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New national and regional biological records for Finland 4. Additions to the knowledge of Finnish aphyllophoroid funga (Basidiomycota)

New national and regional biological records for Finland 4. Additions to the knowledge of Finnish aphyllophoroid funga (Basidiomycota)

Profile image of Krister KarttunenKrister Karttunen

Memoranda - Societatis pro Fauna et Flora Fennica

Byssocorticium efibulatum is reported as new to Finland. New records of following rare or little collected species are presented and notes of their substrates are given: Cristinia gallica, Helicogloea farinacea, Helicogloea lagerheimii, Phlebia femsioeensis, Sistotrema dennisii and Tomentella coerulea. We also list 33 aphyllophoroid fungi as new to some section of the boreal vegetation zone in Finland.

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ISSNFennica 0373-6873 al. • Memoranda Soc. Fauna Flora 94,(print) 2018 Memoranda Soc. Fauna Flora Fennica 94:Juutinen 84–99. et2018 84 Helsinki 21 November 2018 ISSN 1796-9816 (online) New national and regional biological records for Finland 9. Contributions to Bryophyta and Marchantiophyta 8 Riikka Juutinen, Tauno Ulvinen, Timo Kypärä, Sanna Huttunen, Katariina Vuorinen, Tuomo Kuitunen, Ari Parnela, Terhi Korvenpää, Kimmo Syrjänen, Kati Pihlaja, Heribert Köckinger, Nadya Konstantinova, Annikki Kestilä, Esa Ervasti, Xiaolan He, Teppo Häyhä, Krister Karttunen, Sanna Laaka-Lindberg, Jyrki Torniainen, Outi Vainio, Harri Vasander, Esko Vuorinen Juutinen, R., Metsähallitus, Parks and Wildlife Lapland, P.O. Box 8016, FI-96101, Rovaniemi; email: riikka.juutinen@metsa.fi. Ulvinen, T., Botanical Museum, Biodiversity Unit, P.O. Box 3000, FI-90014, University of Oulu. Kypärä, T., Tilkkulantie 23, FI-41900, Petäjävesi. Huttunen, S., Botanical museum, FI-20014, University of Turku. Vuorinen, K., University Museum, Norwegian University of Science and Technology (NTNU), NO-7491 Trondheim, Norway. Kuitunen, T., Nystyrämäentie 12, FI-40420, Jyskä. Parnela, A., Kulmalantie 18, FI-35300, Orivesi. Korvenpää, T., Metsähallitus, Parks and Wildlife Coastal and Metropolitan Area, Kärsämäentie 8, FI-20300, Turku. Syrjänen, K., Finnish Environment Institute SYKE, P.O. Box 140, FI00251, Helsinki. Pihlaja, K., Botanical Museum, FI-20014, University of Turku. Köckinger, H., Roseggergasse 12, A-8741, Weisskirchen, Austria. Konstantinova, N., N. A. Avrorin Polar-Alpine Botanical Garden, Institute of the Kola Sci Center RAS, Fersmana 18a, RU-184209, Apatity, Russia. Kestilä, A., Botanical Museum, Biodiversity Unit, P.O. Box 3000, FI-90014, University of Oulu. Ervasti, E., Vasarasepäntie 38, FI-10330, Pinjainen. He, X., Botany Unit, Finnish Museum of Natural History, University of Helsinki, P.O. Box 7, FI-00014, Helsinki. Häyhä, T., Niementie 80, FI-19600, Hartola. Karttunen, K., Finnish Environment Institute SYKE, P.O. Box 140, FI-00251, Helsinki. Laaka-Lindberg, S., Botany Unit, Finnish Museum of Natural History, University of Helsinki, P.O. Box 7, FI-00014, Helsinki. Torniainen, J., Open Science Centre, P.O. Box 35, FI-40014, University of Jyväskylä. Vainio, O., Kuopio Natural History Museum, Myhkyrinkatu 22, FI-70100, Kuopio. Vasander, H., Department of Forest Sciences, P.O. Box 27, FI-00014, University of Helsinki. Vuorinen, E., Heikinkatu 4, FI-10300, Karjaa. Five species of mosses (Bryophyta: Oncophorus demetrii, Oncophorus integerrimus, Oxyrrhynchium speciosum, Sphagnum divinum and Sphagnum medium) and one of liverworts (Marchantiophyta: Kurzia trichoclados) are presented as new for Finland. Seligeria calcarea and Seligeria pusilla, previously thought to be regionally extinct from Finland, are reported to being found again. New records in biogeographical provinces for 74 species of mosses and 36 species of liverworts are listed. Finally, 17 occurrences in provinces are removed due to misidentifications or missing specimens. Memoranda Soc. Fauna Flora Fennica 94, 2018 • Juutinen et al. Introduction We present here 180 new records of bryophyte taxa for country and for Finnish biogeographical provinces that have come to our attention during 2017-2018. The publication is eighth in line after year 2010 (Ulvinen & Syrjänen 2010, 2011a, Ulvinen et al. 2012, Juutinen et al. 2013, Juutinen & Ulvinen 2014, Juutinen et al. 2015, Juutinen et al. 2016). Results are based on field work, on herbarium studies and on complete checkup of two databases. Several people have made new records on the field and others have confirmed determinations. Tauno Ulvinen has revised Finnish herbarium material of Oncophorus. Sanna Huttunen has revised material of Rhynchostegium riparioides and together with Katariina Vuorinen material of Fissidens bryoides, F. gymnandrus and F. viridulus (Huttunen et al. 2018). According to their observations, these taxa cannot be reliably distinguished from each other with morphological characters that are currently used in identification keys. Below we follow their treatment and recognize these taxa at variety level. Riikka Juutinen has compared the Hertta Eliölajit threatened species database of the Finnish Environmental Administration (content on 25.8.2018) and the Finnish Biodiversity Info Facility (FinBIF, https://laji.fi/en), including Collection Management System Kotka of Finnish herbaria (content on 22.9.-14.10.2018, depending on taxa) with previous distribution tables of Finnish bryophytes (Juutinen & Ulvinen 2017). It has been suggested, that the information on species occurrence in Finnish biogeographical provinces or other areas could be directly derived from databases. However, because of frequent misidentifications both in reported observations and in herbarium specimens, reliable information on species’ distribution will require that bryophyte experts must assess each record before they are accepted. Finally, personnel of herbaria have digitized specimens appearing in this paper and provided information on specimens that are not yet digitized. Harri Vasander came up with the Finnish name for Sphagnum divinum. Riikka Juutinen has compiled all the records. We present six species as new for Finland and report two regionally extinct (RE) species to being found again. Five of the new species are re- 85 sults of a recent splitting. Tauno Ulvinen has redetermined Oncophorus specimens in Finnish herbaria after Lars Hedenäs (2017, 2018) published one new combination and one new species: Oncophorus demetrii and O. integerrimus. Sphagnum divinum and S. medium are results of splitting what was formerly known as S. magellanicum (Hassel et al. 2018). And finally, Oxyrrhynchium speciosum is separated from Rhynchostegium riparioides (Huttunen in prep.). In addition, we present 17 corrections for old regional records. These are mainly corrections of misidentifications or deletions based on absent specimens. Updated distribution tables of all Finnish bryophytes can be found from the Internet (http:// www.ymparisto.fi, Luonto/Lajit/Lajiensuojelutyo/Eliotyoryhmat/Sammaltyoryhma/Suomen_sammalet). Nomenclature in this paper follows Hodgetts (2015) and Söderström et al. (2016), except for the newly split species (Hassel et al. 2018, Hedenäs 2017, 2018, Huttunen et al. 2018). We have only accepted records based on public herbarium specimens. Observations based on field determinations or specimens in private herbaria cannot be confirmed and we regard them uncertain. Main source of information now and in the future will be labelled dried herbarium specimens, which provide invaluable archive for scientific studies including biogeography, ecology, morphological and genetic taxonomy. We had to leave out some interesting and valuable bryophyte observations, including Acaulon muticum (previously regionally extinct RE) and Oleolophozia perssonii in Regio aboënsis (Hertta Eliölajit database 2.11.2018), Cephaloziella elachista in Satakunta (Laaka-Lindberg & Kalinauskaitė 2014), Crossocalyx hellerianus in Karelia ladogensis (Syrjänen et al. 2004) and Grimmia alpestris in Regio kuusamoënsis (Juutinen et al. 2017) because of missing specimens. These will be added later if specimens are deposited in public herbaria. Most of the specimens presented here are digitalized in Kotka Collection Management System and their label data is freely accessible using HTTP URI identifiers. Because specimen data is already accessible through FinBIF, records were not digitalized in Hertta Eliölajit threatened species database, which was done as standard procedure in editions 1-7 of new bryophyte records 86 Juutinen et al. • Memoranda Soc. Fauna Flora Fennica 94, 2018 for Finland. Majority of the labels are originally written in Finnish and are translated into English here. Additions to original label data are marked with square brackets. In many cases, what is original label data, is inferred from how things are written in Kotka. For Cephalozia, Fuscocephaloziopsis, Cephaloziella and Nardia double blind determination was mostly required due to the large proportion of misidentifications in herbarium specimens (Ryömä et al. 2013). Coordinates are given in ETRS-TM35-FIN (EUREF-FIN), Uniform Coordinate System (UCS) or WGS84 degrees. Abbreviations for herbaria follow Index Herbariorum (Thiers 2018): H= Botanical Museum, Finnish