Ceylon Journal of Science (Bio. Sci.) 43(1): 1-36, 2014
Article first published online: August 25, 2014
DOI: http://dx.doi.org/10.4038/cjsbs.v43i1.7280
LEAD ARTICLE
Liverworts and Hornworts of Sri Lanka: a revised checklist
David G. Long1* and Sumudu C.K. Rubasinghe2
1
Royal Botanic Garden, Edinburgh EH3 5LR, UK.
Department of Botany, University of Peradeniya, Sri Lanka.
2
ABSTRACT
A revised checklist of liverworts and hornworts of Sri Lanka is presented based on an extensive review of
published literature sources and incorporating synonymy from numerous floristic reports and taxonomic
revisions. A number of erroneous records and doubtfully recorded taxa have been excluded. In total 82
genera and 327 species of liverworts and 4 genera and 5 species of hornworts are accepted. For each species
literature sources are listed. For synonyms the source of the synonymy is given. Further, names whose type
specimens originated from Sri Lanka are indicated. Two new formal taxonomic changes are propsed:
Bazzania obtusa D.G.Long & Rubasinghe, nom. nov. and Heteroscyphus fleischeri (Steph.) D.G.Long &
Rubasinghe, comb. nov. An index to all accepted names and synonyms is appended.
Keywords: Anthocerotopsida, bryophyte, Ceylon, hornwort, liverwort, Marchantiopsida.
INTRODUCTION
Bryophytes which comprise mosses (Class
Bryopsida), liverworts (Marchantiopsida) and
hornworts (Anthocerotopsida) have long been
recognised as very important components of almost
all the earth's major ecosystems. Although they are
a much smaller group compared to Angiosperms in
terms of species numbers, the significance of their
dominance and biomass in many ecosystems such
as wet forests and peatlands, is now becoming more
widely recognised, particularly in aspects such as
water retention and as a vitally important carbon
sink (Vanderpoorten & Goffinet, 2009).
Yet, in many countries, especially in the tropics,
knowledge of bryophytes is decades behind that of
their Angiosperm cousins. As a conseqequence,
their protection, conservation and utilisation are
often largely ignored or overlooked because of this
lack of knowledge, and the scarcity of means to
identify them. Without taxonomic knowledge and
identification manuals, we cannot apply names and
communicate about the plants. As stressed in the
recent historical review of bryology in Sri Lanka
(Rubasinghe & Long, 2014) "the biggest
impediment to progress in documenting the
bryophytes of Sri Lanka is lack of a modern
Bryophyte Flora, and this should be regarded as a
priority for the future".
*Corresponding Author’s email: d.long@rbge.ac.uk
As a first step towards producing a Bryophyte Flora,
a checklist is a very useful way of gathering
together existing knowledge about the group and
working out the scale of the task. Abeywickrama &
Jensen (1978a, b) compiled the first checklists for
Sri Lanka of liverworts and mosses. Although
excellent at the time, these are now out-of-date in
many ways and require updating. In the case of the
mosses, this was done by O'Shea (2002) who
provided an excellent moss list incorporating much
new data. However, this has not been done for
liverworts and hornworts until now. Abeywickrama
& Jensen (1978a) gleaned their liverwort data from
only 22 publications. Since then, there have been
four significant developments: i, there has been
much more field collecting of liverworts in Sri
Lanka by a range of bryologists, notably J. Eggers,
A. Jayasuriya, C. Ruinard, A. Schäfer-Verwimp and
P. Tixier (Rubasinghe & Long, 2014); ii, there have
been numerous floristic papers published in several
international journals enumerating the results of
these collectors' fieldwork (e.g. Eggers & SchäferVerwimp, 1987; Jayasuriya, 1984; Hattori, 1979;
Onraedt, 1978, 1979a, b, 1981, 1983, 1985; Tixier,
1977); iii, there are important recent published
studies on liverworts of adjacent regions, such as
India (Asthana & Srivastava, 2003; Dey & Singh,
2012; Sharma & Srivastava, 1993; Srivastava &
Srivastava, 2002), China (Zhu & So, 2001) and
Papua New Guinea (Piippo, 1984 a,b; 1985a,b, etc.)
which are highly relevant to species in Sri Lanka;
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Long and Rubasinghe
and iv, the publication of new monographic studies
at both regional and global levels for particular
groups found in Sri Lanka, such as Cololejeunea
(Tixier, 1985), Frullania (Hattori, 1979),
Marchantia (Bischler, 1989), Metzgeria (So,
2003b), Plagiochila (Inoue, 1984), Pleurozia
(Thiers, 1993), Radula (Yamada, 1979a), and
Schistochila (So 2003a).
The present list is also a compilation, largely based
on published literature, but it attempts several
refinements over earlier work, including adopting
new taxonomic placements of genera and species
resulting from DNA-based studies, new synonymy
from regional and global taxonomic revisions and
monographs, a wider trawl of literature old and new
and a critical reassessment of some doubtful records
based on probable misidentifications.
The list is arranged alphabetically by family, genus
and species, and accepted genera and species are
numbered. Families follow Crandall-Stotler et al.
(2009) with some recent refinements such as
segregation
of
Anastrophyllaceae
from
Lophoziaceae. Synonyms, where relevant to Sri
Lanka, are given under the currently accepted
name, and the source of synonymy is cited. Where
names are based on Sri Lankan type specimens this
is indicated. Sources of records are indicated by
numbered references in square brackets and listed
in the bibliography. Two new formal taxonomic
changes are proposed in Appendix I. Accepted
names and synonyms are listed in Appendix II with
a reference to accepted name for each synonym.
LIVERWORTS
Acrobolbaceae
1.1 Marsupidium knightii Mitt. [37, 64]
Adelanthaceae
2.1. Cuspidatula flexicaulis (Nees) Váňa & L.Söderstr.
Jungermannia flexicaulis Nees [148]
Jamesoniella flexicaulis (Nees) Schiffn. [55, 169]
(Jungermannia colorata Lehm. var. ceylanica Gottsche nom. nud. [44] may belong here).
3.1 Denotarisia linguifolia (De Not.) Grolle [55, 169]
Jamesoniella ovifolia (Schiffn.) Schiffn. [1, 2, 110, 227]; syn. fide Grolle (1971).
(Syzygiella elongella (Taylor) K. Feldberg, Váňa, Hentschel & J. Heinrichs
Jamesoniella elongella (Taylor) Steph. [1, 2, 227]. Excluded from Sri Lanka, reports refer to
Cuspidatula flexicaulis, fide Grolle (1971)).
4.1 Syzygiella securifolia (Nees ex Lindenb.) Inoue [118, 213]
Plagiochila securifolia Nees ex Lindenb. [1, 2, 45, 148, 227]; type from Sri Lanka
Plagiochila variegata Lindenb. [148]; syn. fide Inoue (1984)
Syzygiella variegata (Lindenb.) Schiffn. [1, 2, 114, 200]; syn. fide Inoue (1979)
4.2 Syzygiella subintegerrima (Nees) Spruce [2, 115]
Anastrophyllaceae
5.1 Anastrophyllum imbricatum (Wilson ex Gottsche, Lindenb. & Nees) Steph.
Jungermannia imbricata Wilson ex Gottsche, Lindenb. & Nees [45, 148]
5.2 Anastrophyllum piligerum (Reinw., Blume & Nees) Steph. [1, 2, 200, 227]
Jungermannia piligera Reinw., Blume & Nees [148]
6.1 Plicanthus birmensis (Steph.) R.M.Schust.
Chandonanthus birmensis Steph. [37]
6.2 Plicanthus hirtellus (F.Weber) R.M.Schust.
Chandonanthus hirtellus (F.Weber) Mitt. [1, 2, 98, 110, 169, 227, 240]
7.1 Schizophyllopsis bidens (Reinw., Blume & Nees) Váňa & L. Söderstr.
Anastrophyllum bidens (Reinw, Blume & Nees) Steph. [169]
Aneuraceae
8.1 Aneura pinguis (L.) Dumort.
Riccardia pinguis (L.) Gray [1, 2]
9.1 Riccardia canaliculata (Nees) Schiffn. [41, 42]
9.2 Riccardia multifida (L.) Gray [1, 2, 128]
Revised checklist to the Liverworts and Hornworts of Sri Lanka
3
Aytoniaceae
10.1 Plagiochasma rupestre (G.Forst.) Steph. [19, 199]
Plagiochasma nepalense (Lehm. & Bisch.) Steph. [1, 2, 80, 128]; syn. fide Bischler (1979).
11.1 Reboulia hemisphaerica (L.) Raddi [1, 2, 169, 199]
Blasiaceae
12.1 Blasia pusilla L. [80]
Calypogeiaceae
13.1 Calypogeia apiculata (Steph.) Steph. [1, 2, 136, 169, 227]
13.2 Calypogeia arguta Nees & Mont. [169]
13.3 Calypogeia ceylanca S.Hatt. & Mizut. [80, 169]; type from Sri Lanka.
13.4 Calypogeia fissa (L.) Raddi [169, 205]
13.5 Calypogeia fragilis (Steph.) Inoue & H.A.Miller [2]
14.1 Metacalypogeia alternifolia (Nees) Grolle
Calypogeia alternifolia (Nees) Steph. [1, 2]
15.1 Mnioloma fuscum (Lehm.) R.M.Schust.
Calypogeia fusca (Lehm.) Steph. [58, 169]
Calypogeia baldwinii Aust. [52, 53, 134]; syn. fide Grolle (1977b)
Cephaloziaceae
16.1 Nowellia curvifolia (Dicks.) Mitt. [169]
Cephaloziellaceae
(Cephaloziella capillaris (Steph.) Douin [2, 121]; Kitagawa (1969) rejected these reports which may refer
to Cylindrocolea kiaeri)
17.1 Cylindrocolea kiaeri (Austin) Váňa
Cephaloziella kiaeri (Austin) Pearson [169];
Cephalozia willisana Steph. [1, 2, 227]; type from Sri Lanka; syn. fide Váňa et al. (2013)
Cephaloziella willisana (Steph.) N.Kitag. [131]
18.1 Gottschelia schizopleura (Spruce) Grolle [54, 55, 169]
Jamesoniella microphylla Schiffn. [1, 2, 195, 227]; syn. fide Onraedt (1981).
Jungermannia ceylanica Steph. [1, 2]; type from Sri Lanka; syn. fide Grolle (1971)
(Jungermannia ceylanica Gottsche nom. nud. [44]; may belong here).
Note: Gottschelia was formerly placed in Lophoziaceae but Feldberg et al. (2009) showed that was
incorrect; it is provisionally placed in Cephaloziellaceae.
Cyathodiaceae
19.1 Cyathodium cavernarum Kunze [221]
(Cyathodium foetidissimum Schiffn. [199]. Note: not reported from Sri Lanka or India by Srivastava &
Dixit (1996); record considered doubtful).
19.2 Cyathodium smaragdinum Schiffn. ex Keissl. [1, 2, 199, 221]
Dumortieraceae
20.1 Dumortiera hirsuta (Sw.) Nees. sens. lat. [1, 2, 80, 110, 148, 169, 199]
Exormothecaceae
21.1 Exormotheca ceylonensis Meijer [1, 2, 4, 80, 128, 144, 199]
Fossombroniaceae
(Fossombronia wondraczekii (Corda) Dumort. [2, 80, 121, 169]. Excluded from East Asia by Krayesky et
al. (2005) though no Sri Lankan specimens were studied; may refer to an Asiatic species).
Frullaniaceae
22.1 Frullania acutiloba Mitt. [1, 2, 85, 93, 169, 206, 227, 249]
Frullania hampeana Nees var. acutiloba (Mitt.) S.Hatt. [87]; reinstated as a distinct species by
Hattori (1980a).
4
Long and Rubasinghe
22.2 Frullania alstonii Verd. [1, 2, 91, 169, 191, 249, 251]; type from Sri Lanka
Frullania perocellata Onr. [166, 169]; type from Sri Lanka; syn. fide Hattori (1985)
22.3 Frullania apiculata (Reinw., Blume & Nees) Dumort. [1, 2, 93, 148, 169, 249]
22.3a Frullania apiculata (Reinw., Blume & Nees) Dumort. var. goebelii Schiffn. [249]
22.4 Frullania arecae (Spreng.) Gott. [259]
Frullania wallichiana Mitt. [83, 93, 169]; syn. fide Yuzawa (1991)
22.5 Frullania brotheri Steph. [87, 91, 95]
22.6 Frullania campanulata Sande Lac. [169]
22.7 Frullania capilliformis Steph. [227]; type from Sri Lanka.
22.8 Frullania claviloba Steph. [169]
22.9 Frullania ericoides (Nees) Mont. [37, 88, 93, 94, 169]
Frullania squarrosa (Reinw., Blume & Nees) Nees [1, 2, 44, 148, 248]; syn. fide Piippo (1990).
Frullania laciniosa Lehm. [148]; syn. fide Hattori (1975b).
22.10 Frullania gaudichaudii (Nees & Mont.) Nees & Mont. var. ceylanica (Nees) S.Hatt. [83, 94, 169]
Frullania ceylanica Nees [1, 2, 45, 83, 123, 148, 249]; type from Sri Lanka
Frullania nietneri Gottsche ex Steph. [44, 227]; type from Sri Lanka; syn. fide Verdoorn (1930).
22.11 Frullania gracilis (Reinw., Blume & Nees) Dumort. [1, 2, 87, 91, 169, 249]
22.12 Frullania grandistipula Lindenb. [1, 2, 86, 87, 91, 93, 97, 169, 206, 227, 249]
Frullania rugosa Mitt. [1, 2, 148, 227]; syntype and lectotype from Sri Lanka; syn. fide Hattori
(1986)
Frullania grandistipula Lindenb. var. rugosa (Mitt.) S.Hatt. [88]
22.13 Frullania hottana S.Hatt. [97, 169]
22.14 Frullania hypoleuca Nees [169]
22.15 Frullania intermedia (Reinw., Blume & Nees) Dumort. [1, 2, 95, 123, 249]
Frullania explanata Mitt. [148]; type from Sri Lanka; syn. fide Hattori (1975b)
22.16 Frullania meyeniana Lindenb. [88, 93, 169]
22.17 Frullania microauriculata Verd. [1, 2, 169]
22.17a Frullania microauriculata Verd. var. rotundior Verd. [87, 249]; syntype from Sri Lanka
22.18 Frullania moniliata (Reinw., Blume & Nees) Mont. [110, 195, 227, 234]
Frullania tamarisci (L.) Dumort. subsp. moniliata (Reinw., Blume & Nees) Kamimura [129]
22.18a Frullania moniliata (Reinw., Blume & Nees) Mont. subsp. breviramea (Steph.) Verd. [1, 2, 249]
Frullania tamarisci (L.) Dumort. subsp. obscura (Verd.) S.Hatt. var. breviramea (Steph.) S.Hatt.
[169]
22.18b Frullania moniliata (Reinw., Blume & Nees) Mont. subsp. obscura Verd. [1, 2, 234, 249]
Frullania tamarisci (L.) Dumort. subsp. obscura (Verd.) S.Hatt. [82, 88, 93]
22.19 Frullania monocera (Taylor) Taylor
Frullania hampeana Nees [1, 2, 94, 249]; syn. fide Hattori (1985)
22.19a Frullania monocera (Taylor) Taylor var. schiffneri (Verd.) S.Hatt.
Frullania acutiloba Mitt. var. schiffneri Verd. [86]
22.20 Frullania neurota Taylor [1, 2, 83, 93, 169, 206, 229, 249, 259]
Frullania aperta Gottsche [44]; nom. nud.; syn. fide Verdoorn (1930)
22.21 Frullania nodulosa (Reinw., Blume & Nees) Nees [1, 2, 169, 249]
Frullania thuilleri Nees [45, 148]; syntype from Sri Lanka; syn. fide Kamimura (1961)
22.22 Frullania orientalis Sande Lac. [87, 91, 99]
(Frullania pallida Gottsche nom. nud. [44]; identity unknown)
22.23 Frullania polyptera Taylor [93, 94, 96]
22.23a Frullania polyptera Taylor var. angustata (Mitt.) S.Hatt. [88, 169]
Frullania angustata Mitt. [148, 227]; syntypes from Sri Lanka. Onraedt (1981) wrongly gave this
as a synonym of Frullania campanulata.
(Frullania poondeloyae Gottsche nom. nud. [44]; identity unknown)
22.24 Frullania ramuligera (Nees) Mont. [83, 93, 96, 169]
22.25 Frullania riojaneirensis (Raddi) Spruce [1, 99, 96, 229]
Frullania galeata (Reinw., Blume & Nees) Dumort. [87, 249]; syn. fide Svihla (19566).
22.26 Frullania serrata Gottsche [84, 93, 169, 206, 249]
Frullania latistipula Steph. [227]; syntype from Sri Lanka; syn fide Verdoorn (1930)
22.26a Frullania serrata Gottsche forma crispulodentata Verd. [1, 2, 91, 99, 248]
(Frullania tamarisci (L.) Dumort. [148, 206]; records from Asia mostly refer to F. moniliata (L.
Söderström, pers. comm.)
Revised checklist to the Liverworts and Hornworts of Sri Lanka
5
22.27 Frullania ternatensis Gottsche [1, 2, 84, 93, 169, 249]
Frullania pendula Mitt. [148]; type from Sri Lanka; syn. fide Hattori (1975b)
Note: Stephani (1898–1925) treated F. pendula twice, as a distinct species and as a synonym of Frullania
ceylanica.
22.28 Frullania trichodes Mitt.
Frullania vethii Sande Lac. [169]; syn. fide Hattori & Piippo (1986)
Herbertaceae
23.1 Herbertus ceylanicus (Steph.) H.A.Miller [1, 2, 80, 127, 146, 169]
Schisma ceylanicum Steph. [227]; type from Sri Lanka.
23.2 Herbertus dicranus (Taylor ex Gottsche, Lindenb. & Nees) Trevis. [126, 127]
Sendtnera dicrana Taylor ex Gottsche, Lindenb. & Nees [148]
Herbertus perrottetii (Steph.) Abeyw. [1, 2, 146]
Schisma perrottetii Steph. [227]; type from Sri Lanka; synonymised by Juslén (2006)
Schisma wichurae Steph. [1, 2, 169, 227]; type from Sri Lanka; synonymised by Juslén (2006);
according to Reimers (1931) the Wichura type specimen was wrongly cited by Stephani as from
China and is from Sri Lanka.
(Herbertus fragilis (Steph.) H.A.Miller [169]. According to Juslén (2006) this is a synonym of H. aduncus
(Dicks.) Gray which is not reported from Sri Lanka; Juslén states that many specimens labelled H.
fragilis belong to H. dicranus).
23.3 Herbertus ramosus (Steph.) H.A.Mill. [127]
Jackiellaceae
24.1 Jackiella ceylanica Schiffn. ex Steph. [1, 2, 169, 227]; type from Sri Lanka.
24.2 Jackiella javanica Schiffn. [1, 2, 169, 178]
Jubulaceae
25.1 Jubula hutchinsiae (Hook.) Dumort. subsp. javanica (Steph.) Verdoorn [37]
Lejeuneaceae
26.1 Acanthocoleus gilvus (Gottsche) Kruijt [33, 138]
Dicranolejeunea gilva (Gottsche) Steph. [171]
27.1 Acrolejeunea emergens (Mitt.) Steph. [48, 61, 169, 171].
(Acrolejeunea fertilis (Reinw., Blume & Nees) Schiffn. [46, 76, 138, 158]
Lejeunea fertilis (Reinw., Blume & Nees) Mitt., hom. illeg. [148];
Ptychocoleus ustulatus (Taylor) Steph. [1, 2, 110]; all records from Sri Lanka are doubtful and may
refer to A. emergens (Gradstein & Inoue, 1980)
27.2 Acrolejeunea planiuscula (Mitt.) Steph. [76]
27.3 Acrolejeunea pycnoclada (Taylor) Schiffn. [46, 48, 169, 171]
Ptychocoleus pycnocladus (Taylor) Steph. [1, 2, 227, 251]
Lejeunea pycnoclada (Taylor) Mitt. [148]
28.1 Caudalejeunea cristiloba (Steph.) Gradst. [47, 169, 171, 267]
28.2 Caudalejeunea reniloba (Gottsche) Steph. [37, 48, 123, 145, 169, 171, 235]
Caudalejeunea lessonii Steph. [169, 171]; syn. fide Zhu & So (2001).
29.1 Cheilolejeunea ceylanica (Gottsche) R.M.Schust. & Kachroo [37, 63, 157, 159, 169, 193, 232, 267,
268]
Cheilolejeunea trapezia (Nees) Mizut. var. ceylanica (Gottsche) A.E.D.Daniels & Kariyappa [32]
Pycnolejeunea ceylanica (Gottsche) Schiffn. [1, 2, 109, 110, 227, 228]
Lejeunea ceylanica Gottsche [45, 148]; type from Sri Lanka.
