Checklist |
Corresponding author: Lars Söderström ( lars.soderstrom@ntnu.no ) Academic editor: Lyubomir Penev
© 2016 Lars Söderström, Anders Hagborg, Matt von Konrat, Sharon Bartholomew-Began, David Bell, Laura Briscoe, Elizabeth Brown, D. Christine Cargill, Denise Pinheiro da Costa, Barbara J. Crandall-Stotler, Endymion Cooper, Gregorio Dauphin, John Engel, Kathrin Feldberg, David Glenny, S. Robbert Gradstein, Xiaolan He, Jochen Heinrichs, Joern Hentschel, Anna Luiza Ilkiu-Borges, Tomoyuki Katagiri, Nadezhda A. Konstantinova, Juan Larraín, David Long, Martin Nebel, Tamás Pócs, Felisa Puche, Elena Reiner-Drehwald, Matt Renner, Andrea Sass-Gyarmati, Alfons Schäfer-Verwimp, José Segarra-Moragues, Raymond E. Stotler, Phiangphak Sukkharak, Barbara Thiers, Jaime Uribe, Jiří Váňa, Juan Villarreal, Martin Wigginton, Li Zhang, Rui-Liang Zhu.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Söderström L, Hagborg A, von Konrat M, Bartholomew-Began S, Bell D, Briscoe L, Brown E, Cargill DC, Costa DP, Crandall-Stotler BJ, Cooper ED, Dauphin G, Engel JJ, Feldberg K, Glenny D, Gradstein SR, He X, Heinrichs J, Hentschel J, Ilkiu-Borges AL, Katagiri T, Konstantinova NA, Larraín J, Long DG, Nebel M, Pócs T, Felisa Puche F, Reiner-Drehwald E, Renner MAM, Sass-Gyarmati A, Schäfer-Verwimp A, Moragues JGS, Stotler RE, Sukkharak P, Thiers BM, Uribe J, Váňa J, Villarreal JC, Wigginton M, Zhang L, Zhu R-L (2016) World checklist of hornworts and liverworts. PhytoKeys 59: 1-821. https://doi.org/10.3897/phytokeys.59.6261
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A working checklist of accepted taxa worldwide is vital in achieving the goal of developing an online flora of all known plants by 2020 as part of the Global Strategy for Plant Conservation. We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 7486 species in 398 genera representing 92 families from the two phyla. The checklist has far reaching implications and applications, including providing a valuable tool for taxonomists and systematists, analyzing phytogeographic and diversity patterns, aiding in the assessment of floristic and taxonomic knowledge, and identifying geographical gaps in our understanding of the global liverwort and hornwort flora. The checklist is derived from a working data set centralizing nomenclature, taxonomy and geography on a global scale. Prior to this effort a lack of centralization has been a major impediment for the study and analysis of species richness, conservation and systematic research at both regional and global scales. The success of this checklist, initiated in 2008, has been underpinned by its community approach involving taxonomic specialists working towards a consensus on taxonomy, nomenclature and distribution.
Marchantiophyta , Anthocerophyta , nomenclature, taxonomy
This checklist is dedicated to Elizabeth Brown and Ray Stotler, who sadly left us during our work with it.
“……Charles Darwin wrote of his desire to provide financial support ‘for the formation of a perfect M.S. catalogue of all known plants’ (Darwin 1881, letter 13570). It is a personal embarrassment to me, and should be chastening to us all, that more than 120 years later we still have not delivered on that commitment.” (
The natural world is changing fast (
Version 1.0 of The Plant List (http://www.theplantlist.org/) was released in December 2010 aimed to be comprehensive for species of vascular plants (flowering plants, conifers, ferns and their allies) and of bryophytes (mosses, liverworts and hornworts), as a response to the 2010 Target 1 of the GSPC and to a clear global need for such data. The Plant List is a broad collaboration, coordinated by the Royal Botanic Gardens, Kew, and the Missouri Botanical Garden (MO), involving diverse partnerships.
Earlier,
Liverworts and hornworts (Figure
Habit images of selected lineages of Anthocerotophyta (hornworts) and Marchantiophyta (liverworts). APhaeomegaceros squamuliger, photo by J. Hollinger BHaplomitrium mnioides, photo by L. Zhang CConocephalum japonicum, photo by D. G. Long DColura vitiensis, photo by T. Pócs EPleurozia gigantea, photo by L. Söderström FRiccardia spegazziniana, photo by Juan Larraín.
Liverworts, hornworts and mosses offer microhabitats that are critical to the survival of a tremendous diversity of organisms such as single-celled eukaryotes, protozoa, and numerous groups of invertebrates (
There are estimated to exist only 215 hornwort species. By comparison estimates of liverwort species richness have varied considerably, by as much as 50%, with estimates ranging from 4,500 to 9,000 (e.g.
Below we provide context to the worldwide checklist, including a brief historical account and current informational resources, structure and layout, systematic and classification concepts, and methodology, including a brief overview of the underlying data set and nomenclatural elements.
Figure
Data availability and information needs associated with liverworts and hornworts have been reviewed extensively by
A major nomenclatural work is the Index Hepaticarum (http://www.ville-ge.ch/musinfo/bd/cjb/hepatic/index.php), which includes all effectively published liverwort epithets spanning 12 volumes with the closing date of 1973. The indices were prepared as a purely nomenclatural resource and did not claim to express any particular taxonomic concept. Recently,
The application of molecular phylogenetics continues to generate new insights into the evolutionary history of liverworts and hornworts. In recent years, inferences made from these phylogenies have especially revolutionized liverwort classification (
The systematic treatment as outlined here also includes the addition of subfamily and infrageneric classification; ranks not presented by
The checklist is presented in a taxonomic framework as outlined above with four main sections. The first is arranged systematically with taxa ordered alphabetically within the nearest higher rank. A brief discussion for selected families is provided in the main body of the text under the respective families. This does not appear uniformly for all families, but typically for larger more complicated families. Rather, a brief description is provided mainly where concepts depart from
*** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010 (
There are comprehensive footnotes throughout the systematic section of the checklist. These are applied to selected names representing the rank of genus and below only. Footnotes without any literature citation represent observations from the authors of the respective family. Footnotes are wide ranging in their content. Generally, footnotes can be categorized into the following: i) general statements relating to taxonomic opinion, ii) comments on the type specimen, iii) comments about possible conspecificity, including citation of conflicting opinions, and iv) comment about species complexes.
This section is followed by a list of taxa in genera that we do not recognize. Those names represent taxa that have not been studied recently and have not been recognized in any recent treatment. However, many of those names are old and may gain priority over some recognized taxa once their identity is determined.
The next section is an alphabetical list of taxa of the ranks of species and below. The alphabetical list includes the confidence level, the taxon name and authority as well as a reference to the page of the corresponding entry in the systematic section. Thus, the alphabetical list provides a rapid gateway requiring no prior knowledge of higher taxonomy, but also serves as an index to the corresponding name in the systematic section that includes more detailed information.
The last section is a reference list where full bibliographic citations are given to all references for taxa included in the checklist. Titles and references have all been checked and verified, except in a few cases where we could not get hold of the publications.
Significantly, each accepted taxon is qualified using a three level ranking system that summarizes our knowledge about a taxon. The coding convention we are adopting largely follows that described by
The coding convention:
* Serious doubts. There are doubts about the value of the taxon. It can be that there are conflicting views without any substantial evidence in any direction, conflicting views with substantiating evidence supporting one or both positions, or evidence pointing towards synonymization but it is premature to do it. We have adhered to the principle that it is better to keep a taxon with one star (and preferably a note) than to synonymize it too quickly. Example: Bazzania asymmetrica is conspecific with Bazzania macgregorii in
** Knowledge problem. The taxon is not well known by the person evaluating it. It may be a newly described species or a species originally not well described and not restudied recently. Example: Jungermannia erectii Ajit P.Singh et V.Nath was recently described and has not been independently studied by someone with a global overview. It is therefore difficult to evaluate.
*** Accepted. A good taxon as currently understood based on personal experience or on taxonomic revisions that have been convincingly performed. Nomenclature and/or taxonomic position may, however, be questioned. Elements may be excluded from the taxon, but the taxon with the current type will still be accepted. Example: H. Bischler-Causse revised the genus Marchantia worldwide in a series of publications. Although she had a broad species concept, she also recognized infraspecific taxa. The taxa that she recognized without doubt should be accepted unless new evidence against it exists. Adopting a narrower species concept so that many of her subspecies are elevated to species does not change her view of what a good taxon is.
The foundation of the checklist is in the underlying data set from which it is derived. It was briefly described by
There has been an intense systematic effort focusing on data quality. In all but a few cases nomenclatural data has been verified against original publications. Söderström and Hagborg have checked and confirmed almost every original publication (three publications have not been available) for correct author, title and journal/book citation, date of publication as far as possible, page number for the protologue, and if the name is validly and legitimately published according to the International Code of Nomenclature for algae, fungi and plants (Melbourne Code;
The notes, published in the journal Phytotaxa, have included 38 authors from 13 countries. These are the results of several years of revisions of taxonomic groups by taxonomic experts. Following these revisions and working with participants identifying nomenclatural and taxonomic problems that would impact the worldwide checklist, Söderström, Hagborg and von Konrat led editorial efforts in compiling manuscripts with baseline information and nomenclatural or taxonomic issues that required resolving. The work was then coordinated with the taxonomic experts who drove and led the process and often identified further issues. The series of notes on taxonomy and nomenclature of liverworts and hornworts provided updated nomenclature, corrected invalid and illegitimate names and described new taxa based on studies (mainly molecular) that did not draw nomenclatural conclusions. Those notes are all open access and any changes were effective immediately as the Melbourne Code of Botanical Nomenclature allows publishing on internet from 1 Jan. 2012 (
This checklist builds on all published taxonomic and nomenclatural papers, that have come to our attention until June 30, 2015.
We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 8,078 taxa (species and below) in 7,486 species representing 398 genera, 92 families, 20 orders and 7 classes from the two phyla. The list includes 3,533 species with three stars, 2,988 species with two stars and 915 species with one star. The checklist also has extra utility in that it contains 3,106 references in the bibliography that serve as a powerful bibliographic resource for liverwort and hornwort systematic and taxonomic research.
The marked-up publication form of the current checklist by PhytoKeys provides a virtual instrument with a linked environment both internally (e.g. within an article) and externally (GBIF, IPNI, Tropicos, Wikispecies, etc.) that will undoubtedly help accelerate taxonomic research. The published checklist was the first phase in providing a worldwide list of accepted names. The next phase is to establish a generally recognized online repository to augment the huge underlying informational auxiliary data of over 25,000 publications, almost 39,000 published names, and the over 700,000 geographical observations. Several features of Web-based technology make it an essential tool, including 1) a vehicle to facilitate data access, 2) to unify the vastly scattered data on distributional information and nomenclature, 3) offering dynamic rather than static data in a searchable forum. The broader accessibility to the wealth of auxiliary data will help augment monographic and revisional work for many taxonomic groups, aid in identifying the need for increased floristic and survey work in many regions throughout the world, and have broad implications and applications beyond taxonomic research such as conservation science. The current Early Land Plants Today model has strong participation from taxonomic experts and an online resource will provide an opportunity to expand stakeholders to include ecologists, conservationists, scientists from other disciplines and general interest groups. However, such an effort can only be successful if it comes with sustained funding and infrastructure rather than depending on an ad hoc commitment by a few individuals, however dedicated.
The project will help augment monographic and revisional work for many taxonomic groups and aid in identifying the need for increased floristic and survey work in many regions throughout the world. Although there are many challenges ahead to obtain high quality data, quantifying global liverwort diversity is a tractable, multi-faceted and scientifically important goal, and everyone stands to gain by fostering this endeavour. The success of the project will lie on strong collaboration between institutions and the bryological community in general.
by J.C. Villarreal and D.C. Cargill
Notes on nomenclature and taxonomy can also be found in
*** Anthoceros L., Sp. Pl. 1: 1139, 1753 (
*** Anthoceros adscendens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 24, 1832 (
*** Anthoceros agrestis Paton, J. Bryol. 10 (3): 257, 1979 (
*** Anthoceros alpinus Steph., Sp. Hepat. (Stephani) 6: 425, 1923 (
* Anthoceros angustifolius Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 582, 1846 (
*** Anthoceros angustus Steph., Sp. Hepat. (Stephani) 5: 1001, 1916 (
*** Anthoceros bharadwajii Udar et A.K.Asthana, Proc. Indian Natl. Sci. Acad., B 51 (4): 484, 1985 (
** Anthoceros buettneri Steph., Sp. Hepat. (Stephani) 5: 997, 1916 (
*** Anthoceros capricornii Cargill et G.A.M.Scott, J. Hattori Bot. Lab. 82: 55, 1997 (
*** Anthoceros caucasicus Steph., Izv. Kavkazsk. Muz. 8: 87, 1914 (
** Anthoceros cavernosus Steph., Sp. Hepat. (Stephani) 5: 998, 1916 (
* Anthoceros chambensis Kashyap, J. Bombay Nat. Hist. Soc. 25 (2): 281, 1917 (
** Anthoceros chungii Khanna, J. Indian Bot. Soc. 17 (5/6): 316, 1938 (
** Anthoceros crispatus Griff., Not. pl. asiat. 2: 349, 1849 (
** Anthoceros dimorphus Sim, Trans. Roy. Soc. South Africa 15 (1): 114, 1926 (
*** Anthoceros erectus Kashyap, New Phytol. 14 (1): 9, 1915 (
** Anthoceros expansus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 92, 2015 (
* Anthoceros ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 5: 1007, 1916 (
*** Anthoceros fragilis Steph., Sp. Hepat. (Stephani) 5: 1006, 1916 (
*** Anthoceros fusiformis Austin, Bull. Torrey Bot. Club 6 (4): 28, 1875 [1876] (
*** Anthoceros fusiformis var. taiwanensis J.Haseg., Acta Phytotax. Geobot. 44 (2): 100, 1993 (
** Anthoceros gasongorii Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916 (
** Anthoceros granulatus Gottsche, Mexik. Leverm.: 275, 1863 (
** Anthoceros harrisanus (Steph.) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 31, 1962 (
* Anthoceros helmsii Steph., Hedwigia 32 (3): 142, 1893 (
** Anthoceros jamesonii Taylor ex Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 171, 1855 (
* Anthoceros javanicoides H.A.Mill., Phytologia 47 (4): 319, 1981 (
** Anthoceros jungermannioides Schwein., Spec. Fl. Amer. Crypt.: 25, 1821 (
* Anthoceros kajumas (K.I.Goebel) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (
* Anthoceros koshii Khanna, J. Indian Bot. Soc. 15 (4): 237, 1936 (
*** Anthoceros lamellatus Steph., Sp. Hepat. (Stephani) 5: 1000, 1916 (
*** Anthoceros laminifer Steph., J. Linn. Soc., Bot. 29 (201): 266, 1892 (
*** Anthoceros macounii M.Howe, Bull. Torrey Bot. Club 25 (1): 19, 1898 (
*** Anthoceros macrosporus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (
** Anthoceros maritimus Steph., Sp. Hepat. (Stephani) 5: 984, 1916 (
* Anthoceros megasporus Meijer, J. Hattori Bot. Lab. 18: 6, 1957 (
** Anthoceros muscoides Colenso, Trans. & Proc. New Zealand Inst. 16: 361, 1884 (
*** Anthoceros myriandroecius Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 134, 1911 (
** Anthoceros natalensis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913 (
*** Anthoceros neesii Prosk., Leberm. Eur. 2 (9): 1312, 1958 (
** Anthoceros niger Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (
** Anthoceros orizabensis (Steph.) Hässel, Candollea 45 (1): 211, 1990 (
*** Anthoceros pandei Udar et A.K.Asthana, J. Indian Bot. Soc. 64: 305, 1985 (
*** Anthoceros patagonicus Hässel, Candollea 45 (1): 207, 1990 (
*** Anthoceros patagonicus subsp. gremmenii J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 32, 2013 (
** Anthoceros peruvianus Steph., Sp. Hepat. (Stephani) 5: 999, 1916 (
** Anthoceros pinnatus Steph., Bol. Soc. Brot. 4: 154 [182], 1885 [1886] (
*** Anthoceros punctatus L., Sp. Pl. 1: 1139, 1753 (
* Anthoceros pusillus Colenso, Trans. & Proc. New Zealand Inst. 18: 255, 1886 (
*** Anthoceros rosulans J.Haseg., J. Hattori Bot. Lab. 60: 379, 1986 (
*** Anthoceros sambesianus Steph., Sp. Hepat. (Stephani) 5: 996, 1916 (
*** Anthoceros scariosus Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 230, 1869 (
** Anthoceros schroederi Steph., 52 (5): 306, 1912 (
** Anthoceros serratus Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 90, 1911 (
** Anthoceros simulans M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 204, 1934 (
** Anthoceros spongiosus Steph., Sp. Hepat. (Stephani) 5: 1003, 1916 (
*** Anthoceros subtilis Steph., Sp. Hepat. (Stephani) 5: 1003, 1916 (
*** Anthoceros telaganus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (
*** Anthoceros tristanianus J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 33, 2013 (
*** Anthoceros tuberculatus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 25, 1832 (
** Anthoceros venosus Lindenb. et Gottsche, Syn. Hepat. 4: 584, 1846 (
Excluded from the genus
* Anthoceros aethyopicus Gola, Ann. Bot. (Rome) 13 (1): 73, 1914 (
* Anthoceros brunnthaleri Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913 (
* Anthoceros floribundus Steph., Sp. Hepat. (Stephani) 5: 977, 1916 (
* Anthoceros mildbraedii Steph., Sp. Hepat. (Stephani) 6: 428, 1923 (
* Anthoceros parvifrons Steph., 52 (5): 307, 1912 (
* Anthoceros pseudocostus Steph., Sp. Hepat. (Stephani) 5: 977, 1916 (
* Anthoceros rossoi Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916 (
*** Folioceros D.C.Bharadwaj, Geophytology 1 (1): 9, 1971 (
*** Folioceros amboinensis (Schiffn.) Piippo, Acta Bot. Fenn. 148: 36, 1993 (
** Folioceros apiahynus (Steph.) Hässel, Candollea 45 (1): 215, 1990 (
** Folioceros argillaceus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015 (
*** Folioceros assamicus D.C.Bharadwaj, Geophytology 1 (1): 10, 1971 (
** Folioceros dilatatus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015 (
*** Folioceros dixitianus (Mahab.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros fuciformis (Mont.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros glandulosus (Lehm. et Lindenb.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros incurvus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros indicus D.C.Bharadwaj, Geophytology 8 (1): 114, 1978 (
*** Folioceros kashyapii S.C.Srivast. et A.K.Asthana, Bryologist 92 (2): 219, 1989 (
*** Folioceros mangaloreus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros paliformis D.K.Singh, Bull. Bot. Surv. India 29: 176, 1987 [1989] (
*** Folioceros physocladus D.C.Bharadwaj, Geophytology 8 (1): 115, 1978 (
*** Folioceros pinnilobus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros satpurensis D.C.Bharadwaj et K.P.Srivast., Geophytology 8 (1): 112, 1978 (
*** Folioceros udarii A.K.Asthana et S.C.Srivast., Cryptog. Bryol. Lichénol. 7 (2): 151, 1986 (
*** Folioceros verruculosus (J.Haseg.) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 383, 2011 (
by J.C. Villarreal and D.C. Cargill
*** Dendroceros Nees, Syn. Hepat. 4: 579, 1846 (
** subg. Apoceros R.M.Schust., Phytologia 63 (3): 200, 1987 (
*** Dendroceros cavernosus J.Haseg., J. Hattori Bot. Lab. 47: 306, 1980 (
*** Dendroceros difficilis Steph., Sp. Hepat. (Stephani) 5: 1009, 1917 (
*** Dendroceros muelleri Steph., Hedwigia 28 (2): 133, 1889 (
*** Dendroceros ogeramnangus Piippo, Acta Bot. Fenn. 148: 40, 1993 (
*** Dendroceros pedunculatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909 (
** Dendroceros subdifficilis S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 119, 1951 (
** subg. Dendroceros
*** Dendroceros acutilobus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909 (
*** Dendroceros borbonicus Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 31, 1893 [1894] (
*** Dendroceros crispus (Sw.) Nees, Syn. Hepat. 4: 581, 1846 (
*** Dendroceros foliicola J.Haseg., J. Hattori Bot. Lab. 47: 296, 1980 (
*** Dendroceros japonicus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 15, 1909 (
*** Dendroceros javanicus (Nees) Nees, Syn. Hepat. 4: 582, 1846 (
*** Dendroceros subplanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 20, 1909 (
*** Dendroceros tubercularis S.Hatt., Bot. Mag. (Tokyo) 58 (685): 6, 1944 (
*** Dendroceros validus Steph., Sp. Hepat. (Stephani) 5: 1016, 1917 (
** Dendroceros vesconianus Gottsche, J. Bot. (Morot) 12: 150, 1898 (
** Dendroceros wattsianus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909 (
Incertae sedis
* Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 580, 1846 (
*** Dendroceros africanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909 (
** Dendroceros allionii Steph., Sp. Hepat. (Stephani) 5: 1014, 1917 (
** Dendroceros australis Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909 (
** Dendroceros breutelii Nees, Syn. Hepat. 4: 581, 1846 (
** Dendroceros breutelii var. surinamensis Lindenb. et Gottsche, Linnaea 24 (6): 639, 1851 [1852] (
*** Dendroceros cichoraceus (Mont.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 16, 1858 (
* Dendroceros crassicostatus Steph., Sp. Hepat. (Stephani) 5: 1015, 1917 (
** Dendroceros crassinervis (Nees) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 18, 1858 (
*** Dendroceros crispatus (Hook.) Nees, Syn. Hepat. 4: 579, 1846 (
** Dendroceros crispatus var. simplicior Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 574, 1885 (
** Dendroceros cucullatus Steph., Sp. Hepat. (Stephani) 6: 429, 1923 (
* Dendroceros exalatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909 (
* Dendroceros gracilis Steph., Sp. Hepat. (Stephani) 5: 1015, 1917 (
*** Dendroceros granulatus Mitt., Fl. vit.: 419, 1871 [1873] (
** Dendroceros herasii M.Infante, J. Bryol. 32 (4): 285, 2010 (
** Dendroceros humboldtensis Hürl., Bauhinia 1 (3): 254, 1960 (
** Dendroceros paivae C.A.Garcia, Sérgio et J.C.Villarreal, Cryptog. Bryol. 33 (1): 5, 2012 (
* Dendroceros rarus Steph., Sp. Hepat. (Stephani) 5: 1014, 1917 (
* Dendroceros reticulus Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 37, 1950 [1951] (
** Dendroceros rigidus Steph., Sp. Hepat. (Stephani) 5: 1017, 1917 (
*** Dendroceros seramensis J.Haseg., Acta Phytotax. Geobot. 37 (1/3): 10, 1986 (
** Dendroceros subtropicus C.J.Wild, Trans. Nat. Hist. Soc. Queensland 1: 49, 1893 (
** Dendroceros tahitensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 138, 1873 (
*** Megaceros Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907 (
** Megaceros aneuriformis Steph., Sp. Hepat. (Stephani) 5: 949, 1916 (
*** Megaceros austronesophilus Cargill et Seppelt, Austral. Syst. Bot. 26 (5): 372, 2013 (
** Megaceros ciliatus K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 74, 1928 (
*** Megaceros denticulatus (Lehm.) Steph., Sp. Hepat. (Stephani) 5: 956, 1916 (
*** Megaceros flagellaris (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 951, 1916 (
*** Megaceros gracilis (Reichardt) Steph., Sp. Hepat. (Stephani) 5: 955, 1916 (
** Megaceros leptohymenius (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 955, 1916 (
*** Megaceros pellucidus (Colenso) E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 115, 1972 (
** Megaceros tjibodensis Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907 (
Excluded from the genus
* Megaceros flavens (Spruce) Campb., Ann. Bot. (Oxford) 21 (4): 483, 1907 (
* Megaceros jamesonii (Taylor) Steph., Biblioth. Bot. 87 (2): 268, 1916 (
*** Nothoceros (R.M.Schust.) J.Haseg., J. Hattori Bot. Lab. 76: 32, 1994 (
*** Nothoceros aenigmaticus J.C.Villarreal et K.D.McFarland, Bryologist 113 (1): 109, 2010 (
*** Nothoceros canaliculatus (Pagán) J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (
*** Nothoceros endiviifolius (Mont.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (
** Nothoceros fuegiensis (Steph.) J.C.Villarreal, Bryologist 113 (1): 109, 2010 (
*** Nothoceros giganteus (Lehm. et Lindenb.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (
*** Nothoceros minarum (Nees) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014 (
** Nothoceros renzagliensis J.C.Villarreal, L.V.Campos et Uribe, Syst. Bot. 37 (1): 32, 2012 (
*** Nothoceros schizophyllus (Steph.) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014 (
*** Nothoceros superbus J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 280, 2007 (
*** Nothoceros vincentianus (Lehm. et Lindenb.) J.C.Villarreal, Bryologist 113 (1): 111, 2010 (
** Phaeomegaceros R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros coriaceus (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros fimbriatus (Gottsche) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros foveatus (J.Haseg.) J.C.Villarreal, Nova Hedwigia 91 (3/4): 352, 2010 (
*** Phaeomegaceros hirticalyx (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros plicatus (Mitt.) J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 85, 2013 (
*** Phaeomegaceros skottsbergii (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros squamuliger (Spruce) J.C.Villarreal, Nova Hedwigia 91 (3/4): 351, 2010 (
** Phaeomegaceros squamuliger subsp. hasselii J.C.Villarreal, Cargill et Goffinet, Nova Hedwigia 91 (3/4): 352, 2010 (
by J.C. Villarreal and D.C. Cargill
*** Phymatoceros Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 113, 2005 (
*** Phymatoceros bulbiculosus (Brot.) Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 114, 2005 (
*** Phymatoceros phymatodes (M.Howe) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 240, 2007 (
by J.C. Villarreal and D.C. Cargill
*** Notothylas Sull. ex A.Gray, Amer. J. Sci. Arts (ser. 2) 1 (1): 74, 1846 (
** subg. Notothylas
*** Notothylas anaporata Udar et D.K.Singh, Rev. Bryol. Lichénol. 45 (2): 202, 1979 (
*** Notothylas breutelii (Gottsche) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 21, 1858 (
*** Notothylas depressispora J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 26, 1979 (
*** Notothylas dissecta Steph., Sp. Hepat. (Stephani) 5: 1020, 1917 (
*** Notothylas himalayensis Udar et D.K.Singh, J. Bryol. 11 (3): 451, 1981 (
*** Notothylas indica Kashyap, Proc. Lahore Philos. Soc. 4: 54, 1925 (
*** Notothylas javanica (Sande Lac.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 20, 1858 (
*** Notothylas orbicularis (Schwein.) Sull., Amer. J. Sci. Arts (ser. 2) 1 (1): 75, 1846 (
*** Notothylas pandei Udar et V.Chandra, Geophytology 7 (2): 142, 1977 (
** subg. Notothyloides A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 106, 1991 (
*** Notothylas khasiana Udar et D.K.Singh, J. Indian Bot. Soc. 60: 112, 1981 (
*** Notothylas pfleidereri Udar et D.K.Singh, Lindbergia 5 (1): 28, 1979 (
Incertae sedis
*** Notothylas decurva (Mitt.) Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901 (
*** Notothylas flabellata Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901 (
*** Notothylas galapagensis M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 203, 1934 (
** Notothylas irregularis Chantanaorr., Acta Bot. Hung. 56 (3/4): 270, 2014 (
*** Notothylas kashyapii D.K.Singh, Indian J. Forest. 23 (4): 386, 2000 (
*** Notothylas nepalensis D.K.Singh, J. Bombay Nat. Hist. Soc. 84 (3): 650, 1987 [1988] (
*** Notothylas temperata J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 20, 1979 (
*** Notothylas udarii D.K.Singh et Semwal, Phytotaxonomy 1: 35, 2001 (
* Notothylas verdoornii Khanna, Rev. Bryol. Lichénol. 6: 118, 1933 (
*** Notothylas vitalii Udar et D.K.Singh, Misc. Bryol. Lichenol. 8 (9): 173, 1980 (
** Notothylas yunnanensis T.Peng et R.L.Zhu, Phytotaxa 156 (3): 157, 2014 (
* Mesoceros Piippo, Acta Bot. Fenn. 148: 30, 1993 (
* Mesoceros mesophoros Piippo, Acta Bot. Fenn. 148: 30, 1993 (
* Mesoceros porcatus Piippo, Haussknechtia, Beih. 9: 279, 1999 (
** Paraphymatoceros Hässel, Phytologia 88 (2): 208, 2006 (
*** Paraphymatoceros diadematus Hässel, Phytologia 88 (2): 209, 2006 (
*** Paraphymatoceros hallii (Austin) Hässel, Phytologia 88 (2): 209, 2006 (
*** Paraphymatoceros pearsonii (M.Howe) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015 (
*** Paraphymatoceros proskaueri (Stotler, Crand.-Stotl. et W.T.Doyle) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015 (
*** Phaeoceros Prosk., Bull. Torrey Bot. Club 78 (4): 346, 1951 (
** Phaeoceros austroandinus Hässel, Candollea 44 (2): 721, 1989 (
** Phaeoceros bolusii (Sim) S.W.Arnell, Hepat. South Africa: 401, 1963 (
** Phaeoceros brevicapsulus (Steph.) Hässel, Candollea 44 (2): 725, 1989 (
*** Phaeoceros carolinianus (Michx.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (
*** Phaeoceros delicatus E.O.Campb. et Outred, New Zealand J. Bot. 33 (3): 285, 1995 (
*** Phaeoceros dendroceroides (Steph.) Hässel, Beih. Nova Hedwigia 134: 487, 2009 (
*** Phaeoceros engelii Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 248, 2008 (
** Phaeoceros erectus Udar et D.K.Singh, Geophytology 11 (2): 257, 1981 (
*** Phaeoceros evanidus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 245, 2008 (
*** Phaeoceros exiguus (Steph.) J.Haseg., J. Hattori Bot. Lab. 60: 387, 1986 (
*** Phaeoceros flexivalvis (Gottsche et Nees) Hässel, Candollea 44 (2): 728, 1989 (
*** Phaeoceros fulvisporus (Steph.) J.Haseg., Trop. Bryol. 8: 52, 1993 (
*** Phaeoceros gemmifer (Horik.) J.Haseg., J. Hattori Bot. Lab. 57: 252, 1984 (
** Phaeoceros gualaquizanus (Steph.) Gradst., J. Hattori Bot. Lab. 45: 123, 1979 (
*** Phaeoceros himalayensis (Kashyap) Prosk. ex Bapna et G.G.Vyas, J. Hattori Bot. Lab. 25: 88, 1962 (
*** Phaeoceros huebschmannii Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 301, 1986 (
*** Phaeoceros inflatus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 246, 2008 (
*** Phaeoceros kashyapii A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 129, 1991 (
*** Phaeoceros laevis (L.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (
** Phaeoceros maranguensis (Steph.) Bapna, Univ. Udaipur Res. Stud. 3: 138, 1966 (
* Phaeoceros microsporus (Steph.) Hässel, Candollea 44 (2): 721, 1989 (
*** Phaeoceros minutus (Mitt.) S.W.Arnell, Hepat. South Africa: 403, 1963 (
*** Phaeoceros mohrii (Austin) Hässel, Candollea 44 (2): 721, 1989 (
*** Phaeoceros oreganus (Austin) Hässel, Candollea 44 (2): 718, 1989 (
*** Phaeoceros parvulus (Schiffn.) J.Haseg., J. Hattori Bot. Lab. 57: 248, 1984 (
*** Phaeoceros perpusillus Chantanaorr., Acta Bot. Hung. 51 (1/2): 30, 2009 (
** Phaeoceros pichinchensis (Spruce) Hässel, Candollea 44 (2): 725, 1989 (
** Phaeoceros propagulifer (Steph.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (
* Phaeoceros striatisporus J.Haseg., J. Hattori Bot. Lab. 75, 268 (
*** Phaeoceros tenuis (Spruce) Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 303, 1986 (
** Phaeoceros tigrinus (Gola) J.C.Villarreal, Phytotaxa 208 (1): 95, 2015 (
*** Phaeoceros tuberosus (Taylor) Prosk., J. Indian Bot. Soc. 42A: 185, 1964 (
*** Phaeoceros udarii A.K.Asthana et V.Nath, Proc. Natl. Acad. Sci. India, B 63 (4): 461, 1993 (
** Phaeoceros wrightii (Steph.) Hässel, Candollea 44 (2): 722, 1989 (
by J.C. Villarreal and D.C. Cargill
*** Leiosporoceros Hässel, J. Bryol. 14 (2): 255, 1986 (
*** Leiosporoceros dussii (Steph.) Hässel, J. Bryol. 14 (2): 255, 1986 (
by S. Bartholomew-Began
The treatment of Haplomitriaceae follows
*** Haplomitrium Nees, Naturgesch. Eur. Leberm. 1: 109, 1833 (
*** subg. Calobryum (Nees) R.M.Schust., Nova Hedwigia 13 (1/2): 40, 1967 (
*** Haplomitrium blumei (Nees) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (
*** Haplomitrium mnioides (Lindb.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (
*** subg. Haplomitrium
** sect. Archibryum (R.M.Schust.) J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 675, 1981 (
*** Haplomitrium gibbsiae (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (
*** Haplomitrium intermedium Berrie, Proc. Linn. Soc. New South Wales (ser. 2) 87 (399): 191, 1963 (
** sect. Haplomitrium
*** Haplomitrium hookeri (Lyell ex Sm.) Nees, Naturgesch. Eur. Leberm. 1: 111, 1833 (
*** Haplomitrium hookeri var. minutum (E.O.Campb.) Barthol.-Began, Bryophyt. Biblioth. 41: 230, 1991 (
*** Haplomitrium monoicum J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 668, 1981 [1982] (
*** Haplomitrium ovalifolium R.M.Schust., Bryologist 74 (2): 136, 1971 (
by R. Stotler and B.J. Crandall-Stotler
*** Apotreubia S.Hatt. et Mizut., Bryologist 69 (4): 491, 1966 [1967] (
*** Apotreubia hortoniae Konstant., Phytotaxa 76 (3): 33, 2013 (
*** Apotreubia nana (S.Hatt. et Inoue) S.Hatt. et Mizut., Bryologist 69 (4): 492, 1966 [1967] (
** Apotreubia pusilla (R.M.Schust.) Grolle, Acta Bot. Fenn. 125: 63, 1984 (
*** Apotreubia yunnanensis Higuchi, Cryptog. Bryol. Lichénol. 19 (4): 321, 1998 (
*** Treubia K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] (
*** Treubia insignis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] (
** Treubia insignis subsp. bracteata (Steph.) R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 241, 1969 (
** Treubia insignis subsp. caledonica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 243, 1969 (
** Treubia insignis subsp. vitiensis R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 242, 1969 (
*** Treubia lacunosa (Colenso) Prosk., Bryologist 58 (3): 199, 1955 (
** Treubia lacunosoides T.Pfeiff., W.Frey et M.Stech, Nova Hedwigia 75 (1/2): 249, 2002 (
*** Treubia pygmaea R.M.Schust., Phytologia 56 (7): 460, 1985 (
*** Treubia scapanioides R.M.Schust., J. Hattori Bot. Lab. 32: 246, 1969 (
*** Treubia tahitensis (Nadeaud) Besch., J. Bot. (Morot) 12: 147, 1898 (
*** Treubia tasmanica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 248, 1969 (
by K. Feldberg, J. Váňa and J. Heinrichs
The subfamily Jamesonielloideae was excluded from Lophoziaceae/Scapaniaceae by
*** Adelanthoideae K.Feldberg, Heinrichs et Váňa
*** Adelanthus Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 (
** sect. Adelanthus
*** Adelanthus falcatus (Hook.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 (
*** Adelanthus occlusus (Hook.f. et Taylor) Carrington, Trans. Bot. Soc. Edinburgh 10: 381, 1870 (
** sect. Calyptrocolea (R.M.Schust.) Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (
*** Adelanthus aureomarginatus R.M.Schust., Phytologia 39 (4): 250, 1978 (
*** Adelanthus gemmiparus (R.M.Schust.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Biol. Sci. 11 (18): 241, 1970 (
** Adelanthus lingulatus J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 26, 2013 (
*** Adelanthus tenuis J.J.Engel et Grolle, J. Hattori Bot. Lab. 35: 333, 1972 (
** sect. Lindenbergiani Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (
*** Adelanthus carabayensis (Mont.) Grolle, J. Hattori Bot. Lab. 35: 348, 1972 (
*** Adelanthus integerrimus Grolle, J. Hattori Bot. Lab. 35: 340, 1972 (
*** Adelanthus lindenbergianus (Lehm.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 244, 1864 (
** sect. Pittieri Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (
*** Adelanthus pittieri (Steph.) Grolle, J. Hattori Bot. Lab. 35: 337, 1972 (
*** Adelanthus squarrosus Grolle, J. Hattori Bot. Lab. 67: 243, 1989 (
*** Pseudomarsupidium Herzog, Svensk Bot. Tidskr. 47 (1): 42, 1953 (
*** Pseudomarsupidium aureocinctum (R.M.Schust.) J.J.Engel, Novon 17 (3): 312, 2007 (
*** Pseudomarsupidium borneensis (Grolle) Váňa, L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 60, 2012 (
*** Pseudomarsupidium decipiens (Hook.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963 (
*** Pseudomarsupidium piliferum (Steph.) Herzog ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963 (
*** Wettsteinia Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 44, 1898 (
*** Wettsteinia densiretis (Herzog) Grolle, J. Hattori Bot. Lab. 28: 99, 1965 (
*** Wettsteinia inversa (Sande Lac.) Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 45, 1898 (
*** Wettsteinia rotundifolia (Horik.) Grolle, J. Hattori Bot. Lab. 28: 100, 1965 (
*** Wettsteinia schusteriana Grolle, J. Hattori Bot. Lab. 28: 99, 1965 (
** Jamesonielloideae Inoue
*** Cuspidatula Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1141 (124), 1901 (
*** Cuspidatula contracta (Reinw., Blume et Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1141 (124), 1901 (
*** Cuspidatula flaccida (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010 (
*** Cuspidatula flexicaulis (Nees) Váňa et L.Söderstr., Phytotaxa 76 (3): 35, 2013 (
*** Cuspidatula kirkii (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010 (
*** Cuspidatula monodon (Taylor) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1143 (126), 1901 (
*** Cuspidatula orbicularis (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013 (
*** Cuspidatula robusta (Austin) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013 (
** Denotarisia Grolle, Feddes Repert. 82 (1): 6, 1971 (
*** Denotarisia linguifolia (De Not.) Grolle, Feddes Repert. 82 (1): 6, 1971 (
** Nothostrepta R.M.Schust., Phytologia 45 (5): 420, 1980 (
*** Nothostrepta bifida (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980 (
*** Nothostrepta longissima (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980 (
** Pisanoa Hässel, Lindbergia 14 (3): 179, 1988 [1989] (
*** Pisanoa chilensis Hässel, Lindbergia 14 (3): 179, 1988 [1989] (
** Protosyzygiella (Inoue) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980 (
*** Protosyzygiella pseudoconnexa (Herzog) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980 (
*** Syzygiella Spruce, J. Bot. 14: 234, 1876 (
*** subg. Anomalae (Inoue) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella anomala (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 471 (190), 1902 (
*** Syzygiella bilobata Inoue, J. Hattori Bot. Lab. 29: 186, 1966 (
** Syzygiella ciliata Gradst. et A.R.Benitez, Nova Hedwigia 99 (1/2): 115, 2014 (
*** Syzygiella concreta (Gottsche) Spruce, J. Bot. 14: 234, 1876 (
*** Syzygiella manca (Mont.) Steph., Hedwigia 31 (1): 14, 1892 (
*** Syzygiella pectiniformis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 501, 1885 (
*** Syzygiella tonduzana Steph., Sp. Hepat. (Stephani) 6: 118, 1917 (
*** Syzygiella trigonifolia (Steph.) Herzog, Hedwigia 74 (2): 87, 1934 (
*** subg. Cryptochila (R.M.Schust.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella acinacifolia (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella nigrescens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella paludosa (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella pseudocclusa (E.A.Hodgs.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella sonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella spegazziniana (Spruce ex C.Massal.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** subg. Pseudoplagiochila Inoue, J. Hattori Bot. Lab. 29: 182, 1966 (
*** Syzygiella ovalifolia Inoue, J. Hattori Bot. Lab. 29: 191, 1966 (
* Syzygiella securifolia (Nees) Inoue, J. Hattori Bot. Lab. 46: 232, 1979 (
*** Syzygiella subintegerrima (Reinw., Blume et Nees) Spruce, J. Linn. Soc., Bot. 30 (210): 362, 1895 (
*** Syzygiella tasmanica (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** subg. Roivainenia (Perss.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** Syzygiella jacquinotii (Mont.) Hentschel, K.Feldberg, Váňa et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** subg. Syzygiella
*** Syzygiella autumnalis (DC.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** Syzygiella campanulata Herzog, Rev. Bryol. Lichénol. 11 (1): 9, 1938 [1939] (
*** Syzygiella colorata (Lehm.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
** Syzygiella colorata var. collenchymata J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 102, 2013 (
*** Syzygiella contigua Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 470 (189), 1902 (
*** Syzygiella elongella (Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** Syzygiella macrocalyx (Mont.) Spruce, J. Bot. 14: 234, 1876 (
*** Syzygiella nipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (
** Syzygiella oenops (Lindenb. et Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (
*** Syzygiella perfoliata (Sw.) Spruce, J. Bot. 14: 234, 1876 (
*** Syzygiella purpurascens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (
*** Syzygiella rubricaulis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 468 (187), 1902 (
*** Syzygiella setulosa Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 469 (188), 1902 (
*** Syzygiella teres (Carrington et Pearson) Váňa, Phytotaxa 76 (3): 35, 2013 (
*** Syzygiella undata (Mont.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (
Incertae sedis
*** Syzygiella eatonii (Austin) Inoue, J. Jap. Bot. 37 (12): 359, 1962 (
*** Syzygiella uleana Steph., Hedwigia 44 (4): 224, 1905 (
** Vanaea (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989 (
*** Vanaea plagiochiloides (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989 (
by J. Váňa and L. Söderström
Anastrophyllaceae was described by
*** Anastrepta (Lindb.) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (
*** Anastrepta orcadensis (Hook.) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (
*** Anastrophyllum (Spruce) Steph., Hedwigia 32 (3): 139, 1893 (
*** Anastrophyllum alpinum Steph., Sp. Hepat. (Stephani) 6: 103, 1917 (
*** Anastrophyllum assimile (Mitt.) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum auritum (Lehm.) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1137 (120), 1901 (
*** Anastrophyllum ciliatum Steph., Hedwigia 32 (3): 139, 1893 (
*** Anastrophyllum donnianum (Hook.) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum ellipticum Inoue, Bull. Natl. Sci. Mus. Tokyo, B 4 (1): 13, 1978 (
** Anastrophyllum esenbeckii (Mont.) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum fissum Steph., Bull. Herb. Boissier 5 (10): 845, 1897 (
*** Anastrophyllum joergensenii Schiffn., Hedwigia 49 (4): 396, 1910 (
* Anastrophyllum lignicola D.B.Schill et D.G.Long, Ann. Bot. Fenn. 39 (2): 130, 2002 (
*** Anastrophyllum michauxii (F.Weber) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 289, 1932 [1933] (
*** Anastrophyllum nigrescens (Mitt.) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum obtusum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 285, 1950 (
*** Anastrophyllum piligerum (Nees) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum squarrosum Herzog, Ann. Bryol. 5: 72, 1932 (
*** Anastrophyllum stellatum R.M.Schust., Phytologia 39 (4): 243, 1978 (
*** Anastrophyllum tubulosum (Nees) Grolle, J. Hattori Bot. Lab. 28: 101, 1965 (
*** Barbilophozia Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (
** subg. Barbilophozia
*** Barbilophozia barbata (Schmidel ex Schreb.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (
*** Barbilophozia hatcheri (A.Evans) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (
*** Barbilophozia lycopodioides (Wallr.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (
*** Barbilophozia rubescens (R.M.Schust. et Damsh.) Kartt. et L.Söderstr., Ann. Bot. Fenn. 29 (2): 120, 1992 (
** subg. Sudeticae (Schljakov) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Barbilophozia sudetica (Nees ex Huebener) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Biantheridion (Grolle) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Biantheridion undulifolium (Nees) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Chandonanthus Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
*** Chandonanthus squarrosus (Menzies) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
*** Crossocalyx Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939 (
*** Crossocalyx hellerianus (Nees ex Lindenb.) Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939 (
*** Crossocalyx tenuis (Harry Williams) Schljakov, Novosti Sist. Nizš. Rast. 15: 246, 1978 (
*** Gymnocolea (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
*** Gymnocolea borealis (Frisvoll et Moen) R.M.Schust., Lindbergia 12 (1): 7, 1986 (
** Gymnocolea fascinifera Potemkin, Arctoa 2: 76, 1993 (
*** Gymnocolea inflata (Huds.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
* Gymnocolea inflata subsp. acutiloba (Schiffn.) R.M.Schust. et Damsh. ex L.Söderstr. et Váňa, Lindbergia 27 (1): 43, 2002 (
*** Hamatostrepta Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008 (
*** Hamatostrepta concinna Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008 (
** Hattoria R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 69, 1961 (
*** Hattoria yakushimensis (Horik.) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 70, 1961 (
*** Isopaches H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 287, 1932 [1933] (
*** Isopaches alboviridis (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979 (
*** Isopaches bicrenatus (Schmidel ex Hoffm.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933] (
*** Isopaches decolorans (Limpr.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933] (
*** Isopaches pumicicola (Berggr.) Bakalin, Arctoa 17: 162, 2008 [2009] (
*** Neoorthocaulis L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (
*** Neoorthocaulis attenuatus (Mart.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (
*** Neoorthocaulis binsteadii (Kaal.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (
*** Neoorthocaulis floerkei (F.Weber et D.Mohr) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
** Neoorthocaulis hyperboreus (R.M.Schust.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Orthocaulis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 293, 1932 [1933] (
*** Orthocaulis atlanticus (Kaal.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 294, 1932 [1933] (
* Orthocaulis cavifolius H.Buch et S.W.Arnell, Memoranda Soc. Fauna Fl. Fennica 26: 71, 1951 (
** Plicanthus R.M.Schust., Nova Hedwigia 74 (3/4): 484, 2002 (
*** Plicanthus birmensis (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 223, 2002 (
** Plicanthus difficilis (Steph.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 30, 2013 (
** Plicanthus giganteus (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 485, 2002 (
*** Plicanthus hirtellus (F.Weber) R.M.Schust., Nova Hedwigia 74 (3/4): 492, 2002 (
*** Schizophyllopsis Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
** Schizophyllopsis aristata (Herzog ex N.Kitag.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
*** Schizophyllopsis bidens (Reinw., Blume et Nees) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
** Schizophyllopsis lanciloba (Steph.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
*** Schizophyllopsis papillosa (J.J.Engel et Braggins) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
*** Schizophyllopsis sphenoloboides (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 152 (1): 49, 2013 (
*** Schljakovia Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Schljakovia kunzeana (Huebener) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Schljakovianthus Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Schljakovianthus quadrilobus (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Sphenolobopsis R.M.Schust. et N.Kitag., Nova Hedwigia 22: 152, 1971 [1972] (
*** Sphenolobopsis pearsonii (Spruce) R.M.Schust., Nova Hedwigia 22: 153, 1971 [1972] (
*** Sphenolobus (Lindb.) Berggr., New Zealand Hepat.: 22, 1898 (
*** Sphenolobus austroamericanus (Váňa) Váňa, Phytotaxa 81 (1): 30, 2013 (
*** Sphenolobus minutus (Schreb. ex D.Crantz) Berggr., New Zealand Hepat.: 22, 1898 (
*** Sphenolobus saxicola (Schrad.) Steph., Bull. Herb. Boissier (sér. 2) 2 (2): 168 (160), 1902 (
** Tetralophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 13: 227, 1976 (
*** Tetralophozia cavallii (Gola) Váňa, Trop. Bryol. 8: 102, 1993 (
*** Tetralophozia filiformis (Steph.) Urmi, J. Bryol. 12 (3): 394, 1983 (
** Tetralophozia pilifera (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 482, 2002 (
*** Tetralophozia setiformis (Ehrh.) Schljakov, Novosti Sist. Nizš. Rast. 13: 228, 1976 (
*** Zantenia (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013 (
*** Zantenia borneensis (Herzog) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013 (
*** Zantenia denticulata (Grolle) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (
*** Zantenia karstenii (Schiffn.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (
*** Zantenia prionophylla (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (
by J. Váňa with contributions by S.R. Gradstein (Odontoschisma)
Cephaloziaceae was recently studied by
** Alobiella (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (
*** Alobiella husnotii (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (
** Alobiellopsis R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (
*** Alobiellopsis acroscypha (Spruce) R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (
*** Alobiellopsis dominicensis (Spruce) Fulford, Mem. New York Bot. Gard. 11 (3): 350, 1968 (
*** Alobiellopsis heteromorpha (Lehm.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 682, 1969 (
*** Alobiellopsis parvifolia (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 679, 1969 (
*** Alobiellopsis pillansii (Sim) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 683, 1969 (
*** Cephalozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
*** Cephalozia acuminata (Herzog) Váňa, Phytotaxa 112 (1): 11, 2013 (
** Cephalozia acutiloba (Inoue) Váňa, Phytotaxa 112 (1): 11, 2013 (
*** Cephalozia albula Steph., Hedwigia 32 (5): 318, 1893 (
*** Cephalozia ambigua C.Massal., Malpighia 21 (7/8): 310, 1907 (
*** Cephalozia austrigena R.M.Schust. ex J.J.Engel, Novon 17 (3): 312, 2007 (
*** Cephalozia badia (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 483 (313), 1908 (
*** Cephalozia bicuspidata (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
* Cephalozia bicuspidata subsp. lammersiana (Huebener) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 730, 1974 (
*** Cephalozia chilensis (J.J.Engel et R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 29, 2002 (
*** Cephalozia conchata (Grolle et Váňa) Váňa, Syst. Bot. 40 (1): 38, 2015 (
*** Cephalozia crossii Spruce, Cephalozia: 46, 1882 (
*** Cephalozia darjeelingensis Udar et D.Kumar, Geophytology 6 (1): 36, 1976 (
*** Cephalozia drucei (R.M.Schust.) Váňa, Phytotaxa 112 (1): 11, 2013 (
*** Cephalozia fuegiensis Váňa, Phytotaxa 112 (1): 12, 2013 (
*** Cephalozia hamatiloba Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 427 (303), 1908 (
*** Cephalozia hamatiloba subsp. siamensis (N.Kitag.) Váňa, Acta Bot. Fenn. 177: 16, 2004 (
*** Cephalozia lacinulata (J.B.Jack ex Gottsche et Rabenh.) Spruce, Cephalozia: 45, 1882 (
*** Cephalozia lucens (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 496 (326), 1908 (
*** Cephalozia macgregorii (Steph.) Váňa, Phytotaxa 112 (1): 12, 2013 (
*** Cephalozia macounii (Austin) Austin, Hepat. bor.-amer.: 14, 1873 (
*** Cephalozia maxima Steph., Sp. Hepat. (Stephani) 6: 441, 1924 (
*** Cephalozia mollusca (De Not.) Váňa, Phytotaxa 112 (1): 12, 2013 (
* Cephalozia neesiana Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 429 (305), 1908 (
*** Cephalozia nishimurae (N.Kitag.) Váňa, Phytotaxa 112 (1): 12, 2013 (
** Cephalozia pachygyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007 (
** Cephalozia physocaula (Hook.f. et Taylor) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 485 (315), 1908 (
** Cephalozia schusteriana J.J.Engel, Novon 17 (3): 313, 2007 (
*** Cephalozia stolonacea (Herzog) Váňa, Phytotaxa 112 (1): 12, 2013 (
*** Cephalozia tubulata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (
Excluded from the genus
* Cephalozia hians Steph., Sp. Hepat. (Stephani) 6: 441, 1924 (
* Cephalozia indica Udar et D.Kumar, Geophytology 6 (1): 42, 1976 (
* Cephalozia kodaikanalensis G.Asthana et Saumya Srivast., Geophytology 43 (1): 63, 2013 (
* Cephalozia parvifolia Arnell, Rev. Bryol. 25 (1): 1, 1898 (
* Cephalozia tricuspidata (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (
*** Fuscocephaloziopsis Fulford, Mem. New York Bot. Gard. 11 (3): 353, 1968 (
** Fuscocephaloziopsis affinis (Lindb. ex Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis africana (Váňa) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis albescens (Hook.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
* Fuscocephaloziopsis albescens var. islandica (Nees) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis baldwinii (C.M.Cooke) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
** Fuscocephaloziopsis biloba (Herzog) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968 (
*** Fuscocephaloziopsis catenulata (Huebener) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis catenulata subsp. nipponica (S.Hatt.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis connivens (Dicks.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis connivens subsp. fissa (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis connivens subsp. sandwicensis (Mont.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis crassifolia (Lindenb. et Gottsche) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis gollanii (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis leucantha (Spruce) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis loitlesbergeri (Schiffn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis lunulifolia (Dumort.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis macrostachya (Kaal.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis macrostachya subsp. australis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis macrostachya subsp. macrostachya var. spiniflora (Schiffn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis monticola (J.D.Godfrey) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
*** Fuscocephaloziopsis pachycaulis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
*** Fuscocephaloziopsis pleniceps (Austin) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
** Fuscocephaloziopsis pleniceps var. caroliniana (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
*** Fuscocephaloziopsis pulvinata (Steph.) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968 (
* Fuscocephaloziopsis schusteri (Sushil K.Singh et D.K.Singh) Váňa, Phytotaxa 183 (4): 291, 2014 (
*** Fuscocephaloziopsis subintegra Gradst. et Váňa, Cryptog. Bryol. 25 (3): 274, 2004 (
*** Fuscocephaloziopsis zoopsioides (Horik.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
*** Nowellia Mitt., Nat. hist. Azores: 321, 1870 (
** sect. Acronowellia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974 (
*** Nowellia reedii H.Rob., Bryologist 73 (1): 150, 1970 (
*** Nowellia yunckeri Fulford, Mem. New York Bot. Gard. 11 (3): 329, 1968 (
** sect. Metanowellia (Grolle) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974 (
*** Nowellia borneensis (De Not.) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (
*** Nowellia dominicensis Steph., Sp. Hepat. (Stephani) 6: 443, 1924 (
*** Nowellia evansii Grolle, J. Hattori Bot. Lab. 31: 33, 1968 (
*** Nowellia langii Pearson, J. Linn. Soc., Bot. 46 (305): 25, 1922 (
*** Nowellia pusilla Grolle, J. Hattori Bot. Lab. 31: 37, 1968 (
*** Nowellia wrightii (Gottsche ex Spruce) Steph. ex Duss, Enum. musc. Antilles franç., Hép.: 21, 1903 (
** sect. Nowellia
*** Nowellia aciliata (P.C.Chen et P.C.Wu) Mizut., Hikobia 11: 469, 1994 (
*** Nowellia curvifolia (Dicks.) Mitt., Nat. hist. Azores: 321, 1870 (
*** Odontoschisma (Dumort.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (
** sect. Cladopodiella (H.Buch) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (
*** Odontoschisma francisci (Hook.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013 (
** sect. Denudata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 833, 1974 (
*** Odontoschisma brasiliense Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 585 (369), 1908 (
** Odontoschisma cleefii Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (
*** Odontoschisma denudatum (Mart.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (
** Odontoschisma denudatum subsp. naviculare (Steph.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (
** Odontoschisma denudatum subsp. sandvicense (Ångstr.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (
*** Odontoschisma elongatum (Lindb.) A.Evans, Rhodora 14 (157): 13, 1912 (
*** Odontoschisma engelii Gradst. et Burghardt, Fieldiana, Bot. (n.ser.) 47: 194, 2008 (
*** Odontoschisma longiflorum (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877 (
*** Odontoschisma macounii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 92, 1884 (
*** Odontoschisma portoricense (Hampe et Gottsche) Steph., Hedwigia 27 (11/12): 296, 1888 (
** Odontoschisma pseudogrosseverrucosum Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (
*** Odontoschisma purpuratum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 297, 1950 (
*** Odontoschisma soratamum Fulford, Mem. New York Bot. Gard. 11 (3): 338, 1968 (
*** Odontoschisma variabile (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877 (
** Odontoschisma zhui Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1020, 2014 (
** sect. Iwatsukia (N.Kitag.) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (
*** Odontoschisma bifidum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (
*** Odontoschisma jishibae (Steph.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 13, 2013 (
*** Odontoschisma spinosum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (
** sect. Neesia Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (
*** Odontoschisma fluitans (Nees) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013 (
** sect. Odontoschisma
*** Odontoschisma grosseverrucosum Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 593 (377), 1908 (
*** Odontoschisma sphagni (Dicks.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (
** Odontoschisma steyermarkii Gradst. et Ilk.-Borg., Nova Hedwigia 100 (1/2): 38, 2015 [2014 online] (
Incertae sedis
* Odontoschisma obcordatum (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 586 (370), 1908 (
** Schiffneria Steph., Österr. Bot. Z. 44 (1): 1, 1894 (
*** Schiffneria hyalina Steph., Österr. Bot. Z. 44 (1): 1, 1894 (
** Haesselia Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988 (
*** Haesselia acuminata Gradst., Trop. Bryol. 1: 30, 1989 (
*** Haesselia roraimensis Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988 (
** Trabacellula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967 (
*** Trabacellula tumidula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967 (
by J. Váňa
The treatment of Cephaloziellaceae follows what was outlined in
*** Allisoniella E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 80, 1965 (
** sect. Allisoniella
*** Allisoniella nigra (Rodway) R.M.Schust., Nova Hedwigia 22: 137, 1971 [1972] (
** Allisoniella nigra var. acutiloba J.J.Engel, Novon 17 (3): 313, 2007 (
** Allisoniella nigra subsp. novaezelandiae R.M.Schust., Nova Hedwigia 22: 143, 1971 [1972] (
*** Allisoniella recurva R.M.Schust., Nova Hedwigia 22: 146, 1971 [1972] (
*** Allisoniella subbipartita (C.Massal.) R.M.Schust. et J.J.Engel, Nova Hedwigia 22: 147, 1971 [1972] (
*** Allisoniella tasmanica R.M.Schust., Nova Hedwigia 22: 145, 1971 [1972] (
** sect. Protomarsupella (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 150, 1971 (
*** Allisoniella scottii (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 151, 1971 [1972] (
*** Amphicephalozia R.M.Schust., Nova Hedwigia 22: 131, 1971 [1972] (
*** Amphicephalozia africana Váňa et M.Wigginton, J. Bryol. 30 (1): 55, 2008 (
*** Amphicephalozia amplexicaulis R.M.Schust., Nova Hedwigia 22: 133, 1971 [1972] (
*** Amphicephalozia geisslerae Pócs et Váňa, Polish Bot. J. 46 (2): 145, 2001 (
*** Anastrophyllopsis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013 (
*** Anastrophyllopsis involutifolia (Mont. ex Gottsche, Lindenb. et Nees) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013 (
*** Anastrophyllopsis revoluta (Steph.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013 (
*** Anastrophyllopsis subcomplicata (Lehm. et Lindenb.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013 (
** Cephalojonesia Grolle, Rev. Bryol. Lichénol. 37 (4): 763, 1970 [1971] (
*** Cephalojonesia incuba Grolle et Vanden Berghen, Rev. Bryol. Lichénol. 37 (4): 764, 1970 [1971] (
*** Cephalojonesia incuba subsp. mexicana Burghardt, Gradst. et Váňa, J. Hattori Bot. Lab. 100: 35, 2006 (
** Cephalomitrion R.M.Schust., Nova Hedwigia 61 (3/4): 550, 1995 (
*** Cephalomitrion aterrimum (Steph.) R.M.Schust., Nova Hedwigia 61 (3/4): 554, 1995 (
*** Cephaloziella (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (
** subg. Cephaloziella
*** Cephaloziella aenigmatica R.M.Schust., Nova Hedwigia 63 (1/2): 20, 1996 (
*** Cephaloziella anthelioides S.W.Arnell, Bot. Not. 105: 322, 1952 (
*** Cephaloziella arctogena (R.M.Schust.) Konstant., Arctoa 3: 126, 1994 (
** Cephaloziella arenaria (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (
*** Cephaloziella aspericaulis Jørg., Bergens Mus. Skr. (n.ser.) 16: 197, 1934 (
*** Cephaloziella baumgartneri Schiffn., Verh. K.K. Zool.-Bot. Ges. Wien 56 (3): 273, 1906 (
** Cephaloziella breviperianthia C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 208, 1981 (
** Cephaloziella brinkmannii Douin, Mém. Soc. Bot. France 29: 75, 1920 (
*** Cephaloziella capensis (Sim) S.W.Arnell, Bot. Not. 105: 326, 1952 (
*** Cephaloziella capillaris (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920 (
*** Cephaloziella crassigyna (R.M.Schust.) R.M.Schust., Nova Hedwigia 61 (3/4): 556, 1995 (
*** Cephaloziella crispata N.Kitag., J. Hattori Bot. Lab. 32: 301, 1969 (
*** Cephaloziella densifolia R.M.Schust., Nova Hedwigia 22: 199, 1971 [1972] (
** Cephaloziella densifolia var. dubia R.M.Schust., Nova Hedwigia 22: 202, 1971 [1972] (
*** Cephaloziella divaricata (Sm.) Schiffn., Hepat. (Engl.-Prantl): 99, 1893 (
** Cephaloziella divaricata var. scabra (M.Howe) Haynes, Bryologist 12 (4): 68, 1909 (
** Cephaloziella dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 49, 1900 (
*** Cephaloziella elachista (J.B.Jack ex Gottsche et Rabenh.) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900 (
** Cephaloziella elachista var. spinophylla (C.Gao) C.Gao, Fl. Bryoph. Sin. 9: 181, 2003 (
** Cephaloziella elegans (Heeg) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900 (
*** Cephaloziella exigua R.M.Schust., Nova Hedwigia 63 (1/2): 49, 1996 (
*** Cephaloziella exiliflora (Taylor) Douin, Mém. Soc. Bot. France 29: 72, 1920 (
** Cephaloziella fragillima (Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 409, 1976 (
*** Cephaloziella garsidei S.W.Arnell, Rev. Bryol. Lichénol. 23 (1/2): 173, 1954 (
*** Cephaloziella grandiretis (R.M.Schust.) R.M.Schust., Nova Hedwigia 63 (1/2): 35, 1996 (
*** Cephaloziella grimsulana (J.B.Jack ex Gottsche et Rabenh.) Lacout., Hépat. France: 52, 1905 (
*** Cephaloziella hampeana (Nees) Schiffn. ex Loeske, Moosfl. Harz.: 92, 1903 (
** Cephaloziella hebridensis Steph., Hedwigia 32 (5): 316, 1893 (
*** Cephaloziella herzogiana (Pandé et K.P.Srivast.) Udar et D.Kumar, Geophytology 6 (1): 45, 1976 (
** Cephaloziella heteroica (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 76, 1920 (
** Cephaloziella hyalina Douin, Mém. Soc. Bot. France 29: 77, 1920 (
* Cephaloziella hyalina var. rappii (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 56, 1980 (
*** Cephaloziella inaequalis R.M.Schust., Nova Hedwigia 22: 186, 1971 [1972] (
*** Cephaloziella invisa R.M.Schust., Nova Hedwigia 63 (1/2): 30, 1996 (
** Cephaloziella kilohanensis (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 85, 1920 (
*** Cephaloziella longii Váňa, Folia Geobot. Phytotax. 27 (2): 193, 1992 (
*** Cephaloziella lycopodioides (Sim) S.W.Arnell, Bot. Not. 105: 321, 1952 (
** Cephaloziella mammillifera R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (
*** Cephaloziella massalongi (Spruce) Müll.Frib., Lebermoose 2 (17): 191, 1913 (
*** Cephaloziella muelleriana R.M.Schust., Nova Hedwigia 63 (1/2): 24, 1996 (
*** Cephaloziella natalensis (Sim) S.W.Arnell, Hepat. South Africa: 338, 1963 (
** Cephaloziella nicholsonii Douin, Rev. Bryol. 40 (6): 81, 1913 (
*** Cephaloziella nothogena R.M.Schust., Nova Hedwigia 63 (1/2): 37, 1996 (
** Cephaloziella obtusilobula R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 108, 1980 (
** Cephaloziella patulifolia (Steph.) Douin, Mém. Soc. Bot. France 29: 70, 1920 (
*** Cephaloziella phyllacantha (C.Massal. et Carestia) Müll.Frib., Lebermoose 2 (17): 194, 1913 (
*** Cephaloziella polystratosa (R.M.Schust. et Damsh.) Konstant., Bot. Zhurn. (Moscow & Leningrad) 85 (10): 127, 2000 (
*** Cephaloziella pseudocrassigyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007 (
*** Cephaloziella pulcherrima R.M.Schust., Nova Hedwigia 22: 203, 1971 [1972] (
** Cephaloziella pulcherrima subsp. sphagnicola R.M.Schust., Nova Hedwigia 22: 205, 1971 [1972] (
** Cephaloziella pungens Steph. ex Fulford, Mem. New York Bot. Gard. 11 (4): 410, 1976 (
*** Cephaloziella rubella (Nees) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 231, 1902 (
*** Cephaloziella schelpei S.W.Arnell, Bot. Not. 110 (1): 18, 1957 (
*** Cephaloziella spegazziniana (C.Massal.) Douin, Mém. Soc. Bot. France 29: 69, 1920 (
*** Cephaloziella spinicaulis Douin, Mém. Soc. Bot. France 29: 62, 1920 (
*** Cephaloziella spinigera (Lindb.) Jørg., Bergens Mus. Skr. (n.ser.) 16: 189, 1934 (
*** Cephaloziella stellulifera (Taylor ex Carrington et Pearson) Croz., Rev. Bryol. 30 (2): 31, 1903 (
*** Cephaloziella stephanii Schiffn. ex Douin, Mém. Soc. Bot. France 29: 85, 1920 (
** Cephaloziella stolonifera R.M.Schust., Phytologia 39 (6): 426, 1978 (
** Cephaloziella subtilis (Lindenb. et Gottsche) Steph., Hedwigia 32 (5): 318, 1893 (
*** Cephaloziella sumatrana Schiffn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920 (
*** Cephaloziella tabularis S.W.Arnell, Bot. Not. 105: 318, 1952 (
*** Cephaloziella transvaalensis S.W.Arnell, Bot. Not. 110 (1): 19, 1957 (
*** Cephaloziella triplicata S.W.Arnell, Bot. Not. 115: 203, 1962 (
*** Cephaloziella umtaliensis S.W.Arnell, Bot. Not. 110 (1): 20, 1957 (
*** Cephaloziella uncinata R.M.Schust., Meddel. Grønland 199 (1): 316, 1974 (
** Cephaloziella uncinata var. brevigyna R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (
** Cephaloziella uncinata var. sphagnicola R.M.Schust., Meddel. Grønland 199 (1): 323, 1974 (
*** Cephaloziella vaginans Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 119, 1911 (
*** Cephaloziella varians (Gottsche) Steph., Wiss. Ergebn. Schwed. Südpolar-Exped. [1901–1903] 4 (1): 4, 1905 (
*** Cephaloziella verrucosa Steph., Hedwigia 32 (5): 318, 1893 (
* Cephaloziella villaumei (Steph.) Váňa, Phytotaxa 112 (1): 3, 2013 (
** Cephaloziella violacea Schljakov, Novosti Sist. Nizš. Rast. 15: 242, 1978 (
** Cephaloziella welwitschii (Steph.) Douin, Mém. Soc. Bot. France 29: 58, 1920 (
** subg. Dichiton (Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (
*** Cephaloziella calyculata (Durieu et Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (
*** Cephaloziella integerrima (Lindb.) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 232, 1902 (
** subg. Evansia (Douin et Schiffn.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (
*** Cephaloziella antillana (Besch. et Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 405, 1976 (
*** Cephaloziella dentata (Raddi) Steph., Bull. Herb. Boissier 5 (2): 78, 1897 (
*** Cephaloziella hirta (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (
** Cephaloziella squarrosula (Trevis.) Steph., Hedwigia 32 (5): 318, 1893 (
*** Cephaloziella subspinosa R.M.Schust., Nova Hedwigia 22: 210, 1971 [1972] (
** subg. Prionolobus (Spruce) Müll.Frib., Lebermoose 2 (16): 110, 1912 (
*** Cephaloziella acanthophora (S.Hatt.) Horik., Hikobia 1 (2): 79, 1951 (
*** Cephaloziella biokoensis Váňa et Frank Müll., Trop. Bryol. 24: 1, 2003 (
*** Cephaloziella granatensis (J.B.Jack ex Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 411, 1976 (
*** Cephaloziella grisea R.M.Schust., Phytologia 39 (6): 425, 1978 (
* Cephaloziella meghalayensis Udar et Ad.Kumar, Lindbergia 8 (1): 34, 1982 (
*** Cephaloziella microphylla (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920 (
*** Cephaloziella tenuissima (Lehm.) Steph., Hedwigia 32 (5): 318, 1893 (
*** Cephaloziella turneri (Hook.) Müll.Frib., Lebermoose 2 (17): 202, 1913 (
Incertae sedis
** Cephaloziella dentifolia Udar et Ad.Kumar, Lindbergia 8 (1): 30, 1982 (
** Cephaloziella filum (Trevis.) Steph., Hedwigia 32 (5): 318, 1893 (
* Cephaloziella flexuosa C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 88, 1984 (
** Cephaloziella intricata Schiffn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920 (
** Cephaloziella levieri Schiffn. ex Douin, Mém. Soc. Bot. France 29: 80, 1920 (
*** Cephaloziella pellucida R.M.Schust., Nova Hedwigia 63 (1/2): 57, 1996 (
* Cephaloziella pygmaea (Spruce) Váňa, Phytotaxa 183 (4): 290, 2014 (
* Cephaloziella secundifolia Pearson, Ann. Bryol. 4: 106, 1931 (
** Cephaloziella sinensis Douin, Rev. Bryol. 41 (1): 8, 1914 (
** Cephaloziopsis (Spruce) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (
*** Cephaloziopsis intertexta (Gottsche) R.M.Schust., Nova Hedwigia 22: 183, 1971 [1972] (
** Chaetophyllopsis R.M.Schust., J. Hattori Bot. Lab. 23: 69, 1960 [1961] (
*** Chaetophyllopsis whiteleggei (Carrington et Pearson) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 255, 1972 (
** Cylindrocolea R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
** subg. Cylindrocolea
*** Cylindrocolea kiaeri (Austin) Váňa, Phytotaxa 112 (1): 2, 2013 (
*** Cylindrocolea sanctae-helenae M.Wigginton, Polish Bot. J. 58 (1): 107, 2013 (
** sect. Cylindrocolea
*** Cylindrocolea andersonii R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 33, 1980 (
*** Cylindrocolea brasiliensis D.P.Costa, N.D.Santos et Váňa, Bryologist 111 (4): 667, 2008 (
*** Cylindrocolea chevalieri (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
*** Cylindrocolea gittinsii (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972] (
*** Cylindrocolea madagascariensis (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
*** Cylindrocolea nigerica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972] (
*** Cylindrocolea novae-caledoniae (Grolle) R.M.Schust., Nova Hedwigia 22: 161, 1971 [1972] (
*** Cylindrocolea planifolia (Steph.) R.M.Schust., Nova Hedwigia 22: 164, 1971 [1972] (
*** Cylindrocolea sprucei R.M.Schust., Nova Hedwigia 22: 163, 1971 [1972] (
*** Cylindrocolea ugandica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 171, 1971 [1972] (
** sect. Platycaulis R.M.Schust., Nova Hedwigia 22: 173, 1971 (
*** Cylindrocolea recurvifolia (Steph.) Inoue, J. Jap. Bot. 47 (11): 348, 1972 (
*** Cylindrocolea tagawae (N.Kitag.) R.M.Schust., Nova Hedwigia 22: 174, 1971 [1972] (
** subg. Cylindroscyphus (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 20, 1980 (
*** Cylindrocolea abyssinica (Gola) Váňa, Phytotaxa 112 (1): 2, 2013 (
** Cylindrocolea obtusifolia Fulford, Mem. New York Bot. Gard. 11 (4): 401, 1976 (
** Cylindrocolea reticulata Udar et Ad.Kumar, Lindbergia 8 (3): 181, 1982 [1983] (
*** Cylindrocolea rhizantha (Mont.) R.M.Schust., Nova Hedwigia 22: 175, 1971 [1972] (
*** Gottschelia Grolle, J. Hattori Bot. Lab. 31: 13, 1968 (
*** Gottschelia maxima (Steph.) Grolle, J. Bryol. 25 (1): 6, 2003 (
*** Gottschelia schizopleura (Spruce) Grolle, J. Hattori Bot. Lab. 31: 16, 1968 (
** Gymnocoleopsis (R.M.Schust.) R.M.Schust., Phytologia 39 (4): 243, 1978 (
*** Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust., J. Hattori Bot. Lab. 78: 123, 1995 (
*** Gymnocoleopsis cylindriformis (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 78: 126, 1995 (
*** Herzogobryum Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964] (
*** Herzogobryum atrocapillum (Hook.f. et Taylor) Grolle, Österr. Bot. Z. 113 (2): 228, 1966 (
*** Herzogobryum filiforme R.M.Schust., Phytologia 45 (5): 422, 1980 (
*** Herzogobryum molle Grolle, Österr. Bot. Z. 113 (2): 226, 1966 (
*** Herzogobryum vermiculare (Schiffn.) Grolle, J. Hattori Bot. Lab. 28: 103, 1965 (
** Kymatocalyx Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 56, 1950 (
*** Kymatocalyx africanus Váňa et M.Wigginton, Haussknechtia, Beih. 9: 158, 1999 (
*** Kymatocalyx dominicensis (Spruce) Váňa, Österr. Bot. Z. 118 (5): 575, 1970 (
*** Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa, Haussknechtia, Beih. 9: 164, 1999 (
*** Kymatocalyx rhizomaticus (Herzog) Gradst. et Váňa, Haussknechtia, Beih. 9: 166, 1999 (
*** Lophonardia R.M.Schust., Phytologia 39 (4): 244, 1978 (
*** Lophonardia jamesonii (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 19, 2013 (
*** Lophonardia laxifolia (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013 (
*** Lophonardia tristaniana (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013 (
*** Nothogymnomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 43, 1996 (
*** Nothogymnomitrion erosum (Carrington et Pearson) R.M.Schust., J. Hattori Bot. Lab. 80: 44, 1996 (
*** Obtusifolium S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 309, 1956 (
*** Obtusifolium obtusum (Lindb.) S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 133, 1956 (
*** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010 (
** Phycolepidozia R.M.Schust., Bull. Torrey Bot. Club 93 (6): 438, 1966 [1967] (
** subg. Metaphycolepidozia Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 506, 2014 (
** Phycolepidozia indica Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 499, 2014 (
** subg. Phycolepidozia
*** Phycolepidozia exigua R.M.Schust., Bull. Torrey Bot. Club 93 (6): 440, 1966 [1967] (
** Protolophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979 (
** Protolophozia androgyna R.M.Schust. ex Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
** Protolophozia autoica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
** Protolophozia crispata (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia druceae (Grolle et E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia elongata (Steph.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979 (
*** Protolophozia herzogiana (E.A.Hodgs. et Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia lancistipa (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia leucorhiza (Mitt.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia longiflora (Herzog) L.Söderstr. et Váňa, Phytotaxa 76 (3): 51, 2013 (
** Protolophozia monoica (E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
*** Protolophozia multicuspidata (Hook.f. et Taylor) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
** Protolophozia nivicola (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
*** Protolophozia perssoniana (H.A.Mill.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
** Protolophozia subalpina (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
*** Protolophozia tasmanica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
*** Protolophozia verruculosa (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
by L. Söderström and J. Váňa
Lophoziaceae was broadly defined by
*** Andrewsianthus R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 66, 1961 (
** subg. Andrewsianthus
*** Andrewsianthus aberrans (Nees et Mont.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 440, 1963 (
*** Andrewsianthus bidens (Mitt. ex Steph.) R.M.Schust., Nova Hedwigia 15: 492, 1968 (
*** Andrewsianthus bilobus (Mitt.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 437, 1963 (
*** Andrewsianthus carinatus Grolle, Marion Prince Edw. Is: 229, 1971 (
*** Andrewsianthus cavifolius Grolle et Váňa, J. Hattori Bot. Lab. 38: 640, 1974 (
*** Andrewsianthus chimbuensis R.M.Schust., Nova Hedwigia 15: 491, 1968 (
*** Andrewsianthus kinabaluensis N.Kitag., J. Hattori Bot. Lab. 33: 205, 1970 (
*** Andrewsianthus koponenii Váňa et Piippo, Ann. Bot. Fenn. 26 (3): 284, 1989 (
*** Andrewsianthus mizutanii N.Kitag., J. Hattori Bot. Lab. 32: 307, 1969 (
*** Andrewsianthus papillosus N.Kitag., J. Hattori Bot. Lab. 33: 207, 1970 (
*** Andrewsianthus perigonialis (Hook.f. et Taylor) R.M.Schust., Beih. Nova Hedwigia 119: 336, 2002 (
*** Andrewsianthus puniceus (Nees) R.M.Schust. ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 439, 1963 (
*** Andrewsianthus recurvifolius (Nees) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969 (
*** Andrewsianthus sundaicus (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969 (
*** Andrewsianthus zantenii Váňa, J. Hattori Bot. Lab. 38: 645, 1974 (
** subg. Cephalolobus (R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 326, 2002 (
*** Andrewsianthus hodgsoniae (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (
*** Andrewsianthus marionensis (S.W.Arnell) Grolle, Marion Prince Edw. Is: 232, 1971 (
*** Andrewsianthus scabrellus (C.Massal.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (
** Andrewsianthus sphenoloboides (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (
** Gerhildiella Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966 (
*** Gerhildiella rossneriana Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966 (
*** Heterogemma (Jørg.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Heterogemma capitata (Hook.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Heterogemma laxa (Lindb.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Heterogemma patagonica (Herzog et Grolle) L.Söderstr. et Váňa, Phytotaxa 97 (2): 29, 2013 (
*** Lophozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
*** Lophozia ascendens (Warnst.) R.M.Schust., Bryologist 55 (3): 180, 1952 (
* Lophozia austrosibirica Bakalin, Ann. Bot. Fenn. 40 (1): 49, 2003 (
** Lophozia ciliata Damsh., L.Söderstr. et H.Weibull, Lindbergia 25 (1): 3, 2000 (
*** Lophozia guttulata (Lindb. et Arnell) A.Evans, Proc. Wash. Acad. Sci. 2: 302, 1900 (
* Lophozia jamaicensis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 128 (155), 1901 (
** Lophozia lacerata N.Kitag., Hikobia 3 (3): 172, 1963 (
** Lophozia lantratoviae Bakalin, Ann. Bot. Fenn. 40 (1): 47, 2003 (
** Lophozia murmanica Kaal., Rep. Second Norweg. Arctic Exped. 11: 34, 1906 (
** Lophozia pacifica Bakalin, Bryologist 114 (2): 302, 2011 (
* Lophozia pallida (Steph.) Grolle, J. Jap. Bot. 39 (6): 174, 1964 (
** Lophozia savicziae Schljakov, Novosti Sist. Nizš. Rast. 10: 299, 1973 (
* Lophozia schusterana Schljakov, Novosti Sist. Nizš. Rast. 12: 320, 1975 (
*** Lophozia silvicola H.Buch, Beret. 18 Skand. Naturforskarmøde: 228, 1929 (
*** Lophozia silvicoloides N.Kitag., J. Hattori Bot. Lab. 28: 276, 1965 (
* Lophozia subapiculata R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 104, 1974 (
** Lophozia udarii S.Srivast., S.C.Srivast. et K.K.Rawat, Nelumbo 55: 130, 2013 (
*** Lophozia ventricosa (Dicks.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
* Lophozia wenzelii (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 35 (135), 1902 (
Excluded from the genus
* Lophozia serpens (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
** Lophoziopsis Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Lophoziopsis excisa (Dicks.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Lophoziopsis excisa var. elegans (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Lophoziopsis excisa var. infuscata (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Lophoziopsis excisa var. succulenta (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Lophoziopsis longidens (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Lophoziopsis longidens subsp. arctica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 97 (2): 28, 2013 (
*** Lophoziopsis pellucida (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
** Lophoziopsis pellucida var. minor (R.M.Schust.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 28, 2013 (
*** Lophoziopsis polaris (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
** Lophoziopsis polaris var. sphagnorum (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
* Lophoziopsis propagulifera (Gottsche) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
** Lophoziopsis rubrigemma (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
** Pseudocephaloziella R.M.Schust., Phytologia 39 (4): 243, 1978 (
*** Pseudocephaloziella epiphytica R.M.Schust., Phytologia 39 (4): 243, 1978 (
*** Trilophozia (R.M.Schust.) Bakalin, Monogr. Lophozia: 34, 2005 (
*** Trilophozia quinquedentata (Huds.) Bakalin, Monogr. Lophozia: 34, 2005 (
** Trilophozia quinquedentata var. asymmetrica (Horik.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 32, 2013 (
*** Tritomaria Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (
*** Tritomaria exsecta (Schmidel) Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (
** Tritomaria exsecta subsp. novaezelandiae J.J.Engel, Bryologist 109 (1): 61, 2006 (
*** Tritomaria exsectiformis (Breidl.) Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (
** Tritomaria exsectiformis subsp. arctica R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 661, 1969 (
** Tritomaria exsectiformis subsp. camerunensis S.W.Arnell ex Váňa, Phytotaxa 167 (2): 216, 2014 (
*** Tritomaria ferruginea (Grolle) Váňa, Phytotaxa 81 (1): 24, 2013 (
* Tritomaria koreana Bakalin, S.S.Choi et B.Y.Sun, Arctoa 18: 163, 2009 [2010] (
* Tritomaria mexicana Bakalin, Arctoa 17: 162, 2008 [2009] (
*** Tritomaria scitula (Taylor) Jørg., Bergens Mus. Aarbok 1919/20 (7): 3, 1922 (
by J. Váňa
Scapaniaceae, in its classical concept, was recognized as monophyletic and nested within Lophoziaceae s. lat. (cf.