Museum of Natural History, University of Helsinki, JYV= Natural History Collections of Jyväskylä University Open Science Centre, KUO= Kuopio Natural History Museum, OULU= Botanical Museum, University of Oulu, TUR= Herbarium, University of Turku. New records – Bryophyta Aloina rigida (Hedw.) Limpr. FINLAND. Savonia borealis (Sb/PS), Siilinjärvi, Kuuslahti, Jaakonmäki hill, W slope. Rocky slope. UCS 7003: 3537. Coll. Reino Fagerstén, 19.8.1974 (KUO http://tun.fi/ SBR.58259). Anomodon viticulosus (Hedw.) Hook. & Taylor FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Pitkäpohja, Kalliosaari, cliff E from mountain, N part. Between boulders and on small cliffs. Abundant. UCS 68392:36465. Coll. Tuomo Kuitunen 3563, 17.VII.2018, conf. Ari Parnela 2018 (OULU http://id.herb.oulu.fi/GAP.49). Barbula convoluta Hedw. FINLAND. Lapponia enontekiensis (Le/EnL), Enontekiö, Hetta church village, Hetta church S. Tiled stone stairs of the church, S exposition, on mineral soil. Scarce, small patches and single shoots among other bryophytes. EUREF-FIN 7589485:361609. Coll. Timo Kypärä 2651, 1.IX.2018 (OULU http://tun.fi./GAT.736). Brachythecium mildeanum (Schimp.) Schimp. FINLAND. Lapponia inarensis (Li/InL), Utsjoki, Mantojärvi, Church Hut Museum. Shore. UCS 77538:35005. Coll. Juhani Räsänen 16/928s, 28.VI.2016, det. Tauno Ulvinen 2017, conf. Sanna Huttunen 2017 (OULU http:// id.herb.oulu.fi/GAL.850). Bryum tenuisetum Limpr. FINLAND. Tavastia australis (Ta/EH), Pälkäne, Kortteenpohja, Haukijärvi. Sandy bank of a forest road, open and sunny spot. Relatively abundant. UCS 68103: 33796. Coll. Tuomo Kuitunen 3409, 3.VIII.2017, conf. Kimmo Syrjänen 2018 (OULU http://id.herb.oulu.fi/GAP.50). Campyliadelphus chrysophyllus (Brid.) R.S.Chopra FINLAND. Karelia australis (Ka/EK), Miehikkälä, Hauhia, N from Muhikko, W shore of Vaalimaanjoki river, close to Vuorikoski rapids. At foot of a rock wall. UCS 6735:3536. Coll. Reino Fagerstén, 11.IX.1971 (KUO http://tun.fi/SBR.32964). Cirriphyllum piliferum (Hedw.) Grout FINLAND. Lapponia kittilensis (Lkk/KiL), Muonio, Pallastunturit, Rihmakuru gorge, SW part. Herb-rich forest in a slope, N exposition. UCS 7560:3377. Coll. Risto Virtanen, 15.VI.1990, det. Risto Virtanen 2007 (OULU http:// id.herb.oulu.fi/0025490). Cnestrum alpestre (Wahlenb. ex Huebener) Nyholm ex Mogensen FINLAND. Karelia borealis (Kb/PK), Juuka, Tahkovaara, northern gorge of Pirunkirnu. Rocky slope, under a protruding rock outcrop. Scarce, a few cm2. EUREF-FIN 7007279:605183. Coll. Timo Kypärä 2468, 13.IX.2017 (OULU http://tun.fi/GAT.551). Cratoneuron filicinum (Hedw.) Spruce FINLAND. Savonia borealis (Sb/PS), Kuopio, Vuorimäki, upper part of Pölkkypuro stream dell, NW from stream curve. In a spring. UCS 69751:35310. Coll. Reino Fagerstén, 23.VIII.2002 (KUO http://tun.fi/SBR.58083). Dichelyma capillaceum (L. ex Dicks.) Myrin FINLAND. Lapponia sompiensis (Lks/SoL), Sodankylä, Mataraoja brook, upstream the mine, but very close. 1–3 m wide, shaded and stony small brook. EUREF-FIN 7510644:497007. Coll. Minna Kuoppala 24.VIII.2017, det. Krister Karttunen 2018, conf. Kimmo Syrjänen 2018 (H http://id.luomus.fi/HA.H4260297). Dichodontium palustre (Dicks.) M.Stech FINLAND. Ostrobottnia media (Om/KP), Pyhäjärvi, Haapamäki, Rimpimäki, bottom part of Maaselänlahti bay, shore line close to water works. Sandy shore, on mineral soil. Several small patches in an area of 10 m2. EUREFFIN 7041271:453099. Coll. Timo Kypärä 2675, 18.IX. 2018 (OULU http://tun.fi./GAT.760). Dicranella grevilleana (Brid.) Schimp. FINLAND. Tavastia australis (Ta/EH), Orivesi, Eräjärvi, Vihasjärvi, Tiirikainen N, between road and Längelmäve- Memoranda Soc. Fauna Flora Fennica 94, 2018 • Juutinen et al. si lake. Edge of herb rich forest strip, side of a field ditch. UCS 682757:337309. Coll. Ari Parnela, 6.V.2016, det. Ari Parnela 2017 (OULU http://id.herb.oulu.fi/GAL.2103). Dicranella subulata (Hedw.) Schimp. FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Särkisalmi, Oronmylly. Sandy side of a forest road. Scarce. UCS 683735:362624. Coll. Tuomo Kuitunen 3377, 17. VII.2017 (OULU http://id.herb.oulu.fi/GAL.2313). Dicranum fragilifolium Lindb. FINLAND. Savonia australis (Sa/ES), Sulkava, Partalansaari island, P. Isoviita summer place. [UCS 684:357] Coll. Reino Fagerstén, 15.VII.1988 (KUO http://tun.fi/SBR.58056). Dicranum laevidens R.S.Williams FINLAND. Lapponia enontekiensis (Le/EnL), Enontekiö, Kilpisjärvi, Malla Strict Nature Reserve, Iso-Malla (Gihsibákti), SE slope. Oligotrophic Sphagnum fuscum fjell mire, among Sphagnum. Patches, scarce and occasional. UCS 7678986:3249003. Coll. Kimmo Syrjänen, 29.VII. 2016 (TUR http://mus.utu.fi/TBR.122136). Dicranum leioneuron Kindb. FINLAND. Savonia borealis (Sb/PS), [Pieksämäki] Jäppilä, Suonpohjansuo mire. Papillosum fen, severely drained, on hummock. UCS 692838:351443. Coll. Teppo Häyhä, 19.VI.2000, conf. Tauno Ulvinen 2018 (OULU http:// id.herb.oulu.fi/HT.24606). Didymodon fallax (Hedw.) R.H.Zander FINLAND. Tavastia borealis (Tb/PH), Viitasaari, Haapasaari, Viitasaari church, next to churchyard main gate, NE side. Massive, masoned stone fence, on concrete. Few 20– 30 cm2 sized patches in a stretch of two meters. EUREFFIN 6994180:442023. Coll. Timo Kypärä 2557, 5.VII. 2018, conf. Tauno Ulvinen 10/2018 (OULU http://tun.fi./ GAT.642). Didymodon ferrugineus (Schimp. ex Besch.) M.O.Hill FINLAND. Tavastia australis (Ta/EH), Heinola, Mataraniemi-Kiistasniemi, Mäyrämäki, calcareous embankments W from nature protection area. An open steep calcareous meadow slope between two embankments. UCS 6787761: 3453620. Coll. Kimmo Syrjänen, 9.VIII.2016 (TUR http:// mus.utu.fi/TBR.122138). Ditrichum flexicaule (Schwägr.) Hampe FINLAND. Ostrobottnia australis (Oa/EP), Teuva, Luovankylä, Iso-Parra S, middle part of Sikakallio rocks. Cave at foot of a rock face. Probably relatively scarce. EUREF-FIN 6930701:233063. Coll. Timo Kypärä 2427, 16. VIII.2017 (OULU http://tun.fi/GAT.510). 87 Ditrichum zonatum (Brid.) Kindb. FINLAND. Savonia borealis (Sb/PS), Kuopio, Nilsiä, Lastukoski, N end of Pisa hill, Pirunkellari E. Rock face. Relatively abundant, a few dm2. EUREF-FIN 7011643: 566457. Coll. Timo Kypärä 2382, 24.VII.2017 (OULU http://tun.fi/GAT.465); Ostrobottnia kajanensis (Ok/Kn), Suomussalmi, Hossa National Park, S shore of lake Somerjärvi, near Värikallio. Rock face, moist crack, NW exposition. Relatively scarce, a few cm2. EUREF-FIN 7267928: 605456. Coll. Timo Kypärä 2326, 14.VI.2017 (OULU http://tun.fi/GAT.409). Drepanocladus sordidus (Müll.Hal.) Hedenäs FINLAND. Alandia (Al/A), Eckerö, Storby, Posthuset W, S side of a dirt road leading to Postbryggan. Moist Carex panicea depression. UCS 6701955:3087261. Coll. Kimmo Syrjänen, 2.IX.2001 (TUR http://mus.utu.fi/TBR.119873). Encalypta procera Bruch FINLAND. Nylandia (N/U), Helsinki, Västersundom, Tryvik. Picea abies -dominated herb-rich forest, abandoned lime quarry, on wall. UCS 668144:339842. Coll. Juha Pykälä 32286, 18.I.2008. (H http://id.luomus.fi/HA.H4258248). Fissidens bryoides Hedw. var. bryoides (Fissidens bryoides) FINLAND. Nylandia (N/U), Hfors [Helsinki] Humleberg [Humalisto]. On sandy soil on high river bank. [UCS 6676:3384] Coll. Alfred Oswald Kihlman & Ragnar Hult, 17.V.1877, det. Katariina Vuorinen & Sanna Huttunen 2017 (KUO http://tun.fi/SBR.4805); Savonia borealis (Sb/ PS), Kuopio, Puijo, (plot no. 10). On mineral ground at a base of rich forest slope. [UCS 6980: 3534] Coll. Matti Haapasaari 3278 & Reino Fagerstén, 30.VI.1976, det. Katariina Vuorinen & Sanna Huttunen 2017 (KUO http:// tun.fi/SBR.50069); Karelia borealis (Kb/PK), Tohmajärvi, Akkala, Hiidenvaara NE slope. Steep, shaded cliff. UCS 6903: 3671. Coll. Matti Haapasaari & Ilkka Kytövuori, 25.V.1964, det. Katariina Vuorinen & Sanna Huttunen 2017 (KUO http://tun.fi/SBR.21416); Ostrobottnia kajanensis (Ok/Kn), Puolanka, Hepoköngäs, Heinijoki river. Riverside, cavity in a rock wall. Scarce, single shoots among other bryophytes. UCS 7191924: 3541430. Coll. Timo Kypärä 1696 & Niina Onttonen, 21.VIII.2014, det. Sanna Huttunen 2018 (JYV http://tun.fi/LC.20654); Ostrobottnia ultima (Obu/PeP), Tervola, river Louejoki, Myllyköngäs rapids. [UCS 7347:3413] Coll. Mauno J. Kotilainen, 22.VIII.1924, det. Katariina Vuorinen & Sanna Huttunen 2017 (TUR http://mus.utu.fi/TBR.12412); Regio kuusamoënsis (Ks/Ks), Kuusamo, river Kitkajoki, Juuma gorges. [WGS84 66.27153 °N 29.39135 °E] Coll. Viktor Ferdinand Brotherus, 22.VII.1883, det. Katariina Vuorinen & Sanna Huttunen 2017 (H http://id.luomus.fi/HA.H4065672). See also Huttunen et al. (2018). 