The Dzhizak Province is a large administrative region of the Republic of Uzbekistan located in the central part of the country,
between two large rivers, Syrdarya and Zeravschan. There are represented all altitudinal zones and the majority of ecosystems
and vegetation types of Central Asia. The territory of the Dzhizak Province is divided into two physiographical parts. The
northern plain part includes the Kyzylkum Desert, the Aydar-Arnasay lake system, and the Hungry Steppe depression in
the middle reaches of the Syrdarya River. The southern mountainous part includes Turkestan Range and its western spurs,
Malguzar and Nuratau mountains. The highest and lowest points are 4029 m and 240 m above sea level, respectively. This
area is characterized by rich plant diversity with a large number of endemic, endangered species, and species of global
economic importance. The Flora of the Dzhizak Province, Uzbekistan has been compiled for the first time. It includes 1965
species of 613 genera and 105 families of vascular plants, among which 51 species are red-listed at the national level. The
book contains following information of species: life form, habitat, altitudinal zone, distribution within province, conservation
status, and economic use. Photographs of selected plant species and landscapes are provided.
P)数据
图书在版编目(CI
乌兹别克斯坦吉扎克省植物 = Flora of the
Dzhizak Province,Uzbekistam : 英文 / ( 乌兹 ) 托基比
夫·科米尔杰恩等著 . -- 北京 : 中国林业出版社 ,
2020.6
(“一带一路”沿线植物多样性保护研究)
ISBN 978-7-5219-0519-5
Ⅰ . ①乌… Ⅱ . ①托… Ⅲ . ①植物-生物多样性-生
物资源保护-乌兹别克-英文 Ⅳ . ① Q948.536.2
中国版本图书馆 CIP 数据核字 (2020) 第 089026 号
中 国 林 业出 版 社 · 自 然 保 护 分社(国家公园分社)
策划编辑:肖静
责任编辑:肖静
出版发行
发
印
版
印
开
印
字
定
行
刷
次
次
本
张
数
价
中国林业出版社(100009 北京市西城区德内大街刘海胡同 7 号)
https:// www.forestry.gov.cn/lycb.html
电话:(010)83143577
中国林业出版社
河北京平诚乾印刷有限公司
2020 年 9 月第 1 版
2020 年 9 月第 1 版
889mm×1194mm 1/16
32.75
400 千字
500.00 元
未经许可,不得以任何方式复制或抄袭本书的部分或全部内容。
© 版权所有,侵权必究。
FOREWORD
Dzhizak Province is the large region in the central part of
the Republic of Uzbekistan, which has a unique history,
great political, economic and spiritual status, and this
region has a great importance for the development of
science, culture and tourism.
Dzhizak Province is characterized by outstanding diversity
of natural landscapes and wildlife, including a large number
of endemic, rare species, and species of global economic
importance. There are represented different ecosystems
and cultural landscapes: deserts and lakes, foothills and
mountains, croplands and pastures, gardens and vineyards.
The territory has colourful natural landscapes. In the mountains of Zaamin, Bakhmal and Forish districts with
an exotic relief and temperate climate, there are juniper, almond and walnut forests, and valuable medicinal and
ornamental plants.
The plant diversity of the Dzhizak Province is extremely rich, and its detailed study is very
important. This book provides detailed information about almost 2000 wild-growing plant species
of the Dzhizak Province, as well as their images. This is the fruitful result of extensive joint research
performed by Uzbek and Chinese botanists.
Uzbekistan and China are neighbors and partners along the belt and road. In recent years, the
Academy of Sciences of the Republic of Uzbekistan has been working more and more closely with
the Chinese Academy of Sciences. In particular, the Institute of Botany, Academy of Sciences of the
Republic of Uzbekistan and the Kunming Institute of Botany, Chinese Academy of Sciences, within
the framework of projects devoted to the study and protection of plant diversity, sustainable use of
plant resources in the two countries, have made good achievements in joint scientific researches,
personnel training and other aspects. The Global Allium Garden with Kunming center and Tashkent
center has been created in collaboration of Chinese and Uzbek sides. These unique living collections
conserve and display the outstanding global diversity of Allium species, provide resource support
for systematic and thorough research on this group of plants, knowledge dissemination, education,
environmental protection, etc.
Academician Yuldashev Bekhzod Sadikovich
The President of Academy of Sciences
of the Republic of Uzbekistan
4
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
PREFACE
The Republic of Uzbekistan is located in the heart of the Eurasian continent and occupies a position
of great bio-geographical importance. The territory of Uzbekistan characterized by rich biological
and landscape diversity belongs to the Iran-Turanian region in the Tethyan (Ancient Mediterranian)
floristic subkingdom of Holarctic kingdom, which is one of major centers of plant diversity in the
world (Takhtajan, 1986). Ecosystems of Uzbekistan are included in the list of key ecological regions
of our planet (Olson & Dinerstein, 2002). Flora of vascular plants of Uzbekistan is rich with a large
number of crop wild relatives, relicts, endemic and endangered species. Considerable diversity of
plants can be explained by the geographic position and huge area of the country, as well as wide
spectrum of physiographic factors and complicated geological history. The unique centers of origin
and diversity of many genera are revealed on the territory of the country. However, it is necessary to
note that the ecosystems of the country have been influenced by human activities since ancient times;
anthropogenic impact has increased considerably in recent decades.
Floristic surveys in Uzbekistan were started more than 150 years ago, and the first edition of Flora
of Uzbekistan was published in 6 volumes (1941–1962). This treatment includes 4,148 species of
vascular plants (3,663 of them are native for Uzbekistan, and 485 are naturalized, alien and cultivated
taxa). However, at the present stage of our knowledge, this fundamental synopsis and the data on
distribution of many plant species were significantly outdated. Currently, non-published database of
the flora of Uzbekistan counts more than 4,375 species; thus, at least 712 wild-growing species were
added to the national checklist after issue of the first edition of the Flora of Uzbekistan (Sennikov et
al., 2016).
Detailed study of plant diversity became extremely important in the last decades in connection
with current trends and approaches in plant sciences and biodiversity conservation. Protection and
sustainable use of biodiversity are priority directions of state ecological policy in Uzbekistan. The
National Strategy and Action Plan on CBD (1998) includes identification, sampling and monitoring
of flora and vegetation (The Fifth National Report, 2015). Compilation of national and regional
floristic checklists has a special significance, as well as the assessment of populations of economic
valuable and endangered plants does so. The Flora of Uzbekistan Project aimed at taxonomical
revision of the national flora was started in 2012, and the first two volumes of the second edition of
PREFACE
Flora of Uzbekistan were published (Sennikov, 2016, 2017). Synopses of flora were compiled for
some large bio-geographical and administrative regions of the country.
In this book, the results of inventory of flora of the Dzhizak Province, a large administrative region
of the Republic of Uzbekistan, are represented for the first time.
The Dzhizak Province has been chosen as the study area because this territory is located in the
central part of the country and characterized by rich plant diversity with a large number of endemic
and rare species. There are represented all altitudinal zones and the majority of ecosystems of Central
Asia, including deserts, foothills, mountains, highlands and wetlands.
The synopsis of the flora of the Dzhizak Province has been compiled on the basis of the data of our
field surveys, analysis of TASH herbarium collections, publications, and scientific reports. We hope
that this book will become a reliable source for further studies and nature conservation planning,
and will be useful for botanists, ecologists, staff of nature reserves and national parks, students,
educators, and amateur-naturalists.
The study has been performed within the framework of the Uzbek state research projects I5FA-0-17440 and BA-FA-F5-010 and supported by the Second Tibetan Plateau Scientific ExPedition
and Research (STEP) Program (2019QZKK0502), the Strategic Priority Research Program of
Chinese Academy of Sciences (XDA20050203), the International Partnership Program of Chinese
Academy of Sciences (151853KYSB20160021) as well as the International Partnership Program of
Chinese Academy of Sciences (151853KYSB20180009). We would like to extend special thanks
to the Institute of Botany of Academy Sciences of the Republic of Uzbekistan, the Bureau of
International Cooperation of Chinese Academy of Sciences (CAS), the Kunming Institute of
Botany of CAS and the CAS Key Laboratory for Plant Diversity and Biogeography of East Asia for
supporting the study and to the China Forestry Publishing House for publishing this book.
5
6
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
CONTENTS
FOREWORD
/ 3
PREFACE
/ 4
STUDY AREA
/ 7
INVENTORY OF THE FLORA
/ 24
POLYPODIOPHYTA [FERNS]
/
33
GYMNOSPERMAE [GYMNOSPERMS]
/
39
ANGIOSPERMAE [ANGIOSPERMS]
/
47
LILIOPSIDA [MONOCOTS]
MAGNOLIOPSIDA [EUDICOTS]
/
135
REFERENCES
/
499
INDEX OF LATIN NAMES
/
507
STUDY AREA
The Dzhizak Province (Jizzax Viloyati in Uzbek) has been established in 1973. It is a large
administrative region of the Republic of Uzbekistan located in the central part of the country. The
total area is 21.210 km2 (approximately 5% of the country), 23% are occupied with arable lands,
2.2% with gardens and vineyards, 36% with pastures, and 7.7% are forested. The average population
density is 54 people/km2. The province is divided into 12 administrative districts: Arnasay, Bakhmal,
Dustlik, Farish, Gallaaral, Mirzachul, Pakhtakor, Sharaf Rashidov, Yangiabad, Zaamin, Zafarabad
and Zarbdar. The Dzhizak Province borders with the Syrdarya Province to the east and north-east,
with the Navoi Province to the west, with the Samarkand Province and Tajikistan to the south, and
with Kazakhstan to the north. The economy of the province is primarly based on agriculture; cotton
and wheat are the main crops, and extensive irrigation is used. Approximately 80% of population
lives in rural areas. The industry, mining operations and transportation infrastructure also are welldeveloped (www.jizzax.uz; www.stat.uz).
The study area is located between two large rivers, Syrdarya and Zeravschan. The territory of
the Dzhizak Province is divided into two physiographical parts (Figure A). The northern plain
part includes the desert of south-eastern Kyzylkum, the Aydar-Arnasay lake system located
in the large saline depression, and the Hungry Steppe depression in the middle reaches of the
Syrdarya River. The southern mountainous part includes Turkestan Ridge and its western spurs,
Malguzar and Nuratau mountains. The natural boundary between plain and mountainous part
of the Dzhizak Province runs approximately at 380–400 m.s.l.a in the west (piedmonts of the
Nuratau Ridge), and 480–500 m.s.l. in the east (piedmonts of the Turkestan Ridge). The highest
and lowest points are 4029 m (Mt. Shaukartau, Turkestan Ridge) and 240 m.s.l. (the shore of
Aydar-Arnasay lake system), respectively (Aramov, 2012).
The plain part of region has been transformed significantly due to human activity. Up to the middle
of the 20th Century, the Aydar salt marsh and a number of small salty lakes were located in the
large drainless depression in the south-eastern edge of Kyzylkum. With the beginning of intensive
agricultural development of the Hungry Steppe, this depression used to evacuate the drainage
water from irrigated lands. In 1969, the water level was extremely high; more than 21210 km2 of
a
m.s.l.=meter(s) above sea level.
8
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure A Physiographical map of the Dzhizak Province
water was discharged from the Chardara Reservoir, the Aydar salt marsh was flooded completely,
and a large drainless salty Aydar-Arnasay lake system was formed. At the present time, this
anthropogenic water body holds the 4th position in Central Asia in order of size, after the Aral Sea,
Balkhash and Issyk-Kul. In the past, an alluvial-proluvial plain on the left bank of the Syrdarya
River called the Hungry Steppe or Mirzachul was an almost waterless desert with ephemeroid
vegetation and fragments of salt marshes. At present, almost the entire territory of Hungry Steppe
is occupied by irrigated crop lands and settlements.
The climate is continental (Csa and BWk according to the Köppen climate classification),
with high amplitudes of daily, monthly and annual air temperature, low precipitation and high
intensity of solar radiation. In general, the winter is moderately cold, and summer is hot and dry.
There are significant differences among the climates of different natural regions of the province
related to their geomorphology. The continentality of climate and contrast seasonal changes is much
more expressed in the desert areas than in the mountains. The vast arid plain of Kyzylkum, located in
the north and north-west, and huge mountain ranges in the south and southeast have a great influence
on the climate of the Dzhizak Province.
Within the Dzhizak Province, the annual and monthly air temperature decreases from the south to the
north and from the plain to the mountains. In the plain and piedmonts, the mean annual temperature
STUDY AREA
ranges from 12°C to 15°C, the mean temperature of July is +27–+29°C, and the mean temperature of
January ranges from–4.4°C in the nortern part of the province to –0.1–0°C in piedmonts. The average
duration of the no frost period in plain and foothills ranges from 170 to 230 days. The warmest
area of the province is the piedmont plain of the Turkestan, Malguzar and Nuratau ridges (the mean
annual temperature reaches 15.1°C, and the mean temperature of July reaches 29.4°C). The coldest
area is the alpine zone of the Turkestan range (the mean annual temperature does not exceed 5°С,
and the mean temperature of July is 11–16°С). The lowest temperature recorded in the province is
–37°C, and the highest temperature is 48°C (Alibekov & Nishanov, 1978; Williams & Konovalov,
2008).
The amount of precipitation and humidity increases from the northwest to the southeast and from the
plain to the mountains. The mean annual precipitation is 150–200 mm in the plain, and 200–300 mm
in foothills. In the montane and alpine zones, this indicator ranges from 300–400 mm (Nuratau Mts.)
to 400–500 mm (Turkestan Ridge). Precipitation in the desert and foothill zones occurs mainly in
the winter and spring period; the maximum is in March–April, and the dry period is July–September.
In the mountains, the rainfall takes place throughout the year, but the maximum is in April–May
(Alibekov & Nishanov, 1978; Williams & Konovalov, 2008).
The Dzhizak Province is distinguished by outstanding diversity of flora and vegetation determined
by its geographical location, natural conditions and wide variety of landscapes and habitats. The
territory of the province is represented by all altitudinal zones, from the plain to highlands, the
majority of Central Asian vegetation types and most types of ecosystems including sandy and clay
desert, saline lands, wetlands, insular relic mountains, foothills, montane and alpine ecosystems. In
accordance with phytogeographical division of Uzbekistan, the territory of the Dzhizak Province
belongs to Nuratau and Kuhistan dictricts of Mountain Central Asian Province, and Middle Syrdarya
and Kyzylkum districts of the Turan Province (Tojibaev et al., 2016; Tojibaev et al., 2017). The study
area is divided into following phytogeographical regions: Nuratau, Nuratau Relic Mountains and
Aktau regions of the Nuratau district, North Turkestan and Malguzar regions of the Kuhistan dictrict,
Mirzachul region of the Middle Syrdarya district, and Kyzylkum region of the Kyzylkum district
(Figure B).
Northern and north-western part of the Dzhizak Province belongs to the Kyzylkum region of the
Kyzylkum district. This area includes the aeolian plain of the Southeastern Kyzylkum with fixed and semifixed sands, and the Aydar-Arnasay lake system with saline wetlands and small plots of tugay vegetation.
The sand desert is covered with psammophilous woodlands of saxaul, sand acacia and Calligonum
species, psammophilous shrubs, semishrubsand herbs (Zakirov, 1971, 1973, 1976, 1984).
Before 1993-1995, the shore of the Aydar-Arnasay lake system was fringed with a wide belt of tugay
vegetation (reeds and riparian woodlands of bloomy poplar, willow, oleaster and tamarisk). Since
1993, the Chardara Reservoir was overflowed with waters of the Syrdarya River several times. A
huge volume of water was drained into the Aydar-Arnasay lake system, the water surface level was
drastically increased, and large areas were flooded. Currently, tugay vegetation is represented mainly
9
10
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
with local plots of reeds, camel-thorn and tamarisks.
Figure B Phytogeographical regions of the Dzhizak Province (according to Tojibaev et al., 2017)
Notes:I. Central Asian Mountain Province: I-4 Nuratau district (I-4-a Nuratau, I-4-b Aktau, I-4-c Nuratau Relic Mountains). II.
Turan Province: II-2 Middle-Syrdarya district (II-2-b Mirzachul), II-3 Kyzylkum (II-3-a Kyzylkum). Protected areas: 1—Nuratau
Nature Reserve, 2—Zaamin Nature Reserve, 3—Zaamin National Park.
The central and north-eastern part of the Dzhizak Province belongs to the Mirzachul region of the
Middle Syrdarya district. The Mirzachul or Hungry Steppe is an alluvial-proluvial plain on the left
bank of the Syrdarya River in its middle reaches between the throat of the Fergana valley in the
east, the Chardara water reservoir, Aydar-Arnasay lake system and salt river Kly in the west, and
foothills of Turkestan and Malguzar ridges on the south. At present, this region is densely populated
and occupied by anthropogenic landscapes. Small areas of natural ecosystems with ephemeralephemeroid, saline and tugay vegetation can be found on the eastern shore of the Aydar-Arnasay
lake system, in the piedmonts of the Turkestan and Malguzar ridges, and along the irrigation network
(Zakirov, 1971, 1973, 1976, 1984).
The region of Nuratau Relic Mountains include the insular ridges, Pistalitau (557 m.s.l.),
Khanbandytag (476 m.s.l.), Egarbelistag (618 m.s.l.) and Balyklitau (581 m.s.l.), located at the
southern edge of the Kyzylkum, to the north of the Nuratau ridge. They stretch parallel to the Nuratau
Mountains from south-east to north-west. The largest of them is the Pistalitau ridge approximately
40 km long (Aramov, 2012). These small ridges are covered with sagebrush communities and
STUDY AREA
xerophytic shrublands. The general appearance of vegetation and dominant species is similar to
that in the lower belts of the Nuratau and other mountain ridges of the North-Western Pamir-Alay
(Batoshov & Beshko, 2013, 2015; Batoshov, 2016; Tojibaev et al., 2017).
The territory of the Nuratau region includes the northern branch of the Nuratau Mountains (or North
Nuratau Ridge) and is divided into two parts separated by the Saurbel pass: the Nuratau and Koytash
ridges. The length of this middle-altitude mountain chain is approximately 200 km, and the highest
peak named Khayat-bashi is 2169 m.s.l. (Aramov, 2012).
The Nuratau ridge was formed in the late Proterozoic and early Paleozoic as a result of the
Baikal and the Caledonian orogeny. It is one of the oldest mountains of Central Asia, almost
unaffected by the Alpine orogeny. During transgressions of the Tethys Sea (Cretaceous–
Paleogene), it was an isolated island. These circumstances contributed to the evolution of the
autochthonous flora. In general, the vegetation has xerophytic character with an original type
of vertical zonality determined by geographical position and natural conditions of the territory,
but the majority of mountainous Central Asian vegetation types (except for alpine communities)
are represented here. The piedmont plain named the Farish Steppe, and foothills of the Nuratau
ridge (400–800 m.s.l.) are covered with ephemeroid–sagebrush and phlomis–ephemeroid
communities. The stony slopes in the upper part of the foothills are occupied by petrophytic
ephemeral–sagebrush vegetation and xerophytic shrublands. The montane zone (from 800
m.s.l. up to the crest of the ridge) has mosaic vegetation represented by tall grass and tall herb
communities (so-called savannoids), forb-sagebrush, semishrub and dwarf shrub communities
(so-called phryganoids), bunch grass steppes, spiny herbs and pulvinates (so-called mountain
xerophytes), shrubs and open woodlands. This area is the most western point of the distribution
of forests of Juniperus polycarpos var. seravschanica. In the past, juniper woodlands grew in
the Nuratau Mountains, but they were cut down almost entirely by the beginning of the 20th
century and were replaced with scrublands and tall grass communities. Small populations and
solitary trees of Zeravshan juniper were still preserved in some remote places (Zakirov P., 1969,
1971; Zakirov K., 1971, 1973, 1976, 1984; Tojibaev et al., 2017).
The Nuratau ridge is one of the three isolated sites of the relict walnut-fruit forests in Uzbekistan.
The walnut-fruit stands occupy the valleys of the largest streams on the northern slope of the Nuratau
ridge. Currently, they represent an original cultural landscape (so-called forest-garden) that has been
transformed extensively by local people over the centuries-old horticultural activities. These gallery
woodlands with 0.7–0.8 degree of density are composed by Juglans regia L., Malus sieversii (Ledeb.)
M. Roem, Prunus armeniaca L., Morus alba L., Acer semenovii Regel & Herder, Crataegus
turkestanica Pojark., etc. Walnut-fruit forests of the Nuratau ridge accounts approximately 16.7
percent of the national area.
In central part of Nuratau ridge, there is situated the Nuratau Nature Reserve (17.752 hectares,
among which 2.303 are forested), a strictly protected area of Category I IUCN established in 1975.
Flora of the Nuratau Nature Reserve numbers 835 species, 33 of them are red-listed at the national
11
12
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
level (Beshko et al., 2013).
Aktau phytogeographical region includes the southern branch of the Nuratau Mountains, which
consists of several small ridges. Within the study area, it is represented with the north slopes of
Khobduntau and Karachatau ridges (1672 and 1101 m.s.l., respectively) situated in the south-western
part of the Dzhizak Province (Aramov, 2012). Vegetation of of this territory is similar with foothills
and lower part of montane zone of the Nuratau ridge.
The southern and south-eastern part of the Dzhizak Province belongs to the North Turkestan
phytogeographical region. This area covers the northern slope of the Turkestan ridge with the
complete profile of altitudinal zones. The zone of foothills lies here between of 480–500 and 1000–
1300 m.s.l, montane zone is situated between 1000–1300 and 2800–3000 m.s.l., and upper part
of the ridge (up of 2800–3000 m.s.l.) belongs to the alpine zone. The highest peak of the Dzhizak
Province, Mt. Shaukartau (4029 m.s.l.) is located here (Aramov, 2012). This is the most humid area
of the Dzhizak Province. The flora and vegetation of this area is distinguished by the significant
diversity due to the strongly rugged terrain, large amplitude of elevations, and wide range of soil and
climatic conditions. There are represented the semi-arid version of all piedmont and mountainous
types of landscapes of Central Asia and almost all mountain types of vegetation. The piedmont
plain and foothills are characterized by ephemeral-sagebrush and low herb ephemeral–ephemeroid
communities, often with saltworts. Tall grass savannoids and xerophytic open woodlands replace
them at 1000–1300 m.s.l. In river valleys, there are small plots of walnut–fruit forests. A thick belt
of juniper forests begins at 1500–1700 m.s.l. In some areas, the juniper trees form stands with a
stock density of 0.6–1.0. Juniperus polycarpos var. seravschanica(Kom.)Kitam., J. semiglobosa
Regel, and J. pseudosabina Fisch. & C. A. Mey. dominates in the lower, middle, and upper part of
the juniper belt, respectively. Despite the rich diversity of deciduous trees and shrubs represented
with maple, hawthorn, Sievers apple, cherry-plum, honeysuckle, and other species, broad-leaved
woodlands not play a notable role in the landscape.The vegetation of highlands is comprised of
juniper elfin wood, bunchgrass steppes, mountain xerophytes and alpine meadows (Demurina, 1975;
Konnov, 1973, 1990; Zakirov K., 1971, 1973, 1976, 1984; Esankulov, 2012; Khassanov et al., 2013;
Tojibaev, 2017).
The Zaamin Nature Reserve is situated here, it is one of the oldest nature reserves in Central Asia
established in 1926 under the name Guralash Reserve. At the present time, the area of the nature
reserve is 26.840 hectares (22.137 hectares are forested). The check-list of flora of the Zaamin
Nature Reserve includes 1216 plant species (Khassanov et al., 2013). In the east, the reserve borders
with the territory of the Zaamin National Park (23.894 hectares) established in 1976. A synopsis of
the flora of the national park is still incomplete (Beshko et al., 2013a).
The Malguzar phytogeographical region includes the Malguzar ridge (2620 m.s.l.), a northwestern spur of the Turkestan ridge stretching about 80 km and separated from the Turkestan
ridge by the Zaaminsu River valley in the east and the Sanzar river valley in the south (Aramov,
2012). The gorge called Tamerlane’s Gate is the border between the Malguzar and Nuratau
STUDY AREA
ridges. The northern slope of the ridge faces the Hungry Steppe, which is almost completely
occupied by arable land. The vegetation of Malguzar and the northern slope of the Turkestan
ridge is similar (Demurina, 1975; Zakirov K., 1971, 1973, 1976, 1984). There are arid and
semi-arid piedmont and mountain landscapes, and most mountain vegetation types (except
for highlands). As on the Turkestan ridge, the foothills are covered with ephemeroid and
ephemeroid–sagebrush communities. The vegetation of the montane zone is represented with
savannoids, fescue steppes, xerophytic shrubs, juniper woodlands with small plots of broadleaved stands. The natural boundary between foothills and montane zone lies at the same
elevations as in the northern slope of the Turkestan ridge. The differences between the flora and
vegetation of the Malguzar and North Turkestan phytogeographical regions (e.g., the absence of
alpine plants and communities) are explained by the relatively low altitude and xeric conditions
of the Malguzar ridge (Demurina, 1975; Azimova, 2017; Tojibaev et al., 2017).
Uzbekistan is the most densely populated country in Central Asia, and the ecosystems of the Dzhizak
Province have been influenced by human activities since ancient times. Anthropogenic impact has
increased considerably since the second half of the 20th century. Indigenous vegetation communities
have been transformed extensively, except for remote desert and mountain areas. In the Hungry
Steppe, natural ecosystems have been replaced almost completely by arable lands and settlements. In
the piedmonts, large areas are covered by rainfed crops and fallow lands with ruderal vegetation. In
the mountainous part of the Province, the main threats influencing the plant diversity are excessive
grazing and clear-cutting of trees and shrubs. Forests and open woodlands are under serious threat
from over-exploitation, over-grazing and desertification. In this connection, the study and analysis
of historical and current distribution of plant species and communities within this large region of the
country is extremely important.
Landscapes of South-eastern Kyzylkum and sandy desert with a giant Umbelliferae Ferula foetida(photography by N.
Yu. Beshko)
13
14
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Landscapes of South-eastern Kyzylkum and psammophilous woodlands of white saxaul Haloxylon persicum (photography by N. Yu. Beshko)
STUDY AREA
Aydar-Arnasay lake system(photography by N. Yu. Beshko)
Aydar-Arnasay lake system, a view from the ridge Pistalitau (photography by N. Yu. Beshko)
15
16
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
The insular ridge Balyklitau (photography by N. Yu. Beshko)
Piedmonts of the ridge Nuratau (photography by N. Yu. Beshko)
STUDY AREA
Landscapes of the Nuratau ridge (photography by N. Yu. Beshko)
17
18
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Landscapes of the Nuratau ridge and the rock massif Parrandas (photography by N. Yu. Beshko)
Landscapes of the Nuratau ridge and spiny almond communities in foothills (photography by N. Yu. Beshko)
STUDY AREA
Landscapes of the Nuratau ridge and the walnut-fruit gallery forest (photography by N. Yu. Beshko)
19
20
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Landscapes of the Nuratau ridge and tall grass communities and xerophytic shrublands (photography by N. Yu. Beshko)
Landscapes of the Nuratau ridge and mountain xerophytes (photographby N. Yu. Beshko)
STUDY AREA
Landscapes of the Turkestan ridge and juniper forests (photography by N. Yu. Beshko)
21
22
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Landscapes of the Malguzar ridge (photography by N. Yu. Beshko)
STUDY AREA
Landscapes of Mirzachul (Hungry Steppe) and Papaver pavoninum on the fallow lands (photography by N. Yu. Beshko)
Landscapes of Mirzachul (Hungry Steppe) and saline lands (photography by N. Yu. Beshko)
23
24
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
INVENTORY OF THE FLORA
First botanical expeditions in the territory that currently belongs to the Dzhizak Province were
conducted in the Sanzar river basin and surroundings of the town Dzhizak by A. Fedtschenko (1866),
P. Capus and G. Bonvalot (1881), A. Regel (1882), and V. I. Lipsky (1890). Many naturalists and
officials, who worked in Central Asia in the late 19th century, also collected herbarium specimens and
contributed to the accumulation of floristic data.
The systematic and purposeful study of the flora and vegetation of Central Asia began in the 1910s.
Several significant botanical expeditions were carried out in the first half of the 20th century and in
1950 –1970s on the Turkestan, Malguzar and Nuratau Ranges and in the Hungry Steppe. The studies of
this period have been focused on applied purposes, mainly on an investigation of the vegetation cover
and resource plants. The most important collections in this region were made by B. A. Fedtschenko,
E. P. Korovin, V. P. Drobow, M. G. Popov, I. I. Sprygin, M. V. Kultiassov, B. S. Zakrzczewsky, S. N.
Kudrjashev, P. A. Gomolitsky, M. M. Sovetkina, E. M. Demurina, E. E. Korotkova, O. E. Knorring, A. I.
Vvedensky, L. L. Bulgakova, L. I. Nazarenko, P. K. Zakirov, and R. V. Kamelin. Essay on the Vegetation
of the Pistalitau Mountains by M. V. Kultiassov (1923), Plant Formations of the Nurata Valley (1923) and
Vegetation of Middle Asia and South Kazakhstan (1934; 1961; 1962) by E. P. Korovin, Vegetation of the
Khobduntau and Karachatau Mountains by S. N. Kudrjashev (1930), Vegetation of the Guralash Nature
Reserve and Zaamin Forestry by M. G. Popov and N. V. Androssov (1937), Flora and Vegetation of the
Zeravschan River Basin by K. Z. Zakirov (1955, 1962), Vegetation Cover of the Nuratau Mountains (1969)
and Botanical Geography of Low Mountains of Kyzylkum and Nuratau Ridge (1971) by P. K. Zakirov,
Vegetation of the Western Part of Turkestan Ridge and Its Spurs by E. M Demurina (1975), To the
Knowledge of the Flora of Nuratau Mountains (1973) and The Kuhistan District of Mountainous Middle
Asia (1979) by R.V. Kamelin should be cited as the most important publications of the second half of the
20th century dedicated to the plant diversity of this region. Several new studies have been performed since
2000 (Beshko, 2000a, 2000b, 2011; Tirkasheva, 2011; Botirova, 2012; Esankulov, 2012; Khassanov
et al., 2013; Batoshov, 2016; Azimova, 2017), a number of new species have been described from this
territory and dozens of new records were found. Analysis of obtained data allows us to give a summary of
floristic studies in the Dzhizak Province.
The synopsis of flora of the Dzhizak Province of the Republic of Uzbekistan has been compiled for
INVENTORY OF THE FLORA
the first time as a result of analysis of a huge volume of data of our long-term field surveys, revision of
herbarium material stored in TASH, LE and MW, publications, and reports (Azimova, 2017; Batoshov &
Beshko, 2013, 2015; Batoshov, 2016; Belolipov, 1973; Beshko, 2000a, 2000b, 2011; Beshko & Azimova,
2013, 2014; Beshko et al., 2013, 2014; Botirova, 2012; Botschantzev et al., 1961; Conspectus Florae
Asiae Mediae (1963–1993); Demurina, 1975; Esankulov, 2012; Flora of Tajikistan (1957–1991); Flora of
Uzbekistan (1941–1962); Kamelin, 1973a, 1973b, 1979, 1990; Khassanov, 2015; Khassanov et al, 2013;
Konnov, 1973, 1990; Korovin, 1923; Kudrjashev, 1930; Kultiassov, 1923; Popov & Androsov, 1937;
Sennikov, 2016, 2017; Tirkasheva, 2011; Tojibaev & Beshko, 2007, 2015; Tojibaev et al., 2015; Zakirov,
1955, 1962; Zakirov & Burygin, 1956; Zakirov, 1969, 1971).
The Flora of the Dzhizak Province, Uzbekistan contains information about plant species arranged in
the following order: currently accepted scientific name, commonly-used synonymy (optional), life
form, habitat, altitudinal zone, distribution in the Province, character of use, conservation status (for
nationally or globally red-listed species).
Families and genera of ferns in the checklist are arranged in accordance with the modern
classification that was recently proposed by the Pteridophytes Phylogeny Group (Christenhusz et
al., 2011a), Gymnosperms are arranged according to Christenhusz et al. (2011b), flowering plants are
arranged in accordance with the APG system (APG IV, 2016). Species are arranged in alphabetic order.
Accepted names of species are provided in accordance with The Plant List database (www.theplantlist.
org), databases and recently published treatments of different taxonomical groups (Akhani et al., 2007;
Al-Shehbaz et al., 2014; Clayton et al., 2006; Degtjareva et al., 2013; Downie et al., 2010; Egorova,
1999; http://www.e-monocot.org/; Hill, 2017; Horandl & Emadzade, 2012; www.ildis.com; Khassanov
& Rakhimova N. 2012; Lammers, 2007; Lazkov, 2016; Lopez-Vinyallonga et al., 2009, 2011; Podlech
& Zarre, 2013; Salmaki et al., 2012; Sennikov, 2016, 2017; The Global Compositae Checklist, 2009;
The Gymnosperm Database, 2017; The World Umbellifer Database, 1999; Tsvelev, 1983; Wiegleb et al.,
2017). Citation of authorship of taxa corresponds to the Authors of Plants Names (Brummitt & Powell,
1992) and the International Plant Names Index (www.ipni.org).
Altitudinal distribution is provided with the following elevation zones: plains, foothills, montane
zone, and alpine zone (their natural boundaries were described above). The distribution of species
within the Dzhizak Province was indicated according to the scheme of phytogeographical regions of
Uzbekistan (Tojibaev et al., 2017).
Character of economical use of plants is done in accordance with published data (Plant Resources
of the U. S. S. R., 1984–1996; Larin, 1956; Gammerman & Grom, 1976; Nikitin, 1983; Kurmikov &
Belolipov, 2012, etc.).
IUCN RL is the abbreviation for the IUCN Red List (www.iucnredlist.org). UzbRDB is the abbreviation
for the Red Data Book of Uzbekistan (2009), and categories of threatened species used in that book are
employed as follows: 0 (extinct species) — corresponds to EX or EW categories of the IUCN Red List, 1
(endangered, disappearing species) — meets CR or EN categories of IUCN, 2 (rare species) — meets VU
25
26
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
category of IUCN and 3 (declining species) — corresponds to NT category of IUCN.
The checklist presented below includes 1965 species of 613 genera and 105 families of vascular
plants occurring in the wild in the study area including naturalized aliens. Alien taxa are marked with
an asterisk. The assessment of alien status is based on analysis of publications and online databases
(Nikitin, 1983; IUCN/ISSG, 2014; CABI, 2017).
There are 44 species of trees and 116 shrubs, among which 11 species can have a habit of tree or shrub;
108 species of semi-shrubs; 1062 perennial herbs, among which 61 biennial, 516 annual; 46 herbaceous
plants can have different life forms (annual, biennial or perennial). The number of plant species recorded
for different phytogeographical regions of the Dzhizak Province is shown in Table 1. The leading families
and genera are represented in Table 2, and ratio of life forms is shown in Table 3.
Only 5 species of the flora of the Dzhizak Province are assessed as globally threatened (CR, EN, VU)
on the IUCN Red List (www.iucnredlist.org), and 3 species are assessed as Near Threatened (Table
4). Only one species among them (Lonicera paradoxa Pojark.) is red-listed at the national level.
The status of Platycladus orientalis (L.) Franco is uncertain because the species was naturalized in
different Asian countries in ancient times, but many botanists regarded Platycladus in Uzbekistan
as native and Tertiary relic. The list of plants included in the Red Data Book of Uzbekistan is
represented with 51 species, most of them (42 species) occur within protected areas (Table 5).
Following 20 species of the flora of the Dzhizak Province are endemic to the Nuratau Mountains:
Acantholimon nuratavicum Zakirov ex Lincz., Acantholimon subavenaceum Lincz., Acantolimon
zakirovii Beshko, Arctium pallidivirens (Kult.) S. López, Romaschenko, Susanna & N. Garcia [Anura
pallidivirens (Kult.) Tscherneva], Cousinia botschantzevii Juz. ex Tscherneva, Dianthus helenae Vved.,
Dracoсephalum nuratavicum Adylov, Eremurus nuratavicus A. P. Khohkr., Helichrysum nuratavicum
Krasch., Jurinea zakirovii Iljin, Lagochilus olgae Kamelin, Lagochilus proskorjakovii Ikramov, Lappula
nuratavica Nabiev & Zakirov, Lepidium olgae (R. M. Vinogr.) Al-Shehbaz & Mummenhoff (Stubendorffia
olgae R.Vinogradova), Oxytropis pseudorosea Filim., Parrya nuratensis Botsch. & Vved., Phlomis
nubilans Zakirov, Phlomoides anisochila (Pazij & Vved.) Salmaki, Salvia submutica Botsch. & Vved.,
Thymus subnervosus Vved., Nabiev & Tulyag. One species, Ferula helenae Rakhm. & Melibaev, is
endemic to the insular low mountains located to the north of the Nuratau ridge.
One species, Eremurus chloranthus Popov, is endemic to the Turkestan ridge. Two species are
endemic to the Malguzar ridge: Allium levichevii F. O. Khass. & N. Sulejm. and Oxytropis kamelinii
Vassilcz. Following 13 species occurring in Turkestan and Malguzar ridges are endemic to the
Kuhistan phytogeographical district: Allium alexeianum Regel, Astragalus belolipovii Kamelin
ex F. O. Khass. & N. Sulajm., Astragalus russanovii F. O. Khass., Sarybaeva & Esankulov,
Dianthus subscabridus Lincz., Euphorbia rosularis Fed., Ferula ovczinnikovii Pimenov, Jurinea
helichrysifolia Popov ex Iljin, Oxytropis capusii Franch., Oxytropis seravschanica Gontsch. ex
Vassilcz. & B. Fedtsch., Primula iljinskii Fed., Scutellaria schachristanica Juz., Serratula lancifolia
Zakirov, Tanacetopsis urgutensis (Popov ex Tzvelev) Kovalevsk. One species, Bryonia melanocarpa
Nabiev, is endemic to the South-eastern Kyzylkum.
INVENTORY OF THE FLORA
The identification of economically valuable species is an important result of the inventory of the
flora of vascular plants of the Dzhizak Province. The most numerous group among them are fodder
plants (558 species); 446 species are medicinal plants (officinal and used in folk medicine); 407
species are meliferous, 388 ornamental, 154 essential oil, 123 dye, 136 food plants (including wildgrowing vegetables, fruits, berries, nut-bearing, spicy-aromatic species, etc.); 89 species can be
used for afforestation and soil stabilization. At the same time, various ruderal, segetal and pastoral
weeds (355 species), as well as poisonous plants (99 species) and alien species (114) are widespread
in the Dzhizak Province. This fact indicates that ecosystems of study area have been noticeably
transformed by anthropogenic impact.
Table 1. The number of species recorded for different phytogeographical regions of
the Dzhizak Province
Phytogeographical regions
Number of species
Mountain Central Asian Province
North Turkestan
1463
Malguzar
1255
Nuratau
1139
Nuratau Relic Mountains
614
Aktau
541
Turan Province
Mirzachul
590
Kyzylkum
456
Table 2. Leading families and genera of the flora of the Dzhizak Province
Number of
species
Percentage
(%)
107
5.45
Allium
37
1.89
8.82
Cousinia
34
1.73
113
5.76
Artemisia
27
1.38
29
95
4.84
Silene
25
1.27
Apiaceae
41
91
4.64
Ranunculus
22
1.12
Amaranthaceae
29
86
4.38
Oxytropis
21
1.07
Caryophyllaceae
25
83
4.23
Taraxacum
20
1.02
Rosaceae
18
66
3.36
Ferula
19
0.97
Boraginaceae
21
58
2.96
Carex
18
0.92
Ranunculaceae
18
57
2.91
Galium,
Polygonum,
Salsola, Stipa
16
0.82
Number of
genera
Number of
species
Percentage
(%)
Asteraceae
82
281
14.32
Astragalus
Fabaceae
26
210
10.70
Poaceae
58
173
Brassicaceae
57
Lamiaceae
Families
Genera
27
28
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Table 3. The ratio of life forms of the flora of the Dzhizak Province
Life form
Number of species
Percentage(%)
Trees
44
2.24
Shrubs
116
5.91
11
0.56
108
5.50
1064
54.17
Biennial herbs
61
3.11
Annual herbs
516
26.22
46
2.34
Trees or shrubs
Semi-shrubs
Perennial herbs
Annual, biennial or perennial herbs
Table 4. List of plants of the flora of Dzhizak Province included in the IUCN Red List
No
Species
Category
Distribution in the Dzhizak Province
(phytogeographical regions and protected areas)
1
Prunus bucharica
(Korsh.) B. Fedtsch. ex
Rehder[Amygdalus
bucharica Korsh.](Rosaceae)
VU
B2ab(iii,v)
Nuratau, Aktau, North Turkestan, Malguzar. Nuratau
Reserve, Zaamin Reserve, Zaamin National Park.
Dominant species of open woodlands widely
distributed in mountainous part of the province
2
Prunus armeniaca
L.[Armeniaca vulgaris Lam.]
(Rosaceae)
EN B2ab(iii)
Nuratau, North Turkestan, Malguzar. Nuratau Reserve,
Zaamin Reserve, Zaamin National Park
3
Fraxinus sogdiana Bunge
(Oleaceae)
NT
Nuratau, North Turkestan, Malguzar. Nuratau Reserve,
Zaamin Reserve, Zaamin National Park
4
Juglans regia L.
(Juglandaceae)
NT
Nuratau, North Turkestan, Malguzar. Nuratau Reserve,
Zaamin Reserve, Zaamin National Park
5
Lonicera paradoxa Pojark.
(Caprifoliaceae)
EN B2ab(iii,v)
UzbRDB 1
6
Malus sieversii (Ledeb.) M.
Roem (Rosaceae)
VU A2cde
7
Platycladus orientalis (L.)
Franco (Cupressaceae)
NT
8
Pyrus korshinskyi Litv.
(Rosaceae)
CR B2ab(iii,v)
North Turkestan
Nuratau, Aktau, North Turkestan, Malguzar. Nuratau
Reserve, Zaamin Reserve, Zaamin National Park
Nuratau. Nuratau Reserve.One giant “holy” tree in the
valley Madjrum ca. 2000 years old
Malguzar. Only one location is known
INVENTORY OF THE FLORA
Table 5. List of plants of the flora of the Dzhizak Province included in the Red Data Book of
Uzbekistan
No
Species
Category of
UzbRDB
Distribution in the Dzhizak Province
(phytogeographical regions and protected areas)
1
Acantholimon nuratavicum
Zakirov ex Lincz.
(Plumbaginaceae)
2
Nuratau. Nuratau Reserve
2
Aconitum talassicum Popov
(Ranunculaceae)
3
North Turkestan, Malguzar. Zaamin Reserve,
Zaamin National Park
3
Allium isakulii R. M. Fritsch & F.
O. Khass. (Amaryllidaceae)
2
Nuratau, Malguzar. Nuratau Reserve
4
Allium praemixtum Vved.
(Amaryllidaceae)
1
Nuratau, Malguzar. Nuratau Reserve
5
Allochrusa gypsophiloides Regel
(Caryophyllaceae)
3
Nuratau, Malguzar, North Turkestan
6
Arctium haesitabundum (Juz.) S.
López, Romasch., Susanna & N.
Garcia [Cousinia haesitabunda
Juz.] (Asteraceae)
1
Nuratau, Malguzar
7
Arctium pallidivirens (Kult.) S.
López, Romaschenko, Susanna & N.
Garcia [Anura pallidivirens (Kult.)
Tscherneva] (Asteraceae)
1
Nuratau, Malguzar. Nuratau Reserve
8
Astragalus belolipovii Kamelin
ex F. O. Khass. & N. Sulajm.
(Fabaceae)
1
North Turkestan, Malguzar. Zaamin Reserve
9
Astragalus kelleri Popov
(Fabaceae)
2
Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar. Nuratau Reserve
10
Astragalus knorringianus Boriss.
(Fabaceae)
2
Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Nuratau Reserve,
Zaamin Reserve, Zaamin National Park
11
Astragalus leptophysus Vved.
(Fabaceae)
2
Nuratau, North Turkestan. Nuratau Reserve
12
Bryonia melanocarpa Nabiev
(Cucurbitaceae)
2
Kyzylkum
13
Cicer grande (Popov) Korotkova
(Fabaceae)
2
Nuratau. Nuratau Reserve
14
Colchicum kesselringii Regel
(Colchicaceae)
3
Nuratau, Malguzar, North Turkestan. Nuratau
Reserve, Zaamin Reserve, Zaamin National Park
15
Cousinia dshizakensis Kult.
(Asteraceae)
2
Nuratau, Aktau, Malguzar, North Turkestan.
Nuratau Reserve, Zaamin Reserve, Zaamin
National Park
16
Eremurus chloranthus Popov
(Asphodelaceae)
0
North Turkestan. Zaamin Nature Reserve. Only
type specimen collected in 1926 is known
29
30
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
(continued)
No
Species
Category of
UzbRDB
Distribution in the Dzhizak Province
(phytogeographical regions and protected areas)
17
Eremurus lactiflorus O. Fedtsch.
(Asphodelaceae)
2
Nuratau. Nuratau Nature Reserve
18
Eremurus nuratavicus A. P.
Khokhr. (Asphodelaceae)
2
Nuratau. Nuratau Nature Reserve
19
Eremurus robustus (Regel) Regel
(Asphodelaceae)
3
Nuratau, Malguzar, North Turkestan. Nuratau
Reserve, Zaamin Reserve, Zaamin National Park
20
Ferula fedtschenkoana Koso-Pol.
(Apiaceae)
1
Malguzar, North Turkestan. Zaamin Reserve,
Zaamin National Park
21
Ferula moschata (H. Reinsch)
Koso-Pol. [Ferula sumbul
(Kauffm.) Hook. f.] (Apiaceae)
2
Nuratau, Malguzar, North Turkestan. Nuratau
Reserve, Zaamin Reserve, Zaamin National Park
22
Gladiolus italicus Mill. (Iridaceae)
1
North Turkestan. Only one herbarium specimen
collected in 1934 is known
23
Helichrysum nuratavicum Krasch.
(Asteraceae)
2
Nuratau. Nuratau Reserve
24
Jurinea zakirovii Iljin (Asteraceae)
1
Nuratau. Nuratau Reserve
25
Lagochilus inebrians Bunge
(Lamiaceae)
2
Nuratau, Aktau, Malguzar. Nuratau Reserve
26
Lagochilus olgae Kamelin
(Lamiaceae)
2
Nuratau. Nuratau Reserve
27
Lagochilus proskorjakovii Ikramov
(Lamiaceae)
1
Nuratau. Nuratau Reserve
28
Lappula nuratavica Nabiev &
Zakirov (Boraginaceae)
2
Nuratau. Nuratau Reserve
29
Lepidium olgae (R. Vinogradova)
Al-Shehbaz & Mummenhoff
[Stubendorffia olgae R.
Vinogradova] (Brassicaceae)
2
Nuratau. Nuratau Reserve
30
Lepidolopha nuratavica Krasch.
(Asteraceae)
1
Nuratau. Nuratau Reserve
31
Lipskya insignis Nevski (Apiaceae)
3
Malguzar
32
Lonicera paradoxa Pojark.
(Caprifoliaceae)
1
North Turkestan
33
Oxytropis pseudorosea Filim.
(Fabaceae)
2
Nuratau. Nuratau Reserve
34
Paeonia tenuifolia L. [P. hybrida
Pall.] (Paeoniaceae)
3
North Turkestan. Zaamin Nature Reserve. Only one
location is known
35
Phlomoides anisochila (Pazij &
Vved.) Salmaki (Lamiaceae)
1
Nuratau. Nuratau Reserve
36
Phlomis nubilans Zakirov
(Lamiaceae)
2
Nuratau. Nuratau Reserve
INVENTORY OF THE FLORA
(continued)
No
Species
Category of
UzbRDB
Distribution in the Dzhizak Province
(phytogeographical regions and protected areas)
37
Parrya olgae (Regel & Schmalh)
D. A. German & Al. Shehbaz
[Pseudoclausia olgae (Regel &
Schmalh.] (Brassicaceae)
2
Nuratau, Malguzar, North Turkestan. Nuratau
Reserve, Zaamin Reserve, Zaamin National Park
38
Parrya sarawschanica (Regel &
Schmalh.) D. A. German &
Al-Shehbaz [Pseudoclausia
sarawschanica (Regel & Schmalh.)
Botsch.] (Brassicaceae)
2
Nuratau. Nuratau Nature Reserve
39
Pseudosedum campanuliflorum
Boriss. (Crassulaceae)
1
North Turkestan, Malguzar. Zaamin National Park
40
Salsola titovii Botsch.
(Chenopodiaceae)
1
Aktau
41
Salvia submutica Botsch. & Vved.
(Lamiaceae)
2
Nuratau. Nuratau Reserve
42
Saxifraga hirculus L.
(Saxifragaceae)
2
North Turkestan. Zaamin Reserve, Zaamin
National Park
43
Serratula lancifolia Zakirov
(Asteraceae)
1
Malguzar
44
Seseli turbinatum Korovin
(Apiaceae)
2
Nuratau. Nuratau Reserve
45
Silene paranadena Bondarenko &
Vved. (Caryophyllaceae)
2
Nuratau, Malguzar. Nuratau Reserve
46
Tulipa affinis Botschantz
(Liliaceae)
2
Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Nuratau Reserve,
Zaamin Reserve, Zaamin National Park
47
Tulipa dasystemon Regel
(Liliaceae)
2
North Turkestan, Malguzar. Zaamin Reserve,
Zaamin National Park
48
Tulipa korolkowii Regel (Liliaceae)
2
Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Nuratau Reserve,
Zaamin Reserve, Zaamin National Park
49
Tulipa lehmanniana Mercklin
(Liliaceae)
3
Kyzylkum
50
Tulipa micheliana Hoog
(Liliaceae)
2
Nuratau, Nuratau Relic Mountains, Malguzar.
Nuratau Reserve
51
Vitis vinifera L. (Vitaceae)
2
Nuratau. Nuratau Reserve
31
32
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
POLYPODIOPHYTA [FERNS]
POLYPODIOPHYTA
[FERNS]
33
34
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 1. Equisetaceae
Genus 1. Equisetum L.
1. Equisetum arvense L.
Perennial. Banks of rivers, wet places. Plain, foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Medicinal.
2. Equisetum ramosissimum Desf. Figure 1
Perennial. Banks of rivers, wet places, meadows. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, fodder.
Figure 1 Equisetum ramosissimum (photography by Natalya Beshko)
Family 2. Ophioglossaceae
Genus 2. Botrychium Sw.
3. Botrychium lunaria (L.) Sw.
Perennial. Wet meadows, moraines. Montane and alpine zones. North Turkestan.
POLYPODIOPHYTA [FERNS]
Family 3. Marsileaceae
Genus 3. Marsilea L.
4. Marsilea quadrifolia L.
Perennial. Standing and slowly flowing water, rice fields. Plain. Kyzylkum, Mirzachul.
Family 4. Salviniaceae
Genus 4. Salvinia Micheli
5. Salvinia natans (L.) All.
Perennial. Standing and slowly flowing water. Plain. Kyzylkum, Mirzachul.
Family 5. Pteridaceae
Genus 5. Cheilanthes Sw.
6. Cheilanthes persica (Bory) Mett. ex Kuhn. Figure 2
Perennial. Rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
Figure 2 Cheilanthes persica (photography by Natalya Beshko)
35
36
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 6. Adiantum L.
7. Adiantum capillus-veneris L. Figure 3
Perennial. Wet rocks and boulders near waterfalls, stream banks. Montane zone. Nuratau, Malguzar,
North Turkestan. Ornamental, medicinal.
Figure 3 Adiantum capillus-veneris (photography by Natalya Beshko)
Family 6. Aspleniaceae
Genus 7. Cystopteris Berh.
8. Cystopteris fragilis (L.) Bernh. Figure 4
Perennial. Shady wet places. Montane zone. Nuratau, Aktau, Malguzar, North Turkestan.
Figure 4 Cystopteris fragilis (photography by Natalya Beshko)
POLYPODIOPHYTA [FERNS]
Genus 8. Asplenium L.
9. Asplenium ruta-muraria L. Figure 5
Perennial. Rocks. Montane zone. Nuratau, Malguzar, North Turkestan. Ornamental, medicinal.
Figure 5 Asplenium ruta-muraria(photography by Natalya Beshko)
10. Asplenium trichomanes L.
Perennial. Rocks. Montane zone. Nuratau, Malguzar, North Turkestan. Ornamental, medicinal.
Genus 9. Ceterach Gorsault.
11. Ceterach officinarum Willd. Figure 6
Perennial. Rocks. Montane zone. Malguzar, North Turkestan. Ornamental, medicinal.
37
38
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 6 Ceterach officinarum (photography by Natalya Beshko)
Family 7. Polypodiaceae
Genus 10. Dryopteris Adans.
12. Dryopteris filix-mas (L.) Schott.
Perennial. Shady wet places. Montane zone. Malguzar, North Turkestan. Ornamental, medicinal.
GYMNOSPERMAE [GYMNOSPERMS]
GYMNOSPERMAE
[GYMNOSPERMS]
39
40
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 8. Ephedraceae
Genus 11. Ephedra L.
13. Ephedra botschantzevii Pachom.
Shrub. Stony slopes. Foothills, montane zone. Nuratau.
14. Ephedra equisetina Bunge Figure 7
Shrub. Stony and fine-earth slopes, rocks, screes. Plain, foothills, montane and alpine zones. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal.
Figure 7 Ephedra equisetina (photography by Natalya Beshko)
15. Ephedra fedtschenkoae Paulsen
Shrub. Stony slopes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
GYMNOSPERMAE [GYMNOSPERMS]
16. Ephedra foliata Boiss. ex C. A. Mey. [Ephedra kokanica Regel] Figure 8
Shrub. Stony and fine-earth slopes. Foothills, montane zone. Nuratau, Nuratau Relic Mountains,
Malguzar, North Turkestan.
Figure 8 Ephedra foliata (photography by Natalya Beshko)
41
42
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
17. Ephedra intermedia Schrenk & C. A. Mey. Figure 9
Shrub. Stony and fine-earth slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal.
Figure 9 Ephedra intermedia (photography by Natalya Beshko)
GYMNOSPERMAE [GYMNOSPERMS]
18. Ephedra regeliana Florin Figure 10
Shrub. Gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Figure 10 Ephedra regeliana (photography by Natalya Beshko)
43
44
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
19. Ephedra strobilacea Bunge Figure 11
Shrub. Sandy and clay desert, relic mountains. Plain, foothills. Nuratau Relic Mountains, Kyzylkum.
Fodder, afforestation.
Figure 11 Ephedra strobilacea (photography by Natalya Beshko)
Family 9. Cupressaceae
Genus 12. *Platycladus Spach ①
20. *Platycladus orientalis (L.) Franco
Tree. A giant “holy” tree in the valley Madjrum, the Nuratau Nature Reserve, ca. 8 m in diam., ca.
2000 years old. Their status is uncertain because the species was naturalized in many Asian countries
in ancient times, but botanists regarded Platycladus in Uzbekistan as native and Tertiary relic.
Nuratau. Ornamental. IUCN NT.
Genus 13. Juniperus L.
21. Juniperus pseudosabina Fisch. & C. A. Mey. [Juniperus turkestanica Kom.] Figure 12
Tree. Stony and fine-earth slopes. Montane and alpine zones. North Turkestan. Industrial, essential
a
An asterisk means the alien taxon.
GYMNOSPERMAE [GYMNOSPERMS]
oil, medicinal, afforestation.
Figure 12 Juniperus pseudosabina (photography by Natalya Beshko)
22. Juniperus semiglobosa Regel Figure 13
Tree. Stony and fine-earth slopes, screes. Montane and alpine zones. Malguzar, North Turkestan.
Industrial, essential oil, medicinal, afforestation.
Figure 13 Juniperus semiglobosa (photography by Natalya Beshko)
45
46
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
23. Juniperus polycarpos var. seravschanica (Kom.) Kitam. [Juniperus seravschanica Kom.] Figure 14
Tree. Stony and fine-earth slopes, rocks, screes. Montane zone. Nuratau, Malguzar, North Turkestan.
Industrial, essential oil, food (spice-aromatic), medicinal, afforestation.
Figure 14 Juniperus polycarpos var. seravschanica (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
ANGIOSPERMAE
[ANGIOSPERMS]
LILIOPSIDA
[MONOCOTS]
47
48
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 10. Araceae
Genus 14. Lemna L.
24. Lemna gibba L.
Perennial. Standing and slowly flowing water. Plain. Mirzachul, Kyzylkum. Medicinal, fodder, food.
25. Lemna minor L.
Perennial. Standing and slowly flowing water. Plain. Mirzachul, Kyzylkum. Medicinal, fodder, food.
Genus 15. Arum L.
26. Arum korolkowii Regel Figure 15
Perennial. Shady wet places, river valleys, stony slopes, rocks, screes. Montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, North Turkestan. Medicinal, poisonous, ornamental, dye.
Figure 15 Arum korolkowii (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Genus 16. Eminium Schott.
27. Eminium lehmanii (Bunge) O. Kuntze
Perennial. Sandy deserts. Plain. Nuratau Relic Mountains, Kyzylkum. Poisonous, ornamental.
Family 11. Alismataceae
Genus 17. Alisma L.
28. Alisma plantago-aguatica L.
Perennial. Banks of rivers, lakes and canals, shallows. Plain, foothills, montane zone. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, poisonous.
Genus 18. Sagittaria L.
29. Sagittaria trifolia L.
Perennial. Banks of rivers, lakes and canals, shallows. Plain, foothills. Nuratau Relic Mountains,
Malguzar, Mirzachul, Kyzylkum. Medicinal, fodder, ornamental.
Family 12. Butomaceae
Genus 19. Butomus L.
30. Butomus umbellatus L.
Perennial. Banks of rivers, lakes and canals, shallows. Plain, foothills, montane zone. Nuratau Relic
Mountains, Mirzachul, Kyzylkum. Medicinal, fodder, meliferous, ornamental.
Family 13. Hydrocharitaceae
Genus 20. *Vallisneria L.
31. *Vallisneria spiralis L.
Perennial. Standing and slowly flowing water. Plain. Nuratau Relic Mountains, Malguzar, North
49
50
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Turkestan, Mirzachul, Kyzylkum. Fodder (for geese and ducks), ornamental.
Genus 21. Najas L.
32. Najas marina L.
Annual. Standing and slowly flowing water. Plain. Nuratau Relic Mountains, Malguzar, Mirzachul,
Kyzylkum.
33. Najas minor All.
Annual. Standing and slowly flowing water. Plain, foothills. Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum.
Family 14. Juncaginaceae
Genus 22. Triglochin L.
34. Triglochin palustre L.
Perennial. Banks of rivers, lakes and canals, swamp meadows. Plain, foothills, montane zone.
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder, food.
Family 15. Potamogetonaceae
Genus 23. Potamogeton L.
35. Potamogeton crispus L.
Perennial. Standing and slowly flowing water. Plain, foothills. Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Fodder (for geese, ducks and muskrats).
36. Potamogeton filiformis Pers.
Perennial. Standing and slowly flowing water. Plain, foothills, montane zone. Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder (for geese, ducks and
muskrats).
37. Potamogeton lucens L.
Perennial. Standing and slowly flowing water. Plain, foothills, montane and alpine zones. North
Turkestan, Mirzachul, Kyzylkum. Fodder (for geese, ducks and muskrats).
38. Potamogeton natans L.
Perennial. Standing and slowly flowing water. Plain, foothills. Nuratau Relic Mountains, Mirzachul,
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Kyzylkum. Fodder (for geese, ducks and muskrats), medicinal, ornamental.
39. Potamogeton pectinatus L.
Perennial. Standing and slowly flowing water. Plain, foothills. Nuratau Relic Mountains, Mirzachul,
Kyzylkum. Fodder (for geese, ducks and muskrats), medicinal, ornamental.
40. Potamogeton perfoliatus L.
Perennial. Standing and slowly flowing water. Plain, foothills. Nuratau Relic Mountains, Mirzachul,
Kyzylkum. Fodder (for geese, ducks and muskrats), medicinal, ornamental.
41. Potamogeton pusillus L.
Perennial. Standing and slowly flowing water. Plain, foothills. Malguzar, Mirzachul, Kyzylkum.
Fodder (for geese, ducks and muskrats), medicinal, ornamental.
Genus 24. Zannichellia L.
42. Zannichellia pedunculata Rchb.
Perennial. Standing and slowly flowing water, usually saline. Plain. Nuratau Relic Mountains,
Mirzachul, Kyzylkum. Fodder (for geese, ducks and muskrats).
Family 16. Ruppiaceae
Genus 25. Ruppia L.
43. Ruppia maritima L.
Perennial. Standing and slowly flowing water. Plain. Mirzachul, Kyzylkum. Fodder (for geese, ducks
and muskrats).
Family 17. Colchicaceae
Genus 26. Colchicum (Tourn.) L.
44. Colchicum kesselringii Regel Figure 16
Perennial. Wet places, fine-earth, gravelly and stony slopes, places near melting snow. Plain,
foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan. Medicinal, alkaloidbearing, poisonous, ornamental, threatened species. UzbRDB 3.
51
52
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 16 Colchicum kesselringii (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
45. Colchicum luteum Baker Figure 17
Perennial. Fine-earth, gravelly and stony slopes, places near melting snow. Montane and alpine
zones. Malguzar, North Turkestan. Medicinal, alkaloid-bearing, poisonous, ornamental, dye.
Figure 17 Colchicum luteum (photography by Natalya Beshko)
Genus 27. Merendera Ramond.
46. Merendera robusta Bunge Figure 18
Perennial. Sandy and clay deserts, fine-earth slopes. Plain, foothills. Nuratau Relic Mountains,
Kyzylkum. Medicinal, alkaloid-bearing, poisonous, ornamental.
Figure 18 Merendera robusta (photography by N. Yu. Beshko)
53
54
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 18. Liliaceae
Genus 28. Gagea Salisb.
47. Gagea afghanica A. Terracc.
Perennial. Sandy soils. Plain. Nuratau Relic Mountains, Mirzachul, Kyzylkum. Ornamental.
48. Gagea bergii Litv.
Perennial. Sandy deserts. Plain. Kyzylkum. Ornamental.
49. Gagea calyptrifolia Levichev
Perennial. Fine-eart and stony slopes. Montane zone. North Turkestan. Ornamental.
50. Gagea capillifolia Vved.
Perennial. Shady places. Montane and alpine zones. North Turkestan. Ornamental.
51. Gagea capusii A. Terracc. Figure 19
Perennial. Fine-eart and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental.
Figure 19 Gagea capusii (photography by Natalya Beshko)
52. Gagea chomutovae (Pascher) Pascher Figure 20
Perennial. Fine-earth and stony slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Ornamental.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Figure 20 Gagea chomutovae (photography by Natalya Beshko)
53. Gagea divaricata Regel
Perennial. Sandy soils. Plain. Nuratau Relic Mountains, Kyzylkum. Ornamental.
54. Gagea dschungarica Regel
Perennial. Fine-earth and stony slopes, places near melting snow. Montane and alpine zones.
Malguzar, North Turkestan. Ornamental.
55. Gagea filiformis (Ledeb.) Kar. & Kir.
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental.
56. Gagea gageoides (Zucc.) Vved. Figure 21
Perennial. Shady wet places. Montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Ornamental.
55
56
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 21 Gagea gageoides (photography by N. Yu. Beshko)
57. Gagea graminifolia Vved. Figure 22
Perennial. Clay deserts, piedmont plains, fine-eart slopes. Foothills. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Ornamental.
Figure 22 Gagea graminifolia (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
58. Gagea hissarica Lipsky Figure 23
Perennial. Fine-earth and stony slopes, places near melting snow. Montane and alpine zones.
Malguzar, North Turkestan. Ornamental.
Figure 23 Gagea hissarica (photography by N. Yu. Beshko)
59. Gagea kamelinii Levichev Figure 24
Perennial. Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental.
Figure 24 Gagea kamelinii (photography by Natalya Beshko)
60. Gagea ova Stapf.
Perennial. Sandy and clay deserts, piedmont plains, fine-earth and stony slopes. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Ornamental.
57
58
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
61. Gagea liotardii (Sternb.) Schult. & Schult. f. [Gagea emarginata Kar. & Kir] Figure 25
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental.
Figure 25 Gagea liotardii (photography by N. Yu. Beshko)
62. Gagea nabievii Levichev
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental.
63. Gagea olgae Regel Figure 26
Perennial. Clay deserts, piedmont plains, fine-earth slopes. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Ornamental.
Figure 26 Gagea olgae (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
64. Gagea reinhardii Levichev
Perennial. Fine-earth and stony slopes. Fothills, montane zone. Malguzar, North Turkestan.
Ornamental.
65. Gagea reticulata (Pall.) Schult. & Schult. f. [Gegea pseudoreticulata Vved.] Figure 27
Perennial. Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental.
Figure 27 Gagea reticulata (photography by N. Yu. Beshko)
66. Gagea stipitata Merckl. ex Bunge Figure 28
Perennial. Sandy and clay deserts, piedmont plains, fine-earth and stony slopes. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Ornamental.
Figure 28 Gagea stipitata (photography by Natalya Beshko)
59
60
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
67. Gagea stolonifera Popov & Czugaeva
Perennial. Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental.
68. Gagea subtilis Vved. Figure 29
Perennial. Stony slopes, rocks, shady places. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan, Mirzachul. Ornamental.
Figure 29 Gagea subtilis (photography by N. Yu. Beshko)
69. Gagea taschkentica Levichev
Perennial. Clay deserts, piedmont plains, fine-earth and stony slopes. Plain, foothills, montane zone.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Ornamental.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
70. Gagea tenera Pascher
Perennial. Clay deserts, piedmont plains, fine-earth and stony slopes. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Ornamental.
71. Gagea turkestanica Pascher
Perennial. Stony slopes. Foothills. Malguzar, North Turkestan. Ornamental.
72. Gagea vegeta Vved.
Perennial. Fine-earth and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan,
Mirzachul. Ornamental.
73. Gagea vvedenskyi Grossh.
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental.
Genus 29. Tulipa L.
74. Tulipa affinis Botschantz. Figure 30
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Ornamental. Threatened species, UzbRDB 2.
Figure 30 Tulipa affinis (photography by N. Yu. Beshko)
61
62
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
75. Tulipa borszczowii Regel
Perennial. Sandy deserts. Plain. Kyzylkum. Ornamental.
76. Tulipa buhseana Boiss. Figure 31
Perennial. Sandy and clay deserts, piedmont plain, relic mountains. Plain. Nuratau, Nuratau Relic
Mountains, Kyzylkum. Ornamental.
Figure 31 Tulipa buhseana (photography by Natalya Beshko)
77. Тulipa dasystemon Regel
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Malguzar, North
Turkestan. Ornamental. Threatened species, UzbRDB 2.
78. Тulipa dasystemonoides Vved.
Perennial. Fine-earth, gravelly and stony slopes. Alpine zone. North Turkestan. Ornamental.
79. Тulipa korolkowii Regel
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Ornamental. Threatened species, UzbRDB 2.
80. Tulipa lehmanniana Mercklin Figure 32
Perennial. Sandy and clay deserts, relic mountains. Plain, foothills. Nuratau Relic Mountains,
Kyzylkum. Ornamental. Threatened species, UzbRDB 3.
Figure 32 Tulipa lehmanniana (photography by N. Yu. Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
81. Tulipa micheliana Hoog Figure 33
Perennial. Fine-earth, gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Ornamental. Threatened species, UzbRDB 2.
Figure 33 Tulipa micheliana (photography by Natalya Beshko)
63
64
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
82. Тulipa turkestanica Regel
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Ornamental.
Genus 30. Lloydia Salisb.
83. Lloydia serotina (L.) Reichenb
Perennial. Wet stony slopes, rocks, screes, moraines. Alpine zone. North Turkestan. Ornamental.
Genus 31. Fritillaria L.
84. Fritillaria olgae Vved. Figure 34
Perennial. Fine-earth, gravelly and stony slopes, wet shady places. Montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Ornamental, medicinal.
Figure 34 Fritillaria olgae (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Family 19. Orchidaceae
Genus 32. Epipactis Adans.
85. Epipactis latifolia (L.) All.
Perennial. Swamp meadows, river valleys. Montane and alpine zones. Nuratau, North Turkestan.
Ornamental.
Genus 33. Orchis L.
86. Orchis pseudolaxiflora Czerniak.
Perennial. Swamp meadows, river valleys. Montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental, medicinal.
Genus 34. Dactylorhiza Nevski
87. Dactylorhiza umbrosa (Kar. & Kir.) Nevski Figure 35
Perennial. Swamp meadows, river valleys. Foothills, montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Ornamental, medicinal.
Figure 35 Dactylorhiza umbrosa (photography by Natalya Beshko)
65
66
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 20. Ixioliriaceae
Genus 35. Ixiolirion Herb.
88. Ixiolirion tataricum (Pall.) Schult. & Schult. f. Figure 36
Perennial. Fine-earth, gravelly and stony slopes, river vallys, piedmont plains, sandy and clay
deserts. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Ornamental, medicinal.
Figure 36 Ixiolirion tataricum (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Family 21. Iridaceae
Genus 36. Crocus L.
89. Crocus korolkowii Maw & Regel Figure 37
Perennial. Fine-earth and gravelly slopes, places near melting snow. Foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan. Ornamental, medicinal, food (spice).
Figure 37 Crocus korolkowii (photography by Natalya Beshko)
Genus 37. Iris L.
90. Iris linifoliiformis (Khalk.) Tojibaev & Turginov
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan.
Ornamental.
91. Iris loczyi Kanitz
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. North Turkestan.
Ornamental.
92. Iris longiscapa Ledeb. Figure 38
Perennial. Sandy and clay deserts. Plain. Nuratau Relic Mountains, Kyzylkum. Ornamental.
Figure 38 Iris longiscapa (photography by Natalya Beshko)
67
68
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
93. Iris maracandica (Vved.) Wendelbo Figure 39
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Ornamental.
Figure 39 Iris maracandica (photography by Natalya Beshko)
94. Iris narbutii O. Fedtsch. Figure 40
Perennial. Fine-earth and gravelly slopes, clay deserts, piedmont plains. Plain, foothills. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental.
Figure 40 Iris narbutii (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
95. Iris parvula (Vved.) T. Hall & Seisums
Perennial. Fine-earth, gravelly and stony slopes. Alpine zone. North Turkestan. Ornamental.
96. Iris songarica Schrenk Figure 41
Perennial. Sandy and clay deserts, piedmont plains. Plain, foothills. Nuratau, Aktau, Nuratau Relic
Mountains, Kyzylkum. Ornamental, poisonous.
Figure 41 Iris songarica (photography by Natalya Beshko)
97. Iris tadshikorum Vved.
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental.
Genus 38. Gladiolus L.
98. Gladiolus italicus Mill.
Perennial. Fine-earth slopes, meadows, fields. Foothills, montane zone. North Turkestan. Ornamental.
Threatened species, UzbRDB 1.
69
70
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 22. Asphodelaceae
Genus 39. Eremurus M. Bieb.
99. Eremurus anisopterus (Kar. & Kir.) Regel
Perennial. Sandy deserts. Plain. Kyzylkum. Ornamental.
100. Eremurus chloranthus Popov
Perennial. Alpine zone. North Turkestan. Ornamental. Threatened species, UzbRDB 0. Endemic to
the Turkestan ridge.
101. Eremurus fuscus (O. Fedtsch.) Vved.
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Ornamental, meliferous.
102. Eremurus inderiensis (Stev.) Regel
Perennial. Sandy deserts. Plain. Kyzylkum. Ornamental, meliferous.
103. Eremurus kaufmannii Regel Figure 42
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Malguzar, North
Turkestan. Ornamental.
Figure 42 Eremurus kaufmannii (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
104. Eremurus lactiflorus O. Fedtsch. Figure 43
Perennial. Gravelly and stony slopes, rocks, screes. Montane zone. Nuratau. Ornamental. Threatened
species, UzbRDB 2.
Figure 43 Eremurus lactiflorus (photography by N. Yu. Beshko)
105. Eremurus nuratavicus A. P. Khohkr. Figure 44
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau. Ornamental. Threatened
species, UzbRDB 2. Endemic to the Nuratau ridge.
Figure 44 Eremurus nuratavicus (photography by N. Yu. Beshko)
71
72
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
106. Eremurus olgae Regel Figure 45
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Malguzar,
North Turkestan. Ornamental, food, medicinal, dye, meliferous.
Figure 45 Eremurus olgae (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
107. Eremurus regelii Vved. Figure 46
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental, food, medicinal, dye, melliferous.
Figure 46 Eremurus regelii (photography by Natalya Beshko)
73
74
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
108. Eremurus robustus (Regel) Regel Figure 47
Perennial. Fine-earth, gravelly and stony slopes, rocks. Montane zone. Nuratau, Malguzar, North
Turkestan. Ornamental, medicinal, melliferous. Threatened species, UzbRDB 3.
Figure 47 Eremurus robustus (photography by Natalya Beshko)
109. Eremurus sogdianus (Regel) Benth. & Hook. f. Figure 48
Perennial. Gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Malguzar, North
Turkestan. Ornamental.
Figure 48 Eremurus sogdianus (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
110. Eremurus tianschanicus Pazij & Vved. Figure 49
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Ornamental,
meliferous.