*** Diplophyllum (Dumort.) Dumort., Recueil Observ. Jungerm.: 15, 1835 (
*** subg. Austrodiplophyllum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 18, 1968 (
** Diplophyllum recurvifolium C.Massal., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 221, 1928 (
*** Diplophyllum squarrosum Steph., Sp. Hepat. (Stephani) 4: 116, 1910 (
*** Diplophyllum verrucosum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 19, 1968 (
*** subg. Diplophyllum
*** sect. Diplophyllum
*** Diplophyllum albicans (L.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (
*** sect. Protodiplophyllum (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 29, 2013 (
*** Diplophyllum africanum S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 527, 1956 (
*** Diplophyllum andicola R.M.Schust., Phytologia 39 (4): 248, 1978 (
*** Diplophyllum andrewsii A.Evans, Bryologist 25 (2): 28, 1922 (
* Diplophyllum androgynum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 277, 1998 (
* Diplophyllum angustifolium J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 262, 1998 (
*** Diplophyllum apiculatum (A.Evans) Steph., Sp. Hepat. (Stephani) 4: 110, 1910 (
** Diplophyllum dioicum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 20, 1968 (
** Diplophyllum exiguum Steph., Sp. Hepat. (Stephani) 6: 500, 1924 (
* Diplophyllum gemmiparum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 255, 1998 (
* Diplophyllum incrassatum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 265, 1998 (
*** Diplophyllum nanum Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 48, 1950 [1951] (
* Diplophyllum novum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 274, 1998 (
*** Diplophyllum obtusatum (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 215, 1974 (
*** Diplophyllum obtusifolium (Hook.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (
** Diplophyllum obtusifolium subsp. domesticum (Gottsche) Váňa, Phytotaxa 76 (3): 29, 2013 (
*** Diplophyllum serrulatum (Müll.Frib.) Steph., Sp. Hepat. (Stephani) 4: 112, 1910 (
*** Diplophyllum taxifolium (Wahlenb.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (
** Diplophyllum taxifolium var. mucronatum R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 203, 1974 (
*** Diplophyllum trollii Grolle, Khumbu Himal 1 (4): 273, 1966 (
*** Douinia (C.E.O.Jensen) H.Buch, Scapan. N.-Eur. Sib.: 13, 1928 (
*** Douinia imbricata (M.Howe) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013 (
*** Douinia ovata (Dicks.) H.Buch, Scapan. N.-Eur. Sib.: 14, 1928 (
*** Douinia plicata (Lindb.) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013 (
** Pseudotritomaria Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Pseudotritomaria heterophylla (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Saccobasis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 291, 1932 [1933] (
*** Saccobasis polita (Nees) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 292, 1932 [1933] (
** Saccobasis polymorpha (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979 (
*** Scapania (Dumort.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** subg. Ascapania Grolle, Khumbu Himal 1 (4): 268, 1966 (
*** Scapania contorta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861] (
** subg. Gracilidae (H.Buch) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (
*** Scapania ampliata Steph., Bull. Herb. Boissier 5 (2): 106, 1897 (
** Scapania ampliata subsp. queenslandica M.L.Hicks, J. Hattori Bot. Lab. 69: 130, 1991 (
*** Scapania bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 218, 1869 (
*** Scapania gracilis Lindb., Morgonbladet (Helsinki) 1873 (286, 9 Dec): 2, 1873 (
** Scapania macroparaphyllia T.Cao, C.Gao et J.Sun, Acta Phytotax. Sin. 42 (2): 180, 2004 (
*** Scapania maxima Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 223, 1934 (
*** Scapania nipponica (Amakawa et S.Hatt.) Amakawa, J. Hattori Bot. Lab. 30: 319, 1967 (
** Scapania paraphyllia T.Cao, C.Gao, J.Sun et B.R.Zuo, Acta Phytotax. Sin. 45 (3): 311, 2007 (
*** Scapania subnimbosa Steph., Sp. Hepat. (Stephani) 4: 150, 1910 (
** subg. Macroscapania R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 248, 1974 (
* Scapania geppii Steph., Hedwigia 44 (1): 14, 1904 (
*** Scapania portoricensis Hampe et Gottsche, Linnaea 25 (3): 342, 1852 [1853] (
** Scapania portoricensis var. boliviensis (Steph.) Herzog, Ann. Bryol. 1: 110, 1928 (
** Scapania portoricensis var. organensis (Herzog) Herzog, Ann. Bryol. 1: 110, 1928 (
** Scapania portoricensis var. roraimensis Warnst., Hedwigia 63 (2): 109, 1921 (
** subg. Plicaticalyx Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 36, 1903 (
** sect. Planifoliae (Müll.Frib.) Potemkin, J. Hattori Bot. Lab. 85: 56, 1998 (
*** Scapania davidii Potemkin, Ann. Bot. Fenn. 38 (2): 83, 2001 (
** Scapania ferruginaeoides T.Cao, C.Gao et J.Sun, Guihaia 24 (1): 23, 2004 (
** Scapania gaochii X.Fu ex T.Cao, Phytotaxa 97 (2): 26, 2013 (
*** Scapania harae Amakawa, J. Hattori Bot. Lab. 27: 5, 1964 (
*** Scapania nimbosa Taylor, Nov. Stirp. Pug. 8: 6, 1844 (
*** Scapania ornithopodioides (With.) Waddell, Moss Exch. Club Cat. Hepat.: 4, 1897 (
*** Scapania rotundifolia W.E.Nicholson, Symb. Sin. 5: 31, 1930 (
*** Scapania secunda Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 226, 1894 (
*** Scapania zhukovae Potemkin, Arctoa 9: 121, 2000 (
** sect. Plicaticalyx (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 326, 2002 (
*** Scapania ciliatospinosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 222, 1934 (
*** Scapania ferruginea (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 1: 72, 1844 (
*** Scapania hians Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 223, 1905 (
** Scapania hians subsp. salishensis J.D.Godfrey et G.Godfrey, Bryologist 81 (3): 362, 1978 (
*** Scapania orientalis Steph. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 606, 1901 (
*** Scapania pseudocontorta Potemkin, Arctoa 9: 115, 2000 (
*** Scapania sinikkae Potemkin, Ann. Bot. Fenn. 38 (2): 85, 2001 (
*** Scapania spiniloba Potemkin, Arctoa 9: 117, 2000 (
** subg. Pseudomacrodiplophyllum Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (
*** Scapania microdonta (Mitt.) Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 262, 1905 (
*** subg. Scapania
** sect. Aequilobae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 110, 1928 (
*** Scapania aequiloba (Schwägr.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
*** Scapania aspera M.Bernet et Bernet, Cat. hép. Suisse: 42, 1888 (
** sect. Americanae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (
*** Scapania americana Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 44, 1903 (
** sect. Apiculatae H.Buch, Scapan. N.-Eur. Sib.: 53, 1928 (
*** Scapania apiculata Spruce, Hep. Pyr. Exsic.: no. 15, 1847 (
*** Scapania carinthiaca J.B.Jack ex Lindb., Rev. Bryol. 7 (4): 77, 1880 (
*** Scapania carinthiaca var. massalongi Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 598, 1901 (
*** Scapania umbrosa (Schrad.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** sect. Ciliatae Grolle, Khumbu Himal 1 (4): 272, 1966 (
*** Scapania bhutanensis Amakawa, Fl. E. Himalaya 2: 230, 1971 (
*** Scapania ciliata Sande Lac., Prolus. fl. jap.: 209, 1867 (
*** Scapania ciliata subsp. hawaiica (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 321, 2002 (
*** Scapania hirosakiensis Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 120, 1905 (
*** Scapania hollandiae W.S.Hong, Bryologist 83 (1): 56, 1980 (
*** Scapania koponenii Potemkin, Ann. Bot. Fenn. 37 (1): 41, 2000 (
*** Scapania lepida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861] (
*** Scapania sandei Schiffn. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 612, 1901 (
** sect. Compactae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 101, 1928 (
*** Scapania compacta (Roth) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
*** Scapania kaurinii Ryan, Bot. Not. 42: 210, 1889 (
*** Scapania spitsbergensis (Lindb.) Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 607, 1901 (
** sect. Curtae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 55, 1928 (
*** Scapania curta (Mart.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** Scapania curta var. grandiretis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 393, 1974 (
** Scapania curta var. isoloba R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 396, 1974 (
*** Scapania diplophylloides Amakawa et S.Hatt., J. Hattori Bot. Lab. 9: 59, 1953 (
*** Scapania esterhuyseniae S.W.Arnell, Bot. Not. 110 (1): 26, 1957 (
*** Scapania fulfordiae W.S.Hong, Bryologist 83 (1): 46, 1980 (
*** Scapania gamundiae R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 14, 1968 (
*** Scapania helvetica Gottsche, Hepat. Eur., Leberm. 42-44: no. 426, 1868 (
*** Scapania irrigua (Nees) Nees, Syn. Hepat. 1: 67, 1844 (
** Scapania irrigua subsp. rufescens (Loeske) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 471, 1974 (
*** Scapania lingulata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 92, 1916 (
** Scapania lingulata var. microphylla (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 415, 1974 (
** Scapania magadanica S.S.Choi, Bakalin et B.Y.Sun, Bot. Pacifica 1: 46, 2012 (
*** Scapania mucronata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 91, 1916 (
*** Scapania obcordata (Berggr.) S.W.Arnell, Ark. Bot. (n.ser.) 4 (6): 117, 1959 (
** Scapania parvifolia Warnst., Hedwigia 63 (2): 78, 1921 (
*** Scapania praetervisa Meyl., Jahresber. Naturf. Ges. Graubündens (n.f.) 64: 364, 1926 (
*** Scapania scandica (Arnell et H.Buch) Macvicar, Stud. handb. Brit. hepat. (ed. 2): 394, 1926 (
** Scapania scandica var. argutedentata H.Buch, Scapan. N.-Eur. Sib.: 75, 1928 (
** Scapania scandica var. dimorpha R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 453, 1974 (
** Scapania scandica var. grandiretis (Schljakov) Schljakov, Pečen. Mchi Sev. SSSR 4: 152, 1981 (
*** Scapania uliginosa (Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
*** Scapania valdonii Váňa, Bedn.-Ochyra et Cykowska, Nova Hedwigia 89 (1/2): 126, 2009 (
*** Scapania zemliae S.W.Arnell, Svensk Bot. Tidskr. 41: 215, 1947 (
** sect. Cuspiduligerae H.Buch, Scapan. N.-Eur. Sib.: 125, 1928 (
*** Scapania cuspiduligera (Nees) Müll.Frib., Lebermoose 2 (22): 472, 1915 (
** Scapania cuspiduligera var. diplophyllopsis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 361, 1974 (
** sect. Grolleoscapania Potemkin, Ann. Bot. Fenn. 39 (4): 329, 2002 (
*** Scapania karl-muelleri Grolle, Khumbu Himal 1 (4): 270, 1966 (
** sect. Hyperboreae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012 (
*** Scapania hyperborea Jørg., Forh. Vidensk.-Selsk. Kristiania 1894 (8): 56, 1894 (
*** Scapania paludicola Loeske et Müll.Frib., Lebermoose 2 (21): 425, 1915 (
** Scapania paludicola var. viridigemma R.M.Schust., Bull. Natl. Mus. Canada 122: 20, 1950 [1951] (
*** Scapania tundrae (Arnell) H.Buch, Scapan. N.-Eur. Sib.: 99, 1928 (
** sect. Kaalaasia (H.Buch) Jørg., Bergens Mus. Skr. (n.ser.) 16: 210, 1934 (
*** Scapania calcicola (Arnell et J.Perss.) Ingham, Naturalist (Hull) 564: 11, 1904 (
*** Scapania gymnostomophila Kaal., Bot. Not. 49: 21, 1896 (
*** Scapania ligulifolia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 306, 1974 (
*** Scapania pseudocalcicola R.M.Schust., Phytologia 63 (5): 327, 1987 (
* Scapania sect. Muelleria Potemkin, Ann. Bot. Fenn. 39 (4): 320, 2002 (
*** Scapania himalayica Müll.Frib. ex Herzog, Ann. Bryol. 12: 81, 1939 (
*** Scapania schljakovii Potemkin, Ann. Bot. Fenn. 38 (2): 87, 2001 (
** sect. Nemorosae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 152, 1928 (
*** Scapania brevicaulis Taylor, London J. Bot. 5: 272, 1846 (
*** Scapania crassiretis Bryhn, Rev. Bryol. 19 (1): 7, 1892 (
* Scapania degenii Schiffn. ex Müll.Frib., Lebermoose 2 (22): 497, 1915 (
* Scapania glaucoviridis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 221, 1934 (
** Scapania grossidens Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 146, 1905 (
*** Scapania hedbergii S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 556, 1956 (
*** Scapania integerrima Steph., Sp. Hepat. (Stephani) 4: 148, 1910 (
*** Scapania matveyevae Potemkin, Arctoa 9: 97, 2000 (
*** Scapania nemorea (L.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964] (
** Scapania parvidens Steph., Hedwigia 44 (1): 15, 1904 (
*** Scapania parvitexta Steph., Bull. Herb. Boissier 5 (2): 107, 1897 (
*** Scapania rigida Nees, Syn. Hepat. 1: 69, 1844 (
** sect. Scapania
*** Scapania gigantea Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 15, 1931 (
** Scapania grandiloba Steph., Sp. Hepat. (Stephani) 6: 502, 1924 (
*** Scapania komagadakensis Amakawa, J. Hattori Bot. Lab. 31: 96, 1968 (
*** Scapania obscura (Arnell et C.E.O.Jensen) Schiffn., Österr. Bot. Z. 58 (10): 377, 1908 (
** Scapania paludosa (Müll.Frib.) Müll.Frib., Mitt. Bad. Bot. Vereins 4 (182/183): 287, 1902 (
*** Scapania rufidula Warnst., Hedwigia 63 (2): 94, 1921 (
*** Scapania serrulata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 539, 1974 (
*** Scapania subalpina (Nees ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** Scapania subalpina var. haynesiae Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (4): 638, 1946 (
** Scapania subalpina var. muddiae C.D.Bird et W.S.Hong, Bryologist 83 (1): 51, 1980 (
*** Scapania undulata (L.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** sect. Scapaniella (H.Buch) Potemkin, J. Hattori Bot. Lab. 85: 43, 1998 (
*** Scapania glaucocephala (Taylor) Austin, Bull. Torrey Bot. Club 6 (16): 85, 1876 (
*** Scapania glaucocephala var. saxicola (R.M.Schust.) Potemkin, Bryologist 102 (1): 36, 1999 (
* Scapania scapanioides (C.Massal.) Grolle, Feddes Repert. 87 (3/4): 235, 1976 (
** sect. Simmonsiae (R.M.Schust.) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012 (
*** Scapania simmonsii Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 51, 1906 (
** sect. Sphaeriferae Konstant. et Potemkin, Ann. Bot. Fenn. 31 (2): 125, 1994 (
*** Scapania sphaerifera H.Buch et Tuom., Memoranda Soc. Fauna Fl. Fennica 11: 227, 1936 (
** sect. Stephaniae Potemkin, J. Hattori Bot. Lab. 85: 57, 1998 (
*** Scapania griffithii Schiffn., Österr. Bot. Z. 49 (6): 204, 1899 (
*** Scapania javanica Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 575, 1853 (
* Scapania javanica var. scabra Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 397, 1955 (
*** Scapania ligulata Steph., Hedwigia 44 (1): 14, 1904 (
** Scapania ligulata subsp. stephanii (Müll.Frib.) Potemkin, Piippo et T.J.Kop., Ann. Bot. Fenn. 41 (6): 423, 2004 (
** sect. Verrucosae Potemkin, J. Hattori Bot. Lab. 85: 54, 1998 (
*** Scapania udarii S.C.Srivast. et A.Srivast., J. Indian Bot. Soc. 72: 237, 1993 (
*** Scapania verrucosa Heeg, Rev. Bryol. 20 (5): 81, 1893 (
*** Schistochilopsis (N.Kitag.) Konstant., Arctoa 3: 125, 1994 (
*** Schistochilopsis cornuta (Steph.) Konstant., Arctoa 3: 125, 1994 (
*** Schistochilopsis grandiretis (Lindb. ex Kaal.) Konstant., Arctoa 3: 125, 1994 (
** Schistochilopsis hyperarctica Konstant. et L.Söderstr., Phytotaxa 162 (4): 240, 2014 (
*** Schistochilopsis incisa (Schrad.) Konstant., Arctoa 3: 125, 1994 (
* Schistochilopsis nakanishii (Inoue) Konstant., Arctoa 3: 125, 1994 (
** Schistochilopsis opacifolia (Culm. ex Meyl.) Konstant., Arctoa 3: 125, 1994 (
*** Schistochilopsis setosa (Mitt.) Konstant., Arctoa 3: 125, 1994 (
by L. Briscoe and J.J. Engel
** Enigmella G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992 (
** Enigmella thallina G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992 (
*** Acrobolbus Nees, Syn. Hepat. 1: 5, 1844 (
** Acrobolbus africanus (Pearson) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus anisodontus (Hook.f. et Taylor) Briscoe, Phytotaxa 202 (1): 59, 2015 (
** Acrobolbus antillanus R.M.Schust., J. Hattori Bot. Lab. 90: 143, 2001 (
** Acrobolbus azoricus (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus caducifolius R.M.Schust., J. Hattori Bot. Lab. 90: 154, 2001 (
*** Acrobolbus ciliatus (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 86, 1893 (
*** Acrobolbus cinerascens (Lehm. et Lindenb.) Bastow, Pap. & Proc. Roy. Soc. Tasmania 1887: 242, 1888 (
*** Acrobolbus concinnus (Mitt.) Steph., Trans. & Proc. New Zealand Inst. 24: 399, 1892 (
** Acrobolbus cuneifolius (Steph.) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus diversifolius R.M.Schust., J. Hattori Bot. Lab. 90: 150, 2001 (
*** Acrobolbus epiphytus (Colenso) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus flavicans (J.J.Engel et Grolle) Briscoe et J.J.Engel, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus gradsteinii (Grolle) Briscoe, Phytotaxa 202 (1): 59, 2015 (
** Acrobolbus integrifolius (A.Evans) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus knightii (Mitt.) Briscoe, Phytotaxa 202 (1): 59, 2015 (
** Acrobolbus kunkelii (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus laxus (Lehm. et Lindenb.) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus limbatus (Steph.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus lophocoleoides (Mitt.) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
** Acrobolbus madeirensis (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 60, 2015 (
** Acrobolbus mittenii Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 460 (179), 1902 (
*** Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 64, 1961 (
*** Acrobolbus papillosus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus perpusillus (Colenso) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus perpusillus var. denticulatus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus plagiochiloides (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus pseudosaccatus (Grolle) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus renifolius (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (
** Acrobolbus ruwenzorensis (S.W.Arnell) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus saccatus (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (
*** Acrobolbus setulosus (Mitt.) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus spinifolius R.M.Schust., J. Hattori Bot. Lab. 90: 137, 2001 (
*** Acrobolbus sumatranus (Schiffn.) Briscoe, Phytotaxa 202 (1): 61, 2015 (
*** Acrobolbus surculosus (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (
*** Acrobolbus tenellus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (
*** Acrobolbus tenellus var. diversifolius (E.A.Hodgs.) Briscoe, Phytotaxa 202 (1): 61, 2015 (
*** Acrobolbus urvilleanus (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (
*** Acrobolbus viridis (Mitt.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 61, 2015 (
*** Acrobolbus wilsonii Nees, Syn. Hepat. 1: 5, 1844 (
** Acrobolbus wilsonii var. andinus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 522, 1885 (
Excluded from the genus
* Acrobolbus bispinosus (J.B.Jack et Steph.) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 459 (178), 1902 (
** Austrolophozia R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963 (
** Austrolophozia andina R.M.Schust., Nova Hedwigia 15: 495, 1968 (
*** Austrolophozia camensis (Steph.) Grolle ex Hässel et Solari, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 3 (6): 240, 1970 (
*** Austrolophozia paradoxa R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963 (
*** Goebelobryum Grolle, J. Hattori Bot. Lab. 25: 135, 1962 (
*** Goebelobryum grossitextum (Steph.) Grolle, J. Hattori Bot. Lab. 25: 137, 1962 (
*** Goebelobryum unguiculatum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 25: 137, 1962 (
*** Goebelobryum vermiculare J.J.Engel et Glenny, Nova Hedwigia 95 (3/4): 320, 2012 (
*** Lethocolea Mitt., Handb. N. Zeal. fl. 2: 751, 1867 (
** Lethocolea congesta (Lehm.) S.W.Arnell, Bot. Not. 108: 311, 1955 (
*** Lethocolea glossophylla (Spruce) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (
** Lethocolea indica G.Asthana et Maurya, Natl. Acad. Sci. Lett. 37 (6): 535, 2014 (
*** Lethocolea javanica (Schiffn.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (
** Lethocolea naruto-toganensis Furuki, Bryologist 104 (2): 306, 2001 (
*** Lethocolea pansa (Taylor) G.A.M.Scott et K.G.Beckm., Symp. Biol. Hung. 35: 212, 1987 (
*** Lethocolea radicosa (Lehm. et Lindenb.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (
* Lethocolea repens S.Winkl., Mitt. Inst. Colombo-Alemán Invest. Ci. 3: 67, 1969 (
*** Saccogynidium Grolle, J. Hattori Bot. Lab. 23: 43, 1960 [1961] (
** sect. Decurvum Grolle, J. Hattori Bot. Lab. 23: 59, 1960 (
*** Saccogynidium decurvum (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 59, 1960 [1961] (
** sect. Jugata Grolle, J. Hattori Bot. Lab. 23: 55, 1960 (
** Saccogynidium rigidulum (Nees) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961] (
** sect. Saccogynidium
*** Saccogynidium australe (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 49, 1960 [1961] (
** Saccogynidium caldense (Ångstr.) Grolle, J. Hattori Bot. Lab. 23: 44, 1960 [1961] (
** Saccogynidium goebelii (Herzog) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961] (
** Saccogynidium muricellum (De Not.) Grolle, J. Hattori Bot. Lab. 36: 80, 1972 [1973] (
*** Saccogynidium vasculosum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 23: 46, 1960 [1961] (
Incertae sedis
** Saccogynidium chiloscyphoides R.M.Schust., J. Hattori Bot. Lab. 26: 272, 1963 (
** Saccogynidium irregularospinum C.Gao, T.Cao et M.J.Lai, Bryologist 104 (1): 129, 2001 (
*** Anthelia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
*** Anthelia julacea (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
** Anthelia juratzkana (Limpr.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (
*** Arnellia Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889 (
*** Arnellia fennica (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889 (
by J.J. Engel with contribution by J. Váňa (Neesioscyphus)
The placement of Pseudoisotachis in Balantiopsidaceae is only preliminary (cf.
*** Acroscyphella N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (
*** Acroscyphella iwatsukii (N.Kitag.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (
*** Acroscyphella phoenicorhiza (Grolle) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (
*** Acroscyphella tjiwideiensis (Sande Lac.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (
** Pseudoisotachis Váňa, Polish Bot. J. 58 (1): 55, 2013 (
** Pseudoisotachis pocsii Váňa, Polish Bot. J. 58 (1): 55, 2013 (
*** Balantiopsis Mitt., Handb. N. Zeal. fl. 2: 751, 1867 (
*** Balantiopsis asymmetrica (Herzog) J.J.Engel, Nova Hedwigia 16: 93, 1968 (
*** Balantiopsis bisbifida (Steph.) Steph., Sp. Hepat. (Stephani) 4: 101, 1910 (
*** Balantiopsis brasiliensis Steph., Sp. Hepat. (Stephani) 4: 104, 1910 (
*** Balantiopsis cancellata (Nees) Steph., Sp. Hepat. (Stephani) 4: 103, 1910 (
*** Balantiopsis ciliaris S.Hatt., J. Jap. Bot. 41 (5): 129, 1966 (
** Balantiopsis ciliaris subsp. novoguineensis S.Hatt., J. Jap. Bot. 41 (5): 131, 1966 (
*** Balantiopsis convexiuscula Berggr., New Zealand Hepat.: 44, 1898 (
*** Balantiopsis crocea Herzog, Beih. Bot. Centralbl. 60B (1/2): 12, 1939 (
*** Balantiopsis diplophylla (Hook.f. et Taylor) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
** Balantiopsis diplophylla var. hockenii (Berggr.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 11, 1997 (
*** Balantiopsis erinacea (Hook.f. et Taylor) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
*** Balantiopsis lingulata R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 26, 1968 (
*** Balantiopsis montana (Colenso) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 12, 1997 (
** Balantiopsis neocaledonica Pearson, J. Linn. Soc., Bot. 46 (305): 28, 1922 (
** Balantiopsis paucidens Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 29, 1901 (
*** Balantiopsis purpurata Mitt., Rep. Challenger, Bot. 1 (3, 1): 86, 1884 (
*** Balantiopsis rosea Berggr., New Zealand Hepat.: 43, 1898 (
*** Balantiopsis splendens (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 55, 1997 (
*** Balantiopsis tumida Berggr., New Zealand Hepat.: 45, 1898 (
*** Balantiopsis verrucosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 16, 1997 (
*** Isotachis Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 148, 1854 (
*** Isotachis armata (Nees) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (
*** Isotachis aubertii (Schwägr.) Mitt., J. Linn. Soc., Bot. 22 (146): 322, 1886 (
* Isotachis boliviensis Gottsche, Sp. Hepat. (Stephani) 3: 670, 1909 (
** Isotachis chinensis C.Gao, T.Cao et J.Sun, Bryologist 105 (4): 694, 2002 [2003] (
** Isotachis erythrorhiza (Lehm. et Lindenb.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 338, 1885 (
*** Isotachis fragilis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 67, 1911 (
** Isotachis grandis Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1041, 1888 (
*** Isotachis grossidens Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 69, 1911 (
** Isotachis hastatistipula (Steph.) J.J.Engel, Phytotaxa 183 (4): 299, 2014 (
* Isotachis hians Steph., Sp. Hepat. (Stephani) 3: 665, 1909 (
*** Isotachis humectata (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 3: 654, 1909 (
** Isotachis inflata Steph., Arch. Mus. Nac. Rio de Janeiro 13: 113, 1905 (
*** Isotachis intortifolia (Hook.f. et Taylor) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (
** Isotachis japonica Steph., Sp. Hepat. (Stephani) 3: 652, 1909 (
* Isotachis lacustris Herzog, Hedwigia 74 (2): 94, 1934 (
*** Isotachis lopezii (R.M.Schust.) Gradst., Mem. New York Bot. Gard. 84: 66, 1999 (
*** Isotachis lyallii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 149, 1854 (
*** Isotachis minima Pearson, Univ. Calif. Publ. Bot. 10 (4): 322, 1923 (
*** Isotachis montana Colenso, Trans. & Proc. New Zealand Inst. 21: 68, 1889 (
*** Isotachis multiceps (Lindenb. et Gottsche) Gottsche, Mexik. Leverm.: 105, 1863 (
** Isotachis multiceps var. fendleri Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 125, 1864 (
** Isotachis nigella Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 107, 1952 (
*** Isotachis obtusa Steph., Sp. Hepat. (Stephani) 6: 354, 1922 (
** Isotachis olivacea R.M.Schust., J. Hattori Bot. Lab. 83: 207, 1997 (
*** Isotachis plicata J.J.Engel, J. Hattori Bot. Lab. 83: 210, 1997 (
** Isotachis pusilla Steph., Sp. Hepat. (Stephani) 3: 655, 1909 (
** Isotachis riparia Rodway, Tasm. Bryoph.: 63, 1917 (
*** Isotachis serrulata (Sw.) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (
*** Isotachis spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 220, 1885 (
* Isotachis sprucei Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924 (
*** Isotachis stephanii E.S.Salmon, Rev. Bryol. 28 (4): 75, 1901 (
* Isotachis vexans Steph., Sp. Hepat. (Stephani) 3: 662, 1909 (
*** Isotachis westlandica (E.A.Hodgs.) R.M.Schust., Nova Hedwigia 15: 455, 1968 (
*** Neesioscyphus Grolle, Österr. Bot. Z. 111 (1): 19, 1964 (
*** Neesioscyphus allionii (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 185, 1966 (
*** Neesioscyphus argillaceus (Nees) Grolle, Österr. Bot. Z. 111 (1): 24, 1964 (
*** Neesioscyphus bicuspidatus (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 182, 1966 (
*** Neesioscyphus carneus (Nees) Grolle, Österr. Bot. Z. 111 (1): 20, 1964 (
*** Neesioscyphus homophyllus (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 188, 1964 (
*** Ruizanthus R.M.Schust., Phytologia 39 (4): 240, 1978 (
*** Ruizanthus venezuelanus R.M.Schust., Phytologia 39 (4): 240, 1978 (
by J.J. Engel
*** Blepharidophyllum Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 151, 1873 (
*** Blepharidophyllum densifolium (Hook.) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885 (
*** Blepharidophyllum vertebrale (Gottsche) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885 (
*** Clandarium (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (
*** Clandarium clandestinum (Mont.) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (
*** Clandarium gottscheanum (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (
*** Clandarium xiphophyllum (Grolle) R.M.Schust., Phytologia 56 (2): 68, 1984 (
by M.A.M. Renner
Calypogeiaceae is shown to be monophyletic (
*** Calypogeia Raddi, Jungermanniogr. Etrusca: 31, 1818 (
** subg. Asperifoliae (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 115, 1969 (
*** Calypogeia arguta Nees et Mont., Naturgesch. Eur. Leberm. 3: 24, 1838 (
** Calypogeia sullivantii Austin, Hepat. bor.-amer.: 19, 1873 (
** subg. Calypogeia
*** Calypogeia andicola Bischl., Candollea 18: 79, 1962 (
** Calypogeia annabonensis Steph., Sp. Hepat. (Stephani) 6: 447, 1924 (
** Calypogeia azorica Bischl., Rev. Bryol. Lichénol. 37 (1): 116, 1970 (
*** Calypogeia azurea Stotler et Crotz, Taxon 32 (1): 74, 1983 (
*** Calypogeia bidentula (F.Weber) Nees, Syn. Hepat. 2: 199, 1845 (
** Calypogeia falcata Bischl., Candollea 18: 112, 1962 (
*** Calypogeia fissa (L.) Raddi, Jungermanniogr. Etrusca: 33, 1818 (
** Calypogeia fissa subsp. neogaea R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 169, 1969 (
** Calypogeia goebelii (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908 (
** Calypogeia goebelii var. siamensis N.Kitag., Beih. Nova Hedwigia 90: 165, 1988 (
*** Calypogeia grandistipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908 (
*** Calypogeia integristipula Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 662 (394), 1908 (
*** Calypogeia laxa Gottsche et Lindenb., Syn. Hepat. 5: 713, 1847 (
*** Calypogeia lechleri (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908 (
** Calypogeia lechleri var. densifolia (Steph.) Bischl., Candollea 18: 101, 1962 (
** Calypogeia longifolia Steph., Sp. Hepat. (Stephani) 6: 449, 1924 (
** Calypogeia lophocoleoides Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908 (
** Calypogeia mascarenensis Bischl., Rev. Bryol. Lichénol. 37 (1): 89, 1970 (
** Calypogeia microstipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 670 (402), 1908 (
*** Calypogeia miquelii Mont. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 200, 1845 (
*** Calypogeia muelleriana (Schiffn.) Müll.Frib., Beih. Bot. Centralbl. 10 (4/5): 217, 1901 (
** Calypogeia muelleriana subsp. blomquistii R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 187, 1969 (
*** Calypogeia neesiana (C.Massal. et Carestia) Müll.Frib., Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905 (
** Calypogeia neesiana subsp. subalpina (Inoue) Inoue, Mem. Natl. Sci. Mus. (Tokyo) 4: 58, 1971 (
*** Calypogeia oblata Herzog, Svensk Bot. Tidskr. 51 (1): 189, 1957 (
*** Calypogeia peruviana Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838 (
*** Calypogeia rhombifolia (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 667 (399), 1908 (
** Calypogeia rhombifolia var. colombiana Bischl., Candollea 18: 104, 1962 (
*** Calypogeia sphagnicola (Arnell et J.Perss.) Warnst. et Loeske, Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905 (
*** Calypogeia subintegra (Gottsche, Lindenb. et Nees) Bischl., Candollea 18: 75, 1962 (
** Calypogeia subintegra var. dussiana (Steph.) Bischl., Candollea 18: 77, 1962 (
*** Calypogeia suecica (Arnell et J.Perss.) Müll.Frib., Beih. Bot. Centralbl. 17 (2): 224, 1904 (
*** Calypogeia tenax (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 664 (396), 1908 (
** Calypogeia uncinulatula Herzog, Hedwigia 67 (6): 250, 1927 (
Incertae sedis
*** Calypogeia aeruginosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (
* Calypogeia amazonica (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908 (
** Calypogeia angusta Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 663 (395), 1908 (
** Calypogeia apiculata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 668 (400), 1908 (
** Calypogeia asakawana S.Hatt. ex Inoue, J. Jap. Bot. 39 (4): 107, 1964 (
** Calypogeia ceylanica S.Hatt. et Mizut., Candollea 23: 288, 1968 (
** Calypogeia contracta Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 139, 1975 (
** Calypogeia cuspidata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908 (
* Calypogeia decurrens (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 675 (407), 1908 (
** Calypogeia formosana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 186, 1934 (
** Calypogeia fujisana Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 135, 1975 (
** Calypogeia granulata Inoue, J. Jap. Bot. 43 (10/11): 468, 1968 (
** Calypogeia japonica Steph., Sp. Hepat. (Stephani) 6: 448, 1924 (
** Calypogeia khasiana Ajit P.Singh et V.Nath, Taiwania 52 (4): 320, 2007 (
** Calypogeia latissima Steph., Sp. Hepat. (Stephani) 6: 449, 1924 (
*** Calypogeia lunata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (
** Calypogeia marginella Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 106, 1860 [1861] (
** Calypogeia obovata R.M.Schust., Phytologia 39 (4): 242, 1978 (
* Calypogeia steyermarkii Fulford, Mem. New York Bot. Gard. 11 (3): 305, 1968 (
** Calypogeia tosana (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 678 (410), 1908 (
** Calypogeia udarii Sudipa Das et D.K.Singh, Nelumbo 53: 194, 2011 (
** Eocalypogeia (R.M.Schust.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (
** Eocalypogeia quelpaertensis (S.Hatt. et Inoue) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (
** Eocalypogeia schusterana (S.Hatt. et Mizut.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (
** Metacalypogeia (S.Hatt.) Inoue, J. Hattori Bot. Lab. 21: 231, 1959 (
*** Metacalypogeia alternifolia (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 185, 1964 (
** Metacalypogeia cordifolia (Steph.) Inoue, J. Hattori Bot. Lab. 21: 233, 1959 (
** Mizutania Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989 (
*** Mizutania riccardioides Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989 (
*** Mnioloma Herzog, Ann. Bryol. 3: 115, 1930 (
** subg. Caracoma (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 833, 1995 (
** Mnioloma bolivianum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000 (
*** Mnioloma caespitosum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (
*** Mnioloma cellulosum (Spreng.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 836, 1995 (
*** Mnioloma crenulatum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (
*** Mnioloma cyclostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 843, 1995 (
** Mnioloma elliottii (Steph.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 841, 1995 (
** Mnioloma fissistipulum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (
*** Mnioloma fuscum (Lehm.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 848, 1995 (
*** Mnioloma nephrostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (
** Mnioloma novaezelandiae J.J.Engel, Cryptog. Bryol. 27 (1): 111, 2006 (
*** Mnioloma parallelogramum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (
** Mnioloma retusum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (
** Mnioloma stamatotonum M.A.M.Renner et E.A.Br., Fieldiana, Bot. (n.ser.) 47: 173, 2008 (
*** Mnioloma venezuelanum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000 (
** subg. Mnioloma, Fragm. Florist. Geobot. 40 (2): 833, 1995 (
*** Mnioloma rhynchophyllum Herzog, Ann. Bryol. 3: 120, 1930 (
by N.A. Konstantinova
*** Endogemma Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011 (
*** Endogemma caespiticia (Lindenb.) Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011 (
Placement of Geocalycaceae in Jungermanniinae follows
*** Geocalyx Nees, Naturgesch. Eur. Leberm. 1: 97, 1833 (
*** Geocalyx caledonicus Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 205 (265), 1908 (
*** Geocalyx graveolens (Schrad.) Nees, Naturgesch. Eur. Leberm. 2: 397, 1836 (
** Geocalyx lancistipulus (Steph.) S.Hatt., J. Jap. Bot. 28 (8): 234, 1953 (
** Geocalyx orientalis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxix, 1889 [1890] (
by J. Váňa
The treatment of the family follows
*** Acrolophozia R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 259, 1966 (
*** Acrolophozia fuegiana R.M.Schust., Nova Hedwigia 15: 499, 1968 (
*** Acrolophozia pectinata R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 261, 1966 (
*** Acrolophozia sulcata Hässel, J. Bryol. 11 (1): 108, 1980 (
** Nanomarsupella R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013 (
*** Nanomarsupella xenophylla (R.M.Schust.) R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013 (
** Paramomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 134, 1996 (
*** Paramomitrion paradoxum R.M.Schust., J. Hattori Bot. Lab. 80: 135, 1996 (
** Cryptocoleopsis Amakawa, J. Hattori Bot. Lab. 21: 274, 1959 (
*** Cryptocoleopsis imbricata Amakawa, J. Hattori Bot. Lab. 21: 274, 1959 (
*** Gymnomitrion Corda, Gen. hepat.: 651, 1829 (
*** Gymnomitrion adustum Nees, Naturgesch. Eur. Leberm. 1: 120, 1833 (
*** Gymnomitrion africanum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943 (
*** Gymnomitrion alpinum (Gottsche ex Husn.) Schiffn., Österr. Bot. Z. 53 (7): 280, 1903 (
*** Gymnomitrion asperulatum R.M.Schust., Acta Acad. Ped. Agr., Sect. Biol. 24: 114, 2003 (
*** Gymnomitrion atrofilum Váňa, J. Hattori Bot. Lab. 41: 411, 1976 (
*** Gymnomitrion bolivianum (Steph.) Váňa, Novon 20 (2): 225, 2010 (
*** Gymnomitrion brevissimum (Dumort.) Warnst., Hedwigia 53 (3): 196, 1913 (
*** Gymnomitrion commutatum (Limpr.) Schiffn., Magyar Bot. Lapok 13: 304, 1914 [1915] (
*** Gymnomitrion concinnatum (Lightf.) Corda, Gen. hepat.: 651, 1829 (
*** Gymnomitrion corallioides Nees, Naturgesch. Eur. Leberm. 1: 118, 1833 (
*** Gymnomitrion crenatilobum Grolle, Khumbu Himal 1 (4): 278, 1966 (
*** Gymnomitrion crenulatum Gottsche ex Carrington, Trans. Bot. Soc. Edinburgh 7 (3): 444, 1863 (
*** Gymnomitrion crystallocaulon (Grolle) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
** Gymnomitrion incompletum (Gottsche) R.M.Schust. ex Váňa, J. Hattori Bot. Lab. 40: 186, 1976 (
*** Gymnomitrion laceratum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943 (
*** Gymnomitrion miniatum Lindenb. et Gottsche, Syn. Hepat. 4: 617, 1846 (
*** Gymnomitrion minutulum (Hässel) Váňa, Novon 20 (2): 225, 2010 (
** Gymnomitrion moralesae Váňa, J. Hattori Bot. Lab. 48: 230, 1980 (
*** Gymnomitrion mucronulatum (N.Kitag.) N.Kitag., Acta Phytotax. Geobot. 19 (2/3): 53, 1962 (
** Gymnomitrion mucrophorum R.M.Schust., Bryologist 98 (2): 243, 1995 (
*** Gymnomitrion nigrum (Grolle et Váňa) Váňa, Novon 20 (2): 225, 2010 (
*** Gymnomitrion noguchianum S.Hatt., J. Jap. Bot. 27 (2): 55, 1952 (
*** Gymnomitrion obtusilobum N.Kitag., Bull. Univ. Mus. Univ. Tokyo 8: 229, 1975 (
*** Gymnomitrion obtusum Lindb., Morgonbladet (Helsinki) 1877 (30, 6 Feb): 2, 1877 (
*** Gymnomitrion pacificum Grolle, Trans. Brit. Bryol. Soc. 5 (1): 92, 1966 (
*** Gymnomitrion revolutum (Nees) H.Philib., Rev. Bryol. 17 (3): 34, 1890 (
** Gymnomitrion revolutum subsp. novoguineanensis (R.M.Schust.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
*** Gymnomitrion rubidum (Mitt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
*** Gymnomitrion setaceum Grolle et Váňa, J. Hattori Bot. Lab. 41: 411, 1976 (
*** Gymnomitrion sinense Müll.Frib., Rev. Bryol. Lichénol. 20 (1/2): 176, 1951 (
** Gymnomitrion strictum (Berggr.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (
** Gymnomitrion strictum var. inaequale R.M.Schust., J. Hattori Bot. Lab. 80: 118, 1996 (
*** Gymnomitrion subintegrum (S.W.Arnell) Váňa, Novon 20 (2): 225, 2010 (
*** Gymnomitrion truncatoapiculatum Herzog, Hedwigia 74 (2): 81, 1934 (
*** Gymnomitrion verrucosum W.E.Nicholson, Symb. Sin. 5: 10, 1930 (
*** Marsupella Dumort., Commentat. Bot. (Dumortier): 114, 1822 (
*** Marsupella alata S.Hatt. et N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 79, 1960 (
*** Marsupella andreaeoides (Lindb.) Müll.Frib., Feddes Repert. Spec. Nov. Regni Veg. 54 (2/3): 214, 1951 (
*** Marsupella apiculata Schiffn., Österr. Bot. Z. 53 (6): 249, 1903 (
*** Marsupella aquatica (Lindenb.) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 44 (8): 267, 1896 [1897] (
*** Marsupella arctica (Berggr.) Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 26, 1906 (
*** Marsupella boeckii (Austin) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 409, 1893 (
*** Marsupella bolanderi (Austin) Underw., Zoe 1 (12): 365, 1891 (
*** Marsupella condensata (Ångstr. ex C.Hartm.) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 420, 1893 (
*** Marsupella disticha Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 164 (25), 1901 (
*** Marsupella emarginata (Ehrh.) Dumort., Recueil Observ. Jungerm.: 24, 1835 (
** Marsupella emarginata subsp. tubulosa (Steph.) N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 76, 1960 (
** Marsupella emarginata subsp. tubulosa var. apertifolia (Steph.) N.Kitag., J. Hattori Bot. Lab. 26: 89, 1963 (
** Marsupella emarginata subsp. tubulosa var. patens N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 77, 1960 (
** Marsupella emarginata subsp. tubulosa var. tubulosa (Steph.) N.Kitag. ex Váňa et L.Söderstr., Phytotaxa 183 (4): 288, 2014 (
*** Marsupella funckii (F.Weber et D.Mohr) Dumort., Recueil Observ. Jungerm.: 24, 1835 (
*** Marsupella microphylla R.M.Schust., Phytologia 39 (4): 249, 1978 (
** Marsupella minutissima N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 81, 1960 (
*** Marsupella neesii Sande Lac. ex Schiffn., Consp. Hepat. Arch. Ind.: 70, 1898 (
*** Marsupella paroica R.M.Schust., Bryologist 60 (2): 145, 1957 (
*** Marsupella profunda Lindb., Rev. Bryol. 14 (2): 19, 1887 (
*** Marsupella pseudofunckii S.Hatt., J. Hattori Bot. Lab. 4: 63, 1950 (
*** Marsupella sparsifolia (Lindb.) Dumort., Bull. Soc. Roy. Bot. Belgique 13: 128, 1874 (
** Marsupella sparsifolia subsp. childii R.M.Schust., Phytotaxa 183 (4): 288, 2014 (
*** Marsupella sphacelata (Giesecke ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 24, 1835 (
*** Marsupella spiniloba R.M.Schust. et Damsh., Phytologia 63 (5): 326, 1987 (
*** Marsupella sprucei (Limpr.) Bernet, Cat. hép. Suisse: 33, 1888 (
** Marsupella stableri Spruce, Rev. Bryol. 8 (6): 96, 1881 (
*** Marsupella stoloniformis N.Kitag., J. Hattori Bot. Lab. 30: 201, 1967 (
** Marsupella stoloniformis subsp. vermiformis R.M.Schust., J. Hattori Bot. Lab. 80: 72, 1996 (
*** Marsupella yakushimensis (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 80, 1944 (
*** Poeltia Grolle, Khumbu Himal 1 (4): 280, 1966 (
*** Poeltia campylata Grolle, Khumbu Himal 1 (4): 280, 1966 (
*** Prasanthus Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889 (
** Prasanthus jamalicus Potemkin, Ann. Bot. Fenn. 29 (4): 319, 1992 (
*** Prasanthus suecicus (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889 (
*** Nardia Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (
*** Nardia arnelliana Grolle, Bot. Mag. (Tokyo) 77 (914): 297, 1964 (
*** Nardia assamica (Mitt.) Amakawa, J. Hattori Bot. Lab. 26: 23, 1963 (
*** Nardia breidleri (Limpr.) Lindb., Helsingf. Dagbl. 1880 (311, 15 Nov.): 2, 1880 (
*** Nardia compressa (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (
*** Nardia flagelliformis Inoue, J. Jap. Bot. 46 (1): 1, 1971 (
*** Nardia geoscyphus (De Not.) Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 371, 1874 (
* Nardia geoscyphus var. dioica Bakalin, Arctoa 18: 87, 2009 [2010] (
* Nardia geoscyphus var. suberecta (Lindb. ex Kaal.) Váňa, Phytotaxa 76 (3): 37, 2013 (
*** Nardia grollei Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 491, 2009 (
*** Nardia insecta Lindb., Helsingf. Dagbl. 1878 (315, 18 Nov.): 2, 1878 (
*** Nardia japonica Steph., Bull. Herb. Boissier 5 (2): 101, 1897 (
* Nardia kamtschatica Arnell et C.E.O.Jensen, Hedwigia 67 (1/2): 111, 1927 (
* Nardia leptocaulis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981 (
*** Nardia lescurii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 115, 1884 (
** Nardia minutifolia Furuki, Bryol. Res. 9 (3): 73, 2006 (
** Nardia nuda (Lindenb. et Gottsche) Váňa, Folia Geobot. Phytotax. 8 (2): 193, 1973 (
*** Nardia poeltii Váňa, J. Hattori Bot. Lab. 36: 73, 1972 [1973] (
*** Nardia scalaris Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (
* Nardia scalaris var. botryoidea (R.M.Schust.) Váňa, Phytotaxa 76 (3): 38, 2013 (
* Nardia scalaris var. harae (Amakawa) Váňa, Phytotaxa 76 (3): 38, 2013 (
*** Nardia subclavata (Steph.) Amakawa, J. Jap. Bot. 32 (2): 40, 1957 (
*** Nardia succulenta (A.Rich.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 519, 1885 (
*** Nardia unispiralis Amakawa, J. Jap. Bot. 32 (6): 167, 1957 (
Placement of Gyrothyraceae in Jungermanniinae follows
** Gyrothyra M.Howe, Bull. Torrey Bot. Club 24 (4): 201, 1897 (
*** Gyrothyra underwoodiana M.Howe, Bull. Torrey Bot. Club 24 (4): 202, 1897 (
*** Harpanthus Nees, Naturgesch. Eur. Leberm. 2: 351, 1836 (
*** Harpanthus drummondii (Taylor) Grolle, Österr. Bot. Z. 112 (3): 274, 1965 (
*** Harpanthus flotovianus (Nees) Nees, Naturgesch. Eur. Leberm. 2: 353, 1836 (
*** Harpanthus scutatus (F.Weber et D.Mohr) Spruce, Trans. Bot. Soc. Edinburgh 3 (1/4): 209, 1850 (
by N. Konstantinova
The family Hygrobiellaceae was validated by
*** Hygrobiella Spruce, Cephalozia: 73, 1882 (
** Hygrobiella intermedia Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014 (
*** Hygrobiella laxifolia (Hook.) Spruce, Cephalozia: 74, 1882 (
** Hygrobiella squamosa Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014 (
by J. Váňa
The monogeneric status of Jackiella is supported by a molecular study based on three loci by
*** Jackiella Schiffn., Hep. Fl. Buitenzorg: 211, 1900 (
** Jackiella angustifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 296, 1950 (
** Jackiella ceylanica Schiffn. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 212 (272), 1908 (
*** Jackiella curvata E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand 85 (4): 571, 1958 (
*** Jackiella javanica Schiffn., Hep. Fl. Buitenzorg: 212, 1900 (
** Jackiella javanica var. cavifolia Schiffn., Hep. Fl. Buitenzorg: 213, 1900 (
** Jackiella javanica var. cordifolia Schiffn., Hep. Fl. Buitenzorg: 213, 1900 (
** Jackiella renifolia Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901] (
** Jackiella sinensis (W.E.Nicholson) Grolle, Österr. Bot. Z. 111 (2/3): 186, 1964 (
** Jackiella singapurensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901] (
** Jackiella singapurensis var. philippinensis N.Kitag., Misc. Bryol. Lichenol. 9 (1): 9, 1981 (
Excluded from the genus
* Jackiella unica Steph., Sp. Hepat. (Stephani) 6: 318, 1922 (
by J. Váňa
The treatment of Jungermanniaceae follows
** Delavayella Steph., Hedwigia 33 (1): 4, 1894 (
** Delavayella serrata Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 211, 1894 (
** Delavayella serrata var. purpurea P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 38, 1955 (
** Liochlaena Nees, Syn. Hepat. 2: 150, 1845 (
*** Liochlaena lanceolata Nees, Syn. Hepat. 2: 150, 1845 (
*** Liochlaena subulata (A.Evans) Schljakov, Pečen. Mchi Sev. SSSR 4: 71, 1981 (
** Eremonotus Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 200, 1900 (
*** Eremonotus myriocarpus (Carrington) Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 201, 1900 (
*** Jungermannia L., Sp. Pl. 1: 1131, 1753 (
*** Jungermannia atrovirens Dumort., Syll. Jungerm. Europ.: 51, 1831 (
*** Jungermannia borealis Damsh. et Váňa, Lindbergia 4 (1/2): 5, 1977 (
* Jungermannia erectii Ajit P.Singh et V.Nath, Hepat. Khasi Jaintia Hills: E. Himal.: 117, 2007 (
*** Jungermannia exsertifolia Steph., Sp. Hepat. (Stephani) 6: 86, 1917 (
** Jungermannia exsertifolia subsp. cordifolia (Dumort.) Váňa, Folia Geobot. Phytotax. 8 (3): 268, 1973 (
*** Jungermannia gollanii Steph., Sp. Hepat. (Stephani) 6: 86, 1917 (
** Jungermannia konstantinovae Bakalin et Vilnet, Arctoa 18: 161, 2009 [2010] (
*** Jungermannia ovatotrigona (Steph.) Grolle, Feddes Repert. 82 (1): 90, 1971 (
*** Jungermannia polaris Lindb., Öfvers. Kongl. Vetensk.-Akad. Förh. 23 (10): 560, 1866 [1867] (
*** Jungermannia pumila With., Arr. Brit. Pl., ed. 3, 3: 883, 1796 (
Taxa doubtfully belonging to the genus
* Jungermannia amentacea Bertol., Mem. Reale Accad. Sci. Ist. Bologna (ser. 2) 1: 19, 1862 (
* Jungermannia brasiliensis Raddi, Critt. Brasil.: 15, 1822 (
* Jungermannia chinensis Osbeck, Dagb. Ostind. Resa: 221, 1757 (
* Jungermannia cordata Vill., Hist. Pl. Dauphiné (Villars) 3: 923, 1789 (
* Jungermannia crenulata Schmidel, Jungerm. Char.: 20, 1760 (
* Jungermannia creutzeri Kremer, Monogr. hépat. Moselle: 26, 1837 (
* Jungermannia digitata C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 326, 1827 (
* Jungermannia dubioides H.A.Mill., Phytologia 47 (4): 322, 1981 (
* Jungermannia fernandeziana Mitt., Rep. Challenger, Bot. 1 (3, 1): 85, 1884 (
* Jungermannia hexagona Schwägr., Hist. Musc. Hepat. Prodr.: 18, 1814 (
* Jungermannia holandriana Kremer, Monogr. hépat. Moselle: 25, 1837 (
* Jungermannia incerta Gottsche, Abh. Naturwiss. Vereins Bremen 7: 344, 1882 (
* Jungermannia lateriflora Hampe ex Gottsche, Mexik. Leverm.: 82, 1863 (
* Jungermannia lescuriana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 67, 1866 (
* Jungermannia longiretis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxv, 1889 [1890] (
* Jungermannia mastigophora Spreng. Neue Entdeck. Pflanzenk. 2: 99, 1821 (
* Jungermannia michelii Mérat, Nouv. fl. env. Paris (ed. 2) 1: 219, 1821 (
* Jungermannia minima Scop., Fl. Carniol. (ed. 2) 2: 350, 1772 (
* Jungermannia odorata With., Bot. arr. veg. Gr. Brit. 2: 693, 1776 (
* Jungermannia peltata Schmidel, Jungerm. Char.: 14, 1760 (
* Jungermannia quadridigitata Griff., Not. pl. asiat. 2: 314, 1849 (
* Jungermannia sauteri De Not. ex Rabenh., Hedwigia 1 (20): 121, 1857 (
* Jungermannia secunda Hampe ex Gottsche, Mexik. Leverm.: 82, 1863 (
* Jungermannia stereocaulis Bory, Voy. Uranie, Bot. 4: 130, 1827 (
* Jungermannia submersa Kremer, Monogr. hépat. Moselle: 36, 1837 (
* Jungermannia sullivantiana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 66, 1866 (
* Jungermannia supina Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 86, 1795 [1796] (
* Jungermannia tenuis Ehrh., Beitr. Naturk. (Ehrhart) 4: 45, 1789 (
* Jungermannia uncifolia Steph., Hedwigia 34 (2): 51, 1895 (
* Jungermannia vernicosa Cass. ex Mérat, Nouv. fl. env. Paris (ed. 2) 1: 221, 1821 (
*** Mesoptychia (Lindb.) A.Evans, Ottawa Naturalist 17: 15, 1903 (
*** Mesoptychia badensis (Gottsche ex Rabenh.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (
*** Mesoptychia bantriensis (Hook.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (
** Mesoptychia bantriensis subsp. wallfischii (Ştefănuţ) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (
* Mesoptychia chichibuensis (Inoue) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
* Mesoptychia collaris (Nees) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
* Mesoptychia fitzgeraldiae (Paton et A.R.Perry) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
*** Mesoptychia gillmanii (Austin) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
*** Mesoptychia heterocolpos (Thed. ex Hartm.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
** Mesoptychia heterocolpos var. arctica (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
** Mesoptychia heterocolpos var. harpanthoides (Bryhn et Kaal.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
*** Mesoptychia igiana (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
* Mesoptychia mamatkulovii (Duda) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
*** Mesoptychia mayebarae (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
*** Mesoptychia morrisoncola (Horik.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
* Mesoptychia polymorpha Stotler, Crand.-Stotl. et Bakalin, Polish Bot. J. 58 (1): 82, 2013 (
*** Mesoptychia rutheana (Limpr.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
** Mesoptychia rutheana var. laxa (Schiffn. ex Burrell) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
*** Mesoptychia sahlbergii (Lindb. et Arnell) A.Evans, Ottawa Naturalist 17: 15, 1903 (
*** Mesoptychia subcrispa (Herzog) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
*** Mesoptychia turbinata (Raddi) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012 (
* Mesoptychia ussuriensis (Bakalin) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012 (
** Rivulariella D.H.Wagner, Phytoneuron 2013 (10): 2, 2013 (
** Rivulariella gemmipara (A.Evans) D.H.Wagner, Phytoneuron 2013 (10): 2, 2013 (
*** Notoscyphus Mitt., Fl. vit.: 407, 1871 [1873] (
*** Notoscyphus lutescens (Lehm. et Lindenb.) Mitt., Fl. vit.: 407, 1871 [1873] (
*** Saccogyna Dumort., Commentat. Bot. (Dumortier): 113, 1822 (
* Saccogyna ligulata Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 207 (267), 1908 (
** Saccogyna subacuta Steph., Sp. Hepat. (Stephani) 6: 316, 1922 (
*** Saccogyna viticulosa (L.) Dumort., Syll. Jungerm. Europ.: 74, 1831 (
Excluded from the genus
* Saccogyna tridens Steph., Sp. Hepat. (Stephani) 6: 317, 1922 (
by J. Váňa
Solenostomataceae was first defined by
** Aponardia (R.M.Schust.) Váňa, Phytotaxa 65: 46, 2012 (
*** Aponardia huerlimannii (Váňa et Grolle) Váňa, Phytotaxa 65: 46, 2012 (
** Arctoscyphus Hässel, Lindbergia 16 (4): 133, 1990 [1992] (
** Arctoscyphus fuegiensis (C.Massal.) Hässel, Cryptog. Bryol. Lichénol. 17 (3): 164, 1996 (
** Arctoscyphus ronsmithii Hässel, Lindbergia 16 (4): 133, 1990 [1992] (
** Cryptocolea R.M.Schust., Amer. Midl. Naturalist 49 (2): 414, 1953 (
*** Cryptocolea imbricata R.M.Schust., Amer. Midl. Naturalist 49 (2): 417, 1953 (
** Diplocolea Amakawa, J. Jap. Bot. 37 (9): 274, 1962 (
*** Diplocolea sikkimensis Amakawa, J. Jap. Bot. 37 (9): 274, 1962 (
*** Solenostoma Mitt., J. Proc. Linn. Soc., Bot. 8 (29): 51, 1864 [1865] (
** subg. Eucalyx (Lindb.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015 (
*** Solenostoma bilobum (S.Hatt. ex Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 286, 2009 (
*** Solenostoma emarginatum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma flagellatum (S.Hatt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma hokkaidense (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma obovatum (Nees) C.Massal., Epat. erb. critt. ital.: 17, 1903 (
*** Solenostoma obscurum (A.Evans) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1013, 1969 (
*** Solenostoma schusteranum (J.D.Godfrey et G.Godfrey) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma subtilissimum (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969 (
** subg. Metasolenostoma Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015 (
*** Solenostoma fusiforme (Steph.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 944, 1969 (
*** Solenostoma gracillimum (Sm.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 972, 1969 (
*** Solenostoma handelii (Schiffn.) Müll.Frib., Beitr. Kryptogamenfl. Schweiz 10 (2): 38, 1947 (
** Solenostoma lignicola (Schiffn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma limbatifolium (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (
** Solenostoma philippinense Váňa, Phytotaxa 152 (1): 43, 2013 (
*** Solenostoma rubrum (Gottsche) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 975, 1969 (
*** Solenostoma suborbiculatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (
** subg. Plectocolea Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 (
*** Solenostoma balfourii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma borneense (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma callithrix (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 486 (48), 1901 (
*** Solenostoma champawatense (S.N.Srivast. et Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
*** Solenostoma comatum (Nees) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 73, 1981 (
*** Solenostoma comatum var. novae-guineae (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma crenuliforme (Austin) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901 (
*** Solenostoma decolor (Schiffn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma dusenii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma erectum (Amakawa) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 66, 1981 (
* Solenostoma flagellalioides C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981 (
*** Solenostoma flavialbicans (Amakawa et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma fossombronioides (Austin) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969 (
*** Solenostoma glaucum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (
* Solenostoma gongshanense (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (
*** Solenostoma haskarlianum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (
*** Solenostoma hattorianum (Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 287, 2009 (
*** Solenostoma hirticalyx (Steph.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma horikawanum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma hyalinum (Lyell) Mitt., Nat. hist. Azores: 319, 1870 (
*** Solenostoma infuscum (Mitt.) Hentschel, Pl. Syst. Evol. 268 (1/4): 152, 2007 (
*** Solenostoma infuscum var. ovicalyx (Steph.) Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 288, 2009 (
** Solenostoma kurilense (Bakalin) Váňa, Phytotaxa 152 (1): 40, 2013 (
* Solenostoma lixingjiangii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (
*** Solenostoma marginatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969 (
*** Solenostoma micranthum (Mitt.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
** Solenostoma montanum (Steph.) Váňa, Phytotaxa 65: 44, 2012 (
* Solenostoma multicarpum (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
** Solenostoma nilgiriense (A.Alam, Ad.Kumar et S.C.Srivast.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma obliquifolium (Schiffn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma onraedtii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
* Solenostoma orbicularifolium (Piippo ex C.Gao et Bai) Váňa, Phytotaxa 222 (2): 199, 2015 (
*** Solenostoma otianum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 984, 1969 (
*** Solenostoma ovalifolium (Amakawa) Váňa, Phytotaxa 152 (1): 40, 2013 (
*** Solenostoma paroicum (Schiffn.) R.M.Schust., Amer. Midl. Naturalist 49 (2): 402, 1953 (
*** Solenostoma plagiochilaceum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma polyrhizoides (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma radicellosum Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 [1865] (
*** Solenostoma renauldii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma rigidulum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969 (
*** Solenostoma rosulans (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (
*** Solenostoma rotundatum Amakawa, J. Jap. Bot. 31 (2): 50, 1956 (
*** Solenostoma rubripunctatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 898, 1969 (
* Solenostoma rupicola (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (
*** Solenostoma sikkimense (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (
*** Solenostoma tetragonum (Lindenb.) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009 (
* Solenostoma tetragonum var. kodaikanalense A.Alam, D.Sharma et So.Yadav, Phytotaxonomy 12: 70, 2012 (
*** Solenostoma torticalyx (Steph.) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 69, 1981 (
*** Solenostoma truncatum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009 (
** Solenostoma truncatum var. setulosum (Herzog) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
*** Solenostoma tuberculiferum (Herzog) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (
*** Solenostoma unispire (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (
*** Solenostoma virgatum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
*** Solenostoma vulcanicola (Schiffn.) Nyushko ex Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 289, 2009 (
* Solenostoma zangmuii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
** subg. Solenostoma
*** Solenostoma amoenum (Lindenb. et Gottsche) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma amplexifolium (Hampe) Váňa et Schäf.-Verw., Acta Bot. Hung. 51 (3/4): 408, 2009 (
** Solenostoma appalachianum R.M.Schust. ex Bakalin, Arctoa 23: 127, 2014 (
*** Solenostoma appressifolium (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 494, 2009 (
*** Solenostoma appressifolium var. minor (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (
*** Solenostoma appressifolium var. nigricans (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (
*** Solenostoma ariadne (Taylor) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (
*** Solenostoma atrobrunneum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (
*** Solenostoma atrorevolutum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
*** Solenostoma atrovirens Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 493 (55), 1901 (
*** Solenostoma baueri (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 495 (57), 1901 (
*** Solenostoma bengalense (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
** Solenostoma breviflorum Kashyap et R.S.Chopra, Liverworts W. Himal. 2: 85, 1932 (
*** Solenostoma caeleste (Inoue et Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
* Solenostoma caoi (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
*** Solenostoma caucasicum (Váňa) Konstant., Arctoa 1: 123, 1992 (
*** Solenostoma chenianum (C.Gao, Y.H.Wu et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
*** Solenostoma clavellatum Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 491 (53), 1901 (
*** Solenostoma confertissimum (Nees) Schljakov, Novosti Sist. Nizš. Rast. 17: 239, 1980 (
*** Solenostoma coniflorum (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901 (
*** Solenostoma crassulum (Nees et Mont.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901 (
** Solenostoma cryptogynum R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (
*** Solenostoma cyclops (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969 (
*** Solenostoma diversiclavellatum (Amakawa et Grolle) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma dulongense Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 497, 2009 (
*** Solenostoma exsertum (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901 (
*** Solenostoma faurieanum (Beauverd) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969 (
*** Solenostoma flagellare (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma flavorevolutum (Váňa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma grollei (D.G.Long et Váňa) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (
*** Solenostoma grosseverrucosum (Amakawa et S.Hatt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
*** Solenostoma heterolimbatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma hewsoniae (Amakawa et Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma hiugaense Amakawa, J. Jap. Bot. 31 (2): 47, 1956 (
*** Solenostoma indrodayanum (Sushil K.Singh et D.K.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma inundatum (Hook.f. et Taylor) Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901 (
*** Solenostoma javanicum (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901 (
** Solenostoma kanaii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma kashyapii (S.C.Srivast., S.Srivast. et D.Sharma) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma lanigerum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma macrocarpum (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (
*** Solenostoma mamatkulovii (Váňa et Zerov) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
* Solenostoma microphyllum C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 206, 1981 (
* Solenostoma microrevolutum (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma mildbraedii (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 387, 2002 (
*** Solenostoma niveum (Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
* Solenostoma novazelandiae R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (
*** Solenostoma ohbae (Amakawa) C.Gao, Bryofl. Xizang: 495, 1985 (
*** Solenostoma orbiculatum (Colenso) R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (
*** Solenostoma parvitextum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma patoniae (Grolle, D.B.Schill et D.G.Long) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (
*** Solenostoma pocsii (Váňa) Bakalin, Arctoa 18: 160, 2009 [2010] (
*** Solenostoma poeltii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma pseudocyclops (Inoue) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009 (
** Solenostoma pseudopyriflorum Bakalin et Vilnet, Arctoa 18: 159, 2009 [2010] (
*** Solenostoma purpuratum (Mitt.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901 (
*** Solenostoma pyriflorum Steph., Sp. Hepat. (Stephani) 6: 83, 1917 (
*** Solenostoma pyriflorum var. gracillimum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009 (
** Solenostoma pyriflorum var. major (S.Hatt.) Bakalin, Arctoa 17: 230, 2008 [2009] (
*** Solenostoma pyriflorum var. minutissimum (Amakawa) Bakalin, Arctoa 16: 208, 2007 [2008] (
*** Solenostoma raujeanum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (
*** Solenostoma riclefii Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (
*** Solenostoma sanguinolentum (Griff.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901 (
*** Solenostoma schaulianum (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (
*** Solenostoma shimizuanum (S.Hatt. ex Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (
*** Solenostoma speciosum (Horik.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (
** Solenostoma speciosum subsp. villosum (R.M.Schust.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 246, 2009 (
*** Solenostoma sphaerocarpum (Hook.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 499 (61), 1901 (
*** Solenostoma stephanii (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 496 (58), 1901 (
*** Solenostoma stoloniferum (Steph.) S.W.Arnell, Hepat. South Africa: 316, 1963 (
*** Solenostoma strictum (Schiffn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (
*** Solenostoma subacutum (Herzog) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
*** Solenostoma subrubrum (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (
* Solenostoma sunii Bakalin et Vilnet, Bot. Pacifica 3 (2): 15, 2014 (
* Solenostoma totopapillosum (E.A.Hodgs.) R.M.Schust., Bryologist 100 (3): 366, 1997 (
*** Solenostoma udarii (S.C.Srivast. et P.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
* Solenostoma ventroversum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
*** Solenostoma zantenii (Amakawa) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
* Solenostoma zengii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
by J. Váňa
Southbyaceae has traditionally been placed in Arnelliaceae (e.g.
*** Gongylanthus Nees, Naturgesch. Eur. Leberm. 2: 405, 1836 (
*** Gongylanthus dusenii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906 (
*** Gongylanthus ericetorum (Raddi) Nees, Naturgesch. Eur. Leberm. 2: 407, 1836 (
*** Gongylanthus granatensis (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906 (
*** Gongylanthus himalayensis Grolle, Khumbu Himal 1 (4): 285, 1966 (
*** Gongylanthus javanicus Grolle, J. Jap. Bot. 40 (7): 206, 1965 (
*** Gongylanthus liebmanianus (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906 (
*** Gongylanthus limbatus (Herzog) Grolle et Váňa, Folia Geobot. Phytotax. 9 (2): 198, 1974 (
*** Gongylanthus muelleri (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906 (
** Gongylanthus oniscoides (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 386 (42), 1906 (
*** Gongylanthus richardsii E.W.Jones, Trans. Brit. Bryol. Soc. 4 (4): 650, 1964 (
*** Southbya Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849 (
*** Southbya gollanii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 381 (37), 1906 (
*** Southbya nigrella (De Not.) Henriq., Bol. Soc. Brot. 4: 244, 1886 [1887] (
*** Southbya organensis Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 53, 1950 (
*** Southbya tophacea (Spruce) Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849 (
by J. Váňa
In the absence of molecular data, the phylogenetic affinities of Stephaniella and Stephaniellidium remain equivocal, but there is no evidence to support their placement in either the Arnelliaceae or Southbyaceae where they have been placed recently. Thus the family as construed by
*** Stephaniella J.B.Jack, Hedwigia 33 (1): 11, 1894 (
* Stephaniella boliviensis Steph., Biblioth. Bot. 87 (2): 182, 1916 (
*** Stephaniella hamata Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1024 (87), 1901 (
*** Stephaniella paraphyllina J.B.Jack, Hedwigia 33 (1): 11, 1894 (
*** Stephaniella rostrata U.Schmitt, Österr. Bot. Z. 115 (2): 124, 1968 (
*** Stephaniella uncifolia S.Winkl., Österr. Bot. Z. 115 (2): 124, 1968 (
** Stephaniellidium S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983 (
*** Stephaniellidium sleumeri (Müll.Frib.) S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983 (
*** Trichotemnoma R.M.Schust., Nova Hedwigia 15: 440, 1968 (
*** Trichotemnoma corrugatum (Steph.) R.M.Schust., Nova Hedwigia 15: 440, 1968 (
The recognition of Blepharostomataceae follows
*** Blepharostoma (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
** Blepharostoma arachnoideum M.Howe, Mem. Torrey Bot. Club 7: 140, 1899 (
** Blepharostoma indicum G.Asthana, M.Saxena et Maurya, J. Bryol. 35 (4): 267, 2013 (
** Blepharostoma minor Horik., Hikobia 1 (2): 104, 1951 [1952] (
*** Blepharostoma trichophyllum (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
** Blepharostoma trichophyllum subsp. brevirete (Bryhn et Kaal.) R.M.Schust., Bull. Natl. Mus. Canada 164: 16, 1959 (
by L. Söderström, J. Váňa, R. Stotler, B.J. Crandall-Stotler and J.J. Engel
The treatment of Brevianthaceae follows
*** Brevianthus J.J.Engel et R.M.Schust., Phytologia 47 (4): 317, 1981 (
*** Brevianthus flavus (Grolle) J.J.Engel et R.M.Schust., Phytologia 47 (4): 318, 1981 (
** Brevianthus flavus subsp. crenulatus J.J.Engel, Nova Hedwigia 93 (3/4): 406, 2011 (
* Brevianthus hypocanthidium M.A.M.Renner et J.J.Engel, PhytoKeys 50: 46, 2015 (
*** Tetracymbaliella Grolle, Nova Hedwigia 3 (1): 48, 1961 (
** Tetracymbaliella comptonii (Pearson) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 164, 1963 [1964] (
*** Tetracymbaliella cymbalifera (Hook.f. et Taylor) Grolle, Nova Hedwigia 3 (1): 50, 1961 (
*** Tetracymbaliella decipiens (Gottsche) Grolle, Nova Hedwigia 3 (1): 49, 1961 (
*** Tetracymbaliella subsimplex (Austin) J.J.Engel, Phytotaxa 207 (2): 185, 2015 (
** Chonecolea Grolle, Rev. Bryol. Lichénol. 25 (3/4): 294, 1956 [1957] (
** Chonecolea acutiloba (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 321, 1980 (
** Chonecolea andina Grolle et Váňa, J. Hattori Bot. Lab. 48: 229, 1980 (
** Chonecolea doellingeri (Nees) Grolle, Rev. Bryol. Lichénol. 25 (3/4): 295, 1956 [1957] (
** Chonecolea ruwenzorensis E.W.Jones, J. Bryol. 13 (4): 498, 1986 (
** Chonecolea schusteri Udar et Ad.Kumar, Bryologist 85 (3): 315, 1982 (
** Chonecolea verae Potemkin, Proc. int. meeting 90th anniv. Abramova: 165, 2005 (
** Grollea R.M.Schust., Nova Hedwigia 8 (3/4): 288, 1964 (
** Grollea antheliopsis R.M.Schust., Nova Hedwigia 8 (3/4): 288, 1964 (
by D. Bell
*** Herbertus Gray, Nat. Arr. Brit. Pl. 1: 705, 1821 (
*** Herbertus aduncus (Dicks.) Gray, Nat. Arr. Brit. Pl. 1: 705, 1821 (
** Herbertus arcticus (Inoue et Steere) Schljakov, Novosti Sist. Nizš. Rast. 19: 209, 1982 (
** Herbertus armitanus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 324, 1965 (
** Herbertus asparus Tixier, Bryophyt. Biblioth. 18: 64, 1979 (
*** Herbertus bivittatus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 343, 1885 (
** Herbertus borealis Crundw., Trans. Brit. Bryol. Soc. 6 (1): 41, 1970 (
** Herbertus buchii Juslén, Ann. Bot. Fenn. 43 (6): 416, 2006 (
** Herbertus ceylanicus (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 43, 1959 (
** Herbertus circinatus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 318, 1965 (
*** Herbertus delavayi (Steph.) Steph., Hedwigia 34 (2): 43, 1895 (
*** Herbertus dicranus (Gottsche, Lindenb. et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (
** Herbertus durandii (Steph.) Herzog, Rev. Bryol. Lichénol. 11 (1): 25, 1938 [1939] (
* Herbertus evittatus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 327, 1965 (
** Herbertus gaochienii X.Fu, Fl. Bryoph. Sin. 9: 38, 2003 (
** Herbertus gracilis (Mont.) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (
** Herbertus guangdongii P.J.Lin ex Piippo, Bryobrothera 1: 206, 1992 (
** Herbertus hawaiiensis H.A.Mill., J. Hattori Bot. Lab. 28: 317, 1965 (
** Herbertus helleri (Steph.) W.E.Nicholson, Rev. Bryol. Lichénol. 13: 143, 1942 (
** Herbertus herpocladioides E.B.Scott et H.A.Mill., Bryologist 62 (2): 116, 1959 (
*** Herbertus hutchinsiae (Gottsche et Rabenh.) A.Evans, Bull. Torrey Bot. Club 44 (4): 214, 1917 (
*** Herbertus juniperoideus (Sw.) Grolle, Rev. Bryol. Lichénol. 30 (1/2): 80, 1961 (
*** Herbertus juniperoideus subsp. acanthelius (Spruce) K.Feldberg et Heinrichs, Bot. J. Linn. Soc. 151: 326, 2006 (
*** Herbertus juniperoideus subsp. pensilis (Taylor) K.Feldberg et Heinrichs, Bot. J. Linn. Soc. 151: 329, 2006 (
** Herbertus kurzii (Steph.) R.S.Chopra, J. Indian Bot. Soc. 22: 247, 1943 (
** Herbertus leratii (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 327, 1965 (
** Herbertus lonchobasis H.A.Mill., J. Hattori Bot. Lab. 28: 306, 1965 (
** Herbertus longifissus Steph., Hedwigia 34 (2): 44, 1895 (
** Herbertus longispinus J.B.Jack et Steph., Hedwigia 31 (1): 15, 1892 (
** Herbertus mauritianus N.G.Hodgetts, J. Bryol. 30 (4): 247, 2008 (
*** Herbertus norenus D.G.Long, D.Bell et H.H.Blom, Molec. Ecol. Res. 12: 44, 2012 (
** Herbertus oldfieldianus (Steph.) Rodway, Tasm. Bryoph.: 72, 1917 (
** Herbertus pilifer Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 252, 1893 (
** Herbertus pocsii N.G.Hodgetts, J. Bryol. 30 (4): 249, 2008 (
** Herbertus pumilus Steph., Hedwigia 34 (2): 44, 1895 (
** Herbertus ramosus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 314, 1965 (
** Herbertus runcinatus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (
*** Herbertus sendtneri (Nees) Lindb., Hepat. Scand. Exsicc.: no. 4, 1874 (
** Herbertus spicatus N.G.Hodgetts, J. Bryol. 30 (4): 244, 2008 (
*** Herbertus stramineus (Dumort.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (
** Herbertus streimannii M.L.So, Syst. Bot. 28 (1): 13, 2003 (
*** Herbertus tenuis A.Evans, Bull. Torrey Bot. Club 44 (4): 219, 1917 (
** Herbertus udarii D.Kumar et N.Manocha, Geophytology 29 (1/2): 105, 1999 [2000] (
Excluded from the genus
* Herbertus subrotundatus X.Fu et Y.J.Yi, Acta Phytotax. Sin. 39 (1): 89, 2001 (
*** Triandrophyllum Fulford et Hatcher, Bryologist 64 (4): 349, 1961 [1962] (
*** Triandrophyllum eophyllum (R.M.Schust.) Gradst., Mem. New York Bot. Gard. 86: 104, 2001 (
** Triandrophyllum fernandeziense (S.W.Arnell) Grolle ex Fulford et Hatcher, Bryologist 64 (4): 351, 1961 [1962] (
** Triandrophyllum heterophyllum (Steph.) Grolle, J. Jap. Bot. 39 (8): 238, 1964 (
*** Triandrophyllum subtrifidum (Hook.f. et Taylor) Fulford et Hatcher, Bryologist 64 (4): 350, 1961 [1962] (
** Triandrophyllum subtrifidum var. trifidum (Gottsche) Solari, Bol. Soc. Argent. Bot. 15 (2/3): 201, 1973 (
** Triandrophyllum symmetricum J.J.Engel, Haussknechtia, Beih. 9: 115, 1999 (
by M. von Konrat
Lepicoleaceae has conventionally been considered a monogeneric family only containing Lepicolea (e.g.
