Veget Hist Archaeobot (2005) 14:562–570 DOI 10.1007/s00334-005-0060-4 ORIGINAL ARTICLE Andreas G. Heiss · Klaus Oeggl The oldest evidence of Nigella damascena L. (Ranunculaceae) and its possible introduction to central Europe Received: 15 September 2004 / Accepted: 22 December 2004 / Published online: 8 September 2005
Springer-Verlag 2005 Abstract After the beginning of metal processing at the transition from the Neolithic to the Bronze Age, further knowledge of ore mining and smelting had spread from the Near East to central Europe. In the copper ore deposits of Schwaz, in the central part of the Alps, the oldest traces of copper mining derive from the early to middle Bronze Ages. Investigation of a middle to late Bronze Age (1410–920 cal B.C.) slag-washing site in the area revealed a carbonised seed of Nigella damascena (Ranunculaceae) (love-in-a-mist) together with individual other food plants. The plant remains had become incorporated into the slag sediments by chance and had been preserved in an excellent state due to toxic copper salts contained in the soil. Nigella damascena, like N. sativa (black cumin), is traditionally used as a condiment and healing herb in southern Europe and the Near East, but has never grown in the wild in central Europe. Until now, there has been no evidence of prehistoric large-scale cultivation of N. damascena in central Europe. This leads to two possible conclusions: the find may either originate from an exchange of goods with the cultures in the Mediterranean during the Bronze Age, or indicate an introduction of the plant by an immigrant population from that area. Implicating the latter alternative together with the archaeological context of the ore processing site suggests that Nigella damascena had been introduced to the Alps by foreign miners in the course of ore exploitation during the middle to late Bronze Age. Keywords Bronze Age · Nigella damascena · Copper ore mining · Alps A. G. Heiss ()) · K. Oeggl University of Innsbruck, Institute of Botany, Sternwartestrasse 15, A-6020 Innsbruck, Austria e-mail: andreas.heiss@uibk.ac.at Introduction: Investigating Bronze Age metallurgical history in the eastern Alps The first steps in metal processing were taken in Asia Minor during the Neolithic. As of the 9th/8th millennium B.C. solid gold and copper were tempered into objects of ritual and status purposes. Subsequent improvements of metallurgical techniques are reported from the same region: mining and smelting of copper ore, casting solid copper, and creating bronze alloys (Pernicka 1990; Strahm 1994). The spread of metallurgical knowledge throughout Europe, originating from the high cultures in the Aegean and the Near East (Pernicka 1990; Strahm 1994), is documented as of the 5th millennium B.C. Prospectors are believed to have travelled the lands in search of profitable ore deposits, and immigrants brought their metallurgical skills with them (Krause 2004). During the transition from the late Neolithic (the Chalcolithic, or Copper Age) to the Bronze Age the use of the new working material increased greatly, together with the appearance of new and more elaborate techniques of extracting and processing copper. Copper ore deposits of prime importance are located in the central part of the Alps, as in the areas of Schwaz (Tyrol, Austria) and Mitterberg (Salzburg, Austria). The deposits around Schwaz are well-known for their predominance of tetrahedrite-tennantite type ores (Fahlerz, grey copper ore). While the use of copper in the eastern Alps is first documented from the beginning of the 4th millennium B.C. (Matuschik 1997), the oldest evidence of copper extraction from copper ore dates to the early to the middle Bronze Age, as for the Schwaz deposits (Goldenberg 1998, 2001). Prehistoric large-scale cultural and mercantile relations between the eastern Mediterranean and central Europe are already well documented by artefact finds. Della Casa and Primas (1998), for instance, have illustrated the trade between the Aegean and the eastern Adriatic in the period between 2800 and 2700 B.C. Giardino (1998) has studied the role of the Italian area as part of metal trade routes 563 between the Aegean and the rest of Europe: in the gulf of Naples the beginning of trade relations with the Aegean region was dated to Bronze Age B (1700–1500 B.C.). In the Alps, however, direct evidence for cultural and trade bonds to the Aegean region during Bronze Age has only rarely been found: first of all, the finding of several gold objects in the Alpine foothills near Bernstorf (Bavaria, Germany) is to be mentioned. Archaeological and metallurgical analyses of the discovered objects, among them a large crown-like diadem and a golden belt, have shown that this jewellery had been cast in the Aegean region. Moreover, their supposed ritual purpose refers unambiguously to cultures of the eastern Mediterranean (Gebhard 1999). For the area near Schwaz, finds of imported prestige weapons serve as another fine example of Bronze Age large-scale trade: three one-piece handle daggers (“Vollgriffdolche”) have been discovered up to now in the lower Inntal. One of them is of nětice origin, but had been forged mimicking archetypes from Italy (Schwenzer 2004). The traces of prehistoric mining are still visible everywhere on the south bank of the lower Inntal. Palaeoethnobotanical investigation of a slag-washing site near Schwaz has now supplied plant material that proved an unexpected insight into Bronze Age mining history. Fig. 1 Location of the site in Tyrol (Skamen 2002 onwards, modified; small map from Spiess 2002, modified) Materials and methods The site In the years 1997 and 2000 excavations were carried out in the low mountain ranges of the Inntal near Schwaz in Tyrol, Austria (Fig. 1). Archaeological investigation of a small southern tributary valley, the Maukengraben, revealed a huge slag-dump beneath the humus of a Picea abies (spruce) stand at about 900 m a.s.l. The dimensions of the slag-dump were estimated to an area of approximately 1000 m2, its thickness reaching from about 50 cm up to 1 m. Several samples of wood and charcoal were taken from the sediments and sent to the VERA (Vienna Environmental Research Accelerator) laboratory at the Institute for Isotope Research and Nuclear Physics of the University of Vienna for radiocarbon dating. The results (Table 1) gave evidence that the slag heap derived from a middle to late Bronze Age (1410–920 B.C.) slag-washing site where miners had post-processed crude ore. The first excavation campaign in 1997 revealed a wooden trough embedded in the slag layers (Fig. 2). Its walls consisted of wooden planks in an extraordinarily good state of preservation. The two lateral planks were later identified as Larix decidua (larch), but the front plank was not analysed. The bottom of the supposed flotation device consisted of a massive clay layer obviously thick Fig. 2 The excavation of Mauken A, with the partly uncovered wooden trough (photo: G. Goldenberg). The photograph shows transect 5a during 1997 campaign. Letters indicate the different layers (their limits highlighted): a forest soil, b slag dump, c basal clay layer Table 1 Results of the radiocarbon dating. Program used for calibration: OXCAL (Bronk Ramsey 1994) and VERA 0591 Sample no Mauk Mauk Mauk Mauk Mauk A A A A A 38 40 43 44 46 Lab no (VERA-1325) (VERA-1326) (VERA-1327) (VERA-1328) (VERA-1329) Material charcoal wood charcoal charcoal charcoal Method AMS AMS AMS AMS AMS d13C 26.2€1.3 24.7€1.3 26.7€1.3 25.9€1.3 25.1€1.4 14 C age 2860€35 2890€45 3050€40 3000€40 2885€40 Calibrated Age B.P B.P B.P B.P B.P 1 s (68.26%) 2 s (95.44%) 1110–930 B.C 1190–1000 B.C 1390–1260 B.C 1370–1130 B.C 1190–1000 B.C 1190–920 B.C 1260–920 B.C 1410–1130 B.C 1390–1090 B.C 1220–920 B.C 564 Table 2 Important plant remains from Mauken A Preservation Carbonised Useful plants Nigella damascena L. (seed) x Panicum miliaceum L. (caryopsis) x Rubus fruticosus agg. (pyrene) Rubus idaeus L. (pyrenes) Rubus sp. (pyrenes) Sambucus nigra L. (pyrenes) Herbaceous indicators of disturbance, and woody pioneer species Betula sp. (charcoal fragments) Daucus carota L. (schizocarp halves) Moehringia trinervia (L.) Clairv. (seed) Salix caprea L. (leaf) Stellaria media agg. (seeds) Urtica dioica L. (nutlets) Plants of coniferous woods and related heathland* Larix decidua Miller (total finds) L. decidua (wooden trough planks) L. decidua (charcoal fragments) x Picea abies (L.) Karsten (total finds) P. abies (twig fragments) P. abies (bark fragments) P. abies (needles) P. abies (bud) cf. Picea abies (total finds) cf. P. abies (wood fragments) cf. P. abies (charcoal fragments) x Pinus sp. (charcoal fragments) Plants of broad-leaved woods and related brushwood* Abies alba Miller (total finds) A. alba (wood fragments) A. alba (charcoal fragments) x A. alba (twig fragments) A. alba (needles) A. alba (seeds) Fagus sylvatica L. (total finds) Fagus sylvatica (wood fragments) Fagus sylvatica (charcoal fragments) x Fagus sylvatica (twig fragment) Fagus sylvatica (leaves) Fagus sylvatica (buds) Fagus sylvatica (bud scales) Lonicera