PDCA ISSN 2181-2403
PDCA ISSN 2181-239X
REVIEW ARTICLE
Prominent botanists of Central Asia: Vvedensky Aleksei Ivanovich (1898–1972)
Furkat Khassanov1, Olga Cherneva2, Obidjon Turdiboev1, Shokhbozbek Turdiboev1, Komiljon Sh.
Tojibaev1*
1
2
Institute of Botany, Academy of Sciences of the Republic of Uzbekistan, Tashkent 100125, Uzbekistan
Komarov Botanical Institute RAS, St. Petersburg 197376, Russian Federation
ktojibaev@mail.ru
ABSTRACT
This article honours the memory of the outstanding taxonomist and tireless researcher of the flora of
Central Asia, Aleksei Ivanovich Vvedensky, the founder of the most extensive Central Asian
herbarium collection, the author of monographic revisions in ‘Flora of the USSR’, the editor and
author of the six volume edition of ‘Flora of Uzbekistan’, the ‘Conspectus Florae Asiae Mediae’, and
various taxonomic publications Vvedensky described dozens of new genera and hundreds of new
species of vascular plants from Central Asia.
Key words: Allium; Iris, Eremurus; bibliography; Vvedensky; taxonomy; Middle Asia
Aleksei Ivanovich Vvedensky was born on
March 7, 1898, in the city of Penza (Russia) in
the family of a teacher. He discovered his
interest in botany early in school. The famous
Penzini botanists Profs. I.I. Sprygin and M.G.
Popov played an eminent role in the career of
their young and energetic student. Under their
guidance, Aleksei acquired his first skills in
herbarization,
biodocumentation,
and
identification of vascular plants (Butkov et al.
1968).
The professional development of A.I.
Vvedensky took place against the backdrop of
the violent revolutionary events of 1917–1920,
the fall of the Russian Empire as a result of
revolutions, the First World War and the Civil
War. In 1917, Vvedensky, still very young,
became a member of the Penza Society for
Natural Sciences, whose chairman was I.I.
Sprygin. The following year, his first
publication appeared: ‘News and the rarity of
the Penza flora’ (1918). In January 1919,
Vvedensky was appointed head of the
http://doi.org/10.54981/PCDA/vol1_iss2/a1
Received: 30 June 2022; Accepted: 5 August 2022 Academic editor: Sergei Volis
Plant Diversity of Central Asia 2 (2022) 1–25
Herbarium of the Penza Provincial Natural
Historical Museum. A year later, he began
working at the Penza Zoological and Botanical
Garden, which was united with the Museum,
while still heading the herbarium and holding
the full-time position of taxonomist.
In 1920, by order of the government, a
large group of Russian scientists was sent to
Tashkent with the mission of creating a higher
educational institution in Turkestan. Together
with I.I. Sprygin, M.G. Popov, E.P. Korovin,
and four other employees of the Penza
Museum, Vvedensky arrived to Tashkent and,
among the large group of botanists and soil
scientists, became actively involved in
organizing the botanical department. The Penza
botanists who moved to Tashkent were called
the ‘Group of Sprygin’.
The Department of Botany they created
became part of the Physics and Mathematics
Faculty of the Turkestan People's University
(since 1924 the Central Asian State University;
since 1960 Tashkent State University; and
Khassanov et al.
currently the National University of
Uzbekistan). Since the 1920s, many leading
botanists of that time began to work there,
including R.I. Abolin, A.A. Baranov, A.V.
Blagoveschensky, V.P. Drobov, M.G. Popov,
E.P. Korovin, M.V. Kultiasov, N.D. Leonov,
I.A. Raikova, M.M. Sovetkina and others
(Tojibaev et al. 2020). Over time, it was this
department that would later serve as a scientific
centre for the development of plant science in
Central Asia. The Tashkent group of young and
talented botanists introduced the scientific
world to hundreds of new plant species, as well
as to the unique features of the vegetation of
Central Asia.
In Tashkent, throughout the 1920s,
Vvedensky combined organizational work with
scientific research at the university, which may
explain his relatively late graduation.
Vvedensky completed his higher education in
1929 with the thesis entitled ‘Central Asian
Gagea reticulata s.l. alliance and its path of
evolution’ (handwritten manuscript, 1929). At
that time, Vvedensky worked as a researcher at
the Faculty of Physics and Mathematics, head
of the herbarium of the Botanical Garden,
expert in the Botanical Garden, and head of the
herbarium of the Central Asian University
(Lipschitz 1947).
The special contribution of A.I.
Vvedensky was the creation of the University
Herbarium in 1920, originally called Herbarium
Horti Botanici Universitatis Asiae Mediae
(subsequently
Herbarium
Universitatis
Taschkenticae, acronym TAK) which he
himself headed since 1925. From 1923 to 1934,
twenty issues of ‘Shedae ad Herbarium Florae
Asiae Mediae’ were published (Vasilchenko,
Vassilieva, 1975). The herbarium was created
from the collections of the Bureau of Soil and
Botanical Research, established in 1909. Much
later, in early 1988, the University Herbarium
was merged with the Herbarium of the Institute
of Botany of the Academy of Sciences of the
Uzbek SSR (Pratov and Adylov 1996). The
TASH Herbarium (Institute of Botany,
Tashkent) (Pratov and Adylov 1996), which
already existed at that time, was originally
based on the collections of the plant resources
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
sector of the Committee of Sciences created in
1936. At first, in the united herbarium, the
university and academic collections existed in
parallel,
with
their
own
particular
nomenclature, and were located in different
premises. The collections were merged and
reorganized by T.A. Adylov, who was the
curator of the Herbarium from 1972 to 1988, in
a process of painstaking work. Later, A.I.
Vvedensky brought the number of sheets in this
collection to around half a million. Under his
leadership, there was established a practice of
an exchange of duplicates of herbarium
specimens with the then Soviet and foreign
herbaria, such as W (Vienna), P (Paris), K and
BM (London). At that time, this required not
only a huge organizational effort, but also the
firm authority of Vvedensky.
In working with the herbarium
collection, Vvedensky was exceptionally
meticulous and pedantic. According to the
memoirs of T.A. Adylov, when they brought
freshly
collected
plants,
Vvedensky
immediately sent the collectors to dry the plant
material
before proceeding
with
the
identification procedure. At the same time, he
did not trust anyone with a significant part of
the herbarization work, and preferred to
perform it himself, or only with reliable
employees (V.K. Pazij, S.S. Kovalevskaya,
N.B. Nikiforova, and others). Vvedensky also
actively participated in field research and
collected huge numbers of herbarium
specimens. The samples collected by him
personally constituted a significant part of the
material he used in the preparation of his major
scientific publications. So, when working on
the publication ‘Herbarium Florae Asiae
Mediae’ (1924–1928), Vvedensky utilized over
25,000 herbarium sheets, a significant portion
of which were collected in field expeditions led
by Vvedensky.
In 1935, A.I. Vvedensky was awarded
the Candidate of Biological Sciences degree
without the need of a dissertation defense
(based on the totality of his published works)
and was awarded the title of associate
professor. In 1943, he was elected a
corresponding member of the Academy of
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Khassanov et al.
Sciences of Uzbekistan. Vvedensky established
the Systematics of Vascular Plants Laboratory
at the Institute of Botany of the Academy of
Sciences of Uzbekistan, which he headed from
1950 to 1958.
Despite problems with his health that
arose with aging, he continued to lead and
participate in many field expeditions to various
regions throughout Central Asia (Fig. 1.).
Although his mobility decreased with age and
the expeditions became fewer, he nevertheless
was collecting large numbers of samples during
field work. Botanists dealing with the scientific
heritage of A.I. Vvedensky note the broad
spectrum of his floral interests. He was a
known expert for all plant groups, from ferns to
composites.