29.2 Cheilolejeunea decurrens (Sande Lac.) X.-L.He [169]
29.3 Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. [57, 59, 263]
29.4 Cheilolejeunea gardneri (Mitt.) Mizut. [150]
Pycnolejeunea gardneri (Mitt.) Steph. [1, 2, 109, 227, 232]
Lejeunea gardneri Mitt. [148]; type from Sri Lanka.
29.5 Cheilolejeunea imbricata (Nees) S.Hatt. [123, 169]
29.6 Cheilolejeunea intertexta (Lindenb.) Steph. [2, 10, 63, 80, 153, 156, 159, 169]
Cheilolejeunea kurzii Steph. [169]; syn. fide Kitagawa (1981).
29.7 Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. [261]
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Long and Rubasinghe
29.8 Cheilolejeunea lindenbergii (Gottsche) Mizut. [169, 190, 193]
Hygrolejeunea lindenbergii (Gottsche) Steph. [1, 2, 227]
Lejeunea lindenbergii Gottsche [45, 228]; syntype from Sri Lanka.
29.9 Cheilolejeunea longiloba (Steph. ex G.Hoffm.) J.J.Engel & B.C.Tan [169, 267]
29.10 Cheilolejeunea nietneri (Steph.) Mizut. [68, 169]
Harpalejeunea nietneri Steph. [1, 2, 227]; type from Sri Lanka.
(Cheilolejeunea orientalis (Gottsche) Mizut.
Euosmolejeunea orientalis (Gottsche) Steph. [1, 2, 227]; these reports are based on a Nietner
specimen redetermined by Mizutani (1972) as C. trifaria)
29.11 Cheilolejeunea serpentina (Mitt.) Mizut. [10, 63, 150, 169, 231]
Euosmolejeunea serpentina (Mitt.) Steph. [1, 2, 227]
Lejeunea serpentina Mitt. [148]; type from Sri Lanka.
29.12 Cheilolejeunea streimannii Pócs & Ninh [191]
29.13 Cheilolejeunea subopaca (Mitt.) Mizut. [169, 267, 268]
29.14 Cheilolejeunea trapezia (Nees) Mizut. [33]
Pycnolejeunea trapezia (Nees) Steph. [1, 2]
29.15 Cheilolejeunea trifaria (Reinw., Blume & Nees) Mizut. [80, 156, 159]
Euosmolejeunea trifaria (Reinw, Blume & Nees) Spruce [1, 2]
(Cheilolejeunea turgida (Mitt.) W. Ye & R.L. Zhu
Lejeunea turgida Mitt. [148]; syntype from Sri Lanka; this record is erroneous (Grolle & Piippo,
1990)
29.16 Cheilolejeunea vittata (Steph. ex G.Hoffm.) R.M.Schust. & Kachroo [169, 268]
30.1 Cololejeunea amoena Benedix [169]
30.2 Cololejeunea appressa (A.Evans) Benedix [169]
30.3 Cololejeunea ceratilobula (P.C.Chen) R.M.Schust. [9, 33, 267]
30.4 Cololejeunea ceylanica Onr. [9, 33, 37, 168, 169, 264, 267]; type from Sri Lanka.
30.5 Cololejeunea cordiflora Steph. [190]
30.6 Cololejeunea desciscens Steph. [37, 192, 267]
Cololejeunea formosana Mizut. [169]; syn. fide Pócs & Piippo (2011)
30.7 Cololejeunea falcata (Horik.) Benedix [192, 193, 235, 260, 267]
Cololejeunea falcatoides Benedix [145, 169, 234]; syn. fide Zhu & So (2001)]
30.8 Cololejeunea fissilobula Herz. [169 ‘aff’]
30.9 Cololejeunea floccosa (Lehm. & Lindenb.) Steph. [9, 33, 37, 169, 235, 267]
30.9a Cololejeunea floccosa (Lehm. & Lindenb.) Steph. var. aurita Benedix [192, 235, 236]
30.9b Cololejeunea floccosa (Lehm. & Lindenb.) Steph. var. trivitata Tixier [236]
30.10 Cololejeunea fredericii Onr. [166, 169, 238]; type from Sri Lanka
30.11 Cololejeunea gottschei (Steph.) Mizut. [1, 2, 9, 151, 169, 267]
Physocolea gottschei Steph. [227]; syntype from Sri Lanka
Note: this was synonymised under C. longifolia by Tixier (1984, 1985) but maintained by Zhu & So
(2001) and Asthana & Srivastava (2003).
30.12 Cololejeunea gynophthalma Benedix [234]
30.13 Cololejeunea haskarliana (Lehm. & Lindenb.) Steph. [9, 33, 169, 267]
Lejeunea haskarliana Lehm. & Lindenb. [45]
Schiffneriolejeunea haskarliana (Lehm. & Lindenb.) Gradst. [169]
30.14 Cololejeunea hinidumae Onr. [142, 169]; type from Sri Lanka.
30.15 Cololejeunea inflata Steph. [169, 267]
Cololejeunea oshimensis (Horik.) Benedix [235]; syn. fide Zhu & So (2001).
30.16 Cololejeunea inflectens (Mitt.) Benedix ("inflectidens") [1, 2, 12, 169, 193, 260, 267]
Campylolejeunea inflectens (Mitt.) Mizut. [150]
Lejeunea inflectens Mitt. [148]; type from Sri Lanka
Physocolea inflectens (Mitt.) Steph. [227]
30.17 Cololejeunea lanciloba Steph. [33, 37, 169, 267]
Cololejeunea bolombensis (Steph.) Vanden Berghen [169]; syn. fide Zhu & So (2001).
30.18 Cololejeunea leonidens Benedix var. saccata Benedix [169]
Note: C. leonidens was given as a synonym of C. ocelloides (Horik.) S.Hatt. by Zhu & So (2001) .
30.19 Cololejeunea longifolia (Mitt.) Benedix ex Mizut. [9, 23, 237]
30.20 Cololejeunea metzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilkiu-Borges & J. Heinrichs [36]
30.21 Cololejeunea minutissima (Sm.) Schiffn. [169]
Revised checklist to the Liverworts and Hornworts of Sri Lanka
7
30.22 Cololejeunea obliqua (Nees & Mont.) Schiffn.
Cololejeunea scabrifolia Schiffn., nom. inval. [169]; syn. fide Grolle (1995) and Zhu & So (2001) .
30.23 Cololejeunea peraffinis (Schiffn.) Schiffn. [169, 267]
30.24 Cololejeunea perakensis Tixier [235, 238]
Note: not validly published until Tixier (1985).
30.25 Cololejeunea planissima (Mitt.) Abeyw. [1, 2, 9, 33, 149, 160, 169, 192, 238, 267]
Lejeunea planissima Mitt. [148]; type from Sri Lanka
Physcocolea planissima (Mitt.) Steph. [227]
30.26 Cololejeunea pluripunctata Benedix [169]
30.27 Cololejeunea pretiosa Benedix [37]
Note: Tixier (1985) treats this as a synonym of C. hispidissima (Steph.) Pandé, but this is a synonym of C.
haskarliana (Zhu & So, 2001).
30.28 Cololejeunea pseudofloccosa (Horik.) Benedix [33, 169, 192, 234, 267]
Taeniolejeunea pseudofloccosa (Horik.) S.Hatt. [1, 2, 173]
30.29 Cololejeunea schmidtii Steph. [193, 237, 260, 267]
30.30 Cololejeunea serrata (Steph.) Benedix [1, 2, 12, 16, 169, 192, 267]
Physcocolea serrata Steph. [227]; type from Sri Lanka.
30.31 Cololejeunea taprobanea Tixier [238]; type from Sri Lanka
30.32 Cololejeunea tenella Benedix [37, 169, 192, 264, 267]
30.33 Cololejeunea triapiculata (Herzog) Tixier [169, 192, 234, 235, 238]
30.34 Cololejeunea trichomanis (Gottsche) Steph. subsp. cordiflora (Steph.) Pócs [192]
30.35 Cololejeiunea vesicaria (Sande Lac.) Benedix [169]
30.36 Cololejeunea vidaliana Tixier [235, 238, 267]
31.1 Colura acroloba (Mont. ex Steph.) Ast [1, 2, 37, 124, 167, 169, 234, 235, 267]
31.2 Colura ari (Steph.) Steph. [1, 2, 52, 167, 169, 267]
31.3 Colura brevistyla Herzog [103, 124, 167, 169, 193]; type from Sri Lanka
31.4 Colura calyptrifolia (Hook.) Dumort. [167, 267]
31.5 Colura conica (Sande Lac) K.I.Goebel [167, 169]
Colura acutifolia Ast.; syn. fide Zhu & So (2001).
31.6 Colura corynephora (Gottsche, Lindenb. & Nees) Trevis. [167, 169, 267]
31.7 Colura greig-smithii Ast [167]
31.8 Colura javanica Steph. [103]
31.9 Colura meijeri Ast [37]
31.10 Colura ornata K.I.Goebel [167, 169]
31.11 Colura tenuicornis (A.Evans) Steph.
Colura calyptrifolia (Hook.) Dumort. subsp. tenuicornis (A.Evans) Vanden Berghen [169]
31.12 Colura verdoornii Herzog & Ast [37]
(Dicranolejeunea ceylanica Steph. nom. nud. [24]; original material from Sri Lanka; this belongs to
Acrolejeunea but species uncertain (Mizutani, 1980a)).
32.1 Diplasiolejeunea cavifolia Steph. [37]
32.2 Diplasiolejeunea onraedtii Grolle [60, 169]; type from Sri Lanka.
32.3 Diplasiolejeunea rudolphiana Steph. [169, 267]
33.1 Drepanolejeunea angustifolia (Mitt.) Grolle [33, 34, 169, 267]
33.2 Drepanolejeunea fissicornua Steph. [104]
33.3 Drepanolejeunea fleischeri (Steph.) Grolle & R.L.Zhu [8, 33, 34, 75, 267]
Leptolejeunea fleischeri Steph. [1, 2, 227]; type from Sri Lanka
Rhaphidolejeunea fleischeri (Steph.) Herzog [17, 18, 108].
33.4 Drepanolejeunea macrodonta (Mitt.) Steph. [1, 2, 227]
Lejeunea macrodonta Mitt. [148]; type from Sri Lanka.
33.5 Drepanolejeunea pentadactyla (Mont.) Steph.
Drepanolejeunea micholitzii Steph. [169]; syn. fide Piippo (1990).
Drepanolejeunea micholitzii Steph. var. dactylophoroides Herzog [235]
33.6 Drepanolejeunea tenera K.I.Goebel [169]
33.7 Drepanolejeunea ternatensis (Gottsche) Schiffn. [34, 169, 267]
33.8 Drepanolejeunea teysmannii Steph. [37]
33.9 Drepanolejeunea thwaitesiana (Mitt.) Steph. [1, 2, 104, 105, 123, 145, 164, 169, 227, 235]
Lejeunea thwaitesiana Mitt. [148]; type from Sri Lanka
Ceratolejeunea thwaitesiana (Mitt.) Steph. [1, 2, 169, 227, 235]
8
Long and Rubasinghe
Note: Lejeunea thwaitesiana Mitt. was treated twice by Stephani (1898–1925) under both
Drepanolejeunea and Ceratolejeunea.
33.9a Drepanolejeunea thwaitesiana (Mitt.) Steph. var. zhengii R.L.Zhu [267]
33.10 Drepanolejeunea vesiculosa (Mitt.) Steph. [1, 2, 34, 106, 169, 227, 267]
Lejeunea vesiculosa Mitt. [148]; type from Sri Lanka
34.1 Harpalejeunea filicuspis (Steph.) Mizut. [62, 68, 169]
35.1 Lejeunea anisophylla Mont. [33, 191, 267]
Lejeunea borneensis Steph. [169]; syn. fide Zhu & So (2001).
35.2 Lejeunea apiculata Sande Lac [33, 193]
Stenolejeunea apiculata (Sande Lac.) R.M.Schust. [169, 267]
Taxilejeunea apiculata (Sande Lac.) Eifrig nom. illeg. [169]
35.3 Lejeunea cocoes Mitt. [1, 2, 16, 143, 148, 150, 193, 217, 227, 267]; type from Sri Lanka.
35.4 Lejeunea dipterota Eifrig [234]
35.5 Lejeunea discreta Lindenb. [33, 155, 169, 267]
Hygrolejeunea discreta (Lindenb.) Steph. [1, 2, 38, 234]
35.6 Lejeunea flava (Sw.) Nees [33, 169]
35.7 Lejeunea flavida Mitt. [148, 169]; type from Sri Lanka
Pycnolejeunea flavida (Mitt.) Steph. [1, 2, 109, 150, 227].
(Lejeunea grandiloba Gottsche, nom. nud. [44]; identity unknown)
35.8 Lejeunea herzogii Mizut. [169]
35.9 Lejeunea lowriana Steph. [169]
35.10 Lejeunea micholitzii Mizut. [193, 235]
35.11 Lejeunea neelgherriana Gottsche [37, 265, 267]
Harpalejeunea indica Steph. [234]; syn. fide Grolle & Reiner-Drehwald (1999).
35.12 Lejeunea nietneri (Steph.) Steph [1, 2, 17]
Eulejeunea nietneri Steph. [226]; type from Sri Lanka
? Lejeunea nietneri Gottsche nom. nud. [44]; original material from Sri Lanka.
35.13 Lejeunea obfusca Mitt. [1, 2, 148, 150, 169, 227, 267]; type from Sri Lanka.
35.14 Lejeunea obscura Mitt. [33, 169, 267]
35.15 Lejeunea papilionacea Steph. [33]
35.16 Lejeunea riparia Mitt. [148, 150, 196]; type from Sri Lanka
Cheilolejeunea riparia (Mitt.) M.E.Reiner
Rectolejeunea riparia (Mitt.) Steph. [1, 2, 227]
35.17 Lejeunea subacuta Mitt. [33, 150, 155]
35.18 Lejeunea tuberculosa Steph. [143]
35.19 Lejeunea utriculata (Steph.) Mizut.
Pycnolejeunea utriculata Steph. [1, 2, 227]
35.20 Lejeunea wightii Lindenb. [80]
36.1 Lepidolejeunea bidentula (J.B.Jack & Steph.) R.M.Schust. [267]
Pycnolejeunea badia Steph. [169]; syn. fide Zhu & So (2001)
37.1 Leptolejeunea elliptica (Lehm. & Lindenb.) Schiffn. [33, 169]
Leptolejeunea subacuta Steph. [235]; syn. fide Zhu & So (2001)
37.2 Leptolejeunea epiphylla (Mitt.) Steph. [1, 2, 107, 123, 169, 227, 234, 235, 267]
Lejeunea epiphylla Mitt. [148]; type from Sri Lanka
37.3 Leptolejeunea maculata (Mitt.) Schiffn. [1, 2, 37, 56, 107, 123, 169, 227, 267]
Lejeunea maculata Mitt. [148]; type from Sri Lanka.
37.4 Leptolejeunea massartiana Schiffn. ex Herzog [235]
37.5 Leptolejeunea rhombifolia Steph. [235]
37.6 Leptolejeunea vitrea (Nees) Schiffn. [169, 190, 234, 235]
38.1 Leucolejeunea xanthocarpa (Lehm. & Lindenb.) A.Evans [1, 2, 149, 169, 171, 195, 251]
Lejeunea xanthocarpa (Lehm. & Lindenb.) Lehm. & Lindenb. [44]
39.1 Lopholejeunea applanata (Reinw., Blume & Nees) Schiffn. [76, 262]
Lejeunea applanata (Reinw., Blume & Nees) Nees [148] 'adplanata'.
39.2 Lopholejeunea nigricans (Lindenb.) Steph.
Lopholejeunea arbortiva (Mitt.) Steph. [171]; syn. fide Zhu & Gradstein (2005).
39.3 Lopholejeunea ceylanica Steph. [1, 2, 37, 48, 76, 161, 169, 171, 227, 230, 251, 262]; syntype and
lectotype from Sri Lanka.
Lopholejeunea longiloba Steph. [227] [251]; type from Sri Lanka; syn. fide Zhu & Gradstein
Revised checklist to the Liverworts and Hornworts of Sri Lanka
9
(2005)
? Phragmicoma ceylanica Gottsche nom. nud. [44]
39.4 Lopholejeunea eulopha (Taylor) Schiffn. [76, 262]
39.5 Lopholejeunea javanica (Nees) Schiffn. [161, 266]
39.6 Lopholejeunea subfusca (Nees) Schiffn. [1, 2, 48, 52, 76, 80, 149, 169, 171, 227, 234, 250, 251, 262]
Lejeunea subfusca (Nees) Nees & Mont. [148]
39.7 Lopholejeunea zollingeri (Steph.) Schiffn. [76, 262]
Lopholejeunea nipponica Horik. [169, 171]; syn. fide Zhu & Gradstein (2005)
40.1 Mastigolejeunea auriculata (Wilson) Schiffn. [48]
40.2 Mastigolejeunea humilis (Gottsche) Schiffn. [1, 2, 48, 98, 169, 171, 250, 251]
40.2a Mastigolejeunea humilis (Gottsche) Schiffn. var. ciliata Awasthi & Udar [11]
40.3 Mastigolejeunea repleta (Taylor) A.Evans [2, 48, 98, 133]
Lejeunea repleta Taylor [148]
Mastigolejeunea atypos (Schiffn.) Steph. [1, 227]; syn. fide Gradstein & Inoue (1980).
40.4 Mastigolejeunea virens (Ångstr.) Steph. [76, 162]
41.1 Metalejeunea cucullata (Reinw., Blume & Nees) Grolle
Microlejeunea cucullata (Reinw., Blume & Nees) J.B.Jack & Steph. [169, 235]
(Microlejeunea lunulatilobula Horik. was reported by Miller et al. (1983) but no source was given for the
record)
42.1 Microlejeunea punctiformis (Taylor) Spruce [1, 2, 33, 110, 191, 227]
Lejeunea punctiformis Taylor [80, 98, 149, 150, 267]
Lejeunea gracillima Mitt. [148]; type from Sri Lanka; syn. fide Mizutani (1963)
Microlejeunea gracillima (Mitt.) Carringt. & Pearson [1, 2, 21, 227]
43.1 Ptychanthus striatus (Lehm. & Lindenb.) Nees [2, 33, 48, 76, 169, 251]
Ptychanthus striatus (Lehm. & Lindenb.) Nees var. retusus (Reinw., Blume & Nees) Verd. [1, 2];
syn. fide Haerida et al. (2010)
Ptychanthus birmensis Steph. [1, 2]; syn. fide Verdoorn (1934)
Ptychanthus effusus Steph. [1, 2, 227]; syntype from Sri Lanka; syn. fide Verdoorn (1934)
Ptychanthus nietneri (Steph.) Schiffn. [1, 2, 227]; type from Sri Lanka; syn. fide Verdoorn (1934)
Ptycholejeunea nietneri Steph. [226]; type from Sri Lanka; syn. fide Verdoorn (1934)
? Phragmicoma nietneri Gottsche nom. nud. [44] is probably a synonym.
44.1 Pycnolejeunea grandiocellata Steph. [100, 169]
45.1 Schiffneriolejeunea polycarpa (Nees) Gradst. [48, 169, 171]
45.2 Schiffneriolejeunea pulopenangensis (Gottsche) Gradst. [48, 169, 171]
Ptychocoleus pulopenangensis (Gottsche) Trevis. [1, 2, 250, 251]
45.3 Schiffneriolejeunea tumida (Nees) Gradst. [48, 49, 169, 171, 230]
Ptychocoleus peradeniensis (Mitt.) Steph. [1, 2, 227, 250, 251]; syn. fide Gradstein & Inoue (1980)
Lejeunea peradeniensis Mitt. [148]; type from Sri Lanka
Ptychocoleus cumingianus auct. non (Mont.) Trevis. [1, 2]; syn. fide Gradstein & Inoue (1980)
45.3a Schiffneriolejeunea tumida (Nees) Gradst. var. hasskarliana (Gottsche) Gradst. & Terken [49, 171]
Schiffneriolejeunea hasskarliana (Gottsche) Gradst. [169]
46.1 Spruceanthus marianus (Gottsche) Mizut. [48, 169, 171]
Archilejeunea mariana (Gottsche) Steph. [1, 2]
46.2 Spruceanthus semirepandus (Nees) Verd. [1, 2, 48, 76, 169, 171, 251]
Lejeunea semirepanda (Nees) Nees & Mont. [148]
47.1 Taxilejeunea compressiuscula Lindenb. ex Steph. [169]
48.1 Thysananthus convolutus Lindenb. [37]
48.2 Thysananthus spathulistipus (Reinw., Blume & Nees) Lindenb. [1, 2, 48, 169, 171, 251]
Lejeunea spathulistipa (Reinw., Blume & Nees) Mitt. [148]
49.1 Trocholejeunea infuscata (Mitt.) Verd. [1, 2, 33, 48, 163, 251]
49.2 Trocholejenea sandvicensis (Gottsche) Mizut. [33, 48, 169, 171]
Brachiolejeunea sandvicensis (Gottsche) A.Evans [1, 2]
50.1 Xenolejeunea longiloba (Steph. ex G.Hoffm.) Kachroo & R.M.Schust. [235]
Lepicoleaceae
(Lepicolea ochroleuca (Spreng.) Spruce [1, 2]
Sendtnera ochroleuca (Spreng.) Nees [148]; records from Sri Lanka are based on misidentifications
of L. yakusimensis fide Scott (1960)).
