White wood aster

Appearance

''Eurybia divaricata'' is a late summer to fall-flowering herbaceous perennial, typically growing to heights between 30 and 90 cm, though some specimens may be up to 1.2 metres tall. The plant emerges each year from rhizomes and forms dense colonies of clones that lack sterile rosettes. The rhizomes are branched, elongated and become woody with age. One simple erect stem is present per plant. It is flexible and nearly hairless to finely hairy near the base, though densely covered with fine hair towards the extremities. It is very similar to and often confused with ''E. chlorolepis'', ''E. schreberi'' and ''Symphyotrichum cordifolium'', though ''E. schreberi'' differs in having wider leaves with more teeth, while ''E. chlorolepis'' has more rays, longer involucres, and only occurs in the southern U.S. from Virginia to Georgia.

Three types of leaves with differing morphology are present: cauline leaves, or those that appear on the stem from the middle of the plant upwards; basal leaves, or those that are present at the base of the plant; and distal leaves, which are those found on the extremities of the plant. All types are thin and sharply serrated with 6 to 15 pointed teeth per side. They are ciliate, meaning they have small hairy projections emerging from the margins of the leaf, while the apices, or tips of the leaves, are acuminate, meaning they taper to a point. The adaxial surfaces of the leaves are nearly hairless or sparsely hairy, while the abaxial surfaces are sparsely hairy with the veins being more villous, or covered in shaggy hairs.

The basal leaves are ovate, or egg-shaped, with bases that are cordate, or heart-shaped. The blades measure 1.9 to 6.5 cm in length by 1.7 to 6 cm in width and have petioles ranging in length from 2 to 7 cm. They wither when the plant flowers. The cauline leaves have petioles measuring 2.5 to 7 mm long that are often winged. Their blades are also ovate, though the bases may be cordate to rounded. They measure 2 to 20 cm long by 1 to 10 cm wide, making them often much longer than the basal leaves. The distal leaves are typically sessile, meaning that no petiole is present, though they are sometimes subpetiolate, meaning a very short petiole is present. The blades are ovate to lanceolate, meaning lance-shaped, with rounded bases and are 0.5 to 2 cm long by 0.1 to 0.8 cm wide.

The capitula, or flower heads, are arranged in relatively flat-topped corymbiform arrays. The capitula number anywhere from 4 to 50 and up to 100 or more in exceptional cases. The peduncles, i.e. the flower stalks, are up to 1.5 cm in length and are densely covered with non-glandular hairs. Bracts, modified leaves that appear at the axil of a peduncle, are typically absent, though in some cases up to two are present. The involucres, which are the whorls of small, scale-like modified leaves that appear at the base of the capitulum, are in between cylindric and campanulate in shape and measure 4.2 to 6 mm long, making them much shorter than the pappi.


The phyllaries, which are the small leaves that make up the involucre, number from 25 to 30 and are arranged in 4 to 5 series. The inner phyllaries are between linear and lanceolate in shape with a purplish colouration towards the apices, while the outer ones are more oblong. All of the phyllaries are strongly unequal with hardened bases, margins that are somewhat scarious as well as fimbriate-ciliate, meaning fringed with hair at the margins. Their apices are rounded to acute in shape, while the surfaces are sparsely haired, though sometimes sparsely covered in stipular glands. Their chlorophyllous zones, a darker green zone where chlorophyll is concentrated, appear on the upper half of the outer phyllaries, to the upper third or along the outer midveins of the inner phyllaries. The outer phyllaries typically measure 0.7 to 1.5 mm wide with the lengths rarely exceeding 2.5 times the width.

As with most members of the composite family, the actual flowers appear in two different forms: as ray florets, which have strap-like appendages that look like petals and project around the outside of the capitulum, and as disc florets, which appear at the center of the flower head and are very small. The ray florets number between 5 and 10, though as many as 12 may be present. Their straps are white and measure 6 to 12 mm long by 1.5 to 2.2 mm wide. The disc florets number from 12 to 19 and up to 25 and have yellow corollas that are 4.1 to 4.8 and exceptionally 5.5 mm long. Their corollas are abruptly ampliate, or enlarged, with tubes that are longer than their campanulate throats. The tubes measure 2.3 to 2.6 mm while the throats are typically only 0.9 to 1.2 mm long. The lobes, i.e. the friges of the throat, are reflexed and lanceolate in shape, measuring 0.7 to 1.4 mm.

