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WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
SpeciesMapsDocumentsIDAO

Alternanthera sessilis (L.) R.Br. ex DC.

Accepted
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
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Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
/7d46cca0-d9f9-47e8-a261-178c3715e347/956.jpg
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
Alternanthera sessilis (L.) R.Br. ex DC.
🗒 Synonyms
synonymAchyranthes linearifolia Sw. ex Wikstr.
synonymAchyranthes sessilis (L.) Besser
synonymAchyranthes triandra Roxb.
synonymAchyranthes villosa Blanco
synonymAllaganthera forsskaolei Mart.
synonymAlternanthera achyranthes Forssk.
synonymAlternanthera achyranthoides Forssk.
synonymAlternanthera achyranthoides Hiern
synonymAlternanthera angustifolia R.Br.
synonymAlternanthera angustifolia var. lanata Benth.
synonymAlternanthera angustifolia var. macrocephala Domin
synonymAlternanthera angustifolia var. typica Domin
synonymAlternanthera denticulata R.Br.
synonymAlternanthera denticulata var. major Moq.
synonymAlternanthera denticulata var. micrantha Benth.
synonymAlternanthera denticulata var. uliginosa Domin
synonymAlternanthera dubia Moq. [Illegitimate]
synonymAlternanthera ficoides P.Beauv.
synonymAlternanthera ficoides var. minor P.Beauv.
synonymAlternanthera ficoides var. versicolor Lem.
synonymAlternanthera glabra Moq.
synonymAlternanthera major (Benth.) Domin
synonymAlternanthera micrantha (Benth.) Domin [Illegitimate]
synonymAlternanthera nana R.Br.
synonymAlternanthera nana var. major Benth.
synonymAlternanthera nodiflora R.Br.
synonymAlternanthera nodiflora var. lanceolata Moq.
synonymAlternanthera nodiflora var. linearifolia Moq.
synonymAlternanthera polygonoides (L.) R.Br. ex Sweet
synonymAlternanthera polygonoides (L.) R.Br. ex Sweet
synonymAlternanthera prostrata D.Don
synonymAlternanthera repens J.F.Gmel.
synonymAlternanthera sennii Mattei
synonymAlternanthera sessilis f. lanceolata Kuntze
synonymAlternanthera sessilis f. spathulifolia Kuntze
synonymAlternanthera sessilis var. angustifolia Moq.
synonymAlternanthera sessilis var. denticulata (R.Br.) Kuntze
synonymAlternanthera sessilis var. major Moq.
synonymAlternanthera sessilis var. neglecta Kuntze
synonymAlternanthera sessilis var. nodiflora (R.Br.) Kuntze
synonymAlternanthera sessilis var. nodiflora (R.Br.) Kuntze
synonymAlternanthera sessilis var. parviflora Kuntze
synonymAlternanthera sessilis var. stauntonii Moq.
synonymAlternanthera sessilis var. tenuissima (Suess.) Backer
synonymAlternanthera sibirica (De Vest ex Roem. & Schult.) Steud.
synonymAlternanthera tenella Moq. [Illegitimate]
synonymAlternanthera tenuissima Suess.
synonymAlternanthera triandra Lam. [Illegitimate]
synonymAlternanthera triandra var. denticulata (R.Br.) Maiden & Betche
synonymAlternanthera triandra var. nodiflora (R.Br.) Maiden & Betche
synonymAlternanthera uliginosa (Domin) Dinter
synonymGomphrena brasiliensis Moq. [Invalid]
synonymGomphrena polygonoides L.
synonymGomphrena sessilis L.
synonymIllecebrum sessile (L.) L.
synonymParonychia sessilis (L.) Desf.
synonymTelanthera polygonoides (L.) Moq.
🗒 Common Names
Chinese
  • Lián zǐ cǎo, 莲子草
Comorian
  • Mgawa
Creoles and pidgins; French-based
  • Brède emballage, Brède chevrette (La Réunion)
  • Maglwa, Magwa, Vwè klè (Antilles)
English
  • Sessile joyweed
  • Sessile flowered globe amaranth
  • Large purple alternanthera, Dwarf copperleaf, Red sessile joyweed, Sessile alligator weed
French
  • Herbe d'emballage, Magloire
Hindi
  • Garundi, Guroo
Indonesian
  • Bayam keremak merah
Malagasy
  • Anabalaza
Other
  • Hanga feliky tariky, Sary mlalyhapana, Sary mlalyhapana kely (Kibushi, Mayotte)
Spanish; Castilian
  • Colchón de nino, Coyuntura, Hierba de perico, Paja blanca, Sanguinaria
📚 Overview
Overview
Brief

