THE GENUS REBUTIA

1895—1981

REBUTIA MUSCULA

No. 2 of a series of illustrated guide books by:-

Brian Fearn and Leslie Pearcy

Published by Abbey Brook Cactus Nursery

 Published by: Abbey Brook Cactus Nursery,
Old Hackney Lane,
Matlock, Derbyshire.

Copyright ©ABCN 1981

Illustrations: All illustrations are by Brian Fearn

Printed in Great Britain by:

Higham Press Ltd., Shirland, Derby., DE5 6BP
 THE GENUS REBUTIA
1895-1981

An up to date check list of the genus Rebutia

and related groups including Aylostera,
Mediolobivia and Digitorebutia together with
photographs and descriptions of 20 of the
more notable species.

No. 2 of a series of illustrated guide books by:-

Brian Fearn and Leslie Pearcy

Published by ABBEY BROOK CACTUS NURSERY

A 1
 Contents
Preface. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 3
Introduction . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 4
Nomenclature and Classification. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 5
Analytical key to the Rebutia Sections . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 7
Flowers and floral structure . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 8
Geographical Distribution . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 13
Distribution maps and locational data. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 15
Cultivation . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 23
Pests and Diseases. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 26
Descriptions - arranged in alphabetical order 27
Check list of Genera, Subgenera and Sections. . . . . . . . . . . . . . . . . . . . . . . . . 52
Check list of Aylostera Speg.. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 53
Check list of Digitorebutia Frič & Kreuzgr. ex Buin.. . . . . . . . . . . . . . . . . . . . 54
Check list of Mediolobivia Backbg.. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 55
Check list of Genus Rebutia including specific, varietal and name
combinations which have been used in connection with the genus . . . . . . 59
Friedrich Ritter field numbers . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 71
Alfred Lau field numbers. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 73
Walter Rausch field numbers. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 74
Karel Knize field numbers. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 77
Bibliography. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 78
Acknowledgements. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79
Glossary. . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . . 79

2
 Preface
‘The Genus Rebutia 1895-1981’ is the second in a planned series of publications
by Abbey Brook Cactus Nursery. The first booklet ‘50 Choice Mammillarias’ has
been well received and the 1st edition has sold well and is now out of print. Two
years have passed since its introduction and we apologise for the delay in the
production of the 2nd booklet. The original intention was to produce a new booklet
each year. In the meantime another booklet ‘Lithops’ by Brian Fearn has been
published by the National Cactus and Succulent Society and so the new booklet is
really the 3rd that has been produced by Abbey Brook. We are thus still on target.
There seems to be no end to the publication of new general books on Cacti and
other Succulent plants but our intention of producing inexpensive booklets with
detailed information for the serious collector is as relevant now as it was when
booklet No. 1 was published.
Brian Fearn and Leslie Pearcy
Matlock 1981

REBUTIA FLAVISTYLA
Fig. 1 A superb species discovered by Friedrich Ritter and described in 1978
(FR756). Beautiful light orange flowers freely produced.

3
 Introduction
The genus Rebutia is one of the most widely cultivated groups of plants which
belong to the family Cactaceae. Their ease of cultivation and free flowering nature
endears them to all cactophiles. They are plants of the Andean mountain chain in
South America occurring in northern Argentina and Bolivia. One of the reasons for
their popularity is their ease of cultivation and adaptability to neglect. They do not
exactly thrive on neglect, but nevertheless they will still produce their beautiful
flowers in the spring, provided cool, dry conditions have been given during the
winter months. As these are plants from the high Andes, they have the added ability
to survive in an unheated greenhouse or porch when winter temperatures could fall
to well below 0°C (32°F).
Nearly all the species are small growing plants, rarely needing more than a 3½''
pot, with a range of body form from globular to short cylindrical stems. The spination
is very varied and often very colourful. Some plants have snow white spines, others
golden yellow. The flowers are often very large for the size of the plant. In the
majority of cases the flower colours are vivid oranges, reds, yellows and purple but
occasionally bicoloured flowers are produced. There are also a few white flowered
species. These plants are of great horticultural value and no collection of cacti is
complete without some representatives of this genus.

REBUTIA FUSCA
Fig. 2 Described by Friednch Ritter in 1977 (FR940). It produces beautiful dark red
flowers and is probably closely related to R. spegazziniana Backbg.
4
 Nomenclature and Classification
It is doubtful if two people will ever agree on the taxonomy of the genus Rebutia
and as a result the nomenclature is a veritable minefield. This genus has suffered
more than most from the over-enthusiastic naming of new species, revisions, and
changes in names and status. This has resulted in a bewildering array of possible
names and combinations of names. To be quite honest, far too many Rebutia species
have been described because of the demand and commercial pressures.
Consequently there is considerable confusion in this genus in trying to establish
clear relationships between the many taxa. The result of all this activity has been an
unwarranted cluttering of the synonymy of the group. Over 800 possible names and
combinations have been found in a search of the literature and we are sure that there
are many more that we have not found. One really begins to wonder when Rebutia
beryllioides has been described as a species, as a variety of 2 different species, a
subspecies and finally as a form of a third species. Is it any wonder that professional
taxonomists rarely work on a group of plants that are of horticultural value? With
apologies to the late Professor C. D. Darlington the following misquotation springs to
mind ‘cactus taxonomy is the pursuit of the impossible by the incompetent’.
We believe that the main cause of this confusion is that the genus Rebutia s. l. is in
an active state of evolution. This possibility does not appear to have been grasped by
the many individuals who have worked on the genus. It would account for the fact that
although on paper there appear to be 4 main sections which can further be
subdivided into 8 or more subsections, there are nevertheless some plants which
cross the boundaries, having characteristics which are intermediate between two or
more sections or subsections. The knowledge of the existence of these plants is
largely due to the field work of Friedrich Ritter who has found and introduced a large
number of new plants, some of which have intermediate characters. This makes the
reduction from a multigeneric group to a single genus desirable. A further
complication is the fact that many species are self fertile and inbreeding, so that the
occurrence of local races and inbred lines is highly likely.
The activities in the field by Ritter, Rausch, Lau, Knize, Vasques and others have
all helped in the proliferation of names. Although not all are to blame for the
subsequent naming of their introductions, they have all collected in the same areas,
often following in each others footsteps. The consequence of this has been the
introduction of many ‘new’ species with the great possibility that there has been
considerable duplication. Every hilltop or slope seems to be inhabited by yet
another ‘different’ species which has been described at the earliest opportunity.
Some of these plants have never been validly described but have been distributed
with field numbers and/or tentative names. What has made this situation worse has
been the careless subsequent propagation of some of this material. As a result there
is a distinct possibility that the plants when distributed are frequently wrongly
identified or wrongly numbered which renders them scientifically worthless.
It is with all this in mind that we have had a long hard cold look at the group as a
whole and have come to certain conclusions which represent our own current
thoughts on the taxonomy of the genus. It is not our intention to further clutter up the
synonymy, and so in the main we use the original scheme proposed by Bertrand in
1951 together with a number of refinements. We accept the view of Donald (1975)
that Sulcorebutia together with Weingartia have evolved from a separate
evolutionary line and are thus excluded from Rebutia.
5
 In proposing the following arrangement we realise that whatever scheme we
propose it will not be universally accepted. We have not slavishly followed any one
particular system but have attempted to amalgamate the best of a number of different
arrangements. One of the criteria we have used is the breeding system, i.e. whether a
species is self sterile or self fertile. This neatly divides the Section IV Rebutia into two
workable subsections. It also divides the large Backeberg genus Mediolobivia into
two distinct sections, Section II Mediolobivia and Section III Digitorebutia.There will
always be a problem with those plants which bridge the gap between the different
sections (being neither wholly self fertile, nor wholly self sterile), but we believe that
ours is the most workable scheme to date.
The following is an attempt at producing a description for the whole genus and is
based on Buxbaum (1958).
Rebutia K. Sch. emend Buining & Donald.
Dwarf, globular or slightly cylindrical plants usually making offshoots from the base.
Tubercles often distinct, spirally arranged and sometimes formed into distinct ribs.
Spines very variable in length and colour; sometimes short, stiff and bristle-like but
often long and flexuous. Flowers always produced from old areoles, usually
appearing crowded at the base and very large in relation to the size of the plant. The
flowers are brilliantly coloured, red, yellow, purple and occasionally white,
campanulate or funnel-shaped. The receptacular scales are sometimes naked but in
others with copious hairs and bristles. Stamens are produced from the funnel part of
the flower never from a ring in the throat. Occasionally the style is united to the
receptacle wall. The fruit is a small capsule which opens by a circular split so that the
lower half which remains on the plant is like a shallow bowl in which some of the
seeds remain. Seeds small, black and shining but also dull and earth coloured in
some species.

Classification of the genus Rebutia

Rebutia K. Sch. emend. Buining & Donald Type species Rebutia minuscula K. Sch.
Section I Aylostera (type Rebutia deminuta (Web.) Br. & R.)
Section II Mediolobivia (type Rebutia aureiflora Backbg.)
Subsection I Setirebutiae
Subsection II Cylindrorebutiae
Section III Digitorebutia (type Rebutia pygmaea (R. E. Fries) Br. & R.)
Subsection I Pygmaeae
Subsection II Euanthemae
Section IV Rebutia (type Rebutia minuscula K. Sch.)
Subsection I Rebutiae
Subsection II Mediorebutiae
Lists of the species which belong to a particular Section or Subsection, together with
distribution maps and locational data, are given in the chapter devoted to
geographical distribution.
6
 Analytical key to the Rebutia Sections
For reasons which have been explained, this key will not separate those species
which bridge the gap between the different sections i.e. those species with
intermediate characters. It should nevertheless help to delimit the majority of
species. It is based on a key published by Krainz (1967) and amended by B. Fearn.
A Body globular or cylindrical with the tubercles arranged on distinct ribs,
which may be vertical or spirally arranged. Body often suffused with
amethyst particularly during the winter months.
B Body cylindrical, flowers broad funnel-shaped to campanulate. Flowers
self sterile. Section II Mediolobivia Subsection II Cylindrorebutiae
BB Body globular or short cylindrical, flowers narrow funnel shaped,
lower part of receptacle usually fused with the base of style. Flowers self
fertile. Section III Digitorebutia
C Body obviously cylindrical, spines less than 5mm long, strongly
appressed, areoles arranged in vertical rows.
Section III Digitorebutia Subsection I Pygmaeae
CC Body globular, spines 5mm. or longer, always slightly erect never
appressed, areoles arranged spirally, occasionally in vertical rows.
Section III Digitorebutia Subsection II Euanthemae
AA Body globular to flattened globular, with indistinct ribs. Body green. Plants
usually densely spined.
C Receptacular scales with hairs and bristles, seeds ± earth coloured.
D Style not fused to the base of the receptacle.
Section II Mediolobivia Subsection I Setirebutiae
DD Basal portion of style usually fused with the lower portion of the
receptacle. Section I Aylostera
CC Receptacular scales naked (i.e. without hairs and bristles). Seeds black
and glossy, dome shaped usually with a prominent white basal hylum
area. Section IV Rebutia
E Flowers self fertile. Subsection I Rebutiae
EE Flowers self sterile. Subsection II Mediorebutiae
It is beyond the scope of this booklet to include a workable species key.
Flowers
All species in the genus Rebutia are day flowering and insect pollinated. Most
species are self fertile with the notable exception of 2 groups - Section II
Mediolobivia and Section IV Subsection II Mediorebutiae.
In Europe, Rebutias flower in late spring usually during the months of April and
May. The first species to flower is Rebutia marsoneri which usually flowers at the end
of February and beginning of March. It is thus one of the heralds of Spring.
Occasionally it flowers much earlier than this and on one notable occasion a plant of
this species was in flower at Abbey Brook on Christmas Day morning. The last plants
to flower are some members of Section I Aylostera such as R. deminuta, R.
pseudodeminuta and R. fiebrigii. There appears to be a distinct sequence in the
flowering of these plants. This time sequence may be of great biological significance
because in habitat it would maintain biological isolation even when the plants were
growing in close proximity to each other.

7
 In cultivation it is possible to change the flowering times of these plants by the
extension of cool temperatures. This has the effect of delaying the flowering time until
late May or June. Flowering time can also be altered by increasing the minimum
temperature during February thus shortening the winter period. This has the effect of
making the plants flower in March.
Although it is possible to alter the timing of flowering, there is one overall
requirement in the successful cultivation of these plants which appears to determine
whether the plants actually produce or do not produce flowers. In our experience
this is the winter rest period. It appears to be essential for the temperature to fall near
to or below freezing for some considerable time. Having due regard to the high
altitudes in the Andes of the natural habitats of these plants 1500-4000 metres altitude
(4900-13,000ft.) it is not really surprising that this is the case. At Abbey Brook we now
keep all our Rebutia plants in a cold unheated glasshouse during the winter together
with two other genera Lobivia and Oreocereus.
The cold requirement may well account for the poor flowering of these plants in
Southern California and the total lack of flower production in the tropics. We have
already suggested to our friends in the tropics that these plants should be wrapped in
newspaper and placed in a refrigerator (not a deep freeze) for some weeks. We
have not yet received the results of these experiments.

Floral Structure
The floral structure of the genus Rebutia has in the past been used to justify
splitting the single genus Rebutia into a number of separate genera. It is now
appreciated that such a split cannot be justified on the basis of floral structure alone,
as it has been observed that there are plants with a floral structure which is
intermediate between the different sections.
For example, the genus Aylostera was separated from the rest of Rebutia on the
observation that the base of the style was fused to the base of the corolla. In other
groups for example Mediolobivia and Rebutia the style is completely free from the
base of the corolla. Digitorebutia was an intermediate group in which the style was
only partially free. It is now recognised that these differences are not exhibited by all
the related species of a section and cannot therefore be used to justify the
maintenance of a number of separate genera.
Similarly the presence or absence of hairs and bristles in the axils of the
receptacle scales is again not a good universal character for splitting up the genus. It
has been found that there are a number of species which bridge the gap between the
previously recognised generic units. A number of fine illustrations of the floral
characters of the genus have been published Krainz (1967), and Brederoo (Donald &
Brederoo, 1978). The drawings in this booklet are by Brian Fearn and have not been
published previously.
8
 Rebutia senilis f. stuemeri nat. size

Rebutia minuscula f. grandiflora nat size

B 9
Rebutia pygmaea nat. size

Rebutia marsoneri nat. size

10
Rebutia einsteinii f. schmiedcheniana nat. size

Rebutia muscula nat. size

11
 REBUTIA CHRYSACANTHA
Fig. 3 Beautiful large flowered species closely related to R. senilis.

12
 The Geographical Distribution of the genus Rebutia
The geographical distribution of the genus Rebutia occurs in North West
Argentina and the southern provinces of Bolivia on the eastern side of the Andes. The
altitude varies between 1500-4000 metres (4,900-13,000ft. above sea level). The
genus can be split into four main taxonomic sections. Although these sections do not
fall into distinct geographical groupings, certain trends can be detected. Map No. 1
shows the important place names and country boundaries and Maps 2-4 the
geographical distribution of Rebutia species.
Section I Aylostera (Map No. 4)
This section has a very extensive distribution in Argentina and Bolivia. It occurs
from Tuome Province in the south, throughout Salta and Jujuy provinces in Argentina
to Tarija, Potosi, Chuquisaca and Santa Cruz Departments of Bolivia.
a) Rebutia deminuta Weber and allied species. Rebutia deminuta is the most
southerly member of this group coming from Trancas, Tucuman province
Argentina. The closely related form Rebutia pseudominuscula occurs over an
extensive area from central Salta province through Jujuy province and into
Tarija.
b) Rebutia pseudodeminuta Backbg. and allied species. Rebutia pseudo-
deminuta is widely distributed from south Salta to central Tarija and into
Chuquisaca and Potosi. Many of the recently described ‘species’ of Ritter,
Rausch and Lau represent geographical forms of R. pseudodeminuta. (e.g. R.
maxima FR755, R. albiareolata FR761, R. tamboensis FR1142 from Tarija and R.
nitida FR769 from Chuquisaca.)
c) Rebutia kupperiana Böd. and allied species. Rebutia kupperiana is restricted in
habitat to an area around Narvaez in Tarija department Bolivia although several
other related Ritter ‘species’ belonging to this group have been described as
coming from south Chuquisaca. (e.g. R. tuberosa Ritt., and R. rubiginosa Ritt.)
d) Rebutia fiebrigii (Guerke) Br. & R. and allied species. The Rebutia fiebrigii
complex occurs over an extensive area of southern Bolivia and northern
Argentina. The main area of distribution occurs from Tarija and Chuquisaca in
Bolivia to northern Salta in Argentina. There is an outlier of distribution in Santa
Cruz department where Rebutia ithyacantha (Card.) Diers, R. donaldiana Lau
and Rowley and R. vallegrandensis Card. occur. Rebutia spinosissima has a
very extensive distribution from north Salta in Argentina through Tarija and into
south Chuquisaca in Bolivia.
Rebutia fiebrigii itself has by far the largest distribution area of any member of
this group occurring from La Quiaca in Jujuy province in Argentina to central
Chuquisaca in Bolivia.
e) Rebutia spegazziniana Backbg. and allied species. This species is found in
Salta province near to the border with Bolivia and into Tarija department from
Padcaya to Tarija. The closely related ‘species’ R. fusca Ritt., R. vulpina Ritt., and
R. tarvitaensis Ritt., are all found in Chuquisaca department, Bolivia.
f) Rebutia heliosa Rausch. This species has a limited distribution area near
Narvaez in Tarija department, Bolivia. There are a number of ‘species’ closely
related to R.heliosa apart from var.cajasensis and var.conderensis,these are R.
perplexa and R. espinosa. Both of these are found in Tarija near to R. heliosa
and at Southern Cinti R. albopectinata occurs.
13
Section II Mediolobivia
This series has a very limited distribution occurring only in Argentina in an area
of north Salta and south Jujuy provinces.
Subsection I Setirebutiae. Plants in this section all occur near to the Salta - Jujuy
border around Quebrada del Toro. Rebutia aureiflora var. elegans and related forms
have the most southerly distribution of this group occurring around Quebrada
Escoipe.
Subsection II Cylindrorebutiae. There is probably only a single species which
belongs in this section - R. einsteinii. The myriad of different names appear to refer to
geographical variants. These plants have been found in either Salta or Jujuy
provinces.
Section in Digitorebutia. (Map No. 2)
This section has a very extensive distribution from Salta and Jujuy provinces in
Argentina in the south to the Bolivian departments of Tarija, Chuquisaca, Potosi and
Oruro in the north.
There appear to be several separate centres of distribution where a number of
named species occur in close proximity. It is here suggested that these ‘species’
should be subjected to an intensive ecological investigation to determine whether
these populations are biologically isolated, by pollination vectors and flowering
times. Admittedly members of this series are all self fertile, but their highly coloured
flowers would attract pollinating insects. Gene flow between populations would
occur if the populations which were in close proximity flowered at the same time.
We have determined that there are 5 separate areas of distribution for the
majority of members of this Section.
a) Argentina - Bolivia border areas, Jujuy province.
e.g. R. eos Rausch, R. costata Werd., R. pygmaea (R. E. Fries) Br. & R.
b) Mendez province, Tarija, Bolivia.
e.g. R. brunneoradicata Ritt., R. colorea Ritt.
c) Camargo - Culpina region, Cinti province, Chuquisaca, Bolivia,
e.g. R. diersiana Rausch, R. mixta Ritt.
d) South Chichas province, Potosi, Bolivia,
e.g. R. iridescens Ritt., R. rosalbiflora Ritt.
e) Environs of Potosi City, Potosi, Bolivia.
e.g. R. pygmaea, R. costata f. pilifera Buin. & Donald.
The species found in Oruro also belong here. The proliferation of names which
is particularly true of section III, are at best, geographical variants of very few
species.
Section IV Rebutia (Map No. 3)
Subsection I Rebutiae.
This subsection contains three species complexes. These are R. minuscula, R.
senilis and R. xanthocarpa. The R. minuscula complex includes R. grandiflora
Backbg.,R. violaciflora Backbg.and R. kariusiana.The group is totally Argentinian in
distribution ranging from the north Tucuman to Salta and Jujuy provinces. The area is
bounded by Tucuman City, Quebrada Escoipe, Cachipampa and Quebrada del
Toro. R. minuscula occurs throughout this region apparently remaining

14
morphologically constant until reaching the northern limit of its distribution where it
appears to intergrade with R. violaciflora.
Subsection II Mediorebutiae.
This subsection contains two species complexes. These are R. wessneriana and
R. marsoneri. It appears that this subsection has a much wider distribution than
Subsection I Rebutiae, ranging from south Salta province in Argentina e.g. R.
wessneriana Bew. to Huari-Huari, Potosi department, Bolivia e.g. R. binnewaldiana
W. Heinr. and R. fiebigiana W. Heinr. Some doubt has been raised as to the
authenticity of the last two species. The habitat data may be incorrect. Seed
distributed from these habitats has produced plants that resemble species of Series I
Aylostera (Donald, Ashingtonia 2 67). If this is correct then the distribution is
restricted to Jujuy and Salta provinces of Argentina and into Tarija department
Bolivia where R. wessneriana is also found. The R. marsoneri complex is distributed
throughout northern Jujuy province, Argentina.