10
Long and Rubasinghe
(Lepicolea scolopendra (Hook.) Dumort. [1, 2]; records from Sri Lanka are based on misidentifications of
L. yakusimensis fide Scott (1960)).
51.1 Lepicolea yakusimensis (S.Hatt.) S.Hatt. [80, 102, 117, 134, 169, 204].
Lepidoziaceae
52.1 Bazzania callida (Steph.) Abeyw. [1, 130, 134, 169]
Mastigobryum callidum Steph. [2, 224, 227]
Mastigobryum intermedium Mitt. hom. illeg. [148, 227]; original material from Sri Lanka; syn. fide
Stephani (1885).
52.2 Bazzania ceylanica (Mitt.) W.E.Nicholson [1, 2, 123, 152, 169]
Mastigobryum ceylanicum Mitt. [148, 227]; type from Sri Lanka
Bazzania grandistipula (Steph.) Abeyw. [1, 2]
Mastigobryum grandistipulum Steph. [227]; type from Sri Lanka; syn. fide Mizutani (1967a).
Bazzania lacerostipula (Steph.) Abeyw. [1, 2]
Mastigobryum lacerostipulum Steph. [227]; type from Sri Lanka; syn. fide Mizutani (1967a)
Bazzania stricta (Steph.) Abeyw. [1, 2]
Mastigobryum strictum Steph. [225, 227]; syn. fide Mizutani (1967a)
52.3 Bazzania erosa (Nees) Trevis. [134]
52.4 Bazzania fleischeri (Steph.) Abeyw. [1, 2, 152, 169]
Mastigobryum fleischeri Steph. [227]; syntype from Sri Lanka.
52.5 Bazzania gaudichaudii (Gottsche ex Steph.) Schiffn. [1, 2]
Mastigobryum gaudichaudii Gottsche ex Steph. [227]
52.6 Bazzania japonica (Sande Lac.) Lindb. [169]
52.7 Bazzania obtusa D.G.Long & Rubasinghe, nom. nov.
Mastigobryum obtusatum Mitt. [148, 227]; type from Sri Lanka
Bazzania obtusata (Mitt.) Abeyw., hom. illeg. non Bazzania obtusata (Taylor) Kuntze [1, 2, 80,
169]; see Appendix I for formal treatment.
52.8 Bazzania oshimensis (Steph.) Horik. [165, 169, 205]
52.9 Bazzania ovistipula (Steph.) Abeyw. [1, 2, 132, 134, 152, 165, 169, 205]
Mastigobryum ovistipulum Steph. [227]; type from Sri Lanka.
52.10 Bazzania pearsonii Steph. [152, 165, 205]
Bazzania rotundistipula (Steph.) Abeyw. [1, 2, 130]
Mastigobryum rotundistipulum Steph. [227]; syntype from Sri Lanka; syn. fide Mizutani (1967a)
Mastigobryum wichurae Steph. [227]; type from Sri Lanka; syn. fide Mizutani (1967a).
Pleuroschisma wichurae (Steph.) Reimers [195]; according to Reimers (1931) the Wichura type
specimen, wrongly cited by Stephani from China, is from Sri Lanka.
52.11 Bazzania praerupta (Reinw., Blume & Nees) Trevis. [2, 130, 152, 165, 169, 189, 205]
Mastigobryum praeruptum Reinw., Blume & Nees [148]
Bazzania decurva (Nees) Kuntze [1, 2]; syn. fide Onraedt (1981).
Mastigobryum decurvum Nees [45, 148, 227]; syntype from Sri Lanka.
Bazzania sandei (Steph.) Abeyw. [1, 2]
Mastigobryum sandei Steph. [227]; syn. fide Onraedt (1981).
52.12 Bazzania rabenhorstii (Steph.) Abeyw. [1, 2]
Mastigobryum rabenhorstii Steph. [227]; type from Sri Lanka.
52.13 Bazzania spiralis (Reinw., Blume & Nees) Meijer [169]
52.14 Bazzania tridens (Reinw., Blume & Nees) Trevis. [80, 152, 169, 189, 205]
Bazzania oblonga (Mitt.) Schiffn. [1, 2]; syn. fide Onraedt (1981).
52.15 Bazzania vittata (Gottsche) Trevis. [1, 2, 169]
Mastigobryum vittatum Gottsche [148, 227]
53.1 Hygrolembidium boschianum (Sande Lac.) R.M.Schust. [2, 121, 169, 184, 203]
54.1 Lepidozia plumula Herz. [103]; type from Sri Lanka.
54.2 Lepidozia subintegra Lindenb. [154]
55.1 Neolepidozia wallichiana (Gottsche) Fulford & J.Taylor
Telaranea wallichiana (Gottsche) R.M.Schust. [39]
Lepidozia wallichiana Gottsche [1, 2, 37, 148, 169, 177, 205]
56.1 Telaranea major (Herz.) J.J.Engel & G.L.Merr. [39]
Arachniopsis major Herz. [37, 180]
Revised checklist to the Liverworts and Hornworts of Sri Lanka
11
57.1 Tricholepidozia neesii (Lindenb.) E.D.Cooper
Lepidozia neesii Lindenb. [1, 2, 227]
57.2 Tricholepidozia semperiana (Steph.) E.D.Cooper
Telaranea semperiana (Steph.) Del Ros. [39, 169]
Lepidozia semperiana Steph. [1, 2, 227]; syntype from Sri Lanka.
Lophocoleaceae
58.1 Cryptolophocolea ciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler & Váňa
Lophocolea ciliolata (Nees) Gottsche [1, 2, 148, 169, 179, 200, 227]
Chiloscyphus ciliolatus (Nees) J.J.Engel & R.M.Schust. [102]
58.2 Cryptolophocolea fleischeri (Steph.) L.Söderstr. [218]
Lophocolea fleischeri Steph. [1, 2, 227]; type from Sri Lanka
Chiloscyphus ceylonensis J.J.Engel, nom. nov. for Lophocolea fleischeri; type from Sri Lanka
59.1 Heteroscyphus argutus (Reinw., Blume & Nees) Schiffn. [169, 179, 186]
Chiloscyphus argutus (Reinw., Blume & Nees) Nees [80]
59.2 Heteroscyphus fleischeri (Steph.) D.G.Long & Rubasinghe, comb. nov.
Chiloscyphus fleischeri Steph. [1, 2, 227]; type from Sri Lanka; see Appendix I for formal
treatment.
59.3 Heteroscyphus perfoliatus (Mont.) Schiffn.
Chiloscyphus perfoliatus (Mont.) Nees [148, 169]
(Heteroscyphus splendens (Lehm. & Lindenb.) Grolle
Chiloscyphus decurrens (Reinw., Blume & Nees) Nees [1, 2, 123]; syn. fide Piippo (1990); the
cited source of this report (Stephani, 1898–1925) does not contain any record from Sri Lanka.)
59.4 Heteroscyphus tener (Steph.) Schiffn. [51, 187, 219]
59.5 Heteroscyphus tridentatus (Sande Lac.) Grolle [37]
60.1 Lophocolea bidentata (L.) Dumort. sens.lat. [50]
Lophocolea cuspidata (Nees) Limpr. [169]; syn. fide Paton (1999)
60.2 Lophocolea javanica Schiffn. [169, 179]
60.3 Lophocolea kurzii Sande Lac. [1, 2, 133, 169]
Chiloscyphus kurzii (Sande Lac.) J.J.Engel & R.M.Schust. [186]
Lophocolea peradeniensis Steph. [1, 227]; type from Sri Lanka; syn. fide Kitagawa (1973).
60.4 Lophocolea muricata (Lehm.) Nees [169]
(Saccogyna viticulosa (Sm.) Dumort. [1]; a north temperate species known from Europe and Macaronesia
(Paton, 1999); this report is considered to be erroneous.)
Lophoziaceae s.str.
61.1 Andrewsianthus puniceus (Nees) R.M. Schust. ex Grolle [2, 121, 132, 134, 169]
Lunulariaceae
62.1 Lunularia cruciata (L.) Dumort. [1, 2, 169, 199]
Marchantiaceae
63.1 Marchantia acaulis Steph. [20]
63.2 Marchantia emarginata Reinw., Blume & Nees subsp. emarginata [20, 22]
Marchantia amboinensis Nees & Mont. [1, 2, 169, 199]; syn. fide Bischler (1989)
Marchantia palmata Nees [1, 2, 80, 169, 199]; syn. fide Bischler (1989)
(Marchantia geminata Reinw., Blume & Nees [1, 2, 128]; records probably refer to M. acaulis fide
Bischler (1989))
(Marchantia linearis Lehm. & Lindenb. [1, 2, 128, 148]; records from Sri Lanka are considered to be M.
papillata subsp. grossibarba by Bischler (1989))
63.3 Marchantia paleacea Bertol. [199]; not confirmed from Sri Lanka by Bischler (1989).
63.4 Marchantia papillata Raddi subsp. grossibarba (Steph.) Bischl. [20]
63.5 Marchantia pappeana Lehm. subsp. robusta (Steph.) Bischl. [20]
Marchantia robusta Steph. [23]; type from Sri Lanka (Bischler, 1989)], though type locality
erroneously reported as Java by Stephani in Bonner (1953).
63.6 Marchantia polymorpha L. [1, 2, 128]; not confirmed from Sri Lanka by Bischler (1989) and records
not assigned to subspecies.
12
Long and Rubasinghe
Mastigophoraceae
64.1 Mastigophora aequifolia Steph. [1, 2, 227]; type from Sri Lanka.
64.2 Mastigophora diclados (Brid. ex F.Weber) Nees [1, 2, 37, 98, 169]
Sendtnera diclados (Brid. ex F.Weber) Gottsche, Lindenb. & Nees [148]
Mesoptychiaceae
65.1 Liochlaena subulata (A. Evans) Schljakov
Jungermannia subulata A. Evans [74, 134, 169, 242]
Jungermannia leiantha Grolle subsp. stephanii (Amak.) S.Hatt. [80]; syn. fide Váňa (1973).
Metzgeriaceae
(Metzgeria albinea Spruce [140, 182]; reports of this neotropical species probably refer to M. foliicola (So,
2002)).
66.1 Metzgeria ciliata Raddi [30]
Metzgeria decipiens (C.Massal.) Schiffn. & Gottsche [2, 140, 142, 169, 182]; syn. fide Costa
(2008).
66.2 Metzgeria conjugata Lindb. [30]
Metzgeria lindbergii Schiffn. [33, 142, 169, 210]; syn. fide Costa (2008).
66.3 Metzgeria consanguinea Schiffn. [30, 33, 139, 141, 142, 182]
66.4 Metzgeria crassipilis (Lindb.) A.Evans [210];
66.5 Metzgeria foliicola Schiffn. [142, 169]; syn. fide Kuwahara (1984a).
(Metzgeria furcata (L.) Dumort. [33, 142, 169, 199]; records from Sri Lanka probably refer to M. ciliata
(Costa, 2008)).
66.6 Metzgeria latifrons Steph. [1, 2, 128, 227]; syntype from Sri Lanka
66.7 Metzgeria lechleri Steph. [30]
Metzgeria vivipara A.Evans [142, 169]; syn. fide Costa (2008).
66.8 Metzgeria leptoneura Spruce [210]
Metzgeria borneensis Kuwah. [141]; syn. fide So (2003b).
Metzgeria subhamata S.Hatt. [2, 121, 145]; syn. fide So (2003b).
Metzgeria luzonensis Kuwah. [142, 169]; syn. fide So (2003b).
Metzgeria papulosa Steph. [1, 2]; syn. fide So (2003b).
Notoscyphaceae
67.1 Notoscyphus lutescens (Lehm. & Lindenb.) Mitt. [80]
Gymnanthe lutescens (Lehm. & Lindenb.) Mitt. [148]
Notoscyphus paroicus Schiffn. [1, 2, 169, 227, 239]; syn. fide Váňa & Long (2009).
Pallaviciniaceae
68.1 Jensenia decipiens (Mitt.) Grolle [66, 128]
Pallavicinia decipiens (Mitt.) Steph. [1, 2, 227]
Steetzia decipiens Mitt. [148]; type from Sri Lanka.
69.1 Pallavicinia indica Schiffn. [2, 128]
69.2 Pallavicinia lyellii (Hook.) Gray [1, 2, 128, 169, 199]
Steetzia lyellii (Hook.) Lehm. [148]
Plagiochilaceae
(Plagiochila acuta Steph. [1, 2, 227]; these records are based on a misidentification of the Sri Lankan
syntype collected by Nietner which is P. microdonta (Inoue, 1979)).
70.1 Plagiochila arbuscula (Lehm. & Lindenb.) Lindenb
Plagiochila belangeriana Lindenb. [80]; syn. fide Inoue (1984).
70.2 Plagiochila bantamensis (Reinw., Blume & Nees) Mont. [71, 120, 207]
Plagiochila lobulata Schiffn. [80, 118, 169]; syn. fide Inoue (1984)
Plagiochila onraedtii Inoue [118, 169]; type from Sri Lanka; syn. fide So & Grolle (2000a)
70.3 Plagiochila ceylanica Mitt. [1, 2, 118, 148, 169, 227]; syntypes and lectotype (Inoue, 1979) from Sri
Lanka.
Plagiochila macrantha Steph. [1, 2, 227]; type from Sri Lanka; syn. fide Inoue (1979)
Plagiochila neutiana Steph. [1, 2, 116, 227]; type from Sri Lanka; syn. fide Inoue (1979).
70.4 Plagiochila devexa Steph. [194, 207, 215]
Revised checklist to the Liverworts and Hornworts of Sri Lanka
13
(Plagiochila elegans Mitt. [194]; this report cites a specimen from Sri Lanka in Mitten's herbarium (NY)
but Mitten (1860–1861) did not mention any such specimen; a Sino-Himalayan species.)
70.5 Plagiochila exigua (Taylor) Taylor [207]
70.6 Plagiochila flexuosa Mitt. [72, 80, 118, 120, 169, 194, 207]
Plagiochila rufa Steph. [1, 2, 227]; type from Sri Lanka; syn. fide Inoue (1979).
70.7 Plagiochila frondescens (Nees) Lindenb. [1, 2, 118, 120, 181, 227]
70.8 Plagiochila ghatiensis Steph. [207]
Plagiochila acutiloba Inoue [118, 169]; syn. fide Grolle & So (1998).
Tylimanthus indicus Steph.; syn. fide Inoue (1979).
70.9 Plagiochila gracilis Lindenb. & Gottsche [80, 120, 194, 207]
Plagiochila firma Mitt. [118, 169]; syn. fide Inoue (1984)
70.10 Plagiochila gymnoclada Sande Lac. [118, 120, 181]
70.11 Plagiochila integrilobula Inoue [118, 120, 169, 191]
(Plagiochila javanica (Sw.) Nees & Mont. [1, 2, 227]; records should be referred to P. neutiana (Inoue,
1969) which is a synonym of P. ceylanica)
70.12 Plagiochila khasiana Mitt. [73, 148, 194, 207]; syntype from Sri Lanka
70.13 Plagiochila kurzii Steph. [207]
Plagiochila meijeri Inoue [118]; type from Sri Lanka; syn. fide So (2001).
(Plagiochila liebmanniana Lehm. & Lindenb. [1, 35, 227]; these reports were rejected as
misidentifications by Inoue (1979), specimens are P. flexuosa)
70.14 Plagiochila microdonta Mitt. [1, 2, 118, 148, 169, 214, 227]; syntype from Sri Lanka.
Plagiochila aciculifera Steph. [1, 2, 227]; type from Sri Lanka; syn. fide Inoue (1979)
(Plagiochila microphylla Steph. Inoue (1981) gave P. wichurae as a synonym of P. microphylla but it
belongs to P. renitens).
(Plagiochila monticola Gottsche nom. nud. non Schiffn. [44]; identity unknown)
(Plagiochila nobilis Gottsche [1, 2, 227]; record doubtful fide Inoue (1979) and not recorded from Sri
Lanka by Inoue (1984).)
70.15 Plagiochila parvifolia Lindenb. [194]
Plagiochila ventricosa Steph. [1, 120, 207, 227]; syntype from Sri Lanka (Inoue, 1979); syn. fide
So & Grolle (2000)
Plagiochila pseudoventricosa Inoue [2, 113, 118, 169]; type from Sri Lanka; syn. fide Inoue
(1984).]
70.16 Plagiochila peradenyensis Schiffn. [1, 2, 70, 73, 118, 201, 216, 227]; type from Sri Lanka.
Plagiochila ramenticaulis Steph. [1, 2, 227]; type from Sri Lanka; syn. fide Inoue (1979)
70.17 Plagiochila renitens (Nees) Lindenb. [181, 212]
Plagiochila aequalis Mitt. [1, 35, 118, 148, 169, 227]; type from Sri Lanka; syn. fide Inoue (1984)
Plagiochila wichurae Steph. [119]; type from Sri Lanka (Reimers, 1931) though wrongly cited by
Stephani as China; syn. fide So & Grolle (2000a).
70.18 Plagiochila sciophila Nees ex Lindenb. [207, 212]
Plagiochila acanthophylla Gottsche [118, 169]; syn. fide Inoue (1984)
Plagiochila cadens Inoue [118, 120, 169]; type from Sri Lanka; syn. fide So & Grolle (1999,
2000a).
(Plagiochila semidecurrens Lehm. & Lindenb. [1, 2, 227, 194]; this Sino-Himalayan species was recorded
from Sri Lanka as var. major Lindenb.; the specimen may have originated from Hooker (Gottsche
et al., 1844; Inoue, 1965a); records are considered doubtful (Inoue, 1979) as it has not been recollected in Sri Lanka. Inoue (1968) placed P. aequalis as a synonym of P. semidecurrens but later
(Inoue, 1984) transferred it to P. renitens).
70.19 Plagiochila sisparensis Steph. [72]
Plagiochila parvidentata Inoue [37, 118, 169]; type from Sri Lanka; syn. fide Grolle & So (1999b)
(Plagiochila spinulosa (Dicks.) Dumort. [199]; a European and Macaronesian species not known in Asia
(So & Grolle, 2000a); identification considered erroneous)
(Plagiochila wightii Lindenb. [148]; record rejected by So & Grolle (2001))
71.1 Plagiochilion braunianum (Nees) S.Hatt. [2, 111, 118, 169, 181]
Plagiochila brauniana Nees [1, 110, 148, 200, 227]
71.2 Plagiochilion oppositum (Reinw., Blume & Nees) S.Hatt. [2, 80, 111, 118, 169, 181, 193]
Plagiochila opposita (Reinw., Blume & Nees) Lindenb. [1, 35, 44, 110, 148, 195, 227];
Plagiochila zygophylla Taylor [45]; syn. fide So & Grolle (2000a)
Note: a Wichura specimen from Sri Lanka was erroneously given as China by Stephani (1898–1925).
14
Long and Rubasinghe
Pleuroziaceae
72.1 Pleurozia acinosa (Mitt.) Steph. [1, 2, 195, 227, 233]
Physiotium acinosum Mitt. [122, 148]; syntypes and lectotype from Sri Lanka (Thiers, 1993).
72.2 Pleurozia gigantea (F.Weber) Lindb. [1, 2, 52, 98, 169, 176, 195, 233]; the Wichura specimen from
Sri Lanka was erroneously given as China by Stephani (1898–1925) according to Reimers (1931).
Physotium giganteum F.Weber [122]
Physotium sphagnoides A.Rich. ex Nees, nom. illeg. [148]
72.3 Pleurozia subinflata (Austin) Austin [233]
Pleurozia microcarpa (J.B.Jack) Steph. [1, 2, 227]
Physotium microcarpum J.B.Jack [122]; type from Sri Lanka; syn. fide Thiers (1993).
Porellaceae
73.1 Porella acutifolia (Lehm. & Lindenb.) Trevis. [81, 92, 169, 176, 188]
Madotheca acutifolia Lehm. & Lindenb. [1, 2, 148, 195, 227]
73.2a Porella caespitans Steph. subsp. latior (S.Hatt.) S.Hatt. [90]
Porella acutifolia (Lehm. & Lindenb.) Trevis. subsp. latior S.Hatt. [81, 92]; type from Sri Lanka
73.2b Porella caespitans (Steph.) S.Hatt. subsp. latior (S.Hatt.) S.Hatt. var. intermedia (S.Hatt.) S.Hatt.
[169]
Porella acutifolia (Lehm. & Lindenb.) Trevis. subsp. latior S.Hatt. var. intermedia S.Hatt. [81, 92]
73.3 Porella japonica (Sande Lac.) Mitt. [169].
73.4 Porella madgascariensis (Nees & Mont.) Trevis. [79, 81, 92, 169, 188]
Madotheca nilgheriensis Mont. [1, 2]; syn. fide Hattori (1967)
73.5 Porella perottetiana (Mont.) Trevis. [92, 169, 188]
Madotheca perrottetiana Mont. [1, 2, 110, 227]
Madotheca ciliaris Nees [148]; syn. fide Stephani (1898–1925).
