Bryological Note New national and regional bryophyte records, 33 L T Ellis1, H Bednarek-Ochyra2, R Ochyra2, B Cykowska2, M V Dulin3, T Ezer4, R Kara4, J R Flores5, G M Suárez5, C Garcia6, A Martins6, C Sérgio6, R Garilleti7, M Kırmacı8, E Agcagil8, L E Kurbatova9, M Lebouvier10, B Papp11, E Szurdoki11, D A Philippov12, V Plášek13, T Pócs14, M Sabovljević15, J Sawicki16, M Sim-Sim17, P Szücs18, A Bidló18, J Váňa19, B Vigalondo20, F Lara20, I Draper20, V M Virchenko21, G J Wolski22 1 Department of Life Sciences, The Natural History Museum, London, UK, 2Laboratory of Bryology, Institute of Botany, Polish Academy of Sciences, Krakow, Poland, 3Institute of Biology, Komi Science Centre UB RAS, Komi, Russia, 4Niğde University, Faculty of Science, Department of Biology, Niğde, Turkey, 5Facultad de Ciencias Naturales e Institución Miguel Lillo, Tucumán, Argentina, 6Universidade de Lisboa, Museu Nacional de História Natural e da Ciência, Lisboa, Portugal, 7Departamento de Botánica, Facultad de Farmacia, Universidad de Valencia, Burjasot, Spain, 8Adnan Menderes Üniversitesi, Fen Edebiyat Fakültesi, Biyoloji Bölümü, Kepez-Aydın, Turkey, 9Komarov Botanical Institute, Russian Academy of Sciences, Moscow, Russia, 10CNRS UMR 6553, Université de Rennes 1, France, 11Botanical Department, Hungarian Natural History Museum, Budapest, Hungary, 12 I.D. Papanin Institute for Biology of Inland Water RAS, Nekouz, Russia, 13Department of Biology and Ecology, University of Ostrava, Czech Republic, 14Department of Botany, Eszterházy Károly College, Hungary, 15 Institute of Botany and Botanical Garden, Faculty of Biology, University of Belgrade, Serbia, 16Department of Botany and Nature Protection, University of Warmia and Mazury in Olsztyn, Poland, 17Museu Nacional de História Natural, Jardim Botânico, Lisboa, Portugal, 18Department of Forest Site Diagnosis and Classification, University of West Hungary, Sopron, Hungary, 19Department of Botany, Charles University, Praha, Czech Republic, 20Departamento de Biologı́a (Botánica), Universidad Autónoma de Madrid, Spain, 21Department of Lichenology and Bryology, Institute of Botany, National Academy of Sciences of the Ukraine, Kiev, Ukraine, 22 Deptartment of Geobotany and Plant Ecology, University of Łódź, Poland 1. Anastrophyllum michauxii (F.Weber) H.Buch Contributors: M. V. Dulin and D. A. Philippov Russian Federation: Vologda Region, Vytegra district, 23 km north of the settlement of Yanishevo, Andomskaya height, 5 km to south-east of Soydozero Lake, the Soyda River bank, 61u26972.10N, 37u33936.80E, ca 205 m a.s.l., in fern and herb rich spruce forest, on rotten spruce log, with Blepharostoma trichophyllum (L.) Dumort., Riccardia latifrons (Lindb.) Lindb., Crossocalyx hellerianus (Nees ex Lindenb.) Meyl., Calypogeia muelleriana (Schiffn.) Müll.Frib., Lophozia guttulata (Lindb. & Arnell) A.Evans, and Ptilidium pulcherrimum (Weber) Vain., plants with antheridia, 17 August 2010, leg. D.A. Philippov, 10783a (IBIW, SYKO). This is the first report of Anastrophyllum michauxii from the Vologda Region. It is a montane subcircumpolar liverwort species. The species is known in several European countries (Austria, Bulgaria, Correspondence to: Leonard Thomas Ellis, Department of Life Sciences, The Natural History Museum, London. Email: l.ellis@nhm.ac.uk ß British Bryological Society 2012 DOI 10.1179/1743282012Y.0000000030 France, the Czech Republic and Slovakia, Germany, Hungary, the former Yugoslavia, Italy, Romania, Poland, Finland, Sweden, Norway), in Asia (China, Japan) and in a few sites in North America (Damsholt, 2002; Schumacker & Váňa, 2005). In Russia, A. michauxii has been found in mountainous areas: Northern and Middle Urals, Caucasus, Stanovoye Nagor’e, Khamar-Daban, Kuznetskij Alatau, Sayan Mountains, Altai, Sikhote-Alin and also in mountains of Kamchatka, Sakhalin and Kuril Islands (Konstantinova et al., 2009). Findings of the species in the lowland areas are rare, but include records from the Finland Gulf Islands (Potemkin et al., 2008), the Russian Plain and Pechora lowlands (Dulin, 2007). 2. Crossidium aberrans Holz. & E.B.Bartram Contributors: M. Kırmacı and E. Agcagil Turkey: Province Aydın, İmamköy, deep valley, 37u529N, 27u549E, 150 m a.s.l., on soil covering rock, associated with Aloina aloides (Koch ex Schultz) Kindb., Bryum argenteum Hedw., Didymodon luridus Hornsch., D. australasiae (Hook. & Grev.) Journal of Bryology 2012 VOL . 34 NO . 4 281 Bryological notes R.H.Zander, Fissidens bryoides Hedw., 2 March 2008, leg. & det. M. Kırmacı & E. Agcagil, conf. B. Papp (AYDN No. 2484). The new record was collected in an area characterized by degraded maquis (dominated by Quercus coccifera L.), plantation of Olea europaea L. and a few stands of Pinus brutia Ten. remaining after clearance by locals for arable land. The area has a Mediterranean climate, with an average annual precipitation of 672 mm (Aydın Province Environment Situation Report, 2006). The moss was collected from very dry habitats exposed to sun, where the high irradiation results in strong desiccation during dry periods that last nearly seven months in a year. It is well known that plants in the genus Crossidium Jur. are adapted to mainly arid and semiarid areas (Zander & Hedderson, 2011). Crossidium includes some 12 species world-wide, and according to Kürschner and Frey (2011) ten taxa (9 species and 1 variety) are known from southwest Asia. Among