FRULLANIA – synonyms (including doubtful taxa, orth var, nom herb) alphabetically with literature

Gerhard  Winter

Frullania-synonyms (including doubtful taxa, orth var, nom herb) alphabetically with literature FRULLANIA newsletter No6 2022 No 6. http://www.frullania.de/frullnl06.pdf 16.01.2022 Gerhard Winter Extract from my database This is a working document in progress abducens Steph. 1910 = Frullania diversitexta Steph. 1897

FRULLANIA newsletter No 6. No6 2022 http://www.frullania.de/frullnl06.pdf 16.01.2022 Frullania - synonyms (including doubtful taxa, orth var, nom herb) alphabetically with literature Gerhard Winter Extract from my database This is a working document in progress abducens Steph. 1910 Typus: Japan. Chugoku, Yasui -.8.1898, Inoué 24 in hb G G00066970 - n° SIB 125540/1 [typus] (lecto-, designated here, G[G00066970]!) syn. fide Hattori (1952) Basionym: Distribution: spec nov - syn nov - syn nov Stephani Iishiba Kamimura Hattori Kamimura Bonner Choe Stephani 1910 1912 1952 1952 1961 1965 1980 1985 Yuzawa Yamada & Iwatsuki Atwood, Vilnet, Mamontov 2000 2006 2021 = Frullania diversitexta 686 888 873 529 199 655 1171 928 Spec Hepat 4: 396-397 Bot Mag Tokyo 26: 35 [Japan, with key in Japanese] Contrib Fl Hepat Shikokuensem, Kochi: 123 [syn sub Fr. diversitexta] J Hatt Bot Lab 8: 29 [syn nov sub Fr diversitexta] J Hatt Bot Lab 24: 80 [sub F diversitexta] Index Hepat 5: 223 Illust Fl & Fauna Korea 24: 733 [South Korea] Stephani Icon Hepat Inedit [Galeiloba] C-227, microfiche 53 fig 3645 ["Japan, Yasui, Inoué No 24, 1898"] 1659 Natural Environmental Sci Res 13: 10 [Japan, syn sub Fr diversitexta] 1470 J Hatt Bot Lab 99: 28 [Catalog of Japan] 5566 Cryptogamie, Bryologie 42(3): 19-31 [lectotype] Steph. 1897 Page 1 abyssinica Steph. 1910 Typus: Ethiopia: Abyssinica sd, Schimper sn in hb G G00045269 - n° SIB 148708/1 - 17284 [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Cufodontis Bonner Vanden Berghen Stephani 1910 1952 1965 1976 1985 Wigginton 2004 = Frullania trinervis accumbens 686 2562 655 185 928 Spec Hepat 4: 366-367 Phyton 4 (1-3): 76 [Ethiopia] Index Hepat 5: 223 Bull Jard Bot Nat Belg 46 (1/2): 199 Stephani Icon Hepat Inedit [Galeiloba] C- 1, microfiche 49 fig 3347 [from type; "leg Schimper, Abyssinica"] - 3348 1172 Sript Bot Belg 30: 227, fig 149(J) from type [syn sub Fr trinervis, West Africa] (Lehm.) Drège 1843 Steph. 1911 Typus: Philippines. Luzon, Prov Tayabas, Infanta -.8.1909, C.B. Robinson, 9424 Bureau of Sci 9424 in hb G G00066971 - n° SIB 125570/1 Basionym: Distribution: Philippines spec nov - syn nov Stephani Verdoorn Verdoorn Verdoorn Bonner Stephani = Frullania meyeniana acolotis 1911 1929 1929 1930 1965 1985 686 245 548 293 655 928 Spec Hepat 4: 550 Ann Bryol 2: 140 Ann Jard Bot Buitenzorg 40: 144 Ann Bryol Suppl 1: 81 [syn sub Fr. meyeniana] Index Hepat 5: 223 Stephani Icon Hepat Inedit [Thyopsiella] F-159, microfiche 59 fig 4099 ["Luzon, Merrill 9424"] Lindenb. 1845 Nees by Schiffn. 1898 [orth var] Basionym: Distribution: orth var Schiffner Miller, Whittier & Whittier = Frullania aeolotis Frullania riparia 1898 1983 831 609 Conspect Hepat Archip Indici, Batavia: 319 Prod Flora Hepat Polynesiae: 120 Nees 1845 Hampe 1838 Page 2 acrorhiza Nees 1845 [pro syn] Typus: India. Madras sd, Wight Hep 10 in hb G [ex hb NEES] in hb E E00002522 [isotypus] Basionym: Distribution: nom herb - syn nov Nees in G., L. & N. Stephani Stephani Verdoorn Bonner Miller, Whittier & Whittier = Frullania recurvata Typus: Peru 2850 686 686 293 655 609 Syn Hepat: 437 [syn sub Fr. recurvata] Spec Hepat 4: 552 [syn sub Fr. ceylanica] Spec Hepat 4: 685 [ syn sub Fr. recurvata] Ann Bryol Suppl 1: 92 [syn sub Fr. gaudichaudi] Index Hepat 5: 223-224 Prod Flora Hepat Polynesiae: 120 Lehm. 1838 Frullania gaudichaudii aculeata 1845 1911 1911 1930 1965 1983 (Nees & Mont.) Nees & Mont. 1845 [auct. non Taylor 1846] Steph. 1909 {Peru} sd, Lechler sn Basionym: Distribution: Stephani Clark 1909 1956 Uribe & Gradstein 2003 = Frullania peruviana acuminata xxxx (Fr aculata] Bryologist 59: 219 "Stephani (op. Cit., 604, 606) clearly distinguished them, and his unpublished drawings clearly illustrate this difference." 1071 Crypt Bryol 24 (3): 200 "Material from Peru (Lechler s.n.) Identified by Stephani, in 1909, as F. aculeata and described in full by Clark (1956) under that name belongs to F. peruviana Gottsche." 378 Gottsche 1846 [non Gottsche ex Spruce 1884] Steph. 1911 Typus: Peru. Tambo de la Chonta, Monte Rosa, 3000-4000m 1876, G. Wallis, G. 109 in hb G G00069133 - n° SIB 125604/1 - 026717 (hb JACK 109) [holotypus] Basionym: Distribution: Peru, Ecuador, Galapagos Isls (Santa Cruz)[Ecuador] spec nov - syn nov Stephani Clark & Svihla Herzog Bartram & Arnell Bonner Gradstein & Weber Stephani 1911 1949 1952 1961 1965 1982 1985 Kron Grims Uribe & Gradstein Uribe 1988 2001 2003 2008 = Frullania aculeata 686 331 444 932 655 496 928 Spec Hepat 4: 606-607 Bryologist 52: 131-134, fig. 1-21(:133) Svensk Bot Tidskr 46 (1): 86 [Ecuador] Bryologist 64: 248 [Galapagos Isls: Santa Cruz] Index Hepat 5: 224 J Hatt Bot Lab 52: 148 [Galapagos; holotype] Stephani Icon Hepat Inedit [Meteriopsis] E-11, microfiche 55 fig 3780 ["Peruvia, Chimbaraso, Spruce - Monte Rosa - Wallis"] - 3781 510 J Hatt Bot Lab 64: 355, fig 6(d-e) [key] 1136 Moosherbar in hb LINZ, from Ekuador-Galapagosinseln [Internet] 1071 Crypt Bryol 24 (3): 200 [sub Fr aculeata] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Taylor 1846 Page 3 acutifolia Hampe ex Bonner 1965 [nom herb] {Peru} Typus: Peru. Tatanara sd, s coll in hb G in hb S (B22561) Basionym: Distribution: Peru Bonner Menzel Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 1965 1984 2002 655 142 782 Index Hepat 5: 224 Willdenowia 14: 488 [Peru] & 502 [= nom nud] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] = acutiloba Typus: ? Gerola 1947 [non Mitten 1861] {Ethiopia} sd, Vatoxa ? Basionym: Distribution: Ethiopia spec nov - nom nov Gerola 1947 Cufodontis Bonner Vanden Berghen Wigginton & Grolle Wigginton 1952 1965 1976 1996 2002 Wigginton 2004 Hentschel et alii 2015 = Frullania plicata 1532 Lavori di botanica / Istituto Botanica dell'Università di Padova 8: 477-478, fig. 11(a-h) [Ethiopia] 2562 Phyton 4 (1-3): 76 [Ethiopia] 655 Index Hepat 5: 224 185 Bull Jard Bot Nat Belg 46 (1/2): 12-13 [from Ethiopia] 696 Bryophyt Bibl 50: 80 1012 Trop Bryol Res Rep 2: 26 [Ethiopia] [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 28 [Ethiopia] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 4097 Phytotaxa 220 (2): 137 [nom nov Fr plicata ] Hentschel & von Konrat 2015 Page 4 acutiloba Mitt. 1861 [non Gerola 1947] Typus: India, Nilgiri Mts. "Frullania with Frullania Utacamundeana MONT. from Dr. Montagne, Nelgherries, hb Montagne" sd, Perrottet s.n. hb MONTAGNE in hb NY (hb MITTEN) [Hattori 1973] Basionym: Distribution: Japan, China (Guangxi, Yunnan) , Taiwan, India (North: Assam, Meghalaya (Khasi Hills); South: Tamil Nadu (Nilgiri Mts)), Sri Lanka, Indonesia (Java, Sumatra) spec nov - comb nov. Mitten 1861 574 Stephani Verdoorn Verdoorn Verdoorn Verdoorn 1911 1930 1932 1932 1934 686 293 554 553 241 Verdoorn 1937 556 Chopra Abeywickrama Parihar Bonner Hattori 1943 1959 1962 1965 1973 205 200 1584 655 541 Hattori 1975 47 Vanden Berghen Abeywickrama & Jansen Hattori Hattori Onraedt Hattori & Lin Stephani 1976 1978 1979 1980 1981 1985 1985 185 794 129 489 5794 505 928 Piippo Chao & Lin Furuki & Mizutani Parihar, Lal & Katiyar Lin & Chen Nath & Asthana Nath & Asthana Bapna & Kachroo Wu & Wang Bai in Gao & Cao Lin Bai Bai Srivastava & Alam Singh 1990 1992 1994 1994 1997 1998 1998 2000 2000 2000 2000 2001 2002 2002 2002 521 460 1331 2196 1130 683 708 712 2671 1019 1524 1742 1189 1251 4886 Carthy Gao & Lai Zhu & So Glenny, Meagher & Brown 2003 2003 2003 2004 1125 1441 1083 1127 Glenny, Meagher & Brown 2004 1126 Nair et al. Singh, Kumar & Nath 2005 2008 3031 1281 Gao & Wu Wang, Lai & Zhu Long & Rubasinghe Hentschel et alii Hardianty 2010 2011 2014 2015 2018 2349 3125 3994 4097 5036 J Linn Soc Bot 5: 120 [India: Nilgiri] "Hab. In montibus Nilgiri, Perrottet." Spec Hepat 4: 464 "Hab. India orientalis, Nilgherry Montes; Ceylon" Ann Bryol Suppl 1: 46, fig. 29 & 31-34(var. Schiffneri) Nederl Kruidk Arch Jrg 1932 (2): 485 [Java] Bull Jard Bot Buitenzorg, ser 3, 12 (1): 55 [Java] Ann Bryol Suppl 4: 203 [India] "Vorderindien: Shembaganur, Palni Hills, 6000', Foreau 1931" Blumea suppl 1: 211 [South India] "S. India: Kodaikanal, R. P. FOREAU, III. 1932." J Indian Bot Soc 22: 249 [India] Ceylon J Sci (Bio Sci) 2 (1): 58 [Sri Lanka] Univ Allahabad studies, Bot sect 1-2: 18 [India: SI] Index Hepat 5: 224 J Hatt Bot Lab 37: 99-101, fig 53(from type)-54 [Java] "Nilgheries (Nilgiri), herb. Montagne, leg. Perrottet, type in herb. NY." Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 73-74 [comb. nov. = Fr. hampeana var acutiloba (MITT) HATT ] Bull Jard Bot Nat Belg 46 (1/2): 13 MaB Nat Comm Sri Lanka, Publ 1: 5 [Sri Lanka] J Jap Bot 54 (1): 22 [Sri Lanka] J Hatt Bot Lab 47: 103 [acutiloba = distinct species] J Hatt Bot Lab 50: 201 [Sri Lanka] J Hatt Bot Lab 59: 131 [China] Stephani Icon Hepat Inedit [Galeiloba] C-100ab, microfiche 51 fig 3482 ["olim Frull nilgirienses St, Neelgherries, Nilgherry Mtes"] - 3483 ["Ceylon, Poendeloya?, J Nietneri 1868, Jack 1548"] J Hatt Bot Lab 68: 37 [annotated catalogue of China] Yushania 9: 197, fig 1(a-f) [Taiwan] Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] Hepat Anth India, annotated checklist: 21 [South India] Biol Bull 75: 83, fig 1(a-d) [Taiwan, key to species, in Taiwanese] J Bryol 20(1): 233 [South India] J Hatt Bot Lab 85: 72, fig 5(1-15) [South India; key to all spec.] Hepaticology in India II: 165-166 Bryoflora of Hengduan MTS (Southwest China): 115, fig. 49(1-5) Fl Yunnanica 17: 392, fig 195(10-15) [China, Yunnan] Liverwort Flora Taiwan: 155, fig p 156(1-3) [Taiwan] Guizhou Science 19(4): 32 [China, Yunnan] Chenia 7: 2 [Yunnan, China] J Bombay Nat Hist Soc 99(2): 237 [Nilgiri, key to South Indian species] Studies on Liverworts (Bryophyta) of Khasi and Jaintia Hills: Meghalaya. 257-258, pl 51 (1-16), map 11 - Unpublished thesis. Flora Australia Suppl Ser 21: 112 [not in Australia] Illustrations Bryophytes China: 1099, fig 511(10-15) [China] Crypt Bryol 24 (4): 328 [Guangxi Prov, China] Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Bryophytes of Wayanad in Western Ghats: 62 [India, Western Ghats] Taiwania 53 (1): 63, fig 1[map], 5(1-16) [India, Meghalaya, Eastern Himalayas] Genera Hepat. et Anth. Sinicorum: fig. 232(10-15) [key in Chinese] Annales Botanici Fennici 48: <369-395> [Checklist Taiwan] Ceylon Journal of Science 43(1): 3 [Sri Lanka] Phytotaxa 220 (2): [as Fr. monocera var. acutiloba] Frullaniaceae di Taman Wisata Alam Sicike-Cike: 18, photos [Sumatra, with key] Page 5 Daniels & Kariyappa 2019 5109 Bryoflora of Agasthyamalai Biosphere Reserve: 387-389, fig 139(a-m) [India (Western Ghats, keys] = Frullania monocera var. acutiloba (Mitt.) Hentschel & von Konrat 2015 acutiloba [auct. non Mitt. 1861 nec Gerola 1947] Yuzawa & S.Hatt. 1981 {Japan} Basionym: Distribution: - syn nov Yuzawa & Hattori Yuzawa Yamada & Iwatsuki = Frullania monocera 1981 2001 2006 494 J Hatt Bot Lab 50: 253 [Japan] 1660 Natural Environmental Sci Res 14: 23 [Japan, syn sub Fr monocera] 1470 J Hatt Bot Lab 99: 28 [Catalog of Japan, syn sub Fr monocera] (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 acutiloba var. schiffneri Verd. 1929 Typus: Indonesia. Java, Prop. Praeanger, "Kratersee" Telaga Bodas, 1580m 15.2.1894, Schiffner 2002 Basionym: Distribution: China, Taiwan, Sri Lanka, Indonesia (Java) var nov Verdoorn Verdoorn Verdoorn Hattori 1929 1930 1932 1975 245 293 553 47 - comb nov Hattori 1979 486 Hattori & Lin Yamada, Yuzawa, Ogasawara & Lai Kuo & Chiang Piippo Lin & Chen Bapna & Kachroo 1985 1986 505 950 Ann Bryol 2: 123-124, fig 2 Ann Bryol Suppl 1: 48, fig 31-34, 55 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 55 [Java] Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 73-74 [as Fr. hampeana var. acutiloba] J Hatt Bot Lab 46: 120, footnote [comb nov as Fr. monocera var. schiffneri] J Hatt Bot Lab 59: 131 [China] J Taiwan Mus 39 (1): 63 [Taiwan] 1988 1990 1997 2000 948 521 1130 712 Taiwania 33: 11 [Index of Taiwan] J Hatt Bot Lab 68: 37 [annotated catalogue of China] Biol Bull 75: 83 [syn sub Fr acutiloba, in Taiwanese] Hepaticology in India II: 166 = Frullania monocera var. schiffneri (Verd.) S.Hatt. 1979 acutistipula Steph. 1911 Typus: Papua New Guinea: Milne Bay, Cloudy Mountains -.12.1894, Micholitz, W. sn in hb G G00066972 - n° SIB 124870/1 - 14395 [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Clark Bonner Miller, Whittier & Whittier Grolle & Piippo Stephani = Frullania apiculata 1911 1930 1930 1955 1965 1983 1984 1985 686 293 549 369 655 609 143 928 Spec Hepat 4: 550 Ann Bryol Suppl 1: 97 & 101 Nova Guinea 14 (4): 541 [syn sub Fr. apiculata fo. warburgii] Bryologist 58: 68 [syn sub Fr. apiculata] Index Hepat 5: 225 Prod Flora Hepat Polynesiae: 120 Acta Bot Fennica 125: 17 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Thyopsiella] F-160, microfiche 59 fig 4100 ["N Guinea, Cloudy mts, Micholitz"] fo. warburgii Verd. 1930 Frullania apiculata (Reinw., Blume & Nees) Nees 1845 Page 6 aeolatis [auct. non Mont. & Nees] C.Massal. by S.Hatt. & LIN 1985 [orth var] Basionym: Distribution: Hamlin Hattori & Lin Hattori & Lin = Frullania aeolotis 1972 1985 1985 Rec Dominion Mus 7(19): 261 [New Zealand] J Hatt Bot Lab 59: 146 [sub Fr subdilatata] J Hatt Bot Lab 59: 143 [sub Fr. sinensis] [auct. non MONT & Nees] C.Massal. 1897 Frullania sinensis aeolotis 659 505 505 Steph. 1906 Mont. & Nees in Mont. in La Sagra 1842 Basionym: Distribution: China spec nov Montagne in La Sagra Montagne & Nees in Montagne Montagne in La Sagra Parihar Miller, Whittier & Whittier Piippo Yamada & Iwatsuki 1842 1842 1845 1962 1983 1990 2006 26 469 25 1584 609 521 1470 L'ile de Cuba [Franz Ed]: 461 Ann Sci Nat Bot, ser 2, 18: 14 [as Fr.aeolotis var. squarrosa] La Isla de Cuba [Span Ed] 9: 276 Univ Allahabad studies, Bot sect 1-2: 19 [India, syn sub Fr squarrosa] Prod Flora Hepat Polynesiae: 120 J Hatt Bot Lab 68: 37 [annotated catalogue of China] J Hatt Bot Lab 99: 28 [Catalog of Japan] = Frullania aeolotis var. squarrosa Mont. & Nees in La Sagra 1842 Frullania squarrosa (Reinw., Blume & Nees) Nees 1845 Frullania ericoides aeolotis (Nees) Mont. 1839 [auct. non MONT & Nees] C.Massal. 1897 Basionym: Distribution: - syn nov Massalongo Levier Hattori & Lin Hattori & Lin Bapna & Kachroo = Frullania sinensis aeolotis 1897 1906 1985 1985 2000 1914 824 505 505 712 Mem Accad Agr Art Comm Verona, ser 3, 73 (2): 38 Nuovo Giorn Bot Ital NS 13: 349 [syn nov sub Fr. sinensis] J Hatt Bot Lab 59: 143 [as "aeolatis" sub Fr. sinensis] J Hatt Bot Lab 59: 146 [as "aeolatis" sub Fr subdilatata] Hepaticology in India II: 181 [sub Fr. sinensis] Steph. 1906 Nees & Mont. ex Nees 1854 [pro syn] {hb MONT No 125} Basionym: Distribution: Nees in G., L. & N. 1845 2850 Syn Hepat: 418 [syn sub Fr aeolotis var singapurensis] = Frullania aeolotis var. singapurensis Nees 1845 Frullania riparia Hampe 1838 Page 7 aeolotis Nees 1845 Typus: in hb GLM (hb LEHMANN) 05942 (ex hb GOTTSCHE) Basionym: Distribution: Lord Howe Isl spec nov auct - syn nov Nees in G., L. & N. Mitten Gottsche Gottsche Mitten in Seemann Britton Mitten Stephani Evans Sawada 1845 1861 1863 1867 1871 1889 1891 1897 1897 1909 Stephani Müller in Rabenhorst Chopra Frye & Clark Kitagawa Stephani 1910 1915 1943 1947 1973 1985 Hattori 1987 = Frullania riparia aeolotis 2850 574 103 104 637 2441 582 213 237 930 Syn Hepat: 417-418 J Linn Soc Bot 5: 119 [India: Kumaon, Sikkim, Khasi] Separata of Mex. Leverm.: 239 K Danske Vid Selks Skrifter, ser 5, 6: 335 [Mexico] Flora Vitiensis: 417 [as Fr. alolotis; Lord Howe Isl] Catalogue of plants in New Jersey [USA, New Jersey]: 347 Trans Linn Soc London, ser 2, 3: 202 [Japan] Bull Herb Boissier 5: 78 [Japan] Trans Connet Acad Arts Sci 10 (1): 13 [syn sub Fr. riparia] Bot Mag Tokyo 23 (no 268): 178-183 [key to Japanese species, in Japanese] 686 Spec Hepat 4: 433 1064 Krypt Flora (2ed) 6, 2: 629 [sub Fr riparia] 205 J Indian Bot Soc 22: 249 [India] 204 Univ Wash Publ Biol 6 (5): 774 404 Misc Bryol Lichen 6 (6): 102-103 928 Stephani Icon Hepat Inedit [Galeiloba] C- 90a, microfiche 51 fig 3466 ["Fr aeolotis Nees, quoad plant american, America, Austin 101" = Fr. riparia Hampe] 400 Bryologist 90 (4): 370 [not on Lord Howe Isl] Hampe 1838 Mont. & Nees ex Nees 1838 [nom nud] Basionym: Distribution: Montagne & Nees in Nees Evans 1838 1897 261 237 Frye & Clark 1947 204 = Frullania aeolotis Frullania riparia aeolotis Naturg Europ Leberm 3: 210 [footnote with nomen only] Trans Connet Acad Arts Sci 10 (1): 13 [is nom nud, synomy syn sub Fr. riparia] Univ Wash Publ Biol 6 (5): 774 [sub Fr riparia] Nees 1845 Hampe 1838 [auct. non MONT & Nees] Milde 1864 [Gratsch bei Meran] Basionym: Distribution: Milde Milde = Frullania cesatiana Frullania riparia 1864 1865 3795 Botanische Zeitung (Berlin) 22, Beilage 3: 9 & 16 2434 Botanische Zeitung (Berlin) 23 (19): 155 [syn sub Fr cesatiana] De Not. 1865 Hampe 1838 Page 8 aeolotis var. (beta) aberrans C.Massal. 1897 Typus: China. Shensi, Lan-san-huo sd, Giraldi hb MASSALONGO 29, hb LEVIER Basionym: Distribution: China var nov - syn nov Massalongo Bonner Hattori Hattori & Lin Piippo Bapna & Kachroo Mizutani, Hasegawa & Iwatsuki = Frullania muscicola 1897 1965 1974 1985 1990 2000 2009 1914 655 545 505 521 712 2112 Mem Accad Agr Art Comm Verona, ser 3, 73 (2): 39 Index Hepat 5: 225 [syn sub Fr. muscicola] J Hatt Bot Lab 38: 219 [syn sub Fr. dilatata ssp. subdilatata] J Hatt Bot Lab 59: 140-141, fig 3 [syn nov sub Fr. muscicola] J Hatt Bot Lab 68: 37 [annotated catalogue of China] Hepaticology in India II: 175 [syn sub Fr. muscicola] Type sepecimens Hatt Bot Lab (NICH) 20: Isolectotype. China, Schensi. Nov. 1895. R. J. Gira/di (NICH 354618). & Syntype. China, Schensi. Nov. 1895. R. J. Gira/di 31c (NICH 354624). Other twö syntypes: NICH 354641 & NICH 354708. Steph. 1894 aeolotis var. auriculata Mont. 1842 Basionym: Jungermannia squarrosa Distribution: Indonesia (Java) var nov Reinw., Blume & Nees 1824 [hom illeg] [vide Grolle 1995] Montagne in La Sagra Montagne in La Sagra Bonner = Frullania riparia 1842 1845 1965 26 25 655 L'ile de Cuba [Franz Ed]: 462 La Isla de Cuba [Span Ed] 9: 276 Index Hepat 5: 225 Hampe 1838 aeolotis var. biformis Nees 1845 Typus: USA. "prope Baltimore or Ebenezar" sd, Beyrich in hb GLM (hb LEHMAN) 05943 Basionym: Frullania biformis Hampe ex Nees 1845 [pro syn] Distribution: Nees in G., L. & N. Bonner = Frullania riparia 1845 1965 2850 Syn Hepat: 418 655 Index Hepat 5: 225 Hampe 1838 aeolotis var. evoluta Mont. 1842 Basionym: Jungermannia ericoides Distribution: var nov Nees in Martius 1833 Montagne in La Sagra Montagne in La Sagra Bonner = Frullania riparia 1842 1845 1965 26 25 655 L'ile de Cuba [Franz Ed]: 462 La Isla de Cuba [Span Ed] 9: 276 Index Hepat 5: 226 Hampe 1838 Page 9 aeolotis var. laxa Mont. 1842 Basionym: Frullania aeolotis Distribution: var nov Mont. & Nees ex Nees 1838 [nom nud] Montagne in La Sagra Montagne in La Sagra Bonner = Frullania riparia 1842 1845 1965 26 25 655 L'ile de Cuba [Franz Ed]: 461-461 La Isla de Cuba [Span Ed] 9: 276 Index Hepat 5: 226 1845 1965 2850 Syn Hepat: 418 655 Index Hepat 5: 226 1845 1910 1990 2850 Syn Hepat: 417-418 686 Spec Hepat 4: 433 105 Mem NY Bot Gard 63: 40 [Mexico] Hampe 1838 aeolotis var. obliqua Nees 1845 Basionym: Distribution: Nees in G., L. & N. Bonner = Frullania riparia Hampe 1838 aeolotis var. riparia Nees 1845 Typus: in hb GLM (hb LEHMANN) 05944 Basionym: Distribution: - syn nov Nees in G., L. & N. Stephani Fulford & Sharp = Frullania riparia Hampe 1838 aeolotis var. singapurensis Nees 1845 Typus: Singapore. in hb MONTAGNE & hb NEES in hb S (B22567) [typus] Basionym: Distribution: Nees in G., L. & N. Bonner Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot = Frullania riparia 1845 1965 2002 2850 Syn Hepat: 418 655 Index Hepat 5: 226 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] Hampe 1838 Page 10 aeolotis var. squarrosa Mont. & Nees in La Sagra 1842 Basionym: Distribution: var nov Montagne & Nees in La Sagra Montagne & Nees in La Sagra Montagne 1842 1842 1845 - syn nov Nees in G., L. & N. Verdoorn Frye & Clark Kamimura Bapna & Kachroo 1845 1930 1947 1961 2000 = Frullania squarrosa (Reinw., Blume & Nees) Nees 1845 Frullania ericoides africana 26 L'ile de Cuba [Franz Ed]: 461 469 Ann Sci Nat Bot, ser 2, 18: 14 "var squarrosa" 1591 Voyage au pole sud 1837-40, Paris, 3 Botanique, vol 1 Cryptog: 222223 2850 Syn Hepat: 417 [syn sub Fr. ericoides] 293 Ann Bryol Suppl 1: 34 [syn sub Fr. squarrosa] 204 Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] 199 J Hatt Bot Lab 24: 17 [syn sub Fr squarossa] 712 Hepaticology in India II: 170 [syn sub Fr. ericoides] (Nees) Mont. 1839 Steph. 1891 Typus: W Africa. Sao Tomé et Principe "Ins. San Thomé" 1885, A.F. Moller, 38 in hb G G00045270 - n° SIB 148709/1 [holotypus] Basionym: Distribution: Democratic Republic of Congo, San Thomé Isl, Ghana, Guinea, Nigeria, Cameroon, Gabon, Congo spec nov - syn nov Stephani Stephani Pearson Bonner Vanden Berghen Augier 1891 1910 1924 1965 1973 1974 Vanden Berghen Vanden Berghen Jones & Harrington Stephani 1976 1976 1983 1985 Hattori Yuzawa Fischer Wigginton Wigginton 1988 1991 1993 2004 2004 = Frullania rio-janeirensis akisawai 209 686 795 655 1702 821 Hedwigia 30 (6): 269-270, pl 32, 33(8-10) [San Thome Isl] Spec Hepat 4: 322 Arkiv Botanik 19 (5): 9 [Mt. Elgon in Kenia/Uganda] Index Hepat 5: 227 Rev Bryol Lichen 39: 370-372, fig 2(a-e) [Gabon] Annales de la Faculté des Sciences du Cameroun 17: 78, 80-82, 8586 [Cameroon] 185 Bull Jard Bot Nat Belg 46 (1/2): 151-154, fig. 37 850 Distributiones plantarum africanarum 11: 362 [map of Africa] 251 Bull Brit Mus nat Hist (Bot) 11 (3): 249-250 [Ghana] 928 Stephani Icon Hepat Inedit [Chonanthelia] A- 1, microfiche 46 fig 3063 [from type; "S Thome, no 38, leg Moller"] 516 J Hatt Bot Lab 65: 443 [syb syn sub Fr. riojaneirensis] 525 J Hatt Bot Lab 70: [aus 696] 308 Trop Bryol 8: 44 [Checklist: Zaire] 1312 Trop Bryol Res Rep 5: 28 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 226 [syn sub Fr riojaneirensis, West Africa] (Raddi) Ångstr. 1876 Kamim. 1970 Basionym: Distribution: nom nov Kamimura 1970 -syn nov Inoue Hattori & Lin 1972 1985 Yamada & Iwatsuki 2006 = Frullania tamarisci ssp. obscura 1552 Bulletin of Kochi Gakuen Junior College 2: 24 [nom nov for Fr. pendula KAMIMURA 1952; Taiwan] 123 J Jap Bot 47 (11): 349 [syn sub Fr. kamimurae] 505 J Hatt Bot Lab 59: 161 [syn sub F. tamarisci subsp. obscura (Verd.) Hatt] 1470 J Hatt Bot Lab 99: 28 [Catalog of Japan] (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 Page 11 albertii Steph. ex Herzog 1916 Typus: Bolivia, Samaipata -.4.???? / sd, T. Herzog 5127 in hb G G00066973 - n° SIB 124869/1 [holotypus] in hb L [isotypus] Basionym: Distribution: Burundi, Kenya, Bolivia, Colombia, Ecuador spec nov Stephani ex Herzog Stephani Bonner Stephani 1916 1924 1965 1985 Yuzawa 1991 Yuzawa Uribe & Gradstein Gradstein in Luteyn 1991 1998 1999 Uribe & Rangel-Ch Wigginton 2000 2002 Wigginton 2004 = Frullania standaertii allionii 222 687 655 928 Biblioth Bot 87: 237-238, fig 185 [Bolivia, Chonanthelia, "alberti"] Spec Hepat 6: 531 [as spec nov, Bolivia, Chonanthelia, "alberti"] Index Hepat 5: 227 ["alberti"] Stephani Icon Hepat Inedit [Chonanthelia] A- 57, microfiche 47 fig 3151 ["(Chonanthelia) Bolivia Herzog 5127"] - 3152 ["(Galeiloba) Bolivia Herzog 5127"] 525 J Hatt Bot Lab 70: 205-207, fig 13, map 2b(:281) ["albertii"; may be a high-altitude form of Fr standaertii] 525 J Hatt Bot Lab 70: 205 [syn sub Fr. albertii] 707 Bryophyt Bibl 53: 35 [Catalogue Colombia] 994 Mem New York Bot Gard 84: 58 [Paramo 3400-3950m: Colombia, Ecuador] 1090 Colombia, diversidad biótica III, Bogota: 444 [Colombia] 1012 Trop Bryol Res Rep 2: 27 [Burundi, Kenya] [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [Burundi, Kenya] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Steph. 1910 Steph. 1910 Typus: Ecuador. Gualaquiza. "Hacienda Vega" - in arbore silva supra rus antiq. „hacienda Vega“ 6443 13.2.1909, M. Allioni sn, Bryotheca Levier in hb G G00066975 - n° SIB 125427/1 (ex BRYOTHECA E. LEVIER 6443) [lectotypus] in hb G 3 more syntypes [vide Gradstein 2021] Basionym: Distribution: Costa Rica, Ecuador spec nov Stephani Bonner Stephani 1910 1965 1985 - syn nov Gradstein Gradstein & Perez 2021 2021 = Frullania ericoides allophylla 686 655 928 Spec Hepat 4: 394 [Ecuador, Gualaquisa] Index Hepat 5: 227 [Ecuador, Costa Rica] Stephani Icon Hepat Inedit [Galeiloba] C- 48, microfiche 50 fig 3403 [from type; "Ecuador, Allioni 1909, Levier 6443"] 5642 Mem NY Bot Gard 121: 365 [syn sub Fr ericoides] 6056 Candollea 76: 44 [Lectotypus, Syntypi - Allioni -Ecuador] (Nees) Mont. 1839 (Hooker & Taylor) Hooker 1867 Typus: Campbell Is. Basionym: Jungermannia allophylla Hooker & Taylor 1844 Distribution: Campbell Isl [New Zealand] comb nov Hooker Hooker Taylor in Stephani Bonner Hamlin = Lepidolaena allophylla 1867 1867 1909 1965 1972 1154 1154 686 655 659 Handbook New Zealand Flora: 538 [Campbell's Island] Handbook New Zealand Flora: 755 [remove to Polyotus] Spec Hepat 4: 47 [syn sub Lepidolaena] Index Hepat 5: 228 Rec Dominion Mus 7(19): 261 [New Zealand] (Hooker & Taylor) Trevisan 1877 Lepidolaena reticulata (Hooker & Taylor) TREVIS 1877 Page 12 alolotis Nees by Mitt. in Seemann 1871 [orth var] Basionym: Distribution: orth var Mitten in Seemann Miller, Whittier & Whittier = Frullania aeolotis 637 609 Flora Vitiensis: 417 Prod Flora Hepat Polynesiae: 120 Nees 1845 Frullania riparia alpina 1871 1983 Hampe 1838 Steph. 1911 Typus: Ecuador. Tungurahua, Cerro Mulmul - "in Mte. Mulmul, in fruticosum cortice vetusto" sd, Spruce, R., sn in hb G G00066974 - n° SIB 125556/1 - 13582 ("Spruce's Original" ex hb SPRUCE sub Fr. flexicaulis SPR.) Basionym: Distribution: Ecuador spec nov Stephani Bonner Stephani 1911 1965 1985 Yuzawa 1987 = Frullania flexicaulis Spec Hepat 4: 533-534 Index Hepat 5: 228 Stephani Icon Hepat Inedit [Thyopsiella] F-100, microfiche 58 fig 4022 ["olim Frull flexicaulis Spr, in Mte Mulmul (ist aber nicht flexicaulis) leg Spruce"] - 4023 5944 Annotated by Y. Yuzaw 1987: note in hb G: "Frullania alpina STEPH. = Frullania flexicaulis SPRUCE [http://www.villege.ch/musinfo/bd/cjb/chg/adetail.php?id=113730&base=img&lang=fr] Spruce 1884 Frullania paradoxa alstonii 686 655 928 Lehm. & Lindenb. 1845 [auct. non Verd. 1930] Herzog & Noguchi 1955 {Taiwan} Basionym: Distribution: Taiwan Herzog & Noguchi Lai & Wang-Yang 1955 1976 Tan & Engel Kuo & Chiang Piippo 1986 1988 1990 = Frullania densiloba 859 33 J Hatt Bot Lab 14: 43 [Botel Tobago] Taiwania 21 (2): 168 [specimen from Taiwan by Herzog & Noguchi belongs to Fr. densiloba] 4286 J. Hatt. Bot. Lab.. No. 60: 301 [Philippines] 948 Taiwania 33: 11 [Index of Taiwan] 521 J Hatt Bot Lab 68: 139 annotation nr 137 Steph. ex A.Evans 1906 Page 13 alte-mammillata S.Hatt. 1972 Typus: New Guinea: NO-Neuguinea [Chimbu, Western Highlands, Madang], Central Highlands, Mt. Wilhelm, Lake Aunde, in low subalpine forest, 3800 m 2.6.1967, P.M. Schuster 67-6537 in hb NICH [holotypus] in hb JE 04004502 (1967, Schuster,R.M. 67-6537/a( [isotypus] in hb JE 04000761 (1967, Schuster,R.M. 67-6050/a) [paratypus], in hb JE 04000760 (1967, Schuster,R.M. 67-6040/a2) [paratypus] Basionym: Distribution: Borneo, Philippines, New Guinea spec nov - syn nov Hattori Hattori Hattori Hattori Hattori Hattori Grolle & Piippo Hattori & Piippo Tan & Engel Mizutani, Hasegawa & Iwatsuki = Frullania armatifolia amakawana 1972 1973 1974 1975 1976 1982 1984 1986 1986 2009 539 542 546 547 80 495 143 557 4286 2112 J Hatt Bot Lab 36: 435-437, fig. 15 J Hatt Bot Lab 37: 148 [New Guinea] J Hatt Bot Lab 38: 223-224 J Hatt Bot Lab 39: 277-278 [Philippines, Borneo] J Hatt Bot Lab 40: 470-472, fig. 179-180 J Hatt Bot Lab 51: 235 Acta Bot Fennica 125: 17 [Cat. West Melanesia] Acta Bot Fennica 133: 38 [syn sub Fr. armatifolia] J. Hatt. Bot. Lab.. No. 60: 301 [Philippines] Type sepecimens Hatt Bot Lab (NICH) 20: Holotype. New Guinea, Central Highlands, Mt. Wilhelm. 11600 ft. In low subalpine Podocarpus comactus-Rapanea-Prunus pullei-Olearia forest. 2 June 1967. R. M. Schuster 67-653 7 (NICH 309936). Verd. 1932 Kamim. ex Shin 1968 [pro syn] Typus: Ryukyu, Amami Basionym: Distribution: nom nud - syn nov Kamimura ex Shin Shin = Frullania amamiensis amboinensis 1968 1970 844 150 Sci Rep Kagoshima Univ 17: 203 [Ryukyu Arch.] Sci Rep Kagoshima Univ 19: 25 [Ryukyu: Amami Isl] Kamim. 1968 Schiffn. 1890 Typus: Indonesia. Amboina 7.6.1875, Naumann (Gazelle Exped.) Basionym: Distribution: spec nov - comb nov Schiffner Schiffner Schiffner & Stephani in Schumann & Lauterbach Stephani Verdoorn 1889 1898 1901 581 Forschungsreise S.M.S. "Gazelle" 4 (4): 39 831 Conspect Hepat Archip Indici, Batavia: 319 1493 Flora Deutsch Schutzgebiete: 78 1911 1930 686 293 Bonner Hattori Grolle & Piippo 1965 1980 1984 655 490 143 = Frullania intermedia Spec Hepat 4: 588 Ann Bryol Suppl 1: 163-164 [cmb nov as Fr. Intermedia var. amboinensis] Index Hepat 5: 228 J Hatt Bot Lab 47: 187 Acta Bot Fennica 125: 17 [Cat. West Melanesia] (Reinw., Blume & Nees) Nees 1845 Page 14 amentacea Herzog 1952 Typus: Ecuador. Prov. Santiago-Zamora, Mendez, woody hill, 700 m 24.5.1947, Gunnar Harling 2225a pp [Typus] in hb JE [holotypus & isotypus] in hb S B22568,B22569, B22570) [dupl] Basionym: Distribution: Ecuador spec nov - syn nov Herzog Bonner Bisang, Schumacker, Sergio & Grolle Yuzawa Müller & Zündorf = Frullania megalostipa 1952 1965 1989 444 655 276 Svensk Bot Tidskr 46 (1): 83-85, fig. 15 Index Hepat 5: 229 Giorn Bot Ital 122: 1991 2007 525 J Hatt Bot Lab 70: 250, 252 [syn sub Fr. megalostipa] 1442 http://www2.unijena.de/biologie/spezbot/herbar/bryotypes/Uebersicht%20F.htm Spruce 1884 Frullania rio-janeirensis var. megalostipa (Spruce) Gradstein 2020 amoena Steph. 1911 Typus: Bolivia. "Andes Boliviae", Grat des Cerro Amboro, 1450m -.10.1907, Th. Herzog, 58847 in hb G (hb LEVIER 5847) G00069155 - n° SIB 124201/1 Basionym: Distribution: Bolivia, Ecuador, Brazil spec nov - syn nov Stephani Schiffner & Arnell 1911 1964 Bonner Yano Stephani 1965 1984 1985 Yuzawa & Koike Yano Gradstein, Meneses & Arbe Gradstein & Costa Yano Winter Schäfer-Verwimp 1989 1999 2003 2003 2008 2020 = Frullania obcordata amplexicaulis 686 198 Spec Hepat 4: 635-636 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 86-87, pl 13(147) [Brazil] [mit Ocellen] 655 Index Hepat 5: 229 501 J Hatt Bot Lab 56: 503 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Diastaloba] B-27, microfiche 48 fig 3259 [from type; "Bolivia (Dr Herzog legit)"] 519 J Hatt Bot Lab 66: 350-351, fig 6 [from type] [syn sub Fr. gymnotis] 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] 975 J Hatt Bot Lab 91: 52 [Bolivia; syn. sub Fr caulisequa (Nees) Nees]] 984 Mem New York Bot Gard 87: 86 [Brazil] [sub Fr caulisequa] 1684 Boletim Instituto Botanica 19: 89 [syn sub Fr gymnotis; Brazil; catalog] 5742 Frahmia 19: 12 [syn sub. Fr obcordata] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Colenso by E.A.Hodgs. 1949 [orth var] Basionym: Distribution: Hodgson Bonner = Frullania implexicaulis Frullania deplanata 1949 1965 203 655 Trans Roy Soc New Zealand 77 (3): 387 Index Hepat 5: 229 Colenso 1887 Mitt. 1855 Page 15 amplicrania Steph. ex Yoshinaga 1901 [nom nud] Basionym: Distribution: - nom nud Stephani ex Yoshinaga Stephani in Sawada 1901 1909 Hattori Kamimura Kamimura Yamada & Iwatsuki 1944 1952 1961 2006 = Frullania amplicrania anamensis 889 930 Bot Mag Tokyo 15: 92 Bot Mag Tokyo 23 (no 268): 178-183, fig 22 [key to Japanese species, in Japanese] 943 Bull Tokyo Sci Mus 11: 141-142, fig 87[a-m) [= nom nud] 873 Contrib Fl Hepat Shikokuensem, Kochi: 122 199 J Hatt Bot Lab 24: 15 1470 J Hatt Bot Lab 99: 28 [Catalog of Japan] Steph. 1910 Steph. 1911 Typus: Lao People's Democratic Republic. Houaphan, Muong Ham 18.12.1906, Mission Sci d´explortation de l´Indo-Chine 168 in hb Gen PARIS 168 in hb G 00128079 - n° SIB 143803/3 - 14396 Basionym: Distribution: Laos spec nov - syn nov Stephani Bonner Hattori Miller, Whittier & Whittier Stephani 1911 1965 1972 1983 1985 686 655 538 609 928 Konrat & Braggins 2001 838 = Frullania apiculata andina Spec Hepat 4: 551 Index Hepat 5: 230 J Hatt Bot Lab 36: 109, fig. 1 [syn nov sub Fr. apiculata] Prod Flora Hepat Polynesiae: 120 Stephani Icon Hepat Inedit [Thyopsiella] F-161, microfiche 59 fig 4101 ["Laos, Indochina, Herb Gén Paris 168"] J Hatt Bot Lab 91: 256-262 [syn sub Fr. apiculata] (Reinw., Blume & Nees) Nees 1845 Steph. 1910 Typus: Peru. Andes coll. ign. / sd, Spruce, sn in hb G (ex hb Spreng. ) G00066978 - n° SIB 125606/1 [holotypus] Basionym: Distribution: Peru, Bolivia spec nov - syn nov Stephani Stephani Bonner Menzel Stephani 1910 1916 1965 1984 1985 686 222 655 142 928 Yuzawa Yuzawa Gradstein, Meneses & Arbe 1991 1991 2003 525 525 975 = Frullania rio-janeirensis Spec Hepat 4: 326-327 Biblioth Bot 87: 241 [Bolivia] Index Hepat 5: 230 Willdenowia 14: 488 [Peru] Stephani Icon Hepat Inedit [Chonanthelia] A- 17, microfiche 46 fig 3089 [from type; "Andes Herb Sprengel"] J Hatt Bot Lab 70: 255-257, fig. 38 [description of type specimen] J Hatt Bot Lab 70: 253 {syn sub Fr. riojaneirensis (Raddi) Aongstroem] J Hatt Bot Lab 91: 52 [Bolivia; as sub Fr. riojaneirensis (Raddi) Aongstroem] (Raddi) Ångstr. 1876 Page 16 angolensis Steph. 1910 Typus: Angola. Hüilla -.3.1860, Welwitsch 251 in hb G G00045271 - n° SIB 148710/1 - 17303 [holotypus], Basionym: Distribution: Angola spec nov - syn nov Stephani Demaret & Vanden Berghen Arnell Bonner Vanden Berghen Stephani 1910 1946 1963 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania borgenii Spec Hepat 4: 366 Bull Jard Bot Etat Bruxelles 18: 122 [syn sub Fr borgeni] Hepat S Africa: 147 [syn sub Fr. borgenii] Index Hepat 5: 230 Bull Jard Bot Nat Belg 46 (1/2): 145 Stephani Icon Hepat Inedit [Galeiloba] C- 4, microfiche 50 fig 3351 [from type; "Angola, Huilla, leg 1860 Welwitsch 251"] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 233 [syn sub Fr obscurifolia, West Africa] Pearson 1890 Frullania obscurifolia angstroemii 686 1102 822 655 185 928 Mitt. 1879 A.Evans ex Steph. 1910 Typus: Hawaii. Oahu, Luakaha 9.7.1898 C.M. Cooke Jr sn in hb YU (ex hb EVANS) [holotypus] in hb G [isotypus] in S-B22571 , in hb MU [paratypus] Basionym: Distribution: Hawaii (Oahu) spec nov - syn nov Stephani Stephani Verdoorn 1910 1910 1930 Verdoorn Miller Clark Miller 1930 1953 1954 1963 Bonner Bonner Miller, Whittier & Whittier Miller, Whittier & Whittier Stephani 1965 1965 1983 1983 1985 Yuzawa Myrdal So & Wang 1991 2002 2006 = Frullania sandvicensis 686 686 293 Spec Hepat 4: 354 ["angstroemii" EVANS ms.] [as Chonanthelia] Spec Hepat 4: 354-355 [as Chonanthelia] Ann Bryol Suppl 1: 66 [syn nov sub Fr. neurota] was under Fr aongstroemii 551 Nederl Kruidk Arch Jrg 1930 (2): 349 Bryologist 56: 45 [sub Fr neurota] was under Fr aongstroemii 362 Bryologist 57: 29-30 was under Fr aongstroemii 834 Ark Bot, ser 2, 5: 518 [syn sub Fr. neurota] -was under Fr aongstroemii 655 Index Hepat 5: 233 -was under Fr aongstroemii 655 Index Hepat 5: 231 [orth var of Fr aongstroemii] 609 Prod Flora Hepat Polynesiae: 120 609 Prod Flora Hepat Polynesiae: 121 -was under Fr aongstroemii 928 Stephani Icon Hepat Inedit [Chonanthelia] A- 94, microfiche 47 fig 3203-3204 ["Hawaii ..."] -was under Fr aongstroemii 525 J Hatt Bot Lab 70: 221 -was under Fr aongstroemii 1276 Mosskollekter [Frullania types in hb S] -was under Fr aongstroemii 1466 J H B L 100: 426-428 [syn nov sub Fr sandvicensis] Ångstr. 1872 Page 17 angulata var. goetzeana (Steph.) Demaret & Vanden Berghen 1950 Basionym: Frullania goetzeana Steph. 1901 [non Steph. 1911] Distribution: Tanganyika, Nyassaland, Congo, Reunion comb nov - syn nov Demaret & Vanden Berghen Bonner Vanden Berghen Wigginton 1950 1965 1976 2002 Wigginton Wigginton 2004 2004 = Frullania angulata 60 655 185 1012 Bull Jard Bot Etat Bruxelles 20: 9-10 Index Hepat 5: 231 Bull Jard Bot Nat Belg 46 (1/2): 71 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Mitt. 1863 angulata var. laciniata Vanden Berghen 1976 Basionym: Distribution: Madogascar var. nov. Vanden Berghen Grolle 1976 1995 185 250 Bull Jard Bot Nat Belg 46 (1/2): 72-73 Bryophyt Bibl 48: 62 = Frullania angulata var. laciniata Demaret & Vanden Berghen 1950 angulistipa Steph. 1910 [orth var & nom superfl] Typus: Samoa Basionym: Distribution: Stephani Verdoorn Verdoorn Bonner Grolle & Schultze-Motel Miller, Whittier & Whittier Stephani = Frullania angustistipa 1910 1937 1937 1965 1972 1983 1985 686 556 556 655 537 609 928 Spec Hepat 4: 438 [spec nov] Blumea suppl 1: 212 [Samoa] Blumea suppl 1: 212 Index Hepat 5: 231 J Hatt Bot Lab 36: 83 [Samoa] Prod Flora Hepat Polynesiae: 120 [nom superfl] Stephani Icon Hepat Inedit [Galeiloba] C-102, microfiche 51 fig 3487 ["Samoa, leg Rechinger 2346"] Flora Vitiensis: 471 [Samoa] Bot Jahrb 23: 314 [Samoa] Spec Hepat 4: 583 Rev Bryol NS 1: 116, fig 6-7 Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 232 J Hatt Bot Lab 36: 83 [Samoa] J Hatt Bot Lab 47: 187, 191, fig 15 [syn sub Fr. intermedia] Prod Flora Hepat Polynesiae: 120 Stephani Icon Hepat Inedit [Homotropantha] D-15, microfiche 54 fig 3724 [" Viti - Ovalau (leg Graeffe)"] - 3725 J Hatt Bot Lab 60: 221 [New Caledonia; syn sub Fr. intermedia var non-apiculata] Steph. 1908 angulosa Mitt. in Seemann 1871 Typus: Samoa sd, Powell Basionym: Distribution: Samoa spec nov Mitten in Seemann Stephani in Reinecke Stephani Verdoorn Verdoorn Bonner Grolle & Schultze-Motel Hattori Miller, Whittier & Whittier Stephani 1871 1896 1911 1928 1930 1965 1972 1980 1983 1985 637 1044 686 242 551 655 537 490 609 928 - syn nov Hattori 1986 506 = Frullania intermedia var. non-apiculata S.Hatt. 1975 Page 18 angustata Mitt. 1861 Typus: Sri Lanka. Peradenia, "on trees" 1.3.1848, Gardner 1434 in hb NY 1048608 (hb W. MITTEN) [lectotypus by Hattori 1975] Basionym: Distribution: Sri Lanka spec nov - comb nov Mitten Sawada 1861 1909 Paris Stephani Iishiba Verdoorn Clark Hattori 1910 1911 1912 1930 1955 1975 Miller, Whittier & Whittier 1983 574 930 J Linn Soc Bot 5: 122 [Sri Lanka] Bot Mag Tokyo 23 (no 268): 178-183, fig 29 [key to Japanese species, in Japanese] 1455 Rev Bryol 37 (6): 130 [New Caledonia: Panié] 686 Spec Hepat 4: 656 888 Bot Mag Tokyo 26: 35 [Japan, with key in Japanese] 293 Ann Bryol Suppl 1: 97 [syn sub Fr. apiculata] 369 Bryologist 58: 68 [syn sub Fr. apiculata] 47 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 74 [comb nov & lectotypification = Fr polyptera var angustata] 609 Prod Flora Hepat Polynesiae: 120 [New Caledonia, Sri Lanka] = Frullania polyptera var. angustata (Mitt.) S.Hatt. 1975 angustifolia Steph. 1910 Typus: Tanzania: Usambara, Lutindi Hochwald -.7.1893, C. Holst 3471 b in hb G (ex hb B) [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Pearson Arnell Bonner Vanden Berghen Stephani 1910 1924 1956 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania trinervis angustistipula Typus: Mexico 686 795 847 655 185 928 Spec Hepat 4: 323-324 Arkiv Botanik 19 (5): 9 [Mt. Elgon in Kenia/Uganda] Svensk Bot Tidskr 50 (4): 527 [Fernando Po] Index Hepat 5: 232 Bull Jard Bot Nat Belg 46 (1/2): 199 Stephani Icon Hepat Inedit [Chonanthelia] A- 10, microfiche 46 fig 3079 {from type; "Usambara Holst 33471b Juli 1893] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 227, fig 149(Q) from type [syn sub Fr trinervis, West Africa] (Lehm.) Drège 1843 Steph. ex Bonner 1965 [nom herb] {Mexico} 1913, Arsén 7851 in hb G Basionym: Distribution: Mexico Bonner Stephani 1965 1985 655 928 Fulford & Sharp 1990 105 Index Hepat 5: 232 Stephani Icon Hepat Inedit [Chonanthelia] A- 58, microfiche 47 fig 3153 [from type; "Mexico Arsén 7851"] Mem NY Bot Gard 63: 40 [Mexico] = Page 19 aongstroemii A.Evans 1900 [nom inval, Art 32.1(c)] Typus: Basionym: Distribution: Hawaii (Oahu) Evans So & Wang 1900 2006 = Frullania angstroemii A.Evans ex Steph. 1910 Frullania sandvicensis aperta 236 Trans Connet Acad Arts Sci 10 (1): 395-397, pl. 44 (1-11) 1466 J H B L 100: 427 [nom inval, Art 32.1c] Ångstr. 1872 Gottsche ex Verd. 1930 [pro syn] Typus: Sri Lanka: Poondeloya 4000-8000 ft 1868, John Nietner Drawings com. by Jack.: hb no 1552 G. Ms. c. per. - 2 Bl. Basionym: Distribution: - syn nov Gottsche Verdoorn Bonner Miller, Whittier & Whittier = Frullania neurota aperta 1886 1930 1965 1983 2390 293 655 609 Bot. Centralblatt 26 (7): 204 [Sri lanka] Ann Bryol Suppl 1: 66 Index Hepat 5: 233 [nom herb] Prod Flora Hepat Polynesiae: 121 Taylor 1846 Gottsche 1886 [nom nud] Typus: Sri Lanka: Poondeloya 4000-8000 ft 1868, John Nietner Drawings com. by Jack.: hb no 1552 G. Ms. c. per. - 2 Bl. Basionym: Distribution: Gottsche Gottsche = Frullania aperta 0 1886 Gottsche ex Verd. 1930 [pro syn] Frullania neurota apiahyna Icon inedt 2390 Bot. Centralblatt 26 (7): 204 [Sri lanka] Taylor 1846 Gottsche ex Steph. 1916 [nom nud] Typus: Bolivia. Cordillera de Santa Cruz Basionym: Distribution: Gottsche Stephani Yano = Frullania apiahyna 0 1916 1999 Icon inedit 222 Biblioth Bot 87: 241 [sine descript, nom nud] [Bolivia] 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] Gottsche ex Steph. 1924 Frullania glomerata Mont 1842] (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Page 20 apiahyna Gottsche ex Steph. 1924 Typus: Brazil: Apiahy sd, Lorentz in hb G 1606 (ex hb GOTTSCHE) Basionym: Distribution: Bolivia, Brazil spec nov -syn nov Stephani Gottsche ex Stephani Schiffner & Arnell 1916 1924 1964 Bonner Yano Stephani 1965 1984 1985 Gradstein, Meneses & Arbe Gradstein & Costa 2003 2003 Yano 2008 = Frullania glomerata Mont 1842] apiculata 222 687 198 Biblioth Bot 87: 241[sine descript, nom nud] [Bolivia] Spec Hepat 6: 535 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 91, pl. 13(fig. 150 j-k) [Brazil] [? forma of Fr. julacea] 655 Index Hepat 5: 234 501 J Hatt Bot Lab 56: 503 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Galeiloba] C- 49, microfiche 50 fig 3404 [from type; "Brasilia, lg Lorentz"] 975 J Hatt Bot Lab 91: 52 [Bolivia; as syn sub Fr glomerata (L & L) Mont] 984 Mem New York Bot Gard 87: 87 [Brazil] [syn sub Fr glomerata (L & L) Mont] vide 975 1684 Boletim Instituto Botanica 19: 89 [syn sub Fr glomerata; Brazil; catalog] (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec [auct. non (R., B. & N.) Nees] Verd. 1930 Basionym: Distribution: Verdoorn Verdoorn 1930 1932 293 553 Verdoorn 1934 241 Verdoorn 1937 556 = Frullania moniliata ssp. obscura Ann Bryol Suppl 1: 100 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 59 [Sumatra, looks like serratae] Ann Bryol Suppl 4: 204-205 [Sumatra, B.N.Borneo: Mt Kinabalu, Sarawak, Neu Guinea, Fidschi] Blumea suppl 1: 212 [South India, Fiji Isl, Samoa] Verd. 1930 {nördl. Indomalaya} Frullania tamarisci ssp. obscura (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata apiculata (Reinw., Blume & Nees) Mont. 1842 [auct. non (R., B. & N.) Nees] Sim 1926 {South Africa} Basionym: Distribution: - syn nov Sim 1926 252 Vanden Berghen 1976 185 = Frullania trinervis apiculata Trans Roy Soc S Africa 15: 43, fig [South Africa] = diptera / apiculata is mentioned at p. 45 Bull Jard Bot Nat Belg 46 (1/2): 107 - there my be an error; SIM 1926, p. 43 there is a picture of diptera not of apiculata, (Lehm.) Drège 1843 [auct. non (R., B. & N.) Nees] Tixier 1973 {Vanuatu (Aneityum)} Basionym: Distribution: Vanuatu (Aneityum) Tixier 1973 678 Bull Mus Nat Hist Nat Paris, ser 3, 190, Bot 10: 82 [New Hebrides (Anotom) = Vanuatu( Aneityum)] = Page 21 apiculata Nees 1838 [nom inval, Art. 32.1(c)] Basionym: Distribution: Nees 1838 = Frullania apiculata apiculata Typus: Africa 261 Naturg Europ Leberm 3: 239 [Anmerkung 1, Fr. a.] (Reinw., Blume & Nees) Nees 1845 [auct. non (R., B. & N.) Nees] Pearson 1887 okt. 1883 in hb KIAR No. 40 Basionym: Distribution: Pearson Vanden Berghen Wigginton = Frullania serrata apiculata 1887 1976 2004 872 Kristiana Vidensk Selsk Forhandl 9: 4, pl 1(2-13) [Africa] 185 Bull Jard Bot Nat Belg 46 (1/2): 111 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Gottsche 1845 [auct. non (R., B. & N.) Nees] Verd. 1932 {Borneo} Typus: Anambas Isl; Boengoeram; Gunung Ranai, 1000m, an Calophyllum -.4.1928 van Steenis Basionym: Distribution: Verdoorn 1932 554 Nederl Kruidk Arch Jrg 1932 (2): 492 [Anambas Isl, Borneo] = apiculata fo. aculeata Verd. 1930 Typus: Papua New Guinea: Morobe, Sattelberg -.6.1899, Nyman in hb FH Basionym: Distribution: New Guinea spec nov - syn nov Verdoorn Verdoorn Clark Bonner Grolle & Schultze-Motel Grolle & Piippo = Frullania apiculata apiculata 1930 1930 1955 1965 1972 1984 293 549 369 655 537 143 Ann Bryol Suppl 1: 101 Nova Guinea 14 (4): 543 Bryologist 58: 68 [syn sub Fr. apiculata] Index Hepat 5: 234 J Hatt Bot Lab 36: 83 [Samoa] Acta Bot Fennica 125: 17 [Cat. West Melanesia] (Reinw., Blume & Nees) Nees 1845 fo. warburgii Verd. 1930 Typus: Philippines: Mindanao sd, Warburg Basionym: Distribution: Indonesia (Java), Philippines, New Guinea fo nov - syn nov Verdoorn Verdoorn Clark Hattori Grolle & Piippo = Frullania apiculata 1930 1930 1955 1975 1984 293 549 369 66 143 Ann Bryol Suppl 1: 101 Nova Guinea 14 (4): 542-543 Bryologist 58: 68 [syn sub F. apiculata] Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 146-147, fig. 161 Acta Bot Fennica 125: 17 [Cat. West Melanesia] (Reinw., Blume & Nees) Nees 1845 Page 22 apiculata var. explicata (Mont.) Schiffn. 1910 Basionym: Distribution: var nov - syn nov Schiffner Clark = Frullania apiculata 1910 1955 436 369 Österr Bot Zeitschr 60 (11): 431-432 Bryologist 58: 68 [syn sub Fr. apiculata] (Reinw., Blume & Nees) Nees 1845 apiculata var. laxa Nees 1845 Basionym: Frullania apiculata var. nodulosa (Nees) Nees 1845 Distribution: Indonesia (Java) var nov - syn nov Nees in G., L. & N. Sande Lacoste Sande Lacoste in Junghuhn Schiffner 1854 1857 1857 1898 141 430 1490 831 Bonner 1965 655 Syn Hepat: 452 Verh Konikl Akad Wetenschappen 5: 89 Plantae Junghuhnianae, Hepaticae: 541 Conspect Hepat Archip Indici, Batavia: 320 [as syn sub Fr. apiculata var nodulosa] Index Hepat 5: 235 = Frullania apiculata var. nodulosa (Nees) Nees 1845 Frullania apiculata var. laxa Nees 1845 Frullania apiculata var. nodulosa (Nees) Nees 1845 Frullania apiculata var. laxa Nees 1845 Frullania apiculata var. nodulosa (Nees) Nees 1845 apiculata var. nodulosa (Nees) Nees 1845 Typus: Indonesia. Java Basionym: Distribution: - syn nov Nees 1830 Nees in G., L. & N. Schiffner Verdoorn Bonner 1845 1898 1930 1965 258 Enum plant cryptogam Javae 1: 50 [No 66 b] [as Jungermannia apiculata var nodulosa] 2850 Syn Hepat: 452 831 Conspect Hepat Archip Indici, Batavia: 320 [as valid var] 293 Ann Bryol Suppl 1: 100 655 Index Hepat 5: 235 = Frullania apiculata var. laxa Nees 1845 Frullania apiculata var. nodulosa (Nees) Nees 1845 Frullania apiculata var. laxa Nees 1845 Frullania apiculata var. nodulosa (Nees) Nees 1845 Frullania apiculata var. laxa Nees 1845 apiculata var. obscura Nees 1845 Basionym: Distribution: Indonesia (Java) spec nov - syn nov Nees in G., L. & N. Sande Lacoste Schiffner Verdoorn Clark Bonner = Frullania apiculata 1845 1857 1898 1930 1955 1965 2850 430 831 293 369 655 Syn Hepat: 452 Verh Konikl Akad Wetenschappen 5: 89 Conspect Hepat Archip Indici, Batavia: 321 Ann Bryol Suppl 1: 100 Bryologist 58: 68 [syn sub Fr. apiculata] Index Hepat 5: 235 (Reinw., Blume & Nees) Nees 1845 Page 23 apiculata var. obtusata S.Hatt. 1978 Typus: Holotype. India, Andaman, Long Islands. 430 m. Aug. 1963. H. C. Verma 1963-1/ 1 in hb NICH 314287 Basionym: Distribution: Andaman Is. var nov - syn nov Hattori Hattori Parihar, Lal & Katiyar Mizutani, Hasegawa & Iwatsuki = Frullania apiculata 1978 1982 1994 2009 127 495 2196 2112 J Jap Bot 53 (5): 129 J Hatt Bot Lab 51: 212 Hepat Anth India, annotated checklist: 21 [Eastern Himalaya] Type sepecimens Hatt Bot Lab (NICH) 20: Holotype. India, Andaman, Long Islands. 430 m. Aug. 1963. H. C. Verma 1963-1/ 1 (NICH 314287). (Reinw., Blume & Nees) Nees 1845 apiculata var. parva S.Hatt. 1984 Typus: Australia. N Queensland, Devil's Thumb, W of Mossman, 1150m 9.10.1982, M.L. Hicks 11287 in hb NICH [holotypus] in hb M.L. HICKS [isotypus] Basionym: Distribution: Australia (QLD) var nov - syn nov Hattori Hattori Hattori Windolf Hattori Konrat & Braggins Carthy McCarthy Mizutani, Hasegawa & Iwatsuki = Frullania apiculata 1984 1986 1987 1987 1988 2001 2003 2006 2009 682 507 402 1025 429 838 1125 1385 2112 Crypt Bryol Lichen 5(1-2): 177-180, fig 1 [Austalia, QLD] J Hatt Bot Lab 60: 242 Mem NY Bot Gard 45: 544-545 [syn sub Fr. apiculata] Austrobaileya 2(4): 390 [checklist Queensland] Nova Hedwigia Beih 90: 147 [Australia: QLD] J Hatt Bot Lab 91: 256-262 [syn sub Fr. apiculata] Flora Australia Suppl Ser 21: 41-42 [sub Fr apiculata] Checklist Australian Liverworts [Australia, syn sub Fr apiculata] Type sepecimens Hatt Bot Lab (NICH) 20: Holotype. Australia, Queensland, Devil's Thujmb. 1150 m. 90et. 1982. M. L. Hicks 11287 (NICH 385065). (Reinw., Blume & Nees) Nees 1845 apiculata ssp. kaindimontana S.Hatt. 1972 Typus: Papua New Guinea. Morobe, Wau Distr. Mt. Kaindi, ca. 2350 m 1967, Schuster 67-5810b in hb NICH [holotypus] in hb JE 04004492 (1967, Schuster,R.M. 67-5810/b1) [isotypus] (Herb. R. Grolle) in hb MASS [isotypus] Basionym: Distribution: New Guinea ssp nov - syn nov Hattori Hattori Hattori Miller, Whittier & Whittier Grolle & Piippo Mizutani, Hasegawa & Iwatsuki = Frullania apiculata 1972 1973 1974 1983 1984 2009 539 542 546 609 143 2112 J Hatt Bot Lab 36: 417-420, fig. 4-5 J Hatt Bot Lab 37: 148 [New Guinea] J Hatt Bot Lab 38: 224-226 Prod Flora Hepat Polynesiae: 121 Acta Bot Fennica 125: 17 [Cat. West Melanesia] Type sepecimens Hatt Bot Lab (NICH) 20: New Guinea, Wau Distr., Summit of Mt. Kaindi. 2350 m. 28 May 1967. R. M. Schuster 675810/b (NICH 309939). (Reinw., Blume & Nees) Nees 1845 Page 24 apiculiloba Steph. 1910 Typus: Indonesia. Java sd, Paterson in hb G (hb STEPHANI 1) Basionym: Distribution: spec nov - syn nov Stephani Merrill Verdoorn Verdoorn Bonner Miller Miller, Whittier & Whittier Stephani = Frullania grandistipula apiculoba 1910 1914 1929 1930 1965 1968 1983 1985 686 1484 245 293 655 672 609 928 Spec Hepat 4: 440-441 Philippine J Sci C, Bot 9(1): 40 [Guam] Ann Bryol 2: 125 [syn sub Fr. grandistipula] Ann Bryol Suppl 1: 43 Index Hepat 5: 235 Micronesia 4 (1): 74 Prod Flora Hepat Polynesiae: 122 Stephani Icon Hepat Inedit [Galeiloba] C-103, microfiche 51 fig 3488 from type; "Java , leg Paterson"] Lindenb. 1845 [non Steph. 1924] Steph. 1996 [nom herb] in hb US {Guam} Typus: Guam. Marianne Isl sd, R.C. McGregor 597 in hb US Basionym: Distribution: NN 1996 632 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 01 of 35 [in hb US] = apollinarii Steph. 1911 Typus: Colombia. Cordillera de Bogota, 2700 m 1905, Frére Apollinaire sn in hb G G00069136 - n° SIB 126170/1 - G20609 (hb LACOUTURE 198) [holotypus] Basionym: Distribution: Colombia, Ecuador spec nov - syn nov Stephani Bonner Gradstein & Hekking Stephani 1911 1965 1979 1985 Uribe & Gradstein Uribe-M Uribe 1998 2004 2008 Uribe 2011 = Frullania lobatohastata appendiculata 686 655 483 928 Spec Hepat 4: 602-603 Index Hepat 5: 236 J Hatt Bot Lab 45: 112 [Cat Colombia] Stephani Icon Hepat Inedit [Meteriopsis] E-13, microfiche 55 fig 3784 ["Bogota, Herb Lacouture 198"] 707 Bryophyt Bibl 53: 35 [Catalogue Colombia] 1174 Caldasia 26: 89-90, fig1(a-h] [sub sgen Thyopsiella] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] 3230 Caldasia 33(2): 365 [syn sub Fr lobato-hastata] Steph. 1911 Steph. ex Bonner 1965 [nom herb] [non Steph. 1897] {PERU} Typus: Peru: Sachapata Lechler sn hb L 194 in hb G sub Fr. sachapatensis Basionym: Distribution: Bonner 1965 655 Index Hepat 5: 236 = Page 25 appendiculata Steph. 1897 [non Steph. ex Bonner 1965] Typus: Japan. Yezo, Horomanhati [holotype] - Japan. Mt Bandayama, Kochi Pref (Prov Tosa) [isotype] [holotype] - Yoshinaga 31[isotype] 21.8.1892, Tokubuchi 271 in hb G [holotypus] in hb TNS [isotypus?] Basionym: Distribution: Russia (Far East), China, Japan spec nov - syn nov Stephani Inoue Yoshinaga Sawada 1897 1900 1906 1909 213 802 927 930 Stephani Iishiba Verdoorn 1911 1912 1930 Horikawa Hattori Horikawa 1934 1944 1951 Hattori & Noguchi Bonner Hattori Stephani 1960 1965 1972 1985 Inoue Piippo Konstantinova, Potemkin & Schijakov Gambaryan N.N. [National Science Museum Tokyo] Yamada & Iwatsuki Konstantinova & Bakalin 1987 1990 1992 Bull Herb Boissier 5: 78 & 88-89 [spec nov; Japan] Bot Mag Tokyo 14 (no 162): 180 Bot Mag Tokyo 20: 52 [Japan] Bot Mag Tokyo 23 (no 268): 178-183, fig 27 [key to Japanese species, in Japanese] 686 Spec Hepat 4: 547 888 Bot Mag Tokyo 26: 35 [Japan, with key in Japanese] 293 Ann Bryol Suppl 1: 81 [syn sub Fr. moniliata ssp. obscura fo. appendiculata] 34 J Sci Hiroshima Univ, ser B, div 2, 2: 235 [syn sub Fr. moniliata] 943 Bull Tokyo Sci Mus 11: 148 [Japan, key to Japanese species] 1698 J Sci Hiroshima Univ, serB, div 2 (Bot) 6(1): 28 [sub Fr moniliata ssp obscura] 965 Index spec Typ herb Japon, Bryophyta: 16 [syntype] 655 Index Hepat 5: 236 536 J Hatt Bot Lab 35: 216 928 Stephani Icon Hepat Inedit [Thyopsiella] F-131, microfiche 59 fig 4061 ["Japan, Tokubuchi 271 (syn. shiyoshiana?)"] - 4062 1155 Index type specimens in Nat Sci Mus, Tokyo: 12 521 J Hatt Bot Lab 68: 38 [annotated catalogue of China] 651 Arctoa 1:101 [USSR] 1998 2000 1222 Chenia 5: 132 [Russia Far East] 995 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 2006 2009 1470 J Hatt Bot Lab 99: 28 [Catalog of Japan] 2113 Arctoa 18: 14 [Checklist of liverworts of Russia: FAR EAST] = Frullania moniliata ssp. obscura fo. appendiculata (Steph.) Verd. 1930 Frullania tamarisci ssp. obscura (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata applanata (Reinw., Blume & Nees) Mont. 1842 Steph. 1911 Typus: New Caledonia "Nova Caledonica Septentr.", Mt. Panié -.2.1910, LeRat in hb G (hb Gén Paris 441) Basionym: Distribution: New Caledonia spec nov Stephani Verdoorn Verdoorn Bonner Stephani 1911 1930 1930 1965 1985 686 293 551 655 928 - syn nov Hattori 1986 506 Spec Hepat 4: 587 Ann Bryol Suppl 1: 161 [syn sub Fr. intermedia] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 236 Stephani Icon Hepat Inedit [Homotropantha] D-16, microfiche 54 fig 3726 ["N Caledonia, Gén Paris 441"] - 3727 J Hatt Bot Lab 60: 221 [syn sub Fr. intermedia var. non-apiculata] = Frullania intermedia var. non-apiculata S.Hatt. 1975 Page 26 areacae (Spreng.) Gottsche by Kattel 2002 [orth var] Basionym: Distribution: orth var Kattel = Frullania arecae 2002 (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura areaceae 1032 Liverworts of Nepal, Katmandu: 16 (Sw.) Mont. 1840 (Spreng.) Gottsche by Nath & Asthana 1998 [orth var] Basionym: Distribution: Nath & Asthana = Frullania arecae 1998 683 J Bryol 20(1): 233 [South India] [orth var] (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura (Sw.) Mont. 1840 Page 27 arecae (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Typus: Demaret & Vanden Berghen (1948) studied type material without mention of its location; Evans (1914) mentioned the presence of an isotype in W; Yuzawa (1991) did not examine the type.[Gradstein 2012] Basionym: Jungermannia arecae Spreng. 1821 Distribution: Bioko (Fernando Po), Burundi, Cameroon, Democratic Republic of Congo, Ethiopia, Ghana, Kenya, Malawi, Mozambique, Nigeria, Rwanda, Sao Tome, Tanzania, Uganda, Zimbabwe, Madagascar, Mauritius, Reunion, South Africa (Natal), Mexico, Guatemala, Costa Rica, Panama, Jamaica, Haiti, Puerto Rico, Martinique, Dominican Rep, Ecuador, Galapagos Isl (Santa Cruz)[Ecuador], Colombia, Peru, Bolivia, French Guiana, Brazil, Chile, Argentina, China (Yunnan), Taiwan, India (North: Assam, Meghalaya (Shillong); South: Tamil Nadu (Nilgiri Hills, Western Ghats)), Sikkim. Bhutan, Nepal, Sri Lanka, Burma, Norfolk Isl Gottsche Gottsche Stephani Schiffner 1863 1867 1888 1895 103 104 936 1015 Schiffner in Engler & Prantl Cavers Stephani Stephani Evans Stephani Herrera Bryan Clark & Svihla Demaret & Vanden Berghen Cufodontis Herzog Clark Herzog Herzog Clark & Svihla Arnell Svihla Arnell Vanden Berghen Arnell Arnell Schiffner & Arnell Bonner Grolle Robinson Crandall Reed & Robinson Vanden Berghen 1905 1907 1910 1911 1914 1916 1926 1929 1947 1948 1952 1952 1953 1955 1955 1956 1956 1956 1959 1961 1962 1963 1964 1965 1966 1967 1969 1971 1972 809 897 686 721 235 222 2788 175 319 59 2562 444 1106 107 107 376 847 380 448 1507 449 822 198 655 843 771 193 1406 249 Augier 1974 821 Vanden Berghen 1976 185 Vanden Berghen Winkler Vanden Berghen Hegewald & Hegewald Gradstein & Hekking Lemos Michel 1976 1976 1976 1977 1979 1980 185 5 850 172 483 719 Onraedt & Cremers Haarbrink Gradstein & Weber Vanden Berghen Jones & Harrington Yuzawa Yano Yuzawa Menzel 1980 1981 1982 1982 1983 1983 1984 1984 1984 75 49 496 76 251 498 501 502 142 Separata of Mex. Leverm.: 236 K Danske Vid Selks Skrifter, ser 5, 6: 332 Hedwigia 27: 62 [Kilimandscharo] Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 132 Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 132 Naturalist 1907 (Jan/Feb): 12, fig 1E [water sac] Spec Hepat 4: 331 Kungl Svenska Vetenskapsakad Handl 45 (9): 88 [Chile: Chiloe Isl] Trans Connet Acad Arts Sci 18: 338 [Peru] Biblioth Bot 87: 238 [Bolivia] Chloris Cuzcoensis: 14 Bot Gazette 88(3): 338 Bryologist 50: 82-85, fig. 1-13 Bull Jard Bot Etat Bruxelles 19 (1): 73-84, fig. 5 (a-d) Phyton 4 (1-3): 76 [Ethiopia] Svensk Bot Tidskr 46 (1): 83 [Ecuador] Proc California Acad Sci 4 ser, 27 (18): 662 [Galapagos Isls: Isabela] Feddes Repert 57 (1/2): 198 Feddes Repert 57 (1/2): 171 [Colombia] Bryologist 59: 36-38, fig. 1-24 Svensk Bot Tidskr 50 (4): 527 [Fernando Po] Bryologist 59: 277-278 Svensk Bot Tidskr 53 (4): 500 [Martinique] Bull Soc Roy Bot Belg 93 (1-2): 70-71 [Kivu, Urundi, Katanga] Svensk Bot Tidskr 56 (2): 341 [Ecuador] Hepat S Africa: 136-138, fig. 100 (a-e) Österr Akad Wiss, Math Nat Kl, Denkschr 111: 84 [Brazil] Index Hepat 5: 236 Khumbu Himal 4: 266 [Nepal] Bryologist 70: 51 [Colombia] Beih Nova Hedwigia 30: 81 Phytologia 21 (1): 19 [Costa Rica: Monteverde] Expl Hydrobiol Bassin Bangweolo et Luapula 8 (1): 66-68, fig 25(a-f) [Zambia] Annales de la Faculté des Sciences du Cameroun 17:82, 83, 86, 91 [Cameroun] Bull Jard Bot Nat Belg 46 (1/2): 154-160, fig. 38 [synonym with Fr. ecklonii] Bull Jard Bot Nat Belg 46 (1/2): 154-160, fig. 38 Rev Bryol Lichen 42 (3): 811 [Colombia] Distributiones plantarum africanarum 11: 363 [map of Africa] Nova Hedwigia 28: 742 [Peru) J Hatt Bot Lab 45: 112 [Cat Colombia] Diss, Uni Fed do Rio Grande do Sul, Porto Alegre: 81-83, pl 12, map 11, [Brasil, Rio Grande do Sul] Crypt Bryol Lichen 1 (3): 274 [French Guiana] Lindbergia 7 (1): 52-54, fig. 5-6 Colombia, high andes] J Hatt Bot Lab 52: 148 [Galapagos Isls: Santa Cruz] Crypt Bryol Lichen 3 (3): 212 [key to Madagascar] & 222-223 Bull Brit Mus nat Hist (Bot) 11 (3): 250 [Sierra Leone, Ghana] J Hatt Bot Lab 54: 109-112, fig. 3 [diagnosis & synonyms] J Hatt Bot Lab 56: 503 [checklist Brasil] J Hatt Bot Lab 57: 391-394, fig. 8 Willdenowia 14: 488-489 [Peru] Page 28 Hattori & Lin Stephani 1985 1985 505 928 Hattori & Piippo Yuzawa, Mues & Hattori Hattori van Zanten & Gradstein Kuo & Chiang Gradstein Gradstein & Hekking Visnadi & Monteiro Best Piippo Fulford & Sharp Long & Grolle Yuzawa Yuzawa Schäfer-Verwimp Morales Z. Boggan, Funk, Kelloff et al. Costa Fischer Wolf 1986 1987 1988 1988 1988 1989 1989 1990 1990 1990 1990 1990 1991 1991 1991 1991 1992 1992 1993 1993 557 509 516 88 948 20 517 1293 879 521 105 522 525 525 523 307 4771 1001 309 2380 O'Shea Wolf 1993 1993 985 2380 Wolf 1993 2380 Fischer Wolf 1993 1993 308 2379 Yuzawa & Koike Parihar, Lal & Katiyar 1994 1994 38 2196 Tixier Yano Chuah-Petiot Grolle Pérez & Sánchez Wigginton & Grolle Chuah-Petiot Boggan, Funk, Kelloff, Hoff, Cremers & Feuillet Nath & Asthana Hodgetts 1995 1995 1995 1995 1996 1996 1997 1997 730 2095 31 250 297 696 646 1396 1998 1998 Uribe & Gradstein Nath & Asthana 1998 1998 Stotler, Allen, Gradstein, McGuinness, Whittemore & Chung Yano Gradstein in Luteyn 1998 683 J Bryol 20(1): 233 [South India] 1014 http://www.nhm.ac.uk/hosted_sites/bbstbg/ugabook.htm [printed 11.12.03, 12 pages]: 6 [Uganda] 707 Bryophyt Bibl 53: 35 [Catalogue Colombia] 708 J Hatt Bot Lab 85: 64-66, fig 1(1-13) as Fr. wallichiana [South India; key to all spec.] 703 Trop Bryol 15: 178 [Checklist Panama] Pocs & Lye Konrat & Braggins Hodgetts, Matcham & Duckett Wu & Wang Uribe & Rangel-Ch Gradstein, Griffin III, Morales & Nadkarni 1999 1999 1999 2000 2000 2000 1999 1999 J Hatt Bot Lab 59: 151 [China] Stephani Icon Hepat Inedit [Chonanthelia] A- 16, microfiche 46 fig 3088 ["Caracas"] Acta Bot Fennica 133: 55-56, fig. 11 [= range of altitude] J Hatt Bot Lab 63: 425-436 [morphology & systematics] J Hatt Bot Lab 65: 414 [PNG] Nova Hedwigia Beih 90: 56 [experimental dispersal geography] Taiwania 33: 11 [Index of Taiwan] Bryologist 92 (3): 338 [Key to spec of Puerto Rico & Virgin Isl] J Hatt Bot Lab 66: 208 [French Guiana] Hoehnea 17(1): 74 [Brazil; Cidade de Rio Claro, Sao Paulo] Kirkia 13 (2): 296 [Zimbabwe] J Hatt Bot Lab 68: 38 [annotated catalogue of China] Mem NY Bot Gard 63: 40 [Mexico] J Hatt Bot Lab 68: 412-413 [Bhutan, Habitat & distribution] J Hatt Bot Lab 70: 261 [difference of arecae & ecklonii] J Hatt Bot Lab 70: 264-269, fig. 42-44, map 1 a(:280) [= var. arecae] J Hatt Bot Lab 69: 153 [Brazil] Trop Bryol 4: 36 [Costa Rica] Checklist of the plants of the Guianas Acta Botanica Brasilica 6(1): 16-17, fig 6(a-f) [Brazil] Trop Bryol 8: 96 [key to species in eastern Zaire & Rwanda] Phytocoenologia 22(1):41-44 [Colombia, Epiphyte - Lower montane rain forest communities: Frullania arecae - Frullanoides densifoliaSquamidium nigricans community] J Bryol 17: 648 [Malawi] Phytocoenologia 22(1): 40-41 [Colombia, Epiphyte - Lower montane rain forest communities: Diplasiolejeunea pauckertii-Brachiolejeune laxifolia community] Phytocoenologia 22(1): 23 [Colombia, Epiphyte - Lower montane rain forest communities: Frullania arecae - Frullanoides densifoliaSquamidium nigricans community] Trop Bryol 8: 44 [Checklist: Zaire, Rwanda] Phytocoenologia 22(1): 64 [Colombia, Epiphyte - Upper montane rain forest communities: accidental with preference for the Lower Mountain rain forest] J Hatt Bot Lab 75: 193 [Nepal] Hepat Anth India, annotated checklist: 21 [Eastern Himalaya, Western Himalaya, South India] Trop Bryol 11: 18 [ephiphyllous in Zaire & Rwanda] J Hatt Bot Lab 78: 142 [annotated checklist of Brazil] Trop Bryol 10: 51 [Kenya: Mount Kenya] Bryophyt Bibl 48: 63-64 [afr3] Trop Bryol 12: 47 [Bioco] Bryophyt Bibl 50: 83 [Catalogue ... of Sub-Saharan Africa] Trop Bryol 13: 62 [Kenya, Aberdare Range] Checklist Guianas: 10 [French Guiana] 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] 994 Mem New York Bot Gard 84: 58 [Paramo 800-3800m: Colombia, Ecuador] 724 Trop Bryol 17: 30 [Uganda] 725 Trop Bryol 17: 106 [epiphyllous] 754 J Bryol 21: 142 [Natal] 2671 Bryoflora of Hengduan MTS (Southwest China): 129, fig. 57(1-6) 1090 Colombia, diversidad biótica III, Bogota: 444 [Colombia] 1405 Caldasia 23(1): 208 [Costa Rica: Monteverde] Page 29 Bai in Gao & Cao Oliviera-e-Silva & Yano Bapna & Kachroo Hodgetts (Gulf of Guinea conservation group) Bai Wigginton Singh in Nath & Astana Chuah-Petiot Merwin, Gradstein & Nadkarni Wigginton Bytebier & Chuah-Petiot Holz, Gradstein & Heinrichs Wigginton 2000 2000 2000 2000 1019 989 712 784 2001 2001 2001 2001 2001 2001 2002 2002 2002 1742 982 1374 1074 1075 1079 1076 1404 1012 Bai 2002 Oliveira-e-Silva, Milanez & Yano 2002 Kattel 2002 1189 1077 1032 Drehwald 2003 2813 Gradstein & Costa Srivastana & Alam Drehwald 2003 2003 2003 984 1324 2813 Chuah-Petiot 2003 1124 Drehwald 2003 2813 Nöske, Gradstein, Kürschner, Parolly & Torracchi Chuah-Petiot & Pocs Holz 2003 1084 2003 2003 Wigginton Manyanga & Perold Visnadi Holz 2004 2004 2004 2004 Parolly & Kürschner 2004 Hodgetts 2004 Camara & Vital Wigginton 2004 2004 Yano & Camara 2004 Costa, Imbassahy & Silva Dauphin 2005 2005 Camara, Oliveira & Santiago Kürschner & Parolly Ah-Peng & Bardat Holz & Gradstein Costa, Alencar Imbassahy & Silva Camara & Costa Tusiime, Byarujali & Bates Schäfer-Verwimp & Pócs 2005 2005 2005 2005 2005 1082 Acta Bot Hungarica 45 (1-2): 57 [Aberdare Mts., Kenya] 1185 Diversity Ecology Montane Quercus Forest, Costa Rica [Dissertation]: 46, 76 [upper montane oak forest, in mats] 1172 Sript Bot Belg 30: 224-226, fig 146(A-P) [West Africa] 1301 SABONET Report 21: 4 [Zimbabwe] 1158 Acta bot bras 18(1): 93 [Sao Paulo, Brazil] 1184 [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm 1217 Nova Hedwigia 79(3-4): 393, 398, 403, 415, 416, tab 2, 3, 4 & 7 [Podocarpus NP, Ecuador, epiphytic] 1254 http://tring.naturalhistory.museum/hosted_sites/bbstbg/ugaheps.htm [printed 24.08.2004 13 pages]: 6 [Uganda] 1367 Acta bot bras 18(4): 884 [Brazil, Panatanal de Mato Grosso] 1312 Trop Bryol Res Rep 5: 29 [Bioko, Burundi, Cameroun, Ethiopia, Ghana, Kenya, Madagascar, Malawi, Mozambique, Nigeria, Reunion, Rwanda, South Africa, Sao Tome, sie, Tanzania, Uganda, DR Congo, Zimbabwe] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1292 Acta Amazonica 34(4): 455 [Brazil: Manaus, Rio Clara, Belem, Salvador] 1448 Rodriguesia 56 (87): 38 [State of Rio de Janeira] 1360 Tropical Bryology 26: 141-218 [Catalogue of Costa Rican Hepaticae and Anthocerotae] 1359 Trop Bryology 26: 137 [Brazil; Distrito Federal Brasilia] 1362 Nova Hedwigia 80 (1-2): 94 [Ecuador; Podocarpus NP] 1358 Tropical Bryology 26: 108 [Checklist of Reunion Isl] 1408 J Biogeogr 32: 1595 [Costa Rica: Cordillera de Talamanca] 1382 J Hatt Bot Lab 98: 283 [Rio de Janeiro State, Brazil] Gradstein 2009 2006 2007 2009 Fl Yunnanica 17: 423, fig 212(1-8) [China, Yunnan] Boletim Inst Botanica 13: 22-23 [Rio de Janeiro, Brazil] Hepaticology in India II: 184-185 http://www.ggcg.st/botany/liverworts1.htm [Guinea Islands: Bioko, São Tomé] Guizhou Science 19(4): 47 [China, Yunnan] http://www.nhm.ac.uk/hosted-Sites/bbstbg/malmoss.htm [Malawi] Perspectives Indian Bryology: 342 [India & Pantropical] Trop Bryol 20: 36 [Cherangani Hills, Kenya] Trop Bryol 20: 66 [Montverde, Costa Rica] Trop Bryol 20: 94 [foliicolous taxa of Malawi] Trop Bryol 22: 62 [Taita Hills, Kenya] Bryologist 105(3): 336-337 [Costa Rica] Trop Bryol Res Rep 2: 27 [Bioko, Burundi, Cameroun, Ethiopia, Ghana, Kenya, Madagascar, Malawi, Mozambique, Nigeria, Reunion, Rwanda, South Africa, Sao Tome, sie, Tanzania, Uganda, DR Congo, Zimbabwe] [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Chenia 7: 20 [Yunnan, China] Trop Bryol 22: 80 & 92 [Brazil] Liverworts of Nepal, Katmandu: 16 [Nepal, 2000-3000 m; as "areacae" & "arelae"] Cambios en la vegetación briofítica: 292 [Peru (Bosque Protector Alto Mayo)] Mem New York Bot Gard 87: 83 [Brazil] Geophytology 33: 47 [India] Cambios en la vegetación briofítica: 296 [Bolivia (Parque Nacional Amboro)] Mosses, liverworts & hornworts of Kenya: 204, fig 262(a-f) [Kenya; key to species] Cambios en la vegetación briofítica: 283 [Ecuador (Parque Nacional Sangay)] Crypt Bryol 24 (1): 25 [Reserva Biologica San Francisco, Ecuador] 1131 Hoehnea 33(1): 82 [Reserva Ecologica do IGBE, Brazil] 2014 Afr J Ecol, suppl 3: 82 [Uganda, 1700m] 1985 Acta Bot Hungarica 51(3-4): 383 [new to Dominican Republic, Hispaniola] 2161 Bibliotheca Lichenologica 99: 148-149 [Galapagos Isl, Note: a specimen with toothed lobules from Santa Cruz was described by YUZAWA (1991) as F. areceae var. spiniloba (Steph.) Yuzawa.] Page 30 - syn nov Gao & Wu Bordin & Yano Gradstein & Uribe 2010 2010 2011 Wang, Lai & Zhu Söderström, Hagborg, Vana & Konrat Gradstein & Ziemmeck Yano 2011 2011 Gradstein 2012 Schäfer-Verwimp et alii Long & Rubasinghe Daniels & Kariyappa 2013 2014 2019 = Frullania obscura arecae 2012 2012 2349 Genera Hepat. et Anth. Sinicorum: fig. 244(1-8) [key in Chinese] 2266 Pesquisas, sér Bot 61: 47 [Brazil] 3230 Caldasia 33(2): 355, fig 1(f-i) [Colombia; synopsis of Frullaniacae & keys] 3125 Annales Botanici Fennici 48: <369-395> [Checklist Taiwan] 2581 Bryologist 114(1) : 72 [checklist Jamaica] 4845 Checklist of Galapagos Horn- Liverworts [Galapagos] 3678 Pesquisas, série Botânica 63:55-140 [Catálogo ... do Estado do Espírito Santo, Brasil] 3341 Cryptogamie, Bryologie 33 (3 ): 257-262 [Frullania arecae (Spreng.) Gottsche is replaced by the older name Frullania obscura (Sw.) Dumort] 3799 Phytotaxa 128 81): 20 [Ecuador] 3994 Ceylon Journal of Science 43(1): 4 [Sri Lanka] 5109 Bryoflora of Agasthyamalai Biosphere Reserve: 392-394, fig 141 (a-p) [India (Western Ghats, keys] (Sw.) Mont. 1840 [auct. non (Spreng.) Gottsche 1863] Spruce 1884 Basionym: Distribution: - syn nov Spruce Clark & Svihla Demaret & Vanden Berghen Clark & Svihla 1884 1947 1948 1948 Clark & Svihla Arnell Gao & Lai Yano Bordin & Yano 1956 1963 2003 2008 2010 = Frullania ecklonii arecae 243 319 59 325 Trans Proc Bot Soc Edinburgh 15: 12 & 20-21 Bryologist 50: 82-85, fig 1-13 [pp, syn with SPRUCE 1884] Bull Jard Bot Etat Bruxelles 19 (1): 82 [syn nov sub Fr. ecklonii] Bryologist 51: 23 [key to Fr. of Middle America: Mexico, Guatemala, Costa Rica, Jamaica, Haiti, Puerto Rico, Venezuela, Brazil, Ecuador, Peru, Bolivia (Chile, Cape of Good Hope)] 376 Bryologist 59: 36-38, fig 1-24 822 Hepat S Africa: 135 1441 Illustrations Bryophytes China: 1127, fig 528(1-8) [China] 1684 Boletim Instituto Botanica 19: 29-30 [Brazil; catalog] 2266 Pesquisas, sér Bot 61: 47 [Brazil] (Spreng.) Gottsche, Lindenb. & Nees 1845 (Spreng.) Clark & Svihla 1947 [pp] [ pro syn] [ non Spruce 1884] Basionym: Distribution: Clark & Svihla Demaret & Vanden Berghen 1947 1948 319 59 Arnell 1963 822 = Frullania arecae Bryologist 50: 82-85 Bull Jard Bot Etat Bruxelles 19 (1): 77 & 82 [pp syn non Spruce, syn sub Fr. arecae] Hepat S Africa: 136 [syn sub Fr. arecae] (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura (Sw.) Mont. 1840 Page 31 arecae var. spiniloba (Steph.) Yuzawa 1991 Basionym: Frullania spiniloba Steph. 1910 Distribution: Mexico, Panama, Jamaica, Colombia, Ecuador, Bolivia comb nov Yuzawa Uribe M & Gradstein Söderström, Hagborg, Vana & Konrat Hentschel at alii 1991 1998 2011 525 J Hatt Bot Lab 70: 269-271, fig. 45, map 4c(:283) 707 Bryophyt Bibl 53: 35 [Catalogue Colombia] 2581 Bryologist 114(1) : 72 [checklist Jamaica] 2015 4097 Phytotaxa 220 (2): 136 [=Fr obscura var. spiniloba] = Frullania obscura var. spiniloba (Steph.) Hentschel & von Konrat 2015 arelae Mitt. by Kattel 2002 [orth var] Basionym: Distribution: orth err Kattel 2002 = Frullania wallichiana Mitt. 1861 Frullania obscura argentina 1032 Liverworts of Nepal, Katmandu: 16 [Nepal, 1100-3000 m] (Sw.) Mont. 1840 Gottsche ex Bonner 1965 [nom herb] Typus: Argentinia: Tacuman sd, Lorentz 904 in hb G Basionym: Distribution: Argentina Stephani 1910 686 Bonner 1965 655 = Frullania conferta Spec Hepat 4: 346 [syn sub Fr. conferta; according BONNER, not found by STEPHANI] Index Hepat 5: 237 Spruce 1889 Frullania tetraptera Nees & Mont. 1838 Page 32 arietina Taylor 1845 Typus: Surinam:? /?Colipa 8000 ft ?, Weigelt sn /? Liebmann in hb GLM (hb LEHMANN) 05947, 05948 (ex hb GOTTSCHE) Basionym: Distribution: USA ( FL ), Mexico, Honduras, Dominica, Barbados, Guadeloupe, Martinique, Colombia, Peru, Galapagos Isls [Ecuador], Brazil, Chile (Chiloe Isl), Hawaii spec nov - syn nov - syn nov Taylor in G., L. & N. Taylor in G., L. & N. Lindenberg & Gottsche Gottsche Gottsche Austin Spruce Stephani Spruce Stephani Stephani Evans Duss Stephani Stephani Evans in Britton & Millspaugh Fulford Pagan Pagán Frye & Clark Herzog Schiffner & Arnell Bonner Lemos Michel 1845 1847 1851 1863 1867 1874 1884 1888 1895 1897 1897 1897 1904 1910 1911 1920 1939 1939 1942 1947 1955 1964 1965 1980 Yano Menzel Stephani 1984 1984 1985 Fulford Fulford & Sharp Yuzawa Schuster 1987 1990 1991 1992 NN NN Schäfer-Verwimp 2002 2002 2010 = Frullania rio-janeirensis arietina 2850 2848 282 103 104 668 243 938 238 1059 214 237 3132 686 721 1094 660 931 3152 204 107 198 655 719 Syn Hepat: 413 (No 5) Syn Hepat: 771 Linnaea 24 (5): 637 [Suriname] Separata of Mex. Leverm.: 235 K Danske Vid Selks Skrifter, ser 5, 6: 311 [Mexico] Bull Torrey Bot Club 5: 15 [Oahu - Sandwich Isl] Trans Proc Bot Soc Edinburgh 15: 14 & 25-26 (No 14) Hedwigia 27: 280 [Puerto Rico] J Linn Soc Bot 30: 333 [Dominica] Bihang Kgl Sv Vetensk-Akad Handl bot 23 III (2): 18 [Brazil] Bull Herb Boissier 5: 842[:260] [Hawaii] Trans Connet Acad Arts Sci 10 (1): 5-6 (No 1), pl. 1 (fig. 1-6) Flore Cryptogamique des Antilles Françaises: 132 [Martinique] Spec Hepat 4: 327 Kungl Svenska Vetenskapsakad Handl 45 (9): 88 [Chile: Chiloe Isl] Bahama Flora: 519 [Bahamas] Ann Bryol 11: 56 [British Honduras] Bryologist 42: 79 [Puerto Rico] Bryologist 45(4): 107-108 [Guadeloupe] Univ Wash Publ Biol 6 (5): 778 [syn sub Fr. riojaneirensis] Feddes Repert 57 (1/2): 171 [Colombia] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 84 [Brazil] Index Hepat 5: 237 Diss, Uni Fed do Rio Grande do Sul, Porto Alegre: 92-95, pl 15, map 14, [Brasil, Rio Grande do Sul] 501 J Hatt Bot Lab 56: 563 [checklist Brazil] 142 Willdenowia 14: 509 [syn of Fr. riojaneirensis] 928 Stephani Icon Hepat Inedit [Chonanthelia] A- 19, microfiche 46 fig 3091 ["Caripe Moritz"] 99 Mem NY Bot Gard 45: 272 [Dominica] 105 Mem NY Bot Gard 63: 40 [Mexico] 525 J Hatt Bot Lab 70: 252-257 [syn sub Fr. riojaneirensis] 603 xxxx: 254-261 [syn sub Frull riojaneirensis, scarcely separable phenotype: arietina & riojaneirensis, with key] 1266 http://www.natureserve.org/explorer [USA] 1266 http://www.natureserve.org/explorer [USA; Canada] 2460 Crypt Bryol 31(4): 346, 347 [syn sub Frull riojaneirensis, Dominica, West Indies] (Raddi) Ångstr. 1876 [auct. non Taylor 1845] Austin 1874 {Hawaii} Typus: Sandwich Isl. Basionym: Distribution: auct - syn nov Austin Baldwin Evans Miller Miller, Whittier & Whittier = Frullania neurota 1874 1877 1900 1963 1983 668 901 236 834 609 Bull Torrey Bot Club 5: 15 [with Dendroceros clintoni - Sandwich Isl] Hawaiian Almanac: 42 [Hawaii] Trans Connet Acad Arts Sci 10 (1): 399 [syn sub Fr.sandvicensis] Ark Bot, ser 2, 5: 518 [syn nov sub Fr. neurota] Prod Flora Hepat Polynesiae: 122 Taylor 1846 Page 33 arietina fo. laxior Lindenb. & Gottsche 1851 Typus: Suriname. ?, Kegel 747 Basionym: Distribution: Surinam fo nov Lindenberg & Gottsche = Frullania rio-janeirensis 1851 282 Linnaea 24 (5): 637 [Suriname] (Raddi) Ångstr. 1876 arietina var. pluricarinata Spruce 1884 Typus: S America: San Carlos, Flumen Negro sd, Spruce sn ? Basionym: Distribution: ?Brasil (Rio Negro / San Carlos) var nov Spruce Bonner Yuzawa 1884 1965 1991 = Frullania rio-janeirensis armatistipula 243 655 525 Trans Proc Bot Soc Edinburgh 15: 26 Index Hepat 5: 237 J Hatt Bot Lab 70: 252 (syn sub Fr. riojaneirensis] (Raddi) Ångstr. 1876 Steph. 1910 Typus: Tanzania [DeutschOst Africa]: Tanga 1905, Ramlow 18a in hb G 17327 (ex hb B) [holotypus] Basionym: Distribution: East Africa spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1910 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania ericoides armitana 686 655 185 928 Spec Hepat 4: 378-379 Index Hepat 5: 237 Bull Jard Bot Nat Belg 46 (1/2): 163 Stephani Icon Hepat Inedit [Galeiloba] C- 5, microfiche 50 fig 33523553 [from type; "Ostafrica, Tanga, No 18a Ramlow legit 1905"] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 231 [syn sub Fr ericoides, West Africa] (Nees) Mont. 1839 Steph. by S.Hatt. 1980 [orth var] Basionym: Distribution: Hattori Hattori = Frullania armitiana 1980 1982 489 495 J Hatt Bot Lab 47: 107-109 J Hatt Bot Lab 51: 213-214 [as Fr. armitiana] Steph. 1911 armitana var. longe-attenuata (S.Hatt.) S.Hatt. 1980 [orth var] Basionym: Distribution: Hattori Hattori 1980 1982 489 495 J Hatt Bot Lab 47: 107-109 J Hatt Bot Lab 51: 213-214 [as Fr. armitiana var. longe-attenuata] = Frullania armitiana var. longe-attenuata (S.Hatt.) S.Hatt. 1980 Page 34 arnellii Steph. ex Herzog 1916 Typus: Bolivia, Cerro Tunari 4700m 1911, T.Herzog 4930a in hb G 9664 [holotypus] Basionym: Distribution: Bolivia, Colombia spec nov Stephani Stephani Bonner Gradstein & Hekking Stephani 1916 1924 1965 1979 1985 222 687 655 483 928 - syn nov Yuzawa Gradstein, Meneses & Arbe 1991 2003 525 975 = Frullania albertii Steph. ex Herzog 1916 Frullania standaertii arsenii Biblioth Bot 87: 241, fig 189 [Bolivia] Spec Hepat 6: 535 [as spec nov] Index Hepat 5: 238 J Hatt Bot Lab 45: 112 [Cat Colombia] Stephani Icon Hepat Inedit [Chonanthelia] A- 59, microfiche 47 fig 3154 [from type; "Bolivia Herzog 4930a"] J Hatt Bot Lab 70: 205 [syn sub Fr. albertii] J Hatt Bot Lab 91: 52 [Bolivia; as syn sub Fr. albertii] Steph. 1910 Steph. 1924 Typus: Mexico 1913, Arsén 7794 in hb G G00069137 - n° SIB 125177/1 - G20951A (autoica c per, Arsén 7794) [holotypus] in hb G G00128068 - n° SIB 143811/7 (monoica, Arsén 7832) in hb G G00128069 - n° SIB 143812/1 (Arsén 7756) Basionym: Distribution: Mexico, Colombia, Bolivia spec nov syn Stephani Bonner Stephani 1924 1965 1985 687 655 928 Fulford & Sharp Yuzawa 1990 1991 105 525 Uribe & Gradstein Gradstein 1998 2020 707 = Frullania standaertii Spec Hepat 6: 530 Index Hepat 5: 238 Stephani Icon Hepat Inedit [Chonanthelia] A- 60, microfiche 47 fig 3155 [from type; "Mexico Arsén 7794"] Mem NY Bot Gard 63: 40 [Mexico] J Hatt Bot Lab 70: 207-208, fig. 14, map 5a(:284) [key to arsenii & neurota] Bryophyt Bibl 53: 35 [Catalogue Colombia] syn sub Fr standaertii Steph. 1910 asagrayana var. alsophila Howe 1894 Typus: USA. California, Marin Co., Mill Valley & Olemna Basionym: Distribution: var nov - syn nov Howe Evans Bonner = Frullania californica 1894 1897 1965 570 237 655 Erythea 2: 99 Trans Connet Acad Arts Sci 10 (1): 25-26 Index Hepat 5: 233 (M.Howe) A.Evans 1897 Page 35 asagrayana var. californica pp A (Austin) Howe 1894 Basionym: Frullania grayana var. californica pp A Austin ex Underwood 1884 Distribution: - syn nov Pearson Howe 1890 1894 Evans Stephani Frye & Clark Bonner 1897 1911 1947 1965 = Frullania franciscana 1153 List of Canadian Hepaticae: 3 570 Erythea 2: 98-99 [Fr. asagrayana pp = Fr. franciscana HOWE, pp = Fr. californica (AUST) EVANS] 237 Trans Connet Acad Arts Sci 10 (1): 25-28 686 Spec Hepat 4: 570 [as syn sub Fr. californica] 204 Univ Wash Publ Biol 6 (5): 745 655 Index Hepat 5: 238 M. Howe 1894 asagrayana var. californica pp B (Austin) Howe 1894 Basionym: Frullania grayana var. californica pp B Austin ex Underwood 1844 Distribution: nom nov stat nov Pearson Howe Evans Macoun Bonner = Frullania californica asperifolia 1890 1894 1897 1902 1965 1153 570 237 1150 655 List of Canadian Hepaticae: 3 Erythea 2: 98-99 [ppA = Fr. franciscana, ppB = Fr. california]] Trans Connet Acad Arts Sci 10 (1): 25-28 Catalogue Canadian Plants, VII: 45 [sub Fr californica] Index Hepat 5: 238 (M.Howe) A.Evans 1897 Steph. in Steph. & Watts 1914 Typus: Australia. New South Wales, Mt. Wilson, The Avenue, 3500ft 11.1.1991, W. W. Watts 1093 [Hep Aust ex hb WATTS 109] in hb G 18193 in hb NSW Basionym: Distribution: Australia (NSW) spec nov - syn nov Stephani in Stephani & Watts Bonner Hattori Stephani 1914 1965 1979 1985 Scott & Bradshaw Brown, Ramsay & Seur Carthy McCarthy 1986 1992 2003 2006 = Frullania falciloba 220 655 485 928 J Proc Roy Soc New S Wales 48: 105-106 [Australia] Index Hepat 5: 239 J Hatt Bot Lab 45: 323-325, fig 1(a-j) from type Stephani Icon Hepat Inedit [Galeiloba] C-194, microfiche 53 fig 3599 ["Australia N.S.W., Watts 1093"] 56 Brunnonia 8 (1): 33 [Australia] 1140 Telopea 5(1): 74 [types in hb NSW] 1125 Flora Australia Suppl Ser 21: 43 [sub Fr falciloba] 1385 Checklist Australian Liverworts [Australia, syn sub Fr falciloba] Lehm. 1844 Page 36 asperula Mitt. 1861 Typus: India. Khasi Mts, 5000-6000 ft, "on Meteorium" sd, J.D. Hooker & T. Thomson 1440 in hb NY 1048609 (hb W. MITTEN, with drawings) Basionym: Distribution: India (North: Eastern Himalayas: Megalaya), Sri Lanka, Indonesia (Java) spec nov - syn nov spec nov Mitten Stephani Chopra Chopra Parihar Bonner Hattori Hattori 1861 1910 1938 1943 1962 1965 1973 1975 Stephani 1985 Parihar, Lal & Katiyar 1994 Bapna & Kachroo Singh 2000 2002 Singh, Kumar & Nath 2008 = Frullania acutiloba 574 686 933 205 1584 655 541 47 J Linn Soc Bot 5: 119 [India: Khasi] Spec Hepat 4: 445 Proceed Indian Acad Sci, sect B, 7 (5): 248 [South India] J Indian Bot Soc 22: 249 [India] Univ Allahabad studies, Bot sect 1-2: 18 [India: EH IO] Index Hepat 5: 239 J Hatt Bot Lab 37: 98-99, fig. 53 [from type] Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 75 [syn nov sub Fr. hampeana var acutiloba] 928 Stephani Icon Hepat Inedit [Galeiloba] C-104, microfiche 51 fig 3489 ["india orient, Herb München"] 2196 Hepat Anth India, annotated checklist: 21 [Eastern Himalaya, South India] 712 Hepaticology in India II: 165 [syn sub Fr acutiloba] 4886 Studies on Liverworts (Bryophyta) of Khasi and Jaintia Hills: Meghalaya. 284, map 11 - Unpublished thesis. 1281 Taiwania 53 (1): 79, fig 1[map], [India, Meghalaya, Eastern Himalayas] Mitt. 1861 [non Gerola 1947] Frullania monocera var. acutiloba (Mitt.) Hentschel & von Konrat 2015 atchafalagae Hampe ex Hooker & Taylor 1845 [nom nud] {N America} Basionym: Distribution: Hampe in Hooker & Taylor 1845 290 London J Bot 4: 89 [nomen only; species from North America] = aterrima var. rostrata (R.M.Schust.) S.Hatt. 1983 [nom inval (Art 37.1)] Basionym: Neohattoria rostrata Distribution: New Zealand comb nov -syn nov R.M.Schust. 1970 [nom inval (Art 37.1)] Schuster 1970 761 Hattori Glenny Fife & allii Setzepfand 1983 1998 1998 2002 499 1412 2424 1159 Glenny & Malcolm 2005 1410 Konrat, Hentschel, Heinrichs, Braggins & Pócs 2010 2377 J Hatt Bot Lab 33: 281-285, fig 2(1-11), 3 (1-6) [as Neohattoria rostrata] J Hatt Bot Lab 54: 135 [comb nov] Tuhinga 10: 132 [revised checklist of New Zealand] Australasian bryol newsletter 39: 14 [Australia] Epiphytische Vegetation auf Weinmannia racemosa, Neuseeland [Diss]: 117-118 [New Zealand, endemic] [syn sub Fr aterrima lepida, according VON KONRAT] ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand species] Nova Hedwigia 91(3-4): 482 [nom inval (Art 37.1)] = Frullania aterrima var. lepida E.A.Hodgs. 1949 Page 37 atrata [auct. non (Sw.) Nees] Spruce 1884 {Ecuador, Mt. Tunguragua & Mt. Guayrapata} Typus: in hb MANCH 12782, 12775 Basionym: Distribution: auct - syn nov Spruce Stotler in Seaward & FitzGerald Uribe & Gradstein Uribe = Frullania peruviana atrata 1884 1997 2003 2008 243 1092 1071 1513 Trans Proc Bot Soc Edinburgh 15: 38-39 Richard Spruce, Roy Bot Gard Kew: 133, 135, fig 15-21 Crypt Bryol 24 (3): 199 Caldasia 30(1): 75 [syn sub Fr peruviana] [Monography of subgenus Meteoriopsis] Gottsche 1846 [non Gottsche ex Spruce 1884] Nees ex Mont. 1840 {French Guiana} Typus: in hb PC-MONT. LEPRIEUR 270 Basionym: Distribution: - syn nov Montagne Husnot Gradstein & Hekking = Frullania apiculata 1840 1875 1989 467 Ann Sci Nat Bot, ser 2, 14: 333 2435 Rev Bryol 2(1): 4 [Antilles francaises] 517 J Hatt Bot Lab 66: 208 (Reinw., Blume & Nees) Nees 1845 atrata var. fo. forma [auct. non (Sw.) Nees ex Mont.] Spruce 1884 {Ecuador, Quito} Typus: Ecuador. Quito 1847, Jameson 56 Basionym: Distribution: Uribe & Gradstein = Frullania convoluta 2003 1071 Crypt Bryol 24 (3): 199 [auct sub Fr convoluta] Lindenb. & Hampe 1851 atrata var. martiana Nees 1845 Typus: in hb S (B22579, B22580) [possible typus] Basionym: Distribution: Nees in G., L. & N. Bonner Myrdal = Frullania atrata 1845 1965 2002 2850 Syn Hepat: 464 655 Index Hepat 5: 240 1276 Mosskollekter [Frullania types in hb S] (Sw.) Nees ex Mont. 1839 Page 38 atrata var. subtilissima Nees ex Mont. 1840 Typus: Guiana. River Kau -.5.1838, Leprieur "254" (264, corr. Montagne 1855: 184) in hb PC-MONT [holotype] Basionym: Distribution: Montagne Lindenberg in G., L. & N. Montagne Stephani Bonner Gradstein & Hekking Schäfer-Verwimp 1840 1845 1855 1911 1965 1989 2010 Söderström, Hagborg, Vana & Konrat 2011 = Frullania subtilissima 467 2850 475 686 655 517 2460 Ann Sci Nat Bot, ser 2, 14: 333 Syn Hepat: 443 [pro syn sub Fr. subtilissima] Ann Sci Nat Bot, ser 4, 3: 320 [Leprieur 264, not 254] Spec Hepat 4: 631 [not mentioned as syn of Fr. subtilissima] Index Hepat 5: 240 J Hatt Bot Lab 66: 209 [is nom nud, syn sub Fr. subtilissima] Crypt Bryol 31(4): 348 [Basionym Frull atrata var subtilissima, Dominica, West Indies] 2581 Bryologist 114(1) : 72 [checklist Jamaica] (Nees ex Mont.) Lindenb. 1845 atrata var. swartzii Nees 1845 Typus: Jameica. "in montibus Jamaicae australis" sd, Swartz in hb S (B22587, B22588) [possible typus] Basionym: Jungermannia atrata Sw. 1788 Distribution: Nees in G., L. & N. Bescherelle 1845 1893 Bonner Myrdal 1965 2002 = Frullania atrata 2850 Syn Hepat: 464 3815 J Bot Morot 7 (9): 178 [Guadeloupe (Guadeloupe : Bory (Herb. Montagne); L'Herminier; Husnot (Exsiccata n" 221)., Martinique (: Husnot (n° 221).] 655 Index Hepat 5: 241 1276 Mosskollekter [Frullania types in hb S] (Sw.) Nees ex Mont. 1839 atriculata Steph. [nom. herb.] Typus: Samoa L. Rechinger, 1948 [isotype, Identified by S. Hattori 1979] https://www.gbif.org/occurrence/1090985776 Field Museum Basionym: Distribution: = Page 39 atrosanguinea Taylor ex Spruce 1884 Typus: Ecuador. Mt. Pichincha bei Quito; Mt. Tunguragua and Mt. Guayrapata Pichincha) Spruce (Mt. Tunguragua and Mt. Guayrapata) >> see fr0792.doc Jameson (Mt. in hb MANCH MANCH12798 (Mt. Tunguragua, Spruce sn) [lectotypus] in hb FH (Mt. Tunguragua, Spruce sn) [isolectotypus] in hb MANCH MANCH12799 (Mt. Guayrapata, Spruce sn) [paralectotype] in hb E E00002516 (Woods of Esmeralda, Jameson) [isotype, syntype] Basionym: Distribution: Costa Rica, Venezuela, Bolivia, Ecuador, Colombia spec nov - syn nov Taylor ex Spruce Jack & Stephani Stephani Herzog Clark & Svihla Clark Herzog Bonner Robinson Gradstein & Hekking Menzel 1884 1892 1911 1942 1951 1953 1955 1965 1967 1979 1984 243 1089 686 874 340 1106 107 655 771 483 142 Stephani 1985 928 Judd Kron Morales Z. Uribe & Gradstein Uribe & Rangel-Ch Holz 1987 1988 1991 1998 2000 2004 1681 510 307 707 1090 1184 Uribe 2008 1513 = Frullania peruviana atrosanguinea Trans Proc Bot Soc Edinburgh 15: 39-40 Hedwigia 31: 13 [Peru, Neu-Granada] Spec Hepat 4: 600-601 Beih Bot Centralbl 61 B: 577 [Bolivia] Bryologist 54: 72-74, fig 1-18 Proc California Acad Sci 4 ser, 27 (18): 662 [Galapagos Isls: Santiago] Feddes Repert 57 (1/2): 171 Index Hepat 5: 241 Bryologist 70: 51 [Colombia] J Hatt Bot Lab 45: 112 [Cat Colombia] Willdenowia 14: 506 [not in Peru, but in Ecuador - error by Stephani 1911: 601] Stephani Icon Hepat Inedit [Meteriopsis] E-15, microfiche 55 fig 3787 ["Frontino, Nova Granada, G Wallis 1874"] Bull Florida State Mus, Biol Sci 32(1): ?? [Florida, USA] J Hatt Bot Lab 64: 355, fig 6(a-c) [key] Trop Bryol 4: 37 [Costa Rica] Bryophyt Bibl 53: 35 [Catalogue Colombia] Colombia, diversidad biótica III, Bogota: 444 [Colombia] [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm Caldasia 30(1): 75 [syn sub Fr peruviana] [Monography of subgenus Meteoriopsis] Gottsche 1846 [non Gottsche ex Spruce 1884] fo. rotundifolia (Steph.) Herzog 1955 [nom nud] Basionym: Distribution: Colombia Herzog Uribe & Gradstein = Frullania atrosanguinea Frullania peruviana 1955 1998 107 707 Feddes Repert 57 (1/2): 171 [Colombia] Bryophyt Bibl 53: 35 [is Fr. atrosanguinea] Taylor ex Spruce 1884 Gottsche 1846 [non Gottsche ex Spruce 1884] attenuata var. longe-attenuata (S.Hatt.) S.Hatt. 1975 Basionym: Frullania longe-attenuata Distribution: comb nov Hattori Hattori Hattori Grolle & Piippo S.Hatt. 1973 1975 1980 1982 1984 547 489 495 143 J Hatt Bot Lab 39: 280 [comb nov; basionym is Fr. longe-attenuata] J Hatt Bot Lab 47: 107-108 [syn sub Fr. armitiana var. longe-attenuata] J Hatt Bot Lab 51: 214-215 [status as in 1975] Acta Bot Fennica 125: 18 [Cat. West Melanesia] = Frullania armitiana var. longe-attenuata (S.Hatt.) S.Hatt. 1980 Page 40 auritistipula Steph. 1911 Typus: Ecuador. Gualaquiza sd, Allioni in hb G (hb LEVIER 6452) Basionym: Distribution: Ecuador spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 = Frullania montagnei austiniana 686 655 4761 194 928 Spec Hepat 4: 528-529 Index Hepat 5: 241 Nova Hedwigia 18: 452-453 The genus Frullania, separata: 452-453 Stephani Icon Hepat Inedit [Thyopsiella] F-101, microfiche 58 fig 4024 ["Ecuador, Gualaquiza, Allioni legit, Levier 6452"] Gottsche 1845 Pearson ex Steph. 1985 [pro syn] Basionym: Distribution: Stephani = Frullania deflexa australis 1985 928 Stephani Icon Hepat Inedit [Homotropantha] D-19b, microfiche 54 fig 3733 ["syn Frull Austiniana Pearson ms, Viti, leg Seemann, (Frullania pendula Austin = nomen occupatum) (ex Herbario Austin)"] Mitt. 1862 Steph. 1911 Typus: Australia. New South Wales, East Ballina, tree 14.8.1900, W.W. Watts 117 ?? Sn ?? 2638 ?? in hb G 18194 (ex hb LEVIER 2638) in hb NSW Basionym: Distribution: Australia (NSW, QLD) spec nov - syn nov Stephani Bonner Hattori Hattori Hattori Stephani 1911 1965 1979 1983 1984 1985 686 655 486 499 682 928 Windolf Scott & Bradshaw Hattori Windolf Brown, Ramsay & Seur Konrat & Braggins Carthy Glenny, Meagher & Brown 1985 1986 1987 1987 1992 2001 2003 2004 1049 56 402 1025 1140 838 1125 1127 McCarthy 2006 1385 = Frullania hypoleuca Spec Hepat 4: 659-660 Index Hepat 5: 241 J Hatt Bot Lab 46: 122-123 J Hatt Bot Lab 54: 135-137, fig 51 Crypt Bryol Lichen 5(1-2): 180 [Austalia, QLD] Stephani Icon Hepat Inedit [Diastaloba] B-88, microfiche 49 fig 3336 [from type; "N S Wales, W W Watts 2638" & "Name besetzt Frull Wattsiana St nsp"] J Hatt Bot Lab 59: 174 [Sunshine Coast, Queensland] Brunnonia 8 (1): 33 & 142 [Australia: NSW] Mem NY Bot Gard 45: 545, fig 1 [Australia] [syn sub Fr. hypoleuca] Austrobaileya 2(4): 390 [checklist Queensland] Telopea 5(1): 74 [types in hb NSW] J Hatt Bot Lab 91: 250-256 [syn sub Fr. hypoleuca] Flora Australia Suppl Ser 21: 44 [sub Fr hypoleuca] Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Checklist Australian Liverworts [Australia, syn sub Fr hypoleuca] Nees 1843 Page 41 autoica Steph. ex Bonner 1965 [nom herb] {Trinidad} Typus: Trinidad. Marvael 20.11.1897, Crüger in hb G Basionym: Distribution: Trinidad Bonner Stephani 1965 1985 655 928 Index Hepat 5: 241-242 Stephani Icon Hepat Inedit [Chonanthelia] A- 18, microfiche 46 fig 3090 [from type; "Trinidad Crüger legit 20.11.1847 Herb Jack"] = azorica Corbiere ex Bisang, Schumacker, Sergio & Grolle 1988 [pro syn] Typus: in hb AZ Basionym: Distribution: Bisang, Schumacker, Sérgio & Grolle = Frullania cesatiana 276 Giorn Bot Ital 122: 265 [syn sub Fr. cesatiana] De Not. 1865 Frullania riparia bakeri 1989 Hampe 1838 Steph. ex Herzog 1916 Typus: Bolivia, Comarapa, 2600m, Nebelwald -.4.1911, Herzog 4266 in hb G 13588 [holotypus] in hb S (B22589) [typus] Basionym: Distribution: Bolivia, Ecuador spec nov Stephani ex Herzog Stephani Herzog Bonner Stotler Stotler Stephani 1916 1924 1942 1965 1969 1970 1985 - syn nov Uribe 2008 = Frullania phalangiflora baladica 222 687 874 655 4761 194 928 Biblioth Bot 87: 243, fig 194 [sub sgen Frullania] [Bolivia] Spec Hepat 6: 544-545 [as spec nov] Beih Bot Centralbl 61 B: 577 [Ecuador] Index Hepat 5: 242 Nova Hedwigia 18: 477 The genus Frullania, separata: 477 Stephani Icon Hepat Inedit [Meteriopsis] E-16, microfiche 55 fig 3788 ["Bolivia, Herzog 4266"] 1513 Caldasia 30(1): 81 [syn sub Fr phalangiflora] [Monography of subgenus Meteoriopsis] Steph. 1916 Gottsche by Paris 1908 [orth var] Basionym: Distribution: Paris Miller, Whittier & Whittier = Frullania baladina 1908 1983 864 609 Rev Bryol 35: 62 [New Caledonia] Prod Flora Hepat Polynesiae: 122 Gottsche 1894 Page 42 balansae Steph. 1894 Typus: China. Tonkin, Foret du. Bavi 9.6.1888, leg Balansa in hb G 14397 (ex hb PC) Basionym: Distribution: China, Taiwan spec nov Stephani Stephani Verdoorn Verdoorn Inoue Bonner Hattori Lai & Wang-Yang Stephani 1894 1911 1930 1934 1961 1965 1972 1976 1985 228 686 293 240 964 655 536 33 928 Kuo & Chiang 1988 948 Piippo 1990 521 Hedwigia 33: 156 Spec Hepat 4: 551-552 Ann Bryol Suppl 1: 60 Ann Bryol Suppl 4: 195-196 J Jap Bot 36(5): 187 [Formosa] Index Hepat 5: 242 J Hatt Bot Lab 35: 220 Taiwania 21 (2): 168 [Index Taiwan] Stephani Icon Hepat Inedit [Thyopsiella] F-162, microfiche 59 fig 4102 ["Tonkin, Foret du Bavi (Bovi?) Balansa 9/6. 88"] Taiwania 33: 11 [Index of Taiwan] [syn sub Fr. tamarisci ssp. obscura var. balanse] J Hatt Bot Lab 68: 38 [annotated catalogue of China] = Frullania tamarisci ssp. obscura var. balansae (Steph.) S.Hatt. 1972 Frullania moniliata balanse (Reinw., Blume & Nees) Mont. 1842 Steph. by Bapna & Kachroo 2000 [orth var] Basionym: Distribution: Bapna & Kachroo = Frullania balansae 2000 712 Hepaticology in India II: 196 Steph. 1894 Frullania tamarisci ssp. obscura var. balansae (Steph.) S.Hatt. 1972 Frullania moniliata ballinensis (Reinw., Blume & Nees) Mont. 1842 Steph. 1911 Typus: Australia. New South Wales, Heath, East Ballina -.3.1901, W.W. Watts 239 in hb G 18196 (hb LEVIER 3033) Basionym: Distribution: spec nov - syn nov Stephani Bonner Hattori Hattori Stephani 1911 1965 1979 1984 1985 Scott & Bradshaw Carthy McCarthy 1986 2003 2006 = Frullania serrata 686 655 486 682 928 Spec Hepat 4: 559-560 Index Hepat 5: 242-243 J Hatt Bot Lab 46: 124 [syn sub Fr. serrata] Crypt Bryol Lichen 5(1-2): 187 [Austalia, QLD] Stephani Icon Hepat Inedit [Thyopsiella] F-181, microfiche 59 fig 4131 ["N S Wales,, East Ballina, Watts 239, Levier 3033"] & microfiche 60 fig 4132 56 Brunnonia 8 (1): 33 [Australia] 1125 Flora Australia Suppl Ser 21: 47 [sub Fr serrata] 1385 Checklist Australian Liverworts [Australia, syn sub Fr serrata] Gottsche 1845 Page 43 bambusana Steph. 1924 Typus: Papua New Guinea. Madang: "Schraderberg, Gebirgswald", 2070m 5.6.1913, Ledermann 12034 in hb G 3288 [holotypus] Basionym: Distribution: New Guinea spec nov - syn nov Stephani Verdoorn Bonner Hattori 1924 1930 1965 1978 687 549 655 482 Grolle & Piippo Stephani 1984 1985 143 928 Stephani 1985 928 = Frullania curvistipula bangiensis Spec Hepat 6: 552 Nova Guinea 14 (4): 541 & 543, pl 92 fig 4 Index Hepat 5: 243 J Hatt Bot Lab 44: 529-530 [syn nov, nom nud, St Icon ined no 163, non 48] Acta Bot Fennica 125: 18 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Thyopsiella] F-163, microfiche 59 fig 4103 ["Nova Guinea (Ledermann legit)"] Stephani Icon Hepat Inedit [Diastaloba] B-48, microfiche 49 fig 3288 ["12034. 159. (Diastaloba) Nova Guinea (Ledermann legit)"] Steph. 1911 Steph. 1910 Typus: Africa Centralis, Bangui -.8.1902, A. Chevalier 71 in hb G (ex hb CORBIERE 75) [holotypus] in hb PC [isotypus] Basionym: Distribution: Central Africa spec nov - syn nov - syn nov Stephani Stephani in Corbiere Stephani Demaret Demaret & Vanden Berghen Bonner Vanden Berghen Stephani 1910 1912 1916 1942 1947 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania squarrosa Spec Hepat 4: 372-373 Bull Mus Nat Hist Nat Paris 18 (2): 121 Biblioth Bot 87: 241 [Bolivia] Bull Jard Bot Etat Bruxelles 16 (2): 304 Bull Jard Bot Etat Bruxelles 18: 238 [syn sub Fr squarrosa] Index Hepat 5: 243 Bull Jard Bot Nat Belg 46 (1/2): 163 [syn sub Fr. ericoides] Stephani Icon Hepat Inedit [Galeiloba] C- 6ab, microfiche 50 fig 3354 ["Congo, Bongosamona, Dupuis leg, Cardot 95"] - 3355 - 3356 [from type; " Africa centrlis, Bangi, Chevalier 1902, Herb Corbiere 75"] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 231 [syn sub Fr ericoides, West Africa] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides banksiana 686 2097 222 1066 1101 655 185 928 (Nees) Mont. 1839 Colenso 1889 Typus: New Zealand. County of Waipawa, Dannevirke 1888, W. Colenso Basionym: Distribution: New Zealand spec nov Colenso Stephani Bonner Hamlin Hodgson Hamlin Glenny Glenny & Malcolm = Frullania pycnantha 1889 1892 1965 1972 1972 1973 1998 2005 636 210 655 659 3468 656 1412 1410 Trans Proc New Zealand Inst 21: 75 J Linn Soc Bot 29: 274 [syn sub Fr. "pyncnantha", orth error] Index Hepat 5: 243 Rec Dominion Mus 7(19): 261 [New Zealand] J Roy Soc New Zealand 2 (1): 110 [new synonyms] Rec Dominian Mus 8 (10): 143 [New Zealand; a valid species] Tuhinga 10: 132 [sub Fr pycnantha] ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand species] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Page 44 baoleyana Steph. ex Myrdal 2000 [from textfile, Mossor artlist] Basionym: Distribution: Myrdal = Frullania baileyana baptistae 2000 1265 http://www2.nrmse/kbo/saml/mossor/mossnamn.txt [hb S] Steph. 1910 Gottsche 1845 Typus: Brazil. Prov. Minas Geres, Joao Baptista sd, Martius in hb NEES Basionym: Distribution: spec nov - syn nov - syn nov Gottsche in G., L. & N. Stephani Bonner Stotler Stotler Stephani = Frullania intumescens barbeyana 1845 1911 1965 1969 1970 1985 2850 686 655 4761 194 928 Syn Hepat: 455-456 Spec Hepat 4: 491 Index Hepat 5: 243 Nova Hedwigia 18: 444 [syn sub Fr. intumescens] The genus Frullania, separata: 444 [syn sub Fr. intumescens] Stephani Icon Hepat Inedit [Thyopsiella] F- 31, microfiche 57 fig 3913 ["Brasilia, Minas Geraes"] - 3914 ["ex planta boliviana, leg Herzog, Levier 6086"] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. ex Herzog 1916 Typus: Bolivia, Comarapa, 2600 m 1911, Herzog 4197 in hb G [holotypus] Basionym: Distribution: Bolivia spec nov Stephani Stephani Bonner Stephani 1916 1924 1965 1985 222 687 655 928 - syn nov Yuzawa & Koike Gradstein, Meneses & Arbe 1989 2003 519 975 = Frullania beauverdii bartlettii Biblioth Bot 87: 241, fig 190 [Bolivia] Spec Hepat 6: 536 [as spec nov] Index Hepat 5: 244 Stephani Icon Hepat Inedit [Chonanthelia] A- 20, microfiche 46 fig 3092-3093 [from type; "Bolivia Herzog 4197"] J Hatt Bot Lab 66: 351-352, fig. 7 [from type; syn sub Fr beauverdii] J Hatt Bot Lab 91: 52 [Bolivia; as syn sub Fr beauverdii; vide Yusawa 1991?] Steph. 1916 von Konrat & Braggins in von Konrat & Pickering 2003 [nom nud] Typus: New Zealand (North Isl: Northland, Radar Bush) Basionym: Distribution: New Zealand (North Isl: Northland, Radar Bush) Konrat & Pickering 2003 Konrat & Brennan 2004 1000 http://www.discoverlife.org/nh/tx/Plantae/Bryophyta/ [IDnature guide for Liverworts, Frullania, New Zealand] 1164 http://www.liverworts.org/dich_key/frul_key.shtml, 1st of 6 pages [Key to Frullania of New Zealand; Northland; Radar Busch] = Page 45 baumanii S.Hatt. by Miller, Whittier & Whittier 1983 [orth var] Basionym: Distribution: Miller, Whittier & Whittier = Frullania baumannii 1983 609 Prod Flora Hepat Polynesiae: 122 S.Hatt. 1977 baumanii var. elongata S.Hatt. by Miller, Whittier & Whittier 1983 [orth var] Basionym: Distribution: Miller, Whittier & Whittier 1983 609 Prod Flora Hepat Polynesiae: 122 = Frullania baumannii var. elongata S.Hatt. 1977 Frullania baumannii S.Hatt. 1977 baumannii var. elongata S.Hatt. 1977 Typus: New Caledonia, Mt. Mone. 1060 m. 28 July 1951. H. Hurlimann 2794 in hb NICH 425459 Basionym: Distribution: New Caledonia var nov Hattori Miller, Whittier & Whittier Hattori Mizutani, Hasegawa & Iwatsuki - syn nov = Frullania baumannii bella 1977 1983 1984 2009 479 609 503 2112 J Hatt Bot Lab 43: 411-413, fig 3(f-i) [New Caledonia] Prod Flora Hepat Polynesiae: 122 [as "baumanii var. elongata] J Hatt Bot Lab 57: 407 Type sepecimens Hatt Bot Lab (NICH) 21: Holotype. New Caledonia, Mt. Mone. 1060 m. 28 July 1951. H. Hurlimann 2794 (NICH 425459). S.Hatt. 1977 Steph. ex Paris 1910 [nom nud] Basionym: Distribution: nom nud Paris Paris 1910 1910 = Frullania bella benguetensis 911 Rev Bryol 37 (1): 41 [New Caledonia: Isle of Pines] 1455 Rev Bryol 37 (6): 130 [New Caledonia: Panié] Steph. 1911 Steph. 1911 Typus: Philippines.Luzon, Prov Benguet, Panai -.5.1909, E.D. Merrill 6640 in hb G 17187 Basionym: Distribution: Philippines spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Hattori Stephani = Frullania claviloba 1911 1929 1930 1965 1975 1985 686 245 293 655 66 928 Spec Hepat 4: 652 Ann Bryol 2: 143 Ann Bryol Suppl 1: 121 Index Hepat 5: 244 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 154 Stephani Icon Hepat Inedit [Diastaloba] B-59, microfiche 49 fig 3299 [from type; "Luzon, Merrill 6640"] Steph. 1911 Page 46 berggrenii Nicholson 1925 [non Steph. 1916] [hom illeg] Typus: New Zealand. South Isl, Otira Gorge 1874, S. Berggren 3346 in hb LD [Hodgson 1949] Basionym: Distribution: in hb NY [Hattori 1979] spec nov - nom nov /or/ Nicholson Hodgson Bonner Hamlin 1925 1949 1965 1972 = Frullania nicholsonii berggrenii 867 203 655 659 Bryologist 28: 17-19, fig 1-9 Trans Roy Soc New Zealand 77 (3): 368 [nom nov = Fr. nicholsonii] Index Hepat 5: 245 Rec Dominion Mus 7(19): 261 [New Zealand] E.A.Hodgs. 1949 Steph. ex Herzog 1916 [non Nicholson 1925] Typus: Bolivia: Incacorral, 2200 m 1911, Herzog 4957 in hb G 20951b [holotypus] Basionym: Distribution: Bolivia spec nov Stephani Stephani Bonner Yuzawa Stephani 1916 1924 1965 1983 1985 222 687 655 498 928 - syn nov Yuzawa Gradstein, Meneses & Arbe 1991 2003 525 975 Biblioth Bot 87: 238, fig. 186 [Bolivia] Spec Hepat 6: 537 [as spec nov] Index Hepat 5: 245 J Hatt Bot Lab 54: 117-117, fig 7(a-l) [from type] Stephani Icon Hepat Inedit [Chonanthelia] A- 22ab, microfiche 46 fig 3096-3099 [from type, Bolivia , Herzog 4957"] J Hatt Bot Lab 70: 269 [syn sub Fr. arecae var. spiniloba] J Hatt Bot Lab 91: 52 [Bolivia; as yn sub Fr. arecae var. spiniloba] = Frullania arecae var. spiniloba (Steph.) Yuzawa 1991 Frullania obscura var. spiniloba (Steph.) Hentschel & von Konrat 2015 bergmanii var. latiloba S.Hatt. 1974 Typus: New Guinea. PNG, Morobe, Mt. Kaindi 1700m 1968, van Zanten 68-326/c in hb JE [holotypus] in hb NICH [isotypus] Basionym: Distribution: New Guinea var nov - syn nov Hattori Hattori Grolle & Piippo = Frullania bergmanii berkeleyana Typus: Nicobars 1974 1975 1984 546 547 143 J Hatt Bot Lab 38: 229-232, fig 111 J Hatt Bot Lab 39: 280 Acta Bot Fennica 125: 18 [Cat. West Melanesia] S.Hatt. 1974 Gottsche ex Bonner 1965 [nom herb] sd, E. Berkeley 1298 oder 1300 in hb G sub Fr. recurvata (ex hb GOTTSCHE) in G (14398), G (14399); Basionym: Distribution: Niccobar Isl [India] nom herb Gottsche Bonner Hattori = Frullania recurvata 0 1965 1972 655 538 Icon inedit Index Hepat 5: 245 J Hatt Bot Lab 36; 124-127, fig 10(a-d) [syn sub Fr gaudichaudii] Lehm. 1838 Frullania gaudichaudii (Nees & Mont.) Nees & Mont. 1845 Page 47 berteroana Gottsche ex Lehm. & Lindenb. 1845 [pro syn] Typus: Chile: Quillota sd, Bertero Basionym: Distribution: Lehmann & Lindenberg in G., L. 1845 & N. 2850 Syn Hepat: 428 (No 36-) = Frullania trinervis var. berteroana Lehm. & Lindenb. 1845 Frullania trinervis bescherellei (Lehm.) Drège 1843 [auct. non Steph. 1894] in Pearson 1922 {Tahiti} Typus: Tahiti. Basionym: Distribution: Pearson Hattori 1922 1986 = Frullania nadeaudii bescherellei 120 506 J Linn Soc Bot 46 (No 305): 31-32 [Tahiti] J Hatt Bot Lab 60: 218 [specimens from Tahiti may be Fr. nadeaudii] Hedwigia 33: 140-141 Rev Bryol 35: 62 [New Caledonia] Rev Bryol 37 (6): 130 [New Caledonia: Ouaoueoue] Spec Hepat 4: 456-457 J Linn Soc Bot 46 (No 305): 31-32 [specimen from Tahiti = Fr. nadeaudii] Nederl Kruidk Arch Jrg 1930 (2): 158 [New Caledonia] Index Hepat 5: 246 J Hatt Bot Lab 49: 363-365, fig 3(a-l) from type Prod Flora Hepat Polynesiae: 122 Stephani Icon Hepat Inedit [Galeiloba] C-108, microfiche 51 fig 3493 ["Nouvelle Caledonia, Bourail 5/3 69, leg Balansa 918; syn Fr jacquinotii"] J Hatt Bot Lab 60: 217-218 [syn sub Fr. pusilla] Steph. 1911 Steph. 1894 Typus: New Caledonia. Bourail sd, Balansa 918 in hb PC (hb BESCHERELLE) in hb G 18442 Basionym: Distribution: New Caledonia spec nov - syn nov Stephani Paris Paris Stephani Pearson 1894 1908 1910 1911 1922 228 864 1455 686 120 Verdoorn Bonner Hattori Miller, Whittier & Whittier Stephani 1930 1965 1981 1983 1985 551 655 492 609 928 Hattori 1986 506 = Frullania pusilla Mitt. 1871 bescherellei var. explanata Pearson 1922 Typus: Tahiti. Port Ngea sd, Compton 26, 175, 194 Basionym: Distribution: var nov - syn nov Pearson Verdoorn Bonner Hattori Miller, Whittier & Whittier Hattori = Frullania pusilla 1922 1930 1965 1981 1983 1986 120 551 655 492 609 506 J Linn Soc Bot 46 (No 305): 32 [Tahiti] Nederl Kruidk Arch Jrg 1930 (2): 158 [only a modification] Index Hepat 5: 246 J Hatt Bot Lab 49: 365 Prod Flora Hepat Polynesiae: 122 J Hatt Bot Lab 60: 218 [syn sub Fr. pusilla] Mitt. 1871 Page 48 bessonii Steph. in Renauls & Cardot 1893 Typus: Madagaskar sd, Sikora no 56 in hb G (hb RENAULD 310) Basionym: Distribution: Madagascar spec nov Stephani in Renauld & Cardot Stephani Bonner 1893 1911 1965 Vanden Berghen Stephani 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania loricata beyrichiana 935 686 655 Bull Soc Roy Bot Belg 32 (2,I): 31-32 [Madagaskar] Spec Hepat 4: 488 [not mentioned by Fr. loricata] Index Hepat 5: 246 [error: not a syn sub Fr. loricata by STEPH 1910: 488] 185 Bull Jard Bot Nat Belg 46 (1/2): 13 [ ? syn sub Fr. loricata] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 23a, microfiche 56 fig 3901 ["olim Frull Bessonii St, Frull laxepinnata St nsp olim; Madagascar; Sikora 1891, inter 56"] 250 Bryophyt Bibl 48: 66 [syn] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Pearson 1891 (L. & L.) Nees 1838 [nom inval, Art. 32.1(c)] Basionym: Distribution: comb nov Nees 1838 = Frullania beyrichiana 261 Naturg Europ Leberm 3: 210, footnote (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 beyrichiana var. granatensis Gottsche 1864 Typus: Colombia, Fusagasuga 2100m Lindig Basionym: Distribution: Colombia Gottsche Bonner 1864 1965 = Frullania beyrichiana beyrichii 478 655 Ann Sci Nat Bot, ser 5, 1: 174-175 Index Hepat 5: 247 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 (L. & L. in Lehm.) Nees by Spruce 1884 [orth var] Basionym: Distribution: Spruce Stotler Stotler 1884 1969 1970 = Frullania beyrichiana bicornistipula 243 Trans Proc Bot Soc Edinburgh 15: 48 4761 Nova Hedwigia 18: 442 194 The genus Frullania, separata: 442 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. ex Verd. 1930 [pro syn] Basionym: Distribution: nom herb Verdoorn = Frullania intermedia 1930 549 Nova Guinea 14 (4): 541 fo. billardieriana (Nees & Mont. in Mont.) Verd. 1930 Frullania intermedia (Reinw., Blume & Nees) Nees 1845 Page 49 bicornistipula Steph. in Gradstein & Hekking 1989 [error] Basionym: Distribution: Gradstein & Hekking = Frullania bicornistipula 1989 517 J Hatt Bot Lab 66: 208 [Guyana; = Fr. bicornistipula SPRUCE] Spruce 1884 bicornistipula var. bradeana (Steph.) Clark & Svihla 1951 Basionym: Frullania bradeana Distribution: stat nov Steph. 1924 Clark & Svihla 1951 = Frullania bicornistipula 341 Bryologist 54: 124-126, fig. 1-21(:125) Spruce 1884 bicornistipula var. mucroloba (Clark & Svihla) Clark & Svihla 1951 Basionym: Frullania mucroloba Distribution: Colombia stat nov Clark & Svihla 1949 Clark & Svihla Bonner Stotler Stotler Gradstein & Hekking Uribe & Gradstein = Frullania bicornistipula bicornustipula 1951 1965 1969 1970 1979 1998 341 655 4761 194 483 707 Bryologist 54: 124-126 Index Hepat 5: 247 Nova Hedwigia 18: 459-460 The genus Frullania, separata: 459-460 J Hatt Bot Lab 45: 112 [Cat Colombia] Bryophyt Bibl 53: 35 [Catalogue Colombia] Spruce 1884 Steph. 1889 {Neu Guinea} Typus: PNG, Western Fly River branch -.2.1886, Bäuerlen 51 in hb G (ex hb MEL) [holotypus] Basionym: Distribution: spec nov - nom nov Stephani Stephani Stephani Verdoorn Bonner Hattori Grolle & Piippo Stephani 1889 1889 1911 1930 1965 1982 1984 1985 940 941 686 549 655 495 143 928 Hattori & Piippo 1986 557 = Frullania plumaeformis Hedwigia 28: 157-158, pl 4(1-7) [Australia] Hedwigia 28: 278 [as Fr. plumaeformis] Spec Hepat 4: 575 [not mentioned as syn sub Fr. billardieriana] Nova Guinea 14 (4): 541 Index Hepat 5: 248 J Hatt Bot Lab 51: 239 Acta Bot Fennica 125: 18 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Homotropantha] D- 1, microfiche 54 fig 3703 ["olim Frullania plumaeformis (bicornustipula), Nova Guinea, Fly River Branch, Bäuerlen 1885, Herb Melbourne"] - 3704 [missing] Acta Bot Fennica 133: 40 Steph. 1889 Frullania billardieriana Frullania intermedia Nees & Mont. in Mont. 1843 (Reinw., Blume & Nees) Nees 1845 Page 50 bicornustipula Spruce by Steph. 1905 [orth var] Basionym: Distribution: Stephani 1905 = Frullania bicornistipula bicornuta 934 Hedwigia 45: 229 Spruce 1884 Steph. 1911 Typus: Chile. sd, Gay in hb G (hb BESCHERELLE 126) Basionym: Distribution: spec nov Stephani Kühnemann 1911 1937 - syn nov Herzog Herzog in Skottsberg 1938 1942 Kühnemann Bonner Miller, Whittier & Whittier 1949 1965 1983 Stephani 1985 Yuzawa 1988 - syn nov = Frullania stipatiloba bicornuta 686 Spec Hepat 4: 665-666 1031 Revista Centro Estud Doct Ci Nat, Buenos Aires 1 (3): 166 [Patagonia, Tierra del Fuego] 723 Arch Esc Farm Fac Ci Med Cordoba 7: 29 722 Nat Hist Juan Fernandez & Easter Island, 2: 740 [Fr. bicornuta may be or not synonym sub Fr. stipatiloba] 657 Lilloa 19: 354 [Argentina: Patagonia, Tierra del Fuego] 655 Index Hepat 5: 248 609 Prod Flora Hepat Polynesiae: 122 [? = Goebeliella cornigera (MITT) STEPH] 928 Stephani Icon Hepat Inedit [Diastaloba] B-41, microfiche 49 fig 3278 [from type; "Chile (Gay leg) Herb Bescherelle"] 511 J Hatt Bot Lab 64: 441-442, fig 3 [from type] [syn sub Fr. stipatiloba] Steph. 1894 Steph. ex Paris 1910 [nom nud] Basionym: Distribution: Paris = Frullania bicornuta 1910 Steph. 1911 Frullania stipatiloba bidentula 1455 Rev Bryol 37 (6): 130 [New Caledonia: Pic des Sources; mont Mou] Steph. 1894 Steph. in Sawada 1909 [nom nud] Basionym: Distribution: Sawada = Frullania bidentula 1909 930 Bot Mag Tokyo 23 (no 268): 178-183, fig 13, 13' [key to Japanese species, in Japanese] Steph. 1910 Frullania delavayi Frullania inflexa Steph. 1894 Mitt. 1861 Page 51 bidentula Steph. 1910 Typus: Japan. Shikoku, Prov Awa, Mt Tsurugi-san -.4.1900, U. Faurie 664 in hb G in hb KYO [syntypus] in hb HIRO [syntypus] Basionym: Distribution: spec nov - syn nov Sawada 1909 930 Stephani Iishiba Hattori in Nakai Hattori Hattori Kamimura Hattori & Noguchi Kamimura Bonner Stephani 1910 1912 1942 1944 1952 1952 1960 1961 1965 1985 686 888 1706 2791 529 873 965 199 655 928 Wu in Li, Gao, Wu et al. Yuzawa NN 1985 2001 2003 1169 1660 1397 Yamada & Iwatsuki 2006 1470 = Frullania delavayi Steph. 1894 Frullania inflexa bidentula Bot Mag Tokyo 23 (no 268): 178-183, fig 13, 13' [key to Japanese species, in Japanese] Spec Hepat 4: 398 Bot Mag Tokyo 26: 35 [Japan, with key in Japanese] Iconogr Pl Asia Orient 4 (3): 405-406, pl 129(a-r) J Jap Bot 20 (2): 263, fig 47 J Hatt Bot Lab 8: 29 [may be conspecific with Fr. devalayi] Contrib Fl Hepat Shikokuensem, Kochi: 132 Index spec Typ herb Japon, Bryophyta: 16 J Hatt Bot Lab 24: 83 [syn sub Fr. devalayi] Index Hepat 5: 248 Stephani Icon Hepat Inedit [Galeiloba] C-229, microfiche 53 fig 3647 ["Japan, Provinz Tsurugizan, Faurie 664, 1900"] Bryoflora Xizang (Tibet), Science Press, Beijing: 527 [sub Fr delavayi] Natural Environmental Sci Res 14: 43 [Japan, syn sub Fr inflexa] Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] J Hatt Bot Lab 99: 28 [Catalog of Japan] Mitt. 1861 ex Steph. 1985 [pro syn] Basionym: Distribution: Stephani 1985 = Frullania filiformis Stephani Icon Hepat Inedit [Galeiloba] C- 18, microfiche 50 fig 3369 [from type; "Madagascar, Forsyth Major (olim bidentula)"] Steph. 1910 Frullania loricata bilabiata 928 Pearson 1891 Steph. 1911 Typus: Madagascar sd, s coll in hb G (hb HARIOT 38, ex hb PC) [holotypus] Basionym: Distribution: Madagascar spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania loricata 686 655 185 928 Spec Hepat 4: 489-490 [as Fr. bilobiata on p. 471, errata on p. 824] Index Hepat 5: 248 Bull Jard Bot Nat Belg 46 (1/2): 92 & 97 Stephani Icon Hepat Inedit [Thyopsiella] F- 15, microfiche 56 fig 3891 ["Madagascar, com Hariot 38, Herb Paris; auch aus Nyassaland, Mt Malesa, leg Whyte"] - 3892 250 Bryophyt Bibl 48: 67 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Pearson 1891 Page 52 billardieri Nees & Mont. by Steph. 1985 [orth var] Basionym: Distribution: Stephani 1985 = Frullania billardieriana Stephani Icon Hepat Inedit [Homotropantha] D- 1, microfiche 54 fig 3703 ["olim Frullania plumaeformis (bicornustipula), Nova Guinea, Fly River Branch, Bäuerlen 1885, Herb Melbourne"] - 3704 [missing] Nees & Mont. in Mont. 1843 Frullania intermedia billardieriana 928 (Reinw., Blume & Nees) Nees 1845 Nees & Mont. in Mont. 1843 Typus: Indonesia. Mollucas, Amboina sd, La Billardiére, sn in hb S (B22592-B22593) [typus] in hb G (hb STEPHANI 171) Basionym: Distribution: Indonesia (Ambon, Ceram), Philippines, Vanuatu (Aneityum), New Caledonia spec nov Nees & Montagne in Montagne Nees & Montagne in G., L. & N. Montagne Sande Lacoste Schiffner Stephani Stephani in Stephani & Watts Verdoorn Verdoorn Verdoorn Bonner 1843 1845 1856 1864 1898 1911 1914 1928 1930 1930 1965 Hattori Hattori Hattori Miller, Whittier & Whittier Grolle & Piippo Stephani 1980 1982 1982 1983 1984 1985 Hattori Tan & Engel Hattori & Piippo Menzel Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Myrdal 1986 1986 1986 1988 2002 Ann Sci Nat Bot, ser 2, 19: 256 [Amboina] Syn Hepat: 434-435 Sylloge generum specierumque cryptogamaram: 85 Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 313 [Ceram, Nova Guinea] Conspect Hepat Archip Indici, Batavia: 321-322 Spec Hepat 4: 575-576 J Proc Roy Soc New S Wales 48: 106 [Aneityum] Rev Bryol NS 1: 112-113, fig 2 Nova Guinea 14 (4): 546 [comb nov as Fr. intermedia fo. billardieriana] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 248-249 [erroreously included New Guinea; vide Hattori 1982: 263] 490 J Hatt Bot Lab 47: 197-200, fig 20 [as distinct species; review] 495 J Hatt Bot Lab 51: 263 [not in New Guinea] 495 J Hatt Bot Lab 51: 239 [Sarawak specimens = Fr. intermedia ] 609 Prod Flora Hepat Polynesiae: 122-123 143 Acta Bot Fennica 125: 18 [Cat. West Melanesia] 928 Stephani Icon Hepat Inedit [Homotropantha] D- 1, microfiche 54 fig 3703 ["olim Frullania plumaeformis (bicornustipula), Nova Guinea, Fly River Branch, Bäuerlen 1885, Herb Melbourne"] - 3704 [missing] 506 J Hatt Bot Lab 60: 221 4286 J. Hatt. Bot. Lab.. No. 60: 335 557 Acta Bot Fennica 133: 40 512 J Hatt Bot Lab 65: 189, footnote 49 [Bornean record = Fr. intermedia] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 2002 1276 Mosskollekter [Frullania types in hb S] = Frullania intermedia bilobulata Typus: 470 2850 1912 842 831 686 220 242 549 551 655 (Reinw., Blume & Nees) Nees 1845 Steph. ex Merrill 1906 [nom nud] Merrill 6416 Basionym: Distribution: Merrill Verdoorn = Frullania junghuhniana 1906 1930 1496 Philippine J Sci C, Bot 5: 311 [Philippines: Mt. Pulog] 293 Ann Bryol Suppl 1: 134 Gottsche 1845 Page 53 binominata Steph. 1924 Typus: Australia. New South Wales, Mt. Wilson, The Avenue 11.1.1911, W. W. Watts 1094 in hb G 18198 Basionym: Distribution: Australia spec nov - syn nov Stephani Bonner Hattori Hattori Stephani 1924 1965 1979 1983 1985 687 655 485 499 928 Scott & Bradshaw Carthy Glenny, Meagher & Brown 1986 2003 2004 56 1125 1126 Glenny, Meagher & Brown 2004 1127 McCarthy 2006 1385 1911 1925 1965 1969 1970 1984 1985 686 1586 655 4761 194 142 928 Spec Hepat 4: 491-492 Arquivos de botânica do Estado de Sao Paulo 1 (2): 46 [Brazil] Index Hepat 5: 249 Nova Hedwigia 18: 436 The genus Frullania, separata: 436 Willdenowia 14: 509 [syn of Fr. brasiliensis] Stephani Icon Hepat Inedit [Thyopsiella] F- 34, microfiche 57 fig 3919 ["Peru, S Garan (Lechler leg)"] 686 293 204 199 655 609 928 Spec Hepat 4: 459 Ann Bryol Suppl 1: 34 Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] J Hatt Bot Lab 24: 17 [syn sub Fr squarrosa] Index Hepat 5: Prod Flora Hepat Polynesiae: 123 Stephani Icon Hepat Inedit [Galeiloba] C-109, microfiche 51 fig 3494 [from type; "Birma (Berthoumien leg)"] = Frullania falciloba bipinnata Spec Hepat 6: 545 Index Hepat 5: 249 J Hatt Bot Lab 45: 325, fig 2 [propably a weak form of Fr. falciloba] J Hatt Bot Lab 54: 146 Stephani Icon Hepat Inedit [Galeiloba] C-197, microfiche 53 fig 3602 ["Australia N.S.W., Watts legit 1094"] Brunnonia 8 (1): 33 [Australia] Flora Australia Suppl Ser 21: 43 [sub Fr falciloba] NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Checklist Australian Liverworts [Australia, syn sub Fr falciloba] Lehm. 1844 Steph. 1911 Typus: Peru, San Garan sd, Lechler sn in hb G (ex hb STR) Basionym: Distribution: Peru, Chile spec nov - syn nov - syn nov Stephani Herzog Bonner Stotler Stotler Menzel Stephani = Frullania brasiliensis birmensis Typus: Birma: Raddi 1822 Steph. 1910 Berthoumien in hb G 3494 & Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Frye & Clark Kamimura Bonner Miller, Whittier & Whittier Stephani = Frullania squarrosa Frullania ericoides 1911 1930 1947 1961 1965 1983 1985 (Reinw., Blume & Nees) Nees 1845 (Nees) Mont. 1839 Page 54 biroana Steph. 1910 Typus: PNG: Morobe, Huon Golf, Simbang 21.8.1899, Biro 111 in hb G (ex hb B) Basionym: Distribution: New Guinea spec nov - syn nov Stephani Verdoorn Bonner Grolle & Piippo Stephani 1911 1930 1965 1984 1985 686 549 655 143 928 Stephani 1985 928 Spec Hepat 4: 459-460 Nova Guinea 14 (4): 541 Index Hepat 5: 249 Acta Bot Fennica 125: 18 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Galeiloba] C-110, microfiche 51 fig 3495 ["Frullania Galeiloba, Biroana St nsp, N Guinea, Simbang, Huon Golf, Biro 111" - ""] Stephani Icon Hepat Inedit [Galeiloba] C-110, microfiche 51 fig 3495 [from type; "N Guinea, Simbang, Huon Golf, Biro 111" - "Frullania Galeiloba"] = Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides biroana (Nees) Mont. 1839 Steph. in Schumann & Lauterbach 1905 [nom nud] Typus: Kaiser-Wilhelmsland: Simbang auf Baumstämmen 21.8.1899, Biro no 111 & 112 Basionym: Distribution: nom nud Schumann & Lauterbach = Frullania biroana 1905 1492 Nachttrag Flora Deutsch Schutzgebiete: 33 Steph. 1910 Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides blanda (Nees) Mont. 1839 De Not. 1858 [nom nud - nom herb ?] Typus: Italy, sopra Genua -.11.1856, De Notaris, ERBARIO CRITTOGAMICO ITALIANO 14 in hb G Basionym: Distribution: Italy Notaris Notaris 1858 1865 576 Bonner 1965 655 Erbario Crittogamico Italiano No 14 Mem Reale Accad Sci Torino, ser 2, 22 (1): 378 [as Fr tamarisci var blanda] Index Hepat 5: 250 = Frullania tamarisci var. atrovirens Carrington 1863 Page 55 boivini Steph. 1911 Typus: Comores. Grand Comore 1850, Boivin in hb G (hb CORBIERE 132) Basionym: Distribution: spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania serrata boliviana 686 655 185 928 Spec Hepat 4: 481 Index Hepat 5: 250 Bull Jard Bot Nat Belg 46 (1/2): 111 Stephani Icon Hepat Inedit [Thyopsiella] F- 1, microfiche 56 fig 3866 ["Ins Comores, Boivin 1850, ex Herb Corbiere No 132"] 250 Bryophyt Bibl 48: 68 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 221 [syn sub Fr serrata, West Africa] Gottsche 1845 J.B.Jack ex Bonner 1965 [nom herb] [non SPRUCE 1890] Typus: Bolivia. Bolivianische Cuesta, nördl. v. Oran 1873, P. G. Lorentz sn in hb G sub Frullania tetraptera (ex hb JACK) Basionym: Distribution: Ecuador Bonner = Frullania tetraptera boliviana 1965 655 Index Hepat 5: 251 Nees & Mont. 1838 Spruce 1890 [non Jack ex Bonner 1965] Typus: Bolivia, Mapiri 5000ft. 1886, Rusby 3039 in hb NY [lectotypus] Basionym: Distribution: Mexico, Bolivia spec nov - syn nov Spruce Stephani Stephani Bonner Stephani 1890 1910 1916 1965 1985 898 686 222 655 928 Fulford & Sharp Yuzawa Gradstein, Meneses & Arbe 1990 1991 2003 105 525 975 = Frullania cuencensis Mem Torrey Bot Club 1: 118 Spec Hepat 4: 357-358 Biblioth Bot 87: 238 [Bolivia] Index Hepat 5: 250 Stephani Icon Hepat Inedit [Chonanthelia] A- 61, microfiche 47 fig 3156 Mem NY Bot Gard 63: 40 [Mexico] J Hatt Bot Lab 70: 209 [syn sub Fr cuencensis] J Hatt Bot Lab 91: 52 [Bolivia; as syn sub Fr cuencensis] Taylor 1846 Page 56 bomboizana Steph. 