*** Lepicolea Dumort., Recueil Observ. Jungerm.: 20, 1835 (
*** Lepicolea attenuata (Mitt.) Steph., J. Linn. Soc., Bot. 29 (201): 276, 1892 (
*** Lepicolea magellanica (Gola) Solari, Lindbergia 9 (2): 86, 1983 (
*** Lepicolea norrisii Piippo, Ann. Bot. Fenn. 25 (1): 55, 1988 (
*** Lepicolea ochroleuca (Spreng.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 345, 1885 (
*** Lepicolea pruinosa (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 345, 1885 (
*** Lepicolea ramentifissa Herzog, Biblioth. Bot. 88: 30, 1920 [1921] (
*** Lepicolea rara (Steph.) Grolle, Nova Hedwigia 16: 152, 1968 (
*** Lepicolea rigida (De Not.) E.B.Scott, Nova Hedwigia 2: 148, 1960 (
*** Lepicolea scolopendra (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 398, 1877 (
*** Lepicolea yakusimensis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 10: 42, 1953 (
*** Vetaforma Fulford et J.Taylor, Nova Hedwigia 4 (1/2): 81, 1962 (
*** Vetaforma dusenii (Steph.) Fulford et J.Taylor, Nova Hedwigia 4 (1/2): 82, 1962 (
by E.D. Cooper
The classification within Lepidoziaceae is in a state of flux. The treatment provided here follows the interim classification proposed by
** Meinungeria Frank Müll., Bryologist 110 (3): 494, 2007 (
** Meinungeria mouensis Frank Müll., Bryologist 110 (3): 494, 2007 (
*** Acromastigum A.Evans, Bull. Torrey Bot. Club 27 (3): 103, 1900 (
** subg. Acromastigum
*** Acromastigum caledonicum (Steph.) Grolle, Österr. Bot. Z. 111 (2/3): 243, 1964 (
*** Acromastigum cavifolium R.M.Schust., J. Hattori Bot. Lab. 26: 257, 1963 (
** Acromastigum herzogii Grolle, Österr. Bot. Z. 111 (2/3): 250, 1964 (
*** Acromastigum homodictyon (Herzog) Grolle, Österr. Bot. Z. 111 (2/3): 245, 1964 (
** Acromastigum integrifolium (Austin) A.Evans, Bull. Torrey Bot. Club 27 (3): 103, 1900 (
*** Acromastigum stellare N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 109, 1985 (
** Acromastigum stenophyllum R.M.Schust., Nova Hedwigia 15: 465, 1968 (
*** Acromastigum verticale (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 18, 1954 (
** subg. Inaequilatera (Schiffn.) Grolle, J. Hattori Bot. Lab. 44: 2, 1978 (
*** Acromastigum adaptatum Hürl., Bauhinia 7 (4): 263, 1983 (
*** Acromastigum anisostomum (Lehm. et Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 48, 1934 (
** Acromastigum anisostomum var. minutum E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 71, 1965 (
** Acromastigum aurescens A.Evans, Ann. Bryol., Suppl. 3: 45, 1934 (
** Acromastigum bancanum (Sande Lac.) A.Evans, Ann. Bryol., Suppl. 3: 20, 1934 (
** Acromastigum brotheri (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 70, 1934 (
** Acromastigum capillare (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 37, 1934 (
*** Acromastigum colensoanum (Mitt.) A.Evans ex Reimers, Hedwigia 73 (3/4): 142, 1933 (
*** Acromastigum cunninghamii (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 106, 1934 (
** Acromastigum curtilobum A.Evans, Ann. Bryol., Suppl. 3: 97, 1934 (
*** Acromastigum divaricatum (Nees) A.Evans ex Reimers, Hedwigia 73 (3/4): 142, 1933 (
** Acromastigum echinatiforme (De Not.) A.Evans, Ann. Bryol., Suppl. 3: 64, 1934 (
* Acromastigum echinatum (Gottsche) A.Evans, Ann. Bryol., Suppl. 3: 147, 1934 (
*** Acromastigum exiguum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 75, 1934 (
** Acromastigum exile (Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 24, 1934 (
*** Acromastigum filum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 31, 1934 (
*** Acromastigum filum var. papillosum N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 112, 1985 (
** Acromastigum fimbriatum (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 153, 1934 (
** Acromastigum furcatifolium (Steph.) E.A.Br., Phytotaxa 65: 58, 2012 (
*** Acromastigum inaequilaterum (Lehm. et Lindenb.) A.Evans, Ann. Bryol., Suppl. 3: 129, 1934 (
** Acromastigum interstisiale E.A.Br. et M.A.M.Renner, Telopea 17: 268, 2014 (
** Acromastigum laetevirens (Sande Lac. ex Steph.) A.Evans, Ann. Bryol., Suppl. 3: 94, 1934 (
** Acromastigum laevigatum A.Evans, Ann. Bryol., Suppl. 3: 101, 1934 (
** Acromastigum linganum (De Not.) A.Evans, Ann. Bryol., Suppl. 3: 118, 1934 (
** Acromastigum lobuliferum A.Evans, Ann. Bryol., Suppl. 3: 157, 1934 (
** Acromastigum longirete Grolle, J. Hattori Bot. Lab. 44: 11, 1978 (
*** Acromastigum mooreanum (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 19, 1954 (
*** Acromastigum moratii N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 119, 1985 (
** Acromastigum obliquatum (Mitt.) A.Evans, Ann. Bryol., Suppl. 3: 110, 1934 (
** Acromastigum tenax (Steph.) A.Evans, Ann. Bryol., Suppl. 3: 41, 1934 (
Incertae sedis
** Acromastigum fumosum E.A.Br. et M.A.M.Renner, Telopea 17: 274, 2014 (
** Acromastigum leptophyllum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 309, 1950 (
*** Acromastigum marginatum E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 22, 1954 (
** Acromastigum microstictum A.Evans, Ann. Bryol., Suppl. 3: 115, 1934 (
** Acromastigum prismaticale E.A.Br. et M.A.M.Renner, Telopea 17: 281, 2014 (
** Acromastigum pusillum N.Kitag., Acta Phytotax. Geobot. 36 (4/6): 116, 1985 (
** Acromastigum rigidum R.M.Schust., Nova Hedwigia 64 (3/4): 617, 1997 (
** Acromastigum subechinatiforme Hürl., Bauhinia 7 (4): 266, 1983 (
*** Bazzania Gray, Nat. Arr. Brit. Pl. 1: 704, 1821 (
*** Bazzania acanthostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 381, 1885 (
*** Bazzania accreta (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania acinaciformis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 290, 1907 (
** Bazzania acuminata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania acutifolia (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 147, 1898 (
*** Bazzania adnexa (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania adnexa var. aucklandica (Lindenb. et Gottsche) J.J.Engel et G.L.Merr., Bryologist 97 (3): 319, 1994 (
*** Bazzania affinis (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania albifolia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 195, 1934 (
** Bazzania ambigua (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
*** Bazzania amblyphylla Meagher, Nova Hedwigia 92 (3/4): 487, 2011 (
** Bazzania aneityensis (Steph.) Tixier, Bull. Mus. Natl. Hist. Nat. (Sér. 3), Bot. 10 (190): 80, 1973 (
** Bazzania angusta (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 307, 1950 (
** Bazzania angustifalcata Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 166, 1955 (
** Bazzania angustifolia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 198, 1934 (
** Bazzania angustisedens (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 445, 1972 [1973] (
** Bazzania angustistipula N.Kitag., J. Hattori Bot. Lab. 30: 268, 1967 (
*** Bazzania appendiculata (Mitt.) S.Hatt., Fl. E. Himalaya: 505, 1966 (
** Bazzania approximata Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 139, 1977 (
** Bazzania arcuata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania arcuata var. mamillosa Gradst. et A.R.Benitez, Nova Hedwigia 99 (1/2): 113, 2014 (
** Bazzania armatistipula (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 114, 1946 (
** Bazzania asperrima Steph., Rev. Bryol. 34 (3): 48, 1907 (
* Bazzania asymmetrica (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 77, 1979 (
** Bazzania aterrima (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 77, 1977 (
*** Bazzania aurescens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 374, 1885 (
** Bazzania avia Meagher, Nova Hedwigia 97 (3/4): 529, 2013 (
** Bazzania azorica H.Buch et Perss., Bryophyt. Azoren Madeira: 3, 1941 (
** Bazzania baldwinii Austin, Trans. Connecticut Acad. Arts 8 (15): 255, 1891 (
** Bazzania bernieri (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 142, 1965 (
*** Bazzania bescherellei Steph., Hedwigia 32 (4): 204, 1893 (
** Bazzania bhutanica N.Kitag. et Grolle, J. Hattori Bot. Lab. 61: 269, 1986 [1987] (
** Bazzania bicrenata N.Kitag., J. Hattori Bot. Lab. 47: 127, 1980 (
*** Bazzania bidens (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania bidens var. heterodonta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 372, 1885 (
** Bazzania bidentula (Steph.) Yasuda, Shokubutsugaku Kakuron: 711, 1911 (
*** Bazzania bilobata N.Kitag., J. Hattori Bot. Lab. 30: 257, 1967 (
** Bazzania borneensis N.Kitag., J. Hattori Bot. Lab. 37: 263, 1973 (
** Bazzania brasiliensis (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania brighamii (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 255, 1891 (
** Bazzania cadens N.Kitag., J. Hattori Bot. Lab. 47: 129, 1980 (
*** Bazzania calcarata (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 149, 1898 (
** Bazzania callida (Sande Lac. ex Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (
*** Bazzania canelensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 152, 1946 (
** Bazzania caudata (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950 (
*** Bazzania caudistipula (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 141, 1965 (
** Bazzania ceylanica (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 306, 1896 (
** Bazzania chilensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 51, 1946 (
*** Bazzania chimantensis Fulford, Bryologist 63 (2): 89, 1960 (
** Bazzania cincinnata (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
*** Bazzania citharodes Meagher, Nova Hedwigia 86 (3/4): 481, 2008 (
** Bazzania combinata (J.B.Jack et Steph.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 306, 1896 (
** Bazzania commutata (Lindenb. et Gottsche) Schiffn., Consp. Hepat. Arch. Ind.: 149, 1898 (
* Bazzania comorensis Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 197, 1891 [1892] (
** Bazzania confertifolia (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 298, 1950 (
** Bazzania conistipula (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
** Bazzania conophylla (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 150, 1898 (
** Bazzania consanguinea (Hampe et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania consociata (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
* Bazzania corbieri (Steph.) Meagher, Nova Hedwigia 86 (3/4): 483, 2008 (
** Bazzania crassidentata Fulford, Bull. Torrey Bot. Club 86 (5): 338, 1959 (
** Bazzania crassitexta Steph., Hedwigia 32 (4): 205, 1893 (
* Bazzania crenata (Trevis.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania cubensis (Gottsche ex Steph.) Pagán, Bryologist 42 (2): 38, 1939 (
** Bazzania cucullata Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 142, 1977 (
*** Bazzania cuneistipula (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
* Bazzania curvidens Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 197, 1891 [1892] (
** Bazzania debilis N.Kitag., J. Hattori Bot. Lab. 30: 256, 1967 (
** Bazzania deciduifolia Onr., Bull. Jard. Bot. Natl. Belg. 47 (1/2): 144, 1977 (
*** Bazzania decrescens (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania decrescens var. dentistipula Kiaer et Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 6, 1893 (
** Bazzania decrescens subsp. molleri (Steph.) E.W.Jones, J. Bryol. 8 (3): 303, 1975 (
** Bazzania decrescens subsp. pumila (Mitt.) Pócs, Trop. Bryol. 9: 129, 1994 (
** Bazzania densa (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 151, 1898 (
** Bazzania densa var. connata (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 151, 1898 (
*** Bazzania denticulata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania denticulifera Mägd., Nova Hedwigia 38: 53, 1983 (
** Bazzania denudata (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
*** Bazzania deplanchei (Gottsche) Jovet-Ast, Rev. Bryol. Lichénol. 18 (1/2): 83, 1949 (
** Bazzania deplanchei var. filamentosa Tixier, Cryptog. Bryol. Lichénol. 6 (2): 179, 1985 (
** Bazzania desciscens (Steph.) Meijer, Blumea 10 (2): 382, 1960 (
** Bazzania didericiana (Gottsche ex Steph.) Steph., Bull. Herb. Boissier 5 (10): 841, 1897 (
** Bazzania diminuta Herzog, Trans. Brit. Bryol. Soc. 1 (4): 305, 1950 (
* Bazzania distans (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
*** Bazzania diversicuspis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 373, 1885 (
** Bazzania drepanophylla Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950 (
** Bazzania dulitensis Herzog, Trans. Brit. Bryol. Soc. 1 (4): 299, 1950 (
** Bazzania dulongensis L.P.Zhou et Li Zhang, J. Bryol. 34 (1): 22, 2012 (
** Bazzania eggersiana (Steph.) Pagán, Bryologist 42 (2): 39, 1939 (
** Bazzania elmeri (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 446, 1972 [1973] (
** Bazzania elongata Fulford, Bull. Torrey Bot. Club 86 (5): 337, 1959 (
** Bazzania emarginata (Steph.) C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 17, 1904 (
** Bazzania engelii Glenny, Fieldiana, Bot. (n.ser.) 47: 176, 2007 (
*** Bazzania erosa (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania erosa var. pulopenangensis (Lindenb. et Gottsche) Schiffn., Consp. Hepat. Arch. Ind.: 157, 1898 (
*** Bazzania exempta J.J.Engel, J. Hattori Bot. Lab. 99: 197, 2006 (
*** Bazzania falcata (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania falcifolia (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
* Bazzania fallax (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 158, 1898 (
*** Bazzania fasciculata (Steph.) Meagher, Australas. Bryol. Newslett. 46: 6, 2002 (
** Bazzania fauriana (Steph.) S.Hatt., Bot. Mag. (Tokyo) 59 (693/694): 27, 1946 (
** Bazzania filiformis Steph., Hedwigia 28 (2): 131, 1889 (
*** Bazzania flaccida (Dumort.) Grolle, Lindbergia 1 (3/4): 200, 1972 [1973] (
** Bazzania flavescens (Sande Lac. ex Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 158, 1898 (
** Bazzania fleischeri (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (
* Bazzania francana (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 446, 1972 [1973] (
** Bazzania friabilis N.Kitag. et T.Kodama, J. Hattori Bot. Lab. 39: 67, 1975 (
*** Bazzania fuhreri Meagher, Nova Hedwigia 92 (3/4): 488, 2011 (
** Bazzania fuscescens A.Evans, Pap. Michigan Acad. Sci. 17: 85, 1932 [1933] (
*** Bazzania gamscottii Meagher, Nova Hedwigia 92 (3/4): 492, 2011 (
*** Bazzania gedeana (Steph.) Meijer, Blumea 10 (2): 378, 1960 (
*** Bazzania gracilis (Hampe et Gottsche) Steph., Hedwigia 27 (11/12): 279, 1888 (
** Bazzania grandiretis (Steph.) Herzog, Trans. Brit. Bryol. Soc. 1 (4): 298, 1950 (
** Bazzania griffithiana (Steph.) Mizut., J. Hattori Bot. Lab. 30: 82, 1967 (
** Bazzania gunniana (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
** Bazzania hainanensis L.P.Zhou et Li Zhang, J. Bryol. 34 (1): 25, 2012 (
** Bazzania halconiensis (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 447, 1972 [1973] (
** Bazzania hamatifolia (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
** Bazzania harpago (De Not.) Schiffn., Consp. Hepat. Arch. Ind.: 159, 1898 (
** Bazzania hebridensis (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
** Bazzania herminieri (Gottsche ex Steph.) Pagán, Bryologist 45 (4): 90, 1942 (
** Bazzania herzogiana Meijer, Blumea 10 (2): 371, 1960 (
** Bazzania heterostipa (Steph.) Fulford, Bull. Torrey Bot. Club 86 (6): 410, 1959 (
*** Bazzania himalayana (Mitt.) Schiffn., Österr. Bot. Z. 49 (4): 132, 1899 (
*** Bazzania hochstetteri (Reichardt) E.A.Hodgs., Trans. Roy. Soc. New Zealand 82 (1): 11, 1954 (
*** Bazzania hookeri (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania horridula Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 258, 1893 (
** Bazzania inaequabilis Steph., Trans. Connecticut Acad. Arts 12 (1): 21, 1904 (
** Bazzania inaequitexta Steph., Hedwigia 32 (4): 208, 1893 (
** Bazzania incrassata (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 448, 1972 [1973] (
** Bazzania indica (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania indigenarum (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 82, 1977 (
** Bazzania insignis (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
* Bazzania intermedia (Gottsche et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania intermedia var. sarawakiana (De Not.) Schiffn., Consp. Hepat. Arch. Ind.: 162, 1898 (
** Bazzania involuta (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania involuta var. submutica (Lindenb. et Gottsche) J.J.Engel et G.L.Merr., Bryologist 97 (3): 314, 1994 (
** Bazzania involutiformis (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania irregularis (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 163, 1898 (
*** Bazzania jamaicensis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania japonica (Sande Lac.) Lindb., Acta Soc. Sci. Fenn. 10: 224, 1872 [1873] (
** Bazzania japonica var. sumatrana Herzog, Ann. Naturhist. Mus. Wien 53 (1): 366, 1942 [1943] (
** Bazzania javanica (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 163, 1898 (
** Bazzania kernii Steph., Hedwigia 32 (4): 208, 1893 (
** Bazzania kokawana N.Kitag. et T.Kodama, J. Jap. Bot. 50 (1): 11, 1975 (
** Bazzania latifolia Steph., Hedwigia 32 (4): 209, 1893 (
* Bazzania lehmanniana (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania leratii (Beauverd) H.A.Mill., Phytologia 47 (4): 320, 1981 (
** Bazzania lessonii (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (
** Bazzania levieri (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 448, 1972 [1973] (
** Bazzania liebmaniana (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania linearis Herzog, Trans. Brit. Bryol. Soc. 1 (4): 303, 1950 (
** Bazzania linguiformis (Sande Lac.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
*** Bazzania longa (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania longa var. papillata (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 94, 1946 (
*** Bazzania longicaulis (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 165, 1898 (
** Bazzania longicaulis var. latiareata Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 46, 1950 [1951] (
*** Bazzania longistipula (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
*** Bazzania loricata (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania lowii (Sande Lac. ex Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 166, 1898 (
** Bazzania luzonensis (Steph.) Del Ros., Philipp. J. Sci. 100 (3/4): 231, 1971 (
** Bazzania macgregorii Steph., Hedwigia 32 (4): 210, 1893 (
** Bazzania macrostipula Fulford, Bull. Torrey Bot. Club 86 (6): 407, 1959 (
** Bazzania magna Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 197, 1934 (
** Bazzania magnistipula N.Kitag., J. Hattori Bot. Lab. 47: 132, 1980 (
** Bazzania malaccensis (Steph.) Tixier, Gard. Bull. Singapore 25 (3): 342, 1971 (
** Bazzania manillana (Gottsche ex Steph.) S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 113, 1951 (
* Bazzania marginata (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 449, 1972 [1973] (
** Bazzania marginella (Herzog) N.Kitag. et T.Kodama, Bull. Osaka Mus. Nat. Hist. 27: 17, 1973 (
* Bazzania mascarena (Steph.) Herzog, Bot. Not. 100 (4): 334, 1947 (
** Bazzania mayebarae S.Hatt., J. Hattori Bot. Lab. 19: 91, 1958 (
* Bazzania menzelii E.D.Cooper, Phytotaxa 97 (2): 52, 2013 (
** Bazzania merrillana (Steph.) Inoue ex Bonner, Index Hepat. 3: 359, 1963 (
** Bazzania minuta (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 255, 1891 (
** Bazzania minutidens (Steph.) Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 142, 1965 (
** Bazzania minutiserra (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 26 (2): 77, 1977 (
** Bazzania missionum (Herzog) Jovet-Ast, Rev. Bryol. Lichénol. 20 (1/2): 96, 1951 (
** Bazzania mittenii (Steph.) Steph., Hedwigia 28 (2): 132, 1889 (
*** Bazzania monilinervis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania morokensis (Steph.) Grolle, J. Hattori Bot. Lab. 31: 1, 1968 (
*** Bazzania nitida (F.Weber) Grolle, Rev. Bryol. Lichénol. 29 (3/4): 210, 1960 [1961] (
*** Bazzania novae-zelandiae (Mitt.) Besch. et C.Massal., Miss. sci. Cape Horn, Bot. 5: 233, 1889 (
** Bazzania nudicaulis A.Evans, Bryologist 26 (6): 62, 1923 [1924] (
** Bazzania nuuanuensis C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 15, 1904 (
** Bazzania obcuneata (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (
** Bazzania obtusata (Mitt.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (
** Bazzania okaritana Meagher et Glenny, J. Bryol. 29 (1): 60, 2007 (
** Bazzania orbanii Pócs, Acta Biol. Pl. Agr. 1: 16, 2010 [2011] (
** Bazzania ovistipula (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (
** Bazzania pallidevirens (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 42, 1946 (
** Bazzania papillosa S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 67, 1965 (
** Bazzania paradoxa (Sande Lac.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896 (
*** Bazzania parisii (Steph.) N.Kitag., J. Hattori Bot. Lab. 47: 135, 1980 (
** Bazzania parvitexta Steph., Hedwigia 32 (4): 211, 1893 (
** Bazzania patens (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania patentistipa (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 168, 1898 (
** Bazzania paucidens (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (
** Bazzania pearsonii Steph., Hedwigia 32 (4): 212, 1893 (
*** Bazzania pectinata (Lindenb. et Gottsche) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 259, 1893 (
** Bazzania perfalcata N.Kitag., J. Hattori Bot. Lab. 47: 135, 1980 (
** Bazzania perrotana E.W.Jones, J. Bryol. 8 (3): 310, 1975 (
** Bazzania peruviana (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
*** Bazzania phyllobola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 372, 1885 (
*** Bazzania placophylla (Taylor) Grolle, Rev. Bryol. Lichénol. 27 (1/2): 54, 1958 (
** Bazzania platycnema (Schwägr. ex Steph.) H.A.Mill., Bryologist 63 (2): 121, 1960 (
** Bazzania pompeana (Sande Lac.) Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 200, 1891 (
*** Bazzania praerupta (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
* Bazzania praerupta var. obliquata (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 170, 1898 (
** Bazzania praerupta var. thermalis Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 396, 1955 (
** Bazzania pseudovittata N.Kitag. et T.Kodama, J. Hattori Bot. Lab. 39: 69, 1975 (
** Bazzania pusilla (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896 (
** Bazzania pycnophylla (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania quadratistipula H.A.Mill., Phytologia 47 (4): 321, 1981 (
** Bazzania rabenhorstii (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 45, 1959 (
** Bazzania recurva (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
* Bazzania recurva var. major (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 172, 1898 (
** Bazzania reflexa (Gottsche) Steph., Rev. Bryol. 18 (4): 56, 1891 (
** Bazzania reinwardtii (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 172, 1898 (
** Bazzania renistipula Steph., Hedwigia 32 (4): 212, 1893 (
** Bazzania revoluta (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 450, 1972 [1973] (
*** Bazzania rimosa Meagher, Nova Hedwigia 86 (3/4): 489, 2008 (
** Bazzania roccatii Gola, Ann. Bot. (Rome) 6 (2): 273, 1907 (
*** Bazzania roraimensis (Steph.) Fulford, Ann. Cryptog. Phytopathol. 3: 27, 1946 (
** Bazzania sandvicensis (Gottsche ex Steph.) Steph., Bull. Herb. Boissier 5 (10): 841, 1897 (
** Bazzania sauropoda Meagher, Austrobaileya 7 (1): 129, 2005 (
*** Bazzania scalaris Meagher, Telopea 11 (3): 247, 2006 (
*** Bazzania schlimiana (Gottsche) Fulford, Bull. Torrey Bot. Club 86 (6): 401, 1959 (
** Bazzania schultze-motelii N.Kitag., J. Hattori Bot. Lab. 47: 138, 1980 (
** Bazzania schusterana N.Kitag., J. Hattori Bot. Lab. 47: 139, 1980 (
*** Bazzania schwaneckiana (Hampe et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
* Bazzania scutigera (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania semicordata (Lindenb. et Gottsche) Kuntze, Revis. Gen. Pl. 2: 832, 1891 (
* Bazzania serpentina (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania serrapiculata Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 141, 1965 (
** Bazzania serrata Fulford, Bull. Torrey Bot. Club 86 (5): 321, 1959 (
** Bazzania serrulatoides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 200, 1934 (
* Bazzania sikkimensis (Steph.) Herzog, Ann. Bryol. 12: 78, 1939 (
** Bazzania spinosa S.Okamura, J. Coll. Sci. Imp. Univ. Tokyo 38 (4): 2, 1916 (
** Bazzania spiralis (Reinw., Blume et Nees) Meijer, Blumea 10 (2): 381, 1960 (
** Bazzania spruceana Steph., Hedwigia 32 (4): 213, 1893 (
** Bazzania squarrosa (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (
*** Bazzania stolonifera (Sw.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
* Bazzania stolonifera var. granatensis (Gottsche) Fulford, Ann. Cryptog. Phytopathol. 3: 51, 1946 (
** Bazzania stresemannii (Herzog) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 77, 1979 (
** Bazzania subacuta (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 307, 1896 (
** Bazzania subaequitexta (Steph.) N.Kitag., Bull. Nara Univ. Educ., B 28 (2): 81, 1979 (
** Bazzania subintegra (Steph.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 69, 2015 (
** Bazzania sublonga Fulford, Bull. Torrey Bot. Club 86 (5): 334, 1959 (
** Bazzania subserrifolia (Beauverd) H.A.Mill., Phytologia 47 (4): 321, 1981 (
** Bazzania subserrulata A.Evans, Pap. Michigan Acad. Sci. 17: 97, 1932 [1933] (
*** Bazzania subtilis (Sande Lac.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania succulenta N.Kitag., J. Hattori Bot. Lab. 47: 141, 1980 (
** Bazzania sumatrana (Sande Lac. ex Steph.) Steph., Hedwigia 32 (4): 209, 1893 (
* Bazzania sumbavensis (Gottsche ex Steph.) Steph., Hedwigia 32 (4): 204, 1893 (
*** Bazzania taleana (Gottsche) Fulford, Ann. Cryptog. Phytopathol. 3: 54, 1946 (
*** Bazzania tayloriana (Mitt.) Kuntze, Revis. Gen. Pl. 2: 832, 1891 (
** Bazzania temariana (Steph.) H.A.Mill., Phytologia 47 (4): 321, 1981 (
*** Bazzania tessellata Meagher, Nova Hedwigia 92 (3/4): 492, 2011 (
** Bazzania tiaoloensis Mizut. et K.C.Chang, J. Hattori Bot. Lab. 60: 432, 1986 (
*** Bazzania tricrenata (Wahlenb.) Lindb., Musci Fenn. Exsic., fasc. 2: [2 (adnot.)], 1872 (
** Bazzania tricrenata var. fulfordiae W.S.Hong, Bryologist 91 (4): 331, 1988 (
*** Bazzania tridens (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
*** Bazzania tridens var. assamica (Steph.) Pócs, J. Hattori Bot. Lab. 32: 86, 1969 (
** Bazzania tridens var. cornutistipula (Steph.) Pócs, J. Hattori Bot. Lab. 32: 83, 1969 (
*** Bazzania trilobata (L.) Gray, Nat. Arr. Brit. Pl. 1: 704, 1821 (
** Bazzania trilobata var. depauperata (Müll.Frib.) Grolle, Lindbergia 1 (3/4): 197, 1972 [1973] (
*** Bazzania uncigera (Reinw., Blume et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania uncigera var. brevifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 301, 1950 (
** Bazzania uncigera var. gibba (Sande Lac.) Meijer, Blumea 10 (2): 378, 1960 (
** Bazzania undulata Herzog, Trans. Brit. Bryol. Soc. 1 (4): 306, 1950 (
** Bazzania vietnamica Pócs, J. Hattori Bot. Lab. 32: 90, 1969 (
** Bazzania vitiana Mitt., Hedwigia 32 (4): 214, 1893 (
*** Bazzania vittata (Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
** Bazzania vittata var. luxurians (De Not.) Schiffn., Consp. Hepat. Arch. Ind.: 178, 1898 (
** Bazzania wallichiana (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 415, 1877 (
** Bazzania watanabei Inoue, J. Jap. Bot. 34 (9): 269, 1959 (
* Bazzania wattsiana (Steph.) Meagher, Australas. Bryol. Newslett. 50: 8, 2005 (
** Bazzania wiltensii (Sande Lac. ex Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 180, 1898 (
** Bazzania wooroonooran Meagher, Nova Hedwigia 100 (3/4): 549, 2015 (
** Bazzania wrightii (Gottsche ex Steph.) Steph., Hedwigia 27 (11/12): 279, 1888 (
** Bazzania yoshinagana (Steph.) Yasuda, Shokubutsugaku Kakuron: 711, 1911 (
** Bazzania zollingeri (Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 414, 1877 (
*** Bazzania zonulata Meagher, Nova Hedwigia 86 (3/4): 491, 2008 (
* Mastigopelma Mitt., Fl. vit.: 406, 1871 [1873] (
** Mastigopelma fragile (Steph.) N.Kitag., J. Hattori Bot. Lab. 36: 454, 1972 [1973] (
** Mastigopelma pulvinulatum (De Not.) Grolle, J. Hattori Bot. Lab. 33: 39, 1970 (
** Mastigopelma simplex Mitt., Fl. vit.: 406, 1871 [1873] (
** Mastigopelma subfissum Grolle, J. Hattori Bot. Lab. 33: 36, 1970 (
*** Drucella E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (3): 45, 1962 (
*** Drucella integristipula (Steph.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (3): 45, 1962 (
** Dendrolembidium Herzog, Ark. Bot. (n.ser.) 1 (13): 497, 1951 (
*** Dendrolembidium dendroides (Carrington et Pearson) Herzog, Ark. Bot. (n.ser.) 1 (13): 500, 1951 (
*** Dendrolembidium tenax (Grev.) Herzog, Ark. Bot. (n.ser.) 1 (13): 499, 1951 (
** Hygrolembidium R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963 (
*** Hygrolembidium acrocladum (Berggr.) R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963 (
** Hygrolembidium andinum (Herzog) R.M.Schust., Nova Hedwigia 10 (1/2): 23, 1965 (
*** Hygrolembidium australe (Steph.) Grolle, J. Jap. Bot. 41 (8): 229, 1966 (
*** Hygrolembidium boschianum (Sande Lac.) R.M.Schust., J. Hattori Bot. Lab. 26: 277, 1963 (
** Hygrolembidium isophyllum R.M.Schust., Nova Hedwigia 15: 467, 1968 (
*** Hygrolembidium rigidum R.M.Schust. et J.J.Engel, Phytologia 62 (1): 9, 1987 (
*** Hygrolembidium triquetrum J.J.Engel et R.M.Schust., Phytologia 62 (1): 11, 1987 (
** Hygrolembidium ventrosum (Mitt.) Grolle, Marion Prince Edw. Is: 233, 1971 (
** Isolembidium R.M.Schust., Nova Hedwigia 15: 466, 1968 (
*** Isolembidium anomalum (Rodway) Grolle, J. Bryol. 10 (3): 264, 1979 (
** Isolembidium anomalum var. cucullatum (E.A.Hodgs.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 63: 268, 1987 (
*** Kurzia G.Martens, Flora 53 (27): 417, 1870 (0).
** subg. Kurzia
*** Kurzia capillaris (Sw.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 173, 1963 [1964] (
*** Kurzia capillaris subsp. capillaris var. verrucosa (Steph.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 111, 1984 (
*** Kurzia capillaris subsp. paramicola Pócs, Acta Biol. Pl. Agr. 2: 102, 2012 (
*** Kurzia capillaris subsp. stephanii (Renauld ex Steph.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 111, 1984 (
*** Kurzia gonyotricha (Sande Lac.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 167, 1963 [1964] (
** Kurzia nemoides (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 173, 1963 [1964] (
** subg. Micrisophylla (Fulford) J.J.Engel ex R.M.Schust., J. Hattori Bot. Lab. 48: 347, 1980 (
** Kurzia saddlensis (Besch. et C.Massal.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964] (
** subg. Microlepidozia (Spruce) R.M.Schust., J. Hattori Bot. Lab. 48: 355, 1980 (
** Kurzia calcarata (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (
** Kurzia compacta (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (
** Kurzia helophila R.M.Schust., J. Hattori Bot. Lab. 48: 368, 1980 (
** Kurzia helophila var. flaccida R.M.Schust. ex J.J.Engel, Novon 17 (3): 310, 2007 (
*** Kurzia hippurioides (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (
** Kurzia hippurioides var. ornata J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 80: 228, 1996 (
** Kurzia irregularis (Steph.) Grolle, J. Hattori Bot. Lab. 36: 548, 1972 [1973] (
** Kurzia moniliformis J.J.Engel, Cryptog. Bryol. 26 (1): 73, 2005 (
*** Kurzia pauciflora (Dicks.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964] (
Incertae sedis
** Kurzia abbreviata Mizut., J. Hattori Bot. Lab. 38: 379, 1974 (
** Kurzia abietinella (Herzog) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964] (
*** Kurzia bisetula (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964] (
** Kurzia borneensis Mizut., J. Hattori Bot. Lab. 38: 377, 1974 (
** Kurzia brasiliensis (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964] (
** Kurzia brevicalycina (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964] (
* Kurzia caduciloba R.M.Schust., Beih. Nova Hedwigia 118: 273, 2000 (
* Kurzia cucullifolia (Steph.) R.M.Schust., J. Hattori Bot. Lab. 48: 350, 1980 (
*** Kurzia flagellifera (Steph.) Grolle, J. Jap. Bot. 39 (3): 80, 1964 (
*** Kurzia fragilifolia R.M.Schust., J. Hattori Bot. Lab. 48: 364, 1980 (
** Kurzia fragillima (Herzog) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 174, 1963 [1964] (
** Kurzia geniculata Mizut., J. Hattori Bot. Lab. 38: 383, 1974 (
** Kurzia hawaica (C.M.Cooke) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964] (
** Kurzia hispida (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 170, 1963 [1964] (
** Kurzia lateconica (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964] (
** Kurzia lineariloba Mizut., J. Hattori Bot. Lab. 38: 382, 1974 (
*** Kurzia longicaulis Piippo, Acta Bot. Fenn. 131: 174, 1985 (
** Kurzia makinoana (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964] (
** Kurzia mauiensis (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 274, 1967 (
** Kurzia mollis (Steph.) J.J.Engel et R.M.Schust., Bryologist 79 (4): 514, 1976 [1977] (
*** Kurzia nivicola (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 52, 2013 (
*** Kurzia pallescens Grolle, Rev. Bryol. Lichénol. 32 (3/4): 177, 1963 [1964] (
** Kurzia pallida Piippo, Acta Bot. Fenn. 131: 178, 1985 (
*** Kurzia quinquespina J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 80: 217, 1996 (
** Kurzia reversa (Carrington et Pearson) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 175, 1963 [1964] (
** Kurzia setiformis (De Not.) J.J.Engel et R.M.Schust., Bryologist 79 (4): 514, 1976 [1977] (
** Kurzia sexfida (Steph.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (
** Kurzia sinensis K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 83, 1984 (
** Kurzia sylvatica (A.Evans) Grolle, Herzogia 3: 77, 1973 (
*** Kurzia tasmanica (Steph.) E.D.Cooper, Phytotaxa 97 (2): 52, 2013 (
** Kurzia tayloriana (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 274, 1967 (
** Kurzia tenerrima (Mitt.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964] (
** Kurzia touwii N.Kitag., Acta Phytotax. Geobot. 29 (1/5): 56, 1978 (
** Kurzia trichoclados (Müll.Frib.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 171, 1963 [1964] (
*** Kurzia trilobata (R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 118: 270, 2000 (
** Kurzia verticellata (Carrington) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 178, 1963 [1964] (
** Lembidium Mitt., Handb. N. Zeal. fl. 2: 754, 1867 (
*** Lembidium berggrenii Herzog, Ark. Bot. (n.ser.) 1 (13): 485, 1951 (
*** Lembidium longifolium R.M.Schust., Phytologia 45 (5): 420, 1980 (
*** Lembidium nutans (Hook.f. et Taylor) Mitt., Handb. N. Zeal. Fl. 2: 754, 1867 (
** Lembidium nutans var. flagelliferum E.A.Hodgs., Rec. Domin. Mus. 4 (11): 110, 1962 (
** Megalembidium R.M.Schust., J. Hattori Bot. Lab. 26: 258, 1963 (
*** Megalembidium insulanum (W.Martin et E.A.Hodgs.) R.M.Schust., J. Hattori Bot. Lab. 26: 258, 1963 (
** Pseudocephalozia R.M.Schust., Nova Hedwigia 10 (1/2): 21, 1965 (
** sect. Lobulatae R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972 (
** Pseudocephalozia cucullata J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 38: 694, 1974 (
*** Pseudocephalozia lobulata (Herzog) R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972 [1973] (
*** Pseudocephalozia quadriloba (Steph.) R.M.Schust., J. Hattori Bot. Lab. 36: 371, 1972 [1973] (
** sect. Pseudocephalozia
** Pseudocephalozia lepidozioides R.M.Schust., Nova Hedwigia 10 (1/2): 22, 1965 (
** Pseudocephalozia leptodictyon R.M.Schust., J. Hattori Bot. Lab. 36: 369, 1972 [1973] (
*** Pseudocephalozia paludicola R.M.Schust., Nova Hedwigia 10 (1/2): 21, 1965 (
*** Ceramanus E.D.Cooper, Phytotaxa 97 (2): 53, 2013 (
*** Ceramanus centipes (Lindenb. et Gottsche) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (
*** Ceramanus clatritexta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (
*** Ceramanus elegans (Colenso) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (
*** Ceramanus grossiseta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (
*** Ceramanus perfragilis (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (
*** Ceramanus pruinosa (Herzog) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (
*** Ceramanus tuberifera (J.J.Engel et R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 54, 2013 (
*** Lepidozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (
** Lepidozia acantha J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 71, 2001 (
** Lepidozia aequiloba Steph., Sp. Hepat. (Stephani) 6: 319, 1922 (
** Lepidozia africana Steph., Sp. Hepat. (Stephani) 6: 320, 1922 (
*** Lepidozia alstonii Fulford, Mem. New York Bot. Gard. 11 (2): 211, 1966 (
** Lepidozia ambigua De Not., Epat. Borneo: 25, 1874 (
** Lepidozia andicola Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924 (
** Lepidozia appressifolia Steph., Sp. Hepat. (Stephani) 3: 583, 1909 (
*** Lepidozia armata Steph., Sp. Hepat. (Stephani) 3: 567, 1909 (
** Lepidozia asymmetrica Steph., Sp. Hepat. (Stephani) 3: 586, 1909 (
** Lepidozia auriculata Mitt., Sp. Hepat. (Stephani) 3: 579, 1909 (
** Lepidozia australis (Lehm. et Lindenb.) Mitt., Fl. vit.: 406, 1871 [1873] (
** Lepidozia bidens J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 79, 2001 (
** Lepidozia biloba Herzog, Ann. Bryol. 4: 83, 1931 (
*** Lepidozia bisbifida Steph., Sp. Hepat. (Stephani) 3: 593, 1909 (
** Lepidozia borneensis Steph., Sp. Hepat. (Stephani) 3: 625, 1909 (
** Lepidozia bragginsiana E.D.Cooper et M.A.M.Renner, Phytotaxa 173 (2): 118, 2014 (
* Lepidozia brasiliensis Steph., Sp. Hepat. (Stephani) 3: 571, 1909 (
** Lepidozia brevidentata Mitt., Fl. vit.: 406, 1871 [1873] (
** Lepidozia brevifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 104, 1860 [1861] (
** Lepidozia brevifolia var. planifolia Schiffn., Ann. Bryol. 8: 155, 1935 (
** Lepidozia brotheri Steph., Sp. Hepat. (Stephani) 3: 623, 1909 (
** Lepidozia buffalona Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914 (
** Lepidozia bursifera S.Hatt. et Grolle, J. Hattori Bot. Lab. 30: 115, 1967 (
** Lepidozia caespitosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 362, 1885 (
** Lepidozia caledonica Steph., Rev. Bryol. 35 (2): 31, 1908 (
** Lepidozia caledonica var. tenuisecta Hürl., Bauhinia 8 (2): 109, 1985 (
* Lepidozia ceramensis Herzog, Hedwigia 66 (6): 340, 1926 (
** Lepidozia cherydrion Hürl., Bauhinia 8 (2): 109, 1985 (
** Lepidozia chiloensis Steph., Sp. Hepat. (Stephani) 6: 322, 1922 (
** Lepidozia chordulifera Taylor, London J. Bot. 5: 371, 1846 (
*** Lepidozia cladorhiza (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 210, 1845 (
*** Lepidozia coilophylla Taylor, London J. Bot. 5: 370, 1846 (
** Lepidozia coilophylla var. apiculiloba (Steph.) Fulford, Mem. New York Bot. Gard. 11 (2): 194, 1966 (
** Lepidozia communis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914 (
*** Lepidozia concinna Colenso, Trans. & Proc. New Zealand Inst. 18: 244, 1886 (
** Lepidozia cordata Lindenb., Syn. Hepat. 2: 207, 1845 (
* Lepidozia cordistipula Steph., Sp. Hepat. (Stephani) 6: 345, 1922 (
** Lepidozia crassitexta Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 111, 1914 (
*** Lepidozia cupressina (Sw.) Lindenb., Syn. Hepat. 2: 207, 1845 (
** Lepidozia cupressina subsp. natalensis (Steph.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 109, 1984 (
* Lepidozia cupressina subsp. pinnata (Hook.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 109, 1984 (
* Lepidozia cupressina subsp. quinquefida (Steph.) Pócs, Proc. Third Meeting Bryol. C. & E. Europe: 109, 1984 (
** Lepidozia decaisnei Steph., Sp. Hepat. (Stephani) 3: 588, 1909 (
** Lepidozia dendritica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 362, 1885 (
** Lepidozia densa Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 26, 1925 (
*** Lepidozia digitata Herzog, Trans. & Proc. Roy. Soc. New Zealand 68 (1): 45, 1938 (
*** Lepidozia eenii S.W.Arnell, Svensk Bot. Tidskr. 57 (2): 190, 1963 (
** Lepidozia elobata R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 74, 2001 (
** Lepidozia erosa Steph., Sp. Hepat. (Stephani) 3: 621, 1909 (
** Lepidozia erronea Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 725, 1942 (
** Lepidozia everettii Steph., Sp. Hepat. (Stephani) 3: 622, 1909 (
* Lepidozia everettii var. javensis Herzog, Ann. Bryol. 5: 78, 1932 (
** Lepidozia fauriana Steph., Sp. Hepat. (Stephani) 3: 631, 1909 (
*** Lepidozia ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 3: 614, 1909 (
** Lepidozia filamentosa (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 206, 1845 (
** Lepidozia fistulosa Mitt., Fl. vit.: 406, 1871 [1873] (
** Lepidozia flexuosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 103, 1860 [1861] (
** Lepidozia fuegiensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 63, 1911 (
** Lepidozia fugax J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 63, 2001 (
* Lepidozia gedena Steph., Sp. Hepat. (Stephani) 6: 327, 1922 (
*** Lepidozia glaucescens J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 101, 2001 (
*** Lepidozia glaucophylla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 207, 1845 (
** Lepidozia grandifolia Steph., Sp. Hepat. (Stephani) 3: 625, 1909 (
* Lepidozia griseola Herzog, Hedwigia 66 (6): 340, 1926 (
** Lepidozia groenlandica Lehm., Nov. Stirp. Pug. 10: 7, 1857 (
** Lepidozia gwamii Piippo, Ann. Bot. Fenn. 21 (4): 311, 1984 (
** Lepidozia hampeana Lindenb., Syn. Hepat. 2: 208, 1845 (
*** Lepidozia haskarliana (Gottsche, Lindenb. et Nees) Steph., Sp. Hepat. (Stephani) 3: 614, 1909 (
** Lepidozia hastatistipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 113, 1914 (
*** Lepidozia hirta Steph., Sp. Hepat. (Stephani) 3: 599, 1909 (
*** Lepidozia holorhiza (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 210, 1845 (
** Lepidozia holorhiza var. laxa (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 186, 1898 (
** Lepidozia inaequalis (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 209, 1845 (
*** Lepidozia incurvata Lindenb., Syn. Hepat. 2: 203, 1845 (
** Lepidozia infuscata Mitt., Fl. vit.: 406, 1871 [1873] (
** Lepidozia integrifolia Doei, J. Hattori Bot. Lab. 63: 421, 1987 (
*** Lepidozia jamaicensis Steph., Sp. Hepat. (Stephani) 3: 568, 1909 (
** Lepidozia kashyapii D.Singh et D.K.Singh, Nova Hedwigia 94 (1/2): 222, 2012 (
** Lepidozia kinabaluensis Mizut., J. Hattori Bot. Lab. 38: 372, 1974 (
*** Lepidozia kirkii Steph., Sp. Hepat. (Stephani) 3: 598, 1909 (
** Lepidozia lacerifolia Steph., Sp. Hepat. (Stephani) 6: 332, 1922 (
*** Lepidozia laevifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 208, 1845 (
** Lepidozia laevifolia var. acutiloba J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 61, 2001 (
** Lepidozia laevifolia var. alpina R.M.Schust. et J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 62, 2001 (
** Lepidozia lindigiana Steph., Sp. Hepat. (Stephani) 3: 573, 1909 (
** Lepidozia loheri Steph., Sp. Hepat. (Stephani) 3: 621, 1909 (
** Lepidozia longifolia Steph., Sp. Hepat. (Stephani) 3: 606, 1909 (
** Lepidozia loriana Steph., Sp. Hepat. (Stephani) 6: 333, 1922 (
*** Lepidozia macrocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 363, 1885 (
* Lepidozia massartiana Schiffn. ex Steph., Sp. Hepat. (Stephani) 3: 611, 1909 (
*** Lepidozia microphylla (Hook.) Lindenb., Syn. Hepat. 2: 202, 1845 (
** Lepidozia microstipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 114, 1914 (
** Lepidozia minima Steph., Sp. Hepat. (Stephani) 6: 335, 1922 (
* Lepidozia minor (Gottsche, Lindenb. et Nees) Solari, Boll. Mus. Civico Storia Nat. Verona 10: 203, 1983 [1985] (
** Lepidozia miqueliana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 301, 1864 (
** Lepidozia montana Steph., Sp. Hepat. (Stephani) 3: 587, 1909 (
* Lepidozia newtonii Steph., Sp. Hepat. (Stephani) 3: 623, 1909 (
* Lepidozia nova Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 115, 1914 (
*** Lepidozia novae-zelandiae Steph., Sp. Hepat. (Stephani) 3: 595, 1909 (
** Lepidozia novae-zelandiae var. heterostipa R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 70, 2001 (
** Lepidozia novae-zelandiae var. minima R.M.Schust., Fieldiana, Bot. (n.ser.) 42: 71, 2001 (
*** Lepidozia obtusiloba Steph., Sp. Hepat. (Stephani) 3: 598, 1909 (
*** Lepidozia obtusiloba var. parvula J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 48, 2001 (
** Lepidozia omeiensis P.C.Chen ex Mizut. et K.C.Chang, J. Hattori Bot. Lab. 60: 421, 1986 (
*** Lepidozia ornata J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 49, 2001 (
* Lepidozia pallida Steph., Sp. Hepat. (Stephani) 3: 604, 1909 (
* Lepidozia palmicola Steph., Sp. Hepat. (Stephani) 6: 346, 1922 (
** Lepidozia paschalis Steph., Sp. Hepat. (Stephani) 6: 336, 1922 (
*** Lepidozia patens Lindenb., Syn. Hepat. 2: 202, 1845 (
* Lepidozia paucifolia Steph., Sp. Hepat. (Stephani) 3: 610, 1909 (
** Lepidozia paupercula Steph., Sp. Hepat. (Stephani) 6: 337, 1922 (
** Lepidozia pearsonii Spruce, J. Bot. 19: 34, 1881 (
*** Lepidozia pendulina (Hook.) Lindenb., Syn. Hepat. 2: 208, 1845 (
** Lepidozia peruviensis Steph., Sp. Hepat. (Stephani) 3: 575, 1909 (
*** Lepidozia pinnaticruris Spruce ex Steph., Sp. Hepat. (Stephani) 3: 579, 1909 (
* Lepidozia plumula Herzog, Beih. Bot. Centralbl. 38 (2): 331, 1921 (
** Lepidozia portoricensis Fulford, Mem. New York Bot. Gard. 11 (2): 187, 1966 (
*** Lepidozia procera Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 231, 1860 (
* Lepidozia pseudocupressina Schiffn., Krit. Bemerk. Eur. Lebermoose 14: 9, 1919 (
*** Lepidozia pumila J.J.Engel, Fieldiana, Bot. (n.ser.) 42: 76, 2001 (
*** Lepidozia quadridens (Nees) Nees, Syn. Hepat. 2: 209, 1845 (
** Lepidozia quadrifida Lindenb., Syn. Hepat. 2: 203, 1845 (
*** Lepidozia reptans (L.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (
** Lepidozia richardsii Herzog, Trans. Brit. Bryol. Soc. 1 (4): 313, 1950 (
** Lepidozia rigida Steph., Sp. Hepat. (Stephani) 6: 340, 1922 (
** Lepidozia robusta Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 217, 1894 (
* Lepidozia rufescens Steph., Biblioth. Bot. 87 (2): 226, 1916 (
** Lepidozia sandvicensis Lindenb., Syn. Hepat. 2: 201, 1845 (
** Lepidozia schwabei Herzog, Rev. Bryol. Lichénol. 23 (1/2): 49, 1954 (
** Lepidozia sellingiana H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 494, 1963 (
*** Lepidozia septemfida Steph., Sp. Hepat. (Stephani) 3: 588, 1909 (
** Lepidozia serpens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 364, 1885 (
*** Lepidozia serrulata J.J.Engel, J. Hattori Bot. Lab. 96: 273, 2004 (
*** Lepidozia setigera Steph., Sp. Hepat. (Stephani) 3: 599, 1909 (
** Lepidozia sikkimensis Steph., Sp. Hepat. (Stephani) 6: 341, 1922 (
*** Lepidozia spinosissima (Hook.f. et Taylor) Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 146, 1854 (
* Lepidozia squamifolia Steph., Sp. Hepat. (Stephani) 6: 341, 1922 (
*** Lepidozia squarrosa Steph., Sp. Hepat. (Stephani) 3: 573, 1909 (
** Lepidozia stahlii Steph., Sp. Hepat. (Stephani) 3: 616, 1909 (
** Lepidozia stuhlmannii Steph., Bot. Jahrb. Syst. 20 (3): 308, 1895 (
** Lepidozia stuhlmannii var. carnosa (Steph.) Pócs, Lidia 4 (1): 22, 1997 (
** Lepidozia stuhlmannii subsp. pulvinata (Steph.) Pócs, Trop. Bryol. 9: 127, 1994 (
** Lepidozia subdichotoma Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 361, 1885 (
** Lepidozia subintegra Lindenb., Syn. Hepat. 2: 201, 1845 (
** Lepidozia subtransversa Steph., Bull. Herb. Boissier 5 (2): 95, 1897 (
** Lepidozia subtrichodes Steph., Sp. Hepat. (Stephani) 3: 615, 1909 (
** Lepidozia succida Mitt., Trans. Linn. Soc. London 23 (1): 57, 1860 (
** Lepidozia supradecomposita Lindenb., Syn. Hepat. 2: 202, 1845 (
* Lepidozia supradecomposita var. falcifolia Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 191, 1931 (
** Lepidozia suyungii C.Gao et X.L.Bai, J. Hattori Bot. Lab. 92: 192, 2002 (
** Lepidozia terricola Steph., Sp. Hepat. (Stephani) 3: 585, 1909 (
** Lepidozia triangulifolia Steph., Sp. Hepat. (Stephani) 6: 344, 1922 (
*** Lepidozia trichodes (Reinw., Blume et Nees) Nees, Syn. Hepat. 2: 203, 1845 (
* Lepidozia tricuspidata Steph., Sp. Hepat. (Stephani) 6: 344, 1922 (
** Lepidozia tunguraguae Steph., Sp. Hepat. (Stephani) 6: 345, 1922 (
** Lepidozia ubangiensis Steph., Sp. Hepat. (Stephani) 3: 561, 1909 (
*** Lepidozia udarii S.C.Srivast., D.Kumar et D.Sharma, Cryptog. Bryol. Lichénol. 9 (3): 237, 1988 (
*** Lepidozia ulothrix (Schwägr.) Lindenb., Syn. Hepat. 2: 210, 1845 (
** Lepidozia vitrea Steph., Bull. Herb. Boissier 5 (2): 96, 1897 (
** Lepidozia wattsiana Steph., Sp. Hepat. (Stephani) 3: 586, 1909 (
** Lepidozia weymouthiana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 117, 1914 (
Excluded from the genus
* Lepidozia hexiloba Pearson, Ann. Cryptog. Exot. 4 (2): 67, 1931 (
* Lepidozia parvistipa Taylor, London J. Bot. 5: 370, 1846 (
*** Neolepidozia Fulford et J.Taylor, Brittonia 11 (2): 81, 1959 (
*** Neolepidozia aubertii (Jovet-Ast) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (
*** Neolepidozia autoica (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (
*** Neolepidozia capilligera (Schwägr.) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959 (
*** Neolepidozia consobrina (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (
*** Neolepidozia cuneifolia (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959 (
*** Neolepidozia disparata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (
*** Neolepidozia disticha (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959 (
*** Neolepidozia gibbsiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (
*** Neolepidozia heterotexta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (
*** Neolepidozia hodgsoniae (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 55, 2013 (
*** Neolepidozia longitudinalis (Herzog) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (
*** Neolepidozia mamillosa (Schiffn.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (
*** Neolepidozia meridiana (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (
*** Neolepidozia oligophylla (Lehm. et Lindenb.) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959 (
*** Neolepidozia ophiria (Gottsche) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (
*** Neolepidozia palmata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (
*** Neolepidozia paludicola (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (
*** Neolepidozia papulosa (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959 (
*** Neolepidozia parvifolia (Steph.) Fulford et J.Taylor, Brittonia 11 (2): 85, 1959 (
*** Neolepidozia patentissima (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (
*** Neolepidozia patentissima var. ampliata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 56, 2013 (
*** Neolepidozia patentissima var. zebrina (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia pennata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia planifolia (Steph.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia praenitens (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia praenitens var. dentifolia (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia quadristipula (Steph.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia rectangularis (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia seriatitexta (Steph.) Fulford, Mem. New York Bot. Gard. 11 (2): 215, 1966 (
*** Neolepidozia tetrapila (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia tetrapila var. cancellata (Colenso) E.D.Cooper, Phytotaxa 97 (2): 57, 2013 (
*** Neolepidozia tetrapila var. roseana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Neolepidozia tridactylis (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Neolepidozia trifida (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Neolepidozia verruculosa (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Neolepidozia wallichiana (Gottsche) Fulford et J.Taylor, Brittonia 11 (2): 84, 1959 (
*** Tricholepidozia (R.M.Schust.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Tricholepidozia chaetocarpa (Pearson) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Tricholepidozia fernandeziensis (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Tricholepidozia ferruginea (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Tricholepidozia fissifolia (Steph.) E.D.Cooper, Phytotaxa 97 (2): 58, 2013 (
*** Tricholepidozia jowettiana (H.A.Mill.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia kaindina (Grolle) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia kogiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia lawesii (Steph.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia leratii (Steph.) E.D.Cooper, Phytotaxa 167 (2): 218, 2014 (
*** Tricholepidozia lindenbergii (Gottsche) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia lindenbergii var. complanata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia lindenbergii var. mellea (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia lindenbergii var. papillata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia marginata (J.J.Engel et G.L.Merr.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia martinii (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 59, 2013 (
*** Tricholepidozia melanesica (H.A.Mill.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia neesii (Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia octoloba (Del Ros.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia plumulosa (Lehm. et Lindenb.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia pulcherrima (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia pulcherrima var. mooreana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia quadriseta (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia remotifolia (E.A.Hodgs.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia semperiana (Steph.) E.D.Cooper, Phytotaxa 97 (2): 60, 2013 (
*** Tricholepidozia tetradactyla (Hook.f. et Taylor) E.D.Cooper, Phytotaxa 97 (2): 61, 2013 (
*** Tricholepidozia trichocoleoides (Herzog) E.D.Cooper, Phytotaxa 97 (2): 61, 2013 (
** Micropterygium Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 233, 1845 (
** Micropterygium angustistipulum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 385, 1885 (
*** Micropterygium bialatum Fulford, Mem. New York Bot. Gard. 11 (2): 268, 1966 (
*** Micropterygium bolivarense Fulford, Mem. New York Bot. Gard. 11 (2): 263, 1966 (
*** Micropterygium campanense Spruce, Hedwigia 73 (3/4): 157, 1933 (
*** Micropterygium carinatum (Grev.) Reimers, Hedwigia 76 (3): 166, 1936 (
*** Micropterygium conchifolium Reimers, Hedwigia 73 (3/4): 155, 1933 (
*** Micropterygium duidae Reimers, Hedwigia 73 (3/4): 147, 1933 (
** Micropterygium exalatum Steph., Sp. Hepat. (Stephani) 3: 547, 1909 (
** Micropterygium grandistipulum Steph., Trans. Linn. Soc. London, Bot. 6 (1): 98, 1901 (
** Micropterygium laeve H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 252, 1976 (
** Micropterygium lechleri Reimers, Hedwigia 73 (3/4): 184, 1933 (
*** Micropterygium leiophyllum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 386, 1885 (
** Micropterygium longicellulatum Uribe et E.L.Linares, Phytotaxa 213 (3): 297, 2015 (
** Micropterygium parvistipulum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 383, 1885 (
*** Micropterygium pterygophyllum (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 413, 1877 (
* Micropterygium pterygophyllum var. lancifolium Reimers, Hedwigia 73 (3/4): 176, 1933 (
** Micropterygium reimersianum Herzog, Hedwigia 81 (5/6): 226, 1943 (
*** Micropterygium steyermarkii Fulford, Mem. New York Bot. Gard. 11 (2): 270, 1966 (
*** Micropterygium tatei Reimers, Hedwigia 73 (3/4): 150, 1933 (
*** Micropterygium tenax (Steph.) Grolle, J. Bryol. 10 (3): 265, 1979 (
*** Micropterygium trachyphyllum Reimers, Hedwigia 73 (3/4): 186, 1933 (
* Micropterygium trachyphyllum var. brasiliense Reimers, Hedwigia 73 (3/4): 195, 1933 (
* Micropterygium trachyphyllum var. cubense Reimers, Hedwigia 73 (3/4): 188, 1933 (
* Micropterygium trachyphyllum var. guadeloupense Reimers, Hedwigia 73 (3/4): 190, 1933 (
* Micropterygium trachyphyllum var. jamaicense Reimers, Hedwigia 73 (3/4): 190, 1933 (
*** Micropterygium tumidulum Fulford, Mem. New York Bot. Gard. 11 (2): 272, 1966 (
** Mytilopsis Spruce, Cephalozia: 90, 1882 (
*** Mytilopsis albifrons Spruce, Cephalozia: 91, 1882 (
** Protocephalozia (Spruce) K.I.Goebel, Flora 77 (2): 83, 1893 (
*** Protocephalozia ephemeroides (Spruce) K.I.Goebel, Flora 77 (2): 83, 1893 (
** Amazoopsis J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 242, 2004 (
*** Amazoopsis diplopoda (Pócs) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 245, 2004 (
*** Amazoopsis dissotricha (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 247, 2004 (
*** Amazoopsis gracilis J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 246, 2004 (
* Hyalolepidozia S.W.Arnell ex Grolle, Rev. Bryol. Lichénol. 32 (3/4): 179, 1963 [1964] (
** Hyalolepidozia bicuspidata (C.Massal.) S.W.Arnell ex Grolle, Rev. Bryol. Lichénol. 32 (3/4): 179, 1963 [1964] (
* Monodactylopsis (R.M.Schust.) R.M.Schust., Nova Hedwigia 56 (1/2): 45, 1993 (
** Monodactylopsis monodactyla (Spruce) R.M.Schust., Nova Hedwigia 69 (3/4): 523, 1999 (
** Neogrollea E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 70, 1965 (
*** Neogrollea notabilis E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 70, 1965 (
* Odontoseries Fulford, Mem. New York Bot. Gard. 11 (3): 364, 1968 (
* Odontoseries chimantana Fulford, Mem. New York Bot. Gard. 11 (3): 366, 1968 (
*** Paracromastigum Fulford et J.Taylor, Brittonia 13 (4): 336, 1961 (
*** Paracromastigum denticulatum (Steph.) E.D.Cooper, Phytotaxa 97 (2): 61, 2013 (
*** Paracromastigum drucei (R.M.Schust.) R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974 (
*** Paracromastigum dusenii (Steph.) R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974 (
** Paracromastigum fiordlandiae R.M.Schust. et J.J.Engel, Brittonia 48 (2): 167, 1996 (
*** Paracromastigum furcifolium (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963 (
** Paracromastigum granatense (Gottsche) R.M.Schust., J. Hattori Bot. Lab. 48: 341, 1980 (
** Paracromastigum longiscyphum (Taylor) R.M.Schust. et J.J.Engel, Brittonia 48 (2): 167, 1996 (
** Paracromastigum macrostipum (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963 (
** Paracromastigum micromera (Spruce) R.M.Schust., J. Hattori Bot. Lab. 26: 276, 1963 (
** Paracromastigum microphyllum (R.M.Schust. ex J.J.Engel) E.D.Cooper, Phytotaxa 97 (2): 61, 2013 (
*** Paracromastigum pachyrhizum (Nees) Fulford, Mem. New York Bot. Gard. 11 (3): 390, 1968 (
** Paracromastigum ryszardii Váňa, Bedn.-Ochyra et Cykowska, Nova Hedwigia 89 (1/2): 122, 2009 (
** Paracromastigum stipulatum (Herzog) Fulford, Mem. New York Bot. Gard. 11 (3): 390, 1968 (
** Paracromastigum subsimplex (Steph.) Fulford et J.Taylor, Brittonia 13 (4): 336, 1961 (
** Paracromastigum succulentum (Sim) J.J.Engel et G.L.Merr., Bryologist 104 (1): 151, 2001 (
** Paracromastigum tristanianum (R.M.Schust.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 38: 700, 1974 (
** Paracromastigum vastilobum (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 255, 2004 (
** Psiloclada Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 143, 1854 (
*** Psiloclada clandestina Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 143, 1854 (
** Psiloclada clandestina subsp. melanesica R.M.Schust., J. Hattori Bot. Lab. 48: 411, 1980 (
** Psiloclada clandestina subsp. spinosa (S.W.Arnell) R.M.Schust., J. Hattori Bot. Lab. 48: 410, 1980 (
** Pteropsiella Spruce, J. Bot. 14: 161, 1876 (
*** Pteropsiella frondiformis Spruce, J. Bot. 14: 161, 1876 (
** Pteropsiella metzgeriiformis R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (
** Telaranea Spruce ex Schiffn., Hepat. (Engl.-Prantl): 103, 1893 (
* sect. Telaranea
*** Telaranea apiahyna (Steph.) Fulford, Brittonia 15 (1): 71, 1963 (
*** Telaranea bicruris (Steph.) M.Howe, Bull. Torrey Bot. Club 29 (5): 287, 1902 (
*** Telaranea blepharostoma (Steph.) Fulford, Brittonia 15 (1): 73, 1963 (
*** Telaranea breviseta (Herzog) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 131, 2004 (
*** Telaranea chaetophylla (Spruce) Schiffn., Hepat. (Engl.-Prantl): 103, 1893 (
*** Telaranea europaea J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 150, 2004 (
*** Telaranea fragilis Mizut., J. Hattori Bot. Lab. 40: 449, 1976 (
*** Telaranea granulata J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 103, 2004 (
*** Telaranea longifolia (M.Howe) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 163, 2004 (
*** Telaranea nematodes (Gottsche ex Austin) M.Howe, Bull. Torrey Bot. Club 29 (5): 284, 1902 (
*** Telaranea panchoi Del Ros., Philipp. J. Sci. 100 (3/4): 238, 1971 (
*** Telaranea pellucida J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 179, 2004 (
*** Telaranea pseudozoopsis (Herzog) Fulford, Brittonia 15 (1): 71, 1963 (
*** Telaranea redacta (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 186, 2004 (
*** Telaranea rosarioana H.A.Mill., J. Bryol. 14 (2): 240, 1986 (
*** Telaranea setosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 192, 2004 (
*** Telaranea trisetosa (Steph.) Grolle, J. Hattori Bot. Lab. 29: 280, 1966 (
* sect. Tenuifoliae (R.M.Schust.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 112, 2004 (
*** Telaranea anomala R.M.Schust. ex J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 121, 2004 (
** Telaranea bischleriana Pócs, Acta Bot. Hung. 48 (1/2): 120, 2006 (
*** Telaranea coactilis (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 140, 2004 (
*** Telaranea confervoides J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 143, 2004 (
*** Telaranea diacantha (Mont.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 145, 2004 (
*** Telaranea herzogii (E.A.Hodgs.) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 106, 1962 (
*** Telaranea inaequalis R.M.Schust. ex J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 117, 2004 (
** Telaranea maorensis Pócs, Acta Bot. Hung. 48 (1/2): 124, 2006 (
*** Telaranea microstipulata R.M.Schust., Phytologia 39 (4): 241, 1978 (
*** Telaranea monocera (Mitt. ex R.M.Schust. et Grolle) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 168, 2004 (
*** Telaranea pecten (Spruce) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 178, 2004 (
*** Telaranea sejuncta (Ångstr.) S.W.Arnell, Bot. Not. 110 (1): 18, 1957 (
*** Telaranea tenuifolia J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 193, 2004 (
Incertae sedis
** Telaranea azorica (H.Buch et Perss.) Pócs in Schumacker et Váňa, Identif. keys liverw. hornw. Europe: 160, 2005 (
** Telaranea indica (S.C.Srivast. et P.K.Verma) A.E.D.Daniels et P.Daniel, Bull. Bot. Surv. India 49 (1/4): 231, 2007 (
** Telaranea major (Herzog) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 44: 165, 2004 (
** Zoopsidella R.M.Schust., Nova Hedwigia 10 (1/2): 24, 1965 (
** Zoopsidella antillana (Steph.) H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 254, 1976 (
** Zoopsidella antillana subsp. jamaicensis R.M.Schust., Nova Hedwigia 69 (1/2): 132, 1999 (
*** Zoopsidella caledonica (Steph.) R.M.Schust., Taxon 18 (1): 57, 1969 (
** Zoopsidella cynosurandra (Steph.) R.M.Schust., Nova Hedwigia 10 (1/2): 24, 1965 (
*** Zoopsidella integrifolia (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
** Zoopsidella macella (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
*** Zoopsidella serra (Spruce) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
** Zoopsis Hook.f. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 473, 1846 (
** subg. Eozoopsis R.M.Schust., J. Hattori Bot. Lab. 36: 373, 1972 (
*** Zoopsis leitgebiana (Carrington et Pearson) Bastow, Pap. & Proc. Roy. Soc. Tasmania 1887: 269, 1888 (
*** Zoopsis macrophylla R.M.Schust., Nova Hedwigia 68 (1/2): 14, 1999 (
** subg. Zoopsis
*** Zoopsis argentea (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 473, 1846 (
** Zoopsis argentea var. flagelliformis (Colenso) R.M.Schust., Nova Hedwigia 68 (1/2): 38, 1999 (
*** Zoopsis bicruris Glenny et E.A.Br., J. Bryol. 28 (4): 332, 2006 (
** Zoopsis liukiuensis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 65, 1931 (
*** Zoopsis matawaia M.A.M.Renner, J. Bryol. 28 (4): 334, 2006 (
*** Zoopsis nitida Glenny, Braggins et R.M.Schust., J. Bryol. 19 (4): 776, 1997 (
*** Zoopsis setulosa Leitg., Mitt. Naturwiss. Vereins Steiermark 13: 24, 1876 (
Incertae sedis
