Bryological Monograph An annotated checklist of the mosses of ...

Journal of Bryology (2006) 28: 198–267 

Bryological Monograph 

An annotated checklist of the mosses of Europe and Macaronesia 

M. O. HILL 1 , N. BELL 2 , M. A. BRUGGEMAN-NANNENGA 3 , M. BRUGUÉS 4 , M. J. CANO 5 , 

J. ENROTH 2 , K. I. FLATBERG 6 , J.-P. FRAHM 7 , M. T. GALLEGO 5 , R. GARILLETI 8 , 

J. GUERRA 5 , L. HEDENÄS 9 , D. T. HOLYOAK 10 , J. HYVÖNEN 2 , M. S. IGNATOV 11 , 

F. LARA 12 , V. MAZIMPAKA 12 , J. MUÑOZ 13 and L. SÖDERSTRÖM 14 

1 Centre for Ecology and Hydrology, Huntingdon, U.K., 2 University of Helsinki, Finland, 3 Nationaal Herbarium 

Nederland, Utrecht University branch, The Netherlands, 4 Universitat Autònoma de Barcelona, Spain, 

5 Universidad de Murcia, Spain, 6 Museum of Natural History and Archaeology, Trondheim, Norway, 7 Nees 

Institute for Biodiversity of Plants, University of Bonn, Germany, 8 Universidad de Valencia, Spain, 9 Swedish 

Museum of Natural History, Stockholm, 10 Camborne, Cornwall, U.K., 11 Main Botanical Garden, Moscow, 

Russia, 12 Universidad Autónoma de Madrid, Spain, 13 Real Jardín Botánico, Madrid, Spain and 14 Norwegian 

University of Science and Technology, Trondheim, Norway 

SUMMARY 

The moss flora of Europe and Macaronesia comprises 278 genera, 1292 species, 46 subspecies and 118 

varieties. Of the total 1292 species, 53 are confined to Macaronesia and 21 are thought to be non-native. The 

checklist was derived from those for the various component countries and regions. It is based on results 

published up to the end of 2005. Subspecies and varieties are included; hybrids are omitted. The taxonomic 

hierarchy is based on one published by Goffinet & Buck in 2004. While it has been strongly influenced by 

results of modern molecular methods, there are still many remaining uncertainties, even at family level. 

Because of these uncertainties, taxonomic innovation has generally been avoided. There are four new 

combinations and one change of status. 

KEYWORDS: Bryophyta, Sphagnopsida, Andreaeopsida, Oedipodiopsida, Polytrichopsida, Tetraphidopsida, 

Bryopsida. 

INTRODUCTION 

During the last two centuries, authors have at intervals 

produced floras and checklists that summarize the current 

state of knowledge of mosses in Europe. In the past 60 

years, comprehensive lists of European mosses were 

produced by Podpěra (1954), Corley et al. (1981), with a 

substantial update by Corley & Crundwell (1991) and 

CONTENTS 

INTRODUCTION . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .198 

TAXONOMIC HIERARCHY . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .201 

LIST OF TAXA. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .202 

ANNOTATIONS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .227 

ACKNOWLEDGEMENTS . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .244 

REFERENCES. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .244 

INDEX . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . .252 

# British Bryological Society 2006 

DOI: 10.1179/174328206X119998 

Dierssen (2001). It might be thought therefore that the time 

is not yet ripe for another list. However, Dierssen gives 

much information on ecology and phytosociology, but does 

not attempt a critical taxonomic review. Thus about 15 

years have passed since the moss flora was fully revised. 

These years have been very active and many new species 

have been described. A remarkable development, which 

would hardly have been anticipated in 1991, is the use of

DNA sequencing to understand basic relationships. This 

has revolutionized the systematics of land plants and 

resulted in a new classification of mosses. 

The immediate stimulus for producing a new list of 

European mosses came from a meeting of the European 

Committee for Conservation of Bryophytes (ECCB) in 

Valencia, Spain, in September 2004. They intend to produce 

an updated Red List of European bryophytes. For this 

activity it is necessary to have an updated checklist. There 

was no immediate need for a new list of Hepaticae and 

Anthocerotae, which had been revised by Grolle & Long 

(2000). Mosses were the priority. 

A project to make a new checklist 

After some consultation, M.O.H. was persuaded to coordinate 

production of the new list. He proposed a project lasting 

for the year 2005. It was given the name EuroMoss2005 and 

had its own website. A Steering Group consisting of Michele 

Aleffi (Italy), Bill Buck (USA), Montserrat Brugués (Spain), 

Klaus Dierssen (Germany), Misha Ignatov (Russia), Lars 

Söderström (Norway) and Jirˇí Váňa (Czech Republic) 

guided the project. It played a crucial role from the start, 

notably in giving the project sufficient gravitas to attract 

authors who were experts in particular groups. 

The project then proceeded in three stages. The first stage 

was to construct a provisional list, including synonyms, for 

experts to work from. For this, the ECCB supplied M.O.H. 

with an initial checklist, which was cross-referenced to the 

world list of Crosby et al. (1999). National and regional lists 

produced since about 1995 were then searched for names and 

additional taxa. Many lists were available as electronic 

documents, either posted on the internet or sent by e-mail. 

Members of the Steering Group acted as scouts, seeking as 

complete a species list as possible. The output of the first stage 

was a provisional list of names, some of which had authorities, 

together with synonyms that had recently been used in Europe. 

At the next stage, experts (Table 1) were each sent a 

provisional list of names and synonyms for their group, and 

Table 1. Experts contributing text for particular families; for the 

authorship of the paper, contributors other than the coordinator are 

listed alphabetically, with no attempt made to distinguish the magnitude 

of their contribution. 

Family or families Expert or experts 

Sphagnaceae Flatberg 

Polytrichaceae Hyvönen, Bell 

Funariaceae Brugués 

Grimmiaceae (except Schistidium) Muñoz 

Fissidentaceae Bruggeman-Nannenga 

Dicranales (except Fissidentaceae) Frahm 

Pottiaceae Guerra, Cano, Gallego 

Orthotrichaceae Lara, Garilleti, Mazimpaka 

Bryaceae Holyoak 

Amblystegiaceae, Calliergonaceae Hedenäs 

Brachytheciaceae Ignatov 

Neckeraceae, Lembophyllaceae Enroth 

MOSSES OF EUROPE AND MACARONESIA 199 

were asked to convert the provisional list into one 

conforming to their own ideas. This process resulted in 

most of the major groups having been reviewed by one or 

more experts. The gaps were filled by the coordinator and 

Professor Söderström, with advice from other bryologists 

who are thanked in the acknowledgements. 

When the list was nearly complete, doubtful species were 

reviewed in detail by the Steering Group, to ensure that 

they received reasonably consistent treatment. At the final 

stage, the information was loaded into databases (a 

database of names, another of annotations and a third 

with the bibliography) and reformatted for publication. 

Information published in papers after 2005 was not 

accepted for this list, which presents the state of knowledge 

on 31 December 2005. 

The area of the checklist 

Europe is defined basically as that part of the land mass of 

Eurasia bounded in the east by the Ural mountains and 

Ural River, and in the south by the Caucasus mountains 

and the Mediterranean and Black seas (cf. Söderström, 

Urmi & Váňa, 2002). Defined thus, it includes the 

Caucasian parts of the Russian Federation but excludes 

Georgia and Azerbaijan. It includes Turkey-in-Europe, 

western Kazakhstan, and all the Greek islands. It excludes 

Anatolia and Cyprus. In the Arctic, Novaya Zemlaya and 

Franz Josef Land are included (excluded by Corley et al., 

1981), as well as Svalbard and Jan Mayen. The Atlantic 

islands of Iceland and of Macaronesia (Canaries, Madeira 

and Azores but not Cape Verde) are included. 

Nomenclature and taxonomy 

With some exceptions, the names listed here are those 

included in European checklists from about 1995, together 

with names from the world list of Crosby et al. (1999). 

Earlier names are ignored. This means, fortunately, that 

there are exceedingly few cases where a name is ambiguous, 

with different significance depending on authorities. 

The more important subspecies and varieties have been 

included, but there is no intention to include varieties that 

are not generally recognized. 

No attempt has been made to give a comprehensive 

catalogue of moss hybrids. Hybrid taxa are mentioned in 

the notes only if they have been treated in recent lists. 

At the time of writing, moss taxonomy is being strongly 

affected by the results of studies using molecular methods 

and detailed anatomical analysis. In the present list for 

Europe, we have adopted a standpoint that might be 

characterized as moderately conservative. Thus we have 

accepted new generic concepts in Amblystegiaceae (mainly 

due to Hedenäs) and in Brachytheciaceae (mainly due to 

Ignatov and Huttunen). However, in Bryaceae, Hypnaceae, 

Plagiotheciaceae, Pottiaceae and several related small

200 M. O. HILL ET AL. 

families, generic and familial concepts are in turmoil. Our 

response has been to make only a few changes from 

previous European concepts, anticipating that new circumscriptions 

will become clear in future. 

For similar reasons we have avoided splitting traditional 

genera, retaining Dicranoweisia, Dicranum, Grimmia and 

Racomitrium as in Corley et al. (1981). We are aware from 

molecular studies that our circumscriptions of Barbula, 

Hygrohypnum, Hypnum, Polytrichastrum and Trichostomum 

are unsatisfactory, but are not yet able to provide 

new ones. On the other hand, we are happy to accept genera 

and species that are phylogenetically nested within others, 

provided that there are clear morphological distinctions. In all 

of these matters, we have aimed for stability and convenience. 

Author citations 

Correct author citations are difficult to achieve, and we are 

well aware that our own are far from perfect. Given the 

timescale of the project, we have had to rely extensively on 

secondary sources, especially for those families that were not 

revised by experts. For author citations where expert opinion 

was not available, a hierarchy of reliability was applied. At the 

top of the hierarchy were the nomina conservanda set out in the 

International Code of Botanical Nomenclature (Greuter et al., 

2000). At the next stage in the hierarchy, we used the 

catalogue of Polish mosses (Ochyra, Z˙ arnowiec & Bednarek- 

Ochyra, 2003), which contains the most comprehensive 

modern nomenclatural treatment of European taxa. For 

species and higher taxa not found in Poland, we drew on 

many other sources, none of which had overall priority. The 

internet database of the Missouri Botanical Garden, 

W 3 MOST, was frequently consulted. We have not hesitated 

to make further changes where we had a definite opinion. The 

resulting treatment no doubt contains some inconsistencies, 

for example in the citation of Bruch & Schimper or Schimper 

alone in the Bryologia Europaea. 

Table 2. Species confined in Europe to Macaronesia. 

Author abbreviations are those recommended by the 

International Plant Names Index (2005), which are 

accurately based on those of Brummitt & Powell (1992). 

Where the IPNI omits diacritical signs, we have followed it 

exactly; thus Sjors, Martensson and Jaderh. appear in place 

of the more correct Sjörs, Ma˚rtensson and Jäderh. Eva 

Maier is abbreviated E.Maier (as in W 3 MOST); she is not 

the Elva Maria Maier listed by IPNI. Also Robert Scott, 

author of Dicranum scottianum, is cited here as R.Scott, not 

as the cumbersome Scott, Robert in W 3 MOST. 

The starting date for moss nomenclature (the Sphagnaceae 

excepted) is 1 January 1801, which is the notional publication 

date of Species Muscorum. According to article 46.5 of the 

International Code (Greuter et al., 2000), the citation of an 

author who published the name before the starting point of 

the group concerned may be indicated by the use of the word 

‘ex’. However, this is optional and is not followed here for 

names in Hedwig (1801). We have generally cited the pre- 

Hedwigian author in other cases. Thus the form Sw. ex anon. 

is used for several Swartz species that were validated in an 

anonymous review (Anon., 1801), and L. ex Dicks. for a 

Linnaean species that was validated in the index of Dickson 

(1801). All names of species described by Dickson in his earlier 

fascicles were validated in this index if not before (Karttunen, 

1988). Dickson’s fourth fascicle was published on 4 October 

1801 and antedates the fourth edition of Withering’s 

Systematic Arrangement of British Plants, which was published 

on 26 December 1801 (Greene, 1962). Therefore the 

citation of Tayloria tenuis is here given as (Dicks.) Schimp. 

rather than (With.) Schimp. as in Corley et al. (1981). Several 

other citations have been changed similarly. 

New combinations 

For convenience of citation, new combinations are made by 

individual authors, including A.J.E. Smith, who supplied 

the text for his three. Smith’s reasons for making these 

Aloina humilis Echinodium prolixum Microcampylopus laevigatus 

Alophosia azorica Echinodium renauldii Nobregaea latinervis 

Amphidium tortuosum Echinodium setigerum Orthotrichum handiense 

Andoa berthelotiana Echinodium spinosum Pelekium atlanticum 

Brachymenium notarisii Entosthodon krausei Philonotis uncinata 

Brachymenium philonotula Fissidens azoricus Platyhypnidium torrenticola 

Brachythecium percurrens Fissidens coacervatus Rhynchostegiella bourgaeana 

Breutelia azorica Fissidens luisierii Rhynchostegiella macilenta 

Bryoxiphium madeirense Fissidens microstictus Rhynchostegiella trichophylla 

Bryum apiculatum Fissidens nobreganus Syntrichia bogotensis 

Campylopus cygneus Fissidens serratus Thamnobryum fernandesii 

Campylopus flaccidus Fissidens sublimbatus Thamnobryum rudolphianum 

Campylopus incrassatus Fissidens sublineaefolius Thuidiopsis sparsa 

Crossidium davidai Grimmia curviseta Tortella limbata 

Cryptoleptodon longisetus Homalia webbiana Tortula ampliretis 

Daltonia stenophylla Leucobryum albidum Tortula bogosica 

Dicranella campylophylla Leucodon canariensis Trematodon perssoniorum 

Ditrichum punctulatum Leucodon treleasei

Table 3. Non-native mosses in Europe. 

Species Location 

combinations are given in the second edition of his flora 

(Smith, 2004). 

Heterocladium flaccidum (Schimp.) A.J.E.Sm., stat. et 

comb. nov. Basionym: Heterocladium heteropterum var. 

flaccidum Schimp. in Bruch, Schimp. & W.Gümbel, 

Bryologia Europaea 5: 154 (fasc. 49–51). 1852. 

Hypnum cupressiforme var. heseleri (Ando & Higuchi) 

M.O.Hill, stat. et comb. nov. Basionym: Hypnum heseleri 

Ando & Higuchi, Journal of the Hattori Botanical 

Laboratory 75: 98. f. 1–2. 1994. 

Pelekium atlanticum (Hedenäs) Hedenäs, comb. nov. 

Basionym: Thuidium atlanticum Hedenäs, Journal of 

Bryology 16: 387. f. 1–2. 1991. 

Pohlia flexuosa var. pseudomuyldermansii (Arts, 

Nordhorn-Richter & A.J.E.Sm.) A.J.E.Sm., comb. nov. 

Basionym: Pohlia muyldermansii var. pseudomuyldermansii 

Arts, Nordhorn-Richter & A.J.E.Sm. Journal of Bryology 

14: 642. f. 3. 1987. 

Weissia6mittenii (Bruch & Schimp.) Mitt. emend. 

A.J.E.Sm., stat. nov. Basionym: Weissia mittenii (Bruch & 

Schimp.) Mitt., Annals and Magazine of Natural History, 

ser. 2, 8: 317. 1851. 

Planta originis hybridae. Parens feminea Weissia multicapsularis. 

Parens mas forte W. rostellata. 

Composition of the moss flora 

The moss flora comprises 1292 species in 278 genera, 71 

families and 22 orders. In addition, we recognize 46 

subspecies and 118 varieties. Of the total 1292 species, 53 

are confined in Europe to the Atlantic islands of 

Macaronesia (Table 2). In addition, Bryoxiphium norvegicum, 

Didymodon brachyphyllus and Polytrichastrum 


Achrophyllum dentatum Garden, England 

Atrichum crispum Britain and Ireland, widespread (male only) 

Bryum apiculatum Tenerife 

Bryum valparaisense Canary Islands, Portugal 

Calomnion complanatum Garden, Ireland 

Calyptrochaeta apiculata Britain, rare 

Campylopus introflexus C. and W. Europe, widespread 

Hennediella macrophylla Britain, widespread and spreading 

Hennediella stanfordensis Britain and Ireland, widespread and spreading 

Hypopterygium tamarisci Portugal 

Leptophascum leptophyllum Widespread in S. and W. Europe, apparently spreading 

Leptotheca gaudichaudii Garden, Ireland 

Myuroclada maximowiczii Garden, southern Russia 

Orthodontium lineare C. and W. Europe, widespread 

Sematophyllum adnatum Lombardy and Piedmont, several localities 

Splachnobryum obtusum Hot springs, Hungary and Azores; elsewhere in greenhouses 

Syntrichia bogotensis Madeira 

Tortula amplexa Britain, confined to one small area 

Tortula bogosica Azores and Canaries 

Tortula bolanderi S. Europe (rare), Macaronesia 

Thuidiopsis sparsa Madeira, in a former park area 

sphaerothecium are confined to Iceland, and Bryum 

miniatum to the Faeroe islands. 

In the east and south-east, several species are known only 

from Russia, mainly from the Urals and the Caucasus. In 

Asia just outside Europe, Crumia latifolia, Tortula caucasica 

and Trichostomum connivens are known from the 

Caucasus. An additional eight species are listed by 

Kürschner & Erdag˘ (2005) from Anatolia. 

Twenty-onespeciesofmossarethoughttohavebeen 

introduced to Europe by human agency (Table 3), a 

remarkably small number. Mosses must have been 

introduced from the Americas, especially to Macaronesia, 

before moss recording began; but invasions before 

1900 would not have been noticed. No non-native species 

is known to have arrived before 1800, though Corley & 

Frahm (1982) have speculated that Campylopus pyriformis 

is a late 18th century introduction from the Southern 

Hemisphere. Indeed, the only alien known to have reached 

Europe before 1900 is the North American Atrichum 

crispum, which has spread slowly and is probably a single 

clone. There are several other species that may be 

introduced. For example Bryum gemmiferum, now widespread 

in central and west Europe, appears still to be 

spreading, and lacks pre-1940 records (Herman 

Stieperaere, pers. comm.). 

TAXONOMIC HIERARCHY 

The main source for the hierarchy is Goffinet & Buck 

(2004). A few genera have been moved between families to 

take account of suggestions from Ignatov and Hedenäs or 

where there is evidence from newer molecular phylogenies. 

The following classes and orders occur in Europe. Orders 

are included in the list of taxa, below.


Sphagnopsida (Engl.) Ochyra 

Sphagnales Limpr. 

Andreaeopsida (Limpr.) Rothm. 

Andreaeales Limpr. 

Oedipodiopsida (Schimp.) Goffinet & W.R.Buck 

Oedipodiales (Schimp.) Goffinet & W.R.Buck 

Polytrichopsida Ochyra, Z˙ arnowiec & Bednarek-Ochyra 

Polytrichales M.Fleisch. 

Tetraphidopsida (M.Fleisch.) Goffinet & W.R.Buck 

Tetraphidales M.Fleisch. 

Bryopsida (Limpr.) Rothm. 

Buxbaumiales M.Fleisch. 

Diphysciales M.Fleisch. 

Timmiales (M.Fleisch.) Ochyra 

Encalyptales Dixon 

Funariales M.Fleisch. 

Bryoxiphiales H.A.Crum & L.E.Anderson 

Grimmiales M.Fleisch. 

Archidiales Limpr. 

Dicranales H.Philib. ex M.Fleisch. 

Pottiales M.Fleisch. 

Splachnales (M.Fleisch.) Ochyra 

Orthotrichales Dixon 

Hedwigiales Ochyra 

Bryales Limpr. 

Rhizogoniales (M.Fleisch.) Goffinet & W.R.Buck 

Hookeriales M.Fleisch. 

Hypnales (M.Fleisch.) W.R.Buck & Vitt 

LIST OF TAXA 

Sphagnales Limpr. 

Sphagnaceae Dumort. 

1 Sphagnum L. [1] 

section Sphagnum 

1 affine Renauld & Cardot (S. affine var. flagellare 

(Schlieph. ex Röll) L.Söderstr. & Hedenäs, S. imbricatum 

subsp. affine (Renauld & Cardot) Flatberg) 

2 austinii Sull. (S. imbricatum subsp. austinii (Sull.) 

Flatberg) 

3 centrale C.E.O.Jensen (S. palustre var. centrale 

(C.E.O.Jensen) A.Eddy) 

4 magellanicum Brid. 

5 palustre L. 

6 papillosum Lindb. 

section Rigida (Lindb.) Limpr. 

7 compactum Lam. & DC. 

8 strictum Sull. 

section Cuspidata Lindb. 

9 angustifolium (C.E.O.Jensen ex Russow) 

C.E.O.Jensen (S. recurvum var. tenue H.Klinggr.) 

10 annulatum H.Lindb. ex Warnst. 

11 balticum (Russow) C.E.O.Jensen 

12 cuspidatum Ehrh. ex Hoffm. (S. viride Flatberg) [2] 

13 fallax (H.Klinggr.) H.Klinggr. (S. brevifolium (Lindb. 

ex Braithw.) Röll, S. fallax subsp. isoviitae (Flatberg) 

M.O.Hill, S. isoviitae Flatberg, S. recurvum var. 

brevifolium (Lindb. ex Braithw.) Warnst.) [3] 

14 flexuosum Dozy & Molk. 

15 jensenii H.Lindb. 

16 lenense H.Lindb. ex L.I.Savicz [4] 

17 lindbergii Schimp. 

18 majus (Russow) C.E.O.Jensen 

subsp. majus 

subsp. norvegicum Flatberg 

19 obtusum Warnst. 

20 pulchrum (Lindb. ex Braithw.) Warnst. 

21 riparium A˚ ngstr. 

22 tenellum (Brid.) Pers. ex Brid. 

23 troendelagicum Flatberg 

section Subsecunda (Lindb.) Schimp. 

24 auriculatum Schimp. (S. denticulatum Brid. [5], S. 

subsecundum var. rufescens (Nees ex Hornsch.) 

Huebener) 

25 contortum Schultz (S. subsecundum var. contortum 

(Schultz) Huebener) 

26 inundatum Russow (S. denticulatum var. inundatum 

(Russow) Kartt., S. subsecundum subsp. inundatum 

(Russow) Meyl., S. subsecundum var. inundatum 

(Russow) C.E.O.Jensen) 

27 platyphyllum (Lindb. ex Braithw.) Warnst. 

(S. subsecundum var. platyphyllum (Lindb. ex 

Braithw.) Cardot) 

28 pylaesii Brid. 

29 subsecundum Nees 

section Squarrosa (Russow) Schimp. 

30 squarrosum Crome 

31 teres (Schimp.) A˚ ngstr. 

32 tundrae Flatberg [6] 

section Polyclada (C.E.O.Jensen) Horrell 

33 wulfianum Girg. [7] 

section Insulosa Isov. 

34 aongstroemii C.Hartm. [8] 

section Acutifolia Wilson [9, 10] 

35 angermanicum Melin 

36 arcticum Flatberg & Frisvoll 

37 capillifolium (Ehrh.) Hedw. (S. nemoreum Scop., S. 

tenerum auct. eur. non Sull. & Lesq. ex Sull.) [11] 

38 fimbriatum Wilson 

subsp. concinnum (Berggr.) Flatberg & Frisvoll 

subsp. fimbriatum 

39 fuscum (Schimp.) H.Klinggr. 

40 girgensohnii Russow 

41 molle Sull. 

42 olafii Flatberg [12] 

43 quinquefarium (Braithw.) Warnst. 

44 rubellum Wilson (S. andersonianum R.E.Andrus [14], 

S. capillifolium subsp. rubellum (Wilson) M.O.Hill, S. 

capillifolium var. tenellum (Schimp.) H.A.Crum) 

45 rubiginosum Flatberg [13] 

46 russowii Warnst. 

47 skyense Flatberg 

48 subfulvum Sjors

subsp. purpureum Flatberg 

subsp. subfulvum 

49 subnitens Russow & Warnst. 

subsp. ferrugineum Flatberg (S. subnitens var. 

ferrugineum (Flatberg) M.O.Hill) 

subsp. subnitens 

50 warnstorfii Russow 

Andreaeales Limpr. 

Andreaeaceae Dumort. 

2 Andreaea Hedw. [15] 

section Chasmocalyx Lindb. ex Braithw. 

1 nivalis Hook. 

section Nerviae Cardot ex G.Roth 

2 blyttii Schimp. 

3 crassinervia Bruch 

4 frigida Huebener 

5 heinemannii Hampe & Müll.Hal. (A. planinervia 

Lindb. ex G.Roth) 

subsp. crassifolia (Luisier) Sérgio (A. crassifolia 

Luisier) [16] 

subsp. heinemannii 

6 megistospora B.M.Murray 

7 rothii F.Weber & D.Mohr 

subsp. falcata (Schimp.) Lindb. 

subsp. rothii 

section Andreaea 

8 alpestris (Thed.) Schimp. (A. rupestris var. alpestris 

(Thed.) Sharp) 

9 alpina Hedw. 

10 mutabilis Hook.f. & Wilson 

11 obovata Thed. (A. hartmanii Thed.) 

12 rupestris Hedw. 

var. papillosa (Lindb.) Podp. 

var. rupestris 

13 sinuosa B.M.Murray 

Oedipodiales (Schimp.) Goffinet & W.R.Buck 

Oedipodiaceae Schimp. 

3 Oedipodium Schwägr. 

1 griffithianum (Dicks.) Schwägr. 

Polytrichales M.Fleisch. 

Polytrichaceae Schwägr. 

4 Alophosia Cardot 

1 azorica (Renauld & Cardot) Cardot 

5 Atrichum P.Beauv., nom. cons. 

1 angustatum (Brid.) Bruch & Schimp. (A. angustatum 

var. rhystophyllum (Müll.Hal.) P.W.Richards & 

E.C.Wallace) [17] 

2 crispum (James) Sull. 

3 flavisetum Mitt. (A. haussknechtii Jur. & Milde, A. 

undulatum var. gracilisetum Besch.) [18] 

4 tenellum (Röhl.) Bruch & Schimp. 

5 undulatum (Hedw.) P.Beauv. 

6 Oligotrichum DC., nom. cons. 

1 hercynicum (Hedw.) Lam. & DC. 

7 Pogonatum P.Beauv. [19] 

1 aloides (Hedw.) P.Beauv. [20] 


2 dentatum (Menzies ex Brid.) Brid. (P. capillare 

(Michx.) Brid.) 

3 nanum (Hedw.) P.Beauv. [20] 

4 neesii (Müll.Hal.) Dozy [21] 

5 urnigerum (Hedw.) P.Beauv. (P. urnigerum var. 

subintegrifolium (Arnell & C.E.O.Jensen) H.Möller) 

8 Polytrichastrum G.L.Sm. [22] 

1 alpinum (Hedw.) G.L.Sm. (Pogonatum alpinum 

(Hedw.) Rohl., Polytrichastrum alpinum var. 

septentrionale (Sw. ex Brid.) G.L.Sm., P. alpinum var. 

fragile (Bryhn) D.G.Long, P. norwegicum (Hedw.) 

Schljakov, Polytrichum alpinum Hedw., P. alpinum 

var. arcticum (Sw. ex Brid.) Wahlenb.) [23] 

2 formosum (Hedw.) G.L.Sm. (Polytrichum 

formosum Hedw.) 

3 longisetum (Sw. ex Brid.) G.L.Sm. (P. longisetum var. 

anomalum (Milde) Ignatov & G.L.Merr., Polytrichum 

longisetum Sw. ex Brid.) 

4 pallidisetum (Funck) G.L.Sm. (Polytrichum formosum 

var. decipiens (Limpr.) Loeske, Polytrichum 

pallidisetum Funck) 

5 sexangulare (Brid.) G.L.Sm. (Polytrichum sexangulare 

Hedw.) 

6 sphaerothecium (Besch.) J.-P.Frahm (P. sexangulare 

var. vulcanicum (C.E.O.Jensen) G.L.Merr., 

Polytrichum sphaerothecium (Besch.) Müll.Hal.) [24] 

9 Polytrichum Hedw. [25] 

1 commune Hedw. (P. commune var. humile Sw., P. 

commune var. perigoniale (Michx.) Hampe, P. 

perigoniale Michx.) 

2 hyperboreum R.Br. 

3 jensenii I.Hagen (P. commune var. jensenii (I.Hagen) 

Mönk.) 

4 juniperinum Hedw. 

5 piliferum Hedw. (P. piliferum var. hoppei (Hornsch.) 

Haller) 

6 strictum Menzies ex Brid. (P. affine Funck, P. alpestre 

Hoppe) 

7 swartzii Hartm. (P. commune var. swartzii (Hartm.) 

Nyholm) 

8 uliginosum (Wallr.) Schriebl (P. commune var. 

uliginosum Wallr.) 

10 Psilopilum Brid. 

1 cavifolium (Wilson) I.Hagen 

2 laevigatum (Wahlenb.) Lindb. (P. laevigatum var. 

aloma I.Hagen) 

Tetraphidales M.Fleisch. 

Tetraphidaceae Schimp. 

11 Tetraphis Hedw. 

1 pellucida Hedw. 

12 Tetrodontium Schwägr. 

1 brownianum (Dicks.) Schwägr. 

2 ovatum (Funck) Schwägr. 

3 repandum (Funck) Schwägr. 

Buxbaumiales M.Fleisch. 

Buxbaumiaceae Schimp. 

13 Buxbaumia Hedw.


1 aphylla Hedw. 

2 viridis (Moug. ex Lam. & DC.) Brid. ex Moug. & Nestl. 

Diphysciales M.Fleisch. 

Diphysciaceae M.Fleisch. 

14 Diphyscium D.Mohr 

1 foliosum (Hedw.) D.Mohr 

Timmiales (M.Fleisch.) Ochyra 

Timmiaceae Schimp. 

15 Timmia Hedw., nom. cons. 

section Timmiaurea Brassard 

1 austriaca Hedw. 

section Timmia 

2 bavarica Hessl. 

3 megapolitana Hedw. 

section Norvegica Brassard 

4 comata Lindb. & Arnell 

5 norvegica J.E.Zetterst. 

6 sibirica Lindb. & Arnell 

Encalyptales Dixon 

Encalyptaceae Schimp. 

16 Bryobrittonia Williams [26] 

1 longipes (Mitt.) D.G.Horton (Encalypta longipes Mitt.) 

17 Encalypta Hedw. [27] [28] 

section Streptotheca (Kindb.) Broth. 

1 procera Bruch 

2 streptocarpa Hedw. 

section Pyromitrium Wallr. ex Hampe 

3 alpina Sm. 

4 mutica I.Hagen (E. vulgaris var. mutica Brid.) 

section Rhabdotheca Müll.Hal. 

5 intermedia Jur. [29] 

6 obovatifolia Nyholm [30] 

7 rhaptocarpa Schwägr. 

var. leptodon Lindb. (E. rhaptocarpa var. trachymitria 

(Ripart) Wijk & Margad., E. trachymitria Ripart) [31] 

var. rhaptocarpa 

8 spathulata Müll.Hal. (E. rhaptocarpa var. spathulata 

(Müll.Hal.) Husn.) 

9 vulgaris Hedw. 

section Megasporae D.G.Horton 

10 longicolla Bruch [32] 

section Encalypta 

11 affinis R.Hedw. 

subsp. affinis 

subsp. macounii (Austin) D.G.Horton (E. macounii 

Austin) [33] 

12 brevicolla (Bruch & Schimp.) A˚ ngstr. [32] 

13 brevipes Schljakov 

14 ciliata Hedw. 

15 microstoma Bals.-Criv. & De Not. 

Funariales M.Fleisch. 

Funariaceae Schwägr. [34] [35] 

18 Entosthodon Schwägr. [36] 

Subgenus Entosthodon 

1 attenuatus (Dicks.) Bryhn (E. templetonii (Sm.) 

Schwägr., Funaria attenuata (Dicks.) Lindb.) 

2 durieui Mont. (E. durieui var. mustaphae Trab., 

E. mustaphae Trab., E. pallescens Jur., E. pallescens 

var. mitratus (Casares-Gil) Wijk & Margad., 

E. physcomitroides Casares-Gil & Beltrán, Funaria 

pallescens (Jur.) Lindb.) [37] 

3 hungaricus (Boros) Loeske (E. maroccanus (Meyl.) 

Hébr. & Lo Giudice, Funaria hungarica Boros, 

Physcomitrium maroccanum Meyl.) [38] 

4 krausei Besch. (Funaria krausei (Besch.) Geh. & 

Herzog) [39] 

5 obtusus (Hedw.) Lindb. (E. ericetorum (De Not.) 

Müll.Hal., Funaria obtusa (Hedw.) Lindb.) 

Subgenus Plagiodus (Mitt.) Fife 

6 convexus (Spruce) Brugués (Funaria convexa Spruce, 

Funaria pustulosa Zodda) 

7 muhlenbergii (Turner) Fife (Funaria calcarea 

Wahlenb., Funaria dentata Crome, Funaria hibernica 

Hook., Funaria mediterranea Lindb., Funaria 

muhlenbergii Turner) 

8 pulchellus (H.Philib.) Brugués (Funaria pulchella 

H.Philib.) 

9 schimperi Brugués (Funaria algieriensis Lindb., 

Funaria durieui Schimp.) [40] 

Subgenus Murcia Fife 

10 fascicularis (Hedw.) Müll.Hal. (Funaria fascicularis 

(Hedw.) Lindb.) 

11 mouretii (Corb.) Jelenc (Funaria mouretii Corb.) [41] 

19 Funaria Hedw. [36] [39] [42] 

1 aequidens Lindb. ex Broth. 

2 arctica (Berggr.) Kindb. 

3 hygrometrica Hedw. 

4 microstoma Bruch ex Schimp. 

20 Funariella Sérgio 

1 curviseta (Schwägr.) Sérgio (Entosthodon curvisetus 

(Schwägr.) Müll.Hal., Funaria curviseta (Schwägr.) 

Milde) 

21 Goniomitrium Hook.f. & Wilson [43] 

1 seroi Casas [44] 

22 Physcomitrella Bruch & Schimp. 

1 patens (Hedw.) Bruch & Schimp. (Aphanorrhegma 

patens (Hedw.) Lindb., Physcomitrium patens (Hedw.) 

Mitt.) 

23 Physcomitrium (Brid.) Brid. 

1 arenicola Laz. 

2 eurystomum Sendtn. 

subsp. acuminatum (Bruch & Schimp.) Giacom. 

(P. acuminatum Bruch & Schimp.) [45] 

subsp. eurystomum 

3 pyriforme (Hedw.) Bruch & Schimp. 

4 sphaericum (C.F.Ludw. ex Schkuhr) Brid. 

24 Pyramidula Brid. 

1 tetragona (Brid.) Brid. (P. algeriensis Chud. & Douin, 

P. tetragona var. zoddae Bott.) [46] 

Disceliaceae Schimp. 

25 Discelium Brid. 

1 nudum (Dicks.) Brid. 

Gigaspermaceae Lindb.

26 Gigaspermum Lindb. 

1 mouretii Corb. 

27 Oedipodiella Dixon 

1 australis (Wager & Dixon) Dixon (O. australis var. 

catalaunica P.de la Varde) [47] 

Bryoxiphiales H.A.Crum & L.E.Anderson 

Bryoxiphiaceae Besch., nom. cons. 

28 Bryoxiphium Mitt., nom. cons. 

1 madeirense Á.Löve & D.Löve [48] 

2 norvegicum (Brid.) Mitt. 

Grimmiales M.Fleisch. 

Grimmiaceae Arn. 

29 Coscinodon Spreng. 

1 cribrosus (Hedw.) Spruce (C. cribrosus var. brevipila 

M.Fleisch. & Warnst.) 

2 humilis Milde [49] 

30 Grimmia Hedw. 

1 alpestris (F.Weber & D.Mohr) Schleich. 

(Orthogrimmia alpestris (F.Weber & D.Mohr) Ochyra 

&Z˙ arnowiec) [50] 

2 anodon Bruch & Schimp. (G. limprichtii Kern) [51] 

3 anomala Hampe ex Schimp. (Dryptodon anomalus 

(Hampe ex Schimp.) Loeske) 

4 arenaria Hampe (G. donniana var. curvula Spruce, 

Orthogrimmia arenaria (Hampe) Ochyra & 

Z˙ arnowiec) [52] 

5 atrata Miel. ex Hornsch. (Streptocolea atrata (Miel. ex 

Hornsch.) Ochyra & Z˙ arnowiec) 

6 caespiticia (Brid.) Jur. (G. pyrenaica Kern, 

Orthogrimmia caespiticia (Brid.) Ochyra & Z˙ arnowiec) 

7 capillata De Not. [53] 

8 crinita Brid. 

9 curviseta Bouman [54] 

10 decipiens (Schultz) Lindb. (Dryptodon decipiens 

(Schultz) Loeske) 

11 dissimulata E.Maier [55] 

12 donniana Sm. (G. sudetica Spreng. ex Schkuhr, 

Orthogrimmia donniana (Sm.) Ochyra & Z˙ arnowiec) 

13 elatior Bruch ex Bals.-Criv. & De Not. (Dryptodon 

incurvus (Hornsch.) Brid. [56], G. elatior var. 

pseudofunalis Limpr.) 

14 elongata Kaulf. (Dryptodon elongatus (Kaulf.) Hartm.) 

15 funalis (Schwägr.) Bruch & Schimp. (Dryptodon 

funalis (Schwägr.) Brid., G. funalis var. calvescens 

(Kindb.) H.Möller) [57] 

16 fuscolutea Hook. (G. apiculata Hornsch., G. holleri 

Molendo) 

17 hartmanii Schimp. (Dryptodon hartmanii (Schimp.) 

Limpr.) 

18 incurva Schwägr. (Dryptodon contortus (Wahlenb.) 

Brid.) 

19 laevigata (Brid.) Brid. (Guembelia laevigata (Brid.) 

Ochyra & Z˙ arnowiec) 

20 lisae De Not. (G. retracta Stirt.) [58] 

21 longirostris Hook. (G. affinis Hornsch., Guembelia 

longirostris (Hook.) Ochyra & Z˙ arnowiec) 


22 mollis Bruch & Schimp. (Hydrogrimmia mollis (Bruch 

& Schimp.) Loeske) 

23 montana Bruch & Schimp. (G. pilosissima Herzog, 

Orthogrimmia montana (Bruch & Schimp.) Ochyra 

&Z˙ arnowiec) 

24 muehlenbeckii Schimp. (Dryptodon muehlenbeckii 

(Schimp.) Loeske) [59] 

25 nutans Bruch (G. meteorae C.C.Towns.) 

26 orbicularis Bruch ex Wilson (Dryptodon orbicularis 

(Bruch ex Wilson) Ochyra & Z˙ arnowiec) 

27 ovalis (Hedw.) Lindb. (Guembelia ovalis (Hedw.) 

Müll.Hal.) 

28 plagiopodia Hedw. (G. plagiopodia var. arvernica 

(H.Philib.) Boulay) 

29 poecilostoma Cardot & Sebille, nom. cons. prop. 

(G. tergestina var. poecilostoma (Cardot & Sebille) 

Loeske, G. tergestina var. tergestinoides Culm.) [59] 

30 pulvinata (Hedw.) Sm. (Dryptodon pulvinatus (Hedw.) 

Brid., G. pulvinata var. africana (Hedw.) Hook.f., 

G. pulvinata var. obtusa (Brid.) Huebener) [60] 

31 ramondii (Lam. & DC.) Margad. (Dryptodon patens 

(Hedw.) Brid., G. curvata (Brid.) De Sloover) 

32 reflexidens Müll.Hal. (G. grisea Cardot [62], 

G. sessitana De Not., G. subsulcata Limpr. [63], 

Orthogrimmia sessitana (De Not.) Ochyra 

&Z˙ arnowiec) [61] 

33 teretinervis Limpr. 

34 tergestina Tomm. ex Bruch & Schimp. (G. crassifolia 

Lindb. ex Broth. [64], Guembelia tergestina (Tomm. ex 

Bruch & Schimp.) Buyss.) 

35 torquata Drumm. (Dryptodon torquatus (Drumm.) 

Brid.) 

36 trichophylla Grev. (Dryptodon trichophyllus (Grev.) 

Brid., G. austrofunalis auct. eur. non Müll.Hal. [65], 

G. britannica A.J.E.Sm. [66], G. meridionalis (Müll.Hal.) 

E.Maier [67], G. trichophylla var. stirtonii (Schimp.) 

H.Möller, G. trichophylla var. subsquarrosa (Wilson) 

A.J.E.Sm., G. trichophylla var. tenuis (Wahlenb.) Wijk 

& Margad.) 

37 triformis Carestia & De Not. (Orthogrimmia triformis 

(Carestia & De Not.) Ochyra & Z˙ arnowiec) [68] 

38 ungeri Jur. [69] 

39 unicolor Hook. 

31 Indusiella Broth. & Müll.Hal. 

1 thianschanica Broth. & Müll.Hal. [70] 

32 Jaffueliobryum Thér. 

1 latifolium Thér. [71] 

33 Racomitrium Brid. [72] 

Subgenus Racomitrium 

1 lanuginosum (Hedw.) Brid. 

Subgenus Niphotrichum Bednarek-Ochyra 

(Niphotrichum (Bednarek-Ochyra) Bednarek-Ochyra 

& Ochyra) 

2 canescens (Hedw.) Brid. (Niphotrichum canescens 

(Hedw.) Bednarek-Ochyra & Ochyra) 

subsp. canescens


subsp. latifolium (C.E.O.Jensen) Frisvoll 

(Niphotrichum canescens subsp. latifolium 

(C.E.O.Jensen) Bednarek-Ochyra & Ochyra) 

3 elongatum Ehrh. ex Frisvoll (Niphotrichum elongatum 

(Frisvoll) Bednarek-Ochyra & Ochyra, R. canescens 

var. intermedium Venturi & Bott.) 

4 ericoides (Brid.) Brid. (Niphotrichum ericoides (Brid.) 

Bednarek-Ochyra & Ochyra, R. canescens var. 

ericoides (Brid.) Hampe) 

5 panschii (Müll.Hal.) Kindb. (Niphotrichum panschii 

(Müll.Hal.) Bednarek-Ochyra & Ochyra) 

Subgenus Cataracta Vilh. (Codriophorus P.Beauv.) 

6 aciculare (Hedw.) Brid. (Codriophorus acicularis 

(Hedw.) P.Beauv.) 

7 aquaticum (Brid. ex Schrad.) Brid. (Codriophorus 

aquaticus (Brid. ex Schrad.) Bednarek-Ochyra 

& Ochyra) 

8 fasciculare (Hedw.) Brid. (Codriophorus fascicularis 


9 hespericum Sérgio, J.Muñoz & Ochyra (Codriophorus 

hespericus (Sérgio, J.Muñoz & Ochyra) Bednarek- 

Ochyra & Ochyra) [73] 

Subgenus Ellipticodryptodon (Vilh.) Bednarek-Ochyra 

& Ochyra (Bucklandiella Roiv.) 

10 affine (F.Weber & D.Mohr) Lindb. (Bucklandiella 

affinis (F.Weber & D.Mohr) Bednarek-Ochyra 

& Ochyra, R. heterostichum var. affine (F.Weber 

& D.Mohr) Lesq.) 