88 Juutinen et al. • Memoranda Soc. Fauna Flora Fennica 94, 2018 Fissidens bryoides Hedw. var. gymnandrus (Buse) R. Ruthe (F. gymnandrus) FINLAND. Tavastia australis (Ta/EH), Urjala, Menonen, Kivijärvi nature protection area. Swamp depression in a mature forest, on stones and sides of depression. EUREF-FIN 6769995:3307981. Coll. Turkka Korvenpää, 17.VIII.2010, conf. Katariina Vuorinen & Sanna Huttunen 2017 (TUR http://mus.utu.fi/TBR.116455); Karelia borealis (Kb/PK), Tohmajärvi, Kemie, Musko, Havukkavaara E slope, lower part. Picea abies dominated forest, from steep, shaded rock faces. [UCS 6908:670] Coll. Matti Haapasaari 4056, 30.V.1966, det. Katariina Vuorinen & Sanna Huttunen 2017 (KUO http://tun.fi/SBR.21420); Lapponia enontekiensis (Le/EnL), Enontekiö, Markkina, river Jietajoki (Hietajoki). In terra inundata arenosoturfosa. [WGS84 68.53 °N 22.39 °E] Coll. Ritva Soini, 3.VIII.1966, det. Katariina Vuorinen & Sanna Huttunen 2017 (H http://id.luomus.fi/HA.H4065686). See also Huttunen et al. (2018). UCS 681774:362155. Coll. Tuomo Kuitunen 3559, 19. VII.2018, conf. Ari Parnela 2018 (OULU http://id.herb.oulu. fi/GAP.52). Hygrohypnum smithii (Sw.) Broth. FINLAND. Ostrobottnia kajanensis (Ok/Kn), Suomussalmi, Hossa National Park, Korkeakangas, Somerjoki river, Lihapyörre NW. River rock under the waterline. Probably scarce, seen a few shoots. EUREF-FIN 7267594:608621. Coll. Timo Kypärä 2344, 15.VI.2017, conf. Tauno Ulvinen 2017 (OULU http://tun.fi/GAT.427). Hypnum andoi A.J.E.Sm. FINLAND. Karelia borealis (Kb/PK), Kontiolahti, Kyykkä, Kirveskallio, lower part of W slope of forested hill, S end of rock face. Crack of a rock, quartzite. UCS 69752: 36503. Coll. Juhani Räsänen 16/1846s, 31.VIII.2016, det. Tauno Ulvinen 2017 (OULU http://id.herb.oulu.fi/GAL.919). Fontinalis dichelymoides Lindb. FINLAND. Savonia australis (Sa/ES), Puumala, Rokansalo, Syrjälampi pond. Oligo-dystropic pond, submerged. UCS 68275:35559. Coll. Teppo Häyhä, 11.VI.2001 (TUR http://mus.utu.fi/TBR.100595). Grimmia elatior Bruch ex Bals.-Criv. & De Not. FINLAND. Tavastia borealis (Tb/PH), Konnevesi, Siikakoski. UCS 6946:3466. Coll. Veli Saari, 27.VI.1996 (JYV http://tun.fi/LC.20775). Grimmia ovalis (Hedw.) Lindb. FINLAND. Lapponia enontekiensis (Le/EnL), Enontekiö, Porojärvet, Jokashpakti. [UCS 7684:3283] Coll. Laila Ollila & Heikki Roivainen, 29.VII.1955, det. Tauno Ulvinen 2013 (H http://id.luomus.fi/HA.H4070523). Grimmia unicolor Hook. FINLAND. Satakunta (St/St), Eura, Haveri, Sieravuori, Pyhäjärvi. Flat, open rock shore, relatively close to waterfront, on cracks. UCS 67743:32424. Coll. Ari Parnela, 13. X.2012 (OULU http://id.herb.oulu.fi/0035695). Herzogiella seligeri (Brid.) Z.Iwats. FINLAND. Karelia borealis (Kb/PK), Joensuu, Linnunlahti, Vainolahti, between two paths leading from kindergarten to Vainolahti bay. SE end of a shaded depression, on rotten wood. UCS 694724: 363995. Coll. Juhani Räsänen 16/314s b, 17.IV.2016, det. Tauno Ulvinen 2017 (OULU http://id.herb.oulu.fi/GAL.825). Hygrohypnum ochraceum (Turner ex Wilson) Loeske FINLAND. Karelia ladogensis (Kl/LK), Rautjärvi, Koskenkulma, Lohioja. On rapid stones. Abundant, as a mat. Hypnum recurvatum (Lindb. & Arnell) Kindb. FINLAND. Savonia australis (Sa/ES), Savonlinna, Punkaharju, Hiukkajoki, Hannula 400 m E. Abandoned lime quarry, on calcareous boulders. UCS 6855495:3634899. Coll. Juha Pykälä 47636, 9.IX.2014 (H http://id.luomus.fi/ HA.H4258252). Leucodon sciuroides (Hedw.) Schwägr. FINLAND. Karelia lagodensis (Kl/LK), Parikkala, Pitkäpohja, Kalliosaari, cliff E from mountain, N part. Between boulders and on small cliffs. Scarce. UCS 68390:36465. Coll. Tuomo Kuitunen 3564, 17.VII.2018, conf. Ari Parnela 2018 (OULU http://id.herb.oulu.fi/GAP.53). Loeskypnum badium (Hartm.) H.K.G.Paul FINLAND. Savonia australis (Sa/ES), Juva, mire on W shore of lake Lentunen. Mesotrophic tall-sedge pine fen, flark-intermediate surface. UCS 68391:35540. Coll. Teppo Häyhä, 25.VII.2000 (OULU http://id.herb.oulu.fi/HT.24646). Myurella julacea (Schwägr.) Schimp. FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Intsilä, Väntsävaara. Ultramafic rock outcrop, crack in a vertical rock face. Very scarce. UCS 683356:363433. Coll. Tuomo Kuitunen 3403, 20.VII.2017, conf. Ari Parnela 2017 (OULU http://id.herb.oulu.fi/GAL.2317). Neckera pumila Hedw. FINLAND. Nylandia (N/U), Kirkkonummi, Laitamaa, Kakarsberget. Cliff, W exposition, shaded oligotrophic rock wall, on vertical crack. UCS 6683106:3361095. Coll. Esa Ervasti 1181 & Esko Vuorinen, 10.IX.2018 (TUR http://mus.utu.fi/TBR.124257). Memoranda Soc. Fauna Flora Fennica 94, 2018 • Juutinen et al. Oncophorus demetrii (Renauld & Cardot) Hedenäs, ryhmytihkusammal, fjällknölmossa FINLAND. Regio kuusamoënsis (Ks/Ks), Kuusamo, Juuma, Jäkälävuoma gorge. UCS 735[3]:360[9]. Coll. Timo Koponen 2381, 26.VI.1964, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/HA.H4098107); Lapponia enontekiensis (Le/EnL), NW-Le [Enontekiö] Kilpisjärvi, SW-Saana. Paludified place. [UCS 7674:3254] Coll. Laila & Heikki Roivainen, 14.VII.1958, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/HA.H4098231); Lapponia inarensis (Li/ InL), Utsjoki, Kistuskaidi, N slope. Mountain birch zone, on a wet cliff. [WGS84 69.93°N 26.52 °E] Coll. Yrjö Mäkinen, 26.VII.1956, det. Tauno Ulvinen 2018 (TUR http://mus.utu.fi/TBR.42703). New species for Finland. Recently presented as new for Scandinavia (Hedenäs 2018). It’s type locality is in Canada and it has probably a circumpolar distribution. The species grows in northern part of Fennoscandia on moist mountain heaths, wet meadows, fens, wet rocks, brooks and river shores. It is more calciphilous than other Oncophorus species. Oncophorus elongatus (I.Hagen) Hedenäs FINLAND. Alandia (Al/A), Eckerö, Torp, Degersand. Low-lying, flat seashore rock surface, in a crack. UCS 6694061:3089881. Coll. Kimmo Syrjänen, 1.X.2001, det. Tauno Ulvinen 2018 (TUR http://mus.utu.fi/TBR.119923); Regio aboënsis (Ab/V), [Salo] Perniö, Pohjankylä, E from Pohjanjärvi lake. On rotten wood in a moist rock slope. [UCS 6692:3286] Coll. Unto Laine, 22.VIII.1966, det. Tauno Ulvinen 2018 (TUR http://mus.utu.fi/TBR.88785); Nylandia (N/U), Espoo, Metsämaa, stream running from Kringelkärr mire. On rotten wood. [UCS 6691:3376] Coll. Timo Koponen, 12.X.1964, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/HA.H4097889); Karelia australis (Ka/EK), [Hamina] Vehkalahti, Onkamaa, Pirijoki. [UCS 673:352] Coll. Lars Fagerström, 10.VI.1972, det. Tauno Ulvinen 2018 (KUO http://tun.fi/SBR.18490); Satakunta (St/St), Kankaanpää, Luomajärvi, Vääräneva. Margin of a mire. [UCS 6866:3239] Coll. Matti Laurila, 17.VI.1935, det. Tauno Ulvinen 2018 (TUR http://mus.utu.fi/TBR.42264); Tavastia australis (Ta/EH), Orivesi, Pukala, Neejärvi, a stream running from Neejärvi lake, rapids between road and Isolahti bay in Pukala. On rotten wood in a stream. UCS 68454:33512. Coll. Ari Parnela, 5.X.2006, det. Tauno Ulvinen 2018 (OULU http://herb.oulu.fi/GAP.59); Savonia australis (Sa/ES), [Joutsa] Leivonmäki, along the road 5 km from Rutalahti to highway 4, brook crossing. Brooksides. UCS 6875:3450. Coll. Reino Fagerstén 15183, 29.IX.1980, det. Tauno Ulvinen 2018 (KUO http:// tun.fi/SBR.54752); Karelia ladogensis (Kl/LK), [Parikkala] Uukuniemi. [UCS 685:3365] Coll. [Theodor] Simming, 22.VII.1861, det. Tauno Ulvinen 2018 (H http://id.luomus. fi/HA.H4097956); Ostrobottnia australis (Oa/EP), Kauhajoki, Muurahainen, Paholuoma, Ruostetkangas, S from Ruoste- 89 toja. Rotten log that has fallen to the river. Scarce, c. 20 cm2. EUREF-FIN 6901130:254085. Coll. Timo Kypärä 2233, 16.IX.2016, det. Tauno Ulvinen 2018 (OULU http://tun.fi/ GAT.315); Tavastia borealis (Tb/PH), Ähtäri, Keskinen Sappio, NE shore. Boulder field (”pirunpelto”) behind shore cliffs, in a cavity between rocks. UCS 69494:33470. Coll. Ari Parnela & Harri Arkkio, 26.V.2016, det. Tauno Ulvinen 2018 (OULU http://id.herb.oulu.fi/GAL.1131); Savonia borealis (Sb/PS), Tuusniemi, Seinävuori, Pieni Seinälampi pond. On walls of the rocky gorge, bottom of the gorge. UCS 6975:3574. Coll. Reino Fagerstén 4017, 17.VI.1976, det. Tauno Ulvinen 2018 (KUO http://tun.fi/SBR.33979); Karelia borealis (Kb/PK), Ilomantsi, Käenkoski. NE from turnaround of a forest road, east from south projecting part of Leppäsuo mire. On rotten wood close to a pond. UCS 698630:370814. Coll. Juhani Räsänen 17/931s and Leena Pulkkinen, 2.VIII.2017, det. Tauno Ulvinen 2018 (OULU http://id.herb.oulu.fi/GAL.3539); Ostrobottnia media (Om/KP), [Kokkola] Kälviä, Renögrund [Krunni]. On sea shore rocks. [UCS7103:3311] Coll. A.V. Auer, 5.8.1933, det. Tauno Ulvinen 2018 (TUR http://mus.utu.fi/TBR.18495); Ostrobottnia kajanense (Ok/Kn), Vaala, Manamansalo, Martinkanta, NW from the road. Mesotrophic fen, in a hummock at the side of a flark. Abundant. UCS 71450:35070. Coll. Tauno Ulvinen, 12.VIII.1989, det. Tauno Ulvinen 2018 (OULU http://id.herb.oulu.fi/0037530); Ostrobottnia ouluensis (Obo/OP), Pudasjärvi, Korpinen, W from Pieni Huosioislampi pond, a stream running from Ollinlampi pond by the path. Side of a stream, on rotten stump. UCS 72309: 35434. Coll. Tauno Ulvinen, 14.VI.2000, det. Tauno Ulvinen 2018 (OULU http://id.herb.oulu.fi/0037531); Ostrobottnia ultima (Obu/PeP), Ranua, Hosio, Laitakarinoja. Moist forest by the side of brook, on rotten wood. UCS 7313:3448. Coll. Tauno Ulvinen, 17.VII.1972, det. Tauno Ulvinen 2018 (OULU http://id.herb.oulu.fi/GAP.58); Regio kuusamoensis (Ks/Ks), Taivalkoski, Kirvesoja brook, rocky rapids by the edge of a Picea abies forest. Side of a brook. UCS 7248775:3558919. Coll. Risto Virtanen, 19. IX.2007, det. Tauno Ulvinen 2018 (OULU http://id.herb. oulu.fi/0001116). Oncophorus integerrimus (I.Hagen) Hedenäs, silotihkusammal, slät skruvknölmossa FINLAND. Alandia (Al/A), Vårdö, Töftö. In a moist place. [UCS 6698:3129] Coll. Paul Olofsson, 16.IX.1945, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/HA.H4098396); Ostrobottnia kajanensis (Ok/Kn), Suomussalmi, Kianta village, Portti (between Salov[aara] and Saarijärvi). Marshy place. [UCS 7241:3606] Coll. Olli Kyyhkynen, 2.VIII.1917, det. Tauno Ulvinen 2018 (H http://id.luomus. fi/HA.H4098407); Otrobottnia ouluensis (Obo/OP), Pudasjärvi, Nuorunka, Nuukavaara. Springy place on a slope in SW-part of hills Näsiäkummut. UCS 7294:3527. Coll. Tauno Ulvinen & Esteri Ohenoja, 24.VII.1973, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/HA.H4098411); Ostrobottnia ultima (Obu/PeP), Tervola, Vähäjoki, Vääräkoski rapids. [UCS7342:3430] Coll. Mauno J. Kotilainen, 18. 