these, C. squamiferum (Viv.) Jur. var. squamiferum, C. squamiferum var. pottioideum (De. Not.) Mönk., C. crassinerve (De Not.) Jur. and C. laxefilamentosum Frey & Kürschner were previously recorded from Turkey. Crossidium aberrans is easily distinguished from other species of this genus by the cells and ventral filaments of the leaves. The filaments are 1–2 cells high, subglobose and densely papillose (Bai, 2002). It shows a xerothermic Pangaean distribution pattern, related to a former Permo-Triassic continental Pangaean range (Frey & Kürschner, 1988), and has been recorded from North America, Europe, China, North Africa and southwest Asia (Syria, Israel, Saudi Arabia, and Sinai Peninsula) (Bai, 2002; Chavoutier & Hugonnot, 2006; Kürschner & Frey, 2011). This new record from Turkey is not surprising, it completes the distribution pattern for the species between southern Europe and southwest Asia, and raises the number of taxa in Crossidium known to occur in Turkey from four to five (four species and one variety). 3. Dichelyma capillaceum (Dicks.) Myrin Contributors: V. M. Virchenko and R. Ochyra Ukraine: Zhytomyr Province, Ovruch district, Selezivske forestry, in the valley of the Bolotnytsya river near the village of Selezivka, ca 51u329N, ca 28u079E; on bark of willow; 22 April 1973, leg. L. Ya. Partyka s.n. (KRAM, KW). In 197221973, L. Ya. Partyka studied the moss flora of the Polissya Nature Reserve in the northwestern part of Zhytomyr Province in the Ukraine which was established for the protection of the best preserved fragments of the largest European wetland, especially peat bogs and bog pinewoods. Altogether she reported 139 species of moss from the study area 282 Journal of Bryology 2012 VOL . 34 NO . 4 (Partyka, 1974, 1986). A revision of the voucher collections revealed that one specimen she had determined as Sanionia uncinata (Hedw.) Loeske included, as an admixture, a plant she had provisionally named as Drepanocladus (Müll.Hal.)G.Roth. Actually, it represents Dichelyma capillaceum, a species hitherto unknown from the Ukraine. The moss was collected from the bark of willow in a flooded area of the Bolotnytsya river valley. It lacked sporophytes but the male gametangia were present. In continental Europe D. capillaceum is widely distributed, but scattered and localised, and so has been recorded from France, Holland, Germany, Denmark, Poland, Sicily, and Greece (Düll, 1985). However, it has not been seen for nearly a century in most of these countries. The most recent discovery of D. capillaceum in Central Europe was made in 1997 in Rheinland, Germany (Frahm & Stapper, 1998), where the species was refound at a locality originally discovered in 1920 and 1923 (Brasch, 1923). For six subsequent decades the species had not been rediscovered and was considered to be extinct at this site (Düll, 1980). Therefore D. capillaceum is included in Appendix I of the Bern Convention and in Annex 2 of the EC Habitats and Species Directive as a vulnerable species (Martiny, 1995). The information on the occurrence of the species in Romania provided by Düll (1985) and Martiny (1995) was apparently erroneous since the species has not been recorded in the latest catalogue of mosses for this country (Mohan, 1998). However, in 1978 D. capillaceum was recorded from the Munţii Retezat in the Southern Carpathians in Romania (Blockeel et al., 2003). In Europe, D. capillaceum has the main centre of its occurrence in Fennoscandia, where it is particularly frequent in southern Sweden (Hedenäs et al., 1996; Hallingbäck, 1998; Hylander, 1998). In contrast, the species is widely scattered but localised in southern and central Finland (Toivonen, 1972) and it is considered to be endangered in this country (Sallantaus, 2009). The easternmost European station of D. capillaceum is in Russian Karelia in Leningrad Province, but it was found at this site only in 1914 (Czernyadjeva, 2002). However, in 1975 and 1995 the species was discovered at two stations in Estonia (Kannukene et al., 1997; Ingerpuu & Vellak, 2000) and this discovery, along with that in the Ukrainian Polissya, now represents the easternmost occurrences of the species in Europe. Dichelyma capillaceum has maximum occurrence in eastern North America where it ranges from south-eastern Canada in the north (Ireland, 1990) to Florida in the south and Missouri and Arkansas in the west (Welch, 1960). The species was recently found in West Siberia in the middle course of the Ob river area (Czernyadjeva, 2002). This discovery changes the phytogeographical status of D. capillaceum a little. However, it should be Bryological notes still considered as a Euro-North American borealtemperate species weakly penetrating into westernmost boreal Asia. 4. Ditrichum hyalinum (Mitt.) Kuntze Contributors: L. E. Kurbatova and R. Ochyra South Shetland Islands, Nelson Island: (1) Stansbury Peninsula between Edgell Bay and Fildes Strait, in the central valley, 62u149S, 59u009W; 37 m a.s.l.; forming small patches, 223 cm dia., on soil in depressions and cracks in rock, together with Andreaea gainii Cardot, Bartramia patens Brid., Hymenoloma antarcticum (Müll.Hal.) Ochyra, Meesia uliginosa Hedw. and Pohlia cruda (Hedw.) Lindb. in a lichen and moss community, 20 February 2009, leg. L. E. Kurbatova L101-7 (KRAM, LE): (2) same locality, in the flat range near the glacier, on the top of a rock outcrop, 62u149S, 59u019W, 110 m a.s.l., in