08_Colacino.fm Page 1 Mercredi, 6. septembre 2006 4:11 16
Cryptogamie, Bryologie, 2006, 27 (4): 1-•••
© 2006 Adac. Tous droits réservés
Bryophyte flora of Albania: A Preliminary Check-List
Carmine COLACINO a* and Marko SABOVLJEVI± b
a
Laboratorio di Briologia, Dipartimento di Biologia,
Difesa & Biotecnologie Agro-Forestali, Facoltà di Agraria,
Università della Basilicata, Viale dell’Ateneo Lucano 10, 85100 Potenza, Italy
b
Institute of Botany & Botanical Garden, Faculty of Biology,
University of Belgrade, Takovska 43, 11000 Belgrade, Serbia
(marko@bfbot.bg.ac.yu)
(Received 18 April 2005, accepted 31 January 2006)
Résumé – Un inventaire préliminaire de la flore des bryophytes d’Albanie (Sud-Est de
l’Europe) est présenté. La flore des bryophytes de l’Albanie comprend 327 taxons
représentant 141 genres et 62 familles. La liste est basée sur des travaux réalisés à partir
de 1888; elle représente le premier inventaire des bryophytes du pays. Une distinction a
été établie entre les références antérieures à 1950 et celles plus récentes. Quelques taxons
indiqués comme présents en Albanie, mais situés hors des limites territoriales actuelles du
pays, ont été exclus. Les synonymes relevés dans les travaux originels utilisés ont été
listés.
Abstract – A preliminary check-list of the bryoflora of Albania (SE Europe) within its
political boundaries is presented. It consists of 327 taxa representing 141 genera and
62 families. This list is based on literature reports from 1888, and it represents the first
check-list for the bryophytes of this country. Reports before 1950 are distinguished from
those based on more recent records. Some taxa previously reported for Albania, but from
localities actually outside Albanian present-day political borders are indicated and
excluded. Synonyms from the works considered are reported.
Albania / South-Eastern Europe / Balkans / Flora / Bryophytes / Mosses / Liverworts /
Hornworts / Check-list
INTRODUCTION
Albania represents one of the less known areas of Europe in relation to
its bryophyte flora (see, for instance Stewart, 1995; Söderström et al., 1998;
Sabovljevi† et al., 2001; Söderström et al., 2002; as well as Bego & Koni, 1999).
No check-list has ever been prepared for this group of plants, perhaps also
because of a lack of local bryologists. In his studies on European bryophytes,
* Correspondence and reprints: colacino@unibas.it
08_Colacino.fm Page 2 Mercredi, 6. septembre 2006 4:11 16
2
C. Colacino and M. Sabovljevi†
Düll (1983, 1984, 1985, 1992) lists 3 hornworts, 63 liverworts and 109 moss
species in the Albanian flora. Söderström et al. (2002) list 3 hornworts and
89 liverwort species for Albania. A much higher bryophyte diversity is likely to
be present in Albania. All bryological studies have been carried out by foreign
scientists or collectors, often as part of studies covering a wider area, especially
for the older records, in a period when Albania was still an integral part of the
Ottoman Empire, the so called Turkey in Europe. These older works covered
mainly the border area between Albania and Montenegro or Serbia (Kosova/
Kosovo).
This check-list has been prepared because one us (Senior Author) has
collected extensively during recent years in Albania, the need of a check-list
including all previous records from the literature became apparent. It is hoped this
work will fill the gap and stimulate further bryological investigations in Albania.
It should be emphasized that this is not a specimen based list, distribution is based
solely upon the literature references.
MATERIALS AND METHODS
The check-list (Table 1, which includes also the excluded taxa, and notes)
is supplemented by a systematic synopsis of genera (Table 2), and synonyms
(Table 3).
The following symbols are used:
1 = report based on collections made before 1950.
3 = report based on collections made after 1950.
For synonyms and accepted names we have referred to Corley et al.,
1981; Corley et Crundwell, 1991; Cortini Pedrotti, 2001a, 2001b, 2006; Dierßen,
2001; Düll, 1983, 1984, 1985, 1992; Grolle & Long, 2000; Podp{ra, 1954;
Schumacker & Vá≈a, 2005; Smith, 2004; Zander, 1993 (for Pottiaceae), and
consulted the on line data base TROPICOS [http://mobot.mobot.org/W3T/
Search/most.html] of the Missouri Botanical Garden, which is based on the
information contained in Index Muscorum (Wijk, Margadant & Florschuetz, 1959,
1962, 1964, 1967, 1969), Index Muscorum supplementa (Bauer & Crosby 1986;
Crosby 1977, 1979; Crosby & Bauer 1981, 1983, 1987), in the various editions of
Index of Mosses (Crosby et al., 1992; Crosby & Magill, 1994, 1997, 2000, 2004,
2005), and is continually updated.
In the maps (Figs 1, 2 & 3) are shown, respectively, the names of the
administrative division of the country (Prefectures), the number of taxa of
liverworts and hornworts, and the number of taxa of mosses, in parenthesis is
indicated the number of collections (records). The lower administrative
subdivisions (not represented) are: Rreth (District), and Komunë/Bashki
(Commune/Municipality). For most collections an attempt has been made to
determine their localization at the lower administrative level, sometime even
more precisely. These data are not presented here, though.
For each taxon we state the distribution in Albania, the number of
accepted records (collections), literature references, and excluded records
(localities outside Albanian present-day political borders).
Und.
GJI
BER
VLO
KOR
FIE
ELB
TIR
DUR
DIB
LZH
"
"
"
"
"
"
"
"
"
#
#
#
"
"
"
" "
"
"
"
"
"
" "
"
#
"
"
"
"
"
" # #
"
" "
"
" "
"
"
"
" "
" "
" "
"
"
" "
" " "
"
"
"
#
Records
excluded
h, q
l, n, o
c
m
n
q
q
q
c
c
c
c
c
c
q
c
c, l, m
c, h, l, m, n, q
c
c
c
c, h, j, k, m, n, o, q
c
l, m
c, k, n
q1: KS
c
3
"
"
c
c
c, n
N. of
Records References
2
3
2
1
3
1
1
1
1
1
27
1
3
2
1
1
5
11
8
3
13
26
1
2
4
2
Records
excluded
Bryoflora of Albania: A Preliminary Check-List
Und.
#
GJI
BER
VLO
KOR
FIE
ELB
DUR
TIR
DIB
LZH
KUK
SHK
Aneura pinguis (L.) Dumort.
Athalamia hyalina (Sommerf.) S.Hatt.
Athalamia spathysii (Lindenb.) S.Hatt.
Barbilophozia barbata (Schmidel ex Schreb.) Loeske
Barbilophozia lycopodioides (Wallr.) Loeske
Bazzania tricrenata (Wahlenb.) Lindb.
Bazzania trilobata (L.) Gray
Calypogeia azurea Stotler & Crotz
Calypogeia fissa (L.) Raddi
Cephalozia bicuspidata (L.) Dumort.
Cephaloziella baumgartneri Schiffn.
Cephaloziella divaricata (Sm.) Schiffn. var. divaricata (1)
Cephaloziella stellulifera (Taylor ex Spruce) Schiffn.
Cephaloziella turneri (Hook.) Müll.Frib.
Chiloscyphus polyanthos (L.) Corda
Cololejeunea calcarea (Lib.) Schiffn.
Cololejeunea rossettiana (C.Massal.) Schiffn.
Conocephalum conicum (L.) Dumort.
Corsinia coriandrina (Spreng.) Lindb.
Fossombronia angulosa (Dicks.) Raddi
Fossombronia caespitiformis De Not. ex Rabenh.
Frullania dilatata (L.) Dumort.
Frullania fragilifolia (Taylor) Gottsche, Lindenb. & Nees
Frullania inflata Gottsche
Frullania tamarisci (L.) Dumort.
Gongylanthus ericetorum (Raddi) Nees
N. of
Records References
1
1
3
"
Anthoceros punctatus L.
Phaeoceros bulbiculosus (Brot.) Posk.
Phaeoceros laevis (L.) Prosk.
MARCHANTIOPHYTA
1
2
3
4
5
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
24
25
26
KUK
1
2
3
SHK
ANTHOCEROTOPHYTA
08_Colacino.fm Page 3 Mercredi, 6. septembre 2006 4:11 16
Table 1. Check-List of Albanian Bryophytes.
(The distribution is given in relation to the main administrative subdivisions of the country, for their acronyms and localization refer to fig. 1)
"
#
"
#
"
"
"
"
"
"
" "
"
" "
" "
#
#
" "
"
"
#
#
"
"
"
"
"
"
#
"
"
"
#
"
"
" "
" "
"
"
"
"
"
" "
" "
"
"
"
"
" # " " " " " " " "
" "
#
"
"
"
"
"
"
" "
"
" "
#
#
"
"
#
# #
"
" "
" "
6
1
1
1
2
1
1
18
23
1
10
2
1
1
1
46
14
3
2
3
7
1
1
26
9
5
3
2
3
11
2
9
1
16
c, m, n
q
c
q
c, n
n
c
c, m, n
c, l, m, n
q
c, h, o
n, q
n
q
q
c, m, n
c, m
a, c
c
h, n
c, n, q
c
m
c, h, l, m, n, q
c, h
c, n, q
m, n
l, m
a, n
c, h, l, m, n, q
j
a, c, m, n
q
c, j, k, m, n , q
Records
excluded
q1: KS
q1: MN
q1: KS
q2: 2KS
q1: MN
q1: KS
C. Colacino and M. Sabovljevi†
"
" "
"
N. of
Records References
08_Colacino.fm Page 4 Mercredi, 6. septembre 2006 4:11 16
4
Und.
GJI
BER
"
VLO
KOR
FIE
ELB
TIR
"
DUR
DIB
LZH
Jungermannia atrovirens Dumort.
Jungermannia gracillima Sm.
Jungermannia hyalina Lyell
Jungermannia leiantha Grolle
Jungermannia polaris Lindb.
Leiocolea badensis (Gottsche) Jørg.
Leiocolea collaris (Nees) Schljakov
Leiocolea turbinata (Raddi) H.Buch.
Lejeunea cavifolia (Ehrh.) Lindb.
Lophocolea bidentata (L.) Dumort.
Lophocolea heterophylla (Schrad.) Dumort.
Lophocolea minor Nees
Lophozia excisa (Dicks.) Dumort.
Lophozia incisa (Schrad.) Dumort.
Lophozia ventricosa (Dicks.) Dumort.
Lunularia cruciata (L.) Lindb.
Mannia androgyna (L.) A. Evans
Mannia triandra (Scop.) Grolle
Marchantia paleacea Bertol. (2)
Marchantia polymorpha L. (3)
Metzgeria furcata (L.) Dumort.
Oxymitra incrassata (Brot.) Sérgio & Sim-Sim
Pedinophyllum interruptum (Nees) Kaal.
Pellia endiviifolia (Dicks.) Dumort.
Plagiochasma rupestre (J.R.Forst. & G.Forst.) Steph.
Plagiochila asplenioides (L. emend. Taylor) Dumort. (4)
Plagiochila porelloides (Torrey ex Nees) Lindenb. (4)
Porella arboris-vitae (With.) Grolle
Porella cordaeana (Huebener) Moore
Porella platyphylla (L.) Pfeiff.
Porella baueri (Schiffn.) C.E.O.Jens.
Preissia quadrata (Scop.) Nees
Ptilidium ciliare (L.) Hampe
Radula complanata (L.) Dumort.
KUK
27
28
29
30
31
32
33
34
35
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
SHK
MARCHANTIOPHYTA Tab. 1 (cont.)
" "
" "
"
N. of
Records References
5
c, m
64
1
1
#
"
"
"
"
2
2
6
1
"
" "
"
"
3
"
"
2
2
"
"
"
" "
"
1
"
" "
"
"
6
4
3
3
"
"
2
"
3
" "
" "
"
"
1
"
"
#
4
"
2
"
2
1
#
"
"
"
" "
" "
" "
33
" "
"
" "
"
14
26
Und.
GJI
BER
VLO
KOR
FIE
" #
ELB
TIR
#
" #
#
#
" # #
DUR
Aloina ambigua (Bruch & Schimp.) Limpr.
Amblystegium serpens (Hedw.) Schimp.
Anomodon attenuatus (Hedw.) Hueb.
Anomodon longifolius (Schleich. ex Brid.) Hartm.
Anomodon viticulosus (Hedw.) Hook. & Taylor
DIB
LZH
KUK
BRYOPHYTA
"
a, c, h, m, n
c
q
c
c, n
c
n
c
c
c
c
c
c
c
c
c
m
m, n
n
c
c, q
q
c
c, m
c, h
N. of
Records References
1
4
1
1
9
Bryoflora of Albania: A Preliminary Check-List
"
"
"
" "
"
" "
Records
excluded
Records
excluded
f
g, h, q
q
a
a, g, h, i, k, l, n, q
08_Colacino.fm Page 5 Mercredi, 6. septembre 2006 4:11 16
GJI
Und.
"
" "
" "
VLO
BER
KOR
"
FIE
ELB
TIR
DUR
DIB
LZH
"
"
SHK
1
2
3
4
5
Radula lindenbergiana Gottsche ex C.Hartm.
Reboulia hemisphaerica (L.) Raddi
Riccardia multifida (L.) Gray
Riccardia palmata (Hedw.) Carruth.
Riccia bicarinata Lindb.
Riccia bifurca Hoffm.
Riccia ciliata Hoffm.
Riccia ciliifera Link ex Lindenb.
Riccia crozalsii Levier
Riccia crystallina L. emend. Raddi
Riccia glauca L.
Riccia macrocarpa Levier
Riccia michelii Raddi
Riccia nigrella DC.
Riccia sorocarpa Bisch. var. sorocarpa (5)
Riccia subbifurca Warnst. ex Croz.