Figure 49 Eremurus tianschanicus (photography by Natalya Beshko)
111. Eremurus turkestanicus Vved. Figure 50
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental, meliferous.
Figure 50 Eremurus turkestanicus (photography by Natalya Beshko)
75
76
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 23. Amaryllidaceae
Genus 40. Ungernia Bunge
112. Ungernia oligostroma Popov & Vved. Figure 51
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental, medicinal.
Figure 51 Ungernia oligostroma (photography by Natalya Beshko)
Genus 41. Allium L.
113. Allium alexeianum Regel Figure 52
Perennial. Stony and gravelly slopes, screes. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental.
Figure 52 Allium alexeianum (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
114. Allium altissimum Regel Figure 53
Perennial. River valleys. Montane zone. Nuratau, Malguzar, North Turkestan. Ornamental, food.
Figure 53 Allium altissimum (photography by Natalya Beshko)
115. Allium barsczewskii Lipsky
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Ornamental.
116. Allium caesium Schrenk Figure 54
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Foothills, montane zone. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental, food.
Figure 54 Allium caesium (photography by Natalya Beshko)
77
78
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
117. Allium carolinianum DC. Figure 55
Perennial. Fine-earth, gravelly and stony slopes. Alpine zone. North Turkestan. Ornamental.
Figure 55 Allium carolinianum (photography by Natalya Beshko)
118. Allium caspium (Pall.) M. Bieb. Figure 56
Perennial. Sandy deserts. Plain. Nuratau Relic Mountains, Kyzylkum.
Figure 56 Allium caspium (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
119. Allium clausum Vved. Figure 57
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Ornamental.
Figure 57 Allium clausum (photography by Natalya Beshko)
79
80
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
120. Allium cupuliferum Regel subsp. cupuliferum Figure 58
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar. Ornamental.
Figure 58 Allium cupuliferum subsp. cupuliferum (photography by Natalya Beshko)
121. Allium cupuliferum subsp. nuratavicum R. M. Fritsch & Beshko Figure 59
Perennial. Stony slopes. Montane zone. Nuratau. Ornamental.
Figure 59 Allium cupuliferum subsp. nuratavicum (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
122. Allium drepanophyllum Vved.
Perennial. Stony and gravelly slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains. Ornamental.
123. Allium filidens Regel Figure 60
Perennial. Piedmont plains, fine-earth, gravelly and stony slopes, rocks. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental.
Figure 60 Allium filidens (photography by Natalya Beshko)
124. Allium griffithianum Boiss. Figure 61
Perennial. Fine-earth slopes, piedmont plains. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental.
Figure 61 Allium griffithianum (photography by Natalya Beshko)
81
82
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
125. Allium gusaricum Regel Figure 62
Perennial. Stony slopes, screes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental.
Figure 62 Allium gusaricum (photography by Natalya Beshko)
126. Allium inconspicuum Vved.
Perennial. Stony and gravelly slopes. Foothills. Nuratau, Nuratau Relic Mountains. Ornamental.
127. Allium isakulii R. M. Fritsch & F. O. Khass. Figure 63
Perennial. Stony slopes. Foothills, montane zone. Nuratau, Malguzar. Ornamental. UzbRDB 2.
Figure 63 Allium isakulii (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
128. Allium jodanthum Vved. Figure 64
Perennial. Fine-earth and gravelly slopes. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan. Ornamental.
Figure 64 Allium jodanthum (photography by Natalya Beshko)
129. Allium karataviense Regel Figure 65
Perennial. Stony and gravelly slopes, screes. Montane zone. Malguzar, North Turkestan. Ornamental.
Figure 65 Allium karataviense (photography by Natalya Beshko)
83
84
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
130. Allium kaufmannii Regel Figure 66
Perennial. Swamp meadows, wet places. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental.
Figure 66 Allium kaufmannii (photography by Natalya Beshko)
131. Allium kokanicum Regel Figure 67
Perennial. Stony slopes. Alpine zone. North Turkestan. Ornamental.
Figure 67 Allium kokanicum (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
132. Allium komarowii Lipsky Figure 68
Perennial. Screes. Montane zone. Malguzar, North Turkestan. Ornamental.
Figure 68 Allium komarowii (photography by Natalya Beshko)
133. Allium levichevii F. O. Khass., Esankulov & N. Sulejm
Perennial. Stony slopes. Montane zone. Malguzar. Ornamental.Endemic to the Malguzar ridge.
134. Allium longicuspis Regel
Perennial. Ravines, river valleys. Montane zone. Nuratau. Ornamental, food, medicinal.
135. Allium oreodictyum Vved.
Perennial. Stony slopes. Montane zone. Nuratau, North Turkestan. Ornamental.
136. Allium oreophiloides Regel
Perennial. Stony slopes, rocks. Alpine zone. North Turkestan. Ornamental.
137. Allium oreophilum C. A. Mey. Figure 69
Perennial. Stony slopes. Alpine zone. North Turkestan. Ornamental.
Figure 69 Allium oreophilum (photography by Natalya Beshko)
85
86
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
138. Allium oschaninii O. Fedtsch.
Perennial. Rocks, stony slopes. Montane zone. Malguzar, North Turkestan. Ornamental, food.
139. Allium praemixtum Vved. Figure 70
Perennial. Rocks, stony slopes, screes. Montane zone. Nuratau, Malguzar. Ornamental, food.
Threatened species, UzbRDB 1.
Figure 70 Allium praemixtum (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
140. Allium protensum Wendelbo
Perennial. Stony and gravelly slopes. Foothills. Nuratau Relic Mountains. Ornamental.
141. Allium sarawschanicum Regel Figure 71
Perennial. River valleys, fine earth slopes. Montane zone. Malguzar, North Turkestan. Ornamental.
Figure 71 Allium sarawschanicum (photography by Natalya Beshko)
142. Allium stephanophorum Vved.
Perennial. Fine-earth and gravelly slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental.
143. Allium stipitatum Regel Figure 72
Perennial. Fine-earth slopes, wet places, river valleys. Montane zone. Nuratau, Aktau, Malguzar,
North Turkestan. Ornamental, food.
Figure 72 Allium stipitatum (photography by Natalya Beshko)
87
88
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
144. Allium suworowii Regel Figure 73
Perennial. Fine-earth slopes, river valleys, burials, gardens. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Ornamental, food.
Figure 73 Allium suworowii (photography by Natalya Beshko)
145. Allium taeniopetalum Popov & Vved. Figure 74
Perennial. Fine-earth and stony slopes, shady places. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental.
Figure 74 Allium taeniopetalum (photography by Natalya Beshko)
146. Allium talassicum Regel
Perennial. Stony slopes. Montane zone. North Turkestan. Ornamental.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
147. Allium turkestanicum Regel Figure 75
Perennial. Stony slopes, rocks. Foothills, montane zone. Nuratau, Malguzar. Ornamental.
Figure 75 Allium turkestanicum (photography by Natalya Beshko)
148. Allium verticillatum Regel Figure 76
Perennial. Piedmont plains, fine-earth and gravelly slopes. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental.
Figure 76 Allium verticillatum (photography by Natalya Beshko)
89
90
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
149. Allium xiphopetalum Aitch. & Baker
Perennial. Stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Ornamental.
Family 24. Asparagaceae
Genus 42. Polygonatum Adans.
150. Polygonatum sewerzowii Regel Figure 77
Perennial. Fine-earth and stony slopes, shady places. Montane zone. Malguzar, North Turkestan.
Medicinal, food, poisonous, ornamental.
Figure 77 Polygonatum sewerzowii (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Genus 43. Asparagus L.
151. Asparagus brachyphyllus Turcz. Figure 78
Perennial. Saline lands and depressions, sand deserts, alkaline meadows, lake shores. Plain.
Kyzylkum. Medicinal, fodder.
Figure 78 Asparagus brachyphyllus (photography by Natalya Beshko)
152. Asparagus persicus Baker Figure 79
Perennial. River valleys, rocks, fine-earth, gravelly and stony slopes. Foothills, montane zone.
Malguzar, North Turkestan.
Figure 79 Asparagus persicus (photography by Natalya Beshko)
91
92
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 44. Fessia Speta
153. Fessia puschkinioides (Regel) Speta
Perennial. Fine-earth and stony slopes, places near melting snow. Montane and alpine zones.
Malguzar, North Turkestan. Ornamental.
Family 25. Typhaceae
Genus 45. Sparganium L.
154. Sparganium stoloniferum (Buch.-Ham. ex Graebn.) Buch.-Ham. ex Juz.
Perennial. Banks of rivers, lakes and canals. Plain. Mirzachul, Kyzylkum.
Genus 46. Typha L.
155. Typha angustata Bory & Chaub
Perennial. Swamps, banks of rivers, lakes and canals. Plain. Mirzachul, Kyzylkum.
156. Typha angustifolia L. Figure 80
Perennial. Swamps, banks of rivers, lakes and canals. Plain, foothills. Malguzar, North Turkestan,
Mirzachul, Kyzylkum.
Figure 80 Typha angustifolia (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
157. Typha latifolia L.
Perennial. Swamps, banks of rivers, lakes and canals. Plain, foothills, montane zone. Malguzar,
North Turkestan, Mirzachul, Kyzylkum.
158. Typha laxmannii Lepech.
Perennial. Swamps, banks of rivers, lakes and canals. Plain. Mirzachul.
159. Typha minima Funck
Perennial. Swamps, banks of rivers, lakes and canals. Plain. Mirzachul.
Family 26.*Eriocaulaceae
Genus 47.*Eriocaulon L.
160.*Eriocaulon sieboldianum Sieb. ex Zucc.
Annual. Swamps, banks of rivers, lakes and canals, pebbles, rice fields. Plain, foothills, montane and
alpine zones. Nuratau, North Turkestan, Mirzachul. Medicinal, weed.
Family 27. Juncaceae
Genus 48. Juncus L.
161. Juncus articulatus L.
Perennial. Swamps, banks of rivers, lakes and canals, pebbles. Plain, foothills, montane and alpine
zones. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Fodder.
162. Juncus bufonius L.
Annual. Swamp meadows, banks of rivers, lakes and canals, pebbles, saline lands. Plain, foothills,
montane and alpine zones. Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder, medicinal, weed.
163. Juncus compressus Jacq.
Perennial. Banks of rivers and lakes, dry riverbeds, swamp meadows, fields, saline lands. Plain,
foothills, montane and alpine zones. North Turkestan. Fodder, medicinal, industrial.
164. Juncus gerardii Loisel
Perennial. Swamp meadows, banks of rivers, lakes and canals, wet slopes, fallow lands. Plain,
foothills, montane zone. Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder, medicinal,
industrial.
165. Juncus heptopotamicus V. I. Krecz. & Gontsch.
Perennial. Swamp meadows, banks of rivers, wet stony slopes, saline lands. Plain, foothills, montane
and alpine zones. North Turkestan. Fodder.
93
94
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
166. Juncus hybridus Brot.
Annual. Banks of rivers and lakes, dry riverbeds, swamp meadows, fallow lands. Plain, foothills,
montane and alpine zones. North Turkestan, Mirzachul, Kyzylkum. Fodder.
167. Juncus inflexus L.
Perennial. Swamp meadows, banks of rivers, lakes and canals, wet slopes. Plain, foothills, montane
and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Fodder.
168. Juncus jaxarticus V. I. Krecz. & Gontsch.
Perennial. Swamp meadows, banks of rivers, lakes and canals, saline lands, depressions, fields. Plain,
foothills, montane zone. Malguzar, Mirzachul, Kyzylkum. Fodder.
169. Juncus macrantherus V. I. Krecz. & Gontsch.
Perennial. Swamps, banks of rivers, wet slopes. Montane and alpine zones. North Turkestan. Fodder.
170. Juncus persicus subsp. libanoticus (J.Thiébaut) Novikov & Snogerup [Juncus vvedenskyi
V. I. Krecz.]
Perennial. Swamp meadows, banks of rivers and lakes, wet slopes. Plain, foothills, montane zone.
Nuratau. Fodder.
171. Juncus rechingeri Snogerup.
Annual. Swamp meadows, banks of rivers and lakes. Plain, foothills, alpine zone. Nuratau. Fodder.
172. Juncus sphaerocarpus Nees.
Annual. Swamp meadows, banks of rivers and lakes. Plain, foothills, montane and alpine zones.
Nuratau. Fodder.
173. Juncus triglumis L.[Juncus schischkinii Krylov & Sumnev.]
Perennial. Swamp meadows, banks of rivers and lakes. Montane and alpine zones. Malguzar, North
Turkestan. Fodder.
174. Juncus turkestanicus V. I. Krecz. & Gontsch.
Annual. Swamp meadows, banks of rivers and lakes. Plain, foothills, montane zone. Nuratau, North
Turkestan. Fodder.
Genus 49. Luzula DC.
175. Luzula pallescens Sw.
Perennial. Riwer banks, swamps, alpine meadows. Montane and alpine zones. North Turkestan.
Fodder.
176. Luzula spicata (L.) DC.
Perennial. Moraines, rocks, wet stony slopes, swamps, banks of rivers. Alpine zone. North Turkestan.
Fodder.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Family 28. Cyperaceae
Genus 50. Schoenoplectus (Rchb.) Palla
177. Schoenoplectus litoralis (Schrad.) Palla [Scirpus litoralis Schrad.]
Perennial. Banks of rivers, lakes and canals. Plain. Nuratau Relic Mountains, Mirzachul, Kyzylkum.
178. Schoenoplectus triqueter (L.) Palla [Scirpus triqueter L.]
Perennial. Banks of rivers, lakes and canals. Plain. Mirzachul.
Genus 51. Scirpus L.
179. Scirpus triquetriformis (V. Krecz.) T. V. Egorova
Perennial. Banks of rivers, lakes and canals, near springs. Plain, foothills, montane zone. North
Turkestan.
Genus 52. Bolboschoenus (Aschers.) Palla
180. Bolboschoenus maritimus (L.) Palla Figure 81
Perennial. Banks of rivers, lakes and canals, near springs, swamp meadows, saline depressions. Plain,
foothills. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Medicinal, fodder, industrial, weed.
Figure 81 Bolboschoenus maritimus (photography by Natalya Beshko)
95
96
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
181. Bolboschoenus popovii T. V. Egorova Figure 82
Perennial. Banks of rivers, lakes and canals, saline lands. Plain. Kyzylkum. Medicinal, fodder,
industrial, weed.
Figure 82 Bolboschoenus popovii (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Genus 53. Holoschoenus Link.
182. Holoschoenus vulgaris Link. Figure 83
Perennial. Banks of rivers, lakes and canals, pebbles, dry riverbeds, wet depressions, saline lands.
Plain, foothills, montane zone. Nuratau, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Figure 83 Holoschoenus vulgaris (photography by Natalya Beshko)
Genus 54. Blysmus Panz. ex Schult.
183. Blysmus compressus (L.) Panz. ex Link.
Perennial. Swamp meadows, banks of rivers, lakes and canals. Foothills, montane and alpine zones.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Genus 55. Baeothryon A. Dietr.
184. Baeothryon pumilum (Vahl) Á. & D. Löve
Perennial. Banks of rivers, swamps, alpine meadows, wet places, places near melting snow. Foothills,
montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Genus 56. Eleocharis R. Br.
185. Eleocharis argyrolepis Kier.
Perennial. Swamp meadows, banks of rivers, lakes and canals. Plain. Mirzachul. Weed.
97
98
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
186. Eleocharis quinqueflora (Hartmann) O. Schwarz. [Eleocharis meridionalis Zinserl.]
Perennial. Swamp meadows, banks of rivers, lakes and canals, saline depressions. Plain, foothills,
montane and alpine zones. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul.
187. Eleocharis mitracarpa Steud.
Perennial. Banks of rivers, lakes and canals, swamps, wet meadows. Plain, foothills, montane zone.
Malguzar, North Turkestan, Mirzachul.
188. Eleocharis uniglumis (Link.) Schult.
Perennial. Swamp meadows, saline lands, banks of rivers, lakes and canals. Plain, foothills, montane
zone. Nuratau, Malguzar.
Genus 57. Fimbristylis Vahl.
189. Fimbristylis dichotoma (L.) Vahl
Annual. Wet places, banks of rivers, lakes and canals, pebbles. Plain. Mirzachul. Weed.
Genus 58. Cyperus L.
190. Cyperus glaber L.
Annual. Banks of rivers, lakes and canals, pebbles, swamp meadows, wet places near springs. Plain,
foothills, montane zone. Nuratau. Fodder, weed.
191. Cyperus longus L. Figure 84
Perennial. Banks of rivers, lakes and canals, pebbles, swamp meadows, wet places near springs.
Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Medicinal, food (spice-aromatic), fodder, ornamental.
Figure 84 Cyperus longus (photography by Natalya Beshko)
192. Cyperus rotundus L.
Banks of rivers, lakes and canals, pebbles, swamp meadows, wet places near springs, fallow lands.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Medicinal, fodder, weed, ornamental.
Genus 59. Pycreus P. Beauv.
193. Pycreus globosus (All.) Reichenb
Annual. Banks of rivers and canals, pebbles, swamps, wet places near springs, pebbles, alkaline
meadows. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains. Fodder, weed.
Genus 60. Schoenus L.
194. Schoenus nigricans L. Figure 85
Perennial. Swamp meadows, banks of rivers, wet places near springs. Montane zone. Nuratau.
Figure 85 Schoenus nigricans (photography by Natalya Beshko)
Genus 61. Kobresia Willd.
195. Kobresia pamiroalaica Ivanova
Perennial. Wet slopes, banks of rivers, swamps, highlands. Montane and alpine zones. North
Turkestan. Fodder.
196. Kobresia persica Kük. & Bornm.
Perennial. Banks of rivers, swamps, highlands, wet stony slopes, screes. Montane and alpine zones.
North Turkestan. Fodder.
197. Kobresia stenocarpa (Kar. & Kir.) Steud.
Perennial. Swamps, banks of rivers, wet places near springs. Montane and alpine zones. North
Turkestan. Fodder.
99
100
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 62. Carex L.
198. Carex diluta M. Bieb.
Perennial. Swamp meadows, banks of rivers, wet places near springs. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder.
199. Carex divisa Huds.
Perennial. Swamp meadows, banks of rivers, lakes and canals, wet places near springs, depressions.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Fodder.
200. Carex enervis C. A. Mey.
Perennial. Swamp meadows, banks of rivers, wet places near springs. Montane and alpine zones.
North Turkestan. Fodder.
201. Carex karoi (Freyn) Freyn
Perennial. Swamp meadows, banks of rivers, wet places near springs. Montane and alpine zones.
North Turkestan. Fodder.
202. Carex melanantha C. A. Mey. Figure 86
Perennial. Swamp meadows, banks of rivers, wet places near springs. Alpine zone. North Turkestan.
Fodder.
Figure 86 Carex melanantha (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
203. Carex melanostachya М. Bieb. ex Willd.
Perennial. Swamp meadows, fine-earth slopes, banks of rivers, wet places near springs. Plain,
foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Fodder.
204. Carex microglochin Wahlenb
Perennial. Swamp meadows, banks of rivers and canals, wet places near springs. Montane and alpine
zones. North Turkestan. Fodder.
205. Carex orbicularis Boott.
Perennial. Swamp meadows, banks of rivers, lakes and canals, pebbles, wet places near springs.
Plain, foothills, montane and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan. Fodder.
206. Carex pachystilis J. Gay.
Perennial. Clay deserts, piedmont plain, fine-earth, gravelly and stony slopes, fallow lands. Plain,
foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Fodder.
207. Carex parva Nees.
Perennial. Swamps, banks of rivers, wet places near springs. Montane and alpine zones. North
Turkestan. Fodder.
208. Carex physodes M. Bieb. Figure 87
Perennial. Sandy deserts. Plain. Kyzylkum. Fodder.
Figure 87 Carex physodes (photography by Natalya Beshko)
209. Carex pseudofoetida Kük.
Perennial. Swamps, banks of rivers, wet places near springs, alpine meadows. Montane and alpine
zones. North Turkestan. Fodder.
210. Carex serotina Mérat.
Perennial. Swamp meadows, banks of rivers, wet places near springs. Foothills, montane zone.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder.
211. Carex songorica Kar. & Kir.
Perennial. Swamp meadows, river valleys, ravines, pebbles, banks of canals. Plain, foothills,
montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Fodder.
101
102
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
212. Carex stenophylla Wahlenb
Perennial. Piedmont plains, fine-earth slopes, meadows, river valleys, dry riverbeds, banks of canals.
Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan. Fodder.
213. Carex subphysodes Popov ex V. I. Krecz.
Perennial. Sandy deserts. Plain. Kyzylkum. Fodder.
214. Carex stenocarpa Turcz. & Besser.
Perennial. Alpine and subalpine meadows, fine-earth slopes, rocks. Montane and alpine zones,
Malguzar, North Turkestan. Fodder.
215. Carex turkestanica Regel Figure 88
Perennial. Fine-earth and stony slopes, river valleys, rocks, screes. Montane zone. Malguzar, North
Turkestan. Fodder.
Figure 88 Carex turkestanica (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Family 29. Poaceae [Gramineae]
Genus 63. Imperata Cyr.
216. Imperata cylindrica (L.) Raeusch.
Perennial. River valleys, banks of lakes and canals, fallow lands, fine-earth slopes. Plain, foothills,
montane zone. North Turkestan. Fodder, industrial, ornamental.
Genus 64. Saccharum L.
217. Saccharum spontaneum L.
Perennial. Banks of rivers, lakes and canals, pebbles. Plain. Mirzachul, Kyzylkum. Fodder, food,
industrial.
Genus 65. Erianthus Rich.
218. Erianthus ravennae (L.) P. Beauv. Figure 89
Perennial. River valleys, banks of lakes and canals, wet places. Plain, foothills, montane zone.
Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder, industrial, ornamental.
Figure 89 Erianthus ravennae (photography by Natalya Beshko)
103
104
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 66. Botriochloa Kuntze
219. Botriochloa ischaemum (L.) H. Keng
Perennial. Fine-earth and stony slopes, banks of canals, roadsides. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder, medicinal,
weed.
Genus 67. *Sorghum Pers.
220. *Sorghum halepense (L.) Pers.
Perennial. Fields, fallow lands, gardens, river valleys, banks of canals. Plain, foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Fodder, food, medicinal, industrial, weed.
Genus 68. Digitaria Heist.
221. Digitaria ischaemum (Schreb.) Muehl.
Annual. Banks of rivers and canals, fields, fallow lands, gardens, wastelands. Plain, foothills,
montane zone. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Fodder, weeds.
222. Digitaria sanguinalis (L.) Scop.
Annual. Banks of rivers and canals, fields, gardens, wastelands. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder, weed.
Genus 69. *Echinochloa P. Beauv.
223. *Echinochloa crus-galli (L.) P. Beauv.
Annual. Fields, fallow lands, gardens, wastelands, banks of rivers and canals, pebbles. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Fodder, food, medicinal, weed.
Genus 70. Setaria P. Beauv.
224. Setaria lutescens (Weig.) F. T. Hubb.
Annual. Fields, fallow lands, gardens, wastelands, banks of canals. Plain, foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Fodder, weed.
225. *Setaria verticillata (L.) P. Beauv.
Annual. Fields, fallow lands, gardens, wastelands, banks of canals. Plain, foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Fodder, weed.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
226. Setaria viridis (L.) P. Beauv.
Annual. Fields, fallow lands, gardens, wastelands, banks of canals, stony slopes. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Fodder, weed.
Genus 71. Phalaroides Wolf
227. Phalaroides arundinacea (L.) Rauschert
Perennial. Banks of rivers, lakes and canals, swamp meadows, wet places. Plain, foothills, montane
zone. Nuratau, Malguzar, North Turkestan. Fodder, ornamental, weed.
Genus 72. Stipagrostis Nees
228. Stipagrostis karelinii (Trin. & Rupr.) H. Scholz [Aristida karelinii (Trin. & Rupr.)
Roshev.] Figure 90
Perennial. Sandy deserts. Plain. Mirzachul, Kyzylkum. Fodder.
Figure 90 Stipagrostis karelinii (photography by Natalya Beshko)
105
106
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
229. Stipagrostis pennata (Trin.) De Winter [Aristida pennata Trin.]
Perennial. Sandy deserts. Plain. Kyzylkum. Fodder.
Genus 73. Stipa L.
230. Stipa arabica Trin. & Rupr. [Stipa szovitsiana Trin.]
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Fodder.
231. Stipa caragana Trin. [Achnatherum caragana (Trin.) Roshev.]
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Fodder, industrial.
232. Stipa capillata L. Figure 91
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Fodder, ornamental.
Figure 91 Stipa capillata (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
233. Stipa caucasica Schmalh.
Perennial. Stony slopes, screes. Foothills, montane and alpine zones. Nuratau, Malguzar, North
Turkestan. Fodder.
234. Stipa hohenackeriana Trin. & Rupr.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Fodder.
235. Stipa kirghisorum P. A. Smirn.
Perennial. Fine-earth, gravelly and stony slopes, pebbles, rocks, screes. Montane and alpine zones.
North Turkestan. Fodder.
236. Stipa kurdistanica Bor. [Achnatherum turkomanicum (Roshev.) Tzvelev]
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Fodder, industrial.
237. Stipa lessingiana Trin. & Rupr.
Perennial. Fine-earth, gravelly and stony slopes, screes. Montane zone. North Turkestan. Fodder,
ornamental.
238. Stipa lingua Junge
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. North Turkestan. Fodder.
239. Stipa lipskyi Roshev.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Fodder.
240. Stipa margelanica P. A. Smirn.
Perennial. Stony slopes. Foothills, montane zone. North Turkestan. Fodder.
241. Stipa orientalis Trin. ex Ledeb.
Perennial. Stony and gravelly slopes, screes, rocks. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Fodder.
242. Stipa richteriana Kar. & Kir.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Fodder.
243. Stipa sareptana Beck.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Fodder.
244. Stipa splendens Trin. [Achnatherum splendens (Trin.) Nevski]
Perennial. Saline lands, river valleys. Plain, foothills, montane and alpine zones. North Turkestan.
Fodder, industrial.
245. Stipa turkestanica Hack. [Stipa trichoides P. Smirn.]
Perennial. Stony slopes. Montane and alpine zones. North Turkestan. Fodder.
Genus 74. Piptatherum P. Beauv.
246. Piptatherum alpestre (Grig.) Roshev.
Perennial. Stony slopes, screes, moraines, swamp meadows. Alpine zone. North Turkestan. Fodder.
107
108
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
247. Piptatherum ferganense (Litv.) Roshev.
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Fodder.
248. Piptatherum holciforme (M. Bieb.) Roem. & Schult.
Perennial. Stony slopes, screes. Montane zone. North Turkestan. Fodder.
249. Piptatherum laterale (Regel) Nevski
Perennial. Fine-earth, gravelly and stony slopes, screes, rocks. Montane and alpine zones. Malguzar,
North Turkestan. Fodder.
250. Piptatherum latifolium (Roshev.) Nevski
Perennial. Fine-earth and stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
Fodder.
251. Piptatherum pamiralaicum (Grig.) Roshev.
Perennial. Fine-earth, gravelly and stony slopes, moraines. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Fodder.
252. Piptatherum platyanthum Nevski
Perennial. Stony slopes, moraines, pebbles, screes, rocks. Montane and alpine zones. North
Turkestan. Fodder.
253. Piptatherum sogdianum (Grig.) Roshev.
Perennial. Fine-earth, gravelly and stony slopes, screes, rocks. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan.
254. Piptatherum songaricum (Trin. & Rupr.) Roshev.
Perennial. Fine-earth and stony slopes, rocks, screes. Foothills, montane and alpine zones. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder.
255. Piptatherum microcarpum (Pilg.) Tzvelev [Piptatherum vicarium (Grig.) Roshev.]
Perennial. Stony slopes, rocks. Montane zone. North Turkestan. Fodder.
Genus 75. Milium L.
256. Milium vernale M. Bieb.
Annual. Fine-earth and stony slopes, wet places, banks of rivers and canals, gardens. Foothills,
montane zone. Nuratau, North Turkestan. Fodder.
Genus 76. Crypsis Ait.
257. Crypsis schoeoides (L.) Lam.
Annual. River valleys, banks of canals, wastelands, saline depressions. Plain. Nuratau, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Fodder.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Genus 77. Phleum L.
258. Phleum alpinum L. Figure 92
Perennial. Banks of rivers, wet places, alpine meadows. Montane and alpine zone. North Turkestan.
Fodder.
Figure 92 Phleum alpinum (photography by Natalya Beshko)
109
110
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
259. Phleum exaratum Griseb. [Phleum graecum Boiss. & Heldr.]
Annual. Stony slopes. Foothills, montane zone. North Turkestan. Fodder.
260. Phleum paniculatum Huds. Figure 93
Annual. Fine-earth, gravelly and stony slopes, screes, pebbles, fallow lands. Foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder.
Figure 93 Phleum paniculatum (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
261. Phleum phleoides (L.) H. Karst.
Perennial. Fine-earth slopes. Montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan. Fodder.
262. *Phleum pratense L.
Perennial. Meadows, river valleys, banks of canals, swamps. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan. Fodder.
Genus 78. Alopecurus L.
263. Alopecurus arundinaceus Poir.
Perennial. Swamp meadows, banks of rivers, wet screes. Plain, foothills, montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Fodder.
264. Alopecurus himalaicus Hook. f.
Perennial. Fine-earth and gravelly slopes, screes, alpine meadows. Montane and alpine zones. North
Turkestan. Fodder.
265. Alopecurus myosuroides Huds.
Annual. Banks of rivers and canals, meadows, pebbles, fields. Plain, foothills. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder, weed.
266. Alopecurus pratensis L.
Perennial. Swamp meadows, banks of rivers, pebbles. Plain, foothills, montane and alpine zones.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder, ornamental.
Genus 79. Polypogon Desf.
267. Polypogon fugax Nees ex Steud.
Annual, perennial. Saline lands, banks of rivers and canals, fields, swamp meadows. Plain, foothills,
montane zone. Nuratau, Malguzar, North Turkestan. Fodder.
268. Polypogon monspleniensis (L.) Desf.
Annual. Wet places, banks of rivers and canals, fields, swamp meadows. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
269. Polypogon viridis (Gouan) Breistr. [Polypogon semiverticillatus (Forssk.) Hyl.]
Annual, perennial. Banks of rivers and canals, fields, swamp meadows. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder.
Genus 80. Agrostis L.
270. Agrostis gigantea Roth.
Perennial. Wet places, banks of rivers, swamp meadows. Foothills, montane and alpine zones.
111
112
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Malguzar, North Turkestan. Fodder, ornamental.
271. Agrostis hissarica Roshev.
Perennial. River valleys. Alpine zone. Nuratau, Malguzar, North Turkestan. Fodder.
272. Agrostis olympica (Boiss.) Bor. [Pentatherum agrostidiforme (Roshev.) Nevski]
Perennial. Swamps, banks of rivers. Alpine zone. North Turkestan. Fodder.
273. Agrostis stolonifera L.
Perennial. Wet places, banks of rivers, swamp meadows. Foothills, montane and alpine zones.
Nuratau, North Turkestan. Fodder.
Genus 81. Calamagrostis Adans.
274. Calamagrostis anthoxanthoides (Munro) Regel [Calamagrostis laguroides Regel]
Perennial. Stony slopes, screes. Alpine zone. North Turkestan. Fodder.
275. Calamagrostis dubia Bunge
Perennial. River valleys, saline lands, banks of canals, fields. Plain, foothills. Nuratau, Malguzar,
North Turkestan. Fodder.
276. Calamagrostis epigeios (L.) Roth.
Perennial. Banks of rivers and canals, fallow lands, wet fine-earth slopes. Plain, foothills, montane
and alpine zones. Malguzar, North Turkestan. Fodder, medicinal, industrial.
277. Calamagrostis holciformis Jaub. & Spach.
Perennial. Rocks, stony slopes. Alpine zone. North Turkestan. Fodder.
278. Calamagrostis pseudophragmites (Haller f.) Koeler.
Perennial. Banks of rivers, pebbles, meadows, wet stony slopes. Foothills, montane and alpine zones.
Nuratau, North Turkestan. Fodder.
Genus 82. Apera Adans.
279. Apera interrupta (L.) P. Beauv.
Annual. Saline lands, river valleys. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Fodder.
Genus 83. Deschampsia P. Beauv.
280. Deschampsia cespitosa (L.) P. Beauv.
Perennial. River valleys, subalpine meadows. Alpine zone. North Turkestan. Fodder, industrial.
281. Deschampsia koelerioides Regel
Perennial. Alpine meadows, stony slopes. Alpine zone. North Turkestan. Fodder, industrial.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Genus 84. Trisetum Pers.
282. Trisetum spicatum (L.) K. Richt.
Perennial. Fine-earth and gravelly slopes, swamps, subalpine meadows, banks of rivers. Alpine zone.
North Turkestan. Fodder, ornamental.
Genus 85. Avena L.
283. Avena barbata Pott ex Link.
Annual. Fine-earth, gravelly and stony slopes, fields, fallow lands. Foothills. North Turkestan.
Fodder, weed.
284. *Avena fatua L.
Annual. Fine-earth, gravelly and stony slopes, fields, fallow lands. Plain, foothills. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder, food, medicinal, weed.
285. *Avena sterilis subsp. ludoviciana (Durieu) Gillet & Magne [Avena persica Steud.; Avena
trichophylla K. Koch]
Annual. River valleys, fine-earth slopes. Foothills. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Genus 86. Helictotrichon Bess.
286. Helictotrichon asiaticum (Roshev.) Grossh.
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Fodder.
287. Helictotrichon desertorum (Less.) Pilg.
Perennial. Piedmont plain, fine-earth, gravelly and stony slopes. Plain, foothills, montane zone.
Malguzar, North Turkestan. Fodder.
Genus 87. *Cynodon Rich.
288. *Cynodon dactylon (L.) Pers. Figure 94
Perennial. River valleys, fields, gardens, fallow lands, banks of canals, roadsides, wastelands, fineearth slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul. Fodder, medicinal, industrial, weed.
113
114
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 94 Cynodon dactylon (photography by Natalya Beshko)
Genus 88. Enneapogon Desv.
289. Enneapogon persicus Boiss.
Perennial. Fine-earth and stony slopes, dry riverbeds, pebbles, rocks. Foothills, montane zone.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Genus 89. Phragmites Adans.
290. Phragmites australis (Cav.) Trin. & Steud. Figure 95
Perennial. Banks of rivers, lakes and canals, swamps, wet places, saline lands, fields. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Fodder, food, industrial, medicinal, ornamental.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Figure 95 Phragmites australis (photography by Natalya Beshko)
Genus 90. Eragrostis Host.
291. Eragrostis minor Host.
Annual. Fine-earth slopes, pebbles, banks of rivers and canals, fields, fallow lands. Plain, foothills.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder, weed.
292. Eragrostis pilosa (L.) Beauv.
Annual. River valleys, banks of canals, pebbles, fields, fallow lands, gardens, wastelands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Fodder, weed.
Genus 91. Koeleria Pers.
293. Koeleria macrantha (Ledeb.) Schult. [Koeleria cristata (L.) Pers.]
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. North Turkestan. Fodder,
medicinal, ornamental.
115
116
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 92. Cutandia Willk.
294. Cutandia rigescens (Grossh.) Tzvelev
Annual. Wet places, saline lands, banks of rivers, lakes and canals, fields. Plain. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Genus 93. Catabrosa P. Beauv.
295. Catabrosa aquatica (L.) Beauv. [Catabrosa capusii Franch.]
Perennial. Wet places, banks of rivers, lakes and canals, swamps. Montane and alpine zones.
Malguzar, North Turkestan. Fodder.
Genus 94. Melica L.
296. Melica altissima L.
Perennial. Stony slopes. Montane zone. Nuratau, North Turkestan. Poisonous.
297. Melica hohenackeri Boiss.
Perennial. Stony slopes, rocks, screes. Montane zone. Nuratau, Malguzar, North Turkestan.
298. Melica inaequiglumis Boiss. Figure 96
Perennial. Stony slopes, screes, pebbles. Montane zone. Nuratau, Malguzar, North Turkestan.
Figure 96 Melica inaequiglumis (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Genus 95. Aeluropus Trin.
299. Aeluropus intermedius Regel
Perennial. Saline sandy soils, saline lands, depressions. Plain. Nuratau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder.
300. Aeluropus litoralis (Gouan.) Parl.
Perennial. Saline lands, swamp meadows, banks of rivers, lakes and canals. Plain, foothills. Nuratau
Relic Mountains, Mirzachul, Kyzylkum. Fodder.
Genus 96. Dactylis L.
301. Dactylis glomerata L.
Perennial. Meadows, river valleys, fine-earth and stony slopes, gardens, wet places. Plain, foothills,
montane zone. Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder, medicinal, ornamental.
Genus 97. Sclerochloa P. Beauv.
302. Sclerochloa dura (L.) P. Beauv.
Annual. Banks of rivers and canals, roadsides. Plain, foothills, montane zone. Nuratau, Aktau,
Malguzar, North Turkestan. Fodder, weed.
Genus 98. Schismus P. Beauv.
303. Schismus arabicus Nees.
Annual. Sandy and clay deserts, saline lands, piedmont plains, fine-earth and gravelly slopes. Plain,
foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Fodder.
Genus 99. Poa L.
304. Poa alpina L. Figure 97
Perennial. Alpine meadows. Alpine zone. North Turkestan. Fodder.
117
118
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 97 Poa alpina (photography by Natalya Beshko)
305. Poa angustifolia L.
Perennial. Fine-earth and stony slopes, screes, meadows. Montane and alpine zones, Malguzar, North
Turkestan. Fodder.
306. Poa annua L.
Annual or perennial. Banks of rivers and canals, wet places near springs, wastelands, gardens. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Fodder, weed.
307. Poa bactriana Roshev.
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Montane and alpine zones. Malguzar,
North Turkestan. Fodder.
308. Poa bucharica Roshev.
Perennial. Fine-earth, gravelly and stony slopes, meadows. Montane and alpine zones. North
Turkestan. Fodder.
309. Poa bulbosa L. Figure 98
Perennial. This plant has a wide ecological range, and it is found almost everywhere in the Province.
Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Fodder.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Figure 98 Poa bulbosa (photography by Natalya Beshko)
119
120
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
310. Poa diaphora Trin.
Annual. Sandy, clayey, stony and saline soils, banks of rivers and lakes, wet places near springs.
Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Fodder.
311. Poa fragilis Ovcz.
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones, Nuratau, Malguzar,
North Turkestan. Fodder.
312. Poa litvinoviana Ovcz.
Perennial. Stony slopes, rocks. Montane and alpine zones. Nuratau, Malguzar, North Turkestan. Fodder.
313. Poa nemoralis L.
Perennial. River valleys, ravines, meadows, stony slopes, rocks. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Fodder.
314. Poa palustris L.
Perennial. Wet places, river valleys, swamp meadows. Foothills, montane zone. Nuratau, Malguzar,
North Turkestan. Fodder.
315. Poa pratensis L.
Perennial. River valleys, banks of canals, meadows. Plain, foothills, montane and alpine zones.
Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder, ornamental.
316. Poa relaxa Ovcz.
Perennial. Stony, Fine-earth and stony slopes, rocks, screes. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Fodder.
317. Poa supina Schrad.
Annual or perennial. Wet places, swamps, banks of rivers. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Fodder.
318. Poa trivialis L.
Perennial. River valleys, gardens, banks of canals, fallow lands. Plain, foothills, montane zone.
Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder.
Genus 100. Colpodium Trin.
319. Colpodium humile (M. Bieb.) Griseb.
Perennial. Sandy deserts, saline lands, wet places. Plain, foothills. Mirzachul, Kyzylkum. Fodder.
320. Colpodium parviflorum Boiss. & Buhse.
Perennial. Wet places. Plain. Mirzachul. Fodder.
Genus 101. Glyceria R. Br.
321. Glyceria plicata (Fr.) Fr.
Perennial. Banks of rivers, lakes and canals. Plain. Nuratau, Malguzar, North Turkestan. Fodder.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Genus 102. Puccinelia Parl.
322. Puccinelia distans (Jack.) Parl.
Perennial. Banks of rivers, lakes and canals, swamp meadows. Plain, foothills, montane and alpine
zones. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder, weed.
323. Puccinellia poecilantha (K. Koch) Grossh.
Perennial. Swamp meadows, saline lands, banks of rivers, lakes and canals. Plain, foothills. Nuratau,
Malguzar, North Turkestan, Mirzachul. Fodder, weed.
Genus 103. Festuca L.
324. Festuca alaica Drobow.
Perennial. Stony slopes, screes, rocks, moraines. Montane and alpine zones. North Turkestan. Fodder.
325. Festuca amblyodes Krecz. & Bobrov
Perennial. Stony slopes, screes, rocks, moraines. Alpine zone. North Turkestan. Fodder.
326. Festuca arundinacea Schreb.
Perennial. Saline lands, banks of rivers, lakes and canals, gardens, meadows, fields, fallow lands,
cliffs. Plain, foothills, montane zone. Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder.
327. Festuca griffithiana (St.-Yves) Krivot. [Leucopoa karatavica (Bunge) V. Krecz. & Bobrov].
Perennial. Stony slopes, screes, rocks. Montane and alpine zones. North Turkestan. Fodder.
328. Festuca olgae (Regel) Krivot. [Leucopoa olgae (Regel) V. Krecz. & Bobrov]
Perennial. Stony slopes, screes, rocks. Montane and alpine zones. North Turkestan. Fodder.
329. Festuca pratensis Huds.
Perennial. Wet places, gardens, banks of rivers and canals, wet slopes. Plain, foothills, montane zone.
North Turkestan. Fodder.
330. Festuca rubra L.
Perennial. River valleys, pebbles, meadows, swamps, fine-earth and stony slopes. Montane and
alpine zones. Malguzar, North Turkestan. Fodder.
331. Festuca rupicola Heuff.
Perennial. Stony slopes, rocks. Montane and alpine zones. Malguzar, North Turkestan. Fodder.
332. Festuca valesiaca Schleich. & Gaudin. Figure 99
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes, pebbles, moraines. Montane and
alpine zones. Nuratau, Aktau, Malguzar, North Turkestan. Fodder.
121
122
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 99 Festuca valesiaca (photography by Natalya Beshko)
Genus 104. Loliolum V. Krecz. & Bobrov
333. Loliolum subulatum (Banks & Sol.) Eig.
Annual. Piedmont plains, fine-earth, gravelly and stony slopes. Plain, foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan.
Genus 105. Vulpia C. C. Gmel.
334. Vulpia ciliata Dumort.
Annual. Sandy, clayey, gravelly and saline soils, banks of rivers. Plain, foothills. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
335. Vulpia myuros (L.) C. C. Gmel.
Annual. Fine-earth and stony slopes, pebbles, river valleys, saline lands. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder, weed
336. Vulpia persica (Boiss. & Buhse) V. Krecz. & Bobrov
Annual. Clay deserts, fine-earth and gravelly slopes, pebbles, river valleys, saline depressions.
Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Fodder, weed.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
337. Vulpia unilateralis (L). Stace [Nardurus krausei (Regel) V. Krecz. & Bobrov]
Annual. Fine-earth and stony slopes, piedmont plains, sandy and clay deserts, pebbles, river
valleys. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Genus 106. Bromus L.
338. Bromus danthoniae Trin. Figure 100
Annual. Clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, fallow lands, pebbles.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Fodder, weed.
Figure 100 Bromus danthoniae (photography by Natalya Beshko)
123
124
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
339. Bromus inermis Leyss.
Perennial. Meadows, fine-earth and gravelly slopes, river valleys. Montane and alpine zones .
Nuratau, Malguzar, North Turkestan. Fodder.
340. Bromus japonicus Thung.
Annual. Meadows, fine-earth slopes, roadsides, fallow lands, banks of rivers and canals. Foothills,
montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Fodder, weed.
341. Bromus lanceolatus Roth.
Annual. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, fields,
fallow lands, banks of rivers, pebbles. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
342. Bromus oxyodon Schrenk
Annual. Fine-earth, gravelly and stony slopes, fields, fallow lands. Plain, foothills, montane zone.
Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder, weed.
343. Bromus paulsenii Hack.
Perennial. Stony slopes, banks of rivers, screes. Montane and alpine zones. Malguzar, North
Turkestan. Fodder.
344. Bromus popovii Drobow
Annual. Saline lands, river valleys. Plain. Mirzachul. Fodder.
345. Bromus scoparius L.
Annual. Fine-earth, gravelly and stony slopes, fields, fallow lands, river valleys, roadsides, gardens.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan. Fodder, weed.
346. Bromus sericeus Drobow
Annual. Sandy deserts, fine-earth and stony slopes, river valleys. Plain, foothills, montane and alpine
zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder, weed.
347. Bromus sewerzowii Regel
Annual. River valleys, fields, fallow lands, fine-earth and stony slopes. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder, weed.
348. Bromus scoparius L.
Annual. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, meadows,
pebbles, fields, fallow lands, gardens. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
349. Bromus sterilis L.
Annual. River valleys, ravines, fine-earth slopes, meadows, gardens, roadsides, fields. Foothills,
montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Fodder, weed.
350. Bromus tectorum L.
Annual. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, fields, fallow
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
lands, settlements, river valleys. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Genus 107. Boissiera Hochsst & Steud.
351. Boissiera squarrosa (Sol.) Nevski. Figure 101
Annual. Sandy and clay desert, piedmont plains, fine-earth, gravelly and stony slopes, fields,
fallow lands, roadsides. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Figure 101 Boissiera squarrosa (photography by Natalya Beshko)
Genus 108. Brachypodium P. Beauv.
352. Brachypodium sylvaticum P. Beauv.
Perennial. Fine-earth wet slopes, river valleys, ravines. Montane zone. Nuratau, Malguzar, North
Turkestan. Fodder.
125
126
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 109. Lolium L.
353. *Lolium perenne L.
Perennial. Gardens, fields, fallow lands, banks of canals, dry slopes. Plain, foothills, montane zone.
Malguzar, North Turkestan, Mirzachul. Fodder, medicinal.
354. Lolium persicum Boiss. & Hohen.
Annual. Fallow lands, fields, stony slopes, river valleys. Foothills, montane zone. Malguzar, North
Turkestan. Fodder, weed.
355. *Lolium temulentum L.
Annual. Fallow lands, gardens, wet meadows, banks of canals, river valleys, fine-earth slopes.
Foothills, montane zone. Malguzar, North Turkestan. Fodder, medicinal.
Genus 110. Henrardia Hubb.
356. Henrardia persica (Boiss.) C. E. Hubb.
Annual. Dry fine-earth, gravelly and stony slopes. Foothills. Nuratau, Malguzar, North Turkestan.
Fodder, weed.
Genus 111. Psilurus Trin.
357. Psilurus incurvus (Gouan) Schinz & Thell.
Annual. Piedmont plains, fine-earth, gravelly and stony slopes. Plain, foothills. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder.
Genus 112. Agropyron Gaertn.
358. Agropyron badamense Drobow
Perennial. Stony slopes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan. Fodder.
359. Agropyron setuliferum (Nevski) Nevski
Perennial. Fine-earth and stony slopes. Montane zone. Malguzar, North Turkestan. Fodder.
Genus 113. Eremopyrum Jaub. & Spach.
360. Eremopyrum bonaepartis (Spreng.) Nevski Figure 102
Annual. Sandy, clayey and gravelly soils, fallow lands. Plain, foothills. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Figure 102 Eremopyrum bonaepartis (photography by Natalya Beshko)
361. Eremopyrum distans (K. Koch) Nevski Figure 103
Annual. Sandy, clayey, gravelly and saline soils, pebbles, fallow lands. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder, weed.
Figure 103 Eremopyrum distans (photography by Natalya Beshko)
127
128
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
362. Eremopyrum orientale (L.) Jaub. & Spach.
Annual. Sandy, clayey, gravelly and saline soils, pebbles, fallow lands. Plain, foothills. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
363. Eremopyrum triticeum (Gaertn.) Nevski
Annual. Sandy, clayey, gravelly and saline soils, river valleys, fallow lands. Plain, foothills. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Genus 114. Secale L.
364. *Secale cereale L.
Annual. Fields, fallow lands, river valleys, banks of canals, roadsides. Foothills, montane zone.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, food.
365. Secale segetale (Zhuk.) Roshev.
Annual. Fields, fallow lands, river valleys, banks of canals, roadsides. Plain, foothills. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, food.
366. Secale sylvestre Host.
Annual. Sandy and clayey soils. Plain, Mirzachul, Kyzylkum. Fodder, food.
Genus 115. Aegilops L.
367. Aegilops crassa Boiss.
Annual. Fine-earth and stony slopes, fields, fallow lands, river valleys. Plain, foothills, montane
zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder,
weed.
368. Aegilops cylindrica Host.
Annual. Fine-earth and stony slopes, fields, fallow lands, river valleys. Plain, foothills. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder, weed.
369. Aegilops juvenalis (Thell.) Eig.
Annual. Fine earth slopes. Foothills. Nuratau, Aktau, Nuratau Relic Mountains. Fodder, weed.
370. Aegilops triuncialis L. [Aegilops squarrosa L.] Figure 104
Annual. Piedmont plains, fine-earth and stony slopes, fields, fallow lands, river valleys. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Fodder, weed.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Figure 104 Aegilops triuncialis (photography by Natalya Beshko)
Genus 116. Heteranthelium Hochst. ex Jaub. & Spach.
371. Heteranthelium piliferum (Sol.) Hochst. ex Jaub. & Spach.
Annual. Stony slopes, piedmont plains. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder.
129
130
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 117. Taeniatherum Nevski
372. Taeniatherum caput-medusae (L.) Nevski [Taeniatherum asperum (Simonk.) Nevski;
Taeniatherum crinitum (Schreb.) Nevski]
Annual. Fallow lands, fields, roadsides, piedmont plain, fine-earth, gravelly and stony slopes. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Fodder, weed.
Genus 118. Hordeum L.
373. Hordeum bogdanii Wilensky
Perennial. Swamp meadows, saline lands, banks of rivers, lakes and canals. Plain, foothills, montane
zone. Mirzachul, Kyzylkum. Fodder.
374. Hordeum brevisubulatum (Trin.) Link.
Perennial. Banks of rivers and lakes, swamp meadows, stony slopes. Plain, foothills, montane and
alpine zones. Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder.
375. Hordeum bulbosum L. Figure 105
Perennial. Fine-earth, gravelly and stony slopes, river valleys, fields, gardens, banks of canals. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Fodder, food.
Figure 105 Hordeum bulbosum (photography by Natalya Beshko)
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
376. Hordeum marinum subsp. gussoneanum (Parl.) Thell. [Hordeum geniculatum All.]
Annual. River valleys. Plain, foothills. Nuratau, North Turkestan, Mirzachul. Fodder, weed.
377. Hordeum murinum subsp. leporinum (Link.) Arcang. [Hordeum leporinum Link.]
Annual. River valleys, gardens, fields, fallow lands, wastelands, settelments, sandy and clay deserts,
fine-earth, gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
378. Hordeum spontaneum K. Koch Figure 106
Annual. Fields, fallow lands, banks of canals, fine-earth, gravelly and stony slopes. Plain, foothills,
montane zone. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Figure 106 Hordeum spontaneum (photography by Natalya Beshko)
Genus 119. Elymus L.
379. Elymus bungeanus (Trin.) Melderis [Agropyron ferganense Drobow]
Perennial. Fine-earth and stony slopes. Montane zone. Malguzar, North Turkestan. Fodder.
380. Elymus dentatus (Hook. f.) Tzvelev [Agropyron lachnophyllum (Ovcz. & Sidorenko)
Bondarenko; Agropyron ugamicum Drobow]
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
131
132
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Fodder.
381. Elymus drobovii (Nevski) Tzvelev [Agropyron drobovii Nevski]
Perennial. Stony slopes. Montane zone. Nuratau. Fodder.
382. Elymus hispidus (Opiz) Melderis [Agropyron intermedium (Host) Beauv.; Agropyron
trichophorum (Link.) K. Richt.] Figure 107
Perennial. Fine-earth, gravelly and stony slopes. The species is dominant of so-called savannoids (dry
herbaceous steppes), a vegetation type endemic to the Central Asian mountains. Foothills, montane
zone. Nuratau, Aktau, Malguzar, North Turkestan. Fodder.
Figure 107 Elymus hispidus (photography by Natalya Beshko)
383. Elymus lolioides (P. Candargy) Melderis [Agropyron lolioides (Kar. & Kir.) Roshev.]
Perennial. Meadows, fine-earth slopes, banks of rivers and canals. Plain, foothills, montane zone.
Nuratau, Malguzar, North Turkestan. Fodder.
384. Elymus repens (L.) Gould. [Agropyron repens (L.) P. Beauv.]
Perennial. Wet slopes, meadows, river valleys, gardens. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Fodder, medicinal, weed.
385. Elymus uralensis (Nevski) Tzvelev [Agropyron tianschanicum Drobow]
Perennial. Stony slopes, screes. Montane zone. Malguzar, North Turkestan. Fodder.
Genus 120. Leymus Hochst.
386. Leymus alaicus (Korsch.) Tzvelev
Perennial. Stony slopes. Montane zone. North Turkestan. Fodder.
387. Leymus angustus (Trin.) Pilg. Figure 108
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau. Fodder.
ANGIOSPERMAE [ANGIOSPERMS]
LILIOPSIDA [MONOCOTS]
Figure 108 Leymus angustus (photography by Natalya Beshko)
388. Leymus multicaulis (Kar. & Kir.) Tzvelev
Perennial. River valleys, wet meadows, banks of canals. Plain, foothills, montane zone. North
Turkestan. Fodder.
133
134
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
MAGNOLIOPSIDA
[EUDICOTS]
MAGNOLIOPSIDA
[EUDICOTS]
135
136
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 30. Ceratophyllaceae
Genus 121. Ceratophyllum L.
389. Ceratophyllum demersum L.
Perennial. Standing and slowly flowing water. Plain, foothills. Nuratau, Mirzachul, Kyzylkum.
Fodder (for fish), ornamental.
Family 31. Papaveraceae
Genus 122. Papaver L.
390. Papaver dubium L. [Papaver litwinowii Fedde ex Bornm.] Figure 109
Annual. Fine-earth, gravelly and stony slopes, rocks, screes, pebbles, dry riverbeds. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Alkaloidbearing.
Figure 109 Papaver dubium (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
391. Papaver nudicaule L. [Papaver croceum Ledeb.] Figure 110
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes, alpine meadows. Montane and alpine
zones. North Turkestan. Ornamental, poisonous, alkaloid-bearing.
Figure 110 Papaver nudicaule (photography by Natalya Beshko)
137
138
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
392. Papaver pavoninum C. A. Mey. Figure 111
Annual. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, fallow lands,
roadsides, wastelands, settlements, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, dye, ornamental,
weed, poisonous, alkaloid-bearing.
Figure 111 Papaver pavoninum (photography by Natalya Beshko)
Genus 123. Roemeria Medik.
393. Roemeria hybrida (L.) DC. Figure 112
Annual. Sandy and clay deserts, saline depressions, pebbles, fine-earth, gravelly and stony slopes.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Weed.
Figure 112 Roemeria hybrida (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
394. Roemeria refracta DC. Figure 113
Annual. Sandy and clay deserts, fine-earth, gravelly and stony slopes, fallow lands, roadsides,
gardens, settlements, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, dye, ornamental, weed,
poisonous, alkaloid-bearing.
Figure 113 Roemeria refracta (photography by Natalya Beshko)
Genus 124. Glaucium Adans.
395. Glaucium elegans Fisch. & C. A. Mey. Figure 114
Annual. Fine-earth, gravelly and stony slopes, screes, pebbles, fallow lands. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, ornamental,
poisonous, alkaloid-bearing.
Figure 114 Glaucium elegans (photography by Natalya Beshko)
139
140
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
396. Glaucium fimbrilligerum (Trautv.) Boiss. Figure 115
Biennial. Fine-earth, gravelly and stony slopes, dry riverbeds, pebbles, screes, cliffs, fallow lands.
Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Medicinal, ornamental, poisonous, alkaloid-bearing.
Figure 115 Glaucium fimbrilligerum (photography by Natalya Beshko)
397. Glaucium squamigerum Kar. & Kir.
Perennial. Stony slopes, rocks, screes, pebbles, ravines, banks of rivers. Montane and alpine zones.
Malguzar, North Turkestan. Poisonous, alkaloid-bearing.
Genus 125. Hypecoum L.
398. Hypecoum pendulum L. [Hypecoum parviflorum Kar. & Kir.] Figure 116
Annual. Sandy and clay deserts, fine-earth, gravelly and stony slopes, fallow lands, fields. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed, alkaloid-bearing.
Figure 116 Hypecoum pendulum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
399. Hypecoum trilobum Trautv.
Annual. Fine-earth and stony slopes, sandy and clay deserts, saline depressions, wastelands, fallow
lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Weed, alkaloid-bearing.
Genus 126. Corydalis DC.
400. Corydalis aitchisonii Popov [Corydalis sewerzowii Regel] Figure 117
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental, alkaloid-bearing.
Figure 117 Corydalis aitchisonii (photography by Natalya Beshko)
141
142
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
401. Corydalis glaucescens Regel Figure 118
Perennial. Fine-earth and stony slopes, shady wet places. Montane zone. Malguzar, North Turkestan.
Ornamental, medicinal, alkaloid-bearing.
Figure 118 Corydalis glaucescens (photography by Natalya Beshko)
402. Corydalis gortschakovii Schrenk
Perennial. Stony slopes, rocks, screes, subalpine and alpine meadows. Alpine zone. North Turkestan.
Ornamental, alkaloid-bearing.
403. Corydalis ledebouriana Kar. & Kir. Figure 119
Perennial. Fine-earth, gravelly and stony slopes, places near melting snow. Montane and alpine
zones. Nuratau, Malguzar, North Turkestan. Ornamental, medicinal, alkaloid-bearing.
Figure 119 Corydalis ledebouriana (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
404. Corydalis nudicaulis Regel
Perennial. Fine-earth, gravelly slopes, rocks, places near melting snow. Montane and alpine zones.
North Turkestan.
405. Corydalis schelesnowiana Regel & Schmalh.
Perennial. Stony slopes, rocks, screes. Montane zone. North Turkestan.
Genus 127. Fumaria L.
406. Fumaria vaillantii Loisel
Annual. Fine-earth, gravelly and stony slopes, screes, fallow lands, wastelands, roadsides, settlements,
gardens, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Medicinal weed, alkaloid-bearing, poisonous.
Family 32. Berberidaceae
Genus 128. Berberis L.
407. Berberis integerrima Bunge
Shrub. Fine-earth, gravelly and stony slopes, river valleys. Montane zone. Malguzar, North
Turkestan. Medicinal, food, dye, ornamental, meliferous, afforestation.
408. Berberis nummularia Bunge Figure 120
Shrub. Stony and gravelly slopes, rocks, screes, river valleys. Foothills, montane zone. Malguzar,
North Turkestan. Medicinal, food, dye, ornamental, melliferous, afforestation.
Figure 120 Berberis nummularia (photography by Natalya Beshko)
143
144
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
409. Berberis oblonga C. K. Schneid Figure 121
Shrub. Stony slopes, screes, river valleys. Montane zone. Malguzar, North Turkestan. Medicinal,
food, dye, ornamental, melliferous, afforestation.
Figure 121 Berberis oblonga (photography by Natalya Beshko)
Genus 129. Gymnospermium Spach
410. Gymnospermium albertii (Regel) Takht.
Perennial. Fine-earth, gravelly and stony slopes, screes. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Ornamental, alkaloid-bearing, saponin-bearing.
Genus 130. Leontice L.
411. Leontice eversmannii Bunge
Perennial. Fine-earth, gravelly and stony slopes, piedmont plains, river valleys, fallow lands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Ornamental, alkaloid-bearing, saponin-bearing, poisonous.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 131. Bongardia C. A. Mey.
412. Bongardia chrysogonum (L.) Boiss. Figure 122
Perennial. Fine-earth, gravelly and stony slopes, piedmont plains, fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Alkaloid-bearing,
saponin-bearing, ornamental.
Figure 122 Bongardia chrysogonum (photography by Natalya Beshko)
Family 33. Ranunculaceae
Genus 132. Aquilegia L.
413. Aquilegia lactiflora Kar. & Kir.
Perennial. Fine-earth and stony slopes, rocks, shady wet places, swamps, river valleys, places near
melting snow. Montane and alpine zones. North Turkestan. Ornamental, poisonous.
414. Aquilegia vicaria Nevski Figure 123
Perennial. Shady wet places, rocks, banks of rivers, swamps, wet places near springs. Montane and
alpine zones. Malguzar, North Turkestan. Ornamental, poisonous.
145
146
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 123 Aquilegia vicaria (photography by Natalya Beshko)
Genus 133. Isopyrum L.
415. Isopyrum anemonoides Kar. & Kir. Figure 124
Perennial. Shady wet places, fine-earth and stony slopes, rocks, alpine meadows, swamps, banks of
rivers, places near melting snow. Montane and alpine zones. North Turkestan.
Figure 124 Isopyrum anemonoides (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 134. Paraquilegia Drumm. & Hutch.
416. Paraquilegia caespitosa J. R. Drumm. & Hutch.
Perennial. Stony slopes, rocks, screes. Alpine zone. North Turkestan.
Genus 135. Thalictrum L.
417. Thalictrum isopyroides C. A. Mey.
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, alkaloid-bearing.
418. Thalictrum minus L. Figure 125
Perennial. River valleys, fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar,
North Turkestan. Medicinal, alkaloid-bearing.
Figure 125 Thalictrum minus (photography by Natalya Beshko)
419. Thalictrum sultanabadense Stapf.
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Alkaloid-bearing.
Genus 136. Adonis L.
420. *Adonis aestivalis L. Figure 126
Annual. Fine-earth, gravelly and stony slopes, wastelands, fields, fallow lands, roadsides. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Medicinal, weed, poisonous.
147
148
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 126 Adonis aestivalis (photography by Natalya Beshko)
421. Adonis turkestanica (Korsh.) Adolf. Figure 127
Perennial. Fine-earth, gravelly and stony slopes, subalpine meadows. Montane and alpine zones.
North Turkestan. Medicinal, weed, poisonous.
Figure 127 Adonis turkestanica (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 137. Trollius L.
422. Trollius komarovii Pachom. Figure 128
Perennial. Stream banks, swamps, places near melting snow. Alpine zone. North Turkestan.
Figure 128 Trollius komarovii (photography by Natalya Beshko)
Genus 138. Aconitum L.
423. Aconitum zeravschanicum Steinb
Perennial. Fine-earth and stony slopes, screes, subalpine and alpine meadows, places near melting
snow. Montane and alpine zones. Malguzar, North Turkestan. Alkaloid-bearing, medicinal,
poisonous, ornamental.
424. Aconitum talassicum Popov Figure 129
Perennial. Banks of rivers, ravines, wet places. Montane and alpine zones. Malguzar, North
149
150
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Turkestan. Ornamental, alkaloid-bearing, medicinal, poisonous. UzbRDB 3.
Figure 129 Aconitum talassicum (photography by Natalya Beshko)
Genus 139. Consolida S. F. Gray.
425. Consolida camptocarpa (Fisch. & C. A. Mey. ex Ledeb.) Nevski [Delphinium
camptocarpum Fisch. & C. A. Mey.] Figure 130
Annual. Sandy and clay deserts. Plain. Kyzylkum. Poisonous, alkaloid-bearing.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 130 Consolida camptocarpa (photography by Natalya Beshko)
426. Consolida leptocarpa Nevski [Delphinium leptocarpum Nevski] Figure 131
Annual. Sandy and clay deserts, river valleys, fine-earth and stony slopes. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Kyzylkum. Poisonous,
151
152
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
alkaloid-bearing.
Figure 131 Consolida leptocarpa (photography by Natalya Beshko)
427. Consolida rugulosa (Boiss.) Schrödinger [Delphinium rugulosum Boiss.]
Annual. Sandy and clay deserts, saline lands, fine-earth and stony slopes, fallow lands. Plain,
foothills. Nuratau Relic Mountains, Kyzylkum. Alkaloid-bearing, poisonous.
Genus 140. Delphinium L.
428. Delphinium barbatum Bunge Figure 132
Annual. Fine-earth and stony slopes, screes, river valleys, fallow lands, fields, roadsides. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Dye, alkaloidbearing.
Figure 132 Delphinium barbatum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
429. Delphinium batalinii Huth Figure 133
Perennial. Fine-earth and stony slopes, screes, banks of rivers, pebbles. Montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Alkaloid-bearing.
Figure 133 Delphinium batalinii (photography by Natalya Beshko)
153
154
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
430. Delphinium biternatum Huth Figure 134
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes, banks of rivers, subalpine meadows.
Montane and alpine zones. Nuratau, Malguzar. Dye, poisonous, alkaloid-bearing.
Figure 134 Delphinium biternatum (photography by Natalya Beshko)
431. Delphinium confusum Popov
Perennial. Fine-earth, gravelly and stony slopes, alpine meadows, banks of rivers. Montane and
alpine zones. North Turkestan. Dye, alkaloid-bearing, poisonous.
432. Delphinium longipedunculatum Regel & Schmalh. Figure 135
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Dye, alkaloid-bearing, poisonous.
Figure 135 Delphinium longipedunculatum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
433. Delphinium oreophilum Huth
Perennial. Fine-earth and stony slopes, rocks, screes, pebbles, moraines, banks of rivers, swamps,
subalpine and alpine meadows. Montane and alpine zones. Malguzar, North Turkestan. Alkaloidbearing.
434. Delphinium poltoratzkii Rupr.
Perennial. Fine-earth and stony slopes, rocks, screes, ravines, banks of rivers. Montane and alpine
zones. Malguzar, North Turkestan. Alkaloid-bearing.
435. Delphinium propinquum Nevski
Perennial. Fine-earth and stony slopes, places near melting snow. Alpine zone. North Turkestan.
Alkaloid-bearing.
436. Delphinium semibarbatum Bien. ex Boiss. Figure 136
Perennial. Clay deserts, relic mountains, piedmont plains, foothills, fine-earth and stony slopes,
screes, ravines, fallow lands, gardens. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Dye, medicinal, ornamental, poisonous, alkaloid-bearing.
Figure 136 Delphinium semibarbatum (photography by Natalya Beshko)
155
156
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 141. Nigella L.
437. Nigella integrifolia Regel Figure 137
Annual. Sandy and clay deserts, relic mountains, foothills, fine-earth and stony slopes, screes, fallow
lands, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum.
Figure 137 Nigella integrifolia (photography by Natalya Beshko)
Genus 142. Eranthis Salisb.
438. Eranthis longistipitata Regel Figure 138
Perennial. Fine-earth, gravelly and stony slopes, river valleys, places near melting snows. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Figure 138 Eranthis longistipitata (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 143. Callianthemum C. A. Mey.
439. Callianthemum alatavicum Freyn.
Perennial. Fine-earth and stony slopes, screes, rocks, moraines, pebbles, alpine meadows, banks of
rivers, wet places, places near melting snow. Alpine zone. North Turkestan. Alkaloid-bearing.
Genus 144. Anemone L.
440. Anemone biflora DC. var. petiolulosa (Juz.) S. Ziman [Anemone petiolulosa Juz.] Figure 139
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan.
Figure 139 Anemone biflora var. petiolulosa (photography by Natalya Beshko)
441. Anemone narcissiflora subsp. protracta (Ulbr.) Ziman & Fedor. [Anemone protracta
(Ulbr.) Juz.] Figure 140
Perennial. Fine-earth and stony slopes, alpine meadows, places near melting snow. Montane and
alpine zones. Malguzar, North Turkestan. Ornamental, poisonous.
Figure 140 Anemone narcissiflora subsp. protracta (photography by Natalya Beshko)
157
158
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
442. Anemone tschernaewii (Czern.) Regel
Perennial. Fine-earth and stony slopes, shady wet places. Montane zone. Malguzar, North Turkestan.
Genus 145. Clematis L.
443. Clematis orientalis L. Figure 141
Shrub or liana. Banks of rivers, canals and lakes, fine-earth and stony slopes. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Ornamental, alkaloid-bearing, poisonous.
Figure 141 Clematis orientalis (photography by Natalya Beshko)
Genus 146. Pulsatilla Adans.
444. Pulsatilla campanella Fisch. ex Krylov.
Perennial. Fine-earth and stony slopes, moraines. Alpine zone. North Turkestan. Ornamental,
alkaloid-bearing.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 147. Ceratocephala Moench
445. Ceratocephala falcata (L.) Рers. Figure 142
Annual. Sandy and clay desert, pebbles, fine-earth and stony slopes, screes, wastelands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed, poisonous.
Figure 142 Ceratocephala falcata (photography by Natalya Beshko)
446. Ceratocephala testiculata (Crantz) Besser
Annual. Sandy and clay deserts, pebbles, fine-earth and stony slopes, screes, wastelands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed, poisonous.
Genus 148. Halerpestes Green
447. Halerpestes sarmentosa (Adams) Kom.
Perennial. Swamps, banks of rivers. Montane and alpine zones. North Turkestan.
Genus 149. Ranunculus L.
448. *Ranunculus arvensis L.
Annual. Clayey soils, banks of rivers and canals, wastelands, gardens, fields, fallow lands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Weed, poisonous.
159
160
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
449. Ranunculus baldshuanicus Regel & Kom. Figure 143
Perennial. Wet places, meadows, banks of rivers and canals. Plain, foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Alkaloid-bearing.
Figure 143 Ranunculus baldshuanicus (photography by Natalya Beshko)
450. Ranunculus brevirostris Edgew
Perennial. Banks of rivers, wet places. Plain, foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Alkaloid-bearing.
451. Ranunculus komarovii Freyn
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau. Alkaloid-bearing.
452. Ranunculus linearilobus Bunge
Perennial. Sandy, clayey and skeleton soils. Plain, foothills. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, Kyzylkum. Medicinal, poisonous.
453. Ranunculus mindshelkensis B. Fedtsch. Figure 144
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan. Alkaloid-bearing.
Figure 144 Ranunculus mindshelkensis (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
454. Ranunculus natans C. A. Mey.
Perennial. Standing and slowly flowing water. Montane and alpine zones. North Turkestan.
455. Ranunculus olgae Regel
Perennial. Fine-earth and stony slopes, places near melting snow. Montane and alpine zones.
Malguzar, North Turkestan. Alkaloid-bearing.
456. Ranunculus oxyspermus Willd.
Perennial. River valleys, banks of canals, fields, gardens, roadsides, fallow lands, fine-earth and
gravelly slopes. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Weed,
alkaloid-bearing.
457. Ranunculus paucidentatus Schrenk Figure 145
Perennial. Fine-earth and stony slopes, places near melting snow. Montane and alpine zones.
Nuratau, Aktau, Malguzar, North Turkestan. Alkaloid-bearing.
Figure 145 Ranunculus paucidentatus (photography by Natalya Beshko)
458. Ranunculus pinnatisectus Popov Figure 146
Perennial. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, banks of
canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Weed, alkaloid-bearing.
161
162
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 146 Ranunculus pinnatisectus (photography by Natalya Beshko)
459. Ranunculus pulchellus C. A. Mey.
Perennial. Wet places, swamps, alpine meadows, banks of rivers. Alpine zone. North Turkestan.
Alkaloid-bearing.
460. Ranunculus repens L.
Perennial. Wet places, banks of rivers and canals, swamps, gardens. Plain, foothills, montane and
alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal,
alkaloid-bearing, poisonous.
461. Ranunculus rionii Lagger [Batrachium rionii (Lagger) Nym.]
Perennial. Standing and slowly flowing water. Plain, foothills, montane and alpine zones. Nuratau,
North Turkestan, Mirzachul, Kyzylkum.
462. Ranunculus rubrocalyx Regel ex Kom.
Perennial. Fine-earth and stony slopes, rocks, alpine meadows, banks of rivers, swamps, places near
melting snow. Alpine zone. North Turkestan.
463. Ranunculus rufosepalus Franch. Figure 147
Perennial. Fine-earth and stony slopes, screes, moraines, alpine meadows, places near melting snow.
Montane and alpine zones. Malguzar, North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 147 Ranunculus rufosepalus (photography by Natalya Beshko)
464. Ranunculus sceleratus L.
Annual or biennial. Wet places, swamps, banks of rivers and canals. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Weed, poisonous, alkaloid-bearing, weed.
465. Ranunculus sewerzowii Regel Figure 148
Perennial. Fine-earth and stony slopes, piedmont plains, sandy and clay deserts. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Alkaloid-bearing.
Figure 148 Ranunculus sewerzowii (photography by Natalya Beshko)
163
164
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
466. Ranunculus songaricus Schrenk
Perennial. Fine-earth, gravelly and stony slopes, subalpine meadows. Montane and alpine zones.
Malguzar, North Turkestan.
467. Ranunculus sphaerospermus Boiss. & C. I. Blanche [Batrachium pachycaulon Nevski]
Perennial. Standing and slowly flowing water. Plain, foothills, montane and alpine zones. Nuratau,
North Turkestan.
468. Ranunculus tenuilobus Regel & Kom.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
469. Ranunculus trichophyllus Chaix [Batrachium divaricatum (Schrank) Schur; Batrachium
trichophyllum (Chaix) Bosch]
Perennial. Standing and slowly flowing water. Plain, foothills, montane and alpine zones. Nuratau,
North Turkestan, Mirzachul, Kyzylkum.