90 Juutinen et al. • Memoranda Soc. Fauna Flora Fennica 94, 2018 VIII.1924, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/ HA.H4098416); Regio kuusamoënsis (Ks/Ks), Kuusamo parish, Juuma, Jäkälävuoma gorge. Edge of pool, on peat. UCS 7353:3610. Coll. Aune & Timo Koponen 27665, 26. VIII.1980, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/ HA.H4098469); Lapponia kittilensis (Lkk/KiL), Kittilä, Tepsa, Lomajärvi, between Keski- and Ala-Lomajärvi. Eutrophic fen depression in a valley, calcareous ground. UCS 75083448. Coll. Tauno Ulvinen, 13.IX.1983, det. Tauno Ulvinen 2018 (OULU http://id.herb.oulu.fi/GAP.67); Lapponia sompiensis (Lks/SoL), Sodankylä, Rajala, Pahtakoski rapids in Saittajoki river, c. 300 m E from cottage on N side of river. Rich fen, in a moist spruce fen passage. UCS 75036:34663. Coll. Tauno Ulvinen, 17.VII.1996, det. Tauno Ulvinen 2018 (OULU http://id.herb.oulu.fi/GAP.68); Lapponia enontekiensis (Le/EnL), Enontekiö, Kilpisjärvi, W Saana. [UCS 7675:3259] Subalpine herb rich forest. On stone in a rivulet. Coll. Matti Haapasaari 2121, 17.VII. 1964, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/ HA.H4098567); Lapponia inarensis (Li/InL), [Inari] Laanila, 3 km north of Mt. Kaunispää. Along brook in Betula tortuosa forest. [UCS 7597:3519] Coll. Timo Koponen 26506, 29.VII.1975, det. Tauno Ulvinen 2018 (H http://id.luomus. fi/HA.H4098627). New species for Finland. Recently presented as new for Scandinavia (Hedenäs 2017). Majority of specimens earlier named as O. virens belong to this species. Distribution is circumpolar. Additionally, there are specimens collected from the Middle European mountains. In Finland it has a separate locality in Ahvenanmaa (Al/Al), but the main area of distribution is in the northern part of the country north of Kainuu (Kn/Kn). It grows most often in open wetlands, eutrophic fens, moist meadows and shores. Orthotrichum affine Schrad. ex Brid. FINLAND. Ostrobottnia australis (Oa/EP), Teuva, Luovankylä, Iso-Parra S, middle part of Sikakallio rocks. Top part of a fallen aspen. Probably relatively scarce, a few tufts. EUREF-FIN 6930684:233049. Coll. Timo Kypärä 2429, 16.VIII.2017 (OULU http://tun.fi/GAT.512); Karelia borealis (Kb/PK) Ilomantsi, Palopohja, Pötönkangas graveyard. On aspen in the middle of the graveyard. Scarce, one tuft, some cm2. EUREF-FIN 6956301:701748. Coll. Timo Kypärä 2404, 31.VII.2017, conf. Kimmo Syrjänen 2017 (OULU http://tun.fi/GAT.487). Orthotrichum cupulatum Hoffm. ex Brid. FINLAND. Nylandia (N/U), Raasepori, Jussarö E, Österflada SW, hostel yard. On concrete garden tiles. EUREFFIN 6636898:307951. Coll. Timo Kypärä 2437, 30.VIII. 2017 (OULU http://tun.fi/GAT.520). Oxyrrhynchium speciosum (Brid.) Warnst., tursonsammal, strandspötmossa FINLAND. Regio aboënsis (Ab/V), Raasepori, Pohjanpitäjänlahti, Flada. Growing at 1,5 m depth. WGS84 60.05171 °N 23.53702 °E. Coll. Roosa Mikkola & Anna Lyssenko, 7.IX.2016, det. Sanna Huttunen 2017 (TUR http://mus.utu.fi/TBR.121367); Nylandia (N/U), [Raasepori] Tammisaari, Björknäs. [EUREF-FIN 66567:302875] Coll. Hans Luther, 4.IX.1939, det. Sanna Huttunen 2017 (H http://id.luomus.fi/HA.H4130640); Karelia australis (Ka/ EK), Virolahti, Vilkkilä, bay NW from Kirskallionniemi cape (Ruosviikinlahti bay). On loam bottom, c. 50,5 m depth, brackish water. UCS 67146:35395. Coll. Tapio Rintanen, 17.IX.1980, det. Sanna Huttunen 2017 (TUR http:// mus.utu.fi/TBR.107691); Tavastia australis (Ta/EH), Sysmä, lake Päijänne, Majutvesi, Muurisalmi, in lake between the easternmost tip of Pitkäniemi and island Päiväsaari. UCS 6818326:3428049. Coll. Sanna Huttunen 2.VII.2010, det. Sanna Huttunen 2017 (TUR http://mus.utu.fi/TBR.120402); Tavastia borealis (Tb/PH), Konnevesi, Hytölä, Vanginvesi. At the bottom of the lake, at a depth of 3–4 meters. Sparsely in a relatively large area, several hundred m2. EUREF-FIN 6939397:459004. Coll. Timo Kypärä & Johanna Hallman, 6.IX.2015, det. Sanna Huttunen 2017 (TUR http://mus.utu.fi/TBR.121366); Savonia borealis (Sb/ PS), Kuopio, Kolmisoppi lake, E side. Floating on lake bottom 4–6 m deep. EUREF-FIN 6971224:530856. Coll. Krister Karttunen & Pirjo Punju, 17.VI.2009, det. Sanna Huttunen 2017 (TUR http://mus.utu.fi/TBR.106791); Ostrobottnia media (Om/KP), Lappajärvi, Lappajärvi lake, N part, close to Timo house. UCS 7018852:3331320. Coll. Anna-Maria Koivisto & Anssi Teppo, 3.VIII.2009, det. Sanna Huttunen 2017 (OULU http://id.herb.oulu.fi/GAP.69); Ostrobottnia ouluensis (Obo/OP), Oulu, Kintasletto. At 40–60 line. WGS84 65.140115 °N 25.058049 °E. Coll. Jalmari Laurila, 21.VII.2015, det. Sanna Huttunen 2017 (OULU http://id.herb.oulu.fi/GAL.150); Ostrobottnia ultima, (Obu/PeP), Kemi, Sarvenselkä. On rocky bottom. WGS84 65.655626 °N 24.324495 °E. Coll. Essi Keskinen, 1.VIII.2015, det. Sanna Huttunen 2017 (OULU http:// id.herb.oulu.fi/GAL.146); Regio kuusamoënsis, (Ks/Ks), Posio, Kitkajärvi lake, by Suonna village ca. south from mouth of brook Suonna. UCS 7338224:3567134. Coll. Sanna Huttunen, 6.VI.2010, det. Sanna Huttunen 2017 (TUR http://mus.utu.fi/TBR.121362). New species for Finland. Sanna Huttunen is preparing a manuscript about this species. Philonotis caespitosa Jur. FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Joukio, Hotel Laatokanportti, at the shore of the hotel. Thicket of Phragmites australis, on sand. Scarce. UCS 682330:363226. Coll. Tuomo Kuitunen 3401, 19.VII. 2017, det. Ari Parnela 2017, conf. Tauno Ulvinen 2017 (OULU http://id.herb.oulu.fi/GAL.2318). Memoranda Soc. Fauna Flora Fennica 94, 2018 • Juutinen et al. 91 Plagiopus oederianus Pohlia sphagnicola (Bruch & Schimp.) Broth. (Sw.) H.A.Crum & L.E.Anderson FINLAND. Ostrobottnia australis (Oa/EP), Kristiinankaupunki, Lapväärtti, S slope of Pyhävuori mountain. [UCS 692:322] Coll. Matti Haapasaari 6274, 26.X.1965 (KUO http://tun.fi/SBR.7813). FINLAND. Karelia borealis (Kb/PK), Kaavi, Niinivaara, Nevansaari, 70 m NE from the SW side of the open mire. Open mire. UCS 698955:359243. Coll. Juhani Räsänen 17/1063s & Marko Riikonen, 19.VIII.2017, det. Tauno Ulvinen 2018 (OULU http://id.herb.oulu.fi/GAL.3535). Plagiothecium denticulatum (Hedw.) Schimp. var. obtusifolium (Turner) Moore, töppölaakasammal Polytrichum hyperboreum R.Br. FINLAND. Tavastia borealis (Tb/PH). Virrat, Jäähdyspohja, Munkkivuori. UCS 690:34. Coll. Niilo Söyrinki, 5.VIII.1951, det. Tauno Ulvinen 1972 (OULU http:// id.herb.oulu.fi/0011806); Ostrobottnia media (Om/KP). Haapajärvi, Kuona, Matokangas. UCS 707:342. Coll. Heikki Roivainen, 8.VIII.1923, det. Tauno Ulvinen 1998 (OULU http://id.herb.oulu.fi/0011851); Ostrobottnia kajanensis (Ok/Kn). Sotkamo, Haapaniemi. UCS 7118: 3550. Coll. M. Lindqvist, 18.VII.1928, det. Tauno Ulvinen 1967 (OULU http://id.herb.oulu.fi/0011833); Ostrobottnia ouluensis (Obo/OP). [Oulu] Oulujoki, Madekoski, Länsisuo. UCS 720:343. Coll. Martti Ohenoja, 11.VI.1962, det. Tauno Ulvinen 1967 (OULU http://id.herb.oulu.fi/0011876). The taxon is dubious according to Hedenäs (2014) but is included in the Checklist of European bryophytes (Hodgetts 2015). Specimens in other Finnish herbaria besides OULU have not been checked and even specimens presented here are uncertain. The taxon is thought to have a northern distribution. FINLAND. Regio kuusamoënsis (Ks/Ks), Salla, N-end of the top of Pieni Pyhätunturi, sample plot 351. Empetrum – Dicranum – Lichenes heath. [UCS 7409:3578] Coll. Matti Haapasaari, 19.VII.1966 (KUO http://tun.fi/SBR.25136). Polytrichum pallidisetum Funck (Polytrichastrum pallidisetum) FINLAND. Lapponia sompiensis (Lks/SoL), Pelkosenniemi, Pyhä-Luosto National Park, N from fjell Pyhätunturi, mire Tunturiaapa. SphLN – MeVSN. EUREF-FIN 7435175:504961. Coll. Niina Sankari 185/2015, 14.VII. 2015, det. Timo Kypärä 2017 (OULU http://id.herb.oulu.fi/ HT.21135). Pseudoamblystegium subtile (Hedw.) Vanderp. & Hedenäs (Amblystegium subtile) FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Intsilä, Suurmäki, W end. Ultramafic rock outcrop, at foot of a cliff. UCS 683352:364328. Coll. Tuomo Kuitunen 3390, 21.VII.2017 (OULU http://id.herb.oulu.fi/GAL.2308). Pseudotaxiphyllum elegans (Brid.) Z.Iwats. Pohlia annotina (Hedw.) Lindb. FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Tiviä, Lähdelampi pond. Sandy side of a forest road. Very scarce. UCS 68259:36372. Coll. Tuomo Kuitunen 3379, 17.VII.2017, conf. Ari Parnela 2017, Tauno Ulvinen 2017 (OULU http://id.herb.oulu.fi/GAL.2319). FINLAND. Lapponia enontekiensis (Le/EnL), Enontekiö, Ounastunturi, Pahakuru, W part. Rocky low side ravine, in a cavity at the foot on a rock face. C. 2 dm2 sized patch. EUREF-FIN 7571247:373394. Coll. Timo Kypärä 2635, 27.VIII.2018 (OULU http://tun.fi./GAT.720). Psilopilum cavifolium (Wilson) I.Hagen Pohlia elongata Hedw. FINLAND. Savonia borealis (Sb/PS), Vieremä, Riutankallio, Pikku Pöytämäki hill, SW side of a rocky hill near Juortinnotko. Shady rock face of tonalitic gneiss. UCS 707362:350038. Coll. Juhani Räsänen 16/897s, 17.VI. 2016, det. Tauno Ulvinen 2017 (OULU http://id.herb.oulu. fi/GAL.851). FINLAND. Lapponia enontekiensis (Le/EnL), Enontekiö, Käsivarsi wilderness area, E side of fjell Duolljehuhput. Snow bed under a bank, on thin peat. EUREF-FIN 7677380:266490. Coll. Riikka Juutinen 7154, 14.VIII. 2016 (OULU http://tun.fi/GAK.1239, as associated species in Moerckia blyttii specimen). Ptychostomum compactum Hornsch. Pohlia filum (Schimp.) Mårtensson FINLAND. Karelia borealis (Kb/PK), Lieksa, Ruunaa, Juppulansärkkä NE. On moist forest road. UCS 70382:36643. Coll. Juhani Räsänen 16/1277s, 25.VII.2016, det. Tauno Ulvinen 2017 (OULU http://id.herb.oulu.fi/GAL.882). (Bryum algovicum) FINLAND. Karelia borealis (Kb/PK), Kaavi, Niinivaara, W from Onkilampi pond, S end of a rock outcrop, NE exposition. Serpentinite rock wall. UCS 69934:35916, Coll. Juhani Räsänen 16/1715s, 22.VIII.2016, det. Risto Virtanen 2017 (OULU http://id.herb.oulu.fi/GAL.888). 