Riccia trabutiana Steph.
Scapania aequiloba (Schwägr.) Dumort.
Scapania aspera Bernet & M. Bernet
Scapania calcicola (Arnell & J.Perss.) Ingham
Scapania compacta (A.Roth) Dumort.
Scapania nemorea (L.) Grolle
Scapania umbrosa (Schrad.) Dumort.
Southbya nigrella (De Not.) Henriq.
Southbya tophacea (Spruce) Spruce
Targionia hypophylla L.
KUK
61
62
63
64
65
66
67
68
69
70
71
72
73
74
75
76
77
78
79
80
81
82
83
84
85
86
SHK
MARCHANTIOPHYTA Tab. 1 (cont.)
5
N. of
Records References
2
f, n
2
1
f, n
h
q
2
"
"
" "
6
"
1
"
1
#
1
#
#
" "
5
" "
1
"
#
"
"
"
2
"
5
"
"
#
1
#
#
#
"
#
2
2
"
6
"
4
"
#
"
"
"
3
#
"
1
"
#
2
n, q
#
2
a, f
"
1
#
1
1
n
a
o
a, f
q
o
5
h, k, n, q
#
"
1
"
2
#
1
#
"
"
#
"
q1: MN
q1: MN
q1: MN
C. Colacino and M. Sabovljevi†
4
a, q
f, n
f, h
n
q
g
h, q
n
a, m
f, h, n
g, h
n
q
a, f, h, q
f, h, n, q
a, f, n
f, h, n
2
#
Records
excluded
08_Colacino.fm Page 6 Mercredi, 6. septembre 2006 4:11 16
6
Und.
GJI
BER
VLO
Bryum pseudotriquetrum (Hedw.) P.Gaertn., B.Mey. & Scherb.
28 Bryum radiculosum Brid.
29 Bryum schleicheri Schwägr.
30 Bryum schleicheri var. latifolium (Schwägr.) Schimp.
31 Bryum stirtonii Schimp.
32 Bryum torquescens Bruch ex De Not.
33 Buxbaumia aphylla Hedw.
34 Buxbaumia viridis (Moug. ex Lam. & DC.) Moug. & Nestl.
35 Calliergonella cuspidata (Hedw.) Loeske
KOR
"
FIE
#
Bryum gemmiparum De Not.
Bryum kunzei Hornsch.
Bryum pallens Sw.
ELB
Bryum donianum Grev.
24
25
26
27
TIR
# #
DUR
Antitrichia curtipendula (Hedw.) Brid.
Barbula convoluta Hedw.
Barbula unguiculata Hedw.
Bartramia ithyphylla Brid.
Bartramia pomiformis Hedw.
Bartramia stricta Brid.
Brachytheciastrum velutinum (Hedw.) Ignatov & Huttunen
Brachythecium albicans (Hedw.) Schimp.
Brachythecium rivulare Schimp.
Brachythecium rutabulum (Hedw.) Schimp.
Brachythecium salebrosum (Hoffm. ex F.Web. & D.Mohr) Schimp.
Bryoerythrophyllum recurvirostre (Hedw.) P.C.Chen
Bryum algovicum Sendtn. ex Müll.Hal.
Bryum argenteum Hedw.
Bryum caespiticium Hedw.
Bryum canariense var. provinciale (H.Philib.) Husn.
Bryum capillare Hedw.
DIB
6
7
8
9
10
11
12
13
14
15
16
17
18
19
20
21
22
23
LZH
KUK
SHK
BRYOPHYTA (Tab. 1 cont.)
"
"
"
"
#
"
#
"
#
#
"
"
#
#
#
#
#
"
"
"
#
#
#
" "
"
"
"
"
#
"
"
#
" #
"
"
" "
"
#
"
"
"
"
1
2
1
2
1
2
3
1
1
1
7
3
1
4
3
1
1
1
1
1
1
1
7
5
2
3
1
1
5
3
5
3
1
h
n
g
k, q
f
k, n
a, n, q
n
n
k
a, g, n, q
n
k
i, n
a, f
n
h
f
a
q
q
q
a, k, n, q
f, h, n
h
f, h
q
q
f, h, n
f, h
n, q
a, n
h
Records
excluded
q1: KS
q1: KS
q3: MN;2KS
q1: KS
Bryoflora of Albania: A Preliminary Check-List
#
#
#
"
#
" #
"
"
#
"
#
N. of
Records References
q1: KS
q1: KS
08_Colacino.fm Page 7 Mercredi, 6. septembre 2006 4:11 16
"
"
#
Und.
GJI
BER
VLO
KOR
FIE
ELB
TIR
DUR
DIB
LZH
Campyliadelphus chrysophyllus (Brid.) R.S.Chopra
Campylium protensum (Brid.) Kindb.
Campylophyllum sommerfeltii (Myrin) Hedenäs (6)
Ceratodon purpureus (Hedw.) Brid.
Cheilothela chloropus (Brid.) Lindb. ex Broth.
Cinclidotus aquaticus (Hedw.) Bruch & Schimp.
Cinclidotus fontinaloides (Hedw.) P.Beauv.
Cinclidotus fontinaloides var. lorentzianus Mol. (7)
Cinclidotus riparius (Host ex Brid.) Arnott
Coscinodon cribrosus (Hedw.) Spruce
Cratoneuron filicinum (Hedw.) Spruce
Cratoneuron filicinum var. atrovirens (Brid.) Ochyra
Crossidium squamiferum (Viv.) Jur.
Ctenidium molluscum (Hedw.) Mitt.
Dialytrichia mucronata (Brid.) Broth.
Dichodontium pellucidum (Hedw.) Schimp.
Dicranella heteromalla (Hedw.) Schimp.
Dicranella varia (Hedw.) Schimp.
Dicranoweisia cirrata (Hedw.) Lindb. ex Milde
Dicranoweisia crispula (Hedw.) Milde
Dicranum majus Sm.
Dicranum polysetum Sw.
Dicranum scoparium Hedw.
Didymodon acutus (Brid.) K.Saito
Didymodon fallax (Hedw.) Zander
Didymodon luridus Hornsch. ex Spreng.
Didymodon rigidulus Hedw.
Didymodon sinuosus (Mitt.) Delogne
Didymodon tophaceus (Brid.) Lisa
Didymodon vinealis (Brid.) Zander
Distichium capillaceum (Hedw.) Bruch & Schimp.
Ditrichum flexicaule (Schwägr.) Hampe
Drepanocladus aduncus (Hedw.) Warnst.
KUK
36
37
38
39
40
41
42
43
44
45
46
47
48
49
50
51
52
53
54
55
56
57
58
59
60
61
62
63
64
65
66
67
68
SHK
BRYOPHYTA (Tab. 1 cont.)
7
3
1
"
"
"
"
5
6
3
4
4
5
1
1
2
3
2
1
1
1
1
1
6
n, q
a
f
h, n, q
f, h, n, q
n
h, n
a, g, h, q
h
h
n
g
h
n
g
h
n
i
q
n
1
h
2
2
5
6
h
h
a, f, h, k, q
f, h, n
a
q
h
f
f
a, f, h, k, n
f
1
#
"
#
"
"
#
"
"
"
" #
"
"
"
"
"
"
"
"
"
#
"
"
#
"
"
"
#
#
"
"
"
"
"
"
"
#
"
#
" #
" "
"
"
"
1
#
1
#
1
"
1
#
1
#
#
"
#
"
7
1
Records
excluded
q1: KS
q1: KS
C. Colacino and M. Sabovljevi†
"
#
"
N. of
Records References
1
n
q1: MN
q1: KS
08_Colacino.fm Page 8 Mercredi, 6. septembre 2006 4:11 16
8
"
#
Und.
GJI
BER
VLO
KOR
FIE
ELB
TIR
DUR
DIB
LZH
91
92
93
94
95
96
97
98
99
100
101
Encalypta alpina Sm.
Encalypta ciliata Hedw.
Encalypta rhaptocarpa Schwägr.
Encalypta rhaptocarpa var. trachymitria (Ripart) Wijk & Margad.
Encalypta streptocarpa Hedw.
Encalypta vulgaris Hedw.
Encalypta vulgaris var. obtusa Nees & Hornsch.
Eucladium verticillatum (Brid.) Schimp.
Eurhynchium crassinervium (Taylor) Schimp.
Eurhynchium hians (Hedw.) Sande Lac
Eurhynchium hians var. rigidum (Boul.) Düll
Eurhynchium meridionale (Schimp.) De Not.
Eurhynchium praelongum (Hedw.) Schimp.
Eurhynchium pulchellum (Hedw.) Jenn.
Eurhynchium pulchellum var. diversifolium (Schimp.) C.E.O.Jens.
Eurhynchium pumilum (Wils.) Schimp.
Eurhynchium schleicheri (Hedw.f.) Jur.
Eurhynchium striatulum (Spruce) Schimp.
Eurhynchium striatum (Schreb. ex Hedw.) Schimp.
Fissidens bryoides Hedw.
Fissidens dubius P.Beauv.
Fissidens dubius var. mucronatus (Breidl. ex Limpr.) Kartt., Hedenäs &
Söderstr.
Fissidens taxifolius Hedw.
Fontinalis antipyretica Hedw.
Funaria hygrometrica Hedw.
Funaria muhlenbergii Turner
Grimmia alpestris (F. Weber & D. Mohr) Schleich.
Grimmia crinita Brid.
Grimmia hartmanii Schimp.
Grimmia laevigata (Brid.) Brid.
Grimmia muehlenbeckii Schimp.
Grimmia pulvinata (Hedw.) Sm. ex Sm. & Sowerby
Grimmia tergestina Tomm. ex Bruch & Schimp.
KUK
69
70
71
72
73
74
75
76
77
78
79
80
81
82
83
84
85
86
87
88
89
90
SHK
BRYOPHYTA (Tab. 1 cont.)
a
f
n
n
n, q
n
h
i, n
"
2
n
"
1
10
1
1
n
a, h, i, n, q
h
a
2
n
3
1
9
3
2
7
1
10
n, q
a
g, h, i, n, q
k, n
h
g, h, k, l, n
g
a, g, h, i, n, q
i
n, q
n
n
o
i, n, q
i, n, q
n
q
a, f, h
"
"
"
#
"
" #
"
#
#
" "
#
"
# #
"
"
" #
"
" #
#
"
#
"
"
#
" #
#
"
"
"
"
"
#
" "
" "
1
#
"
#
5
"
1
"
"
#
" #
"
#
"
3
"
#
#
"
"
"
1
3
4
1
1
3
Records
excluded
q3:MN;2KS
q2:MN, KS
q1: KS
q1: KS
Bryoflora of Albania: A Preliminary Check-List
"
1
1
1
1
2
1
3
4
#
#
#
N. of
Records References
q1: MN
q1: MN
08_Colacino.fm Page 9 Mercredi, 6. septembre 2006 4:11 16
Und.
GJI
BER
VLO
KOR
FIE
ELB
TIR
DUR
DIB
116
117
118
119
120
121
122
123
124
125
126
127
128
129
130
131
132
133
LZH
111
112
113
114
115
Gymnostomum aeruginosum Sm.
Gymnostomum calcareum Nees & Hornsch.
Gymnostomum viridulum Brid.
Habrodon perpusillus (De Not.) Lindb.
Herzogiella seligeri (Brid.) Z. Iwats. (8)
Homalia besseri Lob.
Homalothecium aureum (Lag. ex Spruce ) H.Rob.
Homalothecium lutescens (Hedw.) H. Rob.
Homalothecium lutescens var. fallax (Philib. ex Schimp.) Hedenäs &
Söderstr.
Homalothecium philippeanum (Spruce) Bruch & Schimp.
Homalothecium sericeum (Hedw.) Schimp.
Hygroamblystegium varium (Hedw.) Mönk.
Hygrohypnum luridum (Hedw.) Jenn.
Hygrohypnum luridum var. subsphaericarpon (Schleich. ex Brid.)
C.E.O.Jens.
Hylocomium splendens (Hedw.) Schimp.
Hymenostylium recurvirostre (Hedw.) Dixon
Hypnum cupressiforme Hedw.
Isothecium alopecuroides (Lam. ex Dubois.) Isov.
Leptodictyum riparium (Hedw.) Warnst.
Leptodon smithii (Hedw.) F.Web. & D.Mohr
Leskea polycarpa Ehrh. ex Hedw.
Leucodon sciuroides (Hedw.) Schwägr.
Leucodon sciuroides var. morensis (Schwägr.) De Not.
Mnium stellare Reichard ex Hedw.
Mnium thomsonii Schimp.
Myurella julacea (Schwägr.) Schimp.
Neckera cephalonica Jur. & Unger.
Neckera complanata (Hedw.) Hueb.
Neckera crispa Hedw.
Orthothecium rufescens (Dicks. ex Brid.) Schimp.
Orthotrichum affine Schrad. ex Brid.
Orthotrichum anomalum Hedw.
KUK
102
103
104
105
106
107
108
109
110
SHK
BRYOPHYTA (Tab. 1 cont.)
9
" #
#
#
#
#
"
#
"
" "
"
"
"
"
#
"
"
#
# #
#
#
#
"
"
#
#
#
"
#
#
#
#
" "
#
#
" " "
"
#
#
"
"
"
"
"
"
"
"
#
# #
"
"
"
"
#
"
a, f
f
f
h
h, i, n, q
i, k
i
a, h, l, n
n
h
n
a, h, n
q
n, q
q
q
q
g, n, q
k, q
q
h, n, q
n, q
q
q
f, h, n
q
f
n
n
n
n
a, n, q
h, k
h
n, q
q1: KS
q1: KS
q1: KS
q1: KS
C. Colacino and M. Sabovljevi†
"
"
2
1
1
1
5
2
2
6
1
1
1
4
1
4
1
1
5
3
2
1
5
3
1
1
6
1
1
3
1
1
2
6
3
2
3
Records
excluded
08_Colacino.fm Page 10 Mercredi, 6. septembre 2006 4:11 16
"
N. of
Records References
10
#
#
#
Und.