*** Zoopsis ceratophylla (Spruce) Hamlin, Rec. Domin. Mus. 7: 311, 1972 (
* Zoopsis ciliata Colenso, Trans. & Proc. New Zealand Inst. 20: 253, 1888 (
* Zoopsis martinicensis Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 268 (282), 1908 (
** Zoopsis setigera K.I.Goebel, Flora 77 (2): 93, 1893 (
** Zoopsis uleana Steph., Hedwigia 44 (4): 225, 1905 (
by B.J. Crandall-Stotler, R. Stotler, J. Váňa, J.J.Engel and L. Söderström
** Bragginsella R.M.Schust., Bryologist 100 (3): 363, 1997 (
*** Bragginsella anomala R.M.Schust., Bryologist 100 (3): 363, 1997 (
*** Chiloscyphus Corda, Gen. hepat.: 651, 1829 (
** Chiloscyphus kashyapii A.Srivast. et S.C.Srivast., Indian Geocalyc.: 34, 2002 (
*** Chiloscyphus pallescens (Ehrh.) Dumort., Syll. Jungerm. Europ.: 67, 1831 (
* Chiloscyphus pallescens var. fragilis (Roth) Müll.Frib., Ber. Deutsch. Bot. Ges. 59 (10): 429, 1942 (
*** Chiloscyphus polyanthos (L.) Corda, Gen. hepat.: 651, 1829 (
* Chiloscyphus polyanthos var. rivularis (Schrad.) Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 24, 1889 (
Incertae sedis
** Chiloscyphus acutus Steph., Sp. Hepat. (Stephani) 6: 302, 1922 (
** Chiloscyphus alpicola J.J.Engel, Phytotaxa 207 (2): 181, 2015 (
** Chiloscyphus beesleyanus Pearson, J. Linn. Soc., Bot. 46 (305): 22, 1922 (
* Chiloscyphus bifidus Schiffn., Hep. Fl. Buitenzorg: 200, 1900 (
** Chiloscyphus breviculus B.Y.Yang et W.C.Lee, Bot. Bull. Acad. Sin. (n.ser.) 5 (2): 190, 1964 (
* Chiloscyphus brevistipulus Steph., Sp. Hepat. (Stephani) 6: 303, 1922 (
** Chiloscyphus chinnarensis Manju, K.P.Rajesh et Madhus., Acta Bot. Hung. 53 (1/2): 152, 2011 (
** Chiloscyphus confertifolius Steph., Sp. Hepat. (Stephani) 6: 304, 1922 (
** Chiloscyphus confertus Steph., Sp. Hepat. (Stephani) 6: 305, 1922 (
** Chiloscyphus cornutistipulus Steph., Sp. Hepat. (Stephani) 6: 303, 1922 (
* Chiloscyphus durus (Steph.) Hässel, Revista Mus. Argent. Ci. Nat. (n.ser.) 1 (2): 122, 1999 (
* Chiloscyphus ernstianus Steph., Sp. Hepat. (Stephani) 6: 306, 1922 (
** Chiloscyphus etesseanus Steph., Bull. Herb. Boissier (sér. 2) 7 (10): 845 (217), 1907 (
** Chiloscyphus francanus Steph., Sp. Hepat. (Stephani) 6: 306, 1922 (
** Chiloscyphus graeffeanus Steph., Sp. Hepat. (Stephani) 6: 307, 1922 (
** Chiloscyphus greenwelliae (H.A.Mill.) H.A.Mill., J. Hattori Bot. Lab. 30: 275, 1967 (
** Chiloscyphus hookeri J.J.Engel, J. Hattori Bot. Lab. 36: 150, 1972 [1973] (
* Chiloscyphus hookeri var. constantifolius J.J.Engel, J. Hattori Bot. Lab. 36: 155, 1972 [1973] (
** Chiloscyphus integerrimus Schiffn., Hep. Fl. Buitenzorg: 197, 1900 (
** Chiloscyphus kehdingianus (Steph.) N.Kitag., Hikobia, Suppl. 1: 68, 1981 (
** Chiloscyphus kilauensis Steph., Sp. Hepat. (Stephani) 6: 309, 1922 (
** Chiloscyphus koeppensis (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 8 (2): 139 (255), 1908 (
** Chiloscyphus laceratus Steph., Sp. Hepat. (Stephani) 6: 310, 1922 (
** Chiloscyphus lambertonii H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 506, 1963 (
** Chiloscyphus latistipus Steph., Sp. Hepat. (Stephani) 6: 309, 1922 (
** Chiloscyphus lepervanchei (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 418, 1984 [1985] (
** Chiloscyphus longifissus Steph., Sp. Hepat. (Stephani) 6: 310, 1922 (
** Chiloscyphus propagulifer Schiffn., Hep. Fl. Buitenzorg: 208, 1900 (
** Chiloscyphus purpureus Steph., Sp. Hepat. (Stephani) 6: 312, 1922 (
** Chiloscyphus quadricilius Steph., Sp. Hepat. (Stephani) 6: 312, 1922 (
*** Chiloscyphus quadridentatus (Spruce) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 422, 1984 [1985] (
** Chiloscyphus rotundifolius Mitt., Rep. Challenger, Bot. 1 (3, 1): 85, 1884 (
** Chiloscyphus rotundiphyllus H.A.Mill., Phytologia 47 (4): 321, 1981 (
** Chiloscyphus scaberulus Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890] (
** Chiloscyphus septatus J.J.Engel, Fieldiana, Bot. (n.ser.) 48: 125, 2010 (
** Chiloscyphus similis Steph., Rev. Bryol. 35 (2): 28, 1908 (
** Chiloscyphus skottsbergianus H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 508, 1963 (
** Chiloscyphus subacuminatus Herzog, Ark. Bot. (n.ser.) 3 (3): 49, 1953 (
** Chiloscyphus subsimilis Steph., Sp. Hepat. (Stephani) 6: 314, 1922 (
* Chiloscyphus tridens Steph., Sp. Hepat. (Stephani) 6: 315, 1922 (
** Chiloscyphus trigonifolius Steph., Sp. Hepat. (Stephani) 6: 316, 1922 (
* Chiloscyphus venustulus Colenso, Trans. & Proc. New Zealand Inst. 21: 60, 1889 (
** Clasmatocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 440, 1885 (
*** Clasmatocolea bisexualis Glenny et J.J.Engel, New Zealand J. Bot. 51 (1): 23, 2013 (
*** Clasmatocolea crassiretis (Herzog) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 69, 1962 [1963] (
*** Clasmatocolea ctenophylla (Schiffn.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 71, 1960 (
*** Clasmatocolea cucullistipula (Steph.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 71, 1960 (
*** Clasmatocolea fasciculata (Nees) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (
*** Clasmatocolea fulvella (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (
*** Clasmatocolea gayana (Mont.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (
*** Clasmatocolea humilis (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (
*** Clasmatocolea humilis var. polymorpha J.J.Engel, Phytologia 41 (5): 309, 1979 (
*** Clasmatocolea humilis var. suspecta (C.Massal.) J.J.Engel, Phytologia 41 (5): 309, 1979 (
*** Clasmatocolea inflexispina (Hook.f. et Taylor) J.J.Engel, Bryologist 94 (4): 436, 1991 (
*** Clasmatocolea marginata (Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 73, 1962 [1963] (
*** Clasmatocolea minutiretis J.J.Engel et Grolle, Phytologia 41 (5): 309, 1979 (
*** Clasmatocolea moniliformis J.J.Engel, Phytologia 41 (5): 310, 1979 (
*** Clasmatocolea navistipula (Steph.) Grolle, Feddes Repert. 82 (1): 88, 1971 (
*** Clasmatocolea navistipula var. parceramosa J.J.Engel, Phytologia 41 (5): 311, 1979 (
*** Clasmatocolea notophylla (Hook.f. et Taylor) Grolle, J. Jap. Bot. 41 (8): 228, 1966 (
*** Clasmatocolea obvoluta (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (
*** Clasmatocolea obvoluta var. cookiana (C.Massal.) J.J.Engel, Phytologia 41 (5): 311, 1979 (
*** Clasmatocolea puccioana (De Not.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 72, 1960 (
*** Clasmatocolea rigens (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 36: 156, 1972 [1973] (
*** Clasmatocolea strongylophylla (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 73, 1960 (
*** Clasmatocolea trachyopa (Hook.f. et Taylor) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 73, 1960 (
*** Clasmatocolea vermicularis (Lehm.) Grolle, Rev. Bryol. Lichénol. 29 (1/2): 78, 1960 (
*** Clasmatocolea verrucosa J.J.Engel, Bryologist 83 (2): 220, 1980 (
*** Conoscyphus Mitt., Fl. vit.: 404, 1871 [1873] (
** Conoscyphus koponenii Piippo, Mamontov et Potemkin, Acta Bryolichenol. Asiat. 5: 20, 2014 (
*** Conoscyphus trapezioides (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 125, 1898 (
*** Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa, Phytotaxa 97 (2): 39, 2013 (
*** Cryptolophocolea aculeata (Mitt.) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (
** Cryptolophocolea chiloscyphoidea (Lindenb.) L.Söderstr. et Crand.-Stotl., Phytotaxa 112 (1): 18, 2013 (
*** Cryptolophocolea ciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler et Váňa, Phytotaxa 97 (2): 39, 2013 (
** Cryptolophocolea compacta (Mitt.) L.Söderstr., Phytotaxa 112 (1): 19, 2013 (
*** Cryptolophocolea connata (Sw.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 19, 2013 (
*** Cryptolophocolea connatifolia (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (
*** Cryptolophocolea costata (Nees) L.Söderstr., Phytotaxa 112 (1): 19, 2013 (
*** Cryptolophocolea edentata (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (
** Cryptolophocolea explanata (Mitt.) Váňa et Crand.-Stotl., Phytotaxa 202 (1): 69, 2015 (
** Cryptolophocolea fleischeri (Steph.) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (
*** Cryptolophocolea guadalupensis (Steph.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 19, 2013 (
*** Cryptolophocolea helmsiana (Steph.) L.Söderstr., Phytotaxa 112 (1): 19, 2013 (
*** Cryptolophocolea leucophylla (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 19, 2013 (
* Cryptolophocolea levieri (Schiffn.) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (
* Cryptolophocolea lilliena (Steph.) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (
*** Cryptolophocolea martiana (Nees) L.Söderstr., Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 20, 2013 (
** Cryptolophocolea martiana subsp. bidentula (Nees) L.Söderstr., Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 20, 2013 (
** Cryptolophocolea martiana var. perissodonta (Spruce) Gradst., Phytoneuron 2015 (22): 1, 2015 (
* Cryptolophocolea massalongoana (Schiffn.) L.Söderstr., Phytotaxa 112 (1): 20, 2013 (
*** Cryptolophocolea mitteniana (Colenso) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (
*** Cryptolophocolea mitteniana var. obtusa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 21, 2013 (
*** Cryptolophocolea mitteniana var. symmetrica (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 22, 2013 (
*** Cryptolophocolea pallida (Mitt.) L.Söderstr., Phytotaxa 112 (1): 22, 2013 (
** Cryptolophocolea pallidovirens (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 20, 2013 (
* Cryptolophocolea proteus (Herzog) L.Söderstr., Phytotaxa 112 (1): 22, 2013 (
* Cryptolophocolea pycnophylla (Spruce) L.Söderstr., Phytotaxa 112 (1): 22, 2013 (
*** Cryptolophocolea regularis (Steph.) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (
*** Cryptolophocolea spinifera (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 20, 2013 (
* Cryptolophocolea stephanii (Schiffn.) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (
*** Cryptolophocolea subopposita (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (
* Cryptolophocolea thermarum (Schiffn.) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (
*** Cryptolophocolea trialata (Gottsche) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (
** Cryptolophocolea tricorata (Hässel) Crand.-Stotl. et Stotler, Phytotaxa 112 (1): 23, 2013 (
*** Cryptolophocolea tuberculata (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 23, 2013 (
* Cryptolophocolea whittieriana (Inoue et H.A.Mill.) L.Söderstr., Phytotaxa 112 (1): 24, 2013 (
*** Deceptifrons J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 54, 2013 (
*** Deceptifrons plagiochiloides J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 54, 2013 (
*** Evansianthus R.M.Schust. et J.J.Engel, Bryologist 76 (4): 516, 1973 (
*** Evansianthus georgiensis (Gottsche) R.M.Schust. et J.J.Engel, Bryologist 76 (4): 518, 1973 (
*** Hepatostolonophora J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 46: 91, 1979 (
*** Hepatostolonophora abnormis (Besch. et C.Massal.) J.J.Engel et R.M.Schust., J. Hattori Bot. Lab. 46: 94, 1979 (
** Hepatostolonophora conica (Steph.) Hässel, Bol. Soc. Argent. Bot. 22 (1/4): 123, 1983 (
*** Hepatostolonophora paucistipula (Rodway) J.J.Engel, J. Hattori Bot. Lab. 46: 103, 1979 (
*** Hepatostolonophora rotata (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 46: 98, 1979 (
*** Hepatostolonophora rotata var. perssonii (R.M.Schust.) J.J.Engel, J. Hattori Bot. Lab. 46: 103, 1979 (
*** Heteroscyphus Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 (
** Heteroscyphus acutangulus (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus allodontus (Hook.f. et Taylor) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 399, 1984 [1985] (
** Heteroscyphus amboinensis (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus ammophilus (Colenso) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980 (
** Heteroscyphus ammophilus var. obtusifolius J.J.Engel et G.L.Merr., Nova Hedwigia 99 (1/2): 158, 2014 (
*** Heteroscyphus argutus (Reinw., Blume et Nees) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus argutus var. brevidens (Schiffn.) Herzog et Nog., J. Hattori Bot. Lab. 14: 40, 1955 (
*** Heteroscyphus aselliformis (Reinw., Blume et Nees) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus assurgentifolius J.J.Engel, Nova Hedwigia 99 (1/2): 158, 2014 (
** Heteroscyphus baduinus (Nees) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 (
** Heteroscyphus balnetii (Herzog) Grolle, Acta Bot. Fenn. 125: 68, 1984 (
*** Heteroscyphus billardierei (Schwägr.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus billardierei var. clasmatocoleoides (J.J.Engel et G.L.Merr.) J.J.Engel, Nova Hedwigia 100 (3/4): 565, 2015 (
** Heteroscyphus brassii (Grolle) Grolle, Acta Bot. Fenn. 125: 68, 1984 (
* Heteroscyphus caesius (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
* Heteroscyphus caledonicus (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus chlorophyllus (Hook.f. et Taylor) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus ciliatus (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
*** Heteroscyphus coalitus (Hook.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus coalitus var. simplicifolius J.J.Engel, Nova Hedwigia 100 (3/4): 572, 2015 (
** Heteroscyphus combinatus (Nees) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
*** Heteroscyphus conjugatus (Mitt.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 399, 1984 [1985] (
*** Heteroscyphus contortuplicatus (Nees et Mont.) Grolle, J. Hattori Bot. Lab. 55: 504, 1984 (
*** Heteroscyphus cuneistipulus (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus darjeelingensis A.Srivast. et S.C.Srivast., Indian Geocalyc.: 115, 2002 (
** Heteroscyphus deceptifrons J.J.Engel, Nova Hedwigia 99 (1/2): 162, 2014 (
*** Heteroscyphus dentammophilus J.J.Engel et G.L.Merr., Polish Bot. J. 58 (1): 96, 2013 (
** Heteroscyphus denticulatus (Mitt.) Schiffn., Beih. Bot. Centralbl. 29: 106, 1912 (
** Heteroscyphus deplanchei (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus diestianus (Sande Lac.) Piippo, Acta Bot. Fenn. 131: 165, 1985 (
** Heteroscyphus divergenticiliatus (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 501, 1976 (
** Heteroscyphus dubius (Gottsche) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
*** Heteroscyphus echinellus (Lindenb. et Gottsche) J.J.Engel et Xiao L.He, Bryologist 113 (1): 155, 2010 (
** Heteroscyphus echinellus var. hyalinus J.J.Engel, Bryologist 113 (1): 161, 2010 (
** Heteroscyphus elliottii (Steph.) Pagán, Bryologist 42 (1): 7, 1939 (
* Heteroscyphus elliottii var. portoricensis Fulford, Mem. New York Bot. Gard. 11 (4): 489, 1976 (
* Heteroscyphus falcifolius (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
*** Heteroscyphus fissistipus (Hook.f. et Taylor) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus fissistipus var. multispinus (E.A.Hodgs. et Allison) J.J.Engel, Nova Hedwigia 99 (1/2): 165, 2014 (
** Heteroscyphus fissistipus var. repandus J.J.Engel, Nova Hedwigia 99 (1/2): 165, 2014 (
** Heteroscyphus flaccidus (Mitt.) A.Srivast. et S.C.Srivast., Indian Geocalyc.: 85, 2002 (
** Heteroscyphus fleischeri (Steph.) D.G.Long et Rubas., Ceylon J. Sci., Biol. Sci. 43 (1): 26, 2014 (
* Heteroscyphus fragilicilius (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus furcistipulus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (
** Heteroscyphus giganteus (Steph.) Hürl., Bauhinia 12 (1/2): 114, 1998 (
** Heteroscyphus graeffei (J.B.Jack et Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 29 (4): 645, 1980 (
** Heteroscyphus grandiflorus (Steph.) Hürl., Bauhinia 12 (1/2): 114, 1998 (
** Heteroscyphus grandistipus (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus gunnianus (Mitt.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (
** Heteroscyphus hastatus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (
** Heteroscyphus hebridensis (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus heterophyllus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (
** Heteroscyphus hyalinus (Steph.) A.Srivast. et S.C.Srivast., Indian Geocalyc.: 118, 2002 (
*** Heteroscyphus integrifolius (Lehm. et Lindenb.) Fulford, Mem. New York Bot. Gard. 11 (4): 495, 1976 (
** Heteroscyphus iwatsukii (S.Hatt.) Piippo, Willdenowia 18 (2): 522, 1989 (
** Heteroscyphus jackii (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
*** Heteroscyphus knightii (Steph.) Grolle, J. Hattori Bot. Lab. 61: 251, 1986 [1987] (
** Heteroscyphus levieri (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus limosus (Carrington et Pearson) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus lingulatus (Colenso) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (
** Heteroscyphus lophocoleoides S.Hatt., Bull. Tokyo Sci. Mus. 11: 45, 1944 (
*** Heteroscyphus lyallii (Mitt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980 (
*** Heteroscyphus magellanicus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (
** Heteroscyphus mamillatus Piippo, Nova Hedwigia 56 (3/4): 360, 1993 (
** Heteroscyphus marginatus (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 491, 1976 (
*** Heteroscyphus menziesii (Mitt.) J.J.Engel, Polish Bot. J. 58 (1): 99, 2013 (
** Heteroscyphus merapiensis (Steph.) Piippo, Ann. Bot. Fenn. 30 (3): 200, 1993 (
** Heteroscyphus miradorensis (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
*** Heteroscyphus mononuculus J.J.Engel, J. Hattori Bot. Lab. 90: 241, 2001 (
** Heteroscyphus mononuculus var. ammophilopsis J.J.Engel, Nova Hedwigia 99 (1/2): 166, 2014 (
** Heteroscyphus mononuculus var. bilobus J.J.Engel, Nova Hedwigia 99 (1/2): 167, 2014 (
** Heteroscyphus montagnei (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 497, 1976 (
** Heteroscyphus multifidus (Steph.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (
** Heteroscyphus multifidus var. subintegerrimus J.J.Engel, Nova Hedwigia 100 (3/4): 562, 2015 (
** Heteroscyphus nadeaudii (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus orbiculatus A.Srivast. et S.C.Srivast., Indian Geocalyc.: 140, 2002 (
** Heteroscyphus palniensis A.Srivast. et S.C.Srivast., Indian Geocalyc.: 130, 2002 (
** Heteroscyphus pandei S.C.Srivast. et A.Srivast., Lindbergia 15 (6): 197, 1989 [1991] (
*** Heteroscyphus parallelifolius J.J.Engel, Polish Bot. J. 58 (1): 99, 2013 (
** Heteroscyphus parvulus (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 (
** Heteroscyphus parvus A.Srivast. et S.C.Srivast., Indian Geocalyc.: 112, 2002 (
** Heteroscyphus perfoliatus (Mont.) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 (
** Heteroscyphus pertusus (Lehm.) Fulford, Mem. New York Bot. Gard. 11 (4): 505, 1976 (
*** Heteroscyphus planiusculus (Hook.f. et Taylor) J.J.Engel, J. Hattori Bot. Lab. 68: 315, 1990 (
** Heteroscyphus planus (Mitt.) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 (
*** Heteroscyphus polyblepharis (Spruce) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
*** Heteroscyphus polychaetus (Spruce) Hentschel et Heinrichs, Taxon 56 (4): 1139, 2007 (
** Heteroscyphus polycladus (Hook.f. et Lév.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980 (
** Heteroscyphus rectangulatus (Herzog) Piippo, Ann. Bot. Fenn. 30 (3): 200, 1993 (
** Heteroscyphus saccogynoides Herzog, J. Hattori Bot. Lab. 14: 40, 1955 (
** Heteroscyphus sarawaketanus Piippo, Acta Bot. Fenn. 131: 143, 1985 (
*** Heteroscyphus sinuosus (Hook.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus spectabilis (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus spinifer C.Gao, T.Cao et Y.H.Wu, J. Bryol. 26 (2): 97, 2004 (
*** Heteroscyphus splendens (Lehm. et Lindenb.) Grolle, Acta Bot. Fenn. 125: 68, 1984 (
** Heteroscyphus splendidus (E.A.Hodgs.) J.J.Engel et R.M.Schust., Nova Hedwigia 39: 400, 1984 [1985] (
*** Heteroscyphus stolonifer J.J.Engel, Polish Bot. J. 58 (1): 102, 2013 (
*** Heteroscyphus succulentus (Gottsche) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 (
** Heteroscyphus supinus (Hook.f. et Taylor) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 248, 1980 (
** Heteroscyphus tener (Steph.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
*** Heteroscyphus thraustus (Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 495, 1976 (
** Heteroscyphus timppae Piippo, Ann. Bot. Fenn. 29 (3): 246, 1992 (
*** Heteroscyphus triacanthus (Hook.f. et Lév.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus triacanthus var. magnistipulatus J.J.Engel, Nova Hedwigia 99 (1/2): 167, 2014 (
** Heteroscyphus tridentatus (Sande Lac.) Grolle, Acta Bot. Fenn. 125: 68, 1984 (
** Heteroscyphus turgidus (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 (
*** Heteroscyphus valdiviensis (Mont.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus varians (Steph.) J.J.Engel, J. Hattori Bot. Lab. 68: 315, 1990 (
** Heteroscyphus wettsteinii (Schiffn.) Schiffn., Österr. Bot. Z. 60 (5): 172, 1910 (
** Heteroscyphus zollingeri (Gottsche) Schiffn., Österr. Bot. Z. 60 (5): 171, 1910 (
*** Lamellocolea J.J.Engel, J. Hattori Bot. Lab. 70: 65, 1991 (
*** Lamellocolea granditexta (Steph.) J.J.Engel, J. Hattori Bot. Lab. 70: 66, 1991 (
*** Lamellocolea integrostia J.J.Engel et Glenny, Bryologist 114 (1): 23, 2011 (
*** Leptophyllopsis R.M.Schust., J. Hattori Bot. Lab. 26: 269, 1963 (
*** Leptophyllopsis laxa (Mitt.) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 284, 1972 (
*** Leptoscyphopsis R.M.Schust., Phytologia 39 (4): 246, 1978 (
*** Leptoscyphopsis paradoxa R.M.Schust., Phytologia 39 (4): 246, 1978 (
*** Leptoscyphus Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (
** subg. Anomylia (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 272, 1980 (
*** Leptoscyphus cuneifolius (Hook.) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (
** Leptoscyphus cuneifolius subsp. fragilis (J.B.Jack et Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 28, 1962 [1963] (
** subg. Austroleptoscyphus Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010 (
*** Leptoscyphus antarcticus (C.Massal.) Solari, Cryptog. Bryol. Lichénol. 7 (3): 219, 1986 (
*** Leptoscyphus australis (Gottsche, Lindenb. et Nees) R.M.Schust., J. Hattori Bot. Lab. 26: 270, 1963 (
*** Leptoscyphus belmoranus (Steph.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993 (
*** Leptoscyphus excipulatus (Steph.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993 (
*** Leptoscyphus innovatus (E.A.Hodgs.) J.J.Engel, J. Hattori Bot. Lab. 74: 33, 1993 (
*** Leptoscyphus longistipulus (Steph.) J.J.Engel, Bryologist 94 (4): 436, 1991 (
** subg. Leptoscyphus
** sect. Hexagonistipa Grolle, Nova Acta Leop. (n.ser.) 25 (161): 46, 1962 (
*** Leptoscyphus gibbosus (Taylor) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (
*** Leptoscyphus gradsteinii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 182, 2010 (
*** Leptoscyphus hexagonus (Nees) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 47, 1962 [1963] (
*** Leptoscyphus jackii (Steph.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 48, 1962 [1963] (
*** Leptoscyphus physocalyx (Hampe et Gottsche) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 33, 1858 (
*** Leptoscyphus sotiauxii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010 (
** sect. Leptoscyphus
*** Leptoscyphus aequatus (Hook.f. et Taylor) Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (
*** Leptoscyphus intermedius Grolle, Nova Acta Leop. (n.ser.) 25 (161): 32, 1962 [1963] (
*** Leptoscyphus lambinonii Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 179, 2010 (
*** Leptoscyphus obcordatus (Spruce) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 33, 1962 [1963] (
*** Leptoscyphus ovatus (Spruce) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 45, 1962 [1963] (
*** Leptoscyphus porphyrius (Nees) Grolle, Österr. Bot. Z. 117 (1): 3, 1969 (
*** Leptoscyphus porphyrius subsp. azoricus (H.Buch et Perss.) Vanderp. et Heinrichs, Taxon 59 (1): 181, 2010 (
** subg. Physoscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 51, 1962 (
** sect. Homaloscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 58, 1962 (
*** Leptoscyphus chilensis (De Not.) Hässel, J. Hattori Bot. Lab. 91: 207, 2001 (
* Leptoscyphus difficilis (Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 534, 1976 (
*** Leptoscyphus diversifolius (Gottsche) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 58, 1962 [1963] (
*** Leptoscyphus expansus (Lehm.) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 60, 1962 [1963] (
*** Leptoscyphus hedbergii (S.W.Arnell) R.M.Schust., Amer. Midl. Naturalist 62 (1): 13, 1959 (
* Leptoscyphus huidobroanus (Mont.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 33, 1858 (
*** Leptoscyphus huonicus Piippo, Acta Bot. Fenn. 131: 152, 1985 (
*** Leptoscyphus magellanicus (Gola) Hässel, J. Hattori Bot. Lab. 91: 214, 2001 (
** sect. Physoscyphus Grolle, Nova Acta Leop. (n.ser.) 25 (161): 51, 1962 (
*** Leptoscyphus amphibolius (Nees) Grolle, Nova Acta Leop. (n.ser.) 25 (161): 54, 1962 [1963] (
*** Leptoscyphus infuscatus (Mitt.) E.W.Jones ex Grolle, Nova Acta Leop. (n.ser.) 25 (161): 52, 1962 [1963] (
** subg. Spinoscyphus Vanderp., Schäf.-Verw. et D.G.Long, Taxon 59 (1): 183, 2010 (
*** Leptoscyphus cleefii Fulford, Mem. New York Bot. Gard. 11 (4): 534, 1976 (
*** Leptoscyphus spectabilis (Steph.) Grolle, J. Bryol. 11 (2): 328, 1980 [1981] (
Incertae sedis
*** Leptoscyphus autoicus (J.J.Engel et Gradst.) Vanderp. et Gradst., J. Bryol. 34 (4): 252, 2012 (
** Leptoscyphus beckettianus (Steph.) R.M.Schust. ex J.J.Engel, Nova Hedwigia 93 (3/4): 402, 2011 (
** Leptoscyphus compactus (Colenso) J.J.Engel, Nova Hedwigia 100 (3/4): 579, 2015 (
** Leptoscyphus erraticus (W.Martin et E.A.Hodgs.) J.J.Engel, Nova Hedwigia 99 (1/2): 168, 2014 (
*** Leptoscyphus horizontalis (Hook.) Kühnem., Revista Centro Estud. Doct. Ci. Nat. 1: 176, 1937 (
** Leptoscyphus normalis (Steph.) J.J.Engel, Nova Hedwigia 100 (3/4): 579, 2015 (
** Leptoscyphus physanthus (Hook.f. et Taylor) J.J.Engel, Nova Hedwigia 99 (1/2): 168, 2014 (
** Leptoscyphus subemarginatus (Hook.f. et Taylor) J.J.Engel, Bryologist 94 (4): 436, 1991 (
*** Leptoscyphus trapezoïdes (Mont.) L.Söderstr., Phytotaxa 112 (1): 27, 2013 (
*** Lophocolea (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
*** Lophocolea aberrans Lindenb. et Gottsche, Syn. Hepat. 5: 696, 1847 (
*** Lophocolea aequifolia Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 55, 1836 (
* Lophocolea angustistipula Steph., Sp. Hepat. (Stephani) 6: 260, 1922 (
*** Lophocolea anisoloba (J.J.Engel et Glenny) L.Söderstr., Phytotaxa 112 (1): 25, 2013 (
** Lophocolea anomala Steph., Sp. Hepat. (Stephani) 6: 300, 1922 (
** Lophocolea anomoda (Mont.) Steph., Hedwigia 32 (5): 327, 1893 (
** Lophocolea apalachicola R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 195, 1980 (
*** Lophocolea aperticaulis (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 25, 2013 (
** Lophocolea aphelophylla (Hässel) Váňa, Phytotaxa 112 (1): 25, 2013 (
** Lophocolea apophylla (Hässel) Váňa, Phytotaxa 112 (1): 25, 2013 (
*** Lophocolea appalachiana R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 208, 1980 (
** Lophocolea ascensionis Steph., Sp. Hepat. (Stephani) 6: 261, 1922 (
** Lophocolea asperrima Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 949 (129), 1906 (
** Lophocolea atra Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 167, 1922 [1923] (
*** Lophocolea attenuata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 34, 1900 (
*** Lophocolea australis Gottsche, Linnaea 28 (5): 553, 1856 [1857] (
** Lophocolea autoica Steph., Sp. Hepat. (Stephani) 6: 262, 1922 (
** Lophocolea baldwinii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 950 (130), 1906 (
** Lophocolea bartlettii H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 506, 1963 (
** Lophocolea bewsii (Sim) Grolle, Trans. Brit. Bryol. Soc. 3 (4): 588, 1959 (
** Lophocolea bicuspidata Steph., Sp. Hepat. (Stephani) 6: 263, 1922 (
*** Lophocolea bidentata (L.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
** Lophocolea bifidistipula Steph., Sp. Hepat. (Stephani) 6: 264, 1922 (
*** Lophocolea bispinosa (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 162, 1845 (
** Lophocolea bootanensis Steph., Sp. Hepat. (Stephani) 6: 265, 1922 (
** Lophocolea boulyana Steph., Sp. Hepat. (Stephani) 6: 264, 1922 (
** Lophocolea bowiena Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 117, 1914 (
*** Lophocolea brookwoodiana Paton et Sheahan, J. Bryol. 28 (3): 163, 2006 (
** Lophocolea caespitans Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 949 (129), 1906 (
*** Lophocolea calcarea Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 884 (109), 1906 (
** Lophocolea caledonica Steph., Sp. Hepat. (Stephani) 6: 267, 1922 (
*** Lophocolea canaliculata (Gottsche, Lindenb. et Nees) Steph., Bull. Herb. Boissier (sér. 2) 6 (9): 786 (86), 1906 (
*** Lophocolea canaliculata var. concava (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (
** Lophocolea cervicornis Steph., Biblioth. Bot. 87 (2): 219, 1916 (
** Lophocolea ciliifera Steph., Bull. Herb. Boissier (sér. 2) 6 (8): 660 (76), 1906 (
* Lophocolea coadunata (Sw.) Mont., Voy. Amér. Mérid., Bot. 7 (1): 76, 1839 (
** Lophocolea concreta Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 350, 1845 (
** Lophocolea convexula Mitt., Fl. vit.: 405, 1871 [1873] (
** Lophocolea corticola Steph., Sp. Hepat. (Stephani) 6: 268, 1922 (
*** Lophocolea decurrens Herzog, Trans. & Proc. Roy. Soc. New Zealand 65 (3): 352, 1936 (
* Lophocolea deningeri Herzog, Beih. Bot. Centralbl. 38 (2): 321, 1921 (
** Lophocolea dentiflora Steph., Bull. Herb. Boissier (sér. 2) 6 (7): 550 (64), 1906 (
** Lophocolea difformis Nees, Syn. Hepat. 2: 166, 1845 (
** Lophocolea discedens (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 2: 167, 1845 (
* Lophocolea dusenii Steph., Cat. Afr. Pl. (Hiern) 2 (2): 314, 1901 (
*** Lophocolea erosa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (
*** Lophocolea excisifolia Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 118, 1914 (
*** Lophocolea fertilis (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (
** Lophocolea flavicans Steph., Sp. Hepat. (Stephani) 6: 300, 1922 (
*** Lophocolea floribunda Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 886 (111), 1906 (
** Lophocolea foliicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 428, 1885 (
** Lophocolea fragillima Steph., Sp. Hepat. (Stephani) 6: 273, 1922 (
*** Lophocolea fragmentissima R.M.Schust., Phytologia 39 (4): 245, 1978 (
*** Lophocolea fragrans (Moris et De Not.) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 166, 1845 (
** Lophocolea glaziovii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 961 (141), 1906 (
** Lophocolea gollanii (Steph.) Váňa, Phytotaxa 112 (1): 26, 2013 (
** Lophocolea granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 126, 1864 (
** Lophocolea griffithiana Steph., Sp. Hepat. (Stephani) 6: 274, 1922 (
** Lophocolea hahnii Steph., Bull. Herb. Boissier (sér. 2) 6 (8): 660 (76), 1906 (
** Lophocolea haskarliana Gottsche, Syn. Hepat. 2: 153, 1845 (
*** Lophocolea hattorii (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (
** Lophocolea hawaica Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 945 (125), 1906 (
** Lophocolea heterodonta Steph., Sp. Hepat. (Stephani) 6: 275, 1922 (
** Lophocolea heteromorpha Steph., Sp. Hepat. (Stephani) 6: 275, 1922 (
*** Lophocolea heterophylla (Schrad.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
** Lophocolea heterophylla subsp. cladogyna R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 223, 1980 (
** Lophocolea horikawana S.Hatt., Bull. Tokyo Sci. Mus. 11: 50, 1944 (
** Lophocolea howeana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 118, 1914 (
* Lophocolea humifusa (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 695, 1847 (
** Lophocolea humistrata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 701, 1847 (
** Lophocolea itoana Inoue, J. Jap. Bot. 31 (11): 340, 1956 (
** Lophocolea javanica Schiffn., Hep. Fl. Buitenzorg: 178, 1900 (
** Lophocolea koponenii (Piippo) Váňa, Phytotaxa 112 (1): 26, 2013 (
** Lophocolea kurzii Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 296, 1864 (
* Lophocolea kurzii var. siamensis N.Kitag., Acta Phytotax. Geobot. 30 (1/3): 33, 1979 (
** Lophocolea laceristipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922 (
** Lophocolea latistipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922 (
*** Lophocolea lauterbachii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 938 (118), 1906 (
* Lophocolea laxissima Herzog, Ann. Bryol. 5: 77, 1932 (
** Lophocolea ledermannii Steph., Sp. Hepat. (Stephani) 6: 300, 1922 (
*** Lophocolea lenta (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 162, 1845 (
*** Lophocolea leptantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 694, 1847 (
*** Lophocolea liebmanniana Gottsche, Mexik. Leverm.: 113, 1863 (
** Lophocolea lindmannii Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 960 (140), 1906 (
*** Lophocolea longiciliata Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 96, 1952 (
** Lophocolea lucida (Spreng.) Mont., Voy. Amér. Mérid., Bot. 7 (2): 78, 1839 (
** Lophocolea madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 344, 1882 (
** Lophocolea magna (Udar et V.Nath) Váňa, Phytotaxa 183 (4): 291, 2014 (
*** Lophocolea mediinfrons (J.J.Engel et Braggins) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (
** Lophocolea micronesica Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 4, 1968 (
* Lophocolea microstipula Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 43, 1900 (
*** Lophocolea minor Nees, Naturgesch. Eur. Leberm. 2: 330, 1836 (
** Lophocolea minutistipula Steph., Sp. Hepat. (Stephani) 6: 283, 1922 (
* Lophocolea mollis (Nees) Nees, Syn. Hepat. 2: 158, 1845 (
* Lophocolea morobeana Piippo, Acta Bot. Fenn. 131: 160, 1985 (
** Lophocolea muhavurensis (S.W.Arnell) S.W.Arnell ex Pócs, Acta Bot. Acad. Sci. Hung. 25 (3/4): 227, 1979 [1980] (
*** Lophocolea muricata (Lehm.) Nees, Syn. Hepat. 2: 169, 1845 (
* Lophocolea muricata var. major Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 10, 1893 (
** Lophocolea nakajimae S.Hatt. et Inoue, J. Hattori Bot. Lab. 21: 221, 1959 (
*** Lophocolea novae-zeelandiae (Lehm. et Lindenb.) Nees, Syn. Hepat. 2: 168, 1845 (
*** Lophocolea novae-zeelandiae var. meridionalis (Steph.) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (
** Lophocolea orbigniana Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 55, 1836 (
** Lophocolea papulimarginata H.A.Mill., Phytologia 47 (4): 323, 1981 (
** Lophocolea parca (Gottsche) Fulford et Sharp, Mem. New York Bot. Gard. 63: 19, 1990 (
** Lophocolea parva Steph., Sp. Hepat. (Stephani) 6: 287, 1922 (
*** Lophocolea parvispinea (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 26, 2013 (
** Lophocolea parvistipula Steph., Sp. Hepat. (Stephani) 6: 287, 1922 (
*** Lophocolea patulistipa Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 50, 1911 (
*** Lophocolea perpusilla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 163, 1845 (
** Lophocolea piacenzai (Gola) Váňa, Phytotaxa 112 (1): 26, 2013 (
** Lophocolea pilistipula Steph., Sp. Hepat. (Stephani) 6: 288, 1922 (
* Lophocolea pinnatistipula Steph., Biblioth. Bot. 87 (2): 220, 1916 (
** Lophocolea platensis C.Massal., Atti Accad. Sci. Med. Nat. Ferrara 80 (3/4): 12, 1906 (
** Lophocolea purpurea Steph., Sp. Hepat. (Stephani) 6: 289, 1922 (
** Lophocolea pusilla Steph., Sp. Hepat. (Stephani) 6: 290, 1922 (
** Lophocolea randii S.W.Arnell, Svensk Bot. Tidskr. 47 (3): 420, 1953 (
** Lophocolea rara Steph., Sp. Hepat. (Stephani) 6: 290, 1922 (
** Lophocolea rectangularis Herzog, Rev. Bryol. Lichénol. 23 (1/2): 43, 1954 (
** Lophocolea rectangulata Mitt., Fl. vit.: 404, 1871 [1873] (
*** Lophocolea rupicola Steph., Bull. Herb. Boissier (sér. 2) 6 (10): 874 (99), 1906 (
*** Lophocolea sabuletorum (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 697, 1847 (
** Lophocolea salacensis Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 943 (123), 1906 (
** Lophocolea savesiana Steph., Bull. Herb. Boissier (sér. 2) 6 (11): 942 (122), 1906 (
*** Lophocolea semiteres (Lehm.) Mitt., J. Linn. Soc., Bot. 16 (91): 188, 1877 (
*** Lophocolea semiteres var. retusa (J.J.Engel) L.Söderstr., Phytotaxa 112 (1): 27, 2013 (
** Lophocolea serrata Mitt., St. Helena: 368, 1875 (
** Lophocolea siamensis Steph., Sp. Hepat. (Stephani) 6: 293, 1922 (
** Lophocolea sikkimensis (Steph.) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 164, 1958 [1959] (
* Lophocolea silvestris Gottsche, Abh. Naturwiss. Vereins Bremen 7: 345, 1882 (
** Lophocolea steetziae De Not., Epat. Borneo: 20, 1874 (
*** Lophocolea striatella (C.Massal.) Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 13, 1890 (
** Lophocolea subbidentata Herzog, Rev. Bryol. Lichénol. 23 (1/2): 43, 1954 (
** Lophocolea subcostata Steph., Sp. Hepat. (Stephani) 6: 295, 1922 (
*** Lophocolea subporosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 137, 1854 (
** Lophocolea subporosa var. inflexifolia (Steph.) L.Söderstr., Phytotaxa 112 (1): 27, 2013 (
** Lophocolea subviridis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 699, 1847 (
** Lophocolea sumatrana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 195, 1900 [1901] (
*** Lophocolea sylvatica Mitt., Rep. Challenger, Bot. 1 (3, 1): 84, 1884 (
** Lophocolea tenera Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 79, 1876 [1877] (
** Lophocolea tenerrima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 439, 1885 (
** Lophocolea teptepensis Piippo, Acta Bot. Fenn. 131: 163, 1985 (
*** Lophocolea textilis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 696, 1847 (
** Lophocolea textiloidea J.J.Engel, Phytologia 41 (5): 311, 1979 (
*** Lophocolea trichocoleoides (Glenny, J.J.Engel et He-Nygrén) L.Söderstr., Phytotaxa 112 (1): 27, 2013 (
** Lophocolea tricuspidata Herzog, Rev. Bryol. Lichénol. 11 (1): 17, 1938 [1939] (
*** Lophocolea tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 19, 1958 (
* Lophocolea undulata Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 351, 1845 (
*** Lophocolea villosa Mitt., Sp. Hepat. (Stephani) 6: 299, 1922 (
** Lophocolea wacei (S.W.Arnell ex J.J.Engel et Váňa) Váňa et L.Söderstr., Phytotaxa 112 (1): 27, 2013 (
*** Lophocolea wambana Piippo, Acta Bot. Fenn. 131: 163, 1985 (
** Lophocolea werthii (J.J.Engel et R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 27, 2013 (
** Lophocolea widgrenii Steph., Bull. Herb. Boissier (sér. 2) 7 (1): 66 (154), 1907 (
** Otoscyphus J.J.Engel, Bardat et Thouvenot, Cryptog. Bryol. 33 (3): 280, 2012 (
*** Otoscyphus crassicaulis (Steph.) J.J.Engel, Bardat et Thouvenot, Cryptog. Bryol. 33 (3): 280, 2012 (
*** Pachyglossa Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 150, 1958 [1959] (
*** Pachyglossa austrigena (Hook.f. et Taylor) L.Söderstr., Phytotaxa 112 (1): 24, 2013 (
*** Pachyglossa austrigena subsp. okaritana (Steph.) L.Söderstr., Phytotaxa 112 (1): 24, 2013 (
*** Pachyglossa boveana (C.Massal.) L.Söderstr., Phytotaxa 112 (1): 24, 2013 (
** Pachyglossa dissitifolia Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 155, 1958 [1959] (
* Pachyglossa exilis (Herzog et Grolle) Hässel et Solari, Transecta botánica de la Patagonia austral: 324, 1985 (
** Pachyglossa fissa (Mitt.) Herzog et Grolle, Rev. Bryol. Lichénol. 28 (3/4): 346, 1959 [1960] (
*** Pachyglossa gottscheoides (Besch. et C.Massal.) L.Söderstr., Phytotaxa 112 (1): 25, 2013 (
** Pachyglossa grolleana Váňa, Cryptog. Bryol. 26 (1): 86, 2005 (
** Pachyglossa otiphylla (Hook.f. et Taylor) Váňa, Phytotaxa 112 (1): 25, 2013 (
** Pachyglossa spegazziniana (C.Massal.) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 159, 1958 [1959] (
** Pachyglossa tenacifolia (Hook.f. et Taylor) Herzog et Grolle, Rev. Bryol. Lichénol. 27 (3/4): 153, 1958 [1959] (
** Perdusenia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 7 (2): 11, 1989 (
** Perdusenia rheophila Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 7 (2): 11, 1989 (
** Pigafettoa C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 237, 1885 (
** Pigafettoa crenulata C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 237, 1885 (
** Platycaulis R.M.Schust., Phytologia 39 (4): 245, 1978 (
*** Platycaulis renifolius R.M.Schust., Phytologia 39 (4): 245, 1978 (
*** Stolonivector J.J.Engel, J. Hattori Bot. Lab. 69: 80, 1991 (
** Stolonivector clasmatocoleoides J.J.Engel, Nova Hedwigia 88 (3/4): 339, 2009 (
*** Stolonivector fiordlandiae (E.A.Hodgs.) J.J.Engel, J. Hattori Bot. Lab. 69: 82, 1991 (
** Stolonivector fiordlandiae var. nodulosus J.J.Engel, Nova Hedwigia 93 (3/4): 403, 2011 (
*** Stolonivector gremmenii (Váňa) Váňa, Phytotaxa 112 (1): 28, 2013 (
** Stolonivector obtusilobus J.J.Engel, Nova Hedwigia 88 (3/4): 337, 2009 (
** Stolonivector waipouensis J.J.Engel, J. Hattori Bot. Lab. 93: 70, 2003 (
** Xenocephalozia R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (
** Xenocephalozia navicularis (Steph.) R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (
** Dendromastigophora R.M.Schust., Mem. New York Bot. Gard. 45: 738, 1987 (
*** Dendromastigophora flagellifera (Hook.) R.M.Schust., Mem. New York Bot. Gard. 45: 738, 1987 (
*** Mastigophora Nees, Naturgesch. Eur. Leberm. 3: 89, 1838 (
* Mastigophora appendiculata Steph., Sp. Hepat. (Stephani) 6: 368, 1922 (
* Mastigophora attenuata (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (
** Mastigophora caledonica Steph., Rev. Bryol. 35 (2): 31, 1908 (
*** Mastigophora diclados (Brid. ex F.Weber) Nees, Naturgesch. Eur. Leberm. 3: 18, 1838 (
** Mastigophora diclados var. borneensis (De Not.) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 251, 1893 (
** Mastigophora diclados var. ramentifissa Herzog, Trans. Brit. Bryol. Soc. 1 (4): 315, 1950 (
* Mastigophora diclados var. villosa Herzog, Ann. Bryol. 5: 82, 1932 (
* Mastigophora guineensis Steph., Sp. Hepat. (Stephani) 6: 369, 1923 (
* Mastigophora humillima (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (
* Mastigophora pyramidana Steph., Sp. Hepat. (Stephani) 6: 369, 1923 (
** Mastigophora sepikiana Piippo, Ann. Bot. Fenn. 23 (1): 2, 1986 (
** Mastigophora tuberculata D.H.Mill. et H.A.Mill., J. Hattori Bot. Lab. 75: 181, 1994 (
* Mastigophora valida Steph., Sp. Hepat. (Stephani) 6: 369, 1923 (
** Mastigophora viridula (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (
*** Mastigophora woodsii (Hook.) Nees, Naturgesch. Eur. Leberm. 3: 95, 1838 (
by L. Söderström
The placement of Pedinophyllopsis in Plagiochilaceae follows
** Acrochila R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963 (
*** Acrochila biserialis (Lehm. et Lindenb.) Grolle, J. Jap. Bot. 39 (8): 236, 1964 (
** Acrochila caledonica (Steph.) Inoue, J. Jap. Bot. 42 (6): 182, 1967 (
** Chiastocaulon Carl, Flora 126: 58, 1931 (
*** Chiastocaulon dendroides (Nees) Carl, Flora 126: 59, 1931 (
** Dinckleria Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 421, 1877 (
*** Dinckleria fruticella (Hook.f. et Taylor) J.J.Engel et Heinrichs, Cryptog. Bryol. 29 (2): 194, 2008 (
*** Dinckleria pleurata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 421, 1877 (
** Pedinophyllopsis R.M.Schust. et Inoue, Phytologia 47 (4): 311, 1981 (
** Pedinophyllopsis abdita (Sull.) R.M.Schust. et Inoue, Phytologia 47 (4): 311, 1981 (
** Pedinophyllum Lindb. ex Nordst., Bot. Not. 1874: 156, 1874 (
*** Pedinophyllum autoicum (Steph.) Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (4): 575, 1966 (
*** Pedinophyllum interruptum (Nees) Kaal., Nyt Mag. Naturvidensk. 33 (1): 190, 1893 (
*** Pedinophyllum monoicum (Steph.) Grolle, Nova Hedwigia 2: 287, 1960 (
** Pedinophyllum truncatum (Steph.) Inoue, J. Hattori Bot. Lab. 23: 35, 1960 [1961] (
*** Plagiochila (Dumort.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
*** Plagiochila heteromalla (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 83, 1840 (
*** sect. Adianthoideae Lindenb., Monogr. hep. gen. Plagiochilae: xx, 1844 (
*** Plagiochila adianthoides (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 77, 1840 (
* Plagiochila adianthoides var. aspergillifera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 474, 1885 (
*** Plagiochila cristata (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 1: 33, 1839 (
*** Plagiochila grandicrista Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 931 (581), 1905 (
*** Plagiochila herminieri Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 748 (547), 1905 (
*** sect. Africanae Heinrichs, Taxon 54 (2): 319, 2005 (
*** Plagiochila barteri Mitt., J. Linn. Soc., Bot. 22 (146): 320, 1886 (
** Plagiochila barteri var. valida (Steph.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 73, 1981 (
*** Plagiochila colorans Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 116, 1911 (
*** sect. Arrectae Carl, Ann. Bryol., Suppl. 2: 52, 1931 (
* Plagiochila arnelliana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 861 (230), 1902 (
*** Plagiochila badia Mitt., Rep. Challenger, Bot. 1 (3, 1): 84, 1884 (
*** Plagiochila bidens Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 322, 1857 (
*** Plagiochila bifaria (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 5: 127, 1843 (
** Plagiochila bifaria var. rosea (R.M.Schust.) Heinrichs, Org. Divers. Evol. 4 (1/2): 112, 2004 (
*** Plagiochila chacabucensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 27, 1911 (
*** Plagiochila emeiensis Grolle et M.L.So, Bryologist 101 (2): 282, 1998 (
* Plagiochila fragilis Taylor, London J. Bot. 7: 198, 1848 (
*** Plagiochila lunata S.W.Arnell, Bot. Not. 115: 204, 1962 (
*** Plagiochila pachyloma Taylor, London J. Bot. 5: 267, 1846 (
* Plagiochila pachyloma var. elatior Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 480, 1885 (
** Plagiochila papillifolia Steph., Biblioth. Bot. 87 (2): 207, 1916 (
*** Plagiochila parviramifera Inoue, J. Hattori Bot. Lab. 46: 317, 1979 (
*** Plagiochila pseudoattenuata S.W.Arnell, Bot. Not. 115: 206, 1962 (
*** Plagiochila punctata (Taylor) Taylor, London J. Bot. 5: 261, 1846 (
*** Plagiochila renauldii Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 156 (408), 1904 (
*** Plagiochila retrorsa Gottsche, Mexik. Leverm.: 67, 1863 (
*** Plagiochila rubescens (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 46, 1840 (
*** Plagiochila sichuanensis Grolle et M.L.So, Bryologist 101 (2): 284, 1998 (
*** Plagiochila spinulosa (Dicks.) Dumort., Recueil Observ. Jungerm.: 15, 1835 (
** Plagiochila sticticola Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 198, 1856 (
*** Plagiochila stricta Lindenb., Sp. Hepat. (Lindenberg) 1: 20, 1839 (
*** Plagiochila tronadoris Herzog, Darwiniana 11 (2): 214, 1957 (
*** Plagiochila uncialis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 628, 1847 (
*** Plagiochila wilmsiana Steph., Sp. Hepat. (Stephani) 6: 240, 1921 (
* sect. Caducifoliae J.J.Engel et G.L.Merr., Nova Hedwigia 96 (3/4): 407, 2013 (
** Plagiochila caducifolia Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 71, 1971 (
* sect. Cardotiae Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 386, 1965 (
*** Plagiochila cumingiana Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 32 (404), 1904 (
*** Plagiochila denticulata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 95, 1860 [1861] (
*** Plagiochila fragillima Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 522 (326), 1903 (
*** Plagiochila pseudorenitens Schiffn., Österr. Bot. Z. 49 (4): 132, 1899 (
*** Plagiochila stevensiana Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 110 (290), 1903 (
* sect. Cobanae Carl, Ann. Bryol., Suppl. 2: 79, 1931 (
** Plagiochila cobana Steph., Sp. Hepat. (Stephani) 6: 138, 1918 (
** Plagiochila detecta M.L.So et Grolle, Nova Hedwigia 71 (3/4): 391, 2000 (
*** Plagiochila singularis Schiffn., Hep. Fl. Buitenzorg: 158, 1900 (
*** Plagiochila tagawae Inoue, J. Hattori Bot. Lab. 38: 561, 1974 (
** Plagiochila tixieri Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 87, 1975 (
*** Plagiochila zhuensis Grolle et M.L.So, Bryologist 102 (2): 200, 1999 (
*** sect. Cucullatae Schiffn., Hep. Fl. Buitenzorg: 107, 1900 (
*** Plagiochila bantamensis (Reinw., Blume et Nees) Mont., Voy. Amér. Mérid., Bot. 7 (2): 82, 1839 (
*** Plagiochila blepharophora (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 102, 1840 (
** Plagiochila chauviniana Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 34, 1849 (
** Plagiochila clavatosaccata Steph., Bull. Herb. Boissier (sér. 2) 4 (1): 25 (397), 1904 (
** Plagiochila grossispina Steph., Sp. Hepat. (Stephani) 6: 162, 1918 (
*** Plagiochila integerrima Steph., Bot. Jahrb. Syst. 8 (2): 83, 1886 (
* Plagiochila integrilobula Schiffn., Hep. Fl. Buitenzorg: 170, 1900 (
** Plagiochila johannis-winkleri Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 187, 1931 (
* Plagiochila kuhliana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 292, 1864 (
** Plagiochila kurzii Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 112 (292), 1903 (
** Plagiochila reischeckiana Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 331 (315), 1903 (
*** Plagiochila sandei Dozy ex Sande Lac., Plagiochila Sandei: 5, 1856 (
*** Plagiochila sciophila Nees, Sp. Hepat. (Lindenberg) 2-4: 100, 1840 (
** Plagiochila sciophila subsp. ciliigera (R.M.Schust.) L.Söderstr., Phytotaxa 208 (1): 84, 2015 (
* Plagiochila stephanii Schiffn., Hep. Fl. Buitenzorg: 166, 1900 (
** Plagiochila subplana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 73, 1840 (
** Plagiochila vitiensis Mitt., Bonplandia 9 (24): 367, 1861 (
*** sect. Denticulatae Schiffn., Hep. Fl. Buitenzorg: 106, 1900 (
*** Plagiochila alternans Lindenb. et Gottsche, Syn. Hepat. 5: 648, 1847 (
** Plagiochila ansata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 649, 1847 (
*** Plagiochila banksiana Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 329, 1857 (
** Plagiochila banksiana var. echinophora Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 62, 1971 (
** Plagiochila chonotica Taylor, London J. Bot. 5: 260, 1846 (
** Plagiochila equitans Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 331, 1857 (
* Plagiochila fragmentissima Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 155, 1971 (
*** Plagiochila gigantea Lindenb., Sp. Hepat. (Lindenberg) 2-4: 115, 1840 (
** Plagiochila gigantea var. inermis J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 513, 2010 (
*** Plagiochila gregaria (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 654, 1847 (
** Plagiochila hookeriana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 81, 1840 (
** Plagiochila latifrons Gottsche et Hampe, Linnaea 27 (5): 553, 1854 (
** Plagiochila minutula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 652, 1847 (
*** Plagiochila nobilis Gottsche, Bot. Zeitung (Berlin) Beil. 16: 37, 1858 (
** Plagiochila obovata Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 33, 1911 (
*** Plagiochila ovata Lindenb. et Gottsche, Syn. Hepat. 5: 656, 1847 (
*** Plagiochila retrospectans Lindenb., Sp. Hepat. (Lindenberg) 5: 123, 1843 (
** Plagiochila riparia Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 34, 1911 (
** Plagiochila rutlandii Steph., Bull. Herb. Boissier (sér. 2) 4 (8): 776 (454), 1904 (
** Plagiochila subpectinata Besch. et C.Massal., Bull. Mens. Soc. Linn. Paris 1 (79): 628, 1886 (
*** sect. Durae Carl, Ann. Bryol., Suppl. 2: 123, 1931 (
** Plagiochila acanthocaulis Sull., Hooker’s J. Bot. Kew Gard. Misc. 2: 317, 1850 (
** Plagiochila angulata Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 26, 1900 (
** Plagiochila bicornuta Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 305, 1896 (
** Plagiochila crozetensis Kaal., Nyt Mag. Naturvidensk. 49 (2/3): 92, 1911 (
*** Plagiochila deltoidea Lindenb., Sp. Hepat. (Lindenberg) 5: 132, 1843 (
** Plagiochila deltoidea var. densa J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 506, 2010 (
** Plagiochila dura De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 214, 1857 (
** Plagiochila heterodonta (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 638, 1847 (
*** Plagiochila ramosissima (Hook.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 87, 1840 (
*** sect. Duseniae Carl, Ann. Bryol., Suppl. 2: 126, 1931 (
** Plagiochila dusenii Steph., Bull. Herb. Boissier (sér. 2) 4 (10): 979 (475), 1904 (
** Plagiochila elata Taylor, London J. Bot. 5: 259, 1846 (
** Plagiochila lechleri Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 325, 1857 (
** Plagiochila validissima Steph., Biblioth. Bot. 87 (2): 214, 1916 (
* sect. Flexicaules Carl, Ann. Bryol., Suppl. 2: 127, 1931 (
** Plagiochila flexicaulis Mont., Syn. Hepat. 5: 629, 1847 (
*** sect. Fruticosae Inoue, Gen. Plagiochila SE Asia: 50, 1984 (
*** Plagiochila assamica Steph., Sp. Hepat. (Stephani) 6: 125, 1917 (
** Plagiochila benitoi Inoue ex Piippo, J. Hattori Bot. Lab. 72: 122, 1992 (
*** Plagiochila frondescens (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 52, 1840 (
*** Plagiochila fruticosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 94, 1860 [1861] (
** Plagiochila mauiensis Steph., Sp. Hepat. (Stephani) 6: 184, 1921 (
*** Plagiochila pulcherrima Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 63, 1931 (
** Plagiochila tamiensis Steph., Sp. Hepat. (Stephani) 6: 235, 1921 (
*** sect. Fuscoluteae Carl, Ann. Bryol., Suppl. 2: 46, 1931 (
*** Plagiochila aerea Taylor, London J. Bot. 5: 263, 1846 (
*** Plagiochila dependula Taylor, London J. Bot. 5: 265, 1846 (
*** Plagiochila fuscolutea Taylor, London J. Bot. 5: 263, 1846 (
** Plagiochila heterophylla Lindenb., Nov. Stirp. Pug. 10: 2, 1857 (
** Plagiochila heterophylla var. beauverdii (Steph.) Heinrichs, Bryophyt. Biblioth. 58: 148, 2002 (
*** Plagiochila longiramea Steph., Biblioth. Bot. 87 (2): 204, 1916 (
*** Plagiochila paraphyllina Herzog, Hedwigia 74 (2): 89, 1934 (
*** Plagiochila rudischusteri H.Rob., Beih. Nova Hedwigia 90: 199, 1988 (
*** Plagiochila subbidentata Taylor, Ann. Mag. Nat. Hist. 20 (135): 381, 1847 (
*** Plagiochila tabinensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 862 (231), 1902 (
*** sect. Glaucescentes Carl, Ann. Bryol., Suppl. 2: 70, 1931 (
*** Plagiochila buchtiniana Steph., Sp. Hepat. (Stephani) 6: 135, 1918 (
*** Plagiochila diversifolia Lindenb. et Gottsche, Syn. Hepat. 5: 640, 1847 (
*** Plagiochila longispina Lindenb. et Gottsche, Syn. Hepat. 5: 642, 1847 (
*** sect. Hylacoetes Carl, Ann. Bryol., Suppl. 2: 50, 1931 (
*** Plagiochila amicta Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 895 (561), 1905 (
*** Plagiochila boryana Gottsche, Bull. Soc. Roy. Bot. Belgique 31: 118, 1892 (
*** Plagiochila breuteliana Lindenb., Sp. Hepat. (Lindenberg) 5: 150, 1843 (
*** Plagiochila canelensis Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 926 (576), 1905 (
*** Plagiochila cucullifolia J.B.Jack et Steph., Hedwigia 31 (1): 24, 1892 (
*** Plagiochila cucullifolia var. anomala Heinrichs et Gradst., Pl. Syst. Evol. 242 (1/4): 208, 2003 (
*** Plagiochila dimorpha Lindenb. et Gottsche, Syn. Hepat. 5: 627, 1847 (
*** Plagiochila dominicensis Taylor, London J. Bot. 5: 270, 1846 (
*** Plagiochila ecuadorica (Inoue) L.Söderstr., Phytotaxa 208 (1): 84, 2015 (
*** Plagiochila ensiformis Taylor, London J. Bot. 5: 265, 1846 (
*** Plagiochila flabelliflora Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 880 (249), 1902 (
** Plagiochila guevarii H.Rob., Bryologist 70 (1): 48, 1967 (
** Plagiochila husnotii Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 178 (506), 1905 (
*** Plagiochila macrostachya Lindenb., Sp. Hepat. (Lindenberg) 2-4: 75, 1840 (
*** Plagiochila patriciae Heinrichs et H.Anton, Bryophyt. Biblioth. 58: 107, 2002 (
*** Plagiochila superba (Nees ex Spreng.) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839 (
*** Plagiochila superba var. macrotricha (Spruce) Heinrichs, Bryophyt. Biblioth. 58: 114, 2002 (
*** Plagiochila turgida Herzog, Hedwigia 72 (6): 196, 1932 (
*** Plagiochila vincentina Lindenb., Sp. Hepat. (Lindenberg) 2-4: 39, 1840 (
* sect. Jacquinotiae Hässel, Nova Hedwigia 89 (1/2): 85, 2009 (
** Plagiochila jacquinotii Mont., Voy. Pole Sud, Bot. 1: 273, 1845 (
* sect. Kaalaasiae Carl, Ann. Bryol., Suppl. 2: 103, 1931 (
** Plagiochila amboynensis Taylor, London J. Bot. 5: 260, 1846 (
* sect. Longiflorae Carl, Ann. Bryol., Suppl. 2: 130, 1931 (
** Plagiochila longiflora Mont., Syn. Hepat. 5: 651, 1847 (
* sect. Oligodontes Carl, Ann. Bryol., Suppl. 2: 130, 1931 (
** Plagiochila lophocoleoides Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 348, 1845 (
*** sect. Peculiares Schiffn., Hep. Fl. Buitenzorg: 107, 1900 (
** Plagiochila aspericaulis Grolle et M.L.So, Syst. Bot. 24 (3): 307, 1999 (
** Plagiochila caulimammillosa Grolle et M.L.So, J. Bryol. 20 (1): 42, 1998 (
*** Plagiochila devexa Steph., Bull. Herb. Boissier (sér. 2) 3 (4): 340 (324), 1903 (
*** Plagiochila durelii Schiffn., Österr. Bot. Z. 49 (4): 131, 1899 (
** Plagiochila durelii subsp. guizhouensis Grolle et M.L.So, Syst. Bot. 24 (3): 304, 1999 (
** Plagiochila grollei Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 384, 1965 (
** Plagiochila hoei Inoue, J. Hattori Bot. Lab. 40: 421, 1976 (
** Plagiochila hyalodermica Grolle et M.L.So, Bryologist 100 (4): 470, 1997 (
** Plagiochila magna Inoue, J. Hattori Bot. Lab. 28: 216, 1965 (
** Plagiochila paraphyllosa Grolle et M.L.So, Syst. Bot. 24 (3): 298, 1999 (
** Plagiochila peculiaris Schiffn., Hep. Fl. Buitenzorg: 157, 1900 (
*** Plagiochila perserrata Herzog, Symb. Sin. 5: 19, 1930 (
** Plagiochila philippinensis Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 526 (330), 1903 (
*** Plagiochila pseudopoeltii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 382, 1965 (
*** Plagiochila renitens (Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 90, 1840 (
** Plagiochila semidecurrens (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 5: 142, 1843 (
** Plagiochila semidecurrens var. alaskana (A.Evans) Inoue, J. Hattori Bot. Lab. 28: 216, 1965 (
*** Plagiochila vexans Schiffn. ex Steph., Sp. Hepat. (Stephani) 6: 237, 1921 (
*** Plagiochila zangii Grolle et M.L.So, Bryologist 100 (4): 467, 1997 (
*** Plagiochila zonata Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 225, 1894 (
*** sect. Plagiochila
** Plagiochila arctica Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 41, 1906 (
* Plagiochila arctica var. intermedia R.M.Schust., Amer. Midl. Naturalist 62 (1): 152, 1959 (
*** Plagiochila asplenioides (L.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
*** Plagiochila bischleriana Grolle et M.L.So, Cryptog. Bryol. Lichénol. 18 (3): 191, 1997 (
** Plagiochila britannica Paton, J. Bryol. 10 (3): 245, 1979 (
*** Plagiochila chinensis Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 223, 1894 (
*** Plagiochila circumdentata Steph., Bull. Herb. Boissier (sér. 2) 4 (8): 778 (456), 1904 (
** Plagiochila circumdentata var. carinata J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 511, 2010 (
** Plagiochila circumserrata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 27, 1979 (
** Plagiochila columbiana A.Evans, Bot. Gaz. 21 (4): 189, 1896 (
*** Plagiochila delavayi Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 224, 1894 (
*** Plagiochila elegans Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861] (
*** Plagiochila hakkodensis Steph., Bull. Herb. Boissier 5 (2): 103, 1897 (
** Plagiochila korthalsiana Molk., Ned. Kruidk. Arch. 3: 416, 1854 [1855] (
** Plagiochila loriana Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 608 (354), 1903 (
** Plagiochila microdentata M.L.So, New Zealand J. Bot. 38 (3): 426, 2000 (
** Plagiochila mundaliensis Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 536 (340), 1903 (
** Plagiochila orbicularis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 3: 26, 1948 [1950] (
*** Plagiochila ovalifolia Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 193, 1891 (
*** Plagiochila porelloides (Torr. ex Nees) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 61, 1840 (
** Plagiochila porelloides var. norvegica (H.H.Blom et Holten) Schumacker et Váňa, Identif. keys liverw. hornw. Europe: 131, 2005 (
** Plagiochila porelloides var. subarctica (Jørg.) Lammes, Fl. Fenn. 6: 54, 1977 (
** Plagiochila schofieldiana Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 15 (1): 183, 1972 (
*** Plagiochila secretifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861] (
** Plagiochila spinosa M.L.So, New Zealand J. Bot. 38 (3): 428, 2000 (
** Plagiochila sumatrana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 183, 1900 [1901] (
** Plagiochila taiwanensis Inoue, Bull. Natl. Sci. Mus. Tokyo, B 8 (4): 136, 1982 (
*** Plagiochila trapezoidea Lindenb., Sp. Hepat. (Lindenberg) 2-4: 112, 1840 (
** Plagiochila uniformis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861] (
** sect. Poeltiae Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 394, 1965 (
*** Plagiochila biondiana C.Massal., Hepat. Shen-si: 15, 1897 (
*** Plagiochila carringtonii (Balf. ex Carrington) Grolle, Trans. Brit. Bryol. Soc. 4 (4): 656, 1964 (
** Plagiochila carringtonii subsp. lobuchensis Grolle, Trans. Brit. Bryol. Soc. 4 (4): 660, 1964 (
*** Plagiochila duthiana Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 527 (331), 1903 (
** Plagiochila erlangensis M.L.So, Haussknechtia, Beih. 9: 350, 1999 (
*** Plagiochila poeltii Inoue et Grolle, Trans. Brit. Bryol. Soc. 4 (4): 656, 1964 (
*** Plagiochila recurvata (W.E.Nicholson) Grolle, Trans. Brit. Bryol. Soc. 4 (4): 654, 1964 (
*** Plagiochila retusa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 96, 1860 [1861] (
*** Plagiochila wallichiana Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 523 (327), 1903 (
** Plagiochila wangii Inoue, J. Jap. Bot. 37 (6): 187, 1962 (
** Plagiochila yulungensis Piippo, Ann. Bot. Fenn. 34 (4): 283, 1997 (
*** sect. Rutilantes Carl, Ann. Bryol., Suppl. 2: 83, 1931 (
*** Plagiochila bicuspidata Gottsche, Mexik. Leverm.: 43, 1863 (
** Plagiochila caduciloba H.L.Blomq., Bryologist 42 (5): 114, 1939 (
** Plagiochila chimborazensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 469, 1885 (
* Plagiochila cuneata Lindenb. et Gottsche, Syn. Hepat. 5: 632, 1847 (
*** Plagiochila debilis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861] (
*** Plagiochila deflexa Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 192, 1856 (
*** Plagiochila defolians Grolle et M.L.So, Syst. Bot. 23 (4): 459, 1998 [1999] (
*** Plagiochila exigua (Taylor) Taylor, London J. Bot. 5: 264, 1846 (
** Plagiochila fracta Pócs, Phytotaxa 195 (2): 183, 2015 (
*** Plagiochila ghatiensis Steph., Sp. Hepat. (Stephani) 6: 159, 1918 (
*** Plagiochila grossa Grolle et M.L.So, Syst. Bot. 23 (4): 461, 1998 [1999] (
*** Plagiochila gymnocalycina (Lehm. et Lindenb.) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839 (
** Plagiochila gymnocalycina var. surinamensis (Molk.) Heinrichs et D.S.Rycroft, J. Hattori Bot. Lab. 100: 139, 2006 (
** Plagiochila loriloba Herzog ex Carl, Ann. Bryol., Suppl. 2: 47, 1931 (
** Plagiochila maderensis Gottsche ex Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 350 (426), 1904 (
** Plagiochila oresitropha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 467, 1885 (
*** Plagiochila pectinata Lindenb., Sp. Hepat. (Lindenberg) 1: 14, 1839 (
*** Plagiochila rutilans Lindenb., Sp. Hepat. (Lindenberg) 2-4: 47, 1840 (
*** Plagiochila rutilans var. moritziana (Lindenb. et Gottsche) Heinrichs, Bryologist 105 (2): 197, 2002 (
*** Plagiochila rutilans var. standleyi (Herzog ex Carl) Heinrichs et D.S.Rycroft, Bryologist 104 (3): 357, 2001 (
* Plagiochila steyermarkii H.Rob., Bryologist 68 (1): 93, 1965 (
*** Plagiochila trichostoma Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 113, 1864 (
* sect. Strombifoliae Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 130, 1971 (
