Bryophytes - Gambar

http://bryophytes.plant.siu.edu/class.
html
A 1994 diagram indicating our understanding of cladistic relationships of the major lineages of green algae and land plants.
(Modified from Mishler, B. D.et al. (1994) Phylogenetic relationships of the "green algae: and "bryophytes."
Ann. Missouri Bot. Gard. 81: 451-483.)
Abbreviations as follows:
Chloro. = Chlorophytes
Pleur. = Pleurophytes;
Ulvo. = Ulvophytes
Charop. = Charophytes
Micromonad.= Micromonadophytes;
Liv. = Liverworts;
Horn. = Hornworts;
Moss = Mosses;
Trach. = Tracheophytes.
Below, a 2006 diagram indicting our understanding of the relationship of the major lineages of green algae and land plants
based upon phylogenomic evidence.

Liverwort Classification at the Rank of Genus and Above
This classification is taken from: Crandall-Stotler, B., R. E. Stotler & D. G.Long. [2008]2009. Morphology
and classification of the Marchantiophyta, pp. 1-54. In B. Goffinet & A. J. Shaw (eds.) Bryophyte Biology,
2nd edition. Cambridge University Press, Cambridge. Modifications have been made based upon the more
recent publication: Crandall-Stotler, B., R. E. Stotler & D. G. Long. 2009. Phylogeny and Classification of
the Marchantiophyta, Edinburgh Journal of Botany 66: 155-198.
PHYLUM MARCHANTIOPHYTA Stotler & Crand.-Stotl., in A. J. Shaw & B. Goffinet, Bryoph. Biol.: 63 (2000).
CLASS: HAPLOMITRIOPSIDA Stotler & Crand.-Stotl., Bryologist 80: 425 (1977). [including
Treubiopsida Stech, J.-P. Frahm, Hilger & W. Frey, Nova Hedwigia 71: 207 (2000)].
SUBCLASS TREUBIIDAE Stotler & Crand.-Stotl., Taxon 57: 290 (2008).
ORDER: TREUBIALES Schljakov, Bot. Zhurn. (Moscow & Leningrad) 57: 499 (1972).
Treubiaceae Verd., Man. Bryol. 427 (1932). Apotreubia S. Hatt. & Mizut., Treubia K. I. Goebel
SUBCLASS HAPLOMITRIIDAE Stotler & Crand.-Stotl., Taxon 57: 290 (2008).
ORDER: Calobryales Hamlin, Rec. Domin. Mus. 7: 315 (1972).

Haplomitriaceae Dedecek, Arch. Naturwiss. Landesdurchf. Böhmen 5(4): 71 (1884). Haplomitrium Nees
nom. cons.
CLASS: MARCHANTIOPSIDA Cronquist, Takht. & W. Zimm., Taxon 15: 132-133 (1966).
("Marchantiatae").
SUBCLASS: BLASIIDAE He-Nygrén, Juslén, Ahonen, Glenny & Piippo, Cladistics 22: 27 (2006).
ORDER: BLASIALES Stotler & Crand.-Stotl. in A. J. Shaw & B. Goffinet, Bryoph. Biol.: 63 (2000).
Blasiaceae H. Klinggr., Höh. Crypt. Preuss.: 14 (1858). Blasia L., Cavicularia Steph.
SUBCLASS: MARCHANTIIDAE Engl. [Unterklasse "Marchantiales"] in: A. Engler & K. Prantl, Nat.

Pflanzenfam. I (3): 1 (1893).
ORDER: SPHAEROCARPALES Cavers, New Phytol. 9: 81 (1910).
Sphaerocarpaceae Heeg, Verh. K. K. Zool.-Bot. Ges. Wien 41: 573

(1891). Sphaerocarpos Boehm., Geothallus Campb.
Riellaceae Engl., Syllabus, Grosse Ausgabe: 45 (1892). Riella Mont.
ORDER: NEOHODGSONIALES D. G. Long, Edinburgh J. Bot. 63: 258 (2006).
Neohodgsoniaceae D. G. Long, Edinburgh J. Bot. 63: 258 (2006). Neohodgsonia Perss.
ORDER: LUNULARIALES D. G. Long, Edinburgh J. Bot. 63: 259 (2006).
Lunulariaceae H. Klinggr., Höh. Crypt. Preuss.: 9 (1858). Lunularia Adans.

ORDER: MARCHANTIALES Limpr. in Cohn, Krypt.-Fl. Schlesien 1: 239, 336 (1877).
Marchantiaceae Lindl., Nat. Syst. Bot. (ed. 2): 412 (1836). Bucegia Radian, Marchantia L., Preissia Corda
Aytoniaceae Cavers, New Phytol. 10: 42 (1911). Asterella P. Beauv., Cryptomitrium Austin ex

Underw., Mannia Opiz nom. cons., Plagiochasma Lehm. & Lindenb. nom. cons., Reboulia Raddi nom. cons.
Cleveaceae Cavers, New Phytol. 10: 42 (1911). Athalamia Falconer, Sauteria Nees, Peltolepis Lindb.
Monosoleniaceae Inoue, Bull. Natl. Sci. Mus. Tokyo 9: 117 (1966). Monosolenium Griff.
Conocephalaceae Müll. Frib. ex Grolle, J. Bryol. 7: 207 (1972). Conocephalum Hill nom. cons.
Cyathodiaceae Stotler & Crand.-Stotl. in A. J. Shaw & B. Goffinet, Bryoph. Biol.: 63

(2000). Cyathodium Kunze
Exormothecaceae Müll. Frib. ex Grolle, J. Bryol. 7: 208

(1972). Aitchisoniella Kashyap, Exormotheca Mitt., Stephensoniella Kashyap
Corsiniaceae Engl., Syllabus, Grosse Ausgabe: 44 (1892). Corsinia Raddi, Cronisia Berk.
Monocarpaceae D. J. Carr ex Schelpe, J. S. African Bot. 35: 110 (1969). Monocarpus D. J. Carr
Oxymitraceae Müll. Frib. ex Grolle, J. Bryol. 7: 215 (1972). Oxymitra Bisch. ex Lindenb.
Ricciaceae Rchb., Bot. Damen: 255 (1828). Riccia L., Ricciocarpos Corda
Wiesnerellaceae Inoue, Ill. Jap. Hepat. 2: 192 (1976). Wiesnerella Schiffn.
Targioniaceae Dumort., Anal. Fam. Pl.: 68, 70 (1829). Targionia L.

Monocleaceae A. B. Frank in Leunis, Syn. Pflanzenk. ed. 2: 1556 (1877). Monoclea Hook.
Dumortieraceae D. G. Long, Edinburgh J. Bot. 63: 260 (2006). Dumortiera Nees
CLASS: JUNGERMANNIOPSIDA Stotler & Crand.-Stotl., Bryologist 80: 425 (1977).
SUBCLASS: PELLIIDAE He-Nygrén, Juslén, Ahonen, Glenny & Piippo, Cladistics 22: 27 (2006).
ORDER: PELLIALES He-Nygrén, Juslén, Ahonen, Glenny & Piippo, Cladistics 22: 27 (2006).
Pelliaceae H. Klinggr, Höh. Crypt. Preuss.: 13 (1858). Noteroclada Taylor ex Hook. & Wilson, Pellia Raddi

nom. cons.
ORDER: FOSSOMBRONIALES Schljakov, Bot. Zhurn. (Moscow & Leningrad) 57: 500 (1972).
SUBORDER: Calyculariineae He-Nygrén, Juslén, Ahonen, Glenny & Piippo, Cladistics 22: 27 (2006).
Calyculariaceae He-Nygrén, Juslén, Ahonen, Glenny & Piippo, Cladistics 22: 27 (2006). Calycularia Mitt.
SUBORDER: Makinoiineae He-Nygrén, Juslén, Ahonen, Glenny & Piippo, Cladistics 22: 27 (2006).
Makinoaceae Nakai, Chosakuronbun Mokuroku [Ord. Fam. Trib. Nov.]: 201 (1943). Makinoa Miyake
SUBORDER: Fossombroniineae R.M.Schust. ex Stotler & Crand.-Stotl. in A. J. Shaw & B. Goffinet, Bryoph.

Biol.: 63 (2000).
Petalophyllaceae Stotler & Crand.-Stotl., Novon 12: 335 (2002). Petalophyllum Nees & Gottsche ex

Lehm., Sewardiella Kashyap
Allisoniaceae Schljakov, Pecenocnye Mchi, Morfol. Filog. Klassif. [Liverwort morphology, phylogeny,
classification]: 119 (1975). Allisonia Herzog
Fossombroniaceae Hazsl. nom. cons., Magyar Bir, Moh.-Fl.: 20, 36 (1885). Fossombronia Raddi

[including Austrofossombronia R.M.Schust.]
ORDER: PALLAVICINIALES W. Frey & M. Stech, Nova Hedwigia 81: 64 (2005).
SUBORDER: Phyllothalliineae R. M. Schust., Trans. Brit. Bryol. Soc. 5: 283 (1967).
Phyllothalliaceae E. A. Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2: 247 (1964). Phyllothallia E. A.
Hodgs.
SUBORDER: Pallaviciniineae R. M. Schust., Phytologia 56: 65 (1984).
Sandeothallaceae R. M. Schust., New Man. Bryol.: 951 (1984). Sandeothallus R. M. Schust.
Moerckiaceae Stotler & Crand.-Stotl., Nova Hedwigia Beih. 131: 54 (2007). Hattorianthus R. M. Schust. &
Inoue, Moerckia Gottsche
Hymenophytaceae R. M. Schust., J. Hattori Bot. Lab. 26: 296 (1963). Hymenophyton Dumort.
Pallaviciniaceae Mig., Krypt.-Fl. Deutschl., Moose: 423

(1904). Greeneothallus Hässel, Jensenia Lindb., Pallavicinia Gray nom.
cons., Podomitrium Mitt., Seppeltia Grolle, Symphyogyna Nees &
Mont., Symphyogynopsis Grolle, Xenothallus R. M. Schust.
SUBCLASS METZGERIIDAE Barthol.-Began, Phytologia 69: 465. (1990[1991]).
ORDER: PLEUROZIALES Schljakov, Bot. Zhurn. (Moscow & Leningrad) 57: 505 (1972).
Pleuroziaceae Müll. Frib., Lebermoose 1: 404 (1909). Pleurozia Dumort. [including Eopleurozia R. M.
Schust.]
ORDER: METZGERIALES Chalaud, Ann. Bryol. 3: 41 (1930).
Metzgeriaceae H. Klinggr., Höh. Crypt. Preuss.: 10 (1858). Metzgeria Raddi

[including Apometzgeria Kuwah. and Austrometzgeria Kuwah.], Steereella Kuwah., Vandiemenia Hewson
Aneuraceae H. Klinggr., Höh. Crypt. Preuss.: 11 (1858). Aneura Dumort.

[including Cryptothallus Malmb.], Riccardia Gray nom. cons., Lobatiriccardia (Mizut. & S. Hatt.)
Furuki, Verdoornia R. M. Schust.
Mizutaniaceae Furuki & Z. Iwats., J. Hattori Bot. Lab. 67: 291 (1989). Mizutania Furuki & Z. Iwats.
SUBCLASS JUNGERMANNIIDAE Engl. [Unterklasse "Jungermanniales"] in A. Engler & K. Prantl, Nat.

Pflanzenfam. I (3): 1 (1893).
ORDER: PORELLALES Schljakov, Bot. Zhurn. (Moscow & Leningrad) 57: 505 (1972).
SUBORDER: Porellineae R. M. Schust., J. Hattori Bot. Lab. 26: 229 (1963).
Porellaceae Cavers nom. cons., New Phytol. 9: 292 (1910). Ascidiota C. Massal., Porella L.

[including Macvicaria W. E. Nicholson]
Goebeliellaceae Verd., Man. Bryol.: 425 (1932). Goebeliella Steph.
Lepidolaenaceae Nakai, Chosakuronbun Mokuroku [Ord. Fam. Trib. Nov.]: 200

(1943). Gackstroemia Trevis., Lepidogyna R. M. Schust., Lepidolaena Dumort., Jubulopsis R. M. Schust.
SUBORDER: Radulineae R. M. Schust., J. Hattori Bot. Lab. 26: 229 (1963).

Radulaceae Müll. Frib., Lebermoose 1: 404 (1909). Radula Dumort. nom. cons.
SUBORDER: Jubulineae Müll. Frib., Lebermoose 1: 403 (1909).
Frullaniaceae Lorch in G. Lindau, Krypt.-Fl. Anf. 6: 174 (1914). Frullania Raddi [including Amphijubula R. M.

Schust., Neohattoria Kamim., Schusterella S. Hatt., Sharp & Mizut., and Steerea S. Hatt. & Kamim.]
Jubulaceae H. Klinggr., Höh. Crypt. Preuss.: 40 (1858). Jubula Dumort. nom. cons., Nipponolejeunea S.

Hatt.
Lejeuneaceae Cavers, New Phytol. 9: 291 (1910). Acanthocoleus R. M. Schust., Acantholejeunea (R. M.

Schust.) R. M. Schust., Acrolejeunea (Spruce) Schiffn. nom. cons., Anoplolejeunea (Spruce)
Schiffn., Aphanotropis Herzog, Archilejeunea (Spruce) Schiffn., Aureolejeunea R. M.
Schust., Austrolejeunea (R. M. Schust.) R. M. Schust.
[including Nephelolejeunea Grolle], Blepharolejeunea S. W. Arnell, Brachiolejeunea (Spruce)
Schiffn., Bromeliophila R. M. Schust., Bryopteris (Nees) Lindenb., Calatholejeunea K. I.
Goebel, Caudalejeunea (Steph.) Schiffn., Cephalantholejeunea (R. M. Schust. & Kachroo) R. M.
Schust., Cephalolejeunea Mizut., Ceratolejeunea (Spruce) J. B. Jack & Steph., Cheilolejeunea (Spruce)
Schiffn. [including Cyrtolejeunea A. Evans], Chondriolejeunea (Benedix) Kis &
Pócs, Cladolejeunea Zwick., Cololejeunea (Spruce) Schiffn. [including Aphanolejeunea A.
Evansand Metzgeriopsis K. I. Goebel], Colura (Dumort.) Dumort., Cyclolejeunea A. Evans, Cystolejeunea A.
Evans, Dactylophorella R. M. Schust., Dendrolejeunea (Spruce) Lacout., Dicranolejeunea (Spruce)
Schiffn., Diplasiolejeunea (Spruce) Schiffn., Drepanolejeunea (Spruce) Schiffn. [including Capillolejeunea S.
W. Arnell and Rhaphidolejeunea Herzog], Echinolejeunea R. M. Schust., Evansiolejeunea Vanden
Berghen, Frullanoides Raddi, Fulfordianthus Gradst., Haplolejeunea Grolle, Harpalejeunea (Spruce)
Schiffn., Hattoriolejeunea Mizut., Kymatolejeunea Grolle, Leiolejeunea A. Evans, Lejeunea Lib. nom. cons.
[including Amblyolejeunea Ast, Amphilejeunea R. M. Schust., Crossotolejeunea (Spruce)
Schiffn., Cryptogynolejeunea R. M. Schust., Dactylolejeunea R. M. Schust., Dicladolejeunea R. M.
Schust., Echinocolea R. M. Schust., Macrolejeunea (Spruce) Schiffn., Neopotamolejeunea E. Reiner
and Taxilejeunea (Spruce) Schiffn. nom. cons.], Lepidolejeunea R. M. Schust., Leptolejeunea (Spruce)
Schiffn., Leucolejeunea A. Evans, Lindigianthus Kruijt & Gradst., Lopholejeunea (Spruce) Schiffn. nom.
cons., Luteolejeunea Piippo, Macrocolura R. M. Schust., Marchesinia Gray nom.
cons., Mastigolejeunea (Spruce)
Schiffn., Metalejeunea Grolle, Microlejeunea Steph., Myriocolea Spruce, Myriocoleopsis Schiffn., Neurolejeu
nea (Spruce) Schiffn., Odontolejeunea (Spruce) Schiffn., Omphalanthus Lindenb. &
Nees, Oryzolejeunea (R. M. Schust.) R. M. Schust., Otolejeunea Grolle &
Tixier, Phaeolejeunea Mizut., Physantholejeunea R. M. Schust., Pictolejeunea Grolle, Pluvianthus R. M.
Schust. & Schäf.-Verw., Prionolejeunea (Spruce) Schiffn., Ptychanthus Nees, Pycnolejeunea (Spruce)
Schiffn., Rectolejeunea A.
Evans, Schiffneriolejeunea Verd., Schusterolejeunea Grolle, Siphonolejeunea Herzog, Sphaerolejeunea Her
zog, Spruceanthus Verd., Stenolejeunea R. M. Schust., Stictolejeunea (Spruce)
Schiffn., Symbiezidium Trevis., Thysananthus Lindenb., Trachylejeunea (Spruce) Schiffn. nom. cons.
[including Potamolejeunea (Spruce) Lacout.], Trocholejeunea Schiffn., Tuyamaella S. Hatt., Tuzibeanthus S.
Hatt., Verdoornianthus Gradst., Vitalianthus R. M. Schust. & Giancotti, Xylolejeunea X-L. He & Grolle
ORDER PTILIDIALES Schljakov, Bot. Zhurn. (Moscow & Leningrad) 57: 501 (1972).
Ptilidiaceae H. Klinggr., Höh. Crypt. Preuss.: 37 (1858). Ptilidium Nees
Neotrichocoleaceae Inoue, Ill. Jap. Hepat. 1: 176 (1974). Neotrichocolea S. Hatt., Trichocoleopsis S.