Family 34. Platanaceae
Genus 150. Platanus L.
470. Platanus orientalis L.
Tree. River valleys. Montane zone. Nuratau, Malguzar, North Turkestan, Mirzachul (planted).
Industrial, ornamental, medicinal, afforestation.
Family 35. Paeoniaceae
Genus 151. Paeonia L.
471. Paeonia tenuifolia L. [Paeonia hybrida Pall.] Figure 149
Perennial. Fine-earth, gravelly and stony slopes, subalpine and alpine meadows, places near melting
snow. Montane and alpine zones. North Turkestan. Ornamental. UzbRDB 3.
Figure 149 Paeonia tenuifolia (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Family 36. Grossulariaceae
Genus 152. Ribes L.
472. Ribes meyeri Maxim. Figure 150
Shrub. Wet slopes, banks of rivers. Montane zone. Malguzar, North Turkestan. Food, ornamental.
Figure 150 Ribes meyeri (photography by Natalya Beshko)
Family 37. Saxifragaceae
Genus 153. Saxifraga L.
473. Saxifraga hirculus L.
Perennial. Banks of rivers, wetstony slopes, screes, moraines, alpine meadows, places near melting
165
166
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
snow. Alpine zone. North Turkestan. UzbRDB 2.
474. Saxifraga sibirica L.
Perennial. Stony slopes, screes, rocks, moraines, alpine meadows. Alpine zone. North Turkestan.
Meliferous.
Family 38. Crassulaceae
Genus 154. Clemensia Rose
475. Clementsia semenovii (Regel & Herder) Boriss. Figure 151
Perennial. Wet stony slopes, pebbles, swamps, banks of rivers. Alpine zone. North Turkestan.
Medicinal, ornamental, dye.
Figure 151 Clementsia semenovii (photography by Natalya Beshko)
Genus 155. Rhodiola L.
476. Rhodiola heterodonta (Hook f. & Thomson) Boriss. Figure 152
Perennial. Stony slopes, rocks, alpine meadows. Alpine zone. North Turkestan. Medicinal,
ornamental.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 152 Rhodiola heterodonta (photography by Natalya Beshko)
Genus 156. Sedum L.
477. Sedum hispanicum L. [Sedum pentapetalum Boriss.]
Annual. Stony slopes. Montane zone. Malguzar.
Figure 153 Sedum hispanicum (photography by Natalya Beshko)
Figure 153
167
168
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
478. Sedum tetramerum Trautv. Figure 154
Annual. Stony, slopes, clay deserts, piedmont plain, pebbles. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar.
Figure 154 Sedum tetramerum (photography by Natalya Beshko)
Genus 157. Pseudosedum (Boiss.) A. Berger
479. Pseudosedum bucharicum Boriss.
Perennial. Stony slopes, rocks. Montane zone. Malguzar, North Turkestan. Ornamental.
480. Pseudosedum campanuliflorum Boriss. Figure 155
Perennial. Stony slopes, rocks. Montane zone. Malguzar, North Turkestan. Ornamental, threatened
species. UzbRDB 1.
Figure 155 Pseudosedum campanuliflorum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
481. Pseudosedum lievenii (Ledeb.) A. Berger.
Perennial. Stony slopes, rocks. Foothills, montane zone. Nuratau, Malguzar. Ornamental.
482. Pseudosedum longidentatum Boriss. Figure 156
Perennial. Stony slopes, rocks, screes. Foothills, montane and alpine zones. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Ornamental.
Figure 156 Pseudosedum longidentatum (photography by Natalya Beshko)
Genus 158. Rosularia (DC.) Stapf.
483. Rosularia radicosa (Boiss. & Hohen.) Eggli [Rosularia paniculata (Regel & Schmalh.) A.
Berger]
Perennial. Stony slopes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
484. Rosularia subspicata (Freyn & Sint.) Boriss.
Perennial. Stony slopes. Montane zone. Nuratau.
Family 39. Haloragaceae
Genus 159. Myriophyllum L.
485. Myriophyllum spicatum L.
Perennial. Standing and slowly flowing water. Plain, foothills, montane zone. Nuratau, Mirzachul.
Medicinal, fodder, dye, ornamental, industrial.
169
170
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 40. Vitaceae
Genus 160. Vitis L.
486. Vitis vinifera L. Figure 157
Shrub. Ravines, mountain valleys. Montane zone, Nuratau. Food, medicinal, ornamental, melliferous.
UzbRDB 2.
Figure 157 Vitis vinifera (photography by Natalya Beshko)
Family 41. Zygophyllaceae
Genus 161. Zygophyllum L.
487. Zygophyllum jaxarticum Popov
Perennial. Saline lands. Plain. Mirzachul.
488. Zygophyllum macrophyllum Regel & Schmalh. Figure 158
Perennial. Saline lands, banks of canals, fields, fallow lands, wastelands. Plain. Mirzachul.
Figure 158 Zygophyllum macrophyllum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
489. Zygophyllum miniatum Cham. Figure 159
Perennial. Clay deserts, relic mountains. Plain, foothills. Nuratau Relic Mountains, Mirzachul.
Figure 159 Zygophyllum miniatum (photography by Natalya Beshko)
490. Zygophyllum oxianum Boriss. Figure 160
Perennial. Saline lands, sandy soils, river valleys, banks of canals, fields, fallow lands, wastelands.
Plain, foothills. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Weed, medicinal,
poisonous.
Figure 160 Zygophyllum oxianum (photography by Natalya Beshko)
Genus 162. Tribulus L.
491. *Tribulus terrestris L.
Annual. Sandy, clayey and saline soils, pebbles, fields, fallow lands, wastelands, roadsides. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
171
172
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Mirzachul, Kyzylkum. Weed, medicinal, poisonous.
Family 42. Fabaceae [Leguminosae]
Genus 163. *Gleditsia L.
492. *Gleditsia caspia Desf.
Tree. River valleys, fine-earth slopes (cultivated and naturalized). Foothills, montane zone. Nuratau.
Ornamental, afforestation, industrial, fodder.
Genus 164. Sophora L.
493. Sophora alopecuroides L. [Vexibia alopecuroides (L.) Yakovlev] Figure 161
Perennial. Fine-earth and gravelly slopes, river valleys, meadows, swamps, wet places, fields, fallow
lands, roadsides, banks of canals. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North
Turkestan. Weed, poisonous.
Figure 161 Sophora alopecuroides (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
494. Sophora lehmanni (Bunge) Yakovlev [Ammothamnus lehmanni Bunge]
Semishrub. Sandy deserts. Plain. Kyzylkum. Poisonous, ornamental, afforestation, melliferous,
insecticide.
495. Sophora pachycarpa C. A. Mey. [Vexibia pachycarpa (C. A. Mey.) Yakovlev] Figure 162
Perennial. Sandy, clayey and saline soils, fine-earth and gravelly slopes, banks of canals, fields,
fallow lands, roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, Mirzachul, Kyzylkum. Weed, poisonous.
Figure 162 Sophora pachycarpa (photography by Natalya Beshko)
173
174
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 165. Ammodendron Fisch.
496. Ammodendron conollyi Bunge ex Boiss. Figure 163
Tree or shrub. Sandy deserts, saline lands. Plain. Kyzylkum. Dye, ornamental, industrial,
afforestation, melliferous.
Figure 163 Ammodendron conollyi (photography by Natalya Beshko)
497. Ammodendron lehmannii Bunge ex Boiss.
Tree or shrub. Sandy deserts. Plain. Kyzylkum, Mirzachul. Ornamental, afforestation, melliferous.
Genus 166. Ononis L.
498. Ononis spinosa subsp. antiquorum (L.) Briq.
Perennial. Fine-earth and stony slopes, river valleys, banks of canals, roadsides, fields. Foothills,
montane zone. Nuratau. Fodder.
Genus 167. Trigonella L.
499. Trigonella adscendens (Nevski) Afan. & Gontsch. [Melissitus adscendens (Nevski) Ikonn.]
Perennial. Fine-earth and stony slopes, pebbles. Montane and alpine zones, Malguzar, North
Turkestan. Fodder.
500. Trigonella aristata (Vassilcz.) Sojak [Melissitus aristatus (Vass.) Latsch.]
Perennial. Stony slopes, dry riverbeds, river valleys. Montane zone. Malguzar, North Turkestan.
Fodder.
501. Trigonella cancellata Desf.
Annual. Fine-earth and stony slopes, pebbles, fallow lands, dry riverbeds. Foothills, montane zone.
North Turkestan. Fodder.
MAGNOLIOPSIDA
[EUDICOTS]
502. Trigonella geminiflora Bunge Figure 164
Annual. Fine-earth and stony slopes, sandy and clay deserts, fallow lands, roadsides, banks of rivers,
dry riverbeds. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Fodder.
Figure 164 Trigonella geminiflora (photography by Natalya Beshko)
503. Trigonella gontscharovii Vassilcz. [Melissitus gontscharovii (Vassilcz.) Latsch.]
Perennial. Stony slopes. Montane and alpine zones. Malguzar, North Turkestan. Fodder.
504. Trigonella grandiflora Bunge Figure 165
Annual. Fine-earth and stony slopes, banks of canals, roadsides, dry riverbeds, fallow lands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Fodder.
Figure 165 Trigonella grandiflora (photography by Natalya Beshko)
175
176
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
505. Trigonella popovii Korovin [Melissitus popovii (Korovin) Golosk.]
Perennial. Fine-earth, gravelly slopes, river valleys. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Fodder.
506. Trigonella verae Sirj.
Annual. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Fodder.
Genus 168. Melissitus Medik.
507. Melissitus pamiricus (Boriss.) Golosk
Perennial. Stony slopes, rocks, screes, pebbles. Montane and alpine zones. Malguzar, North
Turkestan. Fodder.
Genus 169. Medicago L.
508. Medicago lupulina L.
Annual or biennial. Fine-earth, gravelly and stony slopes, river valleys, banks of canals, pebbles,
meadows, gardens, fields. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, medicinal, melliferous.
509. Medicago medicaginoides (Retz.) E. Small [Trigonella arcuata C. A. Mey.]
Annual. Fine-earth and stony slopes, fallow lands, banks of rivers, dry riverbeds. Foothills, montane
zone. Malguzar, North Turkestan.
510. Medicago minima (L.) L.
Annual or biennial. Clay deserts, fine-earth, gravelly and stony slopes, dry riverbeds, river valleys,
fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Fodder, melliferous, weed.
511. Medicago monantha (C. A. Mey.) Trautv. [Trigonella noeana Boiss.]
Annual. Fine-earth and stony slopes, fields, fallow lands, dry riverbeds. Foothills, montane zone.
Malguzar. Fodder, melliferous, weed.
512. Medicago orbicularis (L.) Bartal
Annual. Fine-earth, gravelly and stony slopes, river valleys, pebbles, fields, fallow lands. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Fodder, melliferous, weed.
513. Medicago orthoceras (Kar. & Kir.) Trautv. [Trigonella orthoceras Kar. & Kir.]
Annual. Fine-earth and stony slopes, fields, fallow lands, dry riverbeds, roadsides. Plain, foothills,
montane zone. Nuratau, Malguzar. Fodder, medicinal, dye.
514. Medicago polymorpha L. [Medicago denticulata Willd.]
Annual. Piedmont plains, fine-earth and gravelly slopes, gardens, fields. Plain, foothills, montane
zone. North Turkestan. Fodder, medicinal, melliferous.
MAGNOLIOPSIDA
[EUDICOTS]
515. Medicago rigidula (L.) All.
Annual. Fine-earth, gravelly and stony slopes, piedmont plains, clay deserts, river valleys, pebbles.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Fodder, melliferous, weed.
516. *Medicago sativa L.
Perennial. Fine-earth, gravelly and stony slopes, meadows, screes, river valleys, pebbles, gardens,
pastures, fields, fallow lands, roadsides (naturalized). Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, medicinal,
melliferous.
Genus 170. Melilotus Mill.
517. Melilotus albus Medik.
Biennial. Meadows, river valleys, banks of canals, fine-earth slopes, pebbles, wet places, fallow
lands, fields, gardens. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, medicinal, melliferous, insecticide.
518. Melilotus officinalis (L.) Pall.
Biennial. Fine-earth slopes, meadows, river valleys, roadsides, gardens, fields, fallow lands. Plain,
foothills. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder, medicinal, melliferous, insecticide.
Genus 171. Trifolium L.
519. Trifolium campestre Schreb. [Chrysaspis campestris (Schreb.) Desv.]
Annual. River valleys, meadows, fine-earth slopes, fields. Foothills, montane zone. Malguzar. Fodder,
melliferous.
520. Trifolium fragiferum L.[Trifolium neglectum C. A. Mey.; Amoria bonannii (C. Presl.)
Roskov; Amoria fragifera (L.) Roskov]
Perennial. Wet places, meadows, banks of rivers and canals, swamps, pebbles, fields, fallow
lands. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Fodder,
melliferous.
521. Trifolium pratense L. Figure 166
Perennial. River valleys, banks of canals, fine-earth slopes, wet places, meadows, fields, fallow
lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Fodder, medicinal, essential oil, dye, melliferous.
177
178
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 166 Trifolium pratense (photography by Natalya Beshko)
522. Trifolium repens L.
Perennial. Wet places, meadows, banks of rivers and canals, swamps, pebbles, fields, fallow
lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Fodder, medicinal, melliferous.
Genus 172. Lotus L.
523. Lotus krylovii Schischkin & Serg. [L. sergievskiae Kamelin & Kovalevsk.]
Perennial. River valleys, banks of rivers, lakes and canals, fallow lands, meadows, fields, gardens,
saline lands. Plain, foothills, montane zone. Nuratau, Malguzar, North Turkestan. Fodder, weed.
Genus 173. Cullen Medik.
524. Cullen drupaceum (Bunge) C. H. Stirt. [Psoralea drupacea Bunge] Figure 167
Perennial. Fine-earth slopes, piedmont plain, sandy and clay deserts, banks of rivers, lakes
and canals, fallow lands, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, essential oil,
MAGNOLIOPSIDA
[EUDICOTS]
meliferous, weed.
Figure 167 Cullen drupaceum (photography by Natalya Beshko)
Genus 174. Smirnowia Bunge
525. Smirnowia turkestana Bunge
Shrub. Sandy deserts. Plain. Kyzylkum. Medicinal.
Genus 175. Colutea L.
526. Colutea paulsenii Freyn Figure 168
Shrub. Fine-earth and stony slopes. Montane zone. Malguzar, North Turkestan. Ornamental,
afforestation, melliferous.
Figure 168 Colutea paulsenii (photography by Natalya Beshko)
179
180
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 176. Halimodendron Fisch. ex DC.
527. Halimodendron halodendron (Pall.) Voss. Figure 169
Shrub. Saline lands, banks of rivers, lakes and canals, pebbles. Plain, foothills, montane zone.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Dye, fodder,
ornamental, afforestation, melliferous.
Figure 169 Halimodendron halodendron (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 177. Caragana Fabr.
528. Caragana alaica Pojark.
Shrub. Stony slopes, ravines, river valleys. Montane zone. Malguzar, North Turkestan. Melliferous.
529. Caragana turkestanica Kom.
Shrub. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan. Melliferous.
Genus 178. Chesneya Lindl.
530. Chesneya ternata (Korsh.) Popov Figure 170
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes, pebbles, banks of rivers. Foothills,
montane zone. Nuratau, Malguzar, North Turkestan.
Figure 170 Chesneya ternata (photography by Natalya Beshko)
181
182
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
531. Chesneya turkestanica Franch.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan.
Genus 179. Astragalus L.
532. Astragalus adpressipilosus Gontsch.
Perennial. Stony slopes, pebbles. Montane and alpine zones. Malguzar, North Turkestan.
533. Astragalus alabugensis B.
Fedtsch. Perennial. Stony slopes. Montane zone. Nuratau.
534. Astragalus alopecias Pall. Figure 171
Perennial. Fine-earth slopes, piedmont plains, sandy and clay deserts, screes, fallow lands, banks
of rivers and canals, roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North
Turkestan, Mirzachul. Fodder, ornamental.
Family 171 Astragalus alopecias (photography by Natalya Beshko)
535. Astragalus alpinus L.
Perennial. Fine-earth and gravelly slopes, banks of rivers, subalpine and alpine meadows. Montane
and alpine zones. North Turkestan. Fodder, ornamental, melliferous.
536. Astragalus ammotrophus Bunge
Perennial. Stony and gravelly slopes, clay deserts. Plain, foothills. Nuratau Relic Mountains.
MAGNOLIOPSIDA
[EUDICOTS]
537. Astragalus angreni Lipsky
Perennial. Fine-earth slopes. Montane zone. Malguzar, North Turkestan.
538. Astragalus aphanassjievii Gontsch.
Perennial. Fine-earth and gravelly slopes. Montane and alpine zones. Malguzar, North Turkestan.
539. Astragalus aschuturi B. Fedtsch.
Perennial. Fine-earth, gravelly and stony slopes, screes, dry riverbeds. Montane and alpine zones,
Malguzar, North Turkestan.
540. Astragalus bactrianus Bunge Figure 172
Shrub. Fine-earth, gravelly and stony slopes, pebbles, fallow lands. Foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan.
Figure 172 Astragalus bactrianus (photography by Natalya Beshko)
541. Astragalus bakaliensis Bunge
Annual. Sandy deserts, stony slopes, fields, fallow lands. Plain, foothills, montane zone. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder, melliferous.
542. Astragalus belolipovii Kamelin ex F. O. Khass. & N. Sulajm.
Perennial. Fine-earth and stony slopes, among juniper forests. Montane zone. Malguzar, North
Turkestan. UzbRDB 1.
543. Astragalus camptoceras Bunge
Annual. Fine-earth, gravelly and stony slopes, piedmont plains. Plain, foothills, montane zone.
183
184
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
544. Astragalus campylorrhynchus Fisch. & C. A. Mey. Figure 173
Annual. Fine-earth, gravelly and stony slopes, sandy and clay deserts. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder.
Figure 173 Astragalus campylorrhynchus (photography by Natalya Beshko)
545. Astragalus campylotrichus Bunge
Annual. Fine-earth, gravelly and stony slopes, sandy and clay deserts. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder.
546. Astragalus chodshenticus B. Fedtsch.
Semishrub. River valleys, saline lands. Plain, foothills. Malguzar.
547. Astragalus commixtus Bunge
Annual. Sandy, clayey and sceleton soils. Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder.
548. Astragalus compositus Pavlov
Annual. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Fodder, weed.
549. Astragalus contortuplicatus L.
Annual. Saline lands, banks of rivers, lakes and canals, fields. Plain. Mirzachul. Fodder, weed.
MAGNOLIOPSIDA
[EUDICOTS]
550. Astragalus cornu-bovis Lipsky
Annual. Sandy and clay deserts. Plain. Kyzylkum. Fodder.
551. Astragalus dendroides Kar. & Kir.
Shrub. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan.
552. Astragalus dictamnoides Gontsch.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan.
553. Astragalus dipelta Bunge
Annual. Fine-earth and stony slopes, fields, fallow lands. Foothills, montane zone. Malguzar, North
Turkestan. Fodder.
554. Astragalus dolichocarpus Popov
Semishrub. Fine-earth and gravelly slopes. Foothills, montane zone. Nuratau, Malguzar.
555. Astragalus eximius Bunge Figure 174
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Ornamental.
Figure 174 Astragalus eximius (photography by Natalya Beshko)
556. Astragalus falcigerus Popov
Semishrub. Gravelly and stony slopes. Montane zone. Nuratau.
557. Astragalus farctissimus Lipsky
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Aktau, Malguzar, North
Turkestan.
558. Astragalus ferganensis (Popov) B. Fedtsch. ex A. S. Korol.
Perennial. Stony slopes, pebbles. Foothills. North Turkestan.
559. Astragalus filicaulis Fisch. & C. A. Mey.
Annual. Sandy, clayey and skeleton soils, fields, fallow lands. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder.
185
186
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
560. Astragalus flexus Fisch. Figure 175
Perennial. Sandy deserts. Plain. Kyzylkum. Fodder.
Figure 175 Astragalus flexus (photography by Natalya Beshko)
561. Astragalus floccosifolius Sumnev. Figure 176
Perennial. Fine-earth and stony slopes, river valleys, fallow lands. Montane zone. Malguzar, North
Turkestan.
Figure 176 Astragalus floccosifolius (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
562. Astragalus globiceps Bunge Figure 177
Perennial. Fine-earth, gravelly and stony slopes, fields, fallow lands. Foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan. Fodder, ornamental.
Figure 177 Astragalus globiceps (photography by Natalya Beshko)
563. Astragalus guttatus Bans & Solander [Astragalus striatellus M. Bieb.]
Annual. Fine-earth, gravelly and stony slopes. Foothills. Nuratau. Fodder.
564. Astragalus harpilobus Kar. & Kir.
Annual. Sandy deserts. Plain. Kyzylkum. Fodder, weed.
565. Astragalus inaequalifolius Basil.
Perennial. Stony slopes, rocks, screes. Montane zone. Nuratau.
566. Astragalus intarrensis Franch.
Perennial. Stony slopes. Montane zone. Malguzar, North Turkestan.
567. Astragalus iskanderi Lipsky
Shrub. Fine-earth and stony slopes, river valleys. Montane zone. Nuratau, Malguzar, North
Turkestan.
568. Astragalus isfahanicus Boiss.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan.
569. Astragalus jagnobicus Lipsky
Perennial. Fine-earth, gravelly and stony slopes, pebbles. Montane and alpine zones. Malguzar,
North Turkestan.
570. Astragalus juratzkanus Freyn & Sint. [Astragalus maverranagri Popov]
Semishrub. Fine-earth, gravelly and stony slopes, roadsides. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
571. Astragalus kelleri Popov Figure 178
Perennial. Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar. UzbRDB 2.
187
188
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 178 Astragalus kelleri (photography by Natalya Beshko)
572. Astragalus knorringianus Boriss. Figure 179
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. UzbRDB 2.
Figure 179 Astragalus knorringianus (photography by Natalya Beshko)
573. Astragalus lasiosemius Boiss.
Semishrub. Fine-earth, gravelly and stony slopes, screes. Montane and alpine zones. Malguzar, North
Turkestan.
574. Astragalus lasiostylus Fisch. [Astragalus vladimiri Sirj.]
Shrub. Fine-earth and stony slopes, rocks, screes. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan.
575. Astragalus leiophysa Bunge
Perennial. Sandy deserts. Plain. Kyzylkum.
576. Astragalus lentilobus Kamelin & Kovalevsk.
Perennial. Fine-earth and gravelly slopes, screes, moraines, subalpine and alpine meadows. Montane
and alpine zones. Malguzar, North Turkestan. Fodder.
MAGNOLIOPSIDA
[EUDICOTS]
577. Astragalus leptophysus Vved. Figure 180
Perennial. Stony slopes, screes. Montane zone. Nuratau, North Turkestan, Nuratau Reserve. UzbRDB 2.
Figure 180 Astragalus leptophysus (photography by Natalya Beshko)
189
190
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
578. Astragalus leptostachys Pall. Figure 181
Perennial. Fine-earth, gravelly and stony slopes, pebbles. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Fodder.
Figure 181 Astragalus leptostachys (photography by Natalya Beshko)
579. Astragalus lipskyi Popov Figure 182
Perennial. Fine-earth and gravelly slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Figure 182 Astragalus lipskyi (photography by Natalya Beshko)
580. Astragalus lithophilus Kar. & Kir.
Perennial. Fine-earth, gravelly and stony slopes, pebbles, moraines, screes, alpine meadows.
Montane and alpine zones, North Turkestan.
581. Astragalus macrocladus Bunge Figure 183
Semishrub. Sandy deserts. Plain. Kyzylkum. Fodder, melliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 183 Astragalus macrocladus (photography by Natalya Beshko)
582. Astragalus macronyx Bunge Figure 184
Perennial. Fine-earth, gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan.
Figure 184 Astragalus macronyx (photography by Natalya Beshko)
191
192
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
583. Astragalus macropetalus Schrenk
Perennial. Fine-earth, gravelly slopes, Clay deserts. Plain, foothills. Nuratau.
584. Astragalus macrotropis Bunge
Perennial. Fine-earth, gravelly and stony slopes, fallow lands, rocks, banks of rivers, pebbles.
Foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan.
585. Astragalus marguzaricus Lipsky
Perennial. Fine-earth and stony slopes, banks of rivers, pebbles. Montane zone. Nuratau, Malguzar,
North Turkestan.
586. Astragalus masenderanus Bunge [Astragalus kurdaicus Saposhn. ex Sumn.; Astragalus
skorniakovii B. Fedtsch.] Figure 185
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Fodder.
Figure 185 Astragalus masenderanus (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
587. Astragalus mucidus Bunge ex Boiss. Figure 186
Perennial. Fine-earth, gravelly and stony slopes, piedmont plains. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Figure 186 Astragalus mucidus (photography by Natalya Beshko)
588. Astragalus nematodes Bunge
Perennial. Fine-earth, gravelly and stony slopes, rocks. Foothills. Nuratau.
589. Astragalus nivalis Kar. & Kir.
Perennial. Gravelly and stony slopes, pebbles, banks of rivers, moraines. Montane and alpine zones.
North Turkestan.
590. Astragalus nobilis Bunge ex B. Fedtsch. Figure 187
Perennial. Fine-earth, gravelly and stony slopes, rocks. Montane zone. Nuratau, Aktau, Malguzar,
North Turkestan.
Figure 187 Astragalus nobilis (photography by Natalya Beshko)
193
194
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
591. Astragalus nuciferus Bunge Figure 188
Perennial. Fine-earth and stony slopes, screes. Montane zone. Malguzar, North Turkestan.
Figure 188 Astragalus nuciferus (photography by Natalya Beshko)
592. Astragalus ophiocarpus Benth. ex Boiss.
Annual. Sandy and clay deserts, banks of rivers. Plain. Mirzachul. Fodder.
593. Astragalus orbiculatus Ledeb.
Perennial. Sandy deserts, saline lands, wet places, banks of rivers. Plain. Malguzar, Mirzachul,
Kyzylkum.
594. Astragalus oxyglottis Stev. ex M. Bieb.
Annual. Fine-earth, skeleton, rarely sandy soils. Plain, foothills, montane zone. Nuratau, Nuratau
Relic Mountains, Malguzar, Mirzachul. Fodder.
595. Astragalus patentipilosus Kitam. [Astragalus korovinianus Barneby]
Perennial. Clayey soils, among sagebrush vegetation. Plain, foothills. Nuratau, Mirzachul. Fodder.
596. Astragalus patentivillosus Gontsch.
Perennial. Fine-earth and gravelly slopes, moraines. Montane and alpine zone. North Turkestan.
597. Astragalus paucijugus Schrenk
Shrub. Sandy deserts. Plain. Kyzylkum. Fodder, meliferous.
598. Astragalus pauper Bunge
Perennial. Fine-earth and gravelly slopes, alpine meadows. Alpine zone. North Turkestan. Fodder.
599. Astragalus peduncularis Benth. [Astragalus corydalinus Bunge]
Perennial. Stony slopes, pebbles, screes, wet meadows. Montane and alpine zone. Malguzar, North
Turkestan.
600. Astragalus persipolitanus Boiss. [Astragalus ammophilus Kar. & Kir.]
Annual. Sandy, clayey and skeleton soils, screes. Plain, foothills, montane zone. Nuratau, Nuratau
Relic Mountains, Mirzachul, Kyzylkum. Fodder.
601. Astragalus petunnikovii Litv.
Perennial. Sandy deserts. Plain. Kyzylkum.
MAGNOLIOPSIDA
[EUDICOTS]
602. Astragalus platyphyllus Kar. & Kir.
Perennial. Stony slopes. Montane zone. Malguzar, North Turkestan.
603. Astragalus pterocephalus Bunge [Astragalus stipulosus Boriss.]
Shrub. Fine-earth, gravelly and stony slopes, screes, rocks. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan.
604. Astragalus pulcher Korovin Figure 189
Perennial. Stony slopes, rocks. Montane zone. Nuratau.
Figure 189 Astragalus pulcher (photography by Natalya Beshko)
605. Astragalus quisqualis Bunge
Perennial. Fine-earth, gravelly and stony slopes, screes. Montane zone. Malguzar, North Turkestan.
Fodder.
606. Astragalus retamocarpus Boiss. & Hohen.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Malguzar. Fodder.
607. Astragalus rubromarginatus Czerniak.
Perennial. Sandy deserts. Plain. Kyzylkum.
608. Astragalus russanovii F. O. Khass., Sarybaeva & Esankulov
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan.
609. Astragalus saratagius Bunge
Perennial. Stony slopes, screes. Montane and alpine zones. North Turkestan.
610. Astragalus sarytavicus Popov
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan.
611. Astragalus schachimardanus Basil.
Perennial. Stony slopes, rocks, screes. Montane zone. Malguzar, North Turkestan.
612. Astragalus schmalhausenii Bunge
Annual. Fine-earth, gravelly and stony slopes, fields, fallow lands. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder, weed.
613. Astragalus schrenkianus Fisch. & C. A. Mey.
Perennial. Stony slopes, rocks, screes. Montane zone. Nuratau.
195
196
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
614. Astragalus schugnanicus B. Fedtsch.
Perennial. Fine-earth, gravelly and stony slopes, alpine meadows, stream banks. Alpine zone. North
Turkestan. Fodder.
615. Astragalus sesamoides Boiss.
Annual. Fine-earth, gravelly and stony slopes, pebbles, river valleys, fields, fallow lands. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Fodder.
616. Astragalus sewerzowii Bunge [Astragalus subbarbellatus Bunge]
Perennial. Fine-earth slopes, subalpine meadows. Montane and alpine zones. Nuratau, Aktau,
Malguzar, North Turkestan. Fodder.
617. Astragalus sieversianus Pall. Figure 190
Perennial. Fine-earth, gravelly and stony slopes, fallow lands. Foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Fodder, medicinal, ornamental.
Figure 190 Astragalus sieversianus (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
618. Astragalus sogdianus Bunge Figure 191
Perennial. Fine-earth, gravelly and stony slopes, piedmont plain. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Figure 191 Astragalus sogdianus (photography by Natalya Beshko)
619. Astragalus stalinskyi Širj.
Annual. Fine-earth, gravelly and stony slopes, piedmont plain. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder.
620. Astragalus stenanthus Bunge
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Foothills, montane zone. Nuratau,
Malguzar.
621. Astragalus stenocystis Bunge
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan.
622. Astragalus subinduratus Gontsch.
Perennial. Stony slopes, banks of rivers. Montane and alpine zones. North Turkestan.
623. Astragalus subscaposus Popov ex Boriss.
Perennial. Fine-earth, gravelly and stony slopes, wet places near springs, meadows. Montane and
alpine zones. North Turkestan. Fodder.
624. Astragalus substipitatus Gontsch.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
197
198
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Turkestan.
625. Astragalus subverticillatus Gontsch.
Perennial. Fine-earth, gravelly slopes. Montane zone. Malguzar, North Turkestan.
626. Astragalus tibetanus Benth. ex Bunge
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Montane and alpine zones. Malguzar,
North Turkestan. Fodder.
627. Astragalus titovii Gontsch.
Perennial. Stony slopes, swamps, places near melting snow. Montane and alpine zones. Malguzar,
North Turkestan. Fodder.
628. Astragalus transoxanus Fisch. ex Bunge
Shrub. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau.
629. Astragalus turbinatus Bunge
Perennial. Sandy deserts. Plain. Kyzylkum.
630. Astragalus turczaninovii Kar. & Kir. Figure 192
Perennial. Sandy deserts. Plain. Kyzylkum.
Figure 192 Astragalus turczaninovii (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
631. Astragalus turkestanus Bunge Figure 193
Perennial. Fine-earth and gravelly slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Fodder.
Figure 193 Astragalus turkestanus (photography by Natalya Beshko)
199
200
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
632. Astragalus unifoliolatus Bunge
Semishrub. Sandy deserts. Plain. Kyzylkum. Fodder, melliferous.
633. Astragalus uninodus Popov & Vved.
Annual. River valleys, stony slopes. Montane zone. Malguzar, North Turkestan.
634. Astragalus urgutinus Lipsky
Shrub. Fine-earth, gravelly and stony slopes, rocks, cliffs. Foothills, montane and alpine zones.
Nuratau, Aktau, Malguzar, North Turkestan.
635. Astragalus variegatus Franch. Figure 194
Shrub. Fine-earth, gravelly and stony slopes, banks of rivers, fallow lands. Montane and alpine
zones. Nuratau, Malguzar, North Turkestan.
Figure 194 Astragalus variegatus (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
636. Astragalus vicarius Lipsky
Annual. Fine-earth, gravelly and stony slopes, fallow lands, fields. Foothills, montane zone. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder.
637. Astragalus villosissimus Bunge
Semishrub. Sandy and clay deserts, saline lands. Plain, foothills. Nuratau Relic Mountains,
Mirzachul, Kyzylkum. Fodder, melliferous.
638. Astragalus xanthomeloides Korovin & Popov Figure 195
Perennial. Stony slopes, screes, pebbles. Montane zone. Nuratau, Malguzar, North Turkestan.
Figure 195 Astragalus xanthomeloides (photography by Natalya Beshko)
201
202
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 180. Oxytropis DC.
639. Oxytropis aspera Gontsch. Fedtsch.
Perennial. Stony slopes, places near melting snow. Montane and alpine zones. North Turkestan.
640. Oxytropis capusii Franch. Figure 196
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Malguzar, North
Turkestan.
Figure 196 Oxytropis capusii (photography by Natalya Beshko)
641. Oxytropis chesneyoides Gontsch.
Perennial. Rocks, stony slopes. Foothills, montane zone. Nuratau, Malguzar.
642. Oxytropis humifusa Kar. & Kir.
Perennial. Fine-earth, gravelly and stony slopes, moraines, alpine meadows, places near melting
snow. Alpine zone. North Turkestan.
643. Oxytropis immersa (Baker) B. Fedtsch.
Perennial. Fine-earth, gravelly and stony slopes, rocks, moraines, alpine meadows. Alpine zone.
North Turkestan.
644. Oxytropis integripetala Bunge
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. North Turkestan.
645. Oxytropis kamelinii Vassilcz.
Perennial. Stony slopes. Montane zone. Malguzar. Endemic.
646. Oxytropis lehmanni Bunge
Perennial. Fine-earth and stony slopes, rocks, alpine meadows. Montane and alpine zones. Malguzar,
North Turkestan.
647. Oxytropis leucocyanea Bunge
Semishrub. Stony slopes, screes. Montane and alpine zones. North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
648. Oxytropis lipskyi Gontsch.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan.
649. Oxytropis macrocarpa Kar. & Kir. Figure 197
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Figure 197 Oxytropis macrocarpa (photography by Natalya Beshko)
203
204
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
650. Oxytropis michelsonii B. Fedtsch.
Perennial. Stony slopes, places near melting snow. Montane and alpine zones. North Turkestan.
651. Oxytropis microsphaera Bunge
Perennial. Stony slopes, rocks, places near melting snow. Montane and alpine zones. Malguzar,
North Turkestan.
652. Oxytropis platonychia Bunge
Perennial. Stony slopes, screes, pebbles, alpine meadows. Alpine zone. North Turkestan.
653. Oxytropis pseudorosea Filim. Figure 198
Perennial. Stony slopes. Montane zone. Nuratau, UzbRDB 2.
Figure 198 Oxytropis pseudorosea (photography by N. Yu. Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
654. Oxytropis riparia Litv.
Perennial. Meadows, river valleys, wet slopes. Montane zone. North Turkestan. Endemic to the
Nuratau Mountains.
655. Oxytropis rosea Bunge Figure 199
Perennial. Dry riverbeds, pebbles, gravelly and stony slopes, rocks. Foothills, montane zone.
Malguzar, North Turkestan.
Figure 199 Oxytropis rosea (photography by Natalya Beshko)
656. Oxytropis savellanica Bunge
Semishrub. Stony slopes, screes, rocks, alpine meadows, stream banks. Montane and alpine zones.
North Turkestan.
657. Oxytropis seravschanica Gontsch.
Perennial. Stony slopes, dry riverbeds. Montane and alpine zones. Malguzar, North Turkestan.
658. Oxytropis tachtensis Franch.
Perennial. Fine-earth and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
659. Oxytropis trichocalycina Bunge Figure 200
Perennial. Stony slopes, rocks, screes. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan.
205
206
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 200 Oxytropis trichocalycina (photography by Natalya Beshko)
Genus 181. Glycyrrhiza L.
660. Glycyrrhiza aspera Pall.
Perennial. Banks of rivers, saline lands, fine-earth slopes, fallow lands, fields. Plain, foothills, montane
zone. Nuratau, Nuratau Relic Mountains, Malguzar, Mirzachul. Medicinal, fodder, melliferous.
661. Glycyrrhiza glabra L.
Perennial. Fine-earth slopes, meadows, fields, fallow lands, river valleys, banks of canals. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Medicinal, food, fodder, dye, industrial, melliferous, weed.
662. Glycyrrhiza triphylla Fisch. & C. A. Mey. [Meristotropis triphylla Fisch. & C. A. Mey.]
Perennial. Fine-earth, gravelly and stony slopes, river valleys, pebbles, fallow lands. Foothills,
montane zone. Nuratau. Medicinal, fodder, melliferous.
Genus 182. Hedysarum L.
663. Hedysarum flavescens Regel & Schmalh.
Perennial. Fine-earth, gravelly and stony slopes, screes, rocks, river valleys, alpine meadows.
Montane and alpine zones. North Turkestan. Fodder, ornamental, meliferous.
664. Hedysarum jaxarticum Popov & Kir.
Perennial. Clayey soils. Plain, foothills. North Turkestan. Fodder, ornamental, meliferous.
665. Hedysarum mogianicum (B. Fedtsch.) B. Fedtsch.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau. Fodder, ornamental,
melliferous.
666. Hedysarum montanum (B. Fedtsch.) B. Fedtsch.
Perennial. Pebbles, fine-earth and stony slopes, fallow lands, roadsides. Foothills, montane zone.
Malguzar, North Turkestan. Fodder, ornamental, melliferous.
MAGNOLIOPSIDA
[EUDICOTS]
667. Hedysarum nuratense Popov Figure 201
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Fodder, ornamental, melliferous.
Figure 201 Hedysarum nuratense (photography by Natalya Beshko)
207
208
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
668. Hedysarum plumosum Boiss. & Hausskn. Figure 202
Perennial. Stony slopes, screes, rocks, pebbles. Montane and alpine zones. Malguzar, North
Turkestan. Fodder, ornamental, melliferous.
Figure 202 Hedysarum plumosum (photography by Natalya Beshko)
Genus 183. Onobrychis Hill
669. Onobrychis chorassanica Bunge Figure 203
Perennial. Fine-earth, gravelly slopes, fallow lands. Foothills, montane zone. Nuratau, Aktau,
Malguzar, North Turkestan. Fodder, melliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 203 Onobrychis chorassanica (photography by Natalya Beshko)
670. Onobrychis echidna Lipsky
Shrub. Stony slopes, screes. Montane and alpine zones. Malguzar, North Turkestan. Fodder,
melliferous.
209
210
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
671. Onobrychis gontscharovii Vassilcz.
Perennial. Rocks, stony slopes. Montane zone. Nuratau. Fodder, melliferous.
672. Onobrychis grandis Lipsky Figure 204
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan. Fodder,
melliferous.
Figure 204 Onobrychis grandis (photography by Natalya Beshko)
673. Onobrychis micrantha Schrenk
Annual. Sandy and clay deserts, fine-earth, gravelly and stony slopes, fields, fallow lands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Fodder, meliferous, weed.
674. Onobrychis pulchella Schrenk Figure 205
Annual. Fine-earth and stony slopes, pebbles, fields, fallow lands, piedmont plains, clay deserts.
Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Fodder, meliferous, weed.
Figure 205 Onobrychis pulchella (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
675. Onobrychis saravschanica B. Fedtsch.
Perennial. Fine-earth and stony slopes, screes. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan. Fodder, melliferous.
Genus 184. Alhagi Hill
676. Alhagi canescens (Regel) B. Keller & Shap.
Perennial. Sandy and clay deserts, roadsides, fallow and saline lands, banks of canals. Plain,
foothills. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder, medicinal, melliferous.
677. Alhagi kirghisorum Schrenk Figure 206
Perennial. Sandy and clay deserts, fine-earth, gravelly and stony slopes, roadsides, fallow lands,
fields, banks of canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, medicinal, melliferous.
Figure 206 Alhagi kirghisorum (photography by Natalya Beshko)
678. Alhagi pseudalhagi (M. Bieb.) Desv. ex B. Keller & Shap.
Perennial. Saline wet places, sandy deserts, river valleys, banks of lakes and canals, fallow lands,
fields. Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Fodder, medicinal, melliferous.
Genus 185. Cicer L.
679. Cicer flexuosum Lipsky
Perennial. Stony slopes, screes, pebbles. Montane zone. Nuratau, Malguzar, North Turkestan. Fodder.
680. Cicer grande (Popov) Korotkova Figure 207
Perennial. Fine-earth and stony slopes. Montane zone. Nuratau, Nuratau Reserve. UzbRDB 2. Fodder, food, threatened species.
211
212
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 207 Cicer grande (photography by N. Yu. Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
681. Cicer macracanthum Popov
Perennial. Fine-earth and stony slopes, pebbles. Montane and alpine zones. Malguzar, North
Turkestan.
682. Cicer paucijugum (Popov) Nevski.
Perennial. Fine-earth and stony slopes. Montane and alpine zones. North Turkestan. Fodder.
683. Cicer pungens Boiss.
Perennial. Fine-earth and stony slopes, pebbles. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan.
684. Cicer songaricum DC. Figure 208
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Fodder, food.
Figure 208 Cicer songaricum (photography by Natalya Beshko)
213
214
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 186. Vicia L.
685. Vicia anatolica Turrill [Vicia hajastana Grossh.]
Annual. Stony slopes. Foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North
Turkestan. Fodder.
686.*Vicia ervilia (L.) Willd.
Annual. Fine-earth, gravelly and stony slopes, fallow lands, fields. Foothills, montane zone, Nuratau,
Aktau, Malguzar, North Turkestan. Fodder, food, weed.
687. Vicia hyrcanica Fisch. & C. A. Mey.
Annual. Fine-earth slopes, meadows, river valleys, wastelands, gardens, fields, fallow lands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Fodder, weed.
688. Vicia kokanica Regel & Schmalh.
Perennial. Fine-earth and stony slopes, screes, river valleys. Montane zone. Nuratau, Malguzar,
North Turkestan. Fodder.
689. Vicia michauxii Spreng. Figure 209
Annual. Fine-earth, gravelly and stony slopes, river valleys, meadows, fallow lands, fields, gardens.
Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Fodder, weed.
Figure 209 Vicia michauxii (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
690. *Vicia peregrina L. [Vicia gracilior (Popov) Popov ex B. Fedtsch.]
Annual. Fine-earth, gravelly and stony slopes, river valleys, meadows, screes, fallow lands, fields,
roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar.
Fodder, weed.
691. *Vicia sativa subsp. nigra (L). Ehrh. [Vicia angustifolia L.]
Annual. Fine-earth slopes, meadows, river valleys, wastelands, gardens, fields, fallow lands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Fodder, melliferous.
692. Vicia subvillosa (Ledeb.) Boiss. Figure 210
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Fodder.
Figure 210 Vicia subvillosa (photography by Natalya Beshko)
693. Vicia tenuifolia Roth.
Perennial. Fine-earth, gravelly slopes, river valleys, meadows, fallow lands. Foothills, montane zone.
Malguzar, North Turkestan. Fodder, melliferous.
694. *Vicia villosa Roth.
Annual. Meadows, gardens, fallow lands, river valleys, fine-earth slopes. Plain, foothills, montane
zone. Nuratau, Malguzar, North Turkestan. Fodder, weed, melliferous.
215
216
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 187. Lens Mill.
695. Lens culinaris subsp. orientalis (Boiss.) Ponert [Lens orientalis (Boiss.) Schmalh.]
Annual. Fine-earth, gravelly slopes, fallow lands. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder, food, weed.
Genus 188. Lathyrus L.
696. Lathyrus aphaca L.
Annual. Fine-earth, gravelly and stony slopes, river valleys, banks of canals, pebbles, fallow lands,
gardens, fields, meadows. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Fodder, medicinal, dye, melliferous.
697.*Lathyrus cicera L.
Annual. Fine-earth and stony slopes, river valleys, meadows, pebbles, banks of streams and canals,
fields. Plain, foothills, montane zone. Nuratau, Malguzar. Fodder, medicinal, melliferous.
698. Lathyrus inconspicuus L.
Annual. Fine-earth, gravelly slopes, ravines, fallow lands, fields, gardens, banks of rivers, dry
riverbeds. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Fodder.
699. Lathyrus pratensis L. Figure 211
Perennial. Fine-earth and stony slopes, ravines, banks of rivers, lakes and canals, pebbles, wet meadows.
Plain, foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan, Mirzachul. Fodder,
medicinal, melliferous.
Figure 211 Lathyrus pratensis (photography by Natalya Beshko)
700. *Lathyrus sativus L. [Lathyrus asiaticus (Zalkind) Kudr.]
Annual. River valleys, fields, fallow lands, wastelands, ravines. Plain, foothills, montane and alpine
zones. Nuratau, Nuratau Relic Mountains. Fodder, food, weed.
MAGNOLIOPSIDA
[EUDICOTS]
701. Lathyrus tuberosus L.
Perennial. Fine-earth and stony slopes, river valleys, wet meadows, banks of canals, fallow lands,
fields. Foothills, montane zone. North Turkestan. Fodder, food, medicinal.
Family 43. Polygalaceae
Genus 189. Polygala L.
702. Polygala hybrida DC.
Perennial. Fine-earth and stony slopes, banks of streams, wet places, subalpine meadows, gardens.
Foothills, montane and alpine zones. Malguzar, North Turkestan. Medicinal, ornamental, melliferous.
Family 44. Rosaceae
Genus 190. Spiraea L.
703. Spiraea hypericifolia L. Figure 212
Shrub. River valleys, fine-earth, gravelly and stony slopes, screes, rocks, pebbles. Montane zone.
Malguzar, North Turkestan. Medicinal, afforestation, ornamental, melliferous.
Figure 212 Spiraea hypericifolia (photography by Natalya Beshko)
704. Spiraea pilosa Franch. Figure 213
Shrub. Stony slopes, banks of rivers, rocks, ravines. Montane zone. North Turkestan. Ornamental,
melliferous.
217
218
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 213 Spiraea pilosa (photography by Natalya Beshko)
Genus 191. Cotoneaster Medik.
705. Cotoneaster goloskokovii Pojark.
Shrub. Stony slopes, screes, ravines. Montane zone. North Turkestan. Industrial, afforestation,
ornamental, melliferous.
706. Cotoneaster multiflorus Bunge Figure 214
Shrub. Rocks, stony slopes, ravines. Montane zone. Nuratau, North Turkestan. Industrial,
afforestation, ornamental, melliferous.
Figure 214 Cotoneaster multiflorus (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
707. Cotoneaster nummularioides Pojark. Figure 215
Shrub. Fine-earth, gravelly and stony slopes, rocks, screes, pebbles, banks of rivers, ravines. Montane
zone. Nuratau, Malguzar, North Turkestan. Industrial, afforestation, ornamental, melliferous.
Figure 215 Cotoneaster nummularioides (photography by Natalya Beshko)
708. Cotoneaster nummularius Fisch. & C.A. Mey.
Shrub. Fine-earth, gravelly and stony slopes, rocks, subalpine meadows. Montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Industrial, afforestation, ornamental, melliferous.
709. Cotoneaster oliganthus Pojark. Figure 216
Shrub. Fine-earth, gravelly and stony slopes, rocks. Montane zone. Malguzar, North Turkestan.
Industrial, afforestation, ornamental, melliferous.
Figure 216 Cotoneaster oliganthus (photography by Natalya Beshko)
219
220
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
710. Cotoneaster songaricus (Regel & Herder) Popov Figure 217
Shrub. Fine-earth, gravelly and stony slopes, rocks. Montane and alpine zones. Nuratau, North
Turkestan. Industrial, afforestation, ornamental, melliferous.
Figure 217 Cotoneaster songaricus (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
711. Cotoneaster suavis Pojark.
Shrub. Fine-earth, gravelly and stony slopes, river valleys. Montane zone. Malguzar, North
Turkestan. Industrial, afforestation, ornamental, melliferous.
Genus 192. Pyrus L.
712. *Pyrus communis L.
Tree River valleys (planted and naturalized). Montane zone. Nuratau. Food, ornamental, industrial,
dye, melliferous.
713. Pyrus korshinskyi Litv.
Tree. Fine-earth and stony slopes, river valleys. Montane zone. Malguzar. Ornamental, melliferous.
IUCN CR.
714. Pyrus regelii Rehd. Figure 218
Shrub. Fine-earth, gravelly and stony slopes, river valleys. Montane zone. Nuratau, Malguzar, North
Turkestan. Ornamental, melliferous.
Figure 218 Pyrus regelii (photography by Natalya Beshko)
221
222
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
715. Pyrus turcomanica Maleev.
Tree. Fine-earth, gravelly and stony slopes, river valleys. Foothills, montane zone. Nuratau. Food,
ornamental, industrial, dye, melliferous.
Genus 193. Malus Mill.
716. Malus sieversii (Ledeb.) M. Roem Figure 219
Tree. Fine-earth, gravelly and stony slopes, ravines, river valleys. Foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Food, fodder, industrial, dye, ornamental, melliferous. IUCN VU.
Figure 219 Malus sieversii (photography by Natalya Beshko)
Genus 194. Sorbus L.
717. Sorbus persica Hedl. Figure 220
Tree. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Afforestation, industrial, ornamental, melliferous.
Figure 220 Sorbus persica (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
718. Sorbus tianschanica Rupr. Figure 221
Tree. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, North Turkestan. Medicinal,
food, afforestation, industrial, ornamental, melliferous.
Figure 221 Sorbus tianschanica (photography by Natalya Beshko)
Genus 195. Crataegus L.
719. Crataegus pontica C. Koch Figure 222
Tree. Fine-earth slopes, stream banks. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Food, afforestation, industrial, ornamental, melliferous.
Figure 222 Crataegus pontica (photography by Natalya Beshko)
223
224
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
720. Crataegus songarica C. Koch Figure 223
Tree. Fine-earth, gravelly and stony slopes, screes, banks of rivers. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan. Food, medicinal, afforestation, industrial, ornamental, melliferous.
Figure 223 Crataegus songarica (photography by Natalya Beshko)
721. Crataegus turkestanica Pojark. Figure 224
Tree. Fine-earth, gravelly and stony slopes, rocks, screes, river valleys. Foothills, montane zone.
Nuratau, Malguzar, North Turkestan. Food, medicinal, afforestation, industrial, ornamental,
melliferous.
Figure 224 Crataegus turkestanica (photography by Natalya Beshko)
Genus 196. Rubus L.
722. Rubus caesius L. Figure 225
Shrub. Woodlands, wet stony slopes, screes, pebbles, fallow lands, river valleys, banks of canals,
gardens. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan.
Medicinal, food, dye, ornamental, fodder, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 225 Rubus caesius (photography by Natalya Beshko)
Genus 197. Potentilla L.
723. Potentilla asiatica (Th. Wolf) Juz.
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Montane and alpine zones. Malguzar,
North Turkestan. Medicinal, melliferous.
724. Potentilla asiae-mediae Ovcz. & Koczk.
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Malguzar, North
Turkestan. Melliferous.
725. Potentilla desertorum Bunge
Perennial. Fine-earth, gravelly and stony slopes, rocks. Montane and alpine zones. Nuratau, Malguzar. melliferous.
726. Potentilla flabellata Regel & Schmalh.
Perennial. Wet fine-earth and stony slopes. Alpine zone. North Turkestan. Melliferous.
727. Potentilla gelida C. A. Mey. Figure 226
Perennial. Fine-earth and stony slopes, rocks, moraines, alpine meadows. Montane and alpine zones.
North Turkestan. Medicinal, fodder, melliferous.
225
226
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 226 Potentilla gelida (photography by Natalya Beshko)
728. Potentilla hololeuca Boiss. ex Lehm.
Perennial. Fine-earth, gravelly and stony slopes, river valleys, alpine meadows. Alpine zone. North
Turkestan. melliferous.
729. Potentilla inclinata Vill. [Potentilla impolita Wahlenb.]
Perennial. Fine-earth and stony slopes, fallow lands. Montane and alpine zones. North Turkestan.
melliferous.
730. Potentilla orientalis Juz. Figure 227
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Foothills, montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Fodder, melliferous.
Figure 227 Potentilla orientalis (photography by Natalya Beshko)
731. Potentilla pamiroalaica Juz.
Perennial. Fine-earth and stony slopes, rocks, pebbles, alpine meadows. Alpine zone. Malguzar,
North Turkestan. Melliferous.
732. Potentilla pedata Willd. ex Hornem
Perennial. Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Medicinal, melliferous.
MAGNOLIOPSIDA
[EUDICOTS]
733. Potentilla reptans L. Figure 228
Perennial. Meadows, banks of rivers, lakes and canals, wet places. Plain, foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, melliferous.
Figure 228 Potentilla reptans (photography by Natalya Beshko)
734. Potentilla soongorica Bunge
Perennial. Fine-earth, gravelly slopes. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Melliferous.
735. Potentilla supina L.
Perennial. River valleys, swamps, wet places, banks of canals, roadsides, wastelands, fields, fineearth and stony slopes. Plain, foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Medicinal, fodder, melliferous.
736. Potentilla vvedenskyi Botsch.
Perennial. Rocks, alpine meadows, moraines, stony slopes. Alpine zone. North Turkestan.
Melliferous.
Genus 198. Pentaphylloides Duham.
737. Pentaphylloides parvifolia (Fisch. ex Lehm.) Soják
Shrub. Fine-earth and stony slopes, rocks, pebbles, river valleys, alpine meadows. Montane and
alpine zones. North Turkestan. Medicinal, fodder, ornamental, meliferous.
Genus 199. Sibbaldia L.
738. Sibbaldia tetrandra Bunge
Shrub. Moraines, fine-earth and stony slopes, screes, rocks, alpine meadows, places near melting
snow. Alpine zone. North Turkestan.
227
228
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 200. Geum L.
739. Geum heterocarpum Boiss. [Orthurus heterocarpus (Boiss.) Juz.] Figure 229
Perennial. Fine-earth, gravelly and stony slopes, stream banks, shady wet places. Montane zone.
Nuratau, Malguzar, North Turkestan. Medicinal, meliferous.
Figure 229 Geum heterocarpum (photography by Natalya Beshko)
Genus 201. Orthurus Juz.
740. Orthurus kokanicus (Regel & Schmalh. ex Regel) Juz.
Perennial. Fine-earth and stony slopes, rocks. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan. Medicinal, meliferous.
Genus 202. Alchemilla L.
741. Alchemilla fontinalis Juz.
Perennial. Stream banks, alpine meadows. Alpine zone. North Turkestan.
742. Alchemilla krylovii Juz.
Perennial. Alpine meadows, stream banks, swamps. Alpine zone. North Turkestan.
743. Alchemilla sibirica Zamelis
Perennial. Wet places, stream banks. Montane and alpine zones. North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 203. Agrimonia L.
744. Agrimonia asiatica Juz.
Perennial. Fine-earth and stony slopes, river valleys, meadows, gardens, banks of canals, roadsides.
Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Genus 204. Sanguisorba L.
745. *Sanguisorba minor Scop. [Poterium lasiocarpum Boiss. & Hausskn.]
Perennial. Fine-earth and stony slopes, river valleys, fallow lands, fields, meadows, gardens,
roadsides. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Fodder, meliferous.
746. *Sanguisorba minor subsp. balearica (Bourg. & Nyman) Muoz Garm. & C. Navarro
[Poterium polygamum Waldst. & Kitag.]
Perennial. Fine-earth and stony slopes, river valleys, fallow lands, fields, meadows, gardens,
roadsides. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Fodder, meliferous.
Genus 205. Rosa L.
747. Rosa beggeriana Schrenk & Fisch. ex C. A. Mey. Figure 230
Shrub. Fine-earth slopes, river valleys, banks of canals, roadsides. Plain, foothills, montane zone.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, ornamental.
Figure 230 Rosa beggeriana (photography by Natalya Beshko)
229
230
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
748. Rosa canina L. Figure 231
Shrub. River valleys, fine-earth, gravelly and stony slopes. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, food,
tanniferous, ornamental, afforestation, meliferous.
Figure 231 Rosa canina (photography by Natalya Beshko)
749. Rosa corymbifera Borkh
Shrub. River valleys, fine-earth and stony slopes. Montane zone. Nuratau. Medicinal, food,
ornamental, afforestation, meliferous.
750. Rosa ecae Aitch. Figure 232
Shrub. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Medicinal, food, ornamental, meliferous, weed.
Figure 232 Rosa ecae (photography by Natalya Beshko)
751. Rosa fedtschenkoana Regel Figure 233
Shrub. Fine-earth, gravelly and stony slopes, ravines, river valleys. Montane and alpine zones.
Malguzar, North Turkestan. Medicinal, food, ornamental, afforestation, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 233 Rosa fedtschenkoana (photography by Natalya Beshko)
752. Rosa hissarica Slobodova
Shrub. Stony slopes, screes, rocks. Montane and alpine zones. North Turkestan. Medicinal,
ornamental, meliferous.
753. Rosa lehmanniana Bunge
Shrub. Ravines, river valleys, wet fine-earth slopes. Montane zone. Nuratau. Medicinal, ornamental,
meliferous.
754. Rosa kokanica (Regel) Regel ex Juz. Figure 234
Shrub. Fine-earth, gravelly and stony slopes, ravines, river valleys. Montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Medicinal, ornamental, afforestation, meliferous.
231
232
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 234 Rosa kokanica (photography by Natalya Beshko)
755. Rosa maracandica Bunge Figure 235
Shrub. Fine-earth, gravelly and stony slopes, rocks, pebbles, river valleys. Foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, ornamental, meliferous.
Figure 235 Rosa maracandica (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
756. Rosa nanothamnus Boulenger
Shrub. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan. Medicinal, ornamental, meliferous.
757. Rosa transturkestanica N. F. Russanov.
Shrub. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Ornamental,
meliferous.
Genus 206. Hulthemia Dumort.
758. Hulthemia persica (Michx. & Juss.) Bornm.
Shrub. Wastelands, pebbles, sandy soils, piedmont plains, gravelly and stony slopes, fallow lands,
fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Ornamental, weed, meliferous.
Genus 207. Prunus L.
759. Prunus armeniaca L. [Armeniaca vulgaris Lam.]
Tree. Fine-earth slopes, river valleys. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Food, oil, medicinal, tanniferous, dye, afforestation, industrial, ornamental, meliferous. IUCN EN.
760. Prunus bucharica (Korsh.) B. Fedtsch. ex Rehder [Amygdalus bucharica Korsh.] Figure 236
Tree or shrub. Fine-earth and stony slopes, rocks, screes, river valleys. Foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, food, oil, ornamental, dye, afforestation,
industrial, meliferous. IUCN VU.
Figure 236 Prunus bucharica (photography by Natalya Beshko)
233
234
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
761. *Prunus cerasus L. [Cerasus vulgaris Mill.]
Tree. River valleys (naturalized). Nuratau. Food, medicinal, ornamental, meliferous.
762. Prunus divaricata Ledeb. Figure 237
Tree. Fine-earth, gravelly and stony slopes, screes, river valleys, ravines. Montane zone. Nuratau,
Malguzar, North Turkestan. Medicinal, food, oil, ornamental, afforestation, meliferous.
Figure 237 Prunus divaricata (photography by Natalya Beshko)
763. *Prunus domestica L.
Tree. Fine-earth slopes, river valleys (naturalized). Montane zone. Nuratau. Medicinal, food, oil,
ornamental, afforestation, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
764.*Prunus dulcis (Mill.) D.A. Webb [Amygdalus communis L.]
Tree. Fine-earth and stony slopes, screes, banks of rivers. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan. Medicinal, food, oil, ornamental, afforestation, industrial, meliferous.
765. Prunus erythrocarpa (Nevski) Gilli [Cerasus erythrocarpa Nevski] Figure 238
Shrub. Fine-earth and stony slopes, rocks, screes, ravines, banks of rivers. Foothills, montane zone.
Nuratau, Malguzar, North Turkestan. Ornamental, industrial, afforestation, meliferous.
Figure 238 Prunus erythrocarpa (photography by Natalya Beshko)
235
236
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
766. Prunus mahaleb L. [Cerasus mahaleb (L.) Mill.] Figure 239
Tree. Fine-earth and stony slopes, rocks, screes, ravines, banks of rivers. Montane zone. Nuratau,
Malguzar, North Turkestan. Food, ornamental, afforestation, meliferous.
Figure 239 Prunus mahaleb (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
767. Prunus spinosissima (Bunge) Franch. [Amygdalus spinosissima Bunge] Figure 240
Shrub. Fine-earth, gravelly and stony slopes, rocks. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Oil, dye, ornamental, afforestation,
meliferous.
Figure 240 Prunus spinosissima (photography by Natalya Beshko)
237
238
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
768. Prunus verrucosa Franch.[Cerasus amygdaliflora Nevski, Cerasus verrucosa (Franch.)
Nevski] Figure 241
Shrub. Stony slopes, rocks, screes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Ornamental, afforestation, meliferous.
Figure 241 Prunus verrucosa (photography by Natalya Beshko)
Family 45. Elaeagnaceae
Genus 208. Hippophae L.
769. Hippophae rhamnoides L. Figure 242
Shrub. River valleys, ravines. Montane zone. North Turkestan. Medicinal, dye, food, ornamental,
meliferous, afforestation.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 242 Hippophae rhamnoides (photography by Natalya Beshko)
Genus 209. Elaeagnus L.
770. Elaeagnus angustifolia L. Figure 243
Tree. Banks of lakes and canals, river valleys. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Food, medicinal,
afforestation, industrial, tanniferous, dye, ornamental, meliferous.
Figure 243 Elaeagnus angustifolia (photography by Natalya Beshko)
239
240
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 46. Rhamnaceae
Genus 210. Sageretia Brongn.
771. Sageretia thea (Osbeck) M. S. Johnst. [Sageratia laetevirens (Kom). Gontsch.]
Shrub. Fine-earth and stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
Ornamental, afforestation.
Genus 211. Rhamnus L.
772. Rhamnus cathartica L.
Tree or shrub. Stony and gravelly slopes, river valleys, ravines. Montane zone. Nuratau, Malguzar,
North Turkestan. Medicinal, ornamental, afforestation, industrial, dye, oil, meliferous.
773. Rhamnus coriacea (Regel) Kom. Figure 244
Shrub. Fine-earth, gravelly and stony slopes, screes, cliffs, ravines. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan. Ornamental, afforestation.
Figure 244 Rhamnus coriacea (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Family 47. Ulmaceae
Genus 212. Ulmus L.
774. *Ulmus glabra Huds.
Tree. Fine-earth and stony slopes, river valleys (naturalized), settlements, forest belts, cemeteries
(planted). Foothills, montane zone. Nuratau, Malguzar, North Turkestan. Industrial, ornamental,
afforestation.
775. *Ulmus laevis Pall.
Tree. Fine-earth and stony slopes, river valleys (naturalized), settlements, forest belts, cemeteries
(planted). Plain, foothills, montane zone. Nuratau, North Turkestan. Industrial, ornamental,
afforestation.
776. *Ulmus minor Mill
Tree. Fine-earth and stony slopes, river valleys (naturalized), settlements, forest belts, cemeteries
(planted). Plain, foothills, montane zone. Nuratau, Malguzar. Industrial, ornamental, afforestation.
777. Ulmus pumila L.
Tree or shrub. Fine-earth and stony slopes, river valleys, ravines, rocks. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Industrial, ornamental,
afforestation.
Genus 213. Celtis L.
778. Celtis australis subsp. caucasica (Willd.) C. C. Towns. [Celtis caucasica Willd.]
Tree. Fine-earth and stony slopes, screes, rocks, river valleys. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan. Industrial, ornamental, afforestation, medicinal, tanniferous.
Family 48. Cannabaceae
Genus 214. Cannabis L.
779. *Cannabis sativa L.
Annual. Roadsides, fallow lands, wastelands, banks of canals. Plain, Mirzachul (naturalized).
Narcotic, industrial, oil, weed.
241
242
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 49. *Moraceae
Genus 215. *Morus L.
780. *Morus alba L.
Tree. Fine-earth and stony slopes, river valleys (naturalized), settlements, gardens, forest belts, burials
(planted). Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Food, fodder, medicinal, ornamental, industrial, dye, tanniferous.
781. *Morus nigra L.
Tree. Fine-earth and stony slopes, river valleys (naturalized), settlements, gardens (planted).
Plain, foothills, montane zone. Nuratau, Malguzar, North Turkestan. Food, medicinal, ornamental,
industrial, dye, tanniferous.
Family 50. Urticaceae
Genus 216. Urtica L.
782. Urtica dioica L.
Perennial. River valleys, roadsides, wastelands, gardens, riparian forests, banks of canals. Plain,
foothills, montane and alpine zones. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Medicinal, food, dye.
Genus 217. Parietaria L.
783. Parietaria debilis G. Forst [Parietaria micrantha Ledeb.]
Annual. Rocks, ravines, shady wet places. Montane and alpine zones, Nuratau, Malguzar. Alkaloidbearing.
784. Parietaria lusitanica subsp. serbica (Pančić) P. W. Ball [Parietaria serbica Panćić]
Annual. Ravines, screes, rocks. Montane zone. Malguzar, North Turkestan. Alkaloid-bearing.
Family 51. Juglandaceae
Genus 218. Juglans L.
785. Juglans regia L.
Tree. River valleys, wet slopes. Montane zone. Nuratau, Malguzar, North Turkestan. Food, industrial,
MAGNOLIOPSIDA
[EUDICOTS]
oil, afforestation, ornamental, medicinal, dye. IUCN NT.
Family 52. Betulaceae
Genus 219. Betula L.
786. Betula pendula Roth. Figure 245
Tree. River valleys, wet slopes. Montane zone. North Turkestan. Industrial, ornamental, afforestation.
Figure 245 Betula pendula (photography by Natalya Beshko)
Family 53. Cucurbitaceae
Genus 220. Bryonia L.
787. *Bryonia cretica subsp. dioica (Jacq.) Tutin [Bryonia dioica Jacq.] Figure 246
Perennial. Stony slopes, ravines, shady places. Foothills, montane zone. Nuratau. Medicinal,
243
244
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
essential oil, ornamental, poisonous.
Figure 246 Bryonia cretica subsp. dioica (photography by Natalya Beshko)
788. Bryonia melanocarpa Nabiev Figure 247
Perennial. Sandy deserts. Plain. Kyzylkum. Medicinal, poisonous. Threatened species, UzbRDB 2.
Endemic to the South-eastern Kyzylkum.
Figure 247 Bryonia melanocarpa (photography by N. Yu. Beshko)
Family 54. Datiscaceae
Genus 221. Datisca L.
789. Datisca cannabina L. Figure 248
Perennial. Banks of rivers, swamp meadows, pebbles, wet slopes. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan. Medicinal, dye, ornamental.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 248 Datisca cannabina (photography by Natalya Beshko)
Family 55. Celastraceae
Genus 222. Parnassia L.
790. Parnassia laxmannii Pall. ex Schult.
Perennial. Meadows, stony slopes, rocks, ravines, banks of rivers, swamps. Montane and alpine
zones. North Turkestan.
Family 56. Hypericaceae
Genus 223. Hypericum L.
791. Hypericum elongatum Ledeb. ex Rchb. Figure 249
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes. Montane and alpine zones. Malguzar,
245
246
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
North Turkestan. Medicinal.
Figure 249 Hypericum elongatum (photography by Natalya Beshko)
792. Hypericum perforatum L. Figure 250
Perennial. Fine-earth, gravelly and stony slopes, river valleys, dry riverbeds. Foothills, montane
zone. Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, dye, essential oil, meliferous.
Figure 250 Hypericum perforatum (photography by Natalya Beshko)
793. Hypericum scabrum L. Figure 251
Perennial. Fine-earth, gravelly and stony slopes, screes, rocks, dry riverbeds. Foothills, montane and
alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, dye,
MAGNOLIOPSIDA
[EUDICOTS]
essential oil, meliferous.
Figure 251 Hypericum scabrum (photography by Natalya Beshko)
Family 57. Violaceae
Genus 224. Viola L.
794. Viola alaica Vved.
Perennial. Screes, wet stony slopes, rocks, shady places. Montane zone. Malguzar, North Turkestan.
795. Viola occulta Lehm. Figure 252
Annual. Fine-earth, gravelly and stony slopes, meadows, fallow lands. Foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan.
Figure 252 Viola occulta (photography by Natalya Beshko)
247
248
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
796. Viola suavis M. Bieb. Figure 253
Perennial. Fine-earth, gravelly and stony slopes, stream banks, shady places, gardens. Foothills,
montane zone. Nuratau, North Turkestan. Medicinal, ornamental, meliferous.
Figure 253 Viola suavis (photography by Natalya Beshko)
797. Viola turkestanica Regel & Schmalh.
Perennial. Fine-earth and stony slopes, rocks, screes, shady wet places, meadows. Montane and
alpine zones. Malguzar, North Turkestan.
Family 58. Salicaceae
Genus 225. Populus L.
798. Populus afghanica (Aitch. & Hemsl.) C. K. Schneid. Figure 254
Tree. River valleys. Foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Industrial,
ornamental, afforestation, tanniferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 254 Populus afghanica (photography by Natalya Beshko)
799. Populus alba L. Figure 255
Tree. River valleys, banks of canals. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North
Turkestan, Mirzachul. Industrial, ornamental, afforestation, tanniferous, meliferous.
Figure 255 Populus alba (photography by Natalya Beshko)
249
250
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
800. Populus euphratica Oliv. Figure 256
Tree. River valleys, banks of lakes and canals, riparian forests. Plain, foothills. Kyzylkum,
Mirzachul. Industrial, ornamental, afforestation, tanniferous, meliferous.
Figure 256 Populus euphratica (photography by Natalya Beshko)
801. *Populus nigra L.
Tree. River valleys. Plain, foothills, montane zone (naturalized). Nuratau. Industrial, ornamental,
afforestation, tanniferous, medicinal, meliferous.
802. Populus pruinosa Schrenk Figure 257
Tree. River valleys, banks of lakes and canals, riparian forests. Plain. Kyzylkum, Mirzachul.
Industrial, ornamental, afforestation, tanniferous, dye.
Figure 257 Populus pruinosa (photography by Natalya Beshko)
803. Populus talassica Kom.
Tree. River valleys, fine-earth and stony slopes. Montane and alpine zones. Malguzar, North
MAGNOLIOPSIDA
[EUDICOTS]
Turkestan. Industrial, ornamental, afforestation.
Genus 226. Salix L.
804. Salix alba L.
Tree. Banks of rivers. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Industrial, ornamental, afforestation, tanniferous, medicinal, meliferous.
805. Salix blakii Goerz.
Tree or shrub. Banks of rivers. Foothills, montane zone. Nuratau, Malguzar, North Turkestan, Kyzylkum.
806. Salix olgae Regel
Shrub. Banks of rivers, pebbles. Plain, foothills, montane zone. Malguzar, North Turkestan.
807. Salix pycnostachya Andersson.
Tree or Shrub. Banks of rivers. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
808. Salix rosmarinifolia L.
Shrub. Wet places, river valleys. Montane zone. Nuratau.
809. Salix songarica Andersson
Tree or shrub. River valleys. Plain. Mirzachul. Industrial, ornamental, afforestation, tanniferous, dye,
meliferous.
810. Salix wilhelmsiana M. Bieb.
Shrub. River valleys. Plain, foothills, montane and alpine zones. North Turkestan, Mirzachul.
Industrial, ornamental, afforestation, tanniferous, meliferous.
Family 59. Euphorbiaceae
Genus 227. Andrachne L.
811. Andrachne fedtschenkoi Kossinsky
Semishrub. Stony slopes, rocks. Montane zone, Nuratau, Aktau, Malguzar.
812. Andrachne telephioides L.
Semishrub. Stony slopes, fields. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Medicinal, alkaloid-bearing, dye, fodder.
Genus 228. Chrozophora Neck.
813. Chrozophora tinctoria (L.) A. Juss. [Chrozophora hierosolymitana Spreng.; Chrozophora
obliqua (Vahl.) A. Juss. ex. Spreng.]
Annual. Sandy deserts, wastelands, fine-earth, gravelly and stony slopes, fields, fallow lands,
251
252
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
wastelands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Dye, poisonous.
814. Chrozophora sabulosa Kar. & Kir. [Chrozophora. gracilis Fisch. & C. A. Mey. ex Ledeb.]
Annual. Sandy deserts, banks of rivers, lakes and canals. Plain. Mirzachul, Kyzylkum. Dye, fodder.
Genus 229. Euphorbia L.
815. Euphorbia chamaesyce L. [Chamaesyce canescens (L.) Prokh.] Figure 258
Annual. Pebbles, river valleys, dry riverbeds, fine-earth and stony slopes, fields, wastelands.
Foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Weed.