In 1923, A.I. Vvedensky published the
‘Identification of the Plants of Tashkent: an
illustrated guide to the identification of wild
vascular spore and flowering plants’ (Popov
1923), which focused on the plant diversity of
the mountainous surroundings Tashkent city
(Tashkentskiy Ala-Too). In fact, this
publication served as a textbook for teaching
lab classes on the taxonomy of flowering plants
at the university, which took place at that time
in the vicinity of the village of Kaplanbek (near
Tashkent, now belonging to Kazakhstan). Due
to the additional coverage of areas with high
plant diversity, such as the vicinities of Humsan
(Ugam Range), Chimgan (Chatkal Range), and
Burch Mullah (Koksuv Range), it became an
often used handbook for many Central Asian
botanists. The textbook focused on seven
families (Araceae, Lemnaceae, Juncaceae,
Liliaceae, Amaryllidaceae, Iridaceae and
Orchidaceae), 20 genera and 86 species. In the
following year, Vvedensky (1924) critically
reviewed 150 species for the Herbarium Florae
Asiae Mediae. For the Flora of Turkmenistan
(vol. 1, 1932), Vvedensky contributed the
families Liliaceae (except Tulipa, Eremurus,
Asparagus), Amaryllidaceae and Araceae,
totaling 15 genera and 74 species. Vvedensky
also contributed to the multi-volume Flora of
the USSR (30 volumes, 1934–1964), for which
the team of authors was selected with great
care. He revised the genera Bromus
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
(Vvedensky et al. 1934), Tulipa, Allium,
Ungernia, Iris, and Juno (Vvedensky 1935); the
Rutaceae and the genera Biebersteinia and
Erodium (Vvedensky 1949), Pedicularis
(Vvedensky 1955), contributing to the Flora of
the USSR in total morphological descriptions
of 518 species. To the six-volume edition of the
Flora of Uzbekistan (1941–1963), he
contributed the Eriocaulonaceae, Liliaceae,
Amaryllidaceae, Iridaceae (Schreder 1941),
Caryophyllaceae
(Vvedensky
1953),
Brassicaceae and Rosa (Vvedensky 1955).
Rutaceae,
Lythraceae
and
Onagraceae
(Vvedensky
1959),
Convolvulaceae,
Lamiaceae, Scrophulariaceae, Valerianaceae
and Campanulaceae (Vvedensky 1961), and the
dichotomous identification key for Asteraceae
(Vvedensky 1962). A total of 714 species were
referenced by Vvedensky in the Flora of
Uzbekistan. The Flora of Uzbekistan included
information on 4148 vascular plants of
Uzbekistan and in 1962 was awarded the
prestigious state scientific prize named after
Abu Rayhan Beruni (Sennikov 2016). For the
ten-volume Flora of Tajikistan (Ovchinnikov
1963), Vvedensky contributed the Araceae,
Liliaceae, Amaryllidaceae, and Iridaceae (28
genera, 236 species). For the first two volumes
of the ‘Conspectus Florae Asiae Mediae,’ he
contributed
the
Potamogetonaceae,
Zosteraceae, Najadaceae, and Pteridophyta
(Vvedensky 1968) and Liliaceae (Vvedensky
1971).
Vvedensky devoted much attention to
the ornamental bulbous plants of Central Asia.
Using the living collection of the Tashkent
Botanical Garden (TBG), he monographically
revised the genera Tulipa, Eremurus, Fritillaria
and Iris. He made a thorough revision of the
genus Allium L., a favorite genus that he
worked on throughout his life. The taxonomic
revisions by Vvedenskyi were adopted by
leading specialists from abroad. His revision of
the genus Allium L. was appreciated and
accepted by R.V. Kamelin (1973) and Per
Wendelbo (1971). In his work with Allium,
Vvedensky mainly used fresh plant materials of
the TBG, maintaining close collaboration with
the local staff. One result of this close
3
Khassanov et al.
collaboration was the establishment in the TBG
of a section with bulbous geophytes, mainly
from the genera Tulipa and Allium. At one
point, this collection consisted of about 6,000
wild species and horticultural varieties (Tulipa,
Eremurus, Iris, Juno, Crocus, Colchicum,
Fritillaria, etc.), of which more than 2000 were
wild and garden tulips. In the collection of the
TBG, preference was given to wild Central
Asian tulips (70% of the area was occupied by
wild species and about 30% by various varieties
of cultivated tulips, mainly Dutch (Tojibaev
2011).
An article dedicated to Vvedensky's
70th birthday (Butkov et al. 1968) noted the
high level of his knowledge of the flora of
Central Asia, as well as his great professional
erudition and excellent knowledge of botanical
nomenclature and geography.
According to oral communication by
O.V. Cherneva, A.I. Vvedensky dreamed of
compiling a map of ancient soils (variegate
outcrops) of the Central Asian region, and
‘populating’ the outcrops of such soils in
Central Asia with ancient plant communities.
This task was largely due to Vvedensky’s
specific scientific approach of involving
specialists from all related fields (geographers,
geologists, soil scientists, zoologists).
In 1968, the first volume of a 10-volume
monograph on the flora of Central Asia
appeared (Vvedensky 1968–1993), which was
essentially a critical regional flora overview
summarizing almost 100 years of research, as
well as revisions of the floras of neighbouring
to Central Asia regions Iran and Afghanistan. In
fact, this multi-volume work served as the basis
for the new Flora of Uzbekistan (Sennikov et
al. 2016).
In addition to the abovementioned
multi-volume publications, A.I. Vvedensky was
the editor of the following botanical reports: 18
issues of ‘Botanical materials of Herbarium of
the Institute of Botany of the Academy of
Sciences of the UzSSR’ (1941–1948); vols. III–
XI ‘Flora of Kyrgyzstan’ (1951–1965), ‘Key of
the wild plant species of the Hungary steppe’
(1960).
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
The following monographs were
published under the editorship of Vvedensky:
Lapin A.M. ‘Key of the plants of the
Tashkent oasis, Part 1’ (1938);
Korovin E.P. ‘Illustrated monograph of
the genus Ferula’ (1947);
Bondarenko O.N. ‘Key of the highest
plants of the Karakalpakia’ (1964);
Puchkova E.G. ‘Species of the
Chaetodon section and its position in the genus
Astragalus’ (1967) and many others.
As a university professor, A.I.
Vvedensky supervised undergraduate and
graduate students in the areas of floristics, plant
taxonomy, and phytogeography with his
impressive encyclopedic knowledge. Among
his students are: A.Ya. Butkov – one of the
leading phytocenologists of Uzbekistan; Z.P.
Botschantzeva – a leading scientist on tulips in
Central Asia and adjacent territories; S.S.
Kovalevskaja – a famous expert on many
leading taxonomic groups of the flora of
Central Asia, including Asteraceae; V.K. Pazij
– an eminent university lecturer who described
more then 40 new species from the families
Fabaceae, Poaceae, Euphorbiaceae, and
Lamiaceae; O.V. Cherneva – an expert in
Cousinia Cass. and Jurinea Cass. In working
with his students, Vvedensky had a unique
pedagogical style of teaching the students how
to conduct scientific research and write the
papers. His approach was fascinating and
highly motivating. However, Vvedensky
became notably disappointed when some of his
students, having undergone an extensive
training, moved from Tashkent to other
scientific institutions of the former Soviet
Union republics.
In total, A.I. Vvedensky's botanical
contribution to the flora of Central Asia
comprises the description of 9 new genera, 20
new sections, about 300 new species, more than
50 new combinations, amounting to more than
1,500 original descriptions of wild plants (Fig.
3.) (Table 1, 2) (Butkov et al. 1968). His
invaluable contribution to science was the
creation of the TASH Herbarium. Vvedensky
educated a large number of talented plant
taxonomists. He was the editor of a number of
4
Khassanov et al.
Vvedensky Aleksei Ivanovich
famous botanical publications and was one of
the first to recognize the uniqueness and
diversity of the Central Asian flora and its
connections with other floristic regions of the
world. Without a doubt, Aleksei Ivanovich
Vvedensky deserves enormous gratitude and
respect from botanists around the world.
A.I. Vvedensky was married and had a
daughter. In interpersonal communication A.I.
seems to have been a humorous man who loved
telling funny stories (Fig. 4.). Many photos
have been preserved from happy gatherings
with employees (Fig. 2). Other colleagues
enviously joked that the Vvedensky group
always had the most beautiful women.
Fig. 1. A.I. Vvedensky
Plant Diversity of Central Asia. 2 (2022) 1–25
5
Khassanov et al.
Vvedensky Aleksei Ivanovich
Fig. 2. Social gathering, 12 Feb. 1959. A.Y. Butkov, O.V. Cherneva, A.I. Vvedensky, S.S.
Kovalevskaya, S. Abdullayeva (from left to right)
On July 27, 1972, at age 74, Aleksei
Ivanovich Vvedensky died in Tashkent and was
buried at the Botkin cemetery.
It is difficult to overestimate the
contributions of Aleksei Ivanovich Vvedensky
to the development of plant systematics and the
herbaria of Central Asia.