Okamura
Herzogianthaceae Stotler & Crand.-Stotl. in B. Goffinet & A. J. Shaw, Bryoph. Biol. ed. 2: 54

(2009[2008]). Herzogianthus R. M. Schust.
ORDER JUNGERMANNIALES H. Klinggr., Höh. Crypt. Preuss.: 16 (1858).
SUBORDER: Perssoniellineae R. M. Schust., J. Hattori Bot. Lab. 26: 229-230 (1963).
Perssoniellaceae R. M. Schust. ex Grolle, J. Bryol. 7: 216 (1972). Perssoniella Herzog

Schistochilaceae H. Buch, Commentat. Biol. 3(1): 9 (1928). Gottschea Nees ex Mont.
[including Paraschistochila R. M. Schust.], Pachyschistochila R. M. Schust. & J. J. Engel, Pleurocladopsis R.
M. Schust., Schistochila Dumort.
SUBORDER: Lophocoleineae Schljakov, Bot. Zhurn. (Moscow & Leningrad) 57: 504 (1972). (syn.:
Geocalycineae R. M. Schust., J. Hattori Bot. Lab. 36: 397 (1972 [1973]).
Pseudolepicoleaceae Fulford & J. Taylor, Nova Hedwigia 1: 411 (1960). Archeophylla R. M.

Schust., Blepharostoma (Dumort.) Dumort., Chaetocolea Spruce, Herzogiaria Fulford ex
Hässel, Isophyllaria E. A. Hodgs. & Allison, Pseudolepicolea Fulford & J. Taylor [including Archeochaete R.
M. Schust. and Lophochaete R. M. Schust.], Temnoma Mitt.
Trichocoleaceae Nakai, Chosakuronbun Mokuroku [Ord. Fam. Trib. Nov.]: 201 (1943). Castanoclobus J. J.

Engel & Glenny, Eotrichocolea R. M. Schust., Leiomitra Lindb., Trichocolea Dumort. nom. cons.
Grolleaceae Solari ex R. M.S chust., Phytologia 56: 66 (1984). Grollea R. M. Schust.
Mastigophoraceae R. M. Schust., J. Hattori Bot. Lab. 36: 345. (1972 [1973]). Dendromastigophora R. M.

Schust., Mastigophora Nees nom. cons.
Herbertaceae Müll. Frib. ex Fulford & Hatcher, Bryologist 61: 284 (1958). Herbertus Gray, Olgantha R. M.

Schust., Triandrophyllum Fulford & Hatcher
Vetaformataceae Fulford & J. Taylor, Mem. New York Bot. Gard. 11: 27 (1963). Vetaforma Fulford & J.
Taylor
Lepicoleaceae R. M. Schust., Nova Hedwigia 5: 27 (1963). Lepicolea Dumort.
Phycolepidoziaceae R. M. Schust., Bull. Torrey Bot. Club 93: 442 (1967). Phycolepidozia R. M. Schust.
Lepidoziaceae Limpr. in Cohn, Krypt.-Fl. Schlesien 1: 310 (1877). Acromastigum A. Evans, Amazoopsis J.
J. Engel & G. L. S. Merr., Arachniopsis Spruce, Bazzania Gray nom. cons., Chloranthelia R. M.
Schust., Dendrobazzania R. M. Schust. & W. B. Schofield, Drucella E. A. Hodgs., Hyalolepidozia S. W.
Arnell ex Grolle, Hygrolembidium R. M. Schust., Isolembidium R. M. Schust., Kurzia G.
Martens, Lembidium Mitt. nom. cons., Lepidozia (Dumort.) Dumort. nom. cons., Monodactylopsis (R. M.
Schust.) R. M. Schust., Mastigopelma Mitt., Megalembidium R. M. Schust., Micropterygium Lindenb., Nees
& Gottsche, Mytilopsis Spruce, Neogrollea E. A. Hodgs., Odontoseries Fulford, Paracromastigum Fulford &
J. Taylor, Protocephalozia (Spruce) K. I. Goebel, Pseudocephalozia R. M.
Schust., Psiloclada Mitt., Pteropsiella Spruce, Sprucella Steph., Telaranea Spruce ex
Schiffn., Zoopsidella R. M. Schust., Zoopsis Hook. f. ex Gottsche, Lindenb. & Nees
Lophocoleaceae Vanden Berghen in Robyns, Fl. Gén. Belgique, Bryoph. 1: 208

(1956). Amphilophocolea R. M.
Schust., Chiloscyphus Corda, Clasmatocolea Spruce, Conoscyphus Mitt., Cyanolophocolea R. M.
Schust., Evansianthus R. M. Schust. & J. J. Engel
[including Austrolembidium Hässel], Hepatostolonophora J. J. Engel & R. M.
Schust., Heteroscyphus Schiffn. nom. cons. [including Tetracymbaliella Grolle], Lamellocolea J. J
.Engel, Leptophyllopsis R. M. Schust., Leptoscyphopsis R. M.
Schust., Leptoscyphus Mitt., Lophocolea (Dumort.) Dumort. [including Campanocolea R. M.
Schust.], Pachyglossa Herzog & Grolle [including Invisocaulis R. M. Schust. nom.
inval.], Perdusenia Hässel, Physotheca J. J. Engel & Gradst., Pigafettoa C. Massal., Platycaulis R. M.
Schust., Pseudolophocolea R. M. Schust. & J. J. Engel, Stolonivector J. J. Engel, Xenocephalozia R. M.
Schust.
Brevianthaceae J. J. Engel & R. M. Schust., Phytologia 47: 317 (1981). Brevianthus J. J. Engel & R. M.
Schust.
Chonecoleaceae R. M. Schust. ex Grolle, J. Bryol. 7: 206 (1972). Chonecolea Grolle

Plagiochilaceae Müll. Frib. & Herzog in Müller, Leberm. Eur.: 877 (1956). Acrochila R. M.
Schust., Chiastocaulon Carl, Dinckleria Trevis. [? Proskauera Heinrichs & J. J. Engel], Pedinophyllopsis R.
M. Schust. & Inoue, Pedinophyllum (Lindb.) Lindb., Plagiochila (Dumort.) Dumort. nom. cons.
[including Rhodoplagiochila R. M. Schust., Steereochila Inoue, and Szweykowskia Gradst. & E.
Reiner], Plagiochilidium Herzog, Plagiochilion S. Hatt., Xenochila R.M.Schust.
SUBORDER: Cephaloziineae Schljakov, Bot. Zhurn. (Moscow & Leningrad) 57: 503 (1972). [syn.:
Cephaloziineae R. M. Schust., J. Hattori Bot. Lab. 36: 391. (1972 [1973])].
Adelanthaceae Grolle, J. Hattori Bot. Lab. 35: 327 (1972). Adelanthus Mitt. nom. cons.

[including Pseudomarsupidium Herzog], Calyptrocolea R. M. Schust., Wettsteinia Schiffn.
Jamesoniellaceae He-Nygrén, Juslén, Ahonen, Glenny & Piippo, Cladistics 22: 27 (2006). Anomacaulis (R.

M. Schust.) Grolle, Cryptochila R.M.Schust., Cuspidatula Steph., Denotarisia Grolle, Jamesoniella (Spruce)
Carrington, Nothostrepta R. M. Schust., Pisanoa Hässel, Protosyzygiella (Inoue) R. M.
Schust., Roivainenia Perss., Syzygiella Spruce, Vanaea (Inoue & Gradst.) Inoue & Gradst.
Cephaloziaceae Mig., Krypt.-Fl. Deutschl., Moose: 465 (1904). Alobiella (Spruce) Schiffn., Alobiellopsis R.

M. Schust., Anomoclada Spruce, Cephalozia (Dumort.) Dumort., Cladopodiella H.
Buch, Fuscocephaloziopsis Fulford, Haesselia Grolle & Gradst., Hygrobiella Spruce, Iwatsukia N.
Kitag., Metahygrobiella R. M. Schust., Nowellia Mitt., Odontoschisma (Dumort.)
Dumort., Pleurocladula Grolle, Schiffneria Steph., Schofieldia J. D. Godfrey, Trabacellula Fulford
Cephaloziellaceae Douin, Bull. Soc. Bot. France, Mém. 29: 1, 5, 13 (1920). Allisoniella E. A. Hodgs.

[including Protomarsupella R. M. Schust.], Amphicephalozia R. M.
Schust., Cephalojonesia Grolle, Cephalomitrion R. M. Schust., Cephaloziella (Spruce) Schiffn. nom.
cons., Cephaloziopsis (Spruce) Schiffn., Cylindrocolea R. M. Schust., Gymnocoleopsis (R. M. Schust.) R. M.
Schust., Kymatocalyx Herzog [including Stenorrhipis Herzog]
Scapaniaceae Mig., Krypt.-Fl. Deutschl., Moose: 479 (1904) [including Chaetophyllopsidaceae R. M.
Schust., J. Hattori Bot. Lab. 23: 68. (1960 [1961]), Diplophyllaceae Potemk., Ann. Bot. Fenn. 36: 281 (1999),
and Lophoziaceae Cavers, New Phytol. 9: 293 (1910)]. Anastrepta (Lindb.)
Schiffn., Anastrophyllum (Spruce) Steph., Andrewsianthus R. M. Schust. [including Cephalolobus R. M.
Schust.], Barbilophozia Loeske, Chaetophyllopsis R. M.
Schust., Chandonanthus Mitt., Diplophyllum (Dumort.) Dumort. nom. cons., Douinia (C. N. Jensen) H.
Buch, Gerhildiella Grolle, Gottschelia Grolle, Gymnocolea (Dumort.) Dumort., Hattoria R. M.
Schust., Isopaches H. Buch, Krunodiplophyllum Grolle, Lophozia (Dumort.) Dumort., Macrodiplophyllum (H.
Buch) Perss., Plicanthus R. M. Schust., Pseudocephaloziella R. M. Schust., Scapania (Dumort.) Dumort.
nom. cons., Scapaniella H. Buch, Schistochilopsis (N. Kitag.) Konst., Sphenolobopsis R. M. Schust. & N.
Kitag., Sphenolobus (Lindb.) Berggr., Tetralophozia (R. M. Schust.) Schljakov, Tritomaria Schiffn. ex Loeske
SUBORDER: Jungermanniineae R. M. Schust. ex Stotler & Crand.-Stotl. in A. J. Shaw & B. Goffinet,

Bryoph. Biol.: 64 (2000).
Myliaceae Schljakov, Novosti Sist. Nizsh. Rast. 12: 308 (1975). Leiomylia J. J. Engel &
Braggins, Mylia Gray
Trichotemnomataceae R. M. Schust., J. Hattori Bot. Lab. 36: 340 (1972 [1973]). Trichotemnoma R. M.

Schust.
Balantiopsidaceae H. Buch, Mitt. Thüring. Bot. Ges. 1(2-3): 23 (1955). Anisotachis R. M.

Schust., Acroscyphella N. Kitag. & Grolle [ Austroscyphus R. M. Schust. nom.
illeg.], Balantiopsis Mitt., Eoisotachis R. M. Schust., Hypoisotachis (R. M. Schust.) J. J. Engel & G. L. S.
Merr., Isotachis Mitt., Neesioscyphus Grolle, Ruizanthus R. M. Schust.
Acrobolbaceae E. A. Hodgs., Rec. Domin. Mus. 4: 177 (1962). Acrobolbus Nees, Austrolophozia R. M.

Schust., Enigmella G.A. M.Scott & K. G. Beckm., Goebelobryum Grolle, Lethocolea Mitt. nom. cons.
[including Neoprasanthus S. Winkl.], Marsupidium Mitt., Tylimanthus Mitt.
Blepharidophyllaceae R. M. Schust., Nova Hedwigia Beih. 119: 491
(2002). Blepharidophyllum Ångstr., Clandarium (Grolle) R. M. Schust.
Arnelliaceae Nakai, Chosakuronbun Mokuroku [Ord. Fam. Trib. Nov.]: 200

(1943). Arnellia Lindb., Gongylanthus Nees, Southbya Spruce, Stephaniella J. B. Jack, Stephaniellidium S.
Winkl. ex Grolle
Jackiellaceae R. M. Schust., J. Hattori Bot. Lab. 36: 395. (1972 [1973]). Jackiella Schiffn.
Calypogeiaceae Arnell in Holmberg, Skand. Fl. 2a.: 189 (1928). Calypogeia Raddi nom.

cons., Eocalypogeia (R. M. Schust.) R. M. Schust., Metacalypogeia (S. Hatt.) Inoue, Mnioloma Herzog
Delavayellaceae R. M. Schust., Bryologist 64: 202 (1961). Delavayella Steph., LiochlaenaNees
Jungermanniaceae Rchb., Bot. Damen: 256 (1828).[including Mesoptychiaceae Inoue & Steere, Bull. Natl.
Sci. Mus. Tokyo, B 1: 62 (1975).] Eremonotus Lindb. & Kaal. ex Pearson [including Anomomarsupella R. M.
Schust.], Hattoriella (Inoue) Inoue, Jungermannia L., Leiocolea (Müll. Frib.) H. Buch, Mesoptychia (Lindb.)
A. Evans
Solenostomataceae Stotler & Crand.-Stotl., Edinburgh J. Bot. 66: 190

(2009). Arctoscyphus Hässel, Bragginsella R. M. Schust., Cryptocolea R. M.
Schust., Cryptocoleopsis Amak., Cryptostipula R. M. Schust., Diplocolea Amak., Horikawaella S. Hatt. &
Amakawa, Nardia Gray nom. cons. [including Apotomanthus (Spruce)
Schiffn.], Scaphophyllum Inoue, Solenostoma Mitt. [including Plectocolea (Mitt.) Mitt.]
Geocalycaceae H. Klinggr., Höh. Crypt. Preuss.: 34

(1858). Geocalyx Nees, Harpanthus Nees, Notoscyphus Mitt., Saccogyna Dumort. nom.
cons., Saccogynidium Grolle,
Gyrothyraceae R. M. Schust., Trans. Brit. Bryol. Soc. 6: 87 (1970). Gyrothyra M. Howe

Antheliaceae R. M. Schust., J. Hattori Bot. Lab. 26: 236 (1963). Anthelia (Dumort.) Dumort.
Gymnomitriaceae H. Klinggr., Höh. Crypt. Preuss.: 16 (1858). Acrolophozia R. M.

Schust., Apomarsupella R. M. Schust., Gymnomitrion Corda nom.
cons., Herzogobryum Grolle, Lophonardia R. M. Schust., Marsupella Dumort., Nanomarsupella (R. M.
Schust.) R. M. Schust., Nothogymnomitrion R. M. Schust., Paramomitrion R. M.
Schust., Poeltia Grolle, Prasanthus Lindb.
Haplomitrium mnioides (Lindb.) R. M. Schust.
Photo taken by Li Zhang

China. Hong Kong, August 2002.

Marchantia polymorpha L.
Photo taken by Li Zhang,

China, Hong Kong, June 2002.
Green house cultivated, SIUC, April 2004.

Female
Malaysia. Penang. Cameron Highlands. Brinchang, Country Lodge Resort. 1500 m. On soil wall, 1500 m. Li
Zhang 4474, July 28, 2007. (voucher in SZG)

Male

Malaysia. Penang. Cameron Highlands. Brinchang, Country Lodge Resort. 1500 m. On soil wall, 1500 m. Li
Zhang 4474, July 28, 2007. (voucher in SZG)
Asterella tenella (L.) P. Beauv.

Photo by Li Zhang
Thalli with purple scales at growing tips; note the rounded purple button-like aborted female
gametangiophore bordered by a fringe of small scales near the center of the photo. This branch did not
elongated to elevate the carpocephalum above the thallus.
Found on an exposed sandstone rock glade, April 17, 2005 in southern Illinois at Bell Smith Springs (Pope
County) by Dylan Kosma;
Asterella tenella (L.) P. Beauv.
Photo by Li Zhang
Carpocephala with immature sporophtes; note the reddish tinge on the stalks and pseudoperianth scales
that surround each developing sporophyte.
Found on an exposed sandstone rock glade, April 17, 2005 in southern Illinois at Bell Smith Springs (Pope
County) by Dylan Kosma;

Asterella tenella (L.) P. Beauv. [identified by study of a paradermal section to view the dorsal epidermal cell
walls and the pattern of the cells surrounding the air chamber openings]
Photo by Li Zhang
USA. Illinois. Johnson Co. Ferne Clyffe State Park, Black Jack Oak Trail. On sandy soil on exposed
sandstone rock glade, 180 m, May 31, 2009. Zhang 6331 (voucher in SZG)
Plagiochasma sp.
CHINA. Guizhou. Pu’an Co., Panshui. Dongfeng Reservoir, on rock by path in sparse forest, 1595
m, Zhang 5883, Oct 3, 2008. (voucher at SZG
Reboulia hemisphaerica (L.) Raddi
USA. Southern Illinois, April 2004.

