 140/2lanatum, 142/6leptostomoides, 140/1, Ann265lusitanicum, 140/3, Ann266Anomodon, 277attenuatus, 277/1longifolius, 277/2rostratus, 277/3rugelii, 277/4tristis, 277/5viticulosus, 277/6Antitrichia, 266californica, 266/1curtipendula, 266/2Aongstroemia, 67longipes, 67/1Aphanorrhegmapatens, 22/1Aplodon, 119wormskioldii, 119/1, Ann32Archidium, 40alternifolium, 40/1Arctoa, 55anderssonii, 55/1fulvella, 55/2hyperborea, 55/3Aschisma, 80carniolicum, 80/1cuynetii, 80/2Astomum, 96crispum, 96/5levieri, 96/4mittenii, Ann161multicapsulare, 96/6Atractylocarpus, 71alpinus, 71/1Atrichum, 5angustatum, 5/1, Ann17crispum, 5/2flavisetum, 5/3, Ann18haussknechtii, 5/3rhystophyllum, Ann17tenellum, 5/4undulatum, 5/5, Ann17, Ann18Aulacomnium, 156androgynum, 156/1palustre, 156/2turgidum, 156/3Barbellastrongylensis, 210/6, Ann360Barbula, 99acuta, 104/1adriatica, 104/8asperifolia, 104/2bicolor, 99/1bolleana, 99/2, Ann139commutata, 99/3, Ann167convoluta, 99/3cordata, 104/6crocea, 99/4cylindrica, 104/13ehrenbergii, 99/2enderesii, 99/5fallax, 104/8gigantea, 104/10hornschuchiana, 113/1indica, 99/6insidiosa, 104/22johansenii, 104/14kneuckeri, 104/2lurida, 104/16reflexa, 104/9revoluta, 113/4rigidula, 104/19sardoa, 99/3, Ann167semilimbata, Ann166sinuosa, 104/21spadicea, 104/22tophacea, 104/25unguiculata, 99/7vinealis, 104/27Bartramia, 135breviseta, 135/3, Ann260halleriana, 135/1ithyphylla, 135/4, Ann260, Ann261pomiformis, 135/2, Ann258stricta, 135/6, Ann256subulata, 135/5Blindia, 37acuta, 37/1caespiticia, 37/2Brachydontium, 38trichodes, 38/1, Ann84Brachymenium, 141cellulare, 142/14commutatum, 141/1, Ann267exile, Ann267, Ann270notarisii, 141/2, Ann268paradoxum, 141/3, Ann269philonotula, 141/4, Ann270Brachytheciastrum, 223collinum, 223/1dieckii, 223/2, Ann383fendleri, 223/3, Ann384olympicum, 223/4, Ann384trachypodium, 223/5vanekii, 223/6velutinum, 223/7Brachythecium, 220albicans, 220/1appleyardiae, 221/1, Ann382campestre, 220/2, Ann374capillaceum, 220/3, Ann375cardotii, 210/3, Ann359cirrosum, 220/4collinum, 223/1coruscum, 220/5, Ann376curtum, 219/4dieckii, 223/2, Ann383dovrense, 219/2erythrorrhizon, 220/6, Ann377fendleri, 223/3geheebii, 220/7glaciale, 219/2, Ann372glareosum, 220/8groenlandicum, 220/5, Ann376laetum, 220/9, Ann379latifolium, 219/3mildeanum, 220/10oedipodium, 219/4olympicum, 223/4ornellanum, 219/5oxycladon, 220/9, Ann379percurrens, 220/11, Ann380plumosum, 219/6populeum, 219/7reflexum, 219/8rivulare, 220/12rotaeanum, 220/3, Ann375rutabulum, 220/13, Ann381ryanii, 220/2, Ann374salebrosum, 220/14

254 M. O. HILL ET AL.salteri, 223/2, Ann383starkei, 219/9thedenii, Ann378tommasinii, 220/15trachypodium, 223/5turgidum, 220/16udum, 220/10umbilicatum, Ann373vanekii, 223/6velutinum, 223/7Braunia, 131alopecura, 131/1Breidleria, 228pratensis, 228/1Breutelia, 136azorica, 136/1chrysocoma, 136/2Brotherella, 258lorentziana, 258/1Bruchia, 52flexuosa, 52/1trobasiana, 52/1vogesiaca, 52/2Bryhnia, 217novae-angliae, 217/1, Ann370scabrida, 217/1, Ann370Bryobrittonia, 16longipes, 16/1Bryoerythrophyllum, 100alpigenum, 100/1caledonicum, 100/2campylocarpum, 100/3, Ann168ferruginascens, 100/4inaequalifolium, 100/5lusitanicum, 100/3machadoanum, 100/3recurvirostrum, 100/6rubrum, 100/7Bryohaplocladiumangustifolium, 192/1microphyllum, 192/2virginianum, 192/3Bryoxiphium, 28madeirense, 28/1, Ann48norvegicum, 28/2Bryum, 142acutiforme, 142/10aeneum, 142/41, Ann293affine, 142/15algovicum, 142/1, Ann276alpinum, 142/2, Ann274, Ann291amblyodon, 142/4apiculatum, 142/3, Ann277archangelicum, 142/4, Ann296arcticum, 142/5argenteum, 142/6, Ann279, Ann280axel-blyttii, 142/10badium, 142/9balticum, 142/19barnesii, 142/19bicolor, 142/19, Ann287bimum, 142/43, Ann295blindii, 142/7, Ann281bornholmense, 142/8, Ann282bryoides, 142/5caespiticium, 142/9, Ann283calophyllum, 142/10canariense, 142/11, Ann284capillare, 142/12castaneum, 142/43caucasicum, 142/13, Ann285cavifolium, 142/39cellulare, 142/14, Ann269cernuum, 142/55chibinense, 142/23colombii, Ann274comense, 142/9, Ann283compactum, 142/1concinnatum, 140/1creberrimum, 142/15crispulum, 142/39cruegeri, 142/3cryophilum, 142/16curvatum, 142/4cuspidatum, 142/15cyclophyllum, 142/17demaretianum, 142/18, Ann286demissum, 143/1dichotomum, 142/19dixonii, 142/20donianum, 142/21dunense, 142/19duvalii, 142/59dyffrynense, 142/22, Ann287ekstamii, Ann272elegans, 142/23erythrocarpum, 142/51excurrens, 142/19filiforme, 140/2flaccidum, 142/37funckii, 142/24funkii, 142/24geheebii, Ann274gemmiferum, 142/25gemmilucens, 142/26gemmiparum, 142/27, Ann275gerwigii, Ann275globosum, 142/60icodense, 142/53imbricatum, 142/4inclinatum, 142/4intermedium, 142/28, Ann274julaceum, 140/2juliforme, 140/2klinggraeffii, 142/29knowltonii, 142/30kunzei, 142/31, Ann283, Ann288labradorense, Ann272lacustre, 142/30laevifilum, 142/37lanatum, 142/6lawersianum, 142/5leptostomum, 140/2lisae, 142/15lonchocaulon, 142/42, Ann294longisetum, 142/32mamillatum, 142/58marratii, 142/33mesodon, Ann274microerythrocarpum, 142/51mildeanum, 142/34miniatum, 142/35, Ann289minii, 142/36, Ann290moravicum, 142/37muehlenbeckii, 142/38, Ann291murmanicum, Ann274neodamense, 142/39, Ann292nitens, 142/3nitidulum, 142/28, Ann274obconicum, 142/42, 142/53oblongum, 142/40, Ann281obovatum, 142/21obtusifolium, 142/16oelandicum, 142/58oeneum, 142/41, Ann293ontariense, 144/1pachyloma, 142/21pallens, 142/41, Ann106, Ann279pallescens, 142/42, Ann294pamirense, Ann271platyloma, 142/12provinciale, 142/11, Ann284pseudotriquetrum, 142/43, Ann292, Ann295purpurascens, 142/5pyriferum, 142/56, Ann299radiculosum, 142/44riparium, 142/45roseum, 144/2rubens, 142/46ruderale, 142/47rufifolium, 142/12rutilans, 142/41salinum, 142/48, Ann296sauteri, 142/49savicziae, Ann271, Ann272schleicheri, 142/50, Ann297splachnoides, 142/14, Ann269stenotrichum, 142/4stirtonii, 142/23subapiculatum, 142/51subelegans, 142/37, 142/41