** Plagiochila strombifolia Taylor, Nov. Stirp. Pug. 8: 5, 1844 (
*** sect. Tayloriae Carl, Ann. Bryol., Suppl. 2: 140, 1931 (
*** Plagiochila annotina Lindenb., Sp. Hepat. (Lindenberg) 1: 34, 1839 (
*** Plagiochila baylisii Inoue et R.M.Schust., J. Hattori Bot. Lab. 34: 150, 1971 (
*** Plagiochila bazzanioides J.J.Engel et G.L.Merr., Novon 9 (1): 29, 1999 (Engel and Smith Merrill 1999b).
*** Plagiochila chenii Grolle et M.L.So, Syst. Bot. 25 (1): 6, 2000 (
*** Plagiochila circinalis (Lehm. et Lindenb.) Lehm., Sp. Hepat. (Lindenberg) 5: 124, 1843 (
** Plagiochila circinalis var. hemicardia (Hook.f. et Taylor) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 512, 2010 (
*** Plagiochila colensoi Hook.f. et Taylor, London J. Bot. 5: 269, 1846 (
** Plagiochila colensoi var. quinquespina (Steph.) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 504, 2010 (
** Plagiochila corticola Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 224, 1894 (
** Plagiochila fasciculata Lindenb., Sp. Hepat. (Lindenberg) 1: 7, 1839 (
*** Plagiochila fusca Sande Lac., Ned. Kruidk. Arch. 3: 417, 1854 [1855] (
** Plagiochila fuscella (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 648, 1847 (
** Plagiochila fuscella var. novae-zelandiae (E.A.Hodgs.) J.J.Engel et G.L.Merr., Nova Hedwigia 89 (3/4): 294, 2009 (
*** Plagiochila gracilis Lindenb. et Gottsche, Syn. Hepat. 5: 632, 1847 (
*** Plagiochila gymnoclada Sande Lac., Plagiochila Sandei: 6, 1856 (
*** Plagiochila himalayana Schiffn., Österr. Bot. Z. 49 (4): 131, 1899 (
** Plagiochila incurvicolla (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 651, 1847 (
** Plagiochila incurvicolla var. lonchoscypha (Herzog) J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 505, 2010 (
** Plagiochila microdictyon Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 131, 1854 (
** Plagiochila monospiris Inoue et Grolle, J. Hattori Bot. Lab. 36: 489, 1972 [1973] (
*** Plagiochila nitens Inoue, Willdenowia 18 (2): 561, 1989 (
** Plagiochila pseudocapillaris Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (3): 302, 1968 (
*** Plagiochila pseudofirma Herzog, Symb. Sin. 5: 17, 1930 (
** Plagiochila radiculosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 132, 1854 (
** Plagiochila spathulifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 96, 1860 [1861] (
*** Plagiochila stephensoniana Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 133, 1854 (
*** sect. Trabeculatae S.Hatt. ex Inoue, J. Hattori Bot. Lab. 20: 75, 1958 (
** Plagiochila austinii A.Evans, Rhodora 16 (184): 68, 1914 (
*** Plagiochila flexuosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 94, 1860 [1861] (
** Plagiochila sullivantii Gottsche, Bot. Gaz. 21 (4): 191, 1896 (
** Plagiochila sullivantii var. spinigera R.M.Schust., Amer. Midl. Naturalist 62 (2): 323, 1959 (
*** Plagiochila trabeculata Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 103 (283), 1903 (
*** sect. Vagae Lindenb., Monogr. hep. gen. Plagiochilae: xv, 1844 (
*** Plagiochila abietina (Nees) Mont. et Nees, Voy. Amér. Mérid., Bot. 7 (2): 81, 1839 (
** Plagiochila abrupta Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 106, 1840 (
** Plagiochila aequatorialis Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 334, 1857 (
*** Plagiochila africana Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 973 (263), 1902 (
*** Plagiochila akiyamae Inoue, Bull. Natl. Sci. Mus. Tokyo, B 12 (3): 73, 1986 (
*** Plagiochila angusta Lindenb., Sp. Hepat. (Lindenberg) 5: 148, 1843 (
*** Plagiochila angustitexta Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 977 (267), 1902 (
*** Plagiochila arbuscula (Brid. ex Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 1: 23, 1839 (
** Plagiochila arbuscula var. rekohuensis J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 509, 2010 (
* Plagiochila aspera Steph., Sp. Hepat. (Stephani) 6: 125, 1917 (
** Plagiochila aspleniformis R.M.Schust., Amer. Midl. Naturalist 63 (1): 51, 1960 (
** Plagiochila beddomei Steph., Bull. Herb. Boissier (sér. 2) 3 (10): 876 (361), 1903 (
*** Plagiochila boivinii Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 987 (277), 1902 (
*** Plagiochila bryopteroides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 499, 1885 (
** Plagiochila contigua Gottsche, Mexik. Leverm.: 30, 1863 (
*** Plagiochila corrugata (Nees) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 52, 1836 (
*** Plagiochila cuspidata Steph., Sp. Hepat. (Stephani) 6: 144, 1918 (
** Plagiochila cymata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 95, 1975 (
** Plagiochila deflexirama Taylor, London J. Bot. 5: 262, 1846 (
*** Plagiochila dichotoma (P.Beauv.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 53, 1836 (
* Plagiochila dichotoma var. fluitans Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 490, 1885 (
*** Plagiochila dissecta Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 600 (346), 1903 (
*** Plagiochila disticha (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 107, 1840 (
** Plagiochila distinctifolia Lindenb., Sp. Hepat. (Lindenberg) 1: 17, 1839 (
*** Plagiochila divergens Steph., Hedwigia 30 (6): 268, 1891 (
*** Plagiochila drepanophylla Sande Lac., Syn. hepat. jav.: 103, 1856 [1857] (
* Plagiochila drepanophylla var. minor Gottsche, Abh. Naturwiss. Vereins Bremen 7: 341, 1882 (
*** Plagiochila effusa Steph., Bot. Jahrb. Syst. 20 (3): 310, 1895 (
* Plagiochila effusa var. decurrens Steph., Bot. Jahrb. Syst. 20 (3): 310, 1895 (
*** Plagiochila ericicola Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 590 (441), 1904 (
** Plagiochila exinnovata Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 600 (346), 1903 (
** Plagiochila fastigiata Lindenb. et Gottsche, Syn. Hepat. 5: 657, 1847 (
*** Plagiochila flabellata Steph., Bot. Jahrb. Syst. 8 (2): 82, 1886 (
** Plagiochila floridana A.Evans, Bot. Gaz. 21 (4): 190, 1896 (
*** Plagiochila fordiana Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 104 (284), 1903 (
** Plagiochila francana Steph., Sp. Hepat. (Stephani) 6: 157, 1918 (
*** Plagiochila furcifolia Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 194, 1891 (
*** Plagiochila fusifera Taylor, London J. Bot. 5: 268, 1846 (
*** Plagiochila hampeana Gottsche, Bot. Zeitung (Berlin) Beil. 16: 38, 1858 (
* Plagiochila heterospina Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 128, 1914 (
* Plagiochila heterostipa Steph., Hedwigia 31 (3): 129, 1892 (
** Plagiochila incerta Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 324, 1857 (
*** Plagiochila indica Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 532 (336), 1903 (
** Plagiochila invisa (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 513, 1980 (
*** Plagiochila javanica (Sw.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 52, 1836 (
*** Plagiochila junghuhniana Sande Lac., Ned. Kruidk. Arch. 3: 416, 1854 [1855] (
*** Plagiochila khasiana Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 95, 1860 [1861] (
** Plagiochila kiaeri Gottsche, Abh. Naturwiss. Vereins Bremen 7: 341, 1882 (
** Plagiochila kiaeri var. capensis (Steph.) M.Wigginton et Grolle, Bryophyt. Biblioth. 50: 182, 1996 (
** Plagiochila kiaeri var. myriocarpa (Pearson) Pócs, J. E. Afr. Nat. Hist. 96 (1): 38, 2007 (
*** Plagiochila kunmingensis Piippo, Ann. Bot. Fenn. 34 (4): 281, 1997 (
** Plagiochila kurokawae Inoue, J. Hattori Bot. Lab. 32: 104, 1969 (
*** Plagiochila laetevirens Lindenb., Sp. Hepat. (Lindenberg) 2-4: 101, 1840 (
** Plagiochila lamellistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 491, 1885 (
*** Plagiochila lastii Mitt., J. Linn. Soc., Bot. 22 (146): 320, 1886 (
** Plagiochila latifolia Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 742 (541), 1905 (
* Plagiochila loloënsis Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 166 (418), 1904 (
** Plagiochila manillana Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 189, 1856 (
** Plagiochila massalongoana Schiffn., Hep. Fl. Buitenzorg: 136, 1900 (
** Plagiochila mastigophoroides Inoue, J. Hattori Bot. Lab. 32: 99, 1969 (
** Plagiochila metcalfii Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 533 (337), 1903 (
** Plagiochila micropteryx Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 107, 1864 (
** Plagiochila miradorensis Gottsche, Mexik. Leverm.: 31, 1863 (
** Plagiochila miradorensis var. convoluta R.M.Schust., Amer. Midl. Naturalist 63 (1): 113, 1960 (
* Plagiochila moenkemeyeri Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 160 (412), 1904 (
*** Plagiochila montagnei Nees, Ann. Sci. Nat. Bot. (sér. 2) 5: 53, 1836 (
** Plagiochila morobensis Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 203, 1989 (
** Plagiochila multipinnula Herzog et S.Hatt., J. Hattori Bot. Lab. 14: 36, 1955 (
*** Plagiochila neckeroidea Mitt., Trans. Linn. Soc. London 23 (1): 57, 1860 (
*** Plagiochila nepalensis Lindenb., Sp. Hepat. (Lindenberg) 2-4: 93, 1840 (
** Plagiochila norfolkiensis Steph., Bull. Herb. Boissier (sér. 2) 3 (10): 877 (362), 1903 (
*** Plagiochila obtusa Lindenb., Sp. Hepat. (Lindenberg) 2-4: 42, 1840 (
* Plagiochila owaihiensis Nees et Lindenb., Sp. Hepat. (Lindenberg) 1: 30, 1839 (
** Plagiochila pacifica Mitt., Fl. vit.: 407, 1871 [1873] (
** Plagiochila parallela Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 686 (223), 1902 (
*** Plagiochila parvifolia Lindenb., Sp. Hepat. (Lindenberg) 1: 28, 1839 (
*** Plagiochila patentissima Lindenb., Sp. Hepat. (Lindenberg) 2-4: 64, 1840 (
*** Plagiochila patula (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 1: 21, 1839 (
** Plagiochila patula var. brevifolia Gottsche, Mexik. Leverm.: 10, 1863 (
** Plagiochila paucidens Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 117 (297), 1903 (
** Plagiochila pensilis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 497, 1885 (
** Plagiochila peradenyensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 172, 1900 [1901] (
** Plagiochila perdentata M.L.So et Grolle, Syst. Bot. 26 (3): 460, 2001 (
*** Plagiochila pinniflora Steph., Hedwigia 30 (5): 212, 1891 (
*** Plagiochila praemorsa Steph., Bot. Jahrb. Syst. 8 (2): 92, 1886 (
** Plagiochila propinqua Sande Lac., Plagiochila Sandei: 5, 1856 (
** Plagiochila purpurascens Steph., Sp. Hepat. (Stephani) 6: 197, 1921 (
*** Plagiochila raddiana Lindenb., Sp. Hepat. (Lindenberg) 1: 9, 1839 (
* Plagiochila ragazzii Gola, Ann. Bot. (Rome) 13 (1): 67, 1914 (
*** Plagiochila repanda (Schwägr.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 62, 1840 (
** Plagiochila repanda var. perrotana (Steph.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 65, 1981 (
*** Plagiochila rodriguezii Steph., Bot. Gaz. 15 (11): 290, 1890 (
*** Plagiochila rudolfii Pócs, Beih. Nova Hedwigia 90: 223, 1988 (
*** Plagiochila salacensis Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 576, 1853 (
*** Plagiochila salvadorica Steph., Hedwigia 30 (6): 272, 1891 (
** Plagiochila serrialata Herzog, Hedwigia 72 (6): 212, 1932 (
*** Plagiochila shangaica Steph., Sp. Hepat. (Stephani) 6: 216, 1921 (
*** Plagiochila simplex (Sw.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 54, 1840 (
*** Plagiochila squamulosa Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 165, 1863 (
** Plagiochila squamulosa var. crispulocaudata (Gottsche) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 74, 1981 (
** Plagiochila squamulosa var. sinuosa (Mitt.) Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 51 (1/2): 75, 1981 (
** Plagiochila streimannii Inoue, J. Jap. Bot. 63 (11): 365, 1988 (
*** Plagiochila strictifolia Steph., Hedwigia 30 (5): 210, 1891 (
** Plagiochila subflabellata Colenso, Trans. & Proc. New Zealand Inst. 21: 51, 1889 (
** Plagiochila subjavanica M.L.So, Austral. Syst. Bot. 13 (5): 804, 2000 (
*** Plagiochila subtropica Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 46, 1899 (
** Plagiochila tamariscina Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 685 (222), 1902 (
** Plagiochila tecta Inoue et Grolle, J. Hattori Bot. Lab. 33: 319, 1970 (
*** Plagiochila terebrans Nees et Mont., Sp. Hepat. (Lindenberg) 2-4: 98, 1840 (
*** Plagiochila teysmannii Sande Lac., Plagiochila Sandei: 6, 1856 (
** Plagiochila thyoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 498, 1885 (
** Plagiochila tocarema Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 106, 1864 (
** Plagiochila ulata Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 91, 1975 (
** Plagiochila undata Sull., Amer. J. Sci. Arts (ser. 2) 1 (1): 73, 1846 (
** Plagiochila undata subsp. crispata (Gottsche) R.M.Schust., Amer. Midl. Naturalist 63 (1): 122, 1960 (
** Plagiochila ungarangana Sande Lac., Syn. hepat. jav.: 10, 1856 [1857] (
*** Plagiochila virginica A.Evans, Prelim. cat. fl. W. Virginia: 497, 1892 (
** Plagiochila virginica var. caroliniana R.M.Schust., Amer. Midl. Naturalist 63 (1): 15, 1960 (
** Plagiochila virginica var. euryphylla R.M.Schust., Amer. Midl. Naturalist 63 (1): 21, 1960 (
*** Plagiochila wightii Nees, Sp. Hepat. (Lindenberg) 2-4: 43, 1840 (
** Plagiochila wilhelmina Inoue, J. Hattori Bot. Lab. 33: 317, 1970 (
* sect. Zanteniae (Inoue) Inoue, Gen. Plagiochila SE Asia: 45, 1984 (
** Plagiochila zantenii Inoue, J. Hattori Bot. Lab. 32: 107, 1969 (
Incertae sedis
* Plagiochila abscendens Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 104, 1864 (
** Plagiochila aculeata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 627, 1847 (
** Plagiochila acuta Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 607 (353), 1903 (
* Plagiochila albertii Steph., Biblioth. Bot. 87 (2): 188, 1916 (
* Plagiochila aliena Gottsche, Mexik. Leverm.: 22, 1863 (
** Plagiochila allionii Steph., Sp. Hepat. (Stephani) 6: 120, 1917 (
* Plagiochila ambigua Lindenb. et Hampe, Linnaea 24 (6): 640, 1851 [1852] (
* Plagiochila ampliata Steph., Biblioth. Bot. 87 (2): 189, 1916 (
** Plagiochila andicola Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 187, 1856 (
** Plagiochila andina Steph., Sp. Hepat. (Stephani) 6: 121, 1917 (
* Plagiochila angolensis Steph., Sp. Hepat. (Stephani) 6: 122, 1917 (
* Plagiochila angusteoblonga Steph., Biblioth. Bot. 87 (2): 189, 1916 (
** Plagiochila angustisedens Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 743 (542), 1905 (
** Plagiochila angustispina Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 671 (208), 1902 (
** Plagiochila apicalis Gottsche, Mexik. Leverm.: 29, 1863 (
** Plagiochila appalachiana Inoue, J. Hattori Bot. Lab. 40: 415, 1976 (
* Plagiochila arcta S.Winkl., Rev. Bryol. Lichénol. 42 (3): 818, 1976 (
** Plagiochila arcuata Lindenb., Sp. Hepat. (Lindenberg) 2-4: 91, 1840 (
** Plagiochila arenacea Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 53, 1964 (
* Plagiochila artsii Pócs, J. Hattori Bot. Lab. 100: 334, 2006 (
** Plagiochila atrovirens Taylor, London J. Bot. 5: 266, 1846 (
** Plagiochila balansae Gottsche, Contr. Étude Plagiochila: 149, 1928 (
** Plagiochila baldwinii Austin, Trans. Connecticut Acad. Arts 8 (15): 257, 1891 (
* Plagiochila bamingensis Steph., Bull. Mus. Natl. Hist. Nat. 18 (2): 117, 1912 (
** Plagiochila bancroftii Steph., Sp. Hepat. (Stephani) 6: 127, 1917 (
*** Plagiochila barbadensis Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 897 (563), 1905 (
* Plagiochila barbata Steph., Biblioth. Bot. 87 (2): 190, 1916 (
* Plagiochila barbeyi Steph., Biblioth. Bot. 87 (2): 190, 1916 (
* Plagiochila batava Steph., Sp. Hepat. (Stephani) 6: 128, 1917 (
* Plagiochila berggrenii Steph., Biblioth. Bot. 87 (2): 191, 1916 (
** Plagiochila beskeana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 863 (232), 1902 (
** Plagiochila bialata Mitt., Fl. vit.: 407, 1871 [1873] (
* Plagiochila biapiculata Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 891 (557), 1905 (
* Plagiochila bicaudata Steph., Sp. Hepat. (Stephani) 6: 130, 1918 (
** Plagiochila biciliata Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 529 (333), 1903 (
** Plagiochila bicornis Hampe et Gottsche, Linnaea 25 (3): 338, 1852 [1853] (
* Plagiochila bidentula Steph., Sp. Hepat. (Stephani) 6: 130, 1918 (
** Plagiochila binghamiae A.Evans, Trans. Connecticut Acad. Arts 18 (5): 304, 1914 (
** Plagiochila binominata Steph., Sp. Hepat. (Stephani) 6: 131, 1918 (
** Plagiochila bitexta Dugas, Contr. Étude Plagiochila: 58, 1928 (
** Plagiochila blepharobasis Herzog, Hedwigia 72 (6): 216, 1932 (
** Plagiochila bogotensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 98, 1864 (
* Plagiochila boliviana Spruce, Mem. Torrey Bot. Club 1 (3): 137, 1890 (
* Plagiochila borneensis Steph., Sp. Hepat. (Stephani) 6: 132, 1918 (
** Plagiochila brassii Inoue et Grolle, J. Hattori Bot. Lab. 36: 492, 1972 [1973] (
* Plagiochila brevicalycina Lindenb. et Gottsche, Linnaea 20 (3): 322, 1847 (
** Plagiochila brunneola Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 164 (416), 1904 (
*** Plagiochila bryhnii Steph., Biblioth. Bot. 87 (2): 192, 1916 (
** Plagiochila bunburii Taylor, London J. Bot. 5: 269, 1846 (
* Plagiochila byssacea Hampe, Linnaea 20 (3): 326, 1847 (
** Plagiochila caducidentata R.M.Schust., Phytologia 39 (4): 247, 1978 (
** Plagiochila caldana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 879 (248), 1902 (
** Plagiochila callaensis Steph., Sp. Hepat. (Stephani) 6: 136, 1918 (
* Plagiochila camusii Steph., Biblioth. Bot. 87 (2): 193, 1916 (
* Plagiochila capillicaulis Steph., Biblioth. Bot. 87 (2): 193, 1916 (
** Plagiochila caribbeania R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 412, 1980 (
* Plagiochila cava Steph., Bull. Herb. Boissier (sér. 2) 4 (12): 1213 (501), 1904 (
** Plagiochila ceylanica Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 98, 1860 [1861] (
** Plagiochila chacoënsis Herzog, Feddes Repert. Spec. Nov. Regni Veg. 55 (1): 7, 1952 (
** Plagiochila chiloënsis Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 27, 1900 (
** Plagiochila chinantlana Gottsche, Mexik. Leverm.: 12, 1863 (
* Plagiochila chiovendae Gola, Ann. Bot. (Rome) 13 (1): 67, 1914 (
* Plagiochila cinchonae Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 920 (570), 1905 (
* Plagiochila circumvoluta Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 475, 1947 (
*** Plagiochila cleefii Inoue, Stud. Cryptog. S. Peru: 95, 1987 (
* Plagiochila concava Nees, Sp. Hepat. (Lindenberg) 2-4: 70, 1840 (
** Plagiochila conduplicata Steph., Sp. Hepat. (Stephani) 6: 139, 1918 (
* Plagiochila conferta Steph., Sp. Hepat. (Stephani) 6: 139, 1918 (
** Plagiochila confertissima Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 182 (510), 1905 (
** Plagiochila connata Lindenb. et Gottsche, Syn. Hepat. 5: 645, 1847 (
** Plagiochila contorta Lindenb. et Hampe, Linnaea 24 (3): 301, 1851 [1852] (
** Plagiochila convoluta Steph., Sp. Hepat. (Stephani) 6: 141, 1918 (
** Plagiochila convolutifolia Steph., Sp. Hepat. (Stephani) 6: 142, 1918 (
*** Plagiochila corymbulosa Pearson, Forh. Vidensk.-Selsk. Kristiania 1887 (9): 14, 1887 (
** Plagiochila costariensis Horik., Acta Phytotax. Geobot. 13: 213, 1943 (
** Plagiochila crispabilis Lindenb., Sp. Hepat. (Lindenberg) 1: 15, 1839 (
* Plagiochila crispabilis var. minima Lindenb., Sp. Hepat. (Lindenberg) 5: 155, 1843 (
** Plagiochila cristophylla Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 117 (297), 1903 (
*** Plagiochila cuatrecasii H.Rob., Bryologist 70 (1): 47, 1967 (
** Plagiochila cubensis Steph., Sp. Hepat. (Stephani) 6: 143, 1918 (
** Plagiochila cucullata Lindenb. et Gottsche, Syn. Hepat. 5: 642, 1847 (
* Plagiochila cuervina Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 96, 1864 (
** Plagiochila delapsa Inoue, Beih. Nova Hedwigia 90: 171, 1988 (
* Plagiochila delognei Steph., Biblioth. Bot. 87 (2): 196, 1916 (
** Plagiochila denigrata Inoue, Willdenowia 18 (2): 557, 1989 (
** Plagiochila densa Herzog, Hedwigia 72 (6): 222, 1932 (
** Plagiochila densiflora Herzog, Hedwigia 72 (6): 226, 1932 (
** Plagiochila densiramosa Steph., Biblioth. Bot. 87 (2): 196, 1916 (
** Plagiochila deppeana Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 886 (552), 1905 (
** Plagiochila desciscens Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 867 (236), 1902 (
* Plagiochila dilatata Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 887 (553), 1905 (
* Plagiochila distans Colenso, Trans. & Proc. New Zealand Inst. 19: 283, 1887 (
** Plagiochila divaricata Lindenb., Sp. Hepat. (Lindenberg) 5: 147, 1843 (
* Plagiochila doerfleri Steph., Biblioth. Bot. 87 (2): 196, 1916 (
** Plagiochila dolichoblasta Herzog, Trans. Brit. Bryol. Soc. 1 (4): 286, 1950 (
** Plagiochila dussiana Steph., Symb. Antill. (Urban) 3 (2): 277, 1902 (
** Plagiochila echinata R.M.Schust., Amer. Midl. Naturalist 62 (2): 341, 1959 (
** Plagiochila echinella Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 332, 1857 (
* Plagiochila ecuadorensis Steph., Sp. Hepat. (Stephani) 6: 149, 1918 (
* Plagiochila effuseintricata Steph., Biblioth. Bot. 87 (2): 197, 1916 (
** Plagiochila eggersii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 14 (4): 138, 1988 (
** Plagiochila ekmanii S.W.Arnell, Bryologist 59 (4): 274, 1956 (
* Plagiochila electa Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 739 (538), 1905 (
* Plagiochila elegantula Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 161, 1955 (
* Plagiochila emarginatobidentula Steph., Biblioth. Bot. 87 (2): 197, 1916 (
* Plagiochila erythraeae Herzog, Hedwigia 78 (3/4): 224, 1938 (
** Plagiochila estrellensis Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 23, 1925 (
** Plagiochila eurydictyon Herzog, Hedwigia 72 (6): 207, 1932 (
* Plagiochila excisa Steph., Sp. Hepat. (Stephani) 6: 153, 1918 (
** Plagiochila exesa Lindenb. et Gottsche, Syn. Hepat. 5: 629, 1847 (
* Plagiochila exilis Colenso, Trans. & Proc. New Zealand Inst. 19: 282, 1887 (
** Plagiochila expansa Gottsche, Mexik. Leverm.: 18, 1863 (
** Plagiochila facallonia Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 871 (240), 1902 (
* Plagiochila falcato-oblonga Steph., Biblioth. Bot. 87 (2): 198, 1916 (
** Plagiochila fallax Lindenb. et Hampe, Linnaea 24 (3): 300, 1851 [1852] (
** Plagiochila faxinensis Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 60, 1964 (
** Plagiochila fendleri Mont., Ann. Sci. Nat. Bot. (sér. 4) 6: 198, 1856 (
* Plagiochila filicicola Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 351 (427), 1904 (
** Plagiochila fissidentoides Taylor, London J. Bot. 5: 264, 1846 (
* Plagiochila flabellifrons Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 488, 1885 (
** Plagiochila flava Steph., Sp. Hepat. (Stephani) 6: 156, 1918 (
** Plagiochila flavescens (Gottsche, Lindenb. et Nees) Gottsche, Mexik. Leverm.: 52, 1863 (
* Plagiochila flavorufescens Steph., Biblioth. Bot. 87 (2): 199, 1916 (
** Plagiochila footei A.Evans, Trans. Connecticut Acad. Arts 18 (5): 306, 1914 (
* Plagiochila formosa Nees, Contr. Étude Plagiochila: 147, 1928 (
** Plagiochila fragmentata R.M.Schust., Phytologia 45 (5): 421, 1980 (
* Plagiochila frausa Gottsche, Mexik. Leverm.: 66, 1863 (
* Plagiochila frausa var. boliviana Spruce, Mem. Torrey Bot. Club 1 (3): 134, 1890 (
** Plagiochila frayjorgensis Hässel, Bol. Soc. Argent. Bot. 22 (1/4): 103, 1983 (
* Plagiochila gaffatensis Gottsche ex Schweinf., Beitr. Fl. Aethiop.: 227, 1866 [1867] (
** Plagiochila gaudichaudii Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 193, 1856 (
** Plagiochila geniculata Lindenb., Sp. Hepat. (Lindenberg) 5: 131, 1843 (
* Plagiochila geppii Steph., Biblioth. Bot. 87 (2): 199, 1916 (
** Plagiochila germana Gottsche, Mexik. Leverm.: 34, 1863 (
** Plagiochila germanii Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 938 (588), 1905 (
* Plagiochila gibbiflora Steph., Bull. Herb. Boissier (sér. 2) 4 (6): 590 (441), 1904 (
** Plagiochila gittinsii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 139, 1977 (
* Plagiochila glauca Carl, Ann. Bryol., Suppl. 2: 129, 1931 (
* Plagiochila gracilicaulis Spruce, Mem. Torrey Bot. Club 1 (3): 132, 1890 (
** Plagiochila gracillima Austin, Trans. Connecticut Acad. Arts 8 (15): 256, 1891 (
* Plagiochila granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 111, 1864 (
** Plagiochila granditexta Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 165 (417), 1904 (
** Plagiochila grateloupii Mont., Ann. Sci. Nat. Bot. (sér. 4) 6: 188, 1856 (
** Plagiochila guatemalensis Steph., Sp. Hepat. (Stephani) 6: 163, 1918 (
** Plagiochila guttisquama Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 29, 1979 (
** Plagiochila gymnocalyx Inoue, Trop. Bryol. 1: 34, 1989 (
** Plagiochila haeseliae Inoue, Stud. Cryptog. S. Chile: 97, 1984 (
* Plagiochila hans-meyeri Steph., Sp. Hepat. (Stephani) 6: 164, 1918 (
** Plagiochila harlingii S.W.Arnell, Svensk Bot. Tidskr. 56 (2): 346, 1962 (
** Plagiochila haumanii Herzog, Repert. Spec. Nov. Regni Veg. 41 (14/25): 285, 1937 (
** Plagiochila hawaica Steph., Bull. Herb. Boissier (sér. 2) 3 (7): 598 (344), 1903 (
** Plagiochila heteracantha Steph., Sp. Hepat. (Stephani) 6: 166, 1918 (
* Plagiochila heterofolia Steph., Biblioth. Bot. 87 (2): 200, 1916 (
* Plagiochila hieronymii Steph., Biblioth. Bot. 87 (2): 201, 1916 (
** Plagiochila hiroshiana Pócs, J. Hattori Bot. Lab. 100: 335, 2006 (
** Plagiochila hoehnei Herzog, Hedwigia 72 (6): 220, 1932 (
* Plagiochila holstii Steph., Hedwigia 45 (4): 214, 1906 (
** Plagiochila horrida Gottsche, Mexik. Leverm.: 74, 1863 (
* Plagiochila huatuscana Gottsche, Mexik. Leverm.: 24, 1863 (
** Plagiochila huerlimannii Inoue, J. Hattori Bot. Lab. 33: 307, 1970 (
* Plagiochila humboldtiana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 112, 1864 (
** Plagiochila hyalina Lindenb., Syn. Hepat. 5: 640, 1847 (
* Plagiochila incisa Dugas, Contr. Étude Plagiochila: 112, 1928 (
** Plagiochila inflata Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 961 (376), 1903 (
* Plagiochila informifolia Steph., Biblioth. Bot. 87 (2): 202, 1916 (
** Plagiochila infuscata Mitt., J. Linn. Soc., Bot. 15 (82): 63, 1876 (
* Plagiochila injasutiensis S.W.Arnell, Bot. Not. 110: 404, 1957 (
** Plagiochila inouei Grolle, J. Bryol. 10 (3): 269, 1979 (
** Plagiochila insularia Mitt., St. Helena: 366, 1875 (
** Plagiochila intertexta Hook.f. et Taylor, London J. Bot. 5: 267, 1846 (
* Plagiochila inuensis Steph., Biblioth. Bot. 87 (2): 202, 1916 (
** Plagiochila irregularis Gottsche, Mexik. Leverm.: 14, 1863 (
** Plagiochila itatiajensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 874 (243), 1902 (
** Plagiochila jamaicensis Lindenb. et Hampe, Linnaea 24 (3): 302, 1851 [1852] (
** Plagiochila jaramillii H.Rob., Bryologist 70 (1): 48, 1967 (
* Plagiochila karstenii Steph., Hedwigia 45 (4): 214, 1906 (
* Plagiochila kaulfussiana Steph., Sp. Hepat. (Stephani) 6: 172, 1918 (
** Plagiochila keckiana Steph., Bull. Herb. Boissier (sér. 2) 5 (4): 358 (526), 1905 (
** Plagiochila kerneriana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 60, 1964 (
** Plagiochila koponenii Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 216, 1989 (
* Plagiochila lacerifolia Steph., Biblioth. Bot. 87 (2): 202, 1916 (
* Plagiochila lachenaudii Steph., Biblioth. Bot. 87 (2): 202, 1916 (
* Plagiochila laciniosa Dugas, Contr. Étude Plagiochila: 68, 1928 (
* Plagiochila laevifolia Gola, Ann. Bot. (Rome) 6 (2): 273, 1907 (
** Plagiochila lansbergii Gottsche, Mexik. Leverm.: 13, 1863 (
** Plagiochila latitrigona Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 48, 1964 (
* Plagiochila laxiramea Steph., Biblioth. Bot. 87 (2): 203, 1916 (
* Plagiochila lecontei Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 165 (417), 1904 (
* Plagiochila ledermanniana Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924 (
* Plagiochila ledieui Steph., Bull. Herb. Boissier (sér. 2) 4 (2): 161 (413), 1904 (
** Plagiochila lehmanniana S.W.Arnell, Svensk Bot. Tidskr. 55 (1): 205, 1961 (
** Plagiochila liebmanniana Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 97, 1840 (
* Plagiochila lignicola Spruce, Mem. Torrey Bot. Club 1 (3): 135, 1890 (
* Plagiochila lindaui Steph., Biblioth. Bot. 87 (2): 204, 1916 (
* Plagiochila lindigiana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 105, 1864 (
** Plagiochila lingua Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 677 (214), 1902 (
* Plagiochila longifissa Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 891 (557), 1905 (
* Plagiochila lotsyana Steph., Biblioth. Bot. 87 (2): 205, 1916 (
* Plagiochila luteola Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 175 (503), 1905 (
** Plagiochila lutescens Steph., Sp. Hepat. (Stephani) 6: 179, 1921 (
** Plagiochila luzonensis Grolle et M.L.So, Bryologist 102 (1): 69, 1999 (
* Plagiochila macra Taylor, London J. Bot. 7: 198, 1848 (
** Plagiochila macrifolia Taylor, London J. Bot. 5: 270, 1846 (
* Plagiochila macrifolia var. angustifolia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 470, 1885 (
*** Plagiochila martiana (Nees) Lindenb., Sp. Hepat. (Lindenberg) 1: 12, 1839 (
* Plagiochila matanga Steph., Sp. Hepat. (Stephani) 6: 183, 1921 (
** Plagiochila maunakeana Steph., Sp. Hepat. (Stephani) 6: 186, 1921 (
** Plagiochila maximiliana Gottsche, Mexik. Leverm.: 18, 1863 (
** Plagiochila meghalayensis K.K.Rawat et S.C.Srivast., Geophytology 35 (1/2): 49, 2005 (
** Plagiochila meridana Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 328, 1857 (
** Plagiochila microdonta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 97, 1860 [1861] (
* Plagiochila mildbreadiana Beauverd, Sp. Hepat. (Stephani) 6: 571, 1924 (
** Plagiochila minarum Herzog, Hedwigia 72 (6): 207, 1932 (
** Plagiochila minutiretis Reimers, Repert. Spec. Nov. Regni Veg. 21 (8/20): 264, 1925 (
** Plagiochila miqueliana Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 95, 1840 (
** Plagiochila molliuscula Inoue, Ann. Bot. Fenn. 13 (3): 134, 1976 (
** Plagiochila mollusca Lehm., Nov. Stirp. Pug. 10: 4, 1857 (
** Plagiochila moniliformis R.M.Schust., Phytologia 39 (4): 247, 1978 (
** Plagiochila muelleriana Gottsche, Mexik. Leverm.: 38, 1863 (
* Plagiochila multiflora Steph., Pflanzenw. Ost-Afrikas C: 64, 1895 (
** Plagiochila mutila Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 327, 1857 (
* Plagiochila naranjoënsis Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 687 (224), 1902 (
** Plagiochila neesiana Lindenb., Sp. Hepat. (Lindenberg) 2-4: 71, 1840 (
* Plagiochila neglecta Steph., Bull. Herb. Boissier (sér. 2) 5 (4): 351 (519), 1905 (
* Plagiochila negrensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 466, 1885 (
* Plagiochila negrii Gola, Ann. Bot. (Rome) 13 (1): 66, 1914 (
* Plagiochila nigricaulis Steph., Biblioth. Bot. 87 (2): 206, 1916 (
* Plagiochila notha Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 737 (536), 1905 (
* Plagiochila nova Steph., Sp. Hepat. (Stephani) 6: 189, 1921 (
** Plagiochila nudifolia (Steph.) Grolle, Feddes Repert. 82 (1): 89, 1971 (
** Plagiochila nutans Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 960 (375), 1903 (
* Plagiochila obliquetruncata Steph., Biblioth. Bot. 87 (2): 207, 1916 (
** Plagiochila oblonga Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (3): 398, 1965 (
** Plagiochila oerstediana Lindenb. et Hampe, Linnaea 24 (3): 301, 1851 [1852] (
** Plagiochila olivacea Steph., Bull. Herb. Boissier (sér. 2) 5 (2): 190 (518), 1905 (
* Plagiochila ornata Wilson ex Lindenb., Sp. Hepat. (Lindenberg) 5: 164, 1843 (
* Plagiochila ovato-obconica Steph., Sp. Hepat. (Stephani) 6: 191, 1921 (
* Plagiochila ovifolia Steph., Sp. Hepat. (Stephani) 6: 191, 1921 (
** Plagiochila palangiensis S.C.Srivast., K.K.Rawat et P.K.Verma, Natl. Acad. Sci. Lett. 29 (7/8): 267, 2006 (
** Plagiochila panamensis Inoue, Bull. Natl. Sci. Mus. Tokyo, B 15 (3): 94, 1989 (
** Plagiochila paramicola Herzog, Beih. Bot. Centralbl. 61B (3): 563, 1942 (
* Plagiochila parvitexta Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 674 (211), 1902 (
** Plagiochila parvivittata Inoue, Bull. Natl. Sci. Mus. Tokyo, B 13 (2): 50, 1987 (
* Plagiochila parvula Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 726, 1913 (
* Plagiochila pastasensis Steph., Sp. Hepat. (Stephani) 6: 197, 1921 (
** Plagiochila patentispina Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 877 (246), 1902 (
** Plagiochila patuloides Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 58, 1964 (
** Plagiochila paucidentata Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 197, 1856 (
** Plagiochila paupercula Gottsche, Mexik. Leverm.: 22, 1863 (
* Plagiochila pearceana Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 742 (541), 1905 (
** Plagiochila pellucida Lindenb. et Gottsche, Linnaea 20 (3): 321, 1847 (
** Plagiochila perrottetiana Mont. et Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 6: 195, 1856 (
** Plagiochila perrottetiana var. minor Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 101, 1864 (
** Plagiochila pilifera Steph., Sp. Hepat. (Stephani) 6: 198, 1921 (
** Plagiochila pinnatidens Steph., Sp. Hepat. (Stephani) 6: 199, 1921 (
** Plagiochila pittieri Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 673 (210), 1902 (
** Plagiochila platyphylla Herzog, Hedwigia 72 (6): 223, 1932 (
** Plagiochila polopolensis Herzog, Hedwigia 67 (6): 261, 1927 (
* Plagiochila prostrata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 668 (205), 1902 (
* Plagiochila pseudopatula Herzog, Hedwigia 72 (6): 227, 1932 (
** Plagiochila pseudoradicans Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 24, 1925 (
** Plagiochila ptychanthoidea Steph., Bull. Herb. Boissier (sér. 2) 3 (2): 121 (301), 1903 (
** Plagiochila pulchella Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 871 (240), 1902 (
** Plagiochila pulvinata Steph., Bull. Herb. Boissier (sér. 2) 3 (6): 526 (330), 1903 (
** Plagiochila ratkowskiana Inoue, Brunonia 3 (1): 141, 1980 (
** Plagiochila regeliana Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 675 (212), 1902 (
** Plagiochila remyana Steph., Bull. Herb. Boissier (sér. 2) 3 (11): 963 (378), 1903 (
*** Plagiochila revolvens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (
** Plagiochila rigidula Lindenb. et Gottsche, Linnaea 20 (3): 323, 1847 (
** Plagiochila rosana Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 892 (558), 1905 (
* Plagiochila rubricaulis Steph., Bot. Jahrb. Syst. 20 (3): 311, 1895 (
* Plagiochila rufifolia Steph., Biblioth. Bot. 87 (2): 209, 1916 (
** Plagiochila rufoviridis Spruce, Mem. Torrey Bot. Club 1 (3): 136, 1890 (
* Plagiochila rusbyi Spruce, Mem. Torrey Bot. Club 1 (3): 135, 1890 (
* Plagiochila sabensis Steph., Sp. Hepat. (Stephani) 6: 214, 1921 (
** Plagiochila sachapatensis Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 679 (216), 1902 (
** Plagiochila salazariae Inoue, Bull. Natl. Sci. Mus. Tokyo, B 15 (3): 91, 1989 (
** Plagiochila saltuensis Spruce ex Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 927 (577), 1905 (
** Plagiochila saltuensis var. spinosissima Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 163, 1955 (
* Plagiochila samoana Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 85: 197, 1910 (
** Plagiochila sarmentosa (Lehm. et Lindenb.) Lindenb., Sp. Hepat. (Lindenberg) 2-4: 86, 1840 (
* Plagiochila schiffneri Steph., Biblioth. Bot. 87 (2): 209, 1916 (
* Plagiochila schinzei Steph., Biblioth. Bot. 87 (2): 209, 1916 (
* Plagiochila schmidtii Steph., Biblioth. Bot. 87 (2): 209, 1916 (
* Plagiochila schraderbergii Steph., Sp. Hepat. (Stephani) 6: 244, 1922 (
** Plagiochila schubertiana Steph., Sp. Hepat. (Stephani) 6: 208, 1921 (
** Plagiochila schusteri Inoue et Grolle, J. Hattori Bot. Lab. 33: 326, 1970 (
** Plagiochila scissifolia Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 865 (234), 1902 (
* Plagiochila scotica Macvicar ex Steph., Sp. Hepat. (Stephani) 6: 209, 1921 (
** Plagiochila semiamplexicaulis Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 936 (586), 1905 (
** Plagiochila semiermis Dugas, Contr. Étude Plagiochila: 66, 1928 (
** Plagiochila silvatica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 108, 1864 (
* Plagiochila similis Steph., Biblioth. Bot. 87 (2): 210, 1916 (
** Plagiochila sisparensis Steph., Sp. Hepat. (Stephani) 6: 207, 1921 (
* Plagiochila slateri Steph., Biblioth. Bot. 87 (2): 211, 1916 (
** Plagiochila solitaria Gottsche, Sp. Hepat. (Stephani) 6: 225, 1921 (
*** Plagiochila spinifera Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 113, 1873 (
* Plagiochila sprucei Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 860 (229), 1902 (
* Plagiochila staudtiana Steph., Bull. Herb. Boissier (sér. 2) 4 (4): 352 (428), 1904 (
** Plagiochila stipata Steph., Sp. Hepat. (Stephani) 6: 209, 1921 (
* Plagiochila stolzii Steph., Sp. Hepat. (Stephani) 6: 244, 1922 (
** Plagiochila subcontigua Herzog, Hedwigia 72 (6): 230, 1932 (
* Plagiochila subconvoluta Gottsche, Mexik. Leverm.: 24, 1863 (
* Plagiochila subdenudata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 673 (210), 1902 (
** Plagiochila subedentata Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 679 (216), 1902 (
** Plagiochila subfragilis Inoue, Stud. Cryptog. S. Peru: 98, 1987 (
* Plagiochila subhyalina Steph., Biblioth. Bot. 87 (2): 212, 1916 (
** Plagiochila subligulata Steph., Sp. Hepat. (Stephani) 6: 224, 1921 (
** Plagiochila sublyallii M.L.So, New Zealand J. Bot. 39 (1): 109, 2001 (
* Plagiochila subrara Herzog, Hedwigia 74 (2): 87, 1934 (
* Plagiochila subrotundifolia Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 935 (585), 1905 (
** Plagiochila subundulata Lindenb., Sp. Hepat. (Lindenberg) 5: 137, 1843 (
* Plagiochila suringarii Steph., Sp. Hepat. (Stephani) 6: 208, 1921 (
** Plagiochila sylvicultrix Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 486, 1885 (
* Plagiochila symmetrica Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 876 (245), 1902 (
** Plagiochila tabinana Gottsche, Mexik. Leverm.: 46, 1863 (
** Plagiochila talinayi S.W.Arnell, Svensk Bot. Tidskr. 50 (2): 312, 1956 (
** Plagiochila tambillensis Loitl., Diagn. pl. nov.: 22, 1894 (
** Plagiochila tarapotensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 863 (232), 1902 (
* Plagiochila tenera Steph., Sp. Hepat. (Stephani) 6: 231, 1921 (
** Plagiochila tenuis Lindenb., Sp. Hepat. (Lindenberg) 2-4: 50, 1840 (
* Plagiochila tenuispica Steph., Sp. Hepat. (Stephani) 6: 234, 1921 (
** Plagiochila thamniopsis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890] (
* Plagiochila thollonii Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 980 (270), 1902 (
** Plagiochila thrausta Inoue et Grolle, Bull. Natl. Sci. Mus. Tokyo, B 5 (1): 34, 1979 (
* Plagiochila tonduzana Steph., Sp. Hepat. (Stephani) 6: 227, 1921 (
** Plagiochila tortuosa Lindenb. et Gottsche, Mexik. Leverm.: 70, 1863 (
* Plagiochila tovarina Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 102, 1864 (
* Plagiochila trabeculatospinosa Herzog, Hedwigia 72 (6): 213, 1932 (
* Plagiochila trabutii Steph., Biblioth. Bot. 87 (2): 213, 1916 (
** Plagiochila trianae Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 114, 1864 (
** Plagiochila trichomanes Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cc, 1889 [1890] (
* Plagiochila tricuspis Steph., Sp. Hepat. (Stephani) 6: 229, 1921 (
** Plagiochila tristaniana Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 89, 2013 (
** Plagiochila tristis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 880 (249), 1902 (
** Plagiochila trollii Herzog, Hedwigia 74 (2): 90, 1934 (
** Plagiochila truncata Gottsche, Mexik. Leverm.: 25, 1863 (
** Plagiochila uleana Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 868 (237), 1902 (
** Plagiochila umbrosioides L.Söderstr., Phytotaxa 208 (1): 85, 2015 (
* Plagiochila unduavensis Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 882 (251), 1902 (
* Plagiochila usambarana Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 980 (270), 1902 (
** Plagiochila variedentata Steph., Bull. Herb. Boissier (sér. 2) 2 (10): 887 (256), 1902 (
** Plagiochila vastifolia Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 680 (217), 1902 (
** Plagiochila velata Inoue et Piippo, Ann. Bot. Fenn. 26 (2): 207, 1989 (
* Plagiochila venezuelana Steph., Bull. Herb. Boissier (sér. 2) 5 (8): 749 (548), 1905 (
* Plagiochila ventricosotrigona Steph., Biblioth. Bot. 87 (2): 214, 1916 (
* Plagiochila verrucosa Steph., Bull. Herb. Boissier (sér. 2) 5 (9): 885 (551), 1905 (
* Plagiochila vetustisilva Steph., Sp. Hepat. (Stephani) 6: 245, 1922 (
* Plagiochila viminea Spruce, Mem. Torrey Bot. Club 1 (3): 134, 1890 (
* Plagiochila viridis Steph., Sp. Hepat. (Stephani) 6: 239, 1921 (
** Plagiochila viridonigra (E.A.Hodgs.) Inoue, Bryologist 68 (2): 218, 1965 (
** Plagiochila vittiana Inoue, Beih. Nova Hedwigia 90: 171, 1988 (
** Plagiochila vittifolia Steph., Sp. Hepat. (Stephani) 6: 239, 1921 (
** Plagiochila vulcanica Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 671 (208), 1902 (
** Plagiochila wacei S.W.Arnell ex Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 89, 2013 (
* Plagiochila wallisiana Steph., Bull. Herb. Boissier (sér. 2) 5 (10): 936 (586), 1905 (
** Plagiochila wattsiana J.J.Engel et G.L.Merr., Nova Hedwigia 91 (3/4): 511, 2010 (
** Plagiochila wettsteiniana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 53, 1964 (
* Plagiochila weymouthiana Steph., Biblioth. Bot. 87 (2): 215, 1916 (
** Plagiochila wiemanniana S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 54, 1964 (
*** Plagiochila winteri Steph., Bull. Herb. Boissier (sér. 2) 2 (12): 981 (271), 1902 (
** Plagiochila wrightii Steph., Bull. Herb. Boissier (sér. 2) 2 (8): 681 (218), 1902 (
** Plagiochila xalapensis Gottsche, Mexik. Leverm.: 21, 1863 (
* Plagiochila xanthochroma Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 489, 1885 (
* Plagiochila yoshinagana Steph., Sp. Hepat. (Stephani) 6: 242, 1922 (
** Plagiochila zacuapana Gottsche, Mexik. Leverm.: 20, 1863 (
** Plagiochilidium Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 186, 1931 (
** Plagiochilidium bidentulum (Steph.) Grolle, J. Hattori Bot. Lab. 65: 408, 1988 (
** Plagiochilion S.Hatt., Biosphaera 1 (1): 7, 1947 (
*** Plagiochilion braunianum (Nees) S.Hatt., Biosphaera 1 (1): 7, 1947 (
** Plagiochilion combinatum (Mitt.) Inoue, J. Hattori Bot. Lab. 27: 55, 1964 (
*** Plagiochilion conjugatum (Hook.) R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963 (
** Plagiochilion fimbriatum (Mitt.) Inoue, J. Hattori Bot. Lab. 27: 57, 1964 (
** Plagiochilion giulianettii (Steph.) Inoue, J. Hattori Bot. Lab. 27: 57, 1964 (
** Plagiochilion herzogii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (2): 270, 1971 (
** Plagiochilion intermedium R.M.Schust., Phytologia 45 (5): 421, 1980 (
** Plagiochilion mayebarae S.Hatt., J. Hattori Bot. Lab. 3: 39, 1948 [1950] (
*** Plagiochilion oppositum (Reinw., Blume et Nees) S.Hatt., Biosphaera 1 (1): 7, 1947 (
** Plagiochilion pachycephalum (De Not.) Inoue, J. Hattori Bot. Lab. 27: 55, 1964 (
*** Plagiochilion proliferum (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 285, 1963 (
** Plagiochilion theriotanum (Steph.) Inoue, J. Hattori Bot. Lab. 27: 59, 1964 (
** Pseudolophocolea R.M.Schust. et J.J.Engel, Lindbergia 8 (2): 71, 1982 (
** Pseudolophocolea denticulata R.M.Schust. et J.J.Engel, Lindbergia 8 (2): 73, 1982 (
** Xenochila R.M.Schust., Amer. Midl. Naturalist 62 (1): 15, 1959 (
** Xenochila integrifolia (Mitt.) Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 6 (4): 373, 1963 (
by M. von Konrat
Blepharostoma was recognized as an element within Pseudolepicoleaceae by
*** Archeophylla R.M.Schust., J. Hattori Bot. Lab. 26: 263, 1963 (
** Archeophylla paradoxa R.M.Schust., Trans. Brit. Bryol. Soc. 4 (5): 810, 1965 (
** Archeophylla pungens (Herzog) R.M.Schust., Candollea 21 (1): 86, 1966 (
*** Archeophylla schusteri (E.A.Hodgs. et Allison) R.M.Schust., J. Hattori Bot. Lab. 26: 263, 1963 (
*** Castanoclobos J.J.Engel et Glenny, Novon 17 (4): 424, 2007 (
*** Castanoclobos julaceus (Hatcher ex J.J.Engel) J.J.Engel et Glenny, Novon 17 (4): 425, 2007 (
** Chaetocolea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 346, 1885 (
*** Chaetocolea palmata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 346, 1885 (
*** Herzogiaria Fulford ex Hässel, Lindbergia 7 (1): 23, 1981 (
*** Herzogiaria teres (Steph.) Fulford ex Hässel, Lindbergia 7 (1): 24, 1981 (
** Isophyllaria E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand, Bot. 3 (4): 68, 1965 (
*** Isophyllaria attenuata (Rodway) E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 111, 1972 (
** Isophyllaria fuegiana (Hässel) R.M.Schust., Beih. Nova Hedwigia 118: 141, 2000 (
*** Pseudolepicolea Fulford et J.Taylor, Nova Hedwigia 1 (3/4): 412, 1959 [1960] (
** Pseudolepicolea andoi (R.M.Schust.) Inoue, Bull. Natl. Sci. Mus. Tokyo, B 4 (3): 94, 1978 (
** Pseudolepicolea fryei (Perss.) Grolle et Ando, Hikobia 3 (3): 180, 1963 (
*** Pseudolepicolea grolleana (R.M.Schust.) Grolle, Ann. Bot. Fenn. 21 (1): 30, 1984 (
*** Pseudolepicolea kuehnemannii (R.M.Schust.) Hässel, Fl. Criptog. Tierra del Fuego 15: 125, 1975 (
*** Pseudolepicolea quadrilaciniata (Sull.) Fulford et J.Taylor, Nova Hedwigia 1 (3/4): 413, 1959 [1960] (
*** Pseudolepicolea temnomoides (R.M.Schust.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 92, 2013 (
** Pseudolepicolea trollii (Herzog) Grolle et Ando, Hikobia 3 (3): 177, 1963 (
* Pseudolepicolea trollii var. darjeelingensis S.Hatt. et Mizut., J. Hattori Bot. Lab. 31: 252, 1968 (
*** Temnoma Mitt., Handb. N. Zeal. fl. 2: 750, 1867 (
*** Temnoma angustifolium R.M.Schust., Candollea 21 (2): 279, 1966 [1967] (
*** Temnoma chaetophyllum R.M.Schust., Phytologia 39 (4): 239, 1978 (
** Temnoma palmatum (Lindb. ex Pearson) R.M.Schust., Bryologist 62 (4): 240, 1959 [1960] (
** Temnoma palmatum var. cuneatum R.M.Schust., Candollea 21 (2): 347, 1966 [1967] (
** Temnoma palmatum var. laxifolium R.M.Schust., Candollea 21 (2): 345, 1966 [1967] (
** Temnoma palmatum var. pseudospiniferum R.M.Schust., Candollea 21 (2): 348, 1966 [1967] (
** Temnoma patagonicum R.M.Schust., Candollea 21 (2): 313, 1966 [1967] (
*** Temnoma paucisetigerum R.M.Schust., Candollea 21 (2): 266, 1966 [1967] (
*** Temnoma pilosum (A.Evans) R.M.Schust., Bryologist 62 (4): 240, 1959 [1960] (
*** Temnoma pulchellum (Hook.) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
*** Temnoma quadrifidum (Mitt.) Mitt. ex E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand, Bot. 1 (12): 142, 1962 (
*** Temnoma quadripartitum (Hook.) Mitt., J. Linn. Soc., Bot. 15 (82): 68, 1876 (
** Temnoma quadripartitum var. pseudopungens R.M.Schust., Candollea 21 (2): 312, 1966 [1967] (
** Temnoma quadripartitum var. randii (S.W.Arnell) R.M.Schust., Candollea 21 (2): 307, 1966 [1967] (
*** Temnoma setigerum (Lindenb.) R.M.Schust., Nova Hedwigia 5: 35, 1963 (
** Temnoma setigerum var. hawaiicum Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 17 (3): 228, 1974 (
** Temnoma townrowii R.M.Schust., Candollea 21 (2): 351, 1966 [1967] (
by T. Katagiri
The treatment of Trichocoleaceae follows what was outlined in
*** Eotrichocolea R.M.Schust., J. Hattori Bot. Lab. 26: 252, 1963 (
*** Eotrichocolea furukii T.Katag., Bryologist 115 (4): 519, 2012 (
*** Eotrichocolea polyacantha (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 264, 1963 (
*** Leiomitra Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875 (
** subg. Brachygyna R.M.Schust., Nova Hedwigia 73 (3/4): 480, 2001 (
** Leiomitra mastigophoroides R.M.Schust., Phytologia 45 (5): 416, 1980 (
** subg. Leiomitra, Nova Hedwigia 73 (3/4): 480, 2001 (
*** Leiomitra breviseta (Steph.) R.M.Schust., Beih. Nova Hedwigia 118: 152, 2000 (
*** Leiomitra capillata Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875 (
*** Leiomitra elegans (Lehm.) Hässel, Novon 12 (4): 465, 2002 (
** Leiomitra elliottii (Steph.) R.M.Schust., Nova Hedwigia 73 (3/4): 469, 2001 (
*** Leiomitra flaccida Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 349, 1885 (
** Leiomitra hirticaulis R.M.Schust., Nova Hedwigia 73 (3/4): 472, 2001 (
*** Leiomitra lanata (Hook.) R.M.Schust., Phytologia 45 (5): 417, 1980 (
*** Leiomitra merrillana (Steph.) T.Katag., Bryologist 115 (4): 488, 2012 (
*** Leiomitra paraphyllina Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 350, 1885 (
** Leiomitra robusta (Steph.) R.M.Schust., Nova Hedwigia 73 (3/4): 469, 2001 (
** Leiomitra smaragdina Hässel, Novon 12 (4): 467, 2002 (
*** Leiomitra tomentosa (Sw.) Lindb., Acta Soc. Sci. Fenn. 10: 515, 1875 (
*** Trichocolea Dumort., Commentat. Bot. (Dumortier): 113, 1822 (
** Trichocolea argentea Herzog, Arch. Bot. São Paulo 1 (2): 40, 1925 (
*** Trichocolea brevifissa Steph., Sp. Hepat. (Stephani) 4: 54, 1909 (
** Trichocolea comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 27, 1922 (
*** Trichocolea filicaulis Steph., Sp. Hepat. (Stephani) 4: 59, 1909 (
* Trichocolea floccosa Herzog et Hatcher, Lloydia 20 (3): 148, 1957 [1958] (
** Trichocolea geniculata Pearson, J. Linn. Soc., Bot. 46 (305): 28, 1922 (
** Trichocolea gracillima Austin, Bot. Gaz. 3 (1): 6, 1878 (
*** Trichocolea hatcheri E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 69, 1965 (
*** Trichocolea iriomotensis T.Katag., Hattoria 4: 6, 2013 (
*** Trichocolea japonica T.Katag., Bryologist 114 (4): 744, 2011 (
*** Trichocolea magna T.Katag., Hattoria 4: 7, 2013 (
** Trichocolea minutifolia Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 134, 1914 (
*** Trichocolea mollissima (Hook.f. et Taylor) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 132, 1864 (
*** Trichocolea pluma (Reinw., Blume et Nees) Mont., Voy. Bonite, Bot. 1: 238, 1846 (
*** Trichocolea rigida R.M.Schust., Nova Hedwigia 15: 447, 1968 (
*** Trichocolea rudimentaris Steph., Sp. Hepat. (Stephani) 6: 376, 1923 (
* Trichocolea sprucei Steph., Sp. Hepat. (Stephani) 4: 59, 1909 (
*** Trichocolea tomentella (Ehrh.) Dumort., Syll. Jungerm. Europ.: 67, 1831 (
** Trichocolea udarii D.K.Singh, Bull. Bot. Surv. India 25: 177, 1983 [1985] (
** Trichocolea wattsiana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 135, 1914 (
by L. Söderström
*** Mylia Gray, Nat. Arr. Brit. Pl. 1: 693, 1821 (
** subg. Anomalae (R.M.Schust. ex Potemkin) L.Söderstr., Phytotaxa 202 (1): 70, 2015 (
*** Mylia anomala (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 693, 1821 (
** subg. Mylia
*** Mylia taylorii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 693, 1821 (
*** Mylia verrucosa Lindb., Acta Soc. Sci. Fenn. 10: 236, 1872 [1873] (
*** Mylia verrucosa subsp. nuda (Inoue et B.Y.Yang) Potemkin et Kazan., Arctoa 2: 5, 1993 (
by X. He and D. Glenny
*** Schistochila Dumort., Recueil Observ. Jungerm.: 15, 1835 (
** Schistochila acuminata Steph., Sp. Hepat. (Stephani) 4: 81, 1909 (
*** Schistochila aequiloba Steph., Sp. Hepat. (Stephani) 4: 80, 1909 (
*** Schistochila alata (Lehm.) Schiffn., Hepat. (Engl.-Prantl): 111, 1893 (
*** Schistochila aligera (Nees et Blume) J.B.Jack et Steph., Hedwigia 31 (1): 12, 1892 (
* Schistochila aligera var. laxa (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 212, 1898 (
*** Schistochila altissima E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 85, 1965 (
** Schistochila altissima subsp. polystratosa R.M.Schust. et J.J.Engel, Phytologia 30 (4): 241, 1975 (
*** Schistochila antara Grolle, J. Hattori Bot. Lab. 29: 249, 1966 (
*** Schistochila appendiculata (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (
* Schistochila baileyana Steph., Sp. Hepat. (Stephani) 4: 85, 1909 (
*** Schistochila balfouriana (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 91, 1909 (
*** Schistochila beccariana (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (
*** Schistochila berggrenii (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (
*** Schistochila berteroana (Hook.) Steph., Sp. Hepat. (Stephani) 4: 96, 1909 (
*** Schistochila blumei (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (
** Schistochila brassii Grolle, J. Hattori Bot. Lab. 31: 5, 1968 (
** Schistochila caledonica Steph., Sp. Hepat. (Stephani) 4: 77, 1909 (
*** Schistochila carnosa (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 93, 1909 (
*** Schistochila caudata R.M.Schust. et J.J.Engel, Phytologia 30 (4): 242, 1975 (
*** Schistochila childii (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (
*** Schistochila chlorophylla (Colenso) E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 71 (3): 192, 1941 (
*** Schistochila ciliata (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 87, 1909 (
*** Schistochila compacta (Colenso) E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 71 (3): 192, 1941 (
*** Schistochila conchophylla Herzog ex E.A.Hodgs. et Allison, Trans. & Proc. Roy. Soc. New Zealand 71 (3): 191, 1941 (
** Schistochila conchophylla var. multidentata (J.J.Engel) Xiao L.He et Glenny, Phytotaxa 173 (1): 92, 2014 (
*** Schistochila congoana Steph., Sp. Hepat. (Stephani) 4: 71, 1909 (
** Schistochila cookei (H.A.Mill.) R.M.Schust., J. Hattori Bot. Lab. 42: 274, 1977 (
** Schistochila crinita Grolle, J. Hattori Bot. Lab. 31: 7, 1968 (
*** Schistochila cristata Steph., Hedwigia 28 (4): 274, 1889 (
*** Schistochila cunninghamii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 27, 1901 (
** Schistochila doriae (De Not.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (
** Schistochila engleriana Steph., Sp. Hepat. (Stephani) 4: 69, 1909 (
*** Schistochila exalata Herzog, Rev. Bryol. Lichénol. 29 (3/4): 191, 1960 [1961] (
** Schistochila fijiensis H.Buch et Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 94, 1952 (
*** Schistochila glaucescens (Hook.) A.Evans, Rev. Bryol. 32 (4): 60, 1905 (
** Schistochila hattorii Grolle, J. Hattori Bot. Lab. 29: 247, 1966 (
** Schistochila integerrima Steph., Sp. Hepat. (Stephani) 6: 492, 1924 (
** Schistochila isotachyphylla (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (
*** Schistochila kirkiana Steph., Sp. Hepat. (Stephani) 4: 86, 1909 (
*** Schistochila kunkelii S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 12, 1957 (
** Schistochila lacerata Steph., Sp. Hepat. (Stephani) 6: 492, 1924 (
*** Schistochila lamellata (Hook.) Dumort. ex A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892 (
*** Schistochila laminigera (Hook.f. et Taylor) A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892 (
*** Schistochila latiloba (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (
*** Schistochila lehmanniana (Lindenb.) Steph., Sp. Hepat. (Stephani) 4: 86, 1909 (
*** Schistochila leucophylla (Lehm. ex Gottsche, Lindenb. et Nees) Steph., Sp. Hepat. (Stephani) 4: 98, 1910 (
** Schistochila macrodonta W.E.Nicholson, Symb. Sin. 5: 29, 1930 (
** Schistochila minor C.Gao et Y.H.Wu, J. Hattori Bot. Lab. 95: 267, 2004 (
*** Schistochila monticola R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 623, 1971 (
*** Schistochila muricata E.A.Hodgs. et Allison, Trans. & Proc. Roy. Soc. New Zealand 71 (3): 186, 1941 (
*** Schistochila nadeaudiana Steph., Sp. Hepat. (Stephani) 4: 76, 1909 (
** Schistochila neesii (Mont.) Lindb., J. Linn. Soc., Bot. 13 (67): 194, 1872 [1873] (
*** Schistochila nitidissima R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 27, 1968 (
*** Schistochila nivicola (R.M.Schust. et J.J.Engel) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (
*** Schistochila nobilis (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (
*** Schistochila nuda Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 215, 1934 (
*** Schistochila pachyphylla (Lehm.) Steph., Sp. Hepat. (Stephani) 4: 99, 1910 (
*** Schistochila papillifera R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 27, 1968 (
*** Schistochila parvistipula Rodway, Pap. & Proc. Roy. Soc. Tasmania 1916: 47, 1917 (
*** Schistochila pellucida R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 286, 1985 (
*** Schistochila piligera Steph., Bot. Gaz. 15 (11): 291, 1890 (
*** Schistochila pinnatifolia (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 392, 1877 (
*** Schistochila pluriciliata R.M.Schust. et J.J.Engel, J. Hattori Bot. Lab. 58: 395, 1985 (
*** Schistochila pseudociliata R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 623, 1971 (
*** Schistochila quadrifida A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892 (
** Schistochila ramentacea Steph., Sp. Hepat. (Stephani) 6: 494, 1924 (
*** Schistochila reflexa (Mont.) Steph., Sp. Hepat. (Stephani) 4: 97, 1910 (
*** Schistochila reflexistipula J.J.Engel et R.M.Schust., Phytologia 30 (4): 245, 1975 (
*** Schistochila reinwardtii (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 218, 1898 (
*** Schistochila repleta (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 90, 1909 (
** Schistochila rubriseta Steph., Sp. Hepat. (Stephani) 4: 78, 1909 (
** Schistochila schultzei Steph., Sp. Hepat. (Stephani) 6: 494, 1924 (
*** Schistochila sciophila R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 14 (4): 632, 1971 (
*** Schistochila sciurea (Nees) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 251, 1893 (
*** Schistochila simulans (C.Massal.) Xiao L.He et Yu Sun, Polish Bot. J. 58 (2): 473, 2013 (
*** Schistochila spegazziniana (C.Massal.) Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 60, 1900 (
*** Schistochila sphagnoides (Schwägr.) Lindb. ex Steph., Sp. Hepat. (Stephani) 4: 70, 1909 (
*** Schistochila splachnophylla (Hook.f. et Taylor) Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 28, 1901 (
** Schistochila stratosa (Mont.) A.Evans, Contr. U.S. Natl. Herb. 1 (5): 141, 1892 (
*** Schistochila subhyalina R.M.Schust., Phytologia 30 (4): 246, 1975 (
** Schistochila subhyalina var. grandidentata (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (
*** Schistochila subimmersa J.J.Engel et R.M.Schust., Phytologia 30 (4): 247, 1975 (
*** Schistochila succulenta (J.J.Engel et R.M.Schust.) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (
*** Schistochila tasmanica Steph., Sp. Hepat. (Stephani) 4: 86, 1909 (
*** Schistochila trispiralis R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 28, 1968 (
*** Schistochila tuloides (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 4: 89, 1909 (
*** Schistochila undulatifolia Piippo, Ann. Bot. Fenn. 23 (1): 8, 1986 (
*** Schistochila virescens R.M.Schust., Phytologia 30 (4): 248, 1975 (
*** Schistochila vitreocincta (Herzog) Xiao L.He et Glenny, Austral. Syst. Bot. 23 (4): 237, 2010 (
** Schistochila volans Grolle, J. Hattori Bot. Lab. 29: 244, 1966 (
*** Schistochila yakushimensis N.Ohnishi et Deguchi, Bryologist 106 (3): 451, 2003 (
** Schistochila zantenii Grolle, J. Hattori Bot. Lab. 29: 243, 1966 (
by M. von Konrat, J. Hentschel, J. Uribe, P. Sukkharak, J. Heinrichs, J. Larraín, R. Stotler and L. Zhang
The subgeneric treatment of Frullaniaceae follows
*** Frullania Raddi, Jungermanniogr. Etrusca: 9, 1818 (
*** subg. Chonanthelia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 8, 1884 (
* Frullania flammea Taylor, Trans. & Proc. Bot. Soc. Edinburgh 15: 29, 1884 (
*** Frullania lindmanii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 19, 1897 (
** Frullania paranensis Steph., Sp. Hepat. (Stephani) 4: 607, 1911 (
** Frullania spegazzinii M.E.Reiner, Bol. Soc. Argent. Bot. 25 (3/4): 310, 1988 (
*** sect. Chonanthelia (Spruce) Yuzawa ex Hentschel et von Konrat, Phytotaxa 220 (2): 129, 2015 (
*** Frullania gibbosa Nees, Syn. Hepat. 3: 411, 1845 (
*** sect. Cladocarpicae Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 43, 1884 (
*** Frullania beauverdii Steph., Biblioth. Bot. 87 (2): 241, 1916 (
*** Frullania blepharozia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 18, 1884 (
*** Frullania bogotensis Steph., Sp. Hepat. (Stephani) 4: 327, 1910 (
* Frullania brachycarpa Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cciv, 1889 [1890] (
*** Frullania confertiloba Steph., Sp. Hepat. (Stephani) 4: 326, 1910 (
*** Frullania decidua Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 30, 1884 (
*** Frullania ecklonii (Spreng.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 413, 1845 (
* Frullania ecklonii var. robustior (Gottsche, Lindenb. et Nees) Sim, Trans. Roy. Soc. South Africa 15 (1): 39, 1926 (
* Frullania ecklonii var. rufescens Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 771, 1847 (
* Frullania ecklonii var. tenerior (Gottsche, Lindenb. et Nees) Sim, Trans. Roy. Soc. South Africa 15 (1): 39, 1926 (
*** Frullania holostipula S.Hatt. et D.G.Griffin, Misc. Bryol. Lichenol. 8 (3): 47, 1978 (Hattori and Griffin 1978).
*** Frullania megalostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 15, 1884 (
*** Frullania obscura (Sw.) Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 333, 1840 (
*** Frullania obscura var. spiniloba (Steph.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (
*** Frullania ringens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 17, 1884 (
*** Frullania rio-janeirensis (Raddi) Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 88, 1876 [1877] (
*** Frullania sphaerocephala Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 17, 1884 (
** Frullania tunguraguana L.Clark et Frye, Bryologist 55 (2): 133, 1952 (
*** sect. Pluricarinatae (Yuzawa, Mues et S.Hatt.) Hentschel et von Konrat, Phytotaxa 220 (2): 129, 2015 (
*** Frullania albertii Steph., Biblioth. Bot. 87 (2): 237, 1916 (
*** Frullania arsenii Steph., Sp. Hepat. (Stephani) 6: 530, 1924 (
** Frullania bonariensis M.E.Reiner, Bol. Soc. Argent. Bot. 25 (3/4): 313, 1988 (
*** Frullania cuencensis Taylor, London J. Bot. 5: 406, 1846 (
*** Frullania depressa Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 168, 1863 (
*** Frullania dusenii Steph., Arch. Mus. Nac. Rio de Janeiro 13: 115 (9), 1905 (
*** Frullania gradsteinii Yuzawa, Mues et S.Hatt., J. Hattori Bot. Lab. 63: 429, 1987 (
*** Frullania haematocysta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 54, 1884 (
*** Frullania jelskii Loitl., Diagn. pl. nov.: 18, 1894 (
*** Frullania laxiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 26, 1884 (
* Frullania laxiflora var. crossii Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 27, 1884 (
*** Frullania neurota Taylor, London J. Bot. 5: 400, 1846 (
*** Frullania planifolia Steph., Sp. Hepat. (Stephani) 4: 337, 1910 (
*** Frullania pluricarinata Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 168, 1864 (
** Frullania sandvicensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 28, 1872 (
*** Frullania standaertii Steph., Sp. Hepat. (Stephani) 4: 342, 1910 (
*** Frullania stenostipa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 29, 1884 (
*** Frullania tetraptera Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838 (
** Frullania trinervis (Lehm.) Drège, Flora, Beig. 26: 186, 1843 (
*** Frullania winteri Steph., Sp. Hepat. (Stephani) 4: 338, 1910 (
*** Frullania winteri var. vanderhammenii (Haarbrink) Yuzawa, J. Hattori Bot. Lab. 70: 233, 1991 (
*** subg. Diastaloba Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 55, 1884 (
** Frullania antaresensis S.Hatt., J. Hattori Bot. Lab. 47: 92, 1980 (
* Frullania armatifolia Verd., Bull. Jard. Bot. Buitenzorg (sér. 3) 12 (1): 61, 1932 (
*** Frullania curvilobula Schäf.-Verw., D.F.Peralta et S.M.Siqueira, Phytotaxa 57 (4): 27, 2012 (
** Frullania gracilicaulis S.Hatt., J. Hattori Bot. Lab. 43: 421, 1977 [1978] (
** Frullania humbertii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 24, 1976 (
** Frullania hypoleucula S.Hatt., J. Hattori Bot. Lab. 57: 413, 1984 (
** Frullania incurva S.Hatt., J. Hattori Bot. Lab. 65: 431, 1988 (
** Frullania klotzschii Nees, Sp. Hepat. (Stephani) 4: 558, 1911 (
** Frullania letestui Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 38, 1976 (
* Frullania miradorensis Lindenb. et Gottsche, Syn. Hepat. 5: 781, 1847 (
* Frullania odontostipa Spruce, Mem. Torrey Bot. Club 1 (3): 120, 1890 (
*** Frullania pilibracteola S.Hatt., J. Hattori Bot. Lab. 43: 428, 1977 [1978] (
*** Frullania pilistipula Steph., Sp. Hepat. (Stephani) 4: 648, 1911 (
*** Frullania ramuligera (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 14, 1842 (
** Frullania subpilibracteola S.Hatt., J. Hattori Bot. Lab. 43: 434, 1977 [1978] (
** Frullania subtilissima (Nees ex Mont.) Lindenb., Syn. Hepat. 3: 443, 1845 (
** Frullania vandenberghenii Pócs, Acta Bot. Acad. Sci. Hung. 25 (3/4): 229, 1979 [1980] (
grp. Diastaloba I
** Frullania akiyamae S.Hatt., J. Hattori Bot. Lab. 60: 240, 1986 (
** Frullania apiculata (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 452, 1845 (
** Frullania apiculata var. goebelii Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 222, 1893 (
*** Frullania armitiana Steph., Sp. Hepat. (Stephani) 4: 538, 1911 (
** Frullania armitiana var. inflexula S.Hatt., J. Hattori Bot. Lab. 65: 415, 1988 (
** Frullania aterrima (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 450, 1845 (
*** Frullania attenuata Steph., Sp. Hepat. (Stephani) 4: 538, 1911 (
*** Frullania bella Steph., Sp. Hepat. (Stephani) 4: 643, 1911 (
** Frullania changii S.Hatt. et C.Gao, J. Jap. Bot. 60 (1): 1, 1985 (
* Frullania claviloba Steph., Sp. Hepat. (Stephani) 4: 651, 1911 (
*** Frullania colliculosa von Konrat, Braggins, Hentschel et Heinrichs, Nova Hedwigia 91 (3/4): 494, 2010 (
*** Frullania cordistipula (Reinw., Blume et Nees) Nees, Voy. Amér. Mérid. 7 (2): 68, 1839 (
** Frullania cordistipula var. dentistipula S.Hatt., J. Hattori Bot. Lab. 60: 242, 1986 (
* Frullania cordistipula var. mutica (Gottsche, Lindenb. et Nees) Schiffn., Consp. Hepat. Arch. Ind.: 323, 1898 (
* Frullania crenatiloba Steph., Sp. Hepat. (Stephani) 6: 551, 1924 (
*** Frullania curvistipula Steph., Sp. Hepat. (Stephani) 4: 548, 1911 (
** Frullania curvistipula var. falcatidentata S.Hatt., Misc. Bryol. Lichenol. 9 (6): 124, 1982 (
** Frullania curvistipula var. lamii Verd., Ann. Bryol., Suppl. 1: 91, 1930 (
** Frullania cuspidifolia Steph., Sp. Hepat. (Stephani) 4: 543, 1911 (
* Frullania degelii S.W.Arnell, Svensk Bot. Tidskr. 53 (4): 503, 1959 (
** Frullania dentifera S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 102, 1985 (
*** Frullania dentiloba S.Hatt., J. Jap. Bot. 50 (6): 161, 1975 (
** Frullania durifolia Steph., Hedwigia 33 (3): 162, 1894 (
** Frullania exilis Taylor, London J. Bot. 5: 405, 1846 (
** Frullania gabonensis Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 81, 1976 (
*** Frullania gracilis (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 452, 1845 (
* Frullania gracilis var. brevior (Gottsche, Lindenb. et Nees) Schiffn., Consp. Hepat. Arch. Ind.: 327, 1898 (
** Frullania gracilis var. vittata S.Hatt., J. Hattori Bot. Lab. 60: 243, 1986 (
** Frullania gracilis subsp. zennoskei S.Hatt. et Thaithong, J. Hattori Bot. Lab. 44: 183, 1978 (
*** Frullania hasskarliana Lindenb., Syn. Hepat. 3: 453, 1845 (
** Frullania hasskarliana var. gracilis S.Hatt., J. Hattori Bot. Lab. 60: 243, 1986 (
** Frullania hasskarliana var. parvidentata S.Hatt., J. Hattori Bot. Lab. 60: 245, 1986 (
*** Frullania hattorii von Konrat et Braggins, New Zealand J. Bot. 41 (1): 56, 2003 (
*** Frullania hodgsoniae von Konrat, Braggins, Hentschel et Heinrichs, Nova Hedwigia 91 (3/4): 492, 2010 (
** Frullania hottana S.Hatt., J. Hattori Bot. Lab. 40: 479, 1976 (
*** Frullania inconstans Verd., Ann. Bryol., Suppl. 1: 83, 1930 (
** Frullania inconstans var. grossedentata Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 530, 1973 (
*** Frullania johnsonii Steph., Hedwigia 33 (3): 163, 1894 (
** Frullania macgregorii Steph., Hedwigia 33 (3): 154, 1894 (
** Frullania macgregorii var. rostellula (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 220, 1982 (
** Frullania madens Steph., Sp. Hepat. (Stephani) 6: 553, 1924 (
** Frullania mehrana S.Hatt., Recent Adv. Bot.: 66, 1976 (
** Frullania motoyana Steph., Sp. Hepat. (Stephani) 4: 646, 1911 (
* Frullania multilacera Steph., Sp. Hepat. (Stephani) 4: 650, 1911 (
** Frullania multilacera subsp. gracilior S.Hatt., Mem. New York Bot. Gard. 45: 547, 1987 (
** Frullania multilacera var. lacerissima S.Hatt., J. Hattori Bot. Lab. 39: 294, 1975 (
** Frullania multilaceroides S.Hatt., Mem. New York Bot. Gard. 45: 549, 1987 (
** Frullania neosheana S.Hatt., J. Hattori Bot. Lab. 45: 350, 1979 (
** Frullania papillata Steph., Sp. Hepat. (Stephani) 4: 615, 1911 (
** Frullania pulogensis Steph., Sp. Hepat. (Stephani) 4: 545, 1911 (
*** Frullania purpurea Steph., Sp. Hepat. (Stephani) 4: 626, 1911 (
** Frullania reimersii Verd., Ann. Bryol., Suppl. 1: 84, 1930 (
** Frullania saepisdentata S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 116, 1985 (
** Frullania schiffneri Verd., Ann. Bryol. 2: 150, 1929 (
** Frullania schusterana S.Hatt., J. Hattori Bot. Lab. 36: 411, 1972 [1973] (
** Frullania seriatifolia Steph., Hedwigia 33 (3): 167, 1894 (
** Frullania serrata Gottsche, Syn. Hepat. 3: 453, 1845 (
** Frullania serrata var. ceramensis S.Hatt., J. Hattori Bot. Lab. 60: 247, 1986 (
** Frullania serrata subsp. grolleana (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982 (
** Frullania serrata var. hamatispina (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982 (
** Frullania serrata var. pertenuis (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 342, 1898 (
** Frullania serrata subsp. spinistipula S.Hatt., J. Hattori Bot. Lab. 51: 225, 1982 (
** Frullania setacea S.Hatt., J. Hattori Bot. Lab. 65: 447, 1988 (
** Frullania sheana S.Hatt., J. Hattori Bot. Lab. 45: 356, 1979 (
** Frullania simmondsii Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 109, 1914 (
*** Frullania sinuata Sande Lac., Ned. Kruidk. Arch. 3: 424, 1854 [1855] (
** Frullania steereana S.Hatt., Mem. New York Bot. Gard. 45: 553, 1987 (
** Frullania stipatiloba Steph., Hedwigia 33 (3): 168, 1894 (
** Frullania subdentata Steph., Sp. Hepat. (Stephani) 4: 545, 1911 (
** Frullania subdentata var. latistipula (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 51: 228, 1982 (
** Frullania submultilacera S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 190, 1994 (
** Frullania subocellata S.Hatt., J. Hattori Bot. Lab. 60: 248, 1986 (
*** Frullania taxodiocola R.M.Schust., Phytologia 53 (5): 364, 1983 (
*** Frullania ternatensis Gottsche, Syn. Hepat. 4: 465, 1846 (
** Frullania ternatensis var. non-appendiculata S.Hatt., J. Hattori Bot. Lab. 38: 174, 1974 (
*** Frullania trichodes Mitt., Bonplandia 10 (2): 19, 1862 (
** Frullania vaga Mitt., Fl. vit.: 418, 1871 [1873] (
*** Frullania vaginata (Sw.) Nees, Syn. Hepat. 4: 465, 1846 (
* Frullania vaginata var. nigricans (Gottsche, Lindenb. et Nees) Schiffn., Consp. Hepat. Arch. Ind.: 348, 1898 (
* Frullania van-zantenii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 528, 1973 (
** Frullania venusta S.Hatt., J. Hattori Bot. Lab. 38: 217, 1974 (
** Frullania verdoorniana S.Hatt., J. Hattori Bot. Lab. 37: 122, 1973 (
*** Frullania vitalii Yuzawa et S.Hatt., J. Jap. Bot. 63 (1): 30, 1988 (
** Frullania vittata S.Hatt., J. Hattori Bot. Lab. 38: 270, 1974 (
** Frullania vivipara Pócs, Fieldiana, Bot. (n.ser.) 47: 151, 2008 (
** Frullania wairua von Konrat et Braggins, New Zealand J. Bot. 43 (4): 886, 2005 (
** Frullania warnckeana S.Hatt., J. Hattori Bot. Lab. 38: 213, 1974 (
** Frullania warnckeana var. dentosa S.Hatt., Misc. Bryol. Lichenol. 7 (8): 162, 1977 (
grp. Diastaloba II
*** Frullania baumannii S.Hatt., J. Hattori Bot. Lab. 43: 410, 1977 [1978] (
*** Frullania congesta Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 451, 1845 (
** Frullania ocellata S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 531, 1973 (
*** Frullania ptychantha Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 257, 1843 (
*** Frullania repandistipula Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855] (
** Frullania repandistipula subsp. queenslandica S.Hatt., Mem. New York Bot. Gard. 45: 550, 1987 (
** Frullania repandistipula subsp. spinibractea S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 150, 1975 (
grp. Diastaloba III
** Frullania grossiclava Steph., Sp. Hepat. (Stephani) 4: 384, 1910 (
** Frullania loricata Pearson, Forh. Vidensk.-Selsk. Kristiania 1890 (2): 6, 1891 (
* Frullania loricata var. laxa Pearson, Forh. Vidensk.-Selsk. Kristiania 1890 (2): 8, 1891 (
*** Frullania usambarana Schiffn., Hedwigia 33 (3): 160, 1894 (
** Frullania usambarana var. reducta Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 91, 1976 (
grp. Diastaloba IV
* Frullania brunea (Spreng.) Drège, Flora, Beig. 26: 186, 1843 (
*** Frullania caulisequa (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839 (
*** Frullania grossifolia Steph., Sp. Hepat. (Stephani) 4: 633, 1911 (
*** Frullania hypoleuca Nees, Observ. bot.: 470, 1843 (
*** Frullania lindenbergii Lehm., Nov. Stirp. Pug. 8: 17, 1844 (
* Frullania lindenbergii var. fusca Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 780, 1847 (
** Frullania ponapensis S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 186, 1994 (
* Frullania tricarinata Sande Lac., Plagiochila Sandei: 10, 1856 (
* subg. Diversitextae (Kamim.) S.Hatt., J. Hattori Bot. Lab. 59: 154, 1985 (
*** Frullania diversitexta Steph., Bull. Herb. Boissier 5 (2): 89, 1897 (
*** subg. Frullania
* Frullania amamiensis Kamim., Bull. Kochi Gakuen Jun. Coll. 1: 51, 1970 (
** Frullania amplicrania Steph., Sp. Hepat. (Stephani) 4: 404, 1910 (
*** Frullania ampullifera J.B.Jack et Steph., Hedwigia 33 (3): 139, 1894 (
*** Frullania anderssonii Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 144, 1873 (
** Frullania angustistipa Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 297, 1907 (
** Frullania aposinensis S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 131, 1985 (
** Frullania appendistipula S.Hatt., J. Hattori Bot. Lab. 36: 424, 1972 [1973] (
** Frullania appendistipula var. spinifera S.Hatt., J. Hattori Bot. Lab. 38: 226, 1974 (
** Frullania auriculata S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 11 (1): 11, 1985 (
** Frullania benjaminiana Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 109, 1975 (
** Frullania bergmanii S.Hatt., J. Hattori Bot. Lab. 38: 192, 1974 (
*** Frullania berthoumieui Steph., Hedwigia 33 (3): 140, 1894 (
** Frullania bhutanensis S.Hatt., Fl. E. Himalaya 2: 232, 1971 (
** Frullania blastopetala S.Hatt., J. Hattori Bot. Lab. 57: 407, 1984 (
*** Frullania bonincola S.Hatt., J. Hattori Bot. Lab. 44: 551, 1978 (
* Frullania brevicalycina Steph., Hedwigia 33 (3): 141, 1894 (
** Frullania brittoniae A.Evans, Trans. Connecticut Acad. Arts 10 (1): 15, 1899 (
*** Frullania bullata Steph., Sp. Hepat. (Stephani) 4: 371, 1910 (
*** Frullania caffraria Steph., Hedwigia 33 (3): 141, 1894 (
*** Frullania calcarata Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 137, 1873 (
** Frullania carrii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 520, 1973 (
** Frullania chenii S.Hatt. et P.J.Lin, J. Jap. Bot. 60 (4): 106, 1985 (
** Frullania chilensis Steph., Hedwigia 33 (3): 142, 1894 (
*** Frullania chodatii Beauverd, Sp. Hepat. (Stephani) 6: 533, 1924 (
*** Frullania cobrensis Gottsche, Hedwigia 33 (3): 142, 1894 (
** Frullania consociata Steph., Sp. Hepat. (Stephani) 4: 461, 1911 (
* Frullania contracta Steph., Sp. Hepat. (Stephani) 4: 469, 1911 (
* Frullania cornuta Steph., Sp. Hepat. (Stephani) 4: 467, 1911 (
*** Frullania crassitexta Steph., Sp. Hepat. (Stephani) 4: 423, 1910 (
*** Frullania crispiplicata Yuzawa et S.Hatt., J. Jap. Bot. 58 (2): 43, 1983 (
** Frullania cristata S.Hatt., J. Hattori Bot. Lab. 49: 165, 1981 (
* Frullania cuneiloba Nees, Syn. Hepat. 3: 427, 1845 (
