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Ver. 19.2
April 17th, 2024

024.0000 Family Cyatheaceae Kaulf.; Wes. d. Farrnkr. 119 (1827)
4 genera, ca. 700 spp. Recent monographs tended to lump everything in Cyathea s. lat., but PPG-I separates again the 3 genera, and Dong et al. (2018) additionally reinstate Gymnosphaera, followed by Loiseau et al. 2019.
Species listed in database: 714; hybrids: 18
= Alsophilaceae C.Presl; Gefässbündel Farrn: 22 (footnote), 32 (1847)
= Alsophileae Engl.; Syllabus, ed. 2: 60 (1898)
= Alsophilinae Pfeiff.; Nomencl. Bot. 1(1): 123 (1872)
= Cyatheeae Gaudich.; Freycinet, Voy. Uranie, Bot.: 365 (1829)
= Cyatheinae Dumort.; Anal. Fam. Pl.: 67 (1829)
= Cyatheineae Bessey; Bot. High Schools: 376 (1880)
= Cyatheoideae Endl.; Gen. Pl. [Endlicher] 63 (1836)
= Hemiteliinae Pfeiff.; Nomencl. Bot. 1(2): 1606 (1874)
= Hymenophyllopsidaceae Pic.Serm.; Webbia 24: 712 (1970)
024.0002 Genus Gymnosphaera Blume; Enum. Pl. Javae 2: 242 (1828)
Distribution: Pantropics, mostly Old World (many Madagascar), only 2 neotropical. The genus has been demonstrated to be separate from Alsophila by Loiseau et al. 2019
Species listed in database: 46
species :
Gymnosphaera acrostichoides (Alderw.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 14 (2018)
Distribution: Moluccas (Seram); New Guinea (Irian Jaya, Papua New Guinea, Yapen Isl.)
42 MOL NWG
Synonyms: hidden
= Alsophila acrostichoides Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 2 (1918)
= Cyathea acrostichoides (Alderw.) Domin; Acta Bot. Bohem. 9: 88 (1930)
species :
Gymnosphaera alticola Tardieu; Naturaliste Malgache 3: 76, t. 2 (1951)
Distribution: Madagascar
29 MDG
Conservation Status: IUCN I
Synonyms: hidden
= Alsophila alticola (Tardieu) R.M.Tryon; Contr. Gray Herb. 200: 29 (1970)
= Cyathea alticola (Tardieu) Tindale; Contr. New South Wales Natl. Herb. 2: 331 (1956)
species :
Gymnosphaera andersonii (J.Scott ex Bedd.) Ching & S.K.Wu; Fl. Xizangica 1: 54 (1983)
Distribution: China (Yunnan); Tibet; India (Arunachal Pradesh, Assam State, Meghalaya, Nagaland, Sikkim, West Bengal); Myanmar (Kachin, etc.)
36 CHI TIB 40 IND 41 MYA
Synonyms: hidden
= Alsophila andersonii J.Scott ex Bedd.; Ferns Brit. India t. 310 (1869)
= Alsophila andersonii var. miaoensis Sarn.Singh & Panigrahi; Pl. Sci. Res. (Orissa) 17(1-2): 47 (1995), et Ferns Fern-Allies Arunachal Pradesh 1: 219 (2005)
= Alsophila suprasora Panigrahi & Sarn.Singh; Pl. Sci. Res. (India) 17(1-2): 47 (1995)
= Cyathea andersonii (J.Scott ex Bedd.) Copel.; Philipp. J. Sci., Bot. 4C: 56 (1909)
= Cyathea nayarii T.Bandyop., T.Sen & U.Sen; Pteridol. New Millennium 126 (2003) (fide Fraser-Jenkins 2008)
= Cyathea suprasora Panigrahi & Sarn.Singh; Pl. Sci. Res. (Orissa) 17(1-2): 47 (1995), et in Sarn. Singh & Panigrahi, Ferns Fern-Allies Arunachal Pradesh 1: 225 (2005)
= Gymnosphaera andersonii (J.Scott ex Bedd.) S.R.Ghosh; Pterid. Fl. E. India 198 (2004), isonym
species :
Gymnosphaera andohahelensis (Tardieu) Tardieu; Humbert, Fl. Madag. Fam. 4: 34, f. 4(6-7) (1951)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Alsophila andohahelensis Tardieu; Bull. Soc. Bot. France 88: 683 (1941)
= Cyathea andohahelensis (Tardieu) Tindale; Contr. New South Wales Natl. Herb. 2: 331 (1956)
species :
Gymnosphaera annae (Alderw.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 14 (2018)
Distribution: Sulawesi; Moluccas (Ambon)
42 MOL SUL
Conservation Status: IUCN V (two collections)
Synonyms: hidden
= Alsophila annae Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 23: 3 (1916)
= Cyathea annae (Alderw.) Domin; Acta Bot. Bohem. 9: 90 (1930)
species :
Gymnosphaera atropurpurea (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
Distribution: Philippines (Oriental Mindoro, Agusan del Norte, Davao)
42 PHI
Conservation Status: EN in the Philippines
Synonyms: hidden
= Alsophila atropurpurea (Copel.) C.Chr.; Index Filic., Suppl. (1906-1912): 4 (1913)
= Cyathea atropurpurea Copel.; Philipp. J. Sci., Bot. 3: 354 (1909)