xylosteum L. (seed) Taxus baccata L. (total finds) T. baccata (wood fragments) T. baccata (needle) Total finds Uncarbonised Sum Freq x x x x 1 1 1 28 6 187 1 1 1 3 2 9 x x x x x x 11 35 1 1 2 294 1 2 1 1 2 3 43 3 40 808 31 38 738 1 581 223 358 12 5 2 3 11 6 3 11 1 9 2 9 2 1657 194 947 6 506 4 271 23 229 1 14 2 2 1 7 6 1 10 2 9 1 10 1 8 2 8 1 3 2 1 1 2 1 1 x x x x x x x x x x x x x x x x x x x * Phytosociological class groups according to Ellenberg (1996). Freq. = Frequency enough to keep the water inside. From behind the southern plank, shreds of rough cloth were retrieved—perhaps the remnants of a sieve-like device for fractioning the slag grains. According to the excavation director, G. Goldenberg, the area with the wooden trough had most probably been covered by the sediments of a second slag washing device, situated further uphill, which explains why the trough itself was embedded within the slag. This second site has not been located yet. Sampling and sample processing The material was obtained by purposively sampling the slag layers inside the trough, and in its perimeter, and the basal clay layers were also sampled. Twelve soil samples and seven single macro remains were retrieved from the slag dump, making a total of 21.5 l (21.1 kg) of material. The samples contained both carbonised and uncarbonised plant remains. During sample-taking we were already able to judge the exceptionally good state of preservation of the uncarbonised macro remains, when several intact leaves and large leaf fragments of Fagus sylvatica (beech) in perfect condition became visible in the slag material. The slag’s high content of toxic heavy metal compounds (salts of copper, antimony, and arsenic, for instance) had added anti-microbial properties to the soil layers. Due to direct proximity of the site to a small brook, the Maukenbach, the slag sediments were waterlogged. Both conditions had caused a greatly slowed down decomposition of uncarbonised plant macro remains. All samples were immediately enclosed in airtight polyethylene bags and stored at +5C until analysis. The macro remains were extracted using the flotation method. Staggered sieves at mesh sizes of 0.25, 0.5, 1 and 2 mm were used for separation by particle size. Sub-sampling was done according to standard methods (Jacomet and Kreuz 1999) depending on the amount of plant material. Uncarbonised macro remains were preserved using Strasburger’s preserving agent (Gerlach 1969) consisting of equal amounts of 565 distilled water, 96% ethanol, and glycerine. A small amount of phenol was added (0.4 - 0.6% by weight), serving as a fungicide. Results Remains of wild and cultivated plants were extracted from the samples (Table 2). The species spectra in the soil samples did not give any evidence of an intentional plant deposit (as for example in refuse or storage pits) but indicated a rather “natural” sedimentation of plant remains from the surrounding vegetation. The wood and charcoal findings, as distinct remains of human activity, did not indicate a preference of the miners for certain woody taxa. For this reason the identified wild plants were used to characterise the site’s surrounding vegetation during the middle and late Bronze Age (Heiss 2001). Parts of the area had been covered by a zonal climax forest dominated by Picea abies (spruce), Abies alba (fir), and Fagus sylvatica (beech), with Pinus sp. (pine) and Larix decidua (larch) occurring sporadically. The presence of treeless patches was indicated in the samples by the presence of plants such as Urtica dioica (stinging nettle) and Sambucus nigra (elder), and the woody pioneer taxa Betula sp. (birch) and Salix sp. (willow). Cultivated plants were represented by a carbonised caryopsis of Panicum miliaceum (common millet). Though a cereal of only minor importance, the cultivation of common millet in the eastern Alpine region during the Bronze Age is documented by archaeological evidence (Oeggl 1992; Swidrak and Oeggl 1998). Lacking gluten protein and thus unsuitable for baking bread, P. miliaceum was usually consumed as millet gruel. Among the plant remains a carbonised seed of Nigella damascena (love-in-a-mist) was also found (Fig. 3). It was identified by its characteristic features: a triangularovate seed with tubercled surface, three longitudinal and several transverse ridges (crests) forming a conspicuous reticulum. Besides standard literature, reference material from several Nigella species was used for validating the identification (Fig. 4). However, according to A. Weber (pers. comm.) differentiation between the seeds of N. damascena and N. elata, which are both in the subsection Erobathos (see section on taxonomy and Table 3), is only possible by their surface structure: N. damascena is characterised by the presence of a small, nipple-like elevation on each of the faces’ convex cells (Fig. 3c). General information on the genus Nigella, and on N. damascena in particular Etymology The genus Nigella L. (nigella, fennel flower) was named after its seeds which, as a characteristic, are black in most Nigella species. The Latin term nigellus is diminutive of niger, meaning ‘black’ (Hegi 1975). The ancient Greek name melánqon (melanthion, ‘black [seeded] flower’) for Fig. 3 The seed of N. damascena from Mauken A. a ventral view, b lateral view, c detail of surface structure Nigella, allegedly for N. sativa, derives from the same attribute (Aufmesser 2002). The species epithet of Nigella damascena is named after the Syrian capital Damascus, referring to the plant’s eastern Mediterranean origin. Common names of N. damascena allude either to this geographical context, to the plant’s use as a condiment, or to its morphology (see below). 566 Table 3 Overview of the Nigella taxa Taxon names Sect. Garidella (L.) Spenn N. nigellastrum (L.) Willk N. unguicularis (Lam.) Spenn Sect. Komaroffia (O. Ktze.) Brand Nigella integrifolia Regel Sect. Nigella L Subsect. Erobathos (DC.) Zoh N. damascena L. N. elata Boiss. Subsect. Nigellaria (DC.) Terracc N. arvensis L. var. anatolica Zoh var. arvensis var. assyriaca (Boiss.) Zoh var. beershevensis Zoh var. glauca (Schkuhr) Boiss var. var. var. var. var. var. var. glaucescens Guss involucrata Boiss iranica (Zoh.) longicornis (Zoh.) Townsend microcarpa Boiss multicaulis Zoh negevensis Zoh var. palaestina (Zoh.) Zoh. & Feinbr var. simplicifolia Zoh var.mutica Bornm N. fumariifolia Kotschy N. hispanica L. var. hispanica var. intermedia Coss var. parviflora Coss N. sativa L. var. hispidula Boiss var. sativa N. segetalis Marsch. Bieb. var. armena (Stev.) Boiss var. segetalis N. stellaris Boiss Subsect. Nigellastrum (DC.) Zoh N. ciliaris DC N. orientalis L. N. oxypetala Boiss. Important synonyms Distribution Garidella nigellastrum L. Garidella unguicularis Lam. N. Medit., W. Irano-Turanian E. Medit Komaroffia diversifolia O. Ktze Irano-Turanian Erobathos coarctatus (Gmel.) Spach N. bithynica Azn Medit.; cultivated N.E. Medit N. arvensis subsp. arvensis sensu Fl. Europ. N. deserti Boiss N. carpatha Strid, N. degenii Vierh., N. doerfleri Vierh. N. divaricata Moris N. arvensis subsp. aristata (Sm.) Nyman N. assyriaca var. longicornis Zoh. N. taubertii Brand N. deserti var. latilabris Zoh N. arvensis subsp. negevensis (Zoh.) W.Greuter & Burdet N. divaricata var. palaestina Zoh. N. arvensis var. submutica Bornm. W. Irano-Turanian N. Medit. - Pontic - central European W. Irano-Turanian Saharo-Arabian E. Medit., W. Irano-Turanian W. Medit E. Medit W. Irano-Turanian W. Irano-Turanian E. Medit Medit. - Saharo-Arabian Saharo-Arabian E. Medit W. Irano-Turanian E. Medit N.E. Medit N. amoena Salisb. N. papillosa subsp. atlantica (Murb.) Amich ex G. L
pez N. gallica Jord N.W. Medit S.W. Medit N. truncata Viv N. indica Roxb N. verrucosa C. Koch N. bicolor Boiss N. cilicica Boiss. et Bal. in Boiss N.E. Medit.; cultivated widely cultivated N.E. Medit. - W. Irano-Turanian W. Irano-Turanian W. Irano-Turanian N.E. Medit N. oxypetala subsp. ciliaris Terracc. Nigellastrum flavum Moench N. lancifolia Hub.-Mor., N. latisecta P.H.Davis S.E. Medit. N.E. Medit., W. Irano-Turanian N.E. Medit., W. Irano-Turanian N.W. Medit Taxon names in bold script are accepted sensu Flora Europaea (Tutin et al. 1993). Data on taxonomy and distribution according to Zohary (1983). Medit. = Mediterranean Morphology Nigella (s.l.) species are characterised by the following features: all are annual plants with pinnatisect (rarely simple) leaves, their segments more or less linear. The hermaphrodite flowers are pentamerous and tetracyclic with conspicuous white to bluish or yellowish sepals persisting for a time on the ripening fruit. The nectariferous petals (honey leaves) are of reduced size, and of characteristic shape: each consists of a lower petaloid, bifid lip secreting nectar and an upper scale-like lip acting as a protective “lid” (Fig. 5b). The five carpels are partly united forming aggregate follicles (Hegi 1975; Zohary 1983). Slightly different morphological features can be recognised in Nigella subsection Erobathos, containing the two species N. damascena and N. elata. The number of