11 ellipticum (Turner) Bruch & Schimp. (Bucklandiella 

elliptica (Turner) Bednarek-Ochyra & Ochyra) 

12 heterostichum (Hedw.) Brid. (Bucklandiella 

heterosticha (Hedw.) Bednarek-Ochyra & Ochyra) 

13 himalayanum (Mitt.) A.Jaeger (Bucklandiella 

himalayana (Mitt.) Bednarek-Ochyra & Ochyra) 

14 lamprocarpum (Müll.Hal.) A.Jaeger (Bucklandiella 

lamprocarpa (Müll.Hal.) Bednarek-Ochyra & Ochyra) 

15 lusitanicum Ochyra & Sérgio (Bucklandiella lusitanica 

(Ochyra & Sérgio) Bednarek-Ochyra & Ochyra) [74] 

16 macounii Kindb. (Bucklandiella macounii (Kindb.) 

Bednarek-Ochyra & Ochyra) 

subsp. alpinum (E.Lawton) Frisvoll (Bucklandiella 

macounii subsp. alpina (E.Lawton) Bednarek-Ochyra 

& Ochyra) 

subsp. macounii 

17 microcarpon (Hedw.) Brid. (Bucklandiella microcarpa 


18 obtusum (Brid.) Brid. (Bucklandiella obtusa (Brid.) 

Bednarek-Ochyra & Ochyra) 

19 sudeticum (Funck) Bruch & Schimp. (Bucklandiella 

sudetica (Funck) Bednarek-Ochyra & Ochyra, 

R. heterostichum var. sudeticum (Funck) E.Bauer) 

34 Schistidium Bruch & Schimp., nom. cons. [75] [76] 

1 agassizii Sull. & Lesq. [77] 

2 apocarpum (Hedw.) Bruch & Schimp. 

3 atrofuscum (Schimp.) Limpr. (S. apocarpum var. 

atrofuscum (Schimp.) C.E.O.Jensen) 

4 boreale Poelt 

5 brunnescens Limpr. 

subsp. brunnescens (S. apocarpum var. brunnescens 

(Limpr.) Loeske) 

subsp. griseum (Nees & Hornsch.) H.H.Blom 

6 bryhnii I.Hagen 

7 confertum (Funck) Bruch & Schimp. (S. apocarpum 

var. confertum (Funck) H.Möller) 

8 confusum H.H.Blom 

9 crassipilum H.H.Blom 

10 crenatum H.H.Blom 

11 dupretii (Thér.) W.A.Weber 

12 elegantulum H.H.Blom 

subsp. elegantulum 

subsp. wilsonii H.H.Blom 

13 flaccidum (De Not.) Ochyra (S. pulvinatum var. 

flaccidum (De Not.) De Not.) 

14 flexipile (Lindb. ex Broth.) G.Roth 

15 frigidum H.H.Blom 

var. frigidum 

var. havaasii H.H.Blom 

16 frisvollianum H.H.Blom 

17 grande Poelt 

18 grandirete H.H.Blom 

19 helveticum (Schkuhr) Deguchi (S. singarense (Schiffn.) 

Laz.) [78] 

20 holmenianum Steere & Brassard 

21 lancifolium (Kindb.) H.H.Blom 

22 maritimum (Sm. ex R.Scott) Bruch & Schimp. [79] 

subsp. maritimum 

subsp. piliferum (I.Hagen) B.Bremer (S. maritimum 

var. piliferum (I.Hagen) Loeske) 

23 occidentale (E.Lawton) S.P.Churchill [80] 

24 papillosum Culm. (S. apocarpum subsp. papillosum 

(Culm.) Poelt) 

25 platyphyllum (Mitt.) H.Perss. (S. alpicola auct. non 

(Hedw.) Limpr., S. rivulare subsp. latifolium 

(J.E.Zetterst.) B.Bremer, S. rivulare var. latifolium 

(J.E.Zetterst.) H.A.Crum & L.E.Anderson) [77] 

subsp. abrupticostatum (Bryhn) H.H.Blom 

subsp. platyphyllum 

26 poeltii H.H.Blom 

27 pruinosum (Wilson ex Schimp.) G.Roth 

28 pulchrum H.H.Blom 

29 recurvum H.H.Blom 

30 rivulare (Brid.) Podp. 

31 robustum (Nees & Hornsch.) H.H.Blom 

32 scandicum H.H.Blom 

33 sinensiapocarpum (Müll.Hal.) Ochyra [81] 

34 sordidum I.Hagen 

35 spinosum H.H.Blom & Lüth [82] 

36 strictum (Turner) Loeske ex Martensson 

37 subjulaceum H.H.Blom 

38 submuticum H.H.Blom 

subsp. arcticum H.H.Blom 

subsp. submuticum 

39 tenerum (J.E.Zetterst.) Nyholm 

40 trichodon (Brid.) Poelt

var. nutans H.H.Blom 

var. trichodon 

41 umbrosum (J.E.Zetterst.) H.H.Blom 

42 venetum H.H.Blom 

Ptychomitriaceae Schimp. 

35 Campylostelium Bruch & Schimp. 

1 pitardii (Corb.) E.Maier (Grimmia pitardii Corb.) [83] 

2 saxicola (F.Weber & D.Mohr) Bruch & Schimp. 

3 strictum Solms 

36 Ptychomitrium Fürnr., nom. cons. 

1 incurvum (Schwägr.) Spruce 

2 nigrescens (Kunze) Wijk & Margad. 

3 polyphyllum (Dicks. ex Sw.) Bruch & Schimp. 

Seligeriaceae Schimp. 

37 Blindia Bruch & Schimp. 

1 acuta (Hedw.) Bruch & Schimp. 

2 caespiticia (F.Weber & D.Mohr) Müll.Hal. 

38 Brachydontium Fürnr. 

1 trichodes (F.Weber) Milde (Seligeria transylvanica 

Plam. [84]) 

39 Seligeria Bruch & Schimp. [85] 

Subgenus Seligeria 

1 acutifolia Lindb. 

2 brevifolia (Lindb.) Lindb. 

3 galinae Mogensen & I.Goldberg [86] 

4 pusilla (Hedw.) Bruch & Schimp. 

Subgenus Cyrtoseligeria Vitt 

5 campylopoda Kindb. 

6 diversifolia Lindb. 

7 recurvata (Hedw.) Bruch & Schimp. 

Subgenus Anodus (Bruch & Schimp.) Boulay 

8 calcarea (Hedw.) Bruch & Schimp. 

9 calycina Mitt. ex Lindb. (S. paucifolia auct. non 

(With.) Carruth.) 

10 donniana (Sm.) Müll.Hal. 

Subgenus Blindiadelphus Lindb. 

11 polaris Berggr. 

12 subimmersa Lindb. 

Subgenus Megalosporia Vitt 

13 austriaca T.Schauer 

14 carniolica (Breidl. & Beck) Nyholm (Trochobryum 

carniolicum Breidl. & Beck) 

15 irrigata (H.K.G.Paul) Ochyra & Gos (S. trifaria var. 

irrigata H.K.G.Paul) [87] 

16 oelandica C.E.O.Jensen & Medelius 

17 patula (Lindb.) I.Hagen (S. alpestris T.Schauer, 

S. patula var. alpestris (T.Schauer) Gos & Ochyra, 

S. tristichoides var. patula (Lindb.) Broth.) 

18 trifaria (Brid.) Lindb. 

19 tristichoides Kindb. 

Archidiales Limpr. 

Archidiaceae Schimp. 

40 Archidium Brid. 

1 alternifolium (Hedw.) Mitt. 

Dicranales H.Philib. ex M.Fleisch. 

Fissidentaceae Schimp. 

41 Fissidens Hedw. 


Subgenus Pachyfissidens (Müll.Hal.) Kindb. 

section Pachyfissidens 

1 adianthoides Hedw. 

2 azoricus (P.de la Varde) Bizot 

3 dubius P.Beauv. (F. cristatus Wilson ex Mitt., 

F. decipiens De Not.) [88] 

4 grandifrons Brid. 

5 luisieri P.de la Varde [89] 

6 osmundoides Hedw. 

7 polyphyllus Wilson ex Bruch & Schimp. [90] 

8 serrulatus Brid. [91] 

9 taxifolius Hedw. 

subsp. pallidicaulis (Mitt.) Mönk. (F. pallidicaulis 

Mitt., F. taxifolius var. pallidicaulis (Mitt.) Corb.) 

subsp. taxifolius 

section Amblyothallia (Müll.Hal.) Pursell & 

Brugg.-Nann. 

10 asplenioides Hedw. 

Subgenus Octodiceras (Brid.) Broth. 

11 fontanus (Bach.Pyl.) Steud. (Octodiceras fontanum 

(Bach.Pyl.) Lindb.) 

Subgenus Fissidens [92] 

12 arcticus Bryhn [93] 

13 arnoldii R.Ruthe 

14 bryoides Hedw. [94] 

var. bryoides 

var. caespitans Schimp. (F. bryoides var. curnovii 

(Mitt.) J.J.Amann, F. curnovii Mitt.) [95] 

15 coacervatus Brugg.-Nann. 

16 crassipes Wilson ex Bruch & Schimp. 

subsp. crassipes (F. crassipes var. rufipes Schimp.) 

subsp. warnstorfii (M.Fleisch.) Brugg.-Nann. 

(F. crassipes var. philibertii Besch., F. mouretii Corb.) 

17 crispus Mont. (F. herzogii R.Ruthe ex Herzog, 

F. limbatus Sull., F. minutulus Sull.) [96] 

18 curvatus Hornsch. (F. algarvicus Solms) 

19 gracilifolius Brugg.-Nann. & Nyholm (F. viridulus var. 

tenuifolius (Boulay) A.J.E.Sm.) 

20 gymnandrus Büse [97] 

21 jansenii Sérgio & Pursell [98] 

22 microstictus Dixon & Luisier [99] 

23 monguillonii Thér. 

24 nobreganus Dixon & Luisier [100] 

25 ovatifolius R.Ruthe (F. perssonii P.de la Varde) [101] 

26 pusillus (Wilson) Milde (F. exiguus auct. eur. partim 

[103], F. viridulus var. pusillus Wilson) [102] 

27 rivularis (Spruce) Schimp. 

28 rufulus Bruch & Schimp. 

29 sublimbatus Grout [104] 

30 sublineaefolius (P.de la Varde) Brugg.-Nann. 

31 viridulus (Sw. ex anon.) Wahlenb. [105] [106] 

var. incurvus (Starke ex Röhl.) Waldh. (F. incurvus 

Starke ex Röhl., F. incurvus var. tamarindifolius 

(Turner) Braithw.) [107] 

var. viridulus (F. bambergeri Milde [108], F. bryoides 

var. inconstans (Schimp.) R.Ruthe, F. bryoides var. 

viridulus (Sw. ex anon.) Broth., F. exiguus auct. eur.


partim [92] [103], F. haraldii (Lindb.) Limpr., F. 

limbatus var. bambergeri (Schimp. ex Milde) Düll, F. 

marginatulus Meln. [109], F. viridulus var. bambergeri 

(Schimp. ex Milde) Waldh., F. viridulus var. 

haraldii (Lindb.) C.E.O.Jensen, F. viridulus var. 

intralimbatulus (R.Ruthe) Düll) [92] 

Subgenus Aloma Kindb. 

32 celticus Paton 

33 exilis Hedw. 

34 serratus Müll.Hal. (F. papillosus Sande Lac.) [110] 

Ditrichaceae Limpr., nom. cons. 

42 Ceratodon Brid. 

1 conicus (Hampe) Lindb. (C. purpureus var. conicus 

(Hampe) Husn.) 

2 purpureus (Hedw.) Brid. [111] 

subsp. purpureus (C. purpureus var. brevifolius Milde, 

C. purpureus var. flavisetus Limpr., C. purpureus var. 

pallidisetus Luisier, C. purpureus var. xanthopus Sull.) 

[112] 

subsp. stenocarpus (Bruch. & Schimp. ex Müll.Hal.) 

Dixon (C. corsicus Bruch & Schimp, C. stenocarpus 

Bruch & Schimp. ex Müll.Hal.) [112] 

43 Cheilothela Broth. 

1 chloropus (Brid.) Broth. 

44 Cleistocarpidium Ochyra & Bednarek-Ochyra 

1 palustre (Bruch & Schimp.) Ochyra & Bednarek- 

Ochyra (Pleuridium palustre (Bruch & Schimp.) Bruch 

& Schimp., Sporledera palustris (Bruch & Schimp.) 

Hampe) 

45 Distichium Bruch & Schimp., nom. cons. 

1 capillaceum (Hedw.) Bruch & Schimp. (D. capillaceum 

var. compactum (Huebener) Dalla Torre & Sarnth.) 

2 hagenii Ryan ex H.Philib. 

3 inclinatum (Hedw.) Bruch & Schimp. 

46 Ditrichum Timm ex Hampe, nom. cons. 

1 cornubicum Paton 

2 flexicaule (Schwägr.) Hampe 

3 gracile (Mitt.) Kuntze (D. crispatissimum (Müll.Hal.) 

Paris, D. flexicaule var. longifolium (J.E.Zetterst.) 

I.Hagen, D. flexicaule var. sterile (De Not.) Limpr., 

D. giganteum R.S.Williams) [113] 

4 heteromallum (Hedw.) E.Britton (D. homomallum 

(Hedw.) Hampe) 

5 lineare (Sw.) Lindb. (D. vaginans (Sull.) Hampe) 

6 pallidum (Hedw.) Hampe 

7 plumbicola Crundw. 

8 punctulatum Mitt. (Campylopus brevipilus var. 

marginatulus (Geh.) Paris, Campylopus madeirensis 

Luisier, Campylopus marginatulus Geh.) [114] 

9 pusillum (Hedw.) Hampe (D. tortile (Schrad.) Brockm.) 

10 subulatum Hampe 

11 zonatum (Brid.) Kindb. (D. heteromallum var. zonatum 

(Brid.) Podp., D. zonatum var. scabrifolium Dixon) 

47 Pleuridium Rabenh., nom. cons. [115] 

1 acuminatum Lindb. 

2 subulatum (Hedw.) Rabenh. (P. alternifolium auct. 

non (Dicks. ex Hedw.) Rabenh.) 

48 Pseudephemerum (Lindb.) I.Hagen 

1 nitidum (Hedw.) Loeske (Pleuridium nitidum (Hedw.) 

Rabenh., P. axillare (Dicks.) I.Hagen) 

49 Rhamphidium Mitt. 

1 purpuratum Mitt. [116] 

50 Saelania Lindb. 

1 glaucescens (Hedw.) Broth. 

51 Trichodon Schimp. 

1 cylindricus (Hedw.) Schimp. (Ditrichum cylindricum 

(Hedw.) Grout, Ditrichum cylindricum var. oblongum 

(Lindb.) C.E.O.Jensen) 

Bruchiaceae Schimp. 

52 Bruchia Schwägr. 

1 flexuosa (Schwägr.) Müll.Hal. (B. trobasiana De Not.) 

2 vogesiaca Nestl. ex Schwägr. 

53 Trematodon Michx. 

1 ambiguus (Hedw.) Hornsch. 

2 brevicollis Hornsch. [32] 

3 laetevirens Hakelier & J.-P.Frahm 

4 longicollis Michx. [32] 

5 perssoniorum P.Allorge & Thér. ex V.Allorge [117] 

Rhabdoweisiaceae Limpr. 

54 Amphidium Schimp., nom. cons. 

1 lapponicum (Hedw.) Schimp. 

2 mougeotii (Schimp.) Schimp. 

3 tortuosum (Hornsch.) Cufod. (A. curvipes (Müll.Hal.) 

Broth.) [118] 

55 Arctoa Bruch & Schimp. 

1 anderssonii Wich. (A. fulvella var. anderssonii (Wich.) Grout) 

2 fulvella (Dicks.) Bruch & Schimp. 

3 hyperborea (Gunnerus ex Dicks.) Bruch & Schimp., 

nom. cons. 

56 Cnestrum I.Hagen 

1 alpestre (Wahlenb. ex Huebener) Nyholm ex 

Mogensen (Cynodontium alpestre (Wahlenb. ex 

Huebener) Milde) 

2 glaucescens (Lindb. & Arnell) Holmen ex Mogensen 

& Steere 

3 schisti (F.Weber & D.Mohr) I.Hagen, nom. cons. 

57 Cynodontium Bruch & Schimp., nom. cons. 

1 asperifolium (Lindb. ex Arnell) Paris [119] 

2 bruntonii (Sm.) Bruch & Schimp. (Oreoweisia bruntonii 

(Sm.) Milde) 

3 fallax Limpr. 

4 gracilescens (F.Weber & D.Mohr) Schimp. 

5 jenneri (Schimp.) Stirt. 

6 polycarpon (Hedw.) Schimp. [120] 

7 strumiferum (Hedw.) Lindb. (C. polycarpon var. 

strumiferum (Hedw.) Schimp.) 

8 suecicum (Arnell & C.E.O.Jensen) I.Hagen 

9 tenellum (Schimp.) Limpr. (C. torquescens Limpr.) 

58 Dichodontium Schimp. 

1 flavescens (Dicks.) Lindb. (D. pellucidum var. 

flavescens (Dicks.) Moore) 

2 palustre (Dicks.) M.Stech (Anisothecium palustre 

(Dicks.) I.Hagen, Dicranella palustris (Dicks.) 

Crundw., Diobelonella palustris (Dicks.) Ochyra) [121]

3 pellucidum (Hedw.) Schimp. (D. pellucidum var. 

fagimontanum Brid., D. pellucidum var. 

propaguliferum (Correns) Casares-Gil) 

59 Dicranoweisia Milde [122] 

1 cirrata (Hedw.) Lindb. 

2 compacta (Schleich. ex Schwägr.) Schimp. (D. crispula 

var. compacta (Schwägr.) Lindb., Hymenoloma 

compactum (Schwägr.) Ochyra) 

3 crispula (Hedw.) Milde (Hymenoloma crispulum 

(Hedw.) Ochyra) 

60 Glyphomitrium Brid. 

1 daviesii (Dicks.) Brid. 

61 Kiaeria I.Hagen 

1 blyttii (Bruch & Schimp.) Broth. 

2 falcata (Hedw.) I.Hagen 

3 glacialis (Berggr.) I.Hagen 

4 riparia (H.Lindb.) M.F.V.Corley (Dicranella riparia 

(H.Lindb.) Martensson & Nyholm, K. starkei var. 

riparia I.Hagen, Oncophorus riparius H.Lindb.) 

5 starkei (F.Weber & D.Mohr) I.Hagen 

62 Oncophorus (Brid.) Brid. 

1 elongatus (I.Hagen) Hedenäs (O. wahlenbergii var. 

elongatus I.Hagen) [123] 

2 virens (Hedw.) Brid. 

3 wahlenbergii Brid. 

var. compactus (Bruch & Schimp.) Braithw. 

(O. compactus (Bruch & Schimp.) Kindb.) 

var. wahlenbergii 

63 Oreas Brid. 

1 martiana (Hoppe & Hornsch.) Brid. 

64 Oreoweisia (Bruch & Schimp.) De Not. 

1 torquescens (Hornsch. ex Brid.) Wijk & Margad. 

(O. serrulata (Funck) De Not.) 

65 Rhabdoweisia Bruch & Schimp. 

1 crenulata (Mitt.) H.Jameson 

2 crispata (Dicks.) Lindb. (R. denticulata (Brid.) Bruch 

& Schimp., R. kusenevae Broth. [124]) 

3 fugax (Hedw.) Bruch & Schimp. (R. striata (Schrad.) 

Lindb.) 

Schistostegaceae Schimp. [125] 

66 Schistostega D.Mohr 

1 pennata (Hedw.) F.Weber & D.Mohr 

Dicranaceae Schimp. 

67 Aongstroemia Bruch & Schimp., nom. cons. 

1 longipes (Sommerf.) Bruch & Schimp. 

68 Dicranella (Müll.Hal.) Schimp., nom. cons. 

1 campylophylla (Taylor) A.Jaeger [126] 

2 cerviculata (Hedw.) Schimp. 

3 crispa (Hedw.) Schimp. (Anisothecium vaginale 

(Dicks.) Loeske) 

4 grevilleana (Brid.) Schimp. (Anisothecium grevilleanum 

(Brid.) Lindb.) 

5 heteromalla (Hedw.) Schimp. 

6 howei Renauld & Cardot 

7 humilis R.Ruthe (Anisothecium humile (R.Ruthe) 

Lindb., Anisothecium rigidulum (Hedw.) 

C.E.O.Jensen) 


8 rufescens (Dicks.) Schimp. (Anisothecium rufescens 

(Dicks.) Lindb.) 

9 schreberiana (Hedw.) Dixon (Anisothecium 

schreberianum (Hedw.) Dixon) 

10 staphylina H.Whitehouse (Anisothecium staphylinum 

(H.Whitehouse) Sipman, Rubers & Riemann) 

11 subulata (Hedw.) Schimp. (D. curvata (Hedw.) 

Schimp., D. secunda Lindb.) 

12 varia (Hedw.) Schimp. (Anisothecium rubrum Lindb., 

Anisothecium varium (Hedw.) Mitt.) 

69 Dicranum Hedw. [127] 

section Dicranum 

1 bonjeanii De Not. (D. palustre Bruch & Schimp., 

D. undulatum Turner, hom. illeg.) 

2 crassifolium Sérgio, Ochyra & Séneca [128] 

3 leioneuron Kindb. 

4 majus Sm. 

5 polysetum Sw. ex anon. (D. rugosum Brid., D. undulatum 

Ehrh. ex F.Weber & D.Mohr, hom. illeg.) [106] 

6 scoparium Hedw. 

7 transsylvanicum Lüth [129] 

section Spuria Bruch & Schimp. 

8 acutifolium (Lindb. & Arnell) C.E.O.Jensen (D. bergeri 

var. acutifolium Lindb. & Arnell, D. muehlenbeckii var. 

acutifolium (Lindb. & Arnell) Nyholm) 

9 brevifolium (Lindb.) Lindb. (D. muehlenbeckii var. 

brevifolium Lindb., D. muehlenbeckii var. cirrhatum 

(Schimp.) Lindb.) 

10 dispersum Engelmark [130] 

11 drummondii Müll.Hal. (D. elatum Lindb., D. robustum 

Bruch & Schimp.) 

12 spurium Hedw. 

13 undulatum Schrad. ex Brid. (D. affine Funck, D. 

bergeri Blandow) [131] 

section Fuscescentiformia (Kindb.) Ochyra 

14 flexicaule Brid. (D. congestum Brid., D. fuscescens var. 

congestum (Brid.) Kindb., D. fuscescens var. flexicaule 

(Brid.) Wilson) 

15 fuscescens Sm. 

section Convolutifolia (Kindb.) Ochyra 

16 angustum Lindb. [132] 

17 muehlenbeckii Bruch & Schimp. 

18 spadiceum J.E.Zetterst. (D. muehlenbeckii var. neglectum 

(De Not.) Pfeff., D. muehlenbeckii var. spadiceum 

(J.E.Zetterst.) Podp., D. neglectum Jur. ex De Not.) 

section Elongata I.Hagen 

19 elongatum Schleich. ex Schwägr. (D. elongatum var. 

sendtneri (Limpr.) Mönk., D. sendtneri Limpr. [133]) 

20 fragilifolium Lindb. 

21 groenlandicum Brid. (D. elongatum subsp. 

groenlandicum (Brid.) Mönk.) 

22 laevidens R.S.Williams [132] 

section Crassinervia G.Roth 

23 canariense Hampe ex Müll.Hal. (D. scottianum subsp. 

canariense (Hampe) Corb.) [134] 

24 fulvum Hook. (Orthodicranum fulvum (Hook.) 

G.Roth ex Casares-Gil)


25 scottianum Turner ex R.Scott (Orthodicranum scottianum 

(Turner ex R.Scott) G.Roth ex Casares-Gil) [79] 

26 viride (Sull. & Lesq.) Lindb. 

section Montana Hartm. 

27 flagellare Hedw. (Orthodicranum flagellare (Hedw.) 

Loeske) 

28 montanum Hedw. (Orthodicranum montanum (Hedw.) 

Loeske) 

29 tauricum Sapjegin (D. strictum Schleich. ex D.Mohr, 

hom. illeg., Orthodicranum tauricum (Sapjegin) 

Smirnova) 

70 Paraleucobryum (Limpr.) Loeske 

1 enerve (Thed.) Loeske 

2 longifolium (Hedw.) Loeske 

3 sauteri (Bruch & Schimp.) Loeske (P. longifolium var. 

sauteri (Bruch & Schimp.) C.E.O.Jensen) 

Leucobryaceae Schimp. 

71 Atractylocarpus Mitt., nom. cons. 

1 alpinus (Schimp. ex Milde) Lindb. (Metzlerella alpina 

(Schimp. ex Milde) I.Hagen, Metzleria alpina Schimp. 

ex Milde) 

72 Campylopus Brid. 

1 atrovirens De Not. (C. atrovirens var. adustus (De 

Not.) Husn., C. atrovirens var. falcatus Braithw., C. 

atrovirens var. gracilis Dixon) 

2 brevipilus Bruch & Schimp. 

3 cygneus (Hedw.) Brid. (C. setaceus Cardot) 

4 flaccidus Renauld & Cardot 

5 flexuosus (Hedw.) Brid. 

6 fragilis (Brid.) Bruch & Schimp. 

7 gracilis (Mitt.) A.Jaeger (C. schwarzii Schimp.) 

8 incrassatus Müll.Hal. (C. eximius Reichardt) [135] 

9 introflexus (Hedw.) Brid. 

10 oerstedianus (Müll.Hal.) Mitt. (C. mildei Limpr.) 

11 pilifer Brid. 

12 pyriformis (Schultz) Brid. (C. pyriformis var. azoricus 

(Mitt.) M.F.V.Corley) 

13 schimperi Milde (C. subulatus var. schimperi (Milde) 

Husn.) 

14 setifolius Wilson 

15 shawii Wilson (C. carreiroanus Cardot) 

16 subulatus Schimp. ex Milde 

73 Dicranodontium Bruch & Schimp. 

1 asperulum (Mitt.) Broth. 

2 denudatum (Brid.) E.Britton (D. denudatum var. 

alpinum (Schimp.) I.Hagen) 

3 subporodictyon Broth. (Campylopus subporodictyon 

(Broth.) B.H.Allen & Ireland, Dicranum 

subporodictyon (Broth.) C.Gao & T.Cao) [136] 

4 uncinatum (Harv.) A.Jaeger (D. circinatum (Wilson) 

Schimp.) 

74 Leucobryum Hampe 

1 albidum (P.Beauv.) Lindb. [137] 

2 glaucum (Hedw.) A˚ ngstr. 

3 juniperoideum (Brid.) Müll.Hal. (L. albidum auct. eur. 

non (P.Beauv.) Lindb., L. minus Hampe) 

75 Microcampylopus (Müll.Hal.) M.Fleisch. 

1 laevigatus (Thér.) Giese & J.-P.Frahm 

Calymperaceae Kindb. [138] 

76 Calymperes Sw. ex F.Weber 

1 erosum Müll.Hal. (C. sommieri Bott.) 

Pottiales M.Fleisch. [139] 

Pottiaceae Schimp., nom. cons. [140] 

Timmielloideae R.H.Zander 

77 Timmiella (De Not.) Limpr. 

1 anomala (Bruch & Schimp.) Limpr. 

2 barbuloides (Brid.) Mönk. 

3 flexiseta (Bruch) Limpr. 

Merceyoideae Broth. 

78 Scopelophila (Mitt.) Lindb. 

1 cataractae (Mitt.) Broth. 

2 ligulata (Spruce) Spruce 

Trichostomoideae (Limpr.) Broth. 

79 Anoectangium Schwägr., nom. cons. 

1 aestivum (Hedw.) Mitt. (A. angustifolium Mitt.) 

2 handelii Schiffn. [141] 

80 Aschisma Lindb. 

1 carniolicum (F.Weber & D.Mohr) Lindb. 

2 cuynetii (Bizot & R.B.Pierrot) J.Guerra & M.J.Cano 

(Phascum cuynetii Bizot & R.B.Pierrot) 

81 Ephemerum Hampe, nom. cons. 

1 cohaerens (Hedw.) Hampe 

2 hibernicum Holyoak & V.S.Bryan [142] 

3 minutissimum Lindb. (E. serratum var. 

angustifolium (Bruch & Schimp.) Bruch & Schimp., 

E. serratum var. 

minutissimum (Lindb.) Grout) [143] 

4 recurvifolium (Dicks.) Boulay 

5 serratum (Hedw.) Hampe (E. serratum var. rutheanum 

(Schimp.) Jur.) [143] 

6 sessile (Bruch) Müll.Hal. 

7 spinulosum Bruch & Schimp. ex Schimp. [144] 

8 stellatum H.Philib. 

82 Eucladium Bruch & Schimp. 

1 verticillatum (With.) Bruch & Schimp. [145] 

var. angustifolium Lindb. 

var. verticillatum 

83 Gymnostomum Nees & Hornsch., nom. cons. [146] 

1 aeruginosum Sm. 

var. aeruginosum 

var. obscurum J.Guerra [147] 

2 boreale Nyholm & Hedenäs 

3 calcareum Nees & Hornsch. 

4 lanceolatum M.J.Cano, Ros & J.Guerra [146] 

5 viridulum Brid. 

84 Gyroweisia Schimp., nom. cons. 

1 reflexa (Brid.) Schimp. 

2 tenuis (Hedw.) Schimp. (G. tenuis var. badia Limpr.) 

85 Hymenostylium Brid. 

1 recurvirostrum (Hedw.) Dixon (Gymnostomum 

recurvirostrum Hedw.) [148] 

var. insigne (Dixon) E.B.Bartram (H. insigne (Dixon) 

Podp.) 

var. recurvirostrum

86 Hyophila Brid., nom. cons. 

1 involuta (Hook.) A.Jaeger 

87 Leptobarbula Schimp. 

1 berica (De Not.) Schimp. 

88 Micromitrium Austin 

1 tenerum (Bruch & Schimp.) Crosby 

89 Molendoa Lindb. 

1 hornschuchiana (Hook.) Lindb. ex Limpr. 

(Anoectangium hornschuchianum (Hook.) Funck ex 

Hornsch.) 

2 schliephackei (Schlieph.) R.H.Zander (Anoectangium 

schliephackei (Schlieph.) Paris) 

3 sendtneriana (Bruch & Schimp.) Limpr. 

(Anoectangium sendtnerianum Bruch & Schimp.) 

4 taeniatifolia Herzog (Anoectangium taeniatifolium 

(Herzog) M.O.Hill) 

5 tenuinervis Limpr. (Anoectangium tenuinerve (Limpr.) 

Paris) 

6 warburgii (Crundw. & M.O.Hill) R.H.Zander 

(Anoectangium warburgii Crundw. & M.O.Hill) 

90 Oxystegus (Broth.) Hilp. 

1 hibernicus (Mitt.) Hilp. (Trichostomum hibernicum 

(Mitt.) Dixon) 

2 tenuirostris (Hook. & Taylor) A.J.E.Sm. 

(Trichostomum tenuirostre (Hook. & Taylor) Lindb.) 

91 Pleurochaete Lindb. [149] 

1 squarrosa (Brid.) Lindb. 

92 Pottiopsis Blockeel & A.J.E.Sm. 

1 caespitosa (Brid.) Blockeel & A.J.E.Sm. (Pottia 

caespitosa (Brid.) Müll.Hal., Trichostomum 

caespitosum (Brid.) Jur.) 

93 Splachnobryum Müll.Hal. 

1 obtusum (Brid.) Müll.Hal. (S. delicatulum Broth.) [150] 

94 Tortella (Müll.Hal.) Limpr., nom. cons. [151] 

1 alpicola Dixon [152] 

2 bambergeri (Schimp.) Broth. (T. tortuosa var. 

bambergeri (Schimp.) Düll) [153] 

3 flavovirens (Bruch) Broth. (T. flavovirens var. minor 

Lindb.) 

var. flavovirens (T. flavovirens var. viridiflava (De 

Not.) Casares-Gil) 

var. glareicola (T.A.Chr.) Crundw. & Nyholm (T. 

glareicola T.A.Chr.) 

var. papillosissima Sérgio & Casas 

4 fragilis (Hook. & Wilson) Limpr. 

5 humilis (Hedw.) Jenn. 

6 inclinata (R.Hedw.) Limpr. 

var. densa (Lorentz & Molendo) Limpr. (T. densa 

(Lorentz & Molendo) Crundw. & Nyholm) [154] 

var. inclinata 

7 inflexa (Bruch) Broth. 

8 limbata (Schiffn.) Geh. & Herzog [155] 

9 limosella (Stirt.) P.W.Richards & E.C.Wallace [156] 

10 nitida (Lindb.) Broth. (T. cirrifolia (Mitt.) Broth.) 

[157] 

11 rigens Alberts. 

12 tortuosa (Hedw.) Limpr. 


var. fragilifolia (Jur.) Limpr. 

var. tortuosa 

95 Trichostomum Bruch, nom. cons. [158] 

1 arcticum Kaal. 

2 brachydontium Bruch (Hyophila treleasei Cardot [159]) 

3 crispulum Bruch 

4 triumphans De Not. (T. pallidisetum H.Müll., Weissia 

triumphans (De Not.) M.O.Hill) 

96 Weissia Hedw. (Astomum Hampe) [160] [161] 

1 brachycarpa (Nees & Hornsch.) Jur. (Hymenostomum 

microstomum (Hedw.) R.Br. ex Nees & Hornsch.) 

2 condensa (Voit) Lindb. 

var. armata (Thér. & Trab.) M.J.Cano, Ros & 

J.Guerra (W. papillosissima Laz.) 

var. condensa 

3 controversa Hedw. 

var. controversa 

var. crispata (Nees & Hornsch.) Nyholm (W. fallax 

Sehlm.) 

var. densifolia (Bruch & Schimp.) Wilson 

4 levieri (Limpr.) Kindb. (Astomum levieri Limpr.) 

5 longifolia Mitt. (Astomum crispum (Hedw.) Hampe) 

6 multicapsularis (Sm.) Mitt. (Astomum multicapsulare 

(Sm.) Bruch & Schimp.) 

7 perssonii Kindb. 

8 rostellata (Brid.) Lindb. 

9 rutilans (Hedw.) Lindb. 

10 squarrosa (Nees & Hornsch.) Müll.Hal. 

(Hymenostomum squarrosum Nees & Hornsch.) 

11 sterilis W.E.Nicholson 

12 tyrrhena M.Fleisch. 

13 wimmeriana (Sendtn.) Bruch & Schimp. 

(W. controversa var. wimmeriana (Sendtn.) Blockeel & 

A.J.E.Sm., W. wimmeriana subsp. pallescens (Schimp. 

ex Besch.) Giacom.) [162] 

Pottioideae (Limpr.) Broth. 

97 Acaulon Müll.Hal. 

1 casasianum Brugués & H.A.Crum 

2 dertosense Casas, Sérgio, Cros & Brugués 

3 fontiquerianum Casas & Sérgio 

4 mediterraneum Limpr. (A. muticum var. mediterraneum 

(Limpr.) Sérgio) [163] 

5 muticum (Hedw.) Müll.Hal. 

6 piligerum (De Not.) Limpr. 

7 triquetrum (Spruce) Müll.Hal. 

98 Aloina Kindb., nom. cons. 

1 aloides (Koch ex Schultz) Kindb. 

2 ambigua (Bruch & Schimp.) Limpr. (A. aloides var. 

ambigua (Bruch & Schimp.) E.J.Craig) 

3 bifrons (De Not.) Delgad. 

4 brevirostris (Hook. & Grev.) Kindb. 

5 humilis M.T.Gallego, M.J.Cano & Ros [164] 

6 obliquifolia (Müll.Hal.) Broth. (A. rigida var. 

mucronulata (Bruch & Schimp.) Limpr., A. rigida var. 

obliquifolia (Müll.Hal.) Delgad.) [165] 

7 rigida (Hedw.) Limpr. 

99 Barbula Hedw., nom. cons. [139] [166]


1 bicolor (Bruch & Schimp.) Lindb. 

2 bolleana (Müll.Hal.) Broth. (B. ehrenbergii (Lorentz) 

M.Fleisch.) 

3 convoluta Hedw. (Streblotrichum convolutum (Hedw.) 

P.Beauv.) 

var. convoluta 

var. sardoa Schimp. (B. commutata Jur., B. convoluta 

var. commutata (Jur.) Husn., B. convoluta var. 

uliginosa (Limpr.) Limpr., B. sardoa (Schimp.) 

J.-P.Frahm, nom. inval.) [167] 

4 crocea (Brid.) F.Weber & D.Mohr 

5 enderesii Garov. 

6 indica (Hook.) Spreng. 

7 unguiculata Hedw. 

100 Bryoerythrophyllum P.C.Chen 

1 alpigenum (Venturi) P.C.Chen (B. recurvirostrum var. 

dentatum (Schimp.) H.A.Crum, Steere & 

L.E.Anderson) 

2 caledonicum D.G.Long 

3 campylocarpum (Müll.Hal.) H.A.Crum (B. lusitanicum 

(Cardot & Dixon) M.O.Hill, B. machadoanum (Sérgio) 

M.O.Hill, Hyophila lusitanica Cardot & Dixon, 

Hyophila machadoana Sérgio) [168] 

4 ferruginascens (Stirt.) Giacom. 

5 inaequalifolium (Taylor) R.H.Zander 

6 recurvirostrum (Hedw.) P.C.Chen 

7 rubrum (Jur. ex Geh.) P.C.Chen 

101 Cinclidotus P.Beauv., nom. cons. [169] 

1 aquaticus (Hedw.) Bruch & Schimp. 

2 confertus Lüth [170] 

3 danubicus Schiffn. & Baumgartner 

4 fontinaloides (Hedw.) P.Beauv. 

5 pachylomoides Bizot 

6 riparius (Host ex Brid.) Arn. 

7 vivesii Ederra [171] 

102 Crossidium Jur., nom. cons. 

1 aberrans Holz. & E.B.Bartram 

2 crassinerve (De Not.) Jur. 

3 davidai Catches. [172] 

4 geheebii (Broth.) Broth. [173] 

5 laevipilum Thér. & Trab. [174] 

6 laxefilamentosum W.Frey & Kürschner [175] 

7 squamiferum (Viv.) Jur. 

var. pottioideum (De Not.) Mönk. 

var. squamiferum 

103 Dialytrichia (Schimp.) Limpr. 

1 fragilifolia (Bizot & J.Roux) F.Lara (D. mucronata 

var. fragilifolia Bizot & J.Roux) [176] 

2 mucronata (Brid.) Broth. (Cinclidotus mucronatus 

(Brid.) Guim.) 

104 Didymodon Hedw. [177] [178] [179] 

1 acutus (Brid.) K.Saito (Barbula acuta (Brid.) Brid.) 

2 asperifolius (Mitt.) H.A.Crum, Steere & L.E.Anderson 

(Barbula asperifolia Mitt., Barbula kneuckeri Loeske & 

Osterwald, D. ferrugineus var. kneuckeri (Loeske & 

Osterwald) Düll, D. rufus Lorentz) [180] 

3 australasiae (Hook. & Grev.) R.H.Zander (D. aaronis 

(Lorentz) J.Guerra, D. incrassatus (Lindb.) Broth., 

Trichostomopsis aaronis (Lorentz) S.Agnew & 

C.C.Towns., Trichostomopsis australasiae (Hook. & 

Grev.) H.Rob.) [181] 

4 bistratosus Hébr. & R.B.Pierrot [182] 

5 brachyphyllus (Sull.) R.H.Zander [183] 

6 cordatus Jur. (Barbula cordata (Jur.) Loeske) 

7 erosus J.A.Jiménez & J.Guerra [184] 

8 fallax (Hedw.) R.H.Zander (Barbula adriatica 

Baumgartner, Barbula fallax Hedw.) [180] 

9 ferrugineus (Schimp. ex Besch.) M.O.Hill (Barbula 

reflexa (Brid.) Brid.) 

10 giganteus (Funck) Jur. (Barbula gigantea Funck, 

Geheebia gigantea (Funck) Boulay) [180] 

11 glaucus Ryan (Barbula rigidula subsp. verbana 

(W.E.Nicholson & Dixon) Podp., Barbula rigidula 

var. glauca (Ryan) J.J.Amann) 

12 icmadophilus (Schimp. ex Müll.Hal.) K.Saito 

(Barbula acuta var. icmadophila (Schimp. ex 

Müll.Hal.) H.A.Crum) 

13 insulanus (De Not.) M.O.Hill (Barbula cylindrica 

(Taylor) Schimp., Barbula vinealis var. cylindrica 

(Taylor) Boulay, D. vinealis var. flaccidus (Bruch & 

Schimp.) R.H.Zander) 

14 johansenii (R.S.Williams) H.A.Crum (Barbula 

johansenii R.S.Williams) 

15 lamyanus (Schimp.) Thér. [185] 

16 luridus Hornsch. (Barbula lurida Hornsch., D. trifarius 

auct. non (Hedw.) Röhl.) 

17 maximus (Syed & Crundw.) M.O.Hill (Barbula reflexa 

var. robusta Braithw.) [180] 

18 nicholsonii Culm. 

19 rigidulus Hedw. (Barbula rigidula (Hedw.) Milde, D. 

mamillosus (Crundw.) M.O.Hill) [186] 

20 sicculus M.J.Cano, Ros, García-Zamora & J.Guerra 

[187] 

21 sinuosus (Mitt.) Delogne (Barbula sinuosa (Mitt.) 

Grav.) 

22 spadiceus (Mitt.) Limpr. (Barbula insidiosa Jur. & 

Milde, Barbula spadicea (Mitt.) Braithw., D. 

barbuloides Lib. ex Marchal, D. zetterstedtii Schimp., 

Limneria viridula Stirt.) [180] 

23 subandreaeoides (Kindb.) R.H.Zander (D. rigidulus 

subsp. andreaeoides (Limpr.) Wijk & Margad.) [188] 

24 tomaculosus (Blockeel) M.F.V.Corley 

25 tophaceus (Brid.) Lisa (Barbula tophacea (Brid.) Mitt., 

D. bosniacus Głow., D. spadiceus var. siluricus Velen.) 

[180] 

26 umbrosus (Müll.Hal.) R.H.Zander (D. australasiae 

var. umbrosus (Müll.Hal.) R.H.Zander, D. trivialis 

(Müll.Hal.) J.Guerra, Trichostomopsis trivialis 

(Müll.Hal.) H.Rob., Trichostomopsis umbrosa 

(Müll.Hal.) H.Rob.) [189] 

27 vinealis (Brid.) R.H.Zander (Barbula vinealis Brid.) 

105 Hennediella Paris

1 heimii (Hedw.) R.H.Zander (Desmatodon heimii 

(Hedw.) Mitt., Pottia heimii (Hedw.) Hampe) 

var. arctica (Lindb.) R.H.Zander (Desmatodon heimii 

var. arcticum (Lindb.) H.A.Crum) 

var. heimii 

2 macrophylla (R.Br.bis) Paris [190] 

3 stanfordensis (Steere) Blockeel 

106 Hilpertia R.H.Zander 

1 velenovskyi (Schiffn.) R.H.Zander 

107 Leptodontium (Müll.Hal.) Lindb. 

1 flexifolium (Dicks.) Hampe 

2 gemmascens (Mitt.) Braithw. 

3 styriacum (Jur.) Limpr. 