cracks in soil, associated with Andreaea gainii, A. regularis Müll.Hal., Bartramia patens, Hymenoloma antarcticum, Meesia uliginosa, Pohlia cruda, Polytrichastrum alpinum (Hedw.) G.L.Sm., Schistidium antarctici (Cardot) L.I.Savicz & Smirnova and Syntrichia filaris (Mül.Hal.) R.H.Zander., 20 February 2009, leg. L. E. Kurbatova L100-12 (KRAM, LE). Ditrichum hyalinum is the most frequent and abundant of five species of the genus Ditrichum Hampe in the Antarctic (Ochyra & Lewis Smith, 1998; Ochyra et al., 2008a). For a long time the species was better known under the name D. austrogeorgicum Cardot, but it proved to be synonymous with D. hyalinum (Ochyra, 1999). In the Antarctic biome the species is restricted in its distribution to two periAntarctic archipelagos, namely the South Orkney Islands and the South Shetland Islands situated in the northern part of the maritime Antarctic. There is also a single isolated station on Adelaide Island off the central Antarctic Peninsula (Ochyra et al., 2008b). In the South Shetland Islands D. hyalinum was recorded from almost all major islands, and here its distribution is extended to Nelson Island, a heavily glaciated island situated immediately to the south-west of King George Island, the largest island of the archipelago. 5. Ditrichum lineare (Sw.) Lindb. Contributors: B. Papp, E. Szurdoki and M. Sabovljević Serbia: SE Serbia, Mt Vrtop (Plana) at Okruglica (Vlasina lake), on soil near the peak, 42u47934.10N, 22u22923.60E, 1730 m a.s.l., 21 June 2011, leg. B. Papp s.n. (BP 183642). Ditrichum lineare is a small moss with a scattered distribution in the Northern Hemisphere. It has a subatlantic and montane distribution (Düll, 1984) and prefers acidic and disturbed soils. D. lineare has not been recorded very often in the Balkans. It has probably been overlooked owing to its small size, and there being relatively few appropriate habitats on acidic bedrock. The species can also be easily misidentified due to its significant variability (Frahm et al., 2008). D. lineare was collected on the top of a hill in the surroundings of Vlasina Lake. This region seems to be quite interesting bryologically, as around the lake can be found the biggest peat-bog in south-eastern Europe. It has quite a diverse geology, and a lot of foggy and wet days in an average year compared with other highland areas in the Balkans. Ellis et al. (2012a), already records a novelty for the Serbian bryophyte flora from this area. In Southeast Europe, D. lineare is known only from Bulgaria, Romania and Slovenia (Sabovljević et al., 2008), but could be expected elsewhere with further investigation. 6. Gymnomitrion subintegrum (S.W.Arnell) Váňa Contributors: J. Váňa, R. Ochyra, H. BednarekOchyra, B. Cykowska and M. Lebouvier Île Amsterdam: Central part, Mont de la Dives, south of the summit, 37u5199.6910S, 77u32950.3990E, 748 m a.s.l., on rock, 22 December 2006, leg. M. Lebouvier RO-06-Dives (KRAM). The Palaeotropical montane species Gymnomitrion subintegrum was originally described as Marsupella subintegra S.W.Arnell from the Ruwenzori Mountains of Uganda in Central Africa (Arnell, 1956). The species is characterised by the lack of a perianth and Váňa et al. (2010b) transferred it to the genus Gymnomitrion Corda. This taxonomic decision was supported by the results of molecular studies (Vilnet et al., 2007) which showed that the development or reduction of the perianth and perigynium is an important character in the classification of the Gymnomitriaceae. The species has a bicentric distribution in the Old World. It is widely scattered in Central and East Africa, including in Rwanda (Váňa et al., 1979) and Malawi (Wigginton & Porley, 2001) (but excluding Uganda) and extends to Réunion Island in the Indian Ocean (Váňa, 1985). The second centre of its occurrence is in Malesia where it was first recorded from Mt. Kinabalu as Marsupella integra N.Kitag. (Kitagawa, 1967). Subsequently, it was found on Mt. Wilhelm and Mt. Hagen in New Guinea and on Mt. Korinchi in Sumatra (Váňa, 1976). Additionally the species has penetrated into the temperate zone, where it was found in Nepal in the Himalaya (Hattori, 1975). Herein it is recorded from Île Amsterdam, which is situated in the south-cooltemperate zone in the South Indian Ocean. The hepatic flora of this highly isolated, oceanic volcanic island is still poorly known, and a decade ago merely 15 species of hepatics were reported (Grolle, 2002). However, recent field studies yielded six additional species for the island’s liverwort flora (Blockeel et al., 2009b; Váňa et al., 2010a; Ellis et al., 2012b), so including the present record it consists of 22 species. 7. Metzgeria myriopoda Lindb. Contributors: J. R. Flores and G. M. Suárez. Journal of Bryology 2012 VOL . 34 NO . 