Code

ALRSE

Growth form

Broadleaf

Biological cycle

Annual to vivacious

Habitat

terrestrial

Wiktrop
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ravi luckhun
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    Diagnostic Keys
    Description

    Global description 

    Alternanthera sessilis is a highly branched herbaceous plant common in cultures and ruderal places, in moist soil preferably, but also in dry environments as in flooded areas. Its general appearance varies greatly, depending on the environment. It is a small plant with slender stems in dry environments, it is a creeping, prostrate plant, with erect stems in humid areas and a tall plant in aquifers with flooded areas. The stem is 30 cm to 1 m in length, striated and becoming hollow in flooded areas. Roots emerge at the nodes, at the points of contact with the ground or when the plant is floating. It is generally glabrous. The leaves are simple and opposite, sub sessile, glabrous or nearly. The blade is of very variable shape, and is generally rather narrow and elongated, and lanceolate. The inflorescences, isolated or grouped in globose or ovoid clusters are whitish sessile glomeruli, not spiny, and inserted into the base of the leaves.

    General habit

    Alternanthera sessilis is a highly branched plant with a highly variable size, depending on the degree of humidity of the environment: in dry areas, it is erect and can reach 30 cm long; in humid areas, it is prostrate and then erect, with stems up to 10 cm to 1 m. In flooded areas, it is floating and reaches several meters long.

    Underground system

    The principal root is a taproot.  Adventitious roots arise at the nodes in contact with the ground when the plant is prostrate. In aquatic environments, the nodes emit beams of long white roots
     
    Stem

    The stem is cylindrical and extensively branched; it becomes hollow in flooded situation. It is striated, as well as the branches. It is rigid and fine in dry environments, it can measure 1 cm in diameter in aquatic environment where it is long and floating. It is glabrous, with the exception of two narrow and opposite bands with whitish hairs on the erect parts and tufts of white hairs at the level of the nodes.

    Leaf
     
    The leaves are simple, opposite and decussate with leaf blade generally of the same size. The petiole, slightly distinct, is short (1.5 mm) or even absent. The leaf blade is narrow and elongated and is very variable in shape and size. It is oblanceolate to elliptic-lanceolate or linear-lanceolate, acute to obtuse, shortly acuminate, base angled to attenuate, from 1 to 12 cm long for 0.25 to 3 cm wide. It is glabrous or nearly so, some with very sparse hairs and which are mostly found on the underside on the midrib.

    Inflorescence

    The sessile inflorescences are in axillary position and are isolated or grouped (up to 5). They are subglobose, they measure between 5 and 7 mm in diameter and are white.

    Flower
     
    Each flower is subtended by a scabrous, white, and oval with pointed tip bract , 0.75-1 mm long and 2 bracteoles similar to the bracts. The perianth consists of tepals which are equal, ovate-elliptic, pointed at the top, 1.5 to 2.5 mm long, white, hairless, clearly mucronate and with a slightly denticulate margin. The 5 stamens (Including 2 without anther) are alternate with the pseudostaminodes similar to the filaments, but generally a little smaller. At maturity, they equal the ovary and style. The sub-orbicular ovary, is compressed with short style.

    Fruit

    The fruit is an obcordate to orbicular-obcordate utricle, 2 to 3 mm long, dark brown with paler edge. It falls together with the tepals.

    Seed

    The seed is lenticular, 0.5 to 1 mm in diameter, glossy brown seed coat, slightly reticulate.

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      No Data
      📚 Natural History
      Life Cycle

      Life cycle

      Annual
      Annual

      China: Alternanthera sessilis flowers and fruits from May to July.
      Mayotte: Alternanthera sessilis flowers and fruits all year round.
      Nicaragua: Alternanthera sessilis flowers and fruits all year round.
      West Indies: Alternanthera sessilis flowers and fruits all year round.