108 Leptophascum (Müll.Hal.) J.Guerra & M.J.Cano 

1 leptophyllum (Müll.Hal.) J.Guerra & M.J.Cano 

(Chenia leptophylla (Müll.Hal.) R.H.Zander, Chenia 

rhizophylla (Sakurai) R.H.Zander, Phascum 

leptophyllum Müll.Hal., Tortula rhizophylla 

(Sakurai) Z.Iwats. & K.Saito) 

109 Microbryum Schimp. [191] 

1 curvicollum (Hedw.) R.H.Zander (M. piptocarpum 

(Durieu & Mont.) J.Guerra & M.J.Cano, Phascum 

curvicollum Hedw., Phascum piptocarpum Durieu & 

Mont.) [32] [19] 

2 davallianum (Sm.) R.H.Zander (Pottia commutata 

Limpr., Pottia conica (Schleich. ex Schwägr.) Fürnr. 

ex Paris, Pottia davalliana (Sm.) C.E.O.Jensen, Pottia 

starckeana subsp. conica (Schleich. ex Schwägr.) 

D.F.Chamb., Pottia starckeana subsp. minutula 

(Schwägr.) D.F.Chamb., Pottia starckeana var. 

minutula (Schwägr.) Corb.) [193] 

3 floerkeanum (F.Weber & D.Mohr) Schimp. (Phascum 

floerkeanum F.Weber & D.Mohr) 

4 fosbergii (E.B.Bartram) Ros, O.Werner & Rams (M. 

starckeanum var. fosbergii (E.B.Bartram) R.H.Zander, 

Pottia 6andalusica Ros & R.Oliva) [194] 

5 longipes (J.Guerra, J.J.Martínez & Ros) R.H.Zander 

(Phascum longipes J.Guerra, J.J.Martínez & Ros) 

6 rectum (With.) R.H.Zander (Pottia recta (With.) Mitt.) 

7 starckeanum (Hedw.) R.H.Zander (Pottia mutica 

Venturi, Pottia starckeana (Hedw.) Müll.Hal., Pottia 

starckeana var. brachyodus (Bruch & Schimp.) 

Müll.Hal.) [193] [195] 

110 Paraleptodontium D.G.Long 

1 recurvifolium (Taylor) D.G.Long (Trichostomum 

recurvifolium (Taylor) R.H.Zander) 

111 Phascum Hedw. 

1 cuspidatum Hedw. (Tortula acaulon (With.) 

R.H.Zander, Tortula atherodes R.H.Zander) 

var. cuspidatum (P. cuspidatum var. curvisetum 

(Dicks.) Nees & Hornsch., P. cuspidatum var. 

mitraeforme Limpr., Tortula atherodes var. curviseta 

(Dicks.) R.H.Zander) 

var. papillosum (Lindb.) G.Roth (P. cuspidatum 

subsp. papillosum (Lindb.) J.Guerra & Ros, Tortula 

acaulon var. papillosa (Lindb.) R.H.Zander, Tortula 

atherodes var. papillosa R.H.Zander) 


var. piliferum (Hedw.) Hook. & Taylor (Tortula 

acaulon var. pilifera (Hedw.) R.H.Zander Tortula 

atherodes var. pilifera (Hedw.) R.H.Zander) 

var. retortifolium J.Guerra & Ros 

var. schreberianum (Dicks.) Brid. (Tortula acaulon var. 

schreberiana (Dicks.) R.H.Zander, Tortula atherodes 

var. schreberiana (Dicks.) R.H.Zander) 

2 vlassovii Laz. (Microbryum vlassovii (Laz.) 

R.H.Zander) 

112 Protobryum J.Guerra & M.J.Cano 

1 bryoides (Dicks.) J.Guerra & M.J.Cano (Pottia 

bryoides (Dicks.) Mitt., Tortula protobryoides 

R.H.Zander) 

113 Pseudocrossidium R.S.Williams 

1 hornschuchianum (Schultz) R.H.Zander (Barbula 

hornschuchiana Schultz) 

2 obtusulum (Lindb.) H.A.Crum & L.E.Anderson 

(P. revolutum var. obtusulum (Lindb.) B.C.Tan, 

R.H.Zander & Taylor,Terry) [196] 

3 replicatum (Taylor) R.H.Zander [197] 

4 revolutum (Brid.) R.H.Zander (Barbula revoluta Brid.) 

114 Pterygoneurum Jur., nom. cons. 

1 compactum M.J.Cano, J.Guerra & Ros [198] 

2 crossidioides W.Frey, Herrnst. & Kürschner [199] 

3 kozlovii Laz. 

4 lamellatum (Lindb.) Jur. 

5 ovatum (Hedw.) Dixon (P. ovatum var. incanum Jur.) 

6 papillosum Oesau [200] 

7 sampaianum (Guim.) Guim. 

8 squamosum Segarra & Kürschner [201] 

9 subsessile (Brid.) Jur. 

115 Stegonia Venturi 

1 latifolia (Schwägr.) Venturi ex Broth. (S. latifolia var. 

latifolia, S. latifolia var. pilifera (Brid.) Broth.) 

116 Syntrichia Brid. [202] 

1 bogotensis (Hampe) R.H.Zander [203] 

2 calcicola J.J.Amann (S. densa (Velen.) J.-P.Frahm, 

Tortula calcicolens W.A.Kramer, Tortula densa (Velen.) 

J.-P.Frahm, Tortula ruralis var. calcicola (J.J.Amann) 

Barkman, Tortula ruralis var. densa Velen.) [204] 

3 caninervis Mitt. (Tortula caninervis (Mitt.) Broth.) [205] 

var. abranchesii (Luisier) R.H.Zander (S. abranchesii 

(Luisier) Ochyra, Tortula abranchesii Luisier) [206] 

var. astrakhanica Ignatov, Ignatova & Suragina [207] 

var. caninervis 

var. gypsophila (J.J.Amann ex G.Roth) Ochyra (S. 

caninervis var. spuria (J.J.Amann) R.H.Zander, S. 

ruralis var. gypsophila (J.J.Amann ex G.Roth) 

J.J.Amann, S. ruralis var. spuria (J.J.Amann) Podp., 

Tortula caninervis subsp. spuria (J.J.Amann) 

W.A.Kramer, Tortula caninervis subsp. spuria var. 

gypsophila (J.J.Amann ex G.Roth) W.A.Kramer, 

Tortula ruralis var. gypsophila J.J.Amann ex G.Roth, 

Tortula spuria J.J.Amann) 

4 echinata (Schiffn.) Herrnst. & Ben-Sasson (S. princeps 

subsp. echinata (Schiffn.) Podp., S. princeps var.


echinata (Schiffn.) R.H.Zander, Tortula echinata 

Schiffn.) [202] 

5 fragilis (Taylor) Ochyra (Barbula alpina var. inermis 

Milde, S. mutica Giacom., Tortula fragilis Taylor) 

6 glabra J.-P.Frahm & M.T.Gallego [208] 

7 handelii (Schiffn.) S.Agnew & Vondr. (S. montana 

subsp. handelii (Schiffn.) Podp., Tortula handelii 

Schiffn., Tortula intermedia subsp. handelii (Schiffn.) 

Wijk & Margad.) 

8 laevipila Brid. (S. laevipila var. laevipilaeformis (De 

Not.) J.J.Amann, S. pagorum (Milde) J.J.Amann, 

Tortula laevipila (Brid.) Schwägr., Tortula laevipila 

var. meridionalis (Schimp.) Wijk & Margad., Tortula 

laevipila var. notarisii Barkman, Tortula laevipila var. 

wachteri Barkman, Tortula laevipilaeformis De Not., 

Tortula pagorum (Milde) De Not., Tortula 

saccardoana De Not.) [209] 

9 latifolia (Bruch ex Hartm.) Huebener (Tortula latifolia 

Bruch ex Hartm.) 

10 minor (Bizot) M.T.Gallego, J.Guerra, M.J.Cano, Ros 

&Sánchez-Moya [210] 

11 montana Nees (S. intermedia Brid., Tortula crinita (De 

Not.) De Not., Tortula intermedia (Brid.) Berk., hom. 

illeg. [211]) 

var. calva (Durieu & Sagot ex Bruch & Schimp.) 

J.J.Amann (Tortula crinita var. calva (Durieu & Sagot 

ex Bruch & Schimp.) Nebel & Heinrichs, Tortula 

ruralis var. calva (Durieu & Sagot ex Bruch & 

Schimp.) C.Hartm.) 

var. montana 

12 norvegica F.Weber (Tortula norvegica (F.Weber) 

Lindb.) 

13 papillosa (Wilson) Jur. (Tortula papillosa var. 

meridionalis Warnst. [212], Tortula papillosa 

Wilson) 

14 papillosissima (Copp.) Loeske (S. ruralis var. hirsuta 

(Venturi) Podp., Tortula hirsuta (Venturi) Laz., 

Tortula papillosissima (Copp.) Broth., Tortula ruralis 

subsp. hirsuta (Venturi) W.A.Kramer, Tortula ruralis 

var. hirsuta (Venturi) Paris) [202] 

15 princeps (De Not.) Mitt. (Tortula princeps De Not.) 

[213] 

16 rigescens (Broth. & Geh.) Ochyra (Tortula rigescens 

Broth. & Geh.) [202] 

17 ruralis (Hedw.) F.Weber & D.Mohr (Tortula ruralis 

(Hedw.) P.Gaertn., B.Mey. & Scherb.) 

var. ruraliformis (Besch.) Delogne (S. ruraliformis 

(Besch.) Cardot, S. ruralis var. arenicola J.J.Amann, 

Tortula ruraliformis (Besch.) Ingham, Tortula ruralis 

var. ruraliformis (Besch.) De Wild.) [214] 

var. ruralis (S. ruralis var. glacialis J.J.Amann) 

18 sinensis (Müll.Hal.) Ochyra (Tortula sinensis 

(Müll.Hal.) Broth.) 

19 subpapillosissima (Bizot & R.B.Pierrot ex 

W.A.Kramer) M.T.Gallego & J.Guerra (Tortula 

papillosissima var. submamillosa (W.A.Kramer) 

Heinrichs & Caspari, Tortula ruraliformis var. 

subpapillosissima Bizot & R.B.Pierrot ex 

W.A.Kramer, Tortula ruralis var. submamillosa 

W.A.Kramer) [215] 

20 virescens (De Not.) Ochyra (Tortula virescens (De 

Not.) De Not.) 

117 Tortula Hedw., nom. cons. (Desmatodon Brid., Pottia 

Ehrh. ex Fürnr.) [216] [217] 

1 amplexa (Lesq.) Steere (Syntrichia amplexa (Lesq.) 

R.H.Zander) 

2 ampliretis Crundw. & D.G.Long [218] 

3 atrovirens (Sm.) Lindb. 

4 bogosica (Müll.Hal.) R.H.Zander (Desmatodon 

bogosicus Müll.Hal.) 

5 bolanderi (Lesq. & James) M.Howe (Syntrichia 

bolanderi (Lesq. & James) R.H.Zander) [219] 

6 brevissima Schiffn. [221] 

7 canescens Mont. 

8 cernua (Huebener) Lindb. (Desmatodon cernuus 

(Huebener) Bruch & Schimp.) 

9 cuneifolia (Dicks.) Turner 

10 freibergii Dixon & Loeske 

11 guepinii (Bruch & Schimp.) Broth. (Desmatodon 

guepinii Bruch & Schimp.) 

12 hoppeana (Schultz) Ochyra (Desmatodon latifolius 

(Hedw.) Brid., T. eucalyptrata Lindb., T. euryphylla 

R.H.Zander) [222] 

13 inermis (Brid.) Mont. (Syntrichia inermis (Brid.) 

Bruch) [219] 

14 israelis Bizot & F.Bilewsky (T. baetica (Casas & 

R.Oliva) J.Guerra & Ros, T. muralis var. baetica 

Casas & R.Oliva) [220] 

15 lanceola R.H.Zander (Pottia lanceolata (Hedw.) 

Müll.Hal.) 

16 laureri (Schultz) Lindb. (Desmatodon laureri (Schultz) 


17 leucostoma (R.Br.) Hook. & Grev. (Desmatodon 

leucostoma (R.Br.) Berggr.) 

18 lingulata Lindb. 

19 marginata (Bruch & Schimp.) Spruce (Desmatodon 

meridionalis Luisier) [223] 

20 modica R.H.Zander (Pottia intermedia (Turner) 

Fürnr.) 

21 mucronifolia Schwägr. 

22 muralis Hedw. 

23 obtusifolia (Schwägr.) Mathieu 

24 pallida (Lindb.) R.H.Zander (Pottia pallida Lindb., 

P. cuneifolia Solms ex Schimp., T. zoddae 

R.H.Zander) [224] 

25 randii (Kenn.) R.H.Zander (Desmatodon oxneri Laz., 

Desmatodon randii (Kenn.) Laz.) 

26 revolvens (Schimp.) G.Roth 

27 rhodonia R.H.Zander (Desmatodon wilczekii Meyl.) 

[225] 

28 schimperi M.J.Cano, O.Werner & J.Guerra 

(T. angustata Lindb., hom. illeg., T. subulata var. 

angustata (Schimp.) Limpr.) [226]

29 solmsii (Schimp.) Limpr. (T. marginata subsp. limbata 

(Lindb.) Podp.) [227] 

30 subulata Hedw. (T. subulata var. graeffii Warnst., 

T. subulata var. subinermis (Bruch & Schimp.) Wilson) 

[226] 

31 systylia (Schimp.) Lindb. (Desmatodon systylius 

Schimp.) 

32 truncata (Hedw.) Mitt. (Pottia truncata (Hedw.) Bruch 

& Schimp.) 

33 ucrainica (Laz.) R.H.Zander (Desmatodon ucrainicus 

Laz.) 

34 vahliana (Schultz) Mont. 

35 viridifolia (Mitt.) Blockeel & A.J.E.Sm. (Pottia crinita 


36 wilsonii (Hook.) R.H.Zander (Pottia wilsonii (Hook.) 


118 Triquetrella Müll.Hal. 

1 arapilensis Luisier 

Splachnales (M.Fleisch.) Ochyra 

Splachnaceae Grev. & Arn. 

119 Aplodon R.Br. 

1 wormskioldii (Hornem.) R.Br. [32] 

120 Splachnum Hedw. 

1 ampullaceum Hedw. 

2 luteum Hedw. 

3 melanocaulon (Wahlenb.) Schwägr. 

4 pensylvanicum (Brid.) Grout ex H.A.Crum 

5 rubrum Hedw. 

6 sphaericum Hedw. 

7 vasculosum Hedw. 

121 Tayloria Hook. 

1 acuminata Hornsch. 

2 froelichiana (Hedw.) Mitt. ex Broth. 

3 hornschuchii (Grev. & Arn.) Broth. 

4 lingulata (Dicks.) Lindb. 

5 rudolphiana (Garov.) Bruch & Schimp. 

6 serrata (Hedw.) Bruch & Schimp. 

7 splachnoides (Schleich. ex Schwägr.) Hook. 

8 tenuis (Dicks.) Schimp. (T. serrata var. tenuis (Dicks.) 


122 Tetraplodon Bruch & Schimp. 

1 angustatus (Hedw.) Bruch & Schimp. 

2 blyttii Frisvoll 

3 mnioides (Hedw.) Bruch & Schimp. 

4 pallidus I.Hagen 

5 paradoxus (R.Br.) I.Hagen 

6 urceolatus (Hedw.) Bruch & Schimp. [228] 

123 Voitia Hornsch. 

1 hyperborea Grev. & Arn. 

2 nivalis Hornsch. 

Meesiaceae Schimp. 

124 Amblyodon P.Beauv., nom. cons. 

1 dealbatus (Hedw.) P.Beauv. 

125 Leptobryum (Bruch & Schimp.) Wilson 

1 pyriforme (Hedw.) Wilson 

126 Meesia Hedw., nom. cons. 

1 hexasticha (Funck) Bruch 


2 longiseta Hedw. 

3 triquetra (L. ex Jolycl.) A˚ ngstr. 

4 uliginosa Hedw. 

127 Paludella Brid. 

1 squarrosa (Hedw.) Brid. 

Orthotrichales Dixon 

Orthotrichaceae Arn. 

128 Orthotrichum Hedw. 

Subgenus Orthotrichum [229] 

1 anomalum Hedw. 

2 cupulatum Hoffm. ex Brid. 

var. bistratosum Schiffn. 

var. cupulatum 

var. fuscum (Venturi) Boulay (O. limprichtii I.Hagen) 

[230] 

var. riparium Huebener (O. cupulatum var. nudum 

(Dicks.) Braithw.) 

3 pellucidum Lindb. 

4 urnigerum Myrin 

Subgenus Pulchella (Schimp.) Vitt 

5 alpestre Bruch & Schimp. 

6 casasianum F.Lara, Garilleti & Mazimpaka [231] 

7 consimile Mitt. 

8 crenulatum Mitt. [231] [232] 

9 diaphanum Schrad. ex Brid. 

10 handiense F.Lara, Garilleti & Mazimpaka [233] 

11 hispanicum F.Lara, Garilleti & Mazimpaka [234] 

12 macrocephalum F.Lara, Garilleti & Mazimpaka [235] 

13 microcarpum De Not. 

14 pallens Bruch ex Brid. (O. paradoxum Gronvall) 

15 patens Bruch ex Brid. 

16 philibertii Venturi 

17 pulchellum Brunt. 

18 pumilum Sw. ex anon. [106] 

19 rivulare Turner 

20 rogeri Brid. 

21 scanicum Gronvall (O. lewinskyae F.Lara, Garilleti & 

Mazimpaka) [236] 

22 schimperi Hammar [237] 

23 sprucei Mont. 

24 stellatum Brid. 

25 stramineum Hornsch. ex Brid. (O. rogeri var. defluens 

(Venturi) Venturi) [238] 

26 tenellum Bruch ex Brid. (O. australe Jur.) [239] 

27 vittii F.Lara, Garilleti & Mazimpaka [240] 

Subgenus Callistoma (Z.Iwats. & Sharp) Lewinsky 

28 callistomum Fisch.-Oost. ex Bruch & Schimp. 

Subgenus Orthophyllum Delogne 

29 gymnostomum Bruch ex Brid. 

30 obtusifolium Brid. 

Subgenus Phaneroporum Delogne 

31 laevigatum J.E.Zetterst. (O. arcangelianum Massari) 

[241] 

32 rupestre Schleich. ex Schwägr. 

33 shawii Wilson [242] 

Subgenus Gymnoporus (Braithw.) Limpr. 

34 acuminatum H.Philib.


35 affine Schrad. ex Brid. (O. fastigiatum Bruch ex Brid.) 

36 ibericum F.Lara & Mazimpaka [243] 

37 lyellii Hook. & Taylor 

38 pylaisii Brid. 

39 sordidum Sull. & Lesq. (O. caucasicum Venturi) [244] 

40 speciosum Nees (O. elegans auct. eur. non Schwägr.) 

[245] 

var. brevisetum F.Lara, Garilleti & Mazimpaka [246] 

var. speciosum 

41 striatum Hedw. 

42 tortidontium F.Lara, Garilleti & Mazimpaka [247] 

43 vladikavkanum Venturi [248] 

129 Ulota D.Mohr 

1 bruchii Hornsch. ex Brid. (U. crispa var. norvegica 

(Gronvall) A.J.E.Sm. & M.O.Hill) 

2 calvescens Wilson 

3 coarctata (P.Beauv.) Hammar 

4 crispa (Hedw.) Brid. (U. intermedia Schimp.) 

5 curvifolia (Wahlenb.) Lilj. 

6 drummondii (Hook. & Grev.) Brid. 

7 hutchinsiae (Sm.) Hammar 

8 macrospora E.Bauer & Warnst. [249] 

9 phyllantha Brid. 

10 rehmannii Jur. 

130 Zygodon Hook. & Taylor 

1 conoideus (Dicks.) Hook. & Taylor 

var. conoideus 

var. lingulatus S.R.Edwards 

2 dentatus (Limpr.) Kartt. (Z. viridissimus var. dentatus 

Limpr.) 

3 forsteri (Dicks.) Mitt. (Codonoblepharon forsteri 

(Dicks.) Goffinet, Zygodon madeirensis Dixon & 

Luisier) [250] 

4 gracilis Wilson 

5 rupestris Schimp. ex Lorentz (Z. baumgartneri Malta, 

Z. viridissimus var. rupestris C.Hartm., Z. vulgaris 

Nyholm) 

6 sibiricus Ignatov, Ignatova, Z.Iwats. & B.C.Tan [251] 

7 stirtonii Schimp. ex Stirt. [252] 

8 viridissimus (Dicks.) Brid. 

Hedwigiales Ochyra 

Hedwigiaceae Schimp. 

131 Braunia Bruch & Schimp. 

1 alopecura (Brid.) Limpr. 

132 Hedwigia P.Beauv., nom. cons. 

1 ciliata (Hedw.) P.Beauv. [253] 

var. ciliata 

var. leucophaea Bruch & Schimp. [254] 

2 integrifolia P.Beauv. (Hedwigidium integrifolium 

(P.Beauv.) Dixon) 

3 stellata Hedenäs [255] 

Bryales Limpr. 

Catoscopiaceae Boulay ex Broth. 

133 Catoscopium Brid. 

1 nigritum (Hedw.) Brid. 

Bartramiaceae Schwägr. 

134 Anacolia Schimp., nom. cons. 

1 laevisphaera (Taylor) Flowers [256] 

2 menziesii (Turner) Paris [257] 

3 webbii (Mont.) Schimp. 

135 Bartramia Hedw., nom. cons. 

section Bartramia 

1 halleriana Hedw. 

2 pomiformis Hedw. [258] 

section Pyridium Müll.Hal. [259] 

3 breviseta Lindb. (B. ithyphylla var. breviseta (Lindb.) 

Kindb.) [260] 

4 ithyphylla Brid. [261] 

5 subulata Bruch & Schimp. 

section Strictidium Müll.Hal. 

6 stricta Brid. 

136 Breutelia (Bruch & Schimp.) Schimp. 

1 azorica (Mitt.) Cardot 

2 chrysocoma (Hedw.) Lindb. 

137 Conostomum Sw. ex F.Weber & D.Mohr 

1 tetragonum (Hedw.) Lindb. 

138 Philonotis Brid. 

section Bartramidula (Bruch & Schimp.) Mitt. 

1 cernua (Wilson) D.G.Griffin & W.R.Buck 

section Philonotula (Schimp.) A.Jaeger 

2 rigida Brid. 

section Homomorphae (Kindb.) Ochyra 

3 arnellii Husn. (P. capillaris auct. non Lindb.) 

4 hastata (Duby) Wijk & Margad. 

5 marchica (Hedw.) Brid. (P. marchica var. laxa 

(Limpr.) Loeske & Warnst. [262]) 

6 uncinata (Schwägr.) Brid. [263] 

section Philonotis 

7 caespitosa Jur. 

8 calcarea (Bruch & Schimp.) Schimp. 

9 fontana (Hedw.) Brid. 

10 seriata Mitt. 

11 tomentella Molendo (P. fontana var. pumila (Turner) 

Brid.) 

139 Plagiopus Brid. 

1 oederianus (Sw.) H.A.Crum & L.E.Anderson 

(P. oederi (Brid.) Limpr.) 

var. alpinus (Schwägr.) Ochyra 

var. oederianus 

Bryaceae Schwägr. [264] 

140 Anomobryum Schimp. 

1 concinnatum (Spruce) Lindb. (A. filiforme subsp. 

concinnatum (Spruce) J.J.Amann, A. julaceum var. 

concinnatum (Spruce) J.E.Zetterst., A. leptostomoides 

Schimp., Bryum concinnatum Spruce, Bryum filiforme 

var. concinnatum (Spruce) Boulay, Bryum julaceum 

var. concinnatum (Spruce) Wilson) [265] 

2 julaceum (Schrad. ex P.Gaertn. et al.) Schimp. (A. 

filiforme (Dicks.) Husn., hom. illeg., A. filiforme var. 

juliforme (Solms) Husn., A. juliforme Solms, Bryum 

filiforme Dicks., Bryum julaceum Schrad. ex P.Gaertn. 

et al., Bryum juliforme (Solms) Schimp., Bryum 

leptostomum Schimp.) 

3 lusitanicum (I.Hagen ex Luisier) Thér. [266]

141 Brachymenium Schwägr. 

1 commutatum (Müll.Hal.) A.Jaeger [267] 

2 notarisii (Mitt.) A.J.Shaw (Haplodontium notarisii 

(Mitt.) Broth.) [268] 

3 paradoxum (Herzog) A.J.Shaw (Mielichhoferia 

paradoxa Herzog) [269] 

4 philonotula Broth. [270] 

142 Bryum Hedw. (Imbribryum N.Pedersen) [271] [272] 

[273] [274] [275] 

1 algovicum Sendtn. ex Müll.Hal. [276] 

var. algovicum (B. algovicum var. compactum 

(Hornsch.) Düll, B. compactum (Hornsch.) Kindb.) 

var. rutheanum (Warnst.) Crundw. 

2 alpinum Huds. ex With. (Imbribryum alpinum (Huds. 

ex With.) N.Pedersen) 

3 apiculatum Schwägr.(B. cruegeri Hampe, B. nitens Hook., 

Pohlia apiculata (Schwägr.) H.A.Crum & L.E.Anderson, 

Pohlia cruegeri (Hampe) A.L.Andrews) [277] 

4 archangelicum Bruch & Schimp. (B. amblyodon 

Müll.Hal., B. curvatum Kaurin & Arnell, B. 

imbricatum auct. non? (Schwägr.) Bruch & Schimp., 

B. inclinatum (Brid.) Turton, hom. illeg. non (Hedw.) 

Dicks., B. stenotrichum Müll.Hal.) [278] 

5 arcticum (R.Br.) Bruch & Schimp. (B. bryoides (R.Br.) 

Wijk & Margad., B. lawersianum H.Philib., B. 

purpurascens (R.Br.) Bruch & Schimp.) [278] 

6 argenteum Hedw. (Anomobryum lanatum (P.Beauv.) 

J.R.Spence & H.P.Ramsay, B. argenteum subsp. 

veronense (De Not.) J.J.Amann, B. argenteum var. 

lanatum (P.Beauv.) Hampe [280], B. argenteum var. 

veronense (De Not.) Molendo, B. lanatum (P.Beauv.) 

Brid., B. veronense De Not.) [279] 

7 blindii Bruch & Schimp. [281] 

8 bornholmense Wink. & R.Ruthe [282] 

9 caespiticium Hedw. (B. badium (Brid.) Schimp., B. 

comense Schimp.) [283] 

10 calophyllum R.Br. (B. acutiforme Limpr., B. 

axel-blyttii Kaurin ex H.Philib.) [278] 

11 canariense Brid. (B. provinciale H.Philib.) [284] 

12 capillare Hedw. (B. platyloma Schwägr.,B. rufifolium 

(Dixon) Demaret & R.Wilczek, B. validicostatum 

Cardot & Dixon, Rosulabryum capillare (Hedw.) 

J.R.Spence) [278] 

13 caucasicum (Schimp. ex Broth.) C.J.Cox & Hedd. 

(Mielichhoferia caucasia Schimp. ex Broth., 

Mielichhoferia himalayana Mitt. non Bryum 

himalayanum (Mitt.) Müll.Hal.) [285] 

14 cellulare Hook. (Brachymenium cellulare (Hook.) A.Jaeger, 

B. splachnoides (Harv.) Müll.Hal. non Dicks.) [269] 

15 creberrimum Taylor (B. affine F.W.Schultz non 

J.F.Gmel. ex Broth., B. cuspidatum (Bruch & Schimp.) 

Schimp., B. lisae De Not.) 

16 cryophilum Martensson (B. cyclophyllum var. 

obtusifolium Lindb., B. obtusifolium Lindb., 

hom. illeg.) 

17 cyclophyllum (Schwägr.) Bruch & Schimp. 

(B. tortifolium Funck ex Brid.) 


18 demaretianum Arts [286] 

19 dichotomum Hedw. (B. balticum Nyholm & Hedenäs, 

B. barnesii J.B.Wood ex Schimp., B. bicolor Dicks., 

B. dunense A.J.E.Sm. & H.Whitehouse, B. excurrens 

Lindb., B. versicolor A.Braun ex Bruch & Schimp.) [287] 

20 dixonii Cardot ex W.E.Nicholson (B. bicolor subsp. 

dixonii (Cardot ex W.E.Nicholson) Podp.) 

21 donianum Grev. (B. obovatum Mitt., B. pachyloma 

Cardot) 

22 dyffrynense Holyoak [287] 

23 elegans Nees (B. capillare var. elegans (Nees) Husn., B. 

capillare var. ferchelii (Funck ex Brid.) Bruch & 

Schimp., B. chibinense Schljakov, B. stirtonii Schimp., 

Rosulabryum elegans (Nees) Ochyra) [278] 

24 funckii Schwägr. (B. funkii auct. non Schwägr.) [278] 

25 gemmiferum R.Wilczek & Demaret 

26 gemmilucens R.Wilczek & Demaret 

27 gemmiparum De Not. (B. alpinum subsp. gemmiparum 

(De Not.) Kindb.) 

28 intermedium (Brid.) Blandow (B. nitidulum Lindb.) 

[278] 

29 klinggraeffii Schimp. 

30 knowltonii Barnes (B. lacustre (F.Weber & D.Mohr) 

Blandow non Brid.) 

31 kunzei Hornsch. (B. caespiticium subsp. kunzei 

(Hornsch.) Podp., B. caespiticium var. imbricatum 

Bruch & Schimp., B. caespiticium var. kunzei 

(Hornsch.) Braithw., nom. illeg.) [278] [283] [288] 

32 longisetum Blandow ex Schwägr. (B. inclinatum subsp. 

longisetum (Blandow ex Schwägr.) Podp.) 

33 marratii Hook.f. & Wilson 

34 mildeanum Jur. (B. alpinum var. mildeanum (Jur.) Podp.) 

35 miniatum Lesq. [289] 

36 minii Podp. [290] 

37 moravicum Podp. (B. flaccidum auct. non Brid., B. 

laevifilum Syed, B. subelegans auct. non Kindb., 

Rosulabryum laevifilum (Syed) Ochyra) [278] 

38 muehlenbeckii Bruch & Schimp. (B. alpinum var. 

brevifolium Myrin, Imbribryum muehlenbeckii (Bruch 

& Schimp.) N.Pedersen) [291] 

39 neodamense Itzigs. (B. cavifolium Wilson, nom. nud. in 

synon., B. crispulum Hampe ex Müll.Hal., B. 

neodamense var. ovatum (Lange & C.E.O.Jensen) 

Lindb. & Arnell, B. pseudotriquetrum subsp. 

neodamense (Itzigs.) J.J.Amann, B. pseudotriquetrum 

var. cavifolium Schimp., B. subneodamense Kindb.) 

[292] 

40 oblongum Lindb. (B. blindii subsp. oblongum (Lindb.) 

Kindb., B. blindii var. oblongum (Lindb.) Mönk.) [281] 

41 pallens Sw. ex anon. (B. aeneum Blytt ex Bruch & 

Schimp. [293], B. oeneum Blytt ex Bruch & Schimp. 

emend. Wijk et al., B. rutilans auct. eur. non Brid., B. 

subelegans Kindb.) [106] [278] 

42 pallescens Schleich. ex Schwägr. (B. lonchocaulon 

Müll.Hal. [294], B. obconicum Hornsch. ex Bruch & 

Schimp.)


43 pseudotriquetrum (Hedw.) P.Gaertn. et al. (B. 

castaneum I.Hagen, B. ventricosum Dicks., nom. illeg.) 

var. bimum (Schreb.) Lilj. (B. bimum (Schreb.) Turner, 

B. pseudotriquetrum subsp. bimum (Schreb.) Hartm.) 

[295] 

var. pseudotriquetrum 

44 radiculosum Brid. 

45 riparium I.Hagen 

46 rubens Mitt. 

47 ruderale Crundw. & Nyholm 

48 salinum I.Hagen ex Limpr. [296] 

49 sauteri Bruch & Schimp. 

50 schleicheri DC. (B. schleicheri var. latifolium 

(Schwägr.) Schimp., B. turbinatum subsp. schleicheri 

(Schwägr.) Kindb., B. turbinatum var. latifolium 

(Schwägr.) Bruch & Schimp.) [297] 

51 subapiculatum Hampe (B. erythrocarpum auct. partim, 

B. microerythrocarpum Müll.Hal. & Kindb. ex 

Macoun) 

52 tenuisetum Limpr. 

53 torquescens Bruch & Schimp. (B. capillare subsp. 

icodense (H.Winter) Podp., B. capillare subsp. 

torquescens (Bruch & Schimp.) Kindb., B. capillare 

var. torquescens (Bruch & Schimp.) Husn., B. icodense 

H.Winter, B. obconicum auct. non Hornsch. ex Bruch 

& Schimp., Rosulabryum torquescens (Bruch & 

Schimp.) J.R.Spence) [278] [298] 

54 turbinatum (Hedw.) Turner (B. syriacum Lorentz) 

55 uliginosum (Brid.) Bruch & Schimp. (B. cernuum 

(Hedw.) Bruch & Schimp.) 

56 valparaisense Thér. (B. pyriferum Crundw. & 

H.Whitehouse) [299] 

57 violaceum Crundw. & Nyholm 

58 warneum (Röhl.) Brid. (B. mamillatum Lindb., B. 

oelandicum H.Philib.) [278] 

59 weigelii Spreng. (B. duvalii Voit) 

60 wrightii Sull. & Lesq. (B. globosum Lindb., B. 

mamillatum var. globosum (Lindb.) Berggr.) 

143 Plagiobryum Lindb. 

1 demissum (Hook.) Lindb. (Bryum demissum Hook.) 

2 zieri (Hedw.) Lindb. (Bryum zieri Hedw.) [300] 

144 Rhodobryum (Schimp.) Limpr., nom. cons. 

1 ontariense (Kindb.) Kindb. (Bryum ontariense Kindb., 

R. spathulatum auct. non (Hornsch.) Pócs) [301] 

2 roseum (Hedw.) Limpr. (Bryum roseum (Hedw.) 

P.Gaertn. et al.) 

Mielichhoferiaceae Schimp. (Mniaceae subfam. 

Pohlioideae auct., nom. nud.) [302] 

145 Epipterygium Lindb. 

1 tozeri (Grev.) Lindb. 

146 Mielichhoferia Nees & Hornsch. 

1 elongata (Hoppe & Hornsch. ex Hook.) Hornsch. 

(M. mielichhoferiana var. elongata (Hoppe & Hornsch. 

ex Hook.) Wijk & Margad.) [303] 

2 mielichhoferiana (Funck) Loeske (M. mielichhoferi 

(Hook.) Wijk & Margad., nom. inval.) 

147 Pohlia Hedw. [304] [305] 

section Pohlia 

1 bolanderi (Lesq.) Broth. [306] 

2 cruda (Hedw.) Lindb. 

3 crudoides (Sull. & Lesq.) Broth. 

4 elongata Hedw. 

var. acuminata (Hornsch.) Huebener (P. acuminata 

Hornsch., P. ambigua (Limpr.) Broth.) [288] [307] 

var. elongata 

var. greenii (Brid.) A.J.Shaw (P. elongata var. 

polymorpha (Hornsch.) Nyholm [288] 

5 longicolla (Hedw.) Lindb. [32] 

6 nutans (Hedw.) Lindb. [308] 

subsp. nutans 

subsp. schimperi (Müll.Hal.) Nyholm (P. nutans var. 

purpurascens Latzel, P. schimperi (Müll.Hal.) 

A.L.Andrews) [308] 

7 obtusifolia (Vill. ex Brid.) L.F.Koch 

8 saprophila (Müll.Hal.) Broth. [309] 

9 sphagnicola (Bruch & Schimp.) Broth. 

section Cacodon Lindb. ex Broth. 

10 andalusica (Höhn.) Broth. 

11 andrewsii A.J.Shaw 

12 annotina (Hedw.) Lindb. (P. grandiflora H.Lindb.) 

13 bulbifera (Warnst.) Warnst. 

14 camptotrachela (Renauld & Cardot) Broth. 

15 drummondii (Müll.Hal.) A.L.Andrews 

16 erecta Lindb. (P. defecta (Sanio) A.L.Andrews) 

17 filum (Schimp.) Martensson (P. schleicheri H.A.Crum) 

18 flexuosa Hook. (P. muyldermansii R.Wilczek & 

Demaret) [310] 

var. flexuosa 

var. pseudomuyldermansii (Arts, Nordhorn-Richter & 

A.J.E.Sm.) A.J.E.Sm. (P. muyldermansii var. 

pseudomuyldermansii Arts, Nordhorn-Richter & 

A.J.E.Sm.) [311] 

19 lescuriana (Sull.) Ochi (Mniobryum pulchellum 

(Hedw.) Loeske) 

20 ludwigii (Spreng. ex Schwägr.) Broth. 

21 lutescens (Limpr.) H.Lindb. 

22 proligera (Kindb.) Lindb. ex Broth. 

23 scotica Crundw. 

24 tundrae A.J.Shaw [312] 

section Apalodictyon (Müll.Hal.) Ochyra (section 

Mniobryum Nyholm, nom. inval.) 

25 atropurpurea (Wahlenb.) H.Lindb. 

26 melanodon (Brid.) A.J.Shaw (Mniobryum delicatulum 

(Hedw.) Dixon) 

27 vexans (Limpr.) H.Lindb. 

28 wahlenbergii (F.Weber & D.Mohr) A.L.Andrews 

(Mniobryum wahlenbergii (F.Weber & D.Mohr) Jenn.) 

var. calcarea (Warnst.) E.F.Warb. 

var. glacialis (Brid.) E.F.Warb. 

var. wahlenbergii 

148 Schizymenium Harv. 

1 pontevedrense (Luisier) Sérgio, Casas, Cros & Brugués 

Mniaceae Schwägr. [302] 

149 Mnium Hedw., nom. cons.

section Mnium 

1 hornum Hedw. 

section Spinosa (Kindb.) T.J.Kop. 

2 spinosum (Voit) Schwägr. 

3 spinulosum Bruch & Schimp. 

4 thomsonii Schimp. 

section Laevinervia P.C.Chen ex X.J.Li & M.Zang 

5 lycopodioides Schwägr. (M. ambiguum H.Müll.) [313] 

6 marginatum (Dicks.) P.Beauv. 

var. dioicum (H.Müll.) Crundw. (M. marginatum var. 

riparium (Mitt.) Husn.) 

var. marginatum 

section Stellariformia (Kindb.) T.J.Kop. 

7 blyttii Bruch & Schimp. 

8 heterophyllum (Hook.) Schwägr. 

9 stellare Hedw. 

150 Trachycystis T.J.Kop. 

1 ussuriensis (Maack & Regel) T.J.Kop. (T. immarginata 

(Broth.) Laz.) [314] 

Cinclidiaceae Kindb. (Mniaceae tribus Cinclidieae 

T.J.Kop.) [302] 

151 Cinclidium Sw. 

1 arcticum (Bruch & Schimp.) Schimp. 

2 latifolium Lindb. 

3 stygium Sw. 

4 subrotundum Lindb. 

152 Cyrtomnium Holmen 

1 hymenophylloides (Huebener) T.J.Kop. 

2 hymenophyllum (Bruch & Schimp.) Holmen 

153 Rhizomnium (Broth.) T.J.Kop. 

1 andrewsianum (Steere) T.J.Kop. 

2 gracile T.J.Kop. 

3 magnifolium (Horik.) T.J.Kop. 

4 pseudopunctatum (Bruch & Schimp.) T.J.Kop. 

5 punctatum (Hedw.) T.J.Kop. 

Plagiomniaceae T.J.Kop. (Mniaceae tribus Plagiomnieae 

T.J.Kop.) [302] 

154 Plagiomnium T.J.Kop. 

section Plagiomnium 

1 cuspidatum (Hedw.) T.J.Kop. 

2 drummondii (Bruch & Schimp.) T.J.Kop. 

section Rosulata (Kindb.) T.J.Kop. 

3 affine (Blandow ex Funck) T.J.Kop. 

4 curvatulum (Lindb.) Schljakov (P. medium subsp. 

curvatulum (Lindb.) T.J.Kop.) [315] 

5 elatum (Bruch & Schimp.) T.J.Kop. 

6 ellipticum (Brid.) T.J.Kop. 

7 medium (Bruch & Schimp.) T.J.Kop. 

section Undulata (Kindb.) T.J.Kop. 

8 confertidens (Lindb. & Arnell) T.J.Kop. 

9 undulatum (Hedw.) T.J.Kop. 

var. madeirense T.J.Kop. & Sérgio [316] 

var. undulatum 

section Rostrata (Kindb.) T.J.Kop. 

10 rostratum (Schrad.) T.J.Kop. 

155 Pseudobryum (Kindb.) T.J.Kop. 

1 cinclidioides (Huebener) T.J.Kop. 


Aulacomniaceae Schimp. 

156 Aulacomnium Schwägr., nom. cons. 

1 androgynum (Hedw.) Schwägr. 

2 palustre (Hedw.) Schwägr. 

3 turgidum (Wahlenb.) Schwägr. 

Orthodontiaceae (Broth.) Goffinet 

157 Orthodontium Schwägr. 

1 gracile (Wilson) Schwägr. ex Bruch & Schimp. 

2 lineare Schwägr. (O. australe Hook.f. & Wilson [317]) 

3 pellucens (Hook.) Bruch & Schimp. 

Rhizogoniales (M.Fleisch.) Goffinet & W.R.Buck 

Rhizogoniaceae Broth. 

158 Leptotheca Schwägr. 

1 gaudichaudii Schwägr. [318] 

Calomniaceae Kindb. 

159 Calomnion Hook.f. & Wilson 

1 complanatum (Hook.f. & Wilson) Lindb. [319] 

Hookeriales M.Fleisch. 

Hypopterygiaceae Mitt. 

160 Hypopterygium Brid. 

1 tamarisci (Sw.) Brid. ex Müll.Hal. (H. muelleri 

Hampe) [320] 

Daltoniaceae Schimp. 

161 Achrophyllum Vitt & Crosby 

1 dentatum (Hook.f. & Wilson) Vitt & Crosby [321] 

162 Calyptrochaeta Desv. 

1 apiculata (Hook.f. & Wilson) Vitt (Eriopus apiculatus 

(Hook.f. & Wilson) Mitt.) 

163 Daltonia Hook. & Taylor, nom. cons. 

1 splachnoides (Sm.) Hook. & Taylor 

2 stenophylla Mitt. [322] 

164 Distichophyllum Dozy & Molk. 

1 carinatum Dixon & W.E.Nicholson 

Hookeriaceae Schimp. 

165 Hookeria Sm., nom. cons. 

1 lucens (Hedw.) Sm. 

Leucomiaceae Broth. 

166 Tetrastichium (Mitt.) Cardot 

1 fontanum (Mitt.) Cardot 

2 virens (Cardot) S.P.Churchill (Lepidopilum virens 

Cardot) 

Pilotrichaceae Kindb. 

167 Cyclodictyon Mitt. 

1 laetevirens (Hook. & Taylor) Mitt. [323] 

Hypnales (M.Fleisch.) W.R.Buck & Vitt 

Fontinalaceae Schimp. 

168 Dichelyma Myrin 

1 capillaceum (L. ex Dicks.) Myrin 

2 falcatum (Hedw.) Myrin 

169 Fontinalis Hedw. 

1 antipyretica Hedw. [324] 

subsp. antipyretica 

subsp. bryhnii (Limpr.) Podp. (F. bryhnii Limpr. ex 

I.Hagen) [325] 

subsp. gracilis (Lindb.) Kindb. (F. antipyretica var. 

gracilis (Lindb.) Schimp.)


subsp. kindbergii (Renauld & Cardot) Cardot (F. 

kindbergii Renauld & Cardot) 

2 dalecarlica Schimp. 