carpels is 10, and they are entirely fused into a capsule (Fig. 5c, d; Hegi 1975; Tutin et al. 1993). Another typical feature is the involucre of finely dissected leaves enclosing the inflorescence. Hence originated some of the common names of N. damascena, such as ‘love-in-amist’, or ‘devil-in-the-bush’. Taxonomy The presence of the characteristic honey leaves mentioned above differentiates, among other things, the subtribe Nigellinae from the other Ranunculaceae. The Nigellinae were traditionally placed within the subfamily 567 Fig. 4 Reference seeds of some Nigella species (all in lateral view). a N. arvensis L., b N. damascena L., c N. hispanica L., d N. integrifolia Regel, e N. nigellastrum (L.) Willk., f N. sativa L., g N. orientalis L and G. nigellastrum L.) and monotypic Komaroffia (K. diversifolia Ktze). Illustrating Nigella‘s sub-generic taxonomy is, however, somewhat more intricate. Differing taxonomic approaches have resulted in an assemblage of many synonyms, some of them ambiguous. Linnaeus, when he first described this rather small genus, recorded six species (Linnaeus 1753, cited by Zohary 1983). Referring to Zohary (1983), the number of species names published up to now in the genus Nigella is at least 80 (!). – According to the Flora Europaea (Tutin et al. 1993), 14 native European Nigella species including six subspecies are differentiated today. – Strid (1970, cited by Zohary 1983) investigated many populations of the extremely polymorphic N. arvensis L. in the eastern Mediterranean, and split the taxon into several separate species. – Zohary (1983) described 14 species. But in contrast to the Flora Europaea he “lumped” all three Nigellinae genera into the genus Nigella at the rank of sections and also included some non-European taxa in his work. Furthermore, many species and all of the subspecies were reduced to the rank of varieties. The current work is based on Zohary’s taxonomic point of view. Fig. 5 Line drawing of Nigella damascena L. (Pabst 1887, 1889; modified). a sepal, b petal (honey leaf), c, d capsule, e seeds Helleboroideae, tribe Caltheae (Hegi 1975). However, modern multidisciplinary approaches to Ranunculaceae taxonomy suggest that Nigella and its subtribe are to be placed within the subfamily Ranunculoideae, as done by the Angiosperm Phylogeny Group (APG, Stevens 2001 onwards) and by Frohne and Jensen (1998). Apart from Nigella, the subtribe Nigellinae contains two other genera: Garidella (with G. unguicularis Lam. Table 3 gives an overview of the Nigella taxa treated by Zohary (1983), providing their geographical distribution and their most important synonyms (Tutin et al. 1993; Hooker and Jackson 1893 et seq.). Ecology and distribution All Nigella species are short-lived annuals, and are generally to be found in summer annual vegetation of semi- 568 arid areas. They frequently occur on disturbed soils. The whole subtribe Nigellinae (Nigella s.l.), in its native distribution, is limited to the meridional and southern temperate zones of Eurasia and northern Africa, particularly in the eastern Mediterranean and Near East. Hence this region is believed to be the taxon’s centre of diversity (Strid 1970, cited by Zohary 1983; Kstner et al. 2001). Most of the species can be found between the western Mediterranean-continental areas (Iberian Peninsula) and the Irano-Turanian zone (Turkey, Syria, and Israel). Only the following Nigella taxa occur outside of Europe: – N. glandulifera Freyn: China – N. media Pakh.: Central Asia – N. integrifolia Regel, syn. Komaroffia diversifolia Ktze.: Irano-Turanian zone to central Asia – N. stellaris Boiss.: Syria – N. arvensis s.l.: Morocco to Near East; archaeophyte (introduced by 15th century) in Europe (Kstner et al. 2001). Although being the most dispersed group within the genus, N. arvensis (s.l.) does not occur on the Iberian Peninsula. However, it is the only Nigella taxon growing wild in central Europe. N. damascena, initially a plant with an eastern Mediterranean distribution, is today a common ruderal plant in many parts of the Mediterranean. Outside this area, this species may only form ephemeral populations after escaping from cultivation. Just like N. sativa, N. damascena has been introduced to central Europe. Though neither species is to be found there in the wild (Tutin et al. 1993), both provide satisfying crop yields when cultivated in the area, as shown by modern experiments conducted in Thringen, Germany (Biertmpfel et al. 2001). Prehistoric, historical, and present cultivation and use The time when Nigella