Three genera have been named in
honour of Vvedensky (Vvedenskya Korovin,
Vvedenskyella Botsch., Alexeya Pachom.), one
section of Allium – Vvedenskya Kamelin and 51
species: Centaurea vvedenskyi Popov, Iris
vvedenskyi Nevski ex Woronow & Popov,
Tragopogon vvedenskyi Popov, Psephellus
vvedenskii Sosn., Gagea vvedenskyi Grossh., G.
alexejana Kamelin ex Levichev, G. alexii Ali &
Levichev, Juncus vvedenskyi V.I. Krecz.,
Salsola
vvedenskyi
Iljin
&
Popov,
Haplophyllum vvedenskyi Nevski, Rheum
vvedenskyi Sumnev., Polygonum vvedenskyi
Sumnev., Elytrigia vvedenskyi Drobow,
Ranunculus vvedenskyi Ovcz., Poa vvedenskyi
Drobow, Bupleurum vvedenskyi Manden.,
Astragalus vvedenskyi Popov, A. alexejii
Gontsch., Frankenia vvedenskyi Botsch.,
Plant Diversity of Central Asia. 2 (2022) 1–25
Andrachne vvedenskyi Pazij, Rosa vvedenskyi
Korotkova, Allium vvedenskyanum Pavlov,
Gentiana vvedenskyi Grossh., Acantholimon
vvedenskyi
Lincz.,
Aster
vvedenskyi
Bondarenko, Cousinia vvedenskyi Tscherneva,
Asparagus vvedenskyi Botsch., Ephedra
vvedenskyi Pachom., Hedysarum vvedenskyi
Korotkova, Olgaea vvedenskyi Iljin, Salvia
vvedenskyi Nikitina, Potentilla vvedenskyi
Botsch., Saussurea vvedenskyi Lipsch.,
Scrophularia vvedenskyi Bondarenko &
Filatova, Tulipa vvedenskyi Botschantz.,
Delphinium vvedenskyi Pachom., Strigosella
vvedenskyi Bondarenko & Botsch., Draba
vvedenskyi Kovalevsk., Scaligeria vvedenskyi
Kamelin, Saxifraga vvedenskyi Abdullaeva,
Viola alexejana Kamelin & Junussov, Alexeya
vvedenskyi Pachomova, Rosularia vvedenskyi
U.P. Pratov, Pseudoclausia vvedenskyi
Pachom., Ungernia vvedenskyi Khamidch.,
Valerianella
vvedenskyi
Lincz.,
Scilla
vvedenskyi Pazij, Oxytropis vvedenskyi Filim.,
Gentianа vvedenskyi Grossh., Lagochilus
vvedenskyi Kamelin & Zuckerw., Silene
vvedenskyi Lazkov.
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Khassanov et al.
Vvedensky Aleksei Ivanovich
Fig. 3. Manuscript of A.I. Vvedensky with a protologue of Eremurus brachystemon (archive
of Institute of Botany UAS).
Plant Diversity of Central Asia. 2 (2022) 1–25
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Khassanov et al.
Vvedensky Aleksei Ivanovich
Fig. 4. Friendly cartoon of A.I. Vvedensky, surprisingly found at the Botanical Institute of Komarov in
the office of I.A. Linczewsky (Institute of Botany, Kazakh Academy of Sciences) (author is unknown).
The original is kept at the Botanical Institute of Komarov (keeper is А.К. Sytin, to whom we are
greatly indebted for providing this cartoon).
List of main publications of A.I. Vvedensky
1918
Новости и редкости Пензенской флоры // Труды Пензенского общества любителей
естествознания и краеведения. Вып. 3–4 (1917 г.). С. 159–162.
1923
Новый род из семейства губоцветных Phlomidopsis // Труды Туркестанского Научного общества
при Средне-Азиатском Государственном Университете. Ташкент. Вып. I. С. 119–122.
(Совместно с М. Г. Поповым).
Новые виды туркестанских луков // Труды Туркестанского Научного общества при СреднеАзиатском Государственном Университете. Ташкент. Вып. I. С. 122–126.
Определитель растений окрестностей г. Ташкента, вып. I (Polypodiaceae — Urticaceae) / Под ред.
М. Г. Попова. — Ташкент: Изд-во САГУ. 80 с. (Совместно с В. П. Дробовым, Е. П. Коровиным,
М. В. Культиасовым, М. Г. Поповым, И. А. Райковой).
Определитель растений окрестностей г. Ташкента, вып. II (Urticaceae — Cruciferae) / Под ред. М.
Г. Попова. — Ташкент: Изд-во САГУ. 80 с.
Plant Diversity of Central Asia. 2 (2022) 1–25
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Khassanov et al.
Vvedensky Aleksei Ivanovich
1924
Decas Alliorum novorum ex Asia Media // Ботанические материалы Гербария Главного
ботанического сада РСФСР. Т. V, вып. 6. С. 89–96.
1925
Species novae е montibus Karatau Turkestaniae // Труды Туркестанского Научного общества при
Средне-Азиатском Государственном Университете. Вып. II. С. 29–31.
1932
Семейство Araceae // Флора Туркмении. Т. I. Ленинград: Изд-во Акад. наук СССР. С. 233–236.
Семейство Liliaceae // Флора Туркмении. Т. I. Ленинград: Изд-во Акад. наук СССР. 246–316.
1934
Descriptiones Alliorum novorum // Бюллетень Среднеазиатского государственного университета.
Вып. 19. С. 119–130.
Род Bromus L. // Флора СССР. Т. 2. Ленинград: Изд-во Акад. наук СССР. С. 554–583.
(Совместно с В. И. Кречетовичем, С. А. Невским, В. В. Сочавой).
1935
Род Allium L. // Флора СССР. Т. 4. Ленинград: Изд-во Акад. наук СССР. С. 112–280.
Род Tulipa L. // Флора СССР. Т. 4. Ленинград: Изд-во Акад. наук СССР. С. 320–364.
Род Ungernia Bunge // Флора СССР. Т. 4. Ленинград: Изд-во Акад. наук СССР. С. 481–485.
Род Iris L. (секция Juno) // Флора СССР. Т. 4. Ленинград: Изд-во Акад. наук СССР. С. 557–576.
Tulipae et Junones novae // Бюллетень Среднеазиатского государственного университета.
Ташкент, Вып. 21. С. 147–152.
1937
Новый род Korovinia из сем. Зонтичных // Труды Ботанического института АН СССР. I сер., 4:
272–273. (Совместно с С.А. Невским).
Новый род Platytaenia из сем. Зонтичных // Труды Ботанического института АН СССР. I сер., 4:
270–271. (Совместно с С. А. Невским).
1941
Семейство Eriocaulonaceae // Флора Узбекистана. Т. 1. Ташкент: Изд-во Акад. наук УзССР. С.
381–382.
Семейство Liliaceae // Флора Узбекистана. Т. 1. Ташкент: Изд-во Акад. наук УзССР. С. 394–495.
Семейство Amaryllidaceae // Флора Узбекистана. Т. 1. Ташкент: Изд-во Акад. наук УзССР. С.
495–502.
Семейство Iridaceae // Флора Узбекистана. Т. 1. Ташкент: Изд-во Акад. наук УзССР. С.502–522.
Диагнозы новых видов // Флора Узбекистана. Т. 1. Ташкент: Изд-во Акад. наук УзССР. С. 537,
540–545.
Новые виды растений // Ботанические материалы гербария института Ботаники Академии наук
Узбекской ССР. Т. III. С. 3–20. (Совместно с В. П. Бочанцевым).
Новый род семейства губоцветных // Ботанические материалы Гербария Ботанического
института имени В. Л. Комарова Академии наук СССР. Т. IX, вып. 2. С. 54–55.
Plant Diversity of Central Asia. 2 (2022) 1–25
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Vvedensky Aleksei Ivanovich
1946
Новые лилейные Таджикистана // Ботанические материалы Гербария Ботанического института
имени В. Л. Комарова Академии наук СССР. Т. IX, вып. 4–12. С. 232–247.
Новый вид рода Androsace из Памира // Ботанические материалы Гербария Ботанического
института имени В. Л. Комарова Академии наук СССР. Т. IX, вып. 4–12, С. 213–214. (Совместно
с П. Н. Овчинниковым).
1948
Новые виды рода Haplophyllum // Ботанические материалы Гербария Инстута ботаники и
Зоолигии АН УзССР, 10: 3–5.
Новые крестоцветные из Средней Азии // Ботанические материалы Гербария Инстута ботаники
и Зоолигии АН УзССР, 12: 3–12.
Род Erodium L’Her. // Флора СССР, Т. 14. Ленинград: Изд-во Акад. наук СССР. С. 63–72.
Семейство Biebersteinia Steph. // Флора СССР, Т. 14. Ленинград: Изд-во Акад. наук СССР. С. 74–
76.
Семейство Rutaceae // Флора СССР, Т. 14. Ленинград: Изд-во Акад. наук СССР. С. 198–241.
1952
Новые бобовые из Средней Азии // Ботанические материалы Гербария Инстута ботаники АН
УзССР. 13: 19–25.
Новые лилейные Киргизии // Ботанические материалы Гербария Инстута ботаники АН УзССР.