species :
Gymnosphaera austroyunnanensis (S.G.Lu) S.G.Lu & Chun X.Li; Acta Bot. Yunnan. 27(1): 39 (2005)
Distribution: China (S-Yunnan)
36 CHI
Synonyms: hidden
= Alsophila austroyunnanensis S.G.Lu; Acta Bot. Yunnan 20(1): 45 (1998)
species :
Gymnosphaera bachmaensis S.Y.Dong; J. Syst. Evol. 60(2): 441 (2020)
Distribution: C-Vietnam
41 VIE
species :
Gymnosphaera baileyana (Domin) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 15 (2018)
Name: Wig tree fern
Distribution: Australia (NE-Queensland)
50 AUS
Conservation Status: IUCN R
Synonyms: hidden
= Alsophila baileyana Domin; Biblioth. Bot. 86: 29 (1913)
= Alsophila rebeccae var. commutata W.Bull; Queensl. Fl. Suppl. III: 91 (1890)
= Cyathea baileyana (Domin) Domin; Pteridophyta 262 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
species :
Gymnosphaera biformis (Rosenst.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 99 (1947)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Alsophila biformis Rosenst.; Repert. Spec. Nov. Regni Veg. 9: 423 (1911)
= Alsophila lilianiae R.M.Tryon; Contr. Gray Herb. 200: 34 (1970), nom. nov. for Cyathea arfakensis A. Gepp
= Cyathea arfakensis A.Gepp; Gibbs, Arfak: 69 (1917)
= Cyathea biformis (Rosenst.) Copel.; Philipp. J. Sci., Bot. 6: 364 (1911)
= Cyathea gibbsiae Copel.; Philipp. J. Sci. 38: 129 (1929)
= Hemitelia arfakensis (A.Gepp) Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 26 (1918)
= Polybotrya arfakensis A.Gepp; Gibbs, Dutch N. W. New Guinea 71 (1917)
= Stenochlaena dubia Alderw.; Bull. Dept. agric. Indes Néerl. 18: 26 (1908) (non Cyathea dubia (Bedd.) Domin (1929))
= Thysanobotrya arfakensis (A.Gepp) Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 66, t. 10 (1918)
= Thysanobotrya dubia (Alderw.) Posth.; Proc. Kon. Akad. Wet. Amsterdam 39: 825 (1936)
species :
Gymnosphaera boivinii (Mett. ex Ettingsh.) Tardieu; Humbert, Fl. Madag. Fam. 4: 41 (1951)
Distribution: Madagascar; Comores (Anjouan, Mayotte, Grande Comore)
29 COM MDG
Conservation Status: IUCN I
Synonyms: hidden
= Alsophila boivinii Mett. ex Ettingsh.; Farnkr. Jetztw. 220, t. 155, f. 2-3 (1864), non C. boivinii Mett. (1868)
= Alsophila boiviniiformis (Rakotondr. & Janssen) Christenh.; Global Fl. 1(4): 6 (2018), nom. illeg. superfl.
= Alsophila castanea Baker; J. Bot. 29: 3 (1891)
= Alsophila melanotricha Tardieu; Bull. Soc. Bot. France 88: 684 (1941)
= Cyathea badia Domin; Pteridophyta 262 (1929)
= Cyathea boiviniiformis Rakotondr. & Janssen; Adansonia 29(2): 198 (2007), nom. nov. for Alsophila boivinii Mett. ex Ettingsh.
= Cyathea insularis Domin; Pteridophyta 262 (1929), nom. nov. for Alsophila boivinii Mett. ex Ettingh., non Cyathea boivinii Mett.
= Cyathea melanotricha (Tardieu) Tindale; Contr. New South Wales Natl. Herb. 2: 331 (1956)
= Gymnosphaera melanotricha (Tardieu) Tardieu; Humbert, Fl. Madag. Fam. 4: 36, f. 5(2-3) (1951)
variant :
var. lobata (Rakotondr.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 15 (2018)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Alsophila bullata var. lobata (Rakotondr.) J.P.Roux; Strelitzia 23: 61 (2009)
= Cyathea boiviniiformis var. lobata (Rakotondr.) Rakotondr. & Janssen; Adansonia 29(2): 201 (2007)
= Cyathea bullata var. lobata Rakotondr.; Adansonia ser. 3, 21(1): 147 (1999)
variant :
var. madagascarica (Bonap.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 15 (2018)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Alsophila bullata var. madagascarica (Bonap.) J.P.Roux; Strelitzia 23: 61 (2009)
= Alsophila madagascarica Bonap.; Notes Ptéridol. 5: 53 (1917)
= Cyathea boiviniiformis var. madagascarica (Bonap.) Rakotondr. & Janssen; Adansonia 29(2): 202 (2007)
= Cyathea bullata var. madagascarica (Bonap.) Rakotondr.; Adansonia ser. 3, 21(1): 146 (1999)
= Cyathea malegassica Domin; Acta Bot. Bohem. 9: 133 (1930)
= Gymnosphaera madagascarica (Bonap.) Tardieu; Humbert, Fl. Madag. Fam. 4: 35, f. 6(2) (1951)
species :
