BIBLIOGRAPHY &
ANNOTATED CHECKLIST
of
LIVING ORGANISMS FIRST NAMED
from the
G R A N D C A N Y O N and V I C I N I T Y
(NORTHWESTERN ARIZONA)
compiled and edited by
EARLE E. SPAMER
_____________________________________________________________________________________
COVER ILLUSTRATION Ostrya knowltonii Coville, 1894 (Betulaceae). Coville’s original illustration
of the holotype (US) from the top of the Old Hance Trail, Grand Canyon, Arizona, gathered by J. W.
Toumey, 10 July 1892. (See No. 84 herein, pp. 42–43.)
BACK COVER General view of the type locality of O. knowltonii, looking downward at the top of the
Old Hance Trail. Photo by Nancy Brian during the visit that rediscovered the locality, 23 May 1997.
BIBLIOGRAPHY &
ANNOTATED CHECKLIST
of
LIVING ORGANISMS FIRST NAMED
from the
G R A N D C A N Y O N and V I C I N I T Y
(NORTHWESTERN ARIZONA)
compiled and edited by
EARLE E. SPAMER
RAVE N ’ S P E RCH ME DIA
PHILADELPHIA
1
2022
Copyright © Earle E. Spamer 2022
This publication is a resource manager’s and historian’s reference work documenting
species-level taxa of living organisms that were first scientifically named based wholly or
in part on collections made at, in, or near the Grand Canyon. This is not a work of systematics nor of taxonomic revision. Taxonomic combinations are those of the original authors
or authorities; subsequent revisions of taxonomy and typification are not cited. Publications and pertinent taxa meeting the purpose of the checklist are as known to the compiler
as of November 2022.
RAVEN’S PERCH MEDIA, Philadelphia, Pennsylvania https://ravensperch.org
Bibliographical and Historical Resources in the Grand Canyon and Lower Colorado River Regions
of the United States and Mexico
Published 11 DECEMBER 2022
REPRODUCTION AND FAIR USE • No commercial or for-profit use of this work is allowed
without the author’s permission. Not-for-profit organizations, educational institutions,
government agencies, and Indigenous communities may with credit use this work for
purposes of resource management and interpretation, education, and public outreach; they
also may download the entire document for cataloging and inclusion in their digital
products collections, and they may freely allow users to copy this document under the
conditions stated here. Reuse of this work, in whole or in part, for any purpose, must cite
author, title, publisher, and notice of copyright. Academic or other individual researchers
may copy selections for personal reference and may quote from this work following best
practices of acknowledgment in scholastic research. This statement does not pertain to,
nor supersedes, copyrights of items that still may be in effect among the items cited in the
bibliography.
2
CONTENTS
[In the PDF document, each line in the contents list is a hyperlink]
Map of Approximate Area of Coverage For This Checklist
6
Introduction
7
Commemorative Names
9
Conventions and Styles Used in This Checklist
12
Note on Paleontological Names
16
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
19
Serials Listed in the Bibliography and Annotated Checklist
114
Taxonomic List
117
Kingdom Animalia
119
Phylum Arthropoda
119
Phylum Chordata
128
Phylum Mollusca
130
Phylum Nematoda
131
Kingdom Plantae
132
Class Magnoliopsida
132
Class Pinopsida
137
Kingdom Fungi
138
Phylum (Division) Ascomycota
138
Phylum (Division) Basidiomycota
139
Kingdom Chromista (Phylum Apicomplexa)
140
Kingdom Protista (“Phylum” Hemimastigophora)
140
3
Contents (cont’d)
Localities for Type Material from Grand Canyon National Park
Map of Federally-Designated Boundaries of Grand Canyon National Park and
Predecessor Units (1893–present)
141
142
“Grand Canyon” not discriminated
143
South Rim
144
North Rim
150
Inner Canyon
154
Colorado River Locales
161
Localities for Type Material from Near Grand Canyon
165
Cameron, Arizona, and vicinity
166
Arizona Strip
167
Grand Canyon vicinity undiscriminated
171
Grand Canyon–Parashant National Monument
172
Havasupai Indian Reservation
174
House Rock and House Rock Valley
174
Hualapai Indian Reservation
175
Hualapai Indian Reservation vicinity (off reservation)
177
“Kaibab Forest” (north) undiscriminated
177
“Kaibab Forest” (south) undiscriminated
177
Kaibab Indian Reservation
178
Kaibab National Forest, Tusayan Ranger District
178
Kaibab Plateau (including Kaibab National Forest, North Kaibab Ranger District) 179
Kanab Canyon
184
Lees Ferry
184
Marble Canyon [not on Colorado River]
184
Navajo Nation (westernmost)
185
Red Mountain
185
Index to Nominate Taxa
187
___________________________________________
4
5
APPROXIMATE AREA OF COVERAGE FOR THIS CHECKLIST
UTAH
Kaibab
Indian
Reservation
A R I Z O N A
Lees Ferry
↘
S T R I P
Kaibab National
Forest
(Kaibab Ranger
District)
ARIZONA
NEVADA
(north and west of the Colorado River)
House
Rock
Valley
Kaibab
Plateau
Western extremity of
Havasupai
Indian
Reservation
the Navajo Nation
Tusayan →
(Tusayan Ranger District)
Base map from Google, emended
6
INTRODUCTION
INTRODUCTION
This checklist provides federal and Native American resource managers and historians of
the Grand Canyon region of northwestern Arizona with a census of living (neontological)
organisms that had been scientifically named based on collections made wholly on, or in
part from, the lands that these managers oversee. Their original scientific names are
recorded in this checklist. It does not use updated or otherwise “current” names for taxa
that have been scientifically reidentified (taxonomically synonymized) or that were
systematically transfered to other supra-specific biological groups than those under which
they were first named. Providing such information is beyond the scope of a historicallyand administratively-directed document such as this one; its inclusion would soon make
the records herein out of date. For the same reasons no data regarding the biological
attributes (descriptions) of the organisms are included herein.
Federal administrative units to which this accounting may be of particular interest are:
U.S. Department of the Interior
National Park Service
Grand Canyon National Park
Glen Canyon National Recreation Area (in that area near where it shares a
boundary with the Grand Canyon park at Lees Ferry)
Bureau of Land Management
Arizona Strip District
Joint management by NPS and BLM
Grand Canyon–Parashant National Monument
U.S. Department of Agriculture
U.S. Forest Service
Kaibab National Forest (Tusayan and North Kaibab Ranger Districts only)
Jurisdictions of Native American peoples addressed herein pertain to:
Havasupai Tribe [south side of the Grand Canyon]
Hualapai Indian Tribe [south side of the Grand Canyon]
Kaibab Band of Paiute Indians [on the Arizona Strip]
Navajo Nation [to the east of the Grand Canyon]
7
INTRODUCTION
A few of the collections cited herein seem to have come from privately held lands
that are inholdings within the federal units, and from within the boundaries of particular
municipalities in Arizona.
The bibliography/checklist includes 322 species-level taxa1 named on type specimens2 collected in the greater Grand Canyon region of northwestern Arizona; 215 of those
taxa are based at least in part on specimens from within, or are with good probability from
within, the current boundaries of Grand Canyon National Park. Five taxonomic kingdoms
are represented among them—animals, plants, fungi, photosynthetic eukaryotes, and
protists. Only the name-bearing types from within the geographical boundaries of this
checklist are listed; referred specimens (those not selected to be types) are generally
omitted.
It is probable that there are additional taxa that can be included in this checklist, but
they were not discovered during the course of the several decades of work that has
contributed to it, although hardly little of that work was devoted to this specific purpose.
This is an opportunistic work, condensing facts for an administrative document of interest
to historians, too.
There is far more beyond the geographical boundaries imposed here that might still
be considered to be in the greater Grand Canyon region, which are ignored for the simple
reason that there must be a limit. For example, there is an equal wealth of taxonomic
neology from the lands best recognized now as the Navajo Nation, which does nudge briefly
across this checklist’s eastern boundary; that area is accommodated herein. Then there is
the wholly different zoo and garden of the Mohave Desert to the west, which, while it is
decisively segregated from the Grand Canyon region by physiography and environment,
does in fact sneak into the western part of the canyon along the lower elevations of the
Colorado River corridor; so a few taxa more representative of those regions are included
here. This bibliography and checklist finds its origin in my own researches in history,
science and bibliography of the Grand Canyon vicinity in particular, thus the geographical
bounds are the same.
1
2
“Species-level taxon” means any name that is at the taxonomic rank of species, subspecies, or
variety. [The count includes No. 244a, added late.]
“Type specimens,” more generally called “types”, are the very specimens that were selected by a
species-level taxon’s author as bearing the formal characteristics that uniquely identify the taxon.
There may be just one, or even hundreds, and they can be from different localities (even beyond
the geographical area of interest in this checklist).
8
INTRODUCTION
The checklist is also confined taxonomically to the original names of the organisms
scientifically described for the first time by the authors cited in the bibliography. These
names can disappear through taxonomic synonymy (and in many cases they have) when
researchers in biological systematics have ascertained that one or another organism is
actually the same as another, earlier named one, or if they move species-level taxa to
different taxa than those in which they were originally placed. And in turn, genera can be
forever “fiddled with”, moving them from one Family to another and often carrying its
contained species with it (though this is a bit simplified); and similarly among the higher
taxa, clear up to the Kingdom level. In some measure this is reflected in the highersystematics listed with the taxa herein, which information may not be the same as that
originally assigned by the authors of these taxa.
There is also a chance that some organisms can disappear actually, through extirpation or extinction; and this checklist thus would provide the original data that were communicated about their presence and scientific discovery. Ecological memory is ephemeral,
but taxonomic memory—right, wrong, and revised—is forever, part of the resource manager’s treasury. Thus this checklist serves these managers as a documentary list of organisms first named from the lands that they oversee and the resources that have been used,
and it serves as a contribution to the history of the Grand Canyon and adjacent areas of
northwestern Arizona.
Commemorative Names
When a researcher names a new taxon they usually apply a “scientific” name that describes
some feature or attribute of the organism to which the name is given. Other times, one or
more persons are honored with the epithet, which customarily has one of the common
Latin gender-specific suffixes, -i, -ii, -ae, or -orum. (For example, in this checklist see Ostrya
knowltoni Coville, 1894 [No. 84 in the sequentially enumerated checklist, and also the
cover illustration], an ironwood from the Old Hance Trail, named for Frank H. Knowlton.3
3
The epithet was later corrected to knowltonii. This was a grammatical correction only, which does
not affect the author’s name. Grammatical corrections do not affect the epithet, being mandated
alterations to the Latin suffix only, such as this change from -i to -ii, so the author’s name is not
placed in parentheses. (When a species is moved to a different genus, or a subspecies to a different
species, than the ones under which they were first named, the author’s name is placed within
parentheses. An example of a transfer, from among the listings herein, is Crotalus confluentus
abyssus Klauber, 1930 [No. 183], the “Grand Canyon Rattlesnake”, which was later redescribed as
Crotalus viridis abyssus (Klauber), and has since also been placed in the genus Oreganus.)
9
INTRODUCTION
Often, a place is commemorated, usually with the suffix -ensis or another, grammatically figured suffix like -ense; other formulations are used, too. A few examples from this
checklist are:
238 Oreohelix yavapai angelica Pilsbry & Ferriss, 1911; a land snail from Bright Angel Trail
77
Sclerocactus havasupaiensis Clover, 1942; a cactus from “Havasupai Canyon” (Havasu
Canyon)
158 Opuntia hualpaensis Hester, 1943; a cactus from Route 66 between Peach Springs and Hyde
Park, the name denoting the Hualapai Indian Reservation
245 Panicum mohavense Reeder, 1991; a grass from Mohave County, on the Arizona Strip
109 Echinocereus triglochidiatus var. toroweapensis Fischer, 1991; a cactus from Toroweap
Point
A specially commemorative name may also acknowledge a regional feature closely
associated with the collecting locality; in these two examples, the Colorado River 4:
179 Astragalus kentrophyta var. coloradoensis Jones, 1902; an herb from Lees Ferry
280 Helix (Arionta) coloradoensis Stearns, 1890, a land snail from the Old Hance Trail [the taxon
is now Sonorella coloradoensis (Stearns)]
In the case of the Grand Canyon itself, seven species-level taxa have been erected
with epithets that commemorate the canyon or denote the canyon’s attribute as an “abyss”:
235 Rosa stellata Wooton subsp. abyssa Phillips, 1992; a rose from Twin Point, Shivwits Plateau
159 Opuntia abyssi Hester, 1943; a cactus from Peach Springs Canyon
183 Crotalus confluentus abyssus Klauber, 1930; a rattlesnake from Tanner Trail (the “Grand
Canyon Rattlesnake”)
270 Melanoplus canonicus Scudder, 1897; a grasshopper from Grand Canyon (no specific locale)
54
Aphelonema convergens var. canyonensia Bunn, 1930; a planthopper from Grand Canyon
(no specific locale)
267 Mentzelia canyonensis Schenk, Hodgson & Hufford, 2013; a loasa from Manzanita Canyon
166 Arenivaga grandiscanyonensis Hopkins, 2014; a sand cockroach from Colorado River Mile
211.5 Right
4
When searching for examples like these, bear in mind that such names are contextual. While
Misumenops coloradensis Gertsch, 1933 is listed in this checklist [No. 113] for its paratype specimens from the Grand Canyon, it is not included in this list of examples because the epithet,
coloradensis, was based on the primary type material having come from the State of Colorado.
10
INTRODUCTION
The Kaibab Plateau is the most commonly commemorated place in this checklist,
with 24 taxa named based on at least one type specimen. They are chiefly from the national
forest, but a few are from within the boundary of Grand Canyon National Park. “Kaibab” is
introduced into a Latinized epithet in various ways (not all of which may be in “good” Latin,
depending upon the author’s decision or the context of the name). These are, in alphabetical order by species-level taxon:
117 Loxosceles kaiba Gertsch & Ennik, 1983
217 Agave kaibabensis McKelvey, 1949
253 Andrena (Scaphandrena) kaibabensis Ribble, 1974
48
Branchinecta kaibabensis Belk & Fugate, 2000
164 Catilleja kaibabensis N. Holmgren, 1973
172 Cicindela pussila kaibabensis Johnson, 1990
74
Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986
108 Coryphantha vivipara var. kaibabensis Fischer, 1979
182 Enderleinellus kaibabensis Kim, 1966
227 Felis concolor kaibabensis E. Nelson & Goldman, 1931
258 Lesquerella kaibabensis Rollins, 1982
50
Mischocyttarus flavitaris var. kaibabensis Bequaert, 1932
42
Morchella kaibabensis Beug, T.A. Clem. & T.J. Baroni in Baroni et al., 2018
248 Morsea california kaibabensis Rehn & Grant, 1958
45
Papilio indra kaibabensis Bauer, 1955
115 Phidippus kaibabensis Gertsch, 1934b
155 Polistes canadensis var. kaibabensis Lynn, 1932
220 Sciurus kaibabensis Merriam, 1904
252 Scutellaria potosina var. kaibabensis Rhodes & Ayers, 2010
288 Suillus kaibabensis Thiers, 1976
123 Thomomys fossor kaibabensis Goldman, 1938
3
Tipula (Lunatipula) kaibabensis C. Alexander, 1946
223 Saxinis saucia kaibabiae Moldenke, 1970
180 Astragalus kaibensis Jones, 1902
11
INTRODUCTION
Conventions and Styles Used in This Checklist
The information given here is repeated at the heads of pertinent parts of this volume.
Throughout the volume, Red Numbers indicate and cross-list the sequentially enumerated
taxa in the bibliography.
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
In this checklist the species-level taxa are the names of organisms are as originally erected
by the authors. This is a historical record of nomenclature, not one of taxonomic revision.
For convenience, the systematic levels of phylum, class, order, and family are listed for
each enumerated taxon. Refer to the Taxonomic List herein for other details.
Arrow indicates taxa for which at least one locality reported for its types is in, or with
good probabability within, Grand Canyon National Park. Refer to the bibliography for all
details.
Data were not often provided by the original authors in any concise or systematic
fashion, thus direct quotations have been preferred in this checklist. By quoting, the user
can grasp either the paucity or breadth of data that were conveyed in the original author’s
locality and collecting information for the taxon there named. The information has not been
editorially revised herein to create an artificial sense of conformity; doing so might obscure
original precision, inconsistencies, or confusing data.
Particular localities that lie within the geographical boundaries of this checklist are
displayed in bold in order to bring attention to them, often amidst more detailed data.
Refer to the respective separate lists of localities in this volume for other details.
Data regarding the collectors and dates of collection are provided as published by the
original author, usually as part of the quoted information. However, take note that many
nominate taxa are not necessarily erected based on collections by the author themself, but
on specimens sent to the researcher or are previously collected materials that were
examined in museum collections. The published information thus records the data that
appeared with the examined specimens—although this distinction is not always perfectly
clear.
Information regarding the type specimens themselves is generally as published by the
taxon’s author. The particular codes of taxonomic nomenclature are followed here; there
are independent codes for the animal and botanical kingdoms. In this checklist, zoological
type specimens are indicated as holotype, paratype, allotype, and syntype; botanical type
12
INTRODUCTION
specimens are indicated as holotype, isotype, syntype, and paratype. 5 Other terms (like
topotype) are unessential and have no formal standing. Matters pertaining to lecto- or
neotypifications and similar procedural acts, which are made after the original publication
of a taxon, are beyond the scope of this checklist. In a few instances, older terminology was
used by an author and is updated as needed herein as a parenthetical note; for example,
cotype (= syntype in zoology, or used erroneously in botany to indicate an isotype or
paratype).
Genus and higher taxonomic levels have no type material, although older literature may
indicate such terms as “genotype”, which refers to the “type species” (one species on which
the description of a genus is based); the term has occasionally been used incorrectly to
denote a particular specimen.
When the checklist indicates only para- or isotypical material as having been collected
at a locality within the bounds of the checklist, a note is usually prefixed to the quotation
to indicate where the holotype (and if applicable, allotype) material was collected; so
indicated for the informational value that indicates these primary types were collected
elsewhere.
In the checklist’s quoted data, certain interpretive remarks are inserted within [square
brackets], introduced by the compiler. These usually clarify that the term “Type” or “the
type” refers to the holotype specimen; and when other specimens are cited they may be
interpreted to be (though not specifically mentioned as) material that is paratypical (in
zoology) or iso- or paratypical (in botany). In older literature, prior to the establishment of
formal codes of taxonomic nomenclature, later researchers are generally more liberal in
the acceptance of type status for material that supplements the holotype, except in those
cases where the original author may have precisely identified which specimens are types.
5
“Type specimens,” more generally called “types”, are the very specimens that were selected by a
species-level taxon’s author as bearing the formally defined physical characteristics that uniquely
identify the taxon. In older published descriptions, prior to the creation of the various international
conventions for taxonomic nomenclature, a time when types were not always specially designated,
it is the surviving elements known to have been used by the author in the original description that
now represent the type material for a given taxon. “Holotype,” allotype,” and “isotype” are so-called
primary types; other types supplement them in defining a taxon’s characteristics in such a way that
characteristics not well shown in one specimen may be exhibited by another. “Paratype” has
different meanings in zoology and botany: in zoology it is applied to one or more specimens that
supplement the description of the name-bearing holotype; in botany it is applied to one or more
specimens that were not formally specified to be type material but which were cited along with the
type material. The latter procedure is different from that followed in zoology, where specimens not
formally identified as type material (usually listed as “examined” or “referred” specimens) have no
status at all as types.
13
INTRODUCTION
Today the criteria are more rigorously applied; type material for new species-level taxa
must be uniquely identified.
When repository catalog numbers were published by the original author, they appear
as part of the quoted material. However, since this is a nomenclatural checklist only, no
attempt has been made to canvass repositories to identifiy either the continued existence
of specimens or their catalog numbers. Standard repository acronyms are not spelled out.
Throughout this checklist all text within [square brackets] has been inserted by the
compiler, except where noted otherwise.
TAXONOMIC LIST
This is a nomenclatural record. Taxonomic combinations are as named by the original
authors; later taxonomic reassignments, if any, are not generally noted herein.
The checklist for the greater Grand Canyon region lists 322 taxa first named from
collections in this area of northwestern Arizona, 215 of which were named from collections
made within, or with good proabability within, the present boundaries of Grand Canyon
National Park. [No. 244a was added late, thus the whole-number count stops at 321.]
Higher systematics for the genus to which the nominate taxa were assigned are given
here, as derived from the Integrated Taxonomic Information System (ITIS) online in 2022.
Taxa not included in ITIS have had their higher systematics derived from other sources,
including the National Center for Biotechnology Information Taxonomy Browser (National
Library of Medicine) and the World Flora Online. The systematic placements listed here may
not reflect the most current views, which are in any case bound to vary between researchers.
Again, this is a resource manager’s checklist to nominate taxa, not one of systematics or
revised nomenclature. In any case, for a species to be reassigned to another family does not
happen often, except in significant works of systematic revision, especially those that erect
new families.
The taxonomic ranks cited in this list are: Phylum or Division: Class: Order: Family.
Users of this bibliography and checklist may note that in some titles certain systematic
groups do not correspond to the groups that are listed for each of the taxa cited from a given
publication. This generally reflects a revised pespective of familial and higher groupings;
again, as usually derived for this checklist from ITIS and other sources.
Arrow indicates taxa for which at least one locality reported for its types is within, or is
with good probability from within, the boundaries of Grand Canyon National Park. Refer
to the bibliography for all details.
14
INTRODUCTION
The taxa are not listed in so-called “systematic order” because most users of this
checklist will not likely be attuned to the scientifically proper ordering of taxa within each
Phylum, Class, and so forth. To accommodate these more general users, who are more likely
to be resource managers and not necessarily practicing biologists, all higher taxa are listed
strictly in alphabetical order (the order of Kingdoms is arranged only for convenience). The
nominate taxa within each category are listed in the order in which they appear in the
checklist, arranged numerically to facilitate finding them in that bibliograph/checklist.
Common names, mostly for the Families, are as listed by ITIS or other sources. They
are not regularly adopted by all users and have been added here only a matter of convenience
for those who might be more comfortable referring only to common names.
LOCALITY LISTS
Two lists of localities where type material was collected are provided herein; one for
localities within the present boundaries of Grand Canyon National Park, the other for localities outside of the park and within the bounds defined for this checklist (see map).
Localities for the collection of type material are as identified by the authors of the taxa.
The bibliography/checklist quotes the authors’ data so as to record exactly how the data
were reported. The arrangement of the localities in these lists are, however, in some measure
modified for geographical and contextual sense; but specific determinants as communicated
by the authors of the taxa are retained (for example, differences in altitude at the same
general locality). In a few instances, sufficient historical or bibliographical data have made it
possible to locate a place with greater precision, which information is appended to the basic
locality data, and thus the data for a locality from which some type material came has been
refined herein.
Many localities are identified only to very broad geographical constraints—simply
“Grand Canyon”, for example, as indicated mostly in older literature—which reflects the
original data. In such cases no specifically identifiable type localities can be discriminated. At
the most general, the only locality information is “Grand Canyon”, which does not distinguish
either rim or inner-canyon locales. Except for the earliest years, it is unfortunate that the
records of “Grand Canyon, Ariz.” have not distinguished whether a collection was made at
Grand Canyon village (the place name, “Grand Canyon, Arizona”, which unofficially
sometimes also capitalizes “Village”) or more generally the “Grand Canyon of Arizona”; thus
taxa with this information for locality are separately listed. Only when the village area is
specifically stated, or inferred from other information such as “Bright Angel, Ariz.” (which
refers to the old Bright Angel Hotel), are taxa listed specifically from that locale. Similarly,
potentially misleading data are recorded in localities cited as “North Rim, Grand Canyon” or
similar arrangement, because there is the sometimes unofficial community name, “North
15
INTRODUCTION
Rim”, which while it has been a postal address is otherwise the tourist conclave at Bright
Angel Point on the North Rim. Another potentially confusing locale is “Vermilion Cliffs”,
which may refer to the unofficial place name, a business called “Vermilion Cliffs”, which is
along U.S. Route 89A along the base of the Vermilion Cliffs west of Marble Canyon.
INDEX TO NOMINATE TAXA
Names in bold indicate nominate taxa for which at least one locality reported for its types is
within, or is with good probability from within, the present boundaries of Grand Canyon
National Park. The names are original combinations, as published by the authors. The
taxonomic phylum is appended to each name and author. Refer to the bibliography and
annotated checklist for details.
Note on Paleontological Names
One may also ask about the record of new taxa named based on paleontological collections
from the Grand Canyon and vicinity, the complement to the neontological record presented
by this checklist. Those taxa are the intellectual antecedents of the living ark; in the Grand
Canyon they are the lithologically embalmed life of the Proterozoic Eon to the Permian
Period and the urologically indurated records of the Pleistocene, encroaching on the
Recent Epoch (about 11,700 years before present). (There are also a few nearby records of
fossils from the Triassic Period at Cedar Mountain, and a handful of Pliocene Epoch fossils
from nearby Anita, south of the canyon.) These all have been accounted for in earlier
bibliographies and checklists. Though now the publications are bit dated, lacking a comparatively small number of new taxa named since the early 1990s, they are itemized here for
convenience and to alert interested users to their presence.
A general bibliography of Grand Canyon paleontology (recently updated):
Spamer, Earle E.
2019
Bibliography of Paleontology of the Grand Canyon Region and in the Stratigraphic
Continuity of Grand Canyon Formations : compiled to commemorate the centennial of
Grand Canyon National Park and National Fossil Day 2019. Raven’s Perch Media,
Philadelphia, 120 pp. (PDF). Occasionally upated online at Raven’s Perch Media,
https://ravensperch.org/bibliography-of-paleontology-of-the-grand-canyon-region/.
16
INTRODUCTION
Historical overviews of paleontological research in Grand Canyon National Park:
Spamer, Earle E.
1984
Paleontology in the Grand Canyon of Arizona: 125 years of lessons and enigmas from the
late Precambrian to the present. The Mosasaur (Journal of the Delaware Valley Paleontological Society), Volume 2, pp. 45-128.
2020
History of paleontological work at Grand Canyon National Park. Up and down the long
federal and NGO trails of paleontology in Grand Canyon National Park, 1858-2019.
Chapter 2 in: Santucci, Vincent L., and Tweet, Justin S. (eds.), Grand Canyon National Park:
centennial paleontological resource inventory (non-sensitive version). Fort Collins, Colorado: U.S. National Park Service, Natural Resource Stewardship and Science, pp. 11-44.
(Volume: Natural Resource Report NPS/GRCA/NRR—2020/2103.)
2021
History of paleontological work at Grand Canyon National Park—up and down the long
federal and NGO trails of paleontology in Grand Canyon National Park, 1858-2019. In:
Santucci, Vincent L., and Tweet, Justin S. (eds.), Grand Canyon National Park centennial
paleontological resource inventory; a century of fossil discovery and research. Utah
Geological Association, Special Publication 1, pp. 11-32. [The volume is a reset and bound
reprinting of the Natural Resource Report by Santucci and Tweet (2020).]
Publications that contain information about paleontological taxa first named from
collections made in the Grand Canyon and at nearby localities are listed below. These
microform publications have since been made available as PDFs online by the Geological
Society of America, https://pubs.geoscienceworld.org/books/.
Spamer, Earle E.
1983
Geology of the Grand Canyon: An annotated bibliography, 1857-1982, with an annotated
catalogue of Grand Canyon type fossils. (Foreword by Edwin D. McKee.) Geological
Society of America, Microform Publication 13, 543 [544] pp. on six 98-frame fiche.
1984
Geology of the Grand Canyon: An annotated bibliography, with an annotated catalogue
of Grand Canyon type fossils. Volume 2. Supplement to the annotated bibliography
(1857-1983), Supplement and revisions to the annotated catalogue. Geological Society
of America, Microform Publication 14, 229 pp. on three 98-frame fiche. [The PDF available online is absent pp. 17-48.]
1988
Geology of the Grand Canyon: An annotated bibliography, with an annotated catalogue
of Grand Canyon type fossils. Volume 3. Second supplement (to 1987) with an annotated
bibliography of the world literature on the Grand Canyon type fossil Chuaria circularis
Walcott, 1899, an index fossil for the late Proterozoic. Geological Society of America,
Microform Publication 17, 343 [355] pp. on four 98-frame fiche.
17
INTRODUCTION
1990
Geology of the Grand Canyon: An annotated bibliography, with an annotated catalogue
of Grand Canyon type fossils. Volume 4. Third supplement (to 1989), with supplement
to the annotated bibliography of the world literature on the Grand Canyon type fossil
Chuaria circularis Walcott, 1899. Geological Society of America, Microform Publication
20, 178 pp. on two 98-frame fiche.
1992
Geology of the Grand Canyon: An annotated bibliography, with an annotated catalogue
of Grand Canyon type fossils. Volume 5. Fourth supplement (to 1991) with second supplement to the annotated bibliography of the world literature on the Grand Canyon type
fossil Chuaria circularis Walcott, 1899. Geological Society of America, Microform Publication 23, 234 pp. on three 98-frame fiche.
1992
The Grand Canyon fossil record; a source book on paleontology of the Grand Canyon and
vicinity, northwestern Arizona and southeastern Nevada. Bibliography; indexes to taxa,
stratigraphic records, localities, and repositories of type, figured, and cited specimens.
Geological Society of America, Microform Publication 24, 1,008 pp. on eleven 98-frame
fiche.
18
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
In this checklist the itemized species-level names of organisms are as originally erected by
the authors. This is a historical record of nomenclature, not one of taxonomic revision.
For convenience, the systematic levels of phylum/division, class, order, and family are
listed for each enumerated taxon. Refer to the Taxonomic List herein for other details.
Repository data for type material are as published by the original authors. The standard
acronyms for these repositories, when used, are not spelled out (again, as published).
Because locality data were not provided in any regular format by the original authors,
with wide variances in the amount of data provided, direct quotations have been preferred
in this checklist, which thus communicate the author’s (or publisher’s) style and the precision of data so conveyed. Some authors used the scientific symbols for male ( ♂) and female
(♀), which uses are retained in the quotations. This information has not been editorially
revised in the checklist; doing so would create an artificial sense of conformity that could
obscure the original precision and overall quality of the data
Throughout this checklist all text within [square brackets] has been inserted by the
compiler, except where noted otherwise.
________________________________________________________________________________
FORMAT FOR THE ANNOTATED CHECKLIST
Author(s)
Year
Bibliographical citation.
SEQUENTIAL #
*
Taxonomic name [as published], TAXONOMIC LEVEL (page and illustration
data, if from among many taxa in the publication)
Taxonomic Phylum: Class: Order: Family [placed before the sequential number if there are
multiple taxa under the same family]
Data about types and localities, usually quoted from the original publication. Principal
names of localities are highlighted in bold. (Refer to the respective separate lists of
localities in this volume for other details.) Notes are added where appropriate.
* An
arrow appears in this position if it is a taxon for which at least some of the type material
recorded by the authors is from one or more localities that are within, or are with good probability
from within, the present boundaries of Grand Canyon National Park.
_________________________________________________________________________ _______
19
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Adams, C. F. [Charles Frederick]
1903
Dipterological contributions. Kansas University Science Bulletin, [series 2], 2(2) (June):
21-47.
Ceratopogon dimidiatus, NEW SPECIES (p. 27); not illustrated
1
Arthropoda: Insecta: Diptera: Ceratopogonidae (biting midges)
Syntypes: “Four specimens; Grand Canon, Arizona”; all females.
________________________________________________________________________________
Adams, James K., and Lafontaine, J. Donald
2009
A new species of Plagiomimicus Grote (Noctuidae: Stiriinae) from northern Arizona and
southeastern Utah. Journal of the Lepidopterists’ Society, 63(3): 173-176, figs. 3, 4, 7,
8, 10.
2
Plagiomimicus kathyae,
NEW SPECIES
Arthropoda: Insecta: Lepidoptera: Noctuidae (owlet moths)
Holotype male (CNC [Canadian National Collection]) and 18 paratypes (7 males, 11 females, CNC,
personal collection of J. K. Adams, and USNM). Type material also includes specimens from Moab,
Utah. “The precise location for collection of the Arizona specimens is the Cameron Trading Post,
Cameron, AZ, at 35º 52’ 30” N, 111º 24’ 47” W, just south of the Little Colorado River and just
W of Hwy. 89”. Holotype collected by J. K. Adams, 7 September 1995; paratypes collected by
Adams, 7 September 1995, 1 September 1996. Text of species description (p. 174) indicates that
holotype male is illustrated in fig. 4, but figure legends (p. 175) indicate holotype is in fig. 3 and
a female paratype is in fig. 4.
_________________________________________________________________________ _______
Alexander, Charles P.
1946
Records and descriptions of North American crane-flies (Diptera). Part VI. Tipuloidea of
Arizona, New Mexico and Trans-Pecos Texas, 1. American Midland Naturalist, 35(2)
(March): 484-531.
3
Tipula (Lunatipula) kaibabensis,
NEW SPECIES
(pp. 506-508, fig. 7 [p. 507])
Arthropoda: Insecta: Diptera: Tipulidae (crane flies)
Holotype
male
from
“Kaibab
Plateau,
north
rim
of
the
Grand
Canyon,
Arizona
[undiscriminated], altitude 8,000 ft., June 18, 1942 (C. P. Alexander)”. Three male “paratopotypes
[paratypes] June 17-18, 1942 (C. P. & M. M. Alexander)”.
________________________________________________________________________________
20
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Al-Shehbaz, Ihsan Ali
1973
The biosystematics of the genus Thelypodium (Cruciferae). Contributions from the Gray
Herbarium of Harvard University, (204) (October 1): 3-148.
Thelypodium integrifolium (Nuttall) Endlicher subsp. longicarpum, NEW SUBSPECIES
4
(pp. 111-112, Pl. 7 [p. 36])
Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards)
Holotype (G) from “ca. 0.5 mile below Indian Garden [Havasupai Gardens] on Bright Angel
Trail, Grand Canyon National Park”, September 13, 1969, Ihsan & Mona Al-Shehbaz 6991;
“isotypes to be distributed”.
________________________________________________________________________________
Atwood, N. Duane
2007
Six new species of Phacelia (Hydrophyllaceae) from Arizona and New Mexico. Novon,
17(4): 403-416.
Trachyophyta: Magnoliopsida: Boraginales: Hydrophyllaceae (waterleafs)
5
Phacelia furnissii,
NEW SPECIES
(pp. 408-410, fig. 3 [p. 409])
Holotype and isotypes from “Mohave Co., Arizona Strip, Grand Canyon-Parashant National
Monument, Andrus Canyon at rd. crossing, T32N, R10W, S6 & 7, 27 May 2003, N. D. Atwood
& L. C. Higgins 29469 (holotype, BRY; isotypes, ARIZ, ASU, BRY [5], K, MO, NY, OSC, RM, TEX, UNLV,
UNM, US)”, and paratypes from 22 locales on the Arizona Strip in the vicinities of Andrus
Canyon, Trail Canyon, Whitmore Wash, Parashant Canyon, and Hack Canyon.
6
Phacelia higginsii,
NEW SPECIES
(pp. 410-412, fig. 4 [p. 411])
Holotype and isotypes from “Mohave Co., Arizona Strip, 12.5 mi. SW from head of Trail Canyon
toward Andrus Canyon, 36°11.48′N, 113°19.10′W, 3 Sep. 2003, N. D. Atwood, B. Furniss &
L. C. Higgsins 29729 (holotype, BRY; isotypes, ARIZ, ASU, BM, BRY [3], Dixie State College, K, MO, NY,
RM, TEX, US)”, and paratypes from four locales on the Arizona Strip in the vicinities of Andrus
Canyon, and Parashant Canyon.
7
Phacelia hughesii,
NEW SPECIES
(pp. 412-414, fig. 5 [p. 413])
Holotype and isotypes from “Mohave Co., Arizona Strip, T34N, R11W, S8, E of Salt Spring S
base of Poverty Mtn., 20 Sep. 2001, N. D. Atwood & L. E. Hughes 27814 (holotype, BRY;
isotypes, ARIZ, ASU, BRY [2], Dixie State College, MO, NY, US )”, and numerous paratypes from various
locales on the Arizona Strip.
________________________________________________________________________________
21
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Atwood, N. Duane, and Welsh, Stanley L.
2013
New plant taxa from Colorado, New Mexico, and Arizona. Western North American
Naturalist, 73(1): 113-115.
8
Xylorhiza tortifolia (Torrey & Gray) Greene var. parashantensis,
NEW VARIETY
(p. 115); not
illustrated
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
Holotype from “Grand Canyon–Parashant National Monument, Cunningham Canyon, T33N,
R14W, S30/SE, Yucca, Acacia, Fouquieria, Ferrocactus community, at 900 m elevation, on
limestone ledges and talus, N.D. Atwood & M. Madsen 26586, 29 March 2001. Holotype BRY!”
Paratype from “Grand Canyon–Parashant National Monument, 1.5 mi due west of Cane
Spring, Sandstone Canyon, with Lycium, Yucca, Agave, Rhus, and Fraxinus, at 1000 m
elevation. N.D. Atwood & L.C. Higgins 26927, 12 April 2001, BRY!”
________________________________________________________________________________
Augustson, G. F.
1943
A new subspecies of Orchopeas sexdentatus (Baker) (Siphonaptera: Dolichopsyllidae).
Bulletin of the Southern California Academy of Sciences, 42(1): 49-51, Pl. 5 (p. 51).
Orchopeas sexdentatus neotomae, NEW SUBSPECIES
9
Arthropoda: Insecta: Siphonaptera: Ceratophyllidae (rodent fleas)
“Holotype: a ♀ from Neotoma lepida devia Goldman, collected by R. L. Rutherford, South Entrance
Grand Canyon National Park, Coconino Co., Arizona, June 5, 1942. Deposited in the Allan
Hancock Foundation, University of Southern California. Los Angeles, California.
Allotype: a
♂
collected and deposited with the holotype as above.”
________________________________________________________________________________
Ball, Carleton R.
1921
10
Undescribed willows of the section Cordatae. Botanical Gazette, 71(6) (June): 426-437.
Salix lutea ligulifolia, NEW VARIETY (pp. 428-430); not illustrated
Trachyophyta: Magnoliopsida: Malpighiales: Salicaceae (willows)
“Referred specimens” [= isotypes] include “Grand Canyon, Indian Gardens [Havasupai Gardens],
alt. 3800 ft., E. A. Goldman 2237, August 25, 1913 (N [U.S. National Herbarium], twig densely
pubescent).”
________________________________________________________________________________
22
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Ball, E. D.
1933
The genus Myndus Stal in North America (Homoptera Fulgoridae). Journal of the
Washington Academy of Sciences (Washington, D.C.), 23(10) (October 15): 478-484.
[Myndus Stål.]
11
Myndus yuccandus,
NEW SPECIES
(p. 482); not illustrated
Arthropoda: Insecta: Hemiptera: Cixiidae (planthoppers)
“Holotype
♂ and one paratype male taken from
Yucca at the Grand Canyon Bridge, Ariz., Aug.
30, 1930 by the writer.”
[Note: Kramer (1979) examined the type material, with the following observations regarding the
locality, without further specifics: “Holotype male, Grand Canyon, Arizona, 4 August 1930, E.D.
Ball. The information published with the original description is slightly different from that on the
labels of the holotype; this is probably due to a lapsus by Ball. The previously published locality
was given as ‘Grand Canyon Bridge’ and the date of collection ‘30 August 1930’. One paraty pe
male with same data. Both are in the collection of the USNM.” (no further specifics by Kramer).
Ball’s particular note of “Grand Canyon Bridge” is surely Navajo Bridge, by which name it was
also known. The dates, however, both may be correct, and may reflect a bridge crossing on both
dates as a part of his travels in the field, although it does not resolve the immediate problem with
the dates ascribed to the collection of M. yuccandus. Compare data with Hysteropterum cornutum
var. utahnum Ball, 1935, and with Yucanda miniata Ball, 1937, below.]
[Reference: Kramer,
James P., 1979, Taxonomic study of the planthopper genus Myndus in the Americas (Homoptera:
Fulgoroidea: Cixiidae). American Entomological Society, Transactions, 105 (September): 301389. See Myndus yuccandus Ball (p. 328, figs. 54-56 [p. 327]; figure legend, p. 326, indicates
illustrations from paratype specimen).]
_______________________________________________________________
1935
Some new Issidae, with notes on others—(Homoptera–Fulgoridae). Bulletin of the
Brooklyn Entomological Society, 30(2) (April): 37-41.
12
Hysteropterum cornutum var. utahnum,
NEW VARIETY
(p. 38); not illustrated
Arthropoda: Insecta: Hemiptera: Issidae (planthoppers)
“Holotype
♀, allotype ♂, and 10 paratypes, taken by the writer near the Grand Canyon, August
1, 1930.”
[Note: Regarding the locality and date of collection, compare the comments with Myndus
yuccandus Ball, 1933, above, and with Yucanda miniata Ball, 1937, below, from which we glean
that he was present at Navajo Bridge twice in August 1930. However, it is not possible to discern
from these data whether “near the Grand Canyon” pertains to the North Rim or South R im, or in
fact just where in the area the specimens were taken.]
_______________________________________________________________
23
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1937
Some new Fulgoridae from the western United States. Bulletin of the Brooklyn
Entomological Society, Bulletin, 32(5) (December): 171-183.
13
Yucanda miniata,
NEW SPECIES
(p. 175); not illustrated
Arthropoda: Insecta: Hemiptera: Dictyopharidae (planthoppers)
“Holotype
♀,
allotype
♂,
and four paratypes, September 17, 1932, eight, August 4, 1930, and
two, August 31, 1935, all taken by the writer from a small shrub that looks like a dwarf mesquite,
at the Grand Canyon Bridge, Arizona.”
[Note: Regarding the locality and date of collection, compare the comments with Myndus
yuccandus Ball, 1933, and with Hysteropterum cornutum var. utahnum (above), from which we
glean that he was present at Navajo Bridge twice in August 1930.]
________________________________________________________________________________
Ball, E. D., and Beamer, R. H.
1939
A revision of the genus Athysanella and some related genera (Homoptera-Cicadellidae).
Kansas University Science Bulletin, 26(1) (October 1): 5-82 (including Pls. 1-12).
Arthropoda: Insecta: Hemiptera: Cicadellidae (leafhoppers)
14
Athysanella fredonia, NEW SPECIES (pp. 12-13, Pl. II, fig. 3 [p. 71), Pl. IX, fig. 3 [p. 79])
“Holotype male, allotype female, and 3 males and 2 females, Fredonia, Ariz., August 6, 1930,
E. D. Ball; paratypes as follows: . . . 7 males and 8 females, Grand Canyon, Ariz., E. D. Ball
. . . .” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon
village.]
15
Athysanella globosa, NEW SPECIES (pp. 18-19, Pl. II, fig. 13 [p. 72], Pl. X, fig. 13 [p. 80])
“Holotype male, allotype female, 1 male and 3 female paratypes, Grand Canyon, Ariz., E. D.
Ball, and in his collection.” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or
to Grand Canyon village.]
16
Athysanella (Gladionura) casa, NEW SPECIES (p. 48, Pl. VI, fig. 52 [p. 76], Pl. XI, fig. 52
[p. 81])
(Holotype, allotype, and paratypes are from Flagstaff, Ariz.) “. . . other paratypes as follows: 2
males, 3 females, Grand Canyon, Ariz., August, 1930, E. D. Ball . . . numerous specimens [nontype], Grand Canyon, Ariz., August 11, 1927, R. H. Beamer . . . .” [It is not clear whether “Grand
Canyon, Ariz.” refers to the canyon or to Grand Canyon village.]
________________________________________________________________________________
24
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Baral, Hans-Otto; Weber, Evi; and Marson, Guy
2020
Monograph of Orbiliomycetes (Ascomycota) based on vital taxonomy. Luxembourg:
National Museum of Natural History Luxembourg, Parts I and II, pp. 1-952, 953-1752.
Ascomycota (sac fungi): Orbiliomycetes: Orbiliales: Orbiliaceae
This is a profusely documented and illustrated publication, chiefly comprising the descriptions of
new species (date of publication 30 October 2020). This citation lists those taxa of interest to this
checklist that are typified. Only page numbers are given; pertinent enumerated plates of illustrations are embedded within those page ranges so are not cited separately. Locality data are given
here only in condensed format, documenting just the geographic locale, although each published
description adds information such as altitude, host material, collector, date of collection, and
repository and catalog number of the material.
In Baral et al.’s publication, “included material” is indicated to be distinct from the primary types,
thus in terms of botanical type nomenclature they are “ PARATYPES” (the term differs from the
homonym used in zoological nomenclature). Take note as well that the authors also list material
that is specifically “ not included ”; these references are omitted here. (The “not included”
specimens “deviate from the established taxa only in some minor points . . . [though] mentioned
under that taxon which appears to be most closely related, but they are not included in a
description, nor are they proposed as a separate new species.” [p. 17].)
All localities are specifically noted by Baral et al. as being in Arizona, although some locality
references to “Grand Canyon” are, confusingly, neither within the physiographic canyon nor within
the national park but are on adjacent national forest and other lands, sometimes at quite some
distance from the canyon or the national park. In the tables of localities in the present volume,
some interpretive statements have been included in order to clarify this; specifically,
measurements have been made from a map that reveal that linear distances from geographic
locales are mostly along the principal highways of the area, which information is added
notationally to the original locality data. Also, a few localities northwest of Page, Arizona, when
plotted on the map actually show that they are some distance into Utah, along U.S. Route 89;
these records are thus omitted here as being extralimital to this checklist.
The authorities (authors) of each species are as given in the publication; each thus is in turn “in”
Baral, Weber & Marson, as so noted in the taxonomic and locality lists herein. Species are listed
here in the same order as in which they appear in the publication.
17
Orbilia purshiae Baral & G. Marson, NEW SPECIES (pp. 466-468)
Holotype from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia, 13 km NNW of Jacob
Lake”; included material [paratype(s)] from “93 km SSE of Fredonia, S of North Rim”,
“Coconino Plateau, 7.5 km ESE of Grand Canyon Village, close to South Rim”.
18
Orbilia ocellata Baral, G. Marson & E. Weber,
NEW SPECIES
(pp. 476-481)
Included material [paratype(s)] from “Canyonlands, 45 km SW of Page, 26 km SW of Marble
Canyon” and “Grand Canyon, Coconino Plateau, 3 km N of Tusayan”.
25
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
19
Orbilia magnifica Baral & G. Marson,
NEW SPECIES
(pp. 509-511)
Holotype and isotypes from “Grand Canyon, Kaibab Plateau, 37 km SE of Fredonia, S of Jacob
Lake”; included material [paratype(s)] from “3 km N of Kaibab Lodge”.
20
Orbilia phanosoma Baral & G. Marson, NEW SPECIES (pp. 514-515)
Included material [paratype(s)] from “Grand Canyon, Coconino Plateau, 3 km N of Tusayan”
and “3 km S of Tusayan”.
21
Orbilia multiphanosoma Baral & G. Marson, NEW SPECIES (pp. 515-517)
Holotype from “Grand Canyon, Coconino Plateau, 3 km N of Tusayan”.
22
Orbilia arizonensis Baral & G. Marson, NEW SPECIES (pp. 553-556)
Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia,
13 km NNW of Jacob Lake”, “93 km SSE of Fredonia, S of North Rim”, and “Coconino
Plateau, 7.5 km ESE of Grand Canyon Village”.
23
Orbilia navajoana Baral & G. Marson, NEW SPECIES (pp. 571-573)
Included material [paratype(s)] from “Grand Canyon, Coconino Plateau, 3.5 km N of Tusayan”
and “3 km N of Tusayan”.
24
Orbilia spermoides Baral & G. Marson, NEW SPECIES (pp. 734-737)
Holotype from “Coconino Plateau, 3 km N of Tusayan”; included material [paratype(s)] from
“Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia, 13 km NNW of Jacob Lake”,
“Coconino Plateau, 8.5 km NNE of Tusayan, Mather Point” and “3 km N of Tusayan”.
25
Orbilia ophiosoma Baral & G. Marson, NEW SPECIES (pp. 1050-1052)
Included material [paratype(s)] from “Grand Canyon, Coconino Plateau, 3.5 km N of Tusayan”
and “3 km S of Tusayan”.
26
Orbilia maeandrina Baral & G. Marson, NEW SPECIES (pp. 1075-1078)
Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia,
13 km NNW of Jacob Lake”, “93 km SSE of Fredonia, S of North Rim”, “3 km N of Kaibab
Lodge”, “Coconino Plateau, 15 km ESE of Grand Canyon Village, Grandview Point”, and “8.5
km ESE of Grand Canyon Village, close to South Rim”.
27
Orbilia multimaeandrina Baral & G. Marson,
NEW SPECIES
(pp. 1078-1082)
Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia,
13 km NNW of Jacob Lake”.
28
Orbilia flexisoma Baral & G. Marson, NEW SPECIES (pp. 1095-1098)
Holotype from “Coconino Plateau, 3 km S of Tusayan); included material [paratype(s)] from
“Grand Canyon, Kaibab Plateau, 93 km SSE of Fredonia, S of North Rim”, “Coconino Plateau,
3.5 km N of Tusayan”, and “3 km N of Tusayan”.
26
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Orbilia delphinus Baral & G. Marson, NEW SPECIES (pp. 1105-1110)
29
Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 93 km SSE of Fredonia,
S of North Rim”, “Coconino Plateau, 15 km ESE of Grand Canyon Village, Grandview Point”,
and “3 km N of Tusayan”.
Orbilia macrodelphinus Baral & G. Marson, NEW SPECIES (pp. 1113-1116)
30
Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 28 km ESE of Fredonia,
13 km NNW of Jacob Lake” and “Coconino Plateau, 3.5 km N of Tusayan”.
31
Orbilia multitrapezoidea Baral & G. Marson,
NEW SPECIES
(pp. 1117-1120)
Holotype from “Grand Canyon, Coconino Plateau, 3 km S of Tusayan”.
Orbilia calyptrata Baral & G. Marson, NEW SPECIES (pp. 1187-1189)
32
Included material [paratype(s)] from “Grand Canyon, Coconino Plateau, 3.5 km N of Tusayan”
and “3 km S of Tusayan”.