Distribution maps and locational data

The following is a list of the species together with locational data which have
been used in preparing the distribution maps. It is interesting to note that often
species belonging to the same Rebutia section (i.e. closely related) have often been
collected at the same place. The altitude at the different sites may vary, but the vital
information on genetic isolation is not available.Without this, together with ecological
and experimental taxonomic data the suspicion is that these different ‘species’ are
identical merely representing the range of species variation. For example the
following species numbers 18, 20, 22, 24, 40, 41, 44, 45 have the same location - Mal
Paso, Sth. Chichas, Potosi, Bolivia. Also species numbers 4, 12, 16, 26, 36, 46 occur at -
Camargo - Culpina, Chuquisaca, Bolivia.

Rebutia Section III Digitorebutia (See Map No. 2)

1 brachyantha (Wessn.) Buin. & Donald. Tarija, Tarija, Bolivia.
2 brunescens Rausch. Huari-Huari, Potosi, Bolivia.
3 brunneoradicata Ritter. San Antonio, Mendez, Tarija, Bolivia.
4 canacruzensis Rausch. Camargo-Culpina, Chuquisaca, Bolivia.
5 carmeniana Rausch. Caspala, Jujuy, Argentina.
6 cincinnata Rausch. Potosi, Potosi, Bolivia.
7 colorea Ritter. San Antonio, Mendez, Tarija, Bolivia.
8 costata (Werd.) Krainz. La Quiaca, Jujuy, Argentina.
9 costata f. eucalyptana (Backbg.) Buin. & Donald. Eucaliptos, La Paz, Bolivia.
10 costata f. pilifera Buin. & Donald Potosi, Potosi, Bolivia.
11 densispectinata Ritter nomen nudum. Jucanas, Tarija, Bolivia.
12 diersiana Rausch. Camargo-Culpina, Chuquisaca, Bolivia.
13 eos Rausch. Tafna, Jujuy, Argentina.

15
 14 euanthema (Backbg.) Buin. & Donald. Quebrada de Humahuaca, Jujuy,
Argentina.
15 euanthema f. oculata (Werd.) Buin. & Donald. Humahuaca, Jujuy, Argentina.
16 friedrichiana Rausch. Camargo-Culpina, Chuquisaca, Bolivia.
17 graciliflora Backbg. Chichas, Potosi, Bolivia.
18 gracilispina Ritter. Mal Paso, Sth. Chichas, Potosi, Bolivia.
19 hirsutissima Card. Bueto, Villaserrano, Chuquisaca, Bolivia.
20 iridescens Ritter. Mal Paso, Sth. Chichas, Potosi, Bolivia.
21 iscayachensis Rausch. Iscayachi, Tarija, Bolivia.
22 lanosiflora Ritter. Mal Paso, Sth. Chichas, Potosi, Bolivia.
23 minutissima Ritter. Tarija, Tarija, Bolivia.
24 mixta Ritter. Mal Paso, Sth. Chichas, Potosi, Bolivia.
25 mixticolor Ritter. San Antonio, Mendez, Tarija, Bolivia.
26 mudanensis Rausch. Camargo-Culpina, Chuquisaca, Bolivia.
27 odontopetala Ritter. Cueva, Sth. Cinti, Chuquisaca, Bolivia.
28 patericalyx Ritter. Cueva, Sth. Cinti, Chuquisaca, Bolivia.
29 pauciareolata Ritter. San Antonio, Mendez, Tarija, Bolivia.
30 paucicostata Ritter. San Antonio, Mendez, Tarija, Bolivia.
31 poecilantha Ritter. North Cinti, Chuquisaca, Bolivia.
32 potosina Ritter. Potosi, Potosi, Bolivia.
33 pseudopygmaea Ritter nomen nudum. San Antonio, Mendez, Tarija, Bolivia.
34 pygmaea (R. E. Fries) Br. & R. Yavi, Jujuy, Argentina., Potosi, Potosi, Bolivia.
35 pygmaea f. haefneriana (Cullm.) Buin. & Donald. Oruro, Oruro, Bolivia, Iturbe,
Argentina, Chichas, Potosi, Bolivia.
36 raulii Rausch Camargo-Culpina, Chuquisaca, Bolivia.
37 rauschii Zecher. Huari-Huari, Potosi, Bolivia.
38 ritteri (Wessn.) Buin. & Donald, Iscayachi, Tarija, Bolivia.
39 ritteri f. nigricans (Wessn.) Buin. & Donald. Quebrada del Toro, Salta,
Argentina.
40 rosalbiflora Ritter. Mal Paso, Sth. Chichas Province, Bolivia.
41 rutiliflora Ritter. Mal Paso, Sth. Chichas Province, Bolivia.
42 salpingantha Ritter. Villazon, Potosi, Bolivia.
43 tarvitaensis Ritter. Tarvita, Azurduy, Chuquisaca, Bolivia.
44 torquata Ritter. Mal Paso, Sth. Chichas Province, Bolivia.
45 tropaeoliptica Ritter. Mal Paso, Sth. Chichas Province, Bolivia.
46 violascens Ritter. Camargo, Cinti, Chuquisaca, Bolivia.

Rebutia Section II Mediolobivia Subsection I Setirebutiae (See Map No. 2)

55 R. aureiflora Backbg. Quebrada del Toro, Salta, Argentina.
56 R. aureiflora f. rubelliflora (Backbg.) Buin. & Donald. Quebrada del Toro,
Salta, Argentina.
57 R. aureiflora f. duursmaiana (Backbg.) Buin. & Donald. Yacones, Salta,
Argentina.
58 R. aureiflora f. boedeckeriana (Backbg.) Buin. & Donald. Nr. Chani Volcano,
Jujuy, Argentina.
59 R. aureiflora v. elegans (Backbg.) Buin. & Donald. Quebrada Escoipe, Salta,
Argentina.
16
Rebutia Section IV Rebutia Subsection I Rebutiae (See Map No. 3)
60 fabrisii Rausch. Valle Grande, Salta, Argentina.
61 minuscula K. Sch. Tucuman, Tucuman Prov., Argentina.
62 minuscula var. grandiflora (Backbg.) Buin. & Donald. Quebrada Escoipe,
Salta, Argentina.
63 minuscula var. violaciflora (Backbg.) Buin. & Donald. Quebrada Escoipe,
Salta, Argentina.
64 margarethae Rausch. Iruya, Santa Victoria, Salta, Argentina.
65 padcayensis Rausch. Padcaya, Tarija, Bolivia.
66 senilis Backbg. Quebrada Escoipe, Salta, Argentina.
67 xanthocarpa Backbg. Quebrada del Toro, Salta, Argentina.

Rebutia Section IV Rebutia Subsection II Mediorebutiae (See Map No. 3)

68 binnewaldiana W. Heinr. Huari-Huari, Potosi, Bolivia.
69 fiebigiana W. Heinr. Huari-Huari, Potosi, Bolivia.
70 krainziana Kess. Yavi, Jujuy, Argentina.
71 marsoneri Werd. North Jujuy, Argentina.
72 marsoneri f. sieperdaiana (Buin.) Buin. & Donald. Salta, Salta, Argentina.
74 wessneriana Bewg. Quebrada de Humahuaca, Jujuy, Argentina; Salta,
Salta, Argentina; Tarija, Bolivia and Guachipas, Escoipe, Salta, Argentina.

ebutia Section I Aylostera (See Map No. 4)

R
76 albiflora Ritter & Buin. Rio Pilaya, North East Tarija, Bolivia.
77 albipilosa Ritter. Narvaez, Tarija, Bolivia.
78 albiareolata Ritter. Padcaya, Arce, Tarija, Bolivia.
79 albopectinata Rausch. Culpina, Chuquisaca, Bolivia.
80 aureispina Knize nomen nudum. Jucanas, Tarija, Bolivia.
81 buiningiana Rausch. Nr. Iruya, Salta, Argentina.
82 camargoensis Rausch. Camargo-Culpina, Chuquisaca, Bolivia.
83 cajasensis Ritter. Tarija, Tarija, Bolivia.
84 cintiensis Ritter. Camargo, Sth. Cinti, Chuquisaca, Bolivia.
85 deminuta (Weber.) Br. & R. Trancas, Tucuman, Argentina.
86 donaldiana Lau & Rowley. Pucara, Vallegrande, Santa Cruz, Bolivia.
87 fiebrigii (Guerke) Br. & R. Iscayachi, Tarija, Bolivia.
88 flavistyla Ritter. Tarija, Tarija, Bolivia.
89 froehlichiana Rausch. Yuquina, Cinti, Chuquisaca, Bolivia.
90 fulviseta Rausch. Padcaya, Arce, Tarija, Bolivia.
91 fusca Ritter. Cinti Chuquisaca, Bolivia.
92 heliosa Rausch. Road to Narvaez, Tarija, Bolivia.
93 hoffmannii Rausch. Santa Victoria, Salta, Argentina.
94 huasiensis Rausch. Inca Huasi, Cinti, Chuquisaca, Bolivia.
95 ithyacantha (Card.) Diers. North Comarapa, Valle Grande, Santa Cruz, Bolivia.
96 jujuyana Rausch. Quebrada de Humahuaca, Jujuy, Argentina.
97 jujuyensis Lau. Santa Victoria, Salta, Argentina.
98 kieslingii Rausch. Caspala, Jujuy, Argentina.
99 kupperiana Böd. Narvaez, Tarija, Bolivia.
C 17
 100 lauii Donald. Cajas Pass, Tarija, Bolivia.
101 leucanthema Rausch. Camargo-Culpina, Chuquisaca, Bolivia.
102 mamillosa Rausch. Camargo, Chuquisaca, Bolivia.
103 muscula Ritter & Thiele. Narvaez, Tarija, Bolivia.
104 napina Ritter. South Cinti, Bolivia.
105 nitida Ritter. Tarvita Azurduy, Chuquisaca, Bolivia.
106 nogalensis Ritter. Tarvita Azurduy, Chuquisaca, Bolivia.
107 pseudodeminuta Backbg. South Salta, Argentina - to - Chuquisaca, Bolivia.
108 deminuta f. pseudominuscula (Speg.) Buin. & Donald. Cachi, Salta, Argentina.
109 pulchella Rausch. North of Padilla, Chuquisaca, Bolivia.
110 pulvinosa Ritter & Buin. Narvaez, Tarija, Bolivia.
111 robustispina Ritter. Tarija, Tarija, Bolivia.
112 rubiginosa Ritter. South Cinti, Chuquisaca, Bolivia.
113 schatzliana Rausch. Camargo-Culpina, Chuquisaca, Bolivia.
114 spegazziniana Backbg. Salta, Victoria - to - Tarija, Bolivia.
115 sphaerica Ritter. Tarija, Tarija, Bolivia.
116 spinosissima Backbg. Santa Victoria, Salta, Argentina.
117 supthutiana Rausch. Camargo-Culpina, Chuquisaca, Bolivia.
119 tarijensis Rausch. Tarija, Tarija, Bolivia.
120 tarvitaensis Ritter. Tarvita, Azurduy, Chuquisaca, Bolivia.
121 tuberosa Ritter. South Cinti, Chuquisaca, Bolivia.
122 vallegrandensis Card. Candellaria, Valle Grande, Santa Cruz, Bolivia.
123 vulpina Ritter. Cinti, Chuquisaca, Bolivia.
124 zecheri Rausch. Iscayachi, Tarija, Bolivia.

Rebutia Section II Mediolobivia Subsection II Cylindrorebutiae (See Map No. 4)

125 einsteinii Frič ex Backbg. La Quiaca, Jujuy, Argentina.
127 einsteinii var. gonjianii (Kies.) Donald. Tilcara, Jujuy, Argentina.
128 einsteinii f. schmiedcheniana (Kohl.) Buin. & Donald. Chani Volcano, Jujuy,
Argentina.
129 einsteinii f. rubriviridis (Frič & Kreuzgr. ex. Backbg.) Donald.
Quebrada del Toro, Salta, Argentina.
130 einsteinii f. conoidea (Wessn.) Buin. & Donald. Cham Volcano, Jujuy, Argentina.

18
19
20
21
22
 Cultivation
Cultivation of the majority of species should present no real problems. In the
growing season (March-October) they need a considerable amount of water. This
may mean watering 2 or 3 times a week in hot sunny weather but normally once a
week should be sufficient. Try not to get the soil soaking wet which leads to bloated
plants which are very susceptible to soft rot. Wet soil also encourages Sciara Fly
which can also lead to loss of plants. In the winter (October-March) the watering
requirements are completely different as the plants are not growing. For successful
flower production it appears to be essential for the temperature to fall near to or
below freezing point for some considerable time. During this period the plants must
be kept absolutely dry and you will then be rewarded by the production in early
spring of the superb, brightly coloured flowers produced by all members of this
genus.
All Rebutia species need plenty of light and a good sunny position is essential.
Although these plants can be successfully grown on the window sill, a greenhouse or
conservatory is a better place especially if you wish the plants to flower. We have
found from long experience that plants that are watered and kept warm 7°C (45°F) or
above in the winter are very shy flowering.
We can recommend the following compost although Rebutias can be grown
successfully in virtually any mixture:- 3 or 4 parts of Fisons Levington Potting
Compost and 1 part of coarse lime-free sand.
We grow the majority of our plants in this mixture. It has also been found
beneficial to feed the plants during the growing season with a low nitrogen, high
potash fertiliser such as Fisons Tomorite. We normally feed the plants 3 times during
the growing season i.e. once every 6 weeks using the recommended strength
fertilizer. Repotting is only necessary when the plants have outgrown the container
they are growing in. Repotting is usually done at the beginning of the growing
season. If regular feeding is given as recommended, repotting once in 2 or 3 years
may be all that is necessary, particularly with the slower growing species.
Vegetative Propagation. Rebutias can be propagated by removing and rooting
offsets but it is not a recommended practice as the plants are so easy to grow from
seed. R. albiflora is the only real exception and this species is usually propagated by
rooting offsets as seed is rarely available. Grafting is also sometimes used to
propagate the slower growing species such as R. aureiflora, R. einsteinii and R.
heliosa but it is not really necessary as the plants grow well on their own roots.
Growing from seed. Rebutias are best propagated from seed. If you are collecting
your own seed it is best to collect the fruits when they are fully ripe at the end of the
growing season. Many species produce copious quantities of seed, others have only
a few seeds in each capsule. Carefully separate the seeds from any adhering capsule
wall or debris. They can then be stored or sown immediately.
To be successful you must try to imitate the conditions in which the plants grow
naturally. Cacti are nearly always found under scrub or small bushes, on rocky slopes
or on level ground where the soil is porous. This is because:-
(1) The slope and porous soil provide adequate drainage.
(2) The scrub provides shade for the seedlings and young plants.
(3) The rocks radiate warmth at night and the crevices between them trap
pockets of humid air.
23
When to sow. If you do not have a heated propagator, the best time to sow is in
Spring. i.e. from March until the end of May. Do not sow later than this as the
seedlings will not be large enough to withstand the following winter.
If you have a heated propagator or a warm window sill (Do not use the airing
cupboard) you can sow at any time of the year, but January and February are the best
months.
Compost. We can recommend the following composts.
Either:
(1) Equal parts of Fisons Levington potting compost and coarse lime-free sand.
or (2) One part John Innes No. 1 or 2 and ¼ part lime-free sand.
Sowing. Fill the seed trays or shallow pots to within ¼'' of the top with compost and
level the surface, removing any large lumps. Water with a watering can fitted with a
fine hose. Scatter the seed thinly on the surface of the damp compost. Seeds can be
spaced out and lightly pressed level with the surface using a pencil. Cactus seed
need light before they will germinate, so do not cover the seeds with compost and do
not put the containers in a dark cupboard.
Temperature. Temperatures of 12-22°C (55-70°F) are best for germination. Once
the seedlings are 2-3 weeks old normal growing temperatures are adequate.
Germination. The percentage of seeds that germinate depends on the species, age
of the seed and temperature, but there should be signs of germination within 2-3
weeks. A few seeds may be slow to germinate, and seedlings may not appear until
2-3 months after sowing.
Watering. If possible, water with a fine mist spray, or place the container in a
saucer and water from below. Never let the compost dry out, particularly before
germination has taken place, or the germinating seeds will be killed. Conversely, do
not allow the compost to become waterlogged either, else the seeds will rot. After
germination keep soil moist - not too wet, not too dry. During the first winter the
seedlings can be kept completely dry in a minimum temperature of 2°C (35°F).
Care of the young seedlings. Keep the container in an airy place to prevent
‘damping off. Shade the seedlings with a single thickness of tissue or newspaper for
the first two months, then avoid direct sunlight for a further 6 months. After that time
treat as adult plants. Try to ensure the seedlings remain a healthy deep green (or
occasionally brownish) colour - a bright red colour indicates that too much light has
been given, thin spindly pale green seedlings results from too little light.
Transplanting. Do not transplant seedlings until they begin to touch one another in
the container - this will depend on the rate of growth of individual species. If in doubt,
leave the seedlings in the same container for 1 year, and do not transplant until the
spring of the second year. They will not come to any harm even if they look
overcrowded.
Detailed germination studies on Rebutia xanthocarpa f. salmonea have been
made Fearn (1974). It was found that no seed germinated below 9°C or above 26°C It
was found that this species when compared with other species from South America
e.g. Gymnocalycium spegazzinii, Frailea pumila and Helianthocereus pasacana has
a very restricted temperature range. At least 50% germination occurred between
11∙5°C and 22∙8°C (52°F - 70°F) a range of 11∙3°C.

24
Fig. 4 The germination of R. xanthocarpa f. salmonea has been plotted on a day to
day basis and the figures on the graph indicate the number of days
incubated.

Fig. 5 The curves obtained for H. pasacana and R. xanthocarpa f. salmonea are
shown. On successive days the maximum and minimum temperatures at
which 50% of maximum germination was obtained. The resulting U shaped
curves reflect the temperature span over which 50% germination occurred,
the rate of germination at different temperatures and the position of
maximum and minimum temperature at which 50% of maximum germination
was reached.

D 25
 Pests and Diseases
Mealy Bugs are the most common pests of cacti. They look a bit like tiny wood lice but
with a white waxy coat. They usually aggregate in areas which are not affected by
overhead watering, e.g. in the axis of the tubercles and around the base of the plant.
Spraying with Malathion or a combination with Dimethoate will eradicate them.
Root Mealy Bugs usually remain unseen until the plants are repotted. They occur as
white woolly colonies which feed on the roots. Watering with a systemic insecticide
such as Dimethoate should eradicate them. Repeat the dose after seven or ten days.
Red Spider Mite can be a serious pest. Its presence may remain undetected until the
plants have become severely disfigured with a reddish-brown area on the
epidermis. It is a very tiny reddish creature just visible to the naked eye. Water or
spray the plants with a contact or systemic insecticide Repeat the dose after ten
days.
Sciara Fly is a relative newcomer as far as pests of Cacti are concerned. The first
warning may be the presence of a few tiny black flies near the plants. The real pest is
the larvae which lives just under the soil surface and can cause damage and rot the
base of the plants. Total collapse of the plant may result. Watering with a systemic
insecticide should eradicate the larvae.
Aphids are not an important pest in the cactus collection. They occasionally occur on
flower buds and on young growth and any standard insecticide should deal with
them.