Gymnosphaera bonii (Christ) S.Y.Dong; J. Syst. Evol. 60(2): 442 (2020)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hongkong, Yunnan); Vietnam
36 CHI 41 VIE
Synonyms: hidden
= Cyathea bonii Christ; J. Bot. (Morot) 28(4): 410 (1890)
species :
Gymnosphaera capensis (L.f.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 15 (2018)
subspecies :
subsp. capensis
Name: Forest Tree Fern
Distribution: Tanzania; Malawi; E-Zimbabwe; Mozambique; South Africa (Limpopo, Mpulamanga, KwaZulu-Natal, W-Cape Prov., E-Cape Prov.); Eswatini [Swaziland]
25 TAN 26 MLW MOZ ZIM 27 SAF SWZ
Synonyms: hidden
= Alsophila capensis (L.f.) J.Sm.; Lond. J. Bot. 1: 666 (1842)
= Amphicosmia capensis (L.f.) Klotzsch; Allg. Gartenzeit. 107 (1856)
= Amphicosmia riparia (Willd.) Gardner; Lond. J. Bot. 1: 441, t. 12 (1842)
= Aspidium capense (L.f.) Sw.; J. Bot. (Schrader) 1800(2): 42 (1801)
= Cormophyllum capense (L.f.) Newman; Phytol. 6: 238 (1854)
= Cyathea capensis (L.f.) Sm.; Mém. Acad. Turin 5: 417 (1793)
= Cyathea riparia Willd.; Sp. Pl. ed. 4. 5: 493 (1810)
= Hemitelia capensis (L.f.) Kaulf.; Enum. Filic. 253 (1824)
= Hemitelia riparia (Willd.) Desv.; Mém. Soc. Linn. Paris 6: 322 (1827)
= Polypodium capense L.f.; Suppl. 445 (1781)
= Polystichum capense (L.f.) J.Sm.; Bot. Mag. 72: Comp. 35 (1846)
= Trichomanes cormophyllum Kaulf.; Enum. Filic. 266 (1824)
= Trichomanes incisum Thunb.; Prodr. Fl. Cap. 173 (1800)
subspecies :
subsp. polypodioides (Sw.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 15 (2018)
Distribution: Brazil (Minas Gerais, Paraná, Rio de Janeiro, Rio Grande do Sul, Santa Catarina, São Paulo)
84 BRA
Synonyms: hidden
= Alsophila capensis subsp. polypodioides (Sw.) D.S.Conant; J. Arnold Arbor. 64(3): 369 (1983)
= Cyathea polypodioides Sw.; Kongl. Vetensk. Acad. Handl. 1817: 78 (1817)
= Hemitelia capensis f. nana Sehnem; Fl. Ilustr. Catarin. 26 (1978)
= Hemitelia gardneriana C.Presl; Abh. Königl. Böhm. Ges. Wiss., Math.-Naturw. Cl. V, 5: 320 (1848)
species :
Gymnosphaera commutata (Mett.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 15 (2018)
Distribution: peninsular Malaysia (Perak, Pahang, Selangor, Johor); Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak); Philippines (Mindanao)
42 BOR MLY PHI SUM
Synonyms: hidden
= Alsophila commutata Mett.; Ann. Mus. Bot. Lugduno-Batavi 1: 53 (1863)
= Alsophila heteromorpha Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 1 (1914)
= Alsophila heteromorpha var. decomposita Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 3: 129 (1920)
= Alsophila hewittii (Copel.) Alderw.; Malayan Ferns Fern Allies: 55 (1917)
= Alsophila squamulata (J.Sm.) Hook.; Sp. Fil. [W. J. Hooker] 1: 51 (1844) (p. p.)
= Alsophila subulata Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 1 (1918)
= Cyathea heteromorpha (Alderw.) Domin; Pteridophyta 262 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
= Cyathea hewittii Copel.; Philipp. J. Sci., Bot. 6: 134, t. 14 (1911)
= Cyathea recommutata Copel.; Philipp. J. Sci., Bot. 4: 36 (1909), nom. nov.
= Cyathea subulata (Alderw.) Domin; Acta Bot. Bohem. 9: 164 (1930)
= Cyathea toppingii Copel.; Philipp. J. Sci., Bot. 12: 51 (1917)
= Gymnosphaera hewittii (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
= Gymnosphaera recommutata (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
= Gymnosphaera squamulata J.Sm.; J. Bot. (Hooker) 3: 419 (1841) (non Blume (1828))
species :
Gymnosphaera denticulata (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
Distribution: China (Chongqing, Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Sichuan, Yunnan, Zhejiang); Japan; Ryukyu Isl.; Taiwan
36 CHI 38 JAP NNS TAI
Synonyms: hidden
= Alsophila acaulis Makino; Bot. Mag. (Tokyo) 28: 335 (1914)
= Alsophila denticulata Baker; J. Bot. 102 (1846), non Cyathea denticulata Goldm. 1843
= Cyathea acaulis (Makino) Domin; Pteridophyta 262 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
= Cyathea hancockii Copel.; Philipp. J. Sci., Bot. 4: 37 (1909), nom. nov.