4 283 Bryological notes Uruguay: Cerro Largo, Sierra de Rı́o, 32u11934.10S, 53u51953.20W, 309 m a.s.l., próximos al curso de agua del arroyo, 1 March 2012, leg. G. Suárez 1420 (LIL). As with the mosses, the liverwort flora of Uruguay is poorly known and large areas have not been explored or studied (Ellis et al., 2011b, 2012b). According to Hässel de Menéndez & Rubies (2009) only 30 species of Marchantiophyta are recorded from Uruguay, and the genus Metzgeria Raddi is not represented. Metzgeria myriopoda was identified during the examination of a recent collection of bryophytes from Uruguay. This is an American species, reported in South America from Equador, Peru, Brazil, Bolivia, Argentina and Paraguay (Kuwahara, 1986; Nieva & Schiavone, 2002; Pinheiro da Costa, 2008). In Uruguay, M. myriopoda occured in ‘Cerro Largo’; this is a temperate and warm grassland area with some forested zones where exotic, and a few native, tree species are found. It characteristically has precipitation rates of around 1100–1200 mm/year. Taking into account the climatic regime in Cerro Largo, and those prevailing in the localities of previous records, there appears to be an extensive range of environmental conditions under which M. myriopoda can grow. It occurs on diverse substrates (as an epiphyte, including on bark, and on rocks and soil) in environments with a wide range of temperatures and degrees of disturbance. The examination of M. myriopoda revealed that its characters are highly constant, with 2–8 dorsal epidermal cells and 4–6 ventral epidermal cells in the midrib; convex wings with short, straight geminate hairs, and abundant marginal gemmae. This note constitutes the first record for the genus Metzgeria in Uruguay, as well as extending the distributional range of M. myriopoda. 8. Notoligotrichum trichodon (Hook. & Wilson) G.L.Sm. Contributors: H. Bednarek-Ochyra, R. Ochyra and M. Lebouvier Île Amsterdam: l37u509S, 77u319E, without locality data, 13 March 1970, leg. Noel s.n. (KRAM, PC). Since publication of the first account of mosses on Île Amsterdam (Bescherelle, 1875), information on the moss flora of this remote and highly isolated island, situated in the middle of the South Indian Ocean in the south-cool-temperate zone, has not increased markedly. The only other account of the mosses was published by Tixier (1980) and recently two additional species were added to the island’s flora, namely Ptychomnium densifolium (Brid.) A.Jaeger and Racomitrium lanuginosum (Hedw.) Brid. (Blockeel et al., 2009a). A revision of voucher specimens reported by Tixier (1980) for a planned survey of the moss flora of Île Amsterdam revealed that a specimen 284 Journal of Bryology 2012 VOL . 34 NO . 4 determined as Polytrichum formosum Hedw. actually represented Notoligotrichum trichodon. The plants were in fine fruiting condition; hence the identification of the material did not pose any problems. The characteristic gibbous capsule lacking an apophysis and narrowed at the oblique mouth, making the genus immediately distinct from all polytrichalean congeners. Notoligotrichum trichodon is an amphiatlantic subantarctic species which has the main centre of its occurrence on South Georgia from where it extends eastwards to Îles Kerguelen and westwards to Tierra del Fuego (Ochyra et al., 2008b). Additionally, it penetrates well into the Neotropics along the Andean chain, reaching to the northern Andes of Ecuador and Colombia from whence it was originally described (Ochyra, 2003). 9. Orthotrichum affine Bird. var. bohemicum Plášek & Sawicki Contributors: V. Plášek, G. J. Wolski and J. Sawicki Poland: Western Pomerania, Czersk district, near Brda village, base of Salix fragilis L., GPS coordinates (WGS 84): 53u419540N, 17u499410E, ca 140 m a.s.l., 16 July 1973, leg. E. Filipiak, det. V. Plášek & G.J. Wolski (March 2012) (LOD, #00013186). Orthotrichum affine var. bohemicum is an epiphytic moss from the Czech Republic, recently described by Plášek et al. (2011). The taxon has been noted in five localities in the Czech Republic, but has not yet been reported elsewhere in Europe. One specimen collected by D.H. Vitt in 1976 from the U.S.A., was recorded during a revision of the herbarium collections in St Petersburg (LE) (Ellis et al., 2012c). It can be expected that this variety will, before long, be found in other territories, or through the revision of other herbarium collections. Orthotrichum affine var. bohemicum is easily distinguished from the type variety mainly by the possession of 16 endostome segments which are practically as long as the exostome teeth (not alternately longer and shorter as in O. pallens Brid.). According to the herbarium label for this new record, the moss cushion was collected from the base of Salix fragilis, together with Brachythecium rutabulum (Hedw.) Schimp. and Ceratodon purpureus (Hedw.) Brid. 10. Orthodontium lineare Schwägr. Contributors: P. Szücs and A. Bidló Hungary: Komárom-Esztergom County, Dunaalmás, northern foothills of Gerecse Mts, 47u42959.50N, 18u19922.60E, ca 180 m a.s.l. on pine log (ca 40 cm dia.) in advanced state of decay, in planted old Pinus forest stand, associated with Bryum capillare Hedw., B. moravicum Podp., Campylopus introflexus (Hedw.) Brid., Dicranum montanum Hedw., D. scoparium Hedw., Herzogiella seligeri (Brid.) Z.Iwats, Hypnum Bryological notes cupressiforme Hedw. and Leucobryum glaucum (Hedw.) Ångstr., 14 January 2012, leg. & det. P. Szücs, conf. B. Papp. (BP, P. Szücs private herb.). Orthodontium lineare is a non-native species in Europe (Hill et al., 2006), originating from the southern hemisphere. It was first recorded in Europe from Britain in 1910 (Burrell, 1940). Since then, it has been spreading continuously, and about 30 years ago was detected in Central Europe (Hassel & Söderström, 2005). Orthodontium lineare has reached the following countries in the eastern part of Central Europe: Czech Republic (Kucěra & Vánă, 2003), Slovakia (Hassel & Söderström, 2005), Poland (Ochyra, 1982) and Russia (Kaliningrad Province) (Ignatov et al., 