Reboulia hemisphaerica (L.) Raddi

Monosolenium tenerum Griff.
China, Macau, Macau Peninsula. On soil in flower bed, 10 m., February 15, 2007. (voucher in SZG)
Monosolenium tenerum Griff.
China, Macau, Macau Peninsula. On soil in flower bed, 10 m., February 15, 2007. (voucher in SZG)
Conocephalum conicum (L.) Corda.
Photos taken by Li Zhang
( Note air pores and the arrangement of air chambers)

Green house cultivated, SIUC, April 2004
( Note air pores and the arrangement of air chambers)
Green house cultivated, SIUC, April 2004


( Note female receptacle )
Collected by Dylan Kosma in Southern Illinois, March 2005.

Cyathodium sp.
Malaysia. Kuala Lumpur. University of Malaya Botanical Garden. On moist rock in greenhouse. Li Zhang
4467, July 25, 2007. (voucher in SZG)
Exormotheca sp.

September 2004
Riccia lamellosa Gottsche [NB: the small Riccia to the lower left is R. hirta (Austin) Underw.]
Photo by Li Zhang
USA. Illinois. Johnson Co. Ferne Clyffe State Park. Black Jack Oak Trail. On sandy soil by path, 180 m,
May 31, 2009. ZHANG 6329 (voucher in SZG)
Monoclea gottschei Lindb. subsp. elongata Gradst. & Mues
Green house cultivated, SIUC. (Note yellow oil cells). April 2004.
Monoclea gottschei Lindb. subsp. elongata Gradst. & Mues
Green house cultivated, SIUC. (Note yellow oil cells). April 2004
Dumortiera hirsuta (Swaegr.) Nees ssp. nepalensis (Tay.) Frye & Clark
China, Guangdong, May 2003.

Dumortiera hirsuta (Swaegr.) Nees
China, Guangdong, August 2004.

Noteroclada confluens Taylor ex Hook. & Wilson
Greenhouse cultivated at the University of Göttingen, Göttingen, Germany

(plants originally from Ecuador)
Noteroclada confluens Taylor ex Hook. & Wilson

Greenhouse cultivated at the University of Göttingen, Göttingen, Germany
(plants originally from Brazil)
September 2004
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Petalophyllum americanum C. H. Ford & Crand.-Stotl.
Photo taken by B. Crandall-Stotler
USA. Texas. Buescher State Park.

Allisonia cockaynii (K. I. Goebel) R. M. Schust.
Photo taken with Bill Malcolm
NEW ZEALAND. Mt. Aurther, in sinkhole (voucher in ABSH)

Fossombronia japonica Schiffn.
Photo taken by Li Zhang.
China, Macau, Coloane Is. On soil in flower field, 10 m., January 26, 2007. (voucher in SZG
Phyllothallia navicola
Photo taken with Bill Malcolm
NEW ZEALAND. Nelson

Moerckia blytti (Moerck) Brockm.
September 2004.
Jensenia connivens (Colenso) Grolle
Photo taken by Barbara Crandall-Stotler
New Zealand: From a limestone sinkhole, Mt. Arthur, South Island

Pallavicinia levieri Schiffn.
China, Guangdong, August 2004

Pallavicinia levieri Schiffn.
China, Guangdong, August 2004

Xenothallus vulcanicola R. M. Schust.
Photo by Barbara Crandall-Stotler
New Zealand, South Island. Temple Basin. On soil shaded by overhanging grass.
Pleurozia acinosa (Mitt.) Trev.
CHINA. Hainan, China, November 2002

Aneura pinguis (L.) Dum.
CHINA. Hong Kong, September 2003.
Aneura pinguis (L.) Dumort.

CHINA. Macau, Coloane Is. January 25, 2007. On moist rock by stream in sparse forest, 100 m
Riccardia sp.
Photo taken by Li Zhang,
CHINA. Sichuan, July 2003.
Frullania sp.
China, Yunnan, August 2003

Acrolejeunea pusilla (Steph.) Grolle & Gradst.
CHINA. Macau, Coloane Is. N 22°06.900', E 113°33.259', on trunk, 10 m. Li Zhang 5613, April 12, 2007.
(voucher in SZG)
Schistochila sp.
CHINA. Hainan, November 2002

Trichocolea tomentella (Ehrn.) Dum.
CHINA. Sichuan, July 2003.
Trichocolea tomentella (Ehrh.) Dum.
USA. Southern illinois, April 2005.
Photo by Li Zhang
CHINA. XIZANG (TIBET). Chayu Co., Cibagou National Nature Reserve,
28° 45' 19.79” N, 97° 02' 27.84” E, on humus in swamp, 3010 m., 20 Aug. 2009, Li Zhang 6785 (SZG)
Herbertus sp.

China. Yunnan, August 2003.
Bazzania tridens (Reinw., Blume & Nees) Trevis.
CHINA. Hong Kong, June 2002

Lepidozia cf. trichodes (Reinw., Blume & Nees) Nees
Malaysia. Penang. Cameron Highlands. Brinchang, Mt. Irau. N 04°28', E 101°23'. 1935 m. On humus in
forest. Li Zhang 5720, July 29, 2007. (voucher in SZG)
Lophocolea heterophylla (Schrad.) Dum.
Collected by Dylan Kosma in Southern Illinois, March 2005.

Plagiochila porelloides (Torr.) Lindenb.

Gottschelia schizopleura (Spruce) Grolle
Malaysia. Penang. Cameron Highlands. Brinchang, Mt. Irau. N 04°28', E 101°23'. 1950 m. On rock by road.
Li Zhang 5736, July 29, 2007. (voucher in SZG)
Scapania nemorea (L.) Grolle
A sexually reproductive population; note the elongating seta terminated by a brown, obong-ovoid sporophyte
capsule. Following further seta elongation, the capsule will dehisce, splitting along 4 vertical lines, and
spores are released.
Found growing over moist, protected sandstone, April 17, 2005 in southern Illinois at Bell Smith Springs
(Pope County) by Dylan Kosma; photo by Li Zhang

Found on moist rock , June 3, 2005 in Southern Illinois at Jackson Co., Little Grand Canyon by Li Zhang;

CHINA. INNER MONGOLIA. Hulunbei’er. Genhe. Aokelidiu Mt., Mountbase. N 51° 49' 48.2”, E 122° 03'
01.4”. On moist rock along streamlet in Larix gemelinii forest, 1025 m., July 13, 2010, Li Zhang 7432
(voucher in SZG)
Gyrothyra underwoodiana M. Howe.
Collected by Laura Forrest in Vancouver Island, British Columbia, April 2004.

Gyrothyra underwoodiana M. Howe.
Collected by Laura Forrest in Vancouver Island, British Columbia, April 2004.

THE GENERA OF LIVERWORTS
[Last Update March 2011]
This alphabetical list of all accepted liverwort genera is the first to include the number of species for each
[given in square brackets]. Species numbers follow contemporary taxonomic concepts with the source of
these data referenced parenthetically to the literature listed below. Species numbers based upon our
unpublished studies are given as (S & C-S) for Stotler and Crandall-Stotler. While most of the species
numbers are widely accepted, others represent approximations. For example, the number of species in the
two very largest genera of liverworts, namely Plagiochila and Frullania, was taken from Gradstein et al. (28)
and appears to be very conservative. More recently used generic names regarded as synonyms are given
in bold.
Alphabetical Jump Station

A B C D E F G H I J K L M N O P Q R S T U V W X Y Z

Acanthocoleus R. M. Schust. [7-8] - Lejeuneaceae (28)
Acantholejeunea (R. M. Schust.) R. M. Schust. [monotypic] - Lejeuneaceae (63)
Acrobolbus Nees [10-12] - Acrobolbaceae (28)
Acrochila R. M. Schust. [2] - Plagiochilaceae (46)
Acrolejeunea (Spruce) Schiffn. [15] - Lejeuneaceae (28)
Acrolepidozia R. M. Schust. = Telaranea (65)
Acrolophozia R. M. Schust. [3] - Gymnomitriaceae (99)
Acromastigum A. Evans [35-40] - Lepidoziaceae (66)
Acroscyphella N. Kitag. & Grolle [4] - Balantiopsidaceae (60)

Acroscyphus N. Kitag. = Acroscyphella (105)
Adelanthus Mitt. [15] - Adelanthaceae (28)
Aitchisoniella Kashyap [monotypic] - Exormothecaceae (5)
Allisonia Herzog [monotypic] - Allisoniaceae (98)
Allisoniella E. A. Hodgs. [5] - Cephaloziellaceae (68)
Allorgella Tixier = Otolejeunea (87)
Alobiella (Spruce) Schiffn. [2] - Cephaloziaceae (68)
Alobiellopsis R. M. Schust. [5] - Cephaloziaceae (68)
Amazoopsis J. J. Engel & G. L. S. Merr. [3] - Lepidoziaceae (85)
Amblyolejeunea Ast [monotypic] - Lejeuneaceae (28)

Amphicephalozia R. M. Schust. [monotypic] - Cephaloziellaceae (68)
Amphijubula R. M. Schust. = Frullania (98)
Amphilejeunea R. M. Schust. = Lejeunea (80)
Amphilophocolea R. M. Schust. [monotypic] - Lophocoleaceae
Anastrepta (Lindb.) Schiffn. [monotypic] - Anastrophyllaceae (76; 100)
Anastrophyllum (Spruce) Steph. [35-40] - Anastrophyllaceae (28; 100)
Andrewsianthus R. M. Schust. [30] - Scapaniaceae (28)
Aneura Dumort. [10] - Aneuraceae (28)
Anisotachis R. M. Schust. [monotypic] - Balantiopsidaceae (4)
Anomacaulis (R. M. Schust.) Grolle [monotypic] - Adelanthaceae (68; 102)

Anomoclada Spruce [monotypic] - Cephaloziaceae (28)
Anomomarsupella R. M. Schust. = Eremonotus (98)
Anomylia R. M. Schust. = Leptoscyphus
Anoplolejeunea (Spruce) Schiffn. [monotypic] - Lejeuneaceae (28)
Anoplostoma E. A. Hodgs. & Allison = Herzogianthus
Anthelia (Dumort.) Dumort. [2] - Antheliaceae (66)
Apertithallus Kuwah. = Metzgeria
Aphanolejeunea A. Evans = Cololejeunea (98)
Aphanotropis Herzog [monotypic] - Lejeuneaceae (59)
Apomarsupella R. M. Schust. [4] - Gymnomitriaceae (99)

Apometzgeria Kuwah. = Metzgeria (98)
Apotreubia S. Hatt. & Mizut. [4] - Treubiaceae (44)
Arachniopsis Spruce = Telaranea (85)
Archeochaete R. M. Schust. = Pseudolepicolea (98)
Archeophylla R. M. Schust. [3] - Pseudolepicoleaceae (66)
Archilejeunea (Spruce) Schiffn. [15] - Lejeuneaceae (28)
Arctoscyphus Hässel [1-2] - Jungermanniaceae (66; 42)
Arnellia Lindb. [monotypic] - Arnelliaceae (9)
Ascidiota C. Massal. [monotypic] - Porellaceae (43)
Asterella P. Beauv. [80] - Aytoniaceae (5)

Athalamia Falc. [12] - Cleveaceae (5)
Aureolejeunea R. M. Schust. [5-6] - Lejeuneaceae (28)
Austrofossombronia R. M. Schust. = Fossombronia (98)
Austrolejeunea (R. M. Schust.) R. M. Schust. [9] - Lejeuneaceae (94)
Austrolembidium Hässel = Evansianthus (66)
Austrolophozia R. M. Schust. [3] - Acrobolbaceae (67)
Austrometzgeria Kuwah. = Metzgeria (98)
Austroscyphus R. M. Schust. = Acroscyphella (98)
Top
Balantiopsis Mitt. [11] - Balantiopsidaceae (28)

Barbilophozia Loeske [5] - Anastrophyllaceae (100)
Bazzania Gray [100] - Lepidoziaceae (28)
Biantheridion (Grolle) Konstant. & Vilnet. [monotypic] - Anastrophyllaceae (100; 101)
Blasia L. [monotypic] - Blasiaceae (9)
Blepharidophyllum Ångstr. [5] - Blepharidophyllaceae (68)
Blepharolejeunea S. W. Arnell [5] - Lejeuneaceae (28)
Blepharostoma (Dumort.) Dumort. [3] - Pseudolepicoleaceae (66)
Bonneria Fulford & J. Taylor = Paracromastigum
Brachiolejeunea (Spruce) Schiffn. [6] - Lejeuneaceae (28)
Brachygyna R. M. Schust. = Trichocolea

Bragginsella R. M. Schust. [monotypic] - Jungermanniaceae (68)
Brevianthus J. J. Engel & R. M. Schust. [monotypic] - Brevianthaceae (17)
Bromeliophila R. M. Schust. [2] - Lejeuneaceae (28)
Bryopteris (Nees) Lindenb. [7] - Lejeuneaceae (74)
Bucegia Radian [monotypic] - Marchantiaceae (5)
Byssolejeunea Herzog = Lejeunea
Calatholejeunea K. I. Goebel [monotypic] - Lejeuneaceae (6) Top
Calycularia Mitt. [2] - Calyculariaceae (28)
Calypogeia Raddi [35] - Calypogeiaceae (28)
Calyptrocolea R. M. Schust. [9-10] - Adelanthaceae (68)

Campanocolea R. M. Schust. = Lophocolea (98)
Campylolejeunea S. Hatt. = Cololejeunea
Capillolejeunea S. W. Arnell = Drepanolejeunea (82)
Cardiolejeunea R. M. Schust. & Kachroo = Lejeunea
Carrpos Prosk. = Monocarpus (5)
Castanoclobus J. J. Engel & Glenny [monotypic] - Pseudolepicoleaceae (S & C-S)
Caudalejeunea (Steph.) Schiffn. [15] - Lejeuneaceae (28)
Cavicularia Steph. [monotypic] - Blasiaceae (51)
Cephalantholejeunea (R. M. Schust. & Kachroo) R. M. Schust. [monotypic] - Lejeuneaceae (28)
Cephalojonesia Grolle [monotypic] - Cephaloziellaceae (4)

Cephalolejeunea Mizut. [monotypic] - Lejeuneaceae (47)
Cephalolobus R. M. Schust. = Andrewsianthus (68)
Cephalomitrion R. M. Schust. [monotypic] - Cephaloziellaceae (68)
Cephalozia (Dumort.) Dumort. [24] - Cephaloziaceae (28)
Cephaloziella (Spruce) Schiffn. [50-125] - Cephaloziellaceae (28; 50)
Cephaloziopsis (Spruce) Schiffn. [3] - Cephaloziellaceae (68)
Ceratolejeunea (Spruce) J. B. Jack & Steph. [30] - Lejeuneaceae (28)
Chaetocolea Spruce [monotypic] - Pseudolepicoleaceae (66)
Chaetophyllopsis R. M. Schust. [monotypic] - Scapaniaceae (52)
Chandonanthus Mitt. [monotypic] - Anastrophyllaceae (91; 99)

Cheilolejeunea (Spruce) Schiffn. [60-70] - Lejeuneaceae (28)
Chiastocaulon Carl [ ] - Plaqiochilaceae
Chiloscyphus Corda [ca 100] - Lophocoleaceae (59)
Chloranthelia R. M. Schust. [2] - Lepidoziaceae (66)
Chondriolejeunea (Benedix) Kis & Pócs [ ] - Lejeuneaceae
Chonecolea Grolle [3] - Chonecoleaceae (28)
Cladocolea R. M. Schust. = Schusterolejeunea
Chrondrophyllum Herzog non Kylin = Herzogobryum Grolle
Cladolejeunea Zwick. [monotypic] - Lejeuneaceae (6)
Cladomastigum Fulford = Iwatsukia
Cladopodiella H. Buch [2] - Cephaloziaceae (9)
Clandarium (Grolle) R. M. Schust. [3] - Blepharidophyllaceae (68)
Clasmatocolea Spruce [20] - Lophocoleaceae (11)
Cololejeunea (Spruce) Schiffn. [30-40] - Lejeuneaceae (28)
Colura (Dumort.) Dumort. [80] - Lejeuneaceae (28)
Conocephalum Hill [3]- Conocephalaceae (5)
Conoscyphus Mitt. [monotypic] - Lophocoleaceae (4)
Corsinia Raddi [monotypic] - Corsiniaceae (5)
Cronisia Berk. [1-2] - Corsiniaceae (5)
Crossocalyx Meylan [ ] - Anastrophyllaceae (100)