Orbilia subovoidea Baral, Matočec & E. Weber, NEW SPECIES (pp. 1257-1260)
33
Included material [paratype(s)] from “Grand Canyon, Kaibab Plateau, 93 km SSE of Fredonia,
S of North Rim”.
________________________________________________________________________________
Barber, H. G. [Harry Gardner]
1938
A new genus and species of the subfamily Triatominae (Reduviidae: Hempitera).
Proceedings of the Entomological Society of Washington (Washington, D.C.), 40(4)
(April): 104-105.
Paratriatoma,
34
NEW GENUS
Paratriatoma hirsuta, NEW SPECIES; not illustrated
Arthropoda: Insecta: Hemiptera: Reduviidae (assassin bugs)
(The “type male” [holotype] is from Mojave, Calif., USNM 52747; disposition of other material not
indicated.) “Paratypes: Two males, Phanton Ranch [Phantom Ranch is consistently misspelled
in this paper], Grand Canyon, Ariz., May, 1929 (Vernon Bailey) . . . one nymph, Phanton Ranch,
Grand Canyon, Ariz., May 1929 (Vernon Bailey) . . . .”
________________________________________________________________________________
27
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Barneby, Rupert C.
1944
Alice Eastwood Semi-Centennial Publications. No. 3. Pugillus astragalorum alter.
Proceedings of the California Academy of Sciences, Series 4, 25(3) (June 1): 147-170, Pl.
17.
Astragalus bryantii, NEW SPECIES (pp. 156-157, Pl. 17, figs. 10-18 [pp. 168-169])
35
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
Type [holotype] from “the head of Phantom Canyon in the Grand Canyon of the Colorado River,
Coconino Co., 15 Dec. 1939. Collected by Dr. H. C. Bryant of the National Park Service in whose
honor it is named. Type in Herb. Calif. Acad. Sci. No. 293940. Also collected [= paratype] in
sand at the mouth of Hermit Creek in the Grand Canyon, 10 April 1917, Eastwood 5991 (G, F).”
_______________________________________________________________
1945
Pugillus astragalorum IV: The Section Diplocystium. Leaflets of Western Botany, 4(5)
(March): 65-147 [including Pls. 1-4].
36
Astragalus lentiginosus Dougl. var. oropedii, NEW VARIETY (pp. 135-137); not illustrated
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
“Type [holotype].
Kaibab Trail to Roaring Springs [North Kaibab Trail], Grand Canyon
National Park, Coconino County, Arizona, 22 September 1938, fruct. Eastwood & Howell No. 7064
(Herb. Calif. Acad. Sci. No. 262056). Also ibid., 23 June 1933, flor., Eastwood & Howell No. 1054
(Herb. Calif. Acad. Sci. No. 211208, cotype).”
[Barneby’s use of “cotype” is an older term often
mistakenly used to indicate an isotype or paratype; in this case, the “ibid.” specimen is an
isotype.]
[Take note also of Jason Andrew Alexander (2009, The types of Astragalus Section Diphysi
(Fabaceae),
a complex endemic to
western North
America,
Part
I: Lectotypifications,
epitypifications, and new combinations of several taxa. Botanical Research Institute of Texas,
Journal, 3(1) (July): 211-218). Alexander indicates the following information regarding the type
of A. l. var. oropedii from North Kaibab Trail: effective lectotypification later by Barneby (1989),
epitype designated by Alexander from among Barneby’s syntypes. However, note that Barneby,
1945, did indeed designate the “type” [holotype] and that Alexander may have mistakenly
interpreted Barneby’s “cotype” (an older, often misued term) to mean a syntype; thus the
“effective lectotypification” by Barneby, as noticed by Alexander, likewise was a misinterpretation;
see above).]
[Barneby, 1989, is cited by Alexander as in Cronquist, A., Holmgren, A. H.,
Holmgren, N. H., Reveal, J. L., and Holmgren, P. K. (eds.), Intermountain Flora. Volume III, Part
B (New York Botanical Garden, Bronx, New York).]
_______________________________________________________________
28
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1948
Pugillus astragalorum IX: Novelties in Batidophaca Rydb. Leaflets of Western Botany,
5(5) (February): 82-89.
Astragalus cremnophylax, NEW SPECIES (pp. 83-85); not illustrated
37
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes) [Sentry milkvetch; endemic and
endangered.]
Type locality “about 2 miles west of El Tovar, Coconino Co., alt. 7050 ft., 3 June 1947, fr.,
Ripley & Barneby No. 8473. Type [holotype] in Herb. Calif. Acad. Sci., No. 336060. Grand Canyon
(‘at the end of the railroad on sandy ledges’), Jones in 1903, fl. (CAS, fragments, PO).” (p. 84).
“In its type-locality, A. cremnophylax is confined to a narrow trip of limestone pavement
immediately overlooking the Grand Canyon . . . Whence the name from the Greek, meaning
watchman or overseer of the chasm. ”
_______________________________________________________________
1949
Pugillus astragalorum X: New species, varieties and combinations. American Midland
Naturalist, 41(2) (March): 496-502.
38
Astragalus wootoni var. endopterus,
NEW VARIETY
(p. 498); not illustrated
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
From “sandy bed of the Little Colorado River, at Cameron, Coconino Co., alt. 3700 ft., 5 June
1947, fl. & fr., Ripley & Barneby No. 8491. Type in herb. Calif. Acad. Sci. No. 337308.”
_______________________________________________________________
1979
Dragma hippomanicum IV. New taxa of Astragalus sect. Humillimi. Brittonia, 31(4):
459-463.
39
Astragalus cremnophylax Barneby var. myriorrhaphis,
NEW VARIETY
(p. 463, fig. 2 [p.
462])
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
“Type [holotype]: United States. Arizona. Coconino Co.: locally common in cracks and crevices
of limestone cliff and rimrock, 1890 m (6200 ft), Buckskin Mt. [Kaibab Plateau] S of
Honeymoon Trail, 28 km (17.5 mi) airline distance E of Fredonia (T41N, R2E, S29/30)
[an area of Forest Service roads], 23 May 1979, N. H. Holmgren, P. K. Holmgren & R. C. Barneby
9145 (HOLOTYPE: NY; ISOTYPES: BRY, UTC ad others to be distributed).”
________________________________________________________________________________
29
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Barnes, William, and Benjamin, F. H.
Notes on diurnal Lepidoptera, with additions and corrections to the recent “List of Diurnal
1926
Lepidoptera”. Bulletin of the Southern California Academy of Sciences, 25(3)
(September/December): 88-98.
Coenonympha fureae, NEW SPECIES (p. 90); not illustrated
40
Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals)
“Type locality: Grand Canyon, Ariz.” “Holotype
♂, Allotype ♀, 3 ♀ Paratypes; no dates except
on one female, 1-7 June.” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or
to Grand Canyon village.]
Cercyonis damei , NEW SPECIES (p. 90); not illustrated
41
Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals)
“Type locality: Grand Canyon, Ariz.” “Holotype
♂, 1-7 June; 4 ♂Paratypes, one only dated 8-
15 June.” [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or to Grand Canyon
village.]
_______________________________________________________________________________
Baroni, Timothy J.;
Beug, Michael W.; Cantrell, Sharon A.; Clements, Teresa A.; Iturriaga, Teresa;
Læssøe, Thomas; Holgado Rojas, María E.; Aguilar, Frank M.; Quispe, Miguel O.;
Lodge, D. Jean; and O’Donnell, Kerry
2018
Four new species of Morchella from the Americas. Mycologia, 110(6) (December): 12051221.
42
Morchella kaibabensis Beug, T.A. Clem. & T.J. Baroni,
NEW SPECIES
(pp. 1208-1213, figs. 3A-
D
Ascomycota: Pezizomycetes: Pezizales: Morchellaceae
“Typification: USA. Arizona: Coconino County, along Forest Road 293, Kaibab Plateau, North
Rim, Grand Canyon National Park, 36.40806, –112.26692, 2700 m, with Picea engelmannii
and Picea pungens, 21 May 2017, T.A. Clements & D.M. Fulton TAC-1708 (KOD1793) (holotype
ARIZ AN043595). Ex-type culture: NRRL 66753. GenBank: ITS = MH014728; TEF1 = MH014722;
RPB1 = MH014733; RPB2_a = MH014738; RPB2-b = MH014744.” [Locality is ca. 7.3 miles west
of Kaibab Lodge, on the Kaibab National Forest.]
________________________________________________________________________________
30
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Barr, M. E.
1989
The Venturiaceae in North America: Revisions and additions. Sydowia, 41: 25-40.
Protoventuria arizonica, NEW SPECIES (pp. 35, 37, fig. 17 [p. 36])
43
Ascomycota: Dothidiomycetes: Pleosporales: Venturiaceae
“Holotypus in foliis emortuis Cercocarpi ledifolii NUTT. ‘North Rim of Grand Canyon, Arizona, 18
Aug. 1973’ a M. E. BARR n. 6096a lectis in Herb. MASS depositus.” [translation: Holotype from
decaying leaves of Cercocarpi ledifolii Nutt., from “North Rim of Grand Canyon, Arizona, 18
Aug. 1973”, by M. E. Barr n. 6096a deposited in Herb. MASS.] [It is not certain whether “North
Rim of Grand Canyon” refers to the North Rim of the canyon or to North Rim, Arizona.]
________________________________________________________________________________
Barr, William F.
1972
New species of North American Acmaeodera (Coleoptera: Buprestidae). Arquivos
do Museu Boçage (Lisboa), Series 2, 3(7): 145-201.
Acmaeodera pletura, NEW SPECIES (pp. 164-166, fig. 12 [p. 199])
44
Arthropoda: Insecta: Coleoptera: Buprestidae (jewel beetles)
(Holotype from Lee Canyon, Clark Co., Nevada; California Academy of Sciences.)
Paratypes
include “. . . north rim, Grand Canyon National Park, Arizona, June 15, 1930 (C. Kelley) 1
specimen; south rim of Grand Canyon, Arizona, July 8, 1967, on blossom of Cowania
stansburiana (W. F. Barr), 1 specimen. [All] Paratypes in the collections of the California Academy
of Sciences, R. L. Westcott and W. F. Barr.”
________________________________________________________________________________
Bauer, David L.
1955
A new race of Papilio indra from the Grand Canyon region. The Lepidopterists’ News,
9(2/3): 49-54, plate (p. 52).
45
Papilio indra kaibabensis, NEW SUBSPECIES
Arthropoda: Insecta: Lepidoptera: Papilionidae (swallowtails)
“Holotype male: (expanse 76 mm.) Bright Angel Point, Grand Canyon, Coconino Co., Ariz.,
August 1953, leg. Ernst Christensen.
Allotype female: (expanse 86 mm.) same locality and
collector as holotype, 14 August 1953. Paratypes: 9 males and 3 females. Data as follows (all
but one male from the Grand Canyon National Park): 1 male, North Rim, 3 August 1938, leg.
Louis Schellback [sic, Schellbach]; 1 male, Yavapai Point, South Rim, 22 August 1944, leg. Louis
Schellback; 1 male, Bright Angel Point, North Rim, 13 August 1951, leg. Ernst Christensen; 1
male, Bright Angel Point, North Rim, 7 August 1953, leg. Ernst Christensen; 1 male, Roaring
Springs, North Rim, 4 August 1953, leg. Ernst Christensen; 1 male, near Ryan Ranger Station,
Kaibab Plateau, Coconino Co., Ariz., 1 July 1952, leg. D. L. Bauer; 1 female, data same as for
allotype; 1 female, data same as for holotype.”
31
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Allotype and holotype, respectively, are illustrated in the plate, the third pair of figures (upperside
of specimens) and the fourth pair of figures (underside of specimens).
“The holotype and allotype and four male paratypes will be returned to the Naturalist Work Shop
Collection, Grand Canyon National Park; from there I believe the holotype and allot ype will be
sent to the U. S. National Museum. One male and one female paratype will be returned to Dr.
Rodeck and the University of Colorado Collection, one male paratype to the Yale Peabody Museum,
and one male paratype to the Los Angeles County Museum. The remainder are in the author’s
collection.”
________________________________________________________________________________
Baxter, E. M.
1933
Opuntia aurea—a new species. Cactus and Succulent Journal (Cactus and Succulent
Society of America), 5: 489-490.
46
Opuntia aurea,
NEW SPECIES ;
illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
“Type Locality: ½ mile north of Pipe Springs [Pipe Spring] on the Kaibab Indian
Reservation, Arizona.
Type Specimen [holotype]: Two joints collected in 1930 by Percy and
Helen McCabe at the type lcoality and placed in the Dudley Herbarium at Stanford University as
No. 213750, October, 1933.” [The illustration is a photo of a living plant, credited to “McCabe”;
it is not indicated whether the photo depicts the source plant of the holotype. In addition to the
type from Percy and Helen McCabe, Baxter also credits without further detail (p. 490) “Howard
Gates for a specimen and much information on distribution.”]
Discussion begins: “A new species, introduced first by the McCabe Gardens of San Diego, and
later by Howard Gates of Anaheim, is here described for the first time. It has appeared in the
McCabe catalog both under the name shown and as Opuntia basilaris aurea, and has been sold by
Mr. Gates as ‘yellow-flowered basilaris’. The McCabe name is taken as the specific name for the
species, denoting the gold color of the flower.”
________________________________________________________________________________
Beamer, R. H.
1948
Some new species of Delphacodes (Homoptera, Fulgoridae, Delphacinae); Part IV.
Journal of the Kansas Entomological Society, 21(3) (July): 96-110.
47
Delphacodes apicata, NEW SPECIES (pp. 100-101, Pl. VIII, figs. 37, 37a, 37b [p. 107;
legend p. 106])
Arthropoda: Insecta: Hemiptera: Delphacidae (planthoppers)
(Under the heading, “Brachypterous Form” (pp. 100-101), Beamer designates a holotype,
allotype, and two paratypes from “San Francisco Mt., Ariz.”; deposited in “Snow Entomological
Collections and United States National Museum”.) Under the heading, “Macropterous Form” (p.
101), Beamer lists: “Holomorphotype , Grand Canyon, Ariz., Aug. 11, 1927, R. H. Beamer” with
an “allomorphotype” and “paramorphotypes” from San Francisco Mt. and Flagstaff, Ariz.
32
It is
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
unclear whether these may also be considered paratype material, based on provisions of earlier
editions of the International Code of Zoological Nomenclature. The figures do not indicate which
specimens are illustrated. [It is not clear whether “Grand Canyon, Ariz.” refers to the canyon or
to Grand Canyon village.]
________________________________________________________________________________
Belk, Denton, and Fugate, Michael
2000
Two new Branchinecta (Crustacea: Anostraca) from the southwestern United States.
Southwestern Naturalist, 45(2) (June): 111-117.
Arthropoda: Branchiopoda: Anostraca: Branchinectidae (brine shrimp)
48
Branchinecta kaibabensis,
NEW SPECIES
(pp. 115-117, figs. 2b [p. 114], 3a-e [p. 116]
“Holotype male (USNM 260875) and paratypes of both sexes (USNM 260876) from DB 232 are at
the National Museum of Natural History, Smithsonian Institution, Washington, D.C.”
“Type Locality—A natural pool 0.6 km east of Arizona Highway 67 just northeast of Forest
Road 213 in the Kaibab National Forest . . . (36°30′38″N, 112°07′55″W). Forest Road
213 intersects Arizona Highway 67, 30.6 km south of the city of Jacob Lake.” “DB 232 pool in
depression 0.6 km east of AZ 67 on Forest Road 213, Kaibab National Forest, Coconino Co.,
Arizona, collector D. Belk, 22 May 1974.”
________________________________________________________________________________
Benson, Bernard W.
1957
A new cactus from Arizona. Cactus and Succulent Journal (Cactus and Succulent Society
of America), 29(5) (September/October): 136-137.
49
Pediocactus paradinei,
NEW SPECIES ;
illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
Named as “Pediocactus (?) paradinei B. Benson, sp. nov.” From House Rock Valley (without
further specific locale).
Holotype, “Boyce Thompson Southwestern Arboretum Herbarium, No. Bwb 8-1956-1”. Isotype,
“Herbarium of Pomona College, No. 286,120”.
“Mr. N. A. Paradine collected the first plants in House Rock Valley, Coconino County, Arizona, in
the spring of 1956. These were found under the protection of small sagebrush plants. Later
others were found in clumps of perennial bunch grasses.”
Benson adds: “In August of 1956, a description of this plant was sent to the editor of the Cactus
and Succulent Journal for publication.
In October of 1956, Mr. W. T. Marshall of the Desert
Botanical Garden, Tempe, Arizona, described the same plant in the Saguaroland Bulletin 10: 8991. The author appreciates Mr. Marshall’s courtesy in stepping aside to permit the naming of
Pediocactus paradinei.” The item thus cited is “A New Genus of Cactuses for Arizona”, unsigned
but in Saguaroland Bulletin (W. Taylor Marshall, editor), 10(8) (October 1956): 89-91, wherein it
33
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
appears only as Pediocactus sp., based on gatherings in House Rock Valley by Gus and Sylvia
Hermann.
________________________________________________________________________________
Bequaert, J. [Joseph C.]
1932
The Nearctic social wasps of the subfamily Polybiinae (Hymenoptera; Vespidae).
Entomologica Americana, 13(3) (December): 87-149 (pagination includes Pls. 27-29).
Mischocyttarus flavitarsis var. kaibabensis, NEW VARIETY (p. 133); not illustrated [the
50
plates and text-figs. do not pertain to this new variety]
Arthropoda: Insecta: Hymenoptera: Vespidae (hornets)
“Bright Angel Trail, Grand Canyon, Cococino [sic] Co., one female, holotype (Cornell Univ.) and
one female paratype (M.C.Z[.]); both collected by R. C. Shannon, on the Cornell Biological
Expedition, August 4, 1917.” (The etymology of the varietal name is from its “perfect parallel in
color (or ‘mimic’) of Polistes canadensis var. kaibabensis Hayward, which occurs in the same
district.”) [Refer to Hayward (1932, no. 155 herein).]
________________________________________________________________________________
Beutenmüller, William
1907
Notes on a few North American Cynipidae, with descriptions of new species. Bulletin of
the American Museum of Natural History, 23: 463-466, Pl. 37.
Andricus wheeleri, NEW SPECIES (p. 464, Pl. 37, figs. 7-9)
51
Arthropoda: Insecta: Hymenoptera: Cynipidae (gall wasps)
“Coconino Forest, rim of the Grand Cañon, Arizona; altitude 7,000 feet. (William M. Wheeler.)
Types [syntypes]. Coll. Am. Mus. Nat. Hist.”
________________________________________________________________________________
Bohart, R. M.
1958
A new Priononyx and a key to the North American species (Hymenoptera: Sphecidae).
Bulletin of the Brooklyn Entomological Society, 53(4) (October): 90-93.
52
Priononyx subatrata, NEW SPECIES
Arthropoda: Insecta: Hymenoptera: Sphecidae (sphecid wasps)
(Holotype from Deep Springs, California.) Paratypes include specimens from “Grand Canyon
South Rim (M. A. Evans), Kaibab Forest (M. Wasbauer)”. The “Kaibab Forest” specimen may
or may not be from Kaibab National Forest units near Grand Canyon.
________________________________________________________________________________
34
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Bromley, Stanley W.
1940
New U.S.A. robber flies (Diptera: Asilidae). Bulletin of the Brooklyn Entomological
Society, 35(1) (February): 13-21.
Erax benedicti, NEW SPECIES (pp. 15-16, fig. 1 [p. 20])
53
Arthropoda: Insecta: Diptera: Asilidae (robber flies)
(Holotype from Winslow, Ariz.; allotype from Santa Rita Mountains, Ariz.; both in University of
Kansas Collection.)
Paratypes include “2 females, Grand Canyon, Ariz., June 12, 1937 (W.
Benedict)”. No further data regarding these paratypes. [It is not clear whether “Grand Canyon,
Ariz.” refers to the canyon or to Grand Canyon village.]
________________________________________________________________________________
Bunn, Ralph W.
1930
Notes on the genus Aphelonema Uhler with descriptions of new species (Homoptera,
Fulgoridae). Journal of the Kansas Entomological Society, 3(3) (July): 73-77.
Aphelonema convergens var. canyonensia, NEW VARIETY (p. 76); not illustrated
54
Arthropoda: Insecta: Hemiptera: Caliscelidae (planthoppers)
“Holotype; female, from vicinity of Grand Canyon, Arizona collected by Dr. [R. H.] Beamer
August 11, 1927. Allotype; male, collected at the same time and place by Dr. Beamer. Paratypes;
. . . one female from Grand Canyon, Arizona, taken August 11, 1927 by Dr. P. A. Readio; three
females from Grand Canyon, Arizona taken August 11, 1927 by L. D. Anderson; and six females
from the same locality and collected at the same time by Dr. Beamer. Types deposited in Snow
Entomological Collection.” [It is not clear whether “Grand Canyon, Arizona” refers to the canyon
or to Grand Canyon village.]
________________________________________________________________________________
Burke, H. E.
1907
A new buprestid enemy of Pinus edulis. (Melanophila pini-edulis, n. sp.) Proceedings of
the Entomological Society of Washington (Washington, D.C.), 9(1/4) (March/December):
117-118.
55
Melanophila piniedulis, NEW SPECIES; fig. 6
Arthropoda: Insecta: Coleoptera: Buprestidae (jewel beetles)
“Type [syntypes].—♀ and
♂, No. 11350, U. S. National Museum.”
One syntype is “One female taken by Mr. E. A. Schwarz from pinyon (Pinus edulis) at Bright
Angel Hotel (Grand Canyon P.O.), Arizona, on July 11, 1902.”
________________________________________________________________________________
35
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Casey, Thomas L.
1895
Coleopterological notices. VI. Annals of the New York Academy of Sciences, 8 (July):
435-838.
Arthropoda: Insecta: Coleoptera: Melyridae (soft-winged flower beetles)
56
Trichochrous incipiens,
NEW SPECIES
(pp. 489-490); not illustrated
“Arizona (near the Grand Cañon of the Colorado). Dr. T. Mitchell Prudden. The single type
[holotype] is a female . . . .”
57
Trichochrous reversus,
NEW SPECIES
(pp. 527-528); not illustrated
“Arizona (near the Grand Cañon of the Colorado). The single type [holotype] is a female
. . . ; it was collected and kindly given to me by Dr. T. Mitchell Prudden, of New York.”
_______________________________________________________________
1900
Review of the American Corylophidae, Cryptophagidae, Tritomidae and Dermestidae, with
other studies. Journal of the New York Entomological Society, 8(2) (June): 51-172.
Arthropoda: Insecta: Coleoptera: Dermestidae (dermestid beetles)
Cryptorhopalum pruddeni, NEW SPECIES (p. 156); not illustrated
58
“Arizona (Cañon of the Colorado River)—Dr. T. Mitchell Prudden.”
Orphilus aequalis, NEW SPECIES (p. 164); not illustrated
59
“Arizona (Cañon of the Colorado River)—Dr. Prudden.”
_______________________________________________________________
1907
Notes on Chalcolepidius and the Zopherini. The Canadian Entomologist (London), 39(2)
(February): 29-46.
Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles)
Zopherodes,
NEW GENUS
(p. 38)
Zopherodes lugubris, NEW SPECIES (p. 41); not illustrated
60
“Arizona (Grand Canyon of the Colorado).”
Zopherodes pruddeni, NEW SPECIES (p. 41); not illustrated
61
“Arizona (Grand Canyon of the Colorado), T. Mitchell Prudden.”
_______________________________________________________________
1908
A revision of the tenebrionid subfamily Coniontinæ. Proceedings of the Washington
Academy of Sciences (Washington, D.C.), 10 (April 25): 51-166.
Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles)
62
Discodemus brevipennis, NEW SPECIES (p. 61); not illustrated
“Arizona (Grand Canyon of the Colorado),—T. Mitchell Prudden.”
_______________________________________________________________
36
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1912
Studies in the Longicornia of North America. In: Casey, Thomas L., Memoirs on the
Coleoptera. III. Lancaster, Pennsylvania: New Era Printing Co., pp. 215-386.
Arthropoda: Insecta: Coleoptera: Cerambycidae (long-horned beetles)
Prionus spiculosus, NEW SPECIES (p. 240); not illustrated
63
“Arizona (Grand Cañon of the Colorado),—T. Mitchell Prudden.”
Prionus angustulus, NEW SPECIES (p. 241); not illustrated
64
“Arizona (Cañon of the Colorado River),—Prudden.”
Prionus terminalis, NEW SPECIES (pp. 243-244); not illustrated
65
“Arizona (Cañon of the Colorado River),—Prudden.”
Stenosphenus pruddeni, NEW SPECIES (pp. 346-347); not illustrated
66
“Arizona (Cañon of the Colorado),—T. Mitchell Prudden.”
_______________________________________________________________
1918
A review of the North American Bembidiinæ. In: Casey, Thomas L., Memoirs on the
Coleoptera. VIII. 1918. Lancaster, Pennsylvania: New Era Printing Co., pp. 1-223.
Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles)
Bembidion (Cyclolopha) occultum, NEW SUBGENUS, NEW SPECIES (p. 144); not illustrated
67
“Arizona (Grand Cañon of the Colorado),—T. Mitchell Prudden.”
________________________________________________________________________________
Chamberlin, Ralph V.
1940
New American tarantulas of the family Aviculariidae. Bulletin of the University of Utah,
30(13) (May 6), 39 pp.
Arthropoda: Euchelicerata: Areneae: Theraphosidae (tarantulas)
68
Aphonopelma behlei, NEW SPECIES (pp. 26-27); not illustrated
“Locality: Arizona: Coconino Co., Grand Canyon Village, 7000 ftt. el., 15 Sept., 1939. Two males
[syntypes] taken by Dr. W. H. Behle on a paved highway after a heavy rain.” (Additional syntypes
from Aztec, New Mexico.)
69
Aphonopelma phasmus, NEW SPECIES (p. 28); not illustrated
“Locality: Arizona: Grand Canyon, Phantom Ranch. 26 July. 1934.
One male [holotype]
collected by Dr. Lutz. American Museum collection.”
_______________________________________________________________
37
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1948
On some American spiders of the family Erigonidae. Annals of the Entomological Society
of America, 41(4): 483-562.
70
Tapinocyba kesimba,
NEW SPECIES
(pp. 552-553, Pl. 11, figs. 117-121 [p. 515, legend p.
514])
Arthropoda: Euchelicerata: Areneae: Linyphiidae (dwarf weavers)
“Type locality.—Arizona: Kaibab Forest (V. T. Ranch), September 4, 1931, 2 ♂ 6♀ [syntypes],
R. V. Chamberlin.” (“Other records” include “Kaibab Forest, September 4, 1931, 2 ♂, 6♀, R. V.
Chamberlin.)
________________________________________________________________________________
Chamberlin, Ralph V., and Ivie, Wilton
1942
A hundred new species of American spiders. Bulletin of the University of Utah, 32(13)
(June 30) [Biological Series, 7(1)], 117 pp.
71
Neoantistea coconino,
NEW SPECIES
(pp. 28-29, figs. 59, 60 [p. 95, legend p. 94])
Arthropoda: Euchelicerata: Areneae: Hahniidae (hahniid spiders)
“Type locality: W 112°:N 36°, Kaibab Forest, Arizona;
♂ holotype; June 14, 1934; W. Ivie and
H. Rasmussen, collectors.” [The overly generalized coordinates lie within Grand Canyon National
Park, just west of Grandview Point; in fact, by coincidence on a cliff face(!) just over the rim of the
canyon (https://confluence.org/confluence.php?id=91). The specific epithet and indication of
“Kaibab Forest” may point either to a locality on the nearby Kaibab National Forest or is only a
generalized term for the South Rim forest, sometimes called the Coconino forest, on the Coconino
Plateau.]
________________________________________________________________________________
Chandler, Peter
1993
The Holarctic species of the Mycetophila fungorum (De Geer) group (Diptera:
Mycetophilidae). British Journal of Entomology and Natural History, 6: 5-11.
72
Mycetophila neofungorum, NEW SPECIES (pp. 7-9, figs. 7-9 [p. 8])
Arthropoda: Insecta: Diptera: Mycetophilidae (fungus gnats)
“Holotype male USA, Arizona, Grand Canyon National Park (north rim), 15.vii.1954 (W. L.
Downes, Natural History Museum, London).” [Based on a unique museum specimen.]
________________________________________________________________________________
38
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Christy, Charlotte M.
1997
A new species of Mentzelia Section Bartonia (Loasaceae) from Arizona. Novon, 7(1): 2526.
73
Mentzelia collomiae,
NEW SPECIES ;
not illustrated
Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa)
(“[A]n edaphic endemic species restricted to cinder areas of the San Francisco Volcanic Field”,
holotype and isotypes from “ca. 3.5 mi. due N of Sunset Crater National Monument".) Paratypes
include various localities, the nearest of which pertinent to this bibliography (and at the limit of
its coverage) is “Red Mountain, 35 mi. NW of Flagstaff on Hwy. 180”.
________________________________________________________________________________
Chuang, Tsan Iang, and Heckard, Lawrence R.
1986
Systematics and evolution of Cordylanthus (Scrophulariaceae—Pedicularieae) (including
the taxonomy of subgenus Cordylanthus). Systematic Botany Monographs, Volume 10,
105 pp.
74
Cordylanthus wrightii kaibabensis,
NEW SUBSPECIES
(pp. 87-89, figs. 35i-n [p. 88])
Trachyophyta: Magnoliopsida: Lamiales: Orobanchaceae (broomrapes)
“TYPE: U.S.A. Arizona: Coconino Co., openings in Juniperus monosperma, occasional Pinus
ponderosa with Artemisia tridentata, 7.5 mi E of Jacobs Lake [Jacob Lake], 6400 feet, Heckard,
Bacigalupi & Chuang 3585 (holotype: JEPS!; isotypes: ARIZ! ASU! BRY! COLO! F! GH! ISU! MO! NY!
TEX! US! UTC!).”
“Representative Specimens [= paratypes]. U.S.A. Arizona: Coconino Co., House Rock Valley,
Smith s.n. (NY, PH); Kaibab Plateau near Jacobs Lake, Eastwood & Howell 6441 (DS, POM).”
________________________________________________________________________________
Clark, Curtis
1998
New names and combinations in Encelia frutescens sensu lato (Asteraceae: Heliantheae).
Aliso (Rancho Santa Ana Botanic Garden, Claremont, California), 17(2): 201-202.
75
Encelia resinifera subsp. tenuifolia, NEW SUBSPECIES (p. 202); not illustrated
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
“TYPE: United States. Arizona. Coconino Co: 4 miles down Supai Trail, Havasupai Canyon
[Havasu Canyon]. 3 June 1961. C. and C. Dailey s.n., MNA 2256/ B12,747. holotype: MNA!.”
[Note: The precise gathering locality may be within the boundaries of the Havasupai Indian
Reservation.]
“Subspecies tenuifolia occurs almost entirely within the Grand Canyon of the Colorado River in
northern Arizona, ranging from the Grand Wash Cliffs east to Marble Canyon, evidently following
the 1200 m contour.”
________________________________________________________________________________
39
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Clarke, J. F. Gates
1947
Notes on Oecophoridae, with descriptions of new species. Journal of the Washington
Academy of Sciences (Washington, D.C.), 37(1): 2-18.
Depressaria schellbachi, NEW SPECIES (pp. 10, 13; figs. 6-6a [p. 7], 13 [p. 12])
76
Arthropoda: Insecta: Lepidoptera: Oecophoridae (oecophorid moths)
“Type.— U.S.N.M. No. 58013. Type locality.—Shoshone Point, Grand Canyon, Ariz., elev. 7,050
feet. Food plant.—Lomatium macdougali Coult. and Rose.”
“Described from the
♂ type [holotype], five ♂ and three ♀ paratypes from the Grand Canyon
(June dates, 1944 and 1946, Louis Schellbach, 3d). Paratypes in the U.S. National and British
Museums and in the Grand Canyon National Park Collection.”
________________________________________________________________________________
Clover, Elzada U.
1942
A new species and variety of Sclerocactus from Arizona. American Journal of Botany,
29(2) (February): 172-173.
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
77
Sclerocactus havasupaiensis, NEW SPECIES; illustrated [fig. 1 (p. 172) shows “Actual type
specimen photographed in natural habitat at Havasupai Canyon, Arizona (Clover 5229 in part: there are
several cotypes”]
“The type specimen [holotype] (fig. 1) was collected by William Belknap, Jr., April 26, 1941, on
top of the Supai Formation in Havasupai Canyon [Havasu Canyon], Arizona, where the species
grows abundantly. Specimens [= paratypes] not in flower were previously collected by the author
in July, 1940, in Havasupai Canyon (Clover 5229) near Navajo Falls, and in Hualapai Canyon
[tributary to Havasu Canyon] (Clover 5100) on talus of the Supai Formation. A few plants were
seen on the floor of Cataract Canyon above Supai, September, 1941.”
Note: The precise gathering localities may be within the boundaries of the Havasupai Indian
Reservation.
78
Sclerocactus havasupaiensis var. roseus, NEW VARIETY; illustrated [figs. 3, 4 (p. 173) show
“Actual type specimen [holotype] (Clover 6403) photographed while alive at Botanical Garden, University
of Michigan, now dried and preserved as a herbarium specimen. Additional plants of the same collection
(cotypes [= istotypes]) are in the living collection (Bot. Gard. no. 18079).”]
“Specimen typicum [holotype] legit E. Clover & Wm. Belknap, Jr. (Clover 6403), in Havasupai
Canyon [Havasu Canyon], Arizona, siccatum est in Herbario, Universitatis Michiganensis.”
[comparable text in English not provided].
“This variety was found in association with S.
havasupaiensis on top of the Supai Formation in the same location in May 1941, and are in the
living collection (no. 18079) in the Botanical Gardens, University of Michigan.”
Note: The precise gathering locality may be within the boundaries of the Havasupai Indian
Reservation.
________________________________________________________________________________
40
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Cockerell, T. D. A. [Theodore Dru Alison]
1905
New American bees. Proceedings of the Biological Society of Washington (Washington,
D.C.), 18 (June 29): 177-184.
Triepeolus hopkinsi, NEW SPECIES (p. 184); not illustrated
79
Arthropoda: Insecta: Hymenoptera: Apidae (honey bees)
Apparently named on a single male specimen [holotype].
“Grand Canyon of the Colorado,
Arizona, August 3, 1904. (Webb). Received from Mr. Viereck, to whom it has been returned. It
is named after Professor Hopkins [A. D. Hopkins?], who sent it to Mr. Viereck, and who has done
good work in the region of the Grand Canyon.”
________________________________________________________________________________
Cole, A. C., Jr.
1936
Descriptions of seven new western ants. (Hymenop.: Formicidae.) Entomological News
(Academy of Natural Sciences of Philadelphia, Entomological Section), 47 (May): 118121.
80
Myrmecocystus melliger subsp. semirufus Emery var. romainei,
NEW VARIETY
(p. 120).
[Not valid as a quadrinomial taxon, and no identification of holotype; non Myrmecocystus
romainei Snelling, 1975]
Arthropoda: Insecta: Hymenoptera: Formicidae (ants)
“Described from a series of 54 workers [= syntypes] taken by Miss Marjorie Romaine, at
Cameron, Arizona. The holotype is in the author’s collection, and paratypes are to be deposited
in the collections of the U. S. National Museum and of Dr. C. H. Kennedy.”
________________________________________________________________________________
Coquillett, D. W.
1902
New acalyptrate Diptera from North America. Journal of the New York Entomological
Society, 10(4) (December): 177-191.
81
Trypeta varipennis, new species (p. 180); not illustrated
Arthropoda: Insecta: Diptera: Tephritidae (fruit flies)
“Habitat.—Bright Angel Hotel, brink of Grand Canyon, Coconino Co., Arizona.
A female
specimen collected July 10 by Mr. H. S. Barber. Type [holotype] No. 6635, U.S.N.M.”
________________________________________________________________________________
41
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Coulter, John M.
1896
Preliminary revision of the North American species of Echinocactus, Cereus, and Opun tia.
Contributions from the U.S. National Herbarium, 3(7): 355-462, I-IV [index].
82
Echinocactus polycephalus xeranthemoides,
NEW VARIETY
(p. 358).
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
“Type, Siler of 1881 and 1883 [= syntypes] in Herb. Mo. Bot. Gard.” [Herbarium of the Missouri
Botanical Garden]. “Specimens examined: Utah (Siler of 1883, ‘Kanab Mts.’ [syntype]): Arizona
(Siler of 1881, near the Colorado ‘on the Kanab wash’ [syntype]; Rusby 619, of 1883, at
Peach Springs [paratype]; Evans of 1891, between Gila Bend and Yuma) [paratype].”
________________________________________________________________________________
Coulter, John M., and Rose, J. N.
1900
Monograph of the North American Umbelliferae. Contributions from the U.S. National
Herbarium, 7(1): 9-256, i-vii.
83
Phellopterus multinervatus,
NEW SPECIES
(pp. 169-170); not illustrated
Trachyophyta: Magnoliopsida: Brassicales: Apiaceae (parsleys)
“Type locality, Peach Springs, northern Arizona; collected by Mr. and Mrs. J. G. Lemmon, May,
1884 (distributed as Cymopterus montanus); type specimen [holotype] in U.S. Nat. Herb.”
(Paratypes indicated from San Francisco Mountains and Fort Huachuca, Arizona.)
[Although there is no further data on the locality, the Lemmons visited Peach Springs Canyon,
descending to Diamond Creek and the Colorado River and remained in the vicinity of Peach Springs
for a month. See J. G. Lemmon (1888, Grand Cañon of the Colorado. Overland Monthly, New
Series, 12(69) (September): 244-256). Thus the precise type locality for this species must be
considered to be generalized. Also note that the gathering is likely from what is now the Hualapai
Indian Reservation.]
________________________________________________________________________________
Coville, Frederick Vernon
1894
New or little-known plants. Ostrya Knowltoni, a new species of hop hornbeam. Garden
and Forest, 7(317) (March 21): 114-116.
84
Ostrya knowltoni, NEW SPECIES, fig. 23.
(Subsequent grammatical correction of specific epithet to Ostrya knowltonii.)
Trachyophyta: Magnoliopsida: Fagales: Betulaceae (alders)
“Type specimen in the United States National Herbarium, collected July 10th, 1892 (No. 272) in
Yavapai County [today Coconino Co.], Arizona, within the Grand Cañon of the Colorado River, by
J. W. Toumey.” Holotype illustrated in fig. 23 (p. 115). Type locality at top of Old Hance Trail,
Hance Canyon (see below regarding its recovery). [The original holotype illustration and the
recovered type locality are illustrated on the covers of the present checklist.]
42
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
“In the year 1889 Mr. Frank H. Knowlton was occupied in making a collection of the plants of San
Francisco Mountain, in northern Arizona, and spent a portion of his time in a side trip to the Grand
Cañon at a point known as Cañon Spring, having traversed the road from San Francisco Mountain
by the way of Hull Spring and Red Horse Spring. [. . .] In the vicinity of Cañon Spring Mr. Knowlton
obtained a specimen of a tree which subsequently proved to be an Ostrya, but which was not at
that time in fruit. In 1892 Professor J. W. Toumey, of Tucson, Arizona, botanist of the State
University, visited the same locality and obtained fruiting specimens of this tree, which in January,
1894, came into the writer’s hands.”
“The following communication from Professor Toumey, under date of February 5th, 1894, will
service to direct others to the extact station of the original tree, which is, with little doubt, the
same specimen from which Mr. Knowlton obtained his material:
“The Ostrya, which you inquire further in regard to, and specimens of wich were sent to the
Department of Agriculture a year ago, as found north of Flagstaff, on the rim of the Grand Cañon
of the Colorado River. It is growing at the left, a few rods after begining the descent to the
river over John Hance’s trail.”
Later researchers failed to locate the type locality, and the species was there presumed to have
been extirpated.
However, in 1997 Earle Spamer thought that the oversight was due to the
misunderstanding of “John Hance’s trail,” portions of which were destroyed by landslides in 1895.
Hance thereafter rerouted the upper part of his trail to descend into Red Canyon farther to the
east, which is the “Hance Trail” as known today. The older trail at the head of Hance Canyon, if
referred to at all, is called the “Old Hance Trail,” although it is hardly even a route today. Spamer
asked Nancy Brian (Grand Canyon National Park) to investigate whether the type locality could be
identified at the head of the Old Hance Trail. As reported by Nancy J. Brian and Earle E. Spamer
(2000, Knowlton hop-hornbeam revisited (Ostrya knowltonii Cov.), Bartonia, no. 60: 49-56), p.
54:
“On 23 May 1997, Brian located the head of the Old Hance Trail and descended into the canyon.
She saw the distinctive dark yellow-green color of a deciduous tree nestled amongst the darker
foliage of the surrounding Rocky Mountain Douglas-firs and white firs (Figs. 2A, B [of that paper]).
Such a ‘window box’ habitat is frequently found just below the north-facing slope of the canyon
wall. It is a shaded, moist, steep environment which allows forest-dwelling species to grow well
below their usual elevational range. Scrambling down the steep slope, at ca. 6,800 feet (2,100 m)
elevation, Brian located about 25 trees of Ostrya knowltonii. Other plants of the community include
scattered shrubs such as snowberry and Utah serviceberry, along with an occasional pinyon pine,
ponderosa pine, mutton grass, and phlox. No redbuds were in the immediate vicinity, as Tourney
had described, although Gambel oak was found upslope on the canyon rim. Most of the Knowlton
hop-hornbeams were shrub-like, with up to ten branches originating from ground level. A few trees
were saplings, indicating a measure of reproductive success. Collections were made for the National
Herbarium (US), the University of Arizona (ARIZ), the Desert Botanical Garden (DBG) in Phoenix,
and the Academy of Natural Sciences of Philadelphia (PH; Fig. 3 [of Brian and Spamer]).”
________________________________________________________________________________
43
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Cresson, Ezra T., Jr.
1919
Dipterological notes and descriptions. Proceedings of the Academy of Natural Sciences of
Philadelphia, 71: 171-194.
Bombylius facialis, NEW SPECIES (pp. 187-188); not illustrated
85
Arthropoda: Insecta: Diptera: Bombyliidae (bee flies)
“Type.—♂; Rim of Grand Canyon, Arizona, 7000 feet, alt., [sic] May 23, 1918, (F. M. Jones),
[A. N. S. P. No. 6213].” [Square brackets around catalogue number data are part of quotation.]
________________________________________________________________________________
Crosswhite, Frank S.
1967
Revision of Penstemon Section Habroanthus (Scrophulariaceae). III. Series Virgati.
American Midland Naturalist, 77(1) (January): 28-41.
86
Penstemon virgatus pseudoptutus,
NEW SUBSPECIES
(pp. 35-37) [the call-out for fig. 1 refers
only to a map of distributions].
Trachyophyta: Magnoliopsida: Lamiales: Plantaginaceae (plantains)
“Typus [holotype]: Mead 503 (US) from DeMotte Park, Kaibab Plateau, Coconino County,
Arizona.”
Other “specimens examined” [paratypes] include: “Bright Angel Point, N rim of Grand Canyon,
Pilsbry s.n. (PH). 19 mi N of Bright Angel Point at V.T. Park, Richards s.n. (NY). V.T. Ranger
Station, Kaibab Plateau, Collum s.n. (US) . . . Kaibab Plateau, Kraus s.n. (WIS). Kaibab Natl.
Forest, Goodman & Hitchcock 1630 (F, NY, PH), Johnson s.n. (WIS). Dry Park, Kaibab N.F.,
Eggleston 10227 (US). Hwy. from Jacob Lake to Lodge, Kaibab Forest, Collom 944 (US).
Roadside near Grand Canyon N.P., Kaibab Forest, Kearney & Peebles 13744 (NY). W of Jct.
of Cape Royal & Point Imperial Rds., Kaibab Plateau, Collom s.n. (US) [i.e. in Grand Canyon
National Park]. Marble Flat, N Rim of Grand Canyon, Collom 1049 (US).”
________________________________________________________________________________
44
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Davis, William T.
1921
Records of cicadas from North America with descriptions of new species. Journal of the
New York Entomological Society, 29(1) (March): 1-16.
Tibicen apache, NEW SPECIES (pp. 3-5, Pl. 1, figs. 4-6)
87
Arthropoda: Insecta: Hemiptera: Cicadidae (cicadas)
(Holotype and allotype from Florence, Arizona; illustrated.)
“The following typical material
6
[paratypes ; not illustrated] of apache from Arizona is in the writer’s collection: . . . Grand
Canyon, Lower Bright Angel Trail, August 2, 1917, three males (Dr. Knight) . . . .”
________________________________________________________________________________
DeLong, Dwight M.
1964
A monographic study of the North American species of the genus Ballana (Homoptera:
Cicadellidae). Ohio Journal of Science, 64(5) (September): 305-370.
88
Ballana basala, NEW SPECIES (p. 356, figs. 74 [p. 319], 193 [p. 327], 391 [p. 339])
Arthropoda: Insecta: Hemiptera: Cicadellidae (leafhoppers)
(Holotype and allotype from Walsenburg, Colorado.) Paratypes include “. . . 5 males, 6 females,
Grand Canyon, Arizona, July 28, 1936, R. H. Beamer . . .” Holotype, allotype, and paratypes
(not specified) in DeLong collection; other paratypes (not specified) in U.S. National Museum,
University of Kansas collection, and Ohio State University collection.
________________________________________________________________________________
6
Whereas it seems that Davis’ use of the word “typical” may actually mean “usual”, and not being an
implication of paratypical material, I have always preferred in preparing catalogs and checklists
such as this one to err subjectively on the side of types when they appear +in older literature prior
to the established codes of nomenclature, because occasionally it can be demonstrated through
ancillary evidence, if not simply inferred, that such material is in fact of type status. The checklist
entries can always be dismissed. —E.E.S.
45
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Dempster, Lauramay T., and Ehrendorfer, Friedrich
1965
Evolution of the Galium multiflorum complex in western North America. II. Critical
taxonomic revision. Brittonia, 17 (October): 289-334.
Galium munzii Hilend & Howell subsp. ambivalens, NEW SUBSPECIES (pp. 309-310); not
89
illustrated
Trachyophyta: Magnoliopsida: Gentianales: Rubiaceae (madders)
“Type from ‘about Grand Canyon of the Colorado,’ MacDougal 157 (UC 138660. Isotypes,
ARIZ, F, GH, NY, US)”; distribution notes “Grand Canyon of the Colorado River, and eastward on
the Little Colorado River.”
________________________________________________________________________________
Doering, Kathleen C.
1939
A contribution to the taxonomy of the subfamily Issinae in America North of Mexico
(Fulgoridae, Homoptera). University of Kansas Science Bulletin, 26(2) (October 1): 83167 (including Pls. 13-22).
Bruchomorpha bunni, NEW SPECIES pp. 119-121, Pl. 20, fig. 3, Pl. 21, fig. 15, Pl. 22, figs.
90
4, 4a
Arthropoda: Insecta: Hemiptera: Issidae (planthoppers)
“Holotype male, taken at Grand Canyon, Arizona, on August 11, 1927, by R. H. Beamer; allotype
female, Grand Canyon, August 11, 1927, by P. A. Readio. Two female paratypes and nine male
paratypes, same data.” (Other paratypes from Arizona and New Mexico.) “The types are in Snow
entomological Collection at the University of Kansas.”
________________________________________________________________________________
Durham, Floyd E.
1952
A new pocket gopher from the North Rim of the Grand Canyon, Arizona. Journal of
Mammalogy, 33(4) (November): 498-499.
91
Thomomys bottae boreorarius, NEW SUBSPECIES; not illustrated
Chordata: Mammalia: Rodentia: Geomyidae (pocket gophers)
“Type.—Adult female, skin and skull, number 1184, Hancock Foundation Collections, University of
Southern California, taken at Swamp Point, 7522 ft. alt., 18½ mi. northwestward of Bright Angel
Point, North Rim of the Grand Canyon, Coconino County, Arizona; July 10, 1947; Floyd E. Durham;
original number, 2141.”
________________________________________________________________________________
46
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Eastwood, Alice
1902
New western plants. Bulletin of the Torrey Botanical Club, 29(8) (August): 523-525.
Corydalis wetherillii, NEW SPECIES (p. 524); not illustrated
92
Trachyophyta: Magnoliopsida: Ranunculales: Papaveraceae (poppies)
Listed under the section, “New Species from Colorado and Utah [sic].” “This peculiar species was
collected near Bright Angel Creek, one of the branches of the Colorado River which comes into
the river nearly opposite Hans’ Trail [Hance Trail]. This trail is that taken by tourists who go into
the cañon from the Flagstaff side. It is named in honor of Mr. Alfred Wetherill who collected it in
the summer of 1897.” “The type [holotype] is in the herbarium of the California Academy of
Sciences.”
_______________________________________________________________
1903
New species of western plants. Bulletin of the Torrey Botanical Club, 30(9) (September):
483-502.
Fraxinus macropetala, NEW SPECIES (pp. 494-495); not illustrated
93
Trachyophyta: Magnoliopsida: Lamiales: Oleaceae (olives)
“This was collected by E. O. Wooton, July 9, 1902, in the Grand Cañon of the Colorado, being
no. 1102 of his collection; this specimen is in fruit. The flowers were collected at the same place
[i.e., Grand Canyon], on the Bright Angel Trail, by Dr. C. Hart Merriam, the middle of May,
1903.
These specimens [syntypes] are both in the Herbarium of the California Academy of
Sciences.”
_______________________________________________________________
1937
94
New species of western plants. Leaflets of Western Botany, 2(1) (January): 7-9.
Oreocarya capitata, NEW SPECIES (p. 9); not illustrated
Trachyophyta: Magnoliopsida: Boraginales: Boraginaceae (borage)
“Type [holotype]: No. 232041, Herb. Calif. Acad. Sci., collected by the author, No. 5969, on the
Hermit Trail, Grand Canyon of the Colorado, Arizona, April 9, 1917. This specimen is in flower.