damascena was introduced to central Europe is unknown. Until now no archaeological data at all has been available on that species (Kroll 2004, pers. comm.). The seed from the site near Schwaz is actually the oldest archaeological evidence of N. damascena in central Europe so far. The evaluation of historical documents on plant cultivation and utilisation requires a high degree of caution. Due to inaccurate or (more commonly) completely lacking morphological descriptions it is often hard to identify the genera mentioned by antique and medieval authors. Of the ancient Greek authors, Theophrastos does not give any account about Nigella in his “Enquiry into Plants” from the 4th century B.C. (Hort 1916). In Hippocrates the plant melanthion (see section on etymology) is recommended for a small number of medical uses as, for instance, to promote pregnancy (Fuchs 1895–1900). In the 1st century A.D., Pliny the Elder mentions a plant called git in his “Natural History” (books XVII, XIX, XX; Von Jan 1857, 1859). He accredits various healing properties to the seeds of that plant which today is commonly interpreted as Nigella sativa. He recommends it as a digestive amarum, as an ingredient of antidotes curing snake bites and scorpion stings, as a fumigant for driving away snakes, and also as a condiment in bread. Pliny’s contemporary Dioscorides illustrates similar uses of git, adding those of an emmenagogue, galactagogue, vermifuge, and disinfectant (Aufmesser 2002). It should however be mentioned that he warns against possibly fatal effects of an overdose of git. Consequently it should be considered that Dioscorides might have confused Nigella with a poisonous plant: indeed, it is commonly accepted today that in classical and medieval literature Nigella (and N. sativa in particular) has often been confused with corn cockle, Agrostemma githago (Caryophyllaceae). Though quite dissimilar in growth habit and flowers, this once very common crop weed shares Nigella‘s characteristic of black seeds. This has also caused it share the same name in history. The species epithet of A. githago refers to this ancient connexion: “(seeds) resembling git(h)”, that is Nigella (Flckiger 1871). Medieval documents on useful plants and healing herbs unfortunately challenge the reader with the same inaccuracies: – Albertus Magnus mentions a plant named nigella, but is obviously writing about corn cockle (Flckiger 1871; Fischer-Benzon 1894). – In the 9th century A.D. the famous decree “Capitulare de villis vel curtis imperialibus” by Charlemagne, an assemblage of rules and guidelines for agriculture, lists git among desirable garden herbs (Fischer-Benzon 1894). No description of the plant is added. – In her “Physica”, well-known 12th century abbess Hildegard von Bingen refers to the healing herb (githerum) ratden and warns of harmful effects when eaten (Riethe 1989). The translator interprets that plant as being either Nigella sativa or Lolium temulentum L. (darnel ryegrass). Yet in the Renaissance, Fuchs (1543) clearly differentiates and describes the Nigella species known at that time. He assigns the same healing properties to N. damascena (“schwartz coriander”) as to the other species he mentions (N. arvensis, N. sativa). He recommends Nigella seeds for chiefly the same uses as Dioscorides and Pliny the Elder did, furthermore against ulcers, various pains, pulmonary diseases, and as a carminative agent. Nota bene that Fuchs, too, warns of fatal overdoses. Today N. damascena and N. sativa seeds are widely used in folk medicine in Arabian countries from Morocco to Pakistan, as well as in southern Europe. N. sativa is regarded a panacea (universal remedy) in Islamic folk medicine and used for mostly the same purposes as passed down from Pliny and Dioscorides (Ali and Blunden 2003). N. damascena seeds are used for example in Sicilian folk medicine as an emmenagogue and diuretic 569 (Ballero and Fresu 1993). Modern analyses show oestrogenic activity (Agradi et al. 2002) and analgesic properties (Bekemeier et al. 1967). Apart from medical uses, Nigella seeds are still used as spices today. Both N. sativa and N. damascena are used in bread and cheese, while N. damascena also serves as a condiment in sweets (D’Antuono et al. 2002). N. damascena also has some horticultural importance as it is cultivated in many varieties as an ornamental plant. Discussion and conclusions The Nigella damascena seed that has been retrieved from the Bronze Age slag-washing site near Schwaz is definitely a remarkable find. As far as we currently