69. MENTHA Linnaeus, Sp. Pl. 2: 576. 1753.
69. MENTHA Linnaeus, Sp. Pl. 2: 576. 1753.
69. MENTHA Linnaeus, Sp. Pl. 2: 576. 1753.
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Flora of China 17: 236–239. 1994.<br />
<strong>69.</strong> <strong>MENTHA</strong> <strong>Linnaeus</strong>, <strong>Sp</strong>. <strong>Pl</strong>. 2: <strong>576.</strong> <strong>1753.</strong><br />
薄荷属 bo he shu<br />
Herbs annual or perennial, aromatic, often rhizomatous or stoloniferous. Upper leaves sessile or subsessile; blade margin<br />
dentate, serrate, or crenate. Verticillasters (2–6)- to many flowered; floral leaves similar to stem leaves or reduced; bracts<br />
lanceolate to linear, ± distinct. Flowers bisexual or pistillate. Calyx funnelform to campanulate, 10–13-veined, throat glabrous or<br />
hairy, limb equally 5-toothed or 2-lipped, upper lip 3-toothed, lower lip 2-toothed. Corolla funnelform, ± regular or slightly<br />
irregular; tube generally included, throat slightly dilated or saccate in front, limb 4-lobed; lobes equal, entire, upper lobe slightly<br />
wider, emarginate or 2-lobulate. Stamens 4, subequal, divaricate, erect, exserted in bisexual flowers, included and often<br />
rudimentary in pistillate flowers, posterior 2 slightly longer than anterior 2; filaments glabrous; anther cells 2, parallel. Style<br />
exserted, apex equally 2-cleft. Nutlets ovoid, dry, smooth or slightly tuberculate, apex rounded, rarely hairy.<br />
About 30 species: mainly in north temperate regions, a few in the Southern Hemisphere; six native and six cultivated species in China.<br />
A number of species are widely cultivated for their aromatic properties and many cultivars have been developed.<br />
1a. Calyx tubular or campanulate-tubular, slightly curved, 2-lipped; teeth of upper lip lanceolate-triangular, shorter<br />
than subulate teeth of lower lip; tube hairy at throat inside, closed in fruit, hirtellous outside, conspicuously veined;<br />
corolla saccate at throat in front ............................................................................................................................................................. 12. M. pulegium<br />
1b. Calyx broadly campanulate or funnelform-campanulate, straight, regular; teeth ± equal; tube glabrous at throat,<br />
spreading in fruit, obscurely veined, not hirtellous; corolla throat slightly dilated, not saccate.<br />
2a. Verticillasters widely spaced, inserted in leaf axils, sometimes along entire stem; leaves longer than<br />
verticillasters; corolla throat hairy.<br />
3a. Stems much branched, puberulent at apex and along angles at base; leaves 3–5(–7) cm, sparsely coarsely<br />
dentate-serrate; calyx teeth puberulent; stamens and style usually slightly exserted ........................................ 1. M. canadensis<br />
3b. Stems unbranched or branched at apex, densely pubescent; leaves (2.5–)4–9 cm, irregularly shallow serrate;<br />
calyx teeth pilose; stamens and style usually much exserted ................................................................................... 2. M. sachalinensis<br />
2b. Verticillasters in leafless, terminal, capitate or spicate inflorescences, upper ones contiguous; floral leaves linear<br />
or nearly similar to stem leaves; corolla throat hairy or glabrous inside.<br />
4a. Verticillasters 1–3, in short, capitate, rarely elongated inflorescences; basal verticillasters sometimes remote<br />
from upper, axillary, with large floral leaves; calyx tubular; corolla hairy inside; nutlets glabrous<br />
........................................................................................................................................................................................................................ 3. M. dahurica<br />