Radulaceae
74.1 Radula acuminata Steph. [33, 37, 169, 170, 193, 234]
74.2 Radula amentulosa Mitt. [169, 170, 252, 255, 257, 258]
74.3 Radula assamica Steph. [33, 170, 191, 252, 255, 267]
74.4 Radula ceylanica K.Yamada [169, 170, 252, 255]; type from Sri Lanka.
74.5 Radula cordata Mitt. [37, 227]; earlier reports of this species [1, 2, 252] were based on a specimen of
R. kurzii fide Yamada (1979a).
74.6 Radula formosa (Meissn.) Nees [1, 2, 26, 29, 148, 169, 170, 227, 252, 255, 257, 258]
74.7 Radula javanica Gottsche [1, 2, 37, 45, 148, 169, 170, 197, 211, 252, 255, 258]; syntype from Sri
Lanka
Radula nietneri Steph. [1, 2, 227]; type from Sri Lanka; syn. fide Yamada (1979a)
Radula colliculosa Mitt. [29, 148]; syntypes from Sri Lanka; syn. fide Onraedt (1983).
74.8 Radula kurzii Steph. [28, 29, 37, 123, 169, 170, 252, 255]
Radula speciosa Gottsche ex Steph. [1, 2, 44, 223, 227, 252a]; type from Sri Lanka; syn. fide
Verdoorn (1934).
74.9 Radula madagascariensis Gottsche [169, 170, 252, 255, 258]
74.10 Radula nymanii Steph. [169, 170]
74.11 Radula obscura Mitt. [33, 170, 252, 255]
74.12 Radula onraedtii K.Yamada [169, 170, 256]; type from Sri Lanka.
74.13 Radula retroflexa Taylor [169, 170, 252, 255, 258]
74.14 Radula sandei Steph. [29, 252b]
74.15 Radula subpallens Steph. [1, 2, 27, 29, 170, 227, 255, 258]; syntype from Sri Lanka.
74.16 Radula tabularis Steph. [170]
74.17 Radula tjibodensis K.I.Goebel [33, 169, 170]
74.18 Radula ventricosa Steph. [27, 29, 170, 255, 258]
74.19 Radula yangii K.Yamada [169, 170, 255, 257]
Radula pinnulata Yang nom. illeg. fide Yamada (1979a).
Ricciaceae
75.1 Riccia attenuata Pandé [2, 128]
75.2 Riccia billardieri Mont. & Nees [1, 2, 125, 128, 174]
73.3 Riccia crispatula Mitt. [1, 2, 128, 148, 174, 220, 227]; type from Sri Lanka.
Revised checklist to the Liverworts and Hornworts of Sri Lanka
15
75.4 Riccia discolor Lehm. & Lindenb. [2, 128, 169]
75.5 Riccia huebeneriana Lindenb. [37]
75.6 Riccia sorocarpa Bisch. [169]
Scapaniaceae
76.1 Diplophyllum nanum Herzog [169, 178, 222]
77.1 Scapania lepida Mitt. [1, 2, 148, 169, 227]; type from Sri Lanka.
Schistochilaceae
78.1 Schistochila aligera (Nees & Blume) J.B.Jack & Steph. [1, 2, 110, 123, 169, 176, 209, 227]
Gottschea aligera Nees & Blume [43, 148]
Schistochila commutata Steph. [227]; syntype from Sri Lanka; syn. fide So (2003a)
Schistochila fleischeri Steph. [1, 2, 227]; type from Sri Lanka; syn. fide So (2003a)
Schistochila philippinensis (Mont.) J.B.Jack & Steph. [1, 2, 25, 101, 169, 176]; syn. fide So (2003a)
Gottschea philippinensis Mont. [43]
Note: S. aligera and S. philippinensis are sometimes kept separate (Gao & Wu, 2004).
(Schistochila neesii (Mont.) Trevis. [1, 2, 25]; reports from Sri Lanka are based on specimens of S. aligera
fide So (2003a))
78.2 Schistochila sciurea (Nees) Schiffn. [1, 2, 25, 80, 134, 169, 209]
Schistochila thwaitesii (Mitt.) Steph. [1, 2, 37, 227]
Gottschea thwaitesii Mitt. [148]; type from Sri Lanka; syn. fide So (2003a).
Solenostomataceae
79.1 Solenostoma appressifolium (Mitt.) Váňa & D. G. Long
Jungermannia appressifolia Mitt. [6, 169, 241, 245]
79.2 Solenostoma ariadne (Taylor ex Lehm.) R. M. Schust ex Váňa & D. G. Long
Jungermannia ariadne Taylor ex Lehm. [169]
79.3 Solenostoma borneense (Amakawa) Váňa, Hentschel & J. Heinrichs
Jungermannia borneensis Amakawa [169]
(Solenostoma comatum (Nees) C.Gao
Jungermannia comata Nees [1, 2, 227]; Váňa (1972) rejected all records of this species from Sri
Lanka)
79.4 Solenostoma hasskarlianum (Nees) R.M.Schust. ex Váňa & D.G.Long
Jungermannia hasskarliana (Nees) Steph. [1, 2, 6, 199, 241, 245]
79.5 Solenostoma javanicum (Schiffn.) Steph.
Jungermannia herzogiana Váňa [169, 245]; syn. fide Váňa & Piippo (1989).
(Solenostoma obovatum (Nees) R.M.Schust.
Jungermannia obovata Nees [199]. A north temperate species not known in tropical Asia (Paton,
1999); report considered erroneous.)
79.6 Solenostoma polyrhizoides (Grolle ex Amakawa) Váňa & D. G. Long
Jungermannia polyrhizoides Grolle ex Amakawa [169]
79.7 Solenostoma strictum (Schiffn.) Váňa, Hentschel & J.Heinrichs
Jungermannia stricta (Schiffn.) Steph. [245]
79.8 Solenostoma tetragonum (Lindenb.) R. M. Schust. ex Váňa & D. G. Long
Jungermannia tetragona Lindenb. [6, 169, 241, 245]
79.9 Solenostoma truncatum (Nees) R. M. Schust. ex Váňa & D. G. Long
Jungermannia truncata Nees [5, 134, 245]
Jungermannia polyrhiza Hook. ex Lehm. & Lindenb. [80, 148, 169]; syn. fide Amakawa (1972).
Southbyaceae
80.1 Southbya organensis Herzog [243]
Southbya grollei N.Kitag. [169, 178]; syn. fide Váňa et al. (2012)
Targioniaceae
81.1 Targionia hypophylla L. [1, 2]
Trichocoleaceae
82.1 Trichocolea pluma (Reinw., Blume & Nees) Mont. [1, 2, 123, 169]
16
Long and Rubasinghe
HORNWORTS
Anthocerotaceae
83.1 Anthoceros agrestis Paton
Aspiromitus punctatus (L.) Schljakov subsp. agrestis (Paton) R.M.Schust. [202]
83.2 Anthoceros erectus Kashyap [1, 2, 128]
84.1 Folioceros amboinensis (Schiffn.) Piippo [185]
Anthoceros amboinensis Schiffn. [1, 2, 128]
Folioceros spinisporus (Steph.) D.C. Bharadwaj [7]; syn. fide Piippo (1993a)
Folioceros mamillisporus (D.C.Bharadwaj) D.C.Bharadwaj; syn. fide Bharadwaj (1973)
Anthoceros mamillisporus (D.C.Bharadwaj) D.C.Bharadwaj [2, 128]
Aspiromitus mamillisporus D.C.Bharadwaj [13, 14]; type from Sri Lanka.
85.1 Phaeoceros carolinianus (Michx.) Prosk.
Phaeoceros laevis (L.) Prosk. subsp. carolinianus (Michx.) Prosk. [128, 169]
Anthoceros communis Steph. [1, 2]; syn. fide Hasegawa (1984).
(Phaeoceros laevis (L.) Prosk. [1, 2]; records probably refer to P. carolinianus.)
Dendrocerotaceae
86.1 Dendroceros borbonicus Steph. [77]
CONCLUSIONS
Abeywickrama & Jansen (1978a) published the
only previous checklist of liverworts and hornworts
of Sri Lanka. They reported 66 genera and 183
species of liverworts and 3 genera and 5 species of
hornworts. The present list has increased these
totals to 82 genera and 327 species of liverworts
and 4 genera and 5 species of hornworts. The main
reason for these changes is the numerous
publications resulting from new field investigations
over the past 36 years by many bryologists, mostly
from other countries. These activities have been
summarized by Rubasinghe & Long (2014).
However, a number of early reports were not
available to Abeywickrama & Jansen (1978a) and
these have now been incorporated. Also, a number
of species accepted by Abeywickrama & Jansen
(1978a) have since been relegated to synonymy
(e.g. the synonyms of Folioceros amboinensis) and
others are considered doubtful, such as Saccogyna
viticulosa.
The largest family of liverworts by far is the
Lejeuneaceae with 25 genera and 138 species,
including the largest genus Cololejeunea with 36
species. Outside the Lejeuneaceae the largest
genera are Frullania with 28 species and
Plagiochila with 19 accepted species. Complex
thalloid liverworts are poorly represented with 10
genera and 20 species of which the largest are
Marchantia and Riccia, both with 6 species. Why
the simple thalloid genus Riccardia (with only two
species) is so poorly represented is puzzling, and
surely others have been overlooked. These totals
clearly indicate that the most important habitats in
Sri Lanka for liverworts are the wet evergreen
forests with numerous small species, particularly
epiphyllous Lejeuneaceae. However, further work
in other habitats will surely reveal more species of
hornworts and complex thalloid liverworts.
ACKNOWLEDGMENTS
The library staff at the Royal Botanic Gardens at
Edinburgh and Kew are thanked for bibliographic
help and staff at the National Herbarium,
Peradeniya, for information on early collectors and
their collections at PDA. Lars Söderström,
Department of Biology, Norwegian University of
Science and Technology, Trondheim, Norway, is
thanked for his major input into numerous
taxonomic clarifications and addition of many
published reports, and other valuable comments.
REFERENCES
[1] Abeywickrama, B.A. 1959. The genera of the
liverworts of Ceylon. Ceylon Journal of
Science, Biological Sciences 2: 33–81.
[2] Abeywickrama, B.A. & Jansen, M.A.B. 1978a.
Revised checklist to the Liverworts and Hornworts of Sri Lanka
A check list of the liverworts of Sri Lanka.
UNESCO: Man and the Biosphere National
Committee for Sri Lanka Publication 1: 1–10.
[3] Abeywickrama, B.A. & Jansen, M.A.B. 1978b.
A check list of the mosses of Sri Lanka.
UNESCO: Man and the Biosphere National
Committee for Sri Lanka Publication 2: 1–25.
[4] Alam, A., Vats, S. & Behera, K.K. 2012.
Exormotheca ceylonensis Meijer - a threatened
liverwort in India, rediscovered in Palni Hills,
Tamil Nadu. Journal of threatened taxa 4:
2593–2595.
[5] Amakawa, T. 1972. New or little known Asiatic
species of the family Jungermanniaceae, VII.
Journal of the Hattori Botanical Laboratory 35:
382–390.
[6] Amakawa, T. 1973 '1972'. Miscellaneous notes
on the Asiatic Jungermannioideae II. Journal of
the Hattori Botanical Laboratory 36: 57–74.
[7] Asthana, A.K. & Srivastava, S.C. 1991. Indian
Hornworts (a taxonomic study). Bryophytorum
Bibliotheca 42: 1–158.
[8] Asthana, G. & Shukla, A. 2009. Two
epiphyllous species of Drepanolejeunea
(Spruce) Schiffn. new to the Indian bryoflora.
Journal of Bryology 31: 139–142.
[9] Asthana, G. & Srivastava, S.C. 2003. Indian
Cololejeunea.
A
taxonomic
study.
Bryophytorum Bibliotheca 60: 1–155.
[10] Asthana, G., Srivastava, S.C. & Asthana, A.K.
1995. The genus Cheilolejeunea in India.
Lindbergia 20: 125–143.
[11] Awasthi, U.S. & Udar, R. 1984. The genus
Mastigolejeunea (Spruce) Schiffn. in India.
Proceedings, Indian Academy of Sciences.
Plant Sciences 93: 485–494.
[12] Benedix, E.H. 1953. Indomalayische
Cololejeuneen. Feddes Repertorium Beiheft
134: 1–88.
[13] Bharadwaj, D.C. 1948. Studies in Indian
Hepaticae V. On a new species of Aspiromitus
St. from Kandy, Ceylon. Aspiromitus
mamillispora Bhardwaj. sp. nov. Journal of
Indian Botanical Society 27: 200–205.
[14] Bharadwaj, D.C. 1965. Studies in Indian
Anthocerotaceae IV. On the morphology of
Anthoceros mamillisporus (Bhardw.) emend.
and A. dixitianus (Mahabalé) Prosk. Journal of
the Indian Botanical Society 44: 347–364.
[15] Bharadwaj, D.C. 1973. Taxonomy of some
Indo-Pacific species of Folioceros Bharadwaj.
Geophytology 3: 215–221.
[16] Bischler, H. 1961. Recherches sur l’anatomie
de la tige chez les Lejeuneaceae Paradoxae.
Revue Bryologique et Lichénologique 30: 232–
252.
[17] Bischler, H. 1966. Recherches sur l’anatomie
de la tige chez les Lejeuneaceae. III. Les sous-
17
familles Lejeuneoideae, tribu Lejeuneeae, et
Tuyamaelloideae. Revue Bryologique et
Lichénologique 34: 601–675.
[18] Bischler, H. 1968. Monograph du genre
Rhaphidolejeunea Herzog. Revue Bryologique
et Lichénologique n.s. 36: 56–104.
[19] Bischler, H. 1979. Plagiochasma Lehm. et
Liindenb. III. Les taxa d'Asie et d'Oceanie.
Journal of the Hattori Botanical Laboratory 45:
25–79.
[20] Bischler, H. 1989. Marchantia. The Asiatic
and Oceanic taxa. Bryophytorum Bibliotheca
38: 1–317.
[21] Bischler, H., Miller, H.A. & Bonner, C.E.B.
1961. Studies in Lejeuneaceae III. A historical
account of Lejeunea cucullata (Reinwardt,
Blume & Nees) and its varieties. Nova
Hedwigia 3: 445–462.
[22] Bischler, H., & Piippo, S. 1991. Bryophyte
Flora of the Huon Peninsula, Papua New
Guinea. L. Marchantia (Marchantiaceae,
Hepaticae). Annales Botanici Fennici 28: 277–
301.
[23] Bonner, C.E.B. 1953. De Hepaticis II. An
unpublished section of volume 6 of Stephani's
Species Hepaticarum: the genus Marchantia.
Candollea 14: 102–112.
[24] Bonner, C.E.B. 1965. Index Hepaticarum. Pars
5: Delavayella to Geothallus. Weinheim: J.
Cramer.
[25] Buch, H. 1939. Die Schistochila-Arten der
Inseln Sumatra, Java und Celebes. Annales
Bryologici 12: 1–20.
[26] Castle, H. 1950. A revision of the genus
Radula. Part II. Subgenus Acroradula. Section
2. Amentulosae. Bryologist 53: 253–275.
[27] Castle, H. 1959. A revision of the genus
Radula. Part II. Subgenus Acroradula. Section
3. Dichotomae. Journal of the Hattori Botanical
Laboratory 21: 1–52.
[28] Castle, H. 1965. A revision of the genus
Radula. Part II. Subgenus Acroradula. Section
9. Densifoliae. Revue Bryologique et
Lichénologique 33: 328–398.
[29] Castle, H. 1968. Radula (L.) Dumortier. – A
synopsis of the taxonomic revision of the genus.
Revue Bryologique et Lichénologique n.s. 36:
5–44.
[30] Costa, D.P. da. 2008. Metzgeriaceae
(Hepaticae). Flora Neotropica Monograph 102:
1–169.
[31] Crandall-Stotler, B., Stotler, R. & Long, D.G.
2009. Phylogeny and classification of the
Marchantiophyta. Edinburgh Journal of Botany
66: 1–44.
[32] Daniels, A.E.D. & Kariyappa, K.C. 2013.
Cheilolejeunea trapezia (Nees) Kachroo &
R.M.Schust. ex Mizut. var. ceylanica
18
(Gottsche) A.E.D.Daniels & K.C.Kariyappa
comb. et stat. nov. (Lejeuneaceae) from India.
Taiwania 58: 140–145.
[33] Dey, M. & Singh, D.K. 2012. Epiphyllous
liverworts of Eastern Himalaya. Kolkata:
Botanical Survey of India.
[34] Dey, M., Singh, D. & Singh, D.K. 2013. The
genus Drepanolejeunea (Marchantiophyta;
Lejeuneaceae) in India with anote on three new
additions from Eastern Himalaya. Tropical
Bryology 35: 14–25.
[35] Dugas, M. 1929. Contribution à l’étude du
genre “Plagiochila” Dum. Annales des Sciences
Naturelles, Botanique (série 10) 11: 1–199.
[36] Eggers, J. & Pócs, T. 2010. Epiphylle
Lebermoose aus Samoa. Auf den Spuren von
Eduard Graeffe in der Farn-Abteilung des
Herbarium Hamburgense. Berichte des
Botanischen Vereins zu Hamburg 25: 3–11.
[37] Eggers, J. & Schäfer-Verwimp, A. 1987. Some
liverworts new to Sri Lanka (Ceylon). Journal
of Bryology 14: 531–534.
[38] Eifrig, H. 1937. Monographische Studien über
die Indomalayischen Arten von Taxilejeunea.
Annales Bryologici 9: 73–114.
[39] Engel, J.J. & Merrill, G.L. 2004. Austral
Hepaticae. 35. A taxonomic and phylogenetic
study of Telaranea (Lepidoziaceae), with a
monograph of the genus in temperate
Australasia and commentary on extraAustralasian taxa. Fieldiana, Botany N.S. 44: 1–
265.
[40] Feldberg, K., Hentschel, J., Bombosch, A.,
Long, D.G., Váňa, J. & Heinrichs, J. 2009.
Transfer of Gottschelia grollei, G. patoniae and
Scaphophyllum speciosum to Solenostoma
based on chloroplast DNA rbcL sequences.
Plant Systematics and Evolution 280: 243–250.
[41] Furuki, T. 1995. Taxonomic studies of Asiatic
species of Aneuraceae (Hepaticae). II.
Riccardia subgen Hyaloneura Schust. Journal
of the Hattori Botanical Laboratory 78: 111–
118.
[42] Furuki, T. 2006. Taxonomical studies of the
family Aneuraceae (Hepaticae) based on the
Philippine collections made by Dr. and Mrs A.J.
Sharp and Dr. Z. Iwatsuki. Journal of the
Hattori Botanical Laboratory 100: 89–99.
[43] Gao, C. & Wu, Y.-H. 2004. On seven species
of the Schistochilaceae (Hepaticae) in China,
including one new species and one new
combination. Journal of the Hattori Botanical
Laboratory 95: 263–270.
[44] Gottsche, C.M. 1886. Ueber Lebermoose von
Ceylon. Botanisches Centralblatt 26: 203–205.
[45] Gottsche, C.M., Lindenberg J.B.W. & Nees
von Esenbeck, C.G. 1844. Synopsis
Hepaticarum. Hamburg: Meissner.
Long and Rubasinghe
[46] Gradstein, S.R. 1975. A taxonomic monograph
of the genus Acrolejeunea (Hepaticae).
Bryophytorum Bibliotheca 4: 1–162.
[47] Gradstein, S.R., He, X.-L., Piippo, S. &
Mizutani, M. 2002. Bryophyte flora of the Huon
Peninsula, Papua New Guinea. LXVII.
Lejeuneaceae subfamily Ptychanthoideae
(Hepaticae). Acta Botanica Fennica 174: 1–88.
[48] Gradstein, S.R. & Inoue, H. 1980. Studies on
Lejeuneaceae subfam. Ptychanthoideae, V. A
review of the species from Ceylon. Bulletin of
the National Science Museum, Tokyo, series B,
6: 23–32.
[49] Gradstein, S.R. & Terken, L. 1981. Studies on
Lejeuneaceae subfam. Ptychanthoideae VI. A
revision of Schiffneriolejeunea Sect. Saccatae
from Asia. Occasional Papers from the Farlow
Herbarium 16: 71–81.
[50] Gradstein, S.R. & Váňa, J. 1987. On the
occurrence of Laurasian liverworts in the
Tropics. Memoirs of the New York Botanical
Garden 45: 388–425.
[51] Grolle, R. 1960. Über Saccogyna Dum. und
Saccogynidium, eine neue Lebermoosgattung.
Journal of the Hattori Botanical Laboratory 23:
41–67.
[52] Grolle, R. 1965a. Lebermoose aus Neuguinea.
I. Journal of the Hattori Botanical Laboratory
28: 43–54
[53] Grolle, R. 1965b. Miscellanea hepaticologica
31–40. Journal of the Hattori Botanical
Laboratory 28: 101–106.