The fruit are cypselae, a type of achene, which are brown in colour, slightly compressed and are between cylindric and obovoid, or inversely egg-shaped. They are between 2.6 and 3.8 mm in length and sparsely strigillose, or set with stiff bristly hairs, with 7 to 10 ribs, which themselves are tan to stramineous . The pappi, which are modified sepals, are made up of reddish to cream-coloured bristles that are 3.7 to 5 mm long, making them equal to or longer than the disc corollas in length. The bristles are fine and barbellulate, or barb-like, though they may be sometimes more or less clavate, or club-shaped, towards their apices.

Naming

Several different plants are superficially quite similar to the white wood aster, but close examination as well as knowledge of the plants' differing habitats and ranges can readily distinguish them. The most similar species is the mountain wood aster , which was previously considered conspecfic with ''E. divaricata''. ''E. chlorolepis'' differs in having flower stalks that are longer than 1.5 cm, while those of ''E. divaricata'' are shorter than this. The involucres of the mountain wood aster are generally between 6.5 and 9 mm in length, while those of the white wood aster are normally from 4.2 to 6.5 mm long, but in rare cases they may be as long as 7.5 mm. The mountain wood aster almost always has more rays than the white wood aster: the former usually has 12 to 16, but occasionally as few as 10, while the latter generally has 5 to 10 and never more than 12. The rays are also longer on the mountain wood aster at 17 to 18 mm in most cases, while the white wood aster's are generally only 10 to 15 mm. Lastly, ''E. chlorolepsis'' is present only from southern Virginia south to Georgia in the Appalachian Mountains at altitudes of 1,200 to 2,000 metres , while ''E. divaricata'' can be found from southern Canada south to Alabama, meaning their ranges only overlap in the southern United States, and there only at high elevations.

Another species commonly confused with the white wood aster is Schreber's aster . Schreber's aster is typically found on moister soils, though it can also be found on mesic sites. While much of their ranges overlap, Schreber's aster is not found south of Virginia or Tennessee. Schreber's aster can also be distinguished by the more numerous teeth on the leaves, typically numbering 15 to 30 per side to the white wood aster's 6 to 15 per side. The leaves are wider on Schreber's aster with broader sinuses at the bases. The flower stalks are also densely hairy on the white wood aster, while they are only sparsely haired on Schreber's aster. Lastly, the clones of ''E. schreberi'' have sterile rosettes, while those of ''E. divaricata'' do not.

Two other superficially similar plants include the bigleaf aster and the heartleaf aster , though both usually have blue or pale blue rays, while those of ''E. divaricata'' are always white. Both species sometimes have white rays in rare cases, but ''E. macrophylla'' can still be distinguished by its larger and broader leaves and ''S. cordifolium'' by its straighter and branched stems.

Distribution

''Eurybia divaricata'' is present primarily in the Appalachian Mountains in eastern North America, with some populations in adjacent lowlands. It can be found on dry to mesic sites in eastern deciduous and mixed deciduous woods as well as on edges, clearings, and roadsides. It is most common at altitudes ranging from 0 to 1,200 metres , though it can be found as high as 1,700 metres . In Canada, it is present in Ontario and Quebec, while in the United States it can be found in all states from Maine south as far as Alabama and west to Ohio and Kentucky. It also has been introduced to Europe, particularly to the Netherlands.

The species is known from only 25 locations in Canada, and is considered threatened.

Habitat

''Eurybia divaricata'' is present primarily in the Appalachian Mountains in eastern North America, with some populations in adjacent lowlands. It can be found on dry to mesic sites in eastern deciduous and mixed deciduous woods as well as on edges, clearings, and roadsides. It is most common at altitudes ranging from 0 to 1,200 metres , though it can be found as high as 1,700 metres . In Canada, it is present in Ontario and Quebec, while in the United States it can be found in all states from Maine south as far as Alabama and west to Ohio and Kentucky. It also has been introduced to Europe, particularly to the Netherlands.

The species is known from only 25 locations in Canada, and is considered threatened.