GJI
BER
VLO
KOR
FIE
ELB
TIR
DUR
DIB
LZH
Orthotrichum cupulatum Hoffm. ex Brid.
Orthotrichum cupulatum var. sardagnanum (Vent.) Vent
Orthotrichum diaphanum Schrad. ex Brid.
Orthotrichum shawii Wilson in Schimp.
Orthotrichum speciosum Nees ex Sturm.
Orthotrichum stramineum Hornsch. ex Brid.
Orthotrichum striatum Hedw.
Palustriella commutata (Hedw.) Ochyra
Palustriella commutata var. fluctuans (Schimp.) Ochyra
Palustriella commutata var. ptychodioides (G.Roth) Ochyra
Palustriella decipiens (De Not.) Ochyra
Palustriella falcata (Brid.) Hedenäs
Paraleucobryum longifolium (Ehrh. ex Hedw.) Loeske
Philonotis calcarea (Bruch & Schimp.) Schimp. (9)
Philonotis fontana (Hedw.) Brid.
Philonotis marchica (Hedw.) Brid.
Philonotis seriata Mitt. (9)
Plagiomnium affine (Bland. ex Funck) T.J.Kop.
Plagiomnium cuspidatum (Hedw.) T.J.Kop.
Plagiomnium rostratum (Schrad.) T.J.Kop.
Plagiomnium undulatum (Hedw.) T.J.Kop.
Plagiopus oederianus (Sw.) Crum & Anders.
Plagiothecium nemorale (Mitt.) A.Jaeger
Platydictya subtilis (Hedw.) H.A.Crum
Pleurochaete squarrosa (Brid.) Lindb.
Pleurozium schreberi (Willd. ex Brid.) Mitt.
Pohlia atropurpurea (Wahlenb.) H.Lindb.
Pohlia cruda (Hedw.) Lindb.
Bryum alpinum var. latifolium Mönk.
Pohlia wahlenbergii (F.Web. & D.Mohr) Andr.
Polytrichastrum alpinum (Hedw.) G.L.Sm.
Polytrichum juniperinum Hedw.
Polytrichum piliferum Hedw.
Pseudocrossidium revolutum (Brid.) Zander
Pseudoleskea incurvata (Hedw.) Loeske
KUK
134
135
136
137
138
139
140
141
142
143
144
145
146
147
148
149
150
151
152
153
154
155
156
157
158
159
160
161
162
163
164
165
166
167
168
SHK
BRYOPHYTA (Tab. 1 cont.)
"
5
"
#
#
#
# #
"
"
" "
"
"
#
#
"
"
# #
"
"
"
"
"
"
"
"
"
#
"
"
"
#
#
" "
"
" #
"
"
"
"
" #
"
" "
"
"
"
"
"
"
"
"
"
" "
"
#
" " #
#
q1: KS
q1: MN
q1: MN
11
"
"
#
1
1
3
1
4
1
2
1
2
12
1
4
7
1
1
1
8
1
n
i
i
n
i, k, n, q
a
q
n
h, k, n
a, q
n
h
h, n
g, h
n
g, h
g, h
h
h
l, n, q
n
n, q
n
n
n
f, n
a, h, k, n, q
n
g, h
g, h
n
i
h
a, f, h, k
f
Records
excluded
Bryoflora of Albania: A Preliminary Check-List
"
2
4
1
2
9
1
1
2
5
2
1
1
3
5
1
3
"
#
N. of
Records References
08_Colacino.fm Page 11 Mercredi, 6. septembre 2006 4:11 16
Und.
GJI
BER
VLO
KOR
FIE
ELB
TIR
DUR
DIB
LZH
Pseudoleskea radicosa (Mitt.) Macoun & Kindb.
Pseudoleskea saviana (De Not.) Latzel
Pseudoleskeella catenulata (Brid. ex Schrad.) Kindb.
Pseudoscleropodium purum (Hedw.) M.Fleisch.
Pterigynandrum filiforme Hedw.
Pterogonium gracile (Hedw.) Sm.
Ptilium crista-castrensis (Hedw.) De Not.
Ptychodium plicatum (Schleich. ex F.Web. & D.Mohr) Schimp.
Racomitrium canescens (Hedw. ex Hedw.) Brid.
Racomitrium ericoides (Web. ex Brid.) Web.
Rhizomnium punctatum (Hedw.) T. Kop.
Rhynchostegiella curviseta (Brid.) Limpr.
Rhynchostegiella litorea (De Not.) Limpr.
Rhynchostegiella tenella (Dicks.) Limpr.
Rhynchostegium alopecuroides (Brid.) A.J.E.Smith
Rhynchostegium confertum (Dicks.) Schimp.
Rhynchostegium megapolitanum (Blandow ex F.Web. & D.Mohr) Schimp.
Rhynchostegium megapolitanum var. meridionale Schimp.
Rhynchostegium murale (Hedw.) Schimp.
Rhynchostegium riparioides (Hedw.) Cardot
Rhynchostegium riparioides fo. atlanticum (Brid.) Düll
Rhytidiadelphus triquetrus (Hedw.) Warnst.
Rhytidium rugosum (Hedw.) Kindb.
Saelania glaucescens (Hedw.) Broth. ex Bom. & Broth.
Sanionia uncinata (Hedw.) Loeske
Schistidium agassizii Sull. & Lesq. ex Sull.
Schistidium apocarpum (Hedw.) Bruch & Schimp.
Sciuro-hypnum glaciale var. gelidum (Bryhn) Ochyra & -arnowiec
Scleropodium touretii (Brid.) L.F.Koch
Scorpiurium circinatum (Brid.) M.Fleisch. & Loeske
Scorpiurium deflexifolium (Solms.) M.Fleisch. & Loeske
Sphagnum palustre L.
Syntrichia calcicola J.J. Amann
Syntrichia intermedia Brid.
Syntrichia intermedia var. calva (Dur. & Sag.) Delogne
KUK
169
170
171
172
173
174
175
176
177
178
179
180
181
182
183
184
185
186
187
188
189
190
191
192
193
194
195
196
197
198
199
200
201
202
203
SHK
BRYOPHYTA (Tab. 1 cont.)
"
"
"
#
"
"
"
#
"
"
"
"
#
"
#
"
"
"
" "
"
"
#
"
"
"
#
"
"
#
"
"
#
"
"
"
"
" "
"
"
1
"
#
#
#
"
"
"
# " "
1
1
2
5
3
n
n
a, f, k, n
h, n
n
n
n, q
n
i
n
h
h
n
h
h, i, l, n, q
f
h
f
n
h
f
n
a, f, h
f
n
f
f, h, n
f, h, n
a, f, h
h
k
k
q
f, h
h, q
q1: MN
q1: KS
q1: KS
q1: MN
q1: MN
C. Colacino and M. Sabovljevi†
"
"
2
1
7
2
1
1
2
2
1
1
4
2
1
3
9
1
1
1
2
2
1
1
7
1
1
1
4
4
7
Records
excluded
08_Colacino.fm Page 12 Mercredi, 6. septembre 2006 4:11 16
#
N. of
Records References
12
"
"
"
Und.
GJI
BER
"
VLO
"
KOR
FIE
ELB
TIR
DUR
DIB
LZH
Syntrichia norvegica Web.
Syntrichia ruraliformis (Besch.) Cardot
Syntrichia ruralis (Hedw.) F. Web. & D. Mohr
Thamnobryum alopecurum (Hedw.) Nieuwl. ex Gangulee
Thuidium delicatulum (Hedw.) Schimp.
Thuidium philibertii Limpr.
Thuidium recognitum (Hedw.) Lindb.
Timmia austriaca Hedw.
Timmia bavarica Hessl.
Timmiella barbuloides (Brid.) Mönkem.
Tortella flavovirens (Bruch) Broth.
Tortella flavovirens var. viridiflava (De Not.) Cas.-Gil.
Tortella inclinata (Hedw. f.) Limpr.
Tortella nitida (Lindb.) Broth
Tortella tortuosa (Hedw.) Limpr.
Tortella tortuosa var. fragilifolia (Jur.) Limpr.
Tortula canescens Mont.
Tortula cuneifolia (Dicks. ex With.) Turn. (10)
Tortula eucalyptrata Lindb.
Tortula inermis (Brid.) Mont.
Tortula lanceola var. angustata (Bruch & Schimp.) Zander
Tortula marginata (Bruch & Schimp.) Spruce
Tortula muralis Hedw.
Tortula muralis var. aestiva Brid. ex Hedw.
Tortula obtusifolia (Schwägr.) Mathieu
Tortula solmsii (Schimp.) Limpr.
Tortula subulata Hedw.
Trichostomum brachydontium Bruch
Trichostomum crispulum Bruch
Trichostomum crispulum var. brevifolium (Müll. Hall.) Bruch & Schimp.
Ulota crispa (Hedw.) Brid.
Ulota curvifolia (Wahlenb.) Lilj.
Warnstorfia fluitans (Hedw.) Loeske
Weissia condensa (Voit ex Sturm) Lindb.
Weissia controversa Hedw.
KUK
204
205
206
207
208
209
210
211
212
213
214
215
216
217
218
219
220
221
222
223
224
225
226
227
228
229
230
231
232
233
234
235
236
237
238
SHK
BRYOPHYTA (Tab. 1 cont.)
Bryoflora of Albania: A Preliminary Check-List
13
Locality is in
Reference
1 Bryophyta
KOSOVA
Atrichum undulatum var. minus (Hedw.) Paris
Locality
is in
Reference q
Locality
is
in
Reference
Bryophyta
EXCLUDED
TAXA
KOSOVA
q
Barbulaundulatum
unguiculata
obtusifolia
Mönk.
1 2 Atrichum
KOSOVA
q
var.f.minus
(Hedw.)
Paris
1 Atrichum undulatum var. minus (Hedw.) Paris
KOSOVA
q
Locality
Bryophyta
(11)is in
d
Campylophyllum
(Crundw.
2 3 Barbula
KOSOVA
qReference
unguiculatacalcareum
f. obtusifolia
Mönk.& Nyholm) Hedenäs (6)
2 Barbula unguiculata f. obtusifolia Mönk.
KOSOVA
qq
KOSOVA
Atrichum
undulatum
var. minus
(Hedw.)
Paris Hedenäs (6)
q
KOSOVA
Desmatodon
heimii
(Hedw.)
Mitt.
314 Campylophyllum
(11)
d
calcareum
(Crundw.
& Nyholm)
3 Campylophyllum calcareum (Crundw. & Nyholm) Hedenäs (6)
(11)
dq
KOSOVA
Barbula
unguiculata
f.Jur.
obtusifolia
Mönk.
(11)
d
425 Desmatodon
KOSOVA
q
Didymodon
cordatus
(12)
heimii
(Hedw.)
Mitt.
4 Desmatodon heimii (Hedw.) Mitt.
qd
KOSOVA
Campylophyllum
calcareum
(Crundw.
& Nyholm)
Hedenäs
(6) (13)
(11)
d
536 Didymodon
(11) (11)
d
Didymodon
vinealis
var.
flaccida
(Bruch
& Schimp.)
Zander
cordatus
Jur.
(12)
5 Didymodon cordatus Jur. (12)
(11)
qdq
KOSOVA
Desmatodon
heimii var.
(Hedw.)
Mitt.(Bruch & Schimp.) Zander (13)
647 Didymodon
(11)
d
KOSOVA
flaccida
Fissidens vinealis
adianthoides
Hedw.
6 Didymodon vinealis var. flaccida (Bruch & Schimp.) Zander (13)
(11)
dd
(11)
758 Fissidens
KOSOVA
q
Didymodon
cordatus var.
Jur.
(12)
adianthoides
Hedw.
MONTENEGRO
b,
d
Grimmia
hartmanii
montenegrina
Beck & Szysz. (14)
7 Fissidens adianthoides Hedw.
KOSOVA
qd
(11)
869 Grimmia
MONTENEGRO
b,
d
Didymodon
vinealis
var.
flaccida
(Bruch
&
Schimp.)
Zander
(13)
hartmanii
var.
montenegrina
Beck
&
Szysz.
(14)
(11)
e
Hypnum cupressiforme var. lacunosum Brid. (13)
MONTENEGRO
Grimmia
hartmanii
var.Hedw.
montenegrina
Beck(13)
& Szysz. (14)
98
(11)
e b,qqd
710Hypnum
KOSOVA
cupressiforme
lacunosum
Brid.
Fissidens
adianthoides
KOSOVA
Hypnum
jutlandicum var.
Holmen
& Warncke
(11)
Hypnum
cupressiforme
var. lacunosum
Brid.&
(13)
109
KOSOVA
q b,eqd
jutlandicum
& Warncke
811Hypnum
MONTENEGRO
Grimmia
hartmanii
var.
Beck
Szysz. (14)
MONTENEGRO
Pogonatum
aloides Holmen
var.montenegrina
x minimum
(Crome)
Molendo
KOSOVA
qe
10
Hypnum
jutlandicum
& Warncke
11
MONTENEGRO
q
aloides
var.Holmen
x var.
minimum
(Crome)
Molendo
912Pogonatum
(11)
Hypnum
cupressiforme
lacunosum
Brid.
(13)
KOSOVA
q
Thuidium
abietinum
(Hedw.)
Schimp.
MONTENEGRO
qq
11
Pogonatum
aloides (Hedw.)
var.
x minimum
(Crome) Molendo
12
KOSOVA
q
abietinum
Schimp.
KOSOVA
10
Hypnum
jutlandicum
Holmen
& Warncke
13Thuidium
KOSOVA
q
Thuidium
tamariscinum
(Hedw.)
Schimp.