4b. Verticillasters in spicate continuous or interrupted inflorescences; floral leaves linear or similar to stem<br />
leaves, generally minute; calyx campanulate; corolla throat glabrous inside; nutlets apex hairy.<br />
5a. Leaves crisped, ovate to ovate-lanceolate, incised-serrate; calyx teeth slightly close together ................... 10. M. crispata<br />
5b. Leaves not crisped; calyx teeth not close together in fruit.<br />
6a. Leaves glabrous or subglabrous, dark green or shiny green.<br />
7a. Adaxial stem leaves sessile or subsessile; inflorescences slender, 4–10 cm, interrupted; plants shiny<br />
green ............................................................................................................................................................................................... 9. M. spicata<br />
7b. Adaxial stem leaves petiolate; inflorescences robust, interrupted at base; plants dark green.<br />
8a. Stem leaves lanceolate to ovate-lanceolate, apex acute; calyx teeth ciliate .......................................... 7. M. ×piperita<br />
8b. Stem leaves ovate to elliptic, apex obtuse; calyx teeth not ciliate ................................................................... 8. M. citrata<br />
6b. Leaves hairy at least abaxially.<br />
9a. All leaves sessile, corrugate, margin crenate or crenate-serrate ................................................................ 11. M. suaveolens<br />
9b. Lower leaves sometimes petiolate, leaves not corrugate when all sessile, not crenate or crenate-serrate.<br />
10a. <strong>Pl</strong>ants densely gray tomentose, much branched ................................................................................................... 6. M. vagans<br />
10b. <strong>Pl</strong>ants not entirely densely gray tomentose, not much branched.<br />
11a. Leaves mostly sessile, coarsely serrate-dentate, dark green or green and hairy, sometimes<br />
abaxially densely appressed tomentose-pubescent; inflorescence robust; pedicel tomentosevillous;<br />
calyx as long as corolla tube, spreading hairy, teeth as long as tube; corolla ca. 4 mm<br />
...................................................................................................................................................................................... 4. M. longifolia<br />
11b. Leaves short petiolate or subsessile, remotely shallow unequal dentate, glaucous, finely<br />
minutely tomentose or crisped tomentose; inflorescences rather slender, sometimes<br />
interrupted; pedicel appressed pubescent; calyx shorter than corolla tube, appressed pubescent,<br />
teeth 3/4 as long as tube; corolla 4–5 mm ..................................................................................................... 5. M. asiatica<br />
1. Mentha canadensis <strong>Linnaeus</strong>, <strong>Sp</strong>. <strong>Pl</strong>. 2: 577. <strong>1753.</strong><br />
薄荷 bo he<br />
Mentha arvensis <strong>Linnaeus</strong> var. canadensis (<strong>Linnaeus</strong>)<br />
Maximowicz; M. arvensis f. chinensis Debeaux; M. arvensis<br />
subsp. haplocalyx (Briquet) Briquet; M. arvensis var.<br />
haplocalyx (Briquet) Briquet; M. haplocalyx Briquet;<br />
M. haplocalyx Briquet f. alba X. L. Liu & X. H. Guo;<br />
M. pedunculata Hu & Tsai.<br />
<strong>Pl</strong>ants rhizomatous, perennial. Stems erect, 30–60 cm,<br />
puberulent, much branched. Petiole 2–10 mm; leaf blade<br />
ovate-lanceolate to oblong, 3–5(–7)×0.8–3 cm, puberulent,<br />
base cuneate to rounded, margin remotely coarse dentateserrate<br />
above base, apex acute. Verticillasters axillary,<br />