13. 26–34.
1953
Семейство Nyctaginaceae // Флора Узбекистана, Т. 2. Ташкент: Изд-во Акад. наук УзССР. С.
342–343.
Семейство Aizoaceae // Флора Узбекистана, Т. 2. Ташкент: Изд-во Акад. наук УзССР. С. 344.
Семейство Phytolaccaceae // Флора Узбекистана, Т. 2. Ташкент: Изд-во Акад. наук УзССР. С.
344.
Семейство Basellaceae // Флора Узбекистана, Т. 2. Ташкент: Изд-во Акад. наук УзССР. С. 347.
Семейство Caryophyllaceae // Флора Узбекистана, Т. 2. Ташкент: Изд-во Акад. наук УзССР. С.
349–427.
1954
Новые виды из Средней Азии // Ботанические материалы Гербария Инстута ботаники АН
УзССР. 14. С. 7–12.
1955
Два новых астрагала // Флора Узбекистана, 3. Ташкент: 798–799. (Совместно с С. С.
Ковалевской).
Ключ для определения видов рода Astragalus. // Флора Узбекистана, Т. 3. Ташкент: Изд-во Акад.
наук УзССР. С. 473–487.
Род Pedicularis // Флора СССР, Т. 22. Ленинград: Изд-во Акад. наук СССР. С. 687–795.
Род Calispepla Vved. // Флора Узбекистана, Т. 3. Ташкент: Изд-во Акад. наук УзССР. С. 412.
Семейство Cruciferae // Флора Узбекистана, Т. 3. Ташкент: Изд-во Акад. наук УзССР. С. 65–222.
(Совместно с В.П. Бочанцевым).
Род Rosa L. // Флора Узбекистана, Т. 3. Ташкент: Изд-во Акад. наук УзССР. С. 343–355
(Совместно с В. К. Пазий).
Plant Diversity of Central Asia. 2 (2022) 1–25
10
Khassanov et al.
Vvedensky Aleksei Ivanovich
1957
К дискуссии о виде // Известия Академии наук Узбекской ССР, серия биологическая, 2. 41–48.
1959
Заметки о некоторых среднеазиатских видах рода Pedicularis // Ботанические материалы
Гербария Инстута ботаники АН УзССР, 15: 3–9.
Новая секция рода Astragalus // Ботанические материалы Гербария Инстута ботаники АН
УзССР. 15: 10–12.
Род Epilobium Dill. ex L. // Флора Узбекистана, Т. 4. Ташкент: Изд-во Акад. наук УзССР. С. 243–
248. (Совместно с В.К. Пазий).
Семейство Lythraceae // Флора Узбекистана, Т. 4. Ташкент: Изд-во Акад. наук УзССР. С. 231–
239.
Семейство Onagraceae // Флора Узбекистана, Т. 4. Ташкент: Изд-во Акад. наук УзССР. С. 240–
252.
Семейство Rutaceae // Флора Узбекистана, Т. 4. Ташкент: Изд-во Акад. наук УзССР. С. 62–77.
1960
Определитель дикорастущих растений Голодной Степи. Труды Ташкентского государственного
университета. Вып. 178 (Ботаника). Ташкент: С. 216. (Совместно с В.П. Бочанцев, А.Я. Бутков,
Н.Б. Никифорова, В.К. Пазий).
1961
Antonina - новый род сем. Губоцветных // Ботанические материалы Гербария Инстута ботаники
АН УзССР. Вып. 16. – С. 15–17.
Новый вид из Ixiolirion Средней Азии / Ботанические материалы гербария института Ботаники
АН УзССР. Вып. 16. С. 18–21. [Совместно с С.С. Ковалевская].
О восстановлении Astragalus holargyreus // Ботанические материалы Гербария Института
ботаники АН УзССР. 16. 13–14.
Семейство Convolvulaceae // Флора Узбекистана, Т. 5. Ташкент: Изд-во Акад. наук УзССР. С.
123–139.
Семейство Labiatae // Флора Узбекистана, Т. 5. Ташкент: Изд-во Акад. наук УзССР. С. 263–416.
Семейство Scrophulariaceae // Флора Узбекистана, Т. 5. Ташкент: Изд-во Акад. наук УзССР. С.
446–498. (Совместно с О.Н. Бондаренко, С.С. Ковалевской, В.К. Пазий).
Семейство Valerianaceae // Флора Узбекистана, Т. 5. Ташкент: Изд-во Акад. наук УзССР. С. 578–
590.
Семейство Campanulaceae // Флора Узбекистана, Т. 5. Ташкент: Изд-во Акад. наук УзССР. С.
618–628.
1962
Семейственная характеристика и ключ для определения родов сем. Сложноцветных // Флора
Узбекистана, Т. 6. Ташкент: Изд-во Акад. наук УзССР. С. 23–31.
1963
Семейство Amaryllidaceae // Флора Таджикистана, Т. 2. Москва-Ленинград: Изд-во Акад. наук
СССР. С. 290–370.
Семейство Аrасеае // Флора Таджикистана, Т. 2. Москва-Ленинград: Изд-во Акад. наук СССР.
С. 146–154.
Семейство Liliасеае // Флора Таджикистана, Т. 2. Москва-Ленинград: Изд-во Акад. наук СССР.
С. 178–290.
Plant Diversity of Central Asia. 2 (2022) 1–25
11
Khassanov et al.
Vvedensky Aleksei Ivanovich
Семейство Iridaceae // Флора Таджикистана, Т. 2. Москва-Ленинград: Изд-во Акад. наук СССР.
С. 370–396.
1967
Заметки о среднеазиатских видах рода Eremostachys // Ботанические материалы гербария
института Ботаники АН УзССР. Вып. 18. С. 8–29 [Совместно с Пазый В.К.].
1968
Семейство Potomogetonaceae // Определитель растений Средней Азии. Т. 1. Ташкент: Изд-во
«Фан» УзССР. С. 38–43.
Семейство Zosteraceae // Определитель растений Средней Азии. Т. 1. Ташкент: Изд-во «Фан»
УзССР. С. 43–44.
Семейство Najadaceae // Определитель растений Средней Азии. Т. 1. Ташкент: Изд-во «Фан»
УзССР. С. 44–45.
Ключ для определения родов сем. Gramineae // Определитель растений Средней Азии. Т. 1.
Ташкент: Изд-во «Фан» УзССР. С. 94–109 [Совместно с С.С. Ковалевская, Н.Б. Никифорова].
Семейство Pteridophyta // Определитель растений Средней Азии. Т. 1. Ташкент: Изд-во «Фан»
УзССР. С. 7–19 [Совместно с Е.Г. Пучкова].
1971
Род Eremurus M.Bieb. // Определитель растений Средней Азии. Т. 2. Ташкент: Изд-во «Фан»
УзССР. С. 14–27 [Совместно с С.С. Ковалевская].
Род Gagea Salisb. // Определитель растений Средней Азии. Т. 2. Ташкент: Изд-во «Фан» УзССР.
С. 27–39.
Род Allium L. // Определитель растений Средней Азии. Т. 2. Ташкент: Изд-во «Фан» УзССР. С.
39–89.
Род Tulipa L. // Определитель растений Средней Азии. Т. 2. Ташкент: Изд-во «Фан» УзССР. С.
94–109 [Совместно с С.С. Ковалевская].
Род Juno Tratt. // Определитель растений Средней Азии. Т. 2. Ташкент: Изд-во «Фан» УзССР. С.
132–139.
Table 1. List of new vascular plant genera (9 genera) described by A.I. Vvedensky (including as a coauthor) between 1922–1987 (data from IPNI 2022)
Scientific name
Mesostemma Vved.
Antonina Vved.
Calispepla Vved.
Nevskiella V.I. Krecz. & Vved.
Platytaenia Nevski & Vved.
Stachyopsis Popov & Vved.
Prionotrichon Botsch. & Vved.
Phlomidoschema Vved.
Korovinia Nevski & Vved.
Plant Diversity of Central Asia. 2 (2022) 1–25
Publication
year
1941
1961
1952
1934
1937
1923
1948
1941
1937
Endemic to Central
Asia (POWO)
non-endemic
non-endemic
non-endemic
non-endemic
non-endemic
non-endemic
non-endemic
non-endemic
non-endemic
IPNI Plant Name
Id
956774-1
20666-1
21896-1
18593-1
40406-1
21240-1
12943-1
21112-1
40141-1
12
Khassanov et al.
Vvedensky Aleksei Ivanovich
Table 2. List of new vascular plant species described by A.I. Vvedensky (including as a co-author)
between 1922–1987 (data from IPNI 2022).
Scientific name (taxa)
Publication
year
Acanthophyllum cyrtostegium Vved.