REBUTIA XANTHOCARPA f. VIOLACIFLORA

A charming plant with delicate lilac-rose coloured flowers

26
 Rebutia Descriptions
We have selected the following 20 species as representatives of the range of
variation exhibited by the genus Rebutia. It includes plants from all the Sections and
many of them are old favourites having been in cultivation for some considerable
time e.g. R. kupperiana, and R. minuscula. Other species which have been included
such as R. albiflora, R. muscula and R. heliosa, are among the best of recent
introductions.

REBUTIA ALBIFLORA
Fig. 6 A very distinctive and beautiful species with white flowers.

27
 REBUTIA ALBIFLORA
Rebutia albiflora Ritt. & Buin. Taxon (1963) 12 29
Synonyms Aylostera albiflora (Ritt. & Buin.) Backbg.
Classification Section I Aylostera
Body Individual heads up to 25mm. in diameter but usually much smaller
than this. Plants freely offsetting from the base and eventually
forming large clumps. Epidermis grey-green.
Areoles tiny, more or less circular, pure white
Spines differentiated into approximately 15 radials and 5 centrals, up to
5mm. long, straight and glassy white.
Flowers inner petals white, outer petals often with a pronounced pink
mid-line, 25mm. wide and 30mm. long. The tube is pink and has
short white bristles at the base.
Fruit small, 3-4mm. in diameter, reddish-pink and contains few seeds.
Seed small, matt black.
Type locality Rio Pilaya, N.E. Tarija, Bolivia. Map 4, No. 76.
Discovered by Friedrich Ritter (FR766a)
Notes A very distinctive and beautiful species. It is unusual in both habitat
and flower for a Rebutia species. The free clumping nature of this
species produces beautiful small mounds of white heads. The
white flowers are small, but the unusual colour makes the plant a
real gem.
The majority of plants in cultivation appear to be self sterile and
from the apparent lack of different clones, seed is rare. We believe
that most of the plants in cultivation have been propagated
vegetatively from the clone originally introduced by H. Winter of
West Germany in the late 1950’s.
Recently, an interesting hybrid has been produced with R. heliosa.
The plants have the body form of R. heliosa but with white flowers.
Kakt. u. a. Sukk. (1978) 29 184.
REBUTIA ALBIPILOSA
Rebutia albipilosa Ritter Taxon (1963) 12 29
Synonyms Aylostera albipilosa (Ritt.) Backbg.
Classification Section I Aylostera.
Body up to 40mm. in diameter becoming caespitose with age. Epidermis
dark green.
Areoles elliptical, with white wool, raised on short tubercles and arranged
on ribs.
Spines c. 20 in number, strongly differentiated in centrals and radials,
mostly 7-10mm. long. However, a few spines become extremely
long especially at the base of the plant. These spines are often
30-40mm. long. All spines are white, the longest ones having dark
brown tips.
28
Flowers 45mm. long and 30mm. wide, petals orange-red in colour often with
a darker red midstripe. Tube and ovary with hairs and bristles.
Filaments and style white with a 6 lobed cream coloured stigma.
Self fertile.
Fruit 5mm. in diameter, brownish-green and covered with hair and
bristles.
Seeds dome shaped, dull black.
Type locality Narvaez, Department Tarcia, Bolivia. Map 4, No. 77.
Notes This is another of the Rebutia species found by Friedrich Ritter
(FR754) in the early 1960’s. It is a very variable species particularly
with reference to the length of the spines. Several other ‘species’
are probably referable here for example R. vulpina (FR939) and R.
robustispina (FR763). The original plant depicted by Ritter with
very long and dense white spines appears to be an extreme form.
Nobody has yet recollected or seen another comparable plant.
Donald (Ashingtonia 2 180) has speculated that Ritter’s R. flavistyla
(FR756) is the true R. albipilosa. If this is correct and confirmed by
Ritter it calls into question a whole series of descriptions and
relationships. It is interesting to note that Ritter found and
described both species, (see colour plate and fig. 1.).

REBUTIA AUREIFLORA
Fig. 7 An uncommon, slow growing species with very large flowers.

29
 REBUTIA AUREIFLORA
Rebutia aureiflora Backbg. D. Kaktfrd. (1932) 124
Synonyms Mediolobivia aureiflora (Backbg.) Backbg.
Mediolobivia aureiflora var. albiseta Backbg.
Mediolobivia aureiflora var. albilongiseta Backbg.
Mediolobivia aureiflora var. albispina Backbg. nomen nudum
Mediolobivia spiralisepala Schutz
Rebutia aureiflora var. albilongiseta (Backbg.) Marshall & Bock,
comb. nud.
Rebutia aureiflora var. albiseta (Backbg.) Marshall & Bock. comb,
nud.
Rebutia aureiflora var. longiseta (Backbg.) Marshall & Bock. comb.
nud.
Rebutia aureiflora var. densispina Borg nomen nudum
Rebutia aureiflora subspec. aureiflora (Backbg.) Donald
Classification Section II Mediolobivia Subsection II Setirebutiae
Body 40mm. in diameter becoming caespitose from the base, eventually
forming clumps 100mm. in diameter.
Areoles elliptical with white wool
Spines c. 20, extremely variable not strongly differentiated into centrals
and radials. Seedlings and offshoots on older plants with short
spines 6-8mm. long yellowish at first, becoming all white and
tipped yellowish-brown. Older plants with extremely long thin
flexuous spines 50mm. or more in length, white tipped yellowish-
brown.
Flowers very large, 50mm. long and 45mm. in diameter, orange-yellow
petals, throat white, tube light brownish-orange with small
greenish-brown scales long thin flexuous hair and bristles are
produced up to 10mm. long. Stigma yellow, style and filaments
white, self sterile.
Fruit large, 8mm. in diameter brownish-orange densely covered with
long hairs.
Seeds dull brownish-black
Type locality Quebrada del Toro, Salta, Argentina. Map 2, No. 55.
Notes This species belongs to a group of slow growing plants which are
self sterile. Plants are rarely offered for sale mainly because what
seed is available germinates very erratically. In our experience
10% germination is all that can be expected. There are fourteen
described varieties and in all probability they can all be included
within the variation of the one species. For example var.
albilongiseta has long pure white bristles, var. albiseta has 10mm.
long white spines, var. boedeckeriana has 11-12 thin radial spines
and pale orange flowers, var. rubelliflora with orange-red flowers,
var. rubriflora with red flowers., In our opinion these differences do
not justify varietal status.

30
 REBUTIACOSTATA
Fig. 8 A rare and slow growing species that is shy flowering. The plant in the
photograph has been in cultivation at Abbey Brook for over 20 years.

REBUTIA COSTATA
Rebutia costata Werd. Notizbl. Bot. Gart. u. Mus. Berlin (1934) 12 25
Synonyms Lobivia costata (Werd.) Wessn.
Digitorebutia costata (Werd.) Buin.
Mediolobivia costata (Werd.) Krainz.
Classification Section III Digitorebutia Sub Section II Euanthemae
Body up to 25mm. in diameter simple at first but becoming caespitose
with age. Epidermis deep bluish-green.
Areoles elliptical with brownish wool, raised on tubercles arranged in 8-10
distinct ribs.
31
Spines centrals 0 (old areoles may have 1-2), radials 9-12 (14), pectinate
and strongly projecting up to 7mm. long. White becoming
yellowish-brown at the base.
Flowers up to 35mm. long and 30mm. in diameter, deep orange, white throat
buds green, tube greenish-brown with olive green scales and
copiously hairy with a few long white bristles. Stigma cone shaped
5 lobed, greenish-yellow style greenish filaments lavender pink.
Seeds brownish.
Type locality The type locality is unknown. It has recently been recollected by
W. Rausch at La Quiaca, Jujuy, Argentina (R508).
Notes This is a rare, slow growing species - the plant depicted in fig. 8 has
been in cultivation for over 20 years and is still in a 3ins. pot. It is shy
flowering and rarely sets seed.

REBUTIA DEMINUTA
Fig. 9 An easily flowered and widely grown species that was one of the first species
to be discovered and described.
32
 REBUTIA DEMINUTA
Rebutia deminuta (Web.) Br. & R.
Synonyms Echinopsis deminuta Web. Bull Mus. Hist. Nat. Paris (1904) 386
Echinocactus deminutus (Web.) Gürke
echinorebutia deminuta (Web.) Frič & Krzgr.
Aylostera deminuta (Web.) Backbg.
Classification Section I Aylostera
Body individual heads about 50-60mm. high, dark green. Plants freely
offsetting from the base and eventually forming large clumps more
than 120mm. in diameter.
Areoles elliptical, light brownish.
Spines 10-12, not strongly differentiated into centrals and radials. Spines
white tipped brown and 5-6mm. long. Central when present
usually all brown.
Flowers very free flowering, flowers up to 40mm. long and 30mm. broad.
Petals deep orange-red with pink filaments and an 8 lobed white
stigma. Small greenish scales and white hairs present on flower
tube. Self fertile.
Fruit 5mm. in diameter, greenish-red in colour and covered with
bristles.
Seeds small, matt black.
Type locality Trancas, Tucuman State, N. Argentina. Map 4, No. 85.
Notes Easily flowered and widely grown species.
f. pseudominuscula (Speg.) Buin. & Donald body grey-green, heads
slightly smaller than the type, spines 7-14, longer, up to 8-9mm.,
yellowish white, brown at base and tip. Flowers as in the type.

REBUTIA EINSTEINII f. SCHMIEDCHENIANA

Rebutia einsteinii f. schmiedcheniana (Köhler) Buin. & Donald
Synonyms Lobivia schmiedcheniana Köhler Beitr. z. Skde. u. pflege, (1939) 37
Cylindrorebutia schmiedcheniana (Köhler) Donald
Mediolobivia schmiedcheniana (Köhler) Krainz.
Rebutia schmiedickenana (Köhler) Marsh & Bock comb. nud.
(should be Köhler and schmiedcheniana).
Classification Section II Mediolobivia Sub Section II Cylindrorebutiae
Body up to 35mm. in diameter becoming cylindrical with age. Some
plants become caespitose but this is not always so. The epidermis
is a dark brownish-green often tinged deep purple particularly
during the winter.
Areoles elliptical with sparse brownish wool raised on short tubercles
spirally arranged.
E 33
Spines slightly differentiated into two series, 15 radials appressed,
pectinate, 12 up to 75mm. long, greyish-white, brownish at the
base, upper 3 longer up to 18mm. and projecting. 1 (2) central
spines to the upper three radials. Upper spines and centrals light
brownish.
Flowers 45mm. long and 35-40mm. in diameter, outer petals and sepals
deep golden yellow. Stigma, style and filaments pale yellow tipped
dark brownish yellow. Ovary and tube with copious white hair and
long white bristles.
Fruit large, 7mm, self fertile. Green tinged purple.
Seed large, dome shaped, brownish.
Type locality Cham volcano, Salta, N. Argentina. Map 4, No. 128.
Notes This species has a very large underground tuberous root system
and for this reason needs a gritty porous compost. Plants are most
often seen grafted but this is unnecessary with careful cultivation.

REBUTIA EINSTEINII f. SCHMIEDCHENIANUM

Fig. 10 An uncommon species with a very large tuberous root system and large
golden yellow flowers
34
 REBUTIA EUANTHEMA
Fig. 11 An easily grown popular species with very showy flowers.

REBUTIA EUANTHEMA
Rebutia euanthema (Backbg.) Buin. & Donald
Synonyms Lobivia euanthema Backbg. Blatt. f. Kakt. (1934) 2
Mediolobivia euanthema (Backbg.) Krainz.
Acantholobivia euanthema (Backbg.) Y. Ito
Digitorebutia euanthema (Backbg.) Buin.
Echinolobivia euanthema (Backbg.) Y. Ito
Classification Section III Digitorebutia Subsection II Euanthemae
Body heads 30mm. in diameter, grey-green often tinged amethyst, freely
offsetting from the base and forming clumps 70-80mm. in diameter.
Areoles elliptical, light brownish in colour.
Spines often pectinate, about 10 radials, centrals absent Up to 10mm long,
white tipped brown. Spines always slightly erect never appressed.
This is one of the distinguishing features of Subsection II from
subsection I Pygmaeae. (see analytical key to the Rebutia section)
35
Flowers very free flowering, 45mm. long and 30mm. wide with orange-red
petals. Flower tube very hairy with occasional bristles. Stigma and
style greenish-yellow, 6-lobed, filaments lavender pink.
Fruit brownish-red, densely covered with white hairs.
Seeds dull brownish-black.
Type locality given by Backeberg as South Bolivia - North Argentina. This
species has recently been recollected by W. Rausch at Quebrada
de Humahuaca Jujuy, Argentina (R214).
Notes this is an easily grown popular species with very showy flowers. It
is very closely related to a number of other species belonging to
subsection II namely R. brachyantha and R. ritteri (see colour
plate).

REBUTIA FIEBRIGII
Fig. 12 This photograph depicts Abbey Brook species No. CF2630 originally
collected at Hacienda Ressini and having pure white spines and orange flowers.
36
 REBUTIA FIEBRIGII
Rebutia fiebrigii (Guerke) Br. & R.
Synonyms Echinocactus fiebrigii Guerke: Notizbl. Bot. Gart. u. Mus. Berlin
(1905)4,183
Echinorebutia fiebrigii (Guerke) Frič & Kreuzgr.
Aylostera fiebrigii (Guerke) Backbg.
Rebutia nivosa Ritter nomen nudum
Rebutia fiebrigii f. densiseta Cullm.
Rebutia fiebrigii var. densiseta (Cullm.) Oeser
Classification Section I Aylostera
Body spherical becoming cylindrical with age, often 60mm. in diameter
and 100- 120mm. high. Older plants offsetting from the base.
Areoles prominent, elliptical, whitish in colour.
Spines 30-40 bristle like radial white spines and 2-5 stiff centrals up to
20mm. or more long, white tipped brown, sometimes pure white.
Flowers 35mm. long and 30mm. broad, flame red with white stamens.
Stigma 6 lobed pale yellow. Ovary and tube scaly with a few white
hairs and bristles.
Seed small, matt black.
Type locality Iscayache, Bolivia 3600m. Map 4, No. 87.
Notes Distinctive species and one of the first to be discovered and
cultivated. Several recently described ‘species’ are probably
referable here e.g. R. jujuyana, R. pulchella.
R. fiebrigii, f. densiseta Cullm. (FR390) a form in which the spines
are thinner and denser, spines tipped yellowish-brown, flowers
orange. Originally collected at Hacienda Hura Khathalla ad
Estancia Chujchi, Bolivia. Abbey Brook species No. CF2630, is a
plant originally collected at Hacienda Ressini, Sucre by Gerhard
Frank with longer spines than in the type, softer and all pure white,
flowers clear orange. Attempts have been made to distinguish
between the snow white populations and others having brown
tipped spines. As Donald has pointed out, the original description
by Guerke suggests an almost wholly white spined plant, but it
could accommodate both white spined and the more commonly seen
brown tipped spined plants.

REBUTIA HELIOSA
Rebutia heliosa Rausch (1970) Kakt. u. a. Sukk. 21 30
Synonyms Aylostera heliosa Rausch comb. nud.
Classification Section I Aylostera
Body solitary at first then offsetting profusely, 20mm. high and 25mm. in
diameter.
Areoles 0-5mm. wide, 1mm. long, light brown, felty.
37
Spines radials 24-26, 1mm. long, flattened against tubercles, diverted
downwards, silvery-white, thickened and darker below. No
central spines.
Flowers 50mm. long and 25mm. in diameter, pale orange outer petals with
purplish mid-stripe. The flowers have a very long thin tube (2mm.
in diameter and 20mm. long) making the flower stand away from
the plant body. Tube purplish, filaments white, stigma 6 lobed and
style yellowish.
Fruit tiny 3mm. in diameter with few seeds, (10-20) in each fruit, greenish
later turning violet. Fruit covered with white hair like bristles.
Seed black, spherical to helmet shaped with a papillate testa.
Type locality Road between Tarija City and Narvaez at 2400-2500m., Bolivia.
Map 4, No. 92.

REBUTIA HELIOSA
Fig. 13 One of the most beautiful, distinctive and unusual Rebutia species.

38
Varieties Rebutia heliosa var. cajasensis Lau 405
Rebutia heliosa var.condorensis Lau 401 (Syn.R.solisioides Knize
nomen nudum)
Rebutia heliosa x Rebutia albiflora. An interesting new hybrid, (see
under R. albiflora)
Notes One of the most beautiful and unusual Rebutia species, very
distinctive. The plant seems to grow well on its own roots but plants
are most often seen grafted which does tend to make the plants
elongated and proliferate.
Several ‘species’ are referable here, namely, supthutiana,
albopectinata, perplexa, schatzliana and densipectinata. For an
explanation of the confusion amongst these species an article in
Ashingtonia (1979) 3 by John Donald helps to sort out the situation.

REBUTIA KRAINZIANA
Fig. 14 This is a superb and distinctive plant with very large flowers.
39
 REBUTIA KRAINZIANA
Rebutia krainziana Kesselring: Sukkulentenkunde (1948) II, 23-24
Synonyms Rebutia calliantha var. densiseta Bew. nomen nudum
Rebutia calliantha var. krainziana (Kess.) Buin. & Donald
Rebutia wessneriana var. densiseta nomen nudum
Rebutia wessneriana var. krainziana (Kess.) Buin. & Donald
Classification Section IV Rebutia Subsection II Mediorebutiae
Body up to 50mm. in diameter and becoming caespitose with age
forming large clumps 100mm. or more in diameter.
Areoles very prominent, ellipitical and with white felt.
Spines pectinate, adpressed, up to 12 radials pure white, centrals absent.
Flowers very large and beautiful, 40mm. long and 40mm. wide. Usually dark
red but there is a form (f. breviseta) with clear orange flowers. 5
lobed stigma, style white, filaments yellow. Self sterile.
Fruit large, 8mm. in diameter with small brownish red scales.
Seeds black shiny, dome-shaped with white basal hylum area. Testa near
hylum reticulate but becoming tuberculate at the apex.
Type locality is unknown. It has recently been recollected at Yavi, Jujuy,
Argentina. Map 3, No. 70
Notes This is a superb and striking plant with very large beautiful flowers.
When not in flower the plant is one of the most attractive species.
R. krainziana f. breviseta with orange flowers may be a hybrid
between R. krainziana and R. marsoneri. The following
combinations also belong with this taxa.
R. krainziana var. breviseta (Backbg.) Donald
R. krainziana f. breviseta (Backbg.) Krainz & Haarm.
R. senilis v. breviseta Backbg.
R. senilis f. breviseta (Backbg.) Backbg.

REBUTIA KUPPERIANA
Rebutia kupperiana Böd.: Monatss. D.K.G. (1932) 276-277
Synonyms Aylostera kupperiana (Böd.) Backbg.
Echinorebutia kupperiana (Böd.) Frič & Krzgr.
Classification Section I Aylostera
Body up to 40mm. in diameter and becoming caespitose with age.
Epidermis very dark brownish-green.
Areoles circular with brownish wool, raised on short tubercles.
Spines strongly differentiated into centrals and radials. Centrals 2, upper
12-15mm. long, lower 20mm. long at first dark chocolate brown in
age becoming greyish with a blackish tip. Radials 10-12, strongly
projecting, 6-12mm. long, somewhat flexuous, white with dark
brown tips.
40
Flowers very large and beautiful 50mm. long and 40mm. in diameter, dark
orange-red. Tube 15mm. long greenish with long bristles. Stigma 6
lobed, filaments and style white.
Fruit greenish, 5mm. in diameter.
Seeds greyish black.
Type locality near Tarija at 2500m. altitude, Bolivia. Map 4, No. 99.
Notes This is yet another magnificent species discovered by Friedrich
Ritter in 1931.
Varieties Rebutia kupperiana var. spiniflora Ritter (FR762b)

REBUTIA KUPPERIANA
Fig. 15 This is yet another magnificent species discovered by Friedrich Ritter. It
has dark brown spines and very large dark orange-red flowers.