Crossogyna (R. M. Schust.) Schljakov = Jamesoniella
Crossotolejeunea (Spruce) Schiffn. = Lejeunea (104)
Cryptochila R. M. Schust. [6] = Adelanthaceae (68; 102)
Cryptocolea R. M. Schust. [monotypic] - Solenostomataceae (76; 98)
Cryptocoleopsis Amak. [monotypic] - Solenostomataceae (76; 98)
Cryptogynolejeunea R. M. Schust. = Lejeunea (80)
Cryptolejeunea R. M. Schust. & Kachroo = Echinocolea
Cryptomitrium Austin ex Underw. [3] - Aytoniaceae (5)
Cryptostipula R. M. Schust. [monotypic] - Jungermanniaceae (68)
Cryptothallus Malmb. = Aneura
Cuspidatula Steph. [ ] - Adelanthaceae (102)
Cyanolophocolea (R. M. Schust.) R. M. Schust. [monotypic] - Lophocoleaceae (84)
Cyathodium Kunze ex Lehm [14] - Cyathodiaceae (5)
Cyclolejeunea A. Evans [6-7] - Lejeuneaceae (28)
Cylindrocolea R. M. Schust. [12] - Cephaloziellaceae (28)
Cyrtolejeunea A. Evans = Cheilolejeunea (98)
Cystolejeunea A. Evans [monotypic] - Lejeuneaceae (28)
Dactylolejeunea R. M. Schust. = Lejeunea (104) Top
Dactylophorella R. M. Schust. [monotypic] - Lejeuneaceae (63)
Delavayella Steph. [monotypic] - Delavayellaceae (76)

Dendrobazzania R. M. Schust. & W. B. Schofield [monotypic] - Lepidoziaceae (72)
Dendrolejeunea (Spruce) Lacout. [monotypic] - Lejeuneaceae (27)
Dendrolembidium Herzog = Kurzia
Dendromastigophora R. M. Schust. [monotypic] - Mastigophoraceae (61)
Denotarisia Grolle [monotypic] - Adelanthaceae (68; 102)
Dianthelia R. M. Schust. = Gymnomitrion
Dicladolejeunea R. M. Schust. = Lejeunea (81)
Dicranolejeunea (Spruce) Schiffn. [monotypic] - Lejeuneaceae (28)
Dinckleria Trevis. [2] - Plagiochilaceae (106)
Diplasiolejeunea (Spruce) Schiffn. [62] - Lejeuneaceae (77)

Diplocolea Amak. [monotypic] - Jungermanniaceae (76)
Diplophyllum (Dumort.) Dumort. [20] - Scapaniaceae (28)
Douinia (C. E. O. Jensen) H. Buch [monotypic] - Scapaniaceae (76)
Drepanolejeunea (Spruce) Schiffn. [100] - Lejeuneaceae (28)
Drucella E. A. Hodgs. [monotypic] - Lepidoziaceae (66)
Dumortiera Nees [monotypic] - Dumortieraceae (5)
Echinocolea R. M. Schust. [3-4] - Lejeuneaceae (28) Top
Echinolejeunea R. M. Schust. [monotypic] - Lejeuneaceae (63)
Enigmella G. A. M. Scott & K. G. Beckm. [monotypic] - Acrobolbaceae (2)
Eocalypogeia (R. M. Schust.) R. M. Schust. [2] - Calypogeiaceae (66)

Eoisotachis R. M. Schust. [monotypic] - Balantiopsidaceae (55)
Eopleurozia R. M. Schust. = Pleurozia (98)
Eotrichocolea R. M. Schust. [monotypic] - Trichocoleaceae (66)
Eremonotus Lindb. & Kaal. ex Pearson [monotypic] - Jungermanniaceae (76; 98)
Evansianthus R. M. Schust. & J. J. Engel [monotypic] - Lophocoleaceae (71)
Evansiolejeunea Vanden Berghen [2] - Lejeuneaceae (7)
Exormotheca Mitt. [7] - Exormothecaceae (5)
Fossombronia Raddi [ca 70] - Fossombroniaceae (S & C-S) Top
Frullania Raddi [ca 300+] - Frullaniaceae (28; 98)

Frullanoides Raddi [7] - Lejeuneaceae (28)
Fulfordianthus Gradst. [2] - Lejeuneaceae (28)
Fulfordiella Hässel = Isophyllaria
Fulfordistria H. A. Miller = Schistochila
Fuscocephaloziopsis Fulford [2] - Cephaloziaceae (68)
Gackstroemia Trevis. [6] - Lepidolaenaceae (33) Top
Geocalyx Nees [5-6] - Geocalycaceae (9)
Geothallus Campb. [monotypic] - Sphaerocarpaceae (62)
Gerhildiella Grolle [monotypic] - Scapaniaceae (76)

Goebeliella Steph. [monotypic] - Goebeliellaceae (56)
Goebelobryum Grolle [2] - Acrobolbaceae (31)
Gongylanthus Nees [5-6] - Arnelliaceae (28)
Gottschea Nees ex Mont. [24] - Schistochilaceae (39)
Gottschelia Grolle [2] - Jungermanniaceae (68)
Greenothallus Hässel [monotypic] - Pallaviciniaceae (S & C-S)
Grollea R. M. Schust. [monotypic] - Grolleaceae (66)
Gymnocolea (Dumort.) Dumort. [7] - Anastrophyllaceae (76; 100)
Gymnocoleopsis (R. M. Schust.) R. M. Schust. [3] - Jungermanniaceae (68)
Gymnomitrion Corda [27] - Gymnomitriaceae (99)

Gyrothyra M. Howe [monotypic] - Gyrothyraceae (8)
Haesselia Grolle & Gradst. [monotypic] - Cephaloziaceae (68) Top
Hamatostrepta Vána & D. G. Long [monotypic] - Anastrophyllaceae (100)
Haplochaete R. M. Schust. = Isophyllaria
Haplolejeunea Grolle [2] - Lejeuneaceae (28)
Haplomitrium Nees [8] - Haplomitriaceae (1)
Harpalejeunea (Spruce) Schiffn. [20-25] - Lejeuneaceae (28)
Harpanthus Nees [3] - Geocalycaceae (9)
Hattoria R. M. Schust. [monotypic] - Scapaniaceae (76)

Hattorianthus R. M. Schust. & Inoue [monotypic] - Moerckiaceae (S & C-S)
Hattoriella (Inoue) Inoue [ ] - Jungermanniaceae (98)
Hattoriolejeunea Mizut. [monotypic] - Lejeuneaceae (48)
Hepatostolonophora J. J. Engel & R. M. Schust. [3] - Lophocoleaceae (10)
Herbertus Gray [50] - Herbertaceae (66)
Herzogianthus R. M. Schust. [2] - Herzogiantaceae (61)
Herzogiaria Fulford ex Hässel [monotypic] - Pseudolepicoleaceae (66)
Herzogobryum Grolle [5] - Gymnomitriaceae (99)
Heterogemma (Jørg.) Konstant. & Vilnet [2] - Scapaniaceae (101)
Heteroscyphus Schiffn. [30] - Lophocoleaceae (28)

Hodgsonia Perss. = Neohodgsonia
Horikawaella S. Hatt. & Amakawa [monotypic] - Jungermanniaceae (68)
Hyalolepidozia S. W. Arnell ex Grolle [2-3] - Lepidoziaceae (66)
Hygrobiella Spruce [2]- Hygrobiellaceae (9; 101)
Hygrolembidium R. M. Schust. [8] - Lepidoziaceae (66)
Hymenophyton Dumort. [2] - Hymenophytaceae (S & C-S)
Hypoisotachis (R. M. Schust.) J. J. Engel & G. L. S. Merr. [monotypic] - Balantiopsidaceae (20)
Hypolophozia (R. M. Schust.) Bakalin [2] - Scapaniaceae (99; 107)
Inflatolejeunea S. Arn. = Lejeunea (93) Top

[Invisocaulis R. M. Schust., nom. invalid. [monotypic] - Jungermanniaceae (68) ]
Isolembidium R. M. Schust. [monotypic] - Lepidoziaceae (66)
Isopaches H. Buch [ ] - Anastrophyllaceae (100)
Isophyllaria E. A. Hodgs. & Allison [monotypic] - Pseudolepicoleaceae (66)
Isotachis Mitt. [15] - Balantiopsidaceae (28)
Iwatsukia N. Kitag. [4] - Cephaloziaceae (68)
Jackiella Schiffn. [5-6] - Jackiellaceae (68) Top
Jamesoniella (Spruce) Carrington = Syzygiella (102)
Jensenia Lindb. [8] - Pallaviciniaceae (S & C-S)
Jovetastella Tixier = Cololejeunea
Jubula Dumort. [4] - Jubulaceae (40)
Jubulopsis R. M. Schust. [monotypic] - Jubulopsidaceae (58)
Jungermannia L. [120-125] - Jungermanniaceae (76)
Kingiolejeunea H. Robs. = Lepidolejeunea Top
Krunodiplophyllum Grolle [ ] - Scapaniaceae
Kurzia G. Martins [30-34] - Lepidoziaceae (66)
Kymatocalyx Herzog [4] - Cephaloziellaceae (28)
Kymatolejeunea Grolle [monotypic] - Lejeuneaceae (35)

Lamellocolea J. J. Engel [monotypic] - Lophocoleaceae (14) Top
Leiocolea (Müll. Frib.) H. Buch [ ] - Jungermanniaceae (98)
Leiolejeunea A. Evans [monotypic] - Lejeuneaceae (28)
Leiomitra Lindb. [10] - Trichocoleaceae (91)
Leiomylia J. J. Engels' Braggins [ ] - Jungermanniaceae
Lejeunea Lib. [100+] - Lejeuneaceae (29)
Lembidium Mitt. [2] - Lepidoziaceae (66)
Lepicolea Dumort. [10] - Lepicoleaceae (66)
Lepidogyna R. M. Schust. [3] - Lepidolaenaceae (10)
Lepidolaena Dumort. [9] - Lepidolaenaceae (33; 35)

Lepidolejeunea R. M. Schust. [12] - Lejeuneaceae (28)
Lepidozia (Dumort.) Dumort. [75] - Lepidoziaceae (28)
Lepidoziopsis E. A. Hodgs. = Lepidozia
Leptocolea (Spruce) A. Evans = Cololejeunea
Leptolejeunea (Spruce) Schiffn. [25] - Lejeuneaceae (28)
Leptophyllopsis R. M. Schust. [monotypic] - Lophocoleaceae (53)
Leptoscyphopsis R. M. Schust. [monotypic] - Lophocoleaceae (28)
Leptoscyphus Mitt. [17] - Lophocoleaceae (28)
Leptostictolejeunea R. M. Schust. = Stictolejeunea
Lethocolea Mitt. [5-6] - Acrobolbaceae (28)

Leucolejeunea A. Evans [13] - Lejeuneaceae (28)
Leucosarmentum Fulford = Paracromastigum
Lindigianthus Kruijt & Gradst. [monotypic] - Lejeuneaceae (28)
Liochlaena [ ] - Delavayellaceae (98)
Lobatiriccardia (Mizut. & S. Hatt.) Furuki [4] - Aneuraceae (21)
Lophochaete R. M. Schust. = Pseudolepicolea
Lophocolea (Dumort.) Dumort. [>100; ca 223] - Lophocoleaceae (19; 28)
Lopholejeunea (Spruce) Schiffn. [30] - Lejeuneaceae (108)
Lophonardia R. M. Schust. = Hypolophozia ( )
Lophozia (Dumort.) Dumort. [65-70] - Scapaniaceae (28)

Lunularia Adans. [monotypic] - Lunulariaceae (5)
Luteolejeunea Piippo [monotypic] - Lejeuneaceae (28)
Macrocolura R. M. Schust. [monotypic] - Lejeuneaceae (64) Top
Macrodiplophyllum (H. Buch) Perss. [ ] - Scapaniaceae
Macrolejeunea (Spruce) Schiffn. [3-5] - Lejeuneaceae (28)
Maculia E. A. Hodgs. = Chloranthelia
Macvicaria W. E. Nicholson = Porella (98)
Makednothallus Verd. = Jensenia
Makinoa Miyake [monotypic] - Makinoaceae (5)

Mannia Opiz [15] - Aytoniaceae (5)
Marchantia L. [36] - Marchantiaceae (5)
Marchastra E. O. Campb. = Neohodgsonia
Marchesinia Gray [5] - Lejeuneaceae (28)
Marsupella Dumort. [40-45] - Gymnomitriaceae (68)
Marsupidium Mitt. [12] - Acrobolbaceae (28)
Massula Müll. Frib. ex Schljakov = Lophozia
Mastigolejeunea (Spruce) Schiffn. [15] - Lejeuneaceae (28)
Mastigopelma Mitt. [3] - Lepidoziaceae (66)
Mastigophora Nees [3] - Mastigophoraceae (61)

Megalembidium R. M. Schust. [monotypic] - Lepidoziaceae (66)
Meinungeria Frank Müll. [1] - Lepidoziaceae (97)
Mesoptychia (Lindb.) A. Evans [monotypic] - Jungermanniaceae (76; 98)
Metacalypogeia (S. Hatt.) Inoue [2] - Calypogeiaceae (66)
Metacephalozia Inoue = Cephaloziopsis
Metahygrobiella R. M. Schust. [6] - Cephaloziaceae (68)
Metalejeunea Grolle [monotypic] - Lejeuneaceae (28)
Metzgeria Raddi [100+] - Metzgeriaceae (28)
Metzgeriopsis K. I. Goebel = Cololejeunea ( )
Micrisophylla Fulford = Kurzia
Microlejeunea Steph. [20-30] - Lejeuneaceae (28)
Microlepidozia (Spruce) Jörg. = Kurzia
Micropterygium Lindenb., Nees & Gottsche [18-19] - Lepidoziaceae (28)
Mizutania Furuki & Z. Iwats. [monotypic] - Mizutaniaceae (22)
Mnioloma Herzog [10] - Calypogeiaceae (28)
Moerckia Gottsche [2] - Moerckiaceae (S & C-S)
Monocarpus D. J. Carr [monotypic] - Monocarpaceae (5)
Monoclea Hook. [2] - Monocleaceae (28)
Monodactylopsis (R. M. Schust.) R. M. Schust. [2] - Lepidoziaceae (66; 85)
Monosolenium Griff. [monotypic] - Monosoleniaceae (5)

Mylia Gray [4] - Myliaceae
Myriocolea Spruce [monotypic] - Lejeuneaceae (28)
Myriocoleopsis Schiffn. [2] - Lejeuneaceae (28)
Mytilopsis Spruce [monotypic] - Lepidoziaceae (66)
Nanomarsupella (R. M. Schust.) R. M. Schust. [monotypic] - Gymnomitriaceae (68)

Top
Nardia Gray [16] - Jungermanniaceae (28)
Nardiocalyx Lindb. ex Jörg. = Gymnomitrion
Neesioscyphus Grolle [5] - Balantiopsidaceae (28)
Neogrollea E. A. Hodgs. [monotypic] - Neogrolleaceae (15)

Neohattoria Kamim. = Frullania
Neohodgsonia Perss. [monotypic] - Neohodgsoniaceae (5)
Neolepidozia Fulford & J. Taylor = Telaranea
Neopotamolejeunea E. Reiner = Lejeunea (95)
Neoprasanthus S. Winkl. = Lethocolea
Neotrichocolea S. Hatt. [monotypic] - Neotrichocoleaceae (24)
Nephelolejeunea Grolle = Austrolejeunea (94)
Nesolejeunea Herzog = Lejeunea
Neurolejeunea (Spruce) Schiffn. [4] - Lejeuneaceae (28)
Nipponolejeunea S. Hatt. [2] - Jubulaceae (63)

Noguchia S. Hatt. = Plagiochilion
Noteroclada Taylor ex Hook. & Wils. [monotypic] - Pelliaceae (S & C-S)
Nothogymnomitrion R. M. Schust. [monotypic] - Gymnomitriaceae (68)
Nothostrepta R. M. Schust. [2] - Adelanthaceae (68; 102)
Notoscyphus Mitt. [monotypic] - Jungermanniaceae (68)
Nowellia Mitt. [9] - Cephaloziaceae (68)
Odontolejeunea (Spruce) Schiffn. [3] - Lejeuneaceae (28) Top
Odontoschisma (Dumort.) Dumort. [10-12] - Cephaloziaceae (28)
Odontoseries Fulford [monotypic] - Lepidoziaceae (66)

Olgantha R. M. Schust. [monotypic] - Herbertaceae (66)
Omphalanthus Lindenb. & Nees [4-5] - Lejeuneaceae (28)
Ophthalmolejeunea (R. M. Schust.) R. M. Schust. = Drepanolejeunea
Oreolejeunea R. M. Schust. = Blepharolejeunea
Orthocaulis H. Buch = Barbilophozia
Oryzolejeunea (R. M. Schust.) R. M. Schust. [4] - Lejeuneaceae (28)
Otolejeunea Grolle & Tixier [10] - Lejeuneaceae (28)
Oxymitra Bisch. ex Lindenb. [3] - Oxymitraceae (5)
Pachyglossa Herzog & Grolle [4] - Lophocoleaceae (3) Top
Pachyschistochila R. M. Schust. & J. J. Engel [21] - Schistochilaceae (70)
Pallavicinia Gray [15-20] - Pallaviciniaceae (S & C-S)
Papillolejeunea Pócs = Lejeunea (88)
Paracromastigum Fulford & J. Taylor [9-11] - Lepidoziaceae (66)
Paramomitrion R. M. Schust. [monotypic] - Gymnomitriaceae (68)
Paraschistochila R. M. Schust. = Gottschea
Pedinolejeunea (Benedix ex Mizut.) P. C. Chen & P. C. Wu = Cololejeunea
Pedinophyllopsis R. M. Schust. & Inoue [monotypic] - Plagiochilaceae (69)
Pedinophyllum (Lindb.) Lindb. [4] - Plagiochilaceae (50)
Pellia Raddi [6] - Pelliaceae (5)