know about the history of natural distribution of N. damascena, it can be assumed that the plant had been imported to the area by humans. Considering the archaeological context of an ore processing site, the seed had obviously been brought to its place of discovery by someone involved in mining, ore processing, or metal trade. However, we cannot reliably assign any ethnobotanical relevance to the Nigella seed. Due to poor documentation of historical use of N. damascena as a spice and as a remedy, many questions are left unanswered. The seed may have been part of a Bronze Age miner’s diet, flavouring his millet gruel. Only the single millet caryopsis retrieved from the same place supports this theory. The plant may also have been an ephemeral ruderal plant in the area of the Inntal, imported from the south together with agricultural crops, or spices. In this connexion the presence of other ruderal plants at the mining site (Daucus carota, Stellaria media agg. and Urtica dioica) should be emphasised once again. At any rate, the find of the N. damascena seed in the mining site near Schwaz is clear evidence of the association between Alpine copper ore mining and the Mediterranean during the middle to late Bronze Age. Effectively, the find points to either trade or migration between southern or south-eastern regions and central Europe. These large-scale bonds have been documented before by artefact finds and by evidence of the spread of cultural and technological innovations. It is plausible to assume that N. damascena had been brought to the Alps by the same routes. Acknowledgements The current work is basing on data from the FWF project P12049 supported by the Austrian Science Fund (“Fonds zur Frderung der wissenschaftlichen Forschung”). We thank H. Kroll (Kiel) for his investigations on archaeological finds of N. damascena, and R. Krause (Landesdenkmalamt BadenWrttemberg) for the additional information on archaeometallurgy and artefact finds in the Alps during the Bronze Age. We also thank A. Weber (Wien) for the valuable information on differentiating between N. damascena and N. elata. Further thanks go to H.-P. Stika (Hohenheim) for providing us with seeds of N. integrifolia References Agradi E, Fico G, Cillo F, Francisci C, Tom F (2002) Estrogenic activity of Nigella damascena extracts, evaluated using a recombinant yeast screen. Phytother Res 16:414–416 Ali BH, Blunden G (2003) Pharmacological and Toxicological Properties of Nigella sativa. Phytother Res 17:299–305 Aufmesser M (transl) (2002) Pedanius Dioscorides De materia medica [Fnf Bcher ber die Heilkunde]. Altertumswissenschaftliche Texte und Studien, vol 37. Olms-Weidmann, Hildesheim Ballero M, Fresu I (1993) Le piante di uso officinale nella Barbagia di Seui (Sardegna Centrale). Fitoterapia 64:141–150 Bekemeier H, Leuschner G, Schmollack W (1967) Antipyretische, antidematse und analgetische Wirkung von Damascenin im Vergleich mit Acetylsalicylsure und Phenylbutazon. Archives Internationales de Pharmacodynamie et de Th rapie 168:199– 211 Biertmpfel A, Vetter A, Zirr K (2001) Ergebnisse der Evaluierung von Schwarzkmmelarten und -herknften (Nigella sativa L.) hinsichtlich Ertrag, Inhaltsstoffen und Nutzungsmglichkeiten. Zeitschrift fr Arznei- und Gewrzpflanzen 3/2001:161–164 Bronk Ramsey C (1994) Analysis of chronological information and radiocarbon calibration: the program OxCal. Archaeolog Comput Newsletter 41:11–16 D’Antuono LF, Moretti A, Lovato AFS (2002) Seed yield, yield components, oil content and essential oil content and composition of Nigella sativa L. and Nigella damascena L. Industr Crops Prod 15:59–69 Della Casa PH, Primas M (1998) La production et la circulation des m taux du IIIe au Ier mill naire sur la c te est de l’Adriatique d’aprs les analyses m tallurgiques de Velika Gruda (Mont n gro). In: Mordant, C., Pernot, M., Rychner, V. (eds), L’Atelier du bronzier en Europe du XXe au VIIIe sicle avant notre re. Actes du colloque international Bronze ‘96, Neuch tel et Dijon. Tome I. Paris, CTHS, pp 239–245 Ellenberg H (1996) Vegetation Mitteleuropas mit den Alpen. In kologischer, dynamischer und historischer Sicht. UTB, Stuttgart Fischer-Benzon R (1894) Altdeutsche Gartenflora. Lipsius & Tischer, Kiel/Leipzig. Reprinted 1991 by Sndig Reprint, Vaduz Flckiger FA (1871) Nigella Seeds, or Black Cummin. Am J Pharm 43:1871. In: Kress H (ed) (2000–2004) Henriette’s herbal homepage. Classical texts: journals and periodicals. http:// www.ibiblio.org/herbmed/eclectic/journals/ajp-1871.html Frohne D, Jensen U (1998) Systematik des Pflanzenreichs – Unter besonderer Bercksichtigung chemischer Merkmale und pflanzlicher Drogen, 5th edn. Wissenschaftliche Verlagsgesellschaft, Stuttgart Fuchs L (1543) New Kreterbuch [The New Herbal of 1543]. Annotated reprint 2001. Taschen, Kln Fuchs R (ed) (1895–1900) Hippokrates Corpus Hippocraticum [Smmtliche Werke]. 