3 dichelymoides Lindb. [326] 

4 hypnoides C.Hartm. 

var. duriaei (Schimp.) Kindb. 

var. hypnoides 

5 squamosa Hedw. 

var. curnowii Cardot 

var. dixonii (Cardot) A.J.E.Sm. 

var. squamosa 

Climaciaceae Kindb. 

170 Climacium F.Weber & D.Mohr 

1 dendroides (Hedw.) F.Weber & D.Mohr 

Amblystegiaceae Kindb. [327] 

171 Amblystegium Schimp. [327] 

1 confervoides (Brid.) Schimp. (Platydictya confervoides 

(Brid.) H.A.Crum, Serpoleskea confervoides (Brid.) 

Loeske) 

2 radicale (P.Beauv.) Schimp. (A. saxatile Schimp., 

Campylium radicale (P.Beauv.) Grout) 

3 serpens (Hedw.) Schimp. (A. juratzkanum Schimp.) 

4 subtile (Hedw.) Schimp. (Platydictya subtilis (Hedw.) 

H.A.Crum, Serpoleskea subtilis (Hedw.) Loeske) 

172 Anacamptodon Brid. 

1 splachnoides (Froel. ex Brid.) Brid. 

173 Campyliadelphus (Kindb.) R.S.Chopra 

1 chrysophyllus (Brid.) R.S.Chopra (Campylium 

chrysophyllum (Brid.) Lange) 

2 elodes (Lindb.) Kanda (Campylium elodes (Lindb.) 

Kindb.) 

174 Campylium (Sull.) Mitt. 

1 laxifolium Engelmark & Hedenäs [328] 

2 longicuspis (Lindb. & Arnell) Hedenäs 

3 protensum (Brid.) Kindb. (C. stellatum subsp. 

protensum (Brid.) C.E.O.Jensen, C. stellatum var. 

protensum (Brid.) Bryhn) 

4 stellatum (Hedw.) Lange & C.E.O.Jensen 

(Campyliadelphus stellatus (Hedw.) Kanda) 

175 Conardia H.Rob. [327] 

1 compacta (Drumm. ex Müll.Hal.) H.Rob. 

(Amblystegium compactum (Drumm. ex Müll.Hal.) 

Austin, Rhynchostegiella compacta (Drumm. ex 

Müll.Hal.) Loeske) 

176 Cratoneuron (Sull.) Spruce 

1 curvicaule (Jur.) G.Roth (Callialaria curvicaulis (Jur.) 

Ochyra, Cratoneuron filicinum var. curvicaule (Jur.) 

Mönk.) 

2 filicinum (Hedw.) Spruce 

177 Drepanocladus (Müll.Hal.) G.Roth, nom. cons. 

1 aduncus (Hedw.) Warnst. (D. polycarpos (Blandow ex 

Voit) Warnst., D. simplicissimus Warnst., D. stagnatus 

Z˙ arnowiec) 

2 arcticus (R.S.Williams) Hedenäs (Campylium arcticum 

(R.S.Williams) Broth.) 

3 longifolius (Mitt.) Paris (D. capillifolius (Warnst.) 

Warnst.) 

4 polygamus (Schimp.) Hedenäs (Campyliadelphus 

polygamus (Schimp.) Kanda, Campylium polygamum 

(Schimp.) Lange & C.E.O.Jensen) 

5 sendtneri (Schimp. ex H.Müll.) Warnst. 

6 sordidus (Müll.Hal.) Hedenäs (D. tenuinervis T.J.Kop.) 

178 Hygroamblystegium Loeske, nom. cons. [327] [329] 

1 fluviatile (Hedw.) Loeske (Amblystegium fluviatile 

(Hedw.) Schimp.) 

2 humile (P.Beauv.) Vanderp., Goffinet & Hedenäs 

(Amblystegium humile (P.Beauv.) Crundw., 

Leptodictyum humile (P.Beauv.) Ochyra, 

Leptodictyum kochii (Schimp.) Warnst.) 

3 tenax (Hedw.) Jenn. (Amblystegium tenax (Hedw.) 

C.E.O.Jensen) 

4 varium (Hedw.) Mönk. (Amblystegium varium (Hedw.) 

Lindb., Orthotheciella varia (Hedw.) Ochyra) [330] 

179 Hygrohypnum Lindb. [327] [331] 

1 alpestre (Hedw.) Loeske (Ochyraea alpestris (Hedw.) 

Ignatov & Ignatova) 

2 alpinum (Lindb.) Loeske 

3 cochlearifolium (Venturi) Broth. (Ochyraea 

cochlearifolia (Venturi) Ignatov & Ignatova) 

4 duriusculum (De Not.) D.W.Jamieson (Hygrohypnella 

duriuscula (Turner ex Wilson) Ignatov & Ignatova, H. 

dilatatum (Wilson) Loeske) 

5 eugyrium (Schimp.) Broth. (Pseudohygrohypnum 

eugyrium (Schimp.) Kanda) 

6 luridum (Hedw.) Jenn. (Pictus scoticus C.C.Towns.) 

[332] 

7 molle (Hedw.) Loeske (H. molle var. schimperianum 

(Lorentz) Loeske) 

8 montanum (Lindb.) Broth. (Ochyraea montana 

(Lindb.) Ignatov & Ignatova) 

9 norvegicum (Schimp.) J.J.Amann (Ochyraea norvegica 

(Schimp.) Ignatov & Ignatova) 

10 ochraceum (Turner ex Wilson) Loeske (Hygrohypnella 

ochracea (Turner ex Wilson) Ignatov & Ignatova) 

11 polare (Lindb.) Loeske (Hygrohypnella polaris 

(Lindb.) Ignatov & Ignatova) 

12 smithii (Sw.) Broth. (Ochyraea smithii (Sw.) Ignatov 

& Ignatova) 

13 styriacum (Limpr.) Broth. 

14 subeugyrium (Renauld & Cardot) Broth. 

(Pseudohygrohypnum subeugyrium (Renauld & 

Cardot) Ignatov & Ignatova) 

180 Leptodictyum (Schimp.) Warnst. [333] 

1 riparium (Hedw.) Warnst. (Amblystegium riparium 


181 Ochyraea Váňa 

1 tatrensis Váňa [331] 

182 Palustriella Ochyra 

1 commutata (Hedw.) Ochyra (Cratoneuron 

commutatum (Hedw.) G.Roth, P. commutata var. 

fluctuans (Schimp.) Ochyra, P. commutata var. 

virescens (Schimp.) Ochyra) 

2 decipiens (De Not.) Ochyra (Cratoneuron decipiens 

(De Not.) Loeske)

3 falcata (Brid.) Hedenäs (Cratoneuron falcatum (Brid.) 

G.Roth, P. commutata var. falcata (Brid.) Ochyra, 

P. commutata var. sulcata (Lindb.) Ochyra) 

4 pluristratosa M.Stech & J.-P.Frahm [334] 

183 Pseudocalliergon (Limpr.) Loeske 

1 angustifolium Hedenäs [335] 

2 brevifolium (Lindb.) Hedenäs (Drepanocladus 

brevifolius (Lindb.) Warnst.) [336] 

3 lycopodioides (Brid.) Hedenäs (Drepanocladus 

lycopodioides (Brid.) Warnst.) 

4 trifarium (F.Weber & D.Mohr) Loeske (Calliergon 

trifarium (F.Weber & D.Mohr) Kindb.) 

5 turgescens (T.Jensen) Loeske (Scorpidium turgescens 

(T.Jensen) Loeske) 

184 Sanionia Loeske [327] 

1 georgicouncinata (Müll.Hal.) Ochyra & Hedenäs 

(S. nivalis Hedenäs) 

2 orthothecioides (Lindb.) Loeske 

3 uncinata (Hedw.) Loeske (Drepanocladus uncinatus 

(Hedw.) Warnst.) 

185 Tomentypnum Loeske [327] 

1 nitens (Hedw.) Loeske 

Calliergonaceae (Kanda) Vanderp., Hedenäs, C.J.Cox & 

A.J.Shaw [327] 

186 Calliergon (Sull.) Kindb. [337] 

1 cordifolium (Hedw.) Kindb. 

2 giganteum (Schimp.) Kindb. 

3 megalophyllum Mikut. 

4 richardsonii (Mitt.) Kindb. (C. obtusifolium Karczm.) 

[338] 

187 Hamatocaulis Hedenäs 

1 lapponicus (Norrl.) Hedenäs (Drepanocladus 

lapponicus (Norrl.) Smirnova) 

2 vernicosus (Mitt.) Hedenäs (Drepanocladus vernicosus 

(Mitt.) Warnst.) 

188 Loeskypnum H.K.G.Paul 

1 badium (Hartm.) H.K.G.Paul (Drepanocladus badius 

(Hartm.) G.Roth) 

189 Scorpidium (Schimp.) Limpr. 

1 cossonii (Schimp.) Hedenäs (Drepanocladus cossonii 

(Schimp.) Loeske, Drepanocladus intermedius 

(Lindb.) Warnst., Limprichtia cossonii (Schimp.) 

L.E.Anderson, H.A.Crum & W.R.Buck, Limprichtia 

intermedia (Lindb.) Loeske) 

2 revolvens (Sw. ex anon.) Rubers (Drepanocladus 

revolvens (Sw. ex anon.) Warnst., Limprichtia 

revolvens (Sw. ex anon.) Loeske) [106] 

3 scorpioides (Hedw.) Limpr. 

190 Straminergon Hedenäs 

1 stramineum (Dicks. ex Brid.) Hedenäs (Calliergon 

stramineum (Dicks. ex Brid.) Kindb.) 

191 Warnstorfia Loeske 

1 exannulata (Schimp.) Loeske (Drepanocladus 

exannulatus (Schimp.) Warnst.) 

2 fluitans (Hedw.) Loeske (Drepanocladus fluitans 

(Hedw.) Warnst., Drepanocladus h-schulzei (Limpr.) 

Loeske, W. fluitans var. falcata (Sanio ex 


C.E.O.Jensen) H.A.Crum & L.E.Anderson, W. 

h-schulzei (Limpr.) Loeske) 

3 procera (Renauld & Arnell) Tuom. (Drepanocladus 

procerus (Renauld & Arnell) Warnst.) [339] 

4 pseudostraminea (Müll.Hal.) Tuom. & T.J.Kop. 

(Drepanocladus pseudostramineus (Müll.Hal.) G.Roth) 

5 sarmentosa (Wahlenb.) Hedenäs (Calliergon 

sarmentosum (Wahlenb.) Kindb.) 

6 trichophylla (Warnst.) Tuom. & T.J.Kop. 

(Drepanocladus trichophyllus (Warnst.) Podp.) 

7 tundrae (Arnell) Loeske (Drepanocladus tundrae 

(Arnell) Loeske) 

Leskeaceae Schimp. [340] 

192 Haplocladium (Müll.Hal.) Müll.Hal. 

1 angustifolium (Hampe & Müll.Hal.) Broth. 

(Bryohaplocladium angustifolium (Hampe & 

Müll.Hal.) R.Watan. & Z.Iwats., Thuidium 

angustifolium (Hampe & Müll.Hal.) A.Jaeger) 

2 microphyllum (Hedw.) Broth. (Bryohaplocladium 

microphyllum (Hedw.) R.Watan. & Z.Iwats.) 

3 virginianum (Brid.) Broth. (Bryohaplocladium 

virginianum (Brid.) R.Watan. & Z.Iwats., Thuidium 

virginianum (Brid.) Schimp.) 

193 Lescuraea Schimp. 

1 mutabilis (Brid.) Lindb. ex I.Hagen 

2 saxicola (Schimp.) Molendo 

3 secunda Arnell [341] 

194 Leskea Hedw. 

1 polycarpa Hedw. 

195 Lindbergia Kindb. 

1 brachyptera (Mitt.) Kindb. [342] 

196 Pseudoleskea Schimp. [343] [344] 

1 artariae Thér. (Pseudoleskeopsis artariae (Thér.) Thér.) 

2 incurvata (Hedw.) Loeske (Lescuraea incurvata 

(Hedw.) E.Lawton) 

3 patens (Lindb.) Kindb. (Lescuraea patens Lindb.) 

4 radicosa (Mitt.) Macoun & Kindb. (Lescuraea 

radicosa (Mitt.) Mönk.) 

5 saviana (De Not.) Latzel (Lescuraea saviana (De Not.) 

E.Lawton) 

197 Pseudoleskeella Kindb. [345] 

1 catenulata (Brid. ex Schrad.) Kindb. 

2 nervosa (Brid.) Nyholm (Leskeella incrassata (Lindb. 

ex Broth.) Broth. [346], Leskeella nervosa (Brid.) 

Loeske) 

3 papillosa (Lindb.) Kindb. 

4 rupestris (Berggr.) Hedenäs & L.Söderstr. (P. nervosa 

var. rupestris (Berggr.) Nyholm, P. sibirica (Arnell) 

P.S.Wilson & D.H.Norris) [347] 

5 tectorum (Funck ex Brid.) Kindb. ex Broth. 

198 Ptychodium Schimp. 

1 plicatum (Schleich. ex F.Weber & D.Mohr) Schimp. 

(Lescuraea plicata (Schleich. ex F.Weber & D.Mohr) 

Broth., Pseudoleskea plicata (Schleich. ex F.Weber & 

D.Mohr) Kindb.) 

Thuidiaceae Schimp. [348] 

199 Abietinella Müll.Hal.


1 abietina (Hedw.) M.Fleisch. (Thuidium abietinum 


var. abietina 

var. hystricosa (Mitt.) Sakurai (A. hystricosa (Mitt.) 

Broth., Thuidium abietinum subsp. hystricosum 

(Mitt.) Kindb., Thuidium abietinum var. hystricosum 

(Mitt.) Loeske & Lande) [32] 

200 Helodium Warnst., nom. cons. 

1 blandowii (F.Weber & D.Mohr) Warnst. (Thuidium 

blandowii (F.Weber & D.Mohr) Schimp.) 

201 Pelekium Mitt., nom. cons. [348] 

1 atlanticum (Hedenäs) Hedenäs (Cyrto-hypnum 

atlanticum (Hedenäs) Hedenäs & Sérgio, Thuidium 

atlanticum Hedenäs) [349] 

2 minutulum (Hedw.) Touw (Cyrto-hypnum minutulum 

(Hedw.) W.R.Buck & H.A.Crum, Microthuidium 

minutulum (Hedw.) Warnst., Thuidium minutulum 


202 Thuidiopsis (Broth.) M.Fleisch. 

1 sparsa (Hook.f. & Wilson) Broth. (Cyrto-hypnum 

montei Hedenäs) [350] 

203 Thuidium Schimp. 

1 assimile (Mitt.) A.Jaeger (T. delicatulum var. radicans 

(Kindb.) H.A.Crum, Steere & L.E.Anderson, T. 

philibertii Limpr.) [351] 

2 delicatulum (Hedw.) Schimp. (T. erectum Duby) 

3 recognitum (Hedw.) Lindb. 

4 tamariscinum (Hedw.) Schimp. 

Brachytheciaceae Schimp. [352] 

Eurhynchioideae Milde (Rhynchostegioideae Ignatov & 

Huttunen) [352] 

204 Pseudoscleropodium (Limpr.) M.Fleisch. 

1 purum (Hedw.) M.Fleisch. (Scleropodium purum 

(Hedw.) Limpr.) 

205 Scorpiurium Schimp. 

1 circinatum (Bruch) M.Fleisch. & Loeske (Thamnium 

cossyrense var. melitense Bott.) [353] 

2 deflexifolium (Solms) M.Fleisch. & Loeske 

3 sendtneri (Schimp.) M.Fleisch. (Thamnium cossyrense 

Bott. var. cossyrense, Thamnobryum cossyrense (Bott.) 

A.J.E.Sm.) [353] 

206 Palamocladium Müll.Hal. 

1 euchloron (Müll.Hal.) Wijk & Margad. 

207 Plasteurhynchium M.Fleisch. ex Broth. 

1 meridionale (Schimp.) M.Fleisch. (Eurhynchium 

meridionale (Schimp.) De Not.) 

2 striatulum (Spruce) M.Fleisch. (Eurhynchium 

striatulum (Spruce) Schimp., Isothecium striatulum 

(Spruce) Kindb.) 

208 Eurhynchium Schimp. 

1 angustirete (Broth.) T.J.Kop. (E. striatum subsp. 

zetterstedtii (P.Størmer) Podp., E. striatum var. 

pachycladum G.Roth) 

2 striatum (Hedw.) Schimp. 

209 Platyhypnidium M.Fleisch. 

1 grolleanum Ochyra & Bednarek-Ochyra [354] 

2 lusitanicum (Schimp.) Ochyra & Bednarek-Ochyra 

(P. alopecuroides (Brid.) A.J.E.Sm., Rhynchostegium 

alopecuroides (Brid.) A.J.E.Sm., Rhynchostegium 

lusitanicum (Schimp.) A.J.E.Sm., hom. illeg.) 

3 mutatum Ochyra & Vanderp. [355] 

4 riparioides (Hedw.) Dixon (Rhynchostegium riparioides 

(Hedw.) Cardot) 

5 torrenticola (Ochyra, C.Schmidt & Bültmann) Ochyra 

& Bednarek-Ochyra (Gradsteinia torrenticola Ochyra, 

C.Schmidt & Bültmann) [356] 

210 Rhynchostegium Schimp. 

1 arcticum (I.Hagen) Ignatov & Huttunen (R. murale 

var. arcticum I.Hagen) [357] 

2 confertum (Dicks.) Schimp. (R. surrectum (Mitt.) 

A.Jaeger) [358] 

3 megapolitanum (Blandow ex F.Weber & D.Mohr) 

Schimp. (Brachythecium cardotii H.Winter) [359] 

4 murale (Hedw.) Schimp. 

5 rotundifolium (Scop. ex Brid.) Schimp. 

6 strongylense (Bott.) W.R.Buck & Privitera (Barbella 

strongylensis Bott.) [360] 

Helicodontioideae M.Fleisch. (Rhynchostegielloideae 

Ignatov & Huttunen) [352] [361] 

211 Nobregaea Hedenäs 

1 latinervis Hedenäs [362] 

212 Helicodontium Schwägr. 

1 capillare (Hedw.) A.Jaeger (H. italicum (Schimp.) 

M.Fleisch.) [363] 

213 Rhynchostegiella (Schimp.) Limpr., nom. cons. 

1 bourgaeana (Mitt.) Broth. [364] 

2 curviseta (Brid.) Limpr. (R. curviseta var. curviseta, R. 

curviseta var. laeviseta (W.E.Nicholson & Dixon) 

Podp., R. letourneuxii (Besch.) Broth.) 

3 durieui (Mont.) P.Allorge & Perss. 

4 litorea (De Not.) Limpr. (R. tenella var. litorea (De 

Not.) Mönk.) [365] 

5 macilenta (Renauld & Cardot) Cardot [366] 

6 tenella (Dicks.) Limpr. 

var. meridionalis (Boulay) Zodda 

var. tenella 

7 teneriffae (Mont.) Dirkse & Bouman (R. jacquinii 

(Garov.) Limpr., R. teesdalei (Schimp.) Limpr.) [367] 

8 tenuicaulis (Spruce) Kartt. (Cirriphyllum germanicum 

(Grebe) Loeske & M.Fleisch., Cirriphyllum tenuicaule 

(Spruce) Wijk & Margad.) [368] 

9 trichophylla Dirkse & Bouman [369] 

214 Cirriphyllum Grout 

1 crassinervium (Taylor) Loeske & M.Fleisch. 

(Eurhynchium crassinervium (Taylor) Schimp.) 

2 piliferum (Hedw.) Grout 

215 Oxyrrhynchium (Schimp.) Warnst. 

1 hians (Hedw.) Loeske (Eurhynchium hians (Hedw.) 

Sande Lac., O. swartzii (Turner) Warnst.) 

2 pumilum (Wilson) Loeske (Eurhynchium pumilum 

(Wilson) Schimp., Rhynchostegiella pallidirostris 

(Brid.) Loeske, nom. illeg., Rhynchostegiella pumila 

(Wilson) E.F.Warb.)

3 schleicheri (R.Hedw.) Röll (Eurhynchium schleicheri 

(R.Hedw.) Milde) 

4 speciosum (Brid.) Warnst. (Eurhynchium speciosum 

(Brid.) Jur.) 

Brachythecioideae Engler 

216 Kindbergia Ochyra 

1 praelonga (Hedw.) Ochyra (Eurhynchium praelongum 

(Hedw.) Schimp., Eurhynchium praelongum var. 

stokesii (Turner) Dixon, Eurhynchium stokesii 

(Turner) Schimp., Oxyrrhynchium praelongum (Hedw.) 

Warnst.) 

217 Bryhnia Kaurin 

1 scabrida (Lindb.) Kaurin (B. novae-angliae auct. eur. 

non (Sull. & Lesq.) Grout) [370] 

218 Myuroclada Besch. 

1 maximowiczii (G.G.Borshch.) Steere & W.B.Schofield 

[371] 

219 Sciuro-hypnum Hampe 

1 flotowianum (Sendtn.) Ignatov & Huttunen 

(Cirriphyllum reichenbachianum (Huebener) Wijk & 

Margad., Cirriphyllum velutinoides (Schimp.) Loeske 

& M.Fleisch., nom. illeg., Eurhynchium flotowianum 

(Sendtn.) Kartt.) 

2 glaciale (Schimp.) Ignatov & Huttunen 

(Brachythecium dovrense (Limpr.) Amman, 

Brachythecium glaciale Schimp., Brachythecium 

glaciale var. dovrense Limpr.) [372] 

3 latifolium (Kindb.) Ignatov & Huttunen 

(Brachythecium latifolium Kindb.) 

4 oedipodium (Mitt.) Ignatov & Huttunen 

(Brachythecium curtum (Lindb.) Limpr., 

Brachythecium oedipodium (Mitt.) A.Jaeger, 

Brachythecium starkei var. curtum (Lindb.) Warnst.) 

5 ornellanum (Molendo) Ignatov & Huttunen 

(Brachythecium ornellanum (Molendo) Venturi & 

Bott., Scleropodium apiculigerum (Lindb. & Arnell) 

J.-P.Frahm, Scleropodium ornellanum (Molendo) 

Lorentz) 

6 plumosum (Hedw.) Ignatov & Huttunen, nom. cons. 

(Brachythecium plumosum (Hedw.) Schimp.) 

7 populeum (Hedw.) Ignatov & Huttunen 

(Brachythecium populeum (Hedw.) Schimp.) 

8 reflexum (Starke) Ignatov & Huttunen (Brachythecium 

reflexum (Starke) Schimp.) 

9 starkei (Brid.) Ignatov & Huttunen (Brachythecium 

starkei (Brid.) Schimp., Brachythecium starkei var. 

tromsoeense (Kaurin & Arnell) Nyholm) 

220 Brachythecium Schimp. [373] 

1 albicans (Hedw.) Schimp. 

2 campestre (Müll.Hal.) Schimp. (B. ryanii Kaurin) [374] 

3 capillaceum (F.Weber & D.Mohr) Giacom. (B. 

rotaeanum De Not. [375], B. salebrosum subsp. 

rotaeanum (De Not.) Amman, B. salebrosum var. 

capillaceum (F.Weber & D.Mohr) Lorentz) 

4 cirrosum (Schwägr.) Schimp. (Cirriphyllum cirrosum 

(Schwägr.) Grout) 


5 coruscum I.Hagen (B. groenlandicum (C.E.O.Jensen) 

Schljakov) [376] 

6 erythrorrhizon Schimp. 

subsp. asiaticum Ignatov [377] 

subsp. erythrorrhizon 

var. erythrorrhizon 

var. thedenii (Schimp.) Lindb. [378] 

7 geheebii Milde (Homalothecium geheebii (Milde) Wigh) 

8 glareosum (Bruch ex Spruce) Schimp. 

9 laetum (Brid.) Schimp. (B. oxycladon auct. non (Brid.) 

A.Jaeger) [379] 

10 mildeanum (Schimp.) Schimp. 

var. mildeanum 

var. udum (I.Hagen) Mönk. (B. udum I.Hagen) 

11 percurrens Hedenäs [380] 

12 rivulare Schimp. 

13 rutabulum (Hedw.) Schimp. 

var. atlanticum Hedenäs [381] 

var. rutabulum 

14 salebrosum (Hoffm. ex F.Weber & D.Mohr) Schimp., 

nom. cons. 

15 tommasinii (Sendtn. ex Boulay) Ignatov & Huttunen 

(Cirriphyllum tenuinerve (Lindb.) Wijk & Margad., 

Cirriphyllum tommasinii (Sendtn. ex Boulay) Grout) 

16 turgidum (Hartm.) Kindb. 

Homalothecioideae Ignatov & Huttunen 

221 Scleropodium Bruch & Schimp. 

1 cespitans (Wilson ex Müll.Hal.) L.F.Koch 

(Brachythecium appleyardiae McAdam & A.J.E.Sm.) 

[382] 

2 touretii (Brid.) L.F.Koch [32] 

222 Eurhynchiastrum Ignatov & Huttunen 

1 pulchellum (Hedw.) Ignatov & Huttunen 

(Eurhynchium pulchellum (Hedw.) Jenn.) 

var. diversifolium (Schimp.) Ochyra & Z˙ arnowiec 

(Eurhynchium pulchellum var. diversifolium (Schimp.) 

C.E.O.Jensen) 

var. praecox (Hedw.) Ochyra & Z˙ arnowiec 

(Eurhynchium pulchellum var. praecox (Hedw.) Dixon) 

var. pulchellum 

223 Brachytheciastrum Ignatov & Huttunen 

1 collinum (Schleich. ex Müll.Hal.) Ignatov & Huttunen 

(Brachythecium collinum (Schleich. ex Müll.Hal.) Schimp.) 

2 dieckii (Röll) Ignatov & Huttunen (Brachythecium 

dieckii Röll, Brachythecium salteri Cardot & Dixon) 

[383] 

3 fendleri (Sull.) Vanderp. et al. (Brachythecium fendleri 

(Sull.) A.Jaeger) [384] 

4 olympicum (Jur.) Vanderp. et al. (Brachythecium 

olympicum Jur.) [384] 

5 trachypodium (Brid.) Ignatov & Huttunen (Brachythecium 

trachypodium (Brid.) Schimp., Brachythecium 

trachypodium var. payotianum (Boulay) Bott.) 

6 vanekii (Sˇmarda) Ochyra & Z˙ arnowiec (Brachythecium 

vanekii Sˇmarda) 

7 velutinum (Hedw.) Ignatov & Huttunen 

(Brachythecium velutinum (Hedw.) Schimp.)


var. salicinum (Schimp.) Ochyra & Z˙ arnowiec 

(Brachythecium velutinum var. salicinum (Schimp.) 

Mönk.) 

var. vagans (Milde) Ochyra & Z˙ arnowiec 

(Brachythecium velutinum var. vagans (Milde) Warnst.) 

var. velutinum 

224 Homalothecium Schimp. 

1 aureum (Spruce) H.Rob. 

2 lutescens (Hedw.) H.Rob. (Camptothecium lutescens 


var. fallax H.Philib. ex Schimp. 

var. lutescens 

3 philippeanum (Spruce) Schimp. 

4 sericeum (Hedw.) Schimp. 

Myriniaceae Schimp. 

225 Myrinia Schimp., nom. cons. 

1 pulvinata (Wahlenb.) Schimp. 

Fabroniaceae Schimp. 

226 Fabronia Raddi 

1 ciliaris (Brid.) Brid. 

2 pusilla Raddi 

Hypnaceae Schimp. [340][385] 

227 Andoa Ochyra 

1 berthelotiana (Mont.) Ochyra 

228 Breidleria Loeske 

1 pratensis (W.D.J.Koch ex Spruce) Loeske (Hypnum 

pratense W.D.J.Koch ex Spruce) [386] 

229 Callicladium H.A.Crum 

1 haldanianum (Grev.) H.A.Crum 

230 Calliergonella Loeske 

1 cuspidata (Hedw.) Loeske 

2 lindbergii (Mitt.) Hedenäs (Hypnum lindbergii Mitt.) 

[386] 

231 Campylophyllum (Schimp.) M.Fleisch. [387] 

1 calcareum (Crundw. & Nyholm) Hedenäs 

(Campylidium calcareum (Crundw. & Nyholm) 

Ochyra, Campylium calcareum Crundw. & Nyholm) 

2 halleri (Hedw.) M.Fleisch. (Campylium halleri (Hedw.) 

Lindb.) 

3 sommerfeltii (Myrin) Hedenäs (Campylidium 

sommerfeltii (Myrin) Ochyra, Campylium hispidulum 

auct. eur. non (Brid.) Mitt. [388], Campylium 

sommerfeltii (Myrin) Lange, Campylophyllum 

hispidulum auct. eur. non (Brid.) Hedenäs [388]) 

232 Ctenidium (Schimp.) Mitt. 

1 molluscum (Hedw.) Mitt. [389] 

233 Homomallium (Schimp.) Loeske 

1 incurvatum (Schrad. ex Brid.) Loeske 

234 Hyocomium Bruch & Schimp. 

1 armoricum (Brid.) Wijk & Margad. 

235 Hypnum Hedw., nom. cons. 

1 andoi A.J.E.Sm. (H. cupressiforme var. mammillatum 

Brid., H. mammillatum (Brid.) Loeske, nom. inval.) 

2 bambergeri Schimp. (Stereodon bambergeri (Schimp.) 

Lindb.) 

3 callichroum Brid. (Stereodon callichrous (Brid.) 

Braithw.) 

4 cupressiforme Hedw. [390] 

var. cupressiforme 

var. filiforme Brid. 

var. heseleri (Ando & Higuchi) M.O.Hill (H. heseleri 

Ando & Higuchi) [391] 

var. lacunosum Brid. (H. cupressiforme subsp. 

lacunosum (Brid.) Bertsch, H. lacunosum (Brid.) 

Hoffm. ex Brid.) 

var. resupinatum (Taylor) Schimp. (H. cupressiforme 

subsp. resupinatum (Taylor) Hartm., H. resupinatum 

Taylor) 

var. subjulaceum Molendo 

5 fertile Sendtn. (Stereodon fertilis (Sendtn.) Lindb.) 

6 hamulosum Schimp. (Stereodon hamulosus (Schimp.) 

Lindb.) [392] 

7 holmenii Ando (Stereodon holmenii (Ando) Ignatov 

& Ignatova) [393] 

8 imponens Hedw. 

9 jutlandicum Holmen & E.Warncke (H. cupressiforme 

var. ericetorum Schimp.) 

10 pallescens (Hedw.) P.Beauv. (Stereodon pallescens 

(Hedw.) Mitt.) 

11 plicatulum (Lindb.) A.Jaeger (Stereodon plicatulus 

Lindb.) 

12 procerrimum Molendo (Ctenidium procerrimum 

(Molendo) Lindb.) 

13 recurvatum (Lindb. & Arnell) Kindb. (Drepanium 

recurvatum (Lindb. & Arnell) G.Roth) 

14 revolutum (Mitt.) Lindb. (Stereodon revolutus Mitt.) 

var. dolomiticum (Milde) Mönk. (H. dolomiticum 

Milde) 

var. revolutum 

15 sauteri Schimp. 

16 subimponens Lesq. [394] 

17 uncinulatum Jur. 

18 vaucheri Lesq. (Stereodon vaucheri (Lesq.) Lindb. ex 

Broth.) 

236 Ptilium De Not. 

1 crista-castrensis (Hedw.) De Not. 

237 Pylaisia Schimp., nom. cons. 

1 polyantha (Hedw.) Schimp. (Pylaisiella polyantha 

(Hedw.) Grout) 

2 selwynii Kindb. (Pylaisiella selwynii (Kindb.) 

H.A.Crum, Steere & L.E.Anderson) 

238 Taxiphyllum M.Fleisch. 

1 densifolium (Lindb. ex Broth.) Reimers 

2 wissgrillii (Garov.) Wijk & Margad. 

239 Vesicularia (Müll.Hal.) Müll.Hal. 

1 reimersiana Bizot & P.de la Varde [395] 

Pterigynandraceae Schimp. 

240 Habrodon Schimp. 

1 perpusillus (De Not.) Lindb. 

241 Heterocladium Schimp. 

1 dimorphum (Brid.) Schimp. 

2 flaccidum (Schimp.) A.J.E.Sm. (H. heteropterum var. 

flaccidum Schimp.) [396] 

3 heteropterum (Brid.) Schimp.

4 wulfsbergii I.Hagen (H. heteropterum subsp. 

wulfsbergii (I.Hagen) C.E.O.Jensen & Perss.) [397] 

242 Iwatsukiella W.R.Buck & H.A.Crum 

1 leucotricha (Mitt.) W.R.Buck & H.A.Crum [398] 

243 Pterigynandrum Hedw. 

1 filiforme Hedw. 

var. filiforme 

var. majus (De Not.) De Not. 

Hylocomiaceae (Broth.) M.Fleisch. 

244 Hylocomiastrum Broth. 

1 pyrenaicum (Spruce) M.Fleisch. (Hylocomium 

pyrenaicum (Spruce) Lindb.) 

2 umbratum (Hedw.) M.Fleisch. (Hylocomium umbratum 


245 Hylocomium Schimp., nom. cons. 

1 splendens (Hedw.) Schimp. [399] 

246 Loeskeobryum Broth. 

1 brevirostre (Brid.) M.Fleisch. (Hylocomium brevirostre 

(Brid.) Schimp.) 

247 Pleurozium Mitt., nom. cons. 

1 schreberi (Willd. ex Brid.) Mitt. 

248 Rhytidiadelphus (Limpr.) Warnst. 

1 loreus (Hedw.) Warnst. 

2 squarrosus (Hedw.) Warnst. (Rhytidiastrum 

squarrosum (Hedw.) Ignatov & Ignatova) 

3 subpinnatus (Lindb.) T.J.Kop. (R. squarrosus var. 

calvescens (Kindb.) Warnst., Rhytidiastrum 

subpinnatum (Lindb.) Ignatov & Ignatova) 

4 triquetrus (Hedw.) Warnst. 

Rhytidiaceae Broth. 

249 Rhytidium (Sull.) Kindb. 

1 rugosum (Hedw.) Kindb. 

Plagiotheciaceae (Broth.) M.Fleisch. 

250 Herzogiella Broth. [400] 

1 seligeri (Brid.) Z.Iwats. 

2 striatella (Brid.) Z.Iwats. 

3 turfacea (Lindb.) Z.Iwats. 

251 Isopterygiopsis Z.Iwats. 

1 alpicola (Lindb. & Arnell) Hedenäs (Isopterygium 

alpicola (Lindb. & Arnell) Nyholm) 

2 muelleriana (Schimp.) Z.Iwats. (Isopterygium 

muellerianum (Schimp.) A.Jaeger) 

3 pulchella (Hedw.) Z.Iwats. (Isopterygium pulchellum 

(Hedw.) A.Jaeger) 

252 Myurella Schimp. 

1 julacea (Schwägr.) Schimp. [401] 

2 sibirica (Müll.Hal.) Reimers 

3 tenerrima (Brid.) Lindb. 

253 Orthothecium Schimp., nom. cons. 

1 chryseon (Schwägr.) Schimp. [402] 

2 intricatum (Hartm.) Schimp. 

3 lapponicum (Schimp.) C.Hartm. 

4 rufescens (Dicks. ex Brid.) Schimp. 

5 strictum Lorentz 

254 Plagiothecium Schimp. 

1 berggrenianum Frisvoll 

2 cavifolium (Brid.) Z.Iwats. (P. roeseanum Schimp.) 


3 curvifolium Schlieph. ex Limpr. (P. laetum var. 

curvifolium (Schlieph. ex Limpr.) Mastracci & 

M.Sauer, P. laetum var. secundum (Lindb.) Frisvoll 

et al.) [403] 

4 denticulatum (Hedw.) Schimp. 

var. denticulatum 

var. obtusifolium (Turner) Moore 

var. undulatum R.Ruthe ex Geh. (P. denticulatum var. 

ruthei (Limpr.) Riehm., P. ruthei Limpr.) [404] 

5 laetum Schimp. 

6 latebricola Schimp. 

7 neckeroideum Schimp. (P. neckeroideum subsp. 

noricum (Molendo ex Limpr.) J.J.Amann, P. noricum 

Molendo ex Limpr.) [405] 

8 nemorale (Mitt.) A.Jaeger 

9 piliferum (Sw.) Schimp. 

10 platyphyllum Mönk. 

11 succulentum (Wilson) Lindb. [406] 

12 svalbardense Frisvoll [407] 

13 undulatum (Hedw.) Schimp. (Buckiella undulata 

(Hedw.) Ireland) 

255 Platydictya Berk. 

1 jungermannioides (Brid.) H.A.Crum (Amblystegium 

jungermannioides (Brid.) A.J.E.Sm.) 

256 Pseudotaxiphyllum Z.Iwats. 

1 elegans (Brid.) Z.Iwats. (Isopterygium elegans (Brid.) 

Lindb.) 

2 laetevirens (Dixon & Luisier ex F.Koppe & Düll) 

Hedenäs [408] 

Entodontaceae Kindb., nom. cons. 

257 Entodon Müll.Hal. 

1 challengeri (Paris) Cardot (E. compressus Müll.Hal. 

ex Cardot non (Hedw.) Müll.Hal.) [409] 

2 cladorrhizans (Hedw.) Müll.Hal. 

3 concinnus (De Not.) Paris 

4 schleicheri (Schimp.) Demet. 

Pylaisiadelphaceae Goffinet & W.R.Buck 

258 Brotherella Loeske ex M.Fleisch. 

1 lorentziana (Molendo ex Lorentz) Loeske ex M.Fleisch. 

259 Heterophyllium (Schimp.) Kindb. 

1 affine (Hook.) M.Fleisch. 

260 Isopterygium Mitt. 

1 tenerum (Sw.) Mitt. (I. bottinii (Breidl.) Kindb., 

Sematophyllum bottinii (Breidl.) Podp.) [410] 

261 Platygyrium Schimp., nom. cons. 

1 repens (Brid.) Schimp. 

Sematophyllaceae Broth., nom. cons. 

262 Hageniella Broth. 

1 micans (Mitt.) B.C.Tan & Y.Jia (Hygrohypnum micans 

(Mitt.) Broth., Sematophyllum micans (Mitt.) 

Braithw.) [411] 

263 Sematophyllum Mitt. 

1 adnatum (Michx.) E.Britton [412] 

2 demissum (Wilson) Mitt. 

3 substrumulosum (Hampe) E.Britton 

Cryphaeaceae Schimp. 

264 Cryphaea D.Mohr


1 heteromalla (Hedw.) D.Mohr 

265 Dendrocryphaea Paris & Schimp. ex Broth. 

1 lamyana (Mont.) P.Rao (Cryphaea lamyana (Mont.) 

Müll.Hal.) [413] 

Leucodontaceae Schimp. 

266 Antitrichia Brid. 

1 californica Sull. 

2 curtipendula (Hedw.) Brid. 

267 Leucodon Schwägr. 

1 canariensis (Brid.) Schwägr. 

2 flagellaris Lindb. ex Broth. [414] 

3 immersus Lindb. 

4 pendulus Lindb. [415] 

5 sciuroides (Hedw.) Schwägr. 

var. morensis (Schwägr.) De Not. 

var. sciuroides 

6 treleasei (Cardot) Paris [416] 

268 Pterogonium Sw. 

1 gracile (Hedw.) Sm. 

Neckeraceae Schimp. 

269 Homalia Brid., nom. cons. 

1 lusitanica Schimp. (H. lusitanica var. subrecta (Mitt.) 

Düll-Herm., H. subrecta (Mitt.) A.Jaeger, Neckera 

subrecta Mitt.) 

2 trichomanoides (Hedw.) Brid. [417] 

3 webbiana (Mont.) Schimp. (Neckera webbiana (Mont.) 

Düll) 

270 Neckera Hedw., nom. cons. 

1 besseri (Lobarz.) Jur. (Homalia besseri Lobarz.) 

2 cephalonica Jur. & Unger (N. pennata subsp. 

cephalonica (Jur. & Unger) Giacom.) 

3 complanata (Hedw.) Huebener (Homalia complanata 

(Hedw.) De Not., N. complanata var. longifolia 

Schimp., N. complanata var. secunda Grav., N. 

complanata var. tenella Schimp.) 

4 crispa Hedw. (N. crispa var. falcata Müll.Hal., 

N. pseudopennata (Warnst.) Schlieph. ex Z˙ muda) 

5 intermedia Brid. (N. elegans Jur., N. elegans var. 

laevifolia Schiffn., N. intermedia var. laevifolia 

(Schiffn.) Renauld & Cardot, N. laeviuscula Geh.) 

6 menziesii Drumm. (Metaneckera menziesii (Drumm.) 

Steere) [418] 

7 oligocarpa Bruch (N. pennata subsp. oligocarpa 

(Bruch) Giacom., N. pennata var. tenera Müll.Hal.) 

8 pennata Hedw. 

9 pumila Hedw. (N. fontinaloides Lindb., 

N. fontinaloides var. philippeana (Schimp.) Guim., 

N. philippeana Schimp., N. pumila var. philippeana 

(Schimp.) Milde, N. pumila var. pilifera Jur.) 

271 Thamnobryum Nieuwl. 

1 alopecurum (Hedw.) Gangulee (T. alopecurum var. 

gracillimum Bott., T. mediterraneum (Bott.) G.Roth) 

2 angustifolium (Holt) Nieuwl. 

3 cataractarum N.G.Hodgetts & Blockeel [419] 

4 fernandesii Sérgio (Crassiphyllum fernandesii (Sérgio) 

Ochyra) [420] 

5 maderense (Kindb.) Hedenäs (T. alopecurum var. 

maderense (Kindb.) M.Stech, Ros & O.Werner) [421] 

6 neckeroides (Hook.) E.Lawton [422] 

7 rudolphianum Mastracci [423] 

Echinodiaceae Broth. 

272 Echinodium Jur. 

1 prolixum (Mitt.) Broth. (Scleromnium knyi Jur.) 

2 renauldii (Cardot) Broth. 

3 setigerum (Mitt.) Jur. (E. setigerum var. integrifolium 

Luisier) [424] 

4 spinosum (Mitt.) Jur. (E. madeirense Jur.) 

Leptodontaceae Schimp. 

273 Cryptoleptodon Renauld & Cardot 

1 longisetus (Mont.) Enroth (Leptodon longisetus Mont., 

Neckera longipedunculata Müll.Hal.) [425] 

274 Leptodon D.Mohr, nom. cons. 

1 smithii (Hedw.) F.Weber & D.Mohr 

Lembophyllaceae Broth. 

275 Isothecium Brid. 

1 algarvicum W.E.Nicholson & Dixon (Forsstroemia 

canariensis (Renauld & Cardot) Enroth, I. atlanticum 

Hedenäs, nom. illeg., I. canariense H.Winter, 

Thamnium canariense Renauld & Cardot, 

Thamnobryum canariense (Renauld & Cardot) 

D.G.Long) [426] 

2 alopecuroides (Lam. ex Dubois) Isov. (I. alopecuroides 

var. robustum (Schimp.) Düll, I. circinans Saut., 

I. myurum Brid., I. viviparum Lindb.) 

3 holtii Kindb. (Eurhynchium myosuroides var. rivulare 

(Limpr.) Paris, I. myosuroides var. rivulare Limpr.) 