= Cyathea polyodonta Domin; Pteridophyta 263 (1929)
= Dryopteris denticulata Hayata; Gen. Ind. Fl. Taiwan 101 (1917), non (Sw.) Kuntze
= Dryopteris hancockii (Copel.) Nakai; Bot. Mag. (Tokyo) 41: 74 (1927)
= Dryopteris taitunensis Koidz.; Bot. Mag. (Tokyo) 38: 197 (1924)
= Gymnosphaera hancockii (Copel.) Ching; (1978)
species :
Gymnosphaera dimorpha (Christ) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 16 (2018)
Distribution: C-Sulawesi; SE-Sulawesi
42 SUL
Conservation Status: IUCN I (2 collections)
Synonyms: hidden
= Alsophila dimorpha Christ; Ann. Jard. Bot. Buitenz. 19: 41 (1904)
= Cyathea dimorpha (Christ) Copel.; Philipp. J. Sci., Bot. 4: Bot. 34 (1909)
species :
Gymnosphaera gammiei (R.D.Dixit) S.Y.Dong; J. Syst. Evol. 60(2): 434 (2020)
Distribution: Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Meghalaya, Nagaland, Sikkim, West Bengal); Bangladesh
40 BAN BHU IND NEP
Remarks: Exclusion from Gymnosphaera henryi fide Dong, Haque & Rahman 2020
Synonyms: hidden
= Alsophila gammiei R.D.Dixit; Bull. Bot. Surv. India 34(1-4): 206 (1997) ("1992")
= Cyathea sharmae T.Bandyop., T.Sen & U.Sen; Pteridol. New Millennium 100 (2003), nom. nov. superfl.
species :
Gymnosphaera gigantea (Wall. ex Hook.) S.Y.Dong; Phytotaxa 400(2): 87 (2019)
Distribution: Nepal; ?Bhutan; India (Andhra Pradesh, Arunachal Pradesh, Assam State, Chhatishgarh, Jharkhand, Karnataka, Kerala, Madhya Pradesh, Maharashtra, Manipur, Meghalaya, Mizoram, Nagaland, Odisha, Sikkim, Tamil Nadu, Tripura, West Bengal); Sri Lanka; Bangladesh; Andaman Isl.; Myanmar (Kachin, etc.); Thailand (widespread); Laos; Cambodia; Vietnam; peninsular Malaysia (Langkawi Isl., Penang, Perak); C-Sumatra; W-Java
40 BAN IND NEP SRL 41 AND CBD LAO MYA THA VIE 42 JAW MLY SUM
Synonyms: hidden
= Alsophila balakrishnanii (R.D.Dixit & Tripathi) R.D.Dixit
= Alsophila gigantea Wall. ex Hook.; Sp. Fil. [W. J. Hooker] 1: 53 (1844)
= Alsophila helferiana C.Presl; Abh. Königl. Böhm. Ges. Wiss., Math.-Naturw. Cl. V, 5: 341 (1848)
= Alsophila polycampta Kunze; Bot. Zeitung (Berlin) 4: 475 (1846)
= Alsophila rheosora Christ; Bull. Herb. Boissier 6: 972 (1898)
= Alsophila umbrosa Wall. ex Ridl.; J. Mal. Br. R. As. Soc. 4: 6 (1926) (p. p.)
= Cyathea balakrishnanii R.D.Dixit & A.K.Tripathi; Bull. Bot. Surv. India 26(3-4): 170 (1984 publ. 1986) (fide Fraser-Jenkins 1997)
= Cyathea gigantea (Wall. ex Hook.) Holttum; Gard. Bull. Straits Settlem. 8: 318 (1935)
= Cyathea venulosa Wall.; Numer. List no. 180 (1829), nom. nud.
= Dichorexia gigantea (Wall. ex Hook.) C.Presl; Abh. Königl. Böhm. Ges. Wiss., Math.-Naturw. Cl. V, 5: 344 (1848)
= Gymnosphaera gigantea (Wall.) J.Sm.; Lond. J. Bot. 1: 667 (1842), nom. inval. (based on a nomen nud.)
= Polypodium giganteum Wall.; List 64 n. 321 (1828) (non Desv. (1827))
= Polypodium umbrosum Wall.; Numer. List no. 336 (1829), nomen
species :
Gymnosphaera glabra Blume; Enum. Pl. Javae 2: 242 (1828)
Distribution: Laos; peninsular Malaysia (Perlis, Kedah, Kelantan, Terengganu, Perak, Pahang, Selangor, Negeri Sembilan, Melaka, Johor); +Singapore; Sumatra; Borneo (Mt. Kinabalu, Sabah, Sarawak, Kalimantan); Sulawesi; W-Java; Lesser Sunda Isl. (Flores)
41 LAO 42 BOR JAW LSI MLY SNG SUL SUM
Synonyms: hidden
= Alsophila dubia Bedd.; J. Bot. 26: 1: t. 279 A (1888)
= Alsophila glabra (Blume) Hook.; Sp. Fil. 1: 51 (1844)
= Alsophila reducta Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 1 (1918)
= Alsophila vexans Ces.; Atti Reale Accad. Sci. Napoli 76: 4 (1876)
= Cyathea dubia (Bedd.) Copel.; Sarawak Mus. J. 2: 349 (1917)
= Cyathea glabra (Blume) Copel.; Philipp. J. Sci., Bot. 4: 35 (1909)
= Cyathea reducta Domin; Acta Bot. Bohem. 9: 88-174 (1930)
= Cyathea vexans (Ces.) C.Chr.; Gard. Bull. Straits Settlem. 7: 218 (1934)
= Gymnosphaera vexans (Ces.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
species :
Gymnosphaera henryi (Baker) S.R.Ghosh; Pterid. Fl. E. India 200 (2004)
Distribution: China (Guangdong, Guangxi, Hainan, Yunnan); Myanmar; Thailand
36 CHI 41 MYA THA
Remarks: bona sp. and not a synonym of C. gigantea fide Fraser-Jenkins et al. (2015b, 2017)
Synonyms: hidden
= Alsophila gigantea var. polynervata (R.H.Miao) Q.Xia; Acta Phytotax. Sin. 27: 11 (1989)
= Alsophila henryi Baker; Bull. Misc. Inform. Kew 1898: 229 (1898)
= Cyathea gigantea var. polynervata R.H.Miao; Act. Sci. Nat. Univ. 1: 100 (1980)
= Cyathea henryi (Baker) Copel.; Philipp. J. Sci., Bot. 4C: 38 (1909)
= Cyathea pectinata Ching & S.H.Wu; Ching & Chu H. Wang, Acta Phytotax. Sin. 9: 365 (1964)
= Cyathea petiolulata Ching & S.H.Wu; Ching & Chu H. Wang, Acta Phytotax. Sin. 9: 365 (1964) non H. Karst.