[54] Grolle, R. 1968. Gottschelia – eine neue
Jungermanniales-Gattung der Paläotropis.
Journal of the Hattori Botanical Laboratory 31:
13–19.
[55] Grolle, R. 1971. Jamesoniella und Verwandte.
Feddes Repertorium 82:1–99.
[56] Grolle, R. 1976. Drepanolejeunea subgen.
Kolpolejeunea – eine neue Untergattung aus der
Paläotropis. Journal of the Hattori Botanical
Laboratory 40: 191–216.
[57] Grolle, R. 1977a. Miscellanea hepaticologica
161–170. Journal of Bryology 9: 529–538.
[58] Grolle, R. 1977b. Lebermoose aus Neuguinea
14–15. Calypogeia + Trichocolea. Journal of
the Hattori Botanical Laboratory 43: 63–67.
[59] Grolle, R. 1978a. Miscellanea hepaticologica
161–170. Journal of Bryology 9: 529–538.
[60] Grolle, R. 1978b. Eine neue DiplasiolejeuneaArt aus Sri Lanka. Feddes Repertorium 89:
301–305.
[61] Grolle, R. 1978c. Die Lebermoose der
Seychellen. Wissenschaftlich Zeitschrift der
Friedrich-Schiller-Universität Jena/Thüringen,
Mathematische-naturwissenschaftliche Reihe
27: 7–17.
[62] Grolle, R. 1979a. Lebermoose aus Neuguinea
Revised checklist to the Liverworts and Hornworts of Sri Lanka
17. Harpalejeunea. Journal of the Hattori
Botanical Laboratory 46: 43–47.
[63] Grolle, R. 1979b. Miscellanea hepaticologica
191–200. Journal of the Hattori Botanical
Laboratory 46: 337–355.
[64] Grolle, R. 1989. Miscellanea hepaticologica
271–280. Journal of the Hattori Botanical
Laboratory 67: 249–254.
[65] Grolle, R. 1995. The Hepaticae and
Anthocerotae of the East African Islands. An
annotated catalogue. Bryophytorum Bibliotheca
48: 1–178.
[66] Grolle, R. & Piippo, S. 1986. Bryophyte flora
of the Huon Peninsula, Papua New Guinea.
XVI. Pallaviciniaceae (Hepaticae). Acta
Botanica Fennica 133: 59–79.
[67] Grolle, R. & Piippo, S. 1990. Bryophyte flora
of the Huon Peninsula, Papua New Guinea.
XVII.
Leucolejeunea
(Lejeuneaceae,
Hepaticae). Annales Botanici Fennici 27: 119–
129.
[68] Grolle, R. & Reiner-Drehwald, M.E. 1999.
Review of the genus Harpalejeunea
(Lejeuneaceae) including the description of H.
grandis, a new species from the páramos of
Colombia. Journal of Bryology 21: 31–45.
[69] Grolle, R. & So, M.-L. 1998. Studies on
Plagiochila in China II. Bryologist 101: 282–
294.
[70] Grolle, R. & So, M.-L. 1999a. Studies of
Plagiochila sect. Subtropicae in Asia.
Bryologist 102: 67–75.
[71] Grolle, R. & So, M.L. 1999b. Studies on
Plagiochila in China III. Bryologist 102: 200–
207.
[72] Grolle, R. & So, M.-L. 1999c. Studies on
Plagiochila sect. Tayloriae in Asia. Bryologist
102: 287–293.
[73] Grolle, R. & So, M.-L. 1999d. Studies on
Plagiochila sect. Contiguae in East and South
Asia. Bryologist 102: 294–303.
[74] Grolle, R. & Vàňa. J. 1969. Jungermannia
subulata Evans aus Hawaii – eine verkannte
Sippe
der
Holarktis.
Oesterreichische
botanische Zeitschrift 117: 305–313.
[75] Grolle, R. & Zhu, R.-L. 2000. A study of
Drepanolejeunea subg. Rhaphidolejeunea
(Herzog) Grolle & R.L.Zhu, stat. nov.
(Hepaticae, Lejeuneaceae) in China with notes
on its species elsewhere. Nova Hedwigia 70:
373–396.
[76] Haerida, I., Gradstein, S.R., Tjitrosoedirdjo,
S.S.
2010.
Lejeuneaceae
subfamily
Ptychanthoideae (Hepaticae) in West Java.
Gardens' Bulletin Singapore 62: 53–103.
[77] Hasegawa, J. 1981. Dendroceros borbonicus
Steph. – a new record for Sri Lanka.
Miscellanea Bryologica et Lichenologica 9: 39–
19
41.
[78] Hasegawa, J. 1984. Taxonomical studies on
Asian Anthocerotaceae. IV. A revision of the
genera Anthoceros, Phaeoceros and Folioceros
in Japan. Journal of the Hattori Botanical
Laboratory 57: 241–272.
[79] Hattori, S. 1967. Studies on the Asiatic species
of the genus Porella (Hepaticae). I. Some new
or little known Asiatic species of Porella.
Journal of the Hattori Botanical Laboratory 30:
129–151.
[80] Hattori, S. 1968. Hepaticae collected by F.
Schmid in Ceylon and Pakistan. Candollea 23:
287–294.
[81] Hattori, S. 1969. Studies of the Asiatic species
of the genus Porella (Hepaticae). II. Journal of
the Hattori Botanical Laboratory 32: 319–359.
[82] Hattori, S. 1972a. Frullania tamariscicomplex and the species concept. Journal of the
Hattori Botanical Laboratory 35: 202–251.
[83] Hattori, S. 1972b. Notes on Asiatic species of
the genus Frullania, Hepaticae. I. Journal of the
Hattori Botanical Laboratory 36: 109–140.
[84] Hattori, S. 1973a. Notes on Asiatic species of
the genus Frullania, Hepaticae, II. Journal of
the Hattori Botanical Laboratory 37: 55–84.
[85] Hattori, S. 1973b. Notes on Asiatic species of
the genus Frullania, Hepaticae, III. Journal of
the Hattori Botanical Laboratory 37: 85–120.
[86] Hattori, S. 1974. Notes on the Asiatic species
of the genus Frullania, Hepaticae. V. Journal of
the Hattori Botanical Laboratory 38: 185–221.
[87] Hattori, S. 1975a. Notes on the Asiatic species
of the genus Frullania, Hepaticae. VII. Journal
of the Hattori Botanical Laboratory 39: 277–
313.
[88] Hattori, S. 1975b. A revision of Indian species
of Frullania (Hepaticae) published by W.
Mitten in 1861. Bulletin of the National Science
Museum Tokyo, Series B (Botany) 1: 73–81.
[89] Hattori, S. 1975c. Notes on the Asiatic species
of the genus Frullania, Hepaticae. VIII. Bulletin
of the National Science Museum Tokyo, Series
B (Botany) 1: 141–163.
[90] Hattori, S. 1976a. Studies on the Asiatic
species of the genus Porella (Hepaticae). VI.
Journal of the Hattori Botanical Laboratory 40:
121–138.
[91] Hattori, S. 1976b. Notes on the Asiatic species
of the genus Frullania, Hepaticae. X. Journal of
the Hattori Botanical Laboratory 40: 461–507.
[92] Hattori, S. 1978. Studies on the Asiatic species
of the genus Porella (Hepaticae), VII. A
synopsis of Asiatic Porellaceae. Journal of the
Hattori Botanical Laboratory 44: 91–120.
[93] Hattori, S. 1979. Dr. H. Inoue's collection of
Frullania taxa (Hepaticae) made in Ceylon.
Journal of Japanese Botany 54: 22–26.
20
[94] Hattori, S. 1980a. Notes of the Asiatic species
of the genus Frullania, Hepaticae, XII. Journal
of the Hattori Botanical Laboratory 47: 85–
125.
[95] Hattori, S. 1980b. A revision of the subgenus
Homotropantha of the genus Frullania,
Hepaticae. Journal of the Hattori Botanical
Laboratory 47: 165–236.
[96] Hattori, S. 1985. A preliminary study of
Chinese Frullania flora. Journal of the Hattori
Botanical Laboratory 59: 123–169.
[97] Hattori, S. 1986. Notes on the Asiatic species
of the genus Frullania, Hepaticae XIV. Bulletin
of the National Science Museum Tokyo, Series
B (Botany) 12: 127–138.
[98] Hattori, S. & Mizutani, M. 1969. Hepaticae.
In: Studies in the Flora of Thailand 59. Dansk
Botanisk Arkiv 27: 92–98.
[99] Hattori, S. & Piippo, S. 1986. Bryophyte flora
of the Huon Peninsula, Papua New Guinea. XV.
Frullania (Frullaniaceae, Hepaticae). Acta
Botanica Fennica 133: 25–58.
[100] He, X.L. 1999. A taxonomic monograph of
the genus Pycnolejeunea (Lejeuneaceae,
Hepaticae). Acta Botanica Fennica 163: 1–77.
[101] He, X. & Sun, Y. 2013. Multigene evidence
reveals
the
systematic
position
of
Pleurocladopsis
simulans
(C.Massal.)
R.M.Schust. within Schistochila Dumort.,
Schistochilaceae. Polish Botanical Journal 58:
467–474.
[102] He, Q., Liu, L.-J., Liu, C.-X., Liu, G.-L., Ma,
Y., Gao, D.-L., Wei, Y.-M., Shi, R.-P., Shu, L
& Zhu, R.-L. 2013. Synonymy, taxonomic
notes, and range extensions for several
noteworthy Asian liverworts. Journal of
Bryology 35: 123–128.
[103] Herzog, T. 1921. Die Lebermoose der 2
Freiburger Molukkenexpeditionen und einige
neuen Arten der engeren Indomalaya. Beiheft
zum Botanischen Centralblatt 38: 318–332.
[104] Herzog, T. 1930. Studien über
Drepanolejeunea. Annales Bryologici 3: 126–
149.
[105] Herzog, T. 1937. Studien über
Drepanolejeunea III. Annales Bryologici 9:
115–130.
[106] Herzog, T. 1939. Studien über
Drepanolejeunea, IV. Annales Bryologici 12:
98–122.
[107] Herzog, T. 1942. Revision der
Lebermoosgattung Leptolejeunea Spr. in der
Indomalaya. Flora 135: 377–434.
[108] Herzog, T. 1943. Rhaphidolejeunea Herz.,
eine
neue
Lejeuneaceengattung
der
Indomalaya. Mitteilungen des Thüringischen
Botanischen Vereins 50: 100–105.
[109] Hoffmann, G. 1935. Monographische
Long and Rubasinghe
Studien über die Indomalayischen Arten von
Pycnolejeunea. Annales Bryologici 8: 80–129.
[110] Horikawa, Y. 1934. Monographia
Hepaticarum Australi-Japonicarum. Journal of
Science of the Hiroshima University Series B,
Div. 2 (Botany) 2: 101–325.
[111] Inoue, H. 1964. The genus Plagiochilion.
Journal of the Hattori Botanical Laboratory 27:
51–72.
[112] Inoue, H. 1965a. Contributions to the
knowledge of the Plagiochilaceae of
southeastern Asia VI. Studies on the
Plagiochila semidecurrens complex Journal of
the Hattori Botanical Laboratory 28: 209–218.
[113] Inoue, H. 1965b. Contributions to the
knowledge of the Plagiochilaceae of
Southeastern Asia VII. Synopsis of Plagiochila
Dum. in the Himalayan region. Bulletin of the
National Science Museum, Tokyo 8: 375–403.
[114] Inoue, H. 1966. A monograph of the hepatic
genus Syzygiella Spr. Journal of the Hattori
Botanical Laboratory 29: 171–213.
[115] Inoue, H. 1968. Taxonomic miscellany on the
genera Plagiochila and Syzygiella. Bulletin of
the National Science Museum, Tokyo 11: 299–
307.
[116] Inoue, H. 1969. Contributions to the
knowledge of the Plagiochilaceae of
southeastern Asia. XII. Typification of
Plagiochila javanica (Swartz) Dum. Journal of
the Hattori Botanical Laboratory 32: 50–56.
[117] Inoue, H. 1978. Studies on Taiwan Hepaticae,
III. Subord. Herbertinae and Subord. Ptilidiinae.
Bulletin of the National Science Museum Tokyo,
Series B (Botany) 4: 93–103.
[118] Inoue, H. 1979. Contributions to the
knowledge of the Plagiochilaceae of
southeastern Asia, XVIII. A review of the genus
Plagiochila (Dum.) Dum. in Ceylon. Journal of
the Hattori Botanical Laboratory 46: 193–233.
[119] Inoue, H. 1981. Taxonomic miscellany on
hepatics (1). Journal of Japanese Botany 56:
105–110.
[120] Inoue, H. 1984. The genus Plagiochila (Dum.)
Dum. in southeast Asia. Tokyo: Academia
Scientific Book Inc.
[121] Inoue, H. & Miller, H.A. 1967. Some
noteworthy liverworts from Ceylon. Bulletin of
the National Science Museum, Tokyo 10: 11–
17.
[122] Jack, J.B. 1886. Monographie der
Lebermoosgattung Physiotium. Hedwigia 25:
49–87.
[123] Jayasuriya, A.H.M. 1984. Flora of Ritigala
Strict Natural Reserve. Sri Lanka Forester 16:
61–156.
[124] Jovet-Ast, S. 1953. Le genre Colura.
Hépatiques, Lejeuneaceae, Diplasiae. Revue
Revised checklist to the Liverworts and Hornworts of Sri Lanka
Bryologique et Lichénologique 22: 206–316.
[125] Jovet-Ast, S. 2003. Riccia des sous-genres
Riccia et Ricciella récoltés en Inde et en
Indonésie. Cryptogamie, Bryologie 24: 209–
228.
[126] Juslén, A. 2004. Bryophyte flora of Hunan
Province, China. 7. Herbertus (Herbertaceae,
Hepaticae). Annales Botanici Fennici 41: 393–
404
[127] Juslén, A. 2006. Revision of Asian Herbertus
(Herbertaceae, Marchantiophyta). Annales
Botanici Fennici 43: 409–436.
[128] Kachroo, P., Bapna, K.R. & Dhar, G.L. 1977.
Hepaticae of India. A taxonomic survey and
census. V (concld.). Fossombroniaceae through
Anthocerotaceae. Journal of the Indian
Botanical Society 56: 62–86.
[129] Kamimura, M. 1961. A monograph of
Japanese Frullaniaceae. Journal of the Hattori
Botanical Laboratory 24: 1–109.
[130] Kitagawa, N. 1967. Studies on the Hepaticae
of Thailand. I. The genus Bazzania, with
general introduction. Journal of the Hattori
Botanical Laboratory 30: 249–270.
[131] Kitagawa, N. 1969. Studies on the Hepaticae
of Thailand. II. Journal of the Hattori Botanical
Laboratory 32: 290–306.
[132] Kitagawa, N. 1972. Miscellaneous notes on
little-known species of Hepaticae, 1–25.
Journal of the Hattori Botanical Laboratory 36:
444–454.
[133] Kitagawa, N. 1973. Miscellaneous notes on
little-known species of Hepaticae, 26–50.
Journal of the Hattori Botanical Laboratory 37:
263–273.
[134] Kitagawa, N. 1978. The Hepaticae of
Thailand collected by Dr. A. Touw (I). Acta
Phytotaxonomica et Geobotanica 29: 47–64.
[135] Kitagawa, N. 1981. Miscellaneous notes on
little-known species of Hepaticae, 51–70.
Hikobia, suppl. 1: 67–72
[136] Kitagawa, N. 1988. Studies on the Hepaticae
of Thailand V. The family calypogeiaceae.
Beiheft zur Nova Hedwigia 90: 163–170.
[137] Krayeski, D.M., Crandall-Stotler, B. &
Stotler, R. 2005. A revision of the genus
Fossombronia Raddi in East Asia and Oceania.
Journal of the Hattori Botanical Laboratory 98:
1–45.
[138] Kruijt, R.C. 1988. A monograph of the genera
Dicranolejeunea
and
Acanthocoleus.
Bryophytorum Bibliotheca 36: 1–136.
[139] Kuwahara, Y. 1975. A recent collection of
Metzgeriaceae in the eastern Himalayas.
Journal of the Hattori Botanical Laboratory 39:
363–371.
[140] Kuwahara, Y. 1984a. Synopsis of sect.
Biseria Kuw. subgen. Metzgeria Kuw., gen.
21
Metzgeria Raddi (Hepaticae). Hikobia 9: 31–
42.
[141] Kuwahara, Y. 1984b. Annotated keys to the
species of the Metzgeriaceae from Japan, East
Himalayas, Tropical East Africa, and Tropical
Oceania. Nova Hedwigia 40: 273–285.
[142] Kuwahara, Y. & Onraedt, M. 1979. Notes on
the genus Metzgeria (Hepaticae) in Sri Lanka.
Bulletin du Jardin Botanique National de
Belgique, Brussels 49: 77–80.
[143] Lee, G.E. 2013. A systematic revision of the
genus Lejeunea Lib. (Marchantiophyta:
Lejeuneaceae) in Malaysia. Cryptogamie,
Bryologie 34: 381–484.
[144] Meijer, W. 1956. A new species of
Exormotheca from Ceylon. Journal of the
Hattori Botanical Laboratory 16: 72–74.
[145] Meijer, W. 1972. The discovery of
Ephemeropsis
tjibodensis
Goebel
(Ephemeropsidaceae) in the everwet rainforests
of Ceylon. Journal of the Hattori Botanical
Laboratory 36: 552–555.
[146] Miller, H.A. 1965. A review of Herberta in
the tropical Pacific and Asia Journal of the
Hattori Botanical Laboratory 28: 299–412.
[147] Miller, H.A., Whittier, H.O. & Whittier, B.A.
1983. Prodromus florae hepaticarum Polynesiae
with a key to genera. Bryophytorum Bibliotheca
25: 1–423.
[148] Mitten, W. 1860–1861. Hepaticae Indiae
Orientalis: an enumeration of the Hepaticae of
the East Indies. Journal of the Proceedings of
the Linnean, Society 5: 89–108, 109–128.
[149] Mizutani, M. 1961. A revision of Japanese
Lejeuneaceae. Journal of the Hattori Botanical
Laboratory 24: 115–302.
[150] Mizutani, M. 1963. On some Indian species
of the family Lejeuneaceae described by W.
Mitten. Journal of the Hattori Botanical
Laboratory 26: 171–184.
[151] Mizutani, M. 1965. Studies of little known
Asiatic species of Hepaticae in the Stephani
herbarium. 2. On some little known Southeast
Asiatic species of the genus Cololejeunea.
Journal of the Hattori Botanical Laboratory 28:
107–121.
[152] Mizutani, M. 1967a. Studies of the
Himalayan species of Bazzania. Journal of the
Hattori Botanical Laboratory 30: 71–90.
[153] Mizutani, M. 1967b. Studies of little known
Asiatic species of Hepaticae in the Stephani
herbarium. 3. On some little known species of
Cheilolejeunea,
Euosmolejeunea
and
Pycnolejeunea. Journal of the Hattori
Botanical Laboratory 30: 171–180.
[154] Mizutani, M. 1968. Studies of little known
Asiatic species of Hepaticae in the Stephani
herbarium. 5. On some flagelliform species of
22
Lepidozia. Journal of the Hattori Botanical
Laboratory 31: 152–158.
[155] Mizutani, M. 1971. Lejeunea from the
Himalayan region. Journal of the Hattori
Botanical Laboratory 34: 445–457.
[156] Mizutani, M. 1972. Studies of little known
Asiatic species of Hepaticae in the Stephani
herbarium. 7. Some little known species of the
subfamily Lejeuneoideae of the Lejeuneaceae.
Journal of the Hattori Botanical Laboratory 35:
399–411.
[157] Mizutani, M. 1980a. Notes on the
Lejeuneaceae. 3. Some Asiatic species of the
genus Cheilolejeunea. Journal of the Hattori
Botanical Laboratory 47: 319–331.
[158] Mizutani, M. 1980b. Notes on the
Lejeuneaceae. 4. Asiatic species of the genus
Dicranolejeunea. Journal of the Hattori
Botanical Laboratory 48: 235–242.
[159] Mizutani, M. 1982. Notes on the
Lejeuneaceae. 6. Japanese species of the genus
Cheilolejeunea. Journal of the Hattori
Botanical Laboratory 51: 151–173.
[160] Mizutani, M. 1984. Notes on the
Lejeuneaceae. 9. Cololejeunea lanciloba and its
related species in Japan. Journal of the Hattori
Botanical Laboratory 57: 427–442.
[161] Mizutani, M. 1985. Notes on the
Lejeuneaceae. 10. Some Asiatic species of the
genus Lopholejeunea. Journal of the Hattori
Botanical Laboratory 59: 469–480.
[162] Mizutani, M. 1986. Notes on the
Lejeuneaceae. 12. Mastigolejeunea humilis and
its related species from Asia. Journal of the
Hattori Botanical Laboratory 61: 281–297.