Figure 258 Euphorbia chamaesyce (photography by Natalya Beshko)
816. Euphorbia cheirolepis Fisch. & C. A. Mey. ex Karelin [Cystidospermum cheirolepis (Fisch. &
C. A. Mey. ex Ledeb.) Prokh.]
Annual. Sandy deserts. Plain. Mirzachul, Kyzylkum. Fodder.
817. Euphorbia densa Schrenk
Annual. Sandy deserts, stony slopes. Plain, foothills. Nuratau, Nuratau Relic Mountains, Mirzachul,
Kyzylkum.
MAGNOLIOPSIDA
[EUDICOTS]
818. Euphorbia esula L. [Euphorbia jaxartica Prokh.; Euphorbia glomerulans (Prokh.)
Prokh.] Figure 259
Perennial. River valleys, banks of canals, wet stony slopes, screes, pebbles, fallow lands. Plain,
foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan. Dye, poisonous.
Figure 259 Euphorbia esula (photography by Natalya Beshko)
819. Euphorbia falcata L. Figure 260
Annual. Stony, fine-earth slopes, pebbles, fields. Foothills, montane and alpine zones. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, weed.
Figure 260 Euphorbia falcata (photography by Natalya Beshko)
820. Euphorbia franchetii B. Fedtsch.
Annual. Stony slopes, clay deserts, fields, fallow lands. Foothills, montane and alpine zones.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
253
254
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
821. Euphorbia granulata Forssk. [Chamaesyce turcomanica (Boiss.) Prokh.]
Annual. Gravelly slopes, pebbles, sandy and clay deserts, saline lands. Plains, foothills, montane
zone. Nuratau, Malguzar, North Turkestan, Mirzachul.
822. *Euphorbia helioscopia L. Figure 261
Annual. Stony slopes, roadsides, banks of canals, gardens, fallow lands, wastelands. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Poisonous, dye, weed.
Figure 261 Euphorbia helioscopia (photography by Natalya Beshko)
823. Euphorbia humilis C. A. Mey. ex Ledeb.
Perennial. Stony slopes, rocks, screes. Montane and alpine zones. Malguzar, North Turkestan.
824. Euphorbia inderiensis Less. ex Kar. & Kir. Figure 262
Annual. Fine-earth and stony slopes, sandy soils, saline lands. Plain, foothills, montane zone.
Nuratau, Malguzar, Mirzachul, Kyzylkum.
Figure 262 Euphorbia inderiensis (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
825. Euphorbia kanaorica Boiss. [Euphorbia polytimetica (Prokh.) Prokh.]
Perennial. Rocks, screes, fine-earth and stony slopes. Alpine zone. North Turkestan.
826. Euphorbia rapulum Kar. & Kir. Figure 263
Perennial. Fine-earth, gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Malguzar.
Medicinal, poisonous, rubber-bearing.
Figure 263 Euphorbia rapulum (photography by Natalya Beshko)
827. Euphorbia rosularis Fed.
Perennial. Stony slopes, screes. Montane and alpine zones, Malguzar, North Turkestan.
828. Euphorbia sarawschanica Regel Figure 264
Perennial. Stony slopes, rocks. Montane and alpine zones. Malguzar, North Turkestan. Medicinal,
poisonous.
Figure 264 Euphorbia sarawschanica (photography by Natalya Beshko)
829. Euphorbia sororia Schrenk
Annual. Clayey hills. Foothills. Malguzar.
830. Euphorbia szowitsii Fisch. & C. A. Mey.
Annual. Fine-earth, gravelly slopes, screes, pebbles. Montane zone. Nuratau, Malguzar, North
Turkestan.
255
256
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
831. Euphorbia turczaninowii Kar. & Kir. Figure 265
Annual. Sandy deserts. Plain. Kyzylkum.
Figure 265 Euphorbia turczaninowii (photography by Natalya Beshko)
Family 60. Linaceae
Genus 230. Linum L.
832. Linum corymbulosum Rchb.
Annual. gardens, fallow lands, river valleys, meadows, fine-earth, gravelly and stony slopes. Plain,
foothills, montane zone. Nuratau, Malguzar, North Turkestan.
833. Linum macrorhizum Juz.
Perennial. Fine-earth and stony slopes, screes, fallow lands. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan.
834. Linum olgae Juz.
Perennial. Fine-earth and stony slopes, rocks. Montane and alpine zones. North Turkestan.
Ornamental, oil.
835. *Linum usitatissimum L. [Linum humile Mill.]
Annual. Roadsides, gardens, fallow lands, fields. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Food, medicinal, industrial, oil.
MAGNOLIOPSIDA
[EUDICOTS]
Family 61. Geraniaceae
Genus 231. Geranium L.
836. Geranium collinum Stephan ex Willd. [Geranium regelii Nevski] Figure 266
Perennial. Meadows, river valleys, stream banks, wet places near springs, fine-earth and stony
slopes, rocks. Foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan. Medicinal,
dye, tanniferous, fodder, ornamental.
Figure 266 Geranium collinum (photography by Natalya Beshko)
837. Geranium divaricatum Ehrh. Figure 267
Annual. Shady wet places, fine-earth and stony slopes, screes, wastelands, gardens. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Tanniferous,
ornamental, weed.
257
258
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 267 Geranium divaricatum (photography by Natalya Beshko)
838. Geranium kotschyi subsp. charlesii (Aitch. & Hemsl.) P. H. Davis [Geranium charlesii
(Aitch. & Hemsl.) Vved. ex Nevski] Figure 268
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan.
Figure 268 Geranium kotschyi subsp. charlesii (photography by Natalya Beshko)
839. Geranium linearilobum DC. [Geranium transversale (Kar. & Kir.) Vved. ex Pavlov]
Perennial. Fine-earth, gravelly and stony slopes, piedmont plains. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental.
840. *Geranium pusillum L.
Annual. Fields, fallow lands, roadsides, gardens, wastelands, fine-earth and stony slopes. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
841. Geranium robertianum L.
Annual. Ravines, banks of rivers, shady wet places. Montane zone. Nuratau. Medicinal, ornamental.
842. Geranium rotundifolium L. Figure 269
Annual. Pebbles, fine-earth, gravelly slopes, shady places, gardens. Foothills, montane zone.
MAGNOLIOPSIDA
[EUDICOTS]
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, tanniferous.
Figure 269 Geranium rotundifolium (photography by Natalya Beshko)
843. Geranium saxatile Kar. & Kir. Figure 270
Perennial. Wet places, stony slopes, rocks. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental.
Figure 270 Geranium saxatile (photography by Natalya Beshko)
259
260
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 232. Erodium L’Her.
844. Erodium ciconium (Jusl.) L’Her.
Annual. Sandy and clay deserts, piedmont plains, clayey hills. Plain, foothills. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
845. Erodium cicutarium (L.) L’Her. Figure 271
Annual. Fine-earth, gravelly and stony slopes, river valleys, gardens, banks of canals, fallow
lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Medicinal, fodder, weed, meliferous.
Figure 271 Erodium cicutarium (photography by Natalya Beshko)
846. Erodium hoefftianum C. A. Mey.
Annual. Clayey and skeleton soils. Plain, foothills. Nuratau, Malguzar.
847. Erodium oxyrrhynchum M. Bieb.
Annual or biennial. Stony slopes, sandy soils, pebbles. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum.
MAGNOLIOPSIDA
[EUDICOTS]
Family 62. Lythraceae
Genus 233. Lythrum L.
848. Lythrum hyssopifolia L.
Annual. Saline lands, banks of rivers, lakes and canals. Plain, foothills, montane zone. Nuratau.
Medicinal, ornamental, meliferous.
849. Lythrum nanum Kar. & Kir.
Annual. Saline lands. Plain. Mirzachul.
Family 63. Onagraceae
Genus 234. Epilobium L.
850. Epilobium angustifolium L. [Chamaenerion angustifolium (L.) Scop.] Figure 272
Perennial. Wet fine-earth and stony slopes, river valleys, places near melting snow and springs.
Montane and alpine zones. Malguzar, North Turkestan. Medicinal, ornamental, meliferous.
Figure 272 Epilobium angustifolium (photography by Natalya Beshko)
851. Epilobium hirsutum L. (Epilobium velutinum Nevski) Figure 273
Perennial. Banks of rivers and canals, wet places, swamps, meadows. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal,
ornamental, fodder, meliferous.
261
262
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 273 Epilobium hirsutum (photography by Natalya Beshko)
852. Epilobium komarovii Ovcz.
Perennial. Banks of rivers, swamps, wet places near springs. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan.
853. Epilobium minutiflorum Hausskn. Figure 274
Perennial. Banks of rivers and canals, pebbles, wet places near springs, swamps, alpine meadows.
Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul.
Figure 274 Epilobium minutiflorum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
854. Epilobium palustre L.
Perennial. Meadows, banks of rivers, swamps. Plain, foothills, montane and alpine zones. North
Turkestan. Medicinal, fodder, meliferous.
855. Epilobium roseum subsp. subsessile (Boiss.) P. H. Raven [Epilobium nervosum Boiss.]
Perennial. Wet slopes, banks of rivers. Montane and alpine zones. Malguzar, North Turkestan.
856. Epilobium tetragonum L.
Perennial. Wet meadows, banks of rivers and canals, wet screes, swamps, saline lands, gardens,
fields, fallow lands. Plain, foothills, montane zone. Nuratau, North Turkestan. Medicinal, fodder,
meliferous.
857. Epilobium tianschanicum Pavlov
Perennial. Banks of rivers, wet places near springs, swamps, wet rocks and screes, alpine meadows.
Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
858. Epilobium turkestanicum Pazij & Vved.
Perennial. Wet stony slopes and rocks, wet places near melting snow, swamps, alpine meadows,
pebbles, banks of rivers. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Family 64. Biebersteiniaceae
Genus 235. Biebersteinia Steph.
859. Biebersteinia multifida DC. Figure 275
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Medicinal.
Figure 275 Biebersteinia multifida (photography by Natalya Beshko)
263
264
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 65. Nitrariaceae
Genus 236. Peganum L.
860. Peganum harmala L. Figure 276
Perennial. Saline lands, sandy and clay deserts, roadsides, surroundings of settlements, wells and
sheepyards, wastelands. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, weed, poisonous.
Figure 276 Peganum harmala (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Family 66. Anacaridaceae
Genus 237. Pistacia L.
861. Pistacia vera L. Figure 277
Tree or shrub. Fine-earth, gravelly and stony slopes, rocks, screes. Foothills, montane zone. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Food, medicinal, ornamental, industrial, oil,
dye, afforestation.
Figure 277 Pistacia vera (photography by Natalya Beshko)
Family 67. Sapindaceae
Genus 238. Acer L.
862. Acer pubescens Franch Figure 278
Tree. Fine-earth, gravelly and stony slopes, river valleys. Montane zone. Nuratau, Malguzar, North
265
266
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Turkestan. Ornamental, industrial, afforestation.
Figure 278 Acer pubescens (photography by Natalya Beshko)
863. Acer semenovii Regel & Herder Figure 279
Tree. Fine-earth, gravelly and stony slopes, river valleys. Montane zone. Nuratau, Aktau, Malguzar,
North Turkestan. Ornamental, industrial, afforestation.
Figure 279 Acer semenovii (photography by Natalya Beshko)
864. Acer turkestanicum Pax
Tree. Fine-earth, gravelly and stony slopes, river valleys. Montane zone. Malguzar, North Turkestan.
Ornamental, industrial, afforestation.
Family 68. Rutaceae
Genus 239. Haplophyllum Juss.
865. Haplophyllum acutifolium (DC.) G. Don. [Haplophyllum perforatum (M. Bieb.) Vved.]
Perennial. Sandy and clay deserts, fine-earth, gravelly and stony slopes, pebbles, rocks, fields, fallow
MAGNOLIOPSIDA
[EUDICOTS]
lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Medicinal, dye.
866. Haplophyllum bungei Trautv.
Perennial. Sandy and clay deserts. Plain. Kyzylkum. Medicinal, dye.
867. Haplophyllum lasianthum Bunge [Haplophyllum versicolor Fisch. & C. A. Mey.]
Perennial. Sandy and clay deserts, fine-earth, gravelly and stony slopes, pebbles, fallow lands, fields.
Plain, foothills. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, weed.
868. Haplophyllum latifolium Kar. & Kir. Figure 280
Perennial. Fine-earth, gravelly and stony slopes, fallow lands. Foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal.
Figure 280 Haplophyllum latifolium (photography by Natalya Beshko)
869. Haplophyllum pedicellatum Bunge ex Boiss. Figure 281
Perennial. Sandy and clay deserts, saline lands, fallow lands, fine-earth slopes. Plain, foothills.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal,
poisonous.
267
268
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 281 Haplophyllum pedicellatum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
870. Haplophyllum ramosissimum (Paulsen) Vved.
Semishrub. Sandy deserts. Plain. Kyzylkum.
871. Haplophyllum robustum Bunge
Perennial. Sandy and clay deserts. Plain. Mirzachul, Kyzylkum. Medicinal.
Genus 240. Dictamnus L.
872. Dictamnus angustifolius G. Don. f. & Sweet Figure 282
Perennial. Fine-earth, gravelly and stony slopes, river valleys, stream banks, screes, meadows.
Montane and alpine zones. Malguzar, North Turkestan. Poisonous, medicinal, ornamental,
meliferous.
Figure 282 Dictamnus angustifolius (photography by Natalya Beshko)
Family 69. *Simaroubaceae
Genus 241. *Ailanthus Desf.
873. *Ailantus altissima (Mill.) Swingle
Tree. Introduced alien species. Settlements, forest belts (planted), river valleys, fine-earth slopes
269
270
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
(naturalized). Plain, foothills, montane zone. Nuratau, Malguzar, North Turkestan, Mirzachul.
Ornamental, industrial, afforestation.
Family 70. Malvaceae
Genus 242. Malva L.
874.*Malva neglecta Wallr
Perennial. Fine-earth and skeleton soils, pebbles, wastelands, dry riverbeds. Plain, foothills, montane
and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, food, fodder, meliferous, weed.
Genus 243. Alcea L.
875. Alcea litvinovii (Iljin) Iljin Figure 283
Perennial. Fine-earth, gravelly slopes, river valleys. Plain, foothills, montane zone. Nuratau.
Medicinal, ornamental, meliferous.
Figure 283 Alcea litvinovii (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
876. Alcea nudiflora (Lindl.) Boiss. Figure 284
Perennial. Fine-earth, gravelly and stony slopes, screes, fallow lands, roadsides. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, ornamental,
meliferous.
Figure 284 Alcea nudiflora (photography by Natalya Beshko)
877. Alcea rhyticarpa (Trautv.) Iljin
Perennial. Sandy and clayey soils, fine-earth, gravelly and stony slopes. Plain, foothills, montane
zone. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Medicinal, ornamental, meliferous.
Genus 244. Althaea L.
878. Althaea armeniaca Ten.
Perennial. River valleys, banks of canals, fields. Plain, foothills. Nuratau, Mirzachul. Medicinal,
ornamental, meliferous.
879. Althaea officinalis L.
Perennial. River valleys, fields, fallow lands. Plain, foothills, montane zone. North Turkestan.
Medicinal, ornamental, meliferous.
Genus 245. *Abutilon Mill.
880. *Abutilon therophrasti Medik
Annual. River valleys, fallow lands, banks of canals, roadsides, fields, gardens. Plain, foothills,
271
272
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, oil,
fiber, weed.
Genus 246. *Hibiscus L.
881. *Hibiscus trionum L.
Annual. Banks of canals, fallow lands, fields. Plain. Mirzachul. Medicinal, oil, fiber, weed,
meliferous.
Family 71. Thymelaceae
Genus 247. Thymelaea Mill.
882. Thymelaea passerina (L.) Coss. & Germ.
Annual. Fine-earth and stony slopes, banks of rivers and canals, fields, fallow lands, wastelands.
Montane zone. Nuratau, Malguzar, North Turkestan. Weed, poisonous.
Genus 248. Diarthron Turcz.
883. Diarthron arenaria (Pobed.) Kit Tan [Dendrostellera arenaria Pobed.]
Shrub. Sandy deserts. Plain. Kyzylkum. Fodder, industrial, afforestation.
884. Diarthron vesiculosum (Fisch. & C. A. Mey.) C. A. Mey.
Annual. Stony slopes, fallow lands, fields, piedmont plains, sandy and clay deserts. Plain, foothills,
montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed, poisonous.
Family 72. Cistaceae
Genus 249. Helianthemum Adans.
885. Helianthemum songaricum Schrenk ex Fisch. & C. A. Mey.
Semishrub. Stony slopes, rocks, screes, pebbles, banks of rivers. Foothills, montane and alpine
zones. North Turkestan. Dye, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Family 73. *Resedaceae
Genus 250. *Reseda L.
886. *Reseda lutea L.
Biennial or perennial. Fine-earth, gravelly and stony slopes, fields, fallow lands, dry riverbeds,
banks of rivers, roadsides, wastelands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Dye, oil, weed.
887. *Reseda luteola L.
Biennial. Fine-earth slopes, fields, roadsides, fallow lands, wastelands. Plain, foothills, montane
zone. Aktau, Malguzar, North Turkestan. Dye, oil, fodder, weed.
Family 74. Capparaceae
Genus 251. Capparis L.
888. Capparis spinosa L.
Perennial. Sandy, clayey, gravelly soils, banks of rivers, lakes and canals, fallow lands, screes,
wastelands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Medicinal, food, dye, fodder, ornamental, meliferous,
weed.
Family 75. Cleomaceae
Genus 252. Cleome L.
889. Cleome fimbriata Vicary
Annual. Gypsiferous grey soils, limestones, pebbles, dry riverbeds, fallow lands. Foothills. Nuratau,
Aktau.
273
274
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 76. Brassicaceae [Cruciferae]
Genus 253. Alliaria Scop.
890. Alliaria petiolata (M. Bieb.) Cavara & Grande
Biennial. Fine-earth and stony slopes, ravines, river valleys, shady wet places, gardens. Montane and
alpine zones, Nuratau, Malguzar, North Turkestan. Medicinal, food.
Genus 254. Sisymbrium L.
891. *Sisymbrium altissimum L.
Annual or biennial. Saline lands, wastelands, fallow lands, wastelands, stony slopes, river valleys.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Food, oil, weed.
892. Sisymbrium brassiciforme C. A. Mey.
Biennial. Stony and gravelly slopes, rocks, screes, dry riverbeds, moraines. Foothills, montane and
alpine zones. Nuratau, Malguzar, North Turkestan.
893. Sisymbrium loeselii L.
Annual. Fine-earth slopes, pebbles, meadows, river valleys, settlements, gardens, fields, banks of
canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Food, fodder, oil, weed, meliferous.
Genus 255. Neotorularia Hedge & J. Leonard
894. Neotorularia torulosa (Desf.) Hedge & J. Leonard [Torularia torulosa (Desf.) O. E. Schulz.]
Annual. Roadsides, fields, fallow lands, wastelands, saline lands, sandy deserts. Plain. Mirzachul,
Kyzylkum. Fodder, weed.
Genus 256. Schrenkiella D. A. German & Al-Shehbaz
895. Schrenkiella parvula (Schrenk) D.A. German & Al-Shehbaz [Arabidopsis parvula
(Schrenk) O. E. Schulz.]
Annual. Clayey soils, saline lands. Plain. Mirzachul, North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 257. Olimarabidopsis Al-Shehbaz, O’Kane & Price
896. Olimarabidopsis pumila (Stephan) Al-Shehbaz, O’Kane & R. A. Price [Arabidopsis
pumila (Steph.) N. Busch]
Annual. Clayey soils, hills, limestones, saline lands, banks of canals, gardens, stony slopes,
ravines. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum.
897. Olimarabidopsis umbrosa (Botsch. & Vved.) Al-Shehbaz, O’Kane & R. A. Price [Trichochiton
umbrosum (Kom.) Botsch. & Vved.]
Annual. Shady wet places, stream banks, screes, rocks. Montane and alpine zones. North Turkestan.
Genus 258. Arabidopsis Heynh.
898. Arabidopsis thaliana (L.) Heynh.
Annual or biennial. Pebbles, banks of rivers, shady places, rocks. Plain, foothills, montane zone.
Malguzar, North Turkestan. Food, weed.
Genus 259. Descurainia Webb & Berth.
899. Descurainia sophia (L.) Webb. & Prantl
Annual. Wastelands, settlements, banks of canals, ravines, fields, fallow lands, gardens, saline lands,
roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Weed.
Genus 260. Smelowskia C. A. Mey. ex Ledeb.
900. Smelowskia sisymbrioides (Regel & Herder) Lipsky ex Paulsen [Sophiopsis sisymbrioides
(Regel & Herder) O. E. Schulz]
Biennial. Stony and gravelly slopes, rocks, ravines. Montane zone, Malguzar, North Turkestan.
Genus 261. Erysimum L.
901. Erysimum cyaneum Popov
Biennial. Stony slopes. Montane zone. Malguzar. Poisonous.
902. Erysimum marschallianum Andrz. ex DC.
Biennial. Gardens, fallow lands, fine-earth slopes. Foothills, montane and alpine zones. North
Turkestan. Poisonous.
275
276
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
903. Erysimum violascens Popov [Erysimum nuratense Popovex Botsch. & Vved.]
Biennial. Stony slopes. Montane zone. Nuratau. Medicinal, poisonous.
Genus 262. Barbarea Beck.
904. Barbarea brachycarpa subsp. minor (K. Koch) Parolly & Eren [Barbarea minor C. Koch]
Perennial. Wet stony slopes, meadows, banks of rivers. Alpine zone. North Turkestan.
905. Barbarea vulgaris R. Br.
Biennial. Wet places, banks of rivers and canals, pebbles, meadows, fields, gardens. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, food, fodder, dye, meliferous, weed.
Genus 263. Rorippa Scop.
906. Rorippa palustris (L.) Besser
Annual or biennial. Swamp meadows, banks of rivers, lakes and canals. Plain. Mirzachul. Food, weed.
Genus 264. Nasturtium R. Br.
907. Nasturtium officinale W. T. Aiton [Nasturtium fontanum (Lam.) Aschers.]
Perennial. Swamps, banks of rivers and canals. Plain, foothills. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, food, fodder, meliferous.
Genus 265. Turristis L.
908. Turristis glabra L.
Annual. Fine-earth, gravelly and stony slopes, fallow lands, fields, roadsides, settlements. Foothills,
montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Fodder, weed.
Genus 266. Scapiarabis M. A. Koch, R. Karl, D. A. German & Al-Shehbaz
909. Scapiarabis saxicola (Edgew) M. A. Koch, R. Karl, D. A. German & Al. Shehbaz [Arabis
kokanica Regel & Schmalh.]
Perennial. Stony slopes, rocks. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 267. Arabis L.
910. Arabis auriculata Lam
Annual or biennial. Stony slopes, rocks. Foothills, montane zone. Nuratau, Malguzar. Fodder.
911. Arabis montbretiana Boiss.
Annual, biennial. Stony slopes, rocks. Foothills, montane zone. Nuratau, Aktau, Malguzar. Fodder.
Genus 268. Isatis L.
912. Isatis brevipes (Bunge) Jafri [Pachypterygium brevipes Bunge]
Annual. Stony slopes. Foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan.
913. Isatis emarginata Kar. & Kir. [Isatis violascens Bunge]
Annual. Sandy deserts. Plain. Kyzylkum.
914. Isatis minima Bunge
Annual. Sandy deserts. Plain. Kyzylkum.
915. Isatis multicaulis (Kar. & Kir.) Jafri [Pachypterygium densiflorum Bunge]
Annual. Stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan.
916. *Isatis tinctoria L.
Biennial. River valleys, pebbles, fallow lands, fields, surroundings of settlements (escaped from
culture). Plain, foothills. Nuratau, Malguzar, North Turkestan. Fodder, dye, meliferous.
Genus 269. Tauscheria Fisch.
917. Tauscheria lasiocarpa Fisch. & DC.
Annual. Sandy and clay deserts, fine-earth slopes, banks of rivers, lakes and canals. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Weed.
Genus 270. Goldbachia DC.
918. Goldbachia laevigata (M. Bieb.) DC.
Annual. Sandy deserts, saline lands, fine-earth, gravelly and stony slopes. Plain, foothills. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, oil.
919. Goldbachia sabulosa (Kar. & Kir.) D. A. German & Al-Shehbaz [Spirorrhynchus
sabulosus Kar. & Kir.]
Annual. Sandy desrts. Plain. Kyzylkum. Fodder.
277
278
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
920. Goldbachia torulosa DC.
Annual. Wastelands, fallow lands, fine-earth, gravelly and stony slopes. Foothills. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, oil, weed.
921. Goldbachia verrucosa Kom.
Annual. Fine-earth, gravelly slopes. Foothills, montane zone. Nuratau. Fodder, oil, weed.
Genus 271. Hesperis L.
922. Hesperis sibirica L. Figure 285
Biennial or perennial. Shady wet places, banks of rivers, subalpine and alpine meadows. Montane
and alpine zones. Malguzar, North Turkestan. Fodder, essential oil, ornamental, meliferous.
Figure 285 Hesperis sibirica (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 272. Parrya R. Br.
923. Parrya fruticulosa Regel & Schmalh. [Neuroloma fruticulosum (Regel & Schmalh.)
Botsch.]
Semishrub. Stony slopes, rocks, screes. Montane and alpine zones. Nuratau, Malguzar.
924. Parrya hispida (Regel) D. A. German & Al. Shehbaz [Pseudoclausia hispida (Regel)
Popov ex A.V. Vassil.]
Biennial. Stony slopes, rocks, screes, pebbles. Foothills, montane and alpine zones. Nuratau, Nuratau
Relic Mountains, Malguzar, North Turkestan.
925. Parrya khorasanica (Rech. f. & Aellen) D. A. German & Al. Shehbaz [Pseudoclausia
turkestanica (Lipsky) A.V. Vassil.] Figure 286
Biennial. Fine-earth and stony slopes, shady places, screes, pebbles, banks of rivers, subalpine
meadows. Foothills, montane and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan.
Figure 286 Parrya khorasanica (photography by Natalya Beshko)
926. Parrya nuratensis Botsch. & Vved. [Neuroloma nuratense (Botsch. & Vved.) Botsch.] Figure 287
Semishrub. Fine-earth and stony slopes, rocks. Montane zone. Nuratau. Endemic to the Nuratau
Mountains.
279
280
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 287 Parrya nuratensis (photography by Natalya Beshko)
927. Parrya olgae (Regel & Schmalh.) D. A. German & Al. Shehbaz [Pseudoclausia olgae (Regel &
Schmalh.) Botsch.] Figure 288
Biennial. Rocks, Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. UzbRDB 2.
Figure 288 Parrya olgae (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
928. Parrya sarawschanica (Regel & Schmalh.) D. A. German & Al-Shehbaz [Pseudoclausia
sarawschanica (Regel & Schmalh.) Botsch.]
Biennial. Stony slopes. Montane zone. Nuratau. UzbRDB 2.
Genus 273. Leiospora (C. A. Mey.) Dvorak.
929. Leiospora subscapigera Botsch & Pachom.
Perennial. Moraines, screes, gravelly slopes. Alpine zone. North Turkestan.
Genus 274. Strigosella Boiss.
930. *Strigosella africana (L.) Botsch.
Annual. Sandy and clay deserts, saline lands, ravines, fine-earth and stony slopes, screes, wastelands,
fields, fallow lands, gardens. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Poisonous, weed.
931. Strigosella brevipes (Bunge) Botsch.
Annual. Sandy and clay deserts, fallow lands. Plain. Kyzylkum.
932. Strigosella grandiflora (Bunge) Botsch.
Annual. Sandy soils, fine-earth and stony slopes, fields. Plain, foothills. Nuratau, Kyzylkum.
933. Strigosella intermedia (C. A. Mey.) Botsch.
Annual. Banks of canals, gravelly slopes, fallow lands, saline lands, sandy and clay deserts.
Foothills. Nuratau, Mirzachul, Kyzykum.
934. Strigosella scorpioides (Bunge) Botsch.
Annual. Sandy and clay deserts, dry riverbeds, saline lands, pebbles, fine-earth and stony slopes.
Plain, foothills, montane zone. Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum.
935. Strigosella stenopetala (Fisch. & C. A. Mey.) Botsch.
Annual. Sandy and clay deserts, saline lands, fields, banks of canals. Plain, foothills. Nuratau Relic
Mountains, Mirzachul, Kyzylkum. Weed.
936. Strigosella trichocarpa (Boiss. & Buhse) Botsch.
Annual. Pebbles, Sandy and clay deserts, fine-earth, gravelly and stony slopes, ravines, fields. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed.
937. Strigosella turkestanica (Litv.) Botsch. Figure 289
Annual. Sandy and clay deserts, fine-earth and stony slopes, rocks, pebbles, fields, fallow lands,
dry riverbeds. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
Mirzachul, Kyzylkum. Fodder, weed.
281
282
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 289 Strigosella turkestanica (photography by Natalya Beshko)
Genus 275. Cryptospora Kar. & Kir.
938. Cryptospora falcata Kar. & Kir. [Cryptospora omissa Botsch.]
Annual. Sandy and clay deserts, roadsides, fine-earth and stony slopes, fields, fallow lands. Plain,
foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum.
939. Cryptospora inconspicua (Kom.) O. E. Schulz [Trichochiton inconspicuum Kom.]
Annual. Shady wet places, rocks, screes, pebbles, wet fine-earth and stony slopes, subalpine
meadows, stream banks. Montane and alpine zones. North Turkestan. Fodder.
Genus 276. Matthiola R. Br.
940. Matthiola chorassanica Bunge ex Boiss. [Matthiola integrifolia Kom.]
Perennial. Stony slopes. Montane and alpine zones. Nuratau, Aktau, North Turkestan.
941. Matthiola obovata Bunge
Perennial. Stony and gravelly slopes, rocks. Montane zone. Nuratau, Malguzar.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 277. Tetracme Bunge
942. Tetracme quadricornis (Steph. ex Willd.) Bunge
Annual. Sandy deserts, saline lands, banks of lakes and canals, fallow lands. Plain. Mirzachul,
Kyzylkum. Weed.
943. Tetracme recurvata Bunge
Annual. Sandy and clay deserts, saline lands, dry riverbeds, banks of lakes and canals, fallow lands.
Plain, foothills. Mirzachul, Kyzylkum.
Genus 278. Leptaleum DC.
944. Leptaleum filifolium (Willd.) DC.
Annual. Saline lands, Sandy and clay deserts, banks of rivers, lakes and canals, stony slopes, rocks,
settlements, roadsides, fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Genus 279. Streptoloma Bunge
945. Streptoloma desertorum Bunge
Annual. Sandy deserts, saline lands. Plain, foothills, montane zone. Kyzylkum.
Genus 280. Diptychocarpus Trautv.
946. Diptychocarpus strictus (Fisch. ex M. Bieb.) Trautv.
Annual. Gravelly, Fine-earth slopes, rocks, dry riverbeds, banks of rivers and canals, sandy deserts,
fallow lands, saline lands, fields, gardens. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Mirzachul, Kyzylkum.
Genus 281. Chorispora R. Br.
947. Chorispora bungeana Fisch. & C. A. Mey.
Perennial. Pebbles, screes, stony and gravelly slopes, rocks, moraines, alpine meadows. Alpine zone.
North Turkestan.
948. Chorispora macropoda Trautv.
Perennial. Fine-earth and stony slopes, banks of rivers, pebbles, wet places, screes, rocks, moraines,
subalpine and alpine meadows. Montane and alpine zones. North Turkestan.
283
284
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
949. Chorispora sabulosa Cambess. [Chorispora elegans Cambess.]
Perennial. Alpine meadows, gravelly and stony slopes. Alpine zone. North Turkestan.
950. Chorispora tenella (Pall.) DC.
Annual. Sandy and clay deserts, saline lands, fine-earth, gravelly and stony slopes, banks of rivers,
moraines, fields, fallow lands, dry riverbeds, pebbles, roadsides. Plain, foothills, montane and alpine
zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Fodder, weed.
Genus 282. *Euclidium R. Br.
951. *Euclidium syriacum (L.) R. Br.
Annual. Saline lands, sandy and clay deserts, ravines, banks of rivers, lakes and canals, fine-earth and stony
slopes, shady places, roadsides, fallow lands, fields, wastelands. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Genus 283. Litwinowia Woron.
952. Litwinowia tenuissima (Pall.) Woronow ex Pavlov
Annual. Saline lands, sandy and clay deserts, pebbles, fine-earth, gravelly and stony slopes, rocks,
screes, meadows, settlements, roadsides, banks of canals, gardens, wastelands. Plain, foothills,
montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed.
Genus 284. Octoceras Bunge
953. Octoceras lehmannianum Bunge
Annual. Sandy desert, saline lands, wastelands. Plain. Kyzylkum.
Genus 285. Cithareloma Bunge
954. Cithareloma lehmannii Bunge
Annual. Sandy deserts. Plain. Kyzylkum. Fodder.
Genus 286. Lachnoloma Bunge
955. Lachnoloma lehmanii Bunge
Annual. Sandy and clay deserts, saline lands. Plain. Mirzachul, Kyzylkum.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 287. Alyssum L.
956. Alyssum alyssoides (L.) L. [Alyssum campestre L.] Figure 290
Annual. Fine-earth and stony slopes, piedmont plains, sandy and clay deserts, roadsides, fallow
lands, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Fodder, weed.
Figure 290 Alyssum alyssoides (photography by Natalya Beshko)
957. Alyssum dasycarpum Stephan ex Willd. Figure 291
Annual. Sandy, clayey and skeleton soils. Plain, foothills. Nuratau, North Turkestan, Kyzylkum.
Fodder.
Figure 291 Alyssum dasycarpum (photography by Natalya Beshko)
285
286
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
958. Alyssum desertorum Stapf.
Annual. Sandy and clay deserts, saline lands, banks of rivers, fine-earth and stony slopes, fallow
lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Fodder.
959. Alyssum linifolium Stephan ex Willd. [Meniocus linifolius (Stephan ex Willd.) DC.]
Annual. Pebbles, gravelly and stony slopes, sandy and clay deserts, dry riverbeds. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Fodder.
960. Alyssum szovitsianum Fisch. & C. A. Mey. [Alyssum marginatum Steud.]
Annual. Fine-earth and stony slopes, rocks. Plain, foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Fodder.
Genus 288. Clypeola L.
961. Clypeola jonthlaspi L.
Annual. Pebbles, stony slopes, rocks. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan.
Genus 289. Asperuginoides Rauschert
962. Asperuginoides axillaris [Boiss. & Hohen.] Rauschert [Buchingera axillaris Boiss.]
Annual. Stony and gravelly slopes, shady wet places. Montane zone. Nuratau, Malguzar, North
Turkestan.
Genus 290. Draba L.
963. Draba lanceolata Royle
Perennial. Fine-earth and stony slopes, screes, rocks, subalpine meadows, moraines, wet places near
melting snow. Montane and alpine zones. North Turkestan.
964. Draba melanopus Kom
Biennial or perennial. Fine-earth and stony slopes, screes, rocks, alpine meadows, wet places near
melting snow. Montane and alpine zones. North Turkestan.
965. Draba nemorosa L.
Annual. Fine-earth, gravelly and stony slopes, pebbles, banks of rivers, subalpine meadows, fallow
lands, wet places. Montane zone. Nuratau, Malguzar.
966. Draba nuda (Bel.) Al. Shehbaz & M. Koch [Drabopsis nuda (Bel.) Stapf]
Annual. Stony and fine-earth soils, pebbles. Plain, foothills, montane zone. Nuratau, Aktau,
Malguzar, North Turkestan. Fodder.
MAGNOLIOPSIDA
[EUDICOTS]
967. Draba stenocarpa Hook. f. & Thomson
Annual. Fine-earth, gravelly and stony slopes, subalpine and alpine meadows, stream banks.
Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Genus 291. Erophila DC.
968. Erophila minima C. A. Mey.
Annual. Fine-earth and stony slopes, fallow lands, fields. Foothills. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Fodder.
969. Erophila verna (L.) DC.
Annual. Fine-earth, gravelly and stony slopes, sandy and clay deserts, fallow lands, wastelands.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Fodder.
Genus 292. Brassica L.
970. *Brassica rapa L. [Brassica campestris L.]
Annual. Fields, fallow lands, wastelands, gardens, banks of rivers and canals. Plain, foothills,
montane zone. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Oil, fodder, weed, meliferous.
971. Brassica elongata Ehrh.
Perennial. Fallow lands, fields, roadsides, wastelands. Plain, foothills, montane zone. Nuratau. Oil,
weed.
Genus 293. *Sinapis L.
972. *Sinapis alba L.
Annual. Roadsides, fields, gardens, settlements. Plain, foothills. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Food, medicinal, fodder, oil, weed, meliferous.
973. *Sinapis arvensis L.
Annual. Roadsides, wastelands, fields, gardens, settlements. Plain, foothills. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Food, oil, weed, meliferous.
Genus 294. *Eruca Hill
974. *Eruca vesicaria (L.) Cav. [Eruca sativa Mill.]
Annual. Fields, fallow lands, roadsides, sandy deserts, saline lands, stony slopes. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Oil, food, medicinal, weed.
287
288
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 295. Crambe L.
975. Crambe cordifolia subsp. kotschyana (Boiss.) Jafri [Crambe kotschyana Boiss.]
Perennial. Fine-earth, gravelly and stony slopes, fallow lands. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder, food, meliferous.
Genus 296. Conringia Heist. ex Fabr.
976. Conringia clavata Boiss. Figure 292
Annual. Stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan. Fodder.
Figure 292 Conringia clavata (photography by Natalya Beshko)
977. Conringia orientalis (L.) Dumort.
Annual. Fields, fallow lands, roadsides, stony sopes, rocks. Plain, foothills, montane zone. North
Turkestan.
978. Conringia persica Boiss.
Annual. Stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan. Fodder.
979. Conringia planisiliqua Fisch. & C. A. Mey.
Annual. Fine-earth slopes, river valleys, dry riverbeds, roadsides. Foothills, montane zone. North
Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 297. Chalcanthus Boiss.
980. Chalcanthus renifolius [Boiss. & Hohen.] Boiss. Figure 293
Perennial. Fine-earth and stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
Figure 293 Chalcanthus renifolius (photography by Natalya Beshko)
Genus 298. Lepidium L.
981. Lepidium botschantsevianum Al-Shehbaz [Stroganowia angustifolia Botsch. & Vved.] Figure 294
Perennial. Stony slopes, ravines, rocks. Foothills, montane zone. Nuratau.
Figure 294 Lepidium botschantsevianum (photography by Natalya Beshko)
289
290
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
982. Lepidium draba subsp. chalepense L. (P. Fourn.) [Cardaria repens (Schrenk) Jarm.]
Perennial. Wet places, fallow lands, fields, wastelands, roadsides, river valleys, banks of lakes and
canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Meliferous, weed.
983. Lepidium ferganense Korsh.
Perennial. Stony slopes, rocks, banks of rivers. Foothills, montane zone. Nuratau, Nuratau Relic
Mountains.
984. Lepidium lacerum C. A. Mey.
Perennial. Fine-earth and stony slopes, rocks, banks of rivers. Foothills, montane zone. Nuratau.
985. Lepidium latifolium L.
Perennial. Saline lands, swamps, meadows, banks of rivers and canals, pebbles, stony slopes, fields,
settlements. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul. Medicinal, meliferous, insecticide, weed.
986. Lepidium lipskyi (N. Busch) Al-Shehbaz & Mummenhoff [Stubendorffia lipskyi N. Busch]
Perennial. Fine-earth and stony slopes. Montane zone. Nuratau.
987. Lepidium olgae (R. Vinogradova) Al-Shehbaz & Mummenhoff [Stubendorffia olgae R.
Vinogradova] Figure 295
Perennial. Stony slopes. Montane zone. Nuratau. Medicinal. Threatened species, UzbRDB 2.
Endemic to the Nuratau ridge.
Figure 295 Lepidium olgae (photography by N. Yu. Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
988. Lepidium orientale (Schrenk) Al-Shehbaz & Mummenhoff [Stubendorffia orientalis
Schrenk] Figure 296
Perennial. Fine-earth and stony slopes, rocks, river valleys. Montane zone. Nuratau. Meliferous.
Figure 296 Lepidium orientale (photography by Natalya Beshko)
989. Lepidium perfoliatum L.
Annual or biennial. River valleys, saline lands, fields, roadsides, wastelands. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Medicinal, poisonous, weed.
Genus 299. Megacarpaea DC.
990. Megacarpaea gigantea Regel
Perennial. Fine-earth and stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
Fodder, meliferous.
991. Megacarpaea orbiculata B. Fedtsch.
Perennial. Fine-earth and stony slopes, rocks, ravines, banks of rivers, subalpine meadows. Montane
zone. Nuratau, Malguzar, North Turkestan. Fodder, meliferous.
Genus 300. Hornungia Rchb.
992. Hornungia procumbens (L.) Hayek [Hymenolobus procumbens (L.) Nutt.]
Annual. Saline lands, banks of rivers and canals, gardens, fallow lands. Plain, foothills, montane
zone. Nuratau, Malguzar, North Turkestan, Mirzachul.
291
292
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 301. Iberidella Boiss.
993. Iberidella trinervia (DC.) Boiss. Figure 297
Perennial. Gravelly and stony slopes, rocks. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan.
Figure 297 Iberidella trinervia (photography by Natalya Beshko)
Genus 302. Didymophysa Boiss.
994. Didymophysa fedtschenkoana Regel
Perennial. Screes, pebbles, moraines, stony slopes. Alpine zone. North Turkestan.
Genus 303. Thlaspi L.
995. *Thlaspi arvense L.
Annual. Fine-earth and stony slopes, banks of rivers, wet places, fields, fallow lands, roadsides, dry
riverbeds, river valleys. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Medicinal, oil, weed.
996. Thlaspi perfoliatum L. [Microthlaspi perfoliatum (L.) F. K. Mey.]
Annual. Fine-earth and stony slopes, pebbles, rocks, saline lands, settlements, fields. Plain, foothills,
MAGNOLIOPSIDA
[EUDICOTS]
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Fodder.
Genus 304. Neurotropis (DC.) F. K. Mey.
997. Neurotropis platycarpa (Fisch. & C. A. Mey.) F. K. Mey. [Thlaspi kotschyanum Boiss. &
Hohen.]
Annual. Gravelly and stony slopes, screes. Montane zone. Malguzar, North Turkestan.
Genus 305. Aethionema W. T. Aiton
998. Aethionema carneum (Banks & Soland.) B. Fedtsch. [Campyloptera carnea (Banks &
Soland.) Botsch. & Vved.] Figure 298
Annual. Fine-earth and stony slopes, screes, pebbles, river valleys, dry riverbeds. Foothills, montane
zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Figure 298 Aethionema carneum (photography by Natalya Beshko)
293
294
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 306. Camelina Crantz.
999. Camelina microcarpa Andrz. ex DC. [Camelina sylvestris Wallr.]
Annual. Fine-earth and stony slopes, banks of rivers, lakes and canals, fields, fallow lands, roadsides. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum. Oil, weed.
1000. Camelina rumelica Velen
Annual. Fine-earth and stony slopes, ravines, banks of rivers, wastelands, fallow lands. Plain,
foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, Mirzachul. Oil, weed.
Genus 307. Neslia Desv.
1001. Neslia paniculata subsp. thracica (Velen.) Bornm. [Neslia apiculata Fisch. & C. A. Mey.]
Annual. Fine-earth and stony slopes, ravines, pebbles, saline lands, fields, fallow lands. Foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Dye, weed.
Genus 308. *Capsella Medik.
1002. *Capsella bursa-pastoris (L.) Medik.
Annual. Wastelands, fields, gardens, roadsides, fine-earth and stony slopes, pebbles, banks of rivers.
Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Medicinal.
Family 77. Santalaceae
Genus 309. Arceuthobium M. Bieb.
1003. Arceuthobium oxycedri (DC.) M. Bieb.
Shrub. Parasite of juniper. Montane zone. Malguzar, North Turkestan. Medicinal.
Family 78. Frankeniaceae
Genus 310. Frankenia L.
1004. Frankenia bucharica Basil.
Semishrub. Saline lands, banks of rivers, lakes and canals, fallow lands. Plain. Mirzachul.
MAGNOLIOPSIDA
[EUDICOTS]
1005. Frankenia bucharica subsp. vvedenskyi (Botsch.) Chrtek. [Frankenia vvedenskyi Botsch.]
Semishrub. Saline lands, banks of salt lakes. Plain. Mirzachul, Kyzylkum.
1006. Frankenia hirsuta L.
Semishrub. Saline lands, banks of rivers, lakes and canals. Plain. Kyzylkum.
1007. Frankenia pulverulenta L.
Annual. Saline lands, fallow lands, sandy deserts. Plain. Mirzachul, Kyzylkum.
Family 79. Tamaricaceae
Genus 311. Reaumuria L.
1008. Reaumuria alternifolia (Labill.) Britten [Reaumuria reflexa Lipsky; Reaumuria sogdiana
Kom.; Reaumuria turkestanica Gorschk.]
Semishrub. Saline and gypsiferous soils, sandy desert. Plain, foothills, montane zone. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
1009. Reaumuria fruticosa Boiss.
Shrub. Saline lands, sandy deserts. Plain. Kyzylkum.
Genus 312. Tamarix L.
1010. Tamarix androssowii var. transcaucassica (Bunge) Qaiser [Tamarix litvinowii Gorschk.]
Shrub. Banks of rivers, lakes and canals, saline lands, dry riverbeds. Plain. Mirzachul, Kyzylkum.
Meliferous, ornamental, afforestation, tanniferous.
1011. Tamarix arceuthoides Bunge Figure 299
Shrub. River valleys, pebbles. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Meliferous, ornamental, afforestation.
Figure 299 Tamarix arceuthoides (photography by Natalya Beshko)
295
296
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1012. Tamarix elongata Ledeb.
Shrub. Saline lands, depressions, sandy deserts, dry riverbeds. Plain. Mirzachul, Kyzylkum.
Meliferous, ornamental, afforestation, dye.
1013. Tamarix hispida Willd. Figure 300
Shrub. Saline lands, depressions, banks of rivers, lakes and canals. Plain. Mirzachul, Kyzylkum.
Meliferous, ornamental, afforestation, dye.
Figure 300 Tamarix hispida (photography by Natalya Beshko)
1014. Tamarix laxa Willd. Figure 301
Shrub. Saline lands, banks of rivers, lakes and canals. Plain. Mirzachul, Kyzylkum. Meliferous,
ornamental, afforestation.
Figure 301 Tamarix laxa (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1015. Tamarix ramosissima Ledeb.
Shrub. Banks of rivers, lakes and canals, saline lands, sandy and clay deserts, fallow lands. Plain.
Mirzachul, Kyzylkum. Meliferous, ornamental, afforestation, dye.
1016. Tamarix smyrnensis Bunge [Tamarix hohenackeri Bunge]
Shrub. Banks of rivers. Plain. Mirzachul, Kyzylkum. Meliferous, ornamental, afforestation.
Genus 313. Myricaria Desv.
1017. Myricaria germanica (L.) Desv. [Myricaria bracteata Royle]
Shrub. Banks of rivers, ravines, pebbles. Foothills, montane zone. North Turkestan. Dye, tanniferous,
ornamental.
Family 80. Plumbaginaceae
Genus 314. Acantholimon Boiss.
1018. Acantholimon erythraeum Bunge Figure 302
Semishrub. Stony slopes. Montane and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan.
Figure 302 Acantholimon erythraeum (photography by Natalya Beshko)
297
298
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1019. Acantholimon nuratavicum Zakirov ex Lincz. Figure 303
Semishrub. Stony slopes. Montane zone. Nuratau. UzbRDB 2. Endemic to the Nuratau ridge.
Figure 303 Acantholimon nuratavicum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1020. Acantholimon subavenaceum Lincz. Figure 304
Semishrub. Stony slopes. Montane zone. Nuratau.
Figure 304 Acantholimon subavenaceum (photography by Natalya Beshko)
1021. Acantholimon tataricum Boiss. ex DC. Figure 305
Semishrub. Stony slopes, rocks. Montane and alpine zones. Nuratau, Aktau, Malguzar, North
Turkestan. Endemic to the Nuratau ridge.
Figure 305 Acantholimon tataricum (photography by Natalya Beshko)
1022. Acantolimon zakirovii Beshko
Semishrub. Stony slopes. Montane zone. Nuratau. Endemic to the Nuratau ridge.
299
300
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 315. Chaetolimon (Bunge) Lincz.
1023. Chaetolimon setiferum (Bunge) Lincz. Figure 306
Perennial. Fine-earth and stony slopes, piedmont plains. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan.
Figure 306 Chaetolimon setiferum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 316. Limonium Mill.
1024. Limonium otolepis (Schrenk) Kuntze Figure 307
Perennial. Saline lands, banks of rivers, lakes and canals. Plain. Nuratau Relic Mountains, Mirzachul,
Kyzylkum. Tanniferous, dye, meliferous, ornamental.
Figure 307 Limonium otolepis(photography by Natalya Beshko)
301
302
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1025. Limonium reniforme (Girard) Lincz.
Perennial. Saline lands, banks of rivers, lakes and canals. Plain. Mirzachul. Tanniferous, weed.
1026. Limonium suffruticosum (L.) Kuntze Figure 308
Semishrub. Saline lands. Plain. Mirzachul. Tanniferous, dye, meliferous, fodder.
Figure 308 Limonium suffruticosum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 317. Eremolimon Lincz.
1027. Eremolimon sogdianum (Popov) Lincz.
Perennial. Saline lands. Plain. Mirzachul. Dye.
Genus 318. Psylliostachys (Jaub. & Spach) Nevski
1028. Psylliostachys leptostachya (Boiss.) Roshkova
Annual. Saline lands, fallow lands, fields. Plain, foothills. North Turkestan, Mirzachul. Dye.
1029. Psylliostachys × myosuroides (Regel) Roshkova
Annual. Saline lands. Plain. Mirzachul. Meliferous, ornamental.
1030. Psylliostachys suworowii (Regel) Roshkova
Annual. Saline lands, river valleys, banks of canals, fallow lands, fields. Plain. Mirzachul.
Meliferous, ornamental.
Family 81. Polygonaceae
Genus 319. Oxyria Hill
1031. Oxyria digyna (L.) Hill Figure 309
Perennial. Banks of rivers, pebbles, wet stony slopes and screes. Alpine zone. North Turkestan. Dye,
food.
Figure 309 Oxyria digyna (photography by Natalya Beshko)
303
304
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 320. Rumex L.
1032. Rumex acetosa L.
Perennial. Banks of rivers, wet places, gardens, meadows. Plain, foothills, montane zone. Nuratau,
Mirzachul. Medicinal, food, fodder, dye.
1033. Rumex chalepensis Mill. [Rumex drobovii Korovin]
Perennial. Meadows, wet places, roadsides, banks of rivers and canals, fine-earth and stony slopes,
gardens, fields, wastelands. Plain, foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Tanniferous, weed.
1034. *Rumex conglomeratus Murray
Perennial. Banks of rivers and canals, wet places, swamps, roadsides, wastelands. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal,
tanniferous, weed.
1035. *Rumex crispus L.
Perennial. Meadows, fine-earth and stony slopes, banks of rivers, lakes and canals. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal,
tanniferous, weed, food.
1036. Rumex dentatus subsp. halacsyi (Rech.) Rech. f. [Rumex halacsyi Rech.]
Annual or biennial. River valleys, wet places, saline lands, banks of canals, fallow lands. Plain,
foothills. Malguzar, Mirzachul. Weed.
1037. Rumex marschallianus Rchb.
Annual. Banks of rivers, lakes and canals, saline lands. Plain. Mirzachul.
1038. Rumex pamiricus Rech. f.
Perennial. Fine-earth and stony slopes, screes, wet places, banks of rivers and canals, ravines, fallow lands, wastelands. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Malguzar, North
Turkestan. Tanniferous, weed.
1039. Rumex paulsenianus Rech. f.
Perennial. Banks of rivers, ravines, fine-earth and stony slopes, fallow lands. Montane and alpine
zones. North Turkestan. Tanniferous, weed.
1040. Rumex rectinervis Rech. f.
Perennial. Banks of rivers and canals, swamps. Plain, foothills, montane zone. Nuratau, Aktau,
Malguzar, North Turkestan. Medicinal, tanniferous.
1041. *Rumex syriacus Meissn
Perennial. Banks of rivers and canals, wet places, pebbles, fallow lands, fields, gardens. Plain,
foothills. Nuratau Relic Mountains, Mirzachul. Tanniferous, dye, weed.
1042. Rumex tianschanicus Losinsk.
Perennial. Fine-earth and stony slopes, meadows, banks of rivers. Montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Medicinal, tanniferous, fodder, weed.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 321. Rheum L.
1043. Rheum сordatum Losinsk.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau. Food, medicinal,
tanniferous, dye.
1044. Rheum macrocarpum Losinsk.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Food, tanniferous, dye.
1045. Rheum maximowiczii Losinsk. Figure 310
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Food, medicinal, tanniferous, dye.
Figure 310 Rheum maximowiczii (photography by Natalya Beshko)
305
306
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1046. Rheum turkestanicum Janisch. Figure 311
Perennial. Sandy deserts. Plain. Kyzylkum. Food, medicinal, tanniferous, dye, fodder.
Figure 311 Rheum turkestanicum (photography by Natalya Beshko)
Genus 322. Atraphaxis L.
1047. Atraphaxis compacta Ledeb.
Shrub. Stony slopes. Foothills. Nuratau Relic Mountains. Tanniferous, ornamental, meliferous.
1048. Atraphaxis karataviensis Lipsch. & Pavlov.
Shrub. Stony slopes, dry riverbeds. Montane zone. Nuratau. Tanniferous, ornamental, meliferous.
1049. Atraphaxis pyrifolia Bunge Figure 312
Shrub. Fine-earth and stony slopes, ravines. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Tanniferous, dye, ornamental, meliferous.
Figure 312 Atraphaxis pyrifolia (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1050. Atraphaxis seravschanica Pavlov
Shrub. Fine-earth and stony slopes, rocks, ravines. Montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Tanniferous, ornamental, meliferous.
1051. Atraphaxis spinosa L.
Shrub. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau. Tanniferous, dye,
ornamental, fodder, meliferous.
1052. Atraphaxis virgata (Regel) Krasn. Figure 313
Shrub. Stony slopes, pebbles, dry riverbeds. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Tanniferous, ornamental, meliferous.
Figure 313 Atraphaxis virgata (photography by Natalya Beshko)
Genus 323. Calligonum L.
1053. Calligonum aphyllum (Pall.) Gürke
Shrub. Sandy deserts. Plain. Kyzylkum. Fodder, tanniferous, dye, afforestation, meliferous.
1054. Calligonum caput-medusae Schrenk Figure 314
Shrub. Sandy deserts. Plain. Kyzylkum. Fodder, tanniferous, dye, afforestation, meliferous.
307
308
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 314 Calligonum caput-medusae (photography by Natalya Beshko)
1055. Calligonum leucocladum (Schrenk) Bunge Figure 315
Shrub. Sandy deserts. Plain. Kyzylkum. Fodder, tanniferous, dye, afforestation, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 315 Calligonum leucocladum (photography by Natalya Beshko)
1056. Calligonum macrocarpum I. G. Borshch. Figure 316
Shrub. Sandy deserts. Plain. Kyzylkum. Fodder, afforestation, meliferous.
Figure 316 Calligonum macrocarpum (photography by Natalya Beshko)
309
310
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1057. Calligonum microcarpum I. G. Borshch. Figure 317
Shrub. Sandy deserts. Plain. Kyzylkum. Fodder, afforestation, meliferous.
Figure 317 Calligonum microcarpum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 324. Polygonum L.
1058. Polygonum acetosum M. Bieb.
Annual. Sandy deserts. Plain. Kyzylkum.
1059. Polygonum argyrocoleon Steud. ex Kunze
Annual. Sandy soils, saline lands, wet places, roadsides, fields. Plain, foothills. Nuratau, Aktau,
Mirzachul, Kyzylkum. Medicinal, tanniferous, weed.
1060. Polygonum aviculare L.
Annual. Wastelands, fields, fallow lands, roadsides, banks of canals. Plain, foothills, montane and
alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, dye, food, fodder, weed.
1061. Polygonum biaristatum Aitch. & Hemsl
Semishrub. Stony slopes, screes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Fodder.
1062. Polygonum coriarium Grig. [Aconogonon coriarum (Grig.) Sojăk] Figure 318
Perennial. Fine-earth and stony slopes, subalpine meadows. Montane and alpine zones. North
Turkestan. Medicinal, tanniferous, dye, food, meliferous.
Figure 318 Polygonum coriarium (photography by Natalya Beshko)
1063. Polygonum fibrilliferum Kom.
Perennial. Gravelly and stony slopes. Montane and alpine zones. Malguzar, North Turkestan. Fodder,
311
312
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
weed.
1064. Polygonum hissaricum Popov [Aconogonon hissaricum (Popov) Sojăk]
Perennial. Fine-earth and stony slopes. Alpine zone. North Turkestan. Medicinal, tanniferous, food,
meliferous.
1065. Polygonum inflexum Kom.
Annual. Banks of rivers and canals, fine-earth slopes, saline lands, fallow lands, roadsides, fields.
Plain, foothills. Mirzachul.
1066. Polygonum molliiforme Boiss.
Annual. Stony slopes, pebbles, banks of rivers. Montane and alpine zones. Malguzar, North
Turkestan.
1067. Polygonum paronychioides C. A. Mey.
Semishrub. Stony slopes, screes, rocks, pebbles. Foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Tanniferous.
1068. Polygonum patulum M. Bieb.
Annual. Stony slopes, screes, saline lands, roadsides, fields, fallow lands. Plain, foothills, montane
zone. Nuratau, Malguzar, North Turkestan. Fodder, weed.
1069. Polygonum polychnemoides Jaub. & Spach.
Annual. Stony slopes, screes, pebbles, roadsides, fields. Plain, foothills, montane and alpine zones.
Nuratau, Malguzar, North Turkestan.
1070. Polygonum pulvinatum Kom.
Semishrub. Stony slopes. Montane zone. Nuratau.
1071. Polygonum rottboellioides Jaub. & Spach.
Annual. Fine-earth and stony slopes, pebbles. Plain, foothills, montane zone. Nuratau, Nuratau Relic
Mountains.
1072. Polygonum subaphyllum Sumnev.
Annual. Fine-earth and stony slopes, pebbles, fallow lands, banks of canals, fields. Foothills,
montane zone. Nuratau.
1073. Polygonum vvedenskyi Sumnev. Figure 319
Semishrub. Stony slopes, screes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Figure 319 Polygonum vvedenskyi (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 325. Persicaria Hill
1074. Persicaria amphibia (L.) Delarbre [Polygonum amphibium L]
Perennial. Banks of rivers, lakes, ponds and canals. Plain, foothills, montane and alpine zones.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal,
tanniferous.
1075. Persicaria bistorta (L.) Samp. [Bistortia elliptica (Willd. ex Spreng.) Kom.; Polygonum
nitens (Fish. & C. A. Mey.) V. Petr. ex Kom.] Figure 320
Perennial. Fine-earth and stony slopes, screes, banks of rivers, swamps, subalpine and alpine
meadows. Montane and alpine zones. Malguzar, North Turkestan. Medicinal, tanniferous, dye,
meliferous.
Figure 320 Persicaria bistorta (photography by Natalya Beshko)
1076. Persicaria hydropiper (L.) Delarbre
Annual. Banks of rivers, ponds and canals, swamps. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, dye.
313
314
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1077. Persicaria lapathifolia (L.) Delarbre
Annual. Banks of rivers, ponds and canals, wet places, fields. Plain, foothills, montane zone.
Nuratau, North Turkestan. Medicinal, tanniferous, food, fodder, meliferous.
1078. Persicaria maculata (Raf.) Gray.
Annual. Banks of rivers and canals, wet places, fields, wasteands. Plain, foothills, montane and
alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, essential oil, dye, weed, meliferous.
1079. Persicaria minor (Huds.) Opiz.
Annual. Banks of rivers and canals. Plain, foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Medicinal, weed.
Genus 326. Fallopia Adans.
1080. Fallopia convolvulus (L.) A. Löve
Annual. Woodlands, banks of rivers, fields, gardens, wastelands. Plain, foothills, montane and alpine
zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Medicinal, tanniferous, food, meliferous, weed.
Family 82. Caryophyllaceae
Genus 327. Stellaria L.
1081. Stellaria alsinoides Boiss. & Buhse [Tytthostemma alsinoides (Boiss. & Buhse) Nevski]
Annual. Stony slopes, pebbles. Foothills, montane zone. Nuratau, Malguzar.
1082. Stellaria brachypetala Bunge
Perennial. Wet places, banks of rivers, pebbles, meadows, fields, fine-earth slopes. Montane and
alpine zones. Malguzar, North Turkestan. Poisonous.
1083. *Stellaria media (L.) Vill.
Annual. Roadsides, banks of rivers and canals, shady wet places, settlements, gardens, wastelands.
Plain, foothills, montane zone. Nuratau. Weed, medicinal, dye, saponin-bearing.
1084. Stellaria neglecta Weihe
Annual. River valleys, gardens, fields, wastelands, roadsides, swamps, shady wet places. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Weed.
1085. Stellaria pallida (Dumort.) Crep.
Annual. Shady wet places, gardens, fields, fallow lands. Nuratau.
1086. Stellaria turkestanica Schischk.
Perennial. Screes, fine-earth and stony slopes. Alpine zone. Malguzar, North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 328. Mesostemma Vved.
1087. Mesostemma karatavica (Schischk.) Vved. [Stellaria karatavica Schischk.] Figure 321
Perennial. Stony slopes, rocks. Montane zone. Nuratau, North Turkestan.
Figure 321 Mesostemma karatavica (photography by Natalya Beshko)
315
316
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 329. Cerastium L.
1088. Cerastium arvense L.
Perennial. Fine-earth slopes, meadows. Montane zone. North Turkestan. Medicinal, saponin-bearing, fodder.
1089. Cerastium cerastoides (L.) Britton.
Perennial. Fine-earth, gravelly and stony slopes, rocks, alpine meadows, places near melting snow.
Alpine zone. North Turkestan. Fodder, saponin-bearing.
1090. Cerastium dichotomum L.
Annual. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan.
1091. Cerastium dichotomum subsp. inflatum Cullen [Cerastium inflatum Link. ex Desf.]
Annual. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan.
1092. Cerastium falcatum (Gren.) Bunge ex Fenzl [Cerastium bungeanum Vved.]
Perennial. Fine-earth, gravelly and stony slopes, rocks, saline lands. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
1093. Cerastium fontanum subsp. vulgare (Hartm.) Greuter & Burdet [Cerastium holosteoides Fries]
Annual, biennial, perennial. Montane and alpine zones. North Turkestan. Saponin-bearing.
1094. Cerastium glomeratum Thuill
Annual. River valleys, pebbles, roadsides, gardens, fields, fallow lands. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
1095. Cerastium lithospermifolium Fisch.
Perennial. Screes. Alpine zone. North Turkestan.
1096. Cerastium perfoliatum L.
Annual. Fine-earth and stony slopes, gardens, fields, fallow lands. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Saponin-bearing.
1097. Cerastium pusillum Ser.
Perennial. Stony slopes, banks of rivers, swamps, pebbles. Montane and alpine zones. Malguzar,
North Turkestan.
1098. Cerastium semidecandrum L. [Cerastium pentandrum L.; Cerastium dentatum Möeschl.]
Annual. Clayey soils. Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul.
Genus 330. Holosteum L.
1099. Holosteum umbellatum L.
Annual. Piedmont plains, fine-earth, gravelly and stony slopes, pebbles, fallow lands, fields. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed, saponin-bearing.
MAGNOLIOPSIDA
[EUDICOTS]
1100. Holosteum umbellatum subsp. glutinosum (M. Bieb.) Nyman [Holosteum glutinosum
(M. Bieb.) Fisch. & C. A. Mey.; Holosteum polygamum C. Koch]
Annual. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, pebbles,
fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Weed, saponin-bearing.
Genus 331. Sagina L.
1101. Sagina saginoides (L.) H. Karst.
Perennial. Stony slopes, pebbles, screes, wet places, banks of rivers, swamps. Alpine zone. North
Turkestan.
Genus 332. Bufonia L.
1102. Bufonia oliveriana Ser.
Semishrub. Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Aktau.
Genus 333. Lepyrodiclis Fenzl.
1103. Lepyrodiclis holosteoides (C. A. Mey.) Fenzl ex Fisch. & C. A. Mey.
Annual. River valleys, fine-earth and stony slopes, shady wet places, fallow lands, fields, gardens.
Plain, foothills, montane and alpine zones, Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan. Weed, saponin-bearing.
1104. Lepyrodiclis stellarioides Fisch. & C. A. Mey.
Annual. River valleys, fine-earth and stony slopes, woodlands, shady wet places, gardens. Foothills,
montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Saponin-bearing.
Genus 334. Minuartia L.
1105. Minuartia hamata (Hausskn.) Mattf. [Queria hispanica L.]
Annual. Pebbles, fine-earth, gravelly and stony slopes. Foothills. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan.
1106. Minuartia kryloviana Schischk.
Perennial. Stony slopes, rocks. Alpine zone. North Turkestan.
1107. Minuartia meyeri (Boiss.) Bornm.
Annual. Fine-earth, gravelly and stony slopes, pebbles. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan.
317
318
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 335. Eremogone Fenzl
1108. Eremogone griffithii (Boiss.) Ikonn. [Arenaria griffithii Boiss.] Figure 322
Perennial. Stony slopes, rocks. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Saponin-bearing.
Figure 322 Eremogone griffithii (photography by Natalya Beshko)
Genus 336. Arenaria L.
1109. Arenaria rotundifolia M. Bieb.
Perennial. Stony slopes, banks of rivers, alpine meadows, swamps. Alpine zone. North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
1110. Arenaria serpyllifolia L.
Annual. Fine-earth, gravelly and stony slopes, fields, fallow lands. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Fodder.
1111. Arenaria serpyllifolia subsp. leptoclados (Rchb.) Nyman [Arenaria leptoclados (Rchb.)
Guss.]
Annual. Clay deserts, fine-earth, gravelly and stony slopes. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder.
Genus 337. Spergularia L.
1112. Spergularia diandra (Guss.) Heldr.
Annual. Saline lands. Plain. Mirzachul, Kyzylkum.
1113. *Spergularia marina (L.) Besser [Spergularia salina J. Presl & C. Presl]
Annual, biennial or perennial. Saline lands, banks of rivers, lakes and canals. Plain, foothills.
Nuratau, Mirzachul, Kyzylkum.
1114. Spergularia media (L.) C. Presl.
Annual or biennial. Saline lands, wastelands, banks of canas. Plain. Mirzachul, Kyzylkum. Weed.
1115. *Spergularia rubra J. Presl & C. Presl.
Annual or biennial. Saline lands, wastelands, roadsides, fields, banks of rivers. Plain, foothills.
Nuratau, Mirzachul, Kyzylkum. Weed.
Genus 338. Herniaria L.
1116. Herniaria glabra L.
Annual. Fine-earth, gravelly and stony slopes, banks of rivers, roadsides. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Weed,
poisonous.
1117. Herniaria hirsuta L.
Annual. River valleys, dry riverbeds, pebbles, roadsides. Plain, foothills, montane zone. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Weed, poisonous.
Genus 339. *Agrostemma L.
1118. *Agrostemma githago L.
Annual. Fallow lands, fields, roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, weed, poisonous.
319
320
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 340. Silene L.
1119. Silene brahuica Boiss.
Perennial. Fine-earth, gravelly and stony slopes, ravines, rocks, pebbles. Foothills, montane zone,
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, saponin-bearing.
1120. Silene claviformis Litv.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan.
1121. Silene conica L. Figure 323
Annual. River valleys, pebbles, fine-earth and stony slopes, fallow lands, fields. Foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Saponin-bearing.
Figure 323 Silene conica (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1122. Silene coniflora Nees ex Otth. Figure 324
Annual. Fine-earth, gravelly and stony slopes, river valleys, banks of canals, roadsides. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Figure 324 Silene coniflora (photography by Natalya Beshko)
1123. Silene conoidea L. Figure 325
Annual. Fine-earth, gravelly and stony slopes, river valleys, banks of canals, wastelands, roadsides,
gardens, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Kyzylkum. Weed.
Figure 325 Silene conoidea (photography by Natalya Beshko)
321
322
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1124. Silene graminifolia Otth [Silene schischkinii Sobolevsk.]
Perennial. Fine-earth, gravelly and stony slopes. Alpine zone. North Turkestan.
1125. Silene guntensis Schischk.
Perennial. Stony slopes, rocks, screes, pebbles. Montane and alpine zones. Nuratau, Aktau, Malguzar,
North Turkestan.
1126. Silene incurvifolia Kar. & Kir.
Perennial. Stony slopes, rocks, pebbles. Foothills, montane and alpine zones. North Turkestan.
1127. Silene kuschakewiczii Regel & Schmalh.
Perennial. Stony slopes, rocks, screes. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan.
1128. Silene latifolia Poir. [Melandrium album (Mill.) Garcke]
Annual or biennial. Fine-earth slopes, river valleys, ravines. Montane zone. North Turkestan. Weed,
saponin-bearing.
1129. Silene longicalycina Kom. Figure 326
Perennial. Stony slopes, screes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Figure 326 Silene longicalycina (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1130. Silene nana Kar. & Kir. Figure 327
Annual. Sandy deserts. Plain. Kyzylkum.
Figure 327 Silene nana (photography by Natalya Beshko)
1131. Silene nevskii Schischk.
Perennial. Stony slopes, rocks. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
1132. Silene obtusidentata B. Fedtsch. & Popov
Perennial. Stony slopes. Montane zone. Nuratau.
1133. Silene paranadena Bondarenko & Vved. Figure 328
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar. UzbRDB 2.
Figure 328 Silene paranadena (photography by N. Yu. Beshko)
323
324
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1134. Silene plurifolia Schischk.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar.
1135. Silene praelonga Ovcz.
Perennial. Stony slopes, rocks. Montane zone. Nuratau.
1136. Silene praemixta Popov
Perennial. Sandy and clay deserts, piedmont plains, foothills, pebbles, river valleys, fine-earth,
gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Saponin-bearing.
1137. Silene pugionifolia Popov
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau.
1138. Silene ruinarum Popov [Melandrium ruinarium Popov]
Perennial. Fine-earth, gravelly and stony slopes, river valleys, wet places. Foothills, montane zone.
Nuratau.
1139. Silene stenantha Ovcz.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. North Turkestan.
1140. Silene tachtensis Franch.
Perennial. Stony slopes, rocks, screes. Montane and alpine zones. North Turkestan.
1141. Silene turkestanica Regel [Melandrium turkestanicum Vved.]
Perennial. Stony slopes, screes, woodlands. Montane zone. North Turkestan.
1142. Silene uralensis subsp. apetala (L.) Bocquet [Gastrolychnis apetala (L.) Tolm. &
Kozhanch]
Perennial. Stony slopes, screes, swamps, pebbles. Alpine zone. North Turkestan. Saponin-bearing.
1143. Silene vulgaris (Moench) Garcke Figure 329
Perennial. Fine-earth, gravelly and stony slopes, screes, river valleys, ravines, subalpine meadows.
Montane and alpine zones. Malguzar, North Turkestan. Medicinal, saponin-bearing, ornamental.
Figure 329 Silene vulgaris (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 341. Gastrolychnis (Fenzl.) Rchb.
1144. Gastrolychnis longicarpophora (Kom.) Czerep. [Silene longicarpophora (Kom.) Bocquet]
Perennial. Stony slopes, rocks, screes, alpine meadows, places near melting snow. Alpine zone.
North Turkestan.
Genus 342. Gypsophila L.
1145. Gypsophila cephalotes (Schrenk) F. N. Williams
Perennial. Stony and gravelly slopes, pebbles, river valleys. Montane and alpine zones. Malguzar,
North Turkestan. Saponin-bearing.
1146. Gypsophila herniarioides Boiss.
Perennial. Moraines, screes, stony slopes, pebbles. Alpine zone. North Turkestan.
1147. *Gypsophila paniculata L.
Perennial. Roadsides, dry riverbeds, wastelands. Plain, foothills, montane zone. Malguzar. Medicinal,
poisonous, ornamental.
Genus 343. Bolbosaponaria Bondar.
1148. Bolbosaponaria sewerzowii Bondar. Figure 330
Perennial. Stony slopes, rocks. Foothills, montane zone. North Turkestan. Saponin-bearing.
Figure 330 Bolbosaponaria sewerzowii (photography by Natalya Beshko)
Genus 344. Kuhitangia Ovcz.
1149. Kuhitangia knorringiana Bondarenko
Perennial. Stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan. Saponin-bearing.
325
326
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 345. Petrorhagia (Ser. ex DC.) Link.
1150. Petrorhagia alpina (Hablitz) P. W. Ball & Heywood
Annual or biennial. Stony slopes, screes, river valleys, ravines, rocks, subalpine and alpine meadows.
Montane and alpine zones. North Turkestan. Saponin-bearing.
Genus 346. Acanthophyllum C. A. Mey.
1151. Acanthophyllum aculeatum Schischk.
Perennial. Sandy deserts, piedmont plains, fine-earth and stony slopes, screes. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar. Medicinal, meliferous, saponin-bearing.