Peltolepis Lindb. [monotypic] - Cleveaceae (5)
Perdusenia Hässel [monotypic] - Lophocoleaceae (41)
Perilejeunea (Kachroo & R. M. Schust.) H. Rob. = Lepidolejeunea
Perssoniella Herzog [monotypic] - Perssoniellaceae (70)
Petalophyllum Nees & Gottsche ex Lehm. [5] - Petalophyllaceae (S & C-S)
Phaeolejeunea Mizut. [3] - Lejeuneaceae (24)
Phragmatocolea Grolle = Jungermannia (76)
Phragmilejeunea R. M. Schust. = Schiffneriolejeunea
Phycolepidozia R. M. Schust. [monotypic] - Phycolepidoziaceae (66)
Physotheca J. J. Engel & Gradst. [montypic] - Lophocoleaceae (86)

Phyllothallia E. A. Hodgs. [2] - Phyllothalliaceae (24)
Physantholejeunea R. M. Schust. [monotypic] - Lejeuneaceae (28)
Pictolejeunea Grolle [6] - Lejeuneaceae (92)
Pigafettoa C. Massal. [monotypic] - Lophocoleaceae (16)
Pisanoa Hässel [monotypic] - Adelanthaceae (68; 102)
Placolejeunea Herzog = Cheilolejeunea
Plagiochasma Lehm. & Lindenb. [16] - Aytoniaceae (5)
Plagiochila (Dumort.) Dumort. [ca 400+] - Plagiochilaceae (28)
Plagiochilidium Herzog [monotypic] - Plagiochilaceae (24)
Plagiochilion S. Hatt. [12] - Plagiochilaceae (28)

Plagiolejeunea Mizut. = Lopholejeunea
Platycaulis R. M. Schust. [monotypic] - Lophocoleaceae (28)
Plectocolea (Mitt.) Mitt. = Solenostoma
Pleurocladopsis R. M. Schust. [monotypic] - Schistochilaceae (70)
Pleurocladula Grolle [1-2] - Cephaloziaceae (68)
Pleurozia Dumort. [11] - Pleuroziaceae (28)
Plicanthus R. M. Schust. [3] - Scapaniaceae (91)
Pluvianthus R. M. Schust. & Schäf.-Verw. [monotypic] - Lejeuneaceae (28)
Podomitrium Mitt. [2] - Pallaviciniaceae (S & C-S)
Poeltia Grolle = Marsupella
Porella L. [50-60] - Porellaceae (28)
Potamolejeunea (Spruce) Lacout. = Trachylejeunea (79)
Prasanthus Lindb. [2] - Gymnomitriaceae (76)
Preissia Corda [monotypic] - Marchantiaceae (5)
Prionocolea (R. M. Schust.) R. M. Schust. - not validly published (96)
Prionolejeunea (Spruce) Schiffn. [10-20] - Lejeuneaceae (28)
Proskauera Heinrichs & J. J. Engel = Dinckleria (106)
Protocephalozia (Spruce) K. I. Goebel [monotypic] - Lepidoziaceae (66)
Protolophozia (R. M. Schust.) Schljakov = Lophozia
Protomarsupella R. M. Schust. = Allisoniella (68)

Protosyzygiella (Inoue) R. M. Schust. [monotypic] - Adelanthaceae (76; 102)
Pseudocephalozia R. M. Schust. [6] - Lepidoziaceae (66)
Pseudocephaloziella R. M. Schust. [monotypic] - Scapaniaceae (68)
Pseudolepicolea Fulford & J. Taylor [4] - Pseudolepicoleaceae (66)
Pseudolophocolea R. M. Schust. & J. J. Engel [monotypic] - Lophocoleaceae (69)
Pseudomarsupidium Herzog = Adelanthus (68)
Psiloclada Mitt. [1-2] - Lepidoziaceae (66)
Pteropsiella Spruce [2] - Lepidoziaceae (66)
Pteroriccia R. M. Schust. = Riccia
Ptilidium Nees [3] - Ptilidiaceae (50)

Ptychanthus Nees [2] - Lejeuneaceae (75)
Pycnolejeunea (Spruce) Schiffn. [9] - Lejeuneaceae (28)
Racemigemma S. Hatt. & Inoue = Xenochila Top
Radula Dumort. [150] - Radulaceae (28)
Reboulia Raddi [monotypic] - Aytoniaceae (5)
Rectolejeunea A. Evans [3] - Lejeuneaceae (28)
Regredicaulis Fulford = Zoopsidella
Rhaphidolejeunea Herzog = Drepanolejeunea
Rhizocaulia E. A. Hodgs. = Isotachis

Rhodoplagiochila R. M. Schust. = Plagiochila (89)
Riccardia Gray [>100] - Aneuraceae (28)
Riccia L. [150] - Ricciaceae (5)
Ricciocarpos Corda [monotypic] - Ricciaceae (5)
Riella Mont. [16] - Riellaceae (62)
Roivainenia Perss. = Syzygiella (102)
Ruizanthus R. M. Schust. [monotypic] - Balantiopsidaceae (28)
Saccobasis H. Buch = Tritomaria
Saccogyna Dumort. [monotypic] - Geocalycaceae (50) Top

Saccogynidium Grolle [7] - Geocalycaceae (28)
Sandeothallus R. M. Schust. [monotypic] - Sandeothallaceae (S & C-S)
Sauteria Nees [5] - Cleveaceae (5)
Scapania (Dumort.) Dumort. [90-100] - Scapaniaceae (76)
Scapaniella H. Buch [ ] - Scapaniaceae
Scaphophyllum Inoue [monotypic] - Jungermanniaceae (68)
Schiffneria Steph. [monotypic] - Cephaloziaceae (68)
Schiffneriolejeunea Verd. [14] - Lejeuneaceae (28)
Schistochila Dumort. [27] - Schistochilaceae (70)
Schistochilaster H. A. Miller = Gottschea

Schistochilopsis (N. Kitag.) Konst. [ ] - Scapaniaceae ( )
Schofieldia J. D. Godfrey [monotypic] - Cephaloziaceae (26)
Schusterella S. Hatt., Sharp & Mizut. = Frullania
Schusteria Kachroo = Taxilejeunea
Schusterolejeunea Grolle [monotypic] - Lejeuneaceae (28)
Sendtnerella (Spruce) R. M. Schust. = Triandrophyllum
Seppeltia Grolle [monotypic] - Pallaviciniaceae (38)
Sewardiella Kashyap [monotypic] - Petalophyllaceae (S & C-S)
Siphonolejeunea Herzog [3] - Lejeuneaceae (63)
Solenostoma Mitt. [ ] - Jungermanniaceae
Southbya Spruce [5] - Arnelliaceae (28)
Sphaerocarpos Boehm. [12] - Sphaerocarpaceae (28)
Sphaerolejeunea Herzog [monotypic] - Lejeuneaceae (28)
Sphenolobopsis R. M. Schust. & N. Kitag. [monotypic] - Scapaniaceae (76)
Sphenolobus (Lindb.) Berggr. [ ] - Scapaniaceae ( )
Spruceanthus Verd. [7] - Lejeuneaceae (28)
Sprucella Steph. [2] - Lepidoziaceae (66)
Steerea S. Hatt. & Kamim. = Frullania
Steereella Kuwah. [2] - Metzgeriaceae (28)
Steereochila Inoue [monotypic] - Plagiochilaceae (28)

Steereocolea R. M. Schust. = Balantiopsis
Steereomitrium E. O. Campb. = Haplomitrium
Stenolejeunea R. M. Schust. [3] - Lejeuneaceae (32)
Stenorrhipis Herzog = Kymatocalyx
Stephaniella J. B. Jack [6] - Stephaniellaceae (68)
Stephaniellidium S. Winkl. ex Grolle [monotypic] - Gymnomitriaceae (68)
Stephensoniella Kashyap [monotypic] - Exormothecaceae (5)
Stictolejeunea (Spruce) Schiffn. [3] - Lejeuneaceae (28)
Stolonivector J. J. Engel [monotypic] - Lophocoleaceae (13)
Stolonophora J. J. Engel & R. M. Schust. = Hepatostolonophora

Symbiezidium Trevis. [3] - Lejeuneaceae (28)
Symphyogyna Nees & Mont. [25-30] - Pallaviciniaceae (S & C-S)
Symphyogynopsis Grolle [2] - Pallaviciniaceae (S & C-S)
Syzygiella Spruce [43] - Adelanthaceae (28; 102)
Szweykowskia Gradst. & E. Reiner. [monotypic] - Plagiochilaceae (28) Top

Targionia L. [3-4] - Targioniaceae (5)
Taxilejeunea (Spruce) Schiffn. [20-30] - Lejeuneaceae (28)
Tegulifolium Hässel. = Gottschea
Telaranea Spruce ex Schiffn. [98] - Lepidoziaceae (85)
Temnoma Mitt. [10-11] - Pseudolepicoleaceae (66)

Tetracymbaliella Grolle [4] - Lophocoleaceae (30)
Tetralophozia (R. M. Schust.) Schljakov [3] - Scapaniaceae (91)
Thysananthus Lindenb. [10] - Lejeuneaceae (28)
Torticalyx Perss. = Roivainenia
Trabacellula Fulford [monotypic] - Cephaloziaceae (68)
Trachylejeunea (Spruce) Schiffn. [<10] - Lejeuneaceae (28)
Treubia K. I. Goebel [6] - Treubiaceae (73)
Triandrophyllum Fulford & Hatcher [4-5] - Herbertaceae (66)
Trichocolea Dumort. [12] - Trichocoleaceae (66)
Trichocoleopsis S. Okamura [monotypic] - Neotrichocoleaceae (S & C-S)

Trichotemnoma R. M. Schust. [monotypic] - Trichotemnomataceae (66)
Trilophozia (R. M. Schust.) Bakalin = Tritomaria (107; S & C-S)
Tritomaria Schiffn. ex Loeske [6-7] - Scapaniaceae (68)
Trocholejeunea Schiffn. [3] - Lejeuneaceae (49)
Tuyamaella S. Hatt. [6] - Lejeuneaceae (34)
Tuzibeanthus S. Hatt. [monotypic] - Lejeuneaceae (63)
Tylimanthus Mitt. [20-30] - Acrobolbaceae (28)
Vanaea (Inoue & Gradst.) Inoue & Gradst. [monotypic] - Jungermanniaceae (68)
Top
Vandiemenia Hewson [monotypic] - Vandiemeniaceae (45)

Verdoornia R. M. Schust. [monotypic] - Makinoaceae (S & C-S)
Verdoornianthus Gradst. [2] - Lejeuneaceae (29)
Vetaforma Fulford & J. Taylor [monotypic] - Vetaformataceae (66)
Vitalianthus R. M. Schust. & Giancotti [monotypic] - Lejeuneaceae (29)
Wettsteinia Schiffn. [4] - Adelanthaceae (68) Top
Wiesnerella Schiffn. [monotypic] - Wiesnerellaceae (5)
Xenocephalozia R. M. Schust. [monotypic] - Lophocoleaceae (56)
Xenochila R. M. Schust. [2] - Plagiochilaceae (25)

Xenothallus R. M. Schust. [monotypic] - Pallaviciniaceae (S & C-S)
Xylolejeunea X.-L. He & Grolle [4] - Lejeuneaceae (83)
Zoopsidella R. M. Schust. [8] - Lepidoziaceae (66) Top
Zoopsis (Hook. f. & Taylor) Gottsche, Lindenb. & Nees [8] - Lepidoziaceae (66)
DATA REFERENCES

1 - Bartholomew-Began, S. 1991. Bryophytorum Bibliotheca 41.
2 - Beckmann, K. G. & G. A. M. Scott. 1992. Journal of Bryology 17: 297.
3 - Bednarek-Ochya, J. Vana, R. Ochyra & R. I. Lewis-Smith. 2000. The Liverwort Flora of Antarctica. Polish
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6 - Bonner, C. E. B. 1963. Index Hepaticarum. Pars IV.
7 - Bonner, C. E. B. 1965. Index Hepaticarum. Pars V.
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22 - Furuki, T. & Z. Iwatsuki. 1989. Journal of the Hattori Botanical Laboratory 67: 291.
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Moss Classification at the Rank of Genus and Above
(modified from Goffinet, B., W.R Buck and A.J. Shaw, Bryophyte Biology, 2nd ed. Cambridge University
Press, 2008; Goffinet 2012: <http://www.eeb.uconn.edu/people/goffinet/Classificationmosses.html>)

PHYLUM BRYOPHYTA
SUBPHYLUM TAKAKIOPHYTINA Stech & W. Frey
CLASS TAKAKIOPSIDA Stech & W. Frey
ORDER TAKAKIALES Stech & W. Frey
FAMILY TAKAKIACEAE Stech & W. Frey: Takakia S. Hatt. & Inoue
SUBPHYLLUM SPHAGNOPHYTINA Doweld
CLASS SPHAGNOPSIDA Ochyra
ORDER SPHAGNALES Limpr.
FAMILY SPHAGNACEAE Dumort.: Sphagnum L.
FAMILY FLATBERGIACEAE A.J. Shaw: Flatbergium A.J. Shaw
FAMILY AMBUCHANANIACEAE Seppelt & H. A. Crum: Ambuchanania Seppelt & H. A.

Crum, Eosphagnum A.J. Shaw
SUBPHYLLUM ANDREAEOPHYTINA Goffinet, W. R. Buck & A. J. Shaw
CLASS ANDREAEOPSIDA J. H. Schaffn.
ORDER ANDREAEALES Limpr.
FAMILY ANDREAEACEAE Dumort.: Acroschisma (Hook.f. & Wilson) Lindl., Andreaea Hedw.
CLASS ANDREAEOBRYOPSIDA Goffinet & W. R. Buck
ORDER ANDREAEOBRYALES B. M. Murray
FAMILY ANDREAEOBRYACEAE Steere: Andreaeobryum Steere & B. M. Murray
SUBPHYLLUM BRYOPHYTINA Engler
CLASS OEDIPODIOPSIDA Goffinet & W. R. Buck
ORDER OEDIPODIALES Goffinet & W. R. Buck
FAMILY OEDIPODIACEAE Schimp.: Oedipodium Schwägr.
CLASS POLYTRICHOPSIDA Doweld
ORDER POLYTRICHALES M. Fleisch.
FAMILY POLYTRICHACEAE Schwägr.: Alophozia Cardot, Atrichopsis Cardot, Atrichum P.
Beauv., Bartramiopsis Kindb., Dawsonia R.Br., Dendroligotrichum (Müll. Hal.) Broth., Hebantia G. L.
Sm., Itatiella G. L. Sm., Lyellia R. Br., Meiotrichum (G. L. Sm.) G. L. Sm., Notoligotrichum G. L.
Sm., Oligotrichum Lam. & DC, Plagioracelopus G. L. Sm. Merr., Pogonatum P.
Beauv., Polytrichadelphus (Müll. Hal.) Mitt., Polytrichastrum G. L.
Sm., Polytrichum Hedw., Pseudatrichum Reimers, Psilopilum Brid., Steereobryon G. L. Sm.
CLASS TETRAPHIDOPSIDA Goffinet & W. R. Buck
ORDER TETRAPHIDALES M. Fleisch.
FAMILY TETRAPHIDACEAE Schimp.: Tetraphis Hedw., Tetrodontium Schwägr.
CLASS BRYOPSIDA Rothm.
SUBCLASS BUXBAUMIIDAE Doweld
ORDER BUXBAUMIALES M. Fleisch.
FAMILY BUXBAUMIACEAE Schimp.: Buxbaumia Hedw.
SUBCLASS DIPHYSCIIDAE Ochyra
ORDER DIPHYSCIALES M. Fleisch.
FAMILY DIPHYSCIACEAE M. Fleisch.: Diphyscium D. Mohr
SUBCLASS TIMMIIDAE Ochyra
ORDER TIMMIALES Ochyra
FAMILY TIMMIACEAE Schimp.: Timmia Hedw.
SUBCLASS FUNARIIDAE Ochyra
ORDER GIGASPERMALES Goffinet, Wickett, O. Werner, Ros, A.J. Shaw & C.J. Cox
FAMILY GIGASPERMACEAE Lindb.: Chamaeobryum Thér. &
Dixon, Costesia Thér., Gigaspermum Lindb., Lorentziella Müll. Hal., Oedipodiella Dixon
ORDER ENCALYPTALES Dixon
FAMILY BRYOBARTRAMIACEAE Sainsb.: Bryobartramia Sainsb.
FAMILY ENCALYPTACEAE Schimp.: Bryobrittonia R. S. Williams, Encalypta Hedw.
ORDER FUNARIALES M. Fleisch.
FAMILY FUNARIACEAE Schwägr.: Aphanorrhegma Sull., Brachymeniopsis Broth., Bryobeckettia Fife,
Clavitheca O. Werner, Ros & Goffinet, Cygnicollum Fife &
Magill, Entosthodon Schwägr., Funaria Hedw., Funariella Sérgio, Goniomitrium Hook.f. &
Wilson, Loiseaubryum Bizot, Nanomitriella E. B. Bartram, Physcomitrella Bruch &
Schimp., Physcomitrellopsis Broth. & Wager, Physcomitridium G.Roth, Physcomitrium (Brid.)
Brid., Pyramidula Brid.
FAMILY DISCELIACEAE Schimp.: Discelium Brid.
SUBCLASS DICRANIDAE Doweld
ORDER SCOULERIALES Goffinet & W. R. Buck

FAMILY SCOULERIACEAE S. P. Churchill in Funk & D. R.
Brooks: Scouleria Hook., Tridontium Hook.f.
FAMILY DRUMMONDIACEAE Goffinet: Drummondia Hook.
ORDER BRYOXIPHIALES H. A. Crum & L. E. Anderson
FAMILY BRYOXIPHIACEAE Besch.: Bryoxiphium Mitt.
ORDER GRIMMIALES M. Fleisch.
FAMILY GRIMMIACEAE Arn.: Bucklandiella Roiv., Codriophorus P.
Beauv., Dryptodon Brid., Grimmia Hedw., Leucoperichaetium Magill, Niphotrichum (Bednarek-Ochyra)
Bednarek-Ochyra & Ochyra, Racomitrium Brid., Schistidium Bruch & Schimp.
FAMILY PTYCHOMITRIACEAE Schimp.: Aligrimmia R. S. Williams, Campylostelium Bruch &

Schimp., Indusiella Broth. & Müll.
Hal., Jaffueliobryum Thér., Ptychomitriopsis Dixon, Ptychomitrium Fürnr.
FAMILY SELIGERIACEAE Schimp.: Blindia Bruch &
Schimp., Brachydontium Fürnr., Hymenolomopsis Thér., Notothamia R.Ochyra &
R.D.Seppelt, Seligeria Bruch & Schimp., Trochobryum Breidl. & Beck, Valdonia Ochyra
ORDER ARCHIDIALES Limpr.
FAMILY ARCHIDIACEAE Schimp.: Archidium Brid.
ORDER DICRANALES H. Philib. ex M. Fleisch.