MOSSES OF EUROPE AND MACARONESIA 255subneodamense, 142/39syriacum, 142/54tenuisetum, 142/52teres, Ann274torquescens, 142/53, Ann298tortifolium, 142/17turbinatum, 142/54uliginosum, 142/55, Ann271, Ann272umbratum, Ann272validicostatum, 142/12valparaisense, 142/56, Ann299ventricosum, 142/43vermigerum, Ann273veronense, 142/6, Ann279versicolor, 142/19violaceum, 142/57warneum, 142/58weigelii, 142/59wrightii, 142/60zemliae, Ann272zieri, 143/2, Ann300Buckiellaundulata, 254/13Bucklandiellaaffinis, 33/10elliptica, 33/11heterosticha, 33/12himalayana, 33/13lamprocarpa, 33/14lusitanica, 33/15macounii, 33/16microcarpa, 33/17obtusa, 33/18sudetica, 33/19Buxbaumia, 13aphylla, 13/1viridis, 13/2Callialariacurvicaulis, 176/1Callicladium, 229haldanianum, 229/1Calliergon, 186cordifolium, 186/1giganteum, 186/2megalophyllum, 186/3obtusifolium, 186/4orbiculare-cordatum, Ann337richardsonii, 186/4, Ann337, Ann338sarmentosum, 191/5stramineum, 190/1trifarium, 183/4Calliergonella, 230cuspidata, 230/1lindbergii, 230/2Calomnion, 159complanatum, 159/1, Ann319Calymperes, 76erosum, 76/1sommieri, 76/1Calyptrochaeta, 162apiculata, 162/1Camptotheciumlutescens, 224/2Campyliadelphus, 173chrysophyllus, 173/1elodes, 173/2polygamus, 177/4stellatus, 174/4Campylidiumcalcareum, 231/1sommerfeltii, 231/3Campylium, 174arcticum, 177/2calcareum, 231/1chrysophyllum, 173/1elodes, 173/2halleri, 231/2hispidulum, 231/3laxifolium, 174/1, Ann328longicuspis, 174/2polygamum, 177/4protensum, 174/3radicale, 171/2sommerfeltii, 231/3stellatum, 174/4Campylophyllum, 231calcareum, 231/1halleri, 231/2hispidulum, 231/3, Ann388sommerfeltii, 231/3Campylopus, 72atrovirens, 72/1brevipilus, 72/2carreiroanus, 72/15cygneus, 72/3eximius, 72/8, Ann135flaccidus, 72/4flexuosus, 72/5fragilis, 72/6gracilis, 72/7incrassatus, 72/8, Ann135introflexus, 72/9madeirensis, 46/8marginatulus, 46/8mildei, 72/10oerstedianus, 72/10pilifer, 72/11pyriformis, 72/12schimperi, 72/13schwarzii, 72/7setaceus, 72/3setifolius, 72/14shawii, 72/15subporodictyon, 73/3subulatus, 72/16Campylostelium, 35pitardii, 35/1, Ann83saxicola, 35/2strictum, 35/3Catoscopium, 133nigritum, 133/1Ceratodon, 42antarcticus, Ann111conicus, 42/1corsicus, 42/2heterophyllus, Ann111purpureus, 42/2, Ann111, Ann112stenocarpus, 42/2Cheilothela, 43chloropus, 43/1Chenialeptophylla, 108/1rhizophylla, 108/1Cinclidium, 151arcticum, 151/1latifolium, 151/2stygium, 151/3subrotundum, 151/4Cinclidotus, 101aquaticus, 101/1bistratosus, Ann169confertus, 101/2, Ann170danubicus, 101/3fontinaloides, 101/4mucronatus, 103/2nyholmiae, Ann169pachylomoides, 101/5riparius, 101/6, Ann170vivesii, 101/7, Ann171Cirriphyllum, 214cirrosum, 220/4crassinervium, 214/1germanicum, 213/8piliferum, 214/2reichenbachianum, 219/1tenuicaule, 213/8tenuinerve, 220/15tommasinii, 220/15velutinoides, 219/1Claopodium, 278whippleanum, 278/1Clasmatodonparvulus, Ann361Cleistocarpidium, 44palustre, 44/1Climacium, 170dendroides, 170/1Cnestrum, 56alpestre, 56/1glaucescens, 56/2schisti, 56/3

256 M. O. HILL ET AL.Codonoblepharonforsteri, 130/3Codriophorusacicularis, 33/6aquaticus, 33/7fascicularis, 33/8hespericus, 33/9Conardia, 175compacta, 175/1Conostomum, 137tetragonum, 137/1Coscinodon, 29cribrosus, 29/1humilis, 29/2, Ann49Crassiphyllumfernandesii, 271/4Cratoneuron, 176commutatum, 182/1curvicaule, 176/1decipiens, 182/2falcatum, 182/3filicinum, 176/2Crossidium, 102aberrans, 102/1crassinerve, 102/2davidai, 102/3, Ann172geheebii, 102/4, Ann173laevipilum, 102/5, Ann174laxefilamentosum, 102/6, Ann175seriatum, Ann221squamiferum, 102/7Crumialatifolia, Ann140Cryphaea, 264heteromalla, 264/1lamyana, 265/1, Ann413Cryptoleptodon, 273longisetus, 273/1, Ann425Ctenidium, 232molluscum, 232/1, Ann389procerrimum, 235/12Cyclodictyon, 167laetevirens, 167/1, Ann323Cynodontium, 57alpestre, 56/1asperifolium, 57/1, Ann119bruntonii, 57/2fallax, 57/3, Ann119gracilescens, 57/4jenneri, 57/5polycarpon, 57/6, Ann120strumiferum, 57/7suecicum, 57/8tenellum, 57/9torquescens, 57/9Cyrto-hypnumatlanticum, 201/1minutulum, 201/2montei, 202/1, Ann350Cyrtomnium, 152hymenophylloides, 152/1hymenophyllum, 152/2Daltonia, 163splachnoides, 163/1, Ann322stenophylla, 163/2, Ann322Dendrocryphaea, 265lamyana, 265/1Desmatodon, 117bogosicus, 117/4cernuus, 117/8guepinii, 117/11heimii, 105/1latifolius, 117/12laureri, 117/16leucostoma, 117/17meridionalis, 117/19, Ann223oxneri, 117/25randii, 117/25systylius, 117/31ucrainicus, 117/33wilczekii, 117/27, Ann225Dialytrichia, 103fragilifolia, 103/1, Ann176mucronata, 103/2Dichelyma, 168capillaceum, 168/1falcatum, 168/2Dichodontium, 58flavescens, 58/1palustre, 58/2, Ann121pellucidum, 58/3Dicranella, 68campylophylla, 68/1, Ann126cerviculata, 68/2crispa, 68/3curvata, 68/11grevilleana, 68/4heteromalla, 68/5howei, 68/6humilis, 68/7palustris, 58/2riparia, 61/4rufescens, 68/8schreberiana, 68/9secunda, 68/11staphylina, 68/10subulata, 68/11varia, 68/12Dicranodontium, 73asperulum, 73/1circinatum, 73/4denudatum, 73/2subporodictyon, 73/3, Ann136uncinatum, 73/4Dicranoweisia, 59, Ann122cirrata, 59/1compacta, 59/2crispula, 59/3Dicranum, 69acutifolium, 69/8affine, 69/13angustum, 69/16, Ann132bergeri, 69/13, Ann131bonjeanii, 69/1brevifolium, 69/9canariense, 69/23, Ann134congestum, 69/14crassifolium, 69/2, Ann128dispersum, 69/10, Ann130drummondii, 69/11elatum, 69/11elongatum, 69/19, Ann133flagellare, 69/27flexicaule, 69/14fragilifolium, 69/20fulvum, 69/24fuscescens, 69/15, Ann133groenlandicum, 69/21laevidens, 69/22, Ann132leioneuron, 69/3majus, 69/4montanum, 69/28muehlenbeckii, 69/17neglectum, 69/18palustre, 69/1polysetum, 69/5, Ann106robustum, 69/11rugosum, 69/5scoparium, 69/6scottianum, 69/25, Ann79, Ann134sendtneri, 69/19, Ann133spadiceum, 69/18spurium, 69/12strictum, 69/29subporodictyon, 73/3tauricum, 69/29transsylvanicum, 69/7, Ann129undulatum, 69/13, 69/1, 69/5, Ann131viride, 69/26Didymodon, 104aaronis, 104/3acutus, 104/1asperifolius, 104/2, Ann180australasiae, 104/3, Ann181barbuloides, 104/22bistratosus, 104/4, Ann182bosniacus, 104/25brachyphyllus, 104/5, Ann183cordatus, 104/6erosus, 104/7, Ann184fallax, 104/8, Ann180