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        Reproduction

        Alternanthera sessilis is a herbaceous plant that can be annual or perennial depending on the humidity of the environment. In lowland conditions or in irrigated rice fields, it is most often a perennial. Dissemination by seed or vegetative propagation by cuttings of stem fragments. The fruits are myrmecophilous.

        Thomas Le Bourgeois
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          Morphology

          Growth form

          Prostrated
          Prostrated

          Type of prefoliation

          Leaf ratio medium
          Leaf ratio medium
          Narrow leaf
          Narrow leaf

          Equality of opposite leaves

          Opposite leaves equal
          Opposite leaves equal

          Latex

          Without latex
          Without latex

          Root type

          Taproot
          Taproot
          Fibrous roots
          Fibrous roots

          Hollow or solid stem

          Solid stem
          Solid stem
          Hollow stem
          Hollow stem

          Stipule type

          No stipule
          No stipule

          Leaf attachment type

          sessile
          sessile
          with petiole
          with petiole

          Lamina base

          attenuate
          attenuate

          Lamina apex

          apiculate
          apiculate
          attenuate
          attenuate
          acute
          acute
          obtuse
          obtuse

          Upperface pilosity

          Glabrous
          Glabrous
          Less hairy
          Less hairy

          Simple leaf type

          Lamina elliptic
          Lamina elliptic

          Stem pilosity

          Less hairy
          Less hairy

          Stem hair type

          Aligned pilosity
          Aligned pilosity

          Life form

          Broadleaf plant
          Broadleaf plant
          Look Alikes

          A. sessilis can be confused with A. philoxeroides, the latter species is distinguished by its inflorescences which is also a white globose spikelet, but is held by a 1-9 cm long peduncle.

          A. pungens A. sessillis A. philoxeroides
          Ecology Dewatered soil Very moist soil Wetlands borders
          Growth habit of the plant Creeping erect erect
          Inflorescence (Insertion) Sessile Sessile Stalked
          Inflorescence (appearance) Spiny Non-Spiny Non-spiny
          Inflorescene(size) 10 – 15 mm 5 – 7 mm 10 – 15 mm
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            Ecology

            Alternanthera sessilis prefers moist soils, along ditches, fallow land. The plant grows in moist soils of the lakes, swamps, irrigation canals and rice paddies dams and navigation channels, up to 1200 m altitude.

            Central Africa: Alternanthera sessilis is a widely naturalized exotic species, cosmopolitan and very common, sometimes aquatic or palustrine (riverbanks, ponds, ditches, Echinochloa marshes), sometimes terrestrial on banks and sandy islets (exundated river islets) or anthropophilous: roadsides, villages, crops, rubble.
            China
            : Alternanthera sessilis grows on roadsides, in gardens and swamps.
            Comoros: This species occurs in ruderal areas, along roadsides, in settlements up to the forest of average altitude.
            French Guiana: Species rather present in vegetable zone at the edge of the fields, it develops well in wetlands and in places rich in nitrogen. It is frequent in the ruderal vegetation of wetlands.
            Madagascar: This species prefers hydromorphic and relatively rich moist alluvial soils in conditions more or less shaded, in all climatic zones of Madagascar: edges of canals, marshes and roads. It is also a weed of vegetable crops, taro and rice lowland and upland rice on moist soil, in semi-intensive culture system with organic manure.
            Mauritius: It occupies very often, but not exclusively, humid sites, at the edge of streams, lakes and swamp especially under moist and wet areas.
            Mayotte: Rather common species in the wetlands and on the hydromorphic grounds, especially in the back mangroves, but also in degraded environments.
            Nicaragua: Alternanthera sessilis is a very common species, in disturbed sites, in all parts of the country; from 0 to 750 m altitude.
            Thailand: Weed on marshy land, also along paths, around villages, etc.; up to 1000 m altitude or more.
            West Indies: Alternanthera sessilis is a ruderal species that grows on moist soil, from 0 to 300 m (500m) altitude.