KOSOVA
qq
12
Thuidium
abietinum
(Hedw.)
Schimp.
13
KOSOVA
q
(Hedw.)
Schimp.
MONTENEGRO
11
Pogonatum
aloides subsp.
var.
x montenegrina
minimum
(Crome)
Molendo
14Thuidium
MONTENEGRO
b,
p
Tortula tamariscinum
lingulata
(Breidl.
& Szysz.) Podp.
13
KOSOVA
Thuidium
tamariscinum
(Hedw.)
Schimp.
14
MONTENEGRO
b, p q
lingulata
subsp.
montenegrina
(Breidl. & Szysz.) Podp.
KOSOVA
qq
12
Thuidium
abietinum
(Hedw.)
Schimp.
15Tortula
KOSOVA
Tortula
mucronifolia
Schwägr.
14
MONTENEGRO
b,
p
Tortula
lingulata
subsp.
montenegrina
(Breidl.
&
Szysz.)
Podp.
15
KOSOVA
q f q(15)
mucronifolia
Schwägr.
13
KOSOVA
Thuidium
tamariscinum
(Hedw.)
16Tortula
MONTENEGRO
Weissia
controversa
var.
crispataSchimp.
(Nees & Hornsch.) Nyh.
15
KOSOVA
Tortula
mucronifolia
Schwägr.
16
MONTENEGRO
f (15) b,qp
var.
crispata
(Nees &(Breidl.
Hornsch.)
Nyh. Podp.
14 Weissia
MONTENEGRO
Tortulacontroversa
lingulata subsp.
montenegrina
& Szysz.)
16
MONTENEGRO
f (15)
controversa var.
crispata (Nees & Hornsch.) Nyh.
15 Weissia
KOSOVA
q
Tortula
mucronifolia
Schwägr.
Marchantiophyta
Marchantiophyta
MONTENEGRO
f (15)
161 Weissia
controversa
var. (Thed.
crispataex (Nees
& H.Buch.
Hornsch.) Nyh.
MONTENEGRO
q
Leiocolea
heterocolpos
Hartm.)
Marchantiophyta
1 2 Leiocolea
q (16)
MONTENEGRO
(Thed.
ex Hartm.) H.Buch.
(11)
Porella heterocolpos
obtusata (Taylor)
Trevis
MONTENEGRO
Leiocolea
heterocolpos
21 Porella
(16) q
(11)
obtusata
(Taylor) (Thed.
Trevis ex Hartm.) H.Buch.
3 Marchantiophyta
(17)
(11)
Riccia lamellosa Raddi
32
(11) (11)
(17) (16)
Porella
obtusata
(Taylor)
Trevis
lamellosa
Raddi
1 Riccia
MONTENEGRO
q
Leiocolea
heterocolpos
(Thed.
ex Hartm.) H.Buch.
32 Riccia lamellosa Raddi
(17)
(11)
(11)
(16)
Porella in
obtusata
(Taylor)
References
the check-list
areTrevis
indicated as follows:
References in the check-list are indicated as follows:
(11) (1893); g. Höhnel (1894);
(17) h. Kárpáti &
Riccia lamellosa
a.3 Baumgartner
(1915);Raddi
b. Beck & Szyszy,owicz (1888); c. Bischler & al. (1980); d. Düll (1984); e. Düll (1985); f. Höhnel
in the
check-list
as follows:
a.References
Baumgartner
(1915);
b. Beckare
& indicated
Szyszy,owicz
(1888); c. Bischler & al. (1980); d. Düll (1984); e. Düll (1985); f. Höhnel (1893); g. Höhnel (1894); h. Kárpáti &
Vajda (1961); i. Markgraf (1927a); j. Markgraf (1927b); k. Markgraf (1931); l. Meyer & Grolle (1963); m. Meyer & Grolle (1968); n. Petrov (1960); o. Petrov
Vajda
(1961);ini.the
Markgraf
j.Szyszy,owicz
Markgraf
(1927b);
(1931);
l. Meyer
& Grolle
(1963);
m.(1985);
Meyer &
Grolle (1968);
a. Baumgartner
(1915);
b.(1927a);
Beck &
(1888);k.c.Markgraf
Bischler &
al. (1980);
d. Düll
(1984);
e. Düll
f. Höhnel
(1893); n.
g. Petrov
Höhnel(1960);
(1894);o.h.Petrov
Kárpáti &
References
check-list
indicated
as follows:
(1962); p. Podp.ra
(1954); q.are
Szepesfalvy
(1926).
(1962);
p. Podp.ra
(1954); q.(1927a);
Szepesfalvy
(1926). (1927b); k. Markgraf (1931); l. Meyer & Grolle (1963); m. Meyer & Grolle (1968); n. Petrov (1960); o. Petrov
Vajda
(1961);
i.(1915);
Markgraf
j. Markgraf
a. Baumgartner
b. Beck & Szyszy,owicz
(1888); c. Bischler & al. (1980); d. Düll (1984); e. Düll (1985); f. Höhnel (1893); g. Höhnel (1894); h. Kárpáti &
(1962);(1961);
p. Podp.ra
(1954); q.
Szepesfalvy
(1926).(1927b); k. Markgraf (1931); l. Meyer & Grolle (1963); m. Meyer & Grolle (1968); n. Petrov (1960); o. Petrov
Vajda
i. Markgraf
(1927a);
j. Markgraf
Records
excluded:
References
as above,
the
number following the reference indicates the number of records excluded, the acronym(s) which follow(s) refer to
Records
excluded:
References
as above, the
number following the reference indicates the number of records excluded, the acronym(s) which follow(s) refer to
(1962);
p.
Podp.ra
(1954);
q.
Szepesfalvy
(1926).
Montenegro
(MN),
and
Kosova
(KS).
instance,
MN;2KS” means
3 records
excluded
from
reference
q ( Szepesfalvy
1926) because
localities
are actually
one
Montenegro
(MN),
andReferences
Kosova
(KS).
ForFor
instance,
“q3:“q3:
MN;2KS”
3 records
excluded
from
reference
q ( Szepesfalvy
1926)
localities
arefollow(s)
actually
one
Records
excluded:
as
above,
the
number
followingmeans
the reference
indicates
the
number
of records
excluded,
the because
acronym(s)
which
refer to
in
Montenegro,
and
the
other
2 Kosova.
in Kosova.
inMontenegro
Montenegro,
and
the
other
2
in
(MN), and Kosova (KS). For instance, “q3: MN;2KS” means 3 records excluded from reference q ( Szepesfalvy 1926) because localities are actually one
Records excluded: References as above, the number following the reference indicates the number of records excluded, the acronym(s) which follow(s) refer to
in Montenegro, and the other 2 in Kosova.
Montenegro (MN), and Kosova (KS). For instance, “q3: MN;2KS” means 3 records excluded from reference q ( Szepesfalvy 1926) because localities are actually one
in Montenegro, and the other 2 in Kosova.
08_Colacino.fm Page 13 Mercredi, 6. septembre 2006 4:11 16
EXCLUDED TAXA
Bryophyta TAXA
EXCLUDED
EXCLUDED TAXA
(1)
(2)
(3)
(4)
The variety is indicated in a more recent work (Bischler 2004) and not in the original work cited (Bischler & al. 1980).
Not indicated for Albania in a more recent work, but Albania indicated on distribution map therein (Bischler 2004)
According to Söderström & al. (2002) it should be referred to Marchantia paleacea ssp. ruderalis Bisch. & Boisselier.
In the original work (Bischler & al. 1980) Plagiochila asplenioides (with the indication "incl. var.") was indicated as the species collected, in a more recent
work Plagiochila porelloides instead is indicated as the one collected, while P. asplenoides is reported as present in Albania but not collected (Bischler 2004).
(15) The locality indicated (West coast of Lake Scutari) was within the borders of Albania from 1885 to 1891, presently is in Montenegro.
(16) Reported by M. Shehu in an unpublished thesis (Söderström, pers. comm. ); probably from the Ersekë area (Korçë), no voucher, thesis at Tirana University
(A.Miho, pers. comm. )
(17) Reported by Jovet-Ast 1986 for Albania (with a question mark); Not confirmed by Bischler 2004. Reported as doubtful for Albania by Söderström & al. (2002)
based on Jovet-Ast's paper.
C. Colacino and M. Sabovljevi†
(5) Bischler does not distinguish between varieties of this species in the original work (Bischler & al. 1980), in a more recent one, though, she does differentiate a
var. sorocarpa (which she indicates she did collect) and a var. hegii Schiffn., which she only reports as recorded from Albania (Bischler 2004).
(6) Hypnum sommerfeltii Myr., the species reported by Höhnel, is the basionym of Campylophyllum sommerfeltii (Myrin) Hedenäs, a subarctic species.
According to Smith (2004), Cortini (2006), Düll (1995), though, European authors using that combination may have been rather referring to Campylophyllum
calcareum (Crundw. & Nyholm) Hedenäs. There's a report of C. calcareum for Albania by Düll (1985), but with no indication of locality.
(7) Variety with doubtful taxonomic value (Düll 1992).
(8) In Cortini Pedrotti (2001a): Plagiothecium silesiacum (P.Beauv.) Bryol. eur. = Haplohymenium triste (Ces.) Kindb. (Probably a mistake).
(9) In TROPICOS Philonotis calcarea and Ph. seriata are considered as synonyms of Ph. fontana.
(10) Höhnel (1893) refers also to a var. luteomarginata Höhn. which he mentions he had collected also in Corsica in 1889.
(11) No voucher or reference to locality is available.
(12) Possibly in Albania (Düll, pers. comm.)
(13) Likely to be present in Albania (Düll, pers. comm.)
(14) Düll (1984) and Podp.ra (1954) report it for Albania based on a collection made by G,owacki in 1886 (in Beck and Szyszy,owicz, 1888): the locality is actually
in Montenegro.
08_Colacino.fm Page 14 Mercredi, 6. septembre 2006 4:11 16
14
Notes to table 1:
08_Colacino.fm Page 15 Mercredi, 6. septembre 2006 4:11 16
Bryoflora of Albania: A Preliminary Check-List
15
Table 2. Systematic synopsis of Albanian bryophytes.
Arrangement follows Crandall-Stotler & Stotler (2000) for liverworts (with the addition of Leiocolea within
the Jungermanniaceae), Grolle & Long (2000) for hornworts; Buck and Goffinet (2000) for mosses (with the
addition of Brachytheciastrum and Sciuro-hypnum within the Brachytheciaceae). Bryophytes (s.l.) are
considered as three distinct lineages (Divisions). The number following the genus indicates number of taxa.
Order of genera in families is alphabetical, except for Pottiaceae (*) arranged according to Zander (1993).
HORNWORTS. ANTOCEROTOPHYTA.
ANTHOCEROTOPSIDA
ANTHOCEROTALES
Anthocerotaceae
Anthoceros 1, Phaeoceros 2
LIVERWORTS. MARCHANTIOPHYTA.
MARCHANTIOPSIDA
MARCHANTIALES
Aytoniaceae
Mannia 2, Plagiochasma 1, Reboulia 1
Conocephalaceae
Conocephalum 1
Lunulariaceae
Lunularia 1
Marchantiaceae
Marchantia 2, Preissia 1
Cleveaceae
Athalamia 2
Corsiniaceae
Corsinia 1
Targioniaceae
Targionia 1
Oxymitraceae
Oxymitra 1
Ricciaceae
Riccia 13
JUNGERMANNIOPSIDA
FOSSOMBRIONALES
Fossombroniaceae
Fossombronia 2
Pelliaceae
Pellia 1
METZGERIALES
Aneuraceae
Aneura 1, Riccardia 2
Mezgeriaceae
Metzgeria 1
LEPICOLEALES
Ptilidiaceae
Ptilidium 1
JUNGERMANNIALES
Geocalycaceae
Chiloscyphus 1, Lophocolea 3
Plagiochilaceae
Pedinophyllum 1, Plagiochila 2
Arnelliaceae
Southbya 2, Gongylanthus 1
Calypogeiaceae
Calypogeia 2
Lepidoziaceae
Bazzania 2
Cephaloziaceae
Cephalozia 1
Cephaloziellaceae
Cephaloziella 4
Jungermanniaceae
Barbilophozia 2, Jungermannia 5, Leiocolea 3,
Lophozia 3
Scapaniaceae
Scapania 6
PORELLALES
Porellaceae
Porella 4
Jubulaceae
Frullania 4
Lejeuneaceae
Cololejeunea 2, Lejeunea 1
RADULALES
Radulaceae
Radula 2
MOSSES. BRYOPHYTA
SPHAGNOPSIDA
SPHAGNALES
Sphagnaceae
Sphagnum 1
POLYTRICHOPSIDA
TETRAPHIDALES
Buxbaumiaceae
Buxbaumia 2
Polytrichales
Polytrichaceae
Polytrichastrum 1, Polytrichum 2
BRYALES
Bartramiaceae
Bartramia 4, Philonotis 4, Plagiopus 1
Bryaceae
Bryum 16 (incl. 3 var.)
Mniaceae
Mnium 2, Plagiomnium 4, Pohlia 3,
Rhizomnium 1
HYPNALES
Amblystegiaceae
Amblystegium 1, Hygroamblystegium 1,
Leptodictyum 1
08_Colacino.fm Page 16 Mercredi, 6. septembre 2006 4:11 16
16
C. Colacino and M. Sabovljevi†
Table 2. Systematic synopsis of Albanian bryophytes.
Arrangement follows Crandall-Stotler & Stotler (2000) for liverworts (with the addition of Leiocolea within
the Jungermanniaceae), Grolle & Long (2000) for hornworts; Buck and Goffinet (2000) for mosses (with the
addition of Brachytheciastrum and Sciuro-hypnum within the Brachytheciaceae). Bryophytes (s.l.) are
considered as three distinct lineages (Divisions). The number following the genus indicates number of taxa.