Acanthophyllum leiostegium Vved.
Acanthophyllum ruptile Vved.
Allium alaicum Vved.
Allium albovianum Vved.
Allium alexandrae Vved.
Allium anisotepalum Vved.
Allium aroides Popov & Vved.
Allium brachyscapum Vved.
Allium brevidens Vved.
Allium brevidentiforme Vved.
Allium clausum Vved.
Allium confragosum Vved.
Allium crystallinum Vved.
Allium dasyphyllum Vved.
Allium dictyoscordum Vved.
Allium dodecadontum Vved.
Allium dolichomischum Vved.
Allium dolichostylum Vved.
Allium drepanophyllum Vved.
Allium drobovi Vved.
Allium dshungaricum Vved.
Allium eremoprasum Vved.
Allium eriocoleum Vved.
Allium eugeni Vved.
Allium ferganicum Vved.
Allium filidentiforme Vved.
Allium flavellum Vved.
Allium glaciale Vved.
Allium goloskokovii Vved.
Allium gracillimum Vved.
Allium gypsaceum Popov & Vved.
Allium gypsodictyum Vved.
Allium helicophyllum Vved.
Allium hexaceras Vved.
Allium hissaricum Vved.
Allium inconspicuum Vved.
1941
1941
1941
1934
1935
1924
1934
1934
1934
1924
1971
1971
1971
1934
1925
1924
1971
1971
1934
1934
1923
1971
1924
1952
1934
1924
1952
1946
1934
1971
1946
1924
1941
1934
1963
1963
1924
Plant Diversity of Central Asia. 2 (2022) 1–25
Endemic to
Central Asia
(POWO)
endemic
endemic
non-endemic
endemic
endemic
endemic
endemic
endemic
non-endemic
endemic
endemic
endemic
endemic
endemic
endemic
non-endemic
endemic
endemic
non-endemic
endemic
endemic
non-endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
non-endemic
endemic
endemic
endemic
IPNI Plant
Name Id
Holotype/lectotype
978235-1
978236-1
978237-1
527542-1
527555-1
527559-1
527606-1
527628-1
527710-1
527717-1
527718-1
527844-1
527864-1
527892-1
527914-1
527944-1
527952-1
527954-1
527955-1
527961-1
527962-1
77177072-1
527992-1
527994-1
528001-1
528026-1
528034-1
528047-1
528091-1
528101-1
528110-1
528134-1
528135-1
528146-1
528152-1
528162-1
528191-1
TASH001024
LE
TASH
LE
G00165237
LE
LE
TASH000334
TASH000339
LE
TASH000341
TASH000350
TASH000351
TASH000356
TASH000359
LE
TASH000362
TASH000364
LE
TASH000367
TASH
LE
TASH000380
TASH000381
LE
LE
LE
LE
TASH000384
AA
TASH000386
LE
TASH000388
TASH000391
TASH000392
TASH003028
TASH000395
13
Khassanov et al.
Vvedensky Aleksei Ivanovich
Allium incrustatum Vved.
Allium inops Vved.
Allium insufficiens Vved.
Allium jajlae Vved.
Allium jaxarticum Vved.
Allium jodanthum Vved.
Allium jucundum Vved.
Allium komarovianum Vved.
Allium kopetdagense Vved.
Allium kujukense Vved.
Allium kunthianum Vved.
Allium lasiophyllum Vved.
Allium leptomorphum Vved.
Allium lipskyanum Vved.
Allium litvinovii Drobow ex Vved.
Allium longiradiatum (Regel) Vved.
Allium lutescens Vved.
Allium majus Vved.
Allium margaritiferum Vved.
Allium marschallianum Vved.
1946
1924
1934
1934
1971
1971
1934
1929
1934
1923
1932
1934
1952
1971
1932
1925
1971
1941
1971
1935
endemic
endemic
endemic
non-endemic
endemic
endemic
endemic
non-endemic
non-endemic
endemic
non-endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
non-endemic
528192-1
528196-1
528198-1
528212-1
528219-1
528223-1
528225-1
528258-1
528260-1
528265-1
528267-1
528287-1
528305-1
528321-1
528325-1
528338-1
528352-1
528377-1
528384-1
528389-1
LE
LE
LE
LE
TASH000397
TASH000401
LE
LE
LE
TASH000404
LE
LE
LE
TASH000408
TASH000410
LE
TASH000412
TASH000419
TASH000423
TASH000335
Allium minutum Vved.
Allium mogoltavicum Vved.
Allium monophyllum Vved.
Allium nevskianum Vved.
Allium nothum Vved.
Allium ophiophyllum Vved.
Allium oreodictyum Vved.
Allium oreoprasoides Vved.
Allium oreoscordum Vved.
Allium parvulum Vved.
Allium pauli Vved.
Allium popovii Vved.
Allium praemixtum Vved.
Allium pseudoflavum Vved.
Allium pseudoseravschanicum
Popov & Vved.
Allium pumilum Vved.
Allium rhodanthum Vved.
Allium schachimardanicum Vved.
Allium schugnanicum Vved.
Allium scrobiculatum Vved.
Allium sergii Vved.
Allium sordidiflorum Vved.
1934
1934
1934
1971
1923
1928
1934
1925
1924
1934
1946
1923
1946
1934
1935
endemic
endemic
non-endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
non-endemic
endemic
endemic
non-endemic
non-endemic
528416-1
528426-1
528436-1
528484-1
528500-1
528533-1
528538-1
528541-1
528543-1
528588-1
528596-1
528642-1
528653-1
528675-1
528679-1
LE
TASH000427
LE
TASH000437
TASH000438
TASH000442
LE
TASH000444
LE
LE
LE
TASH000450
TASH000451
LE
LE
1934
1941
1971
1946
1923
1934
1946
non-endemic
endemic
endemic
non-endemic
endemic
endemic
endemic
528688-1
528732-1
528815-1
528827-1
528837-1
528850-1
528885-1
LE
TASH000454
TASH000458
LE
TASH000459
MW
LE
Plant Diversity of Central Asia. 2 (2022) 1–25
14
Khassanov et al.
Allium stephanophorum Vved.
Allium sulphureum Vved.
Allium svetlanae Vved. ex Filim.
Allium taciturnum Vved.
Allium taeniopetalum Popov &
Vved.
Allium trachyscordum Vved.
Allium transvestiens Vved.
Allium tytthanthum Vved.
Allium ugami Vved.
Allium vavilovii Popov & Vved.
Allium victoris Vved.
Alyssum stenostachyum Botsch. &
Vved.
Androsace acrolasia Vved. & Ovcz.
Antonina debilis (Bunge) Vved.
Arabis karatavica (Lipsch.) Botsch.
& Vved.
Arabis popovii Botsch. & Vved.
Asparagus ferganensis Vved.
Asparagus gypsaceus Vved.
Asparagus komarovianus Vved.
Asparagus monoclados Vved.
Astragalus devestitus Pazij & Vved.
Astragalus drobovii Popov & Vved.
Astragalus lachnolobus Kovalevsk.
& Vved.
Astragalus leptophysus Vved.
Astragalus rubescens Kovalevsk. &
Vved.
Astragalus stenophysus Vved. &
Zakirov
Astragalus thaumatothamnus Vved.
Astragalus uninodis Popov & Vved.
Bellevalia inconspicua Vved.
Calispepla aegacanthoides Vved.
Campyloptera carnea (Banks &
Sol.) Botsch. & Vved.
Cerastium bungeanum Vved.
Cerastium taschkendicum Adylov &
Vved.
Chrysanthemum santoanum Krasch.,
Popov & Vved.
Cistanche jodostoma Butkov &
Vved.
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
1924
1946
1982
1971
1934
endemic
endemic
endemic
endemic
endemic
528918-1
528954-1
899127-1
528964-1
528965-1
LE
LE
TASH000466
TASH000471
LE
1925
1935
1934
1923
1934
1971
1941
endemic
non-endemic
endemic
endemic
non-endemic
endemic
non-endemic
529026-1
529029-1
529071-1
529074-1
529098-1
529109-1
978254-1
TASH000473
TASH000480
TASH000482
TASH000485
TASH000488
TASH000490
TASH001324
1946
1961
1957
non-endemic 700154-1
non-endemic 444843-1
endemic
278325-1
TASH002598
LE
LE
1948
1941
1946
1946
1946
1952
1926
1955
endemic
endemic
endemic
non-endemic
endemic
endemic
endemic
endemic
278522-1
531103-1
531133-1
531156-1
531208-1
477126-1
477213-1
478198-1
TASH001245
TASH000563
TASH000567
LE
LE
TASH001651
TASH001660
TASH001714
1959
1955
endemic
endemic
478311-1
479585-1
TASH001726
TASH001788
1952
endemic
479951-1
TASH001797
1952
1922
1946
1952
1955
endemic
endemic
non-endemic
endemic
non-endemic
480190-1
480379-1
531813-1
483026-1
279891-1
TASH001834
TASH001840
LE
TASH003683
LE
1953
1971
non-endemic 152129-1
endemic
152692-1
LE
TASH000931
1928
endemic
LE
1961
non-endemic 661958-1
193733-1
TASH003056
15
Khassanov et al.