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              📚 Habitat and Distribution
              Description

              Origin

              Alternanthera sessilis is native to Central America, the Caribbean, South America and Asia-Oceania (from the Arabian Peninsula to Australia)

              Worldwide distribution

              A pantropical species widespread in America, sub-Saharan Africa, the Middle East, India, Pakistan, China, Southeast Asia, Indonesia, the Pacific Islands and the Indian Ocean. It is also present in Europe (Spain).

              dummy
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                No Data
                📚 Occurrence
                No Data
                📚 Demography and Conservation
                Risk Statement

                Global harmfulness

                Sometimes noxious.

                Local harmfulness

                Burkina Faso: Alternanthera sessilis is frequent and scarce.
                Comoros: A. sessilis is found in association with other weeds in crops, found moderately common in fallow fields and the old cultures of banana, cassava and vanilliers.
                Côte d'Ivoire: frequent and scarce.
                Ghana: rare and scarce.
                French Guiana: A rarely problematic weed, it does however have vigorous taproots that can complicate manual weeding. It is locally abundant.
                Madagascar: Weed common crops in low-lying areas but generally little abundant and associated with other species creeping or not. This rampant species can form a dense population difficult to remove if the weeding is too late. The very moist soil in which it grows makes it difficult to manage with small hand tools. A. sessilis can be troublesome in vegetable crops invading plots from poorly cleaned borders.
                Mauritius: It can become a significant weed in some vegetable crops and rice paddies with a low average aggressive. On the contrary, it is quite rare in sugar cane where it is weakly aggressive.
                Nigeria: rare and scarce.
                Senegal: rare but abundant when it is present.
                Chad: rare and scarce.
                Tanzania: Frequent but not abundant.

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                  📚 Uses and Management
                  Uses

                  Food: The young stems and leaves of Alternanthera sessilis are eaten as a green vegetable.
                  Medicinal: An infusion of the whole plant is used in Indonesia as a remedy against intestinal pain, diarrhea, dysentery, and externally to treat wounds or against inflammation and to decrease fever. A. sessilis is used in local medicine in Taiwan, often mixed with other medicinal plants, to treat hepatitis, chest angina, bronchitis, asthma and lung problems, to stop  bleeding and as capillary stimulating agent. It is also used locally in India against dysentery, as cholagogue, abortive and to treat snake bites, burns and boils and in Thailand and Sri Lanka, as galactogen. Traditional uses as anti-inflammatory and to treat wounds are proven by the anti-inflammatory and anti-hyperglycemic activities of the extracts of aerial parts of the plant. In Comoros it is used externally against skin diseases.

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                    Management
                    Global control

                    For advice on weed control of irrigated and lowland rice in Africa, visit :

                    For advice on weeding perennial broadleaf weed of irrigated and lowland rice in Africa, please visit

                    For advice on weeding perennial broadleaf weed of irrigated and lowland rice in Africa, please visit

                    Biological control: Zettler and Freeman (1972) in their discussion on the biocontrol of aquatic plants, state that in India, Alternanthera sessilis a near relative of alligator weed, is affected by the fungial species Corticium solani, Colletotrichum capsici, Glomerella cingulata, Phoma spp., and Albugo bliti.
                    Chemical: Amitrole is very effective. Reasonably effective are 2,4-D for broadleaved weeds (1-1.5 kg a.i./ha in 200 liters of water) and MCPA (0.5-1 kg a.i./ha in 200 liters of water), but repeated applications are necessary.

                    Local control

                    Comoros: Alternanthera sessilis is a weed easy to pull off, weeding is done manually.
                    Madagascar: Control can be achieved by good soil preparation and manual grubbing or manual tillage with angady for taro crops in the uplands of the Highlands. In cropping systems based on rainfed rice, vegetable crops and taro, the plots are generally very small and the very different crops are side by side making it difficult to use post-emergence herbicides such as 2 , 4-D.
                    Mauritius: It is effectively controlled by post-emergent herbicide mixtures recommended in sugar cane. It must be removed manually or by weeding in other crops.