MOSSES OF EUROPE AND MACARONESIA 257ferrugineus, 104/9giganteus, 104/10, Ann180glaucus, 104/11icmadophilus, 104/12incrassatus, 104/3insulanus, 104/13, Ann185johansenii, 104/14lamyanus, 104/15, Ann185luridus, 104/16mamillosus, 104/19, Ann186maximus, 104/17, Ann180nicholsonii, 104/18, Ann185reedii, Ann178rigidulus, 104/19, Ann186rufus, 104/2sicculus, 104/20, Ann187sinuosus, 104/21soaresii, Ann179spadiceus, 104/22, Ann180subandreaeoides, 104/23, Ann188tectorum, Ann178tomaculosus, 104/24tophaceus, 104/25, Ann180trifarius, 104/16trivialis, 104/26umbrosus, 104/26, Ann189vinealis, 104/27, Ann185zetterstedtii, 104/22Diobelonellapalustris, 58/2Diphyscium, 14foliosum, 14/1Discelium, 25nudum, 25/1Distichium, 45capillaceum, 45/1hagenii, 45/2inclinatum, 45/3Distichophyllum, 164carinatum, 164/1Ditrichum, 46cornubicum, 46/1crispatissimum, 46/3, Ann113cylindricum, 51/1difficile, Ann114flexicaule, 46/2giganteum, 46/3gracile, 46/3, Ann113heteromallum, 46/4homomallum, 46/4lineare, 46/5pallidum, 46/6plumbicola, 46/7punctulatum, 46/8, Ann114pusillum, 46/9subulatum, 46/10tortile, 46/9vaginans, 46/5zonatum, 46/11Drepaniumrecurvatum, 235/13Drepanocladus, 177aduncus, 177/1arcticus, 177/2badius, 188/1brevifolius, 183/2capillifolius, 177/3cossonii, 189/1exannulatus, 191/1fluitans, 191/2h-schulzei, 191/2intermedius, 189/1lapponicus, 187/1longifolius, 177/3lycopodioides, 183/3polycarpos, 177/1polygamus, 177/4procerus, 191/3pseudostramineus, 191/4revolvens, 189/2sendtneri, 177/5simplicissimus, 177/1sordidus, 177/6stagnatus, 177/1tenuinervis, 177/6trichophyllus, 191/6tundrae, 191/7uncinatus, 184/3vernicosus, 187/2Dryptodonanomalus, 30/3contortus, 30/18decipiens, 30/10elongatus, 30/14funalis, 30/15hartmanii, 30/17incurvus, 30/13, Ann56muehlenbeckii, 30/24orbicularis, 30/26patens, 30/31pulvinatus, 30/30torquatus, 30/35trichophyllus, 30/36Echinodium, 272madeirense, 272/4prolixum, 272/1renauldii, 272/2setigerum, 272/3, Ann424spinosum, 272/4Encalypta, 17affinis, 17/11, Ann33alpina, 17/3brevicolla, 17/12brevipes, 17/13ciliata, 17/14, Ann28commutata, Ann29intermedia, 17/5, Ann29longicolla, 17/10longipes, 16/1macounii, 17/11microstoma, 17/15mutica, 17/4obovatifolia, 17/6, Ann30procera, 17/1rhaptocarpa, 17/7, Ann31serbica, Ann28spathulata, 17/8streptocarpa, 17/2trachymitria, 17/7, Ann31vulgaris, 17/9Entodon, 257challengeri, 257/1, Ann409cladorrhizans, 257/2compressus, 257/1, Ann409concinnus, 257/3schleicheri, 257/4Entosthodon, 18attenuatus, 18/1convexus, 18/6curvisetus, 20/1durieui, 18/2, Ann37ericetorum, 18/5fascicularis, 18/10, Ann35handelii, Ann42hungaricus, 18/3, Ann38krausei, 18/4, Ann39maroccanus, 18/3mouretii, 18/11, Ann41muhlenbergii, 18/7mustaphae, 18/2obtusus, 18/5pallescens, 18/2physcomitroides, 18/2pulchellus, 18/8schimperi, 18/9, Ann40subpallescens, Ann34templetonii, 18/1Ephemerum, 81cohaerens, 81/1hibernicum, 81/2, Ann142minutissimum, 81/3, Ann143recurvifolium, 81/4serratum, 81/5, Ann143sessile, 81/6spinulosum, 81/7, Ann144stellatum, 81/8Epipterygium, 145tozeri, 145/1Eriopusapiculatus, 162/1Eucladium, 82

258 M. O. HILL ET AL.verticillatum, 82/1, Ann145Eurhynchiastrum, 222pulchellum, 222/1Eurhynchium, 208angustirete, 208/1crassinervium, 214/1flotowianum, 219/1hians, 215/1meridionale, 207/1praelongum, 216/1pulchellum, 222/1pumilum, 215/2schleicheri, 215/3speciosum, 215/4stokesii, 216/1striatulum, 207/2striatum, 208/2Fabronia, 226ciliaris, 226/1pusilla, 226/2Fissidens, 41adianthoides, 41/1algarvicus, 41/18arcticus, 41/12, Ann93arnoldii, 41/13, Ann99asplenioides, 41/10azoricus, 41/2bambergeri, 41/31, Ann108bryoides, 41/14, Ann94, Ann95, Ann97,Ann102, Ann103, Ann105, Ann107celticus, 41/32coacervatus, 41/15crassipes, 41/16crispus, 41/17, Ann96, Ann108cristatus, 41/3curnovii, 41/14, Ann94curvatus, 41/18decipiens, 41/3dubius, 41/3, Ann88exiguus, 41/26, 41/31, Ann92, Ann94, Ann103exilis, 41/33fontanus, 41/11gracilifolius, 41/19grandifrons, 41/4gymnandrus, 41/20, Ann97haraldii, 41/31herzogii, 41/17incurvus, 41/31, Ann94jansenii, 41/21, Ann98kosaninii, Ann92, Ann94limbatus, 41/17, Ann96, Ann108luisieri, 41/5, Ann89marginatulus, 41/31, Ann109microstictus, 41/22, Ann99minutulus, 41/17, Ann96monguillonii, 41/23mouretii, 41/16nobreganus, 41/24, Ann100osmundoides, 41/6ovatifolius, 41/25, Ann101pallidicaulis, 41/9papillosus, 41/34, Ann110perssonii, 41/25, Ann101polyphyllus, 41/7, Ann90pusillus, 41/26, Ann102, Ann103rivularis, 41/27rufulus, 41/28, Ann98serratus, 41/34, Ann110serrulatus, 41/8, Ann91sublimbatus, 41/29, Ann104sublineaefolius, 41/30, Ann100taxifolius, 41/9viridulus, 41/31, Ann92, Ann94, Ann103,Ann105, Ann107, Ann108, Ann109Fontinalis, 169antipyretica, 169/1, Ann324, Ann325bryhnii, 169/1, Ann325dalecarlica, 169/2dichelymoides, 169/3, Ann326hypnoides, 169/4kindbergii, 169/1squamosa, 169/5, Ann324Forsstroemiacanariensis, 275/1Funaria, 196hybrida, Ann35aequidens, 19/1algieriensis, 18/9anomala, Ann42arctica, 19/2attenuata, 18/1calcarea, 18/7convexa, 18/6curviseta, 20/1dentata, 18/7durieui, 18/9, Ann40fascicularis, 18/10fritzei, Ann39handelii, Ann42hibernica, 18/7hungarica, 18/3hygrometrica, 19/3, Ann35krausei, 18/4maireana, Ann42mediterranea, 18/7microstoma, 19/4mouretii, 18/11muhlenbergii, 18/7obtusa, 18/5pallescens, 18/2pulchella, 18/8pustulosa, 18/6subpallescens, Ann34Funariella, 20curviseta, 20/1Geheebiagigantea, 104/10Gigaspermum, 26mouretii, 26/1Glyphomitrium, 60daviesii, 60/1Goniomitrium, 21acuminatum, Ann44seroi, 21/1, Ann44Gradsteiniatorrenticola, 209/5, Ann356Grimmia, 30affinis, 30/21alpestris, 30/1, Ann50, Ann63anodon, 30/2anomala, 30/3apiculata, 30/16arenaria, 30/4, Ann52atrata, 30/5austrofunalis, 30/36, Ann65britannica, 30/36, Ann66caespiticia, 30/6capillata, 30/7, Ann53crassifolia, 30/34, Ann64crinita, 30/8, Ann53curvata, 30/31curviseta, 30/9, Ann54decipiens, 30/10dissimulata, 30/11, Ann55donniana, 30/12, Ann52, Ann68elatior, 30/13, Ann56elongata, 30/14funalis, 30/15, Ann57fuscolutea, 30/16grisea, 30/32, Ann62hartmanii, 30/17holleri, 30/16incurva, 30/18laevigata, 30/19limprichtii, 30/2, Ann51lisae, 30/20, Ann58longirostris, 30/21maritima, Ann79meridionalis, 30/36, Ann67meteorae, 30/25mollis, 30/22montana, 30/23muehlenbeckii, 30/24, Ann59nutans, 30/25orbicularis, 30/26ovalis, 30/27pilosissima, 30/23pitardii, 35/1plagiopodia, 30/28poecilostoma, 30/29, Ann59, Ann64pulvinata, 30/30, Ann60