1924 Typus: Ecuador: Via Bomboiza - Gualaquiza, in silva secus rivulum Yumara, 950-1000 m 30.1.1910, Allioni 196 in hb G 20952 [holotypus] Basionym: Distribution: Ecuador spec nov Stephani Bonner Yuzawa Stephani 1924 1965 1984 1985 687 655 502 928 - syn nov Yuzawa 1991 525 = Frullania confertiloba boninensis Spec Hepat 6: 530-531 Index Hepat 5: 251 J Hatt Bot Lab 57: 401-403, fig. 14(a-m) from type Stephani Icon Hepat Inedit [Chonanthelia] A- 62, microfiche 47 fig 3157-3158 [from type; "Ecuador Allioni 196 (1912)"] J Hatt Bot Lab 70: 248 Steph. 1910 Horikawa 1934 Typus: Japan. Bonin, Isl. Chichi-jima, Mt. Chu-o " Mt. Chûôsan" 27.7.1930, Y. Horikawa 1954 in hb HIRO [holotypus] Basionym: Distribution: spec nov - syn nov Horikawa Miller Miller Hattori & Noguchi Iverson Bonner Miller Inoue & Iwatsuki Miller, Whittier & Whittier Yuzawa NN 1934 1960 1960 1960 1963 1965 1968 1969 1983 2000 2003 Yamada & Iwatsuki 2006 = Frullania meyeniana borgenii 34 920 920 965 J Sci Hiroshima Univ, ser B, div 2, 2: 239-240, fig 45 Bryologist 63: 121 [Micronesia] Bryologist 63: 121 [Micronesia] Index spec Typ herb Japon, Bryophyta: 16 : 122 655 Index Hepat 5: 251 672 Micronesia 4 (1): 74 [key to Guam & Northern Micronesia] 61 Bull Natn Sci Mus Tokyo 12 (2): 303 [syn sub Fr. meyeniana] 609 Prod Flora Hepat Polynesiae: 123 1659 Natural Environmental Sci Res 13: 18 [Japan, syn sub Fr meyeniana] 1397 Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] 1470 J Hatt Bot Lab 99: 29 [Catalog of Japan] Lindenb. 1845 Gottsche ex Steph. 1985 ["nomen occupatum"] Typus: Stephani Icon Hepat Inedit [Galeiloba] C- 40ab, microfiche 50 fig - 3394 ["Frullania Borgenii Gms (nomen occupatum), Copie nach Gottsche, Madagascar"] Basionym: Distribution: Stephani = Frullania subplana Frullania bullata 1985 928 Stephani Icon Hepat Inedit [Galeiloba] C- 40b, microfiche 50 fig 3394 ["Frullania Borgenii Gms (nomen occupatum), Copie nach Gottsche, Madagascar"] - 3395 ["Madagascar, Borgen"] Gottsche ex Steph. 1910 Steph. 1910 Page 57 borgenii Pearson 1890 Typus: Madagascar. "in montibus Ankaratra" 1879, Borgen in hb G (hb KIAR 99c, ex hb Pearson) in hb O [holotypus] Basionym: Distribution: Angola, Democratic Republic of Congo, Mozambique, Zambia, Zimbabwe, South Africa - syn nov Pearson 1891 Stephani Demaret & Vanden Berghen Arnell Bonner Vanden Berghen 1910 1946 1963 1965 1972 Vanden Berghen Stephani 1976 1985 Best Grolle Wigginton 1990 1995 2002 Manyanga & Perold Wigginton Wigginton 2004 2004 2004 = Frullania obscurifolia borneensis 1921 Christiania Videnskabs-Selskabs Forhandlinger 1890 (2): 8-9, pl 4(116) [Madagascar] 686 Spec Hepat 4: 369 1102 Bull Jard Bot Etat Bruxelles 18: 122-124, fig 7 822 Hepat S Africa: 145-147, fig 107(a-g) 655 Index Hepat 5: 252 249 Expl Hydrobiol Bassin Bangweolo et Luapula 8 (1): 74-76, fig 29(a-i) [Zambia, Zaire] 185 Bull Jard Bot Nat Belg 46 (1/2): 143 [as borgeni] 928 Stephani Icon Hepat Inedit [Galeiloba] C- 7, microfiche 50 fig 3358 ["Diastaloba, Kiar & Pearson, Madagascar"] 879 Kirkia 13 (2): 296 [Zimbabwe] 250 Bryophyt Bibl 48: 67 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1301 SABONET Report 21: 4 [Zimbabwe] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 233 [syn sub Fr obscurifolia, West Africa] Mitt. 1879 Steph. 1910 Typus: Indonesia. Sarawak, Lunda sd, Micholitz in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Stephani = Frullania sublignosa 1910 1929 1930 1965 1985 686 245 293 655 928 Spec Hepat 4: 349 Ann Bryol 2: 122 [syn sub Fr. sublignosa] Ann Bryol Suppl 1: 61-62, fig 91-93 Index Hepat 5: 252 Stephani Icon Hepat Inedit [Chonanthelia] A- 85, microfiche 47 fig 3190 [from type; "Lunda Sarawak Borneo Micholitz"] Steph. 1894 Page 58 bosseana Steph. 1911 Typus: Indonesia. Irian Jaya, Sorong, Waigeoe Insula, Sinus Saripi 1899, Weber van Bosse sn in hb G 16558 (1899, Weber van Bosse sn, hb LEVIER 2840) [lectotypus] - Hattori 1980 in hb G 16559 (hb LEVIER 2834) Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Hattori Miller, Whittier & Whittier Grolle & Piippo Stephani 1911 1930 1930 1965 1976 1983 1984 1985 686 293 549 655 67 609 143 928 Konrat & Braggins 2001 838 = Frullania hypoleuca boveana Spec Hepat 4: 655 Ann Bryol Suppl 1: 126 Nova Guinea 14 (4): 541, 545 [syn nov sub Fr. hypoleuca] Index Hepat 5: 252 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 22 Prod Flora Hepat Polynesiae: 123 Acta Bot Fennica 125: 18 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Diastaloba] B-60, microfiche 49 fig 3300 [from type; N Guinea, Insula Waigeoe, leg Dr Weber v Bosse, Levier 2840, 2834"] J Hatt Bot Lab 91: 250-256 [syn sub Fr. hypoleuca] Nees 1843 [auct. non C.Massal. 1885] Steph. 1911 {Chile} Typus: Chile, Brunswick Peninsula B. Aranz Basionym: Distribution: - syn nov Stephani Engel 1911 1978 = Frullania magellanica bowelliana 721 182 Kungl Svenska Vetenskapsakad Handl 45 (9): 89 Fieldiana Bot 41: 235 F.Weber & Nees 1845 Steph [orth. error in herbar label] Typus: Samoa, Futuila: 600m, Dezember 1894, leg. Dr. Reincke - Baenitz 10.10.1902 https://www.gbif.org/occurrence/436951355 MNHN - Museum national d'Histoire naturelle hb PC PC0104315 https://api.gbif.org/v1/image/unsafe/http%3A%2F%2Fmediaphoto.mnhn.fr%2Fmedia%2F1444584216612GiSxyN1R6 8Z8aqpS Basionym: Distribution: Reinecke = Frullania powelliana Frullania utriculata brachycarpa brachycarpa] 0 :315 [ F. powelliana St. Hedw. 1894 p. 154, Samoa (P., R. n. 63] Steph. 1894 Steph. 1894 Spruce 1884 ex Yano 2008 [type of brachiclada mixed with name of Basionym: Distribution: Yano = Frullania brachyclada 2008 1684 Boletim Instituto Botanica 19: 30 [type of brachiclada mixed with name of brachycarpa; Brazil; catalog] Spruce 1884 [hom illeg] [non Taylor ex Lehm. 1844] Frullania tunguraguana L.Clark & Frye 1952 Page 59 brachyclada Taylor ex Lehm. 1844 [non Spruce 1884] Typus: Tasmania. "Terra van Diemen" 1824, Rob. Neill in hb G (ex hb BESCHERELLE) Basionym: Distribution: spec nov - comb nov Taylor ex Lehmann Gottsche in G., L. & N. Trevisan 1844 1845 1877 Stephani Grolle Hamlin Scott & Bradshaw 1911 1967 1972 1986 = Lepidolaena brachyclada brachyclada 589 Pugillus 8: 21-22 2847 Syn Hepat: 247 [syn sub Polygotus brachycladus] 1576 Mem Reale Ist Lombardo Sci ser 3, Cl Sci Math 4: 393 [ as Lepidolaena brachyclada] Icon Zeitschrift Memorie del Reale Istituto Lombardo di Scienze e Lettere, Classe di Scienze Matematiche e Naturali Milano:, 3.Ser. 4: 393 686 Spec Hepat 4: 50 [sub Lepidolaena] 624 J Hatt Bot Lab 30: 43 [syn sub Lepidolaena brachyclada] 659 Rec Dominion Mus 7(19): 261 [New Zealand] 56 Brunnonia 8 (1): 33 [Australia] (Taylor ex Lehm.) Trevisan 1877 Spruce 1884 [hom illeg] [non Taylor ex Lehm. 1844] Typus: Ecuador. Tunguragua, Mt. Tunguragua sd, Spruce sn in hb G in hb BM [isotypus] in hb US in hb E E00002518 - E00256363 [isotypus] Basionym: Distribution: Ecuador, Bolivia, Argentina spec nov - nom nov - syn nov Spruce Stephani Stephani Kühnemann Clark & Frye 1884 1910 1916 1949 1952 243 686 222 657 343 Bonner Stephani 1965 1985 655 928 Yuzawa, Mues & Hattori Yuzawa NN 1987 1991 1996 509 525 632 = Frullania tunguraguana brachyscypha Trans Proc Bot Soc Edinburgh 15: 11 & 15 (No 1) Spec Hepat 4: 331-332 [Anden] Biblioth Bot 87: 239 [Bolivia] Lilloa 19: 354 [Argentinia: Cordillera] Bryologist 55: 133-136 - nom. illeg. (ICN Art. 53.1, non Lehm. 1844) [≡ Frullania tunguraguana Clark & Frye (1952: 133)]. Index Hepat 5: 252-253 Stephani Icon Hepat Inedit [Chonanthelia] A- 25ab, microfiche 46 fig 3104-3106 ["Bolivia - Bogota 1716 Lindig"] J Hatt Bot Lab 63: 425-436 [Morph/Syst] J Hatt Bot Lab 70: 278, fig. 50 [as tunguraguana], map 4c(:283) gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 05 of 35 [in hb US] L.Clark & Frye 1952 Spruce [nom. herb.] Typus: Mexico, Orzaca 4000 ft, ad rupes, H.G. Galeotti no 6997 https://www.gbif.org/occurrence/436803032 https://www.gbif.org/occurrence/1291545332 NHN - Museum national d'Histoire naturelle hb PC Basionym: Distribution: = Page 60 bradeana Steph. 1924 Typus: Costa Rica 1912, C. Brade sn in hb G Basionym: Distribution: Costa Rica spec nov - stat nov Stephani Clark & Svihla Herzog Bonner Stephani 1924 1951 1953 1965 1985 687 341 640 655 928 Spec Hepat 6: 545-546 Bryologist 54: 124-126 [= Fr. bicornistipula var bradeana] Rev Bryol Lichen 20: 128 [Costa Rica] Index Hepat 5: 253 Stephani Icon Hepat Inedit [Thyopsiella] F- 36, microfiche 57 fig 3922 ["Costa rica, leg Brade"] = Frullania bicornistipula var. bradeana (Steph.) Clark & Svihla 1951 Frullania bicornistipula brasiliensis Spruce 1884 [auct. non RADDI 1822] Pearson 1890 {Madagascar} Basionym: Distribution: Pearson 1891 Vanden Berghen 1976 Grolle 1995 = Frullania longistipula 1921 Christiania Videnskabs-Selskabs Forhandlinger 1890 (2): 6 [Madagascar] 185 Bull Jard Bot Nat Belg 46 (1/2): 73 [Madagascarian specimen of PEARSON = Fr. longistipula] 250 Bryophyt Bibl 48: 66 Steph. 1891 brasiliensis var. mülleriana Gottsche 1863 Typus: Mexiko, Vera Cruz, Volcan Orizaba sd, Fr. Müller Basionym: Distribution: Mexico var nov - syn nov Gottsche Gottsche Fulford & Sharp = Frullania brasiliensis 1863 1867 1990 103 104 105 Separata of Mex. Leverm.: 246 K Danske Vid Selks Skrifter, ser 5, 6: 342 [Mexico] Mem NY Bot Gard 63: 41 [Mexico] Raddi 1822 brasiliensis var. robusta Spruce 1884 Typus: Ecuador. In monte Tunguragua sd, Spruce sn in hb G sub Fr. robusta in hb E E00002580 - E00293968 (1857, Spruce sn) - E00293967 [isotypus] Basionym: Distribution: var nov Spruce 1884 243 - syn nov Stephani Bonner 1894 1965 228 655 = Frullania spruceana Trans Proc Bot Soc Edinburgh 15: 51 [only as "var ? robusta" published] Hedwigia 33: 16 [syn sub Fr. spruceana STEPHANI] Index Hepat 5: 253 Steph. 1894 Frullania robusta Steph. 1911 [non Kiaer ex Renauld & Cardot 1891, nec Spruce [Hepat exsicc] ex Steph. 1911] Frullania brasiliensis Raddi 1822 Page 61 braziliensis Steph. by Kappelle 1996 [orth var] Basionym: Distribution: Costa Rica Kappelle = Frullania brasiliensis brevipinna Steph. 1911 Typus: Jamaica. sd, W. Harris 1996 1409 Los Bosques de Roble, Cordillera de Talamanca, Costa Rica: 257 Raddi 1822 in hb G (ex hb URBAN) Basionym: Distribution: Jamaica spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 Söderström, Hagborg, Vana & Konrat 2011 = Frullania ecuadorensis Frullania kunzei breviplicata 686 655 4761 194 928 Spec Hepat 4: 528 Index Hepat 5: 254 ["brevipinnia"] Nova Hedwigia 18: 471-472 The genus Frullania, separata: 471-472 Stephani Icon Hepat Inedit [Thyopsiella] F-102, microfiche 58 fig 4025 ["Jamaica, (W Harris leg) Herb Urban"] 2581 Bryologist 114(1) : 72 [checklist Jamaica] Steph. 1911 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Glenny, Meagher & Brown 2003 [nom nud] Typus: Basionym: Distribution: Australia (NSW) Carthy Glenny, Meagher & Brown 2003 2003 Glenny, Meagher & Brown 2003 1125 Flora Australia Suppl Ser 21: 112 [Australia (NSW, QLD)] 1126 NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html = brevisulca Mitt. by Chopra 1943 [orth var] Basionym: Distribution: Chopra Chopra Parihar = Frullania breviuscula Frullania neurota 1938 1943 1962 933 Proceed Indian Acad Sci, sect B, 7 (5): 248 [South India] 205 J Indian Bot Soc 22: 249 [India] 1584 Univ Allahabad studies, Bot sect 1-2: 18 [India,syn sub Fr neurota] Mitt. 1861 Taylor 1846 Page 62 breviuscula Mitt. 1861 Typus: India. Khasi Mts, 4000 ft sd, J.D. Hooker & T. Thomson 1551 in hb NY in hb G [isotypus] Basionym: Distribution: China, India (Eastern Himalayas: Megalaya) spec nov - syn nov Mitten Stephani Verdoorn Chopra Bonner Hattori 1861 1910 1930 1943 1965 1975 Miller, Whittier & Whittier Stephani 1983 1985 Piippo Singh, Kumar & Nath 1990 2008 = Frullania neurota Basionym: Frullania truncatifolia Distribution: - syn nov brunnea (Steph.) R.M.Schust. & S.Hatt. in S.Hatt. 1975 Steph. 1910 Hattori Hattori Hattori Hattori Choe Hattori Piippo Yamada & Choe Yuzawa Yamada & Iwatsuki = Frullania muscicola J Linn Soc Bot 5: 120 [India: Khasi mount.] Spec Hepat 4: 350-351 Ann Bryol Suppl 1: 66-68, fig 96-103 [type identical with Fr. neurota] J Indian Bot Soc 22: 249 [India] Index Hepat 5: 253 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 75 [remarks on MITTEN 1861] 609 Prod Flora Hepat Polynesiae: 123 928 Stephani Icon Hepat Inedit [Chonanthelia] A- 95, microfiche 47 fig 33205-3206 ["Khasi Mt (La?orie legit) Original"] 521 J Hatt Bot Lab 68: 38 [annotated catalogue of China] 1281 Taiwania 53 (1): 56, [syn sub Fr neurota] Taylor 1846 brittoniae ssp. truncatifolia comb nov 574 686 293 205 655 47 1975 1979 1980 1980 1980 1981 1990 1997 2001 2006 410 418 130 489 1171 491 521 717 1660 1470 Misc Bryol Lichen 7 (2): 28 Misc Bryol Lichen 8 (4): 71- 73, fig 1 J Jap Bot 55 (5): 134 [Ryukyu, Japan] J Hatt Bot Lab 47: 123-124, fig 240 Illust Fl & Fauna Korea 24: 732 [as valid taxa; South Korea] J Hatt Bot Lab 49: 162-165 [syn sub Fr. muscicola] J Hatt Bot Lab 68: 38 [annotated catalogue of China] J Hatt Bot Lab 81: 287 [sub Fr muscicola]] Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] J Hatt Bot Lab 99: 29 [Catalog of Japan] Steph. 1894 Austin 1846 [hom illeg] [non (Spreng.) Drège 1843] Basionym: Distribution: - syn nov Austin Evans Frye & Clark = Frullania caroliniana Frullania obcordata 1846 1897 1947 237 204 Hep Bor Amer Exsic No 105e Trans Connet Acad Arts Sci 10 (1): 33-34 [syn sub Fr. caroliniana Univ Wash Publ Biol 6 (5): 760 Sull. 1846 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 63 brunnthaleri Steph. 1911 Typus: Africa: Usambara, Amani, 800 m -.8.1909, Brunnthaler sn in hb G (ex hb W) Basionym: Distribution: East Afrika spec nov - syn nov Stephani Stephani in Brunnthaler Bonner Vanden Berghen Stephani 1911 1913 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania apicalis brunnthaleri 686 219 655 185 928 Spec Hepat 4: 623-624 Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 728 [sp. nov.] Index Hepat 5: 255 Bull Jard Bot Nat Belg 46 (1/2): 52 Stephani Icon Hepat Inedit [Diastaloba] B- 5, microfiche 48 fig 3231 [from type; "Africa Usambara (Brunnthaler legit) 1909"] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apicalis, West Africa] Mitt. 1879 Steph. 1913 [nom superfl] Basionym: Distribution: Stephani in Brunnthaler = Frullania brunnthaleri 219 Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 728 [spec nov = nom superfl] Steph. 1911 Frullania apicalis bryhnii 1913 Mitt. 1879 Mueller in Rabenhorst 1915 Typus: Spain. Canary Isl, Teneriffa, Tacoronte, 550m 1908, N. Bryhn in hb S-PA [holotypus] in hb UPS [isotypus] Basionym: Distribution: spec nov - syn nov Müller in Rabenhorst Verdoorn 1915 1928 Schiffner Ade & Koppe Mägdefrau Müller Bonner Grolle 1936 1942 1943 1951 1965 1970 = Frullania polysticta bryhnii 1064 Krypt Flora (2ed) 6, 2: 618-619, fig 174(a-e) 2071 Rec Trav Bot Neerl 25a: 458, fig 7-8 [=Recueil des travaux botaniques néerlandais] 277 Österr Bot Zeitschr 85 (1): 25-26, 21[= nervosa] 206 Hedwigia 81 (1/2): 34 [Tenerifa] 2799 Flora NF 37: 135 [Teneriffa] 641 Rev Bryol Lichen 20: 135 655 Index Hepat 5: 255 615 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 309 [syn of Fr. polysticta) Lindenb. 1845 [auct. non Mueller 1915] Verd. 1928 Basionym: Distribution: Verdoorn Schiffner = Frullania nervosa 1928 1936 2071 Rec Trav Bot Neerl 25A: 452 277 Österr Bot Zeitschr 85 (1): 21 Mont. 1840 [non Mont. in Berthelot & Webb 1844] Frullania tamarisci (L.) Dumort. 1835 Page 64 bullatiflora Steph. 1911 Typus: Colombia. Bogota sd, Apollinaire in hb G (ex hb LACOUTURE) Basionym: Distribution: Mexico, Colombia, Ecuador spec nov - syn nov - syn nov Stephani Clark Bonner Stotler Stotler Menzel Stephani 1911 1954 1965 1969 1970 1984 1985 = Frullania macrocephala 686 364 655 4761 194 142 928 Spec Hepat 4: 513 Bryologist 57: 39-41, fig 1-13 [may be var. ?? ryoliata Herzog] Index Hepat 5: 256 Nova Hedwigia 18: 439 [syn sub Fr. macrocephala] The genus Frullania, separata: 439 [syn sub Fr. macrocephala] Willdenowia 14: 509 Stephani Icon Hepat Inedit [Thyopsiella] F- 39, microfiche 57 fig 3929 ["Bogota, Frére Appollinaire, Herb Lacouture"] - 3930 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 bullatiflora var. labiata Herzog in Clark 1954 [nom herb] Typus: Costa Rica. Prov. San José, La Palma, 1600m 3.2.1924, Paul C. Standley 32953 Basionym: Distribution: Clark = Frullania bullatiflora 1954 Bryologist 57: 39-41, fig 1-13 Steph. 1911 Frullania macrocephala bursicula 364 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1911 Typus: Soputh Africa. Cape Prov., "Cape of God Hope" sd, s coll in hb G (ex hb JACK) [holotypus] Basionym: Distribution: Mauritius, South Africa spec nov Stephani Sim Demaret & Vanden Berghen Arnell Bonner Augier 1911 1926 1947 1963 1965 1974 - syn nov Vanden Berghen Stephani 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania socotrana 686 252 1101 822 655 821 Spec Hepat 4: 678-679 Trans Roy Soc S Africa 15: 45, fig [South Africa] Bull Jard Bot Etat Bruxelles 18: 232-233, fig 34(a-e) Hepat S Africa: 143, fig 105(a-j) Index Hepat 5: 256 Ann Fac Sci Univ Fed Cameroun 17: 83-85, 91, fig 4(1-10) [Cameroun] [as Fr bursicola] 185 Bull Jard Bot Nat Belg 46 (1/2): 139 [syn sub Fr. socotrana] 928 Stephani Icon Hepat Inedit [Galeiloba] C- 9, microfiche 50 fig 3360 [from type; "Cap (Herb Jack)" & "auch Mauritius, leg Renauld"] 250 Bryophyt Bibl 48: 68 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 23 [syn sub Fr socotrana, West Africa] Mitt. 1888 Page 65 bystroemii S.W.Arnell 1961 Typus: Cape Verde Isl. San Vicente, 750m sd, Byström in hb UPS (Stockholm) [holotypus] Basionym: Distribution: spec nov - syn nov Arnell Vanden Berghen Wigginton 1961 1976 2002 Wigginton Wigginton 2004 2004 = Frullania spongiosa caespitosa 54 Bot Not 114 (2): 177-178, fig 1 185 Bull Jard Bot Nat Belg 46 (1/2): 167 &171 [syn sub Fr. spongiosa] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 234, fig 155(D,F,G) from type [syn sub Fr spongiosa,West Africa] Steph. 1894 Beauverd ex Bonner 1965 [nom herb] Basionym: Distribution: Stephani 1924 687 Bonner 1965 655 = Frullania caespitans caledonica Spec Hepat 6: 537 [ Fr. caespitosa is not found as syn of Fr. caespitans]] Index Hepat 5: 256 [syn sub Fr. caespitans according Stephani 1924: 537] Beauverd 1924 Gottsche ex Steph. 1894 [non (Schuster) Schuster 1992] Typus: New Caledonia. Balade sd, Vieillard 1759 in hb G (ex hb BESCHERELLE) Basionym: Distribution: New Caledonia spec nov -syn nov Gottsche ex Stephani Paris Stephani Verdoorn Verdoorn Frye & Clark Bonner Hattori Hattori Miller, Whittier & Whittier Hattori Stephani 1894 1910 1911 1930 1930 1947 1965 1977 1981 1983 1984 1985 Hattori Thouvenot et al. 1986 2011 = Frullania squarrosula 228 1455 686 293 551 204 655 479 492 609 503 928 Hedwigia 33: 156 Rev Bryol 37 (6): 130 [New Caledonia: Ouaoueoue; Dogny] Spec Hepat 4: 460-461 Ann Bryol Suppl 1: 34 (syn sub Fr. squarrosa) Nederl Kruidk Arch Jrg 1930 (2): Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] Index Hepat 5: 257 J Hatt Bot Lab 43: 414 [difference to Fr. squarrosa] [New Caledonia] J Hatt Bot Lab 49: 365-368, fig 4(a-m), 5(a-g), 6(a-d) Prod Flora Hepat Polynesiae: 123 J Hatt Bot Lab 57: 409 Stephani Icon Hepat Inedit [Galeiloba] C-112ab, microfiche 51 fig 3497 ["Nova Caledonia, Balade, Vieillard 1759, Herb Bescherelle"] 3498-3499 ["N Caledonia, Ile des Rus\, 534 A Le Rat, Januar 1902"] 506 J Hatt Bot Lab 60: 208 3452 Cryptogamie, Bryologie, 32(4): 320 (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Page 66 caledonica (R.M.Schust.) R.M.Schust. 1992 [hom illeg] [non Gottsche ex Steph. 1894] Basionym: Neohattoria caledonica Distribution: comb nov - nom nov R.M.Schust. 1970 Schuster Schuster Engel in Engel & Merrill = Frullania neocaledonica 761 603 756 J Hatt Bot Lab 33: 291-292, fig 5(1-5) [as Neohattoria caledonica] Hepat N America 5: [as Fr. caledonica] Novon 9: 344 [nom nov as Fr. neocaledonica] J.J.Engel 1999 Frullania chevalieri cambouena 1970 1992 1999 (R.M.Schust.) R.M.Schust. 1992 Steph. in Renauld & Cardot 1891 Typus: Madagascar. Imerina, Ankavavala, 1500-2000m 1889, Camboué in hb G 12978 (hb RENAULD 159 or 160) [lectotypus ?] in hb PC [holotypus] Basionym: Distribution: Tanzania, Mozambique, Madagascar spec nov - syn nov Stephani in Renauld & Cardot Stephani in Renauld & Cardot Stephani Stephani Bonner Vanden Berghen Stephani 1891 1891 1892 1910 1965 1976 1985 Grolle 1995 Sim-Sim, Fontihna, Mues & Lion 2000 Wigginton 2002 Wigginton = Frullania loricata camerunensis 2004 1602 1453 1121 686 655 185 928 Bull Soc Roy Bot Belg 30 (2): 198 Rev Bryol 18 (4): 56 [Madagascar] [St. n. sp.] Hedwigia 31: 201-202, pl 21, fig 24 & 25 Spec Hepat 4: 383 Index Hepat 5: 258 Bull Jard Bot Nat Belg 46 (1/2): 92 & 97 Stephani Icon Hepat Inedit [Galeiloba] C- 11, microfiche 50 fig 3362 [from type, "Thyopsiella, Ankadanala - Madagascar, Renauld 159, 160 , Camboué"] 250 Bryophyt Bibl 48: 66 749 Nova Hedwigia 71 (1-2): 192 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Pearson 1891 Steph. 1911 Typus: Cameroon: Ekunda Ndene 28.5.1891, P. Dusén sn in hb G (hb STEPHANI a11 & Icon ined 1152) Basionym: Distribution: W Africa spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Konrat & Braggins Wigginton 2001 2002 Wigginton Wigginton 2004 2004 = Frullania apiculata 686 655 185 928 Spec Hepat 4: 620-621 Index Hepat 5: 258 Bull Jard Bot Nat Belg 46 (1/2): 102 [syn nov sub Fr. apiculata] Stephani Icon Hepat Inedit [Diastaloba] B- 2, microfiche 48 fig 3227 [from type; "Kamerun Dusén 1891"] 838 J Hatt Bot Lab 91: 256-262 [syn sub Fr. apiculata] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apiculata, West Africa] (Reinw., Blume & Nees) Nees 1845 Page 67 campanensis Spruce 1884 Typus: Peru: Peruvian Andes, Mi. Campana, 1400 m sd, Spruce sn in hb G [isotypus] Basionym: Distribution: Peru, Colombia spec nov Spruce Stephani Weberbauer Bryan Weberbauer Soukup Bonner Crandall Gradstein & Hekking Menzel Stephani 1884 1910 1911 1929 1945 1955 1965 1969 1979 1984 1985 - syn nov Yuzawa Yuzawa Wigginton 1991 1991 2004 = Frullania obscura campanulata 243 686 792 175 793 171 655 193 483 142 928 Trans Proc Bot Soc Edinburgh 15: 11 & 16-17 (No 3) Spec Hepat 4: 332 Pflanzenwelt Peruan Anden: 238 [Peru] Bot Gazette 88(3): 338 [Peru] Mundo Vegetal Andes Peruanos: 568 [Peru] Biota 1(4): 81 [Peu) Index Hepat 5: 258 Beih Nova Hedwigia 30: 81 J Hatt Bot Lab 45: 112 [Cat Colombia] Willdenowia 14: 489 [Peru] Stephani Icon Hepat Inedit [Chonanthelia] A- 26, microfiche 46 fig 3107-3108 ["Mt Campana leg Spruce"] 525 J Hatt Bot Lab 70: 264 525 J Hatt Bot Lab 70: 267-268, fig. 43 [description of type specimen] 1172 Sript Bot Belg 30: 224 [syn sub Fr arecae, West Africa] (Sw.) Mont. 1840 [auct. non Sande Lac.] Steph. ex Herzog 1916 {Bolivia} Typus: Bolivia. Corani, 1800m 1911, Herzog 4752 in hb G G00069185 - n° SIB 124682/1 Basionym: Distribution: spec nov Stephani in Herzog 1916 - nom nov Stephani 1924 Bonner Stephani 1965 1985 Gradstein, Meneses & Arbe 2003 = Frullania caespitans capensis 4249 Bibliotheca Botanica 87: 242-243 , fig. 192 [Bolivia], as 361. Fr campanulata St. n. sp. (Galeiloba)] 687 Spec Hepat 6: 537 [nom nov: Fr caespitans BEAUVERD in STEPHANI] 655 Index Hepat 5: 259 928 Stephani Icon Hepat Inedit [Galeiloba] C- 51, microfiche 50 fig 3406 [from type; "Bolivia, Herzog 4752"] 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr caespitans] Beauverd 1924 [auct. non Gottsche 1845] Visnadi 1998 {Sao Paulo, Brazil} Basionym: Distribution: Visnadi Gradstein & Costa 1998 2003 Yano 2008 [vide #984: 306] Mem NY Bot Gard 87: 91 [capensis is only African species, not in Brazil] 1684 Boletim Instituto Botanica 19: 30 [Brazil; catalog] 984 = Page 68 capensis [auct. non Gottsche 1845] Sim 1926 Basionym: Distribution: - syn nov Sim Demaret & Vanden Berghen Arnell Vanden Berghen Wigginton 1926 1946 1963 1976 2002 Wigginton Wigginton 2004 2004 = Frullania borgenii 252 1102 822 185 1012 Trans Roy Soc S Africa 15: 46, figs Bull Jard Bot Etat Bruxelles 18: 122 [syn sub Fr borgeni] Hepat S Africa: 145 [syn sub Fr. borgenii] Bull Jard Bot Nat Belg 46 (1/2): 145 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 233 [syn sub Fr obscurifolia, West Africa] Pearson 1890 Frullania obscurifolia Mitt. 1879 capillacea Steph. 1911 Typus: Brasilia 1910, P. Dusén 8171 in hb G Basionym: Distribution: spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 = Frullania ecuadorensis Frullania kunzei capillaris 686 655 4761 194 928 Spec Hepat 4: 682-683 Index Hepat 5: 259 Nova Hedwigia 18: 471-472 The genus Frullania, separata: 471-472 Stephani Icon Hepat Inedit [Thyopsiella] F- 40, microfiche 57 fig 3931 ["Brasilia, leg Dusén 8171, leg 1910"] Steph. 1911 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. ex Paris 1910 [nom nud] Typus: Basionym: Distribution: Paris 1910 = Frullania capillaris capilliformis Typus: Sri Lanka 1455 Rev Bryol 37 (6): 130 [New Caledonia: Table Unio] Steph. 1911 Steph. 1911 [non Steph. 1916] sd, Nietner sn in hb G (hb JACK 1567) Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Clark Bonner Stephani = Frullania apiculata 1911 1930 1954 1965 1985 686 293 369 655 928 Spec Hepat 4: 639-640 Ann Bryol Suppl 1: 97 [syn sub Fr. apiculata] Bryologist 58: 68, 71 [not syn sub Fr. apiculata] Index Hepat 5: 260 Stephani Icon Hepat Inedit [Diastaloba] B-49, microfiche 49 fig 3289 [from type; "Ceylon Nietner Jack 1567"] (Reinw., Blume & Nees) Nees 1845 Page 69 capilliformis Steph. ex Herzog 1916 [hom illeg] [non Steph. 1911] Typus: Bolivia. Comarapa, 2600m 1911, Herzog 3967 in hb G G00128081 - n° SIB 143795/2 - 13669 (sub nom nov Fr. trichomorpha & drawing > see fr0650.jpg) [holotypus] Basionym: Distribution: spec nov - nom nov -syn nov Stephani Stephani Clark Bonner Stephani 1916 1924 1955 1965 1985 Gradstein, Meneses & Arbe Uribe & Gradstein 2003 2003 = Frullania brasiliensis caracensis 222 687 369 655 928 Biblioth Bot 87: 244-245, fig 196 [Bolivia] Spec Hepat 6: 547-548 [nom nov: as Fr. trichomorpha] Bryologist 58: 71 Index Hepat 5: 542 Stephani Icon Hepat Inedit [Meteriopsis] E-17, microfiche 55 fig 3789 ["Bolivia, Dr Herzog 3967 (1913)"] 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. trichomorpha] 1071 Crypt Bryol 24 (3): 200 [syn sub Fr brasiliensis] Raddi 1822 Steph. 1911 Typus: Venezuela. Caracas sd, A. Braun sn in hb G 00069150 - SIB 126180/1 (hb STEPHANI 20610, ex hb Alex BRAUN, JACK misit) [holotypus] in hb G 00069151 - SIB 126182/1 (hb STEPHANI 20610, ex hb Alex BRAUN, JACK misit) [typus] Basionym: Distribution: Venezuela spec nov Stephani Bonner Stephani 1911 1965 1985 - syn nov Uribe & Gradstein Uribe 2003 2008 = Frullania brasiliensis cardotii 686 655 928 Spec Hepat 4: 599 Index Hepat 5: 260 Stephani Icon Hepat Inedit [Meteriopsis] E-18, microfiche 55 fig 3790 ["Caracas, Alex Braun, ex Herb Jack"] - 3791 1071 Crypt Bryol 24 (3): 200 [syn sub Fr brasiliensis] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Raddi 1822 Steph. 1910 Typus: Zaire. "Congo Belge. Mayidi" -.3.1907, Vanderyst 500 in hb G 19719 G00045244 - n° SIB 148725/1 [holotypus]i in hb BR [isotypus] Basionym: Distribution: spec nov - syn nov Stephani Demaret Demaret & Vanden Berghen 1910 1942 1947 Bonner Vanden Berghen Stephani 1965 1976 1985 Wigginton 2002 Wigginton 2004 Wigginton 2004 = Frullania obscurifolia 686 Spec Hepat 4: 365 1066 Bull Jard Bot Etat Bruxelles 16 (2): 304 1101 Bull Jard Bot Etat Bruxelles 18: 233-235, fig 35(a-b) [Congo Belge, Cameroun] 655 Index Hepat 5: 260 185 Bull Jard Bot Nat Belg 46 (1/2): 143 [syn sub Fr cardoti] 928 Stephani Icon Hepat Inedit [Galeiloba] C- 12, microfiche 50 fig 3363 ["Africa Mayidi, leg Pycnaert"] 1012 Trop Bryol Res Rep 2: 27 [cardotii] [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [cardotii] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 233 [syn sub Fr obscurifolia, West Africa] Mitt. 1879 Page 70 caroliniana Sull. 1846 Typus: USA. North Carolina, near Welmington sd, Sullivant sn in hb G 19142 [typus] in hb MICH MICH514763 ( [isotypus] Basionym: Distribution: USA ( NC), Brazil spec nov -s yn nov Sullivant Sullivant Sullivant in G., L. & N. Schiffner 1846 1846 1847 1895 764 1491 2848 1015 Evans Schiffner in Engler & Prantl 1897 1905 237 809 Stephani Wareham Frye & Clark Bonner Wareham 1911 1940 1947 1965 1968 686 2836 204 655 2769 Lemos Michel 1980 719 Lemos Michel Yana Stephani 1983 1984 1985 306 501 928 Yuzawa Behar, Yano & Vallandro Yano Germano & Pôrto Fogel 1988 1992 1995 1996 1997 511 921 2095 299 629 Boas-Bastos & Bastos Yano & de Mello Oliviera-e-Silva & Yano Gradstein & Costa Yano 1998 1999 2000 2003 2008 702 919 989 984 1684 Winter & Schäfer-Verwimp 2020 5742 = Frullania obcordata Musci Alleghaniensis: 64 [Exsicc No 270, reprint of labels] Amer J Sci Arts, ser 2, 1: 74 [No 270] Syn Hepat: 779 [No 68b] Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 134 [syn sub Fr brunnea Austin 1846] Trans Connet Acad Arts Sci 10 (1): 33-34, pl 15 Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 134 [syn sub Fr brunnea Austin 1846] Spec Hepat 4: 668-669 "Noble Fellow" William Starlin Sullivant: 304 [syn sub Fr obcordata] Univ Wash Publ Biol 6 (5): 760 [syn sub Fr. obcordata] Index Hepat 5: 261 "Noble Fellow" William Starlin Sullivant [Reprint]: 304 [syn sub Fr obcordata] Diss, Uni Fed do Rio Grande do Sul, Porto Alegre: 99-103, pl 16, map 15, [Brasil, Rio Grande do Sul] Revta brasil Bot 6 (2): 117-119, fig 1-13 [Brazil; distinct species] J Hatt Bot Lab 56: 504 [Checklist Brazil] Stephani Icon Hepat Inedit [Diastaloba] B-36ab, microfiche 48 fig 3272 ["South Carolina, Cardot No 82"] - 3273 ["Perryville, Demetrio 1885"] J Hatt Bot Lab 64: 444-445 [syn sub Fr. caulisequa] Bol Mus Biol Mello Leitao, new ser 1: 26 [Brazil] J Hatt Bot Lab 78: 142 [annotated checklist of Brazil] Trop Bryol 12: 23 [Brazil] http://141.211.110.91/livrqury.htm [16.11.1997] - Genus=Frullania, printed 5.1.1998 11:11:38: page 1 of 3 Trop Bryol 15: 103 [Brazil, Bahia] Iheringia, ser Bot 52: 70-71 [Brasil] Boletim Inst Botanica 13: 25 [Rio de Janeiro, Brazil] Mem New York Bot Gard 87: 86 [Brazil] [syn. sub Fr caulisequa] Boletim Instituto Botanica 19: 90 [syn sub Fr caulisequa; Brazil; catalog] Frahmia 19: 12 [syn sub Fr. obcordata] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 carrii var. subcarrii S.Hatt. 1980 Typus: Indonesia. West Irian, Jayawijaya, Eipomek-Tal, 1600m 15.3.1976, H. Uiepko & W. Schultze-Motel 2370 [Mus Bot Berlin] in hb NICH [holotypus] in hb JE [isotypus] Basionym: Distribution: New Guinea (Irian Jaya), Indonesia var nov - syn nov Hattori Hattori Grolle & Piippo Hattori & Piippo Mizutani, Hasegawa & Iwatsuki = Frullania carrii 1980 1982 1984 1988 2009 489 495 143 557 2112 J Hatt Bot Lab 47: 112-114, fig 232(a-p) J Hatt Bot Lab 51: 248 Acta Bot Fennica 125: 18 [Cat. West Melanesia] Acta Bot Fennica 133: 44 Type sepecimens Hatt Bot Lab (NICH) 20: Holotype. New Guinea, West-Irian, Eipomek-Tal. 1600 m. 15 Mar. 1976. P. Hiepiko and W. Schultze-Motel 23707 (NICH 431905). Kamim. & S.Hatt. 1973 Page 71 catalinae [auct. non A.Evans 1897] Sutliffe 1932 {Guadalupe} Basionym: Distribution: Sutliffe Clark 1932 1953 = Frullania californica caucasica 1903 Bryologist 35: 42 1106 Proc California Acad Sci 4 ser, 27 (18): 602 [Guadalupe Isl] (M.Howe) A.Evans 1897 Steph. 1910 Typus: UdSSR. Georgien. prope flumen Kura, Borshon, ad ligna putrida 1877, Brotherus 238 in hb H [holotypus] in hb G 7927 [isotypus] Basionym: Frullania tenera Lindb. ex Steph. in Brotherus 1892 [non Spruce 1884] Distribution: Russia, Georgia, Ukraine nom nov - syn nov Stephani Bonner Grolle 1910 1965 1970 686 655 615 Grolle Hattori Stephani 1976 1978 1985 183 482 928 Schuster Konstantinova, Potemkin & Schijakov Bapna & Kachroo Grolle & Long 1992 1992 603 651 2000 2000 712 760 = Frullania parvistipula caudescens Spec Hepat 4: 440 Index Hepat 5: 261 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 311-312 [syn sub Fr. fragilifolia] Feddes Repert 87 (3/4): 193 [type] J Hatt Bot Lab 44: 542-545. fig 213 [syn nov sub Fr. parvistipula] Stephani Icon Hepat Inedit [Galeiloba] C-114, microfiche 51 fig 3501 ["n nov, Frullania tenera Lindb, Kaukasus Borshon prope flumen Kura, ad ligna portr, Broterus 28/7.77"] Hepat N America 5: 143 [stat nov; = Fr. eboracensis subsp. caucasica] Arctoa 1:102 [USSR] Hepaticology in India II: 178 [syn sub Fr. parvistipula] J Bryol 22: 111 [checklist of Europe and Macaronesia] & 124 [syn sub Fr. parvistipula] Steph. 1910 Steph. ex Verd. 1930 [pro syn] Typus: Philippines. Luzon, Bontoc Subprov., Banco 1500m -.1-5.1913, M. van Overbergh in hb G (hb MERRIL 2822,) Basionym: Distribution: Philippines (Luzon) Verdoorn Bonner Hattori Stephani 1930 1965 1980 1985 Tan & Engel 1986 = Frullania orientalis celebica 293 655 Ann Bryol Suppl 1: 152 Index Hepat 5: 261-262 928 Stephani Icon Hepat Inedit [Meteriopsis] E-50, microfiche 56 fig 3844 ["Philippinen, Merril missit 2822"] 4286 J. Hatt. Bot. Lab.. No. 60: 335 Hattori (1980a: 228) stated that the Philippine report could not be confirmed. Sande Lac. 1855 Sande Lac. ex Verd. 1930 [pro syn] Typus: Philippines. Sulawesi in hb SANDE LACOSTE Basionym: Distribution: Indonesia (Sulawesi) Verdoorn Bonner = Frullania gaudichaudii 1930 1965 293 655 Ann Bryol Suppl 1: 92 Index Hepat 5: 262 [nom herb] (Nees & Mont.) Nees & Mont. 1845 Page 72 cerina Steph. 1910 Typus: Brazil, Petropolis 1890, B. Rudolph 202 [102 ?] in hb G G00069189 - n° SIB 125565/1 (hb STEPHANI 20954, ex hb B) [holotypus] Basionym: Distribution: Brazil spec nov - syn nov Stephani Schiffner & Arnell Bonner Yuzawa Yano Stephani 1910 1964 1965 1983 1984 1985 Neto & Yano Yano 1985 2008 = Frullania gibbosa 686 198 655 498 501 928 Spec Hepat 4: 340-341 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 85 [Brazil] Index Hepat 5: 264 J Hatt Bot Lab 54: 109, fig. 2(a-q) [from type] [syn sub Fr. gibbosa] J Hatt Bot Lab 56: 504 [checklist Brazil] Stephani Icon Hepat Inedit [Chonanthelia] A- 63, microfiche 47 fig 3159-3160 [from type; "Petropolis Br Rudolph 1890 (Herb Berlin)"] 1343 Revta brasil Bot 8: 200 [Brazil, Serra de Sao Vicente] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr gibbosa; Brazil; catalog] Nees 1840 Page 73 cesatiana De Not. 1865 Typus: Italy. Lago Maggiore, Ispra 1839, ?? 1837 Cesati in hb RO [lectotypus] - holotype: "ad oras Verbani prope Isptam". MR3, 1837, Cesati (RO) Basionym: Distribution: Switzerland, Yugoslavia, Slowenia, Bulgaria, France (Pyrenees), Italy, Spain (Pyrenees), Canary Isl ( Fu GC Hi Pa Te )[Spain], Iran spec nov De Notaris Milde 1865 1865 Dumortier Schiffner 1874 1895 Dalla Tore & Sarnthein Schiffner in Engler & Prantl Schiffner Stephani Müller in Rabenhorst Dismier Schiffner 1904 1905 1909 1910 1915 1921 1936 Müller in Rabenhorst Bonner Stephani 1958 1965 1985 Bisang, Schumacker, Sérgio & Grolle Casas et al Dirkse, Bouman & Losada-Lima Frey, Frahm, Fischer & Lobin Casas 1989 576 Mem Reale Accad Sci Torino, ser 2, 22 (1): 383-384, pl 5 fig 24 (1-15) 2434 Bot Zeit 23(19): 155 [Frullania cesatiana DeNotaris = F. aeolotis. Nees] 580 Bull Soc Roy Bot Belgique 13: 28 [synonyms] 1015 Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 134 2694 Die Moose (Bryophyta) von Tirol, Vorarlberg und Lichtenstein: 89-90 809 Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 134 968 Österr Bot Zeitschr 59 (12): 467-472, fig 27-32 686 Spec Hepat 4: 434 1064 Krypt Flora (2ed) 6, 2: 629 [syn sub Fr riparia] 1338 Rev Bryol 47: 24-26 [first record in France] as 2105 2561 Krit Bemerk europ Leberm, ser 21: 11-12 [Hepat europ exsicc, nr. 1021 pp] 1028 Krypt Flora (3ed) 6, 2: 1252 [syn sub Fr. riparia] 655 Index Hepat 5: 263 928 Stephani Icon Hepat Inedit [Galeiloba] C- 83abc, microfiche 51 fig 3452 ["Madeira, Fritze leg, vera nach Schiffner"] - 3453 ["Italia (Verona), leg Massalongo ad rupes"] - 3454 ["SüdTirol Meran"] 276 Giorn Bot Ital 122: 258 [key to Europe & Macaronesia] & 262 1989 1993 1995 1998 11 606 996 Casas 1998 997 Casas Sim-Sim, Lloret & Grolle 1998 1999 768 726 Sim-Sim, Lloret & Grolle 1999 726 Kürschner, Papp & Akhani Pando Pando 2000 2000 2000 1216 1273 1273 Grims Sabovljec, Ganeva, Tsakiri & Stefanut Vana Geisller, Bagutti, Bisang, Müller, Schnyder, Schubiger-Bossard & Urmi Thouvenot 2001 2001 1136 1296 2002 2002 1342 1003 2003 Feuerer 2008 Casus, Brugués, Cros, Sérgio, Infante 2009 1218 Bull sci bot Centre-Ouest, ns 34: 330 [first record in France in 1921] as #2105 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm 1983 Handbook Liverw Iberien penins & Bal Isl., 143, fig 52(6-9) [Iberian Peninsula, Mallora, Menorca ] = Frullania riparia [siehe Dirkse ... 1993 = Publ No 11] Crypt Bryol Lichen 14 (1): 4 [Canary Isl: Fu, GC, Hi, Pa, Ten] Kleine Kryptogamenflora 5: 116 [key to european species] Orsis 13: 20 [checklist Spain] http://einstein.uab.es/mbrugues/A%20ChekHep.htm Orsis 13: 20 [checklist Spain] http://einstein.uab.es/mbrugues/hepati.htm Orsis 13: 20 [checklist Spain] Nova Hedwigia 68: 220 [characters of Fr. muscicola, riparia, cesatiana, azorica] Nova Hedwigia 68 (1-2): 217-224, fig 1(a-g), 2(map) [France & Spain] - holotype: "ad oras Verbani prope Isptam". MR3, 1837, Cesati (RO, holotype) Nova Hedwigia 71 (3-4): 510, 512 [new to Iran] Herbario de Criptogamia, Madrid (MA) Herbario de Criptogamia, Madrid (MA) http://161.111.170.202/herb/asp/dos.asp?conque=MAHepat&fnhe=1105&fadic=1 Moosherbar in hb LINZ [Internet] Biol Conservation 101: 79 [threatened, presenting taxonomic problems; conservation interest in Bulgaria & Slovenia] Portug Acta Biol (B) 20: 26-27 [evaluation of taxonomy & Red lists] Herzogia 15: 198-199, fig 7 [distribution & map of Switzerland and Lichtenstein] Hampe 1838 Page 74 cesatiana var. galeata Schiffn. 1936 Typus: Italy. Comersee, Argegno; an Bäumen und Kalkfelsen, ca 250m 9.7.1899, F.A. Artaria - 1020 in hb GZU (2: 1899, Artaria & 1904, Loitlesberger) >> doc in hb MA MA-Hepat 1104 >> pdf in hb E E00002578 - E00002579 - E00414525 in hb NICH 225087 (1899 Artaria) in hb EGER [syntypus] Basionym: Distribution: Italy var nov Schiffner 1936 Bonner Pocs Grolle Pando 1965 1976 1990 2000 Suanjak 2001 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania cesatiana Frullania riparia 2561 Krit Bemerk europ Leberm, ser 21: 9-12 [Hepat europ exsicc, nr. 1019, 1020, 1021 pp] 655 Index Hepat 5: 263 2686 Fol. Hist. nat. Mus. Matr. 4: 19 [types in hb EGER] in hb GZU note: [syn sub Fr. riparia by Grolle] 1273 Herbario de Criptogamia, Madrid (MA) http://161.111.170.202/herb/asp/dos.asp?conque=MAHepat&fnhe=1104&fadic=1 979 http://www.uni-graz.at/botwww/bibliothek/liverworts-types-in-gzu.html [in hb GUZ] 2112 Type specimens Hatt Bot Lab (NICH) 21-22: Syntype. Italien, Cornersee, Argegno. 250 m. An Baumen und Kalkfelsen. 9 July 1899. F. A. Artaria 1020 (NICH 225087). De Not. 1865 Hampe 1838 cesatiana var. muscicola (Steph.) Bisang, Schumacker, Sérgio & Grolle 1988 Basionym: Distribution: comb nov - syn nov Bisang, Schumacker, Sérgio & Grolle Sim-Sim & Sergio Frey, Frahm, Fischer & Lobin Sim-Sim, Lloret & Grolle Schumacker & Vana = Frullania azorica 1989 276 Giorn Bot Ital 122: 258 & 263 [key to Europe & Macaronesia] 1991 1995 1999 2000 1194 606 726 800 IX Simposio Nacional Bot Cript, Salamanca: 252 [Madeira] Kleine Kryptogamenflora 5: 116 [key to european species] Nova Hedwigia 68: 217-224 [Fr. muscicola is a valid species] Doc Stat sci Hautes-Fagnes 31: 61 [syn sub Fr azorica][key to European & Macaronesian species (1st ed)] Sim-Sim, Sérgio, Mues & Kraut 1995 cesatiana var. muscicola [auct. non (Steph.) Bisang, Schumacker, Sérgio & Grolle] Piippo 1990 Typus: Basionym: Distribution: Piippo = Frullania muscicola chilensis 1990 521 J Hatt Bot Lab 68: 44-45 [annotated catalogue of China] Steph. 1894 De Not. [nom herb] Typus: Valparaiso Frullania chilensis De Not. “Valparaiso – Ex Hb De Notar[is] – Lehmann … 53” in hb GLM (hb GOTTSCHE) 05979 - nom herb Basionym: Distribution: = Page 75 chinensis Steph. 1911 [non F. sinensis St. (1906)] Typus: China: Schensi 1894, Giraldi sn in hb G G00048043 - n° SIB 138178/55 (STEPHANI 7928, ex hb LEVIER inter 1791) [lectotypus] in hb G 7964 [syntypus] ??? In hb FH (hb VERDOORN 8679, exs 9) [isolectotypus] Basionym: Distribution: China spec nov - nom nov Stephani Verdoorn in Handel-Mazzetti Hattori Bonner Hattori Hattori Hattori & Lin Stephani 1911 1930 1944 1965 1978 1982 1985 1985 686 550 943 655 482 72 505 928 Piippo Piippo 1990 1990 521 521 = Frullania aposinensis chinlingensis Spec Hepat 4: 469 Symb Sinicae 5: 38 syn sub Fr. muscicola] Bull Tokyo Sci Mus 11: 149 [Japan, key to Japanese species] Index Hepat 5: 264 J Hatt Bot Lab 44: 540-542, fig. 211 [from lectotype, lectotypification] Bull Natn Sci Mus Tokyo, ser B (Bot), 8 (3): 99 [China] J Hatt Bot Lab 59: 131 [nom. nov., key to Chinese Frullania] Stephani Icon Hepat Inedit [Galeiloba] C-115, microfiche 51 fig 3502 [from type; "China, Schensi (Levier 1791 ex parte) Giraldi"] J Hatt Bot Lab 68: 38 [annotated catalogue of China] J Hatt Bot Lab 68: 139, annotation nr 140 about nom nov S.Hatt. & P.J.Lin 1985 Li & Zhang 1983 Typus: China. Shansi, Thai-pei Mts, Pin An Zi, 2740m 22.7.1963, Wie Zhiping 5360, 5361 in hb BOR.-OCC. INST BOT [holotypus] in hb KUN [isotypus] Basionym: Distribution: China spec nov - syn nov Li & Zhang Hattori & Lin Piippo = Frullania dilatata ciliiflora 1983 1985 1990 673 505 521 Acta Bot Yunnanica 5 (4): 387-390, fig 3(1-19) J Hatt Bot Lab 59: 163-164 [maybe conspecific with Fr. dilatata] J Hatt Bot Lab 68: 38 [annotated catalogue of China] (L.) Dumort. 1835 Gottsche ex Bonner 1965 [nom herb] Typus: Brazil. Faxina 1880, Puiggari in hb G Basionym: Distribution: Stephani 1910 686 Bonner 1965 655 = Frullania glomerata Mont 1842] cinchonale Spec Hepat 4: 384 [according BONNER, but not mentioned in STEPHANI] Index Hepat 5: 264 [error: nom herb in STEPHANI 1910: 384] (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Gottsche by Bonner 1965 [orth var] Basionym: Distribution: Bonner = Frullania cinchonae 1965 655 Index Hepat 5: 265 Gottsche 1845 Page 76 cinnamomea Carrington & Pearson 1888 Typus: Australia. New South Wales 1885, Whitelegge 66 in hb G 18200 (ex hb PEARSON, hb STEPHANI 150) [isotypus] Basionym: Distribution: Australia (NSW) spec nov - syn nov Carrington & Pearson Stephani Stephani & Watts Rodway 1888 1910 1914 1916 Bonner Hattori Hattori Stephani 1965 1979 1983 1985 Scott & Bradshaw Carthy McCarthy 1986 2003 2006 = Frullania clavata cinnamomi 1604 686 220 914 Proc Linn Soc New South Wales, ser 2, 2 (4): 1035 Spec Hepat 4: 421-422 J Proc Roy Soc New S Wales 48: 106 [Mt. Wilson] Papers Proceed Roy Soc Tasmania for 1916: 135 [syn sub Fr. proboscifora][Tasmania] 655 Index Hepat 5: 265 485 J Hatt Bot Lab 45: 325-327, fig 3(a-i) from isotype 499 J Hatt Bot Lab 54: 137 928 Stephani Icon Hepat Inedit [Galeiloba] C-198abc, microfiche 53 fig 3603 ["olim Fr falciloba T, Siber Australia, Herb ..."] - 3604 - ["olim Fr falciloba Taylor, Tasmania, Levier 3446, Weymouth 889"] - 3605 ["New South Wales (Carrington legit)"] 56 Brunnonia 8 (1): 33 [Australia] 1125 Flora Australia Suppl Ser 21: 42 [sub Fr clavata] 1385 Checklist Australian Liverworts [Australia, syn sub Fr clavata] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Glenny, Meagher & Brown 2003 [orth var] Typus: Basionym: Distribution: Glenny, Meagher & Brown = Frullania cinnamomea Frullania clavata clavaeflora 2003 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Carrington & Pearson 1888 (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Nees ex Verd. 1930 [pro syn] Basionym: Distribution: Verdoorn = Frullania nodulosa clavellata 1930 293 Ann Bryol Suppl 1: 180 [syn sub fr nodulosa] (Reinw., Blume & Nees) Nees 1845 [auct. non Mitt. 1891] Steph. 1911 {India} Basionym: Distribution: Stephani Bapna & Kachroo 1911 2000 686 712 Spec Hepat 4: 546 Hepaticology in India II: 195 [auct. non MITTEN] = Page 77 clavellata Mitt. 1891 Typus: China. Hong Kong. sd, Bowring Basionym: Distribution: spec nov - syn nov Mitten Yoshinaga Sawada 1891 1900 1909 Stephani Iishiba Verdoorn Reimers Reimers Horikawa Hattori Horikawa 1911 1912 1930 1931 1931 1934 1944 1951 Bonner Hattori 1965 1972 Hattori & Lin 1985 Stephani 1985 Piippo Yuzawa 1990 2000 Yamada & Iwatsuki 2006 = Frullania moniliata ssp. obscura 582 Trans Linn Soc London, ser 2, 3: 203 [Japan] 1685 Bot Mag Tokyo 14: 40 930 Bot Mag Tokyo 23 (no 268): 178-183 [key to Japanese species, in Japanese] 686 Spec Hepat 4: 546-547 888 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] 293 Ann Bryol Suppl 1: 80 [syn sub Fr. moniliata ssp obscura] 229 Hedwigia 71 (1/2): 36 [China] 1024 Hedwigia 70 (6): 362 [Korea] 34 J Sci Hiroshima Univ, ser B, div 2, 2: 235 [syn sub Fr. moniliata] 943 Bull Tokyo Sci Mus 11: 148 [Japan, key to Japanese species] 1698 J Sci Hiroshima Univ, serB, div 2 (Bot) 6(1): 28 [sub Fr moniliata ssp obscura] 655 Index Hepat 5: 266 536 J Hatt Bot Lab 35: 216- [syn sub Fr. tamarisci ssp. obscurae var. obsurae] 505 J Hatt Bot Lab 59: 161 [syn sub Fr. tamarisci ssp. obscura var. balansae] 928 Stephani Icon Hepat Inedit [Thyopsiella] F-134abc, microfiche 59 fig 4067 ["Japan, leg Inoué"] - 4068 ["China, Schensi, Giraldi (Levier 1796)"] - 4069 ["Japan, Kobe, C. Ford 303bis, 250"] 521 J Hatt Bot Lab 68: 38 [annotated catalogue of China] 1659 Natural Environmental Sci Res 13: 18 [Japan, syn sub Fr tamarisci ssp obscura] 1470 J Hatt Bot Lab 99: 29 [Catalog of Japan] Verd. 1930 {nördl. Indomalaya} Frullania tamarisci ssp. obscura (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata clavellifera (Reinw., Blume & Nees) Mont. 1842 Steph. 1911 Typus: Tanzania, Usambara, Amami -.7.1909, Brunnthaler sn [14 ?] in hb G G00045245 - n° SIB 148726/1 (STEPHANI 17559, hb STEPHANI 14) [typus] Basionym: Distribution: East Africa spec nov - syn nov Stephani Stephani in Brunnthaler Bonner Vanden Berghen Stephani 1911 1913 1965 1976 1985 Wigginton 2002 Wigginton 2004 = Frullania lindenbergii 686 219 655 185 928 Spec Hepat 4: 627-628 Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 728 [as "spec nov"] Index Hepat 5: 266 Bull Jard Bot Nat Belg 46 (1/2): 33-38 Stephani Icon Hepat Inedit [Diastaloba] B- 6, microfiche 48 fig 3232 [from type; "Usambara leg Brunnthaler 1909"] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Lehm. 1844 Page 78 clavellifera Steph. 1913 [nom superfl] Basionym: Distribution: Stephani in Brunnthaler = Frullania clavellifera 219 Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 728 [spec nov = nom superfl] Steph. 1911 Frullania lindenbergii clemensii 1913 Lehm. 1844 Verd. 1932 [nom herb] Typus: Malaysia/Sabah. Borneo, Tenompok "Borneo. Kinabalu, Tenompok" 1931, Clemens, J.; Clemens, M.S sn in hb L L0060922 - 992163125 > see fr2415.jpg - http://145.18.162.53:81/c8?ent=300017&rec=69081&sct=1 also iin hb B-W [Berlin] see also Fr clemensiana in hb FH - FH (hb VERDOORN 17491) [typus] - FH (hb VERDOORN 17499) [syntypus] Basionym: Distribution: = Frullania clemensiana clintonii Verd. 1932 Austin ex Eggers 1982 [pro syn] Typus: Azores. Faial 1856, J.W. Higginson in hb BUF Basionym: Distribution: Russell Eggers = Frullania microphylla 1862 1982 680 22 Proc Essex Inst 2: 134-137 [not mentioned] Crypt Bryol Lichen 3 (4): 289 [by Russell as Fr clintonii AUSTIN determinated specimen = Fr microphylla] (Gottsche) Pearson 1894 Page 79 closterantha Spruce 1884 Typus: Ecuador. Rio Verde, Trib fl. Pastasa, 1400m sd, Spruce in hb G (hb STABLER 259) in hb E E00002523 [isotypus] Basionym: Distribution: Costa Rica, Guatemala, El Salvador, Colombia, Peru, Bolivia, Ecuador spec nov Spruce Stephani Weberbauer Weberbauer Clark & Mullen Clark & Svihla 1884 1911 1911 1945 1947 1948 243 686 792 793 320 325 Clark & Svihla Herzog Soukup Bonner Winkler Stotler Stotler Winkler Cole Menzel Stephani 1950 1953 1955 1965 1967 1969 1970 1976 1983 1984 1985 337 640 171 655 3779 4761 194 5 1407 142 928 Yuzawa & Koike 1989 519 nom corr Schuster Gradstein & Pocs 1992 2021 603 5888 -var nov Gradstein 2021 5642 Trans Proc Bot Soc Edinburgh 15: 45 & 51 Spec Hepat 4: 518 Pflanzenwelt Peruan Anden: 242 [Peru] Mundo Vegetal Andes Peruanos: 572 [Peru] Bryologist 50: 86-88, fig 1-14 Bryologist 51: 24 [key to Fr. of Middle America: Guatemala, Costa Rica (Ecuador, Peru, Bolivia)] Bryologist 53: 125, fig 1-8 Rev Bryol Lichen 20: 128 [Costa Rica] Biota 1(4): 81 [Peru] Index Hepat 5: 266 Revue Bryol lichen 35: 313 [El Salvador] Nova Hedwigia 18: 444 [syn sub Fr. intumescens] The genus Frullania, separata: 444 [syn sub Fr. intumescens] Rev Bryol Lichen 42 (3): 811 [Colombia] Brenesia 21: 154 [Costa Rica] Willdenowia 14: 509 [syn of Fr. intumescens] Stephani Icon Hepat Inedit [Thyopsiella] F- 41, microfiche 57 fig 3932 ["Frullania closterantha Spruce vera (Falsch Etikette: Fr tubiflora Spruce), Andes Quitenses, Rio Verde (Pastasa) leg Spruce"] J Hatt Bot Lab 66: 357-358, fig 11 [from type MANCH], [is a proper species] Hepat N America 5: 67 Nova Hedwigia 112(1-2): 58-59 [Venezuela, = Fr. intumescens var clostrantha] Mem Ney York Bot Gard 121: 370 [syn sub Fr. intumescens var sabanetica] = Frullania intumescens var. sabanetica (Gottsche) Gradstein 2021 Frullania intumescens var. closterantha (Spruce) Gradstein & Pócs 2021 closterantha var. hylonoma Spruce 1884 Typus: Ecuador. "In montw Abitagua, Canelos" sd, Spruce Basionym: Distribution: spec nov - syn nov - syn nov Spruce Bonner Stotler Stotler = Frullania intumescens 1884 1965 1969 1970 243 655 4761 194 Trans Proc Bot Soc Edinburgh 15: 51 Index Hepat 5: 267 Nova Hedwigia 18: 444 [syn sub Fr. intumescens] The genus Frullania, separata: 444 [syn sub Fr. intumescens] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 80 coalita Steph. 1911 Typus: Colombia, Merida - ex hb W in hb G G00069144 - n° SIB 131124/1 (STEPHANI 13601, hb BOISSIER, ex hb Lipsiense) [typus] Basionym: Distribution: Bolivia, Colombia spec nov - syn nov - syn nov Stephani Stephani Bonner Stotler Stotler 1911 1916 1965 1969 1970 686 222 655 4761 194 Stephani 1985 928 Gradstein, Meneses & Arbe 2003 975 = Frullania beyrichiana cochleata Spec Hepat 4: 683 Biblioth Bot 87: 243 [Bolivia] Index Hepat 5: 267 Nova Hedwigia 18: 442 [syn nov; sub Fr. beyrichiana (L & L) Nees] The genus Frullania, separata: 442 [syn nov; sub Fr. beyrichiana (L & L) Nees] Stephani Icon Hepat Inedit [Thyopsiella] F- 42, microfiche 57 fig 3933 ["Merida, Herb Lipsiense"] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. beyrichiana {L & L] L & L] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1911 Typus: Papua New Guinea. Mt. Yule -.12.1890, [F.v. Müller, according to Verdoorn 1930: 541] in hb G G00069192 - n° SIB 124906/1 (STEPHANI 11898 (hb GEHEEB 1897, ex hb MEL 16.3.91) [holotypus] Basionym: Frullania nobilis fo. cochleata (Steph.) S.Hatt. 1978 Distribution: spec nov Stephani Verdoorn Verdoorn Bonner Hattori 1911 1930 1930 1965 1975 686 293 549 655 547 - comb nov Hattori Grolle & Piippo Hattori & Streimann 1978 1984 1985 482 143 504 Stephani 1985 928 = Frullania nobilis fo. cochleata (Steph.) S.Hatt. 1978 Frullania nobilis cocosensis Spec Hepat 4: 681 Ann Bryol Suppl 1: 58, fig 79-82 Nova Guinea 14 (4): 541 Index Hepat 5: 267 J Hatt Bot Lab 39: 297, fig 148 [comb nov; as Fr. ornithocephala var. cochleata] J Hatt Bot Lab 44: 535-536 [comb nov; as Fr. nobilis fo. cochleata] Acta Bot Fennica 125: 18 [Cat. West Melanesia] J Hatt Bot Lab 59: 111 [comb nov; as Fr. nobilis var. cochleata] [New Guinea] Stephani Icon Hepat Inedit [Galeiloba] C-116, microfiche 51 fig 35033504 ["N Guinea, Mt Yule Dec 1890, (Ferd v Müller misit"] Steph. 1894 Steph. 1911 Typus: Galapagos. Cocos Island 1902, Pittier in hb G Basionym: Distribution: Cocos Isl [Costa Rica] spec nov - syn nov - syn nov Stephani 1911 Clark & Schultz Clark 1953 1953 Bonner Stotler Stotler Stephani 1965 1969 1970 1985 = Frullania mucronata 686 Spec Hepat 4: 492 [Galapagos, Cocos Isl - Stephani considered Cocos Isl one of the Galapagos group] 354 Bryologist 56: 207-208, fig 1-14 1106 Proc California Acad Sci 4 ser, 27 (18): 662 [Cocos Isl [Costa Rica]; Stephani 1911 considered Cocos Isl one of the Galapagos group] 655 Index Hepat 5: 267 4761 Nova Hedwigia 18: 446 [syn sub Fr. mucronata] 194 The genus Frullania, separata: 446 [syn sub Fr. mucronata] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 43, microfiche 57 fig 3934 ["Galapagos, Cocos Ins (Pittier legit 1902)"] (Lehm. & Lindenb. in Lehm.) Lehm. & Lindenb. 1845 Frullania brasiliensis Raddi 1822 Page 81 cognata Lindenb. 1845 Typus: Marianas Mertens hb LE 21 Basionym: Distribution: Micronesia spec nov - syn nov Lindenberg in G., L. & N. Stephani Verdoorn Kühnemann Kühnemann Miller Kamimura Bonner Miller Miller, Whittier & Whittier Engel = Frullania nodulosa colensoana 1845 1911 1930 1937 1949 1960 1961 1965 1968 1983 1988 2850 686 293 1031 657 920 199 655 672 609 111 Syn Hepat: 433 ["in insulis Marianis legit Mertens"] Spec Hepat 4: 588 Ann Bryol Suppl 1: 177 6 179 [syn sub Fr. nodulosa] Revista Centro Estud Doct Ci Nat, Buenos Aires 1 (3): 166 [Malvinas] Lilloa 19: 354 [Falkland Isl] Bryologist 63: 121 [Micronesia] J Hatt Bot Lab 24: 57 Index Hepat 5: 268 Micronesia 4 (1): 74 Prod Flora Hepat Polynesiae: 123 Fieldiana Bot NS 25: 163 [not at Falkland Isl] (Reinw., Blume & Nees) Nees 1845 Steph. 1910 Typus: New Zealand sd, Colenso 2256 in hb G Basionym: Distribution: New Zealand spec nov - syn nov Stephani Pearson Bonner Hamlin Hattori Stephani = Frullania falciloba colliculosus 1910 1923 1965 1972 1983 1985 686 796 655 659 499 928 Spec Hepat 4: 405 Univ California Publ Bot 10 (4): 325, pl 99 (1-18) [New Zealand] Index Hepat 5: 268 Rec Dominion Mus 7(19): 261 [New Zealand] J Hatt Bot Lab 54: 146 [syn sub Fr falciloba] Stephani Icon Hepat Inedit [Galeiloba] C-257, microfiche 53 fig 3678 ["N Zelandia, Colenso 2256"] Lehm. 1844 von Konrat & Braggins in von Konrat & Pickering 2003 [nom nud] Typus: New Zealand. Subalpine: NW Nelson, Mt Arthur, Arthurs Pass Basionym: Distribution: New Zealand [South Isl: Mt Arthur NW of Nelson] Konrat & Pickering 2003 Konrat & Brennan 2004 = Frullania colliculosa colombica 1000 http://www.discoverlife.org/nh/tx/Plantae/Bryophyta/ [IDnature guide for Liverworts, Frullania, New Zealand] 1164 http://www.liverworts.org/dich_key/frul_key.shtml, 6th of 6 pages [Key to Frullania of New Zealand; Subalpine: NW Nelson, Mt Arthur, Arthurs Pass] von Konrat, Braggins, Hentschel & Heinrichs 2010 Steph. by Stotler 1969 [orth var] Basionym: Distribution: Stotler Stotler = Frullania columbica 1969 1970 4761 Nova Hedwigia 18: 436 194 The genus Frullania, separata: 436 Steph. 1924 Frullania intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 82 colonica Steph. 1910 Typus: Tanzania. Usambara,in monte Kitulwe, caten. Mlinga, 960m -.1.1908, G. Baur 5977 (or sn ?) in hb G G00045246 - n° SIB 148727/1 (STEPHANI 17304 ex Bryotheca E. LEVIER 5977) [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Demaret & Vanden Berghen Bonner Vanden Berghen Stephani 1910 1947 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania spongiosa columbica 686 1101 655 185 928 Spec Hepat 4: 373-374 Bull Jard Bot Etat Bruxelles 18: 238 [syn sub Fr squarrosa] Index Hepat 5: 268 Bull Jard Bot Nat Belg 46 (1/2): 167 & 171 [sub Fr. spongiosa] Stephani Icon Hepat Inedit [Galeiloba] C- 13, microfiche 50 fig 3364 [from type; "Africa, Usambara, leg Bauer, Levier 5977"] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 234, fig 155( C) from type [syn sub Fr spongiosa,West Africa] Steph. 1894 Steph. 1924 Typus: Colombia sd, s. coll. in hb G G00069180 - n° SIB 126164/1 (STEPHANI 13607, ex hb CARDOT 33) Basionym: Distribution: Colombia spec nov - syn nov Stephani Bonner Stotler 1924 1965 1969 - syn nov Stotler 1970 Stephani 1985 Yuzawa 1987 = Frullania intumescens commutata 687 Spec Hepat 6: 546 655 Index Hepat 5: 268 4761 Nova Hedwigia 18: 436 [as Fr. colombica, orth error in print, note in hb G "= Fr columbica] 194 The genus Frullania, separata: 436 [as Fr. colombica, orth error in print, note in hb G "= Fr columbica] 928 Stephani Icon Hepat Inedit [Thyopsiella] F-104, microfiche 58 fig 4027 ["Columbia, Herbier Cardot"] 0 note in hb G: "= Frullania intumescens (not F. brasiliensis), annotated by Y. Yuzawa, Nov 23, 1987" > see fr0149.jpg (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Nees & Hampe ex Gottsche [p. 454] 1845 [pro syn] Basionym: Distribution: Gottsche in G., L. & N. = Frullania serrata commutata 1845 2850 Syn Hepat: 454 [syn sub Fr.serrata] Gottsche 1845 Nees [p. 439] 1845 [pro syn] Typus: East Indies. Basionym: Distribution: Nees in G., L. & N. Bonner 1845 1965 2850 Syn Hepat: 439 655 Index Hepat 5: 269 [nom herb, East Indies] = Frullania tamarisci var. commutata Nees 1845 Page 83 commutata Steph. 1910 Typus: Philippines: Luzon, Benguet 1884/1885, Micholitz 3 in hb G [lectotypus] Basionym: Distribution: Indonesia (Sumatra, Java), Philippines spec nov - syn nov Stephani Verdoorn Verdoorn Svihla Bonner Stephani 1910 1929 1930 1956 1965 1985 686 246 293 380 655 928 Yuzawa 1991 525 = Frullania rio-janeirensis compsotera Spec Hepat 4: 349-350 Ann Bryol 2: 160 Ann Bryol Suppl 1: 68 [syn sub Fr. galeata] Bryologist 59: 278-279 [syn sub Fr. riojaneirensis] Index Hepat 5: 268 Stephani Icon Hepat Inedit [Chonanthelia] A- 86, microfiche 47 fig 3191 [from type; "ex planta luzonensi (Benguet) leg Micholitz No 3 1884/85"] J Hatt Bot Lab 70: 253 [lectotypification] (Raddi) Ångstr. 1876 Spruce 1884 Typus: Ecuador. Andes Quitenses, [ad fl. Pastasa superiorem] - Peru, Campana ?? sd, Spruce s.n. in hb G G00113748 - n° SIB 157630/1 (ex hb auctoris = SPRUCE) [typus] Basionym: Distribution: Ecuador spec nov - syn nov Spruce Stephani Clark Bonner Stephani 1884 1911 1955 1965 1982 Yano Yuzawa Schuster 1984 1988 1992 Winter & Schäfer-Verwimp 2020 = Frullania obcordata comptonii 243 686 372 655 928 Trans Proc Bot Soc Edinburgh 15: 55 & 57-58 Spec Hepat 4: 638 [syn sub Fr. caulisequa] Bryologist 58(3): 201 [syn sub Fr caulisequa] Index Hepat 5: 269 Stephani Icon Hepat Inedit [Diastaloba] B-28, microfiche 48 fig 32603261 ["syn: Frull compsotera Spruce Campana leg Spruce"] 501 J Hatt Bot Lab 56: 504 [checklist] - vide note in hb G 511 J Hatt Bot Lab 64: 444 603 The Hepaticae and Anthocerotae of North America. vol 5: 111 [syn sub Fr. obcordata] 5742 Frahmia 19: 12-13 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Pearson 1922 Typus: New Caledonia, no locality 1914, Compton 1730 Basionym: Distribution: spec nov - syn nov Pearson Pearson Verdoorn Verdoorn Verdoorn Bonner Miller, Whittier & Whittier = Frullania intermedia 1922 1922 1928 1930 1930 1965 1983 120 120 242 293 551 655 609 J Linn Soc Bot 46 (No 305): 32-33 J Linn Soc Bot 46 (No 305): 32-33 Rev Bryol NS 1: 122 [syn sub Fr. intermedia] Ann Bryol Suppl 1: 161 Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 269 Prod Flora Hepat Polynesiae: 123 (Reinw., Blume & Nees) Nees 1845 Page 84 concava Horikawa 1934 Typus: Taiwan. Prov. Taitô, between Shiusuie and Shucho-kyoukai 3.1.1933, Horikawa Y. 10633d in hb HIRO [holotypus] Basionym: Distribution: Taiwan spec nov - syn nov Horikawa Hattori Hattori & Noguchi Yang & Lee 1934 1960 1960 1964 Bonner Lai & Wang-Yang 1965 1976 Hattori Hattori & Lin Kuo & Chiang Piippo NN 1982 1985 1988 1990 2003 Uribe-M. 2011 = Frullania ternatensis conferta 34 J Sci Hiroshima Univ, ser B, div 2, 2: 238, pl 18(19-23) : 16 /aus 948 965 Index spec Typ herb Japon, Bryophyta: 16 961 Bot Bull Acad Sinica 5(2): 186 &193, fig 12(a-d) [close to reipari?] [Taiwan] 655 Index Hepat 5: 269-270 33 Taiwania 21 (2): 169 [Index to Taiwan] [ Steph. 1846 ex Lai & WangYang 1976 = ? written while he was 4 years old] 495 J Hatt Bot Lab 51: 228 [syn sub Fr. ternatensis] 505 J Hatt Bot Lab 59: 162 [syn sub Fr. tern. var non-append.] 948 Taiwania 33: 11 [Index of Taiwan] 521 J Hatt Bot Lab 68: 38 [annotated catalogue of China] 1397 Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html] 2767 Caldasia 33(1): 74 [type study, syn sub Fr ternatensis] Gottsche 1846 Spruce 1889 Typus: Paraguay. ad Guarapi 1880, B. Balanse 4249 in hb G [isotypus] Basionym: Distribution: Brazil, Paraguay, Argentina (San Luis, Córdoba, Entre Ríos, Buenos Aires) spec nov - syn nov Spruce Spruce Stephani Clark Bonner Stephani 1888 1889 1910 1954 1965 1985 Reiner 1988 Yuzawa 1991 = Frullania tetraptera conferta 1062 1122 686 365 655 928 Revue Bryologique 15(3) 34 [Paraguay] Bull Soc Bot France 36 (Suppl) [Congres de Botanique]: CXC [190] Spec Hepat 4: 346 Bryologist 57: 230-232, fig. 1-16 Index Hepat 5: 270 Stephani Icon Hepat Inedit [Chonanthelia] A- 65ab, microfiche 47 fig 3162 [from type; "Paraguay Balansa 4249 Original"] - 3163 ["olim Fr Sellowii Hampe ms Brasilia Sellow"] 1043 Bol Soc Argentina Bot 25 (3-4): 304-307, fig 2(a-o) [Argentina, Prov Buenos Aires] 525 J Hatt Bot Lab 70: 226 [syn sub Fr. tetraptera] Nees & Mont. 1838 Spruce 1888 [nom nud] Basionym: Distribution: Spruce Reiner = Frullania conferta 1888 1988 1062 Rev Bryol 15 (3): 34 [sp nov] [No 4249] 1043 Bol Soc Argentina Bot 25 (3-4): 304 [nom nud] Spruce 1889 Frullania tetraptera Nees & Mont. 1838 Page 85 confertiramea Steph. 1911 Typus: Brazil, Bahia sd, Lesquereux 23 pp in hb G G00066981 - n° SIB 125557/1 - G13619 (ex hb NEES in STR) [typus] Basionym: Distribution: Brazil spec nov - syn nov Stephani Bonner Stotler Yano Stephani 1911 1965 1968 1984 1985 Yuzawa & Koike Gradstein & Costa Yano 1989 2003 2008 = Frullania exilis Spec Hepat 4: 511 [= sgen Frullania] Index Hepat 5: 270 note in hb G: "= subgen Diastaloba" 501 J Hatt Bot Lab 56: 504 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 45, microfiche 57 fig 3937 ["Bahia, leg Lesquereux"] 519 J Hatt Bot Lab 66: 344-345, fig 1 [syn sub Fr exilis] 984 Mem New York Bot Gard 87: 83 [Brazil] [as syn nov sub Fr apiculata] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr excilis; Brazil; catalog] Taylor 1846 Frullania apiculata confluens 686 655 (Reinw., Blume & Nees) Nees 1845 J.B.Jack ex Bonner 1965 [nom herb] {Argentinia} Typus: Argentinia. Cordillerae Argentinenses tropicae 1873, P.C. Lorentz sn in hb G sub Fr. tetraptera (ex hb JACK) Basionym: Distribution: Argentina Bonner 1965 655 Index Hepat 5: 270 1911 1925 1965 1969 1970 1985 686 1586 655 4761 194 928 Spec Hepat 4: 517 Arquivos de botânica do Estado de Sao Paulo 1 (2): 47 [Brazil] Index Hepat 5: 270-271 Nova Hedwigia 18: 452-453 The genus Frullania, separata: 452-453 Stephani Icon Hepat Inedit [Thyopsiella] F- 46, microfiche 57 fig 3938 ["Brasilia, Ule 386"] - 3939 = confusa Typus: Brazil. Steph. 1911 sd, Ule 386 in hb G G00114947 - n° SIB 170397/1 Basionym: Distribution: spec nov - syn nov - syn nov Stephani Herzog Bonner Stotler Stotler Stephani = Frullania montagnei congesta Gottsche 1845 Hooker 1855 {Tehawera, leg Colenso & Auckland, leg Sinclair} Basionym: Distribution: Hooker Konrat, Braggins, Asakawa & Toyota = Frullania rostrata 1855 2006 {Tehawera, leg Colenso & Auckland, leg Sinclair} 1465 J H B L 100: 560 (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Page 86 congesta Mitt. in Hooker 1859 {Tasmania} Basionym: Distribution: Mitten in Hooker Konrat, Braggins, Asakawa & Toyota = Frullania rostrata conistipula 1859 2006 {Tasmania, leg Gunn & leg Archer} 1465 J H B L 100: 560 (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Steph. 1910 Typus: Japan. Prov. Aomori 20.5.1898, Faurie 135 in hb G G00069157 - n° SIB 124199/1 - G22268 [holotypus] [synonyms see fr0908.pdf Basionym: Distribution: Japan spec nov comb nov - syn nov Stephani Iishiba Hattori Kamimura Kamimura Bonner Choe 1910 1912 1952 1952 1961 1965 1980 Stephani 1985 Bapna & Kachroo Yuzawa 2000 2001 Yamada & Iwatsuki 2006 686 888 529 873 199 655 1171 Spec Hepat 4: 399 [as constipula; errata on p. 824] Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] J Hatt Bot Lab 8: 29 [related to muscicola] Contrib Fl Hepat Shikokuensem, Kochi: 133 J Hatt Bot Lab 24: 25 [syn nov sub Fr muscicola var inuena] Index Hepat 5: 271 Illust Fl & Fauna Korea 24: 734 [syn nov sub Fr muscicola var inuenia; South Korea] 928 Stephani Icon Hepat Inedit [Galeiloba] C-230, microfiche 53 fig 3648 ["Japan, Aomori, 205. 1898, Faurie No 135"] 712 Hepaticology in India II: 175 [syn nov sub Fr. muscicola var. inuena] 1660 Natural Environmental Sci Res 14: 26 [Japan, syn sub Fr muscicola var inuena] 1470 J Hatt Bot Lab 99: 29 [Catalog of Japan] = Frullania muscicola var. inuena (Steph.) Kamim. 1961 Frullania muscicola conjugata Steph. 1894 Steph. 1911 Typus: Tanzania. Usambara, Nderema Urwald 24.2.1893, Holst 2282 in hb G G00045646 - n° SIB 148728/1 - G16811 [typus] Basionym: Distribution: spec nov - syn nov Stephani Vanden Berghen Bonner Vanden Berghen Stephani 1911 1953 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania serrata 686 446 655 185 928 Spec Hepat 4: 483 Svensk Bot Tidskr 47 (2): 267 Index Hepat 5: 271 Bull Jard Bot Nat Belg 46 (1/2): 111 Stephani Icon Hepat Inedit [Thyopsiella] F- 19, microfiche 56 fig 3896 ["Usambara, Holst 2282, Nderema Urwald 24/2. 93"] - 3897 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 221 [syn sub Fr serrata, West Africa] Gottsche 1845 Page 87 conjuncta J.B.Jack ex Bonner 1965 [nom herb] {Argentinia} Typus: Argentinia. Siambon bei Tucuman 1873, P.G. Lorentz sn in hb G sub Fr. tetraptera (ex hb JACK) Basionym: Distribution: Argentina Bonner 1965 655 Index Hepat 5: 271 = connata Lindenb. & Gottsche 1847 Typus: Mexico. Veracruz, Hacienda de Fovo sd, Liebmann in hb C in hb NY Basionym: Distribution: Mexico, Guadeloupe, Bolivia spec nov - syn nov - syn nov Lindenberg & Gottsche in G., L. & N. Gottsche Gottsche Stephani Stephani Pagán Bonner Stotler Stotler Stephani 1847 2848 Syn Hepat: 783-784 1863 1867 1911 1916 1942 1965 1969 1970 1985 103 104 686 222 3152 655 4761 194 928 Fulford & Sharp Gradstein, Meneses & Arbe 1990 2003 105 975 = Frullania breuteliana constipula Separata of Mex. Leverm.: 251 K Danske Vid Selks Skrifter, ser 5, 6: 347-348 [Mexico] Spec Hepat 4: 521-522 Biblioth Bot 87: 244 [Bolivia] Bryologist 45(4): 108 [Guadeloupe] Index Hepat 5: 271-272 Nova Hedwigia 18: 453-454 [syn nov; sub Fr breuteliana] The genus Frullania, separata: 453-454 [syn nov; sub Fr breuteliana] Stephani Icon Hepat Inedit [Thyopsiella] F- 47, microfiche 57 fig 3940 ["Mexico, leg Leibold, No 1523"] - 3941 Mem NY Bot Gard 63: 41 [Mexico] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr breuteliana] Gottsche 1845 Steph. by Steph. 1910 [orth var] Basionym: Distribution: Beauverd in Stephani = Frullania conistipula 1924 909 Spec Hepat 6: 570 Steph. 1910 Frullania muscicola var. inuena (Steph.) Kamim. 1961 Frullania muscicola contracta Steph. 1894 Steph. in Paris 1910 [nom nud] Basionym: Distribution: Paris = Frullania contracta 1910 911 Rev Bryol 37 (1): 41 [New Caledonia: Isle of Pines] Steph. 1911 Page 88 controversa Beauverd ex Steph. 1924 Basionym: Frullania herzogiana Distribution: Bolivia nom nov - syn nov Beauverd ex Stephani Bonner Uribe = Frullania convoluta convoluta Steph. ex Herzog 1916 [hom illeg] [non Steph. 1911] 1924 1965 2008 687 Spec Hepat 6: 549 655 Index Hepat 5: 272 1513 Caldasia 30(1): 66 [syn sub Fr convoluta] [Monography of subgenus Meteoriopsis] Lindenb. & Hampe 1851 fo. intermedia Herzog 1953 [nom inval] {Costa Rica} Typus: Costa Rica Basionym: Distribution: Costa Rica Herzog 1953 640 Rev Bryol Lichen 20: 129 [Costa Rica] = cookei A.Evans ex Steph. 1910 Typus: Hawaii Isl. Oahu 1898, C.M. Cooke Jr sn in hb G (ex hb A.W. Evans, Oahu, 1898, Cooke sn) [holotypus] in hb BISH [isotypus] among Fr oahuensis 8 specimens which were collected by Cooke in 1897 & 1898, Identified by Verdoorn as Fr oahuensis Basionym: Distribution: Hawaii spec nov - syn nov Evans ex Stephani Verdoorn Miller Miller Bonner Miller, Whittier & Whittier Stephani 1910 1930 1953 1963 1965 1983 1985 So & Wang 2006 = Frullania oahuensis 686 551 349 834 655 609 928 Spec Hepat 4: 437-438 "Hab. Hawai, Oahu" Nederl Kruidk Arch Jrg 1930 (2): 158-159 [syn nov sub Fr. oahuensis] Bryologist 56: 44 [sub Fr. oahuensis] Ark Bot, ser 2, 5: 518 [sub Fr. oahuensis] Index Hepat 5: 273 Prod Flora Hepat Polynesiae: 124 Stephani Icon Hepat Inedit [Galeiloba] C-119, microfiche 51 fig 3509 [from type; "Oahu, Cooke 1898"] 1466 J H B L 100: 426 [sub Fr oahuensis] Hampe ex Gottsche, Lindenb. & Nees 1843 Page 89 cordata Mitt. 1863 Typus: Cameroon. Cameroons Mountain, 8000ft sd, Mann in hb NY [holotypus] Basionym: Distribution: Cameroon spec nov - syn nov Mitten Stephani in Engler 1863 1895 Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania serrata cordifolia 253 J Linn Soc Bot 7: 168-169 1481 Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [Kilimandscharo, Kamerungebiet] 686 Spec Hepat 4: 477-478 655 Index Hepat 5: 273 185 Bull Jard Bot Nat Belg 46 (1/2): 111 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 2abc, microfiche 56 fig 3867 ["S Thomé 43"] - 3868 ["San Thomé No 19, Moller legit"] - ["St Thomé, Moller legit, inter 50"] 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 221 [syn sub Fr serrata, West Africa] Gottsche 1845 Steph. 1911 Typus: S. Thomé sd, Fr. Quintas sn in hb G G00067924 - n° SIB 140989/3 - G17306 [Typus] "mala" in hb G G00045247 - n° SIB 148729/1 - G17305 [Typus] "optima" Basionym: Distribution: W Africa spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania angulata cordistipula 686 655 185 928 Spec Hepat 4: 683-684 Index Hepat 5: 273 Bull Jard Bot Nat Belg 46 (1/2): 71 [syn sub Fr. angulata] Stephani Icon Hepat Inedit [Thyopsiella] F- 3, microfiche 56 fig 3870 ["San Thomé, Franc Quintas"] - 3871 1012 Trop Bryol Res Rep 2: 27 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Mitt. 1863 Nees 1838 [nom inval; Art. 32.1(c)] Basionym: Distribution: Nees = Frullania cordistipula 1838 261 Naturg Europ Leberm 3: 239 [note 1 to Fr. hispanicum] (Reinw., Blume & Nees) Nees 1845 cordistipula var. foliis submucronatus Nees in Mont. 1855 [pro syn] Typus: in sched, ad celeb Nees ab Esenbeck missa Basionym: Distribution: nom herb Gottsche in G., L. & N. Montagne = Frullania montagnei 1845 1855 2850 Syn Hepat: 456 475 Ann Sci Nat Bot, ser 4, 3: 320 Gottsche 1845 Page 90 cordistipula var. mucronata Nees ex Gottsche 1845 [pro syn] Basionym: Distribution: Gottsche in G., L. & N. = Frullania serrata 1845 2850 Syn Hepat: 454 [syn sub Fr serrata] Gottsche 1845 cordistipula var. scutatum (Nees in Martius) Nees in Mont. in Alcide d'Orbigny 1839 {Brazil} [nom nud] Basionym: Distribution: Nees in Montagne in Alcide d'Orbigny 1839 257 d'Orbigny, Voyage l'Amerique Merid 7 (2): 68 = cordistipula var. taitica Gottsche ex Bonner 1965 [nom herb] Typus: Tahiti sd, Vesco in hb BESCHERELLE, hb PC, in hb G [sub Fr. tahitensis] Basionym: Distribution: Bonner 1965 655 Index Hepat 5: 274 = Frullania cordistipula var. tahitica Gottsche in Bescherelle 1898 cordistipula var. β mutica Gottsche, Lindenb. & Nees 1845 Typus: Java Basionym: Distribution: Indonesia (Java) Nees in G., L. & N. Bonner 1845 1965 2850 Syn Hepat: 454 655 Index Hepat 5: 274 = Frullania cordistipula var. mutica (Gottsche, Lindenb. & Nees) Schiffn. 1898 cordistipula ssp. subapiculata S.Hatt. 1972 Typus: Papua New Guinea. Simbu, Mt. Wilhelm, Lake Aunde, 11600ft 1967, Schuster 67-6522 in hb NICH [holotypus] in hb JE [isotypus] in hb MASS [isotypus] Basionym: Distribution: ssp nov - syn nov Hattori Hattori Hattori Hattori Grolle & Piippo Hattori & Piippo Mizutani, Hasegawa & Iwatsuki = Frullania falsipapuana Frullania papuana 1972 1973 1974 1982 1984 1986 2009 539 542 546 495 143 557 2112 J Hatt Bot Lab 36: 421-423, fig 7 J Hatt Bot Lab 37: 149 [New Guinea] J Hatt Bot Lab 38: 262 [syn sub Fr. reimersii var wilhelmensis] J Hatt Bot Lab 51: 217 [= Fr. falsipapuana] Acta Bot Fennica 125: 18 [Cat. West Melanesia] Acta Bot Fennica 133: 32 [= Fr. papuana] Type sepecimens Hatt Bot Lab (NICH) 22: Holotype. New Guinea, Mt. Wilhelm. 11600 ft. 2 June 1967. R. M. Schuster 67-6522 (NICH 309928). S.Hatt. 1982 [nom inval] Verd. 1930 Page 91 coreana Steph. by Sawada 1909 [orth var] Basionym: Distribution: Sawada 1909 = Frullania koreana Bot Mag Tokyo 23 (no 268): 178-183, fig 20, 20' [key to Japanese species, in Japanese] Steph. 1911 Frullania hamatiloba cornigera 930 Steph. 1910 Mitt. in Hooker 1855 Typus: New Zealand. Northern and Middle Islands, Bay of Islands / Port Cooper Hooker / sd, Lyall sd, Basionym: Distribution: New Zealand spec nov - comb nov Mitten in Hooker 1855 Hooker Stephani Bonner Hamlin Renner 1867 1912 1965 1972 2016 = Goebeliella cornigera cornuta 560 Antarctic voyage, II. Fl Novae-Zelandiae 2: 163, pl 104(8) [New Zealand] 1154 Handbook New Zealand Flora: 535 [New Zealand, N & Middle Islands] 1452 Hedwigia 51: 62 [syn sub Goebeliella] 655 Index Hepat 5: 274-275 659 Rec Dominion Mus 7(19): 261 [New Zealand] 4267 Telopeia 19: 79-99 (Mitt.) Steph. 1911 Steph. ex Paris 1906 [nom nud] Basionym: Distribution: Paris Paris 1906 1910 = Frullania cornuta cornuta 676 Rev Bryol 33 (2): 28 [New Caledonia] 1455 Rev Bryol 37 (6): 130 [New Caledonia: Dogny] Steph. 1911 fo. undulosa S.Hatt. 1984 Typus: New Caledonia. Mont Dzumac, N of Nouméa, 600-850m in hb NICH Basionym: Distribution: fo nov - syn nov Hattori 1984 Hattori 1986 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania cornuta costaricensis Typus: Costarica 503 J Hatt Bot Lab 57: 411 [fo nov] 506 J Hatt Bot Lab 60: 210 [syn sub Fr. cornuta] 2112 Type sepecimens Hatt Bot Lab (NICH) 22: Holotype. New Caledonia, Mont Dzumae. 600-850 m. On bush. 16 July 1982. Z. Iwatsuki 15055 (NICH 386884). Steph. 1911 Steph. ex Steph. 1985 [nom herb] {Costa Rica} 1912, C Brade Basionym: Distribution: Stephani 1985 928 Stephani Icon Hepat Inedit [Galeiloba] C- 55, microfiche 50 fig 3410 ["leg C Brade, Costarica 1912 (com M. Goldschmidt, Gera?"] = Page 92 costaricensis Steph. 1911 Typus: Costa Rica. 1888, H. Pittier 5 in hb G G00067105 - n° SIB 125488/1 - G13612 (ex hb BESCHERELLE) Basionym: Distribution: spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani = Frullania macrocephala crahyrtada 1911 1965 1969 1970 1985 686 655 4761 194 928 Spec Hepat 4: 512-513 Index Hepat 5: 275 Nova Hedwigia 18: 439 [syn sub Fr. macrocephala] The genus Frullania, separata: 439 [syn sub Fr. macrocephala] Stephani Icon Hepat Inedit [Thyopsiella] F- 48, microfiche 57 fig 3942 ["Costarica, Pittier legit 1888, Bescherelle missit"] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Spruce [ nom. herb.] Typus: Cuba, Guantánamo, La Mina, Monte Ronge, 320-370m M. Caluff 15.8.1985, det. D. Reyes 1989 hb IBUNAM: MEXU:BR18567 Basionym: Distribution: Reyes 1989 6099 http://unibio.unam.mx/collections/specimens/urn/IBUNAM:MEXU:BR18 567 = cranialis (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Basionym: Jungermannia cranialis Distribution: spec nov Hooker & Taylor 1845 Taylor ex G,L,N Hooker Weymouth Stephani Rodway Bibby 1845 1867 1903 1910 1916 1955 2850 1154 2090 686 914 868 Hamlin Hattori Hattori Hattori Stephani 1972 1979 1982 1983 1985 659 486 131 499 928 Scott & Bradshaw Carthy Glenny, Meagher & Brown 1986 2003 2004 56 1125 1126 Glenny, Meagher & Brown 2004 1127 Pocs 2008 1812 = Frullania probosciphora Syn Hepat: 419 [in Freto Regis Georgii, unde attulit Cunningham] Handbook New Zealand Flora: 536 [New Zealand, Middle Island] Pap Proc Roy Soc Tasmania for 1902: 125 [Tasmania] Spec Hepat 4: 415 Papers Proceed Roy Soc Tasmania for 1916: 136 [Tasmania] Victorian Naturalist 72: 101-103, fig a-f [ from Steph Icones Hepaticae] [type of Fr. cranialis = Fr. dilatata from Europe] Rec Dominion Mus 7(19): 261 [New Zealand] J Hatt Bot Lab 46: 128-129, fig 33 J Jap Bot 57 ( 9): 257 [syn sub Fr. dilatata] J Hatt Bot Lab 54: 137-140, fig 52 [from type] Stephani Icon Hepat Inedit [Galeiloba] C-199, microfiche 53 fig 3606 ["Original!, Australia, King George Sound, leg Cunningham"] Brunnonia 8 (1): 33 [Australia] Flora Australia Suppl Ser 21: 110 [incorrectly reported from Australia] NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Fieldiana: Botany, New Series 47: 151 [ Taylor 1846 Page 93 cranialis [auct. non (Hooker & Taylor) Taylor in G., L. & N 1845] E.A.Hodgs. 1949 {Australia} Basionym: Distribution: - syn nov Hodgson Bibby 1949 1955 Hattori 1983 Glenny 1998 = Frullania reptans Trans Roy Soc New Zealand 77 (3): 377 [New Zealand] Victorian Naturalist 72: 101-103, fig a-f [ from Steph Icones Hepaticae] [type of Fr. cranialis = Fr. dilatata; conspecific with Fr. probosciphora] 499 J Hatt Bot Lab 54: 139 [may be Fr reptans; Fr reptans = geographical race of Fr probosciphora] 1412 Tuhinga 10: 121-122 [revised checklist of New Zealand] Mitt. in Hooker 1855 Frullania probosciphora crassa 203 868 Taylor 1846 Herzog 1923 Typus: Chile: Valdivia, "an Bäumen im Stadtpark von Valdivia." -.12.1911, T. Herzog sn in hb JE [holotypus?] Basionym: Distribution: Chile, Juan Fernandez Isl [Chile] spec nov Herzog Reimers Herzog Herzog in Skottsberg 1923 1926 1938 1942 - syn nov Herzog Bonner Yuzawa Müller & Zündorf 1954 1965 1991 2007 = Frullania ecklonii crassicaulis 112 113 723 722 Hedwigia 64: 7, fig. 3 [Chile, Valdivia] Hedwigia 66: 46-47 [Chile] Arch Esc Farm Fac Ci Med Cordoba 7: 28 [Chile] Nat Hist Juan Fernandez & Easter Island, 2: 738 [Juan Fernandez: Masatierra = Isla Robinson Crusoe] 202 Rev Bryol Lichen 23: 56 [syn sub Fr. eckloni] 655 Index Hepat 5: 275 525 J Hatt Bot Lab 70: 259 [syn. nov. syn sub Fr. eckloni] 1442 http://www2.unijena.de/biologie/spezbot/herbar/bryotypes/Uebersicht%20F.htm (Spreng.) Gottsche, Lindenb. & Nees 1845 Steph. 1911 Typus: Indonesia. Sumatra. sd, Nyman s.n. in hb G G00069210 - n° SIB 124880/1 [typus] Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Stephani = Frullania hasskarliana 1911 1929 1930 1965 1985 686 246 293 655 928 Spec Hepat 4: 552-553 Ann Bryol 2: 161 Ann Bryol Suppl 1: 88 [syn sub Fr. hasskarliana] Index Hepat 5: 275 Stephani Icon Hepat Inedit [Thyopsiella] F-165, microfiche 59 fig 4106 ["Sumatra, Nyman"] Lindenb. 1845 Page 94 crassiuscula Taylor ex Nees 1845 [pro syn] Typus: Guyana. Demerara sd, Cambell in hb TAYLOR & GOURLIE Basionym: Distribution: Guyana - syn nov Raddi Nees in G., L. & N. Stephani 1829 1845 1910 Bonner 1965 = Frullania gibbosa crawfordii Typus: 2875 Mem Mat Fisica Soc Ital Sci, Modena 20: pl. 1 (fig. 1) dichotoma 2850 Syn Hepat: 411 686 Spec Hepat 4: 344 [syn sub Fr. gibbosa] according BONNER 1965, but not mentioned in STEPHANI 655 Index Hepat 5: 276 Nees 1840 [auct. non STEPH 1894] Steph. & Watts 1914 {Lord Howe Isl} Watts 14 (det Steph), Watts sn (MEL 61530) Basionym: Distribution: auct - syn nov Stephani & Watts Hattori = Frullania fulfordiae creberrima 1914 1987 220 400 J Proc Roy Soc New S Wales 48: 106 [Lord Howe Isl] Bryologist 90 (4): 366 S.Hatt. 1987 Steph. 1911 Typus: Madagascar. Betsileo sd, E. Villaume s.n. in hb G G00045248 - n° SIB 148730/1 - G12980 [typus] Basionym: Distribution: Madagascar spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania loricata crenatiflora 686 655 185 928 Spec Hepat 4: 489 Index hepat 5: 276 Bull Jard Bot Nat Belg 46 (1/2): 92 & 97 Stephani Icon Hepat Inedit [Thyopsiella] F- 18, microfiche 56 fig 3895 ["Madagascar, Betsileo, leg Villaume"] 250 Bryophyt Bibl 48: 67 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Pearson 1891 Pearson by Vanden Berghen 1976 [orth var] Basionym: Distribution: Vanden Berghen = Frullania crenuliflora Frullania trinervis crentiloba 1976 185 Bull Jard Bot Nat Belg 46 (1/2): 13 [orth error] Pearson 1924 (Lehm.) Drège 1843 Steph. 1924 by Bapna & Kachroo 2000 [orth var] Basionym: Distribution: Bapna & Kachroo = Frullania crenatiloba 2000 712 Hepaticology in India II: 191 Steph. 1924 Page 95 crenuliflora Pearson 1924 Typus: Africa. Mt. Elgon, 2000m -.5.1920, Lindblom Basionym: Distribution: Kenya, Uganda spec nov Pearson Bonner Vanden Berghen 1924 1965 1976 795 655 185 Wigginton 2002 1012 Wigginton 2002 1012 Wigginton Wigginton Wigginton Bernal, Gradstein & Celis 2004 2004 2004 2016 1312 1312 1172 4290 = Frullania trinervis crenulifolia Arkiv Botanik 19 (5): 8-9, pl 6(1-14) [Mt. Elgon in Kenia/Uganda] Index Hepat 5: 277 Bull Jard Bot Nat Belg 46 (1/2): 13 [orth error as Fr. crenatiflora; syn sub Fr. trinervis] Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Trop Bryol Res Rep 5: 29 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 227 [syn sub Fr trinervis, West Africa] Catálogo de plantas y líquenes de Colombia. Vol. I. [dubious species] (Lehm.) Drège 1843 J.B.Jack & Steph. 1892 Typus: Colombia. Prov. Antioquia, Nova Granata, prov. Antioquia, Paramo de Sondos, 10.000ft 1872, G. Wallis 6 - Spruce legit ? in hb G G00069161 - n° SIB 124205/1 (ex hb JACK) Basionym: Distribution: Colombia, Ecuador, Argentina spec nov - syn nov Jack & Stephani Stephani Arnell Bonner Fulford Gradstein & Hekking Stephani 1892 1911 1962 1965 1967 1979 1985 Uribe & Gradstein Uribe & Rangel-Ch Gradstein 1998 2000 2021 = Frullania subtilissima crinoidea 1089 686 449 655 1196 483 928 Hedwigia 31: 13, 14-15 [Peru, Neu-Granada] Spec Hepat 4: 631-632 Svensk Bot Tidskr 56 (2): 341 [Ecuador] Index Hepat 5: 277 Acta Bot Venezuelica 2(5-8): 93-94 [Argentina, Auyan-tepui] J Hatt Bot Lab 45: 112 [Cat Colombia] Stephani Icon Hepat Inedit [Diastaloba] B-17, microfiche 48 fig 3246 ["N Granada (Spruce legit) Wallis"] 707 Bryophyt Bibl 53: 36 [Catalogue Colombia] 1090 Colombia, diversidad biótica III, Bogota: 445 [Colombia] 5642 Mem NY Bot Gard 121: 380 [syn sub Fr subtilissima] (Nees ex Mont.) Lindenb. 1845 Steph. ex Clark 1953 Basionym: Distribution: Clark = Frullania crinoidea Frullania atrata 1953 356 Bryologist 56: 213-216, fig 1-29 [as Fr. crinoidea STEPH; difference to Fr. peruviana] Spruce ex Steph. in Steph. 1911 (Sw.) Nees ex Mont. 1839 Page 96 crinoidea Spruce ex Steph. in Steph. 1911 Typus: Peru. Mt Guayrapurina Spruce sn (sub Fr peruviana Spruce, non Gottsche) in hb G G20613 [holotypus], in hb MANCH 14041, 14040 [isotypus] in hb E E00293723 - E00293724 - E00002524 [isotypus] Basionym: Frullania peruviana [auct. non Gottsche 1846] Spruce 1884 Distribution: Guatemala, Colombia, Peru, Ecuador spec nov - syn nov Stephani Herzog Clark 1911 1952 1953 Herzog Bonner Bonner Winkler Gradstein & Hekking Menzel Stephani 1955 1965 1965 1976 1979 1984 1985 Uribe & Gradstein Uribe & Gradstein Uribe 1998 2003 2008 = Frullania atrata crispata 686 444 356 Spec Hepat 4: 604 [Spruce, Hep Amaz And exsicc] Svensk Bot Tidskr 46 (1): 86 [Ecuador] Bryologist 56: 213-216 [as Fr crinoidea Spruce in Stephani 1911; Peru, Ecuador, Guatemala] 107 Feddes Repert 57 (1/2): 198 [Peru] 655 Index Hepat 5: 277 655 Index Hepat 5: 277 5 Rev Bryol Lichen 42 (3): 811 [Colombia] 483 J Hatt Bot Lab 45: 112 [Cat Colombia] 142 Willdenowia 14: 489 [Peru] 928 Stephani Icon Hepat Inedit [Meteriopsis] E-19, microfiche 55 fig 3792 ["(Fr peruviana Hep Am), Mte Guayrapurina, Peruvia, ab ipso auctore"] - 3793 707 Bryophyt Bibl 53: 36 [Catalogue Colombia] 1071 Crypt Bryol 24 (3): 199 [syn sub Fr atrata] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] (Sw.) Nees ex Mont. 1839 Steph. 1910 Typus: Australia: Norfolk Island 1884, Robinson sn in hb G G00069162 - n° SIB 125605/1 - G18465 (ex hb MEL, ded. Dr. Karl Müller, Halle, hb JACK) [lectotypus] in hb G G00066988 - n° SIB 125618/1 - G18466 (F. Müller misit) [syntypus] Basionym: Distribution: Norfolk Isl spec nov - syn nov Stephani Bonner Hattori Miller, Whittier & Whittier Stephani 1910 1965 1979 1983 1985 686 655 485 609 928 Hattori Yuzawa 1988 1991 516 525 = Frullania obscura crispistipula {Bolivia} Spec Hepat 4: 355 Index Hepat 5: 277 [lectotypification] J Hatt Bot Lab 45: 330-332, fig. 6(a-g) from type Prod Flora Hepat Polynesiae: 124 Stephani Icon Hepat Inedit [Chonanthelia] A-104, microfiche 48 fig 3219 ["Norfolk Island Robinson F. Müller missit"] J Hatt Bot Lab 65: 414 [= geographical race of Fr. arecae] J Hatt Bot Lab 70: 264 syn sub Fr. arecae] (Sw.) Mont. 1840 [auct. non Steph. 1894 nec Steph. ex Verd. 1930] Steph. ex Herzog 1916 Basionym: Distribution: Bolivia Stephani = Frullania arecae 1916 222 Biblioth Bot 87: 239 [Bolivia] (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura (Sw.) Mont. 1840 Page 97 crispistipula Steph. ex Verd. 1930 [pro syn] {New Guinea} Basionym: Distribution: Verdoorn 1930 = Frullania novoguineensis crispistipula 549 Nova Guinea 14 (4): 541 Schiffn. 1890 Steph. 1894 {Madagascar} Typus: Madagascra - Madagascar, Andrangoloaka 1891, Sikora 90 pp - Mai 1892, Sikora 333 [hb PC] In hb G G00045249 - n° SIB 148731/1 - G12981 (hb STEPHANI 204, ex hb Wien, leg. Sikora 1891 inter 90,) [holotypus] in hb G in addition 3 specimens G12983, G12988, G12987 [types] >vide fr0307.doc in hb PC PC0102172 (hb F. RENAULD) [isotype] Basionym: Distribution: Abyssinia, Usambara, Madagascar, Mauritius spec nov - syn nov Stephani Stephani in Levier Stephani Pearson Demaret & Vanden Berghen Arnell Bonner Stephani 1894 1901 1910 1924 1948 1963 1965 1985 228 1456 686 795 59 822 655 928 Grolle Wigginton 1995 2002 250 1012 Gradstein, Meneses & Arbe Wigginton Wigginton 2003 2004 2004 975 1312 1172 Feuerer 2008 2517 = Frullania obscura cristicrispula Hedwigia 33: 137 Rev Bryol 28 (5): 94 [Madagascar] Spec Hepat 4: 321-322 [as spec nov] Arkiv Botanik 19 (5): 8, 9 [Mt. Elgon in Kenia/Uganda] Bull Jard Bot Etat Bruxelles 19 (1): 77 [syn nov sub Fr. arecae] Hepat S Africa: 136 [as "crististipula" syn sub Fr. arecae] Index Hepat 5: 278 Stephani Icon Hepat Inedit [Chonanthelia] A- 2, microfiche 46 fig 3064-3065 [from type; "Madagascar Sikora inter 90"] Bryophyt Bibl 48: 63 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. arecae] Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 224 [as Fr cristicrispula, syn sub Fr arecae, West Africa] in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm (Sw.) Mont. 1840 Steph. by Wigginton 2004 [orth var] Basionym: Distribution: Wigginton = Frullania crispistipula Frullania obscura crististipula 2004 1172 Sript Bot Belg 30: 224 [orth var of Fr crispistipula, syn sub Fr arecae, West Africa] Steph. 1894 {Madagascar} (Sw.) Mont. 1840 Steph. by S.W.Arnell 1963 [orth var] Basionym: Distribution: Arnell Wigginton = Frullania crispistipula Frullania obscura 1963 2004 822 Hepat S Africa: 136 1312 Trop Bryol Res Rep 5: 96 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Steph. 1894 {Madagascar} (Sw.) Mont. 1840 Page 98 cruegeri Steph. 1911 Typus: Trinidad sd, Crüger sn in hb G G00067499 - n° SIB 130588/1 - G13615 (ex hb GOTTSCHE) [typus] Basionym: Distribution: Trinidad spec nov Stephani Bonner Stephani 1911 1965 1985 686 655 928 - syn nov Yuzawa & Koike 1989 519 = Frullania exilis Taylor 1846 Frullania apiculata cubensis Spec Hepat 4: 516 [as "crügeri"] Index Hepat 5: 278 Stephani Icon Hepat Inedit [Thyopsiella] F- 49, microfiche 57 fig 3943 ["Frullania crügeri St nsp, Trinidad, Crüger, Herb Gottsche (ohne Namen)"] J Hatt Bot Lab 66: 345-346, fig 2 [from type] (Reinw., Blume & Nees) Nees 1845 Steph. 1910 Typus: Cuba: ? sd, Wright sn in hb G G00069163 - n° SIB 124207/1 - G20958 [holotypus, small stem !!!] Basionym: Distribution: spec nov Stephani Bonner Stephani 1910 1965 1985 686 655 928 - syn nov Yuzawa 1991 525 = Frullania rio-janeirensis Spec Hepat 4: 337-338 Index Hepat 5: 278 Stephani Icon Hepat Inedit [Chonanthelia] A- 28, microfiche 46 fig 3110 {from type; "Cuba Wright"] J Hatt Bot Lab 70: 254 [syn sub Fr. riojaneirensis] (Raddi) Ångstr. 1876 Page 99 cucullata Lindenb. & Gottsche 1847 Typus: Mexico.Veracruz, Mirador & Volcan Orizaba, Hacienda de Mirador, 3000ft sd, Liebmann 207 in hb G, (ex hb BESCHERELLE) in hb S (B22708-22710) [typus] Basionym: Distribution: USA ( FL ), Mexico, Panama, Galapagos, Cocos Isl [Costa Rica], Costa Rica, Peri spec nov - syn nov - syn nov Lindenberg & Gottsche in G., L. & N. Mitten in Seemann Gottsche Gottsche Stephani Evans Clark & Frye Frye & Clark Clark & Svihla 1847 2848 Syn Hepat: 782-783 1857 1863 1867 1911 1914 1945 1947 1948 1494 103 104 686 1902 313 204 325 Clark 1953 1106 Bonner Stotler Stotler Reed & Robinson Cole Stephani 1965 1969 1970 1971 1983 1985 655 4761 194 1406 1407 928 Fulford & Sharp Schuster Stotler, Allen, Gradstein, McGuinness, Whittemore & Chung Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Gradstein, Meneses & Arbe Feuerer 1990 1992 1998 105 603 703 Bot Voyage HMS Herald: 246 [Panama] Separata of Mex. Leverm.: 249 K Danske Vid Selks Skrifter, ser 5, 6: 345-346 [Mexico] Spec Hepat 4: 493 [Mexico, Costa Rica] Bryologist 17: 90 Bryologist 48: 53-56, fig 1-20 Univ Wash Publ Biol 6 (5): 742-745, fig Frc(1-13) Bryologist 51: 24 [key to Fr. of Middle America: Florida, Mexico, Costa Rica, Panama] Proc California Acad Sci 4 ser, 27 (18): 663 [Galapagos Isls: Santiaga, Cocos Isl [Costa Rica]] Index Hepat 5: 278 Nova Hedwigia 18: 444 [syn sub Fr. intumescens] The genus Frullania, separata: 444 [syn sub Fr. intumescens] Phytologia 21 (1): 19 [Costa Rica: Monteverde] Brenesia 21: 154 [Costa Rica] Stephani Icon Hepat Inedit [Thyopsiella] F- 50, microfiche 57 fig 3944 ["olim Frull mollicula Spruce, Bolivia, Original, leg Rusby"] - 3945 ["Mexico (leg Liebmann) Original"] Mem NY Bot Gard 63: 41 [Mexico] Hepat N America 5: 71-77, fig 783-784 [a good species] Trop Bryol 15: 179 [Checklist Panama] 2002 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 2003 2008 Rodriquez et al. 2017 975 J Hatt Bot Lab 91: 53 [Bolivia; as Fr. intumescens] 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm 4629 Arnaldoa 24(1): 255 [Peru, La Libertad] = Frullania intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 cucullata var. (beta) zacuapana Gottsche 1863 Typus: Mexico. Zacuapan, 2500-3000ft sd, Liebmann Basionym: Distribution: Mexico var nov - syn nov - syn nov Gottsche Gottsche Stotler Stotler Fulford & Sharp = Frullania intumescens 1863 1867 1969 1970 1990 103 104 4761 194 105 Separata of Mex. Leverm.: 249 K Danske Vid Selks Skrifter, ser 5, 6: 345-346 [Mexico] Nova Hedwigia 18: 444 [syn sub Fr. intumescens] The genus Frullania, separata: 444 [syn sub Fr. intumescens] Mem NY Bot Gard 63: 41 [Mexico] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 100 cuculliloba Steph. 1911 Typus: South Africa. Cape of Good Hope sd, Ecklon sn in hb G G00069207 - n° SIB 124884/1 - G16773 [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Sim Arnell Bonner Vanden Berghen Stephani 1911 1926 1963 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania obscurifolia cucullistipula 686 252 822 655 185 928 Spec Hepat 4: 622-623 Trans Roy Soc S Africa 15: 47 [South Africa] - not known to SIM Hepat S Africa: 147 [syn sub Fr. borgenii] Index Hepat 5: 278 Bull Jard Bot Nat Belg 46 (1/2): 145 Stephani Icon Hepat Inedit [Diastaloba] B- 7, microfiche 48 fig 3233 ["Cap Ecklon"] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 233 [syn sub Fr obscurifolia, West Africa] Mitt. 1879 Steph. ex Bonner 1965 [nom herb] Typus: New Guinea. Fly River Branch -.2.1886, Bauerlein sn in hb G (sub Fr. nodulosa) ex hb MEL, hb STEPHANI 52, Basionym: Distribution: New Guinea Bonner 1965 655 Index Hepat 5: 279 = cunninghamiana Colenso 1889 Typus: New Zealand. Dannevirke, County of Waipawa 1888, W. Colenso 1349 in hb G (ex hb K) Basionym: Distribution: spec nov - syn nov Colenso Stephani Bonner Hamlin = Frullania squarrosula 1889 1892 1965 1972 636 210 655 659 Trans Proc New Zealand Inst 21: 75 J Linn Soc Bot 29: 274 [syn sub Fr. squarrosula] Index Hepat 5: 279 Rec Dominion Mus 7(19): 261 [New Zealand] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Page 101 curiosissima Horikawa 1936 Typus: Bonin Isl. Chichijima, Byôbu-iwa 7.6.1936, Horikawa in hb HIRO [holotypus] Basionym: Distribution: Bonin Isl spec nov - syn nov Horikawa Miller Hattori & Noguchi Bonner Miller Inoue & Iwatsuki 1936 1960 1960 1965 1968 1969 Miller, Whittier & Whittier Furuki & Hattori Yuzawa 1983 1993 2000 Yamada & Iwatsuki 2006 = Frullania repandistipula curnowii 121 920 965 655 672 61 J Jap Bot 12 (9): 667-668, fig 11(a-g) Bryologist 63: 121 [Micronesia] Index spec Typ herb Japon, Bryophyta: 16 Index Hepat 5: 280 Micronesia 4 (1): 74 [key to Guam & Northern Micronesia] Bull Natn Sci Mus Tokyo 12 (2): 303 [syn sub Fr. uvifera > Fr. ramuligera] 609 Prod Flora Hepat Polynesiae: 124 3 Hikobia 11: 261-263 [syn sub Fr. repandistipula S.L.] 1659 Natural Environmental Sci Res 13: 10 [Japan, syn sub Fr repandistipula] 1470 J Hatt Bot Lab 99: 29 [Catalog of Japan] Sande Lac. 1853 Steph. 1910 Typus: New Zealand sd, Curnow 35 in hb G 3679 Basionym: Distribution: spec nov - syn nov Stephani Hodgson Bonner Hamlin Hattori Hattori Stephani = Frullania deplanata curranii 1910 1949 1965 1972 1978 1983 1985 686 203 655 659 416 499 928 Spec Hepat 4: 409-410 Trans Roy Soc New Zealand 77 (3): 381-382 [syn sub Fr. deplanata] Index Hepat 5: 280 Rec Dominion Mus 7(19): 261 [New Zealand] Misc Bryol Lichen 8 (3): 46 [sub Fr. scandens auct] J Hatt Bot Lab 54: 140-143 [sub Fr. deplanata] Stephani Icon Hepat Inedit [Galeiloba] C-258, microfiche 53 fig 3679 ["Neu Seeland, Curnow 35"] Mitt. 1855 Steph. 1911 Typus: Philippines. Luzon, in Mte Pulag sd, Curran 16430 pp in hb G G00060664 - n° SIB 131168/1 - G22269 (Curran, H. M. 16430pp) Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Bonner Hattori 1911 1930 1965 1982 686 293 655 0 Stephani 1985 928 = Frullania ternatensis Spec Hepat 4: 553 Ann Bryol Suppl 1: 141 [syn sub Fr. ternatensis] Index Hepat 5: 280 note in hb G: "= syn of F. ternastensis GOTT. var. non-appendiculata HATT (a. form)" Aug.1982 Stephani Icon Hepat Inedit [Thyopsiella] F-166, microfiche 59 fig 4107 [Luzon, Mte Pulog, leg Currau 16430 p parte"] Gottsche 1846 Page 102 curvatirostra J.B.Jack ex Bonner 1965 [nom herb] Typus: Fiji "Viti Isles". Ovalau, Mt. Tana-Lailai 1864, E. Graeffe in hb G (ex hb JACK) Basionym: Distribution: Fiji Stephani Bonner 1911 1965 686 655 Miller, Whittier & Whittier 1983 609 = Frullania curvirostris Frullania nutans J.B.Jack & Steph. 1894 [hom illeg] [non Colenso 1889] Steph. 1901 [non Steph. 1910] Frullania caffraria curvirostrata Spec Hepat 4: 457-458 [not mentioned under Fr. curvirostris] Index Hepat 5: 280 [nom herb; error: not as syn sub Fr. curvirostris in STEPH 1911: 457] Prod Flora Hepat Polynesiae: 124 [nom nud] Steph. 1894 J.B.Jack by Miller, Whittier & Whittier 1983 [orth var] Basionym: Distribution: orth var Miller, Whittier & Whittier = Frullania curvatirostra 1983 Prod Flora Hepat Polynesiae: 124 J.B.Jack ex Bonner 1965 [nom herb] Frullania curvirostris J.B.Jack & Steph. 1894 [hom illeg] [non Colenso 1889] Frullania nutans Steph. 1901 [non Steph. 1910] Frullania caffraria curvirostris 609 Steph. 1894 J.B.Jack & Steph. 1894 [hom illeg] [non Colenso 1889] Typus: Fiji. Ovalau, in Monte Tana-lai-lai 1864, Dr. E. Graeffe (ex hb JACK) in hb G G00067405 - n° SIB 130324/1 - G20546 [typus] in hb G G00067408 - n° SIB 130321/1 - G22454 [typus] in hb G G00067406 - n° SIB 130323/1 - G22452 [typus] in hb G G00067407 - n° SIB 130322/1 - G22453 [syntypus ?] Basionym: Distribution: spec nov Jack & Stephani in Stephani Stephani Verdoorn Bonner Campbell Hamlin Hattori 1894 1911 1930 1965 1971 1972 1980 228 686 551 655 674 659 0 - nom nov Hattori 1981 492 Hattori Miller, Whittier & Whittier Stephani 1981 1983 1985 0 609 928 = Frullania nutans Hedwigia 33: 143-144 Spec Hepat 4: 457-458 Nederl Kruidk Arch Jrg 1930 (2): 159-160 [Tahiti] Index Hepat 5: 281 J Roy Soc New Zealand 1 (1): 12 [Fiji] Records of the Dominion Museum 7 (19): 261 note in hb G: " The name should be changed, I propose Fr. vitiensis HATT, nom nov" J Hatt Bot Lab 49: 370-371, fig 8(a-j) from type [nom nov: as Fr. novocurvirostris] note in hb G: "= F. novocurvirostris HATT. (nom nov)" Prod Flora Hepat Polynesiae: 124 Stephani Icon Hepat Inedit [Galeiloba] C-121, microfiche 51 fig 3511 [from type; "Insula Vitienses, Vitien Ovalau, in Mte Tana-lailai, Jack del, Dr E Graeffe"] Steph. 1901 [non Steph. 1910] Frullania caffraria Steph. 1894 Page 103 curvirostris var. setchelii Verd. 1930 Typus: Society Isl. Tahiti Basionym: Distribution: var nov - syn nov Verdoorn Hattori Miller, Whittier & Whittier = Frullania jacquinotii 1930 1981 1983 551 492 609 Nederl Kruidk Arch Jrg 1930 (2): 159-160 J Hatt Bot Lab 49: 371 [conspecific with Fr. jacquinotii] Prod Flora Hepat Polynesiae: 124 Gottsche 1910 curvistipula var. latistipula S.Hatt. 1978 Typus: Indonesia. West Irian, Jayawijaya, "Kam Hellwiggebergte", 2600m 1913, A. Pulle 910a in hb FH [hb VERDOORN 11683 sub Fr. curvistipula) typus], in hb NICH [isotypus] Basionym: Distribution: Indonesia (Irian Jaya), New Guinea var nov - syn nov Hattori Hattori Grolle & Piippo Hattori 1978 1982 1984 1988 Mizutani, Hasegawa & Iwatsuki 2009 482 495 143 516 J Hatt Bot Lab 44: 530-532, fig 204(h-n) J Hatt Bot Lab 51: 228 [comb nov as Fr. subdentata var. latistipula] Acta Bot Fennica 125: 18 [Cat. West Melanesia] J Hatt Bot Lab 65: 419-421, fig 2 [PNG - Mt. Alb. Edward, key to subdentata & curvistipula var. latistipula) 2112 Type sepecimens Hatt Bot Lab (NICH) 22: Isotype. New Guinea, Kam Hellwiggebergte. 2600 m. 1913.A. Pulle 910a (NICH 431913). = Frullania subdentata var. latistipula (S.Hatt.) S.Hatt. 1982 cylindrica Gottsche ex Lehm. 1844 Typus: Mexico F. Leibold in hb S (B22711, 22714) [typus] in hb GLM 05968, 05969, 05970, 05971 Basionym: Distribution: Mexico, Peru, Brazil, Chile spec nov - syn nov Gottsche ex Lehmann Gottsche in G., L. & N. Hampe Hampe Gottsche Gottsche Gottsche Reichardt Ångström Schiffner Stephani Bonner Menzel Stephani 1844 1845 1851 1854 1858 1863 1867 1870 1876 1893 1911 1965 1984 1985 Fulford & Sharp Grims Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 1990 2001 2002 = Frullania brasiliensis 589 2850 281 284 1511 103 104 884 579 572 686 655 142 928 Pugillus 8: 13-14 Syn Hepat: 458-459 (No 107) Linnaea 24 (3): 303 [Costa Rica] Linnaea 27: 444 [Fo Chile, Peru] Bot Zeitung (Berlin) 16 (Beilage): 24 & 29 Separata of Mex. Leverm.: 247 K Danske Vid Selks Skrifter, ser 5, 6: 343 [Mexico] Reise österr Fregatte Novara, Bot Theil, Bd 1: 153 [Brazil] Öfversigt K Vetensk Akad Förh 33 (7): 89 [Brazil] Nova Acta K Leop-Carol Deutsch Akad Naturf Halle 60 (2): 222 Spec Hepat 4: 490 [syn sub Fr. brasiliensis] Index Hepat 5: 282 Willdenowia 14: 509 [syn of Fr. brasiliensis] Stephani Icon Hepat Inedit [Thyopsiella] F- 37d, microfiche 57 fig 3926 ["Frullania cylindrica Gottsche, No 1520, Mexico, leg Leibold"] 105 Mem NY Bot Gard 63: 41 [Mexico] 1136 Moosherbar in hb LINZ, from Kalifornien [Internet] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] Raddi 1822 Page 104 cylindrica var. (beta) Gottsche 1845 Basionym: Distribution: Gottsche in G., L. & N. Gottsche Gottsche = Frullania brasiliensis 1845 1863 1867 2850 Syn Hepat: 458-459 103 Separata of Mex. Leverm.: 247 104 K Danske Vid Selks Skrifter, ser 5, 6: 343 [Mexico] Raddi 1822 cylindrica var. (delta) grandis Gottsche 1847 Basionym: Distribution: Gottsche in G., L. & N. = Frullania brasiliensis 1847 2848 Syn Hepat: 781 Raddi 1822 cylindrica var. (gamma) Gottsche 1845 Basionym: Distribution: Gottsche in G., L. & N. = Frullania brasiliensis 1845 2850 Syn Hepat: 458-459 Raddi 1822 cylindrica var. atro-fusca Gottsche 1847 Basionym: Distribution: Gottsche in G., L. & N Gottsche Gottsche = Frullania brasiliensis 1847 1863 1867 2848 Syn Hepat: 781 103 Separata of Mex. Leverm.: 247 104 K Danske Vid Selks Skrifter, ser 5, 6: 343 [Mexico] Raddi 1822 cylindrica var. granatensis Gottsche 1864 Basionym: Distribution: - syn nov Gottsche Bonner Gradstein & Hekking Uribe & Gradstein = Frullania brasiliensis 1864 1965 1979 1998 478 655 483 707 Ann Sci Nat Bot, ser 5, 1: 171-173 Index Hepat 5: 282 J Hatt Bot Lab 45: 112 [Cat Colombia; as syn sub Fr. brasiliensis] Bryophyt Bibl 53: 38 [Catalogue Colombia] Raddi 1822 Page 105 cylindrica var. lindigii Gottsche 1864 Typus: Colombia. Asseradero, 2500m -.7.1860, Lindig 1725 in hb G Basionym: Distribution: Colombia - comb nov Gottsche Spruce 1864 1884 478 243 Bonner 1965 655 = Frullania brasiliensis Ann Sci Nat Bot, ser 5, 1: 171-173 Trans Proc Bot Soc Edinburgh 15: 20 [ as "subvar lindigii Gotts Hep N Granat 77"] Index Hepat 5: 282 Raddi 1822 cylindrica var. lindigii fo. brevicalycina Gottsche 1864 Typus: Colombia. Asseradero -.7.1860, Lindig Basionym: Distribution: Colombia Gottsche Bonner = Frullania brasiliensis 1864 1965 478 655 Ann Sci Nat Bot, ser 5, 1: 171-173 Index Hepat 5: 282 Raddi 1822 cylindrica var. lindigii fo. laevior Gottsche 1864 Basionym: Distribution: Gottsche Bonner = Frullania brasiliensis 1864 1965 478 655 Ann Sci Nat Bot, ser 5, 1: 171-173 Index Hepat 5: 282 Raddi 1822 cylindrica var. minor Gottsche 1863 Basionym: Distribution: Gottsche Gottsche Husnot = Frullania brasiliensis cyparioides 1863 1867 1875 103 Separata of Mex. Leverm.: 247 104 K Danske Vid Selks Skrifter, ser 5, 6: 343 [Mexico] 2435 Rev Bryol 2(1): 4 [F. cylindrica var minor? Husnot.222 - Sur les arbres, près de la source du Galion (Guad.). Alt. 1150m.] Raddi 1822 Nees 1838 [nom inval, Art. 32.1(c)] Basionym: Distribution: Nees = Frullania cyparioides 1838 261 Naturg Europ Leberm 3: 210 [nomen in foot note] (Schwägr.) Nees 1845 Page 106 cyparioides [auct. non (Schwägr.) Nees 1838] Vanden Berghen 1976 {Mauritius} Typus: Mauritius sd, Sieber in hb STR (hb SIEBER] Basionym: Distribution: Vanden Berghen Grolle = Frullania anderssonii cyperoides 1976 1995 185 250 Bull Jard Bot Nat Belg 46 (1/2): 177 [Mauritius] Bryophyt Bibl 48: 146 Ångstr. 1873 Glenny, Meagher & Brown 2003 [orth var] Basionym: Distribution: Glenny, Meagher & Brown = Frullania cyparioides dalleana 2003 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html (Schwägr.) Nees 1845 Gerola 1947 Typus: Egypt. Abyssinia, Irgalen, Dalle 25.11.1937, Vatora Basionym: Distribution: spec nov - syn nov Gerola 1947 Cufodontis Bonner Vanden Berghen Wigginton 1952 1965 1976 2002 Wigginton 2004 = Frullania socotrana dapitana 1532 Lavori di botanica / Istituto Botanica dell'Università di Padova 12: 477, fig 10(a-c) [Ethiopia] 2562 Phyton 4 (1-3): 76 [Ethiopia] 655 Index Hepat 5: 283 185 Bull Jard Bot Nat Belg 46 (1/2): 13 [syn sub Fr. socotrana] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Mitt. 1888 Steph. 1894 Typus: Philippines, Ins. Dapitam 1884/1885, Micholitz, sn in hb G Basionym: Distribution: spec nov - syn nov Stephani Stephani Merrill Verdoorn Verdoorn Kamimura Bonner Miller Miller, Whittier & Whittier Stephani = Frullania nodulosa 1894 1911 1914 1928 1930 1943 1965 1968 1983 1985 228 686 1484 244 Hedwigia 33: 150 Spec Hepat 4: 583-584 Philippine J Sci C, Bot 9(1): 40 [Guam] Ann Crypt exot 1 (2): 220 2798 655 672 609 928 Nat Sci Mus 14: 293-294, fig 2(A-M) [= Fr nodulosa fo dapitana] Index Hepat 5: 283 Micronesia 4 (1): 74 Prod Flora Hepat Polynesiae: 124 Stephani Icon Hepat Inedit [Homotropantha] D-18, microfiche 54 fig 3730 ["Philippinae, Insula Dapitam, Micholitz"] - 3731 (Reinw., Blume & Nees) Nees 1845 Page 107 darjeelingensis S.Hatt. ex S.Hatt. 1981 [pro syn] Basionym: Distribution: Hattori 1981 491 J Hatt Bot Lab 49: 149 = Frullania pseudoschensiana var. darjeelingensis S.Hatt. 1981 darwinii Gradstein ex Gradstein & Weber 1982 [nom inval; Art 32.1(c)] Basionym: Distribution: Galapagos Isls [Ecuador] spec nov Gradstein & Weber Gradstein in Zizka & Klemmer Gradstein = Frullania darwinii 1982 1994 2000 496 J Hatt Bot Lab 52: 148 [on 4 Galapagos Isls] 1118 Kl Senckenbergreihe 20: 60 [Galapagos] Email from 14.1.2000 [near Fr. peruviana, but with "ungezähntes Involukrum"] Gradst. & Uribe 2004 davurica ssp. jackii (Gottsche) S.Hatt. 1976 [nom inval] Basionym: Frullania jackii Gottsche 1863 Distribution: China (Yunnan), Taiwan comb nov Hattori Kuo & Chiang Konstantinova, Potemkin & Schijakov Piippo, He & Kaponen Rao, Enroth, Piippo & Koponen Grolle & Long 1976 1988 1992 67 948 651 1997 1997 2000 628 685 760 Bai in Gao & Cao Bai Bai 2000 2001 2002 = Frullania jackii Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 19-21 Taiwania 33: 11 [Index of Taiwan] Arctoa 1:102 [USSR] Ann Bot Fennici 34: 52, 61 [Sichuan] Hikobia 12: 183 [checklist Hunan] J Bryol 22: 111 [checklist of Europe and Macaronesia] & 124 [homotypic syn of Fr. jackii] 1019 Fl Yunnanica 17: 414 [China, Yunnan] 1742 Guizhou Science 19(4): 36 [China, Yunnan] 1189 Chenia 7: 6 [Yunnan, China] Gottsche 1863 davurica ssp. jackii var. grandigaleata (Mueller) S.Hatt. 1976 [nom inval] Basionym: Frullania jackii var. grandigaleata Müller 1938 Distribution: Hattori 1976 67 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21, footnote 2 = Frullania jackii var. grandigaleata Müller 1938 Frullania jackii debilis fo. depauperata Grolle 1970 Steph. ex Bonner 1965 [nom herb] {India} [non Jack ex Bonner 1965] Typus: India. Pondicherry sd, s coll ex hb SANDE LACOSTE, in hb G Basionym: Distribution: India Bonner Stephani = Frullania debilis 1965 1985 655 928 Index Hepat 5: 283 Stephani Icon Hepat Inedit [Galeiloba] C-123, microfiche 52 fig 3513 ["Vorder Indien, Herb Sande Lacoste"] Steph. ex S.Hatt. 1974 Page 108 debilis J.B.Jack ex Bonner 1965 [nom herb] {Argentina} [non Steph. ex Bonner 1965] Typus: Argentinia. Siambon near Tucuman 1873, P.G. Lotentz sn hb JACK 1, in hb G (sub Fr. tetraptera) Basionym: Distribution: Argentina Bonner 1965 655 Index Hepat 5: 284 = decipiens Herzog 1923 [non Beauverd in Steph. 1924] Typus: Chile. Valdivia, "an Bäumen im Stadtpark von Valdivia" -.12.1911, Th. Herzog sn in hb JE [holotypus] Basionym: Distribution: spec nov - syn nov Herzog Reimers Herzog in Skottsberg Bonner Müller & Zündorf = Frullania chilensis decipiens 1923 1926 1942 1965 2007 Hedwigia 64: 8, fig 4 [Chile, Valdivia] Hedwigia 66: 48 [syn sub Fr. chilensis] Nat Hist Juan Fernandez & Easter Island, 2: 738 Index Hepat 5: 284 http://www2.unijena.de/biologie/spezbot/herbar/bryotypes/Uebersicht%20F.htm Steph. 1894 Beauverd in Steph. 1924 [hom illeg] [non Herzog 1923] Basionym: Frullania galapagona Distribution: Steph. 1911 [hom illeg] [non AONGSTRÖM 1873] nom nov Beauverd in Stephani 1924 - syn nov - syn nov Bonner Stotler Stotler 1965 1969 1970 = Frullania ecuadorensis Frullania kunzei decurviloba 112 113 722 655 1442 909 Spec Hepat 6: 570 [Frullania decpiens Beauverd, nom. nov. = F . galapagona St., Sp. Hep. IV (1911], 655 Index Hepat 5: 284 4761 Nova Hedwigia 18: 471-472 194 The genus Frullania, separata: 471-472 Steph. 1911 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1910 Typus: New Zealand. sd, Colenso 1346 ? in hb G Basionym: Distribution: spec nov - syn nov Stephani Hodgson Bonner Hamlin Hattori Stephani = Frullania solanderiana 1910 1949 1965 1972 1983 1985 686 203 655 659 499 928 Spec Hepat 4: 412-413 Trans Roy Soc New Zealand 77 (3): 377 Index Hepat 5: 284 Rec Dominion Mus 7(19): 261 [New Zealand] J Hatt Bot Lab 54: 170-172 Stephani Icon Hepat Inedit [Galeiloba] C-259, microfiche 53 fig 3680 ["sub Frull solanderi Col, N Zelandia, Colenso 1346"] Colenso 1889 Page 109 deflexa Steph. by Steph. & Watts 1914 [orth var] Typus: New Hebrides. Aneitum sd, Gunn (sent by Lillie) Basionym: Distribution: Stephani in Stephani & Watts Miller, Whittier & Whittier = Frullania deflexa delavayi Typus: 1914 1983 220 609 J Proc Roy Soc New S Wales 48: 107 [Aneityum] Prod Flora Hepat Polynesiae: 124 Mitt. 1862 [auct. non Steph. 1884] Verd. in Hand.-Mazz. 1930 Handel-Mazzetti 977 Basionym: Distribution: auct - syn nov Verdoorn in Handel-Mazzetti Hattori & Lin Wu in Li, Gao, Wu et al. = Frullania schensiana delavayi 1930 1985 1985 550 Symb Sinicae 5: 42 [Handel-Mazzetti 977] 505 J Hatt Bot Lab 59: 143 1169 Bryoflora Xizang (Tibet), Science Press, Beijing: 527, fig 228(11-16) C.Massal. 1897 Steph. 1894 Typus: China. Yuennan, in mte. Ma-eul-chan, 2500m 9.7.1899, Delavay ex hb BESCHERELLE, in hb PC PC0103042 [isotypus] (sd, Delavay sn) in hb G Basionym: Distribution: Taiwan spec nov - syn nov Stephani Stephani Stephani Verdoorn in Handel-Mazzetti Kamimura Hong Nakanishi Bonner Hattori in Hara Hattori in Hara Kodama Hattori in Ohashi Hattori Wu in Li, Gao, Wu et al. Stephani 1894 1894 1910 1930 1961 1962 1963 1965 1966 1971 1972 1975 1975 1985 1985 228 211 686 550 199 Kuo & Chiang Piippo Parihar, Lal & Katiyar 1988 1990 1994 948 521 2196 Yamada & Choe Nath & Asthana Yuzawa Yamada & Iwatsuki 1997 1998 2001 2006 717 683 1660 1470 = Frullania inflexa 960 655 841 190 1375 187 47 1169 928 Hedwigia 33: 157 [spec nov] Mem Soc Nat Sci, Nat Math Cherbourg 29: 212 [China; spec nov] Spec Hepat 4: 446 Symb Sinicae 5: 42 [China] J Hatt Bot Lab 24: 83, fig 29(1-24), map 9 Thesis of Catholic Medical College 5: 547, fig 54(1- ) Hikobia 3(4): 325 [Taiwan] Index Hepat 5: 288 Flora of Eastern Himalaya: 525 Univ Mus Univ Tokyo Bull 2: 234 [Bhutan] Spec Publ Osaka Mus Nat Hist 4: 120 [Central Japan] Univ Mus Univ Tokyo Bull 8: 233 [Nepal] Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 76-77 [syn sub Fr. inflexa] Bryoflora Xizang (Tibet), Science Press, Beijing: 527, fig 228(11-16) Stephani Icon Hepat Inedit [Galeiloba] C-124, microfiche 52 fig 3514 [from type; "China, Yunnan, Bois de Ma cul chan, 2500m, Delavay 9/7. 89"] Taiwania 33: 11 [Index of Taiwan] J Hatt Bot Lab 68: 39 [annotated catalogue of China] Hepat Anth India, annotated checklist: 21 [Eastern Himalaya, Nepal] new record from India (Hattori 1966, 1975) J Hatt Bot Lab 81: 287 [sub Fr inflata] J Bryol 20(1): 233-237 [South India] Natural Environmental Sci Res 14: 43 [Japan, syn sub Fr inflexa] J Hatt Bot Lab 99: 29 [Catalog of Japan] Mitt. 1861 Page 110 delicatula Colenso 1887 Typus: New Zealand. County of Hawke`s Bay, NW of Napier 1885, A. Hamilton Basionym: Distribution: spec nov - syn nov Colenso Stephani Hamlin = Frullania falciloba deloti 1887 1892 1972 634 210 659 Trans Proc New Zealand Inst 19: 297 J Linn Soc Bot 29: 274 [syn sub Fr. falciloba] Rec Dominion Mus 7(19): 261 [New Zealand] 1975 1983 670 609 Bryophyt Bibl 4: 91 Prod Flora Hepat Polynesiae: 125 [nom nud] Lehm. 1844 N & M in Gradstein 1975 Typus: Gambir Isl, Mangrareva sd, Hombron in hb PC-MONT [holotypus], in hb H, L, M, O PC, PRC, W Basionym: Distribution: Gradstein Miller, Whittier & Whittier = Frullania aeolotis Nees & Mont. ex Nees 1854 [pro syn] {hb MONT No 125} Frullania aeolotis var. singapurensis Nees 1845 Frullania riparia Hampe 1838 densepinnata Steph. 1911 Typus: Magadascar 1896, Forsyth Major sn in Hb G Basionym: Distribution: Madagascar spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania apicalis densepinnata 686 655 185 928 Spec Hepat 4: 480 [spec nov] Index Hepat 5: 285 Bull Jard Bot Nat Belg 46 (1/2): 52 Stephani Icon Hepat Inedit [Thyopsiella] F- 4, microfiche 56 fig 3872 ["Madagascar, Forsyth Major, 491pp, Unicum"] - 3873 250 Bryophyt Bibl 48: 62 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apicalis, West Africa] Mitt. 1879 Steph. in Levier 1901 [nom nud] Basionym: Distribution: nom nud Stephani in Levier = Frullania densepinnata Frullania apicalis 1901 1456 Rev Bryol 28 (5): 94 [Madagascar] Steph. 1911 Mitt. 1879 Page 111 densifolia Steph. 1894 Typus: Burma: Mt. Aracan 1888, Micholitz sn in hb G 14404 Basionym: Distribution: Brazil ?, China (Zhejiang), Burma, India (Eastern Himalayas: Megalaya) spec nov - syn nov Stephani Stephani Verdoorn Bonner Hattori Miller, Whittier & Whittier Yano Stephani 1894 1911 1932 1965 1972 1983 1984 1985 Piippo Piippo Tian Konrat & Braggins Singh, Kumar & Nath Yano 1990 1990 1997 2001 2008 2008 = Frullania apiculata 228 686 554 655 538 609 501 928 Hedwigia 33: 161 [syn sub syn subgen Diastaloba] Spec Hepat 4: 544 [syn sub syn subgen Thyopsiella] Nederl Kruidk Arch Jrg 1932 (2): 493-494 [China] Index Hepat 5: 285 J Hatt Bot Lab 36: 110-112, fig. 2 [syn nov sub Fr. apiculata] Prod Flora Hepat Polynesiae: 125 J Hatt Bot Lab 56: 504 [checklist Brazil] Stephani Icon Hepat Inedit [Thyopsiella] F-138, microfiche 59 fig 4074 ["Birma, Aracan Hills, Micholitz 1888, 277"] 521 J Hatt Bot Lab 68: 139, annotation 139 about synonymy 521 J Hatt Bot Lab 68: 39 [annotated catalogue of China] 1231 Chenia 3-4: 105 [China, Zhejiang Prov.] 838 J Hatt Bot Lab 91: 256-262 [syn sub Fr. apiculata] 1281 Taiwania 53 (1): 57, [syn sub Fr apiculata] 1684 Boletim Instituto Botanica 19: 30 [Brazil; catalog] (Reinw., Blume & Nees) Nees 1845 densifolia ssp. andamana S.Hatt. ex Singh in Nath & Asthana 2001 [nom nud] Typus: India. Andaman Island Basionym: Distribution: Andaman Isl Singh in Nath & Asthana 2001 1374 Perspectives Indian Bryology: 341[endemic to Andaman Island] = densiloba Steph. ex Yoshinaga 1901 [nom nud] Basionym: Distribution: nom nud - nom nud Stephani ex Yoshinaga Horikawa Verdoorn Hattori Kamimura Kamimura = Frullania densiloba densissima 1901 1934 1934 1944 1952 1961 889 34 240 943 873 199 Bot Mag Tokyo 15: 92 J Sci Hiroshima Univ, ser B, div 2, 2: 236 [nom nud] Ann Bryol Suppl 4: 197 Bull Tokyo Sci Mus 11: 142 [nom nud] Contrib Fl Hepat Shikokuensem, Kochi: 122 J Hatt Bot Lab 24: 69 [syn sub Fr densiloba] Steph. ex A.Evans 1906 Spruce ex Spruce 1884 [pro syn] Basionym: Distribution: Spruce 1884 243 Trans Proc Bot Soc Edinburgh 15: 22 = Frullania gibbosa var. densissima Spruce 1884 Frullania gibbosa Nees 1840 Page 112 denticulata Colenso by Bonner 1965 [orth var] Typus: New Zealand. Hawkes Bay, Pohne, NW of Napier 1885, A. Hamilton Basionym: Distribution: New Zealand Colenso Bonner 1887 1965 = Frullania delicatula Trans Proc New Zealand Inst 19: 297 [= delicatula] Index Hepat 5: 286 Colenso 1887 Frullania falciloba denticuloba 634 655 Lehm. 1844 Steph. by S.W.Arnell 1963 [orth var] Basionym: Distribution: Arnell 1963 = Frullania dentilobula Hepat S Africa: 136 Steph. 1894 Frullania obscura dentiloba 822 (Sw.) Mont. 1840 Steph. by Pearson 1924 [orth var] Basionym: Distribution: Pearson 1924 = Frullania dentilobula Arkiv Botanik 19 (5): 9 [Mt. Elgon in Kenia/Uganda] Steph. 1894 Frullania obscura dentilobula 795 (Sw.) Mont. 1840 Steph. 1894 Typus: E Africa: Kilimandjaro, Marangu Station, 1550 m 25.6.1893, G. Volkens 449 ex hb B, in hb G [holotype] Basionym: Distribution: East Africa, Ethiopia, Madagascar spec nov - syn nov Stephani Stephani Stephani in Engler Stephani Gola Gola Demaret Gerola Demaret & Vanden Berghen Arnell Bonner Stephani 1894 1895 1895 1910 1914 1916 1942 1947 1948 1963 1965 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania obscura 228 212 1481 686 786 3469 1066 1532 59 822 655 928 Hedwigia 33: 137-138 Bot Jahrb 20 (3): 314 [as spec nov] Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [Kilimandschar 1550m] Spec Hepat 4: 319-320 Annali di Botanica 13: 72 [Ethiopia] Mem Reale Accad Sci Torino, ser 2. 65: 9 [Kenya] Bull Jard Bot Etat Bruxelles 16 (2): 304 Lavori di botanica / Istituto Botanica dell'Università di Padova: 475 Bull Jard Bot Etat Bruxelles 19 (1): 77 Hepat S Africa: 136 [as "denticuloba" syn sub Fr. areace] Index Hepat 5: 286 Stephani Icon Hepat Inedit [Chonanthelia] A- 3, microfiche 46 fig 3066 [from type; " Volkens 449"] 250 Bryophyt Bibl 48: 63 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 224 [syn sub Fr arecae, West Africa] (Sw.) Mont. 1840 Page 113 dentistipula Steph. ex Bonner 1965 [nom herb] Typus: Japan. Otsu, Omi 23.2.1913, S.O. Kamura 382 in hb G 14405 Basionym: Distribution: nom herb - syn nov Bonner Hattori Stephani 1965 1974 1985 Scott & Bradshaw Yuzawa Carthy 1986 2001 2003 = Frullania muscicola 655 545 928 Index Hepat 5: 286 J Hatt Bot Lab 38: 207-208 [syn sub Fr. muscicola] Stephani Icon Hepat Inedit [Galeiloba] C-231, microfiche 53 fig 3649 ["Japonia, O Kamura 382, leg 1913"] 56 Brunnonia 8 (1): 34 [Australia] 1660 Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] 1125 Flora Australia Suppl Ser 21: 109 [nom nud] Steph. 1894 depitata [nom herb in hb HBG] {Borneo} Typus: Borneo sd, Pfarrer Korstik Basionym: Distribution: Feuerer 2008 Feuerer 2008 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm = deplanata [auct. non Mitt. 1855] S.Hatt. 1978 Basionym: Distribution: auct - syn nov Hattori Hattori 1978 1983 = Frullania scandens 416 499 Misc Bryol Lichen 8 (3): 46 J Hatt Bot Lab 54: 166 Mont. 1843 deplanata var. flavicans Steph. ex Bonner 1965 [nom herb] {New Zealand} Typus: New Zealand. sd, Colenso 723 or 1445 in hb G Basionym: Distribution: Bonner 1965 655 Index Hepat 5: 287 = deplanchei Steph. ex Tixier 1973 [nom herb] {New Caledonia} Typus: New Caledonia. Prony, Sébertville -.12.1903, L. Cribs 1475 Basionym: Distribution: Tixier Miller, Whittier & Whittier 1973 1983 678 609 Bull Mus Nat Hist Nat Paris, ser 3, 190, Bot 10: 77 [New Caledonia] Prod Flora Hepat Polynesiae: 125 [= Fr. spec] = Page 114 dichotoma Raddi 1822 Typus: Brazil. "Montagne d' Estrella" sd, Raddi in hb S (B22764) [lectotypus] in hb PI three numbered collections [no. 2, lectotypus by Stotler] Basionym: Distribution: Raddi Raddi Raddi Montagne in La Sagra Montagne in La Sagra Nees in G., L. & N. Bonner Raddi [Reprint] 1822 1823 1829 1842 1845 1845 1965 1976 Stotler Myrdal Costa 1998 2002 2009 = Bryopteris diffusa diedericiana Typus: Australia 2873 825 2875 26 25 2847 655 166 Crittogame Brasiliane [Seperate print]: 11 (No 26) Mem Mat Fisica Soc Ital Sci, Modena 19: 35 (No 26) Mem Mat Fisica Soc Ital Sci, Modena 20: pl. 1 (fig. 1) L'ile de Cuba [Franz Ed]: 462-463 = Fr. diffusa La isla de Cuba (Span Ed) 9: 277 Syn Hepat: 286 Index Hepat 5: 287 Flora Brasiliana. Memorie 1819-1828 with Reprint of "Crittogame Brasiliane 1822": 107-108 (No 26), pl. 1 note on label [lectotypification: no 2 in hb PI] 1276 Mosskollekter [Frullania types in hb S] 2866 J Bryol 31: 227 (Raddi s.n., types, (Lectotype by Stotler: : no. 2, PI!). (Sw.) Nees 1845 Gottsche ex Bonner 1965 [nom herb] {Australia} Dietrich in hb G [with note by REIMER "lag im hb M als Fr. (Victoria) diedericiana .."] Basionym: Distribution: Bonner Scott & Bradshaw = Frullania dietrichana 655 56 Index Hepat 5: 287 Brunnonia 8 (1): 34 & 143 [Australia: QLD] Steph. 1910 Frullania seriata dietrichana 1965 1986 Gottsche 1889 Steph. 1910 Typus: Australia. Queensland sd, Amalie Dietrich 361 in hb G 18202 [lectotypus], in hb MEL 61554 [isolectotypus], in hb MEL 61556, A.Dietrich 358, 359, 370 [syntypus] Basionym: Distribution: Australia (QLD) spec nov - syn nov Stephani Bonner Hattori Stephani 1910 1965 1979 1985 686 655 485 928 Scott & Bradshaw Hattori Windolf Carthy Carthy McCarthy McCarthy 1986 1987 1987 2003 2003 2006 2006 56 402 1025 1125 1125 1385 1385 = Frullania seriata Spec Hepat 4: 423-424 Index Hepat 5: 288 J Hatt Bot Lab 45: 332-333, fig 7(a-h) from type [as Fr. dietrichiana] Stephani Icon Hepat Inedit [Galeiloba] C-202, microfiche 53 fig 3609 ["olim Frull diedericiana G nsp, Queensland, Amalie Dietrich 361"] Brunnonia 8 (1): 34 & 143 [Australia: QLD] Mem NY Bot Gard 45: 552 [syn sub Fr. seriata] Austrobaileya 2(4): 390 [checklist Queensland] Flora Australia Suppl Ser 21: 47 [as dietrichiana, sub of Fr seriata] Flora Australia Suppl Ser 21: 42 [Australia (QLD)] Checklist Australian Liverworts [Australia, syn sub Fr seriata] Checklist Australian Liverworts [Australia QLD, as valid species] Gottsche 1889 Page 115 differgens (L. & L.) Nees 1838 [nom inval, Art. 32.1(c)] Basionym: Distribution: Nees Montagne 1838 1840 = Frullania divergens Naturg Europ Leberm 3: 210 [nomen in footnote] Ann Sci Nat Bot, ser 2, 14: 333 ["Fr. divergens var. minor"] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Frullania brasiliensis difficilis 261 467 Raddi 1822 Steph. 