2006). No reports of the species have been published to date from the other countries bordering on Hungary (Austria: Grims, 1999; Slovenia, Croatia, Serbia, Romania: Sabovjević et al., 2008; Ukraine: Ignatov et al., 2006). This neophytic species is not included in the recent checklist of the moss flora of Hungary (Papp et al., 2010), and the present report is the first for the country. The first Hungarian report of Campylopus introflexus was found in the same stand of forest, on exactly the same log (Blockeel et al., 2007b). In Hungary, where sandy, peaty banks and siliceous cliffs are rare, decaying wood (Pinus and Quercus spp.) generally provides a suitable substrate for the species. After reaching Hungary, Orthodontium lineare may continue its expansion towards the south and east. The red list status of this new member of the Hungarian bryophyte flora is yet uncertain, but it might be assumed to be moderately spreading. 11. Plagiochila asplenioides (L. emend. Taylor) Dumort. Contributors: M. Sim-Sim, C. A. Garcia, A. Martins and C. Sérgio Portugal: Trás-os-Montes e Alto Douro: Bragança, entre Espinhosela e Parâmio. 29TPG7879039714, 850 m a.s.l., humid rocky slope, 31 January 2012, leg. C. A Garcia, M. Sim-Sim & S. Stow s.n. (LISU249899); Rebordãos. 29TPG7909622082, 979 m a.s.l., slope of the river, 8 May 2012, leg. C. Sérgio, C. A. Garcia, A. Martins & S. Stow s.n. (LISU249898). Plagiochila asplenioides was recently collected in the north-eastern part of Portugal. Portuguese collections formerly named as P. asplenioides had been revised, and their morphological characters found to accord with those of P. porelloides (Torr. ex Nees) Lindenb. (Sérgio et al., 2006; Sim-Sim, 1989). The newly discovered plants of P. asplenioides were large and formed vigorous turfs on a humid acidic slope close to a small stream and on the slopes of river banks, in an area dominated by well conserved Quercus pyrenaica Willd. woodland. The populations found were developing in sheltered, and more or less humid conditions, in a microenvironment dominated by Barbilophozia barbata (Schmidel ex Schreb.) Loeske, Metzgeria furcata (L.) Dumort., Pellia epiphylla (L.) Corda, Plagiochila porelloides, Radula complanata (L.) Dumort., Atrichum undulatum (Hedw.) P.Beauv., Bartramia pomiformis Hedw., Mnium stellare Hedw., Plagiothecium nemorale (Mitt.) A.Jaeger, Platyhypnidium riparioides (Hedw.) Dixon., Pohlia cruda (Hedw.) Lindb., Pseudoscleropodium purum (Hedw.) M.Fleisch. and Thamnobryum alopecurum (Hedw.) Gangulee. Plagiochila asplenioides seems to be rare in Portugal, restricted to sheltered woodland habitats supporting a high diversity of cryptogams and vascular plants. 12. Plagiothecium membranosulum Müll.Hal. Contributor: R. Ochyra, T. Pócs and H. BednarekOchyra Tanzania: (1) Kilimanjaro Mountains, Mweka Route, 3u119210S, 37u219080E, 190022600 m a.s.l., on rotten wood in montane rain forest dominated by Ocotea usambarensis Engl., 6 March 1985, leg. T. Pócs 6998/C (EGR, KRAM). (2) Kilimanjaro Mountains, Marangu Route, near Bismarck (5Mandara) Hut, 3u109550S, 37u299370E, 2700 m a.s.l., epiphyte on tree trunk in montane rain forest, 14 May 1948, leg. Olov Hedberg 1163c (PC, S). Until recently Plagiothecium membranosulum was considered to be a southern African endemic species, known to occur in the Republic of South Africa and Lesotho (O’Shea, 2006). However, its geographical range was markedly extended when it was discovered on Réunion Island in the Indian Ocean (Ellis et al., 2011a). Herein, the species is recorded for the first time in East Africa where it was found in the Kilimanjaro Mountains in Tanzania. Two collections of P. membranosulum were found to grow epiphytically in the montane rain forest at relatively low elevations 190022700 m a.s.l. The specimen collected by O. Hedberg was reported from this site as P. nitens Dixon (Potier de la Varde, 1955) and this species is presumably conspecific with P. membranosulum. 13. Plagiothecium neckeroideum Schimp. Contributors: R. Ochyra and H. Bednarek-Ochyra Africa, Democratic Republic of Congo: Ruwenzori Mountains, ca 0u309N, ca 29u509E, 390024700 m a.s.l., forming extensive mats on ground in alpine zone, July–August 1932, leg. L. Haumann 871 (BM). Plagiothecium neckeroideum is a Eurasian species which occurs in the Alps and Eastern Carpathians in Europe (Düll, 1992), and is common in China (Li & Ireland, 2008) and Japan (Iwatsuki, 1970). In addition, it occurs at alpine outposts in the tropics in Malaysia and Papua New Guinea (Enroth, 1991). The species was once reported from Ethiopia (Miehe & Miehe, 1994) as P. neckeroideum var. javense M.Fleisch. but this material correctly represents P. Journal of Bryology 2012 VOL . 34 NO . 4 285 Bryological notes standleyi (Ellis et al., 2012c). However, P. neckeroideum cannot be excluded from the moss flora of Africa and the first well documented record of the species is provided in this note. Plagiothecium neckeroideum was found at a high elevation of 390024700 m a.s.l. in the Ruwenzori Mountains of the Democratic Republic of Congo. The African plants are characterised by having julaceous stems and branches, and strongly approach P. neckeroideum var. niitakayamae (Toyama) Z.Iwats. which is known from Taiwan and China (Li & Ireland, 2008). 