MOSSES OF EUROPE AND MACARONESIA 259pyrenaica, 30/6ramondii, 30/31reflexidens, 30/32, Ann61, Ann62, Ann63retracta, 30/20, Ann58robusta, Ann66sessitana, 30/32subsulcata, 30/32, Ann63sudetica, 30/12teretinervis, 30/33tergestina, 30/34, Ann51, Ann59, Ann64torquata, 30/35trichophylla, 30/36, Ann59, Ann66, Ann67triformis, 30/37, Ann68ungeri, 30/38, Ann69unicolor, 30/39Guembelialaevigata, 30/19longirostris, 30/21ovalis, 30/27tergestina, 30/34Gymnostomum, 83aeruginosum, 83/1, Ann147boreale, 83/2calcareum, 83/3lanceolatum, 83/4, Ann146mosis, Ann146recurvirostrum, 85/1viridulum, 83/5Gyroweisia, 84reflexa, 84/1tenuis, 84/2Habrodon, 240leucotrichus, Ann398perpusillus, 240/1Hageniella, 262micans, 262/1, Ann411pacifica, Ann411Hamatocaulis, 187lapponicus, 187/1vernicosus, 187/2Haplocladium, 192angustifolium, 192/1microphyllum, 192/2virginianum, 192/3Haplodontiumnotarisii, 141/2Haplohymeniumtriste, 277/5Hedwigia, 132ciliata, 132/1, Ann253, Ann254integrifolia, 132/2stellata, 132/3, Ann253, Ann255Hedwigidiumintegrifolium, 132/2Helicodontium, 212capillare, 212/1, Ann363italicum, 212/1, Ann363Helodium, 200blandowii, 200/1Hennediella, 105antarctica, Ann190heimii, 105/1macrophylla, 105/2, Ann190stanfordensis, 105/3Herzogiella, 250adscendens, Ann400seligeri, 250/1striatella, 250/2turfacea, 250/3Heterocladium, 241dimorphum, 241/1flaccidum, 241/2, Ann396heteropterum, 241/3, Ann396wulfsbergii, 241/4, Ann397Heterophyllium, 259affine, 259/1Hilpertia, 106velenovskyi, 106/1Homalia, 269besseri, 270/1complanata, 270/3lusitanica, 269/1subrecta, 269/1trichomanoides, 269/2, Ann417webbiana, 269/3Homalothecium, 224ureum, 224/1geheebii, 220/7lutescens, 224/2philippeanum, 224/3sericeum, 224/4Homomallium, 233incurvatum, 233/1Hookeria, 165lucens, 165/1Hydrogrimmiamollis, 30/22Hygroamblystegium, 178fluviatile, 178/1humile, 178/2tenax, 178/3varium, 178/4, Ann329, Ann330Hygrohypnelladuriuscula, 179/4ochracea, 179/10polaris, 179/11Hygrohypnum, 179alpestre, 179/1alpinum, 179/2cochlearifolium, 179/3dilatatum, 179/4duriusculum, 179/4eugyrium, 179/5luridum, 179/6, Ann332micans, 262/1molle, 179/7montanum, 179/8norvegicum, 179/9ochraceum, 179/10polare, 179/11mithii, 179/12, Ann331styriacum, 179/13subeugyrium, 179/14Hylocomiastrum, 244pyrenaicum, 244/1umbratum, 244/2Hylocomium, 245alaskanum, Ann399brevirostre, 246/1pyrenaicum, 244/1splendens, 245/1, Ann399umbratum, 244/2Hymenolomacompactum, 59/2crispulum, 59/3Hymenostomummicrostomum, 96/1squarrosum, 96/10Hymenostylium, 85insigne, 85/1recurvirostrum, 85/1, Ann148Hyocomium, 234armoricum, 234/1Hyophila, 86involuta, 86/1lusitanica, 100/3machadoana, 100/3treleasei, 95/2, Ann159Hypnum, 235aemulans, Ann392andoi, 235/1bambergeri, 235/2callichroum, 235/3cupressiforme, 235/4, Ann390, Ann391dolomiticum, 235/14fertile, 235/5hamulosum, 235/6, Ann392heseleri, 235/4, Ann391holmenii, 235/7, Ann393imponens, 235/8jutlandicum, 235/9lacunosum, 235/4, Ann390lindbergii, 230/2, Ann386mammillatum, 235/1pallescens, 235/10plicatulum, 235/11pratense, 228/1, Ann386procerrimum, 235/12recurvatum, 235/13resupinatum, 235/4revolutum, 235/14