Order of genera in families is alphabetical, except for Pottiaceae (*) arranged according to Zander (1993).
BRYOPSIDA
TIMMIALES
Timmiaceae
Timmia 2
ENCALYPTALES
Encalyptaceae
Encalypta 7 (incl. 2 var.)
Funariaceae
Funaria 2
GRIMMIALES
Grimmiaceae
Coscinodon 1, Grimmia 7, Racomitrium 2,
Schistidium 2
DICRANALES
Fissidentaceae
Fissidens 4 (incl. 1 var.)
Dicranaceae
Dicranella 2, Dicranum 3, Paraleucobryum 1
Ditrichaceae
Ceratodon 1, Cheilothela 1, Distichium 1,
Ditrichum 1, Saelania 1
Rhabdoweisiaceae
Dichodontium 1, Dicranoweisia 2
POTTIALES
Pottiaceae (*)
Timmiella 1, Eucladium 1, Trichostomum 3
(incl. 1 var.), Pleurochaete 1, Tortella 6 (incl. 2
var.), Dialytrichia 1, Bryoerythrophyllum 1,
Pseudocrossidium 1, Hymenostylium 1,
Barbula 2, Gymnostomum 3, Didymodon 7,
Weissia 2, Crossidium 1, Aloina 1, Tortula 11
(incl. 2 var.). Syntrichia 6 (incl. 1 var.)
Cinclidotaceae
Cinclidotus 4 (incl. 1 var.)
ORTHOTRICHALES
Orthotrichaceae
Orthotrichum 9 (incl. 1 var.), Ulota 2
Cratoneuraceae
Cratoneuron 2 (incl. 1 var.)
Helodiaceae
Palustriella 5 (incl. 2 var.)
Hylocomiaceae
Hylocomium 1, Pleurozium 1,
Rhytidiadelphus 1
Rhytidiaceae
Rhytidium 1
Leskeaceae
Leskea 1, Pseudoleskea 3, Pseudoleskeella 1,
Ptychodium 1
Pterigynandraceae
Habrodon 1, Myurella 1, Pteryginandrum 1
Thuidiaceae
Thuidium 3
Campyliaceae
Campyliadelphus 1, Campylium 1,
Drepanocladus 1, Hygrohypnum 2 (incl. 1
var.), Sanionia 1, Warnstorfia 1
Brachytheciaceae
Brachytheciastrum 1, Brachythecium 4,
Eurhynchium 11 (incl. 2 var.),
Homalothecium 5 (incl.1 var.), Isothecium 1,
Pseudoscleropodium 1, Rhynchostegiella 3,
Rhynchostegium 7 (incl. 1 var. & 1 form),
Sciuro-hypnum 1, Scleropodium 1,
Scorpiurium 2
Plagiotheciaceae
Plagiothecium 1
Fontinalaceae
Fontinalis 1
Hypnaceae
Campylophyllum 1, Calliergonella 1,
Ctenidium 1, Herzogiella 1, Hypnum 1,
Orthothecium 1, Platydictya 1, Ptilium 1
Leucodontaceae
Antitrichia 1, Leucodon 2 (incl. 1 var.)
Neckeraceae
Homalia 1, Neckera 3, Thamnobryum 1
Leptodontaceae
Leptodon 1
Anomodontaceae
Anomodon 3
08_Colacino.fm Page 17 Mercredi, 6. septembre 2006 4:11 16
Bryoflora of Albania: A Preliminary Check-List
17
Table 3. List of synonyms.
The citations in the original works considered have been corrected when wrong, as far as possible. When more
than two names are present the last name is the one currently used, the others are synonyms or variants.
Synonyms for “excluded taxa” (i.e., those indicated at the end of Table 1) are also included. Some recent
synonyms are also included.
Liverworts
Aneura palmata (Hedw.) Dumort. = Riccardia palmata (Hedw.) Carruth.
Calypogeia trichomanis (L.) Corda = Calypogeia azurea Stotler & Crotz
Chiloscyphus coadunatus (Sw.) J.J.Engel & R.M.Schust. = Lophocolea bidentata (L.) Dumort.
Chiloscyphus minor (Nees) J.J.Engel & R.M.Schust. = Lophocolea minor Nees
Chiloscyphus profundus (Nees) J.J.Engel & R.M.Schust. = Lophocolea heterophylla (Schrad.) Dumort.
Chomiocarpon quadratum (Scop.) Lindb. = Preissia quadrata (Scop.) Nees
Clevea hyalina (Sommerf.) Lindb. = Athalamya hyalina (Sommerf.) S.Hatt.
Dolichotheca silesiaca (F.Web. & D.Mohr) M.Fleisch.= Herzogiella seligeri (Brid.) Z. Iwats.
Fegatella conica (L.) Corda = Conocephalum conicum (L.) Dumort.
Frullania illyrica Grolle = Frullania inflata Gottsche
Haplozia crenulata (Mitt.) Müll. Frib. = Aplozia crenulata (Mitt.) Lindb. = Jungermannia gracillima Sm.
Haplozia lanceolata (Nees) Müll. Frib. = Aplozia lanceolata (L.) Dumort. = Jungermannia leiantha Grolle
Jungermannia tristis Nees = Jungermannia atrovirens Dumort.
Lophozia badensis (Gottsche) Schiffn. = Leiocolea badensis (Gottsche) Jørg.
Lophozia collaris (Nees) Dumort. = Leiocolea collaris (Nees) Schljakov
Lophozia heterocolpos (Thed. ex Hartm.) M.Howe = Leiocolea heterocolpos (Thed. ex Hartm.) H.Buch
Lophozia turbinata (Raddi) Steph. = Leiocolea turbinata (Raddi) H.Buch
Madotheca baueri Schiffn. = Porella baueri (Schiffn.) C.E.O.Jensen
Madotheca cordeana (Hueb.) Dumort. = Porella cordaeana (Hueb.) Moore
Madotheca platyphylla (L.) Dumort. = Porella platyphylla (L.) Pfeiff.
Madotheca rivularis Nees = Porella cordaeana (Hueb.) Moore
Neesiella rupestris (Nees) Schiffn. = Mannia triandra (Scop.) Grolle
Oxymitra paleacea Bisch. ex Lindenb. = Oxymitra incrassata (Brot.) Sérgio & Sim-Sim;
Pellia fabbroniana Raddi = Pellia endiviifolia (Dicks.) Dumort.
Plagiochila asplenioides var. humilis Lindenb. = Plagiochila porelloides (Torrey ex Nees) Lindenb.
Plagiochila asplenioides var. minor Lindenb. = Plagiochila porelloides (Torrey ex Nees) Lindenb.
Pleuroschisma tricrenata (Wahlenb.) Dumort. = Bazzania tricrenata (Wahlenb.) Lindb.
Pleuroschisma trilobatum (L.) Dumort. = Bazzania trilobata (L.) Gray
Porella laevigata (Schrad.) Pfeiff. = Porella arboris-vitae (With.) Grolle
Preissia commutata Nees = Preissia quadrata (Scop.) Nees
Radula lindbergiana Gottsche ex J.B.Jack = Radula lindenbergiana Gottsche ex C. Hartm.
Riccardia pinguis (L.) Gray = Aneura pinguis (L.) Dumort.
Riccia bischoffii Hueb. = Riccia ciliifera Link ex Lindenb.
Scapania nemorosa Dumort. = Scapania nemorea (L.) Grolle
Solenostoma schiffneri (Loitl.) Müll. Frib. = Jungermannia polaris Lindb.
Solenostoma triste (Nees) Müll. Frib. = Jungermannia atrovirens Dumort.
Targionia hypophylla v. fimbriata Müll. Frib. = Targionia hypophylla L.
Mosses
Acrocladium cuspidatum (Hedw.) Lindb. = Calliergonella cuspidata (Hedw.) Loeske
Amblystegium filicinum (Hedw.) De Not. = Cratoneuron filicinum (Hedw.) Spruce
Amblystegium riparium (Hedw.) Schimp. = Leptodictyum riparium (Hedw.) Warnst.
Amblystegium subtile (Hedw.) Schimp. = Platydictya subtilis (Hedw.) Crum
Amblystegium varium (Hedw.) Linbd. = Hygroamblystegium varium (Hedw.) Mönk.
Barbula acuta (Brid.) Brid. = Didymodon acutus (Brid.) K. Saito
Barbula fallax Hedw. = Didymodon fallax (Hedw.) Zander
Barbula gracilis Schwägr. = Didymodon acutus (Brid.) K. Saito
Barbula lurida (Hornsch.) Lindb. (≠ Barbula lurida Hornsch.) = Didymodon luridus Hornsch. ex Spreng.
Barbula revoluta Brid. = Pseudocrossidium revolutum (Brid.) Zander
Barbula sinuosa (Wils.) Braithw. = Barbula sinuosa (Mitt.) Grav. = Didymodon sinuosus (Mitt.) Delogne
08_Colacino.fm Page 18 Mercredi, 6. septembre 2006 4:11 16
18
C. Colacino and M. Sabovljevi†
Table 3. List of synonyms.
The citations in the original works considered have been corrected when wrong, as far as possible. When more
than two names are present the last name is the one currently used, the others are synonyms or variants.
Synonyms for “excluded taxa” (i.e., those indicated at the end of Table 1) are also included. Some recent
synonyms are also included.
Barbula tophacea (Brid.) Mitt. = Didymodon tophaceus (Brid.) Lisa
Barbula unguiculata var. obtusifolia Bruch & Schimp. =Barbula obtusifolia Schultz = Barbula unguiculata f.
obtusifolia Mönk. [Note: Barbula obtusifolia Schultz ≠ Barbula obtusifolia Schwägr. = Tortula obtusifolia
(Schwägr.) Mathieu]
Barbula vinealis Brid. = Didymodon vinealis (Brid.) Zander
Bartramia oederi Brid. = Plagiopus oederianus (Sw.) Crum & Anders.
Brachythecium glaciale var. gelidum (Bryhn) Mönk. = Sciuro-hypnum glaciale var. gelidum (Bryhn) Ochyra &
_arnowiec
Brachythecium velutinum (Hedw.) Schimp. = Brachytheciastrum velutinum (Hedw.) Ignatov & Huttunen
Bryum atropurpureum (Wahlenb.) Wahlenb. = Pohlia atropurpurea (Wahlenb.) H.Lindb.
Bryum caespiticum var. kunzei (Hornsch.) Warnst. = Bryum kunzei Hornsch.
Bryum capillare var. meridionale Schimp. = Bryum capillare subsp. meridionale (Schimp.) Podp. = Bryum
capillare Hedw.
Bryum cirratum (Hedw.) With. = Dicranoweisia cirrata (Hedw.) Lindb. ex Milde
Bryum elegans var. carinthiacum (Schimp.) Breidl. = Bryum stirtonii Schimp.
Bryum murale Wils. ex Hunt. = Bryum radiculosum Brid.
Bryum pendulum (Hornsch.) Schimp. = Bryum algovicum Sendtn. ex Müll. Hal.
Bryum provinciale H.Philib. = Bryum canariense var. provinciale (H.Philib.) Husn.
Bryum ventricosum Dicks. = Bryum pseudotriquetrum (Hedw.) Gaertn. Meyer & Scherb.
Buxbaumia indusiata Brid. = Buxbaumia viridis (Moug. ex Lam. & DC.) Moug. & Nestl.
Calliergon cuspidatum (Hedw.) Kindb. = Calliergonella cuspidata (Hedw.) Loeske
Camptothecium aureum (Lag.) Schimp. = Homalothecium aureum (Lag. ex Spruce ) H.Rob.
Camptothecium lutescens Bruch & Schimp. = Homalothecium lutescens (Hedw.) H.Rob.
Camptothecium lutescens var. fallax (Philib.) Breidl. = Homalothecium lutescens var. fallax (Philib. ex Schimp.)
Hedenäs & Söderstr.
Camptothecium sericeum (Hedw.) Kindb. = Homalothecium sericeum (Hedw.) Schimp.
Camptothecium sericeum f. tenella = Homalothecium sericeum f. tenellum Limpr. = Homalothecium sericeum
(Hedw.) Schimp.
Campylium calcareum Crundw. & Nyh. = Campylophyllum calcareum (Crundw. & Nyh.) Heden.
Campylium protensum f. tenera Mönk. (= f. tenue Podp.) = Campylium protensum (Brid.) Kindb.
Catharinea undulata var. minor F.Weber & D.Mohr = Atrichum undulatum var. minus (Hedw.) Paris
Ceratodon chloropus (Brid.) Brid. = Cheilothela chloropus (Brid.) Lindb. ex Broth.
Chrysohypnum chrysophyllum (Brid.) Loeske = Campyliadelphus chrysophyllus (Brid.) R.S.Chopra
Cinclidotus mucronatus (Brid.) Guim. = Dialytrichia mucronata (Brid.) Broth.
Cirriphyllum crassinervium (Taylor) Loeske & M.Fleisch. = Eurhynchium crassinervium (Taylor) Schimp.
Cratoneuron commutatum (Hedw.) Roth. = Palustriella commutata (Hedw.) Ochyra
Cratoneuron commutatum var. falcatum (Brid.) Mönk. = Palustriella falcata (Brid.) Hedenäs
Cratoneuron commutatum var. irrigatum (Zett.) Mönk. = Palustriella commutata var. fluctuans (Schimp.)
Ochyra
Cratoneuron commutatum var. ptychodioides (G.Roth) Mönk. = Palustriella commutata var. ptychodioides
(G.Roth) Ochyra
Cratoneuron decipiens (De Not.) Loeske = Palustriella decipiens (De Not.) Ochyra
Cratoneuron filicinum f. gracilescens (Schimp.) Mönk. = Cratoneuron filicinum var. gracilescens (Brid.)