1152. Acanthophyllum elatius Bunge
Perennial. Sandy deserts. Plain. Kyzylkum. Saponin-bearing.
1153. Acanthophyllum pungens (Bunge) Boiss. Figure 331
Perennial. Sandy deserts, piedmont plains, fine-earth, gravelly and stony slopes, screes. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Medicinal, meliferous, saponin-bearing.
Figure 331 Acanthophyllum pungens (photography by Natalya Beshko)
Genus 347. Allochrusa Bunge ex Boiss.
1154. Allochrusa gypsophiloides (Regel) Schischk. [Acanthophyllum gypsophiloides Regel]
Perennial. Fine-earth, gravelly and stony slopes, dry riverbeds, fallow lands. Foothills, montane
zone. Nuratau, Malguzar, North Turkestan. Medicinal, food, saponin-bearing. UzbRDB 3.
Genus 348. *Vaccaria Medik.
1155. *Vaccaria hispanica (Mill.) Rauschert
Annual. Fields, fallow lands, banks of canals, meadows, wastelands. Plain, foothills, montane zone.
MAGNOLIOPSIDA
[EUDICOTS]
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Weed, ornamental,
medicinal.
Genus 349. Dianthus L.
1156. Dianthus baldzhuanicus Lincz.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
1157. Dianthus brevipetalus Vved.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan.
1158. Dianthus crinitus subsp. tetralepis (Nevski) Rech.f. [Dianthus tetralepis Nevski]
Perennial. Fine-earth, gravelly and stony slopes, river valleys, piedmont plains. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental,
saponin-bearing.
1159. Dianthus darvazicus Lincz. Figure 332
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar. Saponin-bearing.
Figure 332 Dianthus darvazicus (photography by Natalya Beshko)
327
328
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1160. Dianthus helenae Vved. Figure 333
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Aktau. Endemic to the Nuratau Mountains.
Figure 333 Dianthus helenae (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1161. Dianthus subscabridus Lincz.
Perennial. Stony slopes, rocks, dry riverbeds. Montane and alpine zones. Malguzar, North Turkestan.
Genus 350. Pleioneura Rech. f.
1162. Pleioneura griffithiana (Boiss.) Rech. f. [Saponaria griffithiana Boiss.]
Perennial. Stony slopes, rocks, screes, pebbles. Montane and alpine zones. Malguzar, North
Turkestan.
Genus 351. Velezia L.
1163. Velezia rigida L.
Annual. Fine-earth and stony slopes, screes, pebbles, dry riverbeds. Foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan.
Family 83. Amaranthaceae
Genus 352. *Amaranthus L.
1164. *Amaranthus albus L.
Annual. Fields, fallow lands, wastelands, roadsides, banks of canals. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Weed, fodder.
1165. *Amaranthus blitoides S. Watson
Annual. Fields, fallow lands, wastelands, roadsides. Plain, foothills, montane zone. North Turkestan.
Weed, fodder.
1166. *Amaranthus deflexus L.
Annual. Gardens, fields, fallow lands, wastelands. Plain. Mirzachul, Kyzylkum. Weed.
1167. *Amaranthus retroflexus L.
Annual. Settlements, gardens, fields, fallow lands, roadsides, wastelands. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Weed, fodder, medicinal.
Genus 353. Polycnemum L.
1168. Polycnemum arvense L.
Annual. Stony slopes, pebbles, fallow lands, wastelands. Plain, foothills, montane zone. Nuratau,
329
330
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Malguzar. Weed, fodder.
1169. Polycnemum perenne Litv. Figure 334
Perennial. Stony slopes, rocks, screes. Foothills, montane zone. North Turkestan.
Figure 334 Polycnemum perenne (photography by Natalya Beshko)
Genus 354. Chenopodium L.
1170. Chenopodium album L.
Annual. Wastelands, fields, gardens, fallow lands, roadsides. Plain, foothills, montane and alpine
MAGNOLIOPSIDA
[EUDICOTS]
zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Weed, fodder, medicinal, food, dye, essential oil, meliferous.
1171. Chenopodium botrys L.
Annual. Dry riverbeds, pebbles, stony slopes, wastelands, fields. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Medicinal, dye, essential oil.
1172. Chenopodium chenopodioides (L.) Aellen
Annual. Saline lands. Plain, foothills. Kyzylkum. Weed.
1173. Chenopodium foliosum (Moench) Aschers Figure 335
Annual. Stony slopes, pebbles, screes, rocks, wastelands, roadsides. Plain, foothills, montane and
alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Food, medicinal, dye.
Figure 335 Chenopodium foliosum (photography by Natalya Beshko)
331
332
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1174. Chenopodium glaucum L.
Annual. Saline lands, banks of rivers, lakes and canals, fields, wastelands. Plain, foothills, montane
and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Weed.
1175. *Chenopodium murale L.
Annual. Wastelands. Foothills, montane zone. Nuratau, Malguzar, North Turkestan. Food, medicinal.
1176. Chenopodium rubrum L.
Annual. Saline lands, wastelands, banks of rivers, lakes and canals, river valleys, fields. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Kyzylkum. Food, medicinal, weed.
1177. *Chenopodium vulvaria L.
Annual. Gravelly slopes, pebbles, dry riverbeds, wastelands. Plain, foothills, montane and alpine
zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Kyzylkum. Weed,
medicinal, dye.
Genus 355. Spinacia L.
1178. Spinacia turkestanica Iljin
Annual. Fields, fallow lands, wastelands, banks of canals. Plain, foothills, montane zone. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Food, fodder, weed.
Genus 356. Atriplex L.
1179. Atriplex aucherii Moq.
Annual. Saline lands, roadsides, wastelands, surroundings of wells and sheepyards. Plain, foothills.
Nuratau Relic Mountains, Kyzylkum. Fodder.
1180. Atriplex dimorphostegia Kar. & Kir.
Annual. Sandy deserts, saline lands. Plain. Kyzylkum. Fodder.
1181. Atriplex flabellum Bunge
Annual. Gravelly and fine-earth slopes, roadsides, wastelands, surroundings of wells and sheepyards.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Kyzylkum, Mirzachul. Fodder.
1182. Atriplex micrantha Ledeb.
Annual. Saline lands, banks of canals, wastelands, fallow lands. Plain, foothills. Malguzar,
Mirzachul. Fodder, weed.
1183. Atriplex ornata Iljin
Annual. Saline lands, fallow lands, banks of canals, sandy deserts. Plain. Mirzachul, Kyzyzlkum.
Fodder, weed.
MAGNOLIOPSIDA
[EUDICOTS]
1184. Atriplex prostrata subsp. calotheca (Rafn) M. A. Gust. [Atriplex hastata L.]
Annual. Saline lands, banks of canals, roadsides, fields, fallow lands. Plain. Mirzachul. Fodder, weed.
1185. Atriplex tatarica L.
Annual. Saline lands, banks of canals, roadsides, surroundings of wells and sheepyards, fields,
wastelands. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, food, medicinal.
Genus 357. Krascheninnikovia Gueldenst.
1186. Krascheninnikovia ceratoides (L.) Gueldenst. [Ceratoides latens Reveal & Holmgren;
Eurotia ceratoides (L.) C. A. Mey.] Figure 336
Semishrub. Stony, gravelly and fine-earth slopes, sandy soils, pebbles, dry riverbeds. Plain, foothills,
montane and alpine zones. Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder.
Figure 336 Krascheninnikovia ceratoides (photography by Natalya Beshko)
1187. Krascheninnikovia ewersmanniana (Stschegl. ex Losinsk.) Grubov [Ceratoides
ewersmanniana (Stschegl. ex Losinsk.) Botsch. & Ikonn.; Eurotia ewersmanniana Stschegl. ex
Losinsk.]
Semishrub. Stony slopes, screes, sandy soils, saline lands, dry riverbeds. Plain, foothills, montane
and alpine zones. Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, oil, dye.
Genus 358. Ceratocarpus L.
1188. Ceratocarpus arenarius L. [Ceratocarpus utriculosus Bluket ex Krylov]
Annual. Sandy and clay deserts, saline lands, pebbles, dry riverbeds, fine-earth slopes, roadsides,
wastelands, fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, Mirzachul, Kyzylkum. Weed, fodder.
333
334
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 359. Camphorosma L.
1189. Camphorosma monspeliaca L.
Semishrub. Sandy deserts, saline lands, banks of lakes. Plain. Kyzylkum. Fodder, essential oil,
medicinal.
Genus 360. Bassia All.
1190. Bassia crassifolia (Pall.) Soldano [Suaeda crassifolia Pall.]
Annual. Saline lands, banks of lakes and canals. Plain. Mirzachul. Fodder.
1191. Bassia eriophora (Schrad.) Asch. [Kirilovia eriantha Bunge; Londesia eriantha Fisch. et
C. A. Mey.]
Annual. Sandy and clay deserts, surroundings of wells and sheepyards. Plain. Kyzylkum.
1192. Bassia hyssopifolia (Pall.) Kuntze
Annual. Banks of canals, saline lands, wastelands, fields, surroundings of wells and sheepyards.
Plain. Mirzachul, Kyzylkum. Weed.
Genus 361. Kochia Roth.
1193. Kochia iranica Bornm.
Annual. Sandy soils, saline lands. Plain. Kyzylkum. Fodder.
1194. Kochia prostrata (L.) Schrad.
Semishrub. Sandy soils, saline lands, stony and gravelly slopes. Plain, foothills, montane and alpine
zones, Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Kyzylkum. Fodder.
1195. Kochia scoparia (L.) Schrad.
Annual. Fields, gardens, roadsides, wastelands. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum. Fodder, medicinal, ornamental, weed.
Genus 362. Corispermum L.
1196. Corispermum lehmanninum Bunge
Annual. Sandy deserts. Plain. Kyzylkum. Fodder.
Genus 363. Agriophyllum M. Bieb.
1197. Agriophyllum lateriflorum (Lam.) Moq.
Annual. Sandy deserts. Plain. Kyzylkum. Fodder.
Figure 337
MAGNOLIOPSIDA
[EUDICOTS]
Figure 337 Agriophyllum lateriflorum (photography by Natalya Beshko)
335
336
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 364. Kalidium Moq.
1198. Kalidium caspicum (L.) Ung.-Sternb.
Semishrub. Saline lands, banks of lakes. Plain. Kyzylkum. Industrial.
Genus365. Halopeplis Bunge
1199. Halopeplis pygmaea (Pall.) Bunge ex Ung.-Sternb.
Annual. Saline lands, banks of lakes and canals. Plain. Mirzachul, Kyzylkum.
Genus 366. Halostachys C. A. Mey.
1200. Halostachys belangeriana (Moq.) Botsch. Figure 338
Shrub. Saline lands, banks of lakes and canals, fallow lands. Plain. Nuratau Relic Mountains,
Mirzachul, Kyzylkum. Industrial.
Figure 338 Halostachys belangeriana (photography by Natalya Beshko)
Genus 367. Halocnemum M. Bieb.
1201. Halocnemum strobilaceum (Pall.) M. Bieb.
Semishrub. Saline lands, banks of lakes and canals. Plain. Mirzachul, Kyzylkum. Alkaloid-bearing,
insecticide, industrial.
Genus 368. Salicornia L.
1202. Salicornia europaea L.
Annual. Saline lands, banks of lakes and canals, depressions, fallow lands. Plain. Nuratau Relic
MAGNOLIOPSIDA
[EUDICOTS]
Mountains, Mirzachul, Kyzylkum. Fodder, insecticide, food, medicinal, industrial.
Genus 369. Suaeda Forsk.
1203. Suaeda acuminata (C. A. Mey.) Moq.
Annual. Saline lands, wastelands, fallow lands, fields. Plain. Mirzachul, Kyzylkum. Fodder.
1204. Suaeda altissima (L.) Pall.
Annual. Wastelands, fallow lands, fields, saline lands, banks of canals, roadsides. Plain. Mirzachul,
Kyzylkum. Fodder.
1205. Suaeda arcuata Bunge
Annual. Saline lands, wastelands, fallow lands, fields. Plain. Mirzachul, Kyzylkum. Fodder, alkaloidbearing.
1206. Suaeda dendroides (C. A. Mey.) Moq.
Shrub. Saline lands, banks of lakes and canals. Plain. Mirzachul, Kyzylkum. Fodder, industrial.
1207. Suaeda heterophylla Bunge ex Boiss.
Annual. Saline lands, fallow lands, fields, roadsides, banks of canals, wastelands. Plain. Mirzachul,
Kyzylkum. Fodder, weed.
1208. Suaeda linifolia Pall.
Annual. Saline lands, banks of lakes and canals, fallow lands, fields. Plain. Mirzachul, Kyzylkum.
Fodder, weed.
1209. Suaeda paradoxa (Bunge) Bunge
Annual. Saline lands, wastelands, fallow lands, fields. Plain. Mirzachul, Kyzylkum. Fodder, weed.
1210. Suaeda physophora Pall.
Semishrub. Saline lands, banks of lakes and canals. Plain. Mirzachul. Industrial, poisonous.
Genus 370. Salsola L.
1211. Salsola arbuscula Pall.
Shrub. Sandy deserts, stony and gravelly slopes. Plain, foothills. Nuratau Relic Mountains, Malguzar,
Mirzachul, Kyzylkum. Fodder, dye, tanniferous, afforestation.
1212. Salsola arbusculiformis Drobow
Shrub. Sandy deserts, clayey and gravelly soils, pebbles. Plain, foothills. Nuratau Relic Mountains,
Malguzar, Kyzylkum. Fodder.
1213. Salsola collina Pall.
Annual. Sandy and clay deserts, piedmont plain, fallow lands, saline lands, screes, roadsides,
dry riverbeds, surroundings of settlements, wells and sheepyards. Plain, foothills. Nuratau Relic
Mountains, Kyzylkum. Fodder, medicinal.
1214. Salsola dendroides Pall.
Semishrub. Saline lands, gravelly slopes, river valleys, fields. Plain, foothills. Nuratau Relic
337
338
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Mountains, Mirzachul, Kyzylkum. Fodder, afforestation.
1215. Salsola foliosa (L.) Schrad. ex Schult.
Annual. Sandy and clay deserts, saline lands. Plain. Mirzachul, Kyzylkum. Fodder.
1216. Salsola iberica Sennen & Pau.
Annual. Sandy, clayey and skeleton soils, saline lands, wastelands, fields. Plain, foothills. Nuratau,
Aktau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder.
1217. Salsola leptoclada Gand.
Annual. Sandy, clayey and gravelly soils. Plain. Nuratau Relic Mountains, Mirzachul, Kyzylkum.
Fodder.
1218. Salsola micranthera Botschantz.
Annual. Saline lands, sandy and clay deserts. Plain, foothills. Nuratau Relic Mountains, Kyzylkum.
Fodder.
1219. Salsola orientalis S. G. Gmel.
Semishrub. Clay desert, saline lands, piedmont plains, gravelly slopes. Plain, foothills. Nuratau Relic
Mountains, Mirzachul, Kyzylkum. Fodder.
1220. Salsola paletzkiana Litv.
Tree or shrub. Sandy deserts. Plain. Kyzylkum. Dye, afforestation, ornamental.
1221. Salsola paulsenii Litv. Figure 339
Annual. Sandy and clay deserts, saline lands. Plain. Mirzachul, Kyzylkum. Fodder.
Figure 339 Salsola paulsenii (photography by Natalya Beshko)
1222. Salsola richteri (Moq.) Kar. ex Litv. Figure 340
Tree or shrub. Sandy deserts. Plain. Kyzylkum. Fodder, dye, afforestation, medicinal, alkaloid-bearing.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 340 Salsola richteri (photography by Natalya Beshko)
1223. Salsola sclerantha C. A. Mey.
Annual. Sandy deserts, saline lands. Plain. Mirzachul, Kyzylkum. Fodder.
1224. Salsola titovii Botsch.
Semishrub. Stony slopes. Montane zone. Aktau. UzbRDB 1.
1225. Salsola turkestanica Litv.
Annual. Saline lands. Plain. Mirzachul, Kyzylkum. Fodder.
1226. Salsola vvedenskyi Iljin & Popov
Annual. Saline lands, sandy deserts. Plain. Kyzylkum. Fodder.
Genus 371. Climacoptera Botsch.
1227. Climacoptera lanata (Pall.) Botsch.
Annual. Saline lands, sandy and clay deserts, banks of lakes and canals, fields, fallow lands. Plain,
foothills. Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder.
1228. Climacoptera longistylosa (Iljin) Botsch.
Annual. Saline lands. Plain, foothills. Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder.
339
340
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1229. Climacoptera minkvitziae (Korovin) Botsch.
Annual. Saline lands. Plain. Kyzylkum. Fodder.
1230. Climacoptera obtusifolia (Schrenk) Botsch.
Annual. Saline lands. Plain. Mirzachul, Kyzylkum. Fodder.
1231. Climacoptera olgae (Iljin) Botsch.
Annual. Saline lands. Plain. Mirzachul, Kyzylkum. Fodder.
1232. Climacoptera transoxana (Iljin) Botsch.
Annual. Saline lands, roadsides, banks of lakes and canals, fallow lands, fields. Plain. Mirzachul,
Kyzylkum. Fodder.
Genus 372. Halothamnus Jaub. et Spach
1233. Halothamnus glaucus (M. Bieb.) Botsch. [Aellenia glauca (M. Bieb.) Aellen]
Semishrub. Sandy deserts, saline lands. Plain. Kyzylkum. Fodder, medicinal.
1234. Halothamnus iliensis (Lipsky) Botsch. [Aellenia iliensis (Lipsky) Aellen]
Annual. Saline lands, Sandy and clay deserts. Plain. Kyzylkum. Fodder.
1235. Halothamnus subaphyllus (C.A.Mey.) Botsch. [Aellenia subaphylla (C.A. Mey.) Aellen]
Semishrub. Sandy soils, saline lands. Plain. Kyzylkum. Fodder, medicinal, dye.
Genus 373. Girgensohnia Bunge
1236. Girgensohnia oppositiflora (Pall.) Fenzl
Annual. Saline, sandy, clayey and gravelly soils, fields, roadsides, fallow lands, dry riverbeds. Plain,
foothills. Nuratau, Aktau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder.
1237. Girgensohnia diptera Bunge
Annual. Saline lands. Plain. Kyzylkum. Fodder, dye.
Genus 374. Gamanthus Bunge
1238. Gamanthus gamocarpus (Moq.) Bunge
Annual. Saline, sandy, clayey and gravelly soils, fields, roadsides, fallow lands, dry riverbeds. Plain,
foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum. Fodder.
Genus 375. Anabasis (Schrenk) Benth.
1239. Anabasis eriopoda (Schrenk) Paulsen Figure 341
Semishrub. Gravelly and stony slopes, saline lands. Plain, foothills. Nuratau Relic Mountains,
Kyzylkum. Fodder, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 341 Anabasis eriopoda (photography by Natalya Beshko)
1240. Anabasis jaxartica (Bunge) Benth. ex Iljin
Semishrub. Saline lands. Plain. Mirzachul.
1241. Anabasis salsa (Ledeb.) Benth. ex Volkens Figure 342
Semishrub. Saline lands. Plain. Kyzylkum. Fodder, medicinal.
341
342
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 342 Anabasis salsa (photography by Natalya Beshko)
Genus 376. Haloxylon Bunge
1242. Haloxylon ammodendron (C. A. Mey.) Bunge ex Fenzl [Haloxylon aphyllum (Minkw.)
Iljin] Figure 343
Tree. Saline lands, sandy and clay deserts. Plain, foothills. Kyzylkum. Fodder, afforestation.
Figure 343 Haloxylon ammodendron (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1243. Haloxylon persicum Bunge Figure 344
Tree. Sandy deserts. Plain. Kyzylkum. Fodder, afforestation, tanniferous.
Figure 344 Haloxylon persicum (photography by Natalya Beshko)
343
344
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 377. Nanophyton Botsch.
1244. Nanophyton erinaceum (Pall.) Bunge Figure 345
Semishrub. Gravelly and stony slopes, piedmont plains. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains.
Figure 345 Nanophyton erinaceum (photography by Natalya Beshko)
1245. Nanophyton saxatile Botsch. Figure 346
Semishrub. Gravelly and stony slopes, piedmont plains. Plain, foothills. Nuratau, Malguzar.
Figure 346 Nanophyton saxatile (photography by Natalya Beshko)
Genus 378. Halocharis Moq.
1246. Halocharis hispida (Schrenk) Bunge
Annual. Sandy, clayey and gravelly soils, saline lands, fields, fallow lands, banks of canals,
roadsides. Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder,
weed.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 379. Halimocnemis C. A. Mey.
1247. Halimocnemis mollissima Bunge
Annual. Saline lands. Plain. Mirzachul, Kyzylkum. Fodder.
1248. Halimocnemis villosa Kar. & Kir.
Annual. Saline lands, sandy and clay deserts. Plain. Mirzachul, Kyzylkum. Fodder.
Genus 380. Halogeton C. A. Mey.
1249. Halogeton glomeratus (M. Bieb.) Ledeb. Figure 347
Annual. Saline lands, sandy soils, roadsides. Plain. Mirzachul, Kyzylkum.
Figure 347 Halogeton glomeratus (photography by Natalya Beshko)
Family 84. *Portulacaceae
Genus 381. *Portulaca L.
1250. *Portulaca oleracea L.
Annual. Fallow lands, wastelands, banks of rivers and canals, fields. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Food, medicinal,
weed, alkaloid-bearing.
345
346
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 85. Balsaminaceae
Genus 382. Impatiens L.
1251. Impatiens parviflora DC. Figure 348
Annual. Shady wet places, banks of rivers, ravines, stony slopes, screes, gardens. Foothills, montane
zone. Nuratau, Malguzar, North Turkestan. Medicinal, poisonous, weed.
Figure 348 Impatiens parviflora (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Family 86. Primulaceae
Genus 383. Primula L.
1252. Primula algida Adams
Perennial. Alpine meadows, swamps, moraines, stream banks, places near melting snow. Alpine
zone. North Turkestan. Ornamental.
1253. Primula fedtschenkoi Regel Figure 349
Perennial. Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Ornamental.
Figure 349 Primula fedtschenkoi (photography by Natalya Beshko)
1254. Primula iljinskii Al. Fed. ex Schischk. & Bobrov
Perennial. Wet meadows, swamps, banks of rivers. Montane and alpine zones. North Turkestan.
Ornamental.
1255. Primula matthioli (L.) V. A. Richt. [Cortusa turkestanica Losinsk.] Figure 350
Perennial. Stream banks, shady wet places, subalpine and alpine meadows, swamps. Montane and
alpine zones. Malguzar, North Turkestan. Ornamental.
347
348
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 350 Primula matthioli (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1256. Primula olgae Regel
Perennial. Banks of rivers, swamps, subalpine and alpine meadows, moraines. Montane and alpine
zones, Nuratau, Malguzar, North Turkestan. Ornamental.
1257. Primula pamirica Al. Fed. ex Schischk. & Bobrov
Perennial. Alpine meadows, banks of rivers. Alpine zone. North Turkestan. Ornamental.
1258. Primula turkeviczii V. V. Byalt [Primula lactiflora Turkev.]
Perennial. Rocks, places near melting snow. Montane and alpine zones. North Turkestan. Ornamental.
Genus 384. Androsace L.
1259. Androsace dasyphylla Bunge ex Ledeb.
Perennial. Stony slopes, screes, rocks. Alpine zone. North Turkestan.
1260. Androsace maxima L.
Annual. Pebbles, rocks, fine-earth and stony slopes, meadows, fallow lands, fields, roadsides.
Foothills, montane zone. North Turkestan.
1261. Androsace sericea Ovcz.
Perennial. Fine-earth and stony slopes, alpine meadows, moraines, rocks, screes. Alpine zone. North
Turkestan.
Genus 385. Samolus L.
1262. Samolus valerandi L.
Perennial. Wet places, banks of rivers and canals, fallow lands. Plain, foothills. Mirzachul.
Genus 386. Lysimachia L.
1263. Lysimachia dubia Soland. ex Aiton
Perennial. Wet places, banks of rivers and canals, fields. Plain. Mirzachul.
1264. Lysimachia maritima (L.) Galasso, Banfi & Soldano [Glaux maritima L.]
Perennial. Wet places, saline lands, banks of rivers, swamps, meadows. Plain, foothills, montane and
alpine zones. Nuratau, Nuratau Relic Mountains, North Turkestan. Dye.
Genus 387. *Anagallis L.
1265. *Anagallis arvensis L. [Anagallis foemina Mill.] Figure 351
Annual. Banks of rivers and canals, roadsides, gardens, fields, fallow lands, wastelands, pebbles, dry
riverbeds, stony slopes. Plain, foothills, montane zone. Nuratau, Aktau Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Medicinal, poisonous, weed.
349
350
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 351 Anagallis arvensis (above) and Anagallis arvensis subsp. foemina (below) (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Family 87. Ericaceae
Genus 388. Pyrola L.
1266. Pyrola rotundifolia L.
Perennial. Fine-earth, gravelly and stony slopes, screes, banks of rivers. Plain, foothills, montane and
alpine zones. North Turkestan. Medicinal, ornamental.
Family 88. Rubiaceae
Genus 389. Crucianella L.
1267. Crucianella chlorostachys Fisch. & C. A. Mey.
Annual. Pebbles, stony slopes, fallow lands. Plain, foothills, montane zone. Nuratau, Nuratau Relic
Mountains, North Turkestan.
1268. Crucianella exasperata Fisch. & C. A. Mey.
Annual. Fine-earth, gravelly and stony slopes, rocks, screes, pebbles. Montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan.
1269. Crucianella filifolia Regel & Schmalh.
Annual. Fine-earth, gravelly and stony slopes, banks of canals, pebbles, fallow lands. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, North Turkestan.
Genus 390. Asperula L.
1270. Asperula glabrata Tschern.
Semishrub. Stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
1271. Asperula oppositifolia Regel & Schmalh.
Semishrub. Fine-earth, gravelly and stony slopes, rocks, screes. Montane and alpine zones. North
Turkestan.
1272. Asperula setosa Jaub. & Spach.
Annual. Stony slopes, screes, rocks, alpine meadows, wet places near springs. Montane and alpine
zones. Nuratau, Malguzar, North Turkestan.
1273. Asperula trichodes J. Gay ex DC.
Annual. Fine-earth, gravelly and stony slopes, pebbles, dry riverbeds, banks of rivers. Foothills,
montane zone. North Turkestan.
351
352
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 391. Galium L.
1274. Galium aparine L.
Annual. Fine-earth, gravelly and stony slopes, screes, rocks, woodlands, river valleys, gardens,
fields, fallow lands, wastelands. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, dye, fodder, weed.
1275. Galium ceratopodum Boiss.
Annual. Gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar.
1276. Galium humifusum M. Bieb. [Asperula humifusa (M. Bieb.) Besser]
Perennial. Fine-earth, gravelly and stony slopes, pebbles, saline lands, meadows, swamps, banks
of canals, fields, fallow lands, wastelands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Medicinal, dye, fodder, weed.
1277. Galium karakulense Pobed.
Perennial. Wet places, swamps, banks of rivers. Montane zone. Nuratau.
1278. Galium pamiroalaicum Pobed. Figure 352
Perennial. Fine-earth, gravelly and stony slopes, subalpine and alpine meadows, banks of rivers.
Foothills, montane and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, fodder,
meliferous.
Figure 352 Galium pamiroalaicum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1279. Galium pseudorivale Tzvelev [Asperula aparine M. Bieb.]
Perennial. Woodlands, river valleys, gardens, fine-earth and stony slopes. Foothills, montane and
alpine zones. Nuratau, Malguzar, North Turkestan.
1280. Galium setaceum Lam. [Galium decaisnei Boiss.]
Annual. Fine-earth, gravelly and stony slopes, screes, rocks, banks of rivers, fallow lands. Foothills,
montane and alpine zones. Nuratau, Malguzar, North Turkestan.
1281. Galium songaricum Schrenk
Annual. Juniper forests, wet shady places. Montane and alpine zones. North Turkestan.
1282. Galium spurium L.
Annual. Fine-earth, gravelly and stony slopes, screes, rocks, river valleys, woodlands, gardens, fields,
fallow lands. Plain, foothills, montane and alpine zones, Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan.
1283. Galium spurium subsp. ibicinum (Boiss. & Hausskn.) Ehrend. [Galium ibicinum Boiss.
& Hausskn.]
Annual. Fine-earth slopes, river valleys, wet shady places. Montane and alpine zones. North
Turkestan.
1284. Galium tenuissimum M. Bieb.
Annual. Fine-earth, gravelly and stony slopes, pebbles, river valleys, piedmont plains, clay deserts,
fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul. Fodder.
1285. Galium tianschanicum Popov
Perennial. Gravelly and stony slopes, screes, rocks, moraines, pebbles, dry riverbeds. Montane and
alpine zones. Nuratau, Malguzar, NorthTurkestan.
1286. Galium tricornutum Dandy
Annual. Gravelly and stony sloper, screes, rocks, banks of rivers and canals, fields, fallow lands.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan.
1287. Galium turkestanicum Pobed.
Perennial. Fine-earth slopes, meadows, river valleys, woodlands. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Medicinal.
1288. Galium verticellatum Danthoine ex Lam
Perennial. Fine-earth, gravelly and stony slopes, screes, rocks, fields, fallow lands. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
1289. Galium verum L. [Galium ruthenicum Willd.]
Perennial. Fine-earth, gravelly and stony slopes, dry riverbeds, banks of rivers, subalpine and alpine
meadows, banks of rivers. Foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Medicinal, poisonous, dye, ornamental, meliferous.
353
354
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 392. Cruciata Mill.
1290. Cruciata pedemontana (Bellardi) Ehrend.
Annual. Fine-earth slopes, banks of rivers, fallow lands. Foothills, montane zone. Nuratau.
Genus 393. Rubia L.
1291. Rubia regelii Pojark. Figure 353
Semishrub. Gravelly and stony slopes, screes. Montane zone. Nuratau.
Figure 353 Rubia regelii (photography by Natalya Beshko)
Genus 394. Callipeltis Stev.
1292. Callipeltis cucularis (L.) DC.
Annual. Gravelly and stony sloper, screes, rocks, pebbles. Plain, foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Family 89. Gentianaceae
Genus 395. Centaurium Hill
1293. Centaurium erythraea subsp. turcicum (Velen.) Melderis [Centaurium turcicum (Velen.)
Bornm.] Figure 354
Biennial. Banks of rivers and canals, swamps, meadows. Montane zone. Nuratau. Medicinal,
ornamental.
Figure 354 Centaurium erythraea subsp. turcicum (photography by Natalya Beshko)
1294. Centaurium pulchellum (Sw.) Druce
Annual. Meadows, saline lands, wet places, banks of rivers and canals, sandy deserts, fallow
lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Medicinal, ornamental.
1295. Centaurium spicatum (L.) Fritsch
Annual. Meadows, saline lands, wet places, banks of rivers and canals, fallow lands. Plain.
Mirzachul. Medicinal, ornamental.
355
356
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 396. Gentiana L.
1296. Gentiana leucomelaena Maxim.
Annual or biennial. Meadows, swamps, banks of rivers. Montane and alpine zones. Malguzar, North
Turkestan.
1297. Gentiana olivieri Griseb. Figure 355
Perennial. Fine-earth slopes, piedmont plains, clay deserts, banks of rivers, fallow lands. Plain,
foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Medicinal, ornamental.
Figure 355 Gentiana olivieri (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1298. Gentiana prostrata Haenke
Annual or biennial. Subalpine and alpine meadows, swamps, wet places, juniper forests, banks of
rivers. Montane and alpine zones. North Turkestan.
1299. Gentiana riparia Kar. & Kir.
Annual or biennial. Swamps, meadows, banks of rivers. Montane and alpine zones. North Turkestan.
Genus 397. Gentianella Moench
1300. Gentianella sibirica (Kuzn.) Holub [Gentiana sibirica (Kuzn.) Grossh.]
Annual or biennial. Alpine meadows, moraines, pebbles, shady wet places, banks of rivers. Montane
and alpine zones. North Turkestan.
1301. Gentianella turkestanorum (Gand.) Holub [Gentiana turkestanorum Gand.]
Annual or biennial. Meadows, swamps, banks of rivers, woodlands, wet places near springs.
Montane and alpine zones. Malguzar, North Turkestan. Medicinal, ornamental.
Genus 398. Gentianopsis Ma
1302. Gentianopsis barbata (Froel.) Ma [Gentiana barbata Froel.].
Annual or biennial. Meadows, fine-earth slopes, moraines, swamps, banks of rivers. Montane and
alpine zones. North Turkestan. Medicinal, ornamental.
Genus 399. Lomatogonium A. Braun
1303. Lomatogonium carinthiacum (Wulfen) A. Braun
Annual. Wet stony slopes, alpine meadows, swamps, moraines, banks of streams. Alpine zone. North
Turkestan. Medicinal, meliferous.
Genus 400. Swertia L.
1304. Swertia lactea Bunge
Perennial. Banks of rivers, wet stony slopes and rocks, woodlands, subalpine meadows, swamps,
moraines. Montane and alpine zones. Malguzar, North Turkestan. Ornamental.
357
358
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 90. Apocynaceae
Genus 401. Cynanchum L.
1305. Cynanchum acutum subsp. sibiricum (Willd.) Rech. f. [Cynanchum sibiricum
Willd.] Figure 356
Perennial. Riparian forests, banks of rivers, lakes and canals, wet stony slopes, pebbles, saline lands,
sandy deserts. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Medicinal, weed.
Figure 356 Cynanchum acutum subsp. sibiricum (photography by Natalya Beshko)
Genus 402. Trachomitum Woodson
1306. Trachomitum scabrum (Russanov) Pobed. Figure 357
Perennial. Riparian forests, banks of rivers, lakes and canals, saline lands. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Medicinal, poisonous, industrial, ornamental, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 357 Trachomitum scabrum (photography by Natalya Beshko)
Family 91. Boraginaceae
Genus 403. Heliotropium L.
1307. Heliotropium arguzioides Kar. & Kir.
Perennial. Sandy deserts. Kyzylkum. Poisonous, fodder, meliferous.
1308. Heliotropium bogdanii Czukav.
Annual. Stony banks of rivers, pebbles, conglomerates. Foothills. North Turkestan.
1309. Heliotropium dasycarpum Ledeb.
Perennial. Sandy, clayey and gravelly soils. Plain, foothills. Nuratau, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Poisonous, fodder, weed.
1310. Heliotropium ellipticum Ledeb.
Annual. Pebbles, dry riverbeds, gravelly soils, fields, fallow lands. Plain, foothills. Nuratau.
Poisonous, weed.
1311. Heliotropium lasiocarpum Fisch. & C. A. Mey.
Annual. Sandy, clayey and gravelly soils, wastelands, roadsides, fallow lands. Foothills. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Poisonous,
weed.
359
360
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1312. Heliotropium olgae Bunge
Annual. Sandy, clayey and gravelly soils, pebbles. Plain, foothills. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Poisonous.
Genus 404. Lithospermum L.
1313. Lithospermum officinale L.
Perennial. Fine-earth slopes, river valleys, banks of canals. Plain, foothills, montane zone. Nuratau,
Malguzar, North Turkestan, Mirzachul. Medicinal, dye, weed, fodder.
Genus 405. Buglossoides Moench
1314. Buglossoides arvensis (L.) J. M. Johnst.
Annual. Fine-earth, gravelly and stony slopes, river valleys, fallow lands, fields. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Dye, weed.
Genus 406. Arnebia Forsk.
1315. Arnebia coerulea Schipcz. Figure 358
Annual. Fine-earth, gravelly and stony slopes, sandy deserts, wastelands. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Dye, fodder, ornamental.
Figure 358 Arnebia coerulea (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1316. Arnebia decumbens (Vent.) Coss. & Kralik. Figure 359
Annual. Sandy and clay deserts, gravelly and stony slopes, fields, fallow lands, banks of canals.
Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Dye, fodder, weed.
Figure 359 Arnebia decumbens (photography by Natalya Beshko)
361
362
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1317. Arnebia euchroma (Royle) I. M. Johnst. [Macrotomia euchroma (Royle) Paulsen]
Perennial. Stony slopes, rocks. Alpine zone. North Turkestan. Dye.
1318. Arnebia obovata Bunge
Perennial. Stony slopes. Montane zone. Malguzar, North Turkestan. Dye, fodder.
1319. Arnebia transcaspica Popov Figure 360
Annual. Sandy, clayey and gravelly soils, stony slopes. Plain, foothills. Nuratau, Nuratau Relic
Mountains, Kyzylkum. Dye, fodder.
Figure 360 Arnebia transcaspica (photography by Natalya Beshko)
Genus 407. Onosma L.
1320. Onosma dichroanta Boiss.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Medicinal, dye, meliferous.
1321. Onosma maracandica Zakirov
Perennial. Rocks, screes, stony slopes. Montane zone. North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 408. Echium L.
1322. Echium biebersteinii Lacaita
Biennial. Fields, wastelands, fallow lands, roadsides, fine-earth and stony slopes, river valleys. Foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, dye, weed, meliferous.
1323. *Echium vulgare L.
Biennial. Wastelands, fallow lands, roadsides, fine-earth and stony slopes. Foothills. Malguzar, North
Turkestan. Medicinal, dye, weed, meliferous.
Genus 409. *Anchusa L.
1324. *Anchusa arvensis subsp. orientalis (L.) Nordh. [Lycopsis orientalis L.]
Annual. Fine-earth, gravelly and stony slopes, wastelands, fallow lands, fields, roadsides. Plains,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed.
1325. *Anchusa azurea Mill. [Anchusa italica Retz.] Figure 361
Perennial. Ravines, river valleys, fine-earth, gravelly and stony slopes, pebbles, wastelands, fallow
lands, fields, roadsides, banks of canals. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Dye, medicinal, weed, meliferous,
ornamental.
Figure 361 Anchuza azurea (photography by Natalya Beshko)
363
364
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 410. Nonea Medik.
1326. Nonea caspica (Willd.) G. Don. Figure 362
Annual. Sandy and clay deserts, gravelly and stony slopes, roadsides, fallow lands. Plain, foothills,
montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed.
Figure 362 Nonea caspica (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1327. Nonea melanocarpa Boiss.
Annual. Fields, fallow lands, fields. North Turkestan, Mirzachul. Weed.
1328. Nonea pulla (L.) DC.
Perennial. Fallow lands, wastelands, roadsides, river valleys. North Turkestan (this species can
be found in the Zaamin National Park, in basin of the river Uriklisay, because it was reported from
the neighbouring area of Tajikistan, from the basin of the river Kusavli). Medicinal, meliferous.
Genus 411. Myosotis L.
1329. Myosotis alpestris F. W. Schmidt
Perennial. Subalpine and alpine meadows, gravelly and stony slopes, moraines. Montane and alpine
zones, North Turkestan. Fodder, ornamental.
1330. Myosotis laxa subsp. caespitosa (Schultz) Hyl. ex Nordh. [Myosotis caespitosa Schultz]
Perennial. River valleys, swamps, wet places near springs. Nuratau, Nuratau Relic Mountains,
Malguzar, North Turkestan.
1331. Myosotis refracta Boiss.
Annual. Shady wet places, fine-earth, gravelly and stony slopes, pebbles. Foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Weed.
1332. Myosotis stricta Link. ex Roem. & Schult. [Myosotis micrantha Pall. & Lehm.]
Annual. River valleys, fine-earth, gravelly and stony slopes, pebbles, rocks, fallow lands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Weed.
Genus 412. Lappula Moench
1333. Lappula brachycentra (Ledeb.) Gürke
Annual or biennial. Gravelly and stony slopes. Foothills, montane and alpine zones. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan.
1334. Lappula microcarpa (Ledeb.) Gürke
Annual or biennial. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes,
fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Weed.
1335. Lappula myosotis Moench
Annual or biennial. Wastelands, roadsides, fallow lands, surroundings of wells and sheepyards, fineearth, gravelly and stony slopes. Plain, foothills. Nuratau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Weed.
1336. Lappula nuratavica Nabiev & Zakirov Figure 363
Annual. Stony slopes, rocks. Montane zone. Nuratau. Endemic to the Nuratau ridge. UzbRDB 2.
365
366
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 363 Lappula nuratavica (photography by Natalya Beshko)
1337. Lappula occultata Popov
Annual. Gravelly and stony slopes, screes, rocks. Foothills, montane zone. Malguzar, North Turkestan.
1338. Lappula marginata (M. Bieb.) Gürke [Lappula patula (Lehm.) Asch. ex Gurke; Lappula
semiglabra (Ledeb.) Gürke]
Annual. Gardens, fields, wastelands, roadsides, fallow lands, pebbles, fine-earth and gravelly slopes,
river valleys, sandy and clay deserts. Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Weed.
1339. Lappula sarawschanica (Lipsky) Nabiev
Perennial. Stony slopes, screes, rocks. Montane and alpine zones. Malguzar, North Turkestan.
1340. Lappula semialata Popov
Biennial. Gravelly and stony slopes. Foothills, montane and alpine zones. Malguzar, North
Turkestan.
1341. Lappula sessiliflora Gürke
Annual. Gravelly and stony slopes. Foothills, montane zone. Nuratau, Nuratau Relic Mountains.
1342. Lappula sinaica (A. DC.) Asch. & Schweinf.
Annual. Fine-earth, gravelly and stony slopes, rocks, fallow lands. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Weed.
1343. Lappula spinocarpos (Forssk.) Asch. ex Kuntze
Annual. Sandy and clay deserts, piedmont plains, gravelly and stony slopes. Plain, foothills. North
Turkestan, Mirzachul, Kyzylkum.
1344. Lappula squarrosa (Retz.) Dumort. [Lappula consanguinea (Fisch. & C. A. Mey.) Gürke]
Annual or biennial. Fields, fallow lands, wastelands. Plain, foothills, montane and alpine zones.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Weed.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 413. Asperugo L.
1345. Asperugo procumbens L.
Annual. Fields, river valleys, wastelands, roadsides, settlements, surroundings of wells and
sheepyards, gardens, banks of rivers, lakes and canals, fallow lands, shady wet places. Plain,
foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Medicinal, weed.
Genus 414. Heterocaryum DC.
1346. Heterocaryum laevigatum (Kar. & Kir.) A. DC.
Annual. Gravelly and stony slopes, river valleys. Foothills. Nuratau. Weed.
1347. Heterocaryum macrocarpum Zakirov
Annual. Fine-earth and stony slopes, wastelands, roadsides, fallow lands. Foothills, montane zone.
Malguzar, North Turkestan. Weed.
1348. Heterocaryum subsessile Vatke
Annual. Gravelly slopes, wastelands. Foothills, montane zone. Nuratau, Aktau, Malguzar, North
Turkestan, Mirzachul. Weed.
1349. Heterocaryum szovitsianum (Fisch. & C. A. Mey.) A. DC. [Heterocaryum rigidum A. DC.]
Annual. Fine-earth, gravelly and stony slopes, wastelands, fallow lands, roadsides. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Weed.
Genus 415. Rochelia Rchb.
1350. Rochelia bungei Trautv.
Annual. Fine-earth, gravelly and stony slopes, dry riverbeds, sandy deserts, fields, fallow lands.
Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Kyzylkum. Weed.
1351. Rochelia cardiosepala Bunge
Annual. Fine-earth, gravelly and stony slopes, fallow lands, fields, dry riverbeds, sandy deserts.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Weed.
1352. Rochelia disperma subsp. retorta (Pall.) Kotejowa [Rochelia retorta (Pall.) Lipsky]
Annual. Clay deserts, river valleys, piedmont plains, fine-earth, gravelly and stony slopes, fields,
fallow lands, wastelands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Weed.
1353. Rochelia leiocarpa Ledeb.
Annual. Fine-earth, gravelly and stony slopes, pebbles. Plain, foothills, montane zone. Nuratau,
367
368
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Weed.
1354. Rochelia peduncularis Boiss.
Annual. Fine-earth, gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Aktau,
Malguzar, North Turkestan. Weed.
Genus 416. Rindera Pall.
1355. Rindera tetraspis Pall. Figure 364
Perennial. Rocks, fine-earth, gravelly and stony slopes, piedmont plains. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Figure 364 Rindera tetraspis (photography by Natalya Beshko)
Genus 417. Hackelia Opiz.
1356. Hackelia uncinata (Benth.) C. E. C. Fisch. [Paracaryum glochidiatum (A. DC.) Benth. &
Hook. f.]
Annual. Clayey and gravelly soils. Mirzachul.
Genus 418. Mattiastrum Brand
1357. Mattiastrum himalayense (Klotzsch) Brand [Paracaryum himalayense (Klotzsch) C. B.
Clarke]
Perennial. Stony slopes, screes. Montane and alpine zones. North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 419. Lindelofia Lehm.
1358. Lindelofia olgae Brand
Perennial. Fine-earth, gravelly and stony slopes. Alpine zone. North Turkestan. Dye.
1359. Lindelofia macrostyla (Bunge) Popov Figure 365
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Dye, weed.
Figure 365 Lindelofia macrostyla (photography by Natalya Beshko)
1360. Lindelofia stylosa (Kar. & Kir.) Brand
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. North Turkestan.
Ornamental.
Genus 420. Solenanthus Ledeb.
1361. Solenanthus circinatus Ledeb.
Perennial. Fine-earth, gravelly and stony slopes, shady wet places. Montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental, meliferous.
369
370
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 421. Kuschakewiczia Regel & M. Smirn.
1362. Kuschakewiczia turkestanica Regel & Smirn. [Solenanthus turkestanicus (Regel &
Smirn.) Kuzn.]
Perennial. Clayey and fine-earth soils. Foothills. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Meliferous.
Genus 422. *Cynoglossum L.
1363. *Cynoglossum creticum Mill.
Annual or biennial. Banks of rivers and canals, fallow lands, fields, wastelands. Plain, foothills,
montane zone. Nuratau, North Turkestan. Medicinal, meliferous, weed.
Genus 423. Trichodesma R. Br.
1364. Trichodesma incanum (Bunge) A. DC.
Perennial. Fine-earth, gravelly and stony slopes, pebbles, river valleys, fields, fallow lands,
settlements. Foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Weed, poisonous, medicinal.
Family 92. Convolvulaceae
Genus 424. Cuscuta L.
1365. Cuscuta approximata Bab. [Cuscuta cupulata Engelm.]
Annual. Fields, fallow lands, gardens. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Poisonous. Parasite. Host plants are Medicago
sativa, Setaria viridis, Cynodon dactylon, Convolvulus arvensis, Lactuca scariola. Quarantine
invasive weed.
1366. Cuscuta brevistyla A. Braun ex A. Rich.
Annual. Fine-earth, gravelly and stony slopes. Foothills, montane and alpine zones. Nuratau,
Malguzar, North Turkestanl. Poisonous. Parasite. Host plants are different wild herbs and semishrubs.
Quarantine invasive weed.
1367. *Cuscuta campestris Yunck.
Annual. Gardens, fields, vineyards. Plain, foothills. Nuratau, Aktau, Malguzar, North Turkestan,
Mirzachul. Parasite. Host plants are different crops, vegetables and ornamentals, as well as weeds
MAGNOLIOPSIDA
[EUDICOTS]
and wild species of Polygonum, Zygophyllum, etc. Quarantine invasive weed.
1368. Cuscuta europaea L.
Annual. Woodlands, wet places, banks of rivers, lakes and canals, fine-earth, gravelly and stony
slopes. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Medicinal, poisonous. Parasite. Host plants are different
wild herbs, semishrubs and shrubs (Glycyrrhiza glabra, Ferula sp., Artemisia dracunculus, etc.).
Quarantine invasive weed.
1369. Cuscuta lehmanniana Bunge
Annual. Riparian forests, banks of rivers, lakes and canals, gardens, fields, vineyards. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal. Parasite. Host plants are different cultivated and wild herbs, semishrubs,
shrubs and trees (more than 200 species). Quarantine invasive weed.
1370. Cuscuta monogyna Vahl.
Annual. Woodlands, gardens, vineyards, river valleys, banks of lakes and canals. Plain, foothills,
montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Parasite. Host plants are different cultivated and wild herbs, shrubs and trees (species of
genera Rhamnus, Salix, Ziziphus, Paliurus, Tamarix, Berberis, Rosa, Fraxinus, Pistacia, Vitis, etc.).
Quarantine invasive weed.
1371. Cuscuta pedicellata Ledeb.
Annual. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes. Plain,
foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal. Parasite. Host plants are different wild herbs and shrubs (Poa bulbosa,
Peganum harmala, Alhagi sp., Trifolium sp., Trigonella sp., Galium sp., Artemisia sp., etc.).
Quarantine invasive weed.
1372. Cuscuta pellucida Butkov
Annual. Piedmont plains, fine-earth, gravelly and stony slopes. Plain, foothills, montane zone.
Nuratau, Nuratau Relic Mountains, North Turkestan. Parasite. Host plants are different wild
herbs and semishrubs (Alhagi sp., Artemisia sp., Origanum sp., etc.). Quarantine invasive
weed.
Genus 425. Convolvulus L.
1373. *Convolvulus arvensis L.
Perennial. A weed widespread in different habitats. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, alkaloidbearing, weed, fodder, meliferous.
1374. Convolvulus fruticosus Pall. Figure 366
Semishrub. Sandy deserts. Plain. Mirzachul, Kyzylkum. Meliferous.
371
372
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 366 Convolvulus fruticosus (photography by Natalya Beshko)
1375. Convolvulus hamadae (Vved.) V. Petrov
Semishrub. Sandy and clay deserts, gravelly and stony slopes of relic mountains. Plain, foothills.
Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder, meliferous.
1376. Convolvulus korolkowii Regel & Schmalh.
Perennial. Sandy deserts. Plain. Kyzylkum. Meliferous.
1377. Convolvulus lineatus L. Figure 367
Perennial. Fine-earth, gravelly and stony slopes, fallow lands, roadsides, banks of rivers and canals,
wet places. Foothills, montane and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan. Fodder,
weed, meliferous.
Figure 367 Convolvulus lineatus (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1378. Convolvulus pilosellifolius Desr.
Perennial. Fallow lands, clayey and skeleton soils, banks of canals. Plain. Mirzachul. Fodder,
meliferous.
1379. Convolvulus pseudocantabricus Schrenk ex Fisch. & C. A. Mey.
Perennial. Piedmont plains, fine-earth, gravelly and stony slopes. Foothills, montane and alpine
zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, poisonous,
meliferous.
1380. Convolvulus subhirsutus Regel & Schmalh.
Perennial. Piedmont plains, fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Medicinal, fodder, weed, meliferous.
Genus 426. Calystegia R. Br.
1381. Calystegia sepium (L.) R. Br.
Perennial. Riparian forests, gardens, banks of rivers and canals, stony wet places, settlements. Plain,
foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, poisonous, ornamental, meliferous.
Family 93. Solanaceae
Genus 427. Solanum L.
1382. *Solanum americanum Mill. (Solanum nigrum L.)
Annual. Gardens, fields, fallow lands, settlements, river valleys, wastelands. Plain. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, alkaloid-bearing, oil, poisonous, food, weed, meliferous.
1383. Solanum asiae-mediae Pojark.
Semishrub. Riparian forests, ravines, stream banks. Plain, foothills, montane zone. Nuratau,
Malguzar, North Turkestan, Mirzachul.
1384. Solanum kitagawae Schonb.-Tem.
Semishrub. Riparian forests, stony slopes, ravines, banks of rivers, lakes and canals. Plain, foothills,
montane zone. Nuratau, North Turkestan, Mirzachul.
1385. Solanum olgae Pojark.
Annual. Fields, fallow lands, wastelands, gardens, banks of rivers, lakes and canals, stony slopes.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum.
373
374
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 428. Lycium L.
1386. Lycium dasystemum Pojark.
Shrub. Sandy and clay deserts, saline lands, banks of rivers, lakes and canals. Plain, foothills.
Nuratau Relic Mountains, Mirzachul, Kyzylkum. Medicinal, oil, afforestation, meliferous.
1387. Lycium ruthenicum Murray Figure 368
Shrub. Sandy and clay deserts, saline lands, banks of rivers, lakes and canals. Nuratau Relic
Mountains, Mirzachul, Kyzylkum. Medicinal, oil, afforestation, weed, meliferous.
Figure 368 Lycium ruthenicum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 429. Hyoscyamus L.
1388. *Hyoscyamus niger L. Figure 369
Biennial. Fine-earth, gravelly and stony slopes, wastelands, fallow lands, fields, gardens, settlements,
roadsides. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, poisonous, insecticide, weed, alkaloid-bearing.
Figure 369 Hyoscyamus niger (photography by Natalya Beshko)
1389. Hyoscyamus pusillus L. Figure 370
Annual. Fine-earth, gravelly and stony slopes, screes, rocks, sandy and clay deserts, saline lands,
wastelands, fallow lands. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Poisonous, alkaloid-bearing, weed.
375
376
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 370 Hyoscyamus pusillus (photography by Natalya Beshko)
Genus 430. *Datura L.
1390. *Datura stramonium L.
Annual. Settlements, wastelands, fields, fallow lands, roadsides, banks of canals. Nuratau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, poisonous,
insecticide, weed, alkaloid-bearing.
Genus 431. *Nicandra Adans.
1391. *Nicandra physaloides (L.) Gaertn
Annual. Settlements, wastelands, fields, fallow lands, roadsides, banks of canals. Mirzachul, North
MAGNOLIOPSIDA
[EUDICOTS]
Turkestan. Medicinal, oil, fodder, weed.
Family 94. Oleaceae
Genus 432. Fraxinus L.
1392. Fraxinus sogdiana Bunge
Tree. River valleys, fine-earth slopes. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
IUCN NT. Industrial, afforestation, ornamental, medicinal.
1393. *Fraxinus angustifolia subsp. syriaca (Boiss.) Yalt. [Fraxinus syriaca Boiss.]
Tree. River valleys. Montane zone. Nuratau. Industrial, afforestation, ornamental, medicinal.
Family 95. Plantaginaceae
Genus 433. *Kickxia Dumort.
1394. *Kickxia elatine (L.) Dumort.
Annual. Fine-earth slopes, pebbles, riparian forests, fields, fallow lands. Plain, foothills, montane
zone. North Turkestan, Mirzachul. Medicinal.
Genus 434. Veronica L.
1395. *Veronica anagallis-aquatica L.
Perennial. Wet places near spings, banks of rivers and canals. Plain, foothills, montane zone.
Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, meliferous.
1396. Veronica anagalloides Guss
Perennial. Banks of rivers, lakes and canals, swamps, meadows, fallow lands, pebbles, saline lands.
Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
1397. Veronica arguteserrata Regel & Schmalh.
Perennial. Fine-earth and stony slopes, screes, rocks, piedmont plains, fields, fallow lands. Plain,
foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul.
1398. Veronica arvensis L.
Annual. Fine-earth slopes, wet meadows, river valleys, roadsides, fields, fallow lands. Foothills,
montane zone. North Turkestan. Medicinal.
377
378
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1399. Veronica beccabunga L.
Perennial. Banks of rivers and canals, swamps, wet places near springs. Plain, foothills, montane and
alpine zones. Nuratau, Malguzar, North Turkestan. Medicinal.
1400. Veronica biloba Schreb. ex L.
Annual. Sandy and clay deserts, fine-earth, gravelly and stony slopes, pebbles, rocks, screes,
moraines. Plain, foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan,
Mirzachul, Kyzylkum.
1401. Veronica campylopoda Boiss.
Annual. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, rocks,
pebbles, fields, fallow lands, saline lands, river valleys. Plain, foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
1402. Veronica capillipes Nevski
Annual. Fine-earth, gravelly and stony slopes, screes, rocks, river valleys. Foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
1403. Veronica cardiocarpa (Kar. & Kir.) Walp.
Annual. Sandy and clay deserts, fine-earth, gravelly and stony slopes, rocks, screes. Plain, foothills,
montane zone. Nuratau, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
1404. Veronica hederifolia L.
Annual. Settlements, fields, fallow lands, gardens, wastelands, roadsides, river valleys, banks of
canals, stony slopes. Plain, foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed.
1405. Veronica intercedens Bornm.
Annual. Fine-earth and stony slopes, screes. Foothills, montane and alpine zones. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan.
1406. Veronica oxycarpa Boiss.
Perennial. Banks of rivers, lakes and canals, wet places near springs, gardens, meadows. Plain,
foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul.
1407. *Veronica persica Poir.
Annual. Settlements, fields, fallow lands, wastelands, gardens, banks of canals, river valleys. Plain,
foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Weed.
1408. *Veronica polita Fr.
Annual. Settlements, banks of rivers and canals, fields, fallow lands, fine-earth slopes, pebbles, river
valleys. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum.
1409. Veronica verna L.
Annual. Roadsides, pebbles, meadows, gravelly and fine-earth slopes. Foothills, montane and alpine
zones. Nuratau, North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 435. Lagotis Gaertn.
1410. Lagotis korolkowii (Regel & Schmalh.) Maxim. Figure 371
Perennial. Fine-earth and stony slopes, alpine meadows. Montane and alpine zones. North Turkestan.
Figure 371 Lagotis korolkowii (photography by Natalya Beshko)
Genus 436. Plantago L.
1411. Plantago gentianoides Sm.
Perennial. Swamps, stream banks. Alpine zone. North Turkestan.
1412. Plantago lagocephala Bunge
Annual. Sandy and clay deserts, saline lands. Plain. Kyzylkum.
1413. Plantago lanceolata L.
Perennial. Fine-earth slopes, river valleys, banks of lakes and canals, roadsides, wastelands, gardens,
wet places. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, fodder.
1414. Plantago major L.
Biennial or perennial. Wet places, swamps, meadows, banks of rivers, lakes and canals, riparian
forests, roadsides, settlements, fields, gardens. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, fodder.
379
380
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 437. Hippuris L.
1415. Hippuris vulgaris L.
Perennial. Standing and slowly flowing water. Nuratau Relic Mountains, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, fodder.
Family 96. Scrophulariaceae
Genus 438. Verbascum L.
1416. Verbascum blattaria L.
Annual or biennial. Fine-earth slopes, piedmont plains, sandy and clay deserts, banks of rivers,
lakes and canals, saline lands, riparian forests, fallow lands, fields, gardens, meadows. Plain,
foothills. Nuratau, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, meliferous, weed,
insecticide.
1417. Verbascum erianthum Benth. [Verbascum bactrianum Bunge]
Biennial. Fields, fallow lands, banks of rivers and canals, saline lands, stony slopes, pebbles.
Plain, foothills, montane and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul.
Medicinal, meliferous, insecticide.
1418. Verbascum songaricum Schrenk Figure 372
Biennial. Sandy deserts, roadsides, banks of canals, dry riverbeds, river valleys, ravines, fallow lands,
fields, fine-earth, gravelly and stony slopes. Plain, foothills, montane and alpine zones. Nuratau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, meliferous, insecticide.
Figure 372 Verbascum songaricum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1419. Verbascum turkestanicum Franch.
Annual or biennial. Fine-earth and stony slopes. Montane zone. North Turkestan.
Genus 439. Linaria Mill.
1420. Linaria popovii Kuprian. Figure 373
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Malguzar,
North Turkestan. Medicinal.
Figure 373 Linaria popovii (photography by Natalya Beshko)
1421. Linaria sessilis Kuprian.
Perennial. Stony slopes, screes, pebbles. Montane and alpine zones. Malguzar, North Turkestan.
Genus 440. Scrophularia L.
1422. Scrophularia gontscharovii Gorschk.
Perennial. Stony slopes. Montane and alpine zones. North Turkestan.
1423. Scrophularia griffithii Benth. ex DC.
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. North Turkestan.
1424. Scrophularia heucheriiflora Schrenk
Perennial. Stony and gravelly slopes, meadows, river valleys. Foothills, montane zone. North Turkestan.
1425. Scrophularia incisa Weinm.
Perennial. Stony slopes, pebbles, screes. Plain, foothills, montane and alpine zones. North Turkestan.
Medicinal.
1426. Scrophularia integrifolia Pavlov
Perennial. Gravelly and stony slopes, rocks. Montane zone. North Turkestan.
381
382
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1427. Scrophularia kiriloviana Schischk.
Perennial. Fine-earth, gravelly and stony slopes, screes, ravines, river valleys. Montane zone. North
Turkestan.
1428. Scrophularia leucoclada Bunge
Perennial. Sandy soils, stony and gravelly slopes, dry riverbeds, saline lands. Plain, foothills,
montane zone. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum.
1429. Scrophularia scoparia Pennell
Perennial. Stony and gravelly slopes, pebbles. Foothills, montane and alpine zones. Nuratau,
Malguzar, North Turkestan.
1430. Scrophularia striata Boiss.
Perennial. Fine-earth, gravelly and stony slopes, dry riverbeds, pebbles. Foothills, montane zone.
North Turkestan.
1431. Scrophularia umbrosa Dumort.
Perennial. Fine-earth, gravelly and stony slopes, stream banks, wet places, meadows. Montane zone.
Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, poisonous, meliferous.
1432. Scrophularia vvedenskyi Bondarenko & Filatova
Biennial. Rocks, fine-earth, gravelly and stony slopes, pebbles. Foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan.
1433. Scrophularia xanthoglossa Boiss.
Perennial. Rocks, stony slopes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Family 97. Lentibulariaceae
Genus 441. Utricularia L.
1434. Utricularia minor L.
Perennial. Standing and slowly flowing water. Montane zone. North Turkestan.
Family 98. Verbenaceae
Genus 442. Verbena L.
1435. Verbena officinalis L.
Perennial. River valleys, ravines, gardens, meadows, banks of rivers, lakes and canals, fallow
lands, fine-earth slopes, roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, poisonous, weed.
MAGNOLIOPSIDA
[EUDICOTS]
Family 99. Lamiaceae [Labiatae]
Genus 443. Teucrium L.
1436. Teucrium scordium subsp. scordioides (Schreb.) Arcang. [Teucrium scordioides Schreb.]
Perennial. Wet places, banks of rivers, lakes and canals. Plain. Mirzachul. Medicinal, essential oil,
poisonous, insecticide.
Genus 444. Scutellaria L.
1437. Scutellaria adenostegia Briq.
Perennial. Fine-earth and stony slopes, rocks, screes, dry riverbeds. Montane zone. Nuratau, North
Turkestan.
1438. Scutellaria comosa Juz. Figure 374
Semishrub. Stony slopes, rocks, screes. Montane zone. Nuratau, Aktau, Malguzar, North Turkestan.
Figure 374 Scutellaria comosa (photography by Natalya Beshko)
1439. Scutellaria cordifrons Juz. Figure 375
Semishrub. Stony slopes, rocks, screes, ravines. Montane and alpine zones. Malguzar, North
Turkestan.
383
384
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 375 Scutellaria cordifrons (photography by Natalya Beshko)
1440. Scutellaria galericulata L.
Perennial. Banks of rivers, riparian forests, fields, wet slopes. Plain, foothills, montane zone.
Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, dye, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
1441. Scutellaria glabrata Vved.
Semishrub. Stony slopes, screes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
1442. Scutellaria immaculata Nevski ex Juz. Figure 376
Semishrub. Stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
Figure 376 Scutellaria immaculata (photography by Natalya Beshko)
1443. Scutellaria intermedia Popov
Semishrub. Stony and gravelly slopes, rocks. Montane zone. Aktau, Malguzar, North Turkestan.
1444. Scutellaria oxystegia Juz.
Semishrub. Fine-earth and stony slopes, rocks, screes, banks of rivers. Montane zone. Nuratau,
Aktau, Malguzar. Essential oil, meliferous, alkaloid-bearing.
385
386
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1445. Scutellaria phyllostachya Juz.
Semishrub. Stony slopes, banks of rivers. Montane and alpine zones. North Turkestan.
1446. Scutellaria ramosissima Popov Figure 377
Semishrub. Stony slopes, screes, dry riverbeds. Montane zone. Nuratau, Malguzar, North Turkestan.
Figure 377 Scutellaria ramosissima (photography by Natalya Beshko)
1447. Scutellaria schachristanica Juz.
Perennial. Gravelly and stony slopes, screes. Alpine zone. Malguzar, North Turkestan.
1448. Scutellaria squarrosa Nevski
Semishrub. Rocks, stony slopes, screes. Montane zone. Malguzar, North Turkestan.
Genus 445. Drepanocaryum Pojark.
1449. Drepanocaryum sewerzowii (Regel) Pojark.
Annual. Fine-earth, gravelly and stony slopes, rocks, screes, shady places, gardens. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Weed.
Genus 446. Marrubium L.
1450. Marrubium anisodon K. Koch
Perennial. Fine-earth and stony slopes, rocks, screes, pebbles, fallow lands, roadsides, banks of
canals, wastelands, fields, surroundings of wells and sheepyards. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Medicinal, essential oil, meliferous, weed.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 447. Sideritis L.
1451. Sideritis montana L.
Annual. Fine-earth and stony slopes, screes, rocks, pebbles, fields. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, essential oil, food,
meliferous.
Genus 448. Lophanthus Adans.
1452. Lophanthus ouroumitanensis (Franch.) Kochk. & Zuckerw.
Perennial. Fine-earth and stony slopes, river valleys, juniper forests. Montane zone. Malguzar, North
Turkestan. Essential oil, meliferous.
1453. Lophanthus schtschurowskianus (Regel) Lipsky Figure 378
Perennial. Fine-earth, gravelly and stony slopes, dry riverbeds. Montane zone. Nuratau, Malguzar,
North Turkestan. Essential oil, meliferous.
Figure 378 Lophanthus schtschurowskianus (photography by Natalya Beshko)
1454. Lophanthus subnivalis Lipsky
Perennial. Rocks, stony slopes, pebbles, moraines. Montane and alpine zone. North Turkestan.
Essential oil, meliferous.
Genus 449. Nepeta L.
1455. Nepeta alatavica Lipsky
Perennial. Stony slopes, screes, pebbles. Montane and alpine zones. North Turkestan. Essential oil,
meliferous.
1456. Nepeta bracteata Benth.
Annual. Stony and gravelly slopes. Montane and alpine zones. North Turkestan. Essential oil,
387
388
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
meliferous.
1457. Nepeta cataria L.
Perennial. Meadows, banks of rivers and canals, stony slopes. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Essential oil, meliferous.
1458. Nepeta kokanica Regel
Perennial. Stony slopes, screes, rocks. Alpine zone. North Turkestan. Essential oil, meliferous.
1459. Nepeta mariae Regel
Perennial. Stony slopes, screes, banks of rivers. Montane and alpine zones. Malguzar, North
Turkestan. Essential oil, meliferous.
1460. Nepeta maussarifii Lipsky
Perennial. Stony slopes. Montane zone. North Turkestan. Essential oil, meliferous.
1461. Nepeta micrantha Bunge
Annual. Gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Essential oil, meliferous.
1462. Nepeta nuda L. (Nepeta pannonica L.)
Perennial. Meadows, pebbles, fine-earth slopes. Foothills, montane zone. North Turkestan. Essential
oil, medicinal, meliferous.
1463. Nepeta olgae Regel
Perennial. Stony slopes, pebbles, rocks. Foothills, montane zone. Nuratau, Aktau, Malguzar, North
Turkestan. Essential oil, medicinal, meliferous.
1464. Nepeta podostachys Benth.
Perennial. Stony slopes, rocks, screes, pebbles. Montane and alpine zones. Malguzar, North
Turkestan. Essential oil, meliferous.
1465. Nepeta pungens (Bunge) Benth.
Annual. Stony and gravelly slopes, dry riverbeds. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Essential oil, meliferous.
1466. Nepeta saturejoides Boiss.
Annual. Stony slopes, rocks, screes, pebbles. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Essential oil, meliferous.
1467. Nepeta ucranica L.
Perennial. Pebbles, banks of rivers. Montane zone. Malguzar, North Turkestan. Essential oil,
meliferous.
Genus 450. Dracocephalum L.
1468. Dracocephalum diversifolium Rupr.
Perennial. Fine-earth and stony slopes, screes, pebbles. Montane and alpine zones. Malguzar, North
Turkestan. Medicinal, meliferous.
1469. Dracocephalum imberbe Bunge
Perennial. Rocks, screes, moraines, stony slopes, wet places near melting snow. Alpine zone. North
MAGNOLIOPSIDA
[EUDICOTS]
Turkestan.
1470. Dracocephalum nodulosum Rupr.
Semishrub. Rocks, stony slopes, screes, moraines. Montane and alpine zones, Nuratau, North
Turkestan. Medicinal, meliferous.
1471. Dracocephalum nuratavicum Adylov Figure 379
Semishrub. Stony and gravelly slopes, rocks. Montane zone. Nuratau. Ornamental. Endemic to the
Nuratau Mountains.
Figure 379 Dracocephalum nuratavicum (photography by N. Yu. Beshko)
1472. Dracocephalum scrobiculatum Regel
Semishrub. Stony slopes, moraines. Alpine zone. North Turkestan.
Genus 451. Kudrjaschevia Pojark.
1473. Kudrjaschevia jacubi (Lipsky) Pojark.
Annual. Fine-earth, gravelly and stony slopes, screes. Montane zone. Nuratau, Malguzar, North
Turkestan.
Genus 452. Lallemantia Fisch. & C. A. Mey.
1474. Lallemantia royleana (Benth.) Benth. Figure 380
Annual. Sandy and clay deserts, fine-earth, gravelly and stony slopes, fields, fallow lands. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal,
essential oil.
389
390
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 380 Lallemantia royleana (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 453. Hypogomphia Bunge
1475. Hypogomphia turkestana Bunge [Hypogomphia turkestanica Regel] Figure 381
Annual. Clay deserts, fine-earth, gravelly and stony slopes, river valleys, fields, fallow lands.
Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul.
Figure 381 Hypogomphia turkestana (photography by Natalya Beshko)
Genus 454. Prunella L.
1476. Prunella vulgaris L.
Perennial. Swamps, meadows, banks of rivers, lakes and canals, pebbles, stony and gravelly slopes,
wet places. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
391
392
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Malguzar, North Turkestan, Mirzachul. Medicinal, essential oil, fodder, ornamental, meliferous.
Genus 455. Phlomoides Moench
1477. Phlomoides ambigua (Popov ex Pazij & Vved.) Adylov, Kamelin & Makhm. Figure 382
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Meliferous, ornamental.
Figure 382 Phlomoides ambigua (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1478. Phlomoides anisochila (Pazij & Vved.) Salmaki Figure 383
Perennial. Stony slopes. Montane zone. Nuratau. Endemic to the Nuratau ridge. UzbRDB 1.
Figure 383 Phlomoides anisochila (photography by N. Yu. Beshko)
393
394
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1479. Phlomoides canescens (Regel) Adylov, Kamelin & Makhm. Figure 384
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Malguzar, North
Turkestan. Meliferous, ornamental.
Figure 384 Phlomoides canescens (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1480. Phlomoides eriocalyx (Regel) Adylov, Kamelin & Makhm. Figure 385
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar. Meliferous, ornamental.
Figure 385 Phlomoides eriocalyx (photography by Natalya Beshko)
1481. Phlomoides kaufmanniana (Regel) Adylov, Kamelin & Makhm. Figure 386
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Aktau, Malguzar, North
Turkestan. Meliferous, ornamental.
Figure 386 Phlomoides kaufmanniana (photography by Natalya Beshko)
395
396
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1482. Phlomoides labiosa (Bunge) Adylov, Kamelin & Makhm.
Perennial. Sandy and clay deserts. Plain. Mirzachul, Kyzylkum. Meliferous, ornamental.
1483. Phlomoides napuligera (Franch.) Adylov, Kamelin & Makhm. Figure 387
Perennial. Piedmont plains, gravelly, fine-earth and stony slopes. Plain, foothills. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan. Meliferous, ornamental.
Figure 387 Phlomoides napuligera (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1484. Phlomoides oreophila (Kar. & Kir.) Adylov, Kamelin & Makhm. [Phlomoides dszumrutensis
(Afan.) Adylov, Kamelin & Makhm.]
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Meliferous,
ornamental.
1485. Phlomoides sogdiana (Pazij & Vved.) Salmaki Figure 388
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Meliferous, ornamental.
Figure 388 Phlomoides sogdiana (photography by Natalya Beshko)
1486. Phlomoides speciosa (Rupr.) Adylov, Kamelin & Makhm. Figure 389
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Malguzar,
North Turkestan. Meliferous, ornamental.
Figure 389 Phlomoides speciosa (photography by Natalya Beshko)
1487. Phlomoides uniflora (Regel) Adylov, Kamelin & Makhm. Figure 390
Perennial. Sandy, clayey and gravelly soils. Plain, foothills. Nuratau, Nuratau Relic Mountains,
Mirzachul, Kyzylkum. Meliferous, ornamental.
397
398
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 390 Phlomoides uniflora (photography by Natalya Beshko)
Genus 456. Phlomis L.
1488. Phlomis linearifolia Zakirov Figure 391
Perennial. Fine-earth, gravelly and stony slopes, rocks. Foothills, montane zone. Nuratau, Malguzar,
North Turkestan. Essential oil, meliferous.
Figure 391 Phlomis linearifolia (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1489. Phlomis nubilans Zakirov Figure 392
Perennial. Fine-earth, gravelly and stony slopes. Montane zone, Nuratau. Essential oil, meliferous.
Threatened species, UzbRDB 2. Endemic to the Nuratau Mountains.