260 M. O. HILL ET AL.sauteri, 235/15subimponens, 235/16, Ann394thedenii, Ann378uncinulatum, 235/17vaucheri, 235/18Hypopterygium, 160muelleri, 160/1, Ann320tamarisci, 160/1, Ann320Imbribryum, 142alpinum, 142/2muehlenbeckii, 142/38Indusiella, 31thianschanica, 31/1, Ann70Isopterygiopsis, 251alpicola, 251/1muelleriana, 251/2pulchella, 251/3Isopterygium, 260alpicola, 251/1bottinii, 260/1, Ann410elegans, 256/1muellerianum, 251/2pulchellum, 251/3tenerum, 260/1, Ann410Isothecium, 275algarvicum, 275/1, Ann426alopecuroides, 275/2atlanticum, 275/1canariense, 275/1circinans, 275/2holtii, 275/3, Ann427myosuroides, 275/4, Ann427myurum, 275/2striatulum, 207/2viviparum, 275/2Iwatsukiella, 242leucotricha, 242/1, Ann398Jaffueliobryum, 32latifolium, 32/1, Ann72Kiaeria, 61blyttii, 61/1falcata, 61/2glacialis, 61/3riparia, 61/4starkei, 61/5Kindbergia, 216praelonga, 216/1Lepidopilumvirens, 166/2Leptobarbula, 87berica, 87/1Leptobryum, 125pyriforme, 125/1Leptodictyum, 180humile, 178/2kochii, 178/2kurdicum, Ann333riparium, 180/1Leptodon, 274longisetus, 273/1, Ann425smithii, 274/1Leptodontium, 107flexifolium, 107/1gemmascens, 107/2styriacum, 107/3Leptophascum, 108leptophyllum, 108/1Leptotheca, 158gaudichaudii, 158/1, Ann318Lescuraea, 193incurvata, 196/2mutabilis, 193/1patens, 196/3plicata, 198/1radicosa, 196/4saviana, 196/5saxicola, 193/2secunda, 193/3, Ann341Leskea, 194polycarpa, 194/1Leskeellaincrassata, 197/2, Ann346nervosa, 197/2Leucobryum, 74albidum, 74/1, 74/3, Ann137glaucum, 74/2juniperoideum, 74/3, Ann137minus, 74/3Leucodon, 267canariensis, 267/1, Ann416flagellaris, 267/2, Ann414immersus, 267/3pendulus, 267/4, Ann415sciuroides, 267/5treleasei, 267/6, Ann416Limneriaviridula, 104/22Limprichtiacossonii, 189/1intermedia, 189/1revolvens, 189/2Lindbergia, 195brachyptera, 195/1, Ann342Loeskeobryum, 246brevirostre, 246/1Loeskypnum, 188badium, 188/1Meesia, 126hexasticha, 126/1longiseta, 126/2triquetra, 126/3uliginosa, 126/4Metaneckeramenziesii, 270/6Metzlerellaalpina, 71/1Metzleriaalpina, 71/1Microbryum, 109curvicollum, 109/1, Ann192davallianum, 109/2, Ann193floerkeanum, 109/3fosbergii, 109/4, Ann194longipes, 109/5piptocarpum, 109/1rectum, 109/6starckeanum, 109/7, Ann193, Ann195vlassovii, 111/2Microcampylopus, 75laevigatus, 75/1Micromitrium, 88tenerum, 88/1Microthuidiumminutulum, 201/2Mielichhoferia, 146caucasia, 142/13elongata, 146/1, Ann303himalayana, 142/13mielichhoferi, 146/2mielichhoferiana, 146/2, Ann303paradoxa, 141/3, Ann269Mniobryumdelicatulum, 147/26pulchellum, 147/19wahlenbergii, 147/28Mnium, 149ambiguum, 149/5, Ann313blyttii, 149/7heterophyllum, 149/8hornum, 149/1lycopodioides, 149/5, Ann313marginatum, 149/6spinosum, 149/2spinulosum, 149/3stellare, 149/9thomsonii, 149/4Molendoa, 89hornschuchiana, 89/1schliephackei, 89/2sendtneriana, 89/3, Ann141taeniatifolia, 89/4tenuinervis, 89/5warburgii, 89/6Myrinia, 225pulvinata, 225/1Myurella, 252julacea, 252/1, Ann401sibirica, 252/2tenerrima, 252/3Myurium, 276hebridarum, 276/1

MOSSES OF EUROPE AND MACARONESIA 261hochstetteri, 276/1Myuroclada, 218maximowiczii, 218/1, Ann371Neckera, 270besseri, 270/1cephalonica, 270/2complanata, 270/3crispa, 270/4elegans, 270/5fontinaloides, 270/9intermedia, 270/5laeviuscula, 270/5longipedunculata, 273/1menziesii, 270/6oligocarpa, 270/7pennata, 270/8philippeana, 270/9pseudopennata, 270/4pumila, 270/9subrecta, 269/1webbiana, 269/3Niphotrichumcanescens, 33/2elongatum, 33/3ericoides, 33/4panschii, 33/5Nobregaea, 211latinervis, 211/1, Ann362Ochyraea, 181alpestris, 179/1cochlearifolia, 179/3montana, 179/8norvegica, 179/9smithii, 179/12tatrensis, 181/1, Ann331Octoblepharumalbidum, Ann138Octodicerasfontanum, 41/11Oedipodiella, 27australis, 27/1, Ann47Oedipodium, 3griffithianum, 3/1Oligotrichum, 6hercynicum, 6/1Oncophorus, 62compactus, 62/3elongatus, 62/1, Ann123riparius, 61/4virens, 62/2wahlenbergii, 62/3Oreas, 63martiana, 63/1Oreoweisia, 64bruntonii, 57/2serrulata, 64/1torquescens, 64/1Orthodicranumflagellare, 69/27fulvum, 69/24montanum, 69/28scottianum, 69/25tauricum, 69/29Orthodontium, 157australe, 157/2, Ann317gracile, 157/1lineare, 157/2, Ann317pellucens, 157/3Orthogrimmiaalpestris, 30/1arenaria, 30/4caespiticia, 30/6donniana, 30/12montana, 30/23sessitana, 30/32triformis, 30/37Orthotheciellavaria, 178/4Orthothecium, 253chryseon, 253/1, Ann402intricatum, 253/2lapponicum, 253/3rufescens, 253/4strictum, 253/5Orthotrichum, 128acuminatum, 128/34affine, 128/35alpestre, 128/5anomalum, 128/1arcangelianum, 128/31, Ann241australe, 128/26, Ann239callistomum, 128/28casasianum, 128/6, Ann231caucasicum, 128/39, Ann244consimile, 128/7crenulatum, 128/8, Ann232cupulatum, 128/2diaphanum, 128/9elegans, 128/40, Ann245fastigiatum, 128/35flowersii, Ann232gymnostomum, 128/29handiense, 128/10, Ann233hispanicum, 128/11, Ann234ibericum, 128/36, Ann243laevigatum, 128/31lewinskyae, 128/21, Ann236limprichtii, 128/2, Ann230lyellii, 128/37macrocephalum, 128/12, Ann235microcarpum, 128/13obtusifolium, 128/30pallens, 128/14paradoxum, 128/14patens, 128/15pellucidum, 128/3philibertii, 128/16pulchellum, 128/17pumilum, 128/18, Ann106pylaisii, 128/38rivulare, 128/19rogeri, 128/20, Ann238rupestre, 128/32scanicum, 128/21schimperi, 128/22, Ann237shawii, 128/33, Ann242sordidum, 128/39speciosum, 128/40, Ann245, Ann246sprucei, 128/23stellatum, 128/24stramineum, 128/25striatum, 128/41tenellum, 128/26tortidontium, 128/42, Ann247urnaceum, Ann229urnigerum, 128/4vittii, 128/27, Ann240vladikavkanum, 128/43, Ann248Oxyrrhynchium, 215hians, 215/1praelongum, 216/1pumilum, 215/2schleicheri, 215/3speciosum, 215/4swartzii, 215/1Oxystegus, 90hibernicus, 90/1tenuirostris, 90/2Palamocladium, 206euchloron, 206/1Paludella, 127squarrosa, 127/1Palustriella, 182commutata, 182/1decipiens, 182/2falcata, 182/3, Ann334pluristratosa, 182/4, Ann334Paraleptodontium, 110recurvifolium, 110/1Paraleucobryum, 70enerve, 70/1longifolium, 70/2sauteri, 70/3Pelekium, 201atlanticum, 201/1, Ann349minutulum, 201/2muricatulum, Ann349Phascum, 111curvicollum, 109/1cuspidatum, 111/1cuynetii, 80/2