G.Roth = Cratoneuron filicinum (Hedw.) Spruce
Cratoneuron filicinum var. fallax (Brid.) G.Roth. = Cratoneuron filicinum var. atrovirens (Brid.) Ochyra
Cratoneurum Spruce ex I. Hagen = Cratoneuron (Sull.) Spruce
Crossidium squamigerum (Viv.) Jur. = Crossidium squamiferum (Viv.) Jur. [Orthographic variants]
Desmatodon latifolius (Hedw.) Brid. = Tortula eucalyptrata Lindb.
Dialytrichia brebissonii (Brid.) Limpr. = Dialytrichia mucronata (Brid.) Broth.
Dicranum longifolium Ehrh. ex Hedw. = Paraleucobryum longifolium (Ehrh. ex Hedw.) Loeske
08_Colacino.fm Page 19 Mercredi, 6. septembre 2006 4:11 16
Bryoflora of Albania: A Preliminary Check-List
19
Table 3. List of synonyms.
The citations in the original works considered have been corrected when wrong, as far as possible. When more
than two names are present the last name is the one currently used, the others are synonyms or variants.
Synonyms for “excluded taxa” (i.e., those indicated at the end of Table 1) are also included. Some recent
synonyms are also included.
Dicranum scoparium f. orthophyllum (Brid.) Mönk. = Dicranum scoparium var. orthophyllum Brid.
= Dicranum scoparium Hedw.
Dicranum undulatum Ehrh. = Dicranum polysetum Sw. [≠ Dicranum undulatum Schrad. ex Brid.
= Dicranum bergeri Bland.]
Ditrichium flexicaule (Schleich.) Hampe = Ditrichum flexicaule (Schwägr.) Hampe
Drepanocladus fluitans (Hedw.) Warnst. = Warnstorfia fluitans (Hedw.) Loeske
Encalypta contorta Hoppe ex Lindb. = Encalypta streptocarpa Hedw.
Encalypta rhabdocarpa Schwägr. = Encalypta rhaptocarpa Schwägr. (Orthographic variants)
Encalypta rhabdocarpa var. leptodon Lindb. = Encalypta rhaptocarpa var. trachymitria (Ripart) Wijk &
Margad.
Erythrophyllum rubellum Hilp. = Bryoerythrophyllum recurvirostre (Hedw.) P.C.Chen
Eurhynchium circinatum (Brid.) Schimp. = Scorpiurium circinatum (Brid.) M.Fleisch. & Loeske
Eurhynchium circinatum var. deflexifolium (Solms) Boul. = Scorpiurium deflexifolium (Solms.) M.Fleisch. &
Loeske
Eurhynchium striatum. var. meridionale Schimp. = Eurhynchium meridionale (Schimp.) De Not.
Eurhynchium strigosum (Hoffm. ex F.Weber & D.Mohr) Schimp. = Eurhynchium pulchellum (Hedw.) Jenn.
Eurhynchium strigosum var. diversifolium (Schleich.) Mol. & Lorentz. = Eurhynchium pulchellum var.
diversifolium (Schimp.) C.E.O.Jens.
Eurhynchium swartzii (Turn.) Curn. = Erhynchium hians (Hedw.) Sande Lac.
Eurhynchium swartzii var. abbreviatum Turn. = Eurhynchium schleicheri (Hedw. f.) Jur.
Eurhynchium swartzii var. atrovirens (Brid.) Dixon = Erhynchium hians (Hedw.) Sande Lac.
Eurhynchium swartzii var. robustum Limpr. = Eurhynchium hians var. rigidum (Boul.) Düll
Fissidens cristatus f. mucronata Breidl. = Fissidens dubius var. mucronatus (Breidl. ex Limpr.) Kartt.,
Heden. & Söderstr.
Fissidens cristatus Wils. ex Mitt. = Fissidens dubius P.Beauv.
Funaria dentata Crome = Funaria muhlenbergii Turn.
Funaria mediterranea Lindb. = Funaria muhlenbergii Turn.
Grimmia apocarpa Hedw. = Schistidium apocarpum (Hedw.) Bruch & Schimp.
Grimmia apocarpa var. epilosa (Warnst.) Paris = Grimmia apocarpa Hedw.
Grimmia leucophaea Grev. = Grimmia laevigata (Brid.) Brid.
Gymnostomum calcareum var. viridulum (Brid.) Bruch & Schimp. = Gymnostomum viridulum Brid.
Gymnostomum rupestre Schleich. ex Schwägr. = Gymnostomum aeruginosum Sm.
Herpetineuron toccoae (Sull. & Lesq.) Cardot = Anomodon viticulosus (Hedw.) Hook. & Taylor
Hygrohypnum palustre var. subsphaericarpon (Schleich. ex Brid.) Loeske = Hygrohypnum luridum var.
subsphaericarpon (Schleich. ex Brid.) C.E.O.Jens.
Hylocomium schreberi (Willd. ex Brid.) De Not. = Pleurozium schreberi (Willd. ex Brid.) Mitt.
Hylocomium triquetrum (Hedw.) Schimp. = Rhytidiadelphus triquetrus (Hedw.) Warnst.
Hymenostomum tortile (Schwägr.) Bruch & Schimp. = Weissia condensa (Voit ex Sturm) Lindb.
Hymenostylium curvirostre var. cataractum (Schimp.) Limpr = Hymenostylium recurvirostre (Hedw.) Dixon
Hypnum commutatum Hedw. = Palustriella commutata (Hedw.) Ochyra
Hypnum falcatum Brid. = Palustriella commutata (Hedw.) Ochyra
Hypnum filicinum Hedw. = Cratoneuron filicinum (Hedw.) Spruce
Hypnum sommerfeltii Myr. = Campylophyllum sommerfeltii (Myr.) Heden.
Isothecium filescens (Brid.) Mönk. = Eurhynchium striatulum (Spruce) Schimp.
Isothecium myurum Brid. = Isothecium alopecuroides (Lam. ex Dubois.) Isov.
Lescuraea atrovirens (Dicks. ex Brid.) Kindb. = Pseudoleskea incurvata (Hedw.) Loeske
Lescuraea radicosa (Mitt.) Mönk. = Pseudoleskea radicosa (Mitt.) Macoun & Kindb.
Leskea catenulata (Brid. ex Schrad.) Mitt. = Pseudoleskeella catenulata (Brid. ex Schrad.) Kindb.
Mniobryum albicans (Wahlenb.) Limpr. = Pohlia wahlenbergii (F.Weber & D.Mohr) A.L.Andrews
Mnium affine Bland. = Plagiomnium affine (Bland. ex Funck) T.J.Kop.
Mnium cuspidatum Hedw. = Plagiomnium cuspidatum (Hedw.) T.J.Kop.
08_Colacino.fm Page 20 Mercredi, 6. septembre 2006 4:11 16
20
C. Colacino and M. Sabovljevi†
Table 3. List of synonyms.
The citations in the original works considered have been corrected when wrong, as far as possible. When more
than two names are present the last name is the one currently used, the others are synonyms or variants.
Synonyms for “excluded taxa” (i.e., those indicated at the end of Table 1) are also included. Some recent
synonyms are also included.
Mnium orthorrhynchum Brid. = Mnium thomsonii Schimp.
Mnium punctatum Hedw. = Rhizomnium punctatum (Hedw.) T.J.Kop.
Mnium rostratum Schrad. = Plagiomnium rostratum (Schrad.) T.J.Kop.
Mnium undulatum Hedw. = Plagiomnium undulatum (Hedw.) T.J.Kop.
Myurella julacea var. scabrifolia Lindb. ex Limpr. = Myurella julacea (Schwägr.) Schimp.;
Neckera besseri (Lobarz.) Jur. = Homalia besseri Lobarz.
Orthotrichum leiocarpum Bruch & Schimp. = Orthotrichum striatum Hedw.
Orthotrichum sardagnanum Vent. = Orthotrichum cupulatum var. sardagnanum (Vent.) Vent.
Orthotrichum saxatile Brid. = Orthotrichum anomalum Hedw.
Orthotrichum saxatile Schimp. = Orthotrichum anomalum Hedw.
Orthotrichum striatum var. shawii Wils. = Orthotrichum shawii Wils. in Schimp.
Oxyrrhynchium hians var. rigidum (Boul.) Ochyra & µarnowiec = Eurhynchium hians var. rigidum (Boul.)
Düll
Plagiopus oederi (Brid.) Limpr. = Plagiopus oederianus (Sw.) Crum & Anders.
Plagiothecium silesiacum (F.Weber & D.Mohr) Schimp. = Herzogiella seligeri (Brid.) Z.Iwats.
Plagiothecium silvaticum Bruch & Schimp. in Lindb. = Plagiothecium sylvaticum (Brid.) Bruch & Schimp. =
Plagiothecium nemorale (Mitt.) A. Jaeger
Platyhypnidium riparioides (Hedw.) Podp. = Rhynchostegium riparioides (Hedw.) Cardot
Platyhypnidium rusciforme (Necker) Fleisch. = Rhynchostegium riparioides (Hedw.) Cardot
Platyhypnidium rusciforme var. alopecuroides Brid. = Rhynchostegium alopecuroides (Brid.) A.J.E.Smith
Platyhypnidium rusciforme var. atlanticum Brid. = Rhynchostegium riparioides f. atlanticum (Brid.) Düll
Pogonatum alpinum (Hedw.) Roehl. = Polytrichastrum alpinum (Hedw.) G.L.Sm.
Polytrichum juniperinum Willd. = Polytrichum juniperinum Hedw.
Polytrichum piliferum Schreb. = Polytrichum piliferum Hedw.
Pottia heimii (Hedw.) Schimp. = Desmatodon heimii (Hedw.) Mitt.
Pottia lanceolata var. angustata (Bruch & Schimp.) Müll. Hal = Tortula lanceola var. angustata (Bruch &
Schimp.) Zander
Pseudoleskea atrovirens (Dicks.) Schimp. = Pseudoleskea incurvata (Hedw.) Loeske
Pseudoleskea illyrica G_owacki. = Pseudoleskea saviana (De Not.) Latzel
Racomitrium canescens var. ericoides (Weber) Schimp. = Racomitrium ericoides (Weber ex Brid.) Weber
Rhynchostegiella algiriana (Brid.) Broth. = Rhynchostegiella tenella (Dicks.) Limpr.
Rhynchostegiella algiriana var. litorea (De Not.) Mönk. = Rhynchostegiella litorea (De Not.) Limpr.
Rhynchostegium rusciforme (Neck.) Schimp. = Rhynchostegium riparioides (Hedw.) Cardot
Rhynchostegium tenellum (Dicks.) Schimp. = Rhynchostegiella tenella (Dicks.) Limpr.
Rosulabryum capillare (Hedw.) J.R.Spencer = Bryum capillare Hedw.
Sanionia uncinata f. plumosa (Schimp.) Mönk. = Sanionia uncinata (Hedw.) Loeske
Schistidium alpicola (Hedw.) Limpr. = Schistidium agassizii Sull. & Lesq. ex Sull.
Scleropodium illecebrum Schimp. = Scleropodium touretii (Brid.) L.Koch
Scleropodium purum (Hedw.) Limpr. = Pseudoscleropodium purum (Hedw.) M.Fleisch.
Sphagnum cymbifolium var. pallescens Warnst. = Sphagnum palustre L.
Stereodon cupressiformis (Hedw.) Brid. ex Mitt. = Hypnum cupressiforme Hedw.
Stereodon cupressiformis var. ericetorum Schimp. = Hypnum jutlandicum Holmen & Warncke
Syntrichia inermis (Brid.) Bruch = Tortula inermis (Brid.) Mont.
Syntrichia montana Nees in Raab = Syntrichia intermedia Brid.
Syntrichia ruralis ssp. calcicolens (W.A.Kramer) Düll = Syntrichia calcicola J.J.Amann
Syntrichia ruralis var. calcicola Grebe = Syntrichia calcicola J.J.Amann
Syntrichia ruralis var. ruraliformis (Besch.) Husn. ex T. Durand = Syntrichia ruraliformis (Besch.) Cardot
Syntrichia ruralis var. ruraliformis Dixon = Syntrichia ruraliformis (Besch.) Cardot
Syntrichia subulata (Hedw.) F.Weber & D.Mohr = Tortula subulata Hedw.
Thamnium alopecurum (Hedw.) Schimp. = Thamnobryum alopecurum (Hedw.) Nieuwl. ex Gangulee
Tortella squarrosa (Brid.) Limpr. = Pleurochaete squarrosa (Brid.) Lindb.
08_Colacino.fm Page 21 Mercredi, 6. septembre 2006 4:11 16
Bryoflora of Albania: A Preliminary Check-List
21
Table 3. List of synonyms.
The citations in the original works considered have been corrected when wrong, as far as possible. When more
than two names are present the last name is the one currently used, the others are synonyms or variants.
Synonyms for “excluded taxa” (i.e., those indicated at the end of Table 1) are also included. Some recent
synonyms are also included.
Tortella viridiflava (De Not.) Broth. = Tortella flavovirens var. viridiflava (De Not.) Cas.-Gil.
Tortula aestiva (Brid. ex Hedw.) P.Beauv. = Tortula muralis var. aestiva Brid. ex Hedw.
Tortula montana (Nees) Lindb. (≠ Tortula montana Mitt.) = Syntrichia intermedia Brid.
Tortula montana var. calva (Dur. & Sag.) Limpr. = Syntrichia intermedia var. calva (Dur. & Sag.) Delogn.
Tortula montenegrina (Breidl. & Szysz.) Broth. = Tortula lingulata subsp. montenegrina (Breidl. & Szysz.)
Podp.
Tortula muralis f. incana Sapegen = Tortula muralis Hedw.
Tortula muralis var. rupestris A. Chev. = Barbula muralis var. rupestris Schultz = Tortula muralis Hedw.