Figure 392 Phlomis nubilans (photography by N. Yu. Beshko)
399
400
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1490. Phlomis olgae Regel
Perennial. Fine-earth and stony slopes. Montane zone. Malguzar, North Turkestan. Essential oil,
meliferous.
1491. Phlomis salicifolia Regel
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Essential oil, meliferous.
1492. Phlomis thapsoides Bunge Figure 393
Perennial. Piedmont plains, fine-earth, gravelly and stony slopes. Foothills. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan. Essential oil, meliferous.
Figure 393 Phlomis thapsoides (photography by Natalya Beshko)
Genus 457. Stachyopsis Popov & Vved.
1493. Stachyopsis oblongata (Schrenk) Popov & Vved. Figure 394
Perennial. Stony slopes. Montane and alpine zones. North Turkestan.
Figure 394 Stachyopsis oblongata (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 458. Lamium L.
1494. Lamium album L. Figure 395
Perennial. Shady wet places, banks of rivers and canals, gardens, stony slopes, rocks. Montane and
alpine zones. Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, ornamental, meliferous.
Figure 395 Lamium album (photography by Natalya Beshko)
1495. Lamium amplexicaule L. Figure 396
Annual or biennial. Stony slopes, rocks, screes, fallow lands, fields, gardens, roadsides, banks
of rivers and canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, meliferous.
Figure 396 Lamium amplexicaule (photography by Natalya Beshko)
401
402
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 459. Leonurus L.
1496. Leonurus glaucescens Bunge
Perennial. Fine-earth, gravelly and stony slopes, rocks, ravines, banks of rivers. Montane zone. North
Turkestan.
1497. Leonurus turkestanicus V. I. Krecz. & Kuprian. Figure 397
Perennial. Fine-earth, gravelly and stony slopes, ravines, river valleys, shady wet places. Montane
zone. Nuratau, Malguzar, North Turkestan. Medicinal, meliferous.
Figure 397 Leonurus turkestanicus (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 460. Lagochilus Bunge ex Benth.
1498. Lagochilus inebrians Bunge Figure 398
Semishrub. Clay deserts, piedmont plains, stony slopes, pebbles. Plain, foothills. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar. Medicinal. Threatened species, UzbRDB 2.
Figure 398 Lagochilus inebrians (photography by Natalya Beshko)
403
404
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1499. Lagochilus olgae Kamelin Figure 399
Semishrub. Stony and gravelly slopes. Montane zone. Nuratau. UzbRDB 2. Endemic to the Nuratau
ridge.
Figure 399 Lagochilus olgae (photography by N. Yu. Beshko)
1500. Lagochilus proskorjakovii Ikramov Figure 400
Semishrub. Stony slopes, rocks, screes. Montane zone. Nuratau. Medicinal. Threatened species,
UzbRDB 1. Endemic to the Nuratau ridge.
Figure 400 Lagochilus proskorjakovii (photography by N. Yu. Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1501. Lagochilus seravschanicus Knorring
Semishrub. Stony slopes, screes. Montane zone. Malguzar, North Turkestan. Medicinal.
Genus 461. Stachys L.
1502. Stachys hissarica Regel Figure 401
Perennial. Fine-earth, gravelly and stony slopes, screes, river valleys. Montane and alpine zones.
Malguzar, North Turkestan. Medicinal, essential oil, meliferous.
Figure 401 Stachys hissarica (photography by Natalya Beshko)
1503. Stachys setifera C. A. Mey. Figure 402
Perennial. Wet places, pebbles, river valleys, fine-earth and stony slopes. Foothills, montane zone.
Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, meliferous.
Figure 402 Stachys setifera (photography by Natalya Beshko)
405
406
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 462. Chamaesphacos Schrenk
1504. Chamaesphacos ilicifolius Schrenk
Annual. Sandy deserts. Plain. Kyzylkum.
Figure 403
Figure 403 Chamaesphacos ilicifolius (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 463. Salvia L.
1505. Salvia aequidens Botsch.
Perennial. Stony slopes, rocks. Montane zone. Aktau, North Turkestan.
1506. Salvia deserta Schangin Figure 404
Perennial. Banks of rivers and canals, fine-earth slopes, meadows. Montane zone. Nuratau, Malguzar,
North Turkestan. Medicinal, essential oil, ornamental, meliferous.
Figure 404 Salvia deserta (photography by Natalya Beshko)
1507. Salvia glabricaulis Pobed.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. North Turkestan. Essential oil,
ornamental, meliferous.
1508. Salvia macrosiphon Boiss.
Perennial. River valleys, dry riverbeds, fine-earth, gravelly and stony slopes. Montane zone. Nuratau,
Malguzar, North Turkestan. Essential oil, ornamental, meliferous.
1509. Salvia sarawschanica Regel & Schmalh.
Perennial. Dry riverbeds, rocks, fine-earth, gravelly and stony slopes. Montane zone. Nuratau,
Malguzar, North Turkestan. Medicinal, essential oil, ornamental, meliferous.
1510. Salvia sclarea L. Figure 405
Perennial. River valleys, banks of canals, dry riverbeds, ravines, fine-earth, gravelly and stony slopes,
407
408
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
wastelands, fallow lands, gardens. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Medicinal, essential oil, food, ornamental, meliferous.
Figure 405 Salvia sclarea (photography by Natalya Beshko)
1511. Salvia spinosa L. Figure 406
Perennial. Clayey and gravelly soils, pebbles, dry riverbeds, stony slopes. Plain, foothills,
montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal,
essential oil, ornamental, meliferous.
Figure 406 Salvia spinosa (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1512. Salvia submutica Botsch. & Vved. Figure 407
Perennial. Stony slopes, rocks. Montane zone. Nuratau. Essential oil, ornamental, meliferous.
Threatened species, UzbRDB 2. Endemic to the Nuratau ridge.
Figure 407 Salvia submutica (photography by N. Yu. Beshko)
1513. Salvia virgata Jacq.
Perennial. River valleys, ravines, gardens, meadows, stream banks, fine-earth slopes. Montane zone.
Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, ornamental, meliferous.
Genus 464. Perovskia Kar.
1514. Perovskia angustifolia Kudrjasch.
Semishrub. Dry riverbeds, pebbles, stony slopes. Foothills, montane zone. Nuratau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Medicinal, essential oil, dye, meliferous.
1515. Perovskia botschantzevii Kovalevsk. & Koczk.
Semishrub. Stony slopes, pebbles, river valleys. Foothills, montane zone. Nuratau. Medicinal,
essential oil, dye, meliferous.
1516. Perovskia kudrjaschevii Gorschk. & Pjataeva
Semishrub. Pebbles, banks of rivers, stony slopes. Foothills, montane zone. Nuratau, North
Turkestan. Medicinal, essential oil, dye, meliferous.
1517. Perovskia scrophulariifolia Bunge
Semishrub. Banks of rivers, rocks, screes, stony slopes, pebbles, dry riverbeds. Foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, essential oil,
dye, meliferous.
409
410
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 465. Ziziphora L.
1518. Ziziphora clinopodioides Lam. Figure 408
Semishrub. Rocks, stony slopes, screes, banks of rivers, pebbles. Foothills, montane and alpine
zones. Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, essential oil, food (spice-aromatic),
meliferous.
Figure 408 Ziziphora clinopodioides (photography by Natalya Beshko)
1519. Ziziphora pamiroalaica Juz.
Semishrub. Fine-earth and stony slopes, rocks, screes, pebbles. Plain, foothills, montane and
alpine zones. Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, food (spice-aromatic),
meliferous.
1520. Ziziphora suffruticosa Pazij & Vved.
Semishrub. Stony slopes, pebbles. Montane zone. Malguzar, North Turkestan. Medicinal, essential
oil, food (spice-aromatic), meliferous.
1521. Ziziphora tenuior L.
Annual. Sandy and clay deserts, piedmont plains, river valleys, fine-earth, gravelly and stony
slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Medicinal, essential oil, food (spice-aromatic), meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 466. Clinopodium L.
1522. Clinopodium alpinum (L.) Kuntze [Acinos rotundiflorus Friv. ex Walp.]
Annual. Stony slopes, pebbles. Montane zone. Malguzar, North Turkestan. Medicinal.
1523. Clinopodium debile (Bunge) Kuntze [Antonia debilis (Bunge) Vved.]
Annual. Stony slopes, shady wet places, dry riverbeds. Montane and alpine zones. Malguzar, North Turkestan.
Genus 467. Hyssopus L.
1524. Hyssopus seravschanicus (Dubj.) Pazij Figure 409
Semishrub. Stony slopes, rocks, pebbles, screes. Montane and alpine zones. Malguzar, North
Turkestan. Medicinal, essential oil, food (spice-aromatic), meliferous.
Figure 409 Hyssopus seravschanicus (photography by Natalya Beshko)
Genus 468. Origanum L.
1525. Origanum vulgare subsp. gracile (K. Koch) Ietsw. [Origanum tyttanthum Gontsch.]
Perennial. Fine-earth and stony slopes, river valleys, pebbles. Montane zone. Nuratau, Aktau,
Malguzar, North Turkestan. Medicinal, essential oil, food, meliferous.
Genus 469. Thymus L.
1526. Thymus seravschanicus Klokov Figure 410
Semishrub. Stony and gravelly slopes, screes, pebbles, rocks, alpine meadows. Montane and
411
412
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
alpine zones. Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, food (spice-aromatic),
meliferous.
Figure 410 Thymus seravschanicus (photography by Natalya Beshko)
1527. Thymus subnervosus Vved., Nabiev & Tulyag.
Semishrub. Rocks, stony slopes. Montane zone. Nuratau. Essential oil, meliferous.
Genus 470. Lycopus L.
1528. Lycopus europaeus L.
Perennial. Wet places, swamps, banks of rivers and canals. Plain, foothills, montane zone. Nuratau,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, essential oil, tanniferous, dye, meliferous.
Genus 471. Mentha L.
1529. Mentha longifolia var. asiatica (Boriss.) Rech. f.
Perennial. Banks of rivers and canals, wet places. Plain, foothills, montane zone. Nuratau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, essential oil, food, fodder,
MAGNOLIOPSIDA
[EUDICOTS]
ornamental, meliferous.
1530. Mentha pamiroalaica Boriss.
Perennial. Wet places, banks of rivers. Montane zone. Nuratau, Malguzar, North Turkestan.
Medicinal, essential oil, food, fodder, ornamental, meliferous.
Family 100. Mazaceae
Genus 472. Dodartia L.
1531. Dodartia orientalis L. Figure 411
Perennial. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, river
valleys, wastelands, fallow lands, fields, roadsides. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, meliferous,
ornamental, insecticide.
Figure 411 Dodartia orientalis (photography by Natalya Beshko)
413
414
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Family 101. Orobanchaceae
Genus 473. Orobanche L.
1532. *Orobanche aegyptiaca Pers. [Phelipanche aegyptiaca (Pers.) Pom.]
Perennial. Fields, gardens, fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar,
North Turkestan, Mirzachul. Poisonous, weed. Parasite. Host plants are different vegetables, field
crops, ornamental plants, and rarely wild growing species.
1533. Orobanche amoena C. A. Mey. ex Ledeb.
Perennial. Piedmont plains, fine-earth and stony slopes, fallow lands, rocks, screes. Plain, foothills,
montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Weed. Parasite. Host
plants are representatives of genera Artemisia and Alhagi.
1534. Orobanche cernua Loefl.
Perennial. Sandy deserts, piedmont plains, banks of canals, fields, roadsides, riparian forests, screes, fineearth and stony slopes. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Weed. Parasite. Host plants are sunflowers, Karelinia caspica,
representatives of genera Artemisia, Lactuca, Scariola, Cousinia, Carthamus, and species of Solanaceae.
1535. Orobanche coelestis (Reut.) Boiss. & Reut. ex Beck [Phelipanche coelestis (Reut.) Soják]
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Weed. Parasite. Host plants are representatives of genera Artemisia, Cousinia, Scorsonera
and Centaurea.
1536. Orobanche elatior Sutton
Perennial. Stony slopes, screes, piedmont plains. Plain, foothills, montane zone. Malguzar,
North Turkestan. Weed. Parasite. Host plants are representatives of family Asteraceae (Echinops,
Centaurea, Artemisia, etc.).
1537. Orobanche gigantea (Beck) Gontsch.
Perennial. Rocks, screes, fine-earth and stony slopes. Montane zone. Nuratau. Weed. Parasite. Host
plants are representatives of family Apiaceae (Ferula, Schrenkia, etc.).
1538. Orobanche hansii A. Kern
Perennial. Piedmont plains, fallow lands, pebbles, stony slopes, sandy and clay deserts. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Weed. Parasite. Host plants are representatives of genera Artemisia, Inula,
Tussilago and Cousinia.
1539. Orobanche orientalis Beck [Phelipanche orientalis (Beck) Soják]
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Weed. Parasite. Host plants are wild almonds (Prunus bucharica, Prunus spinosissima).
1540. Orobanche pallens F. W. Schultz [Phelipanche pallens (Bunge) Soják]
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Weed. Parasite. Host plants are species of Artemisia.
MAGNOLIOPSIDA
[EUDICOTS]
1541. Orobanche sulphurea Gontsch.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau. Weed. Parasite.
Host plant is Crambe kotschyana.
Genus 474. Cistanche Hoffmanns. & Link.
1542. Cistanche mongolica Beck
Perennial. Saline lands, banks of lakes and canals. Plain. Mirzachul, Kyzylkum. Weed. Parasite. Host
plants are species of Tamarix.
1543. Cistanche salsa (C. A. Mey.) Beck Figure 412
Perennial. Saline lands, sandy deserts, fallow lands, banks of lakes and canals. Plain. Mirzachul,
Kyzylkum. Medicinal, fodder, weed. Parasite. Host plants are species of Haloxylon, Salsola,
Anabasis, Kalidium, Atriplex, Calligonum.
Figure 412 Cistanche salsa (photography by Natalya Beshko)
415
416
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 475. Leptorhabdos Schrenk ex Fisch. et C. A. Mey.
1544. Leptorhabdos parviflora (Benth.) Benth.
Annual. Banks of rivers, lakes and canals, sandy soils, fine-earth, gravelly and stony slopes, fallow
lands, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan.
Genus 476. Euphrasia L.
1545. Euphrasia pectinata Ten.
Annual. Meadows, fine-earth slopes. Montane and alpine zones. Malguzar, North Turkestan.
Medicinal, meliferous.
1546. Euphrasia regelii Wettst.
Annual. Meadows, fine-earth slopes. Montane and alpine zones. Malguzar, North Turkestan.
Medicinal, meliferous.
Genus 477. Parentucellia Viv.
1547. Parentucellia flaviflora (Boiss.) Nevski Figure 413
Annual. Fine-earth, gravelly and stony slopes, piedmont plains, clay desert. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Figure 413 Parentucellia flaviflora (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 478. Pedicularis L.
1548. Pedicularis dolichorrhiza Schrenk
Perennial. Fine-earth and stony slopes. Montane and alpine zones. North Turkestan. Medicinal.
1549. Pedicularis krylovii Bonati
Perennial. Stony slopes. Montane and alpine zones, North Turkestan.
1550. Pedicularis olgae Regel Figure 414
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Nuratau, Nuratau Relic
Mountains, Malguzar, North Turkestan.
Figure 414 Pedicularis olgae (photography by Natalya Beshko)
417
418
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1551. Pedicularis peduncularis Popov
Perennial. Swamp meadows, swamps. Alpine zone. North Turkestan.
1552. Pedicularis popovii Vved. Figure 415
Perennial. Fine-earth and stony slopes. Montane and alpine zones. North Turkestan.
Figure 415 Pedicularis popovii (photography by Natalya Beshko)
1553. Pedicularis rhinanthoides Schrenk
Perennial. Swamp meadows. Alpine zone. North Turkestan.
1554. Pedicularis sarawschanica Regel
Perennial. Wet stony slopes. Alpine zone. North Turkestan.
1555. Pedicularis waldheimii Bonati
Perennial. Screes. Alpine zone. North Turkestan.
Family 102. Campanulaceae
Genus 479. Campanula L.
1556. Campanula cashmeriana Royle
Perennial. Rocks. Montane and alpine zones. Malguzar, North Turkestan. Medicinal, meliferous,
MAGNOLIOPSIDA
[EUDICOTS]
ornamental.
1557. Campanula fedtschenkoana Trautv.
Perennial. Wet places near melting snow, stony slopes, rocks. Montane and alpine zones. North
Turkestan.
1558. Campanula glomerata L. Figure 416
Perennial. Fine-earth and stony slopes, river valleys, meadows. Montane and alpine zones. Malguzar,
North Turkestan. Medicinal, meliferous, ornamental.
Figure 416 Campanula glomerata (photography by Natalya Beshko)
1559. Campanula lehmanniana Bunge
Perennial. Stony slopes, rocks. Montane and alpine zones. Malguzar, North Turkestan.
Genus 480. Asyneuma Griseb. & Schrenk
1560. Asyneuma argutum (Regel) Bornm. Figure 417
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Medicinal.
419
420
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 417 Asyneuma argutum (photography by Natalya Beshko)
1561. Asyneuma trautvetteri (B. Fedtsch.) Bornm.
Perennial. Fine-earth and stony slopes. Montane and alpine zones. North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 481. Sergia Al. Fed.
1562. Sergia regelii (Trautv.) Al. Fed. Figure 418
Perennial. Rocks. Montane zone. Malguzar, North Turkestan.
Figure 418 Sergia regelii (photography by Natalya Beshko)
421
422
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 482. Cylindrocarpa Regel
1563. Cylindrocarpa sewerzowii (Regel) Regel
Perennial. Gravelly and stony slopes, rocks. Montane zone. Malguzar, North Turkestan.
Genus 483. Codonopsis Wall.
1564. Codonopsis clematidea (Schrenk) C. B. Clarke
Perennial. Banks of streams, stony slopes, wet places near springs, meadows. Montane zone.
Malguzar, North Turkestan. Medicinal.
Family 103. Asteraceae [Compositae]
Genus 484. *Cichorium L.
1565. *Cichorium intybus L.
Perennial. Fallow lands, wastelands, river valleys, pebbles, fine-earth and stony slopes, fields,
gardens, banks of canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, fodder, food, weed, meliferous.
Genus 485. Picris L.
1566. Picris nuristanica Bornm.
Annual or biennial. Pebbles, river valleys, fine-earth, gravelly and stony slopes. Plain, foothills,
montane zone. North Turkestan. Meliferous.
Genus 486. Garhadiolus Jaub. & Spach
1567. Garhadiolus hedypnois Jaub. & Spach [Garhadiolus angulosus Jaub. & Spach]
Annual. Sandy and clay deserts, fine-earth, gravelly and stony slopes, screes, pebbles, river valleys,
fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Weed.
1568. Garhadiolus papposus Boiss. & Buhse
Annual. Saline lands, sandy and clay deserts, fine-earth, gravelly and stony slopes, screes, pebbles,
river valleys, fallow lands, wastelands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Weed.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 487. Launaea Cass.
1569. Launaea korovinii (Popov) Pavlov [Rhabdotheca korovinii (Popov) Kirp.]
Annual. Sandy desert. Plain. Kyzylkum.
1570. Launaea procumbens (Roxb.) Ramayya & Rajagopal [Paramicrorhynchus procumbens
(Roxb.) Kipr.]
Perennial. Sandy and clay deserts, saline lands, banks of rivers, lakes and canals, fields, fallow lands,
roadsides. Plain. Mirzachul, Kyzylkum. Weed.
Genus 488. Chondrilla L.
1571. Chondrilla aspera (Schrad. ex Will.) Poir.
Perennial or Biennial. Clay deserts, fine-earth, gravelly and stony slopes, rocks, screes, pebbles,
banks of rivers and canals. Plain, foothills, montane and alpine zones. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder.
1572. Chondrilla lejosperma Kar. & Kir.
Perennial. Fine-earth, gravelly and stony slopes, screes, pebbles, banks of rivers, lakes and canals.
Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Fodder.
Genus 489. Heteroderis (Bunge) Boiss.
1573. Heteroderis pusilla (Boiss.) Boiss.
Annual. Sandy deserts, saline lands. Plain. Kyzylkum.
Genus 490. Lactuca L.
1574. Lactuca crambifolia (Bunge) Boiss. [Steptorhamphus crambifolius Bunge]
Perennial. Gravelly and stony slopes, screes, rocks. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan.
1575. Lactuca dissecta D. Don.
Annual. Fine-earth, gravelly and stony slopes, screes, rocks, shady wet places, banks of rivers,
roadsides, fallow lands, wastelands. Plain, foothills, montane and alpine zones. North Turkestan.
1576. Lactuca glauciifolia Boiss.
Annual. Fine-earth, gravelly and stony slopes, rocks, pebbles, fallow lands, wastelands. Plain,
foothills, montane and alpine zones. North Turkestan, Mirzachul.
1577. Lactuca orientalis (Boiss.) Boiss. [Scariola orientalis (Boiss.) Sojăk]
Semishrub. Fine-earth, gravelly and stony slopes, screes, rocks, pebbles, dry riverbeds. Plain,
423
424
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan.
1578. Lactuca serriola Torner [Lactuca altaica Fisch. & C. A. Mey.]
Annual or biennial. Fine-earth, gravelly and stony slopes, pebbles, roadsides, banks of rivers, lakes
and canals, fallow lands, wastelands, fields, gardens, settlements. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, food, fodder, oil, weed, rubberbearing.
1579. Lactuca soongorica Regel [Cephalorrhynchus soongoricus (Regel) Kovalevsk.]
Perennial. Stony slopes, screes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
1580. Lactuca spinidens Nevski
Annual or biennial. Stony slopes, fields, fallow lands. Nuratau.
1581. Lactuca tatarica (L.) C. A. Mey.
Perennial. Saline lands, pebbles, banks of rivers, lakes and canals, fields, fallow lands, roadsides.
Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum.
1582. Lactuca undulata Ledeb.
Annual. Sandy and clay deserts, banks of rivers, lakes and canals, saline lands, pebbles, fine-earth,
gravelly and stony slopes, fields, gardens, fallow lands, roadsides. Plain, foothills. Nuratau Relic
Mountains, Mirzachul, Kyzylkum.
Genus 491. Kovalevskiella Kamelin
1583. Kovalevskiella zeravschanica (Popov & Kovalevsk) Kamelin
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. North Turkestan.
Genus 492. Crepidifolium Sennikov
1584. Crepidifolium serawschanicum (B. Fedtsch.) Sennikov [Youngia serawschanica (B.
Fedtsch.) Babc. & Stebbins]
Perennial. Gravelly and stony slopes, screes, rocks, alpine meadows. Montane and alpine zones.
North Turkestan.
Genus 493. *Sonchus L.
1585. *Sonchus arvensis L.
Perennial. Banks of rivers, lakes and canals, meadows, wastelands, fallow lands, fields, gardens,
settlements. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Medicinal, food, fodder, weed, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
1586. *Sonchus asper (L.) Hill
Annual. Fallow lands, fields, roadsides, wastelands, settlements. Nuratau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Food, fodder, weed, meliferous.
1587. *Sonchus oleraceus (L.) L.
Annual. Fallow lands, fields, roadsides, banks of rivers, lakes and canals, wastelands, settlements.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Medicinal, food, fodder, weed, meliferous.
1588. *Sonchus palustris L.
Perennial. Banks of rivers and canals, swamps, wet places, gardens. Plain, foothills, montane zone.
Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Weed,
meliferous.
Genus 494. Taraxacum F. H. Wigg.
1589. Taraxacum bessarabicum (Hornem.) Hand.-Mazz.
Perennial. Meadows, swamps, pebbles, banks of rivers, lakes and canals, fallow lands, wet places. Plain,
foothills, montane zone. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Meliferous.
1590. Taraxacum bicorne Dahlst
Perennial. Meadows, saline lands, depressions, banks of rivers, lakes and canals, fallow lands, fields,
roadsides. Plain, foothills. Nuratau Relic Mountains, Mirzachul, Kyzylkum. Meliferous, weed.
1591. Taraxacum brevirostre Hand.-Mazz.
Perennial. Fine-earth, gravelly and stony slopes, alpine meadows. Alpine zone. North Turkestan.
1592. Taraxacum contristans Kovalevsk.
Perennial. Wet places, saline lands, roadsides, banks of canals, fields, fallow lands. Plain, foothills.
Malguzar, Mirzachul. Meliferous, weed.
1593. Taraxacum ecornutum Kovalevsk.
Perennial. Banks of rivers and canals, roadsides, fields, fallow lands, ravines. Plain, foothills,
montane zone. Nuratau, Nuratau Relic Mountains, North Turkestan, Mirzachul. Meliferous,
weed.
1594. Taraxacum elongatum Kovalevsk.
Perennial. Banks of rivers, meadows, swamps, fine-earth, gravelly and stony slopes. Montane zone.
Malguzar, North Turkestan. Meliferous.
1595. Taraxacum erostre Zakirov
Perennial. Rocks, gravelly and stony slopes, screes, moraines. Montane and alpine zones. Malguzar,
North Turkestan.
1596. Taraxacum glaucivirens Schischk.
Perennial. Fine-earth, gravelly and stony slopes, rocks, moraines. Montane and alpine zones. North
Turkestan.
425
426
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1597. Taraxacum maracandicum Kovalevsk.
Perennial. Wet places, banks of rivers and canals, meadows, fine-earth slopes, roadsides, settlements.
Plain, foothills, Montane zone. Nuratau, Malguzar, North Turkestan.
1598. Taraxacum marginatum (Dahlst.) Raunk. [Taraxacum reflexum (Brenner) Hjelt]
Perennial. Stony slopes, screes, wet places, swamps, meadows. Montane zone. North Turkestan.
1599. Taraxacum minutilobium Popov ex Kovalevsk.
Perennial. Gravelly and stony slopes, moraines, rocks, screes, banks of rivers. Alpine zone.
Malguzar, North Turkestan.
1600. Taraxacum modestum Schischk.
Perennial. Fine-earth, gravelly and stony slopes, wet places near melting snow. Montane and alpine
zones. North Turkestan.
1601. Taraxacum monochlamydeum Hand.-Mazz.
Perennial. Fine-earth slopes, piedmont plains, gardens, fields, fallow lands, roadsides, wastelands,
banks of rivers and canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Meliferous.
1602. Taraxacum nevskii Juz.
Perennial. Gravelly and stony slopes, screes. Montane zone. Malguzar, North Turkestan.
1603. Taraxacum nuratavicum Schischk.
Perennial. Wet places, swamps, banks of rivers. Montane zone. Nuratau. Meliferous.
1604. *Taraxacum officinale (L.) Weber ex F.H. Wigg.
Perennial. Banks of rivers, lakes and canals, fallow lands, wastelands, gardens, settlements. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Medicinal, food, fodder, meliferous.
1605. Taraxacum popovii Kovalevsk. ex Vainberg
Perennial. Gravelly and stony slopes, moraines, alpine meadows, screes, wet places near melting
snow. Montane and alpine zones. Malguzar, North Turkestan.
1606. Taraxacum repandum Pavlov
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Malguzar, North
Turkestan.
1607. Taraxacum sonchoides (D. Don) Sch. Bip. [Taraxacum montanum (C. A. Mey.) DC.]
Perennial. Fine-earth, gravelly and stony slopes, dry riverbeds, pebbles. Montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Meliferous.
1608. Taraxacum strobilocephalum Kovalevsk.
Perennial. Wet places, banks of rivers and canals. Malguzar.
Genus 495. Heteracia Fisch. & C. A. Mey.
1609. Heteracia szovitzii Fisch. & C. A. Mey. [Heteracia epapposa (Regel & Schmalh.) Popov]
Annual. River valleys, sandy and clay deserts, saline lands, piedmont plains, fine-earth, gravelly and
stony slopes, rocks, fields, fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau
MAGNOLIOPSIDA
[EUDICOTS]
Relic Mountains, Malguzar, Mirzachul, Kyzylkum.
Genus 496. Crepis L.
1610. Crepis kotschyana (Boiss.) Boiss. [Barkhausia kotschyana Boiss.]
Annual. Gravelly and stony slopes, rocks. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan.
1611. Crepis multicaulis Ledeb.
Perennial. Gravelly and stony slopes, screes, pebbles, moraines, river valleys, wet places. Montane
and alpine zones. North Turkestan.
1612. Crepis pulchra L. [Phaecasium pulchrum (L.) Reich. f.] Figure 419
Annual. Fine-earth, gravelly and stony slopes, screes, rocks, banks of rivers and canals, shady wet
places, gardens, roadsides, fields, fallow lands. Plain, foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Figure 419 Crepis pulchra (photography by Natalya Beshko)
Genus 497. Pilosella Hill
1613. Pilosella echioides (Lumn.) F. W. Schultz & Sch. Bip. [Hieracium echioides Lumn.]
Perennial. Stony slopes. Montane and alpine zones. Malguzar, North Turkestan. Medicinal.
427
428
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1614. Pilosella procera (Fr.) F. W. Schultz & Sch. Bip. [Hieracium procerum Fr.]
Perennial. Fine-earth, gravelly and stony slopes, meadows, ravines. Montane zone. North Turkestan.
Genus 498. Hieracium L.
1615. Hieracium raddeanum subsp. regelianum (Zahn) Greuter [Hieracium regelianum Zahn]
Perennial. Juniper forests, meadows, river valleys. Montane zone. North Turkestan.
1616. Hieracium robustum Fr.
Perennial. Meadows, fine-earth, gravelly and stony slopes, river valleys. Montane zone. North
Turkestan. Weed.
1617. Hieracium virosum Pall. Figure 420
Perennial. Meadows, fine-earth, gravelly and stony slopes, river valleys. Montane zone. North
Turkestan. Weed.
Figure 420 Hieracium virosum (photography by Natalya Beshko)
Genus 499. Acanthocephalus Kar. & Kir.
1618. Acanthocephalus amplexifolius Kar. & Kir.
Annual. Saline lands, river valleys, dry riverbeds, ravines, fine-earth, gravelly and stony slopes.
Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Weed.
MAGNOLIOPSIDA
[EUDICOTS]
1619. Acanthocephalus benthamianus Regel
Annual. River valleys, banks of canals, gardens, roadsides, ravines, fine-earth, gravelly and stony
slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Weed.
Genus 500. Scorzonera L.
1620. Scorzonera acanthoclada Franch.
Semishrub. Gravelly and stony slopes. Montane and alpine zones. North Turkestan. fodder, rubberbearing.
1621. Scorzonera bracteosa C. Winkl. Figure 421
Perennial. Fine-earth and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
Rubber-bearing, meliferous.
Figure 421 Scorzonera bracteosa (photography by Natalya Beshko)
1622. Scorzonera circumflexa Krasch. & Lipsch.
Perennial. Fine-earth, gravelly and stony slopes, pebbles, clay deserts. Plain, foothills, montane and
alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Rubber-bearing, meliferous.
1623. Scorzonera hissarica C. Winkl.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Malguzar, North
Turkestan.
1624. Scorzonera inconspicua Lipsch.
Perennial. Fine-earth, gravelly and stony slopes, pebbles, banks of rivers. Plain, foothills, montane
zone. Malguzar, North Turkestan. Rubber-bearing, meliferous.
1625. Scorzonera litwinowii Krasch. & Lipsch.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau. Rubber-bearing,
429
430
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
meliferous.
1626. Scorzonera ovata Trautv. Figure 422
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Nuratau Relic
Mountains, Malguzar, North Turkestan. Rubber-bearing, meliferous.
Figure 422 Scorzonera ovata (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1627. Scorzonera songarica (Kar. & Kir.) Lipsch. & Vassilcz.
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Montane zone. North Turkestan.
1628. Scorzonera tragopogonoides Regel & Schmalh.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane and alpine zones, Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Rubber-bearing, meliferous.
Genus 501. Takhtajaniantha Nazarova
1629. Takhtajaniantha pusilla (Pall.) Nazarova [Scorzonera pusilla Pall.] Figure 423
Perennial. Saline lands, sandy and clay deserts, dry riverbeds, stony slopes. Plain, foothills. Nuratau
Relic Mountains, Mirzachul, Kyzylkum. Rubber-bearing, meliferous.
Figure 423 Takhtajaniantha pusilla (photography by Natalya Beshko)
Genus 502. Epilasia (Bunge) Benth. & Hook. f.
1630. Epilasia acrolasia (Bunge) Lipsch.
Annual. Sandy and clay deserts, saline lands, fine-earth and stony slopes, river valleys, fallow lands.
Plain, foothills. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum.
431
432
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1631. Epilasia hemilasia (Bunge) C. B. Clarke Figure 424
Annual. Sandy and clay deserts, saline lands, fine-earth, gravelly and stony slopes, roadsides, fields,
fallow lands. Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum.
Figure 424 Epilasia hemilasia (photography by Natalya Beshko)
1632. Epilasia mirabilis Lipsch.
Annual. Sandy and clay deserts, fields, fallow lands. Plain. Mirzachul.
Genus 503. Tragopogon L.
1633. Tragopogon capitatus S. A. Nikitin
Biennial. Fine-earth, gravelly and stony slopes, river valleys. Foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Rubber-bearing, meliferous.
1634. Tragopogon conduplicatus S.A. Nikitin
Perennial. Gravelly and stony slopes. Montane zone. Nuratau, North Turkestan. Rubber-bearing,
meliferous.
1635. Tragopogon kultiassovii Popov ex S. A. Nikitin
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. North Turkestan. Rubber-
MAGNOLIOPSIDA
[EUDICOTS]
bearing, meliferous.
1636. Tragopogon malikus S. A. Nikitin
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Rubber-bearing, meliferous.
1637. Tragopogon porrifolius subsp. longirostris (Sch. Bip.) Greuter [Tragopogon
kraschennikovii S. A. Nikitin] Figure 425
Biennial. Fine-earth, gravelly and stony slopes, river valleys, fallow lands, fields. Foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Rubber-bearing,
meliferous.
Figure 425 Tragopogon porrifolius subsp. longirostris (photography by Natalya Beshko)
1638. Tragopogon pseudomajor S. A. Nikitin
Biennial. Piedmont plains, fine-earth and stony slopes, river valleys. Plain, foothills, montane zone.
Nuratau, Malguzar, North Turkestan. Rubber-bearing, meliferous.
1639. Tragopogon turkestanicus S. A. Nikitin ex Pavlov
Biennial. Gravelly and stony slopes. Foothills, montane and alpine zones. Nuratau, Malguzar, North
Turkestan. Rubber-bearing, meliferous.
1640. Tragopogon vvedenskyi Popov ex Pavlov
Biennial. Fine-earth, gravelly and stony slopes, ravines. Foothills, montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Rubber-bearing, meliferous.
Genus 504. Koelpinia Pall.
1641. Koelpinia linearis Pall.
Annual. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes. Plain,
foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
433
434
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Turkestan, Mirzachul, Kyzylkum. Fodder, weed, meliferous.
1642. Koelpinia macrantha C. Winkl.
Annual. Sandy deserts, fine-earth, gravelly and stony slopes, river valleys. Plain, foothills. Nuratau,
Kyzylkum. Fodder, weed, meliferous.
1643. Koelpinia tenuissima Pavlov & Lipsch.
Annual. Sandy and clay deserts, saline lands, fine-earth, gravelly and stony slopes. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Fodder, weed, meliferous.
1644. Koelpinia turanica Vassilcz.
Annual. Sandy and clay deserts, saline lands, fine-earth, gravelly and stony slopes. Plain, foothills,
montane zone. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder, weed, meliferous.
Genus 505. Echinops L.
1645. Echinops knorringianus Iljin
Annual. Fine-earth, gravelly and stony slopes. Foothills. North Turkestan.
1646. Echinops leucographus Bunge
Perennial. Fine-earth, gravelly and stony slopes, piedmont plains. Plain, foothills. Nuratau, Nuratau
Relic Mountains.
1647. Echinops maracandicus Bunge
Perennial. Fine-earth, gravelly and stony slopes, pebbles, screes. Foothills, montane and alpine
zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
1648. Echinops nanus Bunge
Annual or biennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. North Turkestan.
1649. Echinops nuratavicus A. D. Li Figure 426
Perennial. Gravelly and stony slopes, rocks. Montane zone. Nuratau.
Figure 426 Echinops nuratavicus (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 506. Chardinia Desf.
1650. Chardinia orientalis (L.) Kuntze
Annual. Clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, rocks. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Genus 507. Xeranthemum L.
1651. Xeranthemum longepapposum Fisch. & C. A. Mey.
Annual. Piedmont plains, fallow lands, pebbles, fine-earth, gravelly and stony slopes, rocks. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul.
Genus 508. Carduus L.
1652. Carduus pycnocephalus subsp. albidus (M. Bieb.) Kazmi [Carduus albidus M. Bieb.]
Annual. River valleys, dry riverbeds, banks of canals, gardens, roadsides, fields, fallow lands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Weed, meliferous.
1653. Carduus nutans L.
Biennial or perennial. Fine-earth, gravelly and stony slopes, rocks, banks of rivers and canals, wet places,
fallow lands, wastelands, roadsides. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, food, weed, meliferous.
Genus 509. Picnomon Adans.
1654. Picnomon acarna (L.) Cass.
Annual or biennial. Gravelly and stony slopes, fallow lands, wastelands, roadsides, banks of
canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Weed.
Genus 510. Onopordum L.
1655. Onopordum acantium L.
Biennial. Gravelly and stony slopes, ravines, wastelands, fallow lands, river valleys, banks of
canals, fields, roadsides. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, oil, weed, fodder, meliferous.
435
436
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1656. Onopordum leptolepis DC.
Biennial. Fine-earth, gravelly and stony slopes, rocks, screes, river valleys, fallow lands, fields,
roadsides, banks of canals. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Weed.
Genus 511. Cirsium Mill.
1657. Cirsium alatum (S. C. Gmel.) Bobrov
Perennial. Meadows, saline lands, wet places, banks of rivers and canals, piedmont plains, ravines,
fields. Plain, foothills, montane zone. Malguzar. Weed.
1658. Cirsium arvense (L.) Scop. [Cirsium ochrolepidium Juz.]
Perennial. Sandy deserts, fallow lands, fields, banks of rivers and canals, wet places, meadows,
riparian forests, fine-earth slopes, gardens. Plain, foothills, montane and alpine zones. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal,
poisonous, oil, weed, meliferous.
1659. Cirsium brevipapposum Czerniak.
Perennial. Swamps, wet places near springs. Montane and alpine zones. Malguzar, North Turkestan.
1660. Cirsium esculentum (Siev.) C. A. Mey.
Perennial. Wet places, banks of rivers, lakes and canals, meadows, fine-earth slopes, gardens. Plain,
foothills, montane and alpine zones. Malguzar, North Turkestan. Medicinal, weed, meliferous.
1661. Cirsium semenovii Regel
Biennial or perennial. Stony slopes, pebbles, screes, moraines. Montane and alpine zones. North
Turkestan.
1662. Cirsium straminispinum C. Jeffrey ex Cufod [Cirsium polyacanthum Hochst. ex Kar. & Kir.]
Biennial or perennial. Fine-earth slopes, ravines, wet places. Foothills, montane and alpine zones.
North Turkestan. Weed, meliferous.
1663. Cirsium turkestanicum (Regel) Petr.
Perennial. Gravelly and stony slopes, rocks, screes, river valleys, swamps, banks of canals. Plain,
foothills, montane and alpine zones. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Weed, meliferous.
1664. Cirsium vulgare (Savi) Ten.
Biennial. Wet places, banks of rivers and canals, fallow lands, fields, wastelands, gravelly and stony
slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Medicinal, food, weed, meliferous.
Genus 512. Arctium L.
1665. Arctium aureum (C. Winkl.) Kuntze [Cousinia aurea C. Winkl.]
Perennial. Gravelly and stony slopes, pebbles. Foothills, montane zone. Nuratau Relic Mountains,
MAGNOLIOPSIDA
[EUDICOTS]
Malguzar, North Turkestan. Fodder.
1666. Arctium chloranthum (Kult.) S. López, Romasch., Susanna & N.Garcia [Cousinia
chlorantha Kult.]
Perennial. Gravelly and stony slopes. Foothills, montane zone. Nuratau, Nuratau Relic Mountains,
Malguzar, North Turkestan.
1667. Arctium haesitabundum (Juz.) S. López, Romasch., Susanna & N.Garcia [Cousinia
haesitabunda Juz.]
Perennial. Gravelly and stony slopes. Montane zone, Nuratau, Malguzar. UzbRDB 1.
1668. Arctium karatavicum (Regel & Schmalh.) Kuntze [Cousinia karatavica Regel &
Schmalh.] Figure 427
Perennial. Gravelly and stony slopes, rocks. Foothills, montane zone. Nuratau, Nuratau Relic
Mountains, Malguzar.
Figure 427 Arctium karatavicum (photography by Natalya Beshko)
1669. Arctium korolkowii (Regel & Schmalh.) Kuntze [Cousinia korolkowii Regel &
Schmalh.]
Perennial. Gravelly and stony slopes. Foothills, montane zone. Nuratau.
1670. Arctium leiospermum Juz. & Ye.V. Serg.
Biennial. River valleys, banks of canals, roadsides, wet places, gardens. Plain, foothills, montane
zone. Nuratau, Malguzar, North Turkestan. Medicinal, food, weed, meliferous.
437
438
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1671. Arctium pallidivirens (Kult.) S. Lopez, Romanschenko, Susanna & N. Garcia [Anura
pallidivirens (Kult.) Tscherneva] Figure 428
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar. UzbRDB 1.
Endemic to the Nuratau Mountains.
Figure 428 Arctium pallidivirens (photography by Natalya Beshko)
Genus 513. Cousinia L.
1672. Cousinia alpina Bunge
Biennial. Fine-earth, gravelly and stony slopes, screes, rocks. Montane and alpine zones. North
Turkestan.
1673. Cousinia ambigens Juz.
Perennial. Gravelly and stony slopes. Foothills. Nuratau.
1674. Cousinia botschantzevii Juz. ex Tscherneva Figure 429
Biennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Aktau. Endemic to the Nuratau Mountains.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 429 Cousinia botschantzevii (photography by Natalya Beshko)
1675. Cousinia bungeana Regel & Schmalh.
Annual. Clay deserts. Plain. Mirzachul.
1676. Cousinia coronata Franch.
Biennial. Fine-earth, gravelly and stony slopes. Montane zone. Aktau, Malguzar, North Turkestan.
1677. Cousinia dissectifolia Kult.
Biennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau.
1678. Cousinia dshisakensis Kult. Figure 430
Biennial. Gravelly and stony slopes. Montane zone. Nuratau, Aktau, Malguzar, North Turkestan.
UzbRDB 2.
439
440
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 430 Cousinia dshisakensis (photography by Natalya Beshko)
1679. Cousinia dubia Popov
Biennial. Gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
1680. Cousinia eriotricha Juz. Figure 431
Perennial. Gravelly and stony slopes, screes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Figure 431 Cousinia eriotricha (photography by Natalya Beshko)
1681. Cousinia franchetii C. Winkl.
Perennial. Stony slopes. Montane zone. North Turkestan.
1682. Cousinia hamadae Juz.
Perennial. Gravelly and stony slopes. Foothills. Nuratau Relic Mountains.
1683. Cousinia horridula Juz.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan.
1684. Cousinia integrifolia Franch.
Biennial. Fine-earth and stony slopes. Montane zone. Malguzar, North Turkestan.
1685. Cousinia lappacea Schrenk
Perennial. Stony slopes, rocks. Montane zone. Malguzar, North Turkestan. Fodder, weed.
MAGNOLIOPSIDA
[EUDICOTS]
1686. Cousinia microcarpa Boiss. Figure 432
Perennial. Fine-earth, gravelly and stony slopes, river valleys, banks of canals, fallow lands, fields,
roadsides, wastelands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Meliferous, weed.
Figure 432 Cousinia microcarpa (photography by Natalya Beshko)
1687. Cousinia mollis Schrenk
Biennial. Fine-earth, gravelly and stony slopes, screes, sandy and clay deserts, fallow lands, river
valleys. Plain, foothills. Nuratau, Nuratau Relic Mountains, North Turkestan, Mirzachul, Kyzylkum.
1688. Cousinia olgae Regel & Schmalh.
Biennial. Fine-earth, gravelly and stony slopes, pebbles, piedmont plains. Plain, foothills. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
1689. Cousinia outichaschensis Franch.
Biennial. Gravelly and stony slopes, rocks. Montane and alpine zones. Malguzar, North Turkestan.
1690. Cousinia oxiana Tscherneva
Biennial. Sandy deserts. Plain. Kyzylkum.
1691. Cousinia polytimetica Tscherneva
Annual. Piedmont plain, river valleys. Plain, foothills. Nuratau, Nuratau Relic Mountains, Malguzar.
1692. Cousinia prolifera Jaub. & Spach.
Annual. Sandy and clay deserts, saline lands, piedmont plains, fields, fallow lands. Plain, foothills.
441
442
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Nuratau Relic Mountains, Mirzachul, Kyzylkum.
1693. Cousinia pseudodshisakensis Tscherneva & Vved.
Biennial. Gravelly and stony slopes, screes, rocks. Foothills, montane zone. Nuratau, Nuratau Relic
Mountains, Malguzar, North Turkestan.
1694. Cousinia pygmaea C. Winkl.
Biennial. Fine-earth and stony slopes, fallow lands. Montane zone. Malguzar, North Turkestan.
1695. Cousinia radians Bunge Figure 433
Biennial. Fine-earth, gravelly and stony slopes, piedmont plains, fallow lands. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Weed, meliferous.
Figure 433 Cousinia radians (photography by Natalya Beshko)
1696. Cousinia regelii C. Winkl.
Biennial. Fine-earth, gravelly and stony slopes. Foothills. Aktau.
1697. Cousinia resinosa Juz.
Perennial. Fine-earth, gravelly and stony slopes, piedmont plains, sandy and clay deserts, roadsides,
fallow lands, wastelands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Fodder, weed, meliferous.
1698. Cousinia schtschurowskiana Regel & Schmalh. Figure 434
Biennial. Gravelly and stony slopes. Montane zone. Nuratau, Aktau.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 434 Cousinia schtschurowskiana (photography by Natalya Beshko)
1699. Cousinia simulatrix C. Winkl.
Biennial. Gravelly and stony slopes, ravines. Montane zone. Nuratau.
1700. Cousinia sogdiana Bornm. Figure 435
Biennial. Sandy deserts. Plain. Kyzylkum.
Figure 435 Cousinia sogdiana (photography by Natalya Beshko)
443
444
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1701. Cousinia spiridonovii Juz.
Perennial. Clay deserts, river valleys, banks of canals, fallow lands. Plain. Nuratau Relic Mountains,
Mirzachul.
1702. Cousinia tenella Fisch. & C. A. Mey.
Annual. Sandy and clay deserts, saline lands, piedmont plains, river valleys, fields, roadsides, fallow
lands. Plain, foothills. Nuratau, Nuratau Relic Mountains, Mirzachul, Kyzylkum.
1703. Cousinia umbrosa Bunge
Perennial. Fine-earth slopes, river valleys, shady wet places. Foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Fodder, weed, meliferous.
1704. Cousinia verticillaris Bunge
Perennial. Gravelly and stony slopes. Alpine zone. Malguzar, North Turkestan.
1705. Cousinia xanthina Bornm.
Perennial. Fine-earth slopes. Alpine zone. North Turkestan.
Genus 514. Lipskyella Juz.
1706. Lipskyella annua (C. Winkl.) Juz.
Annual. Sandy deserts. Plain. Kyzylkum.
Genus 515. Saussurea DC.
1707. Saussurea elegans Ledeb.
Perennial. Fine-earth and stony slopes, meadows, river valleys. Foothills, montane zone. North
Turkestan.
1708. Saussurea salsa (Pall. ex Pall.) Spreng.
Perennial. Saline lands, banks or rivers, lakes and canals. Plain. Mirzachul.
Genus 516. Jurinea Cass.
1709. Jurinea abramowii Regel & Herder
Perennial. Stony slopes, screes. Montane and alpine zones. North Turkestan.
1710. Jurinea androssovii Iljin
Perennial. Gravelly and stony slopes, rocks. Alpine zone. North Turkestan.
1711. Jurinea ferganica (Iljin) Iljin
Perennial. Gravelly and stony slopes, screes, rocks. Foothills, montane zone. Malguzar, North
Turkestan.
1712. Jurinea helichrysifolia Popov ex Iljin Figure 436
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar, NorthTurkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 436 Jurinea helichrysifolia (photography by Natalya Beshko)
1713. Jurinea kokanica Iljin
Perennial. Fine-earth, gravelly and stony slopes, river valleys. Foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental, meliferous.
1714. Jurinea komarovii Iljin
Perennial. Gravelly and stony slopes, pebbles, rocks. Montane and alpine zones. North Turkestan.
1715. Jurinea lasiopoda Trautv.
Perennial. Stony slopes, rocks. Foothills, montane zone. Nuratau, Nuratau Relic Mountains.
445
446
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1716. Jurinea maxima C. Winkl. Figure 437
Perennial. Fine-earth and stony slopes. Montane and alpine zones. Malguzar, North Turkestan.
Ornamental, meliferous.
Figure 437 Jurinea maxima (photography by Natalya Beshko)
1717. Jurinea olgae Regel & Schmalh. Figure 438
Perennial. Rocks, Fine-earth, gravelly and stony slopes, screes. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Ornamental, meliferous.
Figure 438 Jurinea olgae (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1718. Jurinea suffruticosa Regel
Semishrub. Piedmont plains, gravelly and stony slopes, rocks. Plain, foothills, montane zone. Nuratau.
1719. Jurinea trautvetteriana Regel & Schmalh. Figure 439
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Malguzar,
North Turkestan. Ornamental, meliferous.
Figure 439 Jurinea trautvetteriana (photography by Natalya Beshko)
1720. Jurinea zakirovii Iljin Figure 440
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau. UzbRDB 1. Endemic to the
Nuratau Mountains.
Figure 440 Jurinea zakirovii (photography by N. Yu. Beshko)
447
448
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 517. Amberboa (Pers.) Less.
1721. Amberboa turanica Iljin
Annual. Sandy and clay deserts, piedmont plains, fallow lands. Plain, foothills. Nuratau, Nuratau
Relic Mountains, Malguzar, Mirzachul, Kyzylkum.
Genus 518. Centaurea L.
1722. *Centaurea benedicta (L.) L. [Cnicus benedictus L.]
Annual. Clay deserts, wastelands, fallow lands, fields, wet places, banks of rivers and canals.
Plain, foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul.
Medicinal, oil, weed.
1723. Centaurea bruguierana subsp. belangeriana (DC.) Bornm. [Centaurea belangeriana
(DC.) Stapf]
Annual. Sandy, clayey and skeleton soils, fields, fallow lands, wastelands, roadsides. Plain, foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum. Weed.
1724. *Centaurea iberica Trevir. ex Spreng.
Biennial. Wastelands, banks of canals, river valleys, roadsides, surroundings of settlements, fineearth, gravelly and stony slopes. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, weed.
1725. *Centaurea solstitialis L.
Biennial. Fallow lands, wastelands, fields, banks of canals, surroundings of settlements, fine-earth and
gravelly slopes. Plain, foothills. Nuratau, Malguzar, North Turkestan, Mirzachul. Medicinal, weed.
1726. Centaurea virgata subsp. squarrosa (Boiss.) Gugler [Centaurea squarrosa Willd.]
Perennial. Fine-earth, gravelly and stony slopes, piedmont plains, sandy and clay deserts, fields,
fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Medicinal, weed, meliferous.
Genus 519. Cyanus Mill.
1727. Cyanus depressus (M. Bieb.) Soják [Centaurea depressa M. Bieb.]
Annual. Fallow lands, wastelands, fields, banks of canals, fine-earth and gravelly slopes. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul. Medicinal, ornamental, meliferous, weed.
1728. *Cyanus segetum Hill [Centaurea cyanus L.]
Annual. Fallow lands, fields, wastelands, surroundings of settlements, banks of canals. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Medicinal, poisonous, dye, ornamental, meliferous, weed.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 520. Rhaponticum Ludw.
1729. Rhaponticum repens (L.) Hidalgo [Acroptilon repens (L.) DC.]
Perennial. Sandy and clay desets, saline lands, banks of rivers, lakes and canals, fine-earth, gravelly
and stony slopes, fields, fallow lands, roadsides, wastelands, gardens. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Medicinal, poisonous, meliferous, weed. Quarantine invasive species.
Genus 521. Hyalea (DC.) Jaub. & Spach
1730. Hyalea pulchella (Ledeb.) Koch
Annual. Sandy desert. Plain. Kyzylkum.
Figure 441
Figure 441 Hyalea pulchella (photography by Natalya Beshko)
Genus 522. *Stizolophus Cass.
1731. *Stizolophus balsamita (Lam.) K. Koch
Annual. Fields, fallow lands, roadsides, gravelly and stony slopes, banks of rivers, lakes and
canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
449
450
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Turkestan, Mirzachul. Medicinal, dye, weed.
Genus 523. Crupina Cass.
1732. Crupina vulgaris Pers. ex Cass. [Crupina oligantha Tscherneva]
Annual. Gravelly and stony slopes, rocks, screes, pebbles, banks of rivers, gardens, vineyards, fallow
lands, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan. Weed.
Genus 524. Serratula L.
1733. Serratula algida Iljin
Perennial. Rocks, gravelly and stony slopes, moraines. Alpine zone. North Turkestan. Medicinal.
1734. Serratula lancifolia Zakirov
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Malguzar. UzbRDB 1.
Genus 525. Carthamus L.
1735. Carthamus oxyacanthus M. Bieb.
Annual. Fallow lands, wastelands, pebbles, fields, banks of rivers and canals, surroundings of
settlements, stony slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul. Weed.
1736. *Carthamus tinctorius L.
Annual. Fields, fallow lands, wastelands, roadsides, fine-earth slopes (escaped from culture). Plain,
foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul. Dye, oil, food,
industrial.
1737. Carthamus turkestanicus Popov
Annual. Fine-earth, gravelly and stony slopes, wastelands, fields, fallow lands, roadsides, banks
of canals, surroundings of settlements. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Weed.
Genus 526. *Eclipta L.
1738. *Eclipta prostrata (L.) L.
Annual. Fields, fallow lands, banks of canals. Plain. Mirzachul. Weed.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 527. Bidens L.
1739. Bidens tripartita L.
Annual. Wet places, banks of rivers, lakes and canals, fields, fallow lands. Plain, foothills, montane
zone. Nuratau, Malguzar, North Turkestan, Mirzachul. Medicinal, essential oil, dye, weed.
Genus 528. Senecio L.
1740. Senecio franchetii C. Winkl.
Perennial. Fine-earth and stony slopes. Montane zone. North Turkestan.
1741. Senecio olgae Regel & Schmalh.
Perennial. Stony slopes. Montane zone. Malguzar, North Turkestan.
1742. Senecio subdentatus (Bunge) Ledeb. Figure 442
Annual. Sandy deserts, saline lands, banks of rivers and canals, pebbles, fine-earth, gravelly and
stony slopes. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Fodder.
Figure 442 Senecio subdentatus (photography by Natalya Beshko)
451
452
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 529. Ligularia Cass.
1743. Ligularia alpigena Pojark.
Perennial. Fine-earth, gravelly and stony slopes, screes, banks of rivers, swamps. Montane and alpine
zones. North Turkestan. Meliferous.
1744. Ligularia heterophylla Rupr.
Perennial. Fine-earth, gravelly and stony slopes, banks of rivers. Montane zone. Nuratau, Malguzar,
North Turkestan. Meliferous.
1745. Ligularia thomsonii (C. B. Clarke) Pojark. Figure 443
Perennial. Fine-earth, gravelly and stony slopes, screes, pebbles, moraines, subalpine meadows,
swamps, banks of rivers. Montane and alpine zones. Malguzar, North Turkestan. Poisonous, weed,
meliferous.
Figure 443 Ligularia thomsonii (photography by Natalya Beshko)
Genus 530. Tussilago L.
1746. Tussilago farfara L.
Perennial. River valleys, wet places, pebbles, fine-earth slopes. Montane zone. North Turkestan. Medicinal, essential oil, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 531. Karelinia Less.
1747. Karelinia caspia (Pall.) Less. Figure 444
Perennial. Saline lands, banks of rivers, lakes and canals, fields, fallow lands. Plain, foothills.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum. Weed, meliferous.
Figure 444 Karelinia caspia (photography by Natalya Beshko)
Genus 532. Inula L.
1748. Inula britannica L.
Perennial. Fine-earth slopes, wet places, banks of rivers and canals, fields, fallow lands. Plain,
foothills, montane zone. Malguzar, North Turkestan, Mirzachul. Medicinal, fodder, meliferous.
1749. Inula caspica F.K. Blum. ex Ledeb.
Annual or biennial. Saline lands, banks of rivers and canals, fields. Plain, foothills. Malguzar,
Mirzachul. Medicinal, meliferous.
1750. Inula glauca C. Winkl.
Perennial. Gravelly and stony slopes, rocks. Montane zone. Malguzar, North Turkestan.
1751. Inula helenium L.
Perennial. Fine-earth slopes, banks of rivers, wet places. Foothills, montane zone. Malguzar, North
Turkestan. Medicinal, ornamental, dye, meliferous.
1752. Inula orientalis Lam. [Inula macrophylla Kar. & Kir.]
Perennial. Fine-earth, gravelly slopes. Foothills, montane zone. Malguzar, North Turkestan.
Medicinal, dye, meliferous.
1753. Inula rhizocephala Schrenk
Perennial. Fine-earth, gravelly and stony slopes, alpine meadows. Montane and alpine zones.
Malguzar, North Turkestan. Medicinal, fodder.
453
454
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 533. Pulicaria Gaerth.
1754. Pulicaria dysenterica subsp. uliginosa Nyman [Pulicaria uliginosa Steven ex DC.]
Annual. Banks of rivers and canals, wet places. North Turkestan. Poisonous, insecticide.
1755. Pulicaria gnaphalodes (Vent) Boiss.
Perennial. Rocks, gravelly and stony slopes, pebbles, dry riverbeds. Plain, foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar. Essential oil.
1756. Pulicaria salviifolia Bunge
Perennial. Gravelly and stony slopes, pebbles, dry riverbeds, banks of rivers and canals. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Essential oil.
Genus 534. Pentanema Cass.
1757. Pentanema albertoregelia (C. Winkl.) Gorschk. [Vicoa albertoregelia C. Winkl.]
Perennial. Gravelly and stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
1758. Pentanema divaricatum Cass.
Annual. Gravelly and stony slopes, dry riverbeds, fields, wastelands. Plain, foothills. Nuratau.
Genus 535. Helichrysum Mill.
1759. Helichrysum maracandicum Popov Figure 445
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan.
Medicinal, ornamental.
Figure 445 Helichrysum maracandicum (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1760. Helichrysum mussae Nevski
Perennial. Gravelly and stony slopes, rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
1761. Helichrysum nuratavicum Krasch. Figure 446
Perennial. Gravelly and stony slopes, rocks. Montane zone. Nuratau. UzbRDB 2. Endemic to the
Nuratau Mountains.
Figure 446 Helichrysum nuratavicum (photography by N. Yu. Beshko)
1762. Helichrysum tianschanicum Regel
Perennial. Gravelly and stony slopes, pebbles, banks of rivers. Foothills, montane and alpine zones.
North Turkestan.
Genus 536. Laphangium (Hilliard & B. L. Burtt) Tzvelev
1763. Laphangium luteoalbum (L.) Tzvelev [Gnaphalium luteoalbum L.]
Annual. River valleys, wet places, banks of canals, fallow lands fields. Nuratau, Malguzar, North
Turkestan. Medicinal, fodder, ornamental, weed.
Genus 537. Gnaphalium L.
1764. Gnaphalium supinum L.
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Malguzar, North
Turkestan.
Genus 538. Leontopodium R. Br. ex Cass.
1765. Leontopodium fedtschenkoanum Beauverd
Perennial. Fine-earth slopes, meadows, swamps. Montane and alpine zones. North Turkestan.
Ornamental.
455
456
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 539. Filago L.
1766. Filago arvensis Lam.
Annual. Fine-earth, gravelly and stony slopes, piedmont plains, fields, fallow lands, roadsides,
wastelands. Plain, foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Medicinal, ornamental, weed.
1767. Filago filaginoides (Kar. & Kir.) Wagenitz [Evax filaginoides Kar. & Kir.]
Annual. Sandy deserts, saline lands, banks of lakes and canals. Plain. Kyzylkum.
1768. Filago germanica (L.) Huds. (Filago vulgaris Lam.)
Annual. Gravelly and stony slopes, fields, wastelands. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Mirzachul. Medicinal, weed.
1769. Filago pyramidata L.
Annual. Fine-earth, gravelly and stony slopes, piedmont plains, fields, fallow lands, roadsides,
wastelands. Foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, weed.
Genus 540. Cymbolaena Smoljan.
1770. Cymbolaena griffithii (A. Gray) Wagenitz
Annual. Fine-earth, gravelly and stony slopes, piedmont plains, fallow lands, wastelands. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Genus 541. Solidago L.
1771. Solidago kuhistanica Juz.
Perennial. Fine-earth and stony slopes, rocks, banks of rivers, meadows. Montane and alpine zones.
Nuratau, North Turkestan. Ornamental, meliferous.
Genus 542. Pseudolinosyris Novopokr.
1772. Pseudolinosyris grimmii (Regel & Schmalh.) Novopokr.
Semishrub. Fine-earth, gravelly and stony slopes, rocks, screes. Foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan.
Genus 543. Galatella Cass.
1773. Galatella coriacea Novopokr.
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane and alpine zones. Nuratau,
MAGNOLIOPSIDA
[EUDICOTS]
Malguzar, North Turkestan. Weed.
Genus 544. Kalimeris (Cass.) Cass.
1774. Kalimeris altaica (Willd.) Nees ex Fisch. Mey. & Avé-Lall. [Aster canescens (Ness) Fisjun]
Perennial. Fine-earth, gravelly and stony slopes, roadsides, banks of canals, fallow lands, fields.
Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul.
Genus 545. *Tripolium L.
1775. *Tripolium pannonicum subsp. tripolium (L.) Greuter
Annual. Saline lands, banks of canals, roadsides, fields, fallow lands. Plain. Nuratau Relic
Mountains, Mirzachul, Kyzylkum. Fodder, ornamental, meliferous.
Genus 546. Rhinactinidia Novopokr.
1776. Rhinactinidia popovii (Botsch.) Botsch.
Perennial. Rocks, stony slopes. Montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Genus 547. Erigeron L.
1777. Erigeron acris L.
Biennial. Meadows, fallow lands, saline lands, banks of rivers, lakes and canals, rocks, fineearth, gravelly and stony slopes. Plain, foothills, montane and alpine zones. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, weed.
1778. Erigeron bellidiformis Popov
Perennial. Fine-earth, gravelly and stony slopes. Alpine zone. North Turkestan
1779. Erigeron cabulicus (Boiss.) Botsch. [Psychrogeton cabulicus Boiss.]
Perennial. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Malguzar, North
Turkestan. Ornamental.
1780. *Erigeron canadensis L. [Conyza canadensis (L.) Cronquist]
Annual. Gardens, fields, fallow lands, wastelands, banks of rivers, lakes and canals, roadsides. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Medicinal, weed, meliferous.
1781. Erigeron khorossanicus Boiss. [Psychrogeton aucherii (DC.) Grierson]
Biennial. Fine-earth slopes, banks of rivers and canals, fallow lands. Foothills, montane and alpine
zones. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
457
458
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1782. Erigeron pallidus Popov
Perennial. Fine-earth, gravelly and stony slopes, pebbles, moraines. Montane and alpine zones. Malguzar,
NorthTurkestan.
1783. Erigeron petiolaris Vierh. [Erigeron pseudoneglectus Popov]
Perennial. Fine-earth slopes, subalpine and alpine meadows. Montane and alpine zones. North
Turkestan.
1784. Erigeron popovii Botsch.
Biennial or perennial. Meadows, banks of rivers. Montane zone. North Turkestan.
1785. Erigeron pseudoseravschanicus Botsch.
Perennial. Fine-earth and stony slopes, banks of rivers, meadows. Foothills, montane and alpine
zones. North Turkestan.
1786. Erigeron seravschanicus Popov
Perennial. Gravelly and stony slopes, shady wet places, subalpine meadows, moraines. Montane and
alpine zones. North Turkestan.
1787. Erigeron sogdianus Popov
Perennial. Subalpine and alpine meadows, rocks, screes, moraines. Montane and alpine zones. North
Turkestan.
1788. Erigeron umbrosus (Kar. & Kir.) Boiss.
Annual. Gravelly and stony slopes, screes, rocks, shady wet places. Montane and alpine zones. North
Turkestan.
Genus 548. Psychrogeton Boiss.
1789. Psychrogeton amorphoglossus (Boiss.) Novopokr. [Psychrogeton leucophyllus (Bunge)
Novopokr.]
Perennial. Gravelly and stony slopes, rocks. Montane and alpine zones. North Turkestan.
1790. Psychrogeton pseuderigeron (Bunge) Novopokr. ex Nevski
Perennial. Fine-earth, gravelly and stony slopes, rocks, screes, pebbles. Montane and alpine zones.
North Turkestan.
Genus 549. Symphyotrichum Nees
1791. Symphyotrichum ciliatum (Ledeb.) G. L. Nesom [Brachyactis ciliata (Ledeb.) Ledeb.]
Annual. Saline lands, banks of rivers, lakes and canals. Plain. Mirzachul, Kyzylkum.
Genus 550. Lachnophyllum Bunge
1792. Lachnophyllum gossypinum Bunge
Annual. Fine-earth, gravelly and stony slopes, piedmont plains, clay deserts, fallow lands, pebbles,
MAGNOLIOPSIDA
[EUDICOTS]
fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Medicinal, essential oil, fodder.
Genus 551. Anthemis L.
1793. Anthemis deserticola Krasch. & Popov
Annual. Saline lands. Plain. Mirzachul, Kyzylkum.
Genus 552. Achillea L.
1794. Achillea arabica Kotschy [Achillea biebersteinii Afan.] Figure 447
Perennial. Fine-earth, gravelly and stony slopes, pebbles, river valleys, piedmont plains, fallow
lands, wastelands, banks of canals. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal.
Figure 447 Achillea arabica (photography by Natalya Beshko)
1795. Achillea filipendulina Lam. Figure 448
Perennial. Fine-earth, gravelly and stony slopes, screes, pebbles, river valleys, fallow lands, banks
of canals, gardens. Foothills, montane and alpine zones. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan. Medicinal, essential oil, ornamental.
459
460
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 448 Achillea filipendulina (photography by Natalya Beshko)
1796. Achillea millefolium L. Figure 449
Perennial. River valleys, fine-earth slopes, meadows. Montane and alpine zones. Malguzar, North
Turkestan. Medicinal, ornamental.
Figure 449 Achillea millefolium (photography by Natalya Beshko)
1797. Achillea santolinoides subsp. wilhelmsii (K. Koch) Greuter [Achillea kermanica
Gand.] Figure 450
Perennial. Sandy and clay deserts, piedmont plains, fine-earth, gravelly and stony slopes, river
valleys, banks of canals, fallow lands, fields. Plain, foothills, montane zone. Nuratau, Aktau,
MAGNOLIOPSIDA
[EUDICOTS]
Nuratau Relic Mountains, Malguzar, Mirzachul, Kyzylkum. Medicinal.
Figure 450 Achillea santolinoides subsp. wilhelmsii (photography by Natalya Beshko)
461
462
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 553. Handelia Heimerl
1798. Handelia trichophylla Heimerl Figure 451
Perennial. Fine-earth, gravelly slopes, screes, pebbles, river valleys, fields, fallow lands. Plain,
foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal,
ornamental.
Figure 451 Handelia trichophylla (photography by Natalya Beshko)
Genus 554. Pseudohandelia Tzvelev
1799. Pseudohandelia umbellifera (Boiss.) Tzvelev Figure 452
Biennial or perennial. Sandy and clay deserts, fine-earth and gravelly slopes, fallow lands. Plain,
foothills. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Medicinal, ornamental.
Figure 452 Pseudohandelia umbellifera (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 555. Tanacetopsis (Tzvelev) Kovalevsk.
1800. Tanacetopsis karataviensis (Kovalevsk.) Kovalevsk.
Perennial. Stony slopes, rocks. Montane zone. Nuratau.
Figure 453 Tanacetopsis karataviensis (photography by Natalya Beshko)
Figure 453
463
464
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1801. Tanacetopsis mucronata (Regel & Schmalh.) Kovalevsk.
Perennial. Stony slopes, rocks, pebbles. Montane and alpine zones. North Turkestan.
1802. Tanacetopsis urgutensis (Popov ex Tzvel.) Kovalevsk.
Perennial. Gravelly and stony slopes, rocks. Montane and alpine zones. Malguzar, North Turkestan.
Genus 556. Lepidolopsis Poljakov
1803. Lepidolopsis turkestanica (Regel & Schmalh.) Poljakov
Perennial. Fine-earth, gravelly and stony slopes, fallow lands, pebbles, banks of rivers. Foothills,
montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Genus 557. Mausolea Bunge
1804. Mausolea eriocarpa (Bunge) Poljakov ex Podlech. Figure 454
Semishrub. Sandy deserts. Plain. Kyzylkum. Essential oil.
Figure 454 Mausolea eriocarpa (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 558. Artemisia L.
1805. Artemisia absinthium L.
Annual, perennial. River valleys, banks of canals, roadsides, fallow lands, gardens, meadows, wastelands, fine-earth, gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, food (spice-aromatic), essential oil,
poisonous.
1806. *Artemisia annua L.
Annual. River valleys, banks of canals, roadsides, fields, fallow lands, wastelands, gardens,
surroundings of settlements. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, essential oil, insecticide,
dye, weed.
1807. Artemisia cina Berg ex Poljakov
Semishrub. Saline lands, dry riverbeds, sandy soils. Plain. Mirzachul. Medicinal, essential oil,
poisonous.
1808. Artemisia diffusa Krasch. ex Poljakov
Semishrub. Sandy and clay deserts, pebbles, saline lands, fine-earth, gravelly and stony slopes. Plain,
foothills. Nuratau Relic Mountains, Mirzachul, Kyzylkum. Medicinal, essential oil, fodder.
1809. Artemisia dracunculus L.
Perennial. Fine-earth, gravelly and stony slopes, river valleys, fallow lands, wastelands. Plain,
foothills, montane zone. Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, food (spicearomatic), fodder, meliferous, weed, ornamental.
1810. Artemisia ferganensis Krasch. ex Poljakov
Semishrub. Sandy and clay deserts, fallow lands, fine-earth, gravelly and stony slopes, river valleys,
dry riverbeds, pebbles, screes. Plain, foothills, montane and alpine zones. Nuratau, Malguzar, North
Turkestan, Mirzachul. Medicinal, essential oil, fodder.
1811. Artemisia eremophila Krasch. & Butkov
Semishrub. Fine-earth, gravelly and stony slopes, pebbles, piedmont plains. Plain, foothills. Nuratau,
North Turkestan.
1812. Artemisia glanduligera Krasch. ex Poljakov
Semishrub. Pebbles, cliffs, fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan.
1813. Artemisia glaucina Krasch. ex Poljakov
Semishrub. Pebbles, banks of rivers and canals, fine-earth, gravelly and stony slopes, ravines.
Foothills, montane zone. Nuratau, Malguzar, North Turkestan.
1814. Artemisia juncea Kar. & Kir. Figure 455
Semishrub. Dry riverbeds, river valleys, pebbles, fine-earth, gravelly and stony slopes, screes, rocks.
Foothills, montane zone. Nuratau. Medicinal, essential oil.
465
466
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 455 Artemisia juncea (photography by Natalya Beshko)
1815. Artemisia lehmanniana Bunge
Semishrub. Fine-earth, gravelly and stony slopes, rocks. Montane and alpine zones. Malguzar, North
Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
1816. Artemisia leucodes Schrenk Figure 456
Annual or biennial. Sandy and clay deserts, fine-earth, gravelly and stony slopes, river valleys,
banks of canals, roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Medicinal, essential oil, fodder, weed.
Figure 456 Artemisia leucodes (photography by Natalya Beshko)
467
468
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1817. Artemisia persica Boiss.
Semishrub. Fine-earth, gravelly and stony slopes, pebbles, screes. Montane and alpine zones.
Malguzar, North Turkestan. Medicinal, essential oil.
1818. Artemisia porrecta Krasch. ex Poljakov
Semishrub. Banks of canals, roadsides, fallow lands, screes, dry riverbeds, pebbles, fine-earth,
gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, essential oil.
1819. Artemisia prolixa Krasch. ex Poljakov
Semishrub. Fine-earth, gravelly and stony slopes, screes. Foothills, montane zone. NorthTurkestan.
1820. Artemisia rutifolia Stephan ex Spreng.
Semishrub. Gravelly and stony slopes, rocks, pebbles, screes. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Medicinal, essential oil.
1821. Artemisia santolinifolia Turcz. ex Krasch.
Perennial. Dry riverbeds, banks of rivers, piedmont plains, fine-earth, gravelly and stony slopes.
Plain, foothills. Nuratau, North Turkestan. Medicinal, essential oil, fodder.
1822. Artemisia scoparia Waldst. & Kitam. Figure 457
Annual or biennial. Sandy and clay deserts, piedmont plains, river valleys, fine-earth, gravelly and
stony slopes, fields, fallow lands, roadsides, wastelands. Plain, foothills, montane and alpine zones.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
Medicinal, essential oil, insecticide, fodder, weed.