A fruiting specimen, Herb Calif. Acad. Sci. No. 232040 [paratype], was collected by the author on
the same trail, June 18, 1916. It was collected June 23, 1933, on the north side of the Grand
Canyon on the Kaibab Trail to Roaring Springs, Eastwood & Howell No. 1005, Herb. Calif. Acad.
Sci. No. 232039 [paratype].”
________________________________________________________________________________
47
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Ehrhorn, Edw. M.
1899
Five new Coccidæ. The Canadian Entomologist (London), 31(1) (January): 5-7.
Arthropoda: Insecta: Hemiptera: Coccidae (soft scales)
Ripersia arizonensis, NEW SPECIES (p. 5); not illustrated
95
“Hab.—In ants’ nest on the roots of grass and Artemisia, sp., Camp Thurber [Hance Ranch],
Grand Canyon of the Colorado, Arizona.”
Dactylopius formicarii, NEW SPECIES (pp. 6-7); not illustrated
96
“Hab.—In ants’ nests on the roots of Artemisia, sp. Thurber’s Camp [Hance Ranch], Grand
Canyon of the Colorado, Arizona.”
________________________________________________________________________________
Ellis, Don E.
1946
Anthracnose of dwarf mistletoe caused by a new species of Septogloeum. Journal of the
Elisha Mitchell Scientific Society, (June): 25-50, Pls. 4-6.
Septogloeum gillii, NEW SPECIES (pp. 30-38, text-fig. 2)
97
Ascomycota (sac fungi) incertae sedis
“Parasititc on shoots of Aceuthobium campylopodum and A. douglasii in Western United States.”
“The following specimens in the herbarium of the Division of Forest Pathology at Albuquerque,
New Mexico, were examined: (Instances in which the disease was found on herbarium collections
of the host are indicated by an asterick [sic] (*)).”
“On A. campylopodum f. divaricatum (Engelmann) Gill . . . Grand Canyon, Gill, June 6, 1932
(68234)* . . . .”
(Other host species examined from specimens gathered within the bounds of the present checklist
did not exhibit the disease.)
________________________________________________________________________________
Ellis, J. B., and Everhart, B. M.
1900
New species of fungi from various localities with notes on some published species.
Bulletin of the Torrey Botanical Club, 27(2) (February): 49-64.
98
Puccinia circinans, NEW SPECIES (p. 61); not illustrated
Basidiomycota (club fungi): Pucciniomycetes: Pucciniales: Pucciniaceae
“On Penstemon spectabilis, Grand Cañon of the Colorado (Prof. J. W. Toumey).”
________________________________________________________________________________
48
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Esslinger, Theodore L.
2000
A key for the lichen genus Physconia in California, with descriptions for three new species
occurring within the state. Bulletin of the California Lichen Society, 7(1) (Summer): 1-6.
Physconia isidiomuscigena, new species (pp. 5-6, figs. 5, 6)
99
Ascomycota (sac fungi): Lecanoromycetes: Teloschistales: Physciaceae
“Type [holotype]: U.S.A. Arizona. Coconino Co.: Grand Canyon Natl. Park, Grandview Trail,
36°00′N, 111°59′W, 1980 m, Nash 30843 (ASU, holotype; TLE, isotype).” Figs. 5, 6 depict
“part of holotype specimen”. [The geographic coordinates pinpoint a location on the upper part
of Grandview Trail.]
“Selected additional specimens examined ([designated] Paratypes): . . . Grand Canyon Natl.
Park, N rim, junction of paved roads, ca. 6.5 km N of Kaibab Lodge, 36° 16′ N, 112° 03′
W [sic, Grand Canyon Lodge, coordinates pinpoint a location just east of Arizona Route 67 north
of the Cape Royal Road jct], 2470 m, Nash 9943 (ASU, MIN, TLE); Grand Canyon Natl. Park,
South Kaibab Trail, 36° 03′ 45″ N, 112° 03′ 30″ W, 1950 m, Nash 30819 (ASU, MIN). . . .”
[In noting the paper’s title, Esslinger comments, “This species is presently known from only two
collections in California and is apparently much more common in the southern Rocky Mountains.”]
________________________________________________________________________________
Fall, H. C.
1905
Revision of the Ptinidæ of boreal America. Transactions of the American Entomological
Society, 31(2/3): 97-296, Pl. 7.
Arthropoda: Insecta: Coleoptera: Ptinidae (spider beetles)
100 Petalium bistriatum Say var. arizonense, NEW VARIETY (p. 217); not illustrated.
[P.
bistriatum Say described, pp. 216-217.]
“A small series [syntypes] collected by Hubbard or Schwarz at Oracle and Bright Angel, Arizona
. . . .”
101 Catorama grande, NEW SPECIES (p. 236); not illustrated
Species based on specimens [syntypes] seen from Texas and Arizona, including “Bright Angel”.
102 Catorama longulum, NEW SPECIES (p. 244); not illustrated
“Described from four examples [syntypes] taken by Barber and Schwarz at Williams and Bright
Angel. With these is included a nearly identical specimen taken by the same collectors Las Vegas,
New Mexico.”
103 Catorama pingue, NEW SPECIES (p. 250); not illustrated
Described on syntypes from “Arizona (Santa Rita Mountains, Oracle, Bright Angel, Pinal
Mountains, Williams, Ash Fork)”.
_______________________________________________________________
49
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1909
Revision of the species of Diplotaxis of the United States. Transactions of the American
Entomological Society, 35(1) (January): 1-97.
104 Diplotaxis conformis, NEW SPECIES (p. 34); not illustrated
Arthropoda: Insecta: Coleoptera: Scarabaeidae (scarab beetles)
Described on syntypes “from Glenwood Springs, Colorado (Fenyes), and from Prescott and Bright
Angel, Arizona. Specimens in Mr. Dury’s collection are labeled ‘Northern Arizona.’ ”
_______________________________________________________________
1910
Miscellaneous notes and descriptions of North American Coleoptera. Transactions of the
American Entomological Society, 36(2): 89-197.
105 Bruchus perplexus, NEW SPECIES (p. 177); not illustrated
Arthropoda: Insecta: Coleoptera: Chrysomelidae (leaf beetles)
“The type [holotype] is from Alburquerque [sic], New Mexico (Wickham). With it are associated
specimens [paratypes] from Highrolls, New Mexico, Bright Angel, Arizona, Palm Springs,
California, and Claremont, California.”
________________________________________________________________________________
Fernald, M. L.
1934
A new Primula from the Grand Canyon of the Colorado. Rhodora (Journal of the New
England Botanical Club), 36 (April): 117-118, Pl. 282.
106 Primula (Section Farinosae) hunnewellii, NEW SPECIES; Pl. 282, figs. 1, 2
Trachyophyta: Magnoliopsida: Ericales: Primulaceae (primroses)
“Arizona: limestone cliffs, North Rim, Grand Canyon, Coconino Co., August 19, 1928, Francis
Welles Hunnewell, no. 10,883 (TYPE [holotype] in herb. F. W. Hunnewell, duplicate [isotype] in
Gray Herb.).” (Fig. 3 of the plate illustrates a comparative species, P. specuicola Rydb. from its
type locality, Bluff City, Utah.)
________________________________________________________________________________
Ferris, Roxana Stinchfield
1918
Taxonomy and distribution of Adenostegia. Bulletin of the Torrey Botanical Club, 45(10)
(October): 299-423.
107 Adenostegia parviflora, NEW SPECIES (p. 409, Pl. 10, fig. 8; Pl. 11, fig. 4 [plate legends p.
422])
Trachyophyta: Magnoliopsida: Lamiales: Orobanchaceae (broomrapes)
“TYPE LOCALITY: Grand Canyon of the Colorado River, near the San Francisco Mountains,
Arizona, Knowlton 270. TYPE [holotype], No. 48859 of the U. S. National Herbarium.”
________________________________________________________________________________
50
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Fischer, Pierre C.
1979
Coryphantha vivipara var. kaibabensis, a new variety from northern Arizona. Cactus and
Succulent Journal (Cactus and Succulent Society of America), 51(6): 286-287.
108
Coryphantha vivipara var. kaibabensis,
NEW VARIETY .
[Illustrations depict living specimens.]
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
“HOLOTYPE P.C. Fischer 4094, collected on 31/III/69 northwest of Jacob Lake, Coconino Co.,
Arizona. 1890 meters (6200 ft).
Growing with Juniper, Pinyon Pine, Sagebrush (Artemisia
tridentata) and Cliff Rose (Cowania sp.). Growing on dolomite and limestone. Deposited at the
herbarium of the University of California (UC), Berkeley, CA.”
_______________________________________________________________
1991
Echinocereus triglochidiatus variety toroweapensis, a new variety from the Grand
Canyon. Cactus and Succulent Journal (Cactus and Succulent Society of America), 63(4)
(July/August): 194-195.
109 Echinocereus triglochidiatus var. toroweapensis, NEW VARIETY; [live specimens
illustrated]
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
“Holotype. P. C. Fischer 7196 collected April 7, 1990, Toroweap Point, Grand Canyon National
Park, Mohave Co., Arizona; 1,340 meters (4,400 ft). [. . .] Deposited at the herbarium of the
University of Arizona (Ariz.), Tucson.”
________________________________________________________________________________
Foissner, Ilse, and Foissner, Wilhelm
1993
Revision of the family Spironemidae Doflein (Protista, Hemimastigophora), with
description of two new species, Spironema terricola n. sp. and Stereonema geiseri n. g.,
n. sp. Journal of Eukaryotic Microbiology, 40(4): 422-438.
110 Spironema terricola, NEW SPECIES (pp. 424-429, figs. 5-34, 59, 62, 65)
Hemimastigophora: Hemimastigea: Hemimastigida: Spironemidae (hemimastigid protist)
“Type location. Soil from the Grand Canyon, USA (upper entrance to Bright Angel Trail),
36° N, 112° E [sic]. Type specimens. One slide of holotype specimens and three slides of paratype
specimens (all protargol-impregnated) have been deposited in the collection of microscope slides
of the Oberösterreichische Landesmuseum in Linz, Austria. Accession numbers: 44, 45, 46, 47
/1992.” [Not only are the coordinates in error (“E” instead of “W”) but they are overly generalized,
that geographical intersection being 8¾ miles southeast of the head of Bright Angel Trail.]
________________________________________________________________________________
51
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Fox, William J.
1893
New species of fossorial Hymenoptera. The Canadian Entomologist (London), 25(5)
(May): 113-117.
111 Gorytes dentatus, NEW SPECIES (pp. 116-117); not illustrated
Arthropoda: Insecta: Hymenoptera: Crabronidae (crabronid wasps)
Holotype male from “Grand Canyon, Arizona, ‘70 miles North of Flagstaff.’
(C. H. Tyler
Townsend).”
[Townsend visited the canyon in July 1892 at Hance Ranch and trail.
For his general (non-
scientific) account of the trip see Townsend, 1893, A wagon-trip to the Grand Cañon of the
Colorado River, Appalachia, 7(1) (February): 48-63.]
[Note: Townsend (1896, Notes on New Mexico and Arizona Hymenoptera, The Canadian
Entomologist (London), 28(5) (May): 138-142), p. 139 there lists “Gorytes dentatus Fox., n. sp.
—Grand Canyon, Arizona; Hance trail, July 11. One specimen. [. . .] Det., Fox.” It is of course
not a new species therein but the record further substantiates the identification of the locality as
the Old Hance Trail.]
________________________________________________________________________________
Garth, John S.
1949
Studies in Arizona Lepidoptera. I. A new subspecies of Speyeria atlantis (Edwards) from
the Kaibab Plateau, Grand Canyon National Park. Bulletin of the Southern California
Academy of Sciences, 48 (Part 1): 1-4.
112 Speyeria atlantis schellbachi, NEW SUBSPECIES; illustrated
Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals)
“Type material: Male holotype, AHF No. 471, and female allotype, AHF No. 471a, from Neal
Spring, North Rim, Grand Canyon National Park, Coconino County, Arizona, 8,175 feet, July 5,
1947, collected by John S. Garth, Allan Hancock Foundation survey party. Twenty paratypes as
follows: 1 female, North Rim, Grand Canyon, July 29, 1939, Louis Schellbach, collector; 1
female, North Rim, Grand Canyon, August 19, 1942, H. C. Bryant, collector; 1 male, 1 female,
Two River Junction, Grand Canyon, July 28, 1945, Louis Schellbach, collector, the preceding
four paratypes on loan from the Naturalist Workshop, Grand Canyon National Park; 1 male, 1
female, Neal Spring, North Rim, Grand Canyon, August 16, 1946, J. S. Garth, collector; 2
females, Kanabownits Spring, North Rim, Grand Canyon, August 22, 1946, J. S. Garth,
collector; 7 males, 2 females, Neal Spring, North Rim, Grand Canyon, July 5 to 18, 1947, J. S.
Garth, collector; 1 male, Robbers’ Roost Spring, North Rim, Grand Canyon, July 10, 1947, J.
S. Garth, collector; 2 females, Swamp Lake and Swamp Ridge, North Rim, Grand Canyon,
July 12 and 14, 1947, J. S. Garth, collector. The holotype, allotype, and ten paratypes are in
the collection of the Allan Hancock Foundation, the University of Southern California. The
remaining paratypes will be distributed as follows: one male and three females to the Naturalist
52
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Workshop, Grand Canyon National Park, one pair each to the United States National Museum,
the American Museum of Natural History, and the Los Angeles County Museum.”
[The type locality, Neal Spring, is by the junction of Cape Royal Road and Point Imperial Road.]
________________________________________________________________________________
Gertsch, Willis J.
1933
New genera and species of North American spiders. American Museum Novitates, (636),
28 pp.
113 Misumenops coloradensis, NEW SPECIES (p. 17, figs. 15 [p. 12], 46 [p. 24])
Arthropoda: Euchelicerata: Areneae: Thomisidae (crab spiders)
(“Male holotype, female allotype, and paratypes of both sexes from Colorado.”) “Female paratypes
from Grand Canyon (North Rim), Arizona, and Kanab, Utah.”
_______________________________________________________________
1934a
Notes on American Lycosidae. American Museum Novitates, (693), 25 pp.
114 Arctosa mokiensis, NEW SPECIES (p. 8); not illustrated [but see note below]
Arthropoda: Euchelicerata: Areneae: Lycosidae (wolf spiders)
Holotype male. “Type Locality.—Male holotype from Indian Gardens [Havasupai Gardens],
Grand Canyon, Arizona, May 26, 1905.”
In synonymy is “Arctosa notuabanda Gertsch, 1933, American Museum Novitates, No. 637, p. 8,
fig. 11 (not noctuabunda Montgomery)”, which was noted therein only by the name Arctosa
noctuabunda Montgomery in the legend for fig. 11 and is not mentioned in that text. [It is not
clear whether this represents a figure of the type of A. mokiensis.]
_______________________________________________________________
1934b
115
Further notes on American spiders. American Museum Novitates, (726), 26 pp.
Phidippus kaibabensis,
NEW SPECIES
(pp. 13-14, fig. 19 [p. 17])
Arthropoda: Euchelicerata: Areneae: Salticidae (jumping spiders)
“Male holotype from the Kaibab forest, near the north rim of the Grand Canyon, Arizona, July 8,
1931 (Gertsch).”
_______________________________________________________________
53
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1960
The fulva group of the spider genus Steatoda (Araneae, Theridiidae). American Museum
Novitates, (1982), 48 pp.
116
Steatoda variata,
(pp. 24-29, figs. 22-25 [p. 13], 34-44 [pp. 17, 26])
NEW SPECIES
Arthropoda: Euchelicerata: Areneae: Theridiidae (cobweb weavers)
“Type Locality: Male holotype from 25 miles north of The Gap, Navajo Indian Reservation,
Arizona, July 21, 1949 (W. J. and J. W. Gertsch). (Non-type records include “Twenty-five males
north of The Gap . . . male, many females.”)
________________________________________________________________________________
Gertsch, Willis J., and Ennik, Franklin
1983
The spider genus Loxosceles in North America, Central America, and the West Indies
(Araneae, Loxoscelidae). Bulletin of the American Museum of Natural History, 175
(Article 3): 264-360.
117 Loxosceles kaiba, NEW SPECIES (p. 303, figs. 110-113 [p. 299], 133 [p. 300])
Arthropoda: Euchelicerata: Areneae: Sicariidae (recluse spiders)
“Type Data: Male holotype, female and four immatures [paratypes] from Thunder Cave [Thunder
River Cave], 3900 feet, near Monument Point, north rim of Grand Canyon National Park, Coconino
County, Arizona, 15 September 1977 (S. Peck, D. Carlile), deposited in AMNH.
Other Record
[non-type?]: Grand Canyon National Park, Cameron Cave, 5000 ft., 7 Dec. 1954 (R. de Saussure),
♂.”
The specific epithet, kaiba [sic], is named “for Kaibab Plateau on north face of Grand Canyon”
(p. 303).
________________________________________________________________________________
Goldman, Edward A.
1924
Two new kangaroo rats from Arizona. Journal of the Washington Academy of Sciences
(Washington, D.C.), 14(15) (September 19): 372-373.
Chordata: Mammalia: Rodentia: Heteromyidae (kangaroo rats)
118
Dipodomys microps celsus,
NEW SUBSPECIES
(p. 372); not illustrated. (Virgin Valley
Kangaroo Rat)
“Type [holotype] from 6 miles north of Wolf Hole, Arizona (altitude 3,500 feet). No. 243101,
♂adult, U. S. National Museum (Biological Survey Collection) collected by E. A. Goldman, October
16, 1922. Original number 23411.”
119
Dipodomys ordii cupidineus,
NEW SUBSPECIES
(pp. 372-373); not illustrated. (Kaibab
Kangaroo Rat)
Type [holotype] from Kanab Wash, at southern boundary of Kaibab Indian Reservation,
Arizona. No. 243093,
♂ adult, U. S. National Museum (Biological
by E. A. Goldman, October 12, 1922. Original number 23384.”
54
Survey Collection), collected
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
_______________________________________________________________
1929
A new antelope squirrel from Arizona. Journal of the Washington Academy of Sciences
(Washington, D.C.), 19: 435-436.
120 Ammospermophilus leucurus tersus, NEW SUBSPECIES; not illustrated.
(Grand Canyon
Antelope Squirrel)
Chordata: Mammalia: Rodentia: Sciuridae (squirrels)
“Type [holotype].—From lower end of Prospect Valley, Grand Canyon, Hualpai Indian
Reservation [Hualapai Indian Reservation], Arizona (altitude 4,500 feet). No. 202645,
♂
young adult, U. S. National Museum (Biological Survey collection), collected by E. A. Goldman,
October 3, 1913.
Original number, 22269.”
“Specimens examined.—Ten, all from the type
locality.” [Examined specimens not itemized.]
_______________________________________________________________
1932
A new beaver from Arizona. Journal of Mammalogy, 13(3) (August): 266-267.
121 Castor canadensis repentinus, NEW SUBSPECIES; not illustrated.
(Colorado River Beaver)
Chordata: Mammalia: Rodentia: Castoridae (beavers)
“Type [holotype] From Bright Angel Creek, Grand Canyon of the Colorado River, Arizona
(altitude 4,000 feet); no. 161613,
♀ adult, U.S. National Museum (Biological Survey collection);
collected by Clarence Birdseye, September 14, 1909; original number 72.”
Two specimens
examined from type locality (no specimen or collecting data published for the second specimen).
“At the type locality the beavers inhabit Bright Angel Creek, which in short reaches descends the
terraced north side, below the out rim, of the Grand Canyon. None are known from the Colorado
River in that vicinity, and a measure of isolation would seem to be due to the rapids and heavy
current through rock-bound gorges extending for many miles, and affording very few places
suitable for beavers to establish homes.”
_______________________________________________________________
1937
A new canyon mouse of the genus Peromyscus from Arizona. Journal of Mammalogy,
18(1) (February): 92-93.
122 Peromyscus crinitus peridoneus, NEW SUBSPECIES; not illustrated.
(Grand Canyon Mouse)
Chordata: Mammalia: Rodentia: Cricetidae (deer mice)
“Type [holotype].—From Bright Angel Trail, 4800 feet altitude, south side of Grand Canyon,
Coconino County, Arizona. No. 202,424,
♂ adult, skin and skull, U. S. National Museum (Biol.
Surv. coll.); collected by E. A. Goldman, August 19, 1913. Original number 22,145.” [Altitude
on Bright Angel Trail indicates locality is at about the 2nd rest house descending.]
“Specimens examined [non-type].—Total number, 36, as follows: Arizona: Fredonia, 1; Grand
Canyon (near Bass Camp, 3000-5200 feet), 3; near Grand Canyon Spring, 4000-7500 feet, 9;
Betatakin Canyon, 3; Bright Angel Creek, 3 miles above mouth, 1; Bright Angel Trail, south side,
4800-6500 feet, 2; Shinumo Creek, 3000 feet, 12; Nankoweap Valley, 1.
near Fort Cameron, 3; Parowan, 1.”
55
Utah: Beaver River,
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
_______________________________________________________________
1938
New pocket gophers of the genus Thomomys from Arizona and Utah. Journal of the
Washington Academy of Sciences (Washington, D.C.), 28(7) (July 15): 333-343.
Chordata: Mammalia: Rodentia: Geomyidae (pocket gophers)
123
Thomomys fossor kaibabensis,
NEW SUBSPECIES
(pp. 333-334); not illustrated. (Common
name given by the author, “Kaibab Pocket Gopher”)
“Type [holotype].—From DeMotte Park, Kaibab Plateau, Coconino County, Arizona (altitude
9,000 feet).
No. 262891,
♂
adult, skin and skull, U. S. National Museum (Biological Survey
collection), collected by Luther C. Goldman, September 10, 1937. Original number 443.”
“Specimens examined [non-type].—Total number, 23, all from Kaibab Plateau, Arizona, as
follows: Bright Angel Spring, 3; DeMotte Park (type locality), 18; Greenland Spring, 1; Jacob Lake,
1.”
124
Thomomys bottae nicholi,
NEW SUBSPECIES
(pp. 337-338); not illustrated. (Common name
given by the author, “Shivwits Plateau Pocket Gopher”)
“Type [holotype].—From 20 miles south of Wolf Hole (road to Parashouts), Shivwits Plateau,
Mohave County, Arizona (altitude 5,000 feet). No. 262864,
♂ adult, skin and skull, U. S. National
Museum (Biological Survey Collection), collected by Luther C. Goldman, August 6, 1937. Original
number 363.”
“Specimens examined [non-type].—Total number, 19, as follows: Arizona: Diamond Butte (3 miles
northwest), 1; Wolf Hole, 6 (20 miles south, 4; 6 miles north, 2). Utah: St. George, 12.”
________________________________________________________________________________
Gray, Asa
1878
Contributions to the botany of North America. Proceedings of the American Academy of
Arts and Sciences, 13: 361-374.
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
125
Astragalus subcinereus,
NEW SPECIES
(p. 366); not illustrated
“Mokiak Pass in the northwestern part of Arizona, near the Utah boundary, Dr. E. Palmer, 1877.”
126
Astragalus scaposus,
NEW SPECIES
(p. 366); not illustrated
“Among rocks in dry creek-bottoms, in Mokiak Pass, near the northeastern [sic] corner of
Arizona, Dr. E. Palmer, 1877.” [Gray correctly cites the location of Mokiak Pass elsewhere.]
127
Astragalus mokiacensis,
NEW SPECIES
(p. 367); not illustrated
“Rocky ravines, Mokiak Pass, on the borders of Utah and N. W. Arizona, Dr. Palmer, 1877.”
128
Astragalus artipes,
NEW SPECIES
(p. 370); not illustrated
“Mokiak Pass, northwest corner of Arizona, in ravines, Dr. E. Palmer, 1877.”
56
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
129
Actinella biennis,
NEW SPECIES
(p. 373); not illustrated
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
“S. Utah and Arizona, Mokiak Pass south of St. George, Palmer (no. 260 of coll. 1877)” [syntype;
another from Richfield, Utah].
_______________________________________________________________
1886
Contributions to American botany. Proceedings of the American Academy of Arts and
Sciences, 21: 363-413.
130
Mirabilis bigelovii,
NEW SPECIES
(p. 413); not illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Nyctaginaceae (four o’clocks)
“Arizona, and perhaps in California on the Colorado. Common in the Grand Cañon of the
Colorado, below Peach Spring [Peach Springs], collected May 5, 1885, A. G.” [indicating
syntypes].
[Gray descended into Grand Canyon in Peach Springs Canyon to Diamond Creek and the Colorado
River. Today the Grand Canyon localities “below Peach Spring” are in the Hualapai Indian
Reservation.]
________________________________________________________________________________
Gray, Asa
in Professors Gray, Torrey, Thurber, and Dr Engelmann
[in Gray, Asa; Torrey, John; Thurber, George; and Engelmann, George]
1861
Botany. (“By Professors Gray, Torrey, Thurber, and Dr Engelmann”.) In: Ives, Joseph
C., Report upon the Colorado River of the West, explored in 1857 and 1858 by Lieutenant
Joseph C. Ives, Corps of Topographical Engineers, under the direction of the Office of
Explorations and Surveys, A. A. Humphreys, Captain Topographical Engineers, in charge.
By order of the Secretary of War. U.S. 30th Congress, 1st Session, House and Senate
Executive Document 90. Washington, D.C.: U.S. Government Printing Office [1861], Part
4, 30 pp. [Botany section, with 1860 date printed on title leaf, apparently arranged or
edited by John Strong Newberry; see p. 20, footnote.]
The collecting localities noted here are now within the Hualapai Indian Reservation.
131
Phacelia (Coreanthus) ivesiana,
NEW SPECIES
(p. 21); not illustrated
Trachyophyta: Magnoliopsida: Boraginales: Hydrophyllaceae (waterleafs)
One of three localities noted is “Diamond river; April 3” (entire note) [indicating syntypes]. Not
illustrated.
132
“Gilia dactylophyllum, (n. sp.?)” (p. 22); not illustrated
Trachyophyta: Magnoliopsida: Ericales: Polemoniaceae (phlox)
[It is unclear whether this entry refers to G. dactylophyllum Torr. (which itself is now in another
genus), or whether Gray had questionably introduced the name or left his specimens open to
57
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
receive a new name in the future (see his remarks below); and so retained here with question as
well.]
“Mouth of Diamond river; April 3. The specimens [syntypes] are scarcely sufficient for
description. A slender plant, about 3 inches high. Corolla twice as long as the calyz, white; ovules
about 7 in each cell.” (entire entry).
________________________________________________________________________________
Greene, Edward Lee
1890
Bibliographical notes on well known plants.—X. Bulletin of the Torrey Botanical Club,
17(1) (January 15): 13-14.
133 Rhus canadensis Marsh. var. simplicifolia, NEW VARIETY, p. 13 (footnote); not
Trachyophyta: Magnoliopsida: Sapindales: Anacardiaceae
illustrated
(cashews)
“In the deep cañons of Northern Arizona which lead down to the Grand Cañon of the
Colorado”.
[Identity of these canyons uncertain and not resolved here based on ecological
potential, which could well include the side canyons in which the more famous trails descend into
the canyon—Hance Trail, Grandview Trail, Bright Angel Trail.]
_______________________________________________________________
1900
Studies in the Cruciferae.—III. Pittonia, 4 [1899-1901]: 187-207.
Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards)
134
Arabis eremophila,
NEW SPECIES
(pp. 194-195); not illustrated
“Common about Peach Springs, northern Arizona . . . .” [No further details. Syntypic material
from this locality and from Aztec, New Mexico.]
135
Arabis recondita,
NEW SPECIES
(p. 195); not illustrated
“Diamond Creek Cañon, a tributary of the Colorado, in northern Arizona, N. C. Wilson, April
1893.” [Syntypic material from this locality and from Glenwood Springs, Colorado.]
[It is unclear whether “Diamond Creek Cañon” refers to Diamond Creek or more generally to Peach
Springs Canyon, to which Diamond Creek is a tributary close to its confluence with the Colorado
River, the former being the long principal route into Grand Canyon from Peach Spring s.]
_______________________________________________________________
1905
Segregates of the genus Rhus. Leaflets of Botanical Observation and Criticism, 1: 114144. [Date of publication for pp. 129-144, November 29, 1905 (see p. 129).]
Trachyophyta: Magnoliopsida: Sapindales: Anacardiaceae
(cashews)
136 Schmaltzia cissodes, NEW SPECIES (p. 136); not illustrated
“Grand Cañon of the Colorado, Ariz., near Indian Garden [Havasupai Gardens], Bright Angel
Trail, C. H. Merriam, 10 May, 1903.”
58
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
137 Schmaltzia hirtella, NEW SPECIES (p. 137); not illustrated
“Grand Cañon of the Colorado, Ariz., 10 July, 1892, E. O. Wooton.”
_______________________________________________________________
1906
The genus Ptelea in the western and southwestern United States and Mexico.
Contributions from the U.S. National Herbarium, 10(2) (July 16): 149-179.
Trachyophyta: Magnoliopsida: Sapindales: Rutaceae (rues)
138
Ptelea palida,
NEW SPECIES
(p. 70); not illustrated
“Arid rocky hills above Peach Springs, northern Arizona, collected by the writer July 3, 1889;
type in the National Herbarium.”
139
Ptelea straminea,
NEW SPECIES
(p. 70); not illustrated
“Rocky slopes of the Virgin Mountains in extreme northwestern Arizona, C. A. Purpus, 1898; his
no 6165 as in the national Herbarium.”
140 Ptelea nitida, NEW SPECIES (p. 71); not illustrated
“Species known to me only as in the Herbarium of the California Academy from somewhere in the
Grand Canyon of the Colorado in Arizona, collected by E. O. Wooton, July 8, 1892.”
141 Ptelea argentea, NEW SPECIES (p. 71); not illustrated
“This species . . . is known to me only as collected by Dr. F. H. Knowlton, somewhere in the Grand
Canyon of the Colorado, September 10, 1889, the specimens filling two sheets in the national
Herbarium (one the type).”
142 Ptelea triptera, NEW SPECIES (pp. 71-72); not illustrated
“Grand Canyon of the Colorado, Arizona, excellent specimen in the National Herbarium,
collected by J. W. Toumey, July 11, 1892.”
[Very probably gathered near Hance Ranch; compare date and remarks with Ostrya knowltonii
Coville, 1894.]
143 Ptelea lutescens, NEW SPECIES (p. 72); not illustrated
“Species of northwestern Arizona, inhabiting canyons tributary to the Grand Canyon of the
Colorado, the best specimen from Red Canyon Trail [Hance Trail], collected June 10, 1901, by
Lester F. Ward (type [holotype] in the National Herbarium). Younger material, with fruit not
mature, from Bright Angel Trail, by Dr. C. Hart Merriam, May 10, 1903. These last imperfect
specimens have smaller relatively broader leaflets, and may possibly represent another species
[paratypes, uncertainly].”
144 Ptelea elegans, NEW SPECIES (p. 72); not illustrated
“Grand Canyon of the Colorado, Arizona, June 14, 1891, D. T. McDougal [sic], the shrub said
to grow ‘in rich soil and in water.’
The only specimens seen are in the National Herbarium
[syntypes].”
59
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
[Daniel Trembly MacDougal visited Hance Ranch in June 1891 and signed the Hance guest book.
He likely also descended into the canyon on the Old Hance Trail, down which probably is the
type locality for this species, in as much as there is no standing water on the rim of the canyon
except at a reservoir maintained by John Hance for the use of his hotel and riding stock. (The
upper portion of this trail was destroyed by landslides in 1895, after which Hance rerouted his
trail into Red Canyon, which today is the “Hance Trail”, which should not be confused with the
original trail, now barely not even a route and referred to as the Old Hance Trail.) See record of
MacDougal’s guestbook signature in G[eorge]. K. Woods, 1899, Personal impressions of the Grand
Cañon of the Colorado River near Flagstaff, Arizona, as seen through nearly two thousand eyes,
and written in the private visitors’ book of the world-famous guide Capt. John Hance, guide, storyteller, and path-finder San Francisco: Whitaker and Ray Co., for G. K. Woods, Flagstaff, Arizona
Territory. MacDougal added no comments to his signature.]
145 Ptelea confinis, NEW SPECIES (p. 72); not illustrated
“. . . inhabiting the region of the Grand Canyon of the Colorado in northwestern Arizona and
adjacent Utah. The material examined is on four sheets in the National Herbarium, among which
I would name that on sheet 15254 as the type [holotype].”
146
Ptelea saligna,
NEW SPECIES
(p. 73); not illustrated
“Nagle’s Ranch, Arizona, altitude 2,240 meters, M. E. Jones, September 15, 1894, no. 6048,
as in the National Herbarium (type) and the Herbarium of the California Academy. [. . .] The
locality is in northern Arizona, north of the Grand Canyon.”
_______________________________________________________________
1909
New plants from Arizona. Leaflets of Botanical Observation and Criticism, 2: 20-24.
[Date of publication for pp. 1-24, 6 February 1909 (see p. 1).]
147
Senecio stygius,
NEW SPECIES
(p. 21); not illustrated
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
“Grand Cañon of the Colorado, J. G. Lemmon, May, 1884. In U. S. Herb., the sheet [holotype]
not numbered.”
[Lemmon, with his wife, spent a month in the Peach Springs area in May 1884. For a general
(non-scientific) account of their visit, see J. G. Lemmon, 1888, Grand Cañon of the Colorado,
Overland Monthly, new series, 12(69) (September): 244-256.]
_______________________________________________________________
1911
Some southwestern mulberries. Leaflets of Botanical Observation and Criticism, 2: 112121. [Date of publication for pp. 105-120, 6 October 1910 (see p. 105); date for pp.
121-152, 11 May 1911 (see p. 121).]
148
Morus grisea,
NEW SPECIES
(p. 121); not illustrated
Trachyophyta: Magnoliopsida: Rosales: Moraceae (mulberries)
“This fine specimen of this is on U. S. Herb. sheet 41506, and purports to have been collected
forty years ago, by Dr. Edw. Palmer, at ‘Hell Cañon, Arizona’; I presume in the northwestern
60
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
part of the State, in some arm or tributary of the Grand Cañon.” [Date of publication for pp. 105 120, 6 October 1910 (see p. 105); date for pp. 121-152, 11 May 1911 (see p. 121).]
[“Hell Canyon” could have been an informal name for Palmer’s locality, and it is not clear why
Greene would have supposed it to be “in some arm or tributary of the Grand Cañon” other than
through some familiarity with Palmer’s collecting activities. Consulting Will C. Barnes’ Arizona
Place Names, revised and enlarged by Byrd H. Granger (University of Arizona Press, 1960), p.
346, the only “Hell Canyon” recognized therein is in the present Yavapai County, which notes,
“this canyon was part of the route from the old Beale Trail into Chino Valley in the early days”
(i.e. between Ash Fork and Prescott). Greene’s record is retained herein in the event that Palmer’s
“Hell Canyon” is in fact elsewhere in northwestern Arizona.]
________________________________________________________________________________
Grijseels, Sietske;
Nielsen, Jens Christian; Randelovic, Milica; Nielsen, Jens; Nielsen, Kristian Fog;
Workman, Mhairi; and Frisvad, Jens Christian
2016
Penicillium arizonense, a new, genome sequenced fungal species, reveals a high chemical
diversity in secreted metabolites. Scientific Reports (Springer Nature), 6(35112),
doi:10.1038/srep35112, 13 pp.
149 Penicillium arizonense Frisvad, Grijseels &
J.C. Nielsen,
NEW SPECIES
Ascomycota (sac fungi): Eurotiomycetes: Eurotiales: Trichocomaceae (saprobe fungi)
“A new soil-borne species belonging to the Penicillium section Canescentia is described, Penicillium
arizonense sp. nov. (type strain CBS 141311 T=IBT 12289T).”
“Type: Herb. C-F-101845, cultures ex type IBT 12289=CBS 141311, from a sample of dry red
soil, south rim of Grand Canyon, Grand Canyon Village, Arizona, USA (36° 3′ 22.31″ N; 112°
7′ 30.73″ W), Per V. Nielsen [Per Væggemose Nielsen], July 1990”; additional cultures “from
same source as the culture ex type, but not of the same clone.”
[Map coordinates pinpoint a locality just to the west of the footpath between Park Headquarters
and the Rim Trail, at a point half-way between those two places
(https://www.findlatitudeandlongitude.com/, accessed 28 October 2022).]
________________________________________________________________________________
Grissell, E. E.
1983
Boharticus, n. gen., with a review of Rhopalicus Foerster and Dinotiscus Ghesquiere
(Hymenoptera: Pteromalidae). Pan-Pacific Entomologist, 59(1/4): 78-102.
150
Boharticus margaretae,
NEW SPECIES
(pp. 86-87, figs. 2 [p. 81], 13 [p. 85], 17 [p. 85])
Arthropoda: Insecta: Hymenoptera: Pteromalidae (pteromalid wasps)
(Holotype female from Garden of the Gods, El Paso Co., Colorado; USNM 100593.) “Paratypes:
8
♀, same data a holotype, 1 ♀, ARIZONA, Mohave Co., 8 miles SW Peach Springs, 4500′, 7
♀ in British
September 1964, C. W. O’Brien, ex Juniperus. All specimens in USNM, except 1
Museum (Natural History).” [The locality is in the vicinity of Truxton, Arizona.]
61
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
The type species of Boharticus Grissell, new genus, is designated as B. richardi Grissell, new
species (which is not pertinent to this checklist).
________________________________________________________________________________
Hall, E. Raymond, and Davis, William B.
1934
Notes on Arizona rodents. Proceedings of the Biological Society of Washington
(Washington, D.C.), 47(5): 51-56.
151 Thomomys bottae trumbullensis, NEW SUBSPECIES (pp. 51-53); not illustrated
Chordata: Mammalia: Rodentia: Geomyidae (pocket gophers)
“Type [holotype].—Male, adult, skin-and-skull; no. 58588, Mus. Vert. Zool.; three miles south of
Nixon Spring, Mt. Trumbull, Mohave Co., Arizona; May 26, 1933; collected by Seth B. Benson,
original no. 2005.” [The locality, “three miles south of Nixon Spring”, is just to the south of Mount
Trumbull itself, today within the Grand Canyon–Parashant National Monument, about 2¾ miles
north of the boundary of Grand Canyon National Park.]
“Specimens examined [non-type]. — Total number, 31, as follows: From Mohave County, Arizona:
Nixon Spring, 6250 ft., Mt. Trumbull, 8; three miles south of Nixon Spring (volcanic sand), Mt.
Trumbull (type locality), 3; four miles south of Nixon Spring (volcanic sand), Mt. Trumbull, 13;
head of Toroweap Valley, 2; six miles north of Wolf Hole, 4900 ft., 3; south side Virgin River, St.
George, Washington County, Utah, 2.”
________________________________________________________________________________
Hall, Jack C., and Evenhuis, Neal L.
2003
Review of the subgenus Geron (Geron) Meigen in the Nearctic region (Diptera:
Bombyliidae: Toxophorinae). Zootaxa, (181): 1-72.
152 Geron prosopidis, NEW SPECIES
Arthropoda: Insecta: Diptera: Bombyliidae (bee flies)
Paratypes include 11 taken at “S Rim Grand Canyon, 21-22.vii.1932 (R.H. Painter)”.
________________________________________________________________________________
Harvey, Mark S., and Wynne, J. Judson
2014
Troglomorphic pseudoscorpions (Arachnida: Pseudoscorpiones) of northern Arizona, with
the description of two new short-range endemic species. Journal of Arachnology, 42(3):
205-219.
Arthropoda: Euchelicerata: Pseudoscorpiones (pseudoscorpions): Chernetidae
153
Hesperochernes bradybaughi,
NEW SPECIES
(pp. 211-214, figs. 15-26)
“Types. U.S.A.: Arizona: Mohave County: holotype male, PARA-1001 Cave, Grand CanyonParashant National Monument, ca. UTM 0264500 N, 4060700 E, Zone 12S, baited pitfall
trap 3B, 20 August 2007, J.J. Wynne (MNA); 1 female, same data as holotype except baited pitfall
62
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
trap 5A (MNA); 1 female, same data as holotype except opportunistic, mid cave, 13 August 2005
(NMA [sic]).”
154
Tuberochernes cohni,
NEW SPECIES
(pp. 214-217)
“Type: U.S.A.: Arizona: Mohave County: holotype male, PARA-1001 Cave, Grand CanyonParashant National Monument, ca. UTM 0264500 N, 4060700 E, Zone 12S, the deeper
extent of the twilight zone (near the dark zone), opportunistic collecting, 13 August 2005, J.J.
Wynne (MNA).”
________________________________________________________________________________
Hayward, C. Lynn
1932
The paper wasps of Utah; including a description of a new variety of Polistes canadensis
Linn. Proceedings of the Utah Academy of Sciences, 9: 85–101 (pagination includes Pl.
9).
155 Polistes canadensis var. kaibabensis, NEW VARIETY (pp. 89-90, Pl. IX, figs. 4, 12, 14, 16
[p. 101, legend p. 100])
Arthropoda: Insecta: Hymenoptera: Vespidae (hornets)
“HOLOTYPE: Female; Number 28, collection of Brigham Young University, Provo, Utah. Collected
at Point Sublime, north rim of Grand Canyon of the Colorado River, Arizona, July 19, 1927 by
Vasco M. Tanner. ALLOTYPE: Male; Number 29, collection of Brigham Young University, Provo,
Utah. Collected at North Rim, Grand Canyon of the Colorado, Arizona, July 9, 1929 by Lowell
Woodbury. PARATYPES: 22 females; all in collection of Brigham Young University, Provo, Utah;
collected at Point Sublime, Rim of Grand Canyon of the Colorado River, Arizona, July 19, 1927
(Vasco M. Tanner, Clarence Cottom, Irvin Rasmussen, Anson Call).
TYPE LOCALITY: Point
Sublime, Rim of Grand Canyon of Colorado River, Arizona.”
The author notes: “The variety kaibabensis is at present known only from the immediate vicinity
of the type locality. I have had the privilege of examinng a considerable series from the collection
of the University of Utah, Salt Lake City, which were taken in the type locality and could well be
designated paratypes.
I have also seen a very typical specimen from the San Diego Natural
History Museum which was collected at El Tovar, South Rim of the Grand Canyon. This specimen
is now in that museum.”
[In consideration of the fact that the new variety is recorded only from the vicinity of Point
Sublime, the reference to the allotype’s locality as “at North Rim, Grand Canyon of the Colorado”
seems to imply that it is from the place, North Rim, as opposed to generically the canyon’s North
Rim. It seems that the “considerable series” of specimens in the University of Utah should be
excluded as paratypes; and the El Tovar specimen is a referred specimen only.]
________________________________________________________________________________
63
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Henderson, Junius
1914
A new Sonorella from the Grand Canyon, Arizona. The Nautilus (Philadelphia), 27(11)
(March): 122-124.
156 Sonorella betheli, NEW SPECIES; not illustrated
Mollusca: Gastropoda: Stylommatophora: Helminthoglyptidae (shoulderbands)
NOTE: The locality is in error, not of the Grand Canyon, but this species is included in this
nomenclator to account for the published record of a new taxon named upon material ostensibly
collected at Grand Canyon. The type specimens, though also subsequently reidentified
taxonomically, still are the name-bearing types of the taxon Sonorella betheli.
Refer to the
information below.
“Type [holotype] (in Univ. Colo. Museum) . . . . Cotype [paratype] (in Acad. Nat. Sci. Phila.) . . . .
Type locality, Bright Angel Trail, Grand Canyon, Arizona.”
Henderson: “Among some shells recently received from Mr. Ellsworth Bethel, of East Denver High
School, were four dead specimens of Sonorella collected by him on Bright Angel Trail, at Grand
Canyon, Arizona, in 1913. He was collecting fungi, and unfortunately did not note the exact locality
of the snail find, but writes that he followed the trail closely, and thinks he got the shells ‘about
one hundred yards west of the upper limit of the trail and not more than twenty feet below the
top,’ though he cannot be certain and ‘may have gotten them as far down as the half way house.’
He supposed them to be common and made no note of the place. They are much larger than the
common S. coloradoensis [refer to Helix (Arionta) coloradoensis Stearns, 1890, No. 280 in the
present checklist] of that region, and differ in other respects. They did not seem to fit the
description of any other species, but the finding of so large a species along a trail which has been
searched by some of our ablest conchologists and most thorough collectors made me doubt that
it could be new, so I sent two specimens to Dr. Pilsbry, who pronounced them undescribed. ”
Under the description for Helminthoglypta traski (Newcomb), Henry A. Pilsbry (1939, Land
Mollusca of North America (North of Mexico), Academy of Natural Sciences of Philadelphia,
Monograph 3, Volume 1, Part 1, p. 174), made the following comments on this species: “Misled
by a mistaken locality given by the collector, Junius Henderson described rather small specimens
. . . as Sonorella betheli. In 1913 after visiting the Grand Canyon, where he thought that this
shell was picked up, the late Prof. Ellsworth Bethel spent some time in Calfornia. His chief interest
was botanical. Probably these shells were taken near Los Angeles; the type in University of
Colorado Museum and a cotype, 109733 A.N.S.P., both of which I have seen, agree fully with
traski from that vicinity.”
________________________________________________________________________________
64
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Hershler, Robert; Liu, Hsiu-Ping; and Stevens, Lawrence E.
2016
A new springsnail (Hydrobiidae: Pyrgulopsis) from the lower Colorado River basin,
northwestern Arizona. Western North American Naturalist, 76(1): 72-81.
157
Pyrgulopsis hualapaiensis, NEW
SPECIES ;
illustrated (including a color photo of the type
locality, fig. 5)
Mollusca: Gastropoda: Neotaenioglossa: Hydrobiidae (spring snails)
“Types.—Holotype, USNM 1248611 (a dry shell), Upper Peach Springs, outflow just below
concrete weir, Hualapai Indian Reservation, Mohave County, Arizona, 35.55786°N,
113.4314°W, 1280 m elevation, coll. RH and LS, 16 May 2014.
Paratypes (from same lot),
USNM 1266144 (approximately 250 dry shells and alcohol-preserved specimens).”
“Referred material [non-type].—Arizona. Mohave County: topotypes, USNM 1247929 (dry shells),
coll. LS, 28 May 2010; USNM 1248612 (dry shells and alcohol-preserved specimens), coll. LS, 9
January 2014; USNM 1259458 (dry shells and alcohol-preserved specimens), coll. LS, 24 July
2014.”
________________________________________________________________________________
Hester, J. Pinckney
1943
Two new Opuntias from the Grand Canyon region in Arizona. Cactus and Succulent
Journal (Cactus and Succulent Society of America), 15: 191-194.
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
158
Opuntia hualpaensis,
NEW SPECIES
(pp. 191, 193, fig. 94, upper right [p. 192])
“Type [holotype]: Deposited in the Dudley Herbarium, Stanford University, No. 285575, collected
along U. S.-Arizona Highway No. 66, 11 or 12 miles east of Peach Springs and 2 or 3
miles west of Hyde Park, Arizona, by J. Pinckney Hester, July, 1942.” [“Hyde Park”, near mile
marker 117 on Historic Route 66, no longer exists. The gathering locality is near the present-day
Grand Canyon Caverns tourist attraction.]
159
Opuntia abyssi,
NEW SPECIES
(pp. 193-194, fig. 94, upper left and lower [p. 192])
“Type [holotype]: Deposited in the Dudley Herbarium, Stanford University, No. 285624, collected
in Peach Springs Canyon . . . by J. Pinckney Hester, in 1939.”
________________________________________________________________________________
65
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Higgins, Larry C.
1974
Cryptantha atwoodii (Boraginaceae); a new species from Arizona. Southwestern
Naturalist, 19(2) (July 26): 127-130.
160
Cryptantha atwoodii,
NEW SPECIES ;
illustrated
Trachyophyta: Magnoliopsida: Boraginales: Boraginaceae (borage)
“TYPE. Arizona: Coconino County, 7 mi N of junction of hwy 89 and 164 on hwy 89, 20
May 1970.
N. D. Atwood 2624.
Holotype deposited at BRY.
Isotype at WTS.”
“Apparently
confined to the type locality near [T]he Gap . . . .”
________________________________________________________________________________
Hochstätter, Fritz
1995
The genera Pediocactus—Navajoa—Toumeya : revised : in the shadow of the Rocky
Mountains. (Chris Holland, translator.) Mannheim, Germany: Fritz Hochstätter, 168 pp.
[It is not determined here whether this title was originally published as this English translation,
or if it was translated from another item thus far not identified.]
161
Sclerocactus whipplei (Englemann & Bigelow) Britton & Rose ssp. busekii,
NEW SUBSPECIES
(pp. 132-134)
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
House Rock Valley, Arizona.
________________________________________________________________________________
Hodgson, Wendy C.
2001
Taxonomic novelties in American Agave (Agavaceae). Novon, 11(4) (Winter): 410-416.
162 Agave phillipsiana, NEW SPECIES (pp. 410-413, fig. 1 [“Drawn from Hodgson 8357B and
Brian s.n.”])
Trachyophyta: Magnoliopsida: Asparagales: Asparagaceae (agaves)
“TYPE: U.S.A. Arizona: Coconino County, Grand Canyon National Park, Clear Creek Canyon,
ca [0].7 mi. upstream from camping area, 1140 m, 36°07.381′N, 112°00.568′W, 13 Sep. 1999,
W. C. Hodgson 11861 (holotype, DES 44332; isotypes, GCNP, MO photo).” “Paratypes. U.S.A.
Arizona: Coconino County, Grand Canyon National Park, Deer Creek Canyon, 36°23′52.2″N,
112°30′32″W, 731 m, 11 June 1994, Hodgson et al. 8357B (DES, GCNP, MO photo), 14 Aug.
1995, Hodgson 8357A and Brian (DES, GCNP), 26 Sep. 1995, Brian s.n. (ASU, DES, GCNP, MO);
Grand Canyon National Park, Clear Creek Canyon, ca. 0.7-1 mi. upstream form camping area,
36°07.381′N, 112°00.568′W, 1140 m, 29 Apr. 1999, Hodgson et al. 11472 (DES, GCNP, MO
photo); same area only 100 m further upstream, 37°[sic]07.592′N, 112°00.324′W, 1145 m, 29
Apr. 1999, Hodgson et al. 11473 (DES, GCNP, MO photo).”