Tortula obtusifolia var. brevifolia (Schimp.) G. Roth = Tortula obtusifolia (Schwägr.) Mathieu
Tortula ruralis (Hedw.) P.Gaertn., B.Mey. & Scherb. = Syntrichia ruralis (Hedw.) F.Weber & D.Mohr
Tortula ruralis subsp. ruraliformis (Besch.) Dixon = Syntrichia ruraliformis (Besch.) Cardot
Trichostomum brevifolium Sendtn. ex Müll. Hal. = Trichostomum crispulum var. brevifolium (Müll. Hal.)
Bruch & Schimp.
Trichostomum mutabile Bruch = Trichostomum brachydontium Bruch
Ulota americana Mitt. = Ulota curvifolia (Wahlenb.) Lilj.
Weissia crispata (Nees & Hornsch.) Müll. Hal.= Weissia controversa var. crispata (Nees & Hornsch.) Nyh.
Weissia tortilis (Schwägr.) Müll. Hal. = Weissia condensa (Voit ex Sturm) Lindb.
Weissia viridula Hedw. ex Brid. = Weissia controversa Hedw.
The fluctuation of borders in earlier times is one of the reasons for the
exclusion of several records which are outside the present-day borders of Albania
(which have not changed since 1921, however). Many records from Szepesfalvy
(1926), in particular those which are based on the collections made by
Andrasowsky in 1916-17, had to be excluded, as what he called Northern Albania
referred actually to areas in Montenegro or Kosova. Some works which included
in their title Albania (or, more generally, the Balkans) have been consulted also,
but in several cases there were no records of bryophytes for Albania in them.
These have been reported too. For three taxa the only record of their presence in
Albania is by Düll (1984, 1985). There is no original source, and no locality, and
therefore, these have not been included in the list (see also Düll’s notes, later).
The most recent paper on the liverworts of Albania is that of Bischler et al. (1980),
which reports about 59 taxa, some of them indicated as new to Albania, although
they had already been collected at the turn of the 19th Century and reported in
works (Baumgartner, 1915; Höhnel, 1893, 1894) apparently overlooked by
Bischler et al. (1980).
The determination of the actual localities from the spellings used by the
original authors has presented several problems, not least the lack of an official
Albanian alphabet, used by all speakers of Albanian, which was agreed upon only
in 1915, but entered into general use much later. The existence of two main
varieties of Albanian (Gheg and Tosk, the last one becoming the “standard form”
only after 1945) sometimes added to these difficulties. Therefore, many localities
in the works consulted were indicated with their Turkish language equivalents
(e.g., Ipek instead of the currently used Pejë/Pe† [Albanian/Serbian, respectively;
a town in Kosova]), or with spellings based on German or Slavic languages
(Serbian mainly). The use of maps and of gazetteers (for Albania and the
08_Colacino.fm Page 22 Mercredi, 6. septembre 2006 4:11 16
22
C. Colacino and M. Sabovljevi†
Fig. 1. The administrative boundaries of Albanian Prefectures with their names. – The
Prefectures are indicated with the following acronyms, from North to South: SHK = Shkodër;
KUK = Kukës; LZH = Lezhë; DIB = Dibër; DUR= Durrës; TIR = Tiranë; ELB = Elbasan;
FIE = Fier; KOR = Korçë; BER = Berat; VLO = Vlorë; GJI = Gjirokastër. (Boundaries
according to: Harta Administrative [GCC, 1999]).
08_Colacino.fm Page 23 Mercredi, 6. septembre 2006 4:11 16
Bryoflora of Albania: A Preliminary Check-List
23
Fig. 2. Number of liverworts (including hornworts) taxa and in brackets number of collections
(records). Total for Albania 89 [559]. - Und. = Undeterminded, utilized in a few instances when
localization was not possible (e.g., “Everywhere in mountains above 1000 m”).
08_Colacino.fm Page 24 Mercredi, 6. septembre 2006 4:11 16
24
C. Colacino and M. Sabovljevi†
Fig. 3. Number of moss taxa and in brackets number of collections (records); Total for Albania
238 [604]. - Und. = Undeterminded, utilized in a few instances when localization was not possible
(e.g., “Everywhere in mountains above 1000 m”).
08_Colacino.fm Page 25 Mercredi, 6. septembre 2006 4:11 16
Bryoflora of Albania: A Preliminary Check-List
25
bordering countries) has allowed us, however, to locate all localities with the
exception of one. The use of a series of topographic maps of Albania at the scale
1:50,000 published by the Russian Army between 1974 and 1988 also has been
useful, even though the spellings of Albanian names are reported there in their
phonetic rendering in the cyrillic script (Electronic Access from UC Berkeley
Library [http://www.lib.berkeley.edu/EART/albania/albania50k.html]).
A data base of the bryoflora of Albania, based at the Laboratory of
Bryology (Dipartimento di Biologia, Difesa e Biotecnologie Agroforestali) of the
University of Basilicata, has been prepared, and it would be appreciated if all new
records (and related publications) would be communicated to the corresponding
author to keep the data base up to date.
RESULTS AND DISCUSSION
A Check-list of the Bryophytes of Albania is given in Table 1. We report
here 327 taxa (species, subspecies, varieties and in a few instances, forms), which
almost doubles the number of records for Albania as indicated by Düll (1983,
1984, 1985), which consisted of 175 entities. In particular, 238 mosses are reported
(109 in Düll, 1984, 1985, 1992, including the 4 excluded herein), 86 liverworts, and
3 hornworts (63 + 3 in Düll, 1983). The taxa indicated by Söderström et al. (2002),
with the exclusion of three of them, are the same we report here. The check-list
is based on a total of 1207 records (including 44 records from localities reported
in the works considered as in Albania, but actually outside present-day Albanian
borders, and which have been indicated in Table 1 under “Records excluded”).
All records, without a valid location in Albania, have been excluded.
The actual number of bryophytes species in Albania is very likely much
higher than that reported here (Bego & Koni, 1999, estimate it at about
500 species, a number probably not far from reality). This is suggested by the
number of taxa in neighboring countries (see e.g., Düll, 1995; Sabovljevi† et al.,
2001; Blockeel et al., 2002; Pavleti†, 1968, Sabovljevi†, 2004) as well as the floristic
and geographic diversity of the country itself, an important meeting area between
the Mediterranean, the central European, and the Pontic floristic elements (for
more information on these characteristics refer also to Colacino (2004) and the
references indicated therein).
Excluded bryophyte taxa
Nineteen taxa, excluded from the Albanian Flora, are reported at the end
of Table 1. Campylophyllum calcareum (Crundw. & Nyholm) Hedenäs,
Didymodon cordatus Jur., Didymodon vinealis var. flaccida (Bruch & Schimp.)
Zander, and Hypnum cupressiforme var. lacunosum Brid., reported in Düll (1983,
1985), have no indication of any voucher or locality and have therefore been
excluded (see also the notes by Düll at the end of Table 1). Grimmia hartmanii
var. montenegrina Beck & Szysz., reported for Albania by Podp{ra (1954) and
Düll (1984), is actually from Montenegro. The records of Atrichum undulatum
var. minus (Hedw.) Paris, Barbula unguiculata f. obtusifolia Mönk., Desmatodon
heimii (Hedw.) Mitt., Fissidens adianthoides Hedw., Hypnum jutlandicum Holmen
& Warncke, Thuidium abetinum (Hedw.) Schimp., Thuidium tamariscinum
08_Colacino.fm Page 26 Mercredi, 6. septembre 2006 4:11 16
26
C. Colacino and M. Sabovljevi†
(Hedw.) Schimp., and Tortula mucronifolia Schwägr., reported for Albania, are
actually from localities in Kosova. The records of Pogonatum aloides var.
x minimum (Crome) Molendo, Tortula lingulata subsp. montenegrina (Breidl. &
Szysz.) Podp., Weissia controversa var. crispata (Nees & Hornsch.) Nyh., and
Leiocolea heterocolpos (Thed. ex Hartm.) H.Buch., also reported for Albania, are
actually from localities in Montenegro.
Three taxa reported for Albania by Söderström et al. (2002) have also
been excluded: Porella obtusata (Taylor) Trevis, Riccia lamellosa Raddi, both
without indication of any voucher or locality, and Leiocolea heterocolpos (Thed.
ex Hartm.) H.Buch, whose report is based on the same record by Szepesfalvy
(1926), which refers to a locality in Montenegro. For more details refer to the
notes at the end of Table 1.
Excluded Works
The following works refer in their title to the Balkan area, or to Albania, in
none of them, however, records of bryophytes for Albania are to be found. In
particular, Ade & Koppe (1955) report species from Macedonia, Greece, Bulgaria,
Serbia, and southern Italy, near Naples, while none is reported from Albania.
Baumgartner (1914a, 1914b) reports several bryophytes from the Island of Corfu
(Greece), while his collections from localities on the adjacent coast of Albania do
not include any bryophytes. Vilhelm (1923) reports species from northern
Montenegro only. Weiss (1866) reports only Vascular Plants from Albania. Düll
(1996), eventually, cites a work by Unger (1862) for Albania. We have not been
able to locate it; its title, however, seems to refer to areas south of Albania.
Systematic synopsis of Albanian bryophytes (Table 2)
Arrangement follows Crandall-Stotler & Stotler (2000) for liverworts.
Given the state of flux of hornworts taxonomy (Renzaglia & Vaughn, 2000), and
considered there are only two genera of this group in our check-list, we follow the
traditional arrangement of Grolle & Long (2000). For mosses the arrangement is
according to Buck and Goffinet (2000), with the exception of Pottiaceae arranged
according to Zander (1993). We differ from Grolle & Long (2000) in considering
the bryophytes made up by three independent lineages (Divisions): Bryophyta
(Mosses), Marchantiophyta (Liverworts), and Anthocerotophyta (Hornworts).
We report for Albania 141 genera and 62 families of bryophytes (s.l.).
Synonyms (Table 3)
This list of synonyms is based on those found in the original sources, with
a few additions (e.g., recent synonymies). Errors in the original citations have
been corrected, as far as possible.
Discussion
Albania is still a relatively unexplored area in Europe, not all areas within
that country, however, are equally unknown from a bryological standpoint. Coastal
areas, as well as those near Tirana and near Scutari (Shkodër), have been more
08_Colacino.fm Page 27 Mercredi, 6. septembre 2006 4:11 16
Bryoflora of Albania: A Preliminary Check-List
27
frequently visited and collected. The less known areas appears to be the Northern
Alps, the areas near the border with Kosova, and the internal areas of Berat and
Elbasan. Even for those areas with the highest numbers of mosses and liverworts
(and hornworts), as indicated in figures 2 and 3, these have been usually collected
along the roads and near the major inhabited areas (with a few notable exceptions).
A description of the characteristics of the bryophyte flora of Albania (as attempted
in Colacino, 2004) is, therefore, probably still premature, even though it appears
there is a prevalence of moss taxa with a central European distribution, followed by
boreal and sub-boreal ones (Colacino, 2004). This is confirmed also by the results of
Bischler et al. (1980) who found, for liverworts, a high number of central European
taxa also (even though they did collect mainly in coastal areas). This is in opposition
to the characteristics of the vascular flora which is made up prevalently by
Mediterranean (and Balkan) elements (see, for instance Bego & Koni, 1999;
Vangjeli et al., 1995). As reported by Stewart (1995), it is likely that the northern
mountains have a bryophyte flora typical of Balkan mountains, with possibly rare
species and arctic-alpine relicts. Interesting (and bryologically almost unexplored)
areas on the Adriatic Coast (including the sand dunes and lagoon of the National
Park of Divjaka) are under severe pressure because of human exploitation, and it is
not known for how long (and if) they will be effectively protected (see also Leone et
al., 2003, for recent data on Divjaka Lagoon).
According to Stewart (1995), there are nine bryophyte species of
particular conservation interest in Albania: the only locally endemic species, the
rare (R) liverwort Frullania illyrica Grolle, is now included in F. inflata Gottsche,
with a wider distribution, and whose status is Vulnerable (V) according to
Schumacker & Vá≈a (2005). There are four other liverworts, Athalamia spathysii
(Lindenb.) S. Hatt., Mannia triandra (Scop.) Grolle, Riccia trabutiana Steph., all
with the status of rare according to the European Red List, and the insufficiently
known (K) Marchantia palacea Bertol. (but not threatened – NT, according to
Schumacker & Vá≈a, 2005). Out of the four mosses cited therein, Tortula lingulata
Lindb. (K) is known from a single locality which we found to be outside the
present borders of the country (actually we report Tortula lingulata subsp.
montenegrina (Breidl. & Szysz.) Podp., which is considered by some authors (e.g.
Düll, 1984) as synonym of Tortula lingulata Lindb.). The other mosses are
Neckera cephalonica Jur. & Unger. (K), Buxbaumia viridis (Moug. ex Lam. &
DC.) Moug. & Nestl. (V), and Tortula solmsii (Schimp.) Limpr. (R), the latter
collected in Albania only once, more than a century ago (Höhnel, 1893). There
are no data available on bryophyte conservation at the national level, the Red
Data Book for Albania (Vangjeli et al., 1995) did not include bryophytes.
Acknowledgements. We thank the many people who helped us to get some of the
older references, and the referees, R.M. Ros and T.Blockeel, for valuable comments.
REFERENCES
ADE A. & KOPPE F., 1955 — Moose aus dem Balkan, aus Süditalien und von den
Äegäischen Inseln. Acta Musei Macedonici scientiarum naturalium 2: 181-197.
BAUER C.R. & CROSBY M.R., 1986 — Index muscorum supplementum: 1982-1983.
Taxon 35: 416-439.
BAUMGARTNER J., 1914a — Hepaticae. In: Rechinger K, Beiträge zur
Kryptogamenflora der Inseln Korfu nebst einegen Standorten von der
albanischen Küste. I. Teil. Verhandlungen der kaiserlich königlichen zoologischbotanischen Gesellschaft (Wien) 64: 142-143.