globose, ca. 1.8 cm in diam.; peduncle 0–3 mm. Pedicel<br />
slender, 2.5 mm. Calyx tubular-campanulate, ca. 2.5 mm,<br />
puberulent, glandular, obscurely 10-veined; teeth narrowly<br />
triangular-subulate, apex acuminate, ca. 1 mm. Corolla
purplish or white, ca. 4 mm, puberulent; upper lobe largest, 2lobulate;<br />
other lobes subequal, oblong, obtuse. Stamens ca. 5<br />
mm. Nutlets yellow-brown, small pitted. Fl. Jul-Sep, fr. Oct.<br />
Wet areas; 0–3500 m. In all provinces [Cambodia, Japan, Korea,<br />
Laos, Malaysia, Myanmar, Russia, Thailand, Vietnam; North<br />
America].<br />
Source of mint oil. Used in cigarettes, pharmaceuticals, and oral<br />
preparations.<br />
2. Mentha sachalinensis (Briquet ex Miyabe & Miyake)<br />
Kudô, J. Coll. Sci. Imp. Univ. Tokyo 43(10): 47. 1921.<br />
东北薄荷 dong bei bo he<br />
Mentha arvensis <strong>Linnaeus</strong> subsp. haplocalyx Briquet var.<br />
sachalinensis Briquet ex Miyabe & Miyake, Fl. Saghalin 361.<br />
1916; M. arvensis var. piperascens Holmes; M. sachalinensis<br />
(Briquet) Kudô var. arguta Kitagawa.<br />
<strong>Pl</strong>ants rhizomatous, perennial. Stems erect, 50–100 cm,<br />
striate, densely retrorse pubescent along angles, little<br />
branched. Petiole 0.5–1.5 cm; leaf blade elliptic-lanceolate,<br />
(2.5–)4–9×1–3.5 cm, glandular, ciliate, puberulent on veins,<br />
base attenuate, margin shallow serrate, apex acute.<br />
Verticillasters axillary, many flowered, globose, to 1.5 cm in<br />
diam., short pedunculate; floral leaves subsessile,<br />
sublanceolate; bracteoles linear to linear-lanceolate, 3–4 mm,<br />
ciliate. Pedicel ca. 2 mm, glabrous. Calyx campanulate, ca.<br />
1.5 mm, densely pilose, yellow glandular, mouth and teeth<br />
pilose inside; teeth narrowly triangular, ca. 1.5 mm, apex<br />
acute. Corolla purplish or purplish red, ca. 4 mm, pilose; lobes<br />
ovate-oblong, upper emarginate. Anterior stamens slightly<br />
longer, ca. 5 mm. Nutlets yellow-brown, oblong, glabrous,<br />
ribbed. Fl. Jul-Aug, fr. Sep.<br />
Wet grasslands near rivers, lakes; 200–1100 m. Heilongjiang,<br />
Jilin, Liaoning, Nei Mongol [Japan, Russia].<br />
3. Mentha dahurica Fischer ex Bentham, Labiat. Gen. <strong>Sp</strong>ec.<br />
181. 1836.<br />
兴安薄荷 xing an bo he<br />
<strong>Pl</strong>ants rhizomatous, perennial. Stems erect, 30–60 cm,<br />
rarely branched, base leafless, retrorse puberulent, sometimes<br />
purplish. Petiole 7–10 mm; leaf blade ovate to oblong, ca.<br />
3 × 1.3 cm, glabrous or sparsely puberulent, abaxially<br />
glandular on veins, base broadly cuneate to rounded, margin<br />
shallow crenate-serrate to subentire, apex acute to obtuse.<br />
Verticillasters 5–13-flowered, generally 2, in terminal capitate<br />
inflorescences longer than floral leaves; peduncle 2–10 mm,<br />
puberulent; bracteoles linear, curved upward. Pedicel 1–3 mm.<br />
Calyx tubular-campanulate, ca. 2.5 mm, veins conspicuous,<br />
puberulent; teeth broadly triangular, ca. 0.5 mm, apex acute;<br />
fruiting calyx broadly campanulate. Corolla reddish or rose<br />
purplish, ca. 5 mm, glabrous, throat puberulent, gradually<br />
dilated from base; lobes circular, ca. 1 mm, obtuse, upper lobe<br />
conspicuously 2-lobulate. Anterior stamens equal to slightly<br />
longer than corolla. Ovary brown, glabrous. Fl. Jul-Aug.<br />
Meadows; 600 m. Heilongjiang, Jilin, Nei Mongol<br />
[Japan, Russia]<br />
4. Mentha longifolia (<strong>Linnaeus</strong>) Hudson, Fl. Angl. 221. 1762.<br />
欧薄荷 ou bo he<br />
Mentha spicata <strong>Linnaeus</strong> var. longifolia <strong>Linnaeus</strong>, <strong>Sp</strong>. <strong>Pl</strong>.<br />
2: <strong>576.</strong> 1753; M. sylvestris <strong>Linnaeus</strong>, 1763.<br />