Figure 457 Artemisia scoparia (photography by Natalya Beshko)
1823. Artemisia scotina Nevski
Semishrub. Fine-earth, gravelly and stony slopes, piedmont plains. Plain, foothills, montane zone.
Nuratau, Malguzar. Essential oil.
1824. Artemisia serotina Bunge
Semishrub. Dry riverbeds, piedmont plains, fine-earth, gravelly and stony slopes. Foothills, montane
MAGNOLIOPSIDA
[EUDICOTS]
zone. Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, fodder.
1825. Artemisia sieversiana Ehrh.
Annual or biennial. Saline lands, banks of rivers and canals, roadsides, gardens, fields, fallow lands,
wastelands. Plain. Mirzachul. Medicinal, essential oil, fodder.
1826. Artemisia sogdiana Bunge Figure 458
Semishrub. Piedmont plains, fine-earth, gravelly and stony slopes, dry riverbeds. Foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal,
essential oil, fodder.
Figure 458 Artemisia sogdiana (photography by Natalya Beshko)
1827. Artemisia subsalsa Filatova
Semishrub. Saline lands, banks of rivers, lakes ands canals. Plain. Mirzachul.
1828. Artemisia tenuisecta Nevski Figure 459
Semishrub. Fine-earth, gravelly and stony slopes, screes. Montane zone. Nuratau, Aktau, Malguzar,
North Turkestan. Essential oil, fodder.
469
470
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 459 Artemisia tenuisecta (photography by Natalya Beshko)
1829. Artemisia tournefortiana Rchb.
Annual or biennial. River valleys, banks of canals, roadsides, fine-earth slopes, fields, pebbles,
wastelands, gardens, surroundings of settlements. Plain, foothills, montane zone. Nuratau, Aktau,
Malguzar, North Turkestan, Mirzachul. Weed.
1830. Artemisia turanica Krasch. Figure 460
Semishrub. Sandy and clay deserts, saline lands, pebbles, fine-earth, gravelly and stone slopes. Plain,
foothills. Nuratau Relic Mountains, Mirzachul, Kyzylkum. Essential oil, fodder.
Figure 460 Artemisia turanica (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1831. Artemisia vulgaris L.
Perennial. River valleys, ravines, banks of canals, roadsides, fields, fine-earth and stony slopes,
fallow lands, surroundings of settlements. Plain, foothills, montane and alpine zones. Nuratau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, essential oil,
insecticide, fodder, weed.
Genus 559. Trichantemis Regel & Schmalh.
1832. Trichanthemis karataviensis Regel & Schmalh.
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau.
Genus 560. Waldheimia Kar. & Kir.
1833. Waldheimia glabra (Decne) Regel
Perennial. Alpine meadows, moraines, rocks. Alpine zone. North Turkestan.
1834. Waldheimia tomentosa (Decne) Regel
Perennial. Gravelly and stony slopes, screes, rocks, pebbles, banks of rivers. Alpine zone. North
Turkestan.
Genus 561. Pyrethrum Zinn
1835. Pyrethrum galae Popov
Perennial. Fine-earth, gravelly and stony slopes, rocks. Montane zone. Nuratau.
1836. Pyrethrum pyrethroides (Kar. & Kir.) B. Fedtsch. ex Krasch. Figure 461
Perennial. Fine-earth, gravelly and stony slopes, screes, pebbles, moraines, rocks. Montane and
alpine zones. Malguzar, North Turkestan.
Figure 461 Pyrethrum pyrethroides (photography by Natalya Beshko)
471
472
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 562. Tanacetum L.
1837. Tanacetum parthenifolium (Willd.) Sch. Bip. [Pyrethrum parthenifolium Willd.]
Perennial. Ravines, banks of rivers, pebbles, rocks, stony slopes. Montane zone. Malguzar, North
Turkestan. Medicinal, essential oil, insecticide, ornamental.
1838. Tanacetum pseudachillea C. Winkl.
Perennial. Banks of rivers, fine-earth, gravelly and stony slopes. Montane and alpine zones. North
Turkestan. Medicinal, essential oil.
Genus 563. Lepidolopha C. Winkl.
1839. Lepidolopha komarowii C. Winkl.
Shrub. Fine-earth and stony slopes, rocks. Montane zone. Nuratau, Malguzar. Essential oil.
1840. Lepidolopha nuratavica Krasch. Figure 462
Shrub. Fine-earth and stony slopes, rocks. Montane zone. Nuratau. UzbRDB 1.
Figure 462 Lepidolopha nuratavica (photography by N. Yu. Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 564. Microcephala Pobed.
1841. Microcephala lamellata (Bunge) Pobed.
Annual. Sandy and clay deserts, saline lands, fine-earth, gravelly and stony slopes, rocks. Plain,
foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul,
Kyzylkum. Essential oil.
1842. Microcephala turcomanica (C. Winkl.) Pobed.
Annual. Sandy and clay deserts, wastelands. Plain. Mirzachul. Essential oil.
Genus 565. *Tripleurospermum Sch. Bip.
1843. *Tripleurospermum disciforme (C. A. Mey.) Sch. Bip.
Biennial. Fine-earth, gravelly and stony slopes, pebbles, river valleys, banks of canals, meadows,
roadsides, fallow lands, fields. Plain, foothills, montane zone. Malguzar, North Turkestan. Weed.
Genus 566. *Xanthium L.
1844. *Xanthium spinosum L.
Annual. Wastelands, roadsides, fallow lands, banks of canals, surroundings of settlements. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Medicinal, oil, poisonous, weed.
1845. *Xanthium strumarium L.
Annual. River valleys, banks of canals, wastelands, fields, fallow lands, roadsides, surroundings of
settlements. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan, Mirzachul, Kyzylkum. Medicinal, oil, poisonous, weed.
Family 104. Caprifoliaceae
Genus 567. Lonicera L.
1846. Lonicera bracteolaris Boiss. & Buhse. Figure 463
Shrub. Fine-earth, gravelly and stony slopes, river valleys, pebbles, ravines. Montane zone. Nuratau,
North Turkestan. Ornamental, meliferous.
473
474
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 463 Lonicera bracteolaris (photography by Natalya Beshko)
1847. Lonicera humilis Kar. & Kir. [Lonicera altmannii Regel & Schmalh.] Figure 464
Shrub. Fine-earth, gravelly and stony slopes, river valleys, ravines. Montane and alpine zones.
Malguzar, North Turkestan. Ornamental, meliferous.
Figure 464 Lonicera humilis (photography by Natalya Beshko)
1848. Lonicera korolkowii Stapf.
Shrub. Fine-earth, gravelly and stony slopes, river valleys, screes, rocks. Foothills, montane zone.
North Turkestan. Ornamental, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
1849. Lonicera microphylla Willd. ex Schult.
Shrub. Fine-earth and stony slopes, rocks, river valleys, subalpine meadows. Montane zone.
Malguzar, North Turkestan. Medicinal, meliferous, ornamental.
1850. Lonicera nummulariifolia Jaub. & Spach Figure 465
Shrub. Fine-earth, gravelly and stony slopes, river valleys, ravines, rocks. Montane and alpine zones.
Nuratau, Aktau, Malguzar, North Turkestan. Medicinal, meliferous, ornamental.
Figure 465 Lonicera nummulariifolia (photography by Natalya Beshko)
1851. Lonicera olgae Regel & Schmalh.
Shrub. Stony slopes, rocks. Montane and alpine zones. North Turkestan. Ornamental, meliferous.
475
476
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1852. Lonicera paradoxa Pojark.
Shrub. Stony slopes, rocks. Montane and alpine zones. North Turkestan. Ornamental, meliferous.
Threatened species.
1853. Lonicera semenovii Regel
Shrub. Stony slopes, rocks, moraines. Montane and alpine zones. North Turkestan. Ornamental,
meliferous.
1854. Lonicera simulatrix Pojark.
Shrub. Fine-earth, gravelly and stony slopes. Montane and alpine zones. North Turkestan.
1855. Lonicera zeravshanica Pojark.
Shrub. Fine-earth, gravelly and stony slopes, river valleys. Montane and alpine zones. North
Turkestan. Ornamental, meliferous.
Genus 568. Dipsacus L.
1856. Dipsacus dipsacoides (Kar. & Kir.) Botsch. Figure 466
Perennial. Fine-earth, gravelly and stony slopes. Montane zone. Nuratau, Malguzar, North Turkestan,
Mirzachul. Medicinal, ornamental, meliferous.
Figure 466 Dipsacus dipsacoides (photography by Natalya Beshko)
1857. Dipsacus laciniatus L.
Biennial. Fine-earth slopes, river valleys, fields, gardens, fallow lands. Plain, foothills, montane
MAGNOLIOPSIDA
[EUDICOTS]
zone. Nuratau, Malguzar, North Turkestan, Mirzachul. Medicinal, ornamental, meliferous.
Genus 569. *Cephalaria Schrad.
1858. Cephalaria syriaca (L.) Schrad. ex Roem. & Schult.
Annual. Piedmont plains, fallow lands, fields. Plain, foothills, montane zone. Nuratau, Malguzar,
North Turkestan, Mirzachul. Medicinal, oil.
Genus 570. Scabiosa L.
1859. Scabiosa flavida Boiss. & Hausskn.
Annual. Fine-earth, gravelly and stony slopes. Foothills. Nuratau.
1860. Scabiosa olivierii Coult.
Annual. Sandy and clay deserts, piedmont plains, gravelly and stony slopes. Plain, foothills, montane
zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
1861. Scabiosa rhodantha Kar. & Kir.
Annual. Fine-earth, gravelly and stony slopes, banks of rivers, piedmont plains, sandy and clay
deserts, fields, fallow lands. Foothills, montane zone. Nuratau, Nuratau Relic Mountains, Kyzylkum.
1862. Scabiosa songarica Schrenk Figure 467
Perennial. Fine-earth, gravelly and stony slopes. Foothills, montane and alpine zones. Nuratau,
Malguzar, North Turkestan. Medicinal, ornamental, meliferous.
Figure 467 Scabiosa songarica (photography by Natalya Beshko)
477
478
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 571. Valeriana L.
1863. Valeriana chionophila Popov & Kult. Figure 468
Perennial. Fine-earth and stony slopes, rocks, wet places near melting snow. Foothills, montane and
alpine zones. Nuratau, Aktau, Malguzar, North Turkestan. Medicinal.
Figure 468 Valeriana chionophila (photography by Natalya Beshko)
1864. Valeriana fedtschenkoi Coincy
Perennial. Rocks, stony slopes, screes, banks of rivers, subalpine and alpine meadows. Montane and
alpine zones. North Turkestan. Medicinal.
1865. Valeriana ficarifolia Boiss.
Perennial. Shady wet places, banks of rivers, ravines, fine-earth, gravelly and stony slopes, screes.
Foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan.
Genus 572. Valerianella Mill.
1866. Valerianella coronata (L.) DC.
Annual. Fine-earth, gravelly and stony slopes, fallow lands, pebbles. Plain, Foothills. North
Turkestan, Mirzachul.
MAGNOLIOPSIDA
[EUDICOTS]
1867. Valerianella cymbocarpa C. A. Mey.
Annual. Fine-earth, gravelly and stony slopes, dry riverbeds. Plain, foothills, montane zone. Nuratau,
Malguzar, North Turkestan.
1868. Valerianella dactylophylla Boiss. & Hohen.
Annual. Fine-earth and stony slopes, shady places. Montane zone. Nuratau, Malguzar.
1869. Valerianella muricata (Steven ex M. Bieb.) W. H. Baxter
Annual. Fine-earth and stony slopes, screes, rocks, pebbles, fallow lands. Plain, foothills, montane
zone. Nuratau, Malguzar, Mirzachul.
1870. Valerianella oxyrrhyncha Fisch. & C. A. Mey. [Valerianella diodon Boiss.]
Annual. Fine-earth and stony slopes, pebbles, banks of rivers, fields, fallow lands. Plain, foothills,
montane and alpine zones. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul.
1871. Valerianella plagiostephana Fisch. & C. A. Mey.
Annual. Fine-earth and stony slopes, screes, rocks, pebbles, river valleys. Plain, foothills, montane
zone. Malguzar, North Turkestan.
1872. Valerianella szovitsiana Fisch. & C. A. Mey.
Annual. Sandy and clay deserts, saline lands, fine-earth and stony slopes, rocks, screes. Plain,
foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan, Mirzachul, Kyzylkum.
1873. Valerianella turkestanica Regel & Schmalh.
Annual. Fine-earth, gravelly and stony slopes, pebbles, banks of rivers, clay deserts, saline lands,
fields, fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan,
Mirzachul.
1874. Valerianella vvedenskyi Lincz.
Annual. Stony slopes, river valleys, pebbles. Foothills, montane zone. North Turkestan.
Family 105. Apiaceae [Umbelliferae]
Genus 573. Eryngium L.
1875. Eryngium caeruleum M. Bieb.
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, river valleys, banks of canals, gardens,
fallow lands, roadsides, wastelands, fields. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar, North Turkestan, Mirzachul. Medicinal, weed.
1876. Eryngium macrocalyx Schrenk
Perennial, monocarpic. Stony and gravelly slopes, screes, dry riverbeds, fallow lands, roadsides.
Foothills, montane zone. Nuratau, Malguzar, North Turkestan. Medicinal, essential oil.
1877. Eryngium octophyllum Korovin
Perennial, polycarpic. Stony slopes, rocks, dry riverbeds. Foothills, montane zone. Nuratau,
Malguzar. Medicinal, essential oil.
479
480
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 574. Echinophora L.
1878. Echinophora tenuifolia subsp. sibthorpiana (Guss.) Tutin [Echinophora sibthorpiana Guss.]
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, piedmont plains, wastelands, fallow
lands, fields, pebbles, gardens, roadsides, banks of canals. Plain, foothills. Nuratau, Aktau, Nuratau
Relic Mountains, Malguzar. Medicinal, essential oil, food (spice-aromatic), fodder.
Genus 575. Anthriscus Pers.
1879. Anthriscus glacialis Lipsky
Perennial, polycarpic. Stony slopes, stream banks. Montane and alpine zones. Nuratau. Fodder.
Genus 576. Scandix L.
1880. Scandix nodosa L. [Chaerophyllum nodosum (L.) Crantz; Myrrhoides nodosa (L.)
Cannon; Physocaulis nodosus (L.) W. D. J . Koch]
Annual. River valleys, shady wet places. Foothills, montane zone. Nuratau. Food, essential oil.
1881. Scandix pecten-veneris L.
Annual. Fine-earth, gravelly and stony slopes, river valleys, banks of canals, fields, gardens,
fallow lands, wastelands, roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic
Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Weed, medicinal.
1882. Scandix stellata Banks & Sol.
Annual. Fine-earth, gravelly and stony slopes, screes, rocks, pebbles, dry riverbeds, roadsides, fields,
fallow lands. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar,
North Turkestan. Weed.
Genus 577. Kozlovia Lipsky
1883. Kozlovia paleacea Lipsky
Perennial, monocarpic. Fine-earth slopes, fallow lands. Foothills, montane zone. Nuratau, Malguzar,
North Turkestan. Weed.
Genus 578.*Torilis Adans.
1884. *Torilis arvensis (Huds.) Link.
Annual. Fallow lands, fields, gardens, roadsides, wastelands, banks of rivers and canals, fine-earth,
gravelly and stony slopes. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
MAGNOLIOPSIDA
[EUDICOTS]
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Essential oil, weed.
1885. *Torilis leptophylla (L.) Rchb. f.
Annual. Fine-earth, gravelly and stony slopes, wastelands, fallow lands, fields, roadsides, banks of
rivers and canals, gardens, surroundings of settlements. Foothills, montane zone. Nuratau, Aktau,
Nuratau Relic Mountains, Malguzar, North Turkestan, Mirzachul, Kyzylkum. Weed, food.
Genus 579. Turgenia Hoffm.
1886. Turgenia latifolia (L.) Hoffm.
Annual. Fields, fallow lands, sandy and clay deserts, river valleys, fine-earth, gravelly and stony
slopes, dry riverbeds. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Essential oil, weed.
Genus 580. *Orlaya Hoffm.
1887. *Orlaya grandiflora (L.) Hoffm.
Annual. Fields, fallow lands. Plain. Mirzachul. Essential oil, weed.
Genus 581. Daucus L.
1888. Daucus carota L.
Biennial. Roadsides, wastelands, fallow lands, fields, gardens, banks of canals, river valleys,
meadows. Foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul, Kyzylkum. Food, essential oil, medicinal, weed.
Genus 582. *Coriandrum L.
1889. *Coriandrum sativum L.
Annual. Surrounds of settlements, roadsides, fallow lands, fields, river valleys (escaped from
culture). Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North
Turkestan, Mirzachul. Food (spice-aromatic), essential oil, meliferous.
Genus 583. Schrenkia Fisch. & C. A. Mey.
1890. Schrenkia golickeana B. Fedtsch.
Perennial, monocarpic. Fine-earth and stony slopes, pebbles, rocks. Foothills, montane and alpine
zones. Nuratau, Malguzar, North Turkestan. Medicinal.
481
482
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1891. Schrenkia pungens Regel & Schmalh.
Perennial, polycarpic. Stony slopes, rocks, screes, dry riverbeds. Foothills, montane zone. Nuratau,
Malguzar.
1892. Schrenkia vaginata (Ledeb.) Fisch. & C. A. Mey.
Perennial, monocarpic. Stony slopes, rocks, screes, pebbles. Montane and alpine zones. Nuratau,
North Turkestan.
Genus 584. Lipskya (K.-Pol.) Nevski
1893. Lipskya insignis Nevski
Perennial, monocarpic. Stony slopes, rocks. Foothills, montane zone. Malguzar. Essential oil, food
(spice-aromatic). Threatened species, UzbRDB 3.
Genus 585. Schtschurowskia Regel & Schmalh.
1894. Schtschurowskia meifolia Regel & Schmalh.
Perennial, polycarpic. Stony slopes, screes, pebbles. Montane and alpine zones. Malguzar, North
Turkestan.
Genus 586. Korshinskya Lipsky
1895. Korshinskya olgae Lipsky
Perennial, monocarpic. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau,
Malguzar, North Turkestan.
Genus 587. Aulacospermum Ledeb.
1896. Aulacospermum roseum Korovin
Perennial, monocarpic. Gravelly and stony slopes, screes, alpine meadows, stream banks, wet places
near melting snow. Montane and alpine zones. Malguzar, North Turkestan.
1897. Aulacospermum simplex Rupr.
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, screes, rocks. Montane and alpine
zones. Malguzar, North Turkestan.
1898. Aulacospermum tenuisectum Korovin
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, screes, rocks. Montane and alpine
zones. Malguzar, North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Genus 588. Eremodaucus Bunge
1899. Eremodaucus lehmannii Bunge
Annual. Fields, fallow lands, wastelands, roadsides, piedmont plains, fine-earth slopes. Plain,
foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan,
Mirzachul, Kyzylkum. Fodder, weed.
Genus 589. *Conium L.
1900. *Conium maculatum L.
Biennial. Wastelands, gardens, fields, banks of rivers and canals. Foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Medicinal, poisonous, essential oil, weed.
Genus 590. Prangos Lindl.
1901. Prangos didyma (Regel) Pimenov & V. N. Tikhom.
Perennial, monocarpic. Sandy and clay deserts, piedmont plains, dry riverbeds. Plain, foothills.
Nuratau Relic Mountains, Mirzachul, Kyzylkum. Fodder, essential oil, meliferous.
1902. Prangos fedtschenkoi (Regel & Schmalh.) Korovin Figure 469
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, dry riverbeds, pebbles, roadsides,
fields. Foothills, montane zone. Nuratau, Aktau, Malguzar, North Turkestan. Fodder, essential oil,
meliferous.
Figure 469 Prangos fedtschenkoi (photography by Natalya Beshko)
1903. Prangos ornata Kuzmina Figure 470
Perennial, polycarpic. Fine-earth, gravelly and stony slopes, river valleys, dry riverbeds. Foothills,
montane zone. Nuratau. Fodder, essential oil, meliferous.
483
484
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 470 Prangos ornata (photography by Natalya Beshko)
1904. Prangos pabularia Lindl. Figure 471
Perennial, polycarpic. Fine-earth, gravelly and stony slopes, river valleys, dry riverbeds. Montane
and alpine zones. Nuratau, Malguzar, North Turkestan. Fodder, essential oil, medicinal, meliferous.
Figure 471 Prangos pabularia (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
Genus 591. Bupleurum L.
1905. Bupleurum falcatum L. subsp. exaltatum (M. Bieb.) Rouy & Camus [Bupleurum exaltatum
M. Bieb.]
Perennial, polycarpic. Stony slopes, screes, rocks, dry riverbeds, pebbles. Montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Medicinal, fodder.
Genus 592. Elaeosticta Fenzl
1906. Elaeosticta allioides (Regel & Schmalh.) Kljuykov, Pimenov & V. N. Tikhom.
Perennial, monocarpic. Fine-earth and stony slopes, dry riverbeds. Foothills, montane zone. Nuratau,
Aktau, Malguzar, North Turkestan. Essential oil, fodder.
1907. Elaeosticta hirtula (Regel & Schmalh.) Kljuykov, Pimenov & V. N. Tikhom.
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, screes. Foothills, montane and alpine
zones. Nuratau, Malguzar, North Turkestan. Essential oil.
1908. Elaeosticta polycarpa (Korovin) Kljuykov, Pimenov & V. N. Tikhom.
Perennial, polycarpic. Fine-earth slopes. Foothills, montane zone. Nuratau Relic Mountains.
Essential oil.
1909. Elaeosticta vvedenskyi (Kamelin) Kljuykov, Pimenov & V. N. Tikhom.
Perennial, monocarpic. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau.
Essential oil.
Genus 593. Galagania Lipsky
1910. Galagania fragrantissima Lipsky
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, fallow lands. Foothills, montane zone.
Nuratau, Malguzar, North Turkestan. Essential oil, food (spice-aromatic).
1911. Galagania tenuisecta (Regel & Schmalh.) M. G. Vassiljeva & Pimenov
Perennial, monocarpic. Fine-earth, gravelly slopes, dry riverbeds, fallow lands. Foothills, montane
zone. Nuratau, Malguzar, North Turkestan. Essential oil, weed.
Genus 594. Hyalolaena Bunge
1912. Hyalolaena depauperata Korovin
Perennial, monocarpic. Gravelly and stony slopes, rocks. Foothills, montane zone. Nuratau. Essential oil.
1913. Hyalolaena jaxartica Bunge
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, sandy and clay deserts, dry riverbeds,
pebbles, rocks. Plain, foothills. Nuratau, Mirzachul. Essential oil.
485
486
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 595. Oedibasis Koso.-Pol.
1914. Oedibasis apiculata (Kar. & Kir.) Koso.-Pol.
Perennial, monocarpic. Sandy deserts, piedmont plains, fine-earth, gravelly and stony slopes, rocks,
fallow lands. Plain, foothills, montane and alpine zones. Nuratau, Malguzar, North Turkestan.
1915. Oedibasis tamerlanii (Lipsky) Korovin ex Nevski
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, rocks. Foothills, montane zone.
Nuratau, Malguzar, North Turkestan.
Genus 596. Elwendia Boiss.
1916. Elwendia capusii (Franch.) Pimenov & Kljuykov [Bunium capusii (Franch.) Korovin]
Perennial, polycarpic. Piedmont plains, fine-earth slopes, river valleys. Foothills, montane zone.
Nuratau, Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan.
1917. Elwendia chaerophylloides (Regel & Schmalh.) Pimenov & Kljuykov [Bunium
chaerophylloides (Regel & Schmalh.) Drude]
Perennial, polycarpic. Fine-earth, gravelly and stony slopes. Foothills, montane zone. Nuratau,
Aktau, Nuratau Relic Mountains, Malguzar, North Turkestan. Medicinal, food (spice-aromatic),
essential oil.
1918. Elwendia intermedia (Korovin) Pimenov & Kljuykov [Bunium intermedium Korovin]
Perennial, polycarpic. Fine-earth slopes, river valleys, shady wet places. Montane zone. Nuratau,
Malguzar, North Turkestan.
1919. Elwendia persica (Boiss.) Pimenov & Kljuykov [Bunium persicum (Boiss.) B. Fedtsch.]
Perennial, polycarpic. Fine-earth, gravelly and stony slopes, screes. Foothills, montane zone.
Nuratau, Malguzar, North Turkestan. Medicinal, food (spice-aromatic), essential oil.
Genus 597. Falcaria Fabr.
1920. Falcaria vulgaris Bernh.
Perennial, monocarpic. River valleys, banks of canals, fine-earth, gravelly slopes, wet places,
gardens, roadsides. Plain, foothills, montane zone. Nuratau, Nuratau Relic Mountains, Malguzar,
North Turkestan. Medicinal, food, essential oil.
Genus 598. Carum L.
1921. Carum carvi L.
Biennial. Banks of rivers, wet places. Montane and alpine zones. Nuratau, Malguzar, North
Turkestan. Medicinal, essential oil, food (spice-aromatic).
MAGNOLIOPSIDA
[EUDICOTS]
Genus 599. Apium L.
1922. Apium nodiflorum (L.) Lag.
Perennial, polycarpic. Banks of rivers and canals. Nuratau. Medicinal, food.
Genus 600. Cuminum L.
1923. Cuminum setifolium (Boiss.) Koso-Pol.
Annual. Sandy and clay deserts, fine-earth slopes. Nuratau, Kyzylkum. Essential oil, weed.
Genus 601. Aphanopleura Boiss.
1924. Aphanopleura capillifolia (Regel & Schmalh.) Lipsky
Annual. Sandy and clay deserts, dry riverbeds, fine-earth, gravelly and stony slopes, riparian forests,
fallow lands, roadsides. Plain, foothills, montane zone. Nuratau, Aktau, Nuratau Relic Mountains,
Malguzar, North Turkestan, Mirzachul, Kyzylkum. Essential oil, weed.
Genus 602. Pimpinella L.
1925. Pimpinella peregrina L.
Biennial. Banks of rivers and canals, gardens, roadsides, fields, fallow lands, riparian forests.
Foothills, montane zone. Nuratau, Malguzar, North Turkestan. Essential oil, medicinal, fodder,
meliferous.
Genus 603. Sium L.
1926. Sium sisaroideum DC.
Perennial, polycarpic. Banks of rivers and canals, swamps. Plain, foothills, montane zone. Nuratau.
Medicinal, oil.
Genus 604. Berula W. D. J. Koch
1927. Berula erecta (Huds.) Coville
Perennial, polycarpic. Banks of rivers and canals. Foothills, montane zone. Nuratau, Malguzar, North
Turkestan. Medicinal, poisonous.
487
488
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 605. Seseli L.
1928. Seseli calycinum (Korovin) Pimenov & Sdobnina
Perennial, polycarpic. Rocks. Montane zone. Nuratau, Malguzar, North Turkestan.
1929. Seseli korovinii Schischk. Figure 472
Perennial, polycarpic. Screes, rocks, gravey and stony slopes. Montane and alpine zones. Nuratau,
Malguzar, North Turkestan.
Figure 472 Seseli korovinii (photography by Natalya Beshko)
1930. Seseli lehmannianum Boiss.
Perennial, polycarpic. Fine-earth, gravelly and stony slopes, rocks, pebbles. Montane and alpine
zones. Malguzar, North Turkestan.
1931. Seseli mucronatum (Schrenk) Pimenov & Sdobnina
Perennial, polycarpic. Meadows, river valleys, fine-earth and stony slopes, rocks. Montane and
alpine zones. Malguzar, North Turkestan.
1932. Seseli seravschanicum Pimenov & Sdobnina Figure 473
Perennial, monocarpic. Stony slopes, rocks. Montane and alpine zones. North Turkestan.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 473 Seseli seravschanicum (photography by Natalya Beshko)
1933. Seseli tenuisectum Regel & Schmalh.
Perennial, polycarpic. Rocks, screes, pebbles. Montane zone. North Turkestan.
1934. Seseli turbinatum Korovin
Perennial, polycarpic. Stony slopes. Montane zone. Nuratau. UzbRDB 2.
Genus 606. Libanotis Haller ex Zinn
1935. Libanotis schrenkiana C. A. Mey. ex Schischk. [Seseli schrenkianum (C. A. Mey. ex
Schischk.) Pimenov & Sdobnina]
Perennial, polycarpic. Fine-earth and stony slopes, pebbles, rocks. Montane and alpine zones. North
Turkestan.
489
490
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 607. Mediasia Pimenov
1936. Mediasia macrophylla (Regel & Schmalh.) Pimenov Figure 474
Perennial, polycarpic. Fine-earth, gravelly and stony slopes, screes. Montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Medicinal, food (spice-aromatic), essential oil.
Figure 474 Mediasia macrophylla (photography by Natalya Beshko)
Genus 608. Alposelinum Pimenov
1937. Alposelinum albomarginatum (Schrenk) Pimenov
Perennial, monocarpic. Stony slopes, moraines, screes, pebbles, alpine meadows. Montane and
alpine zones. North Turkestan.
Genus 609. Conioselinum Hoffm.
1938. Conioselinum vaginatum (Spreng.) Thell. [Conioselinum tataricum Fisch. ex Hoffm.]
Perennial, polycarpic. Fine-earth, gravelly and stony slopes, ravines, river valleys, pebbles, screes,
shady wet places, swamp meadows. Montane and alpine zones. North Turkestan.
Genus 610. Angelica L.
1939. Angelica archangelica subsp. decurrens (Ledeb.) Kuvaev
Perennial, monocarpic. Stream banks, wet places near melting snow, springs, wet gravelly and
fine-earth slopes, meadows, pebbles. Montane and alpine zones. Nuratau, North Turkestan.
Medicinal, essential oil, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
1940. Angelica brevicaulis (Rupr.) B. Fedtsch. Figure 475
Perennial, monocarpic. Banks of rivers, pebbles, moraines, wet gravelly slopes, screes, swamp
meadows. Montane and alpine zones. North Turkestan. Medicinal, essential oil.
Figure 475 Angelica brevicaulis (photography by Natalya Beshko)
1941. Angelica ternata Regel & Schmalh.
Perennial, polycarpic. Gravelly slopes, screes, moraines, rocks, pebbles. Montane and alpine zones.
North Turkestan. Essential oil, food (spice-aromatic).
Genus 611. Ferula L.
1942. Ferula angreni Korovin Figure 476
Perennial, polycarpic. Fine-earth and gravelly slopes, river valleys, screes. Montane zone. Nuratau,
Malguzar. Essential oil, fodder, meliferous.
Figure 476 Ferula angreni (photography by Natalya Beshko)
491
492
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1943. Ferula bucharica (Lipsky) Koso-Pol. [Ladyginia bucharica Lipsky]
Perennial, polycarpic. Fine-earth, stony and gravelly slopes. Montane zone. Malguzar. Essential oil,
saponin-bearing, meliferous.
1944. Ferula diversivittata Regel & Schmalh. Figure 477
Perennial, monocarpic. Fine-earth slopes, dry riverbeds. Foothills, montane zone. Nuratau.
Medicinal, essential oil, meliferous.
Figure 477 Ferula diversivittata (photography by Natalya Beshko)
1945. Ferula dshizakensis Korovin
Perennial, polycarpic. Stony and gravelly slopes, rocks. Montane zone. Nuratau, Nuratau Relic
Mountains, Malguzar. Essential oil, fodder, meliferous.
1946. Ferula fedtschenkoana Koso-Pol.
Perennial, polycarpic. Fine-earth, gravelly slopes. Montane and alpine zones. Malguzar, North
Turkestan. Essential oil, melliferous. Threatened species, UzbRDB 2.
1947. Ferula ferganensis Lipsky ex Korovin
Perennial, polycarpic. Stony slopes, rocks, screes. Montane zone. Nuratau. Essential oil, meliferous.
1948. Ferula foetida (Bunge) Regel Figure 478
Perennial, monocarpic. Sandy and clay deserts, piedmont plains, fine-earth slopes. Plain, foothills.
Nuratau, Nuratau Relic Mountains, Kyzylkum. Medicinal, essential oil, fodder, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
Figure 478 Ferula foetida (photography by Natalya Beshko)
1949. Ferula helenae Rakhm. & Melibaev
Perennial, monocarpic. Stony slopes. Foothills. Nuratau Relic Mountains. Essential oil, fodder,
meliferous. Threatened species. Endemic to the Nuratau Relic Mountains.
1950. Ferula kokanica Regel & Schmalh. Figure 479
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, screes, pebbles, rocks. Foothills,
montane and alpine zones. Nuratau, Nuratau Relic Mountains, Malguzar, North Turkestan.
Medicinal, essential oil, fodder, meliferous.
Figure 479 Ferula kokanica (photography by Natalya Beshko)
493
494
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
1951. Ferula kuhistanica Korovin Figure 480
Perennial, monocarpic. Fine-earth, gravelly and stony slopes. Montane and alpine zones. Aktau,
Malguzar, North Turkestan. Medicinal, essential oil, fodder, meliferous.
Figure 480 Ferula kuhistanica (photography by Natalya Beshko)
MAGNOLIOPSIDA
[EUDICOTS]
1952. Ferula lehmannii Boiss.
Perennial, monocarpic. Sandy deserts. Plain. Kyzylkum. Essential oil, meliferous.
1953. Ferula mollis Korovin
Perennial, polycarpic. Fine-earth, gravelly and stony slopes. Foothills, montane zone. North
Turkestan. Essentialoil, fodder, meliferous.
1954. Ferula moschata (H. Reinsch) Koso-Pol. [Ferula sumbul (Kauffm.) Hook. f.] Figure 481
Perennial, polycarpic. Fine-earth and stony slopes, screes, rocks, ravines. Montane zone. Nuratau,
Malguzar, North Turkestan. Medicinal, essential oil, meliferous. Threatened species, UzbRDB 2.
Figure 481 Ferula moschata (photography by N. Yu. Beshko)
1955. Ferula ovczinnikovii Pimenov
Perennial, monocarpic. Stony and gravelly slopes, screes. Montane zone. Malguzar. Essential oil,
meliferous.
1956. Ferula ovina (Boiss.) Boiss. Figure 482
Perennial, polycarpic. Fine-earth, gravelly and stony slopes, rocks. Montane and alpine zones.
Nuratau, Aktau, Malguzar, North Turkestan. Essential oil, fodder, meliferous.
495
496
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Figure 482 Ferula ovina (photography by Natalya Beshko)
1957. Ferula penninervis Regel & Schmalh. Figure 483
Perennial, polycarpic. Stony slopes, screes, pebbles, dry riverbeds, rocks. Montane and alpine zones.
Nuratau, Malguzar, North Turkestan. Essential oil, medicinal, fodder, meliferous.
Figure 483 Ferula penninervis (photography by Natalya Beshko)
1958. Ferula samarkandica Korovin
Perennial, monocarpic. Fine-earth, gravelly and stony slopes, dry riverbeds, rocks. Montane and
alpine zones. Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, fodder, meliferous.
MAGNOLIOPSIDA
[EUDICOTS]
1959. Ferula schtschurowskiana Regel & Schmalh.
Perennial, monocarpic. Sandy and clay, deserts, saline lands, gravelly slopes. Plain, foothills,
montane zone. Nuratau, Malguzar, North Turkestan, Mirzachul. Essential oil, fodder, meliferous.
1960. Ferula varia (Schrenk) Trautv.
Perennial, monocarpic. Sandy deserts, fine-earth and stony slopes. Plain, foothills. Nuratau Relic
Mountains. Medicinal, essential oil, fodder, meliferous.
Genus 612. Heracleum L.
1961. Heracleum lehmannianum Bunge Figure 484
Perennial or biennial, monocarpic. Subalpine meadows, river valleys, ravines, pebbles, wet screes.
Montane and alpine zones. Nuratau, Malguzar, North Turkestan. Medicinal, essential oil, fodder,
meliferous.
Figure 484 Heracleum lehmannianum (photography by Natalya Beshko)
1962. Heracleum olgae Regel & Schmalh. [Tetrataenium olgae (Regel & Schmalh.) Manden]
Perennial, monocarpic. Screes, pebbles, fine-earth slopes. Montane and alpine zones. North
Turkestan. Essential oil.
497
498
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Genus 613. Semenovia Regel & Herd.
1963. Semenovia dasycarpa (Regel & Schmalh.) Korovin Figure 485
Perennial, polycarpic. Fine-earth and stony slopes, ravines, stream banks, shady wet places, rocks.
Montane and alpine zones. North Turkestan.
Figure 485 Semenovia dasycarpa (photography by Natalya Beshko)
1964. Semenovia heterodonta Manden
Perennial, polycarpic. Rocks, screes, stony slopes. Alpine zone. North Turkestan.
1965. Semenovia pimpinelloides (Nevski) Manden
Perennial, polycarpic. Stony slopes, rocks. Montane zone. Nuratau.
REFERENCES
REFERENCES
Adylov T. A. 1983. Conspectus Florae Asiae Mediae. Vol. 7. Tashkent: Fan Publishers, 415. (In
Russian)
Adylov T. A. 1987. Conspectus Florae Asiae Mediae. Vol. 9. Tashkent: Fan Publishers, 400. (In
Russian)
Adylov T. A., Zuckerwanik T. I. 1993. Conspectus Florae Asiae Mediae. Vol. 10. Tashkent: Fan
Publishers, 692. (In Russian)
Akhani H., Edwards G. & Roalson E. H. 2007. Diversification of the world Salsoleae s.l.
(Chenopodiaceae): molecular phylogenetic analysis of nuclear and chloroplast datasets and a revised
classification. Int. J. Pl. Sci., 168: 931–956.
Alibekov L. A., Nishanov S. A. 1978. Natural conditions and resources of Dzhizak region. Tashkent:
Uzbekistan Publishers, 254. (In Russian)
Al-Shehbaz I. A., German D.A., Mummenhoff K., Moazzeni H. 2014. Systematics, tribal placements,
and synopses of the Malcolmia s.l. segregates (Brassicaceae). Harvard Papers in Botany, London, 19
(1): 53–71.
Aramov, S. A. 2012. Geographical Atlas of Uzbekistan. Tashkent: Goskomgeodeskadastr, 119. [In
Russian]
Azimova D. E. 2017. Flora of the Malguzar Ridge. Tashkent: Abstr. PhD Diss., 44. (In Uzbek and
Russian)
Batoshov A. R. 2016. Flora of South-East Kyzylkum Relic Mountains. Tashkent: Abstr. Doct. Diss.,
75. (In Uzbek, Russian and English)
Batoshov A. R., Beshko N. Yu. 2013. Characteristics of the flora and plant cover of the relic
mountains of the South Kyzylkum. Arid Ecosystems, 19 (3): 73–78. (In Russian)
Batoshov A. R., Beshko N. Yu. 2015. Comparative analysis of geophytes of the flora of relic
mountains of South-eastern Kyzylkum and Nuratau Mountains. Uzbek. Biol. Journ., 5: 29–33. (In
Russian)
Belolipov I. V. 1973. Interesting floristic findings in the vegetation of Nuratau Mountains. Uzbek.
Biol. Journ., 2: 48–49. (In Russian)
Beshko N. Yu. 2000a. Flora of the Nuratau Nature Reserve. Tashkent: Abstr. Cand. Diss., 26 (In
Russian)
Beshko N. Yu. 2000b. Flora of the planned biosphere reserve Nuratau-Kyzylkum. In: Biodiversity
conservation on the protected areas of Uzbekistan. Tashkent: Chinor ENK, 21–43. (In Russian)
Beshko N. Yu. 2011. Flora of vascular plants of the Nuratau nature reserve. In: Proceedings of nature
reserves of Uzbekistan. Issue 7. Tashkent: Chinor ENK, 19–78. (In Russian)
Beshko N. Yu., Azimova D. E. 2013. New floristic findings on the North-West Pamir-Alay
499
500
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
(Uzbekistan). Turczaninowia, (16) 1: 197–203. (In Russian)
Beshko N. Yu., Azimova D. E. 2014. The genus Astragalus L. in the flora of Nuratau and Malguzar
mountains: the comparative analysis. Uzb. Biol. Journ., Special issue: 20-21. (In Russian)
Beshko N. Yu., Tojibaev K. Sh. & Batoshov A. R. 2013a. Tulips of the Nuratau Mountains and
South-Eastern Kyzylkum (Uzbekistan). Stapfia Reports, 99: 198–204.
Beshko N. Yu., Tojibaev K. Sh., Batoshov A. R. & Azimova D. E. 2014. Botanical-geographical
regionalization of Uzbekistan. Kuhistan and Nuratau districts. Uzbek. Biol. Journ., 3: 30–34. (In
Russian)
Beshko, N. Yu., Zagrebin, V., Popov, A., Khasanov, F., Mitropolskaya, O., Magdiev, Kh. 2013.
Recommendations for Protected Areas System Development in Uzbekistan. UNDP, GEF, Main
Department of Forestry of Ministry of Agriculture and Water Resources of the Republic of
Uzbekistan. Tashkent: Baktria Press, 256. [In Russian].
Bondarenko O. N., Nabiev M. M. 1972. Conspectus Florae Asiae Mediae. Vol. 3. Tashkent: Fan
Publishers, 266. (In Russian)
Botirova L. A. 2012. Vegetation of the Zaaminsu River basin. Tashkent: Abstr. Cand. Diss., 24 (In
Uzbek)
Botschantzev V. P., Butkov A. Ya., Vvedensky A. I., Nikiforova N. B. & Pazij V. K. 1961. An
identification guide of wild plants of the Hungry Steppe. Tashkent:Tashkent State University, 216. (In
Russian)
Brummitt R. K., Powell C. E. 1992. Authors of plant names. Royal Botanic Gardens, Kew. 732 p.
Budantsev A.L. 1996. Plant resources of Russia and adjacent states: Flowering plants, their
chemical composition and use. Part I. Families Lycopodiaceae – Ephedraceae. Part II. Addenda. St.Petersburg: World and Family -95, 571. (In Russian)
CABI. 2017. Invasive Species Compendium. CAB International, Wallingford, UK. Available from:
www.cabi.org/isc.
Christenhusz M. J. M., Reveal J. L., Farjon A., Gardner M. F., Mill R. R. & Chase M. W. 2011a. A
linear sequence of extant families and genera of lycophytes and ferns. Phytotaxa, 19: 7–54.
Christenhusz M. J. M., Reveal J. L., Farjon A., Gardner M. F., Mill R. R. & Chase M. W. 2011b. A
new classification and linear sequence of extant gymnosperms. Phytotaxa, 19: 55-70.
Clayton W.D., Vorontsova M.S., Harman K.T. & Williamson H. 2006. GrassBase - The Online World
Grass Flora. Available from: http://www.kew.org/data/grasses-db.html.
Czukavina A. P. 1984. Flora of the Tajik SSR. Vol. 7. Academy of Sciences of USSR, Leningrad,
563. (In Russian)
Degtjareva G. V., Kljuykov E. V., Samigullin T. H., Valiejo-Roman C. M. & Pimenov M. G. 2013.
ITS phylogeny of Middle Asian geophilic Umbelliferae-Apioideae genera with comments on their
morphology and utility of psbA-trnH sequences. Plant Syst. Evol., 299: 985–1010.
Demurina E. M. 1975. Vegetation of the western part of Turkestan ridge and its spurs. Tashkent: Fan
Publishers, 189. (In Russian)
Downie S. R., Spalik K., Katz-Downie D. S. & Reduron J.-P. 2010. Major clades within Apiaceae
subfamily Apioideae as inferred by phylogenetic analysis of nrDNA ITS sequences. Plant Div. Evol.,
REFERENCES
128: 111–136.
Egorova T. V. 1999. Sedges (Carex L.) of Russia and adjanced states (within the limits of the former
USSR). St. Petersburg: State Chemical-Pharmaceutical Academy, 771.
eMonocot: an online resource for monocot plants. Available from:http://e-monocot.org/.
Esankulov A. S. 2012. Flora of the Zaamin Nature Reserve. Tashkent: Abstr. Cand. Diss., 19 (In
Russian)
Fedorov A. A. 1984. Plant resources of the U.S.S.R.: Flowering plants, their chemical composition
and use. Families Magnoliaceae–Limoniaceae. Leningrad: Science Publishers, 460. (In Russian)
Gammerman A. F., Grom I. I. 1976. Wild medicinal plants of the USSR. Moscow: Medicine
Publishers, 288. (In Russian)
Geltman D. V. 2013. Spurges (Euphorbia L., Euphorbiaceae) of East Europe and the Caucasus in the
mirrow of new system of the genus. Turczaninowia, 16 (2): 30–40. (In Russian)
Hill L. Fritillaria: a list of published names. 2017. Available from: http://www.fritillariaicones.com/
info/names/frit.names.pdf.
Horandl E., Emadzade K. 2012. Evolutionary classification: A case study on the diverse plant genus
Ranunculus L. (Ranunculaceae). Perspectives in Plant Ecology, Evolution and Systematics, 14: 310–324.
ILDIS World Database of Legumes (version 10, 2005). Available from: www.ildis.com.
IUCN. 2020 The IUCN Red List of Threatened Species. Available from: http://www.iucnredlist.org.
IUCN. 2020. The IUCN Red List of Threatened Species. Version 2020-1. Available from: http://
www.iucnredlist.org.
IUCN/ISSC Invasive Species Specialist Group (ISSG). 2014. Global Invasive Species Database
version 2014-2. Available from: http://193.206.192.138/gisd/
Jizzax Viloyat Hokimligi. Official web-site. Available from: http://jizzax.uz.
Kamelin R V. 1973a. Florogenetic analysis of the native flora of the Mountain Middle Asia.
Leningrad: Science Publishers, 133-138. (in Russian)
Kamelin R V. 1973b. To the knowledge of the flora of Nuratau Mountains. Botan. Journ., 58 (5):
625–637. (In Russian).
Kamelin R V. 1990. Flora of Syrdarya Karatau: Materials for floristic regionalization of Middle Asia.
Leningrad: Science Publishers 70–107. (in Russian)
Kamelin R. V. 1979. The Kuhistan district of mountainous Middle Asia: Botanical-geographical
analysis. Leningrad: Science Publishers, 166. (In Russian)
Kamelin R. V., Kovalevskaja S. S., Nabiev M. M. 1981. Conspectus Florae Asiae Mediae. Vol. 6.
Tashkent: Fan Publishers, 395. (In Russian)
Khassanov F. O. 2015. Conspectus Florae Asiae Mediae. Vol. 11. Tashkent: Fan Publishers, 456. (In
Russian)
Khassanov F. O., Esankulov A. S. & Tirkasheva M. B. 2013. Flora of the Zaamin Nature Reserve.
Tashkent: [s.n.], 119. (In Russian)
Khassanov F. O., Rakhimova N. 2012. Taxonomical revision of genus Iris L. (Iridaceae Juss.) in the
Flora of Central Asia. Stapfia: reports, 97: 121-126.
Kinzikaeva G. K. 1988. Flora of the Tajik SSR. Vol. 9. Academy of Sciences of USSR, Leningrad,
501
502
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
568. (In Russian)
Kochkareva T. F. 1986. Flora of the Tajik SSR. Vol. 8. Academy of Sciences of USSR, Leningrad,
519. (In Russian)
Konnov A. A. 1973. Flora of juniper forests of the Shakhristan. Dushanbe: Donish Publishers, 176. (In
Russian)
Konnov A. A. 1990. Juniper formations of Central Asia and adjacent areas. Novosibirsk: Abstr. Doct.
Diss., 34. (In Russian)
Korovin E. P. 1923. Plant formations of the Nurata Valley. In: Proceedings of the Turkestan Scientific
Society. Vol. 1. Tashkent: Turkestan Publishing House, 2–5. (In Russian)
Korovin E. P. 1934. Vegetation of Middle Asia and South Kazakhstan. Moscow & Tashkent: [s.n.],
479. (In Russian)
Korovin E. P. 1961. Vegetation of Middle Asia and South Kazakhstan. Vol. 1. Tashkent: [s.n.], 452. (In
Russian)
Korovin E. P. 1962. Vegetation of Middle Asia and South Kazakhstan. Vol. 2. Tashkent: [s.n.], 547. (In
Russian).
Kovalevskaja S. S. 1963. Conspectus Florae Asiae Mediae. Vol. 1. Tashkent: Fan Publishers, 226. (In
Russian)
Kovalevskaja S. S. 1971. Conspectus Florae Asiae Mediae. Vol. 2. Tashkent: Fan Publishers, 360. (In
Russian)
Kudrjashev, S. N. 1941. Flora of Uzbekistan (Vol.1). Tashkent: Uzbek Department of the Academy
of Sciences of the USSR. [In Russian]
Kudryashev S. N. 1941. Flora of Uzbekistan. Vol. 1. Uzbek Department of the Academy of Sciences
of the USSR, Tashkent, 568. (In Russian)
Kultiassov M. V. 1923. Essay on the vegetation of the Pistalitau Mountains. In: Proceedings of the
Turkestan Scientific Society. Tashkent: Turkestan Publishing House, 89–102. (In Russian)
Kurmikov A. G., Belolipov I. V. 2012. Wild medicinal plants of Uzbekistan. Tashkent: Extremum
Press, 278. (In Russian)
Lammers T. G. 2007. World checklist and bibliography of Campanulaceae. Kew, UK: Royal Botanic
Gardens, 675.
Larin I. V. 1956. Forage plants of hayfields and pastures of the USSR. (In 3 vol). Moskow,
Leningrad: State agricultural publishing house. (In Russian)
Lazkov G. A. 2016. Family Labiatae Juss. in flora of Kyrgyzstan. Pocheon, Republic of Korea:
Geobook, 384.
Lopez-Vinyallonga S., Mehregan I., Garcia-Jacas N., Tscherneva O., Susanna A. & Kadereit
J.W. 2009. Phylogeny and evolution of the Arctium-Cousinia complex (Compositae, CardueaeCarduinae). Taxon, 58 (1): 153–171. (In Russian)
Lopez-Vinyallonga S., Romaschenko K., Susanna A. & Garcia-Jacas N. 2011. Systematics of the
Arctioid group: Disentangling Arctium and Cousinia (Cardueae, Carduinae). Taxon, 60(2): 539–554.
Nabiev M. M. 1986. Conspectus Florae Asiae Mediae. Vol. 8. Tashkent: Fan Publishers, 191. (In
Russian)
REFERENCES
Nikitin V. V. 1983. Weeds of the flora of the USSR. Leningrad: Science Publishers, 454. (In Russian)
Olson D. M., Dinerstein E. 2002. The Global 200: Priority ecoregions for global conservation.
Annals of the Missouri Botanical Garden, 89(2): 199–224.
Ovczinnikov P. N. 1957. Flora of the Tajik SSR. Vol. 1. Academy of Sciences of the USSR, Moscow &
Leningrad, 547. (In Russian)
Ovczinnikov P. N. 1963. Flora of the Tajik SSR. Vol. 2. Academy of Sciences of the USSR, Moscow &
Leningrad, 456. (In Russian)
Ovczinnikov P. N. 1968. Flora of the Tajik SSR. Vol. 3. Academy of Sciences of the USSR,
Leningrad, 711. (In Russian)
Ovczinnikov P. N. 1975. Flora of the Tajik SSR. Vol. 4. Academy of Sciences of the USSR,
Leningrad, 576. (In Russian)
Ovczinnikov P. N. 1978. Flora of the Tajik SSR. Vol. 5. Academy of Sciences of the USSR,
Leningrad, 678. (In Russian).
Ovczinnikov P. N. 1981. Flora of the Tajik SSR. Vol. 6. Academy of Sciences of the USSR,
Leningrad, 727. (In Russian)
Pachomova M. G. 1974. Conspectus Florae Asiae Mediae. Vol. 4. Tashkent: Fan Publishers, 274. (In
Russian)
Pachomova M. G. 1976. Conspectus Florae Asiae Mediae. Vol. 5. Tashkent: Fan Publishers, 375. (In
Russian)
Podlech D. & Zarre S. (with collaboration of Ekici M., Maassoumi A. A. R. & Sytin A.). 2013. A
taxonomic revision of the genus Astragalus L. (Leguminosae) in the Old World (vols. 1–3). Wien:
Naturhistorisches Museum, 2439.
Popov M. G., Androsov N. V. 1937. Vegetation of the Guralash Nature Reserve and Zaamin forestry.
Tashkent: Department of Sciences, 40. (In Russian).
Pratov U. P. 1998. The Red Data Book of the Republic of Uzbekistan. Vol. 1. Plants. Tashkent:
Chinor ENK. 335. (in Uzbek and Russian)
Pratov U. P. 2006. The Red Data Book of the Republic of Uzbekistan. Vol. 1. Plants and Fungi.
Tashkent: Chinor ENK. 250. (in Uzbek, Russian and English)
Pratov U. P., Khassanov F. O. The Red Data Book of the Republic of Uzbekistan. Vol. 1. Plants and
Fungi. 2009. Tashkent: Chinor ENK. 360. (in Uzbek, Russian and English)
Rassulova M. R. 1991. Flora of the Tajik SSR. Vol. 10. Academy of Sciences of USSR, Leningrad,
624. (In Russian)
Sadykov A.S. 1984. The Red Data Book of the Uzbek SSR. Vol. 2. Plants. Tashkent. 150. (in Uzbek
and Russian)
Salmaki Y., Zarre S., Ryding O., Lindqvist C., Schneunert A., Brauchler C. & Heubl G. 2012.
Phylogeny of the tribe Phlomideae (Lamioideae: Lamiaceae) with special focus on Eremostachys
and Phlomoides: New insights from nuclear and chloroplast sequences. Taxon, 61 (1): 161–179.
Sennikov A. N., Tojibaev K. Sh, Khassanov F. O., Beshko N. Yu. 2016. The Flora of Uzbekistan
Project. Phytotaxa, 282 (2): 107–118.
Sennikov, A. N. 2016. Flora of Uzbekistan (Vol.1). Toshkent: Navroz Publishers, xviii + 173. [In
503
504
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Russian]
Sennikov, A. N. 2017. Flora of Uzbekistan (Vol. 2). Toshkent: Navroz Publishers, xii + 200. [In
Russian]
Sokolov P. D. 1986. Plant resources of the U. S. S. R.: Flowering plants, their chemical composition
and use. Families Paeoniaceae – Thymelaeaceae. Leningrad: Science Publishers, 336. (In Russian)
Sokolov P. D. 1987. Plant resources of the U. S. S. R.: Flowering plants, their chemical composition
and use. Families Hydrangeaceae – Haloragaceae. Leningrad: Science Publishers, 326. (In Russian)
Sokolov P. D. 1988. Plant resources of the U. S. S. R.: Flowering plants, their chemical composition
and use. Families Rutaceae – Elaeagnaceae. Leningrad: Science Publishers, 357. (In Russian)
Sokolov P. D. 1990. Plant resources of the U. S. S. R.: Flowering plants, their chemical composition
and use. Families Caprifoliaceae – Plantaginaceae. Leningrad: Science Publishers, 328. (In Russian)
Sokolov P. D. 1991. Plant resources of the U.S.S.R.: Flowering plants, their chemical composition
and use. Families Hippuridaceae – Lobeliaceae. St.-Petersburg: Science Publishers, 200. (In Russian)
Sokolov P. D. 1993. Plant resources of the U. S. S. R.: Flowering plants, their chemical composition
and use. Familia Asteraceae (Compositae). St.-Petersburg: Science Publishers, 352. (In Russian)
Sokolov P. D. 1994. Plant resources of Russia and adjacent states: Flowering plants, their chemical
composition and use. Families Butomaceae – Typhaceae. St.-Petersburg: Science Publishers, 271. (In
Russian)
Takhtajan A. 1986. Floristic regions of the World. Los Angeles, London: Berkeley, 522.
The Angiosperm Phylogeny Group. 2016. An update of the Angiosperm Phylogeny Group
classification for the orders and families of flowering plants: APG IV. Botanical Journal of the
Linnean Society, 181 (1): 1–20.
The Global Compositae Checklist. 2009. Available from: http://www.compositae.org/checklist.
The Gymnosperm Database. 2017. Available from: http://www.conifers.org/.
The International Plant Names Index. 2012. Available from: http://www.ipni.org.
The Plant List (Version 1.1). 2017. Available from: http://www.theplantlist.org/.
The Republic of Uzbekistan: Biodiversity Conservation, National Strategy and Action Plan. 1998.
Tashkent. Available from: https://www.cbd.int/doc/world/uz/uz-nr-01-en.pdf.
The State Committee of the Republic of Uzbekistan on Statistics. Official web-site. 2017. Available
from: http://www.stat.uz/.
The World Umbellifer Database. 1999. Available from: http://www.umbellifers.com/frames.
html?http://rbg-web2.rbge.org.uk/URC/urchomepage.html.
Tirkasheva M. B. 2011. Vegetation of the Sanzar River basin. Tashkent: Abstr. Cand. Diss., 21. (In
Uzbek)
Tojibaev K. Sh., Beshko N. Yu. & Popov V. A. 2016. Botanical-geographical regionalization of
Uzbekistan. Bot. Journ., 101 (10): 1105–1132. (In Russian)
Tojibaev K. Sh., Beshko N. Yu. 2007. The cadaster of rare and endemic plants of Dzhizak and Navoi
provinces of the Republic of Uzbekistan. In: Biodiversity of Uzbekistan — monitoring and use.
Tashkent: [s.n.], 200-208. (In Russian).
Tojibaev K. Sh., Beshko N. Yu., Karimov F. I., Batoshov A. R., Turginov O. T. & Azimova D. 2014a.
REFERENCES
The Data Base of Flora of Uzbekistan. Journal for Arid Land Studies, 24 (1): 157–160.
Tojibaev K. Sh., Beshko N.Yu., Popov V. A., Jang C. G. & Chang K. S. 2017. Botanical Geography
of Uzbekistan. Pocheon, Republic of Korea: Korea National Arboretum, 250.
Tojibaev K.Sh., Beshko N.Yu. 2015. Reassessment of diversity and analysis of distribution in Tulipa
(Liliaceae) in Uzbekistan. Nordic Journal of Botany, 33: 324–334.
Tojibaev K.Sh., Beshko N.Yu., Azimova D. E. & Turginov O. T. 2015. Distribution patterns of
species of the genus Astralalus L. (sect. Macrocystis, Laguropsis and Chaetodon) in the territory of
Mountain Middle Asian province. Turczaninowia, 18 (2): 17–38. (In Russian)
Tsvelev N. N. 1983. Grasses of the Soviet Union. New Delhi: Oxonian Press, 1196.
UNDP, State Committee for Nature Protection, Academy Sciences of the Repubic of Uzbekistan.
2015. The Fifth National Report of the Republic of Uzbekistan on the Biodiversity Conservation.
Tashkent: [s.n.], 58. [In Russian]
Vvedensky A. I. 1953. Flora of Uzbekistan. Vol. 2. Academy of Sciences of the Uzbek SSR,
Tashkent, 549. (In Russian)
Vvedensky A. I. 1955. Flora of Uzbekistan. Vol. 3. Academy of Sciences of the Uzbek SSR,
Tashkent, 825. (In Russian)
Vvedensky A. I. 1959. Flora of Uzbekistan. Vol. 4. Academy of Sciences of the Uzbek SSR,
Tashkent, 507. (In Russian)
Vvedensky A. I. 1961. Flora of Uzbekistan. Vol. 5. Academy of Sciences of the Uzbek SSR,
Tashkent, 667. (In Russian)
Vvedensky A. I. 1962. Flora of Uzbekistan. Vol. 6. Academy of Sciences of the Uzbek SSR,
Tashkent, 630. (In Russian)
Wiegleb G., Bobrov A. A. & Zalewska-Galosz J. 2017. A taxonomic account of Ranunculus section
Batrachium (Ranunculaceae). Phytotaxa, 319 (1): 001–055.
Williams M. W., Konovalov V. G. 2008. Central Asia temperature and precipitation data, 1879-2003:
USA National Snow and Ice Data Center. Available from: http://nsidc.org/data/docs/noaa/g02174_
central_asia_data/index.html.
Zakirov K. Z. 1955. Flora and vegetation of the Zeravschan river basin. (Vol. 1.) Tashkent: [s.n.],
205 p. (In Russian)
Zakirov K. Z. 1962. Flora and vegetation of the Zeravschan river basin. (Vol. 2.) Tashkent: [s.n.],
446 p. (In Russian)
Zakirov K. Z., Burygin V. A. 1956. About some relict plants of the Nuratau ridge. Botan Journ., 41 (9):
1331–1332. (In Russian).
Zakirov P. K. 1969. Vegetation cover of the Nuratau Mountains. Tashkent: Fan Publishers, 142. (In
Russian)
Zakirov P. K. 1971. Botanical geography of low mountains of Kyzylkum and Nuratau ridge.
Tashkent: Fan Publishers. 203. (In Russian)
Zakirov, K. Z. 1971. Vegetation cover of Uzbekistan and the ways of its practical use (I-vol).
Tashkent: Fan Publishers. [In Russian]
Zakirov, K. Z. 1976. Vegetation cover of Uzbekistan and the ways of its practical use (III-vol).
505
506
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Tashkent: Fan Publishers. [In Russian]
Zakirov, K. Z. 1984. Vegetation cover of Uzbekistan and the ways of its practical use (IV-vol).
Tashkent: Fan Publishers. [In Russian]
Zakirov, K. Z., 1973. Vegetation cover of Uzbekistan and the ways of its practical use (II-vol).
Tashkent: Fan Publishers. [In Russian]
INDEX OF LATIN NAMES
INDEX OF LATIN NAMES
A
Abutilon 271
Abutilon therophrasti 271
Acanthocephalus 428
Acanthocephalus amplexifolius 428
Acanthocephalus benthamianus 429
Acantholimon 297
Acantholimon erythraeum 297
Acantholimon nuratavicum 298
Acantholimon subavenaceum 299
Acantholimon tataricum 299
Acanthophyllum 326
Acanthophyllum aculeatum 326
Acanthophyllum elatius 326
Acanthophyllum gypsophiloides 326
Acanthophyllum pungens 326
Acantholimon zakirovii 299
Acer 265
Acer pubescens 265
Acer semenovii 266
Acer turkestanicum 266
Achillea 459
Achillea arabica 459
Achillea biebersteinii 459
Achillea filipendulina 459
Achillea kermanica 460
Achillea millefolium 460
Achillea santolinoides subsp.
wilhelmsii 460
Achnatherum splendens 107
Achnatherum turkomanicum 107
Acinos rotundiflorus 411
Aconitum 149
Aconitum talassicum 149
Aconitum zeravschanicum 149
Aconogonon coriarum 311
Aconogonon hissaricum 312
Acroptilon repens 449
Adiantum 36
Adiantum capillus-veneris 36
Adonis 147
Adonis aestivalis 147
Adonis turkestanica 148
Aegilops 128
Aegilops crassa 128
Aegilops cylindrica 128
Aegilops juvenalis 128
Aegilops squarrosa 128
Aegilops triuncialis 128
Aellenia glauca 340
Aellenia iliensis 340
Aellenia subaphylla 340
Aeluropus 117
Aeluropus intermedius 117
Aeluropus litoralis 117
Aethionema 293
Aethionema carneum 293
Agrimonia 229
Agrimonia asiatica 229
Agriophyllum 334
Agriophyllum lateriflorum 334
Agropyron 126
Agropyron badamense 126
Agropyron drobovii 132
Agropyron ferganense 131
Agropyron intermedium 132
Agropyron lachnophyllum 131
Agropyron lolioides 132
Agropyron repens 132
Agropyron setuliferum 126
Agropyron tianschanicum 132
Agropyron trichophorum 132
Agropyron ugamicum 131
Agrostemma 319
Agrostemma githago 319
Agrostis 111
Agrostis gigantea 111
Agrostis hissarica 112
Agrostis olympica 112
Agrostis stolonifera 112
Ailanthus 269
Ailantus altissima 269
Alcea 270
Alcea litvinovii 270
Alcea nudiflora 271
Alcea rhyticarpa 271
Alchemilla 228
Alchemilla fontinalis 228
Alchemilla krylovii 228
Alchemilla sibirica 228
Alhagi 211
Alhagi canescens 211
Alhagi kirghisorum 211
Alhagi pseudalhagi 211
Alisma 49
Alisma plantago-aguatica 49
Alismataceae 49
Alliaria 274
Alliaria petiolata 274
Allium 76
Allium alexeianum 76
Allium altissimum 77
Allium barsczewskii 77
Allium caesium 77
Allium carolinianum 78
Allium caspium 78
Allium clausum 79
Allium cupuliferum subsp.
cupuliferum 80
Allium cupuliferum subsp.
nuratavicum 80
Allium drepanophyllum 81
Allium filidens 81
Allium griffithianum 81
Allium gusaricum 82
Allium inconspicuum 82
Allium isakulii 82
Allium jodanthum 83
Allium karataviense 83
Allium kaufmannii 84
Allium kokanicum 84
Allium komarowii 85
Allium levichevii 85
Allium longicuspis 85
Allium oreodictyum 85
Allium oreophiloides 85
Allium oreophilum 85
Allium oschaninii 86
Allium praemixtum 86
Allium protensum 87
Allium sarawschanicum 87
Allium stephanophorum 87
Allium stipitatum 87
Allium suworowii 88
Allium taeniopetalum 88
Allium talassicum 88
Allium turkestanicum 89
Allium verticillatum 89
Allium xiphopetalum 90
Allochrusa 326
Allochrusa gypsophiloides 326
Alopecurus 111
Alopecurus arundinaceus 111
Alopecurus himalaicus 111
Alopecurus myosuroides 111
Alopecurus pratensis 111
Alposelinum 490
Alposelinum albomarginatum 490
Althaea 271
Althaea armeniaca 271
Althaea officinalis 271
Alyssum 285
Alyssum alyssoides 285
Alyssum campestre 285
Alyssum dasycarpum 285
Alyssum desertorum 286
Alyssum linifolium 286
Alyssum marginatum 286
Alyssum szovitsianum 286
Amaranthaceae 329
Amaranthus 329
Amaranthus albus 329
Amaranthus blitoides 329
Amaranthus deflexus 329
507
508
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Amaranthus retroflexus 329
Amaryllidaceae 76
Amberboa 448
Amberboa turanica 448
Ammodendron 174
Ammodendron conollyi 174
Ammodendron lehmannii 174
Ammothamnus lehmanni 173
Amoria bonannii 177
Amoria fragifera 177
Amygdalus bucharica 233
Amygdalus communis 235
Amygdalus spinosissima 237
Anabasis 340
Anabasis eriopoda 340
Anabasis jaxartica 341
Anabasis salsa 341
Anacaridaceae 265
Anagallis 349
Anagallis arvensis 349
Anagallis foemina 349
Anchusa 363
Anchusa arvensis subsp. orientalis 363
Anchusa azurea 363
Anchusa italica 363
Andrachne 251
Andrachne fedtschenkoi 251
Andrachne telephioides 251
Androsace 349
Androsace dasyphylla 349
Androsace maxima 349
Androsace sericea 349
Anemone 157
Anemone biflora var. petiolulosa 157
Anemone narcissiflora subsp.
protracta 157
Anemone petiolulosa 157
Anemone protracta 157
Anemone tschernaewii 158
Angelica 490
Angelica archangelica subsp.
decurrens 490
Angelica brevicaulis 490
Angelica ternata 491
Angiospermae 47
Angiosperms 47
Anthemis 459
Anthemis deserticola 459
Anthriscus 480
Anthriscus glacialis 480
Antonia debilis 411
Anura pallidivirens 438
Apera 112
Apera interrupta 112
Aphanopleura 487
Aphanopleura capillifolia 487
Apiaceae 479
Apium 487
Apium nodiflorum 487
Apocynaceae 358
Aquilegia 145
Aquilegia lactiflora 145
Aquilegia vicaria 123 145
Arabidopsis 275
Arabidopsis parvula 274
Arabidopsis pumila 275
Arabidopsis thaliana 275
Arabis 277
Arabis auriculata 277
Arabis kokanica 276
Arabis montbretiana 277
Araceae 48
Arceuthobium 294
Arceuthobium oxycedri 294
Arctium 436
Arctium aureum 436
Arctium chloranthum 437
Arctium haesitabundum 437
Arctium karatavicum 437
Arctium korolkowii 437
Arctium leiospermum 437
Arctium pallidivirens 438
Arenaria 318
Arenaria griffithii 318
Arenaria leptoclados 319
Arenaria rotundifolia 318
Arenaria serpyllifolia 319
Arenaria serpyllifolia subsp.
leptoclados 319
Aristida karelinii 105
Armeniaca vulgaris 233
Arnebia 360
Arnebia coerulea 360
Arnebia decumbens 361
Arnebia euchroma 362
Arnebia obovata 362
Arnebia transcaspica 362
Artemisia 465
Artemisia absinthium 465
Artemisia annua 465
Artemisia cina 465
Artemisia diffusa 465
Artemisia dracunculus 465
Artemisia eremophila 465
Artemisia ferganensis 465
Artemisia glanduligera 465
Artemisia glaucina 465
Artemisia juncea 465
Artemisia lehmanniana 466
Artemisia leucodes 467
Artemisia persica 468
Artemisia porrecta 468
Artemisia prolixa 468
Artemisia rutifolia 468
Artemisia santolinifolia 468
Artemisia scoparia 468
Artemisia scotina 468
Artemisia serotina 468
Artemisia sieversiana 469
Artemisia sogdiana 469
Artemisia subsalsa 469
Artemisia tenuisecta 469
Artemisia tournefortiana 470
Artemisia turanica 470
Artemisia vulgaris 471
Arum 48
Arum korolkowii 48
Asparagaceae 90
Asparagus 91
Asparagus brachyphyllus 91
Asparagus persicus 91
Asperuginoides 286
Asperuginoides axillaris 286
Asperugo 367
Asperugo procumbens 367
Asperula 351
Asperula aparine 353
Asperula glabrata 351
Asperula humifusa 352
Asperula oppositifolia 351
Asperula setosa 351
Asperula trichodes 351
Asphodelaceae 70
Aspleniaceae 36
Asplenium 37
Asplenium ruta-muraria 37
Asplenium trichomanes 37
Aster canescens 457
Asteraceae 422
Astragalus 182
Astragalus adpressipilosus 182
Astragalus alabugensis 182
Astragalus alopecias 182
Astragalus alpinus 182
Astragalus ammophilus 194
Astragalus ammotrophus 182
Astragalus angreni 183
Astragalus aphanassjievii 183
Astragalus aschuturi 183
Astragalus bactrianus 183
Astragalus bakaliensis 183
Astragalus belolipovii 183
Astragalus camptoceras 183
Astragalus campylorrhynchus 184
Astragalus campylotrichus 184
Astragalus chodshenticus 184
Astragalus commixtus 184
Astragalus compositus 184
Astragalus contortuplicatus 184
Astragalus cornu-bovis 185
Astragalus corydalinus 194
Astragalus dendroides 185
Astragalus dictamnoides 185
Astragalus dipelta 185
Astragalus dolichocarpus 185
Astragalus eximius 185
Astragalus falcigerus 185
Astragalus farctissimus 185
Astragalus ferganensis 185
Astragalus filicaulis 185
Astragalus flexus 186
Astragalus floccosifolius 186
Astragalus globiceps 187
Astragalus guttatus 187
Astragalus harpilobus 187
Astragalus inaequalifolius 187
INDEX OF LATIN NAMES
Astragalus intarrensis 187
Astragalus isfahanicus 187
Astragalus iskanderi 187
Astragalus jagnobicus 187
Astragalus juratzkanus 187
Astragalus kelleri 187
Astragalus knorringianus 188
Astragalus korovinianus 194
Astragalus kurdaicus 192
Astragalus lasiosemius 188
Astragalus lasiostylus 188
Astragalus leiophysa 188
Astragalus lentilobus 188
Astragalus leptophysus 189
Astragalus leptostachys 190
Astragalus lipskyi 190
Astragalus lithophilus 190
Astragalus macrocladus 190
Astragalus macronyx 191
Astragalus macropetalus 192
Astragalus macrotropis 192
Astragalus marguzaricus 192
Astragalus masenderanus 192
Astragalus maverranagri 187
Astragalus mucidus 193
Astragalus nematodes 193
Astragalus nivalis 193
Astragalus nobilis 193
Astragalus nuciferus 194
Astragalus ophiocarpus 194
Astragalus orbiculatus 194
Astragalus oxyglottis 194
Astragalus patentipilosus 194
Astragalus patentivillosus 194
Astragalus paucijugus 194
Astragalus pauper 194
Astragalus peduncularis 194
Astragalus persipolitanus 194
Astragalus petunnikovii 194
Astragalus platyphyllus 195
Astragalus pterocephalus 195
Astragalus pulcher 195
Astragalus quisqualis 195
Astragalus retamocarpus 195
Astragalus rubromarginatus 195
Astragalus russanovii 195
Astragalus saratagius 195
Astragalus sarytavicus 195
Astragalus schachimardanus 195
Astragalus schmalhausenii 195
Astragalus schrenkianus 195
Astragalus schugnanicus 196
Astragalus sesamoides 196
Astragalus sewerzowii 196
Astragalus sieversianus 196
Astragalus skorniakovii 192
Astragalus sogdianus 197
Astragalus stalinskyi 197
Astragalus stenanthus 197
Astragalus stenocystis 197
Astragalus stipulosus 195
Astragalus striatellus 187
Astragalus subbarbellatus 196
Astragalus subinduratus 197
Astragalus subscaposus 197
Astragalus substipitatus 197
Astragalus subverticillatus 198
Astragalus tibetanus 198
Astragalus titovii 198
Astragalus transoxanus 198
Astragalus turbinatus 198
Astragalus turczaninovii 198
Astragalus turkestanus 199
Astragalus unifoliolatus 200
Astragalus uninodus 200
Astragalus urgutinus 200
Astragalus variegatus 200
Astragalus vicarius 201
Astragalus villosissimus 201
Astragalus vladimiri 188
Astragalus xanthomeloides 201
Asyneuma 419
Asyneuma argutum 419
Asyneuma trautvetteri 420
Atraphaxis 306
Atraphaxis compacta 306
Atraphaxis karataviensis 306
Atraphaxis pyrifolia 306
Atraphaxis seravschanica 307
Atraphaxis spinosa 307
Atraphaxis virgata 307
Atriplex 332
Atriplex aucherii 332
Atriplex dimorphostegia 332
Atriplex flabellum 332
Atriplex hastata 333
Atriplex micrantha 332
Atriplex ornata 332
Atriplex prostrata subsp. calotheca 333
Atriplex tatarica 333
Aulacospermum 482
Aulacospermum roseum 482
Aulacospermum simplex 482
Aulacospermum tenuisectum 482
Avena 113
Avena barbata 113
Avena fatua 113
Avena persica 113
Avena sterilis subsp. ludoviciana 113
Avena trichophylla 113
B
Baeothryon 97
Baeothryon pumilum 97
Balsaminaceae 346
Barbarea 276
Barbarea brachycarpa subsp.