14. Plagiothecium novogranatense (Hampe) Mitt. Contributors: R. Ochyra, T. Pócs and H. BednarekOchyra Tanzania: Kilimanjaro Mountains, Umbwe Route, 3u089590S, 37u179420E, 2250 m a.s.l., on soil in montane rain forest, 1 March 1985, leg. T. Pócs 6984/F (EGR, KRAM). For a long time, Plagiothecium novogranatense was considered to be a neotropical endemic species occurring at high elevations in the Andes, where it extends from Colombia to Bolivia and occurs in southeastern Brazil (Buck & Ireland, 1989). It was subsequently discovered in the Ruwenzori Mountains of Uganda in Central Africa (Ochyra et al., 2002), and this discovery confirmed the existence of bryophyte disjunctions between montane areas of tropical Central and South America and tropical Central and East Africa. Only a handful of examples of this AfroAmerican distribution pattern have so far been found amongst mosses. Examples include Squamidium brasiliense (Hornsch.) Broth. (Allen & Crosby, 1986), Tristichium mirabile (Müll.Hal.) Herzog (Ochyra & Sharp, 1988), Adelothecium bogotense (Hampe) Mitt. (Ochyra et al., 1992), Rigodium toxarion (Schwägr.) A.Jaeger (Zomlefer, 1993), Calymperes venezuelanum (Mitt.) Broth. ex Pittier (Orbán, 2000), and Plagiothecium lucidum (Hook.f. & Wilson) Paris (Ochyra et al., 2000). Herein, P. novogranatense is recorded for the first time from tropical East Africa where it was found in the Kilimanjaro Mountains of Tanzania. It appears to be relatively rare on the southwestern slope of this highest African massif, growing on ground in the afromontane rain forest at an altitude similar to that in the Ruwenzori Mountains. 15. Pseudocalliergon turgescens (T.Jensen) Loeske Contributors: T. Ezer and R. Kara Turkey: Niğde-Aladağlar National Park-Narpuz Valley: 37u49933.00N, 35u8919.00E, ca 2606 m a.s.l., on rock flushed with melting snow water, 15 October 2011. leg. et det. T. Ezer & R. Kara, Aladağ-20 (Niğde University Herbarium). Plants in the genus Pseudocalliergon (Limpr.) Loeske typically grow in mineral-rich to strongly calcareous habitats and are easily recognized by the colour and gloss of the species, the structure of alar 286 Journal of Bryology 2012 VOL . 34 NO . 4 cells, the structure and colour of the axillary hairs and their ecological preferences (Hedenäs, 1992). According to the Turkish bryological literature (Uyar & Çetin, 2004; Kürschner & Erdağ, 2005; Batan & Özdemir, 2008; Kürschner & Frey, 2011), to date only two species of the genus have been recorded in Turkey, (Pseudocalliergon lycopodioides (Brid.) Hedenäs, P. trifarium F.Weber & D.Mohr) Loeske). Herein, a third species is added to the Turkish bryoflora. This study is based on specimens held in the herbarium of Niğde University, which were collected from Niğde-Aladağlar National ParkNarpuz Valley in October 2011. P. turgescens was collected from rock periodically flushed with melting snow water. It was found together with many associates such as Anoectangium handelii Schiffn., Cinclidotus riparius (Host ex Brid.) Arn. and Philonotis fontana (Hedw.) Brid. Pseudocalliergon turgescens is easily recognized by its sparsely branched and imbricate shoots, strongly concave and apiculate leaves, and stem leaves ovate or broadly ovate (Hedenäs, 1992). It is widely distributed in the mountains of Scandinavia and widely reported in Sweden and around the Baltic Sea. The species occurs in scattered areas throughout the Holarctic region (Crum & Anderson, 1981; Karczmarz, 1971). 16. Sanionia uncinata (Hedw.) Loeske Contributors: H. Bednarek-Ochyra, R. Ochyra and M. Lebouvier Île aux Cochons: 46u69S, 50u149E, without precise details, March 1954, leg. E. Aubert de la Rüe s.n. (KRAM, PC). Île aux Cochons is the third largest island (67 km2) of Îles Crozet, the most extensive archipelago in the South Indian Ocean (Kerguelen) Province in the Subantarctic. It is the westernmost island of the group of three islands (Île aux Cochons, Îlot Pingouins and Îlots des Apôtres) lying about 100 km from Île de la Possession and Île de l’East, which are the two largest islands of the archipelago and situated in its easternmost part. Île aux Cochons is well known as a breeding site for sea birds and it holds the world’s largest King Penguin colony and the largest colony of Wandering Albatrosses in the Indian Ocean. Hitherto, the only records of bryophytes from Îles Crozet originate from Île de la Possession and Île de l’East, whereas no bryophytes have been recorded from the western island group of the archipelago. In April 2012, when checking unstudied bryophyte collections from austral islands deposited in PC, a small packet was found with a moss collected in 1954 by E. Aubert de la Rüe on Île aux Cochons. It contained a large specimen of Sanionia uncinata, one of the most widespread and abundant mosses on subantarctic islands, where it grows Bryological notes in a wide range of habitats, often forming extensive pure stands on flat areas and along streams (Ochyra et al., 2008b). This is the first record of this species from the westernmost part of Îles Crozet and the first record of a moss from an area which is considered to be a ‘blank spot’ on the bryological map in the Subantarctic. 