262 M. O. HILL ET AL.floerkeanum, 109/3leptophyllum, 108/1longipes, 109/5piptocarpum, 109/1, Ann192vlassovii, 111/2Philonotis, 138arnellii, 138/3caespitosa, 138/7calcarea, 138/8capillaris, 138/3cernua, 138/1fontana, 138/9hastata, 138/4laxa, Ann262marchica, 138/5, Ann262rigida, 138/2seriata, 138/10tomentella, 138/11uncinata, 138/6, Ann263Physcomitrella, 226hampei, Ann35patens, 22/1, Ann35Physcomitrium, 23acuminatum, 23/2arenicola, 23/1eurystomum, 23/2, Ann35, Ann45maroccanum, 18/3patens, 22/1pyriforme, 23/3sphaericum, 23/4Pictusscoticus, 179/6, Ann332Plagiobryum, 143demissum, 143/1zieri, 143/2Plagiomnium, 154affine, 154/3confertidens, 154/8curvatulum, 154/4, Ann315cuspidatum, 154/1drummondii, 154/2elatum, 154/5ellipticum, 154/6medium, 154/7, Ann315rostratum, 154/10undulatum, 154/9, Ann316Plagiopus, 139oederi, 139/1oederianus, 139/1Plagiothecium, 254berggrenianum, 254/1cavifolium, 254/2, Ann406curvifolium, 254/3, Ann403denticulatum, 254/4, Ann404laetum, 254/5, Ann403latebricola, 254/6neckeroideum, 254/7, Ann405nemorale, 254/8, Ann406noricum, 254/7, Ann405piliferum, 254/9platyphyllum, 254/10, Ann406roeseanum, 254/2ruthei, 254/4, Ann404succulentum, 254/11, Ann406svalbardense, 254/12, Ann407undulatum, 254/13Plasteurhynchium, 207meridionale, 207/1striatulum, 207/2Platydictya, 255confervoides, 171/1jungermannioides, 255/1subtilis, 171/4Platygyrium, 261repens, 261/1Platyhypnidium, 209alopecuroides, 209/2grolleanum, 209/1, Ann354lusitanicum, 209/2mutatum, 209/3, Ann355riparioides, 209/4, Ann355torrenticola, 209/5, Ann356Pleuridium, 47acuminatum, 47/1alternifolium, 47/2nitidum, 48/1palustre, 44/1serrulatum, Ann115subulatum, 47/2Pleurochaete, 91malacophylla, Ann149squarrosa, 91/1Pleurozium, 247schreberi, 247/1Pogonatum, 7aloides, 7/1, Ann20alpinum, 8/1capillare, 7/2dentatum, 7/2, Ann23inflexum, Ann19nanum, 7/3, Ann20neesii, 7/4, Ann19, Ann21urnigerum, 7/5, Ann23Pohlia, 147acuminata, 147/4, Ann288ambigua, 147/4, Ann307andalusica, 147/10andrewsii, 147/11annotina, 147/12apiculata, 142/3atropurpurea, 147/25bolanderi, 147/1, Ann306bulbifera, 147/13camptotrachela, 147/14cruda, 147/2crudoides, 147/3, Ann305cruegeri, 142/3defecta, 147/16drummondii, 147/15elongata, 147/4, Ann307, Ann309erecta, 147/16filum, 147/17flexuosa, 147/18, Ann305, Ann310, Ann311grandiflora, 147/12lescuriana, 147/19longicolla, 147/5ludwigii, 147/20lutescens, 147/21marchica, Ann308melanodon, 147/26muyldermansii, 147/18, Ann311nutans, 147/6, Ann308obtusifolia, 147/7polymorpha, Ann288proligera, 147/22ramannii, Ann304saprophila, 147/8, Ann309schimperi, 147/6, Ann308schleicheri, 147/17scotica, 147/23sphagnicola, 147/9tundrae, 147/24, Ann312turonensis, Ann308vexans, 147/27wahlenbergii, 147/28Polytrichastrum, 8alpinum, 8/1, Ann23formosum, 8/2longisetum, 8/3norwegicum, 8/1pallidisetum, 8/4sexangulare, 8/5, Ann24sphaerothecium, 8/6, Ann24Polytrichum, 9affine, 9/6alpestre, 9/6alpinum, 8/1commune, 9/1, Ann25formosum, 8/2hyperboreum, 9/2jensenii, 9/3juniperinum, 9/4longisetum, 8/3pallidisetum, 8/4perigoniale, 9/1piliferum, 9/5sexangulare, 8/5sphaerothecium, 8/6strictum, 9/6swartzii, 9/7uliginosum, 9/8

MOSSES OF EUROPE AND MACARONESIA 263Pottia, 1176andalusica, 109/4, Ann194bryoides, 112/1caespitosa, 92/1caucasica, Ann217commutata, 109/2conica, 109/2crinita, 117/35cuneifolia, 117/24, Ann224davalliana, 109/2heimii, 105/1intermedia, 117/20lanceolata, 117/15mutica, 109/7pallida, 117/24, Ann224recta, 109/6starckeana, 109/7truncata, 117/32wilsonii, 117/36Pottiopsis, 92caespitosa, 92/1, Ann158Protobryum, 112bryoides, 112/1Pseudephemerum, 48axillare, 48/1nitidum, 48/1Pseudobryum, 155cinclidioides, 155/1Pseudocalliergon, 183angustifolium, 183/1, Ann335brevifolium, 183/2, Ann336lycopodioides, 183/3trifarium, 183/4turgescens, 183/5Pseudocrossidium, 113hornschuchianum, 113/1obtusulum, 113/2, Ann196replicatum, 113/3, Ann197revolutum, 113/4, Ann196Pseudohygrohypnumeugyrium, 179/5subeugyrium, 179/14Pseudoleskea, 196artariae, 196/1brachyclados, Ann344incurvata, 196/2, Ann344patens, 196/3plicata, 198/1radicosa, 196/4, Ann344saviana, 196/5Pseudoleskeella, 197catenulata, 197/1nervosa, 197/2, Ann346papillosa, 197/3rupestris, 197/4, Ann347sibirica, 197/4, Ann347tectorum, 197/5, Ann361Pseudoleskeopsisartariae, 196/1Pseudoscleropodium, 204purum, 204/1Pseudotaxiphyllum, 256elegans, 256/1laetevirens, 256/2, Ann408Psilopilum, 10cavifolium, 10/1laevigatum, 10/2Pterigynandrum, 243filiforme, 243/1Pterogonium, 268gracile, 268/1Pterygoneurum, 114compactum, 114/1, Ann198crossidioides, 114/2, Ann199kozlovii, 114/3lamellatum, 114/4ovatum, 114/5papillosum, 114/6, Ann200sampaianum, 114/7squamosum, 114/8, Ann201subsessile, 114/9Ptilium, 236crista-castrensis, 236/1Ptychodium, 198plicatum, 198/1Ptychomitrium, 36incurvum, 36/1nigrescens, 36/2polyphyllum, 36/3Pylaisia, 237polyantha, 237/1selwynii, 237/2Pylaisiellapolyantha, 237/1selwynii, 237/2Pyramidula, 24algeriensis, 24/1tetragona, 24/1, Ann46Racomitrium, 33aciculare, 33/6affine, 33/10aquaticum, 33/7canescens, 33/2ellipticum, 33/11elongatum, 33/3ericoides, 33/4fasciculare, 33/8hespericum, 33/9, Ann73heterostichum, 33/12himalayanum, 33/13lamprocarpum, 33/14lanuginosum, 33/1lusitanicum, 33/15, Ann74macounii, 33/16microcarpon, 33/17obtusum, 33/18panschii, 33/5sudeticum, 33/19Rhabdoweisia, 65crenulata, 65/1crispata, 65/2, Ann124denticulata, 65/2fugax, 65/3kusenevae, 65/2, Ann124striata, 65/3Rhamphidium, 49purpuratum, 49/1, Ann116Rhizomnium, 153andrewsianum, 153/1gracile, 153/2magnifolium, 153/3pseudopunctatum, 153/4punctatum, 153/5Rhodobryum, 144ontariense, 144/1, Ann301roseum, 144/2spathulatum, 144/1, Ann301Rhynchostegiella, 213bourgaeana, 213/1, Ann364compacta, 175/1curviseta, 213/2durieui, 213/3jacquinii, 213/7, Ann367letourneuxii, 213/2litorea, 213/4, Ann365macilenta, 213/5, Ann366pallidirostris, 215/2pumila, 215/2teesdalei, 213/7, Ann367tenella, 213/6, Ann364, Ann365teneriffae, 213/7, Ann367tenuicaulis, 213/8, Ann368trichophylla, 213/9, Ann369Rhynchostegium, 210alopecuroides, 209/2arcticum, 210/1, Ann357confertum, 210/2, Ann358lusitanicum, 209/2megapolitanum, 210/3, Ann359murale, 210/4riparioides, 209/4rotundifolium, 210/5strongylense, 210/6surrectum, 210/2, Ann358Rhytidiadelphus, 248loreus, 248/1squarrosus, 248/2subpinnatus, 248/3triquetrus, 248/4Rhytidiastrumsquarrosum, 248/2