66
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Note: The discovery of this species was first noted, without name, in Hodgson (1996, Field work
and leg work; finding a new species, The Sonoran Quarterly (Desert Botanical Garden, Phoenix),
50(4) (December): 10-11).
________________________________________________________________________________
Holmgren, Arthur H., and Holmgren, Noel H.
1988
Euphorbia aaron-rossii (Euphorbiaceae), a new species from Marble and Grand Canyons
of the Colorado River, Arizona. Brittonia, 40(4) (October/December): 357-362.
163 Euphorbia aaronrossii, NEW SPECIES; illustrated
Trachyophyta: Magnoliopsida: Malpighiales: Euphorbiaceae (spurge)
“Type: U.S.A. ARIZONA. Coconino Co.: Marble Canyon, along the Colorado River, 12 river
miles below Lees Ferry, Salt Water Wash, among boulders, 930 m (3050 ft), 5 May 1971, A.
Holmgren, N. Holmgren, P. Holmgren, D. Holmgren & A. Ross 15558 (HOLOTYPE: NY; ISOTYPES:
ARIZ, ASU, BRY, F, TEX, UTC).”
________________________________________________________________________________
Holmgren, Noel H.
1973
Five new species of Castilleja (Scrophulariaceae) from the Intermountain Region. Bulletin
of the Torrey Botanical Club, 100(2): 83-93.
164
Castilleja kaibabensis,
NEW SPECIES
(pp. 89-91, fig. 4 [p. 88])
Trachyophyta: Magnoliopsida: Lamiales: Scrophulariaceae (figworts)
“Type: Arizona, Coconino County, Kaibab Plateau, V T Meadows, 6.5 km south of Kaibab
Lodge, T.34 N., R.3 E., Sec. 7, elevation 2680 m, 7 August 1970, N. Holmgren & P. Holmgren
4682 (Holotype: NY; Isotypes: ARIZ, BRY, C, DAO, JEPS, KANU, NCU, RSA, TENN, US, UTC,
WTU).”
“Paratypes: All on the Kaibab Plateau, Coconino County, Arizona: Marble Flat, just north of
North Rim Ranger Station, elevation 2500 m, 17 July 1941, Collom s.n. (US, GCNP [herbarium
at Grand Canyon National Park Headquarters]); DeMotte Park, 7 August 1929, Mead 998
(GCNP); House Rock Canyon to Jacob Lake, elevation 2400 m, 14 August 1941, Kearney &
Peebles 13648 (US); V T Ranger Station, elevation 2800 m, 12 July 1940, Collom s.n. (GCNP);
V T Park, 30 km north of Bright Angel Point, 14 August 1938, Richards s.n. (NY); Ariz. Highway
67, 3.2 km north of Grand Canyon National Park boundary, T.34 N., R.3 E., Sec. 7,
elevation 2650 m, 12 August 1966, Holmgren & Reveal 2995 (ARIZ, BRY, C, DAO, JEPS, KANU,
NCU, NY, RSA, TENN, US, UTC, WTU); Lookout Canyon Road (422D), 5.6 km from
(northwest of) Grand Canyon Highway (Ariz. 67), T.35 N., R.2 E., Sec. 15, elevation 2700
m, 6 August 1970, N. Holmgren & P. Holmgren 4668 (ARIZ, BRY, C, DAO, JEPS, KANU, NCU, NY,
RSA, TENN, US, UTC, WTU).”
________________________________________________________________________________
67
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Holmgren, Noel H., and Holmgren, Patricia K.
2002
New mentzelias (Loasaceae) from the Intermountain Region of western United States.
Systematic Botany, 27(4) (October/December): 747-762.
165
Mentzelia memorabilis, new species (pp. 753-756; fig. 4)
Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa)
“TYPE: U.S.A. Arizona. Mohave Co., Uinkaret Plateau, just N of the Navajo Trail Road, 1.1
km W of the Clayhole Wash crossing, 27.5 km air distance SSW of Colorado City,
36°49′09″N, 113°11′21″W, T40N R8W S31, 1480 m, in partially barren gypsum-clay soil, 17
Sep 2001, N. H. Holmgren & P K. Holmgren 14580 (holotype: NY; isotypes: ASC, ASU, BRY, COLO,
RM, RSA, UTC).”
________________________________________________________________________________
Hopkins, Heidi
2014
A review of the genus Arenivaga (Rehn) (Blattodea, Corydiidae), with descriptions of new
species and key to the males of the genus. Sofia, Bulgaria; and Moscow, Russia: Pensoft
Publishers, 256 pp. (ZooKeys, 384: 1-256 (Special Issue).)
Arthropoda: Insecta: Blattodea: Polyphagidae (sand cockroaches)
166 Arenivaga grandiscanyonensis, NEW SPECIES (pp. 108-111, figs. 69, 70)
[Note: “rm” indicates Colorado River Mile, by convention measured downstream from Lees Ferry,
Arizona, with Right or Left bank indicated. Localities downstream from RM 164.7, on the left,
could actually be within the boundaries of Hualapai Indian Reservation.]
“Type locality. USA, Arizona, Mohave Co., Colorado River, Grand Canyon.”
“Material examined. Holotype:
♂ in NAUF labeled ‘Mohave Co. AZ, Colorado R. GC, rm211.5R,
4/13/02, Coll. R.J.Delph, Ex: Light, Old High Water, blue label with “3”,’ ‘HOLOTYPE Arenivaga
grandiscanyonensis Hopkins, 2012 [sic]’ [red label with black border].” [Square brackets are part
of the quotation.]
“Paratypes (2): USA: AZ, Coconino Co., Colorado River GC, 5/10/2001, J Rundall, rm160.5L,
ex.light old high water, blue label ‘3’, 1 specimen—NAU 106 (2, NAUF). All paratypes labeled
‘Paratype Arenivaga grandiscanyonensis Hopkins 2012 [sic]’ [blue label with black border].”
[Square brackets other than “sic” are part of the quotation.]
Also 169 specimens examined (non-type).
167 Arenivaga impensa, NEW SPECIES (pp. 138-141, figs. 93, 94)
(Holotype from Kingman, Arizona.)
“Paratypes (14): . . . AZ, Indian Garden [Havasupai
Gardens], Grand Canyon, 7/24/1934, 3800 ft., EL Bell & FE Lutz (2, AMNH); AZ, Coconino Co.,
Grand Canyon, 6/18/1954, M Cazier (1, AMNH) . . . . All paratypes labeled ‘Paratype Arenivaga
impensa Hopkins 2012 [sic]’ [blue label with black border].” [Square brackets other than “sic”
are part of the quotation.]
68
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
168 Arenivaga pagana, NEW SPECIES (pp. 177, 182, figs. 120 [p. 180], 121 [p. 181])
“Type locality. USA, Arizona, Mohave Co., Colorado River, Grand Canyon.”
“Material examined. Holotype:
♂ in NAUF labeled ‘Mohave Co. AZ, Colorado R. GC, blue label
with “3”, rm202.5L, 9/4/03, Coll. R.J.Delph, Ex: Light, New High Water’ ‘HOLOTYPE Arenivaga
pagana Hopkins, 2012 [sic]” [red label with black border].” [Square brackets other than “sic” are
part of the quotation.]
“Paratypes (9): USA: AZ, Mohave Co., Colorado River GC, 9/4/2003, RJ Delph, blue label with
“3”, rm 202.5L, ex.light, new high water (2, NAUF); AZ, Mohave Co., Colorado River GC,
9/9/2001, J Rundall, blue label with “3”, rm 198.0R, ex.light, old high water (1, NAUF); AZ,
Mohave Co., Colorado River GC, 9/9/2002, RJ Delph, blue label with “3”, rm 186.5L, ex.light, old
high water, NAU 107 (1, NAUF); . . . AZ, Coconino Co., Colorado River GC, 9/1/2002, RJ Delph,
blue label with “3”, rm37.3L, ex.light, old high water (1, NAUF); AZ, Coconino Co., Colorado River
GC, 10/18/1982, 855 m, LE Stevens, blue label with “3”, M:53R, Nankowap [sic], sand dunes at
night, Polyphagidae, Arenivaga det. D.Lightfoot (1, NAUF) . . . . All paratypes labeled ‘Paratype
Arenivaga pagana Hopkins 2012 [sic]’ [blue label with black border].” [Square brackets other
than “sic” are part of the quotation.]
________________________________________________________________________________
Horn, George H.
1893
The Galerucini of boreal America. Transactions of the American Entomological Society,
20(2) (April/June): 57-136.
169
Luperodes wickhami,
NEW SPECIES
(pp. 114-115); not illustrated
Arthropoda: Insecta: Coleoptera: Chrysomelidae (leaf beetles)
“Occurs in Arizona at Peach Springs (Wickham).”
________________________________________________________________________________
Hottes, F. C.
1956
Two new species of Cinara from northern Arizona with illustrations of hitherto unfigured
species and notes on Schizolachnus flocculosa (Williams) (Aphidae). Proceedings of the
Biological Society of Washington (Washington, D.C.), 69: 219-224.
Arthropoda: Insecta: Hemiptera: Aphididae (aphids)
170 Cinara grande, NEW SPECIES (pp. 219-220; text-figs. on p. 223)
“Host Abies concolor. Taken on the North Rim of the Grand Canyon, Arizona June 7, 1954.
Collected by J. W. Bongberg. Holotype apterous viviparous female returned to the collection of
The United States National Museum.”
69
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
171
Cinara poketa,
NEW SPECIES
“Holotype apterous
(pp. 220-221; text-figs. on p. 222)
viviparous female. Host, Pinus edulis Aug. 24, 1956. Taken at point where
road to Anita, Arizona branches from Highway 64 leading to Grand Canyon, Arizona.”
[Locality is on the Kaibab National Forest, Tusayan Ranger District, ca. 35.861° N, 112.134° W.]
________________________________________________________________________________
Johnson, Walter N.
1990
A new subspecies of Cicindela pusilla Say from northern Arizona. Cicindela, 22(1)
(March): 1-12.
172
Cicindela pusilla kaibabensis,
NEW SUBSPECIES
(description pp. 4-10, including right elytra of
various subspecies illustrated pp. 8-9 [C. p. kaibabensis, figs. 16-21])
Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles)
“TYPE MATERIAL: Holotype (male) collected 8 miles north of Kaibab Lodge, Coconino Co.,
Arizona, on August 2, 1977, along with 11 male and 13 female paratypes, collected b y author
(Walter N. Johnson). Allotype (female) collected 4 miles north of Kaibab Lodge, Coconino
Co., Arizona, along with 4 male and 2 female paratypes on August 2, 1977, collected by author.
Seven male and 2 female paratypes collected 8 miles north of Kaibab Lodge, Coconino Co.,
Arizona, on July 27, 1981, by author. Forty-eight male and 34 female paratypes collected 8
miles north of Kaibab Lodge, Coconino Co., Arizona, on July 22, 1987, by author. Thirty
male and 14 female paratypes collected 8 miles north of Kaibab Lodge, Coconino Co.,
Arizona, on July 22, 1987, by Todd Lawton.”
________________________________________________________________________________
Johnston, M. Andrew
2019
Phylogenetic revision of the psammophilic Trogloderus Leconte (Coleoptera:
Tenebrionidae), with biogeographic implications for the Intermountain Region. PeerJ, 7:
e8039, doi:10.7717/peerj.8039, 45 pp. + Supplemental Information online.
173
Trogloderus skillmani,
NEW SPECIES
(pp. 24-26, figs. 6D [p. 16], 7F [p. 17])
Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles)
“Type material. Holotype: ‘USA: AZ: Mohave Co./6m E Colorado City/Rosy Canyon Road/1.5
m S UT state line/12-VII-2016/F.W. & S.A. Skillman,’ ‘ARTSYS0007053,’ bearing red holotype
label. Deposited in the ASUHIC, catalog number ASUHIC0101565.
Paratypes: A total of 920
specimens from the western regions of the Colorado Plateau around the Coral Pink Sand Dunes,
Hurricane, and Toquerville Utah, bearing blue paratype labels (see Data S1 and S2 or SCAN for
full specimen data).” The 920 paratypes are listed in the Supplemental Information online.
________________________________________________________________________________
70
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Jones, Marcus E.
1895
Contributions to western botany. No. VII. Proceedings of the California Academy of
Sciences, Series 2, 5: 611-732.
174
Astragalus humistratus var. tenerrimus,
NEW VARIETY
(p. 649); not illustrated
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
[Syntypes]: “No. 6052f. September 17, 1894, Buckskin Mountains, Arizona, 9000° [sic] alt.,
in gravel, under conifers. No. 6064. September 20, 1894, road to Nagle’s Ranch, Buckskin
Mountains, Arizona, 9000° [sic] alt., in gravel. No. 6056gm. Same locality and date.”
175
Oenothera triloba var. ecristata,
NEW VARIETY
(pp. 681-682); not illustrated
Trachyophyta: Magnoliopsida: Myrtales: Onagraceae (evening primroses)
[Syntypes], all from Utah except for: “No. 6056ah.
September 18, 1894, De Motte Park,
Buckskin Mountains, Arizona, 9000° [sic] alt., in gravel.”
176
Bigelovia howardi var. attenuata,
NEW VARIETY
(pp. 691-692); not illustrated
Trachyophyta: Magnoliopsida: Lamiales: Oleaceae (olives)
[Syntypes], all from Utah except for: “No. 6052k. September 17, 1894, Buckskin Mountains,
Arizona, 9000° [sic] alt., in gravel.”
177
Laphamia congesta,
NEW SPECIES
(p. 703); not illustrated
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
[Holotype]: “No. 6063. September 21, 1894, in clefts of rocks on the mesa below the Buckskin
Mountains, Arizona, 7000° [sic] alt.”
178
Laphamia gracilis,
NEW SPECIES
(pp. 703-704)
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
[Holotype]: “No. 6050c. September 15, 1894, below Nagle’s Ranch, on edge of Buckskin
Mountains, in crevices of limestone rocks, at 7000° [sic] alt., in very dry places.”
_______________________________________________________________
1902
Contributions to western botany. No. 10. Robinson, Utah: Mammoth Record Print, 90
pp.; [17], 16 plates.
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
179
Astragalus kentrophyta var. coloradoensis,
NEW VARIETY
(p. 63, Pl. 1 [top row, 5th figure])
“Lee’s Ferry, Arizona, on the Colorado river, June 16, 1890.”
180
Astragalus kaibensis,
NEW SPECIES
(pp. 64-65, Pl. 2 [center row, 2nd figure])
“Gathered at House Rock, near Lee’s Ferry, northern Arizona, June 18, 1890, Jones.”
________________________________________________________________________________
71
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Kavanaugh, David H.
2008
A new species of Nebria Latreille (Coleoptera: Carabidae: Nebriini) from the Grand
Canyon, Arizona. Annals of the Carnegie Museum of Natural History, 77(1) (July): 1-5.
181 Nebria (Reductonebria) georgei, NEW SPECIES; illustrated
Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles)
“Types.—HOLOTYPE, a male, deposited in MSBC, labeled: ‘USA AZ Coconino Co Grand Canyon
Natl. Park, N36.102, W112.163 RM92.3L 06-07 May 2001 coll. Cobb, Brantley, Lightfoot’/
‘shore zone pitfall trap’/ ‘COL CAR 001 006’/ ‘Nebria n. sp. ? det G.E. Ball 2004’/ ‘UNM-29’/
‘HOLOTYPE Nebria georgei Kavanaugh n. sp. des. by D.H. Kavanaugh 2007 [sic]’ [red label]. One
paratype, a female, deposited in CAS, labeled: “USA AZ Coconino Co Grand Canyon Natl. Park,
N36.187, W113.107 RM180.8R 01-02 Sept 2003 coll. Cobb, Brantley, Lightfoot’/ ‘shore zone
pitfall trap’/ ‘COL CAR 001 006’/ ‘UNM-30’/ ‘PARATYPE Nebria georgei Kavanaugh n. sp. des. by
D. H. Kavanaugh 2007 [sic]’ [yellow label.” [Square brackets except for “sic” are part of the
quotation.]
“Type Locality.—Colorado River, Grand Canyon National Park, Coconino County,
Arizona.”
________________________________________________________________________________
Kim, Ke Chung
1966
The species of Enderleinellus (Anoplura, Hoplopleuridae) parisitic on the Sciurini and
Tamiasciurini. Journal of Parasitology, 52(5) (October): 988-1024.
182
Enderleinellus kaibabensis,
NEW SPECIES
(pp. 1000-1002, 1022; figs. 12 [p. 999, legend p.
998], 29 [p. 1001], 54-59 [p. 1005, legend p. 1004])
Arthropoda: Insecta: Psocodea: Enderleinellidae (sucking lice)
“Type data: Holotype male, allotype female, and 6 paratypes (3 males and 3 females), ex Sciurus
kaibabensis Merriam, Kaibab National Forest, Arizona, USA, Ferris Coll. 906 (USNM 168301),
G.F.F. All type specimens except for 2 paratypes deposited in UCB; 1 male and 1 female paratype
(on 1 slide) are deposited in UM.” [Inasmuch as the host is the endemic Kaibab squirrel, the type
locality must be on the North Kaibab Ranger District.]
________________________________________________________________________________
Klauber, Laurence M.
1930
New and renamed subspecies of Crotalus confluentus Say, with remarks on related
species. Transactions of the San Diego Society of Natural History, 6(3) (February 28):
95-144, Pls. 9-12.
183 Crotalus confluentus abyssus, NEW SUBSPECIES (pp. 114-117, Pl. 11, fig. 1 [legend p.
142]). (Common name given by the author, “Grand Canyon Rattlesnake”)
Chordata: Reptilia: Squamata: Viparidae (rattlesnakes)
“Type [holotype].—No. 2216 in collection of L.M.K. Captured alive on the Tanner Trail 300 ft.
below the south rim of the Grand Canyon, Coconino County, Arizona; altitude approximately 7000
72
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
ft. Collected Sept. 15, 1929, by E. D. McKee.” The holotype is illustrated in Pl. 11, fig. 1.
Subspecies also studied on two other living specimens and ten other specimens from the South
Rim and the inner canyon [= paratypes]: “GCNP R37, Tonto Platform near Bright Angel Trail
Alt. 4000 ft.; GCNP R38 [Coconino] Plateau near Navajo Point, Alt. 7000 ft.; GCNP R49
Phantom Ranch, north of Bright Angel Creek [sic] [seen as a skin]; GCNP R50 Desert View
Point [specimen in poor condition]; GCNP R51 Desert View Point [specimen in poor condition];
LMK 2093 Roaring Springs Power Plant; LMK 2215 Roaring Springs Power Plant [specimen
seen alive]; LMK 2272 Grand View Trail, 2000 ft. below South Rim [specimen seen alive];
USNM 32725 Grand Canyon [seen as a skin]; USNM 59747 Shimuno Creek [sic, Shinumo
Creek] [seen as a skin]; USNM 78477 Bright Angel Canyon [seen as a skin]; USNM 78478
Burro Spring [North Rim; seen as a skin].
[Edwin D. McKee wrote of his 1929 sighting and capture of the holotype on Tanner Trail. (McKee,
1976, Discovery of the Grand Canyon Rattlesnake. Journal of Arizona History, 17(1) (Spring): 4749.)]
________________________________________________________________________________
Knight, Harry H.
1921
Monograph of the North American species of Deraeocoris (Heteroptera, Miridae).
Minnesota State Entomologist, 18th Report to the Governor, pp. 77-210, Pl. 8.
[Published June 18, 1921.] [Also published June 19, 1921 as University of Minnesota,
Agricultural Experiment Station, Technical Bulletin 1, retaining original pagination, 77210.]
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
184 Deraeocoris fulvus, NEW SPECIES (pp. 144-145; fig. 18)
(Holotype and allotype from Williams, Arizona.) Paratypes include: “2 ♀ Aug 3 [1917], Grand
View, Grand Canyon”.
185 Deraeocoris bullatus, NEW SPECIES (pp. 147-148; fig. 20)
“Holotype:
♂
August 3 [1917], Grand View, Grand Canyon, Arizona (H. H. Knight); Cornell
University collection. Allotype: taken with the type.”
186 Deraeocoris navajo, NEW SPECIES (pp. 155-156; fig. 24)
“Holotype:
♂
August 3 [1917], Grand View, Grand Canyon, Arizona (H. H. Knight); Cornell
University collection.”
_______________________________________________________________
73
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1925
Descriptions of thirty new species and two new genera of North American Miridae
(Hemiptera). Bulletin of the Brooklyn Entomological Society, 20(1) (February): 33-58.
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
187 Ceratocapsus clavicornis, NEW SPECIES (pp. 47-48)
“Holotype:
♂
August 3, 1917, Grand View, Grand Canyon, Arizona (H. H. Knight); author’s
collection. Allotype: same data as the type. Paratypes: 30
♂♀, taken with the types on Cowania
mexicana.”
188 Phytocoris mellarius, NEW SPECIES (pp. 56-57)
“Holotype:
♂
August 3, 1917, Grand View, Grand Canyon, Arizona (H. H. Knight); author’s
collection. Allotype: same data as the type. Paratypes: 3♂ 6♀, taken with the types. This
species was probably collected on some conifer although I do not find the remarks in my notes.”
_______________________________________________________________
1926
Descriptions of seven new species of Pilophorus (Hemiptera, Miridae). Bulletin of the
Brooklyn Entomological Society, 21(1/2) (February/April): 18-26.
189 Pilophorus fuscipennis, NEW SPECIES (pp. 23-24); not illustrated
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
(Holotype and allotype from near Trinidad, Colorado.) “Paratypes: . . .
♂ ♀, Aug. 3, 1917, Grand
View, Grand Canyon, Arizona (H. H. Knight). Paratypes in collections of Iowa State college and
Colorado Agricultural College.”
_______________________________________________________________
1928
New species of Phytocoris from North America (Hemiptera, Miridae). Bulletin of the
Brooklyn Entomological Society, 23(1) (February): 28-46.
190 Phytocoris hesperius, NEW SPECIES (pp. 44-45); not illustrated
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
(Holotype from near Trinidad, Colorado; allotype from Chiricahua Mountains, Arizona.) P aratype
specimens include “2♂ 1♀ Aug. 2, 1917, at top of Bright Angel trail, Grand Canyon (H. H.
Knight) . . . .”
_______________________________________________________________
1929
Descriptions of five new species of Plagiognathus from North America (Hemip.: Miridae).
Entomological News (Academy of Natural Sciences of Philadelphia, Entomological
Section), 40 (March): 69-74.
191 Plagiognathus tenellus, NEW SPECIES (p. 73); not illustrated
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
“Holotype:
♂ August 2, 1917, top of Bright Angel trail, Grand Canyon, ARIZONA (H. H. Knight);
♂ ♀, taken with the types . . . .”
author's collection. Allotype: taken with the type. Paratypes: 12
_______________________________________________________________
74
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1934
Phytocoris Fallen—twelve new species from the western United States (Hemiptera,
Miridae). Bulletin of the Brooklyn Entomological Society, 29(1) (February): 1-16.
192 Phytocoris varius, NEW SPECIES (pp. 9-11); not illustrated
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
♂ September 6, 1931, Grand Canyon (H. H. Knight); author’s collection. Allotype:
♀, taken with the type. Paratypes: ♂, 3♀, taken with the types by beating on large cedar trees
“Holotype:
(Juniperus sp.) which were found growing behind the cabin camp located at the entrance
gate of the Grand Canyon National Park.” (Additional paratypes from Durango, Colorado and
Chiricahua Mountains, Arizona.)
_______________________________________________________________
1968
Taxonomic review: Miridae of the Nevada Test Site and the western United States.
Brigham Young University Science Bulletin (Biological Series), 9(3), 282 pp.
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
193 Parthenicus cowaniae, NEW SPECIES (pp. 148-149, fig. 211 [p. 147])
♂ Aug. 3, 1917, Grand View, Grand Canyon, Arizona (H. H. Knight), taken on cliff
rose, Cowania stansburiana, which is the host plant of the species. Allotype: ♀ taken with the
type. Paratypes: “26 ♂ 20 ♀ taken with the types on cliff rose . . . ♂ ♀ Sept. 6, 1931, Grand
View, Grand Canyon (H. H. Knight) . . . ♂ ♀ Aug. 1, 1914, Hermit Rim road, Grand Canyon,
“Holotype:
Arizona (L. C. Bradley).”
194
Bolteria juniperi,
NEW SPECIES
(pp. 202-203, fig. 253)
(Holotype, allotype, and paratypes from Scipio, Utah.) Other paratypes include “. . .
♂ June 16,
alt 8000 ft, near Grand Canyon, Arizona.” [Given the altitude data for this paratype, and the
fact that the primary types are from Utah, it is presumably on the northern side of the canyon.]
195 Phytocoris flaviatus, NEW SPECIES (p. 241, fig. 297 [p. 242]
“Holotype:
♂ Sept. 6, 1931, above Bright Angel Trail, Grand Canyon, Arizona (H. H. Knight).”
[No other type material.]
________________________________________________________________________________
Knull, Josef N.
1957
Three new species of Aphricus with a note on Ctenicera (Coleoptera: Elateridae). Ohio
Journal of Science, 57(4) (July): 200-202.
196
Aphricus knowltoni,
NEW SPECIES
(pp. 200-201); not illustrated
Arthropoda: Insecta: Coleoptera: Elateridae (click beetles)
(Holotype and paratypes from Washington, Utah.) Another paratype from “Cameron, Arizona,
Aug. 19, 1950, D. J. and J. N. Knull, all in collection of author.”
________________________________________________________________________________
75
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Kruidenier, Francis J., and Mehra, Krishna N.
1958
Wellcomia perognathi n. sp. (Nematoda: Oxyuridae) from pocket mice, Perognathus
intermedius (Merriam, 1889) of the Grand Canyon, Arizona. Transactions of the Illinois
State Academy of Science, 51(3/4) (July 10): 20-25.
197 Wellcomia perognathi, NEW SPECIES; illustrated
Nematoda: Chromadorea: Ascaridida: Oxyuridae (pinworms)
“Host: Perognathus intermedius (Merriam, 1889). Habitat: Intestine (cecum). Locality: Coconino
County, Arizona (Grand Canyon National Park). Holotypes [sic]: U.S. Nat’l. Mus. Helm. Coll.
No. 38306.” [Limited data, thus. Collection made in 1954; see note with no. 198, below.]
_______________________________________________________________
1959
Aspiculuris ackerti, n. sp, (Nematoda: Oxyruidae) from the wood rats of Arizona.
Proceedings of the Helminthological Society, 26(2) (July): 147-150.
198 Aspiculuris ackerti, NEW SPECIES; illustrated
Nematoda: Chromadorea: Ascaridida: Oxyuridae (pinworms)
“Hosts: Neotoma albigula—U. of Ill. Museum #11650 [and] N. cinerea—U. of Ill. Museum #11688.
Habitat: Intestine. Locality: Coconino County, Arizona. Holotype: U.S. Nat’l Mus. Helm. Coll.
#56163.”
“. . . recovered from a series of neotomid rats examined during a 1954 expedition to the Grand
Canyon, Coconimo [sic] County, Arizona (see Kruidenier and Peebles, 1958).” The cited item does
not appear in the Literature Cited section of this paper, but is identified (here) as the citation with
no. 199, below, which there provides locality data only as “Grand Canyon National Park”.
________________________________________________________________________________
Kruidenier, Francis J., and Peebles, C. R.
1958
Gongylonema of rodents: G. neoplasticum (redefinition); G. dipodomysis n. sp.; and G.
peromyski n. sp. Transactions of the American Microscopical Society, 77(3) (July): 307315.
199 Gongylonema peromysci, NEW SPECIES (pp. 310-312, Pl. 2, figs. 1-6 [p. 313])
Nematoda: Chromadorea: Spiurida: Gongylonematidae (spiurid gullet worms)
“Host: Type: Peromyscus maniculatus rufinus (Merriam, 1890) Osgood, 1909. Univ. Ill. Mus. No.
10832 . . . .
Habitat: Mucosa of cardiac stomach.
Coconimo [sic] County, Arizona.
Locality: Grand Canyon National Park,
Holotypes [sic]: U.S. National Museum Helminthological
Collection No. 38202.” [Collection made in 1954; see note with no. 198, above.]
________________________________________________________________________________
76
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
LaBerge, Wallace E.
1967
A revision of the bees of the genus Andrena of the western hemisphere. Part I.
Callandrena. (Hymenoptera: Andrenidae). Bulletin of the University of Nebraska State
Museum, 7 (October): 1-318.
200 Andrena (Callandrena) utahensis, NEW SPECIES (pp. 258-260, figs. 292-296 [p. 314])
Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees)
(Holotype and allotype from Duchesne, Utah.)
Grand Canyon. 5
Additional paratypes include: “ARIZONA: . . .
♀♀, 1 ♂, June 5, 1940, G. E. Bohart . . . .”
________________________________________________________________________________
Levine, Norman D.; Ivens, Virginia; and Kruidenier, Francis J.
1957a
New species of Eimeria from Arizona rodents. Journal of Protozoology, 4(2) (May): 8088.
Apicomplexa: Conoidasida : Eucoccidiorida: Eimeriidae (coccid parasites)
201 Eimeria tamiasciuri, NEW SPECIES (pp. 80-81, fig. 1 [p. 82])
“Host: Tamiasciurus hudsonicus mogollonensis (U. of Ill. Mus. #10,432) (red or spruce squirrel).
Location: Intestinal contents. Locality: The host animal was trapped by Mr. Woodrow Goodpaster
on Aug. 7, 1954 on Road W-3 on the north rim of Grand Canyon near the entrance to
Grand Canyon National Park, Arizona.”
202 Eimeria cutamiae, NEW SPECIES (pp. 82-83, fig. 3)
“Host: Eutamias dorsalis dorsalis (U. of Ill. Mus. #10,386) (cliff chipmunk). Location: Intestinal
contents. Locality: This species was found in one of two Eutamias d. dorsalis examined in the
Grand Canyon National Park, Arizona. The host chipmunk was trapped by Mr. K. R. Nelson on
July 4, 1954 at Hull Tank, on the south rim of Grand Canyon.”
203 Eimeria thomomysis, NEW SPECIES (p. 83, fig. 4 [p. 82])
“Host: Thomomys bottae fulvus (U. of Ill. Mus. #10,568 and #7990) (pocket gopher). Location:
Intestinal contents.
Locality: This species was found in two out of five T. bottae examined in
Arizona. One animal was trapped by Mr. Donald Hoffmeister on July 4, 1954 at Hearst Ranch
[Grandview] on the south rim of Grand Canyon in Grand Canyon National Park.”
204 Eimeria perognathi, NEW SPECIES (pp. 83-84, fig. 5 [p. 82])
“Host: Perognathus intermedius subsp. (U. of Ill. Mus. #10,640) (rock pocket mouse). Location:
Intestinal contents. Locality: This species was found in one out of two P. intermedius examined
from the Grand Canyon National Park, Arizona. The host animal was trapped by Dr. Donald F.
Hoffmeister on July 23, 1954 along the Plateau Point Trail one mile below [sic] Indian Garden
[Havasupai Gardens] inside Grand Canyon on the south side of the Colorado River.”
77
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
205 Eimeria albigulae, NEW SPECIES (p. 84, fig. 6 [p. 85])
“Host: Neotoma albigula subsp. (U. of Ill. Mus. #11,650) (white-throated woodrat). Location:
Intestinal contents. Locality: This species was found in a N. albigula in Grand Canyon National
Park, Arizona. The host woodrat was collected by Mr. K. R. Nelson on July 11, 1954 on the east
side of Cedar Mountain, on the south rim of Grand Canyon [sic].” [The precise collecting locality
may be within the boundaries of the Navajo Indian Reservation.]
206 Eimeria operculata, NEW SPECIES (pp. 84-85); not illustrated
“Host: Neotoma stephensi stephensi (U. of Ill. Mus. #11,636) (Stephens’ woodrat).
Location:
Intestinal contents. Locality: This species was found in one out of two N. s. stephensi examined
in Grand Canyon National Park, Arizona. The host woodrat was collected by Mr. K. R. Nelson on
July 5, 1954 at Zuni Point, on the south rim of Grand Canyon.”
207 Eimeria peromysci, NEW SPECIES (pp. 85-86, fig. 7)
“Host: Peromyscus truei truei (U. of Ill. Mus. #11,249) (piñon mouse).
Location: Intestinal
contents. Locality: This species was found in one out of four P. t. truei examined in Grand Canyon
National Park, Arizona. The host mouse was collected by Dr. Donald F. Hoffmeister on July 5,
1954 on the west side of Zuni Point, which is on the south rim of Grand Canyon.”
208 Eimeria arizonensis, NEW SPECIES (p. 86, fig. 8 [p. 85])
“Host: Peromyscus truei truei (U. of Ill. Mus. #11,220 and 11,249) (piñon mouse).
Location:
Intestinal contents. Locality: This species was found in two of four P. t. truei examined in Grand
Canyon National Park, Arizona.
The host mouse form which the larger series of oocysts was
obtained was collected by Dr. Donald F. Hoffmeister on August 5, 1954 at Cape Royal on the
north rim of Grand Canyon. The host mouse from which the smaller series of oocysts was obtained
was collected by Dr. Hoffmeister on July 5, 1954 on the west side of Zuni Point, which is on the
south rim of Grand Canyon. This same individual was infected with E. peromysci [see no. 207,
above].”
209 Eimeria eremici, NEW SPECIES (pp. 86-87, fig. 9 [p. 85])
“Host: Peromyscus eremicus eremicus (U. of Ill. Mus. #11,354) (cactus mouse).
Location:
Intestinal contents. Locality: This species was found in one out of two P. eremiscus Grand Canyon
National Park, Arizona. The host mouse was collected by Dr. Donald F. Hoffmeister on July 23,
1954 on the Tonto Plateau along the Plateau Point Trail one mile below [sic] Indian Garden
[Havasupai Gardens]; this location is inside the Grand Canyon, on the south side of the Colorado
River.”
210 Eimeria onychomysis, NEW SPECIES (p. 87, fig. 10 [p. 85])
“Host: Onychomys leucogster subsp. (U. of Ill. Mus. #10,666) (northern grasshopper mouse).
Location: Intestinal contents. Locality: This species was found in one out of two O. leucogaster
examined in Grand Canyon National Park, Arizona. The host mouse was collected by Dr. Donald
F. Hoffmeister on July 19, 1954 along State Highway 64 at the southeast boundary of the
park, south of the canyon.”
_______________________________________________________________
78
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1957b
Isopora citolli n. sp. from the rock squirrel, Citellus variegatus utah. Journal of
Protozoology, 4(3) (August): 143-144.
211 Isopora citolli, NEW SPECIES (p. 87, fig. 1)
Apicomplexa: Conoidasida : Eucoccidiorida: Eimeriidae (coccid parasites)
“The host rock squirrel was trapped by Dr. Hoffmeister on August 9, 1954 one-half mile from
Swamp Point, near Powell Saddle, on the north rim of Grand Canyon at an elevation of 7500
feet. Its skin and skull were preserved by Dr. Hoffmeister, and are in the University of Illinois
Natural History Museum (Museum No. 10220).”
________________________________________________________________________________
Lindsay, Dale R.
1938
New species of Norvellina (Homoptera, Cicadellidae). Journal of the Kansas
Entomological Society, 11(4) (October): 113-123.
212 Norvellina bicolorata (Ball) var. inflata, NEW VARIETY (pp. 113-114); not illustrated
Arthropoda: Insecta: Hemiptera: Cicadellidae (leafhoppers)
(Holotype and allotype from White Sands, New Mexico.) “Numerous paratypes from the following
localities: Grand Canyon, Ariz. . . . . Types and paratypes in the Snow Entomological Collection.
Paratypes in collection of Dr. Ball and U. S. N. M.”
________________________________________________________________________________
Liu, Tong-Xian, and Kosztarab, Michael
1987
Two new species of Chionaspis (Homoptera: Coccoidea: Diaspididae) from North America.
Florida Entomologist, 70(4) (December): 512-520.
213 Chionaspis gilli, NEW SPECIES (pp. 516-519, fig. 2)
Arthropoda: Insecta: Hemiptera: Diaspididae (armored scales)
(Holotype from Las Cruces, New Mexico.) “Paratypes: On Tamarix chinensis, 5 on 4 slides, Grand
Canyon National Park, Colorado River, Coconino Co., Arizona (AZ), VII-23-1984 collected by
L. E. Stevens, deposited in USNM . . . .”
________________________________________________________________________________
Marshall, William B.
1929
Three new land shells of the genus Oreohelix from Arizona. Proceedings of the U.S.
National Museum, 76(2802), 3 pp.
214
Oreohelix yavapai vauxae,
NEW SUBSPECIES
(pp. 1-2, Plate1, figs. 1-3, 11).
Mollusca: Gastropoda: Stylommatophora: Oreohelicidae (mountain snails)
“The study of a collection of Oreohelix made by Mrs. Mary Vaux Walcott in the canyon at Supai,
Coconino County, Ariz., in 1928, and presented by her to the United States National Museum, not
79
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
only proved that they belonged to a new subspecies, but their examination entailed a close
scrutiny of forms long since contained in our collection but not previously adequately studied.”
“The type [holotype] and 14 paratypes come from the Canyon at Supai, Coconino County,
Ariz., and were collected and presented by Mrs. Charles D. Walcott, whose maiden name has
been bestowed upon it.”
A table provides measurements for the “type and those of the paratypes which are adult”, giving
the USNM catalog numbers: 380687 (type), 380688 (eight adult paratypes in one lot; presumably
the sub-adult paratypes are in the same lot).
________________________________________________________________________________
Massey, Calvin L.
1974
Biology and taxonomy of nematode parasites and associates of bark beetles in the United
States. U.S. Forest Service, Agriculture Handbook 446, 233 pp.
215 Parasitorhabditis gracilis, NEW SPECIES (p. 69, fig. 41 [p. 70])
Nematoda: Chromadorea: Rhabditida : Rhabditidae (rhabditid roundworms)
“Type habitat.—Associated with Pseudohylesinus grandis Sw. in white fir. Type locality.—Grand
Canyon, Arizona. Type specimens.—Collection No. 84-J.”
________________________________________________________________________________
McAtee, W. L. [Waldo Lee]
1919
Notes on Nearctic Heteroptera. Bulletin of the Brooklyn Entomological Society, 14(1)
(February): 8-15.
216 Chelinidea vittiger Uhler var. artuflava, NEW VARIETY (pp. 11-12); not illustrated
Arthropoda: Insecta: Hemiptera: Coreidae (leaf-footed bugs)
Syntypes include “Grand Cañon, Ariz., July 10 (U. S. N. M.)”.
[No further data about this
specimen.]
________________________________________________________________________________
McKelvey, Susan Delano
1949
A new Agave from Arizona. Journal of the Arnold Arboretum, 30(3) (July): 227-230, 2
plates.
217 Agave kaibabensis, NEW SPECIES (pp. 227-228, Pl. 1)
“ARIZONA, Coconino County, Kaibab Plateau on the north side of the Grand Canyon of the
Colorado River, McKelvey 4381 (Arnold Arboretum, TYPE [holotype]), 4381A (Arnold Arboretum)
[paratype], May 15, 1934.” [Apparently in Grand Canyon National Park. See notes below, which
suggest the author’s casualness with geographical and records concerning the gathering.]
80
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
McKelvey is not specific about the locality of the type, although she offers somewhat confusing
additional data in her text from which one may infer that the type is from Grand Canyon National
Park and not from the Kaibab National Forest north of the park.
“On May 15, 1934, while visiting the Kaibab Plateau on the northern side of the Grand Canyon of
the Colorado, Coconino County, Arizona, the writer first saw the Agave here described. Two plants
had been collected by the Park authorities at some previous date, but their leaf-rosettes and long
fruiting stalks were still intact. It was not possible to make the descent into the gorge, but a few
scattered plants were discernible some distance below the rim. Flowers were promised at a later
date but were never forthcoming, unfortunately. [¶]The two plants are shown in the included
photograph (McKelvey 152-12); although one was considerably smaller than the other it was
undoubtedly the same species. The dimensions are from the larger plant.” (p. 227).
“The smaller plant of the picture [which is posed (!?) beside the type in Pl. I] is McKelvey 4381A.
The plant, as noted [p. 227?], came originally from some distance below the rim on the north wall
of the Canyon. The date of actual collection is not known; it must have been in the season of
1933; the writer made her specimens on May 15, 1934. Her photographs 152-11, 12, and 1531, 2 are of these plants; reproduced here is 153-2 (Pl. I)” (p. 229). Note that on p. 227 McKelvey
indicates that the photograph of the two plants is her photo 152-12. [Pl. II of this paper depicts
a comparative specimen of Agave utahensis Engelmann, gathered by McKelvey in 1929 “some 13
miles east of Peach Springs, Mohave County, Arizona, in the extreme northwestern corner of
Yavapai County.”]
________________________________________________________________________________
Merriam, C. Hart [Clinton Hart]
1890
Results of a biological survey of the San Francisco Mountain region and the desert of the
Little Colorado River, Arizona. U.S. Department of Agriculture, North American Fauna,
(3), 136 pp., 13 plates, 5 maps.
Regarding the localities noted at Grand Canyon, Merriam describes the trip to the canyon thusly
(p. 4, in its entirety): “TRIP TO THE GRAND CAÑON OF THE COLORADO, SEPTEMBER 9 TO 16,
INCLUSIVE. The usual road was followed from Little Spring to Hull Spring and Red Horse Tank,
and thence to the tank known as Cañon Spring on the Cocanini [sic] Plateau, close to the cañon,
which is here about 1,800 meters (6,000 feet) in depth. Mr. Bailey and myself climbed down into
the cañon and remained in it two days and two nights.” This information is enough to identify the
locale as John Hance’s accommodations at the head of the Old Hance Trail, down which Merriam
and Bailey descended.
218 Hesperomys megalotis, NEW SPECIES (pp. 63-64, Pl. III, figs. 1-4; Pl. IV) [holotype
illustrated]
Chordata: Mammalia: Rodentia: Crecitidae (deer mice)
(Type [holotype] from Black Tank, Desert of the Little Colorado River.)
Paratype specimens
include USNM 17930/24841 (original no. 469), “Grand Cañon, Arizona, Sept. 11, 1889,
81
♀ ad.”
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
219 Spilogale gracilis, NEW SPECIES (pp. 83-84); not illustrated.
(Common name given by the
author, “Little Striped Skunk”)
Chordata: Mammalia: Carnivora: Mephitidae (skunks)
“Type [holotype] No. 17986/24897
♂
ad. U. S. National Museum (Department of Agriculture
collection). From Grand Cañon of the Colorado (altitude 3,500 feet), Arizona, north of San
Francisco Mountain, September 12, 1889. Collected by C. Hart Merriam (Original number 451).”
Also lists USNM 18598/25368 “♂ im.” [paratype] also from Grand Canyon.
Merriam remarked (p. 83): “While asleep near a small spring in the Grand Canon of the Colorado,
September 12, 1889, I was awakened at midnight by a sniffling noise about my head. Rising
suddenly on my elbow, a small animal scampered hurriedly away over the rocks. His form was
only dimly outlined in the dark, but a hasty shot left no doubt as to his identity, and a moment
later I held in my hand the type of a new species of Little Striped Skunk. A day or two afterward
a younger individual was captured among the cliffs at the top of the Cañon. The stomachs of both
contained remains of the cliff mouse Hesperomys erenucus).”
_______________________________________________________________
1904
Two new squirrels of the aberti group. Proceedings of the Biological Society of
Washington (Washington, D.C.) 17: 129-130.
220 Sciurus kaibabensis, NEW SPECIES (pp. 129-130); not illustrated.
Chordata: Mammalia: Rodentia: Sciuridae (squirrels)
“Type [holotype] from head of Bright Angel Creek, top of Kaibab Plateau, north side of Grand
Canyon of Colorado [sic], Arizona. Adult male, No. 130,932, U. S. National Museum, Biological
Survey Collection.
December 1, 1903.
John T. Stewart.”
[This is the widely known “Kaibab
Squirrel”.]
________________________________________________________________________________
Miller, Robert Rush
1946
Gila cypha, a remarkable new species of cyprinid fish from the Colorado River in Grand
Canyon, Arizona. Journal of the Washington Academy of Sciences (Washington, D.C.),
36(12): 409-415.
221 Gila cypha, NEW SPECIES; illustrated
Chordata: Teleostei: Cypriniformes: Cyprinidae (carps)
“Holotype.—The holotype (U.S.N.M. no. 131389) is a specimen about 305 mm in standard length
and was taken by N. N. Dodge near Phantom Ranch in the western end of Grand Canyon National
Park [old boundaries], Arizona. It was caught in swift water on hook and line, presumably in the
nearby Colorado River at or near the mouth of Bright Angel Creek.” [This is the widely
known “Humpback Chub”.]
________________________________________________________________________________
82
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Miller, Walter B.
1984
Three new species of Sonorella (Gastropoda: Pulmonata: Helminthoglyptidae) from
Arizona. Proceedings of the Biological Society of Washington (Washington, D.C.), 97(4):
681-687.
222 Sonorella reederi, NEW SPECIES (pp. 681-684, figs. 1A-C, 2)
Mollusca: Gastropoda: Stylommatophora: Helminthoglyptidae (shoulderbands)
“Disposition of types.—Holotype: USNM 792406. Paratypes: ANSP 356004; CAS 033405; FMNH
206235; UTEP 9051; RLR 298; WBM 5241 & 6307.”
“Type-locality.—Mohave Co., Arizona. Lower Granite Gorge of the Colorado River, in limestone
rockslide just west and below Rampart Cave; elevation ca. 1700 ft., 36°06′N, 113°56′W.”
________________________________________________________________________________
Moldenke, Andrew R.
1970
A revision of the Clytrinae of North America north of the Isthmus of Panama (Coleoptera:
Chrysomelidae). Stanford, California: Stanford University, 310 pp.
223
Saxinis saucia kaibabiae,
NEW SUBSPECIES
(pp. 182-
; illustrated)
Arthropoda: Insecta: Coleoptera: Chrysomelidae (leaf beetles)
Jacob Lake, Kaibab Forest.
________________________________________________________________________________
Monod, M. [Michel]
1983
Monographie taxonomique des Gnomoniaceae (ascomycès de l’ordre des Diaporthales). I.
Beihefte zur Sydowia (Horn, Niederösterreich, Austria), (Annales Mycologici, Ser. II), IX.
Beiheft, 315 pp. [This publication originally was Monod’s doctoral dissertation, Université
de Lausanne”.] [Monograph is in French; species descriptions are in botanical Latin and
in French.]
224 Gnomonia quercusgambellii, NEW SPECIES (pp. 98-99, fig. 13.5 [p. 278])
Ascomycota (sac fungi): Sordariomycetes: Diaporthales: Valsaceae
“. . . sous Gnomonia fasciculata: Quercus gambellii: North Rim of Grand Canyon, Arizona, 18
août 1973, Barr, 6095 (MASS, Holotype).”
________________________________________________________________________________
83
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Muchmore, William B.
1981
Cavernicolous species of Larca, Archeolarca, and Pseudogarypus with notes on the
genera, (Pseudoscorpionida, Gaypidae and Pseudogarypidae). Journal of Arachnology,
9(1): 47-60.
225 Archeolarca cavicola, new species (pp. 55-56, figs. 11, 12 [p. 53])
Arthropoda: Euchelicerata: Pseudoscorpiones (pseudoscorpions): Larcidae
“Holotype female (WM 5398.01001) from Cave of the Domes [Horseshoe Mesa], Grand Canyon
National Park, Coconino County, Arizona, 15 April 1978 (W. Calvin Welbourn). The type is in the
Florida State Collection of Arthropods, Gainesville.”
________________________________________________________________________________
Nabokov, Vladimir
1942
Some new or little known Nearctic Neonympha (Lepidoptera: Satyridae). Psyche,
49(3/4) (September/December): 61-80.
Neonympha dorothea
NEW SPECIES
(pp. 62-63)
226 Neonympha dorothea dorothea, NEW SUBSPECIES (pp. 64-66); not illustrated
Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals)
“Male, holotype, female, allotype, and two males, paratypes, placed in the American Museum of
Natural History. Taken during a brief visit to Grand Cañon, Ariz., South Rim, on June 9th, 1941
(bright cold morning after snow and rain). They were weakly fluttering beside the trail together
with a few Coen[onympha]. tullia furcae almost no other butterflies about. Named in honor of
Miss Dorothy Leuthold who kindly kicked up the first specimen. Female, paratype, labelled ‘Grand
Cy., June 11th ’30’, ex Coll. of C.F. dos Passos, Am. Mus. Nat. Hist.”
[Two additional subspecies of N. dorothea also named therein, which are not pertinent to the
present checklist.]
________________________________________________________________________________
Nelson, Edward W., and Goldman, Edward A.
1931
Three new pumas. Journal of the Washington Academy of Sciences (Washington, D.C.),
21(10) (May 19): 209-212.
227 Felis concolor kaibabensis, NEW SUBSPECIES (pp. 209-210); not illustrated.
(Common
name given by the author, “Kaibab Mountain Lion”)
Chordata: Mammalia: Carnivora: Felidae (cats)
“Type [holotype].—From Powell Plateau, Grand Canyon National Park, Arizona (altitude 8,700
feet). No. 171186
♂ adult, U. S. National Museum (Biological Survey collection), collected by J.
T. Owens, April 15, 1911. X number 8432.”
________________________________________________________________________________
84
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Nelson, G. H., and Westcott, R. L.
1995
Three new species of Acmaeodera Eschscholtz (Coleoptera: Buprestidae) from the United
States and Mexico. The Coleopterists Bulletin, 49(1): 77-87.
228
Acmaeodera navajo,
NEW SPECIES
(pp. 82-84; figs. 2 [p. 78], 7 [p. 80])
Arthropoda: Insecta: Coleoptera: Buprestidae (jewel beetles)
“The holotype
♀ (USNM) and 90 paratypes, Arizona: Coconino Co., 9 mi NW Page, 3-VI-70, R.