[163] Mizutani, M. 1989. Notes of the
Lejeuneaceae. 15. Three species of the genus
Trocholejeunea. Journal of the Hattori
Botanical Laboratory 66: 271–281.
[164] Mizutani, M. 1990. Notes on the
Lejeuneaceae.
16.
Drepanolejeunea
thwaitesiana and its related species from Asia.
Journal of the Hattori Botanical Laboratory 68:
367–380.
[165] Mizutani, M. & Chang, K.C. 1986. A
preliminary study of Cinese Lepidoziaceae
Flora. Journal of the Hattori Botanical
Laboratory 60: 419–437.
[166] Onraedt, M. 1978. Bryophytes de Sri Lanka
(Ceylan). Revue Bryologique et Lichénologique
44: 77–82.
[167] Onraedt, M. 1979a. Bryophytes de Sri Lanka
(Ceylan) IV. Le genre Colura. Revue
Bryologique et Lichénologique 45: 443–452.
[168] Onraedt, M. 1979b. Bryophytes de Sri Lanka
(Ceylan) III. Cololejeunea ceylanica Onr. et
Cololejeunea hinidumae Onr., especes
nouvelles.
Acta
Botanica
Academiae
Long and Rubasinghe
Scientiarum Hungaricae 25: 107–111.
[169] Onraedt, M. 1981. Bryophytes récoltées a Sri
Lanka (Ceylan). V. Journal of the Hattori
Botanical Laboratory 50: 191–216.
[170] Onraedt, M. 1983. Bryophytes de Sri Lanka
(Ceylan) VI. Le genre Radula Dum.
Cryptogamie Bryologique et Lichénologique 4:
71–85.
[171] Onraedt, M. 1985. Bryophytes de Sri Lanka
VII. Lejeuneaceae Holostipae. Cryptogamie
Bryologique et Lichénologique 6: 151–175.
[172] O'Shea. B.J. 2002. Checklist of the mosses of
Sri Lanka. Journal of the Hattori Botanical
Laboratory 92: 125–164.
[173] Pandé, S.K., Srivastava, K.P. & Ahmad, S.
1957. Epiphyllous liverworts of India and
Ceylon – II. Journal of Indian Botanical Society
36: 335–347.
[174] Pandé, S.K. & Udar, R. 1957. Genus Riccia
in India. I. A reinvestigation of the taxonomic
status of the ndian species of Riccia. Journal of
the Indian Botanical Society 36: 564–579.
[175] Paton, J.A. 1999. The Liverwort Flora of the
British Isles. Colchester: Harley Books.
[176] Piippo, S. 1984a. Bryophyte flora of the Huon
Peninsula,
Papua
New
Guinea.
III.
Haplomitriaceae, Lepicoleaceae, Herbertaceae,
Pseudolepicoleaceae,
Trichocoleaceae,
Schistochilaceae,
Balantiopsaceae,
Pleuroziaceae and Porellaceae (Hepaticae).
Annales Botanici Fennici 21: 21–48.
[177] Piippo, S. 1984b. Bryophyte flora of the
Huon Peninsula, Papua New Guinea. VI.
Lepidoziaceae
subfam.
Lepidozioideae,
Calypogeiaceae,
Adelanthaceae,
Cephaloziaceae subfam. Cephalozioideae and
subfam. Odontoschismatoideae and Jubulaceae
(Hepaticae). Annales Botanici Fennici 21: 309–
335.
[178] Piippo, S. 1985a. Bryophyte flora of the Huon
Peninsula,
Papua
New
Guinea.
X.
Jackiellaceae, Scapaniaceae, Arnelliaceae and
Acrobolbaceae (Hepaticae). Acta Botanica
Fennica 131: 89–97.
[179] Piippo, S. 1985b. Bryophyte flora of the
Huon Peninsula, Papua New Guinea. XII.
Geocalycaceae (Hepaticae). Acta Botanica
Fennica 131: 129–167.
[180] Piippo, S. 1985c. Bryophyte flora of the Huon
Peninsula, Papua New Guinea. XIII.
Arachniopsis and Kurzia (Lepidoziaceae
subfam. Lepidozioideae, Hepaticae). Acta
Botanica Fennica 131: 169–179.
[181] Piippo, S. 1989a. Bryophyte flora of the Huon
Peninsula,
Papua
New
Guinea.
X.
Plagiochilaceae (Hepaticae). Annales Botanici
Fennici 26: 183–236.
[182] Piippo, S. 1989b. Bryophyte flora of the
Revised checklist to the Liverworts and Hornworts of Sri Lanka
Huon Peninsula, Papua New Guinea. XIX.
Fossombronia
(Fossombroniaceae)
and
Metzgeria (Metzgeriaceae, Hepaticae). Acta
Botanica Fennica 143: 1–22.
[183] Piippo, S. 1990. Annotated catalogue of
Chinese Hepaticae and Anthocerotae. Journal
of the Hattori Botanical Laboratory 68: 1–192.
[184] Piippo, S. 1991. Bryophyte flora of the Huon
Peninsula, Papua New Guinea. XL.
Acromastigum
and
Hygrolembidium
(Lepidoziaceae subfam Bazzanioideae and
Lembidioideae, Hepaticae). Acta Botanica
Fennica 143: 23–34.
[185] Piippo, S. 1993a. Bryophyte flora of the Huon
Peninsula, Papua New Guinea. LIV.
Anthocerotophyta. Acta Bot. Fennica 148: 27–
51.
[186] Piippo, S. 1993b. On the taxonomy and
nomenclature of SW Asiatic Geocalycaceae.
Annales Botanici Fennici 30: 195–303.
[187] Piippo, S. 1996. Notes on Chinese
Geocalycaceae (Hepaticae). 1. Annales
Botanici Fennici 33: 45–49.
[188] Pócs, T. 1968. The genus Porella in Vietnam.
Journal of the Hattori Botanical Laboratory 31:
65–93.
[189] Pócs, T. 1969. A short survey of the Bazzania
of North Viet-Nam. Journal of the Hattori
Botanical Laboratory 32: 79–94.
[190] Pócs, T., Mizutani, M., Piippo, S. 1994.
Bryophyte flora of the Huon Peninsula, Papua
New Guinea. LXV. Preliminary contributions
on Lejeuneaceae (Hepaticae). Annales Botanici
Fennici 31: 179–190.
[191] Pócs, T. & Ninh, T. 2005. Contribution to the
bryoflora of Vietnam. VI. On the liverwort flora
of Vu Quang Nature Reserve. Acta Botanica
Hungarica 47: 151–171.
[192] Pócs, T. & Piippo, S. 2011. Bryophyte flora
of the Huon Peninsula, Papua New Guinea.
LXXIV.
Cololejeunea
(Lejeuneaceae,
Hepaticae). Acta Bryolichenologica Asiatica 4:
59–137.
[193] Pócs, T., Sass-Gyarmati, A., Naikatini, A.,
Tuiwawa, M., Braggins, J.E., Pócs, S. & von
Konrat,
M.
2011.
New
liverwort
(Marchantiophyta) records for the Fiji Islands.
Telopea 13: 455–494.
[194] Rawat, K.K. & Srivastava, S.C. 2007. Genus
Plagiochila in Eastern Himalaya (India). Dehra
Dun: Bishen Singh Mahendra Pal Singh.
[195] Reimers, H. 1931 Beitrage zur Moosflora
Chinas 1. Hedwigia 71: 1–77.
[196] Reiner-Drehwald, M.E. & Grolle, R. 2012.
Review of the genus Rectolejeunea
(Lejeuneaceae,
Marchantiophyta).
Nova
Hedwigia 95: 451–482.
[197] Renner, M.A.M. 2005. Additions to the
23
Radula (Radulaceae: Hepaticae) floras of New
Zealand and Tasmania. Journal of the Hattori
Botanical Laboratory 97: 39–79.
[198] Rubasinghe, S.C.K. & Long, D.G. 2014.
Bryophytes of Sri Lanka: a review of past
exploration and taxonomic research and
priorities for the future. Journal of Bryology (in
press).
[199] Ruklani, N.C.S. & Rubasinghe, S.C.K. 2013.
A preliminary survey of bryophytes of the
Central Province of Sri Lanka. Ceylon Journal
of Science (Biological Sciences) 42: 67–72.
[200] Schiffner, V. 1900a. Die Hepaticae der Flora
von Buitenzorg. Leiden: E.J. Brill.
[201] Schiffner, V. 1900b. Exposito plantarum in
itinere Suo Indico annis 1893/94 suspecto
collectarum, II. Denkschriften der kaiserlichen
Akademie
der
Wissenschaften,
Wien,
Mathematisch-naturwissenschaftliche Klasse
70: 155–218.
[202] Schuster, R.M. 1992. The Hepaticae and
Anthocerotae of North America east of ther
hundredth meridian. Volume 6. Chicago: Field
Museum of Natural History.
[203] Schuster, R.M. & Engel, J.J. 1987. A
monograph
of
Lepidoziaceae
subfam.
Lembidioideae (Hepaticae). Journal of the
Hattori Botanical Laboratory 63: 247–350.
[204] Scott, E.B. 1960. A monograph of the genus
Lepicolea (Hepaticae). Nova Hedwigia 2: 129–
172.
[205] Sharma, D. & Srivastava, S.C. 1993. Indian
Lepidoziineae
(a
taxonomic
study).
Bryophytorum Bibliotheca 47: 1–353.
[206] Singh, A.P., Kumar, D. & Nath, V. 2008.
Studies on the genera Frullania Raddi and
Jubula Dum. from Meghalaya (India): Eastern
Himalayas. Taiwania 53: 51–84.
[207] So, M.L. 2001. Plagiochila (Hepaticae,
Plagiochilaceae) in China — Systematic Botany
Monographs 60: 1–214.
[208] So, M.L. 2002. Metzgeria (Hepaticae) in
Australasia and the Pacific. New Zealand
Journal of Botany 40: 603–627.
[209] So, M.L. 2003a. The genus Schistochila in
Asia. Journal of the Hattori Botanical
Laboratory 93: 79–100.
[210] So, M.L. 2003b. The genus Metzgeria
(Hepaticae) in Asia. Journal of the Hattori
Botanical Laboratory 94: 159–177.
[211] So, M.L. 2006. Radula (Radulaceae,
Marchantiophyta) in the South Pacific. Journal
of the Hattori Botanical Laboratory 99: 207–
232.
[212] So, M.L. & Grolle, R. 1999. Studies on
Plagiochila in Asia: supplements to sections
Abietinae, Annotinae, Ciliatae, Contiguae,
Cucullatae, Poeltiae, Subtropicae and Zonatae.
24
Cryptogamie, Bryologie 20: 167–179.
[213] So, M.L. & Grolle, R. 2000a. Checklist of
Plagiochila (Hepaticae) in Asia. Journal of the
Hattori Botanical Laboratory 88: 199–243.
[214] So, M.L. & Grolle, R. 2000b. Studies on
Plagiochila sect. Plagiochila (Hepaticae) in
East and South Asia. Journal of Bryology 22:
17–28.
[215] So, M.L. & Grolle, R. 2001. On several littleknown species of Plagiochila (Hepaticae) in
Asia. Journal of Bryology 23: 123–132.
[216] So, M.L. & Grolle, R. 2001. On Plagiochila
subgenus Plagiochila section Abietinae
(Hepaticae). Systematic Botany 26: 459–469.
[217] So, M.L. & Zhu, R.-L. 1998. On six species
of the genus Lejeunea in China, including one
new species. Bryologist 101: 137–143.
[218] Söderström, L., Váňa, J., Crandall-Stotler, B.,
Stotler, R.E., Hagborg, A. & von Konrat, M.
2013. Notes on Early Land Plants Today. 43.
New combinations in Lophocoleaceae
(Marchantiophyta). Phytotaxa 112: 18–32.
[219] Srivastava, A. & Srivastava, S.C. 2002.
Indian
Geocalycaceae
(Hepaticae.
(A
taxonomic study). Dehra Dun: Bishen Singh
Mahendra Pal Singh.
[220] Srivastava, K.P. 1964. Bryophytes of India –
I. Ricciaceae. Bulletin, National Botanic
Gardens, Lucknow 104: 1–103.
[221] Srivastava, S.C. & Dixit, R. 1996. The genus
Cyathodium Kunze. Journal of the Hattori
Botanical Laboratory 80: 149–215.
[222] Srivastava, S.C., Sharma, D. & Shaheen, F.
1990. Diplophyllum nanum Herz. in India.
Lindbergia 16: 55–58.
[223] Stephani, F. 1884. Die Gattung Radula.
Hedwigia 23: 145–159.
[224] Stephani, F. 1885. Hepaticarum species
novae vel minus cognitae. Hedwigia 24: 246–
250
[225] Stephani, F. 1886. Hepaticarum species
novae vel minus cognitae. Hedwigia 25: 233–
249.
[226] Stephani, F. 1896. Hepaticarum species
novae IX. Hedwigia 35: 73–140.
[227] Stephani, F. 1898–1925. Species
Hepaticarum. 6 volumes. Genève, Bale &
Lyon: Herbier Boissier.
[228] Stephani, F. 1890. Die Gattung Lejeunea in
Herbarium Lindenberg. Hedwigia 29: 1–23,
68–99, 133–142.
[229] Svihla, R.D. 1956. The Frullaniaceae of
Burma. I. Bryologist 59: 277–280.
[230] Thiers, B.M. 1985. Branching in
Lejeuneaceae III. Ptychanthoideae. Beiheft zur
Nova Hedwigia 80: 31–61.
[231] Thiers, B.M. 1986. Branching in the
Lejeuneaceae (Hepaticae). IV. Lejeuneoideae.
Long and Rubasinghe
Nova Hedwigia 42: 237–275.
[232] Thiers, B.M. 1992. A re-evaluation of
Cheilolejeunea
subgenus
Xenolejeunea.
Tropical Bryology 5: 11–21.
[233] Thiers, B.M. 1993. A monograph of
Pleurozia
(Hepaticae;
Pleuroziaceae).
Bryologist 96: 517–554.
[234] Tixier, P. 1973. Bryophytes exotiques.
Bulletin du Muséum National d'Histoire
Naturelle (Sér. 3) Botanique 10: 73–86.
[235] Tixier, P. 1977. Lejeuneaceae of Sri Lanka
collected by D.D. Tirvengadum (1974). Ceylon
Journal of Science (Biological Sciences) 12:
85–88.
[236] Tixier, P. 1981. La notion d'espece chez le
genre Cololejeunea. Le complexe Cololejeunea
floccosa (Lehm. & Lindenb.) Schiffn.
Cryptogamie Bryologique et Lichénologique 2:
47–76.
[237] Tixier, P. 1984. La notion d'espece chez le
genre Cololejeunea. Le complexe Cololejeunea
longifolia (Mitt.) Benedix est-il monotypique?
Cryptogamie Bryologique et Lichénologique 5:
111–125.
[238] Tixier, P. 1985. Contribution à la
Connaisscance
des
Cololejeuneoideae.
Bryophytorum Bibliotheca 27: 1–439.
[239] Udar, R. & Kumar, A. 1981. Notoscyphus
Mitt. in India. Journal of the Hattori Botanical
Laboratory 49: 247–260.
[240] Udar, R. & Kumar, A. 1982. Studies in East
Himalayan Hepaticae – II: the genus
Chandonanthus Mitten. Geophytology 12: 59–
64.
[241] Váňa, J. 1972. Miscellaneous notes on the
Asiatic Jungermannioideae II. Journal of the
Hattori Botanical Laboratory 36: 57–74.
[242] Váňa, J. 1973. Studien über die
Jungermannioideae
(Hepaticae)
3.
Jungermannia subg. Liochlaena. Folia
Geobotanica & Phytotaxonomica, Praha 8:
397–416.
[243] Váňa, J., Grolle, R. & Long, D.G. 2012.
Taxonomic realignments and new records of
Gongylanthus and Southbya (Marchantiophyta:
Southbyaceae) from the Sino-Himalayan
region. Nova Hedwigia 95: 183–196.
[244] Vàňa, J. & Long, D.G. 2009.
Jungermanniaceae of the Sino-Himalayan
region. Nova Hedwigia 89: 485–517.
[245] Váňa, J. & Piippo, S. 1989. Bryophyte flora
of the Huon Peninsula, Papua New Guinea. IX.
Jungermanniaceae
and
Gymnomitriaceae
(Hepaticae). Annales Botanici Fennici 26: 107–
125.
[246] Váňa, J., Söderström, L., Hagborg, A. & von
Konrat, M. 2013. Notes on Early Land Plants
Today. 40. Notes on Cephaloziellaceae.
Revised checklist to the Liverworts and Hornworts of Sri Lanka
Phytotaxa 112: 1–6.
[247] Vanderpoorten, A. & Goffinet, B. 2009.
Introduction to Bryophytes. Cambridge:
Cambridge University Press.
[248] Verdoorn, F. 1929. V. Schiffner-expositio
plantarum in itinere suo Indico annis 1893/94
suscepto
collectarum
speciminibusque
exsiccatis
distributarum,
adjectis
descriptionibus novarum. Series tertia (No.
1473–2460). Frullaniaceas continens. Annales
Bryologici 2: 117–154.
[249] Verdoorn, F. 1930. Die Frullaniaceae der
Indomalesischen Inseln. Annales Bryologici
Supplementary Volume 1: 1–187.
[250] Verdoorn, F. 1933. Die von V. Schiffner
(1893–1894) und von Fr. Verdoorn (1930) auf
den indomalesischen Inseln gesammelten
Lejeuneaceae Holostipae. De Frullaniaceis XI.
Recueil des Travaux Botaniques Neerlandais
30: 212–233.
[251] Verdoorn, F. 1934. Studien über Asiatische
Jubuleae. Annales Bryologici Supplementary
Volume 4: 1–231.
[252] Yamada, K. 1975a. Radula collection made
by Dr. H. Inoue in Ceylon. Journal of Japanese
Botany 50: 372–378.
[253] Yamada, K. 1975b. Memoranda of the type
specimens of Radula taxa from the Southeast
Asia (2). Journal of the Hattori Botanical
Laboratory 39: 71–84.
[254] Yamada, K. 1976. Memoranda on the type
specimens of Radula taxa from the south-east
Asia (3). Journal of the Hattori Botanical
Laboratory 40: 453–460.
[255] Yamada, K. 1979a. A revision of Asian taxa
of Radula, Hepaticae. Journal of the Hattori
Botanical Laboratory 45: 201–322.
[256] Yamada, K. 1979b. A new species of Radula
(Hepaticae) from Ceylon. Miscellanea
Bryologica et Lichenologica 8: 113–114.
[257] Yamada, K. 1999. Taxonomy of the section
Amentulosae of genus Radula (Radulaceae,
Hepaticae). Haussknechtia Beiheft 9: 391–397.
[258] Yamada, K. & Piippo, S. 1989. Bryophyte
Flora of the Huon Peninsula, Papua New
Guinea. XII. Radula (Radulaceae, Hepaticae).
Annales Botanici Fennici 26: 349–387.
25
[259] Yuzawa, Y. 1991. A monograph of subgen.
Chonanthelia of gen. Frullania (Hepaticae) of
the world. Journal of the Hattori Botanical
Laboratory 70: 181–291.
[260] Zhu, R.-L. 1995. Notes on some species of
the genus Cololejeunea (Lejeuneaceae,
Hepaticae) in China. Journal of the Hattori
Botanical Laboratory 78: 83–109.
[261] Zhu, R.-L. 2006. Taxonomy and distribution
of
Cheilolejeunea
krakakammae
(Lejeuneaceae,
Jungermanniopsida,
Marchantiophyta), with a description and
illustration of Cheilolejeunea laevicalyx from
Bolivia, Colombia and Ecuador. Nova
Hedwigia 83: 187–198.
[262] Zhu, R.-L. & Gradstein, S.R. 2005.
Monograph of Lopholejeunea (Lejeuneaceae,
Hepaticae) in Asia. Systematic Botany
Monographs 74: 1–98.
[263] Zhu, R.-L. & Lai, M.-J. 2005. Cheilolejeunea
decursiva (Sande Lac.) R.M. Schust. and C.
ventricosa (Schiffn.) X.L.He (Lejeuneaceae,
Hepaticae) new to Australia. Journal of the
Hattori Botanical Laboratory 98: 193–203.
[264] Zhu, R.-L. & So, M.L. 1999. New records of
Cololejeunea (Lejeuneaceae, Hepaticae) for
China and Vietnam. Botanical Bulletin of
Academia Sinica 40: 165–171.
[265] Zhu, R.-L. & So, M.L. 2000a. Notes on the
taxonomy and distribution of Lejeunea
neelgherriana and Tuyamaella serratistipa
(Hepaticae, Lejeuneaceae). Annales Botanici
Fennici 37: 149–153.
[266] Zhu, R.-L. & So, M.L. 2000b. Additions and
corrections
for
Chinese
Lejeuneaceae
(Hepaticae). Botanical Bulletin of Academia
Sinica 41: 243–250.
[267] Zhu, R.-L. & So, M.L. 2001. Epiphyllous
liverworts of China. Nova Hedwigia Beiheft
121: 1–418.