41
 REBUTIA MARSONERI
Fig. 16 Another superb species with golden yellow flowers

REBUTIA MARSONERI
Rebutia marsoneri Werdermann: Kakteenkunde (1937) 2
Synonyms Rebutia marsoneri var. albescens hort. nomen nudum
Rebutia marsoneri var. breviseta hort. nomen nudum
Rebutia marsoneri var. brevispina Donald nomen nudum
Rebutia marsoneri var. grandiflora Donald nomen nudum
Rebutia marsoneri var. spathulata Donald nomen nudum
Rebutia marsoneri f. spathulata Donald nomen nudum
Rebutia marsoneri var. vatteri Donald nomen nudum
Rebutia peruviana hort. nomen nudum
Classification Section IV Rebutia Subsection II Mediorebutiae
Body up to 70mm. in diameter becoming caespitose with age.
42
Areoles ellipitical with dense white wool.
Spines very variable in size and number, c. 10 undifferentiated into
centrals and radials, up to 15mm. long, whitish or tinged yellow. In f.
brevispina all the spines are pectinate, white and less than 5mm.
long.
Flowers large, 40mm. long and 35mm. in diameter. Usually golden yellow
but sometimes lighter yellow, sometimes orange-yellow. Stigma
5-10 lobed, style and filaments golden yellow. The buds
are variable in colour sometimes red and sometimes strikingly
bright green.
Fruit large, 8mm. in diameter.
Seeds black, shiny, dome-shaped with white basal hylum area.
Type locality North Jujuy, Argentina. Map 3, No. 71.
Notes A superb species named after Oreste Marsoner, an Italian who has
collected extensively in Argentina. This species is relatively rare
in cultivation because it is self sterile and seed is not as readily
available as self fertile species. There are a number of varieties
and forms most of which are insufficiently different to warrant
separate classification, (see colour plate).

REBUTIA MINUSCULA
Fig. 17 This is one of the easiest and most widely grown of all species of Cacti.
43
 REBUTIA MINUSCULA
Rebutia minuscula K. Sch.: Monats. f. Kakt. (1895) 5, 102
Synonyms Echinopsis minuscula Web.
Echinocactus minuscula Web.
Rebutia minuscula v. multiflora Frič nomen nudum
Rebutia petersonii Hort. nomen nudum
Classification Section IV Rebutia Subsection I Rebutiae
Body up to 50mm. in diameter and becoming strongly caespitose with
age soon forming large clumps 150mm. or more in diameter.
Epidermis deep green.
Areoles circular with sparse white wool.
Spines numerous, c. 40, undifferentiated into radials and centrals and
arranged like needles on a pin cushion. Spines up to 5mm. long,
whitish, sometimes becoming brownish at the base.
Flowers large, up to 40mm. long and 35mm. in diameter, bright red. 6 lobed
stigma, style pinkish, filaments golden yellow.
Fruit 6mm. in diameter, orange-red.
Seeds black, shining.
Type locality Tucuman, N. Argentina. Map 3, No. 61.
Notes One of the easiest and most widely grown of all species of Cacti. It
can be flowered when only 1 year old from seed. It was the first
species of Rebutia to be described.
There is a form in cultivation R. minuscula f. grandiflora which has a
very elongated flower tube making the flower 70mm. long.
Consult the section on flowers and floral structure.

REBUTIA MUSCULA
Rebutia muscula Ritter & Thiele: Taxon (1963) 12,29
Synonyms Aylostera muscula (Ritter & Thiele) Backbg.
Rebutia nivosa Ritter nomen nudum
Rebutia muscula v. nivosa Knize comb. nud.
Classification Section I Aylostera
Body up to 5cm. in diameter becoming caespitose with age, individual
heads becoming elongated.
Areoles circular with white wool.
Spines pure white, soft bristle like, centrals indistinguishable from the
radials. 6mm. long, 50-55 in number.
44
Flowers 40mm. long and 25mm. in diameter, sepals greenish-orange, petals
flame orange, throat white. Stigma very pale green, style white,
filaments white. Flower tube and ovary with very small scales. Very
thin flexuous white hairs up to 8mm. long produced from the axils of
the scales.
Seeds small, dull black, dome shaped.
Type locality Narvaez, Dept. Tarija, Bolivia. Map 4, No. 103.
Notes Beautiful pure white spined species with gorgeous clear orange
flowers. Flowers are often produced at odd times during the year
but the main flowering time is at the end of May beginning of June.
Plants labelled as R. nivosa are referable here, there is no distinct
difference and it was found at the same locality.
(see illustration on front cover)

REBUTIA PSEUDODEMINUTA
Rebutia pseudodeminuta Backbg. non Frič (1933) D.K.F. 7
Synonyms Aylostera pseudodeminuta (Backbg.) Backbg.
Classification Section I Aylostera
Body globular up to 10cm. high, 5-6cm. wide, light green in colour,
offsetting grousely at first but later forming large clumps 100mm. or
more in diameter.
Areoles small with sparse light brown wool.
Spines radials ten or more, 3-7mm. long, glossy white. Centrals 2-3,
yellowish-white at first later turning brown-tipped, up to 15mm.
long.
Flowers 30mm. long and 30mm. wide, petals red, buds dark red. Style and
filaments white united with the tube, 5 lobed stigma.
Fruit round, small with many bristles.
Seed very small, matt black.
Habitat Salta, N. Argentina. Map 4, No. 107.
Varieties f. albiseta (Backbg.) Buin. & Donald. Central spines entirely white.
f. grandiflora (Backbg.) Buin. & Donald. Flower 4cm. wide and
40mm. long.
f. schneideriana (Backbg.) Buin. & Donald. Central spines up to
35mm. long, dark tipped.
f. schumanniana (Backbg.) Buin. & Donald. Central spines golden
brown, about 12mm. long.
Notes Another of the ‘original’ Rebutias, quite distinct with its prominent
tubercles and spination.

45
 REBUTIA PYGMAEA
Fig. 18 A distinctive species with pectinate spines. It has very large flowers when
compared to the size of the plants.

REBUTIA PYGMAEA
Rebutia pygmaea (R. E. Fries) Br. & R.
Synonyms Echinopsis pygmaea R. E. Fries Nov. Act. Soc. Sci. Upsala (1905) IV
120
Acantholobivia haagei (Frič & Schelle) Y. Ito.
Digitorebutia digitiformis (Backbg.) Buin.
Digitorebutia haagei (Frič & Schelle) Frič
Digitorebutia haagei var. digitiformis (Backbg.) Donald
Digitorebutia haagei var. orurensis (Backbg.) Donald
Digitorebutia haagei var. pectinata (Backbg.) Donald
Digitorebutia orurensis (Backbg.) Buin.
46
 Digitorebutia pectinata (Backbg.) Buin.
Digitorebutia pygmaea (R. E. Fries) Donald
Digitorebutia steinmannii sensu Buin.
Echinolobivia haagei (Frič & Schelle) Y. Ito.
Lobivia neo-haageana Backbg.
Lobivia pygmaea (R. E. Fries) Backbg.
Mediolobivia haagei (Frič & Schelle) Backbg.
Mediolobivia haagei var. digitiformis (Backbg.) Donald
Mediolobivia haagei var. orurensis (Backbg.) Donald
Mediolobivia haagei var. pectinata (Backbg.) Donald
Mediolobivia haageana Borg nomen nudum
Mediolobivia orurensis (Backbg.) Backbg.
Mediolobivia pygmaea (R. E. Fries) Backbg.
Mediolobivia pectinata (Backbg.) Backbg. non Frič
Mediolobivia pectinata var. digitiformis (Backbg.) Backbg.
Mediolobivia pectinata var. orurensis (Backbg.) Backbg.
Mediolobivia pectinata var. neosteinmannii Backbg.
Pygmaeolobivia odierata Hort. nomen nudum
Pygmaeolobivia pygmaea (R. E. Fries)
Rebutia haagei Frič & Schelle
Rebutia orurensis Backbg.
Rebutia pectinata Backbg.
Rebutia pectinata var. digitiformis Backbg.
Rebutia pectinata var. orurensis Backbg.
Classification Section III Digitorebutia Subsection I Pygmaeae
Body up to 25mm. in diameter, simple at first but becoming caespitose
with age. The epidermis bluish-green often tinged deep purple
particularly during the winter.
Areoles elliptical with sparse greyish-brown wool, raised in the tubercles.
Spines arranged in 12 distinct ribs. Centrals absent, radials strongly
pectinate and adpressed, up to 12 in number, 4mm. long, greyish-
white, brownish at the base.
Flowers very variable, pink to salmon often bicoloured, 25mm. long and
20mm. in diameter with a white throat. Stigma 6 lobed green, style
green, filaments white or pinkish. Tube brownish-green with
green scales with copious white wool.
Fruit dull brownish-black, dome shaped, testa finely pitted.
Seed large, 7mm. in diameter, greenish-brown covered with white wool.
Type locality Yavi, Jujuy Province, N. Argentina. Map 2, No. 35.
Notes This species has a very wide geographical range and a large
number of minor variations occur. Many of these minor variations
have been described as ‘new’ species or varieties. There are for
example a very large number of flower colour variants and also
spine colour variants. It is indeed possible that this species could
include 90% of all the species ascribed to the subsection
Pygmaeae!

47
 REBUTIA SENILIS f. STUEMERI
Rebutia senilis f. stuemeri (Backbg.) Buin. & Donald
Synonyms Rebutia senilis v. stuemeri Backbg. D. Kakt. frd. (1932) 131
Rebutia senilis v. stuemeriana Backbg.
Classification Section IV Rebutia Subsection I Rebutiae
Body up to 60mm. in diameter and becoming caespitose with age.
Areoles elliptical with white wool.
Spines pure white, stiff bristle like, centrals hardly distinguishable from
the radials. 15-30 in number, size is variable - at the top of the
plant spines up to 18mm. long. Older areoles at the base of the plant
produce longer spines up to 25mm. long.
Flowers 50mm. long and 40mm. in diameter, clear red with a yellowish
throat. Stigma 5 lobed, style and filaments golden yellow. Self
fertile.
Fruit yellowish, 7mm. in diameter.
Seed black, shiny, elongated dome-shaped with a white basal hylum
area. Testa near hylum more or less reticulate but becoming
tuberculate at the apex.
Type locality Quebrada Escoipe, Salta, Argentina. Map 3, No. 66.
Notes R. almeyeri W. Heinr. probably belongs here. In Backbg. (1977) it is
described as a self fertile species with orange-red flowers with a
yellowish throat and a yellow style, locality unknown.

REBUTIA SPINOSISSIMA
Rebutia spinosissima Backbg. (1935) Kaktus ABC., 275
Synonyms Aylostera spinosissima (Backbg.) Backbg.
Aylostera spinosissima var. brunispina Backbg. nomen nudum
Classification Section I Aylostera
Body up to 35mm. in diameter becoming caespitose with age eventually
forming large clumps.
Areoles circular to elliptical with greyish wool.
Spines soft and bristle like, about 20, not strongly differentiated into
centrals and radials. ‘Radial’ spines up to 6mm. long, all white,
‘centrals’ at first yellow tipped brown later all brown.
Flowers 40mm. long and 30mm. in diameter. Petals orange-red, tube
greenish, 8 lobed stigma, filaments and style white.
Fruit greenish brown 4mm. in diameter with white hairs and bristles.
Seeds elongated dome shaped, dull brownish black.
Type locality Salta Province, N. Argentina. Map 4, No. 116.
Notes This is a beautiful species with soft bristle like spines and orange
flowers. It is slow growing and an asset to any Rebutia collection.
Closely related to R. muscula and R. fiebrigii

48
 REBUTIA SPINOSISSIMA
Fig. 19 A slow growing species with soft bristle-like spines and orange flowers.

REBUTIA WESSNERIANA
Rebutia wessneriana Bew. Sukkulentenkunde Jahrb. Schweiz.
Kakt. Ges. (1948)2 24
Synonyms Rebutia calliantha Bew.
Rebutia hyalacantha (Backbg.) Backbg.
Rebutia senilis var. hyalacantha Backbg.
Rebutia senilis f. hyalacantha (Backbg.) ex Simon
Rebutia wessneriana var. calliantha (Bew.) Krainz
Rebutia wessneriana var. permutata (Heinr.) Buin. & Donald
Rebutia wessneriana subspec. wessneriana f. calliantha (Bew.)
Buin. & Donald
49
 REBUTIA WESSNERIANA
Fig. 20 A superb large growing, large flowering species

Rebutia wessneriana subspec. wessneriana var. gokrausei (Heinr.)

Donald
Rebutia wessneriana subspec. wessneriana var. gokrausei f.
permutata (Heinr.) Donald
Classification Section IV Rebutia Subsection II Mediorebutiae
Body large, up to 100m. in diameter, caespitose and forming large
clumps with age
Areoles elliptical raised up on tubercles with white wool
Spines not differentiated into centrals

50
Flowers very large, 45mm. long and 40mm. in diameter. Petals dark red,
buds dark purplish-red. Stigma 6 lobed, style yellowish-white,
filaments orange. Self sterile.
Fruit large, 8mm. in diameter, yellowish
Seed black, shiny, dome shaped with white basal hylum area.
Type locality Quebrada de Humahuaca, Jujuy, Arg.? Map 3, No. 74.
Notes A superb large growing large flowered species which is
uncommon in collections because the plant is self sterile and seed
is therefore not readily available. Originally R. wessneriana was
separated from R. calliantha by flower colour, scarlet in calliantha
and bluish-red in wessneriana, and on the observation that the
spines cover the stem apex in calliantha but did not hide the stem
apex in wessneriana. Such minor differences are not sustained and
are only observed in extreme plants and are not exhibited by the
population as a whole. (see colour plate).

REBUTIA XANTHOCARPA f. SALMONEA

Rebutia xanthocarpa f. salmonea (Frič ex Backbg.) Buin. & Donald
Synonyms Rebutia xanthocarpa var. salmonea Frič ex Backbg. (1951) Cact.
Succ. J. Amer. 23 83
Rebutia salmonea Frič non Backbg. nomen nudum
Rebutia senilis var. salmonea (Backbg.) Marsh.
Classification Section IV Rebutia Subsection I Rebutiae
Body 40mm. in diameter, very slowly becoming caespitose with age.
Areoles elliptical with greyish brown felt.
Spines not differentiated into centrals and radials, c. 15. Short stiff and
bristle like, 2mm. long occasionally 4mm., white tinged yellow.
Flowers small, 15mm. long and 15mm. in diameter. A beautiful salmon pink.
Tube yellow with greenish yellow scales. Stigma 6 lobed, yellowish
white. Style and filaments white.
Fruit large 7mm. in diameter, golden yellow, (not pale red, Backbg.
Lexicon P. 439)
Seeds small, black shining with white basal hylum area.
Habitat unknown, but probably Salta area, Argentina. Map 3, No. 67.
Notes An easily grown and flowered plant with a very characteristic and
distinctive flower colour. (see colour plate).

51
 Check list of Genera, Subgenera and Sections
The following is a list of the more important genera to which members of the
genus Rebutia have been assigned. There are in addition to the following list a
number of additional generic names which have not been published in accordance
with the code of Botanical Nomenclature and which are probably best forgotten. The
correct Rebutia section for the plants which were originally placed under these
names is indicated in brackets after the name. It follows the taxonomic arrangement
used in this booklet. The original publication and date is given for the most important
of these generic names.

Acantholobivia Y. Ito. non Backbg. (Rebutia Section III) (1957) Expl. Diagr. 130
Aylostera Speg. (Rebutia Section I) (1923) An. Soc. Cient. Argent. 96 75
Cylindrorebutia Frič & Kreuzgr. nomen nudum (Rebutia Section II) (1938) Succulenta
20 55 & 71
Cylindrorebutia Buin. & Donald pro. sect. (Rebutia Section II) (1963)
Sukkulentenkunde Jb. Schweiz Kakt. Ges. 8 98
Digitorebutia Frič & Kreuzgr. nomen nudum (Rebutia Section III) (1938) Succulenta
20 54
Digitorebutia Frič & Kreuzgr. ex. Buin. (Rebutia Section III) (1940) Succulenta 22 51
Digitorebutia Buin. & Donald pro. sect. (Rebutia Section III) (1963) Sukkulenten-
kunde Jb. Schweiz Kakt. Ges. 8 98
Echinocactus Link & Otto (Rebutia Section 1, 11 and IV) (1827) Verh. Ver. Beford.
Gartenb. 3 420
Echinocactus Weber, ex K. Sch. (Rebutia Section 1,11 and IV) (1898) Gesamtbeschr.
Kakt. 396
Echinolobivia Y. Ito. (Rebutia Section III) (1957) Expl. Diagr.
Echinopsis Zucc. (Rebutia Section 1, 11 and IV) (1837) Abh. Bayer. Akad. Wiss.
München 2 675
Echinorebutia Frič & Kreuzgr. nomen nudum (Rebutia Section I) (1935) Verzeichnis
26
Lobivia Br. & R. (Rebutia Section 11 and III) (1922) The Cactaceae III 49
Lobivia subgen. Pygmaeolobivia Backbg. nomen nudum (Rebutia Section II) (1934)
Blatt. f. Kakteenf.
Mediolobivia Backbg. (Rebutia Section I, II and III) (1934) Blatt. f. Kakteenf.
Mediolobivia subgen. Eumediolobivia Backbg. (Rebutia Section II) (1942) Cact. Jb.
D.K.G. 1135
Mediolobivia subgen. Pygmaeolobivia Backbg. (Rebutia Section III) (1941) Cact. Jb.
D.K.G.
Mediorebutia hort. nomen nudum (Rebutia Section IV)
Mediorebutia Buin. & Donald pro, sect. (Rebutia Section IV)
Rebulobivia Frič & Kreuzgr. (Rebutia Section II) (1936) Succulenta 18 104
Rebutia K. Sch. (1895) Monatsschr. f. Kakteenfreunde 5 102
Rebutia K. Sch. emend Buin. & Donald (1963) Sukkulentenkunde 7 96
Rebutia subgen. Eurebutia Backbg. non Frič. (Rebutia Section IV) (1934) Blatt. f.
Kakteenf.
Rebutia subgen. Neorebutia Bew. (Rebutia Section IV) (1949) Sukkulentenkunde Jb.
Schweiz Kakt. Ges. 3 54
Setirebutia Frič & Kreuzgr. nomen nudum (Rebutia Section II) (1935) Verzeichnis 26
52
Setirebutia Frič & Kreuzgr. (Rebutia Section II) (1938) Succulenta 20 53
Setirebutia Buin. & Donald pro. sect. (Rebutia Section II) (1963) Sukkulentenkunde Jb.
Schweiz Kakt. Ges. 7 98

Check list of Genus Aylostera Speg.