08_Colacino.fm Page 28 Mercredi, 6. septembre 2006 4:11 16
28
C. Colacino and M. Sabovljevi†
BAUMGARTNER J., 1914b — Musci. In: Rechinger K, Beiträge zur Kryptogamenflora
der Inseln Korfu nebst einegen Standorten von der albanischen Küste. I. Teil.
Verhandlungen der kaiserlich königlichen zoologisch-botanischen Gesellschaft
(Wien) 64: 141-142.
BAUMGARTNER J., 1915 — Verzeichnis der von I. Dörfler auf seiner Reise im
albanisch-montenegrinischem Grenzgebiete im Jahre 1914 gesammelten Moose.
Österreichische botanische Zeitschrift 65 (10-12): 312-319.
BECK G. & SZYSZYfiOWICZ I., 1888 — Plantae à Dre Ign. Szyszyflowicz in itinere per
Cernagoram et in Albania adjacente anno 1886 lectae. Cracoviae, Universitatis
Jagellonicae.
BEGO F. & KONI M. (Eds), 1999 — Biodiversity Strategy and Action Plan. Tirana,
National Environmental Agency (NEA).
BISCHLER H., JOVET-AST S. & BAUDOIN R., 1980 — Hépatiques de la côte
albanaise. Cryptogamie, Bryologie-Lichénologie 1: 247-267.
BISCHLER H., 2004 — Liverworts of the Mediterranean. Ecology, diversity and
distribution. Bryophytorum bibliotheca 61: 1-252.
BLOCKEEL T.L., ROS R.M., SABOVLJEVI± M., CANO M.J., GALLEGO M.T. &
MUÑOZ J., 2002 — New and interesting bryophyte records for Greece.
Cryptogamie, Bryologie, 23: 149-155.
BUCK W.R. & GOFFINET B., 2000 — Morphology and classification of mosses. In: Shaw
A.J. & Goffinet B. (eds), Bryophyte Biology. Cambridge, Cambridge University
Press, pp. 71-123.
COLACINO C., 2004 — Brioflora dell’Albania: Situazione attuale e considerazioni in
relazione ad una possibile utilizzazione nel biomonitoraggio ambientale.
S.I.S.E.F. Atti 4: 357-364.
CORLEY M.F.V. & CRUNDWELL A.C., 1991 — Additions and amendments to the
mosses of Europe and the Azores. Journal of bryology 16: 337-356.
CORLEY M.F.V., CRUNDWELL A.C., DÜLL R., HILL M.O. & SMITH A.J.E., 1981 —
Mosses of Europe and the Azores: An annotated list of species, with synonyms
from the recent literature. Journal of bryology 11: 609-689.
CORTINI PEDROTTI C., 2001a — New Check-list of the Mosses of Italy. Flora
Mediterranea 11: 23-107.
CORTINI-PEDROTTI C., 2001b — Flora dei muschi d’Italia. Sphagnopsida, Andreaeopsida,
Bryopsida (I parte).Roma, A. Delfino Editore.
CORTINI-PEDROTTI C., 2006 — Flora dei muschi d’Italia. Bryopsida (II parte). Roma,
A. Delfino Editore.
CRANDALL-STOTLER B. & STOTLER R.E., 2000 — Morphology and classification of
the Marchantiophyta. In: Shaw A.J. & Goffinet B. (eds), Bryophyte Biology.
Cambridge, Cambridge University Press, pp. 21-70.
CROSBY M.R., 1977 — Index muscorum supplementum: 1974-1975. Taxon 26: 285-307.
CROSBY M.R., 1979 — Index muscorum supplementum: 1976-1977. Taxon 28: 237-264.
CROSBY M.R. & BAUER C.R., 1981 — Index muscorum supplementum: 1978-1979.
Taxon 30: 667-693
CROSBY M.R. & BAUER C.R., 1983 — Index muscorum supplementum: 1980-1981.
Taxon 32: 670-691.
CROSBY M.R. & BAUER C.R., 1987 — Index muscorum supplementum: 1984-1985.
Taxon 36: 502-527.
CROSBY M.R., MAGILL R.E & BAUER C.R., 1992 — Index of Mosses 1963-1989.
Monographs in systematic botany from the Missouri Botanical Garden 42.
CROSBY M.R. & MAGILL R.E., 1994 — Index of Mosses 1990-1992. Monographs in
systematic botany from the Missouri Botanical Garden 50.
CROSBY M.R. & MAGILL R.E., 1997 — Index of Mosses 1993-1995. Monographs in
systematic botany from the Missouri Botanical Garden 62.
CROSBY MR & MAGILL R.E., 2000 — Index of Mosses 1996-1998. Monographs in
systematic botany from the Missouri Botanical Garden 80.
CROSBY M.R. & MAGILL R.E., 2004 — Index of mosses, 1999-2001. Monographs in
systematic botany from the Missouri Botanical Garden 96.
08_Colacino.fm Page 29 Mercredi, 6. septembre 2006 4:11 16
Bryoflora of Albania: A Preliminary Check-List
29
CROSBY M.R. & MAGILL R.E., 2005 — Index of bryophytes, 2001-2004. Sidre Presse,
Sugar Grove (NC), Missouri Botanical Garden.
DIERßEN K., 2001 — Distribution, ecological amplitude and phytosociological
characterization of European bryophytes. Bryophytorum bibliotheca 56: 1-289.
DÜLL R., 1983 — Distribution of the European and Macaronesian Liverworts
(Hepaticophytina). Bryologische Beiträge 2: 1-114.
DÜLL R., 1984 — Distribution of the European and Macaronesian Mosses (Bryophytina).
Part I. Bryologische Beiträge 4: 1-113.
DÜLL R., 1985 — Distribution of the European and Macaronesian Mosses (Bryophytina).
Part II. Bryologische Beiträge 6: 110-232.
DÜLL R., 1992 — Distribution of the European and Macaronesian Mosses (Bryophytina).
Annotations and Progress. Bryologische Beitraege 8/9: 1-223.
DÜLL R., 1995 — Moose Griechenlands — Bryophytes of Greece. Bryologische Beiträge
10: 1-125.
DÜLL R., 1996 — The current state of bryophyte investigation in the Mediterranean area.
Bocconea 5: 271-278.
GCC, 1999. Republika e Shqipërisë — Harta Administrative. Tiranë. (May 1999:
Administrative Divisions updated to March 1997).
GROLLE R. & LONG, D. G., 2000 — An annotated check-list of the Hepaticae and
Anthocerpthae of Europe and Macaronesia. Journal of bryology 22: 103-140.
HÖHNEL F., 1893 — Beitrag zur Kenntniss der Laubmoosflora des Küstenstriches vom
Görzer Becken bis Skutari in Albanien. Österreiches botanische Zeitschrift (Wien)
43: 381-412.
HÖHNEL F., 1894 — Beitrag zur Kenntniss der Laubmoosflora des Küstenstriches vom
Görzer Becken bis Skutari in Albanien. Österreichische botanische Zeitschrift
(Wien) 44: 23-27.
JOVET-AST S., 1986 — Les Riccia de la région méditerranéenne. Cryptogamie, BryologieLichénologie 7: 283-431.
KÁRPÁTI L. & VAJDA L., 1961 — Beiträge zur Moosflora Albaniens. Fragmenta
botanica musei historico-naturalis Hungarici 1 (1-4): 3-16.
LEONE V., MARCHIORI S., COLACINO C., FASCETTI S., MEDAGLI P.,
SARACINO A., SEMERARI P., 2003 — Habitat della Laguna di Karavasta e
delle zone contermini: considerazioni e restauro. In: Baldassarre G., (ed.),
Salvaguardia e sviluppo sostenibile dell’area lagunare di Karavasta (Albania).
Atti del seminario: Divjaka (Albania) 9 maggio 2003. Università di Bari (Edizioni
dal Sud), Bari, pp. 55-65.
MARKGRAF F., 1927a — Laubmoose. In: Markgraf F, An den Grenzen des
Mittelmeergebiets. Pflanzengeographie von Mittelalbanien. Repertorium
specierum novarum regni vegetabilis 45: 164-166.
MARKGRAF F., 1927b — Lebermoose. In: Markgraf F, An den Grenzen des
Mittelmeergebiets. Pflanzengeographie von Mittelalbanien. Repertorium
specierum novarum regni vegetabilis 45: 164.
MARKGRAF F., 1931 — Pflanzen aus Albanien. Denschriften der kaiserlichen Akademie
der Wißenschaften. Mathematisch-Naturwißenschaftliche Klasse 102: 317-360.
MEYER F. K. & GROLLE R., 1963 — Eine neue Frullania-Art aus Albanien. Feddes
repertorium Bd. 68: 101-107.
MEYER F. K. & GROLLE R., 1968 — Lebermoose aus Albanien, Bulgarien und dem
Kaukasus. Wissenschaftliche Zeitschrift der Friedrich-Schiller-Universität Jena.
Mathematisch-Naturwißenschaftliche Reihe 17(3): 363-367.
PAVLETI± Z., 1968 — Flora mahovina Jugoslavije. Zagreb, Vjesnik.
PETROV S., 1960 — Contribution à la flore bryologique de l’Albanie. Revue Bryologique
et Lichénologique 29: 212-234.
PETROV S., 1962 — Zweiter Beitrag zur Moosflora Albaniens. B’lgarska Akademija na
Naukite — Izvestija na Botaniªeskija Institut 9: 185-189 (In Bulgarian) [Пeтpoв Сл
(1962). Bтopи принос към мъховата флора на Албания. Ъългарска Академия на
Нaуките. Ицвестия на Ъотаническия Институт. кн. ix/1962: 185-189.]
08_Colacino.fm Page 30 Mercredi, 6. septembre 2006 4:11 16
30
C. Colacino and M. Sabovljevi†
PODP}RA J., 1954 — Conspectus Muscorum Europaeorum. Praga, Nakl-Ωeskoslovenské
Akademie V{d, 700p.
RENZAGLIA K.S. & VAUGHN K.C., 2000 — Anatomy, development, and classification
of hornworts. In: Shaw A.J. & Goffinet B. (eds), Bryophyte Biology. Cambridge,
Cambridge University Press, pp. 1-20.
SABOVLJEVI± M., 2004 — Comparison of the bryophyte flora of the three southern
European maindlands: the Iberian, the Apennine and the Balkan peninsulas.
Braun-Blanquetia 34: 21-28.
SABOVLJEVI± M., GANEVA A., TSAKIRI E. & STEFANUT S., 2001 — Bryology and
bryophyte protection in south-eastern Europe. Biological conservation 101: 73-84.
SCHUMACKER R. & VÁ~A J., 2005 — Identification Keys to the Liverworts and
Hornworts of Europe and Macaronesia (Distribution and Status). Ed. 2. Pozna¬,
Sorus.
SMITH, A.J.E., 2004 — The Moss Flora of Britain and Ireland. Ed. 2. Cambridge,
Cambridge University Press.
SÖDERSTRÖM L., HALLINGBÄCK T., HODGETTS N., RAEYMAEKERS G.,
SCHUMACKER R., SÉRGIO C., STEWART N. & VÁ~A J., 1998 — State of
knowledge of the bryoflora of Europe as illustrated by the hepatic flora.
Lindbergia 23: 28-32.
SÖDERSTRÖM L., URMI E. & VÁ~A J., 2002 — Distribution of Hepaticae and
Anthocerotae in Europe and Macaronesia. Lindbergia 27: 3-47.
STEWART N. (ed.), 1995 — Red Data Book of European Bryophytes. Trondheim, ECCB.
SZEPESFALVY J. V., 1926 — Bryophyta. In: Csiki E., Jávorka A., Kümmerle E. B. (eds),
Adatok Albánia Flórájához-Additamenta ad floram Albaniae. Budapest, A
Magyar Tudományos Akadémia Kiadása, pp. 180-196.
UNGER F., 1862 — Wissenschaftliche Ergebnisse einer Reise in Griechenland und in den
ionischen Inseln. Wien.
VANGJELI J., RUCI B. & MULLAJ A., 1995. Libri i kuq: Bimët e kërcënuara dhe të rralla
të Shqipërisë (Red Book: Threatened and rare plants species of Albania). Tiranë,
Akademia e Shkencave Instituti i Kërkimeve Biologjike.
VILHELM J., 1923 — Addi[t]amenta floristica in bryofloram montenegrinam. Acta
botanica Bohemica 2: 46-50.
WEISS E., 1866 — Floristisches aus Istrien, Dalmatien und Albanien. Verhandlugen der
kaiserlich-königlichen zoologisch-botanischen Gesellschaft (Wien) 16: 572-584.
WIJK R.V.D., MARGADANT W.D. & FLORSCHUETZ P.A., 1959 — Index
Muscorum., Vol. 1 (A-C). Regnum Vegetabile 17.
WIJK R.V.D., MARGADANT W.D. & FLORSCHUETZ P.A., 1962 — Index Muscorum,
Vol. 2 (D-Hypno). Regnum Vegetabile 26.
WIJK R.V.D., MARGADANT W.D. & FLORSCHUETZ P.A., 1964 — Index Muscorum,
Vol. 3 (Hypno-O). Regnum Vegetabile 48.
WIJK R.V.D., MARGADANT W.D. & FLORSCHUETZ P.A., 1967 — Index Muscorum,
Vol. 4 (P-S). Regnum Vegetabile 48.
WIJK R.V.D., MARGADANT W.D. & FLORSCHUETZ P.A., 1969 — Index Muscorum,
Vol. 5 (T-Z). Regnum Vegetabile 65.
ZANDER R.H., 1993 — Genera of the Pottiaceae: Mosses of Harsh Environments.
Bulletin of the Buffalo society of natural sciences 32: 1-378.
Marko Sabovljevic