[268] Zhu, R.-L., So, M.L. & Wang, Y.-F. 2002.
The
genus
Cheilolejeunea
(Hepaticae,
Lejeuneaceae) in China. Nova Hedwigia 75:
387–408.
APPENDIX I – FORMAL TAXONOMIC CHANGES
26
Long and Rubasinghe
Bazzania obtusa D.G.Long & Rubasinghe, nom. nov. for Mastigobryum obtusatum Mitt., Journal of the
Proceedings of the Linnean Society 5: 106. 1860.
Syn.: Bazzania obtusata (Mitt.) Abeyw., Ceylon Journal of Science, Biological Sciences 2: 45. 1959, hom.
illeg. non Bazzania obtusata (Taylor) Kuntze, Revisio Generum Plantarum 833. 1891.
The plant described by Mitten (1960–1861) as Mastigobryum obtusatum Mitt. belongs to the genus
Bazzania, and the combination Bazzania obtusata was made by Abeywickrama and Jansen (1978a).
However this is illegitimate due to the existence of the earlier Bazzania obtusata (Taylor) Kuntze. The
substitute name Bazzania obtusa is therefore proposed.
Heteroscyphus fleischeri (Steph.) D.G.Long & Rubasinghe, comb. nov.
Basionym: Chiloscyphus fleischeri Steph., Species Hepaticarum 6: 306. 1922. Type citation: ‘Insula Ceylon
(Fleischer legit)'.
The genus Chiloscyphus Corda has been subdivided in various ways in the past, but the segregate genus
Heteroscyphus Schiffn. is now widely accepted as one of these, with four other species recognised in Sri
Lanka. From Stephani’s description (Stephani 1898–1925) Chilscyphus fleischeri belongs to Heteroscyphus
and is therefore formally transferred to that genus.
APPENDIX II – INDEX OF ACCEPTED NAMES AND SYNONYMS
Accepted names are given in bold italics; synonyms in italics.
Acanthocoleus gilvus (Gottsche) Kruijt 26.1
Acrolejeunea emergens (Mitt.) Steph. 27.1
Acrolejeunea planiuscula (Mitt.) Steph. 27.2
Acrolejeunea pycnoclada (Taylor) Schiffn. 27.3
Anastrophyllum bidens (Reinw, Blume & Nees) Steph. = 7.1
Anastrophyllum imbricatum (Wilson ex Gottsche, Lindenb. & Nees) Steph. 5.1
Anastrophyllum piligerum (Reinw., Blume & Nees) Steph. 5.2
Andrewsianthus puniceus (Nees) R.M. Schust. ex Grolle 61.1
Aneura pinguis (L.) Dumort. 8.1
Anthoceros agrestis Paton 83.1
Anthoceros amboinensis Schiffn. = 84.1
Anthoceros communis Steph. = 85.1
Anthoceros erectus Kashyap 83.2
Anthoceros mamillisporus (D.C.Bharadwaj) D.C.Bharadwaj = 84.1
Arachniopsis major Herz. = 56.1
Archilejeunea mariana (Gottsche) Steph. = 46.1
Aspiromitus mamillisporus D.C.Bharadwaj = 84.1
Aspiromitus punctatus (L.) Schljakov subsp. agrestis (Paton) R.M.Schust. = 83.1
Bazzania callida (Steph.) Abeyw. 52.1
Bazzania ceylanica (Mitt.) W.E.Nicholson 52.2
Bazzania decurva (Nees) Kuntze = 52.11
Bazzania erosa (Nees) Trevis. 52.3
Bazzania fleischeri (Steph.) Abeyw. 52.4
Bazzania gaudichaudii (Gottsche ex Steph.) Schiffn. 52.5
Bazzania grandistipula (Steph.) Abeyw. = 52.2
Bazzania japonica (Sande Lac.) Lindb. 52.6
Bazzania lacerostipula (Steph.) Abeyw. = 52.2
Bazzania oblonga (Mitt.) Schiffn. = 52.14
Bazzania obtusa D.G.Long & Rubasinghe 52.7
Bazzania obtusata (Mitt.) Abeyw. = 52.7
Bazzania oshimensis (Steph.) Horik. 52.8
Bazzania ovistipula (Steph.) Abeyw. 52.9
Bazzania pearsonii Steph. 52.10
Bazzania praerupta (Reinw., Blume & Nees) Trevis. 52.11
Bazzania rabenhorstii (Steph.) Abeyw. 52.12
Bazzania rotundistipula (Steph.) Abeyw. = 52.10
Bazzania sandei (Steph.) Abeyw. = 52.11
Revised checklist to the Liverworts and Hornworts of Sri Lanka
Bazzania spiralis (Reinw., Blume & Nees) Meijer 52.13
Bazzania stricta (Steph.) Abeyw. = 52.2
Bazzania tridens (Reinw., Blume & Nees) Trevis. 52.14
Bazzania vittata (Gottsche) Trevis. 52.15
Blasia pusilla L. 12.1
Brachiolejeunea sandvicensis (Gottsche) A.Evans = 49.2
Calypogeia alternifolia (Nees) Steph. = 14.1
Calypogeia apiculata (Steph.) Steph. 13.1
Calypogeia arguta Nees & Mont. 13.2
Calypogeia baldwinii Aust. = 15.1
Calypogeia ceylanca S.Hatt. & Mizut. 13.3
Calypogeia fissa (L.) Raddi 13.4
Calypogeia fragilis (Steph.) Inoue & H.A.Miller 13.5
Calypogeia fusca (Lehm.) Steph. = 15.1
Campylolejeunea inflectens (Mitt.) Mizut. = 30.16
Caudalejeunea cristiloba (Steph.) Gradst. 28.1
Caudalejeunea lessonii Steph. = 28.2
Caudalejeunea reniloba (Gottsche) Steph. 28.2
Cephalozia willisana Steph. = 17.1
Cephaloziella kiaeri (Austin) Pearson = 17.1
Cephaloziella willisana (Steph.) N.Kitag. = 17.1
Ceratolejeunea thwaitesiana (Mitt.) Steph. = 33.9
Chandonanthus birmensis Steph. = 6.1
Chandonanthus hirtellus (F.Weber) Mitt. = 6.2
Cheilolejeunea ceylanica (Gottsche) R.M.Schust. & Kachroo 29.1
Cheilolejeunea decurrens (Sande Lac.) X.-L. He 29.2
Cheilolejeunea decursiva (Sande Lac.) R.M.Schust. 29.3
Cheilolejeunea gardneri (Mitt.) Mizut. 29.4
Cheilolejeunea imbricata (Nees) S.Hatt. 29.5
Cheilolejeunea intertexta (Lindenb.) Steph. 29.6
Cheilolejeunea krakakammae (Lindenb.) R.M.Schust. 29.7
Cheilolejeunea kurzii Steph. = 29.6
Cheilolejeunea lindenbergii (Gottsche) Mizut. 29.8
Cheilolejeunea longiloba (Steph. ex G.Hoffm.) J.J.Engel & B.C.Tan 29.9
Cheilolejeunea nietneri (Steph.) Mizut. 29.10
Cheilolejeunea riparia (Mitt.) M.E.Reiner 35.16
Cheilolejeunea serpentina (Mitt.) Mizut. 29.11
Cheilolejeunea streimannii Pócs & Ninh 29.12
Cheilolejeunea subopaca (Mitt.) Mizut. 29.13
Cheilolejeunea trapezia (Nees) Mizut. 29.14
Cheilolejeunea trapezia (Nees) Mizut. var. ceylanica (Gottsche) A.E.D.Daniels & Kariyappa = 29.1
Cheilolejeunea trifaria (Reinw., Blume & Nees) Mizut. 29.15
Cheilolejeunea vittata (Steph. ex G.Hoffm.) R.M.Schust. & Kachroo 29.16
Chiloscyphus argutus (Reinw., Blume & Nees) Nees = 59.1
Chiloscyphus ceylonensis J.J.Engel = 58.2
Chiloscyphus ciliolatus (Nees) J.J.Engel & R.M.Schust. = 58.1
Chiloscyphus fleischeri Steph. = 59.2
Chiloscyphus kurzii (Sande Lac.) J.J.Engel & R.M.Schust. = 60.3
Chiloscyphus perfoliatus (Mont.) Nees = 59.3
Cololejeiunea vesicaria (Sande Lac.) Benedix 30.35
Cololejeunea amoena Benedix 30.1
Cololejeunea appressa (A.Evans) Benedix 30.2
Cololejeunea bolombensis (Steph.) Vanden Berghen = 30.17
Cololejeunea ceratilobula (P.C.Chen) R.M.Schust. 30.3
Cololejeunea ceylanica Onr. 30.4
Cololejeunea cordiflora Steph. 30.5
Cololejeunea desciscens Steph. 30.6
Cololejeunea falcata (Horik.) Benedix 30.7
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Long and Rubasinghe
Cololejeunea falcatoides Benedix = 30.7
Cololejeunea fissilobula Herz. 30.8
Cololejeunea floccosa (Lehm. & Lindenb.) Steph. 30.9
Cololejeunea floccosa (Lehm. & Lindenb.) Steph. var. aurita Benedix 30.9a
Cololejeunea floccosa (Lehm. & Lindenb.) Steph. var. trivitata Tixier 30.9b
Cololejeunea formosana Mizut. = 30.6
Cololejeunea fredericii Onr. 30.10
Cololejeunea gottschei (Steph.) Mizut. 30.11
Cololejeunea gynophthalma Benedix 30.12
Cololejeunea haskarliana (Lehm. & Lindenb.) Steph. 30.13
Cololejeunea hinidumae Onr. 30.14
Cololejeunea inflata Steph. 30.15
Cololejeunea inflectens (Mitt.) Benedix 30.16
Cololejeunea lanciloba Steph. 30.17
Cololejeunea leonidens Benedix var. saccata Benedix 30.18
Cololejeunea longifolia (Mitt.) Benedix ex Mizut. 30.19
Cololejeunea metzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilkiu-Borges & J. Heinrichs 30.20
Cololejeunea minutissima (Sm.) Schiffn. 30.21
Cololejeunea obliqua (Nees & Mont.) Schiffn. 30.22
Cololejeunea oshimensis (Horik.) Benedix = 30.15
Cololejeunea peraffinis (Schiffn.) Schiffn. 30.23
Cololejeunea perakensis Tixier 30.24
Cololejeunea planissima (Mitt.) Abeyw. 30.25
Cololejeunea pluripunctata Benedix 30.26
Cololejeunea pretiosa Benedix 30.27
Cololejeunea pseudofloccosa (Horik.) Benedix 30.28
Cololejeunea scabrifolia Schiffn., nom. inval. = 30.22
Cololejeunea schmidtii Steph. 30.29
Cololejeunea serrata (Steph.) Benedix 30.30
Cololejeunea taprobanea Tixier 30.31
Cololejeunea tenella Benedix 30.32
Cololejeunea triapiculata (Herzog) Tixier 30.33
Cololejeunea trichomanis (Gottsche) Steph. subsp. cordiflora (Steph.) Pócs 30.34
Cololejeunea vidaliana Tixier 30.36
Colura acroloba (Mont. ex Steph.) Ast 31.1
Colura acutifolia Ast. = 31.5
Colura ari (Steph.) Steph. 31.2
Colura brevistyla Herzog 31.3
Colura calyptrifolia (Hook.) Dumort. 31.4
Colura calyptrifolia (Hook.) Dumort. subsp. tenuicornis (A.Evans) Vanden Berghen = 31.11
Colura conica (Sande Lac) K.I.Goebel 31.5
Colura corynephora (Gottsche, Lindenb. & Nees) Trevis. 31.6
Colura greig-smithii Ast 31.7
Colura javanica Steph. 31.8
Colura meijeri Ast 31.9
Colura ornata K.I.Goebel 31.10
Colura tenuicornis (A.Evans) Steph. 31.11
Colura verdoornii Herzog & Ast 31.12
Cryptolophocolea ciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler & Váňa 58.1
Cryptolophocolea fleischeri (Steph.) L.Söderstr. 58.2
Cuspidatula flexicaulis (Nees) Váňa & L.Söderstr. 2.1
Cyathodium cavernarum Kunze 19.1
Cyathodium smaragdinum Schiffn. ex Keissl. 19.2
Cylindrocolea kiaeri (Austin) Váňa 17.1
Dendroceros borbonicus Steph. 86.1
Denotarisia linguifolia (De Not.) Grolle 3.1
Dicranolejeunea gilva (Gottsche) Steph. = 26.1
Diplasiolejeunea cavifolia Steph. 32.1
Revised checklist to the Liverworts and Hornworts of Sri Lanka
Diplasiolejeunea onraedtii Grolle 32.2
Diplasiolejeunea rudolphiana Steph. 32.3
Diplophyllum nanum Herzog 76.1
Drepanolejeunea angustifolia (Mitt.) Grolle 33.1
Drepanolejeunea fissicornua Steph. 33.2
Drepanolejeunea fleischeri (Steph.) Grolle & R.L.Zhu 33.3
Drepanolejeunea macrodonta (Mitt.) Steph. 33.4
Drepanolejeunea micholitzii Steph. = 33.5
Drepanolejeunea micholitzii Steph. var. dactylophoroides Herzog = 33.5
Drepanolejeunea pentadactyla (Mont.) Steph. 33.5
Drepanolejeunea tenera K.I.Goebel 33.6
Drepanolejeunea ternatensis (Gottsche) Schiffn. 33.7
Drepanolejeunea teysmannii Steph. 33.8
Drepanolejeunea thwaitesiana (Mitt.) Steph. 33.9
Drepanolejeunea thwaitesiana (Mitt.) Steph. var. zhengii R.L.Zhu 33.9a
Drepanolejeunea vesiculosa (Mitt.) Steph. 33.10
Dumortiera hirsuta (Sw.) Nees. s.l. 20.1
Eulejeunea nietneri Steph. = 35.12
Euosmolejeunea serpentina (Mitt.) Steph. = 29.11
Euosmolejeunea trifaria (Reinw, Blume & Nees) Spruce = 29.15
Exormotheca ceylonensis Meijer 21.1
Folioceros amboinensis (Schiffn.) Piippo 84.1
Folioceros mamillisporus (D.C.Bharadwaj) D.C.Bharadwaj = 84.1
Folioceros spinisporus (Steph.) D.C. Bharadwaj = 84.1
Frullania acutiloba Mitt. 22.1
Frullania acutiloba Mitt. var. schiffneri Verd. = 22.19a
Frullania alstonii Verd. 22.2
Frullania angustata Mitt. = 22.23a
Frullania aperta Gottsche = 22.20
Frullania apiculata (Reinw., Blume & Nees) Dumort. 22.3
Frullania apiculata (Reinw., Blume & Nees) Dumort. var. goebelii Schiffn. 22.3a
Frullania arecae (Spreng.) Gott. 22.4
Frullania brotheri Steph. 22.5
Frullania campanulata Sande Lac. 22.6
Frullania capilliformis Steph. 22.7
Frullania ceylanica Nees = 22.10
Frullania claviloba Steph. 22.8
Frullania ericoides (Nees) Mont. 22.9
Frullania explanata Mitt. = 22.15
Frullania galeata (Reinw., Blume & Nees) Dumort. = 22.25
Frullania gaudichaudii (Nees & Mont.) Nees & Mont. var. ceylanica (Nees) S.Hatt. 22.10
Frullania gracilis (Reinw., Blume & Nees) Dumort. 22.11
Frullania grandistipula Lindenb. 22.12
Frullania grandistipula Lindenb. var. rugosa (Mitt.) S.Hatt. = 22.12
Frullania hampeana Nees = 22.19
Frullania hampeana Nees var. acutiloba (Mitt.) S.Hatt. = 22.1
Frullania hottana S.Hatt. 22.13
Frullania hypoleuca Nees 22.14
Frullania intermedia (Reinw., Blume & Nees) Dumort. 22.15
Frullania laciniosa Lehm. = 22.9
Frullania latistipula Steph. = 22.26
Frullania meyeniana Lindenb. 22.16
Frullania microauriculata Verd. 22.17
Frullania microauriculata Verd. var. rotundior Verd. 22.17a
Frullania moniliata (Reinw., Blume & Nees) Mont. 22.18
Frullania moniliata (Reinw., Blume & Nees) Mont. subsp. breviramea (Steph.) Verd. 22.18a
Frullania moniliata (Reinw., Blume & Nees) Mont. subsp. obscura Verd. 22.18b
Frullania monocera (Taylor) Taylor 22.19
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Long and Rubasinghe
Frullania monocera (Taylor) Taylor var. schiffneri (Verd.) S.Hatt. 22.19a
Frullania neurota Taylor 22.20
Frullania nietneri Gottsche ex Steph. = 22.10
Frullania nodulosa (Reinw., Blume & Nees) Nees 22.21
Frullania orientalis Sande Lac. 22.22
Frullania pendula Mitt. = 22.27
Frullania perocellata Onr. = 22.2
Frullania polyptera Taylor 22.23
Frullania polyptera Taylor var. angustata (Mitt.) S.Hatt. 22.23a
Frullania ramuligera (Nees) Mont. 22.24
Frullania riojaneirensis (Raddi) Spruce 22.25
Frullania rugosa Mitt. = 22.12
Frullania serrata Gottsche 22.26
Frullania serrata Gottsche forma crispulodentata Verd. 22.26a
Frullania squarrosa (Reinw., Blume & Nees) Nees = 22.9
Frullania tamarisci (L.) Dumort. subsp. moniliata (Reinw., Blume & Nees) Kamimura = 22.18
Frullania tamarisci (L.) Dumort. subsp. obscura (Verd.) S.Hatt. = 22.18b
Frullania tamarisci (L.) Dumort. subsp. obscura (Verd.) S.Hatt. var. breviramea (Steph.) S.Hatt. = 22.18a
Frullania ternatensis Gottsche 22.27
Frullania thuilleri Nees = 22.21
Frullania trichodes Mitt. 22.28
Frullania vethii Sande Lac. = 22.28
Frullania wallichiana Mitt. = 22.4
Gottschea aligera Nees & Blume = 78.1
Gottschea philippinensis Mont. = 78.1
Gottschea thwaitesii Mitt. = 78.2
Gottschelia schizopleura (Spruce) Grolle 18.1
Gymnanthe lutescens (Lehm. & Lindenb.) Mitt. = 67.1
Harpalejeunea filicuspis (Steph.) Mizut. 34.1
Harpalejeunea indica Steph. = 35.11
Harpalejeunea nietneri Steph. = 29.10
Herbertus ceylanicus (Steph.) H.A.Miller 23.1
Herbertus dicranus (Taylor ex Gottsche, Lindenb. & Nees) Trevis. 23.2
Herbertus perrottetii (Steph.) Abeyw. = 23.2
Herbertus ramosus (Steph.) H.A.Mill. 23.3
Heteroscyphus argutus (Reinw., Blume & Nees) Schiffn. 59.1
Heteroscyphus fleischeri (Steph.) D.G.Long & Rubasinghe 59.2
Heteroscyphus perfoliatus (Mont.) Schiffn. 59.3
Heteroscyphus tener (Steph.) Schiffn. 59.4
Heteroscyphus tridentatus (Sande Lac.) Grolle 59.5
Hygrolejeunea discreta (Lindenb.) Steph. = 35.5
Hygrolejeunea lindenbergii (Gottsche) Steph. = 29.8
Hygrolembidium boschianum (Sande Lac.) R.M.Schust. 53.1
Jackiella ceylanica Schiffn. ex Steph. 24.1
Jackiella javanica Schiffn. 24.2
Jamesoniella flexicaulis (Nees) Schiffn. = 2.1
Jamesoniella microphylla Schiffn. = 18.1
Jamesoniella ovifolia (Schiffn.) Schiffn. = 3.1
Jensenia decipiens (Mitt.) Grolle 68.1
Jubula hutchinsiae (Hook.) Dumort. subsp. javanica (Steph.) Verdoorn 25.1
Jungermannia appressifolia Mitt. = 79.1
Jungermannia ariadne Taylor ex Lehm. = 79.2
Jungermannia borneensis Amakawa = 79.3
Jungermannia ceylanica Gottsche nom. nud. ? = 18.1
Jungermannia ceylanica Steph. = 18.1
Jungermannia colorata Lehm. var. ceylanica Gottsche nom. nud. ? = 2.1
Jungermannia flexicaulis Nees = 2.1
Jungermannia hasskarliana (Nees) Steph. = 79.4
Revised checklist to the Liverworts and Hornworts of Sri Lanka
Jungermannia herzogiana Váňa = 79.5
Jungermannia imbricata Wilson ex Gottsche, Lindenb. & Nees = 5.1
Jungermannia leiantha Grolle subsp. stephanii (Amak.) S.Hatt. = 65.1
Jungermannia piligera Reinw., Blume & Nees = 5.2