[427] 

4 myosuroides Brid. 

subsp. brevinerve Lindb. (I. myurum var. piliferum 

C.E.O.Jensen, I. myurum var. tenuinerve (Kindb.) 

Limpr.) 

subsp. myosuroides (I. myosuroides var. cavernarum 

Molendo, I. myosuroides var. debile Braithw., I. 

myosuroides var. falcatum Jaap ex G.Roth, I. 

myosuroides var. filescens (Renauld) Warnst., I. 

myosuroides var. filiforme Jaap ex G.Roth, 

I. myosuroides var. integrifolium Papp) 

var. brachythecioides (Dixon) Braithw. [427] 

Myuriaceae M.Fleisch. 

276 Myurium Schimp. 

1 hochstetteri (Schimp.) Kindb. (M. hebridarum Schimp.) 

Anomodontaceae Kindb. 

277 Anomodon Hook. & Taylor 

1 attenuatus (Hedw.) Huebener 

2 longifolius (Schleich. ex Brid.) Hartm. 

3 rostratus (Hedw.) Schimp. 

4 rugelii (Müll.Hal.) Keissl. 

5 tristis (Ces.) Sull. & Lesq. (Haplohymenium triste 

(Ces.) Kindb.) 

6 viticulosus (Hedw.) Hook. & Taylor 

278 Claopodium (Lesq. & James) Renauld & Cardot 

1 whippleanum (Sull.) Renauld & Cardot 

(C. whippleanum var. cavernicola Luisier)

ANNOTATIONS 

1. Ignatov & Afonina (1992) report Sphagnum imbricatum 

Hornsch. ex Russow (section Sphagnum), S. 

orientale L.I.Savicz (section Subsecunda) and S. 

perfoliatum L.I.Savicz (section Subsecunda) from 

arctic European Russia. The record of S. imbricatum 

would, if correct, be likely to refer to S. steerei 

R.E.Andrus, which is seemingly the only taxon of the 

S. imbricatum complex collected in arctic areas of 

North America and Eurasia (Flatberg, 1984), and 

which is listed by Dierssen (2001). However, Afonina 

(pers. comm. to Flatberg) tells us that there are no 

reliable herbarium specimens of European material of 

any of these species. They are therefore omitted. 

2. Sphagnum viride is closely related to S. cuspidatum 

(Flatberg, 1988), and isoenzymatic results indicate 

that gene flow exists between the two morphs 

(Hanssen, Sa˚stad & Flatberg, 2000). More genetic 

data are required to evaluate their taxonomic status. 

3. Sphagnum brevifolium and S. isoviitae belong to a 

small group of closely related taxa in the S. recurvum 

complex with yellow spores (Flatberg, 1992, 1992 

[1993]). In typical morphological appearance they are 

well separated from S. fallax, but seem not clearly 

segregated genetically in sympatric populations 

(Sa˚stad, Stenøien & Flatberg, 1999). Their species 

status can therefore be questioned, and more genetic 

data are required before a decisive taxonomic conclusion 

can be drawn. 

4. The name Sphagnum lenense was published as a 

nomen nudum in 1915, and was not validated until 

1936 (Afonina and Egorova, pers. comm. to Flatberg). 

5. Sphagnum auriculatum is retained here as the name for 

the taxon that has, following a paper by Dirkse & 

Isoviita (1986), been called S. denticulatum in recent 

lists. Sphagnum denticulatum is an aquatic form with 

abnormally large, nearly isophyllous leaves. Although 

most such plants belong to S. auriculatum, similar 

forms of S. inundatum are found not uncommonly 

growing under aquatic conditions in mire pools along 

the western coast of Norway. The type specimen of S. 

denticulatum is therefore unsatisfactory. 

6. Sphagnum tundrae was described from Svalbard by 

Flatberg (1994). 

7. Sphagnum wulfianum is either nested within section 

Acutifolia (Shaw, 2000b) or is sister to section 

Acutifolia, lying between it and section Squarrosa 

(Shaw, Cox & Boles, 2005). It is left here in its own 

section Polyclada. 

8. Sphagnum aongstroemii may be nested within section 

Acutifolia (Shaw, 2000b, Shaw et al., 2005). It is left 

here in its own section Insulosa. 

9. Sphagnum nitidulum Warnst. was described from 

Terceira in the Azores, but type material has 

presumably been destroyed and its identity is obscure. 


10. Sphagnum subtile (Russow) Warnst. (S. capillifolium 

var. subtile (Russow) Kartt., S. rubellum var. subtile 

(Russow) Amann) is a disputed taxon whose distinctness 

can be questioned. Natcheva & Cronberg (2002) 

consider European herbarium material studied to 

belong to S. rubellum. Shaw et al. (2005) claim that 

North American plants named S. subtile cannot be 

separated genetically from S. capillifolium. The name 

is based on European material, but original material 

has not been traced. It is premature to disregard the 

existence of a European taxon underlying this name. 

11. Sphagnum tenerum Sull. & Lesq. ex Sull. (S. capillifolium 

var. tenerum (Sull. & Lesq. ex Sull.) H.A.Crum) 

is an eastern North American species, which according 

to Shaw et al. (2005) is highly differentiated genetically 

from S. capillifolium. European plants labelled S. 

tenerum have been examined by Flatberg, and belong 

to hemi-isophyllous forms of S. capillifolium. 

12. Sphagnum olafii was described from Svalbard by 

Flatberg (1993a). 

13. Sphagnum rubiginosum was described from Norway by 

Flatberg (1993b). 

14. Sphagnum andersonianum has been reported from 

Europe, but Shaw et al. (2005) claim that examined 

material from Europe and North America cannot be 

distinguished by genetic markers from S. rubellum. 

15. Sections of Andreaea follow Murray (1988). 

16. Sérgio (2004) recognized Andreaea heinemannii subsp. 

crassifolia, stating that it does not intergrade morphologically 

with subsp. heinemannii and that they are 

found in one mixed population. The taxon may thus 

merit species status. 

17. Smith (2004) treated the East Asian Atrichum rhystophyllum 

(Müll.Hal.) Paris as a synonym of A. 

angustatum, following Richards & Wallace (1950), 

who treated it as a variety of A. angustatum. The 

relationships of North American, European and East 

Asian representatives of this complex and their 

affinity with A. undulatum are not clear and are in 

need of thorough revision, as stated by Lou & 

Koponen (1986). 

18. A critical, global revision is required to clarify the 

status of Atrichum flavisetum and its relationship to A. 

undulatum. 

19. Pogonatum inflexum (Lindb.) Sande Lac. is listed for 

Turkey by Kürschner & Erdag˘ (2005). Pogonatum 

inflexum is an exclusively East Asian species confined 

to China, Japan, Korea and the Russian Far East. 

Several records from other parts of mainland Asia 

have been confirmed as misidentified, and are mostly 

P. neesii (Hyvönen, 1989). We have not studied any 

Turkish material but are inclined to assume that these 

records also represent P. neesii, a widespread Asian 

species with a disjunct occurrence in the Caucasus 

area (see below). 

20. According to Schratz (1928), Pogonatum aloides var. 

minimum (Crome) Molendo and P. nanum var. long-


isetum Hampe ex Bruch & Schimp. are hybrids 

between P. aloides and P. nanum. 

21. Pogonatum neesii was reported for European Russia 

for the first time by Akatova & Ignatova (2000). 

22. According to the latest phylogenetic analyses 

(Hyvönen et al., 2004, Koskinen & Hyvönen, 2004), 

neither Polytrichastrum nor the traditional Polytrichum 

sensu lato is monophyletic. Furthermore, several 

Polytrichastrum species have been shown to be 

allopolyploids (Derda & Wyatt, 2000; Van der Velde 

& Bijlsma, 2001). However, as Polytrichum sensu 

stricto (Smith, 1971) may be monophyletic, and as the 

paraphyly of Polytrichastrum is due to some taxa 

being more closely related to some Pogonatum species, 

we continue to accept Polytrichastrum until such time 

as more comprehensive phylogenetic analyses have 

been completed. 

23. No infraspecific taxa of Polytrichastrum alpinum are 

recognized here, not even var. fragile, which Long 

(1985) recognized, although with reservations. 

Results obtained by Yli-Rekola (1980) and 

Schriebl (1991) in culture experiments support this 

wide circumscription of the species, although 

neither study included var. fragile. Although this 

taxon is easily identified by its caducous leaves that 

are constricted at the line of dehiscence, this 

tendency may be a result of phenotypic plasticity 

associated with certain habitats or environmental 

stimuli. A similar feature has been observed by 

Long (1988) at the population level in Pogonatum 

urnigerum (which recent phylogenetic analyses suggest 

is closely related to Polytrichastrum alpinum) 

and by both Holmen (1960) and Long (1985) for 

Pogonatum dentatum. As the other distinguishing 

characters of var. fragile are well within the range 

of variation for the species, it would be inconsistent 

to treat this feature as taxonomically significant in 

one taxon only. 

24. Polytrichastrum sphaerothecium was treated by Corley 

et al. (1981) as a synonym of P. sexangulare and by 

Merrill (1992) as a variety of P. sexangulare. 

25. Recent genetic (Van der Velde & Bijlsma, 2000) and 

culture experiments (Schriebl, 1991) based on 

European material support the recognition of 

several closely related species formerly treated as 

varieties of Polytrichum commune. Differences 

between some of these are small, and studies of 

North American material have not supported all of 

them (Derda & Wyatt, 1990). A global study 

including examination of type material is required 

to stabilize the taxonomy. We follow Long (1985) 

and Bijlsma et al. (2000) in reducing P. commune 

var. humile and var. perigoniale to synonymy under 

P. commune. As noted by Long (1985), perichaetial 

leaf length varies significantly in P. commune and 

minor differences in form hardly warrant taxonomic 

recognition. 

26. The generic status of Bryobrittonia is questionable. 

Recent European checklists followed Horton (1983) in 

treating it as a genus, whereas Nyholm (1998) treated 

it as a member of Encalypta section Streptotheca. The 

question is still unresolved, so we follow previous 

checklists by treating Bryobrittonia as a genus. 

27. Encalypta sections follow Horton (1983). 

28. Encalypta serbica Katić was described from Serbia 

in 1906, but has not been refound. Horton (1983) 

was unable to trace a type specimen, but thought from 

the description that it was probably a form of E. 

ciliata. 

29. The occurrence of Encalypta intermedia in Europe has 

hitherto not been well documented. It was described 

from Harput and Berit Dagt in Turkey (D.G. Horton, 

pers. comm.), and is listed for Turkey by Kürschner & 

Erdag˘ (2005). It was added to the European list by 

Corley & Crundwell (1991) because Horton (1983) 

stated that it occurred in Europe. Horton (pers. 

comm.) has confirmed two specimens (B) collected 

by J. Bornmüller in 1907 from Gröden in the South 

Tyrol and determined by him as E. commutata. 

30. Encalypta obovatifolia was described from Gotland 

and Öland by Nyholm (1995 [1996]). 

31. The status of Encalypta rhaptocarpa var. leptodon is 

disputed. Nyholm (1998) and Mogensen (2001) accord 

it species rank as E. trachymitria, while Ignatov & 

Ignatova (2003) treat it as a form of E. rhaptocarpa 

and Kučera & Váňa (2003) treat it as doubtful. 

32. The original spelling of names terminating with -colla 

or -collum is retained, in accordance with article 60.1 

of the International Code (Greuter et al., 2000). The 

argument of Crundwell (1970) in favour of the -collis 

ending is not sustainable. Trematodon brevicollis and 

T. longicollis are, however, correct. We have also 

checked the original spelling of Hypnum 

(Scleropodium) touretii Brid., Splachnum (Aplodon) 

wormskioldii Hornem. and Thuidium hystricosum Mitt. 

(Abietinella abietina var. hystricosa). 

33. Encalypta affinis subsp. macounii is commonly treated 

as a species, but Horton (1983; and pers. comm. to 

Hill) points out that it differs from subsp. affinis only 

in the muticous rather than hair-pointed leaves. 

34. Entosthodon subpallescens Laz. (Funaria subpallescens 

(Laz.) Smirnova) has been included in European lists 

but is Asian. 

35. The hybrid taxa Funaria6hybrida R.Ruthe ex Limpr. 

(Entosthodon fascicularis6Funaria hygrometrica) and 

Physcomitrella6hampei Limpr. (usually interpreted as 

Physcomitrella patens6Physcomitrium eurystomum) 

are omitted from the main list. 

36. The genera Funaria and Entosthodon are distinguished 

according to the revision by Fife (1985). 

37. For synonyms of Entosthodon durieui see Brugués 

(1998). 

38. For synonymy of Entosthodon hungaricus see Cano 

et al. (1999).

39. The poorly known Macaronesian endemic Funaria 

fritzei is treated by Losada-Lima, Dirkse & 

Rodríguez-Núñez (2001) as a synonym of 

Entosthodon krausei. It needs further elucidation. 

40. Entosthodon schimperi Brugués is a new name for 

the moss described by Schimper as Funaria durieui. 

The epithet durieui is not available in Entosthodon. 

The species has been found in Portugal, Spain and 

the Canary Islands (Brugués, Dirkse & Sérgio, 

2001). 

41. Entosthodon mouretii was reported from Spain by 

Brugués et al. (1999). 

42. Funaria maireana Copp., from Greece, is too poorly 

known to be included in the checklist. F. anomala Jur. 

and Funaria handelii Schiffn. (Entosthodon handelii 

(Schiffn.) Laz.) are listed for Turkey by Kürschner & 

Erdag˘ (2005). 

43. Goffinet & Buck (2004) placed Goniomitrium in 

Pottiales on the basis of a molecular study by 

Goffinet & Cox (2000). However, the following 

morphological features favour Funariales: laminal 

cells smooth, large and thin-walled; stomata with 

simple guard cells; calyptra mitriform. 

44. Fife & Seppelt (2001) expressed doubts about the 

distinctness of Goniomitrium seroi from the Australian 

G. acuminatum Hook. & Wilson. A fuller study of 

Australian and Spanish material is required before 

these two species can be treated as synonyms. 

45. Physcomitrium eurystomum subsp. acuminatum is 

treated as a species in the Polish checklist (Ochyra 

et al., 2003) but as a synonym of subsp. eurystomum in 

the alpine countries from which it was described. 

46. For synonymy of Pyramidula tetragona see Brugués, 

Casas & Belmonte (1998). 

47. For synonymy of Oedipodiella australis see Brugués 

(2003). 

48. Bryoxiphium madeirense was described from Madeira 

by Löve & Löve (1953) but ignored by Corley et al. 

(1981) and Corley & Crundwell (1991). It is listed by 

Eggers (1982), Düll (1992) and Dierssen (2001). 

49. Coscinodon humilis was described from Europe, but 

largely ignored by European authors although recognized 

in Japan; European plants lack capsules 

(Greven, 1995). 

50. For typification of Grimmia alpestris, see Muñoz 

(1997). 

51. According to Maier (2002a), Grimmia limprichtii is a 

synonym of G. tergestina. However, Greven (2000) 

treats it as distinct. 

52. Grimmia donniana var. curvula is homotypic with G. 

arenaria. 

53. Grimmia capillata, originally described from Sardinia, 

was generally treated as a variety of G. crinita during 

the 20th century. Both Greven (1995) and Muñoz & 

Pando (2000) recognize it at species rank. 

54. Grimmia curviseta was described from Tenerife by 

Bouman (1991). 


55. Grimmia dissimulata was described from Cyprus by 

Maier (2002b), who also listed localities from Turkey, 

Syria and southern and western Europe. 

56. Dryptodon incurvus is homotypic with Grimmia elatior. 

57. Grimmia funalis var. calvescens is homotypic with G. 

funalis f. epilifera J.E.Zetterst., a heterotypic synonym 

of G. funalis. 

58. Although the name Grimmia retracta remains to be 

typified, it is now treated as a synonym of G. lisae by 

Smith (2004). 

59. Grimmia muehlenbeckii and G. poecilostoma were 

included by Corley et al. (1981) in G. trichophylla 

and G. tergestina, respectively. Both Greven (1995) 

and Muñoz & Pando (2000) recognize them as species. 

60. Grimmia pulvinata var. obtusa is homotypic with G. 

pulvinata var. africana. 

61. The identity of Grimmia reflexidens, which was 

described from Chile, is controversial; Maier (2002a) 

and Ochyra (2004b) treat it as a species of Coscinodon. 

62. Grimmia grisea has been reported from Iceland by 

Greven (1998). This taxon was originally described 

from South Georgia (Cardot, 1906) after materials 

collected by Skottsberg. There are no original 

materials at PC, and all the available syntypes at S 

(Skottsberg original herbarium) and H-BR are conspecific 

with G. reflexidens. 

63. Grimmia subsulcata was considered a synonym of G. 

reflexidens by Muñoz (1998), although he later 

realized (Muñoz & Pando, 2000) that this typification 

was in error (the name is homotypic with G. alpestris 

var. microstoma Bruch & Schimp.). No type of the 

latter name could be found in BM (original 

Schimper’s herbarium) or any of the many herbaria 

to which materials were requested. Pending further 

developments the name is tentatively maintained 

under G. reflexidens. 

64. The status of Grimmia crassifolia is controversial. 

There are no capsules in the holotype, and thus it is 

impossible to decide if it belongs to G. poecilostoma or 

G. tergestina. Muñoz (1999) considered it a synonym 

of G. poecilostoma, but in a more recent paper it was 

placed under G. tergestina (Ignatova & Muñoz, 2005). 

65. Grimmia austrofunalis is an Australasian and South 

American taxon that does not grow in Europe. It is 

characterized by large gemmae (100–200 mm) formed 

on short stalks arising at dorsal costa base, whose 

liberation does not cause any damage to the leaves. 

Other characters, such as very large size for the genus 

(up to 8 cm) and shiny leaves, are more variable 

(Muñoz, 1999). 

66. The name Grimmia britannica, now treated by Smith 

(2004) as a form of G. trichophylla, is a new name for 

G. robusta Braithw., a later homonym for G. robusta 

Nees & Hornsch. (? Schistidium robustum (Nees & 

Hornsch.) H.H.Blom) and thus illegitimate. When 

Braithwaite (1872) published his new species, he 

mentioned four syntypes from different Scotland


localities, collected by Fergusson, Moore, Stirton and 

Hunt, which are, at least Fergusson’s, a mixture of 

several species. Not all possible syntypes have been 

located, and the name remains to be typified. 

67. Grimmia meridionalis, originally described as G. 

trichophylla var. meridionalis Müll.Hal. and raised to 

species rank by Maier (2002b), is listed here as a 

synonym of G. trichophylla, in accordance with the 

opinion of Muñoz (1999). 

68. Grimmia triformis was generally treated as a synonym 

or variety of G. donniana until revised by Muñoz 

(1998). 

69. Grimmia ungeri was originally described from Cyprus. 

Its presence in Europe is disputed by Greven (1995) 

but asserted by Muñoz & Pando (2000). 

70. Indusiella thianschanica was reported by Thériot 

(1918) from Dagestan, Russian Caucasus. 

71. Jaffueliobryum latifolium has been found in the 

Russian Caucasus (Kharzinov et al., 2005). 

72. Bednarek-Ochyra & Ochyra in Ochyra et al. (2003) 

divide Racomitrium into four genera, namely 

Racomitrium Brid., Niphotrichum (Bednarek-Ochyra) 

Bednarek-Ochyra & Ochyra, Codriophorus P.Beauv. 

and Bucklandiella Roiv. In the interests of nomenclatural 

stability, these subdivisions are recognized here 

at the rank of subgenus. 

73. Racomitrium hespericum was described from the 

Iberian Peninsula by Sérgio, Muñoz & Ochyra (1995). 

74. Racomitrium lusitanicum was described from Portugal 

by Ochyra & Sérgio (1992). 

75. Schistidium andreaeopsis (Müll.Hal.) Laz., known 

from Beringia, is probably not European. 

Schistidium marginale, S. pratense, S. scabrum and S. 

subflaccidium are mapped for Austria on the website 

‘Mooskartierung in Österreich’ http://www.sbg.ac.at/ 

bot/, on the basis of specimens determined by H.H. 

Blom (pers. comm.). These names are nomina nuda 

and cannot be included in the checklist although the 

mapped taxa are thought to be good species. 

76. Schistidium apocarpum s.l. has been comprehensively 

revised in northern Europe by Blom (1996). He 

subsequently wrote a substantial flora account of 

Schistidium (Blom in Nyholm, 1998) and has 

described additional species from elsewhere. Our 

treatment of Schistidium follows his flora account 

except where otherwise indicated. 

77. The name Schistidium alpicola (Hedw.) Limpr., nom. 

rejic., was for long applied to the plant now called S. 

platyphyllum. It is a synonym of Schistidium agassizii 

(Blom in Nyholm, 1998) and is now rejected. Blom 

(pers. comm. to Hill) points out that the author 

citation S. platyphyllum (Mitt.) Kindb., often used in 

North America, is incorrect because Kindberg did not 

recognize Schistidium as a genus. 

78. According to Ochyra et al. (2003), Schistidium 

helveticum (Schkuhr) Deguchi is a synonym of S. 

singarense and has priority. Blom (1996) was unable to 

study the lectotype of S. helveticum, which was not 

found at LAU. He has subsequently written (pers. 

comm.) ‘All evidence (including the protologue) 

points towards the taxonomic identity of S. helveticum 

and S. singarense, further strengthened by the article 

by Ochyra et al. (2003)’. Given that there is apparently 

no Schkuhr material at HAL, he concludes that if we 

accept the lectotypification by Deguchi, then we may 

use the name S. helveticum. Therefore the name S. 

helveticum is recommended here, even though a new 

lectotype to replace the lost type specimen chosen by 

Deguchi has not been designated. 

79. For protologues of Grimmia maritima and Dicranum 

scottianum, see Nelson (1997). 

80. Schistidium occidentale, a species previously known 

only from the mountains of southwestern North 

America, was found in the Sierra Nevada, southern 

Spain (Casas, Blom & Cros, 2001). 

81. Schistidium sinensiapocarpum is mapped for two 

localities in Austria on the website ‘Mooskartierung 

in Österreich’ http://www.sbg.ac.at/bot/, on the basis 

of specimens determined by H.H. Blom (pers. comm.). 

82. Schistidium spinosum was described from the Black 

Forest, with localities elsewhere in Germany and 

France (Blom & Lüth, 2002). 

83. Campylostelium pitardii was transferred from Grimmia 

by Maier (1998). 

84. Seligeria transylvanica was synonymized with 

Brachydontium trichodes by Ochyra (2004c). 

85. Subgenera of Seligeria are those of Vitt (1976). 

86. Seligeria galinae was described from the Urals by 

Mogensen & Goldberg (2002). 

87. Seligeria irrigata was raised to species rank by Ochyra 

& Gos (1992). 

88. Fissidens dubius var. mucronatus (Breidl. ex Limpr.) 

Kartt., Hedenäs & L.Söderstr. is included here in var. 

dubius. 

89. For information on Fissidens luisieri refer to Sérgio, 

Iwatsuki & Ederra (1997). 

90. Like Sérgio & Carvalho (2003) we include pending 

further investigation Fissidens polyphyllus var. newtonii 

Husn. in F. polyphyllus var. polyphyllus. 

91. Sérgio et al. (1997) provide a useful discussion of 

Fissidens serrulatus. 

92. Limbidia can be greatly reduced in many limbate 

species. Most taxa that are primarily or only 

characterized by their limbidia restricted to the 

vaginant lamina, e.g. Fissidens kosaninii Latzel, F. 

exiguus Sull. and F. viridulus var. intralimbatus 

(R.Ruthe) Düll, are probably just poorly limbate 

forms of limbate species. 

93. Pursell (2005) provides new insight on the characterization 

of Fissidens arcticus. 

94. The European concept of Fissidens bryoides and 

related species diverges considerably from that of 

North American authors, e.g. Crum & Anderson 

(1981), Pursell (2005). European floras emphasize

differences and recognize several species. North 

American authors stress the presence of intergrading 

forms and recognize only one species, F. bryoides 

Hedw. On both continents many specimens are easily 

identified as F. bryoides sensu stricto (with numerous 

axillary antheridial buds), F. viridulus (most antheridia 

terminal on dwarf male or longer plants) or F. incurvus 

(incurved capsules). Also on both continents sexual 

characters are more or less correlated to other 

characters (extension of the limbidium, shape of the 

leaf apex, sporophyte, etc.). Intergrading specimens 

are also known from both continents and are probably 

most common in southern and northern regions. 

Whether they are more common in North America 

is unknown. In the Netherlands most specimens can 

be easily identified. Variation in sexuality in European 

material of this complex has led to the description of 

many varieties and species. In this checklist, we adopt 

a conservative (from a European point of view) 

perspective, but omit F. exiguus Sull. and F. kosaninii 

Latzel, and reduce F. curnovii and F. incurvus to 

varietal rank. 

95. Cortini Pedrotti (2001) and Smith (2004) treat 

Fissidens bryoides var. caespitans at species level. It is 

here included as variety because, though extreme 

forms are clearly distinct from F. bryoides var. 

bryoides by red rhizoids, a thicker limbidium, inclined 

capsules and a cushion-like growth form, many 

collections cannot be identified with certainty. 

96. In a recent provisional publication Pursell (2005) 

recognizes Fissidens minutulus as a distinct species, 

closely related to, yet distinct from, F. crispus 

(including F. limbatus). We are not convinced that 

the two taxa are distinct in Europe and they are 

treated as synonyms here. 

97. Fissidens gymnandrus is treated by North American 

authors and Ignatov & Afonina (1992) as a synonym 

of F. bryoides. However it differs from F. bryoides 

sensu stricto in being polyoicous, having oblong leaves 

with a wide apex, and by its preference for woody 

substrates in regularly flooded localities. 

98. Fissidens jansenii was described by Sérgio & Pursell 

(2001). The resemblance of this species to Fissidens 

rufulus was overlooked by its authors. Both species 

have small cells and limbidia that reach the insertion 

of the dorsal laminae. F. jansenii has bistratose leaves; 

the leaves of F. rufulus are irregularly bistratose or 

unistratose. 

99. Fissidens microstictus is a relatively unknown species 

characterized by stems with numerous wide, elliptical 

to ovate leaves, broad, acute leaf tips and dorsal 

laminae that are narrowed below and typically reach 

the insertion. All leaves of a stem can be elimbate or 

limbate, or limbate and elimbate leaves may occur on 

the same stem, limbidia are found on the vaginant 

laminae and sometimes also in the middle of the 

dorsal laminae. It is closely related to F. arnoldii from 


which it differs in the broadly acute not acuminate leaf 

tips. 

100. Fissidens nobreganus resembles F. sublineaefolius 

which also has linear leaves and smooth cells. It 

differs from this species in having bud-shaped axillary 

perigonia and perigonia terminal on short axillary 

branches, by smaller, almost isodiametric laminal cells 

and limbidia that are restricted to the upper leaves of 

perichaetial plants. Fissidens sublineaefolius has terminal 

perigonia, larger, hexagonal laminal cells and 

well-developed limbidia on all laminae of all leaves. 

101. Fissidens perssonii was synonymized with F. ovatifolius 

by Dirkse, Bouman & Losada-Lima (1993). 

102. Fissidens pusillus is treated by North American 

authors as an expression of F. bryoides. 

103. European specimens identified as Fissidens exiguus are 

poorly limbate forms of either F. viridulus or of F. 

pusillus. Fissidens exiguus is treated by North 

American authors as a poorly limbate expression of 

F. bryoides. 

104. Fissidens sublimbatus was reported from the Canary 

Islands by Ros et al. (2001). 

105. Fissidens viridulus is treated by North American 

authors as an expression of F. bryoides. 

106. Bryum pallens, Dicranum polysetum, D. viridulum 

(Fissidens), Hypnum revolvens (Scorpidium) and 

Orthotrichum pumilum are Swartz names from before 

Species Muscorum that were validated in a book 

review (Anon., 1801); thus ‘ex anon.’. 

107. Fissidens viridulus var. incurvus is linked to var. 

viridulus by transitional forms. North American 

authors treat both taxa as expressions of F. bryoides. 

108. Re-examination of the isotype of Fissidens bambergeri 

in U showed that it is a poorly limbate form of 

F. viridulus. It differs from F. crispus (F. limbatus) 

with which it has been confused by larger, hardly 

bulging, less than twice as deep as wide, laminal 

cells. 

109. Fissidens marginatulus is not accepted by Ignatov & 

Ignatova (2003) and seems hardly distinct from F. 

viridulus. 

110. Fissidens serratus was reported as F. papillosus, new 

for the Canary Islands, by Dirkse, Bruggeman- 

Nannenga & Bouman (1991). Bruggeman-Nannenga 

& Pursell (1995) treated F. papillosus as a synonym of 

F. serratus. 

111. Ceratodon antarcticus Cardot and C. heterophyllus 

Kindb. have been reported from arctic and alpine 

parts of Europe. According to Ochyra (1998), C. 

antarcticus is merely a developmental phase of C. 

purpureus with very large mid-leaf cells and plane leaf 

margins. European records of C. heterophyllus are 

based on plants that lack capsules and therefore 

according to the account of Burley & Pritchard (1990) 

are not reliable. Nevertheless, the occurrence in cold 

parts of Europe of Ceratodon plants with rounded, 

entire leaf apices is remarkable. Cultivation experi-


ments and additional genetic data are needed to 

elucidate their status. 

112. The assignment of synonyms to the subspecies of 

Ceratodon purpureus follows Burley & Pritchard 

(1990). 

113. Ditrichum gracile was synonymized with D. crispatissimum 

by Allen (1994). 

114. Ditrichum punctulatum is a subantarctic species, 

known in Europe only from Madeira and the 

Azores. Records of D. difficile (Duby) M.Fleisch 

probably belong to this species. 

115. Pleuridium serrulatum Luisier & Dixon, treated as 

doubtful by Corley et al. (1981) but included in recent 

Spanish lists, is represented in Luisier’s herbarium 

(INA) by non-fruiting plants. It therefore remains 

doubtful. 

116. Rhamphidium purpuratum, with peristome resembling 

Saelania, is excluded from Pottiaceae by Zander 

(1993). Its leaves resemble Dicranella and its tubers 

resemble those of Trichodon (Arts, 1989). 

117. Trematodon perssoniorum, endemic to S. Miguel, 

Azores, is closely related to T. longicollis. It was 

described by Allorge (1951) on the basis of suggestions 

made by her husband (died 1944) and Thériot (died 

1947). She makes it absolutely clear that the description 

is newly written by her for the paper. In 

accordance with Article 60.11 of the International 

Code, the epithet perssonorum is corrected here to 

perssoniorum (noting that the form perssonii is used for 

‘of Persson’ in several bryophyte epithets). 

118. Amphidium tortuosum is a tropical montane species, in 

Europe only in Tenerife and Madeira, where it had 

been reported as A. curvipes. Synonymy follows 

Frahm et al. (2000). 

119. Cynodontium asperifolium is similar to C. fallax, with 

rather slight differences. It is known from the Ural 

Mountains (Ignatov & Ignatova, 2003). 

120. The basionym of Cynodontium polycarpon is 

Fissidens polycarpos, which appears incorrectly in the 

index of Species Muscorum (Hedwig, 1801) as F. 

polycarpus. 

121. Although Dichodontium palustre has been variously 

placed in Dicranella, Diobelon or Diobelonella, a 

molecular study by Stech (1999) supports its affiliation 

with Dichodontium; which seems reasonable in view of 

its leaf shape. 

122. The systematic position of Dicranoweisia is controversial. 

Ochyra (1993) argued that its peristome 

places it in Seligeriaceae. He also (in Ochyra 

et al., 2003) divided the European species into 

Dicranoweisia, which occurs in the Northern 

Hemisphere and Africa, and Hymenoloma, whose 

distribution is mainly in the Southern Hemisphere. 

We recognize the merits of his argument but prefer 

to a adopt a conservative position until further 

work has been done. 

123. Oncophorus elongatus, from Norway and Sweden, was 

elevated to the rank of species by Hedenäs (2005). 

124. Rhabdoweisia kusenevae Broth. was included by 

Corley & Crundwell (1991) but treated by Ignatov & 

Afonina (1992) as a synonym of R. crispata. 

125. The placement of Schistostega in Dicranales is 

controversial, but we follow Goffinet & Buck (2004) 

in this. 

126. Dicranella campylophylla, not listed by Corley 

et al. (1981), was reported for Madeira by Eggers 

(1982). 

127. Sections of Dicranum follow those of Nyholm (1987), 

with some nomenclatural changes due to Ochyra et al. 

(2003). 

128. Dicranum crassifolium was described from Portugal 

(Sérgio, Ochyra & Séneca, 1995). 

129. Dicranum transsylvanicum was described from 

Romania by Lüth (2002). 

130. Dicranum dispersum was described from Germany by 

Engelmark (1999). 

131. Dicranum undulatum Schrad. ex Brid. 1801 has 

priority over D. bergeri Blandow 1809. Dicranum 

undulatum Ehrh. ex F.Weber & D.Mohr 1803 is an 

illegitimate homonym. Although many recent 

European authors have avoided the name, it is current 

in North America and should also be used in Europe 

(Ochyra et al., 2003). 

132. Dicranum angustum and D. laevidens are treated as 

distinct species, following Hedenäs & Bisang (2004). 

133. An isotype of Dicranum sendtneri at S consists of a 

mixture of D. fuscescens and D. elongatum. Following 

Hedenäs & Bisang (2004), D. sendtneri is not 

recognized here as a distinct species. 

134. Dicranum canariense, often treated as a subspecies of 

D. scottianum, is known from the Canary Islands and 

Spain. 

135. Campylopus incrassatus was reported under the name 

C. eximius from Madeira by Koppe & Düll (1986), 

having been previously known under other names 

including C. dixonii Luisier. The synonymy of C. 

eximius and C. incrassatus is due to Frahm (1987). 

136. Dicranodontium subporodictyon is only known 

as sterile. Therefore its generic position is still 

doubtful. 

137. Leucobryum albidum is a North American species, in 

Europe only in the Azores. For differences between it 

and L. juniperoideum see Frahm (2005b). The distinction 

between L. albidum and L. juniperoideum is 

controversial; according to Vanderpoorten, Boles & 

Shaw (2003) they should be treated as synonyms. 

138. Octoblepharum albidum Hedw. was reported from 

Tenerife by Düll (1992) on the basis of a 19th century 

herbarium specimen. It is omitted from a recent list for 

the Canary Islands (Losada-Lima et al., 2001). If the 

record is genuine, the species had probably been 

introduced from the tropics.

139. Molecular phylogenies of Pottiaceae by Werner et al. 

(2004a, 2005b) place the European species in the three 

groups recognized here: Merceyoideae (Scopelophila), 

Trichostomoideae (including Splachnobryum, 

Ephemeraceae, Pleuroweiseae sensu Corley et al., 

1981, and some Barbula species, e.g. B. bolleana), 

and Pottioideae (including Pottieae, Cinclidotus and 

Barbuleae sensu Corley et al., 1981). Timmiella does 

not belong in Pottiaceae but is listed here until a better 

place can be found for it. Likewise, those Barbula 

species that belong in Trichostomoideae have been 

retained in Barbula until their correct generic position 

can be ascertained. 

140. Crumia latifolia (Kindb.) W.B.Schofield is known 

from the Caucasus but outside Europe. 

141. Anoectangium handelii is known from the Crimea. It is 

described by Zander & Weber (2005), correcting an 

earlier report by Zander (1977) who had treated it as a 

synonym of Molendoa sendtneriana. 

142. Ephemerum hibernicum was described from Ireland by 

Holyoak & Bryan (2005). 

143. Holyoak has examined much material of Ephemerum 

serratum and E. minutissimum, and finds that in both 

species occasional plants have leaves with a weak 

nerve. He has not seen the type of E. serratum var. 

praecox, which could be a synonym of either of these 

taxa. 

144. Ephemerum spinulosum has been found in Ireland and 

the Basque region of Spain (Holyoak, 2001; Infante & 

Heras, 2005). 

145. Author citation for Eucladium verticillatum follows the 

typification by Ochyra & Zijlstra (2005). 

146. Gymnostomum mosis (Lorentz) Jur. & Milde was 

reported from Spain by Martínez-Sánchez, Ros & 

Guerra (1991), but later Cano, Ros & Guerra (1994) 

described the new species Gymnostomum lanceolatum 

based on this material. 

147. Gymnostomum aeruginosum var. obscurum was 

described by Guerra (2004). 

148. Spelling of species epithet in Hymenostylium recurvirostrum 

follows Hedwig (1801) not Crosby et al. 

(1999). 

149. Pleurochaete malacophylla (Müll.Hal.) Broth. is listed 

for Turkey by Kürschner & Erdag˘ (2005). 

150. Splachnobryum delicatulum is treated by Arts (2001) as 

a synonym of S. obtusum. Splachnobryum obtusum was 

not included in the main list of Corley et al. (1981), 

because these authors considered that its occurrences 

were not sufficiently natural. However, it occurs in the 

open in Hungary and the Azores, and is accordingly 

listed here. 

151. Tortella arctica (Arnold) Crundw. & Nyholm has been 

reported from Svalbard, but the records refer to 

Trichostomum arcticum (Frisvoll & Elvebakk,1996). 

152. Tortella alpicola was reported new for Europe by 

Otnyukova et al. (2004). 


153. Tortella bambergeri was not recognized as a good 

species by Corley et al. (1981), but is generally 

accepted by bryologists in central Europe and also 

by us. 

154. Tortella densa is treated as a variety T. inclinata var. 

densa, following Eckel (1998). 

155. Tortella limbata has been refound on the Canary 

Islands. Dirkse et al. (1993) compared their collection 

with the holotype and noted the distinctive denticulate 

leaf border, composed of long narrow cells, running 

nearly to the leaf apex. 

156. Tortella limosella is known only from the Scottish 

type-locality, and has not been refound since its 

discovery in 1906. 

157. Tortella cirrifolia was synonymized with T. nitida by 

Sjögren (2001). 

158. Molecular studies by Werner et al. (2005b) show that 

the genus Trichostomum as listed here is heterogeneous, 

with T. brachydontium and T. crispulum nested 

within Weissia and T. triumphans very close to 

Pottiopsis caespitosa. Trichostomum connivens 

(Lindb. ex Broth.) Paris is recorded from the 

Caucasus but is not known from Europe. 

Trichostomum contortum (Kunze) Sérgio is probably 

a synonym of T. brachydontium, although Sérgio 

(1985) thought that it was a good species. 

159. For synonymy of Hyophila treleasei, see Sollman 

(1993). 

160. Weissia leptocarpa Schimp. ex Besch. is treated by 

Sérgio & Carvalho (2003) as a doubtful species, but is 

accepted for Turkey by Kürschner & Erdag˘ (2005), in 

spite of the fact that Frey & Kürschner (1991) had 

treated it as doubtful and illegitimate. 

161. Weissia mittenii (Bruch & Schimp.) Mitt. is omitted, 

because it is designated above as a hybrid 

W.6mittenii (Bruch & Schimp.) Mitt. emend. 

A.J.E.Sm. Smith (pers. comm. to Hill) notes that W. 

mittenii has malformed capsules and abnormal spores. 

Following Werner et al. (2005b), Astomum is included 

in Weissia. 

162. A molecular study by Werner et al. (2004b) supports 

the species status of Weissia wimmeriana. 

163. Acaulon mediterraneum is treated by many authors as 

a variety of A. muticum, to which it is undoubtedly 

close. 

164. Aloina humilis was described from the Canary Islands 

by Gallego et al. (1998). 

165. Aloina obliquifolia is accorded species rank following 

Gallego et al. (1999). 

166. The Macaronesian Barbula semilimbata Dixon & 

Luisier was listed by Eggers (1982) but is a poorly 

known taxon, not recorded since it was described. 

167. Barbula convoluta var. sardoa is treated by some at 

species rank. Frahm & Ahmed (2004a) suggested the 

name B. sardoa. However, the epithet sardoa is correct 

at the varietal level but superfluous at the species level.


If species rank is used, the correct name is B. 

commutata (see Frahm & Ahmed, 2004b). 

168. Although Corley & Crundwell (1991) did not accept 

the synonymy of Bryoerythrophyllum campylocarpum, 

Sollman (1990), Zander (1993) and Sérgio & Carvalho 

(2003) all accept it, as we do here. 

169. Cinclidotus bistratosus Kürschner & Lüb.-Nestle and 

C. nyholmiae Çetin have been described from 

Anatolia. 

170. Cinclidotus confertus Lüth was described from northwest 

Greece. Dr Alicia Ederra, who treated 

Cinclidotus for the Flora Briofítica Ibérica, has written 

(pers. comm. to Hill) ‘I had the opportunity to see the 

holotype of Cinclidotus confertus from STU. It is, 

without any doubt, a Cinclidotus; its habit is very 

similar to C. riparius, but, as noted by Lüth, the 

peristomes are different. So, I think they must be 

closely related species.’ She recommends retaining it in 

the list, although it has not been found elsewhere. 

171. Cinclidotus vivesii was described by Ederra in Ederra 

& Guerra (2005). 

172. Crossidium davidai was reported new for our area 

from the Canary Islands by Dirkse & Bouman (1995b) 

and Stern (1995). 

173. Crossidium geheebii was reported from Italy, new to 

Europe, by Privitera & Puglisi (2000a). 

174. Crossidium laevipilum was described in 1931 from 

North Africa and subsequently found in Spain (Casas, 

Cros & Brugués, 1993; Guerra et al., 1993). 

175. Crossidium laxefilamentosum was reported new to 

Europe by Pócs et al. (2004). 

176. Dialytrichia fragilifolia was raised to species rank by 

Lara (2005). 

177. A molecular study by Werner et al. (2005a) shows that 

previous sectional divisions of Didymodon cannot be 

maintained, but does not yet provide enough detail to 

demarcate new sections. 

178. Didymodon reedii H.Rob. (D. tectorum (Müll.Hal.) K. 

Saito), listed by Corley & Crundwell (1991), was 

erroneously reported from Britain (Blockeel & Smith, 

1998). 

179. According to Sérgio & Jiménez Fernández (2003), 

Didymodon soaresii is a non-fruiting species of 

Tortula, of uncertain species. 

180. The synonymy of Didymodon asperifolius, D. fallax, 

D. giganteus, D. maximus, D. spadiceus and D. 

tophaceus follows Jiménez et al. (2005b). 

181. Didymodon australasiae synonymy follows Jiménez 

et al. (2005a). 

182. For description of Didymodon bistratosus, see Hébrard 

(1994). 

183. Didymodon brachyphyllus is reported from Iceland 

(Jóhannsson, 2003). 

184. Didymodon erosus was described by Jiménez et al. 

(2004). 

185. Didymodon lamyanus, from France, has been omitted 

from recent European lists. It was studied by Werner 

et al. (2005a). Its closest relatives appear to be D. 

insulanus, D. nicholsonii and D. vinealis. 

186. Didymodon mamillosus is treated by Jiménez (2004) as 

a synonym of D. rigidulus. 

187. Didymodon sicculus was described by Cano et al. 

(1996). 

188. For synonymy of Didymodon subandreaeoides see 

Kučera & Köckinger (2000). 

189. Didymodon umbrosus synonymy follows Jiménez et al. 

(2005a). 

190. According to Ochyra (1998), Hennediella antarctica 

(A˚ ngstr.) Ochyra & Matteri is a synonym of H. 

macrophylla and has priority. However, there appear 

to be some differences, and we have not changed the 

name applied to European plants pending further 

study of variation in the genus. 