= Cyathea pseudogigantea Ching & S.H.Wu; Ching & Chu H. Wang, Acta Phytotax. Sin. 9: 365 (1964)
= Cyathea tinganensis Ching & S.H.Wu; Ching & Chu H. Wang, Acta Phytotax. Sin. 9: 366 (1964)
= Gymnosphaera gigantea var. polynervata (R.H.Miao) Y.K.Yang & J.K.Wu; Chin. Wild Pl. Resour. 21(6): 4 (2002)
species :
Gymnosphaera hornei (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 99 (1947)
Distribution: New Guinea (Louisiade Arch.: Misima Isl., Sudest Isl., Rossel Isl., Papua New Guinea, Irian Jaya); Solomon Isl. (Bougainville, Guadalcanal, Kolombangara, Makira, New Georgia, Nggatokae, Santa Ysabel, Vella Lavella); Fiji (Viti Levu, Vanua Levu, Ovalau, Taveuni)
42 NWG 43 SOL 60 FIJ
Synonyms: hidden
= Alsophila brunnea Brause; Bot. Jahrb. Syst. 56: 73 (1920)
= Alsophila dissitifolia Baker; J. Bot. 24: 182 (1886)
= Alsophila hornei Baker; J. Bot. 17: 293 (1879)
= Alsophila ledermannii Brause; Bot. Jahrb. Syst. 56: 76 (1920)
= Alsophila melanocaulos Alderw.; Nova Guinea 14: 1 (1924)
= Cyathea brunnea (Brause) Domin; Acta Bot. Bohem. 9: 101 (1930)
= Cyathea dimorphophylla Domin; Acta Bot. Bohem. 9: 111 (1930), nom. nov. for Alsophila ledermannii
= Cyathea dissitifolia (Baker) Domin; Pteridophyta 262 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
= Cyathea hornei (Baker) Copel.; Bull. Bernice P. Bishop Mus. 59: 38 (1929)
= Cyathea melanoclada Domin; Acta Bot. Bohem. 9: 174 (1930), nom. nov. for Alsophila melanocaulos
= Gymnosphaera melanocaulos (Alderw.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 99 (1947)
species :
Gymnosphaera impolita (Rakotondr. & Janssen) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 16 (2018)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Alsophila impolita (Rakotondr. & Janssen) Christenh.; Global Fl. 1(4): 6 (2018)
= Cyathea impolita Rakotondr. & Janssen; Adansonia 29(2): 203 (2007)
variant :
var. michelii (Rakotondr. & Janssen) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 16 (2018)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Cyathea impolita var. michelii Rakotondr. & Janssen; Adansonia 29(2): 204 (2007)
species :
Gymnosphaera khasyana (T.Moore ex Kuhn) Ching; Acta Phytotax. Sin. 22(3): 196 (1984)
Distribution: China (Yunnan); Tibet; Nepal; Bhutan; India (Arunachal Pradesh, Assam State, Manipur, Meghalaya, Mizoram, Nagaland, Sikkim, West Bengal); Bangladesh; Myanmar (Kachin, etc.)
36 CHI TIB 40 BAN BHU IND NEP 41 MYA
Synonyms: hidden
= Alsophila khasyana T.Moore ex Kuhn; Linnaea 36: 154 (1869)
= Alsophila latebrosa var. ornata (J.Scott ex Bedd.) Bedd.; Bull. Misc. Inform. Kew 1909: 423 (1909)
= Alsophila oldhamii Bedd.; Handb. Ferns Brit. India, t. 343 (1870)
= Alsophila ornata J.Scott ex Bedd.; Handb. Ferns Brit. India, t. 342 (1870)
= Alsophila ornata var. sikkimensis (C.B.Clarke & Baker) Bedd.; J. Bot. 31: 225 (1893)
= Alsophila pingbianica Y.K.Yang, Y.M.He & J.K.Wu; J. Wuhan Bot. Res. 17(3): 211 (1999)
= Alsophila scottiana Baker; Gard. Chron. 1872: 699 (1872)
= Alsophila sikkimensis C.B.Clarke & Baker; J. Linn. Soc., Bot. 24: 409 (1888) (non A. ornata var. sikkimensis Bedd., nor A. latebrosa var. ornata Bedd.)