Clausia gracillima Popov ex Botsch.
& Vved.
Clausia tschimganica Popov ex
Botsch. & Vved.
Convolvulus pseudocantabrica
subspec. askabadensis (Bornm. &
Sint.) Vved.
Convolvulus tschimganicus Popov &
Vved.
Cousinia allolepis Tscherneva &
Vved.
Cousinia butkovii Tschern. & Vved.
Cousinia mogoltavica Tschern. &
Vved.
Cousinia praestans Tschern. &
Vved.
Cousinia pseudodshizakensis
Tschern. & Vved.
Cousinia strobilocephala
Tscherneva & Vved.
Crepis distincta Popov & Vved.
Cymatocarpus popovii Botsch. &
Vved.
Dianthus angrenicus Vved.
Dianthus brevipetalus Vved.
Dianthus helenae Vved.
Dianthus ugamicus Vved.
Dictamnus tadshikorum Vved.
Dipcadi turkestanicum Vved.
Eminium regelii Vved.
Epilobium acradenum Pazij & Vved.
Epilobium turkestanicum Pazij &
Vved.
Eremostachys anisochila Pazij &
Vved.
Eremostachys codonocalyx Pazij &
Vved.
Eremostachys eriolarynx Pazij &
Vved.
Eremostachys impressa Pazij &
Vved.
Eremostachys integior Pazij & Vved.
Eremostachys isochila Pazij & Vved.
Eremostachys labiosissima Pazij &
Vved.
Eremostachys leiocalyx Pazij &
Vved.
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
1941
endemic
978246-1
TASH001373
1941
endemic
978247-1
TASH001379
1925
non-endemic 77188795-1
BM000832678
1927
endemic
267145-1
TASH002660
1962
endemic
198647-1
TASH003132
1961
1961
endemic
endemic
198731-1
199109-1
TASH003394
TASH003479
1961
endemic
199211-1
TASH003523
1961
endemic
199222-1
TASH003527
1961
endemic
199350-1
TASH003145
1933
1955
endemic
endemic
199820-1
281908-1
TASH003679
TASH001199
1941
1941
1941
1941
1949
1941
1941
1959
1959
endemic
endemic
endemic
endemic
non-endemic
endemic
endemic
non-endemic
endemic
978238-1
978239-1
978240-1
978241-1
772379-1
534000-1
87000-1
611567-1
612410-1
TASH001031
TASH001035
TASH001042
TASH001056
LE
TASH000557
TASH000205
TASH002164
TASH002166
1961
endemic
446770-1
TASH002863
1961
endemic
446786-1
TASH
1961
endemic
446799-1
TASH002882
1961
endemic
446814-1
TASH000205
1961
1961
1961
endemic
endemic
endemic
446815-1
446816-1
446824-1
LE
TASH000236
TASH002788
1961
endemic
446829-1
TASH002791
16
Khassanov et al.
Eremostachys sogdiana Pazij &
Vved.
Eremurus ambigens Vved.
Eremurus ammophilus Vved.
Eremurus brachystemon Vved.
Eremurus candidus Vved.
Eremurus cristatus Vved.
Eremurus dolichomischus Vved. &
Wendelbo
Eremurus fuscus (O. Fedtsch.) Vved.
Eremurus hilariae Popov & Vved.
Eremurus hissaricus Vved.
Eremurus iae Vved.
Eremurus lachnostegius Vved.
Eremurus micranthus Vved.
Eremurus pubescens Vved.
Eremurus regelii Vved.
Eremurus roseolus Vved.
Eremurus subalbiflorus Vved.
Eremurus tadshikorum Vved.
Eremurus tianschanicus Pazij &
Vved. ex Pavlov
Eremurus zenaidae Vved.
Eremurus zoae Vved.
Erysimum chrysanthum Botsch. &
Vved.
Erysimum clausioides Botsch. &
Vved.
Erysimum ferganicum Botsch. &
Vved.
Erysimum gypsaceum Botsch. &
Vved.
Erysimum jodonyx Botsch. & Vved.
Erysimum microtrichon Botsch. &
Vved.
Erysimum nuratense Popov ex
Botsch. & Vved.
Erysimum popovii Botsch. & Vved.
Fritillaria olgae Vved.
Gagea capillifolia Vved.
Gagea circumplexa Vved.
Gagea delicatula Vved.
Gagea exilis Vved.
Gagea gageoides (Zucc.) Vved.
Gagea graminifolia Vved.
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
1961
endemic
446863-1
TASH002792
1941
1971
1946
1946
1952
1966
non-endemic
endemic
endemic
endemic
endemic
non-endemic
534671-1
534672-1
534682-1
534685-1
534692-1
534693-1
LE
LE
TASH000223
TASH000224
TASH000228
W-2780
1952
1924
1946
1941
1954
1946
1946
1928
1946
1971
1946
1958
non-endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
non-endemic
non-endemic
endemic
endemic
534696-1
534700-1
534702-1
534703-1
534713-1
534718-1
534726-1
534728-1
534730-1
534740-1
534743-1
534747-1
LE
TASH061356
TASH000232
TASH000236
LE
LE
TASH
LE
LE
TASH000243
LE
TASH000248
1952
1971
1941
endemic
endemic
endemic
534753-1
534754-1
978243-1
TASH
TASH000251
TASH001258
1948
endemic
283714-1
TASH001262
1948
endemic
283799-1
TASH001269
1941
endemic
978244-1
TASH001273
1941
1948
endemic
endemic
978245-1
283988-1
TASH001278
TASH001282
1948
endemic
284025-1
TASH001288
1948
1932
1923
1952
1941
1946
1932
1932
endemic
endemic
non-endemic
non-endemic
endemic
non-endemic
non-endemic
non-endemic
284093-1
535249-1
535467-1
535479-1
535495-1
535512-1
535521-1
535530-1
TASH001289
TASH000492
TASH000260
TASH000261
TASH000267
TASH000270
G
TASH000276
17
Khassanov et al.
Gagea gymnopoda Vved.
Gagea incrustata Vved.
Gagea kopetdagensis Vved.
Gagea minutissima Vved.
Gagea paedophila Vved.
Gagea parva Vved. ex Grossh.
Gagea popovii Vved.
Gagea praemixta Vved.
Gagea pseudoreticulata Vved.
Gagea subtilis Vved.
Gagea triquetra Vved.
Gagea vegeta Vved.
Gagea villosula Vved.
Geranium charlesii (Aitch. &
Hemsl.) Vved.
Geranium transversale Vved. ex
Pavlov
Gypsophila ferganica Vved.
Gypsophila knorringiana (Schischk.)
Vved.
Haplophyllum ciscaucasicum (Rupr.)
Grossh. & Vved.
Haplophyllum ferganicum Vved.
Haplophyllum foliosum Vved.
Haplophyllum leptomerum Lincz. &
Vved.
Haplophyllum multicaule Vved.
Haplophyllum ramosissimum
(Paulsen) Vved.
Haplophyllum tenuisectum Lincz. &
Vved.
Hedysarum setosum Vved.
Hyacinthus saviczii (Woron.) Vved.
Hypogomphia bucharica Vved.
Hypogomphia purpurea (Regel)
Vved. ex Kochk.
Iris amankutanica O. Fedtsch. ex
Vved.
Iris brevituba (Maxim.) Vved.
Iris kopetdagensis Vved.
Iris magnifica Vved.
Iris maracandica Vved.
Iris popovii Vved.
Iris subdecolorata Vved.
Iris tadshikorum Vved.