191. Our circumscription of the genus Microbryum follows 

Zander (1993). 

192. Phascum piptocarpum is treated as a synonym of 

Microbryum curvicollum following Ros, Werner & 

Rams (2005). 

193. The synonymy of Microbryum davallianum and M. 

starckeanum follows Ros et al. (1996). 

194. Microbryum fosbergii was raised to species rank and 

synonymized with Pottia6andalusica by Ros et al. 

(2005). 

195. The basionym of Microbryum starckeanum is Weissia 

starckeana, named by Hedwig (1801) after Johan 

Christian Starke. Although Hedwig’s spelling is not 

apt, Article 60.1 of the International Code (Greuter et 

al., 2000) requires that it shall be retained. 

196. Pseudocrossidium obtusulum, originally described from 

Sweden, is widespread in North America, from which 

P. revolutum is absent. Reasons for treating it as a 

distinct species are given by Eckel, Hoy & Elliot (1997) 

and Zander (2002). 

197. Pseudocrossidium replicatum was found in Italy, new 

for the European moss flora (Privitera & Puglisi, 

2000b). 

198. Pterygoneurum compactum was described from Spain 

by Cano, Guerra & Ros (1994). 

199. Pterygoneurum crossidioides, originally described from 

the Dead Sea area of Israel, was found in Hungary 

(Pócs, 1999). However, Erzberger & Papp (2004) 

regard the record as somewhat doubtful. 

200. Pterygoneurum papillosum was described by Oesau 

(2003). 

201. Pterygoneurum squamosum was described from Spain 

by Segarra et al. (1998). 

202. Syntrichia treatment follows Gallego (2005). 

203. Syntrichia bogotensis (Hampe) R.H.Zander has been 

found in Madeira (Gallego, Cano & Sérgio, 2005). 

204. Some recent authors such as Vanderpoorten (2001) 

treat Syntrichia calcicola as a synonym of S. ruralis. 

205. According to a revision by Gallego et al. (2002a), 

Spanish material ascribed to Syntrichia pseudohandelii 

is S. caninervis var. caninervis. These authors treat S.

pseudohandelii as a synonym of the non-European S. 

caninervis var. pseudodesertorum (Vondr.) 

M.T.Gallego, which is found in western Asia, including 

Turkey. 

206. For the status of Syntrichia caninervis var. abranchesii 

refer to Gallego et al. (2002a). 

207. Syntrichia caninervis var. astrakhanica was described 

by Ignatov, Ignatova & Suragina (2002). 

208. Syntrichia glabra was described by Frahm & Gallego 

(2001). 

209. Following Gallego, Cano & Guerra (2004), no 

infraspecific taxa are recognized within Syntrichia 

laevipila, which is synonymized with S. pagorum, 

Tortula laevipilaeformis and T. saccardoana. 

210. Syntrichia minor was recognized at species rank by 

Gallego et al. (2000). 

211. According to Ochyra (1994), Tortula intermedia De 

Not. 1838 is a synonym of Tortella humilis, making 

Tortula intermedia (Brid.) Berk. 1863 an illegitimate 

homonym. 

212. According to Gallego (2005), Tortula papillosa var. 

meridionalis Warnst. should not be recognized as a 

distinct variety. 

213. According to Gallego (2005), Syntrichia princeps var. 

parnassica (Schiffn.) Podp. should not be recognized 

as a distinct variety. 

214. Author citation of Syntrichia ruralis var. ruraliformis 

follows Gallego (2005). 

215. Syntrichia subpapillosissima was recognized at species 

rank by Gallego et al. (2002b). These authors treated 

Tortula ruralis var. submamillosa W.A.Kramer as a 

synonym. 

216. Tortula buyssonii (H.Philib.) Broth., from the 

Auvergne, France, is a little-known taxon that has 

apparently not been found since the 19th century. 

217. Tortula caucasica Lindb. (Pottia caucasica (Lindb.) 

Paris) is known from Georgia but not from Europe. 

Tortula grandiretis Broth. is listed for Turkey by 

Kürschner & Erdag˘ (2005). 

218. Tortula ampliretis, described from the Canary Islands 

in a paper by Crundwell, During & Long (1978), was 

inexplicably not mentioned by Corley et al. (1981). 

219. Tortula bolanderi and T. inermis should be in Tortula 

not Syntrichia (Werner et al., 2003a). 

220. Tortula israelis, described from Spain as T. muralis 

var. baetica, was raised to species rank by Guerra & 

Ros in Guerra, Ros & Carrión (1992) and subsequently 

synonymized with T. israelis by Cano, Guerra 

& Ros (1996). 

221. According to Cano (2004a), European plants attributed 

to Crossidium seriatum Crum & Steere should be 

referred to Tortula brevissima. 

222. The synonymy and nomenclature of Tortula hoppeana 

follow Ochyra (2004a). 

223. For the synonymy of Desmatodon meridionalis see 

Sérgio & Granzow de la Cerda (2001). 


224. According to Sérgio & Carvalho (2003), the type 

material of Pottia cuneifolia corresponds to P. pallida. 

Therefore, P. pallida and Tortula zoddae, which is a 

new name for P. cuneifolia in Tortula, are included in 

the synonymy of T. pallida. 

225. Zander (1993) provided the new name Tortula 

rhodonia for Desmatodon wilczekii, because the epithet 

wilczekii is not available in Tortula. 

226. Cano, Werner & Guerra (2005) raised Tortula subulata 

var. angustata with bistratose border to species rank, on 

the basis of T. angustata Lindb., and provided the new 

name T. schimperi, because the epithet angustata is not 

available at species rank in Tortula. The other varieties 

of T. subulata were sunk in a single variable species. 

227. According to Cano (2004a, b), Tortula marginata 

subsp. limbata is a synonym of T. solmsii. 

228. Tetraplodon urceolatus is a good species which has 

been misunderstood in Scandinavia, where it has been 

confused with forms of T. mnioides. 

229. Orthotrichum urnaceum Müll.Hal. was signified for 

Europe by Lewinsky-Haapasaari (1995) but the 

locality is in Armenia. 

230. Synonymy of Orthotrichum limprichtii follows 

Lewinsky-Haapasaari in Nyholm (1998). 

231. Orthotrichum casasianum is reported only from northern 

Spain (Mazimpaka et al., 1999). 

232. Orthotrichum crenulatum was reported as O. 

flowersii Vitt from the French Alps (Boudier & 

Pierrot, 1992). 

233. Orthotrichum handiense is an endemic from 

Fuerteventura, Canary Islands (Lara et al., 1999a; 

Lara, Garilleti & Mazimpaka, 2003). 

234. Orthotrichum hispanicum was described from Spain by 

Lara, Garilleti & Mazimpaka (2000). 

235. Orthotrichum macrocephalum was described from 

Spain by Lara, Garilleti & Mazimpaka (1994). 

236. For synonymy of Orthotrichum lewinskyae see Draper 

et al. (2003). 

237. Orthotrichum schimperi is recognized as a species in 

accordance with Cortini & Lara in Cortini Pedrotti 

(2001). 

238. For synonymy of Orthotrichum rogeri var. defluens, 

see Garilleti, Lara & Mazimpaka (2002). 

239. For synonymy of Orthotrichum australe, see Lewinsky 

(1993). 

240. Orthotrichum vittii was described from Spain by Lara 

et al. (1999b). 

241. For synonymy of Orthotrichum arcangelianum see 

Lara et al. (2002) and Lewinsky (1993). 

242. Orthotrichum shawii has been reinstated as a distinct 

species (Mazimpaka et al., 2000). 

243. Orthotrichum ibericum was described from Spain and 

Portugal by Lara & Mazimpaka (1993). 

244. For the synonymy of Orthotrichum caucasicum, see 

Lewinsky (1993). 

245. We have excluded Orthotrichum elegans Schwägr. as a 

European species because Nordic material named as


O. speciosum var. elegans (Schwägr.) Warnst. is 

referable to O. speciosum. For differences between 

these two taxa see Vitt & Darigo (1997). 

246. Orthotrichum speciosum var. brevisetum is known from 

Spain & Sicily; see Draper et al. (2003). 

247. Orthotrichum tortidontium was described from 

Morocco and Spain by Lara, Garilleti & Mazimpaka 

(1996). 

248. Orthotrichum vladikavkanum was described from the 

Russian Caucasus by Venturi (1887). It was listed in 

Podpěra’s (1954) Conspectus, but the Caucasus was 

assigned to Asia by Corley et al. (1981). For the 

present enumeration, the Russian part of the Causasus 

is now again included in Europe. 

249. Corley et al. (1981) included Ulota macrospora in U. 

rehmannii, but the two species are distinguished 

here, following Boudier & Pierrot (1996) and Sauer 

(1998). 

250. The inclusion of Zygodon forsteri in Codonoblepharon 

is controversial (Goffinet and Vitt, 1998; Matcham & 

O’Shea, 2005). In our opinion it is best placed in 

Zygodon. Casas et al. (1985) considered Z. madeirensis 

to be close to Z. forsteri; it is treated here as a 

synonym following Düll (1992). Further revision is 

needed to assess its identity. 

251. Zygodon sibiricus was reported from the European 

side of the Urals by Ignatov & Ignatova (2003). 

252. Zygodon stirtonii is recognized at species rank following 

Smith (2004). 

253. Hedwigia ciliata var. viridis Bruch & Schimp. is listed 

by Casas (1991), but Hedenäs (1994) was unable to 

trace the type specimen, so it is uncertain whether the 

variety belongs to H. ciliata or H. stellata. 

254. Hedwigia ciliata var. leucophaea was illustrated and 

mapped by Hedenäs (1994). 

255. Hedwigia stellata was described from Sweden by 

Hedenäs (1994). 

256. Anacolia laevisphaera is reported from Europe in an 

abstract by Quandt & Frahm (2004). It is a tropical 

montane species and German specimens from the 

Mosel region had been incorrectly identified as 

Bartramia stricta. 

257. Anacolia menziesii was found new to Europe in the 

provinces of Almería and Granada in southern Spain 

(García-Zamora et al., 1998). Its distinctness from A. 

webbii was confirmed by molecular data (Werner et 

al., 2003b). 

258. Bartramia pomiformis var. elongata Turner and B. 

pomiformis var. heteromalla (Brid.) Müll.Hal. are 

treated with great scepticism by Nyholm (1998), and 

var. elongata is treated as a synonym of var. 

pomiformis by Smith (2004). These varieties are 

accordingly excluded from the checklist. 

259. Nomenclature for Bartramia section Pyridium (section 

Vaginella Müll.Hal., nom. illeg.) follows Ochyra et al. 

(2003). 

260. Bartramia breviseta was originally described from 

Finnmark, but was treated as a synonym of B. 

ithyphylla in many European checklists. Fransén 

(2004) has demonstrated that it is distinct, differing, 

for example, in the lack of an inner peristome. 

261. Bartramia ithyphylla var. strigosa (Wahlenb.) Hartm. 

was mapped by Söderström (1996). However, Fransén 

(2004) treats it as a synonym of var. ithyphylla, and it 

is accordingly not recognized here. 

262. Philonotis marchica var. laxa, described from 

Switzerland and Germany as P. laxa Limpr., is listed 

by Casas (1991) but is no longer recognized in German 

and Swiss lists. 

263. Philonotis uncinata (Schwägr.) Brid. is listed for the 

Azores by Gabriel et al. (2005). 

264. Molecular data now leave no doubt that the genera 

Anomobryum, Brachymenium, Plagiobryum and 

Rhodobryum cannot be maintained with the circumscription 

used by Corley et al. (1981) since most are 

ingroups within Bryum sensu lato (e.g. Pedersen, Cox 

& Hedenäs, 2003; Pedersen & Hedenäs, 2005). The 

molecular data also show that there have been 

multiple instances of convergent evolution of morphological 

characters in Bryaceae, so that classifications 

based exclusively on the morphological 

characters currently known are unsound. Hence, for 

example, the genus Rosulabryum proposed by Spence 

(1996) is not adopted here because molecular data 

imply it is polyphyletic (Pedersen et al., 2003). A start 

has been made on subdividing Bryum into segregate 

genera (e.g. Imbribryum N.Pedersen) on the basis 

of molecular data from multiple loci used in combination 

with morphological data (Pedersen, 2005; 

Pedersen & Hedenäs, 2005), but their treatment 

cannot yet be applied to European species as a 

whole because molecular data are not available 

for many species. The generic treatment of Corley 

et al. (1981) is therefore used here in an interim 

arrangement. 

265. Loeske (1932 [1933]) long ago interpreted Anomobryum 

leptostomoides as fertile A. concinnatum, and 

numerous fertile specimens examined by Holyoak 

from its type-locality at Mont Dore (France) 

consistently show the gametophytic characters of 

concinnatum. This is therefore the only European 

locality at which A. concinnatum has been found 

with capsules, and it is significant because sporophytes 

of A. leptostomoides differ from those of A. 

julaceum in the erect not inclined to cernuous 

capsules and their greatly reduced peristome, as 

described in detail by Shaw & Fife (1984). 

Furthermore, similar capsules have been described 

from Michigan (USA), again from plants with 

gametophytes indistinguishable from those of A. 

concinnatum. Shaw & Fife (1984) were ‘inclined to 

recognize A. leptostomoides at the specific level’ but 

hesitated to do so because they thought that the

usual non-fertile plants would be impossible to name 

with certainty. However, European bryologists have 

long recognized A. concinnatum without great 

difficulty from non-fertile material, so this form is 

surely best treated at species rank as advocated by 

Loeske. 

266. Sérgio, Cros & Brugués (1996) re-examined Anomobryum 

lusitanicum in detail, retaining it in the genus 

Anomobryum. 

267. Brachymenium commutatum is known in Europe only 

from the Sierra Nevada. It was first reported by 

Thériot (1932) as B. commutatum var. hispanicum 

Thér. and rediscovered in 1997 (Brugués et al., 2003). 

European plants are sterile and less robust than var. 

attenuatum Thér. & Trab., which was described from 

the Hoggar mountains, Algeria. Small forms are also 

found in the Hoggar (Thériot, 1932). Ochi (1972, 

1973) did not examine any of the type material of B. 

commutatum during his review of African Bryoideae. 

However, a specimen assigned to B. commutatum var. 

attenuatum from Sudan and material from Eritrea 

were reidentified by him as the widespread B. exile 

Dozy & Molk. It appears very likely that B. 

commutatum should be placed as a synonym of B. 

exile. 

268. Brachymenium notarisii was placed in the genus 

Haplodontium for many years, within which it is the 

type of subgenus Ateleobryum (Mitt.) Broth. (Wijk, 

Margadant & Florschütz, 1962). It was transferred 

from Haplodontium to Brachymenium by Shaw (1987), 

who commented that ‘it certainly does not belong in 

the Mielichhoferioideae’. 

269. Podpěra (1954) identified Mielichhoferia paradoxa 

with ‘Bryum splachnoides C. M.’, i.e. B. cellulare. 

However, Shaw (1987) reidentified it as a distinct 

species known only from Macedonia that he placed in 

Brachymenium, an arrangement that was followed by 

Corley et al. (1991). The recent checklist of mosses of 

the former Yugoslavia (Pavletic, Martincic & Düll, 

1999) continues to treat Brachymenium paradoxum 

erroneously as Bryum cellulare. 

270. Brachymenium philonotula is known in our area only 

from Madeira (Rocha da Pena, 1928, leg. C. Barreto, 

S). According to Ochi (1972) it is perhaps conspecific 

with B. exile Dozy & Molk. 

271. Bryum pamirense H.Philib. ex Broth. was reported 

new for Europe from Komi Republic in northern 

Russia (Zheleznova & Shubina, 1998). However, 

Ignatov & Afonina (1992) marked it with ?, to denote 

a ‘particularly poorly known and putatively ill-defined 

species’. Bryum pamirense is apparently close to B. 

savicziae and B. uliginosum (Savicz-Ljubitzkaja & 

Smirnova, 1970), but differs from both of these in 

being synoicous rather than autoicous. Like B. 

savicziae, it has the cilia of the endostome well 

developed and appendiculate, whereas B. uliginosum 


has short cilia. It might be a form of B. uliginosum or 

possibly a hybrid. 

272. Bryum savicziae Schljakov was described from the 

Kola peninsula in Russia in 1951 but overlooked by 

Index Muscorum (Wijk, Margadant & Florschütz, 

1959, 1969). It is known only from the type-locality. It 

resembles B. uliginosum but has the cilia of the 

endostome well developed and appendiculate. Spore 

sizes of 19–31 mm given by Savicz-Ljubitzkaja & 

Smirnova (1970) seem rather variable and perhaps 

this suggests a hybrid is involved. Ignatov & Afonina 

(1992) also reported B. ekstamii L.I.Savicz, B. 

labradorense H.Philib., B. umbratum I.Hagen and B. 

zemliae Arnell & Jaderh. from the north of European 

Russia. These little-known species require further 

study. 

273. Bryum vermigerum Arnell & C.E.O.Jensen was 

discovered at Hardanger, Norway in 1915 and 

1916; it is otherwise known only by a recent report 

from Iceland. Bryum vermigerum was treated as a 

valid species by Nyholm (1993) and its combination 

of characters is undoubtedly different from that in 

any other European Bryum species. Nevertheless, 

Nyholm reported an extraordinarily wide range of 

spore sizes (10–20 mm) and, although one capsule 

examined by Holyoak (at S) contained spores of 

less variable size (10–14 mm), these included a 

proportion of small shrunken and presumably 

abortive spores. The rarity of B. vermigerum, its 

peculiar characters and the variable and sometimes 

abortive spores may suggest that an interspecific 

hybrid is involved. 

274. Bryum colombii Meyl. and B. mesodon J.J.Amann are 

listed in the current Swiss list http://www.bryolich.ch/, 

but their identity is uncertain. Bryum murmanicum 

Broth. is on the Red List for the Murmansk Oblast of 

Russia as Data Deficient; its identity is also uncertain. 

Bryum geheebii Müll.Hal. was treated as probably a 

synonym of B. alpinum by Corley et al. (1981), as a 

species by Düll (1985), but very doubtful by Düll 

(1992); its identity is uncertain. The identity of B. teres 

Lindb. is also uncertain; it was treated by Düll (1985) 

as a synonym of B. nitidulum (here treated as a 

synonym of B. intermedium), but this synonymy was 

questioned by Nyholm (1993). 

275. Bryum gerwigii (Müll.Hal.) Limpr. is known from its 

type-locality on limestone rocks above the Rheinfalles 

near Schaffhausen in Switzerland and from a few 

other reports from central Europe. As with other 

submerged Bryum, it is apparently so greatly modified 

by growth in water (and perhaps by low light levels) 

that it is difficult to judge which species was its 

immediate progenitor. This uncertainty provides 

insufficient reason for treating it as a valid species. 

Bryum gemmiparum seems closest in view of the leaf 

shape, occurrence of bulbils and the habitat.


276. Crundwell (1970) recognized the widespread synoicous 

forms of Bryum algovicum as var. rutheanum and 

the less common autoicous ones as var. algovicum (of 

which he treated var. compactum as a synonym). 

Unpublished studies by Holyoak suggest that autoicous 

plants appear to occur mainly if not entirely on 

high ground in the Alps, whereas synoicous plants 

occur throughout the lowlands, from central Europe 

north to Svalbard. The autoicous plants are often 

small and densely tufted, they sometimes have narrow 

capsules, and the endostome processes have narrow 

perforations. They might merit treatment as a separate 

species rather than merely as a variety. 

277. Bryum apiculatum has a Pantropical range extending 

into temperate areas, with a single confirmed record in 

our region, from Tenerife (Ochi, 1972). The synonymy 

of B. apiculatum follows Ochi (1994). 

278. Taxonomic treatment of several Bryum species follows 

Holyoak (2004). 

279. Bryum veronense was treated as a subspecies of B. 

argenteum by Podpěra (1954) and Urmi (1987), as ‘a 

weak species’ by Düll (1992) who noted that verification 

is needed for most specimens, and as a distinct 

species allied to B. argenteum by Nyholm (1993). 

Some of the material referred to B. veronense is a 

modified form of B. argenteum, including plants with 

a proportion of achlorophyllose leaf tips that 

undoubtedly intergrade with that species, and otherwise 

typical B. veronense may have the papillose 

rhizoids of B. argenteum. However, other material 

placed as B. veronense in herbaria differs in various 

respects and may comprise similarly reduced, rather 

featureless, modified forms of other Bryum species, 

e.g. diminutive B. pallens. 

280. Longton (1981) studied intraspecific variation in 

morphology and physiology of Bryum argenteum on 

a global scale by means of cultures grown on agar 

under standardized laboratory conditions. He established 

that var. lanatum is of doubtful taxonomic value 

because the excurrent nerve was not maintained in 

cultivation by clones originating in Canada and 

Hawaii. Ochi (1994) also noted ‘gradations with the 

typical variety’. Nevertheless, lanatum was treated as a 

distinct species by Frahm (2002) and Spence & 

Ramsay (2002), on the basis of its longer acuminate 

leaf point, excurrent nerve and differing habitat 

preferences, but without any mention of Longton’s 

detailed study. Unless better evidence for the genetic 

distinctness of lanatum can be found it should be 

dismissed as an inconstant form of B. argenteum. 

281. Bryum oblongum and B. blindii are treated as separate 

species based on the detailed study by Shaw (1981). 

282. Bryum bornholmense was redescribed by Crundwell & 

Whitehouse (2001). 

283. Corley et al. (1981) suggested that Bryum caespiticium 

is probably an aggregate species, within which B. 

comense and B. kunzei may be good segregates. The 

latter is treated as a valid species here, but the case for 

maintaining B. comense appears weak. Although Düll 

(1985) listed B. comense as a species, he later (Düll, 

1992) expressed doubt. Nyholm (1993) treated it as a 

synonym of B. caespiticium, as we do here. 

284. Demaret & Wilczek (1982) examined the types of B. 

canariense and B. provinciale and concluded that they 

represented different species, but Corley et al. (1991) 

did not follow them because ‘their work was based too 

rigidly on the study of types alone’. Studies by 

Holyoak support the conclusions of Ochi (1972), 

who noted that ‘The form called B. provinciale is 

synoicous, but there are also dioicous plants which 

have no other clear-cut differences from such synoicous 

ones. These two forms should be better included 

in a single taxon’. 

285. Bryum caucasicum is known in Europe only from the 

holotype from the Caucasus (S). It was transferred 

from Mielichhoferia to Bryum by Cox & Hedderson 

(2003). 

286. Bryum demaretianum was described from Belgium by 

Arts (1992). 

287. Treatment of the Bryum bicolor complex follows 

Holyoak (2003), who described B. dyffrynense as a 

new species from Britain. 

288. Bryum kunzei, Pohlia acuminata and P. polymorpha 

were described by Hornschuch (1819), not by Hoppe 

& Hornschuch as commonly stated. 

289. Bryum miniatum is a predominantly North American 

species known in Europe only from the Faeroe 

Islands. 

290. Bryum minii was mainly overlooked until the paper by 

Sérgio et al. (1999). 

291. Bryum muehlenbeckii is probably best regarded as a 

form of B. alpinum that grows where intermittently 

submerged in cold water, since a substantial proportion 

of European specimens intergrade with that 

species (Holyoak, unpublished). Some specimens have 

the characteristic leaves of both B. muehlenbeckii and 

B. alpinum on different parts of the same stem, 

apparently as a result of growth under wet or dry 

conditions, respectively. 

292. Bryum neodamense is almost certainly a recurrent 

form of B. pseudotriquetrum, since intermediate plants 

occur intermixed with most populations of neodamense, 

often in large numbers (Holyoak, in preparation). 

Occurrence of the neodamense morphotype 

appears to be closely associated with intermittent 

flooding by hard water, most often on lake shores, but 

also in basic fens and dune-slacks. Detailed morphological 

analyses and molecular studies have been 

carried out (Holyoak & Hedenäs, in preparation). 

293. Bryum aeneum was later emended to B. oeneum; see 

Wijk et al. (1959). 

294. Zolotov (2000) treats the polyoicous (partly synoicous) 

B. lonchocaulon as a species distinct from the 

autoicous B. pallescens. However, he examined few

autoicous specimens and the differences he reported 

from B. lonchocaulon (other than in sexuality) appear 

inconstant in material from Britain. 

295. Bryum bimum may merit species rank since it differs 

from B. pseudotriquetrum in being synoicous rather 

than autoicous, perhaps in having twice as many 

chromosomes (Smith, 2004), and in average dimensions 

of the leaf lamina cells and thickness of their cell 

walls (Demaret & Empain, 1985; Zolotov, 2000). 

However, they apparently do not differ much, if at all, 

in ranges, frequency or ecology, and preliminary 

molecular data (Hedenäs, pers. comm. to Holyoak) 

suggest they are very close. 

296. Bryum salinum differs from B. archangelicum in only a 

few characters; a few intermediate specimens are 

known. 

297. Bryum schleicheri author citation follows Geissler 

(1985). 

298. Wijk et al. (1959) cited the original description of 

Bryum torquescens as Bruch ex De Not., Syllab. 

Musc., no. 163, 1838. Syed (1973) found this reference 

to be wrong and he discounted ‘Fl. Sard. Exs., 1828’, 

given by other authors because he suspected it is a 

nomen nudum. The original description is therefore 

taken as that by Bruch & Schimper (Bryol. Eur., 4 p. 

119, pl. 20, 1839 (fasc. 6–9, Mon., p. 49, pl. 20)) partly 

because this was the earliest description found by Syed 

and partly because they refer to it as ‘Diese neue Art’. 

299. Arts, Crundwell & Whitehouse (1995) showed that 

Bryum pyriferum described from Tenerife is conspecific 

with B. valparaisense, which was originally 

described from Chile but is now also known from N. 

America, Africa and Portugal. 

300. Hedwig (1801) used the spelling zierii twice, but the 

name Bryum zierii commemorates John Zier as was 

pointed out by Crum & Anderson (1981). More recent 

authors who amend it to zieri are therefore correcting 

it according to Article 60.11 of the International Code 

(Greuter et al., 2000). 

301. Touw (1984) gave reasons for rejecting the name 

Rhodobryum spathulatum in favour of R. ontariense. 

302. The large family Mniaceae of Goffinet & Buck (2004) 

is here divided following Koponen (1988) into the 

three families traditionally included in Mniaceae and 

the remainder, for which the only available name is 

Mielichhoferiaceae. 

303. Mielichhoferia elongata is often treated as a synonym 

or variety of M. mielichhoferiana, but according to 

Shaw (2000a), it is in reality a cryptic species. 

304. Pohlia ramannii Warnst. is recorded from the territory 

that was formerly in Finland and now in Russia 

(Söderström, 1998). It is not mentioned by Ignatov & 

Afonina (1992) and is treated here as doubtful. 

Webera luisieri Dixon and W. maderensis Dixon & 

Luisier, listed by Eggers (1982), are obscure taxa 

whose identity is uncertain. 


305. Sections of Pohlia are based on Shaw (1984), except 

that we follow Smith (2004) in placing P. crudoides in 

section Pohlia and P. flexuosa in section Cacodon. 

306. Pohlia bolanderi was found in the Sierra Nevada, new 

for Europe (Rams et al., 2004). 

307. Pohlia ambigua was described from Austria but 

remains a poorly known taxon. According to Shaw 

(1982) it is a form of P. elongata. It is autoicous and is 

therefore assigned to var. acuminata. 

308. Species limits in the Pohlia nutans group require 

further elucidation. Köckinger, Kučera & Stebel 

(2005) studied P. schimperi and established that the 

only really stable character separating it from P. 

nutans is its red coloration; they accordingly gave it 

subspecies rank and synonymized it with P. nutans 

var. purpurascens. Pohlia marchica Osterwald was 

described from near Berlin and subsequently collected 

on Bornholm. It has not been found for many years, 

and is signified by Nyholm (1993) as very close to P. 

schimperi. Pohlia turonensis J.M.Couderc & Guédès, 

nom. inval. was described from France. It is a close 

relative of P. nutans and has not subsequently been 

validated. 

309. Pohlia saprophila, a mainly Asian species, is reported 

from the Komi Republic, Russia (Zheleznova, 1994). 

It may prove to be a dioicous race of P. elongata, but 

needs further study. 

310. For the synonymy of Pohlia flexuosa see Townsend 

(1995). 

311. Smith (2004) recognized two varieties of Pohlia 

flexuosa in Europe, namely var. flexuosa and var. 

pseudomuyldermansii nom. nud. (P. muyldermansii 

var. pseudomuyldermansii Arts, Nordhorn-Richter & 

A.J.E.Sm.). Townsend (1995) had noted that there is 

continuous variation in bulbil morphology in Nepal 

and India, and argued that the varieties should 

therefore be treated as synonyms. He hypothesized 

that there might be only two clones in Europe, with 

var. muyldermansii possibly a recent introduction. 

However Smith (pers. comm. to Hill) is of the opinion 

that in Europe there has been differentiation into two 

morphologically and ecologically distinct groups of 

plants. In view of the distribution of var. pseudomuyldermansii 

(Austria, British Isles and Switzerland) 

it seems unlikely that the populations concerned 

represent a single clone. The habitat of the varieties 

appears to be markedly different in Europe. For these 

reasons, Smith (above) makes the new combination P. 

flexuosa var. pseudomuyldermansii. 

312. Pohlia tundrae is reported from Central Europe (Düll, 

1991, 1992; Müller, 2004). 

313. Mnium lycopodioides is treated here as a full synonym 

of M. ambiguum following Koponen (1994). 

314. There are a number of records of Trachycystis 

ussuriensis in the Russian Caucasus, e.g. Kharzivov 

et al. (2004).


315. Plagiomnium curvatulum is treated as a (rather cryptic) 

species distinct from P. medium on the basis of a 

genetic study by Wyatt, Odrzykoski & Stoneburner 

(1993). 

316. Plagiomnium undulatum var. madeirense was described 

from Madeira by Koponen & Sérgio (2001). 

317. According to Ochyra & Broughton (2004), 

Orthodontium australe is a synonym of O. lineare. 

318. Leptotheca gaudichaudii is a New Zealand species that 

has been introduced to a garden in Ireland (Smith, 

2004). It is not at present known in the wild in Europe. 

319. Calomnion complanatum is an Australasian species 

that has been introduced to a garden in Ireland 

(Smith, 2004). It is not at present known in the wild in 

Europe. 

320. According to Pfeiffer et al. (2000), Hypopterygium 

tamarisci is a variable and widely distributed species, 

which includes H. muelleri, introduced to Portugal 

from the Southern Hemisphere. We follow Sérgio & 

Carvalho (2003) in using the name H. tamarisci. 

321. Achrophyllum dentatum is an austral species that has 

been introduced to a garden in England (Smith, 2004). 

It is not at present known in the wild in Europe. 

322. Daltonia stenophylla is reported from Azores by 

Sjögren (2001) and Frahm (2004). According to 

Sjögren, Azores records attributed to D. splachnoides 

are actually D. stenophylla; this had been suggested to 

him by R. Schumacker and confirmed by H.A. Crum. 

323. The species epithet of Cyclodictyon laetevirens was 

originally hyphenated as ‘laete-virens’. This is corrected 

here to ‘laetevirens’ in accordance with Article 

60.9 of the International Code (Greuter et al., 2000). 

324. The taxonomy of Fontinalis antipyretica is poorly 

understood and is further complicated by the startling 

results of recent phylogenetic studies using molecular 

methods (Shaw & Allen, 2000; Shaw, 2001). These 

show that European F. antipyretica is more closely 

related to the European endemic F. squamosa than to 

American F. antipyretica, which is morphologically 

indistinguishable from the European species. The 

subspecies retained here are recognized as distinct in 

several European countries, but they may be merely 

ecotypes that have evolved in different locations to 

suit similar selective pressures. Many other infraspecific 

taxa are not included in the main list. Recent 

European checklists have included subsp. lachenaudii 

(Cardot) Podp., subsp. sparsifolia (Limpr.) Kindb., 

var. cymbifolia W.E.Nicholson, and var. gigantea 

(Sull.) Sull. 

325. Fontinalis antipyretica subsp. bryhnii is mapped as a 

species, F. bryhnii, by Söderström (1996), but is 

treated with great scepticism by Nyholm (1960) and 

signified as a possible hybrid by P. Martiny in Stewart 

(1995). In view of the taxonomic difficulties in the F. 

antipyretica aggregate, F. bryhnii is retained here only 

as a subspecies. 

326. Fontinalis dichelymoides is recognized as a species in 

recent lists from Fennoscandia, e.g. Söderström 

(1996). 

327. Generic concepts in Amblystegiaceae and 

Calliergonaceae mostly follow Vanderpoorten et al. 

(2002). Hygroamblystegium is separated from 

Amblystegium according to the results of 

Vanderpoorten et al. (2003) and Serpoleskea is 

retained in Amblystegium. Hygrohypnum is not monophyletic, 

but the genus is retained in its traditional 

sense awaiting ongoing studies regarding the relationships 

of its species. The positions of Conardia, 

Sanionia and Tomentypnum could not be decided by 

Vanderpoorten et al. (2002), and these genera are 

retained in the Amblystegiaceae until further studies 

have been made. 

328. Campylium laxifolium was described from northern 

Sweden by Engelmark & Hedenäs (1990 [1992]). 

329. Vanderpoorten (2004) has argued that the four 

European Hygroamblystegium species should be synonymized 

as H. varium. Further work is needed before 

making changes. 

330. Hygroamblystegium varium is an aptly named species 

in a difficult genus. There is evidence that the 

Hygroamblystegium species recognized here are not 

monophyletic (Vanderpoorten, Cox & Shaw, 2004). 

331. Hygrohypnum as circumscribed here is known to 

comprise heterogeneous elements, probably from 

more than one family. Ochyraea tatrensis would come 

within this circumscription, but a new combination is 

not made, pending a full revision of the group. 

According to Frahm (2005a), O. tatrensis is a mutant 

of Hygrohypnum smithii, but they are superficially 

very different and H. smithii was present at only one of 

the O. tatrensis sites. 

332. The type of Pictus scoticus has been examined by 

Ignatov and others (pers. comm.), who consider it to 

be a form of Hygrohypnum luridum. This view is 

followed here. 

333. Leptodictyum kurdicum (Schiffn.) Broth. is listed for 

Turkey by Kürschner & Erdag˘ (2005). 

334. Palustriella pluristratosa was described from Valais, 

Switzerland (Stech & Frahm, 2001). However, in a 

later publication Frahm (2005a) signifies that P. 

pluristratosa may be a somatic mutation of P. falcata. 

335. Pseudocalliergon angustifolium was described from 

Sweden by Hedenäs (1990 [1992]). 

336. The high-arctic Pseudocalliergon brevifolium was 

transferred to Pseudocalliergon and recognized as a 

species by Hedenäs (1990 [1992]). 

337. Calliergon orbiculare-cordatum (‘orbicularicordatum’) 

(Renauld & Cardot) Broth. was listed by Corley et al. 

(1981) but the specimens are C. richardsonii 

(Söderström, 1996). 

338. For observations on Calliergon richardsonii, see 

Hedenäs (1993b).

339. Warnstorfia procera is recognized as a species following 

Hedenäs (1993b). 

340. The circumscription of Leskeaceae and Hypnaceae 

has been challenged by Gardiner et al. (2005). We 

recognize the merit of their proposals, which would 

necessitate substantial nomenclatural change. We 

prefer not to make the changes until a more general 

overview of families in Hypnales is available. 

341. Lescuraea secunda is reported from European Russia 

in the Urals (Ignatov & Ignatova, 2004). 

342. Lindbergia brachyptera, known from the Russian 

Caucasus, was included by Podpěra (1954) but 

excluded by Corley et al. (1981) because the 

Caucasus was treated by them as part of Asia. 

343. Pseudoleskea was reinstated as a genus distinct from 

Lescuraea by Corley & Crundwell (1991), on the 

basis of peristome differences emphasized by Crum 

& Anderson (1981). The peristome characters now 

seem less important, and Ignatov & Ignatova (2004) 

and Söderström (1998) included Pseudoleskea in 

Lescuraea. A conservative position is adopted 

here, pending more detailed analysis by molecular 

methods. 

344. Pseudoleskea brachyclados (Schwägr.) Kindb., originally 

described from Austria, is listed for Turkey by 

Kürschner & Erdag˘ (2005). According to Podpěra 

(1954) it is a variety of P. radicosa (but if so it should 

have priority at species rank). Other European 

authors have treated it as a variety or synonym of P. 

incurvata. 

345. We follow Wilson & Norris (1989) in including 

Leskeella (Limpr.) Loeske 1903 in Pseudoleskeella 

Kindb. 1897. 

346. Leskeella incrassata type specimen has been examined 

by Ignatov; it is very close to Pseudoleskeella nervosa 

and, if recognized as a species, will prove to be 

common in southern Europe. For the time being the 

two are treated as synonyms. 

347. Pseudoleskeella rupestris was lectotypified and synonymized 

with P. sibirica by Hedenäs & Söderström 

(1991). 

348. In Thuidiaceae, the generic concepts of Touw (2001a) 

are adopted here. In particular Pelekium Mitt. (1868) 

emend. Touw is treated as a synonym of Cyrtohypnum 

(Hampe) Hampe & Lorentz (1869). 

349. Pelekium atlanticum was described from Madeira as 

Thuidium atlanticum (Hedenäs, 1991). The name 

Pelekium atlanticum (Hedenäs) Touw appears in 

Touw (2001a), p. 191, but he failed to validate the 

name by a combination, commenting that P. atlanticum 

is doubtfully distinct from the American P. 

muricatulum (Hampe) Touw. On the other hand, 

Touw was not prepared to treat P. atlanticum as a 

synonym of P. muricatulum. Hedenäs makes the 

necessary combination above. 

350. For the synonymy of Cyrto-hypnum montei with 

Thuidiopsis sparsa, see Touw (2001a, p. 205). Touw 


notes that T. sparsa is known only from a former park 

area in Madeira, and suggests that it is an alien, 

introduced with ornamental plants. 

351. Thuidium assimile has priority over T. philibertii, based 

on the large-scale revision of Touw (2001b). As noted 

by Touw, T. assimile belongs to a complex of poorly 

defined taxa including T. delicatulum. Observations by 

Mrs C.T. van Dorp on European specimens support 

the recognition of two weakly distinguished species, 

rather than treating T. assimile as a variety of T. 

delicatulum. 

352. Subfamilial and generic concepts in Brachytheciaceae 

mostly follow Ignatov & Huttunen (2002). 

These authors overlooked the names Helicodontioideae 

M.Fleisch. and Eurhynchioideae Milde, and 

superfluously described these subfamilies as 

Rhynchostegielloideae and Rhynchostegioideae, 

respectively. 

353. For the taxonomic status of Thamnium cossyrense 

Bott. var. cossyrense and var. melitense Bott. see 

Mastracci (2001). 

354. Platyhypnidium grolleanum was described from Czech 

Republic by Ochyra & Bednarek-Ochyra (1999). 

355. Platyhypnidium mutatum was described by Ochyra & 

Vanderpoorten (1999); molecular studies did not find 

differences between this species and P. riparioides 

(Stech & Frahm, 1999). Frahm (2005a) asserts that it 

is a somatic mutant of P. riparioides. 

356. Platyhypnidium torrenticola was described as 

Gradsteinia torrenticola from the Canary Islands by 

Ochyra, Schmidt & Bultmann (1998) and subsequently 

transferred to the genus Platyhypnidium. 

357. The status of Rhynchostegium arcticum was reevaluated 

by Ignatov & Huttunen (2002). 

358. Rhynchostegium surrectum was synonymized with R. 

confertum by Hedenäs (1992b). 

359. Brachythecium cardotii was synonymized with 

Rhynchostegium megapolitanum by Hedenäs (1992b). 

360. Barbella strongylensis was transferred to Rhynchostegium 

by Buck & Privitera (1999). 

361. Clasmatodon parvulus (Hampe) Sull. is excluded. 

Heras, Infante & Buck (2006) have found that every 

Spanish specimen named as Clasmatodon was 

Pseudoleskeella tectorum. An old German record from 

1851 is poorly localized (Müller, 2004) and is unlikely 

to be C. parvulus. 

362. Nobregaea latinervis is a morphologically very peculiar 

species known from a single locality in Madeira; it was 

described by Hedenäs (1992b). 

363. Synonymy of Helicodontium capillare follows Cortini 

Pedrotti (2006 [2005]), who indicates that a specimen 

of the supposedly endemic H. italicum had been 

identified as H. capillare by W.R. Buck. 

364. Rhynchostegiella bourgaeana is treated by Dirkse & 

Bouman (1995a) as an endemic of the Canary Islands. 

On the other hand, Corley et al. (1991) and Hedenäs 

(1992b) regarded it as a synonym of R. tenella.


365. The status of Rhynchostegiella litorea and its differentiation 

from R. tenella are discussed by Dirkse & 

Bouman (1995a). 

366. Rhynchostegiella macilenta, omitted by Corley et al. 

(1981) and Corley & Crundwell (1991), is treated as a 

full species and illustrated by Dirkse & Bouman 

(1995a). 

367. Rhynchostegiella teesdalei and R. jacquinii were 

synonymized with R. teneriffae by Dirkse & Bouman 

(1995a). 

368. According to Ignatov & Huttunen (2002), 

Rhynchostegiella tenuicaulis does not belong to 

Brachytheciaceae and is more likely a member of 

Amblystegiaceae, but more studies are needed. 

369. Rhynchostegiella trichophylla was described from the 

Canary Islands by Dirkse & Bouman (1995a). 

370. Molecular data do not confirm the traditional view on 

the identity of European Bryhnia scabrida and North 

American B. novae-angliae, so the resurrection of the 

former name was suggested by Ignatov & Huttunen 

(2002). 

371. Popov et al. (2000) showed that Myuroclada maximowiczii 

was reported from Europe on the basis of a 

misidentification and excluded it from the European 

flora, but later Virchenko & Babenko (2001) found it 

as an alien plant in the grounds of the University 

Botanical Garden, Rostov-na-Donu, southern Russia. 

372. The status of Brachythecium glaciale var. dovrense 

needs further study. The taxon may deserve recognition 

as a species. 

373. Brachythecium umbilicatum Jur. & Milde is listed for 

Turkey by Kürschner & Erdag˘ (2005). 

374. Brachythecium ryanii was synonymized with B. 

campestre by Hedenäs (1996). 

375. Brachythecium rotaeanum and B. capillaceum are 

treated as synonyms, as in most recent European 

checklists. However, their identity needs confirmation. 

376. Brachythecium coruscum was recognized as a synonym 

of B. groenlandicum by Nyholm (1965) but the name 

was omitted by Corley et al. (1981). 

377. Brachythecium erythrorrhizon subsp. asiaticum was 

described from the Altai by Ignatov (1998), and is 

known from the Urals. 

378. A typographic error in Wijk et al. (1962) makes it 

seem that Hypnum thedenii (Schimp.) C.Hartm. (1854) 

(not Hartm. 1845 as indicated) was published before 

Brachythecium thedenii Schimp. (1853). 

379. Robinson & Ignatov (1997) showed that 

Brachythecium laetum is the correct name for the 

species often called B. oxycladon. In eastern North 

America this species is very common. It is reported 

from various parts of Europe, but confirmation of the 

identity of European plants and the American one is 

needed. 

380. Brachythecium percurrens was described from Madeira 

by Hedenäs (1992a). Unpublished molecular data by 

Vanderpoorten et al. and Huttunen et al. suggest that 

it belongs to subfamily Helicodontioideae, and may be 

a member of Oxyrrhynchium. 