** Frullania cyparioides (Schwägr.) Nees, Naturgesch. Eur. Leberm. 3: 210, 1838 (
*** Frullania davurica Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 422, 1845 (
** Frullania debilis Steph. ex S.Hatt., J. Hattori Bot. Lab. 38: 190, 1974 (
** Frullania deppii Lehm., Nov. Stirp. Pug. 8: 15, 1844 (
** Frullania diptera (Lehm.) Drège, Flora, Beig. 26: 186, 1843 (
*** Frullania duthiana Steph., Sp. Hepat. (Stephani) 4: 351, 1910 (
** Frullania duthiana var. appendiculata S.Hatt., Fl. E. Himalaya 2: 234, 1971 (
** Frullania duthiana var. laevis S.Hatt., Bull. Univ. Mus. Univ. Tokyo 8: 233, 1975 (
** Frullania duthiana var. szechuanensis S.Hatt. et C.Gao, J. Jap. Bot. 60 (1): 2, 1985 (
** Frullania echinantha S.Hatt., J. Hattori Bot. Lab. 38: 233, 1974 (
** Frullania echinatella S.Hatt., J. Hattori Bot. Lab. 65: 423, 1988 (
* Frullania elegans Lehm., Nov. Stirp. Pug. 10: 16, 1857 (
*** Frullania elephantum S.Hatt., J. Hattori Bot. Lab. 43: 416, 1977 [1978] (
** Frullania epiphylla S.Hatt., J. Hattori Bot. Lab. 38: 235, 1974 (
* Frullania epiphylla subsp. fijiensis S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 11 (1): 14, 1985 (
** Frullania erostrata S.Hatt., J. Hattori Bot. Lab. 38: 237, 1974 (
* Frullania esenbeckiana Beauverd, Sp. Hepat. (Stephani) 6: 570, 1924 (
** Frullania evelynae S.Hatt. et Thaithong, J. Hattori Bot. Lab. 44: 179, 1978 (
* Frullania evelynae var. devendrae Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 109, 2012 (
* Frullania evelynae var. srivastavae Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 112, 2012 (
*** Frullania eymae S.Hatt., J. Hattori Bot. Lab. 39: 284, 1975 (
** Frullania eymae var. crispidentata S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 105, 1985 (
*** Frullania falciloba Lehm., Nov. Stirp. Pug. 8: 20, 1844 (
** Frullania falsicornuta S.Hatt., J. Hattori Bot. Lab. 60: 212, 1986 (
** Frullania fauriana Steph., Hedwigia 33 (3): 144, 1894 (
** Frullania fengyangshanensis R.L.Zhu et M.L.So, Bryologist 100 (3): 356, 1997 (
** Frullania ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 4: 417, 1910 (
* Frullania flexuosa S.Hatt., J. Hattori Bot. Lab. 54: 146, 1983 (
** Frullania fuscovirens Steph., Sp. Hepat. (Stephani) 4: 401, 1910 (
** Frullania fuscovirens var. gemmipara (R.M.Schust. et S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 135, 1985 (
** Frullania gaoligongensis X.L.Bai et C.Gao, Hikobia 13 (1): 87, 1999 (
** Frullania gemmulosa S.Hatt. et Thaithong, J. Hattori Bot. Lab. 43: 449, 1977 [1978] (
*** Frullania gigantea Steph., Sp. Hepat. (Stephani) 4: 467, 1911 (
** Frullania giraldiana C.Massal., Hepat. Shen-si: 41, 1897 (
** Frullania giraldiana var. handelii (Verd.) S.Hatt., J. Hattori Bot. Lab. 36: 123, 1972 [1973] (
** Frullania globosa S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 107, 1985 (
** Frullania grandilobula S.Hatt. et Piippo, Acta Bot. Fenn. 133: 45, 1986 (
*** Frullania grandistipula Lindenb., Syn. Hepat. 3: 430, 1845 (
** Frullania hainanensis S.Hatt. et P.J.Lin, J. Jap. Bot. 61 (10): 307, 1986 (
** Frullania hamatiloba Steph., Sp. Hepat. (Stephani) 4: 400, 1910 (
** Frullania handelii Verd., Symb. Sin. 5: 36, 1930 (
** Frullania handel-mazzettii S.Hatt., J. Hattori Bot. Lab. 49: 150, 1981 (
** Frullania hattoriantha Udar et V.Nath, Misc. Bryol. Lichenol. 9 (2): 44, 1981 (
* Frullania hebridensis Steph., Sp. Hepat. (Stephani) 4: 469, 1911 (
** Frullania hicksiae S.Hatt., Cryptog. Bryol. Lichénol. 5 (1/2): 182, 1984 (
** Frullania higuchii Yuzawa, Koike et S.Hatt., J. Hattori Bot. Lab. 75: 194, 1994 (
** Frullania hiroshii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 6 (1): 34, 1980 (
* Frullania hirtiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 35, 1884 (
*** Frullania howeana Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 107, 1914 (
** Frullania ignatovii Sofronova, Mamontov et Potemkin, Novosti Sist. Nizš. Rast. 47: 335, 2013 (
** Frullania incisoduthiana S.Hatt., J. Hattori Bot. Lab. 46: 391, 1979 (
* Frullania incisoduthiana var. parva S.Hatt., J. Hattori Bot. Lab. 46: 391, 1979 (
* Frullania incisostipula Steph., Sp. Hepat. (Stephani) 6: 541, 1924 (
** Frullania inflexiloba S.Hatt., J. Hattori Bot. Lab. 57: 415, 1984 (
** Frullania inouei S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 6 (1): 36, 1980 (
*** Frullania irregularis S.Hatt. et Piippo, Acta Bot. Fenn. 133: 46, 1986 (
*** Frullania jackii Gottsche, Hepat. Eur., Leberm. 29-30: no. 294, 1863 (
** Frullania jacobsii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (4): 129, 1986 (
* Frullania jacquinotii Gottsche, J. Bot. (Morot) 12: 138, 1898 (
** Frullania jovetiana von Konrat et Hentschel, Phytotaxa 220 (2): 135, 2015 (
** Frullania kagoshimensis Steph., Sp. Hepat. (Stephani) 4: 353, 1910 (
** Frullania kagoshimensis subsp. hunanensis (S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 137, 1985 (
** Frullania kalimantanensis S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (4): 127, 1986 (
** Frullania kashyapii Verd., Ann. Bryol. 5: 162, 1932 (
** Frullania kitagawana S.Hatt., J. Hattori Bot. Lab. 57: 417, 1984 (
** Frullania laeviperiantha X.L.Bai et C.Gao, Phytotaxa 220 (2): 136, 2015 (
* Frullania lancistyla Steph., Sp. Hepat. (Stephani) 4: 389, 1910 (
* Frullania latiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 34, 1884 (
** Frullania latogaleata Herzog, Trans. Brit. Bryol. Soc. 1 (3): 189, 1949 (
** Frullania lepida S.Hatt. et Piippo, Acta Bot. Fenn. 133: 48, 1986 (
* Frullania levieri Steph., Sp. Hepat. (Stephani) 4: 388, 1910 (
** Frullania linii S.Hatt., J. Hattori Bot. Lab. 49: 155, 1981 (
* Frullania longistyla Yuzawa et S.Hatt., J. Jap. Bot. 63 (11): 361, 1988 (
* Frullania ludoviciae Steph., Rev. Bryol. 35 (2): 29, 1908 (
** Frullania lushanensis S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 137, 1985 (
* Frullania macularis Taylor, London J. Bot. 5: 403, 1846 (
** Frullania maymyoensis Svihla, Bryologist 61 (4): 376, 1958 [1959] (
** Frullania microauriculata Verd., Ann. Bryol. 2: 126, 1929 (
** Frullania microauriculata var. rotundior Verd., Ann. Jard. Bot. Buitenzorg 40: 141, 1929 (
** Frullania microrhyncha L.Clark et Svihla, Bryologist 53 (1): 63, 1950 (
** Frullania mizoramensis Sushil K.Singh et Barbhuiya, Taiwania 57 (2): 106, 2012 (
* Frullania montana Steph., Sp. Hepat. (Stephani) 4: 455, 1911 (
** Frullania multituberculata Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (
** Frullania mutilata Steph., Sp. Hepat. (Stephani) 4: 673, 1911 (
** Frullania nadeaudii Steph., Sp. Hepat. (Stephani) 4: 465, 1911 (
*** Frullania nepalensis (Spreng.) Lehm. et Lindenb., Syn. Hepat. 3: 422, 1845 (
*** Frullania nicholsonii E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 368, 1949 (
** Frullania nigricaulis (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 457, 1845 (
* Frullania nigricaulis var. elongata Verd., Ann. Bryol., Suppl. 1: 155, 1930 (
** Frullania nivimontana S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 8 (3): 95, 1982 (
*** Frullania nobilis Steph., Hedwigia 33 (3): 154, 1894 (
** Frullania nobilis var. cochleata (Steph.) S.Hatt., J. Hattori Bot. Lab. 59: 111, 1985 (
** Frullania novocurvirostris S.Hatt., J. Hattori Bot. Lab. 49: 370, 1981 (
** Frullania oahuensis Hampe ex Gottsche, Lindenb. et Nees, Observ. bot.: 471, 1843 (
** Frullania obovata S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 8 (3): 97, 1982 (
** Frullania obtusangula Hentschel et von Konrat, Phytotaxa 220 (2): 137, 2015 (
** Frullania okinawensis Kamim., Misc. Bryol. Lichenol. 9 (4): 90, 1982 (
** Frullania orbicularis Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869 (
*** Frullania orientalis Sande Lac., Plagiochila Sandei: 10, 1856 (
** Frullania orinocensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 30, 1884 (
*** Frullania ornithocephala (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 425, 1845 (
* Frullania ornithocephala var. major (Nees) Schiffn., Consp. Hepat. Arch. Ind.: 336, 1898 (
** Frullania ornithocephala var. pilosa Verd., Ann. Bryol. 2: 129, 1929 (
** Frullania ornithocephala var. tuberculosa S.Hatt., J. Hattori Bot. Lab. 38: 179, 1974 (
** Frullania pachyderma S.Hatt., J. Hattori Bot. Lab. 44: 525, 1978 (
** Frullania pallidevirens Steph., Sp. Hepat. (Stephani) 4: 454, 1911 (
** Frullania pariharii S.Hatt. et Thaithong, J. Jap. Bot. 53 (5): 130, 1978 (
* Frullania parvifolia Steph., Sp. Hepat. (Stephani) 4: 354, 1910 (
** Frullania pauciramea Steph., Sp. Hepat. (Stephani) 4: 458, 1911 (
** Frullania pauciramea var. pauciramella S.Hatt. et Piippo, Acta Bot. Fenn. 133: 50, 1986 (
** Frullania paucirameoides S.Hatt. et Piippo, Acta Bot. Fenn. 133: 51, 1986 (
** Frullania pedicellata Steph., Bull. Herb. Boissier 5 (2): 90, 1897 (
*** Frullania physantha Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 121, 1860 [1861] (
** Frullania piptophylla S.Hatt., J. Hattori Bot. Lab. 47: 87, 1980 (
** Frullania piptophylla var. minor S.Hatt., J. Hattori Bot. Lab. 60: 247, 1986 (
** Frullania piptophylloides S.Hatt., J. Hattori Bot. Lab. 47: 90, 1980 (
* Frullania plicata Hentschel et von Konrat, Phytotaxa 220 (2): 137, 2015 (
** Frullania pocsantha Thaithong et S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 149, 1977 (
** Frullania pran-nathii M.Dey et D.K.Singh, J. Jap. Bot. 83 (5): 281, 2008 (
** Frullania pringlei Fulford et Sharp, Mem. New York Bot. Gard. 63: 45, 1990 (
** Frullania prominula S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 112, 1985 (
** Frullania propaginea S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 114, 1985 (
* Frullania pseudericoides S.Hatt., J. Hattori Bot. Lab. 51: 256, 1982 (
** Frullania pseudoschensiana S.Hatt., J. Hattori Bot. Lab. 47: 101, 1980 (
** Frullania pseudoschensiana var. darjeelingensis S.Hatt., J. Hattori Bot. Lab. 49: 149, 1981 (
** Frullania pullei Verd., Nova Guinea 14: 542, 1930 (
** Frullania pusilla Mitt., Fl. vit.: 417, 1871 [1873] (
*** Frullania pycnantha (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 411, 1845 (
** Frullania queenslandica Steph., Sp. Hepat. (Stephani) 4: 424, 1910 (
** Frullania recurvistipula S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 114, 1975 (
*** Frullania reflexistipula Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855] (
** Frullania reflexistipula var. squarrosa S.Hatt. et Piippo, Acta Bot. Fenn. 133: 54, 1986 (
** Frullania remotidens S.Hatt., J. Hattori Bot. Lab. 51: 257, 1982 (
* Frullania reptans Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 161, 1855 (
*** Frullania retusa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 119, 1860 [1861] (
** Frullania retusa var. gymnantha S.Hatt. et Thaithong, J. Jap. Bot. 53 (5): 131, 1978 (
** Frullania retusa var. hirsuta S.Hatt. et Thaithong, J. Hattori Bot. Lab. 44: 191, 1978 (
*** Frullania rhystocolea Herzog, Symb. Sin. 5: 39, 1930 (
** Frullania rhytidantha S.Hatt., J. Hattori Bot. Lab. 47: 97, 1980 (
** Frullania rigida Steph., Sp. Hepat. (Stephani) 4: 371, 1910 (
*** Frullania riparia Hampe, Nov. Stirp. Pug. 7: 14, 1838 (
** Frullania rizalii Piippo et S.Hatt., J. Hattori Bot. Lab. 72: 119, 1992 (
*** Frullania rostellata Mitt., Handb. N. Zeal. fl. 2: 755, 1867 (
** Frullania rubella Gottsche, Hedwigia 28 (3): 159, 1889 (
** Frullania rubella var. elongata (Steph.) S.Hatt., J. Hattori Bot. Lab. 54: 166, 1983 (
** Frullania rupicola Steph., Sp. Hepat. (Stephani) 6: 534, 1924 (
** Frullania saipanensis S.Hatt. et Koike, J. Hattori Bot. Lab. 75: 188, 1994 (
*** Frullania schensiana C.Massal., Hepat. Shen-si: 40, 1897 (
** Frullania schusteri S.Hatt., Beih. Nova Hedwigia 90: 154, 1988 (
** Frullania scottiana S.Hatt., Mem. New York Bot. Gard. 45: 551, 1987 (
** Frullania setchellii Pearson, Univ. Calif. Publ. Bot. 10 (4): 326, 1923 (
** Frullania shanensis Svihla, Bryologist 60 (4): 359, 1957 (
** Frullania sharpantha Udar et Ad.Kumar, Misc. Bryol. Lichenol. 9 (9): 192, 1983 (
** Frullania sharpii S.Hatt., J. Hattori Bot. Lab. 38: 180, 1974 (
** Frullania sinensis Steph., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 349, 1906 (
** Frullania sinosphaerantha S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 144, 1985 (
** Frullania sphaerantha S.Hatt., J. Hattori Bot. Lab. 47: 99, 1980 (
** Frullania sphaerolobulata S.H.Lin, Tunghai Journal 38: 104, 1997 (
*** Frullania spinifera Taylor, London J. Bot. 5: 407, 1846 (
** Frullania spinigastria S.Hatt., J. Hattori Bot. Lab. 45: 358, 1979 (
** Frullania spiniplica S.Hatt., J. Hattori Bot. Lab. 36: 428, 1972 [1973] (
*** Frullania spongiosa Steph., Hedwigia 33 (3): 147, 1894 (
** Frullania squamuligera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 33, 1884 (
*** Frullania squarrosula (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 412, 1845 (
** Frullania subcaduca S.Hatt., J. Hattori Bot. Lab. 38: 267, 1974 (
** Frullania subclavata Steph., Sp. Hepat. (Stephani) 4: 354, 1910 (
** Frullania subnigricaulis S.Hatt., J. Hattori Bot. Lab. 37: 89, 1973 (
** Frullania subnigricaulis var. subtruncata S.Hatt., J. Hattori Bot. Lab. 39: 308, 1975 (
* Frullania subpedicellata S.Hatt., J. Hattori Bot. Lab. 47: 93, 1980 (
** Frullania subsquarrosa S.Hatt., J. Hattori Bot. Lab. 36: 429, 1972 [1973] (
** Frullania subvalida S.Hatt. et Thaithong, J. Jap. Bot. 53 (6): 173, 1978 (
*** Frullania svihlana S.Hatt., J. Hattori Bot. Lab. 54: 180, 1983 (
* Frullania taiheizana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 241, 1934 (
** Frullania tamsuina Steph., Sp. Hepat. (Stephani) 4: 444, 1910 (
*** Frullania taradakensis Steph., Sp. Hepat. (Stephani) 4: 352, 1910 (
** Frullania tenuirostris Steph., Sp. Hepat. (Stephani) 4: 462, 1911 (
** Frullania togashiana S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 118, 1975 (
** Frullania tubercularis S.Hatt. et P.J.Lin, J. Jap. Bot. 60 (4): 107, 1985 (
*** Frullania usamiensis Steph., Bull. Herb. Boissier 5 (2): 91, 1897 (
** Frullania valdiviensis J.B.Jack et Steph., Hedwigia 33 (3): 149, 1894 (
** Frullania valida Steph., Sp. Hepat. (Stephani) 4: 402, 1910 (
** Frullania variegata Steph., Hedwigia 33 (3): 149, 1894 (
* Frullania victoriensis Steph., Sp. Hepat. (Stephani) 4: 418, 1910 (
*** Frullania vittiana S.Hatt., Bryologist 90 (4): 368, 1987 [1988] (
** Frullania wangii S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 146, 1985 (
*** Frullania yuennanensis Steph., Hedwigia 33 (3): 161, 1894 (
** Frullania yuennanensis var. siamensis (N.Kitag., Thaithong et S.Hatt.) S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 148, 1985 (
** Frullania yuzawana S.Hatt., J. Hattori Bot. Lab. 49: 157, 1981 (
** Frullania zangii S.Hatt. et P.J.Lin, J. Hattori Bot. Lab. 59: 149, 1985 (
** Frullania zennoskeana S.Hatt., J. Jap. Bot. 59 (10): 308, 1984 (
** sect. Acutilobae Verd., Ann. Bryol., Suppl. 1: 44, 1930 (
** Frullania allanii E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 371, 1949 (
*** Frullania clavata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 428, 1845 (
** Frullania hamaticoma Steph., Hedwigia 28 (3): 158, 1889 (
*** Frullania monocera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 418, 1845 (
** Frullania monocera var. acutiloba (Mitt.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (
** Frullania monocera var. depauperata S.Hatt., J. Hattori Bot. Lab. 57: 419, 1984 (
** Frullania monocera var. schiffneri (Verd.) S.Hatt., J. Hattori Bot. Lab. 46: 120, 1979 (
*** Frullania monocera var. subhampeana (E.A.Hodgs.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (
*** Frullania monocera var. undulata (Kamim.) Hentschel et von Konrat, Phytotaxa 220 (2): 136, 2015 (
** Frullania osumiensis (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 16: 87, 1956 (
* Frullania pallidula S.Hatt., Beih. Nova Hedwigia 90: 152, 1988 (
** Frullania pseudomonocera S.Hatt., J. Hattori Bot. Lab. 60: 216, 1986 (
** Frullania seriata Gottsche, Hedwigia 28 (3): 160, 1889 (
** Frullania spinistipula Steph., Sp. Hepat. (Stephani) 4: 463, 1911 (
** Frullania streimannii S.Hatt., J. Hattori Bot. Lab. 54: 176, 1983 (
*** sect. Australes Verd., Ann. Bryol., Suppl. 1: 58, 1930 (
*** Frullania anomala E.A.Hodgs., Trans. & Proc. Roy. Soc. New Zealand 77 (3): 374, 1949 (
* Frullania baileyana Steph., Sp. Hepat. (Stephani) 4: 417, 1910 (
*** Frullania baladina Gottsche, Hedwigia 33 (3): 140, 1894 (
* Frullania belmorensis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 106, 1914 (
*** Frullania campanulata Sande Lac., Ned. Kruidk. Arch. 3: 422, 1854 [1855] (
** Frullania campanulata var. caduca Verd., Ann. Bryol., Suppl. 1: 41, 1930 (
** Frullania campanulata var. malesiaca (Verd.) S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 158, 1975 (
* Frullania cataractarum Steph., Sp. Hepat. (Stephani) 4: 657, 1911 (
* Frullania crawfordii Steph., Hedwigia 33 (3): 143, 1894 (
*** Frullania dentata S.Hatt., J. Hattori Bot. Lab. 38: 231, 1974 (
** Frullania dentata var. secernens S.Hatt., J. Hattori Bot. Lab. 65: 422, 1988 (
*** Frullania errans Verd., Ann. Bryol., Suppl. 1: 59, 1930 (
** Frullania errans var. angulistipula S.Hatt., J. Hattori Bot. Lab. 36: 431, 1972 [1973] (
*** Frullania fugax (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 445, 1845 (
** Frullania fulfordiae S.Hatt., Bryologist 90 (4): 365, 1987 [1988] (
*** Frullania glomerata (Lehm. et Lindenb.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 46, 1838 (
*** Frullania incumbens Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 162, 1855 (
*** Frullania inflexa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 120, 1860 [1861] (
** Frullania media (E.A.Hodgs.) S.Hatt., J. Hattori Bot. Lab. 54: 153, 1983 (
** Frullania mizutanii Kamim. et S.Hatt., J. Hattori Bot. Lab. 37: 524, 1973 (
*** Frullania obscurifolia Mitt., Philos. Trans. 168: 400, 1879 (
*** Frullania patagonica Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 88, 1911 (
** Frullania pentapleura Taylor, London J. Bot. 5: 402, 1846 (
** Frullania polyptera Taylor, London J. Bot. 5: 401, 1846 (
** Frullania polyptera var. angustata (Mitt.) S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (2): 74, 1975 (
** Frullania probosciphora Taylor, London J. Bot. 5: 402, 1846 (
** Frullania pulchella Herzog, Rev. Bryol. Lichénol. 23 (1/2): 60, 1954 (
*** Frullania sinskeana J.J.Engel et B.C.Tan, J. Hattori Bot. Lab. 60: 335, 1986 (
** Frullania socotrana Mitt., Trans. Roy Soc. Edinburgh 31: 335, 1888 (
* Frullania solanderiana Colenso, Trans. & Proc. New Zealand Inst. 21: 75, 1889 (
** Frullania subincumbens S.Hatt., Bryologist 90 (4): 367, 1987 [1988] (
* Frullania subtropica Steph., Sp. Hepat. (Stephani) 4: 416, 1910 (
* Frullania tjibodensis S.Hatt. et Thaithong, J. Jap. Bot. 52 (10): 289, 1977 (
** Frullania tuyamae S.Hatt. et Thaithong, J. Jap. Bot. 53 (6): 175, 1978 (
*** sect. Frullania
*** Frullania appalachiana R.M.Schust., Phytologia 53 (5): 366, 1983 (
*** Frullania azorica Sim-Sim, Sérgio, Mues et Kraut, Cryptog. Bryol. Lichénol. 16 (2): 112, 1995 (
*** Frullania catalinae A.Evans, Trans. Connecticut Acad. Arts 10 (1): 11, 1899 (
*** Frullania dilatata (L.) Dumort., Recueil Observ. Jungerm.: 13, 1835 (
** Frullania dilatata subsp. asiatica S.Hatt., J. Jap. Bot. 57 (9): 258, 1982 (
*** Frullania eboracensis Lehm., Nov. Stirp. Pug. 8: 14, 1844 (
*** Frullania ericoides (Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839 (
* Frullania ericoides var. laxa (Gottsche, Lindenb. et Nees) Schiffn., Consp. Hepat. Arch. Ind.: 324, 1898 (
* Frullania ericoides var. minor Kamim., Bull. Kochi Gakuen Jun. Coll. 2: 22, 1971 (
** Frullania ericoides var. verrucosa (Kamim.) Hentschel et von Konrat, Phytotaxa 220 (2): 134, 2015 (
*** Frullania fragilifolia (Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 437, 1845 (
** Frullania fuegiana Steph., Sp. Hepat. (Stephani) 4: 428, 1910 (
** Frullania hattoriana J.D.Godfrey et G.Godfrey, J. Hattori Bot. Lab. 48: 321, 1980 (
** Frullania koponenii S.Hatt., Ann. Bot. Fenn. 15 (2): 111, 1978 (
*** Frullania muscicola Steph., Hedwigia 33 (3): 146, 1894 (
*** Frullania oakesiana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 225, 1869 (
** Frullania oakesiana subsp. takayuensis (Steph.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 195, 1992 (
*** Frullania parvistipula Steph., Sp. Hepat. (Stephani) 4: 397, 1910 (
** Frullania sabaliana R.M.Schust., Phytologia 53 (5): 365, 1983 (
** Frullania semivillosa Lindenb. et Gottsche, Syn. Hepat. 5: 774, 1847 (
** Frullania stylifera (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 210, 1992 (
* Frullania subdilatata C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 349, 1906 (
*** Frullania virginica Lehm., Nov. Stirp. Pug. 8: 19, 1844 (
** sect. Irregulares E.A.Hodgs. ex S.Hatt., J. Hattori Bot. Lab. 54: 143, 1983 (
* Frullania astrolabea Steph., Sp. Hepat. (Stephani) 4: 460, 1910 (
*** Frullania deplanata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 161, 1855 (
*** Frullania morobensis S.Hatt. et Streimann, J. Hattori Bot. Lab. 59: 109, 1985 (
*** Frullania patula Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 159, 1854 (
*** Frullania scandens Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 258, 1843 (
** sect. Planae R.M.Schust., Phytologia 57 (5): 372, 1985 (
*** Frullania plana Sull., Mem. Amer. Acad. Arts (n.ser.) 4: 175, 1849 (
*** subg. Homotropantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 35, 1884 (
*** Frullania deflexa Mitt., Bonplandia 10 (2): 19, 1862 (
*** Frullania integristipula (Nees) Nees, Syn. Hepat. 3: 431, 1845 (
** Frullania integristipula var. emarginata Verd., Ann. Bryol. 2: 153, 1929 (
** Frullania macrophylla S.Hatt., J. Hattori Bot. Lab. 47: 220, 1980 (
** Frullania sabahana S.Hatt., J. Hattori Bot. Lab. 40: 493, 1976 (
** Frullania sackawana Steph., Bull. Herb. Boissier 5 (2): 91, 1897 (
** Frullania sarawakensis S.Hatt., J. Hattori Bot. Lab. 40: 496, 1976 (
** Frullania umbonata Mitt., Sp. Hepat. (Stephani) 4: 579, 1911 (
*** Frullania utriculata Steph., Hedwigia 33 (3): 152, 1894 (
*** sect. Fallaces Verd., Rev. Bryol. Lichénol. 1: 112, 1928 (
* Frullania fallax Gottsche, Syn. Hepat. 3: 432, 1845 (
*** Frullania intermedia (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 434, 1845 (
** Frullania intermedia subsp. morokensis (Steph.) S.Hatt., J. Hattori Bot. Lab. 47: 194, 1980 (
* Frullania intermedia var. non-apiculata S.Hatt., J. Hattori Bot. Lab. 39: 291, 1975 (
** Frullania novoguineensis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 37, 1890 (
** Frullania regularis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 38, 1890 (
*** sect. Nodulosae Verd., Rev. Bryol. Lichénol. 1: 116, 1928 (
* Frullania brotheri Steph., Hedwigia 33 (3): 150, 1894 (
** Frullania hamata Steph., Sp. Hepat. (Stephani) 4: 582, 1911 (
** Frullania leeuwenii Verd., Nova Guinea 14: 545, 1930 (
*** Frullania nodulosa (Reinw., Blume et Nees) Nees, Syn. Hepat. 3: 433, 1845 (
** sect. Remotilobae Verd., Rev. Bryol. Lichénol. 1: 119, 1928 (
*** Frullania heteromorpha Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 38, 1890 (
** Frullania remotiloba Steph., Hedwigia 33 (3): 152, 1894 (
* subg. Mammillosae S.Hatt., J. Hattori Bot. Lab. 60: 226, 1986 (
** Frullania huerlimannii S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 2 (3): 84, 1976 (
* Frullania huerlimannii var. dioica S.Hatt., J. Hattori Bot. Lab. 57: 413, 1984 (
** Frullania involvens S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 526, 1973 (
** Frullania iriomotensis S.Hatt., J. Jap. Bot. 55 (5): 133, 1980 (
** Frullania mammillosa S.Hatt., J. Hattori Bot. Lab. 43: 424, 1977 [1978] (
*** Frullania meijeri S.Hatt., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 17 (4): 307, 1974 (
** Frullania notarisii Steph., Sp. Hepat. (Stephani) 4: 651, 1911 (
** Frullania papulosa Steph., Sp. Hepat. (Stephani) 4: 654, 1911 (
** Frullania rudolfiana S.Hatt., J. Hattori Bot. Lab. 36: 437, 1972 [1973] (
*** Frullania thiersiae S.Hatt., Beih. Nova Hedwigia 90: 156, 1988 (
** Frullania tixieri S.Hatt., J. Jap. Bot. 51 (7): 193, 1976 (
*** subg. Meteoriopsis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 37, 1884 (
* Frullania caldensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 89, 1876 [1877] (
** Frullania evoluta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861] (
*** Frullania tagawana (S.Hatt. et Thaithong) S.Hatt., J. Hattori Bot. Lab. 59: 160, 1985 (
*** sect. Intumescentes R.M.Schust., Phytologia 57 (5): 370, 1985 (
*** Frullania aculeata Taylor, London J. Bot. 5: 407, 1846 (
** Frullania ambronnii Steph., Biblioth. Bot. 87 (2): 242, 1916 (
*** Frullania atrata (Sw.) Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 12: 51, 1839 (
*** Frullania beyrichiana (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845 (
*** Frullania bicornistipula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 46, 1884 (
*** Frullania brasiliensis Raddi, Critt. Brasil.: 12, 1822 (
** Frullania brasiliensis var. elegantula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 50, 1884 (
* Frullania breuteliana Gottsche, Syn. Hepat. 3: 461, 1845 (
* Frullania compacta Gottsche, Sp. Hepat. (Stephani) 4: 493, 1911 (
* Frullania crenulifolia J.B.Jack et Steph., Hedwigia 31 (1): 14, 1892 (
** Frullania crispiloba Steph., Hedwigia 33 (3): 156, 1894 (
** Frullania curviramea Steph., Sp. Hepat. (Stephani) 4: 684, 1911 (
*** Frullania ecuadorensis Steph., Sp. Hepat. (Stephani) 4: 526, 1911 (
** Frullania formosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 46, 1884 (
* Frullania granatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 173 (79), 1864 (
*** Frullania griffithsiana Gottsche, Syn. Hepat. 4: 466, 1846 (
* Frullania guadalupensis Gottsche, Sp. Hepat. (Stephani) 4: 496, 1911 (
* Frullania gualaquizana Steph., Sp. Hepat. (Stephani) 4: 531, 1911 (
** Frullania hamiflora Herzog et L.Clark, Bryologist 56 (3): 180, 1953 (
* Frullania humilis Spruce, Mem. Torrey Bot. Club 1 (3): 119, 1890 (
*** Frullania intumescens (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845 (
* Frullania laticaulis Spruce, Mem. Torrey Bot. Club 1 (3): 120, 1890 (
*** Frullania lobatohastata Steph., Sp. Hepat. (Stephani) 4: 499, 1911 (
* Frullania longistipula var. apiculata Demaret et Vanden Berghen, Bull. Jard. Bot. État Bruxelles 20 (1): 4, 1950 (
*** Frullania macrocephala (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 460, 1845 (
** Frullania madothecoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 47, 1884 (
** Frullania meridana Steph., Sp. Hepat. (Stephani) 4: 500, 1911 (
* Frullania microcephala Gottsche, Mexik. Leverm.: 251, 1863 (
*** Frullania montagnei Gottsche, Syn. Hepat. 3: 456, 1845 (
** Frullania moritziana Lindenb. et Gottsche, Syn. Hepat. 5: 782, 1847 (
** Frullania osculatiana De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 236, 1857 (
*** Frullania paradoxa Lehm. et Lindenb., Syn. Hepat. 3: 462, 1845 (
** Frullania pearceana Steph., Sp. Hepat. (Stephani) 4: 515, 1911 (
*** Frullania pittieri Steph., Primit. fl. costar.: 113, 1892 [1893] (
** Frullania rigescens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 52, 1884 (
*** Frullania setigera Steph., Hedwigia 33 (3): 159, 1894 (
** Frullania speciosa Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 59, 1950 (
** Frullania supradecomposita (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 431, 1845 (
* Frullania trianae Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 173, 1864 (
** Frullania triquetra Lindenb. et Gottsche, Syn. Hepat. 5: 780, 1847 (
* Frullania trollii Herzog, Beih. Bot. Centralbl. 61B (3): 575, 1942 (
*** Frullania uleana Steph., Hedwigia 33 (3): 155, 1894 (
** sect. Meteoriopsis Uribe, von Konrat et Hentschel, Phytotaxa 220 (2): 132, 2015 (
*** Frullania convoluta Lindenb. et Hampe, Linnaea 24 (3): 303, 1851 [1852] (
* Frullania convoluta var. ampliata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 129, 1951 [1952] (
*** Frullania darwinii Gradst. et Uribe, Cryptog. Bryol. 25 (4): 296, 2004 (
*** Frullania dulimensis Uribe, Cryptog. Bryol. 27 (3): 309, 2006 (
*** Frullania grandifolia Steph., Sp. Hepat. (Stephani) 4: 684, 1911 (
*** Frullania peruviana Gottsche, Syn. Hepat. 4: 465, 1846 (
*** Frullania phalangiflora Steph., Biblioth. Bot. 87 (2): 247, 1916 (
*** Frullania weberbaueri Steph., Sp. Hepat. (Stephani) 4: 510, 1911 (
*** sect. Obtusilobae Verd., Ann. Bryol., Suppl. 1: 81, 1930 (
** Frullania angulata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 169, 1863 (
** Frullania angulata var. laciniata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 72, 1976 (
** Frullania apicalis Mitt., Philos. Trans. 168: 401, 1879 (
** Frullania apicalis var. camerunensis Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 53, 1976 (
* Frullania borbonica Lindenb., Syn. Hepat. 3: 455, 1845 (
*** Frullania capensis Gottsche, Syn. Hepat. 3: 449, 1845 (
** Frullania donnellii Austin, Bull. Torrey Bot. Club 6 (52): 301, 1879 (
** Frullania eplicata Steph., Sp. Hepat. (Stephani) 4: 679, 1911 (
** Frullania imerinensis Steph., Sp. Hepat. (Stephani) 4: 484, 1911 (
*** Frullania kunzei (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 449, 1845 (
** Frullania kunzei var. maritima R.M.Schust., Phytotaxa 220 (2): 135, 2015 (
** Frullania longistipula Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 199, 1891 [1892] (
*** Frullania meyeniana Lindenb., Syn. Hepat. 3: 455, 1845 (
** Frullania meyeniana var. dioica S.Hatt., J. Hattori Bot. Lab. 43: 426, 1977 [1978] (
** Frullania onraedtii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 60, 1976 (
** Frullania papuana Verd., Ann. Bryol., Suppl. 1: 82, 1930 (
*** Frullania schimperi Nees, Syn. Hepat. 3: 454, 1845 (
** Frullania schimperi var. laciniata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 46 (1/2): 59, 1976 (
*** subg. Microfrullania (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 34, 1992 (
*** Frullania fertilis De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 235, 1857 (
*** Frullania knightbridgei von Konrat et de Lange, PhytoKeys 8: 28, 2012 (
*** Frullania magellanica F.Weber et Nees, Syn. Hepat. 3: 446, 1845 (
** Frullania magellanica subsp. tristaniana (S.W.Arnell) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 59, 2013 (
* Frullania matafaoica H.A.Mill., Phytologia 47 (4): 322, 1981 (
*** Frullania toropuku von Konrat, de Lange et Larraín, Polish Bot. J. 58 (2): 439, 2013 (
*** sect. Amphijubula (R.M.Schust.) von Konrat, Hentschel, Heinrichs et Braggins, Bryologist 114 (1): 53, 2011 (
*** Frullania lobulata (Hook.) Hook. et Nees, Syn. Hepat. 3: 445, 1845 (
*** Frullania microcaulis Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 29 (1/4): 172, 1922 [1923] (
*** Frullania truncatistyla von Konrat, Hentschel, Heinrichs et Braggins, Bryologist 114 (1): 63, 2011 (
*** sect. Microfrullania (R.M.Schust.) von Konrat et Hentschel, Phytotaxa 220 (2): 133, 2015 (
*** Frullania chevalieri (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 34, 1992 (
*** Frullania microscopica Pearson, J. Linn. Soc., Bot. 46 (305): 33, 1922 (
** Frullania neocaledonica J.J.Engel, Novon 9 (3): 344, 1999 (
*** Frullania parhamii (R.M.Schust.) R.M.Schust. ex von Konrat, L.Söderstr. et A.Hagborg, Telopea 13 (3): 407, 2011 (
*** sect. Regulares Verd., Ann. Bryol., Suppl. 1: 133, 1930 (
*** Frullania junghuhniana Gottsche, Syn. Hepat. 3: 444, 1845 (
** Frullania junghuhniana var. bisexualis S.Hatt., J. Hattori Bot. Lab. 40: 485, 1976 (
** Frullania junghuhniana var. tenella (Sande Lac.) Grolle et S.Hatt., Misc. Bryol. Lichenol. 9 (6): 123, 1982 (
** Frullania mcveanii S.Hatt., J. Hattori Bot. Lab. 37: 55, 1973 (
** Frullania pseudomeyeniana S.Hatt., J. Hattori Bot. Lab. 60: 231, 1986 (
*** Frullania rostrata (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 445, 1845 (
** Frullania scalaris S.Hatt., J. Hattori Bot. Lab. 43: 432, 1977 [1978] (
** subg. Saccophora Verd., Ann. Bryol. 2: 121, 1929 (
*** Frullania gaudichaudii (Nees et Mont.) Nees et Mont., Syn. Hepat. 3: 435, 1845 (
** Frullania gaudichaudii var. ceylanica (Nees) S.Hatt., J. Hattori Bot. Lab. 47: 104, 1980 (
*** Frullania hedrantha S.Hatt. et Kamim., J. Hattori Bot. Lab. 37: 519, 1973 (
** Frullania immersa Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 315, 1896 (
** Frullania pancheri Gottsche, Hedwigia 33 (3): 159, 1894 (
** Frullania papillilobula S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 141, 1975 (
** Frullania sublignosa Steph., Hedwigia 33 (3): 148, 1894 (
* subg. Steerea (S.Hatt. et Kamim.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 32, 1992 (
*** Frullania clemensiana Verd., Ned. Kruidk. Arch. (ser. 3) 42 (2): 493, 1932 (
*** subg. Thyopsiella Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 41, 1884 (
*** Frullania acicularis Hentschel et von Konrat, Phytotaxa 220 (2): 134, 2015 (
** Frullania alstonii Verd., Ann. Bryol., Suppl. 1: 76, 1930 (
** Frullania aoshimensis Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 4 (1): 64, 1929 (
*** Frullania appendiculata Steph., Bull. Herb. Boissier 5 (2): 88, 1897 (
*** Frullania asagrayana Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 14, 1842 (
*** Frullania calcarifera Steph., Bol. Soc. Brot. 4: 241, 1886 [1887] (
*** Frullania californica (M.Howe) A.Evans, Trans. Connecticut Acad. Arts 10 (1): 25, 1899 (
*** Frullania densiloba Steph. ex A.Evans, Proc. Wash. Acad. Sci. 8: 157, 1906 (
*** Frullania franciscana M.Howe, Erythea 2 (6): 99, 1894 (
** Frullania iwatsukii S.Hatt., J. Hattori Bot. Lab. 35: 240, 1972 (
*** Frullania microphylla (Gottsche) Pearson, J. Bot. 32: 328, 1894 (
*** Frullania moniliata (Reinw., Blume et Nees) Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 13, 1842 (
*** Frullania nisquallensis Sull., Mem. Amer. Acad. Arts (n.ser.) 4: 175, 1849 (
*** Frullania polysticta Lindenb., Syn. Hepat. 3: 440, 1845 (
** Frullania pseudoalstonii Tsudo et J.Haseg., Bryol. Res. 9 (3): 44, 2006 (
** Frullania punctata Reimers, Hedwigia 71 (1/2): 36, 1931 (
** Frullania schaefer-verwimpii Yuzawa et S.Hatt., J. Jap. Bot. 64 (2): 37, 1989 (
** Frullania selwyniana Pearson, List. Canad. Hepat.: 1, 1890 (
*** Frullania sergiae Sim-Sim, Fontinha, Mues et Lion, Nova Hedwigia 71 (1/2): 186, 2000 (
*** Frullania subarctica Vilnet, Borovich. et Bakalin, Phytotaxa 173 (1): 67, 2014 (
*** Frullania tamarisci (L.) Dumort., Recueil Observ. Jungerm.: 13, 1835 (
*** Frullania teneriffae (F.Weber) Nees, Naturgesch. Eur. Leberm. 3: 239, 1838 (
** Frullania trigona L.Clark, Jovet-Ast et Frye, Bryologist 50 (1): 52, 1947 (
Incertae sedis
* Frullania affinis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 257, 1843 (
* Frullania allionii Steph., Sp. Hepat. (Stephani) 4: 394, 1910 (
* Frullania alpina Steph., Sp. Hepat. (Stephani) 4: 533, 1911 (
** Frullania alternans Nees, Syn. Hepat. 3: 430, 1845 (
* Frullania apertilobula Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 478, 1947 (
** Frullania armata Herzog et L.Clark, Bryologist 57 (1): 36, 1954 (
*** Frullania bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 226, 1869 (
*** Frullania boveana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 244, 1885 (
* Frullania caespitans Beauverd, Sp. Hepat. (Stephani) 6: 537, 1924 (
* Frullania canaliculata Gottsche, Sp. Hepat. (Stephani) 4: 391, 1910 (
** Frullania capillaris Steph., Sp. Hepat. (Stephani) 4: 616, 1911 (
* Frullania cavallii Gola, Ann. Bot. (Rome) 6 (2): 275, 1907 (
* Frullania chiapasana Steph., Sp. Hepat. (Stephani) 4: 392, 1910 (
* Frullania chilcootiensis Steph., Bot. Jahrb. Syst. 8 (2): 98, 1886 (
* Frullania chiovendae Gola, Ann. Bot. (Rome) 13 (1): 71, 1914 (
** Frullania ciliata Lindenb. et Gottsche, Syn. Hepat. 5: 775, 1847 (
** Frullania cinchonae Gottsche, Syn. Hepat. 3: 455, 1845 (
* Frullania complicata Steph., Sp. Hepat. (Stephani) 4: 671, 1911 (
* Frullania cordaeana Lindenb., Syn. Hepat. 3: 463, 1845 (
** Frullania cuneatistipula Steph., Sp. Hepat. (Stephani) 6: 538, 1924 (
* Frullania cuspiloba Steph., Sp. Hepat. (Stephani) 4: 394, 1910 (
* Frullania dispar Nees, Syn. Hepat. 3: 429, 1845 (
* Frullania guatemalensis Steph., Sp. Hepat. (Stephani) 4: 497, 1911 (
*** Frullania inflata Gottsche, Syn. Hepat. 3: 424, 1845 (
** Frullania inflata var. communis R.M.Schust., Phytologia 57 (5): 372, 1985 (
** Frullania inflata var. dioica S.Hatt. et Thaithong, J. Hattori Bot. Lab. 44: 185, 1978 (
** Frullania inflata var. mayebarae (S.Hatt.) K.Yamada, Misc. Bryol. Lichenol. 6 (9): 163, 1974 (
* Frullania laetevirens Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 420, 1845 (
* Frullania larjiana Sushil K.Singh et D.K.Singh, J. Bryol. 27 (2): 105, 2005 (
* Frullania larjiana var. didyhatii S.N.Srivast. et M.Rai, Geophytology 41 (1/2): 109, 2011 (
* Frullania leana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869 (
* Frullania madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 364, 1882 (
* Frullania mauritiana Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 227, 1869 (
*** Frullania mirabilis J.B.Jack et Steph., Hedwigia 31 (1): 15, 1892 (
** Frullania monoica Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 67, 1900 (
* Frullania ocanniensis Steph., Sp. Hepat. (Stephani) 6: 542, 1924 (
*** Frullania platycalyx Herzog, Feddes Repert. Spec. Nov. Regni Veg. 55 (1): 10, 1952 (
* Frullania pyricalycina Steph., Sp. Hepat. (Stephani) 4: 394, 1910 (
** Frullania quillotensis (Nees et Mont.) Nees et Mont., Syn. Hepat. 3: 427, 1845 (
** Frullania reicheana Steph., Sp. Hepat. (Stephani) 4: 427, 1910 (
* Frullania sabanetica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 170, 1864 (
* Frullania semienana Gola, Ann. Bot. (Rome) 13 (1): 71, 1914 (
** Frullania serrifolia Steph., Sp. Hepat. (Stephani) 4: 526, 1911 (
** Frullania subpyricalycina Herzog, Rev. Bryol. Lichénol. 23 (1/2): 57, 1954 (
** Frullania subtruncata Steph., Sp. Hepat. (Stephani) 4: 389, 1910 (
** Frullania turfosa Lindenb. et Gottsche, Syn. Hepat. 5: 779, 1847 (
** Frullania udarii V.Nath et Ajit P.Singh, Curr. Sci. 91 (6): 744, 2006 (
** Frullania valparaisiana Lehm., Nov. Stirp. Pug. 10: 17, 1857 (
** Frullania wagneri Steph., Sp. Hepat. (Stephani) 4: 392, 1910 (
by J. Hentschel and M. von Konrat
The systematic placement of Neohattoria herzogii has been contentious since its description six decades ago. It has been interpreted as either a member of the genus Frullania or segregated into its own genus, Neohattoria, due to morphological similarities with both Frullania and Jubula. We follow Larrain et al. (2015) that provided molecular evidence supporting the recognition of the genus Neohattoria and its inclusion within the Jubulaceae, together with Jubula and Nipponolejeunea.
*** Jubula Dumort., Commentat. Bot. (Dumortier): 112, 1822 (
*** Jubula blepharophylla Grolle, J. Hattori Bot. Lab. 28: 47, 1965 (
** Jubula hattorii Udar et V.Nath, Misc. Bryol. Lichenol. 8 (3): 49, 1978 (
** Jubula hattorii var. muthukuzhiana A.E.D.Daniels et P.Daniel, Bryofl. Southernm. W Ghats: 181, 2013 (
** Jubula himalayensis S.C.Srivast. et D.Sharma, Proc. Indian Acad. Sci. Pl. Sci. 100 (2): 85, 1990 (
*** Jubula hutchinsiae (Hook.) Dumort., Syll. Jungerm. Europ.: 36, 1831 (
** Jubula hutchinsiae subsp. australiae Pócs et A.Cairns, Nova Hedwigia 86 (1/2): 232, 2008 (
** Jubula hutchinsiae subsp. bogotensis (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 215, 1928 (
*** Jubula hutchinsiae subsp. caucasica Konstant. et Vilnet, Arctoa 20: 234, 2011 (
*** Jubula hutchinsiae subsp. japonica (Steph.) Horik. et Ando, J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 6: 300, 1954 (
*** Jubula hutchinsiae subsp. javanica (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 216, 1928 (
*** Jubula hutchinsiae subsp. pennsylvanica (Steph.) Verd., Ann. Cryptog. Exot. 1 (2): 215, 1928 (
** Jubula kwangsiensis C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 89, 1984 (
*** Neohattoria Kamim., J. Jap. Bot. 37 (7): 218, 1962 (
*** Neohattoria herzogii (S.Hatt.) Kamim., J. Jap. Bot. 37 (7): 218, 1962 (
*** Nipponolejeunea S.Hatt., Bull. Tokyo Sci. Mus. 11: 124, 1944 (
*** Nipponolejeunea pilifera (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 125, 1944 (
*** Nipponolejeunea subalpina (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 125, 1944 (
by S.R. Gradstein, T. Pócs and R.-L. Zhu with contributions by G. Dauphin (Ceratolejeunea), X. He (Pycnolejeunea), A.-L. Ilkiu-Borges (Prionolejeunea), E. Reiner-Drehwald (Rectolejeunea, Lejeunea), A. Sass-Gyarmati (Lopholejeunea), A. Schäfer-Verwimp (Diplasiolejeunea) and P. Sukkharak (Thysananthus)
The subdivision of Lejeuneaceae follows
** Acanthocoleus R.M.Schust., Bull. Torrey Bot. Club 97 (6): 339, 1970 [1971] (
*** Acanthocoleus aberrans (Lindenb. et Gottsche) Kruijt, Bryophyt. Biblioth. 36: 62, 1988 (
*** Acanthocoleus aberrans var. laevis Gradst., Fl. Neotrop. Monogr. 62: 193, 1994 (
*** Acanthocoleus chrysophyllus (Lehm.) Kruijt, Bryophyt. Biblioth. 36: 72, 1988 (
** Acanthocoleus elgonensis Gyarmati et Pócs, Cryptog. Bryol. 35 (2): 120, 2014 (
*** Acanthocoleus gilvus (Gottsche) Kruijt, Bryophyt. Biblioth. 36: 79, 1988 (
*** Acanthocoleus javanicus (Steph.) Kruijt, Bryophyt. Biblioth. 36: 85, 1988 (
*** Acanthocoleus juddii Kruijt, Bryophyt. Biblioth. 36: 93, 1988 (
** Acanthocoleus madagascariensis (Steph.) Kruijt, Bryophyt. Biblioth. 36: 98, 1988 (
*** Acanthocoleus trigonus (Nees et Mont.) Gradst., Contr. Univ. Michigan Herb. 18: 101, 1992 (
** Acanthocoleus yoshinaganus (S.Hatt.) Kruijt, Bryophyt. Biblioth. 36: 105, 1988 (
*** Blepharolejeunea S.W.Arnell, Svensk Bot. Tidskr. 56 (2): 335, 1962 (
*** Blepharolejeunea chimantaensis van Slageren et Kruijt, Beih. Nova Hedwigia 80: 126, 1985 (
*** Blepharolejeunea fuegiana (Besch. et C.Massal.) Gradst., Beih. Nova Hedwigia 80: 130, 1985 (
*** Blepharolejeunea incongrua (Lindenb. et Gottsche) van Slageren et Kruijt, Beih. Nova Hedwigia 80: 133, 1985 (
*** Blepharolejeunea saccata (Steph.) van Slageren et Kruijt, Beih. Nova Hedwigia 80: 138, 1985 (
*** Blepharolejeunea securifolia (Steph.) R.M.Schust., Phytologia 45 (5): 424, 1980 (
*** Brachiolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Brachiolejeunea conduplicata (Steph.) Gradst., Fl. Neotrop. Monogr. 62: 175, 1994 (
*** Brachiolejeunea fernandeziana S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 18, 1957 (
*** Brachiolejeunea laxifolia (Taylor) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Brachiolejeunea leiboldiana (Gottsche et Lindenb.) Schiffn., Hedwigia 33 (4): 182, 1894 (
*** Brachiolejeunea phyllorhiza (Nees) Kruijt et Gradst., Nova Hedwigia 43 (3/4): 299, 1986 (
*** Brachiolejeunea spruceana (C.Massal.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Dicranolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Dicranolejeunea axillaris (Nees et Mont.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
* Dicranolejeunea bovonei Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 27 (2/4): 248, 1920 (
*** Lindigianthus Kruijt et Gradst., Beih. Nova Hedwigia 80: 165, 1985 (
*** Lindigianthus cipaconeus (Gottsche) Kruijt et Gradst., Beih. Nova Hedwigia 80: 166, 1985 (
*** Odontolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
*** Odontolejeunea decemdentata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 175, 1912 (
*** Odontolejeunea lunulata (F.Weber) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Odontolejeunea rhomalea (Spruce) Steph., Sp. Hepat. (Stephani) 5: 176, 1912 (
*** Neurolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (
*** Neurolejeunea breutelii (Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (1): 13, 1907 (
** Neurolejeunea breutelii var. africana Pócs, Herzogia 28 (1): 63, 2015 (
*** Neurolejeunea catenulata (Nees) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (
*** Neurolejeunea sastreana Gradst., Bryologist 92 (3): 345, 1989 (
*** Neurolejeunea seminervis (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (
*** Stictolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (
*** subg. Leptostictolejeunea R.M.Schust., Phytologia 56 (2): 70, 1984 (
*** Stictolejeunea balfourii (Mitt.) E.W.Jones, J. Bryol. 9 (1): 50, 1976 (
*** Stictolejeunea balfourii var. bekkeri Gradst., Beih. Nova Hedwigia 80: 214, 1985 (
*** Stictolejeunea iwatsukii Mizut., J. Hattori Bot. Lab. 44: 134, 1978 (
*** subg. Stictolejeunea
*** Stictolejeunea squamata (Willd.) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (
** Dactylophorella R.M.Schust., Phytologia 45 (5): 427, 1980 (
*** Dactylophorella muricata (Gottsche) R.M.Schust., Phytologia 45 (5): 427, 1980 (
*** Metalejeunea Grolle, Bryophyt. Biblioth. 48: 17, 1995 (
** Metalejeunea crassitexta (J.B.Jack et Steph.) Pócs, Telopea 13 (3): 456, 2011 (
*** Metalejeunea cucullata (Reinw., Blume et Nees) Grolle, Bryophyt. Biblioth. 48: 100, 1995 (
*** Metalejeunea winkleri R.L.Zhu et Grolle, Nova Hedwigia 74 (3/4): 498, 2002 (
*** Pictolejeunea Grolle, Feddes Repert. 88 (4): 248, 1977 (
*** Pictolejeunea levis Grolle et M.E.Reiner, J. Bryol. 27 (3): 281, 2005 (
*** Pictolejeunea mizutanii Grolle, Feddes Repert. 88 (4): 255, 1977 (
** Pictolejeunea piconii Pócs, Acta Bot. Hung. 49 (1/2): 110, 2007 (
*** Pictolejeunea picta (Steph.) Grolle, Feddes Repert. 88 (4): 252, 1977 (
*** Pictolejeunea reginae Ilk.-Borg., Brittonia 54 (4): 318, 2002 [2003] (
*** Pictolejeunea sprucei Grolle, Feddes Repert. 88 (4): 249, 1977 (
*** Ceratolejeunea (Spruce) J.B.Jack et Steph., Hedwigia 31 (1): 16, 1892 (
*** subg. Ceratolejeunea
*** Ceratolejeunea andringitrae Pócs, Polish Bot. J. 56 (2): 144, 2011 (
** Ceratolejeunea atlantica Alvarenga et Ilk.-Borg., Nova Hedwigia 86 (1/2): 238, 2008 (Ilkiu-Borges and Alvarenga 2008).
** Ceratolejeunea belangeriana (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 396, 1913 (
** Ceratolejeunea beninensis E.W.Jones et Vanden Berghen, Bull. Jard. Bot. État Bruxelles 21 (1/2): 63, 1951 (
*** Ceratolejeunea brevinervis (Spruce) A.Evans, Bull. Torrey Bot. Club 32 (6): 282, 1905 (
*** Ceratolejeunea ceratantha (Nees et Mont.) Schiffn., Bot. Jahrb. Syst. 23 (5): 582, 1897 (
** Ceratolejeunea coalita (Ångstr.) Steph., Sp. Hepat. (Stephani) 5: 402, 1913 (
*** Ceratolejeunea coarina (Gottsche) Schiffn., Hepat. (Engl.-Prantl): 125, 1893 (
*** Ceratolejeunea confusa R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 313, 1956 (
*** Ceratolejeunea cornuta (Lindenb.) Steph., Pflanzenw. Ost-Afrikas C: 65, 1895 (
*** Ceratolejeunea cubensis (Mont.) Schiffn., Hepat. (Engl.-Prantl): 125, 1893 (
*** Ceratolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 407, 1913 (
*** Ceratolejeunea fallax (Lehm. et Lindenb.) Bonner, Candollea 14: 189, 1953 (
*** Ceratolejeunea filaria (Taylor) Steph., Sp. Hepat. (Stephani) 5: 412, 1913 (
** Ceratolejeunea floribunda Steph., Sp. Hepat. (Stephani) 5: 412, 1913 (
*** Ceratolejeunea guianensis (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 416, 1913 (
* Ceratolejeunea karstenii Steph., Sp. Hepat. (Stephani) 5: 420, 1913 (
* Ceratolejeunea kuerschneri Eb.Fisch. et Vanderp., Beih. Nova Hedwigia 138: 87, 2010 (
*** Ceratolejeunea laetefusca (Austin) R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 306, 1956 (
* Ceratolejeunea ledermannii Steph., Sp. Hepat. (Stephani) 6: 399, 1923 (
*** Ceratolejeunea malleigera (Spruce) Steph., Sp. Hepat. (Stephani) 5: 422, 1913 (
** Ceratolejeunea maranhensis Silva Brito et Ilk.-Borg., Nova Hedwigia 95 (3/4): 424, 2012 (
** Ceratolejeunea minor Mizut., J. Hattori Bot. Lab. 49: 311, 1981 (
*** Ceratolejeunea minuta G.Dauphin, Fl. Neotrop. Monogr. 90: 66, 2003 (
** Ceratolejeunea moniliata Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 205, 1931 (
** Ceratolejeunea oculata (Gottsche) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (
*** Ceratolejeunea oxygonia Steph., Sp. Hepat. (Stephani) 5: 429, 1913 (
*** Ceratolejeunea papuliflora Steph., Sp. Hepat. (Stephani) 5: 430, 1913 (
*** Ceratolejeunea patentissima (Hampe et Gottsche) A.Evans, Bull. Torrey Bot. Club 32 (6): 286, 1905 (
*** Ceratolejeunea pungens Steph., Sp. Hepat. (Stephani) 5: 434, 1913 (
*** Ceratolejeunea rubiginosa Steph., Hedwigia 34 (5): 237, 1895 (
*** Ceratolejeunea saroltae Pócs, Polish Bot. J. 56 (2): 150, 2011 (
* Ceratolejeunea sinensis P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 232, 1964 (
** Ceratolejeunea singapurensis (Lindenb.) Schiffn., Consp. Hepat. Arch. Ind.: 273, 1898 (
*** Ceratolejeunea spinosa (Gottsche) Steph., Hedwigia 34 (5): 238, 1895 (
*** Ceratolejeunea temnantha (Spruce) M.E.Reiner, Cryptog. Bryol. 32 (2): 95, 2011 (
*** Ceratolejeunea umbonata Steph., Sp. Hepat. (Stephani) 5: 446, 1913 (
** Ceratolejeunea vitiensis Steph., Sp. Hepat. (Stephani) 5: 448, 1913 (
** Ceratolejeunea zenkeri Steph., Sp. Hepat. (Stephani) 5: 449, 1914 (
*** subg. Ceratophora R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (2): 294, 1956 (
*** Ceratolejeunea desciscens (Sande Lac.) Schiffn., Hepat. (Engl.-Prantl): 126, 1893 (
*** Ceratolejeunea globulifera Herzog, Rev. Bryol. Lichénol. 13: 23, 1942 (
*** Ceratolejeunea grandiloba J.B.Jack et Steph., Hedwigia 31 (1): 16, 1892 (
** Ceratolejeunea grandiloba subsp. inflata (Mizut.) Gradst., Phytotaxa 81 (1): 5, 2013 (
*** Ceratolejeunea szyszylowiczii (Loitl.) Steph., Sp. Hepat. (Stephani) 5: 443, 1913 (
Incertae sedis
** Ceratolejeunea aliena Herzog, Trans. Brit. Bryol. Soc. 2 (1): 71, 1952 (
*** Luteolejeunea Piippo, Acta Bot. Fenn. 132: 56, 1986 (
*** Luteolejeunea herzogii (Buchloh) Piippo, Acta Bot. Fenn. 132: 57, 1986 (
*** Otigoniolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 125, 1893 (
* Otigoniolejeunea crenulata Steph., Sp. Hepat. (Stephani) 6: 408, 1923 (
*** Otigoniolejeunea huctumalcensis (Lindenb. et Gottsche) Y.M.Wei, R.L.Zhu et Gradst., Phytotaxa 162 (4): 237, 2014 (
* Otigoniolejeunea ledermannii Steph., Sp. Hepat. (Stephani) 6: 409, 1923 (
*** Otigoniolejeunea portoricensis (Hampe et Gottsche) Y.M.Wei, R.L.Zhu et Gradst., Phytotaxa 162 (4): 237, 2014 (
* Aureolejeunea R.M.Schust., Phytologia 39 (6): 428, 1978 (
*** Aureolejeunea aurifera R.M.Schust., Phytologia 39 (6): 429, 1978 (
*** Aureolejeunea lumae (Herzog) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 121, 1985 (
*** Aureolejeunea paramicola (Herzog) R.M.Schust., Phytologia 61 (7): 446, 1987 (
*** Aureolejeunea quinquecarinata R.M.Schust., Phytologia 39 (6): 429, 1978 (
*** Aureolejeunea tonduzana (Steph.) Gradst., Phytotaxa 76 (3): 46, 2013 (
*** Cheilolejeunea (Spruce) Steph., Bot. Gaz. 15 (11): 284, 1890 (
** Cheilolejeunea sandvicensis (Prantl) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (
** subg. Cheilolejeunea
*** Cheilolejeunea acanthina (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009 (
*** Cheilolejeunea adnata (Kunze ex Lehm.) Grolle, J. Bryol. 9 (4): 529, 1977 [1978] (
** Cheilolejeunea adnata var. autoica Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 64, 2009 (
** Cheilolejeunea albovirens (Hook.f. et Taylor) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 127, 1962 (
*** Cheilolejeunea aneogyna (Spruce) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 440, 1900 (
** Cheilolejeunea ascensionis (Hook.f. et Taylor) Grolle, Haussknechtia 4: 45, 1988 (
*** Cheilolejeunea asperiflora (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009 (
** Cheilolejeunea australis Solari, Comun. Mus. Argent. Ci. Nat. “Bernardino Rivadavia,” Ci. Bot. 2 (11): 70, 1981 (
** Cheilolejeunea baumannii Hürl., Bauhinia 11 (3): 160, 1995 (
*** Cheilolejeunea beyrichii (Lindenb.) M.E.Reiner, Nova Hedwigia 83 (3/4): 474, 2006 (
** Cheilolejeunea boninensis Mizut., J. Hattori Bot. Lab. 51: 153, 1982 (
*** Cheilolejeunea caducifolia (Gradst. et Schäf.-Verw.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (
** Cheilolejeunea camerunensis S.W.Arnell, Svensk Bot. Tidskr. 52 (1): 63, 1958 (
** Cheilolejeunea celebensis (Steph.) Mizut., J. Hattori Bot. Lab. 36: 157, 1972 [1973] (
** Cheilolejeunea chenii R.L.Zhu et M.L.So, Taxon 48 (4): 663, 1999 (
*** Cheilolejeunea clypeata (Schwein.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (
** Cheilolejeunea compressa (Herzog) Grolle, Bryophyt. Biblioth. 48: 38, 1995 (
*** Cheilolejeunea conchifolia (A.Evans) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (
*** Cheilolejeunea cordigera (Steph.) Grolle, J. Bryol. 9 (4): 530, 1977 [1978] (
** Cheilolejeunea coronalis (Gottsche) R.M.Schust., Phytologia 45 (5): 431, 1980 (
* Cheilolejeunea curvatilobula (Herzog) Grolle, J. Bryol. 21 (1): 41, 1999 (
*** Cheilolejeunea decursiva (Sande Lac.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (
*** Cheilolejeunea discoidea (Lehm. et Lindenb.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea diversifolia Augier, Ann. Fac. Sci. Univ. Féd. Cameroun 11: 66, 1972 (
** Cheilolejeunea ecarinata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (1/2): 11, 1984 (
** Cheilolejeunea erostrata R.M.Schust., Phytologia 39 (6): 427, 1978 (
*** Cheilolejeunea exinnovata E.W.Jones, J. Bryol. 12 (1): 37, 1982 (
*** Cheilolejeunea fragrantissima (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980 (
* Cheilolejeunea fukiensis (P.C.Chen et P.C.Wu) Piippo, J. Hattori Bot. Lab. 68: 133, 1990 (
* Cheilolejeunea galliotii Steph., Sp. Hepat. (Stephani) 5: 656, 1914 (
** Cheilolejeunea germanii (Besch. et Spruce) Grolle, Acta Bot. Fenn. 125: 64, 1984 (
** Cheilolejeunea ghatensis G.Asthana, S.C.Srivast. et A.K.Asthana, Lindbergia 20 (2/3): 132, 1995 [1996] (
*** Cheilolejeunea gradsteinii (Grolle et Piippo) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (
** Cheilolejeunea grandibracteata Steph., Sp. Hepat. (Stephani) 5: 657, 1914 (
** Cheilolejeunea hamlinii Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982 (
** Cheilolejeunea hawaica Steph., Bull. Herb. Boissier 5 (10): 847, 1897 (
* Cheilolejeunea herzogiana Steph., Biblioth. Bot. 87 (2): 267, 1916 (
** Cheilolejeunea implexicaulis (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963 (
*** Cheilolejeunea insecta Grolle et Gradst., Taxon 50 (4): 1071, 2001 [2002] (
** Cheilolejeunea insignis Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 31 (1/2): 25, 1962 (
** Cheilolejeunea intricata (Steph.) J.J.Engel, Bryologist 79 (4): 514, 1976 [1977] (
** Cheilolejeunea invaginata R.M.Schust., Phytologia 39 (6): 427, 1978 (
** Cheilolejeunea jamaicensis Steph., Hedwigia 34 (5): 241, 1895 (
** Cheilolejeunea japonica (Horik.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (
** Cheilolejeunea kitagawae W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (
*** Cheilolejeunea lacerata C.J.Bastos et Gradst., J. Bryol. 28 (2): 133, 2006 (
** Cheilolejeunea larsenii Mizut., Dansk Bot. Ark. 27 (1): 95, 1969 (
* Cheilolejeunea laurentii Steph., Sp. Hepat. (Stephani) 5: 647, 1914 (
** Cheilolejeunea leptophylla (Ångstr.) Steph., Sp. Hepat. (Stephani) 5: 657, 1914 (
* Cheilolejeunea longiflora (Taylor) R.M.Schust., Phytologia 45 (5): 431, 1980 (
** Cheilolejeunea longispina (Herzog) R.M.Schust., Beih. Nova Hedwigia 9: 111, 1963 (
** Cheilolejeunea loriana (Steph.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 280, 2010 (
** Cheilolejeunea ludoviciae Steph., Sp. Hepat. (Stephani) 5: 668, 1914 (
** Cheilolejeunea lurida (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 658, 1914 (
** Cheilolejeunea macroloba (Herzog) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982 (
*** Cheilolejeunea malaccensis (G.Hoffm.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 59, 1996 (
** Cheilolejeunea mammifera R.M.Schust., Phytologia 45 (5): 429, 1980 (
*** Cheilolejeunea mariana (Gottsche) B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 75, 1989 (
** Cheilolejeunea mexicana Steph., Sp. Hepat. (Stephani) 6: 417, 1923 (
* Cheilolejeunea micholitzii (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea microscypha (Hook.f. et Taylor) M.Wigginton, J. Bryol. 34 (4): 269, 2012 (
*** Cheilolejeunea mizutanii W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010 (
*** Cheilolejeunea neblinensis Ilk.-Borg. et Gradst., Nova Hedwigia 87 (3/4): 522, 2008 (
** Cheilolejeunea nietneri (Steph.) Mizut., J. Hattori Bot. Lab. 37: 191, 1973 (
*** Cheilolejeunea norisiae G.Dauphin et Gradst., J. Bryol. 25 (4): 259, 2003 (
** Cheilolejeunea novaezelandiae R.M.Schust., Phytologia 56 (7): 459, 1985 (
*** Cheilolejeunea obcordata Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 209, 1931 (
** Cheilolejeunea obtruncata (Mont.) Solari, J. Hattori Bot. Lab. 54: 539, 1983 (
** Cheilolejeunea obtusa (Herzog) Solari, J. Hattori Bot. Lab. 54: 537, 1983 (
** Cheilolejeunea obtusifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 18: 116, 1957 (
** Cheilolejeunea occlusa (Herzog) T.Kodama et N.Kitag., Bull. Osaka Mus. Nat. Hist. 28: 40, 1974 (
** Cheilolejeunea orientalis (Gottsche) Mizut., J. Hattori Bot. Lab. 35: 399, 1972 (
** Cheilolejeunea ovistipula Steph., Hedwigia 34 (5): 244, 1895 (
** Cheilolejeunea panurensis (Spruce) Steph., Sp. Hepat. (Stephani) 5: 660, 1914 (
** Cheilolejeunea papillata Solari, Comun. Mus. Argent. Ci. Nat. “Bernardino Rivadavia,” Ci. Bot. 2 (11): 72, 1981 (
** Cheilolejeunea paroica Mizut., J. Hattori Bot. Lab. 46: 364, 1979 (
** Cheilolejeunea piriflora Schiffn., Bot. Jahrb. Syst. 23 (5): 592, 1897 (
*** Cheilolejeunea polystachya (Spruce) Gradst. et Ilk.-Borg., Mem. New York Bot. Gard. 76 (4): 62, 2009 (
** Cheilolejeunea renneri (G.Hoffm.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 62, 1996 (
*** Cheilolejeunea rotundistipula (Lindenb. ex Lehm.) Malombe, Acta Bot. Hung. 51 (3/4): 322, 2009 (
** Cheilolejeunea rufescens (Lindenb.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 212, 1982 (
** Cheilolejeunea stenoschiza (Ångstr.) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 436, 1900 (
** Cheilolejeunea subcrenulata (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980 (
* Cheilolejeunea suborbicularis (Herzog) H.A.Mill., Bonner et Bischl., Nova Hedwigia 4: 559, 1962 [1963] (
*** Cheilolejeunea suprema (Grolle et Piippo) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010 (
** Cheilolejeunea surrepens (Mitt.) E.W.Jones, J. Bryol. 9 (1): 49, 1976 (
*** Cheilolejeunea turgida (Mitt.) W.Ye et R.L.Zhu, J. Bryol. 32 (4): 281, 2010 (
* Cheilolejeunea ulugurica Malombe, Eb.Fisch. et Pócs, Acta Biol. Pl. Agr. 1: 24, 2010 [2011] (
*** Cheilolejeunea unciloba (Lindenb.) Malombe, Acta Bot. Hung. 51 (3/4): 325, 2009 (
** Cheilolejeunea upoluensis S.W.Arnell, Svensk Bot. Tidskr. 50 (3): 516, 1956 (
** Cheilolejeunea valenciae (Steph.) Xiao L.He, Ann. Bot. Fenn. 33 (1): 55, 1996 (
** Cheilolejeunea virescens (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 662, 1914 (
* Cheilolejeunea viridis Steph., Sp. Hepat. (Stephani) 5: 673, 1914 (
** Cheilolejeunea warnstorffii (Steph.) Solari, Darwiniana 20 (3/4): 387, 1976 (
* Cheilolejeunea wrightii Steph., Sp. Hepat. (Stephani) 5: 662, 1914 (
*** Cheilolejeunea xanthocarpa (Lehm. et Lindenb.) Malombe, Acta Bot. Hung. 51 (3/4): 326, 2009 (
** Cheilolejeunea xanthophylla (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 663, 1914 (
** subg. Euosmolejeunea (Spruce) Kachroo, Ceylon J. Sci., Biol. Sci. 8 (1): 6, 1968 (
** Cheilolejeunea aciculifera R.M.Schust., Phytologia 45 (5): 428, 1980 (
** Cheilolejeunea acutangula (Nees) Grolle, J. Hattori Bot. Lab. 45: 173, 1979 (
*** Cheilolejeunea asperrima (Steph.) Grolle, J. Hattori Bot. Lab. 58: 197, 1985 (
** Cheilolejeunea birmensis (Steph.) Mizut., J. Hattori Bot. Lab. 27: 139, 1964 (
** Cheilolejeunea campbelliensis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963 (
** Cheilolejeunea cedercreutzii (H.Buch et Perss.) Grolle, Feddes Repert. 87 (3/4): 188, 1976 (
** Cheilolejeunea choachina (Gottsche) Gradst., Mem. New York Bot. Gard. 84: 69, 1999 (
*** Cheilolejeunea clausa (Nees et Mont.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 863, 1980 (
** Cheilolejeunea clavata Mizut., J. Hattori Bot. Lab. 46: 361, 1979 (
*** Cheilolejeunea comans (Spruce) R.M.Schust., Phytologia 45 (5): 431, 1980 (
** Cheilolejeunea convexa (S.W.Arnell) S.W.Arnell, Hepat. South Africa: 209, 1963 (
** Cheilolejeunea cookiensis (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea cordistipula (Steph.) Grolle ex E.W.Jones, J. Bryol. 13 (3): 395, 1985 (
** Cheilolejeunea fischeri Malombe, Acta Bot. Hung. 51 (3/4): 319, 2009 (
** Cheilolejeunea gaoi R.L.Zhu, M.L.So et Grolle, Bryologist 103 (3): 499, 2000 (
** Cheilolejeunea hallingii B.M.Thiers, Brittonia 44 (2): 160, 1992 (
*** Cheilolejeunea inflexa (Hampe) Grolle, J. Hattori Bot. Lab. 45: 174, 1979 (
*** Cheilolejeunea intertexta (Lindenb.) Steph., Bull. Herb. Boissier 5 (2): 79, 1897 (
*** Cheilolejeunea krakakammae (Lindenb.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (
*** Cheilolejeunea laciniata D.F.Peralta et M.E.Reiner, Bryologist 116 (1): 54, 2013 (
*** Cheilolejeunea laevicalyx (J.B.Jack et Steph.) Grolle, J. Hattori Bot. Lab. 65: 403, 1988 (
** Cheilolejeunea laeviuscula (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 668, 1914 (
*** Cheilolejeunea lindenbergii (Gottsche) Mizut., J. Hattori Bot. Lab. 33: 226, 1970 (
** Cheilolejeunea mimosa (Hook.f. et Taylor) R.M.Schust., J. Hattori Bot. Lab. 26: 245, 1963 (
** Cheilolejeunea nana R.M.Schust., Phytologia 39 (6): 426, 1978 (
** Cheilolejeunea ngongensis Malombe et Pócs, Acta Bot. Hung. 51 (3/4): 317, 2009 (
** Cheilolejeunea nipponica (S.Hatt.) S.Hatt., Misc. Bryol. Lichenol. 1 (14): 1, 1957 (
*** Cheilolejeunea omphalogastria Pócs, Trop. Bryol. 9: 131, 1994 (
*** Cheilolejeunea oncophylla (Ångstr.) Grolle et M.E.Reiner, J. Bryol. 19 (4): 781, 1997 (
** Cheilolejeunea ornata C.J.Bastos, J. Bryol. 33 (1): 86, 2011 (
** Cheilolejeunea osumiensis (S.Hatt.) Mizut., Misc. Bryol. Lichenol. 8 (7): 148, 1980 (
** Cheilolejeunea pluriplicata (Pearson) R.M.Schust., Phytologia 45 (5): 430, 1980 (
** Cheilolejeunea pocsii E.W.Jones, J. Bryol. 15 (1): 156, 1988 (
** Cheilolejeunea polyantha A.Evans, Mem. Torrey Bot. Club 8 (2): 141, 1902 (
** Cheilolejeunea polyantha var. caduciloba R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 887, 1980 (
*** Cheilolejeunea revoluta (Herzog) Gradst. et Grolle, J. Hattori Bot. Lab. 74: 59, 1993 (
*** Cheilolejeunea rigidula (Nees ex Mont.) R.M.Schust., Castanea 36 (2): 102, 1971 (
** Cheilolejeunea ruwenzorensis (S.W.Arnell) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (
** Cheilolejeunea ryukyuensis Mizut., J. Hattori Bot. Lab. 51: 162, 1982 (
** Cheilolejeunea subopaca (Mitt.) Mizut., J. Hattori Bot. Lab. 26: 183, 1963 (
*** Cheilolejeunea trifaria (Reinw., Blume et Nees) Mizut., J. Hattori Bot. Lab. 27: 132, 1964 (
* Cheilolejeunea udarii G.Asthana, S.C.Srivast. et A.K.Asthana, Lindbergia 20 (2/3): 142, 1995 [1996] (
** Cheilolejeunea usambarana (Steph.) Grolle, J. Hattori Bot. Lab. 46: 344, 1979 (
** Cheilolejeunea verrucosa Steph., Sp. Hepat. (Stephani) 5: 673, 1914 (
** subg. Renilejeunea R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (
*** Cheilolejeunea montagnei (Gottsche ex Mont.) R.M.Schust., Beih. Nova Hedwigia 9: 112, 1963 (
** subg. Xenolejeunea Kachroo et R.M.Schust., J. Linn. Soc., Bot. 56 (368): 496, 1961 (
** Cheilolejeunea ceylanica (Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea eximia (Jovet-Ast et Tixier) R.L.Zhu et M.L.So, Beih. Nova Hedwigia 121: 114, 2001 (
*** Cheilolejeunea falsinervis (Sande Lac.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea gardneri (Mitt.) Mizut., J. Hattori Bot. Lab. 26: 181, 1963 (
** Cheilolejeunea gigantea (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea huerlimannii Tixier, Misc. Bryol. Lichenol. 9 (9): 184, 1983 (
** Cheilolejeunea hyalomarginata R.L.Zhu et Frank Müll., Bryologist 115 (2): 218, 2012 (
*** Cheilolejeunea incisa (Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea incisa var. teretiflora B.M.Thiers, J. Hattori Bot. Lab. 82: 321, 1997 (
** Cheilolejeunea longidens (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea meyeniana (Nees, Lindenb. et Gottsche) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea obtusilobula (S.Hatt.) S.Hatt., Misc. Bryol. Lichenol. 1 (14): 2, 1957 (
** Cheilolejeunea parvidens B.M.Thiers, J. Hattori Bot. Lab. 82: 323, 1997 (
** Cheilolejeunea streimannii Pócs et Ninh, Acta Bot. Hung. 47 (1/2): 162, 2005 (
*** Cheilolejeunea trapezia (Nees) Kachroo et R.M.Schust., J. Linn. Soc., Bot. 56 (368): 509, 1961 (
** Cheilolejeunea ventricosa (Schiffn. ex P.Syd.) Xiao L.He, Acta Bot. Fenn. 163: 60, 1999 (
*** Cheilolejeunea vittata (Steph. ex G.Hoffm.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 509, 1961 (
Incertae sedis
** Cheilolejeunea gottscheana C.J.Bastos, J. Bryol. 34 (4): 316, 2012 (
* Cheilolejeunea heteroclada (Spruce) Schiffn., Hepat. (Engl.-Prantl): 124, 1893 (
** Cheilolejeunea minutilobula Amakawa, J. Jap. Bot. 35 (12): 365, 1960 (
** Cheilolejeunea norrisii (Grolle) M.A.M.Renner, Bryologist 115 (4): 550, 2012 (
** Cheilolejeunea oscilla M.A.M.Renner, Bryologist 115 (4): 551, 2012 (
** Cheilolejeunea papulosa Schiffn., Hepat. (Engl.-Prantl): 124, 1893 (
*** Cheilolejeunea riparia (Steph.) M.E.Reiner, Nova Hedwigia 95 (3/4): 467, 2012 (
*** Cheilolejeunea rotalis (Hook.f. et Taylor) M.Wigginton, J. Bryol. 34 (4): 270, 2012 (
** Cheilolejeunea tenerrima (Steph.) C.J.Bastos, J. Bryol. 34 (4): 317, 2012 (
** Cheilolejeunea urubuensis (Zartman et I.L.Ackerman) R.L.Zhu et Y.M.Wei, Phytotaxa 152 (1): 50, 2013 (
* Cyrtolejeunea A.Evans, Bull. Torrey Bot. Club 30 (10): 553, 1903 (
*** Cyrtolejeunea holostipa (Spruce) A.Evans, Bull. Torrey Bot. Club. 30 (10): 553, 1903 (
** Cystolejeunea A.Evans, Bull. Torrey Bot. Club 33 (1): 16, 1906) (
*** Cystolejeunea lineata (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club. 33 (1): 17, 1906 (
* Omphalanthus Lindenb. et Nees, Syn. Hepat. 2: 303, 1845 (
** Omphalanthus baracoensis Mustelier, M.E.Reiner et Gradst., J. Bryol. 29 (2): 95, 2007 (
*** Omphalanthus filiformis (Sw.) Nees, Syn. Hepat. 2: 304, 1845 (
** Omphalanthus filiformis var. platycoleus (Herzog) Gradst., Phytotaxa 76 (3): 46, 2013 (
** Omphalanthus filiformis var. wallisii (Prantl) Gradst., Phytotaxa 76 (3): 46, 2013 (
** Omphalanthus huanucensis (Gottsche) Gradst., Beih. Nova Hedwigia 80: 109, 1985 (
*** Omphalanthus jackii (Prantl) Gradst., Proc. Kon. Ned. Akad. Wetensch. C 80: 410, 1977 (
*** Omphalanthus ovalis (Lindenb. et Gottsche) Gradst., Proc. Kon. Ned. Akad. Wetensch. C 80: 411, 1977 (
*** Omphalanthus roccatii (Gola) R.M.Schust., Beih. Nova Hedwigia 9: 96, 1963 (
** Aphanotropis Herzog, Trans. Brit. Bryol. Soc. 2 (1): 63, 1952 (
*** Aphanotropis saxicola Herzog, Trans. Brit. Bryol. Soc. 2 (1): 63, 1952 (
** Calatholejeunea K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 8, 1928 (
*** Calatholejeunea lamii Mizut., J. Hattori Bot. Lab. 56: 334, 1984 (
*** Calatholejeunea paradoxa (Schiffn.) K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 8, 1928 (
*** Cololejeunea (Spruce) Steph., Hedwigia 30 (5): 208, 1891 (
** Cololejeunea micrandroecia (Spruce) M.Menzel, Willdenowia 14: 492, 1984 [1985] (
* subg. Aphanolejeunea (A.Evans) Pócs, Phytotaxa 202 (1): 64, 2015 (
*** Cololejeunea berneckerae Pócs, Polish Bot. J. 54 (1): 4, 2009 (
*** Cololejeunea cingens (Herzog) Bernecker et Pócs, Polish Bot. J. 54 (1): 4, 2009 (
*** Cololejeunea cornutissima (R.M.Schust.) Stotler et Crand.-Stotl., Bryologist 80 (3): 411, 1977 (
*** Cololejeunea costaricensis (Bern.-Lück.) Bernecker et Pócs, Polish Bot. J. 54 (1): 5, 2009 (
*** Cololejeunea cubensis Pócs, Polish Bot. J. 54 (1): 5, 2009 (
*** Cololejeunea gracilis (Jovet-Ast) Pócs, Cryptog. Bryol. 29 (3): 233, 2008 (
** Cololejeunea gracilis var. linearifolia (R.M.Schust.) Pócs, Acta Bot. Hung. 56 (1/2): 189, 2014 (
*** Cololejeunea grossepapillosa (Horik.) N.Kitag., Hikobia, Suppl. 1: 68, 1981 (
** Cololejeunea iwatsukiana (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009 (
*** Cololejeunea jovetastiana (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009 (
** Cololejeunea lisowskii (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009 (
*** Cololejeunea madeirensis Schiffn., Hedwigia 41 (5): 279, 1902 (
*** Cololejeunea microscopica (Taylor) Schiffn., Hepat. (Engl.-Prantl): 122, 1893 (
*** Cololejeunea microscopica var. africana (Pócs) Pócs et Bernecker, Cryptog. Bryol. 29 (3): 234, 2008 (
*** Cololejeunea microscopica var. exigua (A.Evans) Pócs, Mem. New York Bot. Gard. 76 (4): 73, 2009 (
*** Cololejeunea minuscula Pócs, Polish Bot. J. 54 (1): 7, 2009 (
** Cololejeunea moramangae Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 241, 1977 [1979] (
*** Cololejeunea norrisii (Pócs) Pócs, Polish Bot. J. 54 (1): 7, 2009 (
*** Cololejeunea papillosa (K.I.Goebel) Mizut., J. Hattori Bot. Lab. 29: 156, 1966 (
*** Cololejeunea sicifolia (Gottsche ex A.Evans) Pócs et Bernecker, Polish Bot. J. 54 (1): 8, 2009 (
*** Cololejeunea sicifolia subsp. jamaicensis (R.M.Schust.) Bernecker et Pócs, Polish Bot. J. 54 (1): 8, 2009 (
*** Cololejeunea sintenisii (Steph.) Pócs, Cryptog. Bryol. 29 (3): 235, 2008 (
*** Cololejeunea subsphaeroidea (R.M.Schust.) Pócs, Cryptog. Bryol. 29 (3): 235, 2008 (
** Cololejeunea thiersiae (Pócs) Pócs, Polish Bot. J. 54 (1): 9, 2009 (
** Cololejeunea veillonii Tixier, Nova Hedwigia 31: 757, 1979 (
*** Cololejeunea winkleri (M.I.Morales et A.Lücking) Pócs, Mem. New York Bot. Gard. 76 (4): 78, 2009 (
* subg. Austrocololejeunea Tixier, Nova Hedwigia 31: 776, 1979 (
** Cololejeunea australis Tixier, Nova Hedwigia 31: 781, 1979 (
** Cololejeunea caledonica Gottsche, Hedwigia 34 (5): 246, 1895 (
** Cololejeunea sophiana Tixier, Bot. Not. 128: 428, 1975 [1976] (
** Cololejeunea virotana Tixier, Nova Hedwigia 31: 777, 1979 (
* subg. Chlorocolea R.M.Schust., Beih. Nova Hedwigia 9: 178, 1963 (
*** Cololejeunea ceratilobula (P.C.Chen) R.M.Schust., Beih. Nova Hedwigia 9: 179, 1963 (
*** Cololejeunea desciscens Steph., Hedwigia 34 (5): 248, 1895 (
*** Cololejeunea linopteroides H.Rob., Bryologist 67 (4): 457, 1964 (
* Cololejeunea rotundilobula (P.C.Wu et P.J.Lin) Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (
*** Cololejeunea sigmoidea Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 31 (1/2): 27, 1962 (
** Cololejeunea sigmoidea var. dubia Tixier, Bryophyt. Biblioth. 27: 114, 1985 (
*** Cololejeunea standleyi Herzog, Rev. Bryol. Lichénol. 20 (1/2): 172, 1951 [1952] (
*** Cololejeunea stylilobula Tixier, Phytotaxa 202 (1): 65, 2015 (
*** Cololejeunea zangii R.L.Zhu et M.L.So, Syst. Bot. 24 (4): 501, 1999 [2000] (
*** subg. Chlorolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 81, 1953 (
** Cololejeunea lacinulata Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 82, 1953 (
*** Cololejeunea madothecoides (Steph.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 81, 1953 (
** Cololejeunea ombrophila Tixier, Nat. Hist. Bull. Siam Soc. 23 (4): 550, 1970 (
*** Cololejeunea stotleriana Gradst., Ilk.-Borg. et Vanderp., Bryologist 114 (1): 13, 2011 (
*** subg. Chondriolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 75, 1953 (
*** Cololejeunea chinii Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 445, 1973 (
*** Cololejeunea pseudostipulata Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (
*** Cololejeunea shimizui N.Kitag., Acta Phytotax. Geobot. 23 (5/6): 185, 1969 (
*** Cololejeunea shimizui var. phangngana N.Kitag., Acta Phytotax. Geobot. 23 (5/6): 187, 1969 (
*** subg. Cololejeunea,
** Cololejeunea albodentata P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 252, 1964 (
** Cololejeunea armata Tixier, Gard. Bull. Singapore 26 (1): 149, 1972 (
*** Cololejeunea bhutanica Grolle et Mizut., J. Bryol. 15 (2): 281, 1989 (
** Cololejeunea biddlecomiae (Austin) A.Evans, Mem. Torrey Bot. Club 8 (2): 168, 1902 (
* Cololejeunea caihuaella But et P.C.Wu, Hepat. Fl. Hong Kong: 132, 2009 (
*** Cololejeunea calcarea (Lib.) Steph., Bot. Gaz. 17 (6): 171, 1892 (
** Cololejeunea capuronii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 241, 1977 [1979] (
** Cololejeunea dinghuiana R.L.Zhu et Y.F.Wang, J. E. China Norm. Univ., Nat. Sci. Ed. 2: 91, 1992 (
** Cololejeunea dolichodonta Tixier, Bryophyt. Biblioth. 27: 215, 1985 (
*** Cololejeunea dozyana (Sande Lac.) Schiffn., Hedwigia 39 (4): 199, 1900 (
*** Cololejeunea elegans Steph., Hedwigia 30 (5): 208, 1891 (
** Cololejeunea falcidentata R.M.Schust., Nova Hedwigia 15: 507, 1968 (
** Cololejeunea filidens Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 49, 1953 (
* Cololejeunea frahmii Tixier, Trop. Bryol. 11: 63, 1995 (
*** Cololejeunea grolleana Pócs, J. Hattori Bot. Lab. 48: 312, 1980 (
*** Cololejeunea haskarliana (Lehm.) Schiffn., Consp. Hepat. Arch. Ind.: 244, 1898 (
*** Cololejeunea hyalina G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 25, 2003 (
* Cololejeunea kahuziensis Tixier, Trop. Bryol. 11: 63, 1995 (
*** Cololejeunea karnatakensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 26, 2003 (
** Cololejeunea kodamae Kamim., Feddes Repert. Spec. Nov. Regni Veg. 58: 55, 1955 (
*** Cololejeunea kolombangarae Pócs, Acta Bryolichenol. Asiat. 4: 67, 2011 (
** Cololejeunea kolombangarae subsp. sepikensis Pócs, Acta Bryolichenol. Asiat. 4: 68, 2011 (
*** Cololejeunea konratii Pócs, Acta Bot. Hung. 54 (1/2): 156, 2012 (
*** Cololejeunea kuciana Pócs et Schäf.-Verw., Polish Bot. J. 57 (1): 51, 2012 (
*** Cololejeunea longiana Grolle et Mizut., J. Bryol. 15 (2): 284, 1989 (
*** Cololejeunea macounii (Spruce) A.Evans, Mem. Torrey Bot. Club 8 (2): 171, 1902 (
** Cololejeunea magillii Pócs, J. Hattori Bot. Lab. 74: 49, 1993 (
*** Cololejeunea malanjae Steph., Bot. Jahrb. Syst. 30 (2): 261, 1901 (
** Cololejeunea mamillata (Ångstr.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 184, 1967 (
** Cololejeunea mizutaniana Udar et G.Srivast., Misc. Bryol. Lichenol. 9 (7): 138, 1983 (
*** Cololejeunea mocambiquensis S.W.Arnell, Mitt. Thüring. Bot. Ges. 1 (1): 7, 1955 [1956] (
** Cololejeunea nanhutashanensis J.D.Yang et S.H.Lin, Phytotaxa 177 (1): 56, 2014 (
*** Cololejeunea nilgiriensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 27, 2003 (
*** Cololejeunea ornata A.Evans, Bryologist 41 (4): 73, 1938 (
** Cololejeunea parva Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 239, 1977 (
*** Cololejeunea planiflora Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 60, 1953 (
*** Cololejeunea platyneura (Spruce) A.Evans, Mem. Torrey Bot. Club 8 (2): 172, 1902 (
*** Cololejeunea pluridentata P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 105, 1978 (
** Cololejeunea pretiosa Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 62, 1953 (
*** Cololejeunea pseudocristallina P.C.Chen et P.C.Wu, Acta Phytotax. Sin. 9 (3): 257, 1964 (
** Cololejeunea pseudoplagiophylla P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 106, 1978 (
** Cololejeunea pseudoschmidtii Tixier, Gard. Bull. Singapore 26 (1): 145, 1972 (
** Cololejeunea ramromensis Pócs, Biodiv., biogeogr., nat. conserv. Wallacea N. Guinea II: 113, 2014 (
*** Cololejeunea rosellata Mizut., J. Hattori Bot. Lab. 29: 159, 1966 (
*** Cololejeunea rossettiana (C.Massal.) Schiffn., Hepat. (Engl.-Prantl): 122, 1893 (
*** Cololejeunea runssorensis (Steph.) Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 371, 1975 (
** Cololejeunea schaeferi Grolle, J. Bryol. 13 (4): 488, 1986 (
*** Cololejeunea schmidtii Steph., Bot. Tidsskr. 24 (3): 278, 1902 (
** Cololejeunea schmidtii var. acutepapillosa Pócs, Acta Bot. Hung. 54 (1/2): 165, 2012 (
** Cololejeunea selaginellicola Tixier, Gard. Bull. Singapore 26 (1): 149, 1972 (
*** Cololejeunea serrata (Steph.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 53, 1953 (
** Cololejeunea shikokiana (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 101, 1944 (
** Cololejeunea spinosa (Horik.) Pandé et R.N.Misra, J. Indian Bot. Soc. 22 (2/4): 166, 1943 (
*** Cololejeunea stellaris Pócs, Acta Bryolichenol. Asiat. 4: 70, 2011 (
** Cololejeunea subkodamae Mizut., J. Hattori Bot. Lab. 60: 448, 1986 (
** Cololejeunea tamasii Schäf.-Verw., Phytotaxa 60: 9, 2012 (
** Cololejeunea tanneri Pócs, Acta Bot. Hung. 31 (1/4): 126, 1985 (
*** Cololejeunea tenella Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 55, 1953 (
** Cololejeunea tenella var. dentiloba Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 436, 1989 (
** Cololejeunea thiersiana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (
** Cololejeunea tuiwawana Pócs, Acta Bot. Hung. 54 (1/2): 165, 2012 (
*** Cololejeunea verrucosa Steph., Hedwigia 34 (5): 253, 1895 (
** Cololejeunea verrucosa var. rectispina (Herzog) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 55, 1953 (
*** Cololejeunea zenkeri (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 420, 1954 (
* subg. Cryptolejeunea Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 77, 1953 (