albiareolata Ritter comb. nud. = R. albiareolata
albiflora (Ritt. & Buin.) Backbg. = R. albiflora
albipilosa (Ritt.) Backbg. = R. albipilosa
albopectinata Rausch comb. nud. = R. albopectinata
buiningiana Rausch comb, nud. = R. buiningiana
buiningiana Ritter nomen nudum (not R. buiningiana Rausch) = R. spinosissima
caespitosa Frič nomen nudum = ?
camarguensis Rausch comb. nud. = R. camarguensis
deminuta = R. deminuta
deminuta var. pseudominuscula = R. deminuta f. pseudominuscula
densipectinata Ritter comb. nud. = R. densipectinata
diersiana Rausch comb. nud. = R. diersiana
fiebrigii (Guerke) Backbg. = R. fiebrigii
fiebrigii var. castanea Rausch nomen nudum= R. fiebrigii
fiebrigii var. densiseta Cullm. = R. fiebrigii f. densiseta
fiebrigii var. spinosior nomen nudum = R. fiebrigii
flavistyla Ritter comb. nud. = R. flavistyla
fulviseta Rausch comb. nud. = R. fulviseta
fulviseta var. albispina Rausch nomen nudum
graciliflora (Backbg.) Rausch comb. nud. = R. graciliflora
grandiflora Frič nomen nudum = ? R. pseudodeminuta f. grandiflora
heliosa Rausch nomen comb. = R. heliosa
hoffmanii Diers. et Rausch comb. nud. = R. hoffmannii
huasiensis Rausch comb. nud. = R. huasiensis
ithyacantha Card. = R. ithyacantha
jujuyana Rausch comb. nud. = R. jujuyana
kieslingii Rausch comb. nud. = R. kieslingii
kruegerii Card. = Sulcorebutia kruegerii
kupperiana (Böd.) Backbg. = R. kupperiana
lampromeliana Rausch nomen nudum = R. pulchella
leucanthema Rausch comb. nud. = R. leucanthema
mamillosa Rausch comb. nud. = R. mamillosa
maxima Ritter nomen nudum = ?
muscula (Ritt. & Thiele) Backbg. = R. muscula
narvaecense Card. = R. narvaecense
odontopetala Ritter comb. nud. = R. odontopetala
padcayensis Rausch comb. nud. = R. padcayensis
patericalyx Ritter comb. nud. = R. patericalyx
pectinata Frič nomen nudum = R. pygmaea
pseudodeminuta (Backbg.) Backbg. = R. pseudodeminuta
pseudodeminuta var. albiseta Backbg. = R. pseudodeminuta f. albiseta
pseudodeminuta var. grandiflora Backbg. = R. pseudodeminuta f. grandiflora
53
pseudodeminuta f. rubrifilamentosa Buin. & Donald comb. nud. =
R. pseudodeminuta f. rubrifilamentosa
pseudodeminuta var. schneideriana Backbg. = R. pseudodeminuta f. schneideriana
pseudodeminuta var. schumanniana (Backbg.) Backbg. =
R. pseudodeminuta f. schumanniana
pseudominuscula (Speg.) Speg. = R. deminuta f. pseudominuscula
pulchella Rausch comb. nud. = R. pulchella
pulvinosa (Ritt. & Buin.) Backbg. = R. pulvinosa
robustiflora Frič = ?
robustispina (Ritt.) Backbg. = R. robustispina
rubiginosa (Ritt.) Backbg. = R. rubiginosa
sanguinea Ritter comb. nud. = R. sanguinea
schatzliana Rausch comb. nud. = R. schatzliana
sordida Rausch nomen nudum. = ?
spegazziniana (Backbg.) Backbg. = R. spegazziniana
spegazziniana var. atroviridis Backbg. = R. spegazziniana var. atroviridis
spegazziniana var. waltheriana Backbg. nomen nudum = R. spegazziniana
spinosissima (Backbg.) Backbg. = R. spinosissima
spinosissima var. aurea Rausch nomen nudum = R. spinosissima
spinosissima var. brunispina = R. spinosissima
spinosissima var. buiningiana Ritter nomen nudum = R. spinosissima
steinmannii (Solms-Laub.) Backbg. = R. steinmannii
[not R. pygmaea (R. E. Fries) Br. & R.!]
supthutiana Rausch comb. nud. = R. supthutiana
tarijensis Rausch comb. nud. = R. tarijensis
tuberosa (Ritter) Backbg. = R. tuberosa
umbraculiforma Rausch nomen nudum = R. camarguensis
vallegrandensis Card. = R. vallegrandensis
waltheriana (Backbg.) Y. Ito. = ? R. spegazziniana
zavaletae Card. = Sulcorebutia zavaletae (Card.) Backbg.

Check list of Genus Digitorebutia Frič. Kreuzgr. Buin.

albopectinata Rausch comb. nud. = R. albopectinata
atrovirens (Backbg.) = R. pygmaea var. atrovirens
brachyantha (Wessn.) Buin. = R. brachyantha
brachyantha var. ritteri (Wessn.) Donald = R. ritteri
brunescens Rausch comb. nud. = R. brunescens
canacruzensis Rausch comb. nud. = R. canacruzensis
christinae Rausch comb. nud. = R. christinae
cincinnata Rausch comb. nud. = R. cincinnata
costata (Werd.) Buin. = R. costata
costata var. eucalyptana (Backbg.) Donald = R. costata f. eucalyptana
diersiana Rausch comb. nud. = R. diersiana
diersiana var. atrovirens Rausch comb. nud. = R. diersiana v. atrovirens
digitiformis (Backbg.) Buin. = R. pygmaea
donaldiana Rausch nomen nudum = R. steinmannii v. cincinnata
Rausch nomen nudum (R300)

54
eucalyptana (Backbg.) Buin. = R. costata f. eucalyptana
euanthema (Backbg.) Buin. = R. euanthema
euanthema var. friciana Donald & Cullm. comb. nud. = R. euanthema f. fricii
euanthema var. longispina (Backbg.) Donald comb. nud. = R. euanthema
euanthema var. oculata (Werd.) Donald = R. euanthema f. oculata
haagei (Frič & Schelle) Frič = R. pygmaea
haagei var. digitiformis (Backbg.) Donald = R. pygmaea
haagei var. atrovirens (Backbg.) Donald = R. pygmaea v. atrovirens
haagei var. orurensis (Backbg.) Donald = R. pygmaea
haagei var. pectinata (Backbg.) Donald = R. pygmaea
haefneriana (Cullm.) Donald = R. pygmaea f. haefneriana
mudanensis Rausch comb. nud. = R. mudanensis
nazarenoensis Rausch (1979) Succulenta 58 185.
nigricans (Wessn.) Buin. = R. ritteri var. nigricans
orurensis (Backbg.) Buin. = R. pygmaea
oculata (Werd.) Buin. = R. euanthema f. oculata
oculata var. friciana Donald & Cullm. = R. euanthema f. fricii
pectinata (Backbg.) Buin. = R. pygmaea
peterseimii (Frič & Kreuzgr.) Donald comb. nud. =
R. ritteri var. nigricans f. peterseimii
pygmaea (R. E. Fries) Donald = R. pygmaea
rauschii Zecher comb. nud. = R. rauschii
ritteri (Wessn.) Buin. = R. ritteri
steinmannii sensu Buin. = R. pygmaea
c.v. ‘A. V. Frič’ Donald & Cullm. = R. euanthema f. fricii

Check list of Genus Mediolobivia Backbg.

albilongiseta nomen nudum = R. aureiflora
albinidulans Frič nomen nudum = ?
albiscoparia Frič nomen nudum = ?
albiseta Frič nomen nudum = R. aureiflora
albiseta var. aureiflora Frič nomen nudum = R. aureiflora
albiseta var. rubriflora Frič nomen nudum = R. aureiflora f. rubriflora
albispina Frič nomen nudum = ? R. aureiflora
albopectinata Rausch = R. albopectinata
antarctica Hort. nomen nudum = ?
apricusioides Frič nomen nudum = ?
auranitida (Wessn.) Krainz = R. auranitida
auranitida var. flaviflora Backbg. = R. auranitida
auranitida var. gracilis (Wessn.) Backbg. = R. auranitida f. gracilis
aureiflora (Backbg.) Backbg. = R. aureiflora
aureiflora var. albiseta Backbg. = R. aureiflora
aureiflora var. albi-longiseta Backbg. = R. aureiflora
aureiflora var. albispina Backbg. nomen nudum = R. aureiflora
aureiflora var. boedeckeriana (Backbg.) Backbg. = R. aureiflora f. boedeckeriana
aureiflora var. brevispina Backbg. nomen nudum = R. aureiflora

55
aureiflora var. duursmaiana (Backbg.) Backbg. = R. aureiflora f. duursmaiana
aureiflora var. haertlingiana hort. nomen nudum = R. aureiflora f. rubriflora
aureiflora var. leucolutea Backbg. nomen nudum = R. aureiflora
aureiflora var. lilacinostoma Backbg. nomen nudum = R. aureiflora
aureiflora var. longiseta Backbg. nomen nudum = R. aureiflora
aureiflora var. longispina Wenzel nomen nudum = R. aureiflora
aureiflora var. rubelliflora (Backbg.) Backbg. = R. aureiflora f. rubelliflora
aureiflora var. robustior Backbg. nomen nudum = R. aureiflora
aureiflora var. rubriflora (Backbg.) Backbg. = R. aureiflora f. rubriflora
aureiflora var. rubriflora f. kesselringiana (Cullm.) Köhler) = R. aureiflora f. rubriflora
aureiflora var. sarothroides (Werd.) Backbg. = R. aureiflora f. sarothroides
aureiflora var. variegata Backbg. nomen nudum = R. aureiflora
aureiflora subvar. leucolutea (Backbg.) Backbg. = R. aureiflora
aureiflora subvar. lilacinostoma (Backbg.) Backbg. = R. aureiflora
atrovirens (Backbg.) Backbg. = R. pygmaea f. atrovirens
atroviridis hort. (catalogue 1974 Uhlig) U1396 = ?
blossfeldii (Werd.) Buin. = R. aureiflora f. rubriflora
blossfeldii var. campactiflora Wessn. = R. aureiflora f. rubriflora
blossfeldii var. nigrilongiseta Wessn. = R. aureiflora f. rubriflora
boedeckeriana Backbg. = R. aureiflora f. boedeckeriana
brachyantha (Wessn.) Krainz = R. brachyantha
breviseta Frič nomen nudum = ?
brunescens (Rausch) Backbg. comb. nud. = R. brunescens
bruneoradicata Ritter comb. nud. = R. bruneoradicata
brunicentra Frič nomen nudum = ?
brunicentra var. roseiflora Frič nomen nudum = ?
bruniscoparia Frič nomen nudum = ?
bruniscoparia var. chata Frič nomen nudum = ?
bulbispina hort. (catalogue 1974 Uhlig) U2250 = ?
cachiensis Frič nomen nudum = ?
cajasensis (Ritter) Backbg. comb. nud. = R. cajasensis
calenduliflora Frič nomen nudum = ?
calva Frič nomen nudum = ?
calva var. similis Frič nomen nudum = ?
carminata Backbg. ex Krainz nomen nudum = R. aureiflora f. rubriflora
citriaurata Frič nomen nudum = ?
columnaris (Wessn.) Krainz. = R. einsteinii f. columnaris
columnaris Frič nomen nudum = ? R. einsteinii f. columnaris
conoidea (Wessn.) Krainz. = R. einsteinii f. conoidea
conoidea var. columnaris (Wessn.) Backbg. = R. einsteinii f. columnaris
costata (Werd.) Krainz = R. costata
costata var. brachyantha (Wessn.) Donald nomen nudum = R. brachyantha
costata var. eucalyptana (Backbg.) Donald comb. nud. = R. costata f. eucalyptana
cupreata Frič nomen nudum = ?
diersiana Rausch comb. nud. = R. diersiana
digitiformis (Backbg.) Backbg. = R. pygmaea
disciformis Frič nomen nudum = ?
duursmaiana Backbg. = R. aureiflora f. duursmaiana

56
einsteinii = R. einsteinii
einsteinii var. atrovirens = ? R. einsteinii f. rubriviridis
elegans Backbg. = R. aureiflora v. elegans
elegans var. gracilis Backbg. nomen nudum = R. aureiflora v. elegans
eos Rausch comb. nud. = R. eos
eos var. roseiflora Rausch comb. nud. = R. eos
erythrantha Backbg. nomen nudum = R. aureiflora f. rubriflora
escayachensis Knize nomen nudum KK1686
euanthema (Backbg.) Krainz = R. euanthema
euanthema var. fricii (Hort.) Backbg. = R. euanthema f. fricii
euanthema var. pygmaea (Backbg.) Donald comb. nud. =
R. euanthema f. neopygmaea
euanthema var. ritteri (Wessn.) Donald comb. nud. = R. ritteri
eucalyptana (Backbg.) Krainz = R. costata f. eucalyptana
fricii hort. = R. euanthema f. fricii
friedrichiana Rausch comb. nud. = R. friedrichiana
fuauxiana Backbg. = R. fuauxiana
grandiflora Frič nomen nudum = ?
haageana Borg nomen nudum = ? R. pygmaea
haagei (Frič & Schelle) Backbg. = R. pygmaea
haagei var. atrovirens (Backbg.) Donald comb. nud. = R. pygmaea f. atrovirens
haagei var. chamaeleon Frič nomen nudum = R. pygmaea f. flavovirens
haagei var. digitiformis (Backbg.) Donald comb. nud. = R. pygmaea
haagei var. flavovirens (Backbg.) Backbg. = R. pygmaea f. flavovirens
haagei var. flavovirens Frič nomen nudum = R. pygmaea f. flavovirens
haagei var. orurensis (Backbg.) Donald comb. nud. = R. pygmaea
haagei var. pectinata (Backbg.) Donald comb. nud. = R. pygmaea
haagei var. salmonea Frič nomen nudum = R. pygmaea f. flavovirens
haagei var. tricolor Frič nomen nudum = R. pygmaea f. flavovirens
haefneriana Cullm. = R. pygmaea var. haefneriana
hartlingiana Borg = ?
hirsutissima Card. = R. hirsutissima
iscayachensis Rausch comb. nud. = R. iscayachensis
ithyacantha Card. = R. ithyacantha
kesselringiana Cullm. = R. aureiflora f. rubriflora
longiflora Frič nomen nudum = ?
longiseta Backbg. nomen nudum = R. aureiflora
mammillata Frič nomen nudum = ?
mammilosa = ?
melanotricha Frič nomen nudum = ?
miniflora Frič nomen nudum = ?
minimiflora Frič nomen nudum = ?
morenoensis Rausch nomen nudum = ?
mudanensis Rausch comb. nud. = R. mudanensis
multiflora Frič nomen nudum = ?
neopygmaea Backbg. = R. euanthema f. neopygmaea
neosteinmannii (catalogue 1974 Uhlig) U1845 = R. pygmaea v. neosteinmannii
nidulans Frič = ?

57
nigra Frič nomen nudum = ?
nigricans (Wessn.) Krainz. = R. ritteri var. nigricans
nigricans var. peterseimii Frič = R. ritteri f. peterseimii
nigriticentra Frič nomen nudum = ?
orurensis (Backbg.) Backbg. = R. pygmaea
padcayensis Rausch comb. nud. = R. pygmaea
patericalyx Ritter comb. nud. = R. patericalyx
patericalyx var. odontopetala Ritter comb. nud. = R. patericalyx
paucipetala Frič nomen nudum = ?
pectinata (Backbg.) Backbg. non Frič = R. pygmaea
pectinata var. atrovirens (Backbg.) Backbg. = R. pygmaea v. atrovirens
pectinata var. digitiformis (Backbg.) Backbg. = R. pygmaea
pectinata var. gracilis Knize nomen nudum (KK970)
pectinata var. neosteinmannii Backbg. = R. pygmaea
pectinata var. paznaensis Knize nomen nudum (KK969)
pectinata var. orurensis (Backbg.) Backbg. = R. pygmaea
permutata Knize nomen nudum KK1046
pygmaea (R. E. Fries) Krainz = R. pygmaea
pygmaea var. flavovirens (Backbg.) Backbg. = R. pygmaea f. flavovirens
pygmaea var. longispina nomen nudum
puchella Frič nomen nudum = ?
pumilla Frič nomen nudum = ?
pyrimidalis Frič nomen nudum = ?
rauschii Zecher comb. nud. = R. rauschii
reichii Borg = Neoporteria reichii
raulii Rausch comb. nud. = R. raulii
ritteri (Wessn.) Krainz = R. ritteri
ritteri var. pilifera (Frič) Backbg. = R. ritteri
robusta Hort. nomen nudum = ?
rosalbiflora Ritter comb. nud. = R. rosalbiflora
roseiaurata Frič nomen nudum = ?
rubelliflora Backbg. = reddish-orange flowering plant, may be a natural hybrid
between the red flowered R.aureiflora f.rubriflora and the yellow flowered
R. aureiflora = R. aureiflora f. rubelliflora
rubriflora Backbg. = R. aureiflora f. rubriflora
rubriflora var. blossfeldii (Werd.) Krainz = R. aureiflora f. rubriflora
rubriflora var. nigrilongiseta (Wessn.) Krainz = R. aureiflora f. rubriflora
sarothroides Werd. = R. aureiflora f. sarothroides
schmiedcheniana (Kohl.) Krainz. = R. einsteinii f. schmiedcheniana
schmiedcheniana var. einsteinii (Frič) Backbg. = R. einsteinii
schmiedcheniana var. karreri (Frič) Backbg. = R. einsteinii
schmiedcheniana var. rubriviridis Frič & Kreuzgr. ex Backbg. =
R. einsteinii f. rubriviridis
schmiedcheniana var. steineckei Frič & Kreuzgr. ex Backbg. =
R. einsteinii f. steineckei
scoparia Frič nomen nudum = ?
scoparia var. cylindrica Frič nomen nudum = ?
semicolumnaris Frič nomen nudum = ?
spiralisepala Schutz = cultivated form of R. aureiflora

58
spiralisepala Jajo ex Brederoo nomen nudum
steinmannii (Solms-Laub.) Krainz = R. steinmannii
steinmannii var. complanata Rausch nomen nudum
steinmannii var. rotundiseta Rausch nomen nudum
steinmannii var. rubriflora Rausch nomen nudum
stellata Frič nomen nudum = ?
struwelpeter Frič nomen nudum = ?
sulcomammillata Frič nomen nudum = ?
tarvitaensis Ritter comb. nud. = R. tarvitaensis
tarvitensis = R. tarvitaensis
turbiniformis Frič nomen nudum = ?
uniclaricentra Frič nomen nudum = ?
uninigricentra Frič nomen nudum = ?
waltheriana Borg = R. spegazziniana

Check list of Rebutia species

The following is a list of specific and varietal names which have been used in
connection with the genus Rebutia. A large number of these names are invalid,
unfortunately many of them are still in use and have therefore been included. Please
realise that this is not a complete check list as we are sure there are many other
names and combinations which have appeared in obscure publications and which
are not included. Nursery catalogues are a fruitful source of misnames and invalid
combinations. As it is, the list already contains nearly 800 names and combinations.
We do not pretend to know what are all the correct current names for many of
the names in the following list for a variety of reasons.
a) often living plants of many of the named ‘species’ are not in cultivation and in
addition were sometimes inadequately described without photographs. This
means that accurate assessment as to status or indeed relationship to a
currently accepted name is extremely difficult and often impossible.
b) names have sometimes been given to plants which after further field work
appear to have been described from extreme geographical forms. They have
never been recollected and probably never will be again (e.g. R. albipilosa)
c) many of the names are names in the literature described without full
description and so referal to the correct name is often impossible.
d) a number of the names have not been published in accordance with the code of
Botanical Nomenclature and have therefore to be excluded.
The complex and superfluous nature of the names in this genus is admirably
illustrated by the example of Rebutia senilis, where there are no less than 50 possible
combinations.
I Starred * names have accurate locational data and have been used in the
compilation of the distribution maps.
II The accepted species together with the names of plants which are reasonably
distinct and identifiable are set in bold type.
III All other names are set in italic type.