3 volumes. Lneburg, Mnchen Gebhard R (1999) Der Goldfund von Bernstorf bei Kranzberg. Bayerische Vorgeschichtsbltter 64:1–18 Gerlach D (1969) Botanische Mikrotechnik – Eine Einfhrung. Thieme, Stuttgart Giardino C (1998) Tyrrhenian Italy and Sicily in the protohistoric metal trade across the Mediterranean: An archaeometallurgical outline. In: Mordant C, Pernot M, Rychner V (eds) L’Atelier du bronzier en Europe du XXe au VIIIe sicle avant notre re. Actes du colloque international Bronze ‘96, Neuch tel et Dijon. Tome II. Paris, CTHS, pp 157–168 Goldenberg G (1998) L’exploitation du cuivre dans les Alpes Autrichiennes l’ ge du Bronze. In: Mordant C, Pernot M, Rychner V (eds), L’Atelier du bronzier en Europe du XXe au VIIIe sicle avant notre re. Actes du colloque international Bronze ‘96, Neuch tel et Dijon. Tome II. Paris, CTHS, pp 9–24 Goldenberg G (2001) Bronzezeitlicher Kupferbergbau in Nordtirol. In: Brunn A, Steinacker C, Jordan T (eds), Archologie Online. http://www.archaeologie-online.de/thema/2001/02/ Hegi G (ed) (1975). Illustrierte Flora von Mitteleuropa. vol. III/3, 2nd edn., Parey, Berlin 570 Heiss AG (2001). Anthrakologische und paloethnobotanische Untersuchungen im bronzezeitlichen Bergbaugebiet Schwaz – Brixlegg (Tirol). Unpublished diploma thesis. University of Innsbruck Hooker JD, Jackson DB (eds) (1893 et seq) Index Kewensis Plantarum Phanerogamarum, vol 2, 19 supplement. Clarendon Press, Oxford, pp 1901–1991 Hort A (ed) (1916) Theophrastos Per jutkwn storwn [Peri phytikon historion, Enquiry into plants] vol 2 reprinted 1961. Harvard University Press, Cambridge Massachusetts Jacomet S, Kreuz A (1999) Archobotanik. Ulmer, Stuttgart Von Jan L (ed) (1857) C. Plini Secundi Naturalis historiae libri XXXVII. Libb. XVI–XXII. Teubner, Leipzig Von Jan L (ed). (1859) C. Plini Secundi Naturalis historiae libri XXXVII. Libb. XXIII—XXXII. Teubner, Leipzig Kstner A, Jger EJ, Schubert R (2001) Handbuch der Segetalpflanzen Mitteleuropas. Springer, Wien Krause R (2004) Eliten der frhen Bronzezeit zwischen Alpen und Donau. In: Krieger, Frsten und Helden. Katalog Trento 2004 (in press) Kroll H (2004) Literature on archaeological remains of cultivated plants 1981–2004. http://www.archaeobotany.de/database.html Matuschik I (1997) Der neue Werkstoff – Metall. In: Planck D (ed) ALManach 2, Archaeologisches Landesmuseum Baden-Wrttemberg. Theiss, Stuttgart, pp 16–25 Oeggl K (1992) Botanische Untersuchungen zur menschlichen Besiedlung im mittleren Alpenraum whrend der Bronze- und Eisenzeit. In: Metzger IR, Gleirscher P (eds) Die Rter – I Reti. Athesia, Bozen, pp 709–721 Pabst G (ed) (1887, 1889) Khler’s Medizinal-Pflanzen in naturgetreuen Abbildungen mit kurz erluterndem Texte. Gera-Untermhaus, Khler Pernicka E (1990) Gewinnung und Verbreitung der Metalle in prhistorischer Zeit. Jahrbuch Rmisch Germanisches Zentralmuseum 37:21–129 Riethe P (ed) (1989) Hildegard von Bingen Liber subtilitatum diversarum naturarum creaturarum [Naturkunde, Physica]. 4th edn. Mller, Salzburg Schwenzer S (2004) Frhbronzezeitliche Vollgriffdolche: typologische, chronologische und technische Studien auf der Grundlage einer Materialaufnahme von Hans-Jrgen Hundt. Kataloge Vor- u. Frhgeschichtlicher Altertmer, 36, Mainz Skamen J (ed) (2002 onwards) Tiroler Raumordnungs-Informationssystem (tiris), Kartengalerie. http://tiris.tirol.gv.at/kartengal/karteng.cfm Spiess E (ed) (2002) Schweizer Weltatlas. Neuausgabe 2002. Lehrmittelverlag des Kantons Zrich, Zrich. http://www.educeth.ch/geographie/weltatlas/ Stevens PF (2001 onwards) Angiosperm Phylogeny Website. http:// www.mobot.org/MOBOT/research/APweb/ Strahm C (1994) Die Anfnge der Metallurgie in Mitteleuropa. Helv Archaeol 25:2–39 Swidrak I, Oeggl K (1998) Palaeoethnobotanische Untersuchungen von Bodenproben aus der bronzezeitlichen Siedlung von Sotćiastel. In: Tecchiati U (ed) Sotćiastel—Un abitato fortificato dell’et del bronzo in Val Badia. Istitut Cultural Ladin “Micur de R”, Soprintendenza Provinciale ai Beni Culturali di Bolzano, pp 335–346 Tutin TG et al (1964–80) Flora Europea, vol 1 (1964, 2nd edn. 1993). Cambridge University Press, Cambridge Zohary M (1983) The genus Nigella (Ranunculaceae)—a taxonomic revision. Plant Systematics and Evolution 142:71–107