FAMILY FISSIDENTACEAE Schimp.: Fissidens Hedw.
FAMILY HYPODONTIACEAE Stech & W. Frey: Hypodontium Müll. Hal.
FAMILY EUSTICHIACEAE Broth.: Eustichia (Brid.) Brid.
FAMILY DITRICHACEAE Limpr.: Astomiopsis Müll.
Hal., Bryomanginia Thér., Ceratodon Brid., Cheilothela Broth., Chrysoblastella R. S.
Williams, Cladastomum Müll. Hal., Cleistocarpidium Ochyra & Bednarek-Ochyra, Crumuscus W. R.
Buck & Snider, Cygniella H. A. Crum, Distichium Bruch &
Schimp., Ditrichopsis Broth., Ditrichum Hampe, Eccremidium Hook.f. & Wilson, Garckea Müll.
Hal., Kleioweisiopsis Dixon, ×Pleuriditrichum A. L. Andrews & F. J.
Herm., Pleuridium Rabenh., Rhamphidium Mitt., Saelania Lindb., Skottsbergia Cardot, Strombulidens W.
R. Buck, Trichodon Schimp., Tristichium Müll. Hal., Wilsoniella Müll. Hal.
FAMILY BRUCHIACEAE Schimp.: Bruchia Schwägr., Cladophascum Sim, Eobruchia W. R.
Buck, Pringleella Cardot, Trematodon Michx.
FAMILY RHACHITHECIACEAE H. Rob.: Hypnodontopsis Z. Iwats. & Nog., Jonesiobryum B. H. Allen

& Pursell, Rhachitheciopsis P. de la Varde, Rhachithecium Le Jolis, Tisserantiella P. de la
Varde, Uleastrum W. R. Buck, Zanderia Goffinet
FAMILY ERPODIACEAE Broth.: Aulacopilum Wilson, Erpodium (Brid.) Brid., Solmsiella Müll.
Hal., Venturiella Müll. Hal., Wildia Müll. Hal. & Broth.
FAMILY SCHISTOSTEGACEAE Schimp.: Schistostega D. Mohr
FAMILY VIRIDIVELLERACEAE I. G. Stone: Viridivellus I. G. Stone

FAMILY RHABDOWEISIACEAE Limpr.: Amphidium Schimp., Arctoa Bruch &
Schimp., Cynodontium Schimp., Dichodontium Schimp., Dicranoweisia Milde, Glyphomitrium Brid., Hol
odontium (Mitt.) Broth., Hymenoloma Dusén, Kiaeria I. Hagen, Oncophorus (Brid.)
Brid., Oreas Brid., Oreoweisia (Bruch & Schimp.) De Not., PseudohyophilaHilp., Rhabdoweisia Bruch &
Schimp., Symblepharis Mont., Verrucidens Cardot
FAMILY DICRANACEAE Schimp.: Anisothecium Mitt., Aongstroemia Bruch &
Schimp., Aongstroemiopsis M. Fleisch., Braunfelsia Paris, Brotherobryum M. Fleisch., Bryotestua Thér.
& P. de la Varde, Camptodontium Dusén, Campylopodium (Müll. Hal.) Besch., Chorisodontium (Mitt.)
Broth., Cnestrum I. Hagen, Cryptodicranum E. B. Bartram, Dicnemon Schwägr., Dicranella (Müll. Hal.)
Schimp., Dicranoloma (Renauld)
Renauld, Dicranum Hedw., Diobelonella Ochyra, Eucamptodon Mont., Eucamptodontopsis Broth., Holo
mitriopsis H. Rob., Holomitrium Brid., Hygrodicranum Cardot, Leptotrichella (Müll. Hal.)
Lindb., Leucoloma Brid., Macrodictyum (Broth.) E. H. Hegew., Mesotus Mitt., Mitrobryum H.
Rob., Muscoherzogia Ochyra, Orthodicranum (Bruch & Schimp.) Loeske, Paraleucobryum (Limpr.)
Loeske, Parisia Broth., Platyneuron (Cardot)
Broth., Pocsiella Bizot, Polymerodon Herzog, Pseudephemerum (Lindb.) I.
Hagen, Pseudochorisodontium (Broth.) C. H. Gao, Vitt, X. H. Fu & T. Cao, Schliephackea Müll.
Hal., Sclerodontium Schwägr., Sphaerothecium Hampe, Steyermarkiella H. Rob., Wardia Harv. &
Hook., Werneriobryum Herzog
FAMILY MICROMITRIACEAE Smyth ex Goffinet & Budke: Micromitrium Austin
FAMILY LEUCOBRYACEAE Schimp.: Atractylocarpus Mitt., Brothera Müll. Hal., Bryohumbertia P. de
la Varde & Thér., Campylopodiella Cardot, Campylopus Brid., Cladopodanthus Dozy &
Molk., Dicranodontium Bruch & Schimp., Leucobryum Hampe, Microcampylopus (Müll. Hal.)
Fleisch., Ochrobryum Mitt., Pilopogon Brid., Schistomitrium Dozy & Molk.
FAMILY CALYMPERACEAE Kindb.: Arthrocormus Dozy &
Molk., Calymperes Sw., Exodictyon Cardot, Exostratum L. T. Ellis, Leucophanes Brid., Mitthyridium H.
Rob., Octoblepharum Hedw., Syrrhopodon Schwägr.
ORDER POTTIALES M. Fleisch.
FAMILY POTTIACEAE Schimp.: Acaulon Müll. Hal., Acaulonopsis R. H. Zander &

Hedd., Algaria Hedd. & R. H. Zander, Aloina (Müll.Hal)
Kindb., Aloinella Cardot, Anoectangium Schwägr., Aschisma Lindb., Barbula Hedw., Bellibarbula P. C.
Chen, Bryoceuthospora H. A. Crum & L. E. Anderson, Bryoerythrophyllum P. C.
Chen, Calymperastrum I. G. Stone, Calyptopogon (Mitt.) Broth., Chenia R. H.
Zander, Chionoloma Dixon, Cinclidotus P. Beauv., Crossidium Jur., Crumia W. B.
Schofield, Dialytrichia (Schimp.) Limpr., Didymodon Hedw., Dolotortula R. H.
Zander, Ephemerum Schimp., Erythrophyllopsis Broth., Eucladium Bruch & Schimp., Ganguleea R. H.
Zander, Gertrudiella Broth., Globulinella Steere, Guerramontesia M.J.Cano, J.A.Jiménez, M.T.Gallego
& J.F.Jiménez Gymnostomiella M. Fleisch., Gymnostomum Nees &
Hornsch., Gyroweisia Schimp., Hennediella Paris, Hilpertia R. H. Zander, Hymenostyliella E. B.
Bartram, Hymenostylium Brid., Hyophila Brid., Hyophiladelphus (Müll. Hal.) R. H.
Zander, Indopottia A.E.D.Daniels, R.D.A.Raja & P.Daniels, Leptobarbula Schimp., Leptodontiella R. H.
Zander & E. H. Hegew., Leptodontium (Müll. Hal.) Lindb., Ludorugbya Hedd. & R.H.
Zander, Luisierella Thér. & P. de la Varde, Microbryum Schimp., Mironia R. H.
Zander, Molendoa Lindb., Nanomitriopsis Cardot, Neophoenix R. H. Zander &
During, Pachyneuropsis H.A.Mill., Phascopsis I. G. Stone, Picobryum R.H.Zander &
Hedd., Plaubelia Brid., Pleurochaete Lindb., Pottiopsis Blockeel & A. J. E. Sm., Pseudocrossidium R. S.
Williams, Pseudosymblepharis Broth., Pterygoneurum Jur., Quaesticula R. H. Zander, Reimersia P. C.
Chen, Rhexophyllum Herzog, Sagenotortula R. H. Zander, Saitobryum R. H.
Zander, Sarconeurum Bryhn, Scopelophila (Mitt.) Lindb., Splachnobryum Müll.
Hal., Stegonia Venturi, Stonea R. H. Zander, Streptocalypta Müll.
Hal., Streptopogon Mitt., Streptotrichum Herzog, Syntrichia Brid., Teniolophora W. D.
Reese, Tetracoscinodon R. Br. ter, Tetrapterum A. Jaeger, Timmiella (De Not.) Schimp., Tortella (Lindb.)
Limpr., Tortula Hedw., Trachycarpidium Broth., Trachyodontium Steere, Trichostomum Bruch, Triquetre
lla Müll. Hal., Tuerckheimia Broth., Uleobryum Broth., Vrolijkheidia Hedd. & R. H.
Zander, Weisiopsis Broth., Weissia Hedw., Weissiodicranum W. D. Reese, Willia Müll. Hal.
FAMILY PLEUROPHASCACEAE Broth.: Pleurophascum Lindb.
FAMILY SERPOTORTELLACEAE W. D. Reese & R. H. Zander: Serpotortella Dixon
FAMILY MITTENIACEAE Broth.: Mittenia Lindb.
SUBCLASS BRYIDAE Engl.
SUPERORDER BRYANAE (Engl.) Goffinet & W. R. Buck
ORDER SPLACHNALES (M. Fleisch.) Ochyra
FAMILY SPLACHNACEAE Grev. & Arn.: Aplodon R.

Br., Moseniella Broth., Splachnum Hedw., Tayloria Hook., Tetraplodon Bruch &
Schimp., Voitia Hornsch.
FAMILY MEESIACEAE Schimp.: Amblyodon P. Beauv., Leptobryum (Bruch & Schimp.)
Wilson, Meesia Hedw., Neomeesia Deguchi, Paludella Brid.
ORDER BRYALES Limpr.
FAMILY CATOSCOPIACEAE Broth.: Catoscopium Brid.
FAMILY PULCHRINODACEAE D. Quandt, N.E. Bell & Stech: Pulchrinodus B. H. Allen
FAMILY BRYACEAE Schwägr.: Acidodontium Schwägr., Anomobryum Schimp., Brachymenium Schwägr.
, Bryum Hedw., Leptostomopsis (Müll. Hal.) J. R. Spence & H. P.
Ramsay, Mniobryoides Hörmann, Ochiobryum J. R. Spence & H. P. Ramsay, Osculatia De
Not., Perssonia Bizot, Ptychostomum Hornsch., Rhodobryum (Schimp.)
Limpr., Roellobryon R.Ochyra, Rosulabryum J. R. Spence
FAMILY PHYLLODREPANIACEAE Crosby: Mniomalia Müll. Hal., Phyllodrepanium Crosby
FAMILY PSEUDODITRICHACEAE Steere & Z. Iwats.: Pseudoditrichum Steere & Z. Iwats.
FAMILY MNIACEAE Schwägr.: Cinclidium Sw., Cyrtomnium Holmen, Epipterygium Lindb., Leucolepis L
indb., Mielichhoferia Nees & Hornsch., Mnium Hedw.,Orthomnion Wilson, Plagiomnium T. J.
Kop., Pohlia Hedw., Pseudobryum (Kindb.) T. J. Kop., Pseudopohlia R. S.
Williams, Rhizomnium (Broth.) T. J. Kop., Schizymenium Harv., Synthetodontium Cardot, Trachycystis T.
J. Kop.
FAMILY LEPTOSTOMATACEAE Schwägr.: Leptostomum R. Br.
ORDER BARTRAMIALES D. Quandt, N.E. Bell & Stech

FAMILY BARTRAMIACEAE Schwägr.: Anacolia Schimp., Bartramia Hedw., Breutelia (Bruch &
Schimp.) Schimp., Conostomum Sw., Fleischerobryum Loeske, Flowersia D. G. Griffin & W. R.
Buck, Leiomela (Mitt.) Broth., Neosharpiella H. Rob. & Delgad., Philonotis Brid., Plagiopus Brid.
ORDER ORTHOTRICHALES Dixon
FAMILY ORTHOTRICHACEAE Arn.: Cardotiella
Vitt, Ceuthotheca Lewinsky, Codonoblepharon Schwägr., Desmotheca Lindb., Florschuetziella Vitt, Gro
utiella Steere, Leiomitrium Mitt., Leratia Broth. & Paris, Macrocoma (Müll. Hal.)
Grout, Macromitrium Brid., Matteria Goffinet, Orthotrichum Hedw., Pentastichella Müll.
Hal., Pleurorthotrichum Broth., Schlotheimia Brid., Sehnemobryum Lewinsky-Haapasaari &
Hedenäs, Stoneobryum D. H. Norris & H. Rob., Ulota D. Mohr, Zygodon Hook. & Taylor
ORDER HEDWIGIALES Ochyra
FAMILY HEDWIGIACEAE Schimp.: Braunia Bruch & Schimp., Bryowijkia Nog., Hedwigia P.

Beauv., Hedwigidium Bruch & Schimp., Pseudobraunia (Lesq. & James) Broth.
FAMILY HELICOPHYLLACEAE Broth.: Helicophyllum Brid.
FAMILY RHACOCARPACEAE Kindb.: Pararhacocarpus Frahm, Rhacocarpus Lindb.
ORDER RHIZOGONIALES (M. Fleisch.) Goffinet & W. R. Buck
FAMILY RHIZOGONIACEAE Broth.: Calomnion Hook.f. &
Wilson, Cryptopodium Brid., Goniobryum Lindb., Pyrrhobryum Mitt., Rhizogonium Brid.
FAMILY AULACOMNIACEAE Schimp.: Aulacomnium Schwägr., Hymenodontopsis Herzog, Mesochaete
Lindb.
FAMILY ORTHODONTIACEAE Goffinet: Hymenodon Hook.f. &
Wilson, Leptotheca Schwägr., Orthodontium Wilson, Orthodontopsis Ignatov & B. C. Tan
SUPERORDER HYPNANAE W. R. Buck, Goffinet & A. J. Shaw
ORDER HYPNODENDRALES N. E. Bell, A. E. Newton & D. Quandt
FAMILY BRAITHWAITEACEAE N. E. Bell, A. E. Newton & D. Quandt: Braithwaitea Lindb.
FAMILY RACOPILACEAE Kindb.: Powellia Mitt., Powelliopsis Zanten, Racopilum P.
Beauv., Timokoponenia Zanten.
FAMILY PTEROBRYELLACEAE (Broth.) W.R. Buck & Vitt: Cyrtopodendron M.

Fleisch., Pterobryella (Müll. Hal.) A. Jaeger, Sciadocladus Lindb. ex Kindb.
FAMILY HYPNODENDRACEAE Broth.: Bescherellia Duby, Cyrtopus (Brid.) Hook.f., Dendro-
hypnum Hampe, Franciella Thér., Hypnodendron (Müll. Hal.) Mitt., Mniodendron Lindb. ex Dozy &
Molk., Spiridens Nees, Touwiodendron N. E. Bell, A. E. Newton & D. Quandt
ORDER PTYCHOMNIALES W. R. Buck, C. J. Cox, A. J. Shaw & Goffinet
FAMILY PTYCHOMNIACEAE M. Fleisch.: Cladomnion Hook.f. & Wilson, Cladomniopsis M.