17. Scapania crassiretis Bryhn Contributor: M. V. Dulin Russian Federation: Komi Republic, Vorkuta district, southern part of the Polar Urals, Voykarsyninsky ridge, northern branch of the Hardyus mountain, the northern end of Bol’shaja Lagorta Lake, 200 m to the south-west of the source of the Levaja Lagorta River, 66u30954.90N, 63u31943.90E, ca 375 m a.s.l., at the base of the mountain slope, rocky place overgrown by Racomitrium sp., on stones in the runnels of a temporary watercourse, 13 August 2010, leg. M.V. Dulin, 905mvd (SYKO). This is the first report of Scapania crassiretis from the Komi Republic. It is an arctomontane subcircumpolar species with sporadic occurrences in the Holarctic region. S. crassiretis is known from some European countries (Austria, Czech Republic and Slovakia, Switzerland, Italy, the former Yugoslavia, Bulgaria, Poland, Netherlands, Finland, Sweden, and Norway) and from some localities in North America (Alaska and Quebec), China and Japan. It is also recorded from Greenland (Damsholt, 2002; Schumacker & Váňa, 2005; Bakalin, 2009; Ellis et al., 2011c). In Russia, it is recorded in Murmansk Province, Yamalo-Nenets Autonomous Okrug (Yamal Peninsula), Krasnoyarsk Territory (Taymyr Peninsula, Bol’shevik Island, Sayan Mountains), Yakutia Republic, Altai Territory (Altai Mountains), Chukotka Autonomous Okrug, Buryatia Republic (Khamar-Daban Chain, Stanovoye Plateau), Magadan Region (Kolymskoe Plateau), Khabarovsk Territory (Bureya River), Sakhalin Region (Sakhalin Island, Iturup Island), Primorsky Territory (Bakalin, 2009; Konstantinova et al., 2009). 18. Schistidium helveticum (Schkuhr) Deguchi Contributor: C. Sérgio Portugal: Trás-os-Montes e Alto Douro: Pr de Silva, estrada para Campo das Viboras, 29TQF1398, 650 m a.s.l., on wet slope of a crystalline limestone quarry, August 2002, leg. Gabriel Sérgio s.n. (LISU249221). This is the first report of Schistidium helveticum for the Portuguese bryoflora. It is a globally widespread species, mainly in the Mediterranean area (Blom, 1996) but is also known from SW Norway to Asia Minor and North Africa and relatively common in Spain and in Mallorca (Casas et al., 2006). This new occurrence in Portugal fits well with the known distribution of the species, and its habitat conforms to the typical ecological preferences of this Schistidium. 19. Schistidium strictum (Turner) Loeske ex Mårtensson Contributors: C. Sérgio and M. Sim-Sim Portugal, Madeira: Pico Areeiro to Pico Ruivo, on a wet slope and on volcanic rocks, 28SCB1923, 1750 m a.s.l., 5 October 1990, leg. C. Sérgio, M. Sim-Sim, & S. Fontinha 7057, (LISU248236). Schistidium strictum is a hyperoceanic species (Blom, 1996). It occurs in western North America with a disjunct distribution in Europe. In the Mediterranean area and Macaronesia it is rare and is known from a single locality in Spain and one in Madeira (Blom, 1996), where it was collected by Düll in the highest part of the island. The present collection was found in an area of volcanic origin, more or less exposed between 1700 and 1800 m, associated with Anacolia webbii (Mont.) Schimp., Andreaea heinemannii Hampe & Müll.Hal., Leucodon canariensis (Brid.) Schwägr., Antitrichia curtipendula (Hedw.) Brid., and Orthotrichum rupestre Schleich. ex. Schwägr. In the Madeira archipelago it is certainly an uncommon species, occurring in only two localities of the upper belt on the Pico Areeiro area. 20. Willia calobolax (Müll.Hal.) Lightowlers Contributors: R. Ochyra, H. Bednarek-Ochyra and M. Lebouvier Îles Crozet: Île de la Possession, Port Alfred over Baie du Marin, by road to Rivière du Camp, north of the Alfred Faure base, 46u259S, 51u509E, 80 m a.s.l.; on rock on the scarp overgrown with Blechnum penna-marina (Poir.) Kuhn by road to penguin rookery, associated with Muelleriella crassifolia (Hook.f. & Wilson) Dusén and Guembelia kidderi (James) Ochyra & Z̊˙arnowiec, 9 November 2006, leg. R. Ochyra 11/06, 29/06 (with Marc Lebouvier) (KRAM). Willia calobolax is a subantarctic species which is most frequent on the islands in the South Indian Ocean (Kerguelen) Province, including Prince Edward Islands (Ochyra, 2008), Îles Kerguelen (Lightowlers, 1985; Ochyra, 1998) and Heard Island (Ellis et al., 2011a). The present discovery completes the geographical range of the species in this biome. Like elsewhere, W. calobolax grows on Île de la Possession on exposed, dry maritime rocks in association with Muelleriella crassifolia and and Guembelia kidderi, and only sterile plants have been detected. The moss flora of Îles Crozet is still least known of all the islands of this biome and until recently only about 50 species had been recorded from this archipelago. However, one can assume that its species richness is similar to that of the neighbouring Prince Edward Islands, where about 100 species of moss are known to occur (Ochyra, 2008). This assumption is confirmed by a number of Journal of Bryology 2012 VOL . 34 NO . 4 287 Bryological notes phytogeographically remarkable additions to the moss flora of Île de la Possession, the largest island of this group, which were made in the last decade. These included Hennediella antarctica (Ångstr.) Ochyra & Matteri (Blockeel et al., 2006), H. marginata (Hook.f. & Wilson) R.H.Zander (Cano, 2008), Anomobryum julaceum (P.Gaertn., B.Mey. & Scherb.) Schimp. (Blockeel et al., 2007a), Pohlia nutans (Hedw.) Lindb. (Blockeel et al., 2007c), Bucklandiella pachydictyon (Cardot) BednarekOchyra & Ochyra (Blockeel et al., 2008), B. striatipila (Cardot) Bednarek-Ochyra & Ochyra and Hymenoloma immersa (Broth.) Ochyra (Blockeel et al., 2009c), Blindia magellanica Müll.Hal., Ditrichum conicum (Mont.) Mitt. and Drepanocladus longifolius (Mitt.) Paris (Blockeel et al., 2010), and Catagonium nitens (Brid.) Cardot (Ellis et al., 2010). 