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
1941
1941
1932
1946
1946
1935
1925
1977
1932
1941
1923
1946
1946
1937
endemic
endemic
endemic
endemic
non-endemic
non-endemic
endemic
endemic
non-endemic
endemic
endemic
non-endemic
endemic
non-endemic
535538-1
535552-1
535562-1
535589-1
535605-1
535608-1
535621-1
77176982-1
535627-1
535669-1
535669-1
535706-1
535710-1
372883-1
TASH000277
TASH000283
LE
LE
LE
TASH000295
TASH000298
TASH000308
TASH000310
TASH000320
TASH000323
TASH000327
LE
BM000606788
1934
non-endemic 373752-1
MW
1941
1953
endemic
endemic
978233-1
154509-1
TASH001009
LE
1949
non-endemic 773861-1
TASH000205
1948
1948
1949
endemic
773865-1
endemic
773866-1
non-endemic 773877-1
TASH002053
TASH000232
TASH002061
1949
1949
endemic
endemic
773883-1
773889-1
TASH002063
LE
1949
non-endemic 773897-1
TASH002066
1952
1932
1954
1986
non-endemic
non-endemic
endemic
endemic
498104-1
536826-1
447910-1
943708-1
TASH001980
TBI
TASH002852
LE
1924
endemic
77241657-1
LE
1951
1935
1935
1935
1935
1923
1935
non-endemic
non-endemic
endemic
endemic
endemic
endemic
endemic
438439-1
438765-1
438839-1
438845-1
438982-1
439145-1
439159-1
LE
TASH
TASH000609
TASH000612
TASH001288
TASH000617
LE
18
Khassanov et al.
Vvedensky Aleksei Ivanovich
Iris thianschanica (Maxim.) Vved.
Iskandera alaica (Korsh.) Botsch. &
Vved.
Ixiolirion ferganicum Kovalevsk. &
Vved.
Juno baldshuanica (O. Fedtsch. & B.
Fedtsch.) Vved.
Juno bucharica (Foster) Vved.
Juno capnoides Vved.
Juno hippolyti Vved.
Juno inconspicua Vved.
Juno leptorrhiza Vved.
Juno linifolia (O. Fedtsch.) Vved.
Juno narbuti (O. Fedtsch.) Vved.
Juno narynensis (O. Fedtsch.) Vved.
Juno nicolai Vved.
Juno orchioides (Carrière) Vved.
Juno parvula Vved.
Juno rosenbachiana (Regel) Vved.
Juno svetlanae Vved.
Juno tubergeniana (Foster) Vved.
Juno vicaria Vved.
Juno warleyensis (Foster) Vved.
1932
1955
non-endemic 439179-1
endemic
285888-1
LE
LE
1961
endemic
TASH000584
1963
non-endemic 439666-1
LE
1941
1971
1941
1971
1935
1935
1941
1941
1935
1941
1963
1935
1971
1941
1963
1941
non-endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
endemic
non-endemic
endemic
endemic
endemic
endemic
439667-1
439668-1
439678-1
439680-1
439684-1
439685-1
439689-1
439690-1
439691-1
439693-1
439696-1
439704-1
439710-1
439712-1
439714-1
439716-1
Juno willmottiana (Foster) Vved.
1941
endemic
439717-1
Juno zenaidae Vved.
Korovinia tenuisecta (Regel &
Schmalh.) Nevski & Vved.
Lagochilus gypsaceus var.
linearilobus Vved. ex Zuckerw.
Lagochilus gypsaceus Vved.
Lagochilus hirsutissimus Vved.
1971
1937
endemic
439721-1
non-endemic 843908-1
P01846412
TASH000599
TASH0000314
TASH000604
LE00050076
LE
LE00050088
LE00050091
TASH000613
BM001191245
TASH000615
LE00050101
TASH000616
LE00050146
TASH000598
Lectotype: [Icon]
”Iris warleyensis”
in Foster,
Gard.Chron., ser.
3, 31: 386, f. 134.
1902 (lectotype
designated by
Boltenkov 2016:
231)
Lectotype [Icon]:
Iris willmottiana
in Foster (1901:
fig. 100)
(lectotype
designated by
Boltenkov (2016:
231)
TASH000619
LE
1985
endemic
918980-1
TASH
1961
1961
endemic
endemic
448727-1
448728-1
TASH002901
TASH002900
Plant Diversity of Central Asia. 2 (2022) 1–25
65679-1
19
Khassanov et al.
Vvedensky Aleksei Ivanovich
Lagochilus intermedius Vved.
Lagochilus pubescens Vved.
Lagochilus setulosus Vved.
Lepidium subcordatum Botsch. &
Vved.
Linum heterosepalum subsp.
tianschanicum Vved.
Malcolmia malacotricha Botsch. &
Vved.
Malcolmia pamirica Botsch. &
Vved.
Malcolmia spryginioides Botsch. &
Vved.
Matthiola czerniakowskae Botsch. &
Vved.
Melandrium glaberrimum
Bondarenko & Vved.
Melandrium turkestanicum (Regel)
Vved.
Mesostemma karatavicum
(Schischk.) Vved.
Mesostemma kotschyanum (Fenzl ex
Boiss.) Vved.
Micromeria gontscharovii Vved.
Micromeria popovii (B. Fedtsch. &
Gontsch.) Vved.
Mulgedium roseum Popov & Vved.
1961
1954
1955
1941
endemic
endemic
endemic
endemic
448738-1
448756-1
448761-1
978255-1
TASH002903
TASH002905
TASH002907
TASH001159
1941
endemic
978255-1
TASH003956
1952
endemic
286715-1
TASH001309
1952
non-endemic 286721-1
TASH001314
1952
non-endemic 286726-1
TASH001318
1941
endemic
978252-1
TASH001324
1971
endemic
155293-1
TASH001007
1953
endemic
155388-1
LE
1941
endemic
961833-1
LE
1941
non-endemic 961834-1
LE
1961
1961
non-endemic 451781-1
non-endemic 451863-1
LE
LE
1925
TASH003656
Nevskiella gracillima (Bunge) V.I.
Krecz. & Vved.
Onobrychis amoena Popov & Vved.,
Oryzopsis tianschanica Drobow &
Vved.
Otostegia glabricalyx Vved.
Otostegia megastegia Vved.
Oxytropis aulieatensis Vved.
Oxytropis echidna Vved.
Oxytropis ervicarpa Vved. ex Z.N.
Filimonova
Oxytropis jucunda Vved.
Oxytropis pilosissima Vved.
Parrya angrenica Botsch. & Vved.
Parrya crassifolia Botsch. & Vved.
Parrya darvazica Botsch. & Vved.
Parrya minuta Botsch. & Vved.
Parrya nuratensis Botsch. & Vved.
1934
no
233741-1
information
non-endemic 409502-1
1927
1941
non-endemic 509836-1
non-endemic 410235-1
TASH001989
TASH000055
1954
1954
1948
1925
1982
endemic
endemic
endemic
endemic
endemic
453639-1
453646-1
511307-1
511442-1
900445-1
TASH002910
TASH002913
TASH001864
TASH001869
TASH001874
1948
1948
1941
1948
1965
1948
1941
endemic
endemic
endemic
endemic
endemic
endemic
endemic
511565-1
511775-1
978248-1
287963-1
287964-1
287992-1
978249-1
TASH001876
TASH003919
TASH001332
TASH001335
LE
TASH001347
TASH001351
Plant Diversity of Central Asia. 2 (2022) 1–25
LE
20
Khassanov et al.
Parrya pamirica Botsch. & Vved.
Parrya saxifraga Botsch. & Vved.
Parrya subscapigera Botsch. &
Vved.
Parrya villosula Botsch. & Vved.
Pedicularis alatavica Stadlm. ex
Vved.
Pedicularis allorrhampha Vved.
Pedicularis amoeniflora Vved.,
Pedicularis argutiserrata Vved.
Pedicularis chroorrhyncha Vved.
Pedicularis gypsicola Vved.
Pedicularis hyperborea Vved.
Pedicularis inconspicua Vved.
Pedicularis macrochila Vved.
Pedicularis pallasii Vved.
Pedicularis popovii Vved.
Pedicularis pubiflora Vved.
Pedicularis schistostegia Vved.
Pedicularis sibirica Vved.
Pedicularis talassica Vved.
Pedicularis uralensis Vved.
Pedicularis verae Vved.
Pedicularis willdenovii Vved.
Petilium eduardii (Regel) Vved.
Phlomidoschema parviflorum
(Benth.) Vved.
Phlomis hypoleuca Vved.
Phlomis tschimganica Vved.
Phlomis tytthaster Vved.
Phyteuma occultans Popov & Vved.
Platytaenia pamirica (Lipsky)
Nevski & Vved.
Platytaenia tragioides (Boiss.)
Nevski & Vved.
Prionotrichon erysimoides (Kar. &
Kir.) Botsch. & Vved.
Prionotrichon pseudoparrya Botsch.
& Vved.
Pyrethrum santoanum Krasch.,
Popov & Vved.
Rhammatophyllum frutex Botsch. &
Vved.
Rhinopetalum arianum Losinsk. &
Vved.