92 Juutinen et al. • Memoranda Soc. Fauna Flora Fennica 94, 2018 Ptychostomum creberrimum Schistidium trichodon (Brid.) Poelt (Taylor) J.R. Spence & H.P. Ramsay (Bryum creberrimum) FINLAND. Karelia australis (Ka/EK), [Kouvola] Sippola, Hirvelä, in Myllyoja brook. Rapids. [UCS 673:350] Coll. Lars Fagerström, 4.VI.1968 (KUO http://tun.fi/SBR.23906). FINLAND. Tavastia borealis (Tb/PH), Petäjävesi, Heikkilänperä, W part of Sydänmaa village. Moss covered, shaded tile roof of an old sauna. EUREF-FIN 6920242:405461. Coll. Timo Kypärä 2304, 3.VI.2017, conf. Tauno Ulvinen 2017 (OULU http://tun.fi/GAT.386). Scorpidium cossonii (Schimp.) Hedenäs Ptychostomum cyclophyllum (Schwägr.) J.R. Spence (Bryum cyclophyllum) FINLAND. Karelia borealis (Kb/PK), Ilomantsi, NW shore of lake Nuorajärvi, where mire Puohtiinsuo meets bay Honkalahti. Partly sunken log on marshy shore. Relatively scarce, scattered in an area of 2 dm2. EUREF-FIN 6960903:706600. Coll. Timo Kypärä 2405, 1.VIII.2017, conf. Kimmo Syrjänen 2017 (OULU http://tun.fi/GAT.488). Rhabdoweisia crispata (Dicks.) Lindb. FINLAND. Nylandia (N/U), Kirkkonummi, Laitamaa, Kakarsberget, N part. Cliff, NW exposition, steep oligotrophic cliff wall (steps, cracks and holes), moist horizontal cracks. Small patches, at least two separate locations. UCS 6683525:3361208. Coll. Esa Ervasti 1188 & Esko Vuorinen, 10.IX.2018 (TUR http://mus.utu.fi/TBR.124259). FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Intsilä, Mikonvaara hill. Bottom of quarry, periodically wet depression, on mining waste. Abundant. UCS 68331:36417. Coll. Tuomo Kuitunen 3573, 18.VII.2018, det. Ari Parnela 2018 (OULU http://id.herb.oulu.fi/GAP.54). Seligeria calcarea (Hedw.) Bruch & Schimp. FINLAND. Regio kuusamoënsis (Ks/Ks), Salla, S end of gorge Kurtinniittykuru, c. 170 m north from border of Oulanka National Park, east slope. C. 6 m high calcareous wall on a slope, on vertical surface. EUREF-FIN 7370604: 596242. Coll. Riikka Juutinen 6944, 14.VI.2016, conf. Risto Virtanen 2016 (OULU http://tun.fi/GAK.1030). Previously assessed in Finland as regionally extinct (RE) (Syrjänen et al. 2010). Seligeria pusilla (Hedw.) Bruch & Schimp. Rhynchostegium riparioides (Hedw.) Card. FINLAND. Lapponia enontekiensis (Le/EnL), Enontekiö, Somashjärvi. [UCS 769:328] Coll. A.J. Huuskonen, 3. VIII.1966, det. Tauno Ulvinen 1999 (OULU http://id.herb. oulu.fi/GAP.63). Rhytidiadelphus subpinnatus (Lindb.) T.J.Kop. FINLAND. Ostrobottnia australis (Oa/EP), Kristiinankaupunki, Tiukka, W slope of Puskavuori hill, Paratiisilehto. [UCS 692:322] Coll. Matti Haapasaari 6297, 24.X.1965, det. Timo Koponen 1974 (KUO http://tun.fi/SBR.9244). Schistidium robustum (Nees & Hornsch.) H.H.Blom FINLAND. Savonia borealis (Sb/PS), [Kuopio] Nilsiä, Loutteinen. [WGS84 63.2197 °N 28.0218 °E] Coll. Risto Tuomikoski, 13.VII.1946, det. Tauno Ulvinen 2017 (H http://id.luomus.fi/HA.H4219995). Schistidium submuticum H.H.Blom FINLAND. Ostrobottnia ultima (Obu/PeP), Ylitornio, Tengeliö, Tengeliöjoki river, flooding canal of an old hydroelectric power plant. On concrete. UCS 73691:33563. Coll. Juhani Räsänen 16/1060s, 4.VII.2016, det. Tauno Ulvinen 2017 (OULU http://id.herb.oulu.fi/GAL.862). FINLAND. Regio kuusamoënsis (Ks/Ks), Salla, S end of gorge Kurtinniittykuru, close to border of Oulanka National Park, east slope. Separate calcareous bank on a slope, on partially sheltered vertical surface, SW exposition. Altogether 3 dm2 in an area of 0,5 m2, covers patches of less than 1 cm2 to 0,5 dm2. EUREF-FIN 7370450:596201. Coll. Riikka Juutinen 6937, 14.VI.2016, conf. Risto Virtanen 2016 (OULU http://tun.fi/GAK.1023). Previously assessed in Finland as regionally extinct (RE) (Syrjänen et al. 2010). Sphagnum divinum Flatberg & Hassel, punaterärahkasammal FINLAND. Regio aboënsis (Ab/V), Salo, Kisko, Toija, Viiari. Pine mire margin of small oligotrophic lowsedge fen, hummock. UCS 6690392: 3303805. Coll. Sanna Huttunen, 21.IV.2018 (TUR http://mus.utu.fi/TBR.120606); Satakunta (St/St), Huittinen, Puurijärvi-Isosuo National Park, north part of mire Isosuo, by the nature path. Spruce-pine mire, by the edge of pine mire. Here and there. UCS 6801950:3263295. Coll. Terhi Korvenpää, 29.IX.2018 (TUR http://mus.utu.fi/TBR.124263); Tavastia australis (Ta/EH), Tammela, Torronsuo National Park, by the hiking path coming from Kiljamo. Ridge-hollow pine bog, in a ditch. Patches. UCS 6740903:3314031. Coll. Terhi Korvenpää, 29.IX.2018 (TUR http://mus.utu.fi/ TBR.124265); Ostrobottnia kajanensis (Ok/Kn), Kuhmo, irrigated area south from Kilpelänvaara, south edge. Irrigated spruce mire, at the edge of forest and mire. Small patches. UCS 7104596:3662570. Coll. Terhi Korvenpää, Memoranda Soc. Fauna Flora Fennica 94, 2018 • Juutinen et al. 4.IX.2018 (TUR http://mus.utu.fi/TBR.124267); Ostrobottnia ultima (Obu/PeP), Rovaniemi, Pisavaara Strict Nature Reserve, Pitkäjänkä mire. Rubus chamaemorus spruce mire, beside a spring stream. EUREF-FIN 7356413:415414. Coll. Riikka Juutinen 7480, Inkeri Ahonen & Salli Uljas, 25.VIII.2018, det. Riikka Juutinen & Sanna Huttunen 2018 (OULU http://tun.fi/GAK.2492). New species for Finland. Distribution of the species is uncertain due to recent taxonomic changes. Hassel et al. (2018) showed that the name Sphagnum magellanicum belongs to a South American taxon and European specimens treated under that name are either S. medium or S. divinum. Old specimens in Finnish herbaria with name S. magellanicum have not been checked. Typical habitat is drier surface in a minerotrophic fen with Pinus sylvestris, Picea abies or Betula pubescens (Laine et al. 2018). Sphagnum medium Limpr., punarahkasammal FINLAND. Regio aboënsis (Ab/V), Salo, Teijo National Park, south part of mire Kylmässuo, 10 m W from spring pond. Close to a spring pond, on pine mire. Patches here and there. UCS 6686136:3281722. Coll. Terhi Korvenpää, 28.IX.2018 (TUR http://mus.utu.fi/TBR.124262); Satakunta (St/St), Huittinen, Puurijärvi-Isosuo National Park, north part of mire Isosuo, by the nature path, 40 E from bird watching tower. Edge of Sphagnum fuscum bog and lowsedge fen, on hummocks. Patches here and there. UCS 6801518:3263396. Coll. Terhi Korvenpää, 29.IX.2018 (TUR http://mus.utu.fi/TBR.124264); (Ta/EH), Tammela, Torronsuo National Park, by the hiking path coming from Kiljamo. Low-sedge fen, on drier hummocks. Here and there. UCS 6740920:3314225. Coll. Terhi Korvenpää, 29.IX. 2018 (TUR http://mus.utu.fi/TBR.124266); Ostrobottnia ultima (Obu/PeP), Rovaniemi, Pisavaara Strict Nature Reserve, Pitkäjänkä mire. Oligotrophic pine fen. EUREFFIN 7356503:415769. Coll. Riikka Juutinen 7478, Inkeri Ahonen & Salli Uljas, 25.VIII.2018, det. Riikka Juutinen & Sanna Huttunen 2018 (OULU http://tun.fi/GAK.2490); Lapponia sompiensis (Lks/SoL), Sodankylä, Peurasuvanto, Koitelainen, Mätöpeura-aapa mire, 975 m N from map name Niemelänkangas. Pine fen, on hummocks. Large pure patches. EUREF-FIN 7516445: 489479. Coll. Riikka Juutinen 7490, 3.IX.2018 (OULU http://tun.fi/GAK.2502). New species for Finland. Distribution of the species is uncertain due to recent taxonomic changes. Hassel et al. (2018) showed that the name Sphagnum magellanicum belongs to a South American taxon and European specimens treated under that name are either S. medium or S. divinum. Old specimens in Finnish herbaria with name S. magellanicum have not been checked. 