21. Zygodon catarinoi C.A.Garcia, F.Lara, Sérgio & Sim-Sim. Contributors: B. Vigalondo, F. Lara, I. Draper and R. Garilleti Turkey: Southern Anatolia, Adana: Anti-Taurus Mts., from Kozan to Feke, Göreken, Tersakan River, disturbed riparian forest with Platanus orientalis L., Alnus glutinosa ssp. antitaurica Yalt. and A. orientalis ssp. pubescens Dippel, 37u399510N, 35u519320E, 590 m a.s.l., on trunk of Platanus orientalis, 13 July 2006. leg. R. Garilleti et al. s.n. (MAUAM); Muğla, Eskiköceĝiz, eastern shore of Köyceĝiz lake, narrow patch of Liquidambar orientalis Mill. close to the lake, 36u559200N, 28u429220E, 100 m a.s.l., on trunk and base of Liquidambar orientalis, 20 July 2006, leg. R. Garilleti et al. s.n. (MAUAM); Köyceĝiz, western shore of Köyceĝiz lake, Liquidambar orientalis disturbed forest, 36u579310N, 28u419380E, 110 m a.s.l., on trunk and base of Liquidambar orientalis, 20 July 2006, leg. R. Garilleti et al. s.n. (MAUAM); Aydin: Samsun Daği, Dilek National Park, forested maquis close to the seaside, 37u419550N, 27u099590E, at sea level, on trunk of Platanus orientalis, 21 July 2006, leg. R. Garilleti et al. s.n. (MAUAM); Samsun Daği, Dilek National Park, Olukludere Kanyon, open maquis dominated by Styrax officinalis L. and Pinus brutia Ten., 37u419370N, 27u099380E, 50 m a.s.l., on trunk of Platanus orientalis, 21 July 2006, leg. R. Garilleti et al. s.n. (MAUAM). Zygodon catarinoi is well known from the western Mediterranean, being a common species in the Iberian Peninsula and in Morocco (Draper et al., 2006; Garcia et al., 2006; Sérgio et al., 2006; Medina et al., 2008; Medina et al., 2010). In the central Mediterranean region this species has been reported from Libya (Youssef et al., 2009) and in the eastern region from Greece (Calabrese & Muñoz, 2008) and Cyprus (Frahm et al., 2009; Ellis et al., 2011b). The collections here reported from southern Anatolia 288 Journal of Bryology 2012 VOL . 34 NO . 4 represent the first records of Z. catarinoi from Turkey and extend its distribution range in the Eastern Mediterranean to continental Asia. This species is apparently similar to Z. rupestris — also occurring in southern Turkey — from which it differs in the presence of fewer, but commonly longer papillae on upper leaf cells and bistratose patches in the leaf lamina. According to the literature cited above, in the western Mediterranean Zygodon catarinoi occurs in different types of Mediterranean woods, frequently cleared, and commonly dominated by Quercus ilex ssp. ballota (Desf.) Samp. or Q. suber L., but it is also found in those of Juniperus thurifera L., Q. faginea Lam. ssp. faginea, or Q. canariensis Willd. These habitat and phorophyte preferences of Z. catarinoi are similar in Libya, where it has been collected growing on Quercus spp. and Olea europaea L. Information about this moss species habitat for Eastern Mediterranean is scarce: in Greece it has been found on Abies sp., and in Cyprus it grows on Pinus brutia Ten. The results of our field survey in southern Anatolia show that Z. catarinoi seems to prefer humid habitats, growing mainly on Platanus orientalis and Liquidambar orientalis. The localities from Aydin (the most westerly ones) correspond to a north exposed humid maquis close to the sea, favoured by the orographic rainfall induced by the Samsun Daği Mountains. In drier areas it has been found in open riparian forests (Adana), or in groves or disturbed forest along of a lake shoreline (Muğla). Acknowledgements The research of M. V. Dulin had the financial support of the Russian Foundation for Basic Research (projects no. 12-04-01476) and the Program of Presidium of the Russian Academy of Sciences (no. 12-P-4-1018). The research of L. E. Kurbatova was financially supported by grants from the Russian Foundation for Basic Research (project no. 11-04-01247-a). D. A. Philippov is grateful to the Vologda Department of the Russian Geographic Society for support. M. Kırmacı and E. Agcagil are very grateful to TÜBİTAK for providing a student scholarship to the second author and to Beata Papp (Hungarian Natural History Museum) for confirming the species. J. R. Flores and G. M. Suárez had Financial support from the Myndel Botanica Fundation and PIP 2012-2014 CONICET (Consejo Nacional de Investigaciones Cientı́ficas y Técnicas). The contributions by H. Bednarek-Ochyra and R. Ochyra have been financially supported by the Polish Ministry of Science and Higher Education through grants no. N N 303 796 940 for H. Bednarek-Ochyra and no. N N 303 469 338 for R. Ochyra. They also thank the Curators Bryological notes at BM, PC and S for kindly allowing them to examine the herbarium material used in their study. The field work of R. Ochyra and Marc Leboubier on Îles Crozet and Îles Kerguelen was organised within the programme 136 ECOBIO of the French Polar Institute (IPEV). The investigations of P. Szücs and A. Bidló have been carried out with the sponsorship of TÁMOP-4.2.2.B-10/1-2010-0018. The contribution by V. Plášek, G. J. Wolski and J. Sawicki is part of a research project of the Institute of Environmental Technologies, reg. no. CZ.1.05/2.1.00/03.0100, supported by the ‘Research and Development for Innovations’ Operational Programme financed by the Structural Funds of the European Union and by the state budget of the Czech Republic. B. Vigalondo, F. Lara, I. Draper & R. 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