1910 Typus: Australia. New South Wales, Richmond River, Pearce's Creek 2.8.1900, W:W: Watts sn in hb G 18467 (ex hb LEVIER 2632) [lectotypus], in hb MEL 61558, in hb NSW Basionym: Distribution: Australia (NSW, QLD) spec nov ?- syn nov Stephani Stephani & Watts Bonner Hattori Stephani 1910 1914 1965 1979 1985 686 220 655 486 928 Windolf Scott & Bradshaw Hattori 1985 1986 1987 1049 56 402 Windolf Hattori Brown, Ramsay & Seur Carthy McCarthy 1987 1988 1992 2003 2006 1025 429 1140 1125 1385 = Frullania seriata diffusa Spec Hepat 4: 425 J Proc Roy Soc New S Wales 48: 107 [Mt. Wilson] Index Hepat 5: 288 J Hatt Bot Lab 46: 129-130, fig 34(a-k) from type Stephani Icon Hepat Inedit [Galeiloba] C-203, microfiche 53 fig 3610 ["Australia, N.S.Wales, inter Levier 2632"] - 3611 J Hatt Bot Lab 59: 174 [Sunshine Coast, Queensland] Brunnonia 8 (1): 34 & 143 [Australia: NSW] Mem NY Bot Gard 45: 553 [syn sub Fr. seriata] - note GWI: may not be synonym Austrobaileya 2(4): 390 [checklist Queensland] Nova Hedwigia Beih 90: 148 [Australia: QLD] Telopea 5(1): 74 [types in hb NSW] Flora Australia Suppl Ser 21: 47 [sub Fr seriata] Checklist Australian Liverworts [Australia, syn sub Fr seriata] Gottsche 1889 Steph. 1911 [hom illeg] [non (SW.) Nees 1838, nec Colenso 1889] Typus: Brazil. Brasilia, St. Vincent 1897, Horeau hb THÉRIOT 23, in hb G Basionym: Distribution: French Guiana, Brazil spec nov - syn nov - syn nov Stephani Herzog Arnell Schiffner & Arnell Bonner Stotler Stotler Stephani 1911 1925 1959 1964 1965 1969 1970 1985 686 1586 448 198 655 4761 194 928 Gradstein & Hekking 1989 517 = Frullania breuteliana Spec Hepat 4: 516-517 Arquivos de botânica do Estado de Sao Paulo 1 (2): 47 [Brazil] Svensk Bot Tidskr 53 (4): 505 [French Guiana] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 88 [Brazil] Index Hepat 5: 288 Nova Hedwigia 18: 453-454 [syn sub Fr. breuteliana] The genus Frullania, separata: 453-454 [syn sub Fr. breuteliana] Stephani Icon Hepat Inedit [Thyopsiella] F- 52, microfiche 57 fig 3947 ["Brasilia, S Vicente, Horeau 1897 leg, Herb Thériot 23"] - 3948 J Hatt Bot Lab 66: 208 Gottsche 1845 Page 116 diffusa (Sw.) Nees 1838 [non Ccolenso 1889, nec Steph. 1911] Basionym: Distribution: comb nov Nees Montagne in La Sagra Montagne in La Sagra Nees in G., L. & N. - syn nov = Bryopteris diffusa diffusa 1838 1842 1845 1845 261 26 25 2847 Naturg Europ Leberm 3: 211 L'ile de Cuba [Franz Ed]: 463 La isla de Cuba (Span Ed) 9: 277 Syn Hepat: 286 (Sw.) Nees 1845 Colenso 1889 [hom illeg] [non (SW.) Nees 1838, nec Steph. 1991] Typus: New Zealand. County of Waipawa 1988, W. Colenso 1347 in hb G Basionym: Distribution: spec nov - syn nov Colenso Stephani Bonner Hamlin Stephani = Frullania pycnantha dilatata 1889 1892 1965 1972 1985 636 210 655 659 928 Trans Proc New Zealand Inst 21: 74-75 J Linn Soc Bot 29: 274 [syn sub Fr. "pyncnantha", orth error] Index Hepat 5: 283 Rec Dominion Mus 7(19): 261 [New Zealand] Stephani Icon Hepat Inedit [Galeiloba] C-269ab, microfiche 54 fig 3693 ["N Zelandia, Colenso 1357"] - 3694 ["sub Fr diffusa Co., N Zelandia, Colenso 1347"] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 [auct. non (L.) Dumort. 1835] Zinovjeva 1973 {Ural} Typus: Russia Basionym: Distribution: Zinovjeva 1973 Konstantinova & Bakalin 2009 = Frullania bolanderi dilatata 2429 Uchonye Zapiski Permskogo Gosudarstvennogo Universiteta, Botanika. 263: 14-137 2113 Arctoa 18: 37 [Fr dilatata described by Zinovjeva were referred by E. Andrejeva to Fr bolanderi] Austin 1869 [auct. non (L.) Dumort. 1835] Oliviera-E-Silva, Milanez & Yano 2002 {Brazil} Typus: Brazil Basionym: Distribution: Oliveira-e-Silva, Milanez & Yano 2002 = Frullania glomerata Mont 1842] dilatata 1077 Trop Bryol 22: 81 & 93 [Brazil] (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec [auct. non (L.) Dumort. 1835] Goeppert 1853 {Fossil in amber} Basionym: Distribution: Göppert 1853 973 Grolle 1985 621 Ber Bekanntm Verh Königl Preuss Akad Wiss 1853: 456 [Fossil in amber] Prace Muz Ziemi z 37: 97-98 [1 specimen = Radula, 3 = Frullania, non conspecific with Fr dilatata] = Page 117 dilatata [auct. non (L.) Dumort. 1835] Mitt. 1891 {Japan} Typus: Japan Basionym: Distribution: auct - syn nov Mitten Iishiba Horikawa Kamimura Bapna & Kachroo Yamada & Iwatsuki = Frullania muscicola dilatata 1891 1912 1935 1961 2000 2006 582 888 1282 199 712 1470 Trans Linn Soc London, ser 2, 3: 202 [Japan] Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Bot Mag Tokyo 49: 52 [Japan] J Hatt Bot Lab 24: 21 [syn nov sub Fr. muscicola] Hepaticology in India II: 175 [syn sub Fr. muscicola] J Hatt Bot Lab 99: 29 [Catalog of Japan] Steph. 1894 [auct. non (L.) Dumort. 1835] Levier 1906 Typus: hb Massalongo 138 Basionym: Distribution: auct - syn nov Levier Hattori & Lin Bapna & Kachroo = Frullania sinensis dilatata 1906 1985 2000 824 505 712 Nuovo Giorn Bot Ital NS 13: 348 [specimen in hb MASSALONGO 138] J Hatt Bot Lab 59: 143 [syn nov sub Fr. sinensis] Hepaticology in India II: 181 [syn sub Fr. sinensis] Steph. 1906 [auct. non (L.) Dumort. 1835] S.Hatt. & Thaithong 1978 {India} Typus: India Basionym: Distribution: India (North: West Bengal (Darjeeling)) -syn nov Hattori & Thaithong 1978 70 Hattori & Thaithong 1978 481 Hattori 1982 131 = Frullania dilatata ssp. asiatica dilatata Bull Natn Sci Mus Tokyo, ser B (Bot), 4 (2): 64 [India. Kashmir, Pahalgam; 12.10.1952, Svihla no 1247, 1251, 1259] J Hatt Bot Lab 44: 178-179, fig 1(a-k) [India. Darjeeling] [? 1965, Sharp & Iwatsuki ] J Jap Bot 57 (9): 258 [as Fr. dilatata sspec asiatica, subspec nov] S.Hatt. 1982 [auct. non (L.) Dumort. 1835] Underwood 1890 Basionym: Distribution: - syn nov Underwood Evans Frye & Clark = Frullania brittoniae dilatata 1890 1897 1947 237 204 Gray's Manual of Botany 6: 706 Trans Connet Acad Arts Sci 10 (1): 15 Univ Wash Publ Biol 6 (5): 773 [sub Fr brittoniae] 615 482 603 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 316 J Hatt Bot Lab 44: 542-545, fig 213 [syn of Fr. parvistipula] Hepat N America 5: 143 [syn of Fr. eboracensis subsp. caucasica] A.Evans 1897 fo. decidua [?] Grolle 1970 Basionym: Distribution: - syn nov Grolle Hattori Schuster = Frullania parvistipula 1970 1978 1992 Steph. 1910 Page 118 dilatata var. fo. viridis De Not. 1865 Basionym: Distribution: Notaris Bonner Grolle 1865 1965 1970 576 655 615 Mem Reale Accad Sci Torino, ser 2, 22 (1): 376 Index Hepat 5: 290 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 310 ? = Frullania microphylla var. deciduifolia Grolle 1970 dilatata var. (beta) microphylla Hampe ex Taylor ex Gottsche, Lindenb. & Nees 1845 [pp] [pro syn] Basionym: Distribution: Taylor in G., L. & N. = Frullania fragilifolia 1845 2850 Syn Hepat: 437 [syn sub Fr fragilifolia] (Taylor) Gottsche, Lindenb. & Nees 1845 [non Austin 1879] dilatata var. blanda De Not. by Bornmüller 1931 [orth var] : Basionym: Distribution: Müller in Rabenhorst Bornmüller Frey & Kürschner 1915 1931 1991 1064 Krypt Flora (2ed) 6, 2: 613 1053 Magyar Bot Lapok 30 (1-2): 21 [Kleinasien] 1033 Bryophyt Bibl 39: 30 [Turkey; may be Fr tamarisci var blanda; Grolle pers comm.] = Frullania tamarisci var. blanda De Not. 1865 Frullania tamarisci var. atrovirens Carrington 1863 dilatata var. briziana C.Massal. 1889 Typus: Italy. Basionym: Distribution: var nov - syn nov Massalongo Müller in Rabenhorst 1889 1915 Müller in Rabenhorst Bonner 1958 1965 1922 Bull Soc Bot Ital: 519-520 1064 Krypt Flora (2ed) 6, 2: 627 [syn sub Fr dilatata var anomala], 628 [Italien, Frankreich] 1028 Krypt Flora (3ed) 6, 2: 1245 655 Index Hepat 5: 289 = Frullania dilatata var. anomala Corbiere 1889 dilatata var. lusitanica Steph. 1887 Typus: Portugal. sd, Prof Henriques Basionym: Distribution: var nov Stephani Sim-Sim = Frullania dilatata 1887 1989 1115 Hedwigia 26 (1): 2 6 Portug Acta Biol (B) 15: 363 [Portugal] 363 [syn sub Fr. dilatata] (L.) Dumort. 1835 Page 119 dilatata var. microphylla (Wallroth) Nees 1838 [pp] Basionym: Jungermannia dilatata var. microphylla Wallroth 1831 Distribution: Madeira [Portugal], Canary Isl ( GC Te )[Spain] - syn nov Nees Nees in G., L. & N. Pokorny Notaris Dumortier Pabst Schiffner Dalla Tore & Sarnthein Luisier Schiffner 1838 1845 1854 1865 1874 1877 1902 1904 1910 1936 261 2850 2263 576 580 1165 905 2694 789 2561 Ade & Koppe Bonner Grolle 1942 1965 1970 206 655 615 Eggers 1982 22 Naturg Europ Leberm 3: 219 Syn Hepat: 415 Verh Zool-Bot Ver Wien. Abhandl 4: 123 [Austria, Unter-Österreich] Mem Reale Accad Sci Torino, ser 2, 22 (1): 376 [= Fr. dilatata fo Bull Soc Roy Bot Belgique 13: 27 Cryptogamen-Flora Deutschlands, Lebermoose: 34 Hedwigia 41: 280 Die Moose (Bryophyta) von Tirol, Vorarlberg und Lichtenstein: 89 Broteria, ser Bot 9 (1): 56 Krit Bemerk europ Leberm, ser 21: 9 [Hepat europ exsicc, nr. 10171018] Hedwigia 81 (1/2): 34 ? Index Hepat 5: 289 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 310 [syn sub Fr. microphylla var deciduifolia] Crypt Bryol Lichen 3 (4): 288 = Frullania microphylla var. deciduifolia Grolle 1970 dilatata var. var 1 [auct. non (L.) Dumort. 1835] Sull. 1846 {USA} Basionym: Distribution: Sullivant 1846 764 Musci Alleghaniensis: 63 [Exsicc No 267, reprint of labels] = dilatata var. var 2 [auct. non (L.) Dumort. 1835] Sull. 1846 {USA} Basionym: Distribution: Sullivant 1846 764 Musci Alleghaniensis: 63 [Exsicc No 268, reprint of labels] = dilatata var. var 3 [auct. non (L.) Dumort. 1835] Sull. 1846 {USA} Basionym: Distribution: Sullivant 1846 764 Musci Alleghaniensis: 63-64 [Exsicc No 269, reprint of labels] = dilatata ssp. appalachiana R.M.Schust. 1962 [nom nud] Basionym: Distribution: Schuster Stotler & Crandall-Stotler = Frullania appalachiana 1962 1977 398 Transactions of the British Bryological Society 4: <230-246> Bryologist 80: 420 [= nom nud] R.M.Schust. 1983 Page 120 dilatata ssp. subdilatata (C.Massal.) S.Hatt. 1974 Basionym: Distribution: comb nov Hattori Hattori Piippo = Frullania subdilatata diplota 1974 1982 1990 545 131 521 J Hatt Bot Lab 38: 219, fig 105 [comb nov pro Fr. subdilatata] J Jap Bot 57 ( 9): 258-259 J Hatt Bot Lab 68: 40 [annotated catalogue of China] [Fr subdilatata is a good species] C.Massal. 1906 [auct. non Taylor 1846] Steph. 1901 Typus: Patagonia; Tierra de Fuego Basionym: Distribution: = diplota [auct. non Taylor 1846] Steph. 1901 [pp] Typus: Punta Arenas / Porvenir / Rio Azopardo sd, Dusen in hb UPS Basionym: Distribution: - syn nov Stephani 1901 799 Engel 1978 182 = Frullania magellanica Bihang Kgl Sv Vetensk-Akad Handl bot 26 III (17): 32 [from South Patagonia] Fieldiana Bot 41: 242 F.Weber & Nees 1845 Page 121 diplota Taylor 1846 Typus: Australia. New South Wales 1836, R. Cunningham in hb MEL, typus in hb S (B22765-22766) in hb GLM 05997 Basionym: Distribution: spec nov - syn nov Taylor Taylor in G., L. & N. Bastow 1846 1847 1888 Stephani Stephani 1889 1901 Stephani Rodway Bonner Hattori & Mizutani Stephani 1911 1916 1965 1982 1985 Scott & Bradshaw Beckmann Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Carthy Glenny, Meagher & Brown 1986 2002 2002 Glenny, Meagher & Brown 2004 McCarthy 2006 = Frullania rostrata diplota 2003 2004 292 London J Bot 5: 405 2848 Syn Hepat: 780 845 Papers Proceed Roy Soc Tasmania for 1887: 265-266, pl 36 [Tasmania] 940 Hedwigia 28: 158 [Australia] [synonym with Fr. magellanica, Fr. patagonica] Frullania diplota TAYLOR 1846 686 Spec Hepat 4: 658-659 914 Papers Proceed Roy Soc Tasmania for 1916: 137 655 Index Hepat 5: 290 495 J Hatt Bot Lab 52: 447 928 Stephani Icon Hepat Inedit [Diastaloba] B-91, microfiche 49 fig 3339 ["Australia, Mt William, D. Sullivan 1882"] 56 Brunnonia 8 (1): 34 [Australia] 971 Australasian bryol Newsletter 45(May 2002): 9 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 1125 Flora Australia Suppl Ser 21: 46 [sub Fr rostrata] 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1126 NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1385 Checklist Australian Liverworts [Australia, syn sub Fr rostrata] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 [auct. non Taylor 1846] Steph. 1901 [pp] Typus: Punta Arenas / Porvenir / Rio Azopardo sd, Dusen in hb UPS, hb S-PA Basionym: Distribution: Stephani 1901 799 Engel 1978 182 = Frullania patagonica diptera Bihang Kgl Sv Vetensk-Akad Handl bot 26 III (17): 32 [from South Patagonia] Fieldiana Bot 41: 242 Steph. 1910 [auct. non (L. & L.) L. & L.] Sande Lac. 1857 Typus: hb SANDE LACOSTE in hb LUGD BAT Basionym: Distribution: - syn nov Sande Lacoste Verdoorn = Frullania squarrosa Frullania ericoides 1857 1929 430 246 Verh Konikl Akad Wetenschappen 5: 79 Ann Bryol 2: 156 [Fr. diptera ex Java by SL 1857 = squarrosa] (Reinw., Blume & Nees) Nees 1845 (Nees) Mont. 1839 Page 122 dissitifolia Steph. 1911 Typus: New Zealand. sd, Colenso ex hb CURNOW, in hb G 20551 Basionym: Distribution: New Zealand spec nov - syn nov Stephani Hodgson Bonner Hamlin Hattori & Mizutani Stephani 1911 1949 1965 1972 1982 1985 Glenny Glenny & Malcolm 1998 2005 Konrat, Braggins, Asakawa & Toyota 2006 = Frullania congesta divergens Spec Hepat 4: 659 Trans Roy Soc New Zealand 77 (3): 383 [syn nov sub Fr. rostrata] Index Hepat 5: 291 Rec Dominion Mus 7(19): 261 [New Zealand] J Hatt Bot Lab 52: 447 [syn sub Fr. rostrata] Stephani Icon Hepat Inedit [Diastaloba] B-92, microfiche 49 fig 3340 ["N Zealand, Ins meridion, int Winteralose, W. N Beckett 461"] 1412 Tuhinga 10: 132 [sub Fr rostrata] 1410 ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr rostrata] 1465 J H B L 100: 553-576 [lectotypification; syn nov sub Fr congesta] Gottsche, Lindenb. & Nees 1845 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Basionym: Jungermannia divergens Distribution: Brazil comb nov 686 203 655 659 495 928 Lehm. & Lindenb. 1833 Lehmann & Lindenberg in G., L. & N. Reichardt Stephani Dusen Stephani Massalongo Schiffner & Arnell Bonner Stephani = Frullania brasiliensis 1845 2850 Syn Hepat: 459-460 (No 108) 1870 1897 1905 1911 1911 1964 1965 1985 884 1059 710 686 903 198 655 928 Reise österr Fregatte Novara, Bot Theil, Bd 1: 153 [Brazil] Bihang Kgl Sv Vetensk-Akad Handl bot 23 III (2): 18 [Brazil] Arch Mus Nac Rio de Janeiro 13: 115 [Brasil] Spec Hepat 4: 494 Rev Bryol 38: 11 [Brasil] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 88 [Brazil] Index Hepat 5: 291 Stephani Icon Hepat Inedit [Thyopsiella] F- 53, microfiche 57 fig 3949 ["Brasilia (Horeau legit)"] Raddi 1822 divergens var. minor Mont. 1840 [nom herb] Typus: sd, Lepr. 2 Basionym: Distribution: nom herb Montagne = Frullania brasiliensis 1840 467 Ann Sci Nat Bot, ser 2, 14: 333 Raddi 1822 Page 123 diversifolia Typus: Java Steph. 1911 sd, s in hb STR (ex hb NEES) in hb G Basionym: Distribution: Indonesia (Java), New Caledonia spec nov - syn nov Stephani Verdoorn Bonner Miller, Whittier & Whittier Stephani = Frullania junghuhniana drummondii 1911 1930 1965 1983 1985 686 293 655 609 928 Spec Hepat 4: 640-641 Ann Bryol Suppl 1: 134 [syn sub Fr. junghuhniana] Index Hepat 5: 291 Prod Flora Hepat Polynesiae: 125 Stephani Icon Hepat Inedit [Diastaloba] B-51, microfiche 49 fig 3291 [from type; "Java Herb Nees"] Gottsche 1845 Taylor 1846 Typus: USA. Lousiana sd, Drummond Basionym: Distribution: spec nov - syn nov Taylor Taylor in G., L. & N. Evans Frye & Clark = Frullania kunzei dubia 1846 1847 1897 1947 292 2848 237 204 London J Bot 5: 401 Syn Hepat: 777 Trans Connet Acad Arts Sci 10 (1): 30 Univ Wash Publ Biol 6 (5): 759 [sub Fr kunzei] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Lindenb. & Hampe in Hampe 1851 Typus: Costa Rica. Lacis montalis ? Oersted typus in hb S (B22767) Basionym: Distribution: Mexico, Costa Rica spec nov - syn nov - syn nov Lindenberg & Hampe in Hampe Stephani Clark Bonner Stotler Stotler Stephani 1851 1911 1958 1965 1969 1970 1985 281 686 388 655 4761 194 928 Fulford & Sharp Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 1990 2002 105 782 = Frullania intumescens Linnaea 24 (6): 641 [Costa Rica] Spec Hepat 4: 494-495 Bryologist 61: 149-151, fig 1-13 Index Hepat 5: 292 Nova Hedwigia 18: 444 [syn sub Fr. intumescens] The genus Frullania, separata: 444 [syn sub Fr. intumescens] Stephani Icon Hepat Inedit [Thyopsiella] F- 54, microfiche 57 fig 3950 [""Costarica (Oersted leg)"] Mem NY Bot Gard 63: 42 & 43 [Mexico] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 124 dupuisana Steph. ex Demaret & Vanden Berghen 1947 [pro syn] Typus: Congo Belge, Bangosameno 1893, P. Dupuis sn. hb CARDOT 95, in hb G Basionym: Distribution: pro syn Stephani 1910 686 Demaret & Vanden Berghen Bonner 1947 1965 1101 655 Stephani 1985 928 Wigginton 2002 1012 Wigginton Wigginton 2004 2004 1312 1172 = Frullania squarrosa (Reinw., Blume & Nees) Nees 1845 Frullania ericoides duriaei Spec Hepat 4: 372 [syn sub Fr. bangiensis]; according BONNER 1965, but is not correct Bull Jard Bot Etat Bruxelles 18: 238 [syn sub Fr squarrosa] Index Hepat 5: 292 [error by Bonner: not in STEPH 1910; as nom herb syn sub Fr. bangiensis] Stephani Icon Hepat Inedit [Galeiloba] C- 6a, microfiche 50 fig 3354 [Frull bangiensis St nsp (olim Dupuisana"] Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 231 [syn sub Fr ericoides, West Africa] (Nees) Mont. 1839 Nees ex Bonner 1965 [nom herb] Typus: Spain. Asturias sd, Durieu ex hb MONTAGNE, in hb G Basionym: Distribution: Stephani Bonner = Frullania hispanica 1911 1965 686 655 Spec Hepat 4: 560 [not mentioned as syn] Index Hepat 5: 292 [error: not as syn sub Fr. hispanica in STEPH 1911: 560] Nees 1838 [non Hampe ex Lindenb. in G., L. & N. 1845] Frullania tamarisci var. sardoa (De Not.) De Not. 1865 duricaulis Steph. ex Steph. 1965 [nom herb] Typus: [1] Caripe pr. Para - [2] Cayenne [1] leg Spruce - [2] Leprieur sn in hb G 23573 Basionym: Distribution: Stephani 1985 928 Gradstein & Hekking 1989 517 = Frullania apiculata Stephani Icon Hepat Inedit [Diastaloba] B-29, microfiche 48 fig 3262 ["Caripe pr. Para leg Spruce auch Cayenne (Leprieur legit)"] J Hatt Bot Lab 66: 208 (Reinw., Blume & Nees) Nees 1845 Page 125 duricaulis Spruce 1884 Typus: Brazil. Caripe near Para sd, Spruce in hb G Basionym: Distribution: spec nov - syn nov Spruce Stephani Bonner Yano Yuzawa & Koike Stotler in Seaward & FitzGerald = Frullania gymnotis 1884 1911 1965 1984 1989 1996 243 686 655 501 519 1092 Trans Proc Bot Soc Edinburgh 15: 56 & 59 Spec Hepat 4: 636 Index Hepat 5: 292 J Hatt Bot Lab 56: 504 [checklist Brazil] J Hatt Bot Lab 66: 349-359, fig 5 [from type] [syn sub Fr. gymnotis] Richard Spruce, Roy Bot Gard Kew: 133-134, fig 17 [subgen Diastaloba] Nees & Mont. in Mont. 1843 Frullania obcordata var. armata Lindenb. & Gottsche 1851 Frullania obcordata dusenii (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. ex S.Hatt. 1972 [pro syn] {India} [non Steph. in Dusen 1905] Typus: India orientalis Pfleiderer in hb G (14542) [Pfleiderer K 12, holotypus], in hb G (19544)[K 12], in hb G (14543)[K 51] Basionym: Distribution: - syn nov Hattori Stephani 1972 1985 538 928 J Hatt Bot Lab 36: 119-120 Stephani Icon Hepat Inedit [Thyopsiella] F-167, microfiche 59 fig 4108 ["India orientalis (leg Pfleiderer) K 12, K 21, K 31"] = Frullania alstonii var. pfleidereri S.Hatt. 1972 dusenii 1905] Schiffn. ex Vanden Berghen 1976 [pro syn] {Cameroon} [non Steph. in Dusen Typus: Cameroon. Ekundu N'dene sd, Dusen sn in hb W 4998 [holotypus] Basionym: Distribution: - syn nov Schiffner in Vanden Berghen Wigginton Wigginton = Frullania spongiosa dussiana 1976 2004 2004 185 Bull Jard Bot Nat Belg 46 (1/2): 167 & 171 [syn sub Fr. spongiosa] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 234 [syn sub Fr spongiosa,West Africa] Steph. 1894 Steph. 1910 Typus: Martinique: -.11.1900, Duss 339 ex hb URBAN, in hb G [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Bonner Stephani 1910 1965 1985 686 655 928 Yuzawa Yuzawa 1991 1991 525 525 = Frullania rio-janeirensis Spec Hepat 4: 332-333 Index Hepat 5: 293 Stephani Icon Hepat Inedit [Chonanthelia] A- 29, microfiche 46 fig 3111 [from type; "Martinique 339"] J Hatt Bot Lab 70: 257, fig. 39 [redescription of type specimen] J Hatt Bot Lab 70: 254 [syn sub Fr. riojaneirensis] (Raddi) Ångstr. 1876 Page 126 duthieana [auct. non Steph. 1910] S.Hatt. in Hara 1966 Typus: Basionym: Distribution: - syn nov Hattori in Hara Hattori in Hara 1966 1971 841 190 Flora of Eastern Himalaya: 525 [Himalaya (Kidarkanta)] Univ Mus Univ Tokyo Bull 2: 234 = Frullania duthieana var. appendiculata S.Hatt. in Hara 1971 Frullania duthieana Steph. 1910 duthieana var. appendiculata S.Hatt. in Hara 1971 Typus: Bhutan, Shodu-Barshong, 3600m, abies forest 24.05.1967 in hb NICH 287259 [typus] Basionym: Distribution: India (North: Himachal Pradesh (Simla), Uttaranchal (Dwali, Kumaon), West Bengal (Darjeeling)), Sikkim, Bhutan var nov - syn nov Hattori in Hara Hattori Hattori in Ohashi Parihar, Lal & Katiyar 1971 1973 1975 1994 Bapna & Kachroo 2000 Nath & Saxena 2002 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania duthieana 190 541 187 2196 Univ Mus Univ Tokyo Bull 2: 234-236, fig 24 [Sikkim, Bhutan] J Hatt Bot Lab 37: 94-96, fig 50(a-l) Univ Mus Univ Tokyo Bull 8: 233 [Nepal] [syn sub Fr. duthiana] Hepat Anth India, annotated checklist: 21 [Eastern Himalaya] - new variety from Eastern Himalayas (Hattori 1971) 712 Hepaticology in India II: 169 2572 WCB Lucknow: 57 [Bhutan] 2112 Type sepecimens Hatt Bot Lab (NICH) 23: Holotype. Bhutan, Shodu Barshong. 3600 m. 24 May 1967. H. Kanai et al. (NICH 287259). Steph. 1910 eboracensis var. caduciloba R.M.Schust. [ex Swedish Museum Nat Hist] 2000 [nom herb in hb WS] Typus: USA in hb WS (= MARION OWNBEY Herbarium at Washington State University, Pullman, USA) [holotypus] Basionym: Distribution: USA Internet Internet (WS) 2000 2003 783 980 Internet - Swedish Museum of Natural History http://www.wsu.edu/~wsherb/typecoll.html = eboracensis var. maior Gottsche 1845 Basionym: Distribution: - syn nov Gottsche in G., L. & N. Winter = Frullania eboracensis 1845 3000 2850 Syn Hepat: 423 Lehm. 1844 Page 127 eboracensis ssp. caucasica (Steph.) R.M.Schust. 1992 [nom superfl] Basionym: Distribution: Caucasus, Ukraine, Spain, Switzerland, Italy - syn nov Schuster 1992 Winter 3000 = Frullania caucasica 603 Hepat N America 5: 143 [ev. conspecific with Fr. eboracensis subsp parvistipula] Steph. 1910 Frullania parvistipula eboracensis ssp. parvistipula Steph. 1910 (Steph.) R.M.Schust. 1992 Basionym: Distribution: Italy, Spain, Switzerland, ? Alaska ?, Russia (Caucasus, Siberia, Far East, Altai), Sakhalin, China (Hunan), Japan, India, Bhutan, Thailand nom superfl Schuster 1992 603 Frey, Frahm, Fischer & Lobin Rao, Enroth, Piippo & Koponen Grolle & Long Winter 1995 1997 2000 3000 606 685 760 = Frullania parvistipula echinata Hepat N America 5: 142 [stat nov: = Fr. eboracensis subsp parvistipula, excl Fr. caucasica] Kleine Kryptogamenflora 5: 116 [key to european species] Hikobia 12: 183 [checklist Hunan] J Bryol 22: 111 [checklist of Europe and Macaronesia] & 124 [Basionym is Fr. parvistipula sensu HATTORI 1978, incl Fr. caucasica] Steph. 1910 S.Hatt. by S.Hatt. 1982 [orth var] Basionym: Distribution: orth var Hattori Hattori 1982 1982 = Frullania echinantha echinella 422 495 Misc Bryol Lichen 9 (6): 124 J Hatt Bot Lab 51:249 S.Hatt. 1974 Colenso 1888 Typus: New Zealand. County od Waipawa, Dannevirke 1887, W. Colenso sn in hb E E00002569 [isotypus] Basionym: Distribution: spec nov - syn nov Colenso Stephani Bonner Hamlin = Frullania pycnantha 1888 1892 1965 1972 635 210 655 659 Trans Proc New Zealand Inst 20: 252 J Linn Soc Bot 29: 274 [syn sub Fr. "pyncnantha", orth error] Index Hepat 5: 294 Rec Dominion Mus 7(19): 261 [New Zealand] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Page 128 ecklonii [auct. non (SPR.) SPR.] Mont.] 1855 French Guyana Typus: Guyana gallica sd, Leprieur sn Basionym: Distribution: French Guyana - syn nov Montagne 1855 475 Gradstein & Hekking 1989 517 = Frullania arecae (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura ecklonii Ann Sci Nat Bot, ser 4, 3: 319 [Cryptogamia Guyanensis, Guyana gallica] J Hatt Bot Lab 66: 208 [French Guiana: Montagne 1840 as F. obscura; Montagne 1855 as F. hians and F. ecklonii] F. hians and F. ecklonii; (Sw.) Mont. 1840 SIM 1926 [hom illeg] [non (Spreng.) Gottsche, Lindenb. & Nees 1845] Basionym: Distribution: - syn nov Sim Demaret & Vanden Berghen Arnell = Frullania arecae 252 59 822 Trans Roy Soc S Africa 15: 38-39, figs [South Africa] Bull Jard Bot Etat Bruxelles 19 (1): 77 [syn sub Fr. arecae] Hepat S Africa: 136 [syn sub Fr. arecae] (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura ecklonii 1926 1948 1963 (Sw.) Mont. 1840 fo. a robustior Gottsche, Lindenb. & Nees 1847 Typus: Mexico. hb LIEBMANN 200d Basionym: Distribution: South Africa, Mexico -stat nov Gottsche, Lindenb. & Nees Gottsche Gottsche Sim 1847 1863 1867 1926 171 103 104 252 Synopsis Hept: 771 Separata of Mex. Leverm.: 236 [alpha robustior, Mexico] K Danske Vid Selks Skrifter, ser 5, 6: 332 [alpha robustior, Mexico] Trans Roy Soc S Africa 15: 39 [var. robustior] = Frullania ecklonii var. robustior (Gottsche, Lindenb. & Nees) Sim 1926 ecklonii fo. b tenerior Gottsche, Lindenb. & Nees 1847 Basionym: Distribution: South Africa - stat nov Gottsche, Lindenb. & Nees Sim 1847 1926 2848 Syn Hepat: 771 [ad 6 ß, Promontorio Bonae Spei] 252 Trans Roy Soc S Africa 15: 39 [South Africa] = Frullania ecklonii var. tenerior (Gottsche, Lindenb. & Nees) Sim 1926 ecklonii var. huitamalcensis Gottsche 1847 Typus: Mexico.Vera Cruz, Huitamalco, 2500-3000ft hb LIEBMANN 443 Basionym: Distribution: Mexico Gottsche Gottsche Gottsche = Frullania ecklonii 1847 1863 1867 141 103 104 Synopsis Hepat: 771 Separata of Mex. Leverm.: 236 K Danske Vid Selks Skrifter, ser 5, 6: 332 [Mexico] (Spreng.) Gottsche, Lindenb. & Nees 1845 Page 129 ecostata S.Hatt. by S.Hatt. 1978 [orth var] Basionym: Distribution: orth var Hattori Grolle & Piippo = Frullania erostrata ecuadorensis 1978 1984 482 143 J Hatt Bot Lab 44: 538-539 [perianth] Acta Bot Fennica 125: 18 [Cat. West Melanesia] S.Hatt. 1974 Steph. 1911 Typus: Ecuadoe. San José sd, Allioni in hb G (hb LEVIER 6551) Basionym: Distribution: Costa Rica, Cocos Isl [Costa Rica], Panama, Jamaica, Ecuador, Galapagos Isls ?? [Ecuador], Colombia, Venezuela, Peru, Brazil spec nov - syn nov Stephani Bonner Stotler Stotler Gradstein & Hekking Mägdefrau Yano Stephani 1911 1965 1969 1970 1979 1983 1984 1985 Morales Z. Schuster Wolf 1991 1992 1993 Stotler in Seaward & FitzGerald Stotler, Allen, Gradstein, McGuinness, Whittemore & Chung Uribe & Gradstein Dauphin Dauphin & Ilkiu-Borges Gradstein & Costa Parolly, Kürschner, SchäferVerwimp & Gradstein Holz 1996 1998 Spec Hepat 4: 526 [=sgen Frullania] Index Hepat 5: 294 Nova Hedwigia 18: 471-472, pl 35 (297-303] The genus Frullania, separata: 471-472, pl 35 (297-303] J Hatt Bot Lab 45: 112 [Cat Colombia] Nova Hedwigia 13: 21 [Colombia] J Hatt Bot Lab 56: 504 [checklist Brazil] Stephani Icon Hepat Inedit [Thyopsiella] F-106, microfiche 58 fig 4030 ["Ecuador, Allioni legit, Unicum, Levier 6551 ex parte"] 307 Trop Bryol 4: 37 [Costa Rica] 603 Hepat N America 5: 43 2380 Phytocoenologia 22(1): 29 [Colombia, Epiphyte - Lower montane rain forest communities: rare taxa ] 1092 Richard Spruce, Roy Bot Gard Kew: 134, fig 8 [subgen Chonanthelia] 703 Trop Bryol 15: 179 [Checklist Panama] 1998 1999 2002 2003 2004 707 1087 1072 984 1170 2004 Yano Gradstein 2008 2009 Söderström, Hagborg, Vana & Konrat Gradstein & Uribe Graham et al Bernal, Gradstein & Celis Gradstein & Perez 2011 1184 [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm 1684 Boletim Instituto Botanica 19: 31 [Brazil; catalog] 2161 Bibliotheca Lichenologica 99: 158 [Galapagos Isl ?, leg. Pittier (STOTLER 1969); since Pittier is not known to have collected on Galapagos, the specimen is probably mislabelled.] 2581 Bryologist 114(1) : 72 [checklist Jamaica] 2011 2015 2016 2021 3230 4766 4290 6056 = Frullania kunzei 686 655 4761 194 483 1206 501 928 Bryophyt Bibl 53: 36 [Catalogue Colombia] Rev Biol Trop 47 (3): 323 [Cocos Isl, Costa Rica] Trop Bryol 22: 118-119 [Cerro Venamo, Venezuela] Mem New York Bot Gard 87: 91 [Brazil] Crypt Bryol 25 (3): 275 [Ecuador] Caldasia 33(2): 361 [Colombia; synopsis of Frullaniacae & keys] Acta Biologica Plantarum Agriensis 4: 16 [Peru] – und andere Seiten Catálogo de plantas y líquenes de Colombia. Vol. I. Candollea 76: 44 [syn sub Fr kunzei - Ecuador] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 130 effusa Steph. 1924 Typus: Mexico 1913, Arsén 8001 in hb G 7795 [holotypus] Basionym: Distribution: Mexico spec nov - syn nov Stephani Bonner Stephani 1924 1965 1985 Fulford & Sharp Uribe & Gradstein Uribe 1990 2003 2008 = Frullania brasiliensis elegantissima Typus: Samoa 687 655 928 Spec Hepat 6: 548 Index Hepat 5: 294 Stephani Icon Hepat Inedit [Meteriopsis] E-20, microfiche 55 fig 3794 ["Mexico, legit Arsén 8001"] 105 Mem NY Bot Gard 63: 42 [Mexico] 1071 Crypt Bryol 24 (3): 204 [syn sub Fr brasiliensis] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Raddi 1822 Steph. 1911 sd, Rechinger, 3316pp Basionym: Distribution: Philippines (Luzon), Samoa (Tutuila) spec nov Stephani Pearson 1911 1924 686 863 - syn nov Verdoorn Verdoorn Bonner Grolle & Schultze-Motel Miller, Whittier & Whittier Stephani 1928 1930 1965 1972 1983 1985 242 551 655 537 609 928 = Frullania angulosa Spec Hepat 4: 575 Papers Dept Marine Biol Carnegie Inst Wash 20 (=341): 140, fig 14(610) (Samoa: Tutuila) Rev Bryol NS 1: 120 [syn sub Fr. angulosa] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 295 J Hatt Bot Lab 36: 83 [sub Fr. angulosa] [Samoa] Prod Flora Hepat Polynesiae: 125 Stephani Icon Hepat Inedit [Homotropantha] D- 2, microfiche 54 fig 3705 ["Samoa, Rechinger 3316pp"] - 3706 Mitt. in Seemann 1871 Frullania intermedia var. non-apiculata S.Hatt. 1975 elegantula Typus: Peru Spruce ex Bonner 1965 [nom herb] sd, J.C. Mutis in hb US Basionym: Distribution: Peru Bonner NN 1965 1996 655 632 Index Hepat 5: 295 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 13 of 35 [in hb US] = Frullania brasiliensis var. elegantula Spruce 1884 Page 131 elliotii Steph. 1894 [non Steph. 1911] Typus: Madagascar: Betsileo sd, Elliot 255 in hb G G00045252 - n° SIB 148734/1 - 13610 (ex hb K, hb STEPHANI 61 ) Basionym: Distribution: Madagascar spec nov - syn nov Stephani Stephani in Levier Stephani Demaret & Vanden Berghen Bonner Vanden Berghen Stephani 1894 1901 1911 1950 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania angulata elliottiana 228 1456 686 60 655 185 928 Hedwigia 33: 153 Rev Bryol 28 (5): 94 [Madagascar] Spec Hepat 4: 594-594 [vide # 420] Bull Jard Bot Etat Bruxelles 20: 5 Index Hepat 5: 295 Bull Jard Bot Nat Belg 46 (1/2): 70 Stephani Icon Hepat Inedit [Meteriopsis] E- 2, microfiche 54 fig 3763 ["Madagascar, Elliot (Kew)"] - 3764 ["aus dem Ex 596 von Forsyth Major"] 250 Bryophyt Bibl 48: 62 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Mitt. 1863 Steph. 1911 Typus: Dominica sd, Elliott, Hep dominicenses 1352 ex hb BM, in hb G Basionym: Distribution: spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 Fulford Schäfer-Verwimp 1987 2010 = Frullania neesii 686 655 4761 194 928 Spec Hepat 4: 527-528 Index Hepat 5: 295 Nova Hedwigia 18: 450, in syn The genus Frullania, separata: 450, in syn Stephani Icon Hepat Inedit [Thyopsiella] F-107, microfiche 58 fig 4031 ["Dominica, leg Elliot 1352"] 99 Mem NY Bot Gard 45: 272 [Dominica] 2460 Crypt Bryol 31(4): 347 [syn sub Frull kunzei, Dominica, West Indies] Lindenb. 1845 Frullania kunzei (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 132 elliptica Steph. 1911 Typus: Hawaii. Maui, Nabiku -.9.1909, Faurie 568 holotypus: in hb G-20552 Basionym: Distribution: Hawaii spec nov - syn nov Stephani Stephani Verdoorn 1911 1924 1930 Verdoorn Miller Miller Bonner Miller, Whittier & Whittier Stephani 1930 1953 1963 1965 1983 1985 Bapna & Kachroo So & Wang 2000 2006 = Frullania sandvicensis elongata 686 687 293 Spec Hepat 4: 677 Spec Hepat 6: 538 Ann Bryol suppl 1: 37 [syn sub Fr. sandvicensis; and Fr sandvicensis may be syn sub Fr squarrosa] 551 Nederl Kruidk Arch Jrg 1930 (2): 164 [sub Fr squarrosa] 349 Bryologist 56: 45 [sub Fr squarrosa] 834 Ark Bot, ser 2, 5: 518 [syn nov sub Fr. squarrosa] 655 Index Hepat 5: 295 609 Prod Flora Hepat Polynesiae: 125 928 Stephani Icon Hepat Inedit [Galeiloba] C-125, microfiche 52 fig 3347 ["Hawaii, Faurie 547"] 712 Hepaticology in India II: 170 [sub Fr. ericoides] 1466 J H B L 100: 428 [syn sub Fr sandvicensis] Ångstr. 1872 Steph. 1910 [non (Lehm. & Lindenb.) Gottsche IN G., L. & N. 1845] Typus: Australia. New South Wales, Richmond River, Pearce's Creek - NSW, Belmore Falls [syntype] -NSW Richmond River [isolectotype] 2.8.1900, W.W. Watts 96 - 5.10.1908, W.W. Watts 931 [syntype] - 11.7.1900, W.W. Watts 83 [isolectotype] in hb G 18203 (ex hb LEVIER 2629) - in hb NSW 249560 [syntypus] - in hb NSW 250712 [isolectotypus] Basionym: Distribution: Australia (NSW) spec nov - comb nov Stephani Bonner Hattori Hattori Hattori Stephani 1910 1965 1979 1979 1983 1985 686 655 485 485 499 928 Scott & Bradshaw Brown, Ramsay & Seur NN 1986 1992 2000 56 1140 1085 Carthy Glenny, Meagher & Brown 2003 2004 1125 1127 McCarthy 2006 1385 Spec Hepat 4: 423 Index Hepat 5: 296 J Hatt Bot Lab 45: 354 [syn sub Fr. rubella] J Hatt Bot Lab 45: 333-335, fig 8(a-j) from type J Hatt Bot Lab 54: 155-156 [comb nov as Fr. rubella var elongata] Stephani Icon Hepat Inedit [Galeiloba] C-204, microfiche 53 fig 3612 ["N.S.Wales, leg Watts 2629"] Brunnonia 8 (1): 34 [Australia] Telopea 5(1): 74-75 [types in hb NSW] http://plantnet.rbgsyd.nsw.gov.au/HerbLink/NSWtypes/srch.html [isolectotype & syntype in NWS] Flora Australia Suppl Ser 21: 46 [sub Fr rubella var elongata] Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Checklist Australian Liverworts [Australia, syn sub Fr "pubella" var elongata]] = Frullania rubella var. elongata (Steph.) S.Hatt. 1983 elongata (Lehm. & Lindenb.) Gottsche 1845 [pro syn] Basionym: Distribution: Lehmann & Lindenberg in G., L. 1845 & N. Bonner 1965 Wigginton 2004 2850 Syn Hepat: 428 655 Index Hepat 5: 296 1172 Sript Bot Belg 30: 227, fig 149(C,G,H) from type [syn sub Fr trinervis, West Africa] = Frullania trinervis var. elongata (L. & L.) Lehm. & Lindenb. 1845 Frullania trinervis (Lehm.) Drège 1843 Page 133 elongatistipula (Verd.) S.Hatt. 1972 Basionym: Frullania moniliata ssp. obscura var. elongatistipula Verd. in Hand.-Mazz. 1930 Distribution: comb nov - comb nov Hattori Hattori & Lin Piippo 1972 1985 1990 536 505 521 J Hatt Bot Lab 35: 241-243, fig 16 J Hatt Bot Lab 59: 161 J Hatt Bot Lab 68: 40 [annotated catalogue of China] = Frullania tamarisci ssp. obscura var. elongatistipula (Verd.) S.Hatt. 1985 emarginatula Steph. 1910 Typus: Indonesia. Java, Cibodas sd, Karsten in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Stephani 1910 1929 1930 1965 1985 = Frullania nigricaulis emergens 686 246 293 655 928 Spec Hepat 4: 447 Ann Bryol 2: 160 Ann Bryol Suppl 1: 153 [syn sub Fr. nigricaulis] Index Hepat 5: 296 Stephani Icon Hepat Inedit [Galeiloba] C-126, microfiche 52 fig 3516 [from type; "Java Tjibodas, leg G Karsten, Herb Jack ..."] (Reinw., Blume et Nees) Nees 1845 Mitt. 1863 Typus: Cameroons: Mt. Cameroon, 8000 ft -.1.1862, Mann sn in hb G [isotypus] Basionym: Distribution: Cameroons spec nov - syn nov Mitten Stephani Sim Demaret & Vanden Berghen Arnell Stephani 1863 1910 1926 1948 1963 1985 Yuzawa Wigginton 1991 2002 Wigginton 2004 = Frullania ecklonii emergens 253 686 252 59 822 928 J Linn Soc Bot 7: 168 Spec Hepat 4: 320-321 Trans Roy Soc S Africa 15: 38-39 [syn nov sub Fr. eckloni] Bull Jard Bot Etat Bruxelles 19 (1): 77 Hepat S Africa: 138 [syn sub Fr. ecklonii] Stephani Icon Hepat Inedit [Chonanthelia] A- 5, microfiche 46 fig 3070 [from type; "Kamerun leg Mann 1862"] 525 J Hatt Bot Lab 70: 259 [as "emargens" syn. nov., syn sub Fr. eckloni] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] (Spreng.) Gottsche, Lindenb. & Nees 1845 Steph. by Pearson 1924 [orth var] Basionym: Distribution: Pearson = Frullania emergens Frullania ecklonii 1924 795 Arkiv Botanik 19 (5): 9 [Mt. Elgon in Kenia/Uganda] Mitt. 1863 (Spreng.) Gottsche, Lindenb. & Nees 1845 Page 134 engelii S.Hatt. 1983 Typus: Tasmania. Florentine River, 380m 5.3.1977, John J. Engel 15080 in hb NICH [holotypus] in hb F 1032742 [isotypus] Basionym: Distribution: Tasmania, Kermadec Isl [New Zealand] spec nov -syn sov Hattori Scott & Bradshaw Ratkowsky 1983 1986 1987 499 56 695 Jarman & Fuhrer Glenny Carthy Glenny, Meagher & Brown 1995 1998 2003 2004 78 1412 1125 1127 Glenny & Malcolm 2005 1410 McCarthy 2006 Mizutani, Hasegawa & Iwatsuki 2009 1385 2112 Hentschel et alii 4097 = Frullania setchellii engleri 2015 J Hatt Bot Lab 54: 143-146, fig 55 Brunnonia 8 (1): 34 & 143 [Tasmania] Papers Proceed Roy Soc Tasmania 121: 157 [Checklist Tasmania] & 158 [Hattori per comm (II): may be conspecific with Fr setchellii & Fr pycnantha, endemic to NZ] Mosses liverworts rainforest Tasmania: 123 Tuhinga 10: 121, 132 [revised checklist of New Zealand] Flora Australia Suppl Ser 21: 43 [Australia (TAS)] Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand species] Checklist Australian Liverworts [Australia TAS] Type sepecimens Hatt Bot Lab (NICH) 23: Holotype. Tasmania, Florentina River. 380 m. 5 Mar. 1977. J. J. Engel 15080 (NICH 430199). Phytotaxa 220 (2): 137 [ syn sub Fr setchellii] Pearson 1923 Steph. ex Bonner 1965 [nom herb] Typus: India orientalis sd, Pfleiderer K 20 in hb G Basionym: Distribution: India nom herb - syn nov Bonner Hattori Stephani = Frullania apiculata engleriana 1965 1972 1985 655 538 928 Index Hepat 5: 296 [nom nud ?] J Hatt Bot Lab 36: 112-113, fig. 3 Stephani Icon Hepat Inedit [Thyopsiella] F-139, microfiche 59 fig 4075 ["India orientalis, Pfleiderer K 20"] (Reinw., Blume & Nees) Nees 1845 Steph. ex Steph. 1985 [nom herb] {New Guinea} Typus: in hb G Basionym: Distribution: New Guinea Stephani 1985 928 Stephani Icon Hepat Inedit [Thyopsiella] F-140, microfiche 59 fig 4076 [Nova Guinea (Ledermann legit)"] = Page 135 epiphylla Vanden Berghen 1976 [hom illeg] [non S.Hatt. 1974] Typus: Madagaskar: , 900 m 30.3.1974, Cremers 3035 in hb ONRAEDT [holotypus] Basionym: Distribution: Madagascar spec nov - nom nov nom nov Vanden Berghen Bizot & Pocs Hattori Wigginton & Grolle Wigginton 1976 1979 1982 1996 2002 Wigginton 2004 = Frullania vandenberghenii eplicata 185 791 495 696 1012 Bull Jard Bot Nat Belg 46 (1/2): 30-32, fig 7(A-H) Acta Bot Acad Sci Hung 25 (3/4): 229 as Fr. vandenberghenii J Hatt Bot Lab 51: 249 (foot note) as Fr. vanden-berghenii Bryophyt Bibl 50: 86 [nom illeg] Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Pócs 1979 Steph. 1913 [nom superfl] Basionym: Distribution: Stephani in Brunnthaler = Frullania eplicata equadorensis 1913 219 Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 728 [spec nov = nom superfl] Steph. 1911 Steph. by Cole 1983 [orth var] Basionym: Distribution: Cole 1983 = Frullania ecuadorensis Frullania kunzei ericicola 1407 Brenesia 21: 154 [Central America] Steph. 1911 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. ex Bonner 1965 [nom herb] Typus: Kilimandscharo, 3300m 1901, Uhlig 5 in hb G Basionym: Distribution: - syn nov Bonner Vanden Berghen 1965 1976 Stephani 1985 Wigginton 2004 = Frullania variegata ericioides 655 185 Index Hepat 5: 297 [East Africa] Bull Jard Bot Nat Belg 46 (1/2): 13 [Stephani Icon ined no 1722; syn sub Fr. variegata] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 20, microfiche 56 fig 3898 ["Kilimandscharo, Uhlig No 5, 3300m"] 1312 Trop Bryol Res Rep 5: 30 [perhaps syn sub Fr trinervis] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Steph. 1894 Nees in Verd. 1930 [orth var] Basionym: Distribution: Verdoorn = Frullania ericoides 1930 293 Ann Bryol Suppl 1: 34 [ericoides in all combinations written as "ericioides"] (Nees) Mont. 1839 Page 136 ericoides var. minor Kamim. 1970 Typus: Japan Basionym: Distribution: var nov - syn nov Kamimura Hattori & Lin Bapna & Kachroo Yuzawa Zhu & So 1970 1985 2000 2001 2001 Hong Yamada & Iwatsuki 2003 2006 = Frullania ericoides 1552 505 712 1660 808 Bulletin of Kochi Gakuen Junior College 2: 22-23, fig. 3 (a-l) [Japan] J Hatt Bot Lab 59: 133 [syn nov sub Fr. ericoides] Hepaticology in India II: 170 [syn sub Fr. ericoides] Natural Environmental Sci Res 14: 13 [Japan, syn sub Fr ericoides] Nova Hedwigia Beiheft 121: 73-5, fig 32(a-j) [Epiphyllous liverworts of China] 1416 Lindbergia 28: 145 [syn sub Fr ericoides] 1470 J Hatt Bot Lab 99: 29 [Catalog of Japan] (Nees) Mont. 1839 ericoides var. planescens (Verd.) S.Hatt. 1984 Basionym: Frullania squarrosa var. planescens Verd. 1929 Distribution: China (Yunnan, Hunan, Xizang, Guangdong), Malaysia (Malay Pen), Indonesia (Sumatra, Java, Mollucas), New Guinea, Bismarck Arch (New Britain [=Neupommern]), New Caledonia comb nov - syn nov Hattori Hattori & Lin Hattori Piippo Rao, Enroth, Piippo & Koponen Piippo, He, Koponen, Refearn & Li Bapna & Kachroo Bai in Gao & Cao Bai Bai = Frullania ericoides 1984 1985 1986 1990 1997 1998 503 505 506 521 685 697 J Hatt Bot Lab 57: 412 J Hatt Bot Lab 59: 134 [China] J Hatt Bot Lab 60: 212 J Hatt Bot Lab 68: 40 [annotated catalogue of China] Hikobia 12: 183 [checklist Hunan] J Hatt Bot Lab 84: 143 [checklist of Yunnan, China] 2000 2000 2001 2002 712 1019 1742 1189 Hepaticology in India II: 170 [syn sub Fr. ericoides] Fl Yunnanica 17: 386-387 [China, Yunnan] Guizhou Science 19(4): 36 [China, Yunnan] Chenia 7: 7 [Yunnan, China] (Nees) Mont. 1839 ericoides var. squarrosa (Nees) Mont. 1839 Typus: Java Basionym: Jungermannia squarrosa Distribution: Peru var nov Reinw., Blume & Nees 1824 Montagne Schiffner = Frullania ericoides 1839 1898 465 831 Ann Sci Nat Bot, ser 2, 12: 51 (No 105, Java) Conspect Hepat Archip Indici, Batavia: 344 [syn sub Fr. squarrosa] (Nees) Mont. 1839 ericoides var. ß laxa Gottsche, Lindenb. & Nees 1845 [p. 417, Wight, Indien] Typus: in hb GLM (hb LEHMANN) 05992 Basionym: Frullania squarrosa var. γ laxa Gottsche, Lindenb. & Nees 1847 [p. 417, Wight, India - pro syn.] Distribution: var nov Gottsche, Lindenb. & Nees Sande Lacoste Schiffner Bonner 1845 1857 1898 1965 2850 430 831 655 Syn Hepat: 417 [Wight, Indien] Verh Konikl Akad Wetenschappen 5: 79 Conspect Hepat Archip Indici, Batavia: 324 Java] Index Hepat 5: 297 [ericoides var. laxa: India, Java] = Frullania ericoides var. ß laxa (Gottsche, Lindenb. et Nees) Schiffn. 1898 Page 137 erosolimbata Steph. 1924 Typus: Tansania. Ukambani sd, G. Scheffler 215a hb WILMS 215a, ex hb LINDAU, in hb G 17307 [holotypus] & Basionym: Distribution: spec nov Stephani Bonner Vanden Berghen Stephani 1924 1965 1976 1985 Wigginton 2002 Wigginton 2004 Spec Hepat 6: 538-539 Index Hepat 5: 297 Bull Jard Bot Nat Belg 46 (1/2): 178 & 184 Stephani Icon Hepat Inedit [Galeiloba] C- 16, microfiche 50 fig 3367 [from type; "Africa orientalis, Ukambani, leg Georg Scheffler, ex Herb Dr Wilms 215a"] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 686 655 185 928 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 Spec Hepat 4: 672 Index Hepat 5: 298 Bull Jard Bot Nat Belg 46 (1/2): 206 Stephani Icon Hepat Inedit [Chonanthelia] A- 12, microfiche 46 fig 3081-3082 [from type; "Madagascar 1877 .."] 250 Bryophyt Bibl 48: 65 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 226 [syn sub Fr depressa, West Africa] 1911 1965 1985 686 655 928 = Frullania caffraria erronea 687 655 185 928 Steph. 1894 Steph. 1911 Typus: Madagascar 1877, Borgen 986 ex hb PEARSON, in hb G 13012 [holotypus], G: Basionym: Distribution: Madagascar spec nov - syn nov = Frullania depressa evansii Typus: Cuba Mitt. 1863 Steph. 1911 sd, s coll & n ex hb KUNZE, ex hb LZ, in Hb G Basionym: Distribution: Cuba spec nov Stephani Bonner Stephani = Frullania kunzei Spec Hepat 4: 634 Index Hepat 5: 298 Stephani Icon Hepat Inedit [Diastaloba] B-30, microfiche 48 fig 3263 ["Cuba (Herb Univ Lipsiensis) (sub Fr. kunzei)"] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 138 evoluta Mitt. 1861 [HOOKER No 1519 in hb G(15401)] vide S.Hatt. 1974 Typus: Himalaya, 7-8000f sd, J. D. Hooker No 1519 No 1519 = in hb G15401, in hb KEW Basionym: Distribution: spec nov - syn nov Mitten Stephani Stephani Hattori 1861 1911 1911 1974 574 686 686 545 Hattori 1975 47 = Frullania nepalensis J Linn Soc Bot 5: 122 Spec Hepat 4: 557 [= Fr. evoluta, sgen Frullania Spec Hepat 4: 451 [= Fr. nepalensis, sgen Trachycolea] J Hatt Bot Lab 38: 215-217, fig. 103 [HOOKER No 1519 in hb G(15401)] Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 76 [about Mitten´s specimen] (Spreng.) Lehm. & Lindenb. 1845 evoluta var. tagawana S.Hatt., Thaithong & Kitagawa 1977 Typus: Thailand: Chiangrai, Doi Phahompok, 2100 m sd, Kitagawa 15340a in hb NICH [typus] Basionym: Distribution: India (North: West Bengal (Darjeeling), Assam, Meghalaya), Thailand, Laos var nov comb et stat nov spec nov Hattori, Thaithong & Kitagawa Hattori & Thaithong Hattori & Thaithong Hattori & Lin 1977 1978 1978 1985 480 128 481 505 Singh 2002 4886 Studies on Liverworts (Bryophyta) of Khasi and Jaintia Hills: Meghalaya. 274-275, map 11 - Unpublished thesis. 1281 Taiwania 53 (1): 75, fig 1[map], [India, Meghalaya, Eastern Himalayas] 2112 Type sepecimens Hatt Bot Lab (NICH): 24: Holotype. Thailand, Chiangdrai, Doi Phahompok. 2100 m. On tree trunk. 12 Sept. 1967. N. Kitagawa 15340a (NICH). Singh, Kumar & Nath 2008 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania tagawana evolutiloba J Hatt Bot Lab 43: 441-443, fig. 1 [Thailand] J Jap Bot 53 (6): 172 [Laos] J Hatt Bot Lab 44: 182-183, fig. 4(a-i) J Hatt Bot Lab 59: 160 [as Fr. tagawana] (S.Hatt. & Thaithong) S.Hatt. 1985 Steph. ex Herzog 1916 Typus: Bolivia. Florida de San mateo, 2200m 1911, T. Herzog 3671 in hb G Basionym: Distribution: spec nov - syn nov - syn nov Stephani Stephani Stotler Stotler Stephani 1916 1924 1969 1970 1985 222 687 4761 194 928 Gradstein, Meneses & Arbe 2003 975 = Frullania intumescens Biblioth Bot 87: 244, fig 195 [Bolivia] Spec Hepat 6: 546-547 [as spec nov] Nova Hedwigia 18: 444 [syn nov: sub Fr. intumescens] The genus Frullania, separata: 444 [syn nov: sub Fr. intumescens] Stephani Icon Hepat Inedit [Thyopsiella] F- 55, microfiche 57 fig 3951 ["Bolivia, Dr Herzog 3671 (1913)"] - 3952 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. intumescens] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 139 excisostipula Steph. ex Bonner 1965 [nom herb] Typus: India. South Canara sd, Pfleiderer K2 in hb G 14408 Basionym: Distribution: nom herb - syn nov Bonner Hattori 1965 1976 655 67 Hattori & Lin Stephani 1985 1985 505 928 Bapna & Kachroo 2000 712 Index Hepat 5: 299 [nom nud ?, nom herb ?] Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21 [as Fr. tamarisci ssp. obscura var. breviramea] J Hatt Bot Lab 59: 161 [syn sub Fr. tamarisci ssp. obscura] Stephani Icon Hepat Inedit [Thyopsiella] F-141, microfiche 59 fig 4077 ["India orientalis, South Canara, Pfleiderer K 2"] Hepaticology in India II: 196 [syn sub Fr. tamarisci ssp obscura] = Frullania tamarisci ssp. obscura var. breviramea (Steph.) S.Hatt. 1972 Frullania moniliata excisula (Reinw., Blume & Nees) Mont. 1842 Steph. in Steph. & Watts 1914 Typus: Australia. NSW, Mt. Wilson, Yarrawa [syntype1] - NSW, Horseshoe Falls, Jungle, Blackheath "rock by track" [syntype2] W. W. Watts 1071 [syntype1] - 7.1.1911, W.W. Watts 1030 [syntype2] 11.1.1911, in hb G [isosyntypus1] - in hb NSW 250735 [syntypus1] - in hb NSW 250716 [syntypus2] - in hb MEL Basionym: Distribution: Australia (NSW) spec nov Stephani in Stephani & Watts Bonner Hattori 1914 1965 1979 220 655 485 - syn nov Hattori Stephani 1983 1985 499 928 Scott & Bradshaw Brown, Ramsay & Seur NN 1986 1992 2000 56 1140 1085 Carthy McCarthy 2003 2006 1125 1385 = Frullania falciloba exciuscula J Proc Roy Soc New S Wales 48: 107 [Australia] Index Hepat 5: 299 ["Watts 1017" error = 1071] J Hatt Bot Lab 45: 335-336, fig 9(a-i) from type [orth error as "exciuscula"] J Hatt Bot Lab 54: 146 Stephani Icon Hepat Inedit [Galeiloba] C-205, microfiche 53 fig 3613 ["Australia, Watts No 1071"] Brunnonia 8 (1): 34 [Australia] Telopea 5(1): 75 [types in hb NSW & hb SYD] http://plantnet.rbgsyd.nsw.gov.au/HerbLink/NSWtypes/srch.html [syntypes in NWS; as syn sub Fr squarrosula] Flora Australia Suppl Ser 21: 43 [sub Fr falciloba] Checklist Australian Liverworts [Australia, ?? syn sub Fr falciloba] Lehm. 1844 Steph. by S.Hatt. 1979 [orth var] Basionym: Distribution: Hattori = Frullania excisula 1979 485 J Hatt Bot Lab 45: 335-336, fig 9(a-i) from type Steph. in Steph. & Watts 1914 Frullania falciloba Lehm. 1844 Page 140 exigua Steph. 1910 Typus: South Africa. Transvaal ? Vanderyst 140 ex hb Brotherus Basionym: Distribution: spec nov - syn nov ? / Zaire. "Congo Belge, Mayidi" , Bas-Zaire ? / -.3.1907, / in hb BR [isotypus] Stephani Stephani in Brunnthaler 1910 1913 686 219 Sim Demaret Demaret & Vanden Berghen Arnell Vanden Berghen Stephani 1926 1942 1947 1963 1976 1985 252 1066 1101 822 185 928 Scott & Bradshaw Windolf 1986 1987 56 1025 Wigginton & Grolle Wigginton 1996 2002 696 1012 Carthy Wigginton 2003 2004 1125 1172 Glenny, Meagher & Brown 2004 1127 Wigginton McCarthy 2004 2006 1312 1385 = Frullania cardotii sd, Mac Lea 6063 Spec Hepat 4: 367 Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 729 [Natal, Kapland, Deutsch-Ostafrika] Trans Roy Soc S Africa 15: 45 [not from Transvaal] Bull Jard Bot Etat Bruxelles 16 (2): 304 Bull Jard Bot Etat Bruxelles 18: 233 [syn sub Fr cardoti] Hepat S Africa: 153-154, fig 112(a-e) Bull Jard Bot Nat Belg 46 (1/2): 145 Stephani Icon Hepat Inedit [Galeiloba] C- 17, microfiche 50 fig 3368 [from type; "Transvaal, Mac Lea No 60 u 63 (Herb Brotherius)"] Brunnonia 8 (1): 34 [Australia] Austrobaileya 2(4): 390 [checklist Queensland; certain a misidentification by BAILEY 1890 for Australia] Bryophyt Bibl 50: 87 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Flora Australia Suppl Ser 21: 43 [Australia (QLD, NSW)] Sript Bot Belg 30: 233, fig 153(A) from type [syn sub Fr obscurifolia, West Africa] Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Checklist Australian Liverworts [Australia QLD NSW] Steph. 1910 Frullania obscurifolia Mitt. 1879 Page 141 exilis Taylor 1846 Typus: British Guiana (=Guyana), Georgtown, Demerara river [Demarara bei BONNER] - hb E "Growing on Jung. aurita, allied to J. tamarisci" in hb GREVILLE, typus in hb S (B22826) in hb E E00002530 [holotypus] in hb GLM (hb GREVILLE) 05993 Basionym: Distribution: Costa Rica, Puerto Rico, Trinidad, Bolivia; Colombia, Guyana, French Guiana, Brazil spec nov syn nov - syn nov Taylor Taylor in G., L. & N. Spruce Schiffner 1846 1847 1884 1895 Schiffner in Engler & Prantl Stephani Pagan Clark & Svihla Clark Clark Arnell Schiffner & Arnell Fulford, Crandall & Stotler Yano Stephani 1905 1911 1939 1949 1955 1955 1959 1964 1971 1984 1985 Gradstein Yuzawa & Koike Gradstein & Hekking Boggan, Funk, Kelloff et al. Weaver Gradstein, Lücking, Morales Z. & Dauphin Sillett. Gradstein & Griffin Ill Yano Boggan, Funk, Kelloff, Hoff, Cremers & Feuillet Yano Gradstein, Griffin III, Morales & Nadkarni Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Gradstein, Meneses & Arbe Gradstein & Costa Holz 1989 1989 1989 1992 1994 1994 London J Bot 5: 405-406 Syn Hepat: 778-779 Trans Proc Bot Soc Edinburgh 15: 55-57 Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 134 809 Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 134 686 Spec Hepat 4: 629-630 931 Bryologist 42: <1-12, 37-50, 71-82> [Puerto Rico] 330 Bryologist 52: 97-99, fig 1-19 369 Bryologist 58: 72 [Fr apiculata & Fr exilis may be one species] 369 Bryologist 58: 70 & 72 [similar to Fr. apiculata] 448 Svensk Bot Tidskr 53 (4): 505 [French Guiana] 198 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 86 [Brazil] 662 J Arnold Arboret 52: 451 [Puerto Rico] 501 J Hatt Bot Lab 56: 504 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Diastaloba] B-18, microfiche 48 fig 3247 ["Para, leg Spruce"] -3248 ["Brasilia, Berchtold, Jack 60"] 20 Bryologist 92 (3): 345 [ exilis sensu Pagán 1939 = Fr. apiculata] 519 J Hatt Bot Lab 66: 349-359 517 J Hatt Bot Lab 66: 208 [may be syn to Fr. apiculata] 4771 Checklist of the plants of the Guianas 4271 Bano de Oro Natural Area Luquillo Mountains, Puerto Rico 4 Lindbergia 19: 76 [Costa Rica] [may be conspecific with Fr. apiculata] 1995 1995 1997 2855 The Bryologist 98: 251-260 [Catalogue of Costa Rica] 2095 J Hatt Bot Lab 78: 142 [annotated checklist of Brazil] 1396 Checklist Guianas: 10 [Guyana, French Guiana] 1999 2000 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] 1405 Caldasia 23(1): 208 [Costa Rica: Monteverde] 2002 782 2003 2003 2004 Dauphin 2005 Yano Gradstein & Ilkiu-Borges 2008 2009 Gradstein & Uribe-M 2011 Gradstein 2013 Bernal, Gradstein & Celis Lima et al. Lima 2016 2018 2019 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. apiculata] 984 Mem New York Bot Gard 87: 83 [Brazil] [syn of Fr. apiculata] 1184 [checklist of Costa Rica; Gradstein & Solano 9517 (U,USJ)] http://asbot.botanik.uni-greifswald.de/personal/holz/bryo_cr/blank.htm 1360 Tropical Bryology 26: 141-218 [Catalogue of Costa Rican Hepaticae and Anthocerotae] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr apiculata; Brazil; catalog] 1920 Memoirs of The New York Botanical Garden 76: 43 [The synonymy of F. apiculata and F. exilis is provisional , needs further study] 3230 Caldasia 33: 354 [Fr exilis and Fr. apiculata has been considered a synonym, needs confirmation] 3864 Polish Botanical Journal 58: 176. [F. apiculata by its much smaller size and may be referred to F. exilis Taylor., needs further study] 4290 Catálogo de plantas y líquenes de Colombia. Vol. I. 5025 Rodriguésia 69 (3) 974-977, fig 2(a-i), 3(a-g), key 5121 Frullania no Brasil [Thesis]: 142-145, fig 9(a-q) fig. 10 [distribution] = Frullania apiculata 292 2848 243 1015 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] (Reinw., Blume & Nees) Nees 1845 Page 142 exipulifera Spruce ex Spruce 1884 [nom herb] Basionym: Distribution: Spruce 1884 243 Trans Proc Bot Soc Edinburgh 15: 21 = Frullania viminicola var. bicolor Spruce 1884 Frullania ecklonii expansa (Spreng.) Gottsche, Lindenb. & Nees 1845 Steph. 1897 Typus: Brazil, Rio de Janeiro 1892, C.A.M. Lindman 21 in h G (ex hb S-PA) G00067442 [holotypus] Basionym: Distribution: Brazil spec nov - syn nov Stephani Stephani Schiffner & Arnell Bonner Yano Stephani 1897 1910 1964 1965 1984 1985 Yano Gradstein & Costa Yano 1999 2003 2008 = Frullania kunzei exparsa 1059 686 198 655 501 928 Bihang Kgl Sv Vetensk-Akad Handl bot 23 III (2): 18-19 [Brazil] Spec Hepat 4: 386 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 91 [Brazil] Index Hepat 5: 300 J Hatt Bot Lab 56: 504 [checklist Brazil] Stephani Icon Hepat Inedit [Galeiloba] C- 59, microfiche 50 fig 3415 [from type; "Rio Janeiro, C A M Lindman leg No 21"] 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] 984 Mem New York Bot Gard 87: 87 [Brazil] [syn sub Fr kunzei] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr kunzei; Brazil; catalog] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. by Yano 1984 [orth var] Basionym: Distribution: Yano = Frullania expansa Frullania kunzei explanata 1984 501 J Hatt Bot Lab 56: 504 Steph. 1897 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Mitt. 1861 Typus: Sri Lanka [Insula Ceylon], Pas-dum-Korle -.12.1848, Gardner 1527 in hb NY 1048674 (hb MITTEN) [holotypus] in hb K Basionym: Distribution: spec nov - syn nov Mitten Stephani Verdoorn Bonner Hattori = Frullania intermedia 1861 1911 1930 1965 1975 574 686 293 655 47 J Linn Soc Bot 5: 121 [Sri Lanka] Spec Hepat 4: 589 Ann Bryol Suppl 1: 161 [syn sub Fr. intermedia] Index Hepat 5: 300 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 76 [Ann Bryolout Mitten´s specimen] (Reinw., Blume & Nees) Nees 1845 Page 143 explicata Mont. 1843 Typus: Sandwich Isles "Ad fronds Filicium in insulis Sandwich a celeb. Gaudichaud lecta" / Hawaii, Bonite ?? sd, Gaudichaud sn in hb PC 0052627 [holotypus] in hb G & PC-0033376 [isotypus] Basionym: Distribution: Philippines, Society (Tahiti), Hawaii spec nov - syn nov Montagne Montagne in G., L. & N. Montagne Reichardt Mitten in Seemann Stephani Bescherelle Evans Merrill Schiffner Stephani Verdoorn Miller Miller Bonner Miller, Whittier & Whittier Stephani 1843 1845 1856 1870 1871 1897 1898 1900 1906 1910 1911 1930 1953 1963 1965 1983 1985 Tan & Engel 1986 Bapna & Kachroo Konrat & Braggins So & Wang 2000 2001 2006 = Frullania apiculata 470 2850 1912 884 637 214 871 236 1496 436 686 293 349 834 655 609 928 Ann Sci Nat Bot, ser 2, 19: 256-257 Syn Hepat: 452 Sylloge generum specierumque cryptogamaram: 87 Reise österr Fregatte Novara, Bot Theil, Bd 1: 153 [Tahiti] Flora Vitiensis: 417 [Sandwich Isl] Bull Herb Boissier 5: 842[:260] [Hawaii] J Bot Morot 12: 138 [Tahiti] Trans Connet Acad Arts Sci 10 (1): 400-401 [syn nov sub Fr. apiculata] Philippine J Sci C, Bot 5: 311 [Philippines: Mt. Pulog] Österr Bot Zeitschr 60 (11): 431-432 Spec Hepat 4: 542 Ann Bryol Suppl 1: 96 Bryologist 56: 45 [sub Fr apiculata] Ark Bot, ser 2, 5: 519 [syn sub Fr. hypoleuca] Index Hepat 5: 300 Prod Flora Hepat Polynesiae: 125 Stephani Icon Hepat Inedit [Thyopsiella] F-129a, microfiche 58 fig 4056 ["Frull explicata Netze, Hawai, Gaudichaud, Original von explicata"] 4286 J. Hatt. Bot. Lab.. No. 60: 335 Hattori (1985: in !itt.) believes that the Philippine records of F. explicata are erroneous. 712 Hepaticology in India II: 191 [sub Fr. apiculata] 838 J Hatt Bot Lab 91: 256-262 [sub Fr. apiculata] 1466 J H B L 100: 420 [types] (Reinw., Blume & Nees) Nees 1845 eymae var. dentistipula S.Hatt. 1978 Typus: New Guinea [PNG]. Morobe: East of Bulolo, 800-1000m 7.12.1974, G. Shea 6572 in hb NICH [holotypus], in hb SHEA [isotypus] Basionym: Distribution: New Guinea [endemic] var nov - syn nov Hattori Hattori Grolle & Piippo Hong Mizutani, Hasegawa & Iwatsuki = Frullania eymae 1978 1982 1984 2003 2009 482 495 143 1416 2112 J Hatt Bot Lab 44: 527-529, fig 202 J Hatt Bot Lab 51: 250 Acta Bot Fennica 125: 19 [Cat. West Melanesia] Lindbergia 28: 138 [identification key to taxa of Korean paninsula] Type sepecimens Hatt Bot Lab (NICH) 24: Holotype. Papua New Guinea, Bulolo. 800-1000 m. 7 Dec. 1974. C. Shea 6572 (NICH 345212). S.Hatt. 1975 Page 144 falcata Steph. 1910 Typus: New Zealand sd, Knight 601 Basionym: Distribution: New Zealand spec nov - syn nov Stephani Martin 1910 1946 Martin 1946 Bonner Hamlin Hattori Stephani 1965 1972 1983 1985 = Frullania falciloba falciflora 686 Spec Hepat 4: 410 1423 Trans Roy Soc New Zealand 76 (1): 49 [New Zealand, South Isl: Arthur Pass NP] 1423 Transact Roy Soc NZ 76(1): 49 [New Zealang; Arthur Pass National Park] 655 Index Hepat 5: 300 659 Rec Dominion Mus 7(19): 261 [New Zealand] 499 J Hatt Bot Lab 54: 146 [syn sub Fr falciloba] 928 Stephani Icon Hepat Inedit [Galeiloba] C-261ab, microfiche 54 fig 3682 ["olim Frull knightiana, N Zealand, Knight 601"] - 3683 ["Frull falcata Stz, Frull knightiana St, Frull scabriseta Col, Trans NZ Inst vol 19, Hepa 1382, ob scandens?"] Lehm. 1844 Herzog ex Bonner 1965 [nom herb] {Costa Rica} Typus: Costa Rica. Prov S. José, La Palma, 1600m 17.3.1924, P. C. Standley 38016 in hb US Basionym: Distribution: Costa Rica Bonner NN 1965 1996 Jolley & Milne 1998 655 632 Index Hepat 5: 300 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 14 of 35 [in hb US] 2423 Australasian bryol newsletter 39: 10 [Australia, Victoria, Grampians NP] = falciloba [auct. non Taylor ex Lehm. 1844] Mitt. in Seemann 1857 {Panama} Basionym: Distribution: Mitten in Seemann 1857 1494 Bot Voyage HMS Herald: 246 [Panama] = falciloba Colenso ex Bonner 1965 [nom herb] Basionym: Distribution: New Zealand - syn nov Stephani Bonner = Frullania colensoana Frullania falciloba 1910 1965 686 655 Spec Hepat 4: 405 [not as syn of Fr. colensoana] Index Hepat 5: 301 [error: not as syn sub Fr. colensoana in STEPH 1910] Steph. 1910 Lehm. 1844 Page 145 falciloba [auct. non Taylor ex Lehm. 1844] Sande Lac. 1857 {Java} Typus: hb SANDE LACOSTE, in hb LUGD BAT Basionym: Distribution: Sande Lacoste Verdoorn 1857 1929 430 246 Verh Konikl Akad Wetenschappen 5: Ann Bryol 2: 157 [Fr. falciloba ex Java = Fr. squarrosa var. planescens] = Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides (Nees) Mont. 1839 falciloba var. setchellii (Pearson) E.A.Hodgs. 1949 [nom inval] Basionym: Frullania setchellii Distribution: New Zealand Pearson 1923 Hodgson Hamlin Hattori Glenny Glenny & Malcolm = Frullania setchellii fallax 1949 1973 1983 1998 2005 203 656 499 1412 1410 Trans Roy Soc New Zealand 77 (3): 367-368 [comb et stat nov] Rec Dominian Mus 8 (10): 143 [New Zealand] J Hatt Bot Lab 54: 168-170 Tuhinga 10: 132 [sub Fr setchellii] ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr setchellii] Pearson 1923 Steph. 1911 [hom illeg] [non Gottsche in G., L. & N. 1845] {Bolivia, Guatemala} Typus: Bolivia sd, Herzog 7 hb LEVIER 5848 & 5854 in hb G Basionym: Distribution: spec nov Stephani Stephani Bonner Stotler Stotler Gradstein, Meneses & Arbe -syn nov = Frullania intumescens fallax 1911 1924 1965 1969 1970 2003 686 909 655 4761 194 975 Spec Hepat 4: 495 [= sgen Frullania] Spec Hepat 6: 570 [Fr fusco-brunnea St. Mss in herb = Fr fallax] Index Hepat 5: 301 Nova Hedwigia 18: [syn sub Fr intumescens ?] vide 975 The genus Frullania, separata: [syn sub Fr intumescens ?] vide 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr intumescens] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Gottsche ex Steph. 1911 [non Gottsche in G., L. & N. 1845] {Java} Typus: Java Basionym: Distribution: Gottsche ex Stephani = Frullania fallax 1911 686 Spec Hepat 4: 577 Gottsche 1845 [non Austin ex Steph. 1911] Page 146 fallax [auct. non Gottsche] Sande Lac. 1857 {Java} Typus: Fr. fallax ex Java, hb SANDE LACOSTE in hb LUGD BAT Basionym: Distribution: - syn nov Sande Lacoste Schiffner 1857 1898 430 831 Verdoorn 1929 246 Verh Konikl Akad Wetenschappen 5: [specimen ex Java] Conspect Hepat Archip Indici, Batavia: 325 [steril specimen from Java may belong to another species] Ann Bryol 2: 157 = Frullania integristipula var. emarginata Verd. 1929 fallax var. magnistipula S.Hatt. 1975 Typus: Neu Guinea. Indonesia: West Irian, Tjenderawasih, Biak 1952, Brongersman c.s. G-17 hb L [HLB 952.24-098] [holotypus], NICH [isotypus] Basionym: Distribution: New Guinea var nov - syn nov Hattori Hattori Grolle & Piippo = Frullania utriculata falsa Typus: 1975 1980 1984 547 490 143 J Hatt Bot Lab 39: 286-287, fig. 143 J Hatt Bot Lab 47: 183 Acta Bot Fennica 125: 19 [Cat. West Melanesia] Steph. 1894 [auct. non STEPH 1910] Steph. & Watts 1914 Watts sn (det Steph) Basionym: Distribution: - syn nov Stephani & Watts Hattori = Frullania fulfordiae falsa 1914 1987 220 400 J Proc Roy Soc New S Wales 48: 107 [Lord Howe Isl] Bryologist 90 (4): 366 S.Hatt. 1987 Steph. 1910 Typus: Tasmania. in hb G 3616 [lectotypus] in hb G 18205 & 18210 [syntypus] Basionym: Distribution: Australia, Tasmania, Lord Howe Isl spec nov - syn nov Stephani Stephani & Watts Rodway 1910 1914 1916 Bonner Hattori 1965 1979 Hattori Miller, Whittier & Whittier Stephani 1979 1983 1985 Scott & Bradshaw Konrat & Braggins Carthy McCarthy 1986 2001 2003 2006 = Frullania probosciphora 686 220 914 Spec Hepat 4: 413-414 J Proc Roy Soc New S Wales 48: 107 [Australia] Papers Proceed Roy Soc Tasmania for 1916: 137 [syn sub Fr. pentapleura] 655 Index Hepat 5: 301-302 486 J Hatt Bot Lab 46: 131-132, fig 35(a-h) from syntype [as Fr. pentapleura ssp. falsa] 486 J Hatt Bot Lab 46: 139 [ comb nov as Fr. pentapleura ssp. falsa] 609 Prod Flora Hepat Polynesiae: 126 928 Stephani Icon Hepat Inedit [Galeiloba] C-207, microfiche 53 fig 3616 ["olim Fr reptans Mitten, Tasmania"] - 3617 56 Brunnonia 8 (1): 34 [Australia] 838 J Hatt Bot Lab 91: 245-250 [syn nov sub Fr. probosciphora] 1125 Flora Australia Suppl Ser 21: 45 [sub Fr probosciphora] 1385 Checklist Australian Liverworts [Australia, syn sub Fr probosciphora] Taylor 1846 Page 147 falsiloba von Konrat & Brennan 2004 [orth var] Basionym: Distribution: Konrat & Brennan = Frullania falciloba falsipapuana 2004 1164 http://www.liverworts.org/dich_key/frul_key.shtml, 3rd of 6 pages [Key to Frullania of New Zealand; throughout] Lehm. 1844 S.Hatt. 1982 [nom inval] Basionym: Frullania papuana var. latistipula S.Hatt. 1974 Distribution: Hattori Hattori Grolle & Piippo Hattori & Piippo = Frullania papuana falsisinuata 1982 1982 1984 1986 - syn nov S.Hatt. & Piippo 1986 S.Hatt. 1975 Hattori & Piippo 1986 557 Hattori Hentschel et alii 1988 2015 516 J Hatt Bot Lab 65: 426-427 [PNG] 4097 Phytotaxa 220 (2): 137 [syn sub Fr sinuata] = Frullania sinuata falsisinuata Misc Bryol Lichen 9 (6): 123 J Hatt Bot Lab 51: 217 [stat et nom nov] Acta Bot Fennica 125: 19 [Cat. West Melanesia] Acta Bot Fennica 133: 32 [= nom inval] Verd. 1930 Basionym: Frullania sinuata ssp. novoguinensis Distribution: New Guinea (endemic) stat et nom nov 422 495 143 557 Acta Bot Fennica 133: 28-29 Sande Lac. 1853 fo. parvistylata (S.Hatt.) S.Hatt. & Piippo 1986 Basionym: Frullania sinuata ssp. novoguinensis fo. parvistylata S.Hatt. 1975 Distribution: New Guinea comb nov Hattori & Piippo = Frullania sinuata faurieana 1986 557 Acta Bot Fennica 133: 29 (forms with small, filiform styli of stemleaves] Sande Lac. 1853 Steph. 1894 [non Steph. 1910] {Philippines} Typus: Philippinen leg. Micholitz as "fauriana", in honor of U.J. Faurie (1847-1915) and corrected in accordance with Arts. 60.8 & 60.9 (Shenzhen Code) Basionym: Distribution: syn nov - syn nov Stephani Stephani Kamimura Hattori Tan & Engel = Frullania berthoumieui 1894 1910 1961 1974 1986 228 686 199 545 4286 Hedwigia 33: 144 {Philippines} Spec Hepat 4: 402 J Hatt Bot Lab 24: 43 [identical with Fr osumiensis according Hattori] J Hatt Bot Lab 38: 188 [syn sub Fr berthoumieui] J. Hatt. Bot. Lab.. No. 60: 335 lectotype was not explicitly selected, Hattori 1974: the extra-Japanese specimens so named by Stephani are F. berthoumieui Steph. 1894 Page 148 faurieana Steph. 1910 [hom illeg] [non Steph. 1894] Typus: New Guinea, PNG: New Britain [Neu Pommern], Gazellen Halbinsel, between Paparatawa & Toma sd, Rechinger 4598 in hb G as "fauriana", in honor of U.J. Faurie (1847-1915) and corrected in accordance with Arts. 60.8 & 60.9 (Shenzhen Code) Basionym: Distribution: Bismarck Arch (New Britain [=Neupommern]) spec nov nom nov - syn nov Stephani Bonner Miller Grolle & Piippo Hattori & Streimann 1910 1965 1981 1984 1985 217 655 439 143 504 Denkschr K Akad Wiss, Math Nat Kl, Wien 85: 196-197 Index Hepat 5: 302 [New Pommerania] Phytologia 47 (4): 322 [nom nov, changed to Fr. novopommeriensis] Acta Bot Fennica 125: 19 [Cat. West Melanesia] J Hatt Bot Lab 59: 120-121 [syn sub Fr. ericoides var. planescens] [New Guinea] = Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides (Nees) Mont. 1839 faurieana var. yoshinagana S.Hatt. 1944 [nom nud] Basionym: Distribution: - syn nov Hattori Hattori Bonner Gao & Chang Aur & Zhang Bapna & Kachroo Yuzawa Yamada & Iwatsuki = Frullania truncatifolia 943 529 655 1021 1022 712 1660 1470 Bull Tokyo Sci Mus 11: 155 [nom nud] J Hatt Bot Lab 8: 31 Index Hepat 5: 302 Fl Hep Chinae bor-orient: 153-154 [sub Fr muscicola] Fl Heilongjiangensis 1: 162 [sub Fr muscicola] Hepaticology in India II: 175 [as "yoshinngana", sub Fr. muscicola] Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] J Hatt Bot Lab 99: 29 [Catalog of Japan] Steph. 1910 Frullania muscicola feana 1944 1952 1965 1981 1985 2000 2001 2006 Steph. 1894 Steph. 1910 Typus: Burma: prope Bhamo -.7.1886, S. Fea sn ex hb LEVIER 45, hb STEPHANI 10, in hb G 14410 Basionym: Distribution: Burma spec nov - syn nov Stephani Verdoorn 1911 1930 686 293 Bonner Hattori Stephani 1965 1973 1985 655 542 928 = Frullania ericoides Spec Hepat 4: 452 Ann Bryol Suppl: 42 [steht zwischen Frullania squarrosa und Frullania microauriculata] Index Hepat 5: 302-303 J Hatt Bot Lab 37: 130-131 [syn sub Fr. ericoides] Stephani Icon Hepat Inedit [Galeiloba] C-129, microfiche 52 fig 3519 [from type; "Birma, leg L Fea 1886, Levier 45"] (Nees) Mont. 1839 Page 149 fendleri Steph. 1894 Typus: Venezuela: Valencia sd, Fendler sn ex hb GOTTSCHE 8, in Hb G 19794 [holotypus] Basionym: Distribution: Venezuela spec nov - syn nov Stephani Stephani Yuzawa Stephani = Frullania laxiflora fijeana Typus: Fiji 1894 1910 1984 1985 228 686 502 928 Hedwigia 33: 138 Spec Hepat 4: 338 J Hatt Bot Lab 57: 396-398 (No 11), fig 11 (a-o) Stephani Icon Hepat Inedit [Chonanthelia] A- 30, microfiche 46 fig 3112-3113 [from type; "Valencia Venezuela Fendler"] Spruce 1884 [nom herb in hb HBG] {Fiji} sd, sc sn in hb HBG "(?) Fidji Isl. ohne Autor ohne Datum ohne Sammler (fehlt im Index Hepat. und in MILLER & al. 1983)" Basionym: Distribution: Feuerer 2008 Feuerer 2008 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm = filicina (Sw.) Raddi 1822 Typus: in hb FI 8 numbered samples in hb PI one sample Basionym: Jungermannia filicina Distribution: comb nov - syn nov Sw. 1788 Raddi Raddi Montagne Nees in G., L. & N. Bonner Raddi [Reprint] 1822 1823 1840 1845 1965 1976 Grolle Costa 1995 2009 = Bryopteris filicina 2873 825 467 2847 655 166 Crittogame Brasiliane [Seperate print]: 11-12 (No 27) Mem Mat Fisica Soc Ital Sci, Modena 19: 35-36 (No 27) Ann Sci Nat Bot, ser 2, 14: 334 Syn Hepat: 284-285 Index Hepat 5: 303 Flora Brasiliana. Memorie 1819-1828 with Reprint of "Crittogame Brasiliane 1822": 108 [No 27) 250 Bryophyt Bibl 48: 141 [Sieber from Flora Mauritius: 30 ??] 2866 J Bryol 31: 227 (Raddi s.n., about samples in hb PI & FI, types, studied by R.E. Stotler] (Sw.) Nees 1845 Page 150 filiformis Steph. 1910 Typus: Madagascar sd, Forsyth Major in hb G 13013 & Basionym: Distribution: Madagascar spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1910 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania loricata filipendra 686 655 185 928 Spec Hepat 4: 370 Index Hepat 5: 304 Bull Jard Bot Nat Belg 46 (1/2): 92 & 97 Stephani Icon Hepat Inedit [Galeiloba] C- 18, microfiche 50 fig 3369 [from type; "Madagascar, Forsyth Major (olim bidentula)"] 250 Bryophyt Bibl 48: 67 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Pearson 1891 Steph. by S.Hatt. 1979 [orth var] Basionym: Distribution: Hattori Windolf = Frullania filipendula 1979 1987 485 J Hatt Bot Lab 45: 339 1025 Austrobaileya 2(4): 391 [syn sub Fr rubella] Steph. 1910 Frullania rubella var. elongata (Steph.) S.Hatt. 1983 filipendula [auct. non STEPH 1910] Steph. & Watts 1914 {Lord Howe Isl} Typus: Australia. [NSW] Lord Howe Island. Northern hills and Saddleback Watts 41 b Basionym: Distribution: Stephani & Watts Miller, Whittier & Whittier Hattori = Frullania vittiana 1914 1983 1987 220 609 400 J Proc Roy Soc New S Wales 48: 107 [Lord Howe Island] Prod Flora Hepat Polynesiae: 126 Bryologist 90 (4): 369-370 S.Hatt. 1987 Page 151 filipendula Steph. 1910 Typus: Australia. New South Wales, E Ballina 17.6.1902, W.W. Watts 468 in hb G 18212 (ex hb LEVIER 3416), in hb NSW Basionym: Distribution: Australia (NSW) spec nov Stephani Stephani & Watts Bonner Hattori 1910 1914 1965 1979 686 220 655 485 - syn nov Hattori Stephani 1983 1985 499 928 Scott & Bradshaw Windolf Brown, Ramsay & Seur Carthy McCarthy 1986 1987 1992 2003 2006 56 1025 1140 1125 1385 Spec Hepat 4: 419-420 J Proc Roy Soc New S Wales 48: 107 Index Hepat 5: 304 J Hatt Bot Lab 45: 339-340, fig 12(a-g) from type & 354 [syn sub Fr. rubella] J Hatt Bot Lab 54: 155-156 [syn sub Fr. rubella var. elongata] Stephani Icon Hepat Inedit [Galeiloba] C-209, microfiche 53 fig 3619 ["Australia, N.S.Wales, East Ballina (Watts) 468, Levier 3416"] - 3620 Brunnonia 8 (1): 34 [Australia] Austrobaileya 2(4): 391 [as "filipendra" syn sub Fr rubella] Telopea 5(1): 75 [types in hb NSW & hb SYD] Flora Australia Suppl Ser 21: 46 [sub Fr rubella var rubella] Checklist Australian Liverworts [Australia, syn sub Fr "pubella "] = Frullania rubella var. elongata (Steph.) S.Hatt. 1983 fissistipula Steph. 1894 Typus: Australia: Bellender Ker Range 1887, Sayer W.A. Sn in hb G 18486 - G00069809 - n° SIB 136150/2 Basionym: Distribution: Australia (QLD) spec nov comb nov syn nov - comb nov Stephani Stephani Bonner Hattori Grolle Stephani 1894 1911 1965 1979 1984 1985 Scott & Bradshaw Grolle 1986 1987 Glenny, Meagher & Brown 2004 = Lepidolaena fissistipula flammea 228 686 655 486 2274 928 Hedwigia 33: 162 Spec Hepat 4: 662 Index Hepat 5: 304 J Hatt Bot Lab 46: 132 [is a species of genus Lepidolaena] J Hatt Bot Lab 55: 504 [syn nov sub Lepidolaena reticulata] Stephani Icon Hepat Inedit [Diastaloba] B-93, microfiche 49 fig 3341 [Australia orientalis, Bellender Ker Mts, Sayer 1887 leg"] 56 Brunnonia 8 (1): 34 & 143 [Australia: QLD] 1156 Hausknechtia 3: 65-69, fig 1-2 [comb nov as Lepidolaena fissistipula (Steph) Grolle 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html (Steph.) Grolle 1987 Taylor 1884 Typus: Ecuador. Andes Quitenses, prope Cuenca sd, Jameson in hb MANCH Basionym: Distribution: Ecuador spec nov - syn nov Taylor in Spruce Stephani Bonner Gradstein = Frullania decidua 1884 1911 1965 2021 243 686 655 5642 Trans Proc Bot Soc Edinburgh 15: 14 & 29-30 Spec Hepat 4: 670 & 672 Index Hepat 5: 304 Mem NY Bot Gard 121: 364 [syn sub Frullania decidua] Spruce 1884 Page 152 fleischeri Steph. 1911 Typus: Indonesia. Java, Mura Enim, in Mte Merapi sd, M. Fleischer sn hb LEVIER 3984, in hb G 17192 Basionym: Distribution: Indonesia (Java), New Caledonia spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Hattori Miller, Whittier & Whittier Stephani 1911 1929 1930 1965 1976 1983 1985 686 245 293 655 67 609 928 Bapna & Kachroo 2000 712 Spec Hepat 4: 463-464 Ann Bryol 2: 134 [syn sub Fr. squarrosa var planescens] Ann Bryol Suppl 1: 38 Index Hepat 5: 304 [Java, New Caledonia] Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21 Prod Flora Hepat Polynesiae: 126 Stephani Icon Hepat Inedit [Galeiloba] C-130, microfiche 52 fig 35203521 ["Java, in monte Merapi, Fleischer legit"] Hepaticology in India II: 170 [sub Fr. ericoides] = Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides fleischeri (Nees) Mont. 1839 Steph. ex Paris 1910 [nom nud] Basionym: Distribution: Paris = Frullania fleischeri 1910 1455 Rev Bryol 37 (6): 130 [New Caledonia: Dogny] Steph. 1911 Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides (Nees) Mont. 1839 Page 153 flexicaulis Spruce 1884 Typus: Ecuador. Andes Quitenses, Mt. Mulmul, 3000m - Ecuador. Tungurahua, Cerro Mulmul - "in Mte. Mulmul, in fruticosum cortice vetusto" sd, Jameson in hb MANCH in hb G G00066974 - n° SIB 125556/1 - G13582 ("Spruce's Original" ex hb SPRUCE sub Fr. flexicaulis SPR.) > see fr0331.pdf Basionym: Distribution: Mexico, Jamaica, Colombia, Ecuador, Bolivia, Peru, Galapagos Isls ? [Ecuador], Brazil spec nov -syn nov Spruce Stephani Weberbauer Stephani Weberbauer Soukup Bonner Stotler Stotler Gradstein & Hekking Cole Menzel Stephani 1884 1910 1911 1916 1945 1955 1965 1969 1970 1979 1983 1984 1985 243 686 792 222 793 171 655 4761 194 483 1407 142 928 Fulford & Sharp Buck Uribe & Gradstein Yano Gradstein & Costa 1990 1994 1998 1999 2003 105 28 707 1263 984 Yano Gradstein 2008 2009 1684 2161 Söderström, Hagborg, Vana & Konrat Hentschel et alii 2011 2581 2015 4097 Phytotaxa 220 (2): 137 [syn sub Fr paradoxa] = Frullania paradoxa floribunda Trans Proc Bot Soc Edinburgh 15: 45 & 53-54 Spec Hepat 4: 387-388 [as Trachycolea] Pflanzenwelt Peruan Anden: 238 [Peru] Biblioth Bot 87: 242 [Bolivia] Mundo Vegetal Andes Peruanos: 568 [Peru] Biota 1(4): 81 [Peru] Index Hepat 5: 305 Nova Hedwigia 18: 463-464, pl 27(235-243) The genus Frullania, separata: 463-464, pl 27(235-243) J Hatt Bot Lab 45: 112 [Cat Colombia] Brenesia 21: 154 [Central America] Willdenowia 14: 489 [Peru] Stephani Icon Hepat Inedit [Galeiloba] C- 60, microfiche 50 fig 3416 ["Bolivia, Undari ..."] Mem NY Bot Gard 63: 42 [Mexico] Bryologist 97 (4): 422 [Vainio's specimen from Brazil at TUR] Bryophyt Bibl 53: 36 [Catalogue Colombia] Briophyta do estado de São Paulo. Lista provisória [Brazil] Mem New York Bot Gard 87: 86-87 [Brazil] [may be a synonym of Fr deppii ="autoicous form"; Fr flexicaulus ="heteroicous form" ] Boletim Instituto Botanica 19: 31 [Brazil; catalog] Bibliotheca Lichenologica 99: 158 [Galapagos Isl ?, Galapagos islands, without locality, Taylor 1013 (STOTLER 1969).] Bryologist 114(1) : 72 [checklist Jamaica] Lehm. & Lindenb. 1845 Steph. 1894 Typus: Society Isl. Tahiti 1855, Vieillard & Panchet ex hb BESCHERELLE, in hb G Basionym: Distribution: spec nov Stephani Bescherelle Paris Stephani Verdoorn Verdoorn Bonner Miller, Whittier & Whittier Stephani 1894 1898 1906 1911 1930 1930 1965 1983 1985 228 871 676 686 293 551 655 609 928 - syn nov Hattori 1986 506 Hedwigia 33: 144-145 J Bot Morot 12: 138 [spec nov] [Tahiti] Rev Bryol 33 (2): 28 [New Caledonia] Spec Hepat 4: 455-456 Ann Bryol Suppl 1: 49 [syn sub Fr. reflexistipula] Nederl Kruidk Arch Jrg 1930 (2): 160 Index Hepat 5: 305 Prod Flora Hepat Polynesiae: 126 Stephani Icon Hepat Inedit [Galeiloba] C-131, microfiche 52 fig 3522 ["Taiti, Vieillard & Panchet 1855"] - 3523 ["ex planta tahitense a Nadeaud lecte No 197"] J Hatt Bot Lab 60: 218 [syn sub Fr. reflexistipula var. novocaledonica] = Frullania reflexistipula var. novocaledonica S.Hatt. 1977 Page 154 floribundaria Steph. by Bescherelle ex Miller, Whittier & Whittier 1963 [orth var] Typus: Society Isl Basionym: Distribution: Society Isl Stephani in Bescherelle 1898 871 Verdoorn Miller, Whittier & Whittier 1930 1983 293 609 = Frullania floribunda J Bot Morot 12: 138 ["floribundaria" not existing, orth error by Miller et al 1893] Ann Bryol Suppl 1: 49 [not mentioned, error by Miller et al 1983] Prod Flora Hepat Polynesiae: 126 Steph. 1894 Frullania reflexistipula var. novocaledonica S.Hatt. 1977 fluminensis Gottsche ex Steph. 1894 Typus: Brazil. Rio Janeiro sd, Gaudichaud ex hb GOTTSCHE, ex hb BESCHERELLE, hb STEPHANI 214, in hb G Basionym: Distribution: Brazil spec nov Gottsche in Stephani Stephani Stephani Schiffner & Arnell Bonner Stephani = Frullania beyrichiana fnouena 1894 1897 1911 1964 1965 1985 228 1059 686 198 655 928 Hedwigia 33: 157-158 Bihang Kgl Sv Vetensk-Akad Handl bot 23 III (2): 19 [Brazil] Spec Hepat 4: 495 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 88 [Brazil] Index Hepat 5: 305 Stephani Icon Hepat Inedit [Thyopsiella] F- 56, microfiche 57 fig 3953 ["Rio Janeiro, Gaudichaud leg, Herb Gottsche"] - 3954 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. by Choe 1980 [orth var] Basionym: Distribution: Choe 1980 1171 Illust Fl & Fauna Korea 24: 734 [South Korea] = Frullania muscicola var. inuena (Steph.) Kamim. 1961 Frullania muscicola formosa Steph. 1894 Steph. ex Iishiba 1912 Typus: Japan Basionym: Distribution: Iishiba = Frullania formosae 1912 888 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Steph. 1924 {Taiwan specimen, sensu S.Hatt. & LIN 1985} Frullania obtusangula Hentschel & von Konrat 2015 Page 155 formosa Steph. 1911 [nom herb] [non Spruce 1884] Typus: Brazil Basionym: Distribution: nom herb Stephani Bonner 1911 1965 Yano 1995 = Frullania confusa Spec Hepat 4: 517 [not mentioned here], but at page 522 Index Hepat 5: 305-306 [Brazil; error - not as syn sub Fr. confusa in STEPH 1911: 517] 2095 J Hatt Bot Lab 78: 142 [annotated checklist of Brazil] Steph. 1911 Frullania montagnei formosae 686 655 Gottsche 1845 Steph. ex Sawada 1909 [nom nud] Basionym: Distribution: Sawada 1909 = Frullania formosae Bot Mag Tokyo 23 (no 268): 178-183, fig 11 [key to Japanese species, in Japanese] Steph. 1924 {Taiwan specimen, sensu S.Hatt. & LIN 1985} Frullania obtusangula formosae 930 Hentschel & von Konrat 2015 Steph. 1924 {Hawaii specimen} Typus: Oaho. Honolulu sd, Fleischer 106 in hb G - n° SIB 90540/2 [lectotypus] Basionym: Distribution: spec nov syn nov syn nov syn nov - syn nov Stephani Verdoorn Miller Miller Lai & Wang-Yang Hattori & Lin 1924 1930 1953 1963 1976 1985 687 551 349 834 33 505 Bapna & Kachroo 2000 712 = Frullania oahuensis Spec Hepat 6: 539 Nederl Kruidk Arch Jrg 1930 (2): Bryologist 56: 45 [syn nov sub Fr squarrosa] Ark Bot, ser 2, 5: 518 [sub Fr. squarrosa] Taiwania 21 (2): 169 [sub Fr. tamsuina] J Hatt Bot Lab 59: 134 [Hawaiian specimen = syn nov sub Fr. oahuensis] Hepaticology in India II: 170 [sub Fr. ericoides] Hampe ex Gottsche, Lindenb. & Nees 1843 Page 156 formosae Steph. 1924 {Taiwan specimen, sensu S.Hatt. & LIN 1985} Typus: Formosa, Tamsui 22.5.1903, U. Faurie 44 in hb G [lectotypification by HATTORI & LIN 1985]; in hb KYO, TNS [isotypus] Basionym: Distribution: Taiwan spec nov syn nov -syn nov Stephani Verdoorn 1924 1930 Horikawa 1934 Hattori Hattori & Noguchi Kamimura Bonner Hattori Lai & Wang-Yang Hattori & Lin 1952 1960 1961 1965 1974 1976 1985 Stephani 1985 Inoue Kuo & Chiang Piippo Bapna & Kachroo N.N. [National Science Museum Tokyo] Wang, Lai & Zhu Hentschel et alii 1987 1988 1990 2000 2000 = Frullania obtusangula formosana 2011 2015 687 293 Spec Hepat 6: 539 [Formosa & Honolulu] Ann Bryol Suppl 1: 37 [= syn sub Fr. sandvicensis = syn of Fr. neurota] 34 J Sci Hiroshima Univ, ser B, div 2, 2: 103 [syn = Fr. squarrosa , as pointed out by Verdoorn] 529 J Hatt Bot Lab 8: 31 [= syn sub Fr. squarrosa] 965 Index spec Typ herb Japon, Bryophyta: 16 [isotypus] 199 J Hatt Bot Lab 24: 19 ["isotype" = Hawaiian specimen = Fr. tamsuina] 655 Index Hepat 5: 306 545 J Hatt Bot Lab 38: 185 [sub Fr. tamsuina] 33 Taiwania 21 (2): 169 [Index to Taiwan, = syn sub Fr. tamsuina] 505 J Hatt Bot Lab 59: 134 [Taiwan, lectotypification, Hawaiian specimen = syn sub Fr. oahuensis] 928 Stephani Icon Hepat Inedit [Galeiloba] C-132, microfiche 52 fig 3524 ["hierzu fehlt Exemplar aus Formosa, Faurie 204x (1903)" - "Exemlar aus Honolulu, Fleischer 108 (1913)" - "Insula Formosa, leg (Faurie)"] 1155 Index type specimens in Nat Sci Mus, Tokyo: 12 948 Taiwania 33: 12 [Index of Taiwan] 521 J Hatt Bot Lab 68: 40 [annotated catalogue of China] 712 Hepaticology in India II: 170 [syn sub Fr. ericoides] 995 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 3125 Annales Botanici Fennici 48: <369-395> [Checklist Taiwan] 4097 Phytotaxa 220 (2): 137 [nom. nov. pro Frullania formosae Steph., Sp. Hepat. (Stephani) 6: 539, 1924 (Stephani 1924), nom. illeg. (ICN Art. 53.1; hom. illeg.). Hentschel & von Konrat 2015 Steph. by Kamim. 1961 [orth var] Basionym: Distribution: orth var Kamimura Hattori = Frullania formosae 1961 1974 199 545 J Hatt Bot Lab 24: 19 J Hatt Bot Lab 38: 185 Steph. 1924 {Taiwan specimen, sensu S.Hatt. & LIN 1985} Frullania obtusangula Hentschel & von Konrat 2015 Page 157 forsythiana Steph. 1910 Typus: Australia. Shoalhaven River Distr., Barker`s Creek -.9.1899, W. Forsyth 10 ex hb NSW, in hb G 19213 Basionym: Distribution: Australia spec nov - syn nov Stephani Stephani & Watts Bonner Hattori Stephani 1910 1914 1965 1979 1985 686 220 655 485 928 Scott & Bradshaw Carthy Glenny, Meagher & Brown 1986 2003 2004 56 1125 1126 Glenny, Meagher & Brown 2004 1127 McCarthy 2006 1385 = Frullania falciloba fortunati Spec Hepat 4: 419 J Proc Roy Soc New S Wales 48: 107 [Australia] Index Hepat 5: 306 J Hatt Bot Lab 45: 340-342, fig 13(a-k) from type Stephani Icon Hepat Inedit [Galeiloba] C-210, microfiche 53 fig 3621 ["Australia orient., Shoalhaven River, W. Forsyth, Sept 1899"] Brunnonia 8 (1): 34 [Australia] Flora Australia Suppl Ser 21: 43 [sub Fr falciloba] NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Checklist Australian Liverworts [Australia, syn sub Fr falciloba] Lehm. 1844 Steph. ex Verd. 1930 [pro syn] Typus: China. Prov. Kony Tcheou, Pin-fa 30.1.1904, Fortunat hb STEPHANI 9, in hb G [sub Fr. sanguinea] Basionym: Distribution: China - syn nov Theriot Verdoorn Bonner Stephani 1909 1930 1965 1985 3357 293 655 928 Piippo 1990 521 = Frullania nepalensis fragilifolia [erwähnt] Ann Bryol Suppl 1: 51 [syn sub Fr. nepalensis] Index Hepat 5: 306 Stephani Icon Hepat Inedit [Thyopsiella] F-168, microfiche 59 fig 4109 ["China, Prov. Köng Tschen, Fortunat 30/1. 1904, Herb Thériot"] J Hatt Bot Lab 68: 40 [annotated catalogue of China] (Spreng.) Lehm. & Lindenb. 1845 [auct. non (Taylor) Taylor] ADE & KOPPE 1942 {Canary Isl} Typus: Canary Island. Teneriffe. "Mercedeswald" in hb KOPPE (Bielefeld) Basionym: Distribution: auct - syn nov Ade & Koppe 1942 Grolle 1970 Dirkse, Bouman & Losada-Lima 1993 206 615 11 Hedwigia 81 (1/2): 34 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 310 Crypt Bryol Lichen 14 (1): 4-5 = Frullania microphylla var. deciduifolia Grolle 1970 Page 158 fragilifolia [auct. non (Taylor) Taylor 1845] Yoshinaga 1898 {Japan} Typus: Japan. Tosa -.4.1899, Yoshinaga in hb BEROL Basionym: Distribution: - syn nov Yoshinaga Reimers Hattori Kamimura Kamimura Choe Hattori & Lin Yamada & Iwatsuki = Frullania densiloba fragilifolia 1898 1931 1952 1952 1961 1980 1985 2006 887 229 529 873 199 1171 505 1470 Bot Mag Tokyo 12: 211 Hedwigia 71 (1/2): 38 [syn sub Fr. densiloba] J Hatt Bot Lab 8: 29 [Fr. fragilifolia not in this region] Contrib Fl Hepat Shikokuensem, Kochi: 134 J Hatt Bot Lab 24: 69 [syn sub Fr densiloba] Illust Fl & Fauna Korea 24: 733 [South Korea] J Hatt Bot Lab 59: 158 [sub Fr. densiloba] J Hatt Bot Lab 99: 29 [Catalog of Japan] Steph. ex A.Evans 1906 [auct. non (Taylor) Taylor 1845] Austin 1879 {USA} Basionym: Distribution: spec nov - syn nov Austin Evans Frye & Clark = Frullania selwyniana 1879 1897 1947 667 237 204 Bull Torrey Bot Club 6 [No 52]: 301 Trans Connet Acad Arts Sci 10 (1): 29-30 [syn sub Fr selwyniana] Univ Wash Publ Biol 6 (5): 755 [sub Fr selwyniana] Pearson 1890 fragilifolia var. autoica C.Massal. ex Levier 1906 [nom nud] {China} Typus: China. Prov Schensi, Tui-kio-san -.8.1896, Giraldi Basionym: Distribution: var nov - syn nov Massalongo ex Levier Müller in Rabenhorst 1906 1915 Reimers Hsu Wu et alii Hattori & Lin Hattori & Lin Wu & Zheng Piippo 1931 1979 1984 1985 1985 1988 1990 = Frullania punctata 824 Nuovo Giorn Bot Ital NS 13: 348 (Hepat Schensiana) 1064 Krypt Flora (2ed) 6, 2: 622 [specimen from China belongs to another species] 229 Hedwigia 71 (1/2): 38 [belongs to another species] Index of bryol list China (134 p.) A glossary of terms and names of bryophytes. Peking [124 p.] : 348 505 J Hatt Bot Lab 59: 159 [may be Fr punctata] 521 J Hatt Bot Lab 68: 40 [annotated catalogue of China] Reimers 1931 Page 159 fragilifolia var. microphylla Schiffn. ex Grolle 1970 [nom herb] Typus: Italy. Südtirol: Schlern Südseite, 1440-1500m 23.7.1899, Schiffner [sterile Probe] in hb FH in hb JE in hb S-PA Basionym: Distribution: Italy (Südtirol) - syn nov Grolle Hattori 1970 1978 615 482 Bisang & Schumacker Bisang, Schumacker, Sérgio & Grolle 1987 1989 612 276 = Frullania dilatata Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 316 J Hatt Bot Lab 44: 545 [caucasica + parvistipula + SCHIFFNER's specimen = probably 1 species] Bot Helvetica 97: 311 Giorn Bot Ital 122: 265 fo. decidua [?] Grolle 1970 Frullania parvistipula Steph. 1910 fragilifolia var. parvistipula Schiffn. in Bisang, Schumacker, Sergio & Grolle 1988 Basionym: Distribution: Bisang, Schumacker, Sérgio & Grolle = Frullania parvistipula fragilis 1989 276 Giorn Bot Ital 122: 265 [syn sub Fr. parvistipula] Steph. 1910 Steph. ex Bonner 1965 [nom herb] Typus: Japan, Jyo, Mt. Onigajo 28.12.1909, S. Okamura 360 in hb G 14412 Basionym: Distribution: Japan nom herb - syn nov Bonner Hattori Stephani 1965 1974 1985 Yuzawa 2001 = Frullania muscicola 655 545 928 Index Hepat 5: 306-307 J Hatt Bot Lab 38: 207-208 [syn sub Fr. muscicola] Stephani Icon Hepat Inedit [Galeiloba] C-234, microfiche 53 fig 3654 ["Japonia, O Kamura No 360"] 1660 Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] Steph. 1894 fugax var. caduca S.Hatt. 1982 [nom herb] Typus: New Zealand. North Island, Wellington, Wairarapa, Carters Bush - on bark of tree 4.5.1968, Braggins in hb AKU 062493 in hb WELT H008831 Basionym: Distribution: Hattori 1982 0 note in hb WELT / AKU http://collections.tepapa.govt.nz/objectdetails.aspx?oid=405848&term= frullania = Page 160 fugax var. media E.A.Hodgs. 1949 Typus: New Zealand sd, K.W. Allison H 510 Basionym: Distribution: New Zealand var noc - syn nov Hodgson Bonner Hamlin Hattori Glenny Glenny & Malcolm = Frullania media fukushimensis 1949 1965 1972 1983 1998 2005 203 655 659 499 1412 1410 Trans Roy Soc New Zealand 77 (3): 375 Index Hepat 5: 307 Rec Dominion Mus 7(19): 262 [New Zealand] J Hatt Bot Lab 54: 153-156 Tuhinga 10: 132 [sub Fr media] ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr media]] (E.A.Hodgs.) S.Hatt. 1983 Kamim. 1982 Typus: Japan. Fuku-shima-ken, 100m 5.5.1962, Y. Yuzawa 1094 in hb YUZAWA & NICH [isotypus] Basionym: Distribution: spec nov - syn nov Kamimura Hattori 1982 1986 Yuzawa 2001 Yamada & Iwatsuki 2006 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania hamatiloba fukuzawana 420 74 Misc Bryol Lichen 9 (4): 86-88, fig 1 Bull Natn Sci Mus Tokyo, ser B (Bot), 12 (4): 138 [syn sub Fr. hamatiloba] 1660 Natural Environmental Sci Res 14: 16 [Japan, syn sub Fr hamatiloba] 1470 J Hatt Bot Lab 99: 29 [Catalog of Japan] 2112 Type sepecimens Hatt Bot Lab (NICH) 24: Isotype. Japan, Honshu, Fukushima.,.ken, Soma-gun, Odaka-machi, Kanabusa. 100 m. On bark of tree. 5 May 1962 . Y. Yuzawa 1094 (NICH 237411) Steph. 1910 Steph. ex Bonner 1965 [nom herb] Typus: Japan. Fukuzawana, Awa 2.2.1913, S. Okamura 422 in hb G 14413 Basionym: Distribution: Japan nom herb - syn nov Bonner Hattori Stephani 1965 1974 1985 = Frullania muscicola funchalensis Typus: Funchal 655 545 928 Index Hepat 5: 307-308 J Hatt Bot Lab 38: 207-208 [syn sub Fr. muscicola] Stephani Icon Hepat Inedit [Galeiloba] C-235, microfiche 53 fig 3655 [Japonia, O Kamura leg 422"] Steph. 1894 Gottsche ex Bonner 1965 [nom herb] sd, Fritze in hb G 17558 Basionym: Distribution: nom herb Stephani Bonner 1910 1965 686 655 - syn nov Vanden Berghen 1976 185 = Frullania dilatata Spec Hepat 4: 434 [not mentioned] Index Hepat 5: 308 [error: not as syn sub Fr. cesatina in STEPH 1910: 434] Bull Jard Bot Nat Belg 46 (1/2): 191 [syn sub Fr. dilatata, nom nud] (L.) Dumort. 1835 Page 161 fusco-brunnea Steph. 1924 Basionym: Frullania fallax Distribution: Bolivia Steph. 1911 [hom illeg] [non Gottsche in G., L. & N. 1845] {Bolivia, Guatemala} nom nov Stephani 1924 - syn nov - syn nov Bonner Stotler Stotler Stephani 1965 1969 1970 1985 Gradstein, Meneses & Arbe 2003 = Frullania intumescens fuscopurpurea 909 Spec Hepat 6: 570 [Ad Vol. Ivum: Frullania fusco-brunnea St. mss. in herb. = F. fallax St. in Spec. Hep. IV [1910],] 655 Index Hepat 5: 308 4761 Nova Hedwigia 18: 444 [syn sub Fr. intumescens] 194 The genus Frullania, separata: 444 [syn sub Fr. intumescens] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 57, microfiche 57 fig 3955 ["Bolivia, Herzog 1907, Levier 5848, 5854"] 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. intumescens] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1911 Typus: Indonesia. Java sd, s coll hb SANDE LACOSTE 8911, in hb G 17193 Basionym: Distribution: Indonesia (Java) spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Stephani = Frullania minor 1911 1929 1930 1965 1985 Spec Hepat 4: 643 Ann Bryol 2: 162 Ann Bryol Suppl 1: 117 [syn sub Fr. minor] Index Hepat 5: 308 Stephani Icon Hepat Inedit [Diastaloba] B-62, microfiche 49 fig 3302 [from type; "Java, Herb Sande-Lacoste"] (Sande Lac.) Verd. 1929 Frullania gracilis ssp. minor Frullania gracilis fusiflora 686 246 293 655 928 (Sande Lac.) S.Hatt. 1975 (Reinw., Blume & Nees) Nees 1845 Steph. 1924 [nom superfl] Basionym: Distribution: spec nov Stephani = Frullania fusiflora 1924 687 Spec Hepat 6: 531-532 [as spec nov; Mexico & Bolivia] Steph. ex Herzog 1916 Frullania obscura (Sw.) Mont. 1840 Page 162 fusiflora Steph. ex Herzog 1916 Typus: Bolivia: Florida de San Mateo, 2000 m 1913, Herzog 3658 in hb G 20962 [holotypus]; Basionym: Distribution: Mexico, Costa Rica, Bolivia spec nov - syn nov Stephani Stephani Bonner Yuzawa Stephani 1916 1924 1965 1983 1985 222 687 655 498 928 Fulford & Sharp Morales Z. Yuzawa Gradstein, Meneses & Arbe 1990 1991 1991 2003 105 307 525 975 = Frullania obscura fusiloba Biblioth Bot 87: 239, fig 187 [Bolivia] Spec Hepat 6: 531-532 [as spec nov; Mexico & Bolivia] Index Hepat 5: 308 J Hatt Bot Lab 54: 109-113, fig 3 (a-p) [from type] [syn sub Fr. arecae] Stephani Icon Hepat Inedit [Chonanthelia] A- 32, microfiche 46 fig 3115-3116 [from type; "Bolivia Dr Herzog No 3658 (1913)"] Mem NY Bot Gard 63: 42 [Mexico] Trop Bryol 4: 37 [Costa Rica] J Hatt Bot Lab 70: 264 [sub Fr. areacae] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. arecae; according Yuzawa 1991 ?] (Sw.) Mont. 1840 Steph. 1924 Typus: Mexico: sd, Arsén 7850 in hb G [holotypus]; Basionym: Distribution: Mexico spec nov - syn nov Stephani Bonner Stephani 1924 1965 1985 687 655 928 Fulford & Sharp Yuzawa 1990 1991 105 525 = Frullania obscura galapagona Spec Hepat 6: 532 Index Hepat 5: 309 Stephani Icon Hepat Inedit [Chonanthelia] A- 31, microfiche 46 fig 3114 [from type; "Mexico Arsèn 7850"] Mem NY Bot Gard 63: 42 [Mexico] J Hatt Bot Lab 70: 264 (Sw.) Mont. 1840 Ångstr. 1873 [non Steph. 1911] Typus: Galapagos Isl. Charles Isl. [Floreana] sd, N.J. Anderson in hb G [isotypus] Basionym: Distribution: Galapagos Isls [Ecuador] spec nov - syn nov - Ångström 1873 NN Evans Stephani Bonner Gradstein & Weber Stephani 1875 1902 1911 1965 1982 1985 Yuzawa Winter & Schäfer-Verwimp 1988 2020 = Frullania obcordata 578 Öfversigt K Vetensk Akad Förh 30 (5): 116-117 ["Charles-ön (Galapagos-öarne)"] 1495 Hedwigia 14: 85-86 [Galapagos-Insel] 665 Proc American Acad Sci 38: 100 [Galapagos Isls: Floreana] 686 Spec Hepat 4: 636-637 655 Index Hepat 5: 309 496 J Hatt Bot Lab 52: 148 [Galapagos Isls] ( syn sub Fr. obcordata] 928 Stephani Icon Hepat Inedit [Diastaloba] B-31, microfiche 48 fig 3264 [from type; "Galapagos Insae Anderson Original"] 511 J Hatt Bot Lab 64: 442-444, fig 4(from type) [syn sub Fr. caulisequa] 5742 Frahmia 19: 13 [syn sub Fr obcordata] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 163 galapagona Steph. 1911 [hom illeg] [non AONGSTRÖM 1873] Typus: Galagagos. Cocos Island 1902, Pittier in hb G Basionym: Distribution: Galapagos Isls [Ecuador] spec nov - nom nov Stephani Beauverd in Stephani Bonner Stephani = Frullania ecuadorensis Frullania kunzei galeata 1911 1924 1965 1985 686 909 655 928 Spec Hepat 4: 532-533 Spec Hepat 6: 570 [syn sub Fr. decipiens] Index Hepat 5: 309 Stephani Icon Hepat Inedit [Thyopsiella] F-108, microfiche 58 fig 4032 ["Galapagos, Cocos Island, Pittier 1902, Am. tropica"] Steph. 1911 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 (Reinw., Blume & Nees) Verd. 1929 Typus: (Reinw., Blume & Nees) Nees 1845 Basionym: Distribution: China (Yunnan), Taiwan, Sri Lanka, Indonesia (Sumatra, Java, Sulawesi), Philippines comb nov - syn nov Dumortier Lindenberg in G., L. & N. 1835 1845 Sande Lacoste 1857 Stephani 1910 Verdoorn Verdoorn 1929 1929 Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Svihla Bonner Hattori Hattori Hattori Hattori Hattori Hattori, Thaithong & Kitagawa Hattori & Thaithong Yamada, Yuzawa, Ogasawara & Lai Hattori Miller, Whittier & Whittier Grolle & Piippo Kuo & Chiang Piippo Bai Bai Gradstein et alii Kornochalert 1930 1930 1932 1932 1932 1934 1937 1956 1965 1972 1974 1974 1974 1975 1977 1978 1981 256 Rec d´Obs Jungerm., Tournay, 1: 13 2850 Syn Hepat: 413 [as Jungermannia galeata syn sub Fr. sebastianopolitana var galeata] 430 Verh Konikl Akad Wetenschappen 5: 77 [as Fr. sebastianopolitana var galeata] 686 Spec Hepat 4: 349 [syn sub Fr. commutata; according BONNER 1965, not found in STEPHANI] 245 Ann Bryol 2: 119-120 [Java, Sumatra] 246 Ann Bryol 2: 156 [Fr. sebastianopolitana var galeata = Fr. galeata (R BL NEES) DUM] 551 Nederl Kruidk Arch Jrg 1930 (2): 293 Ann Bryol Suppl 1: 68-70, fig. 105-109(:49) 232 Ann Bryol 5: 126 [Hepat select crit, ser 3 & 4 no 166; Java] 554 Nederl Kruidk Arch Jrg 1932 (2): 492 [Sumatra, Java] 553 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 57 [Java] 241 Ann Bryol Suppl 4: 204 [Sumatra] 556 Blumea suppl 1: 212 [Rapa, Society Islands] 380 Bryologist 59: 278-279 [syn sub Fr. riojaneirensis] 655 Index Hepat 5: 310 538 J Hatt Bot Lab 36: 135, fig. 20-21 544 J Hatt Bot Lab 38: 177 [Philippines] 544 J Hatt Bot Lab 38: 177 [Luzon] 546 J Hatt Bot Lab 38: 240 547 J Hatt Bot Lab 39: 287 480 J Hatt Bot Lab 43: 447 [Thailand] 70 Bull Natn Sci Mus Tokyo, ser B (Bot), 4 (2): 66 [Burma] 949 Quart J Taiwan Mus 34 (1,2): 113 [new to Taiwan] 1982 1983 1984 1988 1990 2001 2001 2005 2006 495 609 143 948 521 1742 1742 1381 2327 Aryanti & Gradstein 2007 = Frullania rio-janeirensis J Hatt Bot Lab 51: 261-262 Prod Flora Hepat Polynesiae: 126 Acta Bot Fennica 125: 19 [Cat. West Melanesia] Taiwania 33: 12 [Index of Taiwan] J Hatt Bot Lab 68: 41 [annotated catalogue of China] Guizhou Science 19(4): 47-49 [China, Yunnan] Guizhou Science 19(4): 47-49 [China, Yunnan] J Hatt Bot Lab 98: 239 [Sulawesi, Indonesia] Thesis: Thailand, Doi Suthep-Pui National Park, Chiang Mai Province: 111-112, fig 32(1-9) & 65 (foto) 1476 Cryptogamie Bryologie 28(1): 7 [Sulawesi, Asiatic, montane] (Raddi) Ångstr. 1876 Page 164 garsidei S.W.Arnell 1953 Typus: S Africa: Cape Province, Knysna, Deepwalls Forest sd, S. Arnell 1835 /? 1494[Vanden Berghen 1976] in hb BOLUS, Cape Town, in hb S-PA [holotypus] Basionym: Distribution: South Africa spec nov Arnell Arnell Bonner Vanden Berghen Wigginton 1953 1963 1965 1976 2002 Wigginton Wigginton 2004 2004 = Frullania variegata 445 822 655 185 1012 Svensk Bot Tidskr 47 (1): 116-118, fig. 6(a-o) Hepat S Africa: 158-159, fig 116(a-o) Index Hepat 5: 310 Bull Jard Bot Nat Belg 46 (1/2): 128 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 235, fig 156(C,D,F-H,L,P) from type [syn sub Fr variegata, West Africa] Steph. 1894 gaudichaudii var. ceylanica (Nees) S.Hatt. 1980 Basionym: Frullania ceylanica Nees 1845 Distribution: India (Tamil Nadu), Sri Lanka comb nov -syn nov Hattori Hattori & Lin Nath & Asthana 1980 1985 1998 Yuzawa Bapna & Kachroo Daniels & Daniel 2000 2000 2008 Long & Rubasinghe 2014 = Frullania ceylanica ssp nov (Steph.) S.Hatt. 1975 [nom inval] Steph. 1896 Hattori Hattori Miller, Whittier & Whittier Grolle & Piippo = Frullania immersa J Hatt Bot Lab 47: 103-104 [comb nov] J Hatt Bot Lab 59: 154 [syn sub Fr. gaudichaudi] J Hatt Bot Lab 85: 72- 75, fig 6(1-12) as Fr "guadichaudii, gaudichanudii" 1659 Natural Environmental Sci Res 13: 14 [Japan, syn sub Fr gaudichaudii] 712 Hepaticology in India II: 189 1987 Indian J Forestry 31(4): 637-639, fig (a-i) [India (Tamil Nadu)] [is a good species] 3994 Ceylon Journal of Science 43(1): 4 [Sri Lanka] Nees 1845 gaudichaudii ssp. immersa Basionym: Frullania immersa Distribution: Samoa 489 505 708 1975 1981 1983 1984 66 492 609 143 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 142-144, fig 159 J Hatt Bot Lab 49: 373-375 [immersa is a distinct species] Prod Flora Hepat Polynesiae: 127 Acta Bot Fennica 125: 19 [Cat. West Melanesia] Steph. 1896 Page 165 geheebii Steph. 1911 Typus: Papua New Guinea. Milne Bay, Mt. Dayman 1894, Armit jun. in hb GEHEEB, in hb G 14235 [holotypus] Basionym: Distribution: New Guinea, Solomon Isl spec nov - syn nov Stephani Verdoorn Bonner Hattori Miller, Whittier & Whittier Grolle & Piippo Stephani 1911 1930 1965 1972 1983 1984 1985 686 293 655 538 609 143 928 Hattori & Piippo 1986 557 = Frullania gaudichaudii gemmipara Spec Hepat 4: 553-554 Ann Bryol Suppl 1: 96, fig 163 Index Hepat 5: 310 J Hatt Bot Lab 36: 126-127 Prod Flora Hepat Polynesiae: 127 Acta Bot Fennica 125: 19 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Thyopsiella] F-169, microfiche 59 fig 4110 ["N Guinea, Mt Dayman, 1894, leg Armit junior"] Acta Bot Fennica 133: 4 (Nees & Mont.) Nees & Mont. 1845 R.M.Schust. & S.Hatt. 1978 Typus: China. Prov. Yunnan, prope Yunanfu, ad templum Schili-ngan, about 2200m 27.11.1914, Dr. H.F. Handel-Mazzetti 302 hb VERDOORN 10645 (as Fr. muscicola), in hb FH, in hb NICH [isotypus] Basionym: Distribution: spec nov - comb nov Verdoorn in Handel-Mazzetti 1930 Schuster & Hattori in Hattori Hattori & Lin Piippo Mizutani, Hasegawa & Iwatsuki 1978 1985 1990 2009 550 Symb Sinicae 5: 38 [described it under Fr. muscicola, without naming it] 482 J Hatt Bot Lab 44: 547-548, fig 215 505 J Hatt Bot Lab 59: 135 [comb nov; as Fr. fuscovirens var gemmipara] 521 J Hatt Bot Lab 68: 41 [annotated catalogue of China] 2112 Type sepecimens Hatt Bot Lab (NICH) 24: Isotype. China, Yunnan. 2200 m. 27 Feb. 1914. H. F. v. Handel Mazzetti 302 (NICH 431929). Another isotype specimen: NICH 431930. = Frullania fuscovirens var. gemmipara (R.M.Schust. & S.Hatt.) S.Hatt. & P.J. Lin 1985 Page 166 germana (Taylor) Taylor 1845 Typus: in hb TAYLOR [FH] one specimen "at Turk near Killarney, 1832" as Fr. germana; according GROLLE 1970 in hb GLM (hb LEHMANN) 06029 Basionym: Jungermannia germana Taylor 1843 Distribution: British Isl, Ireland, Portugal, Norway, Madeira [Portugal], Canary Isl [Spain] spec nov Taylor in G., L. & N. Dumortier Lindberg Schiffner 1845 1874 1875 1895 2850 580 1582 1015 Schiffner in Engler & Prantl Bryhn Luisier Stephani Müller in Rabenhorst Schiffner 1905 1908 1910 1911 1915 1936 809 1585 913 686 1064 2561 Schiffner Ade & Koppe Mägdefrau Jelenc Jovet-Ast Arnell Müller in Rabenhorst Arnell Bonner Grolle 1936 1942 1943 1950 1956 1956 1958 1961 1965 1970 437 206 2799 1707 955 43 1028 180 655 615 Hattori Grolle Vanden Berghen Hu & Wang Grolle Stephani 1972 1976 1976 1981 1983 1985 536 183 185 Piippo Wang & Tau Ros, Cano & Guerra Ros, Cano & Guerra 1990 1992 1999 1999 521 144 928 759 1013 Syn Hepat: 450 Bull Soc Roy Bot Belgique 13: 29 [synonyms] Acta Societatis Scientiarum Fennicae 10: 475 Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 134 Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 134 Kgl Norske Vid Selks Skrift 8: 12 [Bryoph Archipel Canar, Teneriffa] Broteria, ser Bot 8 (1): 34 [Madeira] Spec Hepat 4: 565 Krypt Flora (2ed) 6, 2: 616-618 Krit Bemerk europ Leberm, ser 21: 3 [Hepat europ exsicc, nr. 1002, 1003 pp] Österr Bot Zeitschr 85 (2): 126 [Ireland: Killarney] Hedwigia 81 (1/2): 4 & 34 [Madeira] Flora NF 35: 136 [Tenerife] Soc Geogr Arch Prov d'Oran 74 (fasc 228): 72 [North Africa] Rev Bryol Lichen NS 25 (1/2): 136-158 [Maroc] Moss Flora of Fennoscandia: 257 [Norway] Krypt Flora (3ed) 6, 2: 1236-1238 Svensk Bot Tidskr 55 (2): 392 [Tenerife] Index Hepat 5: 311 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 313-314 [may be syn of Fr. teneriffae] J Hatt Bot Lab 35: 243-244 [syn of Fr. teneriffae] Feddes Repert 87 (3/4): 195 [types] Bull Jard Bot Nat Belg 46: <xx> J East China Normal Univ (Nat Sci) 1981 (1): 85-104[aus 698] J Bryol 12: ? <403-459> Stephani Icon Hepat Inedit [Thyopsiella] F-193, microfiche 60 fig 4151 ["Hiberina, ...."] - 4152 J Hatt Bot Lab 68: 41 [annotated catalogue of China] [aus 698] J Bryol 21: 235 [checklist of Northern Africa] http://www.um.es/dp-biologia-vegetal/lab-briologia/Check-list.PDF:40 [checklist of Northern Africa] = Frullania teneriffae var. germana (Taylor) Bisang, Schumacker, Sérgio & Grolle 1988 germana var. explanata Macvicar 1912 Typus: Great Britain. Isl of Mull Basionym: Distribution: Great Britain - syn nov Macvicar Müller in Rabenhorst Schiffner Müller in Rabenhorst Bonner = Frullania germana 1912 1915 1936 1958 1965 1064 2561 1028 655 Stud Handbook Brit Hepat: 437 Krypt Flora (2ed) 6, 2: 616 Krit Bemerk europ Leberm, ser 21: 3 [Hepat europ exsicc, nr. 1003pp] Krypt Flora (3ed) 6, 2: 1237, fig 481 [only oecological modification] Index Hepat 5: 311 (Taylor) Taylor 1845 Frullania teneriffae var. germana (Taylor) Bisang, Schumacker, Sérgio & Grolle 1988 Page 167 gibbosa var. densissima Spruce 1884 Typus: Peru. Peruvian Andes, Tarapoto sd, Spruce in hb MANCH in hb G, hb BM [isotypus] Basionym: Distribution: Peru var nov Spruce Bonner Menzel Yuzawa - syn nov 1884 1965 1984 1984 = Frullania gibbosa gigantea 243 655 142 502 Trans Proc Bot Soc Edinburgh 15: 22-23 Index Hepat 5: 311 Willdenowia 14: 489 [Peru] J Hatt Bot Lab 57: 393-394, fig. 9(a-q) [syn sub Fr. gibbosa] Nees 1840 Steph. ex Theriot 1910 [nom nudum] Typus: in hb PH 3406 > see fr2396.png Basionym: Distribution: nom nud Stephani ex Theriot = Frullania gigantea gigantiloba 1910 1915 Bull Acad Int Geogr Bot 20: 104 Steph. 1911 Steph. ex Bonner 1965 [nom herb] Basionym: Distribution: Bonner Miller, Whittier & Whittier Hattori 1965 1983 1986 655 609 506 Stephani Bonner Yano Yuzawa Stephani 1910 1965 1984 1984 1985 686 655 501 502 928 Yano 2008 = Frullania gigantea glauca Index Hepat 5: 312 Prod Flora Hepat Polynesiae: 127 J Hatt Bot Lab 60: 214 [nom nud] Steph. 1911 Steph. 1910 Typus: Brazil: Brasilia sd, Ule 338 in hb G 20964 Basionym: Distribution: Brazil spec nov - syn nov = Frullania obscura Spec Hepat 4: 333 Index Hepat 5: 312 J Hatt Bot Lab 56: 504 [checklist Brazil] J Hatt Bot Lab 57: 391-393, fig. 8(a-p) Stephani Icon Hepat Inedit [Chonanthelia] A- 33, microfiche 46 fig 3117 from type; "Ue 338"] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr arecae; Brazil; catalog] (Sw.) Mont. 1840 Page 168 glomerata {Java} [auct. non (L.& L. in L.) Lehm. & Lindenb. ex Mont. 1838] Sande Lac. 1857 Typus: hb SANDE LACOSTE in hb LUGD BAT Basionym: Distribution: - syn nov Sande Lacoste Schiffner Verdoorn Dauphin 1857 1898 1929 2005 = Frullania squarrosa Verh Konikl Akad Wetenschappen 5: Conspect Hepat Archip Indici, Batavia: 344 Ann Bryol 2: 156 [Fr. glomerata ex Java by SL 1857 = squarrosa] Tropical Bryology 26: 141-218 [Catalogue of Costa Rican Hepaticae and Anthocerotae] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides glomerata 430 831 246 1360 (Nees) Mont. 1839 Mont. 1840 ex Lehm. & Lindenb. 1845 [pro syn] Basionym: Distribution: - syn nov Montagne 1840 Lehmann & Lindenberg in G., L. 1845 & N. 467 Ann Sci Nat Bot, ser 2, 14: No 34 2850 Syn Hepat: 428 [syn sub Fr. trinervis] = Frullania trinervis var. berteroana Lehm. & Lindenb. 1845 Frullania trinervis glomerata South} (Lehm.) Drège 1843 [auct. non (L.& L. in L.) Lehm. & Lindenb. ex Mont. 1838] Mont. 1842 {India, Typus: India. Nilgiri Hills Basionym: Distribution: India (South) Montagne Bapna & Kachroo 1842 2000 469 712 Ann Sci Nat Bot, ser 2, 18: 13 Hepaticology in India II: 197 [distr: South India & America] = glomerata var. sehnemii (Lusier & Herzog) Herzog 1952 Basionym: Frullania sehnemii Distribution: Brazil, Argentina var nov - syn nov Herzog Bonner Reiner But = Frullania ericoides Luisier & Herzog in Herzog 1936 1952 1965 1988 2004 803 655 1043 147 Feddes Repert 55 (1): 10 [Argentina] Index Hepat 5: 312 Bol Soc Argentina Bot 25 (3-4): 318 [syn sub Fr. ericoides] Checklist of Hong Kong (Nees) Mont. 1839 Page 169 goebeliana Steph. ex Herzog 1916 Typus: Bolivia. Tablas, 3400m 1911, Herzog 4569/a in hb G sn [holotypus] Basionym: Distribution: Bolivia spec nov Stephani Stephani Bonner Stephani 1916 1924 1965 1985 - syn nov Uribe & Gradstein Uribe 2003 2008 = Frullania brasiliensis goetzeana 222 687 655 928 Biblioth Bot 87: 245, fig 197 [Bolivia] Spec Hepat 6: 548-549 [as spec nov] Index Hepat 5: 312-313 Stephani Icon Hepat Inedit [Meteriopsis] E-21, microfiche 55 fig 3795 ["Bolivia, Herzog No 4569a"] 1071 Crypt Bryol 24 (3): 204 ["goebelliana", syn sub Fr brasiliensis] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Raddi 1822 Steph. 1901 [non Steph. 1911] Typus: E Africa: Uasfua, Porotoberg, 2300 m 17.6.1899, W. Goetze 1047a in hb G (ex hb B) in hb L L0060928 (1898, Goetze, W. 1047 a - Deutsch-Ostafrika: Usafua) in hb E E00002533 [isotypus] Basionym: Distribution: spec nov comb nov Stephani Stephani Demaret & Vanden Berghen Bonner Stephani 1901 1911 1950 1965 1985 Wigginton 2004 Wigginton Uribe 2004 2008 = Frullania angulata goetzeana 215 686 60 655 928 Bot Jahrb 30: 261 Spec Hepat 4: 595 [nom superfl] Bull Jard Bot Etat Bruxelles 20: 9-10 [as Fr. angulata var goetzeana] Index Hepat 5: 313 Stephani Icon Hepat Inedit [Meteriopsis] E- 3ab, microfiche 55 fig 3765 ["Africa orient, Usafua, Poroto Berg, 1047a"] - 3766 - 3767 [Usambara Kesai, 2000m, Albers No 405"] 1172 Sript Bot Belg 30: 214, fig 136(G,K,L) from type [syn sub Fr angulata, West Africa] 1172 Sript Bot Belg 30: 214-215, fig 136(A-T) [West Africa] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Mitt. 1863 Steph. 1911 [nom superfl] [non Steph. 1902] Basionym: Distribution: Stephani Uribe = Frullania goetzeana Frullania angulata 1911 2008 686 Spec Hepat 4: 595 [spec nov = nom superfl] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Steph. 1901 [non Steph. 1911] Mitt. 1863 Page 170 gollani Steph. 1910 Typus: India. E Punjab, Simla Distr, summit of Mt. Jako, 8000ft 29.8.1904, W. Gollan hb LEVIER 5700, in hb G 7935 Basionym: Distribution: India spec nov - syn nov Stephani Kashyap Chopra Parihar Bonner Hattori Stephani 1910 1932 1943 1962 1965 1978 1985 Parihar, Lal & Katiyar Bapna & Kachroo Singh, Kumar & Nath 1994 2000 2008 = Frullania polyptera gottscheana Typus: Brazil 686 226 205 1584 655 482 928 Spec Hepat 4: 445-446 Liverworts of the western Himalayas, Part 2: 17 J Indian Bot Soc 22: 249 [India] Univ Allahabad studies, Bot sect 1-2: 18 [India: WH] Index Hepat 5: 313 J Hatt Bot Lab 44: 546-547 [syn nov sub Fr. polyptera] Stephani Icon Hepat Inedit [Galeiloba] C-135, microfiche 52 fig 3527 [from type; "Himalaya, Simla, Gollan legit, Levier 5700"] 2196 Hepat Anth India, annotated checklist: 21 [Western Himalaya] 712 Hepaticology in India II: 178 [syn sub Fr. polyptera] 1281 Taiwania 53 (1): 70, [syn sub Fr muscicola] Taylor 1846 Lindenb. 1846 sd, Beyrich in hb S (B22861) in hb GLM 05986 in hb STR Basionym: Distribution: Brazil spec nov - syn nov Lindenberg in G., L. & N. Stephani Bonner Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot = Frullania griffithsiana gracilis 1846 1911 1965 2002 2849 686 655 782 Syn Hepat: 466 Spec Hepat 4: 496 [syn sub Fr. griffithsiana] Index Hepat 5: 313 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] Gottsche 1846 [auct. non (Reinwardt, Blume & Nees) Gottsche 1845] YANO 1996 {Brazil] Basionym: Distribution: Yano Gradstein & Costa Yano 1996 2003 2008 2094 Bol Inst Bot, Sao Paulo 10: <47-232> 984 Mem New York Bot Gard 87: 91 [not in Brazil] 1684 Boletim Instituto Botanica 19: 31 [Brazil; catalog] = gracilis Hampe ex Gottsche 1845 [pro syn] Basionym: Distribution: Gottsche in G., L. & N. = Frullania junghuhniana 1845 2850 Syn Hepat: 444 [syn sub Fr junghuhniana] Gottsche 1845 Page 171 gracilis fo. distincte-ocellata S.Hatt. 1975 Typus: Singapore 04.06.1952 Basionym: Distribution: Singapore, Indonesia (Sumatra), Borneo fo nov - syn nov Hattori Hattori Hattori Hattori Mizutani, Hasegawa & Iwatsuki = Frullania gracilis gracilis 1975 1975 1976 1982 2009 65 547 80 495 2112 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (3): 112 J Hatt Bot Lab 39: 288-289, fig 144 [fo nov] J Hatt Bot Lab 40: 476 J Hatt Bot Lab 51: 217 Type sepecimens Hatt Bot Lab (NICH) 25; Holotype. Sumatra, between Sumato and Bukit Tinggi, Sipisang. On bark. 04 June 1952. R. van der Wijk 1320A (NICH 432545). (Reinw., Blume & Nees) Nees 1845 fo. subintegra S.Hatt. 1974 Typus: Philippines. Luzon Basionym: Distribution: fo nov - syn nov Hattori 1974 Hattori 1982 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania gracilis 544 J Hatt Bot Lab 38: 174-176, fig 3 495 J Hatt Bot Lab 51: 217 [syn sub Fr. gracilis] 2112 Type sepecimens Hatt Bot Lab (NICH) 25: Holotype. Philippines, Luzon I., Mt. Bulusan. 450-900 m. On tree trunk. 31 May 1965. Z. lwatsuki et al. 13766 (NICH 265935). (Reinw., Blume & Nees) Nees 1845 gracilis var. lacerifolia (Steph.) Verd. 1929 Basionym: Frullania lacerifolia Distribution: comb nov - syn nov Steph. 1911 Verdoorn Verdoorn Verdoorn Bonner Hattori = Frullania gracilis 1929 1930 1932 1965 1982 245 293 553 655 495 Ann Bryol 2: 144 [comb nov] [Java] Ann Bryol Suppl 1: 114-115 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 59 [Sumatra] Index Hepat 5: 314 J Hatt Bot Lab 51: 217 [syn sub Fr. gracilis] (Reinw., Blume & Nees) Nees 1845 gracilis var. subapiculata Schiffn. 1900 [nom nud] Typus: Java. Cibodas Basionym: Distribution: nom nud Schiffner 1900 904 Hedwigia 39: 207 = Frullania gracilis var. subapiculata Schiffn. ex Verd. 1929 Frullania gracilis (Reinw., Blume & Nees) Nees 1845 Page 172 gracilis var. subapiculata Schiffn. ex Verd. 1929 Typus: Indonesia. Java sd, Schiffner 2317 - 2323 Basionym: Distribution: var nov - syn nov Verdoorn Verdoorn Verdoorn Bonner Hattori = Frullania gracilis gracilis ssp. minor Basionym: Frullania minor Distribution: Hattori Hattori & Lin = Frullania gracilis 1929 1930 1932 1965 1982 245 293 553 655 495 Ann Bryol 2: 145 [Java] Ann Bryol Suppl 1: 115, fig 211-212 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 59 [Malay Peninsula, Java] Index Hepat 5: 314 J Hatt Bot Lab 51: 217 [syn sub Fr. gracilis] (Reinw., Blume & Nees) Nees 1845 (Sande Lac.) S.Hatt. 1975 Sande Lac. 1854 [non Raddi 1818] 1976 1985 67 505 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 10-13, fig 172-173 J Hatt Bot Lab 59: 159 [syn sub Fr. gracilis] (Reinw., Blume & Nees) Nees 1845 gracilis ssp. minor var. exorta (Verd.) S.Hatt. 1976 Basionym: Frullania minor var. exorta Verd. 1929 Distribution: Hattori Hattori & Lin = Frullania gracilis 1976 1985 67 505 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 13 J Hatt Bot Lab 59: 158 [syn sub Fr. gacilis] (Reinw., Blume & Nees) Nees 1845 gracilis ssp. minor var. subsinuata (Schiffn. in Verd.) S.Hatt. 1976 Basionym: Frullania minor var. subsinuata Schiffn. in Verd. 1929 Distribution: Hattori Hattori & Lin = Frullania gracilis 1976 1985 67 505 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 13 J Hatt Bot Lab 59: 158 [syn sub Fr. gracilis] (Reinw., Blume & Nees) Nees 1845 Page 173 gracillima Steph. 1910 {from India} [non Steph. ex Steph. 1985] Typus: India. "India orientalis" sd, Kurz 338 in hb G 7937 Basionym: Distribution: India, ? Australia (NSW) spec nov - syn nov Stephani Verdoorn Kashyap Verdoorn Chopra Chopra Parihar Bonner Stephani 1910 1930 1932 1932 1938 1943 1962 1965 1985 Parihar, Lal & Katiyar 1994 Bapna & Kachroo McCarthy 2000 2006 = Frullania polyptera graeffeana 686 293 226 554 933 205 1584 655 928 Spec Hepat 4: 437 Ann Bryol Suppl 1: 41 [may be identical with Fr. campanulata] Liverworts of the western Himalayas, Part 2: 14-15, pl 1(13-15) Nederl Kruidk Arch Jrg 1932 (2): 486-487 [S-Indien, Himalaya] Proceed Indian Acad Sci, sect B, 7 (5): 248 [South India] J Indian Bot Soc 22: 249 [India] Univ Allahabad studies, Bot sect 1-2: 18 [India: WH SI] Index Hepat 5: 314 Stephani Icon Hepat Inedit [Galeiloba] C-136, microfiche 52 fig 3528 ["India orient (Kurz), (sine loci natali)"] 2196 Hepat Anth India, annotated checklist: 41 [as grassilima syn sub Fr polyptera] 712 Hepaticology in India II: 178 [syn nov sub Fr. polyptera] 1385 Checklist Australian Liverworts [Australia xxx] Taylor 1846 Steph. ex Paris 1910 [nom nud] Basionym: Distribution: Paris 1910 = Frullania graeffeana 1455 Rev Bryol 37 (6): 130 [New Caledonia: Malaoni] Steph. 1911 Frullania intermedia var. non-apiculata S.Hatt. 1975 graeffeana Typus: Samoa Steph. 1911 sd, Graeffe in hb G Basionym: Distribution: New Caledonia, Samoa spec nov - syn nov Stephani Verdoorn Verdoorn Verdoorn Bonner Grolle & Schultze-Motel Hattori Stephani 1911 1928 1930 1930 1965 1972 1980 1985 686 242 293 551 655 537 Hattori 1986 506 928 Spec Hepat 4: 585 Rev Bryol NS 1: 120 [syn sub Fr. intermedia] Ann Bryol Suppl 1: 161 Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 314 J Hatt Bot Lab 36: 83 [sub Fr. intermedia] [Samoa] : 193 Stephani Icon Hepat Inedit [Homotropantha] D-20, microfiche 54 fig 3734 ["Samoa, Graeffe"] J Hatt Bot Lab 60: 221 [syn sub Fr. intermedia var. non-apiculata] = Frullania intermedia var. non-apiculata S.Hatt. 1975 grandiclada Steph. ex Paris 1910 [nom nud] Basionym: Distribution: Paris Miller, Whittier & Whittier = Frullania grandiclava 1910 1983 1455 Rev Bryol 37 (6): 130 [New Caledonia: Entre Farino et Table Unio] 609 Prod Flora Hepat Polynesiae: 127 Steph. 1911 Page 174 grandifolia Steph. ex Steph. & Watts 1914 [pro syn] Basionym: Distribution: Stephani & Watts Stephani 1914 1985 Bernal, Gradstein & Celis 2016 = Frullania howeana grandiloba 220 928 J Proc Roy Soc New S Wales 48: 107 Stephani Icon Hepat Inedit [Galeiloba] C-213, microfiche 53 fig 3625 ["Lord Howe Island (Australia), legit Watts (olim grandifolia St, Name besetzt)"] - 3626 4290 Catálogo de plantas y líquenes de Colombia. Vol. I. [new to Colombia] Steph. 1914 Steph. 1924 Typus: Mexico: ? 1913, Arsén 7807 in hb G 20965 [holotypus] Basionym: Distribution: Mexico spec nov - syn nov Stephani Bonner Yuzawa Stephani 1924 1965 1984 1985 687 655 502 928 Fulford & Sharp 1990 105 = Frullania pluricarinata Spec Hepat 6: 532 Index Hepat 5: 315 J Hatt Bot Lab 57: 395-396, fig. 10(a-p) [syn sub Fr. pluricarinata] Stephani Icon Hepat Inedit [Chonanthelia] A- 70, microfiche 47 fig 3170 [from type; "Mexico Arsén 7897"] Mem NY Bot Gard 63: 42 [Mexico] Gottsche 1864 grandistipula var. rugosa (Mitt.) S.Hatt. 1975 Basionym: Frullania rugosa Mitt. 1861 Distribution: India (North: Assam, Meghalaya), Sri Lanka, Indonesia (Sumatra, Java) comb nov Hattori 1975 47 Hattori 1986 74 Bapna & Kachroo 2000 712 = Frullania grandistipula Lindenb. 1845 [non Steph. 1924] grandistipula [pp from Bolivia] Typus: Bolivia Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 78-81, fig 2 from lectotype [About mitten´s specimen] - [type: 1441, Matale, Ceylon, dec 1846, lectotype in hb NY] Bull Natn Sci Mus Tokyo, ser B (Bot), 12 (4): 136 [syn sub Fr. grandistipula; rugosa as extrem habitat modification] Hepaticology in India II: 172 [as valid var; Khasi - Assam, Ceylon, Java, Sumatra] Steph. 1924 [hom illeg] [non Lindenb. in G., L. & N. 1845] sd, Herzog 2608 pp in hb G Basionym: Distribution: Bolivia Stephani Bonner Stephani 1924 1965 1985 687 655 928 Fulford & Sharp Gradstein, Meneses & Arbe 1990 2003 105 975 = Frullania chodatii Spec Hepat 6: 533 [syn sub Fr. chodatii] [vide #189] Index Hepat 5: 315-316 Stephani Icon Hepat Inedit [Chonanthelia] A- 71, microfiche 47 fig 3171-3172 ["Bolkivia leg D Herzog 2608 (ex parte)"] Mem NY Bot Gard 63: 43 [Mexico] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr chodatii; according Yuzawa 1991 ?] Beauverd 1924 Page 175 grandistipula [pp from Mexico] Typus: Mexico Steph. 1924 [hom illeg] [non Lindenb. in G., L. & N. 1845] sd, Arsén 7783 in hb G Basionym: Distribution: Stephani 1924 = Frullania glomerata Mont 1842] grandistipulata 687 Spec Hepat 6: 533 [syn sub Fr. chodatii] [vide #188] (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Lindenb. by Verd. 1932 [orth var] Basionym: Distribution: orth var Verdoorn 1932 = Frullania grandistipula grandistyla 553 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 55 [Java] Lindenb. 1845 [non Steph. 1924] Steph. in Steph. 