Fleisch., Dichelodontium Broth., Endotrichellopsis During, EuptychiumSchimp., Garovaglia Endl., Glyph
otheciopsis Pedersen & A. E. Newton, Glyphothecium Hampe, Hampeella Müll. Hal., Ombronesus N. E.
Bell, Pedersen & A. E. Newton, Ptychomniella (Broth.) W. R. Buck, C. J. Cox, A. J. Shaw &
Goffinet, Ptychomnion (Hook.f. & Wilson) Mitt., Tetraphidopsis Broth. & Dixon
ORDER HOOKERIALES M. Fleisch.
FAMILY HYPOPTERYGIACEAE Mitt.: Arbusculohypopterygium Stech, T. Pfeiffer & W.

Frey, Canalohypopterygium W. Frey & Schaepe, Catharomnion Hook.f. & Wilson, Cyathophorum P.
Beauv., Dendrocyathophorum Dixon, Dendrohypopterygium Kruijer, Hypopterygium Brid., Lopidium Ho
ok.f. & Wilson
FAMILY SAULOMATACEAE W. R. Buck, C. J. Cox, A. J. Shaw &

Goffinet: Ancistrodes Hampe, Sauloma (Hook.f. & Wilson) Mitt., Vesiculariopsis Broth.
FAMILY DALTONIACEAE Schimp.: Achrophyllum Vitt &
Crosby, Adelothecium Mitt., Beeveria Fife, Benitotania H. Akiyama, T. Yamag. &
Suleiman, Bryobrothera Thér., Calyptrochaeta Desv., Crosbya Vitt, Daltonia Hook. &
Taylor, Distichophyllidium M. Fleisch., Distichophyllum Dozy & Molk., Ephemeropsis K. I.
Goebel, Leskeodon Broth., Leskeodontopsis Zanten, Metadistichophyllum Nog. & Z. Iwats.
FAMILY SCHIMPEROBRYACEAE W. R. Buck, C. J. Cox, A. J. Shaw &

Goffinet: Schimperobryum Margad.
FAMILY HOOKERIACEAE Schimp.: Crossomitrium Müll. Hal., Hookeria Sm.
FAMILY LEUCOMIACEAE Broth.: Leucomium Mitt., Rhynchostegiopsis Müll. Hal., Tetrastichium (Mitt.)
Cardot
FAMILY PILOTRICHACEAE Kindb.: Actinodontium Schwägr., Amblytropis (Mitt.)
Broth., Brymela Crosby & B. H. Allen, Callicostella (Müll. Hal.)
Mitt., Callicostellopsis Broth., Cyclodictyon Mitt., Diploneuron E. B. Bartram, Helicoblepharum (Mitt.)
Broth., Hemiragis (Brid.) Besch., Hookeriopsis (Besch.) A. Jaeger, Hypnella (Müll. Hal.) A.
Jaeger, Lepidopilidium (Müll. Hal.) Broth., Lepidopilum (Brid.) Brid., Neohypnella E. B.
Bartram, Philophyllum Müll. Hal., Pilotrichidium Besch., Pilotrichum P. Beauv., Stenodesmus (Mitt.) A.
Jaeger, Stenodictyon (Mitt.) A. Jaeger, Thamniopsis (Mitt.) M. Fleisch., Trachyxiphium W. R. Buck
ORDER HYPNALES (M. Fleisch.) W. R. Buck & Vitt
FAMILY RUTENBERGIACEAE M. Fleisch.: Neorutenbergia Bizot &

Pócs, Pseudocryphaea Broth., Rutenbergia Besch.
FAMILY TRACHYLOMATACEAE W. R. Buck & Vitt: Trachyloma Brid.
FAMILY FONTINALACEAE Schimp.: Brachelyma Cardot, Dichelyma Myrin, Fontinalis Hedw.
FAMILY Climaciaceae Kindb.: Climacium F. Weber & D. Mohr, Pleuroziopsis E. Britton
FAMILY AMBLYSTEGIACEAE G.
Roth: Amblystegium Schimp., Anacamptodon Brid., Bryostreimannia Ochyra, Campyliadelphus (Kindb.)
R.S. Chopra, Campylium (Sull.) Mitt., Conardia H. Rob., Cratoneuron (Sull.)
Spruce, Cratoneuropsis (Broth.) M. Fleisch., Drepanocladus (Müll. Hal.) G.
Roth, Gradsteinia Ochyra, HygroamblystegiumLoeske, Hygrohypnella Ignatov &
Ignatova, Hygrohypnum Lindb., Hypnobartlettia Ochyra, Koponenia Ochyra, Leptodictyum (Schimp.)
Warnst., Limbella (Müll. Hal.) Müll.
Hal., Limprichtia Loeske, Ochyraea Váňa, Palustriella Ochyra, Pictus C. C.
Towns., Pseudoamblystegium Vanderpoorten & Hedenäs, Pseudo-calliergon (Limpr.)
Loeske, Pseudocampylium Vanderpoorten &
Hedenäs, Pseudohygrohypnum Kanda, Sanionia Loeske, Sasaokaea Broth., Sciaromiella Ochyra, Sciaro
miopsis Broth., Scorpidium(Schimp.) Limpr., Sinocalliergon Sakurai, Serpoleskea (Limpr.)
Loeske, Vittia Ochyra
FAMILY CALLIERGONACEAE Vanderpoorten, Hedenäs, C. J. Cox & A. J. Shaw: Calliergon (Sull.)

Kindb., Hamatocaulis Hedenäs, Loeskypnum H. K. G. Paul, Straminergon Hedenäs, Warnstorfia Loeske
FAMILY Helodiaceae Ochyra: Actinothuidium (Besch.) Broth., Bryochenea C. H. Gao & K. C.

Chang, Helodium Warnst.
FAMILY LESKEACEAE Schimp.: Claopodium (Lesq. & James) Renauld & Cardot, Fabronidium Müll.

Hal., Haplocladium (Müll. Hal.) Müll.
Hal., Hylocomiopsis Cardot, Leptocladium Broth., Leptopterigynandrum Müll. Hal., Lescuraea Bruch &
Schimp., Leskea Hedw., Leskeadelphus Herzog, Leskeella (Limpr.)
Loeske, Lindbergia Kindb., Mamillariella Laz., Orthoamblystegium Dixon & Sakurai, Platylomella A. L.
Andrews, Pseudoleskea Bruch &
Schimp., Pseudoleskeella Kindb., Pseudoleskeopsis Broth., Ptychodium Schimp., Rigodiadelphus Dixon,
Rozea Besch., Schwetschkea Müll. Hal.
FAMILY THUIDIACEAE Schimp.: Abietinella Müll. Hal., Boulaya Cardot, Cyrto-hypnum (Hampe)
Hampe & Lorentz, Fauriella Besch., Pelekium Mitt., Rauiella Reimers, Thuidiopsis (Broth.) M.
Fleisch., Thuidium Bruch & Schimp.
FAMILY REGMATODONTACEAE Broth.: Regmatodon Brid., Yunnanobryon Shevock, Ochyra, S.He &

D.G.Long
FAMILY STEREOPHYLLACEAE W. R. Buck &
Ireland: Catagoniopsis Broth., Entodontopsis Broth., Eulacophyllum W. R. Buck &
Ireland, Juratzkaea Lorentz, Pilosium(Müll. Hal.) M. Fleisch., Sciuroleskea Broth., Stenocarpidium Müll.
Hal., Stereophyllum Mitt.
FAMILY BRACHYTHECIACEAE G. Roth: Aerobryum Dozy & Molk., Aerolindigia M.

Menzel, Brachytheciastrum Ignatov &
Huttunen, Brachythecium Schimp., BryhniaKaurin, Bryoandersonia H.
Rob., Cirriphyllum Grout, Clasmatodon Hook.f. & Wilson, Donrichardsia H. A. Crum & L. E.
Anderson, Eriodon Mont., Eurhynchiadelphus Ignatov & Huttunen, Eurhynchiastrum Ignatov &
Huttunen, Eurhynchiella M. Fleisch., Eurhynchium Bruch &
Schimp., Flabellidium Herzog, Frahmiella Ignatov, Y.F.Wang & Vanderpoorten, Hedenaesia Huttunen &
Ignatov, Hedenasiastrum Ignatov & Vanderpoorten, Helicodontium Schwägr., Homalotheciella (Cardot)
Broth., Homalothecium Schimp., Juratzkaeella W. R.
Buck, Kindbergia Ochyra, Lindigia Hampe, Mandoniella Herzog, Meteoridium (Müll. Hal.)
Manuel, Microeurhynchium Ignatov &
Vanderpoorten , Myuroclada Besch., Nobregaea Hedenäs, Okamuraea Broth., Oxyrrhynchium (Schimp.)
Warnst., Palamocladium Müll. Hal., Plasteurhynchium Broth., Platyhypnidium M.
Fleisch., Pseudopleuropus Takaki, Pseudorhynchostegiella Ignatov &
Vanderpoorten , Pseudoscleropodium (Limpr.) M. Fleisch., Remyella Müll.
Hal., Rhynchostegiella (Schimp.) Limpr., Rhynchostegium Bruch & Schimp., Sainthelenia Ignatov & M.
Wigginton, Schimperella Thér., Sciuro-hypnum (Hampe) Hampe, Scleropodium Bruch &
Schimp., Scorpiurium Schimp., Squamidium (Müll. Hal.) Broth., Stenocarpidiopsis M.
Fleisch., Tomenthypnum Loeske, Zelometeorium Manuel
FAMILY METEORIACEAE Kindb.: Aerobryidium M. Fleisch., Aerobryopsis M. Fleisch., Barbella M.
Fleisch., Barbellopsis Broth., Chrysocladium M. Fleisch., Cryptopapillaria M.
Menzel, Diaphanodon Renauld & Cardot, Duthiella Renauld, Floribundaria M.
Fleisch., Lepyrodontopsis Broth., Meteoriopsis Broth., Meteorium (Brid.) Dozy &
Molk., Neodicladiella W. R. Buck, Neonoguchia S. H. Lin, Pseudospiridentopsis (Broth.) M.
Fleisch., Pseudotrachypus P. de la Varde & Thér., Sinskea W. R. Buck, Toloxis W. R.
Buck, Trachycladiella (M. Fleisch.) M. Menzel & W. Schultze-Motel, Trachypodopsis M.
Fleisch., Trachypus Reinw. & Hornsch.
FAMILY MYRINIACEAE Schimp.: Austinia Müll. Hal., Macgregorella E. B.
Bartram, Merrilliobryum Broth., Myrinia Schimp., Nematocladia W. R. Buck
FAMILY FABRONIACEAE Schimp.: Dimerodontium Mitt., Fabronia Raddi, Ischyrodon Müll.
Hal., Levierella Müll. Hal., Rhizofabronia (Broth.) M. Fleisch.
FAMILY HYPNACEAE Schimp.: Acritodon H. Rob., Andoa Ochyra, Austrohondaella Z.Iwats.,
H.P.Ramsay & Fife, Bardunovia Ignatov & Ochyra, Breidleria Loeske, Bryocrumia L. E.
Anderson, Buckiella Ireland, Callicladium H. A. Crum, Calliergonella Loeske, Campylophyllopsis W. R.
Buck, Campylophyllum (Schimp.) M. Fleisch., Caribaeohypnum Ando & Higuchi, Chryso-
hypnum (Hampe) Hampe, Crepidophyllum Herzog, Ctenidiadelphus M.
Fleisch., Cyathothecium Dixon, Dacryophyllum Ireland, Ectropotheciella M.
Fleisch., Ectropotheciopsis (Broth.) M. Fleisch., Ectropothecium Mitt., Elharveya H. A.
Crum, Elmeriobryum Broth., Entodontella M. Fleisch., Eurohypnum Ando, Foreauella Dixon & P. de la
Varde, Gammiella Broth., Giraldiella Müll.
Hal., Gollania Broth., Hageniella Broth., Herzogiella Broth., Homomallium (Schimp.)
Loeske, Hondaella Dixon & Sakurai, Horridohypnum W. R. Buck, Hyocomium Bruch &
Schimp., Hypnum Hedw., Irelandia W. R. Buck, Isopterygiopsis Z.
Iwats., Leiodontium Broth., Leptoischyrodon Dixon, Macrothamniella M. Fleisch., Mahua W. R.
Buck, Microctenidium M. Fleisch., Mittenothamnium Henn., Nanothecium Dixon & P. de la
Varde, Orthothecium Bruch & Schimp., Phyllodon Bruch &
Schimp., Plagiotheciopsis Broth., Platydictya Berk., Platygyriella Cardot, Podperaea Z. Iwats. &
Glime, Pseudohypnella (M. Fleisch.) Broth., Pseudotaxiphyllum Z. Iwats., Ptilium De
Not., Pylaisia Schimp., Rhacopilopsis Renauld & Cardot, Rhizohypnella M.
Fleisch., Sclerohypnum Dixon, Stenotheciopsis Broth., Stereodon (Brid.) Mitt., Stereodontopsis R. S.
Williams, Syringothecium Mitt., Taxiphyllopsis Higuchi & Deguchi, Taxiphyllum M.
Fleisch., Vesicularia (Müll. Hal.) Müll. Hal., Wijkiella Bizot & Lewinsky
FAMILY CATAGONIACEAE W. R. Buck & Ireland: Catagonium Broth.
FAMILY PTERIGYNANDRACEAE Schimp.: Habrodon Schimp., Heterocladium Bruch &
Schimp., Iwatsukiella W. R. Buck & H. A. Crum, Myurella Bruch &
Schimp., Pterigynandrum Hedw., Trachyphyllum A. Gepp
FAMILY HYLOCOMIACEAE M. Fleisch.: Ctenidium Mitt., Hylocomiastrum Broth., Hylocomium Bruch
& Schimp., Leptocladiella M.
Fleisch., Leptohymenium Schwägr., Loeskeobryum Broth., Macrothamnium M. Fleisch., Meteoriella S.
Okamura, Neodolichomitra Nog., Orontobryum M. Fleisch., Pleurozium Mitt., Puiggariopsis M.
Menzel, Rhytidiadelphus (Limpr.) Warnst., Rhytidiopsis Broth., Schofieldiella W. R. Buck
FAMILY RHYTIDIACEAE Broth.: Rhytidium (Sull.) Kindb.
FAMILY SYMPHYODONTACEAE M. Fleisch.: Chaetomitriopsis M. Fleisch., Chaetomitrium Dozy &
Molk., Dimorphocladon Dixon, Symphyodon Mont., Trachythecium M. Fleisch., Unclejackia Ignatov, T.
Kop. & D. Norris
FAMILY PLAGIOTHECIACEAE M. Fleisch.: Plagiothecium Bruch & Schimp., Struckia Müll. Hal.
FAMILY ENTODONTACEAE Kindb.: Entodon Müll.
Hal., Erythrodontium Hampe, Mesonodon Hampe, Pylaisiobryum Broth.
FAMILY PYLAISIADELPHACEAE Goffinet & W. R. Buck: Aptychella (Broth.) Herzog, Brotherella M.