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                      No Data
                      📚 Information Listing
                      References
                      1. Grard, P., Homsombath, K., Kessler, P., Khuon, E., Le Bourgeois, T., Prospéri, J., Risdale, C. 2006. Oswald V.1.0: A multimedia identification system of the major weeds of rice paddy fields of Cambodia and Lao P.D.R. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom. ISBN 978-2-87614-653-2.
                      1. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                      1. Holm, L., Doll, J., Holm, E., Pancho, J., Herberger, J. 1997. World Weeds : Natural Histories and Distribution. John Wiley & Sons, Inc., New York, USA.
                      1. Holm, L.G., Plucknett, D., L., Pancho, J.V., Herberger, J.P. 1977. The World's Worst Weeds. University Press of Hawaii, Honolulu, Hawaii.
                      1. Johnson, D.E. 1997. Les adventices en riziculture en Afrique de l'Ouest. ADRAO/WARDA, Bouaké, Côte-d'Ivoire.
                      1. Pancho, J.V., Obien, S.R. 1995. Manual of Ricefield Weeds in the Philippines. Philippine Rice Research Institute, Munoz, Nueva Ecija, Philippines.
                      1. Radanachaless, T., Maxwell, J.F. 1994. Weeds of soybean fields in Thailand. Multiple Cropping Center ed., Chiang Mai Univ., Chiang Mai, Thailand.
                      1. Soerjani, M., Kostermans, A.J.G.H., Tjitrosemito, G. 1987. Weeds of rice in Indonesia. Balai Puskata, Jakarta, Indonesia.
                      1. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Cirad, Gondwana éditions, Montpellier, France.
                      2. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                      1. Bosser, J., Fergusson, I.K., Soopramanien, C. Mult. an. Flore des Mascareignes. La Réunion, Maurice, Rodrigues. MSIRI, IRD, Kew.
                      2. Marnotte, P. and A. Carrara. (2007). "Plantes des rizières de Guyane." from http://plantes-rizieres-guyane.cirad.fr/. http://plantes-rizieres-guyane.cirad.fr/dicotyledones/amaranthaceae/alternanthera_sessilis
                      3. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186p. https://portal.wiktrop.org/document/show/173
                      4. Plants of the World Online https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:1069792-2
                      5. Flora of China http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200006977
                      6. Plants of the World Online http://www.worldfloraonline.org/taxon/wfo-0000528825
                      1. Beleu National Museum (Republic of Palau. Koror.). 1968. Voucher specimen #BNM574(Blackburn 231).
                      2. Anon. 1988. Weed list for American Samoa by cropping system. Checklist dated 07/13/88, origin unknown. 3 pp.
                      1. CAVACO A. 1954. Flore de Madagascar et des Comores, 67è Famille AMARANTHACEES 56 pages
                      2. A.J.G.H., SUBHASHINI T, KRISHNAVENI B AND SRINIVAS REDDY C. 2010. Anti- Inflammatory Activity of Leaf Extracts of Alternanthera sessilis HYGEIA. J .D.MED Vol.2, No.1,54-56 March-Aug, 2010.
                      3. JALALPURE SUNIL S., AGRAWAL N., PATIL MB., CHIMKODE R. & TRIPATHI A. 2008. Antimicrobial and wound healing activities of leaves of Alternanthera sessilis L. International Journal of Green Pharmacy Vol. 2, Issue 3, page 141-144.
                      Information Listing > References
                      1. ISSG http://www.issg.org/database/species/ecology.asp?si=767&fr=1&sts=sss&lang=EN
                      2. Grard, P., Homsombath, K., Kessler, P., Khuon, E., Le Bourgeois, T., Prospéri, J., Risdale, C. 2006. Oswald V.1.0: A multimedia identification system of the major weeds of rice paddy fields of Cambodia and Lao P.D.R. In Cirad [ed.]. Cirad, Montpellier, France. Cdrom. ISBN 978-2-87614-653-2.