Jungermannia polyrhiza Hook. ex Lehm. & Lindenb. = 79.9
Jungermannia polyrhizoides Grolle ex Amakawa = 79.6
Jungermannia stricta (Schiffn.) Steph. = 79.7
Jungermannia subulata A. Evans = 65.1
Jungermannia tetragona Lindenb. = 79.8
Jungermannia truncata Nees = 79.9
Lejeunea anisophylla Mont. 35.1
Lejeunea apiculata Sande Lac 35.2
Lejeunea applanata (Reinw., Blume & Nees) Nees = 39.1
Lejeunea borneensis Steph. = 35.1
Lejeunea ceylanica Gottsche = 29.1
Lejeunea cocoes Mitt. 35.3
Lejeunea dipterota Eifrig 35.4
Lejeunea discreta Lindenb. 35.5
Lejeunea epiphylla Mitt. = 37.2
Lejeunea flava (Sw.) Nees 35.6
Lejeunea flavida Mitt. 35.7
Lejeunea gardneri Mitt. = 29.4
Lejeunea gracillima Mitt. = 42.1
Lejeunea haskarliana Lehm. & Lindenb. = 30.13
Lejeunea herzogii Mizut. 35.8
Lejeunea inflectens Mitt. = 30.16
Lejeunea lindenbergii Gottsche = 29.8
Lejeunea lowriana Steph. 35.9
Lejeunea macrodonta Mitt. = 33.4
Lejeunea maculata Mitt. = 37.3
Lejeunea micholitzii Mizut. 35.10
Lejeunea neelgherriana Gottsche 35.11
Lejeunea nietneri (Steph.) Steph 35.12
Lejeunea nietneri Gottsche nom. nud. ? = 35.12
Lejeunea obfusca Mitt. 35.13
Lejeunea obscura Mitt. 35.14
Lejeunea papilionacea Steph. 35.15
Lejeunea peradeniensis Mitt. = 45.3
Lejeunea planissima Mitt. = 30.25
Lejeunea punctiformis Taylor = 42.1
Lejeunea pycnoclada (Taylor) Mitt. = 27.3
Lejeunea repleta Taylor = 40.3
Lejeunea riparia Mitt. = 35.16
Lejeunea semirepanda (Nees) Nees & Mont. = 46.2
Lejeunea serpentina Mitt. = 29.11
Lejeunea spathulistipa (Reinw., Blume & Nees) Mitt. = 48.2
Lejeunea subacuta Mitt. 35.17
Lejeunea subfusca (Nees) Nees & Mont. = 39.6
Lejeunea thwaitesiana Mitt. = 33.9
Lejeunea tuberculosa Steph. 35.18
Lejeunea utriculata (Steph.) Mizut. 35.19
Lejeunea vesiculosa Mitt. = 33.10
Lejeunea wightii Lindenb. 35.20
Lejeunea xanthocarpa (Lehm. & Lindenb.) Lehm. & Lindenb. = 38.1
Lepicolea yakusimensis (S.Hatt.) S.Hatt. 51.1
Lepidolejeunea bidentula (J.B.Jack & Steph.) R.M.Schust. 36.1
Lepidozia neesii Lindenb. = 57.1
Lepidozia plumula Herz. 54.1
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Lepidozia semperiana Steph. = 57.2
Lepidozia subintegra Lindenb. 54.2
Lepidozia wallichiana Gottsche = 55.1
Leptolejeunea elliptica (Lehm. & Lindenb.) Schiffn. 37.1
Leptolejeunea epiphylla (Mitt.) Steph. 37.2
Leptolejeunea fleischeri Steph. = 33.3
Leptolejeunea maculata (Mitt.) Schiffn. 37.3
Leptolejeunea massartiana Schiffn. ex Herzog 37.4
Leptolejeunea rhombifolia Steph. 37.5
Leptolejeunea subacuta Steph. = 37.1
Leptolejeunea vitrea (Nees) Schiffn. 37.6
Leucolejeunea xanthocarpa (Lehm. & Lindenb.) A.Evans 38.1
Liochlaena subulata (A. Evans) Schljakov 65.1
Lophocolea bidentata (L.) Dumort. 60.1
Lophocolea ciliolata (Nees) Gottsche = 58.1
Lophocolea cuspidata (Nees) Limpr. = 60.1
Lophocolea fleischeri Steph. = 58.2
Lophocolea javanica Schiffn. 60.2
Lophocolea kurzii Sande Lac. 60.3
Lophocolea muricata (Lehm.) Nees 60.4
Lophocolea peradeniensis Steph. = 60.3
Lopholejeunea applanata (Reinw., Blume & Nees) Schiffn. 39.1
Lopholejeunea arbortiva (Mitt.) Steph. = 39.2
Lopholejeunea ceylanica Steph. 39.3
Lopholejeunea eulopha (Taylor) Schiffn. 39.4
Lopholejeunea javanica (Nees) Schiffn. 39.5
Lopholejeunea longiloba Steph. = 39.3
Lopholejeunea nigricans (Lindenb.) Steph. 39.2
Lopholejeunea nipponica Horik. = 39.7
Lopholejeunea subfusca (Nees) Schiffn. 39.6
Lopholejeunea zollingeri (Steph.) Schiffn. 39.7
Lunularia cruciata (L.) Dumort. 62.1
Madotheca acutifolia Lehm. & Lindenb. = 73.1
Madotheca ciliaris Nees = 73.5
Madotheca nilgheriensis Mont. = 73.4
Madotheca perrottetiana Mont. = 73.5
Marchantia acaulis Steph. 63.1
Marchantia amboinensis Nees & Mont. = 63.2
Marchantia emarginata Reinw., Blume & Nees subsp. emarginata 63.2
Marchantia paleacea Bertol. 63.3
Marchantia palmata Nees = 63.2
Marchantia papillata Raddi subsp. grossibarba (Steph.) Bischl. 63.4
Marchantia pappeana Lehm. subsp. robusta (Steph.) Bischl. 63.5
Marchantia polymorpha L. 63.6
Marchantia robusta Steph. = 63.5
Marsupidium knightii Mitt. 1.1
Mastigobryum callidum Steph. = 52.1
Mastigobryum ceylanicum Mitt. = 52.2
Mastigobryum decurvum Nees = 52.11
Mastigobryum fleischeri Steph. = 52.4
Mastigobryum gaudichaudii Gottsche ex Steph. = 52.5
Mastigobryum grandistipulum Steph. = 52.2
Mastigobryum intermedium Mitt. hom. illeg. = 52.1
Mastigobryum lacerostipulum Steph. = 52.2
Mastigobryum obtusatum Mitt. = 52.7
Mastigobryum ovistipulum Steph. = 52.9
Mastigobryum praeruptum Reinw., Blume & Nees = 52.11
Mastigobryum rabenhorstii Steph. = 52.12
Revised checklist to the Liverworts and Hornworts of Sri Lanka
Mastigobryum rotundistipulum Steph. = 52.10
Mastigobryum sandei Steph. = 52.11
Mastigobryum strictum Steph. = 52.2
Mastigobryum vittatum Gottsche = 52.15
Mastigobryum wichurae Steph. = 52.10
Mastigolejeunea atypos (Schiffn.) Steph. = 40.3
Mastigolejeunea auriculata (Wilson) Schiffn. 40.1
Mastigolejeunea humilis (Gottsche) Schiffn. 40.2
Mastigolejeunea humilis (Gottsche) Schiffn. var. ciliata Awasthi & Udar 40.2a
Mastigolejeunea repleta (Taylor) A.Evans 40.3
Mastigolejeunea virens (Ångstr.) Steph. 40.4
Mastigophora aequifolia Steph. 64.1
Mastigophora diclados (Brid. ex F.Weber) Nees 64.2
Metacalypogeia alternifolia (Nees) Grolle 14.1
Metalejeunea cucullata (Reinw., Blume & Nees) Grolle 41.1
Metzgeria borneensis Kuwah. = 66.8
Metzgeria ciliata Raddi 66.1
Metzgeria conjugata Lindb. 66.2
Metzgeria consanguinea Schiffn. 66.3
Metzgeria crassipilus (Lindb.) A.Evans 66.4
Metzgeria decipiens (C.Massal.) Schiffn. & Gottsche = 66.1
Metzgeria foliicola Schiffn. 66.5
Metzgeria latifrons Steph. 66.6
Metzgeria lechleri Steph. 66.7
Metzgeria leptoneura Spruce 66.8
Metzgeria lindbergii Schiffn. = 66.2
Metzgeria luzonensis Kuwah. = 66.8
Metzgeria papulosa Steph. = 66.8
Metzgeria subhamata S.Hatt. = 66.8
Metzgeria vivipara A.Evans = 66.7
Microlejeunea cucullata (Reinw., Blume & Nees) J.B.Jack & Steph. = 41.1
Microlejeunea gracillima (Mitt.) Carringt. & Pearson = 42.1
Microlejeunea punctiformis (Taylor) Spruce = 42.1
Mnioloma fuscum (Lehm.) R.M.Schust. 15.1
Neolepidozia wallichiana (Gottsche) Fulford & J.Taylor 55.1
Notoscyphus lutescens (Lehm. & Lindenb.) Mitt. 67.1
Notoscyphus paroicus Schiffn. = 67.1
Nowellia curvifolia (Dicks.) Mitt. 16.1
Pallavicinia decipiens (Mitt.) Steph. = 68.1
Pallavicinia indica Schiffn. 69.1
Pallavicinia lyellii (Hook.) Gray 69.2
Phaeoceros carolinianus (Michx.) Prosk. 85.1
Phaeoceros laevis (L.) Prosk. subsp. carolinianus (Michx.) Prosk. = 85.1
Phragmicoma ceylanica Gottsche nom. nud. ? = 39.3
Phragmicoma nietneri Gottsche nom. nud. ? = 43.1
Physcocolea planissima (Mitt.) Steph. = 30.25
Physcocolea serrata Steph. = 30.30
Physiotium acinosum Mitt. = 72.1
Physocolea gottschei Steph. = 30.11
Physocolea inflectens (Mitt.) Steph. = 30.16
Physotium giganteum F.Weber = 72.2
Physotium microcarpum J.B.Jack = 72.3
Physotium sphagnoides A.Rich. ex Nees, nom. illeg. = 72.2
Plagiochasma nepalense (Lehm. & Bisch.) Steph. = 10.1
Plagiochasma rupestre (G.Forst.) Steph. 10.1
Plagiochila acanthophylla Gottsche = 70.18
Plagiochila aciculifera Steph. = 70.14
Plagiochila acutiloba Inoue = 70.8
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Long and Rubasinghe
Plagiochila aequalis Mitt. = 70.17
Plagiochila arbuscula (Lehm. & Lindenb.) Lindenb. 70.1
Plagiochila bantamensis (Reinw., Blume & Nees) Mont. 70.2
Plagiochila belangeriana Lindenb. = 70.1
Plagiochila brauniana Nees = 71.1
Plagiochila cadens Inoue = 70.18
Plagiochila ceylanica Mitt. 70.3
Plagiochila devexa Steph. 70.4
Plagiochila exigua (Taylor) Taylor 70.5
Plagiochila firma Mitt. = 70.9
Plagiochila flexuosa Mitt. 70.6
Plagiochila frondescens (Nees) Lindenb. 70.7
Plagiochila ghatiensis Steph. 70.8
Plagiochila gracilis Lindenb. & Gottsche 70.9
Plagiochila gymnoclada Sande Lac. 70.10
Plagiochila integrilobula Inoue 70.11
Plagiochila khasiana Mitt. 70.12
Plagiochila kurzii Steph. 70.13
Plagiochila lobulata Schiffn. = 70.2
Plagiochila macrantha Steph. = 70.3
Plagiochila meijeri Inoue = 70.13
Plagiochila microdonta Mitt. 70.14
Plagiochila neutiana Steph. = 70.3
Plagiochila onraedtii Inoue = 70.2
Plagiochila opposita (Reinw., Blume & Nees) Lindenb. = 71.2
Plagiochila parvidentata Inoue = 70.19
Plagiochila parvifolia Lindenb. 70.15
Plagiochila peradenyensis Schiffn. 70.16
Plagiochila pseudoventricosa Inoue = 70.15
Plagiochila ramenticaulis Steph. = 70.16
Plagiochila renitens (Nees) Lindenb. 70.17
Plagiochila rufa Steph. = 70.6
Plagiochila sciophila Nees ex Lindenb. 70.18
Plagiochila securifolia Nees ex Lindenb. = 4.1
Plagiochila sisparensis Steph. 70.19
Plagiochila variegata Lindenb. = 4.1
Plagiochila ventricosa Steph. = 70.15
Plagiochila wichurae Steph. = 70.17
Plagiochila zygophylla Taylor = 71.2
Plagiochilion braunianum (Nees) S.Hatt. 71.1
Plagiochilion oppositum (Reinw., Blume & Nees) S.Hatt. 71.2
Pleuroschisma wichurae (Steph.) Reimers = 52.10
Pleurozia acinosa (Mitt.) Steph. 72.1
Pleurozia gigantea (F.Weber) Lindb. 72.2
Pleurozia microcarpa (J.B.Jack) Steph. = 72.3
Pleurozia subinflata (Austin) Austin 72.3
Plicanthus birmensis (Steph.) R.M.Schust. 6.1
Plicanthus hirtellus (F.Weber) R.M.Schust. 6.2
Porella acutifolia (Lehm. & Lindenb.) Trevis. 73.1
Porella acutifolia (Lehm. & Lindenb.) Trevis. subsp. latior S.Hatt. = 73.2a
Porella acutifolia (Lehm. & Lindenb.) Trevis. subsp. latior S.Hatt. var. intermedia S.Hatt. 73.2b
Porella caespitans (Steph.) S.Hatt. subsp. latior (S.Hatt.) S.Hatt. var. intermedia (S.Hatt.) S.Hatt. 73.2b
Porella caespitans Steph. subsp. latior (S.Hatt.) S.Hatt. 73.2a
Porella japonica (Sande Lac.) Mitt. 73.3
Porella madgascariensis (Nees & Mont.) Trevis. 73.4
Porella perottetiana (Mont.) Trevis. 73.5
Ptychanthus birmensis Steph. = 43.1
Ptychanthus effusus Steph. = 43.1
Revised checklist to the Liverworts and Hornworts of Sri Lanka
Ptychanthus nietneri (Steph.) Schiffn. = 43.1
Ptychanthus striatus (Lehm. & Lindenb.) Nees 43.1
Ptychanthus striatus (Lehm. & Lindenb.) Nees var. retusus (Reinw., Blume & Nees) Verd. = 43.1
Ptychocoleus cumingianus auct. non (Mont.) Trevis. = 45.3
Ptychocoleus peradeniensis (Mitt.) Steph. = 45.3
Ptychocoleus pulopenangensis (Gottsche) Trevis. = 45.2
Ptychocoleus pycnocladus (Taylor) Steph. = 27.3
Ptycholejeunea nietneri Steph. = 43.1
Pycnolejeunea badia Steph. = 36.1
Pycnolejeunea ceylanica (Gottsche) Schiffn. = 29.1
Pycnolejeunea flavida (Mitt.) Steph. = 35.7
Pycnolejeunea gardneri (Mitt.) Steph. = 29.4
Pycnolejeunea grandiocellata Steph. 44.1
Pycnolejeunea trapezia (Nees) Steph. = 29.14
Pycnolejeunea utriculata Steph. = 35.19
Radula acuminata Steph. 74.1
Radula amentulosa Mitt. 74.2
Radula assamica Steph. 74.3
Radula ceylanica K.Yamada 74.4
Radula colliculosa Mitt. = 74.7
Radula cordata Mitt. 74.5
Radula formosa (Meissn.) Nees 74.6
Radula javanica Gottsche 74.7
Radula kurzii Steph. 74.8
Radula madagascariensis Gottsche 74.9
Radula nietneri Steph. = 74.7
Radula nymanii Steph. 74.10
Radula obscura Mitt. 74.11
Radula onraedtii K.Yamada 74.12
Radula pinnulata Yang nom. illeg. = 74.19
Radula retroflexa Taylor 74.13
Radula sandei Steph. 74.14
Radula speciosa Gottsche ex Steph. = 74.8
Radula subpallens Steph. 74.15
Radula tabularis Steph. 74.16
Radula tjibodensis K.I.Goebel 74.17
Radula ventricosa Steph. 74.18
Radula yangii K.Yamada 74.19
Reboulia hemisphaerica (L.) Raddi 11.1
Rectolejeunea riparia (Mitt.) Steph. = 35.16
Rhaphidolejeunea fleischeri (Steph.) Herzog = 33.3
Riccardia canaliculata (Nees) Schiffn. 9.1
Riccardia multifida (L.) Gray 9.2
Riccardia pinguis (L.) Gray = 8.1
Riccia attenuata Pandé 75.1
Riccia billardieri Mont. & Nees 75.2
Riccia crispatula Mitt. 75.3
Riccia discolor Lehm. & Lindenb. 75.4
Riccia huebeneriana Lindenb. 75.5
Riccia sorocarpa Bisch. 75.6
Scapania lepida Mitt. 77.1
Schiffneriolejeunea haskarliana (Lehm. & Lindenb.) Gradst. = 30.13
Schiffneriolejeunea hasskarliana (Gottsche) Gradst. = 45.3a
Schiffneriolejeunea polycarpa (Nees) Gradst. 45.1
Schiffneriolejeunea pulopenangensis (Gottsche) Gradst. 45.2
Schiffneriolejeunea tumida (Nees) Gradst. 45.3
Schiffneriolejeunea tumida (Nees) Gradst. var. hasskarliana (Gottsche) Gradst. & Terken 45.3a
Schisma ceylanicum Steph. = 23.1
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36
Long and Rubasinghe
Schisma perrottetii Steph. = 23.2
Schisma wichurae Steph. = 23.2
Schistochila aligera (Nees & Blume) Jack & Steph. 78.1
Schistochila commutata Steph. = 78.1
Schistochila fleischeri Steph. = 78.1
Schistochila philippinensis (Mont.) J.B.Jack & Steph. = 78.1
Schistochila sciurea (Nees) Schiffn. 78.2
Schistochila thwaitesii (Mitt.) Steph. = 78.2
Schizophyllopsis bidens (Reinw., Blume & Nees) Váňa & L. Söderstr. 7.1
Sendtnera diclados (Brid. ex F.Weber) Gottsche, Lindenb. & Nees = 64.2
Sendtnera dicrana Taylor ex Gottsche, Lindenb. & Nees = 23.2
Solenostoma appressifolium (Mitt.) Váňa & D. G. Long 79.1
Solenostoma ariadne (Taylor ex Lehm.) R. M. Schust ex Váňa & D. G. Long 79.2
Solenostoma borneense (Amakawa) Váňa, Hentschel & J. Heinrichs 79.3
Solenostoma hasskarlianum (Nees) R.M.Schust. ex Váňa & D.G.Long 79.4
Solenostoma javanicum (Schiffn.) Steph. 79.5
Solenostoma polyrhizoides (Grolle ex Amakawa) Váňa & D. G. Long 79.6
Solenostoma strictum (Schiffn.) Váňa, Hentschel & J.Heinrichs 79.7
Solenostoma tetragonum (Lindenb.) R. M. Schust. ex Váňa & D. G. Long 79.8
Solenostoma truncatum (Nees) R. M. Schust. ex Váňa & D. G. Long 79.9
Southbya grollei N.Kitag. = 80.1
Southbya organensis Herzog 80.1
Spruceanthus marianus (Gottsche) Mizut. 46.1
Spruceanthus semirepandus (Nees) Verd. 46.2
Steetzia decipiens Mitt. = 68.1
Steetzia lyellii (Hook.) Lehm. = 69.2
Stenolejeunea apiculata (Sande Lac.) R.M.Schust. = 35.2
Syzygiella securifolia (Nees ex Lindenb.) Inoue 4.1
Syzygiella subintegerrima (Nees) Spruce 4.2
Syzygiella variegata (Lindenb.) Schiffn. = 4.1
Taeniolejeunea pseudofloccosa (Horik.) S.Hatt. = 30.28
Targionia hypophylla L. 81.1
Taxilejeunea apiculata (Sande Lac.) Eifrig nom. illeg. = 35.2
Taxilejeunea compressiuscula Lindenb. ex Steph. 47.1
Telaranea major (Herz.) J.J.Engel & G.L.Merr. 56.1
Telaranea semperiana (Steph.) Del Ros. = 57.2
Telaranea wallichiana (Gottsche) R.M.Schust. = 55.1
Thysananthus convolutus Lindenb. 48.1
Thysananthus spathulistipus (Reinw., Blume & Nees) Lindenb. 48.2
Trichocolea pluma (Reinw., Blume & Nees) Mont. 82.1
Tricholepidozia neesii (Lindenb.) E.D.Cooper 57.1
Tricholepidozia semperiana (Steph.) E.D.Cooper 57.2
Trocholejenea sandvicensis (Gottsche) Mizut. 49.2
Trocholejeunea infuscata (Mitt.) Verd. 49.1
Tylimanthus indicus Steph. = 70.8
Xenolejeunea longiloba (Steph. ex G.Hoffm.) Kachroo & R.M.Schust. 50.1
Revised checklist to the Liverworts and Hornworts of Sri Lanka
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