L. Westcott, on either purple-flowered Opuntia or flowers Sphaeralcea. Other paratypes are as
follows: . . . Arizona: Coconino Co., Vermillion [sic] Cliffs [Vermilion Cliffs], 9-VII-67, J. H. & J.
M. Davidson, M. Cazier (1).”
Non-type specimens examined include “Navajo Indian Reservation, Hwy. 89, 15 mi S Page, 26 VI-80, L. L. Lampert, on Linum lewisii; 8 mi. SW Cliff Dwellers Lodge, 11-VII-67, M. A. Cazier, J.
H. and J. M. Davidson, on Sphaeralcea sp. . . . .”
[It is unclear whether “Vermillion Cliffs” refers to the geographical feature or to the place name,
Vermilion Cliffs, although note that the authors also specially mention nearby Cliff Dwellers
Lodge.]
_____________________________________________________________________________ ___
Oberholser, Harry C.
1937
Descriptions of three new screech owls from the United States. Journal of the
Washington Academy of Sciences (Washington, D.C.), 27(8) (August 15): 354-357.
229 Otus asio mychophilus, NEW SUBSPECIES (pp. 356-357); not illustrated.
(Common name
given by the author, “Grand Canyon Screech Owl”)
Chordata: Aves: Strigiformes: Strigidae (typical owls)
“Type [holotype].—Adult female, No. 340593, U. S. National Museum, Biological Survey
collection; south rim of Grand Canyon, 6,900 feet altitude, Grand Canyon Village, Arizona;
January 28, 1935; Russell K. Grater, original number, 23.”
________________________________________________________________________________
Pape, Robert B.; Thomas, Donald B.; and Aalbu, Rolf L.
2007
A revision of the genus Eschatomoxys Blaisdell (Tenebrionidae: Pimeliinae: Edrotini) with
notes on the biology. The Coleopterists Bulletin, 61(4): 519-540.
230 Eschatomoxys pholeter Thomas & Pape, NEW SPECIES (pp. 525-527, figs. 1 [p. 521], 5 [p.
523], 8A,B [p. 526])
Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles)
“Holotype. Male. ARIZONA: Mohave Co., Grand Canyon N.P., Bat Cave, 29-III-96, R.B. Pape
coll. Allotype. Female. Labeled same data as holotype. Paratypes: Three males, one female,
labeled same data as holotype; Four males, six females: AZ. Mohave Co., Bat Cave, Gr. Canyon,
GRTCA Cave SCI-0043, 26-X-01, R.B. Pape (deposited CAS and DB Thomas collection).
One
female: AZ. Mohave Co., Eldel Cave [Grand Canyon–Parashant National Monument], twilight, IX-
85
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
23-05, J. Judson Wynne col[l].
Three females: ARIZONA: Mohave Co., Grand Canyon N.P.,
Rampart Cave, teneb on 7,200 yr old sloth shit [sic!].”
________________________________________________________________________________
Parfitt, Bruce D.
1997
Opuntia pinkavae (Cactaceae), a new species from Arizona and Utah. Rhodora (Journal
of the New England Botanical Club), 99(899) (Summer): 223-228.
231 Opuntia pinkavae, NEW SPECIES; illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
“HOLOTYPE: USA, Arizona: Mohave Co., northwest of Bulrush Canyon south of Pipe Spring,
1400 m, n =44, 30 May 1980, B. D. Parfitt 2874 (ASU 111287!).”
“PARATYPES: United States.
Arizona: Coconino Co., House Rock Valley, N of Rock Canyon,
Parfitt 2859 (ASU!); 1 mi N of Fredonia, Brown 651 & Parfitt (ASU!); Mohave Co., 1.2 mi N of
hwy 389 on road to Moccasin, Brown 657 & Parfitt (ASU!); W of Kaibab Indian Reservation,
Parfitt 3958 & Roberts (ASU!); Main Street Valley, W of Hurricane Cliffs, Palmer & Hodgson
4620 (ASU!, DES); Navaho Trail near Hurricane Rim, Gierisch 5132 (ASU!); 4.2 mi SW of
Wolf Hole, Brown 851 & Parfitt (ASU!).” (Additional paratypes from southern Utah.)
________________________________________________________________________________
Peck, Stewart Blaine
1973
A systematic revision and the evolutionary biology of the Ptomaphagus (Adelops) beetles
of North America (Coleoptera; Leiodidae; Catopinae), with emphasis on cave-inhabiting
species. Bulletin of the Museum of Comparative Zoology, 145(2): 29-162.
232 Ptomaphagus cocytus, NEW SPECIES (p. 106, figs. 39, 40 [both p. 93, legend p. 92], 117
[p. 99, legend p. 98], 164 [p. 103, legend p. 102], 199 [p. 105, legend p. 104])
Arthropoda: Insecta: Coleoptera: Leiodidae (round fungus beetles)
“Holotype male and allotype female in MCZ (no. 31893). Type locality: Arizona, Coconino County,
Kaibab Plateau, Roaring Springs Cave. Type data: 1.i.1965, Gregory Lane leg. Paratypes, 2
females, same locality, 16.iv.1965, L. Ball and G. T. Lane leg., in SBP.”
________________________________________________________________________________
Peck, Stewart B., and Wynne, J. Judson
2013
Ptomaphagus parashant Peck and Wynne, new species (Coleoptera: Leiodidae:
Cholevinae: Ptomaphagini): the most troglomorphic cholevine beetle known from western
North America. The Coleopterists Bulletin, 67(3): 309-317.
233
Ptomaphagus parashant,
NEW SPECIES ;
illustrated
Arthropoda: Insecta: Coleoptera: Leiodidae (round fungus beetles)
“Type Specimens. Holotype female and allotype male deposited in Canadian Museum of Nature
(CMN), Ottawa, Canada.
Type locality: USA, Arizona, Mohave County, Parashant National
86
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Monument [Grand Canyon-Parashant National Monument], PARA-1001 cave, N36°39′,
W113°37′. Type label data: female holotype; 2.ix.2011, J.J. Wynne, sweet potato bait #4, site
2, deep zone; and male allotype, same area but chicken liver bait #1.
Our red label states
‘Ptomaphagus parashant Peck & Wynne, holotype.’ Paratypes, all with yellow paratype labels,
seven with same locality data and five from site 2, direct intuitive searching, and two from ‘sweet
potato bait #4’ are deposited at the CMN, the Los Angeles County Museum, California Academy
of Sciences, and the U.S. National Museum of Natural History (Smithsonian Institution).”
________________________________________________________________________________
Pennell, Francis W.
1940
New species of Scrophulariaceae from Arizona. Notulae Naturae of the Academy of
Natural Sciences of Philadelphia, (43) (May 15), 10 pp.
234 Cordylanthus tenuifolis, NEW SPECIES (pp. 9-10); not illustrated
Trachyophyta: Magnoliopsida: Lamiales: Orobanchaceae (broomrapes)
“Type [holotype].—Grand Canyon, Coconino County, Arizona, collected in flower August 24,
1919, by W. W. Eggleston, no. 15677a; in Herb. Academy of Natural Sciences of Philadelphia.”]
________________________________________________________________________________
Phillips, Arthur M., III
1992
A new subspecies of Rosa stellata (Rosaceae) from northwestern Arizona. Madroño,
39(1): 31-35.
235 Rosa stellata Wooton subsp. abyssa, NEW SUBSPECIES; illustrated
Trachyophyta: Magnoliopsida: Rosales: Roasaceae (roses)
“TYPE: USA, Arizona, Mohave Co.: SW edge of Shivwits Plateau, along W rim of Twin Point, 18
km S of Oak Grove, T30N R12W NW¼ sect. 7, 36°01′N, 113°37′W, 1823 m, in sandy to
gravelly soils with limestone chips, derived form Kaibab limestone, in first 100 m from edge of
plateau, open Great Basin conifer woodland, 15 Jun 1980, A. M. Phillips, III, 80-103 (holotype,
ARIZ; isotypes, ASC, ASU, DES, MNA, MO, NY, UNLV, UNM, US, Lake Mead National Recreation
Area herbarium).” [The locality is today within Grand Canyon-Parashant National Monument.]
“Paratypes. USA, AZ, Mohave Co.: type locality . . . ; 27 Jul 1975, Holland 690 (UNLV); W rim
of Kanab Canyon, gravelly soil, T38N R3W sect. 30, 1585 m., 2 Aug 1977, Gierisch 3978 (ASC,
ASU); 8 Jun 1978, Gierisch 4388 (ARIZ, ASU, USDA Forest Service Herb., Albuquerque, NM); W
rim of Kanab Canyon in small drainage 30 m from edge, restricted to Kaibab limestone
conglomerate, T38N R3W NW¼ sect. 29, 1550 m, 13 Jun 1979, A. Phillips and B. Phillips 79624 (MNA); W rim of Kanab Canyon S of Water Canyon, in depression caused by breccia
pipe collapse, T38N R3W NW¼ SW¼ sect. 8, 1525 m, 22 May 1980, A. Phillips 80-91 (ARIZ,
ASC, ASU, MNA, UNLV, UNM, BLM Arizona Strip District herbarium); Grand Canyon National
Monument (Park) [now in the enlarged Grand Canyon National Park], head of SB Trail, 1372 m,
24 May 1958, Riffey s.n. (UNLV, COLO); between SB Point and Hades Knoll, side canyon,
1675 m, 30 May 1978, Reichhardt 123 (MNA). Coconino Co.: Mesa Eremita [Eremita Mesa] S
rim of Grand Canyon, 1980 m, 12 Jun 1935, Hawbecker s.n. (Grand Canyon National Park Study
87
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Collection); Dutton Point, N rim of Grand Canyon, dry ledge overlooking canyon, 2285 m, 17 Jul
1947, Bryant and Cooper s.n. (Grand Canyon National Park Study Collection, 2 specimens).”
________________________________________________________________________________
Pilsbry, Henry A., and Ferriss, James H.
1911
Mollusca of the southwestern states, V: The Grand Canyon and northern Arizona.
Proceedings of the Academy of Natural Sciences of Philadelphia, 63: 174-199.
236 Oreohelix yavapai profundorum, NEW SUBSPECIES (pp. 182-184, text-fig. 4; Pl. XII, figs.
1-14 [legend p. 198])
Mollusca: Gastropoda: Stylommatophora: Oreohelicidae (mountain snails)
“Grand Canyon, in ‘Spectacle Cove,’ Station A, the head of a recess in the cross-bed
sandstone [Coconino Sandstone] south of where the Mystic Spring or Bass Trail [South
Bass Trail] zigzags down, in a talus resting on the red sandstone forming the Le Conte
Plateau. Elevation about 5,700 feet. Cotypes [syntypes], No. 103,234 A. N. S. P., collected by
Ferriss and Pilsbry, October, 1906.” (Text-fig. 5 is a sketch view of “Spectacle Cove (Station A),
from opposite side below Bass Trail. Type locality of Oreohelix yavapai profundorum on the mound
at left end of talus slope.”)
237 Oreohelix yavapai extremitatis, NEW SUBSPECIES (pp. 184-185, Pl. XII, figs. 15-21 [legend
p. 198])
“At Station 2, near Bass’s Trail [South Bass Trail], about 200 feet below the rim of the
Grand Canyon [. . .] Cotypes [syntypes] are No. 103,236 A. N. S. P., collected by Ferriss and
Pilsbry, 1906.”
238 Oreohelix yavapai angelica, NEW SUBSPECIES (pp. 185-186, Pl. XII, figs. 22-25 [legend p.
198])
“On the Bright Angel Trail, at Grand Canyon, from 100 to 400 feet below the rim [. . .]
Cotypes [syntypes] are No. 103,239 A. N. S. P., collected by Ferriss and Pilsbry, 1906.”
239 Pupilla syngenes avus, NEW SUBSPECIES (p. 196, text-fig. 9)
Mollusca: Gastropoda: Stylommatophora: Pupillidae (chrysalis snails)
“Types [syntypes] No. 94,220 A. N. S. P., from upper slope of the Grand Canyon along the
Mystic Spring or Bass Trail [South Bass Trail], about 200 feet below the rim, Station 2
. . . .”
________________________________________________________________________________
88
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Pogue, Michael G.
2004
A new species of Schinia Hübner from riparian habitats in the Grand Canyon
(Lepidoptera: Noctuidae: Heliothinae). Zootaxa, (788): 1-4.
240 Schinia immaculata, NEW SPECIES; illustrated
Arthropoda: Insecta: Lepidoptera: Noctuidae (owlet moths)
“Type material. HOLOTYPE:
♂, Arizona, Coconino Co., Colorado River, Grand Canyon, river mile
23.0 L, shore, 36.6003° N, 111.7668° W, May 2, 2003, R. J. Delph, genitalia slide USNM
47867, USNM ENT 00219964. The holotype is provisionally deposited at the USNM, Washington,
DC, pending mutual resolution and agreement with the National Park Service regarding specimen
deposition.”
“PARATYPES (3°): U.S.A.: ARIZONA: COCONINO CO. 1° same data as holotype except: USNM
ENT 00210120 (NAU); river mile 166.5 L, old high water, 36.2542 N, 112.8996 W, 14 Apr.
2003 (1°), R. J. Delph, USNM ENT 00219965 (USNM); river mile 202 R, new high water,
36.0526 N, 113.3489 W, 15 May 2001 (1°), J. Rundall, USNM ENT 00210119 (NAU). Paratypes
deposited in the National Museum of Natural History, Washington, DC (USNM) and Northern
Arizona University, Flagstaff, AZ (NAU).”
paratypes may be a typographical error for
[Note: The symbol ° as which appears in this list of
♂ (the female of the species being “unknown”); note
as well that it is omitted from the geographical coordinates and that these symbols are correctly
printed in the data for the holotype (above).]
________________________________________________________________________________
Poppius, B. [Bertil Robert]
1914
Übersicht der Pilophorus Arten nebst Beschreibung verwandter Gattungen (Hem. Het.).
Annales de Société Entomologique de Belgique, 58: 237-254. [Article is in German.]
[Volume 58, 1914-1919; signature 16 of this volume, which includes the two species
listed here, is dated 20 August 1914.]
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
[The localities for the type specimens, cited peculiarly, refer to Williams, Arizona, and Bright
Angel, Arizona (Grand Canyon village).]
241 Pilophorus americanus, NEW SPECIES (p. 243)
“Ver. Staaten von N. Amerika : Williams, Ar.!, 21.VII; Br’t Angel, Ar.!, 12.VII, H. S. Barber, 2
♂♂ (U. S. Nat. Mus.).”
[Syntypes.]
242 Pilophorus exiguus, NEW SPECIES (pp. 246-247)
“Vereinigte Staaten von N. Amerika : Br’t Angel, Ar.!, 10 VII, Barber, 1
♀, 2 ♂♂
(U. S. Nat.
Mus., Mus. Helsingf.)” [Syntypes.]
________________________________________________________________________________
89
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Raven, Peter H.
1962
The systematics of Oenothera subgenus Chylismia. University of California Publications in
Botany, 34(1): 1-122.
Trachyophyta: Magnoliopsida: Myrtales: Onagraceae (evening primroses)
243 Oenothera confertiflora, NEW SPECIES (pp. 80-81); not illustrated
“Type.—In cinder soil, on east side, base of Vulcan’s Throne, Toroweap Valley, at about 4,200 feet
elevation, Grand Canyon National Monument [now in the enlarged Grand Canyon National Park],
Mohave County, Arizona, April 30, 1952, E. McClintock 52-294 (CAS 373,481; isotypes, ARIZ, NY,
WS).” “Known only from the type locality.”
244 Oenothera specuicola, NEW SPECIES (p. 87); not illustrated
Oenothera specuicola subsp. specuicola, NEW SUBSPECIES (p. 87); not illustrated
“Type [holotype].—Debris slides and crevices of Redwall limestone, Kaibab Trail [South Kaibab
Trail] at 5,300 feet elevation, south rim of Grand Canyon, Coconino County, Arizona, June 8,
1958, P. H. Raven 13119 (RSA; isotypes to be distributed).”
244a
Oenothera specuicola subsp. hesperia,
NEW SUBSPECIES
(p. 88); not illustrated
“Type [holotype].—Near Mooney Falls, Havasu Canyon, Coconino County, Arizona, May 23,
1950, J. T. Howell 26486 (CAS 359,774); isotype, ARIZ).” [Locality is in the Havasupai Indian
Reservation.]
[No. 244a added late to this checklist.]
________________________________________________________________________________
Reeder, John R.
1991
A new species of Panicum (Gramineae) from Arizona. Phytologia, 71(4) (October): 300303.
245
Panicum mohavense,
NEW SPECIES ;
illustrated
Trachyophyta: Magnoliopsida: Poales: Poaceae (grasses)
“TYPE: UNITED STATES. Arizona: Mohave Co., Arizona Strip: Main Street Valley, ca. 1.5 km S
of the jct. of the Colorado City road with the Main Street Valley road. Low hills with
limestone terraces, ca. 1525 m, 11 Oct 1990, J.R. & C.G. Reeder 8630 (HOLOTYPE: ARIZ;
Isotypes: US,K,MO).
________________________________________________________________________________
90
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Rehder, Alfred
1917
The genus Fraxinus in New Mexico and Arizona. Proceedings of the American Academy of
Arts and Sciences, 53(2) (October): 199-212.
Trachyophyta: Magnoliopsida: Lamiales: Oleaceae (olives)
246 Fraxinus cuspidata var. macropetala, NEW VARIETY (pp. 201-202); not illustrated
[Syntypes include:] “Arizona. Coconino Co.: Grand Canyon, July 9, 1892, E. O. Wooton (U. S.
Nat. Herb. No. 738265), July 13, 1892, J. W. Tourney (No. 273), June 1905, C. A. Purpus, June
13, 1913, A. E. Hitchcock (Nos. 76, 78); Red Canyon Trail [Hance Trail], June 10, 1901, L. F.
Ward (U. S. Nat. Herb. No. 410119); Bright Angel Trail, alt. 1400-2200 m., 1909, E. W. Nelson
(No. 108), May 31 and August 19, 1913, E. A. Goldman (Nos. 2067, 2223), alt. 1400-2000 m.,
July 19, 1914, A. Rehder (No. 106); Hermit Trail, June 18, 1916, Alice Eastwood (No. 5822);
Grand View Trail, June 16, 1916, Alice Eastwood (No. 5693).”
247 Fraxinus lowellii Sargent, NEW SPECIES (p. 211); not illustrated
(Type [holotype] from Oak Creek Canyon, Arizona.) [Paratype material includes:] “Mohave Co.:
Peach Springs, 1884, J. G. Lemmon (No. 3242).”
[This species description in Rehder is separately signed by C[harles]. S[prague]. Sargent, thus
Fraxinus lowellii Sargent in Rehder, 1917.] [J. G. Lemmon, with his wife, spent a month in the
Peach Springs area in May 1884. For a general (non-scientific) account of their visit, see J. G.
Lemmon, 1888, Grand Cañon of the Colorado, Overland Monthly, new series, 12(69) (September):
244-256.]
________________________________________________________________________________
Rehn, James A. G., and Grant, Harold J., Jr.
1958
A revision of the genus Morsea (Orthoptera; Acridoidea; Eumastacidae). Transactions of
the American Entomological Society, 84(3/4) (September/December): 217-259.
248
Morsea california kaibabensis,
NEW SUBSPECIES ;
(pp. 250-254, text-figs. 23-26)
Arthropoda: Insecta: Orthoptera: Eumastacidae (monkey grasshoppers)
“Type [holotype].—
♂;
Northwest escarpment slope of Kaibab Plateau, Coconino County,
Arizona. Elevation, 6700 feet. September 3, 1926. (Rehn and Hebard.) [Academy of Natural
Sciences of Philadelphia, Type no. 5810].” [square brackets are part of quotation].
________________________________________________________________________________
91
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Reinhard, H. J.
1953
New muscoid Diptera from the western United States. Pan-Pacific Entomologist, 29(1)
(January): 49-59.
249
Fabriciella evanida,
NEW SPECIES
(pp. 53-54); not illustrated
Arthropoda: Insecta: Diptera: Tachinidae (tachinid flies)
(Holotype and allotype from Lake City, Colorado.) Paratypes include: “1
♂, S. Grand Canyon,
Ariz., August 17, 1949 (P. R. Fitzgerald).”
________________________________________________________________________________
Reveal, James L.
1974a
Two shrubby novelties in Eriogonum (Polygonaceae) from the deserts of Utah and
Arizona. Brittonia, 26 (January/March): 90-94.
250
Eriogonum mortonianum,
NEW SPECIES
(pp. 91-92, fig. 1)
Trachyophyta: Magnoliopsida: Caryophyllales: Polygonaceae (knotweeds)
“Type: UNITED STATES: ARIZONA: Mohave Co.: along Arizona Hwy. 389, 4.5 mi SW of U.S.
Hwy. 89A at Fredonia (Coconino Co.) and ca. 9 mi ENE of Pipe Springs [Pipe Spring], on red
clay hills, associated with Eriogonum corymbosumn and Atriplex, at 4650 ft (1420 m), 16 Aug
1972, Reveal 2904 (HOLOTYPE: US!; ISOTYPES: ARIZ, ASU, BRY, CAS, GH, K, MO, NY, OKL, RM,
RSA, SMU, TEX, UC, UTC, WTU, and elsewhere).
_______________________________________________________________
1974b
Two new varieties of Eriogonum (Polygonaceae) from the Intermountain Region. Great
Basin Naturalist, 34(3): 245-246.
251
Eriogonum thompsonae S. Wats. var. atwoodii,
NEW VARIETY
(pp. 245-246); not illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Polygonaceae (knotweeds)
“Typus: Arizona: Mohave Co.: Along Arizona Highway 389, 4.3 miles west of the junction
of U.S. Highway 89A at Fredonia, on rolling reddish clay hills, associated with Atriplex, Ephedra,
Stanleya, and Eriogonum mortonianum, at about 4700 feet elevation, 15 August 1973, Reveal &
Reveal 5211. Holotypus, US! Isotypi, 15 duplicates to be distributed from US.”
________________________________________________________________________________
Rhodes, S. L., and Ayers, T. J.
2010
Two new taxa of Scutellaria Section Resinosa (Lamiaceae) from northern Arizona.
Journal of the Botanical Research Institute of Texas, 4(1): 19-26.
252
Scutellaria potosina var. kaibabensis,
NEW VARIETY
(pp. 20, 24, figs. 2A-E, 3A-C)
Trachyophyta: Magnoliopsida: Lamiales: Lamiaceae (mints)
“Type: U.S.A. Arizona: Coconino Co.: East rim of the Kaibab Plateau, North Canyon Trail #4,
36°25′04″N 112°04′17″W (NAD 27), 2133 m (7000 ft), 9 Jul 1998, Suzanne Rhodes 9811 with
92
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Tina Ayes (holotype ASC; isotypes: ARIZ, ASU).”
“Paratype: U.S.A.
Arizona. Coconino Co.:
Kaibab Plateau, East Rim View Point along Trail #7 ca. 2¼ mi down from trailhead, in
North Canyon; 36°25′18″N, 112°04′22″W, 2133 m (7000 ft); 1 Jul 1993, W. Hodgson 7334
(DES, ASU).”
________________________________________________________________________________
Ribble, D. W.
1974
A revision of the bees of the genus Andrena of the western hemisphere; subgenus
Scaphandrena. Transactions of the American Entomological Society, 100(2) (June): 101189.
253 Andrena (Scaphandrena) kaibabensis, NEW SPECIES (pp. 160-161); not illustrated
Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees)
“Type Material.—The holotype female (UCD) and 3 ♀♀ paratypes (UCS, INHS) were collected on
the North Rim of Grand Canyon National Park, June 21, 1949, by W. H. Lang. One female
paratype (AMNH) was collected 20 miles south of Jacob Lake, Arizona [in the Kaibab National
Forest], June 19, 1958, by W. J. and J. W. Gertsch.” (Also another paratype from Red Canyon,
Garfield Co., Utah.)
________________________________________________________________________________
Robinson, Benjamin Lincoln
1899
Three new Choripetalae from North America and Mexico. Botanical Gazette, 28(2)
(August): 134-136.
254 Silene rectiramea, NEW SPECIES (p. 134); not illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Caryophyllaceae (pinks)
“Collected by Professor D. T. MacDougal about the Grand Cañon of the Colorado in Arizona,
altitude 2150 m, 28 June, 1898, no. 181. Type in herb. Gray.”
________________________________________________________________________________
Robinson, Benjamin Lincoln, and Fernald, M. L.
1894
Contributions from the Gray Herbarium of Harvard University, New Series, No. VIII. New
plants collected by Messrs. C. V. Hartman and C. E. Lloyd upon an archæological
expedition to northwestern Mexico under the direction of Dr. Carl Lumholtz. Proceedings
of the American Academy of Arts and Sciences, 30: 114-123.
255 Crossosoma parviflora, NEW SPECIES (p. 114); not illustrated
Trachyophyta: Magnoliopsida: Crossosomatales: Crossosomataceae (crossomas)
[Syntypes:] “Collected in the Grand Cañon of the Colorado, by Dr. Gray, February to May,
1885, but referred to Glossopetalon Nevadense, Gray; and at La Tinaja, Sonora, at 3,700 feet, by
Mr. Hartman, 19 November, 1890 (no. 245).”
93
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
[In May 1885 Gray descended into Grand Canyon in Peach Springs Canyon to Diamond Creek and
the Colorado River.]
________________________________________________________________________________
Robinson, Harold
1967
New species of Dolichopodidae from the United States and Mexico (Diptera). Proceedings
of the Entomological Society of Washington (Washington, D.C.), 69(2) (June): 114-127.
256 Chrysotimus arizonicus, NEW SPECIES (p. 125, fig. 29 [p. 118])
Arthropoda: Insecta: Diptera: Dolichopodidae (longlegged flies)
“Holotype
♂, allotype ♀, and 1♀ paratype, from Grand Canyon National Park (north rim),
Arizona, July 15, 1954, W. L. Downes, Jr. Holotype (no. 69341) and allotype in USNM, paratype
in the Downes collection at the Department of Entomology, University of Illinois.”
________________________________________________________________________________
Rollins, Reed C.
1953
A new Sisymbrium from Arizona. Leaflets of Western Botany, 7(1): 15-16.
257 Sisymbrium kearneyi, NEW SPECIES; not illustrated
Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards)
“Type in the Gray Herbarium, collected along trail into S. B. Canyon, Grand Canyon National
Monument [now in the enlarged Grand Canyon National Park], Arizona, May 8, 1952, Elizabeth
McClintock 481a. Another collection (in flower) from the same locality but found growing under
shelter of overhanging cliffs is Elizabeth McClintock 52-481 (OH). Duplicates in Herb. Calif. Acad.
Sci.”
_______________________________________________________________
1982
Species of Draba, Lesquerella and Sibara (Cruciferae). Contributions from the Gray
Herbarium of Harvard University, (211) (January 15): 107-113.
258
Lesquerella kaibabensis Rollins,
NEW SPECIES
(pp. 110-111); not illustrated
Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards)
“Holotype in the Gray Herbarium, collected on a limestone-clay knoll, 18.6 miles south of Jacob
Lake on road to the north entrance of Grand Canyon National Park, Coconino County,
Arizona, June 8, 1979, Reed C. and Kathryn W. Rollins 79191. (Isotypes to be distributed.)”
________________________________________________________________________________
94
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Rydberg, Per Axel
1909
Studies on the Rocky Mountain flora—XX. Bulletin of the Torrey Botanical Club, 36(12)
(December): 675-698.
259 Penstemon coccinatus, NEW SPECIES (p. 691); not illustrated
Trachyophyta: Magnoliopsida: Lamiales: Plantaginaceae (plantains)
“Arizona: Grand Cañon of the Colorado, 1898, MacDougal 173 (type [holotype], in herb. N. Y.
Bot. Gard.) . . . .”
Paratype material includes: “Red Cañon Trail [Hance Trail], June 10, 1901,
L. F. Ward”. (Other paratypes from Oak Creek, Arizona, and localities in Utah.)
_______________________________________________________________
1913
Studies on the Rocky Mountain flora—XXIX. Bulletin of the Torrey Botanical Club, 40(9)
(September): 461-485.
260
Amsonia eastwoodiana,
NEW SPECIES
(p. 465)
Trachyophyta: Magnoliopsida: Gentianales: Apocynaceae (dogbane)
(Type [holotype] from Moab, Utah.) Paratype material includes “Lee’s Ferry, 1890, M. E. Jones”.
(Other paratype material from east of Holbrook, Arizona, and from Willow Creek Canyon, Utah.)
________________________________________________________________________________
Rydberg, Per Axel, and Pennell, Francis Whittier
1919
North American flora. Volume 24, Part 1. (Rosales) Fabaceae, Psoraleae. Per Axel
Rydberg. Eysenhardtia. Francis Whittier Pennell. New York: New York Botanical Garden,
pp. 1-64. [Pennell’s contribution on the genus Eysenhardtia restricted to pp. 34-40.]
Pediomelum Rydberg,
261
NEW GENUS
(p. 17)
Pediomelum retrorsum Rydberg,
NEW SPECIES
(p. 22); not illustrated
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
“Type [holotype] collected at Peach Spring [Peach Springs] near Big Bend of the Colorado,
Arizona, June, 1884, J. G. Lemmon & wife (herb. Columbia Univ.).”
[Lemmon, with his wife, spent a month in the Peach Springs area in May 1884. For a general
(non-scientific) account of their visit, see J. G. Lemmon, 1888, Grand Cañon of the Colorado,
Overland Monthly, new series, 12(69) (September): 244-256.]
________________________________________________________________________________
95
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Sargent, C. S. [Charles Sprague]
1919
Notes on North American trees. IV. Botanical Gazette, 67(3): 208-242.
262 Juniperus utahensis var. megalocarpa, NEW VARIETY (p. 208); not illustrated
Trachyophyta: Pinopsida: Pinales: Cupressaceae (cypress)
[Syntypes include:] “Arizona: rim of the Grand Canyon, C. S. Sargent, September 9, 1904;
Angel, near Flagstaff, Percival Lowell, September 4.”
[Inasmuch as there seems to be no
separate place name in Arizona by the simple name of “Angel”, Lowell’s gathering is presumably
from “Bright Angel” (Grand Canyon village).]
263 Populus fremontii var. toumeyi, NEW VARIETY (p. 214); not illustrated
Trachyophyta: Magnoliopsida: Malpighiales: Salicaceae (willows)
(Type [holotype] from Tucson, Arizona.)
Paratype material includes “Hermit Creek, Grand
Canyon of the Colorado, Coconino County, Alice Eastwood, April 10, 1917 (no. 6002)”.
________________________________________________________________________________
Scarbrough, Aubrey G.; Stevens, Lawrence E.; and Nelson, C. Riley
2012
The albibarbis-complex of Efferia Coquillett, 1910 from the Grand Canyon region,
southwestern U.S.A., with three new species and new distribution records (Diptera:
Asilidae). Pan-Pacific Entomologist, 88(1) (January): 58-86.
264 Efferia tapeats, NEW SPECIES (pp. 75-78, figs. 25-30)
Arthropoda: Insecta: Diptera: Asilidae (robber flies)
[In this entry all square brackets are part of the quotation, with awkward typography thus.
“GCNP” = Grand Canyon National Park. “HualIR/Peach Spr. Wash” = Hualapai Indian Reservation/
Peach Springs Wash.]
“Types. Arizona: Coconino Co., GCNP/Clear Cr., 4 km up [from confluence with Colorado
River]/12.ix.2005; 775 m/L.E. Stevens (holotype
♂, MNA).
Paratypes: same data as holotype
(1♀, MNA); GCNP/Colo. R. [Colorado River] Mi 108R [north bank] resting on Aster
[Chloracantha] spinosus flower; 20.ix.2009; 670 m/L.E. Stevens (1♂, MNA); GCNP/S. [South]
Bass Tr [Trail] in Supai Fmn. [Formation]/8.vii.2009; 1500 m/L.E. Stevens (1 ♀, MNA);
GCNP/Colo. R. Mi 108R/Shinumo Cr. Mouth/21.ix.2009; 665 m/L.E. Stevens (1 ♀, MNA);
GCNP/Colo. R. M. 108 L/12.vii.2007; 670 m/L.E. Stevens (1 ♂, MNA); Awatubi/Canyon (RM
58.2R) in Grand/Canyon Natl. Pk. 8.v.[19]92: M.E. Douglas, ASU (1 ♂, ASUT); GCNP/S. [South]
Bass Trail, base of Tapeats (Fmn) [formation]/12.vii.2007; 900 m/L.E. Stevens (1 ♀, MNA);
GCNP/Shinumo Cr. 3 km fr mouth/11.vii.2007; 810 m/L.E. Stevens (3♀, MNA); GCNP/S.
[South] Bass Trail in Redwall Fmn/12.vii.2007; 1200 m/L.E. Stevens (1 ♀, MNA); same data
except 13.vii.2007 (1♀, MNA); Mohave Co., HualIR/Peach Spr. Wash 4 km fr. CR [from
Colorado River]/08.x.2008; 380 m/L.E. Stevens (1 ♀, MNA).” (Also a paratype from San Juan
Spring, Glen Canyon National Recreation Area, Utah.)
________________________________________________________________________________
96
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Schell, Dorothydean Viets
1943
The Ochteridae (Hemiptera) of the western hemisphere. Journal of the Kansas
Entomological Society, 16(1) (January): 29-36, (2) (April): 37-47.
265 Ochterus barberi, NEW SPECIES (p. 32, Pl. 1, fig. 8 [p. 35]; pp. 41-42)
Arthropoda: Insecta: Hemiptera: Ochteridae (velvety shore bugs)
Species description and figure appear in the January issue; additional descriptive data and
itemization of type materials appear in the April issue.
“The holotype and allotype are deposited in the United States National Museum. The holotype, a
male, was taken at Colorado Canyon, Arizona; the allotype at Hot Springs, Arizona. Another
specimen in the National Museum bears the label, Orizaba, Mexico. Two specimens, paratypes,
were received from the American Museum of Natural History, collected at Grand Canyon,
Arizona. A series of 61 paratypes were collected at Indian Hot Springs and Castle Hot Springs,
Arizona by Doctor R. H. Beamer and his collecting party in August, 1941. They are now in the
Francis Huntington Snow collection.”
[No further itemized data regarding type material; data
quoted in their entirety here in order to embrace the disparate locality and collecting data as
published.]
________________________________________________________________________________
Schenk, John J.; Hodgson, Wendy; and Hufford, Larry
2010
A new species of Mentzelia section Bartonia (Loasaceae) from the Grand Canyon, Arizona.
Brittonia, 62(1): 1-6.
266 Mentzelia hualapaiensis, NEW SPECIES; illustrated
Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa)
“Type: U.S.A. Arizona: Mohave County, Grand Canyon National Park, along the Colorado River
near the Mile 158 camp site on river right, 339451 E, 4018170 N, NAD 83, 36°, 17′, 42.6″
N; 112°, 47′, 16.6″ W, 990 m, 19 Apr 2008, L. Makarick, D. Hall, and J. Cipra 2008-13 (holotype:
DES; isotype: GCNP).”
_______________________________________________________________
2013
Mentzelia canyonensis (Loasaceae), a new species endemic to the Grand Canyon,
Arizona, U.S.A. Brittonia, 65(4) (December): 408-416.
267 Mentzelia canyonensis, NEW SPECIES; illustrated
Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa)
“Type: U.S.A. Arizona. Coconino Co.: Grand Canyon National Park, Manzanita Canyon, within
¼ mile of Bright Angel Creek and North Kaibab Trail, UTM 12S 4005015 N, 407532 E,
4830 ft, 27 Sep. 2010, W. Hodgson 25718 (holotype: DES; isotypes: GCNP, NY, WS).”
________________________________________________________________________________
97
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Scudder, Samuel Hubbard
1878
Notice of the butterflies collected by Dr. Edward Palmer in the arid regions of southern
Utah and northern Arizona during the summer of 1877. U.S. Geological and Geographical
Survey of the Territories, Bulletin, 4: 253-258.
268 Neominois dionysus, NEW SPECIES (p. 254); not illustrated
Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals)
Syntype material comprises eight males and five females from “Juniper Mountains [extralimital to
this checklist] June 4; Mount Trumbull, June 7-10”.
269
Synchloe thoosa, NEW
SPECIES
(p. 257); not illustrated
Arthropoda: Insecta: Lepidoptera: Pieridae (orange-tips)
“A single female [holotype] was taken at Mokiak Pass, April 28-30 or June 2.”
_______________________________________________________________
1897
Revision of the orthopteran group Melanopli (Acridiidæ), with special reference to North
American forms. Proceedings of the U.S. National Museum, 20: 1-421, plates 1-26.
[Family Acrididae spelled Acridiidae in title.]
270 Melanoplus canonicus, NEW SPECIES (pp. 300-301, Pl. 20, fig. 1)
Arthropoda: Insecta: Orthoptera: Acrididae (short-horned grasshoppers)
“One male, 1 female. Grand Canyon of the Colorado, Arizona, July 10 (L. Bruner).”
________________________________________________________________________________
Shaw, F. R.
1951
Some new Mycetophilidae from the western United States. Bulletin of the Brooklyn
Entomological Society, 46(3) (June): 65-70.
271 Mycetophila denningi, NEW SPECIES (p. 67, Pl. 3, fig. 4)
Arthropoda: Insecta: Diptera: Mycetophilidae (fungus gnats)
“Described from one male [holotype] collected by D. G. Denning at Grand Canyon, Arizona on
June 18, 1949.”
________________________________________________________________________________
98
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Shear, William A.; Taylor, Steven J.; Wynne, J. Judson; and Krejca, Jean K.
2009
Cave millipeds of the United States. VIII. New genera and species of polydesmidan
millipeds from caves in the southwestern United States (Diplopoda, Polydesmida,
Macrosternodesmidae). Zootaxa, (2151): 47-65.
Pratherodesmus Shear,
272
NEW GENUS
Pratherodesmus voylesi Shear,
(p. 50)
NEW SPECIES
(pp. 51-54, figs. 2, 6, 9, 11–14, 19 [p. 56], 33
[p. 61])
Arthopoda: Diplopoda: Polydesmida: Macrosternodesmidae (presumed troglobitic polydesmidan
millipeds)
“Types: Male holotype, five male and seven female paratypes from Millipede Cave, Mojave [sic]
Co., Arizona, collected 16 February 2003 by K. Voyles, J. Jasper, M. Porter and K. Dittmar de la
Cruz, deposited in FMNH.
Four male and sixteen female paratypes from October Gyp Cave,
collected 27 March 2004 by same collectors. All specimens deposited in FMNH and CPMAB.”
________________________________________________________________________________
Sinclair, B. J.
2006
A new species of Wiedemannia Zetterstedt from Grand Canyon National Park, with notes
on additional Nearctic species (Diptera: Empididae). Journal of the Entomological Society
of Ontario, 137: 25-30.
273 Wiedemannia digna, NEW SPECIES (pp. 27-29, fig. 1)
Arthropoda: Insecta: Diptera: Empididae (balloon flies)
“Type Material. Holotype male labelled: ‘USA:AZ:Grand Canyon NP/ Vasey’s Paradise/
10.v.1998/ J.F. MacDonald’ (USNM). Paratype: Same data as holotype,
♂, USNM.”
________________________________________________________________________________
Small, John Kunkel
1906
Studies in North American Polygonaceae—II. Bulletin of the Torrey Botanical Club, 33(1)
(January): 51-57.
274
Eriogonum fusiforme,
NEW SPECIES
(p. 56); not illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Polygonaceae (knotweeds)
(Type [holotype] from Grand Junction, Colorado.) Paratype material includes “ARIZONA: Peach
Springs, June 28, I887, Tracy & Evans 315.”
________________________________________________________________________________
99
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Smiley, Robert L., and Moser, John C.
1974
New tarsonemids associated with bark beetles (Acarina: Tarsonemidae). Annals of the
Entomological Society of America, 67(4) (July): 639-665.
275 Heterotarsonemus nakaharai, NEW SPECIES (pp. 645-647, figs. 24, 25)
Arthropoda: Euchelicerata: Trombidiformes: Tarsonemidae (tarsonemid mites)
“Holotype.—Female, Canadian National Collection no. 13177, collected from Ips knausi Swaine in
Pinus ponderosa Douglas at Grand Canyon, Arizona.
The collector and date collected are
unknown.”
________________________________________________________________________________
Stacey, J. W.
1937
Notes on Carex—VIII. Leaflets of Western Botany, 2(1) (January): 13-16.
276 Carex curatorium [sic], NEW SPECIES (pp. 13-14); not illustrated.
Carex curatorum.
[The species was named for the collectors, Alice Eastwood, curator, and J. T. Howell, assistant
curator (Department of Botany, California Academy of Sciences); accordingly noticed thereafter
in literature with the Latin suffix of the specific epithet grammatically corrected, as Carex
curatorum Stacey.]
Trachyophyta: Magnoliopsida: Poales: Cyperaceae (sedges)
“Types: staminate plant, Herb. Calif. Acad. Sci., No. 204973, Eastwood and Howell No. 1101;
pistillate plant, Herb. Calif. Acad. Sci., No. 204974, Eastwood and Howell No. 1100, collected June
23, 1933, on Kaibab Trail to Roaring Springs [North Kaibab Trail], Grand Canyon National
Park, Arizona. Two other collections made with the type of the pistillate plant bear the numbers
1045 and 1089. Duplicates of some of these collections were distributed as C. pseudoscirpoidea
Rydb., an early determination reported in the literature (Leafl. West. Bot. 1: 142, 1934).”
________________________________________________________________________________
Stahnke, Herbert L.
1940
277
The scorpions of Arizona. Iowa State College Journal of Science, 15 (October): 101-103.
Vejovis [sic] aquilonalis,
NEW SPECIES
(p. 101); not illustrated. Vaejovis aquilonalis.
Arthropoda: Euchelicerata: Scorpiones: Vaejovidae (vaejovid scorpions)
“The specimen, a male, was taken 37 miles south of the Grand Canyon on highway 64.” [see
note below].
The type locality is probably a typographical error, as clarified by W. David Sissom and Oscar F.
Francke (1981), Scorpions of the genus Paruroctonus from New Mexico and Texas (Scorpiones,
Vaejovidae), Journal of Arachnology, 9(1) (Winter): 93-108, under their discussion of
Paruroctonus aquilonalis (Stahnke), p. 94: “Type data.—Holotype male from 30 miles south of
the Grand Canyon, Arizona, 8 August 1938 (Kay Anderson). Deposited in the collection of H. L.
Stahnke; examined. [. . .] The discrepancy in the type locality data seems to be a typographical
error, for the label with the holotype clearly reads ‘30 mi. South.’ ” [Note: There is no indication
100
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
that the label with the specimen is a field label; if written for the collection drawer, it could itself
be in error, obfuscating everything. —E.E.S.]
________________________________________________________________________________
Stallings, Don B., and Turner, J. R.
1957
Four new species of Megathymus (Lepidoptera, Rhopalocera, Megathymidae).
Entomological News (Academy of Natural Sciences of Philadelphia, Entomological
Section), 68(1) (January): 1-16.
278
Megathymus alliae,
NEW SPECIES
(pp. 1-5, Pl. 1)
Arthropoda: Insecta: Lepidoptera: Hesperiidae (skippers)
“Described from 62 specimens (35 males and 27 females) collected 15 miles west of Cameron,
Ariz., along the canyon of the Little Colorado River, elevation 5000 ft. All ex-larvae or expupae emerging from Aug. 25 to Oct. 5, during 1953, 1954 and 1955. Collected by Dr. and Mrs.
R. C. Turner, Dee, Jack, Don, and Viola Stallings.” [Quite probably the type locality is at or near
the Little Colorado River Gorge Overlook, along Arizona Route 64.]
“Holotype, female, Sept. 23, 1955, 15 miles west of Cameron, ARIZONA, el. 5000 ft. (Turner);
allotype, male, Sept. 17, 1954, 15 miles west of Cameron, Ariz., el. 5000 ft. (Turner) are in the
collection of the authors. Paratypes of both sexes will be placed in the following collections: H. A.
Freeman, C. L. Remington, U. S. Nat. Museum, American Mus. of Nat. History, Los Angeles County
Museum.”
________________________________________________________________________________
Standley, Paul C.
1909
The Allioniaceae of the United States with notes on Mexican species. Contributions from
the U.S. National Herbarium, 12(Part 8): 303-389, plates. (As part of the Bulletin of the
U.S. National Museum.)
279 Allionia pachyphylla, NEW SPECIES (p. 346); not illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Nyctaginaceae (four o’clocks)
“Type [holotype] U. S. National Herbarium no. 211717, collected in Arizona at the Grand Canyon,
1892, Toumey 485 [see note following]; cotype [isotype] in the herbarium of the University of
Arizona. Other specimens seen [paratypes]: Arizona: Red Canyon Trail [Hance Trail], Grand
Canyon, 1901, Ward; Grand Canyon, 1892, Wooton; Camp Verde, 1891, MacDougal.”
[Toumey 485 is very likely from the Old Hance Trail, which locality he visited in 1892; see notes
accompanying Coville (1894) in the present volume. Similarly, the Wooton gathering is likely
from the same locale, inasmuch as Wooton accompanied Toumey and company on the trip to the
Grand Canyon. For a general (non-scientific) account of the expedition, see C. H. Tyler Townsend
(1893), A wagon-trip to the Grand Cañon of the Colorado River, Appalachia, 7(1) (February): 4863. Note that the Ward gathering in 1901 is from the newer Hance Trail, which replaced the Old
Hance Trail that was destroyed by landslides in 1895.]
________________________________________________________________________________
101
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Stearns, Robert E. C.
1890
Scientific results of explorations by the U.S. Fish Commissions [sic] steamer Albatross.
No. XVII.—Descriptions of new west American land, fresh-water, and marine shells, with
notes and comments. Proceedings of the U.S. National Museum, 13: 205-225.
280 Helix (Arionta) coloradoensis, NEW SPECIES (p. 206, Pl. XV, figs. 6, 8, 12 [not 6-8; see
note below], legend p. 225)
Mollusca: Gastropoda: Stylommatophora: Helminthoglyptidae (shoulderbands)
“Grand Cañon of the Colorado, opposite the Kaibab plateau, at an elevation of 3,500 feet.
(Mus. No. 104100.)”
[The collection was made during the 1889 biological survey of C. Hart Merriam, thus the locality
is the Old Hance Trail; also note that the elevation indicates the collection was made along the
trail inside the canyon. See remarks with C. Hart Merriam in the present volume. The species
was later reassigned to the helminthoglyptid genus Sonorella.]
[Figures 7 and 12 of Pl. XV were transposed (fide H. A. Pilsbry, 1893, Manual of conchology :
structural and systematic.
Second series: Pulmonata.
Vol. VIII, Part 4, Helicidæ, Vol. VI.
Philadelphia: Academy of Natural Sciences of Philadelphia, Conchological Section, p. 226).]
________________________________________________________________________________
Stickel, William H.
1938
The snakes of the genus Sonora in the United States and Lower California. Copeia, 1938:
182-190.
281 Sonora semiannulata gloydi, NEW SUBSPECIES (pp. 186-187); not illustrated
Chordata: Reptilia: Squamata: Colubridae (typical snakes)
“Type [holotype].—MZUM 83754, collected by Mr. William Holzmark in 1936 on the Bright Angel
Trail, Lower Sonoran level of the Grand Canyon, Grand Canyon National Park, Arizona.
Paratypes.—Grand Canyon National Park, Arizona: Grand Canyon National Park collection 107,
vicinity of Indian Gardens [Havasupai Gardens]; UCLA 32, mouth of Garden Creek; MVZ
1758, Bright Angel Trail.”
________________________________________________________________________________
Stutz, Howard C.; Chu, Ge-Lin; and Sanderson, Stewart C.
1994
Atriplex asterocarpa (Chenopodiaceae), a new species from southern Utah and northern
Arizona. Madroño, 41(3): 199-204.
282 Atriplex asterocarpa, NEW SPECIES; illustrated
Trachyophyta: Magnoliopsida: Caryophyllales: Amaranthaceae (pigweed)
(Holotype from Garfield Co., Utah.) Paratypes include: “Arizona, Coconino Co., vic. Vermilion
Lodge [Vermilion Cliffs], steep rocky south slope, 4200 ft elevation, 18 Apr 1978, R. K. Gierish
102
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
4194 (ASC); Lee's Ferry, south-facing slope, Chinle Formation, 23 May 1993, H. C. Stutz 95792
(BRY).”
________________________________________________________________________________
Suksdorf, Wilhelm
1931
Untersuchungen in der Gattung Amsinckia. Werdenda (Beiträge zur Pflanzenkunde)
(Bingen, Washington), 1(5/8) (December 31): 47-113. [In German.]
Trachyophyta: Magnoliopsida: Boraginales: Boraginaceae (borage)
283 Amsinckia nana, NEW SPECIES (pp. 84-85); not illustrated
Based on a single specimen (holotype): “Ariz.; Hermit Creek, Grand Canyon of the Colorado
River, 10. April 1917, Alice Eastwood, nr. 6016 (C. A.)”
284 Amsinckia macrosepala, NEW SPECIES (p. 108); not illustrated
Syntypes include: “Grand Canyon, 1885, Gray (G. H.)”.
[Gray descended into Grand Canyon in Peach Springs Canyon to Diamond Creek and the Colorado
River.]
________________________________________________________________________________
Tanner, Wilmer W.
1953
A study of taxonomy and phylogeny of Lampropeltis pyromelana Cope. Great Basin
Naturalist, 13(1/2) (September 12): 47-66.
285 Lampropettis [sic] pyromelana Cope infralabials [sic], NEW SUBSPECIES (pp. 56-60, Pl.
2, fig. 1). (Common name given by the author, “Utah Ringed Snake”) Lampropeltis
pyromelana infralabialis.
Chordata: Reptilia: Squamata: Colubridae (typical snakes)
(Holotype from Beaver Co., Utah.) Paratypes include: “Arizona: Coconino County, Grand Canyon
Collection R3, R260-2, R291, R372, and UU 21 all from or near Bright Angel Point, North Rim
of Grand Canyon.”