59
 Check list of Rebutia Sensu Lato

c.v. ‘alabaster’ (hybrid R. minuscula v. violaciflora f.

kariusiana x R. c.v. stirnaldel’s meisterstuck.)
*albiareolata Ritter (FR761) (1977) Kakt. u. a. Sukk. 28 78
*albiflora Ritter & Buin. (FR766a) (1963) Taxon 12 29
*albipilosa Ritter (FR754) (1963) Taxon 12 29
albiseta Graessn. nomen nudum = ? aureiflora
*albopectinata Rausch (1972) Kakt. u. a. Sukk. 23 236
allegraiana Borg nomen nudum = senilis var. elegans
almeyeri W. Heinr. = ? senilis f. stuemeri
archibuingiana Ritter (1978) Ashingtonia 3 14
arenacea Card. = Sulcorebutia arenacea (Card.) Ritter
auranitida (Wessn.) Buin. & Donald (1937) Kakt. u. a. Sukk. 9 130
" var. flaviflora Backbg. = auranitida
" var. gracilis (Wessn.) Donald = auranitida f. gracilis
" f. gracilis (Wessn.) Buin. & Donald (1937) Kakt. u. a. Sukk. 9 130
aurantiaca hort. (catalogue 1974 Uhlig) U1343 = ? U1343

*aureiflora Backbg. (1932) Deutsch Kakteenfreunde 124

" var. albilongiseta (Backbg.) Marshall & Bock. comb. nud. = aureiflora
" var. albiseta (Backbg.) Marshall & Bock. comb. nud. = aureiflora
" f. boedeckeriana (Backbg.) Buin. & Donald (1934) Blatt. f. Kakteenf. 2
" var. densispina Borg nomen nudum = aureiflora
* " var. elegans (Backbg.) Buin. & Donald (1934) Blatt. f. Kakteenf. 9
* " f. duursmaiana (Backbg.) Buin. & Donald (1934) Blatt. f. Kakteenf. 9
" f. kesselringiana (Cullm.) Köhler = aureiflora f. rubriflora
" var. longiseta (Backbg.) Marshall & Bock. comb. nud. = aureiflora
" f. rubelliflora (Backbg.) Buin. & Donald (1935) Kaktus ABC. 247
" var. rubriflora (Backbg.) Buin. & Donald
* " f. rubriflora (Backbg.) Buin. & Donald (1935) Kaktus ABC. 247
" var. sarothroides (Werd.) Buin. & Donald (1936) Blüh. Sukk. f. 106
" var. sarothroides (Werd.) Buin. & Donald (1936) Blüh. Sukk. f. 106
" var. subspec. aureiflora (Backbg.) Donald = aureiflora
" var. subspec. aureiflora var. Blossfeldii (Werd.) Donald =
aureiflora f. rubriflora
" var. subspec. elegans (Backbg.) Donald = aureiflora var. elegans
" var. subspec. elegans var. sarothroides (Werd.) Donald =
aureiflora f. sarothroides

aurescens hort. (catalogue 1974 Uhlig) U1344 = ?

aureicantha hort. (catalogue 1974 Uhlig) U1756 = ?
*aureispina Knize nomen nudum KK843 (also KK1694)
beryllioides Donald nomen nudum = wessneriana var. beryllioides
" var. densiseta nomen nudum = wessneriana var. beryllioides
" var. longiseta nomen nudum = wessneriana var. beryllioides
*binnewaldiana W. Heinr. = ? wessneriana

60
 binghamiana Borg nomen nudum = Lobivia binghamiana
blossfeldii Werd. = aureiflora f. rubriflora
blossfeldiana Werd. ex Backbg. nomen nudum = ? aureiflora f. rubriflora
boedeckeriana (Backbg.) Marshall & Bock, nomen nudum =
aureiflora f. boedeckeriana
boliviensis hort. (catalogue 1974 Köhres) = ?
*brachyantha (Wessn.) Buin. & Donald (1937) Kakt. u. a. Sukk. 207
brachyantha Card. = Sulcorebutia breviflora (Card.) Backbg.
breviflora Card. nomen Illeg. = Sulcorebutia breviflora (Card.) Backbg.
*brunescens Rausch (R480) (1972) Kakt. u. a. Sukk. 23 235
*brunneoradicata Ritter (FR1109) Kakt. u. a. Sukk. 23 98
buiningiana Ritter nomen nudum = ? spinosissima
*buiningiana Rausch (R511) (1972) Kakt. u. a. Sukk. 23 98
cabratai Borg nomen nudum = Lobivia densispina
caineana Card. = Sulcorebutia caineana (Card.) Donald
cajasensis Ritter (FR1141) (1977) Succulenta 56 64
calliantha Bew. nomen nudum
calliantha Wessn. = wessneriana
" var. breviseta Backbg. = wessneriana
" var. beryllioides Buin. & Donald = wessneriana var. beryllioides
" var. beryllioides f. breviseta (Backbg.) Buin. & Donald = wessneriana
" f. hyalacantha Buin. & Donald = wessneriana
" var. densiseta Bew. nomen nudum = krainziana
" var. kariusiana Buin. & Donald = minuscula var. violaciflora f. kariusiana
" var. krainziana (Kess.) Buin. & Donald = krainziana
*camarguensis Rausch (R311) (1976) Succulenta 55 42
*canacruzensis Rausch (1976) Kakt. u. a. Sukk. 27 49 = a form of pygmaea
canaletas Knize nomen nudum KK1565
candiae Card. = Sulcorebutia candiae (Card.) Buin. & Donald
canigueralii Card. = Sulcorebutia canigueralii (Card.) Buin. & Donald
caracarensis Card. = Sulcorebutia caracarensis (Card.) Donald.
*carmeniana Rausch (1978) Kakt. u. a. Sukk. 29 105
carminea Buin. = minuscula var. violaciflora
christinae Rausch (R492a) (1975) Kakt. u. a. Sukk. 26 145
chrysacantha Backbg. (1935) Kaktus ABC, 416
" var. densispina nomen nudum
" var. elegans (Backbg.) Backbg. = senilis var. elegans
" var. iseliniana (Krainz) Donald comb. nud. =
senilis var. iseliniana
" var. kesselringiana (Bew.) Donald comb. nud. =
senilis var. kesselringiana
*cincinnata Rausch (R300) (1976) Kakt. u. a. Sukk. 27 4
*cintiensis Ritter(FR938) (1978) Ashingtonia 2 206
citricarpa Frič & Backbg. = xanthocarpa f. citricarpa
" var. salmonea Frič nomen nudum = xanthocarpa f. salmonea
*colorea Ritter (FR1106?) (1977) Kakt. u. a. Sukk. 28 78
conoidea Wessn. = einsteinii f. conoidea
" var. columnaris Wessn. = einsteinii f. columnaris
corroana Card. = Sulcorebutia corroana (Card.) Don. & Bred.

61
*costata Werd. (1934) Notizbl. Bot. Gait. u. Mus. Berlin 12 25
* " f. eucalyptana (Backbg.) Buin. & Donald (1935) Kaktus ABC, 414
" f. pilifera Buin. & Donald (1965) Cact. Succ. J. Gt. Brit. 27 40
dasyphrissa Werd. = xanthocarpa f. dasyphrissa
*deminuta (Web.) Br. & R. (1904) Bull. Mus. Hist. Nat. Paris, 386
* " f. pseudominuscula (Speg.) Buin. & Donald (1905) Anal Mus. Nac.
Buenos Aires Ser. III, 488
*densipectinata Ritter (FR758) nomen nudum = ? form of R. albopectinata
*diersiana Rausch (R631) (1975) Kakt. u. a. Sukk. 26 25
" var. atrovirens Rausch (R633) (1975) Kakt. u. a. Sukk. 26 26
" var. minor Rausch nomen nudum
donaldiana Lau & Rowley (Lau 348) (1974) Ashingtonia 1 76
durispina Frič nomen nudum = ?
" var. violrosea Frič nomen nudum = ?
duursmaiana (Backbg.) Marshall & Bock. comb. nud. = aureiflora f. duursmaiana

*einsteinii Frič ex Backbg. (1956) Descr. Cact. Nov. 30

" var. atrovirens Frič = ? einsteinii f. rubriviridis
" var. columnaris (Wessn.) Buin. & Donald = einsteinii f. columnaris
" f. karreri (Backbg.) Buin. & Donald = einsteinii
" var. columnaris f. conoidea (Wessn.) Buin. & Donald =
einsteinii f. conoidea
* " f. columnaris (Wessn.) Buin. & Donald (1940) Beitr. Sukk. u. pflege 4
" var. conoidea (Wessn.) Buin. & Donald = einsteinii f. conoidea
" f. conoidea (Wessn.) Buin. & Donald (1940) Beitr. Sukk. u. pflege 3
* " var. gonjianii (Kies.) Donald (1973) Bol. Soc. Argentina Bot. 15 132
" var. rubriviridis (Frič & Kreuzgr. ex Backbg.) Buin. & Donald =
einsteinii f. rubriviridis
" f. rubriviridis (Frič & Kreuzgr. ex Backbg.) Donald (1956)
Descr. Cact. Nov. 30
* " f. schmiedcheniana (Kohl.) Buin. & Donald (1939) Beitr. Sukk. u. pflege 37
" var. steineckei (Frič & Kreuzgr. ex. Backbg.) Buin. & Donald =
einsteinii f. steineckei
" f. steineckei (Frič & Kreuzgr. ex. Backbg.) Buin. & Donald (1956)
Descr. Cact. Nov. 30
elegans Backbg. = aureiflora var. elegans
" var. gracilis Backbg. = aureiflora
*eos Rausch (R333) (1972) Succulenta 51 2
" var. roseiflora Rausch nomen nudum (R333a)
espinosae Knize nomen nudum KK1518 = R. narvaesensis
euanthema (Backbg.) Buin. & Donald (1956) Backbg. Descr. Cact. Nov. 30
" f. fricii (Backbg.) Buin. & Donald (1956) Backbg. Descr. Cact. Nov. 30
" f. neopygmaea (Backbg.) Buin. & Donald (1956) Backbg.
Descr. Cact. Nov. 30
" f. oculata (Werd.) Buin. & Donald (1935) Blüh. Kakt. u. a. Sukk. 99
eucalyptana (Backbg.) Donald = costata f. eucalyptana (Backbg.) Buin. & Donald
*fabrisii Rausch (R688) (1977) Kakt. und Sukk. 28 52
" var. aureiflora Rausch (R687) (1977) Kakt. u. a. Sukk. 28 53
famatimensis (Speg.) Speg. = Lobivia densispina

62
*fiebigiana W. Heinr. = ? wessneriana
*fiebrigii (Guerke) Br. & R. (1905) Notizbl. Bot. Gart. u. Mus. Berlin 4 183
" var. densiseta (Cullm.) Oeser = fiebrigii f. densiseta
" var. densiseta Cullm.
*flavistyla Ritter (FR756) (1978) Ashingtonia 3 12
friciana Donald & Cullm. nomen nudum = euanthema f. fricii
fricii (Hort.) Backbg. nomen nudum = euanthema f. fricii
*friedrichiana Rausch (R630) (R642) (1976) Succulenta 55 101
*froehlichiana Rausch (R649) (1975) Succulenta 54 225
fuauxiana (Backbg.) Backbg. = R. pygmaea f. fuauxiana
*fulviseta Rausch (R583) (R319) (1970) Kakt. u. a. Sukk. 21 29
" var. albispina Rausch nomen nudum (R495)
*fusca Ritter (FR940) (1977) Kakt. u. a. Sukk. 28 78
fusca Hort. nomen nudum
gibbulosa Knize nomen nudum KK1563
glanduliflora Card. nom. prov. = Sulcorebutia spec.?
glaucescens Backbg. nomen nudum = ?
glomeriseta Card. = Sulcorebutia glomeriseta (Card.) Ritter
glomeriseta Card. = Sulcorebutia glomeriseta (Card.) Buin. & Donald
gonjianii Kiesl. = einsteinii var. gonjianii
*graciliflora Backbg. (1963) Descr. Cact. Nov. 3 13
graciliflora var. borealis Ritter nomen nudum FR34la
gracilis Borg nomen nudum Hybrid - intermediate form between R. senilis and
R. xanthocarpa
*gracilispina Ritter (FR1118) (1977) Kakt. u. a. Sukk. 28 76
graessneri Frič nomen nudum = minuscula var. violaciflora
grandiflora Backbg. non Frič = minuscula var. grandiflora
x grandilacea Johnson hybrid (grandiflora x violaciflora)
haageana Frič ex Backbg. = R. pygmaea
haagei Frič & Schelle (1930) Kaktusar 88 = R. pygmaea
haefneriana Cullm. = pygmaea f. haefneriana
hahniana Werd. = ? ritteri var. nigricans f. hahniana
hahnii Werd. = ?
hahnii Frič. nomen nudum = ?
haseltonii Card. = Sulcorebutia haseltonii (Card.) Donald
heinrichiana hort. nomen nudum (catalogue 1977 Uhlig) = ?
*heliosa Rausch (R314) (1970) Kakt. und Sukk. 21 30
" var. cajasensis Donald (Lau401) (1979) Ashingtonia 5 144
" var. condorensis Donald (Lau405) (1979) Ashingtonia 5 143
heliosa x albiflora An interesting new hybrid
hertrichiana comb. nud. = Lobivia hertrichiana
*hirsutissima Card. (1971) Cact. Succ. J. Am. 43 244
*hoffmannii Diers & Rausch (R521a) (1977) Kakt. u. a. Sukk. 28 105
*huasiensis Rausch (R313) (1977) Kakt. u. a. Sukk. 28 25
hyalacantha (Backbg.) Backbg. = wessneriana
inflexiseta Card. = Sulcorebutia inflexiseta (Card.) Donald
*iridescens Ritter (FR1434) (1977) Kakt. u. a. Sukk. 28 76
*iscayachensis Rausch (R335b) (1977) Succulenta 56 1
*ithyacantha (Card.) Diers. (1972) Kakt. u. a. Sukk. 23 12

63
*jujuyana Rausch (R220) (1973) Kakt. u. a. Sukk. 24 147
kariusiana Wessn. Locality unknown - selected from 200 field collected plants of
R. senilis and R. marsoneri imported by Uebelmann. (1963) Kakt. u. a. Sukk. 14 149 =
minuscula v. violaciflora f. kariusiana
*kieslingii Rausch (R694) (1977) Kakt. u. a. Sukk. 28 177
knuthiana Backbg. (1935) Kaktus ABC 416 = minuscula var. violaciflora f. knuthiana
*krainziana Kesselring (1948) Sukkulentenkunde II Jb. Schweiz Kakt. Ges. 23
" var. breviseta (Backbg.) Donald (1935) = ? Hybrid
krainziana x marsoneri Kaktus ABC 416
" f. breviseta (Backbg.) Krainz & Haarm.
" var. breviseta f. beryllioides (Buin. & Donald) Krainz & Haarm. =
wessneriana var. beryllioides
" var. hyalacantha (Backbg.) Buchheim = wessneriana
" var. longiseta nomen nudum = wessneriana
" var. wessneriana (Bew.) Krainz & Haarm. = wessneriana
" var. wessneriana f. calliantha (Bew.) Krainz & Haarm. = wessneriana
kruegeri (Card.) Backbg. = Sulcorebutia kruegeri (Card.) Ritter
*kupperiana Bod (1932) Monats. D. K. Gesell. 276
*kupperiana var. spiniflora Ritter (FR762b) (1977) Kakt. u. a. Sukk. 28 78
*lanosiflora Ritter (FR1116) (1977) Kakt. u. a. Sukk. 28 77
lasseniana hort. nomen nudum = ?
*lauii Donald (Lau416)
lateritia Knize nomen nudum KK1519
*leucanthema Rausch (R305) (1975) Kakt. u. a. Sukk. 26 125
" var. coccinifera Ritter (1977) Succulenta 56 63
longiflora Frič nomen nudum = ?
longispina Frič nomen nudum = ?
lucida Frič nomen nudum = ?
*mamillosa Rausch R302 (1972) Succulenta 51 69
" var. australis Rausch (1977) Kakt. u. a. Sukk. 28 77 = ? var. or form of
R. fulviseta
" var. orientalis Ritter (1977) Kakt. u. a. Sukk. 28 77 = ? var. or form of
R. spegazziniana
maresii Borg nomen nudum = Lobivia densispina
*margarethae Rausch (R521) (1972) Kakt. und Sukk. 23 4
*marsoneri Werd. (1937) Kakteenkunde 2
" var. albescens hort. nomen nudum
" var. breviseta hort. nomen nudum
" var. brevispina Donald nomen nudum
" var. grandiflora Donald nomen nudum
" var. sieperdaiana (Buin.) Donald = marsoneri f. sieperdaiana
* " f. sieperdaiana (Buin.) Buin. & Donald (1941) Succulenta 23 15
" var. spathulata hort. nomen nudum
" f. spathulata Donald nomen nudum
" c.v. spatula Donald
" var. vatteri Donald nomen nudum
maximiflora Frič nomen nudum = ?
c.v. meisterstuck Hort. White flowered hybrid produced by Stirnadel.
Rebutia spec x Echinopsis spec (polyancistra?)

64
 melachlora Ritter & Buin. nomen nudum = leucanthema v. cocciniflora
melanea Borg nomen nudum = Lobivia densispina ?
*minuscula K. Schum. (1895) Monatsschte Kakteenkde. 5 102
" f. bruneo-aurantiaca Simon nom. prov. = minuscula
" var. citricarpa (Frič) Simon = xanthocarpa f. citricarpa
" var. coerulescens (Backbg.) Simon = xanthocarpa f. coerulescens
" var. grandiflora Krainz nomen nudum
" var. grandiflora Marsh. nom. prov.
* " var. grandiflora (Backbg.) Buin. & Donald
" var. subspec. grandiflora (Backbg.) Donald = minuscula var. grandiflora
" f. knuthiana (Backbg.) Buin. & Donald = minuscula var.
violaciflora f. knuthiana
" var. multiflora Frič nomen nudum = minuscula
" f. kariusiana (Wessn.) Buin. & Donald = minuscula var.
violaciflora f. kariusiana
" var. salmonea (Frič) Simon = xanthocarpa f. salmonea
" var. senilis (Backbg.) Simon = senilis
" f. violaciflora (Backbg.) Buin. & Donald = minuscula var. violaciflora
* " var. violaciflora (Backbg.) Buin. & Donald (1935) Blatt. f. Kakt. 8
" var. violaciflora f. kariusiana (Wessn.) Donald (1963) u. a. Sukk. 14 149
" var. violaciflora f. rosiflora nom. prov.
" var. violaciflora f. knuthiana (Backbg.) Buin. & Donald (1935) Kaktus
ABC 277 & 416
" var. subspec. violaciflora (Backbg.) Donald = minuscula var. violaciflora
" var. subspec. violaciflora f. kariusiana (Wessn.) Donald = minuscula var.
violaciflora f. kariusiana
*minutissima Ritter (FR1124) (1977) Kakt. u. a. Sukk. 28 78
*mixta Ritter (FR1429) (1977) Kakt. u. a. Sukk. 28 76
*mixticolor Ritter (FR1108) (1977) Kakt. u. a. Sukk. 27 169
*mudanensis Rausch (R689) (1976) Kakt. u. a. Sukk. 27 169
multicolor Ritter nomen nudum (FR1108)
multiplex (Pfeiff.) v. Roeder (1938) Kakteenkunde 124 = ? Echinopsis multiplex
multicostata hort. nomen nudum = ?
muricata Frič nomen nudum = minuscula var. violaciflora
*muscula Ritter & Thiele (FR753) (1963) Taxon 12 29 1 -> 29
" var. nivosa Knize comb. nud. KK1301
*napina Ritter (FR942) nomen nudum (nom. prov.?)
narvaecensis (Card.) Donald (1971) Cact. & Succ. J. Am. 43 245
neopygmaea Backbg. = euanthema f. neopygmaea
nicolaii Frič nomen nudum = ? (1932) Mollers Deutsch. Gartnerzeit. 47 422
nidulans Borg nomen nudum = ?
nigra Frič nomen nudum = ?
nigricans (Wessn.) Backbg. = ritteri var. nigricans
*nitida Ritter (FR769) (1978) Ashingtonia 3 14
nivea nomen nudum = ? muscula
nivosa Ritter nomen nudum = ? muscula
*nogalensis Ritter (FR768) (1977) Kakt. und Sukk. 28 78
oculata Werd. (1935) Blüh. Kakt. u. a. Sukk. 99 = euanthema f. oculata
*odontopetala Ritter (FR757a) (1977) Kakt. u. a. Sukk. 28 76