<strong>Pl</strong>ants rhizomatous, perennial. Stems erect, to 1 m, much<br />
branched, whitish, striate, retrorse short tomentosevillous.<br />
Leaf blade ovate to oblong-lanceolate, to 6×1.5 cm,<br />
appressed tomentose-villous, subglabrescent, base rounded to<br />
shallow cordate, margin coarsely irregular serrate-dentate,<br />
apex acute. Verticillasters in cylindric terminal spikes 3–<br />
8 cm, lower ones somewhat lax; floral leaves linear-subulate,<br />
mostly shorter than verticillasters. Pedicel to 2 mm. Calyx<br />
campanulate, ca. 2 mm, tomentose-villous, obscurely 5veined;<br />
teeth linear-subulate, ca. 1 mm, close together in fruit.<br />
Corolla purplish, ca. 4 mm, sparsely puberulent; tube ca. 2<br />
mm, gradually dilated upward; lobes oblong, obtuse; upper<br />
lobe oblong-ovate, emarginate. Ovary glabrous. Fl. Jul-Sep.<br />
Nanjing Shi, Shanghai Shi, and other cities [Russia; SW Asia,<br />
Europe].<br />
Cultivated in China. A polymorphic aromatic herb used<br />
medicinally.<br />
Mentha asiatica, M. vagans, and the Himalayan<br />
M. royleana Bentham all are very closely related to and perhaps<br />
doubtfully distinct from M. longifolia. Further work is needed to<br />
fully assess their correct status.<br />
5. Mentha asiatica Borissova-Bekrjasheva, Bot. Mater. Gerb.<br />
Bot. Inst. Komarova Akad. Nauk SSSR 16: 280. 1954.<br />
假薄荷 jia bo he<br />
Herbs rhizomatous, perennial. Stems erect, (30–)50–<br />
120(–150) cm, few branched, densely minutely tomentose.<br />
Petiole 0.1–0.5 mm; leaf blade oblong or elliptic to oblonglanceolate,<br />
3–8×1–2.5 cm, sometimes folded and curved<br />
downward, sometimes glaucous, appressed crisped minutely<br />
tomentose, glandular, base rounded to broadly cuneate,<br />
margin remotely unequal dentate, apex acute. Verticillasters<br />
in terminal cylindric spikes 3–8×1–1.4 cm; bracts linear to<br />
subulate, 5–7 mm; bracteoles subulate, nearly as long as calyx.<br />
Pedicel ca. 1 mm. Calyx ± purple-red, campanulate to<br />
funnelform, 1.5–2 mm, appressed pubescent, obscurely<br />
veined; teeth linear, close together in fruit. Corolla purple-red,<br />
4–5 mm, pilose, tube gradually dilated, lobes ca. 1 mm; upper<br />
lobe oblong-ovate, ca. 2×1.5 mm, apex emarginate. Nutlets<br />
brown, ca. 1 mm, apex pilose, pitted. Fl. Jul-Aug, fr. Aug-Oct.<br />
Riverbanks, landfills, waste areas, wet valleys; 0–3100 m.<br />
Sichuan, Xinjiang, Xizang [Kazakhstan, Kyrgyzstan, Russia,<br />
Tajikistan, Turkmenistan, Uzbekistan; SW Asia].<br />
6. Mentha vagans Borissova-Bekrjasheva, Bot. Mater. Gerb.<br />
Bot. Inst. Komarova Akad. Nauk SSSR 16: 282. 1954.<br />
灰薄荷 hui bo he<br />
<strong>Pl</strong>ants rhizomatous, perennial. Stems erect, 40–80 cm,<br />
densely tomentose, glabrescent, purple-red, base with<br />
exfoliating bark; branches many, long, divaricate. Petiole ca.<br />
1 mm; leaf blade elliptic to oblong, 1–2.5 cm×5–13 mm,<br />
generally shorter than internodes, sometimes curvedconduplicate,<br />
densely gray tomentose, base rounded to<br />
shallow cordate, margin serrate-dentate, apex acute to<br />
subobtuse. Verticillasters in cylindric spikes 2–2.5 cm×ca.<br />
8 mm, lower spikes ± separated; bracts filiform. Pedicel ca.<br />
1 mm. Calyx campanulate, to 2 mm, crisped pilose, obscurely<br />
5-veined; teeth lanceolate, ca. 0.5 mm, apex spinescent, close<br />
together in fruit. Corolla 3–3.5 mm, lobes ovate, subequal,<br />
pilose; upper lobe longer, oblong-ovate, ca. 1.5 mm,<br />
emarginate. Nutlets ovoid, ca. 0.6×0.5 mm, brown, sparsely<br />
hairy, pitted. Fl. Jul-Aug.<br />
Riverbanks. Xinjiang [Tajikistan, Turkmenistan; SW Asia].