1985 [nom herb] {New Caledonia} Typus: New Caledonia Basionym: Distribution: New Caledonia Stephani 1985 928 Stephani 1985 928 Stephani Icon Hepat Inedit [Galeiloba] C-133, microfiche 52 fig 3525 ["(Syn: grandistyla St)" = syn sub Fr gigantea] = error Stephani Icon Hepat Inedit [Chonanthelia] A- 87, microfiche 47 fig 3193 [from type; "NCaledonia Herb Lenormand"] = grassilima Steph. {from India} by Parihar, Lal & Katiyar 1994 [orth var] Basionym: Distribution: orth var Parihar, Lal & Katiyar = Frullania gracillima 2196 Hepat Anth India, annotated checklist: 41 [syn sub Fr polyptera] Steph. 1910 {from India} [non Steph. ex Steph. 1985] Frullania polyptera gravana 1994 Taylor 1846 Steph. by S.Hatt. 1978 [orth var] Basionym: Distribution: Hattori = Frullania grayana 1978 416 Misc Bryol Lichen 8 (3): 46 Steph. 1924 [hom illeg] [non Mont. corr Sullivant 1856] Frullania deplanata Mitt. 1855 Page 176 grayana Steph. 1924 [hom illeg] [non Mont. corr Sullivant 1856] Typus: New Zealand. Mummer`s Bush, Waerarapa -.8.1910, W. Gray hb LILLIE B 3946, in hb G Basionym: Distribution: Australia, New Zealand - syn nov Stephani Hodgson Bonner Hamlin Hattori Hattori Stephani 1924 1949 1965 1972 1978 1983 1985 Scott & Bradshaw Piippo Carthy McCarthy 1986 1990 2003 2006 = Frullania deplanata grayana Spec Hepat 6: 539-540 Trans Roy Soc New Zealand 77 (3): 381-382 [syn sub Fr deplanata] Index Hepat 5: 316 Rec Dominion Mus 7(19): 262 [New Zealand] Misc Bryol Lichen 8 (3): 46 [sub Fr. scandens auct =Fr. deplanata] J Hatt Bot Lab 54: 140-142 [sub Fr. deplanata] Stephani Icon Hepat Inedit [Galeiloba] C-263, microfiche 54 fig 3685 [Nova Zelandia, leg W Gray 1810, Herb Lillie No 202 1912"] 56 Brunnonia 8 (1): 35 [Australia] 521 J Hatt Bot Lab 68: 40 [annotated catalogue of China] 1125 Flora Australia Suppl Ser 21: 42 [sub Fr deplanata] 1385 Checklist Australian Liverworts [Australia, syn sub Fr deplanata] Mitt. 1855 Mont. corr. Sull. 1856 [non Steph. 1924] [nom inval] Basionym: Frullania asagrayana Distribution: USA, Antilles - syn nov 687 203 655 659 416 499 928 Mont. 1842 Sullivant Lehnert Stephani Frye & Clark Bonner = Frullania asagrayana 1856 1886 1911 1947 1965 1499 2089 686 204 655 Gray`s Manual of Botany, 2. ed.: 697 <607-737> pl 1-8, Cambridge Proceed Biol Soc Washington 3: 117 [Washington andd vicinity] Spec Hepat 4: 566 [not mentioned as syn of Fr. asagrayana) Univ Wash Publ Biol 6 (5): 751 Index Hepat 5: 316 [N-America, Antilles] Mont. 1842 grayana var. californica pp A Austin ex Underwood 1884 Basionym: Distribution: var nov Austin ex Underwood Frye & Clark Hattori in Hara = Frullania tricarinata 1884 1947 1966 1708 Bull Illinois State Lab Nat Hist 2: 67 204 Univ Wash Publ Biol 6 (5): 753 841 Flora of Eastern Himalaya: 528 Sande Lac. 1856 grayana var. californica pp B Austin ex Underwood 1844 Typus: California sd, Bolander in hb G, in hb US ? Basionym: Distribution: var nov Austin ex Underwood Howe Frye & Clark Bonner Hattori in Hara NN = Frullania californica 1884 1894 1947 1965 1966 1996 1708 570 204 655 841 632 Bull Illinois State Lab Nat Hist 2: 67 Erythea 2: 98-99 Univ Wash Publ Biol 6 (5): 753 [sub Fr franciscana] Index Hepat 5: 316 Flora of Eastern Himalaya: 528 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 02 of 35 [in hb US] (M.Howe) A.Evans 1897 Page 177 grebeana Steph. 1911 Typus: Malaysia. Borneo. Tamelayang sd, G. Grebe in hb G [holoype] Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Horikawa Kamimura Bonner Stephani 1911 1930 1934 1961 1965 1985 Uribe-M. 2011 = Frullania tenuicaulis Frullania vethii Typus: Nepal. Spec Hepat 4: 537 Ann Bryol Suppl 1: 151 J Sci Hiroshima Univ, ser B, div 2, 2: 236 [syn sub Fr. tenuicaulis] J Hatt Bot Lab 24: 61 [syn sub Fr tenuicaulis] Index Hepat 5: 316 Stephani Icon Hepat Inedit [Thyopsiella] F-142, microfiche 59 fig 4078 ["Borneo, Tamelayang, com C. Grebe"] 2767 Caldasia 33(1): 74 [type study, syn sub Fr trichodes] Mitt. 1891 Sande Lac. 1884 Frullania trichodes grevilleana 686 293 34 199 655 928 Mitt. 1862 Taylor 1845 sd, Wallich sn in hb GREVILLE (sub nomine "Fr. nepalensis") in hb E E00002565 (sd, Wallich sn) [holotypus] - E00293983 (sd, Greville sn) [isotypus] - E00293982 (14.6.1843, Greville sn) [isotypus] in hb GLM 05984, 06019 Basionym: Distribution: India (Eastern Himalayas: Megalaya) spec nov - syn nov Taylor in G., L. & N. Mitten Stephani Kashyap Chopra 1845 1861 1911 1932 1938 2850 574 686 226 1648 Chopra Parihar Bonner Hattori in Hara Grolle Hattori in Hara Hattori in Mizutani 1943 1962 1965 1966 1966 1971 1979 205 1584 655 841 843 190 487 Hattori Stephani 1982 1985 495 928 Piippo Parihar, Lal & Katiyar 1990 1994 521 2196 Bapna & Kachroo Singh, Kumar & Nath Pradhan, Joshi Rawat, Verma, Alam 2000 2008 2009 2015 712 1281 1941 4266 = Frullania nepalensis Syn Hepat: 421-422 J Linn Soc Bot 5: 121 [syn nov sub Fr. nepalensis] Spec Hepat 4: 450 Liverworts of the western Himalayas, Part 2: 16-17, pl 2(4-7) Proceed Indian Acad Sci, sect B, 8: 437 [Sikkim Himalayas and Bengal] J Indian Bot Soc 22: 249 [India] Univ Allahabad studies, Bot sect 1-2: 18 [India: EH WH] Index Hepat 5: 316-317 Flora of Eastern Himalaya: 526 [most common in E. Himalaya] Khumbu Himal 4: 266 [Nepal] Univ Mus Univ Tokyo Bull 2: 236-237 [Bhutan] J Hatt Bot Lab 46: 322-323 [Easter Nepal] [syn sub Fr. nepalensis var. grevilleana] J Hatt Bot Lab 51: 250 [syn sub Fr. nepalensis] Stephani Icon Hepat Inedit [Galeiloba] C-139, microfiche 52 fig 3533 ["Nepal, Originale: Hook. Thomson 1526, 1542"] J Hatt Bot Lab 68: 41 [annotated catalogue of China] Hepat Anth India, annotated checklist: 41 [syn sub Fr nepalensis var grevilleana] Hepaticology in India II: 176 [syn sub Fr. nepalensis] Taiwania 53 (1): 63, [syn sub Fr nepalensis] Botanica Orientalis 6(6) <69-75> [Nepal] Plant Science Today 2(4) <179-183> (Spreng.) Lehm. & Lindenb. 1845 Page 178 grevilleana var. densa Schiffn. ex Verd. 1932 Typus: Sikkim. Kurseong, 6800ft -.10.1898, Decoly Type:—INDIA. Sikkim: Kurseong, October 1898, Decoly Basionym: Distribution: var nov - syn nov Verdoorn Chopra 1932 1938 Bonner Hentschel et alii 1965 2015 = Frullania nepalensis 554 Nederl Kruidk Arch Jrg 1932 (2): 487-488 1648 Proceed Indian Acad Sci, sect B, 8: 437 [Sikkim Himalayas and Bengal] 655 Index Hepat 5: 317 4097 Phytotaxa 220 (2): 136 [syn sub Fr. nepalensis] (Spreng.) Lehm. & Lindenb. 1845 grevilleana var. gebaueri Verd. 1932 Typus: India. Darjeeling, Kataphar, 2500m -.4.1915, Gebauer in hb VERDOORN & hb W Basionym: Distribution: var nov -syn nov Verdoorn Bonner Hentschel et alii = Frullania nepalensis griffiithiana 1932 1965 2015 552 Ann Bryol 5: 164 655 Index Hepat 5: 317 4097 Phytotaxa 220 (2): 136 [syn sub Fr nepalensis] (Spreng.) Lehm. & Lindenb. 1845 Gottsche by Steph. 1911 [orth var] Basionym: Distribution: Stephani Stephani = Frullania griffithsiana grolleana 1911 1985 686 928 Spec Hepat 4: 496-497 Stephani Icon Hepat Inedit [Thyopsiella] F- 58ab, microfiche 57 fig 3956 ["syn.Frull Gottscheana, Am merid (griff leg) Herb Jack"] - 3957 Gottsche 1846 S.Hatt. 1972 Typus: PNG. New Guinea. Wau Distr, Summit of Mt. Kaindi, 2200-2300m 26-27.5.1967, R.M. Schuster 67-6158 in hb NICH [holotypus] in hb JE, MASS [isotypus] Basionym: Distribution: New Guinea (Papua New Guinea) spec nov Hattori Hattori Hattori Grolle & Piippo Hattori 1972 1973 1982 1984 1988 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania serrata ssp. grolleana 539 542 495 143 516 J Hatt Bot Lab 36: 416-417, fig 3 J Hatt Bot Lab 37: 149 [New Guinea] J Hatt Bot Lab 51: 225 [as Fr. serrata ssp. grolleana] [comb et stat nov] Acta Bot Fennica 125: 19 [Cat. West Melanesia] J Hatt Bot Lab 65: 428-431, fig 6 [grolleana as distinct species, key to grolleana & serrata] 2112 Type sepecimens Hatt Bot Lab (NICH) 25: Holotype. New Guinea, Wau District, Summit of Mt. Kaindi. 2200-2350 m. May 1967. R. M. Schuster 67-6158 (NICH 309938). (S.Hatt.) S.Hatt. 1982 Page 179 grollei Hamlin 1972 [nom superfl.] Basionym: Distribution: New Zealand spec nov - syn nov Hamlin Hamlin Hamlin 1972 1972 1973 Hattori & Mizutani Glenny Glenny & Malcolm 1982 1998 2005 Konrat, Braggins, Asakawa & Toyota 2006 = Frullania congesta grossiloba 659 659 656 Records of the Dominion Museum 7 (19): 262 Rec Dominion Mus 7(19): 262 [New Zealand] Rec Dominian Mus 8 (10): 143 [New Zealand; syn nov sub Fr. congesta] 495 J Hatt Bot Lab 52: 447 [syn sub Fr. rostrata] 1412 Tuhinga 10: 132 [sub Fr rostrata] 1410 ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr rostrata] 1465 J H B L 100: 555, 563 [syn nov sub Fr congesta; nom superfl.] Gottsche, Lindenb. & Nees 1845 Steph. 1910 [non Steph. 1911] Typus: Tasmania. Tasman peninsula, Carnarvon 1.2.1899, W.A. Weymouth hb LEVIER 2167, in hb G 13631 Basionym: Distribution: Tasmania spec nov - syn nov Stephani Stephani & Watts Bonner Hattori Hattori Stephani 1910 1914 1965 1979 1979 1985 Scott & Bradshaw Carthy McCarthy 1986 2003 2006 = Frullania monocera grossiloba 686 220 655 485 486 928 Spec Hepat 4: 424 J Proc Roy Soc New S Wales 48: 107 [Australia] Index Hepat 5: 318 J Hatt Bot Lab 45: 342-343, fig 14(a-g) from type J Hatt Bot Lab 46: 134 [syn sub Fr. monocera] Stephani Icon Hepat Inedit [Galeiloba] C-211, microfiche 53 fig 3622 ["Tasmania, Levier 2167"] - 3623 56 Brunnonia 8 (1): 35 [Australia] 1125 Flora Australia Suppl Ser 21: 44 [sub Fr monocera] 1385 Checklist Australian Liverworts [Australia; syn sub Fr monocera] (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Steph. 1911 [hom illeg] [non Steph. 1910] Typus: Mexico ex hb NEES, ex hb JACK, in hb G Basionym: Distribution: Mexico Stephani Stephani Bonner Stephani 1911 1924 1965 1985 686 909 655 928 Fulford & Sharp 1990 105 = Frullania esenbeckiana Spec Hepat 4: 531-532 [spec nov = hom illeg] Spec Hepat 6: 570 Index Hepat 5: 318 Stephani Icon Hepat Inedit [Thyopsiella] F-112, microfiche 58 fig 4037 ["Mexico, ex Herb Nees, missit Jack"] Mem NY Bot Gard 63: 43 [Mexico] Beauverd 1924 Page 180 grossispica Steph. 1911 Typus: Fiji. Viti Levu ? 1860, B. Seemann 834b in hb G Basionym: Distribution: Fiji spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Miller, Whittier & Whittier Stephani = Frullania serrata grossistipula 1911 1930 1930 1965 1983 1985 686 293 551 655 609 928 Spec Hepat 4: 541 Ann Bryol Suppl 1: 85 Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 318 Prod Flora Hepat Polynesiae: 128 Stephani Icon Hepat Inedit [Thyopsiella] F-143, microfiche 59 fig 4079 ["Insula Vitiensis (leg Dr B Seemann 1860) 834b"] Gottsche 1845 Steph. 1911 Typus: Bolivia. Prov Cochachamba, bei Incacorral, 2200m -.1.1908, T. Herzog hb LEVIER 6030, in hb G Basionym: Distribution: Bolivia spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 686 655 4761 194 928 Gradstein, Meneses & Arbe 2003 975 = Frullania macrocephala guadichaudii Spec Hepat 4: 511-512 Index Hepat 5: 318 Nova Hedwigia 18: 439 [syn sub Fr. macrocephala] The genus Frullania, separata: 439 [syn sub Fr. macrocephala] Stephani Icon Hepat Inedit [Thyopsiella] F- 59, microfiche 57 fig 3958 ["Bolivia, Levier 6030 ex parte"] - 3959 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. macrocephala] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 (Nees & Mont.) Nees & Mont. 1845 [orth var] in Shin 1970 Basionym: Distribution: Shin 1970 = Frullania gaudichaudii gualaquizana 150 Sci Rep Kagoshima Univ 19: 25 (Nees & Mont.) Nees & Mont. 1845 Steph. 1911 Typus: Ecuador. Prov. del Oriente, Gualaquiza -.2.1909, M. Alloni hb LEVIER 6480 pp, in hb G Basionym: Distribution: Ecuador spec nov Stephani Bonner Stephani 1911 1965 1985 -syn nov Gradstein & Perez 2021 = Frullania montagnei 686 655 928 Spec Hepat 4: 531 Index Hepat 5: 531 Stephani Icon Hepat Inedit [Thyopsiella] F-113, microfiche 58 fig 4038 ["Ecuador, Alloni, Levier 6480 ex parte"] 6056 Candollea 76: 44-45 [ Lectotypus, syn sub Fr montagnei - Ecuador] Gottsche 1845 Page 181 gunniana Steph. ex Bonner 1965 [nom herb] Typus: New Hebrides. Aneityum -.3.1911, Dr. Gunn hb WATTS 25, in hb G 20580 Basionym: Distribution: Vanuatu Bonner Hattori Miller, Whittier & Whittier Stephani = Frullania gaudichaudii 1965 1981 1983 1985 655 492 609 928 Index Hepat 5: 319 J Hatt Bot Lab 49: 372 [nom nud, syn sub Fr. gaudichaudii] Prod Flora Hepat Polynesiae: 128 Stephani Icon Hepat Inedit [Galeiloba] C-140, microfiche 52 fig 3534 [from type; "Novae Hebridae, Watts No 25, (Dr Gunn legit) 1911"] (Nees & Mont.) Nees & Mont. 1845 Page 182 gymnotis Nees & Mont. in Mont. 1843 Typus: Guyana. sd, Leprieur 360 or 366 [in one packet] in hb PC (hb MONTAGNE) Basionym: Distribution: USA ( FL, LA), Honduras, Nicaragua, Costa Rica, Cocos Isl [Costa Rica], Panama, Jamaica, Guadeloupe, Venezuela, Colombia, Galapagos Isls [Ecuador], Surinam, Brazil, Hawaii spec nov - sn nov Montagne Montagne & Nees in G., L. & N. Montagne Montagne Stephani Pagán Clark & Svihla Clark 1843 1845 1855 1856 1911 1942 1948 1953 Jovet-Ast Clark Schiffner & Arnell Bonner Winkler Gradstein & Hekking Lemos Michel 1957 1958 1964 1965 1976 1979 1980 Miller, Whittier & Whittier Lemos Michel Yano Stephani 1983 1983 1984 1985 Gradstein & Hekking Schuster Yano Germano & Pôrto Stotler, Allen, Gradstein, McGuinness, Whittemore & Chung Costa & Yano Yano & de Mello Yano Oliviera-e-Silva & Yano Gradstein & Costa Yano Porto, Germano & Borges Yano 1989 1992 1995 1996 1998 Ann Sci Nat Bot, ser 2, 19: 257 Syn Hepat: 444 Ann Sci Nat Bot, ser 4, 3: 320 Sylloge generum specierumque cryptogamaram: 83 Spec Hepat 4: 635 Bryologist 45(4): 108 [Guadeloupe] Bryologist 51: 239-242, fig 1-17 Proc California Acad Sci 4 ser, 27 (18): 663 [Galapagos Isls: Santa Cruz, Cocos Isl] 878 Acta Bot Neerlandica 6: 608 [Suriname] 385 Bryologist 61: 67 198 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 85-86 [Brazil] 655 Index Hepat 5: 319 5 Rev Bryol Lichen 42 (3): 811 [Colombia] 483 J Hatt Bot Lab 45: 112 [Cat Colombia] 719 Diss, Uni Fed do Rio Grande do Sul, Porto Alegre: 103-105, pl 17, map 16, [Brasil, Rio Grande do Sul] 609 Prod Flora Hepat Polynesiae: 128 [= syn of Fr. hypoleuca NEES p.p.] 306 Revta brasil Bot 6 (2): 119-121, fig 14-26 [Brazil] 501 J Hatt Bot Lab 56: 505 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Diastaloba] B-32, microfiche 48 fig 3265 ["Surinam (Wullschlägel legit)"] 517 J Hatt Bot Lab 66: 220 [syn sub Fr. obcordata var armata] 603 Hepat N America 5: 118-120, fig ? 2095 J Hatt Bot Lab 78: 142 [annotated checklist of Brazil] 299 Trop Bryol 12: 23 [Brazil] 703 Trop Bryol 15: 179 [Checklist Panama] 1998 1999 1999 2000 2003 2004 2004 2008 757 919 1263 989 984 1450 1446 1684 Yano Bordin & Yano Lavocat Bernard & SchäferVerwimp Yano 2008 2010 2011 Winter & Schäfer-Verwimp 2020 2012 470 2850 475 1912 686 3152 328 1106 Hoehnea 25 (2): 102 [Brasil] Iheringia, ser Bot 52: 77-78 [Brasil] Briophyta do estado de São Paulo. Lista provisória [Brazil] Boletim Inst Botanica 13: 27 [Rio de Janeiro, Brazil] Mem New York Bot Gard 87: 86 [Brazil] [syn sub Fr caulisequa] Acta Amazonica 34(4): 23 [Brazil] Brejos de Altitude em Pernambuco e Paraiba: 83 [Brazil] Boletim Instituto Botanica 19: 90 [syn sub Fr caulisequa; Brazil; catalog] 1684 Boletim Instituto Botanica 19: 31 [Brazil; catalog] 2266 Pesquisas, sér Bot 61: 49 [Brazil] 3130 Cryptogamie, Bryologie 32(3): 243 [Guadeloupe] "as Fr caulisequa" 3678 Pesquisas, série Botânica 63:55-140 [Catálogo ... do Estado do Espírito Santo, Brasil] 5742 Frahmia 19: 13 [syn sub Fr obcordata] = Frullania obcordata var. armata Lindenb. & Gottsche 1851 Frullania obcordata (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 183 gymnotis [auct. non Nees & Mont. in Mont. 1843] Winkler 1976 Basionym: Distribution: Winkler Uribe & Gradstein = Frullania caulisequa 1976 1998 5 707 Rev Bryol Lichen 42 (3): 811 [Colombia] Bryophyt Bibl 53: 35 [CESAR, MAGDALENA: Winkler (1976) 811 as F. gymnotis Nees & Mont - = Fr. obcordata] (Nees) Mont. 1839 gymnotis var. integriloba Yuzawa 1987 [nom. herb.] Typus: Brazil Schäfer-Verwimp & Verwimp 1986 hb Schäfer-Verwimp 7199 Basionym: Distribution: = haeckeriana Typus: Peru. Lindenb. 1846 sd, Haecker sn ex hb BESCHERELLE, in hb PC & G, holotye in hb W, isotypus in hb G (G20614) Basionym: Distribution: Guatemala, Peru, Bolivia, Ecuador spec nov Lindenberg in G., L. & N. Stephani Stephani Bonner Menzel Stephani 1846 1911 1916 1965 1984 1985 - syn nov Uribe 2008 = Frullania peruviana hallii 2849 686 222 655 142 928 Syn Hepat: 465 [In cortice Chinae Locbae legit cl. Haecker] Spec Hepat 4: 603-604 [Hab. Peruvia, Guatemala] Biblioth Bot 87: 245 [Bolivia] Index Hepat 5: 320 Willdenowia 14: 489 [Peru] Stephani Icon Hepat Inedit [Meteriopsis] E-22, microfiche 55 fig 3796 ["Peruvia, leg Haecker, Herb Mus Paris"] - 3797 1513 Caldasia 30(1): 75 [syn sub Fr peruviana] [Monography of subgenus Meteoriopsis] Gottsche 1846 [non Gottsche ex Spruce 1884] Austin 1875 Typus: USA. Oregon, Salem E. Hall Basionym: Distribution: USA spec nov - syn nov Austin 1875 Pearson Evans Macoun Sanborn Frye & Clark Bonner Stephani 1890 1897 1902 1929 1947 1965 1985 = Frullania bolanderi 2471 Bull Torrey Bot Club 6: [No 3] :20-21 [On trees ; Salem, Oregon, F. Hall.] 1153 List of Canadian Hepaticae: 2 237 Trans Connet Acad Arts Sci 10 (1): 8-10 1150 Catalogue Canadian Plants, VII: 45 [sub Fr bolanderi] 3503 Univ Oregon publ Pl Biol 1(1): 10 [Oregon] 204 Univ Wash Publ Biol 6 (5): 768 [sub Fr bolanderi] 655 Index Hepat 5: 320 928 Stephani Icon Hepat Inedit [Galeiloba] C- 80b, microfiche 51 fig 3447 ["Frull Hallii Austin, Rocky Mts, ex Herb Austin, dedit Pearson 1894"] Austin 1869 Page 184 hamachiloba by Toyota, Nagashima & Asakawa 1988 [orth var] Basionym: Distribution: orth err Toyota, Nagashima & Asakawa 1988 = Frullania hamatiloba hamatiloba 2251 Phytochemistry, 27(6): 1789. 1793 Steph. 1910 Steph. ex Yoshinaga 1906 [nom nud] Basionym: Distribution: Japan, China (Yunnan) - nom nud Stephani in Yoshinaga Sawada 1906 1909 927 930 Hattori Bai Bai Gao & Lai Yamada & Iwatsuki 1944 2001 2002 2003 2006 943 1742 1189 1441 1470 = Frullania hamatiloba hamatiloba Bot Mag Tokyo 20: 53 [Japan] Bot Mag Tokyo 23 (no 268): 178-183, fig 19, 19' [key to Japanese species, in Japanese] Bull Tokyo Sci Mus 11: 143-144 [= nom nud] Guizhou Science 19(4): 38-39 [China, Yunnan] Chenia 7: 9 [Yunnan, China] Illustrations Bryophytes China: 1108, fig 516(7-12) [China] J Hatt Bot Lab 99: 29 [Catalog of Japan] Steph. 1910 fo. grosscappendiculata S.Hatt. by Choe 1980 [orth var] Basionym: Distribution: Korea (South) Choe = Frullania hamatiloba 1980 fo. grosse-appendiculata S.Hatt. 1944 Frullania hamatiloba hamatiloba 1171 Illust Fl & Fauna Korea 24: 733 Steph. 1910 fo. grosse-appendiculata S.Hatt. 1944 Typus: Japan. Honsyu, Miyagi, Pref., Prov Rikuzen, Mt. Kinkawasan -.6.1903, U. Faurie 1077 in hb KYO [holotypus], in hb TNS [isotypus] Basionym: Distribution: Taiwan, Japan stat nov - syn nov Hattori Hattori & Noguchi Kamimura Kamimura Bonner Choe Inoue Kuo & Chiang N.N. [National Science Museum Tokyo] Yuzawa Yamada & Iwatsuki = Frullania hamatiloba 1944 1960 1961 1961 1965 1980 1987 1988 2000 1911 965 199 199 655 1171 1155 948 995 J Jap Bot 20 (2): 155 Index spec Typ herb Japon, Bryophyta: 17 J Hatt Bot Lab 24: J Hatt Bot Lab 24: 33 Index Hepat 5: 321 Illust Fl & Fauna Korea 24: 733 [syn sub Fr hamatiloba; South Korea] Index type specimens in Nat Sci Mus, Tokyo: 12 Taiwania 33: 12 [Index of Taiwan] http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 2001 2006 1660 Natural Environmental Sci Res 14: 16 [Japan, syn sub Fr hamatiloba] 1470 J Hatt Bot Lab 99: 29 [Catalog of Japan] Steph. 1910 Page 185 hamatiloba var. latistipula S.Hatt. 1975 Typus: Japan 11.6.1961, Y. Jkegami 63458 in hb NICH [typus] Basionym: Distribution: Japan - syn nov Hattori Yuzawa & Hattori Furuki & Mizutani Yuzawa 1975 1981 1994 2001 Yamada & Iwatsuki 2006 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania hamatiloba hamatirostra 411 494 1331 1660 Misc Bryol Lichen 7 (3): 49-50, fig J Hatt Bot Lab 50: 255 [Japan] Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] Natural Environmental Sci Res 14: 16, fig 13(a-o) [Japan, syn nov sub Fr hamatiloba] 1470 J Hatt Bot Lab 99: 29 [Catalog of Japan] 2112 Type sepecimens Hatt Bot Lab (NICH) 25: Holotype. Japan, Honshu, Niigata-ken, Sasagami-mura, Deyu. lIJune 1961. Y. lkegami 63458 (NICH 71706). Steph. 1910 Gerola 1947 Typus: Socotra or Uonde, Torrente Hallo 5.11.1937, Vatova or 22.11.1937, Vatova Basionym: Distribution: spec nov Gerola 1947 Cufodontis Bonner Vanden Berghen Wigginton 1952 1965 1976 2002 Wigginton 2004 = Frullania ericoides hamatirostra 1532 Lavori di botanica / Istituto Botanica dell'Università di Padova 12: 476, fig 8(a-d) [Ethiopia] 2562 Phyton 4 (1-3): 76 [Ethiopia] 655 Index Hepat 5: 321 185 Bull Jard Bot Nat Belg 46 (1/2): 13 [syn sub Fr. ericoides] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] (Nees) Mont. 1839 fo. evoluta Gerola 1947 Typus: Prov. Uonde, Torrente Hallo 22.11.1937, Vatova Basionym: Distribution: fo nov Gerola 1947 Cufodontis Bonner Vanden Berghen 1952 1965 1976 = Frullania ericoides 1532 Lavori di botanica / Istituto Botanica dell'Università di Padova 12: 477 [Ethiopia] 2562 Phyton 4 (1-3): 76 [Ethiopia] 655 Index Hepat 5: 321 185 Bull Jard Bot Nat Belg 46 (1/2): 13 [syn sub Fr. ericoides] (Nees) Mont. 1839 Page 186 hampeana Nees 1845 Typus: "Nova Hollandia" in hb HAMPE Basionym: Distribution: Taiwan, Japan, Bonin Isl, Sri Lanka, Indonesia (Sumatra, Java), Australia, Tasmania, New Zealand, New Caledonia, Society Isl (Tahiti) spec nov - syn nov Nees in G., L. & N. Mitten in Hooker Mitten in Hooker Hooker Gottsche in Mueller Bailey Bastow Stephani Sawada 1845 1855 1860 1867 1880 1883 1888 1889 1909 2850 560 561 1154 1489 1104 845 940 930 Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Hattori Hattori Kamimura Kamimura Tatuno Abeywickrama Inoue Kamimura Bonner Kodama 1930 1930 1932 1932 1934 1944 1952 1952 1952 1952 1959 1961 1961 1965 1972 551 293 553 554 241 943 529 769 873 1697 200 964 199 655 1375 Hamlin Iwatsuki & Mizutani Allison & Child Lai & Wang-Yang Hattori Abeywickrama & Jansen Hattori Hattori Miller, Whittier & Whittier Miller, Whittier & Whittier Scott & Bradshaw Windolf Kuo & Chiang Simmonds Piippo Parihar, Lal & Katiyar Glenny Bapna & Kachroo Yuzawa Carthy Glenny & Malcolm 1972 1972 1975 1976 1976 1978 1979 1980 1983 1983 1986 1987 1988 1988 1990 1994 1998 2000 2001 2003 2005 659 966 690 33 80 794 486 489 609 609 56 1025 948 1036 521 2196 1412 712 1660 1125 1410 McCarthy 2006 1385 Yamada & Iwatsuki 2006 1470 = Frullania monocera Syn Hepat: 426-427 Antarctic voyage, II. Fl Novae-Zelandiae 2: 160 [New Zealand] Antarctic voyage, III. Fl Tasmaniae 2: 236 [Tasmania] Handbook New Zealand Flora: 536 [New Zealand, N Island] Frag Phyt Australia 11, suppl Hepaticae :64-65 Syn Queensland Flora: 738 Papers Proceed Roy Soc Tasmania for 1887: 264 [Tasmania] Hedwigia 28: 159 [Australia] Bot Mag Tokyo 23 (no 268): 178-183, fig 6, 6' [key to Japanese species, in Japanese] Nederl Kruidk Arch Jrg 1930 (2): Ann Bryol Suppl 1: 44-45, fig 28, 51-52 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 55 [Java] Nederl Kruidk Arch Jrg 1932 (2): 488 [Java] Ann Bryol Suppl 4: 222, fig 32 [map] Bull Tokyo Sci Mus 11: 144 [Japan, key to Japanese species] J Hatt Bot Lab 8: 29 Acta Phytotax Geobot 14 (4): 109 Contrib Fl Hepat Shikokuensem, Kochi: 124 J Sci Hiroshima Univ ser B, div 2 (Bot), 6(2): 51-57 [sex chromosomes] Ceylon J Sci (Bio Sci) 2 (1): 58 [Sri Lanka] J Jap Bot 36(5): 187 [Formosa] J Hatt Bot Lab 24: 49-50, fig 9(26), map 11 Index Hepat 5: 322 Spec Publ Osaka Mus Nat Hist 4: 125, fig 62(a-f), map p.241 [Central Japan] Rec Dominion Mus 7(19): 262 [New Zealand] Colour Illust Bryo Japan, Osaka: 332, pl 45 [715] Liverw New Zealand: 210-211, fig(a-d) Taiwania 21 (2): 168 [Index to Taiwan] J Hatt Bot Lab 40: 476 MaB Nat Comm Sri Lanka, Publ 1: 5 [Sri Lanka] J Hatt Bot Lab 46: 132-134, fig 36(a-j) [syn nov sub Fr. monocera] J Hatt Bot Lab 47: 103 [syn nov; sub Fr. monocera] Prod Flora Hepat Polynesiae: 131 Prod Flora Hepat Polynesiae: 128 Brunnonia 8 (1): 35 [Australia] Austrobaileya 2(4): 390 [syn sub Fr. monocera var depauperata] Taiwania 33: 12 [Index of Taiwan] Proc Roy Soc Queensland 5 (5): 178 [Queensland] J Hatt Bot Lab 68: 41 [annotated catalogue of China] Hepat Anth India, annotated checklist: 22 [Eastern Himalaya] Tuhinga 10: 132 [sub Fr monocera] Hepaticology in India II: 174 [syn sub Fr. monocera] Natural Environmental Sci Res 14: 23 [Japan, syn sub Fr monocera] Flora Australia Suppl Ser 21: 44 [sub Fr monocera var depauperata] ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr monocera] Checklist Australian Liverworts [Australia, syn sub Fr monocera var depauperata] J Hatt Bot Lab 99: 30 [Catalog of Japan] (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Page 187 hampeana var. acutiloba (Mitt.) S.Hatt. 1975 Basionym: Frullania acutiloba Distribution: comb nov - syn nov Mitt. 1861 [non Gerola 1947] Hattori 1975 47 Hattori Hattori, Thaithong & Kitagawa Hattori Bapna & Kachroo 1975 1977 1980 2000 547 480 489 712 = Frullania acutiloba Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 73-74 [Basionym: Fr. acutiloba MITTEN 1861] J Hatt Bot Lab 39: 289 [India] J Hatt Bot Lab 43: 451 [Thailand] J Hatt Bot Lab 47: 103 [syn nov sub Fr. hampeana] Hepaticology in India II: 165 [syn sub Fr. hampeana] Mitt. 1861 [non Gerola 1947] Frullania monocera var. acutiloba (Mitt.) Hentschel & von Konrat 2015 hampeana var. osumiensis S.Hatt. 1944 Typus: Japan. Kagoshima Pref, Kyushu, Prov. Osumi, Kimotsuki-gun, Tashiro-mura -.4.1939, Hattori 1849 in hb TNS [holotypus] Basionym: Distribution: var nov - stat nov Hattori 1944 Hattori Hattori in Iwatsuki & Hattori Hattori & Noguchi Kamimura Bonner Inoue N.N. [National Science Museum Tokyo] Yuzawa 1952 1956 1960 1961 1965 1987 2000 Yamada & Iwatsuki 2006 = Frullania osumiensis 2001 943 Bull Tokyo Sci Mus 11: 144-146, fig 90(a-n) [Japan, key to Japanese species] 529 J Hatt Bot Lab 8: 29 861 J Hatt Bot Lab 16: 87 [stat nov - Fr osumiensis] 965 Index spec Typ herb Japon, Bryophyta: 17 199 J Hatt Bot Lab 24: 39 655 Index Hepat 5: 322 1155 Index type specimens in Nat Sci Mus, Tokyo: 13 995 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 1660 Natural Environmental Sci Res 14: 28 [Japan, basionym of Fr osumiensis] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] (S.Hatt.) S.Hatt. 1956 hampeana var. undulata (Kamim.) Inoue 1966 Basionym: Frullania undulata Distribution: comb nov - syn nov Kamim. 1961 Inoue Yamada & Iwatsuki = Frullania monocera hariotana 1966 2006 1703 J Jap Bot 41 (1): 15 [comb nov] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Steph. 1911 Typus: Chile, Fuegia, Cape Horn, Ile Hermite sd, Hariot 77 pp ex hb BESCHERELLE, in hb G Basionym: Distribution: Chile (Cape Horn) spec nov Stephani Bonner Stephani - syn nov Hässel de Menéndez & Rubies 2009 = Gackstroemia hariotana 1911 1965 1985 686 655 928 Spec Hepat 4: 667 Index Hepat 5: 322 Stephani Icon Hepat Inedit [Diastaloba] B-44, microfiche 49 fig 3282 [from type; "Fuegia Insula Hermite Hariot No 77 ex parte"] 2773 Nova Hedwigia Beiheft 134: 140 [syn sub Gackstroemia hariotiana, Fr hariotana fo pusilla] (Besch. & C. Massal.) Grolle 1967 Page 188 hariotana fo. pusilla Gola 1923 [nom. nud.] Typus: Chile, Isole del Laberinto De Gaspari C41 Gola Basionym: Distribution: fo nov Gola 1923 Hässel de Menéndez & Rubies 2009 823 Nuovo Giornale botanico italiano. Ns 2, 19: 172 [Fr Hariotiana Steph. F. pusilla - Isole del Labirinto] 2772 Nova Hedwigia Beiheft 134: 140 [Fr hariotana fo pusilla Gola 1923] = hariotiana fo. pusilla Gola 1923 [nom nud] Typus: SOUTH AMERICA: Tierra del Fuego "Isole del Labirintio" [nom. inval. (art. 32.1(c))(2000, code St-Louis)] leg. G.B. de Gaspari Basionym: Distribution: Golla 1923 823 Nuovo Giorn Bot Ital ser 2, 29: 172 = harpantha Herzog 1942 Typus: Colombia / Cundinamarca. Tequendama, Bergwald, 2200-2400 m 1929, C. Troll 2140 in hb JE 04000975 [holotypus] in hb JE 04000974 isotypus] Basionym: Distribution: Colombia spec nov - syn nov Herzog Bonner Gradstein & Hekking Uribe & Gradstein Hentschel et alii Bernal, Gradstein & Celis = Frullania ambronnii hasseltii 1942 1965 1979 1998 2015 2016 874 655 483 707 4907 4290 Beih Bot Centralbl 61 B: 571-573, fig 7(a-h), 8(a-c) [Columbia] Index Hepat 5: 322 J Hatt Bot Lab 45: 112 [Cat Colombia] Bryophyt Bibl 53: 36 [Catalogue Colombia] Phytotaxa 220 (2): 134 Catálogo de plantas y líquenes de Colombia. Vol. I. [dubious species] Steph. 1916 Sande Lac. 1884 Typus: Indonesia. Sumatra in hb L L0060927 ( sub Fr. gaudichaudii, 1878, Indonesia, Sumatra) [typus] Basionym: Distribution: spec nov Sande Lacoste in Veth 1884 807 -comb nov Verdoorn 1930 293 = Frullania gaudichaudii Midden Sumatra, Reizen en Onderzoekingen der Sumatra Expeditie, 4 (2): 44, pl 7(=Hepat pl 1) 1-7 Ann Bryol Suppl 1: 92, 94-95, fig. 140 [ Sumatra, Philippinen (Mindanao: Davao, Panay, Antique)] fo. hasseltii (Sande Lac.) Verd. 1930 Page 189 hasskarliana fo. integribractea Verd. 1930 Typus: Indonesia. Java Basionym: Distribution: fo nov Verdoorn Bonner Hattori 1930 1965 1975 293 655 547 Ann Bryol Suppl 1: 89 Index Hepat 5: 323 J Hatt Bot Lab 39: 290 [comb nov] = Frullania hasskarliana var. integribractea (Verd.) S.Hatt. 1975 hasskarliana var. edentata [nom herb in hb L L0060929] Typus: Indonesia. Sulawesi, Rantemario "Celebes. Dub div. Eurekang. Rante Mario. Ridge above Tinabang. Fine; large tufts; black, with bluish purple shine. Common; against dead wood" 17/6/1937, Eyma, P.J. 770 in hb L L0060929 [typus] - http://145.18.162.53:81/c8?ent=300017&rec=69089&sct=1 Basionym: Distribution: = hasskarliana var. nonappiculata [nom herb in hb L L0060930] Typus: Indonesia. Java. Malang. "Java: bij Malang." -/10/1939, Groenhart, P. sn in hb L L0060930 [typus] - http://145.18.162.53:81/c8?ent=300017&rec=69090&sct=1 Basionym: Distribution: = hasskarliana var. subacuta [nom herb in hb L L0060931] Typus: Indonesia. Java. Tjibeber . "Java: S of Tjibeber. Estate of Tjiharum. Earthern precipice." 20/2/1952, Borssum Waalkes, J. van 1263 in hb L L0060931 [typus] - http://145.18.162.53:81/c8?ent=300017&rec=69090&sct=1 Basionym: Distribution: = hasskarliana ssp. queenslandica S.Hatt. 1984 Typus: Australia. Queensland, Devil's Thumb, W of Mossman, 1150m 9.10.1982, M.L. Hicks 11258 in hb NICH [holotypus], in hb M.L. HICKS [isotypus] Basionym: Distribution: Australia (QLD) ssp nov - syn nov Hattori Windolf Hattori Carthy McCarthy Mizutani, Hasegawa & Iwatsuki = Frullania johnsonii 1984 1987 1988 2003 2006 2009 682 1025 429 1125 1385 2112 Crypt Bryol Lichen 5(1-2): 180-182, fig 2(a-s) [Austalia, QLD] Austrobaileya 2(4): 390 [syn sub Fr. johnsonii] Nova Hedwigia Beih 90: 151 Flora Australia Suppl Ser 21: 44 [sub Fr johnsonii] Checklist Australian Liverworts [Australia, syn sub Fr johnsonii] Type sepecimens Hatt Bot Lab (NICH) 25: Holotype. Australia, Queensland, Deyil's Thumb. 1150 m. On rock outcrop. 9 Oct. 1982. M. L. Hicks 11258 (NICH 385068). Steph. 1894 Page 190 hasskarlii [auct. non Lindenb.] J.B.Jack & Steph. 1892 {Philippines} Basionym: Distribution: Jack & Stephani = Frullania serrata hastata 1892 1089 Hedwigia 31: 12 [orth var; Philippines] Gottsche 1845 Steph. 1911 Typus: Bolivia: Andes Boliviae or., Bergurwald des Cerro Amborro, 1200 m -.10.1901, Herzog sn hb LEVIER 5859, in hb G [holotypus] Basionym: Distribution: Bolivia spec nov Stephani Bonner Stephani 1911 1965 1985 686 655 928 - syn nov Yuzawa Gradstein, Meneses & Arbe 1991 2003 525 975 = Frullania laxiflora hastatistipula Spec Hepat 4: 672-673 Index Hepat 5: 323 Stephani Icon Hepat Inedit [Chonanthelia] A- 34, microfiche 46 fig 3118-3119 [from type; "Bolivia leg Herzog Levier 5859"] J Hatt Bot Lab 70: 235 [syn sub Fr. laxiflora] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. laxiflora] Spruce 1884 Steph. ex Herzog 1916 Typus: Bolivia. Abra de San Mateo, 3000m 1911, T. Herzog 3724 in hb G sn [holotypus] Basionym: Distribution: Bolivia spec nov Stephani Stephani Bonner Stephani 1916 1924 1965 1985 - syn nov Uribe & Gradstein Uribe 2003 2008 = Frullania brasiliensis hattoriana 222 687 655 928 Biblioth Bot 87: 245-246, fig 198 [Bolivia] Spec Hepat 6: 549-550 [as spec nov] Index Hepat 5: 323 Stephani Icon Hepat Inedit [Meteriopsis] E-23, microfiche 55 fig 3798 ["Bolivia, später auch 4210 - Dr Herzog 3724 (1913)"] 1071 Crypt Bryol 24 (3): 204 [syn sub Fr brasiliensis] 1513 Caldasia 30(1): 90, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Raddi 1822 J.D.Godfrey & G.Godfrey 1980 Typus: Canada. British Columbia, Vancouver, Mt. Seymour Prov. Pk. sd, Godfrey 1485 in hb GODFREY & GODFREY [holotypus], in hb UBC, hb NICH [isotypus] Basionym: Distribution: Canada ( BC ) spec nov -syn nov Godfrey & Godfrey Hong Qian & Klinka Mamontov et al. Atwwod & Mamontov = Frullania chilcootiensis 1980 1989 1998 2020 2020 310 689 2262 5641 5816 J Hatt Bot Lab 48: 321-327, fig 1(1-7), 2(1-18) Bryologist 92 (3): 363-367 [in key as #7, but text of #7 missing] Plants of British Columbia: 339 [Canada] Nova Hedwigia, Beiheft 150: 223-225 , fig 8(a-n) Botanica Pacifica 9(2):191-195, fig 1g[ syn sub Fr chilcootiensis] Steph. 1887 Page 191 hautuscana Steph. by Stotler 1969 [orth var] Basionym: Distribution: Stotler Stotler = Frullania huatuscana 1969 1970 Steph. 1911 Frullania intumescens hawaica 4761 Nova Hedwigia 18: 444 [syn sub Fr. intumescens] 194 The genus Frullania, separata: 444 [syn sub Fr. intumescens] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. ex Steph. 1985 [pro syn] Typus: Hawai leg Lauterbach 204a Basionym: Distribution: Stephani = Frullania sandvicensis hawaiiensis 1985 928 Stephani Icon Hepat Inedit [Galeiloba] C-178, microfiche 52 fig 3580 ["olim Frull hawaica St ms, Hawai, leg Lauterbach 204a"] Ångstr. 1872 Miller 1953 Typus: USA. Hawaii. Oahu, Honolulu, East Makaleha Valley, 1800ft=549m [holotype] - USA. Hawaii, Maui Isl, above Lahaina, 1800ft [paratype1] - Oahu, Pupukea-Kahuku, Summit Trail [paratype2] 15.9.1950, Miller 1565 [holotype] - 27.11.1950 Miller 1885 [paratype1] 2.8.1951 Lohman 2002 [paratype2] in hb H.A. MILLER [isotypus] in hb BISH 500125 (15.09.1950, Miller 1565) [holotypus] in hb BISH 99534 (02.08.1951, Miller 2233a) [paratypus], BISH 706700 (02.08.1951, M.L. Lohmann 2002) [paratypus] Basionym: Distribution: Hawaii (Oahu, Molokai, Maui) - syn nov Miller Miller Bonner Miller, Whittier & Whittier Inoue O'Connor, Gowing, Medler et alii N.N. [National Science Museum Tokyo] So & Wang Staples & Imada = Frullania crenatiloba hebridensis 1953 1963 1965 1983 1987 1998 2000 349 834 655 609 1155 1274 995 Bryologist 56: 41-44, fig a-n Ark Bot, ser 2, 5: 519 [Hawaii] Index Hepat 5: 323 Prod Flora Hepat Polynesiae: 128 Index type specimens in Nat Sci Mus, Tokyo: 13 Type specimen catalog hb BISH http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 2006 2006 1466 J H B L 100: 422 [syn nov sub Fr crenatiloba] 1716 Trop bryol 28: 27 [Hawaii] Steph. 1924 Steph. ex Paris 1906 [nom nud] Basionym: Distribution: Paris = Frullania hebridensis 1906 676 Rev Bryol 33 (2): 28 [New Caledonia] Steph. 1911 Page 192 hedbergii Vanden Berghen 1953 Typus: Kenya, Aberdare range, near w part of Nyeri track 3070m 18.7.1948, Hedberg 1652a in hb S-PA [holotypus] Basionym: Distribution: Burundi, Kenya spec nov - syn nov Vanden Berghen Arnell Arnell Vanden Berghen Bonner Vanden Berghen Vanden Berghen Yuzawa 1953 1956 1956 1961 1965 1976 1976 1991 Wigginton & Grolle 1996 Wigginton 2002 Wigginton 2004 = Frullania albertii Svensk Bot Tidskr 47 (2): 272-274, fig. 3(a-e) Svensk Bot Tidskr 50 (3): 521 [Kenya] ANGABE FALSCH !!!! 836 Ark Botanik 3 (16): 531 [East African mountains] 1507 Bull Soc Roy Bot Belg 93 (1-2): 70 [Urundi, Kenya] 655 Index Hepat 5: 324 [Kenya] 185 Bull Jard Bot Nat Belg 46 (1/2): 211-213, fig 51 [Kenya] 850 Distributiones plantarum africanarum 11: 372 [map of Africa] 525 J Hatt Bot Lab 70: 205 [syn sub Fr. albertii, only according to figure of Vanden Berghen] 696 Bryophyt Bibl 50: 81 [Catalogue ... of Sub-Saharan Africa] [syn sub Fr. albertii] 1012 Trop Bryol Res Rep 2: 28 [syn sub Fr. albertii] [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [syn sub Fr. albertii] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Steph. ex Herzog 1916 Frullania standaertii helleri 446 Steph. 1910 Steph. 1911 [hom illeg] [non Steph. 1897, nec Steph. 1910] Typus: Hawaii Basionym: Distribution: Stephani = Frullania helleri 1911 686 Spec Hepat 4: 554 [spec nov = nom superfl] Steph. 1897 [non Steph. 1910, nec Steph. 1911] Frullania meyeniana Lindenb. 1845 Page 193 helleri Steph. 1897 [non Steph. 1910, nec Steph. 1911] Typus: Hawaii, Oahu. Nuuana 9.4.1895 [or 23.3.1895], A.A. Heller 2111 in hb G [holotypus] in hb MICH MICH514765 (09.04.1895, A.A. Heller 2111, USA, Hawaii, Honululu: Nuuanu, on trees) [isotypus ?], in hb BISH BISH 99533 (23.03.1895, Heller 2111) [isotypus] in hb E E00002541 [isotypus] Basionym: Distribution: Hawaii spec nov Stephani 1897 - syn nov Evans Stephani Hattori Miller Kamimura Miller Bonner Hattori Miller, Whittier & Whittier Stephani 1900 1911 1944 1953 1961 1963 1965 1972 1983 1985 NN 1996 Fogel 1997 O'Connor, Gowing, Medler et alii 1998 Yamada & Iwatsuki 2006 So & Wang 2006 = Frullania meyeniana helleri 214 Bull Herb Boissier 5: 842[:260] & 845-846 [:263-264] [Hawaii] "Heller 2111" 236 Trans Connet Acad Arts Sci 10 (1): 402 [syn nov sub Fr. meyeniana] 686 Spec Hepat 4: 554 [spec nov = nom superfl] 943 Bull Tokyo Sci Mus 11: 146 [Japan, key to Japanese species] 349 Bryologist 56: 46 [sub Fr meyeniana] 199 J Hatt Bot Lab 24: 73 [sub F meyeniana] 834 Ark Bot, ser 2, 5: 519 [syn sub Fr. meyeniana] 655 Index Hepat 5: 324 538 J Hatt Bot Lab 36: 115 609 Prod Flora Hepat Polynesiae: 128 928 Stephani Icon Hepat Inedit [Thyopsiella] F-170, microfiche 59 fig 4111 ["Sandwich Insel, A A Heller, Herb Underwood"] 632 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 17 of 35 [in hb US] 629 http://141.211.110.91/livrqury.htm [16.11.1997] - Genus=Frullania, printed 5.1.1998 11:11:38: page 1-2 of 3 1274 Type specimen catalog hb BISH 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] 1466 J H B L 100: 424 [sub Fr meyeniana] Lindenb. 1845 Steph. 1910 [nom illeg (Art 53.1)] [non Steph. 1897, nec Steph. 1911] Typus: Hawaii. Kilauea volcano, 1400m 1905, K. Rechinger 2563 [? & 2520, 3131] in hb G 20583 Basionym: Distribution: Hawaii spec nov spec nov Stephani Verdoorn Kamimura Bonner So & Wang = Frullania meyeniana herminieri 1910 1930 1961 1965 2006 217 551 199 655 1466 Denkschr K Akad Wiss, Math Nat Kl, Wien 85: 193 [Hawaii] Nederl Kruidk Arch Jrg 1930 (2): J Hatt Bot Lab 24: 73 [sub F meyeniana] Index Hepat 5: 324 J H B L 100: 424 [sub Fr meyeniana] Lindenb. 1845 Steph. ex Bonner 1965 [nom herb] Typus: Guadeloupe. inter fr. atratam sd, L´Herminier hb STEPHANI 115, in hb G sub Fr. gibbosa Basionym: Distribution: Guadeloupe Bonner Stephani = Frullania gibbosa 1965 1985 655 928 Index Hepat 5: 324 Stephani Icon Hepat Inedit [Chonanthelia] A- 69ab, microfiche 47 fig 3169 ["olim Frull Herminieri Steph Guadeloupe 609 e Mexico 943"] Nees 1840 Page 194 herzogiana Steph. ex Herzog 1916 [hom illeg] [non Steph. 1911] Typus: Bolivia. Tablas, 3400m 1911, Herzog 2829 in hb G Basionym: Distribution: Bolivia spec nov - nom nov - syn nov Stephani Stephani Herzog Bonner Stephani 1916 1924 1942 1965 1985 Gradstein, Meneses & Arbe Uribe 2003 2008 = Frullania controversa Biblioth Bot 87: 246-247, fig 199 [Bolivia] Spec Hepat 6: 549 [nom nov as Fr. controversa] Beih Bot Centralbl 61 B: 575 Index Hepat 5: 325 Stephani Icon Hepat Inedit [Meteriopsis] E-43, microfiche 56 fig 3832 ["Bolivia No 2829 (1913) Herzog leg"] - 3833 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr xxx] 1513 Caldasia 30(1): 66 [syn sub Fr convoluta] [Monography of subgenus Meteoriopsis] Beauverd ex Steph. 1924 Frullania convoluta herzogiana 222 687 874 655 928 Lindenb. & Hampe 1851 Steph. 1911 [non Steph. 1916] Typus: Bolivia. Velasco Prov., Urwald des Rio Blanca, 120m -.8.1907, Th. Herzog hb LEVIER 5892, in hb G Basionym: Distribution: Bolivia spec nov Stephani Bonner Stephani 1911 1965 1985 - syn nov Uribe 2008 = Frullania convoluta herzogii 686 655 928 Spec Hepat 4: 673 Index Hepat 5: 325 Stephani Icon Hepat Inedit [Chonanthelia] A- 35, microfiche 46 fig 3120 [from type; "Bolivia Velasco Rio Blanco Herzog leg 1907 Levier 5892"] 1513 Caldasia 30(1): 66 [syn sub Fr convoluta] [Monography of subgenus Meteoriopsis] Lindenb. & Hampe 1851 S.Hatt. 1955 Typus: Japan. Honsyu, Jwate Pref., Mt. Hayachinae, ca. 1200m 4.9.1953, Shimizu in hb NICH [holotypus] Basionym: Distribution: Japan spec nov - nom nov Hattori Hattori Hattori & Noguchi Kamimura Kamimura 1955 1957 1960 1961 1962 Mizutani Kamimura Bonner Stotler & Crandall-Stotler 1962 1962 1965 1987 Piippo Grolle & Meister 1990 2004 Yamada & Iwatsuki 2006 Mizutani, Hasegawa & Iwatsuki 2009 = Neohattoria herzogii 108 858 965 199 199 Feddes Repert 58: 53-54, fig 1-3 [Japan] J Hatt Bot Lab 18: 117 Index spec Typ herb Japon, Bryophyta: 17 [as Hattoria herzogii] J Hatt Bot Lab 24: 94, fig 33(1-21), map 10 [comb nov as Hattoria herzogii] J Hatt Bot Lab 24: 141, fig III(22-29) [as Hattoria herzogii] J Jap Bot 37: 26 [as Neohattoria herzogii] 655 Index Hepat 5: 325 767 Memoirs of The New York Botanical Garden 45: 535-543 [as subgen Dentatiloba] 521 J Hatt Bot Lab 68: 41 [not in China; annotated catalogue of China] 1321 Liverwort Baltic & Bitterfeld amber, Jena: 22-23 [subgen Dentatiloba with second species] 1470 J Hatt Bot Lab 99: 30 [; as Neohattoria herzogii - Catalog of Japan] 2112 Type sepecimens Hatt Bot Lab (NICH) 26: Holotype. Japan, Honshu, Iwate-ken, Mt. Hayachine. 1200 m. On bark of Thujopsis dolabrata. 4 Sept. 1954. D. Shimizu sn (NICH 55212). Kamim. 1962 Page 195 heteroides Mitten ex Verdon [nom. inval.] Typus: Dent de Saint-Vincent, Nouvelle-Calédonie, 1400m Le Rat 1909-07 https://www.gbif.org/occurrence/1211206741 Université de Montréal Biodiversity Basionym: Distribution: = heteromorpha fo. mamillosa Verd. 1930 Typus: PNG. East Sepik, Hauptlager Malu, Alluvialwald, Sepik Exped. 28.3.1912, Ledermann 6801 in hb FH Basionym: Distribution: fo nov -syn nov Verdoorn Verdoorn Bonner Hattori Grolle & Piippo = Frullania heteromorpha heterostipa 1930 1930 1965 1980 1984 293 549 655 490 143 Ann Bryol Suppl 1: 170-171 Nova Guinea 14 (4): 545 Index Hepat 5: 325 J Hatt Bot Lab 47: 176 [syn sub Fr. heteromorpha] Acta Bot Fennica 125: 20 [Cat. West Melanesia] Schiffn. 1889 Steph. ex Bonner 1965 [nom herb] Typus: Brazil. Moraima sd, Ule 645 in hb G in hb HBG Basionym: Distribution: Brazil Bonner Stephani 1965 1985 Feuerer 2008 655 928 Index Hepat 5: 326 Stephani Icon Hepat Inedit [Meteriopsis] E-24, microfiche 55 fig 3799 ["Brasilia, Roraime, Ule No 645"] 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm = hewsoniae Kamim. [nom herb in hb L L0060932] Typus: Papua New Guinea. New Guinea. Keglsugl "Loc.: Keglsugl. State: E.H.D. Granite rock." in hb LL0060932 [typus] - http://145.18.162.53:81/c8?ent=300017&rec=69092&sct=1 Basionym: Distribution: = Page 196 5/8/1965, Hewson, H.J. 111 hians (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Typus: in hb GLM (hb GOTTSCHE) 06018 Basionym: Distribution: Panama Lehmann & Lindenberg in G., L. & N. Hampe Mitten in Seemann Jack & Stephani Schiffner 1845 2850 Syn Hepat: 414 1851 1857 1892 1895 281 1494 1089 1015 Dusen Schiffner in Engler & Prantl Herzog Yano 1905 1905 1953 1984 710 809 640 501 = Frullania hians d'Orbigny 1839 Linnaea 24 (3): 303 [Costa Rica] Bot Voyage HMS Herald: 246 [Panama] Hedwigia 31: 13 [Peru, Neu-Granada] Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 132 Arch Mus Nac Rio de Janeiro 13: 116 & 119 [Brasil] Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 132 Rev Bryol Lichen 20: 128 [Costa Rica] J Hatt Bot Lab 56: 505 [checklist Brazil] (Lehm. & Lindenb. in Lehm.) Mont. & Nees in Mont. in Alcide Frullania obscura (Sw.) Mont. 1840 Page 197 hians (Lehm. & Lindenb. in Lehm.) Mont. & Nees in Mont. in Alcide d'Orbigny 1839 Typus: Mexico, Xalapa Schiede s.n. (G, S!, STR, W) Basionym: Jungermannia hians Lehm. & Lindenb. 1832 Distribution: Mexico, Nicaragua, Ecuador, Peru, Galapagos Isls [Ecuador], Bolivia, Brazil, Argentina comb nov syn syn syn - syn nov Montagne & Nees in Montagne in Alcide d'Orbigny Lehmann & Lindenberg in G., L. & N. Hampe Montagne Sande Lacoste Gottsche Gottsche Gottsche Ångström Spruce Dusen Dusen Stephani Evans Stephani Bryan Pagan Herrera Demaret & Vanden Berghen Kühnemann Clark & Svihla Herzog Arnell Bonner Yana Menzel Stephani 1839 257 1845 2850 Syn Hepat: 414 (No 7) 1851 1855 1857 1863 1864 1867 1876 1884 1905 1905 1910 1914 1916 1929 1939 1941 1948 1949 1950 1952 1962 1965 1984 1984 1985 281 475 430 103 478 104 579 243 710 710 686 235 222 175 931 2789 59 657 332 444 449 655 501 142 928 Schuster Fulford & Sharp Yuzawa Buck Grolle Wigginton 1985 1990 1991 1994 1995 2002 441 105 525 28 250 1012 Gradstein, Meneses & Arbe 2003 975 Wigginton Wigginton Gradstein 2004 2004 2012 1312 1172 3341 = Frullania obscura d'Orbigny, Voyage l'Amerique Merid 7 (2): 69 Linnaea 24 (3): 303 [Nicaragua] Ann Sci Nat Bot, ser 4, 3: 319 Verh Konikl Akad Wetenschappen 5: 77-78 (No 208) Separata of Mex. Leverm.: 237 Ann Sci Nat Bot, ser 5, 1: 166-167 (No 4) K Danske Vid Selks Skrifter, ser 5, 6: 333 [Mexico] Öfversigt K Vetensk Akad Förh 33 (7): 88 [Brazil] Trans Proc Bot Soc Edinburgh 15: 12 & 19-20 (No 7) Arch Mus Nac Rio de Janeiro 13: 116 [Brasil] Arch Mus Nac Rio de Janeiro 13: 119 [Brasil] Spec Hepat 4: 333-334 Trans Connet Acad Arts Sci 18: 333-338, fig. 10(a-c) [Peru] Biblioth Bot 87: 239 [Bolivia] Bot Gazette 88(3): 338-339 [Peru] Bryologist 42: 80 [Puerto Rico] Sinopsis de la Flora del Cuzco: 70 [Peru, Cuzco] Bull Jard Bot Etat Bruxelles 19 (1): 77 [syn sub Fr arecae] Lilloa 19: 354 [Argentina: Tucuman] Bryologist 53: Svensk Bot Tidskr 46 (1): 83 [Ecuador] Svensk Bot Tidskr 56 (2): 341 [Ecuador] Index Hepat 5: 326 J Hatt Bot Lab 56: 505 [checklist Brazil] Willdenowia 14: 509 [syn sub Fr. arecae] Stephani Icon Hepat Inedit [Chonanthelia] A- 36, microfiche 46 fig 3121 [Brasilia] Phytologia 57 (5): 373 [sect Pluriplicatae - type species for nov sect] Mem NY Bot Gard 63: 43 [Mexico] [syn sub Fr. arecae ?] Bryologist 97 (4): 422 [Vainio's specimen from Brazil at TUR] Bryophyt Bibl 48: 63 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. arecae; according Yuzawa 1991 ?}] Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 224 [syn sub Fr arecae, West Africa] Cryptogamie, Bryologie 33(3): 257-262 [type, syn sub Fr obscura] (Sw.) Mont. 1840 hians var. (delta) Lehm. & Lindenb. 1847 Basionym: Distribution: Lehmann & Lindenberg in G., L. 1847 & N. = Frullania arecae 2848 Syn Hepat: 772 (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura (Sw.) Mont. 1840 Page 198 hians var. (gamma) Lehm. & Lindenb. 1847 Basionym: Distribution: Lehmann & Lindenberg in G., L. 1847 & N. Gottsche 1863 Gottsche 1867 = Frullania arecae 2848 Syn Hepat: 772 103 104 Separata of Mex. Leverm.: 237 K Danske Vid Selks Skrifter, ser 5, 6: 333 [Mexico] (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura (Sw.) Mont. 1840 hians var. conferta Lehm. & Lindenb. 1847 Typus: Mexico. Zacuapan, 2500-3000ft Leibold Basionym: Distribution: Mexico var nov - syn nov Lehmann & Lindenberg in G., L. & N. Gottsche Gottsche Bonner Fulford & Sharp Yuzawa = Frullania arecae 1847 2848 Syn Hepat: 772 1863 1867 1965 1990 1991 103 104 655 105 525 Separata of Mex. Leverm.: 237 K Danske Vid Selks Skrifter, ser 5, 6: 333 [Mexico] Index Hepat 5: 326 Mem NY Bot Gard 63: 43 [Mexico] J Hatt Bot Lab 70: 264 [syn sub Fr. arecae ] (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura (Sw.) Mont. 1840 hians var. crispa Spruce 1884 Basionym: Distribution: Ecuador var nov Spruce Bonner Yuzawa spec nov 1884 1965 1991 = Frullania arecae 243 655 525 Trans Proc Bot Soc Edinburgh 15: 20 Index Hepat 5: 326 J Hatt Bot Lab 70: 264 [syn sub Fr. arecae] (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura (Sw.) Mont. 1840 hians var. javanica Gottsche ex Lehm. & Lindenb. 1845 Typus: Indonesia. Java sd, Hasskarl in hb NEES Basionym: Distribution: Indonesia (Java) spec nov - syn nov Lehmann & Lindenberg in G., L. & N. Sande Lacoste Sande Lacoste in Junghuhn Reichardt Verdoorn Demaret & Vanden Berghen Bonner = Frullania arecae 1845 2850 Syn Hepat: 414 (No 7) 1857 1857 1870 1930 1948 1965 430 1490 884 293 59 655 Verh Konikl Akad Wetenschappen 5: 78 (No 205 b) Plantae Junghuhnianae, Hepaticae: 536 Reise österr Fregatte Novara, Bot Theil, Bd 1: 154 [Java] Ann Bryol Suppl 1: 70-72 [syn sub Fr. wallichiana] Bull Jard Bot Etat Bruxelles 19 (1): 77 [syn sub Fr. arecae] Index Hepat 5: 326 (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura (Sw.) Mont. 1840 Page 199 hiantis Lehm. & Lindenb. in Schiffn. & S.W.Arnell 1964 Basionym: Distribution: Schiffner & Arnell = Frullania arecae 198 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 84 [as syn of Fr. arecae] [Brazil] (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura hieronymi 1964 (Sw.) Mont. 1840 Steph. ex Bonner 1965 [nom herb] Typus: India orientalis sd, Pfleiderer K 93 in hb G 14417 Basionym: Distribution: India nom herb - syn nov Bonner Hattori 1965 1972 655 538 Hattori in Ohashi Stephani 1975 1985 187 928 Index Hepat 5: 326-327 J Hatt Bot Lab 36: 133-134, fig 18(a-g) [syn sub Fr. neurota var pfleidereri] Univ Mus Univ Tokyo Bull 8: 234 [sub Fr. neurota] Stephani Icon Hepat Inedit [Chonanthelia] A- 88, microfiche 47 fig 3194 [from type; "India orientalis Pfleiderer K 93"] = Frullania neurota var. pfleiderei S.Hatt. 1972 Frullania neurota hilariana Taylor 1846 Mont. 1838 Typus: Brazil. "prope Uba, Provinciae Rio de Janeiro" sd, Aug. d. St. Hilaire in hb MONTAGNE 162, in hb G Basionym: Distribution: Costa Rica, Brazil spec nov - syn nov Montagne Montagne Lehmann & Lindenberg in G., L. & N. Montagne Stephani Bonner Yano Stephani 1838 1839 1845 464 Ann Sci Nat Bot, ser 2, 9: 45-46 465 Ann Sci Nat Bot, ser 2, 12: 51 [Brasil, spec nov] 2850 Syn Hepat: 421 1856 1910 1965 1984 1985 1912 686 655 501 928 Morales Z. Gradstein & Costa Holz 1991 2003 2004 = Frullania glomerata Mont 1842] Sylloge generum specierumque cryptogamaram: 87 Spec Hepat 4: 384 Index Hepat 5: 327 J Hatt Bot Lab 56: 505 [checklist Brazil] Stephani Icon Hepat Inedit [Galeiloba] C- 61abcd, microfiche 50 fig 3417 ["(olim Frull hilariana)"] 307 Trop Bryol 4: 38 [Costa Rica] 984 Mem New York Bot Gard 87: 87 [Brazil] [syn sub Fr glomerata] 1184 [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Page 200 hildebrandtii Steph. 1910 Typus: Madagaskar: Imerina sd, Hildbrandt sn ex hb B, in hb G [holotypus], Basionym: Distribution: spec nov - syn nov Stephani Stephani in Brunnthaler Sim Vanden Berghen Arnell Bonner Vanden Berghen Stephani 1910 1913 1926 1953 1963 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania trinervis hinoi 686 219 252 446 822 655 185 928 Spec Hepat 4: 324-325 [Madagascar] Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 729 [Kapland] Trans Roy Soc S Africa 15: 41 [does not differ from trinervis] Svensk Bot Tidskr 47 (2): 272 [ev. syn sub trinervis] Hepat S Africa: 141 [syn sub Fr. trinervis] Index Hepat 5: 327 Bull Jard Bot Nat Belg 46 (1/2): 199 Stephani Icon Hepat Inedit [Chonanthelia] A- 13, microfiche 46 fig 3083 ["Madagascar Bongen legit"] 250 Bryophyt Bibl 48: 69 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 227 [syn sub Fr trinervis, West Africa] (Lehm.) Drège 1843 Kamim. 1982 Typus: Japan. Kagoshima Pref, Isl Amami-oshima, Sumiyo-mura 20.6.1961, I. Hino 97 in hb NICH 67741 Basionym: Distribution: Japan - syn nov Kamimura Furuki & Mizutani Yuzawa 1982 1994 2001 Yamada & Iwatsuki 2006 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania kagoshimensis hirtelliflora 420 Misc Bryol Lichen 9 (4): 89-90, fig 2 1331 Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] 1660 Natural Environmental Sci Res 14: 19, fig 17(a-l) [Japan, syn nov sub Fr kagoshimensis] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] 2112 Type sepecimens Hatt Bot Lab (NICH) 26: Holotype. Japan, Kyushu, Kagoshima-ken, Amami Oshima I., Sumiyo-mura. 30 June 1961. 1. Hino 97 (NICH 67741). Steph. 1910 Gottsche ex Steph. 1910 Typus: Brazil. Apiahy sd, Puiggari 105 in hb G 1612 Basionym: Distribution: Brazil spec nov - syn nov Gottsche Gottsche in Stephani Schiffner & Arnell 0 1910 1964 Bonner Yano Stephani 1965 1984 1985 Reiner Yano Gradstein & Costa Yano 1988 1999 2003 2008 = Frullania glomerata Mont 1842] Icon inedit Spec Hepat 4: 390-391 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 91, pl 13 fig 150(h-g) [Brazil] [may be fo syn sub Fr. julaceaa] 655 Index Hepat 5: 327 501 J Hatt Bot Lab 56: 505 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Galeiloba] C- 62, microfiche 50 fig 3423 [from type; "Apiahy ... "] 1043 Bol Soc Argentina Bot 25 (3-4): 319 [syn sub Fr. glomerata] 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] 984 Mem New York Bot Gard 87: 87 [Brazil] [sub Fr glomerata] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr glomerata; Brazil; catalog] 686 198 (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Page 201 hirtiflora Spruce 1884 Typus: Ecuador. "Andes Quitenses", Banos, Mt. Tunguragua, 1800m sd, Spruce in hb G Basionym: Distribution: Ecuador spec nov Spruce Stephani Bonner Stephani 1884 1910 1965 1985 - syn nov Gradstein 2021 = Frullania glomerata Mont 1842] hirtiflora 243 686 655 928 Trans Proc Bot Soc Edinburgh 15: 33 & 35 Spec Hepat 4: 390 Index Hepat 5: 327-328 Stephani Icon Hepat Inedit [Galeiloba] C- 63, microfiche 50 fig 3424 [from type; "Andes quit, Banos, Spruce"] 5642 Mem NY Bot Gard 121: 367 [syn sub Fr glomerata] (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Gottsche ex Bonner 1965 [nom herb] Typus: New Zealand. sd, Knight 602 in hb G (ex hb GOTTSCHE) [sub Fr. pycnantha] Basionym: Distribution: Bonner = Frullania pycnantha hispanica 1965 655 Index Hepat 5: 328 (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Hampe ex Schiffn. 1936 [pro syn] [hb LINDENBERG No 7361] Basionym: Distribution: Schiffner 1936 277 Österr Bot Zeitschr 85 (1): 22 [hispanica HAMPE - Hist austr - hb LINDENBERG Nr 7361] = Frullania nervosa var. hampei Schiffn. 1936 Frullania tamarisci var. nervosa (Mont.) S.W.Arnell 1961 Frullania tamarisci hispanica (L.) Dumort. 1835 Nees 1838 [non Hampe ex Lindenb. in G., L. & N. 1845] Typus: Spain. Asturiae monte, Mt. Gillon 30.7.18.., Durieu ex hb NOTARIS, ex hb MEISSNER in hb G (hb LINDENBERG 7362), typus in hb S (B24079, 24080) Basionym: Distribution: spec nov - syn nov -syn nov Nees Montagne in Barker-Webb & Berthelot Nees in G., L. & N. Stephani Müller in Rabenhorst Schiffner Müller in Rabenhorst Bonner Pocs Stephani 1838 1844 261 910 Naturg Europ Leberm 3: 236-239 Hist Nat Iles Canariensis 3,2,4: 54-55 1845 1911 1915 1936 1958 1965 1976 1985 2850 686 1064 277 1028 655 2686 928 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 2002 782 Syn Hepat: 439 [stat nov] Spec Hepat 4: 560-561 Krypt Flora (2ed) 6, 2: 610, 612 [syn sub Fr tamarisci var sardoa] Österr Bot Zeitschr 85 (1): 26-27 Krypt Flora (3ed) 6, 2: 1234 [syn sub Fr tamarisci] Index Hepat 5: 328 Fol. Hist. nat. Mus. Matr. 4: 19 [types in hb EGER] Stephani Icon Hepat Inedit [Thyopsiella] F-194, microfiche 60 fig 4153 ["Asturia, Durieu legit 1845"] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] = Frullania tamarisci var. sardoa (De Not.) De Not. 1865 Page 202 hispanica Hampe ex Lindenb. 1845 [pro syn] Typus: Spain. in hb W (hb LINDENBERG 7361) [= paratypus to Fr. polysticta] Basionym: Distribution: Lindenberg in G., L. & N. Schiffner 1845 1936 Müller 1951 Grolle 1970 Bisang, Schumacker, Sérgio & Grolle 1989 = Frullania polysticta hispanica 2850 Syn Hepat: 440 [syn of Fr. polysticta] 277 Österr Bot Zeitschr 85 (1): 21 [nom nov, as Fr. polysticta var hampei, = syn of Fr. bryhnii] 641 Rev Bryol Lichen 20: 134-135 [in hb NEES as Fr. nervosa var polysticta] 615 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 309 [syn of Fr. polysticta] 276 Giorn Bot Ital 122: 264 [syn sub Fr. teneriffae] Lindenb. 1845 Hampe ex MÜLLER 1951 [pro syn] [non Nees 1838] Basionym: Distribution: Schiffner 1936 277 Müller 1951 641 = Frullania nervosa Mont. in Berthold & Webb 1844 [hom illeg] [non Mont. 1840] Frullania polysticta hispanicam Österr Bot Zeitschr 85 (1): 19-21 [hispanica HAMPE - Hist austr - hb LINDENBERG Nr 7361; nervosa syn sub Fr. ploysticta] vide Müller Rev Bryol Lichen 20: 134 Lindenb. 1845 Nees Basionym: Distribution: syn nov Beauverd in Stephani = Frullania decipiens 909 Spec Hepat 6: 570 [P. 560, No 173, ad F. hispanicam Nees, adde : « Syn. F. tamarisci Dum. var. sardoa] Beauverd in Steph. 1924 [hom illeg] [non Herzog 1923] Frullania ecuadorensis Frullania kunzei hispida 1924 Steph. 1911 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Gottsche ex Bonner 1965 [nom herb] [non Steph. 1911] Typus: New Zealand. sd, Knight 596 in hb G (ex hb GOTTSCHE) [sub Fr. pycnantha] Basionym: Distribution: New Zealand Bonner = Frullania pycnantha 1965 655 Index Hepat 5: 328 (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Page 203 hispida Steph. 1911 [non Gottsche ex Bonner 1965] Typus: Madagascar sd, Forsyth Major 485 pp in hb G ; Stephani, Icones Hepat Inedit 1703 Basionym: Distribution: Madagascar spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania angulata hoehneliana 686 655 185 928 Spec Hepat 4: 685 Index Hepat 5: 328 Bull Jard Bot Nat Belg 46 (1/2): 71 [syn sub Fr. angulata] Stephani Icon Hepat Inedit [Thyopsiella] F- 5, microfiche 56 fig 3874 ["Madagascar, Dr Forsyth Major 1896"] 250 Bryophyt Bibl 48: 62 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Mitt. 1863 Steph. 1894 Typus: Kenya. Leikipia, "in monte Kenya" 1887, von Hoehnel, Teleki Exped. in hb G Basionym: Distribution: Ethiopia spec nov - syn nov Stephani Stephani Stephani in Engler Stephani Gola Gola Cufodontis Vanden Berghen Arnell Bonner Vanden Berghen Stephani 1894 1895 1895 1910 1914 1916 1952 1953 1963 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania caffraria holokelensis 228 212 1481 686 786 3469 2562 446 822 655 185 928 Hedwigia 33: 145 Bot Jahrb 20 (3): 316 [as spec nov] [Massaihochland] Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [Massaihochland (L.)] Spec Hepat 4: 382 ["höhneliana"] Annali di Botanica 13: 72 [Ethiopia} Mem Reale Accad Sci Torino, ser 2. 65: 9 [Kenya] Phyton 4 (1-3). 76 [Ethiopia] Svensk Bot Tidskr 47 (2): 271-272 [syn nov sub Fr. caffraria] Hepat S Africa: 148 [syn nov sub Fr. caffraria] Index Hepat 5: 328-329 Bull Jard Bot Nat Belg 46 (1/2): 178 & 184 Stephani Icon Hepat Inedit [Galeiloba] C- 20ab, microfiche 50 fig 3371 [from type; "Leikipia (Kenia), v Höhnel, 1887"] - 3372 ["olim Frull abyssinica, Schoah prope Aukabar, Herb Levier inter No 58"] 250 Bryophyt Bibl 48: 64 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 229 [syn sub Fr caffraria, West Africa] Steph. 1894 Steph. ex Bonner 1965 [nom herb] Typus: Hawaii. Kauai, Holokele 15.3.1910, Faurie 126 in hb G Basionym: Distribution: Hawaii Bonner Miller, Whittier & Whittier Stephani 1965 1983 1985 So & Wang 2006 = Frullania sandvicensis 655 609 928 Index Hepat 5: 329 [nom hb ?] Prod Flora Hepat Polynesiae: 128 Stephani Icon Hepat Inedit [Galeiloba] C-143, microfiche 52 fig 3537 [from type; "Hawai, Kanai, Faurie 1910 - 126"] 1466 J H B L 100: 428 [syn sub Fr sandvicensis] [nom herb, nom illeg (Art 53.1)] Ångstr. 1872 Page 204 hololekensis Steph. in Verd. 1930 [orth var] [pro syn] Basionym: Distribution: - pro syn Verdoorn Verdoorn Miller 1930 1930 1953 = Frullania holokelensis Ann Bryol Suppl 1: 66 ["STEPH in sched", as syn of Fr. neurota ] Ann Bryol Suppl 1: 66 Bryologist 56: 45 [sub Fr neurota] Steph. ex Bonner 1965 [nom herb] Frullania sandvicensis holstii 293 293 349 Ångstr. 1872 Steph. 1894 Typus: E Africa: Usambara, Hochwald Tewe Bangarra -.7.1893, Holst 3289 [& 9042] ex hb B, in hb G Basionym: Distribution: E Africa spec nov - syn nov Stephani Stephani Stephani in Engler 1894 1895 1895 Gola in Savoia Stephani Gola Bonner Vanden Berghen Stephani 1909 1911 1916 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania angulata hondurae 228 Hedwigia 33: 158 212 Bot Jahrb 20 (3): 315 [sp. nov.] [Usambara] 1481 Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [Usambara 1600m Gebirgsbuschwald] 832 Il Ruwenzori, parte scientifica 1: 529 686 Spec Hepat 4: 482 3469 Mem Reale Accad Sci Torino, ser 2. 65: 9 [Kenya] 655 Index Hepat 5: 329 185 Bull Jard Bot Nat Belg 46 (1/2): 70 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 6ab, microfiche 56 fig 3875 ["olim laxepinnata, Usambara, Holst 9042"] - 3876 - 3877 ["Usambara, Holst 3289"] - 3878 ["substricta, Africa, Usambara, Brunthaler 1909"] - 3879 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Mitt. 1863 Herzog & Clark in Clark & Svihla 1953 Typus: Costa Rica. Prov. San José, vicinity of La Hondura, 1300-1700m 2-4.3.1924, P. C. Standley 36559 in hb US Basionym: Distribution: Costa Rica spec nov - syn nov - syn nov Herzog & Clark in Clark & Svihla Bonner Stotler Stotler NN = Frullania brasiliensis 1953 1965 1969 1970 1996 360 655 4761 194 632 Bryologist 56: 289-292, fig. 1-20 Index Hepat 5: 329 Nova Hedwigia 18: 436 The genus Frullania, separata: 436 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 18 of 35 [in hb US] Raddi 1822 Page 205 hongkongensis Verd. 1932 Basionym: Distribution: spec nov syn nov - syn nov Verdoorn Verdoorn Hattori & Lin Piippo 1932 1934 1985 1990 554 240 505 521 Nederl Kruidk Arch Jrg 1932 (2): 494-495 Ann Bryol Suppl 4: 195 [syn sub Fr balansae] J Hatt Bot Lab 59. 161-162 [sub Fr J Hatt Bot Lab 68: 42 [annotated catalogue of China] = Frullania tamarisci ssp. obscura var. balansae (Steph.) S.Hatt. 1972 Frullania moniliata horikawana (Reinw., Blume & Nees) Mont. 1842 Verd. in Hand.-Mazz. 1930 Typus: China: Yunnan, zwischen Haba und Dugwantsun se von Dschungdien [Chungtien], 3700 m 24.6.1915. Handel-Mazzetti 1352 Basionym: Distribution: China, Taiwan spec nov Verdoorn in Handel-Mazzetti Yang Bonner Hattori in Hara 1930 1960 1965 1971 - syn nov Hattori Hattori in Ohashi Kuo & Chiang Piippo Mizutani, Hasegawa & Iwatsuki 1973 1975 1988 1990 2009 = Frullania yuennanensis hornschuchiana 550 962 655 190 Symb Sinicae 5: 36-37, fig. 14(4-7) Quart J Taiwan Mus 13 (3&4): 233 [Taiwan?] Index Hepat 5: 329 Univ Mus Univ Tokyo Bull 2: 234 [? syn sub Fr. duthiana var appendiculata] 542 J Hatt Bot Lab 37: 130 [syn sub Fr. yunnanensis] 187 Univ Mus Univ Tokyo Bull 8: 236 [sub Fr. yuennanensis] 948 Taiwania 33: 12 [Index of Taiwan] 521 J Hatt Bot Lab 68: 42 [annotated catalogue of China] 2112 Type sepecimens Hatt Bot Lab (NICH) 26: Isotype. China, Prov. Yunnan. 24 June 1915. Handel-Mazzetti 1352 (NICH 218334). Steph. 1894 Nees 1845 Typus: Nepal in hb HORNSCHUCH in hb E E00002535 ( as "hornshuchiana - sd, Wallich sn) [isotypus] in hb GLM (hb LEHMANN) 06017 Basionym: Distribution: spec nov - syn nov Nees in G., L. & N. Mitten Verdoorn Bonner = Frullania nepalensis hornshuchiana Typus: Nepal 1845 1861 1930 1965 2850 574 293 655 Syn Hepat: 422 J Linn Soc Bot 5: 121 [syn sub Fr. nepalensis] Ann Bryol Suppl 1: 50 Index Hepat 5: 330 (Spreng.) Lehm. & Lindenb. 1845 Nees 1845 [orth var in hb E E00002535] sd, Wallich sn in hb HORNSCHUCH in hb E E00002535 ( as "hornshuchiana - sd, Wallich sn) [isotypus] Basionym: Distribution: = Frullania hornschuchiana Frullania nepalensis Nees 1845 (Spreng.) Lehm. & Lindenb. 1845 Page 206 horostipula S.Hatt. & Griffin III by Mizutani, Hasegawa & Iwarsuki 2009 [orth var] Basionym: Distribution: orth var Mizutani, Hasegawa & Iwatsuki 2009 = Frullania holostipula hosseana Typus: Siam: 2112 Type sepecimens Hatt Bot Lab (NICH) 26: [as horostipula] Holotype. Venezuela, Bocono Distr., Guaramacal. 2500-2600 m. Aug. 1975.D. Griffin III et at. 994 (NICH 432002). S.Hatt. & D.G.Griffin 1978 Steph. 1910 sd, Hossens sn in hb G 7947 [holotypus] Basionym: Distribution: spec nov Stephani Bonner Hattori Stephani 1910 1965 1982 1985 Singh, Kumar & Nath 2008 = Frullania wallichiana Spec Hepat 4: 348 Index Hepat 5: 330 J Hatt Bot Lab 51: 262 [syn sub Fr. wallichiana] Stephani Icon Hepat Inedit [Chonanthelia] A- 89, microfiche 47 fig 3195-3196 [from type; "Siam Dr Hossens"] 1281 Taiwania 53 (1): 54 [syn sub Fr wallichiana] Mitt. 1861 Frullania obscura huatuscana 686 655 495 928 (Sw.) Mont. 1840 Steph. 1911 Typus: Mexico. Huatusco sd, Liebmann in hb G Basionym: Distribution: Mexico spec nov spec nov - syn nov - syn nov Stephani Stephani Bonner Stotler Stotler Stephani 1911 1911 1965 1969 1970 1985 686 686 655 4761 194 928 Stephani 1985 928 Fulford & Sharp 1990 105 = Frullania intumescens Spec Hepat 4: 531 Spec Hepat 4: 511 Index Hepat 5: 330 [Mexico] Nova Hedwigia 18: 444 [syn sub Fr. intumescens] The genus Frullania, separata: 444 [syn sub Fr. intumescens] Stephani Icon Hepat Inedit [Thyopsiella] F- 62, microfiche 57 fig 3962 ["Mexico, Huatusco, leg Liebmann (Herb Gottsche sine nomine)"] Stephani Icon Hepat Inedit [Thyopsiella] F-114, microfiche 58 fig 4039 ["Mexico, Huatusca, Liebmann"] Mem NY Bot Gard 63: 43 [Mexico] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 huerlimannii var. latiloba S.Hatt. 1977 Basionym: Distribution: var nov - syn nov Hattori Miller, Whittier & Whittier Hattori Mizutani, Hasegawa & Iwatsuki = Frullania huerlimannii 1977 1983 1984 2009 479 609 503 2112 J Hatt Bot Lab 43: 422-423, fig 9 [var nov] [New Caledonia] Prod Flora Hepat Polynesiae: 129 J Hatt Bot Lab 57: 413 Type sepecimens Hatt Bot Lab (NICH) 27: Holotype. New Caledonia, Mt. Dzumac. 800 m. 29 Mar. 1951. H. Hurlimann 2446a (NICH 353790). S.Hatt. 1976 Page 207 hunanensis S.Hatt. 1981 Typus: China. Hunan, .. , 1200-1300m 8.8.1917, Handel-Mazzetti 11190 (Diary No 2206 in hb NICH [holotypus], in hb W [isotypus] Basionym: Distribution: spec nov - comb nov Hattori Hattori & Lin Piippo Mizutani, Hasegawa & Iwatsuki 1981 1985 1990 2009 491 505 521 2112 = Frullania kagoshimensis ssp. hunanensis hutchinsiae J Hatt Bot Lab 49: 152-155, fig 244(a-l) J Hatt Bot Lab 59:137 [comb. nov - Fr. kagoshimensis ssp. huanensis] J Hatt Bot Lab 68: 42 [annotated catalogue of China] Type sepecimens Hatt Bot Lab (NICH) 27: Holotype. China, Mt. Yunshan. 1200-1300 m. 8 Aug. 1917. H. F. v. Handel-Mazzetti 11190 (NICH 380845, 380846). (S.Hatt.) S.Hatt. & P.J.Lin 1985 [auct. non (Hooker) Nees 1838] Sull. 1846 Typus: USA. South Carolina, Pigeon river falls Basionym: Distribution: - comb nov Sullivant Spruce 1846 1884 764 243 Musci Alleghaniensis: 64 [Exsicc No 271, reprint of labels] Trans Proc Bot Soc Edinburgh 15: 62 = hutchinsiae var. (alpha) (Hooker) Nees 1838 {European specimen} Basionym: Distribution: Japan Nees Montagne Nees in G., L. & N. Sande Lacoste in Junghuhn Gottsche Gottsche Lindberg Piippo Yamada & Iwatsuki = Jubula hutchinsiae 1838 1842 1845 1857 1863 1867 1875 1990 2006 261 469 2850 1490 103 104 1582 521 1470 Naturg Europ Leberm 3: 240-244 Ann Sci Nat Bot, ser 2, 18: 14 Syn Hepat: 426 Plantae Junghuhnianae, Hepaticae: 537 Separata of Mex. Leverm.: 242 K Danske Vid Selks Skrifter, ser 5, 6: 338 [Mexico] Acta Societatis Scientiarum Fennicae 10: 475 J Hatt Bot Lab 68: 42 [annotated catalogue of China] J Hatt Bot Lab 99: 30 [Catalog of Japan] (Hooker) Dumort. 1822 Page 208 hutchinsiae var. (beta) (Nees) Nees 1838 {Javanese specimen} Basionym: Jungermannia hutchinsiae var. (beta) Nees 1830 Distribution: Nees Nees in G., L. & N. Hooker in G., L. & N. Sande Lacoste in Junghuhn Mitten Sande Lacoste 1838 1845 1847 1857 1861 1864 261 2850 2848 1490 574 842 Sande Lacoste in Veth 1884 807 Mitten Iishiba Sawada 1891 1909 1909 582 890 930 Guerke 1978 181 = Jubula hutchinsiae ssp. javanica Naturg Europ Leberm 3: 240-244 [Java] Syn Hepat: 426 Syn Hepat: 775 Plantae Junghuhnianae, Hepaticae: 537 J Linn Soc Bot 5: 122 [India: Khasi] Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 312 [Java, Sumatra, Japan] Midden Sumatra, Reizen en Onderzoekingen der Sumatra Expeditie, 4 (2): 44 [Sumatra] Trans Linn Soc London, ser 2, 3: 203 [Japan] Bot Mag Tokyo 23: 363 [Japan] Bot Mag Tokyo 23 (no 268): 178-183 [key to Japanese species, in Japanese] Bryophyt Bibl 17: 82-83 (Steph.) Verd. 1928 hutchinsiae var. ? Underwood 1884 Basionym: Distribution: Underwood 1884 = Jubula pennsylvania 1708 Bull Illinois State Lab Nat Hist 2: 65 (=Bulletin of the Illinois State Laboratory of Natural History) (Steph.) Evans 1905 hutchinsiae var. bogotensis (delta) Gottsche 1864 Typus: Manzanos, 2700m sd, Lindigii Basionym: Distribution: Gottsche 1864 = Jubula pennsylvania ssp. bogotensis 478 Ann Sci Nat Bot, ser 5, 1: 167 (Steph.) Guerke 1978 hutchinsiae var. integrifolia (beta) Lindb. 1875 Basionym: Distribution: Lindberg Brotherus Guerke = Jubula pennsylvania 1875 1892 1978 1582 Acta Societatis Scientiarum Fennicae 10: 474 922 Acta Soc Sci Fennicae 19 (12): 144 [Caucasus] 181 Bryophyt Bibl 17: 82-83 (Steph.) Evans 1905 Page 209 hutchinsiae var. mexicana (gamma) Nees 1847 Basionym: Distribution: Nees in G., L. & N. Gottsche Gottsche Guerke Fulford & Sharp 1847 1863 1867 1978 1990 = Jubula pennsylvania ssp. bogotensis 2848 103 104 181 105 Syn Hepat: 775 Separata of Mex. Leverm.: 242 K Danske Vid Selks Skrifter, ser 5, 6: 338 [Mexico] Bryophyt Bibl 17: 84 Mem NY Bot Gard 63: 43 [Mexico] (Steph.) Guerke 1978 hutchinsiae var. sullivantii Spruce by GUERKE 1978 [orth var] Basionym: Distribution: Guerke 1978 181 Bryophyt Bibl 17: 83 [error; = Jubula not Frullania] = Jubula hutchinsiae var. sullivantii Spruce 1884 hutchinsiae var. β compacta Carrington 1863 Typus: Type:—IRELAND. Eagle’s Nest; Cromaglan (MANCH, syntypes). Basionym: Distribution: Carringtion Hentschel et alii = Jubula hutchinsiae hypogyna 1863 2015 Trans. Bot. Soc. Edinburgh 7: 456 4097 Phytotaxa 220 (2): 137 [syn sub Jubula hutchinsiae (Hook.) Dumort.] (Hooker) Dumort. 1822 Steph. 1894 Typus: Philippines: Luzon, Benguet 1884/1885, Micholitz sn hb STEPHANI 28, in hb G [holotypus] Basionym: Distribution: spec nov - syn nov - syn nov Stephani Stephani Verdoorn Svihla Bonner Hattori Stephani = Frullania rio-janeirensis 1894 1910 1930 1956 1965 1972 1985 228 686 293 380 655 538 928 Hedwigia 33: 138 Spec Hepat 4: 347 Ann Bryol Suppl 1: 70 Bryologist 59: 278-279 [syn sub Fr. riojaneirensis] Index Hepat 5: 331 J Hatt Bot Lab 36: 135-140, fig 20(a-j) [syn sub Fr. galeata] Stephani Icon Hepat Inedit [Chonanthelia] A- 90, microfiche 47 fig 3197 [from type; "Luzon Benguet Micholitz No 28 1884/85"] (Raddi) Ångstr. 1876 Page 210 hypogyna [auct. non Steph. 1894] Steph. ex Steph. 1985 [pro syn] {Bolivia} Typus: Bolivia hb BESCHERELLE Basionym: Distribution: Stephani = Frullania complicata hypoleuca 1985 928 Stephani Icon Hepat Inedit [Chonanthelia] A- 64, microfiche 47 fig 3161[from type; "olim Frullania hypogyna Steph Bolivia Herb Bescherelle"] Steph. 1911 Taylor ex Watts 1901 [nom nud] Basionym: Distribution: Watts Scott & Bradshaw Carthy = Frullania hypoleuca ichthyostoma 1901 1986 2003 1603 Proc Linn Soc New South Wales 26 (2): 215 56 Brunnonia 8 (1): 35 [Australia] 1125 Flora Australia Suppl Ser 21: 109 [nom nud] Nees 1843 Colenso 1889 Typus: New Zealand. County of Waipawa, Dannevirke 1888, W. Colenso 1350 in hb G 20596 Basionym: Distribution: spec nov - syn nov Colenso Stephani Bonner Hamlin = Frullania squarrosula ichthystoma 1889 1892 1965 1972 636 210 655 659 Trans Proc New Zealand Inst 21: 77 ["ichthyostoma"] J Linn Soc Bot 29: 274 [syn sub Fr. squarrosula] Index Hepat 5: 331[as ichthystoma, orth err] Rec Dominion Mus 7(19): 262 [New Zealand] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Colenso by Bonner 1965 [orth var] Basionym: Distribution: Bonner = Frullania ichthyostoma Frullania squarrosula 1965 655 Index Hepat 5: 331 Colenso 1889 (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Page 211 illyrica Grolle in Meyer & Grolle 1963 Typus: Albania.Tepelena, Luzati Tal, Kalkfelsen, ca. 400m 9.11.1961, F.K. Meyer 6062 / 6602 ? in hb JE (hb GROLLE) [holotypus] Basionym: Distribution: Albania spec nov - comb nov - syn nov Grolle in Meyer & Grolle Grolle 1963 1970 Grolle Bischler, Jovet-Ast & Baudoin Grolle Schuster Düll Bisang, Schumacker, Sérgio & Grolle Schuster Frey, Frahm, Fischer & Lobin Grolle & Long 1976 1980 1983 1983 1983 1989 Sabovljec, Ganeva, Tsakiri & Stefanut Vana 2001 = Frullania inflata implexicaulis 1992 1995 2000 2002 109 615 Feddes Repert 68: 101-107, fig 1-3 [Albania] Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 316-317 [key to europa & macaronesia] 183 Feddes Repert 87 (3/4): 194 [types] 149 Crypt Bryol Lichen 1 (3): 255 [Albanien] 144 J Bryol 12: 421 [checklist Europe] 440 Phytologia 53 (5): 366 [comb nov as Fr. inflata ssp illyrica] 620 Bryol Beitr 2: 67 [Europe & Macaronesia] 276 Giorn Bot Ital 122: 259. 264 [key to Europe & Macaronesia, an Albanian endemit] 603 Hepat N America 5: [comb nov as Fr inflata var illyrica] 606 Kleine Kryptogamenflora 5: 116 [key to european species] 760 J Bryol 22: 111 [checklist of Europe and Macaronesia] & 124 [syn sub Fr. inflata] 1296 Biol Conservation 101: 79 [rare; conservation interest in Albania & Turkey] 1342 Portug Acta Biol (B) 20: 27 [evaluation of taxonomy] Gottsche 1845 [non Herzog 1921] Colenso 1887 Typus: New Zealand. County of Waipawa, Norsewood 1886, Colenso 1383 ex hb K, in hb G Basionym: Distribution: spec nov - syn nov Colenso Stephani Stephani Bonner 1887 1892 1910 1965 634 210 686 655 Hamlin 1972 659 = Frullania deplanata indica Trans Proc New Zealand Inst 19: 299 J Linn Soc Bot 29: 273 [syn sub Fr. deplanata] Spec Hepat 4: 407-408 [not as syn sub Fr. deplanata] Index Hepat 5: 332 [error: not as syn sub Fr. deplanata in STEPH 1910] Rec Dominion Mus 7(19): 262 [New Zealand] Mitt. 1855 Steph. 1910 Typus: India: South Canara, W Ghats, Mt. Kudremukh 1905, Pfleiderer sn ex hb LEVIER 5031, in hb G [holotypus] Basionym: Distribution: India spec nov - syn nov Stephani Verdoorn Chopra Herzog Svihla Parihar Bonner Stephani 1910 1930 1943 1955 1956 1962 1965 1985 Singh, Kumar & Nath 2008 = Frullania wallichiana Frullania obscura 686 293 205 107 380 1584 655 928 Spec Hepat 4: 347 Ann Bryol Suppl 1: 70-72 [syn sub Fr. wallichiana] J Indian Bot Soc 22: 249 [India] Feddes Repert 57 (1/2): 198 [Peru] Bryologist 59: 277-278 Univ Allahabad studies, Bot sect 1-2: 19 [India, syn sub Fr wallichiana] Index Hepat 5: 333 Stephani Icon Hepat Inedit [Chonanthelia] A- 91, microfiche 47 fig 3198-3199 [from type; "West Ghats South Canara ... India Rev Pfleiderer Mai 1905 Levier 5031"] 1281 Taiwania 53 (1): 54 [syn sub Fr wallichiana] Mitt. 1861 (Sw.) Mont. 1840 Page 212 indica Schiffn. ex Verd. 1930 [pro syn] Basionym: Distribution: Verdoorn Miller, Whittier & Whittier = Frullania neurota inflata 1930 1983 293 609 Ann Bryol Suppl 1: 66 Prod Flora Hepat Polynesiae: 129 Taylor 1846 Herzog 1921 [hom illeg] [non Gottsche 1845] Typus: Bolivia. "an Bäumchen bei" [on small tree by] bei Incacorral, 2200 m -.6.1911, T. Herzog sn in hb JE [holotypus] Basionym: Distribution: USA, Bolivia spec nov - syn nov Herzog Bonner Yuzawa Gradstein, Meneses & Arbe Müller & Zündorf 1921 1965 1991 2003 2007 Yano 2008 = Frullania rio-janeirensis 223 655 525 975 1442 Biblioth Bot 88: 31, fig. 18 Index Hepat 5: 333 J Hatt Bot Lab 70: 254 [syn sub Fr. riojaneirensis] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. riojaneirensis] http://www2.unijena.de/biologie/spezbot/herbar/bryotypes/Uebersicht%20F.htm 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr riojaneirensis; Brazil; catalog] (Raddi) Ångstr. 1876 inflata var. illyrica (Grolle in MEYER & Grolle) R.M.Schust. 1992 Basionym: Frullania illyrica Distribution: Albania, Italy Grolle in Meyer & Grolle 1963 Schuster 1992 Grolle & Long Düll Papp & Erzberger 2000 2003 2007 = Frullania illyrica 603 Hepat N America 5: 203, 213 [Fr illyrica, comb nov as Fr. inflata var inflata], 760 J Bryol 22: 124 [Fr illyrica syn with Fr inflata] 618 Bryol Beitr 4: xxxx [a relict Mediterranean-montane species] 2678 Studia bot hung 38 : 83-84 [Montenegro, we prefer keeping this name pending further research] Grolle in Meyer & Grolle 1963 Frullania inflata Gottsche 1845 [non Herzog 1921] inflata var. magnicorpa Yamada 1974 Typus: Japan Basionym: Distribution: Japan, Korea var nov -syn nov Yamada Choe Yamada & Iwatsuki = Frullania inflata 1974 1980 2006 406 J Hatt Bot Lab 44: 185 1171 Illust Fl & Fauna Korea 24: 734 [syn nov sub Fr inflata; South Korea] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] Gottsche 1845 [non Herzog 1921] Page 213 inflata var. mayebarae (S. Hatt.) R.M.Schust. 1992 Basionym: Frullania mayebarae Distribution: comb nov S.Hatt. 1952 Schuster Yuzawa Yamada & Iwatsuki Katagiri & Furuki 1992 2001 2006 2018 603 1660 1470 4966 Hepat N America 5: 203 Natural Environmental Sci Res 14: 16 [Japan, syn sub Fr inflata] J Hatt Bot Lab 99: 30 [Catalog of Japan] Hattoria 9: 66 [Checklist of Japanese liverworts and hornworts] [annotation 106] = Frullania inflata var. mayebarae (S.Hatt.) Yamada 1974 Frullania inflata Gottsche 1845 [non Herzog 1921] inflata var. mayebarae (S.Hatt.) Yamada 1974 Basionym: Frullania mayebarae Distribution: Japan comb nov - syn nov S.Hatt. 1952 Yamada Choe Hattori & Lin Schuster Bapna & Kachroo Yuzawa = Frullania inflata 1974 1980 1985 1992 2000 2001 406 1171 505 603 712 1660 Misc Bryol Lichen 6 (9): 163-164, fig 1(d-e) [comb nov] Illust Fl & Fauna Korea 24: 734 [South Korea] J Hatt Bot Lab 59: 136 [syn sub Fr. inflata] Hepat N America 5: 203 [comb nov] Hepaticology in India II: 173 [syn sub Fr. inflata] Natural Environmental Sci Res 14: 16 [Japan, syn sub Fr inflata] Gottsche 1845 [non Herzog 1921] inflata var. stylifera R.M.Schust. 1983 Typus: USA. Minnesota -, RMS 14205 Basionym: Distribution: var nov - comb nov Schuster Schuster 1983 1992 = Frullania stylifera inflata ssp. illyrica - comb nov Phytologia 53 (6): 366 [USA] Hepat N America 5: 210 [as Fr. stylifera] (R.M.Schust.) R.M.Schust. 1992 (Grolle in Meyer & Grolle) R.M.Schust. 1983 Basionym: Frullania illyrica Distribution: comb nov 440 603 Grolle in Meyer & Grolle 1963 Schuster Bisang, Schumacker, Sérgio & Grolle Schuster Grolle & Long 1983 1988 440 276 1992 2000 603 760 Phytologia 53 (5): 366 [Fr illyrica, comb nov as Fr. inflata ssp. illyrica] Giorn Bot Ital 122: 264 [savoir si les oléocorps peuvent varier dans une telle mesure ou si,] Hepat N America 5: 196 [Fr illyrica, comb nov as Fr. inflata var inflata], J Bryol 22: 111 [checklist of Europe and Macaronesia] & 124 [Fr illyrica syn sub Fr. inflata] = Frullania inflata var. illyrica (Grolle in MEYER & Grolle) R.M.Schust. 1992 Frullania illyrica Frullania inflata Grolle in Meyer & Grolle 1963 Gottsche 1845 [non Herzog 1921] Page 214 inflatiloba Steph. 1910 Typus: Mauritius 1904, Völtzkow in hb G 10704 [holotypus] in hb UPS [isotypus] Basionym: Distribution: Mauritius spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1910 1965 1976 1985 Gotteche Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania diptera inflatistipula 686 655 185 928 Spec Hepat 4: 378 Index Hepat 5: 334 Bull Jard Bot Nat Belg 46 (1/2): 194 & 197 Stephani Icon Hepat Inedit [Galeiloba] C- 21, microfiche 50 fig 3373 [from type; "Maurice, leg Völtzkow 1904, inter alias Frull"] 250 Bryophyt Bibl 48: 65 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 229, fig 151( C)from type [syn sub Fr diptera,West Africa] (Lehm.) Drège 1843 J.B.Jack ex Bonner 1965 [nom herb] Typus: Fiji. Viti - Levu 1864, E. Graeffe 585 in hb G (ex hb JACK) sub Fr. secundiflora Basionym: Distribution: Bonner Miller, Whittier & Whittier = Frullania secundiflora 655 609 Index Hepat 5: 334 Prod Flora Hepat Polynesiae: 129 Mont. 1843 [non Taylor 1845] Frullania nodulosa inopinata 1965 1983 (Reinw., Blume & Nees) Nees 1845 Verd. 1932 Typus: China. Prov. Kwangtung, Lofaushan (Loh Fau Mt.) 1917 or -.8.1927, E.D. Merrill 11150 [with Fr. kashyapii] in hb VERDOORN, in hb PNH, in hb FH [typus], in hb Manila [isotypus] Basionym: Distribution: spec nov Verdoorn Verdoorn Bonner Hattori 1932 1932 1965 1981 552 554 655 491 - syn nov Hattori & Lin Piippo 1985 1990 505 521 = Frullania motoyana Ann Bryol 5: 164 Nederl Kruidk Arch Jrg 1932 (2): 497 [China] Index Hepat 5: 334 J Hatt Bot Lab 49: 147 [may be conspecific with Fr. warnckeana or Fr. gracilis-complex] J Hatt Bot Lab 59: 159 [syn sub Fr. motoyana] J Hatt Bot Lab 68: 42 [annotated catalogue of China] Steph. 1911 Page 215 inouena Steph. ex Steph. 1985 [nom herb] {Japan} Typus: Japan. Tosa 1898, Inoue no 12 in hb G Basionym: Distribution: Japan - syn nov Choe Stephani 1980 1985 1171 Illust Fl & Fauna Korea 24: 734 [South Korea] 928 Stephani Icon Hepat Inedit [Galeiloba] C-240, microfiche 53 fig 3660 [Japonia, Tosa, Inoué No 12, 1898"] = Frullania muscicola var. inuena (Steph.) Kamim. 1961 Frullania muscicola insciflora Steph. 1894 Gottsche ex Steph. 1985 [pro syn] Basionym: Distribution: Stephani 1985 = Frullania franciscana insularis - syn nov Stephani Icon Hepat Inedit [Thyopsiella] F-192c, microfiche 60 fig 4150 ["syn Frull insciflora G ms, California coast, Bolander, ... Evans"] M. Howe 1894 Beauverd in Steph. 1924 Basionym: Frullania renauldii Distribution: Mauritius nom nov 928 Steph., Spec Hepat 4: 627 (17 Jul 1911) 1911 [non Steph., Spec Hepat 4: 487 (28 Feb 1911)] Beauverd in Stephani 1924 Bonner Vanden Berghen Grolle Wigginton 1965 1976 1995 2002 Wigginton Wigginton 2004 2004 = Frullania apicalis integristipula 909 Spec Hepat 6: 570 [Frullania insularis Beauverd, nom. nov. = F. Renauldii Stephani in Spec. Hep., vol. IV [1911 ], p. 627 655 Index Hepat 5: 334 185 Bull Jard Bot Nat Belg 46 (1/2): 52 250 Bryophyt Bibl 48: 62 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apicalis, West Africa] Mitt. 1879 [auct. non (Nees) Nees 1845] Nicholson 1942 Typus: Hawaii. Basionym: Distribution: Hawaii - syn nov Nicholson Miller Miller, Whittier & Whittier = Frullania hypoleuca 1942 1963 1983 679 834 609 Rev Bryol Lichen 13: 144 Ark Bot, ser 2, 5: 519-520 [syn nov sub Fr hypoleuca] Prod Flora Hepat Polynesiae: 129 Nees 1843 Page 216 integristipula var. reflexistipuloides Verd. 1929 Typus: Indonesia. Java, Prov. Preanger, Mt. Pangerango, in horto montano Tjibodas, 1420m 4.5.1894, Schiffner 2442 Basionym: Distribution: var nov - syn nov Verdoorn Verdoorn Verdoorn Bonner Hattori = Frullania integristipula intermedia 1929 1930 1934 1965 1980 245 293 241 655 490 Ann Bryol 2: 153-154, fig 10 Ann Bryol Suppl 1: 176, fig 277 Ann Bryol Suppl 4: 210 [Java] Index Hepat 5: 335 J Hatt Bot Lab 47: 228 [syn sub Fr. integristipula] (Nees) Nees 1845 fo. billardieriana (Nees & Mont. in Mont.) Verd. 1930 Basionym: Frullania billardieriana Nees & Mont. in Mont. 1843 Distribution: Indonesia (Ceram, Ambon), Philippines, New Guinea, Borneo, Vanuatu (Aneityum), New Caledonia, Fiji (Viti Levu) comb nov - syn. nov. Verdoorn Verdoorn 1930 1930 293 549 Bonner Campbell Hattori Miller, Whittier & Whittier 1965 1971 1976 1983 655 674 80 609 = Frullania intermedia intermedia Ann Bryol Suppl 1: 164, fig 268(:159) [ Nova Guinea 14 (4): 546 [comb nov; without basionym] [New Guinea ?] Index Hepat 5: 336 J Roy Soc New Zealand 1 (1): 13 [Fiji] J Hatt Bot Lab 40: 483, fig 186(e-i) [Sarawak; = Fr. intermedia] Prod Flora Hepat Polynesiae: 130 (Reinw., Blume & Nees) Nees 1845 fo. billardieriana [auct. non (N. & M. in M.) Verd.] S.Hatt. 1976 Basionym: Distribution: Malaysia (Sarawak), Borneo auct - syn nov Hattori Hattori Grolle & Piippo = Frullania intermedia intermedia 1976 1982 1984 80 495 143 J Hatt Bot Lab 40: 483, fig 186(e-i) [Sarawak, new to Borneo] J Hatt Bot Lab 51: 239 [syn sub Fr. intermedia] Acta Bot Fennica 125: 20 [Cat. West Melanesia] (Reinw., Blume & Nees) Nees 1845 fo. fallacoides Verd. 1930 Basionym: Distribution: fo nov - syn nov Verdoorn Bonner Hattori = Frullania intermedia 1930 1965 1980 293 655 490 Ann Bryol Suppl 1: 164 Index Hepat 5: 336 J Hatt Bot Lab 47: 187 (Reinw., Blume & Nees) Nees 1845 intermedia var. amboinensis (Schiffn.) Verd. 1930 Basionym: Frullania amboinensis Distribution: comb nov - syn nov Verdoorn Bonner Hattori Grolle & Piippo = Frullania intermedia Schiffn. 1890 1930 1965 1980 1984 293 655 490 143 Ann Bryol Suppl 1: 163-164 Index Hepat 5: 335 J Hatt Bot Lab 47: 187 Acta Bot Fennica 125: 20 [Cat. West Melanesia] (Reinw., Blume & Nees) Nees 1845 Page 217 intermedia var. subcommutata Steph. in Miller, Whittier & Whittier 1983 Basionym: Distribution: Miller, Whittier & Whittier = Frullania subcommutata Frullania angulosa 1983 609 Prod Flora Hepat Polynesiae: 129 Steph. 1908 Mitt. in Seemann 1871 Frullania intermedia var. non-apiculata S.Hatt. 1975 intermedia var. submorokensis S.Hatt. 1976 Typus: Indonesia. Sarawak, (Borneo), Mt. Kinabalu Basionym: Distribution: var nov - syn nov Hattori 1976 Hattori 1982 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania intermedia 80 J Hatt Bot Lab 40: 483-484, fig 187 495 J Hatt Bot Lab 51: 239 2112 Type sepecimens Hatt Bot Lab (NICH) 27-28: Holotype. Borneo, Mt.Kinabalu. 600-700 m. 29 May 1963. M. Mizutani 3081 (NICH 253081). (Reinw., Blume & Nees) Nees 1845 intumescens var. sabanetica (Gottsche) Gradstein 2021 Basionym: Frullania sabanetica Distribution: Colombia, Ecuador invalid comb nov - nom corr Gottsche 1864 Gradstein Gradstein 2020 2021 Gradstein 2021 Gradstein & Pocs 2021 Frahmia 17: 16 [ invalid, basionym not cited] 5642 Mem NY Bot Gard 121: 370 [key -Ecuador, Colombia] as Frullania intumescesns var. sabanetica (syn for Fr sabanetica, closterantha) 5642 Mem NY Bot Gard 121: 370 [key -Ecuador, Colombia, comb nov, nom nov by Gradtsein & Pócs, Nova Hedwigia 112] 5888 Nova Hedwigia 112(1-2): 58-59 [Venezuela, = Fr. intumescens var closterantha] = Frullania intumescens var. closterantha (Spruce) Gradstein & Pócs 2021 inuena Steph. 1910 Typus: Japan. Tosa, Ogawa -.4.1898, Inoue 12 in hb G Basionym: Distribution: spec nov comb nov Stephani Iishiba Hattori Kamimura Kamimura Bonner Bapna & Kachroo = Frullania muscicola 1910 1912 1952 1952 1961 1965 2000 686 888 529 873 199 655 712 Spec Hepat 4: 398 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] J Hatt Bot Lab 8: 29 Contrib Fl Hepat Shikokuensem, Kochi: 133 J Hatt Bot Lab 24: 25 [basionym of Fr muscicola var inuena] Index Hepat 5: 336 Hepaticology in India II: 175 [syn sub Fr. muscicola var inuena] Steph. 1894 Page 218 involuta Hampe ex Husnot 1875 [nom inval] Typus: Antilles. Martinique [Sur les rochers du bord de la route d'Alma prés des Pitons du Carbet (Mart.). Alt 500m] 1899 !?!, Husnot 223 in hb G 1298 [lectotypus by Uribe-M & Gradstein 2003], in hb BM 664353 [isolectotypus] Basionym: Distribution: Husnot 1875 = Frullania involuta Hampe ex Steph. 1911 Frullania uleana involuta 2435 Rev Bryol 2(1): 4 ["Frullania involuta Hpe., H. 223 - Sur les rochers du bord de la route d'Alma prés des Pitons du Carbet (Mart.). Alt 500m Steph. 1894 Hampe ex Steph. 1911 Typus: Antilles. Martinique 1899, Husnot 223 in hb G 1298 [lectotypus by Uribe-M & Gradstein 2003], in hb BM 664353 [isolectotypus] Basionym: Distribution: Mexico, Panama, Cuba, Haiti, Martinique, Guadeloupe, Venezuela, Trinidad, Galapagos Isl [Ecuador], Guyana, Surinam, Brazil spec nov syn nov. Husnot 1875 Bescherelle 1893 Duss Hampe ex Stephani Pagán Clark & Svihla Clark & Svihla 1904 1911 1942 1947 1948 Schiffner & Arnell Bonner Gradstein & Weber Mägdefrau Yano Stephani 1964 1965 1982 1983 1984 1985 Kron Fulford & Sharp Yano Grims Uribe & Gradstein 1988 1990 1995 2001 2003 Martinez Alvarenga, Silva et alii Uribe 2005 2007 2008 Yano Gradstein 2008 2009 Lavocat Bernard & SchäferVerwimp Gradstein & Ziemmeck Schäfer-Verwimp Lima Gradstein 2011 2435 Rev Bryol 2(1): 4 ["Frullania involuta Hpe., H. 223 - Sur les rochers du bord de la route d'Alma prés des Pitons du Carbet (Mart.). Alt 500m 3815 J Bot Morot 7 (9): 179 [Guadeloupe : L'Heminier, Martinique : Rochers de la route d'Alma; Husnot (n° 223).] 3132 Flore Cryptogamique des Antilles Françaises: 132 [Martinique] 686 Spec Hepat 4: 595-596 3152 Bryologist 45(4): 108 [Guadeloupe] 321 Bryologist 50: 271-275, fig 1-27 325 Bryologist 51: 24 [key to Fr. of Middle America: Haiti, Martinique, Trinidad (Dutch Guiana)] 198 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 89 [Brazil] 655 Index Hepat 5: 337 496 J Hatt Bot Lab 52: 148 [on 2 Galapagos Isls] 1206 Nova Hedwigia 13: 21 [Venezuela] 501 J Hatt Bot Lab 56: 505 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Meteriopsis] E-25, microfiche 55 fig 3800 ["Antillen, Husnot leg"] - 3801 510 J Hatt Bot Lab 64: 355, fig 6(f-n) 105 Mem NY Bot Gard 63: 44 [Mexico] 2095 J Hatt Bot Lab 78: 142 [annotated checklist of Brazil] 1136 Moosherbar in hb LINZ, from Brasilien [Internet] 1071 Crypt Bryol 24 (3): 204-206, fig 6(a-h) [from type] [belongs to subgen Thyopsiella] 2307 Cuba: Parque Nacional “Alejandro de Humboldt", p. 222 1485 Acta bot bras 21(2): 351, fig 17-22 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] 1684 Boletim Instituto Botanica 19: 32 [Brazil; catalog] 2161 Bibliotheca Lichenologica 99: 149 [Galapagos Isl, Note: the material from Galapagos needs further study, identification tentative.] 3130 Cryptogamie, Bryologie 32(3): 243 [Guadeloupe], 251 [Martinique] 2012 2014 2019 2020 4845 Checklist of Galapagos Horn- Liverworts [Galapagos] 3977 Phytotaxa 172(3):213 [Panama] 5121 Frullania no Brasil [Thesis]: 160-162, fig. 19 [distribution] [syn sub Fr. uleana] = Frullania uleana Steph. 1894 Page 219 involuta var. subintegra Steph. ex Bonner 1965 [nom herb] {Suriname} Typus: Suriname. "Dutch Guiana" Eustarius Isl 1889, Suringar 141 in hb G Basionym: Distribution: Surinam Bonner 1965 655 Index Hepat 5: 337 = itatiaja Steph. 1910 Typus: Brazil: Brasilia, Sierra da Itatiaja, 2000 m -.3.1894, Ule 453 in hb G [holotypus] Basionym: Distribution: Brazil spec nov Stephani Bonner Yano Stephani 1910 1965 1984 1985 686 655 501 928 Yuzawa Yuzawa 1991 1991 525 525 = Frullania obscura itoana Spec Hepat 4: 334 Index Hepat 5: 337 J Hatt Bot Lab 56: 505 [checklist Brazil] Stephani Icon Hepat Inedit [Chonanthelia] A- 37, microfiche 46 fig 3122 [from type; "Brasilia Ule 453 - ob eine Form von Fr. decidua Spruce?"] J Hatt Bot Lab 70: 268, fig. 44 [redescription of type specimen] J Hatt Bot Lab 70: 264 [syn sub Fr. arecae] (Sw.) Mont. 1840 Kamim. 1961 Typus: Japan. Kochi-Pref., Amatsubo-ruuro 12.8.1943, Kamim. 7271 in hb NICH 46876 Basionym: Distribution: Japan, China (Yunnan) spec nov - syn nov Kamimura Bonner Inoue in Inoue & Iwatsuki Hattori Lemos Michel Schuster Yuzawa Konrat & Braggins Bai Yamada & Iwatsuki Mizutani, Hasegawa & Iwatsuki = Frullania hypoleuca iwatsukii ssp. viernamica 1961 1965 1970 1981 1983 1992 2000 2001 2001 2006 2009 199 655 62 492 306 603 1659 838 1742 1470 2112 J Hatt Bot Lab 24: 77-79, fig 28(1-35), map 8 Index Hepat 5: 337 Bull Natn Sci Mus Tokyo 13 (3): 478, fig 1 [syn sub Fr. uvifera] J Hatt Bot Lab 49: 372-373 [syn sub Fr. hypoleuca] Revta brasil Bot 6 (2): 117 [syn sub Fr. caroliniana] Hepat N America 5: 118 [valid species] Natural Environmental Sci Res 13: 4 [Japan, syn sub Fr hypoleuca] J Hatt Bot Lab 91: 250-256 [syn sub Fr. hypoleuca] Guizhou Science 19(4): 49 [syn sub Fr hypoleuca] J Hatt Bot Lab 99: 30 [Catalog of Japan] Type sepecimens Hatt Bot Lab (NICH) 28: Holotype. Japan, Shikoku, Kochi-ken, Amatubo-mura. On bark. 12 Aug. 1943. M Kamimura 7271 (NICH H46876). Nees 1843 S.Hatt. by X.L.Bai in C.Gao & Cao 2000 [orth var] Basionym: Distribution: orth var Bai in Gao & Cao 2000 = Frullania iwatsukii ssp. vietnamica 1019 Fl Yunnanica 17: 434 S.Hatt. 1976 Frullania tamarisci ssp. obscura var. vietnamica (S.Hatt.) S.Hatt. 1985 Page 220 iwatsukii ssp. vietnamica S.Hatt. 1976 Typus: Vietnam. Mts. Hoang-Lien-Son, 1650m 13.11.1960, Polonicus in hb EGR [holotypus], in hb NICH [isotypus] Basionym: Distribution: Vietnam ssp nov comb nov Hattori Hattori in Hattori & Lin 1976 1985 Bai in Gao & Cao 2000 Kis 2004 Mizutani, Hasegawa & Iwatsuki 2009 412 505 Misc. Bryol. Lichenol. 7: 87- 89, fig 88(a-h) Fr iwatsukii ssp vietnamica J. Hatt. Bot. Lab. 59: 162 F. tamarisci subsp. obscura var vietnamica (comb. nov) für F. iwatsukii Hatt. subsp. vietnamica 1019 Fl Yunnanica 17: 434 [as Fr iwatsukii var vietnamica] 1377 F Hist Nat Mus Matraensis 28: 15 [holotype in EGR] 2112 Type sepecimens Hatt Bot Lab (NICH) 28: Isotype. Vietnam, Montes Hoang-Lien-Son. 13 Nov. 1960. Polonicus (NICH 345479). = Frullania tamarisci ssp. obscura var. vietnamica (S.Hatt.) S.Hatt. 1985 jackiana Beauverd in Steph. 1924 ex Bonner 1965 [nom nud] Basionym: Distribution: Mexico nom nov Stephani Bonner 1924 1965 = Frullania esenbeckiana jackii 909 655 Spec Hepat 6: 570 [not in this paper, as stated by BONNER] Index Hepat 5: 337 Beauverd 1924 [auct. non Gottsche 1863] S.Hatt. 1955 {from Japan} Basionym: Distribution: Hattori Hattori Yamada & Iwatsuki = Frullania davurica jackii 1955 1976 2006 1683 J Hatt Bot Lab 15: 91 [on serpentine rocks] 67 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21 [syn sub Fr. davurica] 1470 J Hatt Bot Lab 99: 30 [jackii auct HATTORI 1955 = davurica - Catalog of Japan] Hampe ex Gottsche, Lindenb. & Nees 1845 [auct. non Gottsche in G. & R.] Verd. 1930 {India} Basionym: Distribution: - syn nov Verdoorn 1930 Verdoorn 1932 Mues, Hattori, Asakawa & Grolle 1984 Söderström, Hassel & Weibull 2002 = Frullania retusa 500 J Hatt Bot Lab 56: 235-236 2672 Distribution maps - Northwestern Europe, vol 1: 20 [map] Mitt. 1861 Page 221 jackii Typus: [auct. non Gottsche in G. & R.] Verd. 1930 {China} Mazzetti 6771 Basionym: Distribution: Verdoorn in Handel-Mazzetti 1930 Nicholson 1930 Reimers 1931 Hong & Kim 1961 Inoue 1976 Mues, Hattori, Asakawa & Grolle 1984 Hattori & Lin 1985 Konstantinova & Bakalin 2009 = Frullania davurica jackii 550 Symb Sinicae 5: 37 1117 Ann Bryol 3: 152-153 [Yunnan] 229 Hedwigia 71 (1/2): 34-35 [China] Hikobia 2: <244-252> [aus 717] 734 Illust Jap Hep 2: 62-63, pl 31(1-6), foto, map of Japan 500 J Hatt Bot Lab 56: 235-236 505 J Hatt Bot Lab 59: 132 2113 Arctoa 18: 15 [Checklist of liverworts of Russia: EUR, CAUC] Hampe ex Gottsche, Lindenb. & Nees 1845 fo. umbrosa Schiffn. ex Pando 2000 [nom herb in hb MA(-Hepat 1108)] Typus: Austria. Tirol, Prosegger-Klamm bei Windisch-Matrei, an beschatteten Schieferfelsen, ca 1100m 15.9.1902, J Baumgartner in hb MA MA-Hepat 1108 > see fr2325.pdf Basionym: Distribution: Schiffner 1936 Pando 2000 = Frullania jackii 2561 Krit Bemerk europ Leberm, ser 21: 14-15 [Hepat europ exsicc, nr. 1023pp] 1273 Herbario de Criptogamia, Madrid (MA) http://161.111.170.202/herb/asp/dos.asp?conque=MAHepat&fnhe=1108&fadic=1 Gottsche 1863 jackii var. chichibuensis Kamim. 1961 Typus: Japan. Saitama Pref, Mts. Chichibu, Shiraiwa Peak, 1800-1900m 16.7.1953, M. Inoue 2714 in hb TNS [holotypus] Basionym: Distribution: Japan, Taiwan var nov Kamimura Bonner Lai & Wang-Yang 1961 1965 1976 199 655 33 - syn nov Piippo 1990 N.N. [National Science Museum 2000 Tokyo] 521 995 J Hatt Bot Lab 24: 53, fig 19(18-32), map 5 Index Hepat 5: 338 Taiwania 21 (2): 168 [Index to Taiwan] [as Fr. ssp. japonica var chichibuensis KAMIM.] J Hatt Bot Lab 68: 42 [annotated catalogue of China] http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] = Frullania davurica var. chichibuensis (Kamim.) S.Hatt. 1976 Page 222 jackii var. grandigaleata Müller 1938 Typus: Germany. Baden Württemberg, Seebuck (Nordwand) am Feldberg 1937 Herzog or 1938 Herzog & Müller Basionym: Distribution: var nov Müller 1939 - fo nov Müller in Rabenhorst Bonner Grolle 1958 1965 1970 Hattori 1976 = Frullania jackii 642 Mitt bad Landesver Naturkunde Naturschutz NF 3 (29/30): 439-440 [Feldberg, Black Forest in Germany] 1028 Krypt Flora (3ed) 6, 2: 1249-1250 655 Index Hepat 5: 338 615 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 314 [fo nov = fo. depauperata] 67 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21 [as var., Fr. davurica ssp. jackii var. grandigaleata] fo. depauperata Grolle 1970 jackii var. grandigaleata Müller 1908 [nom inval] Basionym: Distribution: var nov Müller 1908 787 Mitt bad bot Ver 225: 189-204 = Frullania jackii var. grandigaleata Müller 1938 Frullania jackii fo. depauperata Grolle 1970 jackii var. japonica (Sande Lac.) S.Hatt. 1957 Basionym: Distribution: var nov Hattori Kamimura Bonner Kodama 1957 1961 1965 1972 Choe Yamada & Iwatsuki 1980 2006 = Frullania jackii ssp. japonica Frullania davurica 858 199 655 1375 J Hatt Bot Lab 18: 117 J Hatt Bot Lab 24: 52 [syn sub Fr jackii ssp japonica] Index Hepat 5: 338 Spec Publ Osaka Mus Nat Hist 4: 126, fig 62(g-p), map p. 242 [Central Japan] 1171 Illust Fl & Fauna Korea 24: 734 [valid taxon; South Korea] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] (Sande Lac.) S.Hatt. 1959 Hampe ex Gottsche, Lindenb. & Nees 1845 jackii var. rotundata S.W.Arnell 1898 Basionym: Distribution: var nov - syn nov Arnell Schiffner Hattori = Frullania davurica 1898 1936 1976 1454 Rev Bryol 25 (1): 1 2561 Krit Bemerk europ Leberm, ser 21: 16 [Hepat europ exsicc, nr. 1027] 67 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21 [footnote] Hampe ex Gottsche, Lindenb. & Nees 1845 Page 223 jackii ssp. japonica (Sande Lac.) S.Hatt. 1959 Basionym: Frullania curvirostris Colenso 1889 [non Jack & Steph. 1894] Distribution: China (Anhui, Guizhou), Korea (South) comb nov var nov - syn nov Hattori Kamimura Kamimura Hong Bonner Hong Iwatsuki & Mizutani Choe & Yamada Lai & Wang-Yang Hattori Choe Wu in Li, Gao, Wu et al. Liu Li Kuo & Chiang Piippo Lin & Wu Yamada & Choe Peng, Enroth, Koponen & Piippo Cai = Frullania davurica jackii ssp. retusa Kamim. 1952] 1959 1961 1961 1962 1965 1966 1972 1974 1976 1976 1980 1985 1985 1985 1988 1990 1994 1997 2000 2002 857 199 199 655 1107 966 405 33 67 1171 1169 2801 948 521 1229 717 776 1192 J Hatt Bot Lab 21: 128 [Japan] J Hatt Bot Lab 24: [aus 717] J Hatt Bot Lab 24: 52-53, fig 19(1-17), map 5 [aus 717] Index Hepat 5: 338 Bryologist 69 (1): 411 [South Korea] Colour Illust Bryo Japan, Osaka: 332, pl 45 [716] Misc Bryol Lichen 6 (9): 161 [checklist Korea] Taiwania 21 (2): 168 [Index to Taiwan] Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21 [syn sub Fr. davurica] Illust Fl & Fauna Korea 24: 734 [valid taxa; South Korea] Bryoflora Xizang (Tibet), Science Press, Beijing: 525, fig 228(8-10) Invest Stud Nat 5: <133-155> [aus 698] [aus 717] Taiwania 33: 12 [Index of Taiwan] J Hatt Bot Lab 68: 42 [annotated catalogue of China] Chenia 2: 83 [China, Guizhou Prov.] J Hatt Bot Lab 81: 287 [Korean Peninsula] Hikobia 13: 196 [China: Hubei Prov] Chenia 7: 151 [China, Anhui Prov.] Hampe ex Gottsche, Lindenb. & Nees 1845 Kamim. 1952 ex Yamada & Iwatsuki 2006 [nom not mentioned by Basionym: Distribution: Kamimura Kamimura Hattori Yamada & Iwatsuki 1952 1961 1976 2006 873 199 67 1470 Contrib Fl Hepat Shikokuensem, Kochi: 125 [vide #1470 p. 30] J Hatt Bot Lab 24: [= Fr jackii ssp japonica ??] Bull Natn Sci Mus, ser B (Bot) 2(1): 7-22 ???? J Hatt Bot Lab 99: 30 [jackii ssp retusa = davurica - Catalog of Japan] = jacquinoti Typus: Thaiti Gottsche in Bescherelle 1898 sd, Jacquinot, Vernier, Dr. Savaier Basionym: Distribution: spec nov Gottsche in Bescherelle = Frullania jacquinotii jaegüs 1898 871 J Bot Morot 12: 138 Gottsche 1910 Gottsche ex Pando 2000 [nom herb in hb MA MA-Hepat 2229-1] Typus: Austria. Steiermark, Schladming, 1100 m.s.m. 1880, J. Breidler in hb MA MA-Hepat 2229-1 ("hb Caseres Gil. Caja VIII,4") > see fr2326.pdf Basionym: Distribution: Pando 2000 1273 Herbario de Criptogamia, Madrid (MA) http://data.gbif.org/occurrences/29494345/ = Page 224 jamaicensis Steph. 1911 Typus: Jamaica. sd, Börgesen in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn 1911 1928 1928 686 242 244 Bonner Stephani 1965 1985 655 928 = Jubula pennsylvania ssp. bogotensis jamesoni Spec Hepat 4: 574 Rev Bryol NS 1: 120 Ann Crypt exot 1 (2): 213-218 [syn sub Jubula hutchinsiae ssp. bogotensis] Index Hepat 5: 339 Stephani Icon Hepat Inedit [Homotropantha] D- 4, microfiche 54 fig 3708 ["Jamaica, leg Börgesen"] (Steph.) Guerke 1978 ex Spruce 1884 [nom inval] Basionym: Distribution: Spruce 1884 243 Trans Proc Bot Soc Edinburgh 15: 21 = japonica Typus: [auct. non Sande Lac.] Verd. in Hand.-Mazz. 1930 sd, Mazzetti 6570 & 7829 Basionym: Distribution: Verdoorn in Handel-Mazzetti Hattori & Lin = Frullania davurica japonica 1930 1985 550 505 Symb Sinicae 5: 37-38 [Mazzetti-6570, Mazzetti-7829] J Hatt Bot Lab 59: 132 [syn sub Fr. davurica] Hampe ex Gottsche, Lindenb. & Nees 1845 [auct. non Sande Lac.] Verd. in Hand.-Mazz. 1930 Basionym: Distribution: auct - syn nov Verdoorn in Handel-Mazzetti Hattori & Lin = Frullania schensiana 1930 1985 550 505 Symb Sinicae 5: 37 [Handel-Mazzetti 11125] J Hatt Bot Lab 59: 143 C.Massal. 1897 Page 225 japonica Sande Lac. 1864 Typus: Japan sd, Siebold sn in hb L L0059226 - 910287487 [hb VON SIEBOLD) [holotypus] Basionym: Distribution: China, Korea, Japan [Honshiu, Shikoku, Kiushiu], Taiwan spec nov comb nov Sande Lacoste Mitten Stephani Inoue Yoshinaga Sawada 1864 1891 1897 1900 1906 1909 842 582 213 802 927 930 Stephani Iishiba Verdoorn in Handel-Mazzetti Verdoorn 1910 1912 1930 1932 686 888 550 232 Horikawa Schiffner Horikawa in Asahina Kamimura Hattori Hattori Hong Hong & Yoo Kamimura Hong Hattori, Hong & Inoue Kitagawa Choe 1934 1936 1939 1952 1952 1957 1960 1961 1961 1962 1962 1973 1980 34 277 2872 873 529 858 Wu in Li, Gao, Wu et al. 1985 1169 Stephani 1985 928 Kuo & Chiang Piippo Yamada & Choe Yamada & Iwatsuki 1988 1990 1997 2006 948 521 717 1470 199 876 404 1171 Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 311-312 [Japan] Trans Linn Soc London, ser 2, 3: 202 [Japan] Bull Herb Boissier 5: 78 [Japan] Bot Mag Tokyo 14 (no 162): 180 Bot Mag Tokyo 20: 53 [Japan] Bot Mag Tokyo 23 (no 268): 178-183, fig 5 [key to Japanese species, in Japanese] Spec Hepat 4: 400 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Symb Sinicae 5: 37-38 [China] Ann Bryol 5: 125 [Hepat select crit, ser 3 & 4 no 159; Japonia] & 138 [Fr. japonica not identical with Fr. jackii] J Sci Hiroshima Univ, ser B, div 2, 2: 237 [Taiwan] Österr Bot Zeitschr 85 (1): 18-19 [jackii & japonica = jackii] Nippon Inkwasyokubutu Dukan: [aus 717] Contrib Fl Hepat Shikokuensem, Kochi: 124-125 J Hatt Bot Lab 8: 29 J Hatt Bot Lab 18: 117 [comb nov as Fr. jackii var japonica] [aus 717] Hikobia 2: <244-252> [aus 717] J Hatt Bot Lab 24: 52 [syn sub Fr jackii ssp japonica] [aus 717] J Hatt Bot Lab 25: 284 [Korea] Misc Bryol Lichen 6 (6): 102 Illust Fl & Fauna Korea 24: 734 [sub Fr jackii ssp japonica; South Korea] Bryoflora Xizang (Tibet), Science Press, Beijing: 525 [sub Fr jackii ssp japonica] Stephani Icon Hepat Inedit [Galeiloba] C-238, microfiche 53 fig 3658 ["Japan, Mioshi legit"] Taiwania 33: 12 [Index of Taiwan] J Hatt Bot Lab 68: 42 [annotated catalogue of China] J Hatt Bot Lab 81: 287 [Korean Peninsula] J Hatt Bot Lab 99: 30 [Catalog of Japan] = Frullania jackii var. japonica (Sande Lac.) S.Hatt. 1957 Frullania jackii ssp. japonica Frullania davurica japonica ssp. emarginata (Sande Lac.) S.Hatt. 1959 Hampe ex Gottsche, Lindenb. & Nees 1845 Kamim. 1952 Typus: Japan. Prov Tosa, Taisyo-machi -.2.1930, T. Wada 1378 in hb Kamim. [holotypus] Basionym: Distribution: Japan - syn nov Kamimura Hattori & Noguchi Kamimura Bonner Hattori Choe Yamada & Iwatsuki 1952 1960 1961 1965 1976 1980 2006 = Frullania jackii ssp. japonica Frullania davurica 873 965 199 655 67 1171 1470 Contrib Fl Hepat Shikokuensem, Kochi: 125, fig 25 Index spec Typ herb Japon, Bryophyta: 17 J Hatt Bot Lab 24: 52 [syn sub Fr jackii ssp japonica] Index Hepat 5: 339 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21 [syn sub Fr. davurica] Illust Fl & Fauna Korea 24: 734 [sub Fr jackii ssp japonica] J Hatt Bot Lab 99: 30 [Catalog of Japan] (Sande Lac.) S.Hatt. 1959 Hampe ex Gottsche, Lindenb. & Nees 1845 Page 226 javanica Steph. 1910 Typus: Indonesia. Java coll ignot in hb G G00067113 - n° SIB 125478/1 - 14574 (ex hb M) Basionym: Distribution: Philippines spec nov - syn nov Stephani Goebel Verdoorn Bonner Hattori Miller, Whittier & Whittier Stephani 1911 1928 1929 1965 1974 1983 1985 686 234 245 655 0 609 928 Spec Hepat 4: 464 Ann Jard Bot Buitenzorg 39: 49 [Brutkörper, gemmulae] Ann Bryol 2: 134 [syn sub Fr. squarrosa var planescens] Index Hepat 5: 339 [Java] note in hb G: "syn of F ericoides" VIII, 1974 Prod Flora Hepat Polynesiae: 131 Stephani Icon Hepat Inedit [Galeiloba] C-145, microfiche 52 fig 3539 [from type; "Java, Herb München"] = Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides jishibae (Nees) Mont. 1839 Steph. 1924 Typus: Japan. Sendai 10.11.1907, E. Uematsu 81 in hb G Basionym: Distribution: China (Yunnan), Japan spec nov Stephani Verdoorn in Handel-Mazzetti Horikawa Kamimura Bonner Stephani 1924 1930 1934 1961 1965 1985 Piippo Yamada & Iwatsuki 1990 2006 = Frullania japonica 687 550 34 199 655 928 Spec Hepat 6: 540-641 Symb Sinicae 5: 37 [syn sub Fr. japonica] J Sci Hiroshima Univ, ser B, div 2, 2: 237 [syn sub Fr. japonica] J Hatt Bot Lab 24: 52 [syn sub Fr jackii ssp japonica] Index Hepat 5: 340 Stephani Icon Hepat Inedit [Galeiloba] C-239, microfiche 53 fig 3659 ["Japonia, loco Negano, 1908, Uematsu 118"] 521 J Hatt Bot Lab 68: 42 [annotated catalogue of China] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] Sande Lac. 1864 Frullania jackii var. japonica (Sande Lac.) S.Hatt. 1957 Frullania jackii ssp. japonica Frullania davurica johannensis (Sande Lac.) S.Hatt. 1959 Hampe ex Gottsche, Lindenb. & Nees 1845 Steph. 1911 Typus: Comores. Anjouan "Ins. Johanna" 1875, Hildebrandt in hb G 13032 Basionym: Distribution: Comores spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle 1995 Wigginton 2002 Wigginton Wigginton 2004 2004 686 655 185 928 Spec Hepat 4: 477 Index Hepat 5: 340 Bull Jard Bot Nat Belg 46 (1/2): 112 Stephani Icon Hepat Inedit [Thyopsiella] F- 7, microfiche 56 fig 3880 ["Insula Johanna, Comoren, Hildebrandt legit"] 250 Bryophyt Bibl 48: 68 [type syn sub Fr. serrata var. pertenuis, bur syn to Fr. serrata] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 221 [syn sub Fr serrata, West Africa] = Frullania serrata var. pertenuis (Nees) Gottsche 1845 Page 227 josephinae Gottsche ex Steph. 1911 [nom herb] Typus: Cuba. Josephina sd, Wright 296 in hb G Basionym: Distribution: spec nov - syn nov - syn nov Gottsche ex Stephani Bonner Stotler Stotler Stephani = Frullania intumescens 1911 1965 1969 1970 1985 686 655 4761 194 928 Spec Hepat 4: 498 Index Hepat 5: 340 Nova Hedwigia 18: 444 [syn sub Fr. intumescens] The genus Frullania, separata: 444 [syn sub Fr. intumescens] Stephani Icon Hepat Inedit [Thyopsiella] F- 64, microfiche 57 fig 3965 ["Cuba, Josephina (Wright leg)"] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 josephinae var. compacta Gottsche ex Bonner 1965 [nom herb] Typus: Cuba sd, Wright 295 in hb G Basionym: Distribution: Cuba Bonner 1965 = Frullania intumescens julacea 655 Index Hepat 5: 341 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Spruce 1889 Typus: Paraguay. Caaguazú sd, Balansa 3723 in hb G Basionym: Distribution: Brazil, Paraguay spec nov - syn noc Spruce 1889 Stephani Schiffner & Arnell Bonner Yano Stephani 1910 1964 1965 1984 1985 Reiner Yano Gradstein & Costa 1988 1999 2003 = Frullania ericoides 1122 Bull Soc Bot France 36 (Suppl) [Congres de Botanique]: CXC-CXCI [190-191] 686 Spec Hepat 4: 395 198 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 91 [Brazil] 655 Index Hepat 5: 341 501 J Hatt Bot Lab 56: 505 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Galeiloba] C- 65, microfiche 50 fig 3426xx [from type; "Paraquay, leg Balansa 3723"] 1043 Bol Soc Argentina Bot 25 (3-4): 317 [syn sub Fr. ericoides] 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] 984 Mem New York Bot Gard 87: 86 [Brazil] [may be ? syn sub Fr ericoides] (Nees) Mont. 1839 Page 228 junghuhniana var. minutissima Schiffn. ex Verd. 1929 Typus: Indonesia. Java, Cibodas "Urwald von Tjibodas" -.1.1895, Schiffner 1199 Basionym: Distribution: Indonesia (Java), ?Samoa (Tutuila) var nov - syn nov Schiffner ex Verdoorn Verdoorn Bonner Hattori Hattori Hattori Hattori Hattori Hattori & Mizutami Grolle & Piippo 1929 1930 1965 1975 1975 1975 1976 1982 1982 1984 245 293 655 65 66 547 80 422 495 143 Ann Bryol 2: 147 Ann Bryol Suppl 1: 135 Index Hepat 5: 341 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (3): 112 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 148 J Hatt Bot Lab 39: 291-292 [Sumatra, New Guinea] J Hatt Bot Lab 40: 485 Misc Bryol Lichen 9 (6): 123-124 [F. junghuhnia var. tenella] J Hatt Bot Lab 52: 447 Acta Bot Fennica 125: 20 [Cat. West Melanesia] = Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 junghuhniana var. minutissima fo. monoica (S.Hatt.) S.Hatt. 1976 Basionym: Frullania perversa var. monoica S.Hatt. 1974 Distribution: comb nov Hattori Hattori, Thaithong & Kitagawa Hattori & Mizutani Hattori 1976 1977 1982 1982 80 480 495 495 J Hatt Bot Lab 40: 485 J Hatt Bot Lab 43: 445 [Thailand] J Hatt Bot Lab 52: 447 J Hatt Bot Lab 51: 234 = Frullania junghuhniana var. tenella fo. monoica (S.Hatt.) S.Hatt. 1986 junghuhniana var. monoica (S.Hatt.) S.Hatt. 1975 Basionym: Frullania perversa var. monoica S.Hatt. 1974 Distribution: comb nov - comb nov Hattori Hattori Grolle & Piippo 1975 1982 1984 547 495 143 J Hatt Bot Lab 39: 292 J Hatt Bot Lab 51: 234 Acta Bot Fennica 125: 20-21 [Cat. West Melanesia] = Frullania junghuhniana var. minutissima fo. monoica (S.Hatt.) S.Hatt. 1976 Frullania junghuhniana var. tenella fo. monoica (S.Hatt.) S.Hatt. 1986 junghuhniana var. perversa (Steph.) Horikawa 1943 Basionym: Frullania perversa Distribution: comb nov - comb nov Horikawa Schuster Hattori = Neohattoria perversa Steph. 1894 1943 1970 1973 1283 Acta Phytotax Geobot 13: 213 761 J Hatt Bot Lab 33: 288 549 J Hatt Bot Lab 37: 57 [sub Fr. perversa] (Steph.) R.M.Schust. 1970 Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 Page 229 junghuhniana ssp. denticulilobula S.Hatt. 1972 Basionym: Distribution: ssp nov Hattori Hattori Hattori Hattori Hattori Grolle & Piippo Hattori & Piippo Mizutani, Hasegawa & Iwatsuki 1972 1973 1974 1975 1982 1984 1986 2009 539 542 546 547 495 143 557 2112 J Hatt Bot Lab 36: 438-440, 17 J Hatt Bot Lab 37: 149 [New Guinea] J Hatt Bot Lab 38: 241-242 J Hatt Bot Lab 39: 291 [sub Fr. junghuhniana var minutissima] J Hatt Bot Lab 51: 233 Acta Bot Fennica 125: 21 [Cat. West Melanesia] Acta Bot Fennica 133: [= nom inval, type not indicated] Type sepecimens Hatt Bot Lab (NICH) 28: Syntype. New Guinea, Central Highlands, Mt. Wilhelm. 10700-11000 ft. 05 June , 1967. R. M. Schuster 67-6040/a (NICH 309916). Other 6 syntypes: NICH 309917, 314843, 314847,314848,314849&314850. = Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 junghuhniana ssp. wattsii Typus: Australia. Fitzroy Falls S.Hatt. 1972 -. 10.1908, leg w.w. Watts 929 in hb G 15007 Basionym: Distribution: Australia spec nov - syn nov Hattori Hattori Hattori Hattori & Mizutami Scott & Bradshaw Carthy McCarthy = Frullania rostrata jungneri 1972 1979 1982 1982 1986 2003 2006 538 485 495 495 56 1125 1385 J Hatt Bot Lab 36: 130-132, fig 17 J Hatt Bot Lab 45: 352-353 [orth var as "wattii"] J Hatt Bot Lab 51: 232 J Hatt Bot Lab 52: 447 Brunnonia 8 (1): 35 [Australia] Flora Australia Suppl Ser 21: 46 [sub Fr rostrata] Checklist Australian Liverworts [Australia, syn sub Fr rostrata] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Steph. 1911 Typus: Cameroons [Kamerun] , 8000 ft sd, Jungner sn hb STEPHANI 244, in hb G Basionym: Distribution: Cameroon spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania angulata 686 655 185 928 Spec Hepat 4: 482 Index Hepat 5: 341 Bull Jard Bot Nat Belg 46 (1/2): 71 Stephani Icon Hepat Inedit [Thyopsiella] F- 8, microfiche 56 fig 3881 ["Kamerun, leg Jungner, 8000f, 244"] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Mitt. 1863 Page 230 kagoshimensis Steph. ex Sawada 1909 [nom nud] Basionym: Distribution: Sawada 1909 = Frullania kagoshimensis 930 Bot Mag Tokyo 23 (no 268): 178-183, fig 1, 1' [key to Japanese species, in Japanese] Steph. 1910 kagoshimensis var. minor Kamim. 1961 Basionym: Distribution: China, Taiwan, Japan Ryukyu (Okinawa) - syn nov Kamimura Bonner Shin Lai & Wang-Yang Hattori Yamada, Yuzawa, Ogasawara & Lai Hattori & Lin Kuo & Chiang Piippo Furuki & Mizutani Yuzawa 1961 1965 1970 1976 1980 1981 199 655 302 33 958 949 1985 1988 1990 1994 2001 505 948 521 1331 1660 Furuki Yamada & Iwatsuki 2006 2006 = Frullania osumiensis kalawensis J Hatt Bot Lab 24: 45-47, fig 11(19-25), map 4 Index Hepat 5: 342 Sci Rep Kagoshima Univ 19: 37 [Ryukyu: Okinawa] Taiwania 21 (2): 168 [Index to Taiwan] Bull Natn Sci Mus Tokyo, ser B (Bot), 6 (1): 38 [new to Formosa] Quart J Taiwan Mus 34 (1,2): 114 [Taiwan] J Hatt Bot Lab 59: 137 Taiwania 33: 12 [Index of Taiwan] [syn sub Fr. kagoshimensis] J Hatt Bot Lab 68: 42 [annotated catalogue of China] Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] Natural Environmental Sci Res 14: 32, fig 26(a-m) [Japan, syn nov sub Fr osumiensis] 1503 Hikobia 14: 407 [Izu Isls., Northern Izu-Mariana Arc] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] (S.Hatt.) S.Hatt. 1956 Svihla 1968 Typus: Burma. Southern Shan States, Kalaw, about 3800ft 22.4.1953, Svihla 1594 in hb WTU Basionym: Distribution: spec nov - syn nov Svihla Bonner Hattori & Thaithong 1958 1965 1978 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania shanensis 389 655 127 Bryologist 61: 373-376, fig 1-17 Index Hepat 5: 342 J Jap Bot 53 (5): 132 footnote [Fr. kalawensis is hygrophilous form of Fr. shanensis] 2112 Type sepecimens Hatt Bot Lab (NICH) 28: Isotype. Burma, Shan State. 3800 ft. On bark. 22 Apr. 1953. R. D. Svihla 1954 (NICH 218380). Svihla 1957 Page 231 kaluensis Steph. 1924 Typus: Hawai. Kalua -.6.1909, Faurie 430 holotypus: in hb G-22740; isotypus in hb BR Basionym: Distribution: Hawaii spec nov Stephani Verdoorn 1924 1930 - syn nov Verdoorn 1930 Miller Miller Bonner Miller, Whittier & Whittier Stephani 1953 1963 1965 1983 1985 Bapna & Kachroo So & Wang 2000 2006 = Frullania sandvicensis kamimurae 687 293 Spec Hepat 6: 541-542 Ann Bryol suppl 1: 37 [syn sub Fr. sandvicensis; and Fr sandvicensis may be syn sub Fr squarrosa] 551 Nederl Kruidk Arch Jrg 1930 (2): 161 [syn nov sub Fr. sandvicensis AONGSTRÖM, dioicous plants] 349 Bryologist 56: 45 [sub Fr squarrosa] 834 Ark Bot, ser 2, 5: 518 [sub Fr. squarrosa] 655 Index Hepat 5: 342 609 Prod Flora Hepat Polynesiae: 130 928 Stephani Icon Hepat Inedit [Galeiloba] C-146, microfiche 52 fig 3540 [from type; "Hawai, Kalua, Faurie 1909 No 430"] 712 Hepaticology in India II: 170 [sub Fr. ericoides] 1466 J H B L 100: 428 [sub Fr sandvicensis] Ångstr. 1872 Inoue 1966 Basionym: Distribution: nom nov - syn nov Inoue Inoue Lai & Wang-Yang Hattori & Lin 1966 1972 1976 1985 Kuo & Chiang Piippo Yamada & Iwatsuki 1988 1990 2006 = Frullania tamarisci ssp. obscura 1703 123 33 505 J Jap Bot 41 (1): 15 [nom nov pro Fr pendula Kamim. 1952} J Jap Bot 47 (11): 349-350 Taiwania 21 (2): 168 [Index to Taiwan] J Hatt Bot Lab 59: 161 [syn sub F. tamarisci subsp. obscura (Verd.) Hatt] 948 Taiwania 33: 12 [Index of Taiwan] 521 J Hatt Bot Lab 68: 42 [annotated catalogue of China] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata karstenii (Reinw., Blume & Nees) Mont. 1842 Schiffn. 1893 Typus: Amboina, Wawani Hila sd, G. Karsten Basionym: Distribution: Indonesia (Ambon) spec nov - syn nov Schiffner 1893 572 Schiffner Stephani Verdoorn Verdoorn Bonner Stephani 1898 1911 1929 1930 1965 1985 831 686 245 293 655 928 Nova Acta K Leop-Carol Deutsch Akad Naturf Halle 60 (2): 223-225, pl 6(1-27) Conspect Hepat Archip Indici, Batavia: 330 Spec Hepat 4: 539 Ann Bryol 2: 139 Ann Bryol Suppl 1: 97, 101 Index Hepat 5: 342 Stephani Icon Hepat Inedit [Thyopsiella] F-144, microfiche 59 fig 4080 ["Amboina, Dr Karsten, Copie nach Schiffner"] = Frullania apiculata var. goebelii Schiffn. 1893 Page 232 kehdingiana Steph. 1894 Typus: Indonesia. Sumatra sd, Kehding, sn. in hb G Basionym: Distribution: Andaman Isl [India], Malaysia (Malay Pen), Indonesia (Sumatra), New Caledonia, Fiji spec nov - syn nov Stephani Schiffner Stephani Verdoorn Verdoorn Bonner Stephani 1894 1898 1911 1928 1930 1965 1985 = Frullania intermedia khedingiana 228 831 686 242 293 655 928 Hedwigia 33: 150-151 ["khedingiana"] Conspect Hepat Archip Indici, Batavia: 330 ["khedingiana"] Spec Hepat 4: 577 [kehdingiana] Rev Bryol NS 1: 120 [syn sub Fr. intermedia] Ann Bryol Suppl 1: 161 Index Hepat 5: 343 Stephani Icon Hepat Inedit [Homotropantha] D-25, microfiche 54 fig 3741 ["Sumatra, Kehding"] - 3742 (Reinw., Blume & Nees) Nees 1845 Steph. 1894 [orth var] Basionym: Distribution: Stephani 1894 = Frullania kehdingiana Hedwigia 33: 150-151 ["khedingiana"] Steph. 1894 Frullania intermedia kilimandjarica 228 (Reinw., Blume & Nees) Nees 1845 Vanden Berghen 1953 Typus: Tanganyika: Kilimandjaro, 3225 m 26.6.1948, O. Hedberg 1373 b in hb S-PA Basionym: Distribution: Kenya, Tanganyika, Uganda spec nov - syn nov Vanden Berghen Arnell Bonner Vanden Berghen Wigginton 1953 1956 1965 1976 2002 Wigginton 2004 Wigginton 2004 = Frullania apicalis kilimanjarica 446 836 655 185 1012 Svensk Bot Tidskr 47 (2): 267-269, fig. 1[a-d) Ark Botanik 3 (16): 531 [East African mountains] Index Hepat 5: 343 Bull Jard Bot Nat Belg 46 (1/2): 52 Trop Bryol Res Rep 2: 28 [as kilimanjarica; http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [as kilimanjarica; http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217, fig 137(B,N,P,U) from type [syn sub Fr apicalis, West Africa] Mitt. 1879 Vanden Berghen by Wigginton 2002 [orth var] Basionym: Distribution: Wigginton 2002 Wigginton 2004 = Frullania kilimandjarica Frullania apicalis 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Vanden Berghen 1953 Mitt. 1879 Page 233 kinangopii Gola 1916 Typus: Kenya. Mt. Kinangop, 4400 sm. 26 VII 1908 (P. Savio) Basionym: Distribution: Kenya spec nov syn nov ? Gola Vanden Berghen Wigginton = Frullania apicalis kirkii 1916 1976 2004 3469 Mem Reale Accad Sci Torino, ser 2. 65: 9-10, fig 29-32 [Kenya] 185 Bull Jard Bot Nat Belg 46 (1/2): 13 [perhaps syn of Fr. apicalis] 1312 Trop Bryol Res Rep 5: 30 [perhaps syn of Fr apicalis] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Mitt. 1879 Steph. 1894 Typus: New Zealand. The Bluff Hill sd, Prof. Kirk 526 in hb G 20584 Basionym: Distribution: New Zealand spec nov - syn nov Stephani Stephani Rodway 1894 1910 1916 228 686 914 Martin 1946 1423 Martin 1946 1423 Hodgson 1949 203 Bonner Hamlin Hattori Stephani 1965 1972 1983 1985 655 659 499 928 Glenny Glenny & Malcolm 1998 2005 1412 1410 = Frullania patula kizarensis Hedwigia 33: 145 Spec Hepat 4: 407 Papers Proceed Roy Soc Tasmania for 1916: 135 [syn sub Fr. proboscifora] [Tasmania] Trans Roy Soc New Zealand 76 (1): 49 [New Zealand, South Isl: Arthur Pass NP] Transact Roy Soc NZ 76(1): 49 [New Zealang; Arthur Pass National Park] Trans Roy Soc New Zealand 77 (3): 380 [may be Fr. scandens, according Hattori 1983] Index Hepat 5: 343 Rec Dominion Mus 7(19): 262 [New Zealand] J Hatt Bot Lab 54: 152-153, fig 59 [from type] Stephani Icon Hepat Inedit [Galeiloba] C-265, microfiche 54 fig 3688 ["N Zelandia, The Bluff Hill"] Tuhinga 10: 132 [sub Fr patula] ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr patula]] Mitt. 1855 Steph. 