65
 oderata Hort. nomen nudum = wessneriana
orurensis Backbg. = pygmaea
*padcayensis Rausch (R322) Succulenta 56 233
pallida Rausch (1977) Succulenta 56 233
*patericalyx Ritter (FR757) (1977) Kakt. u. a. Sukk. 28 74
" var. odontopetala Ritter nomen nudum
*pauciareolata Ritter (FR1121) (1977) Kakt. u. a. Sukk. 28 77
*paucicostata Ritter (FR936) (1977) Kakt. u. a. Sukk. 28 77
pectinata Backbg. = pygmaea
" var. atrovirens = pygmaea var. atrovirens
" var. digitiformis Backbg. = pygmaea
" var. neosteinmannii Backbg. = pygmaea f. neosteinmannii
" var. orurensis Backbg. = pygmaea
periferia hort. nomen nudum = xanthocarpa (f. salmonea?)
permutata Heinr. = wessneriana
" var. gokrausei Heinr. = wessneriana
peruviana hort. nomen nudum = marsoneri
perplexa Donald (Lau 329a) (not narvaecense (Card. Donald) (1979)
Ashingtonia 3 150
petersainii Borg nomen nudum = ritteri
petersonii hort. nomen nudum = minuscula
pilayensis Knize nomen nudum (KK863)
pilifera (Buin. & Donald) Rausch nomen nudum = costata f. pilifera
pilosa Knize nomen nudum (KK858)
polymorpha Card. = Sulcorebutia polymorpha (Card.) Backbg.
*potosina Ritter (FR1428) (1977) Kakt. u. a. Sukk. 28 77
prolifera Rausch nomen nudum = fabrisii
pseudodeminuta Backbg. non Frič (1933) D.K.F. 7
" f. albiseta (Backbg.) Buin. & Donald (1951) Cact. Succ. J. Amer. 23 82
" var. longiseta nomen nudum
" f. grandiflora (Backbg.) Buin. & Donald (1951) Cact. Succ. J. Amer. 23 82
" f. rubrifilamentosa Buin. & Donald (1963) Sukkulentenkunde 7 102
" f. schneideriana (Backbg.) Buin. & Donald (1951)
Cact. Succ. J. Amer. 23 82
" f. schumanniana (Backbg.) Buin. & Donald (1933) Der Kakteenfr. 7
pseudoheliosa nomen nudum = albopectinata (Lau 401)
pseudokrainziana nomen nudum
pseudominuscula (Speg.) Br. & R. = deminuta f. pseudominuscula
*pseudopygmaea Ritter (FR1122) nomen nudum
*pulchella Rausch (1972) Kakt. u. a. Sukk. 23 340
pulchera Card. = Sulcorebutia pulchera (Card.) Donald
*pulvinosa Ritter & Buin. (FR766) (1963) Taxon 12 29
pulvispina Knize nomen nudum KK1568
pseudograessneri hort. (catalogue 1974 Uhlig) U844 = ?
*pygmaea (R. E. Fries) Br. & R. (1905) Nov. Act. Soc. Sci. Upsala IV 120
" f. atrovirens (Backbg.) Buin. & Donald (1935) Kaktus ABC 242
" f. flavovirens (Backbg.) Buin. & Donald (1935) Kaktus ABC 243
" f. fuauxiana (Backbg.) Buin. & Donald (1956) Descr. Cact. Nov. 131
* " f. haefneriana (Cullm.) Buin. & Donald (1955) Kakt. u. a. Sukk. 6

66
 " f. neosteinmannii (Backbg.) Buin. & Donald (1956) Descr. Cact. Nov. 130
" var. tupizensis Rausch nomen nudum (R676)
*raulii Rausch (R485) (1980) Kakt. u. a. Sukk. 31 170
*rauschii Zecher (R297) (1977) Kakt. u. a. Sukk. 28 73
reichii Borg nomen nudum = ? Neoporteria reichii
residua Knize nomen nudum (KK1517)
rigidispina Frič nomen nudum = ?
*ritteri (Wessn.) Buin. & Donald (1938) Beitr. z. Skde. u. pflege 3
* " var. nigricans (Wessn.) Buin. & Donald (1938) Beitr. z. Sukk. u. Pfl. 51
" var. nigricans f. hahniana Buin. & Donald (1965) Cact. Succ. J. Gt. Brit. 27 40
" var. nigricans f. peterseimii Buin. & Donald (1965)
Cact. Succ. J. Gt. Brit. 27 40
" f. peterseimii Buin. & Donald = ritteri var. nigricans f. peterseimii
" f. hahniana Buin. & Donald = ritteri var. nigricans f. hahniana
ritteriana hort. (catalogue 1974 Andreae) = ? ritteri
*robustispina Ritter (FR618) (1977) Succulenta 56 64
" var. minor Ritter nomen nudum

*rosalbiflora Ritter (FR1115) (1977) Kakt. u. a. Sukk. 28 76

" var. amblypetala Ritter (FR1119) (1977) Kakt. u. a. Sukk. 28 76
rubelliflora (Backbg.) Marshall & Bock. comb. nud. = aureiflora f. rubelliflora
*rubiginosa Ritter (FR767) (1963) Taxon 12 29
rubispina Graessner nomen nudum = ?
rubrispina Backbg. = ?
rubriflora (Backbg.) Marshall & Bock. comb. nud. = aureiflora f. rubriflora

*rutiliflora Ritter (FR1113) (1977) Kakt. u. a. Sukk. 28 76

sieperdaiana Buin. = marsoneri f. sieperdaiana
*salpingantha Ritter (FR937) (1977) Kakt. u. a. Sukk. 28 77
salmonea Frič non Backbg. nomen nudum catalogue name = ?
sanguinea Ritter (1977) Succulenta 56 65
" var. minor Ritter nomen nudum
sarothroides Werd. = aureiflora f. sarothroides
scarlatea Frič nomen nudum = minuscula
" var. brevispina Frič nomen nudum = minuscula
*schatzliana Rausch (R640) Kakt. u. a. Sukk. 26 244
schmiedickeana (Kohl) Marsh. & Bock. comb. nud. = einsteinii f. schmiedcheniana

schmiedcheniana var. karreri Backbg. = einsteinii

" var. rubriviridis Frič = einsteinii f. rubriviridis
" var. steineckei Frič = einsteinii f. steineckei
*senilis Backbg. (1932) D. Kaktfrd. 123
" f. subspec. senilis Backbg. = senilis
" f. subspec. senilis f. lilacinorosea (Backbg.) Buin. & Donald =
senilis f. lilacinorosea
" f. subspec. senilis f. stuemeri (Backbg.) Buin. & Donald =
senilis f. stuemeri
" f. subspec. chrysacantha (Backbg.) Donald = chrysacantha
67
" f. subspec. chrysacantha f. iseliniana (Krainz) Buin. & Donald =
senilis var. iseliniana
" f. subspec. chrysacantha f. kesselringiana (Backbg.) Buin. & Donald =
senilis var. kesselringiana
" f. subspec. chrysacantha f. elegans (Backbg.) Buin. & Donald =
senilis var. elegans
" var. aurescens Backbg. = ? This taxon probably belongs with
R. wessneriana rather than with R. senilis. In our experience
it is self sterile and fits better with Subsection II rather than Subsection I.
" f. blossfeldii nomen nudum = senilis
" var. breviseta Backbg. = krainziana
" f. breviseta Backbg. = krainziana
" var. calva Frič nomen nudum = ?
" var. cana Borg nomen nudum = ?
" var. cana Backbg. = senilis
" f. cana Backbg. ex Simon. = senilis
" var. chrysacantha (Backbg.) Donald comb. nud. = chrysacantha
" var. chrysacantha f. elegans (Backbg.) Buin. & Donald =
senilis var. elegans
" var. chrysacantha f. iseliniana (Krainz) Buin. & Donald =
senilis var. iseliniana
" var. chrysacantha f. kesselringiana (Bew.) Buin. & Donald =
senilis var. kesselringiana
" var. citricarpa (Frič) Marsh = xanthocarpa f. citricarpa
" var. colinsii hort. (catalogue 1974 Uhlig) U2667 = ?
" var. dasyphrissa (Werd.) Marshall & Bock. comb. nud. =
xanthocarpa f. dasyphrissa
" var. dasyphrissa (Werd.) Donald = xanthocarpa f. dasyphrissa
" var. gracilis nomen nudum = ?
" var. elegans (Backbg.) Backbg. (1951) Cact. Succ. J. Amer. 23 83
" var. erecta hort. nomen nudum = ?
" var. hyalacantha Backbg. (Backbg.) Donald = wessneriana
" var. hyalacantha f. hyalacantha (Backbg.) ex Simon = wessneriana
" var. hyalacantha f. blossfeldiana Köhler nomen nudum = ?
" var. iseliniana Krainz (1946) Schweizer Garten 284
" var. iseliniana f. chrysacantha (Backbg.) Buin. & Donald = chrysacantha
" var. iseliniana f. elegans (Backbg.) Buin. & Donald = senilis var. elegans
" var. iseliniana f. kesselringiana (Bew.) Buin. & Donald =
senilis var. kesselringiana
" var. kesselringiana Bew. (1947) Sukkulentenkunde 19
" f. lilacinorosea (Backbg.) Buin. & Donald (1935) Kaktus ABC 416
" var. lilacinorosea Backbg. = senilis f. lilacinorosea
" var. longiflora nomen nudum = ?
" var. longispina Frič nomen nudum = ?
" var. luteirosea (Backbg.) Marsh = xanthocarpa var. luteirosea
" var. pallidior (Backbg.) Donald nomen comb. subnud. = ?
" var. pallidior (Backbg.) Donald comb. nud. = ? xanthocarpa
" var. sieperdaiana (Buin.) Backbg. = marsoneri f. sieperdaiana

68
 " var. schieliana Bew. = senilis f. schieliana
" f. schieliana (Bew.) Donald (1957) Kakt. u. a. Sukk. 8 105
" var. semperflorens Poind. = ?
" var. stuemeri Backbg. = senilis f. stuemeri
" f. stuemeri (Backbg.) Buin. & Donald (1932) Deutsch Kakt. frd. 131
" var. stuemeriana Backbg. = senilis f. stuemeri
" var. violaciflora Backbg. nomen nudum = ?
" var. xanthocarpa (Backbg.) Marsh = xanthocarpa
singularis Ritter FR1423 (1978) Ashingtonia 3 12
solisioides Knize nomen nudum = heliosa var. condorensis
*spegazziniana Backbg. (1934) Blätter, f. Kakteen forschg. 2
" atroviridis (Backbg.) Backbg. (1951) Cact. Succ. J. Am. 23 82
*sphaerica Ritter (FR1140) nomen nudum
*spinosissima Backbg. (1935) Blatt. f. Kakteenf. 1935-8
" var. brunispina Backbg. nomen nudum = spinosissima
" var. aurea Rausch nomen nudum (R318)
" var. similis Frič nomen nudum = spinosissima
spiralisepala Jajo nomen nudum
*steinmannii (Solms-Laub.) Br. & R. (1907) Bt. Zeit. 65 153 (Section I Aylostera!)
" var. cincinnata Rausch nomen nudum (R300) = ?
" var. nigricans Wessn. = ritteri var. nigricans
" var. pilifera (Buin. & Donald) Rausch comb. nud. = costata f. pilifera
*supthutiana Rausch (R629) (1976) Kakt. u. a. Sukk. 27 121
steinbachii Werd. = Sulcorebutia steinbachii (Werd.) Backbg.
" rosiflora Backbg. = Sulcorebutia steinbachii v. rosiflora
" var. violaciflora Backbg. = Sulcorebutia steinbachii v. violaciflora
" var. violacifera Backbg. = Sulcorebutia steinbachii v. violacifera
steineckei Frič nomen nudum (1932) Mollers Deutsch Gartner Zeit. 47 422 = ?
*tamboensis Ritter (FR1142) (1978) Ashingtonia 2 207
taratensis Card. = Sulcorebutia taratensis (Card.) Buin. & Donald
*tarijensis Rausch (R87) (1975) Kakt. u. a. Sukk. 26 195
*tarvitaensis Ritter (FR773) (1977) Kakt. u. a. Sukk. 28 78
tiraquensis Card. = Sulcorebutia tiraquensis (Card.) Ritter
" var. longiseta Card. = Sulcorebutia tiraquensis (Card.) Donald
*torquata Ritter & Buin. (FR1117) (1977) Succulenta 56 63
totorensis Card. = Sulcorebutia totorensis (Card.) Ritter
*tropaeolipicta Ritter (FR1114) (1977) Kakt. u. a. Sukk. 28 78
tuberculato-chrysacantha Card. = Sulcorebutia tuberculato-chrysacantha
(Card.) Bred. & Donald
*tuberosa Ritter (FR770) (1963) Taxon 12 28
tunariensis Card. = Sulcorebutia tunariensis (Card.) Buin. & Donald
turbinata hort. nomen nudum Winter catalogue name = ? wessneriana
*vallegrandensis Card. (1970) Cact. Succ. J. Amer. 42 35
violacea Frič hort. nomen nudum = xanthocarpa var. violaciflora
" var. albispina Frič nomen nudum = ?
" var. densispina Frič nomen nudum = ?
" var. paucipetala Frič nomen nudum = ?
" var. rosea Frič nomen nudum = ?

69
 " var. stellata Frič nomen nudum = ?
*violascens Ritter (FR352) (1977) Kakt. u. a. Sukk. 28 76
violaciflora Backbg. = minuscula var. violaciflora
" var. brunispina hort. (catalogue 1974 Uhlig) U1175
" var. carminea (Burning) Donald = minuscula var. violaciflora
" var. densispina nomen nudum = minuscula var. violaciflora
" var. graessneri nomen nudum = minuscula var. violaciflora
" var. knuthiana (Backbg.) Donald comb. nud. =
minuscula var. violaciflora f. knuthiana
" var. luteispina (Backbg.) Borg hort. nomen nudum =
minuscula var. violaciflora
" var. rosea hort. (catalogue 1974 Uhlig) Ul173
violbrunispina Frič nomen nudum = ?
violsalmonea Frič nomen nudum = ?
vizcarrae Card. = Sulcorebutia vizcarrae (Card.) Donald
volcanensis hort. (1974 catalogue Andraea) = ?
*vulpina Ritter (FR939) (1977) Succulenta 56 66
waltheriana Backbg. nomen nudum = ? spegazziniana
*wessneriana Bew. (1948) Sukkulentenkunde Jahrb. Schweiz. Kakt. Ges. 2 24
" var. beryllioides Buin. & Donald
" var. calliantha (Bew.) Krainz = wessneriana
" var. densiseta nomen nudum = krainziana
" var. krainziana (Kess.) Buin. & Donald = krainziana
" var. permutata (Heinr.) Buin. & Donald = wessneriana
" subspec. beryllioides Buin. & Donald = wessneriana var. beryllioides
" subspec. wessneriana f. calliantha (Bew.) Buin. & Donald =
wessneriana
" subspec. wessneriana var. gokrausei (Heinr.) Donald = wessneriana
" subspec. wessneriana var. gokrausei f. permutata (Heinr.) Donald =
wessneriana
c.v. 'Ruby' Donald
c.v. 'Turbine' Donald
winterae hort. nomen nudum = ?
winteriana hort. nomen nudum = ?
*xanthocarpa Backbg. (1932)D.K.F 131
" var. citricarpa Frič ex Backbg. = f. citricarpa
" f. citricarpa (Frič ex Backbg.) Buin. & Donald (1951) Cact. & Succ.
J Amer. 23 83
" var. coerulescens (Backbg.) Buin. & Donald = f. coerulescens
" f. coerulescens (Backbg.) Buin. & Donald (1956) Backbg. Descr. Cact.
Nov. 31
" var. dasyphrissa (Werd.) Backbg. = f. dasyphrissa
" f. dasyphrissa (Werd.) Buin. & Donald (1935) Blüh. Kakt. u. a. Sukk. 103
" var. elegans Backbg. = senilis var. elegans
" f. graciliflora (Backbg.) Donald (1963) Backbg. Descr. Cact. Nov. 3 13
" var. kesselringiana comb. nud. = senilis var. kesselringiana
" var. lilacinorosea nomen nudum = f. violaciflora
" var. luteirosea Backbg. (1951) Cact. Succ. J. Amer. 23 83
" var. magentaflora Hort. nomen nudum = f. violaciflora

70
 " var. pallidior Backbg. = ? xanthocarpa
" var. ruficeps hort. nomen nudum = ?
" var. salmonea Frič ex Backbg. = f. salmonea
" f. salmonea (Frič ex Backbg.) Buin. & Donald (1951) Cact. Succ. ]. Amer.
23 83
" f. sanguinea nomen nudum = ?
" var. violaciflora (Backbg.) Backbg. = f. violaciflora
" f. violaciflora (Backbg.) Buin. & Donald (1956) Backbg. Descr. Cact.
Nov. 31
zavaletae Card. = Sulcorebutia zavaletae (Card.) Backbg.
*zecheri Rausch (R650) (1977) Succulenta 56 29
Digitorebutia nazarenoensis Rausch. This new species was described by Walter
Rausch in 1979. As it has not been described as a Rebutia the following combination
has been made:-
Rebutia nazarenoensis (Rausch). Fearn & Pearcy comb. nov.
Digitorebutia nazarenoensis Rausch (1979) Succulenta 58 185.

Field Data
The following lists of field numbers are those which have been assigned to
Rebutia plants collected by Friedrich Ritter, Walter Rausch, Alfred Lau and Karel
Knize, all of whom have collected extensively in Bolivia and Argentina.

Friedrich Ritter Field Numbers

(All these field numbers are prefixed by the letters FR)
57 Digitorebutia haagei
57a " brachyantha (pygmaea?)
57b " haagei form
58 Rebutia calliantha
83 fiebrigii densiseta (nivosa)
84 " fiebrigii
84a " fiebrigii var.
84b " fiebrigii var. vulpes
337 " spec. (brachyantha?)
339 Mediolobivia orurensis
340 " eucalyptana
341 " haefneriana steinmannii
341a Rebutia (mamillosa v. australis) graciliflora v. borealis
341b " mamillosa v. orientalis
351 " haagei steinmannii
352 " (violascens) haefneriana
390 " fiebrigii v. densiseta
402 Mediolobivia pilifera
404 Aylostera spegazzinii v. atroviridis
405 Mediolobivia sarothroides
406 Rebutia euanthema
407 Mediolobivia aureiflora
71
 618 Aylostera robustispina
622 " spec.
752 " pseudodeminuta
753 " muscula
754 " albipilosa
755 " maxima
756 " flavistyla
757 " patericalyx
757a " var. odontopetala
758 " densipectinata (dark form)
759 " densipectinata (pale form)
760 " sanguinea spegazziniana
760a " sanguinea var. minor
761 " albiareolata
762 Rebutia kupperiana
762a " " form
762b " " var. spiniflora
763 " robustispina
763a " " var. minor
765 " spec.
766 Aylostera pulvinosa
766a " albiflora
767 " rubiginosa
768 Rebutia nogalensis
769 " nitida
770 " tuberosa
771 " spec.
772 " spec.
773 " tarvitaensis
935 " melachlora
936 " ritteri v. paucicostata
937 " salpingantha costata
938 " cintiensis
939 " vulpina
939a " vulpina var.
940 " fusca
941 " senilis
941a " spec.
942 Rebutia napina
1101 " gracilis
1106 " colorea
1106a " colorea var.
1107 " pygmaea
1108 " mixticolor
1109 " brunneoradicata
1110 " mamillosa var. australis - grandiflora
1113 " rutiliflora
1113a " spec.
1114 " tropaeolipicta

72
 1115 " rosalbiflora
1115a " spec.
1116 " lanosiflora
1117 " torquata
1118 " gracilispina
1118a " gracilispina var.
1119 " rosalbiflora v. amblypetala
1120 " spec.
1121 " pauciareolata
1122 " pseudopygmaea
1123 " ritteri
1124 " minutissima
1130 " spec.
1138 " mamillosa v. orientalis
1138a " mamillosa var.
1139 " poecilantha
1140 " sphaerica
1141 Mediolobivia cajasensis
1142 Rebutia tamboensis
1142a " pseudodeminuta
1142b " pseudodeminuta form
1423 " singularis
1428 " potosina
1429 " mixta
1434 " iridescens

Rebutias collected by Alfred Lau

(All these field numbers are prefixed by the word LAU)
323 R. fiebrigii forma nova. 2700m Mina Asientos, Cochabamba.
329 R. fiebrigii var. densiseta. 2600m Mina Asientos, Cochabamba.
329a R. perplexa
346 R. fiebrigii form, shorter and darker centrals. 2400m Pucara-La Higuera,
Santa Cruz.
346a R. spinosissima var. nova. 2400m Pucara-La Higuera, Santa Cruz.
348 R. donaldiana. 2500m Pucara-Vallegrande, Santa Cruz.
348a R. fiebrigii. 2500m Pucara-Vallegrande, Santa Cruz.
350 R. ithyacantha. 2600m Pucara-Vallegrande, Santa Cruz.
351 R. vallegrandensis. 2100m Pucara-Vallegrande, Santa Cruz.
353 R. vallegrandensis. 2600m Pucara-Vallegrande Pass, Santa Cruz.
383 R. brunescens. 3500m Tarabuco, Chuquisaca.
393 R. fiebrigii form. 2900m. Calle Calle Mts., Chuquisaca.
401 R. albopectinata. 2450m Condor Pass, Tarija.
402 R. rubiginosa. 1600m Narvaez, Tarija.
402a R. fiebrigii form. 1600m Narvaez, Tarija.
404 R. buiningiana Ritt. nomen nudum (R. spinosissima?) 2500m Cara del Diablo,
Tarija.
405 R. albopectinata form. 2400m Cajas Pass, Tarija.