381. Brachythecium rutabulum var. atlanticum was 

described from Madeira by Hedenäs (1992a). 

382. Brachythecium appleyardiae is a synonym of 

Scleropodium cespitans (Blockeel et al., 2005). 

383. Brachythecium salteri was synonymized with B. dieckii 

by Hedenäs (1993a). 

384. The combinations Brachytheciastrum fendleri and B. 

olympicum were made by Vanderpoorten et al. (2005). 

385. There is still much uncertainty about the correct 

circumscription of Hypnaceae and related families. 

The classification adopted here is provisional, and we 

are well aware that it will be changed in future. 

386. The placement of Hypnum lindbergii and H. pratense 

in Calliergonella and Breidleria, respectively, follows 


387. The circumscription of Campylophyllum is that of 


388. Campylophyllum hispidulum (Brid.) Hedenäs is an 

American species not found in Europe (Crundwell & 

Nyholm, 1962). 

389. Ctenidium molluscum is a very variable species, for 

which numerous infraspecific taxa have been 

described. The following varieties are recognized in 

more than one European country: var. condensatum 

(Schimp.) E.Britton, var. molluscum, and var. robustum 

Boulay. The genetic basis of this variation 

requires further elucidation. 

390. Hypnum cupressiforme has such a complex pattern 

of variation that the delimitation of infraspecific 

taxa remains highly controversial. The varieties in 

the main list are those recognized in current checklists 

of at least four European countries. Other names, 

some of which may have priority over more popular 

ones, are var. brevisetum Schimp., var. imbricatum 

Boulay, var. julaceum Brid., var. tectorum Brid., 

var. tenue Hook. & Taylor, and var. tectorum (Brid.) 

J.-P.Frahm. 

391. Hypnum heseleri was described from Saarland in 

Germany and two localities in the Netherlands 

(Ando & Higuchi, 1994). van Zanten & Hofman 

(1994) grew spores from capsules of H. heseleri and 

demonstrated that the progeny were about 50% H. 

heseleri and 50% H. cupressiforme. Allozyme-electrophoresis 

provided further convincing evidence that H. 

heseleri is a locally derived mutant of H. cupressiforme. 

Accordingly, H. heseleri is treated as a variety 

of H. cupressiforme in a new combination, made 

above. 

392. Hypnum aemulans Breidl. ex Limpr., originally 

described from Austria, is treated as a good 

species by Ando (1994) but is listed without 

comment as a synonym of H. hamulosum by 

European authors (Koperski et al., 2000; Ulvinen, 

Syrjänen & Anttila, 2002). European occurrences need 

further investigation.

393. Hypnum holmenii was described from Canada, 

Greenland and Finland (Ando, 1994), and has been 

found in the Arkhangelsk Oblast of Russia (Ignatov & 

Ignatova, 2004). 

394. Hypnum subimponens is reported from European 

Russia by Afonina (2004). 

395. Vesicularia sphaerocarpa (A.Jaeger) Broth. was 

reported from Malta by Reimers in 1934 on the basis 

of a 19th century collection by Schweinfurth. The 

specimen (B) was destroyed in the Second World War, 

but Reimers in Bizot & Potier de la Varde (1952) was 

confident that it belonged to the taxon described by 

these authors as V. reimersiana. According to Corley 

et al. (1981), V. sphaerocarpa is a synonym of V. 

galerulata (Duby) Broth. Düll (1985) listed the 

Maltese plant as V. reimersiana, but also asserted that 

V. galerulata had been found in mainland Italy. 

Corley & Crundwell (1991) accordingly added V. 

reimersiana to the list, retaining V. galerulata. 

However, no species of Vesicularia is listed for Italy 

by Cortini Pedrotti (2006 [2005]). It seems likely that 

V. galerulata has never been found in Europe. 

396. Smith (2004) treated Heterocladium flaccidum as a 

species, but provided only a nomen nudum. He has 

made the necessary new combination above for this 

checklist, noting (pers. comm. to Hill) that consultation 

with a number of British field bryologists revealed 

that, with one exception, none had encountered 

intermediates between it and H. heteropterum. 

397. Heterocladium wulfsbergii is discussed in detail by 

Crundwell & Smith (2000). 

398. Iwatsukiella leucotricha was listed for Europe by 

Podpěra (1954) under the name Habrodon leucotrichus 

(Mitt.) Perss., with a locality in the southern Urals. It 

was omitted without comment by Corley et al. 

(1981), but is indicated from the European side of 

the Urals by Dierssen (2001) and Ignatov & Ignatova 

(2004). 

399. Alpine and arctic forms of Hylocomium splendens have 

been variously treated. Var. alpinum Schlieph. ex 

Limpr. is mapped by Söderström (1996), but is 

doubted and treated as possibly a synonym of var. 

obtusifolium (Geh.) Paris by Koperski et al. (2000). 

Var. obtusifolium is commonly treated as a synonym 

of H. alaskanum (Lesq. & James) Austin, whose 

distinguishing characters have been shown by Ross et 

al. (2001) to be mainly under environmental rather 

than genetic control. It is not clear whether any of 

these forms is sufficiently distinct genetically to 

deserve taxonomic recognition. 

400. Herzogiella adscendens (Lindb.) Z.Iwats. & W.B.Schofield 

has been reported from Svalbard (Corley & 

Crundwell, 1991; Düll, 1985), but the record 

was subsequently rejected (Frisvoll & Elvebakk, 

1996). 

401. Myurella julacea var. ciliata (Chal.) Ochyra & 

Bednarek-Ochyra (syn. M. julacea var. scabrifolia 


Lindb. ex Limpr.) is treated as distinct in some 

European countries, but intergrades completely with 

var. julacea. It is not recognized here. 

402. Orthothecium chryseon is incorrectly listed as O. 

chryseum by Crosby et al. (1999). It was described as 

Hypnum chryseon by Schwägrichen (1804). 

403. Plagiothecium curvifolium is sometimes treated as a 

variety (in Europe) or synonym (in North America) 

(Ireland, 2003) of P. laetum, to which it is undoubtedly 

very close. As a variety, its correct name is P. laetum 

var. secundum. 

404. Plagiothecium denticulatum var. undulatum is treated 

in many European lists as a species, P. ruthei, but in 

North America merely as a synonym of P. denticulatum 

(Ireland, 2003). It is not clearly distinct from var. 

denticulatum, and we follow Ignatov & Ignatova 

(2004), Koperski et al. (2000) and Smith (2004) in 

treating it as a variety. 

405. Plagiothecium noricum was described from the 

Pinzgau, Austria and was thought by Molendo 

(but not Limpricht) to intergrade with P. neckeroideum. 

Grims (1999) listed it but noted that Düll 

(1992) had thought it a weak species, possibly 

only a variety of P. neckeroideum. It is not recognized 

here. 

406. Plagiothecium succulentum is a difficult taxon, 

intergrading on the one hand with P. nemorale 

(Hemeric, 1989) and on the other with P. cavifolium 

(Nyholm, 1965). It is also very close to P. platyphyllum. 

It is retained here, because it cannot satisfactorily 

be subordinated to any of these species without 

treating the whole group as a single, rather-broad 

species. 

407. Plagiothecium svalbardense was described by Frisvoll 

in Frisvoll & Elvebakk (1996). 

408. Pseudotaxiphyllum laetevirens was raised to species 

rank by Hedenäs (1992a). 

409. Entodon challengeri occurs in European Russia 

(Ignatov & Ignatova, 2004) and is the correct name 

for the plant formerly known as E. compressus 

(Iwatsuki & Tan, 2001). 

410. According to Ochyra & Ireland (2004), Isopterygium 

tenerum is absent from Europe but present in sub- 

Saharan Africa. However, Schumacker in Stewart 

(1995), Cortini Pedrotti (2006 [2005]) and Gabriel 

et al. (2005) list it for Italy and the Azores. 

Isopterygium bottinii was excluded from Sematophyllum 

by Guerra & Gallego (2005), and is included 

here in I. tenerum. 

411. Hageniella micans, previously known from Europe 

and North America, was synonymized with H. 

pacifica Broth., which occurs in China and Taiwan 

(Tan & Jia, 1999). 

412. Sematophyllum adnatum was found in Italy, where it is 

thought to be an introduction from North America 

(Brusa in Blockeel et al., 2000).


413. Cryphaea lamyana was transferred to the otherwise 

tropical and Australasian genus Dendrocryphaea by 

Rao (2001). 

414. Leucodon flagellaris author citation is based on 

synonymy in Podpěra (1954). 

415. Leucodon pendulus, whose main distribution is in 

eastern Asia, has been found in the Ivanovo Oblast 

NE of Moscow (Ignatov & Ignatova, 2004). 

416. Leucodon treleasei was treated as a synonym of L. 

canariensis by Corley et al. (1981) but as a distinct 

species by Hedenäs (1992b). 

417. Homalia trichomanoides author citation follows that in 

the International Code (Greuter et al., 2000) Appendix 

IIIa, rather than Corley & Crundwell (1991). 

418. The characters emphasized by Steere (1941) do 

not justify generic status for Metaneckera. There 

are, especially in Asia, several species of Neckera that 

have some or all of those chracters (J. Enroth, pers. 

obs.). 

419. Thamnobryum cataractarum was described from 

England by Hodgetts & Blockeel (1992). 

420. Thamnobryum fernandesii was described from 

Madeira by Sérgio (1981). 

421. Thamnobryum maderense, signified by Hedenäs 

(1992b) as a good species, was reduced to a variety 

of T. alopecurum by Stech, Ros & Werner (2001) on 

the basis of molecular data. We are not fully 

convinced by their argument and prefer to retain T. 

maderense as a species for the time being. 

422. Thamnobryum neckeroides was reported from Czech 

Republic, Germany and Italy by Mastracci (2003). 

423. Thamnobryum rudolphianum was described from the 

Azores by Mastracci (2004). 

424. The Madeiran endemic Echinodium setigerum was not 

mentioned by Corley et al. (1981) but was recognized 

as a good species by Churchill (1986). 

425. Cryptoleptodon longisetus was omitted by Corley et al. 

(1981) but listed as Leptodon longisetus by Düll (1985). 

It was transferred to Cryptoleptodon by Enroth (1992). 

426. Synonymy of Isothecium algarvicum follows Enroth & 


427. According to Smith (2004), Isothecium myosuroides 

var. brachythecioides and I. holtii, although distinct in 

their typical expressions, are linked to I. myosuroides 

by intermediate forms. 

ACKNOWLEDGEMENTS 

Financial support for publication of this checklist came 

from the British Bryological Society (through its Bequest 

Fund), from English Nature and from the Centre for 

Ecology and Hydrology (CEH). We are grateful to those 

members of the Steering Committee who are not authors, 

namely Michele Aleffi, Bill Buck, Klaus Dierssen and Jirˇí 

Váňa, for wise advice during the project. Francis Rowland 

of CEH designed an attractive website. We thank Ron 

Porley (Newbury, England) for encouraging us to realize 

the ECCB’s proposal to make a new European checklist. 

Bernard Goffinet (Connecticut) supplied an electronic copy 

of the Goffinet & Buck (2004) taxonomic hierarchy. John 

Tweddle (London) checked the author abbreviations 

against IPNI recommendations. We made extensive use of 

data on websites, especially the W 3 MOST database of 

Missouri Botanical Garden, and the author query facility of 

the International Plant Names Index. For help on 

particular points, we thank Olga Afonina (St Petersburg, 

Sphagnum), Hans Blom (Bergen, Schistidium), Alicia 

Ederra (Navarra, Cinclidotus), Gerard Dirkse (Nijmegen, 

Tortella limbata), T.V. Egorova (St Petersburg, Sphagnum), 

Len Ellis (London, Conardia), Tomas Hallingbäck 

(Uppsala, various taxa), Patxi Heras & Marta Infante 

(Vitoria-Gasteiz, Clasmatodon), Diana Horton (Iowa, 

Encalypta), Juan Antonio Jiménez (St Louis, Didymodon), 

Eva Maier (Geneva, Grimmia), Rayna Natcheva (Lund, 

Bulgarian mosses), Ron Pursell (Pennsylvania, Fissidens), 

Cecília Sérgio (Lisbon, Portuguese and Macaronesian 

mosses), Tony Smith (north Wales, British mosses), Alain 

Vanderpoorten (Liège, Macaronesian mosses). Holyoak’s 

Bryum research was supported by the HIGH LAT 

RESOURCE under the EC-funded IHP programme, 

Plantlife International, and facilitated by Jeff Duckett 

(London). Finally, we thank two anonymous referees for 

helpful and detailed comments, which have resulted in 

many corrections and improvements. 

TAXONOMIC ADDITIONS and CHANGES: Heterocladium 

flaccidum (Schimp.) A.J.E.Sm., stat. et comb. nov. 

(Heterocladium heteropterum var. flaccidum Schimp.); 

Hypnum cupressiforme var. heseleri (Ando & Higuchi) 

M.O.Hill, stat. et comb. nov. (Hypnum heseleri Ando & 

Higuchi); Pelekium atlanticum (Hedenäs) Hedenäs, comb. 

nov. (Thuidium atlanticum Hedenäs); Pohlia flexuosa var. 

pseudomuyldermansii (Arts, Nordhorn-Richter & 

A.J.E.Sm.) A.J.E.Sm., comb. nov. (Pohlia muyldermansii 

var. pseudomuyldermansii Arts, Nordhorn-Richter & 

A.J.E.Sm.); Weissia6mittenii (Bruch & Schimp.) Mitt. 

emend. A.J.E.Sm., stat. nov. (Weissia mittenii (Bruch & 

Schimp.) Mitt.). 

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INDEX 

Accepted names are given in roman type. Synonyms and names 

mentioned only in the annotations are in italics. The numbers following 

accepted names and synonyms are those of the genus and species in the 

main list. For names appearing only in the annotations, the annotation 

number is given. 