minor 276
Barbarea minor 276
Barbarea vulgaris 276
Barkhausia kotschyana 427
Bassia 334
Bassia crassifolia 334
Bassia eriophora 334
Bassia hyssopifolia 334
Batrachium divaricatum 164
Batrachium pachycaulon 164
Batrachium rionii 162
Batrachium trichophyllum 164
Berberidaceae 143
Berberis 143
Berberis integerrima 143
Berberis nummularia 143
Berberis oblonga 144
Berula 487
Berula erecta 487
Betula 243
Betula pendula 243
Betulaceae 243
Bidens 451
Bidens tripartita 451
Biebersteinia 263
Biebersteinia multifida 263
Biebersteiniaceae 263
Bistortia elliptica 313
Blysmus 97
Blysmus compressus 97
Boissiera 125
Boissiera squarrosa 125
Bolbosaponaria 325
Bolbosaponaria sewerzowii 325
Bolboschoenus 95
Bolboschoenus maritimus 95
Bolboschoenus popovii 96
Bongardia 145
Bongardia chrysogonum 145
Boraginaceae 359
Botriochloa 104
Botriochloa ischaemum 104
Botrychium 34
Botrychium lunaria 34
Brachyactis ciliata 458
Brachypodium 125
Brachypodium sylvaticum 125
Brassica 287
Brassica campestris 287
Brassica elongata 287
Brassica rapa 287
Brassicaceae 274
Bromus 123
Bromus danthoniae 123
Bromus inermis 124
Bromus japonicus 124
Bromus lanceolatus 124
Bromus oxyodon 124
Bromus paulsenii 124
Bromus popovii 124
Bromus scoparius 124
Bromus scoparius 124
Bromus sericeus 124
Bromus sewerzowii 124
Bromus sterilis 124
Bromus tectorum 124
Bryonia 243
Bryonia cretica subsp. dioica 243
Bryonia dioica 243
509
510
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Bryonia melanocarpa 244
Buchingera axillaris 286
Bufonia 317
Bufonia oliveriana 317
Buglossoides 360
Buglossoides arvensis 360
Bunium capusii 486
Bunium chaerophylloides 486
Bunium intermedium 486
Bunium persicum 486
Bupleurum 485
Bupleurum exaltatum 485
Bupleurum falcatum subsp.
exaltatum 485
Butomaceae 49
Butomus 49
Butomus umbellatus 49
C
Calamagrostis 112
Calamagrostis anthoxanthoides 112
Calamagrostis dubia 112
Calamagrostis epigeios 112
Calamagrostis holciformis 112
Calamagrostis laguroides 112
Calamagrostis pseudophragmites 112
Callianthemum 157
Callianthemum alatavicum 157
Calligonum 307
Calligonum aphyllum 307
Calligonum caput-medusae 307
Calligonum leucocladum 308
Calligonum macrocarpum 309
Calligonum microcarpum 310
Callipeltis 354
Callipeltis cucularis 354
Calystegia 373
Calystegia sepium 373
Camelina 294
Camelina microcarpa 294
Camelina rumelica 294
Camelina sylvestris 294
Campanula 418
Campanula cashmeriana 418
Campanula fedtschenkoana 419
Campanula glomerata 419
Campanula lehmanniana 419
Campanulaceae 418
Camphorosma 334
Camphorosma monspeliaca 334
Campyloptera carnea 293
Cannabaceae 241
Cannabis 241
Cannabis sativa 241
Capparaceae 273
Capparis 273
Capparis spinosa 273
Caprifoliaceae 473
Capsella 294
Capsella bursa-pastoris 294
Caragana 181
Caragana alaica 181
Caragana turkestanica 181
Cardaria repens 290
Carduus 435
Carduus albidus 435
Carduus nutans 435
Carduus pycnocephalus subsp.
albidus 435
Carex 100
Carex diluta 100
Carex divisa 100
Carex enervis 100
Carex karoi 100
Carex melanantha 100
Carex melanostachya 101
Carex microglochin 101
Carex orbicularis 101
Carex pachystilis 101
Carex parva 101
Carex physodes 101
Carex pseudofoetida 101
Carex serotina 101
Carex songorica 101
Carex stenocarpa 102
Carex stenophylla 102
Carex subphysodes 102
Carex turkestanica 102
Carthamus 450
Carthamus oxyacanthus 450
Carthamus tinctorius 450
Carthamus turkestanicus 450
Carum 486
Carum carvi 486
Caryophyllaceae 314
Catabrosa 116
Catabrosa aquatica 116
Catabrosa capusii 116
Celastraceae 245
Celtis 241
Celtis australis subsp. caucasica 241
Centaurea 448
Centaurea belangeriana 448
Centaurea benedicta 448
Centaurea bruguierana subsp.
belangeriana 448
Centaurea cyanus 448
Centaurea depressa 448
Centaurea iberica 448
Centaurea solstitialis 448
Centaurea squarrosa 448
Centaurea virgata subsp. squarrosa 448
Centaurium 355
Centaurium erythraea subsp.
turcicum 355
Centaurium pulchellum 355
Centaurium spicatum 355
Centaurium turcicum 355
Cephalaria 477
Cephalaria syriaca 477
Cephalorrhynchus soongoricus 424
Cerastium 315
Cerastium arvense 315
Cerastium bungeanum 316
Cerastium cerastoides 316
Cerastium dentatum 316
Cerastium dichotomum 316
Cerastium dichotomum subsp.
inflatum 316
Cerastium falcatum 316
Cerastium fontanum subsp.
vulgare 316
Cerastium glomeratum 316
Cerastium holosteoides 316
Cerastium inflatum 316
Cerastium lithospermifolium 316
Cerastium pentandrum 316
Cerastium perfoliatum 316
Cerastium pusillum 316
Cerastium semidecandrum 316
Cerasus amygdaliflora 238
Cerasus erythrocarpa 235
Cerasus mahaleb 236
Cerasus verrucosa 238
Cerasus vulgaris 234
Ceratocarpus 333
Ceratocarpus arenarius 333
Ceratocarpus utriculosus 333
Ceratocephala 159
Ceratocephala falcata 159
Ceratocephala testiculata 159
Ceratoides ewersmanniana 333
Ceratoides latens 333
Ceratophyllaceae 136
Ceratophyllum 136
Ceratophyllum demersum 136
Ceterach 37
Ceterach officinarum 37
Chaerophyllum nodosum 480
Chaetolimon 300
Chaetolimon setiferum 300
Chalcanthus 289
Chalcanthus renifolius 289
Chamaenerion angustifolium 261
Chamaesphacos 406
Chamaesphacos ilicifolius 406
Chamaesyce canescens 252
Chamaesyce turcomanica 254
Chardinia 435
Chardinia orientalis 435
Cheilanthes 35
Cheilanthes persica 35
Chenopodium 330
Chenopodium album 330
Chenopodium botrys 331
Chenopodium chenopodioides 331
Chenopodium foliosum 331
Chenopodium glaucum 332
Chenopodium murale 332
Chenopodium rubrum 332
Chenopodium vulvaria 332
Chesneya 181
Chesneya ternata 181
Chesneya turkestanica 182
Chondrilla 423
Chondrilla aspera 423
INDEX OF LATIN NAMES
Chondrilla lejosperma 423
Chorispora 283
Chorispora bungeana 283
Chorispora elegans 284
Chorispora macropoda 283
Chorispora sabulosa 284
Chorispora tenella 284
Chrozophora 251
Chrozophora gracilis 252
Chrozophora hierosolymitana 251
Chrozophora obliqua 251
Chrozophora sabulosa 252
Chrozophora tinctoria 251
Chrysaspis campestris 177
Cicer 211
Cicer flexuosum 211
Cicer grande 211
Cicer macracanthum 213
Cicer paucijugum 213
Cicer pungens 213
Cicer songaricum 213
Cichorium 422
Cichorium intybus 422
Cirsium 436
Cirsium alatum 436
Cirsium arvense 436
Cirsium brevipapposum 436
Cirsium esculentum 436
Cirsium ochrolepidium 436
Cirsium polyacanthum 436
Cirsium semenovii 436
Cirsium straminispinum 436
Cirsium turkestanicum 436
Cirsium vulgare 436
Cistaceae 272
Cistanche 415
Cistanche mongolica 415
Cistanche salsa 415
Cithareloma 284
Cithareloma lehmannii 284
Clematis 158
Clematis orientalis 158
Clemensia 166
Clementsia semenovii 166
Cleomaceae 273
Cleome 273
Cleome fimbriata 273
Climacoptera 339
Climacoptera lanata 339
Climacoptera longistylosa 339
Climacoptera minkvitziae 340
Climacoptera obtusifolia 340
Climacoptera olgae 340
Climacoptera transoxana 340
Clinopodium 411
Clinopodium alpinum 411
Clinopodium debile 411
Clypeola 286
Clypeola jonthlaspi 286
Cnicus benedictus 448
Codonopsis 422
Codonopsis clematidea 422
Colchicaceae 51
Colchicum 51
Colchicum kesselringii 51
Colchicum luteum 53
Colpodium 120
Colpodium humile 120
Colpodium parviflorum 120
Colutea 179
Colutea paulsenii 179
Compositae 422
Conioselinum 490
Conioselinum tataricum 490
Conioselinum vaginatum 490
Conium 483
Conium maculatum 483
Conringia 288
Conringia clavata 288
Conringia orientalis 288
Conringia persica 288
Conringia planisiliqua 288
Consolida 150
Consolida camptocarpa 150
Consolida leptocarpa 151
Consolida rugulosa 152
Convolvulaceae 370
Convolvulus 371
Convolvulus arvensis 371
Convolvulus fruticosus 371
Convolvulus hamadae 372
Convolvulus korolkowii 372
Convolvulus lineatus 372
Convolvulus pilosellifolius 373
Convolvulus pseudocantabricus 373
Convolvulus subhirsutus 373
Conyza canadensis 457
Coriandrum 481
Coriandrum sativum 481
Corispermum 334
Corispermum lehmanninum 334
Cortusa turkestanica 347
Corydalis 141
Corydalis aitchisonii 141
Corydalis glaucescens 142
Corydalis gortschakovii 142
Corydalis ledebouriana 142
Corydalis nudicaulis 143
Corydalis schelesnowiana 143
Corydalis sewerzowii 141
Cotoneaster 218
Cotoneaster goloskokovii 218
Cotoneaster multiflorus 218
Cotoneaster nummularioides 219
Cotoneaster nummularius 219
Cotoneaster oliganthus 219
Cotoneaster songaricus 220
Cotoneaster suavis 221
Cousinia 438
Cousinia alpina 438
Cousinia ambigens 438
Cousinia aurea 436
Cousinia botschantzevii 438
Cousinia bungeana 439
Cousinia chlorantha 437
Cousinia coronata 439
Cousinia dissectifolia 439
Cousinia dshisakensis 439
Cousinia dubia 440
Cousinia eriotricha 440
Cousinia franchetii 440
Cousinia haesitabunda 437
Cousinia hamadae 440
Cousinia horridula 440
Cousinia integrifolia 440
Cousinia karatavica 437
Cousinia korolkowii 437
Cousinia lappacea 440
Cousinia microcarpa 441
Cousinia mollis 441
Cousinia olgae 441
Cousinia outichaschensis 441
Cousinia oxiana 441
Cousinia polytimetica 441
Cousinia prolifera 441
Cousinia pseudodshisakensis 442
Cousinia pygmaea 442
Cousinia radians 442
Cousinia regelii 442
Cousinia resinosa 442
Cousinia schtschurowskiana 442
Cousinia simulatrix 443
Cousinia sogdiana 443
Cousinia spiridonovii 444
Cousinia tenella 444
Cousinia umbrosa 444
Cousinia verticillaris 444
Cousinia xanthina 444
Crambe 288
Crambe cordifolia subsp.
kotschyana 288
Crambe kotschyana 288
Crassulaceae 166
Crataegus 223
Crataegus pontica 223
Crataegus songarica 224
Crataegus turkestanica 224
Crepidifolium 424
Crepidifolium serawschanicum 424
Crepis 427
Crepis kotschyana 427
Crepis multicaulis 427
Crepis pulchra 427
Crocus 67
Crocus korolkowii 67
Crucianella 351
Crucianella chlorostachys 351
Crucianella exasperata 351
Crucianella filifolia 351
Cruciata 354
Cruciata pedemontana 354
Crupina 450
Crupina oligantha 450
Crupina vulgaris 450
Crypsis 108
Crypsis schoeoides 108
511
512
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Cryptospora 282
Cryptospora falcata 282
Cryptospora inconspicua 282
Cryptospora omissa 282
Cucurbitaceae 243
Cullen 178
Cullen drupaceum 178
Cuminum 487
Cuminum setifolium 487
Cupressaceae 44
Cuscuta 370
Cuscuta approximata 370
Cuscuta brevistyla 370
Cuscuta campestris 370
Cuscuta cupulata 370
Cuscuta europaea 371
Cuscuta lehmanniana 371
Cuscuta monogyna 371
Cuscuta pedicellata 371
Cuscuta pellucida 371
Cutandia rigescens 116
Cutandia Willk 116
Cyanus 448
Cyanus depressus 448
Cyanus segetum 448
Cylindrocarpa 422
Cylindrocarpa sewerzowii 422
Cymbolaena 456
Cymbolaena griffithii 456
Cynanchum 358
Cynanchum acutum subsp.
sibiricum 358
Cynanchum sibiricum 358
Cynodon 113
Cynodon dactylon 113
Cynoglossum 370
Cynoglossum creticum 370
Cyperaceae 95
Cyperus 98
Cyperus glaber 98
Cyperus longus 98
Cyperus rotundus 98
Cystidospermum cheirolepis 252
Cystopteris 36
Cystopteris fragilis 36
D
Dactylis 117
Dactylis glomerata 117
Dactylorhiza 65
Dactylorhiza umbrosa 65
Datisca 244
Datisca cannabina 244
Datiscaceae 244
Datura 376
Datura stramonium 376
Daucus 481
Daucus carota 481
Delphinium 152
Delphinium barbatum 152
Delphinium batalinii 153
Delphinium biternatum 154
Delphinium confusum 154
Delphinium leptocarpum 151
Delphinium longipedunculatum 154
Delphinium oreophilum 155
Delphinium poltoratzkii 155
Delphinium propinquum 155
Delphinium rugulosum 152
Delphinium semibarbatum 155
Dendrostellera arenaria 272
Deschampsia 112
Deschampsia cespitosa 112
Deschampsia koelerioides 112
Descurainia 275
Descurainia sophia 275
Dianthus 327
Dianthus baldzhuanicus 327
Dianthus brevipetalus 327
Dianthus crinitus subsp. tetralepis 327
Dianthus darvazicus 327
Dianthus helenae 328
Dianthus subscabridus 329
Dianthus tetralepis 327
Diarthron 272
Diarthron arenaria 272
Diarthron vesiculosum 272
Dictamnus 269
Dictamnus angustifolius 269
Didymophysa 292
Didymophysa fedtschenkoana 292
Digitaria 104
Digitaria ischaemum 104
Digitaria sanguinalis 104
Dipsacus 476
Dipsacus dipsacoides 476
Dipsacus laciniatus 476
Diptychocarpus 283
Diptychocarpus strictus 283
Dodartia 413
Dodartia orientalis 413
Draba 286
Draba lanceolata 286
Draba melanopus 286
Draba nemorosa 286
Draba nuda 286
Draba stenocarpa 287
Drabopsis nuda 286
Dracocephalum 388
Dracocephalum diversifolium 388
Dracocephalum imberbe 388
Dracocephalum nodulosum 389
Dracocephalum nuratavicum 389
Dracocephalum scrobiculatum 389
Drepanocaryum 386
Drepanocaryum sewerzowii 386
Dryopteris 38
Dryopteris filix-mas 38
E
Echinochloa 104
Echinochloa crus-galli 104
Echinophora 480
Echinophora sibthorpiana 480
Echinophora tenuifolia subsp.
sibthorpiana 480
Echinops 434
Echinops knorringianus 434
Echinops leucographus 434
Echinops maracandicus 434
Echinops nanus 434
Echinops nuratavicus 434
Echium 363
Echium biebersteinii 363
Echium vulgare 363
Eclipta 450
Eclipta prostrata 450
Elaeagnaceae 238
Elaeagnus 239
Elaeagnus angustifolia 239
Elaeosticta 485
Elaeosticta allioides 485
Elaeosticta hirtula 485
Elaeosticta polycarpa 485
Elaeosticta vvedenskyi 485
Eleocharis 97
Eleocharis argyrolepis 97
Eleocharis mitracarpa 98
Eleocharis quinqueflora 98
Eleocharis uniglumis 98
Elwendia 486
Elwendia capusii 486
Elwendia chaerophylloides 486
Elwendia intermedia 486
Elwendia persica 486
Elymus 131
Elymus bungeanus 131
Elymus dentatus 131
Elymus drobovii 132
Elymus hispidus 132
Elymus lolioides 132
Elymus repens 132
Elymus uralensis 132
Eminium 49
Eminium lehmanii 49
Enneapogon 114
Enneapogon persicus 114
Ephedra 40
Ephedra botschantzevii 40
Ephedra equisetina 40
Ephedra fedtschenkoae 40
Ephedra foliata 41
Ephedra intermedia 42
Ephedra regeliana 43
Ephedra strobilacea 44
Ephedraceae 40
Epilasia 431
Epilasia acrolasia 431
Epilasia hemilasia 432
Epilasia mirabilis 432
Epilobium 261
Epilobium angustifolium 261
Epilobium hirsutum 261
Epilobium komarovii 262
Epilobium minutiflorum 262
Epilobium nervosum 263
INDEX OF LATIN NAMES
Epilobium palustre 263
Epilobium roseum subsp.
subsessile 263
Epilobium tetragonum 263
Epilobium tianschanicum 263
Epilobium turkestanicum 263
Epilobium velutinum 261
Epipactis 65
Epipactis latifolia 65
Equisetaceae 34
Equisetum 34
Equisetum arvense 34
Equisetum ramosissimum 34
Eragrostis 115
Eragrostis minor 115
Eragrostis pilosa 115
Eranthis 156
Eranthis longistipitata 156
Eremodaucus 483
Eremodaucus lehmannii 483
Eremogone 318
Eremogone griffithii 318
Eremolimon 303
Eremolimon sogdianum 303
Eremopyrum 126
Eremopyrum bonaepartis 126
Eremopyrum distans 127
Eremopyrum orientale 128
Eremopyrum triticeum 128
Eremurus 70
Eremurus anisopterus 70
Eremurus chloranthus 70
Eremurus fuscus 70
Eremurus inderiensis 70
Eremurus kaufmannii 70
Eremurus lactiflorus 71
Eremurus nuratavicus 71
Eremurus olgae 72
Eremurus regelii 73
Eremurus robustus 74
Eremurus sogdianus 74
Eremurus tianschanicus 75
Eremurus turkestanicus 75
Erianthus 103
Erianthus ravennae 103
Ericaceae 351
Erigeron 457
Erigeron acris 457
Erigeron bellidiformis 457
Erigeron canadensis 457
Erigeron khorossanicus 457
Erigeron pallidus 458
Erigeron petiolaris 458
Erigeron popovii 458
Erigeron pseudoneglectus 458
Erigeron pseudoseravschanicus 458
Erigeron seravschanicus 458
Erigeron sogdianus 458
Erigeron umbrosus 458
Eriocaulaceae 93
Eriocaulon 93
Eriocaulon sieboldianum 93
Erodium 260
Erodium ciconium 260
Erodium cicutarium 260
Erodium hoefftianum 260
Erodium oxyrrhynchum 260
Erophila 287
Erophila minima 287
Erophila verna 287
Eruca 287
Eruca sativa 287
Eruca vesicaria 287
Eryngium 479
Eryngium caeruleum 479
Eryngium macrocalyx 479
Eryngium octophyllum 479
Erysimum 275
Erysimum cyaneum 275
Erysimum marschallianum 275
Erysimum nuratense 276
Erysimum violascens 276
Euclidium 284
Euclidium syriacum 284
Eudicots 135
Euphorbia 252
Euphorbia chamaesyce 252
Euphorbia cheirolepis 252
Euphorbia densa 252
Euphorbia esula 253
Euphorbia falcata 253
Euphorbia franchetii 253
Euphorbia glomerulans 253
Euphorbia granulata 254
Euphorbia helioscopia 254
Euphorbia humilis 254
Euphorbia inderiensis 254
Euphorbia jaxartica 253
Euphorbia kanaorica 255
Euphorbia rapulum 255
Euphorbia rosularis 255
Euphorbia sarawschanica 255
Euphorbia sororia 255
Euphorbia szowitsii 255
Euphorbia turczaninowii 256
Euphorbiaceae 251
Euphrasia 416
Euphrasia pectinata 416
Euphrasia regelii 416
Eurotia ceratoides 333
Eurotia ewersmanniana 333
Evax filaginoides 456
F
Fabaceae 172
Falcaria 486
Falcaria vulgaris 486
Fallopia 314
Fallopia convolvulus 314
Ferns 33
Ferula 491
Ferula angreni 491
Ferula bucharica 491
Ferula diversivittata 492
Ferula dshizakensis 492
Ferula fedtschenkoana 492
Ferula ferganensis 492
Ferula foetida 492
Ferula helenae 493
Ferula kokanica 493
Ferula kuhistanica 494
Ferula lehmannii 495
Ferula mollis 495
Ferula moschata 495
Ferula ovczinnikovii 495
Ferula ovina 495
Ferula penninervis 496
Ferula samarkandica 496
Ferula schtschurowskiana 497
Ferula sumbul 495
Ferula varia 497
Fessia 92
Fessia puschkinioides 92
Festuca 121
Festuca alaica 121
Festuca amblyodes 121
Festuca arundinacea 121
Festuca griffithiana - 121
Festuca olgae 121
Festuca pratensis 121
Festuca rubra 121
Festuca rupicola 121
Festuca valesiaca 121
Filago 456
Filago arvensis 456
Filago filaginoides 456
Filago germanica 456
Filago pyramidata 456
Filago vulgaris 456
Fimbristylis 98
Fimbristylis dichotoma 98
Frankenia 294
Frankenia bucharica 294
Frankenia bucharica subsp.
vvedenskyi 295
Frankenia hirsuta 295
Frankenia pulverulenta 295
Frankenia vvedenskyi 295
Frankeniaceae 294
Fraxinus 377
Fraxinus angustifolia subsp.
syriaca 377
Fraxinus sogdiana 377
Fraxinus syriaca 377
Fritillaria 64
Fritillaria olgae 64
Fumaria 143
Fumaria vaillantii 143
G
Gagea 54
Gagea afghanica 54
Gagea bergii 54
Gagea calyptrifolia 54
Gagea capillifolia 54
Gagea capusii 54
513
514
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Gagea chomutovae 54
Gagea divaricata 55
Gagea dschungarica 55
Gagea filiformis 55
Gagea gageoides 55
Gagea graminifolia 56
Gagea hissarica 57
Gagea kamelinii 57
Gagea liotardii 58
Gagea nabievii 58
Gagea olgae 58
Gagea ova 57
Gagea reinhardii 59
Gagea reticulata 59
Gagea stipitata 59
Gagea stolonifera 60
Gagea subtilis 60
Gagea taschkentica 60
Gagea tenera 61
Gagea turkestanica 61
Gagea vegeta 61
Gagea vvedenskyi 61
Galagania 485
Galagania fragrantissima 485
Galagania tenuisecta 485
Galatella 456
Galatella coriacea 456
Galium 352
Galium aparine 352
Galium ceratopodum 352
Galium decaisnei 353
Galium humifusum 352
Galium ibicinum 353
Galium karakulense 352
Galium pamiroalaicum 352
Galium pseudorivale 353
Galium ruthenicum 353
Galium setaceum 353
Galium songaricum 353
Galium spurium 353
Galium spurium subsp. ibicinum 353
Galium tenuissimum 353
Galium tianschanicum 353
Galium tricornutum 353
Galium turkestanicum 353
Galium verticellatum 353
Galium verum 353
Gamanthus 340
Gamanthus gamocarpus 340
Garhadiolus 422
Garhadiolus angulosus 422
Garhadiolus hedypnois 422
Garhadiolus papposus 422
Gastrolychnis 325
Gastrolychnis apetala 324
Gastrolychnis longicarpophora 325
Gegea pseudoreticulata 59
Gentiana 356
Gentiana barbata 357
Gentiana leucomelaena 356
Gentiana olivieri 355 356
Gentiana prostrata 357
Gentiana riparia 357
Gentiana sibirica 357
Gentiana turkestanorum 357
Gentianaceae 355
Gentianella 357
Gentianella sibirica 357
Gentianella turkestanorum 357
Gentianopsis 357
Gentianopsis barbata 357
Geraniaceae 257
Geranium 257
Geranium collinum 257
Geranium divaricatum 257
Geranium kotschyi subsp. charlesii 258
Geranium linearilobum 258
Geranium pusillum 258
Geranium regelii 257
Geranium robertianum 258
Geranium rotundifolium 258
Geranium saxatile 259
Geum 228
Geum heterocarpum 228
Girgensohnia 340
Girgensohnia diptera 340
Girgensohnia oppositiflora 340
Gladiolus 69
Gladiolus italicus 69
Glaucium 139
Glaucium elegans 139
Glaucium fimbrilligerum 140
Glaucium squamigerum 140
Glaux maritima 349
Gleditsia 172
Gleditsia caspia 172
Glyceria 120
Glyceria plicata 120
Glycyrrhiza 206
Glycyrrhiza aspera 206
Glycyrrhiza glabra 206
Glycyrrhiza triphylla 206
Gnaphalium 455
Gnaphalium luteoalbum 455
Gnaphalium supinum 455
Goldbachia 277
Goldbachia laevigata 277
Goldbachia sabulosa 277
Goldbachia torulosa 278
Goldbachia verrucosa 278
Gramineae 103
Grossulariaceae 165
GYMNOSPERMAE 39
Gymnospermium 144
Gymnospermium albertii 144
GYMNOSPERMS 39
Gypsophila 325
Gypsophila cephalotes 325
Gypsophila herniarioides 325
Gypsophila paniculata 325
H
Hackelia 368
Hackelia uncinata 368
Halerpestes 159
Halerpestes sarmentosa 159
Halimocnemis 345
Halimocnemis mollissima 345
Halimocnemis villosa 345
Halimodendron 180
Halimodendron halodendron 180
Halocharis 344
Halocharis hispida 344
Halocnemum 336
Halocnemum strobilaceum 336
Halogeton 345
Halogeton glomeratus 345
Halopeplis 336
Halopeplis pygmaea 336
Haloragaceae 169
Halostachys 336
Halostachys belangeriana 336
Halothamnus 340
Halothamnus glaucus 340
Halothamnus iliensis 340
Halothamnus subaphyllus 340
Haloxylon 342
Haloxylon ammodendron 342
Haloxylon aphyllum 342
Haloxylon persicum 343
Handelia 462
Handelia trichophylla 462
Haplophyllum 266
Haplophyllum acutifolium 266
Haplophyllum bungei 267
Haplophyllum lasianthum 267
Haplophyllum latifolium 267
Haplophyllum pedicellatum 267
Haplophyllum perforatum 266
Haplophyllum ramosissimum 269
Haplophyllum robustum 269
Haplophyllum versicolor 267
Hedysarum 206
Hedysarum flavescens 206
Hedysarum jaxarticum 206
Hedysarum mogianicum 206
Hedysarum montanum 206
Hedysarum nuratense 207
Hedysarum plumosum 208
Helianthemum 272
Helianthemum songaricum 272
Helichrysum 454
Helichrysum maracandicum 454
Helichrysum mussae 455
Helichrysum nuratavicum 455
Helichrysum tianschanicum 455
Helictotrichon 113
Helictotrichon asiaticum 113
Helictotrichon desertorum 113
Heliotropium 359
Heliotropium arguzioides 359
Heliotropium bogdanii 359
Heliotropium dasycarpum 359
Heliotropium ellipticum 359
Heliotropium lasiocarpum 359
Heliotropium olgae 360
INDEX OF LATIN NAMES
Henrardia 126
Henrardia persica 126
Heracleum 497
Heracleum lehmannianum 497
Heracleum olgae 497
Herniaria 319
Herniaria glabra 319
Herniaria hirsuta 319
Hesperis 278
Hesperis sibirica 278
Heteracia 426
Heteracia epapposa 426
Heteracia szovitzii 426
Heteranthelium 129
Heteranthelium piliferum 129
Heterocaryum 367
Heterocaryum laevigatum 367
Heterocaryum macrocarpum 367
Heterocaryum subsessile 367
Heterocaryum szovitsianum 367
Heteroderis 423
Heteroderis pusilla 423
Hibiscus 272
Hibiscus trionum 272
Hieracium 428
Hieracium echioides 427
Hieracium procerum 428
Hieracium raddeanum subsp.
regelianum 428
Hieracium regelianum 428
Hieracium robustum 428
Hieracium virosum 428
Hippophae 238
Hippophae rhamnoides 238
Hippuris 380
Hippuris vulgaris 380
Holoschoenus 97
Holoschoenus vulgaris 97
Holosteum 316
Holosteum glutinosum 317
Holosteum umbellatum 316
Holosteum umbellatum subsp.
glutinosum 317
Hordeum 130
Hordeum bogdanii 130
Hordeum brevisubulatum 130
Hordeum bulbosum 130
Hordeum geniculatum 131
Hordeum leporinum 131
Hordeum marinum subsp.
gussoneanum 131
Hordeum murinum subsp.
leporinum 131
Hordeum spontaneum 131
Hornungia 291
Hornungia procumbens 291
Hulthemia 233
Hulthemia persica 233
Hyalea 449
Hyalea pulchella 449
Hyalolaena 485
Hyalolaena depauperata 485
Hyalolaena jaxartica 485
Hydrocharitaceae 49
Hymenolobus procumbens 291
Hyoscyamus 375
Hyoscyamus niger 375
Hyoscyamus pusillus 375
Hypecoum 140
Hypecoum parviflorum 140
Hypecoum pendulum 140
Hypecoum trilobum 141
Hypericaceae 245
Hypericum 245
Hypericum elongatum 245
Hypericum perforatum 246
Hypericum scabrum 246
Hypogomphia 391
Hypogomphia turkestana 391
Hypogomphia turkestanica 391
Hyssopus 411
Hyssopus seravschanicus 411
I
Iberidella 292
Iberidella trinervia 292
Impatiens 346
Impatiens parviflora 346
Imperata 103
Imperata cylindrica 103
Inula 453
Inula britannica 453
Inula caspica 453
Inula glauca 453
Inula helenium 453
Inula macrophylla 453
Inula orientalis 453
Inula rhizocephala 453
Iridaceae 67
Iris 67
Iris linifoliiformis 67
Iris loczyi 67
Iris longiscapa 67
Iris maracandica 68
Iris narbutii 68
Iris parvula 69
Iris songarica 69
Iris tadshikorum 69
Isatis 277
Isatis brevipes 277
Isatis emarginata 277
Isatis minima 277
Isatis multicaulis 277
Isatis tinctoria 277
Isatis violascens 277
Isopyrum 146
Isopyrum anemonoides 146
Ixioliriaceae 66
Ixiolirion 66
Ixiolirion tataricum 66
J
Juglandaceae 242
Juglans 242
Juglans regia 242
Juncaceae 93
Juncaginaceae 50
Juncus 93
Juncus articulatus 93
Juncus bufonius 93
Juncus compressus 93
Juncus gerardii 93
Juncus heptopotamicus 93
Juncus hybridus 94
Juncus inflexus 94
Juncus jaxarticus 94
Juncus macrantherus 94
Juncus persicus subsp. libanoticus 94
Juncus rechingeri 94
Juncus schischkinii 94
Juncus sphaerocarpus 94
Juncus triglumis 94
Juncus turkestanicus 94
Juncus vvedenskyi 94
Juniperus 44
Juniperus polycarpos var.
seravschanica 46
Juniperus pseudosabina 44
Juniperus semiglobosa 45
Jurinea 444
Jurinea abramowii 444
Jurinea androssovii 444
Jurinea ferganica 444
Jurinea helichrysifolia 444
Jurinea kokanica 445
Jurinea komarovii 445
Jurinea lasiopoda 445
Jurinea maxima 446
Jurinea olgae 446
Jurinea suffruticosa 447
Jurinea trautvetteriana 447
Jurinea zakirovii 447
K
Kalidium 336
Kalidium caspicum 336
Kalimeris 457
Kalimeris altaica 457
Karelinia 453
Karelinia caspia 453
Kickxia 377
Kickxia elatine 377
Kirilovia eriantha 334
Kobresia 99
Kobresia pamiroalaica 99
Kobresia persica 99
Kobresia stenocarpa 99
Kochia 334
Kochia iranica 334
Kochia prostrata 334
Kochia scoparia 334
Koeleria 115
Koeleria macrantha 115
Koelpinia 433
Koelpinia linearis 433
Koelpinia macrantha 434
515
516
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Koelpinia tenuissima 434
Koelpinia turanica 434
Korshinskya 482
Korshinskya olgae 482
Kovalevskiella 424
Kovalevskiella zeravschanica 424
Kozlovia 480
Kozlovia paleacea 480
Krascheninnikovia 333
Krascheninnikovia ceratoides 333
Krascheninnikovia ewersmanniana 333
Kudrjaschevia 389
Kudrjaschevia jacubi 389
Kuhitangia 325
Kuhitangia knorringiana 325
Kuschakewiczia 370
Kuschakewiczia turkestanica 370
L
Labiatae 383
Lachnoloma 284
Lachnoloma lehmanii 284
Lachnophyllum 458
Lachnophyllum gossypinum 458
Lactuca 423
Lactuca altaica 424
Lactuca crambifolia 423
Lactuca dissecta 423
Lactuca glauciifolia 423
Lactuca orientalis 423
Lactuca serriola 424
Lactuca soongorica 424
Lactuca spinidens 424
Lactuca tatarica 424
Lactuca undulata 424
Ladyginia bucharica 491
Lagochilus 403
Lagochilus inebrians 403
Lagochilus olgae 404
Lagochilus proskorjakovii 404
Lagochilus seravschanicus 405
Lagotis 379
Lagotis korolkowii 379
Lallemantia 389
Lallemantia royleana 389
Lamiaceae 383
Lamium 401
Lamium album 401
Lamium amplexicaule 401
Laphangium 455
Laphangium luteoalbum 455
Lappula 365
Lappula brachycentra 365
Lappula consanguinea 366
Lappula marginata 366
Lappula microcarpa 365
Lappula myosotis 365
Lappula nuratavica 365
Lappula occultata 366
Lappula patula 366
Lappula sarawschanica 366
Lappula semialata 366
Lappula semiglabra 366
Lappula sessiliflora 366
Lappula sinaica 366
Lappula spinocarpos 366
Lappula squarrosa 366
Lathyrus 216
Lathyrus aphaca 216
Lathyrus asiaticus 216
Lathyrus cicera 216
Lathyrus inconspicuus 216
Lathyrus pratensis 216
Lathyrus sativus 216
Lathyrus tuberosus 217
Launaea 423
Launaea korovinii 423
Launaea procumbens 423
Leguminosae 172
Leiospora 281
Leiospora subscapigera 281
Lemna 48
Lemna gibba 48
Lemna minor 48
Lens 216
Lens culinaris subsp. orientalis 216
Lentibulariaceae 382
Leontice 144
Leontice eversmannii 144
Leontopodium 455
Leontopodium fedtschenkoanum 455
Leonurus 402
Leonurus glaucescens 402
Leonurus turkestanicus 402
Lepidium 289
Lepidium botschantsevianum 289
Lepidium draba subsp. chalepense 290
Lepidium ferganense 290
Lepidium lacerum 290
Lepidium latifolium 290
Lepidium lipskyi 290
Lepidium olgae 290
Lepidium orientale 291
Lepidium perfoliatum 291
Lepidolopha 472
Lepidolopha komarowii 472
Lepidolopha nuratavica 472
Lepidolopsis 464
Lepidolopsis turkestanica 464
Leptaleum 283
Leptaleum filifolium 283
Leptorhabdos 416
Leptorhabdos parviflora 416
Lepyrodiclis 317
Lepyrodiclis holosteoides 317
Lepyrodiclis stellarioides 317
Leymus 132
Leymus alaicus 132
Leymus angustus 132
Leymus multicaulis 133
Libanotis 489
Libanotis schrenkiana 489
Ligularia 452
Ligularia alpigena 452
Ligularia heterophylla 452
Ligularia thomsonii 452
Liliaceae 54
Liliopsida 47
Limonium 301
Limonium otolepis 301
Limonium reniforme 302
Limonium suffruticosum 302
Linaceae 256
Linaria 381
Linaria popovii 381
Linaria sessilis 381
Lindelofia 369
Lindelofia macrostyla 369
Lindelofia olgae 369
Lindelofia stylosa 369
Linum 256
Linum corymbulosum 256
Linum humile 257
Linum macrorhizum 256
Linum olgae 256
Linum usitatissimum 256
Lipskya 482
Lipskya insignis 482
Lipskyella 444
Lipskyella annua 444
Lithospermum 360
Lithospermum officinale 360
Litwinowia 284
Litwinowia tenuissima 284
Lloydia 64
Lloydia serotina 64
Loliolum 122
Loliolum subulatum 122
Lolium 126
Lolium perenne 126
Lolium persicum 126
Lolium temulentum 126
Lomatogonium 357
Lomatogonium carinthiacum 357
Londesia eriantha 334
Lonicera 473
Lonicera altmannii 474
Lonicera bracteolaris 473
Lonicera humilis 474
Lonicera korolkowii 474
Lonicera microphylla 475
Lonicera nummulariifolia 475
Lonicera olgae 475
Lonicera paradoxa 476
Lonicera semenovii 476
Lonicera simulatrix 476
Lonicera zeravshanica 476
Lophanthus 387
Lophanthus ouroumitanensis 387
Lophanthus schtschurowskianus 387
Lophanthus subnivalis 387
Lotus 178
Lotus krylovii 178
Lotus sergievskiae 178
Luzula 94
Luzula pallescens 94
INDEX OF LATIN NAMES
Luzula spicata 94
Lycium 374
Lycium dasystemum 374
Lycium ruthenicum 374
Lycopus 412
Lycopus europaeus 412
Lysimachia 349
Lysimachia dubia 349
Lysimachia maritima 349
Lythraceae 261
Lythrum 261
Lythrum hyssopifolia 261
Lythrum nanum 261
M
Macrotomia euchroma 362
Magnoliopsida 135
Malus 222
Malus sieversii 222
Malva 270
Malva neglecta 270
Malvaceae 270
Marrubium 386
Marrubium anisodon 386
Marsilea 35
Marsilea quadrifolia 35
Marsileaceae 35
Matthiola 282
Matthiola chorassanica 282
Matthiola integrifolia 282
Matthiola obovata 282
Mattiastrum 368
Mattiastrum himalayense 368
Mausolea 464
Mausolea eriocarpa 464
Mazaceae 413
Mediasia 490
Mediasia macrophylla 490
Medicago 176
Medicago denticulata 176
Medicago lupulina 176
Medicago medicaginoides 176
Medicago minima 176
Medicago monantha 176
Medicago orbicularis 176
Medicago orthoceras 176
Medicago polymorpha 176
Medicago rigidula 177
Medicago sativa 177
Megacarpaea 291
Megacarpaea gigantea 291
Megacarpaea orbiculata 291
Melandrium album 322
Melandrium ruinarium 324
Melandrium turkestanicum 324
Melica 116
Melica altissima 116
Melica hohenackeri 116
Melica inaequiglumis 116
Melilotus 177
Melilotus albus 177
Melilotus officinalis 177
Melissitus 176
Melissitus adscendens 174
Melissitus aristatus 174
Melissitus gontscharovii 175
Melissitus pamiricus 176
Melissitus popovii 176
Meniocus linifolius 286
Mentha 412
Mentha longifolia subsp. asiatica 412
Mentha pamiroalaica 413
Merendera 53
Merendera robusta 53
Meristotropis triphylla 206
Mesostemma 315
Mesostemma karatavica 315
Microcephala 473
Microcephala lamellata 473
Microcephala turcomanica 473
Microthlaspi perfoliatum 292
Milium 108
Milium vernale 108
Minuartia 317
Minuartia hamata 317
Minuartia kryloviana 317
Minuartia meyeri 317
Monocots 47
Moraceae 242
Morus 242
Morus alba 242
Morus nigra 242
Myosotis 365
Myosotis alpestris 365
Myosotis caespitosa 365
Myosotis laxa subsp. caespitosa 365
Myosotis micrantha 365
Myosotis refracta 365
Myosotis stricta 365
Myricaria 297
Myricaria bracteata 297
Myricaria germanica 297
Myriophyllum 169
Myriophyllum spicatum 169
N
Najas 50
Najas marina 50
Najas minor 50
Nanophyton 344
Nanophyton erinaceum 344
Nanophyton saxatile 344
Nardurus krausei 123
Nasturtium 276
Nasturtium fontanum 276
Nasturtium officinale 276
Neotorularia 274
Neotorularia torulosa 274
Nepeta 387
Nepeta alatavica 387
Nepeta bracteata 387
Nepeta cataria 388
Nepeta kokanica 388
Nepeta mariae 388
Nepeta maussarifii 388
Nepeta micrantha 388
Nepeta nuda 388
Nepeta olgae 388
Nepeta pannonica 388
Nepeta podostachys 388
Nepeta pungens 388
Nepeta saturejoides 388
Nepeta ucranica 388
Neslia 294
Neslia apiculata 294
Neslia paniculata subsp. thracica 294
Neuroloma fruticulosum 279
Neuroloma nuratense 279
Neurotropis 293
Neurotropis platycarpa 293
Nicandra 376
Nicandra physaloides 376
Nigella 156
Nigella integrifolia 156
Nitrariaceae 264
Nonea 364
Nonea caspica 364
Nonea melanocarpa 365
Nonea pulla 365
O
Octoceras 284
Octoceras lehmannianum 284
Oedibasis 486
Oedibasis apiculata 486
Oedibasis tamerlanii 486
Oleaceae 377
Olimarabidopsis 275
Olimarabidopsis pumila 275
Olimarabidopsis umbrosa 275
Onagraceae 261
Onobrychis 208
Onobrychis chorassanica 208
Onobrychis echidna 209
Onobrychis gontscharovii 210
Onobrychis grandis 210
Onobrychis micrantha 210
Onobrychis pulchella 210
Onobrychis saravschanica 211
Ononis 174
Ononis spinosa subsp. antiquorum 174
Onopordum 435
Onopordum acantium 435
Onopordum leptolepis 436
Onosma 362
Onosma dichroanta 362
Onosma maracandica 362
Ophioglossaceae 34
Orchidaceae 65
Orchis 65
Orchis pseudolaxiflora 65
Origanum 411
Origanum tyttanthum 411
Origanum vulgare subsp. gracile 411
Orlaya 481
Orlaya grandiflora 481
517
518
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Orobanchaceae 414
Orobanche 414
Orobanche aegyptiaca 414
Orobanche amoena 414
Orobanche cernua 414
Orobanche coelestis 414
Orobanche elatior 414
Orobanche gigantea 414
Orobanche hansii 414
Orobanche orientalis 414
Orobanche pallens 414
Orobanche sulphurea 415
Orthurus 228
Orthurus heterocarpus 228
Orthurus kokanicus 228
Oxyria 303
Oxyria digyna 303
Oxytropis 202
Oxytropis aspera 202
Oxytropis capusii 202
Oxytropis chesneyoides 202
Oxytropis humifusa 202
Oxytropis immersa 202
Oxytropis integripetala 202
Oxytropis kamelinii 202
Oxytropis lehmanni 202
Oxytropis leucocyanea 202
Oxytropis lipskyi 203
Oxytropis macrocarpa 203
Oxytropis michelsonii 204
Oxytropis microsphaera 204
Oxytropis platonychia 204
Oxytropis pseudorosea 204
Oxytropis riparia 205
Oxytropis rosea 205
Oxytropis savellanica 205
Oxytropis seravschanica 205
Oxytropis tachtensis 205
Oxytropis trichocalycina 205
P
Pachypterygium brevipes 277
Pachypterygium densiflorum 277
Paeonia 164
Paeonia hybrida 164
Paeonia tenuifolia 164
Paeoniaceae 164
Papaver 136
Papaver croceum 137
Papaver dubium 136
Papaver litwinowii 136
Papaver nudicaule 137
Papaver pavoninum 138
Papaveraceae 136
Paracaryum glochidiatum 368
Paracaryum himalayense 368
Paramicrorhynchus procumbens 423
Paraquilegia 147
Paraquilegia caespitosa 147
Parentucellia 416
Parentucellia flaviflora 416
Parietaria 242
Parietaria debilis 242
Parietaria lusitanica subsp. serbica
Parietaria micrantha 242
Parietaria serbica 242
Parnassia 245
Parnassia laxmannii 245
Parrya 279
Parrya fruticulosa 279
Parrya hispida 279
Parrya khorasanica 279
Parrya nuratensis 279
Parrya olgae 280
Parrya sarawschanica 281
Pedicularis 417
Pedicularis dolichorrhiza 417
Pedicularis krylovii 417
Pedicularis olgae 417
Pedicularis peduncularis 418
Pedicularis popovii 418
Pedicularis rhinanthoides 418
Pedicularis sarawschanica 418
Pedicularis waldheimii 418
Peganum 264
Peganum harmala 264
Pentanema 454
Pentanema albertoregelia 454
Pentanema divaricatum 454
Pentaphylloides 227
Pentaphylloides parvifolia 227
Perovskia 409
Perovskia angustifolia 409
Perovskia botschantzevii 409
Perovskia kudrjaschevii 409
Perovskia scrophulariifolia 409
Persicaria 313
Persicaria amphibia 313
Persicaria bistorta 313
Persicaria hydropiper 313
Persicaria lapathifolia 314
Persicaria maculata 314
Persicaria minor 314
Petrorhagia 326
Petrorhagia alpina 326
Phaecasium pulchrum 427
Phalaroides 105
Phalaroides arundinacea 105
Phelipanche aegyptiaca 414
Phelipanche coelestis 414
Phelipanche orientalis 414
Phelipanche pallens 414
Phleum 109
Phleum alpinum 109
Phleum exaratum 110
Phleum paniculatum 110
Phleum phleoides 111
Phleum pratense 111
Phlomis 398
Phlomis linearifolia 398
Phlomis nubilans 399
Phlomis olgae 400
Phlomis salicifolia 400
Phlomis thapsoides 400
242
Phlomoides 392
Phlomoides ambigua 392
Phlomoides anisochila 393
Phlomoides canescens 394
Phlomoides dszumrutensis 397
Phlomoides eriocalyx 395
Phlomoides kaufmanniana 395
Phlomoides labiosa 396
Phlomoides napuligera 396
Phlomoides oreophila 397
Phlomoides sogdiana 397
Phlomoides speciosa 397
Phlomoides uniflora 397
Phragmites 114
Phragmites australis 114
Picnomon 435
Picnomon acarna 435
Picris 422
Picris nuristanica 422
Pilosella 427
Pilosella echioides 427
Pilosella procera 428
Pimpinella 487
Pimpinella peregrina 487
Piptatherum 107
Piptatherum alpestre 107
Piptatherum ferganense 108
Piptatherum holciforme 108
Piptatherum laterale 108
Piptatherum latifolium 108
Piptatherum microcarpum 108
Piptatherum pamiralaicum 108
Piptatherum platyanthum 108
Piptatherum sogdianum 108
Piptatherum songaricum 108
Piptatherum vicarium 108
Pistacia 265
Pistacia vera 265
Plantaginaceae 377
Plantago 379
Plantago gentianoides 379
Plantago lagocephala 379
Plantago lanceolata 379
Plantago major 379
Platanaceae 164
Platanus 164
Platanus orientalis 164
Platycladus 44
Platycladus orientalis 44
Pleioneura 329
Pleioneura griffithiana 329
Plumbaginaceae 297
Poa 117
Poa alpina 117
Poa angustifolia 118
Poa annua 118
Poa bactriana 118
Poa bucharica 118
Poa bulbosa
98 118
Poa diaphora 120
Poa fragilis 120
Poa litvinoviana 120
INDEX OF LATIN NAMES
Poa nemoralis 120
Poa palustris 120
Poa pratensis 120
Poa relaxa 120
Poa supina 120
Poa trivialis 120
Poaceae 103
Polycnemum 329
Polycnemum arvense 329
Polycnemum perenne 330
Polygala 217
Polygala hybrida 217
Polygalaceae 217
Polygonaceae 303
Polygonatum 90
Polygonatum sewerzowii 90
Polygonum 311
Polygonum acetosum 311
Polygonum amphibium 313
Polygonum argyrocoleon 311
Polygonum aviculare 311
Polygonum biaristatum 311
Polygonum coriarium 311
Polygonum fibrilliferum 311
Polygonum hissaricum 312
Polygonum inflexum 312
Polygonum molliiforme 312
Polygonum nitens 313
Polygonum paronychioides 312
Polygonum patulum 312
Polygonum polychnemoides 312
Polygonum pulvinatum 312
Polygonum rottboellioides 312
Polygonum subaphyllum 312
Polygonum vvedenskyi 312
Polypodiaceae 38
Polypodiophyta 33
Polypogon 111
Polypogon fugax 111
Polypogon monspleniensis 111
Polypogon semiverticillatus 111
Polypogon viridis 111
Populus 248
Populus afghanica 248
Populus alba 249
Populus euphratica 250
Populus nigra 250
Populus pruinosa 250
Populus talassica 250
Portulaca 345
Portulaca oleracea 345
Portulacaceae 345
Potamogeton 50
Potamogeton crispus 50
Potamogeton filiformis 50
Potamogeton lucens 50
Potamogeton natans 50
Potamogeton pectinatus 51
Potamogeton perfoliatus 51
Potamogeton pusillus 51
Potamogetonaceae 50
Potentilla 225
Potentilla asiae-mediae 225
Potentilla asiatica 225
Potentilla desertorum 225
Potentilla flabellata 225
Potentilla gelida 225
Potentilla hololeuca 226
Potentilla impolita 226
Potentilla inclinata 226
Potentilla orientalis 226
Potentilla pamiroalaica 226
Potentilla pedata 226
Potentilla reptans 227
Potentilla soongorica 227
Potentilla supina 227
Potentilla vvedenskyi 227
Poterium lasiocarpum 229
Poterium polygamum 229
Prangos 483
Prangos didyma 483
Prangos fedtschenkoi 483
Prangos ornata 483
Prangos pabularia 484
Primula 347
Primula algida 347
Primula fedtschenkoi 347
Primula iljinskii 347
Primula lactiflora 349
Primula matthioli 347
Primula olgae 349
Primula pamirica 349
Primula turkeviczii 349
Primulaceae 347
Prunella 391
Prunella vulgaris 391
Prunus 233
Prunus armeniaca 233
Prunus bucharica 233
Prunus cerasus 234
Prunus divaricata 234
Prunus domestica 234
Prunus dulcis 235
Prunus erythrocarpa 235
Prunus mahaleb 236
Prunus spinosissima 237
Prunus verrucosa 238
Pseudoclausia hispida 279
Pseudoclausia olgae 280
Pseudoclausia sarawschanica 281
Pseudoclausia turkestanica 279
Pseudohandelia 462
Pseudohandelia umbellifera 462
Pseudolinosyris 456
Pseudolinosyris grimmii 456
Pseudosedum 168
Pseudosedum bucharicum 168
Pseudosedum campanuliflorum 168
Pseudosedum lievenii 169
Pseudosedum longidentatum 169
Psilurus 126
Psilurus incurvus 126
Psoralea drupacea 178
Psychrogeton 458
Psychrogeton amorphoglossus 458
Psychrogeton aucherii 457
Psychrogeton leucophyllus 458
Psychrogeton pseuderigeron 458
Psylliostachys 303
Psylliostachys leptostachya 303
Psylliostachys suworowii 303
Psylliostachys × myosuroides 303
Pteridaceae 35
Puccinelia 121
Puccinelia distans 121
Puccinellia poecilantha 121
Pulicaria 454
Pulicaria dysenterica subsp.
uliginosa 454
Pulicaria gnaphalodes 454
Pulicaria salviifolia 454
Pulicaria uliginosa 454
Pulsatilla 158
Pulsatilla campanella 158
Pycreus 99
Pycreus globosus 99
Pyrethrum 471
Pyrethrum galae 471
Pyrethrum parthenifolium 472
Pyrethrum pyrethroides 471
Pyrola 351
Pyrola rotundifolia 351
Pyrus 221
Pyrus communis 221
Pyrus korshinskyi 221
Pyrus regelii 221
Pyrus turcomanica 222
Q
Queria hispanica
317
R
Ranunculaceae 145
Ranunculus 159
Ranunculus arvensis 159
Ranunculus baldshuanicus 160
Ranunculus brevirostris 160
Ranunculus komarovii 160
Ranunculus linearilobus 160
Ranunculus mindshelkensis 160
Ranunculus natans 161
Ranunculus olgae 161
Ranunculus oxyspermus 161
Ranunculus paucidentatus 161
Ranunculus pinnatisectus 161
Ranunculus pulchellus 162
Ranunculus repens 162
Ranunculus rionii 162
Ranunculus rubrocalyx 162
Ranunculus rufosepalus 162
Ranunculus sceleratus 163
Ranunculus sewerzowii 163
Ranunculus songaricus 164
Ranunculus sphaerospermus 164
Ranunculus tenuilobus 164
Ranunculus trichophyllus 164
519
520
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Reaumuria 295
Reaumuria alternifolia 295
Reaumuria fruticosa 295
Reseda 273
Reseda lutea 273
Reseda luteola 273
Resedaceae 273
Rhabdotheca korovinii 423
Rhamnaceae 240
Rhamnus 240
Rhamnus cathartica 240
Rhamnus coriacea 240
Rhaponticum 449
Rhaponticum repens 449
Rheum 305
Rheum macrocarpum 305
Rheum maximowiczii 305
Rheum turkestanicum 306
Rheum сordatum 305
Rhinactinidia 457
Rhinactinidia popovii 457
Rhodiola 166
Rhodiola heterodonta 166
Ribes 165
Ribes meyeri 165
Rindera 368
Rindera tetraspis 368
Rochelia 367
Rochelia bungei 367
Rochelia cardiosepala 367
Rochelia disperma subsp. retorta
Rochelia leiocarpa 367
Rochelia peduncularis 368
Roemeria 138
Roemeria hybrida 138
Roemeria refracta 139
Rorippa 276
Rorippa palustris 276
Rosa 229
Rosa beggeriana 229
Rosa canina 230
Rosa corymbifera 230
Rosa ecae 230
Rosa fedtschenkoana 230
Rosa hissarica 231
Rosa kokanica 231
Rosa lehmanniana 231
Rosa maracandica 232
Rosa nanothamnus 233
Rosa transturkestanica 233
Rosaceae 217
Rosularia 169
Rosularia paniculata 169
Rosularia radicosa 169
Rosularia subspicata 169
Rubia 354
Rubia regelii 354
Rubiaceae 351
Rubus 224
Rubus caesius 224
Rumex 304
Rumex acetosa 304
Rumex chalepensis 304
Rumex conglomeratus 304
Rumex crispus 304
Rumex dentatus halacsyi 304
Rumex drobovii 304
Rumex halacsyi 304
Rumex marschallianus 304
Rumex pamiricus 304
Rumex paulsenianus 304
Rumex rectinervis 304
Rumex syriacus 304
Rumex tianschanicus 304
Ruppia 51
Ruppia maritima 51
Ruppiaceae 51
Rutaceae 266
S
367
Saccharum 103
Saccharum spontaneum 103
Sageretia 240
Sageretia thea 240
Sagina 317
Sagina saginoides 317
Sagittaria 49
Sagittaria trifolia 49
Salicaceae 248
Salicornia 336
Salicornia europaea 336
Salix 251
Salix alba 251
Salix blakii 251
Salix olgae 251
Salix pycnostachya 251
Salix rosmarinifolia 251
Salix songarica 251
Salix wilhelmsiana 251
Salsola 337
Salsola arbuscula 337
Salsola arbusculiformis 337
Salsola collina 337
Salsola dendroides 337
Salsola foliosa 338
Salsola iberica 338
Salsola leptoclada 338
Salsola micranthera 338
Salsola orientalis 338
Salsola paletzkiana 338
Salsola paulsenii 338
Salsola richteri 338
Salsola sclerantha 339
Salsola titovii 339
Salsola turkestanica 339
Salsola vvedenskyi 339
Salvia 407
Salvia aequidens 407
Salvia deserta 407
Salvia glabricaulis 407
Salvia macrosiphon 407
Salvia sarawschanica 407
Salvia sclarea 407
Salvia spinosa 408
Salvia submutica 409
Salvia virgata 409
Salvinia 35
Salvinia natans 35
Salviniaceae 35
Samolus 349
Samolus valerandi 349
Sanguisorba 229
Sanguisorba minor 229
Sanguisorba minor subsp.
balearica 229
Santalaceae 294
Sapindaceae 265
Saponaria griffithiana 329
Saussurea 444
Saussurea elegans 444
Saussurea salsa 444
Saxifraga 165
Saxifraga hirculus 165
Saxifraga sibirica 166
Saxifragaceae 165
Scabiosa 477
Scabiosa flavida 477
Scabiosa olivierii 477
Scabiosa rhodantha 477
Scabiosa songarica 477
Scandix 480
Scandix nodosa 480
Scandix pecten-veneris 480
Scandix stellata 480
Scapiarabis 276
Scapiarabis saxicola 276
Scariola orientalis 423
Schismus 117
Schismus arabicus 117
Schoenoplectus 95
Schoenoplectus litoralis 95
Schoenoplectus triqueter 95
Schoenus 99
Schoenus nigricans 99
Schrenkia 481
Schrenkia golickeana 481
Schrenkia pungens 482
Schrenkia vaginata 482
Schrenkiella 274
Schrenkiella parvula 274
Schtschurowskia 482
Schtschurowskia meifolia 482
Scirpus 95
Scirpus triquetriformis 95
Sclerochloa 117
Sclerochloa dura 117
Scorzonera 429
Scorzonera acanthoclada 429
Scorzonera bracteosa 429
Scorzonera circumflexa 429
Scorzonera hissarica 429
Scorzonera inconspicua 429
Scorzonera litwinowii 429
Scorzonera ovata 430
Scorzonera pusilla 431
Scorzonera songarica 431
INDEX OF LATIN NAMES
Scorzonera tragopogonoides 431
Scrophularia 381
Scrophularia gontscharovii 381
Scrophularia griffithii 381
Scrophularia heucheriiflora 381
Scrophularia incisa 381
Scrophularia integrifolia 381
Scrophularia kiriloviana 382
Scrophularia leucoclada 382
Scrophularia scoparia 382
Scrophularia striata 382
Scrophularia umbrosa 382
Scrophularia vvedenskyi 382
Scrophularia xanthoglossa 382
Scrophulariaceae 380
Scutellaria 383
Scutellaria adenostegia 383
Scutellaria comosa 383
Scutellaria cordifrons 383
Scutellaria galericulata 384
Scutellaria glabrata 385
Scutellaria immaculata 385
Scutellaria intermedia 385
Scutellaria oxystegia 385
Scutellaria phyllostachya 386
Scutellaria ramosissima 386
Scutellaria schachristanica 386
Scutellaria squarrosa 386
Secale 128
Secale cereale 128
Secale segetale 128
Secale sylvestre 128
Sedum 167
Sedum hispanicum 167
Sedum pentapetalum 167
Sedum tetramerum 168
Semenovia 498
Semenovia dasycarpa 498
Semenovia heterodonta 498
Semenovia pimpinelloides 498
Senecio 451
Senecio franchetii 451
Senecio olgae 451
Senecio subdentatus 451
Sergia 421
Sergia regelii 421
Serratula 450
Serratula algida 450
Serratula lancifolia 450
Seseli 488
Seseli calycinum 488
Seseli korovinii 488
Seseli lehmannianum 488
Seseli mucronatum 488
Seseli schrenkianum 489
Seseli seravschanicum 488
Seseli tenuisectum 489
Seseli turbinatum 489
Setaria 104
Setaria lutescens 104
Setaria verticillata 104
Setaria viridis 105
Sibbaldia 227
Sibbaldia tetrandra 227
Sideritis 387
Sideritis montana 387
Silene 320
Silene brahuica 320
Silene claviformis 320
Silene conica 320
Silene coniflora 321
Silene conoidea 321
Silene graminifolia 322
Silene guntensis 322
Silene incurvifolia 322
Silene kuschakewiczii 322
Silene latifolia 322
Silene longicalycina 322
Silene longicarpophora 325
Silene nana 323
Silene nevskii 323
Silene obtusidentata 323
Silene paranadena 323
Silene plurifolia 324
Silene praelonga 324
Silene praemixta 324
Silene pugionifolia 324
Silene ruinarum 324
Silene schischkinii 322
Silene stenantha 324
Silene tachtensis 324
Silene turkestanica 324
Silene uralensis subsp. apetala 324
Silene vulgaris 324
Simaroubaceae 269
Sinapis 287
Sinapis alba 287
Sinapis arvensis 287
Sisymbrium 274
Sisymbrium altissimum 274
Sisymbrium brassiciforme 274
Sisymbrium loeselii 274
Sium 487
Sium sisaroideum 487
Smelowskia 275
Smelowskia sisymbrioides 275
Smirnowia 179
Smirnowia turkestana 179
Solanaceae 373
Solanum 373
Solanum americanum 373
Solanum asiae-mediae 373
Solanum kitagawae 373
Solanum olgae 373
Solenanthus 369
Solenanthus circinatus 369
Solenanthus turkestanicus 370
Solidago 456
Solidago kuhistanica 456
Sonchus 424
Sonchus arvensis 424
Sonchus asper 425
Sonchus oleraceus 425
Sonchus palustris 425
Sophiopsis sisymbrioides 275
Sophora 172
Sophora alopecuroides 172
Sophora lehmanni 173
Sophora pachycarpa 173
Sorbus 222
Sorbus persica 222
Sorbus tianschanica 223
Sorghum 104
Sorghum halepense 104
Sparganium 92
Sparganium stoloniferum 92
Spergularia 319
Spergularia diandra 319
Spergularia marina 319
Spergularia marina 319
Spergularia media 319
Spergularia rubra 319
Spinacia 332
Spinacia turkestanica 332
Spiraea 217
Spiraea hypericifolia 217
Spiraea pilosa 217
Spirorrhynchus sabulosus 277
Stachyopsis 400
Stachyopsis oblongata 400
Stachys 405
Stachys hissarica 405
Stachys setifera 405
Stellaria 314
Stellaria alsinoides 314
Stellaria brachypetala 314
Stellaria karatavica 315
Stellaria media 314
Stellaria neglecta 314
Stellaria pallida 314
Stellaria turkestanica 314
Steptorhamphus crambifolius 423
Stipa 106
Stipa arabica 106
Stipa capillata 106
Stipa caragana 106
Stipa caucasica 107
Stipa hohenackeriana 107
Stipa kirghisorum 107
Stipa kurdistanica 107
Stipa lessingiana 107
Stipa lingua 107
Stipa lipskyi 107
Stipa margelanica 107
Stipa orientalis 107
Stipa richteriana 107
Stipa sareptana 107
Stipa splendens 107
Stipa trichoides 107
Stipa turkestanica 107
Stipagrostis 105
Stipagrostis karelinii 105
Stipagrostis pennata 106
Stizolophus 449
Stizolophus balsamita 449
Streptoloma 283
521
522
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN
Streptoloma desertorum 283
Strigosella 281
Strigosella africana 281
Strigosella brevipes 281
Strigosella grandiflora 281
Strigosella intermedia 281
Strigosella scorpioides 281
Strigosella stenopetala 281
Strigosella trichocarpa 281
Strigosella turkestanica 281
Stroganowia angustifolia 289
Stubendorffia lipskyi 290
Stubendorffia olgae 290
Stubendorffia orientalis 291
Suaeda 337
Suaeda acuminata 337
Suaeda altissima 337
Suaeda arcuata 337
Suaeda crassifolia 334
Suaeda dendroides 337
Suaeda heterophylla 337
Suaeda linifolia 337
Suaeda paradoxa 337
Suaeda physophora 337
Swertia 357
Swertia lactea 357
Symphyotrichum 458
Symphyotrichum ciliatum 458
T
Taeniatherum 130
Taeniatherum asperum 130
Taeniatherum caput-medusae 130
Takhtajaniantha 431
Takhtajaniantha pusilla 431
Tamaricaceae 295
Tamarix 295
Tamarix androssowii var.
transcaucassica 295
Tamarix arceuthoides 295
Tamarix elongata 296
Tamarix hispida 296
Tamarix hohenackeri 297
Tamarix laxa 296
Tamarix ramosissima 297
Tamarix smyrnensis 297
Tanacetopsis 463
Tanacetopsis karataviensis 463
Tanacetopsis mucronata 464
Tanacetopsis urgutensis 464
Tanacetum 472
Tanacetum parthenifolium 472
Tanacetum pseudachillea 472
Taraxacum 425
Taraxacum bessarabicum 425
Taraxacum bicorne 425
Taraxacum brevirostre 425
Taraxacum contristans 425
Taraxacum ecornutum 425
Taraxacum elongatum 425
Taraxacum erostre 425
Taraxacum glaucivirens 425
Taraxacum maracandicum 426
Taraxacum marginatum 426
Taraxacum minutilobium 426
Taraxacum modestum 426
Taraxacum monochlamydeum 426
Taraxacum montanum 426
Taraxacum nevskii 426
Taraxacum nuratavicum 426
Taraxacum officinale 426
Taraxacum popovii 426
Taraxacum reflexum 426
Taraxacum repandum 426
Taraxacum sonchoides 426
Taraxacum strobilocephalum 426
Tauscheria 277
Tauscheria lasiocarpa 277
Tetracme 283
Tetracme quadricornis 283
Tetracme recurvata 283
Tetrataenium olgae 497
Teucrium 383
Teucrium scordium subsp.
scordioides 383
Thalictrum 147
Thalictrum isopyroides 147
Thalictrum minus 147
Thalictrum sultanabadense 147
Thlaspi 292
Thlaspi arvense 292
Thlaspi kotschyanum 293
Thlaspi perfoliatum 292
Thymelaceae 272
Thymelaea 272
Thymelaea passerina 272
Thymus 411
Thymus seravschanicus 411
Thymus subnervosus 412
Torilis 480
Torilis arvensis 480
Torilis leptophylla 481
Trachomitum 358
Trachomitum scabrum 358
Tragopogon 432
Tragopogon capitatus 432
Tragopogon conduplicatus 432
Tragopogon kraschennikovii 433
Tragopogon kultiassovii 432
Tragopogon malikus 433
Tragopogon porrifolius subsp.
longirostris 433
Tragopogon pseudomajor 433
Tragopogon turkestanicus 433
Tragopogon vvedenskyi 433
Tribulus 171
Tribulus terrestris 171
Trichantemis 471
Trichanthemis karataviensis 471
Trichochiton inconspicuum 282
Trichodesma 370
Trichodesma incanum 370
Trifolium 177
Trifolium campestre 177
Trifolium fragiferum 177
Trifolium neglectum 177
Trifolium pratense 177
Trifolium repens 178
Triglochin 50
Triglochin palustre 50
Trigon ella gontscharovii 175
Trigonella 174
Trigonella adscendens 174
Trigonella arcuata 176
Trigonella aristata 174
Trigonella cancellata 174
Trigonella geminiflora 175
Trigonella grandiflora 175
Trigonella noeana 176
Trigonella orthoceras 176
Trigonella popovii 176
Trigonella verae 176
Tripleurospermum 473
Tripleurospermum disciforme 473
Tripolium 457
Tripolium pannonicum subsp.
tripolium 457
Trisetum 113
Trisetum spicatum 113
Trollius 149
Trollius komarovii 149
Tulipa 61
Tulipa affinis 61
Tulipa borszczowii 62
Tulipa buhseana 62
Tulipa lehmanniana 62
Tulipa micheliana 63
Turgenia 481
Turgenia latifolia 481
Turristis 276
Turristis glabra 276
Tussilago 452
Tussilago farfara 452
Tulipa 61
Tulipa affinis 61
Tulipa borszczowii 62
Tulipa buhseana 62
Тulipa dasystemon 62
Тulipa dasystemonoides 62
Тulipa korolkowii 62
Тulipa turkestanica 64
Typha 92
Typha angustata 92
Typha angustifolia 92
Typha latifolia 93
Typha laxmannii 93
Typha minima 93
Typhaceae 92
Tytthostemma alsinoides 314
U
Ulmaceae 241
Ulmus 241
Ulmus glabra 241
Ulmus laevis 241
Ulmus minor 241
INDEX OF LATIN NAMES
Ulmus pumila 241
Umbelliferae 479
Ungernia 76
Ungernia oligostroma 76
Urtica 242
Urtica dioica 242
Urticaceae 242
Utricularia 382
Utricularia minor 382
V
Vaccaria 326
Vaccaria hispanica 326
Valeriana 478
Valeriana chionophila 478
Valeriana fedtschenkoi 478
Valeriana ficarifolia 478
Valerianella 478
Valerianella coronata 478
Valerianella cymbocarpa 479
Valerianella dactylophylla 479
Valerianella muricata 479
Valerianella oxyrrhyncha 479
Valerianella plagiostephana 479
Valerianella szovitsiana 479
Valerianella turkestanica 479
Valerianella vvedenskyi 479
Vallisneria 49
Vallisneria spiralis 49
Velezia 329
Velezia rigida 329
Verbascum 380
Verbascum bactrianum 380
Verbascum blattaria 380
Verbascum erianthum 380
Verbascum songaricum 380
Verbascum turkestanicum 381
Verbena 382
Verbena officinalis 382
Verbenaceae 382
Veronica 377
Veronica anagallis-aquatica 377
Veronica anagalloides 377
Veronica arguteserrata 377
Veronica arvensis 377
Veronica beccabunga 378
Veronica biloba 378
Veronica campylopoda 378
Veronica capillipes 378
Veronica cardiocarpa 378
Veronica hederifolia 378
Veronica intercedens 378
Veronica oxycarpa 378
Veronica persica 378
Veronica polita 378
Veronica verna 378
Vexibia alopecuroides 172
Vexibia pachycarpa 173
Vicia 214
Vicia anatolica 214
Vicia angustifolia 215
Vicia ervilia 214
Vicia gracilior 215
Vicia hajastana 214
Vicia hyrcanica 214
Vicia kokanica 214
Vicia michauxii 214
Vicia peregrina 215
Vicia sativa subsp. nigra 215
Vicia subvillosa 215
Vicia tenuifolia 215
Vicia villosa 215
Vicoa albertoregelia 454
Viola 247
Viola alaica 247
Viola occulta 247
Viola suavis 248
Viola turkestanica 248
Violaceae 247
Vitaceae 170
Vitis 170
Vitis vinifera 170
Vulpia 122
Vulpia ciliata 122
Vulpia myuros 122
Vulpia persica 122
Vulpia unilateralis 123
W
Waldheimia 471
Waldheimia glabra 471
Waldheimia tomentosa 471
X
Xanthium 473
Xanthium spinosum 473
Xanthium strumarium 473
Xeranthemum 435
Xeranthemum longepapposum 435
Y
Youngia serawschanica 424
Z
Zannichellia 51
Zannichellia pedunculata 51
Ziziphora 410
Ziziphora clinopodioides 410
Ziziphora pamiroalaica 410
Ziziphora suffruticosa 410
Ziziphora tenuior 410
Zygophyllaceae 170
Zygophyllum 170
Zygophyllum jaxarticum 170
Zygophyllum macrophyllum 170
Zygophyllum miniatum 171
Zygophyllum oxianum 171
523
524
FLORA OF THE DZHIZAK PROVINCE, UZBEKISTAN