Found by Rainer Wahl south of Cerro Lorayo, Tarija, Bolivia, at an altitude of 2500m.

Found by Rudolf Slaba at Culpina, Chuquisaca, Bolivia.

Found by Ralf Hillmann near Camargo, Chuquisaca, Bolivia.

Found by Ladislav Fischer at Inca Huasi, Salta, Argentina.

Found by Ralf Hillmann at Alfarsito, Salta, Argentina.

Found by Omar Ferrari at Jujuy, Argentina.

Found by Walter Rausch in Bolivia; location not certain. This is the type plant for this species.

Found by Ralf Hillmann north of Camargo, Chuquisaca, Bolivia at an altitude of 3500m.

Found by Ralf Hillmann at Carichi Mayu, Tarija, Bolivia, at an altitude of 2450m.

Found by Ralf Hillmann between San Lorenzo and Rio Pilaya, Tarija, Bolivia, at an altitude of 2950m.

Found by Martin Lowry at Inca Huasi, Chuquisaca, Bolivia, at an altitude of 2950m. Note that originally this plant was