1911 Typus: Africa: Usambara, Kizara sd, Holst 2339 pp [sub Fr. varia] in hb G [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Demaret & Vanden Berghen Bonner Stephani 1911 1950 1965 1985 686 60 655 928 Wigginton 2002 1012 Wigginton Wigginton Uribe 2004 2004 2008 1312 1172 1513 = Frullania angulata Spec Hepat 4: 594 Bull Jard Bot Etat Bruxelles 20: 5 [syn sub Fr. angulata] Index Hepat 5: 343 Stephani Icon Hepat Inedit [Meteriopsis] E- 4, microfiche 55 fig 3768 ["Africa or (Kizara in Usambara) Holst 2339 ex parte"] Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Mitt. 1863 Page 234 knightiana Steph. 1924 Typus: Australia. Kings Island 1882. E. Spong in hb G 18218 Basionym: Distribution: Australia spec nov - syn nov Stephani Bonner Hattori Hattori Stephani 1924 1965 1979 1983 1985 Stephani 1985 Scott & Bradshaw Carthy McCarthy 1986 2003 2006 = Frullania falciloba kochiensis 687 655 485 499 928 Spec Hepat 6: 542 Index Hepat 5: 344 J Hatt Bot Lab 45: 346, fig 17(a-h) from type [syn sub Fr. falciloba] J Hatt Bot Lab 54: 154 Stephani Icon Hepat Inedit [Galeiloba] C-214, microfiche 53 fig 3627 ["Australia, Kings Island, leg E Spong 1882"] 928 Stephani Icon Hepat Inedit [Galeiloba] C-261ab, microfiche 54 fig 3682 ["olim Frull knightiana, N Zealand, Knight 601"] - 3683 ["Frull falcata Stz, Frull knightiana St, Frull scabriseta Col, Trans NZ Inst vol 19, Hepa 1382, ob scandens?"] 56 Brunnonia 8 (1): 35 [Australia] 1125 Flora Australia Suppl Ser 21: 43 [sub Fr falciloba] 1385 Checklist Australian Liverworts [Australia, syn sub Fr falciloba] Lehm. 1844 Steph. 1910 Typus: Japan. Shikoku, Tsurugizan, -.6.1900 U Faurie 650 in hb G, in hb NICH, in hb TNS [isotypus] in hb HIRO [isosyntypus?] Basionym: Distribution: Japan (Honshu, Shikoku, Kyushu), Ryukyu (Ishigaki, Kume) spec nov - syn nov - comb nov Stephani Iishiba Hattori Kamimura 1910 1912 1951 1952 686 888 528 873 Hattori & Noguchi Bonner Shin Choe Gao & Chang 1960 1965 1970 1980 1981 965 655 302 1171 1021 Aur & Zhang Inoue N.N. [National Science Museum Tokyo] NN 1985 1987 2000 1022 1155 995 2003 Yamada & Iwatsuki 2006 1397 Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] 1470 J Hatt Bot Lab 99: 30 [Catalog of Japan] = Frullania pedicellata Spec Hepat 4: 400 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] J Hatt Bot Lab 5: 67 [syn sub Fr. pedicellata] Contrib Fl Hepat Shikokuensem, Kochi: 130 [comb nov; as Fr. pedicellata fo. kochiensis] Index spec Typ herb Japon, Bryophyta: 17 [isotypus] Index Hepat 5: 344 Sci Rep Kagoshima Univ 19: 37 [Ryukyu: (Ishigaki, Kume) Illust Fl & Fauna Korea 24: 735 [sub Fr pedicellata; South Korea] Fl Hep Chinae bor-orient: 152 [syn sub Fr. pedicellata, #1612 or #2110] Fl Heilongjiangensis 1: 163 [sub Fr pedicellata] Index type specimens in Nat Sci Mus, Tokyo: 13 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] fo. kochiensis (Steph.) Kamim. 1952 Frullania pedicellata Steph. 1897 Page 235 kochiensis Steph. ex Yoshinaga 1903 [nom nud] Typus: Japan. Sikoku, Prov Tosa: Kochi Park -.-.1900, T. Yoshinaga 39 in hb TNS [isotypus] Basionym: Distribution: Stephani Stephani in Yoshinaga Sawada 1897 1903 1909 -syn nov Hattori Hattori & Noguchi 1951 1960 - syn nov Kamimura Stephani 1961 1985 Inoue Yuzawa 1987 2001 nom nud = Frullania pedicellata koreana 213 801 930 Bull Herb Boissier 5 (2): 90 [vide #965, but not found in this Bot Mag Tokyo 17 (no 193): 37 [nom nud] Bot Mag Tokyo 23 (no 268): 178-183, fig 10 [key to Japanese species, in Japanese] 528 J Hatt Bot Lab 5: 67 [syn sub Fr. pedicellata] 965 Index spec Typ herb Japon, Bryophyta: 17 [isotypus, should be in in #213, 1897] 199 J Hatt Bot Lab 24: 40 [syn sub Fr kochiensis] 928 Stephani Icon Hepat Inedit [Galeiloba] C-241, microfiche 53 fig 3661 ["Japan, KochoiPark, Yoshinaga 39"] 1155 Index type specimens in Nat Sci Mus, Tokyo: 13 1660 Natural Environmental Sci Res 14: 35 [Japan, syn sub Fr pedicellata] Steph. 1897 Steph. 1911 Typus: Korea. Quen-san -.8.1901, U. Faurie 1047, 1036 in hb KYO (Faurie 1036) [isotypus] in hb HIRO (Faurie 1036) [isotypus] in hb G (Faurie 1036) [isotypus] Basionym: Distribution: Korea spec nov - syn nov Stephani Iishiba Hattori & Noguchi Kamimura Bonner Hattori 1911 1912 1960 1961 1965 1980 Choe Choe Stephani 1980 1983 1985 Hong Yamada & Choe NN 1997 1997 2003 Hong 2003 = Frullania hamatiloba kotohirana 686 888 965 199 655 489 Spec Hepat 4: 468 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Index spec Typ herb Japon, Bryophyta: 17 [isotype] J Hatt Bot Lab 24: 34 [regarded as a variety of Fr hamatiloba] Index Hepat 5: 344 J Hatt Bot Lab 47: 120-121, fig 237(a-h) from isotype [close to or conspecific with hamatiloba] 1171 Illust Fl & Fauna Korea 24: 734 [South Korea] J Sci Educ 15: <147-156> [aus 717] 928 Stephani Icon Hepat Inedit [Galeiloba] C-147, microfiche 52 fig 3541 [from type; "Korea, Faurie 1047"] 693 Lindbergia 22: 138 [Korea] & 139 Annotation #26 717 J Hatt Bot Lab 81: 287 [Korean Peninsula][syn sub of Fr. hamatiloba] 1397 Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] 1416 Lindbergia 28: 138 [excluded from Korean paninsula] Steph. 1910 Steph. ex Bonner 1965 [nom herb] Typus: Japan. Kotohira, Sanuki -.1.1911, S. Okamura 405 in hb G Basionym: Distribution: Japan Bonner Stephani 1965 1985 655 928 Index Hepat 5: 344 Stephani Icon Hepat Inedit [Galeiloba] C-242, microfiche 53 fig 3662 ["japan, Kotohira, O Kamura leg 405"] = Page 236 kunsei Lehm. & Lindenb. by Pinzon, Linares & Uribe-M 2003 [orth var] Basionym: Distribution: Pinzon, Linares & Uribe-M = Frullania kunzei kunzeana 2003 1135 Caldasia 25(2): 300 [Colombia] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Lehm. & Lindenb. by Duss 1904 [orth var] Basionym: Distribution: Duss = Frullania kunzei kunzei 1904 3132 Flore Cryptogamique des Antilles Françaises: 132 [Martinique] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Lehm. & Lindenb. ex Gottsche 1845 [pro syn] Basionym: Distribution: Gottsche in G., L. & N. = Frullania capensis kunzei 1845 2850 Syn Hepat: 449 [syn sub Fr capensis] Gottsche 1845 [auct. non () Lehm. & Lindenb.] Austin 1874 Basionym: Distribution: auct - syn nov Austin Stephani Evans Hattori Kamimura Miller Miller, Whittier & Whittier = Frullania meyeniana 1874 1897 1900 1944 1961 1963 1983 668 214 236 943 199 834 609 Bull Torrey Bot Club 5: 15 [Oahu - Sandwich Isl] Bull Herb Boissier 5: 842[:260] [Hawaii] Trans Connet Acad Arts Sci 10 (1): 402 [syn nov sub Fr. meyeniana] Bull Tokyo Sci Mus 11: 146 [Japan, key to Japanese species] J Hatt Bot Lab 24: 73 [sub F meyeniana] Ark Bot, ser 2, 5: 519 [syn sub Fr. meyeniana] Prod Flora Hepat Polynesiae: 131 Lindenb. 1845 kunzei var. maritima R.M.Schust. 1991 Typus: USA. Florida, Collier Co, Collier-Seminole-State Park, on Lagenaria sd, RMS 76-390 Basionym: Distribution: USA ( FL ) var nov Schuster Schuster Hentschel et alii 1991 1992 2015 1151 J Hatt Bot Lab 70: 144-145 603 Hepat N America 5: 97-99, fig 789(6-8) 4097 Phytotaxa 220 (2): 135 [var nov. - as nom. inval. (ICN Art. 37.7; no herbarium indicated).] = Frullania kunzei var. maritima R.M.Schust. 2015 Page 237 kurzii Steph. 1910 Typus: Indonesia. Java sd, Kurz 419 pp in hb G Basionym: Distribution: Indonesia (Java) spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Stephani 1911 1929 1930 1965 1985 = Frullania grandistipula lacerata 686 245 293 655 928 Spec Hepat 4: 454-455 Ann Bryol 2: 125 [syn sub Fr. grandistipula] Ann Bryol Suppl 1: 43 Index Hepat 5: 345 Stephani Icon Hepat Inedit [Galeiloba] C-148, microfiche 52 fig 3542 [from type; "Java, Kurz 419"] Lindenb. 1845 [non Steph. 1924] Steph. 1911 Typus: Madagascar sd, Forsyth Major 580 in hb G Basionym: Distribution: Madagascar spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania serrata lacerifolia 686 655 185 928 Spec Hepat 4: 481 Index Hepat 5: 345 Bull Jard Bot Nat Belg 46 (1/2): 111 Stephani Icon Hepat Inedit [Thyopsiella] F- 9, microfiche 56 fig 3882 ["Madagascar, Forsyth Major 580"] - 3883 250 Bryophyt Bibl 48: 68 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 221 [syn sub Fr serrata, West Africa] Gottsche 1845 Steph. 1911 Typus: Indonesia. Java &, Amboina ? sd, leg Karsten Basionym: Distribution: spec nov - comb nov Stephani Verdoorn Bonner Stephani = Frullania gracilis 1911 1929 1965 1985 686 245 655 928 Spec Hepat 4: 648-649 Ann Bryol 2: 144, 159 [comb nov] Index Hepat 5: 345 Stephani Icon Hepat Inedit [Diastaloba] B-66, microfiche 49 fig 33083309 ["Amboina, Karsten"] (Reinw., Blume & Nees) Nees 1845 Page 238 laceriloba Steph. 1911 Typus: St. Thomé sd, Fr. Quintas sn in hb G Basionym: Distribution: Annobon, W Africa spec nov - syn nov Stephani Arnell Bonner Vanden Berghen Stephani 1911 1956 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania apicalis 686 847 655 185 928 Spec Hepat 4: 484 Svensk Bot Tidskr 50 (4): 532 [Annobon] Index Hepat 5: 345 Bull Jard Bot Nat Belg 46 (1/2): 52 Stephani Icon Hepat Inedit [Thyopsiella] F- 22, microfiche 56 fig 3900 ["Africa, Ins San Thomé, leg Quintas"] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apicalis, West Africa] Mitt. 1879 laceriloba var. acutifolia S.W.Arnell 1956 Typus: Bioko (Fernando Po), Moka sd, Bystrom 837 in hb UPS Basionym: Distribution: Bioko (Fernando Po) var nov - syn nov Arnell Bonner Vanden Berghen Wigginton 1956 1965 1976 2002 Wigginton Wigginton 2004 2004 = Frullania apicalis lacerostipula 847 655 185 1012 Svensk Bot Tidskr 50 (4): 528 [Fernando Po] Index Hepat 5: 345 Bull Jard Bot Nat Belg 46 (1/2): 52 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217, fig 137(G) from type [syn sub Fr apicalis, West Africa] Mitt. 1879 Steph. in Levier 1906 Typus: China. Prov Schensi, Thae-pei-san -.9.1897, J. Giraldi hb LEVIER 1796, in hb G Basionym: Distribution: spec nov spec nov Stephani in Levier Levier Stephani Bonner 1906 1906 1911 1965 824 824 686 655 - syn nov Hattori 1973 542 Hattori & Lin Piippo 1985 1990 505 521 = Frullania tamarisci ssp. obscura Nuovo Giorn Bot Ital NS 13: 348 Nuovo Giorn Bot Ital NS 13: 348 Spec Hepat 4: 546 [not mentioned as syn] Index Hepat 5: 346 [error: not as syn sub Fr. clavellata in STEPH 1911: 546] J Hatt Bot Lab 37: 121-122, fig 67 [from type] [syn sub Fr. tamarisci subsp obscura] J Hatt Bot Lab 59: 161 J Hatt Bot Lab 68: 42 [annotated catalogue of China] (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 Page 239 laciniaeflora Pearson 1923 Typus: New Zealand. Kahikanui, near Whakepara 8.7.1904, Setchell 50 in hb UC 213701 in hb UC [typus, lost] lost on loan to ? Late Frans Verdoorn 6.viii.1929 at Utrecht; New Zealand Basionym: Distribution: spec nov - syn nov Pearson Hodgson Bonner Hamlin N.N. 1923 1949 1965 1972 2003 = Frullania pycnantha laciniosa Univ California Publ Bot 10 (5): 378-379 Trans Roy Soc New Zealand 77 (3): 366 [syn sub Fr. pycnantha] Index Hepat 5: 346 Rec Dominion Mus 7(19): 262 [New Zealand] http://ucjeps.berkeley.edu/bryolab/moss_types.html [type in hb UC; lost] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Gottsche in Lehm. 1844 Typus: Sri Lanka. Peradenia, in hb NY Basionym: Distribution: spec nov 797 203 655 659 978 / / Indonesia, Java 1.3.1847, Gardner 1426 / Haskarl typus in hb S (B24090, 24091) Gottsche in Lehmann Mitten Sande Lacoste Sande Lacoste in Veth 1844 1861 1864 1884 Schiffner Sawada 1898 1909 Stephani Iishiba Verdoorn Chopra Kamimura 1911 1912 1930 1943 1961 Parihar Hattori 1962 1975 Stephani 1985 Piippo Bapna & Kachroo Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 1990 2000 2002 = Frullania squarrosa Frullania ericoides 589 574 842 807 Pugillus 8: 18 J Linn Soc Bot 5: 119 [Sri Lanka] Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 312 [Celebes] Midden Sumatra, Reizen en Qnderzoekingen der Sumatra Expeditie, 4 (2): 44 [Sumatra] 831 Conspect Hepat Archip Indici, Batavia: 330 930 Bot Mag Tokyo 23 (no 268): 178-183, fig 7, 7' [key to Japanese species, in Japanese] 686 Spec Hepat 4: 458 888 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] 551 Nederl Kruidk Arch Jrg 1930 (2): 205 J Indian Bot Soc 22: 249 [India] 199 J Hatt Bot Lab 24: 17 [Fr laciniosa auct. non GOTTSCHE: LEHMAN... syn sub Fr squarossa] 1584 Univ Allahabad studies, Bot sect 1-2: 19 [India, syn sub Fr squarrosa] 47 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 77 [Ann Bryolout Mitten´s specimen] 928 Stephani Icon Hepat Inedit [Galeiloba] C-149, microfiche 52 fig 3543 ["Java"] 521 J Hatt Bot Lab 68: 42 [annotated catalogue of China] 712 Hepaticology in India II: 169 [syb sub Fr ericoides] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] (Reinw., Blume & Nees) Nees 1845 (Nees) Mont. 1839 Page 240 lacouturei Steph. 1911 Typus: Panama. Chirique sd, F. Helion hb LACOUTURE 421, in hb G Basionym: Distribution: Panama spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 = Frullania mucronata Spec Hepat 4: 499 Index Hepat 5: 346 Nova Hedwigia 18: 446 [syn sub Fr. mucronata] The genus Frullania, separata: 446 [syn sub Fr. mucronata] Stephani Icon Hepat Inedit [Thyopsiella] F- 65, microfiche 57 fig 3966 ["Panama, Frere Hélion, Lacouture 421"] (Lehm. & Lindenb. in Lehm.) Lehm. & Lindenb. 1845 Frullania brasiliensis laevicalyx 686 655 4761 194 928 Raddi 1822 Steph. ex Bonner 1965 [nom herb / orth var] Basionym: Distribution: Bonner 1965 = Frullania levicalyx Index Hepat 5: 347 [vide levicalyx] Steph. 1910 Frullania trinervis laeviperiantha 655 (Lehm.) Drège 1843 X.L.Bai & C.Gao 2000 Typus: China, Yunnan, Ruili 8.7.1977, Xing-jiang Li 89 in hb IFSBH, HKAS [holotypus] Basionym: Distribution: China (Yunnan) spec nov nom inval Bai & Gao Bai Bai Hentschel et alii 2000 2001 2002 2015 = Frullania laeviperiantha laeviscypha 735 1742 1189 4097 Nova Hedwigia 70 (1-2): 135-137, fig 1-9 Guizhou Science 19(4): 39-41, fig 6(1-9) [China, Yunnan] Chenia 7: 11, 25 ["Endemic to Yunnan"], fig 7(1-9) Phytotaxa 220 (2): 136 [Frullania laeviperiantha Bai & Gao 2000 nom. inval. (ICN Art. 37.7; no herbarium indicated). X.L.Bai & C.Gao 2015 Taylor 1846 Typus: USA. Ohio W: Columbus sd, S. Sullivant 104 Basionym: Distribution: spec nov - syn nov Taylor Taylor in G., L. & N. Evans Stephani Frye & Clark = Frullania eboracensis 1846 1847 1897 1910 1947 292 2848 237 686 204 London J Bot 5: 403 Syn Hepat: 778, No 59e Trans Connet Acad Arts Sci 10 (1): 18 Spec Hepat 4: 431 [syn sub Fr eboracensis] Univ Wash Publ Biol 6 (5): 771 [sub Fr eboracensis] Lehm. 1844 Page 241 lagenifera Schwägr. ex Gottsche, Lindenb. & Nees 1845 Typus: Ins. Marianae sd, Gaudichaud sn in hb M [Montagne] (hb BESCHERELLE 219) in hb G G00114992 - n° SIB 240818/5 [isotypus] in hb PC 0103106 [holotypus] Basionym: Distribution: Easter Isl [Chile], Micronesia spec nov - syn nov Schwaegrichen in G., L. & N. Stephani Verdoorn 1845 1910 1930 Kühnemann Herzog in Skottsberg 1937 1942 Kühnemann Miller Hattori Miller, Whittier & Whittier Stephani 1949 1968 1981 1983 1985 Engel Bapna & Kachroo Grolle 1988 2000 2002 2850 Syn Hepat: 420 (No 17) 686 Spec Hepat 4: 430 293 Ann Bryol Suppl 1: 38, footnote [syn sub Fr. squarrosa fo. ericoides] [Guam, leg M.S. Clemens 1911] 1031 Revista Centro Estud Doct Ci Nat, Buenos Aires 1 (3): 167 [Malvinas] 722 The Natural History of Juan Fernandez and Easter Island, 2: 739 [Isla de Pascua: Auf der Innenseite des Kraters Rano Kau, untere Hänge, 125m - no 161, 163, 164, 169, 173, 178] 657 Lilloa 19: 354 [Falkland Isl] 672 Micronesia 4 (1): 74 [syn sub Fr. squarrosa fo. ericoides] 492 J Hatt Bot Lab 49: 371-372 [syn sub Fr. ericoides] 609 Prod Flora Hepat Polynesiae: 131 928 Stephani Icon Hepat Inedit [Galeiloba] C- 95, microfiche 51 fig 3476 [from type; "Insulae Marianae, Gaudichaud, Falkland Inseln"] 111 Fieldiana Bot NS 25: 164 [Mariana, not at Falkland Is. ] 712 Hepaticology in India II: 169 [as lanigera; syn sub Fr. ericoides] 1105 Bryologist 105 (1):126 [Easter Isl] = Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides lanciloba (Nees) Mont. 1839 Steph. 1910 Typus: Japan. Chateu de Ttottori 22.5.1899, Faurie 582 in hb G; in hb KYO [isotypus: Honsyu, Yamakita (8.5.1899, Faurie 389)] Basionym: Distribution: spec nov - syn nov Stephani Iishiba Verdoorn Hattori Hattori & Noguchi Kamimura Bonner Stephani 1910 1912 1930 1944 1960 1961 1965 1985 Bapna & Kachroo Yuzawa Yamada & Iwatsuki 2000 2001 2006 = Frullania hampeana Spec Hepat 4: 404-405 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Ann Bryol Suppl 1: 44 [syn nov sub Fr. hampeana] Bull Tokyo Sci Mus 11: 144 [syn sub Fr. hampeana] Index spec Typ herb Japon, Bryophyta: 17 [isotype] J Hatt Bot Lab 24: 49 [syn sub Fr hampeana] Index Hepat 5: 347 Stephani Icon Hepat Inedit [Galeiloba] C-243, microfiche 53 fig 3663 ["Japonia, Provenzia Tottori, Fauri 582"] 712 Hepaticology in India II: 174 [syn sub Fr. monicera] 1660 Natural Environmental Sci Res 14: 23 [Japan, syn sub Fr monocera] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] Nees 1845 Frullania monocera lanigera 686 888 293 943 965 199 655 928 (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Schwägr. in Gott. & al., by S.Hatt. 1986 [orth var] Basionym: Distribution: Hattori Bapna & Kachroo = Frullania ericoides 1986 2000 506 712 J Hatt Bot Lab 60: 212 [only listed as synonym] Hepaticology in India II: 169 [only listed as synonym] (Nees) Mont. 1839 Page 242 latilobula Herzog 1942 Typus: Bolivia: Nor Yungas, Polopolo bei Coroico, 1300 m -.10.1912, O. Buchtien 39 in hb JE [holotypus] Basionym: Distribution: Bolivia spec nov - syn nov Herzog Bonner Yuzawa Gradstein, Meneses & Arbe Müller & Zündorf = Frullania rio-janeirensis latistipula 1942 1965 1991 2003 2007 874 655 525 975 1442 Beih Bot Centralbl 61 B: 570-571, fig 6(d-k) [Bolivia] Index Hepat 5: 348 J Hatt Bot Lab 70: 254 [syn sub Fr. riojaneirensis] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. riojaneirensis] http://www2.unijena.de/biologie/spezbot/herbar/bryotypes/Uebersicht%20F.htm (Raddi) Ångstr. 1876 Steph. 1911 Typus: "India Orient" in hb G G00069228 - n° SIB 126630/1 (ex hb M) Basionym: Distribution: Sri Lanka, Indonesia (Sumatra, Java), Philippines, New Caledonia, Society Isl (Tahiti) spec nov - syn nov Bescherelle Stephani Verdoorn Verdoorn Verdoorn Bonner Stephani = Frullania serrata latistipula 1898 1911 1929 1930 1930 1965 1985 871 686 246 293 551 655 928 J Bot Morot 12: 138 [spec nov sine decript; = nom nud] [Tahiti] Spec Hepat 4: 544-545 Ann Bryol 2: 161 Ann Bryol Suppl 1: 85 [syn sub Fr. serrata] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 348 Stephani Icon Hepat Inedit [Thyopsiella] F-145, microfiche 59 fig 4081 ["Nova Caledonia, India orientalis, an eingelegten Pflanzen, Herb München"] - 4082 Gottsche 1845 Steph. ex Bescherelle 1898 [nom nud] Basionym: Distribution: Stephani in Bescherelle = Frullania latistipula Frullania serrata 1898 871 J Bot Morot 12: 138 [spec nov sine decript; = nom nud] Steph. 1911 Gottsche 1845 Page 243 lauterbachii Steph. 1910 Typus: Indonesia. Java, Ardjoeno, 2000 m sd, Lauterbach 450 in hb G [isotypus] Basionym: Distribution: Himalayas, Sri Lanka, Indonesia (Java) spec nov - syn nov Stephani Verdoorn Verdoorn Chopra Parihar Bonner Miller, Whittier & Whittier Stephani = Frullania neurota laxepinnata 1910 1929 1930 1943 1962 1965 1983 1985 686 245 293 205 1584 655 609 928 Spec Hepat 4: 350 Ann Bryol 2: 120 [Java] Ann Bryol Suppl 1: 66 [syn sub Fr. neurota] J Indian Bot Soc 22: 249 [India] Univ Allahabad studies, Bot sect 1-2: 18 [India, syn sub Fr neurota] Index Hepat 5: 349 Prod Flora Hepat Polynesiae: 131 Stephani Icon Hepat Inedit [Chonanthelia] A- 92, microfiche 47 fig 3200 [from type; "Java Ardjoeno 2000m Lauterbach 450 ; Ceylon; India orientalis Sikkim Himalaya"] Taylor 1846 Steph. 1895 [nom nud] [non Steph. 1911] Typus: Africa: Usambara 1893, Holst 9092 in hb BP [Budapest] 11152 Basionym: Distribution: Stephani in Engler Vanden Berghen Wigginton 1895 1976 2002 Wigginton Wigginton 2004 2004 = Frullania angulata laxepinnata 1481 Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [nom nud] [ [Usambara] 185 Bull Jard Bot Nat Belg 46 (1/2): 71 [= nom. nud., syn sub Fr. angulata] 1012 Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 30 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Mitt. 1863 Steph. 1911 [non Steph. 1895] Typus: India. Darjeeling, Chiabhanjan, 10320ft 18.11.1907, "indigenus pro E. Long" hb LEVIER 5675, in hb G 7950 Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Chopra Parihar Bonner Stephani 1911 1930 1943 1962 1965 1985 686 293 205 1584 655 928 Bapna & Kachroo 2000 712 = Frullania nepalensis laxipinnata Spec Hepat 4: 457 Ann Bryol Suppl 1: 51 [syn sub Fr. nepalensis] J Indian Bot Soc 22: 249 [India] Univ Allahabad studies, Bot sect 1-2: 18 [India, syn sub Fr nepalensis] Index Hepat 5: 349 Stephani Icon Hepat Inedit [Galeiloba] C-150, microfiche 52 fig 3544 8from type; "Himalaya, Sikkim, Darjeeling, Levier 5675"] Hepaticology in India II: 176 [syn sub Fr. nepalensis] (Spreng.) Lehm. & Lindenb. 1845 Steph. by Chopra 1943 [orth var] Basionym: Distribution: Chopra = Frullania laxepinnata Frullania nepalensis 1943 205 J Indian Bot Soc 22: 249 [India] Steph. 1911 [non Steph. 1895] (Spreng.) Lehm. & Lindenb. 1845 Page 244 laxiramea Steph. ex Bonner 1965 [nom herb] Typus: New Hebrides. Aneitum -.3.1911, Gunn hb WATTS 26, in hb G 20586 Basionym: Distribution: Vanuatu (Aneityum) Bonner Hattori Miller, Whittier & Whittier Stephani = Frullania gaudichaudii lechleri 1965 1981 1983 1985 655 492 609 928 Index Hepat 5: 349 J Hatt Bot Lab 49: 372 [nom nud; syn sub Fr. gaudichaudii] Prod Flora Hepat Polynesiae: 131 Stephani Icon Hepat Inedit [Galeiloba] C-151, microfiche 52 fig 3545 [from type; "Novae Hebridae, leg Gunn, Watts misit No 26"] (Nees & Mont.) Nees & Mont. 1845 Steph. 1894 Typus: Peru. Tatanara sd, Lechler 2550 in hb G G00069232 - n° SIB 126531/1 (SLATER misit, sub Fr. subaculeata) [typus] in hb MANCH 17176 [holotypus] Basionym: Distribution: Peru, Bolivia, Colombia, Brazil spec nov - syn nov Stephani Stephani Irmscher Stephani Bonner Gradstein & Hekking Menzel Stephani 1894 1911 1914 1916 1965 1979 1984 1985 Uribe & Gradstein Uribe & Gradstein 1998 2003 Uribe 2008 = Frullania atrata ledermannii 228 686 Hedwigia 33: 153-154 Spec Hepat 4: 606 vide 483 222 Biblioth Bot 87: 247 [Bolivia] 655 Index Hepat 5: 350 483 J Hatt Bot Lab 45: 113 [Cat Colombia] 142 Willdenowia 14: 489 [Peru] 928 Stephani Icon Hepat Inedit [Meteriopsis] E-26ab, microfiche 55 fig 3802 ["Peruvia, leg Lechler 2550"] - 3803 ["Peru, Tatanara, Lechler"] 3804 707 Bryophyt Bibl 53: 36 [Catalogue Colombia] 1071 Crypt Bryol 24 (3): 196 [syn sub Fr atrata, belongs to subgen. Thyopsielle = Frullania] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] (Sw.) Nees ex Mont. 1839 Steph. 1924 Typus: PNG. East Sepik:[Nova Guinea] Hauptlager Malu der Sepik-Exped. 1912, Ledermann, 8094 in hb G 3743 [holotypus], Basionym: Distribution: spec nov Stephani Verdoorn Bonner Grolle & Piippo Stephani = Frullania nodulosa 1924 1930 1965 1984 1985 687 549 655 143 928 Spec Hepat 6: 552-553 Nova Guinea 14 (4): 541 [syn sub Fr. nodulosa] Index Hepat 5: 350 Acta Bot Fennica 125: 21 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Homotropantha] D-26, microfiche 54 fig 3743 ["Nova Guinea (Ledermann legit)"] (Reinw., Blume & Nees) Nees 1845 Page 245 leeuwenii fo. obtusiuscula S.Hatt. 1973 [nom inval] Basionym: Distribution: Hattori 1973 542 J Hatt Bot Lab 37: 149 = lehmanniana Gottsche ex Steph. 1910 Typus: Ecuador: Cuenca, 9000 ft 1878, C. Lehmann 1301 in hb G [holotypus]; Basionym: Distribution: Mexico, Ecuador, Colombia spec nov Gottsche ex Stephani Bonner Gradstein & Hekking Yuzawa Stephani 1910 1965 1979 1984 1985 686 655 483 502 928 Fulford & Sharp Yuzawa 1990 1991 105 525 = Frullania sphaerocephala leiboldiana Typus: Mexico Spec Hepat 4: 334 Index Hepat 5: 356 J Hatt Bot Lab 45: 113 [Cat Colombia] J Hatt Bot Lab 57: 399-401 (No 13), Fig 13 (a-n) from type Stephani Icon Hepat Inedit [Chonanthelia] A- 39ab, microfiche 46 fig 3125-3126 ["Ecuador Cuenca FC Lehmann 1878"] Mem NY Bot Gard 63: 44 [Mexico] J Hatt Bot Lab 70: 276-278, fig. 49 Spruce 1884 Steph. 1911 sd, Leibold sn ex hb RABENHORST, in hb G 20616 [holotypus] Basionym: Distribution: Mexico spec nov - syn nov Stephani Bonner Stephani 1911 1965 1985 Fulford & Sharp Uribe & Gradstein Uribe 1990 2003 2008 = Frullania atrata leiboldii 686 655 928 Spec Hepat 4: 609 Index Hepat 5: 351 Stephani Icon Hepat Inedit [Meteriopsis] E-44, microfiche 56 fig 3834 ["Mexico, leg Leibold"] - 3835 105 Mem NY Bot Gard 63: 44 [Mexico] 1071 Crypt Bryol 24 (3): 199 [syn sub Fr atrata] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] (Sw.) Nees ex Mont. 1839 Lindenb. & Gottsche 1847 Typus: Mexico. Veracruz, Zacuapan Leibold Basionym: Distribution: Mexico spec nov - syn nov Lindenberg & Gottsche in G., L. & N. Gottsche Gottsche Stephani Bonner Stotler Stotler Stephani 1847 2848 Syn Hepat: 782 (No 107c) 1863 1867 1911 1965 1969 1970 1985 103 104 686 655 4761 194 928 Fulford & Sharp 1990 105 = Frullania brasiliensis Separata of Mex. Leverm.: 248 K Danske Vid Selks Skrifter, ser 5, 6: 344 [Mexico] Spec Hepat 4: 490 [syn sub Fr. brasiliensis] Index Hepat 5: 351 Nova Hedwigia 18: 436 The genus Frullania, separata: 436 Stephani Icon Hepat Inedit [Thyopsiella] F- 37c, microfiche 57 fig 3925 ["sub Fr Leiboldii, Costarica, ..."] Mem NY Bot Gard 63: 44 [Mexico] Raddi 1822 Page 246 lepervanchii Steph. 1911 Typus: [1] La Reunion - [2] Madagascar [1] sd, Lapervanche sn - [2] 1896, Forsyth Major [489 pp] hb BESCHERELLE 102, in hb G Basionym: Distribution: Madagascar, Reunion Isl spec nov - syn nov Stephani Arnell Bonner Vanden Berghen Stephani 1911 1963 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania lindenbergii leprieurii 686 822 655 185 928 Spec Hepat 4: 628 Hepat S Africa: 133 [syn nov syn sub Fr. lindenbergii] Index Hepat 5: 351 Bull Jard Bot Nat Belg 46 (1/2): 33-38 Stephani Icon Hepat Inedit [Diastaloba] B- 9, microfiche 48 fig 3236 ["Madagascar (489 ex parte) (Forsyth Major legit) 1896" - auch Reunion leg Lepervanche"] 250 Bryophyt Bibl 48: 66 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Lehm. 1844 Lindenb. 1845 [hom illeg] [non Nees & Mont. in Mont. 1840] Typus: French Guiana. Mt. Badeul near Cayenne. -.5.1857, Leprieur 273 Basionym: Distribution: spec nov Lindenberg in G., L. & N. Montagne Stephani Stephani Stephani 1845 1855 1897 1911 1912 2850 475 1059 686 1636 nom nov Clark & Svihla Schiffner & Arnell Bonner Stephani 1952 1964 1965 1985 347 198 655 928 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 2002 782 = Frullania neesii Lindenb. 1845 Frullania kunzei leprieurii Syn Hepat: 449 Ann Sci Nat Bot, ser 4, 3: 320 Bihang Kgl Sv Vetensk-Akad Handl bot 23 III (2): 19 [Brazil] Spec Hepat 4: 525 Spec Hepat 5: 127 [ Phragnicoma corticalis syn sub Brachiolejeunea corticalis] Bryologist 55: 289-191 [as Fr. magniflora, nom nov] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 89 [Brazil] Index Hepat 5: 351 Stephani Icon Hepat Inedit [Thyopsiella] F-115, microfiche 58 fig 4040 [" ... "] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Nees & Mont. in Mont. 1840 [non Lindenb. in G., L. & N. 1845] Typus: "ad radices montis Baduel propé Cayennam 1837, Majo hb LEPRIEUR 273 & 277 Basionym: Distribution: spec nov Nees & Montagne in Montagne 1840 Lehmann & Lindenberg in G., L. 1845 & N. Montagne 1856 = Phragmicoma corticalis 467 Ann Sci Nat Bot, ser 2, 14: 333-334, pl 20(1) 2847 Syn Hepat: 297 [syn sub Phragmicoma corticalis] 1912 Sylloge generum specierumque cryptogamaram: 86 (Lehm. & Lindenb. in Lehm.) Lehm. & Lindenb. 1845 Page 247 leprieurii var. wrightii Gottsche ex Bonner 1965 [nom herb] Typus: Cuba. sd, Wright 118 in hb G (sub Fr. serrifolia), in hb US Basionym: Distribution: Bonner Stotler Stotler NN = Frullania neesii 1965 1969 1970 1996 Index Hepat 5: 351 Nova Hedwigia 18: 450 The genus Frullania, separata: 450 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 20 of 35 [in hb US] Lindenb. 1845 Frullania kunzei leptomitra 655 4761 194 632 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Spruce 1884 Typus: Ecuador: Andes Quitenses, Flum Bombonasa sd, Spruce sn in hb G [holotypus] Basionym: Distribution: Ecuador spec nov Spruce Stephani Bonner Stephani 1884 1910 1965 1985 243 686 655 928 - syn nov Yuzawa 1991 525 = Frullania rio-janeirensis leptophylla Trans Proc Bot Soc Edinburgh 15: 13 & 25 (No 13) Spec Hepat 4: 328 Index Hepat 5: 352 Stephani Icon Hepat Inedit [Chonanthelia] A- 40, microfiche 46 fig 3127 [from type; "Bombonasa leg Spruce"] J Hatt Bot Lab 70: 252 [syn sub Fr. riojaneirensis] (Raddi) Ångstr. 1876 De Not. 1857 Typus: Brazil. Serra dees Orgaos. sd, Casaretto Basionym: Distribution: Argentina spec nov - syn nov Notaris 1855 Notaris 1857 Stephani Bonner Fulford Yano Stephani 1911 1965 1967 1984 1985 Yuzawa & Koike Winter & Schäfer-Verwimp 1989 2020 = Frullania obcordata leratae 5341 Jungermanniearum Americanarum pugillus [Seperata]: 28-29, fig 19(1-5) [spec nov] 558 Mem Reale Accad Sci Torino, ser 2, 16: 234-235, fig 19(1-5) [spec nov] 686 Spec Hepat 4: 633 655 Index Hepat 5: 352 1196 Acta Bot Venezuelica 2(5-8): 94 [Argentina, Auyan-tepui] 501 J Hatt Bot Lab 56: 505 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Diastaloba] B-20, microfiche 48 fig 3250 ["Brasilia"] 519 J Hatt Bot Lab 66: 347-349, fig 4(from type) [syn sub Fr. caulisequa] 5742 Frahmia 19: 13 [syn sub Fr obcordata] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. ex Bonner 1965 [nom herb, specimen Le Rat 534] Typus: New Caledonia. "Nov. Caledon, Mt. Mon -.7.109, Le Rat 534 in hb G [as Fr. leratae] Basionym: Distribution: Hattori = Frullania pusilla 1986 506 J Hatt Bot Lab 60: 217-218 [= Fr. pusilla, hb G, Le Rat 534] Mitt. 1871 Page 248 leratae Steph. ex Bonner 1965 [nom herb, specimen Le Rat 112] Typus: New Caledonia. "Nov. Caledon, Mt. Mon -.7.1909, Le Rat 112 in hb G Basionym: Distribution: New Caledonia nom herb - syn nov Bonner Miller, Whittier & Whittier Hattori = Frullania caledonica 655 609 506 Index Hepat 5: 352 Prod Flora Hepat Polynesiae: 131 J Hatt Bot Lab 60: 208 [nom nud; as syn sub Fr. caledonica] Gottsche ex Steph. 1894 [non (Schuster) Schuster 1992] Frullania squarrosula levicalyx 1965 1983 1986 (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Steph. 1910 Typus: ? Africa: Bangi 10.8.1902, A. Chevalier sn ex hb CORBIERE 72 pp in hb G [holotypus], in hb PC [isotypus] Basionym: Distribution: spec nov - syn nov Stephani Stephani in Corbiere Bonner Vanden Berghen Stephani 1910 1912 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania trinervis levieri 686 2097 655 185 928 Spec Hepat 4: 372 Bull Mus Nat Hist Nat Paris 18 (2): 120 [as spec nov] Index Hepat 5: 352 Bull Jard Bot Nat Belg 46 (1/2): 199 [syn sub Fr trinervis] Stephani Icon Hepat Inedit [Galeiloba] C- 22, microfiche 50 fig 3374 [from type; "Trachycolea, Africa Bangi, Aug 1902, Auguste Chevalier, Corbiere 72 pp"] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 227, fig 149(B) from type [syn sub Fr trinervis, West Africa] (Lehm.) Drège 1843 Schiffn. ex Verd. 1930 [pro syn] [non Steph. 1910] Basionym: Distribution: Verdoorn 1930 293 Ann Bryol Suppl 1: 115 = Frullania gracilis var. subapiculata Schiffn. ex Verd. 1929 Frullania gracilis leyboldii var. (Reinw., Blume & Nees) Nees 1845 Lindenb. & Gottsche by Bonner 1965 [orth var] Basionym: Distribution: Bonner = Frullania leiboldii 1965 655 Index Hepat 5: 353 Lindenb. & Gottsche 1847 Frullania brasiliensis Raddi 1822 Page 249 libera Nees in Flotow, Göppert & Nees 1842 Typus: Brazil. "on bark of Pào Pereira " = Pau-pereira-do-mato (Aspidosperma subincanum Mart., Apocynaceae)" - * "in Pao Pereiro, Brasiliae" [NEES 1845] is not a locality vide NEES 1842 sd, nn sn in hb G (ex hb NEES) in hb STR ?? Basionym: Distribution: Brazil spec nov - syn nov Flotow, Göppert & Nees 1842 Nees in G., L. & N. 1845 Stephani Bonner Yano 1911 1965 1984 Stephani 1985 Gradstein & Costa 2003 Yano 2008 = Frullania glomerata Mont 1842] liebmaniana 2475 Archiv Pharmazie 80 (3): 324, "on bark of Pào Pereira (Aspidosperma subincanum Mart., Apocynaceae)" 2850 Syn Hepat: 430-431, "in Pào Pereiro, Brasiliae" [NEES 1845] is not a locality vide NEES 1842 686 Spec Hepat 4: 675-676 "Hab. Brasilia, Pào Pereiro" 655 Index Hepat 5: 353 "Type: Brazil, Pào Pereiro, ex hb Nees, in hb G" 501 J Hatt Bot Lab 56: 505 [checklist Brazil] "B: Pào Pereira (Stephani 1911) 928 Stephani Icon Hepat Inedit [ [Galeiloba] C- 61abcd, microfiche 50 fig 3420 [sub Fr glomerata] 984 Mem New York Bot Gard 87: 87 [Brazil] [syn sub Fr glomerata, fide Stephani, in sched] 1684 Boletim Instituto Botanica 19: 90 [Brazil; catalog - "syn sub Fr glomerata (Gradstein & Costa 2003"] (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Lindenb. & Gottsche 1847 Typus: Mexico. Veracruz, Mirador, Aguas Santas sd, Liebman sn typus in hb S (B24125), lectotypus by Uribe-M & Gradstein 2003: in hb W 7572 Basionym: Distribution: Mexico spec nov - syn nov Lindenberg & Gottsche in G., L. & N. Gottsche Gottsche Stephani Clark Bonner Stephani 1847 2848 Syn Hepat: 784 1863 1867 1911 1953 1965 1985 103 104 686 357 655 928 Fulford & Sharp Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Uribe & Gradstein Uribe 1990 2002 105 782 2003 2008 1071 Crypt Bryol 24 (3): 196 [syn sub Fr atrata] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] = Frullania atrata ligulaefolia Separata of Mex. Leverm.: 253 K Danske Vid Selks Skrifter, ser 5, 6: 349-350 [Mexico] "liebmanniana" Spec Hepat 4: 607 Bryologist 56: 217-219, fig 1-15 Index Hepat 5: 353 Stephani Icon Hepat Inedit [Meteriopsis] E- , microfiche 56 fig 3836 ["Mexico"] Mem NY Bot Gard 63: 44 ["liebmanniana", Mexico] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] (Sw.) Nees ex Mont. 1839 Steph. ex Theriot 1910 [nom nudum] Basionym: Distribution: nom nud Paris = Frullania ligulifolia 1908 864 Rev Bryol 35: 62 [as Fr. ligularis (=nom nud)] Steph. ex Paris 1908 [nom nud] Frullania ligulifolia Frullania pusilla Steph. 1911 Mitt. 1871 Page 250 ligularis Steph. ex Paris 1908 [nom nud] Typus: New Caledonia Basionym: Distribution: Stephani in Paris Miller, Whittier & Whittier Hattori = Frullania ligulifolia 864 609 506 Rev Bryol 35: 62 Prod Flora Hepat Polynesiae: 131 J Hatt Bot Lab 60: 218 [= nom nudum] Steph. 1911 Frullania pusilla ligulifolia 1908 1983 1986 Mitt. 1871 Steph. ex Paris 1908 [nom nud] Basionym: Distribution: Paris Stephani ex Theriot Paris nom nud = Frullania ligulifolia 1455 Rev Bryol 37 (6): 130 [New Caledonia: Poindimié; mont. de Crémon] 1915 Bull Acad Int Geogr Bot 20: 104 [nom nudum] 911 Rev Bryol 37 (1): 41 [New Caledonia: Isle of Pines] Steph. 1911 Frullania pusilla ligulifolia 1910 1910 1910 Mitt. 1871 Steph. 1911 Typus: New Caledonia. 1905, Etesse in hb E.G. PARIS 23, in hb G Basionym: Distribution: spec nov Stephani Verdoorn Bonner Hattori Miller, Whittier & Whittier Hattori Stephani 1911 1930 1965 1977 1983 1984 1985 686 551 655 479 609 503 928 - syn nov Hattori 1986 506 = Frullania pusilla limbatiflora Spec Hepat 4: 466 Nederl Kruidk Arch Jrg 1930 (2): 161-162 [New Caledonia] Index Hepat 5: 353 J Hatt Bot Lab 43: 424 [New Caledonia] Prod Flora Hepat Polynesiae: 131 J Hatt Bot Lab 57: 419 Stephani Icon Hepat Inedit [Galeiloba] C-152, microfiche 52 fig 3546 [from type; " NCaledonia, Etesse leg"] J Hatt Bot Lab 60: 217 [syn sub Fr. pusilla] Mitt. 1871 Steph. ex Bonner 1965 [nom herb] Typus: Japan. Mishino, Aki -.12.1910, S. Okamura 358 in hb G Basionym: Distribution: Japan Bonner Stephani 1965 1985 655 928 Index Hepat 5: 353 Stephani Icon Hepat Inedit [Galeiloba] C-244, microfiche 53 fig 3664 ["Japonia, O Kamura No 358"] = Page 251 lindbergii Gottsche by Ångstr. 1876 [orth var] Basionym: Distribution: Brazil orth var Ångström Stephani Bonner Yano 1876 1910 1965 1984 = Frullania lindbergiana lindeniana 579 686 655 501 Öfversigt K Vetensk Akad Förh 33 (7): 88 [orth err] [Brazil] Spec Hepat 4: 384 [as Fr. lindbergii syn sub Fr. glomerata] Index Hepat 5: 353-354 [Fr. lindbergii = nom herb] J Hatt Bot Lab 56: 505 [as Fr. lindbergii p.p.; checklist Brazil] Gottsche 1863 Steph. 1911 Typus: Cuba. sd, Linden 2310 hb L939, in hb G 20617 Basionym: Distribution: Cuba spec nov Stephani Bonner Stephani 1911 1965 1985 - syn nov Uribe & Gradstein Uribe 2003 2008 = Frullania atrata lobangensis 686 655 928 Spec Hepat 4: 603 Index Hepat 5: 354 Stephani Icon Hepat Inedit [Meteriopsis] E-27, microfiche 55 fig 3805 ["Cuba, leg Linden, Herb Leiden 939"] 1071 Crypt Bryol 24 (3): 199 [syn sub Fr atrata] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] (Sw.) Nees ex Mont. 1839 Steph. 1910 Typus: Angola. Chella Mts, Lobango, 2200m sd, Newton 1883 in hb G 17309 [holotypus], G 17310 [isotypus] Basionym: Distribution: spec nov Stephani Bonner Vanden Berghen Stephani 1910 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania caffraria lochensis 686 655 185 928 Spec Hepat 4: 379 Index Hepat 5: 354 Bull Jard Bot Nat Belg 46 (1/2): 178 Stephani Icon Hepat Inedit [Galeiloba] C- 23, microfiche 50 fig 3375 [from type; "Africa Lobango, Newton legit"] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 229, fig 150(P,Q) [prthaps syn sub Fr caffraria, West Africa] Steph. 1894 Hampe ex Lehm. & Lindenb. 1845 [pro syn] Basionym: Distribution: Peru - syn nov Lehmann & Lindenberg in G., L. 1845 & N. Bonner 1965 2850 Syn Hepat: 462 [syn sub Fr mucronata var submutica] 655 Index Hepat 5: 355 = Frullania mucronata var. submutica Lehm. & Lindenb. 1845 Page 252 longe-attenuata S.Hatt. 1973 Typus: New Guinea. Central Highlands ... 10.7.1967, R.M. Schuster 67-5953 in hb NICH [holotypus] in hb MASS & hb JE [isotypus] Basionym: Distribution: spec nov - comb nov Hattori Hattori Hattori Hattori 1973 1973 1974 1975 Hattori 1980 Grolle & Piippo 1984 Hattori & Piippo 1986 Mizutani, Hasegawa & Iwatsuki 2009 540 542 546 547 J Hatt Bot Lab 37: 66-68, fig 29 J Hatt Bot Lab 37: 149 [New Guinea] J Hatt Bot Lab 38: 242-243 [New Guinea] J Hatt Bot Lab 39: 280 [comb nov; as Fr. attenuata var longeattenuata] 489 J Hatt Bot Lab 47: 107-109 [Fr. longe-attenuata is conspecific with Fr. armitiana & Fr. attenuata] 143 Acta Bot Fennica 125: 21 [Cat. West Melanesia] 557 Acta Bot Fennica 133: 27 2112 Type sepecimens Hatt Bot Lab (NICH) 29: Holotype. New Guinea, Central Highland District, from Kundiawa to Goroka. 10 June 1967. R. M. Schuster 67-5953 (NICH 309919). = Frullania attenuata var. longe-attenuata (S.Hatt.) S.Hatt. 1975 Frullania armitiana var. longe-attenuata (S.Hatt.) S.Hatt. 1980 longicollis Lindenb. & Gottsche 1847 Typus: Mexico. Oaxaca,? Ad Yavesea, 8000ft sd, Liebmann typus in hb S (B4130, 24131) Basionym: Distribution: Mexico, Costa Rica, Venezuela, Peru spec nov - syn nov - syn nov Lindenberg & Gottsche in G., L. & N. Gottsche Gottsche Stephani in Brown 1847 2848 Syn Hepat: 783 1863 1867 1901 Stephani Herzog Clark Bonner Stotler Stotler Menzel Stephani 1911 1953 1958 1965 1969 1970 1984 1985 Fulford & Sharp Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 1990 2002 103 Separata of Mex. Leverm.: 250 104 K Danske Vid Selks Skrifter, ser 5, 6: 346-347 [Mexico] 1284 Trans Linn Soc London, ser 2 (Bot) 4: 100 [Summit of Roraima, British Guiana] 686 Spec Hepat 4: 498-499 640 Rev Bryol Lichen 20: 128 [Costa Rica] 386 Bryologist 61: 143-146, fig 1-19 655 Index Hepat 5: 355 4761 Nova Hedwigia 18: 444 [syn sub Fr. intumescens] 194 The genus Frullania, separata: 444 [syn sub Fr. intumescens] 142 Willdenowia 14: 510 [syn of Fr. intumescens] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 67ab, microfiche 57 fig 3969 ["olim Frull robusta Spruce ms (Tunguragua) Andes"] - 3970 ["ex planta Wagneri, Panama"] - 3971 ["Mexico, Original"] - 3972 105 Mem NY Bot Gard 63: 44 [Mexico] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] = Frullania intumescens (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 253 longipinna Steph. 1910 Typus: Colombia. Paramo de Ocana sd, Pehlke sn in hb G (Holotypus G 13643) Basionym: Distribution: Colombia spec nov - syn nov Stephani Bonner Stotler Stotler Gradstein & Hekking Stephani 1911 1965 1969 1970 1979 1985 Uribe & Gradstein Uribe 1998 2008 = Frullania convoluta longipinnata 686 655 4761 194 483 928 Spec Hepat 4: 534 [=sgen Frullania] Index Hepat 5: 355 Nova Hedwigia 18: 477 [as Fr. longipinnata] The genus Frullania, separata: 477 [as Fr. longipinnata] J Hatt Bot Lab 45: 113 [Cat Colombia] Stephani Icon Hepat Inedit [Thyopsiella] F-116, microfiche 58 fig 4041 ["Columbia (leg Pehlke)"] - 4042 707 Bryophyt Bibl 53: 36 [Catalogue Colombia] 1513 Caldasia 30(1): 66 [syn sub Fr convoluta] [Monography of subgenus Meteoriopsis] Lindenb. & Hampe 1851 Steph. by Stotler 1969 [orth var] Basionym: Distribution: Stotler Stotler 1969 1970 = Frullania longipinna Steph. 1910 Frullania convoluta longirostris 4761 Nova Hedwigia 19: 477 194 The genus Frullania, separata: 477 Lindenb. & Hampe 1851 Steph. 1894 Typus: Central Africa. "in monte celeberrimo Runssoro" Ruwenzori sd, Stuhlmann 2413B ex hb B, in hb G [holotypus] Basionym: Distribution: Central Africa spec nov - syn nov Stephani Stephani in Engler 1894 1895 Stephani Stephani Stephani Stephani in Mildbraed 1895 1902 1910 1911 Stephani in Mildbraed 1914 Pearson Demaret Hauman Demaret & Vanden Berghen Arnell Arnell Bonner Stephani 1924 1942 1942 1948 1956 1963 1965 1985 Yuzawa 1991 Wigginton 2002 Wigginton 2004 = Frullania ecklonii 228 Hedwigia 33: 139 1481 Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [Seengebiet, (R., 3300m) Hochwald] 212 Bot Jahrb 20 (3): 314-315 [as spec nov] 215 Bot Jahrb 30: 260 [Kingagebirge] 686 Spec Hepat 4: 320 [as spec nov] Wiss Ergebn Deutsch Z Afr Exped 1907-1908, (Separata): ? 22-23, fig 45(a-e) [as spec nov] 820 Wiss Ergebn Deutsch Z Afr Exped 1907-1908, (Bot=2): 132-133, fig 45(a-e) [as spec nov] 795 Arkiv Botanik 19 (5): 9 [Mt. Elgon in Kenia/Uganda] 1066 Bull Jard Bot Etat Bruxelles 16 (2): 304-305 1067 Bull Jard Bot Etat Bruxelles 16 (4): 352 [Ruwenzori] 59 Bull Jard Bot Etat Bruxelles 19 (1): 74-84 [:82 syn sub Fr. ecklonii] 836 Ark Botanik 3 (16): 531 [East African mountains] 822 Hepat S Africa: 138 [syn sub Fr. ecklonii] 655 Index Hepat 5: 355-356 928 Stephani Icon Hepat Inedit [Chonanthelia] A- 6ab, microfiche 46 fig 3071-3073 [from type; "Stuhlmann 2413 - Milbraed 2647, 831 - "] 525 J Hatt Bot Lab 70: 259, fig. 40 [from type specimen] [syn. nov. syn sub Fr. ecklonii] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] (Spreng.) Gottsche, Lindenb. & Nees 1845 Page 254 longispica Steph. 1910 Typus: Indonesia. Timor, 1100 m -.11.1897, Francis Newton sn in hb G Basionym: Distribution: Indonesia (Java, Timor), New Caledonia spec nov - syn nov Stephani Verdoorn Frye & Clark Kamimura Bonner Miller, Whittier & Whittier Stephani 1911 1930 1947 1961 1965 1983 1985 686 293 204 199 655 609 928 Bapna & Kachroo 2000 712 = Frullania squarrosa (Reinw., Blume & Nees) Nees 1845 Frullania ericoides longispica Spec Hepat 4: 454 Ann Bryol Suppl 1: 34 [syn sub Fr. squarrosa] Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] J Hatt Bot Lab 24: 17 [syn sub Fr squarrosa] Index Hepat 5: Prod Flora Hepat Polynesiae: 131 Stephani Icon Hepat Inedit [Galeiloba] C-153, microfiche 52 fig 3547 [from type; "Insa Timor, Fr Newton, Lisboa 1897"] Hepaticology in India II: 170 [syn sub Fr. ericoides] (Nees) Mont. 1839 Steph. ex Paris 1910 [nom nud] Basionym: Distribution: Paris = Frullania longispica 1910 Steph. 1910 Frullania squarrosa (Reinw., Blume & Nees) Nees 1845 Frullania ericoides loxensis 1455 Rev Bryol 37 (6): 130 [New Caledonia: Dogny] (Nees) Mont. 1839 Hampe ex Raddi ex Gottsche, Lindenb. & Nees 1845 [pp] [pro syn] Basionym: Distribution: Raddi in G., L. & N. = Frullania brasiliensis loxensis 1845 2850 Syn Hepat: 458 (No 106) Raddi 1822 Hampe ex Stotler 1969 [pp] [nom herb] Basionym: Distribution: Stotler Stotler = Frullania macrocephala ludviciae 1969 1970 4761 Nova Hedwigia 18: 439 194 The genus Frullania, separata: 439 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. by S.Hatt. 1986 [orth var] Basionym: Distribution: Hattori = Frullania ludoviciae 1986 506 J Hatt Bot Lab 60: 215-216 Steph. 1908 Page 255 luzonensis Steph. 1910 Typus: Philippines: Luzon, 1500 m 1906, Loher sn hb LEVIER 5128, in hb G Basionym: Distribution: Philippines (Luzon) spec nov - syn nov Stephani Verdoorn Frye & Clark Kamimura Bonner Stephani 1910 1930 1947 1961 1965 1985 Wu in Li, Gao, Wu et al. 1985 Bapna & Kachroo 2000 = Frullania squarrosa Spec Hepat 4: 447 Ann Bryol Suppl 1: 34 Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] J Hatt Bot Lab 24: 17 [syn sub Fr squarrosa] Index Hepat 5: Stephani Icon Hepat Inedit [Galeiloba] C-155, microfiche 52 fig 3549 [from type; "Luzon (Loher legit) Levier 5128"] 1169 Bryoflora Xizang (Tibet), Science Press, Beijing: 523 [sub Fr squarrosa] 712 Hepaticology in India II: 170 [syn sub Fr. ericoides] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides maccraeana 686 293 204 199 655 928 (Nees) Mont. 1839 Mueller 1858 [orth var] Basionym: Distribution: Mueller Scott & Bradshaw = Frullania maccrantica 1858 1986 Gottsche 1856 Frullania probosciphora maccrantica Fourth Syst Index Pl Victoria 12: Brunnonia 8 (1): 35 [presumably an error for Fr maccrantica GOTTSCHE] 56 Taylor 1846 Gottsche 1856 Typus: Australia. "in insula Maccraes et in Wilsons Promontory" sd, Ferd v . Müller Basionym: Distribution: Australia (Maccraes Isl) spec nov - syn nov Gottsche Bibby Bonner Scott & Bradshaw Carthy McCarthy = Frullania probosciphora 1856 1955 1965 1986 2003 2006 23 868 655 56 1125 1385 Linnaea 28: 559 Victorian Naturalist 72: [syn sub Fr. probosciphora] Index Hepat 5: 357 Brunnonia 8 (1): 35-36 [Australia] Flora Australia Suppl Ser 21: 45 [sub Fr probosciphora] Checklist Australian Liverworts [Australia, syn sub Fr probosciphora] Taylor 1846 Page 256 maccreana F. v. Muell. ex Steph. 1880 [orth var] Typus: Australia: Maccreas Insula Basionym: Distribution: F. v. Muell. ex Steph. Stephani Bibby Bonner Stephani 1880 1910 1955 1965 1985 Scott & Bradshaw Carthy McCarthy 1986 2003 2006 = Frullania maccrantica Frag Phyt Australia 11, suppl Hepaticae :65 Spec Hepat 4: 355-356 [as Chonanthelia] Victorian Naturalist 72:[syn sub Fr. probosciphora] Index Hepat 5: 357 Stephani Icon Hepat Inedit [Chonanthelia] A-105, microfiche 48 fig 3220 ["Australia Maccreas Ins .."] 56 Brunnonia 8 (1): 35 [Australia] 1125 Flora Australia Suppl Ser 21: 45 [sub Fr probosciphora] 1385 Checklist Australian Liverworts [Australia, syn sub Fr probosciphora] Gottsche 1856 Frullania probosciphora macrocephala 1489 686 868 655 928 Taylor 1846 (Lehm. & Lindenb.) Nees 1838 [nom inval, Art. 32.1(c)] Basionym: Distribution: Nees Bernal, Gradstein & Celis = Frullania macrocephala macroloba 1838 2016 261 Naturg Europ Leberm 3: 210 [nomen in footnote] 4290 Catálogo de plantas y líquenes de Colombia. Vol. I. (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1910 Typus: Madagascar sd, Forsyth Major sn in hb G 13046 Basionym: Distribution: Madagascar, Seychellen spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1910 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania anderssonii 686 655 185 928 Spec Hepat 4: 377-378 Index Hepat 5: 358 Bull Jard Bot Nat Belg 46 (1/2): 172 Stephani Icon Hepat Inedit [Galeiloba] C- 24, microfiche 50 fig 3376 [rom type; "Madagascar, Forsyth Major leg"] 250 Bryophyt Bibl 48: 62 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Ångstr. 1873 Page 257 macrorhyncha Herzog & Clark in Clark & Schultz 1953 Typus: Costa Rica. San Jose, Zurqui, 2000-2500m 13.2.1926, Standley & Vaerio 48135 in hb US Basionym: Distribution: Costa Rica Herzog & Clark in Clark & Schultz Bonner Stotler Stotler NN = Frullania hamiflora macrostachya 1953 358 Bryologist 56: 283-285, fig 1-15 1965 1969 1970 1996 655 4761 194 632 Index Hepat 5: 358 Nova Hedwigia 18: 474-475 [syn sub Fr. hamiflora] The genus Frullania, separata: 474-475 [syn sub Fr. hamiflora] gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 21 of 35 [in hb US][as macrorrhyncha, orth err] Herzog & L.Clark 1953 J.B.Jack ex Bonner 1965 [nom herb] Basionym: Distribution: Bonner 1965 = Frullania tetraptera macrostipula 655 Index Hepat 5: 358 Nees & Mont. 1838 Steph. 1911 Typus: Indonesia. Java, Cibodas 1889, Karstens ex hb STITZENBERGER, in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Stephani 1911 1929 1930 1965 1985 Uribe 2008 = Frullania nigricaulis macrotus 686 246 293 655 928 Spec Hepat 4: 611 Ann Bryol 2: 162 Ann Bryol Suppl 1: 153 [syn sub Fr. nigricaulis] Index Hepat 5: 359 Stephani Icon Hepat Inedit [Meteriopsis] E-53, microfiche 56 fig 3849 ["Java, Tjibodas, Dr G Karsten 1889 (Herb Jack)"] 1513 Caldasia 30(1): 90, [belongs to subgen Frullania] [Monography of subgenus Meteoriopsis] (Reinw., Blume et Nees) Nees 1845 Lindenb. 1845 Typus: Chile, [ad ramos] sd, Bongard ex hb LE, in hb LEHMANNIANUM Basionym: Distribution: Chile spec nov - syn nov Lindenberg in G., L. & N. Stephani Stephani Bonner = Frullania gibbosa 1845 1910 1910 1965 2850 686 686 655 Syn Hepat: 425 (No 30) Spec Hepat 4: 344 [syn sub Fr. gibbosa] Spec Hepat 4: 430 [as Trachycolea] Index Hepat 5: 359 Nees 1840 Page 258 magellanica var. diminutiva Herzog 1938 Typus: Chile. Prov Corral, Quitaluto, auf Rinde zwischen Lophocolea payana (Prov. Cautin: Pucon) or or 522 (Prov. Chiloa: Petrohue) -.2.1935, C.C.Hossens 679 (Prov. Valdivia: Quitaluto) or 204 in hb JE (Hossens 679) [holotypus] CHILE. Valdivia: Quitaluto, Herzog 679 ex p. (JE!, lectotype here designated); Hentschel et alii, 2015 Basionym: Distribution: Chile (Valdivia, Cautin, Chiloe) var nov - syn nov Herzog & Hosseus 1938 Bonner Müller & Zündorf 1965 2007 Hentschel et alii 2015 = Frullania fertilis magniflora 723 Arch Esc Farm Fac Ci Med Cordoba 7: 29-31, fig 10(a-c) [Chile: Valdivia, Cautin] 655 Index Hepat 5: 360 1442 http://www2.unijena.de/biologie/spezbot/herbar/bryotypes/Uebersicht%20F.htm 4097 Phytotaxa 220 (2): 135 [=Frullania (subg. Microfrullania) fertilis De Not] De Not. 1855 Clark & Svihla 1952 [nom nov] Basionym: Frullania leprieurii Distribution: Lindenb. 1845 [hom illeg] [non Nees & Mont. in Mont. 1840] Clark & Svihla Stotler Stotler = Frullania neesii 1952 1969 1970 347 Bryologist 55: 289-291 [nom nov] 4761 Nova Hedwigia 18: 450 [syn sub Fr. neesii] 194 The genus Frullania, separata: 450 [syn sub Fr. neesii] Lindenb. 1845 Frullania kunzei (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 magniflora var. wrightii Clark & Svihla 1952 Typus: Cuba. sd, C. Wright Basionym: Distribution: var nov Clark & Svihla Clark 1952 1955 - syn nov - syn nov Stottler Stottler 1969 1970 = Frullania neesii Bryologist 55: 291-293, fig 1-14 Bryologist 58: 149 [difference to Fr. strobilantha, intumescens, mollicula] 4761 Nova Hedwigia 18: 450 [syn sub Fr. neesii] 194 The genus Frullania, separata: 450 [syn sub Fr. neesii] Lindenb. 1845 Frullania kunzei magnistyla 347 371 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Glenny, Meagher & Brown 2003 [nom nud] Basionym: Distribution: Australia Carthy Glenny, Meagher & Brown 2003 2003 1125 Flora Australia Suppl Ser 21: 112 [nom nud] 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html = Page 259 maior Raddi 1818 Basionym: Distribution: - syn nov Raddi Raddi Raddi 1818 1820 1820 Nees Raddi in Nees Nees Müller in Rabenhorst Bonner Stotler Piippo Wang & Tau Zaho, Fan, Wie & Lu 1838 1841 1841 1915 1965 1968 1990 1992 1993 = Frullania tamarisci makinoana Jungermanniografia etrusca [Seperate print]: 9 1651 Mem Mat Fisica Soc Ital Sci, Modena 18 (1): 20-21 2874 Preprint of Mem Mat Fisica Soc Ital Sci, Modena 18: (1): 2021 261 Naturg Europ Leberm 3: 229 [syn sub Fr. tamarisci] 164 Jungermannogr Etrusca [Seperate print with remarks by Nees]: 5 164 Reprint of Jungerm Etrusca: 5, pl. 2(2a-d) [= Fr. tamarisci] 1064 Krypt Flora (2ed) 6, 2: 610 [sub Fr. tamarisci] 655 Index Hepat 5: 360 ["cf. Fr. dilatata"] 197 Taxon 17: 635-638 [= Fr. tamarisci] 521 J Hatt Bot Lab 68: 43 [annotated catalogue of China] [aus 698] 1224 Chenia 1: 120 [China, Shandong Prov] (L.) Dumort. 1835 Steph. 1897 Typus: Japan. Chiba Pref., Prov. Awa, Mt. Kiyozumi-Yama 8.4.1896, T. Makino 141 in hb G; in hb TNS [isotypus] Basionym: Distribution: spec nov - syn nov Stephani Evans Sawada 1897 1906 1909 Stephani Iishiba Horikawa Verdoorn Hattori Hattori Kamimura Hattori Kamimura Hattori & Noguchi Bonner Hattori Stephani 1911 1912 1929 1932 1944 1951 1952 1952 1952 1960 1965 1972 1985 Inoue Kuo & Chiang Piippo N.N. [National Science Museum Tokyo]N.N. [National Science Museum Tokyo] Yamada & Iwatsuki 1987 1988 1990 2000 Bull Herb Boissier 5: 78 & 89-90 (spec nov; Japan] Proc Wash Acad Sci 8: 159 Bot Mag Tokyo 23 (no 268): 178-183 [key to Japanese species, in Japanese] 686 Spec Hepat 4: 551 888 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] 924 Sci Rep Tohoku Imp Univ, ser 4, Biol 4: 68-70, fig 6(1-8) [Japan] 554 Nederl Kruidk Arch Jrg 1932 (2): 495 943 Bull Tokyo Sci Mus 11: 146 [Japan, key to Japanese species] 528 J Hatt Bot Lab 5: 64 769 Acta Phytotax Geobot 14 (4): 109 529 J Hatt Bot Lab 8: 29 873 Contrib Fl Hepat Shikokuensem, Kochi: 125-126 965 Index spec Typ herb Japon, Bryophyta: 17 [isoype] 655 Index Hepat 5: 360 536 J Hatt Bot Lab 35: 216 [syn sub Fr tamarisci sp obscura var obscura 928 Stephani Icon Hepat Inedit [Thyopsiella] F-178, microfiche 59 fig 4127 ["Japan, Makino 141"] - 4128 1155 Index type specimens in Nat Sci Mus, Tokyo: 13 948 Taiwania 33: 12 [Index of Taiwan] 521 J Hatt Bot Lab 68: 43 [annotated catalogue of China] 995 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 2006 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] = Frullania tamarisci ssp. obscura 213 849 930 (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 Page 260 malesiaca Verd. 1930 Typus: Indonesia. Java, Gunung Sumbing, 3350m 15.5.1927, vyn Leeuwen hb VERDOORN 8828, in hb FH Basionym: Distribution: spec nov - var nov Verdoorn Hattori 1930 1975 293 66 Ann Bryol Suppl 1: 59, fig 83-88 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 157-158 [ modification under alpine conditions] = Frullania campanulata var. malesiaca (Verd.) S.Hatt. 1975 manii Steph. by Bonner 1965 [orth var] Basionym: Distribution: Andaman Isl [India] orth var Bonner Stephani 1965 1985 Singh in Nath & Astana 2001 = Frullania mannii Index Hepat 5: 360 Stephani Icon Hepat Inedit [Thyopsiella] F-175b, microfiche 59 fig 4120 ["olim Frull manii St nsp, Andaman, Leg Man 4/8. 1890, Levier 92"] 1374 Perspectives Indian Bryology: 341[endemic to Andaman Island] Steph. 1894 Frullania recurvata Lehm. 1838 Frullania gaudichaudii mannii 655 928 (Nees & Mont.) Nees & Mont. 1845 Steph. 1894 Typus: Andaman Isl. near Port Blair 4.8.1890, E.H. Man ex hb LEVIER 92, in hb G Basionym: Distribution: Andaman Isl (Sarangani)[India] spec nov - syn nov Stephani Stephani Verdoorn Bonner Miller, Whittier & Whittier = Frullania recurvata 1894 1911 1930 1965 1983 228 686 549 655 609 Hedwigia 33: 151 Spec Hepat 4: 685 [as "manii", syn sub Fr. recurvata] Nova Guinea 14 (4): 541 Index Hepat 5: 360 Prod Flora Hepat Polynesiae: 131-132 Lehm. 1838 Frullania gaudichaudii (Nees & Mont.) Nees & Mont. 1845 Page 261 maritima Steph. 1911 [pp] {specimen from Acores; incl lectotype} Typus: in hb G 13922 [lectotypus] Basionym: Distribution: spec nov - syn nov Stephani Arnell Arnell Bonner Grolle 1911 1961 1961 1965 1970 Liu Bisang, Schumacker, Sérgio & Grolle Piippo Wang & Tau 1985 1989 = Frullania tamarisci maritima 1990 1992 686 180 180 655 615 Spec Hepat 4: 568 Svensk Bot Tidskr 55 (2): 392 [Tenerife] Svensk Bot Tidskr 55 (2): 392 [Canary Isl] Index Hepat 5: 361 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 312-313 [lectotypification; pp. incl lectotype] 2801 Invest Stud Nat 5: <133-155> [aus 698] 276 Giorn Bot Ital 122: 264 [syn sub Fr. tamarisci] 521 J Hatt Bot Lab 68: 43 [annotated catalogue of China] [aus 698] (L.) Dumort. 1835 von Konrat & Braggins in von Konrat & Pickering 2003 [nom nud] Basionym: Distribution: New Zealand (Stewart Isl), Auckland Isl [New Zealand] Konrat & Pickering 2003 Konrat & Brennan 2004 1000 http://www.discoverlife.org/nh/tx/Plantae/Bryophyta/ [IDnature guide for Liverworts, Frullania, New Zealand] 1164 http://www.liverworts.org/dich_key/frul_key.shtml, 5 of 6 pages [Key to Frullania of New Zealand; Stewart Isl, Auckland Isl] as Fr "maritimus" = maritima Steph. 1911 [pp] {specimen from Scotland} Typus: Scotland. Moidart 25.4.1900, Macvicar hb STEPHANI 2, in hb G sub Fr. fragilifolia Basionym: Distribution: spec nov - syn nov Stephani Müller in Rabenhorst Müller in Rabenhorst Bonner Grolle Stephani 1911 1915 1958 1965 1970 1985 Damsholt 2002 = Frullania fragilifolia maritimus 686 1064 1028 655 615 928 Spec Hepat 4: 568 Krypt Flora (2ed) 6, 2: 619-622 [syn sub Fr fragilifolia] Krypt Flora (3ed) 6, 2: 1240 Index Hepat 5: 361 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 312-313 Stephani Icon Hepat Inedit [Thyopsiella] F-195, microfiche 60 fig 4154 ["Ascotia, Maidart, Macvicar 25/4. 1900"] 1020 Illust Fl Nordic Liverworts & Hornworts: 604-601, fig 243, pl 230(1-9) [Northern Europe] (Taylor) Gottsche, Lindenb. & Nees 1845 [non Austin 1879] von Konrat & Brennan 2004 [orth var] Basionym: Distribution: Konrat & Brennan = Frullania maritima 2004 1164 http://www.liverworts.org/dich_key/frul_key.shtml, 5 of 6 pages [Key to Frullania of New Zealand; Stewart Isl, Auckland Isl] von Konrat & Braggins in von Konrat & Pickering 2003 [nom nud] Page 262 martiana Gottsche 1845 Typus: Brazil. Minas Geraes. sd, Martius in hb NEES, typus in hb S (B24133, 24134) Basionym: Distribution: spec nov - syn nov Gottsche in G., L. & N. Stephani Frye & Clark Schiffner & Arnell Bonner Yano Stephani 1845 1911 1947 1964 1965 1984 1985 Yano Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Gradstein & Costa Yano 1999 2002 = Frullania obcordata martii 2003 2008 2850 686 204 198 655 501 928 Syn Hepat: 448 Spec Hepat 4: 637 Univ Wash Publ Biol 6 (5): 760 [syn sub Fr obcordata] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 87 [Brazil] Index Hepat 5: 361 J Hatt Bot Lab 56: 505 [checklist Brazil] Stephani Icon Hepat Inedit [Diastaloba] B-33, microfiche 48 fig 32663267 ["Brasilia Petropolis, Rudolph leg. 1890"] 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 984 Mem New York Bot Gard 87: 86 [Brazil] [sub Fr caulisequa] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr caulisequa; Brazil; catalog] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Gottsche by Ångstr. 1876 [orth var] Basionym: Distribution: Ångström = Frullania martiana 1876 Öfversigt K Vetensk Akad Förh 33 (7): 89 [Brazil] Gottsche 1845 Frullania obcordata martinica 579 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1910 Typus: Martinique. -.8.1901, Duss 662 pp in hb G 20971 [holotypus] Basionym: Distribution: Mexico, Martinique spec nov - syn nov Stephani Bonner Yuzawa Stephani 1910 1965 1983 1985 686 655 498 928 Fulford & Sharp Yuzawa 1990 1991 105 525 = Frullania rio-janeirensis Spec Hepat 4: 335 Index Hepat 5: 361 J Hatt Bot Lab 54: 115-116, fig 6(a-m) [from type] Stephani Icon Hepat Inedit [Chonanthelia] A- 41, microfiche 46 fig 3128 [from type; "Martinique Duss 662 pp Aug 1901"] Mem NY Bot Gard 63: 44 [Mexico] J Hatt Bot Lab 70: 254 [syn sub Fr. riojaneirensis] (Raddi) Ångstr. 1876 Page 263 mastigophoroides S.Hatt. & Kamim. 1971 Typus: Borneo, Sabah. Mt. Kinabalu, on fallen tree branches along open and dry trail-side between Tenompock and Kambaranga Radio Station, 4500 ft 16/5/1963, Z. Iwatsuki 3412 in hb NICH 253412 Basionym: Distribution: Hattori & Kamimura 1971 532 Hattori 1975 409 Hattori Schuster 1976 1992 80 603 = Frullania clemensiana mathanii J Hatt Bot Lab 34: 429-436, fig. 1-5 [Steerea mastigophoroides sp. nov. from Mt. Kinabalu, Borneo] Misc Bryol Lichen 7 (1): 13-14 [comb. nov. Steerea clemensiana as. syn. nov. Sub St. Mastigophoroides] J Hatt Bot Lab 40: 469-470 [Mt. Kinabalu, as Steerea clemensiana] Hepat N America 5: 32 [sg. nov. Steerea (HATT & KAM) SCHUSTER} Verd. 1932 Steph. 1911 Typus: Ecuador 1896, Mathan sn ex hb LEVIER 4242, in hb G [holotypus G 20618] Basionym: Distribution: Ecuador, Galapagos Isls [Ecuador], Colombia spec nov - syn nov Barbour Stephani Irmscher Bonner Gradstein & Hekking Gradstein & Weber Menzel 1902 1911 1914 1965 1979 1982 1984 Stephani 1985 Gradstein in Zizka & Klemmer Uribe & Gradstein Uribe & Rangel-Ch Uribe 1994 1998 2000 2008 = Frullania peruviana 2233 Bryologist 5(1): 4 [USA] 686 Spec Hepat 4: 602 [Peru, Ecuador] vide 483: 997 655 Index Hepat 5: 361 483 J Hatt Bot Lab 45: 113 [Cat Colombia] 496 J Hatt Bot Lab 52: 148 [on 3 Galapagos Isls] 142 Willdenowia 14: 506 [not in Peru, but in Ecuador - error by Stephani 1911: 602] 928 Stephani Icon Hepat Inedit [Meteriopsis] E-28, microfiche 55 fig 3806 ["Ecuadoe, Mathan, Levier 4242"] - 3807 1118 Kl Senckenbergreihe 20: 60 [Galapagos] 707 Bryophyt Bibl 53: 37 [Catalogue Colombia] 1090 Colombia, diversidad biótica III, Bogota: 445 [Colombia] 1513 Caldasia 30(1): 75 [syn sub Fr peruviana] [Monography of subgenus Meteoriopsis] Gottsche 1846 [non Gottsche ex Spruce 1884] Page 264 mayebarae S.Hatt. 1952 Typus: Japan. Kyusyu, Prov. Higo, Hitoyoshi, prope flum Kuma, 80m 13.7.1947, K. Mayebara 500 in hb NICH [holotypus] in hb GZU; materia originalis in EGR ?? "Kiushiu. Kumamoto: Ohno, sides of River Kuma, on rocks often wet or submerged. Coll. K. Mayebara, VII. 1950, Det. S.H." in hb US - in hb COLO Basionym: Distribution: Korea (South), Japan, India (West Himalaya) spec nov Hattori Hattori Hattori Hattori Vanden Berghen Hattori & Noguchi Kamimura 1950 1951 1952 1952 1960 1960 1961 Bonner Hong Yamada Hattori Kodama Iwatsuki & Mizutani Yamada 1965 1966 1970 1971 1972 1972 1974 655 1107 Choe Hattori & Lin Piippo 1980 1985 1990 1171 505 521 Parihar, Lal & Katiyar NN 1994 1996 2196 632 Bapna & Kachroo Suanjak 2000 2001 712 979 860 926 529 1733 965 199 1375 966 406 Kis 2004 Yamada & Iwatsuki 2006 Mizutani, Hasegawa & Iwatsuki 2009 1377 1470 2112 NN 2627 = Frullania inflata 2011 Exsicc Hepaticae Japonicae, ser 3, No 125 J Hatt Bot Lab 5: 82, pl 7(17) [oil bodies] Bot Mag Tokyo 65: 13-14, fig 42(a-k) J Hatt Bot Lab 8: 30 [Honshu, Kyushu] Bull Chichibu Mus Nat Hist 10: 85-87 [in <Chichibu] Index spec Typ herb Japon, Bryophyta: 17 [holotypus] J Hatt Bot Lab 24: 85-87, fig 30(1-7), map 9 [Type Joc. Hitoyoshi, Kumamoto Pref., Japan.] Index Hepat 5: 361 Bryologist 69 (1): 411-412 [South Korea] Misc. Bryol.Lichen. 5 (8): 144 [form of oil-body] Exsicc Hepaticae Japonicae, ser 18, No 866 Spec Publ Osaka Mus Nat Hist 4: 127, fig 66(g-h [Central Japan] Colour Illust Bryo Japan, Osaka: 334, fig 171 [722] Misc Bryol Lichen 6 (9): 163-164, fig 1(a-e) [comb nov as Fr. inflata var mayebarae] Illust Fl & Fauna Korea 24: 733 [sub Fr inflata] J Hatt Bot Lab 59: 136 J Hatt Bot Lab 68: 43 [syn sub Fr. inflata; annotated catalogue of China] Hepat Anth India, annotated checklist: 22 [Western Himalaya] gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 22 of 35 [in hb US] Hepaticology in India II: 173 [syn sub Fr. inflata] http://www.kfunigraz.ac.at/botanik/gzu-hepaticae-typen2.htm [in hb GUZ] F Hist Nat Mus Matraensis 28: 15 [materia originalis in EGR] J Hatt Bot Lab 99: 31 [Catalog of Japan] Type sepecimens Hatt Bot Lab (NICH) 29: Holotype. Japan, Kyushu, Kumamoto-ken, Prov. Higo, Hitoyoshi. On moist rock. 13 July 1947. K. Mayebara 500 (NICH 11567). in hb COLO [isotype] B-27653 BarCode: 963561 Gottsche 1845 [non Herzog 1921] mayebarae var. concava Kamim. 1961 Typus: Japan. Tosa, Erazawa -.4.1947, Kamim. 14843 in hb NICH Basionym: Distribution: var nov - syn nov Kamimura Bonner Choe Hattori & Lin Bapna & Kachroo Yuzawa Yamada & Iwatsuki Mizutani, Hasegawa & Iwatsuki = Frullania inflata 1961 1965 1980 1985 2000 2001 2006 2009 199 655 1171 505 712 1660 1470 2112 J Hatt Bot Lab 24: 87, fig 30(8-13), map 9 Index Hepat 5: 362 Illust Fl & Fauna Korea 24: 734 [syn nov sub Fr inflata] J Hatt Bot Lab 59: 136 Hepaticology in India II: 173 [syn sub Fr. inflata] Natural Environmental Sci Res 14: 16 [Japan, syn sub Fr inflata] J Hatt Bot Lab 99: 31 [Catalog of Japan] Type sepecimens Hatt Bot Lab (NICH) 29: Holotype. Japan, Shikoku, Kochi-ken, Takaoka-machi. On bark. 5 Jan. 1937. M Kamimura 14843 (NICH 46877). Gottsche 1845 [non Herzog 1921] Page 265 megalocarpa Taylor 1846 Typus: Tasmania. "Van Diemen's Land" in hb GREVILLE Basionym: Distribution: spec nov - syn nov Taylor Taylor in G., L. & N. Mitten in Hooker Gottsche in Mueller Bastow Stephani Bonner Hattori Scott & Bradshaw Hattori Carthy McCarthy = Frullania rostrata megalostipa 1846 1847 1860 1880 1888 1911 1965 1983 1986 1989 2003 2006 292 2848 561 1489 845 686 655 499 56 499 1125 1385 London J Bot 5: 404 Syn Hepat: 779 Antarctic voyage, III. Fl Tasmaniae 2: 237-238 [Tasmania] Frag Phyt Australia 11, suppl Hepaticae :65 [Tasmania] Papers Proceed Roy Soc Tasmania for 1887: 265 [Tasmania] Spec Hepat 4: 560 Index Hepat 5: 362 J Hatt Bot Lab 54: <133-182> Brunnonia 8 (1): 36 [Australia] J Hatt Bot Lab 54: 164 [syn sub Fr. rostrata] Flora Australia Suppl Ser 21: 46 [sub Fr rostrata] Checklist Australian Liverworts [Australia, syn sub Fr rostrata] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Spruce 1884 Typus: Ecuador: Andes Quitenses, flum. Pastasa, Antombos sd, Spruce sn in hb G [isotypus] Basionym: Distribution: Panama, Ecuador, Galapagos Isl, Venezuela, Bolivia spec nov comb nov Spruce Stephani Stephani Bonner Stephani 1884 1910 1916 1965 1985 Yuzawa Atwood, Espinoza-Prieto & Gradstein Gradstein 1991 2018 243 686 222 655 928 Trans Proc Bot Soc Edinburgh 15: 11 & 15-16 (No 2) Spec Hepat 4: 335 Biblioth Bot 87: 240 [Bolivia] Index Hepat 5: 362 Stephani Icon Hepat Inedit [Chonanthelia] A- 42ab, microfiche 46 fig 3129-3130 [from isotype; Andes Quitenses Original leg Spruce Ecuador Hb Renauld] 525 J Hatt Bot Lab 70: 250-252, fig. 36, map 6 a(:285) 5013 Bry. Div. Evo. 40 (2): 68-78 [Key to Frull sect. Cladocarpicae, p. 75) 2020 in press [as Fr. rio-janeirensis var. megalostipa] = Frullania rio-janeirensis var. megalostipa (Spruce) Gradstein 2020 mehrana S.Hatt. 1974 [nom nud] {Philippines} Basionym: Distribution: Hattori 1974 544 J Hatt Bot Lab 38: 171 [as "in press"] [Philippines, Luzon, Mt Pulog] = meridiana Steph. by Stotler 1969 [orth var] Basionym: Distribution: Stotler Stotler = Frullania meridana 1969 1970 4761 Nova Hedwigia 18: 469-470 194 The genus Frullania, separata: 469-470 Steph. 1911 Page 266 merrillana Steph. 1911 Typus: Philippines. Luzon sd, Merrill 19351pp in hb G [holotype] Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Kamimura Bonner Stephani 1911 1930 1961 1965 1985 Uribe 2008 Uribe-M. 2011 = Frullania tenuicaulis Frullania vethii Spec Hepat 4: 613 Ann Bryol Suppl 1: 151 [syn sub Fr tenuicaulis] J Hatt Bot Lab 24: 61 [syn sub Fr tenuicaulis] Index Hepat 5: 363 Stephani Icon Hepat Inedit [Meteriopsis] E-55, microfiche 56 fig 3851 ["Luzon, leg Merrill 19351 pp"] 1513 Caldasia 30(1): 90, [belongs to subgen Diastaloba] [Monography of subgenus Meteoriopsis] 2767 Caldasia 33(1): 76 [type study, syn sub Fr trichodes] "syn. Fide Kamimura (1961). Type: Philippines, Luzon, without collector (holotype, G!)." Mitt. 1891 Sande Lac. 1884 Frullania trichodes merrilleana 686 293 199 655 928 Mitt. 1862 Steph. by Horikawa 1934 [orth var] Basionym: Distribution: Horikawa = Frullania merrillana 1934 Mitt. 1891 Sande Lac. 1884 Frullania trichodes mertensiana J Sci Hiroshima Univ, ser B, div 2, 2: 236 Steph. 1911 Frullania tenuicaulis Frullania vethii 34 Mitt. 1862 [auct. non Lindenb. & Gottsche] Steph. 1910 {Hawaii} Basionym: Distribution: auct - syn nov Stephani Miller Miller, Whittier & Whittier = Frullania hypoleuca 1910 1963 1983 217 834 609 Denkschr K Akad Wiss, Math Nat Kl, Wien 85: 193 [Hawaii] Ark Bot, ser 2, 5: 519 [syn nov sub Fr. hypoleuca] Prod Flora Hepat Polynesiae: 132 Nees 1843 Page 267 mertensiana Lindenb. 1845 Typus: Mariana: "In Insula Marianis & Carolinis" sd, Mertens sn in hb LE (Acad Petropol) 20, in hb S (B24137, B24138) [typus] Basionym: Distribution: Northern Marianas, Micronesia [=Caroline Isl] spec nov - syn nov Lindenberg in G., L. & N. Jardin 1845 1858 Stephani Verdoorn Kühnemann Kühnemann Clark Miller Bonner Miller Miller, Whittier & Whittier Stephani 1911 1930 1937 1949 1955 1960 1965 1968 1983 1985 Engel Myrdal 1988 2002 = Frullania apiculata mesophila 2850 Syn Hepat: 446-447 (No 77) 3821 Essai d'une flore de l'archipel des Marquises: 12 [Marquesas Islands, French Polynesia, France] 686 Spec Hepat 4: 664 [Falkland Is. = error, vide Engel 1988] 293 Ann Bryol Suppl 1: 97 [syn sub Fr. apiculata] 1031 Revista Centro Estud Doct Ci Nat, Buenos Aires 1 (3): 167 [Malvinas] 657 Lilloa 19: 355 [Falkland Isl] 369 Bryologist 58: 68 920 Bryologist 63: 121 [Micronesia] 655 Index Hepat 5: 363 [Falkland = error] 672 Micronesia 4 (1): 74 [key to Guam & Northern Micronesia] 609 Prod Flora Hepat Polynesiae: 132 928 Stephani Icon Hepat Inedit [Diastaloba] B-45, microfiche 49 fig 3283 ["Falkland Inseln (Marianen) - Marianen (Mertens legit)"] 111 Fieldiana Bot NS 25: 164 [Mariana, not Falkland Is.] 1276 Mosskollekter [Frullania types in hb S] (Reinw., Blume & Nees) Nees 1845 Gradstein ex Gradstein & Weber 1982 [nom nud] Typus: Ecuador. Galapagos Isl. Santa Cruz, along the old trail to Bella Vista, dense tufts on twigs of Zanthoxylum fagara Gradstein & W.A. Weber H3 11/4/1976, S.R. in hb F C0000138F - Cryptogamic Herbarium 1037140 - INR 1767015 & 44932 [holotypus] > see fr1368.jpg Basionym: Distribution: Galapagos Isls [Ecuador] nom nud Gradstein & Weber Yuzawa, Mues & Hattori Yuzawa Gradstein = Frullania gradsteinii meteoroides Typus: Fiji. 1982 1987 1988 2000 496 509 0 J Hatt Bot Lab 52: 148 [on 2 Galapagos Isls] J Hatt Bot Lab 63: 429, fig. 4 [as Fr gradsteinii published] note in hb F: ""Holotype Fr. gradsteinii" July 19, 1988 see fr1368.jpg Email from 14.1.2000 [wurde als Fr. gradsteinii YUZAWA et alii 1987] beschrieben Yuzawa, Mues & S.Hatt. 1987 Mitt. 1862 1860, B. Seemann 834 [Milne in hb HOOKER ?], in hb G Basionym: Distribution: Indonesia (Java), Philippines, Samoa (Savaii, Upolu), Fiji spec nov - syn nov Mitten Mitten in Seemann Stephani in Reinecke Stephani Stephani Verdoorn Verdoorn Bonner Grolle & Schultze-Motel Stephani = Frullania serrata 1862 1871 1896 1908 1911 1930 1930 1965 1972 1985 573 637 1044 216 686 293 551 655 537 928 Bonplandia 10 (2): 19 [Fiji] Flora Vitiensis: 417-418 [Fiji] Bot Jahrb 23: 315 [Samoa] Denkschr K Akad Wiss, Math Nat Kl, Wien 81: 298 Spec Hepat 4: 540-541 Ann Bryol Suppl 1: 85 Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 363 J Hatt Bot Lab 36: 84 [Samoa] [sub Fr. serrata] Stephani Icon Hepat Inedit [Thyopsiella] F-147, microfiche 59 fig 4084 ["Viti, Graeffe"] - 4085 Gottsche 1845 Page 268 meteriopsis Colombia)] ex Pando 2000 [nom herb in hb MA(-Hepat 851: Brazil, Hepat 917: Typus: in hb MA MA-Hepat 851: Brazil, Wasum 1987 & 917: Colombia, Cuarecasas 1932) Basionym: Distribution: Pando 2000 1273 Herbario de Criptogamia, Madrid (MA) = micholitzii Steph. 1894 Typus: Philippines. Luzon, Benguet 1884/1885, Micholitz 13 in hb G Basionym: Distribution: Vietnam, Philippines (Luzon), Solomon Isl spec nov - syn nov Stephani Stephani Verdoorn 1894 1911 1930 Bonner Pocs, Tixier & Jovet-Ast Hattori Miller, Whittier & Whittier Stephani 1965 1967 1982 1983 1985 = Frullania gracilis microceras Typus: Chile. 228 686 293 Hedwigia 33: 164 Spec Hepat 4: 645-646 Ann Bryol Suppl 1: 115-116, fig 181-183 [may be Fr. minor of Philippines] 655 Index Hepat 5: 364 1710 Bot Kozlem 54 (1): 30 [North Vietnam] 495 J Hatt Bot Lab 51: 217 [syn sub Fr. gracilis] 609 Prod Flora Hepat Polynesiae: 132 928 Stephani Icon Hepat Inedit [Diastaloba] B-67, microfiche 49 fig 33103311 [from type; "Luzon Benguet, Micholitz No 13 1884/85"] (Reinw., Blume & Nees) Nees 1845 Mont. ex Steph. 1894 [pro syn] sd, Gray in hb MONTAGNE, (? hb BESCHERELLE 222), in hb G [sub Fr. chilensis] Basionym: Distribution: - syn nov Stephani 1894 228 Bonner Stephani 1965 1985 655 928 = Frullania chilensis microloba Hedwigia 33: 142 [in hb MONTAGNE syn sub nomine: Frull. micoceras Mont ms] Index Hepat 5: 364-365 Stephani Icon Hepat Inedit [Galeiloba] C- 92a, microfiche 51 fig 3472 ["(Frull microceras Mont ms, circum Valdivia communis)" syn of Fr chilensis] Steph. 1894 Mont. ex Bonner 1965 [nom herb] [non Steph. 1910] Typus: Chile. Corral, Valdivia Krause original material in hb S (B24144, 24145) Basionym: Distribution: Bonner Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot = Frullania stipatiloba 1965 2002 655 782 Index Hepat 5: 365 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] Steph. 1894 Page 269 microloba Steph. 1910 [non Mont. ex Bonner 1965] Typus: New Caledonia 1906, Franc hb LACOUTURE 16, in hb G 18448 Basionym: Distribution: New Caledonia spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Hattori Miller, Whittier & Whittier Stephani = Frullania caledonica 1910 1930 1930 1965 1981 1983 1985 686 293 551 655 492 609 928 Spec Hepat 4: 437 Ann Bryol Suppl 1: 43 [syn sub Fr. squarrosa] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 365 J Hatt Bot Lab 49: 365-368, fig 5 [syn sub Fr. caledonica] Prod Flora Hepat Polynesiae: 132 Stephani Icon Hepat Inedit [Galeiloba] C-156, microfiche 52 fig 3550 [from type; "Nova Caledonia, lg Franc 1906, No 16 Herb Lacouture"] Gottsche ex Steph. 1894 [non (Schuster) Schuster 1992] Frullania squarrosula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 microphylla [auct. non (Gottsche in Gottsche & Rabenhorst) Pearson] Boesen, Lewinsk & Rasmussen 1975 {Faeroe Isl} Basionym: Distribution: - syn nov Boesen, Lewinsky & Rasmussen 1975 Damsholt 2002 Söderström, Hassel & Weibull Lindbergia 3 (1/2): <69-78> 1020 Illust Fl Nordic Liverworts & Hornworts: 604 [syn sub Fr tamarisci var cornubica] 2672 Distribution maps - Northwestern Europe, vol 1 : 52 [syn sub Fr tamarisci var cornubica, see K. Damsholt] 2002 = Frullania tamarisci var. cornubica Carrington 1878 microphylla fo. corticola Schiffn. 1936 Typus: Ireland. "Bei Killarney, an Arbutus-Bäumen" -.8.1911, D.A. Jones & J.B. Duncan sn in hb EGER [as fo. "corticola", isotypus] Basionym: Distribution: fo nov Schiffner Pocs = Frullania microphylla microscypha Typus: N-Amerika 1936 1976 2561 Krit Bemerk europ Leberm, ser 21: 4 [Hepat europ exsicc, nr. 1006 pp] 2686 Fol. Hist. nat. Mus. Matr. 4: 19 [types in hb EGER] fo. corticicola Schiffn. 1936 Taylor 1846 sd, J. Drummond in hb HOOKER in hb GLM (hb LEHMANN) 06025 Basionym: Distribution: spec nov - syn nov Taylor Taylor in G., L. & N. Evans Stephani Frye & Clark = Frullania eboracensis 1846 1847 1897 1910 1947 292 2848 237 686 204 London J Bot 5: 402 Syn Hepat: 777, No 59c Trans Connet Acad Arts Sci 10 (1): 18 Spec Hepat 4: 431 [syn sub Fr. eboracensis] Univ Wash Publ Biol 6 (5): 771 [sub Fr eboracensis] Lehm. 1844 Page 270 microstipula Steph. 1894 Typus: Burma. in monte Aracan 1888, Micholitz in hb G 14549 Basionym: Distribution: Burma spec nov Stephani Stephani Bonner Hattori 1894 1911 1965 1976 228 686 655 67 Stephani 1985 928 = Frullania gracilis microta Hedwigia 33: 164 Spec Hepat 4: 646 Index Hepat 5: 366 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 9-10, fig 171 [from type] [syn sub Fr. gracilis] Stephani Icon Hepat Inedit [Diastaloba] B-68, microfiche 49 fig 3312 [from type; "Birma, Aracan Hills, Micholitz 1888"] (Reinw., Blume & Nees) Nees 1845 C.Massal. 1897 Typus: China. Schensi, Lun-san-huo -.11.1895, J. Giraldi ex hb LEVIER, in hb G Basionym: Distribution: China spec nov - syn nov Massalono 1897 Levier Stephani Verdoorn in Handel-Mazzetti Horikawa Kamimura Bonner Choe 1906 1910 1930 1934 1961 1965 1980 Stephani 1985 Wu in Li, Gao, Wu et al. 1985 Piippo 1990 Yamada & Iwatsuki 2006 = Frullania japonica 1914 Mem Accad Agr Art Comm Verona, ser 3, 73 (2): 42-44, pl 12 fig 17(18) 824 Nuovo Giorn Bot Ital NS 13: 348-349 686 Spec Hepat 4: 448 550 Symb Sinicae 5: 37 [syn sub Fr. japonica] 34 J Sci Hiroshima Univ, ser B, div 2, 2: 237 [syn sub Fr. japonica] 199 J Hatt Bot Lab 24: 52 [syn sub Fr jackii ssp japonica] 655 Index Hepat 5: 366 1171 Illust Fl & Fauna Korea 24: 734 [sub Fr jackii ssp japonica; South Korea] 928 Stephani Icon Hepat Inedit [Galeiloba] C-157, microfiche 52 fig 3551 ["China, Giraldi (No 30) - ist wohl retusa Mitten ... Himalayas"] 1169 Bryoflora Xizang (Tibet), Science Press, Beijing: 525 [sub Fr jackii ssp japonica] 521 J Hatt Bot Lab 68: 43 [syn sub Fr. davurica; annotated catalogue of China] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] Sande Lac. 1864 Frullania jackii var. japonica (Sande Lac.) S.Hatt. 1957 Frullania jackii ssp. japonica Frullania davurica (Sande Lac.) S.Hatt. 1959 Hampe ex Gottsche, Lindenb. & Nees 1845 microta var. microphylla C.Massal. 1897 Typus: China. Schensi Prov., Pon-o-li -.10.1894, J. Tsan & J. Giraldi in hb G (ex hb LEVIER) Basionym: Distribution: var nov - comb nov Massalongo Levier Bonner Hattori & Lin 1897 1906 1965 1985 1914 824 655 505 Piippo 1990 521 = Frullania davurica Mem Accad Agr Art Comm Verona, ser 3, 73 (2): 43 Nuovo Giorn Bot Ital NS 13: 349 Index Hepat 5: 366 J Hatt Bot Lab 59: 133 [comb nov - davurica fo. microphylla (Mass.) Hatt. & Lin, comb. nov.] J Hatt Bot Lab 68: 43 [annotated catalogue of China] fo. microphylla (C.Massal.) S.Hatt. & P.J.Lin 1985 Page 271 mikawana Steph. ex S.Hatt. 1952 [nom herb] Typus: Japan: Okazaki, Mikawa 23.12.1912, S. Okamura 359 in hb G Basionym: Distribution: - syn nov Hattori Kamimura Bonner Stephani 1952 1961 1965 1985 Yuzawa 2001 = Frullania squarrosa J Hatt Bot Lab 8: 31 [syn sub Fr. squarrosa] J Hatt Bot Lab 24: 17 [syn sub Fr. squarrosa] Index Hepat 5: 366-367 Stephani Icon Hepat Inedit [Galeiloba] C-158, microfiche 52 fig 35523553 [from type; "Japonia, Okamura 359 - (Okamura legit)"] 1660 Natural Environmental Sci Res 14: 13 [Japan, syn sub Fr ericoides] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides mildbraedii 529 199 655 928 (Nees) Mont. 1839 Steph. 1911 Typus: Uganda. Ruwenzori, Batagu Tal, 2700m 1907-1908, A. F. v Mecklenburg Exped, Mildbraed 2659 in hb G Basionym: Distribution: spec nov Stephani in Mildbraed 1911 Stephani Stephani in Mildbraed 1911 1914 686 820 Demaret Bonner Vanden Berghen Stephani 1942 1965 1976 1985 1066 655 185 928 Wigginton 2002 1012 Wigginton Wigginton 2004 2004 1312 1172 = Frullania serrata minima Wiss Ergebn Deutsch Z Afr Exped 1907-1908, (Separata): ? 23-24, fig 47(a-b) [as spec nov] Spec Hepat 4: 480 Wiss Ergebn Deutsch Z Afr Exped 1907-1908, (Bot=2): 133-134, fig 47(a-b) [as spec nov] Bull Jard Bot Etat Bruxelles 16 (2): 305 Index Hepat 5: 367 Bull Jard Bot Nat Belg 46 (1/2): 111 Stephani Icon Hepat Inedit [Thyopsiella] F- 10, microfiche 56 fig 3884 ["Ruwenzoni, 2700m, .... , Exp A Fr v Mbg, Milbraed 2659"] Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 221 [syn sub Fr serrata, West Africa] Gottsche 1845 Steph. 1911 Typus: Guadeloupe. sd, Duss 197 in hb G Basionym: Distribution: Guadeloupe spec nov - syn nov - syn nov Stephani Pagán Bonner Stotler Stotler Stephani 1911 1942 1965 1969 1970 1985 Lavocat Bernard & SchäferVerwimp 2011 = Frullania neesii 686 3152 655 4761 194 928 Spec Hepat 4: 532 Bryologist 45(4): 109 [Guadeloupe] Index Hepat 5: 367 Nova Hedwigia 18: 450 [syn sub Fr. neesii] The genus Frullania, separata: 450 [syn sub Fr. neesii] Stephani Icon Hepat Inedit [Thyopsiella] F-117, microfiche 58 fig 4043 ["Guadeloupe, Duss No 197 (Prof Urban misit"] 3130 Cryptogamie, Bryologie 32(3): 243 [Guadeloupe] Lindenb. 1845 Frullania kunzei (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 272 minor Raddi 1818 [non Sande Lacoste 1854] Basionym: Distribution: Raddi Raddi Raddi Nees Nees Raddi in Nees Bonner Stotler = Frullania dilatata minor 1818 1820 1820 1838 1841 1841 1965 1968 1651 2874 261 164 164 655 197 Jungermanniografia etrusca [Seperate print]: 10 Mem Mat Fisica Soc Ital Sci, Modena 18 (1): 21 Preprint of Mem Mat Fisica Soc Ital Sci, Modena 18 : (1): 21 Naturg Europ Leberm 3: 217 [syn sub Fr. dilatata] Reprint of Jungerm Etrusca: 5, pl 2(3a-e) Jungermannogr Etrusca [Seperate print with remarks by Nees]: 5 Index Hepat 5: 367 Taxon 17: 635-638 (L.) Dumort. 1835 (Sande Lac.) Verd. 1929 Basionym: Frullania minor Distribution: Sande Lac. 1854 [non Raddi 1818] Verdoorn Verdoorn Verdoorn Verdoorn Hattori 1929 1930 1932 1932 1976 = Frullania gracilis ssp. minor Frullania gracilis minor 245 293 553 554 67 Ann Bryol 2: 147 Ann Bryol Suppl 1: 117-118 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 59-60 [Malay Peninsula, Java] Nederl Kruidk Arch Jrg 1932 (2): 497-498 [Borneo] Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 10-13, fig 172-173 (Sande Lac.) S.Hatt. 1975 (Reinw., Blume & Nees) Nees 1845 Sande Lac. 1854 [non Raddi 1818] Typus: Indonesia. Java in hb JUNGHUHN Basionym: Distribution: Indonesia (Java) spec nov emendavit Sande Lacoste Sande Lacoste in Dozy Sande Lacoste Sande Lacoste in Junghuhn Schiffner Schiffner Stephani Verdoorn Verdoorn Bonner Kamimura Hattori Stephani 1855 1856 1857 1857 1898 1900 1911 1929 1930 1965 1974 1975 1985 2038 564 430 1490 831 904 686 245 293 655 3692 547 928 Menzel 1988 512 = Frullania gracilis Nederl Kruidk Arch 3: 424 Plagiochila sandei Dozy: 11-12 Verh Konikl Akad Wetenschappen 5: 89-90, pl 18(1-10) Plantae Junghuhnianae, Hepaticae: 541-542 Conspect Hepat Archip Indici, Batavia: 330-331 Hedwigia 39: 207 [Java] Spec Hepat 4: 649-650 Ann Bryol 2: 147 Ann Bryol Suppl 1: 117-118 Index Hepat 5: 3676 Bull Kochi Gakuen J Coll 5: 82 [Java, Bogor Mt. Tjibodas] J Hatt Bot Lab 39: 293 [Malay Peninsula] Stephani Icon Hepat Inedit [Diastaloba] B-69, microfiche 49 fig 3313 ["Java"] J Hatt Bot Lab 65: 164 [Cat Borneo] (Reinw., Blume & Nees) Nees 1845 Page 273 minor var. exorta Verd. 1929 Typus: Indonesia. Java, Preanger Prov, supra Tjiburrum, prope Tjibodas, 2140m 2.5.1894, Schiffner 2332 Basionym: Distribution: var nov comb nov Verdoorn Verdoorn Verdoorn Bonner Hattori = Frullania gracilis 1929 1930 1932 1965 1976 245 293 553 655 67 Ann Bryol 2: 148, fig 7 [var nov] [Java] Ann Bryol Suppl 1: 118, fig 148 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 60 [Java] Index Hepat 5: 367 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 13 [comb nov] (Reinw., Blume & Nees) Nees 1845 minor var. integribracteola S.Hatt. 1975 Typus: Indonesia. Sumatra, Mt. Singgalang near BukitTingi -.7.1955, Meijer 7023 in hb L [typus], in hb NICH [isotypus] Basionym: Distribution: Indonesia (Sumatra) var nov Hattori 1975 Mizutani, Hasegawa & Iwatsuki 2009 - syn nov Hentschel et alii = Frullania gracilis 2015 547 J Hatt Bot Lab 38: 293 2112 Type sepecimens Hatt Bot Lab (NICH) 30: Isotype. Sumatra, W. Sumatra, Mt. Singfalang. July 1955. W. Meijer 7023 (NICH 43201). 4097 Phytotaxa 220 (2): 135 [sub Fr gracilis] (Reinw., Blume & Nees) Nees 1845 minor var. subsinuata Schiffn. in Verd. 1929 Typus: Indonesia. Java, Tjibodas -.1.1895, Massart 1198pp Basionym: Distribution: Verdoorn Bonner Hattori = Frullania gracilis 1929 1965 1976 246 655 67 Ann Bryol 2: 159 Index Hepat 5: 367-368 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 13 [comb nov] (Reinw., Blume & Nees) Nees 1845 minor var. subsinuata Schiffn. 1900 [nom nud] Basionym: Distribution: nom nud Schiffner 1900 904 Hedwigia 39: 207 = Frullania minor var. subsinuata Schiffn. in Verd. 1929 Frullania gracilis minor ssp. recurviloba (Reinw., Blume & Nees) Nees 1845 S.Hatt. 1975 Typus: Indonesia. Sumatra, Mt. Korincji, 1400m 29.7.1956, Meijer B-9055 in hb L [typus], in hb NICH [isotypus] Basionym: Distribution: Indonesia (Sumatra] ssp nov Hattori 1975 Mizutani, Hasegawa & Iwatsuki 2009 - syn nov Hentschel et alii = Frullania gracilis 2015 547 J Hatt Bot Lab 39: 293-294, fig 146 2112 Type sepecimens Hatt Bot Lab (NICH) 30: Isotype. Sumatra, Mt. Korinchi. 1400 m. 29 July 1956. W. Meijer B9055 (NICH 432020). 4097 Phytotaxa 220 (2): 135 [syn sub Fr gracilis] (Reinw., Blume & Nees) Nees 1845 Page 274 minuta Steph. 1911 [non Gottsche ex Bonner 1965] Typus: Philippines. Luzon, Benguet Prov, Mt. Pulog -.5.1909, E.D. Merrill 6416 in hb G 16563 Basionym: Distribution: Philippines (Luzon) spec nov - syn nov Stephani Verdoorn Bonner Hattori Miller, Whittier & Whittier Stephani = Frullania junghuhniana minuta 1911 1930 1965 1976 1983 1985 686 293 655 67 609 928 Spec Hepat 4: 640 Ann Bryol Suppl 1: 134 [syn sub Fr. junghuhniana] Index Hepat 5: 368 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21-22 Prod Flora Hepat Polynesiae: 132 Stephani Icon Hepat Inedit [Diastaloba] B-53, microfiche 49 fig 3293 [from type; "Philippinen, Luzon in monte Pulog, Merrill legit. Mai 1909"] Gottsche 1845 Gottsche ex Bonner 1965 [nom herb] [non Steph. 1911] Typus: Madagascar. Imerina sd, Hildebrandt sn ex hb B, in hb G [sub Fr. rutenbergii] Basionym: Distribution: Bonner 1965 = Frullania rutenbergii Index Hepat 5: 368 Gottsche 1882 Frullania apicalis minutifolia 655 Mitt. 1879 Steph. 1911 Typus: Japan. Tosa, Godaisan 26.11.1905, Uematsu 7 in hb G Basionym: Distribution: spec nov - syn nov Stephani Iishiba Hattori Hattori Kamimura Bonner Hattori & Lin Stephani 1911 1912 1951 1951 1961 1965 1985 1985 Yuzawa Yamada & Iwatsuki 2000 2006 = Frullania motoyana 686 888 528 528 199 655 505 928 Spec Hepat 4: 650-651 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] J Hatt Bot Lab 5: 65-66 J Hatt Bot Lab 5: 65-66 J Hatt Bot Lab 24: 75 [syn sub Fr. motoyana] Index Hepat 5: 368 J Hatt Bot Lab 59: 159 Stephani Icon Hepat Inedit [Diastaloba] B-70, microfiche 49 fig 3314 [from type; "Japan, lg Uematsu"] 1659 Natural Environmental Sci Res 13: 6 [Japan, syn sub Fr motoyana] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] Steph. 1911 Page 275 minutiloba Steph. 1911 Typus: Japan. Lyukyu Arch, Amami Isl, Oshima prope Naze 17.3.1898, J.B. Ferrie hb LEVIER 1418, in hb G Basionym: Distribution: spec nov Stephani Iishiba Hattori Bonner Hattori Stephani 1911 1912 1952 1965 1982 1985 Yamada & Iwatsuki 2006 = Frullania gaudichaudii minutissima 686 888 1508 655 495 928 Spec Hepat 4: 555 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] J Jap Bot 27(10): 315 [Japan] Index Hepat 5: 368 J Hatt Bot Lab 51: 242 Stephani Icon Hepat Inedit [Thyopsiella] F-179, microfiche 59 fig 4129 ["Japonia, leg Ferrié 1898 (Levier 1418)"] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] (Nees & Mont.) Nees & Mont. 1845 Colenso 1887 [pp] [non Pearson 1924, nec Schiffner 1900] Typus: New Zealand. County od Waipawa, banks of river Mangatawhainui, near Norsewood 1886, W. Colenso Basionym: Distribution: New Zealand spec nov - syn nov Colenso Stephani 1887 1892 634 210 Bonner Hamlin 1965 1972 655 659 = Frullania falciloba minutissima Trans Proc New Zealand Inst 19: 298 J Linn Soc Bot 29: 274 [pp syn sub Fr. falciloba & pp syn sub Fr. rostrata] Index Hepat 5: 368-369 Rec Dominion Mus 7(19): 262 [New Zealand] Lehm. 1844 Pearson 1924 [hom illeg] [non Colenso 1887, nec Schiffner 1900] Typus: Samoa. Tutuila,. Trail to Mt. Matafao -.6/7.1920, M.C. Collarino 382a Basionym: Distribution: Samoa (Tutuila) spec nov Pearson 1924 863 - nom nov Grolle & Schultze-Motel Miller Miller, Whittier & Whittier 1972 1981 1983 537 439 609 = Frullania matafaoica minutissima Papers Dept Marine Biol Carnegie Inst Wash 20 (=341): 140-141, fig 15(1-15) [Samoa: Tutuila] J Hatt Bot Lab 36: 83 [Samoa] [hom illeg] Phytologia 47 (4): 322 Prod Flora Hepat Polynesiae: 132 H.A.Mill. 1981 Colenso 1887 [pp] [non Pearson 1924, nec Schiffner 1900] Typus: New Zealand. River Mangatawhainui, Norsewood 1886, Colenso a 1500 ex hb K, in hb G Basionym: Distribution: spec nov - syn nov Colenso Stephani 1887 1892 634 210 Stephani Bonner Hamlin Hattori 1911 1965 1972 1979 686 655 659 485 = Frullania rostrata Trans Proc New Zealand Inst 19: 298 [?? nom nud ??] J Linn Soc Bot 29: 274 [another specimen of Fr. minutissima as syn sub Fr. falciloba] Spec Hepat 4: 661 [not mentioned there] Index Hepat 5: 368-369 Rec Dominion Mus 7(19): 262 [New Zealand] J Hatt Bot Lab 45: 352-353, fig 21(a-h) from type [syn sub Fr. rostrata] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Page 276 minutissima Schiffn. 1900 [nom nud] [non Colenso 1887, nec Pearson 1924] Typus: Indonesia. Java, Cibodas "Zjibodas" -.1.1895, Massarat 1199 Basionym: Distribution: nom nud - syn nov Schiffner Verdoorn 1900 1930 904 293 Bonner Hattori Hattori 1965 1982 1982 655 422 495 Hedwigia 39: 207 Ann Bryol Suppl 1: 134 [nom nud; syn sub Fr. junghuhniana var. minutissima] Index Hepat 5: 369 Misc Bryol Lichen 9 (6): 123-124 J Hatt Bot Lab 51: 233 = Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 minutistipula Steph. in Steph. & Watts 1914 Typus: Australia. NSW, Rodriquez Pass, Blackheath "mixed with Fr gracillima & Radula wattsiana" Watts 1001c sd, W.W. Watts 1001b - 4.1.1911 W.W. in hb G G00069216 - n° SIB 126393/1 - 18219 (sd, W.W. Watts 1001b) in hb NSW 251030 [isolectotypus] Basionym: Distribution: spec nov - syn nov Stephani in Stephani & Watts Bonner Hattori Hattori & Mizutani Stephani 1914 1965 1979 1982 1985 220 655 485 495 928 Scott & Bradshaw Brown, Ramsay & Seur NN 1986 1992 2000 56 1140 1085 Carthy McCarthy 2003 2006 1125 1385 = Frullania rostrata miradoremis J Proc Roy Soc New S Wales 48: 108 Index Hepat 5: 369 J Hatt Bot Lab 45: 352-353 J Hatt Bot Lab 52: 447 Stephani Icon Hepat Inedit [Diastaloba] B-71, microfiche 49 fig 3315 [from type; "Australie, Rodriquez Pass, Watts 1001b"] Brunnonia 8 (1): 36 [Australia] Telopea 5(1): 75 [types in hb NSW & hb SYD] http://plantnet.rbgsyd.nsw.gov.au/HerbLink/NSWtypes/srch.html [isolectotype in NWS] Flora Australia Suppl Ser 21: 46 [sub Fr rostrata] Checklist Australian Liverworts [Australia, syn sub Fr rostrata] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Steph. by Steph. 1911 [orth var] Basionym: Distribution: Stephani Beauverd in Stephani Fulford & Sharp = Frullania miradorensis N. 1847] Frullania obcordata 1911 1924 1990 686 909 105 Spec Hepat 4: 824 [errata] Spec Hepat 6: 571 [sphalm.] Mem NY Bot Gard 63: 44 [Mexico] Steph. 1911 [hom illeg] [non Lindenb. & Gottsche in G., L. & (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 277 miradorensis Steph. 1911 [hom illeg] [non Lindenb. & Gottsche in G., L. & N. 1847] Typus: Mexico. Veracruz, Mirador Liebmann 92 in hb G (ex hb GOTTSCHE); Basionym: Distribution: Mexico spec nov - syn nov Stephani Bonner Stephani 1911 1965 1985 Yuzawa Fulford & Sharp Schuster Winter & Schäfer-Verwimp 1988 1990 1992 2020 = Frullania obcordata moelleri 686 655 928 Spec Hepat 4: 632 [as "miradoremis"] Index Hepat 5: 369 Stephani Icon Hepat Inedit [Diastaloba] B-21, microfiche 48 fig 3251 [from type; "Mexico Mirador Liebmann 92 Herb Gottsche"] 511 J Hatt Bot Lab 64: 447-449, fig 7(from type) [syn sub Fr. caulisequa] 105 Mem NY Bot Gard 63: 44 [Mexico] 603 Hepat N America 5: 120 5742 Frahmia 19: 13 [syn sub Fr obcordata] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. by S.W.Arnell 1956 [orth var] Basionym: Distribution: Arnell = Frullania molleri 1956 Svensk Bot Tidskr 50 (4): 528 [Fernando Po] Steph. 1894 Frullania caffraria molleri 847 Steph. 1894 Steph. 1894 Typus: San Thome Isl. Coimbra sd, A. Moller 45 pp in hb G 17311 Basionym: Distribution: spec nov - syn nov? Stephani Stephani Arnell Bonner Vanden Berghen Stephani 1894 1910 1956 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania caffraria 228 686 847 655 185 928 Hedwigia 33: 146 Spec Hepat 4: 370 Svensk Bot Tidskr 50 (4): 528 [Fernando Po] Index Hepat 5: 370 Bull Jard Bot Nat Belg 46 (1/2): 185 Stephani Icon Hepat Inedit [Galeiloba] C- 25, microfiche 50 fig 3377 [from type; "San Thomé, Moller inter 45"] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 229, fig 150( E) [syn sub Fr caffraria, West Africa] Steph. 1894 Page 278 mollicula Spruce 1890 Basionym: Frullania rigescens var. mollicula Spruce 1884 Distribution: stat nov - syn nov - syn nov Spruce Stephani Clark & Frye Clark Bonner Stotler Stotler Stephani 1890 1911 1953 1955 1965 1969 1970 1985 898 686 355 371 655 4761 194 928 Stephani 1985 928 Gradstein, Meneses & Arbe 2003 975 = Frullania triquetra moluccensis Mem Torrey Bot Club 1: 119-120 Spec Hepat 4: 501 Bryologist 56: 210-213, fig 1-17 Bryologist 58: 149 Index Hepat 5: 370 Nova Hedwigia 18: 457-458 [syn sub Fr triquetra] The genus Frullania, separata: 457-458 [syn sub Fr triquetra] Stephani Icon Hepat Inedit [Thyopsiella] F- 72, microfiche 57 fig 3978 ["Bolivia, Rusby leg, Herb Slater"] Stephani Icon Hepat Inedit [Thyopsiella] F- 50, microfiche 57 fig 3944 ["olim Frull mollicula Spruce, Bolivia, Original, leg Rusby"] - 3945 ["Mexico (leg Liebmann) Original"] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr triquetra] Lindenb. & Gottsche 1847 Steph. 1911 [nom herb] Basionym: Distribution: Stephani 1911 686 Spec Hepat 4: 824 [im errata: page 749. No 17 Frullania moluccensis (nicht molucensis)] 1900 802 Bot Mag Tokyo 14 (no 162): 181 = moniliata Nees in Inoue 1900 Basionym: Distribution: Inoue = Frullania moniliata moniliata (Reinw., Blume & Nees) Mont. 1842 [auct. non (Reinw. et al. ) Mont.] Steph. 1897 Typus: Japan. Tosa Basionym: Distribution: auct - syn nov Stephani Levier Reimers Chen & Wu 1897 1906 1931 1965 Lai & Wang-Yang Gao & Chang Hattori & Lin 1976 1981 1985 = Frullania moniliata 213 824 229 Bull Herb Boissier 5: 79 [Japan, Tosa] Nuovo Giorn Bot Ital NS 13: 349 Hedwigia 71 (1/2): 35 [China] Observ Fl Hwangshan: 11 = Observationes ad Florulam Hwangshanicum 33 Taiwania 21 (2): 168 [Index to Taiwan] 1021 Fl Hep Chinae bor-orient: 152, fig 66(10-16) [China] 505 J Hatt Bot Lab 59: 161 (Reinw., Blume & Nees) Mont. 1842 Page 279 moniliata [auct. non (REINW. et al.) MONT] ex Konstantinova et al. 1992 Basionym: Distribution: Konstantinova, Potemkin & Schijakov = Frullania appendiculata 1992 651 Arctoa 1:102 [USSR] Steph. 1897 [non Steph. ex Bonner 1965] Frullania moniliata ssp. obscura fo. appendiculata (Steph.) Verd. 1930 Frullania tamarisci ssp. obscura (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 moniliata var. beta Gottsche, Lehm. & Nees 1845 Typus: Java Basionym: Distribution: Gottsche, Lehmann & Nees Schiffner = Frullania moniliata 1845 1898 2850 Syn Hepat: 440 831 Conspect Hepat Archip Indici, Batavia: 332 (Reinw., Blume & Nees) Mont. 1842 moniliata var. commutata Nees 1845 Basionym: Distribution: Ness in G., L. & N. Bapna & Kachroo 1845 2000 2850 Syn Hepat: 440 712 Hepaticology in India II: 195 [syn sub Fr. tamarisci ssp obscura] = Frullania tamarisci ssp. obscura var. vietnamica (S.Hatt.) S.Hatt. 1985 moniliata var. ocellata Herzog ex Zwickel 1932 [nom herb] Typus: Indonesia. Java. Cibodas ("Tjibodas") -.10.1924, Goebel in hb HERZOG, Jena Basionym: Distribution: Zwickel 1932 663 Beih Bot Centralbl 49: 637 [Ozellen] = Page 280 moniliata var. rotundata Horikawa 1935 Typus: Japan. Honshu, miyajima Isl, Aki Prov 15.4.1929, Y. Horikawa 169 Basionym: Distribution: var nov Horikawa 1935 - syn nov Kamimura Kamimura 1952 1952 - syn nov Sawaraki Sawaragi 1955 1955 Bonner Bonner Hattori & Lin Yamada & Iwatsuki 1965 1965 1985 2006 - syn nov = Frullania moniliata ssp. obscura 1675 Bot Mag Tokyo 49: 675 [Species F. moniliatae affinis, sed folia rotundata et apice rotundato-obtusa] 769 Acta Phytotax Geobot 14 (4): 109 873 Contrib Fl Hepat Shikokuensem, Kochi: 125-126 [syn sub Fr moniliatat ssp obscura] 1844 Jap J Ecol 5(1): 14-18 [Fr moniliata ssp. obscura] 1844 Ecol Soc Japan 5(1): 14-18 [syn sub Fr. moniliata ssp. obscura] [Japan] 655 Index Hepat 5: 370 655 Index Hepat 5: 372 505 J Hatt Bot Lab 59: 161 [syn sub Fr. tamarisci ssp. obscura] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] Verd. 1930 {nördl. Indomalaya} Frullania tamarisci ssp. obscura (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata moniliata ssp. breviramea (Reinw., Blume & Nees) Mont. 1842 [auct. non (Steph.) Verd.] Herzog & Noguchi 1955 {Taiwan} Basionym: Distribution: auct - syn nov Herzog & Noguchi Hattori & Lin Kuo & Chiang 1955 1985 1988 = Frullania tamarisci ssp. obscura 859 505 948 J Hatt Bot Lab 14: 43 [Taiwan] J Hatt Bot Lab 59: 161 [= Fr tamarisci ssp obscura] Taiwania 33: 12 [Index of Taiwan] (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata moniliata ssp. breviramea (Reinw., Blume & Nees) Mont. 1842 (Steph.) Verd. 1930 {Java, Sumatra} Basionym: Frullania breviramea Steph. 1911 Distribution: Sri Lanka, Indonesia (Sumatra, Java) comb nov Verdoorn 1930 Verdoorn 1931 Verdoorn Verdoorn Verdoorn Parihar Bonner Hattori Abeywickrama & Jansen Long & Rubasinghe 1932 1932 1934 1962 1965 1972 1978 2014 293 Ann Bryol Suppl 1: 76-80, fig 104 [map], 110, 114-119 {nördl. Indomalaya} 231 Ann Bryol 4: 140 [Hepat select crit, ser 2 no 81-84; Sumatra, Java] & 148, fig 5 553 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 58 [Sumatra, Java] 554 Nederl Kruidk Arch Jrg 1932 (2): 495-496 [Borneo Mt. Kinabalu] 240 Ann Bryol Suppl 4: 194-195 1584 Univ Allahabad studies, Bot sect 1-2: 18 [India: IO] 655 Index Hepat 5: 370 536 J Hatt Bot Lab 35: 22 [= Fr. tamarisci ssp. obscura var. breviramea] 794 MaB Nat Comm Sri Lanka, Publ 1: 5 [Sri Lanka] 3994 Ceylon Journal of Science 43(1): 4 [Sri Lanka] = Frullania tamarisci ssp. obscura var. breviramea (Steph.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 Page 281 moniliata ssp. obscura Verd. 1930 {nördl. Indomalaya} Basionym: Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 Distribution: China, Taiwan, Japan, India (South), Sri Lanka, Philippines ssp nov Verdoorn Verdoorn in Handel-Mazzetti Verdoorn Verdoorn Hattori Horikawa Hattori Kamimura Hattori Sawaragi Hattori Hattori, Iwatsuki & Kanno Iwatsuki & Hattori Hattori Hattori Abeywickrama Inoue Bonner Pocs, Tixier & Jovet-Ast Tixier Lai & Wang-Yang Abeywickrama & Jansen Choe Hattori & Lin Kuo & Chiang Piippo Yamada & Choe Yamada & Iwatsuki Long & Rubasinghe 1930 1930 1931 1934 1944 1951 1951 1952 1952 1955 1955 1956 1956 1957 1959 1959 1961 1965 1967 1973 1976 1978 1980 1985 1988 1990 1997 2006 2014 = Frullania tamarisci ssp. obscura 293 550 230 241 943 1698 528 873 529 1844 1683 862 861 858 857 200 964 655 1710 678 33 794 1171 505 948 521 717 1470 3994 Ann Bryol Suppl 1: 80-81, fig 104, 112-113, map {nördl. Indomalaya} Symb Sinicae 5: 42 Ann Bryol 4: 123 [Hepat select crit, ser 1 no 36; Japonia] Ann Bryol Suppl 4: 205 [Indien] Bull Tokyo Sci Mus 11: 148 [Japan, key to Japanese species] J Sci Hiroshima Univ, serB, div 2 (Bot) 6(1): 27-38 J Hatt Bot Lab 5: 64 Contrib Fl Hepat Shikokuensem, Kochi: 126-127 J Hatt Bot Lab 8: 30 Ecol Soc Japan 5(1): 14-18, fig 1-5 [Japan] J Hatt Bot Lab 15: 91-92 [on serpentine rocks] J Hatt Bot Lab 16: 92 J Hatt Bot Lab 16: 86 J Hatt Bot Lab 18: 118 J Hatt Bot Lab 21: 128 [Japan] Ceylon J Sci (Bio Sci) 2 (1): 58 [Sri Lanka] J Jap Bot 36(5): 187 [Formosa] Index Hepat 5: 371 Bot Kozlem 54 (1): 30 [North Vietnam] Bull Mus Nat Hist Nat Paris, ser 3, 190, Bot 10: 86 [Sri Lanka] Taiwania 21 (2): 168 [Index to Taiwan] MaB Nat Comm Sri Lanka, Publ 1: 5 [Sri Lanka] Illust Fl & Fauna Korea 24: 734 [South Korea] J Hatt Bot Lab 59: 161 [= Fr. tamarisci ssp. obscura] Taiwania 33: 12 [Index of Taiwan] J Hatt Bot Lab 68: 44 [annotated catalogue of China] J Hatt Bot Lab 81: 287 [Korean Peninsula] J Hatt Bot Lab 99: 31 [Catalog of Japan] Ceylon Journal of Science 43(1): 4 [Sri Lanka] (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 moniliata ssp. obscura fo. alpina Verd. 1930 Typus: China. Yunnan, Likiang /or/ Chungtien 1914/1916, Handel-Mazzetti 3583 /or/ 1915, Handel-Mazzetti 6941 in hb E E00002542 (1.7.1914, Handel-Mazzetti 3585) [isosyntypus] Basionym: Distribution: fo nov - syn nov Verdoorn in Handel-Mazzetti Bonner Hattori & Lin Piippo = Frullania tamarisci 1930 1965 1985 1990 550 655 505 521 Symb Sinicae 5: 42 Index Hepat 5: 371 J Hatt Bot Lab 59: 161 J Hatt Bot Lab 68: 44 [annotated catalogue of China] fo. alpina Verd. ex Piippo 1990 [pro syn] Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 Page 282 moniliata ssp. obscura fo. appendiculata (Steph.) Verd. 1930 Basionym: Frullania appendiculata Distribution: comb nov - syn nov Steph. 1897 [non Steph. ex Bonner 1965] Verdoorn Verdoorn in Handel-Mazzetti Hattori Horikawa 1930 1930 1944 1951 293 550 943 1698 Sawaragi Bonner Gao & Chang Hattori & Lin Aur & Zhang Piippo Ji, Liu, Zhang, Chen & Lio 1955 1965 1981 1985 1985 1990 1999 1844 655 1021 505 1022 521 1208 = Frullania tamarisci ssp. obscura Ann Bryol Suppl 1: 81 Symb Sinicae 5: 42 [China] Bull Tokyo Sci Mus 11: 148 [Japan, key to Japanese species] J Sci Hiroshima Univ, serB, div 2 (Bot) 6(1): 28 [sub Fr moniliata ssp obscura] Jap J Ecol 5(1): 18 Index Hepat 5: 371 Fl Hep Chinae bor-orient: 152 [syn sub Fr. moniliata, #1612 or #1428] J Hatt Bot Lab 59: 161 [syn sub Fr tamarisci ssp obscura] Fl Heilongjiangensis 1: 163 [sub Fr tamarisci ssp moniliata] J Hatt Bot Lab 68: 44 [annotated catalogue of China] Chenia 6 (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 moniliata ssp. obscura fo. obtusiloba Verd. 1930 Basionym: Distribution: fo nov - syn nov Verdoorn in Handel-Mazzetti Bonner Hattori & Lin Piippo 1930 1965 1985 1990 = Frullania tamarisci ssp. obscura 550 655 505 521 Symb Sinicae 5: 43 Index Hepat 5: 371-372 J Hatt Bot Lab 59: 161 J Hatt Bot Lab 68: 44 [annotated catalogue of China] (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 moniliata ssp. obscura fo. parva Verd. 1930 Basionym: Frullania clavellata Distribution: fo nov - syn nov Mitt. 1891 Verdoorn Verdoorn in Handel-Mazzetti Sawaragi 1930 1930 1955 Bonner Hattori & Lin Piippo 1965 1985 1990 = Frullania moniliata ssp. obscura 293 Ann Bryol Suppl 1: 80 [fo nov] 550 Symb Sinicae 5: 43 [fo nov]] 1844 Ecol Soc Japan 5(1): 14-18 [syn sub Fr. moniliata ssp. obscura] [Japan] 655 Index Hepat 5: 371 505 J Hatt Bot Lab 59: 161 [syn nov] 521 J Hatt Bot Lab 68: 44 [annotated catalogue of China] Verd. 1930 {nördl. Indomalaya} Frullania tamarisci ssp. obscura (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 Page 283 moniliata ssp. obscura var. elongatistipula Verd. in Hand.-Mazz. 1930 Typus: China. Yunnan, 3700m (Tannenstämme .der ktp .St. des Rückens zwischen Haba und Dugwantsun se von Dschngdien, 3700 m, 24. VI. 1915 (5979) 24.6.1915, Handel-Mazzetti 5979 & 23.10.1915 Handel-Mazzetti 8331 Basionym: Distribution: var nov Verdoorn in Handel-Mazzetti Bonner Hattori Hattori & Lin 1930 1965 1972 1985 Piippo Bai 1990 2002 = Frullania elongatistipula 550 655 536 505 Symb Sinicae 5: 42, fig 14(11) Index Hepat 5: 371 J Hatt Bot Lab 35: 241-243 [syn sub Fr. elongatistipula] J Hatt Bot Lab 59: 161 [= Fr. tamarisci ssp. obscura var. elongatistipula] 521 J Hatt Bot Lab 68: 44 [annotated catalogue of China] 1189 Chenia 7: 23 [Yunnan, China] (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. elongatistipula (Verd.) S.Hatt. 1985 moniliata ssp. obscura var. elongatistipula fo. obtusiloba Verd. in Hand.-Mazz. 1930 Basionym: Distribution: fo nov - syn nov Verdoorn in Handel-Mazzetti Verdoorn = Frullania balansae 1930 1934 550 240 Symb Sinicae 5: 43 Ann Bryol Suppl 4: 195 [sub Fr balansae] Steph. 1894 Frullania tamarisci ssp. obscura var. balansae (Steph.) S.Hatt. 1972 Frullania moniliata monoliata (Reinwardt, Blume & Nees) Mont. by Parihar, Lal & Katiyar 1994 [ort var] Basionym: Jungermannia moniliata Distribution: orth var Reinw., Blume & Nees 1824 Parihar, Lal & Katiyar = Frullania moniliata mooreana (Reinw., Blume & Nees) Mont. 1842 1994 2196 Hepat Anth India, annotated checklist: 41 [syn sub Fr tamarisci] (Reinw., Blume & Nees) Mont. 1842 Steph. 1910 Typus: Tasmania. Wanderer River -.2.1900, J.B. Moore hb LEVIER 5795, in hb G 18223 Basionym: Distribution: Tasmania spec nov - syn nov Stephani Rodway 1910 1916 Bonner Hattori Stephani 1965 1979 1985 Scott & Bradshaw Ratkowsky 1986 1987 Hattori Carthy Glenny, Meagher & Brown 1988 2003 2004 McCarthy 2006 = Frullania falciloba 686 914 Spec Hepat 4: 422 Papers Proceed Roy Soc Tasmania for 1916: 136 [syn sub Fr. scandens] [Tasmania] 655 Index Hepat 5: 373 485 J Hatt Bot Lab 45: 349-350, fig 19(A-k) [from type /? syn ? 928 Stephani Icon Hepat Inedit [Galeiloba] C-216, microfiche 53 fig 3630 ["Tasmania, Wanderer River, J.B. Moore (Levier 5795)"] 56 Brunnonia 8 (1): 36 [Australia] 695 Papers Proceed Roy Soc Tasmania 121: 158 [Hattori per comm (IV): may be conspecific with Fr scandens; not correct] 429 Nova Hedwigia Beih 90: 148 [syn sub Fr falciloba; Australia] 1125 Flora Australia Suppl Ser 21: 43 [sub Fr falciloba] 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1385 Checklist Australian Liverworts [Australia; ß syn sub Fr falciloba] Lehm. 1844 Page 284 morokensis Steph. 1911 Typus: PNG. "Nova Guinea austro-orient. Brit., in montosis Mo-roka, 1300m. (distr. Moresby)Cental [British New Guinea]", Moroka Mts., 1300 m -.7.1893 - -8.1893, Lamberto Loria 853 in hb G G00067206 - n° SIB 129116/1 - 14237 (hb E. LEVIER 852) [holotypus] in hb JE (Loria 853) [isotypus] in hb FH (hb VERDOORN 3455) Basionym: Distribution: New Guinea (Papua New Guinea) spec nov Stephani Verdoorn Verdoorn Bonner Hattori 1911 1928 1930 1965 1976 686 242 293 655 80 - comb nov Hattori Grolle & Piippo Stephani 1980 1984 1985 490 143 928 = Frullania intermedia ssp. morokensis mucroloba Spec Hepat 4: 578 Rev Bryol NS 1: 114 Ann Bryol Suppl 1: 165, fig 289 Index Hepat 5: 373 J Hatt Bot Lab 40: 483-484 [difference to Fr. intermedia var. submorokensis] J Hatt Bot Lab 47: 194-195, fig 17(a-j) Acta Bot Fennica 125: 21 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Homotropantha] D- 5, microfiche 54 fig 3709 ["N Guinea, Moroka, legit Loria"] (Steph.) S.Hatt. 1980 Clark & Svihla 1949 Typus: COLOMBIA. Departamento de Norte de Santander, eastern Cordillera, Barrio Chucarima, Cinchona forest east of Rio Valegra, elevation 6,000 to 7,000 ft. 16.11.1942, Steere 7386 in hb WTU [type] in hb MICH [co-type] Basionym: Distribution: Colombia spec nov - stat nov Clark & Svihla Clark & Svihla Bonner 1949 1951 1965 332 341 655 Bryologist 52: 134-136, fig. 1-18(:135) Bryologist 54: 124-126 [= Fr bicornistipula var mucroloba] Index Hepat 5: 373-374 = Frullania bicornistipula var. mucroloba (Clark & Svihla) Clark & Svihla 1951 Frullania bicornistipula mucronata Spruce 1884 (Lehm. & Lindenb. in Lehm.) Nees in Mont. in Alcide d'Orbigny 1839 Basionym: Distribution: - syn nov Nees in Montagne in Alcide d'Orbigny Gottsche in G., L. & N. Yano Costa, Alencar Imbassahy & Silva = Frullania cylindrica 1839 257 d'Orbigny, Voyage l'Amerique Merid 7 (2): 68-69 1845 1995 2005 2850 Syn Hepat: 458-459 [syn sub Fr. cylindrica] 2095 J Hatt Bot Lab 78: 142 [annotated checklist of Brazil] 1382 J Hatt Bot Lab 98: 284 [Rio de Janeiro State, Brazil] Gottsche ex Lehm. 1844 Frullania brasiliensis Raddi 1822 Page 285 mucronata (Lehm. & Lindenb. in Lehm.) Lehm. & Lindenb. 1845 Basionym: Jungermannia mucronata Lehm. & Lindenb. 1834 Distribution: Guatemala, Costa Rica, Cocos Isl [Costa Rica], Panama, Colombia, Venezuela, Peru, Ecuador,, Guyana, Surinam, Brazil, Chile, Argentina comb nov - syn nov Lehmann & Lindenberg in G., L. & N. Stephani Schiffner & Arnell Stotler Stotler Gradstein & Hekking Cole Menzel Yano Stephani 1845 2850 Syn Hepat: 461-462 1911 1964 1969 1970 1979 1983 1984 1984 1985 686 198 4761 194 483 1407 142 501 928 Gradstein & Florschütz-de Gradstein & Hekking Cornelissen & Gradstein Pôrto Morales Z. Buck Boggan, Funk, Kelloff, Hoff, Cremers & Feuillet Uribe & Gradstein Bastos, Stadmann & BoasBastos Stotler, Allen, Gradstein, McGuinness, Whittemore & Chung Yano Gradstein Dauphin Gradstein & Costa Nöske, Gradstein, Kürschner, Parolly & Torracchi Uribe & Gradstein Holz 1989 1989 1990 1990 1991 1994 1997 Spec Hepat 4: 514-515 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 88 [Brazil] Nova Hedwigia 18: 446-447, pl 13 The genus Frullania, separata: 446-447, pl 13 J Hatt Bot Lab 45: 113 [Cat Colombia] Brenesia 21: 154 [Central America] Willdenowia 14: 490 [Peru] J Hatt Bot Lab 56: 505 [checklist Brazil] Stephani Icon Hepat Inedit [Thyopsiella] F- 75, microfiche 57 fig 3981 ["Brasilia, .... , leg Pabst 285"] 1237 Trop Bryol 1: 42 [Venezuela, Mt Roraima] 517 J Hatt Bot Lab 66: 209 [Guyana, Suriname] 729 Trop Bryol 3 (2): 33 [Guyana] 77 Crypt Bryol Lichen 11 (2): 122-123 [Brazil] 307 Trop Bryol 4: 38 [Costa Rica] 28 Bryologist 97 (4): 422 [Vainio's specimen from Brazil at TUR] 1396 Checklist Guianas: 10 [Guyana, Surinam] 1998 1998 707 701 Bryophyt Bibl 53: 37 [Catalogue Colombia] Trop Bryol 15: 19 [Brazil, Bahia] 1998 703 Trop Bryol 15: 179 [Checklist Panama] 1999 1999 1999 2003 2003 1263 1215 1087 984 1084 Briophyta do estado de São Paulo. Lista provisória [Brazil] Bryobrothera 5: 85 [Costa Rica on cacao trees] Rev Biol Trop 47 (3): 323 [Cocos Isl, Costa Rica] Mem New York Bot Gard 87: 89, fig 27(e-h) [Brazil] Crypt Bryol 24 (1): 25 [Reserva Biologica San Francisco, Ecuador] 2003 2004 Nöske Porto, Germano & Borges Parolly & Kürschner 2004 2004 2004 Kürschner & Parolly 2005 Costa, Imbassahy & Silva Yano 2005 2008 1071 Crypt Bryol 24 (3): 200 [syn sub Fr brasiliensis] 1184 [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm 1383 Kryptogam Epipyten S-Ecuador [Thesis Goettingen]: 124 & 132 1446 Brejos de Altitude em Pernambuco e Paraiba: 83 [Brazil] 1217 Nova Hedwigia 79(3-4): 386-389, 392, 402, 414, 416 tab 1, 2, 4 & 7 [Podocarpus NP, Ecuador, epiphytic: Fr mucronata community] 1362 Nova Hedwigia 80 (1-2): 92, 94 [Frullania mucronata community diagnostic species; Ecuador; Podocarpus NP] 1448 Rodriguesia 56 (87): 38 [State of Rio de Janeira] 1684 Boletim Instituto Botanica 19: 32 [Brazil; catalog] = Frullania brasiliensis mucronata Raddi 1822 (Lehm. & Lindenb. in Lehm.) Nees 1838 [nom inval; Art. 32.1(c)] Basionym: Distribution: - syn nov Nees 1838 Kamimura Boggan, Funk, Kelloff et al. 1961 1992 = Frullania mucronata 261 Naturg Europ Leberm 3: 210 [nomen in footnote], 239 [note #1 ad Fr. hispanica] 199 J Hatt Bot Lab 24: 21 [syn sub Fr. muscicola] 4771 Checklist of the plants of the Guianas (Lehm. & Lindenb. in Lehm.) Lehm. & Lindenb. 1845 Frullania brasiliensis Raddi 1822 Page 286 multiflora Typus: Steph. ex Verd. 1930 [pro syn] (? Peru) / (Brasilien) in hb HBG Basionym: Distribution: Verdoorn 1930 Feuerer 2008 = Frullania gaudichaudii multiflora 293 Ann Bryol Suppl 1: 92 [Frullania multiflora STEPH in sched. - syn sub Fr. gaudichaudii] 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm (Nees & Mont.) Nees & Mont. 1845 Steph. 1985 {Borneo} Basionym: Distribution: Stephani 1985 = Frullania gaudichaudii multiramosa 928 Stephani Icon Hepat Inedit [Thyopsiella] F-175d, microfiche 59 fig 4122 ["olim Frull multiflora nsp, Borneo, leg A Everett, No 37"] - 4123 ["Everett 37"] (Nees & Mont.) Nees & Mont. 1845 Steph. 1894 Typus: [1] Reunion Isl. Bourbon - [2] Madagascar [1] sd, Le Pervanche - [2] leg Besson hb BESCHERELLE et hb RENAULD 287, in hb G 13051 [holotypus] Basionym: Distribution: Madagascar spec nov - syn nov Stephani Stephani Bonner Vanden Berghen Stephani 1894 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania capensis 228 686 655 185 928 Hedwigia 33: 165 Spec Hepat 4: 624-625 Index Hepat 5: 374 Bull Jard Bot Nat Belg 46 (1/2): 42 Stephani Icon Hepat Inedit [Diastaloba] B-12, microfiche 48 fig 32393240 ["Bourbon leg Le Pervanche (Bescherelle) - Madagascar leg Dr. Besson Renauld 27 - Abbildg ist nach dem Ex vom Llapervanche (Bourbon)"] 250 Bryophyt Bibl 48: 64 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Gottsche 1845 Page 287 mundiana Lindenb. & Gottsche 1847 Typus: S Africa: Cape of Good Hope "Promont Bonae Spei" sd, Mund sn in hb HOOKER 14 in hb S (B24157, B24158) [typus] in hb GLM (hb LEHMANN) 06010 Basionym: Distribution: South Africa spec nov - syn nov Lindenberg & Gottsche in G., L. & N. Gottsche Stephani in Renauld & Cardot Schiffner 1847 2848 Syn Hepat: 772 (No 6 b) 1882 1891 1893 Sim Demaret & Vanden Berghen Arnell Bonner Grolle Wigginton 1926 1948 1963 1965 1995 2002 Myrdal Wigginton Wigginton 2002 2004 2004 254 Abh Naturwiss Ver Bremen 7 (3): 362-363 [Madagascar] 1453 Rev Bryol 18 (4): 56 [Madagascar] 572 Nova Acta K Leop-Carol Deutsch Akad Naturf Halle 60 (2): 226 [syn sub Fr. ecklonii] 252 Trans Roy Soc S Africa 15: 38 [syn sub Fr. ecklonii] [South Africa] 59 Bull Jard Bot Etat Bruxelles 19 (1): 77 [syn nov sub Fr. arecae] 822 Hepat S Africa: 136 [syn sub Fr. arecae] 655 Index Hepat 5: 374-375 250 Bryophyt Bibl 48: 63 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1276 Mosskollekter [Frullania types in hb S] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 224 [syn sub Fr arecae, West Africa] = Frullania obscura muscicola (Sw.) Mont. 1840 [auct. non Steph. 1894] Verd. in Hand.-Mazz. 1930 Basionym: Distribution: Verdoorn in Handel-Mazzetti Schuster & Hattori in Hattori - syn nov = Frullania gemmipara 1930 1978 550 482 Symb Sinicae 5: 38 J Hatt Bot Lab 44: 547-548 R.M.Schust. & S.Hatt. 1978 Frullania fuscovirens var. gemmipara (R.M.Schust. & S.Hatt.) S.Hatt. & P.J. Lin 1985 muscicola Typus: [auct. non Steph. 1894] Verd. in Hand.-Mazz. 1930 - , Handel-Mazzetti 11404 Basionym: Distribution: auct - syn nov Verdoorn in Handel-Mazzetti Hattori & Lin 1930 1985 550 505 = Frullania kagoshimensis ssp. hunanensis muscicola Symb Sinicae 5: 38 [Handel-Mazzetti 11404] J Hatt Bot Lab 59: 137 (S.Hatt.) S.Hatt. & P.J.Lin 1985 [auct. non Steph. 1894] Verd. in Hand.-Mazz. 1930 Basionym: Distribution: auct - syn nov Verdoorn in Handel-Mazzetti Hattori & Lin = Frullania sinensis 1930 1985 550 505 Symb Sinicae 5: 38 [specimen Handel-Mazzetti 989] J Hatt Bot Lab 59: 143 Steph. 1906 Page 288 muscicola Typus: [auct. non Steph. 1894] Verd. in Hand.-Mazz. 1930 v. Handel-Mazzetti 989 Basionym: Distribution: - syn nov Verdoorn in Handel-Mazzetti Hattori & Lin = Frullania aposinensis muscicola 1930 1985 550 505 Symb Sinicae 5: 38 J Hatt Bot Lab 59: 131 S.Hatt. & P.J.Lin 1985 [auct. non Steph. 1894] Verd. in Hand.-Mazz. 1930 Basionym: Distribution: - syn nov Verdoorn in Handel-Mazzetti Hattori & Lin = Frullania lushanensis muscicola 1930 1985 550 505 Symb Sinicae 5: 38 J Hatt Bot Lab 59: 139 S.Hatt. & P.J.Lin 1985 [auct. non Steph. 1894] SÈRGIO 1985 {Macaronesia} Basionym: Distribution: - syn nov Sergio 1985 Bisang, Schumacker, Sérgio & 1989 Grolle Sim-Sim, Sergio, Mues & Kraut 1995 = Frullania azorica muscicola 10 276 1 Portug Acta Biol (B) 14: 183 [Portugal] Giorn Bot Ital 122: [Asian & Macaronesian specimens as var nov: Fr cesatiana var muscicola ] Crypt Bryol Lichen 16 (2): 111-123 [belong to Fr. azorica] Sim-Sim, Sérgio, Mues & Kraut 1995 fo. C [auct. non Steph. 1894] Kamim. 1961 Basionym: Distribution: Kamimura = Frullania parvistipula 1961 199 J Hatt Bot Lab 24: 29 [Fr. parvistipula = Fr. muscicola fo. C] Steph. 1910 muscicola var. chungii Verd. in Hand.-Mazz. 1930 Typus: China. Fukien Prov, Buong Kang Yenping, 700-800m 29.6.1925, H.H. Chung B103, B108, F168 in hb FH (hb VERDOORN 10625 = B103) [lectotypus] & (hb VERDOORN 10651 = B168) [isotypus], [lectotypification by HATT 1974] in hb MICH MICH514767 > see fr1101.jpg Basionym: Distribution: var nov - syn nov Verdoorn in Handel-Mazzetti Bonner Hattori Hattori Piippo Fogel 1930 1965 1974 1982 1990 1997 550 655 545 495 521 629 Bapna & Kachroo 2000 712 = Frullania nepalensis Symb Sinicae 5: 38 Index Hepat 5: 375 J Hatt Bot Lab 38: 207 [syn nov sub Fr. nepalensis] [lectotypification] J Hatt Bot Lab 51: 250 J Hatt Bot Lab 68: 45 [annotated catalogue of China] http://141.211.110.91/livrqury.htm [16.11.1997] - Genus=Frullania, printed 5.1.1998 11:11:38: page 2 of 3 Hepaticology in India II: 176 [syn sub Fr. nepalensis] (Spreng.) Lehm. & Lindenb. 1845 Page 289 muscicola var. inaeana (Steph.) Kamim. by Parihar, Lal & Katiyar 1994 [orth var] Basionym: Frullania inuena Distribution: orth var Steph. 1910 Parihar, Lal & Katiyar = Frullania muscicola 1994 2196 Hepat Anth India, annotated checklist: 22 [Central India] - new record from India (Lal & Parihar 1979) Steph. 1894 muscicola var. inuena (Steph.) Kamim. 1961 Basionym: Frullania inuena Steph. 1910 Distribution: Japan, Korea (South), India (Central: Madhya Pradesh), Thailand comb nov - syn nov Kamimura Bonner Hong Kodama Tixier Choe & Yamada Choe Yuzawa & Hattori 1961 1965 1966 1972 1973 1974 1980 1981 Parihar, Lal & Katiyar 1994 Yamada & Choe 1997 Bapna & Kachroo Yuzawa 2000 2001 Yamada & Iwatsuki 2006 = Frullania muscicola 199 655 1107 1375 427 405 1171 494 J Hatt Bot Lab 24: 25-27, fig 7(16-35), map 2 Index Hepat 5: 375 Bryologist 69 (1): 412 [South Korea] Spec Publ Osaka Mus Nat Hist 4: 130 [Central Japan] Nat Hist Bull Siam Soc 24 (3/4): 456 [Thailand] Misc Bryol Lichen 6 (9): 161 [checklist Korea] Illust Fl & Fauna Korea 24: 734 [valid taxon; South Korea] J Hatt Bot Lab 50: 256 [seems to be identical with Fr. muscicola var. muscicola] 2196 Hepat Anth India, annotated checklist: 22 [Central India] as var. "inaeana" 717 J Hatt Bot Lab 81: 288 [Korean Peninsula] [syn sub Fr. muscicola var. muscicola] 712 Hepaticology in India II: 175-176 1660 Natural Environmental Sci Res 14: 26, fig 21(a-h) [Japan, keys to species; in Japanese] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] Steph. 1894 muscicola ssp. brittoniae R.M.Schust. & S.Hatt. in S.Hatt. 1981 Basionym: Distribution: comb nov Hattori Schuster = Frullania brittoniae myosota 1981 1992 491 603 J Hatt Bot Lab 49: 165 Hepat N America 5: 157- , fig 776(6), 797, 798(1-9) [Fr. brittoniaeis a valid species] A.Evans 1897 [auct. non (Hooker & Taylor) Mitt. 1855] Spruce 1884 Typus: Andes Quitenses Basionym: Distribution: Spruce Bonner = Frullania compsotera Frullania obcordata 1884 1965 243 655 Trans Proc Bot Soc Edinburgh 15: 57-58 Index Hepat 5: 375-376 [from New Zealand] Spruce 1884 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 290 myosota (Hook. f. & Taylor) Mitt. 1855 Basionym: Jungermannia myosota Distribution: - syn nov Mitten in Hooker 1855 Mitten in Hooker Spruce 1867 1884 Schiffner 1895 Schiffner in Engler & Prantl Winter & Schäfer-Verwimp 1905 2020 = Frullania obcordata nana Hook. f. & Taylor 1844 560 Antarctic voyage, II. Fl Novae-Zelandiae 2: 163 [as Frullania myosota syn sub Fr. ptychantha] 1154 Handbook New Zealand Flora: 537 [syn sub Fr. ptychantha] 243 Trans Proc Bot Soc Edinburgh 15: 57-58 [syn sub Fr. compsotera, not Fr. ptychantha] 1015 Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 134 809 Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 134 5742 Frahmia 19: 13 [syn sub Fr. obcordata] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Taylor 1846 Typus: N-Amerika. Lake Superior, "on Neckera pumila, HEDW." in hb GREVILLE, typus in hb S (B4159) Basionym: Distribution: spec nov - syn nov Taylor Taylor in G., L. & N. Evans Stephani Frye & Clark Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot = Frullania eboracensis natalensis 1846 1847 1897 1910 1947 2002 292 2848 237 686 204 782 London J Bot 5: 404 Syn Hepat: 773, No 11b Trans Connet Acad Arts Sci 10 (1): Spec Hepat 4: 431 [syn sub Fr. eboracensis] Univ Wash Publ Biol 6 (5): 771 [sub Fr eboracensis] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] Lehm. 1844 Sim 1926 Typus: South Africa. Natal, Wellington, Rosetta sd, Sim 8007 in hb PRE (Pretoria) [holotypus] Basionym: Distribution: Zimbabwe spec nov - syn nov Sim Arnell Arnell Bonner Vanden Berghen Best Wigginton 1926 1955 1963 1965 1976 1990 2002 Manyanga & Perold Wigginton Wigginton 2004 2004 2004 = Frullania sylvestris 252 1690 822 655 185 879 1012 Trans Roy Soc S Africa 15: 44-45, figs [South Africa] Bot Not 108: 309 [syn sub Fr. sylvestris] Hepat S Africa: 138 [as "nathalensis" syn sub Fr sylvestris] Index Hepat 5: 376 Bull Jard Bot Nat Belg 46 (1/2): 206 Kirkia 13 (2): 296 [Zimbabwe] Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1301 SABONET Report 21: 4 [Zimbabwe] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 226 [syn sub Fr depressa, West Africa] Sim 1926 [non silvestris Steph. 