7. Mentha ×piperita <strong>Linnaeus</strong>, <strong>Sp</strong>. <strong>Pl</strong>. 2: <strong>576.</strong> <strong>1753.</strong><br />
辣薄荷 la bo he<br />
Stems erect-ascending, 30–100 cm tall, branched,<br />
purple-red, glabrous or angles sparsely minutely hispid.<br />
Petiole 1–2 mm, purple; leaf blade lanceolate to ovatelanceolate,<br />
2.5–3×0.8–2 cm, glabrous or abaxially minutely<br />
hispid on veins, densely glandular, base rounded to shallow<br />
cordate, margin unequally acute serrate, apex acute.<br />
Verticillasters in cylindric terminal spikes, interrupted at base;<br />
bracts linear-lanceolate, slightly longer than calyx, ciliate.<br />
Pedicel ca. 1 mm. Calyx tubular, tinged purple, puberulent or<br />
subglabrous, glandular outside; teeth 5, linear-subulate, ca. 1<br />
mm, ciliate. Corolla white, tinged red on lobes, ca. 4 mm,<br />
tube nearly as long as calyx; lobes subequal, upper emarginate.<br />
Nutlets brown, obovoid, ca. 0.7 mm, apex glandular. Fl. Jul, fr.<br />
Aug.<br />
Beijing Shi, Nanjing Shi, and other cities [India, Japan,<br />
Kyrgyzstan, Russia, Tukmenistan; SW Asia, Europe, North America].<br />
Cultivated in China. Source of peppermint oil, an essential oil<br />
used for flavoring confectionery, candies, liqueur, and chewing gum.<br />
The dried plant is used medicinally as a carminative, nervine, and<br />
stimulant, while menthol is used in nasal sprays.<br />
A hybrid: Mentha aquatica <strong>Linnaeus</strong>× M. spicata <strong>Linnaeus</strong>.<br />
8. Mentha citrata Ehrhart, Beitr. Naturk. 7: 150. 1792.<br />
柠檬留兰香 ning meng liu lan xiang<br />
Mentha ×piperita <strong>Linnaeus</strong> var. citrata (Ehrhart) Briquet.<br />
Herbs perennial, bearing leafy stolons, glabrous or<br />
subglabrous throughout. Petiole of middle stem leaves ca.<br />
2 mm; leaf blade broadly ovate to elliptic, 2–4×1.5–3 cm,<br />
base rounded to shallow cordate, margin remotely serrate,<br />
apex obtuse; upper leaves reduced, sometimes sublanceolate,<br />
apex acute. Verticillasters in terminal spikes 2.5–4 cm; bracts<br />
linear-lanceolate, shorter than verticillasters. Pedicel ca. 1 mm.<br />
Calyx tubular, ca. 1.5 mm, glandular; teeth linear-subulate, ca.<br />
0.5 mm. Corolla purplish, ca. 4 mm, tube ca. 2.5 mm. Fl. Jul-<br />
Aug.<br />
Beijing Shi, Hangzhou Shi, Nanjing Shi, and other cities<br />
[Europe].<br />
Cultivated in China.<br />
Perhaps best regarded as a cultivar of Mentha ×piperita.<br />
9. Mentha spicata <strong>Linnaeus</strong> <strong>Sp</strong>. <strong>Pl</strong>. 2: <strong>576.</strong> <strong>1753.</strong><br />
留兰香 liu lan xiang<br />
Mentha spicata var. viridis <strong>Linnaeus</strong>; M. viridis<br />
(<strong>Linnaeus</strong>) <strong>Linnaeus</strong>.<br />
Herbs stoloniferous. Stems erect, 40–130 cm, glabrous or<br />
subglabrous, striate. Leaves sessile or subsessile, ovateoblong<br />
to oblong-lanceolate, 3–7×1–2 cm, base broadly<br />
cuneate to rounded, margin irregularly serrate, apex acute.<br />
Verticillasters in cylindric terminal spikes 4–10 cm, spikes<br />
interrupted at base; bracteoles linear, 5–8 mm. Pedicel ca.<br />
2 mm. Calyx campanulate, ca. 2 mm, glandular, obscurely 5veined;<br />
teeth triangular-lanceolate, ca. 1 mm. Corolla purplish,<br />
ca. 4 mm, glabrous, tube ca. 2 mm; lobes subequal, apex<br />
emarginate. Ovary brown, glabrous. Fl. Jul-Sep.<br />
Guangdong, Guangxi, Guizhou, Hebei, Hubei, Jiangsu, Sichuan,<br />
Xizang, Yunnan, Zhejiang [Russia, Turkmenistan; Africa, SW Asia;<br />
Europe].<br />
Cultivated in China. Source of spearmint oil, an essential oil<br />
used for flavoring candies, toothpaste, chewing gum. Dried tops and<br />
leaves are used medicinally as a stimulant, carminative, and nervine.<br />
It contains carvone.<br />