Abietinella, 199 

abietina, 199/1 

hystricosa, 199/1, Ann32 

Acaulon, 97 

casasianum, 97/1 

dertosense, 97/2 

fontiquerianum, 97/3 

mediterraneum, 97/4, Ann163 

muticum, 97/5, Ann163 

piligerum, 97/6 

triquetrum, 97/7 

Achrophyllum, 161 

dentatum, 161/1, Ann321 

Aloina, 98 

aloides, 98/1 

ambigua, 98/2 

bifrons, 98/3 

brevirostris, 98/4 

humilis, 98/5, Ann164 

obliquifolia, 98/6, Ann165 

rigida, 98/7 

Alophosia, 4 

azorica, 4/1 

Amblyodon, 124 

dealbatus, 124/1 

Amblystegium, 171 

compactum, 175/1 

confervoides, 171/1 

fluviatile, 178/1 

humile, 178/2 

jungermannioides, 255/1 

juratzkanum, 171/3 

radicale, 171/2 

riparium, 180/1 

saxatile, 171/2 

serpens, 171/3 

subtile, 171/4 

tenax, 178/3 

varium, 178/4 

Amphidium, 54 

curvipes, 54/3, Ann118 

lapponicum, 54/1 

mougeotii, 54/2 

tortuosum, 54/3, Ann118 

Anacamptodon, 172 

splachnoides, 172/1 

Anacolia, 134 

laevisphaera, 134/1, Ann256 

menziesii, 134/2, Ann257 

webbii, 134/3, Ann257 

Andoa, 227 

berthelotiana, 227/1 

Andreaea, 2 

alpestris, 2/8 

alpina, 2/9 

blyttii, 2/2 

crassifolia, 2/5 

crassinervia, 2/3 

frigida, 2/4 

hartmanii, 2/11 

heinemannii, 2/5, Ann16 

megistospora, 2/6 

mutabilis, 2/10 

nivalis, 2/1 

obovata, 2/11

planinervia, 2/5 

rothii, 2/7 

rupestris, 2/12 

sinuosa, 2/13 

Anisothecium 

grevilleanum, 68/4 

humile, 68/7 

palustre, 58/2 

rigidulum, 68/7 

rubrum, 68/12 

rufescens, 68/8 

schreberianum, 68/9 

staphylinum, 68/10 

vaginale, 68/3 

varium, 68/12 

Anoectangium, 79 

aestivum, 79/1 

angustifolium, 79/1 

handelii, 79/2, Ann141 

hornschuchianum, 89/1 

schliephackei, 89/2 

sendtnerianum, 89/3 

taeniatifolium, 89/4 

tenuinerve, 89/5 

warburgii, 89/6 

Anomobryum, 140 

concinnatum, 140/1, Ann265 

filiforme, 140/2 

julaceum, 140/2, Ann265 

juliforme, 140/2 

lanatum, 142/6 

leptostomoides, 140/1, Ann265 

lusitanicum, 140/3, Ann266 

Anomodon, 277 

attenuatus, 277/1 

longifolius, 277/2 

rostratus, 277/3 

rugelii, 277/4 

tristis, 277/5 

viticulosus, 277/6 

Antitrichia, 266 

californica, 266/1 

curtipendula, 266/2 

Aongstroemia, 67 

longipes, 67/1 

Aphanorrhegma 

patens, 22/1 

Aplodon, 119 

wormskioldii, 119/1, Ann32 

Archidium, 40 

alternifolium, 40/1 

Arctoa, 55 

anderssonii, 55/1 

fulvella, 55/2 

hyperborea, 55/3 

Aschisma, 80 

carniolicum, 80/1 


cuynetii, 80/2 

Astomum, 96 

crispum, 96/5 

levieri, 96/4 

mittenii, Ann161 

multicapsulare, 96/6 

Atractylocarpus, 71 

alpinus, 71/1 

Atrichum, 5 

angustatum, 5/1, Ann17 

crispum, 5/2 

flavisetum, 5/3, Ann18 

haussknechtii, 5/3 

rhystophyllum, Ann17 

tenellum, 5/4 

undulatum, 5/5, Ann17, Ann18 

Aulacomnium, 156 

androgynum, 156/1 


turgidum, 156/3 

Barbella 

strongylensis, 210/6, Ann360 

Barbula, 99 

acuta, 104/1 

adriatica, 104/8 

asperifolia, 104/2 

bicolor, 99/1 

bolleana, 99/2, Ann139 

commutata, 99/3, Ann167 

convoluta, 99/3 

cordata, 104/6 

crocea, 99/4 

cylindrica, 104/13 

ehrenbergii, 99/2 

enderesii, 99/5 

fallax, 104/8 

gigantea, 104/10 

hornschuchiana, 113/1 

indica, 99/6 

insidiosa, 104/22 

johansenii, 104/14 

kneuckeri, 104/2 

lurida, 104/16 

reflexa, 104/9 

revoluta, 113/4 

rigidula, 104/19 

sardoa, 99/3, Ann167 

semilimbata, Ann166 

sinuosa, 104/21 

spadicea, 104/22 

tophacea, 104/25 

unguiculata, 99/7 

vinealis, 104/27 

Bartramia, 135 

breviseta, 135/3, Ann260 

halleriana, 135/1 

ithyphylla, 135/4, Ann260, Ann261 

pomiformis, 135/2, Ann258 

stricta, 135/6, Ann256 

subulata, 135/5 

Blindia, 37 

acuta, 37/1 

caespiticia, 37/2 

Brachydontium, 38 

trichodes, 38/1, Ann84 

Brachymenium, 141 

cellulare, 142/14 

commutatum, 141/1, Ann267 

exile, Ann267, Ann270 

notarisii, 141/2, Ann268 

paradoxum, 141/3, Ann269 

philonotula, 141/4, Ann270 

Brachytheciastrum, 223 

collinum, 223/1 

dieckii, 223/2, Ann383 

fendleri, 223/3, Ann384 

olympicum, 223/4, Ann384 

trachypodium, 223/5 

vanekii, 223/6 

velutinum, 223/7 

Brachythecium, 220 

albicans, 220/1 

appleyardiae, 221/1, Ann382 

campestre, 220/2, Ann374 

capillaceum, 220/3, Ann375 

cardotii, 210/3, Ann359 

cirrosum, 220/4 

collinum, 223/1 

coruscum, 220/5, Ann376 

curtum, 219/4 

dieckii, 223/2, Ann383 

dovrense, 219/2 

erythrorrhizon, 220/6, Ann377 

fendleri, 223/3 

geheebii, 220/7 

glaciale, 219/2, Ann372 

glareosum, 220/8 

groenlandicum, 220/5, Ann376 

laetum, 220/9, Ann379 

latifolium, 219/3 

mildeanum, 220/10 

oedipodium, 219/4 

olympicum, 223/4 

ornellanum, 219/5 

oxycladon, 220/9, Ann379 

percurrens, 220/11, Ann380 

plumosum, 219/6 

populeum, 219/7 

reflexum, 219/8 

rivulare, 220/12 

rotaeanum, 220/3, Ann375 

rutabulum, 220/13, Ann381 

ryanii, 220/2, Ann374 

salebrosum, 220/14


salteri, 223/2, Ann383 

starkei, 219/9 

thedenii, Ann378 

tommasinii, 220/15 

trachypodium, 223/5 

turgidum, 220/16 

udum, 220/10 

umbilicatum, Ann373 

vanekii, 223/6 

velutinum, 223/7 

Braunia, 131 

alopecura, 131/1 

Breidleria, 228 

pratensis, 228/1 

Breutelia, 136 

azorica, 136/1 

chrysocoma, 136/2 

Brotherella, 258 

lorentziana, 258/1 

Bruchia, 52 

flexuosa, 52/1 

trobasiana, 52/1 

vogesiaca, 52/2 

Bryhnia, 217 

novae-angliae, 217/1, Ann370 

scabrida, 217/1, Ann370 

Bryobrittonia, 16 


Bryoerythrophyllum, 100 

alpigenum, 100/1 

caledonicum, 100/2 

campylocarpum, 100/3, Ann168 

ferruginascens, 100/4 

inaequalifolium, 100/5 

lusitanicum, 100/3 

machadoanum, 100/3 

recurvirostrum, 100/6 

rubrum, 100/7 

Bryohaplocladium 


microphyllum, 192/2 

virginianum, 192/3 

Bryoxiphium, 28 

madeirense, 28/1, Ann48 

norvegicum, 28/2 

Bryum, 142 

acutiforme, 142/10 

aeneum, 142/41, Ann293 

affine, 142/15 

algovicum, 142/1, Ann276 

alpinum, 142/2, Ann274, Ann291 

amblyodon, 142/4 

apiculatum, 142/3, Ann277 

archangelicum, 142/4, Ann296 

arcticum, 142/5 

argenteum, 142/6, Ann279, Ann280 

axel-blyttii, 142/10 

badium, 142/9 

balticum, 142/19 

barnesii, 142/19 

bicolor, 142/19, Ann287 

bimum, 142/43, Ann295 

blindii, 142/7, Ann281 

bornholmense, 142/8, Ann282 

bryoides, 142/5 

caespiticium, 142/9, Ann283 

calophyllum, 142/10 

canariense, 142/11, Ann284 

capillare, 142/12 

castaneum, 142/43 

caucasicum, 142/13, Ann285 

cavifolium, 142/39 

cellulare, 142/14, Ann269 

cernuum, 142/55 

chibinense, 142/23 

colombii, Ann274 

comense, 142/9, Ann283 


concinnatum, 140/1 

creberrimum, 142/15 

crispulum, 142/39 

cruegeri, 142/3 

cryophilum, 142/16 

curvatum, 142/4 

cuspidatum, 142/15 

cyclophyllum, 142/17 

demaretianum, 142/18, Ann286 

demissum, 143/1 

dichotomum, 142/19 

dixonii, 142/20 

donianum, 142/21 

dunense, 142/19 

duvalii, 142/59 

dyffrynense, 142/22, Ann287 

ekstamii, Ann272 

elegans, 142/23 

erythrocarpum, 142/51 

excurrens, 142/19 


flaccidum, 142/37 

funckii, 142/24 

funkii, 142/24 

geheebii, Ann274 

gemmiferum, 142/25 

gemmilucens, 142/26 

gemmiparum, 142/27, Ann275 

gerwigii, Ann275 

globosum, 142/60 

icodense, 142/53 

imbricatum, 142/4 

inclinatum, 142/4 

intermedium, 142/28, Ann274 

julaceum, 140/2 

juliforme, 140/2 

klinggraeffii, 142/29 

knowltonii, 142/30 

kunzei, 142/31, Ann283, Ann288 

labradorense, Ann272 

lacustre, 142/30 

laevifilum, 142/37 

lanatum, 142/6 

lawersianum, 142/5 

leptostomum, 140/2 

lisae, 142/15 

lonchocaulon, 142/42, Ann294 

longisetum, 142/32 

mamillatum, 142/58 

marratii, 142/33 

mesodon, Ann274 

microerythrocarpum, 142/51 

mildeanum, 142/34 

miniatum, 142/35, Ann289 

minii, 142/36, Ann290 

moravicum, 142/37 

muehlenbeckii, 142/38, Ann291 

murmanicum, Ann274 

neodamense, 142/39, Ann292 

nitens, 142/3 

nitidulum, 142/28, Ann274 

obconicum, 142/42, 142/53 

oblongum, 142/40, Ann281 

obovatum, 142/21 

obtusifolium, 142/16 

oelandicum, 142/58 

oeneum, 142/41, Ann293 

ontariense, 144/1 

pachyloma, 142/21 

pallens, 142/41, Ann106, Ann279 

pallescens, 142/42, Ann294 

pamirense, Ann271 

platyloma, 142/12 

provinciale, 142/11, Ann284 

pseudotriquetrum, 142/43, Ann292, Ann295 

purpurascens, 142/5 

pyriferum, 142/56, Ann299 

radiculosum, 142/44 


roseum, 144/2 

rubens, 142/46 

ruderale, 142/47 

rufifolium, 142/12 

rutilans, 142/41 

salinum, 142/48, Ann296 

sauteri, 142/49 

savicziae, Ann271, Ann272 

schleicheri, 142/50, Ann297 

splachnoides, 142/14, Ann269 

stenotrichum, 142/4 

stirtonii, 142/23 

subapiculatum, 142/51 

subelegans, 142/37, 142/41

subneodamense, 142/39 

syriacum, 142/54 

tenuisetum, 142/52 

teres, Ann274 

torquescens, 142/53, Ann298 

tortifolium, 142/17 

turbinatum, 142/54 

uliginosum, 142/55, Ann271, Ann272 

umbratum, Ann272 

validicostatum, 142/12 

valparaisense, 142/56, Ann299 

ventricosum, 142/43 

vermigerum, Ann273 

veronense, 142/6, Ann279 

versicolor, 142/19 

violaceum, 142/57 

warneum, 142/58 

weigelii, 142/59 

wrightii, 142/60 

zemliae, Ann272 

zieri, 143/2, Ann300 

Buckiella 

undulata, 254/13 

Bucklandiella 

affinis, 33/10 

elliptica, 33/11 

heterosticha, 33/12 

himalayana, 33/13 

lamprocarpa, 33/14 

lusitanica, 33/15 

macounii, 33/16 

microcarpa, 33/17 

obtusa, 33/18 

sudetica, 33/19 

Buxbaumia, 13 

aphylla, 13/1 

viridis, 13/2 

Callialaria 

curvicaulis, 176/1 

Callicladium, 229 

haldanianum, 229/1 

Calliergon, 186 

cordifolium, 186/1 

giganteum, 186/2 

megalophyllum, 186/3 


orbiculare-cordatum, Ann337 

richardsonii, 186/4, Ann337, Ann338 

sarmentosum, 191/5 

stramineum, 190/1 

trifarium, 183/4 

Calliergonella, 230 

cuspidata, 230/1 

lindbergii, 230/2 

Calomnion, 159 

complanatum, 159/1, Ann319 

Calymperes, 76 


erosum, 76/1 

sommieri, 76/1 

Calyptrochaeta, 162 

apiculata, 162/1 

Camptothecium 

lutescens, 224/2 

Campyliadelphus, 173 

chrysophyllus, 173/1 

elodes, 173/2 

polygamus, 177/4 

stellatus, 174/4 

Campylidium 

calcareum, 231/1 

sommerfeltii, 231/3 

Campylium, 174 



chrysophyllum, 173/1 

elodes, 173/2 

halleri, 231/2 

hispidulum, 231/3 

laxifolium, 174/1, Ann328 

longicuspis, 174/2 

polygamum, 177/4 

protensum, 174/3 

radicale, 171/2 


stellatum, 174/4 

Campylophyllum, 231 


halleri, 231/2 

hispidulum, 231/3, Ann388 


Campylopus, 72 

atrovirens, 72/1 

brevipilus, 72/2 

carreiroanus, 72/15 

cygneus, 72/3 

eximius, 72/8, Ann135 

flaccidus, 72/4 

flexuosus, 72/5 

fragilis, 72/6 

gracilis, 72/7 

incrassatus, 72/8, Ann135 

introflexus, 72/9 

madeirensis, 46/8 

marginatulus, 46/8 

mildei, 72/10 

oerstedianus, 72/10 

pilifer, 72/11 

pyriformis, 72/12 

schimperi, 72/13 

schwarzii, 72/7 

setaceus, 72/3 

setifolius, 72/14 

shawii, 72/15 

subporodictyon, 73/3 

subulatus, 72/16 

Campylostelium, 35 

pitardii, 35/1, Ann83 

saxicola, 35/2 

strictum, 35/3 

Catoscopium, 133 

nigritum, 133/1 

Ceratodon, 42 

antarcticus, Ann111 

conicus, 42/1 

corsicus, 42/2 

heterophyllus, Ann111 

purpureus, 42/2, Ann111, Ann112 

stenocarpus, 42/2 

Cheilothela, 43 

chloropus, 43/1 

Chenia 

leptophylla, 108/1 

rhizophylla, 108/1 

Cinclidium, 151 



stygium, 151/3 

subrotundum, 151/4 

Cinclidotus, 101 

aquaticus, 101/1 

bistratosus, Ann169 

confertus, 101/2, Ann170 

danubicus, 101/3 

fontinaloides, 101/4 

mucronatus, 103/2 

nyholmiae, Ann169 

pachylomoides, 101/5 

riparius, 101/6, Ann170 

vivesii, 101/7, Ann171 

Cirriphyllum, 214 

cirrosum, 220/4 

crassinervium, 214/1 

germanicum, 213/8 

piliferum, 214/2 

reichenbachianum, 219/1 

tenuicaule, 213/8 

tenuinerve, 220/15 

tommasinii, 220/15 

velutinoides, 219/1 

Claopodium, 278 

whippleanum, 278/1 

Clasmatodon 

parvulus, Ann361 

Cleistocarpidium, 44 


Climacium, 170 

dendroides, 170/1 

Cnestrum, 56 

alpestre, 56/1 

glaucescens, 56/2 

schisti, 56/3


Codonoblepharon 

forsteri, 130/3 

Codriophorus 

acicularis, 33/6 

aquaticus, 33/7 

fascicularis, 33/8 

hespericus, 33/9 

Conardia, 175 

compacta, 175/1 

Conostomum, 137 

tetragonum, 137/1 

Coscinodon, 29 

cribrosus, 29/1 


Crassiphyllum 

fernandesii, 271/4 

Cratoneuron, 176 

commutatum, 182/1 

curvicaule, 176/1 

decipiens, 182/2 

falcatum, 182/3 

filicinum, 176/2 

Crossidium, 102 

aberrans, 102/1 

crassinerve, 102/2 

davidai, 102/3, Ann172 

geheebii, 102/4, Ann173 

laevipilum, 102/5, Ann174 

laxefilamentosum, 102/6, Ann175 

seriatum, Ann221 

squamiferum, 102/7 

Crumia 

latifolia, Ann140 

Cryphaea, 264 

heteromalla, 264/1 

lamyana, 265/1, Ann413 

Cryptoleptodon, 273 

longisetus, 273/1, Ann425 

Ctenidium, 232 

molluscum, 232/1, Ann389 

procerrimum, 235/12 

Cyclodictyon, 167 

laetevirens, 167/1, Ann323 

Cynodontium, 57 


asperifolium, 57/1, Ann119 

bruntonii, 57/2 

fallax, 57/3, Ann119 

gracilescens, 57/4 

jenneri, 57/5 

polycarpon, 57/6, Ann120 

strumiferum, 57/7 

suecicum, 57/8 

tenellum, 57/9 

torquescens, 57/9 

Cyrto-hypnum 

atlanticum, 201/1 

minutulum, 201/2 

montei, 202/1, Ann350 

Cyrtomnium, 152 

hymenophylloides, 152/1 

hymenophyllum, 152/2 

Daltonia, 163 

splachnoides, 163/1, Ann322 

stenophylla, 163/2, Ann322 

Dendrocryphaea, 265 

lamyana, 265/1 

Desmatodon, 117 

bogosicus, 117/4 

cernuus, 117/8 

guepinii, 117/11 

heimii, 105/1 

latifolius, 117/12 

laureri, 117/16 

leucostoma, 117/17 

meridionalis, 117/19, Ann223 

oxneri, 117/25 

randii, 117/25 

systylius, 117/31 

ucrainicus, 117/33 

wilczekii, 117/27, Ann225 

Dialytrichia, 103 

fragilifolia, 103/1, Ann176 

mucronata, 103/2 

Dichelyma, 168 

capillaceum, 168/1 

falcatum, 168/2 

Dichodontium, 58 

flavescens, 58/1 

palustre, 58/2, Ann121 

pellucidum, 58/3 

Dicranella, 68 

campylophylla, 68/1, Ann126 

cerviculata, 68/2 

crispa, 68/3 

curvata, 68/11 

grevilleana, 68/4 

heteromalla, 68/5 

howei, 68/6 

humilis, 68/7 

palustris, 58/2 

riparia, 61/4 


schreberiana, 68/9 

secunda, 68/11 

staphylina, 68/10 

subulata, 68/11 

varia, 68/12 

Dicranodontium, 73 

asperulum, 73/1 

circinatum, 73/4 

denudatum, 73/2 

subporodictyon, 73/3, Ann136 

uncinatum, 73/4 

Dicranoweisia, 59, Ann122 

cirrata, 59/1 


crispula, 59/3 

Dicranum, 69 

acutifolium, 69/8 

affine, 69/13 

angustum, 69/16, Ann132 

bergeri, 69/13, Ann131 

bonjeanii, 69/1 

brevifolium, 69/9 

canariense, 69/23, Ann134 

congestum, 69/14 

crassifolium, 69/2, Ann128 

dispersum, 69/10, Ann130 

drummondii, 69/11 

elatum, 69/11 

elongatum, 69/19, Ann133 

flagellare, 69/27 

flexicaule, 69/14 

fragilifolium, 69/20 

fulvum, 69/24 

fuscescens, 69/15, Ann133 

groenlandicum, 69/21 

laevidens, 69/22, Ann132 

leioneuron, 69/3 

majus, 69/4 

montanum, 69/28 

muehlenbeckii, 69/17 

neglectum, 69/18 


polysetum, 69/5, Ann106 

robustum, 69/11 

rugosum, 69/5 

scoparium, 69/6 

scottianum, 69/25, Ann79, Ann134 

sendtneri, 69/19, Ann133 

spadiceum, 69/18 

spurium, 69/12 


subporodictyon, 73/3 

tauricum, 69/29 

transsylvanicum, 69/7, Ann129 

undulatum, 69/13, 69/1, 69/5, Ann131 

viride, 69/26 

Didymodon, 104 

aaronis, 104/3 

acutus, 104/1 

asperifolius, 104/2, Ann180 

australasiae, 104/3, Ann181 

barbuloides, 104/22 

bistratosus, 104/4, Ann182 

bosniacus, 104/25 

brachyphyllus, 104/5, Ann183 

cordatus, 104/6 

erosus, 104/7, Ann184 

fallax, 104/8, Ann180

ferrugineus, 104/9 

giganteus, 104/10, Ann180 

glaucus, 104/11 

icmadophilus, 104/12 

incrassatus, 104/3 

insulanus, 104/13, Ann185 

johansenii, 104/14 

lamyanus, 104/15, Ann185 

luridus, 104/16 

mamillosus, 104/19, Ann186 

maximus, 104/17, Ann180 

nicholsonii, 104/18, Ann185 

reedii, Ann178 

rigidulus, 104/19, Ann186 

rufus, 104/2 

sicculus, 104/20, Ann187 

sinuosus, 104/21 

soaresii, Ann179 

spadiceus, 104/22, Ann180 

subandreaeoides, 104/23, Ann188 

tectorum, Ann178 

tomaculosus, 104/24 

tophaceus, 104/25, Ann180 

trifarius, 104/16 

trivialis, 104/26 

umbrosus, 104/26, Ann189 

vinealis, 104/27, Ann185 

zetterstedtii, 104/22 

Diobelonella 


Diphyscium, 14 

foliosum, 14/1 

Discelium, 25 

nudum, 25/1 

Distichium, 45 

capillaceum, 45/1 

hagenii, 45/2 

inclinatum, 45/3 

Distichophyllum, 164 

carinatum, 164/1 

Ditrichum, 46 

cornubicum, 46/1 

crispatissimum, 46/3, Ann113 

cylindricum, 51/1 

difficile, Ann114 

flexicaule, 46/2 

giganteum, 46/3 

gracile, 46/3, Ann113 

heteromallum, 46/4 

homomallum, 46/4 

lineare, 46/5 

pallidum, 46/6 

plumbicola, 46/7 

punctulatum, 46/8, Ann114 

pusillum, 46/9 

subulatum, 46/10 

tortile, 46/9 


vaginans, 46/5 

zonatum, 46/11 

Drepanium 

recurvatum, 235/13 

Drepanocladus, 177 

aduncus, 177/1 

arcticus, 177/2 

badius, 188/1 

brevifolius, 183/2 

capillifolius, 177/3 

cossonii, 189/1 

exannulatus, 191/1 

fluitans, 191/2 

h-schulzei, 191/2 

intermedius, 189/1 

lapponicus, 187/1 

longifolius, 177/3 

lycopodioides, 183/3 

polycarpos, 177/1 

polygamus, 177/4 

procerus, 191/3 

pseudostramineus, 191/4 

revolvens, 189/2 

sendtneri, 177/5 

simplicissimus, 177/1 

sordidus, 177/6 

stagnatus, 177/1 

tenuinervis, 177/6 

trichophyllus, 191/6 

tundrae, 191/7 

uncinatus, 184/3 

vernicosus, 187/2 

Dryptodon 

anomalus, 30/3 

contortus, 30/18 


elongatus, 30/14 

funalis, 30/15 


incurvus, 30/13, Ann56 


orbicularis, 30/26 

patens, 30/31 

pulvinatus, 30/30 

torquatus, 30/35 

trichophyllus, 30/36 

Echinodium, 272 

madeirense, 272/4 

prolixum, 272/1 

renauldii, 272/2 

setigerum, 272/3, Ann424 

spinosum, 272/4 

Encalypta, 17 

affinis, 17/11, Ann33 

alpina, 17/3 

brevicolla, 17/12 

brevipes, 17/13 

ciliata, 17/14, Ann28 

commutata, Ann29 

intermedia, 17/5, Ann29 

longicolla, 17/10 



microstoma, 17/15 

mutica, 17/4 

obovatifolia, 17/6, Ann30 

procera, 17/1 

rhaptocarpa, 17/7, Ann31 

serbica, Ann28 

spathulata, 17/8 

streptocarpa, 17/2 

trachymitria, 17/7, Ann31 

vulgaris, 17/9 

Entodon, 257 

challengeri, 257/1, Ann409 

cladorrhizans, 257/2 

compressus, 257/1, Ann409 

concinnus, 257/3 

schleicheri, 257/4 

Entosthodon, 18 

attenuatus, 18/1 

convexus, 18/6 

curvisetus, 20/1 

durieui, 18/2, Ann37 

ericetorum, 18/5 

fascicularis, 18/10, Ann35 

handelii, Ann42 

hungaricus, 18/3, Ann38 

krausei, 18/4, Ann39 

maroccanus, 18/3 

mouretii, 18/11, Ann41 

muhlenbergii, 18/7 

mustaphae, 18/2 

obtusus, 18/5 

pallescens, 18/2 

physcomitroides, 18/2 

pulchellus, 18/8 

schimperi, 18/9, Ann40 

subpallescens, Ann34 

templetonii, 18/1 

Ephemerum, 81 

cohaerens, 81/1 

hibernicum, 81/2, Ann142 

minutissimum, 81/3, Ann143 

recurvifolium, 81/4 

serratum, 81/5, Ann143 

sessile, 81/6 

spinulosum, 81/7, Ann144 


Epipterygium, 145 

tozeri, 145/1 

Eriopus 

apiculatus, 162/1 

Eucladium, 82


verticillatum, 82/1, Ann145 

Eurhynchiastrum, 222 

pulchellum, 222/1 

Eurhynchium, 208 

angustirete, 208/1 

crassinervium, 214/1 

flotowianum, 219/1 

hians, 215/1 

meridionale, 207/1 

praelongum, 216/1 


pumilum, 215/2 


speciosum, 215/4 

stokesii, 216/1 

striatulum, 207/2 

striatum, 208/2 

Fabronia, 226 

ciliaris, 226/1 

pusilla, 226/2 

Fissidens, 41 

adianthoides, 41/1 

algarvicus, 41/18 

arcticus, 41/12, Ann93 

arnoldii, 41/13, Ann99 

asplenioides, 41/10 

azoricus, 41/2 

bambergeri, 41/31, Ann108 

bryoides, 41/14, Ann94, Ann95, Ann97, 

Ann102, Ann103, Ann105, Ann107 

celticus, 41/32 

coacervatus, 41/15 

crassipes, 41/16 

crispus, 41/17, Ann96, Ann108 

cristatus, 41/3 

curnovii, 41/14, Ann94 

curvatus, 41/18 


dubius, 41/3, Ann88 

exiguus, 41/26, 41/31, Ann92, Ann94, Ann103 

exilis, 41/33 

fontanus, 41/11 

gracilifolius, 41/19 

grandifrons, 41/4 

gymnandrus, 41/20, Ann97 

haraldii, 41/31 

herzogii, 41/17 

incurvus, 41/31, Ann94 

jansenii, 41/21, Ann98 

kosaninii, Ann92, Ann94 

limbatus, 41/17, Ann96, Ann108 

luisieri, 41/5, Ann89 

marginatulus, 41/31, Ann109 

microstictus, 41/22, Ann99 

minutulus, 41/17, Ann96 

monguillonii, 41/23 

mouretii, 41/16 

nobreganus, 41/24, Ann100 

osmundoides, 41/6 

ovatifolius, 41/25, Ann101 

pallidicaulis, 41/9 

papillosus, 41/34, Ann110 

perssonii, 41/25, Ann101 

polyphyllus, 41/7, Ann90 

pusillus, 41/26, Ann102, Ann103 

rivularis, 41/27 

rufulus, 41/28, Ann98 

serratus, 41/34, Ann110 

serrulatus, 41/8, Ann91 

sublimbatus, 41/29, Ann104 

sublineaefolius, 41/30, Ann100 

taxifolius, 41/9 

viridulus, 41/31, Ann92, Ann94, Ann103, 

Ann105, Ann107, Ann108, Ann109 

Fontinalis, 169 

antipyretica, 169/1, Ann324, Ann325 

bryhnii, 169/1, Ann325 

dalecarlica, 169/2 

dichelymoides, 169/3, Ann326 

hypnoides, 169/4 

kindbergii, 169/1 

squamosa, 169/5, Ann324 

Forsstroemia 

canariensis, 275/1 

Funaria, 19 

6hybrida, Ann35 

aequidens, 19/1 

algieriensis, 18/9 

anomala, Ann42 

arctica, 19/2 

attenuata, 18/1 

calcarea, 18/7 

convexa, 18/6 

curviseta, 20/1 

dentata, 18/7 

durieui, 18/9, Ann40 

fascicularis, 18/10 

fritzei, Ann39 

handelii, Ann42 

hibernica, 18/7 

hungarica, 18/3 

hygrometrica, 19/3, Ann35 

krausei, 18/4 

maireana, Ann42 

mediterranea, 18/7 

microstoma, 19/4 


muhlenbergii, 18/7 

obtusa, 18/5 


pulchella, 18/8 

pustulosa, 18/6 

subpallescens, Ann34 

Funariella, 20 


Geheebia 

gigantea, 104/10 

Gigaspermum, 26 


Glyphomitrium, 60 

daviesii, 60/1 

Goniomitrium, 21 

acuminatum, Ann44 

seroi, 21/1, Ann44 

Gradsteinia 

torrenticola, 209/5, Ann356 

Grimmia, 30 

affinis, 30/21 

alpestris, 30/1, Ann50, Ann63 

anodon, 30/2 

anomala, 30/3 


arenaria, 30/4, Ann52 

atrata, 30/5 

austrofunalis, 30/36, Ann65 

britannica, 30/36, Ann66 


capillata, 30/7, Ann53 

crassifolia, 30/34, Ann64 

crinita, 30/8, Ann53 

curvata, 30/31 

curviseta, 30/9, Ann54 


dissimulata, 30/11, Ann55 

donniana, 30/12, Ann52, Ann68 

elatior, 30/13, Ann56 

elongata, 30/14 

funalis, 30/15, Ann57 

fuscolutea, 30/16 

grisea, 30/32, Ann62 


holleri, 30/16 

incurva, 30/18 

laevigata, 30/19 

limprichtii, 30/2, Ann51 

lisae, 30/20, Ann58 

longirostris, 30/21 

maritima, Ann79 

meridionalis, 30/36, Ann67 

meteorae, 30/25 

mollis, 30/22 

montana, 30/23 

muehlenbeckii, 30/24, Ann59 

nutans, 30/25 

orbicularis, 30/26 

ovalis, 30/27 

pilosissima, 30/23 

pitardii, 35/1 

plagiopodia, 30/28 

poecilostoma, 30/29, Ann59, Ann64 

pulvinata, 30/30, Ann60

pyrenaica, 30/6 

ramondii, 30/31 

reflexidens, 30/32, Ann61, Ann62, Ann63 

retracta, 30/20, Ann58 

robusta, Ann66 

sessitana, 30/32 

subsulcata, 30/32, Ann63 

sudetica, 30/12 

teretinervis, 30/33 

tergestina, 30/34, Ann51, Ann59, Ann64 

torquata, 30/35 

trichophylla, 30/36, Ann59, Ann66, Ann67 

triformis, 30/37, Ann68 

ungeri, 30/38, Ann69 

unicolor, 30/39 

Guembelia 

laevigata, 30/19 

longirostris, 30/21 

ovalis, 30/27 

tergestina, 30/34 

Gymnostomum, 83 

aeruginosum, 83/1, Ann147 

boreale, 83/2 


lanceolatum, 83/4, Ann146 

mosis, Ann146 

recurvirostrum, 85/1 

viridulum, 83/5 

Gyroweisia, 84 

reflexa, 84/1 

tenuis, 84/2 

Habrodon, 240 

leucotrichus, Ann398 

perpusillus, 240/1 

Hageniella, 262 

micans, 262/1, Ann411 

pacifica, Ann411 

Hamatocaulis, 187 

lapponicus, 187/1 

vernicosus, 187/2 

Haplocladium, 192 


microphyllum, 192/2 


Haplodontium 

notarisii, 141/2 

Haplohymenium 

triste, 277/5 

Hedwigia, 132 

ciliata, 132/1, Ann253, Ann254 

integrifolia, 132/2 

stellata, 132/3, Ann253, Ann255 

Hedwigidium 

integrifolium, 132/2 

Helicodontium, 212 

capillare, 212/1, Ann363 

italicum, 212/1, Ann363 


Helodium, 200 

blandowii, 200/1 

Hennediella, 105 

antarctica, Ann190 

heimii, 105/1 

macrophylla, 105/2, Ann190 

stanfordensis, 105/3 

Herzogiella, 250 

adscendens, Ann400 

seligeri, 250/1 

striatella, 250/2 

turfacea, 250/3 

Heterocladium, 241 

dimorphum, 241/1 

flaccidum, 241/2, Ann396 

heteropterum, 241/3, Ann396 

wulfsbergii, 241/4, Ann397 

Heterophyllium, 259 

affine, 259/1 

Hilpertia, 106 

velenovskyi, 106/1 

Homalia, 269 

besseri, 270/1 

complanata, 270/3 


subrecta, 269/1 

trichomanoides, 269/2, Ann417 

webbiana, 269/3 

Homalothecium, 224 

ureum, 224/1 

geheebii, 220/7 


philippeanum, 224/3 

sericeum, 224/4 

Homomallium, 233 

incurvatum, 233/1 

Hookeria, 165 

lucens, 165/1 

Hydrogrimmia 

mollis, 30/22 

Hygroamblystegium, 178 

fluviatile, 178/1 

humile, 178/2 

tenax, 178/3 

varium, 178/4, Ann329, Ann330 

Hygrohypnella 

duriuscula, 179/4 

ochracea, 179/10 

polaris, 179/11 

Hygrohypnum, 179 


alpinum, 179/2 

cochlearifolium, 179/3 

dilatatum, 179/4 

duriusculum, 179/4 

eugyrium, 179/5 

luridum, 179/6, Ann332 

micans, 262/1 

molle, 179/7 


norvegicum, 179/9 

ochraceum, 179/10 

polare, 179/11 

mithii, 179/12, Ann331 

styriacum, 179/13 

subeugyrium, 179/14 

Hylocomiastrum, 244 

pyrenaicum, 244/1 

umbratum, 244/2 

Hylocomium, 245 

alaskanum, Ann399 

brevirostre, 246/1 

pyrenaicum, 244/1 

splendens, 245/1, Ann399 

umbratum, 244/2 

Hymenoloma 


crispulum, 59/3 

Hymenostomum 

microstomum, 96/1 

squarrosum, 96/10 

Hymenostylium, 85 

insigne, 85/1 

recurvirostrum, 85/1, Ann148 

Hyocomium, 234 

armoricum, 234/1 

Hyophila, 86 

involuta, 86/1 


machadoana, 100/3 

treleasei, 95/2, Ann159 

Hypnum, 235 

aemulans, Ann392 

andoi, 235/1 

bambergeri, 235/2 

callichroum, 235/3 

cupressiforme, 235/4, Ann390, Ann391 

dolomiticum, 235/14 

fertile, 235/5 

hamulosum, 235/6, Ann392 

heseleri, 235/4, Ann391 

holmenii, 235/7, Ann393 

imponens, 235/8 

jutlandicum, 235/9 

lacunosum, 235/4, Ann390 

lindbergii, 230/2, Ann386 

mammillatum, 235/1 


plicatulum, 235/11 

pratense, 228/1, Ann386 

procerrimum, 235/12 

recurvatum, 235/13 

resupinatum, 235/4 

revolutum, 235/14


sauteri, 235/15 

subimponens, 235/16, Ann394 

thedenii, Ann378 

uncinulatum, 235/17 

vaucheri, 235/18 

Hypopterygium, 160 

muelleri, 160/1, Ann320 

tamarisci, 160/1, Ann320 

Imbribryum, 142 

alpinum, 142/2 


Indusiella, 31 

thianschanica, 31/1, Ann70 

Isopterygiopsis, 251 

alpicola, 251/1 

muelleriana, 251/2 

pulchella, 251/3 

Isopterygium, 260 

alpicola, 251/1 

bottinii, 260/1, Ann410 


muellerianum, 251/2 


tenerum, 260/1, Ann410 

Isothecium, 275 

algarvicum, 275/1, Ann426 

alopecuroides, 275/2 

atlanticum, 275/1 

canariense, 275/1 

circinans, 275/2 

holtii, 275/3, Ann427 

myosuroides, 275/4, Ann427 

myurum, 275/2 


viviparum, 275/2 

Iwatsukiella, 242 

leucotricha, 242/1, Ann398 

Jaffueliobryum, 32 

latifolium, 32/1, Ann72 

Kiaeria, 61 

blyttii, 61/1 

falcata, 61/2 

glacialis, 61/3 

riparia, 61/4 

starkei, 61/5 

Kindbergia, 216 

praelonga, 216/1 

Lepidopilum 

virens, 166/2 

Leptobarbula, 87 

berica, 87/1 

Leptobryum, 125 

pyriforme, 125/1 

Leptodictyum, 180 

humile, 178/2 

kochii, 178/2 

kurdicum, Ann333 


Leptodon, 274 

longisetus, 273/1, Ann425 

smithii, 274/1 

Leptodontium, 107 

flexifolium, 107/1 

gemmascens, 107/2 

styriacum, 107/3 

Leptophascum, 108 

leptophyllum, 108/1 

Leptotheca, 158 

gaudichaudii, 158/1, Ann318 

Lescuraea, 193 

incurvata, 196/2 

mutabilis, 193/1 

patens, 196/3 

plicata, 198/1 

radicosa, 196/4 

saviana, 196/5 

saxicola, 193/2 

secunda, 193/3, Ann341 

Leskea, 194 

polycarpa, 194/1 

Leskeella 

incrassata, 197/2, Ann346 

nervosa, 197/2 

Leucobryum, 74 

albidum, 74/1, 74/3, Ann137 

glaucum, 74/2 

juniperoideum, 74/3, Ann137 

minus, 74/3 

Leucodon, 267 

canariensis, 267/1, Ann416 

flagellaris, 267/2, Ann414 

immersus, 267/3 

pendulus, 267/4, Ann415 

sciuroides, 267/5 

treleasei, 267/6, Ann416 

Limneria 

viridula, 104/22 

Limprichtia 


intermedia, 189/1 


Lindbergia, 195 

brachyptera, 195/1, Ann342 

Loeskeobryum, 246 

brevirostre, 246/1 

Loeskypnum, 188 

badium, 188/1 

Meesia, 126 

hexasticha, 126/1 

longiseta, 126/2 

triquetra, 126/3 

uliginosa, 126/4 

Metaneckera 

menziesii, 270/6 

Metzlerella 

alpina, 71/1 

Metzleria 

alpina, 71/1 

Microbryum, 109 

curvicollum, 109/1, Ann192 

davallianum, 109/2, Ann193 

floerkeanum, 109/3 

fosbergii, 109/4, Ann194 


piptocarpum, 109/1 

rectum, 109/6 

starckeanum, 109/7, Ann193, Ann195 

vlassovii, 111/2 

Microcampylopus, 75 

laevigatus, 75/1 

Micromitrium, 88 

tenerum, 88/1 

Microthuidium 


Mielichhoferia, 146 

caucasia, 142/13 

elongata, 146/1, Ann303 

himalayana, 142/13 

mielichhoferi, 146/2 

mielichhoferiana, 146/2, Ann303 

paradoxa, 141/3, Ann269 

Mniobryum 

delicatulum, 147/26 


wahlenbergii, 147/28 

Mnium, 149 

ambiguum, 149/5, Ann313 

blyttii, 149/7 

heterophyllum, 149/8 

hornum, 149/1 

lycopodioides, 149/5, Ann313 

marginatum, 149/6 

spinosum, 149/2 

spinulosum, 149/3 

stellare, 149/9 

thomsonii, 149/4 

Molendoa, 89 

hornschuchiana, 89/1 

schliephackei, 89/2 

sendtneriana, 89/3, Ann141 

taeniatifolia, 89/4 

tenuinervis, 89/5 

warburgii, 89/6 

Myrinia, 225 

pulvinata, 225/1 

Myurella, 252 

julacea, 252/1, Ann401 

sibirica, 252/2 

tenerrima, 252/3 

Myurium, 276 

hebridarum, 276/1

hochstetteri, 276/1 

Myuroclada, 218 

maximowiczii, 218/1, Ann371 

Neckera, 270 

besseri, 270/1 

cephalonica, 270/2 

complanata, 270/3 

crispa, 270/4 


fontinaloides, 270/9 


laeviuscula, 270/5 

longipedunculata, 273/1 

menziesii, 270/6 

oligocarpa, 270/7 

pennata, 270/8 

philippeana, 270/9 

pseudopennata, 270/4 

pumila, 270/9 

subrecta, 269/1 

webbiana, 269/3 

Niphotrichum 

canescens, 33/2 

elongatum, 33/3 

ericoides, 33/4 

panschii, 33/5 

Nobregaea, 211 

latinervis, 211/1, Ann362 

Ochyraea, 181 


cochlearifolia, 179/3 


norvegica, 179/9 

smithii, 179/12 

tatrensis, 181/1, Ann331 

Octoblepharum 

albidum, Ann138 

Octodiceras 

fontanum, 41/11 

Oedipodiella, 27 

australis, 27/1, Ann47 

Oedipodium, 3 

griffithianum, 3/1 

Oligotrichum, 6 

hercynicum, 6/1 

Oncophorus, 62 

compactus, 62/3 

elongatus, 62/1, Ann123 

riparius, 61/4 

virens, 62/2 


Oreas, 63 

martiana, 63/1 

Oreoweisia, 64 

bruntonii, 57/2 

serrulata, 64/1 



Orthodicranum 

flagellare, 69/27 

fulvum, 69/24 


scottianum, 69/25 

tauricum, 69/29 

Orthodontium, 157 

australe, 157/2, Ann317 

gracile, 157/1 

lineare, 157/2, Ann317 

pellucens, 157/3 

Orthogrimmia 


arenaria, 30/4 


donniana, 30/12 


sessitana, 30/32 

triformis, 30/37 

Orthotheciella 

varia, 178/4 

Orthothecium, 253 

chryseon, 253/1, Ann402 

intricatum, 253/2 

lapponicum, 253/3 



Orthotrichum, 128 

acuminatum, 128/34 

affine, 128/35 


anomalum, 128/1 

arcangelianum, 128/31, Ann241 

australe, 128/26, Ann239 

callistomum, 128/28 

casasianum, 128/6, Ann231 

caucasicum, 128/39, Ann244 

consimile, 128/7 

crenulatum, 128/8, Ann232 

cupulatum, 128/2 

diaphanum, 128/9 

elegans, 128/40, Ann245 

fastigiatum, 128/35 

flowersii, Ann232 

gymnostomum, 128/29 

handiense, 128/10, Ann233 

hispanicum, 128/11, Ann234 

ibericum, 128/36, Ann243 

laevigatum, 128/31 

lewinskyae, 128/21, Ann236 

limprichtii, 128/2, Ann230 

lyellii, 128/37 

macrocephalum, 128/12, Ann235 

microcarpum, 128/13 


pallens, 128/14 

paradoxum, 128/14 

patens, 128/15 

pellucidum, 128/3 

philibertii, 128/16 


pumilum, 128/18, Ann106 

pylaisii, 128/38 


rogeri, 128/20, Ann238 

rupestre, 128/32 

scanicum, 128/21 


shawii, 128/33, Ann242 

sordidum, 128/39 

speciosum, 128/40, Ann245, Ann246 

sprucei, 128/23 



striatum, 128/41 


tortidontium, 128/42, Ann247 

urnaceum, Ann229 

urnigerum, 128/4 

vittii, 128/27, Ann240 

vladikavkanum, 128/43, Ann248 

Oxyrrhynchium, 215 

hians, 215/1 

praelongum, 216/1 

pumilum, 215/2 


speciosum, 215/4 

swartzii, 215/1 

Oxystegus, 90 

hibernicus, 90/1 

tenuirostris, 90/2 

Palamocladium, 206 

euchloron, 206/1 

Paludella, 127 

squarrosa, 127/1 

Palustriella, 182 

commutata, 182/1 


falcata, 182/3, Ann334 

pluristratosa, 182/4, Ann334 

Paraleptodontium, 110 


Paraleucobryum, 70 

enerve, 70/1 

longifolium, 70/2 

sauteri, 70/3 

Pelekium, 201 

atlanticum, 201/1, Ann349 


muricatulum, Ann349 

Phascum, 111 

curvicollum, 109/1 


cuynetii, 80/2


floerkeanum, 109/3 

leptophyllum, 108/1 


piptocarpum, 109/1, Ann192 

vlassovii, 111/2 

Philonotis, 138 

arnellii, 138/3 

caespitosa, 138/7 


capillaris, 138/3 

cernua, 138/1 

fontana, 138/9 

hastata, 138/4 

laxa, Ann262 

marchica, 138/5, Ann262 

rigida, 138/2 

seriata, 138/10 

tomentella, 138/11 

uncinata, 138/6, Ann263 

Physcomitrella, 22 

6hampei, Ann35 

patens, 22/1, Ann35 

Physcomitrium, 23 


arenicola, 23/1 

eurystomum, 23/2, Ann35, Ann45 

maroccanum, 18/3 

patens, 22/1 

pyriforme, 23/3 

sphaericum, 23/4 

Pictus 

scoticus, 179/6, Ann332 

Plagiobryum, 143 


zieri, 143/2 

Plagiomnium, 154 

affine, 154/3 

confertidens, 154/8 

curvatulum, 154/4, Ann315 



elatum, 154/5 

ellipticum, 154/6 

medium, 154/7, Ann315 

rostratum, 154/10 

undulatum, 154/9, Ann316 

Plagiopus, 139 

oederi, 139/1 

oederianus, 139/1 

Plagiothecium, 254 

berggrenianum, 254/1 

cavifolium, 254/2, Ann406 

curvifolium, 254/3, Ann403 

denticulatum, 254/4, Ann404 

laetum, 254/5, Ann403 

latebricola, 254/6 

neckeroideum, 254/7, Ann405 

nemorale, 254/8, Ann406 

noricum, 254/7, Ann405 


platyphyllum, 254/10, Ann406 

roeseanum, 254/2 

ruthei, 254/4, Ann404 

succulentum, 254/11, Ann406 

svalbardense, 254/12, Ann407 

undulatum, 254/13 

Plasteurhynchium, 207 

meridionale, 207/1 


Platydictya, 255 


jungermannioides, 255/1 

subtilis, 171/4 

Platygyrium, 261 

repens, 261/1 

Platyhypnidium, 209 


grolleanum, 209/1, Ann354 


mutatum, 209/3, Ann355 

riparioides, 209/4, Ann355 

torrenticola, 209/5, Ann356 

Pleuridium, 47 


alternifolium, 47/2 

nitidum, 48/1 


serrulatum, Ann115 

subulatum, 47/2 

Pleurochaete, 91 

malacophylla, Ann149 


Pleurozium, 247 

schreberi, 247/1 

Pogonatum, 7 

aloides, 7/1, Ann20 

alpinum, 8/1 


dentatum, 7/2, Ann23 

inflexum, Ann19 

nanum, 7/3, Ann20 

neesii, 7/4, Ann19, Ann21 

urnigerum, 7/5, Ann23 

Pohlia, 147 

acuminata, 147/4, Ann288 

ambigua, 147/4, Ann307 

andalusica, 147/10 

andrewsii, 147/11 

annotina, 147/12 


atropurpurea, 147/25 

bolanderi, 147/1, Ann306 

bulbifera, 147/13 

camptotrachela, 147/14 

cruda, 147/2 

crudoides, 147/3, Ann305 

cruegeri, 142/3 

defecta, 147/16 


elongata, 147/4, Ann307, Ann309 

erecta, 147/16 

filum, 147/17 

flexuosa, 147/18, Ann305, Ann310, Ann311 

grandiflora, 147/12 

lescuriana, 147/19 

longicolla, 147/5 

ludwigii, 147/20 


marchica, Ann308 

melanodon, 147/26 

muyldermansii, 147/18, Ann311 

nutans, 147/6, Ann308 

obtusifolia, 147/7 

polymorpha, Ann288 

proligera, 147/22 

ramannii, Ann304 

saprophila, 147/8, Ann309 



scotica, 147/23 

sphagnicola, 147/9 

tundrae, 147/24, Ann312 

turonensis, Ann308 

vexans, 147/27 


Polytrichastrum, 8 

alpinum, 8/1, Ann23 

formosum, 8/2 


norwegicum, 8/1 

pallidisetum, 8/4 

sexangulare, 8/5, Ann24 

sphaerothecium, 8/6, Ann24 

Polytrichum, 9 

affine, 9/6 

alpestre, 9/6 

alpinum, 8/1 

commune, 9/1, Ann25 

formosum, 8/2 

hyperboreum, 9/2 

jensenii, 9/3 

juniperinum, 9/4 



perigoniale, 9/1 


sexangulare, 8/5 

sphaerothecium, 8/6 

strictum, 9/6 

swartzii, 9/7 

uliginosum, 9/8

Pottia, 117 

6andalusica, 109/4, Ann194 


caespitosa, 92/1 

caucasica, Ann217 

commutata, 109/2 

conica, 109/2 

crinita, 117/35 

cuneifolia, 117/24, Ann224 

davalliana, 109/2 

heimii, 105/1 


lanceolata, 117/15 

mutica, 109/7 

pallida, 117/24, Ann224 

recta, 109/6 

starckeana, 109/7 

truncata, 117/32 

wilsonii, 117/36 

Pottiopsis, 92 

caespitosa, 92/1, Ann158 

Protobryum, 112 


Pseudephemerum, 48 

axillare, 48/1 

nitidum, 48/1 

Pseudobryum, 155 

cinclidioides, 155/1 

Pseudocalliergon, 183 

angustifolium, 183/1, Ann335 

brevifolium, 183/2, Ann336 

lycopodioides, 183/3 

trifarium, 183/4 

turgescens, 183/5 

Pseudocrossidium, 113 

hornschuchianum, 113/1 

obtusulum, 113/2, Ann196 

replicatum, 113/3, Ann197 

revolutum, 113/4, Ann196 

Pseudohygrohypnum 

eugyrium, 179/5 

subeugyrium, 179/14 

Pseudoleskea, 196 

artariae, 196/1 

brachyclados, Ann344 

incurvata, 196/2, Ann344 

patens, 196/3 

plicata, 198/1 

radicosa, 196/4, Ann344 

saviana, 196/5 

Pseudoleskeella, 197 

catenulata, 197/1 

nervosa, 197/2, Ann346 

papillosa, 197/3 

rupestris, 197/4, Ann347 

sibirica, 197/4, Ann347 

tectorum, 197/5, Ann361 


Pseudoleskeopsis 

artariae, 196/1 

Pseudoscleropodium, 204 

purum, 204/1 

Pseudotaxiphyllum, 256 


laetevirens, 256/2, Ann408 

Psilopilum, 10 

cavifolium, 10/1 

laevigatum, 10/2 

Pterigynandrum, 243 


Pterogonium, 268 


Pterygoneurum, 114 

compactum, 114/1, Ann198 

crossidioides, 114/2, Ann199 

kozlovii, 114/3 

lamellatum, 114/4 

ovatum, 114/5 

papillosum, 114/6, Ann200 

sampaianum, 114/7 

squamosum, 114/8, Ann201 

subsessile, 114/9 

Ptilium, 236 

crista-castrensis, 236/1 

Ptychodium, 198 

plicatum, 198/1 

Ptychomitrium, 36 

incurvum, 36/1 

nigrescens, 36/2 

polyphyllum, 36/3 

Pylaisia, 237 

polyantha, 237/1 

selwynii, 237/2 

Pylaisiella 

polyantha, 237/1 

selwynii, 237/2 

Pyramidula, 24 

algeriensis, 24/1 

tetragona, 24/1, Ann46 

Racomitrium, 33 

aciculare, 33/6 

affine, 33/10 

aquaticum, 33/7 


ellipticum, 33/11 

elongatum, 33/3 

ericoides, 33/4 

fasciculare, 33/8 

hespericum, 33/9, Ann73 

heterostichum, 33/12 

himalayanum, 33/13 

lamprocarpum, 33/14 

lanuginosum, 33/1 

lusitanicum, 33/15, Ann74 


microcarpon, 33/17 

obtusum, 33/18 

panschii, 33/5 

sudeticum, 33/19 

Rhabdoweisia, 65 

crenulata, 65/1 

crispata, 65/2, Ann124 

denticulata, 65/2 

fugax, 65/3 

kusenevae, 65/2, Ann124 

striata, 65/3 

Rhamphidium, 49 

purpuratum, 49/1, Ann116 

Rhizomnium, 153 

andrewsianum, 153/1 


magnifolium, 153/3 

pseudopunctatum, 153/4 

punctatum, 153/5 

Rhodobryum, 144 

ontariense, 144/1, Ann301 

roseum, 144/2 

spathulatum, 144/1, Ann301 

Rhynchostegiella, 213 

bourgaeana, 213/1, Ann364 



durieui, 213/3 

jacquinii, 213/7, Ann367 

letourneuxii, 213/2 

litorea, 213/4, Ann365 

macilenta, 213/5, Ann366 

pallidirostris, 215/2 

pumila, 215/2 

teesdalei, 213/7, Ann367 

tenella, 213/6, Ann364, Ann365 

teneriffae, 213/7, Ann367 

tenuicaulis, 213/8, Ann368 

trichophylla, 213/9, Ann369 

Rhynchostegium, 210 


arcticum, 210/1, Ann357 

confertum, 210/2, Ann358 


megapolitanum, 210/3, Ann359 

murale, 210/4 

riparioides, 209/4 

rotundifolium, 210/5 

strongylense, 210/6 

surrectum, 210/2, Ann358 

Rhytidiadelphus, 248 

loreus, 248/1 

squarrosus, 248/2 

subpinnatus, 248/3 

triquetrus, 248/4 

Rhytidiastrum 

squarrosum, 248/2


subpinnatum, 248/3 

Rhytidium, 249 

rugosum, 249/1 

Rosulabryum 



laevifilum, 142/37 


Saelania, 50 

glaucescens, 50/1 

Sanionia, 184 

georgicouncinata, 184/1 

nivalis, 184/1 

orthothecioides, 184/2 

uncinata, 184/3 

Schistidium, 34 

agassizii, 34/1, Ann77 

alpicola, 34/25, Ann77 

andreaeopsis, Ann75 

apocarpum, 34/2, Ann76 

atrofuscum, 34/3 

boreale, 34/4 

brunnescens, 34/5 

bryhnii, 34/6 

confertum, 34/7 

confusum, 34/8 

crassipilum, 34/9 

crenatum, 34/10 

dupretii, 34/11 

elegantulum, 34/12 

flaccidum, 34/13 

flexipile, 34/14 

frigidum, 34/15 

frisvollianum, 34/16 

grande, 34/17 

grandirete, 34/18 

helveticum, 34/19, Ann78 

holmenianum, 34/20 

lancifolium, 34/21 

marginale, Ann75 

maritimum, 34/22 

occidentale, 34/23, Ann80 

papillosum, 34/24 

platyphyllum, 34/25, Ann77 

poeltii, 34/26 

pratense, Ann75 

pruinosum, 34/27 

pulchrum, 34/28 

recurvum, 34/29 


robustum, 34/31, Ann66 

scabrum, Ann75 

scandicum, 34/32 

sinensiapocarpum, 34/33, Ann81 

singarense, 34/19, Ann78 

sordidum, 34/34 

spinosum, 34/35, Ann82 


subflaccidium, Ann75 

subjulaceum, 34/37 

submuticum, 34/38 

tenerum, 34/39 

trichodon, 34/40 

umbrosum, 34/41 

venetum, 34/42 

Schistostega, 66 

pennata, 66/1 

Schizymenium, 148 

pontevedrense, 148/1 

Sciuro-hypnum, 219 

flotowianum, 219/1 

glaciale, 219/2 


oedipodium, 219/4 


plumosum, 219/6 

populeum, 219/7 

reflexum, 219/8 

starkei, 219/9 

Scleromnium 

knyi, 272/1 

Scleropodium, 221 

apiculigerum, 219/5 

cespitans, 221/1, Ann382 


purum, 204/1 

touretii, 221/2, Ann32 

Scopelophila, 78 

cataractae, 78/1 

ligulata, 78/2 

Scorpidium, 189 



scorpioides, 189/3 

turgescens, 183/5 

Scorpiurium, 205 

circinatum, 205/1 

deflexifolium, 205/2 

sendtneri, 205/3 

Seligeria, 39 

acutifolia, 39/1 


austriaca, 39/13 

brevifolia, 39/2 


calycina, 39/9 

campylopoda, 39/5 

carniolica, 39/14 

diversifolia, 39/6 

donniana, 39/10 

galinae, 39/3, Ann86 

irrigata, 39/15, Ann87 

oelandica, 39/16 

patula, 39/17 

paucifolia, 39/9 

polaris, 39/11 

pusilla, 39/4 

recurvata, 39/7 

subimmersa, 39/12 

transylvanica, 38/1, Ann84 

trifaria, 39/18 

tristichoides, 39/19 

Sematophyllum, 263 

adnatum, 263/1, Ann412 

bottinii, 260/1 


micans, 262/1 

substrumulosum, 263/3 

Serpoleskea 


subtilis, 171/4 

Sphagnum, 1 

affine, 1/1 

andersonianum, 1/44, Ann14 

angermanicum, 1/35 


annulatum, 1/10 

aongstroemii, 1/34, Ann8 


auriculatum, 1/24, Ann5 

austinii, 1/2 

balticum, 1/11 

brevifolium, 1/13, Ann3 

capillifolium, 1/37, Ann10, Ann11 

centrale, 1/3 


contortum, 1/25 

cuspidatum, 1/12, Ann2 

denticulatum, 1/24, Ann5 

fallax, 1/13, Ann3 

fimbriatum, 1/38 

flexuosum, 1/14 

fuscum, 1/39 

girgensohnii, 1/40 

imbricatum, Ann1 

inundatum, 1/26, Ann5 

isoviitae, 1/13, Ann3 

jensenii, 1/15 

lenense, 1/16, Ann4 

lindbergii, 1/17 

magellanicum, 1/4 

majus, 1/18 

molle, 1/41 

nemoreum, 1/37 

nitidulum, Ann9 

obtusum, 1/19 

olafii, 1/42, Ann12 

orientale, Ann1 

palustre, 1/5 

papillosum, 1/6 

perfoliatum, Ann1

platyphyllum, 1/27 

pulchrum, 1/20 

pylaesii, 1/28 

quinquefarium, 1/43 

recurvum, Ann3 


rubellum, 1/44, Ann10, Ann14 

rubiginosum, 1/45, Ann13 

russowii, 1/46 

skyense, 1/47 

squarrosum, 1/30 

steerei, Ann1 

strictum, 1/8 

subfulvum, 1/48 

subnitens, 1/49 

subsecundum, 1/29 

subtile, Ann10 


tenerum, 1/37, Ann11 

teres, 1/31 

troendelagicum, 1/23 

tundrae, 1/32, Ann6 

viride, 1/12, Ann2 

warnstorfii, 1/50 

wulfianum, 1/33, Ann7 

Splachnobryum, 93 

delicatulum, 93/1, Ann150 

obtusum, 93/1, Ann150 

Splachnum, 120 

ampullaceum, 120/1 

luteum, 120/2 

melanocaulon, 120/3 

pensylvanicum, 120/4 

rubrum, 120/5 

sphaericum, 120/6 

vasculosum, 120/7 

Sporledera 


Stegonia, 115 

latifolia, 115/1 

Stereodon 

bambergeri, 235/2 

callichrous, 235/3 

fertilis, 235/5 

hamulosus, 235/6 

holmenii, 235/7 


plicatulus, 235/11 

revolutus, 235/14 

vaucheri, 235/18 

Straminergon, 190 


Streblotrichum 

convolutum, 99/3 

Streptocolea 

atrata, 30/5 

Syntrichia, 116 


abranchesii, 116/3 

amplexa, 117/1 

bogotensis, 116/1, Ann203 

bolanderi, 117/5 

calcicola, 116/2, Ann204 

caninervis, 116/3, Ann205, Ann206, Ann207 

densa, 116/2 

echinata, 116/4 


glabra, 116/6, Ann208 

handelii, 116/7 

inermis, 117/13 


laevipila, 116/8, Ann209 


minor, 116/10, Ann210 


mutica, 116/5 


pagorum, 116/8, Ann209 

papillosa, 116/13 

papillosissima, 116/14 

princeps, 116/15, Ann213 

pseudohandelii, Ann205 

rigescens, 116/16 

ruraliformis, 116/17 

ruralis, 116/17, Ann204, Ann214 

sinensis, 116/18 

subpapillosissima, 116/19, Ann215 

virescens, 116/20 

Taxiphyllum, 238 

densifolium, 238/1 

wissgrillii, 238/2 

Tayloria, 121 

acuminata, 121/1 

froelichiana, 121/2 

hornschuchii, 121/3 

lingulata, 121/4 

rudolphiana, 121/5 

serrata, 121/6 

splachnoides, 121/7 

tenuis, 121/8 

Tetraphis, 11 

pellucida, 11/1 

Tetraplodon, 122 

angustatus, 122/1 

blyttii, 122/2 

mnioides, 122/3, Ann228 

pallidus, 122/4 

paradoxus, 122/5 

urceolatus, 122/6, Ann228 

Tetrastichium, 166 

fontanum, 166/1 

virens, 166/2 

Tetrodontium, 12 

brownianum, 12/1 

ovatum, 12/2 

repandum, 12/3 

Thamnium 


cossyrense, Ann353 

Thamnobryum, 271 

alopecurum, 271/1, Ann421 



cataractarum, 271/3, Ann419 

cossyrense, 205/3 

fernandesii, 271/4, Ann420 

maderense, 271/5, Ann421 

mediterraneum, 271/1 

neckeroides, 271/6, Ann422 

rudolphianum, 271/7, Ann423 

Thuidiopsis, 202 

sparsa, 202/1, Ann350 

Thuidium, 203 

abietinum, 199/1 


assimile, 203/1, Ann351 

atlanticum, 201/1, Ann349 

blandowii, 200/1 

delicatulum, 203/2, Ann351 

erectum, 203/2 


philibertii, 203/1, Ann351 

recognitum, 203/3 

tamariscinum, 203/4 


Timmia, 15 

austriaca, 15/1 

bavarica, 15/2 

comata, 15/4 

megapolitana, 15/3 


sibirica, 15/6 

Timmiella, 77 

anomala, 77/1 

barbuloides, 77/2 

flexiseta, 77/3 

Tomentypnum, 185 

nitens, 185/1 

Tortella, 94 

alpicola, 94/1, Ann152 

arctica, Ann151 

bambergeri, 94/2, Ann153 

cirrifolia, 94/10, Ann157 

densa, 94/6, Ann154 

flavovirens, 94/3 


glareicola, 94/3 


inclinata, 94/6, Ann154 

inflexa, 94/7 

limbata, 94/8, Ann155 

limosella, 94/9, Ann156


nitida, 94/10, Ann157 

rigens, 94/11 

tortuosa, 94/12 

Tortula, 117 

abranchesii, 116/3 

acaulon, 111/1 

amplexa, 117/1 

ampliretis, 117/2, Ann218 

angustata, 117/28, Ann226 

atherodes, 111/1 

atrovirens, 117/3 

baetica, 117/14 

bogosica, 117/4 

bolanderi, 117/5, Ann219 

brevissima, 117/6, Ann221 

buyssonii, Ann216 

calcicolens, 116/2 


caninervis, 116/3 

caucasica, Ann217 

cernua, 117/8 

crinita, 116/11 

cuneifolia, 117/9 

densa, 116/2 

echinata, 116/4 

eucalyptrata, 117/12 

euryphylla, 117/12 


freibergii, 117/10 

grandiretis, Ann217 

guepinii, 117/11 

handelii, 116/7 

hirsuta, 116/14 

hoppeana, 117/12, Ann222 

inermis, 117/13, Ann219 

intermedia, 116/11, Ann211 

israelis, 117/14, Ann220 

laevipila, 116/8 

laevipilaeformis, 116/8, Ann209 

lanceola, 117/15 


laureri, 117/16 

leucostoma, 117/17 

lingulata, 117/18 

marginata, 117/19, Ann227 

modica, 117/20 

mucronifolia, 117/21 

muralis, 117/22, Ann220 


obtusifolia, 117/23 

pagorum, 116/8 

pallida, 117/24 

papillosa, 116/13, Ann212 


princeps, 116/15 

protobryoides, 112/1 

randii, 117/25 


rhizophylla, 108/1 

rhodonia, 117/27, Ann225 

rigescens, 116/16 

ruraliformis, 116/17 

ruralis, 116/17, Ann215 

saccardoana, 116/8, Ann209 


sinensis, 116/18 

solmsii, 117/29, Ann227 

spuria, 116/3 

subulata, 117/30, Ann226 

systylia, 117/31 

truncata, 117/32 

ucrainica, 117/33 

vahliana, 117/34 

virescens, 116/20 

viridifolia, 117/35 

wilsonii, 117/36 

zoddae, 117/24, Ann224 

Trachycystis, 150 

immarginata, 150/1 

ussuriensis, 150/1, Ann314 

Trematodon, 53 

ambiguus, 53/1 

brevicollis, 53/2, Ann32 

laetevirens, 53/3 

longicollis, 53/4, Ann32, Ann117 

perssoniorum, 53/5, Ann117 

Trichodon, 51 

cylindricus, 51/1 

Trichostomopsis 

aaronis, 104/3 

australasiae, 104/3 

trivialis, 104/26 

umbrosa, 104/26 

Trichostomum, 95, Ann158 

arcticum, 95/1, Ann151 

brachydontium, 95/2, Ann158 

caespitosum, 92/1 

connivens, Ann158 

contortum, Ann158 

crispulum, 95/3, Ann158 

hibernicum, 90/1 



tenuirostre, 90/2 

triumphans, 95/4, Ann158 

Triquetrella, 118 

arapilensis, 118/1 

Trochobryum 

carniolicum, 39/14 

Ulota, 129 

bruchii, 129/1 

calvescens, 129/2 

coarctata, 129/3 

crispa, 129/4 

curvifolia, 129/5 


hutchinsiae, 129/7 


macrospora, 129/8, Ann249 

phyllantha, 129/9 

rehmannii, 129/10, Ann249 

Vesicularia, 239 

galerulata, Ann395 

reimersiana, 239/1, Ann395 

sphaerocarpa, Ann395 

Voitia, 123 

hyperborea, 123/1 

nivalis, 123/2 

Warnstorfia, 191 

exannulata, 191/1 

fluitans, 191/2 

h-schulzei, 191/2 

procera, 191/3, Ann339 

pseudostraminea, 191/4 

sarmentosa, 191/5 

trichophylla, 191/6 

tundrae, 191/7 

Webera 

luisieri, Ann304 

maderensis, Ann304 

Weissia, 96 

brachycarpa, 96/1 

condensa, 96/2 

controversa, 96/3 

fallax, 96/3 

leptocarpa, Ann160 

levieri, 96/4 

longifolia, 96/5 

mittenii, Ann161 

multicapsularis, 96/6 


perssonii, 96/7 

rostellata, 96/8 

rutilans, 96/9 


sterilis, 96/11 

triumphans, 95/4 

tyrrhena, 96/12 

wimmeriana, 96/13, Ann162 

Zygodon, 130 

baumgartneri, 130/5 

conoideus, 130/1 

dentatus, 130/2 

forsteri, 130/3, Ann250 

gracilis, 130/4 

madeirensis, 130/3, Ann250 

rupestris, 130/5 

sibiricus, 130/6, Ann251 

stirtonii, 130/7, Ann252 

viridissimus, 130/8 

vulgaris, 130/5


MARK O. HILL, Centre for Ecology and Hydrology, Monks Wood, Abbots Ripton, Huntingdon PE28 2LS, U.K. E-mail: moh@ceh.ac.uk 

NEIL BELL, Botanical Museum, Finnish Museum of Natural History, PO Box 7, FIN-00014 University of Helsinki, Finland. 

E-mail: neil.bell@helsinki.fi 

IDA BRUGGEMAN-NANNENGA, Griffensteijnseplein 23, 3703 BE Zeist, The Netherlands. 

MONTSERRAT BRUGUÉS, Departament de Biologia Animal, Vegetal i Ecologia, Unitat de Botànica, Universitat Autònoma de Barcelona, 

08193 Bellaterra, Barcelona, Spain. E-mail: Montserrat.Brugues@uab.es 

MARíA J. CANO, Departamento de Biología Vegetal (Botánica), Facultad de Biología, Universidad de Murcia, Campus de Espinardo, 30100 

Murcia, Spain. E-mail: mcano@um.es 

JOHANNES ENROTH, Department of Biological and Environmental Sciences and Botanical Museum, PO Box 7, FI-00014 University of 

Helsinki, Finland. E-mail: enroth@touko.helsinki.fi 

KJELL I. FLATBERG, Section of Natural History, Museum of Natural History and Archaeology, Norwegian University of Science and 

Technology, N-9471 Trondheim, Norway. E-mail kjell.flatberg@vm.ntnu.no 

JAN-PETER FRAHM, Nees Institut für Biodiversität der Pflanzen, Meckenheimer Allee 170, D-53115 Bonn, Germany. 

E-mail: frahm@uni-bonn.de 

MARíA TERESA GALLEGO, Departamento de Biología Vegetal (Botánica), Facultad de Biología, Universidad de Murcia, Campus de 

Espinardo, 30100 Murcia, Spain. 

RICARDO GARILLETI, Departamento de Botánica, Facultad de Farmacia, Universidad de Valencia, Avda. Vicente Andrés Estellés s/n, 

E-46100 Burjasot, Spain 

JUAN GUERRA, Departamento de Biología Vegetal (Botánica), Facultad de Biología, Universidad de Murcia, Campus de Espinardo, 30100 

Murcia, Spain. E-mail: jguerra@um.es 

LARS HEDENÄS, Swedish Museum of Natural History, Department of Cryptogamic Botany, Box 50007, SE-104 05, Stockholm, Sweden. 

E-mail: lars.hedenas@nrm.se 

DAVID. T.HOLYOAK, 8 Edward Street, Tuckingmill, Camborne, Cornwall TR14 8PA, UK. E-mail: david@holyoak9187.fsnet.co.uk 

JAAKKO HYVÖNEN, Plant Biology, PO Box 65 and Botanical Museum, PO Box 7, FIN-00014 University of Helsinki, Finland. 

E-mail: jaakko.hyvonen@helsinki.fi 

MICHAEL S. IGNATOV, Main Botanical Garden, Russian Academy of Sciences, Botanicheskaya 4, 127276 Moscow, Russia. 

E-mail: misha_ignatov@list.ru 

FRANCISCO LARA, Departamento de Biología (Botánica), Facultad de Ciencias, Universidad Autónoma de Madrid, C/Darwin, 2, E-28049 

Madrid, Spain. E-mail: francisco.lara@uam.es 

VICENTE MAZIMPAKA, Departamento de Biología (Botánica), Facultad de Ciencias, Universidad Autónoma de Madrid, C/Darwin, 2, 

E-28049 Madrid, Spain. E-mail: vicente.mazimpaka@uam.es 

JESÚS MUÑOZ, Real Jardín Botánico (CSIC), Plaza de Murillo 2, E-28014 Madrid, Spain. E-mail: jmunoz@ma-rjb.csic.es 

LARS SÖDERSTRÖM, Department of Biology, Norwegian University of Science and Technology, N-7491 Trondheim, Norway. 

E-mail: lars.soderstrom@bio.ntnu.no

Bryological Monograph An annotated checklist of the mosses of ...

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