264 M. O. HILL ET AL.subpinnatum, 248/3Rhytidium, 249rugosum, 249/1Rosulabryumcapillare, 142/12elegans, 142/23laevifilum, 142/37torquescens, 142/53Saelania, 50glaucescens, 50/1Sanionia, 184georgicouncinata, 184/1nivalis, 184/1orthothecioides, 184/2uncinata, 184/3Schistidium, 34agassizii, 34/1, Ann77alpicola, 34/25, Ann77andreaeopsis, Ann75apocarpum, 34/2, Ann76atrofuscum, 34/3boreale, 34/4brunnescens, 34/5bryhnii, 34/6confertum, 34/7confusum, 34/8crassipilum, 34/9crenatum, 34/10dupretii, 34/11elegantulum, 34/12flaccidum, 34/13flexipile, 34/14frigidum, 34/15frisvollianum, 34/16grande, 34/17grandirete, 34/18helveticum, 34/19, Ann78holmenianum, 34/20lancifolium, 34/21marginale, Ann75maritimum, 34/22occidentale, 34/23, Ann80papillosum, 34/24platyphyllum, 34/25, Ann77poeltii, 34/26pratense, Ann75pruinosum, 34/27pulchrum, 34/28recurvum, 34/29rivulare, 34/30robustum, 34/31, Ann66scabrum, Ann75scandicum, 34/32sinensiapocarpum, 34/33, Ann81singarense, 34/19, Ann78sordidum, 34/34spinosum, 34/35, Ann82strictum, 34/36subflaccidium, Ann75subjulaceum, 34/37submuticum, 34/38tenerum, 34/39trichodon, 34/40umbrosum, 34/41venetum, 34/42Schistostega, 66pennata, 66/1Schizymenium, 148pontevedrense, 148/1Sciuro-hypnum, 219flotowianum, 219/1glaciale, 219/2latifolium, 219/3oedipodium, 219/4ornellanum, 219/5plumosum, 219/6populeum, 219/7reflexum, 219/8starkei, 219/9Scleromniumknyi, 272/1Scleropodium, 221apiculigerum, 219/5cespitans, 221/1, Ann382ornellanum, 219/5purum, 204/1touretii, 221/2, Ann32Scopelophila, 78cataractae, 78/1ligulata, 78/2Scorpidium, 189cossonii, 189/1revolvens, 189/2scorpioides, 189/3turgescens, 183/5Scorpiurium, 205circinatum, 205/1deflexifolium, 205/2sendtneri, 205/3Seligeria, 39acutifolia, 39/1alpestris, 39/17austriaca, 39/13brevifolia, 39/2calcarea, 39/8calycina, 39/9campylopoda, 39/5carniolica, 39/14diversifolia, 39/6donniana, 39/10galinae, 39/3, Ann86irrigata, 39/15, Ann87oelandica, 39/16patula, 39/17paucifolia, 39/9polaris, 39/11pusilla, 39/4recurvata, 39/7subimmersa, 39/12transylvanica, 38/1, Ann84trifaria, 39/18tristichoides, 39/19Sematophyllum, 263adnatum, 263/1, Ann412bottinii, 260/1demissum, 263/2micans, 262/1substrumulosum, 263/3Serpoleskeaconfervoides, 171/1subtilis, 171/4Sphagnum, 1affine, 1/1andersonianum, 1/44, Ann14angermanicum, 1/35angustifolium, 1/9annulatum, 1/10aongstroemii, 1/34, Ann8arcticum, 1/36auriculatum, 1/24, Ann5austinii, 1/2balticum, 1/11brevifolium, 1/13, Ann3capillifolium, 1/37, Ann10, Ann11centrale, 1/3compactum, 1/7contortum, 1/25cuspidatum, 1/12, Ann2denticulatum, 1/24, Ann5fallax, 1/13, Ann3fimbriatum, 1/38flexuosum, 1/14fuscum, 1/39girgensohnii, 1/40imbricatum, Ann1inundatum, 1/26, Ann5isoviitae, 1/13, Ann3jensenii, 1/15lenense, 1/16, Ann4lindbergii, 1/17magellanicum, 1/4majus, 1/18molle, 1/41nemoreum, 1/37nitidulum, Ann9obtusum, 1/19olafii, 1/42, Ann12orientale, Ann1palustre, 1/5papillosum, 1/6perfoliatum, Ann1

MOSSES OF EUROPE AND MACARONESIA 265platyphyllum, 1/27pulchrum, 1/20pylaesii, 1/28quinquefarium, 1/43recurvum, Ann3riparium, 1/21rubellum, 1/44, Ann10, Ann14rubiginosum, 1/45, Ann13russowii, 1/46skyense, 1/47squarrosum, 1/30steerei, Ann1strictum, 1/8subfulvum, 1/48subnitens, 1/49subsecundum, 1/29subtile, Ann10tenellum, 1/22tenerum, 1/37, Ann11teres, 1/31troendelagicum, 1/23tundrae, 1/32, Ann6viride, 1/12, Ann2warnstorfii, 1/50wulfianum, 1/33, Ann7Splachnobryum, 93delicatulum, 93/1, Ann150obtusum, 93/1, Ann150Splachnum, 120ampullaceum, 120/1luteum, 120/2melanocaulon, 120/3pensylvanicum, 120/4rubrum, 120/5sphaericum, 120/6vasculosum, 120/7Sporlederapalustris, 44/1Stegonia, 115latifolia, 115/1Stereodonbambergeri, 235/2callichrous, 235/3fertilis, 235/5hamulosus, 235/6holmenii, 235/7pallescens, 235/10plicatulus, 235/11revolutus, 235/14vaucheri, 235/18Straminergon, 190stramineum, 190/1Streblotrichumconvolutum, 99/3Streptocoleaatrata, 30/5Syntrichia, 116abranchesii, 116/3amplexa, 117/1bogotensis, 116/1, Ann203bolanderi, 117/5calcicola, 116/2, Ann204caninervis, 116/3, Ann205, Ann206, Ann207densa, 116/2echinata, 116/4fragilis, 116/5glabra, 116/6, Ann208handelii, 116/7inermis, 117/13intermedia, 116/11laevipila, 116/8, Ann209latifolia, 116/9minor, 116/10, Ann210montana, 116/11mutica, 116/5norvegica, 116/12pagorum, 116/8, Ann209papillosa, 116/13papillosissima, 116/14princeps, 116/15, Ann213pseudohandelii, Ann205rigescens, 116/16ruraliformis, 116/17ruralis, 116/17, Ann204, Ann214sinensis, 116/18subpapillosissima, 116/19, Ann215virescens, 116/20Taxiphyllum, 238densifolium, 238/1wissgrillii, 238/2Tayloria, 121acuminata, 121/1froelichiana, 121/2hornschuchii, 121/3lingulata, 121/4rudolphiana, 121/5serrata, 121/6splachnoides, 121/7tenuis, 121/8Tetraphis, 11pellucida, 11/1Tetraplodon, 122angustatus, 122/1blyttii, 122/2mnioides, 122/3, Ann228pallidus, 122/4paradoxus, 122/5urceolatus, 122/6, Ann228Tetrastichium, 166fontanum, 166/1virens, 166/2Tetrodontium, 12brownianum, 12/1ovatum, 12/2repandum, 12/3Thamniumcanariense, 275/1cossyrense, Ann353Thamnobryum, 271alopecurum, 271/1, Ann421angustifolium, 271/2canariense, 275/1cataractarum, 271/3, Ann419cossyrense, 205/3fernandesii, 271/4, Ann420maderense, 271/5, Ann421mediterraneum, 271/1neckeroides, 271/6, Ann422rudolphianum, 271/7, Ann423Thuidiopsis, 202sparsa, 202/1, Ann350Thuidium, 203abietinum, 199/1angustifolium, 192/1assimile, 203/1, Ann351atlanticum, 201/1, Ann349blandowii, 200/1delicatulum, 203/2, Ann351erectum, 203/2minutulum, 201/2philibertii, 203/1, Ann351recognitum, 203/3tamariscinum, 203/4virginianum, 192/3Timmia, 15austriaca, 15/1bavarica, 15/2comata, 15/4megapolitana, 15/3norvegica, 15/5sibirica, 15/6Timmiella, 77anomala, 77/1barbuloides, 77/2flexiseta, 77/3Tomentypnum, 185nitens, 185/1Tortella, 94alpicola, 94/1, Ann152arctica, Ann151bambergeri, 94/2, Ann153cirrifolia, 94/10, Ann157densa, 94/6, Ann154flavovirens, 94/3fragilis, 94/4glareicola, 94/3humilis, 94/5, Ann211inclinata, 94/6, Ann154inflexa, 94/7limbata, 94/8, Ann155limosella, 94/9, Ann156