93 Typical habitat is low hummock, lawn or carpet in an ombrotrophic mire (Laine et al. 2018). Sphagnum subfulvum Sjörs FINLAND. Karelia australis (Ka/EK), [Kouvola] Sippola, the NE-shore of Mölöjärvi lake [UCS 6752:3518]. Coll. Lars Fagerström, 11.VIII.1969 (KUO http://tun.fi/ SBR.23079). Tayloria lingulata (Dicks.) Lindb. FINLAND. Tavastia borealis (Tb/PH), Konnevesi, Lummukankylä, Karpinharju, Etelä-Konnevesi National Park, Haukkaloilo south from Haukkalahti bay. Large flat rock near the shore c. 20-30 cm above water surface. EUREFFIN 6936880:477026. Coll. Timo Kypärä 2302, 29.V.2017 (OULU http://tun.fi/GAT.384). Thuidium delicatulum (Hedw.) Schimp. FINLAND. Tavastia borealis (Tb/PH), Virrat, Suonijärvet, rock by Myllyoja river. Rock shelf at the foot of a rock wall, somewhat herb-rich spot. UCS 69069: 33304. Coll. Ari Parnela & Tuomo Kuitunen, 15.VIII.2018, det. Ari Parnela 2018 (OULU http://id.herb.oulu.fi/GAP.56). Timmia austriaca Hedw. FINLAND. Ostrobottnia australis (Oa/EP), Teuva, Luovankylä, Iso-Parra S, N end of Sikakallio rocks. Foot of a rock face, NW exposition. Relatively abundant, a patch of c. 3 dm2. EUREF-FIN 6930821:233110. Coll. Timo Kypärä 2421, 16.VIII.2017 (OULU http://tun.fi/GAT.504). Ulota bruchii Hornsch. ex Brid. FINLAND. Karelia borealis (Kb/PK), Lieksa, Hatunkylä, Ukonsärkkä Natura 2000 area, Pieni Hemmivaara, base of W slope. Old OMT forest, on large aspen. EUREF-FIN 7027203:687006. Coll. Timo Kypärä 2455, 7.IX.2017 (OULU http://tun.fi/GAT.538). New records – Marchantiophyta Aneura maxima (Schiffn.) Steph. FINLAND. Savonia borealis (Sb/PS), Lapinlahti, Varpaisjärvi, Korpisenniemi, Kuikkasuo mire. On spring seepage, on detritus. Small patches in an area of a few m2. EUREFFIN 7021726:532855. Coll. Timo Kypärä 2518, 27.IX. 2017 (OULU http://tun.fi.GAT.601). Aneura mirabilis (Malmb.) Wickett & Goffinet (Cryptothallus mirabilis) FINLAND. Savonia borealis (Sb/PS), Suonenjoki, Käpylä, Lintharju, rivulet running out from Onkilampi pond, E bank between road and power line. Spring water influ- 94 Juutinen et al. • Memoranda Soc. Fauna Flora Fennica 94, 2018 enced bank near waterline. EUREF-FIN 6946217:501701. Coll. Timo Kypärä 2305, 13.VI.2017 (OULU http://tun.fi/ GAT.387). Bazzania tricrenata (Wahlenb.) Lindb. FINLAND. Savonia australis (Sa/ES), Savonlinna, Pellossalo, rock hill between Vääräjärvi and Paskolammi. N cliff. UCS 68444:36125-6. Coll. Harri Arkkio, 23.VI.2002 (OULU http://id.herb.oulu.fi/0016962). Fuscocephaloziopsis albescens (Hook.) Váňa & L.Söderstr. (Pleurocladula albescens) FINLAND. Karelia borealis (Kb/PK), Lieksa, Ruunaa, road turning point SW from Tökrösensuo mire, S side. In a forest road ditch, on bare broken soil. UCS 7037489:366651. Coll. Juhani Räsänen 16/2029s-5, 9.X.2016, det. Xiaolan He & Sanna Laaka-Lindberg 2016 (H http:// id.luomus.fi/HA.H4259588, Laaka-Lindberg et al. 2016). Fuscocephaloziopsis lunulifolia Blepharostoma trichophyllum (L.) Dumort. subsp. brevirete (Bryhn & Kaal.) R.M.Schust. FINLAND, Regio Kuusamoënsis (Ks/Ks), Salla, Tuntsa, SE side of Auermavaara-Vittoiva-Kuusivaara forested hill, 1,5 km NW from Pirunkirkko. On vertical furrow of serpentinite boulder. EUREF-FIN 7492091:609437. Coll. Riikka Juutinen 7193, 31.VIII.2016 (OULU http:// tun.fi/GAK.1889, as associated species in Weissia controversa specimen). Cephalozia ambigua C.Massal. FINLAND. Ostrobottnia media (Om/KP), Pyhäjärvi, Pyhäsalmi, Pyhäsalmi mine. Brook at the foots of the mine tailings. EUREF-FIN 7059521:451895. Coll. Anna-Liisa Ruotsalainen, Piippa Wäli & Kaisa Lehosmaa, 14.IX. 2017, det. Riikka Juutinen 2017, Timo Kypärä 2017 (blind determination) (OULU http://id.herb.oulu.fi/GAL.2457). (Dumort.) Váňa & L.Söderstr. (Cephalozia lunulifolia) FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Pyhäjärvi, Petäjäsaari island. Herb rich forest. Fairly abundant. UCS 6855393:3652983. Coll. Tuomo Takala 1209, 20.IX. 2012, det. Tuomo Takala 2012, conf. Riikka Juutinen 2013 (not a blind determination) (KUO http://tun.fi/SBR.58148). Fuscocephaloziopsis pleniceps (Austin) Váňa & L.Söderstr. (Cephalozia pleniceps) FINLAND. Karelia ladogensis (KL/LK), Parikkala, Tiviä, Lähdelampi pond. Bryophyte cover at the margin of a spring. Occasional. UCS 682604:363738. Coll. Tuomo Kuitunen 3380, 17.VII.2017, det. Tuomo Kuitunen 2017, Riikka Juutinen 2018 (blind determination) (OULU http:// id.herb.oulu.fi/GAL.2345). Geocalyx graveolens (Schrad.) Nees Cephaloziella integerrima (Lindb.) Warnst. FINLAND. Lapponia enontekiensis (Le/EnL), Enontekiö, Ounastunturi, Pahakuru, W part. Moist, shaded rock face of a rocky ravine. Relatively abundant, a patch of almost one dm2. EUREF-FIN 7571267:373419. Coll. Timo Kypärä 2637, 27.VIII.2018, conf. Kimmo Syrjänen 2018 (not a blind determination) (OULU http://tun.fi./GAT.722). FINLAND. Lapponia inarensis (Li/InL), Inari, Urho Kekkonen National Park, Luujänkä, headwater stream of Luuoja running N from Luulampi pond, under a bridge of a hiking path. Bank of a river, on peat. Among other bryophytes. UCS 7587111:3522518. Coll. Riikka Juutinen 6401, 25.IX.2013 (OULU http://id.herb.oulu.fi/0038552). Hygrobiella laxifolia (Hook.) Spruce Clevea hyalina (Sommerf.) Lindb. (Athalamia hyalina) FINLAND. Karelia borealis (Kb/PK), Tohmajärvi, Luosovaara, N cliff of Jalajanvaara hill. A cliff shaded by a mature forest cover, horizontal crack in a cliff face at the upper part of the slope, NE exposition. C. 10 cm × 30 cm patch. EUREF-FIN 6902047:672325. Coll. Timo Kypärä 2413, 3.VIII.2017, conf. Kimmo Syrjänen 2017 (OULU http://tun.fi/GAT.496). Conocephalum conicum (L.) Dumort. FINLAND. Lapponia enontekiensis (Le/EnL), Enontekiö, Kilpisjärvi, Malla Strict Nature Reserve, fjell Pikku-Malla (Šilismalla danjege, Gilbbesmalla), base of E slope. Bank of a stream in fjell birch forest. Patches along a few meters. UCS 7676930:3251405. Coll. Kimmo Syrjänen, 23. VII.2016 (TUR http://mus.utu.fi/TBR.122124). FINLAND. Tavastia australis (Ta/EH), Jämsä, Hassi, Kähönjoki, Hassinkoski rapids. On wet, slightly slurry rock in a stream. UCS 68491:34034. Coll. Ari Parnela, 27.VII. 2018, conf. Riikka Juutinen 2018 (OULU http://id.herb.oulu. fi/GAP.57). Jungermannia pumila With. FINLAND. Nylandia (N/U), Kirkkonummi, Laitamaa, rivulet running downward from Kakarlampi pond. Pristine rivulet, on top of a rapid rock. 4 cm × 4 cm patch. UCS 66832693361069. Coll. Esa Ervasti 1179 & Esko Vuorinen, 10.IX.2018 (TUR http://mus.utu.fi/TBR.124256); Karelia ladogensis (Kl/LK), Rautjärvi, Koskenkulma, Lohioja. On top of rapid stones. Abundant. UCS 681774:362155. Coll. Tuomo Kuitunen 3549, 19.VII.2018, conf. Ari Parnela 2018 (OULU http://id.herb.oulu.fi/GAP.55). Memoranda Soc. Fauna Flora Fennica 94, 2018 • Juutinen et al. Kurzia trichoclados (K. Müller) Grolle, lännenviiksisammal, västlig fingerfliksmossa FINLAND. Savonia borealis (Sb/PS), Kuopio, Nilsiä, Lastukoski, N part of Pisa hill, Pirunkellari E. Cavity at foot of a rock face. One dense patch seen, c. 30 cm2. EUREF-FIN 7011644:566487. Coll. Timo Kypärä 2380, 24.VII.2017, det. Kimmo Syrjänen 2017 (OULU http://tun.fi/GAT.463). New species for Finland. This species has suboceanic-montane distribution. It is common along the southwest coast of Norway, also occurs in Sweden and in many parts of Central Europe (Damsholt 2009). It is assessed as threatened in Sweden, Italy, Spain, Czech Republic, Slovakia and Switzerland (Hodgetts 2015). In the Nordic countries it occurs from the sea level to 500 m but in Central Europe it is montane and grows above 780-2550 m (Damsholt 2009). Marchantia polymorpha L. subsp. polymorpha FINLAND. Lapponia kittilensis (Lkk/KiL), Kolari, Kalkkikangas, Ruonajänkkä mire. Eutrophic fen, in small spring brook. Scarce. UCS 7490: 3356. Coll. Veli Haikonen 30583, 25.VIII.2016 (H http://id.luomus.fi/HA.H4259424). Marchantia polymorpha L. subsp. ruderalis Bischl. & Boissel.-Dub. FINLAND. Tavastia australis (Ta/EH), Heinola, Sonnanen. New burn-clearing area of Vaccinium vitis-idaea type forest, bottom of sandy pit. Homogenous patch. UCS 6797:3469. Coll. Veli Haikonen 30603, 3.IX.2016 (H http://id.luomus.fi/HA.H4259427); Karelia ladogensis (Kl/ LK), Rautjärvi, Simpele, Peipvuori, side of the village road. Side of a road, soil landfill, on moist sand. Relatively abundant. UCS 681554:362445. Coll. Tuomo Kuitunen 3392 (OULU http://id.herb.oulu.fi/GAL.2327); Lapponia kittilensis (Lkk/KiL), Muonio, Rauhalanmaa, Päämaa. Empetrum – Myrtillus type forest, on campfire base. Scarce. UCS 7547:3363. Coll. Veli Haikonen 30580, 27.VIII.2016 (H http://id.luomus.fi/HA.H4259425). Marsupella sparsifolia (Lindb.) Dumort. FINLAND. Lapponia kittilensis (Lkk/KiL), Muonio, Pallastunturi fjells, Vatikuru gorge. On stone in a rivulet. Mixed with other species in an area of 3 dm2. UCS 7555318:3377443. Coll. Riikka Juutinen, 4.VI.2013 (OULU http://id.herb.oulu.fi/0038877). Marsupella sphacelata (Gieseke ex Lindenb.) Dumort. FINLAND. Karelia borealis (Kb/PK), Juuka, Ruokolanvaara, between Toivala and Ruohonkorpi, Toivala NE. On rock in stream. Relatively abundant, on two rocks c. 4 dm2. EUREF-FIN 7003755:608187. Coll. Timo Kypärä 2459, 12.IX.2017, conf. Kimmo Syrjänen 2017 (OULU http:// tun.fi/GAT.542). 