10. Mentha crispata Schrader ex Willdenow, Enum.<br />
<strong>Pl</strong>. 608. 1809.<br />
皱叶留兰香 zhou ye liu lan xiang<br />
Herbs rhizomatous. Stems erect, 30–60 cm, purplish,<br />
glabrous. Leaves sessile or subsessile, ovate to ovatelanceolate,<br />
2–3×1.2–2 cm, papery, base rounded to shallow<br />
cordate, margin serrate, apex acute. Verticillasters in terminal<br />
spikes 2.5–3×ca. 1 cm, continuous or ± interrupted at base;<br />
bracts linear-lanceolate, slightly longer than calyx. Pedicel ca.<br />
1 mm. Calyx campanulate, ca. 1.5 mm, subglabrous,<br />
glandular, obscurely 5-veined; teeth triangular-lanceolate, ca.<br />
0.1 mm, ciliate. Corolla purplish, ca. 3.5 mm, glabrous, tube<br />
ca. 2 mm; lobes subequal, apex emarginate. Ovary brown,<br />
glabrous. Nutlets brown, triquetrous, ovoid, ca. 0.7 mm,<br />
sparsely glandular.<br />
Beijing Shi, Hangzhou Shi, Kunming Shi, Nanjing Shi,<br />
Shanghai Shi [Russia; Europe].<br />
Cultivated in China. Source of German spearmint oil. Young<br />
shoots and leaves edible.<br />
A form/cultivar of M. spicata <strong>Linnaeus</strong>.<br />
11. Mentha suaveolens Ehrhart, Beitr. Naturk. 7: 149. 1792.<br />
圆叶薄荷 yuan ye bo he<br />
Herbs perennial, rhizomatous, stoloniferous. Stems erect,<br />
30–80 cm, striate, crisped villous, pyramidally<br />
branched. Leaves usually sessile, circular to oblong-ovate, 2–<br />
4.5×1.5–3 cm, villous, base cordate, margin crenate or<br />
crenate-serrate, apex obtuse. Verticillasters in terminal, dense<br />
cylindric spikes 2–4×ca. 0.6 cm, lower 1 or 2 ± widely spaced;<br />
bracts lanceolate, ca. 3 mm, crisped. Pedicel ca. 1 mm,<br />
glabrous. Calyx broadly campanulate, ca. 2.5 mm, pubescent,<br />
subglobose, dilated in fruit, not constricted at throat,<br />
obscurely veined; teeth lanceolate-subulate, ca. 1 mm. Corolla<br />
white, purplish, bluish or purple, ca. 2.5 mm, glabrous, tube<br />
ca. 1.5 mm; lobes subequal, upper emarginate. Ovary<br />
glabrous. Nutlets unknown.<br />
Beijing Shi, Kunming Shi, Nanjing Shi, Shanghai Shi, Yunnan<br />
(Lijiang Xian) [Europe].<br />
Cultivated in China.<br />
Most widely known as Mentha rotundifolia (<strong>Linnaeus</strong>) Hudson,<br />
but the holotype of that species has been shown to be a different plant<br />
of hybrid origin.<br />
12. Mentha pulegium <strong>Linnaeus</strong>, <strong>Sp</strong>. <strong>Pl</strong>. 2: 577. <strong>1753.</strong><br />
唇萼薄_ chun e bo he<br />
Pulegium vulgare Miller.
Herbs. Stolons with scalelike leaves. Stems ascending,<br />
rarely erect or trailing, 15–30(–50) cm, hirtellous, striate,<br />
tinged red-purple, much branched, internodes generally longer<br />
than leaves. Leaf blade ovate-orbicular to ovate, 0.8–1.3<br />
cm×5–7 mm, herbaceous, puberulent, base rounded, margin<br />
entire or sometimes remotely crenate, apex obtuse.<br />
Verticillasters 10–30-flowered, globose, 1–1.5 cm in diam.,<br />
few, widely spaced; floral leaves sessile, recurved, shorter<br />
than verticillasters. Pedicel 2–3 mm. Calyx tubular, 2-lipped,<br />
2.5–3 mm, hirtellous, glandular, throat bearded,<br />
± conspicuously 10-veined, tube ca. 1.5 mm; upper lip 3toothed,<br />
teeth lanceolate-triangular, ca. 1 mm; lower lip 2toothed,<br />
teeth subulate, ca. 1.5 mm. Corolla rose, purple or<br />
rarely white, ca. 4.5 mm, puberulent; tube ca. 3 mm, abruptly<br />
saccate at apex; lobes oblong, ca. 1.5 mm, entire, upper<br />
lanceolate. Ovary glabrous. Fl. Sep.<br />
Beijing Shi, Nanjing Shi, and other cities [Russia,<br />
Tajikistan, Turkmenistan; SW Asia, Europe].<br />
Cultivated in China. Source of oil of pennyroyal. Employed<br />
in the manufacture of soaps and synthetic menthol. Used<br />
medicinally for flatulent colic, stomach ailments, and as a<br />
diaphoretic.