= Cyathea khasyana (T.Moore ex Kuhn) Domin; Pteridophyta: 262 (1929)
= Cyathea oldhamii (Bedd.) Domin; Acta Bot. Bohem. 9: 88-174 (1930)
= Cyathea ornata (J.Scott ex Bedd.) Copel.; Philipp. J. Sci., Bot. 4: 59 (1909)
= Cyathea sikkimensis (C.B.Clarke & Baker) Cretz.; Repert. Spec. Nov. Regni Veg. 60: 316 (1941) (fide Fraser-Jenkins 1997)
= Gymnosphaera hongheensis Y.K.Yang, Z.Y.Yu & J.K.Wu
= Gymnosphaera pingbianica (Y.K.Yang, Y.M.He & J.K.Wu) Y.K.Yang & J.K.Wu
species :
Gymnosphaera lurida (Blume) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 17 (2018)
Distribution: peninsular Malaysia (Perak, Pahang); Sumatra; Borneo (Sabah); Sulawesi; Philippines (Mindoro); W-Java
42 BOR JAW MLY PHI SUL SUM
Synonyms: hidden
= Alsophila bakeri Zeiller; Bull. Soc. Bot. France 82: 72 (1885)
= Alsophila heterophylla (Domin) Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 2 (1914)
= Alsophila kingii C.B.Clarke ex Bedd.; Handb. Ferns Brit. India: 475 (1883)
= Alsophila lurida (Blume) Hook.; Sp. Fil. 1: 55 (1844)
= Alsophila melanorhachis Copel.; Philipp. J. Sci., C2: 146 (1907)
= Alsophila subdimorpha (Copel.) Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 16: 2 (1914)
= Chnoophora lurida Blume; Enum. Pl. Javae 2: 244 (1828)
= Cyathea heterophylla Domin; Pteridophyta 262 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
= Cyathea kingii (C.B.Clarke ex Bedd.) Copel.; Philipp. J. Sci., C 4: 56 (1909)
= Cyathea lurida (Blume) Copel.; Philipp. J. Sci., Bot. 4: 45 (1901)
= Cyathea melanorachis (Copel.) Copel.; Philipp. J. Sci., Bot. 4: 38 (1909)
= Cyathea subdimorpha Copel.; Philipp. J. Sci., Bot. 8: 140, t. 2 (1913)
= Gymnosphaera kingii (C.B.Clarke ex Bedd.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 99 (1947)
= Gymnosphaera melanorhachis (Copel.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
species :
Gymnosphaera metteniana (Hance) Tagawa; Acta Phytotax. Geobot. 14: 94 (1951)
Distribution: China (Chongqing, Fujian, Guangdong, Guizhou, Sichuan, Zhejiang); Japan; Ryukyu Isl.; Taiwan; Vietnam
36 CHI 38 JAP NNS TAI 41 VIE
Synonyms: hidden
= Alsophila formosana Baker; Ann. Bot. (Oxford) 5: 190 (1891)
= Alsophila lamprocaulis (Christ) Ching; Sinensia 2: 36 (1931)
= Alsophila metteniana Hance; J. Bot. 6: 175 (1868)
= Alsophila metteniana var. subglabra Ching & Q.Xia; Acta Phytotax. Sin. 27(1): 14 (1989)
= Aspidium lamprocaulon Christ; Bull. Acad. Int. Géogr. Bot., sér. 3, 16: 117 (1906)
= Cyathea formosana (Baker) Copel.; Philipp. J. Sci., Bot. 4: 36 (1909)
= Cyathea lamprocaulis (Christ) Ching; Bull. Fan Mem. Inst. Biol. Bot. 2, 1: 288 (1949)
= Cyathea metteniana (Hance) C.Chr. & Tardieu; Bull. Mus. Natl. Hist. Nat., sér. 2,6: 450 (1934)
= Dryopteris lamprocaulis (Christ) C.Chr.; Index Filic., Suppl. (1906-1912): 34 (1913)
= Gymnosphaera formosana (Baker) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
= Gymnosphaera lamprocaulis (Christ) Ching ex L.K.Ling
= Gymnosphaera metteniana var. subglabra (Ching & Q.Xia) Y.K.Yang & J.K.Wu
species :
Gymnosphaera mildbraedii (Brause) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 17 (2018)
Distribution: D.R.Congo [Zaire] (Ruwenzori); Uganda
23 ZAI 25 UGA
Synonyms: hidden
= Alsophila mildbraedii Brause; Deutsche Zentralafr. Exp. 2: 2, t. 1, f. A-C (1910)
= Cyathea mildbraedii (Brause) Domin; Pteridophyta 263 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
species :
Gymnosphaera nicklesii Tardieu & F.Ballard ex Tardieu; Notul. Syst. (Paris) 14: 329 (1953)
Distribution: Central African Republic
23 CAF
Synonyms: hidden
= Alsophila nicklesii (Tardieu & F.Ballard) R.M.Tryon; Contr. Gray Herb. 200: 31 (1970)
= Cyathea nicklesii (Tardieu & F.Ballard) Tindale; Contr. New South Wales Natl. Herb. 2: 331 (1956)
species :
Gymnosphaera ogurae (Hayata) Tagawa; Acta Phytotax. Geobot. 14: 94 (1951)
Distribution: Bonin Isl. (Chichijima)
38 OGA
Conservation Status: VU in Japan
Synonyms: hidden
= Alsophila ogurae Hayata; Bot. Mag. (Tokyo) 39: 149 (1925)
= Cyathea ogurae (Hayata) Domin; Acta Bot. Bohem. 9: 88-174 (1930)
species :