95 Mesoptychia rutheana (Limpr.) L.Söderstr. & Váňa (Leiocolea rutheana) FINLAND. Ostrobottnia australis (Oa/EP), Kauhajoki, Ikkeläjärvi, Rytiperä, Harjanneva W. Herb rich sedge fen, spring seepage. EUREF-FIN 6921891:270616. Coll. Timo Kypärä 2353, 19.VI.2017 (OULU http://tun.fi/GAT.436). Moerckia flotoviana (Nees) Schiffn. FINLAND. Tavastia australis (Ta/EH), Hattula, Alajoki, 250 m W from Sahakoski. Flooding Scorpidium flark fen. UCS 67682:33502. Coll. Teppo Häyhä, 3.XI.2000, det. Riikka Juutinen 2015 (OULU http://id.herb.oulu.fi/ GAL.1851); Karelia borealis (Kb/PK), Juuka, Tahkovaara hill. In rich fen. UCS 7010:3605. Coll. Stephan Gradstein 2080 & J. de Molenaar, 5.VIII.1972, det. Riikka Juutinen 2016 (KUO http://tun.fi/SBR.27740); Ostrobottnia ouluensis (Obo/OP), [Oulu] Kiiminki, Iso Juuanjärvi, SW shore of Juuanjoki river rapids upstream from line path. On submerged rock at rapids shore. UCS 722347:345288. Coll. Martti Ohenoja, 20.9.1964, det. Riikka Juutinen 2017 (OULU http://id.herb.oulu.fi/GAL.1855); Regio kuusamoënsis (Ks/Ks), Kuusamo, Kitkajoki, Käyläkoski rapids. Riverbank. [UCS 7358:3596] Coll. Viktor Ferdinand Brotherus, 30.VII.1883, det. A. Potemkin 2008 (H http://id.luomus. fi/HA.H4258903): Lapponia kittilensis (Lkk/KiL), Kolari, Äkäsjokisuu, Kalkkikangas, Aittamaa, S from Ruonajänkkä, W from Mannajärvi, W side of quarry. Old track, on periodically wet soil. In a strech of a few meters. UCS 7490368:3355889. Coll. Kimmo Syrjänen, 21.VII.2016 (TUR http://mus.utu.fi/TBR.122195); Lapponia sompiensis (Lks/SoL), Pelkosenniemi, Kätkävaara N, on snow mobile route near forest edge. Base of a hummock, rich fen. Altogether 2 cm2 in a stretch of about 50 meters. EUREF-FIN 7450867:539388. Coll. Mari Kekäläinen 5/2016, 17.VIII.2016, det. Mari Kekäläinen 2017 (supervised by Riikka Juutinen) (OULU http://tun.fi/GAK.1825); Lapponia enontekiensis (Le/EnL), Enontekiö, Malla Strict Nature Reserve, Pikku-Malla E. Foot of a wet calcareous cliff, on soil of dolomite chippings, partly covered by boulders, shaded. Two dm2 among other bryophytes. UCS 7676842:3251205. Coll. Riikka Juutinen 3816, 30. VI.2011, det. Riikka Juutinen 2015 (OULU http://id.herb. oulu.fi/GAL.1858); Lapponia inarensis (Li/InL), Inari, Lemmenjoki National Park, Pikku-Kietsemäjärvi lake, W shore. Lake shore, on open nutrient rich mineral soil. A few patches, altogether a few dm2 in an area of about two m2. UCS 7613384:3422371. Coll. Riikka Juutinen 4376, 19.VIII.2010, det. Riikka Juutinen 2015 (OULU http:// id.herb.oulu.fi/GAL.1859). See also Juutinen (2017). Moerckia hibernica (Hook.) Goettsche sensu stricta FINLAND. Tavastia borealis (Tb/PH), Toivakka, Paloinen, Sääkslahti, Vookelinselkä. Lake shore with rich fen features. Probably scarce. UCS 6883959:3454808. Coll. Johanna Hallman & Timo Kypärä 365, 29.V.2008, det. Riikka Juutinen 2015 (OULU http://id.herb.oulu.fi/GAL.1860); 96 Juutinen et al. • Memoranda Soc. Fauna Flora Fennica 94, 2018 Ostrobottnia media (Om/KP), Raahe, Saloinen, N from Rytilampi pond, E part of small rich fen patch. On bare peat. In an area of 20 cm × 5 cm. UCS 7171803:3377290. Coll. Tauno Ulvinen, 10.X.2003, det. Riikka Juutinen 2017 (OULU http://id.herb.oulu.fi/GAL.1864); Ostrobottnia kajanensis (Ok/Kn), Puolanka, Lylykylä, about one kilometer from Naulala farm. Carex flava – Trichophorum – Campylium stellatum vegetation type, undertype Calliergon trifarium. UCS 7208:3545. Coll. Paavo Havas Kn 27, 19.VIII.1956, det. Riikka Juutinen 2015 (OULU http:// id.herb.oulu.fi/GAL.1861); Ostrobottnia ouluensis (Obo/OP), Oulu, Kiiminki, Iso Juuanjärvi, SW shore of Juuanjoki river, rapids upstream from line path. Shaded, wet rock near river shore. UCS 722347:345288. Coll. Martti Ohenoja CXVIII a), 20.IX.1964, det. Riikka Juutinen 2017 (OULU http://id.herb.oulu.fi/GAL.1862, could not find conducting strands despite exceptional effort); Regio kuusamoënsis (Ks/Ks), Kuusamo, Oulanka National Park, Tulipuro stream. On stream bank at steep part of the stream rapids. One patch of two sq. dm. UCS 73657:36096. Coll. Risto Virtanen, 29.VIII.1997, det. Riikka Juutinen 2015 (OULU http://id.herb.oulu.fi/GAL.1866); Lapponia sompiensis (Lks/ SoL), Savukoski, Kemihaara wilderness area, N from Jaaraoja stream. Rich fen by a stream. UCS 7545235: 3564689. Coll. Pia Kangas, 11.IX.2012, det. Riikka Juutinen 2016 (OULU http://id.herb.oulu.fi/0036700). See also Juutinen (2017). ate area. Presumably scarce, only a few thalli were found among other bryophytes. EUREF-FIN 6999620:642326. Coll. Timo Kypärä 2617, 17.VIII.2018 (OULU http://tun. fi/GAT.702). Pellia epiphylla (L.) Corda (subsp. epiphylla) FINLAND. Karelia ladogensis (Kl/LK), Rautjärvi, Koskenkulma, Lohioja. On rock in rapids, on depression. Scarce. UCS 681774:362155. Coll. Tuomo Kuitunen 3558, 19.VII.2018, conf. Timo Kypärä 2018 (OULU http://id.herb.oulu.fi/GAP.66). Pellia neesiana (Gottsche) Limpr. FINLAND. Karelia ladogensis (Kl/LK), Rautjärvi, Simpele, Peipvuori, E side. Rock outcrop, under a wet cliff at side of a puddle. Abundant. UCS 68157:36244. Coll. Tuomo Kuitunen 3388, 19.VII.2017, det. Timo Kypärä 2018 (OULU http://id.herb.oulu.fi/GAL.2343). Riccardia palmata (Hedw.) Carruth. FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Särkilahti, Oronmylly, Heralamminhete mire. At the edge of a rivulet flowing through Heralamminhete. Scarce. UCS 36255:68373. Coll. Tuomo Kuitunen 3550, 20.VII.2018, conf. Ari Parnela 2018, Timo Kypärä 2018 (OULU http:// id.herb.oulu.fi/GAP.65). Riccia beyrichiana Hampe Mylia taylorii (Hook.) Gray FINLAND. Karelia borealis (Kb/PK), Lieksa, Koli, Koli National Park, rocky ridge E from Kolinuuro in Pieni-Koli. Moist rock face, NE exposition. Pure c. 30 cm × 30 cm patch. EUREF-FIN 6998408:642482. Coll. Timo Kypärä 2616, 17.8.2018 (OULU http://tun.fi./GAT.701). Nardia geoscyphus (De Not.) Lindb. FINLAND. Karelia ladogensis (Kl/LK), Parikkala, Särkisalmi, Oronmylly. Sandy side of a forest road. Abundant. UCS 683750:362627. Coll. Tuomo Kuitunen 3383, 18.VII.2017, det Tuomo Kuitunen 2017 & Riikka Juutinen 2018 (blind determination) (OULU http://id.herb.oulu. fi/GAL.2344). Odontoschisma macounii (Austin) Underw. FINLAND. Lapponia kittilensis (LKK/KiL), Kittilä, Venejoki, about 10 km SE of Kittilä Village. Venejoki serpentinite rock. UCS 74987(14):33980(38). Coll. Jari Teeriaho 103.4/04, 5.VIII.2004, conf. Sanna Laaka-Lindberg 2014 (H http://id.luomus.fi/HA.H4257909). Pellia endiviifolia (Dicks.) Dumort. FINLAND. Karelia borealis (Kb/PK), Lieksa, Koli, Koli National Park, parking lot for downhill skiing slopes, upper edge of parking lot. Sand and mineral soil, open ruder- FINLAND. Savonia borealis (Sb/PS), Joroinen. In a field ditch, on sandy soil. WGS84 62°15´N 27°20´E. Coll. G. Grotenfelt, 17.VIII.1876 (H http://id.luomus.fi/HA.H4205417). This specimen was earlier thought to be misidentified due to mix up between two herbarium specimens. Grotenfelt has collected also Riccia ciliata (H http://id.luomus.fi/ HA.H4205330) from Joroinen in 1877. The specimen http:// id.luomus.fi/HA.H4205417 is present in H and contains no additional determination notes (Sanna Laaka-Lindberg, pers. comm. 22.10.2018). Scapania apiculata Spruce FINLAND. Ostrobottnia ultima (Obu/PeP), Rovaniemi, Kemijoki river, between Oikarainen and Tennilä villages, N shore of the river, a ravine situated between houses Rotkola and Rinne, 280 m from the main road downstream. Shaded, narrow ravine with a rivulet, on 10 cm diameter fallen, hard and barkless Betula. As a mat in 1 m length of the trunk, altogether 4–5 dm2. EUREF-FIN 7365654: 466855. Coll. Riikka Juutinen 7519, 24.IX.2018, conf. Timo Kypärä 2018 (OULU http://tun.fi/GAK.2531). Scapania mucronata H.Buch FINLAND. Karelia australis (Ka/EK), [Hamina] Vehkalahti, Turkia, Salakkaoja, Rysäkoski, the lower rapids. [UCS 67362:35139] Coll. Lars Fagerström, 23.VI.1965, det. T. Laine 1966. (H http://id.luomus.fi/HA.H4237447). Memoranda Soc. Fauna Flora Fennica 94, 2018 • Juutinen et al. Scapania nemorea (L.) Grolle FINLAND. Karelia borealis (Kb/PK), Kaavi, Luikonlahti, Kekäleinen bay. UCS 6980:3583. Coll. Heikki Roivainen, 26.VIII.1922 (KUO http://tun.fi/SBR.13422) Scapania spitsbergensis (Lindb.) Müll.Frib. FINLAND. Karelia borealis (Kb/PK), Joensuu, Pieni-Valkea. At bottom of a rocky kettle hole, cold cavities between rocks. UCS 6954239:3672010. Coll. Jari Teeriaho 49.1/2015, 29.VI.2015, det. Kimmo Syrjänen 2017 (OULU http://id.herb.oulu.fi/GAP.40); Lapponia sompiensis (Lks/SoL), Savukoski, Pyhäkuru. Boulder field at the bottom of a gorge, partly still frozen cavities between boulders. UCS 74676:35654. Coll. Jari Teeriaho 49.3/2014, 2. VII.2014, det. Kimmo Syrjänen 2017 (OULU http://id.herb. oulu.fi/GAP.41). Also, another specimen from Savukoski (OULU http://id.herb.oulu.fi/GAL.1357). Scapania uliginosa (Lindenb.) Dumort. FINLAND. Karelia australis (Ka/EK), [Hamina] Vehkalahti, between Reitkalli and Ylämaa. In a puddle at the edge of a spruce fen meadow. [WGS84 60°36´N 27°01´E] Coll. V. Kujala, 11.VII.1922 (H http://id.luomus.fi/HA.H4217366). Tritomaria exsectiformis (Breidl.) Schiffn. ex Loeske FINLAND. Karelia borealis (Kb/PK), Nurmes, Mujejärvi, along a brook running to N end of Ylä-Sammaljärvi lake. Spruce fen, on an old dead coniferous tree in the brook. Scarce, single shoots among other bryophytes in an area of two dm2. EUREF-FIN 7077861:623867. Coll. Timo Kypärä 2579, 23.7.2018 (OULU http://tun.fi./GAT.664). Corrections – Bryophyta Gymnostomum calcareum Nees & Hornsch. FINLAND. Regio aboënsis (Ab/V), Paimio, Varkaankellarinmäki. EUREF-FIN 6701594:268574. Coll. Kimmo Syrjänen, Sanna Laaka-Lindberg, Terhi Korvenpää & Turkka Korvenpää, 15.V.2007 (see Syrjänen 2009). The specimen could not be checked as it is not present in TUR. Occurrence is uncertain, probably erroneous; Ostrobottnia kajanensis (Ok/Kn). Misidentification, previous voucher (TUR http://mus.utu.fi/TBR.87824) is Gymnostomum aeruginosum (det. Heribert Köckinger 2017). Occurrence in Ok is deleted. Only correctly identified specimens from Finland are from Karelia borealis (Kb/PK) (conf. Heribert Köckinger 2017). All specimens in OULU and TUR were checked by Heribert Köckinger on 2017. 