[The misspelled generic and subspecific epithets are unfortunate and do not affect the taxonomy
of this snake, in no way forcing a new taxon into existence with these spellings. The genus
“Lampropettis” is simply a typographical error and is in fact spelled correctly (Lampropeltis) in the
title of the paper and elsewhere. The subspecies “infralabials” is spelled correctly (infralabialis) in
the key to subspecies (p. 60) and elsewhere throughout the paper.]
________________________________________________________________________________
103
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Theroux, Michael E.; Pinkava, D. J.; and Keil, D. J.
1977
A new species of Flaveria (Compositae: Flaveriinae) from Grand Canyon, Arizona.
Madroño, 24(1): 13-17.
286 Flaveria mcdougallii, NEW SPECIES; illustrated
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
“Type: United States, Arizona, Mohave Co., Grand Canyon National Park, Cove Canyon (174.2
mi below Lee’s Ferry), 27 Jan 1976, M. E. Theroux 1675 (Holotype: US; isotypes: ARIZ, ASC,
ASU, DES, GCNP, GH, MNA, NY, RSA, SRSC, TEX, UC).
Paratypes: United States, Arizona,
Coconino Co., Grand Canyon National Park, Matkatamiba Canyon (148.8 [sic, 147.8] mi
below Lee's Ferry), 1 Oct 1975, M. E. Theroux 1519 (MNA); Mohave Co., Grand Canyon National
Park, Cove Canyon, 5 Oct 1975, M. E. Theroux 1567 (MNA).”
________________________________________________________________________________
Thiers, Harry D.
1976
Boletes of the southwestern United States. Mycotaxon, 3(2) (January/March): 261-273.
[In this paper the type information is provided only within the portion written in botanical Latin.
No English equivalent was published.]
287
Boletus barrowsii Thiers & Smith ex Thiers,
NEW SPECIES
(pp. 262-264); not illustrated.
[Smith = Alexander H. Smith.]
Basidiomycota (club fungi): Agaricomycetes: Boletales: Boletaceae
“Holotypus (Thiers 27816) lectus prope Jacob Lake, Kaibab National Forest, Coconino County,
Arizona, Aug. 21, 1971; in herbario San Francisco State University (SFSU) conservatus.”
288
Suillus kaibabensis Thiers,
NEW SPECIES
(pp. 266-268); not illustrated
Basidiomycota (club fungi): Agaricomycetes: Boletales: Suillaceae
“Holotypus (Thiers 27813) lectus prope Jacob Lake, Kaibab National Forest, Coconino County,
Arizona, Aug. 21, 1971; in herbario San Francisco State University (SFSU) conservatus.”
289 Suillus occidentalis Thiers, NEW SPECIES (pp. 268-270); not illustrated
Basidiomycota (club fungi): Agaricomycetes: Boletales: Suillaceae
“Holotypus (Thiers 27775) lectus prope Point Sublime, North Rim, Grand Canyon National Park,
Coconino County, Arizona, Aug. 19, 1971; in herbario San Francisco State University (SFSU)
conservatus.”
________________________________________________________________________________
104
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Timberlake, P. H. [Philip H.]
1928
Bees of the genus Perdita Smith in the American Museum of Natural History
(Hymenoptera). American Museum Novitates, (321), 13 pp.
290 Perdita wheeleri, new species (pp. 5-7); not illustrated
Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees)
“1
♂ (holotype) collected in May, 1905, in
Indian Gardens [Havasupai Gardens] in the Grand
Cañon, Arizona (W. M. Wheeler).”
_______________________________________________________________
1956
A revisional study of the bees of the genus Perdita F. Smith, with special reference to the
fauna of the Pacific coast (Hymenoptera, Apoidea); Part II. University of California
Publications in Entomology, 11(5): 247-350.
291
Perdita opacella,
NEW SPECIES
(p. 324)
Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees)
A single female, holotype, from Marble Canyon, near Lee’s Ferry, Arizona (collected by G. D.
Butler, June 5, 1953, on Stanleya.
_______________________________________________________________
1962
A revisional study of the bees of the genus Perdita F. Smith, with special reference to the
fauna of the Pacific coast (Hymenoptera, Apoidea); Part V. University of California
Publications in Entomology, 28(1), 123 pp. [including 13 plates].
Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees)
292 Perdita inornata, NEW SPECIES (pp. 49-50, figs. 787, 788 [p. 115], 872 [p. 122])
(Holotype, allotype, and paratypes from Inyo Co., California.)
“Additional paratypes [include]
ARIZONA, Coconino Co.: 4 females, 14 males, South Rim of the Grand Canyon, July 26, 1934
(H. E. and M. A. Evans); 1 male, Grand Canyon, July 28, 1949 (W. H. Lange).” “Types at present
in collection of the Citrus Experiment Station, Riverside; paratypes in collections of the University
of Kansas, the University of California at Berkeley and Davis, and Cornell University.”
_______________________________________________________________
1968
A revisional study of the bees of the genus Perdita F. Smith, with special reference to the
fauna of the Pacific coast (Hymenoptera, Apoidea); Part VII (including index to Parts I to
VII). University of California Publications in Entomology, (49), 196 pp.
293
Perdita depressa,
NEW SPECIES
(pp. 48-50, plate figs. 1217, 1218 [p. 167], 1312 [p. 175])
Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees)
(Holotype and allotype from Monument Valley, Utah.) Paratypes include: “1 female, Supai, 3,500
feet, Havasu Canyon, Grand Canyon National Park, Coconino Co., Arizona, Aug. 2, 1934 (F. C.
105
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Lutz).” “Types in collection of the University of California, the female paratype in collection of the
American Museum of Natural History.”
________________________________________________________________________________
Townsend, C. H. Tyler [Charles Haskins Tyler]
1915
New western and southwestern Muscoidea. Journal of the New York Entomological
Society, 23(4) (December): 216-234. [Item signed “Charles H. T. Townsend”.]
Microsciasma
NEW GENUS
(p. 234)
294 Microsciasma minuta, NEW SPECIES (p. 234); not illustrated
Arthropoda: Insecta: Diptera: Tachinidae (tachinid flies)
“One male, Grand Canyon, Arizona, near Colorado River on Hance Trail [Old Hance Trail],
about 3,500 ft., July 10, 1892 (Townsend).” “Holotype, No. 19579 U. S. N. M.”
[Regarding the 1892 expedition see a general (non-scientific) account by Townsend (1893), A
wagon-trip to the Grand Cañon of the Colorado River, Appalachia, 7(1) (February): 48-63.]
________________________________________________________________________________
Triplehorn, Charles A.
1975
A subgenus of Eleodes with three new cave-inhabiting species (Coleoptera:
Tenebrionidae). The Coleopterists Bulletin, 29(1): 39-43.
Caverneleodes,
NEW SUBGENUS
(p. 39)
295 Eleodes (Caverneleodes) leptoscelis, NEW SPECIES (pp. 42-43); not illustrated
Arthropoda: Insecta: Coleoptera: Staphylinidae (rove beetles)
“Types: holotype (female) and 2 paratypes: Arizona: Coconino County, Cave of Domes [Cave
of the Domes, Horseshoe Mesa], 16-X-1953; 1 paratype, same data except 14-X-1954; 2
paratypes, Coconino County, Cave 68-Olje [Tse’an Olje, Cremation Canyon], 14-X-1954; 1
paratype, Coconino County, Tse-an-cho [Tse’an Cho, Horseshoe Mesa], 7-XI-1953. Holotype
(USNM #73090) and paratypes in United States National Museum; paratypes in The Ohio State
University Collection of Insects and Spiders.
The following information on cave locations was
supplied by Paul S. Bartholomew in 1957: Cave of Domes and Tse-an-Cho are both caves on
Horseshoe Mesa, about 12 miles east of South Village, which is on the south rim of the Grand
Canyon; Cave 68-Olje (also called Tse-an-Olje) is a cave east of South Village [Grand Canyon
village]; the 3 caves are at approximately 4500 feet elevation (T. J. Spilman, in litt.). ”
________________________________________________________________________________
106
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Triplehorn, Charles A., and Brown, Kirby W.
1971
A synopsis of the species of Asidina in the United States with description of a new species
from Arizona (Coleoptera: Tenebrionidae). The Coleopterists Bulletin, 25(3): 73-86.
296 Asidina rugicollis, NEW SPECIES (pp. 76-77, figs. 1, 7)
Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles)
“Holotype, male . . . Allotype, female . . . U. S. A., ARIZONA, COCONINO Co., Grand Canyon
National Park, South Rim, 19-VII-68, W. E., C. A., and B. W. Triplehorn [Ohio State University
Collection of Insects and Spiders].
Paratypes [include]: 33 specimens, all from ARIZONA:
Coconino Co.: same data as holotype, 3 males, 2 females [OSUC]; 3 males, 3 females [KWBC];
1 male, 1 female [USNM]; Grand Canyon, 26-27-VII-26, E. C. Van Dyke, 3 males, 5 females,
[CASC]; 1 male, 1 female [ASUT]; 1 male, 1 female [BMNH]; 1 male, 1 female [MHNP] . . .
northwest slope of Kaibab Plateau, 3-IX-26, 6700′, R. & H., 1 female [ANSP] . . . .” [Square
brackets are part of the quotation.]
________________________________________________________________________________
Troubridge, J. T., and Crabo, L. G.
2002
A review of the Nearctic species of Hadena Schrank, 1802 (Lepidoptera: Noctuidae) with
descriptions of six new species. Fabreries (Association des Entomologistes Amateurs du
Québec), 27(2) (December): 109-154 (“Section en français”, pp. 135-145). [Article is
principally in English, with an abbreviated section (not an abstract) in French.]
297
Hadena (Hadena) lafontainei,
NEW SPECIES
(pp. 114-117, figs. 3 [p. 146], 17 [p. 148], 33
[p. 153])
Arthropoda: Insecta: Lepidoptera: Noctuidae (owlet moths)
(Holotype from Albany Co., Wyoming.) Paratype material of 85 specimens includes “2 ♀♀, Kaibab
Plateau, nr. Jacob Lk., 8,300 ft [2,550 m], 18.VII.1980, D.C. Ferguson” (square brackets are
part of quotation). [“nr. Jacob Lk.” = near Jacob Lake.]
________________________________________________________________________________
Tuthill, L. D.
1939
New species of Psyllidae from the western United States. Iowa State College Journal of
Science, 13(2) (January): 181-186.
298 Trioza rubra, NEW SPECIES (pp. 185-186); not illustrated
Arthropoda: Insecta: Hemiptera: Psyllidae (jumping plantlice)
(Holotype female, allotype male, and nine paratypes from Creede, Colorado.) Additional paratypes
include: “one male, Grand Canyon, Arizona, Aug. 11, 1927, R. H. Beamer”. Holotype and
allotype in author’s collection; paratypes in author’s collecton and in Snow Collection, University
of Kansas, and U.S. National Museum (not specified).
________________________________________________________________________________
107
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Usinger, Robert L.
1931
A new species of Platylygus (Miridae, Hemiptera). Pan-Pacific Entomologist, 7(3)
(January): 129-130.
299 Platylygus vanduzeei, NEW SPECIES; text-fig.
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
“Holotype, male. No. 2997, Mus. Calif. Acad. Sciences, collected by C. D. Duncan, June 17, 1921,
at the Grand Canyon, Arizona, and allotype, female. No. 2998, Mus. Calif. Acad. Sciences,
collected by the author in Grand Canyon National Park, Arizona, South Rim, June 29, 1930.”
(Also 12 paratype females from Mesa Verde National Park, Colorado.)
________________________________________________________________________________
Van Duzee, M. C.
1927
North American species of Polymedon. (Diptera Dolichopodidæ). Annals of the
Entomological Society of America, 20(1) (March): 123-126.
Arthropoda: Insecta: Diptera: Dolichopodidae (longlegged flies)
300 Polymedon dilaticosta, NEW SPECIES (pp. 124-125); not illustrated
“Described from nine males and three females, taken on the Bright Angel Trail, Grand Canon,
Arizona, and one female taken in Oak Creek Canon, Arizona, August, at an elevation of 6,000
feet; the last one was taken by F. H. Snow. Type and allotype in the Canadian National Collection.”
[Distinguishing data for primary types not indicated, thus all specimens are originally syntypes.]
301 Polymedon nitidus, NEW SPECIES (p. 125); not illustrated
“Described from one female [holotype], taken on the Bright Angel Trail, Grand Canon, Arizona.
Type in the Canadian National Collection.”
________________________________________________________________________________
Wang, Daqing, and Holsinger, John R.
2001
Systematics of the subterranean amphipod genus Stygobromus (Crangonyctidae) in
western North America, with emphasis on species of the hubbsi group. Amphipacifica
(Journal of Aquatic Systematic Biology), 3(2) (November 15): 39-147.
302 Stygobromus blinni, new species (pp. 74-76, fig. 21)
Arthropoda: Malacostraca: Amphipoda: Crangonyctidae (crangonyctid amphipods)
Female holotype and two female paratypes from “Roaring Springs Cave, on the north rim of the
Grand Canyon near Bright Angel Trail and Bright Angel Creek” [sic] (p. 74), “about 1.6 km inside
the cave” (p. 76); collected by Dean Blinn, 28 September 1994.]
________________________________________________________________________________
108
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Watson, Sereno
1873
New plants of northern Arizona and the region adjacent. American Naturalist, 7(5)
(May): 299-303. [Also published as offprint, with header, “Contributions to American
Botany. I.”, “Printed at the Salem Press. May, 1873.”, 7 pp.]
“In the botanical collections made in 1871-’72, mainly in the southern portion of the Great Basin,
in northwestern Arizona and the adjacent desert section of California, by Mr. Ferd. Bischoff and
others, under the direction of Lieut. G. M. Wheeler, Corps of Engineers, in the course of his
exploration of that region, several new species have been found which are here described, by
consent, in anticipation of the fuller report now in preparation. With these are given some others
occurring in a small collection made by Mrs. Ellen P. Thompson near Kanab, Southern Utah, during
the last summer while accompanying her brother, Maj. [John Wesley] Powell, in his survey of the
Colorado.” [Below are listed only those specimens gathered by Thompson in “Northern Arizona”,
inasmuch as these were clearly obtained while on relatively short journeys into the Arizona Strip
from her base in Kanab, Utah.]
303
Dalea amoena,
NEW SPECIES
(p. 300); not illustrated
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
“Northern Arizona (Mrs. E. P. Thompson). In damp places; April.”
304
Oenothera (Chylisma) multijuga,
NEW SPECIES
(p. 300); not illustrated
Trachyophyta: Magnoliopsida: Myrtales: Onagraceae (evening primroses)
“Northern Arizona (Mrs. E. P. Thompson). In dry places; March.”
305
Cymopterus purpureus,
NEW SPECIES
(pp. 300-301); not illustrated
Trachyophyta: Magnoliopsida: Brassicales: Apiaceae (parsleys)
“Northern Arizona (Mrs. E. P. Thompson). In damp, shaded soil; March.
306
Peucedanum newberryi,
NEW SPECIES
(p. 301); not illustrated
Trachyophyta: Magnoliopsida: Brassicales: Apiaceae (parsleys)
“Northern Arizona, on stony soil (Mrs. E. P. Thompson). April.”
307
Brickellia (Clavigera) longifolia,
NEW SPECIES
(p. 301); not illustrated
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
“Northern Arizona (Mrs. E. P. Thompson). In a damp cañon; April.”
308
Calochortus flexuosus,
NEW SPECIES
(p. 303); not illustrated
Trachyophyta: Magnoliopsida: Liliales: Liliaceae (lilies)
“Southern Utah and Northern Arizona (Mrs. E. P. Thompson); April and May. The bulbs, as of
other species, are eaten by the Indians.”
109
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
309
Androstephium breviflorum,
NEW SPECIES
(p. 303); not illustrated
Trachyophyta: Magnoliopsida: Asparagales: Asparagaceae (agaves)
“Southern Utah and Northern Arizona (Mrs. E. P. Thompson); April and May. Bulbs also eaten.”
_______________________________________________________________
1888
Contributions to American botany. Proceedings of the American Academy of Arts and
Sciences, 23(2): 249-287.
310
Lesquerella arizonica,
NEW SPECIES
(p. 254); not illustrated
Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards)
Syntypic material includes “Arizona . . . at Peach Springs (4177 Lemmon, 64 Jones), and at
Mokiak Pass (43 Palmer, 1877).”
[J. G. Lemmon, with his wife, spent a month in the Peach Springs area in May 1884. For a general
(non-scientific) account of their visit, see J. G. Lemmon, 1888, Grand Cañon of the Colorado,
Overland Monthly, new series, 12(69) (September): 244-256.]
________________________________________________________________________________
Welsh, Stanley L., and Atwood, N. Duane
2001
New taxa and nomenclatural proposals in miscellaneous families—Utah and Arizona.
Rhodora (Journal of the New England Botanical Club), 103: 71-95.
311
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis,
NEW VARIETY
(pp. 81-82, fig.
3 [p. 83])
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
[Localities are within the present boundaries of Grand Canyon-Parashant National Monument.]
“Type: U.S.A. Arizona: Mohave Co., Mohave Strip, Andrus Canyon, 3 mi. W of Andrus Pt.,
T32N R10W S6, wash bottom with Mortonia, Purshia, Ephedra, Agave, etc., 26 Apr 1999, N.
D. Atwood & B. Furniss 24293 (HOLOTYPE: BRY; six isotypes to be distributed).”
“Additional specimens (paratypes; all BRY!): Arizona: Mohave Co., Parashant (Trail) Canyon,
11 mi. S of Mt. Trumbull Village, 26 Apr 1974, Atwood 6029, 6029a; ditto, Whitmore Canyon,
1.25 km (7.8 mi.) [sic] S of Mount Trumbull (abandoned town), T34N R9W S29, 25 May
1979, Holmgren & Barneby 9172; Daneill Canyon W of Andrus Canyon, 3 mi. W of Andrus
Pt., T32N R11W S12, 26 Apr 1999, Atwood & Furniss 24300; ditto, Andrus Canyon, 1 mi. W
of Andrus Pt„ T32N R10W S10, 26 Apr 1999, Atwood & Furniss 24302; ditto, Whitmore
Canyon, T32N R9W S1, 27 May 1999, Higgins 20226; ditto, 2 mi. S of Mt. Trumbull school
house, near Griffith Knoll, T34S R10W S1-2, 27 May 1999, Higgins 20277.”
________________________________________________________________________________
110
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Welsh, Stanley L., and Licher, Max
2010
Pediomelum Rydberg (Leguminosae) in Arizona and two previously undescribed species.
Western North American Naturalist, 70(1): 9-18.
312
Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood,
NEW SPECIES
(pp. 14-15,
fig. 3). (Common name given by the author, “Poverty Mountain Breadroot”)
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
“Type.—USA, Arizona, Mohave Co., T35N, R12W, Sec. 35/36, SW of Poverty Mountain, near
Dewdrop Spring; 1756 m, in gravelly lime- stone soil in pinyon-juniper community, L.C. Higgins
23135, 25 May 2001, holotype BRY, isotypes distributed previously as P. mephiticum.”
“Paratypes.—Arizona, Mohave Co., T34N, R11W, Sec 4, SE of Poverty Mountain, Arizona
Strip, SW of Mt. Trumbull, 5500 ft, Atwood & Franklin 18901, 16 May 1993, BRY! — . . . — BLM
Road, S of Poverty Mountain and SW of Salt Spring, T34N, R11W [sic], 5000-5500 ft,
Salywon et al. 1107, 21 May 2001, BRY!”
[The abstract of this paper erroneously indicates P. pauperitense is from Coconino County,
Arizona.]
________________________________________________________________________________
Welsh, Stanley L., and Thorne, K. H.
1977
Plants of Arizona: a new species of Astragalus from the Kaibab Plateau. Great Basin
Naturalist, 37(1): 103-104.
313
Astragalus pinionis,
NEW SPECIES ;
illustrated
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
“Holotype: Arizona, Coconino County, 17 miles southeast of Fredonia Sawmill, along logging
road to Ryan, pinyon-juniper-cowania-big sagebrush community, N. D. Atwood 6794, 15 June
1976 (BRY, isotypes to be distributed).”
________________________________________________________________________________
Wheeler, William Morton
1899
New species of Dolichopodidæ from the United States. Proceedings of the California
Academy of Sciences, Series 3, Zoology, 2(1), 85 pp. [including 4 plates].
314 Polymedon castus, NEW SPECIES, pp. 6-8, Pl. 1, fig. 8.
Arthropoda: Insecta: Diptera: Dolichopodidae (longlegged flies)
“One specimen labelled ‘Grand Cañon, Arizona,’ from the collections of the University of
Kansas.” Specimen (holotype) is a female.
_______________________________________________________________
111
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1906
The ants of the Grand Cañon. Bulletin of the American Museum of Natural History, 22:
329-345.
Arthropoda: Insecta: Hymenoptera: Formicidae (ants)
Pheidole desertorum,
NEW SPECIES
(pp. 337-339). [Type material from Fort Davis, Texas.]
315 Pheidole desertorum var. maricopa, NEW VARIETY (pp. 339-340); not illustrated
“Described from several soldiers and workers taken from a single colony under a stone in the
Grand Cañon just above the Indian Garden [Havasupai Gardens] (3876 feet.)” [Syntypes?]
316 Formica moki, NEW SPECIES (pp. 343-344); not illustrated
“Described from 25 workers taken from a small colony under a stone on the wall of the Grand
Cañon at an altitude of about 5500 feet and 2 workers taken on the rim in the Kohonino
[sic] Forest (7000 feet) about three miles west of the Bright Angel Hotel.” [The “wall of
the Grand Cañon” locale is in all probability on Bright Angel Trail.] [Syntypes?]
(Note: Wheeler, 1913, p. 343 [see citation below], placed the species Formica moki in
Neoformica,
NEW SUBGENUS
[pp. 400-401].)
_______________________________________________________________
1913
A revision of the ants of the genus Formica (Linné) Mayr. Bulletin of the Museum of
Comparative Zoology, 53(10): 379-565.
317 Formica fusca fusca var. gelida, NEW VARIETY (pp. 505-507); not illustrated.
[Not valid as
a quadrinomial taxon.]
Arthropoda: Insecta: Hymenoptera: Formicidae (ants)
(“Type locality.—[State of] Colorado: Ward, 9,000 ft. (T. D. A. Cockerell).”)
Other material
(= paratypes) from numerous, widespread continental localities, including: “Arizona: Coconino
Forest, Grand Canyon, 7,000 ft. (Wheeler).”
_______________________________________________________________
1914
New and little known harvesting ants of the genus Pogonomyrmex. Psyche, (October):
149-157.
318 Pogonomyrmex californicus Buckley subsp. maricopa, NEW SUBSPECIES (pp. 155-156);
not illustrated
Arthropoda: Insecta: Hymenoptera: Formicidae (ants)
[Based on syntypes although with the indication, “New Mexico: Alamogordo, type locality (G. von
Krockow)”, without further correlation to specimens.] Other material noted from New Mexico,
Arizona, California, Texas, and Mexico, including from Arizona “Grand Canyon (Wheeler)”.
_______________________________________________________________
112
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
1915
Some additions to the North American ant-fauna. Bulletin of the American Museum of
Natural History, 34: 389-421.
319 Camponotus acutirostris Wheeler clarigaster, NEW VARIETY (p. 420); not illustrated
Arthropoda: Insecta: Hymenoptera: Formicidae (ants)
“A single specimen [holotype] taken at an altitude of about 3000 ft. on the Bright Angel Trail
in the Grand Cañon, Arizona.”
________________________________________________________________________________
Wickham, Henry Frederick
1899
The habits of American Cicindelidae. Proceedings of the Davenport Academy of Natural
Sciences, 7: 206-228.
320 Cicindela rufiventris var. arizonae, NEW VARIETY (pp. 226-227); not illustrated
Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles)
[Syntypes:] “Mr. Roland Hayward sent me specimens labelled ‘Grand Cañon of the Colorado’
and I have since received the same thing collected by Professor Townsend, who writes
[underscored for emphasis in this checklist]: ‘I collected the species in the Grand Cañon during
the first two weeks of July, 1892. It occurred on sandy places or along paths by a stream going
down a side cañon, from 2500 feet above, down to the level of the Colorado River. Or from 5000
feet above sea (at Hance's stone cabin, 2500 feet below the rim and about two or two and a half
miles down the trail from river) quite to the bottom of the cañon, which is here 2500 feet above
the sea. The insect is not especially wary.’ ”
[Regarding the 1892 expedition see a general (non-scientific) account by Townsend (1893), A
wagon-trip to the Grand Cañon of the Colorado River, Appalachia, 7(1) (February): 48-63. The
locality was on the Old Hance Trail, the upper portion of which was destroyed in 1895 by landslides
after which John Hance re-routed his trail down Red Canyon (the present Hance Trail).]
________________________________________________________________________________
Wilcox, J.
1965
New Heteropogon Loew, with a key to the species and the description of a new genus
(Diptera: Asilidae). Bulletin of the Southern California Academy of Sciences, 64(4): 207222.
321 Heteropogon stonei, NEW SPECIES (pp. 220-222); not illustrated
Arthropoda: Insecta: Diptera: Asilidae (robber flies)
(Holotype and allotype from Hualapai Mountains, Arizona.)
“Paratypes: . . . one
♀,
Grand
Canyon, Arizona, 4 June 1963 (J. Wilcox) . . . .”
________________________________________________________________________________
113
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
SERIALS LISTED IN THE
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Aliso (Rancho Santa Ana Botanic Garden, Claremont, California)
American Journal of Botany
American Midland Naturalist
American Museum Novitates (American Museum of Natural History)
American Naturalist
Amphipacifica (Journal of Aquatic Systematic Biology)
Annales de Société Entomologique de Belgique
Annals of the Carnegie Museum of Natural History
Annals of the Entomological Society of America
Annals of the New York Academy of Sciences
Arquivos do Museu Boçage (Lisboa)
Beihefte zur Sydowia (Horn, Niederösterreich, Austria)
Botanical Gazette
Brigham Young University Science Bulletin (Biological Series)
British Journal of Entomology and Natural History
Brittonia
Bulletin of the American Museum of Natural History
Bulletin of the Brooklyn Entomological Society
Bulletin of the California Lichen Society
Bulletin of the Museum of Comparative Zoology
Bulletin of the Southern California Academy of Sciences
Bulletin of the Torrey Botanical Club
Bulletin of the University of Nebraska State Museum
Bulletin of the University of Utah
Cactus and Succulent Journal (Cactus and Succulent Society of America)
The Canadian Entomologist (London)
Cicindela
The Coleopterists Bulletin
Contributions from the Gray Herbarium of Harvard University
Contributions from the U.S. National Herbarium
Contributions to Western Botany
Copeia
114
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Entomologica Americana
Entomological News (Academy of Natural Sciences of Philadelphia, Entomological Section)
Fabreries (Association des Entomologistes Amateurs du Québec)
Florida Entomologist
Garden and Forest
Great Basin Naturalist
Iowa State College Journal of Science
Journal of Arachnology
Journal of Eukaryotic Microbiology
Journal of Mammalogy
Journal of Parasitology
Journal of Protozoology
Journal of the Arnold Arboretum
Journal of the Botanical Research Institute of Texas
Journal of the Elisha Mitchell Scientific Society
Journal of the Entomological Society of Ontario
Journal of the Kansas Entomological Society
Journal of the Lepidopterists’ Society
Journal of the New York Entomological Society
Journal of the Washington Academy of Sciences (Washington, D.C.)
Kansas University Science Bulletin
Leaflets of Botanical Observation and Criticism
Leaflets of Western Botany
The Lepidopterists’ News
Madroño
Minnesota State Entomologist, Report to the Governor
Mycologia
Mycotaxon
The Nautilus (Philadelphia)
Notulae Naturae of the Academy of Natural Sciences of Philadelphi a
Novon
Ohio Journal of Science
Pan-Pacific Entomologist
PeerJ
Phytologia
Pittonia
Proceedings of the Academy of Natural Sciences of Philadelphia
Proceedings of the American Academy of Arts and Sciences
115
BIBLIOGRAPHY AND ANNOTATED CHECKLIST
Proceedings of the Biological Society of Washington (Washington, D.C.)
Proceedings of the California Academy of Sciences
Proceedings of the Davenport Academy of Natural Sciences
Proceedings of the Entomological Society of Washington (Washington, D.C.)
Proceedings of the Helminthological Society
Proceedings of the U.S. National Museum
Proceedings of the Utah Academy of Sciences
Proceedings of the Washington Academy of Sciences (Washington, D.C.)
Psyche
Rhodora (Journal of the New England Botanical Club)
Scientific Reports (Springer Nature)
Southwestern Naturalist
Sydowia
Systematic Botany
Systematic Botany Monographs
Transactions of the American Entomological Society
Transactions of the American Microscopical Society
Transactions of the Illinois State Academy of Science
Transactions of the San Diego Society of Natural History
University of California Publications in Botany
University of California Publications in Entomology
University of Kansas Science Bulletin
U.S. Department of Agriculture, North American Fauna
U.S. Forest Service Agriculture Handbook
U.S. Geological and Geographical Survey of the Territories, Bulletin
Werdenda (Beiträge zur Pflanzenkunde) (Bingen, Washington)
Western North American Naturalist
ZooKeys
Zootaxa
116
TAXONOMIC LIST
TAXONOMIC LIST
This is a nomenclatural record. Taxa are as named by the original authors; later taxonomic
reassignments, if any, are not generally noted herein.
There are 322 taxa first named from collections made in the greater Grand Canyon
region of northwestern Arizona, 215 of which have type material that is wholly or partly
from within, or with good proabability within, the present boundaries of Grand Canyon
National Park. [No. 244a was added late, thus the whole-number count stops at 321.]
Higher systematics for the original genus are given here, as derived from the Integrated
Taxonomic Information System (ITIS) online in 2022. Taxa not included in ITIS have had
their higher systematics derived from other sources, including the National Center for
Biotechnology Information Taxonomy Browser (National Library of Medicine) and the
World Flora Online. The systematic placements listed here may not reflect the most current
views, which are in any case bound to vary between researchers and over time. Again, this is
a resource manager’s historical checklist to nominate taxa, not one of systematics or revised
nomenclature.
The taxonomic ranks cited in this list are: Phylum or Division: Class: Order: Family.
The taxa in this list are not in so-called “systematic order” because some users of the
checklist will not be accustomed to the methodical order of taxa within Phyla, Classes, and
so forth. To accommodate these users, who are more likely to be resource managers and
historically inclined users who are not necessarily scientists, all higher taxa are listed
strictly in alphabetical order (the order of Kingdoms is arranged only for convenience
based on the relative number of entries). The nominate taxa within each category are listed
in the order in which they appear in the checklist, arranged numerically to facilitate finding
them in the bibliography/ checklist.
Common names are mostly as listed by ITIS or other sources. They should not be taken
to be names regularly adopted by all users, who will understand other common names for
these taxa. The names have been added here only a matter of convenience for those users,
principally non-scientists, who prefer or need to refer in some way to common names.
Arrow indicates taxa for which at least one locality reported for its types is within, or is
with good probability from within, the present boundaries of Grand Canyon National
Park. Refer to the bibliography for all details.
Red Numbers along the left margin of the Taxonomic List indicate and cross-list the
sequentially enumerated taxa in the bibliography (they reflect the alphabetical order by
publication author).
117
TAXONOMIC LIST
TOTAL NUMBERS OF NOMINATE TAXA
/ NO. FROM GRAND CANYON NATIONAL PARK
[In the PDF all lines below are hyperlinks to those categories in the Taxonomic List]
Kingdom Animalia (animals)
Phylum Arthropoda (arthropods)
Total
GRCA
Page
172
133
119
141
107
119
Class Branchiopoda (branchiopod crustaceans)
1
119
Class Diplopoda (millipedes)
1
119
Class Euchelicerata (spiders/mites/scorpions/etc.)
14
7
119
124
99
121
1
1
128
18
14
128
1
1
128
13
9
128
Class Reptilia (reptiles)
3
3
129
Class Teleostei (fishes)
1
1
130
Class Insecta (insects)
Class Malacostraca (malacostracan crustaceans)
Phylum Chordata (vertebrates)
Class Aves (birds)
Class Mammalia (mammals)
Phylum Mollusca (mollusks)
130
Class Gastropoda (snails)
9
7
130
4
5
131
111
49
132
110
48
132
1
1
137
27
22
138
23
20
138
4
2
139
Kingdom Chromista (photosynthetic eukaryotes)
11
11
140
Phylum Apicomplexa (parasitic alveolates)
11
11
140
Phylum Nematoda (roundworms)
Class Chromadorea (chromadorid roundworms)
Kingdom Plantae (plants)
Phylum (Division) Trachyophyta (vascular plants)
Class Magnoliopsida (flowering plants)
Class Pinopsida (conifers)
Kingdom Fungi (fungi)
Phylum (Division) Ascomycota (sac fungi)
Phylum (Division) Basidiomycota (club fungi)
Kingdom Protista (protists)
“Phylum” Hemimastigophora (single-celled eukaryotes)
118
1
140
1
140
TAXONOMIC LIST
KINGDOM ANIMALIA
(animals)
Phylum ARTHROPODA
(arthropods)
Class Branchiopoda
(branchiopod crustaceans)
Arthropoda: Branchiopoda: Anostraca: Branchinectidae (brine shrimp)
48
Branchinecta kaibabensis Belk & Fugate, 2000
________________________________________________________________________________
Class Diplopoda
(millipedes)
Arthopoda: Diplopoda: Polydesmida: Macrosternodesmidae (presumed troglobitic
polydesmidan millipeds)
272
Pratherodesmus voylesi Shear in Shear, Taylor, Wynne & Krejca, 2009
________________________________________________________________________________
Class Euchelicerata
(spiders/mites/scorpions/etc.)
Arthropoda: Euchelicerata: Areneae: Hahniidae (hahniid spiders)
71
Neoantistea coconino Chamberlin & Ivie, 1942
________________________________________________________________________________
Arthropoda: Euchelicerata: Areneae: Linyphiidae (dwarf weavers)
70
Tapinocyba kesimba Chamberlin, 1948
________________________________________________________________________________
Arthropoda: Euchelicerata: Areneae: Lycosidae (wolf spiders)
114
Arctosa mokiensis Gertsch, 1934a
________________________________________________________________________________
Arthropoda: Euchelicerata: Areneae: Salticidae (jumping spiders)
115
Phidippus kaibabensis Gertsch, 1934b
________________________________________________________________________________
119
TAXONOMIC LIST
Arthropoda: Euchelicerata: Areneae: Sicariidae (recluse spiders)
117
Loxosceles kaiba Gertsch & Ennik, 1983
________________________________________________________________________________
Arthropoda: Euchelicerata: Areneae: Theraphosidae (tarantulas)
68
69
Aphonopelma behlei Chamberlin, 1940
Aphonopelma phasmus Chamberlin, 1940
________________________________________________________________________________
Arthropoda: Euchelicerata: Areneae: Theridiidae (cobweb weavers)
116
Steatoda variata Gertsch, 1960
________________________________________________________________________________
Arthropoda: Euchelicerata: Areneae: Thomisidae (crab spiders)
113
Misumenops coloradensis Gertsch, 1933
________________________________________________________________________________
Arthropoda: Euchelicerata: Pseudoscorpiones (pseudoscorpions): Chernetidae
153
154
Hesperochernes bradybaughi Harvey & Wynne, 2014
Tuberochernes cohni Harvey & Wynne, 2014
________________________________________________________________________________
Arthropoda: Euchelicerata: Pseudoscorpiones (pseudoscorpions): Larcidae
225
Archeolarca cavicola Muchmore, 1981
________________________________________________________________________________
Arthropoda: Euchelicerata: Scorpiones: Vaejovidae (vaejovid scorpions)
277
Vaejovis aquilonalis Stahnke, 1940
________________________________________________________________________________
Arthropoda: Euchelicerata: Trombidiformes: Tarsonemidae (tarsonemid mites)
275
Heterotarsonemus nakaharai Smiley & Moser, 1974
________________________________________________________________________________
120
TAXONOMIC LIST
Class Insecta
(insects)
Arthropoda: Insecta: Blattodea: Polyphagidae (sand cockroaches)
166
167
168
Arenivaga grandiscanyonensis Hopkins, 2014
Arenivaga impensa Hopkins, 2014
Arenivaga pagana Hopkins, 2014
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Buprestidae (jewel beetles)
44
55
228
Acmaeodera pletura W. Barr, 1972
Melanophila piniedulis Burke, 1907
Acmaeodera navajo G. Nelson & Westcott, 1995
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Carabidae (ground beetles)
67
172
181
320
Bembidion (Cyclolopha) occultum Casey, 1918
Cicindela pusilla kaibabensis Johnson, 1990
Nebria (Reductonebria) georgei Kavanaugh, 2008
Cicindela rufiventris var. arizonae Wickham, 1899
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Cerambycidae (long-horned beetles)
63
64
65
66
Prionus spiculosus Casey, 1912
Prionus angustulus Casey, 1912
Prionus terminalis Casey, 1912
Stenosphenus pruddeni Casey, 1912
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Chrysomelidae (leaf beetles)
105
169
223
Bruchus perplexus Fall, 1910
Luperodes wickhami Horn, 1893
Saxinis saucia kaibabiae Moldenke, 1970
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Dermestide (dermestid beetles)
58
59
Cryptorhopalum pruddeni Casey, 1900
Orphilus aequalis Casey, 1900
________________________________________________________________________________
121
TAXONOMIC LIST
Arthropoda: Insecta: Coleoptera: Elateridae (click beetles)
196
Aphricus knowltoni Knull, 1957
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Leiodidae (round fungus beetles)
232
233
Ptomaphagus cocytus Peck, 1973
Ptomaphagus parashant Peck & Wynne, 2013
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Melyridae (soft-winged flower beetles)
56
57
Trichochrous incipiens Casey, 1895
Trichochrous reversus Casey, 1895
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Staphylinidae (rove beetles)
295
Eleodes (Caverneleodes) leptoscelis Triplehorn, 1975
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Tenebrionidae (darkling beetles)
60
61
62
173
230
296
Zopherodes lugubris Casey, 1907
Zopherodes pruddeni Casey, 1907
Discodemus brevipennis Casey, 1908
Trogloderus skillmani Johnston, 2019
Eschatomoxys pholeter Thomas & Pape in Pape, Thomas & Aalbu, 2007
Asidina rugicollis Triplehorn & Brown, 1971
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Ptinidae (spider beetles)
100
101
102
103
Petalium bistriatum Say var. arizonense Fall, 1905
Catorama grande Fall, 1905
Catorama longulum Fall, 1905
Catorama pingue Fall, 1905
________________________________________________________________________________
Arthropoda: Insecta: Coleoptera: Scarabaeidae (scarab beetles)
104
Diplotaxis conformis Fall, 1909
________________________________________________________________________________
122
TAXONOMIC LIST
Arthropoda: Insecta: Diptera: Asilidae (robber flies)
53
264
321
Erax benedicti Bromley, 1940
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
Heteropogon stonei Wilcox, 1965
________________________________________________________________________________
Arthropoda: Insecta: Diptera: Bombyliidae (bee flies)
85
152
Bombylius facialis Cresson, 1919
Geron prosopidis Hall & Evenhuis, 2003
________________________________________________________________________________
Arthropoda: Insecta: Diptera: Ceratopogonidae (biting midges)
1
Ceratopogon dimidiatus C. Adams, 1903
________________________________________________________________________________
Arthropoda: Insecta: Diptera: Dolichopodidae (longlegged flies)
256
300
301
314
Chrysotimus arizonicus H. Robinson, 1967
Polymedon dilaticosta Van Duzee, 1927
Polymedon nitidus Van Duzee, 1927
Polymedon castus Wheeler, 1899
________________________________________________________________________________
Arthropoda: Insecta: Diptera: Empididae (balloon flies)
273
Wiedemannia digna Sinclair, 2006
________________________________________________________________________________
Arthropoda: Insecta: Diptera: Mycetophilidae (fungus gnats)
72
271
Mycetophila neofungorum Chandler, 1993
Mycetophila denningi Shaw, 1951
________________________________________________________________________________
Arthropoda: Insecta: Diptera: Tachinidae (tachinid flies)
249
294
Fabriciella evanida Reinhard, 1953
Microsciasma minuta Townsend, 1915
________________________________________________________________________________
Arthropoda: Insecta: Diptera: Tephritidae (fruit flies)
81
Trypeta varipennis Coquillett, 1902
________________________________________________________________________________
123
TAXONOMIC LIST
Arthropoda: Insecta: Diptera: Tipulidae (crane flies)
3
Tipula (Lunatipula) kaibabensis C. Alexander, 1946
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Aphididae (aphids)
170
171
Cinara grande Hottes, 1956
Cinara poketa Hottes, 1956
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Cicadellidae (leafhoppers)
14
15
16
88
212
Athysanella fredonia Ball & Beamer, 1939
Athysanella globosa Ball & Beamer, 1939
Athysanella (Gladionura) casa Ball & Beamer, 1939
Ballana basala DeLong, 1964
Norvellina bicolorata (Ball) var. inflata Lindsay, 1938
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Cicadidae (cicadas)
87
Tibicen apache Davis, 1921
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Cixiidae (planthoppers)
11
Myndus yuccandus E. Ball, 1933
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Caliscelidae (planthoppers)
54
Aphelonema convergens var. canyonensia Bunn, 1930
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Coccidae (soft scales)
95
96
Ripersia arizonensis Ehrhorn, 1899
Dactylopius formicarii Ehrhorn 1899
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Coreidae (leaf-footed bugs)
216
Chelinidea vittiger Uhler var. artuflava McAtee, 1919
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Delphacidae (planthoppers)
47
Delphacodes apicata Beamer, 1948
________________________________________________________________________________
124
TAXONOMIC LIST
Arthropoda: Insecta: Hemiptera: Diaspididae (armored scales)
213
Chionaspis gilli Liu & Kosztarab, 1987
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Dictyopharidae (planthoppers)
13
Yucanda miniata E. Ball, 1937
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Issidae (planthoppers)
12
90
Hysteropterum cornutum var. utahnum E. Ball, 1935
Bruchomorpha bunni Doering, 1939
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Miridae (jumping tree bugs)
184
185
186
187
188
189
190
191
192
193
194
195
241
242
299
Deraeocoris fulvus Knight, 1921
Deraeocoris bullatus Knight, 1921
Deraeocoris navajo Knight, 1921
Ceratocapsus clavicornis Knight, 1925
Phytocoris mellarius Knight, 1925
Pilophorus fuscipennis Knight, 1926
Phytocoris hesperius Knight, 1928
Plagiognathus tenellus Knight, 1929
Phytocoris varius Knight, 1934
Parthenicus cowaniae Knight, 1968
Bolteria juniperi Knight, 1968
Phytocoris flaviatus Knight, 1968
Pilophorus americanus Poppius, 1914
Pilophorus exiguus Poppius, 1914
Platylygus vanduzeei Usinger, 1931
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Ochteridae (velvety shore bugs)
265
Ochterus barberi Schell, 1943
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Psyllidae (jumping plantlice)
298
Trioza rubra Tuthill, 1939
________________________________________________________________________________
Arthropoda: Insecta: Hemiptera: Reduviidae (assassin bugs)
34
Paratriatoma hirsuta Barber, 1938
________________________________________________________________________________
125
TAXONOMIC LIST
Arthropoda: Insecta: Hymenoptera: Andrenidae (andrenid bees)
200
253
290
291
292
293
Andrena (Callandrena) utahensis LaBerge, 1967
Andrena (Scaphandrena) kaibabensis Ribble, 1974
Perdita wheeleri Timberlake, 1928
Perdita opacella Timberlake, 1956
Perdita inornata Timberlake, 1962
Perdita depressa Timberlake, 1968
________________________________________________________________________________
Arthropoda: Insecta: Hymenoptera: Apidae (honey bees)
79
Triepeolus hopkinsi Cockerell, 1905
________________________________________________________________________________
Arthropoda: Insecta: Hymenoptera: Crabronidae (crabronid wasps)
111
Gorytes dentatus Fox, 1893
________________________________________________________________________________
Arthropoda: Insecta: Hymenoptera: Cynipidae (gall wasps)
51
Andricus wheeleri Beutenmüller, 1907
________________________________________________________________________________
Arthropoda: Insecta: Hymenoptera: Formicidae (ants)
80
Myrmecocystus melliger subsp. semirufus Emery var. romainei Cole, 1936 [not valid as a
quadrionomial taxon, and no identification of holotype; non Myrmecocstus romainei Snelling,
1975]
315
316
317
318
319
Pheidole desertorum var. maricopa Wheeler, 1906
Formica moki Wheeler, 1906
Formica fusca fusca var. gelida Wheeler, 1913 [not valid as a quadrinomial taxon]
Pogonomyrmex californicus Buckley subsp. maricopa Wheeler, 1914
Camponotus acutirostris Wheeler clarigaster Wheeler, 1915
________________________________________________________________________________
Arthropoda: Insecta: Hymenoptera: Pteromalidae (pteromalid wasps)
150
Boharticus margaretae Grissell, 1983
________________________________________________________________________________
Arthropoda: Insecta: Hymenoptera: Sphecidae (sphecid wasps)
52
Priononyx subatrata Bohart, 1958
________________________________________________________________________________
126
TAXONOMIC LIST
Arthropoda: Insecta: Hymenoptera: Vespidae (hornets)
50
155
Mischocyttarus flavitarsis var. kaibabensis Bequaert, 1932
Polistes canadensis var. kaibabensis Lynn, 1932
________________________________________________________________________________
Arthropoda: Insecta: Lepidoptera: Hesperiidae (skippers)
278
Megathymus alliae Stallings & Turner, 1957
________________________________________________________________________________
Arthropoda: Insecta: Lepidoptera: Noctuidae (owlet moths)
2
240
297
Plagiomimicus kathyae J. Adams & Lafontaine, 2009
Schinia immaculata Pogue, 2004
Hadena (Hadena) lafontainei Troubridge & Crabo, 2002
________________________________________________________________________________
Arthropoda: Insecta: Lepidoptera: Nymphalidae (admirals)
40
41
112
226
268
Coenonympha fureae Barnes & Benjamin, 1926
Cercyonis damei Barnes & Benjamin, 1926
Speyeria atlantis Edwards schellbachi Garth, 1949
Neonympha dorothea dorothea Nabokov, 1942
Neominois dionysus Scudder, 1878
________________________________________________________________________________
Arthropoda: Insecta: Lepidoptera: Oecophoridae (oecophorid moths)
76
Depressaria schellbachi Gates, 1947
________________________________________________________________________________
Arthropoda: Insecta: Lepidoptera: Papilionidae (swallowtails)
45
Papilio indra kaibabensis Bauer, 1955
________________________________________________________________________________
Arthropoda: Insecta: Lepidoptera: Pieridae (orange-tips)
269
Synchloe thoosa Scudder, 1878
________________________________________________________________________________
Arthropoda: Insecta: Orthoptera: Acrididae (short-horned grasshoppers)
270
Melanoplus canonicus Scudder, 1897
________________________________________________________________________________
127
TAXONOMIC LIST
Arthropoda: Insecta: Orthoptera: Eumastacidae (monkey grasshoppers)
248
Morsea california kaibabensis Rehn & Grant, 1958
________________________________________________________________________________
Arthropoda: Insecta: Psocodea: Enderleinellidae (sucking lice)
182
Enderleinellus kaibabensis Kim, 1966
________________________________________________________________________________
Arthropoda: Insecta: Siphonaptera: Ceratophyllidae (rodent fleas)
9
Orchopeas sexdentatus neotomae Augustson, 1943
________________________________________________________________________________
Class Malacostraca
(malacostracan crustaceans)
Arthropoda: Malacostraca: Amphipoda: Crangonyctidae (crangonyctid amphipods)
302
Stygobromus blinni Wang & Holsinger, 2001
________________________________________________________________________________
Phylum CHORDATA
Class Aves
(birds)
Chordata: Aves: Strigiformes: Strigidae (typical owls)
229
Otus asio mychophilus Oberholser, 1937
________________________________________________________________________________
Class Mammalia
(mammals)
Chordata: Mammalia: Carnivora: Felidae (cats)
227
Felis concolor kaibabensis E. Nelson & Goldman, 1931
________________________________________________________________________________
Chordata: Mammalia: Carnivora: Mephitidae (skunks)
219
Spilogale gracilis Merriam, 1890
________________________________________________________________________________
128
TAXONOMIC LIST
Chordata: Mammalia: Rodentia: Castoridae (beavers)
121
Castor canadensis repentinus Goldman, 1932
________________________________________________________________________________
Chordata: Mammalia: Rodentia: Crecitidae (deer mice)
122
218
Peromyscus crinitus peridoneus Goldman, 1937
Hesperomys megalotis Merriam, 1890
________________________________________________________________________________
Chordata: Mammalia: Rodentia: Geomyidae (pocket gophers)
91
123
124
151
Thomomys bottae boreorarius Durham, 1952
Thomomys fossor kaibabensis Goldman, 1938
Thomomys bottae nicholi Goldman, 1938
Thomomys bottae trumbullensis Hall & Davis, 1934
________________________________________________________________________________
Chordata: Mammalia: Rodentia: Heteromyidae (kangaroo rats)
118
119
Dipodomys microps celsus Goldman, 1924
Dipodomys ordii cupidineus Goldman, 1924
________________________________________________________________________________
Chordata: Mammalia: Rodentia: Sciuridae (squirrels)
120
220
Ammospermophilus leucurus tersus Goldman, 1929
Sciurus kaibabensis Merriam, 1904
________________________________________________________________________________
Class Reptilia
(reptiles)
Chordata: Reptilia: Squamata: Colubridae (typical snakes)
281
285
Sonora semiannulata gloydi Stickel, 1938
Lampropeltis pyromelana Cope infralabialis Tanner, 1953
________________________________________________________________________________
Chordata: Reptilia: Squamata: Viparidae (rattlesnakes)
183
Crotalus confluentus abyssus Klauber, 1930
________________________________________________________________________________
129
TAXONOMIC LIST
Class Teleostei
(fishes)
Chordata: Teleostei: Cypriniformes: Cyprinidae (carp)
221
Gila cypha R. Miller, 1946
________________________________________________________________________________
Phylum MOLLUSCA
(mollusks)
Class Gastropoda
(snails)
Mollusca: Gastropoda: Neotaenioglossa: Hydrobiidae (spring snails)
157
Pyrgulopsis hualapaiensis Herschler, Liu & Stevens, 2016
________________________________________________________________________________
Mollusca: Gastropoda: Stylommatophora: Helminthoglyptidae (shoulderbands)
156
Sonorella betheli Henderson, 1914 [an erroneous account; see details in the bibliography
herein]
222
280
Sonorella reederi W. Miller, 1984
Helix (Arionta) coloradoensis Stearns, 1890
________________________________________________________________________________
Mollusca: Gastropoda: Stylommatophora: Oreohelicidae (mountain snails)
214
236
237
238
Oreohelix yavapai vauxae Marshall, 1929
Oreohelix yavapai profundorum Pilsbry & Ferriss, 1911
Oreohelix yavapai extremitatis Pilsbry & Ferriss, 1911
Oreohelix yavapai angelica Pilsbry & Ferriss, 1911
________________________________________________________________________________
Mollusca: Gastropoda: Stylommatophora: Pupillidae (chrysalis snails)
239
Pupilla syngenes avus Pilsbry & Ferriss, 1911
________________________________________________________________________________
130
TAXONOMIC LIST
Phylum NEMATODA
(roundworms)
Class Chromadorea
(chromadorid roundworms)
Nematoda: Chromadorea: Ascaridida: Oxyuridae (pinworms)
197
198
Wellcomia perognathi Kruidenier & Mehra, 1958
Aspiculuris ackerti Kruidenier & Mehra, 1959
________________________________________________________________________________
Nematoda: Chromadorea: Rhabditida : Rhabditidae (rhabditid roundworms)
215
Parasitorhabditis gracilis Massey, 1974
________________________________________________________________________________
Nematoda: Chromadorea: Spiurida: Gongylonematidae (spiurid gullet worms)
199
Gongylonema peromysci Kruidenier & Peebles, 1958
________________________________________________________________________________
131
TAXONOMIC LIST
KINGDOM PLANTAE
(plants)
Phylum (Division) TRACHYOPHYTA
(vascular plants)
Class Magnoliopsida
(flowering plants)
Trachyophyta: Magnoliopsida: Asparagales: Asparagaceae (agaves)
162
217
309
Agave phillipsiana Hodgson, 2001
Agave kaibabensis McKelvey, 1949
Androstephium breviflorum Watson, 1873
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Asterales: Asteraceae (sunflowers)
8
75
129
147
177
178
286
307
Xylorhiza tortifolia (Torrey & Gray) Greene var. parashantensis Atwood & Welsh, 2013
Encelia resinifera subsp. tenuifolia Curtis, 1998
Actinella biennis A. Gray, 1878
Senecio stygius Greene, 1909
Laphamia congesta Jones, 1895
Laphamia gracilis Jones, 1895
Flaveria mcdougallii Theroux, Pinkava & Keil, 1977
Brickellia (Clavigera) longifolia Watson, 1873
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Boraginales: Boraginaceae (borage)
94
160
283
284
Oreocarya capitata Eastwood, 1937
Cryptantha atwoodii Higgins, 1974
Amsinckia nana Suksdorf, 1931
Amsinckia macrosepala Suksdorf, 1931
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Boraginales: Hydrophyllaceae (waterleafs)
5
6
7
131
Phacelia furnissii Atwood, 2007
Phacelia higginsii Atwood, 2007
Phacelia hughesii Atwood, 2007