Gymnosphaera olivacea (Brause) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 17 (2018)
Distribution: New Guinea (Irian Jaya, Papua New Guinea)
42 NWG
Synonyms: hidden
= Alsophila olivacea Brause; Bot. Jahrb. Syst. 56: 74 (1920)
= Cyathea olivacea (Brause) Domin; Acta Bot. Bohem. 9: 143 (1930)
species :
Gymnosphaera phankelocii S.Y.Dong & C.W.Chen; Bot. J. Linn. Soc. 204(1): 68 (2023)
Distribution: Vietnam
41 VIE
species :
Gymnosphaera phlebodes (Lehnert & Coritico) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 17 (2018)
Distribution: New Guinea (Irian Jaya)
42 NWG
Synonyms: hidden
= Alsophila phlebodes Lehnert & Coritico; Syst. Bot. 38(4): 882 (2014)
species :
Gymnosphaera podophylla (Hook.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
Distribution: China (Fujian, Guangdong, Guangxi, Guizhou, Hainan, Yunnan); Japan; Ryukyu Isl.; Taiwan; Thailand (widespread); Laos; Cambodia; Vietnam
36 CHI 38 JAP NNS TAI 41 CBD LAO THA VIE
Synonyms: hidden
= Alsophila kohchangensis C.Chr.; Bot. Tidsskr. 32: 341 (1916)
= Alsophila podophylla Hook.; Hooker´s J. Bot. Kew Gard. Misc. 9: 334 (1857)
= Alsophila rheosora Baker; J. Bot. 28: 292 (1890)
= Cyathea kohchangensis Domin; Pteridophyta 262 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
= Cyathea podophylla (Hook.) Copel.; Philipp. J. Sci., Bot. 4: 33 (1909)
= Cyathea podophylla var. occulata C.Chr. & Tardieu; Bull. Mus. Hist. Nat. (Paris) 10: 449 (1934)
= Cyathea rheosora Copel.; Philipp. J. Sci., Bot. 4: 34 (1909)
= Dryopteris subconjuncta Christ; Notul. Syst. (Paris) 1: 185 c. fig. (1910)
species :
Gymnosphaera poolii (C.Chr.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 18 (2018)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Alsophila poolii C.Chr.; Index Filic. 46 (1905)
= Alsophila vestita (C.Chr.) Baker; J. Linn. Soc., Bot. 16: 412 (1876) (non C. Presl (1849))
= Cyathea poolii (C.Chr.) Domin; Pteridophyta 263 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
species :
Gymnosphaera ramispina (Hook.) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 98 (1947)
Distribution: Borneo (Mt. Kinabalu, Sabah, Sarawak, Kalimantan)
42 BOR
Synonyms: hidden
= Alsophila amaiambitensis Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 1 (1918)
= Alsophila burbidgei Christ; Ann. Jard. Bot. Buitenz. 20: 138 (1906) (p. p., non Baker)
= Alsophila hallieri Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 2 (1918) (non Rosenst.)
= Alsophila kenepaiana Alderw.; Bull. Jard. Bot. Buitenzorg, ser. 3, 2: 129 (1920)
= Alsophila ramispina Hook.; Hook., Syn. Fil. 42 (1866)
= Cyathea amaiambitensis (Alderw.) Domin; Acta Bot. Bohem. 9: 90 (1930)
= Cyathea kenepaiana Domin; Acta Bot. Bohem. 9: 127 (1930)
= Cyathea ramispina (Hook.) Copel.; Philipp. J. Sci., Bot. 4: 36 (1909)
species :
Gymnosphaera ramispinoides (M.Kato) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 18 (2018)
Distribution: Moluccas (Seram)
42 MOL
Synonyms: hidden
= Alsophila ramispinoides (M.Kato) Lehnert; Syst. Bot. 38(4): 883 (2014), et 40(2): 386 (2015)
= Cyathea ramispinoides M.Kato; J. Fac. Sci. Univ. Tokyo, Sect. 3, Bot. 14(5): 378 (1990)
species :
Gymnosphaera rebeccae (F.Muell.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 18 (2018)
Distribution: Lesser Sunda Isl. (Flores); Australia (Queensland)
42 LSI 50 AUS
Synonyms: hidden
= Alsophila rebeccae F.Muell.; Fragm. 5: 33 (1865), 53, 36: 117 (1896)
= Alsophila rebeccae var. lobulata Domin; Biblioth. Bot. 20: 29 (1914)
= Cyathea rebeccae (F.Muell.) Domin; Pteridophyta: 263 (1929), et Acta Bot. Bohem. 9: 88-174 (1929 publ. 1930)
species :
Gymnosphaera rouhaniana (Rakotondr. & Janssen) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 18 (2018)
Distribution: Madagascar
29 MDG
Synonyms: hidden
= Alsophila rouhaniana (Rakotondr. & Janssen) Christenh.; Global Fl. 1(4): 6 (2018)
= Cyathea rouhaniana Rakotondr. & Janssen; Adansonia 29(2): 207 (2007)
species :
Gymnosphaera rubella (Holttum) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 18 (2018)
Distribution: New Guinea (Papua New Guinea, d´Entrecasteaux Arch.)