*** Cololejeunea angustiflora (Steph.) Mizut., J. Hattori Bot. Lab. 28: 113, 1965 (
** Cololejeunea drepanolejeuneoides (Horik.) R.M.Schust., Beih. Nova Hedwigia 9: 174, 1963 (
** Cololejeunea hattoriana Mizut. et Pócs, Ann. Bot. Fenn. 31 (3): 188, 1994 (
*** Cololejeunea inflectens (Mitt.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 79, 1953 (
** Cololejeunea mouensis (Tixier) H.A.Mill., Phytologia 47 (4): 322, 1981 (
*** Cololejeunea vesicaria (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 247, 1898 (
* subg. Diaphanae R.M.Schust., J. Elisha Mitchell Sci. Soc. 72 (1): 103, 1956 (
*** Cololejeunea amphibola B.M.Thiers, Beih. Nova Hedwigia 90: 130, 1988 (
** Cololejeunea antillana Pócs, Polish Bot. J. 54 (1): 3, 2009 (
* Cololejeunea augieri Tixier, Trop. Bryol. 11: 48, 1995 (
*** Cololejeunea azorica V.Allorge et Jovet-Ast, Mitt. Thüring. Bot. Ges. 1 (2/3): 17, 1955 (
*** Cololejeunea camillii (Lehm.) A.Evans, Bryologist 15 (4): 59, 1912 (
*** Cololejeunea contractiloba A.Evans, Amer. J. Bot. 5 (3): 131, 1918 (
*** Cololejeunea cornuta E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 436, 1954 (
** Cololejeunea crenata (A.Evans) Pócs, Polish Bot. J. 54 (1): 5, 2009 (
*** Cololejeunea diaphana A.Evans, Bull. Torrey Bot. Club 32 (4): 184, 1905 (
*** Cololejeunea erostrata (Herzog) Bernecker et Pócs, Polish Bot. J. 54 (1): 5, 2009 (
** Cololejeunea gradsteinii M.J.Lai et R.L.Zhu, Ann. Bot. Fenn. 45 (5): 334, 2008 (
** Cololejeunea koponenii (Pócs) Pócs, Polish Bot. J. 54 (1): 6, 2009 (
*** Cololejeunea lanceolata E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 428, 1954 (
*** Cololejeunea moralesiae (Bern.-Lück.) Bernecker et Pócs, Polish Bot. J. 54 (1): 7, 2009 (
*** Cololejeunea morobensis (Pócs) Pócs, Polish Bot. J. 54 (1): 7, 2009 (
*** Cololejeunea obtusifolia (E.W.Jones) Tixier, Trop. Bryol. 11: 42, 1995 (
* Cololejeunea obtusifolia var. madecassa (Tixier) Pócs, Phytotaxa 202 (1): 64, 2015 (
*** Cololejeunea papilliloba (Steph.) Steph., Hedwigia 34 (5): 250, 1895 (
** Cololejeunea papulosa R.M.Schust., Phytologia 45 (5): 433, 1980 (
*** Cololejeunea paucifolia (Spruce) Bernecker et Pócs, Polish Bot. J. 54 (1): 8, 2009 (
*** Cololejeunea peponiformis Mizut., J. Hattori Bot. Lab. 33: 258, 1970 (
** Cololejeunea pseudocuspidata Tixier, Bauhinia 8 (4): 228, 1987 (
*** Cololejeunea pterocolea Herzog, Svensk Bot. Tidskr. 46 (1): 103, 1952 (
*** Cololejeunea pusilla Steph., Hedwigia 34 (5): 251, 1895 (
** Cololejeunea spathulata Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 36, 1960 (
** Cololejeunea taurifolia Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 152, 1965 (
*** Cololejeunea wightii Steph., Hedwigia 34 (5): 253, 1895 (
*** Cololejeunea yelitzae Pócs et Bernecker, Acta Bot. Hung. 55 (3/4): 386, 2013 (
* subg. Leptocolea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 122, 1893 (
** Cololejeunea acuminata Mizut., J. Hattori Bot. Lab. 33: 261, 1970 (
*** Cololejeunea aequabilis (Sande Lac.) Schiffn., Consp. Hepat. Arch. Ind.: 242, 1898 (
** Cololejeunea altimontana Pócs, Acta Bryolichenol. Asiat. 4: 78, 2011 (
*** Cololejeunea amaniensis Pócs, Acta Bot. Hung. 31 (1/4): 120, 1985 (
*** Cololejeunea angustibracteata Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (
*** Cololejeunea apiculata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 174, 1963 (
* Cololejeunea arrectifolia (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (
** Cololejeunea aurantia (Tixier) Thouvenot, Cryptog. Bryol. 32 (4): 290, 2011 (
** Cololejeunea bandamiae Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 220, 1977 [1979] (
** Cololejeunea bebourensis Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 229, 1977 [1979] (
** Cololejeunea bergmansiana Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 440, 1989 (
** Cololejeunea bidentula (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 423, 1954 (
** Cololejeunea bifalcata Pócs, Acta Bot. Hung. 54 (1/2): 146, 2012 (
*** Cololejeunea blepharophylla Pócs, Acta Bot. Hung. 54 (1/2): 149, 2012 (
** Cololejeunea borhidiana Pócs, J. Hattori Bot. Lab. 48: 305, 1980 (
** Cololejeunea bosseriana Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 235, 1977 [1979] (
** Cololejeunea calcarata E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 235, 1983 (
* Cololejeunea camusii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 227, 1977 [1979] (
*** Cololejeunea ceatocarpa (Ångstr.) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (
*** Cololejeunea ceylanica Onr., Acta Bot. Acad. Sci. Hung. 25 (1/2): 107, 1979 (
* Cololejeunea chamlongiana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 181, 1970 (
*** Cololejeunea chenii Tixier, Bryophyt. Biblioth. 27: 219, 1985 (
*** Cololejeunea ciliata Pócs, Ann. Bot. Fenn. 31 (3): 182, 1994 (
** Cololejeunea comptonii (Pearson) H.A.Mill., Phytologia 47 (4): 321, 1981 (
** Cololejeunea cookei A.Evans, Trans. Connecticut Acad. Arts 10 (8): 447, 1900 (
** Cololejeunea cordiflora Steph., Hedwigia 34 (5): 246, 1895 (
** Cololejeunea crateris Pócs, Acta Bot. Hung. 54 (1/2): 153, 2012 (
* Cololejeunea crenulata (Pearson) H.A.Mill., Phytologia 47 (4): 321, 1981 (
** Cololejeunea cucullifolia (Herzog) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 127, 1962 (
** Cololejeunea cuneifolia Steph., Hedwigia 31 (4): 166, 1892 (
* Cololejeunea dadeuniana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 186, 1970 (
** Cololejeunea decemplicata (Steph.) Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 213, 1977 [1979] (
** Cololejeunea decliviloba Steph., Hedwigia 34 (5): 247, 1895 (
** Cololejeunea dentata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 175, 1963 (
** Cololejeunea denticulata (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 99, 1944 (
* Cololejeunea dentilobula (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963 (
*** Cololejeunea dianae M.Wigginton, J. Bryol. 28 (4): 364, 2006 (
** Cololejeunea dilatata (Steph.) Mizut., J. Hattori Bot. Lab. 28: 113, 1965 (
*** Cololejeunea diplasiolejeunoides Tixier, Bryologist 82 (4): 602, 1979 (
*** Cololejeunea distalopapillata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963 (
* Cololejeunea effusa (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (
*** Cololejeunea elephantorum Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 185, 1970 (
** Cololejeunea ellipsoidea R.M.Schust., Nova Hedwigia 15: 508, 1968 (
*** Cololejeunea equialbi Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 178, 1970 (
*** Cololejeunea filicis (Herzog) Piippo, J. Hattori Bot. Lab. 68: 133, 1990 (
* Cololejeunea fischeri Tixier, Trop. Bryol. 11: 59, 1995 (
** Cololejeunea flavicans (Steph.) Mizut., J. Hattori Bot. Lab. 28: 115, 1965 (
* Cololejeunea fredericii Onr., Rev. Bryol. Lichénol. 44 (1): 80, 1978 (
** Cololejeunea fusca (Steph.) Mizut., J. Hattori Bot. Lab. 28: 120, 1965 (
** Cololejeunea gottschei (Steph.) Pandé, K.P.Srivast. et Ahmad, J. Indian Bot. Soc. 36 (3): 345, 1957 (
*** Cololejeunea grossestyla M.Wigginton, J. Bryol. 28 (4): 369, 2006 (
*** Cololejeunea grushvitzkiana Pócs, Bot. Zhurn. (Moscow & Leningrad) 56 (5): 674, 1971 (
*** Cololejeunea hainanensis R.L.Zhu, J. Hattori Bot. Lab. 78: 87, 1995 (
*** Cololejeunea harrisii Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 357, 1975 (
*** Cololejeunea hildebrandii (Austin) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (
*** Cololejeunea hirta Steph., Bull. Misc. Inform. Kew 1899 (151/152): 125, 1899 (
** Cololejeunea hodgsoniae (Herzog) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (11): 184, 1967 (
*** Cololejeunea horikawana (S.Hatt.) Mizut., J. Hattori Bot. Lab. 24: 254, 1961 (
*** Cololejeunea huerlimannii Tixier, Nova Hedwigia 31: 773, 1979 (
** Cololejeunea inflexifolia R.M.Schust., Phytologia 56 (7): 458, 1985 (
*** Cololejeunea iradieri M.Infante et Heras, Trop. Bryol. 17: 14, 1999 (
** Cololejeunea irianensis Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 440, 1989 (
*** Cololejeunea johannis-winkleri (Herzog) R.L.Zhu, Nova Hedwigia 79 (3/4): 528, 2004 (
** Cololejeunea kegelii Steph., Hedwigia 34 (5): 249, 1895 (
** Cololejeunea khanii Tixier, Dacca Univ. Stud., B 15: 9, 1967 (
* Cololejeunea kohkongensis Tixier, Bryophyt. Biblioth. 27: 302, 1985 (
*** Cololejeunea lichenyae R.D.Porley, N.G.Hodgetts et M.Wigginton, J. Bryol. 29 (1): 7, 2007 (
* Cololejeunea lobulilineata Tixier, Trop. Bryol. 11: 42, 1995 (
*** Cololejeunea longifolia (Mitt.) Benedix ex Mizut., J. Hattori Bot. Lab. 26: 184, 1963 (
*** Cololejeunea magna (Tixier) M.Infante et Heras, Trop. Bryol. 17: 17, 1999 (
** Cololejeunea magnifica Pócs, Acta Bryolichenol. Asiat. 4: 88, 2011 (
** Cololejeunea magnilobula (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 99, 1944 (
* Cololejeunea micronesica H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 63, 1963 (
** Cololejeunea mooreaensis Tixier, Bauhinia 8 (4): 226, 1987 (
* Cololejeunea ninguana Tixier, Nova Hedwigia 31: 776, 1979 (
** Cololejeunea obcordata (Austin) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 448, 1900 (
*** Cololejeunea obliqua (Nees et Mont.) Schiffn., Bot. Jahrb. Syst. 23 (5): 586, 1897 (
* Cololejeunea oblongiperianthia (P.C.Wu et J.S.Lou) Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (
** Cololejeunea oleana Sim, Trans. Roy. Soc. South Africa 15 (1): 49, 1926 (
** Cololejeunea ovalifolia A.Evans, Trans. Connecticut Acad. Arts 10 (8): 450, 1900 (
** Cololejeunea panchoana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 183, 1970 (
*** Cololejeunea paniensis (Tixier) Grolle, J. Bryol. 8 (4): 485, 1975 (
** Cololejeunea papuliflora Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 85: 199, 1910 (
** Cololejeunea pentagona (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (
*** Cololejeunea plagiochiliana Tixier, Bot. Not. 128: 428, 1975 [1976] (
** Cololejeunea planiuscula Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (
*** Cololejeunea pseudoserrata Tixier, Nova Hedwigia 31: 770, 1979 (
* Cololejeunea pteroporum Tixier, Bryophyt. Biblioth. 27: 271, 1985 (
** Cololejeunea pulchella (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963 (
** Cololejeunea pulchella var. stylifera R.M.Schust., Phytologia 56 (7): 458, 1985 (
*** Cololejeunea quadridentata (S.Hatt.) Grolle, Acta Bot. Fenn. 125: 65, 1984 (
** Cololejeunea retusula (Mitt.) H.A.Mill., Phytologia 47 (4): 322, 1981 (
** Cololejeunea salgadoi Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 433, 1989 (
*** Cololejeunea sanctae-helenae M.Wigginton, J. Bryol. 28 (4): 366, 2006 (
** Cololejeunea serrulata Steph., Hedwigia 34 (5): 252, 1895 (
*** Cololejeunea setiloba A.Evans, Bryologist 16 (4): 51, 1913 (
*** Cololejeunea siangensis G.Asthana et S.C.Srivast., Bryophyt. Biblioth. 60: 57, 2003 (
*** Cololejeunea skottsbergii Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 749, 1942 (
*** Cololejeunea societatis Tixier, Bauhinia 8 (4): 230, 1987 (
* Cololejeunea spathulifolia (Steph.) H.A.Mill., Phytologia 47 (4): 322, 1981 (
** Cololejeunea spruceana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (
** Cololejeunea stenophylla Herzog, Bot. Not. 100 (4): 330, 1947 (
*** Cololejeunea streimannii Pócs, Acta Bryolichenol. Asiat. 4: 96, 2011 (
** Cololejeunea streimannii subsp. solomonensis Pócs, Acta Bryolichenol. Asiat. 4: 96, 2011 (
*** Cololejeunea subalpina Pócs, Acta Bryolichenol. Asiat. 4: 98, 2011 (
** Cololejeunea subcristata A.Evans, Bryologist 20 (2): 24, 1917 (
* Cololejeunea takamakae Tixier, Bryophyt. Biblioth. 27: 319, 1985 (
*** Cololejeunea tanzaniae Pócs, J. Hattori Bot. Lab. 48: 312, 1980 (
** Cololejeunea tenuiparietata Tixier, Trop. Bryol. 11: 56, 1995 (
** Cololejeunea teurnoumensis Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 183, 1970 (
*** Cololejeunea timoi Pócs, Acta Bryolichenol. Asiat. 4: 98, 2011 (
** Cololejeunea touwii Pócs, Acta Bryolichenol. Asiat. 4: 101, 2011 (
*** Cololejeunea tranninhiana Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 97, 1974 (
*** Cololejeunea trichomanis (Gottsche) Besch., Rev. Bryol. 19 (1): 14, 1892 (
* Cololejeunea tuksapiana Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 186, 1970 (
** Cololejeunea vulcania Tixier, Ann. Fac. Sci. Univ. Phnom Penh 3: 181, 1970 (
*** Cololejeunea yoshinagana (S.Hatt.) Mizut., J. Hattori Bot. Lab. 24: 250, 1961 (
* subg. Metzgeriopsis (K.I.Goebel) Pócs, Acta Bryolichenol. Asiat. 4: 106, 2011 (
*** Cololejeunea metzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilk.-Borg. et Heinrichs, Bot. J. Linn. Soc. 151 (3): 306, 2006 (
*** subg. Pedinolejeunea Benedix ex Mizut., J. Hattori Bot. Lab. 24: 240, 1961 (
** Cololejeunea abnormis Mizut., J. Hattori Bot. Lab. 33: 260, 1970 (
*** Cololejeunea adhaesiva (Mitt.) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963 (
** Cololejeunea adnata Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 194, 1977 [1979] (
*** Cololejeunea africana (Steph.) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963 (
** Cololejeunea ambeliensis Tixier, Bryophyt. Biblioth. 27: 142, 1985 (
** Cololejeunea amieuensis Tixier, Nova Hedwigia 31: 748, 1979 (
** Cololejeunea andapania Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 178, 1977 [1979] (
*** Cololejeunea angulata (Steph.) Mizut., J. Hattori Bot. Lab. 28: 108, 1965 (
*** Cololejeunea ankaiana Tixier, Bryophyt. Biblioth. 27: 62, 1985 (
** Cololejeunea attilana Pócs, Magyar Bot. Kut. Ezredf. Tanul. Borhidi: 186, 2002 (
*** Cololejeunea auriculata (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963 (
** Cololejeunea autoica (Steph.) Grolle, Bryophyt. Biblioth. 48: 43, 1995 (
*** Cololejeunea bekkeri Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (
*** Cololejeunea bischleriana Tixier, Bradea 3 (6): 36, 1980 (
** Cololejeunea bolovenensis Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 442, 1973 (
*** Cololejeunea borbonica Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 188, 1977 [1979] (
* Cololejeunea brunelii Tixier, Dacca Univ. Stud., B 15: 10, 1967 (
*** Cololejeunea cardiocarpa (Mont.) A.Evans, Mem. Torrey Bot. Club 8 (2): 172, 1902 (
* Cololejeunea chittagongensis Tixier, Bryophyt. Biblioth. 27: 97, 1985 (
*** Cololejeunea cocoscola Tixier, Cryptog. Bryol. Lichénol. 14 (3): 353, 1993 (
** Cololejeunea cremersii Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (
** Cololejeunea cristata (Steph.) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963 (
** Cololejeunea cuneata (Lehm. et Lindenb.) Herzog, Bot. Not. 100 (4): 320, 1947 (
* Cololejeunea deroinii Tixier, Cryptog. Bryol. Lichénol. 14 (3): 355, 1993 (
* Cololejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 227, 1977 (
* Cololejeunea dzumacensis Tixier, Nova Hedwigia 31: 754, 1979 (
*** Cololejeunea ecuadoriensis Pócs, Acta Bot. Hung. 44 (3/4): 372, 2002 (
* Cololejeunea epiphylla G.Asthana et A.Shukla, Cryptog. Bryol. 31 (3): 218, 2010 (
** Cololejeunea fissilobula Herzog, Trans. Brit. Bryol. Soc. 1 (4): 323, 1950 (
** Cololejeunea florencei Tixier, Cryptog. Bryol. Lichénol. 14 (3): 355, 1993 (
** Cololejeunea foliicola S.C.Srivast. et G.Srivast., Proc. Indian Acad. Sci. Pl. Sci. 99 (2): 86, 1989 (
** Cololejeunea fructumarginata Tixier, Bryophyt. Biblioth. 27: 58, 1985 (
*** Cololejeunea furcilobulata (Berrie et E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 178, 1963 (
*** Cololejeunea geissleriana Tixier, Bradea 3 (6): 37, 1980 (
** Cololejeunea georgiana Tixier, Bryophyt. Biblioth. 27: 145, 1985 (
*** Cololejeunea guadelupensis Tixier, Cryptog. Bryol. Lichénol. 16 (3): 229, 1995 (
*** Cololejeunea hebridensis Tixier, Bot. Not. 128: 425, 1975 [1976] (
*** Cololejeunea hinidumae Onr., Acta Bot. Acad. Sci. Hung. 25 (1/2): 109, 1979 (
* Cololejeunea hoabinhiana Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 91, 1974 (
** Cololejeunea hoeana Tixier, Bryophyt. Biblioth. 27: 56, 1985 (
** Cololejeunea hungii Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 91, 1974 (
*** Cololejeunea indosinica Tixier, Bryophyt. Biblioth. 27: 63, 1985 (
** Cololejeunea inoueana Mizut., J. Hattori Bot. Lab. 57: 440, 1984 (
** Cololejeunea japonica (Schiffn.) Mizut., J. Hattori Bot. Lab. 24: 241, 1961 (
*** Cololejeunea jonesii Pócs, Acta Bot. Acad. Sci. Hung. 21 (3/4): 361, 1975 (
*** Cololejeunea kapingaensis H.A.Mill., Bryologist 59 (3): 170, 1956 (
** Cololejeunea kiriromensis Tixier, Bryophyt. Biblioth. 27: 147, 1985 (
*** Cololejeunea kulenensis Tixier, Bryophyt. Biblioth. 27: 71, 1985 (
** Cololejeunea laevigata (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 241, 1963 (
*** Cololejeunea lanciloba Steph., Hedwigia 34 (5): 250, 1895 (
*** Cololejeunea latilobula (Herzog) Tixier, Bryophyt. Biblioth. 27: 156, 1985 (
*** Cololejeunea latistyla R.L.Zhu, Hikobia 11: 544, 1994 (
*** Cololejeunea leloutrei (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 173, 1963 (
** Cololejeunea leloutrei var. microlobulata Tixier, Bryophyt. Biblioth. 27: 78, 1985 (
** Cololejeunea leloutrei var. ulugurica Pócs ex Tixier, Bryophyt. Biblioth. 27: 74, 1985 (
*** Cololejeunea lemuriana Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 191, 1977 [1979] (
** Cololejeunea littoralis Tixier, Bryophyt. Biblioth. 27: 163, 1985 (
** Cololejeunea longistylis A.Evans, Trans. Connecticut Acad. Arts 10 (8): 453, 1900 (
*** Cololejeunea magnistyla (Horik.) Mizut., J. Hattori Bot. Lab. 24: 243, 1961 (
* Cololejeunea malaccensis Tixier, Bryophyt. Biblioth. 27: 42, 1985 (
** Cololejeunea malayana Tixier, Bryophyt. Biblioth. 27: 154, 1985 (
*** Cololejeunea marginata (Lehm. et Lindenb.) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892 (
* Cololejeunea maritima Tixier, Nova Hedwigia 31: 752, 1979 (
*** Cololejeunea minutilobula Herzog, Rev. Bryol. Lichénol. 20 (1/2): 171, 1951 [1952] (
*** Cololejeunea nigerica (E.W.Jones) R.M.Schust., Beih. Nova Hedwigia 9: 177, 1963 (
** Cololejeunea occidentalis (E.W.Jones) Vanden Berghen, Rev. Bryol. Lichénol. 44 (4): 449, 1978 (
** Cololejeunea onraedtii Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 202, 1977 [1979] (
*** Cololejeunea pacifica Pócs, Acta Bot. Hung. 54 (1/2): 158, 2012 (
** Cololejeunea panamensis G.Dauphin et Pócs, Trop. Bryol. 27: 76, 2006 (
*** Cololejeunea paucimarginata Tixier, Bryophyt. Biblioth. 27: 100, 1985 (
** Cololejeunea perakensis Tixier, Bryophyt. Biblioth. 27: 95, 1985 (
*** Cololejeunea plagiophylla Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 73, 1953 (
*** Cololejeunea planissima (Mitt.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 73, 1959 (
** Cololejeunea planissima var. chagosensis Pócs, J. Bryol. 28 (1): 14, 2006 (
** Cololejeunea praeruptorum Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 205, 1977 [1979] (
** Cololejeunea producta (Mitt.) S.Hatt., Fl. E. Himalaya: 533, 1966 (
* Cololejeunea punctata (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 9, 1893 (
*** Cololejeunea raduliloba Steph., Hedwigia 34 (5): 251, 1895 (
* Cololejeunea reineckeana Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (
** Cololejeunea saltuum Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 188, 1977 [1979] (
** Cololejeunea sambiroana Tixier, Bryologist 82 (4): 608, 1979 (
*** Cololejeunea saroltae Pócs, Acta Bot. Hung. 54 (1/2): 160, 2012 (
*** Cololejeunea schusteri Pócs, Acta Bot. Hung. 44 (3/4): 376, 2002 (
** Cololejeunea schwabei Herzog, J. Hattori Bot. Lab. 14: 54, 1955 (
*** Cololejeunea selangorensis Tixier, Bryophyt. Biblioth. 27: 166, 1985 (
*** Cololejeunea shibiensis Mizut., J. Hattori Bot. Lab. 57: 437, 1984 (
*** Cololejeunea smitinandii Tixier, Bryophyt. Biblioth. 27: 131, 1985 (
** Cololejeunea stoniana Tixier, Bot. Not. 128: 429, 1975 [1976] (
*** Cololejeunea stylosa Steph., Trans. Connecticut Acad. Arts 10 (8): 454, 1900 (
*** Cololejeunea subcardiocarpa Tixier, Bradea 3 (6): 39, 1980 (
** Cololejeunea subinflata Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 182, 1977 [1979] (
*** Cololejeunea submarginata Tixier, Bradea 3 (6): 40, 1980 (
** Cololejeunea subminutilobula Mizut., J. Hattori Bot. Lab. 24: 282, 1961 (
** Cololejeunea subscariosa (Spruce) Pócs, Acta Bot. Hung. 56 (1/2): 197, 2014 (
** Cololejeunea subtriapiculata Tixier, Nova Hedwigia 31: 744, 1979 (
*** Cololejeunea succinea Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 194, 1977 [1979] (
*** Cololejeunea surinamensis Tixier, Bradea 3 (6): 42, 1980 (
** Cololejeunea tahitensis Tixier, Cryptog. Bryol. Lichénol. 14 (3): 359, 1993 (
** Cololejeunea tamatavensis Tixier, Bull. Acad. Malgache (n.ser) 55 (1/2): 191, 1977 [1979] (
* Cololejeunea taprobanea Tixier, Bryophyt. Biblioth. 27: 158, 1985 (
* Cololejeunea thailandensis Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 441, 1973 (
** Cololejeunea triapiculata (Herzog) Tixier, Gard. Bull. Singapore 25 (3): 344, 1971 (
* Cololejeunea tribracteata Tixier, Trop. Bryol. 11: 46, 1995 (
* Cololejeunea tridentata Tixier, Bryophyt. Biblioth. 27: 83, 1985 (
* Cololejeunea uchimae Amakawa, J. Jap. Bot. 33 (5): 142, 1958 (
*** Cololejeunea verwimpii Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995 (
*** Cololejeunea vidaliana Tixier, Nat. Hist. Bull. Siam Soc. 24 (3/4): 444, 1973 (
** Cololejeunea vietnamensis Tixier, Bryophyt. Biblioth. 27: 127, 1985 (
** Cololejeunea vitaliana Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995 (
*** Cololejeunea yakusimensis (S.Hatt.) Mizut., J. Hattori Bot. Lab. 57: 430, 1984 (
* subg. Protocolea R.M.Schust., Beih. Nova Hedwigia 9: 171, 1963 (
** Cololejeunea chuahiana Pócs, Polish Bot. J. 47 (1): 11, 2002 (
** Cololejeunea dauphinii R.L.Zhu, J. Bryol. 28 (3): 277, 2006 (
** Cololejeunea disciflora Tixier, Bryologist 82 (4): 604, 1979 (
*** subg. Taeniolejeunea (Zwickel) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 21, 1953 (
* Cololejeunea amoena Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 25, 1953 (
*** Cololejeunea appressa (A.Evans) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 31, 1953 (
* Cololejeunea bachmaensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 560, 1969 [1970] (
* Cololejeunea bontocensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 583, 1969 [1970] (
* Cololejeunea crassipapillata Tixier, Rev. Bryol. Lichénol. 36 (3/4): 565, 1969 [1970] (
** Cololejeunea eustacei Pócs, J. Bryol. 29 (2): 83, 2007 (
*** Cololejeunea falcata (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 29, 1953 (
** Cololejeunea falcata var. madecassa Tixier, Bryologist 82 (4): 606, 1979 (
* Cololejeunea flavida P.C.Wu et J.S.Lou, Acta Phytotax. Sin. 16 (4): 103, 1978 (
** Cololejeunea flavovittata Pócs, Acta Bryolichenol. Asiat. 4: 120, 2011 (
*** Cololejeunea floccosa (Lehm. et Lindenb.) Schiffn., Consp. Hepat. Arch. Ind.: 243, 1898 (
* Cololejeunea floccosa var. amoenoides Tixier, Cryptog. Bryol. Lichénol. 2 (1): 59, 1981 (
** Cololejeunea floccosa var. angustibracteata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 65, 1981 (
** Cololejeunea floccosa var. aurita Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 24, 1953 (
** Cololejeunea floccosa var. conivens Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 24, 1953 (
* Cololejeunea floccosa var. ocellata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 69, 1981 (
** Cololejeunea floccosa var. plicata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 65, 1981 (
** Cololejeunea floccosa var. trivittata Tixier, Cryptog. Bryol. Lichénol. 2 (1): 62, 1981 (
** Cololejeunea gresicola Tixier, Rev. Bryol. Lichénol. 36 (3/4): 569, 1969 [1970] (
** Cololejeunea gynophthalma Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 32, 1953 (
*** Cololejeunea inflata Steph., Hedwigia 34 (5): 249, 1895 (
** Cololejeunea khiavensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 557, 1969 [1970] (
** Cololejeunea koratensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 585, 1969 [1970] (
* Cololejeunea manlinensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 563, 1969 [1970] (
** Cololejeunea maquilingensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 579, 1969 [1970] (
** Cololejeunea mutabilis Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 27, 1953 (
** Cololejeunea nakajimae S.Hatt., J. Hattori Bot. Lab. 10: 57, 1953 (
*** Cololejeunea ocellata (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 38, 1953 (
*** Cololejeunea ocelloides (Horik.) Mizut., J. Hattori Bot. Lab. 24: 277, 1961 (
*** Cololejeunea peraffinis (Schiffn.) Schiffn., Consp. Hepat. Arch. Ind.: 245, 1898 (
* Cololejeunea peraffinis var. ciconiae Tixier, Bull. Jard. Bot. Natl. Belg. 59 (3/4): 444, 1989 (
** Cololejeunea peraffinis var. elegans Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 35, 1953 (
** Cololejeunea peraffinis var. serrulata Schiffn. ex Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 35, 1953 (
* Cololejeunea polisiana Tixier, Rev. Bryol. Lichénol. 36 (3/4): 550, 1969 [1970] (
*** Cololejeunea pseudofloccosa (Horik.) Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 36, 1953 (
*** Cololejeunea pseudostephanii Tixier, Rev. Bryol. Lichénol. 36 (3/4): 571, 1969 [1970] (
** Cololejeunea setosa Mizut., J. Hattori Bot. Lab. 29: 163, 1966 (
*** Cololejeunea sharpii Mizut., J. Hattori Bot. Lab. 39: 258, 1975 (
*** Cololejeunea siamensis Steph., Bot. Tidsskr. 24 (3): 279, 1902 (
*** Cololejeunea sphaerodonta Mizut., J. Hattori Bot. Lab. 29: 165, 1966 (
*** Cololejeunea stephanii Schiffn. ex Benedix, Feddes Repert. Spec. Nov. Regni Veg. Beih. 134: 40, 1953 (
** Cololejeunea subfloccosa Mizut., J. Hattori Bot. Lab. 57: 168, 1984 (
** Cololejeunea subocelloides Mizut., J. Hattori Bot. Lab. 57: 163, 1984 (
* Cololejeunea tamdaoensis Tixier, Rev. Bryol. Lichénol. 36 (3/4): 567, 1969 [1970] (
** Cololejeunea verdoornii (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 17: 75, 1956 [1957] (
** Cololejeunea yipii R.L.Zhu, Beih. Nova Hedwigia 121: 346, 2001 (
** Cololejeunea zantenorum Pócs, Acta Bryolichenol. Asiat. 4: 127, 2011 (
Incertae sedis
** Cololejeunea conchifolia (Gottsche) Gradst., J. Hattori Bot. Lab. 45: 109, 1979 (
** Cololejeunea dankiaensis Tixier, Phytotaxa 220 (2): 199, 2015 (
** Cololejeunea ensifera Tixier, Phytotaxa 220 (2): 199, 2015 (
** Cololejeunea hamata Steph., Hedwigia 34 (5): 248, 1895 (
** Cololejeunea herzogii K.I.Goebel, Biblioth. Bot. 87 (2): 269, 1916 (
** Cololejeunea jamesii (Austin) M.E.Reiner et Pócs, Phytotaxa 208 (1): 98, 2015 (
** Cololejeunea sublatistyla Jian Wang bis et R.L.Zhu, Phytotaxa 161 (2): 165, 2014 (
** Cololejeunea tixieri Onr., Bull. Jard. Bot. Natl. Belg. 59 (3/4): 436, 1989 (
* Cololejeunea variifolia (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 309, 1896 (
*** Colura (Dumort.) Dumort., Recueil Observ. Jungerm.: 12, 1835 (
** subg. Colura
** sect. Colura
*** Colura berghenii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 245, 1953 [1954] (
*** Colura calyptrifolia (Hook.) Dumort., Recueil Observ. Jungerm.: 12, 1835 (
*** Colura hedbergiana Pócs, J. Bryol. 14 (3): 499, 1987 (
*** Colura humbertii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 251, 1953 [1954] (
*** Colura irrorata (Spruce) Heinrichs, Y.Yu, Schäf.-Verw. et Pócs, Phytotaxa 66: 58, 2012 (
** Colura junghuhniana (Prantl) Steph., Sp. Hepat. (Stephani) 5: 938, 1916 (
** Colura medusa J.Eggers et Pócs, Chenia 11: 22, 2013 (
** Colura mizutanii Pócs, Chenia 11: 22, 2013 (
*** Colura rhynchophora Jovet-Ast, Rev. Bryol. Lichénol. 17 (1/4): 27, 1948 [1949] (
*** Colura tenuicornis (A.Evans) Steph., Sp. Hepat. (Stephani) 5: 942, 1916 (
** sect. Gamolepis Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 207, 1983 (
*** Colura cristata Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 291, 1953 [1954] (
*** Colura greig-smithii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 293, 1953 [1954] (
*** Colura inflata K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 11, 1928 (
*** Colura jovet-astiae Grolle, J. Hattori Bot. Lab. 28: 44, 1965 (
*** Colura meijeri Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 290, 1953 [1954] (
*** Colura obvoluta Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 207, 1983 (
*** Colura ornata K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 26, 1890 [1891] (
*** Colura palawanensis Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 305, 1953 [1954] (
*** Colura valida Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 6, 1961 (
*** Colura verdoornii Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 288, 1953 [1954] (
** sect. Harmophyllum Grolle, J. Hattori Bot. Lab. 28: 44, 1965 (
*** Colura andoi Gradst. et Jovet-Ast, Hikobia 9: 355, 1986 (
*** Colura ari (Steph.) Steph., Sp. Hepat. (Stephani) 5: 936, 1916 (
*** Colura australiensis Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 260, 1953 [1954] (
*** Colura brevistyla Herzog, Beih. Bot. Centralbl. 38 (2): 331, 1921 (
*** Colura calderae Pócs, J. Bryol. 29 (2): 84, 2007 (
*** Colura clementis Grolle, J. Hattori Bot. Lab. 28: 45, 1965 (
*** Colura conica (Sande Lac.) K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 3, 1928 (
*** Colura corynophora (Nees, Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877 (
*** Colura crenulata Grolle, J. Hattori Bot. Lab. 28: 46, 1965 (
*** Colura crispiloba Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 205, 1983 (
** Colura cylindrica Herzog, Svensk Bot. Tidskr. 46 (1): 106, 1952 (
*** Colura cymbalifera Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 268, 1953 [1954] (
*** Colura digitalis (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 931, 1916 (
** Colura digitalis var. mucronata Pócs, Fragm. Florist. Geobot. 40 (1): 263, 1995 (
*** Colura dusenii Steph., Sp. Hepat. (Stephani) 5: 931, 1916 (
*** Colura fastigiata Jovet-Ast, Rev. Bryol. Lichénol. 27 (1/2): 28, 1958 (
*** Colura fistulosa Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 211, 1983 (
*** Colura hattoriana Pócs, J. Hattori Bot. Lab. 74: 47, 1993 (
*** Colura heimii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 275, 1953 [1954] (
*** Colura hemisphaerica Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 267, 1953 [1954] (
*** Colura herzogii Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 261, 1953 [1954] (
*** Colura inuii Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 68, 1931 (
** Colura koponenii Pócs, Chenia 11: 29, 2013 (
** Colura mauritiana Pócs, Cryptog. Bryol. Lichénol. 18 (3): 196, 1997 (
*** Colura maxima Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 284, 1953 [1954] (
*** Colura mosenii Steph., Sp. Hepat. (Stephani) 5: 940, 1916 (
** Colura norrisii Pócs, Chenia 11: 29, 2013 (
*** Colura obesa Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 273, 1953 [1954] (
*** Colura pallida Steph., Sp. Hepat. (Stephani) 5: 941, 1916 (
*** Colura pluridentata Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 265, 1953 [1954] (
*** Colura speciosa Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 307, 1953 [1954] (
** Colura streimannii Pócs, Polish Bot. J. 60 (1): 7, 2015 (
*** Colura superba (Mont.) Steph., Sp. Hepat. (Stephani) 5: 941, 1916 (
*** Colura thomeensis Pócs, Bryologist 114 (2): 363, 2011 (
*** Colura tortifolia (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877 (
*** Colura tutuilana (Pearson) H.A.Mill., Phytologia 47 (4): 322, 1981 (
*** Colura ulei Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 270, 1953 [1954] (
*** Colura vitiensis Pócs et J.Eggers, Polish Bot. J. 52 (2): 88, 2007 (
** sect. Oidocorys Jovet-Ast ex Grolle, J. Hattori Bot. Lab. 32: 140, 1969 (
*** Colura itatyana Steph., Sp. Hepat. (Stephani) 5: 932, 1916 (
*** Colura naumannii (Schiffn.) Steph., Sp. Hepat. (Stephani) 5: 935, 1916 (
*** Colura ornithocephala Herzog, Svensk Bot. Tidskr. 46 (1): 107, 1952 (
*** Colura pulcherrima Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 235, 1953 [1954] (
** Colura pulcherrima var. bartlettii Jovet-Ast, Cryptog. Bryol. Lichénol. 1 (3): 278, 1980 (
*** Colura saccophylla E.A.Hodgs. et Herzog, Trans. & Proc. Roy. Soc. New Zealand 77 (2): 253, 1949 (
*** Colura schusteri Grolle, J. Hattori Bot. Lab. 32: 140, 1969 (
** subg. Glotta Grolle et R.L.Zhu, J. Hattori Bot. Lab. 92: 187, 2002 (
** sect. Glotta Grolle et R.L.Zhu, J. Hattori Bot. Lab. 92: 187, 2002 (
*** Colura bicornis Jovet-Ast, Rev. Bryol. Lichénol. 25 (3/4): 272, 1956 [1957] (
*** Colura bisvoluta Herzog et Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 228, 1953 [1954] (
*** Colura inornata Jovet-Ast, Rev. Bryol. Lichénol. 25 (3/4): 275, 1956 [1957] (
*** Colura karstenii K.I.Goebel, Pflanzenbiol. Schilderungen 2 (1): 153, 1891 (
*** Colura queenslandica B.M.Thiers, Brittonia 39 (2): 175, 1987 (
*** Colura saroltae Pócs, J. Bryol. 14 (3): 497, 1987 (
*** Colura strophiolata Jovet-Ast, Rev. Bryol. Lichénol. 42 (4): 917, 1976 (
*** Colura usambarica E.W.Jones, J. Bryol. 14 (3): 495, 1987 (
** sect. Heterophyllum Jovet-Ast, Cryptog. Bryol. Lichénol. 4 (3): 213, 1983 (
*** Colura acroloba (Prantl) Jovet-Ast, Rev. Bryol. Lichénol. 22 (2/3): 297, 1953 [1954] (
*** Colura corniantha Grolle, J. Hattori Bot. Lab. 28: 44, 1965 (
** Colura denticulata Jovet-Ast, Rev. Bryol. Lichénol. 23 (1/2): 2, 1954 (
*** Colura galeata Jovet-Ast, Rev. Bryol. Lichénol. 35 (1/4): 146, 1967 [1968] (
** Colura imperfecta Steph., Sp. Hepat. (Stephani) 5: 938, 1916 (
*** Colura siamensis Jovet-Ast, Rev. Bryol. Lichénol. 35 (1/4): 139, 1967 [1968] (
*** Colura tixieri Jovet-Ast, Rev. Bryol. Lichénol. 30 (1/2): 9, 1961 (
*** Colura vietnamensis Jovet-Ast et Tixier, Rev. Bryol. Lichénol. 27 (3/4): 205, 1958 [1959] (
Incertae sedis
** Colura amboinensis Steph., Sp. Hepat. (Stephani) 5: 935, 1916 (
*** Colura hirta Steph., Sp. Hepat. (Stephani) 5: 932, 1916 (
*** Diplasiolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 121, 1893 (
*** Diplasiolejeunea utriculata Steph., Sp. Hepat. (Stephani) 5: 920, 1916 (
** subg. Austrolejeuneopsis R.M.Schust., Bull. Torrey Bot. Club 97 (6): 349, 1970 (
*** Diplasiolejeunea alata Jovet-Ast, Rev. Bryol. Lichénol. 17 (1/4): 31, 1948 [1949] (
*** Diplasiolejeunea eggersii Pócs, Bryologist 109 (3): 408, 2006 (
*** Diplasiolejeunea erostrata Schäf.-Verw., Cryptog. Bryol. 25 (1): 3, 2004 (
* Diplasiolejeunea guadalupensis Steph., Sp. Hepat. (Stephani) 5: 924, 1916 (
* Diplasiolejeunea heimii Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 39, 1960 (
** Diplasiolejeunea involuta S.Winkl., Rev. Bryol. Lichénol. 35 (1/4): 320, 1967 [1968] (
** Diplasiolejeunea involuta subsp. andicola Pócs, Cryptog. Bryol. Lichénol. 19 (1): 12, 1998 (
*** Diplasiolejeunea johnsonii A.Evans, Bull. Torrey Bot. Club 39 (12): 603, 1912 [1913] (
** Diplasiolejeunea johnsonii var. mexicana Jovet-Ast, Rev. Bryol. Lichénol. 29 (1/2): 39, 1960 (
* Diplasiolejeunea montecristensis S.Winkl., Rev. Bryol. Lichénol. 35 (1/4): 321, 1967 [1968] (
*** Diplasiolejeunea papilionacea R.M.Schust., Phytologia 39 (6): 431, 1978 (
*** Diplasiolejeunea pauckertii (Nees) Steph., Sp. Hepat. (Stephani) 5: 924, 1916 (
*** Diplasiolejeunea pluridentata Schäf.-Verw., Haussknechtia 8: 71, 2001 (
*** Diplasiolejeunea replicata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 926, 1916 (
** Diplasiolejeunea rudolphiana Steph., Hedwigia 35 (3): 79, 1896 (
*** Diplasiolejeunea runssorensis Steph., Bot. Jahrb. Syst. 20 (3): 318, 1895 (
*** Diplasiolejeunea unidentata (Lehm. et Lindenb.) Schiffn., Bot. Jahrb. Syst. 23 (5): 583, 1897 (
** subg. Diplasiolejeunea
*** Diplasiolejeunea armatiloba Steph., Hedwigia 35 (3): 80, 1896 (
*** Diplasiolejeunea borhidiana Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 177, 1982 [1983] (
*** Diplasiolejeunea brunnea Steph., Sp. Hepat. (Stephani) 5: 922, 1916 (
** Diplasiolejeunea caribea Tixier, Bryophyt. Biblioth. 27: 377, 1985 (
*** Diplasiolejeunea cavifolia Steph., Sp. Hepat. (Stephani) 5: 918, 1916 (
*** Diplasiolejeunea cobrensis Steph., Sp. Hepat. (Stephani) 5: 923, 1916 (
** Diplasiolejeunea cobrensis subsp. antsirananae Pócs, Candollea 56 (1): 72, 2001 (
** Diplasiolejeunea cogoensis M.Infante, Heras et Pócs, Trop. Bryol. 17: 9, 1999 (
** Diplasiolejeunea cubensis Tixier, Bryophyt. Biblioth. 27: 389, 1985 (
* Diplasiolejeunea glaziovii Tixier, Bryophyt. Biblioth. 27: 397, 1985 (
*** Diplasiolejeunea ingekarolae Schäf.-Verw., Herzogia 19: 239, 2006 (
** Diplasiolejeunea jonesii Tixier, Acta Bot. Hung. 30 (1/2): 21, 1984 (
*** Diplasiolejeunea leiocarpa Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 34, 1947 [1948] (
*** Diplasiolejeunea malleiformis (A.Evans) Tixier, Bryophyt. Biblioth. 27: 351, 1985 (
** Diplasiolejeunea palustrium Tixier, Acta Bot. Hung. 30 (1/2): 19, 1984 (
*** Diplasiolejeunea pellucida (C.F.W.Meissn. ex Spreng.) Schiffn., Hepat. (Engl.-Prantl): 121, 1893 (
** Diplasiolejeunea phyllarthronii Tixier, Acta Bot. Hung. 30 (1/2): 19, 1984 (
*** Diplasiolejeunea pocsii Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 173, 1982 [1983] (
** subg. Physolejeunea R.M.Schust., Bull. Torrey Bot. Club 97 (6): 348, 1970 (
*** Diplasiolejeunea andringitrae Schäf.-Verw., Cryptog. Bryol. 27 (4): 447, 2006 (
*** Diplasiolejeunea cornuta Steph., Sp. Hepat. (Stephani) 5: 918, 1916 (
*** Diplasiolejeunea jovet-astiae Grolle, Feddes Repert. 73 (2): 84, 1966 (
*** Diplasiolejeunea kraussiana (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 919, 1916 (
*** Diplasiolejeunea ornata Pócs et Schäf.-Verw., Cryptog. Bryol. 27 (4): 440, 2006 (
*** Diplasiolejeunea patelligera Herzog, Svensk Bot. Tidskr. 42 (3): 240, 1948 (
*** Diplasiolejeunea plicatiloba (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 45: 175, 1979 (
*** Diplasiolejeunea pusilla Grolle, Feddes Repert. 86 (1/2): 79, 1975 (
*** Diplasiolejeunea ranomafanae Pócs, Cryptog. Bryol. 27 (4): 444, 2006 (
*** Diplasiolejeunea villaumei Steph., Sp. Hepat. (Stephani) 5: 921, 1916 (
*** Diplasiolejeunea zakiae Tixier, Lindbergia 4 (1/2): 123, 1977 (
Incertae sedis
*** Diplasiolejeunea albifolia (Taylor) E.W.Jones, Trans. Brit. Bryol. Soc. 2 (3): 393, 1954 (
*** Diplasiolejeunea aulae E.W.Jones, J. Bryol. 7 (4): 552, 1973 [1974] (
** Diplasiolejeunea auriculata Tixier, Rev. Bryol. Lichénol. 45 (2): 210, 1979 (
*** Diplasiolejeunea buckii Grolle, Beitr. Phytotax. 15: 105, 1992 (
** Diplasiolejeunea columbica Tixier, Cryptog. Bryol. Lichénol. 4 (3): 233, 1983 (
* Diplasiolejeunea comorensis Tixier, Rev. Bryol. Lichénol. 45 (2): 213, 1979 (
** Diplasiolejeunea cyanguguensis Tixier, Trop. Bryol. 11: 67, 1995 (
*** Diplasiolejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 217, 1977 (
** Diplasiolejeunea ensifera Tixier, Rev. Bryol. Lichénol. 45 (2): 215, 1979 (
* Diplasiolejeunea evansii Tixier, Bryophyt. Biblioth. 27: 360, 1985 (
** Diplasiolejeunea gradsteinii Tixier, Trop. Bryol. 11: 67, 1995 (
*** Diplasiolejeunea grandirostrata Schäf.-Verw., Cryptog. Bryol. 25 (1): 7, 2004 (
*** Diplasiolejeunea grolleana Reyes Montoya, Acta Bot. Acad. Sci. Hung. 28 (1/2): 175, 1982 [1983] (
** Diplasiolejeunea hamata Tixier, Rev. Bryol. Lichénol. 45 (2): 217, 1979 (
** Diplasiolejeunea insignis Tixier, Lindbergia 4 (1/2): 120, 1977 (
* Diplasiolejeunea integerrima Tixier, Rev. Bryol. Lichénol. 45 (2): 219, 1979 (
*** Diplasiolejeunea lanceolata Grolle, Beitr. Phytotax. 15: 107, 1992 (
*** Diplasiolejeunea latipuensis Tixier, Cryptog. Bryol. Lichénol. 16 (3): 230, 1995 (
** Diplasiolejeunea lemuriana Tixier, Lindbergia 4 (1/2): 120, 1977 (
** Diplasiolejeunea longilobula Herzog, Trans. Brit. Bryol. Soc. 1 (4): 325, 1950 (
** Diplasiolejeunea magnistipula Tixier, Ann. Fac. Sci. Yaoundé 20: 7, 1975 (
*** Diplasiolejeunea mayaykuensis Schäf.-Verw. et Heinrichs, Polish Bot. J. 58 (1): 143, 2013 (
*** Diplasiolejeunea onraedtii Grolle, Feddes Repert. 89 (5/6): 301, 1978 (
** Diplasiolejeunea ramicola Tixier, Rev. Bryol. Lichénol. 45 (2): 222, 1979 (
** Diplasiolejeunea riclefgrollei Schäf.-Verw., Cryptog. Bryol. 26 (1): 37, 2005 (
** Diplasiolejeunea subcornuta Tixier, Lindbergia 4 (1/2): 122, 1977 (
*** Diplasiolejeunea symoensii Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 92: 126, 1960 (
** Haplolejeunea Grolle, J. Hattori Bot. Lab. 39: 205, 1975 (
*** Haplolejeunea cucullata (Steph.) Grolle, J. Hattori Bot. Lab. 45: 176, 1979 (
** Haplolejeunea sticta Grolle, J. Hattori Bot. Lab. 39: 205, 1975 (
** Macrocolura R.M.Schust., J. Hattori Bot. Lab. 75: 233, 1994 (
*** Macrocolura sagittistipula (Spruce) R.M.Schust., J. Hattori Bot. Lab. 75: 233, 1994 (
*** Myriocoleopsis Schiffn., Hedwigia 81 (5/6): 234, 1944 (
*** Myriocoleopsis fluviatilis (Steph.) M.E.Reiner et Gradst., J. Bryol. 19 (3): 639, 1997 (
*** Myriocoleopsis gymnocolea (Spruce) M.E.Reiner et Gradst., J. Bryol. 19 (3): 640, 1997 (
*** Myriocoleopsis minutissima (Sm.) R.L.Zhu, Y.Yu et Pócs, Phytotaxa 183 (4): 293, 2014 (
*** Myriocoleopsis minutissima subsp. myriocarpa (Nees et Mont.) R.L.Zhu, Y.Yu et Pócs, Phytotaxa 183 (4): 294, 2014 (
** Myriocoleopsis vuquangensis (Pócs et Ninh) Pócs, Trop. Bryol. 31: 124, 2010 (
** Nephelolejeunea Grolle, J. Hattori Bot. Lab. 37: 252, 1973 (
** Nephelolejeunea bidentata B.M.Thiers ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 65, 2015 (
** Nephelolejeunea carcharias M.A.M.Renner, Syst. Bot. 34 (4): 621, 2009 (
*** Nephelolejeunea conchophylla Grolle, J. Hattori Bot. Lab. 48: 164, 1980 (
** Nephelolejeunea fragilis (R.M.Schust.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 65, 2015 (
*** Nephelolejeunea hamata Grolle, J. Hattori Bot. Lab. 48: 167, 1980 (
*** Nephelolejeunea hispida R.M.Schust. ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (
*** Nephelolejeunea jarmaniana Grolle ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (
*** Nephelolejeunea nudipes (Hook.f. et Taylor) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (
** Nephelolejeunea occidentalis Pócs ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (
*** Nephelolejeunea papillosa Glenny, New Zealand J. Bot. 34 (2): 195, 1996 (
*** Nephelolejeunea radulifolia (C.Massal.) L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (
** Nephelolejeunea secunda M.A.M.Renner ex L.Söderstr. et A.Hagborg, Phytotaxa 202 (1): 66, 2015 (
*** Nephelolejeunea talinayi (S.W.Arnell) Grolle, J. Hattori Bot. Lab. 37: 253, 1973 (
** Schusterolejeunea Grolle, J. Bryol. 11 (1): 105, 1980 (
*** Schusterolejeunea inundata (Spruce) Grolle, J. Bryol. 11 (1): 105, 1980 (
** Siphonolejeunea Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 744, 1942 (
** Siphonolejeunea elegantissima (Steph.) Grolle, J. Hattori Bot. Lab. 41: 405, 1976 (
** Siphonolejeunea neesii (Mont.) Bischl., Nova Hedwigia 17: 338, 1969 (
*** Siphonolejeunea schiffneri (Schiffn.) Herzog, Svensk Bot. Tidskr. 42 (3): 230, 1948 (
** Tuyamaella S.Hatt., J. Hattori Bot. Lab. 5: 60, 1951 (
** Tuyamaella angulistipa (Steph.) R.M.Schust. et Kachroo, J. Linn. Soc., Bot. 56 (368): 508, 1961 (
** Tuyamaella borneensis Tixier, Rev. Bryol. Lichénol. 39 (2): 241, 1973 (
** Tuyamaella hattorii Tixier, Rev. Bryol. Lichénol. 31 (3/4): 188, 1962 [1963] (
** Tuyamaella jackii (Steph.) Tixier, Rev. Bryol. Lichénol. 39 (2): 235, 1973 (
*** Tuyamaella molischii (Schiffn.) S.Hatt., J. Hattori Bot. Lab. 5: 62, 1951 (
** Tuyamaella molischii var. brevistipa P.C.Wu et P.J.Lin, Acta Phytotax. Sin. 16 (2): 65, 1978 (
** Tuyamaella molischii var. taiwanensis R.L.Zhu et M.L.So, Nova Hedwigia 70 (1/2): 190, 2000 (
** Tuyamaella serratistipa S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 118, 1951 (
*** Bromeliophila R.M.Schust., J. Hattori Bot. Lab. 75: 226, 1994 (
*** Bromeliophila helenae Gradst., Cryptog. Bryol. Lichénol. 18 (3): 218, 1997 (
*** Bromeliophila natans (Steph.) R.M.Schust., J. Hattori Bot. Lab. 75: 226, 1994 (
*** Cyclolejeunea A.Evans, Bull. Torrey Bot. Club 31 (4): 193, 1904 (
*** Cyclolejeunea accedens (Gottsche) A.Evans, Bull. Torrey Bot. Club 31 (4): 201, 1904 (
*** Cyclolejeunea chitonia (Taylor) A.Evans, Bull. Torrey Bot. Club 31 (4): 194, 1904 (
*** Cyclolejeunea convexistipa (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club 31 (4): 198, 1904 (
* Cyclolejeunea ecuadorensis Steph., Sp. Hepat. (Stephani) 5: 194, 1913 (
*** Cyclolejeunea foliorum (Nees) Grolle, J. Hattori Bot. Lab. 65: 403, 1988 (
* Cyclolejeunea integerrima (Steph.) Steph., Sp. Hepat. (Stephani) 5: 195, 1913 (
*** Cyclolejeunea luteola (Spruce) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 33: 761, 1984 (
*** Cyclolejeunea peruviana (Lehm. et Lindenb.) A.Evans, Bull. Torrey Bot. Club 31 (4): 196, 1904 (
* Cyclolejeunea spectabilis Steph., Sp. Hepat. (Stephani) 5: 193, 1913 (
*** Prionolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
*** Prionolejeunea aemula (Gottsche) A.Evans, Bull. Torrey Bot. Club 31 (4): 219, 1904 (
*** Prionolejeunea ampliretis Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 178, 1955 (
*** Prionolejeunea arguta (Nees) Steph., Sp. Hepat. (Stephani) 5: 204, 1913 (
*** Prionolejeunea decora (Taylor) Steph., Sp. Hepat. (Stephani) 5: 207, 1913 (
*** Prionolejeunea denticulata (F.Weber) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
*** Prionolejeunea diversitexta (Hampe et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 209, 1913 (
*** Prionolejeunea exauriculata A.Evans, Bull. Torrey Bot. Club 31 (4): 223, 1904 (
*** Prionolejeunea galliotii Steph., Sp. Hepat. (Stephani) 6: 388, 1923 (
*** Prionolejeunea grata (Gottsche) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
*** Prionolejeunea grollei Ilk.-Borg. et Schäf.-Verw., Cryptog. Bryol. 26 (1): 29, 2005 (
*** Prionolejeunea guadalupensis (Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 211, 1913 (
*** Prionolejeunea limpida Herzog, Hedwigia 67 (6): 252, 1927 (
*** Prionolejeunea magnistipula Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 176, 1955 (
*** Prionolejeunea meissneri (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 212, 1913 (
*** Prionolejeunea mucronata (Sande Lac.) Steph., Hedwigia 35 (3): 119, 1896 (
*** Prionolejeunea muricatoserrulata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 223, 1913 (
*** Prionolejeunea principensis Vanden Berghen, Rev. Bryol. Lichénol. 29 (1/2): 65, 1960 (
*** Prionolejeunea recurvula (Spruce) Steph., Sp. Hepat. (Stephani) 5: 224, 1913 (
*** Prionolejeunea scaberula (Spruce) Steph., Sp. Hepat. (Stephani) 5: 214, 1913 (
*** Prionolejeunea schlimiana (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 214, 1913 (
*** Prionolejeunea trachyodes (Spruce) Steph., Sp. Hepat. (Stephani) 5: 215, 1913 (
Excluded from the genus
* Prionolejeunea corbisieri Pearson, Natuurw. Tijdschr. 4 (5/6): 128, 1922 (
* Prionolejeunea maculata Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 180, 1955 (
** Capillolejeunea S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 69, 1965 (
** Capillolejeunea geisslerae (Pócs) R.L.Zhu, Qiong He, Y.M.Wei et Pócs, Phytotaxa 175 (3): 167, 2014 (
** Capillolejeunea mascarena S.W.Arnell, Svensk Bot. Tidskr. 59 (1): 69, 1965 (
*** Drepanolejeunea (Spruce) Steph., Hedwigia 30 (5): 209, 1891 (
** subg. Acantholejeunea R.M.Schust., Beih. Nova Hedwigia 9: 115, 1963 (
** Drepanolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 343, 1913 (
** Drepanolejeunea spinistipula Herzog, Svensk Bot. Tidskr. 42 (3): 238, 1948 (
*** subg. Drepanolejeunea
*** Drepanolejeunea aculeata Bischl., Rev. Bryol. Lichénol. 33 (1/2): 68, 1964 (
** Drepanolejeunea anderssonii (Ångstr.) A.Evans, Trans. Connecticut Acad. Arts 10 (8): 429, 1900 (
*** Drepanolejeunea andina Herzog, Svensk Bot. Tidskr. 51 (1): 196, 1957 (
*** Drepanolejeunea angustifolia (Mitt.) Grolle, J. Jap. Bot. 40 (7): 206, 1965 (
*** Drepanolejeunea ankasica E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 245, 1983 (
*** Drepanolejeunea anoplantha (Spruce) Steph., Sp. Hepat. (Stephani) 5: 325, 1913 (
*** Drepanolejeunea appalachiana R.M.Schust., J. Elisha Mitchell Sci. Soc. 83 (4): 219, 1967 (
*** Drepanolejeunea araucariae Steph., Hedwigia 35 (3): 80, 1896 (
** Drepanolejeunea araucariae var. chilensis Herzog, Acta Horti Gotob. 15: 161, 1943 (
** Drepanolejeunea aucklandica Steph., Sp. Hepat. (Stephani) 5: 358, 1913 (
*** Drepanolejeunea aurita Bischl., Rev. Bryol. Lichénol. 33 (1/2): 146, 1964 (
** Drepanolejeunea bakeri Herzog, Ann. Bryol. 3: 148, 1930 (
*** Drepanolejeunea bidens (Prantl) A.Evans, Bull. Torrey Bot. Club 30 (1): 29, 1903 (
*** Drepanolejeunea biocellata A.Evans, Bull. Torrey Bot. Club 30 (1): 22, 1903 (
** Drepanolejeunea blumei Steph., Hedwigia 35 (3): 81, 1896 (
** Drepanolejeunea brunnea Mizut., J. Hattori Bot. Lab. 33: 231, 1970 (
** Drepanolejeunea caledonica Steph., Sp. Hepat. (Stephani) 5: 342, 1913 (
*** Drepanolejeunea campanulata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 328, 1913 (
** Drepanolejeunea canceroides H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 53, 1963 (
** Drepanolejeunea capulata (Taylor) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892 (
** Drepanolejeunea capulata var. flagellifera S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 100, 1964 (
** Drepanolejeunea ciliata Mizut., J. Hattori Bot. Lab. 33: 232, 1970 (
*** Drepanolejeunea crassiretis A.Evans, Bull. Torrey Bot. Club 30 (1): 25, 1903 (
*** Drepanolejeunea crucianella (Taylor) A.Evans, Bull. Torrey Bot. Club 30 (1): 33, 1903 (
*** Drepanolejeunea cultrella (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 324, 1913 (
** Drepanolejeunea cutervoensis (Loitl.) Grolle, J. Hattori Bot. Lab. 69: 186, 1991 (
** Drepanolejeunea decurviloba Steph., Sp. Hepat. (Stephani) 6: 397, 1923 (
* Drepanolejeunea deslooveri Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 47 (1/2): 210, 1977 (
* Drepanolejeunea dissitifolia A.Evans, Bull. Torrey Bot. Club 30 (1): 28, 1903 (
** Drepanolejeunea elegans Herzog, Ann. Bryol. 9: 128, 1936 [1937] (
** Drepanolejeunea erecta (Steph.) Mizut., J. Hattori Bot. Lab. 40: 442, 1976 (
** Drepanolejeunea evansii Bischl. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 47, 2012 (
*** Drepanolejeunea fragilis Bischl. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 47, 2012 (
** Drepanolejeunea fulfordiae L.Söderstr., Phytotaxa 208 (1): 98, 2015 (
*** Drepanolejeunea granatensis (Prantl) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 150, 1964 (
** Drepanolejeunea grandis Herzog, Ann. Bryol. 12: 113, 1939 (
*** Drepanolejeunea grollei M.E.Reiner et Schäf.-Verw., Candollea 51 (2): 475, 1996 (
** Drepanolejeunea grossidens Steph., Sp. Hepat. (Stephani) 5: 358, 1913 (
*** Drepanolejeunea hamatifolia (Hook.) Schiffn., Hepat. (Engl.-Prantl): 126, 1893 (
** Drepanolejeunea hamulata Steph., Sp. Hepat. (Stephani) 5: 331, 1913 (
*** Drepanolejeunea helenae Pócs, Cryptog. Bryol. Lichénol. 18 (3): 198, 1997 (
** Drepanolejeunea herzogii R.L.Zhu et M.L.So, Beih. Nova Hedwigia 121: 181, 2001 (
*** Drepanolejeunea inchoata (C.F.W.Meissn.) Steph., Primit. fl. costar.: 115, 1892 [1893] (
** Drepanolejeunea inchoata var. palmicola Pócs, Acta Bot. Hung. 51 (3/4): 381, 2009 (
** Drepanolejeunea inchoata var. roraimae (Steph. ex Zwickel) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 48, 1964 (
*** Drepanolejeunea infundibulata (Spruce) A.Evans, Bull. Torrey Bot. Club 30 (1): 35, 1903 (
*** Drepanolejeunea integribracteata Bischl., Rev. Bryol. Lichénol. 33 (1/2): 142, 1964 (
** Drepanolejeunea laevis (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913 (
* Drepanolejeunea lancifolia (Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892 (
*** Drepanolejeunea lichenicola (Spruce) Steph., Sp. Hepat. (Stephani) 5: 335, 1913 (
** Drepanolejeunea longii Grolle et R.L.Zhu, Ann. Bot. Fenn. 36 (2): 115, 1999 (
** Drepanolejeunea macrodonta (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913 (
** Drepanolejeunea mawtmiana Ajit P.Singh et V.Nath, Hepat. Khasi Jaintia Hills: E. Himal.: 251, 2007 (
** Drepanolejeunea microcarpa Pearson, J. Linn. Soc., Bot. 46 (305): 36, 1922 (
** Drepanolejeunea moluccensis Herzog, Ann. Bryol. 7: 88, 1934 (
*** Drepanolejeunea mosenii (Steph.) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 118, 1967 [1968] (
* Drepanolejeunea obliqua Steph., Hedwigia 35 (3): 82, 1896 (
** Drepanolejeunea obtriangulata T.Kodama, J. Hattori Bot. Lab. 41: 381, 1976 (
** Drepanolejeunea obtusifolia T.Yamag., J. Jap. Bot. 59 (11): 332, 1984 (
*** Drepanolejeunea orthophylla (Nees et Mont.) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 102, 1967 [1968] (
*** Drepanolejeunea palmifolia (Nees) Schiffn., Hepat. (Engl.-Prantl): 126, 1893 (
*** Drepanolejeunea pentadactyla (Mont.) Steph., Sp. Hepat. (Stephani) 5: 357, 1913 (
** Drepanolejeunea perissodonta (Spruce) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 73, 1964 (
*** Drepanolejeunea physifolia (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 8, 1892 (
** Drepanolejeunea pinnatiloba Schiffn., Bot. Jahrb. Syst. 23 (5): 591, 1897 (
** Drepanolejeunea pleiodictya Herzog, Ann. Bryol. 7: 89, 1934 (
** Drepanolejeunea propagulifera Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 204, 1931 (
** Drepanolejeunea pseudoneura (A.Evans) Grolle, J. Hattori Bot. Lab. 65: 405, 1988 (
** Drepanolejeunea pterocalyx (Herzog) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 114, 1967 [1968] (
** Drepanolejeunea pungens Bischl., Rev. Bryol. Lichénol. 33 (1/2): 104, 1964 (
** Drepanolejeunea ramentiflora Steph., Sp. Hepat. (Stephani) 5: 338, 1913 (
* Drepanolejeunea ruandensis Vanden Berghen, Bull. Soc. Roy. Bot. Belgique 93: 63, 1961 (
*** Drepanolejeunea senticosa Bischl., Rev. Bryol. Lichénol. 33 (1/2): 96, 1964 (
** Drepanolejeunea sikkimensis (Udar et U.S.Awasthi) Grolle, J. Hattori Bot. Lab. 55: 503, 1984 (
** Drepanolejeunea spinosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 185, 1955 (
** Drepanolejeunea spinosocornuta Steph., Sp. Hepat. (Stephani) 5: 351, 1913 (
* Drepanolejeunea subdissitifolia Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 65, 1950 (
** Drepanolejeunea submuricata R.M.Schust., Phytotaxa 208 (1): 98, 2015 (
** Drepanolejeunea subquadrata (Mitt.) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 311, 1896 (
** Drepanolejeunea subvittata (Herzog) Grolle, J. Hattori Bot. Lab. 69: 186, 1991 (
* Drepanolejeunea tenax K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 15, 1928 (
** Drepanolejeunea tenera K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 20, 1928 (
* Drepanolejeunea tenera var. litoceras Herzog, Ann. Bryol. 7: 85, 1934 (
*** Drepanolejeunea ternatensis (Gottsche) Schiffn., Hepat. (Engl.-Prantl): 126, 1893 (
** Drepanolejeunea teysmannii (Gottsche) Steph., Hedwigia 35 (3): 84, 1896 (
** Drepanolejeunea tridactyla (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 354, 1913 (
*** Drepanolejeunea trigonophylla Steph., Hedwigia 35 (3): 85, 1896 (
** Drepanolejeunea tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 7, 1958 (
** Drepanolejeunea tuyamae S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 116, 1951 (
** Drepanolejeunea ualanensis Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 148, 1965 (
** Drepanolejeunea ungulata (Steph.) Grolle, J. Hattori Bot. Lab. 69: 187, 1991 (
** Drepanolejeunea urceolata R.M.Schust., Phytologia 39 (6): 427, 1978 (
** Drepanolejeunea valiae Jovet-Ast, Rev. Bryol. Lichénol. 18 (1/2): 38, 1949 (
** Drepanolejeunea vandenberghenii Buchb. et Eb.Fisch., J. Bryol. 26 (4): 273, 2004 (
*** Drepanolejeunea vesiculosa (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 356, 1913 (
** Drepanolejeunea yulensis Steph., Sp. Hepat. (Stephani) 5: 356, 1913 (
*** subg. Kolpolejeunea Grolle, J. Hattori Bot. Lab. 40: 193, 1976 (
*** Drepanolejeunea intermedia Zwickel, Ann. Bryol. 6: 119, 1933 (
*** Drepanolejeunea lyrata Grolle, J. Hattori Bot. Lab. 40: 199, 1976 (
*** Drepanolejeunea madagascariensis (Steph.) Grolle, Lindbergia 2 (3/4): 232, 1974 (
*** Drepanolejeunea pocsii Grolle, J. Hattori Bot. Lab. 40: 209, 1976 (
** Drepanolejeunea symoensii Vanden Berghen et Grolle, J. Hattori Bot. Lab. 49: 86, 1981 (
** Drepanolejeunea symoensii var. minor Tixier, Trop. Bryol. 11: 25, 1995 (
*** Drepanolejeunea trematodes (Nees) Bischl., Rev. Bryol. Lichénol. 35 (1/4): 125, 1967 [1968] (
** subg. Pristolejeunea Grolle, J. Hattori Bot. Lab. 40: 193, 1976 (
** Drepanolejeunea actinogyna Inuthai, R.L.Zhu et Chantanaorr., Bryologist 117 (2): 165, 2014 (
*** Drepanolejeunea fissicornua Steph., Sp. Hepat. (Stephani) 5: 344, 1913 (
* Drepanolejeunea hampeana Steph., Sp. Hepat. (Stephani) 5: 345, 1913 (
** Drepanolejeunea laciniata Qiong He et R.L.Zhu, Cryptog. Bryol. 33 (3): 292, 2012 (
*** Drepanolejeunea levicornua Steph., Sp. Hepat. (Stephani) 5: 347, 1913 (
* Drepanolejeunea longicornua (Herzog) Mizut., J. Hattori Bot. Lab. 68: 368, 1990 (
* Drepanolejeunea nymanii Steph., Sp. Hepat. (Stephani) 5: 348, 1913 (
** Drepanolejeunea pulla (Mitt.) Grolle, J. Hattori Bot. Lab. 46: 349, 1979 (
* Drepanolejeunea serricalyx Herzog, Ann. Bryol. 9: 126, 1936 [1937] (
*** Drepanolejeunea thwaitesiana (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 350, 1913 (
** Drepanolejeunea thwaitesiana var. zhengii R.L.Zhu, Beih. Nova Hedwigia 121: 197, 2001 (
*** Drepanolejeunea tricornua Herzog, Ann. Bryol. 9: 124, 1936 [1937] (
*** subg. Rhaphidolejeunea (Herzog) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 376, 2000 (
** Drepanolejeunea bidoupensis Pócs, Cryptog. Bryol. 34 (3): 293, 2013 (
*** Drepanolejeunea bischlerae (Grolle) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 391, 2000 (
*** Drepanolejeunea commutata Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 377, 2000 (
*** Drepanolejeunea cyclops (Sande Lac.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 391, 2000 (
*** Drepanolejeunea fleischeri (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 379, 2000 (
** Drepanolejeunea foliicola Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 85, 1932 (
** Drepanolejeunea longicruris (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 392, 2000 (
*** Drepanolejeunea polyrhiza (Nees) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 392, 2000 (
*** Drepanolejeunea siamensis (Bischl.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 393, 2000 (
*** Drepanolejeunea spicata (Steph.) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 384, 2000 (
*** Drepanolejeunea tibetana (P.C.Wu et J.S.Lou) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 386, 2000 (
*** Drepanolejeunea yunnanensis (P.C.Chen) Grolle et R.L.Zhu, Nova Hedwigia 70 (3/4): 388, 2000 (
Incertae sedis
*** Drepanolejeunea dactylophora (Nees, Lindenb. et Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 12, 1892 (
*** Drepanolejeunea dactylophora var. submuricata Herzog, Ann. Bryol. 4: 92, 1931 (
* Drepanolejeunea devendrae Sushil K.Singh et M.Dey, Nelumbo 54: 20, 2012 (
* Drepanolejeunea integerrima Herzog, Rev. Bryol. Lichénol. 20 (1/2): 146, 1951 [1952] (
** Vitalianthus R.M.Schust. et Giancotti, Nova Hedwigia 57 (3/4): 447, 1993 (
*** Vitalianthus bischlerianus (K.C.Pôrto et Grolle) R.M.Schust. et Giancotti, Nova Hedwigia 57 (3/4): 448, 1993 (
** Vitalianthus guangxianus R.L.Zhu, Qiong He et Y.M.Wei, J. Bryol. 34 (1): 32, 2012 (
*** Anoplolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 131, 1893 (
*** Anoplolejeunea conferta (C.F.W.Meissn. ex Spreng.) A.Evans, Bull. Torrey Bot. Club 35 (4): 175, 1908 (
*** Echinolejeunea R.M.Schust., Beih. Nova Hedwigia 9: 187, 1963 (
*** Echinolejeunea papillata (Mitt.) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 260, 1972 (
** Kymatolejeunea Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 32 (6): 1005, 1984 (
*** Kymatolejeunea bartlettii Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 32 (6): 1005, 1984 (
** Leiolejeunea A.Evans, Bull. Torrey Bot. Club 35 (8): 377, 1908 (
*** Leiolejeunea grandiflora A.Evans, Bull. Torrey Bot. Club 35 (8): 378, 1908 (
*** Harpalejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 126, 1893 (
** subg. Cleefiolejeunea Grolle et M.E.Reiner, J. Bryol. 21 (1): 33, 1999 (
*** Harpalejeunea grandis Grolle et M.E.Reiner, J. Bryol. 21 (1): 32, 1999 (
** subg. Harpalejeunea
* Harpalejeunea acuta S.Winkl., Rev. Bryol. Lichénol. 42 (3): 812, 1976 (
*** Harpalejeunea ancistrodes (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
** Harpalejeunea buenaventurae Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 183, 1955 (
*** Harpalejeunea cinchonae (Nees) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
** Harpalejeunea cinchonae var. strigulosa Herzog, Svensk Bot. Tidskr. 51 (1): 192, 1957 (
** Harpalejeunea decurvicuspis (Besch. et C.Massal.) P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (
** Harpalejeunea emarginata Jovet-Ast, Rev. Bryol. Lichénol. 16 (1/2): 38, 1947 [1948] (
** Harpalejeunea exocellata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 144, 1951 [1952] (
** Harpalejeunea grandistipula R.M.Schust., J. Hattori Bot. Lab. 87: 290, 1999 (
*** Harpalejeunea harpaphylla (Herzog) Bischl., Rev. Bryol. Lichénol. 33 (1/2): 164, 1964 (
** Harpalejeunea herzogii Jovet-Ast, Feddes Repert. Spec. Nov. Regni Veg. 58: 19, 1955 (
** Harpalejeunea longibracteata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 254, 1913 (
** Harpalejeunea marginalis (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 271, 1913 (
*** Harpalejeunea molleri (Steph.) Grolle, Taxon 38 (1): 89, 1989 (
** Harpalejeunea molleri subsp. integra (R.M.Schust.) Damsh., Ill. Fl. Nord. Liverw. Hornw.: 615, 2002 (
*** Harpalejeunea oxyphylla (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 255, 1913 (
*** Harpalejeunea parasitica (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 268, 1913 (
** Harpalejeunea pinaundensis Grolle, J. Hattori Bot. Lab. 46: 44, 1979 (
** Harpalejeunea reflexula A.Evans, Bull. Torrey Bot. Club 35 (8): 375, 1908 (
** Harpalejeunea scabra Gradst. et Schäf.-Verw., Cryptog. Bryol. 32 (2): 102, 2011 (
*** Harpalejeunea schiffneri S.W.Arnell, Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 102, 1964 (
* Harpalejeunea solitaria (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 246, 1913 (
*** Harpalejeunea stricta (Lindenb. et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 259, 1913 (
** Harpalejeunea subacuta A.Evans, Bull. Torrey Bot. Club 30 (10): 547, 1903 (
* Harpalejeunea tenuicuspis (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
*** Harpalejeunea tridens (Besch. et Spruce) Steph., Sp. Hepat. (Stephani) 5: 263, 1913 (
** Harpalejeunea uncinata Steph., Hedwigia 35 (3): 97, 1896 (
* Harpalejeunea uncinata var. setulosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 183, 1955 (
Incertae sedis
* Harpalejeunea grossearmata Steph., Biblioth. Bot. 87 (2): 256, 1916 (
* Harpalejeunea renneri Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 54, 1950 [1951] (
* Harpalejeunea spruceana Steph., Biblioth. Bot. 87 (2): 257, 1916 (
* Harpalejeunea vitrea Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 94, 1952 (
** Hattoriolejeunea Mizut., J. Hattori Bot. Lab. 61: 303, 1986 [1987] (
** Hattoriolejeunea akiyamae Mizut., J. Hattori Bot. Lab. 61: 303, 1986 [1987] (
*** Lejeunea Lib., Ann. Gen. Sci. Phys. 6: 373, 1820 (
** Lejeunea tunquiniensis M.E.Reiner et Drehwald, Nova Hedwigia 100 (3/4): 584, 2015 (
** subg. Lejeunea
*** Lejeunea abyssinica (Gola) Cufod., Phyton (Horn) 4: 75, 1952 (
*** Lejeunea acanthogona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 177, 1884 (
** Lejeunea acuminata (Lehm. et Lindenb.) Lehm., Nov. Stirp. Pug. 7: 22, 1838 (
** Lejeunea acuta Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 167, 1863 (
** Lejeunea acutata (Steph.) Solari, J. Hattori Bot. Lab. 54: 541, 1983 (
*** Lejeunea adpressa Nees, Repert. Pharm. 76: 45, 1842 (
*** Lejeunea aethiopica E.W.Jones, J. Bryol. 13 (3): 387, 1985 (
** Lejeunea alaskana (R.M.Schust. et Steere) Inoue et Steere, J. Hattori Bot. Lab. 44: 330, 1978 (
*** Lejeunea alata Gottsche, Syn. Hepat. 3: 406, 1845 (
** Lejeunea alata var. patriciae Pócs, Candollea 56 (1): 72, 2001 (
*** Lejeunea albescens (Steph.) Mizut., J. Hattori Bot. Lab. 33: 245, 1970 (
* Lejeunea albiflora Colenso, Trans. & Proc. New Zealand Inst. 21: 72, 1889 (
** Lejeunea aloba Sande Lac., Plagiochila Sandei: 10, 1856 (
** Lejeunea alobifolia H.A.Mill., Phytologia 47 (4): 323, 1981 (
** Lejeunea amaniensis E.W.Jones, J. Bryol. 13 (3): 392, 1985 (
** Lejeunea ambigua Lindenb. et Gottsche, Syn. Hepat. 5: 764, 1847 (
* Lejeunea amentulifera Steph., Sp. Hepat. (Stephani) 5: 707, 1915 (
** Lejeunea androgyna R.M.Schust., Phytologia 45 (5): 432, 1980 (
* Lejeunea angulifolia Mitt., Philos. Trans. 168: 400, 1879 (
*** Lejeunea angusta (Lehm. et Lindenb.) Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 469, 1842 (
** Lejeunea anisophylla Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 263, 1843 (
*** Lejeunea anomala Lindenb. et Gottsche, Linnaea 24 (6): 636, 1851 [1852] (
* Lejeunea antillana Steph., Hedwigia 27 (11/12): 281, 1888 (
** Lejeunea aphanes Spruce, J. Bot. 19: 36, 1881 (
*** Lejeunea apiculata Sande Lac., Ned. Kruidk. Arch. 3: 421, 1854 [1855] (
** Lejeunea aquatica Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 5 (4): 643, 1929 [1930] (
** Lejeunea aquatica var. apiculata S.Hatt., Bull. Tokyo Sci. Mus. 11: 108, 1944 (
** Lejeunea armitii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 768, 1915 (
* Lejeunea asperifolia Steph., Sp. Hepat. (Stephani) 5: 708, 1915 (
*** Lejeunea asperrima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 160, 1884 (
** Lejeunea asperula (Steph.) Mizut., J. Hattori Bot. Lab. 33: 236, 1970 (
** Lejeunea asprella Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 175, 1884 (
* Lejeunea atheatostipa Spruce, J. Bot. 33: 83, 1895 (
** Lejeunea barbata (Herzog) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 376, 2011 (
*** Lejeunea bermudiana (A.Evans) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1105, 1980 (
** Lejeunea bidentula Herzog, Symb. Sin. 5: 51, 1930 (
** Lejeunea biformis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 162 (68), 1864 (
** Lejeunea blepharogona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 178, 1884 (
** Lejeunea blomquistii R.M.Schust., J. Elisha Mitchell Sci. Soc. 78 (1): 64, 1962 (
*** Lejeunea boliviensis (Steph.) R.L.Zhu et M.E.Reiner, Bryologist 107 (2): 237, 2004 (
** Lejeunea boryana Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838 (
** Lejeunea brenanii E.W.Jones, J. Bryol. 10 (4): 391, 1979 (
*** Lejeunea calcicola R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 404, 1957 (
** Lejeunea calcicola var. mexicana R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 408, 1957 (
** Lejeunea canariensis (Steph.) Steph., Sp. Hepat. (Stephani) 5: 802, 1915 (
*** Lejeunea cancellata Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 472, 1842 (
** Lejeunea cantabrigiensis E.W.Jones, J. Bryol. 15 (4): 669, 1989 (
*** Lejeunea capensis Gottsche, Syn. Hepat. 3: 374, 1845 (
* Lejeunea caroliniana Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876 (
*** Lejeunea catinulifera Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 233, 1884 (
*** Lejeunea caulicalyx (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 13, 2000 (
*** Lejeunea cavifolia (Ehrh.) Lindb., Revis. crit. icon.: 43, 1871 (
** Lejeunea caviloba (Steph.) Besch., J. Bot. (Morot) 12: 140, 1898 (
*** Lejeunea cerina (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 391, 1845 (
** Lejeunea chaishanensis S.H.Lin, Yushania 9: 7, 1992 (
** Lejeunea cladogyna A.Evans, Amer. J. Bot. 5 (3): 134, 1918 (
** Lejeunea clavata Lindenb., Syn. Hepat. 3: 379, 1845 (
** Lejeunea claviformis Lindenb. ex Steph., Sp. Hepat. (Stephani) 5: 727, 1915 (
** Lejeunea cochleata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 273, 1884 (
** Lejeunea cocoes Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 114, 1860 [1861] (
*** Lejeunea colensoana (Steph.) M.A.M.Renner, Austral. Syst. Bot. 23 (6): 455, 2010 (
** Lejeunea compacta (Steph.) Steph., Sp. Hepat. (Stephani) 5: 771, 1915 (
** Lejeunea concinnula Spruce et Steph., J. Bot. 25: 39, 1887 (
** Lejeunea connatistipula (Steph.) Steph., Sp. Hepat. (Stephani) 5: 772, 1915 (
** Lejeunea contracta Mizut., J. Hattori Bot. Lab. 33: 248, 1970 (
** Lejeunea controversa Gottsche, Hepat. Eur., Leberm. 56-57: no 556, 1873 (
** Lejeunea convexiloba M.L.So et R.L.Zhu, Bryologist 101 (1): 137, 1998 (
** Lejeunea corcovadae (Steph.) Bischl., Nova Hedwigia 5 (1/2): 406, 1963 (
** Lejeunea cordiflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 283, 1884 (
** Lejeunea corralensis A.Evans, Ann. Bryol. 3: 86, 1930 (
*** Lejeunea corynantha Spruce, J. Linn. Soc., Bot. 30 (210): 344, 1895 (
** Lejeunea crassiretis Mitt., Fl. vit.: 414, 1871 [1873] (
*** Lejeunea cristulata (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 21, 2000 (
*** Lejeunea cristuliflora (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 19, 2000 (
** Lejeunea cuspidistipula (Steph.) Steph. ex Watts, Proc. Linn. Soc. New South Wales (ser. 2) 27 (108): 493, 1903 (
** Lejeunea cyanomontana R.M.Schust., Phytologia 45 (5): 432, 1980 (
** Lejeunea cyanophora R.M.Schust., J. Hattori Bot. Lab. 26: 246, 1963 (
** Lejeunea cyathearum E.W.Jones, J. Bryol. 8 (1): 86, 1974 (
*** Lejeunea cyathophora Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 359, 1851 (
** Lejeunea denticalyx (Steph.) Steph., Sp. Hepat. (Stephani) 5: 794, 1915 (
** Lejeunea denticuspis (Steph.) Mizut., J. Hattori Bot. Lab. 36: 160, 1972 [1973] (
** Lejeunea denudata (Pearson) J.J.Engel, Bryologist 78 (3): 361, 1975 (
*** Lejeunea deplanata Nees, Syn. Hepat. 3: 368, 1845 (
** Lejeunea deplanata var. cuspidata (Steph.) M.E.Reiner, Nova Hedwigia 91 (3/4): 529, 2010 (
** Lejeunea diaphana Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 168, 1884 (
** Lejeunea dimorpha T.Kodama, J. Hattori Bot. Lab. 41: 384, 1976 (
** Lejeunea dipterocarpa E.W.Jones, J. Bryol. 7 (1): 44, 1972 (
*** Lejeunea discreta Lindenb., Syn. Hepat. 3: 361, 1845 (
* Lejeunea disjecta Spruce, J. Linn. Soc., Bot. 30 (210): 347, 1895 (
** Lejeunea diversicuspis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 176, 1884 (
** Lejeunea drummondii Taylor, London J. Bot. 5: 400, 1846 (
** Lejeunea ecarinata (Steph.) J.M.Coult., Barnes et Arthur, Bot. Gaz. 15 (12): 349, 1890 (
** Lejeunea eckloniana Lindenb., Syn. Hepat. 3: 381, 1845 (
** Lejeunea eifrigii Mizut., J. Hattori Bot. Lab. 33: 244, 1970 (
* Lejeunea elongella Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 161, 1864 (
*** Lejeunea erostrata M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 25, 2000 (
*** Lejeunea exilis (Reinw., Blume et Nees) Grolle, J. Hattori Bot. Lab. 46: 353, 1979 (
** Lejeunea exilis var. abnormis (Herzog) G.E.Lee, Polish Bot. J. 58 (1): 61, 2013 (
** Lejeunea fernandeziana S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 16, 1957 (
** Lejeunea firma Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 112, 1860 [1861] (
** Lejeunea fissistipula (Steph.) Steph., Sp. Hepat. (Stephani) 5: 775, 1915 (
** Lejeunea flagellaris Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 273, 1884 (
*** Lejeunea flava (Sw.) Nees, Naturgesch. Eur. Leberm. 3: 277, 1838 (
** Lejeunea flava subsp. moorei (Lindb.) R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 161, 1957 (
** Lejeunea flava subsp. orientalis R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 161, 1957 (
** Lejeunea flava var. pellucida Lindenb. et Gottsche, Linnaea 24 (6): 634, 1851 [1852] (
** Lejeunea flava subsp. tabularis (Spreng.) S.W.Arnell, Hepat. South Africa: 199, 1963 (
*** Lejeunea flavovirens Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 144, 1873 (
** Lejeunea fleischeri (Steph.) Mizut., J. Hattori Bot. Lab. 33: 238, 1970 (
** Lejeunea floridana A.Evans, Bull. Torrey Bot. Club 32 (4): 185, 1905 (
*** Lejeunea fulfordiae (Jovet-Ast) R.L.Zhu, Syst. Bot. 33 (4): 617, 2008 (
** Lejeunea fusagasugana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 158, 1864 (
** Lejeunea galeata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 172, 1884 (
** Lejeunea gayana Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864 (
** Lejeunea gibbiloba (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (
*** Lejeunea glaucescens Gottsche, Syn. Hepat. 3: 378, 1845 (
** Lejeunea glaucescens var. acrogyna R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 400, 1957 (
* Lejeunea glaucescens var. obsoleta R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 395, 1957 (
** Lejeunea globosiflora (Steph.) Steph., Sp. Hepat. (Stephani) 5: 795, 1915 (
** Lejeunea gomphocalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 174, 1884 (
** Lejeunea gracilicaulis Spruce, Mem. Torrey Bot. Club 1 (3): 126, 1890 (
** Lejeunea gracilipes (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 213, 1884 (
** Lejeunea gracilis Steph., Sp. Hepat. (Stephani) 5: 777, 1915 (
** Lejeunea gradsteiniana Pócs, Acta Biol. Pl. Agr. 1: 55, 2010 [2011] (
** Lejeunea gradsteinii G.E.Lee, Damanhuri et Latiff, Acta Biol. Pl. Agr. 1: 29, 2010 [2011] (
* Lejeunea grossecristata (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 5 (3): 556, 1968 (
*** Lejeunea grossiretis (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 27, 2000 (
** Lejeunea grossistipula Steph., Sp. Hepat. (Stephani) 5: 739, 1915 (
*** Lejeunea grossitexta (Steph.) M.E.Reiner et Goda, J. Hattori Bot. Lab. 89: 29, 2000 (
* Lejeunea grossiuscula Gottsche ex Steph., Sp. Hepat. (Stephani) 5: 739, 1915 (
** Lejeunea hahnii Solari, J. Hattori Bot. Lab. 54: 543, 1983 (
* Lejeunea haitica Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 263, 1843 (
** Lejeunea hawaikiana M.A.M.Renner et de Lange, New Zealand J. Bot. 49 (3): 431, 2011 (
** Lejeunea helenae Pearson, Forh. Vidensk.-Selsk. Kristiania 1886 (3): 6, 1886 (
** Lejeunea helmsiana (Steph.) Steph., Sp. Hepat. (Stephani) 5: 796, 1915 (
*** Lejeunea hepaticola (Steph.) Steph., Sp. Hepat. (Stephani) 5: 714, 1915 (
** Lejeunea hibernica Bischl., H.A.Mill. et Bonner ex Grolle, Lindbergia 3 (1/2): 48, 1975 [1976] (
** Lejeunea hodgsoniana Grolle ex R.J.Lewington, Bever. et M.A.M.Renner, PhytoKeys 29: 2, 2013 (
** Lejeunea howeana Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 31 (2): 219, 1982 (
** Lejeunea hui R.L.Zhu, Beih. Nova Hedwigia 121: 134, 2001 (
** Lejeunea humefacta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 275, 1884 (
** Lejeunea hyalina (Steph.) L.Söderstr. et A.Hagborg, Phytotaxa 220 (2): 188, 2015 (
** Lejeunea ibadana A.J.Harr. et E.W.Jones, J. Bryol. 12 (1): 40, 1982 (
** Lejeunea immersa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 186, 1884 (
** Lejeunea increscens Spruce, Mem. Torrey Bot. Club 1 (3): 124, 1890 (
* Lejeunea inflatiloba (Steph.) H.A.Mill., Bonner et Bischl., Nova Hedwigia 14 (1): 67, 1967 (