Phacelia (Coreanthus) ivesiana A. Gray in Gray et al., 1861
________________________________________________________________________________
132
TAXONOMIC LIST
Trachyophyta: Magnoliopsida: Brassicales: Apiaceae (parsleys)
83
305
306
Phellopterus multinervatus Coulter & Rose, 1900
Cymopterus purpureus Watson, 1873
Peucedanum newberryi Watson, 1873
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Brassicales: Brassicaceae [Cruciferae] (mustards)
4
134
135
257
258
310
Thelypodium integrifolium (Nuttall) Endlicher subsp. longicarpum Al-Shehbaz, 1973
Arabis eremophila Greene, 1900
Arabis recondita Greene, 1900
Sisymbrium kearneyi Rollins, 1953
Lesquerella kaibabensis Rollins, 1982
Lesquerella arizonica Watson, 1888
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Caryophyllales: Amaranthaceae (pigweed)
282
Atriplex asterocarpa Stutz, Chu & Sanderson, 1994
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Caryophyllales: Cactaceae (cacti)
46
49
77
78
82
108
109
158
159
231
Opuntia aurea Baxter, 1933
Pediocactus paradinei Benson, 1957
Sclerocactus havasupaiensis Clover, 1942
Sclerocactus havasupaiensis var. roseus Clover, 1942
Echinocactus polycephalus var. xeranthemoides Coulter, 1896
Coryphantha vivipara var. kaibabensis Fischer, 1979
Echinocereus triglochidiatus var. toroweapensis Fischer, 1991
Opuntia hualpaensis Hester, 1943
Opuntia abyssi Hester, 1943
Opuntia pinkavae Parfitt, 1997
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Caryophyllales: Caryophyllaceae (pinks)
254
Silene rectiramea Robinson, 1899
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Caryophyllales: Nyctaginaceae (four o’clocks)
130
279
Mirabilis bigelovii A. Gray, 1886
Allionia pachyphylla Standley, 1909
________________________________________________________________________________
133
TAXONOMIC LIST
Trachyophyta: Magnoliopsida: Caryophyllales: Polygonaceae (knotweeds)
250
251
274
Eriogonum mortonianum Reveal, 1974a
Eriogonum thompsonae S. Wats. var. atwoodii Reveal, 1974b
Eriogonum fusiforme Small, 1906
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Cornales: Losaceae (loasa)
73
165
266
267
Mentzelia collomiae Christy, 1997
Mentzelia memorabilis N. Holmgren & P. Holmgren, 2002
Mentzelia hualapaiensis Schenk, Hodgson & Hufford, 2010
Mentzelia canyonensis Schenk, Hodgson & Hufford, 2013
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Crossosomatales: Crossosomataceae (crossomas)
255 Crossosoma parviflora B. Robinson & Fernald, 1894
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Ericales: Polemoniaceae (phlox)
“Gilia dactylophyllum, (n. sp.?)”, A. Gray in Gray et al., 1861 [see note in bibliography herein]
________________________________________________________________________________
132
Trachyophyta: Magnoliopsida: Ericales: Primulaceae (primroses)
Primula (Section Farinosae) hunnewellii Fernald, 1934
________________________________________________________________________________
106
Trachyophyta: Magnoliopsida: Fabales: Fabaceae (legumes)
35
36
37
38
39
125
126
127
128
174
179
180
261
303
311
312
313
Astragalus bryantii Barneby, 1944
Astragalus lentiginosus Dougl. var. oropedii Barneby, 1945
Astragalus cremnophylax Barneby, 1948
Astragalus wootoni var. endopterus Barneby, 1949
Astragalus cremnophylax Barneby var. myriorrhaphis Barneby, 1979
Astragalus subcinereus A. Gray, 1878
Astragalus scaposus A. Gray, 1878
Astragalus mokiacensis A. Gray, 1878
Astragalus artipes A. Gray, 1878
Astragalus humistratus var. tenerrimus Jones, 1895
Astragalus kentrophyta var. coloradoensis Jones, 1902
Astragalus kaibensis Jones, 1902
Pediomelum retrorsum Rydberg in Rydberg & Pennell, 1919
Dalea amoena Watson, 1873
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001
Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010
Astragalus pinionis Welsh & Thorne, 1977
________________________________________________________________________________
134
TAXONOMIC LIST
Trachyophyta: Magnoliopsida: Fagales: Betulaceae (birches)
84
Ostrya knowltonii Coville, 1894
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Gentianales: Apocynaceae (dogbane)
260
Amsonia eastwoodiana Rydberg, 1913
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Gentianales: Rubiaceae (madders)
89
Galium munzii Hilend & Howell subsp. ambivalens Dempster & Ehrendorfer, 1965
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Lamiales: Lamiaceae (mints)
252
Scutellaria potosina var. kaibabensis Rhodes & Ayers, 2010
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Lamiales: Oleaceae (olives)
93
176
246
247
Fraxinus macropetala Eastwood, 1903
Bigelovia howardi var. attenuata Jones, 1895
Fraxinus cuspidata var. macropetala Rehder, 1917
Fraxinus lowellii Sargent in Rehder, 1917
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Lamiales: Orobanchaceae (broomrapes)
74
107
234
Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986
Adenostegia parviflora Ferriss, 1918
Cordylanthus tenuifolis Pennell, 1940
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Lamiales: Plantaginaceae (plantains)
86
259
Penstemon virgatus pseudoptutus Crosswhite, 1967
Penstemon coccinatus Rydberg, 1909
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Lamiales: Scrophulariaceae (figworts)
164
Castilleja kaibabensis N. Holmgren, 1973
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Liliales: Liliaceae (lilies)
308
Calochortus flexuosus Watson, 1873
________________________________________________________________________________
135
TAXONOMIC LIST
Trachyophyta: Magnoliopsida: Malpighiales: Euphorbiaceae (spurge)
163
Euphorbia aaronrossii A. Holmgren & N. Holmgren, 1988
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Malpighiales: Salicaceae (willows)
10
263
Salix lutea var. ligulifolia C. Ball, 1921
Populus fremontii var. toumeyi Sargent, 1919
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Myrtales: Onagraceae (evening primroses)
175
243
244
244a
304
Oenothera triloba var. ecristata Jones, 1895
Oenothera confertiflora Raven, 1962
Oenothera specuicola subsp. specuicola Raven, 1962
Oenothera specuicola subsp. hesperia Raven, 1962
Oenothera (Chylisma) multijuga Watson, 1873
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Poales: Cyperaceae (sedges)
276
Carex curatorum Stacey, 1937
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Poales: Poaceae (grasses)
245
Panicum mohavense Reeder, 1991
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Ranunculales: Papaveraceae (poppies)
92
Corydalis wetherillii Eastwood, 1902
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Rosales: Moraceae (mulberries)
148
Morus grisea Greene, 1911
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Rosales: Roasaceae (roses)
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
________________________________________________________________________________
136
TAXONOMIC LIST
Trachyophyta: Magnoliopsida: Sapindales: Anacardiaceae
133
136
137
(cashews)
Rhus canadensis Marsh. var. simplicifolia Greene, 1890
Schmaltzia cissodes Greene, 1905
Schmaltzia hirtella Green, 1905
________________________________________________________________________________
Trachyophyta: Magnoliopsida: Sapindales: Rutaceae (rues)
138
139
140
141
142
143
144
145
146
Ptelea palida Green, 1906
Ptelea straminea Green, 1906
Ptelea nitida Green, 1906
Ptelea argentea Green, 1906
Ptelea triptera Green, 1906
Ptelea lutescens Green, 1906
Ptelea elegans Green, 1906
Ptelea confinis Green, 1906
Ptelea saligna Green, 1906
________________________________________________________________________________
Class Pinopsida
(conifers)
Trachyophyta: Pinopsida: Pinales: Cupressaceae (cypress)
262
Juniperus utahensis var. megalocarpa Sargent, 1919
________________________________________________________________________________
137
TAXONOMIC LIST
KINGDOM FUNGI
(fungi)
Phylum (Division) ASCOMYCOTA
(sac fungi)
Ascomycota: Dothidiomycetes: Pleosporales: Venturiaceae
43
Protoventuria arizonica M. Barr, 1989
________________________________________________________________________________
Ascomycota: Eurotiomycetes: Eurotiales: Trichocomaceae (saprobe fungi)
149
Penicillium arizonense Frisvad, Grijseels & J.C. Nielsen in Grijseels et al., 2016
________________________________________________________________________________
Ascomycota: Lecanoromycetes: Teloschistales: Physciaceae
99
Physconia isidiomuscigena Esslinger, 2000
________________________________________________________________________________
Ascomycota: Orbiliomycetes: Orbiliales: Orbiliaceae
17
18
19
20
21
22
23
24
25
26
27
28
29
30
31
32
33
Orbilia purshiae Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia ocellata Baral, G. Marson & E. Weber in Baral, Weber & Marson, 2020
Orbilia magnifica Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia phanosoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia multiphanosoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia arizonensis Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia navajoana Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia spermoides Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia ophiosoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia multimaeandrina Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia delphinus Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia macrodelphinus Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia multitrapezoidea Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia calyptrata Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia subovoidea Baral, Matočec & E. Weber in Baral, Weber & Marson, 2020
________________________________________________________________________________
138
TAXONOMIC LIST
Ascomycota: Pezizomycetes: Pezizales: Morchellaceae
42
Morchella kaibabensis Beug, T.A. Clem. & T.J. Baroni in Baroni et al., 2018
________________________________________________________________________________
Ascomycota: Sordariomycetes: Diaporthales: Valsaceae
224
Gnomonia quercusgambellii Monod, 1983
________________________________________________________________________________
Ascomycota incertae sedis
97
Septogloeum gillii D. Ellis, 1946
________________________________________________________________________________
Phylum (Division) BASIDIOMYCOTA
(club fungi)
Basidiomycota: Agaricomycetes: Boletales: Boletaceae
287
Boletus barrowsii Thiers & Smith in Thiers, 1976
________________________________________________________________________________
Basidiomycota: Agaricomycetes: Boletales: Suillaceae
288
289
Suillus kaibabensis Thiers, 1976
Suillus occidentalis Thiers, 1976
________________________________________________________________________________
Basidiomycota: Pucciniomycetes: Pucciniales: Pucciniaceae
98
Puccinia circinans J. Ellis & Everhart, 1900
________________________________________________________________________________
139
TAXONOMIC LIST
KINGDOM CHROMISTA
(photosynthetic eukaryotes)
Phylum APICOMPLEXA
(parasitic alveolates)
Apicomplexa: Class : Eucoccidiorida: Eimeriidae (coccid parasites)
201
202
203
204
205
206
207
208
209
210
211
Eimeria tamiasciuri Levine, Ivens & Kruidenier, 1957a
Eimeria cutamiae Levine, Ivens & Kruidenier, 1957a
Eimeria thomomysis Levine, Ivens & Kruidenier, 1957a
Eimeria perognathi Levine, Ivens & Kruidenier, 1957a
Eimeria albigulae Levine, Ivens & Kruidenier, 1957a
Eimeria operculata Levine, Ivens & Kruidenier, 1957a
Eimeria peromysci Levine, Ivens & Kruidenier, 1957a
Eimeria arizonensis Levine, Ivens & Kruidenier, 1957a
Eimeria eremici Levine, Ivens & Kruidenier, 1957a
Eimeria onychomysis Levine, Ivens & Kruidenier, 1957a
Isopora citolli Levine, Ivens & Kruidenier, 1957b
________________________________________________________________________________
KINGDOM PROTISTA
(protists)
“Phylum” HEMIMASTIGOPHORA
(single-celled eukaryotes)
Hemimastigophora: Hemimastigea: Hemimastigida: Spironemidae (hemimastigid protist)
110
Spironema terricola Foissner & Foissner, 1993
________________________________________________________________________________
140
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK *
[In the PDF document the lines immediately below are hyperlinks]
page
“Grand Canyon” not discriminated
143
South Rim
144
North Rim
150
Inner Canyon
154
Colorado River Locales
161
Locality data are as described in the original publications; thus some minor variances will be
noted for specific sites or areas that are otherwise in very close proximity to each other.
Refinements or special notes appear within [non-bold square brackets].
Red Numbers indicate and cross-list the sequentially enumerated taxa in the bibliography
(thus they reflect the alphabetical order by publication author).
_________________
* Regarding localities that are today within the Havasupai Indian Reservation or the Hualapai Indian
Reservation, but which lie within the physiographic Grand Canyon, see pertinent entries in the
LOCALITIES FOR TYPE MATERIAL FROM NEAR GRAND CANYON list.
141
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Federally-Designated Boundaries of
Grand Canyon National Park and Predecessor Units
1893–Present
map by Stephanie Smith, Grand Canyon Trust, 2019
Grand Canyon Trust
https://www.grandcanyontrust.org/grand-canyon-historic-boundaries-map
non-commercial permission to use indicated on webpage; last accessed 29 November 2022
142
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
“GRAND CANYON”
Grand Canyon or Colorado Canyon [without specific locale]
1
58
59
60
61
62
63
64
65
66
67
79
93
97
98
107
111
133
137
140
141
167
183
192
234
246
259
265
270
292
298
314
318
7
Ceratopogon dimidiatus Adams, 1903
Cryptorhopalum pruddeni Casey, 1900
Orphilus aequalis Casey, 1900
Zopherodes lugubris Casey, 1907
Zopherodes pruddeni Casey, 1907
Discodemus brevipennis Casey, 1908
Prionus spiculosus Casey, 1912
Prionus angustulus Casey, 1912
Prionus terminalis Casey, 1912
Stenosphenus pruddeni Casey, 1912
Bembidion (Cyclolopha) occultum Casey, 1918
Triepeolus hopkinsi Cockerell, 1905
Fraxinus macropetala Eastwood, 1903
Septogloeum gillii D. Ellis, 1946
Puccinia circinans J. Ellis & Everhart, 1900
Adenostegia parviflora Ferriss, 1918
Gorytes dentatus Fox, 1893
Rhus canadensis Marsh. var. simplicifolia Greene, 1890
Schmaltzia hirtella Green, 1905
Ptelea nitida Greene, 1906
Ptelea argente Greene, 1906
Arenivaga impensa Hopkins, 2014
Crotalus confluentus abyssus Klauber, 1930
Phytocoris varius Knight, 1934
Cordylanthus tenuifolis Pennell, 1940
Fraxinus cuspidata var. macropetala Rehder, 1917
Penstemon coccinatus Rydberg, 1909
Ochterus barberi Schell, 1943
Melanoplus canonicus Scudder, 1897
Perdita inornata Timberlake, 1962
Trioza rubra Tuthill, 1939
Polymedon castus Wheeler, 1899
Pogonomyrmex californicus Buckley subsp. maricopa Wheeler, 1914
_________________________________________________________________________________
7
Most of these undiscriminated “Grand Canyon” records are likely based on collections on the far
more usually visited South Rim but are listed thusly for their lack of further data. See also “Grand
Canyon, Arizona” and (under the “South Rim” heading below) “Grand Canyon village”. Many taxa
are represented by material collected prior to the creation of the national monument or park.
For localities described as in the “vicinity of”, “near”, or “about” the Grand Canyon, see the
LOCALITIES OF TYPE MATERIAL NEAR GRAND CANYON list.
143
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
“Grand Canyon National Park” [without specific locale]
197
199
Wellcomia perognathi Kruidenier & Mehra, 1958
Gongylonema peromysci Kruidenier & Peebles, 1958
_________________________________________________________________________________
“Grand Canyon, Arizona” [the canyon or Grand Canyon village not discriminated]
14
15
16
40
41
47
53
54
88
90
200
212
215
216
271
275
299
Athysanella fredonia Ball & Beamer, 1939
Athysanella globosa Ball & Beamer, 1939
Athysanella (Gladionura) casa Ball & Beamer, 1939
Coenonympha fureae Barnes & Benjamin, 1926
Cercyonis damei Barnes & Benjamin, 1926
Delphacodes apicata Beamer, 1948
Erax benedicti Bromley, 1940
Aphelonema convergens var. canyonensia Bunn, 1930
Ballana basala DeLong, 1964
Bruchomorpha bunni Doering, 1939
Andrena (Callandrena) utahensis LaBerge, 1967
Norvellina bicolorata (Ball) var. inflata Lindsay, 1938
Parasitorhabditis gracilis Massey, 1974
Chelinidea vittiger Uhler var. artuflava McAtee, 1919
Mycetophila denningi Shaw, 1951
Heterotarsonemus nakaharai Smiley & Moser, 1974
Platylygus vanduzeei Usinger, 1931
_________________________________________________________________________________
SOUTH RIM
[Grand Canyon National Park]
“Grand Canyon South Rim” [without specific locale]
44
52
152
226
292
Acmaeodera pletura W. Barr, 1972
Priononyx subatrata Bohart, 1958
Geron prosopidis Hall & Evenhuis, 2003
Neonympha dorothea dorothea Nabokov, 1942
Perdita inornata Timberlake, 1962
_________________________________________________________________________________
“Grand Canyon National Park, South Rim” [without specific locale]
299
Platylygus vanduzeei Usinger, 1931
_________________________________________________________________________________
144
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
“Rim of Grand Canyon” [without specific locale]
85
262
Bombylius facialis Cresson, 1919
Juniperus utahensis var. megalocarpa Sargent, 1919
_________________________________________________________________________________
Arizona Route 64 at southeast boundary of Grand Canyon National Park
[near Desert View]
210
Eimeria onychomysis Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
Bright Angel, Arizona [Grand Canyon village]
see also Grand Canyon village
100
101
102
103
104
105
241
242
262
Petalium bistriatum Say var. arizonense Fall, 1905
Catorama grande Fall, 1905
Catorama longulum Fall, 1905
Catorama pingue Fall, 1905
Diplotaxis conformis Fall, 1909
Bruchus perplexus Fall, 1910
Pilophorus americanus Poppius, 1914
Pilophorus exiguus Poppius, 1914
Juniperus utahensis var. megalocarpa Sargent, 1919
_________________________________________________________________________________
Bright Angel Hotel [Grand Canyon village]
see also Grand Canyon village
55
81
Melanophila piniedulis Burke, 1907
Trypeta varipennis Coquillett, 1902
_________________________________________________________________________________
Bright Angel Trail (upper entrance)
see also Bright Angel Trail under “Inner Canyon” listings
110
Spironema terricola Foissner & Foissner, 1993
_________________________________________________________________________________
Bright Angel Trail (above)
195
Phytocoris flaviatus Knight, 1968
_________________________________________________________________________________
145
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Bright Angel Trail (at top)
190
191
Phytocoris hesperius Knight, 1928
Plagiognathus tenellus Knight, 1929
_________________________________________________________________________________
Cataract Canyon, above Supai
[The precise locality may be within the boundaries of the Havasupai Indian Reservation]
77
Sclerocactus havasupaiensis Clover, 1942
_________________________________________________________________________________
Cedar Mountain, east side
[The precise locality may be within the boundaries of the Navajo Nation]
205
Eimeria albigulae Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
Coconino Forest, Grand Canyon (7000 feet)
317
Formica fusca fusca var. gelida Wheeler, 1913 [not valid as a quadrinomial taxon]
_________________________________________________________________________________
“on the rim in the Kohonino [Coconino] Forest (7000 feet) about three miles west of the
Bright Angel Hotel”
316
Formica moki Wheeler, 1906
_________________________________________________________________________________
Coconino Forest, rim of Grand Canyon
51
Andricus wheeleri Beutenmüller, 1907
_________________________________________________________________________________
Desert View Point
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
El Tovar (ca. 2 miles west of)
37
Astragalus cremnophylax Barneby, 1948
_________________________________________________________________________________
Eremita Mesa
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
_________________________________________________________________________________
146
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Grand Canyon village
see also listings under “Bright Angel, Arizona” and “Bright Angel Hotel” above
68
229
Aphonopelma behlei Chamberlin, 1940
Otus asio mychophilus Oberholser, 1937
_________________________________________________________________________________
Grand Canyon village, 36° 03′ 22.31″ N; 112° 07′ 30.73″ W
[coordinates pinpoint a locality just to the west of the footpath between Park Headquarters and the
Rim Trail, at a point half-way between those two places]
149
Penicillium arizonense Frisvad, Grijseels & J.C. Nielsen in Grijseels et al., 2016
_________________________________________________________________________________
Grand Canyon village (7.5 km east-southeast of), close to South Rim
[Grand Canyon National Park, along Arizona Route 64]
17
22
Orbilia purshiae Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia arizonensis Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
Grand Canyon village (8.5 km east-southeast of), close to South Rim
[Grand Canyon National Park, along Arizona Route 64]
26
Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
Grand View [Grandview]
see also Hearst Ranch
184
185
186
187
188
189
193
Deraeocoris fulvus Knight, 1921
Deraeocoris bullatus Knight, 1921
Deraeocoris navajo Knight, 1921
Ceratocapsus clavicornis Knight, 1925
Phytocoris mellarius Knight, 1925
Pilophorus fuscipennis Knight, 1926
Parthenicus cowaniae Knight, 1968
_________________________________________________________________________________
Grandview Point
26
29
Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia delphinus Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
147
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Hance Ranch
see also Thurber’s Camp
142
Ptelea triptera Green, 1906 [type locality for this species refined here; see entry in
bibliography]
_________________________________________________________________________________
Hearst Ranch
see also Grandview
203
Eimeria thomomysis Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
Hermit Rim Road
193
Parthenicus cowaniae Knight, 1968
_________________________________________________________________________________
Hualapai Canyon (tributary of Havasu Canyon)
[The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.]
77
Sclerocactus havasupaiensis Clover, 1942
_________________________________________________________________________________
Hull Tank
202
Eimeria cutamiae Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
Mather Point
24
Orbilia spermoides Baral & G. Marson, in Baral, Weber & Marson, 2020
_________________________________________________________________________________
Navajo Point (near) (Coconino Plateau)
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
Shoshone Point
76
Depressaria schellbachi Gates, 1947
_________________________________________________________________________________
148
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
South entrance to Grand Canyon National Park
Orchopeas sexdentatus neotomae Augustson, 1943
9
_________________________________________________________________________________
“behind the cabin camp located at the entrance gate of the Grand Canyon National Park”
192
Phytocoris varius Knight, 1934
_________________________________________________________________________________
Thurber’s Camp [Hance Ranch]
95
96
Ripersia arizonensis Ehrhorn, 1899
Dactylopius formicarii Ehrhorn 1899
_________________________________________________________________________________
Tusayan (3.5 km north of)
[Grand Canyon National Park, on Arizona Route 64]8
23
25
28
30
32
Orbilia navajoana Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia ophiosoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia macrodelphinus Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia calyptrata Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
Tusayan (3 km north of)
[Grand Canyon National Park, on Arizona Route 64]8
18
19
19
20
21
23
24
28
29
Orbilia ocellata Baral, G. Marson & E. Weber in Baral, Weber & Marson, 2020
Orbilia cucumispora Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia magnifica Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia phanosoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia multiphanosoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia navajoana Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia spermoides Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia delphinus Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
8
See the TYPE LOCALITIES NEAR GRAND CANYON list for localities south of Tusayan, Arizona.
149
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Yavapai Point
45
Papilio indra kaibabensis Bauer, 1955
_________________________________________________________________________________
Zuni Point
206
Eimeria operculata Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
Zuni Point, west side
207
208
Eimeria peromysci Levine, Ivens & Kruidenier, 1957a
Eimeria arizonensis Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
NORTH RIM
[Grand Canyon National Park]
Grand Canyon (North Rim of) [without specific locale]
44
72
170
224
253
256
Acmaeodera pletura W. Barr, 1972
Mycetophila neofungorum Chandler, 1993
Cinara grande Hottes, 1956
Gnomonia quercusgambellii Monod, 1983
Andrena (Scaphandrena) kaibabensis Ribble, 1974
Chrysotimus arizonicus H. Robinson, 1967
_________________________________________________________________________________
Grand Canyon (North Rim)
[“Grand Canyon North Rim” or place name “North Rim, Arizona”, not discriminated]
see also “North Rim”
43
45
106
112
113
Protoventuria arizonica M. Barr, 1989
Papilio indra kaibabensis Bauer, 1955
Primula (Section Farinosae) hunnewellii Fernald, 1934
Speyeria atlantis schellbachi Garth, 1949
Misumenops coloradensis Gertsch, 1933
_________________________________________________________________________________
Kaibab Plateau [within Grand Canyon National Park]
217
Agave kaibabensis McKelvey, 1949 [see entry in bibliography herein for notes concerning the
locality]
_________________________________________________________________________________
150
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Kaibab Plateau, North Rim of Grand Canyon [undiscriminated]
3
Tipula (Lunatipula) kaibabensis C. Alexander, 1946
_________________________________________________________________________________
Bright Angel Creek (head of), Kaibab Plateau
220
Sciurus kaibabensis Merriam, 1904
_________________________________________________________________________________
Bright Angel Point
45
86
Papilio indra kaibabensis Bauer, 1955
Penstemon virgatus pseudoptutus Crosswhite, 1967
_________________________________________________________________________________
Bright Angel Point (“from or near”)
285
Lampropeltis pyromelana Cope infralabialis Tanner, 1953
_________________________________________________________________________________
Burro Spring
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
Cape Royal
208
Eimeria arizonensis Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
Cape Royal Road and Point Imperial Road, W of junction
86
Penstemon virgatus pseudoptutus Crosswhite, 1967
_________________________________________________________________________________
Grand Canyon National Park, North Rim, junction of paved roads, ca. 6.5 km north of Kaibab
Lodge, 36° 16′ N, 112° 03′ W
[actually Grand Canyon Lodge in Grand Canyon National Park; coordinates pinpoint a location just
east of Arizona Route 67 north of the Cape Royal Road junction]
99
Physconia isidiomuscigena Esslinger, 2000
_________________________________________________________________________________
Dutton Point
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
_________________________________________________________________________________
151
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Mount Trumbull
268
Neominois dionysus Scudder, 1878
_________________________________________________________________________________
Neal Spring
[junction of Cape Royal Road and Point Imperial Road]
112
Speyeria atlantis schellbachi Garth, 1949
_________________________________________________________________________________
North Rim [the place, “North Rim, Arizona”]
155
Polistes canadensis var. kaibabensis Lynn, 1932 [see note in bibliography]
_________________________________________________________________________________
North Rim (S of) (93 km south-southeast of Fredonia)
[Grand Canyon National Park, North Rim, Arizona]
17
22
26
28
29
33
Orbilia purshiae Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia arizonensis Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia delphinus Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia subovoidea Baral, Matočec & E. Weber in Baral, Weber & Marson, 2020
_________________________________________________________________________________
Phantom Canyon (head of)
35
Astragalus bryantii Barneby, 1944
_________________________________________________________________________________
Point Sublime
155
289
Polistes canadensis var. kaibabensis Lynn, 1932
Suillus occidentalis Thiers, 1976
_________________________________________________________________________________
Powell Plateau
227
Felis concolor kaibabensis E. Nelson & Goldman, 1931
_________________________________________________________________________________
Road W-3 on the north rim of Grand Canyon near the entrance to Grand Canyon National
Park
201
Eimeria tamiasciuri Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
152
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Robbers’ Roost Spring
112
Speyeria atlantis schellbachi Garth, 1949
_________________________________________________________________________________
S B Canyon, Grand Canyon National Monument
[now in the enlarged Grand Canyon National Park]
257
Sisymbrium kearneyi Rollins, 1953
_________________________________________________________________________________
S B Point and Hades Knoll (side canyon between)
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
_________________________________________________________________________________
S B Trail (head of)
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
_________________________________________________________________________________
Swamp Lake
112
Speyeria atlantis schellbachi Garth, 1949
_________________________________________________________________________________
Swamp Point
91
112
Thomomys bottae boreorarius Durham, 1952
Speyeria atlantis schellbachi Garth, 1949
_________________________________________________________________________________
Swamp Point (½ mile from) (near Powell Saddle)
211
Isopora citolli Levine, Ivens & Kruidenier, 1957b
_________________________________________________________________________________
Swamp Ridge
112
Speyeria atlantis schellbachi Garth, 1949
_________________________________________________________________________________
Toroweap Point
109
Echinocereus triglochidiatus var. toroweapensis Fischer, 1991
_________________________________________________________________________________
153
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Two River Junction
112
Speyeria atlantis schellbachi Garth, 1949
_________________________________________________________________________________
Vulcan’s Throne, Toroweap Valley
243
Oenothera confertiflora Raven, 1962
_________________________________________________________________________________
INNER CANYON
[Grand Canyon National Park]
Bat Cave [interior]
230
Eschatomoxys pholeter Thomas & Pape in Pape, Thomas & Aalbu, 2007
_________________________________________________________________________________
Bright Angel Canyon
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
Bright Angel Creek
121
Castor canadensis repentinus Goldman, 1932
_________________________________________________________________________________
Bright Angel Creek (near)
92
Corydalis wetherillii Eastwood, 1902
_________________________________________________________________________________
Bright Angel Trail [without specific locale]
see also Bright Angel Trail under “South Rim” listings (above); and “Garden Creek” and “Indian
Garden” (below)
50
93
143
156
281
300
301
Mischocyttarus flavitarsis var. kaibabensis Bequaert, 1932
Fraxinus macropetala Eastwood, 1903
Ptelea lutescens Greene, 1906
Sonorella betheli Henderson, 1914 [an erroneous account; see details in the bibliography]
Sonora semiannulata gloydi Stickel, 1938
Polymedon dilaticosta Van Duzee, 1927
Polymedon nitidus Van Duzee, 1927
_________________________________________________________________________________
154
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Bright Angel Trail, 100-400 feet below rim
238
Oreohelix yavapai angelica Pilsbry & Ferriss, 1911
_________________________________________________________________________________
“wall of the Grand Cañon at an altitude of about 5500 feet”
[in all probability on Bright Angel Trail]
316
Formica moki Wheeler, 1906
_________________________________________________________________________________
Bright Angel Trail, 4800 ft altitude
[altitude indicates locality is at about the 2nd rest house descending]
122
Peromyscus crinitus peridoneus Goldman, 1937
_________________________________________________________________________________
Bright Angel Trail, 1400-2200 m altitude
246
Fraxinus cuspidata var. macropetala Rehder, 1917
_________________________________________________________________________________
Bright Angel Trail, 1400-2000 m altitude
246
Fraxinus cuspidata var. macropetala Rehder, 1917
_________________________________________________________________________________
Bright Angel Trail, about 3000 ft altitude
319
Camponotus acutirostris Wheeler clarigaster Wheeler, 1915
_________________________________________________________________________________
Bright Angel Trail, near Indian Garden [Havasupai Gardens]
136
Schmaltzia cissodes Greene, 1905
_________________________________________________________________________________
Bright Angel Trail, ca. 0.5 mile below Indian Garden [Havasupai Gardens]
4
Thelypodium integrifolium (Nuttall) Endlicher subsp. longicarpum Al-Shehbaz, 1973
_________________________________________________________________________________
Bright Angel Trail, lower
87
Tibicen apache Davis, 1921
_________________________________________________________________________________
155
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Bright Angel Trail, Lower Sonoran level
281
Sonora semiannulata gloydi Stickel, 1938
_________________________________________________________________________________
Cave of the Domes [interior]
[on Horseshoe Mesa]
225
295
Archeolarca cavicola Muchmore, 1981
Eleodes (Caverneleodes) leptoscelis Triplehorn, 1975
_________________________________________________________________________________
Clear Creek, 4 km up from Colorado River confluence
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
Clear Creek Canyon
162
Agave phillipsiana Hodgson, 2001
_________________________________________________________________________________
Deer Creek Canyon
162
Agave phillipsiana Hodgson, 2001
_________________________________________________________________________________
Garden Creek (mouth)
= Colorado River Mile 88.9 Left
281
Sonora semiannulata gloydi Stickel, 1938
_________________________________________________________________________________
Grandview Trail
99
246
Physconia isidiomuscigena Esslinger, 2000
Fraxinus cuspidata var. macropetala Rehder, 1917
_________________________________________________________________________________
Grandview Trail, 2000 feet below rim
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
156
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Hance Trail, Red Canyon [not Old Hance Trail]
143
246
259
279
Ptelea lutescens Green, 1906
Fraxinus cuspidata var. macropetala Rehder, 1917
Penstemon coccinatus Rydberg, 1909
Allionia pachyphylla Standley, 1909
_________________________________________________________________________________
Havasu Canyon, 4 miles down “Supai Trail”
[The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.]
75
Encelia resinifera subsp. tenuifolia Curtis, 1998
_________________________________________________________________________________
Havasu Canyon, near Navajo Falls
[The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.]
77
Sclerocactus havasupaiensis Clover, 1942
_________________________________________________________________________________
Havasu Canyon, on top of Supai formation
[The precise gathering locality may be within the boundaries of the Havasupai Indian Reservation.]
77
78
Sclerocactus havasupaiensis Clover, 1942
Sclerocactus havasupaiensis var. roseus Clover, 1942
_________________________________________________________________________________
Hermit Creek
263
283
Populus fremontii var. toumeyi Sargent, 1919
Amsinckia nana Suksdorf, 1931
_________________________________________________________________________________
Hermit Creek (mouth)
35
Astragalus bryantii Barneby, 1944
_________________________________________________________________________________
Hermit Trail
94
246
Oreocarya capitata Eastwood, 1937
Fraxinus cuspidata var. macropetala Rehder, 1917
_________________________________________________________________________________
157
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Indian Garden [Havasupai Gardens]
10
114
167
290
Salix lutea var. ligulifolia C. Ball, 1921
Arctosa mokiensis Gertsch, 1934a
Arenivaga impensa Hopkins, 2014
Perdita wheeleri Timberlake, 1928
_________________________________________________________________________________
Indian Garden (just above) [Havasupai Gardens]
315
Pheidole desertorum var. maricopa Wheeler, 1906
_________________________________________________________________________________
Indian Garden (vicinity) [Havasupai Gardens]
see also “Bright Angel Trail, near Indian Garden”
281
Sonora semiannulata gloydi Stickel, 1938
_________________________________________________________________________________
Manzanita Canyon, within ¼ mile of Bright Angel Creek and North Kaibab Trail, UTM 12S
4005015 N, 407532 E
267
Mentzelia canyonensis Schenk, Hodgson & Hufford, 2013
_________________________________________________________________________________
North Kaibab Trail
36
94
276
Astragalus lentiginosus Dougl. var. oropedii Barneby, 1945
Oreocarya capitata Eastwood, 1937
Carex curatorum Stacey, 1937
_________________________________________________________________________________
Old Hance Trail [not Hance Trail, Red Canyon]
[the published type localities for these taxa are more precisely identified herein as Old Hance Trail;
see main entries in bibliography]
84
144
218
219
279
280
294
320
Ostrya knowltonii Coville, 1894
Ptelea elegans Greene, 1906
Hesperomys megalotis Merriam, 1890
Spilogale gracilis Merriam, 1890
Allionia pachyphylla Standley, 1909
Helix (Arionta) coloradoensis Stearns, 1890
Microsciasma minuta Townsend, 1915
Cicindela rufiventris var. arizonae Wickham, 1899
_________________________________________________________________________________
158
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Phantom Ranch
34
69
183
Paratriatoma hirsuta Barber, 1938
Aphonopelma phasmus Chamberlin, 1940
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
Plateau Point Trail
204
209
Eimeria perognathi Levine, Ivens & Kruidenier, 1957a
Eimeria eremici Levine, Ivens & Kruidenier, 1957a
_________________________________________________________________________________
Rampart Cave [interior]
230
Eschatomoxys pholeter Thomas & Pape in Pape, Thomas & Aalbu, 2007
_________________________________________________________________________________
Rampart Cave (just west of and below)
222
Sonorella reederi W. Miller, 1984
_________________________________________________________________________________
Roaring Springs
45
Papilio indra kaibabensis Bauer, 1955
_________________________________________________________________________________
Roaring Springs Cave [interior]
232
Ptomaphagus cocytus Peck, 1973
_________________________________________________________________________________
Roaring Springs Cave, ca. 1.6 km inside
302
Stygobromus blinni Wang & Holsinger, 2001
_________________________________________________________________________________
Roaring Springs Power Plant
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
Shinumo Creek
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
159
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Shinumo Creek, 3 km from confluence with Colorado River
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
South Bass Trail, about 200 feet below the rim
237
239
Oreohelix yavapai extremitatis Pilsbry & Ferriss, 1911
Pupilla syngenes avus Pilsbry & Ferriss, 1911
_________________________________________________________________________________
South Bass Trail (“the head of a recess in the cross-bed sandstone [Coconino Sandstone]
south of where the Mystic Spring or Bass Trail [South Bass Trail] zigzags down, in a talus
resting on the red sandstone forming the Le Conte Plateau”)
[Pilsbry & Ferris 1911 also illustrate the appearance of the locality in a sketch]
236
Oreohelix yavapai profundorum Pilsbry & Ferriss, 1911
_________________________________________________________________________________
South Bass Trail, in Supai formation
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
South Bass Trail, in Redwall Formation
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
South Bass Trail, base of Tapeats Formation
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
South Kaibab Trail, 5300 feet elevation, in debris slides and crevices of Redwall Limestone
244
Oenothera specuicola subsp. specuicola Raven, 1962
_________________________________________________________________________________
South Kaibab Trail, 36° 03′ 45″ N, 112° 03′ 30″ W
99
Physconia isidiomuscigena Esslinger, 2000
_________________________________________________________________________________
Tanner Trail
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
160
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
Thunder River Cave [interior]
117
Loxosceles kaiba Gertsch & Ennik, 1983
_________________________________________________________________________________
Tonto Plateau, near Bright Angel Trail
183
Crotalus confluentus abyssus Klauber, 1930
_________________________________________________________________________________
Tse’an Cho [interior]
[cave on Horseshoe Mesa]
295
Eleodes (Caverneleodes) leptoscelis Triplehorn, 1975
_________________________________________________________________________________
Tse’an Olje [interior]
[cave in Cremation Canyon]
295
Eleodes (Caverneleodes) leptoscelis Triplehorn, 1975
_________________________________________________________________________________
COLORADO RIVER LOCALES
[Grand Canyon National Park]
Colorado River [without specific locale]
213
Chionaspis gilli Liu & Kosztarab, 1987
_________________________________________________________________________________
Marble Canyon, along the Colorado River, 12 river miles below Lees Ferry, Salt Water Wash
163
Euphorbia aaronrossii A. Holmgren & N. Holmgren, 1988
_________________________________________________________________________________
River Mile 23.0 Left
240
Schinia immaculata Pogue, 2004
_________________________________________________________________________________
Vasey’s Paradise [River Mile 31.8 Right]
273
Wiedemannia digna Sinclair, 2006
_________________________________________________________________________________
161
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
River Mile 37.3 Left
168
Arenivaga pagana Hopkins, 2014
_________________________________________________________________________________
River Mile 53 Right, Nankoweap
168
Arenivaga pagana Hopkins, 2014
_________________________________________________________________________________
River Mile 58.2 Right, Awatubi Canyon
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
Colorado River, at or near mouth of Bright Angel Creek
[Bright Angel Creek confluence is at River Mile 87.7 Right]
221
Gila cypha R. Miller, 1946
_________________________________________________________________________________
River Mile 88.9 Left see “Garden Creek (mouth)” under “Inner Canyon” listings, above
_________________________________________________________________________________
River Mile 92.3 Left
181
Nebria (Reductonebria) georgei Kavanaugh, 2008
_________________________________________________________________________________
River Mile 108 Right, Shinumo Creek (mouth)
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
River Mile 108 Left
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
River Mile 147.8 Left, Matkatamiba Canyon
286
Flaveria mcdougallii Theroux, Pinkava & Keil, 1977
_________________________________________________________________________________
River Mile 158 Right (campsite near), 339451 E, 4018170 N, NAD 83, 36°17′42.6″ N;
112°47′16.6″ W
266
Mentzelia hualapaiensis Schenk, Hodgson & Hufford, 2010
_________________________________________________________________________________
162
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
River Mile 160.5 Left
166
Arenivaga grandiscanyonensis Hopkins, 2014
_________________________________________________________________________________
Note: Localities between River Miles 164.7 and 273.3, on the left,
could be within the boundaries of the Hualapai Indian Reservation
_________________________________________________________________________________
River Mile 166.5 Left
240
Schinia immaculata Pogue, 2004
_________________________________________________________________________________
River Mile 174.2 Right, Cove Canyon
286
Flaveria mcdougallii Theroux, Pinkava & Keil, 1977
_________________________________________________________________________________
River Mile 180.8 Right
181
Nebria (Reductonebria) georgei Kavanaugh, 2008
_________________________________________________________________________________
River Mile 186.5 Left
168
Arenivaga pagana Hopkins, 2014
_________________________________________________________________________________
River Mile 198.0 Right
168
Arenivaga pagana Hopkins, 2014
_________________________________________________________________________________
River Mile 202.5 Left
168
Arenivaga pagana Hopkins, 2014
_________________________________________________________________________________
River Mile 202 Right
240
Schinia immaculata Pogue, 2004
_________________________________________________________________________________
River Mile 211.5 Right
166
Arenivaga grandiscanyonensis Hopkins, 2014
_________________________________________________________________________________
163
LOCALITIES FOR TYPE MATERIAL FROM
GRAND CANYON NATIONAL PARK
164
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
LOCALITIES FOR TYPE MATERIAL FROM NEAR
GRAND CANYON NATIONAL PARK
See also various headings within this list for further refinements
[In the PDF document the lines immediately below are hyperlinks]
page
Cameron, Arizona, and vicinity
166
Arizona Strip
167
Grand Canyon vicinity undiscriminated
171
Grand Canyon–Parashant National Monument
172
Havasupai Indian Reservation
174
House Rock and House Rock Valley
174
Hualapai Indian Reservation
175
Hualapai Indian Reservation vicinity (off reservation)
177
“Kaibab Forest” (north) undiscriminated
177
“Kaibab Forest” (south) undiscriminated
177
Kaibab Indian Reservation
178
Kaibab National Forest, Tusayan Ranger District
178
Kaibab Plateau (including Kaibab National Forest, North Kaibab Ranger District)
179
Kanab Canyon
184
Lees Ferry
184
Marble Canyon [not on Colorado River]
184
Navajo Nation (westernmost)
185
Red Mountain
185
165
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Locality data are as described in the original publications; thus some minor variances will be
noted for specific sites or areas that are otherwise in very close proximity to each other.
Refinements or special notes appear within [non-bold square brackets].
Red Numbers indicate and cross-list the sequentially enumerated taxa in the bibliography
(thus they reflect the alphabetical order by publication author).
Some taxa listed here may have been collected from the physiographic Grand Canyon, but
outside of the national park boundaries.
Note: Baral et al. (2020) did not demonstrate a precise understanding of the geography of
the Grand Canyon region. All of their localities are listed as being in Arizona (a few actually
are in Utah), but some locality references to “Grand Canyon” are, confusingly, neither within
the physiographic canyon nor within the national park but are on adjacent national forest
and other lands, sometimes at quite some distance from the canyon or the national park.
Some interpretive statements have been included in this locality list in order to clarify these
discrepancies; specifically, measurements have been made from a map that reveal that linear
distances from geographic locales are mostly along the principal highways of the area, which
information is added notationally to the original locality data.
CAMERON, ARIZONA
and vicinity
Cameron
80
Myrmecocystus melliger subsp. semirufus Emery var. romainei Cole, 1936 [not valid as a
196
Aphricus knowltoni Knull, 1957
quadrinomial taxon]
_________________________________________________________________________________
Cameron Trading Post
2
Plagiomimicus kathyae J. Adams & Lafontaine, 2009
_________________________________________________________________________________
Little Colorado River (bed of) at Cameron
38
Astragalus wootoni var. endopterus Barneby, 1949
_________________________________________________________________________________
166
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Cameron (15 miles W of, along the canyon of the Little Colorado River)
[quite probably at or near the Little Colorado River Gorge Overlook, along Arizona Route 64]
278
Megathymus alliae Stallings & Turner, 1957
_________________________________________________________________________________
ARIZONA STRIP
9
“Northern Arizona” [general vicinity nearest to Kanab, Utah; without specific locale]
303
304
305
306
307
308
309
Dalea amoena Watson, 1873
Oenothera (Chylisma) multijuga Watson, 1873
Cymopterus purpureus Watson, 1873
Peucedanum newberryi Watson, 1873
Brickellia (Clavigera) longifolia Watson, 1873
Calochortus flexuosus Watson, 1873
Androstephium breviflorum Watson, 1873
_________________________________________________________________________________
Arizona Route 389, 4.3 mi west of U.S. Route 89A
[near Fredonia, Arizona]
251
Eriogonum thompsonae S. Wats. var. atwoodii Reveal, 1974b
_________________________________________________________________________________
Arizona Route 389, 4.5 mi southwest of U.S. Route 89A
[near Fredonia, Arizona]
250
Eriogonum mortonianum Reveal, 1974a
_________________________________________________________________________________
Bulrush Canyon (northwest of, south of Pipe Spring)
231
Opuntia pinkavae Parfitt, 1997
_________________________________________________________________________________
Colorado City (6 miles east of; Rosy Canyon Road, 1.5 miles south of Utah state line)
173
Trogloderus skillmani Johnston, 2019
_________________________________________________________________________________
9
Refer also separately in this list to: Grand Canyon–Parashant National Monument; House Rock
Valley; Kaibab Indian Reservation; Kaibab National Forest; Kaibab Plateau; Kanab Canyon; Lees
Ferry.
167
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Fredonia, Arizona
Athysanella fredonia Ball & Beamer, 1939
14
_________________________________________________________________________________
Fredonia (1 mile north of)
231
Opuntia pinkavae Parfitt, 1997
_________________________________________________________________________________
Hack Canyon (vicinity)
Phacelia higginsii Atwood, 2007
6
_________________________________________________________________________________
“Hell Canyon”
[see bibliography for comments on the locality]
148
Morus grisea Greene, 1911
_________________________________________________________________________________
Kaibab Indian Reservation (west of)
231
Opuntia pinkavae Parfitt, 1997
_________________________________________________________________________________
Millipede Cave [interior]
[Bureau of Land Management lands]
272
Pratherodesmus voylesi Shear in Shear, Taylor, Wynne & Krejca, 2009
_________________________________________________________________________________
Mokiak Pass
[20 miles southeast of St. George, Utah
125
126
127
128
129
269
310
10
]
Astragalus subcinereus A. Gray, 1878
Astragalus scaposus A. Gray, 1878
Astragalus mokiacensis A. Gray, 1878
Astragalus artipes A. Gray, 1878
Actinella biennis A. Gray, 1878
Synchloe thoosa Scudder, 1878
Lesquerella arizonica Watson, 1888
_________________________________________________________________________________
10
Location of Mokiak Pass is as identified by S. H. Scudder (1878, Notice of the butterflies collected
by Dr. Edward Palmer in the arid regions of southern Utah and northern Arizona during the
summer of 1877, U.S. Geological and Geographical Survey of the Territories, Bulletin, 4).