42 NWG
Synonyms: hidden
= Alsophila rubella (Holttum) R.M.Tryon; Contr. Gray Herb. 200: 36 (1970)
= Cyathea rubella Holttum; Kew Bull. 16: 61 (1962)
species :
Gymnosphaera salletii (Tardieu & C.Chr. ex C.Chr. & Tardieu) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 18 (2018)
Distribution: Vietnam
41 VIE
Synonyms: hidden
= Alsophila salletii (Tardieu & C.Chr.) R.M.Tryon; Contr. Gray Herb. 200: 32 (1970)
= Cyathea salletii Tardieu & C.Chr. ex C.Chr. & Tardieu; Bull. Mus. Natl. Hist. Nat., sér. 2,6: 450 (1934)
species :
Gymnosphaera salvinii (Hook.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 18 (2018)
Distribution: Mexico (Chiapas, Oaxaca, Veracruz); Guatemala; Belize; El Salvador; Honduras; Nicaragua; Costa Rica; Panama; Peru; SE-Brazil (Minas Gerais)
79 MEX 80 BLZ COS ELS GUA HON NIC PAN 83 PER 84 BRA
Remarks: a true Gymnosphaera based on molecular results
Synonyms: hidden
= Alsophila munchii Christ; Bull. Herb. Boissier, sér. 2, 5: 734 (1905)
= Alsophila salvinii Hook.; Hook., Syn. Fil. 36 (1866)
= Cyathea salvinii (Hook.) Domin; Pteridophyta 263 (1929), et Acta Bot. Bohem. 9: 88-174 (1929 publ. 1930)
species :
Gymnosphaera saxicola S.Y.Dong & Z.Y.Zuo; J. Syst. Evol. 60(2): 441 (2020)
Distribution: China (SW-Yunnan)
36 CHI
species :
Gymnosphaera scandens (Brause) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 18 (2018)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: IUCN I (one collection)
Synonyms: hidden
= Alsophila scandens Brause; Bot. Jahrb. Syst. 56: 77 (1920)
= Cyathea scandens (Brause) Domin; Acta Bot. Bohem. 9: 156 (1930)
= Thysanobotrya scandens (Brause) Posth.; Proc. Kon. Akad. Wet. Amsterdam 39: 826 (1936)
species :
Gymnosphaera schlechteri (Brause) Copel.; Ann. Cryptog. Phytopathol. 5 [Gen. Fil.] 99 (1947)
Distribution: New Guinea (Papua New Guinea)
42 NWG
Conservation Status: IUCN R (one collection)
Synonyms: hidden
= Alsophila schlechteri Brause; Bot. Jahrb. Syst. 49: 16, f. 1 D (1912)
= Cyathea schlechteri (Brause) Domin; Pteridophyta 263 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
species :
Gymnosphaera schliebenii (Reimers) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 19 (2018)
Distribution: Tanzania (Uluguru Mts., Ukaguru Mts.)
25 TAN
Synonyms: hidden
= Alsophila schliebenii Reimers; Notizbl. bot. Gart. Berlin-Dahlem 11: 918 (1933)
= Cyathea fadenii Holttum; Kew Bull. 36(3): 470 (1981), nom. nov.
species :
Gymnosphaera subdubia (Alderw.) S.Y.Dong; Ann. Missouri Bot. Gard. 103(1): 19 (2018)
Distribution: C-Sumatra; W-Java
42 JAW SUM
Synonyms: hidden
= Alsophila persquamulata Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 28: 1 (1918)
= Alsophila subdubia Alderw.; Bull. Jard. Bot. Buitenzorg, sér. 2, 20: 3 (1915)
= Cyathea persquamulata (Alderw.) Domin; Acta Bot. Bohem. 9: 88-174 (1930)
= Cyathea subdubia (Alderw.) Domin; Pteridophyta 263 (1929), et Acta Bot. Bohem. 9: 88-174 (1930)
species :
Gymnosphaera vietnamensis S.Y.Dong & C.W.Chen; Bot. J. Linn. Soc. 204(1): 71 (2023)
Distribution: Vietnam
41 VIE
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A complete, synonymic checklist of the Higher Plants of the World


Version 19.2 (April 17th, 2024)

Covering 354,696 accepted species,
8,644 hybrids, 47,758 accepted infraspecific taxa and 996,379 synonyms,
in total 1,407,477* names

[*about 25,000 dubious, "unplaced" names and about 4,000 invalid names (Art. 24.3) like "typicus" are deliberately omitted]


With complete, detailed distribution data (countries and regions) and literature references

Arranged in linear phylogenetic sequence reflecting true relationships

Fully searchable and continously updated

With 38,290 pictures of 15,532 species

Assembled through 40 years independently from other databases

In cooperation with Catalogue of Life