1908] Page 291 naumannii Steph. ex Verd. 1930 [pro syn] Typus: Indonesia, West Irian, Sorong 23.6.1875, Naumann 16 in hb G 14048 [= isolectotypus of Fr. regularis] Basionym: Distribution: Verdoorn Bonner Hattori Stephani 1930 1965 1982 1985 549 655 495 928 Hattori & Piippo 1986 557 = Frullania regularis Nova Guinea 14 (4): 541 [nom herb syn sub Fr. regularis] Index Hepat 5: 376 [nom herb?] J Hatt Bot Lab 51: 240 Stephani Icon Hepat Inedit [Homotropantha] D-10, microfiche 54 fig 3716 [" olim Frull Naumannii St nsp, Nova Guinea, Naumann 75, Warnstorf 16"] - 3717 Acta Bot Fennica 133: 41 Schiffn. 1890 Page 292 neesii Lindenb. 1845 Typus: Brazil. Minas Gerais, Villa Ricca sd, Martius in hb NEES (W) Basionym: Distribution: Mexico, Honduras, Guatemala, Costa Rica, Cuba, Jamaica, Dominican Rep, Guadeloupe, Dominica, Venezuela, Guyana, Surinam, French Guiana, Brazil spec nov - syn nov Lindenberg in G., L. & N. Duss Stephani Bonner Stotler Stotler Griffin III Lemos Michel 1845 1904 1911 1965 1969 1970 1979 1980 2850 3132 686 655 4761 194 988 719 Yano Stephani 1984 1985 501 928 Fulford Gradstein & Florschütz-de Gradstein & Hekking Fulford & Sharp Pôrto Schäfer-Verwimp Morales Z. Schuster 1987 1989 1989 1990 1990 1991 1991 1992 99 1237 517 105 77 523 307 Behar, Yano & Vallandro Boggan, Funk, Kelloff et al. Sillett. Gradstein & Griffin Ill Visnadi & Vital Yano Boggan, Funk, Kelloff, Hoff, Cremers & Feuillet Costa & Yano Leon, Pocs & Rico Boas-Bastos & Bastos Yano Yano & de Mello Oliviera-e-Silva & Yano Visnadi & Vital Molinario & Costa Oliveira-e-Silva, Milanez & Yano Adiego, Boldrin, Vieira, Monteiro, Barbosa Martins & Mello Costa & Silva 1992 1992 1995 1995 1995 1997 921 4771 2855 32 2095 1396 Syn Hepat: 450 Flore Cryptogamique des Antilles Françaises: 131 [Martinique] Spec Hepat 4: 530-531 Index Hepat 5: 377 Nova Hedwigia 18: 450-452, pl 16 [Dominican Rep] The genus Frullania, separata: 450-452, pl 16 [Dominican Rep] Acta Amazonica 9 (3): 34 [Brazil, Manaus] Diss, Uni Fed do Rio Grande do Sul, Porto Alegre: 62-65, pl 8, map 7, [Brasil, Rio Grande do Sul] J Hatt Bot Lab 56: 506 [checklist Brazil] Stephani Icon Hepat Inedit [Thyopsiella] F-118, microfiche 58 fig 4044 ["Brasilia, Beyrich"]- 4045 Mem NY Bot Gard 45: 272 [Dominica, Dominican Rep] Trop Bryol 1: 42 [Venezuela, Mt Roraima] J Hatt Bot Lab 66: 209 [Suriname, French Guiana] Mem NY Bot Gard 63: 45 [Mexico, Dominican Rep ??] Crypt Bryol Lichen 11 (2): 123 [Brazil] J Hatt Bot Lab 69: 153 [Brazil] Trop Bryol 4: 38 [Costa Rica] The Hepaticae and Anthocerotae of North America, East of the Hundredth Meridian. Vol. VI. – Field Museum of Natural History, Chicago, 937 pp. Bol Mus Biol Mello Leitao, new ser 1: 27-28, fig 2 [Brazil] Checklist of the plants of the Guianas The Bryologist 98: 251-260 [Catalogue of Costa Rica] Trop Bryol 10: 71 [Brazil: Setiba State Park] J Hatt Bot Lab 78: 142 [annotated checklist of Brazil] Checklist Guianas: 10 [Guyana, Surinam, French Guiana] 1998 1998 1998 1999 1999 2000 2001 2001 2002 2002 757 706 702 1263 919 989 1120 1086 1077 986 Hoehnea 25 (2): 103-103 [Brasil] Crypt Bryol Lichen 19 (1): 15 [Venezuela, Andes; new to Venezuela] Trop Bryol 15: 103 [Brazil, Bahia] Briophyta do estado de São Paulo. Lista provisória [Brazil] Iheringia, ser Bot 52: 78-80 [Brasil] Boletim Inst Botanica 13: 28-29 [Rio de Janeiro, Brazil] Acta bot bras 15 (2): 260 [Brazil] Rodriguesia 52 (81): 113 [Brazil, Bot Gard in Rio de Janeiro] Trop Bryol 22: 81 & 93 [Brazil] http://www.asee.org/international/INTERTECH2002/617.pdf [Brazil] 2003 Gradstein & Costa Holz 2003 2004 Porto, Germano & Borges Visnadi Camara, Oliveira & Santiago Yano Yano Schäfer-Verwimp & Pócs Bordin & Yano Schäfer-Verwimp Yano 2004 2004 2005 2008 2008 2009 2010 2010 2012 1440 Bol Mus Biol Mello Leitao (N ser) 16: 25 [Reserva Natural da Vale do Rio Doce, Espirito Santo, Brazil] 984 Mem New York Bot Gard 87: 87 [Brazil] [syn nov sub Fr kunzei] 1184 [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm 1446 Brejos de Altitude em Pernambuco e Paraiba: 83 [Brazil] 1158 Acta bot bras 18(1): 93 [Sao Paulo, Brazil] 1359 Trop Bryology 26: 137 [Brazil; Distrito Federal Brasilia] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr kunzei; Brazil; catalog] 1684 Boletim Instituto Botanica 19: 32 [Brazil; catalog] 1985 Acta Bot Hungarica 51(3-4): 385 [Dominican Republic, Hispaniola] 2266 Pesquisas, sér Bot 61: 50 [Brazil] 2460 Crypt Bryol 31(4): 347 [syn sub Frull kunzei, Dominica, West Indies] 3678 Pesquisas, série Botânica 63:55-140 [Catálogo ... do Estado do Espírito Santo, Brasil] = Frullania kunzei (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 293 neocaledonica J.J.Engel 1999 Basionym: Frullania caledonica Distribution: New Caledonia (R.M.Schust.) R.M.Schust. 1992 [hom illeg] [non Gottsche ex Steph. 1894] Schuster Engel in Engel & Merrill Konrat, Braggins, Asakawa & Toyota Larrain et all nom nov - syn nov = Frullania chevalieri nepalensis 1970 1999 2006 761 J Hatt Bot Lab 33: 291-292, fig 5(1-5) [as Neohattoria caledonica] 756 Novon 9: 344 [Frullania neocaledonica ] 1500 Bryologist 109(2): 154 [reassessment of Schusterella] 2018 5012 Bry. Div. Evo. 40 (2): 110 [syn sub Frull chevalieri, with key] (R.M.Schust.) R.M.Schust. 1992 [auct. non Steph.] Verd. 1930 {Japan} Typus: Japan. Kiyoto, Miyadzu Faurie-1000 Basionym: Distribution: Verdoorn = Frullania japonica 1930 293 Ann Bryol Suppl 1: 52 [Faurie-1000, Kiyoto, Miyadzu] Sande Lac. 1864 Frullania jackii var. japonica (Sande Lac.) S.Hatt. 1957 Frullania jackii ssp. japonica Frullania davurica nepalensis (Sande Lac.) S.Hatt. 1959 Hampe ex Gottsche, Lindenb. & Nees 1845 fo. rotundata Verd. in Hand.-Mazz. 1930 Typus: China. Yunnan, Schilunga near Yunnanfu, 2100m 20.2.1914, Handel-Mazzetti 208 Basionym: Distribution: fo nov - syn nov Verdoorn in Handel-Mazzetti Bonner Hattori & Lin Piippo Bapna & Kachroo = Frullania giraldiana 1930 1965 1985 1990 2000 550 655 505 521 712 Symb Sinicae 5: 41 [fo nov] Index Hepat 5: 377 J Hatt Bot Lab 59: 159 [syn sub Fr. giraldiana] J Hatt Bot Lab 68: 45 [annotated catalogue of China] Hepaticology in India II: 171 C.Massal. 1897 nepalensis var. grevilleana (Taylor) S.Hatt. in Mizutani 1979 Basionym: Frullania grevilleana Taylor 1845 Distribution: Nepal, Sikkim, Bhutan, India (North: West Bengal (Darjeeling), NW Himal) comb nov - syn nov Long Hattori in Mizutani Hattori Parihar, Lal & Katiyar 1979 1979 1982 1994 Bapna & Kachroo Kattel 2000 2002 = Frullania nepalensis 50 487 495 2196 Lindbergia 5: 60 [Bhutan] J Hatt Bot Lab 46: 322-323 [Easter Nepal] J Hatt Bot Lab 51: 250 [syn nov sub Fr. nepalensis] Hepat Anth India, annotated checklist: 22 [Nepal] - new new combination (Mizutani 1979) 712 Hepaticology in India II: 176 [syn sub Fr. nepalensis] 1032 Liverworts of Nepal, Katmandu: 17-18 [Nepal, 1100-3900 m] (Spreng.) Lehm. & Lindenb. 1845 Page 294 nepalensis var. nishiyamensis (Steph.) S.Hatt. in Nakai 1941 Basionym: Frullania nishiyamensis Steph. 1897 Distribution: China, Japan, Korea, Taiwan comb nov - syn nov Hattori in Nakai Hattori Hattori Hattori Kamimura Bonner Choe 1941 1943 1944 1951 1961 1965 1980 Hattori Wu in Li, Gao, Wu et al. 1982 1985 Kuo & Chiang Piippo Yamada & Choe Bapna & Kachroo Yuzawa Zhu & So Yamada & Iwatsuki 1988 1990 1997 2000 2001 2001 2006 = Frullania nepalensis nervosa 1705 925 943 528 199 655 1171 Iconogr Pl Asia Orient 4 (1): 331-333, pl 111(a-m) Bot Mag Tokyo 57: 360 & 363, fig 4(h-r) Bull Tokyo Sci Mus 11: 149 [Japan, key to Japanese species] J Hatt Bot Lab 5: 66-67 J Hatt Bot Lab 24: 37 Index Hepat 5: 377 Illust Fl & Fauna Korea 24: 735 [syn nov sub Fr nepalensis; South Korea] 495 J Hatt Bot Lab 51: 250 [syn nov sub Fr. nepalensis] 1169 Bryoflora Xizang (Tibet), Science Press, Beijing: 525 [sub Fr nishiyamensis] 948 Taiwania 33: 13 [Index of Taiwan] 521 J Hatt Bot Lab 68: 45 [annotated catalogue of China] 717 J Hatt Bot Lab 81: 288 [Korean Peninsula] 712 Hepaticology in India II: 176 [syn sub Fr. nepalensis] 1660 Natural Environmental Sci Res 14: 28 [Japan, syn sub Fr nepalensis] 808 Nova Hedwigia Beih 121: 79 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] (Spreng.) Lehm. & Lindenb. 1845 Mont. in Berthold & Webb 1844 [hom illeg] [non Mont. 1840] Basionym: Distribution: spec nov Montagne in Barker-Webb & Berthelot Ade & Koppe 1844 910 1942 206 Jovet-Ast Müller 1946 1951 Müller in Rabenhorst Augier & Noailles Grolle Grolle & Long 1958 1969 1976 2000 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 2002 = Frullania polysticta Hist Nat Iles Canariensis 3,2,4: 55 Hedwigia 81 (1/2): 34 [Fr. nervosa MONT. = Fr. polysticta LINDB.] [Azores, Madeira, Teneriffa] 886 Mem Soc Biogeogr 8: 366 [Cape Verde Isl] 641 Rev Bryol Lichen 20: 124-135 [Portugal; Fr. nervosa has priority against Fr. polysticta] 1028 Krypt Flora (3ed) 6, 2: 1238-1240, fig 482(f-g) 829 Ann Fac Sci Univ Fed Cameroun 2: 33, 36 [Canary Isl: Tenerife] 183 Feddes Repert 87 (3/4): 194 760 J Bryol 22: 111 [checklist of Europe and Macaronesia] & 123 [syn sub Fr. tamarisci ??] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] Lindenb. 1845 Page 295 nervosa Mont. 1840 [non Mont. in Berthelot & Webb 1844] Typus: Canary Isl. sd, Berthelott & Webb in hb W (Lindenberg Hep 7359), in hb L, S-PA [isotypus], isotypus in hb S (B24165) Basionym: Distribution: Canary Isl [Spain] spec nov - syn nov Mont. in Barker-Webb & Berhold Montagne in G., L. & N. Montagne Stephani Verdoorn 1840 1845 1856 1911 1928 Verdoorn Schiffner Mädgefrau Müller in Rabenhorst Arnell Arnell Bonner Grolle 1932 1936 1943 1958 1961 1961 1965 1970 Vanden Berghen Stephani 1976 1985 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Wigginton 2002 Wigginton 2004 = Frullania tamarisci nervosa 2002 2637 2850 1912 686 2071 Hist. Nat. Iles Canaries 4: 55 Syn Hepat: 439 [No 61] Sylloge generum specierumque cryptogamaram: 84 Spec Hepat 4: 564 Rec Trav Bot Neerl 25A: 452-460, fig 1 [=Recueil des travaux botaniques néerlandais] 554 Nederl Kruidk Arch Jrg 1932 (2): 496 [Azoren] 277 Österr Bot Zeitschr 85 (1): 19-21 [nervosa syn sub Fr. ploysticta] 2799 Flora NF 37: 135 [Teneriffa] 1028 Krypt Flora (3ed) 6, 2: 1238-1240 54 Bot Not 114 (2): 177 [Cape Verde Isl] 180 Svensk Bot Tidskr 55 (2): 392 [Canary Isl] 655 Index Hepat 5: 378 615 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 312 [syn sub Fr. tamarisci] 185 Bull Jard Bot Nat Belg 46 (1/2): 127 928 Stephani Icon Hepat Inedit [Thyopsiella] F-197, microfiche 60 fig 4156 [Canar Inseln, Bolle legit, Bescherelle misit"] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] (L.) Dumort. 1835 [auct. non MONT.] Mueller 1958 [pp] Typus: Canary island. La Palma, Cumbre nueva, 1000-1400m 17.04.1901, J. Bornmüller in hb GZU Basionym: Distribution: Grolle Suanjak = Frullania polysticta nervosa 1976 2001 183 979 Feddes Repert 87 (3/4): 194 [types] http://www.kfunigraz.ac.at/botanik/gzu-hepaticae-typen2.htm [in hb GUZ] Lindenb. 1845 fo. minor Schiffn. 1936 [Hep europ exs 1000b] Typus: Canary island. La Palma, Cumbre nueva, 1000-1400m 17.04.1901, J. Bornmüller in hb MA MA-Hepat 1721 > see fr2329.pdf Basionym: Distribution: Pando = Frullania polysticta 2000 1273 Herbario de Criptogamia, Madrid (MA) http://161.111.170.202/herb/asp/dos.asp?conque=MAHepat&fnhe=1721&fadic=1 Lindenb. 1845 Page 296 nervosa var. hampei Schiffn. 1936 Basionym: Distribution: Schiffner 1936 277 Österr Bot Zeitschr 85 (1): 22 = Frullania tamarisci var. nervosa (Mont.) S.W.Arnell 1961 Frullania tamarisci (L.) Dumort. 1835 nervosa var. microcellularis MUELLER in Rabenhorst 1957 Typus: Portugal. Monserrate 1952, Rauh Basionym: Distribution: Portugal var nov - syn nov Müller in Rabenhorst Sim-Sim 1958 1989 1028 Krypt Flora (3ed) 6, 2: 1239-1240 6 Portug Acta Biol (B) 15: 364 [Portugal] = Frullania tamarisci var. nervosa (Mont.) S.W.Arnell 1961 Frullania tamarisci (L.) Dumort. 1835 nervosa var. polysticta Nees ex Lindenb. 1845 [pro syn] Basionym: Distribution: - pro syn Lindenberg in G., L. & N. Müller 1845 1951 Bonner 1965 = Frullania polysticta 2850 Syn Hepat: 440 641 Rev Bryol Lichen 20: 134-135 [specimen in hb NEES; syn sub Fr polysticta] 655 Index Hepat 5: 378 [nom herb] Lindenb. 1845 neurota var. pfleiderei S.Hatt. 1972 Typus: India orientalis Pfleiderer K-93 Basionym: Distribution: var nov - syn nov Hattori Hattori in Ohashi Yuzawa = Frullania neurota 1972 1975 1991 538 187 525 J Hatt Bot Lab 36: 133, fig. 18 Univ Mus Univ Tokyo Bull 8: 234 [syn sub Fr. neurota] J Hatt Bot Lab 70: 222 [sub Fr. neurota] Taylor 1846 Page 297 newtoni Steph. 1911 Typus: Fernando Po, 1000 m -.2.1894, F. Newton sn ex hb BROTHERUS; in hb G [holotypus] Basionym: Distribution: Fernando Po spec nov - syn nov Stephani Demaret & Vanden Berghen Bonner Stephani 1911 1950 1965 1985 Wigginton Wigginton Uribe 2004 2004 2008 = Frullania angulata nicholsonii 686 60 655 928 Spec Hepat 4: 592 Bull Jard Bot Etat Bruxelles 20: 5 [syn sub Fr. angulata] Index Hepat 5: 378 Stephani Icon Hepat Inedit [Meteriopsis] E- 6ab, microfiche 55 fig 3771 ["Africa occ., Fernando Po, F Newton 1894"] - 3772 ["Africa, Fernando Po (Newton legit)"] 1312 Trop Bryol Res Rep 5: 31 [perhaps syn of Fr angulata] 1172 Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Mitt. 1863 Verd. ex S.Hatt. 1981 [nom herb] [non Hodgson 1949] Basionym: Distribution: Hattori 1981 491 J Hatt Bot Lab 49: 149 = Frullania pseudoschensiana var. darjeelingensis S.Hatt. 1981 nicobarica Gottsche ex Verd. 1930 [pro syn] Typus: Nicobar Isll. sd, E. Berkeley 1299 in hb G [sub Fr. intermedia] Basionym: Distribution: Andaman & Nicobar Islands [India] Gottsche Verdoorn Bonner Singh in Nath & Astana = Frullania gaudichaudii nidulans 0 1930 1965 2001 Icon inedit 293 Ann Bryol Suppl 1: 92 655 Index Hepat 5: 379 1374 Perspectives Indian Bryology: 341[endemic to Andaman & Nicobar Islands] (Nees & Mont.) Nees & Mont. 1845 Steph. ex Bonner 1965 [nom herb] Typus: Brazil. Estado de Ceara, "in ramis nidulans" sd, Ule 660 in hb G in hb HBG Basionym: Distribution: Brazil Bonner Stephani 1965 1985 Feuerer 2008 655 928 Index Hepat 5: 379 Stephani Icon Hepat Inedit [Meteriopsis] E-45, microfiche 56 fig 3837 ["Brasilia, Ule leg 1910 (No 660)"] 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm = Page 298 nietneri Gottsche 1886 [nom nud] Typus: Sri Lanka: Poondeloya 4000-8000 ft 1868, John Nietner Drawings com. by Jack.:hb no 1577 G. Ms. c. per. - 4 Bl. Basionym: Distribution: Gottsche Gottsche 0 1886 = Frullania nietneri Steph. 1911 Frullania ceylanica nietneri Icon inedit 2390 Bot. Centralblatt 26: 204 Nees 1845 Steph. 1911 Typus: Sri Lanka. Poondeloya, 4000-8000ft sd, Nietner 1540 in hb G 16350 (ex hb JACK) Basionym: Distribution: spec nov - syn nov Gottsche Stephani Verdoorn Bonner Hattori Stephani 1886 1911 1930 1965 1973 1985 Uribe 2008 = Frullania ceylanica nigashiyamensis 2300 686 293 655 Bot Centeralblatt 26: 2004 Spec Hepat 4: 610 Ann Bryol Suppl 1: 95 Index Hepat 5: 379 vide #1513 928 Stephani Icon Hepat Inedit [Meteriopsis] E-56, microfiche 56 fig 3852 ["Ceylon (Nietner legit)"] - 3853 1513 Caldasia 30(1): 90, [belongs to subgen Fusiorielligerae] [Monography of subgenus Meteoriopsis] Nees 1845 fo. validam Steph. ex Pando 2000 [nom herb in hb MA-Hepat 3479] Typus: JPN. : Mt. Ryumon, Kokawa in Naga distr., Wakayama Prefecture , 720 m on esposed serpentine rocks T.Kodama in hb MA MA-Herb 3479 Basionym: Distribution: Pando 2000 1273 Herbario de Criptogamia, hb MA = Frullania nepalensis var. nishiyamensis (Steph.) S.Hatt. in Nakai 1941 Frullania nepalensis nigra (Spreng.) Lehm. & Lindenb. 1845 Steph. 1911 Typus: Brazil. Brasilia orientalis, prope Rio Janeiro sd, s. coll. in hb G (ex hb STR, hb STEPHANI 3) Basionym: Distribution: Brazil spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Yano Stephani = Frullania brasiliensis 1911 1965 1969 1970 1984 1985 686 655 4761 194 501 928 Spec Hepat 4: 502 Index Hepat 5: 379 Nova Hedwigia 18: 436 [syn sub Fr. brasiliensis] The genus Frullania, separata: 436 [syn sub Fr. brasiliensis] J Hatt Bot Lab 56: 537 [syn sub Fr. ecuadorensis] Stephani Icon Hepat Inedit [Thyopsiella] F- 76, microfiche 57 fig 3962 ["Rio (Her Massal.) (1908)"] - microfiche 58 fig 3963 Raddi 1822 Page 299 nigricans Steph. 1911 Typus: Bourbon / Reunion coll ignot ex hb STR, in hb G 10705 Basionym: Distribution: Reunion Isl spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania apiculata nigrita 686 655 185 928 Spec Hepat 4: 476-477 Index Hepat 5: 379 Bull Jard Bot Nat Belg 46 (1/2): 102 Stephani Icon Hepat Inedit [Thyopsiella] F- 11, microfiche 56 fig 3885 ["Bourbon, Herb Strassburg"] 250 Bryophyt Bibl 48: 63 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apiculata, West Africa] (Reinw., Blume & Nees) Nees 1845 Hampe ex Taylor ex Gottsche, Lindenb. & Nees 1845 [pro syn] Basionym: Distribution: Taylor in G., L. & N. Bonner = Frullania fragilifolia nilgirienses 1845 1965 2850 Syn Hepat: 437 [syn sub Fr fragilifolia] 655 Index Hepat 5: 380 (Taylor) Gottsche, Lindenb. & Nees 1845 [non Austin 1879] Steph. ex Steph. 1985 [pro syn] Basionym: Distribution: nom herb Stephani = Frullania acutiloba 1985 928 Stephani Icon Hepat Inedit [Galeiloba] C-100ab, microfiche 51 fig 3482 ["olim Frull nilgirienses St, Neelgherries, Nilgherry Mtes" = Fr acutiloba] Mitt. 1861 [non Gerola 1947] Frullania monocera var. acutiloba (Mitt.) Hentschel & von Konrat 2015 Page 300 nishiyamensis Steph. 1897 Typus: Japan. Sikoku. Kochi Pref., Prov. Tosa, Sakawa, Nishiyama or/? Prov Tosa, Ogawa-mura, Nishiyama -.4.1896, T. Inoue 56 in hb G [holotypus], in hb TNS [isotypus] Basionym: Distribution: China (Huan, Kwangsi, Anhui), Japan, Taiwan, Vietnam spec nov - comb nov Stephani Inoue Iishiba Sawada 1897 1898 1909 1909 Stephani Iishiba Reimers Verdoorn Horikawa Horikawa Hattori in Nakai 1910 1912 1931 1932 1934 1935 1941 213 945 890 930 Bull Herb Boissier 5: 79 & 90 [Japan] Bot Mag Tokyo 12: 74 [Japan] Bot Mag Tokyo 23: 363 [Japan] Bot Mag Tokyo 23 (no 268): 178-183, fig 8, 8' [key to Japanese species, in Japanese] 686 Spec Hepat 4: 401-402 888 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] 229 Hedwigia 71 (1/2): 33 [China] 554 Nederl Kruidk Arch Jrg 1932 (2): 489 [China] 34 J Sci Hiroshima Univ, ser B, div 2, 2: 242-243 [Taiwan] 1282 Bot Mag Tokyo 49: 53 [Korea] 1705 Iconogr Pl Asia Orient 4 (1): 331-333, pl 111(a-m) [as Fr nepalensis var nishiyamensis] Hattori Hattori Chen Hattori & Noguchi Kamimura Inoue Bonner Chen & Wu Hattori in Hara 1944 1952 1958 1960 1961 1961 1965 1965 1966 943 529 1530 965 199 964 655 Pocs, Tixier & Jovet-Ast Kodama 1967 1972 1710 1375 Choe & Yamada Lai & Wang-Yang Choe Stephani 1974 1976 1980 1985 405 33 1171 928 Liu Wu in Li, Gao, Wu et al. Inoue 1985 1985 1987 2801 1169 1155 Kuo & Chiang Piippo Parihar, Lal & Katiyar 1988 1990 1994 948 521 2196 Yamada & Choe Zhu, So & Ye Konrat & Braggins Bapna & Kachroo N.N. [National Science Museum Tokyo] Zhu & So Yuzawa Cai Yamada & Iwatsuki 1997 1998 1999 2000 2000 717 698 725 712 995 Iconographia plantarum Asiae orientalis 4 (3): 332 [comb nov as Fr. nepalensis var nishiyanensis] Bull Tokyo Sci Mus 11: 149 [Japan, key to Japanese species] J Hatt Bot Lab 8: 30 Acta Phytotaxonomica Sinica 7 (4): 275 Index spec Typ herb Japon, Bryophyta: 17 J Hatt Bot Lab 24: 37, fig 20(14-22), map 6 J Jap Bot 36(5): 187 [Formosa] Index Hepat 5: 380 Observ Fl Hwangshan: 11 Flora of Eastern Himalaya: 527 [close related, or conspecific with Fr nepalensis] Bot Kozlem 54 (1): 30 [North Vietnam] Spec Publ Osaka Mus Nat Hist 4: 131-132, fig 64(a, c-f) [Central Japan] Misc Bryol Lichen 6 (9): 161 [checklist Korea] Taiwania 21 (2): 168 [Index to Taiwan] Illust Fl & Fauna Korea 24: 734 [sub Fr. nepalensis; South Korea] Stephani Icon Hepat Inedit [Galeiloba] C-245, microfiche 53 fig 3665 ["Japan"] Invest Stud Nat 5: <133-155> [aus 698] Bryoflora Xizang (Tibet), Science Press, Beijing: 525, fig 228(17-20) Index type specimens in Nat Sci Mus, Tokyo: 13 [Prov Tosa, Ogawamura, Nishiyama] Taiwania 33: 13 [Index of Taiwan] J Hatt Bot Lab 68: 45 [annotated catalogue of China] Hepat Anth India, annotated checklist: 22 [Eastern Himalaya] - new record from India (Hattori 1966) J Hatt Bot Lab 81: 288 [Korean Peninsula] J Hatt Bot Lab 84: 167 [syn sub Fr. nepalensis] Trop Bryol 17: 106 [epiphyllous] Hepaticology in India II: 176 [syn sub Fr. nepalensis] http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 2001 2001 2002 2006 808 1660 1192 1470 Nova Hedwigia Beih 121: 79 [syn sub Fr. nepalensis] Natural Environmental Sci Res 14: 28 [Japan, syn sub Fr nepalensis] Chenia 7: 151 [China, Anhui Prov.] J Hatt Bot Lab 99: 31 [Catalog of Japan] = Frullania nepalensis 841 (Spreng.) Lehm. & Lindenb. 1845 Page 301 nishiyamensis var. onoi S.Hatt. 1952 [nom nud] Basionym: Distribution: Hattori Hattori Yamada & Iwatsuki 1952 1952 2006 = Frullania taradakensis nobilis 1508 J Jap Bot 27(10): <315-318> 1508 J Jap Bot 27(10): 317 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] Steph. 1910 fo. cochleata (Steph.) S.Hatt. 1978 Basionym: Frullania cochleata Distribution: comb nov var nov Steph. 1911 Hattori Hattori Grolle & Piippo Hattori & Streimann = Frullania nobilis 1978 1982 1984 1985 482 495 143 504 J Hatt Bot Lab 44: 535-536 [key to Fr. ornithocephala & Fr. nobilis] J Hatt Bot Lab 51: 251-252 {syn sub Fr. nobilis] Acta Bot Fennica 125: 21 [Cat. West Melanesia] J Hatt Bot Lab 59: 111 [comb nov] [New Guinea] Steph. 1894 nobilis var. intermedia S.Hatt. 1973 Typus: Papua New Guinea. Morobe, Wau Distr 1967, Schuster 67-5879 in hb NICH [holotypus] in hb MASS & JE [isotypus] Basionym: Distribution: Papua New Guinea var nov comb nov Hattori Hattori Hattori Hattori 1973 1973 1974 1975 Hattori Grolle & Piippo Hattori & Streimann Hattori, Piippo 1982 1984 1985 1986 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania pullei 541 542 546 547 J Hatt Bot Lab 37: 116-120, fig 64-66 J Hatt Bot Lab 37: 150 [New Guinea] J Hatt Bot Lab 38: 245 J Hatt Bot Lab 39: 297-298 [comb nov as Fr. ornithocephala var. intermedia] 495 J Hatt Bot Lab 51: 253 143 Acta Bot Fennica 125: 21 [Cat. West Melanesia] 504 J Hatt Bot Lab 59: 116, footnote [New Guinea] 557 Acta Bot Fennica 133: 53 [syn nov for F. nobilis Steph. var. intermedia Hatt.] 2112 Type sepecimens Hatt Bot Lab (NICH) 31: Holotype. New Guinea, Wau Distr. 5000-5400 ft. 25 May 1967. R. M. Schuster 67-5879 (NICH 314853). Verd. 1930 Frullania pauciramea Steph. 1911 nobilis ssp. pauciramea (Steph.) S.Hatt. 1978 Basionym: Frullania pauciramea Distribution: Steph. 1911 comb nov Hattori Hattori Grolle & Piippo = Frullania pauciramea 1978 1982 1984 482 495 143 J Hatt Bot Lab 44: 536 [comb nov] J Hatt Bot Lab 51: 253 [pauciramea is an own species] Acta Bot Fennica 125: 21 [Cat. West Melanesia] Steph. 1911 Page 302 nodulosa {Japan} [auct. non (Reinwardt, Blume & Nees) Nees in G., L., & N.1846] Kamim. 1943 Basionym: Distribution: - syn nov Kitagawa Yamada & Iwatsuki = Frullania brotheri nodulosa 1974 2006 2382 Proc Bryol Soc Japan 1(7): 75 [syn nov sub Fr brotheri] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] Steph. 1894 fo. brachyantha Herzog 1953 [nom inval] Typus: Honduras Basionym: Distribution: Honduras Herzog 1953 640 Rev Bryol Lichen 20: 130 [Honduras] = nodulosa fo. dapitana (Steph.) Verd. 1928 Basionym: Frullania dapitana Distribution: fo nov - syn nov Herzog 1927 174 Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Bonner Hattori Grolle & Piippo 1928 1929 1930 1930 1931 1932 1965 1980 1984 244 245 293 549 231 553 655 490 143 = Frullania nodulosa nodulosa Steph. 1894 Mit Inst Bot Hamburg 7(1): 196 [West-Borneo; f transiens in f. dapitanam] Ann Crypt exot 1 (2): 220 Ann Bryol 2: 154 [Sumatra] Ann Bryol Suppl 1: 182 Nova Guinea 14 (4): 546 Ann Bryol 4: 140 [Hepat select crit, ser 2 no 100; Borneo] & 150 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 64 [Java] Index Hepat 5: 382 J Hatt Bot Lab 47: 209, 213 [syn sub Fr. nodulosa] Acta Bot Fennica 125: 22 [Cat. West Melanesia] (Reinw., Blume & Nees) Nees 1845 fo. irreflexa Verd. 1928 Basionym: Distribution: fo nov - syn nov Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Hattori Miller, Whittier & Whittier Grolle & Piippo = Frullania nodulosa 1928 1929 1930 1931 1931 1932 1932 1980 1983 1984 244 245 293 230 231 553 554 490 609 143 Ann Crypt exot 1 (2): 220 Ann Bryol 2: 154 [Singapore, Sumatra] Ann Bryol Suppl 1: 182 Ann Bryol 4: 123 [Hepat select crit, ser 1 no 44; Bangka] Ann Bryol 4: 140 [Hepat select crit, ser 2 no 99; Java] & 150 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 64 [Malay Peninsula, Molukken] Nederl Kruidk Arch Jrg 1932 (2): 500 [Sumatra] J Hatt Bot Lab 47: 209, 212 Prod Flora Hepat Polynesiae: 134 Acta Bot Fennica 125: 22 [Cat. West Melanesia] (Reinw., Blume & Nees) Nees 1845 Page 303 nodulosa var. nipponica S.Hatt. & Kamim. in Kamim. 1943 Typus: Japan. Sikoku, Kochi Pref, Prov. Tosa, Amatsubo-mura, 500m 7284 12.8.1942, Kamim., 7268 [type] - also no. 7270， 7276， 7281， in hb NICH 15120 (13.8.1942, Kamim. 7268) [holotypus] Vide Inoue 1987 in hb TNS ((M. Kamim., 12. VIII. 1942) [holotypus] Vide Hattori & Noguchi 1960 in hb EGR (Kamim. 11.1949 - Exsiccata: Hepat. Jap. 128) [topotypus] Vide Kis 2004 Basionym: Distribution: Japan var nov - syn nov Kamimura Kamimura Tatuno Hattori & Noguchi 1943 1952 1952 1960 2798 873 1697 965 Kamimura Bonner Kitagawa Hattori Inoue 1961 1965 1974 1980 1987 199 655 2382 490 1155 N.N. [National Science Museum 2000 Tokyo] Yuzawa 2000 Kis 2004 995 Yamada & Iwatsuki 2006 Mizutani, Hasegawa & Iwatsuki 2009 1470 2112 = Frullania brotheri 1659 1377 Nat Sci Mus Tokyo 14: 292-293, fig 1(A-M) Contrib Fl Hepat Shikokuensem, Kochi: 129 J Sci Hiroshima Univ ser B, div 2 (Bot), 6(2): 51-57 [sex chromosomes] Index spec Typ herb Japon, Bryophyta: 17 [Holotype in hb TNS: Sikoku. Prov. Tosa: Amatsubo-mura (M. Kamimura, 12. VIII. 1942)] J Hatt Bot Lab 24: 57 [syn sub Fr nodulosa] Index Hepat 5: 381 Proc Bryol Soc Japan 1 (7): 75-76 [syn sub Fr. brotheri] J Hatt Bot Lab 47: 213-215, fig 30 [Monography of sg Homotropantha] Index type specimens in Nat Sci Mus, Tokyo]: 14 [Holotype in hb TNS: Amatsubo-mura, Kochi Pref., Japan. Leg. M. KAMIMURA, no. 7268.] http://research.kahaku.go.jp/botany/koke/indexf.htm [Holotype in hb TNS] [ M. Kamimura 7268] Natural Environmental Sci Res 13: 2 [Japan, syn sub Fr brotheri] F Hist Nat Mus Matraensis 28: 15-16 [Topotype in hb EGR: ( Kamimura 11.1949 - Exsiccata: Hepat. Jap. 128. ] J Hatt Bot Lab 99: 31 [Catalog of Japan] Type sepecimens Hatt Bot Lab 31: [Holotype in NICH: Japan, Shikoku, Kochi-ken, Prov. Tosa, Amatubo-mura. Saxicola. 13 Aug. 1942. M. Kamimura 7268 (NICH 15120). Steph. 1894 nodulosa var. plana Schiffn. 1890 Typus: Amboina 7.6.1875, Naumann Basionym: Distribution: Australia (QLD) var nov - syn nov Schiffner Schiffner 1889 1898 Verdoorn Verdoorn Bonner Hattori Grolle & Piippo Scott & Bradshaw Windolf Carthy McCarthy 1930 1932 1965 1980 1984 1986 1987 2003 2006 = Frullania nodulosa 581 831 Forschungsreise S.M.S. "Gazelle" 4 (4): 37 Conspect Hepat Archip Indici, Batavia: 334-335 [may be a different species, which is identical with Fr. cognata] 293 Ann Bryol Suppl 1: 182 554 Nederl Kruidk Arch Jrg 1932 (2): 500 [Borneo, New Guinea] 655 Index Hepat 5: 381 [Amboina, Moluccas, Australia] 490 J Hatt Bot Lab 47: 209 [syn sub Fr. nodulosa] 143 Acta Bot Fennica 125: 22 [Cat. West Melanesia] 56 Brunnonia 8 (1): 36 & 144 [Australia: QLD] 1025 Austrobaileya 2(4): 390 [checklist Queensland] 1125 Flora Australia Suppl Ser 21: 45 [as valid var; Australia (QLD)] 1385 Checklist Australian Liverworts [Australia QLD] (Reinw., Blume & Nees) Nees 1845 nodulosa var. secundiflora (Mont.) Verd. 1930 Basionym: Frullania secundiflora Distribution: - syn nov Mont. 1843 [non Taylor 1845] Verdoorn Hattori Miller, Whittier & Whittier = Frullania nodulosa 1930 1980 1983 293 490 609 Ann Bryol Suppl 1: 183, fig 304 [em et comb nov] J Hatt Bot Lab 47: 209 [syn sub Fr. nodulosa] Prod Flora Hepat Polynesiae: 134 (Reinw., Blume & Nees) Nees 1845 Page 304 nova Steph. ex Steph. 1985 [nom herb] {New Zealand} Typus: New Zealand. in hb G Basionym: Distribution: Stephani 1985 928 Stephani Icon Hepat Inedit [Galeiloba] C-266, microfiche 54 fig 3689 ["Frullania nova St nsp, Frull deplanata Mitten, Frull falciloba Taylot, Neu Seeland, Colenso, Curnow 1"] = novae-zealandiae Colenso 1887 Typus: New Zealand. County of Hawke`s Bay, Glenross 1886, D.P. Balfour in hb WELT Basionym: Distribution: New Zealand spec nov - syn nov Colenso Stephani Stephani Bonner Hamlin Hamlin Hattori Stephani 1887 1892 1910 1965 1972 1973 1979 1985 Glenny Glenny & Malcolm 1998 2005 = Frullania hampeana (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 S.Hatt. 1972 Basionym: Neohattoria australis Distribution: New Zealand nom nov - syn nov R.M.Schust. 1970 Schuster Hattori Hattori & Mizutani Glenny Glenny & Malcolm = Frullania rostrata noviapiculata Trans Proc New Zealand Inst 19: 296-297 J Linn Soc Bot 29: 274 [syn sub Fr. hampeana] Spec Hepat 4: 412 [as valid species] Index Hepat 5: 382 Rec Dominion Mus 7(19): 262 [New Zealand] Rec Dominian Mus 8 (10): 143 [New Zealand] J Hatt Bot Lab 46: 134 [= Fr. hampeana] Stephani Icon Hepat Inedit [Galeiloba] C-267, microfiche 54 fig 3690 ["Frullania Novae Zelandiae Col., Neu Seeland, Colenso legit"] 1412 Tuhinga 10: 132 [sub Fr monocera] 1410 ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr monocera] Nees 1845 Frullania monocera novaustralis 634 210 686 655 659 656 486 928 1970 1972 1982 1998 2005 761 539 495 1412 1410 J Hatt Bot Lab 33: 285-288, fig 4(5-9) [as Neohattoria australis] J Hatt Bot Lab 36: 443 J Hatt Bot Lab 52: 447 Tuhinga 10: 132 [sub Fr rostrata] ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr rostrata] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 S.Hatt. ex Parihar, Lal & Katiyar 1994 [nom nud] Basionym: Distribution: nom nud Parihar, Lal & Katiyar 1994 2196 Hepat Anth India, annotated checklist: 22 [South India] = Page 305 novopommeriensis Miller ex S.Hatt. & Streimann 1985 [nom inval] Basionym: Frullania faurieana Distribution: nom nov Steph. 1910 [hom illeg] [non Steph. 1894] Miller Miller, Whittier & Whittier Grolle & Piippo Hattori & Streimann 1981 1983 1984 1985 439 609 143 504 Hattori & Piippo 1988 557 Phytologia 47 (4): 322 [nom nov, pro Fr. fauriana] Prod Flora Hepat Polynesiae: 134 Acta Bot Fennica 125: 22 [Cat. West Melanesia] J Hatt Bot Lab 59: 120-121 [nom inval; as Fr. fauriana is syn sub Fr. ericoides var planescens] Acta Bot Fennica 133: 44 = Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides nutans (Nees) Mont. 1839 Steph. 1901 [non Steph. 1910] Typus: Tanzania. Ukinga-Berge "Kinyikaberg, im Schluchtenwald an Baumstämmen", 2600m 26.5.1899, W. Goetze hb B 951K, in hb G 17312 & Basionym: Distribution: spec nov - syn nov? Stephani Stephani Bonner Vanden Berghen Stephani 1901 1910 1965 1976 1985 215 686 655 185 928 Wigginton 2002 1012 Wigginton Wigginton Wigginton 2004 2004 2004 1312 1312 1172 = Frullania caffraria nutans Bot Jahrb 30: 260 Spec Hepat 4: 370 [nom superfl] Index Hepat 5: 383 Bull Jard Bot Nat Belg 46 (1/2): 185 [syn with doubt] Stephani Icon Hepat Inedit [Galeiloba] C- 26, microfiche 50 fig 3378 [from type; "Africa orient, Ukingaberge, Goetze 951 k"] Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: xx [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 229, fig 150( R) [perhaps syn sub Fr caffraria, West Africa] Steph. 1894 Steph. 1910 [nom superfl] [non Steph. 1901] Basionym: Distribution: Stephani = Frullania nutans 1910 Spec Hepat 4: 370 [spec nov = nom superfl] Steph. 1901 [non Steph. 1910] Frullania caffraria nymanii 686 Steph. 1894 Steph. 1911 Typus: Indonesia. Java, Salak sd, Nyman sn in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Bonner Stephani 1911 1929 1965 1985 686 245 655 928 Spec Hepat 4: 464-465 ["nymannii"] Ann Bryol 2: 134 Index Hepat 5: 383 [Java] Stephani Icon Hepat Inedit [Galeiloba] C-164, microfiche 52 fig 3562 [from type; "Java, Salak, leg Nyman"] = Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides (Nees) Mont. 1839 Page 306 oakesiana [auct. non Austin 1870] Kitagawa 1963 Typus: Sachalin ex KYO in JE Basionym: Distribution: Sakhalin Kitagawa Grolle Furuki & Mizutani = Frullania takayuensis 1963 1970 1994 626 Acta Phytotax Geobot 19: 150 [Sakhalin] 615 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 318 1331 Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] Steph. 1910 Frullania oakesiana ssp. takayuensis obconica (Steph.) R.M.Schust. 1992 Steph. 1910 Typus: Madagascar: Imerina sd, P. Camboué 115 in hb G, Basionym: Distribution: spec nov - syn nov Stephani Arnell Bonner Vanden Berghen Stephani 1910 1963 1965 1976 1985 Yuzawa Grolle Wigginton 1991 1995 2002 Wigginton Wigginton 2004 2004 = Frullania trinervis 686 822 655 185 928 Spec Hepat 4: 380-381 Hepat S Africa: 141 [syn nov sub Fr. trinervis] Index Hepat 5: 383-384 Bull Jard Bot Nat Belg 46 (1/2): 199 [syn sub Fr. trinervis] Stephani Icon Hepat Inedit [Galeiloba] C- 27, microfiche 50 fig 3379 ["Madagascar, leg Rotereau, Herb Dismier"] 525 J Hatt Bot Lab 70: 230 250 Bryophyt Bibl 48: 69 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 227 [syn sub Fr trinervis, West Africa] (Lehm.) Drège 1843 obcordata var. armata Lindenb. & Gottsche 1851 Typus: Suriname. "ad Poelebantje apud Paramaribo lecta" sd, Kegel sn Basionym: Distribution: Surinam Lindenberg & Gottsche Bonner Boggan, Funk, Kelloff, Hoff, Cremers & Feuillet -syn nov = Frullania obcordata oblicua 1851 1965 1997 282 Linnaea 24 (5): 638 [Suriname] 655 Index Hepat 5: 384 1396 Checklist Guianas: 10 [specimens of Guianas identified as F. caulisequa] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. by Uribe 2008 [orth var] Basionym: Distribution: Uribe = Frullania obliqua 2008 1513 Caldasia 30(1): 90, [orth var] Steph. 1911 [non Hampe ex Nees in G., L. & N. 1845] Frullania orientalis Sande Lac. 1855 Page 307 obliqua Steph. 1911 [non Hampe ex Nees in G., L. & N. 1845] Typus: Neu Guinea. PNG. Central, Moroka, 1300m -.7-8.1893, L. Loria sn in hb G (hb LEVIER 1280) [holotypus] Basionym: Distribution: New Guinea spec nov - syn nov Stephani Verdoorn Bonner Miller, Whittier & Whittier Grolle & Piippo Stephani 1911 1930 1965 1983 1984 1985 Uribe 2008 Uribe-M. 2011 = Frullania orientalis obliqua 686 549 655 609 143 928 Spec Hepat 4: 616-617 Nova Guinea 14 (4): 541 [syn sub Fr. orientalis] Index Hepat 5: 384 Prod Flora Hepat Polynesiae: 134 Acta Bot Fennica 125: 22 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Meteriopsis] E-58, microfiche 56 fig 3855 ["N Guinea, Moroka (Moresby) leg Loria 1893"] 1513 Caldasia 30(1): 90, [belongs to subgen Frullania] [Monography of subgenus Meteoriopsis] 2767 Caldasia 33(1): 71, fig 1(g-l) [type study, syn sub Fr orientalis] Sande Lac. 1855 Hampe ex Nees 1845 [pro syn] [non Steph. 1911] Typus: USA. Virginia, Ebenezar near Baltimore sd, Beyrich in hb HAMPE Basionym: Distribution: Nees in G., L. & N. Bonner 1845 1965 2850 Syn Hepat: 418 655 Index Hepat 5: 384 = Frullania aeolotis var. obliqua Nees 1845 Frullania riparia obscura Hampe 1838 Mont. 1840 {French Guiana: Leprieur No 265} Basionym: Distribution: French Guiana comb nov Montagne 1840 467 - syn nov Gradstein & Hekking 1989 517 = Frullania arecae (Spreng.) Gottsche 1863 [non (Spreng.) Spruce 1884] Frullania obscura obscura Ann Sci Nat Bot, ser 2, 14: 333 (French Guiana: Fr. obscura, Lepr coll n 265; auct, = Fr. areacae] J Hatt Bot Lab 66: 208 (Sw.) Mont. 1840 Steph. 1910 [hom illeg] [non (Sw.) Dumort. 1835] Typus: 1. Madagascar, Betsileo 2. Bourbon 3. La Reunion 1. sd, Montaut 374pp 2. A. Delessert 3. Rodriquex 194 1. in hb G 010706 2. in hb G 17313 3. in hb G 010708, /in hb UPS [isotypus] Basionym: Distribution: Madagascar, Reunion Isl spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1910 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania anderssonii 686 655 185 928 Spec Hepat 4: 375 Index Hepat 5: 385 Bull Jard Bot Nat Belg 46 (1/2): 172 Stephani Icon Hepat Inedit [Galeiloba] C- 28, microfiche 50 fig 3380 ["Bourbon, leg Delessert, auch Madagascar, leg Montaut"] 250 Bryophyt Bibl 48: 61 [nom illeg: Art 53.1] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Ångstr. 1873 Page 308 obscura var. costata Nees ex Lehm. & Lindenb. 1845 [pro syn] {Chile} Typus: Chile: ad Quillota sd, Bertero NEES in hb Berteroana apud Hochstetter Basionym: Distribution: var nov Lehmann & Lindenberg in G., L. 1845 & N. Bonner 1965 2850 Syn Hepat: 428 655 Index Hepat 5: 385 [nom herb; sub Fr trinervis var berteroana] = Frullania trinervis var. berteroana Lehm. & Lindenb. 1845 Frullania trinervis obtusifolia (Lehm.) Drège 1843 Pearson 1923 [orth var] Basionym: Distribution: Pearson = Frullania obtusiloba 1923 Univ California Publ Bot 10 (5): 379-380, pl 107(1-17) & 108(1-9) Pearson 1923 Frullania solanderiana obtusifolia 797 Colenso 1889 Steph. ex Steph. & Watts 1914 [nom nud] {Lord Howe Isl} Typus: Australia. [NSW] Lord Howe Isl, 1911, Watts Basionym: Distribution: Lord Howe Isl Stephani & Watts Miller, Whittier & Whittier Hattori 1914 1983 1987 220 609 400 J Proc Roy Soc New S Wales 48: 108 [Lord Howe Isl] Prod Flora Hepat Polynesiae: 134 [= nom nud] Bryologist 90 (4): 370 [= nom nud] = obtusiloba Pearson 1923 Typus: New Zealand. Island Bay 5.6.1904, Setchell 43 in hb UC 213700 in hb UC [typus, lost] lost on loan to ? late Frans Verdoorn 6.viii.1929 at Utrecht; New Zealand Basionym: Distribution: spec nov - syn nov Pearson Bonner Hamlin Hattori N.N. = Frullania solanderiana 1923 1965 1972 1983 2003 797 655 659 499 978 Univ California Publ Bot 10 (5): 379-380, pl 107(1-17) & 108(1-9) Index Hepat 5: 385 Rec Dominion Mus 7(19): 262 [New Zealand] J Hatt Bot Lab 54: 170 http://ucjeps.berkeley.edu/bryolab/moss_types.html [type in hb UC; lost] Colenso 1889 Page 309 ocanniensis Steph. 1924 Typus: Colombia.Antioquía, páramo de Ocaña, Pehlke s.n., “misit Irmscher nr. 12” 1912, Irmscher 12 [Pehlke legit] in hb G-00069231 Basionym: Distribution: Colombia spec nov - syn nov Stephani Bonner Gradstein & Hekking Stephani 1924 1965 1979 1985 Uribe & Gradstein Gradstein 1998 2021 = Frullania ericoides occanniencis 687 655 483 928 Spec Hepat 6: 542-543 Index Hepat 5: 385-386 J Hatt Bot Lab 45: 113 [Cat Colombia] [as Fr. occanniencis] Stephani Icon Hepat Inedit [Galeiloba] C- 70, microfiche 51 fig 3432 ["Columbia Paramo de Ocana, leg Pehlke, Herb Berlin"] 707 Bryophyt Bibl 53: 37 [Catalogue Colombia] [as Fr. occanniencis] 5642 Mem NY Bot Gard 121: 366 [syn sub Fr ericoides] (Nees) Mont. 1839 Steph. by Uribe & Gradstein 1998 [orth var] Basionym: Distribution: Uribe & Gradstein Bernal, Gradstein & Celis = Frullania ocanniensis Frullania ericoides occulta 1998 2016 707 Bryophyt Bibl 53: 37 [Catalogue Colombia] 4290 Catálogo de plantas y líquenes de Colombia. Vol. I. [dubious species] Steph. 1924 (Nees) Mont. 1839 Milde ex Bonner 1965 [nom herb] Typus: Italy. Meran sd, Milde 60 in hb G Basionym: Distribution: Italy Bonner 1965 655 Index Hepat 5: 386 = occulta Milde 1864 [pro syn, Gratsch bei Meran] Basionym: Distribution: Milde = Frullania aeolotis 1864 Botanische Zeitung (Berlin) 22, Beilage 3: 16 [auct. non MONT & Nees] Milde 1864 [Gratsch bei Meran] Frullania cesatiana De Not. 1865 Frullania riparia Hampe 1838 Page 310 oceanica Mitt. in Seemann 1871 Typus: ? 1 Hawaii 2 Sandwich Isl 3 Samoa ? 1 Beechey; Macrae 2 Hillebrand 3 Powell in hb NY Basionym: Distribution: Samoa, Hawaii spec nov Mitten in Seemann Stephani in Reinecke Stephani Evans Schiffner Stephani Stephani Verdoorn Verdoorn Miller Clark Miller Bonner Grolle & Schultze-Motel Miller, Whittier & Whittier Stephani So & Wang - syn nov = Frullania apiculata onoi 1871 1896 1897 1900 1910 1911 1911 1930 1930 1953 1955 1963 1965 1972 1983 1985 2006 637 1044 214 236 436 686 686 551 293 349 369 834 655 537 609 928 1466 Flora Vitiensis: 417 [Hawaii, Sandwich Isl, Samoa] Bot Jahrb 23: 314 [Samoa] Bull Herb Boissier 5: 842[:260] [Hawaii] Trans Connet Acad Arts Sci 10 (1): 400-402 [syn nov sub Fr. apiculata] Österr Bot Zeitschr 60 (11): 432 Spec Hepat 4: 642-643 Spec Hepat 4: 542 [sub Fr. apiculata] Nederl Kruidk Arch Jrg 1930 (2): Ann Bryol Suppl 1: 97 Bryologist 56: 45 [sub Fr apiculata] Bryologist 58: 71 Ark Bot, ser 2, 5: 518 [sub Fr. apiculata] Index Hepat 5: 386 J Hatt Bot Lab 36: 83 [sub Fr. apiculata] [Samoa] Prod Flora Hepat Polynesiae: 134 Stephani Icon Hepat Inedit Icon n. 3488, 4056-8 J H B L 100: 420 [sub Fr apiculata] (Reinw., Blume & Nees) Nees 1845 S.Hatt. ex S.Hatt. 1952 [nom herb] Basionym: Distribution: Hattori Yamada & Iwatsuki 1952 2006 = Frullania taradakensis ontakensis 529 J Hatt Bot Lab 8: 31 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] Steph. 1910 Steph. ex Sawada 1909 [nom nud] Basionym: Distribution: Sawada = Frullania ontakensis 1909 930 Bot Mag Tokyo 23 (no 268): 178-183, fig 4, 4' [key to Japanese species, in Japanese] Steph. 1910 Frullania schensiana C.Massal. 1897 Page 311 ontakensis Steph. 1910 Typus: Japan. Honshu, Nagano Pref, Prov Kiso, Mt. Ontake, 2000m -.9.1905, U. Faurie 1713?, 1715 in hb G in hb KYO in hb TNS (U. Faurie 1715) [isotypus] in hb HIRO (U. Faurie 1715 )[isotypus] Basionym: Distribution: spec nov - syn nov Sawada 1909 Stephani Iishiba Reimers Hattori Hattori Hattori & Noguchi Kamimura Bonner Hattori 1910 1912 1931 1951 1951 1960 1961 1965 1975 Choe Stephani 1980 1985 Hattori & Lin Inoue Piippo N.N. [National Science Museum Tokyo] Bapna & Kachroo NN 1985 1987 1990 2000 Yamada & Iwatsuki 2006 = Frullania schensiana ontakensis 2000 2003 930 Bot Mag Tokyo 23 (no 268): 178-183, fig 4, 4' [key to Japanese species, in Japanese] 686 Spec Hepat 4: 403-404 [Ontake, Nankai, Takeo] 888 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] 229 Hedwigia 71 (1/2): 34 [China] 860 J Hatt Bot Lab 5: 83, pl 2(34) [oilbodies] 528 J Hatt Bot Lab 5: 67 965 Index spec Typ herb Japon, Bryophyta: 18 199 J Hatt Bot Lab 24: 31 [syn sub Fr schensiana] 655 Index Hepat 5: 387 66 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4); 163 [not syn to schensiana, better a ssp. of schensiana] 1171 Illust Fl & Fauna Korea 24: 735 [sub Fr schensiana] 928 Stephani Icon Hepat Inedit [Galeiloba] C-246, microfiche 53 fig 3666 ["Japan, Ontako, Sept 1905, Faurie No 1713"] 505 J Hatt Bot Lab 59: 143 [sub Fr. schensiana] 1155 Index type specimens in Nat Sci Mus, Tokyo: 14 521 J Hatt Bot Lab 68: 45 [annotated catalogue of China] 995 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 712 Hepaticology in India II: 181 [sub Fr. schensiana] 1397 Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] C.Massal. 1897 fo. punctata S.Hatt. 1944 Typus: Japan. Shikoku, Jyo prov., Matsuyama 9.3.1896, Matsudaira in hb TNS [holotypus] Basionym: Distribution: Japan stat nov Hattori Hattori & Noguchi Kamimura Bonner Yuzawa 1944 1960 1961 1965 2001 Yamada & Iwatsuki 2006 = Frullania schensiana orientalis 1911 965 199 655 1660 J Jap Bot 20 (2): 156 /? 156, fig 35 Index spec Typ herb Japon, Bryophyta: 18 [holotype] J Hatt Bot Lab 24: 33 Index Hepat 5: 386-387 Natural Environmental Sci Res 14: 35 [Japan, basionym of sub Fr schensiana var punctata] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] C.Massal. 1897 [auct. non Sande Lac. 1855] S.Hatt. 1985 Typus: Fiji Basionym: Distribution: Hattori = Frullania capillaris 1985 73 Bull Natn Sci Mus Tokyo, ser B (Bot), 11 (1): 13 Steph. 1911 Page 312 ornithocephala [auct. non (R B N) Nees] S.Hatt. 1974 Basionym: Distribution: auct - syn nov Hattori Hattori Hattori 1974 1975 1982 = Frullania pauciramea 546 547 495 J Hatt Bot Lab 38: 245-250, fig 120 - 123 J Hatt Bot Lab 39: 297 J Hatt Bot Lab 51: 253 [specimens from New Guinea] Steph. 1911 ornithocephala var. cochleata (Steph.) S.Hatt. 1975 Basionym: Frullania cochleata Distribution: comb nov - comb nov Steph. 1911 Hattori Hattori Grolle & Piippo = Frullania nobilis 1975 1978 1984 547 482 143 J Hatt Bot Lab 39: 297-298, fig 148 J Hatt Bot Lab 44: 536 [as Fr. nobilis fo. cochleata] Acta Bot Fennica 125: 22 [Cat. West Melanesia] fo. cochleata (Steph.) S.Hatt. 1978 Frullania nobilis Steph. 1894 ornithocephala var. intermedia (S.Hatt.) S.Hatt. 1975 [nom inval] Basionym: Frullania nobilis var. intermedia S.Hatt. 1973 Distribution: comb nov Hattori Hattori Grolle & Piippo Hattori 1975 1982 1984 1988 547 495 143 516 J Hatt Bot Lab 39: 297-298 [non inval] J Hatt Bot Lab 51: 252-253 [syn sub Fr. nobilis var intermedia] Acta Bot Fennica 125: 22 [Cat. West Melanesia] J Hatt Bot Lab 65: 439-441 [PNG] = Frullania nobilis var. intermedia S.Hatt. 1973 Frullania pullei Verd. 1930 Frullania pauciramea Steph. 1911 ornithocephala var. major (Nees) Schiffn. 1898 Typus: Indonesia. Java Basionym: Jungermannia ornithocephala var. β major Nees 1830 Distribution: Indonesia (Java) - syn nov Schiffner Schiffner 1893 1898 572 831 Bonner 1965 655 = Frullania panamana Nova Acta K Leop-Carol Deutsch Akad Naturf Halle 60 (2): 226 Conspect Hepat Archip Indici, Batavia: 335-336 [syn sub Frullania ornithocephala] Index Hepat 5: 387-388 Steph. 1911 Frullania brasiliensis Raddi 1822 ornithocephala var. minor Nees 1845 Typus: Indonesia. Java Basionym: Distribution: Indonesia (Java) Nees in G., L. & N. Bonner = Frullania ornithocephala 1845 1965 2850 Syn Hepat: 425 655 Index Hepat 5: 388 (Reinw., Blume & Nees) Nees 1845 Page 313 osculantiara De Not. 1855 [orth. var.] Basionym: Distribution: Rodriquez et al. 2017 = Frullania osculatiana De Not. 1855 4629 Arnaldoa 24(1): 255 [Peru, La Libertad] osumiensis var. dentata Kamim. 1970 Typus: Japan. Fukushima Pref. Iwaki, Toono-machi, Gozaisho Pass Y. Yusawa 1003-a ? [holotypus], in hb TNS [isotypus] Basionym: Distribution: Japan var nov - syn nov Kamimura Hattori & Lin Inoue Furuki & Mizutani N.N. [National Science Museum Tokyo] Yuzawa 1970 1985 1987 1994 2000 1552 505 1155 1331 995 2001 Yamada & Iwatsuki 2006 1660 Natural Environmental Sci Res 14: 32, fig 27(a-m) [Japan, syn nov sub Fr osumiensis] 1470 J Hatt Bot Lab 99: 31 [Catalog of Japan] = Frullania osumiensis Bulletin of Kochi Gakuen Junior College 2: 21-22, fig 2(m-x) [Japan] J Hatt Bot Lab 59: 141-142 [syn sub Fr osumiensis] Index type specimens in Nat Sci Mus, Tokyo: 14 Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] (S.Hatt.) S.Hatt. 1956 osumiensis var. orbiculata Kamim. 1961 Typus: Japan. Ehima, Mt. Iwayasan -.8.1899, K. Okudaira 43 in hb NICH Basionym: Distribution: Korea (South) var nov - syn nov Kamimura Bonner Hong Choe Choe Hattori & Lin Yamada & Choe Yuzawa Hong Yamada & Iwatsuki = Frullania osumiensis 1961 1965 1966 1980 1983 1985 1997 2001 2003 2006 199 655 1107 1171 J Hatt Bot Lab 24: 39, fig 15(6-11) Index Hepat 5: 389 Bryologist 69 (1): 412 [South Korea] Illust Fl & Fauna Korea 24: 735 [South Korea] J Sci Educ 15: <147-156> [aus 717] 505 J Hatt Bot Lab 59: 141-142 [syn sub Fr osumiensis] 717 J Hatt Bot Lab 81: 288 [Korean Peninsula] 1660 Natural Environmental Sci Res 14: 32 [Japan, syn sub Fr osumiensis] 1416 Lindbergia 28: 145 [syn sub Fr osumiensis] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] (S.Hatt.) S.Hatt. 1956 Page 314 ovalifolia Kiaer & Pearson in Pearson 1891 Typus: Madagascar: Ankarata -.5.1877, Borgen sn Hb KIAER 107 in hb O Basionym: Distribution: Madagascar spec nov - syn nov Kiaer & Pearson in Pearson 1891 Stephani in Levier Stephani Bonner Vanden Berghen Stephani 1901 1910 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania anderssonii ovifolia 1921 Christiania Videnskabs-Selskabs Forhandlinger 1890 (2): 4-5, pl. 1(114) [Madagascar] 1456 Rev Bryol 28 (5): 94 [Madagascar] 686 Spec Hepat 4: 360 ["ovalifolia"], 376-377 ["ovifolia"] 655 Index Hepat 5: 389 185 Bull Jard Bot Nat Belg 46 (1/2): 172 [syn sub Fr. anderssonii] 928 Stephani Icon Hepat Inedit [Galeiloba] C- 30, microfiche 50 fig 3382 ["Madagascar, leg Borgen"] 250 Bryophyt Bibl 48: 61 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Ångstr. 1873 Kiaer & Pearson by Steph. 1910 [orth var] Basionym: Distribution: Stephani Beauverd in Stephani 1910 1924 Vanden Berghen Wigginton 1976 2002 Wigginton 2004 = Frullania ovalifolia Spec Hepat 4: 360 ["ovalifolia"], 376-377 ["ovifolia"] Spec Hepat 6: 570 [sphalm. = orth var.] [Frullania ovalifolia Kiaer.et Pearson » non P. ovifolia (sphalm.)] 185 Bull Jard Bot Nat Belg 46 (1/2): 172 [orth var] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Kiaer & Pearson in Pearson 1891 Frullania anderssonii ovistipula 686 909 Ångstr. 1873 Steph. ex Herzog 1916 Typus: Bolivia, Vallegrande, Cerro Pampalarga, 2300 m 1911, Herzog 4144 in hb G Basionym: Distribution: Bolivia spec nov - syn nov Stephani Stephani Bonner Stephani 1916 1924 1965 1985 222 687 655 928 Yuzawa, Mues & Hattori Yuzawa Yuzawa Gradstein, Meneses & Arbe 1987 1991 1991 2003 509 525 525 975 = Frullania beauverdii Biblioth Bot 87: 240, fig. 188 [Bolivia] Spec Hepat 6: 533-534 [as spec nov] Index Hepat 5: 389 Stephani Icon Hepat Inedit [Chonanthelia] A- 44, microfiche 46 fig 3132 [from type; "Bolivia Herzog 4144 (1913)"] J Hatt Bot Lab 63: 425-436 [morphology / systematics] J Hatt Bot Lab 70: 273-274, fig. 47 [redescription of the type specimen] J Hatt Bot Lab 70: 271 [syn sub Fr. beauverdii] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. beauverdii] Steph. 1916 Page 315 pabstiana Steph. 1911 Typus: Brazil. Santa Catherina sd, Pabst in hb G Basionym: Distribution: spec nov - sny nov Stephani Herzog Schiffner & Arnell Bonner Yano Stephani 1911 1925 1964 1965 1984 1985 Yuzawa Schuster Winter & Schäfer-Verwimp 1988 1992 2020 = Frullania obcordata pachoensis 686 1586 198 655 501 928 Spec Hepat 4: 629 Arquivos de botânica do Estado de Sao Paulo 1 (2): 47 [Brazil] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 86 [Brazil] Index Hepat 5: 389 J Hatt Bot Lab 56: 506 [checklist Brazil] Stephani Icon Hepat Inedit [Diastaloba] B-23ab, microfiche 48 fig 3253-3254 [from type; "Sta Catharina Brasilia leg Pabst"] 511 J Hatt Bot Lab 64: 445-447, fig 6(from type) [syn sub Fr. caulisequa] 603 Hepat N America 5: 120 5742 Frahmia 19: 13 [syn sub Fr. obcordata] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1911 Typus: Colombia. Pacho sd, Lindig in hb G Basionym: Distribution: Colombia spec nov - syn nov Stephani Bonner Gradstein & Hekking Menzel 1911 1965 1979 1984 686 655 483 142 Stephani 1985 928 Yuzawa & Koike 1989 519 = Frullania intumescens pacifica Spec Hepat 4: 503-504 Index Hepat 5: 390 J Hatt Bot Lab 45: 113 [Cat Colombia] Willdenowia 14: 506 [not in Peru, but in Colombia - error by Stephani 1911: 504] Stephani Icon Hepat Inedit [Thyopsiella] F- 77, microfiche 58 fig 3984 ["N Granada, Pacho, leg Lindig"] J Hatt Bot Lab 66: 355-356, fig 9 [from type], 10 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Taylor ex Mitt. by Seemann 1871 [orth var] Basionym: Distribution: Mitten in Seemann Merrill Stephani Stephani & Watts 1871 1906 1911 1914 637 1496 686 220 Zwickel 1932 663 = Frullania pacificae Flora Vitiensis: 417 [orth error; as "pacifica] Philippine J Sci C, Bot 5: 311 [Philippines: Mt. Pulog] Spec Hepat 4: 539-540 [orth error; as "pacifica"] J Proc Roy Soc New S Wales 48: 108 [Futuna; orth error; as "pacifica"] Beih Bot Centralbl 49: 637 [Ozellen] Taylor 1846 Frullania apiculata fo. aculeata Verd. 1930 Frullania apiculata (Reinw., Blume & Nees) Nees 1845 Page 316 pacificae Taylor 1846 Typus: Pacific Isles sd, Nightingale 384 ex hb BESCHERELLE , in hb G, ? hb HOOKER Basionym: Distribution: New Guinea, Australia, Vanuatu (Futuna), Samoa (Manua), Fiji (?Viti Levu) spec nov - syn nov Taylor Tayler in G., L. & N. Mitten in Seemann Stephani in Reinecke Stephani Stephani & Watts Verdoorn Verdoorn Zwickel Bonner Grolle & Schultze-Motel Miller, Whittier & Whittier Grolle & Piippo Stephani = Frullania apiculata 292 2848 637 1044 686 220 549 551 663 655 537 609 143 928 London J Bot 5: 406 Syn Hepat: 778 Flora Vitiensis: 417 [orth error; as "pacifica] Bot Jahrb 23: 315 [Samoa] [as pacifica] Spec Hepat 4: 539-540 [orth error; as "pacifica" J Proc Roy Soc New S Wales 48: 108 "pacifica" [Futuna] Nova Guinea 14 (4): 541 Nederl Kruidk Arch Jrg 1930 (2): Beih Bot Centralbl 49: 637 [Ozellen] Index Hepat 5: 390 J Hatt Bot Lab 36: 83 [Samoa] [sub Fr. apiculata] Prod Flora Hepat Polynesiae: 135 [orth err; as "pacifica"] Acta Bot Fennica 125: 22 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Thyopsiella] F-149, microfiche 59 fig 4088 ["Pacific Isles, Nightinggale legit", 232] fo. aculeata Verd. 1930 Frullania apiculata pallens 1846 1847 1871 1896 1911 1914 1930 1930 1932 1965 1972 1983 1984 1985 (Reinw., Blume & Nees) Nees 1845 Steph. 1894 Typus: Indonesia. Java sd, Paterson ex hb BROTHERUS, in hb G Basionym: Distribution: spec nov - syn nov Stephani Schiffner Stephani Verdoorn Verdoorn Bonner Stephani = Frullania meyeniana pallida 1894 1898 1911 1929 1930 1965 1985 228 831 686 245 293 655 928 Hedwigia 33: 165 Conspect Hepat Archip Indici, Batavia: 337 Spec Hepat 4: 556 Ann Bryol 2: 140-141 [Java] Ann Bryol Suppl 1: 81 [syn sub Fr. meyeniana] Index Hepat 5: 390 Stephani Icon Hepat Inedit [Thyopsiella] F-173, microfiche 59 fig 4115 ["Java (Paterson leg) publ Hedwigia 1894, p 165"] - 4116 Lindenb. 1845 Gottsche ex Verd. 1930 [pro syn] Basionym: Distribution: Verdoorn = Frullania ceylanica 1930 293 Ann Bryol Suppl 1: 95 [syn sub Fr. ceylanica] Nees 1845 Page 317 pallida Steph. 1911 Typus: Indonesia. Java coll ignot in hb G Basionym: Distribution: Indonesia (Java), Vanuatu (Aneityum) spec nov Stephani Stephani & Watts Verdoorn Verdoorn Clark Bonner Miller, Whittier & Whittier Stephani - syn nov 1911 1914 1929 1930 1955 1965 1983 1985 686 220 246 293 369 655 609 928 Spec Hepat 4: 641 J Proc Roy Soc New S Wales 48: 109 [Aneityum] Ann Bryol 2: 162 Ann Bryol Suppl 1: 97, 101 Bryologist 58: 68 [syn sub Fr. apiculata] Index Hepat 5: 390 Prod Flora Hepat Polynesiae: 135 Stephani Icon Hepat Inedit [Diastaloba] B-54, microfiche 49 fig 3294 ["Java"] = Frullania apiculata var. goebelii Schiffn. 1893 pallida Gottsche 1886 [nom nud] Typus: Sri Lanka: Poondeloya 4000-8000 ft 1868, John Nietner Drawings com. by Jack.: hb no 1540. Frullania pallida G. M. c. per. - 2 Blätter Basionym: Distribution: Gottsche Gottsche 0 1886 Icon inedit 2390 Bot. Centralblatt 26 (7): 204 [Sri lanka] Stephani Verdoorn Verdoorn Verdoorn Bonner Miller, Whittier & Whittier Stephani 1911 1928 1930 1930 1965 1983 1985 686 242 293 551 655 609 928 Hattori 1986 506 = pallidissima Steph. 1911 Typus: New Caledonia. hb L 107, in hb G Basionym: Distribution: spec nov - syn nov = Frullania heteromorpha palvistipula Spec Hepat 4: 578 Rev Bryol NS 1: 115 Ann Bryol Suppl 1: 170 [syn sub Fr. heteromorpha] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 391 Prod Flora Hepat Polynesiae: 135 Stephani Icon Hepat Inedit [Homotropantha] D- 7, microfiche 54 fig 3712 ["N Caledonia, Herb Leiden 107"] J Hatt Bot Lab 60: 220, fig 7 [from type] Schiffn. 1889 Steph. ex Ishiba 1908 [nom nud] Basionym: Distribution: = Frullania parvistipula Steph. 1910 Page 318 palvistipula Steph. ex Ishiba 1908 [nom nud] Typus: Japan Basionym: Distribution: Stephani in Ihsiba Stephani in Ihsiba Hattori = Frullania parvistipula 1908 1909 1944 929 890 943 Bot Mag Tokyo 22: 277 [Japan] Bot Mag Tokyo 23: 364 [Japan] Bull Tokyo Sci Mus 11: 150 [nom nud] 686 655 4761 194 928 Spec Hepat 4: 529 Index Hepat 5: 391 Nova Hedwigia 18: 436 The genus Frullania, separata: 436 Stephani Icon Hepat Inedit [Thyopsiella] F-120, microfiche 58 fig 4047 ["Panama, Dr Wagner leg"] Steph. 1910 panamana Steph. 1911 Typus: Panama sd, Wagner in hb G Basionym: Distribution: Panama spec nov Stephani Bonner Stotler Stotler Stephani = Frullania brasiliensis pancheriana Typus: Tahiti 1911 1965 1969 1970 1985 Raddi 1822 Steph. ex Bescherelle 1898 [nom nud] sd, Vieilland & Pancher Basionym: Distribution: Stephani in Bescherelle = Frullania pancheri panchetiana Typus: Tahiti 1898 871 J Bot Morot 12: 138 [spec nov sine desript.; = nom nud] Gottsche 1894 Steph. ex Verd. 1930 [pro syn] 1855, Vieillard & Panchet 91 in hb G (sub calcarata) Basionym: Distribution: Society Isl (Tahiti) nom herb - syn nov Verdoorn Verdoorn Bonner Miller, Whittier & Whittier Stephani 1930 1930 1965 1983 1985 Wiggington 2004 = Frullania calcarata 293 549 655 609 928 Ann Bryol Suppl 1: 62 Nova Guinea 14 (4): 540 Index Hepat 5: 391 [nom herb] Prod Flora Hepat Polynesiae: 135 Stephani Icon Hepat Inedit [Galeiloba] C-111, microfiche 51 fig 3496 ["Frullania calcarata Angstr, olim Frull panchetiana, Tahiti, Vieillard ... Panchat 1865"] 1172 Sript Bot Belg 30: 229 [syn sub Fr caffraria, West Africa] FALSCH !!!!!! muss eine andere Art sein Ångstr. 1873 Page 319 pandanicola Steph. 1924 Typus: Papua New Guinea: Madang, Schraderberg, Gebirgswald, 2070 m 27.5.1913, Ledermann 11631 in hb G 4089 & Basionym: Distribution: New Guinea spec nov - syn nov Stephani Verdoorn Clark Bonner Miller, Whittier & Whittier Grolle & Piippo Stephani 1924 1930 1955 1965 1983 1984 1985 Gao & Lai 2003 = Frullania apiculata Spec Hepat 6: 553 Nova Guinea 14 (4): 541 Bryologist 58: 68 Index Hepat 5: 351 Prod Flora Hepat Polynesiae: 135 Acta Bot Fennica 125: 22 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Thyopsiella] F-150, microfiche 59 fig 4089 [Nova Guinea (Ledermann legit)"] 1441 Illustrations Bryophytes China: 1117, fig 521(1-9) [China] fo. warburgii Verd. 1930 Frullania apiculata pandicola 687 549 369 655 609 143 928 (Reinw., Blume & Nees) Nees 1845 Steph. by Miller, Whittier & Whittier 1983 [orth var] Basionym: Distribution: Miller, Whittier & Whittier = Frullania pandanicola 609 Prod Flora Hepat Polynesiae: 135 [as "pandicola"] Steph. 1924 Frullania apiculata fo. warburgii Verd. 1930 Frullania apiculata papilliloba 1983 (Reinw., Blume & Nees) Nees 1845 Steph. 1911 Typus: New Caledonia "Nova Caledonica". Ins. Pinorum -.7.1908, Placide hb E.G. Paris 181, in hb G Basionym: Distribution: New Caledonia nom nud spec nov - syn nov Paris Stephani Verdoorn Verdoorn Verdoorn Verdoorn Bonner Hattori Miller, Whittier & Whittier Stephani = Frullania intermedia papilliloba 1910 1911 1928 1930 1930 1930 1965 1980 1983 1985 911 686 242 293 293 551 655 609 928 Rev Bryol 37 (1): 41 [New Caledonia: Isle of Pines] Spec Hepat 4: 575 Rev Bryol NS 1: 120 [syn sub Fr intermedia] Ann Bryol Suppl 1: 161 Ann Bryol Suppl 1: 161 Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 392 : 190 Prod Flora Hepat Polynesiae: 135 Stephani Icon Hepat Inedit [Homotropantha] D- 8, microfiche 54 fig 3713 ["N Caledonia, leg Placide, Gen Paris 181"] (Reinw., Blume & Nees) Nees 1845 Steph. ex Paris 1910 [nom nud] Basionym: Distribution: nom nud Paris = Frullania papilliloba 1910 911 Rev Bryol 37 (1): 41 [New Caledonia: Isle of Pines] Steph. 1911 Frullania intermedia (Reinw., Blume & Nees) Nees 1845 Page 320 papuana var. latistipula S.Hatt. ex S.Hatt. 1973 [nom nud] Typus: Papua New Guinea, Central Highlands, Mt. Wilhelm, Lake Pinde, 11850-12150ft RMS 67-6633 [holotype] Basionym: Distribution: Hattori Hattori Grolle & Piippo 1973 1974 1984 542 546 143 J Hatt Bot Lab 37: 150 J Hatt Bot Lab 38: 251 [nom nud] Acta Bot Fennica 125: 22 [Cat. West Melanesia] = Frullania papuana var. latistipula S.Hatt. 1974 Frullania papuana Verd. 1930 papuana var. latistipula S.Hatt. 1974 Typus: Papua New Guinea. Mt. Hagen, 2600m "New Guinea: Western Highlands, Mt. Hagen, near saw-mill. Nothofagus forest; + F.subdentata Steph. + F.spinosistipula Hatt." 21/10/1968, native collector in hb L L0060934 (hb VAN ZANTEN 68-3186/a) [holotypus] in hb JE - in hb NICH [ isotypus] in hb NICH [isotypus]; RMS 67-6633 [paratypus] Basionym: Distribution: var nov - syn nov Hattori Hattori Hattori Hattori & Piippo Mizutani, Hasegawa & Iwatsuki = Frullania papuana papulirostra 1974 1975 1982 1986 2009 546 547 495 557 2112 J Hatt Bot Lab 38: 251-254, fig 125-126 J Hatt Bot Lab 39: 304 [basionym of Fr. sinuata ssp. subapiculata] J Hatt Bot Lab 51: 217 [basionym of Fr. falsipapuana] Acta Bot Fennica 133: 32 Type sepecimens Hatt Bot Lab (NICH) 32: Holotype. New Guinea, Mt. Hagen. 2600 m. 1968. B. 0. van Zanten 68-3186/a ex herb. R. Grolle (NICH 431943). Verd. 1930 Steph. 1911 Typus: Hawaii. Kauai, Kilauea 1910, Faurie 43 in hb G-20589 [holotypus] Basionym: Distribution: Hawaii (Kauai) spec nov - syn nov Stephani Verdoorn Verdoorn Miller Miller Bonner Miller, Whittier & Whittier Stephani 1911 1930 1930 1953 1963 1965 1983 1985 Konrat & Braggins So & Wang 2001 2006 = Frullania hypoleuca 686 293 551 349 834 655 609 928 Spec Hepat 4: 652-653 Ann Bryol Suppl 1: 126 [syn sub Fr. hypoleuca] Nederl Kruidk Arch Jrg 1930 (2): Bryologist 56: 47 [sub Fr hypoleuca] Ark Bot, ser 2, 5: 519 [sub Fr. hypoleuca] Index Hepat 5: 392 Prod Flora Hepat Polynesiae: 135 Stephani Icon Hepat Inedit [Diastaloba] B-75, microfiche 49 fig 3319 [from type; "Hawai, leg Faurie 1910, Insula Hawai No 43"] 838 J Hatt Bot Lab 91: 250-256 [sub Fr. hypoleuca] 1466 J H B L 100: 423 [sub Fr. hypoleuca] Nees 1843 Page 321 paraensis Steph. 1911 Typus: Brazil: Para sd, Spruce sn in hb G 23572 [lectotypus by Bonner 1965, Gradstein & Hekking 1989] Basionym: Distribution: Guyana, Brazil spec nov Stephani Bonner Yana Stephani 1911 1965 1984 1985 686 655 501 928 - syn nov Yuzawa 1988 511 Gradstein & Hekking Gradstein & Hekking Gradstein & Costa 1989 1989 2003 517 517 984 = Frullania exilis Taylor 1846 Frullania apiculata paranaensis Spec Hepat 4: 630 [Brasilia, Guiana] Index Hepat 5: 393 J Hatt Bot Lab 56: 506 [checklist Brazil] Stephani Icon Hepat Inedit [Diastaloba] B-24, microfiche 48 fig 3255 ["Para (Amazonas)"] J Hatt Bot Lab 64: 437-439, fig. 1 [from type] [syn nov syn sub Fr. exilis TAYLOR 1846] J Hatt Bot Lab 66: 220 [syn sub Fr. apiculata] J Hatt Bot Lab 66: 208 [Guianas] Mem New York Bot Gard 87: 83 [Brazil] [sub Fr apiculata] (Reinw., Blume & Nees) Nees 1845 Steph. by Yano 1984 [orth var] Basionym: Distribution: Yano 1984 = Frullania paranensis parasitica 272} 501 J Hatt Bot Lab 56: 506 [checklist Brazil] Steph. 1911 [auct. non Hampe ex Lehm. 1838] Mont. 1840 {French Guiana: Leprieur No Typus: Basionym: Distribution: Guadeloupe -syn nov - syn nov Montagne Montagne Stotler Stotler Gradstein & Costa Yano = Frullania neesii 1840 1855 1969 1970 2003 2008 Ann Sci Nat Bot, ser 2, 14: 333 Ann Sci Nat Bot, ser 4, 3: 320 [syn sub Fr leprieurii] Nova Hedwigia 18: 450 [syn sub Fr. neesii] The genus Frullania, separata: 450 [syn sub Fr. neesii] Mem New York Bot Gard 87: 87 [Brazil] [syn sub Fr kunzei] Boletim Instituto Botanica 19: 90 [syn sub Fr neesii; Brazil; catalog] Lindenb. 1845 Frullania kunzei parasitica 467 475 4761 194 984 1684 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Lehm. & Lindenb. by Herzog 1953 [orth var] Basionym: Distribution: Herzog = Frullania parasitica 1953 640 Rev Bryol Lichen 20: 129 [Costa Rica] Hampe ex Lehm. 1838 Frullania obcordata (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Page 322 parasitica Hampe ex Lehm. 1838 Typus: Guiana "America tropica, in Frullanaia gibbosa parasitans"? ["Crescit parasitica ad Frullaniam obscuram"] sd, Leprieur 272 in hb HAMPE, typus in hb S (B24186), in hb G (isotypus) in hb GLM 06015 Basionym: Distribution: Guatemala, Costa Rica, Panama, Guadeloupe, Martinique, Dominica, French Guiana, Brazil spec nov - syn nov Hampe ex Lehmann Hampe in G., L. & N. Lindenberg in G., L. & N. Duss 1838 1845 1845 1904 Stephani Clark & Svihla 1911 1948 Herzog Arnell Arnell Bonner Yano Stephani 1953 1959 1959 1965 1984 1985 Fulford Gradstein & Hekking Boggan, Funk, Kelloff et al. Boggan, Funk, Kelloff, Hoff, Cremers & Feuillet Stotler, Allen, Gradstein, McGuinness, Whittemore & Chung Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Gradstein & Costa 1987 1989 1992 1997 Pugillus 7: 11-12 Syn Hepat: 444-445 Syn Hepat: 449 [syn sub Fr. leprieuri] Flore Cryptogamique des Antilles Françaises: 168 [Bois des BainsJaunes] 686 Spec Hepat 4: 631 325 Bryologist 51: 26 [as Fr. parasitica, key to Fr. of Middle America: Brazil, Guiana (Dominica)] 640 Rev Bryol Lichen 20: 129 [Costa Rica] 448 Svensk Bot Tidskr 53 (4): 500 [Martinique] 448 Svensk Bot Tidskr 53 (4): 505 [French Guiana] 655 Index Hepat 5: 393-394 501 J Hatt Bot Lab 56: 506 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Diastaloba] B-25, microfiche 48 fig 3256 ["America tropica Original"] 99 Mem NY Bot Gard 45: 272 [Dominica] 517 J Hatt Bot Lab 66: 209 [French Guiana] 4771 p. 16 - Checklist of the plants of the Guianas 1396 Checklist Guianas: 10 [French Guiana] 1998 703 Trop Bryol 15: 180 [Checklist Panama] 2002 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 2003 984 Yano 2008 1684 Schäfer-Verwimp 2010 2460 Lavocat Bernard & SchäferVerwimp Winter & Gradstein 2011 3130 2020 5742 Mem New York Bot Gard 87: 86 [Brazil] [syn sub Fr caulisequa - no statement] Boletim Instituto Botanica 19: 90 [syn sub Fr caulisequa; Brazil; catalog] Crypt Bryol 31(4): 346 [syn sub Frull caulisequa, Dominica, West Indies] Cryptogamie, Bryologie 32(3): 251 [Martinique] "as Fr parasitica Arnell 1959" Frahmia 19: 13 [syn sub Fr obcordata] = Frullania obcordata 588 2850 2850 3132 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 parhamii (R.M.Schust.) R.M.Schust. 1992 [nom inval: (Art. 34.4, basionym not cited, McNeill et al. 2006] Basionym: Neohattoria parhamii Distribution: Fiji comb nov R.M.Schust. 1963 Schuster 1963 Schuster Schuster Konrat, Braggins, Asakawa & Toyota 1970 1992 2006 = Frullania parhamii Hagborg 2011 745 J Hatt Bot Lab 26: 243 [as Neohattoria parhami; key to Neohattoria in New Caledonia] 761 J Hatt Bot Lab 33: 292-295, fig 6 [as Neohattoria parhami] 603 Hepat N America 5: 34 1500 Bryologist 109(2): 154 [reassessment of Schusterella] (R.M.Schust.) R.M.Schust. ex von Konrat, L. Söderstr. & A. Page 323 parisii Steph. in Paris 1908 [nom nud] Typus: Timbo Basionym: Distribution: spec nov Stephani in Paris Vanden Berghen Wigginton = Frullania parisii 790 Rev Bryol 35: 6 [Stephani in litt] 185 Bull Jard Bot Nat Belg 46 (1/2): 199 [parisii 1908 = nom nud] 1172 Sript Bot Belg 30: 227 [syn sub Fr trinervis, West Africa] Steph. 1910 Frullania trinervis parisii 1908 1976 2004 (Lehm.) Drège 1843 Steph. 1910 Typus: French Guiana: Guinea gallica, Seguiri -.10.1903, Pobeguin 6 ex hb E. B. (Paris) in hb G 17314 Basionym: Distribution: spec nov - syn nov Stephani Arnell Bonner Vanden Berghen Stephani 1910 1963 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania trinervis parva 686 822 655 185 928 Spec Hepat 4: 381 [as spec nov] Hepat S Africa: 141 [syn nov sub Fr. trinervis] Index Hepat 5: 394 Bull Jard Bot Nat Belg 46 (1/2): 199 [parisii 1908 = nom nud] Stephani Icon Hepat Inedit [Galeiloba] C- 31, microfiche 50 fig 3383 [from type; "Africa, Seguiri - Guinea gallica, leg H Pobeguin 1903"] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 227 [syn sub Fr trinervis, West Africa] (Lehm.) Drège 1843 Steph. 1924 Typus: Mexico 1913, Arsén 7789 in hb G 20973 [holotypus]; Basionym: Distribution: Mexico spec nov - syn nov Stephani Bonner Yuzawa Stephani 1924 1965 1983 1985 687 655 498 928 Fulford & Sharp Yuzawa 1990 1991 105 525 = Frullania cuencensis Spec Hepat 6: 534 Index Hepat 5: 394 J Hatt Bot Lab 54: 114-115 (No 5), fig. 5 (a-q) [from type] Stephani Icon Hepat Inedit [Chonanthelia] A- 75, microfiche 47 fig 3177 [from type; "Mexico Arsén 7789"] Mem NY Bot Gard 63: 45 [Mexico] J Hatt Bot Lab 70: 209 [syn sub Fr. cuencencis] Taylor 1846 Page 324 parviflora Steph. 1911 [non Steph. 1910] Typus: Mexico, in rupibus sd, s coll. ex hb NEES, ex hb JACK, in hb G Basionym: Distribution: Mexico spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 686 655 4761 194 928 Fulford & Sharp 1990 105 = Frullania flexicaulis Spruce 1884 Frullania paradoxa parvilobula Spec Hepat 4: 503 Index Hepat 5: 394 Nova Hedwigia 18: 463-464 The genus Frullania, separata: 463-464 Stephani Icon Hepat Inedit [Thyopsiella] F- 78, microfiche 58 fig 3985 ["Mexico (Herb Jack) ex Nees"] - 3986 Mem NY Bot Gard 63: 45 [Mexico] Lehm. & Lindenb. 1845 Steph. 1911 Typus: Hawaii. Kalihi -.10.1909, Faurie 181 in hb G 20590 [holotypus] Basionym: Distribution: Hawaii spec nov - syn nov Stephani Verdoorn Miller Miller Bonner Hattori Miller, Whittier & Whittier Stephani 1911 1930 1953 1963 1965 1981 1983 1985 Konrat & Braggins So & Wang 2001 2006 = Frullania hypoleuca parvistipula 686 293 349 834 655 492 609 928 Spec Hepat 4: 642 Ann Bryol Suppl 1: 126 [syn sub Fr. hypoleuca] Bryologist 56: 47 [sub Fr hypoleuca] Ark Bot, ser 2, 5: 519 [sub Fr. hypoleuca] Index Hepat 5: 394-395 J Hatt Bot Lab 49: 373 [only Fr. neurota in type material] Prod Flora Hepat Polynesiae: 135 Stephani Icon Hepat Inedit [Diastaloba] B-55, microfiche 49 fig 3295 ["Hawai Baldwin Levier 2467, auch (Faurie No 181 1909 olim Fr. hypoleuca"] 838 J Hatt Bot Lab 91: 250-256 [syn nov sub Fr. hypoleuca] 1466 J H B L 100: 423 [sub Fr. hypoleuca] Nees 1843 Steph. ex Ihsiba 1908 [nom nud] Basionym: Distribution: Ihshiba = Frullania parvistipula 1908 927 Bot Mag Tokyo 22: 227 Steph. 1910 Page 325 patens Lindenb. 1845 Typus: Brazil sd, Martius in hb LINDENBERG (sub Jungermannia atrata), typus in hb S (B24188, 24189) in hb GLM (hb LEHMANN 53) 06021 Basionym: Distribution: Brasil spec nov Lindenberg in G., L. & N. Dusen Stephani Massalongo Schiffner & Arnell 1845 1905 1911 1911 1964 - syn nov - syn nov Stotler Stotler Stephani 1969 1970 1985 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 2002 = Frullania mucronata Syn Hepat: 462 Arch Mus Nac Rio de Janeiro 13: 119 [Brasil] Spec Hepat 4: 519 Rev Bryol 38: 10 [Brasil] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 88 [syn sub Fr. divergens] 4761 Nova Hedwigia 18: 446 [syn sub Fr. mucronata] 194 The genus Frullania, separata: 446 [syn sub Fr. mucronata] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 79, microfiche 58 fig 3987 - 3988 [Brasilia (Pabst legit)"] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] (Lehm. & Lindenb. in Lehm.) Lehm. & Lindenb. 1845 Frullania brasiliensis patentiloba 2850 710 686 903 198 Raddi 1822 Steph. 1911 Typus: Chile, Prov. Chiloé, J Guafo 25.7.1908, Cta. Samuel in hb HALLE 118 [lectotypus by Engel 1978], hb UPS [cum per. + Sporo] / in G 2833 [lectotypus by YUZAWA 1988] Basionym: Distribution: Chile (Chiloe, Valdivia, Magallanes), Argentina (Patagonia, Tierra del Fuego), Falkland Isl [Argentinia] spec nov spec nov - syn nov Stephani Stephani Herzog Kühnemann 1911 1911 1923 1937 686 721 112 1031 Kühnemann Bonner Engel 1949 1965 1978 657 655 182 Stephani 1985 928 Engel Yuzawa 1988 1988 111 511 = Frullania boveana pauciexiliens Spec Hepat 4: 666 Kungl Svenska Vetenskapsakad Handl 45 (9): 89, fig 35(d-g) Hedwigia 64: 8 [Chile, Valdivia] Revista Centro Estud Doct Ci Nat, Buenos Aires 1 (3): 167 [Malvinas, Tierra del Fuego] Lilloa 19: 355 [Argentina:Tierra del Fuego] Index Hepat 5: 395 Fieldiana Bot 41: 235 [syn. nov. = boveana / lectotype nov: HALLE 118] Stephani Icon Hepat Inedit [Diastaloba] B-46, microfiche 49 fig 3284 ["Chile, Corral, 5/6 96 Dusen 170 (olim stipatiloba"] Fieldiana Bot NS 25: 162 [sub Fr boveana] J Hatt Bot Lab 64: 439-440, fig. [from type] [syn. nov. = stipatiloba / lectotype nov: G20833] C.Massal. 1885 von Konrat & Braggins in Konrat & Pickering 2004 [nom nud] Basionym: Distribution: New Zealand (South Isl: Coromandel, Table Mountain) Konrat & Pickering 2003 Konrat & Brennan 2004 1000 http://www.discoverlife.org/nh/tx/Plantae/Bryophyta/ [IDnature guide for Liverworts, Frullania, New Zealand] 1164 http://www.liverworts.org/dich_key/frul_key.shtml, 6th of 6 pages [Key to Frullania of New Zealand; Coromandel (Table Land), South Island] = Page 326 paucipinna Steph. 1911 Typus: Bolivia, im Urwlad des Rio Blanco, 15 ° lat S -.8.1907, T. Herzog in hb G (hb LEVIER 5853) Basionym: Distribution: Bolivia spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 686 655 4761 194 928 Gradstein, Meneses & Arbe 2003 975 = Frullania triquetra pedicellata Spec Hepat 4: 502-503 Index Hepat 5: 396 Nova Hedwigia 18: 457-458 [syn sub Fr triquetra] The genus Frullania, separata: 457-458 [syn sub Fr triquetra] Stephani Icon Hepat Inedit [Thyopsiella] F- 80, microfiche 58 fig 3989 ["Bolivia, leg Herzog, Levier 5853"] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr triquetra] Lindenb. & Gottsche 1847 [auct. non Steph. 1897] C.Gao & Chuang 1981 {China} Basionym: Distribution: - syn nov - syn nov Gao & Chang 1981 Liu Hattori & Lin Aur & Zhang Xu Bapna & Kachroo 1985 1985 1985 1989 2000 = Frullania muscicola pedicellata Steph. 1894 fo. kochiensis (Steph.) Kamim. 1952 Basionym: Frullania kochiensis Distribution: Japan, Korea comb nov - syn nov 1021 Fl Hep Chinae bor-orient: 152, fig 67(9-17) [China] [syn sub Fr. pedicellata] 2801 Invest Stud Nat 5: <133-155> [aus 698] 505 J Hatt Bot Lab 59: 140 [syn sub Fr. muscicola] 1022 Fl Heilongjiangensis 1: 163, fig 56(10-18) [China] [aus 698] 712 Hepaticology in India II: 175 [syn sub Fr. muscicola] Steph. 1910 Kamimura Kamimura Bonner Choe 1952 1961 1965 1980 873 199 655 1171 Gao & Chang 1981 1021 Aur & Zhang Yuzawa Yamada & Iwatsuki 1985 2001 2006 1022 1660 1470 = Frullania pedicellata Contrib Fl Hepat Shikokuensem, Kochi: 130 J Hatt Bot Lab 24: 40 [syn nov sub Fr kochiensis] Index Hepat 5: 396-397 Illust Fl & Fauna Korea 24: 735 [syn nov sub Fr pedicellata; South Korea] Fl Hep Chinae bor-orient: 152 [syn sub Fr. pedicellata; #1612 or #2110] Fl Heilongjiangensis 1: 163 [sub Fr pedicellata] Natural Environmental Sci Res 14: 35 [Japan, syn sub Fr pedicellata] J Hatt Bot Lab 99: 32 [Catalog of Japan] Steph. 1897 pedicellata var. hamatiloba (Steph.) S.Hatt. 1951 [nom inval] Basionym: Frullania hamatiloba Distribution: Japan, Taiwan Steph. 1910 Hattori Bonner Kuo & Chiang Piippo 1951 1965 1988 1990 Yamada & Iwatsuki 2006 = Frullania hamatiloba 528 655 948 521 J Hatt Bot Lab 5: 67 [comb nov] Index Hepat 5: 397 Taiwania 33: 13 [Index of Taiwan] J Hatt Bot Lab 68: 46 [basionym of Fr. hamatiloba; annotated catalogue of China] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] Steph. 1910 Page 327 pehlkeana Steph. 1910 Typus: Colombia, Bogota 8800ft -.1.1898, Pehlke 2 ex hb WINKELMANN, in hb G [holotypus] Basionym: Distribution: Colombia spec nov Stephani Herzog Bonner Gradstein & Hekking Stephani 1910 1955 1965 1979 1985 686 107 655 483 928 - syn nov Yuzawa Zanoni Schäfer-Verwimp & Pócs 1991 1993 2009 525 = Frullania cuencensis pendrostyla Spec Hepat 4: 340 Feddes Repert 57 (1/2): 171 [? same as Fr. cuencensis] Index Hepat 5: 397 J Hatt Bot Lab 45: 113 [Cat Colombia] Stephani Icon Hepat Inedit [Chonanthelia] A- 76, microfiche 47 fig 3178 [from type; "Bogota 8800ft 1898 leg Pehlke"] J Hatt Bot Lab 70: 209 [syn sub Fr cuencensis] 1985 Acta Bot Hungarica 51(3-4): 385 [Dominican Republic, Hispaniola] Taylor 1846 Steph. by S.Hatt. 1973 [orth var] Basionym: Distribution: Hattori 1973 = Frullania pendulostyla Frullania mirabilis pendula 124 J Jap Bot 48 (4): 107 Steph. 1924 J.B.Jack & Steph. 1892 Austin 1872 [hom illeg] [non Mitten 1861, nec Kamim. 1952] Typus: Fiji. "Viti Isles" 1860, Seemann 834pp in hb SULLIVANT Basionym: Distribution: Fiji spec nov - syn nov Austin Stephani Bonner Inoue Miller, Whittier & Whittier Stephani = Frullania deflexa 1872 1911 1965 1966 1983 1985 666 686 655 1703 609 928 Bull Torrey Bot Club 3 [No 3]: 16-17 Spec Hepat 4: 587 [syn sub Fr. deflexa] Index Hepat 5: 397 J Jap Bot 41 (1): 15 Prod Flora Hepat Polynesiae: 135 Stephani Icon Hepat Inedit [Homotropantha] D-19a, microfiche 54 fig 3732 ["syn: Frull pendula Austin 1872, syn: Frull Austiniana Pearson ms, Insulae vitienses, Seemann Original"] Mitt. 1862 Page 328 pendula [auct. non Mitt. 1861, nec Austin 1872] Kamim. 1952 {Japan} Typus: Japan. Honshu, Hyogo Pref, Prov. Settsu, Mt. Rokko -.10.1936, K. Uno 3719 in hb Kamim. [holotypus], in hb TNS [isotypus] Basionym: Distribution: spec nov nom nov nom nov - syn nov Kamimura Hattori & Noguchi Bonner Inoue Kamimura Inoue Hattori & Lin Inoue N.N. [National Science Museum Tokyo] Yamada & Iwatsuki Mizutani, Hasegawa & Iwatsuki = Frullania tamarisci ssp. obscura 1952 1960 1965 1966 1970 1972 1985 1987 2000 769 965 655 1703 1552 123 505 1155 995 Acta Phytotax Geobot 14 (4): 108-109, fig 2(a-q) Index spec Typ herb Japon, Bryophyta: 18 [holotype} Index Hepat 5: 398 J Jap Bot 41 (1): 14-15 [new name Fr kamimurae] Bulletin of Kochi Gakuen Junior College 2: 24 (new name Fr akisawai] J Jap Bot 47 (11): 349 [syn sub Fr. kamimurae] J Hatt Bot Lab 59: 161 [syn sub Fr. tamarisci ssp. obscura] Index type specimens in Nat Sci Mus, Tokyo: 14 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 2006 2009 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] 2112 Type sepecimens Hatt Bot Lab (NICH) 32: Isotype. Japan, Hoshu, Hyogo-ken, Prov. Settsu, Mt. Rokko. s.d. N.Ui 3719 (NICH H16556). (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata pendula (Reinw., Blume & Nees) Mont. 1842 Mitt. 1861 [non Austin 1872, nec Kamim. 1952] Typus: Sri Lanka [Ceylon] sd, Gardner 134 in hb NY 1048703 (hb W. MITTEN - sd, Collector unspecified 134) [holotypus] in hb G [holotypus] ex hb K 1556 in hb BM [isotypus] in hb FH [isotypus] Basionym: Distribution: Sri Lanka spec nov - syn nov Mitten Stephani Stephani Verdoorn Bonner Inoue Hattori 1861 1911 1911 1930 1965 1966 1975 Miller, Whittier & Whittier Stephani 1983 1985 Uribe 2008 Uribe-M. 2011 = Frullania ternatensis pendulosa 574 686 686 293 655 1703 47 J Linn Soc Bot 5: 122 [Sri Lanka] Spec Hepat 4: 552 [syn sub Fr. ceylanica] Spec Hepat 4: 614 [= sgen Meteoriopsis] Ann Bryol Suppl 1: 141 [syn nov sub Fr. ternatensis] Index Hepat 5: 397 J Jap Bot 41 (1): 15 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 77-78, fig 1(a-d) [About Mitten´s specimen] 609 Prod Flora Hepat Polynesiae: 135 928 Stephani Icon Hepat Inedit [Meteriopsis] E-61ab, microfiche 56 fig 3858 ["Ceylon, Gardner 134, Original"] - 3859 ["Mindanao, Mt Malindang 4801"] - 3860 [missing] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella,; syn sub Fr ternatensis] [Monography of subgenus Meteoriopsis] 2767 Caldasia 33(1): 74 [type study, belongs to ternatensis] Gottsche 1846 Steph. by Cole 1983 [orth var] Basionym: Distribution: Cole = Frullania pendulostyla Frullania mirabilis 1983 1407 Brenesia 21: 154 [Costa Rica] Steph. 1924 J.B.Jack & Steph. 1892 Page 329 pendulostyla Steph. 1924 Typus: Ecuador. Gualaquiza Oriente, V. Bomboiza, 1200m 23.9.1910, Allioni 600bis in hb G 13655 Basionym: Distribution: Costa Rica, Venezuela, Colombia, Ecuador, Guyana spec nov - syn nov Stephani Bonner Stotler Stotler Hattori Gradstein & Hekking Mägdefrau Stephani 1924 1965 1969 1970 1973 1979 1983 1985 Kron Gradstein & Florschütz-de Morales Z. Boggan, Funk, Kelloff, Hoff, Cremers & Feuillet Holz 1988 1989 1991 1997 Uribe 2008 Gradstein & Perez 2021 = Frullania mirabilis pennsylvanica 2004 687 655 4761 194 124 483 1206 928 Spec Hepat 6: 547 Index Hepat 5: 398 Nova Hedwigia 18: 473-474, pl 37(311-317) The genus Frullania, separata: 473-474, pl 37(311-317) J Jap Bot 48 (4): 104-109, fig 2 [syn sub Fr. mirabilis] J Hatt Bot Lab 45: 113 [Cat Colombia] Nova Hedwigia 13: 21 [Venezuela] Stephani Icon Hepat Inedit [Thyopsiella] F- 82, microfiche 58 fig 3991 ["Ecuador, Allioni 600bis (1912)"] 510 J Hatt Bot Lab 64: 354 1237 Trop Bryol 1: 42 [Venezuela, Mt Roraima] 307 Trop Bryol 4: 38 [Costa Rica] 1396 Checklist Guianas: 10 [Guyana] 1184 [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] 6056 Candollea 76: 43, fig 2 (Label of holotype of Frullania pendulostyla Steph. at G.) [Ecuador] J.B.Jack & Steph. 1892 Steph. 1883 Typus: USA. Pennsylvania, Carbon Cty, Stony Creek sd, E.A.Rau 1470 in hb G G-012313 [lectotypus]; G-16756 & FH-34217 [lectoisotypus] Basionym: Distribution: spec nov spec nov - comb nov Stephani Stephani Evans Stephani Verdoorn 1883 1883 1905 1911 1928 848 1123 1451 686 244 Bonner Guerke Guerke Fulford & Sharp 1965 1978 1978 1990 655 181 181 105 = Jubula hutchinsiae ssp. pennsylvanica pentapleura Bull Torrey Bot Club 10 (12): 132 Hedwigia 22 (10): 147-148 Rhodora 7 (75): 55-58 Spec Hepat 4: 690 Ann Crypt exot 1 (2): 215 [= Jubula hutchinsiae subsp. pennsylvanica (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 215, 1928] Index Hepat 5: 398 Bryophyt Bibl 17: 83 Bryophyt Bibl 17: 79-92 [lectotype of Jubula pennsylvanica] Mem NY Bot Gard 63: 45 [Mexico] (Steph.) Verd. 1928 [auct. non Taylor 1846] Rodway 1916 {Tasmania} Basionym: Distribution: Rodway Hattori = Frullania probosciphora 1916 1983 914 499 Papers Proceed Roy Soc Tasmania for 1916: 137 [Tasmania] J Hatt Bot Lab 54: 159 Taylor 1846 Page 330 pentapleura Typus: [auct. non Taylor 1846] Steph. & Watts 1914 {Lord Howe Isl} WATTS 1 (det Steph), WATTS 7 (det Steph), WATTS 36 & 45 (det Steph) Basionym: Distribution: Stephani & Watts Hattori = Frullania fulfordiae pentapleura 1914 1987 220 400 J Proc Roy Soc New S Wales 48: 109 [Lord Howe Isl] Bryologist 90 (4): 366 S.Hatt. 1987 [auct. non Taylor 1846] Mitt. in Hooker 1855 {New Zealand} Basionym: Distribution: auct - syn nov Mitten in Hooker Hooker Hodgson Hodgson Hattori Glenny 1855 1867 1949 1965 1983 1998 = Frullania solanderiana pentapleura ssp. falsa 560 Antarctic voyage, II. Fl Novae-Zelandiae 2: 162 [New Zealand] 1154 Handbook New Zealand Flora: 537 [New Zealand, N Island] 203 Trans Roy Soc New Zealand 77 (3): 376 Trans Proc Roy Soc New Zealand, Bot 3 (4): 89 499 J Hatt Bot Lab 54: 159 & 170 1412 Tuhinga 10: 122 [New Zealand specimens are all Fr solanderiana] Colenso 1889 (Steph.) S.Hatt. 1988 Basionym: Frullania falsa Steph. 1910 Distribution: Australia (NSW), Tasmania, Lord Howe Isl comb nov -syn nov Ratkowsky Hattori Konrat & Braggins Carthy McCarthy 1987 1988 2001 2003 2006 = Frullania probosciphora pentaplicata "5plicata" Typus: New Zealand 695 429 838 1125 1385 Papers Proceed Roy Soc Tasmania 121: 157 [Checklist Tasmania] Nova Hedwigia Beih 90: 153 [perianth with 8-9 plicae] J Hatt Bot Lab 91: 245-250 [syn nov sub Fr. probosciphora] Flora Australia Suppl Ser 21: 45 [sub Fr probosciphora] Checklist Australian Liverworts [Australia, syn sub Fr probosciphora] Taylor 1846 Steph. ex Steph. 1985 [nom herb] {New Zealand} sd, leg Knight Basionym: Distribution: New Zealand Stephani 1985 928 Stephani Icon Hepat Inedit [Chonanthelia] A-107a, microfiche 48 fig 3223 ["Frullania 5plicata St New Zealand leg Knight"] = perminata var. angulistipula S.Hatt. by Mizutani, Hasegawa & Iwatsuki 2009 [orth var] Basionym: Distribution: orth var Mizutani, Hasegawa & Iwatsuki 2009 2112 ype sepecimens Hatt Bot Lab (NICH) 32: Isotype. New Guinea, Eastern Highlands, Mt. Wilhelm. 4200-4300 m. 1968. B. 0. van Zanten 68-795/a (NICH 432024). = Frullania perminuta var. angulistipula S.Hatt. 1974 Frullania rudolfiana S.Hatt. 1972 Page 331 perminuta S.Hatt. 1972 Typus: Papua New Guinea, Morobe, Wau Distr, Summit of Mt. Kaindi, 2200-2350m 26.5.1967 - 27.5.1967, R.M. Schuster 67-6158 in hb NICH [holotypus] in hb JE, MASS [isotypus] Basionym: Distribution: spec nov - syn nov Hattori Hattori Hattori Hattori Hattori Hattori Grolle & Piippo Mizutani, Hasegawa & Iwatsuki 1972 1973 1975 1975 1976 1982 1984 2009 539 542 66 547 80 495 143 2112 J Hatt Bot Lab 36: 440-443, fig 18 J Hatt Bot Lab 37: 151 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 141-142, fig 158 J Hatt Bot Lab 39: 299 J Hatt Bot Lab 40: 492-493, fig 191(a-d) [Borneo] J Hatt Bot Lab 51: 233 [syn sub Fr. junghuhniana var. minutissima] Acta Bot Fennica 125: 22 [Cat. West Melanesia] Type sepecimens Hatt Bot Lab (NICH) 32: Holotype. New Guinea, Wau Distr., Mt. Kaindi. 2200-2350 m. 26-27 May 1967. R. M. Schuster 67-6158 (NICH 309937). = Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 perminuta var. angulistipula S.Hatt. 1974 Typus: Papua New Guinea. Simbu, Mt. Wilhelm, 4200-4300m "New Guinea: Eastern Highlands, Mt. Wilhelm below summit. On branch of Coprosma-shrub, exposed. " 8/7/1968, Mc Vean 68795a in hb L L0060935 ( hb VAN ZANTEN 68-795/a) [typus] in hb JE [holotypus] in hb NICH [isotypus] Basionym: Distribution: var nov - syn nov Hattori Hattori Grolle & Piippo Mizutani, Hasegawa & Iwatsuki = Frullania rudolfiana 1974 1982 1984 2009 546 495 143 2112 J Hatt Bot Lab 38: 255-256, fig 128 J Hatt Bot Lab 51: 234 Acta Bot Fennica 125: 22 [Cat. West Melanesia] Type sepecimens Hatt Bot Lab (NICH) 32: Isotype. New Guinea, Eastern Highlands, Mt. Wilhelm. 4200-4300 m. 1968. B. 0. van Zanten 68-795/a (NICH 432024). S.Hatt. 1972 perocellata Onraedt 1978 Typus: Sri Lanka 4.3.1976, Onraedt 76.L.3156/b in hb ONRAEDT [holotypus] Basionym: Distribution: spec nov - syn nov Onraedt Onraedt Hattori & Lin = Frullania alstonii 1978 1981 1985 443 Rev Bryol Lichen 44 (1): 77-79, fig. 1 [key to indo-malay species] 5794 J Hatt Bot Lab 50: 202 [Sri Lanka] 505 J Hatt Bot Lab 59: 159 [syn sub Fr. alstonii] Verd. 1930 Page 332 perrotana Steph. 1911 Typus: Madagascar: Silva Fito, Tamatare -.7.1897, Perrot sn hb LEVIER 1202, in hb G & Icon ined 1152 Basionym: Distribution: spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2004 = Frullania apiculata peruviana 686 655 185 928 Spec Hepat 4: 621 Index Hepat 5: 398-399 Bull Jard Bot Nat Belg 46 (1/2): 102 Stephani Icon Hepat Inedit [Diastaloba] B- 3ab, microfiche 48 fig 3228 ["Madagascar Perrot 1897 Levier No 1202 ex ..."] - 3229 ["Madagascar Tamatare leg Perrot"] 250 Bryophyt Bibl 48: 63 1172 Sript Bot Belg 30: 217 [syn sub Fr apiculata, West Africa] (Reinw., Blume & Nees) Nees 1845 [auct. non Gottsche 1846] Spruce 1884 Typus: Peru. "Peruvian Andes", Mt. Guayrapurna sd, Spruce Basionym: Distribution: - syn nov Spruce 1884 243 Spruce in Stephani Bonner Menzel Stephani 1911 1965 1984 1985 686 655 142 928 = Frullania crinoidea Trans Proc Bot Soc Edinburgh 15: 38 & 40 [No 29] [not identical with Fr. peruviana GOTTSCHE] Spec Hepat 4: 604 [syn sub Fr. crinoidea Spruce, Hepat. exsiccatae]] Index Hepat 5: 399 Willdenowia 14: 489 [hom illeg] & 510 [syn of Fr. crinoidea] Stephani Icon Hepat Inedit [Meteriopsis] E-19, microfiche 55 fig 3792 ["(Fr peruviana Hep Am), Mte Guayrapurina, Peruvia, ab ipso auctore"] - 3793 Spruce ex Steph. in Steph. 1911 Frullania atrata (Sw.) Nees ex Mont. 1839 peruviana var. ß Gottsche 1865 Basionym: Distribution: Gottsche Uribe & Gradstein = Frullania peruviana 1864 1998 478 707 Ann Sci Nat Bot, ser 5, 1: 175 Bryophyt Bibl 53: 37 Gottsche 1846 [non Gottsche ex Spruce 1884] Page 333 perversa Steph. 1894 Typus: Philippines. 1870, G. Wallis in hb G 15011 (ex hb JACK) Basionym: Distribution: Malaysia (Sabah), Borneo, Philippines spec nov - comb nov Stephani Stephani Verdoorn Horikawa 1894 1911 1930 1943 Bonner Schuster 1965 1970 Hattori Hattori Hattori 1973 1974 1974 Kamimura 1974 Stephani 1985 228 686 293 1283 Hedwigia 33: 165-166 Spec Hepat 4: 639 Ann Bryol Suppl 1: 136, fig 209 Acta Phytotax Geobot 13: 213 [comb nov: Fr. junghuhniana var. perversa] 655 Index Hepat 5: 399 761 J Hatt Bot Lab 33: 288 [comb nov: Neohattoria (Microfrullania) perversa] 540 J Hatt Bot Lab 37: 57-58, fig 23 544 J Hatt Bot Lab 38: 176-177 [Luzon] 546 J Hatt Bot Lab 38: 241-242 [syn sub Fr. junghuhniana subsp denticulilobula] 3692 Bull Kochi Gakuen J Coll 5: 83 [new to Borneo, Mt. Kinabalu; near Fr perminuta] 928 Stephani Icon Hepat Inedit [Diastaloba] B-56, microfiche 49 fig 3296 [from type; "Ins Philippinae, G Wallis 1870 in Physiotio repens"] = Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 perversa var. monoica S.Hatt. 1974 Basionym: Distribution: var nov - comb nov Hattori 1974 Hattori 1975 Mizutani, Hasegawa & Iwatsuki 2009 544 J Hatt Bot Lab 38: 177 [Luzon] 547 J Hatt Bot Lab 39: 292 [as Fr. junghuhniana var monoica] 2112 Type sepecimens Hatt Bot Lab (NICH) 32: Holotype. Philippines, Luzon, Prov. Mountain, Mt. Pulog. 2700-2900 m. On branches. 11 June 1965. Z. Iwatsuki and A. J. Sharp 244 (NICH 267450). = Frullania junghuhniana var. monoica (S.Hatt.) S.Hatt. 1975 Frullania junghuhniana var. minutissima fo. monoica (S.Hatt.) S.Hatt. 1976 Frullania junghuhniana var. tenella fo. monoica (S.Hatt.) S.Hatt. 1986 petalumensis Gottsche ex Bol. 1870 [nom nud] Basionym: Distribution: - syn nov Gottsche ex Bolander Gottsche ex Bolander Evans Frye & Clark = Frullania bolanderi 1870 1870 1897 1947 3472 1669 237 204 California Medical Gazette 2: 184 [nom nud] Cat Plants San Francisco: 40 [name only] Trans Connet Acad Arts Sci 10 (1): 8-10 Univ Wash Publ Biol 6 (5): 768 [syn of Fr bolanderi] Austin 1869 Page 334 pfefferi Steph. ex Herzog 1916 [nom nud] {Bolivia, Meteoriopsis?} Typus: Bolivia. Comarapa, 2600m 1911, T. Herzog 4207 in hb G Basionym: Distribution: Stephani Bonner Stephani 1916 1965 1985 222 655 928 Biblioth Bot 87: 247 [sine descript, nom nud] [Bolivia] Index Hepat 5: 399 Stephani Icon Hepat Inedit [Meteriopsis] E-31, microfiche 55 fig 3813 [Bolivia, Herzog 4207"] = philippiensis Steph. ex Paris 1910 [nom nud] Basionym: Distribution: Paris 1910 = Frullania philippinensis philippii 1455 Rev Bryol 37 (6): 130 [New Caledonia: St. Panié] Steph. 1911 Steph. ex Bonner 1965 [nom herb] Typus: Chile. Valdivia sd, Lechler 140 in hb G [sub Fr. chilensis] Basionym: Distribution: Chile Bonner Stephani = Frullania chilensis philippinensis 1965 1985 655 928 Index Hepat 5: 400 Stephani Icon Hepat Inedit [Galeiloba] C- 92a, microfiche 51 fig 3472 ["(olim Frull Philippii St ms)" syn of Fr chilensis] Steph. 1894 [auct. non Steph. 1911] S.Hatt. 1974 Basionym: Distribution: Merrill Hattori Hattori Hattori 1906 1974 1974 1982 1496 544 546 495 Philippine J Sci C, Bot 5: 311 [Philippines: Mt. Pulog] J Hatt Bot Lab 38: 180 [Luzon] J Hatt Bot Lab 38: 256-257 J Hatt Bot Lab 51: 252 = Frullania nobilis var. intermedia S.Hatt. 1973 Frullania pullei Verd. 1930 Frullania pauciramea phyllobola Steph. 1911 Glenny, Meagher & Brown 2003 [nom nud] Basionym: Distribution: Australia Carthy Glenny, Meagher & Brown 2003 2003 1125 Flora Australia Suppl Ser 21: 112 [nom nud] 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html = Page 335 picta [auct. non Steph. 1894] Vanden Berghen 1976 {from Mauritius} Typus: Mauritius. Curepipe leg. Vaughan in hb UPS Basionym: Distribution: Vanden Berghen 1976 Grolle Wigginton 1995 2002 = Frullania purpurea picta 185 Bull Jard Bot Nat Belg 46 (1/2): 80 [Mauritius specimen = Fr. purpurea] 250 Bryophyt Bibl 48: 147 [not Mauritius] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Steph. 1911 [auct. non Steph. 1894] Svihla 1959 {femal specimen from Singapore; Svihla 2914} Typus: Singapore. Botanical Garden, on Casuarina from Amboina 23.3.1958, Svihla 2914 Basionym: Distribution: Svihla 1959 394 Hattori & Thaithong 1978 70 = Frullania gracilis picta Bryologist 62: 193-195, fig 1-10 [male & female specimen from Singapore; Svihla2914= Fr gracilis] Bull Natn Sci Mus Tokyo, ser B (Bot), 4 (2): 66 [syn sub Fr. gracilis] (Reinw., Blume & Nees) Nees 1845 [auct. non Steph. 1894] Yoshinaga 1898 {from Japan} Typus: Japan. Prov. Tosa (Shikoku) -.3.1897, Yoshinaga 18 Basionym: Distribution: - syn nov Yoshinaga Iishiba Sawada 1898 1909 1909 Iishiba Hattori Hattori, Iwatsuki & Kanno Kamimura Hattori & Lin Yamada & Iwatsuki 1912 1952 1956 1961 1985 2006 = Frullania motoyana 887 890 930 Bot Mag Tokyo 12: 221 [Prov. Tosa (Shikoku) ] Bot Mag Tokyo 23: 363 [Japan] Bot Mag Tokyo 23 (no 268): 178-183, fig 30 [key to Japanese species, in Japanese] 888 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] 529 J Hatt Bot Lab 8: 31 [not probable in Japan] 862 J Hatt Bot Lab 16: 93 & 94 199 J Hatt Bot Lab 24: 75 [Yoshinaga 18 syn sub Fr. motoyana] 505 J Hatt Bot Lab 59: 159 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] Steph. 1911 Page 336 picta Steph. 1894 Typus: Malaysia. Amboina. Wawani Hia / Hila ? 1889, G. Karsten 230 in hb G [holotype] Basionym: Distribution: spec nov - syn nov Stephani Schiffner Verdoorn Verdoorn Reimers Verdoorn Verdoorn Herzog Svihla 1894 1898 1929 1930 1931 1932 1934 1949 1955 Svihla 1959 Kamimura Bonner Arnell Hattori & Lin 1961 1965 1965 1985 Stephani 1985 Hattori & Lin Grolle Wigginton 1985 1995 2004 Yamada & Iwatsuki Uribe-M. 2006 2011 = Frullania tenuicaulis Frullania vethii Hedwigia 33: 166 [sterile specimen; Amboina, leg Dr G Karstens] Conspect Hepat Archip Indici, Batavia: 337 Ann Bryol 2: 162 Ann Bryol Suppl 1: 122-123, fig 214-216 Hedwigia 71 (1/2): 38-39 [China] Bull Jard Bot Buitenzorg, ser 3, 12 (1): 61 [Sumatra, Java] Ann Bryol Suppl 4: 206-207 [close to Fr vethii] Trans Brit Bryol Soc 1 (3): 317 [Borneo, Sarawak] Bryologist 58: 72-76, fig 1-24 [male specimen; Burma, Pulo Penang, Amboina= Fr vethi; critically about Stephani ] 394 Bryologist 62: 193-195, fig 1-10 [male & female specimen from Singapore; Svihla2914= Fr gracilis] 199 J Hatt Bot Lab 24: 61 [syn sub Fr tenuicaulis] 655 Index Hepat 5: 400 : 74 505 J Hatt Bot Lab 59: 163 [Fr picta syn sub Fr trichodes, except female & Japanese specimen = Fr motoyana] 928 Stephani Icon Hepat Inedit [Diastaloba] B-77, microfiche 49 fig 33213322 [from type; "Amboina, Wawani Hia, D Karsten 1889" & "ex planta 168, O Kamiera1905"] 505 J Hatt Bot Lab 59: 163 [picta = Fr motoyana] 250 Bryophyt Bibl 48: 147 [not Mauritius] 1312 Trop Bryol Res Rep 5: 31 [Excluded from Madagascar] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] 2767 Caldasia 33(1): 74 [type study, syn sub Fr trichodes] Mitt. 1891 Sande Lac. 1884 Frullania trichodes picta 228 831 246 293 229 553 241 18 370 Mitt. 1862 [auct. non Steph. 1894] Svihla 1955 {male specimen from Burma; Svihla1414} Typus: Burma. Thawbawleik Tin Mines, 30 km NE of Tenasserim 15.3. 1953, Svihla 1414 Basionym: Distribution: auct - syn nov Svihla 1955 370 Kamimura 1961 199 Hattori & Thaithong 1978 70 = Frullania vethii Bryologist 58: 72-76, fig 1-24 [male specimen; Burma, Pulo Penang, Amboina; critically about Stephani ] J Hatt Bot Lab 24: 61-63 [may be a habitat modification of Fr. tenuicaulis] Bull Natn Sci Mus Tokyo, ser B (Bot), 4 (2): 69 [Svihla 1414 from Burma = poorly developed form of Fr. vethii] Sande Lac. 1884 Frullania trichodes Mitt. 1862 Page 337 picta Steph. 1911 {from Japan} Typus: Japan. Kyushu, Takeo -.4.1903, U. Faurie 1332 in hb HIRO (U. Faurie 1332) [isosyntypus?] Basionym: Distribution: spec nov - syn nov Stephani Kamimura Hattori & Lin 1911 1961 1985 NN 2003 = Frullania motoyana piliflora 686 199 505 Spec Hepat 4: 644-645 [pp. specimen from Japan, Prov Tosa]] J Hatt Bot Lab 24: 75 [F 1331 syn sub Fr. motoyana] J Hatt Bot Lab 59: 163 [Fr picta syn sub Fr trichodes, except female & Japanese specimen = Fr motoyana] 1397 Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] Steph. 1911 Steph. 1911 Typus: Costa Rica. Cerro de las Vueltas, 3000m 1911, A. Tonduz /? Sanduz in hb G [holotypus G sn] Basionym: Distribution: Costa Rica, Galapagos Isls [Ecuador] spec nov - syn nov Stephani Clark Herzog Bonner Stephani 1911 1953 1953 1965 1985 Morales Z. Kappelle Holz 1991 1996 2004 Uribe 2008 = Frullania peruviana piligera 686 359 640 655 928 Spec Hepat 4: 598-599 Bryologist 56: 286-289, fig 1-15 [similar to Fr.paradoxa] Rev Bryol Lichen 20: 129 [Costa Rica] Index Hepat 5: 401 Stephani Icon Hepat Inedit [Meteriopsis] E-33, microfiche 55 fig 3816 ["Costarica, Tonduz 1911, 3000m Höhe"] 307 Trop Bryol 4: 38 [Costa Rica] 1409 Los Bosques de Roble, Cordillera de Talamanca, Costa Rica: 257 1184 [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm 1513 Caldasia 30(1): 75 [syn sub Fr peruviana] [Monography of subgenus Meteoriopsis] Gottsche 1846 [non Gottsche ex Spruce 1884] (Nees) Austin 1879 Basionym: Jungermannia hutchinsiae var. (beta) Nees 1830 Distribution: Hawaii nom nov - syn nov Austin 1879 667 Stephani Verdoorn Kamimura Miller Bonner Miller, Whittier & Whittier 1911 1930 1961 1963 1965 1983 686 551 199 834 655 609 = Jubula hutchinsiae ssp. javanica pilistipula Bull Torrey Bot Club 6 [No 52]: 301 [nom nov pro Fr hutchinsiae var beta - Synop Hepat, p. 426] Spec Hepat 4: 690 [syn sub Fr. pennsylvanica] Nederl Kruidk Arch Jrg 1930 (2): J Hatt Bot Lab 24: 89 [syn sub Jubula hutchinsiae] Ark Bot, ser 2, 5: 520 [syn nov sub Jubula hutchinsiae ssp javanica] Index Hepat 5: 401 Prod Flora Hepat Polynesiae: 136 (Steph.) Verd. 1928 Steph. ex Theriot 1910 [nom nudum] Basionym: Distribution: nom nud Stephani ex Theriot = Frullania pilistipula 1910 1915 Bull Acad Int Geogr Bot 20: 104 [nom nud] Steph. 1911 Page 338 pinicola Steph. ex Bonner 1965 [nom herb] Typus: New Caledonia. Iles des Pins 1902, A. Le Rat 534 in hb G 18498 Basionym: Distribution: New Caledonia (Isle of Pines) - syn nov Bonner Hattori Miller, Whittier & Whittier = Frullania caledonica 655 492 609 Index Hepat 5: 401 J Hatt Bot Lab 49: 365-368 Prod Flora Hepat Polynesiae: 136 Gottsche ex Steph. 1894 [non (Schuster) Schuster 1992] Frullania squarrosula pinnata 1965 1981 1983 (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Feldberg, Schäf.-Verw. & Krings 2017 Typus: Basionym: Distribution: spec nov soec nov - syn nov Heinrichs & alii Heinrichs & alii Li, Li Feldberg et al = Frullania baerlocheri Schmidt 2011 pinnulata 2017 2017 2021 4615 Cretaceous Research (2017), doi: 10.1016/j.cretres.2017.05.030. 4626 Cretaceous Research 78: 56-60 5879 Cretaceous Research 124, 104803: 1-10 [syn sub Fr baerlocheri] Heinrichs, Reiner, Feldberg, von Konrat, Hentschel, Vána & Sande Lac. 1853 Basionym: Distribution: Indonesia (Java) - syn nov Sande Lacoste Sande Lacoste Sande Lacoste in Dozy Sande Lacoste Sande Lacoste in Junghuhn Schiffner Stephani Verdoorn Verdoorn Bonner Stephani = Frullania ramuligera planifolia 1855 1856 1856 1857 1857 1898 1911 1929 1930 1965 1985 2038 1894 564 430 1490 831 686 246 293 655 928 Nederl Kruidk Arch 3: 423 Natuurk Tijdschr v Nederl Indie 10: 398 Plagiochila sandei Dozy: 11 Verh Konikl Akad Wetenschappen 5: 86, pl 16(1-7) Plantae Junghuhnianae, Hepaticae: 539-540 Conspect Hepat Archip Indici, Batavia: 337 Spec Hepat 4: 656 Ann Bryol 2: 157 Ann Bryol Suppl 1: 130 Index Hepat 5: 402 Stephani Icon Hepat Inedit [Diastaloba] B-79, microfiche 49 fig 3324 ["Syn Hep Javan" & "Java"] - 3325 (Nees) Mont. 1843 Steph. ex Steph. 1985 [pro syn] Typus: Luzon Basionym: Distribution: Stephani = Frullania recurvata 1985 928 Stephani Icon Hepat Inedit [Thyopsiella] F-175c, microfiche 59 fig 4121 ["olim Frull planifolia St nsp, Luzon, Sarangani, No 61, Micholitz 1884/85"] Lehm. 1838 Frullania gaudichaudii (Nees & Mont.) Nees & Mont. 1845 Page 339 platyflora Pearson 1922 Typus: Zaire, Uele, Ituri, montagnes des environs de Nandia 4.10.1916, Lacomblez 65pp [en mélange avec Acrolejeunea orchidae Pears] in hb BR [holotypus] Basionym: Distribution: spec nov - syn nov Pearson Demaret Demaret & Vanden Berghen Bonner Vanden Berghen Wigginton 1922 1942 1947 1965 1976 2002 Wigginton Wigginton 2004 2004 = Frullania eplicata plumaeformis Steph. 1889 Steph. 1889 {Neu Guinea} Stephani Stephani Verdoorn Bonner Hattori Stephani 1889 1911 1930 1965 1980 1985 = Frullania billardieriana Typus: Mexico 941 686 549 655 490 928 Hedwigia 28: 278 Spec Hepat 4: 575 [syn sub Fr. billardieriana] Nova Guinea 14 (4): 541 Index Hepat 5: 403 J Hatt Bot Lab 47: 197-200 Stephani Icon Hepat Inedit [Homotropantha] D- 1, microfiche 54 fig 3703 ["olim Frullania plumaeformis (bicornustipula), Nova Guinea, Fly River Branch, Bäuerlen 1885, Herb Melbourne"] - 3704 [missing] Nees & Mont. in Mont. 1843 Frullania intermedia pluriplicata Natuurwet Tijdschr 4: 118-119, pl 1(1-16) [Begian Congo] Bull Jard Bot Etat Bruxelles 16 (2): 305 Bull Jard Bot Etat Bruxelles 18: 235-236 [Congo Belge] Index Hepat 5: 403 Bull Jard Bot Nat Belg 46 (1/2): 83 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 218 [syn sub Fr eplicata, West Africa] Steph. 1911 Basionym: Frullania bicornustipula Distribution: nom nov - syn nov 3476 1066 1101 655 185 1012 (Reinw., Blume & Nees) Nees 1845 Steph. ex Bonner 1965 [nom herb] {Mexico} sd, s coll ex hb NEES, ex hb JACK, in hb G; Basionym: Distribution: Mexico nom herb Bonner Stephani 1965 1985 655 928 Fulford & Sharp 1990 105 Index Hepat 5: 403 Stephani Icon Hepat Inedit [Chonanthelia] A- 46, microfiche 46 fig 3134 & microfiche 47 fig 3135 [from type; "Mexico (ex Herb Nees) Jack missit"] Mem NY Bot Gard 63: 45 [Mexico] = pocsiantha Thaithong & S.Hatt. by Mizutani, Hasegawa & Iwatsuki 2009 [orth var] Basionym: Distribution: orth var Mizutani, Hasegawa & Iwatsuki 2009 = Frullania pocsantha 2112 Type sepecimens Hatt Bot Lab (NICH) 33: [as pocsiantha] Holotype. Vietnam, Mt. Hoang Lien Son, Pr. opp. Sapa. 1927. A. Petelot 32 NICH (345476). Thaithong & S.Hatt. 1977 Page 340 poizana Gottsche ex Bonner 1965 [nom herb] Typus: Madeira. sd, Fritze 993 or Fritze 89 in hb G Basionym: Distribution: Madeira [Portugal] Bonner 1965 = Frullania trabutiana Index Hepat 5: 404 Steph. 1910 Frullania dilatata polilloensis 655 (L.) Dumort. 1835 Steph. 1911 Typus: Philippines. Island of Polillo -.10.1909 - -.11.1909, R.C. McGregor Bureau of Sci, No 10497, in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Hattori Stephani 1911 1928 1930 1965 1980 1985 Porto, Germano & Borges 2004 = Frullania intermedia Spec Hepat 4: 586 Rev Bryol NS 1: 120-121 Ann Bryol Suppl 1: 161-164 [syn sub Fr. intermedia] Index Hepat 5: 404 J Hatt Bot Lab 47: 187 [syn sub Fr. intermedia fo. fallacoides] Stephani Icon Hepat Inedit [Homotropantha] D-28, microfiche 54 fig 3746 ["PhilippinenPolillo Insel, leg Macgregor 10497"] - 3747 1446 Brejos de Altitude em Pernambuco e Paraiba: 83 [Brazil] fo. fallacoides Verd. 1930 Frullania intermedia polypetera 686 242 293 655 490 928 (Reinw., Blume & Nees) Nees 1845 [auct. non Taylor 1846] Yoshinaga by Choe 1980 [orth var] Basionym: Distribution: Choe = Frullania polyptera 1980 [auct. non Taylor 1846] Yoshinaga 1900 {specimen from Japan} Frullania diversitexta polyptera 1171 Illust Fl & Fauna Korea 24: 733 [South Korea] Steph. 1897 [auct. non Taylor] Verd. in Hand.-Mazz. 1930 Typus: China. Yunnan, 3000-3400m 25.5.1915, Handel-Mazzetti No 6528 in hb NICH, W, WU Basionym: Distribution: auct - comb nov Verdoorn in Handel-Mazzetti Hattori & Lin Hattori & Lin Bapna & Kachroo = Frullania zangii 1930 1985 1985 2000 550 505 505 712 Symb Sinicae 5: 38-39 [Handel-Mazzetti No 6528] J Hatt Bot Lab 59: 150-151 [as Fr. zangii] J Hatt Bot Lab 59: 142 [syn sub Fr. pallide-virens] Hepaticology in India II: 176 [syn sub Fr. pallide-virens] S.Hatt. & P.J.Lin 1985 Page 341 polyptera Typus: [auct. non Taylor 1846] S.Hatt. & P.J.Lin 1985 1906, Levier [det by Stephani] Basionym: Distribution: Hattori & Lin 1985 = Frullania muscicola polyptera 505 J Hatt Bot Lab 59: 141 Steph. 1894 [auct. non Taylor 1846] Yoshinaga 1900 {specimen from Japan} Basionym: Distribution: Yoshinaga Kamimura Choe 1900 1961 1980 = Frullania diversitexta ponapena 1685 Bot Mag Tokyo 14: 40 199 J Hatt Bot Lab 24: 81 [sub F diversitexta] 1171 Illust Fl & Fauna Korea 24: 733 [South Korea] Steph. 1897 Steph. 1924 Typus: New Guinea, Ponape 1913, Ledermann sn. hb G 3748, Basionym: Distribution: New Guinea, Micronesia [=Caroline Isl] (Ponape) spec nov - syn nov Stephani Verdoorn Bonner Hattori Miller, Whittier & Whittier Grolle & Piippo Stephani 1924 1930 1965 1980 1983 1984 1985 687 293 655 490 609 143 928 Spec Hepat 6: 553 Ann Bryol Suppl 1: 168-169 Index Hepat 5: 405 J Hatt Bot Lab 47: 183 Prod Flora Hepat Polynesiae: 136 Acta Bot Fennica 125: 22 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Homotropantha] D-29, microfiche 54 fig 3748 ["Ponape (C Ledermann leg)"] = Frullania utriculata var. powelliana (Steph.) Verd. 1930 Frullania utriculata poondeloyae Steph. 1894 Gottsche 1886 [nom nud] Typus: Sri Lanka: Poondeloya 4000-8000 ft 1868, John Nietner Drawings com. by Jack.: hb no 1547. Frullania Poondeloyae G. Ms. C. per. - 3 Bl. & hb no 1550. Frullania Poondeloyae simplicior. C. per. - 2 Bl. Basionym: Distribution: Gottsche Gottsche 0 1886 Icon inedt 2390 Bot. Centralblatt 26 (7): 204 [Sri lanka] = Page 342 poondeloyae ssp. simplicior Gottsche 1886 [nom nud] Typus: Sri Lanka: Poondeloya 4000-8000 ft 1868, John Nietner Drawings com. by Jack.: hb no 1547. Frullania Poondeloyae G. Ms. C. per. - 3 Bl. & no 1550. Frullania Poondeloyae simplicior. C. per. - 2 Bl. Basionym: Distribution: Gottsche 1886 2390 Bot. Centralblatt 26 (7): 204 [Sri lanka] = powelliana Steph. 1894 Typus: Samoa -.5.1885, T. Powell sn. ex hb K, hb STEPHANI 256, in hb G Basionym: Distribution: Vanuatu (Aneityum, Futuna), New Caledonia, Samoa spec nov - syn nov Stephani Stephani in Reinecke Paris Stephani Stephani & Watts Verdoorn Verdoorn 1894 1896 1910 1911 1914 1928 1930 228 1044 1455 686 220 242 293 Verdoorn Bonner Grolle & Schultze-Motel Hattori Miller, Whittier & Whittier Stephani 1930 1965 1972 1980 1983 1985 551 655 537 490 609 928 = Frullania utriculata primigenia Hedwigia 33: 151-152 Bot Jahrb 23: 315 [Samoa] Rev Bryol 37 (6): 130 [New Caledonia: Panié] Spec Hepat 4: 574 J Proc Roy Soc New S Wales 48: 109 [Aneityum, Futuna] Rev Bryol NS 1: 114-115 Ann Bryol Suppl 1: 169-170 [comb nov; as Fr. utriculata var. powellianan] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 405 J Hatt Bot Lab 36: 84 [Samoa] [sub Fr. utriculata var. powelliana] J Hatt Bot Lab 47: 183 [syn sub Fr. utriculata] Prod Flora Hepat Polynesiae: 136 Stephani Icon Hepat Inedit [Homotropantha] D- 9, microfiche 54 fig 3714 ["Samoa (Rev T Powell leg) 1885, Herb Kew"] - 3715 Steph. 1894 Caspary 1887 Typus: FOSSIL. Basionym: Distribution: Fossil (Eocene) in Baltic amber, Germany spec nov - syn nov Caspary Caspary & Klebs Grolle = Frullania varians proboscidiphora 1887 1907 1985 977 622 621 Schriften phys-ökonom Gesell Königsberg 27: 4, pl1 fig 14-15 Abh königl Preuß Geol Landesanstalt, NF 4: 35-37, Atlas fig 30(a-e) Prace Muz Ziemi z 37: 97 [syn sub Fr. varians] Caspary 1887 Taylor by Steph. 1910 [orth var] Basionym: Distribution: orth var Stephani Stephani 1910 1985 686 928 Stephani 1985 928 = Frullania probosciphora Spec Hepat 4: 356-357 Stephani Icon Hepat Inedit [Chonanthelia] A-106a, microfiche 48 fig 3221 ["olim Frull dentistipula Steph W.S.Wales Woronoka River (E.Theel.) Oct 1901 Sydney Bot Gard 92"] - 3222 Stephani Icon Hepat Inedit [Chonanthelia] A-106b, microfiche 48 fig 3221 - 3222 ["Icones Hookeri / Mitten Flora Tasmanica Hepat Hooker antarctica Voyage"] Taylor 1846 Page 343 proboscidophora keeled] [auct. non Taylor 1846] Mitt. in Hooker 1860 [perianth ventral one- Typus: Tasmania Basionym: Distribution: auct Mitten in Hooker 1860 - syn nov Gottsche in Mueller Bibby Scott & Bradshaw 1880 1955 1986 = Frullania clavata probosciphora 561 Antarctic voyage, III. Fl Tasmaniae 2: 236-237, pl m180(5) as "proboscidiphora" [Tasmania] 1489 Frag Phyt Australia 11, suppl Hepaticae :65 [Tasmania] 868 Victorian Naturalist 72: 102-103 [syn sub Fr. clavata] 56 Brunnonia 8 (1): 36 & 144 [Australia] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 [auct. non Taylor] E.A.Hodgs. {Australia} Basionym: Distribution: Hattori 1983 Glenny 1998 = Frullania reptans J Hatt Bot Lab 54: 139 [sub Fr reptans; Fr reptans = geographical race of Fr probosciphora] 1412 Tuhinga 10: 121-122 [revised checklist of New Zealand] Mitt. in Hooker 1855 Frullania probosciphora propagulifera 499 Taylor 1846 Schiffn. 1900 [nom nud] Typus: Indonesia. Sumatra, Kampong Sungei-Beramei -.3.1895, J. Massart Basionym: Distribution: nom nud Schiffner Verdoorn Verdoorn Bonner = Frullania campanulata pruniflora 1900 1929 1930 1965 904 246 293 655 Hedwigia 39: 208 Ann Bryol 2: 160 [syn with campanulata] Ann Bryol Suppl 1: 40 [nom nudum] Index Hepat 5: 405 Sande Lac. 1853 Steph. 1911 Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Bonner Grolle & Piippo Hattori & Streimann Stephani 1911 1930 1965 1984 1985 1985 686 293 655 143 504 928 Menzel 1988 512 Spec Hepat 4: 465 Ann Bryol Suppl 1: 32 [syn sub Fr. squarroas var. planescens] Index Hepat 5: 405 Acta Bot Fennica 125: 22 [Cat. West Melanesia] J Hatt Bot Lab 59: 121 [syn sub Fr. ericoides] Stephani Icon Hepat Inedit [Galeiloba] C-170, microfiche 52 fig 3571 ["Borneo & Nova Pomerina, leg Micholitz"] J Hatt Bot Lab 65: 164 [Cat Borneo] = Frullania squarrosa var. planescens Verd. 1929 Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides (Nees) Mont. 1839 Page 344 pseudopendula Steph. by S.Hatt. 1973 [orth var] Basionym: Distribution: Hattori 1973 = Frullania pendulostyla J Jap Bot 48 (4): 107 Steph. 1924 Frullania mirabilis ptychantha 124 J.B.Jack & Steph. 1892 [auct. non Mont. 1843] Baldwin 1877 Basionym: Distribution: Hawaii - syn nov Baldwin Miller Miller, Whittier & Whittier = Frullania hypoleuca 1877 1963 1983 901 834 609 Hawaiian Almanac: 42 [Hawaii] Ark Bot, ser 2, 5: 519 [syn nov sub Fr hypoleuca] Prod Flora Hepat Polynesiae: 136 Nees 1843 pubella var. elongata S.Hatt. by McCarthy 2006 [orth var] Typus: Basionym: Distribution: New Zealand (North Isl) = Frullania rubella var. elongata (Steph.) S.Hatt. 1983 pubella var. rubella Gottsche ex Steph. 1899 by McCarthy 2006 [orth var] Basionym: Distribution: McCarthy = Frullania rubella pulcherrima 2006 1385 Checklist Australian Liverworts [orth var for Fr rubella] Gottsche 1889 Steph. 1911 Typus: Surinam: Junhi, flumen Para 1885, Surinagar sn hb E. G. Paris 140, in hb G 13658 [holotypus] Basionym: Distribution: Guyana spec nov Stephani Bonner Stephani 1911 1965 1985 686 655 928 - syn nov Gradstein & Hekking Yuzawa & Koike 1989 1989 517 519 = Frullania apiculata Spec Hepat 4: 518-519 Index Hepat 5: 406 Stephani Icon Hepat Inedit [Thyopsiella] F- 84, microfiche 58 fig 3994 ["Suriname, leg Surinagar, Gén Paris 140"] J Hatt Bot Lab 66: 208 & 220 [syn nov syn sub Fr. apiculata] J Hatt Bot Lab 66: 346-347, fig. 3 [from type] [syn nov syn sub Fr. exilis TAYLOR 1846] (Reinw., Blume & Nees) Nees 1845 Page 345 pullei Verd. 1930 Typus: Indonesia. Irian Jaya, Jayawijaya, "Orangebegirgte", 3800m -.2.1913, Pulle 2526 in hb FH Basionym: Distribution: New Guinea (Papua New Guinea, Irian Jaya), Indonesia, Solomon Isl spec nov - syn nov Verdoorn Verdoorn Bonner Hattori Hattori Hattori Grolle & Piippo Hattori & Streimann Hattori & Piippo = Frullania pauciramea pulvinata 1930 1930 1965 1973 1973 1982 1984 1985 1986 293 549 655 541 540 495 143 504 557 Ann Bryol Suppl 1: 48, fig 35-36 Nova Guinea 14 (4): 542, pl 92(1-3) Index Hepat 5: 406 J Hatt Bot Lab 37: 116-120, (figs. 64-66). as "F. nobilis var. intermedia" J Hatt Bot Lab 37: 69-71, fig 31-32 as "F. phi!ippinensis" J Hatt Bot Lab 51: 253-255 [as syn sub Fr. pauciramea STEPH 1911 Acta Bot Fennica 125: 22 [Cat. West Melanesia] J Hatt Bot Lab 59: 116 [footnote synonym list] [New Guinea] Acta Bot Fennica 133: 53-54, fig 11(map) 8, key to pullei & nobilis] Steph. 1911 Colenso 1889 Typus: New Zealand, County of Waipawa, Dannevirke 1888, W. Colenso 1345 in hb G 20598 Basionym: Distribution: spec nov - syn nov Colenso Stephani Bonner Hamlin Stephani = Frullania squarrosula pungens 1889 1892 1965 1972 1985 636 210 655 659 928 Trans Proc New Zealand Inst 21: 77 J Linn Soc Bot 29: 274 [syn sub Fr. squarrosula] Index Hepat 5: 407 Rec Dominion Mus 7(19): 262 [New Zealand] Stephani Icon Hepat Inedit [Galeiloba] C-273b, microfiche 54 fig 3701 ["sub Frull pulvinata Col, Nova Zelandia, Colenso a 1345"] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 (Wilson) Steph. 1911 Basionym: Jungermannia pungens Wilson ex Hooker 1847 Distribution: Galapagos Isls [Ecuador] comb nov - syn nov Stephani Clark Bonner = Frullania aculeata purpurascens 1911 1956 1965 686 378 655 Spec Hepat 4: 549 Bryologist 59: 218 [syn sub Fr. aculeata] Index Hepat 5: 407 Taylor 1846 Steph. ex Beauverd in Steph. 1924 [pro syn] {Japan} [non Steph. 1911] Basionym: Distribution: Japan, Korea - nom nov Beauverd in Stephani Hattori Kamimura Choe Yuzawa = Frullania uematsuana 1924 1953 1961 1980 2001 687 530 199 1171 1660 Spec Hepat 6: 543 [as Fr uematsuana] J Hatt Bot Lab 10: 38 [syn sub Fr muscicola] J Hatt Bot Lab 24: 21 [syn sub Fr muscicola] Illust Fl & Fauna Korea 24: 732 [South Korea] Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] Beauverd 1924 Frullania truncatifolia Frullania muscicola Steph. 1910 Steph. 1894 Page 346 purpurascens Steph. 1911 Typus: Ecuador. Prov. Oriente, Gualaquiza -.5.1909, A. Allioni hb LEVIER 6463, in hb G Basionym: Distribution: Ecuador spec nov - syn nov - syn nov Stephani Stephani Bonner Stotler Stotler Stephani 1911 1916 1965 1969 1970 1985 Gradstein, Meneses & Arbe Yamada & Iwatsuki 2003 2006 = Frullania montagnei purpurascens 686 222 655 4761 194 928 Spec Hepat 4: 525-526 Biblioth Bot 87: 244 [Bolivia] Index Hepat 5: 407 Nova Hedwigia 18: 452-453 [syn sub Fr. montagnei] The genus Frullania, separata: 452-453 [syn sub Fr. montagnei] Stephani Icon Hepat Inedit [Thyopsiella] F-122, microfiche 58 fig 4049 [Ecuador, Allioni, Levier 6463 ex parte"] 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. montagnei] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] Gottsche 1845 [auct. non Steph. 1911] Wu in Li, C.Gao, Wu et alii 1985 {China} Basionym: Distribution: Wu in Li, Gao, Wu et al. = Frullania muscicola purpureopicta 1985 1169 Bryoflora Xizang (Tibet), Science Press, Beijing: 523 [sub Fr muscicola] Steph. 1894 Steph. 1910 Typus: Madagascar. Andranohinaly 1904, Voltzkow in hb G 13688 [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Demaret & Vanden Berghen Arnell Bonner Vanden Berghen Stephani 1910 1947 1963 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania socotrana pycnantha 686 1101 822 655 185 928 Spec Hepat 4: 367 Bull Jard Bot Etat Bruxelles 18: 236 [syn sub Fr socotrana] Hepat S Africa: 143 [syn sub Fr. socotrana] Index Hepat 5: 408 Bull Jard Bot Nat Belg 46 (1/2): 139 & 142 [ub Fr. socotrana] Stephani Icon Hepat Inedit [Galeiloba] C- 32, microfiche 50 fig 3384 [from type; "S W Madagascar, Völtzkow 1904"] 250 Bryophyt Bibl 48: 69 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 23 [syn sub Fr socotrana, West Africa] Mitt. 1888 [auct. non (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845] Steph. 1892 Basionym: Jungermannia pycnantha Distribution: Hodgson = Frullania banksiana Hooker & Taylor 1844 1972 3468 J Roy Soc New Zealand 2 (1): 110 [syn sub Fr. banksiana] Colenso 1889 Frullania pycnantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Page 347 pycnophylla S.Hatt. 1973 Typus: Indonesia. West Irian. Jayavijaya: Idenburg Riber, Bernhard Camp, 1600m 1939, Brass 12758 in hb JE [holotypus] in hb NICH [isotypus] Basionym: Distribution: spec nov - syn nov Hattori Hattori Grolle & Piippo Schuster 1973 1982 1984 1985 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania curvistipula pyncnantha 540 495 143 441 J Hatt Bot Lab 37: 60-61, fig 25 J Hatt Bot Lab 51: 215 Acta Bot Fennica 125: 23 [Cat. West Melanesia] Phytologia 57 (5): 370 [sect Frullania, ssect Pycnophylla - type species for nov ssect] 2112 Type sepecimens Hatt Bot Lab (NICH) 33: Isotype. New Guinea, Idembung River. 1600 m. On stem of tall palm in mossy forest. 1939. Brass 12758 (NICH 315689). Steph. 1911 Colenso by Steph. 1892 [orth var] Basionym: Distribution: Stephani Hamlin = Frullania pycnantha pyriflora 1892 1972 210 659 J Linn Soc Bot 29: 274 [orth error] Rec Dominion Mus 7(19): 262 [New Zealand] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Steph. 1910 Typus: India. Nilgiri Hills, Kotagiri, St. Katherines Fall, 6500ft 12.7.1907, Lüthy hb LEVIER 6128, in hb G 14555 Basionym: Distribution: India (North: Kashmir), Nepal spec nov - syn nov Stephani Kashyap 1910 1932 Kashyap Chopra Parihar Bonner Hattori 1932 1943 1962 1965 1974 Srivastava Stephani 1979 1985 Parihar, Lal & Katiyar Bapna & Kachroo Kattel 1994 2000 2002 = Frullania polyptera 686 Spec Hepat 4: 443 1529 Liverworts of the western Himalayas, Part 1, Suppl.: 10 [added to Kashmir Valley] 226 Liverworts of the western Himalayas, Part 2: 15-16, pl 2(1-3) 205 J Indian Bot Soc 22: 249 [India] 1584 Univ Allahabad studies, Bot sect 1-2: 19 [India: WH SI] 655 Index Hepat 5: 408 545 J Hatt Bot Lab 38: 208-211, fig 100 [from type] [syn nov sub Fr. polyptera] 24 Nova Hedwigia Beih 63: 335 [India, Kashmir] 928 Stephani Icon Hepat Inedit [Galeiloba] C-171, microfiche 52 fig 3572 [from type; "Nilgherry Mts (Lüthy) Levier 6128"] 2196 Hepat Anth India, annotated checklist: 41 [syn sub F rpolyptera] 712 Hepaticology in India II: 178 [syn sub Fr. polyptera] 1032 Liverworts of Nepal, Katmandu: 19 [Nepal, 2500 m] Taylor 1846 Page 348 quinqueplicata Steph. 1910 Typus: New Zealand. sd, Knight 600 in hb G Basionym: Distribution: spec nov - syn nov Stephani Hodgson Bonner Hamlin Yamada & Iwatsuki 1910 1949 1965 1972 2006 = Frullania sebastionopolitana Lindenb. 1845 Frullania rio-janeirensis Typus: Mexico Spec Hepat 4: 356 [as Chonanthelia] Trans Roy Soc New Zealand 77 (3): 376 Index Hepat 5: 409 Rec Dominion Mus 7(19): 262 [New Zealand] J Hatt Bot Lab 99: 29 [Catalog of Japan] Lindenb. by Buck 1994 [orth var] Frullania sebastianopolitana rabenhorstii 686 203 655 659 1470 (Raddi) Ångstr. 1876 Steph. 1911 sd, Leibold sn ex hb RABENHORST, in hb G (holotypus G 20621) Basionym: Distribution: Mexico, Guatemala spec nov - syn nov Stephani Bonner Stephani 1911 1965 1985 Fulford & Sharp Uribe 1990 2008 = Frullania grandifolia racemosa 686 655 928 Spec Hepat 4: 601 Index Hepat 5: 409 Stephani Icon Hepat Inedit [Meteriopsis] E-34, microfiche 55 fig 3817["Mexico (lg Leibold) (Herb Dr Winter)"] - 3818 105 Mem NY Bot Gard 63: 45 [Mexico] 1513 Caldasia 30(1): 74 [syn sub Fr. grandifolia] [Monography of subgenus Meteoriopsis] Steph. 1911 Lehm. & Lindenb. ex Nees 1845 [pro syn] Basionym: Distribution: Nees in G., L. & N. = Frullania tamarisci raddiana 1845 2850 Syn Hepat: 438 (L.) Dumort. 1835 Lindb. ex Holz 2004 [nom nud] Basionym: Distribution: Costa Rica Holz 2004 Dauphin 2005 1184 [checklist of Costa Rica] http://asbot.botanik.unigreifswald.de/personal/holz/bryo_cr/blank.htm 1360 Tropical Bryology 26: 141-218 [Catalogue of Costa Rican Hepaticae and Anthocerotae] = Page 349 rechingeri Steph. 1908 Typus: Samoa. Upolu Isl, Lanutov 1905, K. Rechinger 3057 in hb G 17501 [lectotypus] Basionym: Distribution: Vanuatu (Aneityum, Futuna), Samoa (Savaii, Upola), Fiji (Viti Levu) spec nov - syn nov Stephani Stephani Stephani & Watts Verdoorn Verdoorn Bonner Campbell Grolle & Schultze-Motel Hattori 1908 1911 1914 1930 1930 1965 1971 1972 1977 Hattori Miller, Whittier & Whittier Stephani 1981 1983 1985 Uribe 2008 = Frullania immersa rechingeri 216 686 220 293 551 655 674 537 479 Denkschr K Akad Wiss, Math Nat Kl, Wien 81: 298 Spec Hepat 4: 611 [spec nov = nom superfl] J Proc Roy Soc New S Wales 48: 105 [Futuna, Aneityum] Ann Bryol Suppl 1: 153 Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 410 J Roy Soc New Zealand 1 (1): 12 [Fiji] J Hatt Bot Lab 36: 84 [Samoa] J Hatt Bot Lab 43: 416, footnote 5 [syn sub Fr. immersa] [New Caledonia] 492 J Hatt Bot Lab 49: 373-375, fig 9(a-i) from type 609 Prod Flora Hepat Polynesiae: 136 928 Stephani Icon Hepat Inedit [Meteriopsis] E-62, microfiche 56 fig 3861 ["Samoa, Rechinger 3057"] 1513 Caldasia 30(1): 90, [belongs to subgen Fusiorielligerae] [Monography of subgenus Meteoriopsis] Steph. 1896 Steph. 1911 [nom superfl] Basionym: Distribution: Stephani = Frullania rechingeri Frullania immersa 1911 686 Spec Hepat 4: 611 Steph. 1908 Steph. 