168
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Main Street Valley, ca. 1.5 km south of the junction of the Colorado City road with the Main
Street Valley road
245
Panicum mohavense Reeder, 1991
_________________________________________________________________________________
Main Street Valley, west of Hurricane Cliffs
231
Opuntia pinkavae Parfitt, 1997
_________________________________________________________________________________
Navaho Trail, near Hurricane Rim
231
Opuntia pinkavae Parfitt, 1997
_________________________________________________________________________________
North Canyon (Kaibab Plateau, East Rim View Point along Trail #7 ca. 2¼ mi down from
trailhead, in North Canyon; 36°25′18″N, 112°04′22″W)
252
Scutellaria potosina var. kaibabensis Rhodes & Ayers, 2010
_________________________________________________________________________________
North Canyon Trail #4, 36°25′04″N 112°04′17″W
[east rim of Kaibab Plateau]
252
Scutellaria potosina var. kaibabensis Rhodes & Ayers, 2010
_________________________________________________________________________________
October Gyp Cave [interior]
[Bureau of Land Management lands]
272
Pratherodesmus voylesi Shear in Shear, Taylor, Wynne & Krejca, 2009
_________________________________________________________________________________
Page (9 miles northwest of)
228
Acmaeodera navajo G. Nelson & Westcott, 1995
_________________________________________________________________________________
Pipe Spring (ca. 9 miles east-northeast of)
see also Kaibab Indian Reservation
250
Eriogonum mortonianum Reveal, 1974a
_________________________________________________________________________________
169
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Uinkaret Plateau, just north of the Navajo Trail Road, 1.1 km W of the Clayhole Wash
crossing, 27.5 km air distance south-southweest of Colorado City, 36°49′09″N,
113°11′21″W, T40N R8W S31
165
Mentzelia memorabilis N. Holmgren & P. Holmgren, 2002
_________________________________________________________________________________
Vermilion Cliffs [geographical feature or place name undiscriminated11]
228
Acmaeodera navajo G. Nelson & Westcott, 1995
_________________________________________________________________________________
Vermilion Lodge (vicinity) [Vermilion Cliffs]
282
Atriplex asterocarpa Stutz, Chu & Sanderson, 1994
_________________________________________________________________________________
Virgin Mountains
139
Ptelea straminea Green, 1906
_________________________________________________________________________________
Wolf Hole
118
Dipodomys microps celsus Goldman, 1924
_________________________________________________________________________________
Wolf Hole (4.2 miles southwest of)
231
Opuntia pinkavae Parfitt, 1997
_________________________________________________________________________________
Wolf Hole (20 miles south of)
124
Thomomys bottae nicholi Goldman, 1938
_________________________________________________________________________________
11
In their paper, Nelson & Westcott also specially mention “Cliff Dwellers Lodge”, which is not far
from another business by the name of Vermilion Cliffs, thus it is unclear whether “Vermilion Cliffs”
refers to that place, or to the natural geographical feature Vermilion Cliffs that align in this area
along U.S. Route 89 Alternate.
170
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
GRAND CANYON VICINITY 12
undiscriminated
“about Grand Canyon” or “region of the Grand Canyon”
89
145
254
Galium munzii Hilend & Howell subsp. ambivalens Dempster & Ehrendorfer, 1965
Ptelea confinis Greene, 1906
Silene rectiramea Robinson, 1899
_________________________________________________________________________________
Grand Canyon (“near”)
12
56
57
194
Hysteropterum cornutum var. utahnum E. Ball, 1935
Trichochrous incipiens Casey, 1895
Trichochrous reversus Casey, 1895
Bolteria juniperi Knight, 1968
_________________________________________________________________________________
Grand Canyon, Ariz. (“vicinity of”)
Aphelonema convergens var. canyonensia Bunn, 1930
54
_________________________________________________________________________________
Grand Canyon (south of)
249
Fabriciella evanida Reinhard, 1953
_________________________________________________________________________________
Grand Canyon (30 miles south of)
277
Vaejovis aquilonalis Stahnke, 1940
_________________________________________________________________________________
12
It is not clear whether materials collected with the indefinite locality descriptions here include
among all specimens many, several, or one that may have come from the Grand Canyon itself.
Because of this uncertainty, the taxa listed here are included in the “near Grand Canyon National
Park” list of type specimens rather than in the national park list itself.
171
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
GRAND CANYON–PARASHANT NATIONAL MONUMENT 13
12.5 miles southwest from head of Trail Canyon toward Andrus Canyon, 36°11.48′N,
113°19.10′W
Phacelia higginsii Atwood, 2007
6
_________________________________________________________________________________
Andrus Canyon, 1 mile west of Andrus Point, T32N R10W S10
311
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001
_________________________________________________________________________________
Andrus Canyon, 3 miles west of Andrus Point, T32N R10W S6, wash bottom
311
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001
_________________________________________________________________________________
Andrus Canyon at road crossing, T32N, R10W, S6 & 7
Phacelia furnissii Atwood, 2007
5
_________________________________________________________________________________
Cunningham Canyon
Xylorhiza tortifolia (Torrey & Gray) Greene var. parashantensis Atwood & Welsh, 2013
8
_________________________________________________________________________________
Daneill Canyon, west of Andrus Canyon, 3 miles west of Andrus Pt., T32N R11W S12
311
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001
_________________________________________________________________________________
Eldel Cave [interior]
230
Eschatomoxys pholeter Thomas & Pape in Pape, Thomas & Aalbu, 2007
_________________________________________________________________________________
13
Localities are as determined based on boundaries at the time of publication of this checklist. Prior
to the creation of Grand Canyon–Parashant National Monument (in 2000), and depending upon
original dates of collection, some material may originally be said to have been from Grand Canyon
National Monument, Lake Mead National Recreation Area (that is, its northeastern section), or the
Arizona Strip generally.
172
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Mount Trumbull, 3 miles south of Nixon Spring
[The locality, “three miles south of Nixon Spring”, is just to the south of Mount Trumbull itself, today
within the Grand Canyon–Parashant National Monument, about 2¾ miles north of the boundary of
Grand Canyon National Park.]
151
Thomomys bottae trumbullensis Hall & Davis, 1934
_________________________________________________________________________________
Mt. Trumbull school house (2 miles south of ), near Griffith Knoll, T34S R10W S1-2
311
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001
_________________________________________________________________________________
PARA-1001 Cave, ca. UTM 0264500 N, 4060700 E [interior]
153
154
233
Hesperochernes bradybaughi Harvey & Wynne, 2014
Tuberochernes cohni Harvey & Wynne, 2014
Ptomaphagus parashant Peck & Wynne, 2013
_________________________________________________________________________________
Parashant (Trail) Canyon, 11 miles south of Mount Trumbull Village
311
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001
_________________________________________________________________________________
Poverty Mountain (south base)
7
Phacelia hughesii Atwood, 2007
_________________________________________________________________________________
Poverty Mountain (southeast of), southwest of Mt. Trumbull, T34N, R11W, Sec 4
312
Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010
_________________________________________________________________________________
Poverty Mountain (southwest of) near Dewdrop Spring, T35N, R12W, Sec. 35/36
312
Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010
_________________________________________________________________________________
Poverty Mountain (BLM road south of) and southwest of Salt Spring, T34N, R11W [sic]
312
Pediomelum pauperitense S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010
_________________________________________________________________________________
Sandstone Canyon (1.5 mile due west of Cane Spring)
8
Xylorhiza tortifolia (Torrey & Gray) Greene var. parashantensis Atwood & Welsh, 2013
_________________________________________________________________________________
173
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Twin Point (west rim of) (18 km south of Oak Grove, T30N R12W NW¼ sect. 7, 36°01′N,
113°37′W)
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
_________________________________________________________________________________
Whitmore Canyon, T32N R9W S1
311
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001
_________________________________________________________________________________
Whitmore Canyon, 1.25 km (7.8 miles) [sic] south of Mount Trumbull (abandoned town),
T34N R9W S29
311
Astragalus lentiginosus Douglas ex Hook. var. trumbullensis Welsh & Atwood, 2001
_________________________________________________________________________________
HAVASUPAI INDIAN RESERVATION
See also entries under “Havasu” in the LOCALITIES FOR TYPE MATERIAL IN GRAND CANYON
NATIONAL PARK list noted as possibly being within the Havasupai Reservation. These localities are
within the physiographic Grand Canyon.
“Canyon at Supai, Coconino County, Ariz.”
214
Oreohelix yavapai vauxae Marshall, 1929
_________________________________________________________________________________
Mooney Falls, near
244a Oenothera specuicola subsp. hesperia Raven, 1962
_________________________________________________________________________________
Supai, Havasu Canyon
293
Perdita depressa Timberlake, 1968
_________________________________________________________________________________
HOUSE ROCK and HOUSE ROCK VALLEY
House Rock, near Lees Ferry
180
Astragalus kaibensis Jones, 1902
_________________________________________________________________________________
174
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
House Rock Valley [without specific locale]
49
74
161
Pediocactus paradinei Benson, 1957
Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986
Sclerocactus whipplei (Englemann & Bigelow) Britton & Rose ssp. busekii Hochstätter, 1995
_________________________________________________________________________________
House Rock Valley, north of Rock Canyon
231
Opuntia pinkavae Parfitt, 1997
_________________________________________________________________________________
45 km southwest of Page, 26 km southwest of Marble Canyon
[House Rock valley vicinity]
18
Orbilia ocellata Baral, G. Marson & E. Weber in Baral, Weber & Marson, 2020
_________________________________________________________________________________
HUALAPAI INDIAN RESERVATION
These localities, with the exception perhaps of some collections from the “vicinity” of, or southerly
from, Peach Springs are within the physiographic Grand Canyon.
Diamond Creek Canyon
135
Arabis recondita Greene, 1900
_________________________________________________________________________________
Diamond River [Diamond Creek]
131
Phacelia (Coreanthus) ivesiana A. Gray in Gray et al., 1861
_________________________________________________________________________________
Diamond River [Diamond Creek] (mouth of)
132
“Gilia dactylophyllum, (n. sp.?)”, A. Gray in Gray et al., 1861 [see remarks in bibliography]
_________________________________________________________________________________
Grand Cañon of the Colorado, below Peach Springs
130
Mirabilis bigelovii A. Gray, 1886
_________________________________________________________________________________
175
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Peach Springs
169
247
274
310
Luperodes wickhami Horn, 1893
Fraxinus lowellii Sargent in Rehder, 1917
Eriogonum fusiforme Small, 1906
Lesquerella arizonica Watson, 1888
_________________________________________________________________________________
Peach Springs (above)
138
Ptelea palida Green, 1906
_________________________________________________________________________________
Peach Springs (vicinity)
82
83
134
147
Echinocactus polycephalus var. xeranthemoides Coulter, 1896
Phellopterus multinervatus Coulter & Rose, 1900
Arabis eremophila Greene, 1900
Senecio stygius Greene, 1909 [type locality for this species refined here; see main entry in
bibliography]
261
Pediomelum retrorsum Rydberg in Rydberg & Pennell, 1919
_________________________________________________________________________________
Peach Springs Canyon (Peach Springs Wash)
159
255
Opuntia abyssi Hester, 1943
Crossosoma parviflora B. Robinson & Fernald, 1894 [type locality for this species refined here;
see entry in bibliography]
284
Amsinckia macrosepala Suksdorf, 1931 [type locality for this species refined here; see entry in
bibliography]
_________________________________________________________________________________
Peach Springs Wash, 4 km from confluence with Colorado River
264
Efferia tapeats Scarbrough, Stevens & Nelson, 2012
_________________________________________________________________________________
Prospect Valley (lower end)
120
Ammospermophilus leucurus tersus Goldman, 1929
_________________________________________________________________________________
Upper Peach Springs, outflow just below concrete weir, 35.55786°N, 113.4314°W
157
Pyrgulopsis hualapaiensis Herschler, Liu & Stevens, 2016
_________________________________________________________________________________
176
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
HUALAPAI INDIAN RESERVATION vicinity (off reservation)
Peach Springs (8 miles southwest of) [vicinity of Truxton, Arizona]
150
Boharticus margaretae Grissell, 1983
_________________________________________________________________________________
U.S.-Arizona Highway No. 66, 11 or 12 miles east of Peach Springs and 2 or 3 miles west of
Hyde Park, Arizona
[“Hyde Park”, near mile marker 117 on historic Route 66, no longer exists; the gathering locality is
near the present-day Grand Canyon Caverns tourist attraction.]
158
Opuntia hualpaensis Hester, 1943
_________________________________________________________________________________
“KAIBAB FOREST” (north, undiscriminated)
[see also “Kaibab Plateau”]
“Kaibab forest, near the north rim of the Grand Canyon”
115
Phidippus kaibabensis Gertsch, 1934b
_________________________________________________________________________________
“KAIBAB FOREST” (south, undiscriminated)
[see also “Kaibab National Forest, Tusayan Ranger District”]
“W 112°:N 36°, Kaibab Forest, Arizona”
[The overly generalized coordinates lie within Grand Canyon National Park, just west of Grandview
Point; the precise intersection is in fact, by coincidence, on a cliff face(!) just over the rim of the
canyon (https://confluence.org/confluence.php?id=91). The specific epithet of the taxon noted here,
and indication of “Kaibab Forest,” may point either to a locality on the nearby Kaibab National Forest
or is only a generalized term for the South Rim forest, sometimes called the Coconino forest, on the
Coconino Plateau.]
71
Neoantistea coconino Chamberlin & Ivie, 1942
_________________________________________________________________________________
177
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
KAIBAB INDIAN RESERVATION
Kanab Wash, at southern boundary of Kaibab Indian Reservation
119
Dipodomys ordii cupidineus Goldman, 1924
_________________________________________________________________________________
Pipe Spring (½ mile north of)
Opuntia aurea Baxter, 1933
46
_________________________________________________________________________________
KAIBAB NATIONAL FOREST, TUSAYAN RANGER DISTRICT
[see also “Kaibab Forest (south, undiscriminted)”]
Arizona Route 64, junction of road to Anita
[road junction coordinates ca. 35.861° N, 112.134° W]
171
Cinara poketa Hottes, 1956
_________________________________________________________________________________
Tusayan (3 km south of)
[locality is on the Kaibab National Forest on Arizona Route 64]14
20
25
28
31
32
Orbilia phanosoma Baral & [G.] Marson in Baral, Weber & Marson, 2020
Orbilia ophiosoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia flexisoma Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia multitrapezoidea Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia calyptrata Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
14
See the LOCALITIES FOR TYPE MATERIAL IN GRAND CANYON NATIONAL PARK list for localities north
of Tusayan that are within Grand Canyon National Park.
178
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
KAIBAB PLATEAU
[also as Buckskin Mountains]
including KAIBAB NATIONAL FOREST, NORTH KAIBAB RANGER DISTRICT
[see also “Kaibab Forest (north, undiscriminated)”]
Kaibab Plateau [without specific locale]
112
Speyeria atlantis schellbachi Garth, 1949
_________________________________________________________________________________
Kaibab National Forest [without specific locale but on North Kaibab Ranger District]
182
Enderleinellus kaibabensis Kim, 1966
_________________________________________________________________________________
Kaibab Plateau, North Rim of Grand Canyon [undiscriminated]
3
Tipula (Lunatipula) kaibabensis C. Alexander, 1946
_________________________________________________________________________________
Kaibab Plateau / Buckskin Mountains [without specific locale]
174
176
Astragalus humistratus var. tenerrimus Jones, 1895
Bigelovia howardi var. attenuata Jones, 1895
_________________________________________________________________________________
Buckskin Mountains (mesa below)
177
Laphamia congesta Jones, 1895
_________________________________________________________________________________
Kaibab Plateau (northwest escarpment slope, 6700 feet)
248
296
Morsea california kaibabensis Rehn & Grant, 1958
Asidina rugicollis Triplehorn & Brown, 1971
_________________________________________________________________________________
Arizona Route 67, 3.2 km north of Grand Canyon National Park boundary, T.34 N., R.3 E.,
Sec. 7
164
Castilleja kaibabensis N. Holmgren, 1973
_________________________________________________________________________________
179
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
DeMotte Park
86
123
164
175
Penstemon virgatus pseudoptutus Crosswhite, 1967
Thomomys fossor kaibabensis Goldman, 1938
Castilleja kaibabensis N. Holmgren, 1973
Oenothera triloba var. ecristata Jones 1895
_________________________________________________________________________________
Dry Park
86
Penstemon virgatus pseudoptutus Crosswhite, 1967
_________________________________________________________________________________
Fredonia (28 km (17.5 miles) airline distance east of), T41N, R2E, S29/30
[an area of Forest Service roads]
39
Astragalus cremnophylax Barneby var. myriorrhaphis Barneby, 1979
_________________________________________________________________________________
Fredonia (28 km east-southeast of Fredonia, 13 km NNW of Jacob Lake)
[along U.S. Route 89A]
17
22
24
26
27
30
Orbilia purshiae Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia arizonensis Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia spermoides Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia multimaeandrina Baral & G. Marson in Baral, Weber & Marson, 2020
Orbilia macrodelphinus Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
Fredonia Sawmill (17 miles southeast of), along logging road to Ryan
313
Astragalus pinionis Welsh & Thorne, 1977
_________________________________________________________________________________
Hwy. from Jacob Lake to Lodge, Kaibab Forest
[Arizona Route 67, between Jacob Lake and Kaibab Lodge]
86
Penstemon virgatus pseudoptutus Crosswhite, 1967
_________________________________________________________________________________
House Rock Canyon to Jacob Lake
164
Castilleja kaibabensis N. Holmgren, 1973
_________________________________________________________________________________
180
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Jacob Lake
223
287
288
Saxinis saucia kaibabiae Moldenke, 1970
Boletus barrowsii Thiers & Smith in Thiers, 1976
Suillus kaibabensis Thiers, 1976
_________________________________________________________________________________
Jacob Lake (near)
74
297
Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986
Hadena (Hadena) lafontainei Troubridge & Crabo, 2002
_________________________________________________________________________________
Jacob Lake (northwest of)
108
Coryphantha vivipara var. kaibabensis Fischer, 1979
_________________________________________________________________________________
Jacob Lake (7.5 miles east of)
74
Cordylanthus wrightii kaibabensis Chuang & Heckard, 1986
_________________________________________________________________________________
Jacob Lake (south of), 37 km southeast of Fredonia
[on Arizona Route 67]
19
Orbilia magnifica Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
Jacob Lake (18.6 miles south of, on Arizona Route 67)
258
Lesquerella kaibabensis Rollins, 1982
_________________________________________________________________________________
Jacob Lake (20 miles south of)
253
Andrena (Scaphandrena) kaibabensis Ribble, 1974
_________________________________________________________________________________
Kaibab Forest, near Grand Canyon National Park
86
Penstemon virgatus pseudoptutus Crosswhite, 1967
_________________________________________________________________________________
Kaibab Lodge (3 km north of)
[on Arizona Route 67]
26
Orbilia maeandrina Baral & G. Marson in Baral, Weber & Marson, 2020
_________________________________________________________________________________
181
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Kaibab Lodge (4 miles north of)
172
Cicindela pusilla kaibabensis Johnson, 1990
_________________________________________________________________________________
Kaibab Lodge (8 miles north of)
172
Cicindela pusilla kaibabensis Johnson, 1990
_________________________________________________________________________________
Kaibab National Forest, 0.6 km east of Arizona Highway 67 just northeast of Forest Road
213 (36°30′38″N, 112°07′55″W)
[FR 213 intersects AZ 67 30.6 km south of Jacob Lake]
48
Branchinecta kaibabensis Belk & Fugate, 2000
_________________________________________________________________________________
Kaibab National Forest, Forest Road 293, 36.40806° N, 112.26692° W
[ca. 7.3 miles west of Kaibab Lodge]
42
Morchella kaibabensis Beug, T.A. Clem. & T.J. Baroni in Baroni et al., 2018
_________________________________________________________________________________
Lookout Canyon Road (422D), 5.6 km from (northwest of) Grand Canyon Highway (Arizona
Route 67), T.35 N., R.2 E., Sec. 15
164
Castilleja kaibabensis N. Holmgren, 1973
_________________________________________________________________________________
Marble Flat
86
Penstemon virgatus pseudoptutus Crosswhite, 1967
_________________________________________________________________________________
Marble Flat, just north of North Rim Ranger Station
164
Castilleja kaibabensis N. Holmgren, 1973
_________________________________________________________________________________
182
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
Nagle’s Ranch15
146
Ptelea saligna Green, 1906
_________________________________________________________________________________
Nagle’s Ranch15 (below, on edge of Buckskin Mountains)
178
Laphamia gracilis Jones, 1895
_________________________________________________________________________________
Nagle’s Ranch15 (road to)
174
Astragalus humistratus var. tenerrimus Jones, 1895
_________________________________________________________________________________
Ryan Ranger Station (near)
Papilio indra kaibabensis Bauer, 1955
45
_________________________________________________________________________________
VT Meadows, 6.5 km south of Kaibab Lodge, T.34 N., R.3 E., Sec. 7
164
Castilleja kaibabensis N. Holmgren, 1973
_________________________________________________________________________________
VT Park
86
164
Penstemon virgatus pseudoptutus Crosswhite, 1967
Castilleja kaibabensis N. Holmgren, 1973
_________________________________________________________________________________
VT Ranch
Tapinocyba kesimba Chamberlin, 1948
70
_________________________________________________________________________________
VT Ranger Station
86
164
Penstemon virgatus pseudoptutus Crosswhite, 1967
Castilleja kaibabensis N. Holmgren, 1973
_________________________________________________________________________________
15
“Nagle’s Ranch was about 60 miles south of Kanab, Utah, on the west slope of the Buckskin
Mountains (i. e. the Kaibab Plateau), and was the first watering place on the old wagon road to the
Grand Canyon from Kanab. [. . .] The present wagon road now ascends the plateau many miles
farther north.” (Kearney, Thomas H., and Peebles, Robert H. (and collaborators), 1942, Flowering
plants and ferns of Arizona, U.S. Department of Agriculture, Miscellaneous Publication 423, p. 986.)
183
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
KANAB CANYON
Kanab Canyon (west rim, south of Water Canyon, in depression caused by breccia pipe
collapse, T38N R3W NW¼ SW¼ sect. 8)
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
_________________________________________________________________________________
Kanab Canyon (west rim, in small drainage 30 m from edge, T38N R3W NW¼ sect. 29
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
_________________________________________________________________________________
Kanab Canyon (west rim, T38N R3W sect. 30)
235
Rosa stellata Wooton subsp. abyssa Phillips, 1992
_________________________________________________________________________________
Kanab Wash “near the Colorado” [Arizona]
82
Echinocactus polycephalus var. xeranthemoides Coulter, 1896
_________________________________________________________________________________
LEES FERRY
Lees Ferry
179
260
282
Astragalus kentrophyta var. coloradoensis Jones, 1902
Amsonia eastwoodiana Rydberg, 1913
Atriplex asterocarpa Stutz, Chu & Sanderson, 1994
_________________________________________________________________________________
Lees Ferry (near)
291
Perdita opacella Timberlake, 1956
_________________________________________________________________________________
MARBLE CANYON
Navajo Bridge
11
13
Myndus yuccandus E. Ball, 1933
Yucanda miniata E. Ball, 1937
_________________________________________________________________________________
184
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
NAVAJO NATION (westernmost)
See also entries for Cameron (above). See also “Cedar Mountain” in the LOCALITIES FOR TYPE
MATERIAL IN GRAND CANYON NATIONAL PARK list, noted as possibly being within the boundaries of
the Navajo Nation.
The Gap
116
Steatoda variata Gertsch, 1960
_________________________________________________________________________________
The Gap (near)
160
Cryptantha atwoodii Higgins, 1974
_________________________________________________________________________________
RED MOUNTAIN
Red Mountain, 35 miles northwest of Flagstaff on U.S. Route 180
73
Mentzelia collomiae Christy, 1997
_________________________________________________________________________________
185
LOCALITIES FOR TYPE MATERIAL FROM
NEAR GRAND CANYON NATIONAL PARK
186
INDEX TO NOMINATE TAXA
INDEX TO NOMINATE TAXA
Names in bold indicate nominate taxa for which at least one locality reported for its types is
within, or is with good probability from within, the present boundaries of Grand Canyon
National Park.
The names are original combinations, as published by the authors. The taxonomic phylum is
appended to each name and author. Refer to the bibliography and annotated checklist for
details.
Red numbers at left indicate and cross-list the sequentially enumerated taxa in the bibliography (thus they reflect the alphabetical order by publication author).
163
235
159
183
198
59
278
89
241
303
238
64
87
47
277
141
320
149
100
22
95
310
43
256
128
216
aaronrossii, Euphorbia A. Holmgren & N. Holmgren, 1988 (Trachyophyta)
abyssa, Rosa stellata Wooton subsp. Phillips, 1992 (Trachyophyta)
abyssi, Opuntia Hester, 1943 (Trachyophyta)
abyssus, Crotalus confluentus Klauber, 1930 (Chordata)
ackerti, Aspiculuris Kruidenier & Mehra, 1959 (Nematoda)
aequalis, Orphilus Casey, 1900 (Arthropoda)
alliae, Megathymus Stallings & Turner, 1957 (Arthropoda)
ambivalens, Galium munzii Hilend & Howell subsp. Dempster & Ehrendorfer, 1965 (Trachyophyta)
americanus, Pilophorus Poppius, 1914 (Arthropoda)
amoena, Dalea Watson, 1873 (Trachyophyta)
angelica, Oreohelix yavapai Pilsbry & Ferriss, 1911 (Mollusca)
angustulus, Prionus Casey, 1912 (Arthropoda)
apache, Tibicen Davis, 1921 (Arthropoda)
apicata, Delphacodes Beamer, 1948 (Arthropoda)
aquilonalis, Vaejovis Stahnke, 1940 (Arthropoda)
argentea, Ptelea Green, 1906 (Trachyophyta)
arizonae, Cicindela rufiventris var. Wickham, 1899 (Arthropoda)
arizonense, Penicillium Frisvad, Grijseels & J.C. Nielsen in Grijseels et al., 2016 (Ascomycota)
arizonense, Petalium bistriatum Say var. Fall, 1905 (Arthropoda)
arizonensis, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
arizonensis, Ripersia Ehrhorn, 1899 (Arthropoda)
arizonica, Lesquerella Watson, 1888 (Trachyophyta)
arizonica, Protoventuria M. Barr, 1989 (Ascomycota)
arizonicus, Chrysotimus H. Robinson, 1967 (Arthropoda)
artipes, Astragalus A. Gray, 1878 (Trachyophyta)
artuflava, Chelinidea vittiger Uhler var. McAtee, 1919 (Arthropoda)
187
INDEX TO NOMINATE TAXA
282
176
160
251
46
239
asterocarpa, Atriplex Stutz, Chu & Sanderson, 1994 (Trachyophyta)
265
287
88
68
53
156
129
130
302
91
153
309
62
35
185
90
barberi, Ochterus Schell, 1943 (Arthropoda)
32
270
54
267
94
16
314
225
118
98
136
319
187
259
71
232
154
73
113
179
calyptrata, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
attenuata, Bigelovia howardi var. Jones, 1895 (Trachyophyta)
atwoodii, Cryptantha Higgins, 1974 (Trachyophyta)
atwoodii, Eriogonum thompsonae S. Wats. var.
Reveal, 1974b (Trachyophyta)
aurea, Opuntia Baxter, 1933 (Trachyophyta)
avus, Pupilla syngenes Pilsbry & Ferriss, 1911 (Mollusca)
barrowsii, Boletus Thiers & Smith in Thiers, 1976 (Basidiomycota)
basala, Ballana DeLong, 1964 (Arthropoda)
behlei, Aphonopelma Chamberlin, 1940 (Arthropoda)
benedicti, Erax Bromley, 1940 (Arthropoda)
betheli, Sonorella Henderson, 1914 (Mollusca)
biennis, Actinella A. Gray, 1878 (Trachyophyta)
bigelovii, Mirabilis A. Gray, 1886 (Trachyophyta)
blinni, Stygobromus Wang & Holsinger, 2001 (Arthropoda)
boreorarius, Thomomys bottae Durham, 1952 (Chordata)
bradybaughi, Hesperochernes Harvey & Wynne, 2014 (Arthropoda)
breviflorum, Androstephium Watson, 1873 (Trachyophyta)
brevipennis, Discodemus Casey, 1908 (Arthropoda)
bryantii, Astragalus Barneby, 1944 (Trachyophyta)
bullatus, Deraeocoris Knight, 1921 (Arthropoda)
bunni, Bruchomorpha Doering, 1939 (Arthropoda)
canonicus, Melanoplus Scudder, 1897 (Arthropoda)
canyonensia, Aphelonema convergens var. Bunn, 1930 (Arthropoda)
canyonensis, Mentzelia Schenk, Hodgson & Hufford, 2013 (Trachyophyta)
capitata, Oreocarya Eastwood, 1937 (Trachyophyta)
casa, Athysanella (Gladionura) Ball & Beamer, 1939 (Arthropoda)
castus, Polymedon Wheeler, 1899 (Arthropoda)
cavicola, Archeolarca Muchmore, 1981 (Arthropoda)
celsus, Dipodomys microps Goldman, 1924 (Chordata)
circinans, Puccinia J. Ellis & Everhart, 1900 (Basidiomycota)
cissodes, Schmaltzia Greene, 1905 (Trachyophyta)
clarigaster, Camponotus acutirostris Wheeler, 1915 (Arthropoda)
clavicornis, Ceratocapsus Knight, 1925 (Arthropoda)
coccinatus, Penstemon Rydberg, 1909 (Trachyophyta)
coconino, Neoantistea Chamberlin & Ivie, 1942 (Arthropoda)
cocytus, Ptomaphagus Peck, 1973 (Arthropoda)
cohni, Tuberochernes Harvey & Wynne, 2014 (Arthropoda)
collomiae, Mentzelia Christy, 1997 (Trachyophyta)
coloradensis, Misumenops Gertsch, 1933 (Arthropoda)
coloradoensis, Astragalus kentrophyta var. Jones, 1902 (Trachyophyta)
188
INDEX TO NOMINATE TAXA
280
243
145
104
177
193
37
119
276
221
coloradoensis, Helix (Arionta) Stearns, 1890 (Mollusca)
132
41
29
271
111
293
273
300
1
268
226
“dactylophyllum”, Gilia “(n. sp.?)” A. Gray in Gray et al., 1861 (Trachyophyta)
260
175
144
38
134
249
242
237
eastwoodiana, Amsonia Rydberg, 1913 (Trachyophyta)
85
195
28
308
96
14
184
40
5
189
274
facialis, Bombylius Cresson, 1919 (Arthropoda)
confertiflora, Oenothera Raven, 1962 (Trachyophyta)
confinis, Ptelea Green, 1906 (Trachyophyta)
conformis, Diplotaxis Fall, 1909 (Arthropoda)
congesta, Laphamia Jones, 1895 (Trachyophyta)
cowaniae, Parthenicus Knight, 1968 (Arthropoda)
cremnophylax, Astragalus Barneby, 1948 (Trachyophyta)
cupidineus, Dipodomys ordii Goldman, 1924 (Chordata)
curatorium, Carex Stacey, 1937 (Trachyophyta)
cypha, Gila R. Miller, 1946 (Chordata)
damei, Cercyonis Barnes & Benjamin, 1926 (Arthropoda)
delphinus, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
denningi, Mycetophila Shaw, 1951 (Arthropoda)
dentatus, Gorytes Fox, 1893 (Arthropoda)
depressa, Perdita Timberlake, 1968 (Arthropoda)
digna, Wiedemannia Sinclair, 2006 (Arthropoda)
dilaticosta, Polymedon Van Duzee, 1927 (Arthropoda)
dimidiatus, Ceratopogon C. Adams, 1903 (Arthropoda)
dionysus, Neominois Scudder, 1878 (Arthropoda)
dorothea, Neonympha dorothea Nabokov, 1942 (Arthropoda)
ecristata, Oenothera triloba var. Jones, 1895 (Trachyophyta)
elegans, Ptelea Green, 1906 (Trachyophyta)
endopterus, Astragalus wootoni var. Barneby, 1949 (Trachyophyta)
eremophila, Arabis Greene, 1900 (Trachyophyta)
evanida, Fabriciella Reinhard, 1953 (Arthropoda)
exiguus, Pilophorus Poppius, 1914 (Arthropoda)
extremitatis, Oreohelix yavapai Pilsbry & Ferriss, 1911 (Mollusca)
flaviatus, Phytocoris Knight, 1968 (Arthropoda)
flexisoma, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
flexuosus, Calochortus Watson, 1873 (Trachyophyta)
formicarii, Dactylopius Ehrhorn 1899 (Arthropoda)
fredonia, Athysanella Ball & Beamer, 1939 (Arthropoda)
fulvus, Deraeocoris Knight, 1921 (Arthropoda)
fureae, Coenonympha Barnes & Benjamin, 1926 (Arthropoda)
furnissii, Phacelia Atwood, 2007 (Trachyophyta)
fuscipennis, Pilophorus Knight, 1926 (Arthropoda)
fusiforme, Eriogonum Small, 1906 (Trachyophyta)
189
INDEX TO NOMINATE TAXA
317
181
213
97
15
281
178
215
219
101
170
166
148
gelida, Formica fusca fusca var. Wheeler, 1913 (Arthropoda)
77
244a
190
6
34
137
79
266
157
158
7
106
havasupaiensis, Sclerocactus Clover, 1942 (Trachyophyta)
240
167
56
212
285
292
99
131
immaculata, Schinia Pogue, 2004 (Arthropoda)
194
juniperi, Bolteria Knight, 1968 (Arthropoda)
117
217
253
48
164
172
74
kaiba, Loxosceles Gertsch & Ennik, 1983 (Arthropoda)
georgei, Nebria (Reductonebria) Kavanaugh, 2008 (Arthropoda)
gilli, Chionaspis Liu & Kosztarab, 1987 (Arthropoda)
gillii, Septogloeum D. Ellis, 1946 (Ascomycota)
globosa, Athysanella Ball & Beamer, 1939 (Arthropoda)
gloydi, Sonora semiannulata Stickel, 1938 (Chordata)
gracilis, Laphamia Jones, 1895 (Trachyophyta)
gracilis, Parasitorhabditis Massey 1974 (Nematoda)
gracilis, Spilogale Merriam, 1890 (Chordata)
grande, Catorama Fall, 1905 (Arthropoda)
grande, Cinara Hottes, 1956 (Arthropoda)
grandiscanyonensis, Arenivaga Hopkins, 2014 (Arthropoda)
grisea, Morus Greene, 1911 (Trachyophyta)
hesperia, Oenothera specuicola subsp. Raven, 1962 (Trachyophyta)
hesperius, Phytocoris Knight, 1928 (Arthropoda)
higginsii, Phacelia Atwood, 2007 (Trachyophyta)
hirsuta, Paratriatoma Barber, 1938 (Arthropoda)
hirtella, Schmaltzia Green, 1905 (Trachyophyta)
hopkinsi, Triepeolus Cockerell, 1905 (Arthropoda)
hualapaiensis, Mentzelia Schenk, Hodgson & Hufford, 2010 (Trachyophyta)
hualapaiensis, Pyrgulopsis Herschler, Liu & Stevens, 2016 (Mollusca)
hualpaensis, Opuntia Hester, 1943 (Trachyophyta)
hughesii, Phacelia Atwood, 2007 (Trachyophyta)
hunnewellii, Primula (Section Farinosae) Fernald, 1934 (Trachyophyta)
impensa, Arenivaga Hopkins, 2014 (Arthropoda)
incipiens, Trichochrous Casey, 1895 (Arthropoda)
inflata, Norvellina bicolorata (Ball) var. Lindsay, 1938 (Arthropoda)
infralabialis, Lampropeltis pyromelana Cope
Tanner, 1953 (Chordata)
inornata, Perdita Timberlake, 1962 (Arthropoda)
isidiomuscigena, Physconia Esslinger, 2000 (Ascomycota)
ivesiana, Phacelia (Coreanthus) A. Gray in Gray et al., 1861 (Trachyophyta)
kaibabensis, Agave McKelvey, 1949 (Trachyophyta)
kaibabensis, Andrena (Scaphandrena) Ribble, 1974 (Arthropoda)
kaibabensis, Branchinecta Belk & Fugate, 2000 (Arthropoda)
kaibabensis, Castilleja N. Holmgren, 1973 (Trachyophyta)
kaibabensis, Cicindela pusilla Johnson, 1990 (Arthropoda)
kaibabensis, Cordylanthus wrightii Chuang & Heckard, 1986 (Trachyophyta)
190
INDEX TO NOMINATE TAXA
108
182
227
258
50
42
248
45
115
155
220
252
288
123
3
223
180
2
257
70
196
84
kaibabensis, Coryphantha vivipara var. Fischer, 1979 (Trachyophyta)
297
295
10
4
307
102
247
60
143
lafontainei, Hadena (Hadena) Troubridge & Crabo, 2002 (Arthropoda)
30
93
246
284
26
19
150
315
318
286
262
macrodelphinus, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
kaibabensis, Enderleinellus Kim, 1966 (Arthropoda)
kaibabensis, Felis concolor E. Nelson & Goldman, 1931 (Chordata)
kaibabensis, Lesquerella Rollins, 1982 (Trachyophyta)
kaibabensis, Mischocyttarus flavitarsis var. Bequaert, 1932 (Arthropoda)
kaibabensis, Morchella Beug, T.A. Clem. & T.J. Baroni in Baroni et al., 2018 (Ascomycota)
kaibabensis, Morsea california Rehn & Grant, 1958 (Arthropoda)
kaibabensis, Papilio indra Bauer, 1955 (Arthropoda)
kaibabensis, Phidippus Gertsch, 1934b (Arthropoda)
kaibabensis, Polistes canadensis var. Lynn, 1932 (Arthropoda)
kaibabensis, Sciurus Merriam, 1904 (Chordata)
kaibabensis, Scutellaria potosina var. Rhodes & Ayers, 2010 (Trachyophyta)
kaibabensis, Suillus Thiers, 1976 (Basidiomycota)
kaibabensis, Thomomys fossor Goldman, 1938 (Chordata)
kaibabensis, Tipula (Lunatipula) C. Alexander, 1946 (Arthropoda)
kaibabiae, Saxinis saucia Moldenke, 1970 (Arthropoda)
kaibensis, Astragalus Jones, 1902 (Trachyophyta)
kathyae, Plagiomimicus J. Adams & Lafontaine, 2009 (Arthropoda)
kearneyi, Sisymbrium Rollins, 1953 (Trachyophyta)
kesimba, Tapinocyba Chamberlin, 1948 (Arthropoda)
knowltoni, Aphricus Knull, 1957 (Arthropoda)
knowltoni, Ostrya Coville, 1894 (Trachyophyta)
leptoscelis, Eleodes (Caverneleodes) Triplehorn, 1975 (Arthropoda)
ligulifolia, Salix lutea var. C. Ball, 1921 (Trachyophyta)
longicarpum, Thelypodium integrifolium (Nuttall) Endlicher subsp. Al-Shehbaz, 1973 (Trachyophyta)
longifolia, Brickellia (Clavigera) Watson, 1873 (Trachyophyta)
longulum, Catorama Fall, 1905 (Arthropoda)
lowellii, Fraxinus Sargent in Rehder, 1917 (Trachyophyta)
lugubris, Zopherodes Casey, 1907 (Arthropoda)
lutescens, Ptelea Green, 1906 (Trachyophyta)
macropetala, Fraxinus Eastwood, 1903 (Trachyophyta)
macropetala, Fraxinus cuspidata var. Rehder, 1917 (Trachyophyta)
macrosepala, Amsinckia Suksdorf, 1931 (Trachyophyta)
maeandrina, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
magnifica, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
margaretae, Boharticus Grissell, 1983 (Arthropoda)
maricopa, Pheidole desertorum var. Wheeler, 1906 (Arthropoda)
maricopa, Pogonomyrmex californicus Buckley subsp. Wheeler, 1914 (Mollusca)
mcdougallii, Flaveria Theroux, Pinkava & Keil, 1977 (Trachyophyta)
megalocarpa, Juniperus utahensis var. Sargent, 1919 (Trachyophyta)
191
INDEX TO NOMINATE TAXA
218
188
165
13
294
245
316
127
114
250
304
27
83
21
31
229
39
megalotis, Hesperomys Merriam, 1890 (Chordata)
275
283
228
186
23
72
9
306
124
140
301
nakaharai, Heterotarsonemus Smiley & Moser, 1974 (Arthropoda)
289
67
18
291
25
36
occidentalis, Suillus Thiers, 1976 (Basidiomycota)
279
168
138
49
233
8
107
pachyphylla, Allionia Standley, 1909 (Trachyophyta)
mellarius, Phytocoris Knight, 1925 (Arthropoda)
memorabilis, Mentzelia N. Holmgren & P. Holmgren, 2002 (Trachyophyta)
miniata, Yucanda E. Ball, 1937 (Arthropoda)
minuta, Microsciasma Townsend, 1915 (Arthropoda)
mohavenss, Panicum Reeder, 1991 (Trachyophyta)
moki, Formica Wheeler, 1906 (Arthropoda)
mokiacensis, Astragalus A. Gray, 1878 (Trachyophyta)
mokiensis, Arctosa Gertsch, 1934a (Arthropoda)
mortonianum, Eriogonum Reveal, 1974a (Trachyophyta)
multijuga, Oenothera (Chylisma) Watson, 1873 (Trachyophyta)
multimaeandrina, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
multinervatus, Phellopterus Coulter & Rose, 1900 (Trachyophyta)
multiphanosoma, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
multitrapezoidea, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
mychophilus, Otus asio Oberholser, 1937 (Chordata)
myriorrhaphis, Astragalus cremnophylax Barneby var. Barneby, 1979 (Trachyophyta)
nana, Amsinckia Suksdorf, 1931 (Trachyophyta)
navajo, Acmaeodera G. Nelson & Westcott, 1995 (Arthropoda)
navajo, Deraeocoris Knight, 1921 (Arthropoda)
navajoana, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
neofungorum, Mycetophila Chandler, 1993 (Arthropoda)
neotomae, Orchopeas sexdentatus Augustson, 1943 (Arthropoda)
newberryi, Peucedanum Watson, 1873 (Trachyophyta)
nicholi, Thomomys bottae Goldman, 1938 (Chordata)
nitida, Ptelea Green, 1906 (Trachyophyta)
nitidus, Polymedon Van Duzee, 1927 (Arthropoda)
occultum, Bembidion (Cyclolopha) Casey, 1918 (Arthropoda)
ocellata, Orbilia Baral, G. Marson & E. Weber in Baral, Weber & Marson, 2020 (Ascomycota)
opacella, Perdita Timberlake, 1956 (Arthropoda)
ophiosoma, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
oropedii, Astragalus lentiginosus Dougl. var. Barneby, 1945 (Trachyophyta)
pagana, Arenivaga Hopkins, 2014 (Arthropoda)
palida, Ptelea Green, 1906 (Trachyophyta)
paradinei, Pediocactus Benson, 1957 (Trachyophyta)
parashant, Ptomaphagus Peck & Wynne, 2013 (Arthropoda)
parashantensis, Xylorhiza tortifolia (Torrey & Gray) Greene var. Atwood & Welsh, 2013 (Trachyophyta)
parviflora, Adenostegia Ferriss, 1918 (Trachyophyta)
192
INDEX TO NOMINATE TAXA
255
312
122
197
199
105
20
69
162
230
103
55
313
231
44
171
236
152
58
66
61
86
305
17
parviflora, Crossosoma B. Robinson & Fernald, 1894 (Trachyophyta)
224
quercusgambellii, Gnomonia Monod, 1983 (Ascomycota)
135
254
222
121
261
57
80
78
298
296
recondita, Arabis Greene, 1900 (Trachyophyta)
146
126
76
112
133
173
saligna, Ptelea Green, 1906 (Trachyophyta)
pauperitense, Pediomelum S.L. Welsh, M. Licher, & N. D. Atwood in Welsh & Licher, 2010 (Trachyophyta)
peridoneus, Peromyscus crinitus Goldman, 1937 (Chordata)
perognathi, Wellcomia Kruidenier & Mehra, 1958 (Nematoda)
peromysci, Gongylonema Kruidenier & Peebles, 1958 (Nematoda)
perplexus, Bruchus Fall, 1910 (Arthropoda)
phanosoma, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
phasmus, Aphonopelma Chamberlin, 1940 (Arthropoda)
phillipsiana, Agave Hodgson, 2001 (Trachyophyta)
pholeter, Eschatomoxys Thomas & Pape in Pape, Thomas & Aalbu, 2007 (Arthropoda)
pingue, Catorama Fall, 1905 (Arthropoda)
piniedulis, Melanophila Burke, 1907 (Arthropoda)
pinionis, Astragalus Welsh & Thorne, 1977 (Trachyophyta)
pinkavae, Opuntia Parfitt, 1997 (Trachyophyta)
pletura, Acmaeodera W. Barr, 1972 (Arthropoda)
poketa, Cinara Hottes, 1956 (Arthropoda)
profundorum, Oreohelix yavapai Pilsbry & Ferriss, 1911 (Mollusca)
prosopidis, Geron Hall & Evenhuis, 2003 (Arthropoda)
pruddeni, Cryptorhopalum Casey, 1900 (Arthropoda)
pruddeni, Stenosphenus Casey, 1912 (Arthropoda)
pruddeni, Zopherodes Casey, 1907 (Arthropoda)
pseudoptutus, Penstemon virgatus Crosswhite, 1967 (Trachyophyta)
purpureus, Cymopterus Watson, 1873 (Trachyophyta)
purshiae, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
rectiramea, Silene Robinson, 1899 (Trachyophyta)
reederi, Sonorella W. Miller, 1984 (Mollusca)
repentinus, Castor canadensis Goldman, 1932 (Chordata)
retrorsum, Pediomelum Rydberg in Rydberg & Pennell, 1919 (Trachyophyta)
reversus, Trichochrous Casey, 1895 (Arthropoda)
romainei, Myrmecocystus melliger subsp. semirufus Emery var. Cole, 1936 (Arthropoda)
roseus, Sclerocactus havasupaiensis var. Clover, 1942 (Trachyophyta)
rubra, Trioza Tuthill, 1939 (Arthropoda)
rugicollis, Asidina Triplehorn & Brown, 1971 (Arthropoda)
scaposus, Astragalus A. Gray, 1878 (Trachyophyta)
schellbachi, Depressaria Gates, 1947 (Arthropoda)
schellbachi, Speyeria atlantis Garth, 1949 (Arthropoda)
simplicifolia, Rhus canadensis Marsh. var. Greene, 1890 (Trachyophyta)
skillmani, Trogloderus Johnston, 2019 (Arthropoda)
193
INDEX TO NOMINATE TAXA
244
24
63
321
139
147
52
125
33
specuicola, Oenothera Raven, 1962 (Trachyophyta)
264
191
174
75
234
65
110
120
269
109
263
142
311
151
tapeats, Efferia Scarbrough, Stevens & Nelson, 2012 (Arthropoda)
200
12
utahensis, Andrena (Callandrena) LaBerge, 1967 (Arthropoda)
299
116
81
192
214
272
vanduzeei, Platylygus Usinger, 1931 (Arthropoda)
92
51
290
169
wetherillii, Corydalis Eastwood, 1902 (Trachyophyta)
82
xeranthemoides, Echinocactus polycephalus var. Coulter, 1896 (Trachyophyta)
11
yuccandus, Myndus E. Ball, 1933 (Arthropoda)
spermoides, Orbilia Baral & G. Marson in Baral, Weber & Marson, 2020 (Ascomycota)
spiculosus, Prionus Casey, 1912 (Arthropoda)
stonei, Heteropogon Wilcox, 1965 (Arthropoda)
straminea, Ptelea Green, 1906 (Trachyophyta)
stygius, Senecio Greene, 1909 (Trachyophyta)
subatrata, Priononyx Bohart, 1958 (Arthropoda)
subcinereus, Astragalus A. Gray, 1878 (Trachyophyta)
subovoidea, Orbilia Baral, Matočec & E. Weber in Baral, Weber & Marson, 2020 (Ascomycota)
tenellus, Plagiognathus Knight, 1929 (Arthropoda)
tenerrimus, Astragalus humistratus var. Jones, 1895 (Trachyophyta)
tenuifolia, Encelia resinifera subsp. Curtis, 1998 (Trachyophyta)
tenuifolis, Cordylanthus Pennell, 1940 (Trachyophyta)
terminalis, Prionus Casey, 1912 (Arthropoda)
terricola, Spironema Foissner & Foissner, 1993 (Hemimastigophora)
tersus, Ammospermophilus leucurus Goldman, 1929 (Chordata)
thoosa, Synchloe Scudder, 1878 (Arthropoda)
toroweapensis, Echinocereus triglochidiatus var. Fischer, 1991 (Trachyophyta)
toumeyi, Populus fremontii var. Sargent, 1919 (Trachyophyta)
triptera, Ptelea Green, 1906 (Trachyophyta)
trumbullensis, Astragalus lentiginosus Douglas ex Hook. var. Welsh & Atwood, 2001 (Trachyophyta)
trumbullensis, Thomomys bottae Hall & Davis, 1934 (Chordata)
utahnum, Hysteropterum cornutum var. E. Ball, 1935 (Arthropoda)
variata, Steatoda Gertsch, 1960 (Arthropoda)
varipennis, Trypeta Coquillett, 1902 (Arthropoda)
varius, Phytocoris Knight, 1934 (Arthropoda)
vauxae, Oreohelix yavapai Marshall, 1929 (Mollusca)
voylesi, Pratherodesmus Shear in Shear, Taylor, Wynne & Krejca, 2009 (Arthropoda)
wheeleri, Andricus Beutenmüller, 1907 (Arthropoda)
wheeleri, Perdita Timberlake, 1928 (Arthropoda)
wickhami, Luperodes Horn, 1893 (Arthropoda)
____________________________________________________________________________________
194