97 Hymenostylium gracillimum (Nees & Hornsch.) Köckinger & J.Kučera (Gymnostomum boreale) FINLAND. Lapponia enontekiensis (Le/EnL). Specimens in OULU and TUR were checked by Heribert Köckinger on 2017. All specimens from Kilpisjärvi area in Enontekiö are Gymnostomum aeruginosum. Does not occur in Finland. Only confirmed herbarium specimens from North Europe are from Kulmakkapuro in Paanajärvi (Ks/Rs, Russia) (Köckinger, pers.comm. 2017). There are two unchecked specimens of Gymnostomum boreale in H (http:// id.luomus.fi/HA.H4071783, http://id.luomus.fi/HA.H4071784) collected from Salla, Kuolajärvi, Kutsa Nature Reserve (Ks/Rs, Russia) in 1934 by Antero Vaarama (det. Tauno Ulvinen, conf. Lars Hedenäs; Ulvinen 1996). Oncophorus virens (Hedw.) Brid. FINLAND. No deletions despite majority of herbarium specimens earlier named as O. virens are in fact O. integerrimus (det. Tauno Ulvinen 2018). Oncophorus wahlenbergii Brid. FINLAND. Karelia australis (Ka/EK), [Hamina] Vehkalahti, Onkamaa, Riissuonsilta. [UCS 672:351] Coll. Lars Fagerström, 4.VII.1969 (H, specimen not digitalized); Karelia ladogensis (Kl/LK), [Parikkala] Uukuniemi. [UCS 685:3365] Coll. [Theodor] Simming, 22.VII.1861, det. Tauno Ulvinen 2018 (H http://id.luomus.fi/HA.H4097956, specimen not digitalized); Ostrobottnia australis (Oa/EP), [Vaasa] Vasa, Boskär. [UCS 701:322] Coll. V.F. Brotherus, 15.VII.1880 (H, specimen not digitalized); Ostrobottnia media (Om/KP), [Kokkola] Kälviä, Helluskari [Helluskeri]. [UCS 7099:7099] Coll. A.V. Auer, 10.8.1933 (H, specimen not digitalized). Misidentifications. Above mentioned previous vouchers of Oncophorus wahlenbergii are all Oncophorus elongatus (det. Tauno Ulvinen 2018). There are no other correctly identified herbarium specimens in H, JYV, KUO, OULU or TUR. Occurrences in Ka, Kl, Oa and Om are deleted. Rhynchostegium riparioides (Hedw.) Card. (Platyhypnidium riparioides) FINLAND. Karelia australis (Ka/EK), Virolahti, Vilkkilä, bay NW from Kirskallionniemi cape (Ruosviikinlahti bay). On loam bottom, c. 50,5 m depth, brackish water. UCS 67146:35395. Coll. Tapio Rintanen, 17.IX.1980 (TUR http://mus.utu.fi/TBR.107691). This specimen is Oxyrrhynchium speciosum (det. Sanna Huttunen 2017). This was the only specimen from Ka, so occurrence is deleted; Savonia borealis (Sb/PS), Kuopio, Kolmisopenlampi pond. [UCS 6974:3531] Coll. Reino Fagerstén 9181, 6.8.1982 (KUO http://tun.fi/SBR.57047). This specimen is Oxyrrhynchium speciosum (det. Sanna Huttunen 2018). There are no other correctly identified herbarium specimens in H, JYV, KUO, OULU or TUR. Occurrence in Sb is deleted; Karelia borealis (Kb/PK), Liperi, Paskoläh- 98 Juutinen et al. • Memoranda Soc. Fauna Flora Fennica 94, 2018 de. [UCS 69544:36391] Coll. Reino Fagerstén 9184, 28.VIII.1983 (KUO http://tun.fi/SBR.57049). This specimen is Brachythecium rivulare (det. Sanna Huttunen 2018). There are no other correctly identified herbarium specimens in H, JYV, KUO, OULU or TUR. Occurrence in Kb is deleted; Ostrobottnia media (Om/KP), Lappajärvi, Lappajärvi lake, N-part of lake, Piihamina. Coll. Anna-Maria Koivisto & Anssi Teppo, 3.VIII.2009 (OULU 10000854, specimen not digitalized). This specimen is Oxyrrhynchium speciosum (det. Sanna Huttunen 2017). This was the only specimen from Om, so occurrence is deleted. Sphagnum inundatum Russow FINLAND. Ostrobottnia media (Om/KP). No specimens found from herbaria H, OULU or TUR. Occurrence in Om is deleted. Sphagnum rubiginosum Flatberg FINLAND. Ostrobottnia australis (Oa/EP). Misidentification. Previous voucher (OULU http://id.herb.oulu.fi/0035792; Juutinen et al. 2013) is Sphagnum girgensohnii (det. Kjell Ivar Flatberg) (Flatberg, pers.comm. 23.2.2018). The specimen in question is collected from Kauhajoki by Raimo Heikkilä & Tapio Lindholm during 1994. It is not digitalized (despite having HTTP URI identifier) and has not been returned to OULU. Occurrence in Oa is deleted. To present knowledge, the species does not occur in Finland. from Sammuttijänkä – Vaijoenjänkä mire protection area in Inari is Shljakovianthus quadrilobus (Lindb.) Konstant. & Vilnet var. collenchymatica (Barbilophozia quadriloba (Lindb.) Loeske var. collenchymatica (R.M. Schust.) Stotler & Crand.-Stotl.). Occurrence in Li is deleted. Nadya Konstantinova has checked all Finnish material in OULU and TUR. The only confirmed identifications from Finland are by Martti Ohenoja on 7.VII.1962 (TUR http://mus.utu.fi/TBR.67260, OULU http://id.herb.oulu. fi/0001741, conf. Nadya Konstantinova 2017) and by Heikki Roivainen on 13.VIII.1935 (H http://id.luomus.fi/ HA.H4205570, conf. Nadya Konstantinova 2010) from Saana in Enontekiö commune. All other specimens in OULU were misidentified (nine specimens from the 21st century). Of the two specimens in H, the one collected from Ridnitsohkka (http://id.luomus.fi/HA.H4225301) has not been checked by Konstantinova (Sanna Laaka-Lindberg, pers. comm. 20.3.2018). Possible specimens in JYV or KUO were not checked. Scapania uliginosa (Lindenb.) Dumort. FINLAND. Nylandia (N/U). Previous voucher by V. Kujala on 11.VII.1922 from [Hamina] Vehkalahti (Ka/EK) (H http://id.luomus.fi/HA.H4217366) has been misplaced in province N since 2011 (Ulvinen & Syrjänen 2011b). Occurrence in N is deleted. References Corrections – Marchantiophyta Harpanthus scutatus (F.Weber & D.Mohr) Spruce FINLAND. Ostrobottnia ultima (Obu/PeP), Rovaniemi, Pisavaara Strict Nature Reserve, Isolomanoja. Spruce mire along Isolomanoja brook in eastern part of the Nature Reserve. UCS 7357:3417. Coll. Sanna Laaka-Lindberg, 29.VIII.1989 (H http://id.luomus.fi/HA.H4254775). Misidentification. Specimen is Lophozia sp. (det. Sanna LaakaLindberg & Xiaolan He 2018). This was the only specimen from Obu, so occurrence is deleted. Orthocaulis cavifolius H.Buch & S.W.Arnell (Anastrophyllum cavifolium) FINLAND. Lapponia sompiensis (Lks/SoL), Sodankylä, Rajala, Saittajoki river, Pahtakoski rapids. [UCS 75036: 34661] Coll. Mauno J. Kotilainen, 15.VIII.1951, det. Martti Ohenoja 2001 (H or OULU, specimen not digitized). The specimen could not be checked as it is no longer present in either H or OULU. Occurrence is uncertain, probably erroneous; Lapponia inarensis (Li/InL). Both specimens (OULU http://id.herb.oulu.fi/0001740, http://id.herb. oulu.fi/0028843) were erroneous (det. Nadya Konstantinova 2018). The first one by Risto Virtanen on 29.VI.2005 from Rišnjárvarri in Utsjoki commune is Sphenolobus minutus (Schreb.) Berggr. var. grandis (Gottsche ex Lindb.) Jørg. and the second by Riikka Juutinen on 9.VIII.2011 Alava, R., Laine, U. & Huhtinen, S. 2004: Edvard August Vainio’s travels to Finnish and Russian Karelia and to Finnish Lapland. – Publications from the Herbarium University of Turku 9: 1–62. Turku. Univ. of Turku. Damsholt, K. 2002: Illustrated Flora of Nordic Liverworts and Hornworts. – Nordic Bryological Society, Lund. p. 443–445. Hassel, K., Kyrkjeeide, M.O., Yousefi, N., Prestø, T., Stenøien, H.K., Shaw, J.O. & Flatberg, K.I. 2018: Sphagnum divinum (sp. nov.) and S. medium Limpr. and their relationship to S. magellanicum Brid. – Journal of Bryology 2018: 1–26. https://doi.org/10.1080/037 36687.2018.1474424 Hedenäs, L. 2014: Plagiothecium denticulatum skogssidenmossa p. 306–307. – In: Nationalnyckeln till Sveriges flora och fauna. Bladmossor: Skirmossor-baronmossor. Bryophyta: Hookeria-Anomodon. ArtDatabanken, SLU, Uppsala. 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Laaka-Lindberg, S., Räsänen, J. & Xiaolan, H. 2016: Metsäteiden maksasammalia Pohjois-Karjalassa ja huomioita aiemmin heimoon Jungermanniaceae kuuluneiden lajien taksonomiasta. – Bryobrotherella 19: 62–70. Laine, J., Flatberg, K.I., Harju, P., Timonen, T., Minkkinen, K., Laine, A., Tuittila, E.-S. & Vasander, H. 2018: Sphagnum mosses – The Stars of European Mires. – University of Helsinki, Department of Forest Sciences, Sphagna Ky, Helsinki. p. 40–48. Rassi, P., Hyvärinen, E., Juslén, A. & Mannerkoski, I. (eds.) 2010: The Red List of Finnish Species. – Ym- 99 päristöministeriö & Suomen ympäristökeskus, Helsinki. P. 224. Ryömä, R., Oldén, A., Xiaolan, H. & Laaka-Lindberg, S. 2013: Suomen puutteellisesti tunnetut maksasammalsuvut: siiransammalet Nardia, rahtusammalet Cephaloziella ja pihtisammalet Cephalozia. – Suomen ympäristö 7/2013. Suomen ympäristökeskus. Helsinki. 61 s. 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L., Brown, E., Cargill, D.C., Costa, D.P., Crandall-Stotler, B.J., Cooper, E.D., Dauphin, G., Engel, J.J., Feldberg, K., Glenny, D., Gradstein, S.R., He, X., Heinrichs, J., Hentschel, J., Ilkiu-Borges, A.L., Katagiri, T., Konstantinova, N.A., Larraín, J., Long, D.G., Nebel, M., Pócs, T., Felisa Puche, F., Reiner-Drehwald, E., Renner, M.A.M., Sass-Gyarmati, A., Schäfer-Verwimp, A., Moragues, J.G.S., Stotler, R.E., Sukkharak, P., Thiers, B.M., Uribe, J., Váňa, J., Villarreal, J.C., Wigginton, M., Zhang, L., Zhu, R-L. 2016: World checklist of hornworts and liverworts. – PhytoKeys 59: 1–821. https://doi.org/10.3897/phytokeys.59.6261 Thiers, B. 2018: Index Herbariorum: A global directory of public herbaria and associated staff. New York Botanical Garden’s Virtual Herbarium. http://sweetgum.nybg. org/science/ih/ Ulvinen, T. 1996: Bryophytes of the former Kutsa Nature Reserve. – Oulanka Reports 16: 53–62. 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