1896 Page 350 recurvata Lehm. 1838 Typus: India. near Madras sd, Wight sn hb HOOKER, in hb NY, typus in hb S (B24217-24219) in hb E E00293863 [isotypus] in hb GLM (hb LEHMANN 53) 05959 Basionym: Distribution: spec nov - syn nov Lehmann Lehmann & Lindenberg in G., L. & N. Sande Lacoste in Junghuhn Sande Lacoste Mitten Sande Lacoste Schiffner Stephani Herzog Verdoorn Verdoorn Verdoorn Chopra Parihar 1838 1845 588 Pugillus 7: 12-13 2850 Syn Hepat: 436-437 1857 1857 1861 1864 1898 1911 1927 1929 1930 1930 1943 1962 1490 430 574 842 831 686 174 245 549 293 205 1584 Hattori Hattori 1972 1975 Miller, Whittier & Whittier Stephani 1983 1985 Stephani 1985 Stephani 1985 Stephani 1985 Parihar, Lal & Katiyar Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 1994 2002 = Frullania gaudichaudii reductiloba Plantae Junghuhnianae, Hepaticae: 538 Verh Konikl Akad Wetenschappen 5: 84 J Linn Soc Bot 5: 121 [South India: Madras] Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 313 [Java] Conspect Hepat Archip Indici, Batavia: 337-338 Spec Hepat 4: 685 [sub sgen Frullania] Mit Inst Bot Hamburg 7(1): 196 [West-Borneo] Ann Bryol 2: 141 [Java] Nova Guinea 14 (4): 541 Ann Bryol Suppl 1: 91 [syn sub Fr. gaudichaudii] J Indian Bot Soc 22: 249 [India] Univ Allahabad studies, Bot sect 1-2: 18 [India; syn sub Fr gaudichaudii] 538 J Hatt Bot Lab 36: 124-126, fig 10 47 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 78-81, fig 1(e-f) from lectotype 609 Prod Flora Hepat Polynesiae: 136-137 928 Stephani Icon Hepat Inedit [Thyopsiella] F-175c, microfiche 59 fig 4121 ["olim Frull planifolia St nsp, Luzon, Sarangani, No 61, Micholitz 1884/85"] 928 Stephani Icon Hepat Inedit [Thyopsiella] F-175a, microfiche 59 fig 4119 ["Ins Sarangani - Philippinen, Micholitz 1884/85"] 928 Stephani Icon Hepat Inedit [Thyopsiella] F-175b, microfiche 59 fig 4120 ["olim Frull manii St nsp, Andaman, Leg Man 4/8. 1890, Levier 92"] 928 Stephani Icon Hepat Inedit [Thyopsiella] F-175d, microfiche 59 fig 4122 ["olim Frull multiflora nsp, Borneo, leg A Everett, No 37"] - 4123 ["Everett 37"] 2196 Hepat Anth India, annotated checklist: 41 [syn sub Fr gaudichaudii] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] (Nees & Mont.) Nees & Mont. 1845 Verd. 1930 Typus: Indonesia. Irian Jaya "Dutch New Guinea", Jayawijaya, Mt. Goliath 1950-3000m 1911, De Kock 7 pp in hb FH (hb VERDOORN 19807) Basionym: Distribution: spec nov - syn nov Verdoorn Verdoorn Bonner Hattori Grolle & Piippo 1930 1930 1965 1975 1984 293 549 655 66 143 Ann Bryol Suppl 1: 121-122, fig 217(:127) Nova Guinea 14 (4): 545, pl 93(24-25) Index Hepat 5: 410 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 147-148 Acta Bot Fennica 125: 23 [Cat. West Melanesia] = Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 Page 351 reflexa Typus: Brazil Ångstr. 1876 sd, S. Henschen typus in hb S (B24220) Basionym: Distribution: Brazil spec nov - syn nov Ångström Stephani Schiffner & Arnell Bonner Yano Yano Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Gradstein & Costa Yano = Frullania atrata 1876 1911 1964 1965 1984 1999 2002 579 686 198 655 501 1263 782 Öfversigt K Vetensk Akad Förh 33 (7): 89-90 [Brazil] Spec Hepat 4: 523 [= sgen Frullania] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 89-90 [Brazil] Index Hepat 5: 410 J Hatt Bot Lab 56: 506 [checklist Brazil] Briophyta do estado de São Paulo. Lista provisória [Brazil] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 2003 2008 984 Mem New York Bot Gard 87: 83 [Brazil] [syn sub Fr atrata] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr atrata; Brazil; catalog] (Sw.) Nees ex Mont. 1839 reflexistipula var. caledonica S.Hatt. by S.Hatt. 1982 [orth var] Basionym: Distribution: - syn nov Hattori Hattori 1977 1982 = Frullania reflexistipula rehmannii 479 495 J Hatt Bot Lab 43: 431-432, fig 14 [as Fr. novocaledonica] J Hatt Bot Lab 51: 256 [oth err & syn sub Fr. reflexistipula] Sande Lac. 1853 Steph. 1911 Typus: South Africa. Montague pass sd, A. Rehmann 152 hb STEPHANI & Icon ined 1727 in G, in hb {British Museum} Basionym: Distribution: Mozambique, Zambia, Zimbabwe, South Africa spec nov - syn nov Stephani Sim Arnell Arnell Bonner Vanden Berghen Stephani 1911 1926 1955 1963 1965 1976 1985 Best Wigginton 1990 2002 Manyanga & Perold Wigginton 2004 2004 686 252 1690 822 655 185 928 Spec Hepat 4: 485-486 (as "rehmanii") Trans Roy Soc S Africa 15: 46 [South Africa] - not known to SIM Bot Not 108: 309 [is different from Fr capensis] Hepat S Africa: 154-156, fig 114(a-h) Index Hepat 5: 411 Bull Jard Bot Nat Belg 46 (1/2): 42 [syn sub Fr. capensis G.] Stephani Icon Hepat Inedit [Thyopsiella] F- 24, microfiche 56 fig 3904 ["Africa australis, Montagupass, leg Rehmann"] 879 Kirkia 13 (2): 296 [Zimbabwe] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1301 SABONET Report 21: 4 [Zimbabwe] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] = Frullania capensis Gottsche 1845 reimersii var. meijeri [nom herb in hb L L0060936 & L0060937] Typus: Malaysia/Sabah. Borneo, Tambuyokan, Gunung "N.Borneo, West Coast Res.; Mt. Tambuyokon, c. 15 miles N.E. of Kinabalu Peak; Puntjak (=summit of) Tambuyokon, c. 15 miles N.E. of Kinabalu Peak" 5/7/1961, Meijer, W. B 11170 in hb L L0060936 & L0060937 [typus] - http://145.18.162.53:81/c8?ent=300017&rec=69096&sct=1 Basionym: Distribution: = Page 352 reimersii var. rostellula S.Hatt. 1974 Typus: Papua New Guinea. Wstern Highlands, Mt. Hagen 3300-3400m 1968, van Zanten 68-2987/a in hb JE [holotypus] in hb NICH [isotypus] Basionym: Distribution: var nov - comb nov Hattori Hattori Grolle & Piippo 1974 1982 1984 546 495 143 J Hatt Bot Lab 38: 258-261, fig 130-131 J Hatt Bot Lab 51: 220 Acta Bot Fennica 125: 23 [Cat. West Melanesia] = Frullania macgregorii var. rostellula (S.Hatt.) S.Hatt. 1982 reimersii var. wilhelmensis (S.Hatt.) S.Hatt. 1974 Basionym: Frullania verdoorniana var. wilhelmensis S.Hatt. 1973 Distribution: comb nov - syn nov Hattori Hattori 1974 1974 546 546 Hattori Hattori Grolle & Piippo 1975 1982 1984 547 495 143 J Hatt Bot Lab 38: 261 [comb nov] J Hatt Bot Lab 38: 243 [New Guinea] [may be conspecific with Fr. macgregorii J Hatt Bot Lab 39: 299-300 J Hatt Bot Lab 51: 219 [syn nov] Acta Bot Fennica 125: 23 [Cat. West Melanesia] 961 aus # 961 Bot Bull Acad Sinica 5(2): 193 = Frullania macgregorii reiparia Steph. 1894 by Conard 1956 [orth var] Basionym: Distribution: Conard Yang & Lee = Frullania riparia relicta 1956 1964 Hampe 1838 (Nees) Spruce 1884 [orth var] in Miller, Whittier & Whittier 1983 Basionym: Distribution: Miller, Whittier & Whittier = Frullania replicata 1983 609 Prod Flora Hepat Polynesiae: 137 Spruce ex Miller 1968 Frullania replicata (Nees) Mont. 1840 Frullania nodulosa (Reinw., Blume & Nees) Nees 1845 Page 353 relicta Steph. 1911 Typus: Indonesia. Java Hasskarl ex hb NEES, ex hb JACK, in hb G Basionym: Distribution: Indonesia (Java) spec nov - syn nov Stephani Verdoorn Verdoorn Clark Bonner Stephani 1911 1929 1930 1955 1965 1985 686 246 293 369 655 928 Spec Hepat 4: 543-544 Ann Bryol 2: 161 Ann Bryol Suppl 1: 97, 101 Bryologist 58: 68 [syn sub Fr. apiculata] Index Hepat 5: 412 Stephani Icon Hepat Inedit [Thyopsiella] F-152, microfiche 59 fig 4091 ["Java, (Hasskarl leg) Herb Nees 1854 (misit Jack)"] = Frullania apiculata var. goebelii Schiffn. 1893 remotifolia Steph. 1911 Typus: Ecuador. "inter Chunchi at jugam Azuay, 3000m 25.1.1909, Allioni sn ex hb LEVIER 6439, in hb G [holotypus G 20622] Basionym: Distribution: Ecuador spec nov Stephani Herzog Bonner Stephani 1911 1957 1965 1985 - syn nov Uribe 2008 = Frullania peruviana 686 447 655 928 Spec Hepat 4: 601 Svensk Bot Tidskr 51 (1): 190 [Ecuador] Index Hepat 5: 412 Stephani Icon Hepat Inedit [Meteriopsis] E-35, microfiche 55 fig 3819 ["Ecuador, Azuay, leg Allioni, Levier 6439 ex parte"] - 3820 1513 Caldasia 30(1): 75 [syn sub Fr peruviana] [Monography of subgenus Meteoriopsis] Gottsche 1846 [non Gottsche ex Spruce 1884] renauldii Steph., Spec Hepat 4: 627 (17 Jul 1911) 1911 [non Steph., Spec Hepat 4: 487 (28 Feb 1911)] Typus: Mauritius [Ins. Maurice] sd, Renauld 99 in hb G 10709 Basionym: Distribution: Mauritius spec nov - syn nov Stephani Stephani Bonner Vanden Berghen Stephani 1911 1924 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania insularis Frullania apicalis 686 909 655 185 928 Spec Hepat 4: 627 Spec Hepat 6: 570 [syn sub Fr. insularis] Index Hepat 5: 412 Bull Jard Bot Nat Belg 46 (1/2): 52 [syn sub Fr. apicalis] Stephani Icon Hepat Inedit [Diastaloba] B-14, microfiche 48 fig 32423243 [from type; Ins Maurice Renauld 99"] 250 Bryophyt Bibl 48: 62 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apicalis, West Africa] Beauverd in Steph. 1924 Mitt. 1879 Page 354 renauldii Steph., Spec Hepat 4: 487 (28 Feb 1911) 1911 [non Steph., Spec Hepat 4: 627 (17 Jul 1911)] Typus: Bourbon sd, Rodriguez hb RENAULD 128, in hb G 10710 [holotypus], in hb UPS [isotypus] Basionym: Distribution: Reunion Isl spec nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania variegata renistipula 686 655 185 928 Spec Hepat 4: 487 Index Hepat 5: 412 Bull Jard Bot Nat Belg 46 (1/2): 128 Stephani Icon Hepat Inedit [Thyopsiella] F- 25, microfiche 57 fig 3905 ["Bourbon, leg Rodriquez, Renauld 128"] 250 Bryophyt Bibl 48: 70 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 235 [syn sub Fr variegata, West Africa] Steph. 1894 Steph. 1910 {Madagascar} Typus: Madagaskar sd, Forsyth Major, 485pp in hb G 13067 [holotypus] in hb JE 04004147 (Herb. R. Grolle ex FI ex Herb. E. Levier (no. 1982) (1894, Forsyth-Major,C.I. 460 p.p., Madagascar, S.E.-Tanala, 1350-1440 m, Ambohimitombo Forest) [isotypus] Basionym: Distribution: Madagascar spec nov - syn nov Stephani Bonner Vanden Berghen Grolle Wigginton 1910 1965 1976 1995 2002 Wigginton 2004 = Frullania anderssonii renistipula 686 655 185 250 1012 Spec Hepat 4: 378 Index Hepat 5: 413 [Madagascar] Bull Jard Bot Nat Belg 46 (1/2): 172 [syn sub Fr. anderssonii] Bryophyt Bibl 48: 62 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Ångstr. 1873 Steph. in Levier 1901 [nom nud] {Madagascar} Typus: Madagascar sd, Forsyth Major 485 pp in hb G 13067 [holotypus], Basionym: Distribution: nom nud Stephani in Levier Stephani Vanden Berghen Stephani = Frullania renistipula 1901 1910 1976 1985 1456 686 185 928 Rev Bryol 28 (5): 94 [Madagascar] Spec Hepat 4: 378 Bull Jard Bot Nat Belg 46 (1/2): 172 [syn sub Fr. anderssonii] Stephani Icon Hepat Inedit [Galeiloba] C- 33, microfiche 50 fig 3385 [from type; "Madagascar, Forsyth Major 485 pp"] Steph. 1910 {Madagascar} Frullania anderssonii Ångstr. 1873 Page 355 renistipula Steph. 1924 [pro syn] {Neu Guinea} [non Steph. 1910, from Madagascar] Typus: Neu Guinea. sd, Ledermann Basionym: Distribution: - nom nov Schiffner Stephani Stephani 1893 1924 1985 572 687 928 Stephani 1985 928 = Frullania ultima Beauverd 1924 [non Steph. 1924] {Neu Guinea} Frullania heteromorpha fo. mamillosa Verd. 1930 Frullania heteromorpha repanda Nova Acta K Leop-Carol Deutsch Akad Naturf Halle 60 (2): 226 Spec Hepat 6: 554 Stephani Icon Hepat Inedit [Thyopsiella] F-153, microfiche 59 fig 4092 ["Nova Guinea (Ledermann legit"] Stephani Icon Hepat Inedit [Homotropantha] D-31, microfiche 54 fig 3751 ["Nova Guinea (Ledermann legit)"] Schiffn. 1889 Gottsche 1864 Typus: Colombia. New Granada, Tequendama, Fusagasuganis sd, Triana in hb GOTTSCHE 2 Basionym: Distribution: Colombia spec nov - syn nov Gottsche Jack & Stephani Stephani Bonner Gradstein & Hekking Stephani 1864 1892 1911 1965 1979 1985 Uribe & Gradstein Uribe 1998 2008 Hentschel et alii Bernal, Gradstein & Celis 2015 2016 = Frullania microauriculata 478 1089 686 655 483 928 Ann Sci Nat Bot, ser 5, 1: 175-176 Hedwigia 31: 13 [Peru, Neu-Granada] Spec Hepat 4: 504 Index Hepat 5: 413 J Hatt Bot Lab 45: 113 [Cat Colombia] Stephani Icon Hepat Inedit [Thyopsiella] F- 85, microfiche 58 fig 3995 ["N Granada, Wallis"] - 3996 707 Bryophyt Bibl 53: 37 [Catalogue Colombia] 1513 Caldasia 30(1): 91, [type was not available] [Monography of subgenus Meteoriopsis] 4097 Phytotaxa 220 (2): 136 [syn sub Fr.macrocephala] 4290 Catálogo de plantas y líquenes de Colombia. Vol. I. [dubious species] Verd. 1929 repandistipula var. dioica S.Hatt. 1974 Typus: Philippines. Luzon Basionym: Distribution: Philippines var nov - syn nov Hattori 1974 Hattori 1975 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania alte-mammillata Frullania armatifolia repandistipula ssp. spinifera 544 J Hatt Bot Lab 38: 176 547 J Hatt Bot Lab 39: 227-228 2112 Type sepecimens Hatt Bot Lab (NICH) 34: Holotype. Philippines, Luzon, Mt. Pulog. 2700-2900 m. On tree. I1June 1965. Z. Iwatsuki and A. J Sharp 272 (NICH 267469). S.Hatt. 1972 Verd. 1932 S.Hatt. by S.Hatt. 1976 [orth var] Basionym: Distribution: orth var Hattori 1976 80 J Hatt Bot Lab 40: 493 = Frullania repandistipula ssp. spinibractea S.Hatt. 1975 Page 356 repestris Steph. by Mizutani, Hasegawa & Iwatsuki 2009 [orth var] Basionym: Distribution: orth var Mizutani, Hasegawa & Iwatsuki 2009 = Frullania rupestris Steph. 1894 Frullania schimperi replicata Typus: 2112 Type sepecimens Hatt Bot Lab (NICH) 34: [as repestris] Isotype. Africa occid., Camerunia, Buea. 2425 m. 21 July 1891. P. Dusen (NICH 218484). Another isotype: NICH 218485. Nees 1845 (Nees) Mont. 1840 Lepr coll n. 267 & 279 Basionym: Jungermannia replicata Distribution: comb nov - syn nov Nees 1833 Montagne Gottsche, Lindenberg & Nees Nees in G., L. & N. Montagne Spruce Stephani Schiffner 1840 1843 1845 1855 1884 1888 1895 Schiffner 1898 Stephani Schiffner in Engler & Prantl Paris Stephani Verdoorn Verdoorn Miller Bonner Yano Stephani 1905 1905 1910 1911 1928 1930 1960 1965 1984 1985 Stotler in Seaward & FitzGerald 1996 Gradstein & Costa 2003 = Frullania nodulosa replicata 467 568 2850 475 243 938 1015 Ann Sci Nat Bot, ser 2, 14: 333 Verh K Leop-Carol Akad Naturforsch 19, suppl 1: 471-472 Syn Hepat: 434 [syn sub Fr. thuillerii] Ann Sci Nat Bot, ser 4, 3: 319 [syn sub Fr. thuillerii] Trans Proc Bot Soc Edinburgh 15: 36-37 (No 26) Hedwigia 27: 280 [Puerto Rico] Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 134 831 Conspect Hepat Archip Indici, Batavia: 339 [according Spruce identical with Fr. nodulosa] 934 Hedwigia 45: 229 809 Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 134 1455 Rev Bryol 37 (6): 130 [New Caledonia: Mé Areinbo] 686 Spec Hepat 4: 581 244 Ann Crypt exot 1 (2): 218-220 [syn sub Fr. nodulosa] 293 Ann Bryol Suppl 1: 177 920 Bryologist 63: 121 [Micronesia] 655 Index Hepat 5: 413 501 J Hatt Bot Lab 56: 506 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Homotropantha] D-32, microfiche 54 fig 3752 ["syn Frull Thuilleri Nees, leg Weigelt, Surinam"] - 3753 1092 Richard Spruce, Roy Bot Gard Kew: 131 [subgen Homotropantha] 984 Mem New York Bot Gard 87: 89 [Brazil] [as Fr replicata (Nees) Spruce, sub Fr nodulosa] (Reinw., Blume & Nees) Nees 1845 Spruce ex Miller 1968 Basionym: Distribution: Kamimura Miller Yano = Frullania replicata 1961 1968 2008 199 J Hatt Bot Lab 24: 57 672 Micronesia 4 (1): 74 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr nodulosa; Brazil; catalog] (Nees) Mont. 1840 Frullania nodulosa (Reinw., Blume & Nees) Nees 1845 Page 357 replicata (Nees) Steph. 1911 [nom superfl] Basionym: Jungermannia replicata Distribution: Nees 1833 Stephani = Frullania replicata 1911 686 Spec Hepat 4: 581 (Nees) Mont. 1840 Frullania nodulosa (Reinw., Blume & Nees) Nees 1845 replicata var. convoluta Gottsche, Lindenb. & Nees 1843 Typus: Philippines.Manila, Laguna da Bay ["In Manila insula ad Laguna da Bay, Octobri 1831, c. perianth"] -.10.1831 hb NEES Basionym: Distribution: var nov - syn nov Gottsche, Lindenberg & Nees G., L. & N. Bonner = Frullania secundiflora Frullania nodulosa reptans 1843 1845 1965 568 Verh K Leop-Carol Akad Naturforsch 19, suppl 1: 472 2850 Syn Hepat: 433 655 Index Hepat 5: 413 [may be nom herb] Mont. 1843 [non Taylor 1845] (Reinw., Blume & Nees) Nees 1845 Mitt. in Hooker 1855 Typus: New Zealand, N Island, east coast sd, Colenso sn in hb E E00002574 [isotypus] Basionym: Distribution: Tasmania, New Zealand spec nov - syn nov Mitten in Hooker 1855 560 Mitten in Hooker Hooker Gottsche in Mueller Bastow Stephani Rodway 1860 1867 1880 1888 1910 1916 561 1154 1489 845 686 914 Bonner Hamlin Stephani 1965 1972 1985 655 659 928 Scott & Bradshaw Windolf Glenny Beckmann 1986 1987 1998 2002 56 1025 1412 971 Carthy Glenny & Malcolm 2003 2005 1125 1410 McCarthy 2006 1385 = Frullania probosciphora Antarctic voyage, II. Fl Novae-Zelandiae 2: 161, pl 104(4) [New Zealand] Antarctic voyage, III. Fl Tasmaniae 2: 237 [Tasmania] Handbook New Zealand Flora: 536-537 [New Zealand, N Island] Frag Phyt Australia 11, suppl Hepaticae :65 [Tasmania] Papers Proceed Roy Soc Tasmania for 1887: 266, pl 29 [Tasmania] Spec Hepat 4: 405-406 Papers Proceed Roy Soc Tasmania for 1916: 137 [syn sub Fr. pentapleura] Index Hepat 5: 414 Rec Dominion Mus 7(19): 262 [New Zealand] Stephani Icon Hepat Inedit [Galeiloba] C-270, microfiche 54 fig 3695 ["New Zealand, legit Chesseman, Levier No 40"] Brunnonia 8 (1): 36 [Australia] Austrobaileya 2(4): 390 [syn sub Fr. probosciphora] Tuhinga 10: 132 [revised checklist of New Zealand] Australasian bryol Newsletter 45(May 2002): 9 [syn sub Fr. probosciphora] Flora Australia Suppl Ser 21: 45 [sub Fr probosciphora] ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand species] Checklist Australian Liverworts [Australia, syn sub Fr probosciphora] Taylor 1846 Page 358 reticulata (Hooker & Taylor) Hooker 1867 Basionym: Jungermannia reticulata Hooker & Taylor 1844 Distribution: Auckland Isl [New Zealand] comb nov Hooker Hooker Stephani Bonner Hamlin Grolle 1867 1867 1909 1965 1972 1976 = Lepidolaena reticulata rhystiantha 1154 1154 686 655 659 624 Handbook New Zealand Flora: 755 [remove to Polyotus] Handbook New Zealand Flora: 538-539 [Lord Auckland's group] Spec Hepat 4: 46 [syn sub Lepidolaeana] Index Hepat 5: 414 Rec Dominion Mus 7(19): 263 [New Zealand] J Hatt Bot Lab 30: 46 (Hooker & Taylor) TREVIS 1877 S.Hatt. by Mizutani, Hasegawa & Iwatsuki 2009 [orth var] Basionym: Distribution: orth var Mizutani, Hasegawa & Iwatsuki 2009 = Frullania rhytidantha 2112 Type sepecimens Hatt Bot Lab (NICH) 33: [as rhystiantha] Holotype. India, Assam, near Schillong. 5000 ft. On boulder. 8 Apr. 1965. A. J. & Evelyn Sharp 6131 b (NICH 257551). S.Hatt. 1980 rhystocolea var. parva S.Hatt. 1971 Typus: Bhutan. Bhotokha-Rinchu, 1450-1500m 8.5.1976, in hb NICH 287397 [holotypus] Basionym: Distribution: Bhutan var nov - syn nov Hattori in Hara 1971 Bapna & Kachroo 2000 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania rhystocolea rhytidontha 190 Univ Mus Univ Tokyo Bull 2: 237-238, fig 25 712 Hepaticology in India II: 180 [syn nov sub Fr. rhystocolea] 2112 Type sepecimens Hatt Bot Lab (NICH) 34: Holotype. Bhutan, Bhotokha-Rinchu. 1450-1500 m. 8 May 1967. 0. Tanaka (NICH 287397). Herzog 1930 S.Hatt. by Bapna & Kachroo 2000 [orth var] Basionym: Distribution: orth var Bapna & Kachroo = Frullania rhytidantha 2000 712 Hepaticology in India II: 181 243 898 142 Trans Proc Bot Soc Edinburgh 15: 52 Mem Torrey Bot Club 1: 119-120 Willdenowia 14: 510 [syn of Fr triquetra] S.Hatt. 1980 rigescens var. mollicula Spruce 1884 Typus: Peruvian Andes, Mt. Campana sd, Spruce Basionym: Distribution: Peru var nov stat nov Spruce Spruce Menzel = Frullania triquetra 1884 1890 1984 Lindenb. & Gottsche 1847 Page 359 rioblancana Steph. 1910 Typus: Bolivia, Rio Blanca coll. ign. in hb G 19734 (ex hb JACK) [holotypus] Basionym: Distribution: Bolivia spec nov Stephani Stephani Bonner Yuzawa Stephani 1910 1916 1965 1983 1985 686 222 655 498 928 - syn nov Yuzawa Gradstein, Meneses & Arbe 1991 2003 525 975 = Frullania decidua riobrancana Spec Hepat 4: 343 Biblioth Bot 87: 240 [Bolivia] Index Hepat 5: 415 J Hatt Bot Lab 54: 112-113, fig. 4 (a-m) [from type Stephani Icon Hepat Inedit [Chonanthelia] A- 78, microfiche 47 fig 3181 [from type; "Bolivia Rio Blanco (Herb Jack)"] J Hatt Bot Lab 70: 211 [syn sub Fr. decidua] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. decidua] Spruce 1884 Steph. by Yuzawa 1991 [orth var] Basionym: Distribution: Yuzawa 1991 = Frullania rioblancana Frullania decidua rio-janeirensis 525 J Hatt Bot Lab 70: 211-212 Steph. 1910 Spruce 1884 [auct. non (Raddi) Ångstr.] Frye & Clark 1947 Basionym: Distribution: - syn nov Frye & Clark Stotler & Crandall-Stotler = Frullania arietina 1947 1977 204 398 Univ Wash Publ Biol 6 (5): 778-780, fig. Bryologist 80: 420 Taylor 1845 Frullania rio-janeirensis (Raddi) Ångstr. 1876 riparia [auct. non Hampe ex Lehm. 1838] MUELLER in Rabenhorst 1915 {Europe: Austria, Switzerland} Basionym: Distribution: Austria, Switzerland Müller in Rabenhorst Dismier Müller in Rabenhorst Bisang, Schumacker, Sergio & Grolle Thouvenot = Frullania cesatiana Frullania riparia 1915 1921 1958 1989 1064 1338 1028 276 Krypt Flora (2ed) 6, 2: 629-631, fig 177(a-e) Rev Bryol 47: 24-26 [first record in France] Krypt Flora (3ed) 6, 2: 1252-1253, fig 488 Giorn Bot Ital 122: 263 2003 1218 Bull sci bot Centre-Ouest, ns 34: 330 [France] De Not. 1865 Hampe 1838 Page 360 riparia [auct. non Hampe in Lehm. 1838] Yoshinaga 1906 {Japan} Basionym: Distribution: auct - syn nov Yoshinaga Iishiba Kamimura Bapna & Kachroo Yuzawa Yamada & Iwatsuki = Frullania muscicola riparia 1906 1912 1961 2000 2001 2006 927 888 199 712 1660 1470 Bot Mag Tokyo 20: 55 [Japan] Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] J Hatt Bot Lab 24: 21 [syn nov sub Fr. muscicola] Hepaticology in India II: 175 [syn nov sub Fr. muscicola] Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] J Hatt Bot Lab 99: 32 [Catalog of Japan] Steph. 1894 [auct. non Hampe ex Lehm. 1838] S.W.Arnell 1961 {Cape Verde Isl} Basionym: Distribution: Canary Isl ( Te )[Spain], Cape Verde Isl Arnell Grolle Eggers Wigginton 1961 1970 1984 2002 Wigginton 2004 = Frullania ericoides riparia 54 615 22 1012 Bot Not 114 (2): 177 [from Cape Verde Isl] Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 315-316 Crypt Bryol Lichen 3 (4): 288 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] (Nees) Mont. 1839 [auct. non Hampe ex Lehm. 1838] Bapna & Kachroo 2000 {North India, Sikkim} Basionym: Distribution: Bapna & Kachroo 2000 712 Hepaticology in India II: 197 [distr: North India, Europe, Japan, N. America] = riparia [auct. non Hampe ex Lehm. 1838] S.W.Arnell 1961 {Canary Isl.} Basionym: Distribution: - syn nov Casas 0 Arnell 1961 Casas, Brugués, Cros & Sérgio 1989 = Frullania azorica 1989 180 Svensk Bot Tidskr 55 (2): 391 [Canary Isl] 2670 [belongs to Fr. azorica] Sim-Sim, Sérgio, Mues & Kraut 1995 Page 361 robillardii Steph. in Renauld & Cardot 1893 Typus: Mauritius [Maurice] sd, Robillard ex hb BESCHERELLE, in hb G 13672 [holotypus], in hb PC [isotypus] Basionym: Distribution: Mauritius spec nov - syn nov Stephani in Renauld & Cardot Stephani Stephani Bonner Vanden Berghen Stephani 1893 1894 1910 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania anderssonii robinsonii 935 228 686 655 185 928 Bull Soc Roy Bot Belg 32 (2,I): 32 [Reunion] Hedwigia 33: 146-147 [as spec nov] Spec Hepat 4: 377 Index Hepat 5: 416 Bull Jard Bot Nat Belg 46 (1/2): 172 Stephani Icon Hepat Inedit [Galeiloba] C- 35, microfiche 50 fig 3387 [from type; "Maurice, Robillard, Herb Bescherelle"] 250 Bryophyt Bibl 48: 61 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Ångstr. 1873 Steph. ex Verd. 1930 [pro syn] Typus: Philippines. Luzon, Ifugao, Mt. Polig -.2.1913, R.G. McGregor in hb G Basionym: Distribution: - syn nov Verdoorn Bonner Stephani 1930 1965 1985 = Frullania nepalensis 293 655 928 Ann Bryol Suppl 1: 51 [syn sub Fr. nepalensis] Index Hepat 5: 416 [nom herb?] Stephani Icon Hepat Inedit [Galeiloba] C-174, microfiche 52 fig 3575 [from type; "Philippinen Merril 19912"] (Spreng.) Lehm. & Lindenb. 1845 robusta Steph. 1911 [non Kiaer ex Renauld & Cardot 1891, nec Spruce [Hepat exsicc] ex Steph. 1911] Typus: Ecuador. Tunguvagua sd, Spruce in hb US Basionym: Distribution: Costa Rica, Ecuador - syn nov Stephani Herzog Clark 1911 1952 1953 686 444 352 Herzog Bonner NN 1953 1965 1996 640 655 632 = Frullania brasiliensis Spec Hepat 4: 505-506 Svensk Bot Tidskr 46 (1): 85 [Ecuador] Bryologist 56: 142 [Fr. robusta SPRUCE = not correct name, as Spruce Hep. exsiccatae were never published] Rev Bryol Lichen 20: 129 [Costa Rica] Index Hepat 5: 416 [cited as "robusta (SPRUCE) SPRUCE 1892"] gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 27 of 35 [in hb US] Raddi 1822 Page 362 robusta Kiaer ex Renauld & Cardot 1891 [pro syn] [non SPRUCE [Hepat exsicc] ex Steph. 1911] Typus: Madagascar. Mt. Ankaratra sd, M. Borgen 97H hb Renauld 170, in hb G sub Fr. loricata Basionym: Distribution: Madagascar - syn nov Stephani in Renauld & Cardot Bonner Stephani 1891 1965 1985 Grolle Wigginton 1995 2002 Wigginton 2004 = Frullania loricata robusta 1453 Rev Bryol 18 (4): 57 [Madagascar] 655 Index Hepat 5: 417 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 23c, microfiche 56 fig 3903 ["olim Fr robusta Kiaer, ex Herb Renauld; Madagascar, leg Borgen"] 250 Bryophyt Bibl 48: 66 [syn sub Fr. loricata] 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Pearson 1891 Spruce [Hepat exsicc] ex Steph. 1911 [non Kiaer ex Renauld & Cardo 1891] Basionym: Distribution: Stephani Clark 1911 1953 686 352 Bartram & Arnell Stephani 1961 1985 932 928 Stephani 1985 928 Stephani 1985 928 Spec Hepat 4: 505-506 Bryologist 56: 142 [Fr. robusta SPRUCE = not correct name, as Spruce Hep. exsiccatae were never published] Bryologist 64: 249 [Galapagos Isls: Santa Cruz, Pinzón] Stephani Icon Hepat Inedit [Thyopsiella] F- 87a, microfiche 58 fig 3999 ["olim Frull Spruceana Steph, olim Fr brasil beta achetra? Spr, Tunguragua Spruce"]- 4000 Stephani Icon Hepat Inedit [Thyopsiella] F- 67ab, microfiche 57 fig 3969 ["olim Frull robusta Spruce ms (Tunguragua) Andes"] - 3972 ["ex planta Wagneri, Panama"] Stephani Icon Hepat Inedit [Thyopsiella] F- 87b, microfiche 58 fig 4001 [""olim Fr Spruceana St, Peru, Agapata"] = Frullania robusta Steph. 1911 [non Kiaer ex Renauld & Cardot 1891, nec Spruce [Hepat exsicc] ex Steph. 1911] Frullania brasiliensis robusta Raddi 1822 fo. spinuliflora Herzog 1951 Basionym: Distribution: Costa Rica Herzog Bonner = Frullania spruceana 1951 1965 640 655 Rev Bryol Lichen 20: 129 Index Hepat 5: 417 Steph. 1894 Frullania robusta Steph. 1911 [non Kiaer ex Renauld & Cardot 1891, nec Spruce [Hepat exsicc] ex Steph. 1911] Frullania brasiliensis Raddi 1822 Page 363 rodriguezii Steph. 1894 Typus: Reunion. Ins. Bourbon 1889, Rodriguez sn ex hb RENAULD, in hb G 10711 Basionym: Distribution: Reunion Isl spec nov - syn nov Stephani Stephani Bonner Vanden Berghen Stephani 1894 1911 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania apicalis rogosa 228 686 655 185 928 Hedwigia 33: 166-167 Spec Hepat 4: 625 Index Hepat 5: 416 Bull Jard Bot Nat Belg 46 (1/2): 52 Stephani Icon Hepat Inedit [Diastaloba] B-15, microfiche 48 fig 3244 [from type; "Bourbon lg Rodriquez"] 250 Bryophyt Bibl 48: 62 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apicalis, West Africa] Mitt. 1879 Mitt. by Parihar, Lal & Katiyar 1994 [orth var] Basionym: Distribution: orth var Parihar, Lal & Katiyar = Frullania rugosa 1994 Mitt. 1861 Frullania grandistipula rostrata 2196 Hepat Anth India, annotated checklist: 22 [Eastern Himalaya] Lindenb. 1845 [non Steph. 1924] [auct. non (H. & T.) Hooker & Taylor 1845] Ångstr. 1872 {Tierra del Fuego} Basionym: Distribution: Ångström 1872 577 Massalongo Engel 1885 1978 875 182 = Frullania magellanica rostrata_plicata Öfversigt K Vetensk Akad Förh 29 (4): 12 ["Porte Famine vid Magelhaens sund"] Nuovo Giorn Bot Ital 17: 264 [Tierra del Fuego] Fieldiana Bot 41: 243 F.Weber & Nees 1845 von Konrat & Braggins in von Konrat & Pickering 2003 [nom nud] Basionym: Distribution: Konrat & Pickering 2003 1000 http://www.discoverlife.org/nh/tx/Plantae/Bryophyta/ [IDnature guide for Liverworts, Frullania, New Zealand] = Frullania pubella var. rubella Gottsche ex Steph. 1899 by McCarthy 2006 [orth var] Frullania rubella Gottsche 1889 Page 364 rotundifolia Colenso 1887 Typus: New Zealand. County os Waipawa, Norsewood 1886, W. Colenso 1379 in hb G 20599 Basionym: Distribution: spec nov - syn nov Colenso Stephani Bonner Hamlin Stephani 1887 1892 1965 1972 1985 = Frullania squarrosula 634 210 655 659 928 Trans Proc New Zealand Inst 19: 297-298 J Linn Soc Bot 29: 274 [syn sub Fr. squarrosula] Index Hepat 5: 418 Rec Dominion Mus 7(19): 263 [New Zealand] Stephani Icon Hepat Inedit [Galeiloba] C-273a, microfiche 54 fig 3699 ["(Frull rotundifolia Colenso) N Zelandia, Colenso 1379"] - 3700 (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 rotundifolia Steph. ex Steph. 1985 [nom herb] {Meteoriopsis, Colombia} Typus: Colombia leg Stübel Basionym: Distribution: Colombia Stephani 1985 928 Stephani Icon Hepat Inedit [Meteriopsis] E-36, microfiche 55 fig 3821 ["Columbia (Stübel legit)"] = rotundiloba Steph. 1911 Typus: "India orient, in cortice" in hb G G00069116 - n° SIB 125408/1 - 007962 (ex hb M) > see fr0465.jpg Basionym: Distribution: Himalayas spec nov - syn nov Stephani Verdoorn Chopra Frye & Clark Kamimura Parihar Bonner Hattori 1911 1930 1943 1947 1961 1962 1965 1982 686 293 205 204 199 1584 655 0 Stephani 1985 928 Bapna & Kachroo 2000 712 = Frullania squarrosa (Reinw., Blume & Nees) Nees 1845 Frullania ericoides rotundistipula Spec Hepat 4: 679 Ann Bryol Suppl 1: 34 [syn sub Fr. squarrosa] J Indian Bot Soc 22: 249 [India] Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] J Hatt Bot Lab 24: 17 [syn sub Fr squarrosa] Univ Allahabad studies, Bot sect 1-2: 19 [India, syn sub Fr squarrosa] Index Hepat 5: 418 note in hb G: "A form of Fr ericoides: leaf-lobes are not strongly squarrose and -lobules with incurved beaks" oct 1982 Stephani Icon Hepat Inedit [Galeiloba] C-175, microfiche 52 fig 3576 ["Sikkim Himalaya, Herbarium München"] Hepaticology in India II: 170 [syn sub Fr. ericoides] (Nees) Mont. 1839 [auct. non Steph. 1894] Verd. in Hand.-Mazz. 1930 Basionym: Distribution: Verdoorn in Handel-Mazzetti Hattori & Lin = Frullania davurica 1930 1985 550 505 Symb Sinicae 5: 39 J Hatt Bot Lab 59: 133 fo. microphylla (C.Massal.) S.Hatt. & P.J.Lin 1985 Page 365 rotundistipula Steph. 1894 Typus: China. Yuennan, Hokin 4.8.1885, Delavay 1644 in hb PC; ex hb BESCHERELLE in hbh G Basionym: Distribution: China (Yunnan, Szechwan), India (Eastern Himalayas: Megalaya) spec nov - syn nov Stephani Stephani Stephani Verdoorn in Handel-Mazzetti Reimers 1894 1894 1910 1930 1931 Verdoorn Bonner Hattori Hattori Stephani 1932 1965 1973 1976 1985 Piippo Singh 1990 2002 Singh & Nath Singh, Kumar & Nath 2004 2008 = Frullania davurica rubrimacula 211 228 686 550 229 Mem Soc Nat Sci, Nat Math Cherbourg 29: 213 [China; spec nov] Hedwigia 33: 147 [spec nov] Spec Hepat 4: 439-440 Symb Sinicae 5: 39 [China] Hedwigia 71 (1/2): 34-35 [may be identical with Fr jackii and Fr davurica] 554 Nederl Kruidk Arch Jrg 1932 (2): 491 [China] 655 Index Hepat 5: 418 541 J Hatt Bot Lab 37: 91-92, fig 47-48 [from holotype] 67 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21 928 Stephani Icon Hepat Inedit [Galeiloba] C-176, microfiche 52 fig 35773578 ["China, Hokin, Yunnan, Delavay 1644 .... ] 521 J Hatt Bot Lab 68: 46 [annotated catalogue of China] 4886 Studies on Liverworts (Bryophyta) of Khasi and Jaintia Hills: Meghalaya. 264-266, pl 53 (1-12), map 11 - Unpublished thesis. 1589 Geophytology 32 (1&2): 73-77 [India, Meghalaya] 1281 Taiwania 53 (1): 68-70, fig 1[map], 7(1-12) [India, Meghalaya, Eastern Himalayas] Hampe ex Gottsche, Lindenb. & Nees 1845 Goebel by Uribe-M. 2011 [orth var] Basionym: Distribution: Uribe-M. = Frullania rubromaculata Frullania tenuicaulis Frullania vethii 2011 Goebel 1928 Mitt. 1891 Sande Lac. 1884 Frullania trichodes rubrimaculata 2767 Caldasia 33(1): 76 [as "rubrimacula", type study, syn sub Fr trichodes] Mitt. 1862 Goebel by Verd. 1930 [orth var] Basionym: Distribution: Verdoorn Horikawa Kamimura = Frullania rubromaculata Frullania tenuicaulis Frullania vethii 1930 1934 1961 293 34 199 Ann Bryol Suppl 1: 151 [syn sub Fr tenuicaulis] J Sci Hiroshima Univ, ser B, div 2, 2: 236 : 61 [syn sub Fr tenuicaulis] Goebel 1928 Mitt. 1891 Sande Lac. 1884 Frullania trichodes Mitt. 1862 Page 366 rubromaculata Goebel 1928 Typus: Indonesia. Sumatra, Fort de Kock 1925, Goebel sn Basionym: Distribution: spec nov - syn nov Goebel Verdoorn 1928 1929 Verdoorn 1929 Verdoorn Horikawa Kamimura Bonner Uribe-M. 1930 1934 1961 1965 2011 = Frullania tenuicaulis Ann Jard Bot Buitenzorg 39: 51-53, pl 9(fig 90-95) Ann Crypt exot 2 (1): 74 ["Über Frullania rubromaculata werde ich an anderer Stelle berichten (de Frullaniaceis VI).] 548 Ann Jard Bot Buitenzorg 40: 139 [GOEBEL (1928: 53) lenkte neuerdings noch die Aufmerksamkeit auf diese Vitta durch Aufstellung einer neuen Art, Frullania rubromaculata, welche jedoch mit Frullania tenuicaulis MITT. Identisch ist. 293 Ann Bryol Suppl 1: 151 [as "rubrimaculata, syn sub Fr tenuicaulis] 34 J Sci Hiroshima Univ, ser B, div 2, 2: 236 ["rubrimaculata"] 199 J Hatt Bot Lab 24: 61 [as "rubimaculata", syn sub Fr tenuicaulis] 655 Index Hepat 5: 418 "rubromaculata" 2767 Caldasia 33(1): 76 [as "rubrimacula", type study, syn sub Fr trichodes] Mitt. 1891 Frullania vethii Sande Lac. 1884 Frullania trichodes rufa 234 247 Mitt. 1862 Steph. 1911 Typus: Venezuela sd, Fendler 136 in hb G Basionym: Distribution: Costa Rica, Venezuela spec nov - syn nov - syn nov Stephani Herzog Bonner Stotler Stotler Stephani 1911 1953 1965 1969 1970 1985 = Frullania brasiliensis rufescens 686 640 655 4761 194 928 Spec Hepat 4: 505-506 Rev Bryol Lichen 20: 129 [Costa Rica] Index Hepat 5: 419 Nova Hedwigia 18: 436 The genus Frullania, separata: 436 Stephani Icon Hepat Inedit [Thyopsiella] F- 88, microfiche 58 fig 4002 ["Venezuela, Fendler"] - 4003 686 198 4761 194 501 928 Spec Hepat 4: 534-535 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 89 [Brazil] Nova Hedwigia 18: 463-464 [syn sub Fr. turbata] The genus Frullania, separata: 463-464 [syn sub Fr. turbata] J Hatt Bot Lab 56: 537 [syn sub Fr. flexicaulis] Stephani Icon Hepat Inedit [Thyopsiella] F-123, microfiche 58 fig 4050 ["Brasilia, Blumenau, Herb Dr Winkelmann"] Raddi 1822 Steph. 1911 Typus: Brasilia. Blumenau sd, Winkelmann 9 in hb G Basionym: Distribution: spec nov - syn nov - syn nov Stephani Schiffner & Arnell Stotler Stotler Yano Stephani = Frullania turbata Frullania atrata 1911 1964 1969 1970 1984 1985 Steph. 1911 (Sw.) Nees ex Mont. 1839 Page 367 rugosa Mitt. 1861 Typus: Sri Lanka. Matale -.12.1846, Nr. 1441[lectotype by Hattori 1975] in hb NY [4 syntypus] Basionym: Distribution: India, Sri Lanka,Indonesia (Java) spec nov - syn nov Mitten 1861 Sande Lacoste 1864 Schiffner Schiffner Stephani Verdoorn Chopra Abeywickrama Parihar 1898 1900 1911 1929 1943 1959 1962 Hattori 1975 Abeywickrama & Jansen Miller, Whittier & Whittier Stephani 1978 1983 1985 Parihar, Lal & Katiyar 1994 = Frullania grandistipula rupestris 574 J Linn Soc Bot 5: 120 [India: Khasi mount,, Sri Lanka: Matale, CochinChina] 842 Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 312 [Java, Sumatra, Halmaheira] 831 Conspect Hepat Archip Indici, Batavia: 341 904 Hedwigia 39: 208 [Java] 686 Spec Hepat 4: 453 245 Ann Bryol 2: 125 [specimens from Java = Fr. grandistipula] 205 J Indian Bot Soc 22: 249 [India] 200 Ceylon J Sci (Bio Sci) 2 (1): 58 [Sri Lanka] 1584 Univ Allahabad studies, Bot sect 1-2: 18 [India; syn sub Fr grandistipula] 47 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 78-81, fig 2(a-f) from lectotype [about mitten´s specimen] 794 MaB Nat Comm Sri Lanka, Publ 1: 5 [Sri Lanka] 609 Prod Flora Hepat Polynesiae: 137 928 Stephani Icon Hepat Inedit [Galeiloba] C-177, microfiche 52 fig 3579 ["Ceylon, Nietner"] 2196 Hepat Anth India, annotated checklist: 22 [Eastern Himalaya] Lindenb. 1845 [non Steph. 1924] Steph. 1894 Typus: Cameroon. Buea, 2426m -.7.1891, P. Dusen 278 or 284 [holotype] - 21.7.1891, Dusen sn [syntype] in hb L [holotypus] in hb BISH 456523 [syntypus] Basionym: Distribution: Cameroon spec nov - syn nov Stephani Stephani Bonner Vanden Berghen Stephani 1894 1911 1965 1976 1985 228 686 655 185 928 O'Connor, Gowing, Medler et alii 1998 Wigginton 2002 1274 1012 Wigginton 2004 Wigginton 2004 Mizutani, Hasegawa & Iwatsuki 2009 1312 1172 2112 = Frullania schimperi Hedwigia 33: 159 Spec Hepat 4: 486 Index Hepat 5: 419 Bull Jard Bot Nat Belg 46 (1/2): 59 Stephani Icon Hepat Inedit [Thyopsiella] F- 26, microfiche 57 fig 3906 ["Kamerun, Dusen 278.284"] - 3907 Type specimen catalog hb BISH Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 221 [syn sub Fr schimperi, West Africa] Type sepecimens Hatt Bot Lab (NICH) 34: [as repestris] Isotype. Africa occid., Camerunia, Buea. 2425 m. 21 July 1891. P. Dusen (NICH 218484). Another isotype: NICH 218485. Nees 1845 Page 368 rutenbergii Gottsche 1882 Typus: Madagascar. "inter alias Hepaticas in silva Ambaranavaranututa" 6.12.1877, Rutenberg 1502 in hb G Basionym: Distribution: Madagascar spec nov Gottsche Stephani in Renauld & Cardot Pearson - syn nov 254 Abh Naturwiss Ver Bremen 7 (3): 363 [Madagascar] 1453 Rev Bryol 18 (4): 56 [Madagascar] 1921 Christiania Videnskabs-Selskabs Forhandlinger 1890 (2): 6 [Madagascar] Stephani in Levier Stephani Bonner Vanden Berghen Stephani 1901 1911 1965 1976 1985 1456 686 655 185 928 Grolle Wigginton 1995 2002 250 1012 Wigginton Wigginton 2004 2004 1312 1172 = Frullania apicalis sabanetica 1882 1891 1891 Icon Zeitschrift Forhandlinger i Videnskaps-Selskabet i Kristiania 1890 (2): 4 Rev Bryol 28 (5): 94 [Madagascar] Spec Hepat 4: 479-480 Index Hepat 5: 420 Bull Jard Bot Nat Belg 46 (1/2): 52 Stephani Icon Hepat Inedit [Thyopsiella] F- 12, microfiche 56 fig 3886 ["leg Rutenberg 1503, Madagascar, 37"] Bryophyt Bibl 48: 62 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: 31 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 217 [syn sub Fr apicalis, West Africa] Mitt. 1879 Gottsche 1864 Typus: Colombia. Prov. Bogota, Sabaneta sd, Lindig 1725 Basionym: Distribution: Colombia, Ecuador spec nov -var nov Gottsche Spruce Stephani Herzog Herzog Bonner Gradstein & Hekking Menzel 1864 1884 1911 1952 1957 1965 1979 1984 Stephani 1985 Uribe & Gradstein Uribe & Rangel-Ch Rangel-Ch. Gradstein 1998 2000 2000 2020 478 243 686 444 447 655 483 142 Ann Sci Nat Bot, ser 5, 1: 170-171 Trans Proc Bot Soc Edinburgh 15: 45 & 53 Spec Hepat 4: 506 Svensk Bot Tidskr 46 (1): 85 [Ecuador] Svensk Bot Tidskr 51 (1): 190 [Ecuador] Index Hepat 5: 420 J Hatt Bot Lab 45: 113 [Cat Colombia] Willdenowia 14: 506 [not in Peru, but in Colombia - error by Stephani 1991: 506] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 89, microfiche 58 fig 4004 [Nova Granada, Lindig legit"] - 4005 707 Bryophyt Bibl 53: 37 [Catalogue Colombia] 1090 Colombia, diversidad biótica III, Bogota: 445 [Colombia] 1091 Colombia, diversidad biótica III, Bogota: 800 [Vulnerable] as Fr. intumescens var. sabanetica = Frullania intumescens var. sabanetica (Gottsche) Gradstein 2021 Frullania intumescens var. closterantha (Spruce) Gradstein & Pócs 2021 sabatenica Gottsche ex Myrdal 2000 [orth var] [from textfile, Mossor artlist] Basionym: Distribution: Bernal, Gradstein & Celis = Frullania sabanetica 1756 4290 Catálogo de plantas y líquenes de Colombia. Vol. I. [dubious species] Gottsche 1864 Frullania intumescens var. sabanetica (Gottsche) Gradstein 2021 Frullania intumescens var. closterantha (Spruce) Gradstein & Pócs 2021 Page 369 sabatenica var. parviloba Herzog ex Myrdal 2000 [orth var] [from textfile, Mossor artlist] Basionym: Distribution: Myrdal 2000 1265 http://www2.nrmse/kbo/saml/mossor/mossnamn.txt [hb S] = Frullania sabanetica var. parviloba Herzog ex Myrdal 2000 [from textfile, Mossor artlist] saccata Verd. 1934 [nom nud] Basionym: Distribution: nom nud - syn nov Verdoorn Verdoorn Grolle & Piippo Hattori & Piippo 1934 1937 1984 1986 = Frullania sublignosa sachapatensis 241 556 143 557 Ann Bryol Suppl 4: 220, fig 28 [map], [no description] Blumea suppl 1: 212 [nom nud; syn of Fr sublignosa] Acta Bot Fennica 125: 23 [Cat. West Melanesia] Acta Bot Fennica 133: 42 Steph. 1894 Steph. 1911 Typus: Peru. Sachapata sd, Lechler hb L 194, in hb G sn Basionym: Distribution: Peru spec nov - syn nov Stephani Bonner Hattori Menzel Stephani 1911 1965 1973 1984 1985 Uribe 2008 = Frullania mirabilis 686 655 Spec Hepat 4: 598 Index Hepat 5: 420 vide #1513 142 Willdenowia 14: 490 [Peru] 928 Stephani Icon Hepat Inedit [Meteriopsis] E-37, microfiche 55 fig 3822 ["Peru, Sachapata, leg Lechler"] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] J.B.Jack & Steph. 1892 sackawana var. minor Kamim. 1982 Typus: Japan. Kochi pref., Tosayamada-cho, Amatsubo, ca. 300m 3.11.1970, M. Kamim. 701131, 701132, 701133 [holotype], 701134 in hb NICH (Kamim. 701133) [holotypus], in hb Kamim. [isotypus] Basionym: Distribution: Japan - syn nov Kamimura Furuki & Mizutani Yuzawa 1982 1994 2000 Yamada & Iwatsuki 2006 = Frullania sackawana 420 Misc Bryol Lichen 9 (4): 91-93, fig 4 1331 Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] 1659 Natural Environmental Sci Res 13: 4, fig 3(a-j) [Japan, syn nov sub Fr sackawana] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] Steph. 1897 Page 370 sagittistipula Lindenb. & Gottsche 1847 Typus: Mexico. Mirador, Vercruz sd, Sartorius ex hb STR, in hb G Basionym: Distribution: Mexico, Costa Rica, Guatemala, Bolivia spec nov - syn nov - syn nov Lindenberg & Gottsche in G., L. & N. Gottsche Gottsche Stephani Bonner Stotler Stotler Fulford & Sharp Gradstein, Meneses & Arbe = Frullania breuteliana 1847 2848 Syn Hepat: 784 1863 1867 1911 1965 1969 1970 1990 2003 103 104 686 655 4761 194 105 975 Separata of Mex. Leverm.: 252 K Danske Vid Selks Skrifter, ser 5, 6: 348-349 [Mexico] Spec Hepat 4: 520 Index Hepat 5: 420 Nova Hedwigia 18: 453-454 [syn sub Fr breuteliana] The genus Frullania, separata: 453-454 [syn sub Fr breuteliana] Mem NY Bot Gard 63: 46 [Mexico] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr breuteliana] Gottsche 1845 sagittistipula var. (beta) laxior Lindenb. & Gottsche 1847 Typus: Mexico. Veracruz, ad Colipa or Misantla, 500-800ft Liebmann Basionym: Distribution: Mexico Lindenberg & Gottsche in G., L. & N. Gottsche Gottsche Bonner Stotler Stotler Stephani 1847 2848 Syn Hepat: 784 ["only "simplicior"] 1863 1867 1965 1969 1970 1985 103 104 655 4761 194 928 Fulford & Sharp 1990 105 = Frullania breuteliana sandei Separata of Mex. Leverm.: 252 K Danske Vid Selks Skrifter, ser 5, 6: 349 [Mexico] Index Hepat 5: 420-421 Nova Hedwigia 18: 453-454 The genus Frullania, separata: 453-454 Stephani Icon Hepat Inedit [Thyopsiella] F- 90, microfiche 58 fig 4006 ["Mexico, Liebmann No 138 (vel 38)"] - 4007 Mem NY Bot Gard 63: 46 [Mexico] Gottsche 1845 Kiaer in Steph. 1894 Typus: Magadascar. Betsileo 31.1.1879, M. Borgen 96H ex hb RENAULD, hb STEPHANI 173, in hb G 13674 [holotypus] & Basionym: Distribution: Madagascar spec nov - syn nov Stephani in Renauld & Cardot Kiaer in Stephani Stephani Bonner Vanden Berghen Stephani 1891 1894 1910 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton 2004 Wigginton 2004 = Frullania caffraria 1453 228 686 655 185 928 Rev Bryol 18 (4): 57 [Madagascar] Hedwigia 33: 147 Spec Hepat 4: 379-380 Index Hepat 5: 421 Bull Jard Bot Nat Belg 46 (1/2): 185 Stephani Icon Hepat Inedit [Galeiloba] C- 36, microfiche 50 fig 3388 [from type; "Madagascar, Borgen 96H"] 250 Bryophyt Bibl 48: 64 1012 Trop Bryol Res Rep 2: 29 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 31 [synonymy is doubtfull] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 229 [syn sub Fr caffraria, West Africa] Steph. 1894 Page 371 sanguifolia Steph. ex Steph. 1985 [nom herb] {New Guinea} Typus: New Guinea. leg Ledermann in hb G Basionym: Distribution: Stephani 1985 928 Stephani Icon Hepat Inedit [Galeiloba] C-179, microfiche 52 fig 3581 [""Nova Guinea, Ledermann legit"] = sanguinea Steph. 1911 Typus: China. Yunnan, fromtier of Burma -.11.1898, Micholitz in hb G 7963 Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Bonner Stephani 1911 1930 1965 1985 686 293 655 928 Piippo Bapna & Kachroo 1990 2000 521 712 = Frullania nepalensis sanukiensis Spec Hepat 4: 453 Ann Bryol Suppl 1: 51 [syn sub Fr. nepalensis] Index Hepat 5: 421 Stephani Icon Hepat Inedit [Galeiloba] C-180, microfiche 52 fig 3582 ["Birma - Yünnan Grenze, leg Micholitz Nov 1898"] J Hatt Bot Lab 68: 46 [annotated catalogue of China] Hepaticology in India II: 176 [sub Fr. nepalensis] (Spreng.) Lehm. & Lindenb. 1845 Steph. ex Bonner 1965 [nom herb] {Japan} Typus: Japan. Kotohira, Sanuki -.1.1911, S. Okamura 403 in hb G Basionym: Distribution: Japan Bonner Stephani 1965 1985 655 928 Index Hepat 5: 421-422 Stephani Icon Hepat Inedit [Galeiloba] C-250, microfiche 53 fig 3670 ["Kotohira, Sanuki, O Kamura legit No 403"] 1924 1930 1934 1961 1965 1985 687 550 34 199 655 928 Spec Hepat 6: 543 Symb Sinicae 5: 35 [syn sub Fr. japonica] J Sci Hiroshima Univ, ser B, div 2, 2: 237 [syn sub Fr. japonica] J Hatt Bot Lab 24: 52 [syn sub Fr jackii ssp japonica] Index Hepat 5: 422 Stephani Icon Hepat Inedit [Galeiloba] C-251, microfiche 53 fig 3671 ["Japonia, Marioka, leg K Sawada 1906, No 32"] = sawadae Steph. 1924 Typus: Japan. Marioka 1906, K. Sawada 32 in hb G Basionym: Distribution: Japan spec nov Stephani Verdoorn in Handel-Mazzetti Horikawa Kamimura Bonner Stephani = Frullania japonica Sande Lac. 1864 Frullania jackii var. japonica (Sande Lac.) S.Hatt. 1957 Frullania jackii ssp. japonica Frullania davurica (Sande Lac.) S.Hatt. 1959 Hampe ex Gottsche, Lindenb. & Nees 1845 Page 372 saxatilis Lindenb. 1845 Typus: USA. Massachusetts sd, Asa Gray hb LINDENBERG 9 in hb GLM 05960 Basionym: Distribution: spec nov Lindenberg in G., L. & N. Sullivant 1845 1849 - syn nov Evans Stephani Frye & Clark Bonner 1897 1910 1947 1965 = Frullania eboracensis scabriseta 2850 Syn Hepat: 424 248 Mem Amer Acad Arts Sci, ser 2, 4 (1): 175 [may be conspecific with Fr. eboracensis] 237 Trans Connet Acad Arts Sci 10 (1): 18-19 [syn sub Fr. eboracensis] 686 Spec Hepat 4: 431 [syn sub Fr. eboracensis] 204 Univ Wash Publ Biol 6 (5): 771 [sub Fr eboracensis] 655 Index Hepat 5: 422 Lehm. 1844 Colenso 1887 Typus: New Zealand. County of Waipawa, Norsewood. 1886, Colenso 1382 ex hb K, in hb G [sub Fr. deplanata] Basionym: Distribution: spec nov - syn nov Colenso Stephani Bonner Hamlin Stephani = Frullania deplanata scandens 1887 1892 1965 1972 1985 634 210 655 659 928 Trans Proc New Zealand Inst 19: 298-299 J Linn Soc Bot 29: 273 [syn sub Fr. deplanata] Index Hepat 5: 422 Rec Dominion Mus 7(19): 263 [New Zealand] Stephani Icon Hepat Inedit [Galeiloba] C-261ab, microfiche 54 fig 3682 ["olim Frull knightiana, N Zealand, Knight 601"] - 3683 ["Frull falcata Stz, Frull knightiana St, Frull scabriseta Col, Trans NZ Inst vol 19, Hepa 1382, ob scandens?"] Mitt. 1855 [auct. non Mont.] S.Hatt. 1978 Basionym: Distribution: Hattori Hattori Tan & Engel = Frullania deplanata schensiana 1978 1983 1986 416 Misc Bryol Lichen 8 (3): 46 [Auckland Isls] 499 J Hatt Bot Lab 54: 140 4286 J. Hatt. Bot. Lab.. No. 60: 301 [Philippines] Mitt. 1855 [auct. non C.Massal. 1897] S.Hatt. 1974 Typus: Philippines. Luzon sd, Is 413, Is 442a, Is 13927a Basionym: Distribution: - syn nov Hattori Hattori = Frullania microauriculata 1974 1975 544 547 J Hatt Bot Lab 38: 180 [Philippines; Luzon, Is 413, Is 442a, Is 13927a J Hatt Bot Lab 39: 293 Verd. 1929 Page 373 schensiana var. formosana S.Hatt. ex Inoue 1961 [nom nud] Typus: Taiwan. Ikenohata, Prov Karenko 1922, K.S. May Basionym: Distribution: nom nud Inoue Lai & Wang-Yang Kuo & Chiang Piippo Bapna & Kachroo 1961 1976 1988 1990 2000 = Frullania schensiana 964 33 948 521 712 J Jap Bot 36(5): 187 [Formosa] Taiwania 21 (2): 168 [Index to Taiwan] Taiwania 33: 13 [Index of Taiwan] J Hatt Bot Lab 68: 47 [annotated catalogue of China] Hepaticology in India II: 181 C.Massal. 1897 schensiana ssp. ontakensis Basionym: Frullania ontakensis Distribution: (Steph.) S.Hatt. 1975 [nom inval] Steph. 1910 Hattori 1975 Hattori & Lin 1985 Bapna & Kachroo Yuzawa Yamada & Iwatsuki 2000 2001 2006 = Frullania schensiana 66 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 163 [comb nov; ontakensis not synonym with schensiana, better treated as ssp.] 505 J Hatt Bot Lab 59: 143 [ontakensis is syn sub fr. schensiana; therefor "nom invalid" for ssp. ontakensis] 712 Hepaticology in India II: 181 [syn sub Fr. schensiana] 1660 Natural Environmental Sci Res 14: 35 [Japan, syn sub Fr schensiana] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] C.Massal. 1897 schensiana ssp. ontakensis var. punctata (S.Hatt.) S.Hatt. 1975 Basionym: Frullania ontakensis Distribution: comb nov Hattori Yamada & Iwatsuki = Frullania schensiana scutula fo. punctata S.Hatt. 1944 1975 2006 66 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 163 [comb nov] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] C.Massal. 1897 Steph. ex Bonner 1965 [nom herb] {Brasil} Typus: Brazil. Caldas sd, Mosen Im pp in hb G Basionym: Distribution: Brazil nom herb Bonner Stephani 1965 1985 655 928 Index Hepat 5: 423 Stephani Icon Hepat Inedit [Galeiloba] C- 72, microfiche 51 fig 3434 [from type; "Brasilia, Mosén legit"] 1957 1981 1506 Berichte Deutsch Bot Ges 70: 434. pl 12, fig 2-4 1058 Occas Pap Farlow Herb 16: 101-110, fig 1-5 = scyphoides Mägdefrau 1957 Basionym: Distribution: Mägdefrau Grolle = Frullania schumannii (Caspary) Grolle 1981 Page 374 sebastianopolitana Lindenb. 1845 Typus: in hb GLM 05961 (hb Lehmann) Basionym: Jungermannia obscura var. (alpha) Nees in Martius 1833 Distribution: Mexico, Peru, Brazil spec nov - syn nov Lindenberg in G., L. & N. Gottsche Sande Lacoste Gottsche Sande Lacoste Gottsche Stephani Verdoorn Frye & Clark Svihla Bonner Yano Menzel Stephani 1845 1863 1864 1864 1867 1867 1910 1929 1947 1956 1965 1984 1984 1985 2850 103 842 478 430 104 686 246 204 380 655 501 142 928 Fulford & Sharp Buck 1990 1994 105 28 = Frullania rio-janeirensis Syn Hepat: 412-413 (No 4) Separata of Mex. Leverm.: 234 Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 311 [Java, Sumatra] Ann Sci Nat Bot, ser 5, 1: 166 (No 2) Verh Konikl Akad Wetenschappen 5: 77 K Danske Vid Selks Skrifter, ser 5, 6: 330-331 [Mexico] Spec Hepat 4: 329 [syn sub Fr. riojaneirensis] Ann Bryol 2: 156 [syn sub Fr. galeata] Univ Wash Publ Biol 6 (5): 778 [sub Fr riojaneirensis] Bryologist 59: 278-279 [sub Fr. riojaneirensis] Index Hepat 5: J Hatt Bot Lab 56: 506 [checklist Brazil] Willdenowia 14: 510 [syn of Fr. riojaneirensis] Stephani Icon Hepat Inedit [Chonanthelia] A- 48, microfiche 47 fig 3138 ["Fr. riojaneirensis olim Fr. sebastianopolitana ... Pabst"] Mem NY Bot Gard 63: 46 [Mexico] Bryologist 97 (4): 422 [Vainio's specimen from Brazil at TUR] (Raddi) Ångstr. 1876 sebastianopolitana var. (c) tumidula Lindenb. 1847 Typus: Colombia: Prov. Merida sd, Moritz 156 in hb REG BEROL Basionym: Distribution: Lindenberg in G., L. & N. = Frullania rio-janeirensis 1847 2848 Syn Hepat: 771 [ad 4, (c)] (Raddi) Ångstr. 1876 sebastianopolitana var. (delta) Gottsche 1864 Basionym: Distribution: Mexico var nov - syn nov Gottsche Spruce Gradstein & Hekking = Frullania rio-janeirensis 1864 1884 1979 478 243 483 Ann Sci Nat Bot, ser 5, 1: 166 Trans Proc Bot Soc Edinburgh 15: 23 [syn sub Fr. riojaneirensis] J Hatt Bot Lab 45: 113 [Cat Colombia; Lindig specimens need further study] (Raddi) Ångstr. 1876 sebastianopolitana var. (ß) galeata (R., B. & N.) Lindenb. 1845 Basionym: Jungermannia galeata Distribution: Indonesia (Java) - syn nov Reinwardt, Blume & Nees 1824 Lindenberg in G., L. & N. Sande Lacoste Verdoorn Verdoorn = Frullania galeata 1845 1857 1929 1930 2850 430 246 293 Syn Hepat: 413 (No 4 ß) Verh Konikl Akad Wetenschappen 5: 77 Ann Bryol 2: 156 [syn sub Fr. galeata] Ann Bryol Suppl 1: 68 (Reinw., Blume & Nees) Verd. 1929 Frullania rio-janeirensis (Raddi) Ångstr. 1876 Page 375 sebastionopolitana Lindenb. by Buck 1994 [orth var] Basionym: Distribution: Buck 1994 = Frullania sebastianopolitana Bryologist 97 (4): 422 [Vainio's specimen from Brazil at TUR] Lindenb. 1845 Frullania rio-janeirensis secundiflora 28 (Raddi) Ångstr. 1876 Taylor ex Gottsche in G., L. & N 1845 [nom herb] Typus: Guyana "British Guiana". Demerara sd, Campbell Basionym: Distribution: Gottsche in G., L. & N. = Frullania taylori 1845 2850 Syn Hepat: 436 Gottsche 1845 Frullania gaudichaudii (Nees & Mont.) Nees & Mont. 1845 secundiflora Mont. 1843 [non Taylor 1845] Typus: Philippines sd, Cuming typus in hb S (B24243, 24244) Basionym: Distribution: spec nov Montagne Montagne Montagne Mitten Mitten Schiffner Schiffner & Stephani in Schumann & Lauterbach Stephani Merrill Verdoorn Verdoorn Kamimura Miller 1843 1845 1856 1885 1885 1898 1901 470 1596 1912 1744 1745 831 1493 1911 1914 1928 1930 1961 1968 686 1484 242 293 199 672 Choe Hattori Miller, Whittier & Whittier Grolle & Piippo Stephani 1980 1980 1983 1984 1985 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Feuerer 2002 = Frullania nodulosa 2008 Ann Sci Nat Bot, ser 2, 19: 258-259 London J Bot 4: 7 Sylloge generum specierumque cryptogamaram: 85 Rep Challenger Exp Botany 1 (3): 214 [South-eastern Moluccas] Rep Challenger Exp Botany 1 (3): 263 [Admirality Islands] Conspect Hepat Archip Indici, Batavia: 341 Flora Deutsch Schutzgebiete: 79 Spec Hepat 4: 580 Philippine J Sci C, Bot 9(1): 40 [Guam] Rev Bryol NS 1: 116-117, fig 9a Ann Bryol Suppl 1: 183, fig 304 J Hatt Bot Lab 24: 57 Micronesia 4 (1): 74 [syn sub Fr. nodulosa; key to Guam & Northern Micronesia] 1171 Illust Fl & Fauna Korea 24: 732 [syn of Fr brotheri; South Korea] 490 J Hatt Bot Lab 47: 209 609 Prod Flora Hepat Polynesiae: 137 143 Acta Bot Fennica 125: 23 [Cat. West Melanesia] 928 Stephani Icon Hepat Inedit [Homotropantha] D-34ab, microfiche 54 fig 3756 ["Andaman, leg Man, Levier 82"] - 3757 ["Manila, Meyen"] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm [Andaman Isl.] (Reinw., Blume & Nees) Nees 1845 Page 376 sehnemii Luisier & Herzog in Herzog 1936 Typus: Brazil. Sao Leopoldo, "auf alten Baumstämmen" sd, L. Sehnem ? typus in hb S (B24245) in hb JE Basionym: Distribution: Brazil, Argentina spec nov - syn nov Luisier & Herzog in Herzog Herzog 1937 1952 Bonner Yano Reiner Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot Gradstein & Costa Yano 1965 1984 1988 2002 Broteria, Série trimestral: Ciências Naturais 6: 18-20, fig 2 Feddes Repert 55: 10 [comb nov as Fr glomerata var sehnemii; Argentina] 655 Index Hepat 5: 424 501 J Hatt Bot Lab 56: 506 [checklist Brazil] 1043 Bol Soc Argentina Bot 25 (3-4): 318 [syn sub Fr. ericoides] 782 http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] 2003 2008 984 Mem New York Bot Gard 87: 86 [Brazil] [sub Fr ericoides] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr ericoides; Brazil; catalog] = Frullania ericoides sellowii 55 803 (Nees) Mont. 1839 Hampe ex Steph. 1985 [pro syn] Typus: Brasilia sd, Sellow Basionym: Distribution: Stephani = Frullania conferta 1985 Stephani Icon Hepat Inedit [Chonanthelia] A- 65ab, microfiche 47 fig 3163 ["olim Fr Sellowii Hampe ms Brasilia Sellow"] Spruce 1889 Frullania tetraptera semiaperta 928 Nees & Mont. 1838 Gottsche ex Clark 1954 [nom herb] Basionym: Distribution: Clark = Frullania neurota 1954 362 Bryologist 57: 30 Taylor 1846 Page 377 semiconnata Lindenb. & Gottsche 1847 Typus: Mexico: Chinantla Distr, Oaxaca sd, Liebmann sn in hb W [ hb LINDENBERG 7255] [holotypus], in hb G [isotypus]; Basionym: Distribution: Mexico, Colombia, Argentina spec nov - syn nov Lindenberg & Gottsche in G., L. 1847 & N. Gottsche 1863 Gottsche 1867 2848 Syn Hepat: 776 (No 38 b) Spruce Stephani Kühnemann Schiffner & Arnell 1884 1910 1949 1964 243 686 657 198 Gottsche Bonner Haarbrink Stephani 1964 1965 1981 1985 478 655 49 928 Fulford & Sharp Gradstein, Meneses & Arbe 1990 2003 105 975 = Frullania tetraptera 103 104 Separata of Mex. Leverm.: 242 K Danske Vid Selks Skrifter, ser 5, 6: 338-339 [as alpha: foliis imbricatis][Mexico] Trans Proc Bot Soc Edinburgh 15: 14 & 27-28 (No 16) Spec Hepat 4: 342 [syn sub Fr. tetraptera] Lilloa 19: 355 [Argentina: Tucuman, Salta] Österr Akad Wiss, Math Nat Kl, Denkschr 111: 85, pl. 13 (fig. 146) [syn with Fr. tetraptera] [Brazil] Ann Sci Nat Bot, ser 5, 1: 168-169 (No 8) Index Hepat 5: 425 Lindbergia 7 (1): 49 [Colombia, High Andes] Stephani Icon Hepat Inedit [Chonanthelia] A- 80ab, microfiche 47 fig 3183 ["Merida Funck & Sohlim legit"] - 3184 ["Icones Gottschei!"] Mem NY Bot Gard 63: 46 [Mexico] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. tetraptera; acording Yuzawa 1991 ?] Nees & Mont. 1838 semiconnata var. ( ) granatensis Gottsche 1864 Typus: Mt. Morro or Cipaquira sd, Lindig Basionym: Distribution: var nov Gottsche Bonner Gradstein & Hekking = Frullania semiconnata Frullania tetraptera 1864 1965 1979 478 655 483 Ann Sci Nat Bot, ser 5, 1: 169-170 (No 8-) Index Hepat 5: 425 J Hatt Bot Lab 45: 113 [Cat Colombia] Lindenb. & Gottsche 1847 Nees & Mont. 1838 semiconnata var. (beta) laxior Gottsche 1863 Basionym: Distribution: Gottsche Gottsche = Frullania semiconnata Frullania tetraptera 1863 1867 103 104 Separata of Mex. Leverm.: 242 K Danske Vid Selks Skrifter, ser 5, 6: 338 [Mexico] Lindenb. & Gottsche 1847 Nees & Mont. 1838 semiconnata var. (gamma) tenella Gottsche 1863 Basionym: Distribution: Gottsche Gottsche = Frullania semiconnata Frullania tetraptera 1863 1867 103 104 Separata of Mex. Leverm.: 242 K Danske Vid Selks Skrifter, ser 5, 6: 338 [Mexico] Lindenb. & Gottsche 1847 Nees & Mont. 1838 Page 378 semiconnata var. coronata Gottsche 1864 Typus: Colombia: Prov. Bogota, Laches, 2650 m -.11.1859, Lindig 1714 Basionym: Distribution: var nov Gottsche Bonner Gradstein & Hekking = Frullania semiconnata 1864 1965 1979 478 655 483 Ann Sci Nat Bot, ser 5, 1: 169-170 (No 8 ß), pl. 20 (fig. 51) Index Hepat 5: 425 J Hatt Bot Lab 45: 113 [Cat Colombia] Lindenb. & Gottsche 1847 Frullania tetraptera Nees & Mont. 1838 semiconnata var. suborbiculata Spruce 1884 Typus: Ecuador: below Banos sd, Spruce Basionym: Distribution: var nov Spruce Bonner 1884 1965 = Frullania semiconnata Trans Proc Bot Soc Edinburgh 15: 28 Index Hepat 5: 425 Lindenb. & Gottsche 1847 Frullania tetraptera semivillosa 243 655 Nees & Mont. 1838 [auct. non L. & G.] Sande Lac. 1857 Typus: Indonesia. Java hb SANDE LACOSTE in hb LUGD BAT Basionym: Distribution: - syn nov Sande Lacoste Schiffner Verdoorn 1857 1898 1929 = Frullania grandistipula sendaica 430 831 246 Verh Konikl Akad Wetenschappen 5: Conspect Hepat Archip Indici, Batavia: 341 [syn sub Fr. rugosa] Ann Bryol 2: 156 [syn sub Fr. grandistipula] Lindenb. 1845 [non Steph. 1924] Steph. 1924 Typus: Japan. Sendai Prov. 1907, Jishiba 52 in hb G (lecto-, designated here, G[G00069109]!) syn. fide Hattori (1952) Basionym: Distribution: spec nov - syn nov - syn nov Stephani Hattori Hattori Kamimura Bonner Stephani 1924 1952 1952 1961 1965 1985 Yuzawa Yamada & Iwatsuki Atwood, Vilnet, Mamontov 2000 2006 2021 = Frullania diversitexta 687 529 1508 199 655 928 Spec Hepat 6: 552 J Hatt Bot Lab 8: 29 [syn nov] J Jap Bot 27(10): 315 [Japan] [syn nov] J Hatt Bot Lab 24: 81 [sub F diversitexta] Index Hepat 5: 426 Stephani Icon Hepat Inedit [Thyopsiella] F-180, microfiche 59 fig 4130 ["Japonia, Provinz Sendai, leg Iishiba 1907, No 52"] 1659 Natural Environmental Sci Res 13: 10 [Japan, syn sub Fr diversitexta] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] 5566 Cryptogamie, Bryologie 42(3): 19-31 [lectotype] Steph. 1897 Page 379 serrata fo. crispulodentata [auct. non Verd. 1930] S.Hatt. 1982 Basionym: Distribution: auct - syn nov Hattori Hattori 1982 1988 = Frullania serrata ssp. spinistipula serrata 495 516 J Hatt Bot Lab 51: 224 [quod pl. Novoguinea] J Hatt Bot Lab 65: 447 (S.Hatt.) S.Hatt. 1982 fo. gracilior S.Hatt. 1974 Typus: Philippines. Luzon 8.6.1965, Schuster ISE 13825a in hb NICH [typus], in hb TENN [isotypus] Basionym: Distribution: fo nov - syn nov Hattori Hattori Yuzawa Yamada & Iwatsuki Mizutani, Hasegawa & Iwatsuki = Frullania serrata 1974 1982 2000 2006 2009 544 495 1659 1470 2112 J Hatt Bot Lab 38: 172, fig 2 J Hatt Bot Lab 51: 223 Natural Environmental Sci Res 13: 18 [Japan, syn sub Fr serrata] J Hatt Bot Lab 99: 32 [Catalog of Japan] Type sepecimens Hatt Bot Lab (NICH) 35: Holotype. Philippines, Luzon I., Mountain Prov., Mt. Data. 2100-2300 m. On branches. 07 June 1965. Z. Iwatsuki, A. J and Evelyn Sharp 13825a (NICH 266083). Gottsche 1845 serrata var. grolleana (S.Hatt.) S.Hatt. 1982 [nom inval] Basionym: Frullania grolleana Distribution: New Guinea S.Hatt. 1972 Hattori Hattori & Streimann Hattori & Piippo Hattori = Frullania grolleana 1982 1985 1986 1988 495 504 557 516 J Hatt Bot Lab 51: 225 [comb et stat nov] J Hatt Bot Lab 59: 118 [New Guinea] Acta Bot Fennica 133: 34 J Hatt Bot Lab 65: 428-431 [Fr. grolleana is a distinct species] S.Hatt. 1972 Frullania serrata ssp. grolleana serrata ssp. hamatispina (S.Hatt.) S.Hatt. 1982 S.Hatt. 1974 Typus: Papua New Guinea. Western Highlands. "New Guinea: Western Highlands, road from Mt. Hagen to Mendi, Nebilyer River. On tree, on edge of Nothofagus-forest exposed" 23/10/1968, Zanten, B.O. van 68-3264/a in hb L L0060939 [holotypus] in hb L L0060940 (25/19/1968, Zanten, B.O. van 68-3313/f - New Guinea: Papua, N. slopes of the Mount Gilowe, above 'cavern'."[typus] in hb JE [holotypus] - in hb NICH [isotypus] Basionym: Distribution: ssp nov - syn nov Hattori Hattori Hattori Grolle & Piippo Mizutani, Hasegawa & Iwatsuki 1974 1975 1982 1984 2009 546 547 495 143 2112 J Hatt Bot Lab 38: 262-265, fig 132-134 J Hatt Bot Lab 39: 300-301 J Hatt Bot Lab 51: 225 Acta Bot Fennica 125: 24 [Cat. West Melanesia] Type sepecimens Hatt Bot Lab (NICH) 35: Isotype. New Guinea, W. Highlands, road from Mt. Hagen to Medi. 2300 m. 1968. B. a. van Zanten 68-32641a (NICH). = Frullania serrata var. hamatispina (S.Hatt.) S.Hatt. 1982 Page 380 serrata ssp. triangularis [nom herb in hb L L0060941] Typus: Solomon Islands. Guadalcanal "Br. Solomon Islands: Guadalcanal, Tasi-camp. Hanging from exposed branches of rainforest undergrowth." 24/9/1968, Zanten, B.O. van 68-2430 in hb L L0060941 [typus] - http://145.18.162.53:81/c8?ent=300017&rec=69099&sct=1 Basionym: Distribution: = serrata ssp. van-zantenii (Kamim. & S.Hatt.) S.Hatt. 1980 Basionym: Frullania van-zantenii Distribution: Kamim. & S.Hatt. 1973 comb nov Hattori 1980 489 Hattori Grolle & Piippo Grolle & Piippo 1982 1984 1984 495 143 143 = Frullania van-zantenii setulosa J Hatt Bot Lab 47: 114-116, fig. 233 [Fr. van-zantenii as ssp. to Fr. serrata] J Hatt Bot Lab 51: 229, fig. 2 [Fr. van-zantenii again as species] Acta Bot Fennica 125: 24 [Cat. West Melanesia] Acta Bot Fennica 125: 24 [Cat. West Melanesia] Kamim. & S.Hatt. 1973 Ångstr. 1873 Typus: Society Isl. Tahiti 1.11.1852, N.J. Anderson in hb G Basionym: Distribution: spec nov Ångström NN Bescherelle Stephani Verdoorn Verdoorn Bonner Miller, Whittier & Whittier Stephani 1873 1875 1898 1910 1930 1930 1965 1983 1985 578 1495 871 686 293 551 655 609 928 - syn nov Hattori 1986 506 Öfversigt K Vetensk Akad Förh 30 (5): 136-137 [Tahiti] Hedwigia 14: 89-90 [Tahiti] J Bot Morot 12: 138 [Tahiti] Spec Hepat 4: 448-449 Ann Bryol Suppl 1: 49 [syn sub Fr. reflexistipula] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 428 Prod Flora Hepat Polynesiae: 138 Stephani Icon Hepat Inedit [Galeiloba] C-182, microfiche 52 fig 3584 ["Tahiti, Anderson"] J Hatt Bot Lab 60: 218 [syn sub Fr. reflexistipula var. novocaledonica] = Frullania reflexistipula var. novocaledonica S.Hatt. 1977 setulosa Steph. ex Miller, Whittier & Whittier 1983 [not existing; error] Basionym: Distribution: Stephani Verdoorn Miller, Whittier & Whittier 1908 1930 1983 870 293 609 Hattori Bapna & Kachroo 1986 2000 506 712 = Frullania ericoides Rev Bryol 35: 29 [setulosa not existing] Ann Bryol Suppl 1: 43 [not mentioned there] Prod Flora Hepat Polynesiae: 138 [syn sub Fr. squarrosa according Verdoorn 1930] J Hatt Bot Lab 60: 212 [syn sub Fr. ericoides] Hepaticology in India II: 169 [syn sub Fr. ericoides] (Nees) Mont. 1839 Page 381 shimperi Nees by Chuah-Petiot 1995 [orth var] Typus: Basionym: Distribution: Chuah-Petiot = Frullania schimperi shinii 1995 31 Trop Bryol 10: 51 [Kenya: Mount Kenya] Nees 1845 S.Hatt. 1982 [nom nud] (tentative name) Typus: Morobe Province. Mt. Kaindi 2400 m alt. on bark in Castanopsis mossy forest, coll.H.Tagawano.471.herb.Shin-28257. Basionym: Distribution: Shin 1982 3778 Mim.KagoshimaUniv.Res.CenterS.Pac 3(1): 26 = siamensis N.Kitag., Thaithong & S.Hatt. 1977 Typus: China: Chiangrai, Doi Phacho, 1700 m ?,Kitagawa 3709 in hb NICH [holotypus], in hb KYO [dubl.] Basionym: Distribution: China spec nov - comb nov Kitagawa, Thaithong & Hattori 1977 in Hattori, Thaithong & Kitagawa Hattori 1982 Hattori & Lin 1985 Mizutani, Hasegawa & Iwatsuki 2009 480 J Hatt Bot Lab 43: 452-455, fig. 3-4(a-c) [Thailand] 72 505 Bull Natn Sci Mus Tokyo, ser B (Bot), 8 (3): 99 [China] J Hatt Bot Lab 59: 148-149 [comb nov - Fr yuennanensis var. siamensis (Kitag. et al.) Hatt. & Lin, comb. nov.] 2112 Type sepecimens Hatt Bot Lab (NICH) 36: Holotype. Thailand, Payap, Chiangrai, Mt. Doi Pacho (Doi Langka). 1700 m. On tree-trunk at ist northern ridge. 27 Dec. 1965. N. Kitagawa T3709 (NICH 354397). = Frullania yuennanensis var. siamensis (N.Kitag., Thaithong & S.Hatt.) S.Hatt. & P.J.Lin 1985 siamensis 1977 fo. retusa Thaithong, Kitagawa & S.Hatt. in S.Hatt., Thaithong & Kitagawa Typus: Thailand. ? ? Basionym: Distribution: Thailand fo nov Thaithong, Kitagawa & Hattori 1977 in Hattori, Thaithong & Kitagawa Hattori & Lin 1985 Sahut 1997 Mizutani, Hasegawa & Iwatsuki 2009 Mizutani, Hasegawa & Iwatsuki 2009 480 J Hatt Bot Lab 43: 455, fig. 4(d-k) [Thailand] 505 J Hatt Bot Lab 59: 148-149 1390 http://mail2.sc.chula.ac.th/~thaiplants/RU/pdf/abstract_BS/SP_40_Sah ut.pdf 2112 Type sepecimens Hatt Bot Lab (NICH) 36:Other 26 syntype specimens: NICH 215463, 297295, 297354, 297543, 297577, 297582, 297591, 297615, 297850,318155, 354378 - 1354393). 2112 Type sepecimens Hatt Bot Lab (NICH) 36: Syntype. Thailand, Chiangrai, Mt. Doi Tung, north of Changrai. 1200-1450. On treebranches in moist evergreen forest at the higher elevation of the mountain. 25 Sept. 1967. N. Kitagawa T12449 (NICH 354377). = Frullania yuennanensis var. siamensis (N.Kitag., Thaithong & S.Hatt.) S.Hatt. & P.J.Lin 1985 Page 382 silvestris Steph. 1908 [non sylvestris Sim 1926] Typus: New Caledonia. Forets du Col d'Annien 1907, Le Rat in hb E.G. Paris 199, in hb G 18448 Basionym: Distribution: spec nov Stephani Paris Stephani Verdoorn Verdoorn Frye & Clark Kamimura Bonner Hattori Miller, Whittier & Whittier Stephani - syn nov = Frullania caledonica 1908 1910 1911 1930 1930 1947 1961 1965 1981 1983 1985 870 1455 686 293 551 204 199 655 492 609 928 Rev Bryol 35: 29 Rev Bryol 37 (6): 130 [New Caledonia: St. Vicent] Spec Hepat 4: 455 [spec nov] Ann Bryol Suppl 1: 34 [syn sub Fr. squarrosa] Nederl Kruidk Arch Jrg 1930 (2): Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa, as "sylvestris"] J Hatt Bot Lab 24: 17 [syn sub Fr. squarrosa] Index Hepat 5: 428 J Hatt Bot Lab 49: 365-368, fig 6 [syn sub Fr. caledonica] Prod Flora Hepat Polynesiae: 138 Stephani Icon Hepat Inedit [Galeiloba] C-183, microfiche 52 fig 3585 ["N Caledonia (Lerat), Herb Gen Paris No 199"] Gottsche ex Steph. 1894 [non (Schuster) Schuster 1992] Frullania squarrosula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 sinensis [auct. non Steph. in Levier 1906] Verd. in Hand.-Mazz. 1930 {Yunnan, leg Gebauer sn} Basionym: Distribution: auct - syn nov Verdoorn in Handel-Mazzetti Hattori & Lin 1930 1985 550 505 = Frullania kagoshimensis ssp. hunanensis sinensis 6668] Typus: Symb Sinicae 5: 40 [Gebauer sine numero] J Hatt Bot Lab 59: 137 (S.Hatt.) S.Hatt. & P.J.Lin 1985 [auct. non Steph. in Levier 1906] Verd. in Hand.-Mazz. 1930 [Handel-Mazzetti Handel-Mazzetti 627 & 6668 Basionym: Distribution: Verdoorn in Handel-Mazzetti Hattori & Lin = Frullania handel-mazzettii sinensis 1930 1985 550 505 Symb Sinicae 5: 40 J Hatt Bot Lab 59: 135-136 S.Hatt. 1981 [auct. non Steph.] Verd. in Hand.-Mazz. 1930 Basionym: Distribution: Verdoorn in Handel-Mazzetti Hattori & Lin = Frullania consociata 1930 1985 550 505 Symb Sinicae 5: 40 [No 10203] J Hatt Bot Lab 59: 132 [compared with Fr. consociata], 144 [compared with Fr. sinoshaerantha] Steph. 1910 Page 383 sinensis 10203] [auct. non Steph. in Levier 1906] Verd. in Hand.-Mazz. 1930 [Handel-Mazzetti Basionym: Distribution: auct - syn nov Verdoorn in Handel-Mazzetti Hattori & Lin Hattori & Lin = Frullania sinosphaerantha Frullania bhutanensis sinosphaerantha 1930 1985 1985 550 505 505 Symb Sinicae 5: 40 [specimen Handel-Mazzetti 10203] J Hatt Bot Lab 59: 132 [syn sub Fr. consociata] J Hatt Bot Lab 59: 144 [syn sub Fr. sinosphaerantha] S.Hatt. & P.J.Lin 1985 S.Hatt. 1971 S.Hatt. & P.J.Lin 1985 Typus: China. Yunnan, Gong Shan Xian, .., 200m 3.9.1982, M.Z. Wang 11509b in hb NICH [holotypus] Basionym: Distribution: China (Yunnan) spec nov - syn nov Hattori & Lin Piippo Piippo, He, Koponen, Refearn & Li Bai in Gao & Cao Wu & Wang Bai Bai Gao & Lai NN Cao, Zhu, Tan, Guo, Gao, Wu & Li Mizutani, Hasegawa & Iwatsuki 1985 1990 1998 505 521 697 J Hatt Bot Lab 59: 144-146, fig 4 J Hatt Bot Lab 68: 47 [annotated catalogue of China] J Hatt Bot Lab 84: 144 [checklist of Yunnan, China] 2000 2000 2001 2002 2003 2004 2006 1019 2671 1742 1189 1441 1399 1414 Fl Yunnanica 17: 395, fig 197(1-10) [China, Yunnan] Bryoflora of Hengduan MTS (Southwest China): 124-127, fig. 56(1-5) Guizhou Science 19(4): 45 [China, Yunnan] Chenia 7: 19 [Yunnan, China] Illustrations Bryophytes China: 1103, fig 513(1-9) [China] Red List of Chinese Endangered Bryophytes J Hatt Bot Lab 99: 285 [Red list of Chinese endangered bryophytes] 2009 Gao & Wu Schäfer-Verwimp & Winter 2010 2020 2112 Type sepecimens Hatt Bot Lab (NICH) 36: Holotype. China, Yunnan, Gong Shan Xian, Do Lang Jiang Gong She, Siong Dang - Long Di Geng. 2000 m. In road side. 3 Sept. 1982. M. Z. Wang 11509b (NICH 401061). 2349 Genera Hepat. et Anth. Sinicorum: fig. 239(1-10) [key in Chinese] 5751 Frahmia 20: 9-11, fig 9-11 [syn sub Fr bhutanensis] = Frullania bhutanensis S.Hatt. 1971 sinuata var. tenella (Sande Lac.) Sande Lac. 1857 Basionym: Frullania tenella Distribution: Sande Lac. 1854 [non Caspary 1887, nec Steph. 1910] var nov Sande Lacoste Sande Lacoste in Junghuhn Stephani Verdoorn Bonner Hattori 1857 1857 1911 1930 1965 1975 430 1490 686 293 655 66 - comb nov Hattori 1982 422 Miller, Whittier & Whittier Stephani 1983 1985 609 928 Verh Konikl Akad Wetenschappen 5: 87-88, pl 17(1-9) Plantae Junghuhnianae, Hepaticae: 540-541 Spec Hepat 4: 644 [syn sub Fr. sundaica] Ann Bryol Suppl 1: 134 [syn sub Fr. junghuhniana] Index Hepat 5: 429 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 148-149 [syn sub Fr. junghuhniana var. minutissima] Misc Bryol Lichen 9 (6): 123-124 [comb nov: Fr. junghuhniana var. tenella] Prod Flora Hepat Polynesiae: 138 Stephani Icon Hepat Inedit [Diastaloba] B-84, microfiche 49 fig 3331 ["Sande Lacoste, Hep Javana tab 27"] = Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 Page 384 sinuata ssp. novoguineensis var. latistipula S.Hatt. 1975 [nom inval] Basionym: Frullania papuana var. latistipula S.Hatt. 1974 Distribution: comb nov Hattori Hattori Grolle & Piippo Hattori & Piippo = Frullania papuana 1975 1982 1984 1986 547 495 143 557 J Hatt Bot Lab 39: 304 J Hatt Bot Lab 51: 217 Acta Bot Fennica 125: 24 [Cat. West Melanesia] Acta Bot Fennica 133: 32 [nom invalid] Verd. 1930 sinuata ssp. novoguinensis [nom herb in hb L L0060942] Typus: Indonesia. New Guinea, Wilhelmina Mountain. "Dutch New Guinea, 2 km E. of Wilhelminatop. Epiphytic. On Drimys brassii A.C.Sm." -/9/1938, Brass, L.J.; Meyer-Drees, E. 10126 in hb L L0060942 [holotypus] - http://145.18.162.53:81/c8?ent=300017&rec=69100&sct=1 Basionym: Distribution: = solanderi Colenso by Steph. 1892 [orth var] Basionym: Distribution: Stephani 1892 = Frullania solanderiana solmsiana 210 J Linn Soc Bot 29: 274 Colenso 1889 Steph. 1911 Typus: Indonesia. Java sd, Solms 47 in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Bonner Stephani = Frullania cordistipula 1911 1930 1965 1985 686 293 655 928 Spec Hepat 4: 541-542 Ann Bryol Suppl 1: 88 [syn sub Fr. cordistipula] Index Hepat 5: 429 Stephani Icon Hepat Inedit [Thyopsiella] F-154, microfiche 59 fig 4093 ["Java, Graf Solms No 47"] (Nees) Nees 1839 {Brazil} Page 385 spathulistipa Steph. 1910 [non (Reinw., Blume & Nees) Lindenb. 1838] Typus: Australia. "New England", Mt. Duval near Armidale 4.11.1903, W:W: Watts 760pp hb LEVIER 4785, in hb G 18450 [typus] Basionym: Distribution: Australia (NSW), Tasmania spec nov -nom nov Stephani Watts Bonner Hattori Stephani 1910 1912 1965 1979 1985 686 1940 655 486 928 Scott & Bradshaw Tan & Engel 1986 1986 56 4286 Wigginton 2002 1012 Carthy Glenny, Meagher & Brown 2003 2004 1125 1126 Glenny, Meagher & Brown 2004 1127 McCarthy 2006 1385 = Frullania sinskeana spathulistipa Spec Hepat 4: 415 [as Trachycolea] unpublished Msc Index Hepat 5: 430 J Hatt Bot Lab 46: 143-144, fig 43(a-h) from type Stephani Icon Hepat Inedit [Galeiloba] C-222, microfiche 53 fig 3639 ["Australia orient, Watts legit, Levier 4785"] Brunnonia 8 (1): 36 & 144 [Australia: NSW] J. Hatt. Bot. Lab.. No. 60: 335, [nom. nov. Frullania sinskeana J.J.Engel et B.C.Tan 1986, Australia] Trop Bryol Res Rep 2: 28 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Flora Australia Suppl Ser 21: 47 [Australia (NSW, TAS)] NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Checklist Australian Liverworts [Australia NSW TAS] J.J.Engel & B.C.Tan 1986 (Reinw., Blume & Nees) Lindenb. 1838 [non Steph. 1910] Basionym: Distribution: Tan & Engel = Thysananthus spathulistipus spatiosa 1986 4286 J. Hatt. Bot. Lab.. No. 60: 335, [nom. nov. Frullania sinskeana J.J.Engel et B.C.Tan 1986, Australia] (Reinw., Blume & Nees) Lindenb. 1845 Steph. ex Steph. 1985 [nom herb] {Micronesia} Basionym: Distribution: Stephani 1985 928 Stephani Icon Hepat Inedit [Homotropantha] D-35, microfiche 54 fig 3758 ["Ponape, Ost Karolinen (Ledermann legit"] = Page 386 spatuliflora Typus: Spruce 1895 Dominica. Laudat 1892, W.R. Elliott 857 in hb MANCH Basionym: Distribution: Dominica, Martinique spec nov - syn nov - syn nov Spruce Stephani Arnell Bonner Stotler Stotler Stephani 1895 1911 1959 1965 1969 1970 1985 Fulford Schäfer-Verwimp Lavocat Bernard & SchäferVerwimp 1987 2010 2011 = Frullania neesii J Linn Soc Bot 30: 333, pl 20(1-4) Spec Hepat 4: 524-525 Svensk Bot Tidskr 53 (4): 500 [Martinique] Index Hepat 5: 430 Nova Hedwigia 18: 450 [syn sub Fr. neesii] The genus Frullania, separata: 450 [syn sub Fr. neesii] Stephani Icon Hepat Inedit [Thyopsiella] F-125, microfiche 58 fig 4052 ["Dominica, Elliot legit"] 99 Mem NY Bot Gard 45: 272 [Dominica] 2460 Crypt Bryol 31(4): 347 [syn sub Frull kunzei, Dominica, West Indies] 3130 Cryptogamie, Bryologie 32(3): 251 [Martinique] "asFr spathuliflora Arnell 1959" Lindenb. 1845 Frullania kunzei sphaeroflora 238 686 448 655 4761 194 928 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1911 Typus: Madagascar. betsileo sd, La Couture 172 in hb G [Steph Icon ined No 1265 & 1630] Basionym: Distribution: Madagascar spec nov - syn nov Stephani Demaret & Vanden Berghen Bonner Augier Vanden Berghen Stephani 1911 1950 1965 1974 1976 1985 Piippo Grolle Wigginton 1990 1995 2002 Wigginton Uribe 2004 2008 = Frullania bullata 686 60 655 821 185 928 Spec Hepat 4: 591 Bull Jard Bot Etat Bruxelles 20: 2-3, fig 1 Index Hepat 5: 430 Annales de la Faculté des Sciences du Cameroun 17: 85 [Cameroun] Bull Jard Bot Nat Belg 46 (1/2): 46: 190 Stephani Icon Hepat Inedit [Meteriopsis] E- 7, microfiche 55 fig 3773 ["Madagascar, 160.172 (Lacouture legit)"] 521 J Hatt Bot Lab 68: 38 [annotated catalogue of China] 250 Bryophyt Bibl 48: 64 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1513 Caldasia 30(1): 90, [belongs to subgen Frullania] [Monography of subgenus Meteoriopsis] Steph. 1910 Page 387 spicata Steph. 1910 [hom illeg] [non (Lehm. & Lindenb.) Lehm. & Lindenb. 1845] Typus: Mexico. Puebla State, Honey Station, 7000ft 20.6.1908, C.G. Pringle 15-333 in hb G G00060666 - n° SIB 131169/1 [holotypus] in hb MICH MICH514769 [isotypus] Basionym: Distribution: Mexico spec nov - syn nov Stephani Clark & Frye Clark & Svihla Bonner Stephani 1910 1947 1948 1965 1985 Yuzawa 1987 Yuzawa & Koike Fulford & Sharp Fogel 1989 1990 1997 = Frullania glomerata Mont 1842] spiniflora 686 322 325 655 928 519 105 629 Spec Hepat 4: 392-393 Bryologist 50: 313-316, fig 1-19 Bryologist 51: 26 [key to Fr. of Middle America: Mexico] Index Hepat 5: 431 Stephani Icon Hepat Inedit [Galeiloba] C- 74, microfiche 51 fig 3439 [from type; "Mexico, Honey Station, 20/6. 1908 Pringle"] hb annotation by Yuzawa - type of Frullania spicata STEPH, Dez 21 1987 J Hatt Bot Lab 66: 352-355, fig 8(a-r) [syn sub Fr. glomerata] Mem NY Bot Gard 63: 46 nom nov as Fr pringlei, Mexico] http://141.211.110.91/livrqury.htm [16.11.1997] - Genus=Frullania, printed 5.1.1998 11:11:38: page 2-3 of 3 (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Glenny, Meagher & Brown 2003 [nom nud] Basionym: Distribution: Australia (NSW) Carthy Glenny, Meagher & Brown 2003 2003 Glenny, Meagher & Brown 2003 1125 Flora Australia Suppl Ser 21: 112 [nom nud] 1126 NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html = spiniloba Steph. 1910 Typus: Ecuador coll. ign. in hb G G00067857 - n° SIB 141500/39 - 20976 (ex hb MÜLLER, Ecuador, leg ?) [lectotypus]; in hb G G00067858 - n° SIB 141499/38 (Panama, Chirigue, Frère Hélion - Lacouture 414) Basionym: Distribution: Mexico, Panama, Ecuador, [Ecuador u/o Colombia - Gradstein in press] spec nov comb nov - comb nov Stephani Stephani Clark & Svihla Bonner Schuster Stephani 1910 1916 1950 1965 1985 1985 686 222 334 655 441 928 Spec Hepat 4: 336-337 Biblioth Bot 87: 240 [Bolivia] Bryologist 53: 66-69, fig. 1-17 Index Hepat 5: 431 Phytologia 57 (5): 372-373 [sect Spinilobae - type species for nov sect] Stephani Icon Hepat Inedit [Chonanthelia] A- 50, microfiche 47 fig 3141 ["Ecuador leg .... / auch Panama ..... Herb Müller"] J Hatt Bot Lab 63: 425-436 [morphology / systematics] Mem NY Bot Gard 63: 46 [Mexico] J Hatt Bot Lab 70: 269-271 [scom as: Fr arecae var. spiniloba] Trop Bryol 15: 180 [Checklist Panama] Yuzawa, Mues & Hattori Fulford & Sharp Yuzawa Stotler, Allen, Gradstein, McGuinness, Whittemore & Chung Gradstein, Meneses & Arbe Schäfer-Verwimp Hentschel et alii 1987 1990 1991 1998 509 105 525 703 2003 2014 2015 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr arecae var. spiniloba] 3977 Phytotaxa 172(3):214 [Panama] 4097 Phytotaxa 220 (2): 136 [comb nov as Fr obscura var. spiniloba] = Frullania obscura var. spiniloba (Steph.) Hentschel & von Konrat 2015 Page 388 spinistipula Steph. ex Paris 1910 [nom nud] Basionym: Distribution: nom nud Stephani in Paris Hattori 1910 1944 = Frullania spinistipula spinistipula 911 943 Rev Bryol 37 (1): 41 [New Caledonia: Isle of Pines] Bull Tokyo Sci Mus 11: 144 [syn nov sub Fr. hampeana] Steph. 1911 [non S.Hatt. 1972] Steph. ex Theriot 1910 [nom nudum] Basionym: Distribution: nom nud Stephani ex Theriot 1910 = Frullania spinistipula spinosistipula - syn nov Steph. 1911 [non S.Hatt. 1972] S.Hatt. 1973 Basionym: Frullania spinistipula Distribution: nom nov S.Hatt. 1972 [hom illeg] [non Steph. 1911] Hattori Hattori Hattori Hattori Grolle & Piippo 1973 1974 1975 1982 1984 = Frullania serrata ssp. spinistipula sprengelii 1915 Bull Acad Int Geogr Bot 20: 104 [nom nud] 542 546 547 495 143 J Hatt Bot Lab 37: 151 [nom nov] J Hatt Bot Lab 38: 267 [New Guinea] J Hatt Bot Lab 39: 304-305 J Hatt Bot Lab 51: 225-226 Acta Bot Fennica 125: 24 [Cat. West Melanesia] (S.Hatt.) S.Hatt. 1982 Gottsche 1845 [non Steph. 1894] Basionym: Distribution: - comb nov Lehmann & Lindenberg in G., L. 1845 & L. 2850 Syn Hepat: 428 [nom inval, pro syn sub Frtrinervis (beta) berteroana] = Frullania trinervis var. Sprengelii Lehm. & Lindenb. 1845 Frullania trinervis sprengelii (Lehm.) Drège 1843 Steph. 1894 [non Gottsche 1845] Typus: Magellan Straits, Christmans Sounds, "in Berberide ilicifolia" 20.12.1775, Forsters ex hb Spreng., hb STEPHANI 128, in hb G 15014 Basionym: Distribution: spec nov - syn nov Stephani Stephani Stephani Kühnemann 1894 1911 1911 1937 228 686 721 1031 Kühnemann Bonner Hattori Hässel de Menéndez Stephani 1949 1965 1975 1983 1985 657 655 411 48 928 = Gackstroemia hariotiana Hedwigia 33: 167-168 Spec Hepat 4: 663 Kungl Svenska Vetenskapsakad Handl 45 (9): 90 Revista Centro Estud Doct Ci Nat, Buenos Aires 1 (3): 167 [Estrecho de Magellanes] Lilloa 19: 355 [Argentina: Magellan Strait] Index Hepat 5: 432 Misc Bryol Lichen 7 (3): 47-48, fig 2 [sect nov. = Sprengelia] Lindbergia 9: 97 [= Gackstroemia hattorianta] Stephani Icon Hepat Inedit [Diastaloba] B-40, microfiche 49 fig 3277 ["cum fr. Magellanica in Berberide ilicifolia Herbar Sprengel"] (Bescherelle & C.Massal.) Grolle 1967 Page 389 spruceana Steph. 1894 Typus: Peru. In Tunuragua mountains Basionym: Frullania robusta Distribution: Peru spec nov - syn nov sd, Spruce Steph. 1911 [non Kiaer ex Renauld & Cardot 1891, nec Spruce [Hepat exsicc] ex Steph. 1911] Stephani Stephani Clark 1894 1911 1953 Bonner Stotler Stotler Winkler Morales-Z 1965 1969 1970 1976 1991 228 686 352 Hedwigia 33: 160 Spec Hepat 4: 505 [syn sub Fr. robusta in Spruce, Hep. exsiccatae.] Bryologist 56: 140-142, fig(1-14) [correct name, as "Spruce, Hep. exsiccatae" were never published] 655 Index Hepat 5: 432 4761 Nova Hedwigia 18: 436 [syn sub Fr. brasiliensis] 194 The genus Frullania, separata: 436 [syn sub Fr. brasiliensis] 5 Rev Bryol Lichen 42 (3): 811 [Colombia] 307 Tropical bryology 4: 36 [Costa Rica; sub Fr brasiliensis] = Frullania robusta Steph. 1911 [non Kiaer ex Renauld & Cardot 1891, nec Spruce [Hepat exsicc] ex Steph. 1911] Frullania brasiliensis squamuligera Raddi 1822 Spruce 1884 [non Goebel 1928] Typus: Ecuador. "Andes Quitenses, ad cataract Agoyan", 5500ft 1857, Spruce s.n. in hb G? - in hb MANCH Basionym: Distribution: Ecuador, Bolivia spec nov Spruce Stephani Stephani Bonner Stephani - syn nov 1884 1910 1916 1965 1985 Stotler in Seaward & FitzGerald 1996 Gradstein 2021 = Frullania glomerata Mont 1842] squamuligera 243 686 222 655 928 Trans Proc Bot Soc Edinburgh 15: 32-34 Spec Hepat 4: 387 Biblioth Bot 87: 242 [Bolivia] Index Hepat 5: 432 Stephani Icon Hepat Inedit [Galeiloba] C- 75, microfiche 51 fig 3440 [from type; "Andes quitenses, Spruce - ad cataratas Agoyan 5500ft"] 1092 Richard Spruce, Roy Bot Gard Kew: 131, fig 11 [subgen Trachycolea] 5642 Mem NY Bot Gard 121: 367 [syn sub Fr glomerata] (Lehm. & Lindenb.) Nees & Mont. 1838 [non Mont 1840, nec Goebel 1928 [hom illeg] [non Spruce 1884] Typus: Indonesia. Sumatra ? Basionym: Distribution: Indonesia (Sumatra) Goebel Verdoorn Verdoorn 1928 1929 1930 234 247 293 Bonner 1965 655 Ann Jard Bot Buitenzorg 39: 48, pl 9(89) [without description] Ann Crypt exot 2 (1): 74 Ann Bryol Suppl 1: 40 [may be syn with Fr. squarrosa var. planescens fo. chonanthelioides] Index Hepat 5: 432 = Frullania squarrosa var. planescens fo. chonanthelioides Verd. 1930 Frullania ericoides (Nees) Mont. 1839 Page 390 squarrosa [auct. non (Reinw., Blume & Nees) Nees 1845] Austin 1874 (Sandwich Isl) Basionym: Distribution: - syn nov Austin Austin Evans = Frullania densepinnata 1874 1874 1900 Bull Torrey Bot Club 5: 15 [Oahu - Sandwich Isl] Bull Torrey Bot Club 5: 17 [Sandwich Isl] Trans Connet Acad Arts Sci 10 (1): 399 Steph. in Levier 1901 [nom nud] Frullania densepinnata Frullania apicalis 668 669 236 Steph. 1911 Mitt. 1879 Page 391 squarrosa (Reinw., Blume & Nees) Nees 1845 Typus: Indonesia. Java Blume in hb STR in hb Graz in hb Leyden in hb GLM 05963, 05964. 05965 Basionym: Jungermannia squarrosa Reinw., Blume & Nees 1824 [hom illeg] [vide Grolle 1995] Distribution: Madeira [Portugal], Canary Isl [Spain], Cape Verde Isl, Annobon, Democratic Republic of Congo, Ethiopia, Uganda, Zambia, Zimbabwe, South Africa; USA ( CT RI VT ), Mexico, Costa Rica, Bolivia, Venezuela, Surinam, Brazil, Argentina, China (Anhui), Taiwan, Korea (South), Ryukyu (Ishigaki), India, Sri Lanka, Thailand, Vietnam, Nepal, Indonesia (Sumatra, Java, Ceram), Borneo, New Guinea, Yemen Rep, Australia (NSW, QLD, VIC), Hawaii, Henderson Isl stat nov auct Dumortier Nees ex G., L. & N. Nees ex G., L. & N. Lindenberg & Gottsche Sande Lacoste in Junghuhn Sande Lacoste Mitten Mitten Gottsche Gottsche Sande Lacoste 1835 1845 1847 1851 1857 1857 1861 1863 1863 1864 1864 256 2850 2848 282 1490 430 574 253 103 478 842 Gottsche 1867 104 Mitten in Seemann Austin Ångström Baldwin Mitten Sande Lacoste in Veth 1871 1874 1876 1877 1879 1884 637 668 579 901 1913 807 Spruce Gottsche 1884 1886 243 2390 Stephani Mitten in Balfour Wild Stephani Britton Stephani Stephani Pearson 1886 1888 1888 1888 1889 1889 1891 1891 944 2093 1038 937 2441 940 209 1921 Rec d´Obs Jungerm., Tournay, 1: 13 Syn Hepat: 416 (No 10) Syn Hepat: 772 Linnaea 24 (5): 637 [Suriname] Plantae Junghuhnianae, Hepaticae: 536 Verh Konikl Akad Wetenschappen 5: 78 J Linn Soc Bot 5: 119 [Nepal, Sri Lanka] J Linn Soc Bot 7: 169 [Prince's Island] Separata of Mex. Leverm.: 238 Ann Sci Nat Bot, ser 5, 1: 167 (No 5) Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 312 [Java, Sumatra, Borneo, Ceram] K Danske Vid Selks Skrifter, ser 5, 6: 334-335 [as Fr. squarrosa vulgaris] [Mexico] Flora Vitiensis: 417 [Sandwich Isl] Bull Torrey Bot Club 5: 15 [Oahu - Sandwich Isl] Öfversigt K Vetensk Akad Förh 33 (7): 88 [Brazil] Hawaiian Almanac: 42 [Hawaii] Phil Trans Roy Soc Lond 168: 401 [Isle of Rodriques] Midden Sumatra, Reizen en Onderzoekingen der Sumatra Expeditie, 4 (2): 44 [Sumatra] Trans Proc Bot Soc Edinburgh 15: 32-33 (No 22) Bot. Centralblatt 26: 204 [Drawings com. by Jack, hb no. 1545 - 2Bl. Mit vergleichenden Zeichnungen aus Oahu (Didrichsen), Krakakamma (Ecklon), Mexico (Leibold), Trinidad (Crüger). Bol Soc Broteriana 4: 180 [152] [South Africa] Botany of Socotra: 336 Proc Roy Soc Queensland 5 (3): 120 [Queensland] Hedwigia 27: 109 [Principe] Catalogue of plants in New Jersey [USA, New Jersey]: 347 Hedwigia 28: 160-161 [Australia] Hedwigia 30 (6): 267 [Kilimandscharo-Gebiet] Christiania Videnskabs-Selskabs Forhandlinger 1890 (2): 4 [Madagascar] Icon Zeitschrift Forhandlinger i Videnskaps-Selskabet i Kristiania 1890 (2): 4 Rev Bryol 18 (4): 57 [Madagascar, Maurice] Separata aus Engler-Prantl, Natürliche Pflanzenfamilien, Teil 1, Abt 3: 134 Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [Verbreitet - Gebirgsbuschwald, Hochwald] Bihang Kgl Sv Vetensk-Akad Handl bot 23 III (2): 20 [Brazil] Bull Herb Boissier 5: 79 [Japan] Bull Herb Boissier 5: 842[:260] [Hawaii] Trans Connet Acad Arts Sci 10 (1): 14-15, pl. 6 Conspect Hepat Archip Indici, Batavia: 343-345 Bot Mag Tokyo 14 (no 162): 181 Hedwigia 39: 208 [Java] Cat Afr Plants Welwitsch 2 (2): 316 [Angola] Flora Deutsch Schutzgebiete: 79 Stephani in Renauld & Cardot Schiffner 1891 1895 1453 1015 Stephani in Engler 1895 1481 Stephani Stephani Stephani Evans Schiffner Inoue Schiffner Stephani Schiffner & Stephani in Schumann & Lauterbach Barbour Schiffner in Engler & Prantl Paris 1897 1897 1897 1897 1898 1900 1900 1901 1901 1059 213 214 237 831 802 904 883 1493 1902 1905 1906 2233 Bryologist 5(1): 4 [USA] 809 Natürliche Pflanzenfamilien, Teil 1, Abt 3, 1: 134 676 Rev Bryol 33 (2): 28 [New Caledonia] Page 392 Sawada 1909 930 Gola in Savoia Stephani Paris Paris Massalongo Iishiba Stephani in Brunnthaler Evans Stephani & Watts Gola Stephani Gola Evans in Britton Evans in Britton & Millspaugh Sim Massalongo Verdoorn Verdoorn Verdoorn in Handel-Mazzetti Verdoorn Verdoorn 1909 1910 1910 1910 1911 1912 1913 1914 1914 1914 1916 1916 1918 1920 1926 1927 1929 1930 1930 1930 1930 832 686 911 1455 903 888 219 235 220 786 222 3469 1093 1094 252 1060 245 551 550 549 293 Verdoorn 1932 232 Kashyap Verdoorn Kurz & Little 1932 1932 1933 226 553 1349 Verdoorn Verdoorn Verdoorn Chopra 1933 1934 1937 1938 233 241 556 1648 Chopra Pagan Herrera Chopra Hattori 1938 1939 1941 1943 1944 933 931 2789 205 943 Demaret & Vanden Berghen 1947 1101 Frye & Clark Clark & Svihla 1947 1948 204 325 Kühnemann Clark & Svihla Herzog Hattori Cufodontis Kamimura Herzog Clark 1949 1950 1952 1952 1952 1952 1953 1953 657 337 803 529 2562 873 640 1106 Miller Schuster Vanden Berghen McGregor Arnell Arnell Arnell Abeywickrama Vanden Berghen Kamimura 1953 1953 1953 1955 1956 1956 1956 1959 1961 1961 349 604 446 2240 836 847 836 200 1507 199 Bot Mag Tokyo 23 (no 268): 178-183, fig 12 [key to Japanese species, in Japanese] Il Ruwenzori, parte scientifica 1: 529 Spec Hepat 4: 388 Rev Bryol 37 (1): 41 [New Caledonia: Isle of Pines] Rev Bryol 37 (6): 130 [New Caledonia: Wagap; Lifou] Rev Bryol 38: 11 [Brasil] Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 729 [Tanzania] Trans Connet Acad Arts Sci 18: 343 [Peru] J Proc Roy Soc New S Wales 48: 109 [Lord Howe Isl]] Annali di Botanica 13: 72 [Ethiopia] Biblioth Bot 87: 242 [Bolivia] Mem Reale Accad Sci Torino, ser 2. 65: 9 [Kenya] Flora Bermuda: 468 [Bermuda] Bahama Flora: 520 [Bahamas] Trans Roy Soc S Africa 15: 41-42, figs [South Africa] Atti Real Ist Veneto Sci 87 (2): 218 [Argentina, Spegazzinius legebat] Ann Bryol 2: 131-134 [Java, Sumatra] Nederl Kruidk Arch Jrg 1930 (2): Symb Sinicae 5: 40 [China] Nova Guinea 14 (4): 541 [New Guinea] Ann Bryol Suppl 1: 34-37, fig. 18-21, 42-45, 130-132 [squarrosa = ericoides] Ann Bryol 5: 126 [Hepat select crit, ser 3 & 4 no 163-165; Java, India] & 138 Liverworts of the western Himalayas, Part 2: 11-13, pl. 1(1-5) Bull Jard Bot Buitenzorg, ser 3, 12 (1): 53 [Java] Liverworts North & Central Florida: 14, fig 22 [key to species of Florida, USA] Ann Bryol 6: 96 [Hepat select crit, ser 5 & 6 no 296; America sept.] Ann Bryol Suppl 4: 204 [Sumatra, Lombok] & 212, fig 11 [map] Blumea suppl 1: 212 [Henderson Island] Proceed Indian Acad Sci, sect B, 8: 437 [Sikkim Himalayas and Bengal] Proceed Indian Acad Sci, sect B, 7 (5): 248 [South India] Bryologist 42: 80 [Puerto Rico] Sinopsis de la Flora del Cuzco: 71 [Peru, Cuzco] J Indian Bot Soc 22: 249 [India] Bull Tokyo Sci Mus 11: 152-153, fig 93(a-h) [Japan, key to Japanese species] Bull Jard Bot Etat Bruxelles 18: 237-239 [Annobon, Oubangui-Chari, Kenya, Tanganyika, Congo Belge, Afrique australe, Madagascar] Univ Wash Publ Biol 6 (5): 776-778, fig Fr.s.(1-9) Bryologist 51: 27 [key to Fr. of Middle America: United States, Mexico (Puerto Rico, Central America, Oceanica, Australia, Ceylon, Asia, Africa, Princes Island, Bermuda] Lilloa 19: 355 [Argentina: Buenos Aires] Bryologist 53: 126-128 [male inflorescences] fig. 9-11 Feddes Repert 55: 10 [Argentina] J Hatt Bot Lab 8: 31 Phyton 4 (1-3): 77 [Ethiopia] Contrib Fl Hepat Shikokuensem, Kochi: 131 Rev Bryol Lichen 20: 128 [Costa Rica] Proc California Acad Sci 4 ser, 27 (18): 663-663 [Galapagos Isls: Floreana, Santa Cruz, Isabela; Nicaragua, Mexico] Bryologist 56: 45 Amer Midl Nat 49 (2): 569, pl 72(11) Svensk Bot Tidskr 47 (2): 269 [Uganda] Univ Kansas Sci, Bull 37(3): 83 [USA, Kansas] Ark Botanik 3 (16): 532 [East African mountains] Svensk Bot Tidskr 50 (4): 532 [Annobon] Ark Botanik 3 (16): 532 [East African mountains] Ceylon J Sci (Bio Sci) 2 (1): 58 [Sri Lanka] Bull Soc Roy Bot Belg 93 (1-2): 69 Kivu, Urundi, Katanga] J Hatt Bot Lab 24: 17-19, fig. 4 (1-8), map 3 Page 393 - syn nov Arnell Arnell Inoue Parihar Miller Miller Nakanishi Arnell Schiffner & Arnell Grolle Hong Hattori in Hara Pocs, Tixier & Jovet-Ast Grolle 1961 1961 1961 1962 1963 1963 1963 1963 1964 1966 1966 1966 1967 1968 54 180 964 1584 834 834 960 822 198 843 1107 841 1710 613 Hattori & Mizutani Inoue in Inoue & Iwatsuki Shin Vanden Berghen 1969 1970 1970 1972 106 62 302 249 Vanden Berghen Augier Augier 1973 1974 1974 1702 821 821 Vanden Berghen Abeywickrama & Jansen Mägdefrau Miller, Whittier & Whittier Grolle & Piippo Menzel Wu in Li, Gao, Wu et al. Stephani 1976 1978 1983 1983 1984 1984 1985 1985 185 794 1206 609 143 142 1169 928 Stephani 1985 928 Stephani 1985 928 Scott & Bradshaw Frey & Kürschner Reiner 1986 1988 1988 56 1119 1043 Kuo & Chiang Fulford & Sharp Best Piippo Frey & Kürschner Parihar, Lal & Katiyar Buck Grolle Dilg & Frahm Bapna & Kachroo Pant Negi, Nath & Asthana 1988 1990 1990 1990 1991 1994 1994 1995 1997 2000 2001 2001 948 105 879 521 1033 2196 28 250 645 712 1369 1373 Greene 2001 1255 Grims NN Srivastava & Alam 2001 2002 2002 1136 1266 1251 Cai NN Gradstein & Costa Gradstein, Meneses & Arbe Manyanga & Perold Wigginton 2002 2002 2003 2003 2004 2004 1192 1266 984 975 1301 1172 Bot Not 114 (2): 178-179 [Cape Verde Is.] Svensk Bot Tidskr 55 (2): 351 [Canary Is.] J Jap Bot 36(5): 187 [Formosa] Univ Allahabad studies, Bot sect 1-2: 19 [India: WH EH SI] Ark Bot, ser 2, 5: 518 [Hawaii] Ark Bot, ser 2, 5: 518 [Hawaii] Hikobia 3(4): 319 [Taiwan] Hepat S Africa: 150-153, fig 111(a-i) Österr Akad Wiss, Math Nat Kl, Denkschr 111: 90 [Brazil] Khumbu Himal 4: 266 [Nepal] Bryologist 69 (1): 412 [South Korea] Flora of Eastern Himalaya: 527-528 Bot Kozlem 54 (1): 30 [North Vietnam] Trans Brit Bryol Soc 5 (3): 541 [lectotypification, Madeira] [published 1846 not 1845] Dansk Bot Arkiv 27 (1): 95 [Thailand] Bull Natn Sci Mus Tokyo 13 (3): 478 Sci Rep Kagoshima Univ 19: 37-38 [Ryukyu: Ishigaki] Expl Hydrobiol Bassin Bangweolo et Luapula 8 (1): 82-84, fig 32(a-k) [Zaire, Zambia] Rev Bryol Lichen 39: 372 [Gabon] Ann Fac Sci Univ Fed Cameroun 17: 82, 86 [Cameroun] Annales de la Faculté des Sciences du Cameroun 17: <67-93> [Cameroun] Bull Jard Bot Nat Belg 46 (1/2): 163 MaB Nat Comm Sri Lanka, Publ 1: 5 [Sri Lanka] Nova Hedwigia 13: 22 [Venezuela] Prod Flora Hepat Polynesiae: 138-139 Acta Bot Fennica 125: 24 [Cat. West Melanesia] Willdenowia 14: 510 [syn of Fr. ericoides] Bryoflora Xizang (Tibet), Science Press, Beijing: 523, fig 227(1-5) Stephani Icon Hepat Inedit [Galeiloba] C- 76ab, microfiche 51 fig 34413442 ["H Ex No 1517, Mexico, leg Leibold"] Stephani Icon Hepat Inedit [Galeiloba] C-185, microfiche 52 fig 3587 ["India orientalis, Ghates montes, André 1908"] Stephani Icon Hepat Inedit [Galeiloba] C- 91-, microfiche 51 fig 3470 (no figure) ["auch Am sept, Conneticut Ohio et, vide America tropica"] Brunnonia 8 (1): 37 & 144 [Australia: NSW, QLD, VIC] Nova Hedwigia 46 (1-2): 77 [Arabian Peninsula & Socotra] Bol Soc Argentina Bot 25 (3-4): 323 [not in Argentina, Prov Buenos Aires] Taiwania 33: 13 [Index of Taiwan] Mem NY Bot Gard 63: 46 [Mexico] Kirkia 13 (2): 296 [Zimbabwe] J Hatt Bot Lab 68: 47 [annotated catalogue of China] Bryophyt Bibl 39: 31 [Yemen Rep] Hepat Anth India, annotated checklist: 41 [syn sub Fr ericoides] Bryologist 97 (4): 422 [Vainio's specimen from Brazil at TUR] Bryophyt Bibl 48: 65 Trop Bryol 13: 38 [South Africa, Drakensberge] Hepaticology in India II: 170 [syn sub Fr ericoides] Perspectives Indian Bryology: 14 [incorporated in bird-nests] Perspectives Indian Bryology: 296 [India; terricolous in Garhwal Himalayas] Liverworts & hornworts of New England: 7 of 9 [USA, New England (CT RI VT)] Moosherbar in hb LINZ, from Ekuador-Galapagosinseln [Internet] http://www.natureserve.org/explorer [USA. Ge, Te] J Bombay Nat Hist Soc 99(2): 235-236 [Nilgiri, key to South Indian species] Chenia 7: 151 [China, Anhui Prov.] http://www.natureserve.org/explorer [USA] Mem New York Bot Gard 87: 86 [Brazil] [sub Fr ericoides] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr xxx] SABONET Report 21: 4 [Zimbabwe] Sript Bot Belg 30: 229 [syn sub Fr ericoides, West Africa] Page 394 Glenny, Meagher & Brown 2004 Glenny, Meagher & Brown 2004 Yamada & Iwatsuki Yano Singh, Kumar & Nath 2006 2008 2008 = Frullania ericoides squarrosa 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1126 NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr ericoides; Brazil; catalog] 1281 Taiwania 53 (1): 62, [syn sub Fr ericoides] (Nees) Mont. 1839 [auct. non (Reinw., Blume & Nees) Nees 1845] Pearson 1890 Basionym: Distribution: Pearson 1891 Stephani = Frullania sandei 1894 228 Icon Zeitschrift Forhandlinger i Videnskaps-Selskabet i Kristiania 1890 (2): 4 Hedwigia 33: 147 [syn sub Fr. sandei] Kiaer in Steph. 1894 Frullania caffraria squarrosa 1921 Christiania Videnskabs-Selskabs Forhandlinger 1890 (2): Steph. 1894 fo. ericioides (Nees) Verd. 1929 Basionym: Distribution: comb nov Verdoorn Verdoorn Verdoorn in Handel-Mazzetti Verdoorn Verdoorn Verdoorn Verdoorn Bonner Miller Bapna & Kachroo = Frullania ericoides squarrosa 1929 1930 1930 1932 1932 1934 1937 1965 1968 2000 245 293 550 553 554 241 556 655 672 712 Ann Bryol 2: 134 [Java] Ann Bryol Suppl 1: 38 Symb Sinicae 5: 40 [China] Bull Jard Bot Buitenzorg, ser 3, 12 (1): 54 [Java] Nederl Kruidk Arch Jrg 1932 (2): 492 [Java] Ann Bryol Suppl 4: 204 [N Formosa] Blumea suppl 1: 212 [Austral Islands] Index Hepat 5: [Brazil, Cuba, Java] Micronesia 4 (1): 74 [key to Guam & Northern Micronesia] Hepaticology in India II: 170 [ as fo "ericoldes", syn sub ericoides] (Nees) Mont. 1839 fo. ericoides Verd. by Bapna & Kachroo 2000 [orth var] Basionym: Distribution: Bapna & Kachroo = Frullania squarrosa 2000 712 Hepaticology in India II: 170 fo. ericioides (Nees) Verd. 1929 Frullania ericoides (Nees) Mont. 1839 squarrosa var. (beta) Nees 1845 [pro syn] Typus: Wright Hep no 8 Basionym: Distribution: Nees in G., L. & N. 1845 2850 Syn Hepat: 417 [syn sub Fr ericoides] = Frullania ericoides var. ß laxa (Gottsche, Lindenb. et Nees) Schiffn. 1898 Page 395 squarrosa var. ericoides (Nees) Schiffn. 1899 Basionym: Distribution: Schiffner Stephani Frye & Clark Kamimura Bonner 1899 1910 1947 1961 1965 1510 686 204 199 655 Piippo 1990 521 = Frullania squarrosa Österr Bot Zeitschr 49 (11): 391 [Japan] Spec Hepat 4: 388 [not as syn sub Fr. squarrosa] Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] J Hatt Bot Lab 24: 17 [syn sub Fr squarrosa] Index Hepat 5: 433 [error: not as syn sub Fr. squarrosa in STEPH] [Brazil, Cuba, Java] J Hatt Bot Lab 68: 47 [annotated catalogue of China] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides (Nees) Mont. 1839 squarrosa var. gracilior Gottsche 1863 Typus: Mexico. Hacienda de Mirador Liebmann 493 Basionym: Distribution: Gottsche Gottsche = Frullania ericoides 1863 1867 103 104 Separata of Mex. Leverm.: 238 K Danske Vid Selks Skrifter, ser 5, 6: 334-335 [Mexico] (Nees) Mont. 1839 squarrosa var. laevior Nees 1847 Typus: Mexico: Cabrestra Leibold Basionym: Distribution: Nees in G., L. & N. Gottsche Gottsche Sim Bonner = Frullania ericoides 1847 1863 1867 1926 1965 2848 103 104 252 655 Syn Hepat: 772 Separata of Mex. Leverm.: 238 K Danske Vid Selks Skrifter, ser 5, 6: 334 [Mexico] Trans Roy Soc S Africa 15: 42, fig [South Africa] Index Hepat 5: 433 [Mexico] (Nees) Mont. 1839 squarrosa var. mülleri Gottsche 1863 Typus: Mexico. hb MÜLLER 2343 Basionym: Distribution: Gottsche Gottsche = Frullania ericoides 1863 1867 103 104 Separata of Mex. Leverm.: 238 K Danske Vid Selks Skrifter, ser 5, 6: 334-335 [Mexico] (Nees) Mont. 1839 Page 396 squarrosa var. planescens Verd. 1929 Basionym: Distribution: China, Indonesia (Java), Philippines (Luzon) var nov - comb nov comb nov Verdoorn Verdoorn Verdoorn Verdoorn in Handel-Mazzetti Verdoorn Verdoorn Grolle & Piippo Hattori Tan & Engel 1929 1930 1930 1930 1932 1937 1984 1984 1986 245 293 549 550 553 556 143 503 4286 Piippo Bapna & Kachroo 1990 2000 521 712 Ann Bryol 2: 134-136 Ann Bryol Suppl 1: 38-40, fig. 44 Nova Guinea 14 (4): 541 Symb Sinicae 5: 40 [Chian] Bull Jard Bot Buitenzorg, ser 3, 12 (1): 53-54 [Java] Blumea suppl 1: 212 [Philippine Islands] Acta Bot Fennica 125: 24 [Cat. West Melanesia] J Hatt Bot Lab 57: 412 [comb nov as Fr. ericoides var planescens] J. Hatt. Bot. Lab.. No. 60: 335 Frullania ericoides (Nees) Mont. var. planescens (Verd.) Hatt., comb. nov. J Hatt Bot Lab 68: 47 [annotated catalogue of China] Hepaticology in India II: 170 [syn sub Fr. ericoides] = Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides (Nees) Mont. 1839 squarrosa var. planescens fo. campanuloides Verd. 1930 Basionym: Distribution: fo nov - syn nov Verdoorn Verdoorn Hattori 1930 1932 1982 293 553 495 Ann Bryol Suppl 1: 38 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 54 [Java] J Hatt Bot Lab 51: 249 [syn sub Fr. ericoides var planescens] = Frullania ericoides var. planescens (Verd.) S.Hatt. 1984 Frullania ericoides (Nees) Mont. 1839 squarrosa var. planescens fo. chonanthelioides Verd. 1930 Basionym: Distribution: fo nov - syn nov Verdoorn Verdoorn Hattori Bapna & Kachroo = Frullania ericoides 1930 1932 1982 2000 293 553 495 712 Ann Bryol Suppl 1: 40 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 54 [Java] J Hatt Bot Lab 51: 249 Hepaticology in India II: 170 [syn sub Fr. ericoides] (Nees) Mont. 1839 squarrosa var. subjulacea Spruce 1888 [nom nud] Typus: Paraguay Basionym: Distribution: Paraguay - syn nov Spruce Reiner = Frullania ericoides 1888 1988 1062 Rev Bryol 15 (3): 34 [No 3712, 3723, 4247, Paraguay] 1043 Bol Soc Argentina Bot 25 (3-4): 316, 318 [sub Fr. ericoides, Buenos Aires] (Nees) Mont. 1839 Page 397 squarrosa var. verrucosa Kamim. 1961 Basionym: Distribution: var nov - syn nov Kamimura Hattori Bapna & Kachroo Yuzawa Yamada & Iwatsuki Mizutani, Hasegawa & Iwatsuki 1961 1982 2000 2001 2006 2009 - comb nov Hentschel et alii 2015 199 495 712 1660 1470 2112 J Hatt Bot Lab 24: 19, fig. 4(9-19) [var nov] J Hatt Bot Lab 51: 249 [syn nov sub Fr. ericoides] Hepaticology in India II: 170 [syn sub Fr. ericoides] Natural Environmental Sci Res 14: 13 [Japan, syn sub Fr ericoides] J Hatt Bot Lab 99: 32 [Catalog of Japan] Type sepecimens Hatt Bot Lab (NICH) 36-37: Isotype. Japan, Shikoku, Kochi-ken, Takaoka-machi, Hasuike-mura, Mt. Shiroyama. 20 Feb. 1948. N. Okubo 13215 (NICH 46849). 4097 Phytotaxa 220 (2): 134-135 [Frullania ericoides var. verrucosa (Kamim.) comb. nov.] = Frullania ericoides var. verrucosa (Kamim.) Hentschel & von Konrat 2015 squarrosa var. vulgaris Nees 1847 Typus: Mexico: Pico de Orizaba [formam communem] Cl. Liebmann Basionym: Distribution: Nees in G., L. & N. Gottsche Gottsche Sim Bonner = Frullania squarrosa 1847 1863 1867 1926 1965 2848 103 104 252 655 Syn Hepat: 772 Separata of Mex. Leverm.: 238 K Danske Vid Selks Skrifter, ser 5, 6: 334 [Mexico] Trans Roy Soc S Africa 15: 42 [South Africa] Index Hepat 5: 434 [Mexico] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides (Nees) Mont. 1839 squarrosa var. γ laxa Gottsche, Lindenb. & Nees 1847 [p. 772, Liebmann, Mexico] Typus: Mexico: Chiantla Liebmann Basionym: Distribution: Mexico var nov Gottsche, Lindenb. & Nees Gottsche Gottsche Sim Bonner 1845 1863 1867 1926 1965 2850 103 104 252 655 Syn Hepat: 772 [Liebmann, Mexico] Separata of Mex. Leverm.: 238 [Hacienda de Mirador] K Danske Vid Selks Skrifter, ser 5, 6: 334 [Mexico] Trans Roy Soc S Africa 15: 42 Index Hepat 5: 433 [squarrosa var. laxa: Mexico] = Frullania ericoides var. ß laxa (Gottsche, Lindenb. et Nees) Schiffn. 1898 squarrosa var. γ laxa Gottsche, Lindenb. & Nees 1847 [p. 417, Wight, India - pro syn.] Typus: in Peninsula Indiae orient Wight Basionym: Distribution: pro syn Gottsche, Lindenb. & Nees Sim 1845 1926 2850 Syn Hepat: 772 [Liebmann, Mexico] 252 Trans Roy Soc S Africa 15: 42 = Frullania ericoides var. ß laxa Gottsche, Lindenb. & Nees 1845 [p. 417, Wight, Indien] Frullania ericoides var. ß laxa (Gottsche, Lindenb. et Nees) Schiffn. 1898 Page 398 squarrulosa (Hooker & Taylor) Taylor by Carrington & Pearson 1888 [orth var] Basionym: Distribution: Carrington & Pearson Scott & Bradshaw Downing = Frullania squarrosula standleyi 1888 1986 1992 1604 Proc Linn Soc New South Wales, ser 2, 2 (4): 1037 56 Brunnonia 8 (1): 37 [Australia] 1138 Bryologist 95(1): 9-10 [Australia] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Herzog in hb US 1996 [nom herb] {Costa Rica} Typus: Costa Rica. Volcan de Turrialba, near Finca del Volcan de Turrialba 22.2.1924, Standley 35239 in hb US Basionym: Distribution: Costa Rica NN 1996 632 Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 686 655 185 928 Wigginton 2002 Wigginton 2004 Wigginton 2004 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 30 of 35 [in hb US] = staudtiana Steph. 1911 Typus: Cameroon. Lolodorf as, Staudt 307 in hb G Basionym: Distribution: Cameroon spec nov = Frullania apiculata stellatitexta Spec Hepat 4: 685-686 [sub sgen Frullania] Index Hepat 5: 435 Bull Jard Bot Nat Belg 46 (1/2): 13 [may be syn of Fr. apiculata] Stephani Icon Hepat Inedit [Thyopsiella] F- 28, microfiche 57 fig 3910 ["Kamerun, Staudt 307 ex parte, Lolodorf"] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [probably Fr apiculata] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apiculata, West Africa] (Reinw., Blume & Nees) Nees 1845 Steph. 1911 Typus: Fiji "Viti-Inseln". Qvalau, "Gebirgessstation" 1864, Dr. E. Graeffe 582 in hb G 18256 (ex hb JACK) Basionym: Distribution: spec nov Stephani Verdoorn Verdoorn Miller Bonner Hattori Miller, Whittier & Whittier Stephani 1911 1928 1930 1953 1965 1980 1983 1985 - syn nov Hattori Staples & Imada 1986 2006 686 242 551 349 655 490 609 928 Spec Hepat 4: 585-586 ["Insulae vitienses"] Rev Bryol NS 1: 115-116 Nederl Kruidk Arch Jrg 1930 (2): Bryologist 56: 48 [doubtfull in Hawaii] Index Hepat 5: 435 J Hatt Bot Lab 47: 187, 192-193, fig 16 Prod Flora Hepat Polynesiae: 139 Stephani Icon Hepat Inedit [Homotropantha] D-36, microfiche 54 fig 3759 ["Ins Viti: Ovalau, leg Graeffe 582 (auch Nova Caledonia, Lerat No 441)"] 506 J Hatt Bot Lab 60: 221 [syn sub Fr. intermedia var. non-apiculata] 1716 Trop bryol 28: 38 [Hawaii dubious record] = Frullania intermedia var. non-apiculata S.Hatt. 1975 Page 399 stephanii Schiffn. 1893 Typus: San Thome Isl. 1888, Quintas in hb G 17325 Basionym: Distribution: W Africa, San Thome Isl spec nov - syn nov? Schiffner Schiffner 1893 1893 Stephani Bonner Vanden Berghen Stephani 1910 1965 1976 1985 Wigginton 2002 Wigginton 2004 Wigginton 2004 Reprint: 10 Nova Acta K Leop-Carol Deutsch Akad Naturf Halle 60: 226-227, pl 7(1-22) 686 Spec Hepat 4: 381 655 Index Hepat 5: 435 185 Bull Jard Bot Nat Belg 46 (1/2): 185 928 Stephani Icon Hepat Inedit [Galeiloba] C- 39, microfiche 50 fig 3391 [from type; "S. Thomé Fr Quintas, Copie nach Schiffner"] - 3392 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [perhaps Fr caffraria] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 229 [perhaps syn sub Fr caffraria, West Africa] Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 686 655 185 928 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania caffraria 572 Steph. 1894 stipatiflora Steph. 1911 Typus: Reunion sd, Dusen sn hb STEPHANI 47, in hb G 10714 Basionym: Distribution: Reunion Isl spec nov - syn nov = Frullania apicalis stipatiloba Mitt. 1879 fo. bicornuta (Steph.) Herzog 1942 [nom inval] Basionym: Frullania bicornuta Distribution: Steph. 1911 Herzog in Skottsberg = Frullania stipatiloba striata Spec Hepat 4: 483-484 Index Hepat 5: 436 Bull Jard Bot Nat Belg 46 (1/2): 52 Stephani Icon Hepat Inedit [Thyopsiella] F- 29, microfiche 57 fig 3911 ["Africa, Réunion, Dusén legit"] 250 Bryophyt Bibl 48: 62 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 217 [syn sub Fr apicalis, West Africa] 1942 722 Nat Hist Juan Fernandez & Easter Island, 2: 740 Steph. 1894 (Lehm. & Lindenb. in Lehm.) Mont. 1842 Basionym: Distribution: comb nov Montagne = Ptychanthus striatus 1842 469 Ann Sci Nat Bot, ser 2, 18: 13 (Lehm. & Lindenb. in Lehm.) Nees 1844 Page 400 stricta (F. Weber) Lindenb. 1845 Basionym: Jungermannia atrata var. stricta Weber 1815 Distribution: Magadascar, Reunion Isl, Comores Lindenberg in G., L. & N. Pearson 1845 1891 Stephani in Renauld & Cardot Stephani Gola Bonner Vanden Berghen Stephani 1891 1911 1916 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania serrata strobilantha 2850 Syn Hepat: 453 1921 Christiania Videnskabs-Selskabs Forhandlinger 1890 (2): 5-6, pl 2(121) [Madagascar] 1453 Rev Bryol 18 (4): 56 [Madagascar] 686 Spec Hepat 4: 478-479 3469 Mem Reale Accad Sci Torino, ser 2. 65: 9 [Kenya] 655 Index Hepat 5: 436 185 Bull Jard Bot Nat Belg 46 (1/2): 111 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 14, microfiche 56 fig 3889 ["Bourbon, Renauld 34.32"] - 3890 250 Bryophyt Bibl 48: 68 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 221 [syn sub Fr serrata, West Africa] Gottsche 1845 Spruce 1884 Typus: Peru. Campana, "flumen Pastasa infra Banos, 1500m" "6°26'S 78°23'W" sd, Spruce sn in hb US in hb E E00256431 - E00256432 - E00002586 [isosyntypus] Basionym: Distribution: Panama, Peru, Ecuador spec nov - syn nov - syn nov Spruce Stephani Clarl 1884 1911 1955 Bonner Stotler Stotler Menzel Stephani 1965 1969 1970 1984 1985 NN 1996 = Frullania macrocephala stuebeli 243 686 371 Trans Proc Bot Soc Edinburgh 15: 44 & 48 Spec Hepat 4: 514 Bryologist 58: 146-149, fig 1-20 [difference to Fr. intumescens, mollicula, magniflora var wrightii] 655 Index Hepat 5: 436 4761 Nova Hedwigia 18: 439 [syn sub Fr. macrocephala] 194 The genus Frullania, separata: 439 [syn sub Fr. macrocephala] 142 Willdenowia 14: 510 [syn of Fr. macrocephala] 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 92, microfiche 58 fig 4010 ["Andes peruviani, Campana, leg Spruce, Original"] - 4011 ["ex planta ecuadorensis Levier 6516"] 632 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 31 of 35 [in hb US] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. ex Steph. 1985 [nom herb] {Colombia} Typus: in hb G Basionym: Distribution: Colombia Stephani 1985 928 Stephani Icon Hepat Inedit [Thyopsiella] F- 93, microfiche 58 fig 4012 [Columbia (Stübel legit)"] = Page 401 stylata Gerola 1947 Typus: Ethiopia. Sciasciamanne 26.3.1938, Vatova Basionym: Distribution: Abyssinia spec nov Gerola 1947 Cufodontis Bonner Vanden Berghen Wigginton 1952 1965 1976 2002 Wigginton 2004 = Frullania trinervis subaculeata Typus: Peru. 1532 Lavori di botanica / Istituto Botanica dell'Università di Padova 12: 475-476, fig 7(a-d) [Ethiopia] 2562 Phyton 4 (1-3): 77 [Ethiopia] 655 Index Hepat 5: 436 185 Bull Jard Bot Nat Belg 46 (1/2): 13 [syn sub Fr. trinervis] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [perhaps Fr trinervis] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] (Lehm.) Drège 1843 Spruce 1890 sd, Lechler 2550 in hb MANCH 17176 [holotypus] Basionym: Distribution: Peru, Ecuador, Bolivia spec nov Spruce 1890 Stephani Stephani Herzog Arnell Bonner Menzel Stephani 1911 1916 1952 1962 1965 1984 1985 Uribe & Gradstein Uribe 2003 2008 = Frullania atrata subalpina 1122 Bull Soc Bot France 36 (Suppl) [Congres de Botanique]: CCII-CCIII [202-203] 686 Spec Hepat 4: 605 222 Biblioth Bot 87: 247 [Bolivia] 444 Svensk Bot Tidskr 46 (1): 86 [Ecuador] 449 Svensk Bot Tidskr 56 (2): 341 [Ecuador] 655 Index Hepat 5: 437 142 Willdenowia 14: 490 [Peru] 928 Stephani Icon Hepat Inedit [Meteriopsis] E-38, microfiche 55 fig 3823 [Ecuador, Allioni 1909, Levier 6415"] - 3824 1071 Crypt Bryol 24 (3): 196 [syn sub Fr atrata; published in 1890] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] (Sw.) Nees ex Mont. 1839 Spruce 1884 Typus: Ecuador: Mt. Pichincho, suora pagum Nono, 3350 m sd, Spruce Basionym: Distribution: Mexico, Ecuador spec nov - syn nov Spruce Stephani Stephani Bonner Stephani 1884 1910 1910 1965 1985 243 686 686 655 928 Fulford & Sharp 1990 105 = Frullania pluricarinata Trans Proc Bot Soc Edinburgh 15: 14 & 28-29 (No 17) Spec Hepat 4: 341 [syn sub Fr. pluricarinata] Spec Hepat 4: 346 Index Hepat 5: 437 Stephani Icon Hepat Inedit [Chonanthelia] A- 77, microfiche 47 fig 3179 ["Bogota Lindig - Fr pluricarinata identisch mit Fr subalpina, G. hat Priorität"] Mem NY Bot Gard 63: 46 [Mexico] Gottsche 1864 Page 402 subatrata Steph. 1911 Typus: 1. St Thomé 2. Saint Thomas 1. 1885, Newton 2. sd, Moller sn in hb G sn; Stephani, Icones Hepat Inedit 1631 Basionym: Distribution: spec nov - syn nov Stephani Demaret & Vanden Berghen Bonner Stephani 1911 1950 1965 1985 686 60 655 928 Wigginton 2002 1012 Wigginton Wigginton Uribe 2004 2004 2008 1312 1172 1513 = Frullania angulata subcommutata Spec Hepat 4: 592-593 Bull Jard Bot Etat Bruxelles 20: 5 [syn sub Fr. angulata] Index Hepat 5: 437 Stephani Icon Hepat Inedit [Meteriopsis] E- 8, microfiche 55 fig 3774 ["Africa occ, Ins San Thomé (Newton leg) 1885)"] - 3775 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Mitt. 1863 Steph. 1908 Typus: Samoa, Insel Upolu, Mt. Lanutoo, 700m -.8.1905, K. & L. Rechinger, 5401 in hb G Basionym: Distribution: spec nov spec nov - syn nov Stephani Stephani Stephani Verdoorn Verdoorn Bonner Grolle & Schultze-Motel Hattori Miller, Whittier & Whittier Stephani = Frullania angulosa 1908 1915 1924 1928 1930 1965 1972 1980 1983 1985 216 221 687 242 551 655 537 490 609 928 Denkschr K Akad Wiss, Math Nat Kl, Wien 81: 298 Denkschr K Akad Wiss, Math Nat Kl, Wien 91: 166 Spec Hepat 6: 551 [as spec nov] Rev Bryol NS 1: 121 [syn sub Fr. angulosa] Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 437-438 J Hatt Bot Lab 36: 83 [sub Fr. angulosa] [Samoa] J Hatt Bot Lab 47: 193 [syn sub Fr. intermedia] Prod Flora Hepat Polynesiae: 139 Stephani Icon Hepat Inedit [Homotropantha] D-11, microfiche 54 fig 3718 ["Samoa, Insula Upolu, Rechinger legit"] Mitt. in Seemann 1871 Frullania intermedia var. non-apiculata S.Hatt. 1975 subcordistipula Typus: Mexico Steph. 1924 sd, Arsén 7802 & 7808 in hb G 20977 (= 7802) [lectotypus by YUZAWA 1984]; Basionym: Distribution: Mexico spec nov - syn nov Stephani Bonner Yuzawa 1924 1965 1984 687 655 502 Stephani 1985 928 Fulford & Sharp Yuzawa 1990 1991 105 525 = Frullania cuencensis Spec Hepat 6: 534-535 Index Hepat 5: 438 J Hatt Bot Lab 57: 398-399 (No 12), fig. 12 (a-n) [as female plant ev. conspecific with Fr. cuencensis] Stephani Icon Hepat Inedit [Chonanthelia] A- 83, microfiche 47 fig 3188 [from type; "Mexico Arsén"] Mem NY Bot Gard 63: 46 [Mexico] J Hatt Bot Lab 70: 209 Taylor 1846 Page 403 subcrenulata Schiffn. ex Verd. 1930 [pro syn] Basionym: Distribution: Verdoorn = Frullania claviloba subcylindrica 1930 293 Ann Bryol Suppl 1: 120 Steph. 1911 Steph. 1922 Typus: Brazil. Brasilia, Rio Grande sd, Lindmann B120 pp. in hb G Basionym: Distribution: spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 = Frullania brasiliensis 686 655 4761 194 928 Spec Hepat 4: 506-507 Index Hepat 5: 438 Nova Hedwigia 18: 436 The genus Frullania, separata: 436 Stephani Icon Hepat Inedit [Thyopsiella] F- 94, microfiche 58 fig 4013 ["Rio Grande, Lindman B 120 ex parte"] Raddi 1822 subdentata ssp. solomonensis [nom herb in hb L L0060944] Typus: Solomon Islands. Guadalcanal, Popemanaseu Mt. "British Solomon Isl.: Guadalcanal, Mt. Popemanaseu, just below summit area. On exposed tree-branch in subalpine shrubbery." 30/9/1968, Zanten, B.O. van 68-2730/b in hb L L0060944 [typus] - http://145.18.162.53:81/c8?ent=300017&rec=69102&sct=1 Basionym: Distribution: = subdeplanata Steph. 1910 Typus: New Zealand. sd, Colenso 977 in hb G Basionym: Distribution: Antipode Isl [New Zealand] spec nov Stephani Hodgson & Sainsbury 1910 1948 Hodgson Bonner Hamlin Stephani 1949 1965 1972 1985 Godley 1989 = Frullania scandens 686 885 Spec Hepat 4: 408 Svensk Bot Tidskr 42 (3): 277-278 [Antipode Isl] [probably syn sub Fr. scandens] 203 Trans Roy Soc New Zealand 77 (3): 381 655 Index Hepat 5: 438 659 Rec Dominion Mus 7(19): 263 [New Zealand] 928 Stephani Icon Hepat Inedit [Galeiloba] C-274, microfiche 54 fig 3702 ["(Colenso legit) n Zealand"] 1334 New Zealand J Bot 27: 537 [Antipode Isl] Mont. 1843 Page 404 subhampeana E.A.Hodgs. 1949 Typus: New Zealand. Maungapoike Falls, Wairoa .3.1947, A.P.Druca 7408* Kinneroa, near Woodville* -.12.1937, E.A. Hodgson 1472 in hb MPN [lectotypus], in hb MPN [*=syntypus] Basionym: Distribution: Australia (NSW, QLD), New Zealand Hodgson Bonner Hamlin Hodgson Hamlin Hattori Hattori Hattori Scott & Bradshaw Windolf Hattori Glenny Carthy Glenny, Meagher & Brown 1949 1965 1972 1972 1973 1979 1983 1984 1986 1987 1988 1998 2003 2004 203 655 659 3468 656 486 499 682 56 1025 429 1412 1125 1126 Glenny, Meagher & Brown 2004 1127 Glenny & Malcolm 2005 1410 McCarthy Hentschel et von Konrat 2006 2015 1385 4097 Trans Roy Soc New zealand 77 (3): 370-371, pl 37(7) Index Hepat 5: 439 Rec Dominion Mus 7(19): 263 [New Zealand] J Roy Soc New Zealand 2 (1): 110 [syn sub Fr novae-zelandiae] Rec Dominian Mus 8 (10): 143 [New Zealand] J Hatt Bot Lab 46: 145, fig 44(a-d) from lectotype J Hatt Bot Lab 54: 178-180, fig 71 Crypt Bryol Lichen 5(1-2): 187 [Australia, QLD] Brunnonia 8 (1): 37 & 145 [Australia: NSW] Austrobaileya 2(4): 391 [checklist Queensland] Nova Hedwigia Beih 90: 155 [Australia: QLD, NSW] Tuhinga 10: 132 [revised checklist of New Zealand] Flora Australia Suppl Ser 21: 47 [Australia (NSW, QLD)] NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand species] Checklist Australian Liverworts [Australia QLD NSW] Phytotaxa 220 (2): 136, 2015, Bas.: Frullania subhampeana E.A.Hodgs = Frullania monocera var. subhampeana (E.A.Hodgs.) Hentschel & von Konrat 2015 subinflata Mitt. 1861 Typus: India. Madras,in caespite Parmeliae leucomelae sd, collector unspezified 1530 in hb NY 1048735 (hb W. MITTEN; with drawing) , hb K 1530 Basionym: Distribution: spec nov - syn nov Mitten Stephani Stephani Verdoorn Verdoorn Chopra Parihar 1861 1910 1911 1929 1930 1943 1962 Bonner Hattori Hattori 1965 1974 1975 Hattori Miller, Whittier & Whittier Stephani 1975 1983 1985 Parihar, Lal & Katiyar Bapna & Kachroo Singh, Kumar & Nath 1994 2000 2008 = Frullania polyptera 574 686 686 245 293 205 1584 J Linn Soc Bot 5: 119 [India: Madras] Spec Hepat 4: 443 [syn sub Fr. grandistipula] Spec Hepat 4: 677-678 [as own species] Ann Bryol 2: 125 [syn sub Fr. grandistipula] Ann Bryol Suppl 1: 43 [syn sub Fr. grandistipula] J Indian Bot Soc 22: 249 [India] Univ Allahabad studies, Bot sect 1-2: 18 [India; syn sub Fr grandistipula] 655 Index Hepat 5: 439 545 J Hatt Bot Lab 38: 208-211, fig 99 [syn nov sub Fr. polyptera] 47 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 81 [Ann Bryol out Mitten´s specimen from India] 47 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (2): 81 609 Prod Flora Hepat Polynesiae: 139 928 Stephani Icon Hepat Inedit [Galeiloba] C-138, microfiche 52 fig 3531 [syn mit Fr gandistipula Ldg " stimmt mit dem Original aus Madras ganz überein, Ceylon, Nietner 1556c"] - 3532 [" Java, Hasskarl (Copie nach Gottsche) Syn p 430" = Fr grandstipula] 2196 Hepat Anth India, annotated checklist: 22 [South India] 712 Hepaticology in India II: 178 [syn sub Fr. polyptera] 1281 Taiwania 53 (1): 79, [syn sub Fr grandistipula] Taylor 1846 Page 405 - subnigra Steph. 1924 Typus: Japan: Ise, Isegi 31.12.1907, Uematsu 114 in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Frye & Clark Hattori Kamimura Stephani 1924 1930 1947 1952 1961 1985 Bapna & Kachroo Yamada & Iwatsuki 2000 2006 = Frullania squarrosa Spec Hepat 6: 544 Ann Bryol Suppl 1: 34 Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] J Hatt Bot Lab 8: 31 J Hatt Bot Lab 24: 17 [syn sub Fr squarrosa] Stephani Icon Hepat Inedit [Galeiloba] C-252, microfiche 53 fig 3672 ["Japonia, leg Uematsu 1907, No 114"] 712 Hepaticology in India II: 170 [syn sub Fr. ericoides] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides subplana 687 293 204 529 199 928 (Nees) Mont. 1839 Gottsche in Engler 1895 [nom nud] Basionym: Distribution: nom nud Stephani in Engler 1895 Vanden Berghen Stephani 1976 1985 = Frullania subplana Gottsche ex Steph. 1910 Frullania bullata subplana 1481 Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [Usambara - Hochwald] [nom nud, vide Vanden Berghen 1976] 185 Bull Jard Bot Nat Belg 46 (1/2): 186 928 Stephani Icon Hepat Inedit [Galeiloba] C- 40ab, microfiche 50 fig 3393 ["Madagascar Betsileo, leg Rev Montant 1893, Herb Renauld 372 1891?"] - 3394 ["Frullania Borgenii Gms (nomen occupatum), Copie nach Gottsche, Madagascar"] - 3395 ["Madagascar, Borgen"] Steph. 