Fleisch., Clastobryopsis M. Fleisch., Clastobryum Dozy & Molk., Heterophyllium (Schimp.)
Kindb., Isocladiella Dixon , Isopterygium Mitt., Mastopoma Cardot, Platygyrium Bruch &
Schimp., Pterogonidium Broth., Pseudotrismegistia H. Akiyama &
Tsubota, Pylaisiadelpha Cardot, Taxitheliella Dixon, Taxithelium Mitt., Trismegistia (Müll. Hal.) Müll.
Hal., Wijkia H. A. Crum, Yakushimabryum H.Akiyama, Y.Chang, Yamagushi & B.C.Tan.
FAMILY SEMATOPHYLLACEAE Broth.: Acanthorrhynchium M.
Fleisch., Acroporium Mitt., Allioniellopsis Ochyra, Aptychopsis (Broth.) M.
Fleisch., Chionostomum Müll. Hal., Clastobryella M. Fleisch., Clastobryophilum M.
Fleisch., Colobodontium Herzog, Donnellia Austin, Hydropogon Brid., Hydropogonella Cardot, Macrohy
menium Müll. Hal., Meiotheciella B. C. Tan, W. B. Schofield & H. P.
Ramsay, Meiothecium Mitt., Papillidiopsis (Broth.) W. R. Buck & B. C. Tan, Paranapiacabaea W. R.
Buck & Vital,Potamium Mitt., Pterogoniopsis Müll. Hal., Piloecium (Müll. Hal.) Broth., Radulina W. R.
Buck & B. C. Tan, Rhaphidostichum M. Fleisch., Schraderella Müll.
Hal., Schroeterella Herzog, Sematophyllum Mitt., Timotimius W. R.
Buck, Trichosteleum Mitt., Trolliella Herzog, Warburgiella Müll. Hal.
FAMILY CRYPHAEACEAE Schimp.: Cryphaea D. Mohr, Cryphaeophilum M.
Fleisch., Cryphidium (Mitt.) A. Jaeger, Cyptodon (Broth.) M.
Fleisch., Cyptodontopsis Dixon, Dendroalsia E. Britton, Dendrocryphaea Broth., Dendropogonella E.
Britton, Pilotrichopsis Besch., Schoenobryum Dozy & Molk., Sphaerotheciella M. Fleisch.
FAMILY PRIONODONTACEAE Broth.: Prionodon Müll. Hal.
FAMILY LEUCODONTACEAE Schimp.: Antitrichia Brid., Dozya Sande Lac., Eoleucodon H. A. Mill. &

H. Whittier, Leucodon Schwägr., Nogopterium Crosby & W.R.buck, Pterogoniadelphus M.
Fleischer, Scabridens E. B. Bartram
FAMILY PTEROBRYACEAE Kindb.: Calyptothecium Mitt., Cryptogonium (Müll. Hal.)
Hampe, Henicodium (Müll. Hal.) Kindb., Hildebrandtiella Müll. Hal., Horikawaea Nog., Jaegerina Müll.
Hal., Micralsopsis W. R. Buck, Muellerobryum M. Fleisch., Neolindbergia M.
Fleisch., Orthorrhynchidium Renauld &
Cardot, Orthostichidium Dusén, Orthostichopsis Broth., Osterwaldiella Broth., Penzigiella M.
Fleisch., Pireella Cardot, Pseudopterobryum Broth., Pterobryidium Broth. &
Watts, Pterobryon Hornsch., Pterobryopsis M. Fleisch., Renauldia Müll.
Hal., Rhabdodontium Broth., Spiridentopsis Broth., Symphysodon Dozy & Molk., Symphysodontella M.
Fleisch.
FAMILY PHYLLOGONIACEAE Kindb.: Phyllogonium Brid.
FAMILY ORTHORRHYNCHIACEAE S. H. Lin: Orthorrhynchium Reichardt
FAMILY LEPYRODONTACEAE Broth.: Lepyrodon Hampe
FAMILY NECKERACEAE Schimp.: Alleniella S.Olsson, Enroth & D. Quandt,Baldwiniella M.
Fleisch., Bryolawtonia D. H. Norris & Enroth, Caduciella Enroth, CirculifoliumS. Olsson, Enroth &
D.Quandt, Crassiphyllum Ochyra, Cryptoleptodon Renauld &
Cardot, Curvicladium Enroth, Dixonia Horik. & Ando, Dolichomitra Broth., Echinodiopsis S.Olsson,
Enroth & D. Quandt, Exsertotheca S.Olsson, Enroth & D.
Quandt,Handeliobryum Broth., Himantocladium (Mitt.) M. Fleisch., Homalia (Brid.) Bruch &
Schimp., Homaliodendron M. Fleisch., Hydrocryphaea Dixon, Isodrepanium (Mitt.) E.
Britton, Metaneckera Steere, Neckera Hedw., Neckeropsis Reichardt, Neomacounia Ireland, Noguchioden
dron Ninh & Pócs, Orthostichella Müll. Hal., Pendulothecium Enroth & S. He, Pinnatella M.
Fleisch., Porotrichodendron M. Fleisch., Porotrichopsis Broth. & Herzog, Porotrichum (Brid.)
Hampe, Shevockia Enroth & Ji, Thamnobryum Nieuwl., Thamnomalia S.Olsson, Enroth & D.
Quandt,Touwia Ochyra
FAMILY ECHINODIACEAE Broth.: Echinodium Jur.
FAMILY LEPTODONTACEAE Schimp.: Alsia Sull., Forsstroemia Lindb., Leptodon D.
Mohr, Taiwanobryum Nog.
FAMILY LEMBOPHYLLACEAE Broth.: Acrocladium Mitt., Bestia Broth., Camptochaete Reichardt, D
olichomitriopsis S. Okamura, Fallaciella H. A. Crum, Fifea H. A.
Crum, Isothecium Brid., Lembophyllum Lindb., Looseria (Thér.) D. Quandt, S. Huttunen, Tangney &
Stech, Neobarbella Nog., Pilotrichella (Müll. Hal.) Besch., Rigodium Schwägr., Tripterocladium (Müll.
Hal.) A. Jaeger, Weymouthia Broth.
FAMILY MYURIACEAE M.
Fleisch.: Eumyurium Nog., Myurium Schimp., Oedicladium Mitt., Palisadula Toyama
FAMILY ANOMODONTACEAE Kindb.: Anomodon Hook. & Taylor, Bryonorrisia L. R. Stark & W. R.
Buck, Chileobryon Enroth, Curviramea H. A. Crum, HaplohymeniumDozy &
Molk., Herpetineuron (Müll. Hal.) Cardot, Schwetschkeopsis Broth.
FAMILY Miyabeaceae Enroth, S. Olsson, Buchbender, Hedenäs, Huttunen & D.

Quandt: Bissetia Broth., Homaliadelphus Dixon & P. de la Varde, Miyabea Broth.
FAMILY THELIACEAE M. Fleisch.: Thelia Sull.
FAMILY MICROTHECIELLACEAE H. A. Mill. & A. J. Harr.: Microtheciella Dixon
FAMILY SORAPILLACEAE M. Fleisch.: Sorapilla Spruce & Mitt.
Hornwort Classification at the Rank of Genus and Above

PHYLUM (DIVISION) ANTHOCEROTOPHYTA Stotl. & Crand.-Stotl. Bryologist 80: 425. 1977.
CLASS LEIOSPOROCEROTOPSIDA Stotl. & Crand.-Stotl., Bryologist 108: 24. 2005
ORDER LEIOSPOROCEROTALES Hässel, J. Hatt. Bot. Lab. 64: 82. 1988.
FAMILY LEIOSPOROCEROTACEAE Hässel, J. Bryol. 14: 255. 1986.
Leiosporoceros Hässel, J. Bryol. 14: 255. 1986.

CLASS ANTHOCEROTOPSIDA Jancz. ex Stotl. & Crand.-Stotl., Bryologist 108: 24. 2005.
ORDER ANTHOCEROTALES Limpricht in Cohn, Krypt. Fl. von Schlesien: 239, 345. Breslau. 1877.
FAMILY ANTHOCEROTACEAE Dumort., Analys. Fam. Pl. 68-69. Tournay. 1829.
Anthoceros L., Sp. Plt.: 1139. Stockholm. 1753.
Folioceros D. C. Bharadwaj, Geophytology 1: 9. 1971.
Sphaerosporoceros Hässel, J. Hattori Bot. Lab. 64: 79. 1988.
ORDER NOTOTHYLADALES Hyvönen & Piippo, J. Hatt. Bot. Lab. 74: 117. 1993. "Notothylales".
FAMILY NOTOTHYLADACEAE (Milde) Müll. Frib. ex Prosk., Phytomorphology 10: 10. 1960.
SUBFAMILY NOTOTHYLADOIDEAE Grolle, J. Bryol. 7: 215. 1972.
Notothyas Sull. ex A. Gray, Amer. J. Sci. Arts, ser. 2, 1: 74. 1846.
SUBFAMILY PHAEOCEROTOIDEAE Hässel, J. Hatt. Bot. Lab. 64: 81. 1988.
Hattorioceros (Hasegawa) Hasegawa, J. Hattori Bot. Lab. 76: 32. 1994.
Mesoceros Piippo, Acta Bot. Fenn. 148: 30. 1993.
Phaeoceros Prosk., Bull. Torrey Bot. Cl. 78: 346. 1951.
Phymatoceros Stotler, W. T. Doyle & Crand.-Stotl., Phytologia 87: 113. 2005.

FAMILY DENDROCEROTACEAE (Milde) Hässel, J. Hatt. Bot. Lab. 64: 82. 1988.
SUBFAMILY DENDROCEROTOIDEAE R. M. Schust., Phytologia 63: 195, 200.
Dendroceros Nees in Gottsche, Lindenberg & Nees, Syn. Hep.: 579. 1846.
SUBFAMILY MEGACEROTOIDEAE Stotl. & Crand.-Stotl., Bryologist 108: 24. 2005.
Megaceros Campb., Ann. Bot. (London) 21: 484. 1907.
Nothoceros (R. M. Schust.) Hasegawa, J. Hatt. Bot. Lab. 76: 32. 1994. "Notoceros"
SOURCE:
Stotler, R. E. & B. Crandall-Stotler. 2005. A revised classification of the Anthocerotophyta and a checklist of
the hornworts of North America, north of Mexico. Bryologist 108: 16-26.

Anthoceros fusiformis Austin
Photo taken in Carbondale [not on natural substrate] by Li Zhang, July, 2004.
Collected 29 June, 2004 by David Wagner (Wagner m1386) along the South Santiam River, Oregon.
Growing on soil next to a seepy area. (voucher in ABSH)
Folioceros fuciformis (Mont.) Bharadw.

CHINA. Hong Kong, February 2002
Nothothylas javanica (Sande Lac.) Gottsche
Photo taken by Li Zhang.
CHINA. Macau, Coloane Is., on soil in green belt, 30 m., October 12, 2006. (voucher in SZG)
Phaeoceros carolinianus (Michx.) Prosk.
Collected 29 June, 2004 by David Wagner (Wagner m1389) along the South Santiam River, Oregon.
Growing in a constantly flowing sheet of water, on rock, next to a small waterfall. [voucher ABSH].
Photo taken in Carbondale [not on natural substrate] by Li Zhang, July, 2004.

Dehiscent capsule; Note yellow spore color
Photo taken by Li Zhang, July, 2004.
Dehiscent capsule; Note yellow spore color
Photo taken by Li Zhang, July, 2004.
Phaeoceros cf. carolinianus (Michx.) Prosk.
Found on soil along a path in sparse forest, June 3, 2005 in Southern Illinois at Jackson Co., Little Grand
Canyon by Li Zhang; photo taken by Li Zhang
Dendroceros sp.
Malaysia. Penang. Cameron Highlands. Brinchang, Mt. Irau. N04°28', E101°23'. 1950 m. On tree trunk in
forest. Li Zhang 5729, July 29, 2007. (voucher in SZG)

Bryophytes - Gambar

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Bryophytes - Gambar

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Ann. Missouri Bot. Gard. 81: 451-483.)

based upon phylogenomic evidence.

SUBCLASS TREUBIIDAE Stotler & Crand.-Stotl., Taxon 57: 290 (2008).

Treubiaceae Verd., Man. Bryol. 427 (1932). Apotreubia S. Hatt. & Mizut., Treubia K. I. Goebel

SUBCLASS HAPLOMITRIIDAE Stotler & Crand.-Stotl., Taxon 57: 290 (2008).

ORDER: Calobryales Hamlin, Rec. Domin. Mus. 7: 315 (1972).

Blasiaceae H. Klinggr., Höh. Crypt. Preuss.: 14 (1858). Blasia L., Cavicularia Steph.

SUBCLASS: MARCHANTIIDAE Engl. [Unterklasse "Marchantiales"] in: A. Engler & K. Prantl, Nat.

ORDER: SPHAEROCARPALES Cavers, New Phytol. 9: 81 (1910).

Sphaerocarpaceae Heeg, Verh. K. K. Zool.-Bot. Ges. Wien 41: 573

Riellaceae Engl., Syllabus, Grosse Ausgabe: 45 (1892). Riella Mont.

ORDER: NEOHODGSONIALES D. G. Long, Edinburgh J. Bot. 63: 258 (2006).

Neohodgsoniaceae D. G. Long, Edinburgh J. Bot. 63: 258 (2006). Neohodgsonia Perss.

ORDER: LUNULARIALES D. G. Long, Edinburgh J. Bot. 63: 259 (2006).

Lunulariaceae H. Klinggr., Höh. Crypt. Preuss.: 9 (1858). Lunularia Adans.

Marchantiaceae Lindl., Nat. Syst. Bot. (ed. 2): 412 (1836). Bucegia Radian, Marchantia L., Preissia Corda

Aytoniaceae Cavers, New Phytol. 10: 42 (1911). Asterella P. Beauv., Cryptomitrium Austin ex

Cleveaceae Cavers, New Phytol. 10: 42 (1911). Athalamia Falconer, Sauteria Nees, Peltolepis Lindb.

Monosoleniaceae Inoue, Bull. Natl. Sci. Mus. Tokyo 9: 117 (1966). Monosolenium Griff.

Conocephalaceae Müll. Frib. ex Grolle, J. Bryol. 7: 207 (1972). Conocephalum Hill nom. cons.

Cyathodiaceae Stotler & Crand.-Stotl. in A. J. Shaw & B. Goffinet, Bryoph. Biol.: 63

Exormothecaceae Müll. Frib. ex Grolle, J. Bryol. 7: 208

Corsiniaceae Engl., Syllabus, Grosse Ausgabe: 44 (1892). Corsinia Raddi, Cronisia Berk.

Monocarpaceae D. J. Carr ex Schelpe, J. S. African Bot. 35: 110 (1969). Monocarpus D. J. Carr

Oxymitraceae Müll. Frib. ex Grolle, J. Bryol. 7: 215 (1972). Oxymitra Bisch. ex Lindenb.

Ricciaceae Rchb., Bot. Damen: 255 (1828). Riccia L., Ricciocarpos Corda

Wiesnerellaceae Inoue, Ill. Jap. Hepat. 2: 192 (1976). Wiesnerella Schiffn.

Targioniaceae Dumort., Anal. Fam. Pl.: 68, 70 (1829). Targionia L.

Dumortieraceae D. G. Long, Edinburgh J. Bot. 63: 260 (2006). Dumortiera Nees

CLASS: JUNGERMANNIOPSIDA Stotler & Crand.-Stotl., Bryologist 80: 425 (1977).

Pelliaceae H. Klinggr, Höh. Crypt. Preuss.: 13 (1858). Noteroclada Taylor ex Hook. & Wilson, Pellia Raddi

Calyculariaceae He-Nygrén, Juslén, Ahonen, Glenny & Piippo, Cladistics 22: 27 (2006). Calycularia Mitt.

Makinoaceae Nakai, Chosakuronbun Mokuroku [Ord. Fam. Trib. Nov.]: 201 (1943). Makinoa Miyake

SUBORDER: Fossombroniineae R.M.Schust. ex Stotler & Crand.-Stotl. in A. J. Shaw & B. Goffinet, Bryoph.

Petalophyllaceae Stotler & Crand.-Stotl., Novon 12: 335 (2002). Petalophyllum Nees & Gottsche ex

Fossombroniaceae Hazsl. nom. cons., Magyar Bir, Moh.-Fl.: 20, 36 (1885). Fossombronia Raddi

ORDER: PALLAVICINIALES W. Frey & M. Stech, Nova Hedwigia 81: 64 (2005).

SUBORDER: Phyllothalliineae R. M. Schust., Trans. Brit. Bryol. Soc. 5: 283 (1967).

SUBORDER: Pallaviciniineae R. M. Schust., Phytologia 56: 65 (1984).

Sandeothallaceae R. M. Schust., New Man. Bryol.: 951 (1984). Sandeothallus R. M. Schust.

Hymenophytaceae R. M. Schust., J. Hattori Bot. Lab. 26: 296 (1963). Hymenophyton Dumort.

Pallaviciniaceae Mig., Krypt.-Fl. Deutschl., Moose: 423

SUBCLASS METZGERIIDAE Barthol.-Began, Phytologia 69: 465. (1990[1991]).

ORDER: METZGERIALES Chalaud, Ann. Bryol. 3: 41 (1930).

Metzgeriaceae H. Klinggr., Höh. Crypt. Preuss.: 10 (1858). Metzgeria Raddi

Aneuraceae H. Klinggr., Höh. Crypt. Preuss.: 11 (1858). Aneura Dumort.

SUBCLASS JUNGERMANNIIDAE Engl. [Unterklasse "Jungermanniales"] in A. Engler & K. Prantl, Nat.

SUBORDER: Porellineae R. M. Schust., J. Hattori Bot. Lab. 26: 229 (1963).

Porellaceae Cavers nom. cons., New Phytol. 9: 292 (1910). Ascidiota C. Massal., Porella L.

Goebeliellaceae Verd., Man. Bryol.: 425 (1932). Goebeliella Steph.

Lepidolaenaceae Nakai, Chosakuronbun Mokuroku [Ord. Fam. Trib. Nov.]: 200

SUBORDER: Radulineae R. M. Schust., J. Hattori Bot. Lab. 26: 229 (1963).

SUBORDER: Jubulineae Müll. Frib., Lebermoose 1: 403 (1909).

Frullaniaceae Lorch in G. Lindau, Krypt.-Fl. Anf. 6: 174 (1914). Frullania Raddi [including Amphijubula R. M.

Jubulaceae H. Klinggr., Höh. Crypt. Preuss.: 40 (1858). Jubula Dumort. nom. cons., Nipponolejeunea S.

Lejeuneaceae Cavers, New Phytol. 9: 291 (1910). Acanthocoleus R. M. Schust., Acantholejeunea (R. M.

Ptilidiaceae H. Klinggr., Höh. Crypt. Preuss.: 37 (1858). Ptilidium Nees

Neotrichocoleaceae Inoue, Ill. Jap. Hepat. 1: 176 (1974). Neotrichocolea S. Hatt., Trichocoleopsis S.

Herzogianthaceae Stotler & Crand.-Stotl. in B. Goffinet & A. J. Shaw, Bryoph. Biol. ed. 2: 54

ORDER JUNGERMANNIALES H. Klinggr., Höh. Crypt. Preuss.: 16 (1858).

SUBORDER: Perssoniellineae R. M. Schust., J. Hattori Bot. Lab. 26: 229-230 (1963).