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Studies on the distribution of Verbascum showed that 96 % of the Persian species are Irano-Turanian elements. Taxonomy of the genus has been revised. Based on the results, 42 species of Verbascum are listed from Iran. V. kermanense Hub.-Mor. and V. straussi (Bornm.) Hub.-Mor. were regarded as a synonymy of V. gabrielae Bornm. and V. nudicaule (Wydl.) Takht. respectively.
2007 •
A new endemic species, Verbascum shahsavarensis, is described from north of Iran. Based on two unique characters of this species, these being two anterior longitudinally obliquely inserted anthers and an ebracteolate pedicel, we compared it to species described by Murbeck in 1933. The new species can be easily distinguished from the other Iranian species by having bi-colored hairs on the stamen filaments and the unique anther and pedicel characters. A distribution map is provided. The name of new species comes from Shahsavar, the ancient name of Tonekabon, an old coastal city on the Caspian Sea.
The high morphological diversity among species of the genus Verbascum causes problems in the delimitation of the species. Therefore, there is a difficulty in offering a natural and well organized classification for the species of the genus. This research was based on the leaf anatomy of 20 species of Verbascum growing in north and middle of Iraq. Leaf samples from 20 species of Verbascum were cleared and stained, was then to examine by light microscopy. Idioblasts occurred both adaxially and abaxially and have a thin primary wall and were empty at maturity, the increase of idioblasts leads to irregular the mesophyll thus the shape of blade in cross section and can divided the species in to irregular mesophyll and regular mesophyll. All species of Verbascum have crystals appeared very clear in the epidermis.we recorded stomatal type and found all the species under study have anomocytic type of stomata and characterized by the fact uniformly amphistomatic (stomata both abaxial and adaxial) and the stomatal apparatus lacked accessory (subsidiary) cells also. The shape of the guard cell on the upper and lower surface are reniform in all species under study, But there are differences in the ordinary cells surrounding the stomata and the cell shaped where are appeared as two types, the first type is the ordinary cells straight walls and ribbed and the cell shapes are rectangular to oval and the second type is the ordinary cells are undulate and irregular walls or wavy edge and the cell shape are rectangular to polygonal.Study showed that the trichomes are normally divided into two groups depending on that in Verbascum species under study to glandular trichomes and unglandular trichomes and the last one also can divided in to two types of trichomes, the dendroid trichomes and asteroid trichomes, clear from result of study the species V. agrimoniifolium is smooth from upper surface and the trichomes appear just in the lower surface.
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