266 M. O. HILL ET AL.nitida, 94/10, Ann157rigens, 94/11tortuosa, 94/12Tortula, 117abranchesii, 116/3acaulon, 111/1amplexa, 117/1ampliretis, 117/2, Ann218angustata, 117/28, Ann226atherodes, 111/1atrovirens, 117/3baetica, 117/14bogosica, 117/4bolanderi, 117/5, Ann219brevissima, 117/6, Ann221buyssonii, Ann216calcicolens, 116/2canescens, 117/7caninervis, 116/3caucasica, Ann217cernua, 117/8crinita, 116/11cuneifolia, 117/9densa, 116/2echinata, 116/4eucalyptrata, 117/12euryphylla, 117/12fragilis, 116/5freibergii, 117/10grandiretis, Ann217guepinii, 117/11handelii, 116/7hirsuta, 116/14hoppeana, 117/12, Ann222inermis, 117/13, Ann219intermedia, 116/11, Ann211israelis, 117/14, Ann220laevipila, 116/8laevipilaeformis, 116/8, Ann209lanceola, 117/15latifolia, 116/9laureri, 117/16leucostoma, 117/17lingulata, 117/18marginata, 117/19, Ann227modica, 117/20mucronifolia, 117/21muralis, 117/22, Ann220norvegica, 116/12obtusifolia, 117/23pagorum, 116/8pallida, 117/24papillosa, 116/13, Ann212papillosissima, 116/14princeps, 116/15protobryoides, 112/1randii, 117/25revolvens, 117/26rhizophylla, 108/1rhodonia, 117/27, Ann225rigescens, 116/16ruraliformis, 116/17ruralis, 116/17, Ann215saccardoana, 116/8, Ann209schimperi, 117/28, Ann226sinensis, 116/18solmsii, 117/29, Ann227spuria, 116/3subulata, 117/30, Ann226systylia, 117/31truncata, 117/32ucrainica, 117/33vahliana, 117/34virescens, 116/20viridifolia, 117/35wilsonii, 117/36zoddae, 117/24, Ann224Trachycystis, 150immarginata, 150/1ussuriensis, 150/1, Ann314Trematodon, 53ambiguus, 53/1brevicollis, 53/2, Ann32laetevirens, 53/3longicollis, 53/4, Ann32, Ann117perssoniorum, 53/5, Ann117Trichodon, 51cylindricus, 51/1Trichostomopsisaaronis, 104/3australasiae, 104/3trivialis, 104/26umbrosa, 104/26Trichostomum, 95, Ann158arcticum, 95/1, Ann151brachydontium, 95/2, Ann158caespitosum, 92/1connivens, Ann158contortum, Ann158crispulum, 95/3, Ann158hibernicum, 90/1pallidisetum, 95/4recurvifolium, 110/1tenuirostre, 90/2triumphans, 95/4, Ann158Triquetrella, 118arapilensis, 118/1Trochobryumcarniolicum, 39/14Ulota, 129bruchii, 129/1calvescens, 129/2coarctata, 129/3crispa, 129/4curvifolia, 129/5drummondii, 129/6hutchinsiae, 129/7intermedia, 129/4macrospora, 129/8, Ann249phyllantha, 129/9rehmannii, 129/10, Ann249Vesicularia, 239galerulata, Ann395reimersiana, 239/1, Ann395sphaerocarpa, Ann395Voitia, 123hyperborea, 123/1nivalis, 123/2Warnstorfia, 191exannulata, 191/1fluitans, 191/2h-schulzei, 191/2procera, 191/3, Ann339pseudostraminea, 191/4sarmentosa, 191/5trichophylla, 191/6tundrae, 191/7Weberaluisieri, Ann304maderensis, Ann304Weissia, 96brachycarpa, 96/1condensa, 96/2controversa, 96/3fallax, 96/3leptocarpa, Ann160levieri, 96/4longifolia, 96/5mittenii, Ann161multicapsularis, 96/6papillosissima, 96/2perssonii, 96/7rostellata, 96/8rutilans, 96/9squarrosa, 96/10sterilis, 96/11triumphans, 95/4tyrrhena, 96/12wimmeriana, 96/13, Ann162Zygodon, 130baumgartneri, 130/5conoideus, 130/1dentatus, 130/2forsteri, 130/3, Ann250gracilis, 130/4madeirensis, 130/3, Ann250rupestris, 130/5sibiricus, 130/6, Ann251stirtonii, 130/7, Ann252viridissimus, 130/8vulgaris, 130/5

MOSSES OF EUROPE AND MACARONESIA 267MARK O. HILL, Centre for Ecology and Hydrology, Monks Wood, Abbots Ripton, Huntingdon PE28 2LS, U.K. E-mail: moh@ceh.ac.ukNEIL BELL, Botanical Museum, Finnish Museum of Natural History, PO Box 7, FIN-00014 University of Helsinki, Finland.E-mail: neil.bell@helsinki.fiIDA BRUGGEMAN-NANNENGA, Griffensteijnseplein 23, 3703 BE Zeist, The Netherlands.MONTSERRAT BRUGUÉS, Departament de Biologia Animal, Vegetal i Ecologia, Unitat de Botànica, Universitat Autònoma de Barcelona,08193 Bellaterra, Barcelona, Spain. E-mail: Montserrat.Brugues@uab.esMARíA J. CANO, Departamento de Biología Vegetal (Botánica), Facultad de Biología, Universidad de Murcia, Campus de Espinardo, 30100Murcia, Spain. E-mail: mcano@um.esJOHANNES ENROTH, Department of Biological and Environmental Sciences and Botanical Museum, PO Box 7, FI-00014 University ofHelsinki, Finland. E-mail: enroth@touko.helsinki.fiKJELL I. FLATBERG, Section of Natural History, Museum of Natural History and Archaeology, Norwegian University of Science andTechnology, N-9471 Trondheim, Norway. E-mail kjell.flatberg@vm.ntnu.noJAN-PETER FRAHM, Nees Institut für Biodiversität der Pflanzen, Meckenheimer Allee 170, D-53115 Bonn, Germany.E-mail: frahm@uni-bonn.deMARíA TERESA GALLEGO, Departamento de Biología Vegetal (Botánica), Facultad de Biología, Universidad de Murcia, Campus deEspinardo, 30100 Murcia, Spain.RICARDO GARILLETI, Departamento de Botánica, Facultad de Farmacia, Universidad de Valencia, Avda. Vicente Andrés Estellés s/n,E-46100 Burjasot, SpainJUAN GUERRA, Departamento de Biología Vegetal (Botánica), Facultad de Biología, Universidad de Murcia, Campus de Espinardo, 30100Murcia, Spain. E-mail: jguerra@um.esLARS HEDENÄS, Swedish Museum of Natural History, Department of Cryptogamic Botany, Box 50007, SE-104 05, Stockholm, Sweden.E-mail: lars.hedenas@nrm.seDAVID. T.HOLYOAK, 8 Edward Street, Tuckingmill, Camborne, Cornwall TR14 8PA, UK. E-mail: david@holyoak9187.fsnet.co.ukJAAKKO HYVÖNEN, Plant Biology, PO Box 65 and Botanical Museum, PO Box 7, FIN-00014 University of Helsinki, Finland.E-mail: jaakko.hyvonen@helsinki.fiMICHAEL S. IGNATOV, Main Botanical Garden, Russian Academy of Sciences, Botanicheskaya 4, 127276 Moscow, Russia.E-mail: misha_ignatov@list.ruFRANCISCO LARA, Departamento de Biología (Botánica), Facultad de Ciencias, Universidad Autónoma de Madrid, C/Darwin, 2, E-28049Madrid, Spain. E-mail: francisco.lara@uam.esVICENTE MAZIMPAKA, Departamento de Biología (Botánica), Facultad de Ciencias, Universidad Autónoma de Madrid, C/Darwin, 2,E-28049 Madrid, Spain. E-mail: vicente.mazimpaka@uam.esJESÚS MUÑOZ, Real Jardín Botánico (CSIC), Plaza de Murillo 2, E-28014 Madrid, Spain. E-mail: jmunoz@ma-rjb.csic.esLARS SÖDERSTRÖM, Department of Biology, Norwegian University of Science and Technology, N-7491 Trondheim, Norway.E-mail: lars.soderstrom@bio.ntnu.no

Bryological Monograph An annotated checklist of the mosses of ...

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