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
1965
1941
1948
non-endemic 287995-1
endemic
978250-1
endemic
288018-1
LE
TASH001349
TASH001355
1941
1955
endemic
endemic
978251-1
806750-1
TASH001357
H
1961
1955
1955
1955
1959
1955
1955
1925
1955
1955
1955
1955
1955
1955
1955
1955
1955
1941
1941
endemic
endemic
non-endemic
non-endemic
endemic
non-endemic
endemic
endemic
non-endemic
endemic
endemic
non-endemic
non-endemic
endemic
non-endemic
non-endemic
non-endemic
non-endemic
non-endemic
806758-1
806768-1
806793-1
806908-1
807121-1
807164-1
807178-1
807314-1
186447-2
807486-1
807526-1
807614-1
807638-1
807699-1
807773-1
807781-1
186481-2
539769-1
453773-1
TASH003042
LE
LE
LE
TASH003047
LE
TASH003049
TASH003050
LE
TASH003051
TASH003052
LE
LE
TASH003054
LE
LE
LE
LE
M
1954
1954
1954
1925
1937
endemic
non-endemic
non-endemic
endemic
non-endemic
453913-1
454096-1
454100-1
144436-1
847195-1
TASH002813
TASH002828
TASH002827
TASH003123
LE
1937
non-endemic 847201-1
G-BOISS
1948
endemic
288191-1
MW
1948
endemic
288193-1
TASH001256
1928
endemic
240929-1
TASH003261
1952
endemic
288565-1
TASH001251
1935
non-endemic 540388-1
LE
21
Khassanov et al.
Rhinopetalum gibbosum (Boiss.)
Losinsk. & Vved.
Salvia submutica Botsch. & Vved.
Saponaria gypsacea Vved.
Saponaria spathulifolia (Fenzl)
Vved.
Saussurea hypargyrea Lipsch. &
Vved.
Scutellaria andrachnoides Vved.
Scutellaria angrenica Juz. & Vved.
Scutellaria glabrata Vved.
Scutellaria heterotricha Juz. &
Vved.
Scutellaria nevskii Juz. & Vved.
Scutellaria popovii Vved.
Scutellaria rubromaculata Juz. &
Vved.
Scutellaria urticifolia Juz. & Vved.
Scutellaria velutina Juz. & Vved.
Silene acutidentata Bondarenko &
Vved.
Silene eviscosa Bondarenko & Vved.
Silene excedens Bondarenko &
Vved.
Silene fedtschenkoi Bondarenko &
Vved.
Silene megalantha Bondarenko &
Vved.
Silene monantha Bondarenko &
Vved.
Silene paranadena Bondarenko &
Vved.
Silene pubicalyx Bondarenko &
Vved.
Silene schischkinii (Popov) Vved.
Sophiopsis micrantha Botsch. &
Vved.
Spergularia adenophora Vved.
Spergularia glaucophylla Vved.
Spergularia microsperma (Kindb.)
Vved.
Spergularia microspermoides Vved
Spergularia nematopoda Vved.
Stachyopsis lamiiflora (Rupr.) Popov
& Vved.
Stachyopsis oblongata (Schrenk)
Popov & Vved.
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
1935
non-endemic 540392-1
G-BOISS
1954
1941
1935
endemic
endemic
endemic
TASH002972
TASH001137
LE
1954
non-endemic 242312-1
LE
1954
1954
1954
1951
endemic
endemic
endemic
endemic
458127-1
458129-1
458287-1
458325-1
TASH002738
TASH003775
TASH002744
LE
1951
1954
1951
endemic
endemic
endemic
458489-1
458578-1
458625-1
LE
TASH002762
LE
1954
1954
1971
endemic
endemic
endemic
458738-1
458746-1
156737-1
LE
TASH002778
TASH000943
1971
1971
endemic
157315-1
non-endemic 157319-1
TASH000969
TASH000968
1971
endemic
157338-1
TASH000973
1971
endemic
157792-1
LE
1971
endemic
157848-1
LE
1971
endemic
158010-1
TASH000991
1971
endemic
158160-1
TASH000996
1953
1952
endemic
158325-1
non-endemic 289972-1
TASH003915
TASH001241
1941
1941
1953
non-endemic 957737-1
non-endemic 978230-1
non-endemic 158952-1
TASH000937
LE
P00388961
1941
1941
1923
non-endemic 978231-1
non-endemic 978232-1
non-endemic 459252-1
LE
LE
LE
1923
non-endemic 459254-1
LE
457324-1
978242-1
156659-1
22
Khassanov et al.
Stroganowia angustifolia Botsch. &
Vved.
Stroganowia longifolia (Boiss.)
Botsch. & Vved.
Stroganowia minor Botsch. & Vved.
Stroganowia tianschanica Botsch. &
Vved.
Stubendorffia curvinervia Botsch. &
Vved.
Stubendorffia pterocarpa Botsch. &
Vved.
Tetracmidion glochidiatum Botsch.
& Vved.
Thymus subnervosus Vved. Nabiev
& Tulyag.
Trichanthemis radiata Krasch. &
Vved.
Trichochiton umbrosum Botsch. &
Vved.
Tulipa anisophylla Vved.
Tulipa bifloriformis Vved.
Tulipa binutans Vved.
Tulipa carinata Vved.
Tulipa dasystemonoides Vved.
Tulipa dubia Vved.
Tulipa dykesiana Vved.
Tulipa ferganica Vved.
Tulipa hissarica Popov & Vved.
Tulipa karatavica (Regel) Vved. ex
Lipsch.
Tulipa korshinskyi Vved.
Tulipa mogoltavica Popov & Vved.
Tulipa orithyioides Vved.
Tulipa orthopoda Vved.
Tulipa paradasystemon Vved.
Tulipa platystemon Vved.
Tulipa prolongata Vved.
Tulipa rosea Vved.
Tulipa subbiflora Vved.
Tulipa subpraestans Vved.
Tulipa subquinquefolia Vved.
Tulipa zenaidae Vved.
Ungernia ferganica Vved. ex
Artjush.
Ungernia minor Vved.
Plant Diversity of Central Asia. 2 (2022) 1–25
Vvedensky Aleksei Ivanovich
1948
endemic
290225-1
TASH001172
1948
non-endemic 290235-1
G
1941
1948
endemic
endemic
978256-1
290242-1
TASH001178
TASH001187
1952
endemic
290250-1
TASH001192
1941
endemic
978257-1
TASH001193
1941
endemic
978253-1
TASH001307
1987
934225-1
TASH002985
1929
no
information
endemic
256198-1
TASH003359
1948
non-endemic 290864-1
TASH001321
1935
1971
1952
1971
1935
1935
1935
1935
1935
1932
endemic
endemic
non-endemic
non-endemic
non-endemic
endemic
non-endemic
endemic
endemic
non-endemic
542823-1
542848-1
542850-1
542871-1
542898-1
542902-1
542903-1
542911-1
542948-1
542961-1
LE
TASH000502
TASH000503
TASH000506
TASH000507
LE
LE
TASH000508
TASH
LE
1935
1971
1935
1971
1935
1935
1971
1971
1971
1971
1946
1935
1970
endemic
non-endemic
endemic
endemic
non-endemic
endemic
non-endemic
endemic
endemic
endemic
endemic
non-endemic
endemic
542969-1
543007-1
543025-1
543029-1
543032-1
543039-1
543047-1
543063-1
543100-1
928395-1
543103-1
543153-1
66973-1
LE
TASH000513
TASH000514
TASH000516
TASH
LE
LE
TASH000520
TASH000523
TASH000524
LE
TASH000555
TASH000513
1935
endemic
66975-1
LE
23
Khassanov et al.
Ungernia oligostroma Popov &
Vved.
Ungernia tadschicorum Vved. ex
Artjush.
Ungernia victoris Vved. ex Artjush.
Veronica mogoltavica Popov ex
Vved.
Veronica nanella Vved.
Veronica stylophora Popov ex Vved.
Viola alaica Vved.
Xiphion winkleri (Regel) Vved.
Ziziphora pedicellata Pazij & Vved.
Ziziphora suffruticosa Pazij & Vved.
Vvedensky Aleksei Ivanovich
1959
endemic
66977-1
TASH000578
1970
endemic
66979-1
TASH000529
1970
1925
endemic
66983-1
non-endemic 812325-1
TASH000349
LE
1961
1925
1928
1941
1961
1961
non-endemic
endemic
endemic
endemic
endemic
endemic
TASH003025
TASH003028
LE
LE
TASH002979
TASH002981
Acknowledgments
This study was supported by the «Taxonomic
revision of polymorphic plant families of the
flora of Uzbekistan» (FZ-20200929321). We
are greatly indebted to Theodor C.H. Cole (FU
Berlin, Germany) for English language editing
and for valuable comments. All photos are from
the private archive of O.V. Cherneva.
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Obidjon Turdiboev