73
 407 R. rubiginosa form. 2400m. 10km. W. Narvaez, Tarija.
409 R. spec. nova. 3500m Tarija to Iscayachi.
410 R. tuberosa. 3500m Tarija to Iscayachi.
411 R. christinae. Tarija to Iscayachi.
411a R. spegazziniana var. atroviridis 3000m Tarija to Iscayachi.
412 R. spegazziniana var. nova. Road to Cajas, Tarija.
413 R. padcayensis. Cara del Diablo, Tarija.
415 = 405 R. albopectinata. 3200m Iscayachi, Tarija.
416 R. lauii. 2400m Cajas Pass, Tarija.
422a R. spec. nova. 3350m Otavi-Camargo, Chuquisaca.
422b R. spec. nova. 3350m Otavi-Camargo, Chuquisaca.
424 R. spec. 3300m Camargo, Chuquisaca.
431 R. pygmaea. 3800m El Aguilar, Jujuy.
442 R. minuscula var. 1500m Sierra Medina, Tucuman Province,
476 R. einsteinii var. gonjianii 3500m Quebrada del Toro, Jujuy.
477 R. einsteinii var. gonjianii 3200m Quebrada del Toro, Jujuy.
519 R. haagei. 3100m Humahuaca, Jujuy.
520 R. spec, (similar to R. costata and R. fuauxiana). 3000m East of Volcan, Jujuy.
522 R. pseudodeminuta. 2700m East of Volcan, Jujuy.
533 R. spec. nova (possibly R. auranitida var. gracilis). 2700m above Tastil, Salta.
541 R. spec.(nigricans?) 3800m Tafne, Jujuy.
541a R. spec. (costata?) 3600m Tafne, Jujuy.
541b R. eos. 3600m Tafne, Jujuy.
541c R. eos var. nova? 3600m. Tafne, Jujuy.
544 R. margarethae. 2100-3000m Santa Victoria, Salta.
544a R. margarethae. 2400m Santa Victoria, Salta.
545 R. jujuyensis. 2300m Santa Victoria, Salta.
546 R. spec. nova. 2200m Santa Victoria, Salta.
546a R. spec. nova. 2200m Santa Victoria, Salta.
547 R. spegazziniana v. atroviridis 2300-3000m Santa Victoria, Salta.
550 R. margarethae. 2800m Road from La Quiaca, Santa Victoria Pass, Salta.
550a R. pseudodeminuta form? 2800m Road from La Quiaca, Santa Victoria Pass,
Salta.
554 R. margarethae. 3500m Road from La Quiaca, Santa Victoria Pass, Salta.
557 R. wessneriana v. beryllioides. 2400m Leon, Jujuy Province.
560 R. xanthocarpa v. lilacinorosea. 10km North West of Antilla, Salta.
561 R. pseudodeminuta form. 10km North West of Antilla, Salta.

Walter Rausch Field Numbers

(All these field numbers are prefixed by the letter R)
1 Rebutia xanthocarpa
11 Aylostera pseudominuscula
14 Mediolobivia nigricans var. Piedra de Molinos
35 " haagei
67 Aylostera ithyacantha
71 Mediolobivia steinmannii var.
87 " tarijensis (Ritter?)

74
 88 Aylostera robustispina
89 " fiebrigii v. densiseta
91 Mediolobivia spec.
158 " aureiflora
162 " nigricans
163 " einsteinii v. atrovirens (viridis?)
167 Rebutia senilis
208 Mediolobivia steinmannii var.
208a " haefneriana
210 " orurensis
214 " euanthema
220 Aylostera sordida/jujuyana
234 Rebutia wessneriana
274 Aylostera vallegrandensis
295 Rebutia rosalbiflora Potosi
295a " spec. Iscayache
296 Mediolobivia steinmannii v. pilifera
297 " rauschii
300 " steinmannii v. cincinnata
301 " pectinata (yellow)
302 Rebutia mammillosa
304 Mediolobivia pectinata (white)
305 Rebutia leucanthema
311 Aylostera umbraculiforma = camarguensis
312 Mediolobivia albopectinata
313 Rebutia huasiensis
314 Aylostera heliosa
317 " spinosissima
318 " var. aurea? var. buiningiana?
319 " fulviseta
320 " lampromeliana = pulchella
321 " fiebrigii v. castanea
322 " padcayensis
324 " kupperiana
333 Mediolobivia eos
333a " eos var. roseiflora
334 " steinmannii v. rotundipetala
335 " pygmaea (Villazon)
335a " pygmaea (Iscayachi)
335b " iscayachensis (Corda)
454 " eucalyptana
480 Rebutia brunescens (Tarabuco?)
484 Mediolobivia mudanensis
485 " rauhii (Rio Honda)
486 " steinmannii v. complanata (Rio Honda)
492 Aylostera graciliflora (Iscayachi)
492a Rebutia christinae (Santa Victoria)
493 " brunescens var.
494 Mediolobivia steinmannii v. rubriflora

75
 495 Rebutia fulviseta v. albispina
496 Aylostera spegazziniana
501 Mediolobivia haagei
502 " haagei
503 Aylostera fiebrigii
503a " fiebrigii f. brown spine
506 Mediolobivia pygmaea var.
507 " haagei var. (Santa Victoria-Purmamarca)
507a " haagei var.
508 " costata
508a " pygmaea v. longispina
509 " einsteinii
511 Rebutia buiningiana
515 Mediolobivia haefneriana
518 Rebutia margarethae
520 Mediolobivia ritteri
521 Rebutia margarethae
521a " hoffmannii
522 Mediolobivia morenoensis
576 " pygmaea
577 " nigricans
578 Rebutia einsteinii v. gonjianii
579 Mediolobivia pygmaea
583 Rebutia fulviseta
597 " pulchella var.
629 " supthutiana
630 Rebutia friedrichiana (pygmaea?)
631 " diersiana
632 " haefneriana
633 " diersiana var. atrovirens
640 " schatzliana
641 " schatzliana var.
642 " friedrichiana
643 " spinosissima
644 " steinmannii var.
646 Mediolobivia friedrichiana
649 Rebutia froehlichiana
650 Aylostera zecheri
654 " pseudominuscula var.
655 " pseudominuscula
660 Rebutia violascens
661 " senilis
676 " pygmaea v. tupizensis
679 Aylostera pseudodeminuta var.
680 Rebutia aureiflora var. duursmaiana
681 " violaciflora
687 " fabrisii var. aureiflora
688 " fabrisii
689 Mediolobivia mudanensis

76
 690 Rebutia pseudominuscula (carmeniana)
694 " kieslingii
697 " pseudodeminuta
700 " oculata

Karel Knize Field Numbers

(All these numbers are prefixed by the letters KK)
597 Rebutia senilis f. Huahata, 2200m, Arg.
598 " pseudominuscula. Catamarca, 2800m, Arg.
637 Mediolobivia pectinata. Oruro, 4000m, Bolivia.
797 " haefneriana. Oruro, Capulla, 3000m, Bolivia.
798 " pectinata var. Oruro, Machacamarca, 4000m, Bolivia.
839 Rebutia albipilosa. Tarcia, Tarija, 2800m, Bolivia.
840 " violaciflora. Salta, Escoipe, 3100m, Arg.
841 " narvaecensis f. Narvaez, Tarija, 2300m, Bolivia.
842 " muscula. Piedra, Larga, 2300m, Bolivia.
843 " aureispina. Jucanas, Tarija, 3000m, Bolivia.
844 " heliosa. Jucanas, 3000m, Bolivia.
845 " senilis f. Salta, Escoipe, 3200m, Arg.
846 " pseudodeminuta. Piedra Larga, Tarija, 2800m, Bolivia,
847 " deminuta f. Narvaez, Tarija, 3000m, Bolivia.
848 " densiseta? Jucanas, Tarija, 3000m, Bolivia.
849 " densipectinata. Jucanas, Tarija, 3000m, Bolivia
852 " albopectinata. Sama, Tarija, 3800m, Bolivia.
853 " fiebrigii v. densiseta. Iscayache, Tarija, 3500m, Bolivia.
854 Mediolobivia conoidea. Iscayache, Tarija, 3000m, Bolivia
855 Rebutia fiebrigii. El Molle, Tarija, 3000m, Bolivia.
856 " spegazzinii. Iscayache, 3500m, Bolivia.
858 " pilosa. Jucanas, Las Cajas, 2800m, Bolivia.
859 " ritteri/tuberosa. Sama, Tarija, 3500m, Bolivia.
860 " buiningiana. Las Cajas, Tarija, 2800m, Bolivia.
861 " rubiginosa. San Antonio, Iscayache, 3200m, Bolivia.
862 " tuberosa. Sama, Tarija, 3500m, Bolivia.
863 " pilayensis. El Nogal, Tarija, 2500m, Bolivia.
871 Mediolobivia euanthema. Sama, Iscayache, 4300m, Bolivia.
872 " auranitida? Sama, Iscayache, 4300m, Bolivia.
873 " auranitida v. gracilis. Sama, Tarija, 4200m, Bolivia.
968 " orurensis. Oruro, Paria, 4100m, Bolivia.
969 " pectinata. v. paznaensis. Pazna, 4100m, Bolivia.
970 " pectinata. v. gracilis. Pazna, 4300m, Bolivia.
971 " eucalyptana. Eucalyptos, 4100m, Bolivia.
972 " pectinata v. challa. Challapata, 4000m, Bolivia.
973 " euanthema v. oculata. Challapata, 4100m, Bolivia.
974 " pectinata v. Poopo, 4200m, Bolivia.
978 " spec. Huari, 4000m, Bolivia.
1046 " permutata?

77
 1052 Rebutia ithyacantha. La Cueva, Tarija, 2500m, Bolivia.
1096 " violaciflora v. knuthiana. Salta, El Aguilar, 2800m, Arg.
1150 " narvaecensis. Piedra Larga, 2500m, Bolivia.
1151 " muscula v. nivosa. Piedra Larga, 2600m, Bolivia.
1152 " spec.
1232 Mediolobivia ritteri. Iscayache, 3800m, Bolivia.
1242 Rebutia grandiflora f. Salta, Escoipe, 3300m, Arg.
1243 " senilis f. Salta, Iscayache, 3500m, Arg.
1244 " xanthocarpa. Salta, El Aguilar, 3800m, Arg.
1301 " muscula v. nivosa. Narvaez, Tarija, 3000m, Bolivia.
1302 " narvaecensis. Narvaez, Tarija, 2800m, Bolivia.
1464 Mediolobivia elegans. Salta, 3800m, Arg.
1465 " pygmaea. El Aguilar, 4000m, Arg.
1466 " nigricans. Salta, 3800m, Arg.
1467 " pectinata v. digitiformis. Escoipe, 3800m, Arg.
1468 " auranitida var. Sierra Padjoja, 4100m, Bolivia.
1469 " auranitida var. Sierra Padjoja, 4100m, Bolivia.
1516 Rebutia cintiensis
1517 " residua
1518 " espinosae
1519 " lateritia
1563 " gibbulosa
1565 " canaletas
1568 " pulvispina
1686 Mediolobivia iscayachensis
1694 Rebutia aureispina
1695 " potosina

Bibliography
The following is a list of the most important publications
concerning the genus Rebutia
Backeberg C. (1932) Der Kakteenfreund
Backeberg C. (1951) Some Results of Twenty Years of Cactus Research.
Cactus Succ. J. Amer. 23 81-83
Backeberg C. (1959) Die Cactaceae Vol. II Fischer Verlag, Jena.
Backeberg C. (1961) Die Cactaceae Vol. VI Fischer Verlag, Jena.
Backeberg C. (1966) Kakteen Lexikon Fischer Verlag, Jena.
Backeberg C. (1977) Cactus Lexicon Blandford Press, Poole.
Backeberg C. & Knuth F.M. (1935) Kaktus ABC Gyldendalske Boghandel,
Denmark.
Berger A. (1929) Kakteen, Stuttgart
Bertrand A. (1951) Genre 66 Rebutia K. Sch. emend A. Bertrand. Cactus (Paris) 28 53
Brederoo A.J. (1955) Rebutiae Succulenta, 36
Brederoo A.J. (1956) Aylostera Succulenta, 56
Brederoo A.J. (1956) Mediolobivia Succulenta, 92
Brederoo A.J. (1957) Digitorebutia Succulenta, 53
Brederoo A.J. (1957) Mediolobivia spiralisepala Jajo Succulenta, 92
78
Brederoo A.J. (1959) Cylindrorebutia Succulenta, 121
Brinkman K.H. (1976) Die Gattung Sulcorebutia Kakt. u. a. Sukk.
Britton N.L. & Rose J.N. (1923) The Cactaceae 1st Ed. Carnegie Institute,
Washington Publication No. 248
Borg J. (1937) Cacti. Macmillan, London
Buining A.F.H. & Donald J.D. (1963) Die Gattung Rebutia Sukkulentenkunde Jb.
Schweiz Kakt. Ges. 97
Buxbaum F. (1938) Was ist Rebutia, was Aylostera? Beitr. Sukk. u. Pflege, 69.
Buxbaum F. (1958) Cactus Culture based on Biology. Blandford, London
Buxbaum F. (1967) Rebutia, Krainz Die Kakteen 37
Donald J.D. (1950) An introduction to some new genera and subgenera proposed
by Backeberg. Natn. Cactus Succ. J. 5 67
Donald J.D. (1955) A Rebutia Reference and Synonymy Brit. Sect. IOS Publ. No. 2.
Donald J.D. (1956) Rebutiae, Succulenta 84
Donald J.D. (1957) Guide to the Rebutiae I The Nomenclature of the Rebutiae,
Natn. Cactus Succ. J. 12 3
Donald J.D. (1957) Guide to the Rebutiae II Key to the Rebutiae, Natn. Cactus Succ.
J. 12 9
Donald J.D. (1957) Notes and observations on New Cacti, I Rebutia krainziana
and its varieties, Natn. Cactus Succ. J. 12 27
Donald J.D. (1957) Was ist Rebutia violaciflora und was Rebutia carminea,
Kakt. u. a. Sukk. 8 24
Donald J.D. (1959) New plants Nr. 2 Rebutia calliantha and Rebutia wessneriana
Natn. Cactus Succ. J. 14 4
Donald J.D. (1973) Rebutia albopectinata Ashingtonia 1 23
Donald J.D. (1974) The Rebutias From the Lau expeditions to Bolivia and Argentina,
Ashingtonia 1 64
Donald J.D. (1976) Studies over Rebutia, Echinopsis en Lobivia, Succulenta 55 235
Donald J.D. (1974-1978) The Classification of the Rebutias, Ashingtonia 1-3
Donald J.D. (1977) Echinopsis, Lobivia, Rebutia. Where to draw the line?
Cact. Succ. J. Gt. Brit. 39 11
Donald J.D. (1977) Further observations on the Rebutia, Sulcorebutia, Weingartia
problem, IOS Bulletin 3 167
Donald J.D. & Brederoo A.J. (1978) The systematics of Rebutia Pt. 7. Ashingtonia 3
Fearn B. (1974) An investigation into the effect of temperature on the seed
germination of nine species of Cacti using thermal gradient bars.
Cactus Succ. J. Amer. 46 215
Fearn B. (1977) An investigation into the effect of age on the germination potential
of seeds of 600 species of Cacti. Excelsa 7 103
Fritz G. (1979) Rebutia albopectinata Rausch. Kakt. u. a. Sukk. 30 12
Fritz G. (1980) Nochmals zu Rebutia spec. Lau 401/405. Kakt. u. a. Sukk. 31 81
Glass C. & Foster R. (1980) Whats New. Cactus Succ. J. Am. 52 164
Goemas W. Maretlmans L. & Wacker Fde. (1959) Rebutia, Succulenta 38 30 & 53
Köhler von U. (1957) Die rosaviolett-Blühenden Rebutien, Kakt. u. a. Sukk. 8 28
Köhler von U. (1961) Medio (Pygmaea) Lobivia pygmaea (Fries) Backbg. Kakt.
u. a. Sukk. 12 105
Köhler von U. (1963) Zweibrücker Hybride 1962 15 113
Krainz H. (1967) Rebutia, Die Kakteen.
Marshall W.T. & Bock T.M. (1941) Cactaceae. Abbey Garden Press, Pasadena.

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Marshall W.T. & Woods R.S. (1945) Glossary of Succulent Plant Terms
Abbey Gdn. Press Pasadena.
Rausch W. (1975) Walter Rausch comments Ashingtonia 1 128
Ritter F. (1977) Nieuwe cactussen uit Zuid-Amerika Succulenta 66
Stace C.A. (1980) Plant Taxonomy and Biosystematics, Contemporary Biology,
Edward Arnold.
Schumann K. (1897) Gesemtbeschreibung der Kakteen, Neudamm.
Schumann K. & Guerke M. (1898) Blühende Kakteen, Neudamm.

Acknowledgements
We sincerely wish to thank all the various persons who have helped in the
preparation of this booklet. The descriptions are based on plants in cultivation at
Abbey Brook.
For technical suggestions and helpful advice to Mr. N. Wall of Higham Press and
Carol and Ann at Abbey Brook for typing the original manuscript. Also to the
Librarian, Royal Botanic Gardens, Kew for the use of facilities, to Dr. Gillian Fearn for
help in proof reading and to Thelma Stuart for photographic assistance.

Glossary
acicular needle shaped.
apomixis a biological system which can produce viable seeds by a process
other than by direct fertilisation of the egg cell,
apomictic a species which can produce seeds by the process of apomixis.
areole a restricted area from which spines and/or flowers are produced,
bristle hair-like spine,
caespitose growing in clumps,
campanulate bell-shaped,
central spine the spine or spines arising from or near the centre of an areole,
usually the largest spine or spines,
filament the stalk upon which is borne the anther or pollen sac.
funnel-form a flower which gradually widens from the base,
globose shaped like a globe,
hilum the scar at the base of the seed making the point of attachment to the
funicle.
spine a pointed more or less rigid structure which is homologous with a
leaf.
stamen the male floral organ which bears the pollen grains. It consists of a
stalk called the filament and an anther or pollen sac.
stigma the part of the flower that is the pollen receptor,
style the stalk joining the stigma to the ovary,
type locality the place where a type specimen was collected.

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 The Cactus Centre of England
Abbey Brook is the oldest established Cactus Nursery in the U. K. and we celebrated
our 25th Anniversary in 1981. We have a vast range of plants for sale (well over 2,000
species) and more than 500,000 plants on view.
Plants and seeds of a wide range of Rebutia species available.

Comprehensive colour illustrated plant list available, stamp appreciated.

We also produce a separate seed list.

Nursery Visits
The nursery is open to visitors 6 days a week throughout the year including Saturdays
and Sundays.
2p.m. - 6p.m. (Closed all day Tuesdays)
We are open on all Bank Holidays except Christmas Day, Boxing Day and New Years
Day.
The nursery is situated just north of Matlock on the A6 trunk road.

Abbey Brook Cactus Nursery

Old Hackney Lane,
Matlock, Derbyshire, England.
☎ National 0629 55360
☎ International + 44 629 55360

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R. euanthema ▲ ▲ R. xanthocarpa f. salmonea
R. wessneriana ▼ ▼ R. albipilosa

R. senilis v. elegans ▼ ▼ R. marsoneri

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