                      3. Grard, P., T. Le Bourgeois, J. Rodenburg, P. Marnotte, A. Carrara, R. Irakiza, D. Makokha, G. kyalo, K. Aloys, K. Iswaria, N. Nguyen and G. Tzelepoglou (2012). AFROweeds V.1.0: African weeds of rice. Cédérom. Montpellier, France & Cotonou, Bénin, Cirad-AfricaRice eds.
                      4. Holm, L., Doll, J., Holm, E., Pancho, J., Herberger, J. 1997. World Weeds : Natural Histories and Distribution. John Wiley & Sons, Inc., New York, USA.
                      5. Holm, L.G., Plucknett, D., L., Pancho, J.V., Herberger, J.P. 1977. The World's Worst Weeds. University Press of Hawaii, Honolulu, Hawaii.
                      6. Johnson, D.E. 1997. Les adventices en riziculture en Afrique de l'Ouest. ADRAO/WARDA, Bouaké, Côte-d'Ivoire.
                      7. Pancho, J.V., Obien, S.R. 1995. Manual of Ricefield Weeds in the Philippines. Philippine Rice Research Institute, Munoz, Nueva Ecija, Philippines.
                      8. Radanachaless, T., Maxwell, J.F. 1994. Weeds of soybean fields in Thailand. Multiple Cropping Center ed., Chiang Mai Univ., Chiang Mai, Thailand.
                      9. Soerjani, M., Kostermans, A.J.G.H., Tjitrosemito, G. 1987. Weeds of rice in Indonesia. Balai Puskata, Jakarta, Indonesia.
                      10. Fournet, J. 2002. Flore illustrée des phanérogames de Guadeloupe et de Martinique. Cirad, Gondwana éditions, Montpellier, France.
                      11. Barthelat, F. 2019. La Flore illustrée de Mayotte. Meze, Paris, France, Collection Inventaires et Biodiversité, Biotope – Muséum national d’Histoire naturelle. 487 p.
                      12. Flora of Zimbabwe http://www.zimbabweflora.co.zw/speciesdata/species.php?species_id=122550
                      13. HEAR http://www.hear.org/pier/species/alternanthera_sessilis.htm
                      14. SNTC http://www.sntc.org.sz/alienplants/speciesinfo.asp?spid=40
                      15. Bosser, J., Fergusson, I.K., Soopramanien, C. Mult. an. Flore des Mascareignes. La Réunion, Maurice, Rodrigues. MSIRI, IRD, Kew.
                      16. Marnotte, P. and A. Carrara. (2007). "Plantes des rizières de Guyane." from http://plantes-rizieres-guyane.cirad.fr/. http://plantes-rizieres-guyane.cirad.fr/dicotyledones/amaranthaceae/alternanthera_sessilis
                      17. Berton, A. (2020). Flore spontanée des cultures maraichères et fruitières de Guyane. Guide de reconnaissance des 140 adventices les plus communes des parcelles cultivées. Cayenne, Guyane, FREDON Guyane: 186p. https://portal.wiktrop.org/document/show/173
                      18. Plants of the World Online https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:1069792-2
                      19. Flora of China http://www.efloras.org/florataxon.aspx?flora_id=2&taxon_id=200006977
                      20. Plants of the World Online http://www.worldfloraonline.org/taxon/wfo-0000528825
                      21. Beleu National Museum (Republic of Palau. Koror.). 1968. Voucher specimen #BNM574(Blackburn 231).
                      22. Anon. 1988. Weed list for American Samoa by cropping system. Checklist dated 07/13/88, origin unknown. 3 pp.
                      23. CAVACO A. 1954. Flore de Madagascar et des Comores, 67è Famille AMARANTHACEES 56 pages
                      24. A.J.G.H., SUBHASHINI T, KRISHNAVENI B AND SRINIVAS REDDY C. 2010. Anti- Inflammatory Activity of Leaf Extracts of Alternanthera sessilis HYGEIA. J .D.MED Vol.2, No.1,54-56 March-Aug, 2010.
                      25. JALALPURE SUNIL S., AGRAWAL N., PATIL MB., CHIMKODE R. & TRIPATHI A. 2008. Antimicrobial and wound healing activities of leaves of Alternanthera sessilis L. International Journal of Green Pharmacy Vol. 2, Issue 3, page 141-144.

                      Effet de l’époque de coupe de la canne à sucre sur l’enherbement et le cycle biologique des adventices - A La Réunion -

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                        WIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areasWIKTROP - Weed Identification and Knowledge in the Tropical and Mediterranean areas
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