1910 Gottsche ex Steph. 1910 Typus: Madagascar. Betisleo 1893, Montaut hb RENAULD 372, in hb G 13102 & Basionym: Distribution: Madagascar spec nov - syn nov Gottsche ex Stephani Bonner Vanden Berghen Grolle Wigginton 1910 1965 1976 1995 2002 Wigginton 2004 = Frullania bullata 686 655 185 250 1012 Spec Hepat 4: 383 Index Hepat 5: 439-440 Bull Jard Bot Nat Belg 46 (1/2): 186 Bryophyt Bibl 48: 64 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Steph. 1910 Page 406 subrostrata S.Hatt. 1972 [nom inval (Art 37.1)] Basionym: Neohattoria rostrata Distribution: New Zealand nom nov syn nov syn nov R.M.Schust. 1970 [nom inval (Art 37.1)] Hattori Hattori & Mizutani Glenny Konrat & Brennan 1972 1982 1998 2004 Glenny & Malcolm 2005 Konrat, Hentschel, Heinrichs, Braggins & Pócs 2010 539 495 1412 1164 J Hatt Bot Lab 36: 443 J Hatt Bot Lab 52: 446 Tuhinga 10: 132 [sub Fr aterrima var rostrata] http://www.liverworts.org/dich_key/frul_key.shtml, 6th of 6 pages [Key to Frullania of New Zealand; throughout] 1410 ABRS Ident Ser, Key Australasian Liverwort & Hornwort Genera - CDRom [New Zealand, sub Fr aterrima var rostrata] 2377 Nova Hedwigia 91(3-4): 482 [nom inval (Art 37.1)] = Frullania aterrima var. rostrata (R.M.Schust.) S.Hatt. 1983 [nom inval (Art 37.1)] Frullania aterrima var. lepida E.A.Hodgs. 1949 substricta Steph. 1913 Typus: Tansania. Usambara, Lutindi Mt, 1400m -.8.1909, Brunnthaler in hb G (sub Fr. holstii) Basionym: Distribution: Tanzania spec nov Stephani in Brunnthaler Bonner Vanden Berghen Stephani 1913 1965 1976 1985 Wigginton 2004 = Frullania angulata subtilis 219 655 185 928 Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 729 [Tanzania] Index Hepat 5: 440 Bull Jard Bot Nat Belg 46 (1/2): 13 [may be syn of Fr. angulata] Stephani Icon Hepat Inedit [Thyopsiella] F- 6b, microfiche 56 fig 3878 ["substricta, Africa, Usambara, Brunthaler 1909"] - 3879 1312 Trop Bryol Res Rep 5: 32 [prpbably syn of Fr angulata] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Mitt. 1863 Steph. 1911 Typus: Papua New Guinea. Northern, Mt. Suckling sd, Mc Gregor in hb G 14655 [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Hattori Hattori & Mizutani Hattori Grolle & Piippo 1911 1930 1930 1965 1973 1982 1982 1984 Stephani 1985 Yang J-D 2016 686 293 549 655 540 497 495 143 Spec Hepat 4: 647 [Fr. subtilis] Ann Bryol Suppl 1: 136-138, fig 220 [asFr. subtilis] Nova Guinea 14 (4): 545 [asFr. subtilis] Index Hepat 5: 440 [as Fr. subtilis] J Hatt Bot Lab 37: 58-59, f 24(a-f) [as Fr subtilis] J Hatt Bot Lab 52: 447 [sub Fr junghuhniana var. minutissima] J Hatt Bot Lab 51: 233 [sub Fr junghuhniana var. minutissima] Acta Bot Fennica 125: 24 [Cat. West Melanesia] [= F. junghuhniana var. tenella] 928 Stephani Icon Hepat Inedit [Diastaloba] B-83, microfiche 49 fig 3330 [from type; "N Guinea, Mt Suckling, Sin d Macgregor"] as Fr. subtilis 4541 BryoString 4: 1-6 (Taiwan, as Fr. subtilis] = Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 Page 407 subtilissima [auct. non Lindenb. 1845] Clark & Svihla 1948 Basionym: Distribution: Clark & Svihla 1948 325 Gradstein & Hekking 1989 517 Bryologist 51: 23-24 [key to Fr. ofMiddle America] [may be syn with Fr. atrata; they examined a misidentified specimen, belonging to Sgen. Meteoriopsis (Gradstein & Hekking )] J Hatt Bot Lab 66: 209 = sullivantiae Austin 1872 Typus: USA. Ohio, Urbana, cedar swamp sd, s coll ex hb AUSTIN, in hb G Basionym: Distribution: spec nov - syn nov Austin Evans 1872 1897 666 237 Stephani Frye & Clark Stephani 1911 1947 1985 686 204 928 = Frullania selwyniana sullivantii Bull Torrey Bot Club 3 [No 3]: 16 Trans Connet Acad Arts Sci 10 (1): 29-30 [preferred selwyniana, because sullivantiae sounds like sullivantii] Spec Hepat 4: 568 & 669 [as syn sub Fr. selwyniana] Univ Wash Publ Biol 6 (5): 755 [sub Fr selwyniana] Stephani Icon Hepat Inedit [Thyopsiella] F-200a, microfiche 60 fig 4162 ["olim Frull Sullivantiae Aust, Cedar Swamp near Urbana ot 1844, syn Frull selwyniana Pears Canad Hep 1890"] Pearson 1890 Austin 1869 Typus: USA. Georgia sd, Sullivant Basionym: Distribution: N-America spec nov spec nov Austin Austin Evans Stephani Frye & Clark Stephani 1869 1870 1897 1910 1947 1985 1339 237 686 204 928 - syn nov Schuster 1992 603 = Frullania eboracensis ssp. virginica sundaica Proc Acad Nat Sci Philadelphia 1869 (Separata): 9 Proc Acad Nat Sci Philadelphia 21: 226 Trans Connet Acad Arts Sci 10 (1): 17-18 [syn sub Fr. virginica] Spec Hepat 4: 431 [syn sub Fr. eboracensis] Univ Wash Publ Biol 6 (5): 771 [sub Fr eboracensis] Stephani Icon Hepat Inedit [Galeiloba] C- 86abc, microfiche 51 fig 3459 ["South Carolina, Curtis, herb Lammig, Pearson dedit Hepat N America 5: 147 [syn sub Fr. eboracensis ssp. virginica] (Gottsche in Lehm.) R.M.Schust. 1992 Steph. 1911 Typus: Indonesia. Sumatra, Panerango sd, scoll in hb G 17194 Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Bonner Hattori Miller, Whittier & Whittier Stephani = Frullania junghuhniana 1911 1930 1965 1976 1983 1985 686 293 655 67 609 928 Spec Hepat 4: 644 Ann Bryol Suppl 1: 134 [syn sub Fr. junghuhniana] Index Hepat 5: 441 Bull Natn Sci Mus Tokyo, ser B (Bot), 2 (1): 21-22 Prod Flora Hepat Polynesiae: 139 Stephani Icon Hepat Inedit [Diastaloba] B-84, microfiche 49 fig 3331 [from type; "Sumatra"] Gottsche 1845 Page 408 supradecomposita (L. & L.) Nees 1838 [nom inval, Art. 32.1(c)] Basionym: Distribution: Nees 1838 = Frullania supradecomposita surinamensis 261 Naturg Europ Leberm 3: 210 [nomen in footnote] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Tayler ex Gottsche, Lindenb. & Nees 1845 [pro syn] Basionym: Distribution: Taylor in G., L. & N. = Frullania arietina 1845 Taylor 1845 Frullania rio-janeirensis surinamensis Typus: Surinam 2850 Syn Hepat: 413 [syn sub Fr. arientina] (Raddi) Ångstr. 1876 Steph. 1911 sd, coll. ign. hb L 205, in hb G 23576 [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Bonner Stephani 1911 1965 1985 686 655 928 Gradstein & Hekking Yuzawa 1989 1991 517 525 = Frullania rio-janeirensis suringarii Spec Hepat 4: 674 Index Hepat 5: 442 Stephani Icon Hepat Inedit [Chonanthelia] A- 84, microfiche 47 fig 3189 [from type; "Surinam Herb Leiden 205"] J Hatt Bot Lab 66: 208 & 221 [syn sub Fr. gibbosa] J Hatt Bot Lab 70: 254 [syn sub Fr. riojaneirensis] (Raddi) Ångstr. 1876 Steph. ex Bonner 1965 [nom herb] Typus: Guiana: Paramaribo 1885, Suringar sn in hb G 23575 Basionym: Distribution: nom herb Bonner Stephani 1965 1985 655 928 - syn nov Gradstein & Hekking 1989 517 = Frullania ericoides sylevestris Index Hepat 5: 442 [Dutch Guiana] Stephani Icon Hepat Inedit [Galeiloba] C- 78, microfiche 51 fig 3444 ["from type; "publicanda, ist wohl besser zu Chonanthelia zu stellen, Paramaribo, Suringar 1885"] J Hatt Bot Lab 66: 2008 & 221 [nom. nudum, syn nov sub Fr. ericoides] (Nees) Mont. 1839 Steph. by Bapna & Kachroo 2000 [orth var] Basionym: Distribution: Bapna & Kachroo = Frullania ericoides 2000 712 Hepaticology in India II: 170 [as "sylevestris", syn sub Fr. ericoides] (Nees) Mont. 1839 Page 409 symmetrica Steph. 1908 Typus: New Caledonia: Foret du Col d´Annieu. -/2/1907, Le Rat 211 in hb G [holotypus] in hb L L0060946 [isotypus] Basionym: Distribution: spec nov - syn nov Stephani Paris Paris Stephani Verdoorn Verdoorn Frye & Clark Bonner Miller, Whittier & Whittier Stephani 1908 1910 1910 1911 1930 1930 1947 1965 1983 1985 870 911 1455 686 293 551 204 655 609 928 Bapna & Kachroo 2000 712 = Frullania squarrosa (Reinw., Blume & Nees) Nees 1845 Frullania ericoides tahitensis Rev Bryol 35: 29-30 Rev Bryol 37 (1): 41 [New Caledonia: Isle of Pines] Rev Bryol 37 (6): 130 [New Caledonia: Dogny] Spec Hepat 4: 462-463 [spec nov] Ann Bryol Suppl 1: 34 [syn sub Fr. squarrosa] Nederl Kruidk Arch Jrg 1930 (2): Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] Index Hepat 5: Prod Flora Hepat Polynesiae: 139 Stephani Icon Hepat Inedit [Galeiloba] C-188, microfiche 52 fig 3590 ["N Caledonia, Lerat"] Hepaticology in India II: 170 [syn sub Fr. ericoides] (Nees) Mont. 1839 Steph. 1911 Typus: Society Isl. Thaiti 1847, Vesco ex hb PC, hb STEPHANI 18, in hb G Basionym: Distribution: Society Isl (Tahiti), Marquesas spec nov - syn nov Stephani Verdoorn Verdoorn Bonner Miller, Whittier & Whittier Stephani = Frullania serrata taiheizana 1911 1930 1930 1965 1983 1985 686 293 551 655 609 928 Spec Hepat 4: 548 Ann Bryol Suppl 1: 85 Nederl Kruidk Arch Jrg 1930 (2): Index Hepat 5: 442 Prod Flora Hepat Polynesiae: 139 Stephani Icon Hepat Inedit [Thyopsiella] F-156, microfiche 59 fig 4095 ["Tahiti (leg Nadeaud)"] - 4096 Gottsche 1845 [auct. non Horikawa 1934] Kamim. 1952 {Japan} Basionym: Distribution: Kamimura Kamimura Choe -syn nov = Frullania taradakensis takayensis 1952 1961 1980 873 Contrib Fl Hepat Shikokuensem, Kochi: 132 199 J Hatt Bot Lab 24: 47 [syn sub Fr taradakensis] 1171 Illust Fl & Fauna Korea 24: 735 [South Korea] Steph. 1910 Steph. by Grolle 1970 [orth var] Basionym: Distribution: Grolle = Frullania takayuensis 1970 615 Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 313 [orth error] Steph. 1910 Frullania oakesiana ssp. takayuensis (Steph.) R.M.Schust. 1992 Page 410 takayuensis Steph. 1910 Typus: Japan. Yamagata Pref., Prov. Tosa, Takayu in hb G [holotypus], in hb KYO, hb TNS [isotypus] Basionym: Frullania oakesiana ssp. takayuensis Distribution: Japan spec nov - syn nov (Steph.) R.M.Schust. 1992 Sawada 1909 Stephani Iishiba Hattori Hattori Hattori & Noguchi Kamimura Bonner Hattori Yuzawa & Hattori Stephani 1910 1912 1953 1959 1960 1961 1965 1974 1981 1985 Inoue Schuster Casa, Brugues & Sergio N.N. [National Science Museum Tokyo] Yuzawa 1987 1992 1999 2000 Yamada & Iwatsuki Mamontov et al. 2006 2020 2001 = Frullania oakesiana ssp. takayuensis takayuensis 24.6.1904, U. Faurie 1443 930 Bot Mag Tokyo 23 (no 268): 178-183, fig 18, 18' [key to Japanese species, in Japanese] 686 Spec Hepat 4: 399 888 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] 530 J Hatt Bot Lab 10: 37-38, fig 48(a-f) 857 J Hatt Bot Lab 21: 128-129 [Japan] 965 Index spec Typ herb Japon, Bryophyta: 18 [isotype] 199 J Hatt Bot Lab 24: 84-85, fig 29(25-30), map 9. [Japan] 655 Index Hepat 5: 443 545 J Hatt Bot Lab 38: 203 [syn nov of Fr. oakesiana]} 494 J Hatt Bot Lab 50: 256 [syn of Fr. oakesiana] 928 Stephani Icon Hepat Inedit [Galeiloba] C-253, microfiche 53 fig 3673 ["Japan, Takayu, 24/6. 1904, Faurie 1443"] 1155 Index type specimens in Nat Sci Mus, Tokyo: 14 603 Hepat N America 5: 195 [as ssp of Fr. oakesiana] 1279 Crypt Bryol 20(3): 205 [Japan, sub Fr oakesiana] 995 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 1660 Natural Environmental Sci Res 14: 28 [Japan, basionym of Fr oakesiana ssp takayuensis] 1470 J Hatt Bot Lab 99: 32 [Catalog of Japan] 5641 Nova Hedwigia, Beiheft 150, p. 201-242 [belong to subg. Frullaniopsis subgen. nov.] (Steph.) R.M.Schust. 1992 Steph. ex Sawada 1909 [nom nud] Basionym: Distribution: Sawada = Frullania takayuensis 1909 930 Steph. 1910 Frullania oakesiana ssp. takayuensis tamarisci Bot Mag Tokyo 23 (no 268): 178-183, fig 18, 18' [key to Japanese species, in Japanese] (Steph.) R.M.Schust. 1992 [auct. non (L.) DUM] ..... ? S.Hatt. 1972 {quod plant. americae pacificae boreal} Basionym: Distribution: Hattori = Frullania nisquallensis tamarisci 1972 536 J Hatt Bot Lab 35: 235 Sull. 1849 Bol. 1870 Basionym: Distribution: - syn nov Bolander Evans Frye & Clark = Frullania californica 1870 1897 1947 3472 California Medical Gazette2: 184 237 Trans Connet Acad Arts Sci 10 (1): 25-26 204 Univ Wash Publ Biol 6 (5): 745 [sub Fr californica] (M.Howe) A.Evans 1897 Page 411 tamarisci [auct. non (L.) DUM.] Mont. 1840 Typus: in hb PC-MONT; Leprieur 260 Basionym: Distribution: Montagne Gradstein & Hekking = Frullania obcordata tamarisci 1840 1989 467 517 Ann Sci Nat Bot, ser 2, 14: 333 J Hatt Bot Lab 66: 209 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 [auct. non (L.) DUM] C.Massal. 1897 Basionym: Distribution: - syn nov Massalongo Levier Verdoorn 1897 1906 1934 = Frullania moniliata ssp. breviramea 1914 Mem Accad Agr Art Comm Verona, ser 3, 73 (2): 43 824 Nuovo Giorn Bot Ital NS 13: 350 240 Ann Bryol Suppl 4: 193 (Steph.) Verd. 1930 {Java, Sumatra} Frullania tamarisci ssp. obscura var. breviramea (Steph.) S.Hatt. 1972 Frullania moniliata tamarisci (Reinw., Blume & Nees) Mont. 1842 [auct. non (L.) DUM] Lehnert 1886 Basionym: Distribution: Canada, USA Lehnert Pearson Haynes Macoun Evans Evans Brown Hattori 1886 1890 1901 1902 1907 1908 1936 1972 = Frullania tamarisci ssp. asagrayana Frullania asagrayana 2089 Proceed Biol Soc Washington 3: 117 [Washington andd vicinity] 1153 List of Canadian Hepaticae: 3 [from Newfoundland] [from Maine] 1150 Catalogue Canadian Plants, VII: 47 [from Newfoundland] [from Conneticut] [from Massachusetts] [from Nova Scotia] 536 J Hatt Bot Lab 35: 232 (Mont.) S.Hatt. in Hara 1966 Mont. 1842 Page 412 tamarisci [auct. non (L.) Dum.] Sande Lac. 1864 {Japan} Basionym: Distribution: India (Eastern Himalayas: Megalaya) - syn nov Sande Lacoste Mitten Massalongo Stephani Levier Iishiba Iishiba Hattori Horikawa 1864 1891 1897 1897 1906 1909 1912 1944 1951 Hattori & Lin Singh, Kumar & Nath 1985 2008 = Frullania moniliata ssp. obscura 842 582 1914 213 824 890 888 943 1698 Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 313 [Japan] Trans Linn Soc London, ser 2, 3: 202 [Japan] Mem Accad Agr Art Comm Verona, ser 3, 73 (2): 43 Bull Herb Boissier 5: 79 [Japan: Tosa] Nuovo Giorn Bot Ital NS 13: 350 Bot Mag Tokyo 23: 363 [Japan] Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Bull Tokyo Sci Mus 11: 148 [Japan, key to Japanese species] J Sci Hiroshima Univ, serB, div 2 (Bot) 6(1): 28 [sub Fr moniliata ssp obscura] 505 J Hatt Bot Lab 59: 161 1281 Taiwania 53 (1): 60-62, fig 1[map], 4(1-12) [India, Meghalaya, Eastern Himalayas] Verd. 1930 {nördl. Indomalaya} Frullania tamarisci ssp. obscura (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata tamarisci (Reinw., Blume & Nees) Mont. 1842 fo. alpina Verd. ex Piippo 1990 [pro syn] Basionym: Distribution: Piippo 1990 = Frullania moniliata tamarisci 521 J Hatt Bot Lab 68: 48 [annotated catalogue of China] (Reinw., Blume & Nees) Mont. 1842 fo. irrigata Schiffn. 1936 Typus: Norway. Söndre Trandhjemsamt Opdal, Kjellan 680m 17.7.1904, J. Hagen 997 in hb E E00002592 [isotypus] in hb EGER [isotypus] Basionym: Distribution: Norway fo nov Schiffner Bonner Hattori Pocs = Frullania tamarisci 1936 1965 1972 1976 992 655 536 2686 Krit Bemerk europ Leberm, ser 20: 23 [Hepat europ exsicc, nr. 997] Index Hepat 5: 443 [Norway] J Hatt Bot Lab 35: 226 Fol. Hist. nat. Mus. Matr. 4: 19 [types in hb EGER] (L.) Dumort. 1835 tamarisci var. americana Nees 1845 Typus: USA. "N-America" Beyrich hb HAMPE 164 Basionym: Distribution: USA Nees in G., L. & N. Bonner Hattori 1845 1965 1972 = Frullania tamarisci ssp. asagrayana Frullania asagrayana 2850 Syn Hepat: 439 655 Index Hepat 5: 443 536 J Hatt Bot Lab 35: 232 (Mont.) S.Hatt. in Hara 1966 Mont. 1842 Page 413 tamarisci var. azorica Frahm 2006 Basionym: Distribution: Azores (Terceira, Pico) [Portugal] var nov - nom nov Frahm Heinrichs Frahm Hentschel et alii = Frullania acicularis 2006 2010 2011 2015 1718 Trop Bryol 27: 101-105 [new var] [Portugal, Azores] 2834 Phylogenetics and Evolution 56 :1105-1114 3415 Archive for Bryology 116: 2-3, fig. 9-10 4097 Phytotaxa 220 (2): 134 [Frullania (subg. Thyopsiella) acicularis Hentschel et von Konrat, nom. nov.] Hentschel & von Konrat 2015 tamarisci var. blanda De Not. 1865 Typus: Italy, S. Eusebio in Bisagno, sopra Genova in hb G :2222 Basionym: Frullania blanda Distribution: Italy - syn nov -.11.1856, De Notaris De Not. 1858 [nom nud - nom herb ?] Notaris Müller in Rabenhorst Bonner Hattori 1865 1915 1965 1972 576 1064 655 536 Mem Reale Accad Sci Torino, ser 2, 22 (1): 378-380, pl 4, fig 21(1-34) Krypt Flora (2ed) 6, 2: 613 [syn sub Fr tamarisci var atrovirens] Index Hepat 5: 444 J Hatt Bot Lab 35: 226 = Frullania tamarisci var. atrovirens Carrington 1863 tamarisci var. chlorella De Not. ex Bonner 1965 Typus: Sardinia. Monte dei Setre Fratelli -.1.1858, Notaris Basionym: Distribution: Notaris 1865 576 Bonner Hattori 1965 1972 655 536 = Frullania tamarisci Mem Reale Accad Sci Torino, ser 2, 22 (1): 382 [chlorella as forma of Fr tamarisci sardoa] Index Hepat 5: 444 J Hatt Bot Lab 35: 226 (L.) Dumort. 1835 tamarisci var. cornubica fo. corticicola Schiffn. 1936 Typus: Ireland. Killarney -.8.1911, D.A. Jones & J.B. Duncan in hb E E00007479 ( as "corticola" ) [isotypus] in hb EGER [isotypus] Basionym: Distribution: fo nov Schiffner 1936 fo nov Schiffner Bonner Hattori Pocs 1936 1965 1972 1976 - syn nov 992 Krit Bemerk europ Leberm, ser 20: 22-23 [Hepat europ exsicc, nr. 996] 437 Österr Bot Zeitschr 85 (2): 132-134, fig 3 [Ireland: Killarney] 655 Index Hepat 5: 445 536 J Hatt Bot Lab 35: 226 2686 Fol. Hist. nat. Mus. Matr. 4: 19 [types in hb EGER] = Frullania dilatata var. blanda De Not. by Bornmüller 1931 [orth var] Frullania tamarisci var. blanda De Not. 1865 Frullania tamarisci var. atrovirens Carrington 1863 Page 414 tamarisci var. cornubica fo. corticola Schiffn. 1936 [orth var in hb E E00007479] Typus: in hb E E00007479 (isotypus of Frullania tamarisci (L.) Dumort. var. cornubica Carr. forma corticola Schiffn.) Basionym: Distribution: Schiffner 1936 992 Krit Bemerk europ Leberm, ser 20: 22 [Hepat europ exsicc, nr. 996] = Frullania tamarisci var. cornubica fo. corticicola Schiffn. 1936 Frullania dilatata var. blanda De Not. by Bornmüller 1931 [orth var] Frullania tamarisci var. blanda De Not. 1865 Frullania tamarisci var. atrovirens Carrington 1863 tamarisci var. cornubica fo. major, acutiloba Schiffn. 1936 Typus: Ireland. Killarney -.8.1911, D.A. Jones, J.B. Duncan & Wilson Basionym: Distribution: Schiffner 1936 992 Krit Bemerk europ Leberm, ser 20: 22 [Hepat europ exsicc, nr. 994] = Frullania dilatata var. blanda De Not. by Bornmüller 1931 [orth var] Frullania tamarisci var. blanda De Not. 1865 Frullania tamarisci var. atrovirens Carrington 1863 tamarisci var. cornubica fo. obtusiloba Schiffn. 1936 Typus: Ireland. Killarney, Upper Lake -.8.1911, D.A. Jones & J.B. Duncan & Wilson 995 in hb E E00007480 [isotypus] in hb EGER [isotypus] Basionym: Distribution: fo nov - syn nov Schiffner Schiffner Bonner Hattori Pocs = Frullania tamarisci 1936 1936 1965 1972 1976 992 437 655 536 2686 Krit Bemerk europ Leberm, ser 20: 22 [Hepat europ exsicc, nr. 995] Österr Bot Zeitschr 85 (2): 132, fig 2 [Ireland: Killarney] Index Hepat 5: 445 J Hatt Bot Lab 35: 226-227 [syn sub Fr. tamarisci] Fol. Hist. nat. Mus. Matr. 4: 19 [types in hb EGER] (L.) Dumort. 1835 tamarisci var. elongatistipula (Verd.) S.Hatt. 1985 [error] Basionym: Distribution: Piippo Gao & Wu 1990 2010 521 J Hatt Bot Lab 68: 47 & 142 [annotated catalogue of China] 2349 Genera Hepat. et Anth. Sinicorum: fig. 247(1-10) [key in Chinese] = Frullania tamarisci ssp. obscura var. elongatistipula (Verd.) S.Hatt. 1985 Page 415 tamarisci var. ericetorum Jorgensen 1934 Typus: Norway. Basionym: Distribution: var nov Jorgensen Arnell Bonner Hattori Damsholt = Frullania tamarisci 1934 1956 1965 1972 2002 225 43 655 536 1020 Bergens Mus Skrift 16: 328-329 [var nov] Moss Flora of Fennoscandia: 257 Index Hepat 5: 445 J Hatt Bot Lab 35: 226 Illust Fl Nordic Liverworts & Hornworts: 602 [key to var., Northern Europe] (L.) Dumort. 1835 tamarisci var. explanata Kaalaas 1893 Typus: Norway. S. Bergenhus sd, Kaalaas Basionym: Distribution: - syn nov Müller in Rabenhorst Jorgensen Bonner 1915 1934 1965 1064 Krypt Flora (2ed) 6, 2: 613 [syn sub Fr tamarisci var atrovirens] 225 Bergens Mus Skrift 16: 329 655 Index Hepat 5: 445 = Frullania tamarisci var. atrovirens Carrington 1863 tamarisci var. germana (Taylor) Carrington 1863 Basionym: Jungermannia germana Distribution: Europe Taylor 1843 Carrington 1863 Stephani Müller in Rabenhorst Bonner Bisang, Schumacker, Sérgio & Grolle 1911 1915 1965 1989 = Frullania teneriffae Trans Proc Bot Soc Edinburgh 7: 457 "Transactions and Proceedings of the Botanical Society of Edinburgh" 686 Spec Hepat 4: 565 [not mentioned as syn by Fr. germana] 1064 Krypt Flora (2ed) 6, 2: 616 [syn sub Fr. germana] 655 Index Hepat 5: 446 276 Giorn Bot Ital 122: 264 [syn sub Fr. teneriffae] (F.Weber) Nees 1838 tamarisci var. heterophylla Corbiere 1889 Typus: France. Environs de la Fauconniere pres Cherbourg or ? le Mesril-au-Val Basionym: Distribution: var nov - syn nov Corbiere Müller in Rabenhorst Schiffner Bonner Hattori 1889 1915 1936 1965 1972 1704 1064 992 655 536 Mem Soc Nat Sci Nat Math Cherbourg 26: 345 Krypt Flora (2ed) 6, 2: 613 [syn sub Fr tamarisci var atrovirens] Krit Bemerk europ Leberm, ser 20: 15 [Hepat europ exsicc, nr. 983] Index Hepat 5: 446 J Hatt Bot Lab 35: 226 = Frullania tamarisci var. atrovirens Carrington 1863 Page 416 tamarisci var. heterophylla fo. brunnea Schiffn. 1936 Typus: France. Manche, Mesnil-au-VAl 26.2.1902, L. Corbiere 984 in hb E E00002591 [isotypus] Basionym: Distribution: Schiffner 1936 992 Bonner 1965 655 = Frullania tamarisci Krit Bemerk europ Leberm, ser 20: 15-16 [Hepat europ exsicc, nr. 984-985] Index Hepat 5: 446 (L.) Dumort. 1835 tamarisci var. hispanica (Nees) Nees 1845 Basionym: Distribution: Nees in G., L. & N. Stephani Bonner 1845 1911 1965 Hattori 1972 = Frullania tamarisci 2850 Syn Hepat: 439 686 Spec Hepat 4: 560 [not mentioned as syn of Fr. hispanica] 655 Index Hepat 5: 446 [error: not as syn of Fr. hispanica in STEPH 1911: 560] 536 J Hatt Bot Lab 35: 226 (L.) Dumort. 1835 tamarisci var. hispanica (Nees) Nees 1845 [pp] Basionym: Distribution: - syn nov Nees in G., L. & N. Notaris Jelenc 1845 1865 1950 2850 Syn Hepat: 439 576 Mem Reale Accad Sci Torino, ser 2, 22 (1): 380 1707 Soc Geogr Arch Prov d'Oran 73 (fasc 228): 72 [North Africa] = Frullania tamarisci var. sardoa (De Not.) De Not. 1865 tamarisci var. mediterranea fo. robustior Schiffn. 1936 Typus: Corsica. Fango bei Bastia, 200m 15.4.1905, Schiffner 987 in hb E E00002596 [isotypus] in hb EGER [isotypus] Basionym: Distribution: fo nov Schiffner 1936 Bonner Pocs 1965 1976 992 Krit Bemerk europ Leberm, ser 20: 18-20 [Hepat europ exsicc, nr. 987, 989b] 655 Index Hepat 5: 447 [Corsica, Southern France] 2686 Fol. Hist. nat. Mus. Matr. 4: 19 [types in hb EGER] = Frullania tamarisci var. mediterranea (De Not.) Nees 1845 Page 417 tamarisci var. nervosa (Mont.) S.W.Arnell 1961 Basionym: Distribution: Portugal, Madeira [Portugal], Azores [Portugal], Canary Isl ( Fu La ) Arnell Grolle 1961 1970 180 615 Düll 1983 620 Malme Sim-Sim 1988 1989 625 6 = Frullania tamarisci Svensk Bot Tidskr 55 (2): 392 [may be a variety of Fr tamarisci] Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 19 (3): 312-313 [Fr. nervosa as syn of Fr. tamarisci] Bryol Beitr 2: 68 & 78 [seems to be a good var or subsp, not a syn of Fr. tamarisci] Sommerfeltia 7: 16 [Fuerteventura, Lanzarote] Portug Acta Biol (B) 15: 364 [Portugal] (L.) Dumort. 1835 tamarisci var. paradoxa Hampe ex Lehm. & Lindenb. 1845 [pro syn] Basionym: Distribution: Lehmann & Lindenberg in G., L. 1845 & N. = Frullania paradoxa 2850 Syn Hepat: 463 [syn sub Fr paradoxa] Lehm. & Lindenb. 1845 tamarisci var. sardoa fo. chlorella De Not. 1865 Typus: Sardinia. Sardegna meridionale, Monte dei Sette Fratelli -.6.1858, De Notaris Basionym: Distribution: De Notaris Schiffner 1865 1936 576 992 Bonner 1965 655 Mem Reale Accad Sci Torino, ser 2, 22 (1): 382, pl 5 fig 22(1-2) Krit Bemerk europ Leberm, ser 20: 21 [Hepat europ exsicc, nr. 991b pp.] Index Hepat 5: 448 = Frullania tamarisci var. sardoa (De Not.) De Not. 1865 tamarisci var. viernamica (S.Hatt.) S.Hatt. by X.L.Bai in C.Gao & Cao 2000 [orth var] Typus: Basionym: Distribution: Bai in Gao & Cao 2000 1019 Fl Yunnanica 17: 434 [China, Yunnan] = Frullania tamarisci var. vietnamica Hatt. (Hatt.) in Piippo, He & Koponen 1997 [error] Frullania tamarisci ssp. obscura var. vietnamica (S.Hatt.) S.Hatt. 1985 Page 418 tamarisci var. vietnamica Hatt. (Hatt.) in Piippo, He & Koponen 1997 [error] Basionym: Distribution: China (Sichuan, Yunnan), Vietnam error Piippo, He & Koponen 1997 Piippo, He, Koponen, Refearn & 1998 Li Bai in Gao & Cao 2000 Bai 2001 Bai 2002 628 Ann Bot Fennici 34: 52, 61 [Sichuan] [error, as Fr. tamarisci var. vietnamica] 697 J Hatt Bot Lab 84: 144 [checklist of Yunnan, China, [as Fr. tamarisci var. vietnamica] 1019 Fl Yunnanica 17: 434 [China, Yunnan, as Fr. tamarisci var. vietnamica] 1742 Guizhou Science 19(4): 51 [China, Yunnan, as Fr. tamarisci var. vietnamica] 1189 Chenia 7: 23 [Yunnan, China, as Fr. tamarisci var. vietnamica] = Frullania tamarisci ssp. obscura var. vietnamica (S.Hatt.) S.Hatt. 1985 tamarisci ssp. asagrayana Basionym: Frullania asagrayana Distribution: comb nov - syn nov (Mont.) S.Hatt. in Hara 1966 Mont. 1842 Hattori in Hara Hattori Crandall-Stotler, Stotler & Geissler Asakawa, Sono, Wakamatsu, Kondo, Hattori & Mizutani Schuster Timme & Redfearn 1966 1972 1987 841 536 399 1991 1682 Phytochemistry 30(7): 2297 [World map of Fr tamarisci subspecies] 1992 1997 603 631 Timme & Redfearn 1997 Lyman 2003 = Frullania asagrayana Flora of Eastern Himalaya: 528 J Hatt Bot Lab 35: 232-235, fig. 5, 12, 13 Bryologist 90 (4): 287-308 Hepat N America 5: 60-65, fig. 780-781 http://science.smsu.edu/~csre/herbarium/liverw_t.htm [4.6.97] printed 15.1.98 12:34 page 3 of 6 [USA] 1134 http://biology.smsu.edu/herbarium/liverw_t.htm [printed 20.6.2004 page 10 of 19; USA] 1268 Checklist of Wisconsin bryophytes [USA] Mont. 1842 Page 419 tamarisci ssp. moniliata [auct. non (RH.) Mont.] Kamim. 1961 Basionym: Distribution: Sakhalin, China (Guizhou, Heilongjiang), Taiwan, Ryukyu (Amami), Korea (South) auct - syn nov Kamimura Hattori, Hong & Inoue Nakanishi Kitagawa Hong Shin Shin Lai & Wang-Yang Hu & Wang Li & Gao Hattori & Lin Aur & Zhang Wu in Li, Gao, Wu et al. Kuo & Chiang Piippo Zhu & Hu Wang & Tau Yamada & Choe Ji & Liu Yamada & Iwatsuki 1961 1962 1963 1963 1966 1968 1970 1976 1981 1983 1985 1985 1985 1988 1990 1991 1992 1997 1998 2006 = Frullania tamarisci ssp. obscura 199 876 960 626 1107 844 150 33 2800 505 1022 1169 948 521 717 1333 1470 J Hatt Bot Lab 24: 65, fig 23(1-38) 24(1-44), map 7 J Hatt Bot Lab 25: 284 [Korea] Hikobia 3(4): 321, 322, 324 [Taiwan] Acta Phytotax Geobot 19: 150 [Sakhalin] Bryologist 69 (1): 412-413 [South Korea] Sci Rep Kagoshima Univ 17: 204 [Ryukyu Arch.] Sci Rep Kagoshima Univ 19: 26 [Ryukyu: Amami Isl] Taiwania 21 (2): 168 [Index to Taiwan] J East China Normal Univ (Nat Sci) 1981 (1): 85-104[aus 698] Invest Stud Nat 1: <19-26> [aus 698] J Hatt Bot Lab 59: 161 Fl Heilongjiangensis 1: 166, fig 58(1-19) [China] Bryoflora Xizang (Tibet), Science Press, Beijing: 525-527, fig 228(1-7) Taiwania 33: 13 [Index of Taiwan] J Hatt Bot Lab 68: 47 [annotated catalogue of China] J East China Normal Univ (Nat Sci) 1991 (3): <98-103> [aus 698] [aus 698] J Hatt Bot Lab 81: 288 [Korean Peninsula] Acta mus hist Nat Sinica 16: 13 [China] J Hatt Bot Lab 99: 33 [Catalog of Japan] (Verd.) S.Hatt. 1972 Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata tamarisci ssp. nisquallensis (Reinw., Blume & Nees) Mont. 1842 (Sull.) S.Hatt. in Hara 1966 Basionym: Distribution: comb nov Schuster & Steere Steere Hattori in Hara Hattori Hattori Murray Abramova, Afonina & Duda Schljakov 1958 1965 1966 1972 1973 1974 1974 1975 Steere Steere & Inoue Konstantinova, Potemkin & Schijakov Qian & Klinka McCune, Rosentreter, Ponzetti & Shaw Peck & Muir 1976 1978 1992 = Frullania nisquallensis 1998 2000 2001 vide 633 vide 633 841 Flora of Eastern Himalaya: 528 536 J Hatt Bot Lab 35: 235-239, fig 13-15 403 Misc Bryol Lichen 6 (5): 72-73, fig a-f [Lena, Sibiria] vide 633 1336 Nov Syst Plant non Vascul 11: 317 [UdSSR] 1337 Nov Syst Plant non Vascul 12: 322 [UdSSR] [as Fr. tamarisci nisquallensis] vide 633 633 J Hatt Bot Lab 44: 330-331, fig 21(1-11) [Alaska] 651 Arctoa 1:102 [USSR] 2262 Plants of British Columbia: 339 [Canada] 915 Bryologist 103 (3): 420 [epiphytic in old conifer forest, USA: Washington] 917 Bryologist 104 (2): 183-190 [harvestable epiphytic bryophytes, Western Oregon, USA] Sull. 1849 Page 420 tamarisci ssp. obscura (Verd.) S.Hatt. 1972 Basionym: Frullania moniliata ssp. obscura Distribution: Japan, Korea (South) comb nov var nov Verd. 1930 {nördl. Indomalaya} Hattori 1972 536 Hattori Hattori Choe & Yamada Lai & Wang-Yang Hattori & Thaithong Hattori in Mizutani Hattori Hattori Choe Inoue Yuzawa & Hattori Hattori & Lin Tan & Engel Kuo & Chiang Piippo Long & Grolle Asakawa, Sono, Wakamatsu, Kondo, Hattori & Mizutani Konstantinova, Potemkin & Schijakov Wang Hashimoto, Asakawa, Nakashima & Tori Furuki & Mizutani Furuki & Higuchi 1973 1974 1974 1976 1978 1979 1979 1980 1980 1981 1981 1985 1986 1988 1990 1990 1991 J Hatt Bot Lab 35: 216, fig 1, 2(a-l), 3(a-h), 4(a-e), 6, 8, 13(q-y) [no basionym clarified] 542 J Hatt Bot Lab 37: 121-122, fig 67 544 J Hatt Bot Lab 38: 173 [Luzon] 405 Misc Bryol Lichen 6 (9): 161 [checklist Korea] 33 Taiwania 21 (2): 168 [Index to Taiwan] 481 J Hatt Bot Lab 44: 193 [Assam] 487 J Hatt Bot Lab 46: 323 [Easter Nepal] 129 J Jap Bot 54 (1): 25 [Sri Lanka] 958 Bull Natn Sci Mus Tokyo, ser B (Bot), 6 (1): 39 [Formosa] 1171 Illust Fl & Fauna Korea 24: 735 [South Korea] 963 Mem Natn Sci Mus Tokyo 14: 65 [Japan, Mt. Fuji] 494 J Hatt Bot Lab 50: 256-257 [Japan] 505 J Hatt Bot Lab 59: 161 [China, synonyms] 4286 J. Hatt. Bot. Lab.. No. 60: 305 [Philippines] 948 Taiwania 33: 13 [Index of Taiwan] 521 J Hatt Bot Lab 68: 47 [annotated catalogue of China] 522 J Hatt Bot Lab 68: 413 [Bhutan] 1682 Phytochemistry 30(7): 2297 [World map of Fr tamarisci subspecies] 1992 651 1993 1993 1223 Chenia 1: 130 [south China, Guangxi Prov.] 1199 J Hatt Bot Lab 74: 125 [Chemical constituents] 1994 1997 Yamada & Choe Hasegawa Yuzawa 1997 1998 2000 Bapna & Kachroo Wu & Wang Peng, Enroth, Koponen & Piippo Iwatsuki 2000 2000 2000 2001 Song & Yamada Yamada & Iwatsuki 2006 2006 1331 Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] 1234 Bull Natn Sci Mis, Tokyo, ser B 23(3): 89, pl 8(3) [oil bodies & oil droplets] 717 J Hatt Bot Lab 81: 288 [Korean South & North] 880 Bull Fac Horticult Minaanikyushu Univ 28 (A): 29 [Kyushu] 1659 Natural Environmental Sci Res 13: 18-21, fig 16(a-r) [Japan, keys to species; in Japanese] 712 Hepaticology in India II: 195-196 2671 Bryoflora of Hengduan MTS (Southwest China): 131, fig. 58(9-12) 776 Hikobia 13: 197 [China: Hubei Prov] 957 Mosses and Liverworts of Japan, Heibonsha (Tokyo): 280, pl 162(1-2) [with key to Japan, in Japanese] 1445 J H B L 100: 446 [from Jeju (Cheju) Isl, Korea] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] Arctoa 1:102 [USSR] = Frullania tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Frullania moniliata (Reinw., Blume & Nees) Mont. 1842 tamarisci ssp. obscura var. balansae (Steph.) S.Hatt. 1972 Basionym: Frullania balansae Steph. 1894 Distribution: China (Hainan, Hongkong), Taiwan, India, Thailand syn nov Hattori Lai & Wang-Yang Hattori, Thaithong & Kitagawa Hattori & Lin Crandall-Stotler, Stotler & Geissler Piippo Bapna & Kachroo = Frullania moniliata 1972 1976 1977 1985 1987 536 33 480 505 399 J Hatt Bot Lab 35: 220-222, fig 7 Taiwania 21 (2): 168 [Index to Taiwan] J Hatt Bot Lab 43: 444 [Thailand] J Hatt Bot Lab 59: 161 [China; synonymys] Bryologist 90 (4): 287-308 1990 2000 521 712 J Hatt Bot Lab 68: 47 [annotated catalogue of China] Hepaticology in India II: 196-197 (Reinw., Blume & Nees) Mont. 1842 Page 421 tamarisci ssp. obscura var. breviramea (Steph.) S.Hatt. 1972 Basionym: Frullania breviramea Steph. 1911 Distribution: Taiwan , India, Thailand , Indonesia (Java), Borneo comb nov - syn nov Hattori Kamimura Hattori Hattori Hattori, Thaithong & Kitagawa Hattori Onraedt Tan & Engel Crandall-Stotler, Stotler & Geissler Menzel Kuo & Chiang = Frullania moniliata 1972 1974 1975 1976 1977 1979 1981 1986 1987 536 3692 547 80 480 129 5794 4286 399 J Hatt Bot Lab 35: 222-226, fig 9 [comb nov] Bull Kochi Gakuen J Coll 5: 78-79 [Java, Bogor Mt. Tjibodas] J Hatt Bot Lab 39: 309-310 J Hatt Bot Lab 40: 498, fig 200(t) J Hatt Bot Lab 43: 444 [Thailand] J Jap Bot 54 (1): 25 [Sri Lanka] J Hatt Bot Lab 50: 203 [Sri Lanka] J. Hatt. Bot. Lab.. No. 60: 305 [Philippines] Bryologist 90 (4): 287-308 [Frullania moniliata] 1988 1988 512 948 J Hatt Bot Lab 65: 165 [Cat Borneo] Taiwania 33: 13 [Index of Taiwan] (Reinw., Blume & Nees) Mont. 1842 tamarisci ssp. obscura var. minshanensis S.Hatt. 1978 Typus: China. SE Kansu, Min-Shan Range, .. 23.7.1930, Dr.D. Hummel B-273 in hb NICH [holotypus], in hb COLO [isotypus] Basionym: Distribution: var nov - comb nov Hattori Hattori in Hattori & Lin Piippo 1978 1985 1990 482 505 521 J Hatt Bot Lab 44: 539-540, fig 210 J Hatt Bot Lab 59: 161 [Fr tamarisci ssp obscura fo. minshanensis] J Hatt Bot Lab 68: 47 [annotated catalogue of China] = Frullania tamarisci ssp. obscura fo. minshanensis (S.Hatt.) S.Hatt. in S.Hatt. & Lin 1985 tamarisci ssp. obscura var. obscura (Verd.) S.Hatt. 1972 Basionym: Distribution: - syn nov comb nov Hattori = Frullania moniliata 0 1974 339 544 J Hatt Bot Lab 38: 173 [Luzon] (Reinw., Blume & Nees) Mont. 1842 Page 422 tamsuina fo. reniformia Kamim. 1952 Typus: Japan. Shikoku, Prov. Jyo, Mt. Ishizuchi -.7.1938, Kamim. 5570 in hb Kamim. [holotypus] Basionym: Distribution: Japan fo nov - syn nov Kamimura Kamimura Hattori & Noguchi Kamimura Bonner Wu in Li, Gao, Wu et al. Furuki & Mizutani Yuzawa Yamada & Iwatsuki = Frullania delavayi 873 873 965 199 655 1169 1331 1660 1470 Contrib Fl Hepat Shikokuensem, Kochi: 133, fig 31 Contrib Fl Hepat Shikokuensem, Kochi: 133 Index spec Typ herb Japon, Bryophyta: 18 [holotype] J Hatt Bot Lab 24: 83 [syn sub Fr. devalayi] Index Hepat 5: 448 Bryoflora Xizang (Tibet), Science Press, Beijing: 527 [sub Fr delavayi] Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] Natural Environmental Sci Res 14: 43 [Japan, syn sub Fr inflexa] J Hatt Bot Lab 99: 33 [Catalog of Japan] Steph. 1894 Frullania inflexa tatanarivensis 1952 1952 1960 1961 1965 1985 1994 2001 2006 Mitt. 1861 [auct. non Steph. 1894] Steph. in Brunnthaler 1913 Typus: "Natal. Drakensberge, Van Reenen-Paß" 1909, Brunnthaler in hb G Basionym: Distribution: -syn nov Stephani in Brunnthaler Sim 1913 1926 Demaret & Vanden Berghen 1947 = Frullania squarrosa Typus: Madagascar Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 729 [Natal] Trans Roy Soc S Africa 15: 44 [BRUNNTHALER identification from South Africa may be wrong] 1101 Bull Jard Bot Etat Bruxelles 18: 238 [synsub Fr squarrosa] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides tatanarivensis 219 252 (Nees) Mont. 1839 Steph. 1894 sd, Renauld 172 in hb G & BM Basionym: Distribution: spec nov - syn nov Stephani Stephani Demaret & Vanden Berghen Bonner Vanden Berghen Stephani 1894 1910 1947 1965 1976 1985 Grolle Wigginton 1995 2002 Wigginton Wigginton 2004 2004 = Frullania ericoides 228 686 1101 655 185 928 Hedwigia 33: 148-149 Spec Hepat 4: 373 Bull Jard Bot Etat Bruxelles 18: 239 [Madagascar] Index Hepat 5: 449 [Madagascar] Bull Jard Bot Nat Belg 46 (1/2): 163 Stephani Icon Hepat Inedit [Galeiloba] C- 41, microfiche 50 fig 3396 ["Madagascar, Renauld"] 250 Bryophyt Bibl 48: 65 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 231 [syn sub Fr ericoides, West Africa] (Nees) Mont. 1839 Page 423 taxodiocola R.M.Schust. 1983 Typus: USA. Florida, Monroe Co., Cypress swamp 25.12.1958, Schuster 42241 in hb FM [only typus collection] Basionym: Distribution: USA ( FL ) spec nov - syn nov Schuster Schuster Gradstein = Frullania subtilissima taylori 1983 1992 2021 440 Phytologia 53 (6): 364-365 [close to Fr. obchordata - gymnotis] 603 Hepat N America 5: 104-111, fig 790 5642 Men NY Bot Gard 121: 380 [syn sub Fr subtilissima], the type material cannot be found (L. Briscoe, pers. comm.) (Nees ex Mont.) Lindenb. 1845 Gottsche 1845 Typus: Guyana. Georgtown, Demerara sd, Campbell sn in hb S (B24309, 24310) [typus] in hb E00002587 (sd, s coll sn) [possible typus of Frullania taylorii Gott. et al.] Basionym: Distribution: spec nov Gottsche in G., L. & N. Stephani Verdoorn Stephani 1845 1911 1930 1985 2850 686 293 928 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 2002 782 = Frullania gaudichaudii tenella Syn Hepat: 436 Spec Hepat 4: 524 Ann Bryol Suppl 1: 92 [syn sub Fr. gaudichaudi] Stephani Icon Hepat Inedit [Thyopsiella] F-126, microfiche 58 fig 4053 ["Demerara (Campbell legit)"] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] (Nees & Mont.) Nees & Mont. 1845 [auct. non Sande Lac. 1854] Verd. 1930 Basionym: Distribution: Verdoorn 1930 293 Ann Bryol Suppl 1: 135 = Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 Page 424 tenella Sande Lac. 1854 [non Caspary 1887, nec Steph. 1910] Basionym: Distribution: spec nov - comb nov Sande Lacoste Sande Lacoste Sande Lacoste in Dozy Sande Lacoste 1855 1856 1856 1857 2038 1894 564 430 Sande Lacoste in Veth 1884 807 Schiffner Stephani Verdoorn Bonner Hattori 1898 1911 1930 1965 1975 831 686 293 655 66 Grolle & Hattori in Hattori 1982 422 Miller, Whittier & Whittier 1983 609 Nederl Kruidk Arch 3: 423 Natuurk Tjidschr v Nederl Indie 10: 398 Plagiochila sandei Dozy: 11 Verh Konikl Akad Wetenschappen 5: 87 [comb nov: Fr. sinuata var. tenella] Midden Sumatra, Reizen en Onderzoekingen der Sumatra Expeditie, 4 (2): 44 [Sumatra] Conspect Hepat Archip Indici, Batavia: 345-346 Spec Hepat 4: 644 [not mentioned as syn sub Fr. sundaica] Ann Bryol Suppl 1: 134 [syn sub Fr. junghuhniana] Index Hepat 5: 449 Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (4): 148-149, fig 162 [syn sub Fr. junghuhniana var. minutissima] Misc Bryol Lichen 9 (6): 123-124 [comb nov: Fr. junghuhniana var. tenella] Prod Flora Hepat Polynesiae: 140 = Frullania sinuata var. tenella (Sande Lac.) Sande Lac. 1857 Frullania junghuhniana var. minutissima Schiffn. ex Verd. 1929 Frullania junghuhniana var. tenella (Sande Lac.) Grolle & S.Hatt. 1982 tenella Steph. ex Yoshinaga 1900 [nom nud] {specimen from Japan} Basionym: Distribution: Yoshinaga Inoue Sawada = Frullania tenella 1887] 1900 1900 1909 Steph. 1910 [hom illeg] [non Sande Lacoste 1854, nec Caspary Frullania diversitexta tenella 1685 Bot Mag Tokyo 14: 40 [F. tenella Steph. n. sp.] 802 Bot Mag Tokyo 14 (no 162): 181 930 Bot Mag Tokyo 23 (no 268): 178-183, fig 33 [key to Japanese species, in Japanese] Steph. 1897 Caspary 1887 [hom illeg] [non Sande Lacoste 1854, nec Steph. 1910] Typus: FOSSIL. Basionym: Distribution: Fossil (Eocene) in Baltic amber, Germany spec nov Caspary Caspary & Klebs Grolle = Frullania casparyi 1887 1907 1985 977 622 621 Schriften phys-ökonom Gesell Königsberg 27: 5, pl 1 fig 21 Abh königl Preuß Geol Landesanstalt, NF 4: 42, Atlas fig 34(a-b) Prace Muz Ziemi z 37: 92, 97 [may be syn of Fr caspary Grolle] Grolle 1985 Page 425 tenella Steph. 1910 [hom illeg] [non Sande Lacoste 1854, nec Caspary 1887] Typus: Japan. Sikoku, Tosa, Jasui -.8.1898, Inoue 14 in hb G; in hb TNS [isotypus] (lecto-, designated here, G[G00066936]!) syn. fide Hattori (1952) Basionym: Distribution: Japan spec nov syn syn Sawada 1909 930 Stephani Iishiba Horikawa Kamimura Hattori Hattori & Noguchi Kamimura Bonner Choe Stephani 1910 1912 1943 1952 1952 1960 1961 1965 1980 1985 686 888 1283 873 529 965 199 655 1171 928 Yamada & Iwatsuki Atwood, Vilnet, Mamontov 2006 2021 1470 5566 = Frullania diversitexta tenera Bot Mag Tokyo 23 (no 268): 178-183, fig 33 [key to Japanese species, in Japanese] Spec Hepat 4: 397 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Acta Phytotax Geobot 13: 213 [nom nov pro Fr tenella STEPH 1910] Contrib Fl Hepat Shikokuensem, Kochi: 123 [syn sub Fr. diversitexta] J Hatt Bot Lab 8: 29 [syn sub Fr diversitexta] Index spec Typ herb Japon, Bryophyta: 18 J Hatt Bot Lab 24: 81 [sub F diversitexta] Index Hepat 5: 449 Illust Fl & Fauna Korea 24: 733 [sub Fr diversitexta] Stephani Icon Hepat Inedit [Galeiloba] C-254, microfiche 53 fig 3674 ["Japan, Tosa, Jasui, Inoué No 14"] J Hatt Bot Lab 99: 33 [Catalog of Japan] Cryptogamie, Bryologie 42(3): 19-31 Steph. 1897 Spruce 1884 [hom illeg] [non Lindb. in Brotherus 1892] Typus: Peru. San Martin, Trapoto. " Andes Peruviae, Flum. Huallaga" sd, Spruce sn in hb BM [isotypus] in hb US in hb E E00002588 [isosyntypus] Basionym: Distribution: Mexico, Peru spec nov Spruce Stephani Clark Bonner Menzel Stephani 1884 1910 1959 1965 1984 1985 243 686 393 655 142 928 Fulford & Sharp Yuzawa NN 1990 1991 1996 105 525 632 = Frullania rio-janeirensis Trans Proc Bot Soc Edinburgh 15: 13 & 23-24 (No 12) Spec Hepat 4: 330 Bryologist 62: 190-193, fig. 1-18 Index Hepat 5: 450 Willdenowia 14: 490 [Peru] Stephani Icon Hepat Inedit [Chonanthelia] A- 51, microfiche 47 fig 3142-3143 ["Andes Peruviae Taraoto leg Spruce"] Mem NY Bot Gard 63: 46 [Mexico] J Hatt Bot Lab 70: 252 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 32 of 35 [in hb US] (Raddi) Ångstr. 1876 Page 426 tenera Lindb. ex Steph. in Brotherus 1892 [non Spruce 1884] Typus: Georgien. "Carthalinia", Borshom prope flumen Kura, ad lignum putridum 28.7.1877, A. H. & V. F. Brotherus 238 in hb H [holotypus] in hb G 7927 [isotypus] Basionym: Distribution: Caucasus spec nov - nom nov Stephani in Brotherus Stephani Bonner Hattori 1892 1910 1965 1978 922 686 655 482 Bisang, Schumacker, Sérgio & Grolle 1989 276 = Frullania parvistipula teneriffae Acta Soc Sci Fennicae 19 (12): 144 [Caucasus] Spec Hepat 4: 440 [= Fr. caucasica] Index Hepat 5: 450 J Hatt Bot Lab 44: 542-545. fig 213 [Fr. caucasica = syn nov sub Fr. parvistipula] Giorn Bot Ital 122: 265 [syn sub Fr. parvistipula] Steph. 1910 Mont. ex Lindenb. 1845 [pro syn] Basionym: Distribution: Lindenberg in G., L. & N. = Frullania polysticta 1845 2850 Syn Hepat: 440 Lindenb. 1845 teneriffae var. decipiens Schiffn. 1902 Basionym: Distribution: var nov syn nov Schiffner Schiffner = Frullania teneriffae 1902 1936 905 277 Hedwigia 41 : 281 Österr Bot Zeitschr 85 (1): 28 (F.Weber) Nees 1838 Page 427 tenuicaulis Mitt. 1891 Typus: China. Quantung Prov., Manshan Mts, near Ookaisa, 2400ft 1863, Dr. Tate in hb G (ex hb K) in hb NY [holotype] Basionym: Distribution: China, Taiwan, Thailand, Indonesia (Sumatra, Java, Sulawesi, Mollucas), Borneo, Malaysia (Sarawak, Sabah), Philippines spec nov Mitten Stephani Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Horikawa Verdoorn Herzog Kamimura Hattori, Iwatsuki & Kanno Hattori Kamimura Bonner Tixier Kamimura 1891 1911 1929 1929 1930 1932 1932 1934 1934 1937 1949 1952 1956 1956 1961 1965 1973 1974 - syn nov Hattori Lai & Wang-Yang Hattori Stephani 1975 1976 1982 1985 Kuo & Chiang Piippo Yamada & Iwatsuki Uribe 1988 1990 2006 2008 Uribe-M. 2011 = Frullania vethii Trans Linn Soc London, ser 2, 3: 203 [Japan] Spec Hepat 4: 613-614 Ann Jard Bot Buitenzorg 40: 139 Ann Bryol 2: 164 Ann Bryol Suppl 1: 151, fig 244-247, 254-256 Bull Jard Bot Buitenzorg, ser 3, 12 (1): 63 [Malay Peninsula, Java] Nederl Kruidk Arch Jrg 1932 (2): 499 [Sumatra, Java, Borneo] Ann Bryol Suppl 4: 206 [B.N. Borneo: Mt Kinabalu, Sarawak] J Sci Hiroshima Univ, ser B, div 2, 2: 236 [Taiwan] Blumea suppl 1: 213 [Key Islands] Trans Brit Bryol Soc 1 (3): 318 [Borneo, Sarawak] Contrib Fl Hepat Shikokuensem, Kochi: 134 J Hatt Bot Lab 16: 93 & 94 J Jap Bot 31: 202 J Hatt Bot Lab 24: 61, fig 22(1-31), map 5 Index Hepat 5: 450 Nat Hist Bull Siam Soc 24 (3/4): 456 [Thailand] Bull Kochi Gakuen J Coll 5: 77-78 [Borneo, Kinabalu area; Java, Bogor Mt. Tjibodas] 547 J Hatt Bot Lab 39: 311-312 33 Taiwania 21 (2): 169 [Index to Taiwan] 495 xxxxx [syn sub Fr xxxxxx] 928 Stephani Icon Hepat Inedit [Meteriopsis] E-63, microfiche 56 fig 3862 ["China, Province Quantung, Manshan Mts, 2400f, Dr Tate 1863"] 948 Taiwania 33: 13 [Index of Taiwan] 521 J Hatt Bot Lab 68: 48 [annotated catalogue of China] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] 1513 Caldasia 30(1): 90, [belongs to subgen Diastaloba] [Monography of subgenus Meteoriopsis] 2767 Caldasia 33(1): 74, fig 3(g-n) from type [type study,syn sub Fr trichodes, holotypus in hb NY] Sande Lac. 1884 Frullania trichodes tenuis 582 686 548 246 293 553 554 241 34 556 18 873 862 2792 199 655 427 3692 Mitt. 1862 Hampe ex Lehm. 1838 Typus: Ecuador. Loja sd, ex hb HAMPE "crescit ad corticem cinchonae (Chinae fuscae, loxae)", in hb BM 6643343 [isotypus] Basionym: Distribution: Venezuela spec nov - syn nov Hampe ex Lehmann Hampe in G., L. & N. Stephani Bonner Menzel 1838 1845 1911 1965 1984 Stephani 1985 Uribe & Gradstein Uribe 2003 2008 = Frullania brasiliensis 588 2850 686 655 142 Pugillus 7: 13-14 Syn Hepat: 461 (No 113) Spec Hepat 4: 598 Index Hepat 5: 450-451 Willdenowia 14: 506 [not in Peru, only in Venezuela - error by Hampe 1845: 461] 928 Stephani Icon Hepat Inedit [Meteriopsis] E-39, microfiche 55 fig 3825 ["Merida, Moritz"] - 3826 1071 Crypt Bryol 24 (3): 200 [syn sub Fr brasiliensis] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Raddi 1822 Page 428 tenuistipula Steph. ex Tatuno 1952 [nom nud] Basionym: Distribution: Tatuno 1952 1697 J Sci Hiroshima Univ ser B, div 2 (Bot), 6(2): 51-57 [sex chromosomes] = teres Sande Lac. 1857 Typus: Indonesia. Java, "in monte Ungarang" sd, Teysmann Basionym: Distribution: spec nov - syn nov. comb nov Sande Lacoste Sande Lacoste in Junghuhn Schiffner 1857 1857 1898 Stephani Verdoorn Verdoorn Bonner Stephani 1911 1929 1930 1965 1985 = Frullania ornithocephala ternatensis 430 Verh Konikl Akad Wetenschappen 5: 76-77, pl 15(1-11) 1490 Plantae Junghuhnianae, Hepaticae: 536 831 Conspect Hepat Archip Indici, Batavia: 336 [syn sub Fr. ornithocephala] 686 Spec Hepat 4: 451-452 246 Ann Bryol 2: 156 [notes to S.-L.] 293 Ann Bryol Suppl 1: 52, 55 655 Index Hepat 5: 451 928 Stephani Icon Hepat Inedit [Galeiloba] C-191abc, microfiche 52 fig 3593 ["Java, ex Herb Sande Lacoste, Original"] - 3595 ["Java, (Jung obscura Swz) Herb Nees"] - 3596 ["Birma, Berthoumier"] (Reinw., Blume & Nees) Nees 1845 [auct. non Gottsche 1846] S.Hatt. 1973 {Taiwan} Basionym: Distribution: Taiwan - syn nov Hattori Hattori & Lin Windolf Kuo & Chiang 1973 1985 1987 1988 540 505 1025 948 J Hatt Bot Lab 37: 8184, fig 40 [from type of Fr. concava] J Hatt Bot Lab 59: 162 Austrobaileya 2(4): 391 [checklist Queensland] Taiwania 33: 14 [Index of Taiwan] = Frullania ternatensis var. non-appendiculata S.Hatt. 1974 ternatensis var. non-appendiculata [auct. non S.Hatt. 1974] S.Hatt. 1976 [pp] Basionym: Distribution: auct non - syn nov Hattori Hattori = Frullania akiyamae tertuosa 1976 1986 80 507 J Hatt Bot Lab 40: 498-502, fig. 195-196, 198(e-g), 200(v-y) J Hatt Bot Lab 60: 240 [syn sub Fr. akiyamae] S.Hatt. 1986 Verd. by Kuo & Chiang 1988 [orth var] Basionym: Distribution: orth var Kuo & Chiang = Frullania tortuosa 1988 948 Taiwania 33: 14 [Index of Taiwan] Verd. 1929 Frullania hampeana Frullania monocera Nees 1845 (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Page 429 tetraptera [auct. non Nees & Mont. in Mont. 1838] Yuzawa 1991 {Brasil} Basionym: Distribution: Gradstein & Costa = Frullania cuencensis thomeensis 2003 984 Mem New York Bot Gard 87: 91 [not in Brazil] [syn sub Fr cuencensis] by Yuzawa 1991 Taylor 1846 Steph. 1910 Typus: San Thomé sd, Quintas sn in hb G [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Pearson Bonner Vanden Berghen Stephani 1910 1924 1965 1976 1985 Wigginton 2002 Wigginton 2004 Wigginton 2004 = Frullania obscura thuillerii 686 795 655 185 928 Spec Hepat 4: 321 Arkiv Botanik 19 (5): 9 [Mt. Elgon in Kenia/Uganda] Index Hepat 5: 451 Bull Jard Bot Nat Belg 46 (1/2): 155 Stephani Icon Hepat Inedit [Chonanthelia] A- 7, microfiche 46 fig 3074-3075 [from type; "San Thome Quintas leg"] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [probably Fr arecae] [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 224 [probably syn sub Fr arecae, West Africa] (Sw.) Mont. 1840 Nees 1845 Basionym: Jungermannia thuillerii [Schwägr. in hb Weigelt] Nees 1845 Distribution: Costa Rica, Panama, Surinam, ?Sri Lanka, ?Singapore, ?Micronesia spec nov - syn nov Nees in G., L. & N. Hampe Lindenberg & Gottsche Montagne Mitten in Seemann Mitten Sande Lacoste Schiffner Stephani Verdoorn Miller Bonner Miller Miller, Whittier & Whittier Stephani 1845 1851 1851 1855 1857 1861 1864 1898 1911 1928 1960 1965 1968 1983 1985 2850 281 282 475 1494 574 842 831 686 244 920 655 672 609 928 Gradstein & Costa 2003 984 = Frullania replicata Syn Hepat: 434 Linnaea 24 (3): 303 [Costa Rica] Linnaea 24 (5): 638 [Suriname] Ann Sci Nat Bot, ser 4, 3: 319 [Guiana] Bot Voyage HMS Herald: 246 [Panama] J Linn Soc Bot 5: 121 [Singapore, Sri Lanka] Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 313 [Sumatra, Amboina] Conspect Hepat Archip Indici, Batavia: 339 [syn sub Fr. replicata] Spec Hepat 4: 581 [syn sub Fr. replicata] Ann Crypt exot 1 (2): 218 [thuillerii = nom nov pro replicata] Bryologist 63: 121 [Micronesia] Index Hepat 5: 452 Micronesia 4 (1): 74 Prod Flora Hepat Polynesiae: 140 Stephani Icon Hepat Inedit [Homotropantha] D-32, microfiche 54 fig 3752 ["syn Frull Thuilleri Nees, leg Weigelt, Surinam"] - 3753 Mem New York Bot Gard 87: 89 [Brazil] [syn sub Fr nodulosa] (Nees) Mont. 1840 Frullania nodulosa (Reinw., Blume & Nees) Nees 1845 Page 430 thuillesii (Schwägr.) Nees by Bonner 1965 [orth var] Basionym: Distribution: Bonner Yano = Frullania thuillerii 1965 1984 Index Hepat 5: 452 J Hatt Bot Lab 56: 507 [checklist Brazil] Nees 1845 Frullania replicata (Nees) Mont. 1840 Frullania nodulosa thuillieri 655 501 (Reinw., Blume & Nees) Nees 1845 (Schwägr.) Nees by Verd. 1930 [orth var] Basionym: Distribution: Verdoorn = Frullania thuillerii 1930 Ann Bryol Suppl 1: 177 Nees 1845 Frullania replicata (Nees) Mont. 1840 Frullania nodulosa thuillierii 293 (Reinw., Blume & Nees) Nees 1845 Nees by Hampe & Gottsche 1852 [orth var] Basionym: Distribution: Hampe & Gottsche = Frullania thuillerii 1852 Linnaea 25: 357 Nees 1845 Frullania replicata (Nees) Mont. 1840 Frullania nodulosa tixierana 283 (Reinw., Blume & Nees) Nees 1845 Vanden Berghen 1976 Typus: La Reunion sd, Cailleux 58 in hb TIXIER [holotypus] in hb BR BR5040033553888 (sd, Cailleux 58, Reunion, Tevelave - as "tixieriana", ) [isotypus] Basionym: Distribution: Madagascar, Reunion Isl, Seychelles spec nov - syn nov Vanden Berghen Vanden Berghen Hattori & Vanden Berghen in Grolle Vanden Berghen Wigginton 1976 1976 1978 Wigginton 2004 = Frullania repandistipula 1982 2002 185 850 285 Bull Jard Bot Nat Belg 46 (1/2): 27-30, fig 6(A-I) Distributiones plantarum africanarum 11: 337 [map of Africa] Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 27 (1): 11 ["tixieriana", syn nov vide HATTORI + VANDEN BERGHEN in lit.] 76 Crypt Bryol Lichen 3 (3): 214 [sub Fr repandistipula] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sande Lac. 1853 Page 431 tixieriana Vanden Berghen by Grolle 1978 [orth var] Typus: in hb BR BR5040033553888 (coll sd sn, Reunion, Tevelave, Cailleux 58) [isotypus] Basionym: Distribution: orth var Grolle 1978 = Frullania tixierana Wiss Z Fried Schiller Univ Jena, Math Nat Reihe 27 (1): 11 ["tixieriana" insted of "tixierana"] Vanden Berghen 1976 Frullania repandistipula tolimana 285 Sande Lac. 1853 Steph. 1910 Typus: Colombia. "Nova Granada", Tolima, Ibaque, 1268 m 1901, Micholitz in hb G [holotypus] Basionym: Distribution: Costa Rica, Guatemala, Grenada, Colombia, Ecuador, Bolivia, Brazil spec nov - syn nov Stephani Bonner Gradstein & Hekking 1910 1965 1979 686 655 483 Stephani 1985 928 Yuzawa, Mues & Hattori Schäfer-Verwimp Schäfer-Verwimp Yuzawa Vital & Visnadi Yano Gradstein, Meneses & Arbe Gradstein & Costa Yano 1987 1989 1991 1991 1994 1995 2003 2003 2008 509 1345 523 525 2245 2095 975 984 1684 Stephani Stephani Hattori Kamimura Bonner Hattori Stephani 1894 1911 1952 1961 1965 1972 1985 228 686 529 199 655 538 928 Yuzawa Yamada & Iwatsuki 2000 2006 = Frullania gibbosa tonkinensis Spec Hepat 4: 339-340 Index Hepat 5: 452 J Hatt Bot Lab 45: 114 [Cat Colombia; doubtful record because based on coll Micholitz, who collected in Indomalesia] Stephani Icon Hepat Inedit [Chonanthelia] A- 52, microfiche 47 fig 3144-3145 [from type; "Andes Ibaque Tolima Micholitz 1901 1268 m."] J Hatt Bot Lab 63: 425-436 [morphology / systematic] J Hatt Bot Lab 67: 315 [new to Brazil] J Hatt Bot Lab 69: 153 [Brazil] J Hatt Bot Lab 70: 216 [syn sub Fr gibbosa] Trop Bryol 9: 70 [Brazil] J Hatt Bot Lab 78: 143 [annotated checklist of Brazil] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr gibbosa] Mem New York Bot Gard 87: 87 [Brazil] [syn sub Fr gibbosa] Boletim Instituto Botanica 19: 90 [syn sub Fr gibbosa; Brazil; catalog] Nees 1840 Steph. 1894 Typus: China. Tonkin sd, Balansa ex hb PC, in hb G Basionym: Distribution: China (Tonkin) spec nov - syn nov = Frullania meyeniana Hedwigia 33: 168-169 Spec Hepat 4: 649 J Hatt Bot Lab 8: 50 [syn sub Fr. meyeniana] J Hatt Bot Lab 24: 73 [sub F meyeniana] Index Hepat 5: 452 J Hatt Bot Lab 36: 115-16, fig 5 Stephani Icon Hepat Inedit [Diastaloba] B-85, microfiche 49 fig 3332 [from type; "Tonkin, Balansa leg"] 1659 Natural Environmental Sci Res 13: 18 [Japan, syn sub Fr meyeniana] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] Lindenb. 1845 Page 432 tortuosa Verd. 1929 Typus: Indonesia. Java, Prop. Praeanger, Mt. Pangerango, 1420m 27.4.1894, Schiffner 2001 Basionym: Distribution: Taiwan, Indonesia (Java) spec nov - syn nov Verdoorn Verdoorn Horikawa Hattori Kamimura Lai & Wang-Yang Miller, Whittier & Whittier Kuo & Chiang Piippo Bapna & Kachroo Yuzawa = Frullania hampeana 245 293 34 943 199 33 609 948 521 712 1660 Ann Bryol 2: 136-137, fig 5 Ann Bryol Suppl 1: 44-45 [syn nov sub Fr. hampeana] J Sci Hiroshima Univ, ser B, div 2, 2: 239 [Taiwan] Bull Tokyo Sci Mus 11: 144 [syn sub Fr. hampeana] J Hatt Bot Lab 24: 49 [syn sub Fr hampeana] Taiwania 21 (2): 168 [Index to Taiwan] Prod Flora Hepat Polynesiae: 140 Taiwania 33: 14 [Index of Taiwan] [as "tertuosa"] J Hatt Bot Lab 68: 48 [annotated catalogue of China] Hepaticology in India II: 174 [as "tortusa" syn sub Fr. monocera] Natural Environmental Sci Res 14: 23 [Japan, syn sub Fr monocera] Nees 1845 Frullania monocera tortusa 1929 1930 1934 1944 1961 1976 1983 1988 1990 2000 2001 (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Verd. by Bapna & Kachroo 2000 [orth var] Basionym: Distribution: Bapna & Kachroo = Frullania tortuosa 712 Hepaticology in India II: 174 Verd. 1929 Frullania hampeana Frullania monocera tosana 2000 Nees 1845 (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Horikawa 1943 Basionym: Distribution: Japan nom nov - syn nov Horikawa Hattori Kamimura Choe Yamada & Iwatsuki = Frullania tenella 1887] 1943 1952 1961 1980 2006 1283 529 199 1171 1470 Acta Phytotax Geobot 13: 213 [nom nov pro Fr tenella STEPH 1910] J Hatt Bot Lab 8: 29 [syn nov sub Fr diversitexta] J Hatt Bot Lab 24: 81 [sub F diversitexta] Illust Fl & Fauna Korea 24: 733 [sub Fr diversitexta] J Hatt Bot Lab 99: 33 [Catalog of Japan] Steph. 1910 [hom illeg] [non Sande Lacoste 1854, nec Caspary Frullania diversitexta Steph. 1897 Page 433 trabutiana Steph. 1910 Typus: Algeria, Portugal, Madeira ? Basionym: Distribution: Portugal, Madeira [Portugal], Algeria spec nov - syn nov Stephani Müller in Rabenhorst Müller in Rabenhorst Bonner Vanden Berghen Stephani 1910 1915 1958 1965 1976 1985 686 1064 1028 655 185 928 Sim-Sim 1989 6 = Frullania dilatata tranninhiana Spec Hepat 4: 368 Krypt Flora (2ed) 6, 2: 625, 628 [syn sub Fr dilatata] Krypt Flora (3ed) 6, 2: 1245 Index Hepat 5: 452 Bull Jard Bot Nat Belg 46 (1/2): 191 Stephani Icon Hepat Inedit [Galeiloba] C- 42, microfiche 50 fig 3397 ["Frull dilatata var, Coimbra, Moller 1885" - "auch Madeira (Fritze)" "auch Algier (Trabut) 1903"] Portug Acta Biol (B) 15: 363 [Portugal] 363 [syn sub Fr. dilatata] (L.) Dumort. 1835 Pocs ex Ninh 2003 [nom nud] Typus: Vietnam Basionym: Distribution: Vietnam - syn nov Ninh Pócs 2003 2019 = Frullania junghuhniana transfalcicula 1363 ASEAN Biodiversty 3 (1-2): 30 [new to Vietnam] 5243 pers. communication [= Fr junghuhniana, never effectively and validly published] Gottsche 1845 Clark & Svihla 1950 Typus: Ecuador, Prov. del Pichincha, in páramo, east slope of Volcán de Pichincha .., 10000-12000 ft Steere 9310 Basionym: Distribution: Ecuador, Colombia spec nov - syn nov Clark & Svihla Winkler Gradstein & Hekking Yuzawa = Frullania dusenii traversiana 1950 1976 1979 1991 335 5 483 525 Bryologist 53: 70-72, fig. 1-18 Rev Bryol Lichen 42 (3): 811 [Colombia] J Hatt Bot Lab 45: 114 [Cat Colombia] J Hatt Bot Lab 70: 214 Steph. 1905 Steph. 1910 Typus: Abyssinia, Schoah, Aferbaini prope Ankober 1885, Traversi 225 /? 255 pp in hb G, Basionym: Distribution: Ethiopia spec nov - syn nov Stephani Gola Cufodontis Bonner Vanden Berghen Stephani 1910 1914 1952 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania depressa 686 786 2562 655 185 928 Spec Hepat 4: 325 Annali di Botanica 13: 71 [Ethiopia] Phyton 4 (1-3): 77 [Ethiopia] Index Hepat 5: 453 Bull Jard Bot Nat Belg 46 (1/2): 206 Stephani Icon Hepat Inedit [Chonanthelia] A- 14, microfiche 46 fig 3084 [from type; "Abysinia Schoah leg Da Traversi 225"] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 226, fig 147(Q) from type [syn sub Fr depressa, West Africa] Mitt. 1863 Page 434 treubiana Schiffn. 1900 [nom nud] Typus: Indonesia. Java, Cibodas -.1.1895, Massart 1250 pp Basionym: Distribution: nom nud - syn nov Schiffner Verdoorn Verdoorn Bonner 1900 1929 1930 1965 = Frullania nepalensis trichomorpha - syn nov Beauverd 1924 Steph. ex Herzog 1916 [hom illeg] [non Steph. 1911] Beauverd in Stephani Bonner Uribe & Gradstein Uribe = Frullania brasiliensis trifaria Hedwigia 39: 208 [= Fr. nepalensis quoad pl. Javan.] Ann Bryol 2: 160 Ann Bryol Suppl 1: 51 [syn sub Fr. nepalensis] Index Hepat 5: 453 (Spreng.) Lehm. & Lindenb. 1845 Basionym: Frullania capilliformis Distribution: Bolivia nom nov 904 246 293 655 1924 1965 2003 2008 687 655 1071 1513 Spec Hepat 6: 547-548 Index Hepat 5: 454 Crypt Bryol 24 (3): 200 [syn sub Fr brsiliensis] Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Raddi 1822 (Reinw., Blume & Nees) Dumort. 1835 Basionym: Distribution: Dumortier Bonner Miller, Whittier & Whittier = Cheirolejeunea trifaria 1835 1965 1983 256 655 609 Rec d´Obs Jungerm., Tournay, 1: 13 Index Hepat 5: 454 [Tropical & Subtropical Asia & Oceania] Prod Flora Hepat Polynesiae: 140 (Reinw., Blume & Nees) Mizutani 1974 trinervis var. berteroana Lehm. & Lindenb. 1845 Basionym: Distribution: Lehmann & Lindenberg in G., L. 1845 & N. Bonner 1965 Yuzawa 1991 = Frullania trinervis 2850 Syn Hepat: 428 (No 36-) 655 525 Index Hepat 5: 454 J Hatt Bot Lab 70: 230 [syn sub Fr. trinervis] (Lehm.) Drège 1843 trinervis var. elongata (L. & L.) Lehm. & Lindenb. 1845 Basionym: Jungermannia elongata Distribution: Lehm. & Lindenb. 1835 Lehmann & Lindenberg in G., L. 1845 & N. Bonner 1965 Yuzawa 1991 = Frullania trinervis 2850 Syn Hepat: 428 (No 36) 655 525 Index Hepat 5: 455 J Hatt Bot Lab 70: 230 [syn sub Fr. trinervis] (Lehm.) Drège 1843 Page 435 trinervis var. obscura (L. & L.) Lehm. & Lindenb. 1845 Basionym: Distribution: Lehmann & Lindenberg in G., L. 1845 & N. Bonner 1965 = Frullania trinervis 2850 Syn Hepat: 428 (No 36 -) 655 Index Hepat 5: 455 (Lehm.) Drège 1843 trinervis var. Sprengelii Lehm. & Lindenb. 1845 Basionym: Frullania sprengelii Distribution: Gottsche 1845 [non Steph. 1894] Lehmann & Lindenberg in G., L. 1845 & N. Bonner 1965 = Frullania trinervis tristaniana 2850 Syn Hepat: 428 (No 36 -) 655 Index Hepat 5: 455 (Lehm.) Drège 1843 S.W.Arnell 1958 Typus: Tristan da Cunha: Round Hill, on Phylica 600m 18.1.1938 Christophersen & Mejland 1225 holotype: O, Isotypes JE, S Basionym: Distribution: - comb nov Arnell Myrdal 1958 2000 Vana & Engel 2013 = Frullania magellanica ssp. tristaniana trivaria 3902 Results Norwegian Sci Exped Tristan da Cunha 42:9-10, fig 5(a-d) 1265 http://www2.nrmse/kbo/saml/mossor/mossnamn.txt [hb S] [from textfile, Mossor artlist] 3868 Memoirs New York Bot Gard 105: 59-60 [Tristan da Cunha Isl.] (S.W. Arnell) Váňa & J.J. Engel 2013 (R., Blume & Nees) Steph. 1914 Basionym: Distribution: Stephani Bonner = Euosmolejeunea trivaria 1914 1965 1636 Spec Hepat 5: 592 [syn sub Euosmolejeunea] 655 Index Hepat 5: 592 (R., B. & N.) Steph. 1914 Page 436 trollii Herzog 1942 Typus: Colombia. Feuchter Buschgürtel über Bogota, 3000-3200m 1929, C. Troll 2086 in hb JE [isotypus] Basionym: Distribution: Colombia spec nov syn nov Herzog Bonner Gradstein & Hekking Uribe & Gradstein Uribe & Rangel-Ch Müller & Zündorf 1942 1965 1979 1998 2000 2007 -syn nov Bernal, Gradstein & Celis Gradstein 2016 2021 = Frullania paradoxa truncata 874 655 483 707 1090 1442 Beih Bot Centralbl 61 B: 575-577, fig 10(a-m) [Columbia] Index Hepat 5: 455 J Hatt Bot Lab 45: 114 [Cat Colombia] Bryophyt Bibl 53: 38 [Catalogue Colombia] Colombia, diversidad biótica III, Bogota: 446 [Colombia] http://www2.unijena.de/biologie/spezbot/herbar/bryotypes/Uebersicht%20F.htm [Typusbelege; syn sub Fr. paradoxa L & L, annotation Stotler] 4290 Catálogo de plantas y líquenes de Colombia. Vol. I. [dubious species] 5642 Nem NY Bot Gard 121: 375 [syn sub Frullania paradoxa - remained unpublished] Lehm. & Lindenb. 1845 von Konrat & Braggins in von Konrat & Pickering 2003 [nom nud] Basionym: Distribution: New Zealand (South Isl: Coromandel, Table Mt & Westland, Stewart Isl) Konrat & Pickering 2003 Asakawa, Toyota, Konrat & Braggins Konrat & Brennan 2003 2004 1000 https://www.discoverlife.org/mp/20q?search=Frullania+truncata&guide =Plant_genera#http://liverworts.org/jubulaceae/frullania/truncata/trunca ta_prof.shtml#desc [New Zealand] 1054 Phytochemestry 62: 444 [chemosystematic; undescribed species] 1164 http://www.liverworts.org/dich_key/frul_key.shtml, 6th of 6 pages [Key to Frullania of New Zealand; Coromandel (Table Mt), Westland, Stewart Isl]] = Page 437 truncatifolia Steph. 1910 Typus: Japan. Sikoku, Kochi Pref., Prov Tosa: Takaoka-gun, Mt. Yokogura -.8.1901, T. Yoshinaga 28 in hb G [holotypus], in hb TNS [isotypus] in hb HIRO ? Japan. Honshu, Aomori, oct. 1902, U. Faurie 1248 [Isotypus?] [=Frullania muscicola Steph.] Basionym: Distribution: spec nov - syn nov Stephani Iishiba Oti / Ochi Tatuno Kamimura Hattori Hattori Hattori Hattori & Noguchi Bonner Hattori Stephani 1910 1912 1950 1952 1952 1952 1953 1955 1960 1965 1975 1985 Inoue Bapna & Kachroo N.N. [National Science Museum Tokyo] Yuzawa NN 1987 2000 2000 Yamada & Iwatsuki 2006 = Frullania muscicola truncatifolia 2001 2003 686 888 Spec Hepat 4: 398-399 Bot Mag Tokyo 26: 36 [Japan, with key in Japanese] Shizen to Simmon / Jinmon 1: 70 [according p#873] 1697 J Sci Hiroshima Univ ser B, div 2 (Bot), 6(2): 51-57 [sex chromosomes] 873 Contrib Fl Hepat Shikokuensem, Kochi: 131 529 J Hatt Bot Lab 8: 31 530 J Hatt Bot Lab 10: 38-39, fig 50(A-T) 1683 J Hatt Bot Lab 15: 92 [on serpentine rocks] 965 Index spec Typ herb Japon, Bryophyta: 18 655 Index Hepat 5: 455-456 410 Misc Bryol Lichen 7 (2): 28 928 Stephani Icon Hepat Inedit [Galeiloba] C-255, microfiche 53 fig 3675 ["Japan, leg Yoshinaga 28, Mt Yokogura"] - 3676 ["1752 (Faurie)"] 1155 Index type specimens in Nat Sci Mus, Tokyo: 15 712 Hepaticology in India II: 175 [sub Fr. muscicola] 995 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 1660 Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] 1397 Types of Nat Hist mus Hiroshima http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] Steph. 1894 Steph. ex Yoshinaga 1903 [nom nud] Basionym: Distribution: Stephani in Yoshinaga Sawada 1903 1909 Kamimura Choe Wu in Li, Gao, Wu et al. 1961 1980 1985 Yamada & Iwatsuki 2006 = Frullania truncatifolia 801 930 Bot Mag Tokyo 17 (no 193): 38 [nom nud] Bot Mag Tokyo 23 (no 268): 178-183, fig 26 [key to Japanese species, in Japanese] 199 J Hatt Bot Lab 24: 21 [syn sub Fr. muscicola] 1171 Illust Fl & Fauna Korea 24: 732 [sub Fr brittoniae truncatifolia] 1169 Bryoflora Xizang (Tibet), Science Press, Beijing: 523 [sub Fr muscicola] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] Steph. 1910 Frullania muscicola Steph. 1894 truncatifolia var. fo. fragilis Ochi ? Oti 1950 Typus: Japan. Prov. Jyo, Saijo City Basionym: Distribution: Oti / Ochi Oti / Ochi Kamimura Bonner = Frullania muscicola 1950 1950 1952 1965 873 655 Shizen to Simmon / Jinmon 1: 71 [according p#873] Shizen to Simmon / Jinmon 1: 71 [according p#655] Contrib Fl Hepat Shikokuensem, Kochi: 132 Index Hepat 5: 456 Steph. 1894 Page 438 truncatifolia var. transiens S.Hatt. ex Kamim. 1952 [nom nud] Typus: Japan. Prov. Tosa, Inabu-muro sd, H. Inoue Basionym: Distribution: nom nud - syn nov Kamimura Kamimura Bonner Bapna & Kachroo Yuzawa = Frullania muscicola truncatiloba 1952 1961 1965 2000 2001 873 199 655 712 1660 Contrib Fl Hepat Shikokuensem, Kochi: 131 [nom nud] J Hatt Bot Lab 24: 21 [syn nov sub Fr. muscicola] Index Hepat 5: 456 Hepaticology in India II: 175 [syn sub Fr. muscicola] Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] Steph. 1894 Steph. 1910 [non Vanden Berghen 1961] Typus: Guinea. Guinea gallica [French Guinea] sd, Pobeguin 175 pp in hb G 1689 [holotypus] Basionym: Distribution: French Guinea, Cameroon, Uganda, Bolivia spec nov - syn nov Stephani Stephani Pearson Bonner Augier Vanden Berghen Stephani 1910 1916 1924 1965 1974 1976 1985 Hattori Wigginton 1988 2002 Gradstein, Meneses & Arbe 2003 Wigginton Wigginton 2004 2004 = Frullania africana Spec Hepat 4: 322 Biblioth Bot 87: 240 [Bolivia] Arkiv Botanik 19 (5): 9 [Mt. Elgon in Kenia/Uganda] Index Hepat 5: 456 Annales de la Faculté des Sciences du Cameroun 17: 85 [Cameroun] Bull Jard Bot Nat Belg 46 (1/2): 151 [syn sub Fr. africana STEPH 1891] Stephani Icon Hepat Inedit [Chonanthelia] A- 8, microfiche 46 fig 3076-3077 [from type; "Guinea gallica leg Pobeguin 175"] 516 J Hatt Bot Lab 65: 443 [syn nov, syn sub Fr. riojaneirensis] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 975 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. riojaneirensis; according Yuzawa 1991] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 226 [syn sub Fr riojaneirensis, West Africa] Steph. 1891 Frullania rio-janeirensis truncatiloba 686 222 795 655 821 185 928 (Raddi) Ångstr. 1876 [auct non Steph. 1910] Vanden Berghen 1961 Basionym: Distribution: - syn nov Vanden Berghen Vanden Berghen Wigginton 1961 1976 2002 Wigginton Wigginton 2004 2004 = Frullania depressa 1507 Bull Soc Roy Bot Belg 93 (1-2): 71-72, fig. 6(a-g) [Urundi] 185 Bull Jard Bot Nat Belg 46 (1/2): 206 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 226 [syn sub Fr depressa, West Africa] Mitt. 1863 Page 439 tsukushiensis Horikawa 1929 Typus: Japan. Kyushu. Prov Hyuga, Isl. Aoshima, Mt. Aoidake 9.4.1927, Y. Horikawa 452 - Y. Horikawa 532 in hb HIRO (Y. Horikawa 452) [lectotypus by Zhu & So 2001] in hb HIRO (Y. Horikawa 532) [syntypus] Basionym: Distribution: Taiwan, Japan spec nov - syn nov Horikawa Verdoorn Verdoorn Horikawa Hattori Hattori & Noguchi Kamimura Bonner Hattori & Lin Kuo & Chiang Piippo 1929 1932 1934 1934 1944 1960 1961 1965 1985 1988 1990 Piippo Yuzawa Zhu & So NN 1990 2000 2001 2003 NN 2006 Yamada & Iwatsuki 2006 = Frullania aoshimensis tuberculosa 924 554 240 34 943 965 199 655 505 948 521 Sci Rep Tohoku Imp Univ, ser 4, Biol 4: 66-68, fig 5(1-10) [Japan] Nederl Kruidk Arch Jrg 1932 (2): 495 Ann Bryol Suppl 4: 197-198 [syn sub Fr. aoshimensis] J Sci Hiroshima Univ, ser B, div 2, 2: 237 [Japan, Taiwan] Bull Tokyo Sci Mus 11: 142 [syn sub Fr. aoshimensis] Index spec Typ herb Japon, Bryophyta: 18 [holotype] J Hatt Bot Lab 24: 69 [syn sub Fr aoshimiensis] Index Hepat 5: 456 J Hatt Bot Lab 59: 157 [syn sub Fr. aoshimiensis] Taiwania 33: 14 [Index of Taiwan] J Hatt Bot Lab 68: 139 annotation nr 138 [according #505 as syn sub Fr. aoshimiensis] 521 J Hatt Bot Lab 68: 48 [annotated catalogue of China] 1659 Natural Environmental Sci Res 13: 14 [Japan, syn sub Fr aoshimensis] 808 Nova Hedwigia Beih 121: 73 [syn sub Fr aoshimiensis] 1397 Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] http://www.digital-museum.hiroshimau.ac.jp/~museum/plant/index.php?Frullania_tsukushiensis_type 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] Horik. 1929 Gottsche ex Ångstr. 1876 [nom inval; Art 32.1(c)] {Brazil} Typus: Brasil, Minas Gerais, Caldas sd, G.A. Lindberg & S. Henschen invenerunt sn Basionym: Distribution: Brazil nom inval Ångström Yano Gradstein & Costa Yano 1876 1984 2003 2008 579 501 984 1684 Öfversigt K Vetensk Akad Förh 33 (7): 88 [Brazil] J Hatt Bot Lab 56: 507 [checklist Brazil] Mem New York Bot Gard 87: 91 [Brazil] Boletim Instituto Botanica 19: 33 [Brazil; catalog] = tubifera Taylor ex Gottsche 1845 [pro syn] Typus: in hb TAYLOR Basionym: Distribution: Gottsche in G., L. & N. = Frullania cylindrica 1845 2850 Syn Hepat: 458 [syn sub Fr. cylindrica) Gottsche ex Lehm. 1844 Frullania brasiliensis Raddi 1822 Page 440 tubiflora Spruce 1884 Typus: Ecuador. "Flumen Pastasa, Antombos, 1500m" sd, Spruce sn in hb G (hb STEPHANI 86) in hb E E00002589 [isotypus] Basionym: Distribution: Ecuador spec nov - syn nov - syn nov Spruce Stephani Bonner Stotler Stotler Stephani 1884 1911 1965 1969 1970 1985 = Frullania macrocephala tuerckheimii 243 686 655 4761 194 928 Trans Proc Bot Soc Edinburgh 15: 44 & 49 Spec Hepat 4: 509 Index Hepat 5: 456-457 Nova Hedwigia 18: 439 [syn sub Fr. macrocephala] The genus Frullania, separata: 439 [syn sub Fr. macrocephala] Stephani Icon Hepat Inedit [Thyopsiella] F- 96, microfiche 58 fig 4016 ["Andes quitenses, leg Spruce, H.a.a. p 49"] (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Steph. 1911 Typus: Guatemala. Baja Verapaz, Santa Rosa, 1600m - "in truncis quercinis, inter Macromitria." -.12.1906, H.v.Türckheim 5057 in hb NY 1047688 (ex BRYOTHECA E. LEVIER 5054 - Dec 1906, H. v. Türckheim sn) hb LEVIER 5054, in hb G [Stotler 1969: #5057 in hb G as typus, #5054 in hb NY] Basionym: Distribution: spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Stephani 1911 1965 1969 1970 1985 686 655 4761 194 928 Gradstein, Meneses & Arbe 2003 975 = Frullania montagnei turbata Typus: Brazil Spec Hepat 4: 508 [as "türckheimii"] Index Hepat 5: 457 Nova Hedwigia 18: 452-453 [syn sub Fr. montagnei] The genus Frullania, separata: 452-453 [syn sub Fr. montagnei] Stephani Icon Hepat Inedit [Thyopsiella] F- 97, microfiche 58 fig 4017 ["Guatemala (Baja Verpaz), H. Türckheim Dez 1906, Levier 5054"] 4018 J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. montagnei] Gottsche 1845 Steph. 1911 1910, P. Dusen 8461 in hb G 20625 [holotypus] Basionym: Distribution: Ecuador, Brazil spec nov -syn nov Stephani Herzog Bonner Yano Stephani 1911 1925 1965 1984 1985 Yano Gradstein & Costa Gradstein & Pinheiro da Silva Uribe 1999 2003 2003 2008 Yano 2008 = Frullania atrata 686 1586 655 501 928 Spec Hepat 4: 597 Arquivos de botânica do Estado de Sao Paulo 1 (2): 47 [Brazil] Index Hepat 5: 457 J Hatt Bot Lab 56: 507 [checklist Brazil] Stephani Icon Hepat Inedit [Meteriopsis] E-40, microfiche 55 fig 3827 ["Brasilia, Dusén 8461"] 1263 Briophyta do estado de São Paulo. Lista provisória [Brazil] 984 Mem New York Bot Gard 87: 83 [Brazil] [syn sub Fr. atrata] vide #1513 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr atrata; Brazil; catalog] (Sw.) Nees ex Mont. 1839 Page 441 ubangiensis Steph. 1911 Typus: Africa: Afrique centrale, bords de l´Oubangui -.8.1902, A. Chevalier sn hb CORBIERE 81, in hb G 5195 [holotypus] & hb PC [isotypus] Basionym: Distribution: spec nov - syn nov Stephani Bonner Vanden Berghen Stephani 1911 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania trinervis udarii 686 655 185 928 Spec Hepat 4: 674-675 Index Hepat 5: 455 Bull Jard Bot Nat Belg 46 (1/2): 199 Stephani Icon Hepat Inedit [Chonanthelia] A- 9, microfiche 46 fig 3078 [from type?; "leg Chevalier"] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 227 [syn sub Fr trinervis, West Africa] (Lehm.) Drège 1843 Ajit P.Singh 2002 [not effectively published] Basionym: Distribution: spec nov Singh 2002 Crosby & Magill 2006 = Frullania udarii uematsuana 4886 Studies on Liverworts (Bryophyta) of Khasi and Jaintia Hills: Meghalaya. 268-271, pl 54(1-18), map 11 - Unpublished thesis. 1429 Index of Bryophytes: 3 [not effectively published] V.Nath & Ajit P.Singh 2006 Beauverd 1924 Typus: Japan. Sendai 12.4.1907, Uematsu 49 in hb G Basionym: Distribution: Japan, Korea spec nov - syn nov Beauverd in Stephani Hattori Bonner Stephani 1924 1953 1965 1985 Bapna & Kachroo Yuzawa Yamada & Iwatsuki 2000 2001 2006 = Frullania truncatifolia Spec Hepat 6: 543 [nom nov pro hb specimen] J Hatt Bot Lab 10: 38 [syn sub Fr. truncatifolia] Index Hepat 5: 455 Stephani Icon Hepat Inedit [Galeiloba] C-249, microfiche 53 fig 3669 ["Japonia, Sendai, leg Uematsu No 49, 1907"] 712 Hepaticology in India II: 175 [ as "uematuana" syn sub Fr. muscicola] 1660 Natural Environmental Sci Res 14: 25 [Japan, syn sub Fr muscicola] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] Steph. 1910 Frullania muscicola uematuana 687 530 655 928 Steph. 1894 Beauverd ex Steph. by Kamim. 1961 [orth var] Basionym: Distribution: Kamimura & Hattori Bapna & Kachroo = Frullania uematsuana 1961 2000 199 712 J Hatt Bot Lab 24: 21 [syn sub Fr. muscicola] Hepaticology in India II: 175 Beauverd 1924 Frullania truncatifolia Frullania muscicola Steph. 1910 Steph. 1894 Page 442 ulotricha Taylor 1846 Typus: A America. sd, Humboldt 102 in hb HOOKER, in hb FH, W, in hb S (B24322-24324) [typus] in hb GLM (hb LEHMANN, ex hb HOOKER) 05955 Basionym: Distribution: spec nov - syn nov Taylor Taylor in G., L. & N. Stephani Stephani 1846 1847 1911 1985 292 2848 686 928 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 2002 782 = Frullania paradoxa ultima London J Bot 5: 405 Syn Hepat: 780 Spec Hepat 4: 533 Stephani Icon Hepat Inedit [Thyopsiella] F-121, microfiche 58 fig 4048 ["syn: Frull ulotricha Tayl, Orig ex Herb Gottsche, Peruvia"] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] Lehm. & Lindenb. 1845 Beauverd 1924 [non Steph. 1924] {Neu Guinea} Typus: PNG: [Nova] Guinea sd, Ledermann, sn hb G [holotypus] as "Frullania renistipula STEPH" Basionym: Frullania renistipula Steph. 1924 [pro syn] {Neu Guinea} [non Steph. 1910, from Madagascar] Distribution: nom nov - syn nov Beauverd in Stephani Verdoorn Bonner Grolle & Piippo = Frullania heteromorpha 1924 1930 1965 1984 Spec Hepat 6: 554 Nova Guinea 14 (4): 541 [syn sub Fr. heteromorpha fo. mammillosa] Index Hepat 5: 458 Acta Bot Fennica 125: 24 [Cat. West Melanesia] fo. mamillosa Verd. 1930 Frullania heteromorpha unciflora 687 549 655 143 Schiffn. 1889 Gottsche ex Bonner 1965 [nom herb] {N America} Typus: N. America [ Basionym: Distribution: N-America Bonner 1965 655 Index Hepat 5: 458-459 = unciflora var. californica Gottsche ex Bol. 1870 [nom nud] Basionym: Distribution: - syn nov Gottsche ex Bolander Gottsche ex Bolander Evans Frye & Clark Hattori in Hara = Frullania franciscana 1870 1870 1897 1947 1966 3472 1669 237 204 841 California Medical Gazette 2: 184 Cat Plants San Francisco: 40 [name only] Trans Connet Acad Arts Sci 10 (1): 26 Univ Wash Publ Biol 6 (5): 753 [sub Fr franciscana] Flora of Eastern Himalaya: 528 [nom nud] M. Howe 1894 Page 443 uncifolia Gottsche ex Steph. 1911 Typus: Tahiti sd, Vesco sn ex hb BESCHERELLE, in hb PC & G Basionym: Distribution: Society Isl (Tahiti) spec nov - syn nov Gottsche ex Stephani Verdoorn Verdoorn Clark Bonner Miller, Whittier & Whittier Stephani = Frullania apiculata undulata 1911 1930 1930 1955 1965 1983 1985 686 293 551 369 655 609 928 Spec Hepat 4: 540 Ann Bryol Suppl 1: 97 [syn sub Fr. apiculata] Nederl Kruidk Arch Jrg 1930 (2): Bryologist 58: 68 [syn sub Fr. apiculata] Index Hepat 5: 459 Prod Flora Hepat Polynesiae: 140 Stephani Icon Hepat Inedit [Thyopsiella] F-157, microfiche 59 fig 4097 ["Tahiti, Vesco, Bescherelle misit (Original)"] (Reinw., Blume & Nees) Nees 1845 Kamim. 1961 Typus: Japan. Shikoku, Kochi Pref., Mt. Kajigamori, Nishitoyonaga-mura, Nagaoka-gun.1100 m 3.7.1950, M. Hara 1365, 1385 in hb NICH, in hb KOCH (Hara-1385) [isotypus] Basionym: Distribution: Japan spec nov - comb. nov. Kamimura Bonner Inoue NN 1961 1965 1966 1997 Yamada & Iwatsuki Hentschel et von Konrat 2006 2015 199 655 1703 639 J Hatt Bot Lab 24: 50-52, fig. 18(1-11) Index Hepat 5: 459 J Jap Bot 41 (1): 15 [basionym of Fr hampeana var undulata] http://plants.cc.kochi-u.ac.jp/www/herbarium [accessed 27.5.2011] [isotype] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] 4097 Phytotaxa 220 (2): 136, 2015, Bas.: Frullania undulata Kamim = Frullania monocera var. undulata (Kamim.) Hentschel & von Konrat 2015 unduliloba Herzog in hb US 1996 [nom herb] {Costa Rica} Typus: Costa Rica. Cartago, Pejivalle 7.2.1927-8.2.1996, Standley 46743 in hb US Basionym: Distribution: Costa Rica NN 1996 632 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 33 of 35 [in hb US] = Page 444 urbanii Steph. 1911 Typus: Guadeloupe. sd, Duss 329 in hb G Basionym: Distribution: Guadeloupe spec nov - syn nov - syn nov Stephani Pagán Bonner Stotler Stotler Stephani 1911 1942 1965 1969 1970 1985 Lavocat Bernard & SchäferVerwimp 2011 = Frullania neesii Spec Hepat 4: 527 Bryologist 45(4): 109 [Guadeloupe] Index Hepat 5: 459 Nova Hedwigia 18: 450 [syn sub Fr. neesii] The genus Frullania, separata: 450 [syn sub Fr. neesii] Stephani Icon Hepat Inedit [Thyopsiella] F-127, microfiche 58 fig 4054 ["Guadeloupe, leg Duss 329 ex parte"] 3130 Cryptogamie, Bryologie 32(3): 243 [Guadeloupe] Lindenb. 1845 Frullania kunzei usagara 686 3152 655 4761 194 928 (Lehm. & Lindenb.) Lehm. & Lindenb. 1845 Mitt. 1886 Typus: East Africa: Usagara Mts. sd, Hannington in hb NY [holotypus], in hb G: Basionym: Distribution: spec nov - syn nov Mitten Stephani in Engler Stephani Demaret & Vanden Berghen Bonner Vanden Berghen Stephani 1886 1895 1910 1947 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania squarrosa J Linn Soc Bot 22: 326, fig. 19(12-14) Die Pflanzenwelt Ost-Afrikas 2 (C): 65 [Usagara, Ugogo] Spec Hepat 4: 372 Bull Jard Bot Etat Bruxelles 18: 238 [syn sub Fr squarrosa] Index Hepat 5: 459 Bull Jard Bot Nat Belg 46 (1/2): 163 Stephani Icon Hepat Inedit [Galeiloba] C- 43, microfiche 50 fig 3398 ["J L Soc vol 22"] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 229, fig 152(K,L,N) from type [syn sub Fr ericoides, West Africa] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides usambarensis 1910 1481 686 1101 655 185 928 (Nees) Mont. 1839 Steph. 1910 Typus: Tanzania. Usambara Mts sd, G. Bauer sn hb LEVIER 5119 & 5122, in hb G 17326 [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Stephani in Brunnthaler Bonner Vanden Berghen Stephani 1910 1913 1965 1976 1985 Wigginton 2002 Wigginton Wigginton 2004 2004 = Frullania spongiosa 686 219 655 185 928 Spec Hepat 4: 375-376 Denkschr K Akad Wiss, Math Nat Kl, Wien 88: 729 [Tansania Index Hepat 5: 460 Bull Jard Bot Nat Belg 46 (1/2): 167 & 171 [syn sub Fr. spongiosa] Stephani Icon Hepat Inedit [Galeiloba] C- 44, microfiche 50 fig 3399 [from type; "Usambara, G. Baur, Levier 5119 5122"] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 234 [syn sub Fr spongiosa,West Africa] Steph. 1894 Page 445 usamiensis Steph. 1910 [nom superfl] Basionym: Distribution: Stephani 1910 = Frullania usamiensis usamiensis Spec Hepat 4: 351-352 [spec nov = nom superfl] Steph. 1897 fo. yakushimensis (Horikawa) S.Hatt. 1951 Basionym: Frullania yakushimensis Distribution: Japan comb nov - syn nov 686 Horikawa 1934 Hattori Hattori Kamimura Bonner Choe Yuzawa Yamada & Iwatsuki = Frullania usamiensis utacamundeana 1951 1952 1961 1965 1980 2001 2006 528 529 199 655 1171 1660 1470 J Hatt Bot Lab 5: 67 J Hatt Bot Lab 8: 31 [syn sub Fr. usamiensis] J Hatt Bot Lab 24: 43 [syn sub Fr usamiensis] Index Hepat 5: 460 Illust Fl & Fauna Korea 24: 736 [sub Fr. usamiensis; South Korea] Natural Environmental Sci Res 14: 38 [Japan, syn sub Fr usamiensis] J Hatt Bot Lab 99: 33 [Catalog of Japan] Steph. 1897 [nom herb] Basionym: Distribution: Hattori 1975 Bull. Natn. Sci. Mus., Ser. B (Bot.), 1 (2): 73 = utriculata var. powelliana (Steph.) Verd. 1930 Basionym: Frullania powelliana Steph. 1894 Distribution: Vanuatu, New Caledonia, Micronesia (Ponape), Samoa (?Savaii, ?Tutuila) comb nov - syn nov Verdoorn Miller Grolle & Schultze-Motel Hattori Miller, Whittier & Whittier Grolle & Piippo = Frullania utriculata 1930 1960 1972 1980 1983 1984 293 920 537 490 609 143 Ann Bryol Suppl 1: 169-170 Bryologist 63: 121 [Micronesia] J Hatt Bot Lab 36: 84 [Samoa] J Hatt Bot Lab 47: 183 Prod Flora Hepat Polynesiae: 141 Acta Bot Fennica 125: 25 [Cat. West Melanesia] Steph. 1894 Page 446 uvifera Horikawa 1934 Typus: Japan. Bonin, Isl. Chichijima, Mt. Chu-o. "Mt. Chûôsan" 27.7.1930, Y. Horikawa 1949 in hb HIRO [holotypus] Basionym: Distribution: Japan (Honshu, Kyushu, Yakushima), Ryukyu (Amami, Okinawa, Tokunoshima), Bonin Isl (Chichijima, Hahajima) spec nov - syn nov Horikawa Hattori 1934 1944 34 943 Hattori Hattori Iwatsuki & Hattori Hattori, Iwatsuki & Kanno Hattori & Noguchi Bonner Shin Inoue & Iwatsuki Shin Hattori Kodama Miller, Whittier & Whittier Yuzawa NN 1951 1952 1956 1956 1960 1965 1968 1969 1970 1972 1972 1983 2000 2003 528 529 861 862 965 655 844 61 302 538 1375 609 1659 1397 Yamada & Iwatsuki 2006 1470 = Frullania ramuligera uvifera J Sci Hiroshima Univ, ser B, div 2, 2: 239, pl 18(24-29) Bull Tokyo Sci Mus 11: 153-155, fig 97(a-h) [Japan, key to Japanese species] J Hatt Bot Lab 5: 68 J Hatt Bot Lab 8: 31 J Hatt Bot Lab 16: 87 J Hatt Bot Lab 16: 93 Index spec Typ herb Japon, Bryophyta: 18 Index Hepat 5: 461 Sci Rep Kagoshima Univ 17: 204 [Ryukyu Arch.] Bull Natn Sci Mus Tokyo 12 (2): 303-304 Sci Rep Kagoshima Univ 19: 38 [Ryukyu: Okinawa] J Hatt Bot Lab 36: 117-119, fig 6 [syn sub Fr. ramuligera] Spec Publ Osaka Mus Nat Hist 4: 137-138, fig 66(a-f) [Central Japan] Prod Flora Hepat Polynesiae: 141 [as a valid species] Natural Environmental Sci Res 13: 10 [Japan, syn sub Fr ramuligera] Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] J Hatt Bot Lab 99: 33 [Catalog of Japan] (Nees) Mont. 1843 [auct. non Horikawa] Kamim. 1952 Basionym: Distribution: auct - syn nov Kamimura Kamimura Iwatsuki & Hattori Hattori, Iwatsuki & Kanno Kamimura Hattori = Frullania hypoleuca 1952 1952 1956 1956 1961 1982 873 873 861 862 199 495 Contrib Fl Hepat Shikokuensem, Kochi: 132 Contrib Fl Hepat Shikokuensem, Kochi: 132 J Hatt Bot Lab 16: 87 J Hatt Bot Lab 16: 93 J Hatt Bot Lab 24: 79-80, fig 26(19-45), map 8 J Hatt Bot Lab 51: 236 Nees 1843 vaginata var. β nigricans Gottsche, Lindenb. & Nees 1846 Basionym: Distribution: Indonesia (Java) var nov - syn nov Nees in G., L. & N. Sande Lacoste Schiffner 1846 1857 1898 2849 Syn Hepat: 465 430 Verh Konikl Akad Wetenschappen 5: 92 (No 238 b) 831 Conspect Hepat Archip Indici, Batavia: 347-348 [syn nov] = Frullania vaginata var. nigricans (Gottsche, Lindenb. & Nees) Schiffn. 1898 valadina Gottsche by Miller, Whittier & Whittier 1983 [orth var] Basionym: Distribution: Miller, Whittier & Whittier = Frullania baladina 1983 609 Prod Flora Hepat Polynesiae: 140 [not "150") Gottsche 1894 Page 447 valida var. fusco-virens (Steph.) S.Hatt. 1980 Basionym: Distribution: comb nov - syn nov Hattori Hattori & Lin Yamada & Choe = Frullania fuscovirens vandenhammenii 1980 1985 1997 489 505 717 J Hatt Bot Lab 47: 119-120, fig 236 [comb. nov pro Fr. fusco-virens] J Hatt Bot Lab 59: 134 J Hatt Bot Lab 81: 289 [Korean Peninsula] Steph. 1910 Haarbrink 1981 Typus: COLOMBIA, BOYACÁ, Localidad: Carretera Arcabuco-Villa de Leiva; entre La Capilla y Los Naranjos; 500 m arriba de la carretera, Bosque con Quercetum, 2550 m 1.9.1967, Jaramillo-M., R.; Murillo, M. T.; Van der Hammen 2482 in hb COL COL000294377 - 203099 [holotypus] in hb U Basionym: Distribution: Colombia spec nov comb nov Haarbrink Yuzawa 1981 1991 49 525 Lindbergia 7 (1): 56-57, fig. 14-20 [Colombia, High Andes] J Hatt Bot Lab 70: 233-234, fig 27 [Frullania winteri Steph. var. vanderhammenii (Haarbrink) Yuzawa, comb. nov.] = Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa 1991 vandercystii Steph. by S.W.Arnell 1963 [orth var] Basionym: Distribution: Arnell 1963 = Frullania vanderystii Hepat S Africa: 147 Steph. 1910 Frullania cardotii Steph. 1910 Frullania obscurifolia vanderystii 822 Mitt. 1879 Steph. 1910 Typus: Zaire. "Congo Belge, Mayidi" 1907, Vanderyst 140 hb CARDOT 140, in hb G 5196 [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Demaret & Vanden Berghen Arnell Bonner Vanden Berghen Stephani 1910 1947 1963 1965 1976 1985 Wigginton & Grolle Wigginton 1996 2002 Wigginton Wigginton 2004 2004 = Frullania cardotii 686 1101 822 655 185 928 Spec Hepat 4: 366 Bull Jard Bot Etat Bruxelles 18: 233 [sub Fr cardoti] Hepat S Africa: 147 [syn sub Fr. borgenii] Index Hepat 5: 462 Bull Jard Bot Nat Belg 46 (1/2): 143 Stephani Icon Hepat Inedit [Galeiloba] C- 45, microfiche 50 fig 3400 [from type; "olim Frullania nigra Steph, Congo, Mayidi, leg Vanderyst 1907 (Cardot 140)"] 696 Bryophyt Bibl 50: 93 [= syn sub Fr. cardoti = syn sub Fr. obscurifolia] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 233 [syn sub Fr obscurifolia, West Africa] Steph. 1910 Frullania obscurifolia Mitt. 1879 Page 448 varia Gottsche 1882 Typus: Madagascar. "in siva Ambaranavaranututa stirps unica" 6.12.1899, Rutenberg 1504 in hb G sn; Stephani, Icones Hepat Inedit 1632 Basionym: Distribution: East Africa, Madagascar spec nov - syn nov Gottsche Stephani in Renauld & Cardot Pearson 254 Abh Naturwiss Ver Bremen 7 (3): 363-364 [Madagascar] 1453 Rev Bryol 18 (4): 56 [Madagascar] 1921 Christiania Videnskabs-Selskabs Forhandlinger 1890 (2): 6 [Madagascar] Stephani in Levier Stephani Demaret & Vanden Berghen Bonner Vanden Berghen Stephani 1901 1911 1950 1965 1976 1985 1456 686 60 655 185 928 Grolle Wigginton 1995 2002 250 1012 Wigginton Wigginton Uribe 2004 2004 2008 1312 1172 1513 = Frullania angulata varians 1882 1891 1891 Icon Zeitschrift Forhandlinger i Videnskaps-Selskabet i Kristiania 1890 (2): 4 Rev Bryol 28 (5): 94 [Madagascar] Spec Hepat 4: 593 Bull Jard Bot Etat Bruxelles 20: 5 Index Hepat 5: 462 Bull Jard Bot Nat Belg 46 (1/2): 70 Stephani Icon Hepat Inedit [Meteriopsis] E- 9, microfiche 55 fig 3776 ["Madagascar, Rutenberg, Chenagon legit"] - 3777 Bryophyt Bibl 48: 62 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] Sript Bot Belg 30: 214 [syn sub Fr angulata, West Africa] Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Mitt. 1863 J.B.Jack ex Bonner 1965 [nom herb] Typus: Fiji: Viti-Inseln, Ovalau, Mont. reg. 1864, E. Graeffe 655 in hb G (ex hb JACK) [sub Fr. deflexa] Basionym: Distribution: Fiji Bonner Miller, Whittier & Whittier = Frullania deflexa verdiana 1965 1983 655 609 Index Hepat 5: 463 Prod Flora Hepat Polynesiae: 141 [? syn sub Fr. deflexa] Mitt. 1862 S.Hatt. by von Konrat & Pickering 2003 [orth var] Basionym: Distribution: = Frullania verdoorniana S.Hatt. 1973 Page 449 verdoorniana var. wilhelmensis S.Hatt. 1973 Typus: Papua New Guinea. Simbu, Mt. Wilhelm, 13500-13800ft 6.6.1967, R.M. Schuster 67-6807 in hb NICH [holotypus] in hb MASS & hb JE [isotypus] Basionym: Distribution: var nov Hattori Hattori Grolle & Piippo Mizutani, Hasegawa & Iwatsuki = Frullania macgregorii vesiculosa 1973 1974 1984 2009 542 546 143 2112 J Hatt Bot Lab 37: 124-125, fig 69 J Hatt Bot Lab 38: 262 Acta Bot Fennica 125: 25 [Cat. West Melanesia] Type sepecimens Hatt Bot Lab (NICH) 38: Holotype. New Guinea, Central Highlands, Mt. Wilhelm, rocky alpine tussock-grassland.. 13500-13800 ft. 8 Aug. 1967. R. M. Schuster 67-6807 (NICH 314858). Steph. 1894 Steph. 1911 Typus: Indonesia. Java sd, Seubert sn in hb G Basionym: Distribution: spec nov - syn nov Stephani Verdoorn 1911 1930 Verdoorn Frye & Clark Kamimura Bonner Miller, Whittier & Whittier Stephani 1930 1947 1961 1965 1983 1985 Bapna & Kachroo Yuzawa 2000 2001 = Frullania squarrosa Frullania ericoides 686 293 Spec Hepat 4: 457 Ann Bryol Suppl 1: 34 & 38 [syn sub Fr. squarrosa var planescens fo campanuloides] 551 Nederl Kruidk Arch Jrg 1930 (2): 204 Univ Wash Publ Biol 6 (5): 776 [sub Fr squarrosa] 199 J Hatt Bot Lab 24: 17 [syn sub Fr. squarrosa] 655 Index Hepat 5: [Java, Fiji, Japan] 609 Prod Flora Hepat Polynesiae: 141 [syn to Fr. squarrosa] 928 Stephani Icon Hepat Inedit [Galeiloba] C-192, microfiche 52 fig 3597 ["Java, leg Seubert (Herb Jack)"] 712 Hepaticology in India II: 170 [syn sub Fr. ericoides] 1660 Natural Environmental Sci Res 14: 13 [Japan, syn sub Fr ericoides] (Reinw., Blume & Nees) Nees 1845 (Nees) Mont. 1839 Page 450 vethii Sande Lac. 1884 Typus: Indonesia. Sumatra. "Barisan Bebirge bei Alahan Pandjang" 1877/79, Veth & van Hasselt sn in hb L (hb LUGD BAT 910.291-178) [lectotypus, vide HATT 1974 and HATT & ST 1985] in hb L [holotype fide Uribe-M. 2011] Basionym: Distribution: Taiwan, Philippines spec nov - syn nov Sande Lacoste in Veth 1884 807 Verdoorn Verdoorn Verdoorn Verdoorn 1930 1932 1932 1934 293 554 553 241 Herzog Hattori Hattori Hattori Hattori Hattori, Thaithong & Kitagawa Hattori & Thaithong 1949 1973 1974 1975 1976 1977 1978 18 542 545 547 80 480 70 Onraedt Hattori Grolle & Piippo Hattori Hattori & Streimann Tan & Engel Kuo & Chiang Piippo Yuzawa Yamada & Iwatsuki Uribe 1981 1982 1984 1985 1985 1986 1988 1990 2000 2006 2008 5794 495 143 73 504 4286 948 521 1659 1470 1513 Uribe-M. 2011 2767 = Frullania trichodes vethii Midden Sumatra, Reizen en Qnderzoekingen der Sumatra Expeditie, 4 (2): 44, pl 8 [= Hepat pl 2] 1-13 Ann Bryol Suppl 1: 123--124, fig 194-196 Nederl Kruidk Arch Jrg 1932 (2): 498-499 [Borneo] Bull Jard Bot Buitenzorg, ser 3, 12 (1): 61-62 [Sumatra] Ann Bryol Suppl 4: 209-210, fig 8-10 [B.N. Borneo: Mt Kinabalu, Sarawak] Trans Brit Bryol Soc 1 (3): 317 [Borneo, Sarawak] J Hatt Bot Lab 37: 151 J Hatt Bot Lab 38: 185-188, fig 86 [lectotypification] J Hatt Bot Lab 39: 311-312 J Hatt Bot Lab 40: 503, fig 200(v-y) J Hatt Bot Lab 43: 444 [note to fig 216 bei VERDOORN] [Thailand] Bull Natn Sci Mus Tokyo, ser B (Bot), 4 (2): 69 [Burma; vide Svihla 1955 as Fr. picta] J Hatt Bot Lab 50: 203 [Sri Lanka] J Hatt Bot Lab 51: 230-231 Acta Bot Fennica 125: 25 [Cat. West Melanesia] Bull Natn Sci Mus Tokyo, ser B (Bot), 11 (1): 16 [Fiji] J Hatt Bot Lab 59: 119, footnote [lectotypification ?] J. Hatt. Bot. Lab.. No. 60: 305 [Philippines] Taiwania 33: 14 [Index of Taiwan] J Hatt Bot Lab 68: 48 [annotated catalogue of China] Natural Environmental Sci Res 13: 21 [Japan, syn sub Fr trichodes] J Hatt Bot Lab 99: 33 [Catalog of Japan] Caldasia 30(1): 91, [type was not available] [Monography of subgenus Meteoriopsis] Caldasia 33(1): 74 [type study, syn sub Fr trichodes] Mitt. 1862 fo. acutiloba S.Hatt. 1976 Typus: Malaysia/Sabah. "N Borneo. Tawau Res., Lahad Datu" 18.9.1960, W. Meijer B-10522 or B10552 in hb L L0060949 [holotypus] in hb NICH [isotypus] Basionym: Distribution: Borneo fo nov - syn nov Hattori 1976 Hattori 1982 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania vethii 80 J Hatt Bot Lab 40: 503-504, fig 189 495 J Hatt Bot Lab 51: 230 2112 Type sepecimens Hatt Bot Lab (NICH) 38: Holotype. Borneo, Lahad Datu. 18 Sept. 1960. W. Meijer B10522 (NICH 432041). Sande Lac. 1884 Frullania trichodes Mitt. 1862 Page 451 vieillardi Gottsche ex Verd. 1930 [pro syn] Basionym: Distribution: Verdoorn Bonner Miller, Whittier & Whittier = Frullania intermedia villosa 1930 1965 1983 293 655 609 Ann Bryol Suppl 1: 161 Index Hepat 5: 464 [nom herb] Prod Flora Hepat Polynesiae: 141 (Reinw., Blume & Nees) Nees 1845 Steph. 1911 Typus: Ecuador: "in andibus" 1896, de Mathan sn in hb G(20627) [holotype] (hb LEVIER 4246) Basionym: Distribution: Ecuador, Bolivia spec nov Stephani Stephani Herzog Bonner Stephani 1911 1916 1942 1965 1985 - syn nov Uribe-M Uribe 2004 2008 = Frullania apollinarii Spec Hepat 4: 608 Biblioth Bot 87: 248 [Bolivia] Beih Bot Centralbl 61 B: 577 [Ecuador] Index Hepat 5: 464 Stephani Icon Hepat Inedit [Meteriopsis] E-48, microfiche 56 fig 3841 ["Ecuador, Levier 4246 ex parte"] 1174 Caldasia 26: 89-90, fig 1a-g) [sny nov] 1513 Caldasia 30(1): 91, [belongs to subgen Thyopsiella] [Monography of subgenus Meteoriopsis] Steph. 1911 Frullania lobatohastata viminicola 686 222 874 655 928 Steph. 1911 Spruce 1884 Typus: Ecuador:Andes Quitensis, Pichincha, 3000 m Jameson sn in hb G [isotypus] Basionym: Distribution: Ecuador, Bolivia spec nov Spruce Stephani Stephani Bonner Stephani 1884 1910 1916 1965 1985 243 686 222 655 928 - syn nov Yuzawa Gradstein, Meneses & Arbe 1991 2003 525 975 = Frullania ecklonii Trans Proc Bot Soc Edinburgh 15: 12-13 & 21-22 (No 9) Spec Hepat 4: 330 Biblioth Bot 87: 240 [Bolivia] Index Hepat 5: 464 Stephani Icon Hepat Inedit [Chonanthelia] A- 53, microfiche 47 fig 3146 from isotype; "Pichincha 3000 m ad arbores ramis legit Jameson ex herb Spruce"] J Hatt Bot Lab 70: 259 [syn sub Fr. ecklonii] J Hatt Bot Lab 91: 53 [Bolivia; as syn of Fr. ecklonii] (Spreng.) Gottsche, Lindenb. & Nees 1845 viminicola var. bicolor Spruce 1884 Typus: Peru: Mt Campana, 1200 m sd, Spruce Basionym: Distribution: Peru var nov Spruce Bonner = Frullania ecklonii 1884 1965 243 655 Trans Proc Bot Soc Edinburgh 15: 21-22 Index Hepat 5: 464 (Spreng.) Gottsche, Lindenb. & Nees 1845 Page 452 virginica [auct. non Lehm.] Russell 1858 Basionym: Distribution: Russell 1858 = Frullania eboracensis virginica 680 Proc Essex Inst 2: 134 [Fayal, Azores] Lehm. 1844 [auct. non Gottsche] A.Evans 1897 Basionym: Distribution: Evans Schuster = Frullania inflata 1897 1992 237 603 Trans Connet Acad Arts Sci 10 (1): 17-18 Hepat N America 5: 132 & 147 Gottsche 1845 [non Herzog 1921] viridis [auct. non Colenso 1888, nec Horikawa 1930] Horikawa 1934 {Taiwan specimen: YH 9043} Basionym: Distribution: - syn nov Horikawa Lin & Wang-Yang Hattori Hattori & Lin Kuo & Chiang = Frullania monocera viridis 1934 1976 1980 1985 1988 34 33 489 505 948 J Sci Hiroshima Univ, ser B, div 2, 2: 240 [Bonin Isl & Formosa] Taiwania 21 (2): 169 [Index Formosa] J Hatt Bot Lab 47: 104 [syn sub Fr. monocera] J Hatt Bot Lab 59: 139 Taiwania 33: 14 [Index of Taiwan] (Hook. f. & Taylor) Gottsche, Lindenb. & Nees 1845 Colenso 1888 [non Horikawa 1930, nec Horikawa 1934] Typus: New Zealand, County of Waipawa, Dannevirke 1887, W. Colenso in hb G Basionym: Distribution: spec nov - syn nov Colenso Stephani Bonner Hamlin = Frullania squarrosula 1888 1892 1965 1972 635 210 655 659 Trans Proc New Zealand Inst 20: 252 J Linn Soc Bot 29: 274 [syn sub Fr. squarrosula] Index Hepat 5: 465 Rec Dominion Mus 7(19): 263 [New Zealand] (Hook.f. et Taylor) Gottsche, Lindenb. et Nees 1845 Page 453 viridis Horikawa 1930 [hom illeg] [non Colenso 1888, nec Horikawa 1934] {Bonin} Typus: Japan. Bonin, Isl. Chichijima; on old trunk of tree fern Cyathea boninsimensis -.2.1927, T Shibata in hb Y. Horikawa 259 in hb HIRO [holotypus] Basionym: Distribution: Taiwan, Bonin Isl spec nov - nom nov Horikawa Horikawa Hattori & Noguchi Miller Miller Iverson Bonner Miller Inoue & Iwatsuki Inoue in Inoue & Iwatsuki Lai & Wang-Yang Hattori 1930 1934 1960 1960 1960 1963 1965 1968 1969 1970 1976 1978 Hattori Piippo Yuzawa NN 1980 1990 2001 2003 Yamada & Iwatsuki 2006 = Frullania bonincola 947 34 965 920 920 Sci Rep Tohoku Imp Univ, ser 4, Biol, 5 (4): 646-648, fig 13(1-16) J Sci Hiroshima Univ, ser B, div 2, 2: 240, fig 45 [Bonin Isl, & Taiwan] Index spec Typ herb Japon, Bryophyta: 18 Bryologist 63: 121 [Micronesia] Bryologist 63: 121 [Micronesia] : 124 655 Index Hepat 5: 465 672 Micronesia 4 (1): 74 [key to Guam & Northern Micronesia] 61 Bull Natn Sci Mus Tokyo 12 (2): 305 62 Bull Natn Sci Mus Tokyo 13 (3): 478 33 Taiwania 21 (2): 168 [Index to Taiwan] 482 J Hatt Bot Lab 44: 551-553, fig 218 [nom nov = Fr. bonincola /Bonin Is; specimen from Taiwan = Fr. monocera] 489 J Hatt Bot Lab 47: 104 . & monocera /Formosa] 521 J Hatt Bot Lab 68: 48 [annotated catalogue of China] 1660 Natural Environmental Sci Res 14: 7 [Japan, basionyn to Fr bonincola] 1397 Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] S.Hatt. 1978 viridis [auct. non Colenso 1888, nec Horikawa 1930] Horikawa 1934 {Taiwan specimen: YH 10316} Basionym: Distribution: - syn nov Horikawa Lin & Wang-Yang Hattori Hattori & Lin Piippo 1934 1976 1980 1985 1990 34 33 489 505 521 J Sci Hiroshima Univ, ser B, div 2, 2: 240 [Bonin Isl & Formosa] Taiwania 21 (2): 169 [Index Formosa] J Hatt Bot Lab 47: 104 [Taiwan specimens syn sub Fr. monocera] J Hatt Bot Lab 59: 139 [syn sub Fr. monocera var. depauperata] J Hatt Bot Lab 68: 139-149, annotation nr 142 about status of Formosa specimens] = Frullania monocera var. depauperata S.Hatt. 1984 Page 454 virillana Steph. 1911 Typus: Costa Rica. Rio Virilla sd, Tonduz in hb G (hb LEVIER 6207) Basionym: Distribution: Costa Rica, Venezuela spec nov - syn nov - syn nov Stephani Bonner Stotler Stotler Lisboa Yano Stephani 1911 1965 1969 1970 1976 1984 1985 Gradstein & Costa Yano 2003 2008 = Frullania montagnei vueltensis 686 655 4761 194 644 501 928 Spec Hepat 4: 528 Index Hepat 5: 465 Nova Hedwigia 18: 452-453 [syn sub Fr. montagnei] The genus Frullania, separata: 452-453 [syn sub Fr. montagnei] Acta Amazonica 6 (2): 175 [Amazonas] J Hatt Bot Lab 56: 507 [checklist Brazil] Stephani Icon Hepat Inedit [Thyopsiella] F-128, microfiche 58 fig 4055 ["Costarica, rio Virilla, Tonduz, Levier 6207"] 984 Mem New York Bot Gard 87: 89 [Brazil] [syn sub Fr montagnei] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr montagnei; Brazil; catalog] Gottsche 1845 Herzog ex Bonner 1965 [nom herb] {Costa Rica} Typus: Costa Rica: Prov. San Jose, Cerro de las Vueltas, 2700-3000 m 29.12.1925 - 1.1.1926, P.C. Standley 43851 in hb G, in hb US (2x) Basionym: Distribution: Costa Rica Bonner NN 1965 1996 655 632 Index Hepat 5: 465 gopher://nmnhgoph.si.edu:70/77/.index/nonflwr [2.4.1996] keyword=Frullania, printed 5.1.1998, # 34-35 of 35 [in hb US] = vuquangensis Pocs ex Ninh 2003 [nom nud] Typus: Vietnam. Basionym: Distribution: Vietnam - syn nov Ninh Pócs = Frullania hamatiloba 2003 2019 1363 ASEAN Biodiversty 3 (1-2): 30 [new to Vietnam] 5243 pers. communication [= Fr hamatiloba, never effectively and validly published] Steph. 1910 Page 455 wallichiana Mitt. 1861 Typus: Sikkim: Himalaya 4000-7000 ft J. D. Hooker sn in hb NY [lectotypus] Basionym: Distribution: China, Taiwan, Himalayas, India (North: Eastern Himalayas: Megalaya, West Bengal (Darjeeling), Assam; South: Madras; Nilgiri Mts), Nepal, Sikkim, Burma, North Vietnam, Indonesia (Sumatra), Philippines, New Guinea, Australia spec nov - syn nov Mitten Sande Lacoste in Veth 1861 1884 574 807 Schiffner Stephani Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Verdoorn Chopra 1898 1910 1929 1930 1931 1932 1932 1934 1938 831 686 245 293 230 553 554 241 1648 Chopra Demaret & Vanden Berghen Svihla Parihar Arnell Bonner Hattori in Hara Grolle Pocs, Tixier & Jovet-Ast Hattori Hattori Hattori Hattori Hattori Hattori in Ohashi Hattori & Thaithong Hattori & Thaithong Hattori & Thaithong Hattori & Thaithong Hattori & Thaithong Hattori Onraedt Hattori Grolle & Piippo Hattori & Lin Stephani 1943 1948 1956 1962 1963 1965 1966 1966 1967 1972 1974 1974 1975 1975 1975 1978 1978 1978 1978 1978 1979 1981 1982 1984 1985 1985 205 59 380 1584 822 655 841 843 1710 538 544 546 65 547 187 128 70 127 481 70 129 5794 495 143 505 928 Kuo & Chiang Piippo Parihar, Lal & Katiyar 1988 1990 1994 948 521 2196 Nath & Asthana Singh 1998 2002 708 4886 Srivastava & Alam Singh, Kumar & Nath 2002 2008 1251 1281 Hardianty 2018 5036 = Frullania obscura J Linn Soc Bot 5: 118-119 [Nepal, Sikkim, Khasi, Madras] Midden Sumatra, Reizen en Onderzoekingen der Sumatra Expeditie, 4 (2): 44 [Sumatra] Conspect Hepat Archip Indici, Batavia: 348 Spec Hepat 4: 348 Ann Bryol 2: 120-121 [Java, Sumatra] Ann Bryol Suppl 1: 70-72, fig. 133-136 Ann Bryol 4: 123 [Hepat select crit, ser 1 no 35; Java] Bull Jard Bot Buitenzorg, ser 3, 12 (1): 57-58 [Sumatra, Java] Nederl Kruidk Arch Jrg 1932 (2): 492 [Java, Bali] Ann Bryol Suppl 4: 204 [Sumatra] Proceed Indian Acad Sci, sect B, 8: 437 [Sikkim Himalayas and Bengal] J Indian Bot Soc 22: 249 [India] Bull Jard Bot Etat Bruxelles 19 (1): 77 [syn nov sub Fr. arecae] Bryologist 59: 277-278 [syn sub Fr. arecae] Univ Allahabad studies, Bot sect 1-2: 19 [India: EH WH SI] Hepat S Africa: 136 [syn sub Fr. arecae] Index Hepat 5: 466 Flora of Eastern Himalaya: 528 Khumbu Himal 4: 266 [Nepal] [syn sub Fr. arecae] Bot Kozlem 54 (1): 30 [North Vietnam] J Hatt Bot Lab 36: 134-135 J Hatt Bot Lab 38: 177-178 [Luzon] J Hatt Bot Lab 38: 273-274 [New Guinea] Bull Natn Sci Mus Tokyo, ser B (Bot), 1 (3): 120 J Hatt Bot Lab 39: 312 Univ Mus Univ Tokyo Bull 8: 236 [Nepal] J Jap Bot 53 (6): 177 [Laos] Bull Natn Sci Mus Tokyo, ser B (Bot), 4 (2): J Jap Bot 53 (5): 132 J Hatt Bot Lab 44: 193 [Assam, Darjeeling] Bull Natn Sci Mus Tokyo, ser B (Bot), 4 (2): 69 [Darjeeling, Burma] J Jap Bot 54 (1): 25-26 [Sri Lanka] J Hatt Bot Lab 50: 203 [Sri Lanka] J Hatt Bot Lab 51: 262 Acta Bot Fennica 125: 25 [Cat. West Melanesia] J Hatt Bot Lab 59: 151 [sub Fr. arecae] Stephani Icon Hepat Inedit [Chonanthelia] A- 93, microfiche 47 fig 3201-3202 ["Sikkim Himalaya (leg Bretandeau 1894"] Taiwania 33: 14 [Index of Taiwan] J Hatt Bot Lab 68: 48 & 139, annotation nr 141 about synonymy Hepat Anth India, annotated checklist: 22 [Eastern Himalaya, Western Himalaya, Nepal; South India] J Hatt Bot Lab 85: 64-66, fig 1(1-13) [South India; key to all spec.] Studies on Liverworts (Bryophyta) of Khasi and Jaintia Hills: Meghalaya. 244-246, pl 48(1-23), map 11 - Unpublished thesis. J Bombay Nat Hist Soc 99(2): 235 [Nilgiri, key to South Indian species] Taiwania 53 (1): 54-56, fig 1[map], 2(1-22) [India, Meghalaya, Eastern Himalayas] Frullaniaceae di Taman Wisata Alam Sicike-Cike: 37, photos [Sumatra, with key] (Sw.) Mont. 1840 Page 456 wallichiana var. hosseana (Steph.) S.Hatt. 1972 Basionym: Frullania hosseana Distribution: Thailand comb nov - syn nov Steph. 1910 Hattori Yamada, Yuzawa, Ogasawara & Lai Hattori = Frullania wallichiana 538 949 J Hatt Bot Lab 36: 134-135, fig. 19 (i-q) Quart J Taiwan Mus 34 (1,2): 114 [new to Taiwan] 1982 495 J Hatt Bot Lab 51: 262 [syn sub Fr. wallichiana] Mitt. 1861 Frullania obscura warmingiana 1972 1981 (Sw.) Mont. 1840 Steph. 1910 Typus: Brazil: Rio Janeiro, Fraitas 29.5.1863, C. Warming sn in hb G (ex hb GOTTSCHE) [holotypus] Basionym: Distribution: Venezuela, Brazil spec nov - syn nov Stephani Schiffner & Arnell 1910 1964 Bonner Yano Stephani 1965 1984 1985 Yuzawa Gradstein & Costa Yano 1991 2003 2008 = Frullania rio-janeirensis wattsiana 686 198 Spec Hepat 4: 330-331 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 85 [? syn to Fr. riojaneirensis] [Brazil] 655 Index Hepat 5: 466 501 J Hatt Bot Lab 56: 507 [checklist Brazil] 928 Stephani Icon Hepat Inedit [Chonanthelia] A- 54, microfiche 47 fig 3147 [from type; "Fraitas Rio Janeiro Brsilia lg E Warming 1863 -Herb Gottsche-"] 525 J Hatt Bot Lab 70: 254 [syn sub Fr. riojaneirensis] 984 Mem New York Bot Gard 87: 89 [Brazil] [syn sub Fr riojaneirensis] 1684 Boletim Instituto Botanica 19: 90 [syn sub Fr riojaneirensis; Brazil; catalog] (Raddi) Ångstr. 1876 Steph. 1911 Typus: Australia. New South Wales. Blackheath, Head of Neate´s Glen 7.4.1903, W.W. Watts 688 hb LEVIER 4429, in hb G Basionym: Distribution: Australia (NSW) spec nov - syn nov Stephani Stephani & Watts Bonner Hattori Stephani 1911 1914 1965 1979 1985 Scott & Bradshaw Carthy McCarthy 1986 2003 2006 = Frullania falciloba 686 220 655 485 928 Spec Hepat 4: 559 J Proc Roy Soc New S Wales 48: 109 [Australia] Index Hepat 5: 466 J Hatt Bot Lab 45: 361-361, fig 27(a-h) from type [syn sub Fr. falciloba] Stephani Icon Hepat Inedit [Thyopsiella] F-184, microfiche 60 fig 4135 ["N S Wales, Watts 688, Levier 4429"] 56 Brunnonia 8 (1): 37 [Australia] 1125 Flora Australia Suppl Ser 21: 43 [sub Fr falciloba] 1385 Checklist Australian Liverworts [Australia, syn sub Fr falciloba] Lehm. 1844 Page 457 welwitschii Steph. 1901 Typus: Angola: Golungo Alto, mountains of Altra Queta -.10.1854, Welwitsch 299 ex hb BM, in hb G 5197 Basionym: Distribution: spec nov - syn nov Stephani Stephani Arnell Arnell Bonner Vanden Berghen Stephani 1901 1910 1962 1963 1965 1976 1985 Wigginton & Grolle Wigginton 1996 2002 Wigginton Wigginton 2004 2004 = Frullania squarrosa Cat Afr Plants Welwitsch 2 (2): 316-317 [Angola] Spec Hepat 4: 380 Svensk Bot Tidskr 56 (1): <55-60> [syn sub Fr. squarrosa] 822 Hepat S Africa: 150 [syn sub Fr. squarrosa] 655 Index Hepat 5: 466-467 185 Bull Jard Bot Nat Belg 46 (1/2): 163 928 Stephani Icon Hepat Inedit [Galeiloba] C- 47, microfiche 50 fig 3402 [from type; "Golungo alto - Angola, Welwitsch 299, ... Oct 1854"] 696 Bryophyt Bibl 50: 94 [= syn sub Fr. squarrosa = syn sub Fr. ericoides] 1012 Trop Bryol Res Rep 2: 30 [http://www.oshea.demon.co.uk/tbr/afrheps.pdf] 1312 Trop Bryol Res Rep 5: 32 [http://www.oshea.demon.co.uk/tbr/tbrr5.pdf] 1172 Sript Bot Belg 30: 231 [syn sub Fr ericoides, West Africa] (Reinw., Blume & Nees) Nees 1845 Frullania ericoides welwitschii 883 686 (Nees) Mont. 1839 Steph. 1910 [nom superfl] Basionym: Distribution: Stephani 1910 = Frullania welwitschii Spec Hepat 4: 380 [spec nov = nom superfl] Steph. 1901 Frullania squarrosa Frullania ericoides weymouthiana 686 (Reinw., Blume & Nees) Nees 1845 (Nees) Mont. 1839 Steph. 1911 Typus: Tasmania. Ida Bay. "on Sassafras bark" -.4.1811, Weymouth 1139 in hb G 18490 [holotypus] in hb MEL 61913 [isotypus] Basionym: Distribution: Tasmania spec nov - syn nov Stephani Rodway 1911 1916 Bonner Hattori Hattori Hattori Stephani 1965 1979 1983 1983 1985 Scott & Bradshaw Carthy McCarthy 1986 2003 2006 = Frullania aterrima 686 914 Spec Hepat 4: 660-661 Papers Proceed Roy Soc Tasmania for 1916: 137 [syn sub Fr. congesta] 655 Index Hepat 5: 486 J Hatt Bot Lab 46: 149-150, fig 47(a-j) from type [= Fr. rostrata ?] 499 J Hatt Bot Lab 54: 164 [= rostrata] 499 J Hatt Bot Lab 54: 133-135 [syn sub Fr. aterrima] 928 Stephani Icon Hepat Inedit [Diastaloba] B-98, microfiche 49 fig 3346 [from type; "Tasmania, leg Weymouth 1139"] 56 Brunnonia 8 (1): 37 [Australia] 1125 Flora Australia Suppl Ser 21: 42 [sub Fr aterrima] 1385 Checklist Australian Liverworts [Australia, syn sub Fr aterrima] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Page 458 weymouthiana [auct. non Steph. 1911] S.Hatt. 1979 Basionym: Distribution: auct - syn nov Hattori Hattori = Frullania rostrata wichurae 1979 1983 486 499 J Hatt Bot Lab 46: 149-150, fig 47(a-j) from type J Hatt Bot Lab 54: 164 [syn sub Fr. rostrata] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 Schiffn. ex Verd. 1930 [pro syn] Basionym: Distribution: Verdoorn 1930 = Frullania junghuhniana wightii 293 Ann Bryol Suppl 1: 134 [syn sub Fr Junghuhniana] Gottsche 1845 Gottsche ex Verd. 1930 [pro syn] Basionym: Distribution: Verdoorn Miller, Whittier & Whittier = Frullania nodulosa wildii 1930 1983 293 609 Ann Bryol Suppl 1: 180 Prod Flora Hepat Polynesiae: 141 (Reinw., Blume & Nees) Nees 1845 Steph. 1894 Typus: Australia: Queensland, Toowoomba, on trees sd, C. Wild 453 hb BROTHERUS No 22 in hb G 18234 Basionym: Distribution: Australia (NSW, QLD) spec nov -syn nov Stephani Stephani Watts Bailey Bonner Hattori 1894 1910 1912 1913 1965 1979 Miller, Whittier & Whittier Stephani 1983 1985 Scott & Bradshaw Hattori Windolf Konrat & Braggins Downing, Oldfield & FairbairnWilson Carthy Glenny, Meagher & Brown 1986 1987 1987 2001 2002 Glenny, Meagher & Brown 2004 McCarthy 2006 = Frullania pentapleura 2003 2004 228 686 1940 3470 655 486 Hedwigia 33: 169 [Diastaloba] Spec Hepat 4: 414-415 unpublished Msc Compreh catalogue Queensland plants: 675 [Queensland] Index Hepat 5: 467 J Hatt Bot Lab 46: 150-151, fig. 48(a-c) from type [close to Fr. pentapleura] 609 Prod Flora Hepat Polynesiae: 141 [also at Lord Howe Isl] 928 Stephani Icon Hepat Inedit [Galeiloba] C-226, microfiche 53 fig 3644 ["Queensland, Toowooomba, C Wild 453, Herb Brotherus 22"] 56 Brunnonia 8 (1): 37 & 144 [Australia: NSW, QLD] 400 Bryologist 90 (4): 370 [not Lord Howe Isl.] 1025 Austrobaileya 2(4): 391 [checklist Queensland] 838 J Hatt Bot Lab 91: 238-245 [syn nov sub Fr. pentapleura] 1145 Cunninghamia 7(3): 533 [Australia, NSW, Mt Canobolas] 1125 Flora Australia Suppl Ser 21: 45 [sub Fr pentapleura] 1126 NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1127 Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html 1385 Checklist Australian Liverworts [Australia, syn sub Fr pentapleura] Taylor 1846 Page 459 wildii [auct. non STEPH 1824] Steph. & Watts 1914 {Lord Howe Isl} Typus: Australia. [NSW] Lord Howe Isl WATTS 4 (det Steph) Basionym: Distribution: Lord Howe Isl Stephani & Watts Hattori = Frullania fulfordiae willkommii Typus: Spain 1914 1987 220 400 J Proc Roy Soc New S Wales 48: 109 [Lord Howe Isl] Bryologist 90 (4): 366 S.Hatt. 1987 Steph. 1911 sd, Willkomm sn in hb G G00066774 - n° SIB 132013/1 (ex hb JACK) [holotypus] > see fr2001.jpg in hb S (B24328) [isotypus] Basionym: Distribution: Spain spec nov - syn nov Stephani Müller in Rabenhorst Müller Frye & Clark Müller in Rabenhorst Bonner Stephani 1911 1915 1915 1947 1958 1965 1985 686 1064 0 204 1028 655 928 Internet - Swedisch Musum of Nat Hist, Dep Crypt Bot 2002 782 Spec Hepat 4: 570-571 Krypt Flora (2ed) 6, 2: 610 note in hb G: "ist form der F. tamarisci" > see fr2001.jpg Univ Wash Publ Biol 6 (5): 748 [syn sub Fr tamarisci] Krypt Flora (3ed) 6, 2: 1234 [syn sub Fr tamarisci var mediterranea] Index Hepat 5: 467 Stephani Icon Hepat Inedit [Thyopsiella] F-203, microfiche 60 fig 4168 ["Spanien, leg Willkomm, Herb Jack"] http://www.nrm.se/cgi-bon/Sfgate [type collection in hb S] = Frullania tamarisci var. mediterranea (De Not.) Nees 1845 wrightii Austin 1872 Typus: USA. New Mexico 1861, C. Wright in hb SULLIVANT Basionym: Distribution: spec nov - syn nov Austin Evans Stephani Frye & Clark Mamontov et al = Frullania riparia 1872 1897 1911 1947 2020 666 237 686 204 5641 Bull Torrey Bot Club 3 [No 3]: 15-16 Trans Connet Acad Arts Sci 10 (1): 34-35 Spec Hepat 4: 682 Univ Wash Publ Biol 6 (5): 774 [syn sub Fr riparia] Nova Hedwigia, Beiheft 150, p. 201-242 [belong to subg. Frullaniopsis subgen. nov.] Hampe 1838 Page 460 wullschlaegeli Steph. 1910 Typus: Suriname "Dutch Guiana". Paramaribo sd, Wullschlaegel 706 in hb G G00067426 - n° SIB 131989/1 - 20980 (ex hb M) [holotypus] Basionym: Distribution: Surinam spec nov - syn nov Stephani Schiffner & Arnell Bonner Yuzawa Yano Stephani 1910 1964 1965 1983 1984 1985 686 198 655 498 501 928 Gradstein & Hekking 1989 517 = Frullania gibbosa yakushimensis Spec Hepat 4: 338 Österr Akad Wiss, Math Nat Kl, Denkschr 111: 85, pl 13 (145) [Brazil] Index Hepat 5: 468 J Hatt Bot Lab 54: 107-109, fig. 1(a-n) [from type] [syn sub Fr. gibbosa] J Hatt Bot Lab 56: 507 [checklist Brazil] Stephani Icon Hepat Inedit [Chonanthelia] A- 56, microfiche 47 fig 3150 [from type; "Paramaribo Wullschlägel No 706"] J Hatt Bot Lab 66: 208 & 221 [syn nov sub Fr. gibbosa] Nees 1840 Horikawa 1934 Typus: Japan. Kyushu, Isl. Yaku, between Ishizuka and Hanano-ego 11.7.1933, Y. Horikawa 11912 in hb HIRO [holotypus] Basionym: Distribution: Japan spec nov comb nov - syn nov Horikawa Hattori Hattori Hattori & Noguchi Kamimura Bonner Choe Yuzawa NN 1934 1951 1952 1960 1961 1965 1980 2001 2003 Yamada & Iwatsuki 2006 = Frullania usamiensis 34 528 529 965 199 655 1171 1660 1397 J Sci Hiroshima Univ, ser B, div 2, 2: 237-238, fig 44 J Hatt Bot Lab 5: 67 [as usamiensis fo. yakushimensis] J Hatt Bot Lab 8: 31 [syn nov sub Fr. usamiensis] Index spec Typ herb Japon, Bryophyta: 19 J Hatt Bot Lab 24: 43 [syn sub Fr usamiensis] Index Hepat 5: 468 Illust Fl & Fauna Korea 24: 736 [sub Fr usamiensis] Natural Environmental Sci Res 14: 38 [Japan, syn sub Fr usamiensis] Types of Nat Hist mus Hiroshima [http://www.digitalmuseum.hiroshimau.ac.jp/~museum/types/pages/hepa/f/hirotypeHepaF.html ] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] Steph. 1897 Page 461 yorkiana Steph. 1911 Typus: Australia: Queensland, Cape York -.8.1897, Micholitz sn in hb G G00066920 - n° SIB 146456/1 - 18235 [holotypus] Basionym: Distribution: Australia (QLD) spec nov - syn nov Stephani Bonner Hattori Hattori 1911 1965 1976 1979 686 655 0 486 Hattori Stephani 1984 1985 682 928 Scott & Bradshaw Windolf Scott Hattori Carthy Glenny, Meagher & Brown 1986 1987 1987 1988 2003 2004 56 1025 1700 429 1125 1126 Glenny, Meagher & Brown 2004 1127 McCarthy Hentschel et alii 2006 2015 1385 4097 = Frullania fugax yuennanensis Typus: Spec Hepat 4: 558-559 Index Hepat 5: 468 note in hb G: "probably conspecific with Fr. fugax" Nov 2, 1976 J Hatt Bot Lab 46: 151-152, fig. 49(A-e) from type [noted the similarity with F. fugax and stated that it possibly is a synonym] Crypt Bryol Lichen 5(1-2): 189 [Australia, QLD] Stephani Icon Hepat Inedit [Thyopsiella] F-185, microfiche 60 fig 4136 [Cape York, Queensland, legit Micholitz"] Brunnonia 8 (1): 37 & 145 [Australia: QLD] Austrobaileya 2(4): 391 [checklist Queensland] Queensland Naturalist 28(1-4): 64 [from Cape York peninsula] Nova Hedwigia Beih 90: 157 [Australia: QLD] Flora Australia Suppl Ser 21: 48 [Australia (QLD)] NSW Liverworts [Australia, NSW] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Austalian Liverworts [Australia] http://plantnet.rbgsyd.nsw.gov.au/PlantNet/Liverworts/index.html Checklist Australian Liverworts [Australia QLD] Phytotaxa 220 (2): 135 [sub Fr fugax] (Hook.f. & Taylor) Gottsche, Lindenb. & Nees 1845 fo. refleximarginata Verd. 1930 ?, Gebauer 648, 595, 6985 ? Basionym: Distribution: China fo nov Verdoorn in Handel-Mazzetti Hattori & Lin Piippo = Frullania yuennanensis yulensis 1930 1985 1990 550 505 521 Symb Sinicae 5: 42 [China] J Hatt Bot Lab 59: 148 J Hatt Bot Lab 68: 49 [annotated catalogue of China] Steph. 1894 Steph. 1911 Typus: Papua New Guinea. Cental, Mt. Yule -.12.1890, McGregor sn in hb G G00066917 - n° SIB 124204/1 - 14243 (hb GEHEEB (1897) ex hb MEL (16.3.1891) [holotypus] Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Verdoorn Hattori Grolle & Piippo Stephani 1911 1930 1930 1973 1984 1985 Hattori & Piippo Uribe 1986 2008 Uribe-M. 2011 = Frullania seriatifolia 686 293 549 540 143 928 Spec Hepat 4: 536-537 Ann Bryol Suppl 1: 146, fig 261-262, 291 Nova Guinea 14 (4): 545 J Hatt Bot Lab 37: 79-80, fig 39 [syn sub Fr. seriatifolia] Acta Bot Fennica 125: 25 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Thyopsiella] F-158, microfiche 59 fig 4098 ["N Guinea, Mt Yule, Herb Melbourne 1890"] 557 Acta Bot Fennica 133: 33, fig 2 1513 Caldasia 30(1): 90, [belongs to subgen Diastaloba] [Monography of subgenus Meteoriopsis] 2767 Caldasia 33(1): 72 [type study, syn sub Fr seriatifolia] Steph. 1894 Page 462 yunnanensis Typus: Taiwan [auct. non Steph.] S.Hatt. 1980 {from Taiwan, Inoue nr 14542} Inoue nr 14542 Basionym: Distribution: - syn nov Hattori 1980 958 Hattori & Lin 1985 505 = Frullania nepalensis yunnanensis Bull Natn Sci Mus Tokyo, ser B (Bot), 6 (1): 40 [specimens from Taiwan] J Hatt Bot Lab 59: 148 (Spreng.) Lehm. & Lindenb. 1845 Steph. by S. Hatt. 1966 [orth var] Basionym: Distribution: Hattori in Hara Hattori in Ohashi Hattori Bapna & Kachroo Bai in Gao & Cao Kattel Kattel 1966 1975 1978 2000 2000 2002 2002 = Frullania yuennanensis yusawae 841 187 482 712 1019 1032 1032 Flora of Eastern Himalaya: 528-529 Univ Mus Univ Tokyo Bull 8: 236 [Nepal] J Hatt Bot Lab 44: 548-550, fig. 216 Hepaticology in India II: 184 Fl Yunnanica 17: 417, fig 209(1-9) [China, Yunnan] Liverworts of Nepal, Katmandu: 20 [Nepal, 1150-4000 m] Liverworts of Nepal, Katmandu: 20 [Nepal] Steph. 1894 Kamim. by Furuki & Mizutani 1994 [orth var] Basionym: Distribution: Japan Yuzawa & Hattori Inoue N.N. [National Science Museum Tokyo] Yamada & Iwatsuki = Frullania yusawai 494 J Hatt Bot Lab 50: 257 [Japan] 1155 Index type specimens in Nat Sci Mus, Tokyo: 15 995 http://research.kahaku.go.jp/botany/koke/indexf.htm [in hb TNS] 2006 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] Kamim. 1970 Frullania hamatiloba yusawai 1981 1987 2000 Steph. 1910 Kamim. 1970 Typus: Japan. Fukushima Pref., Kawamae-mura, Setogaro, ca 150 m (Yuzawa 901) & Tono-machi, Gosaisho-toge (Yuzawa 1027, 1030, 843) 15.12.1961 (Yusawa 901), 25.3.1962 (Yuzawa 1027, 1030), 22.10.1961 (Yuzawa 843) in hb NICH (Y. Yusawa 1027, 1030 [typus] in hb TNS (Y. Yusawa 901) [isotypus] Basionym: Distribution: Japan spec nov Kamimura 1970 - syn nov Furuki & Mizutani Yuzawa 1994 2001 Yamada & Iwatsuki 2006 = Frullania hamatiloba 1552 Bulletin of Kochi Gakuen Junior College 2: 21, fig 2(a-l) [Japan, as Fr. yusawai, close to Fr pedicellata] 1331 Proc Bryol Soc Japan 6 (5): 78 [Japan; checklist] 1660 Natural Environmental Sci Res 14: 16, fig 14(a-o) [Japan, syn sub Fr hamatiloba] 1470 J Hatt Bot Lab 99: 33 [Catalog of Japan] as "yusawae" Steph. 1910 Page 463 zahnii Steph. 1911 Typus: Neu Guinea. PNG [German New Guinea], Morobe. "An Bäumen auf dem Sattelberg" 53 ??] -.7.1903, H. Zahn 53 [ex hb KAULFUSS no in hb G 17502 - G00060707 - n° SIB 124218/1 - (-/7/1903, Zahn 53, ex hb J. KAULFUSS) [holotypus] > see jpg in hb PH 3115 (-/9/1905, Zahn sn, Nürnberger Botanischer Tauschverein, J. Kaulfuss) > see png in hb NICH, hb L, hb FH, hb HBG, hb JE Basionym: Distribution: spec nov - syn nov Stephani Verdoorn Bonner Grolle & Piippo Stephani 1911 1930 1965 1984 1985 Feuerer 2008 Mizutani, Hasegawa & Iwatsuki 2009 = Frullania regularis zhenjingensis 686 549 655 143 928 Spec Hepat 4: 578-579 Nova Guinea 14 (4): 541 [syn sub Fr. regularis] Index Hepat 5: Acta Bot Fennica 125: 25 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Homotropantha] D-14, microfiche 54 fig 3723 ["N Guinea, Sattelberg, leg Miss Zahn"] 2517 in hb HBG: http://www.biologie.unihamburg.de/bzf/herbar/hbg_bryophytes.htm 2112 Type sepecimens Hatt Bot Lab (NICH) 39: Isotype. New Guinea (Nova Guinea Germanica), Sattelberg. An Baumen. July 1903. H. Zahn (NICH 354857). Schiffn. 1890 (C. Gao & G.C. Zhang) Y.Huan Wu 2010 Typus: China. "Zhejiang: Mt. Nanyandangshan . . . 250 m . . . ." Jul 28, 1960 Institution(s): Holotype: ISPH [=IFP] Basionym: Neohattoria zhenjingensis Distribution: comb nov zippelii C. Gao & G.C. Zhang Wu 2010 = Frullania zhenjingensis Gao Chien 835 2349 Genera Hepaticopsida et Anthocerotopsida Sinicorum 380. 2010 [*invalid, basionym citation incomplete] (C. Gao & G.C. Zhang) Y. Jia & S. He 2013 Sande Lac. 1864 Typus: New Guinea. sd, Zippelius hb LUGD BATAV Basionym: Distribution: spec nov - syn nov Sande Lacoste 1864 842 Stephani Stephani & Watts Verdoorn Verdoorn Bonner Grolle & Schultze-Motel Miller, Whittier & Whittier Grolle & Piippo Stephani 1911 1914 1930 1930 1965 1972 1983 1984 1985 686 220 293 549 655 537 609 143 928 Hattori & Piippo 1986 557 = Frullania gaudichaudii Ann Mus Bot Lugduno-Batavi [ed Miquel] 1: 313, pl 8(1-6) [Nova Guinea] Spec Hepat 4: 557 J Proc Roy Soc New S Wales 48: 109 [Lord Howe Isl] Ann Bryol Suppl 1: 92 [syn sub Fr. gaudichaudi] Nova Guinea 14 (4): 544 Index Hepat 5: 469 J Hatt Bot Lab 36: 83 [Samoa] [sub Fr. gaudichaudii] Prod Flora Hepat Polynesiae: 141 Acta Bot Fennica 125: 25 [Cat. West Melanesia] Stephani Icon Hepat Inedit [Thyopsiella] F-176, microfiche 59 fig 4124 ["Nova Guinea"] Acta Bot Fennica 133: 41 (Nees & Mont.) Nees & Mont. 1845 Page 464 This is a working document in progress. The first number is the year of publication, the second the number in my database If something is missing, if there are mistakes or errors, or if you are interested in the literature on which this paper is based, please send me an email. © Gerhard Winter - email: gerhard.winter@senckenberg.de Page 465

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