*** Lejeunea inflexiloba Prantl, Hedwigia 31: xvi, 1892 (
*** Lejeunea intricata Prantl, Hedwigia 31: xvi, 1892 (
*** Lejeunea isophylla E.W.Jones, Trans. Brit. Bryol. Soc. 5 (3): 559, 1968 (
** Lejeunea japonica Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 203, 1891 (
** Lejeunea jardinii Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: ccv, 1889 [1890] (
** Lejeunea julacea Steph., Sp. Hepat. (Stephani) 5: 715, 1915 (
*** Lejeunea jungneri (Steph.) Steph., Cat. Afr. Pl. (Hiern) 2 (2): 318, 1901 (
** Lejeunea kashyapii M.Dey, D.K.Singh et D.Singh, J. Bryol. 30 (2): 126, 2008 (
* Lejeunea kilimandjarensis Steph., Sp. Hepat. (Stephani) 5: 716, 1915 (
** Lejeunea kinabalensis Mizut., J. Hattori Bot. Lab. 33: 246, 1970 (
** Lejeunea kodamae Ikegami et Inoue, J. Jap. Bot. 36 (1): 7, 1961 (
** Lejeunea konosensis Mizut., J. Hattori Bot. Lab. 71: 127, 1992 (
** Lejeunea kuerschneriana Pócs, Beih. Nova Hedwigia 138: 100, 2010 (
*** Lejeunea laeta (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 380, 1845 (
** Lejeunea laii R.L.Zhu, J. Bryol. 30 (2): 173, 2008 (
*** Lejeunea lamacerina (Steph.) Schiffn., Hedwigia 41 (5): 278, 1902 (
** Lejeunea lamacerina subsp. gemminata R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (1): 168, 1957 (
** Lejeunea latilobula (Herzog) R.L.Zhu et M.L.So, J. Bryol. 24 (2): 168, 2002 (
** Lejeunea leptalea Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 272, 1884 (
* Lejeunea leratii (Steph.) Mizut., J. Hattori Bot. Lab. 33: 243, 1970 (
** Lejeunea leucosis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxviii, 1889 [1890] (
* Lejeunea litoralis Steph., Sp. Hepat. (Stephani) 5: 778, 1915 (
** Lejeunea lomana E.W.Jones, Bull. Brit. Mus. (Nat. Hist.), Bot. 11 (3): 257, 1983 (
** Lejeunea longicollis S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 12, 1958 (
** Lejeunea longilobula Pócs, Beih. Nova Hedwigia 138: 112, 2010 (
** Lejeunea lowriana Steph., Sp. Hepat. (Stephani) 5: 779, 1915 (
*** Lejeunea lumbricoides (Nees) Nees, Syn. Hepat. 3: 342, 1845 (
** Lejeunea lunatigastria Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 98, 1974 (
** Lejeunea lyratiflora Prantl, Hedwigia 31: xvi, 1892 (
** Lejeunea magohukui Mizut., Misc. Bryol. Lichenol. 7 (7): 133, 1977 (
** Lejeunea mandonii (Steph.) Müll.Frib., Leberm. Eur. 2 (9): 1281, 1958 (
*** Lejeunea masoalae Pócs, Beih. Nova Hedwigia 138: 103, 2010 (
** Lejeunea massalongoana (Schiffn. ex P.Syd.) Solari, J. Hattori Bot. Lab. 54: 542, 1983 (
** Lejeunea megalantha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 172, 1884 (
** Lejeunea mehrana M.Dey, D.K.Singh et D.Singh, J. Bryol. 30 (2): 128, 2008 (
*** Lejeunea meridensis Ilk.-Borg., Nova Hedwigia 80 (1/2): 59, 2005 (
** Lejeunea micholitzii Mizut., J. Hattori Bot. Lab. 33: 236, 1970 (
** Lejeunea microloba Taylor, London J. Bot. 5: 399, 1846 (
*** Lejeunea mimula Hürl., Bauhinia 11 (1): 12, 1993 (
*** Lejeunea minutiloba A.Evans, Bull. Torrey Bot. Club 44 (11): 525, 1917 (
** Lejeunea minutiloba var. heterogyna R.M.Schust., J. Elisha Mitchell Sci. Soc. 73 (2): 425, 1957 (
** Lejeunea mizutanii Grolle, J. Hattori Bot. Lab. 45: 178, 1979 (
** Lejeunea molkenboeriana Sande Lac., Ned. Kruidk. Arch. 3: 421, 1854 [1855] (
*** Lejeunea monimiae (Steph.) Steph., Sp. Hepat. (Stephani) 5: 747, 1915 (
*** Lejeunea multidentata M.E.Reiner et Mustelier, J. Bryol. 26 (2): 103, 2004 (Reiner-Drehwald and Mustelier 2004).
* Lejeunea musae (Spreng.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845 (
** Lejeunea muscicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 281, 1884 (
* Lejeunea muscicola var. palmicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884 (
** Lejeunea neelgherriana Gottsche, Syn. Hepat. 3: 354, 1845 (
** Lejeunea nemoralis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 159 (65), 1864 (
** Lejeunea nepalensis Steph., Sp. Hepat. (Stephani) 5: 780, 1915 (
* Lejeunea nesiotica Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxviii, 1889 [1890] (
** Lejeunea neumanniana Nees, Repert. Pharm. 76: 44, 1842 (
** Lejeunea nietneri (Steph.) Steph. ex Watts, Proc. Linn. Soc. New South Wales (ser. 2) 26 (104): 633, 1902 (
** Lejeunea notata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 361, 1882 (
** Lejeunea nymannii Steph., Sp. Hepat. (Stephani) 5: 781, 1915 (
** Lejeunea obfusca Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 114, 1860 [1861] (
** Lejeunea obidensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 277, 1884 (
* Lejeunea obscura Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 112, 1860 [1861] (
** Lejeunea obtusata Gottsche, Abh. Naturwiss. Vereins Bremen 7: 354, 1882 (
** Lejeunea okomuensis E.W.Jones, Trans. Brit. Bryol. Soc. 5 (4): 787, 1969 (
*** Lejeunea oligoclada Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcix, 1889 [1890] (
*** Lejeunea oracola M.A.M.Renner, Austral. Syst. Bot. 23 (6): 448, 2010 (
** Lejeunea osculatiana De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 16: 233, 1857 (
** Lejeunea otiana S.Hatt., Bot. Mag. (Tokyo) 65 (763/764): 15, 1952 (
** Lejeunea ovalifolia Steph., Sp. Hepat. (Stephani) 5: 751, 1915 (
** Lejeunea pacifica Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 262, 1843 (
** Lejeunea pallescens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 360, 1851 (
** Lejeunea pallida Lindenb. et Gottsche, Syn. Hepat. 5: 762, 1847 (
* Lejeunea pallidissima Gola, Nuovo Giorn. Bot. Ital. (n.ser.) 27 (2/4): 247, 1920 (
*** Lejeunea papilionacea Prantl, Hedwigia 31: xvii, 1892 (
* Lejeunea paratropa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 176, 1884 (
** Lejeunea patagonica (Steph.) Steph., Sp. Hepat. (Stephani) 5: 797, 1915 (
** Lejeunea patens Lindb., Acta Soc. Sci. Fenn. 10: 482, 1875 (
*** Lejeunea patersonii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 784, 1915 (
*** Lejeunea patriciae Schäf.-Verw., Candollea 56 (1): 64, 2001 (
*** Lejeunea paucidentata (Steph.) Grolle, J. Hattori Bot. Lab. 69: 191, 1991 (
*** Lejeunea pectinella Mizut., J. Hattori Bot. Lab. 33: 239, 1970 (
** Lejeunea perigonialis Gottsche, Mexik. Leverm.: 223, 1863 (
*** Lejeunea perpapillosa M.E.Reiner et K.C.Pôrto, Nova Hedwigia 85 (3/4): 542, 2007 (Reiner-Drehwald and Pôrto 2007).
* Lejeunea pertusa (Corda ex Nees et Mont.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845 (
*** Lejeunea phyllobola Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 471, 1842 (
* Lejeunea phyllobola var. turgidula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 259, 1884 (
*** Lejeunea planiloba A.Evans, Proc. Wash. Acad. Sci. 8: 147, 1906 (
** Lejeunea polilloensis Steph., Sp. Hepat. (Stephani) 5: 786, 1915 (
** Lejeunea praetervisa Steph., Sp. Hepat. (Stephani) 5: 752, 1915 (
** Lejeunea primordialis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 375, 1845 (
** Lejeunea princeps (Steph.) Mizut., J. Hattori Bot. Lab. 34: 454, 1971 (
** Lejeunea procumbens Mitt., Fl. vit.: 413, 1871 [1873] (
** Lejeunea propagulifera Gradst., Phytotaxa 9: 54, 2010 (
** Lejeunea pteridis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxvii, 1889 [1890] (
*** Lejeunea ptosimophylla C.Massal., Nuovo Giorn. Bot. Ital. 13 (2): 123, 1881 (
*** Lejeunea puiggariana Steph., Sp. Hepat. (Stephani) 5: 754, 1915 (
*** Lejeunea pulverulenta (Steph.) M.E.Reiner, Cryptog. Bryol. 26 (1): 60, 2005 (
*** Lejeunea pulvinata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 61, 1836 (
*** Lejeunea raddiana Lindenb., Syn. Hepat. 3: 342, 1845 (
** Lejeunea radicans Lindenb. et Gottsche, Syn. Hepat. 5: 766, 1847 (
** Lejeunea ramosissima Steph., Bot. Jahrb. Syst. 8 (2): 88, 1886 (
** Lejeunea rara Steph., Sp. Hepat. (Stephani) 5: 798, 1915 (
* Lejeunea ravenelii Austin, Bot. Bull. (Hanover) 1 (8): 35, 1876 (
** Lejeunea recurva M.E.Reiner, Polish Bot. J. 58 (2): 423, 2013 (
*** Lejeunea reflexistipula (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 3: 335, 1845 (
** Lejeunea reflexistipula var. costaricensis (Steph.) M.E.Reiner, Nova Hedwigia 81 (3/4): 408, 2005 (
*** Lejeunea reinerae Ilk.-Borg., Nova Hedwigia 80 (1/2): 61, 2005 (
** Lejeunea resupinata (Steph.) Steph., Sp. Hepat. (Stephani) 5: 757, 1915 (
** Lejeunea reticulata Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 747, 1942 (
*** Lejeunea rhigophila M.A.M.Renner, Austral. Syst. Bot. 23 (6): 453, 2010 (
** Lejeunea rhodesiae (Sim) R.M.Schust., J. Hattori Bot. Lab. 25: 71, 1962 (
*** Lejeunea rionegrensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 579, 1885 (
** Lejeunea riparia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861] (
* Lejeunea rothii (Schwägr.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 407, 1845 (
*** Lejeunea rotundifolia Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 359, 1851 (
** Lejeunea sanctae-helenae M.Wigginton, J. Bryol. 29 (1): 12, 2007 (
** Lejeunea schusteri Grolle, Haussknechtia 8: 60, 2001 (
** Lejeunea semiscabrida Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 154 (60), 1864 (
** Lejeunea semperi Steph., Sp. Hepat. (Stephani) 5: 788, 1915 (
** Lejeunea seriata Lindenb. et Gottsche, Syn. Hepat. 5: 762, 1847 (
** Lejeunea sessiliflora (Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 37: 171, 1988 (
** Lejeunea setacea (Steph.) Steph., Sp. Hepat. (Stephani) 5: 719, 1914 (
** Lejeunea setiloba Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 281, 1884 (
** Lejeunea sharpii (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 991, 1980 (
** Lejeunea siccata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 284, 1884 (
** Lejeunea silvatica Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 159 (65), 1864 (
** Lejeunea smaragdina Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxii, 1889 [1890] (
*** Lejeunea soae R.L.Zhu, Y.M.Wei, L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 81 (1): 1, 2013 (
** Lejeunea solanicola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 280, 1884 (
*** Lejeunea sordida (Nees) Nees, Naturgesch. Eur. Leberm. 3: 278, 1838 (
*** Lejeunea spiniloba Lindenb. et Gottsche, Syn. Hepat. 5: 770, 1847 (
** Lejeunea spinuliflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 177, 1884 (
** Lejeunea sporadica Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxx, 1889 [1890] (
* Lejeunea squarrosa (Steph.) Steph., Sp. Hepat. (Stephani) 5: 719, 1915 (
** Lejeunea squarrosula (Herzog) Solari, J. Hattori Bot. Lab. 54: 543, 1983 (
* Lejeunea stephaniana Mizut., J. Hattori Bot. Lab. 27: 143, 1964 (
** Lejeunea stevensiana (Steph.) Mizut., J. Hattori Bot. Lab. 34: 452, 1971 (
** Lejeunea subacuta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861] (
* Lejeunea subaquatica Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 397, 1955 (
* Lejeunea subbifida Steph. ex Duss, Enum. musc. Antilles franç., Hép.: 10, 1903 (
** Lejeunea subigiensis (Steph.) Steph., Hedwigia 35 (3): 94, 1896 (
** Lejeunea subrufula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 289, 1884 (
** Lejeunea subsessilis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884 (
*** Lejeunea subspathulata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 173, 1884 (
** Lejeunea succulenta Herzog, Rev. Bryol. Lichénol. 20 (1/2): 171, 1951 [1952] (
** Lejeunea suffruticola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 279, 1884 (
** Lejeunea syoshii Inoue, Bull. Natl. Sci. Mus. Tokyo, B 3 (4): 143, 1977 (
*** Lejeunea talamancensis M.E.Reiner et Schäf.-Verw., Nova Hedwigia 87 (3/4): 414, 2008 (
*** Lejeunea tamasii M.E.Reiner, N.Salazar et C.Chung, Polish Bot. J. 58 (2): 420, 2013 (
** Lejeunea tamaspocsii G.E.Lee, Polish Bot. J. 58 (1): 66, 2013 (
*** Lejeunea tapajosensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 223, 1884 (
** Lejeunea tarapotensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 282, 1884 (
** Lejeunea tenella Taylor, London J. Bot. 5: 398, 1846 (
** Lejeunea thallophora (Eifrig) Gradst., Phytotaxa 9: 54, 2010 (
*** Lejeunea tonduzana (Steph.) M.E.Reiner, Polish Bot. J. 58 (2): 421, 2013 (
** Lejeunea trachygona Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 175, 1884 (
*** Lejeunea trinitensis Lindenb., Syn. Hepat. 3: 381, 1845 (
** Lejeunea trukensis H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 56, 1963 (
** Lejeunea tuberculosa Steph., Sp. Hepat. (Stephani) 5: 790, 1915 (
** Lejeunea tumida Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 157, 1854 (
** Lejeunea uleana (Steph.) Steph., Sp. Hepat. (Stephani) 5: 765, 1915 (
*** Lejeunea umbilicata (Nees) Nees, Observ. bot.: 472, 1843 (
** Lejeunea utriculata (Steph.) Mizut., J. Hattori Bot. Lab. 33: 242, 1970 (
** Lejeunea vesicata Mitt., Fl. vit.: 415, 1871 [1873] (
*** Lejeunea villaumei (Steph.) Grolle, J. Bryol. 9 (4): 536, 1977 [1978] (
** Lejeunea vojtkoi Pócs, Beih. Nova Hedwigia 138: 107, 2010 (
** Lejeunea vulgariformis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 355, 1882 (
** Lejeunea wattsiana (Steph.) H.A.Mill., Bonner et Bischl., Nova Hedwigia 14 (1): 66, 1967 (
* Lejeunea wichurae (Steph.) Steph., Sp. Hepat. (Stephani) 5: 792, 1915 (
** Lejeunea wightii Lindenb., Syn. Hepat. 3: 379, 1845 (
*** Lejeunea xiphophylla (Herzog) M.E.Reiner, Nova Hedwigia 87 (3/4): 409, 2008 (
** subg. Nanolejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1092, 1980 (
** Lejeunea curviloba Steph., Sp. Hepat. (Stephani) 5: 774, 1915 (
*** Lejeunea laetevirens Nees et Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 469, 1842 (
** Lejeunea pallidevirens S.Hatt., J. Hattori Bot. Lab. 12: 80, 1954 (
** Lejeunea parva (S.Hatt.) Mizut., Misc. Bryol. Lichenol. 5 (10/12): 178, 1971 (
*** Lejeunea prionoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 238, 1884 (
*** Lejeunea ramulosa Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 274, 1884 (
*** subg. Neopotamolejeunea (M.E.Reiner) Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 487, 2007 (
** Lejeunea juruana Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 488, 2007 (
*** Lejeunea polyantha Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 350, 1856 (
** Lejeunea tenera (Sw.) Gottsche, Lindenb. et Nees, Syn. Hepat. 3: 406, 1845 (
*** Lejeunea topoensis Gradst. et M.E.Reiner, Syst. Bot. 32 (3): 488, 2007 (
** subg. Papillolejeunea (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (
** Lejeunea balazsii (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (
** Lejeunea candida (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (
** Lejeunea falcata (Pócs et J.Eggers) Pócs, Phytotaxa 208 (1): 99, 2015 (
** Lejeunea koponenii (Pócs et J.Eggers) Pócs, Phytotaxa 208 (1): 99, 2015 (
** Lejeunea papuana (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (
** Lejeunea touwii (Pócs) R.M.Schust., J. Hattori Bot. Lab. 85: 84, 1998 (
Incertae sedis
* Lejeunea apiahyna (Steph.) Sushil K.Singh, Phytotaxa 96 (1): 63, 2013 (
* Lejeunea aptycta Gottsche, Syn. Hepat. 3: 369, 1845 (
** Lejeunea asthenica Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 222, 1884 (
* Lejeunea bethanica Gottsche, Syn. Hepat. 3: 381, 1845 (Gottsche et al. 1945b)
** Lejeunea bombonasensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 222, 1884 (
** Lejeunea bornmuelleri (Steph.) M.E.Reiner, Nova Hedwigia 95 (3/4): 471, 2012 (
** Lejeunea caracensis Lindenb., Syn. Hepat. 3: 355, 1845 (
** Lejeunea caripensis Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847 (
** Lejeunea chamissonis Lindenb., Syn. Hepat. 3: 378, 1845 (
** Lejeunea chimborazensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 215, 1884 (
*** Lejeunea combuensis O.S.Moura, Ilk.-Borg. et M.E.Reiner, Nova Hedwigia 95 (1/2): 198, 2012 (
** Lejeunea concava Lindenb. et Gottsche, Syn. Hepat. 5: 759, 1847 (
*** Lejeunea conformis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 260, 1843 (
** Lejeunea cordistipula Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847 (
** Lejeunea cyrtotis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 229, 1884 (
*** Lejeunea debilis (Lehm. et Lindenb.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 60, 1836 (
** Lejeunea devendrae (Sushil K.Singh) P.K.Verma et K.K.Rawat, J. Bryol. 36 (2): 161, 2014 (
** Lejeunea dictyocalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 218, 1884 (
** Lejeunea dipterota (Eifrig) G.E.Lee, Polish Bot. J. 58 (1): 61, 2013 (
*** Lejeunea drehwaldii Heinrichs et Schäf.-Verw., Phytotaxa 69: 14, 2012 (
** Lejeunea duncaniae (Sim) M.E.Reiner, Phytotaxa 208 (1): 98, 2015 (
** Lejeunea edentata L.Söderstr., Phytotaxa 208 (1): 98, 2015 (
* Lejeunea emarginuliflora Gottsche ex Steph., Sp. Hepat. (Stephani) 5: 734, 1915 (
* Lejeunea epibrya Taylor, London J. Bot. 7: 199, 1848 (
** Lejeunea estrellamontana M.A.M.Renner et Pócs, Phytotaxa 81 (1): 9, 2013 (
* Lejeunea fawcettiae D.J.Carr, Proc. Roy. Soc. Victoria 117 (2): 325, 2005 (
** Lejeunea flaccida Lindenb. et Gottsche, Syn. Hepat. 5: 758, 1847 (
** Lejeunea flavida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 113, 1860 [1861] (
** Lejeunea florida Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 221, 1884 (
** Lejeunea fulva Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 237, 1884 (
** Lejeunea gottscheana Lindenb., Syn. Hepat. 3: 382, 1845 (
** Lejeunea graminicolor Spruce, J. Linn. Soc., Bot. 30 (210): 343, 1895 (
*** Lejeunea grolleana (Bernecker) R.L.Zhu et W.Ye, J. Syst. Evol. 51 (4): 472, 2013 (
*** Lejeunea herminieri (Steph.) R.L.Zhu, Phytotaxa 208 (1): 99, 2015 (
** Lejeunea heterocheila Taylor, London J. Bot. 5: 394, 1846 (
* Lejeunea hygrophila Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864 (
** Lejeunea laevicalyx Gottsche, Mexik. Leverm.: 221, 1863 (
* Lejeunea laxa (Nees) Lindenb., Syn. Hepat. 3: 378, 1845 (
** Lejeunea leiantha Spruce, J. Linn. Soc., Bot. 30 (210): 345, 1895 (
** Lejeunea leptoscypha Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: ccv, 1889 [1890] (
*** Lejeunea lusoria (Lindenb. et Gottsche) Steph., Hedwigia 29 (3): 141, 1890 (
** Lejeunea luzonensis (Steph.) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 376, 2011 (
** Lejeunea macrorhyncha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 220, 1884 (
** Lejeunea malangensis (Herzog) R.L.Zhu et Y.M.Wei, J. Bryol. 34 (4): 319, 2012 (
** Lejeunea marasmodes Spruce, Mem. Torrey Bot. Club 1 (3): 125, 1890 (
** Lejeunea morobensis (Grolle) M.A.M.Renner et Pócs, Phytotaxa 81 (1): 9, 2013 (
** Lejeunea novoguineensis Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 238, 1893 (
*** Lejeunea obtusangula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 221, 1884 (
* Lejeunea oerstediana Lindenb. et Hampe, Linnaea 24 (6): 641, 1851 [1852] (
** Lejeunea paraensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 224, 1884 (
** Lejeunea parviloba Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 87, 1876 [1877] (
* Lejeunea pfleidereri Sushil K.Singh, Phytotaxa 96 (1): 63, 2013 (
* Lejeunea prominula Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 157, 1864 (
*** Lejeunea pterigonia (Lehm. et Lindenb.) Mont., Ann. Sci. Nat. Bot. (sér. 2) 14: 337, 1840 (
** Lejeunea quinqueumbonata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 230, 1884 (
** Lejeunea quinqueumbonata var. rotundata (Herzog) Sushil K.Singh, Phytotaxa 96 (1): 64, 2013 (
** Lejeunea remotifolia Hampe ex Steph., Sp. Hepat. (Stephani) 5: 756, 1915 (
*** Lejeunea ruthii (A.Evans) R.M.Schust., J. Hattori Bot. Lab. 25: 23, 1962 (
** Lejeunea ruthii var. alata R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 1063, 1980 (
** Lejeunea sikorae (Steph.) Steph., Bull. Soc. Roy. Bot. Belgique, Mém. 32: 120, 1893 [1894] (
** Lejeunea srivastavae P.K.Verma et K.K.Rawat, Taiwania 58 (1): 8, 2013 (
** Lejeunea stenodentata M.A.M.Renner et Pócs, Phytotaxa 81 (1): 8, 2013 (
*** Lejeunea subelobata Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1039, 1888 (
** Lejeunea subolivacea Mizut., J. Hattori Bot. Lab. 28: 121, 1965 (
** Lejeunea subplana (Steph.) C.J.Bastos, J. Bryol. 36 (3): 249, 2014 (
*** Lejeunea sulphurea (Lehm. et Lindenb.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 217, 1884 (
** Lejeunea terricola Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxci, 1889 [1890] (
** Lejeunea urbanii (Steph.) Steph., Sp. Hepat. (Stephani) 5: 766, 1915 (
** Lejeunea venezuelana (R.M.Schust.) R.L.Zhu et W.Ye, J. Syst. Evol. 51 (4): 473, 2013 (
** Lejeunea viridis R.M.Schust. ex L.Söderstr. et A.Hagborg, Phytotaxa 208 (1): 99, 2015 (
* Lejeunea zacuapana (Steph.) Prantl, Hedwigia 29: xviii, 1890 (
Excluded from the genus
* Lejeunea elegans Gottsche, Syn. Hepat. 3: 364, 1845 (
* Lejeunea hieronymii Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cciii, 1889 [1890] (
* Lejeunea proboscidea Gottsche, Mexik. Leverm.: 225, 1863 (
* Lejeunea scabriflora Loitl., Diagn. pl. nov.: 22, 1894 (
* Lejeunea trochantha Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcii, 1889 [1890] (
** Microlejeunea (Spruce) Steph., Hedwigia 27 (2): 61, 1888 (
*** Microlejeunea acutifolia Steph., Hedwigia 35 (3): 113, 1896 (
** Microlejeunea africana Steph., Hedwigia 27 (2): 61, 1888 (
** Microlejeunea aligera (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 827, 1915 (
** Microlejeunea ankasica E.W.Jones, J. Bryol. 10 (4): 394, 1979 (
** Microlejeunea aphanella (Spruce) Steph., Sp. Hepat. (Stephani) 5: 816, 1915 (
** Microlejeunea atsuana Steph., Hedwigia 35 (3): 113, 1896 (
** Microlejeunea bischlerae (B.M.Thiers) B.M.Thiers, Phytotaxa 65: 59, 2012 (
*** Microlejeunea bullata (Taylor) Steph., Hedwigia 29 (2): 90, 1890 (
* Microlejeunea byssoides (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892 (
** Microlejeunea capillaris (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 819, 1915 (
** Microlejeunea cochlearifolia Steph., Hedwigia 27 (3/4): 113, 1888 (
** Microlejeunea colombiana Bischl., Nova Hedwigia 5 (1/2): 373, 1963 (
** Microlejeunea constricta (Grolle) Grolle, J. Bryol. 21 (1): 41, 1999 (
** Microlejeunea crenulifolia (Gottsche) Steph., Hedwigia 35 (3): 114, 1896 (
*** Microlejeunea cystifera Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 68, 1950 (
** Microlejeunea dispar Jovet-Ast, Rev. Bryol. Lichénol. 27 (3/4): 191, 1959 (
*** Microlejeunea epiphylla Bischl., Nova Hedwigia 5 (1/2): 378, 1963 (
*** Microlejeunea filicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013 (
*** Microlejeunea fischeri (Tixier) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013 (
** Microlejeunea fissistipula Steph., Sp. Hepat. (Stephani) 5: 810, 1915 (
*** Microlejeunea globosa (Spruce) Steph., Sp. Hepat. (Stephani) 5: 821, 1915 (
** Microlejeunea herzogiana Steph., Biblioth. Bot. 87 (2): 266, 1916 (
** Microlejeunea indica (Udar et U.S.Awasthi) Y.M.Wei et R.L.Zhu, Phytotaxa 97 (2): 63, 2013 (
** Microlejeunea inflata Steph., Sp. Hepat. (Stephani) 5: 811, 1915 (
** Microlejeunea kinabaluensis (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999 (
*** Microlejeunea latitans (Hook.f. et Taylor) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 52, 2013 (
** Microlejeunea lunulatiloba Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 27, 1931 (
* Microlejeunea magnilobula Gola, Ann. Bot. (Rome) 6 (2): 274, 1907 (
** Microlejeunea mammillosa (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999 (
** Microlejeunea minutissima (Mizut.) Grolle, J. Bryol. 21 (1): 42, 1999 (
* Microlejeunea minutistipula Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907–08), Bot. 2: 131, 1911 (
*** Microlejeunea moniliata (Mizut.) R.L.Zhu et Y.M.Wei, Cryptog. Bryol. 34 (3): 308, 2013 (
* Microlejeunea nepalensis Steph., Sp. Hepat. (Stephani) 5: 832, 1915 (
** Microlejeunea nyandaruensis Pócs, Polish Bot. J. 47 (1): 14, 2002 (
** Microlejeunea oblongistipula (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892 (
** Microlejeunea ocellata (Herzog) Grolle, Haussknechtia 8: 60, 2001 (
* Microlejeunea ovistipula Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 131, 1911 (
** Microlejeunea papulosa (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 9, 1892 (
** Microlejeunea perpusilla (Spruce) Steph., Hedwigia 35 (3): 113, 1896 (
** Microlejeunea punctiformis (Taylor) Steph., Hedwigia 29 (2): 90, 1890 (
** Microlejeunea pyriformis (Lindenb. et Gottsche) Steph., Sp. Hepat. (Stephani) 5: 824, 1915 (
** Microlejeunea spinosa (Mizut.) Grolle, J. Bryol. 21 (1): 43, 1999 (
*** Microlejeunea squarrosa (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong, Phytotaxa 85 (2): 51, 2013 (
** Microlejeunea strasbergii Bardat et Ah-Peng, Bryologist 114 (4): 669, 2011 (
*** Microlejeunea subulistipa Steph., Hedwigia 35 (3): 115, 1896 (
* Microlejeunea szechuanensis P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 46, 1955 (
** Microlejeunea udarii P.K.Verma et S.C.Srivast., J. Bombay Nat. Hist. Soc. 108 (2): 122, 2011 [2012] (
*** Microlejeunea ulicina (Taylor) Steph., Hedwigia 29 (2): 88, 1890 (
* Microlejeunea usambarensis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 730, 1913 (
** Microlejeunea valenciana Steph., Sp. Hepat. (Stephani) 5: 815, 1915 (
* Microlejeunea victoriensis D.J.Carr, Proc. Roy. Soc. Victoria 117 (2): 322, 2005 (
*** Microlejeunea wallichiana (Lehm.) R.L.Zhu et Y.M.Wei, Cryptog. Bryol. 34 (3): 308, 2013 (
* Taxilejeunea (Spruce) Steph., Hedwigia 28 (4): 262, 1889 (
** Taxilejeunea acutifolia (Steph.) Steph., Sp. Hepat. (Stephani) 5: 481, 1914 (
** Taxilejeunea antillana Steph., Sp. Hepat. (Stephani) 5: 482, 1914 (
** Taxilejeunea apiculata (Gottsche) J.B.Jack et Steph., Hedwigia 31 (1): 13, 1892 (
** Taxilejeunea argentina Steph., Sp. Hepat. (Stephani) 5: 482, 1914 (
** Taxilejeunea arsenii Steph., Sp. Hepat. (Stephani) 6: 400, 1923 (
** Taxilejeunea auriculata Steph., Sp. Hepat. (Stephani) 5: 459, 1914 (
** Taxilejeunea berteroana Steph., Sp. Hepat. (Stephani) 5: 483, 1914 (
** Taxilejeunea beyrichiana Steph., Sp. Hepat. (Stephani) 5: 460, 1914 (
** Taxilejeunea biapiculata Steph., Sp. Hepat. (Stephani) 5: 460, 1914 (
* Taxilejeunea boliviana Steph., Biblioth. Bot. 87 (2): 260, 1916 (
** Taxilejeunea brasiliensis Steph., Hedwigia 35 (3): 132, 1896 (
** Taxilejeunea coilantha Herzog, Rev. Bryol. Lichénol. 20 (1/2): 159, 1951 [1952] (
* Taxilejeunea compressiuscula Steph., Sp. Hepat. (Stephani) 5: 501, 1914 (
* Taxilejeunea convoluta Herzog, Biblioth. Bot. 88: 31, 1920 [1921] (
* Taxilejeunea cuervi (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 486, 1914 (
* Taxilejeunea cuneistipula Steph., Sp. Hepat. (Stephani) 6: 401, 1923 (
* Taxilejeunea cuspidata Steph., Biblioth. Bot. 87 (2): 260, 1916 (
** Taxilejeunea decurrens Steph., Sp. Hepat. (Stephani) 5: 487, 1914 (
** Taxilejeunea deflexa Steph., Sp. Hepat. (Stephani) 5: 502, 1914 (
** Taxilejeunea densiflora A.Evans, Bull. Torrey Bot. Club 48 (4): 121, 1921 (
** Taxilejeunea diaphana (Lehm.) Steph., Sp. Hepat. (Stephani) 5: 463, 1914 (
* Taxilejeunea dissitifolia Steph., Symb. Antill. 2: 472, 1901 (
** Taxilejeunea eggersiana Schiffn., Bot. Jahrb. Syst. 23 (5): 579, 1897 (
* Taxilejeunea elobulata Sim, Trans. Roy. Soc. South Africa 15 (1): 66, 1926 (
** Taxilejeunea fissistipula Steph., Sp. Hepat. (Stephani) 5: 488, 1914 (
** Taxilejeunea foliicola Steph., Sp. Hepat. (Stephani) 5: 466, 1914 (
** Taxilejeunea furcicornuta Grolle, J. Bryol. 8 (1): 93, 1974 (
** Taxilejeunea fuscorufa Steph., Hedwigia 35 (3): 133, 1896 (
** Taxilejeunea galapagensis Onr., Misc. Bryol. Lichenol. 9 (6): 117, 1982 (
* Taxilejeunea ghatensis P.K.Verma et S.C.Srivast., Proc. Natl. Acad. Sci. India, B 77 (2): 211, 2007 (
* Taxilejeunea giulianettii Steph., Sp. Hepat. (Stephani) 5: 502, 1914 (
** Taxilejeunea gomphocalyx Herzog, Beih. Bot. Centralbl. 61B (3): 580, 1942 (
* Taxilejeunea grandifolia Steph., Biblioth. Bot. 87 (2): 261, 1916 (
* Taxilejeunea grandistipula Steph., Sp. Hepat. (Stephani) 5: 504, 1914 (
* Taxilejeunea hamatifolia Steph., Biblioth. Bot. 87 (2): 261, 1916 (
** Taxilejeunea himalayensis Herzog, Ann. Bryol. 12: 86, 1939 (
* Taxilejeunea immersa Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 96, 1937 (
** Taxilejeunea irregularis Steph., Sp. Hepat. (Stephani) 5: 490, 1914 (
** Taxilejeunea jamaicensis A.Evans, Bull. Torrey Bot. Club 48 (4): 117, 1921 (
** Taxilejeunea jeringii Steph., Hedwigia 35 (3): 134, 1896 (
** Taxilejeunea killipii Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 190, 1955 (
* Taxilejeunea laevis (Gottsche) Steph., Hedwigia 44 (4): 229, 1905 (
* Taxilejeunea langiana Pearson, Ann. Bryol. 4: 102, 1931 (
** Taxilejeunea lindbergiana Steph., Sp. Hepat. (Stephani) 5: 491, 1914 (
** Taxilejeunea linguifolia Steph., Sp. Hepat. (Stephani) 5: 471, 1914 (
* Taxilejeunea maxima Steph., Sp. Hepat. (Stephani) 5: 473, 1914 (
** Taxilejeunea mexicana Steph., Sp. Hepat. (Stephani) 6: 402, 1923 (
* Taxilejeunea microstipula Steph., Sp. Hepat. (Stephani) 5: 493, 1914 (
* Taxilejeunea mucronata Steph., Sp. Hepat. (Stephani) 5: 473, 1914 (
** Taxilejeunea multiflora Steph., Hedwigia 35 (3): 135, 1896 (
* Taxilejeunea muscicola Steph., Biblioth. Bot. 87 (2): 263, 1916 (
* Taxilejeunea nilgiriensis P.K.Verma et S.C.Srivast., Proc. Natl. Acad. Sci. India, B 77 (2): 207, 2007 (
** Taxilejeunea nymannii Steph., Sp. Hepat. (Stephani) 5: 507, 1914 (
** Taxilejeunea obtusifolia Steph., Sp. Hepat. (Stephani) 5: 474, 1914 (
** Taxilejeunea papuliflora Steph., Sp. Hepat. (Stephani) 5: 494, 1914 (
** Taxilejeunea parvibracteata Steph., Hedwigia 35 (3): 136, 1896 (
** Taxilejeunea parvistipula Steph., Sp. Hepat. (Stephani) 6: 403, 1923 (
* Taxilejeunea paucidens Steph., Biblioth. Bot. 87 (2): 263, 1916 (
* Taxilejeunea pendula Steph., Biblioth. Bot. 87 (2): 263, 1916 (
* Taxilejeunea peruviana Steph., Sp. Hepat. (Stephani) 5: 475, 1914 (
** Taxilejeunea planilobula Herzog, Rev. Bryol. Lichénol. 20 (1/2): 160, 1951 [1952] (
** Taxilejeunea pulchriflora Pearson, Ark. Bot. 19 (5): 15, 1924 (
* Taxilejeunea pusilla Steph., Biblioth. Bot. 87 (2): 264, 1916 (
** Taxilejeunea renistipula (Lindenb.) Steph., Hedwigia 29 (3): 142, 1890 (
* Taxilejeunea rufescens Steph., Biblioth. Bot. 87 (2): 264, 1916 (
*** Taxilejeunea serpillifolioides (Raddi) D.P.Costa, J. Bryol. 31 (4): 230, 2009 (
** Taxilejeunea setchellii Pearson, Amer. Samoa: 146, 1924 (
** Taxilejeunea speciosa Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 188, 1955 (
* Taxilejeunea splendida Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 88, 1937 (
** Taxilejeunea stephanii Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 90, 1937 (
** Taxilejeunea steyermarkii H.Rob., Phytologia 34 (1): 67, 1976 (
** Taxilejeunea surinamensis (Lindenb. et Gottsche) Steph., Hedwigia 29 (3): 142, 1890 (
* Taxilejeunea suringarii Steph., Sp. Hepat. (Stephani) 5: 479, 1914 (
** Taxilejeunea tenerrima Steph., Sp. Hepat. (Stephani) 6: 406, 1923 (
** Taxilejeunea tenuiplica Steph., Sp. Hepat. (Stephani) 5: 480, 1914 (
* Taxilejeunea tjibodensis (Steph.) Eifrig, Monogr. Stud. Indomal. Art. Taxilejeunea: 91, 1937 (
** Taxilejeunea tonduzana Steph., Sp. Hepat. (Stephani) 5: 498, 1914 (
* Taxilejeunea uleana Steph., Hedwigia 35 (3): 136, 1896 (
* Taxilejeunea umbonata Steph., Sp. Hepat. (Stephani) 5: 481, 1914 (
* Taxilejeunea urbanii Steph., Symb. Antill. (Urban) 3 (2): 278, 1902 (
** Taxilejeunea vallis-gratiae Steph., Hedwigia 35 (3): 137, 1896 (
*** Lepidolejeunea R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963 (
** subg. Kingiolejeunea (H.Rob.) R.M.Schust., Phytologia 45 (5): 425, 1980 (
** Lepidolejeunea auriculata Schäf.-Verw. et Heinrichs, Taxon 64 (2): 224, 2015 (
*** Lepidolejeunea cordifissa (Taylor) M.E.Reiner, Nova Hedwigia 83 (3/4): 478, 2006 (
*** Lepidolejeunea grossepapulosa (Steph.) Piippo, Acta Bot. Fenn. 132: 49, 1986 (
*** Lepidolejeunea involuta (Gottsche) Grolle, J. Hattori Bot. Lab. 55: 504, 1984 (
** subg. Lepidolejeunea
*** Lepidolejeunea bidentula (Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980 (
*** Lepidolejeunea bidentula var. novae-caledoniae Piippo, Acta Bot. Fenn. 132: 26, 1986 (
*** Lepidolejeunea falcata (Herzog) R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963 (
*** Lepidolejeunea graeffei (J.B.Jack et Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980 (
*** Lepidolejeunea integristipula (J.B.Jack et Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980 (
** subg. Perilejeunea (Kachroo et R.M.Schust.) R.M.Schust., Phytologia 45 (5): 424, 1980 (
** Lepidolejeunea cuspidata (Gottsche) Heinrichs et Schäf.-Verw., Taxon 64 (2): 224, 2015 (
*** Lepidolejeunea delessertii (Nees et Mont.) Grolle, J. Hattori Bot. Lab. 55: 505, 1984 (
*** Lepidolejeunea eluta (Nees) R.M.Schust., Beih. Nova Hedwigia 9: 139, 1963 (
*** Lepidolejeunea serrulata (Steph.) Grolle, J. Hattori Bot. Lab. 55: 505, 1984 (
*** Lepidolejeunea sullivantii (Gottsche) M.E.Reiner, Nova Hedwigia 83 (3/4): 479, 2006 (
Incertae sedis
*** Lepidolejeunea borneensis (Steph.) R.M.Schust., Phytologia 45 (5): 425, 1980 (
*** Lepidolejeunea longilobula (Amakawa) Xiao L.He, Acta Bot. Fenn. 163: 59, 1999 (
*** Otolejeunea Grolle et Tixier, Nova Hedwigia 32: 609, 1980 (
*** subg. Allorgella (Tixier) Grolle, Haussknechtia 2: 53, 1985 (
*** Otolejeunea australiensis B.M.Thiers, Brittonia 44 (2): 162, 1992 (
*** Otolejeunea hoana (Tixier) Grolle, Haussknechtia 2: 54, 1985 [1986] (
*** Otolejeunea rabenorii Tixier, Nova Hedwigia 46 (3/4): 376, 1988 (
*** Otolejeunea schmidii (Tixier) Grolle, Haussknechtia 2: 54, 1985 [1986] (
*** Otolejeunea schnellii (Tixier) R.L.Zhu et M.L.So, Ann. Bot. Fenn. 34 (4): 287, 1997 (
*** Otolejeunea semperiana (Steph.) Grolle, Haussknechtia 2: 53, 1985 [1986] (
** Otolejeunea subana Pocs, Acta Acad. Ped. Agr., Sect. Biol. 25: 50, 2004 (Pocs 2004).
*** Otolejeunea zantenii Grolle, Haussknechtia 2: 54, 1985 [1986] (
*** subg. Otolejeunea
*** Otolejeunea moniliata Grolle, Nova Hedwigia 32: 609, 1980 (
*** subg. Phoxolejeunea Grolle, Haussknechtia 2: 49, 1985 (
** Otolejeunea philippinensis R.L.Zhu et M.L.So, Syst. Bot. 23 (2): 231, 1998 (
*** Otolejeunea streimannii Grolle, Haussknechtia 2: 49, 1985 [1986] (
*** Rectolejeunea A.Evans, Bull. Torrey Bot. Club 33 (1): 8, 1906 (
* Rectolejeunea colombiana R.M.Schust., J. Hattori Bot. Lab. 89: 146, 2000 (
*** Rectolejeunea emarginuliflora (Schiffn.) A.Evans, Bull. Torrey Bot. Club 33 (1): 14, 1906 (
*** Rectolejeunea flagelliformis A.Evans, Bull. Torrey Bot. Club 33 (1): 9, 1906 (
* Rectolejeunea flagelliformis subsp. hamata R.M.Schust., J. Hattori Bot. Lab. 89: 137, 2000 (
*** Rectolejeunea queenslandica (B.M.Thiers) Xiao L.He, Ann. Bot. Fenn. 34 (2): 129, 1997 (
** Rectolejeunea truncatilobula C.J.Bastos, J. Bryol. 34 (2): 144, 2012 (
*** Rectolejeunea versifolia (Schiffn.) L.Söderstr. et A.Hagborg, Phytotaxa 220 (2): 188, 2015 (
Excluded from the genus
* Rectolejeunea lindenbergii Steph., Sp. Hepat. (Stephani) 5: 689, 1914 (
* Rectolejeunea lindigiana Steph., Sp. Hepat. (Stephani) 5: 690, 1914 (
* Rectolejeunea pachyderma R.M.Schust., J. Hattori Bot. Lab. 89: 148, 2000 (
*** Leptolejeunea (Spruce) Steph., Hedwigia 30 (6): 270, 1891 (
*** Leptolejeunea amphiophthalma Zwickel, Ann. Bryol. 6: 117, 1933 (
** Leptolejeunea apiculata (Horik.) S.Hatt., J. Hattori Bot. Lab. 5: 46, 1951 (
** Leptolejeunea arunachalensis Sudipa Das et D.K.Singh, J. Jap. Bot. 83 (6): 343, 2008 (
*** Leptolejeunea astroidea (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 363, 1913 (
** Leptolejeunea australis Steph., Sp. Hepat. (Stephani) 5: 389, 1913 (
** Leptolejeunea balansae Steph., Hedwigia 35 (3): 105, 1896 (
* Leptolejeunea borneensis Herzog, Flora 135: 394, 1942 (
** Leptolejeunea brasiliensis Bischl., Nova Hedwigia 17: 301, 1969 (
* Leptolejeunea convexistipa Bischl., Nova Hedwigia 17: 325, 1969 (
* Leptolejeunea curvatifolia Steph., Sp. Hepat. (Stephani) 6: 398, 1923 (
** Leptolejeunea denticulata Steph., Sp. Hepat. (Stephani) 5: 389, 1913 (
** Leptolejeunea dentistipula Steph., Sp. Hepat. (Stephani) 5: 379, 1913 (
** Leptolejeunea diversilobulata Bischl., Nova Hedwigia 17: 313, 1969 (
** Leptolejeunea dolabriformis Pearson, J. Linn. Soc., Bot. 46 (305): 37, 1922 (
*** Leptolejeunea elliptica (Lehm. et Lindenb.) Besch., Rev. Bryol. 19 (1): 14, 1892 (
* Leptolejeunea emarginata (Horik.) S.Hatt., J. Hattori Bot. Lab. 5: 46, 1951 (
*** Leptolejeunea epiphylla (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 380, 1913 (
*** Leptolejeunea exocellata (Spruce) A.Evans, Bull. Torrey Bot. Club 29 (8): 498, 1902 (
** Leptolejeunea foliicola Steph., Hedwigia 35 (3): 106, 1896 (
** Leptolejeunea integristipula Steph., Sp. Hepat. (Stephani) 6: 398, 1923 (
* Leptolejeunea jamaicensis R.M.Schust., J. Elisha Mitchell Sci. Soc. 83 (4): 229, 1967 (
** Leptolejeunea lancifolia Steph., Sp. Hepat. (Stephani) 5: 382, 1913 (
** Leptolejeunea latifolia Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 58, 1950 [1951] (
** Leptolejeunea lepinii Steph., Sp. Hepat. (Stephani) 5: 383, 1913 (
** Leptolejeunea ligulata Herzog, Flora 135: 429, 1942 (
*** Leptolejeunea maculata (Mitt.) Schiffn., Consp. Hepat. Arch. Ind.: 275, 1898 (
* Leptolejeunea massartiana Schiffn. ex Herzog, Flora 135: 421, 1942 (
** Leptolejeunea micronesica Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 149, 1965 (
** Leptolejeunea minima Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 60, 1950 [1951] (
** Leptolejeunea mirikana M.Dey et D.K.Singh, Taiwania 55 (4): 355, 2010 (
*** Leptolejeunea moniliata Steph., Sp. Hepat. (Stephani) 5: 371, 1913 (
*** Leptolejeunea obfuscata (Spruce) Steph., Sp. Hepat. (Stephani) 5: 373, 1913 (
* Leptolejeunea punctata Herzog, Flora 135: 432, 1942 (
*** Leptolejeunea radicosa (Nees ex Mont.) Grolle, J. Hattori Bot. Lab. 45: 178, 1979 (
* Leptolejeunea renneri Herzog, Flora 135: 422, 1942 (
* Leptolejeunea revoluta P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 44, 1955 (
** Leptolejeunea rosulans Steph., Sp. Hepat. (Stephani) 5: 390, 1913 (
** Leptolejeunea serratifolia Schiffn., Bot. Jahrb. Syst. 23 (5): 594, 1897 (
** Leptolejeunea serrulata Herzog, Flora 135: 426, 1942 (
** Leptolejeunea spinistipula (Mizut.) Xiao L.He, Ann. Bot. Fenn. 34 (2): 127, 1997 (
** Leptolejeunea subdentata Schiffn. ex Herzog, Flora 135: 403, 1942 (
*** Leptolejeunea subrotundifolia Herzog, Flora 135: 424, 1942 (
*** Leptolejeunea tridentata Bischl., Nova Hedwigia 17: 335, 1969 (
* Leptolejeunea trigonostipa (Spruce) Steph., Sp. Hepat. (Stephani) 5: 376, 1913 (
** Leptolejeunea tripuncta (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 388, 1913 (
** Leptolejeunea truncatifolia Steph., Sp. Hepat. (Stephani) 5: 388, 1913 (
* Leptolejeunea udarii M.Dey et D.K.Singh, Taiwania 55 (4): 359, 2010 (
*** Leptolejeunea vitrea (Nees) Schiffn., Hepat. (Engl.-Prantl): 126, 1893 (
** Pycnolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 124, 1893 (
* Pycnolejeunea anotomensis Tixier, Bull. Soc. Hist. Nat. Afrique N. 63: 10, 1972 (
** Pycnolejeunea borneensis Steph., Sp. Hepat. (Stephani) 5: 632, 1914 (
* Pycnolejeunea cavistipula (Steph.) Mizut., J. Hattori Bot. Lab. 36: 161, 1972 [1973] (
* Pycnolejeunea connivens Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (
*** Pycnolejeunea contigua (Nees) Grolle, J. Hattori Bot. Lab. 45: 179, 1979 (
* Pycnolejeunea convexifolia (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 636, 1914 (
*** Pycnolejeunea decurviloba Steph., Hedwigia 35 (3): 125, 1896 (
*** Pycnolejeunea densistipula (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 602, 1914 (
** Pycnolejeunea gradsteinii Ilk.-Borg., Bol. Inst. Bot. (São Paulo) 21 (1): 1, 2011 (
** Pycnolejeunea grandiocellata Steph., Bot. Tidsskr. 24 (3): 279, 1902 (
* Pycnolejeunea grossiloba Steph., Sp. Hepat. (Stephani) 5: 629, 1914 (
*** Pycnolejeunea macroloba (Nees et Mont.) Schiffn., Hepat. (Engl.-Prantl): 124, 1893 (
*** Pycnolejeunea monophthalma (R.M.Schust.) Xiao L.He, Acta Bot. Fenn. 163: 52, 1999 (
** Pycnolejeunea novae-caledoniae (Steph.) Horik., Acta Phytotax. Geobot. 13: 214, 1943 (
* Pycnolejeunea palmicola Steph., Sp. Hepat. (Stephani) 6: 413, 1923 (
*** Pycnolejeunea papillosa Xiao L.He, Acta Bot. Fenn. 163: 55, 1999 (
** Pycnolejeunea porrectilobula C.J.Bastos et O.Yano, Nova Hedwigia 74 (3/4): 440, 2002 (
** Pycnolejeunea retusa R.M.Schust., J. Hattori Bot. Lab. 100: 402, 2006 (
* Pycnolejeunea schlimiana Steph., Sp. Hepat. (Stephani) 5: 615, 1914 (
*** Pycnolejeunea schwaneckei (Steph.) Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19: 246, 1894 (
** Pycnolejeunea sphaeroides (Sande Lac.) J.B.Jack et Steph., Bot. Centralbl. 60 (4): 107, 1894 (
*** Xylolejeunea Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 27, 2001 (
*** Xylolejeunea aquarius (Spruce) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 29, 2001 (
*** Xylolejeunea crenata (Nees et Mont.) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 36, 2001 (
*** Xylolejeunea grolleana (Pócs) Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 32, 2001 (
*** Xylolejeunea muricella Xiao L.He et Grolle, Ann. Bot. Fenn. 38 (1): 34, 2001 (
*** Symbiezidium Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 402, 1877 (
*** Symbiezidium barbiflorum (Lindenb. et Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (11): 540, 1907 [1908] (
*** Symbiezidium dentatum Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 175, 1955 (
*** Symbiezidium madagascariense Steph., Sp. Hepat. (Stephani) 5: 99, 1912 (
*** Symbiezidium transversale (Sw.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 403, 1877 (
*** Symbiezidium transversale var. hookerianum (Gottsche, Lindenb. et Nees) Gradst. et J.Beek, Beih. Nova Hedwigia 80: 237, 1985 (
*** Acrolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Acrolejeunea sandvicensis (Gottsche) Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 312, 1896 (
** sect. Acrolejeunea
*** Acrolejeunea emergens (Mitt.) Steph., Pflanzenw. Ost-Afrikas C: 65, 1895 (
*** Acrolejeunea emergens var. confertissima (Steph.) Gradst., Bryophyt. Biblioth. 4: 76, 1975 (
*** Acrolejeunea heterophylla (A.Evans) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974 (
*** Acrolejeunea torulosa (Lehm. et Lindenb.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
** sect. Minores (Verd.) L.Söderstr. et A.Hagborg, Bryophyte Diversity Evol. 36 (1): 41, 2014 (
*** Acrolejeunea arcuata (Nees) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974 (
** Acrolejeunea arcuata subsp. gradsteinii M.A.M.Renner, Phytotaxa 83 (1): 42, 2013 (
*** Acrolejeunea fertilis (Reinw., Blume et Nees) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Acrolejeunea parvula (Mizut.) Gradst., Bryophyt. Biblioth. 4: 115, 1975 (
*** Acrolejeunea pycnoclada (Taylor) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Acrolejeunea pycnoclada subsp. latistipula Gradst., Bryophyt. Biblioth. 4: 113, 1975 (
*** Acrolejeunea tjibodensis (Verd.) Grolle et Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (
** sect. Pusillae Gradst., Bryophyt. Biblioth. 4: 59, 1975 (
*** Acrolejeunea pusilla (Steph.) Grolle et Gradst., J. Hattori Bot. Lab. 38: 332, 1974 (
*** Acrolejeunea sikkimensis (Mizut.) Gradst., Bryophyt. Biblioth. 4: 83, 1975 (
** sect. Recurvatae Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014 (
*** Acrolejeunea recurvata Gradst., Bryophyt. Biblioth. 4: 79, 1975 (
** sect. Regulares (Verd.) Gradst., Bryophyt. Biblioth. 4: 63, 1975 (
*** Acrolejeunea allisonii Gradst., Bryophyt. Biblioth. 4: 103, 1975 (
*** Acrolejeunea aulacophora (Mont.) Steph., Bot. Jahrb. Syst. 20 (3): 317, 1895 (
*** Acrolejeunea mollis (Hook.f. et Taylor) Schiffn., Hedwigia 33 (4): 178, 1894 (
*** Acrolejeunea securifolia (Nees) Steph., Hedwigia 34 (2): 59, 1895 (
*** Acrolejeunea securifolia subsp. caledonica (Steph.) Gradst., Bryophyt. Biblioth. 4: 100, 1975 (
*** Acrolejeunea securifolia subsp. hartmannii (Steph.) Gradst., Bryophyt. Biblioth. 4: 99, 1975 (
*** Acrolejeunea securifolia subsp. pallida (Ångstr.) Gradst., Bryophyt. Biblioth. 4: 101, 1975 (
** sect. Trocholejeunea (Schiffn.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014 (
*** Acrolejeunea crassicaulis (Steph.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014 (
*** Acrolejeunea infuscata (Mitt.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 38, 2014 (
** Acrolejeunea meghalayensis (Ajit P.Singh et V.Nath) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014 (
** Acrolejeunea sinensis (Jian Wang bis, R.L.Zhu et Gradst.) Jian Wang bis et Gradst., Bryophyte Diversity Evol. 36 (1): 39, 2014 (
Excluded from the genus
* Acrolejeunea abnormis (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 4, 1892 (
* Acrolejeunea comptonii Pearson, J. Linn. Soc., Bot. 46 (305): 33, 1922 (
* Acrolejeunea inflexa (Gottsche) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 4, 1892 (
*** Archilejeunea (Spruce) Steph., Hedwigia 27 (3/4): 113, 1888 (
*** Archilejeunea abbreviata (Mitt.) Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 117, 1951 (
** Archilejeunea africana Steph., Sp. Hepat. (Stephani) 4: 705, 1911 (
** Archilejeunea alata Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 33, 1893 [1894] (
** Archilejeunea amakawana Inoue, J. Jap. Bot. 41 (1): 16, 1966 (
*** Archilejeunea auberiana (Mont.) Steph., Hedwigia 29 (3): 134, 1890 (
*** Archilejeunea autoica Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 119, 1951 (Vanden Bergen 1951b).
*** Archilejeunea badia (Spruce) Steph., Sp. Hepat. (Stephani) 4: 711, 1911 (
* Archilejeunea bilabiata (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 723, 1911 (
*** Archilejeunea bischleriana Gradst., Fl. Neotrop. Monogr. 62: 62, 1994 (
** Archilejeunea bongardii Steph., Hedwigia 29 (1): 20, 1890 (
** Archilejeunea brachyantha J.B.Jack et Steph., Bot. Centralbl. 60 (4): 104, 1894 (
** Archilejeunea brevilobula Steph., Sp. Hepat. (Stephani) 4: 706, 1911 (
*** Archilejeunea crispistipula (Spruce) Steph., Sp. Hepat. (Stephani) 4: 712, 1911 (
* Archilejeunea eberhardtii Steph., Sp. Hepat. (Stephani) 4: 725, 1911 (
** Archilejeunea elobulata Steph., Sp. Hepat. (Stephani) 4: 707, 1911 (
** Archilejeunea gradsteinii X.Q.Shi et R.L.Zhu, Nova Hedwigia 100 (3/4): 592, 2015 (Shi and Zhu 2015).
** Archilejeunea incrassata Steph., Rev. Bryol. 35 (2): 30, 1908 (
** Archilejeunea jonesii Vanden Berghen, Rev. Bryol. Lichénol. 20 (1/2): 116, 1951 (
*** Archilejeunea juliformis (Nees) Gradst., Bryophyt. Biblioth. 4: 127, 1975 (
*** Archilejeunea kiushiana (Horik.) Verd., Ann. Bryol., Suppl. 4: 46, 1934 (
*** Archilejeunea ludoviciana (De Not.) P.Geissler et Gradst., J. Hattori Bot. Lab. 75: 202, 1994 (
*** Archilejeunea ludoviciana subsp. porelloides (Spruce) Gradst., Fl. Neotrop. Monogr. 62: 58, 1994 (
** Archilejeunea nebeliana Gradst. et Schäf.-Verw., Cryptog. Bryol. 33 (2): 108, 2012 (
*** Archilejeunea olivacea (Hook.f. et Taylor) Steph., Hedwigia 29 (3): 134, 1890 (
*** Archilejeunea parviflora (Nees) Steph., Hedwigia 29 (3): 134, 1890 (
** Archilejeunea planifolia (Horik.) Mizut., J. Hattori Bot. Lab. 73: 175, 1993 (
*** Archilejeunea planiuscula (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 731, 1911 (
Excluded from the genus
* Archilejeunea negrensis Steph., Sp. Hepat. (Stephani) 4: 716, 1911 (
* Archilejeunea ovata Herzog, Rev. Bryol. Lichénol. 20 (1/2): 130, 1951 [1952] (
*** Bryopteris (Nees) Lindenb., Syn. Hepat. 2: 284, 1845 (
*** Bryopteris diffusa (Sw.) Nees, Syn. Hepat. 2: 286, 1845 (
*** Bryopteris filicina (Sw.) Nees, Syn. Hepat. 2: 284, 1845 (
* Bryopteris fissiloba Steph., Sp. Hepat. (Stephani) 6: 568, 1924 (
*** Bryopteris gaudichaudii Gottsche, Ann. Sci. Nat. Bot. (sér. 4) 8: 340, 1857 (
*** Caudalejeunea Schiffn., Hepat. (Engl.-Prantl): 129, 1893 (
** subg. Acaudalejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 779, 1980 (
*** Caudalejeunea grolleana Gradst., Acta Bot. Neerl. 23 (3): 334, 1974 (
** subg. Caudalejeunea
** Caudalejeunea africana (Steph.) Schiffn., Hepat. (Engl.-Prantl): 129, 1893 (
*** Caudalejeunea hanningtonii (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 129, 1893 (
** Caudalejeunea katangensis Vanden Berghen, Explor. Hydrobiol. Lac Bangweolo Luapula: 94, 1972 (
*** Caudalejeunea lehmanniana (Gottsche) A.Evans, Bull. Torrey Bot. Club 34 (11): 554, 1907 [1908] (
** Caudalejeunea lewallei Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 42 (4): 434, 1972 (
** subg. Vermilejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 778, 1980 (
*** Caudalejeunea cristiloba (Steph.) Gradst., Acta Bot. Neerl. 23 (3): 340, 1974 (
*** Caudalejeunea cristiloba subsp. samoana (Steph.) Gradst., Acta Bot. Neerl. 23 (3): 343, 1974 (
** Caudalejeunea dusenii Steph., Sp. Hepat. (Stephani) 5: 11, 1912 (
*** Caudalejeunea reniloba (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 16, 1912 (
** Caudalejeunea yangambiensis (Vanden Berghen) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 192, 1957 (
Incertae sedis
** Caudalejeunea acutifolia Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 478, 1947 (
* Caudalejeunea mauritiana Horik., Acta Phytotax. Geobot. 13: 214, 1943 (
** Caudalejeunea streimannii Gyarmati, Trop. Bryol. 22: 129, 2002 (
** Caudalejeunea tridentata R.L.Zhu, Y.M.Wei et Qiong He, Bryologist 114 (3): 469, 2011 (
** Cephalantholejeunea R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 798, 1980 (
*** Cephalantholejeunea temnanthoides (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 807, 1980 (
** Cephalolejeunea Mizut., J. Hattori Bot. Lab. 46: 359, 1979 (
*** Cephalolejeunea parvilobula Mizut., J. Hattori Bot. Lab. 46: 359, 1979 (
*** Frullanoides Raddi, Critt. Brasil.: 13, 1822 (
*** Frullanoides bahamensis (A.Evans) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 81, 1985 (
*** Frullanoides corticalis (Lehm. et Lindenb.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 84, 1985 (
*** Frullanoides densifolia Raddi, Critt. Brasil.: 14, 1822 (
*** Frullanoides densifolia subsp. grandidentata (L.Clark) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 95, 1985 (
*** Frullanoides laciniatiflora (Loitl.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 100, 1985 (
*** Frullanoides liebmaniana (Lindenb. et Gottsche) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 102, 1985 (
*** Frullanoides mexicana van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 106, 1985 (
*** Frullanoides tristis (Steph.) van Slageren, Meded. Bot. Mus. Herb. Rijks Univ. Utrecht 544: 110, 1985 (
*** Fulfordianthus Gradst., Bryologist 95 (1): 44, 1992 (
*** Fulfordianthus evansii (Fulford) Gradst., Bryologist 95 (1): 46, 1992 (
*** Fulfordianthus pterobryoides (Spruce) Gradst., Bryologist 95 (1): 44, 1992 (
*** Lopholejeunea (Spruce) Steph., Bot. Gaz. 15 (11): 285, 1890 (
** subg. Lopholejeunea
** Lopholejeunea borbonica Steph., Hedwigia 35 (3): 109, 1896 (
** Lopholejeunea jonesii Vanden Berghen, Bull. Jard. Bot. État Bruxelles 20 (2): 178, 1950 (
** Lopholejeunea laciniata E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 194, 1957 (
** Lopholejeunea minima Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 437, 1984 (
* Lopholejeunea obtusilacera Herzog, Bull. Jard. Bot. État Bruxelles 20 (2): 172, 1950 (
* Lopholejeunea paramultilacera Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 435, 1984 (
* Lopholejeunea quinquecarinata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 408, 1984 (
** Lopholejeunea renistipula (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 94, 1912 (
** Lopholejeunea revoluta E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 207, 1957 (
** sect. Eulophae Verd., Ann. Bryol., Suppl. 4: 87, 1934 (
*** Lopholejeunea applanata (Reinw., Blume et Nees) Schiffn., Hepat. (Engl.-Prantl): 129, 1893 (
*** Lopholejeunea borneensis (Steph.) Verd., Ann. Bryol., Suppl. 4: 83, 1934 (
*** Lopholejeunea erugata B.M.Thiers, Brittonia 36 (2): 174, 1984 (
*** Lopholejeunea eulopha (Taylor) Schiffn., Hepat. (Engl.-Prantl): 129, 1893 (
*** Lopholejeunea evansiana Verd., Nova Guinea 18: 4, 1934 (
*** Lopholejeunea grollei R.L.Zhu et Gradst., Monogr. Syst. Bot. Missouri Bot. Gard. 74: 36, 2005 (
*** Lopholejeunea herzogiana Verd., Recueil Trav. Bot. Neerl. 30: 217, 1933 (
*** Lopholejeunea hispidissima Steph., Sp. Hepat. (Stephani) 5: 80, 1912 (
*** Lopholejeunea loheri Steph., Sp. Hepat. (Stephani) 5: 77, 1912 (
*** Lopholejeunea minuta R.L.Zhu et Gradst., Nova Hedwigia 78 (3/4): 436, 2004 (
*** Lopholejeunea nigricans (Lindenb.) Schiffn., Consp. Hepat. Arch. Ind.: 293, 1898 (
*** Lopholejeunea plicatiscypha (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 96, 1912 (
* Lopholejeunea proxima Steph., Sp. Hepat. (Stephani) 5: 89, 1912 (
*** Lopholejeunea streimannii B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 37, 1989 (
** Lopholejeunea vojtkoana Gyarmati, Nova Hedwigia 87 (3/4): 480, 2008 (
** sect. Lopholejeunea
*** Lopholejeunea ceylanica Steph., Sp. Hepat. (Stephani) 5: 86, 1912 (
*** Lopholejeunea horticola Schiffn., Ann. Bryol. 6: 133, 1933 (
*** Lopholejeunea latilobula Verd., Nova Guinea 18: 4, 1934 (
*** Lopholejeunea magna Mizut., J. Hattori Bot. Lab. 32: 131, 1969 (
*** Lopholejeunea recurvata Mizut., J. Hattori Bot. Lab. 46: 369, 1979 (
*** Lopholejeunea soae R.L.Zhu et Gradst., Monogr. Syst. Bot. Missouri Bot. Gard. 74: 69, 2005 (
*** Lopholejeunea subfusca (Nees) Schiffn., Bot. Jahrb. Syst. 23 (5): 593, 1897 (
** Lopholejeunea subfusca var. elongata Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 445, 1984 (
*** Lopholejeunea wiltensii Steph., Hedwigia 35 (3): 112, 1896 (
* Lopholejeunea yapensis Steph., Sp. Hepat. (Stephani) 5: 81, 1912 (
*** Lopholejeunea zollingeri (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 296, 1898 (
** subg. Pholianthus B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 25, 1989 (
*** Lopholejeunea colensoi Steph., Sp. Hepat. (Stephani) 5: 97, 1912 (
*** Lopholejeunea pocsii Gyarmati, Cryptog. Bryol. 26 (4): 404, 2005 (
** subg. Pteryganthus B.M.Thiers, Brittonia 35 (1): 85, 1983 (
** Lopholejeunea grandicrista Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 34, 1893 [1894] (
*** Lopholejeunea leioptera Gyarmati, Candollea 56 (1): 80, 2001 (
** Lopholejeunea onraedtii Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 452, 1984 (
** Lopholejeunea sphaerophora (Lehm. et Lindenb.) Steph., Sp. Hepat. (Stephani) 5: 68, 1912 (
* Lopholejeunea tixieriana Vanden Berghen, Bull. Jard. Bot. Natl. Belg. 54 (3/4): 454, 1984 (
** Lopholejeunea utriculata Steph., Sp. Hepat. (Stephani) 5: 69, 1912 (
Incertae sedis
** Lopholejeunea lepidoscypha Kiaer et Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (8): 5, 1892 (
* Lopholejeunea multilacera Steph., Bot. Gaz. 15 (11): 285, 1890 (
** Lopholejeunea udarii M.Dey et D.K.Singh, Nelumbo 53: 197, 2011 (
Excluded from the genus
* Lopholejeunea aberrantia Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 256, 1934 (
* Lopholejeunea vietnamica Tixier, Ann. Hist.-Nat. Mus. Natl. Hung. 66: 99, 1974 (
*** Marchesinia Gray, Nat. Arr. Brit. Pl. 1: 689, 1821 (
*** subg. Marchesinia
*** Marchesinia mackaii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 689, 1821 (
*** subg. Marchesiniopsis R.M.Schust., J. Hattori Bot. Lab. 72: 358, 1992 (
*** Marchesinia bongardiana (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 405, 1877 (
*** Marchesinia brachiata (Sw.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Marchesinia deslooveri Vanden Berghen, Rev. Bryol. Lichénol. 42 (4): 926, 1976 (
*** Marchesinia excavata (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
*** Marchesinia languida (Nees et Mont.) Steph., Sp. Hepat. (Stephani) 5: 149, 1912 (
*** Marchesinia nobilis (Gottsche) X.Q.Shi, R.L.Zhu et Gradst., Phytotaxa 195 (3): 249, 2015 (
*** Marchesinia robusta (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 128, 1893 (
** Mastigolejeunea (Spruce) Steph., Hedwigia 30 (5): 206, 1891 (
*** Mastigolejeunea auriculata (Wilson et Hook.) Steph., Bot. Gaz. 17 (6): 171, 1892 (
** Mastigolejeunea auriculata var. rhodesica (Vanden Berghen) Sukkharak et Gradst., Nova Hedwigia 99 (3/4): 297, 2014 (
*** Mastigolejeunea calcarata (Steph.) Verd., Blumea 1 (1): 230, 1934 (
*** Mastigolejeunea florea (Mitt.) Paris, Rev. Bryol. 33 (3): 42, 1906 (
*** Mastigolejeunea frauenfeldii (Reichardt) Verd., Blumea 1 (1): 230, 1934 (
** Mastigolejeunea gradsteinii Sukkharak, J. Bryol. 36 (1): 56, 2014 (
*** Mastigolejeunea humilis (Gottsche) Schiffn., Hepat. (Engl.-Prantl): 129, 1893 (
*** Mastigolejeunea indica Steph., Sp. Hepat. (Stephani) 4: 776, 1912 (
*** Mastigolejeunea innovans (Spruce) Steph., Sp. Hepat. (Stephani) 4: 765, 1912 (
*** Mastigolejeunea ligulata (Lehm. et Lindenb.) Schiffn., Hepat. (Engl.-Prantl): 129, 1893 (
*** Mastigolejeunea nigra Steph., Hedwigia 30 (5): 206, 1891 (
*** Mastigolejeunea plicatiflora (Spruce) Steph., Sp. Hepat. (Stephani) 4: 766, 1912 (
*** Mastigolejeunea recondita (Steph.) Mizut., J. Hattori Bot. Lab. 32: 134, 1969 (
*** Mastigolejeunea repleta (Taylor) A.Evans, Mem. Torrey Bot. Club 8 (2): 131, 1902 (
*** Mastigolejeunea truncata Mizut., J. Hattori Bot. Lab. 61: 292, 1986 [1987] (
** Mastigolejeunea turgida Steph., Hedwigia 31 (4): 170, 1892 (
*** Mastigolejeunea virens (Ångstr.) Steph., Sp. Hepat. (Stephani) 4: 776, 1912 (
** Phaeolejeunea Mizut., J. Hattori Bot. Lab. 31: 130, 1968 (
*** Phaeolejeunea amicorum (Hürl.) Pócs, Fieldiana, Bot. (n.ser.) 47: 140, 2008 (
** Phaeolejeunea etesseana (Steph.) Mizut., J. Hattori Bot. Lab. 31: 133, 1968 (
** Phaeolejeunea inermis (Steph.) Mizut., J. Hattori Bot. Lab. 31: 134, 1968 (
*** Phaeolejeunea latistipula (Schiffn. ex P.Syd.) Mizut., J. Hattori Bot. Lab. 31: 131, 1968 (
*** Ptychanthus Nees, Naturgesch. Eur. Leberm. 3: 211, 1838 (
** Ptychanthus africanus Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 131, 1911 (
* Ptychanthus stephensonianus (Mitt.) Steph., Sp. Hepat. (Stephani) 4: 754, 1912 (
*** Ptychanthus striatus (Lehm. et Lindenb.) Nees, Naturgesch. Eur. Leberm. 3: 212, 1838 (
** Ptychanthus striatus var. intermedius (Gottsche) Verd., Ann. Bryol., Suppl. 4: 122, 1934 (
*** Schiffneriolejeunea Verd., Ann. Bryol. 6: 89, 1933 (
** sect. Pappeanae R.M.Schust. ex Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 174, 1985 (
*** Schiffneriolejeunea fragilis Gradst. et E.W.Jones, J. Bryol. 12 (1): 45, 1982 (
*** Schiffneriolejeunea madagascariensis (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (
*** Schiffneriolejeunea pappeana var. bidentata Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 182, 1985 (
*** Schiffneriolejeunea pappeana var. integra Gradst. et Vanden Berghen, Beih. Nova Hedwigia 80: 182, 1985 (
*** Schiffneriolejeunea parviloba (Steph.) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (
*** Schiffneriolejeunea pappeana (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (
** sect. Schiffneriolejeunea, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 72, 1981 (
*** Schiffneriolejeunea altimontana Vanden Berghen, Rev. Bryol. Lichénol. 42 (4): 923, 1976 (
*** Schiffneriolejeunea amazonica Gradst., Beih. Nova Hedwigia 80: 25, 1985 (
*** Schiffneriolejeunea cumingiana (Mont.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (
*** Schiffneriolejeunea ferruginea (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (
*** Schiffneriolejeunea occulta (Steph.) Gradst., J. Hattori Bot. Lab. 38: 333, 1974 (
*** Schiffneriolejeunea omphalanthoides Verd., Ann. Bryol. 6: 91, 1933 (
*** Schiffneriolejeunea tumida (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (
*** Schiffneriolejeunea tumida var. haskarliana (Gottsche) Gradst. et Terken, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 77, 1981 (
*** Schiffneriolejeunea nymannii (Steph.) Gradst. et Terken, Occas. Pap. Farlow Herb. Cryptog. Bot. 16: 79, 1981 (
*** Schiffneriolejeunea polycarpa (Nees) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (
*** Schiffneriolejeunea pulopenangensis (Gottsche) Gradst., J. Hattori Bot. Lab. 38: 335, 1974 (
** Spruceanthus Verd., Ann. Bryol., Suppl. 4: 151, 1934 (
*** Spruceanthus macrostipulus (Steph.) Gradst., Trop. Bryol. 4: 13, 1991 (
** Spruceanthus mamillilobulus (Herzog) Verd., Hepat. Select. Crit. 9: no. 447, 1936 (
*** Spruceanthus pluriplicatus (Steph.) Gradst., Schriftenreihe Mensch, Kultur Umwelt z. Bergl. W Neug. 7: 14, 1981 (
*** Spruceanthus polymorphus (Sande Lac.) Verd., Ann. Bryol., Suppl. 4: 155, 1934 (
*** Spruceanthus semirepandus (Nees) Verd., Ann. Bryol., Suppl. 4: 153, 1934 (
*** Spruceanthus sulcatus (Nees) Gradst., Beih. Nova Hedwigia 80: 26, 1985 (
*** Spruceanthus theobromae (Spruce) Gradst., Beih. Nova Hedwigia 80: 26, 1985 (
*** Spruceanthus thozetianus (Gottsche et F.Muell.) B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 62, 1989 (
*** Thysananthus Lindenb., Nov. Stirp. Pug. 8: 24, 1844 (
** sect. Thysananthus
** subsect. Anguiformes Sukkharak, Phytotaxa 193 (1): 37, 2015 (
*** Thysananthus anguiformis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 289, 1845 (
*** Thysananthus pancheri (Steph.) Hürl., Bauhinia 9 (2): 167, 1989 (
** subsect. Thysananthus
*** Thysananthus aculeatus Herzog, Ann. Bryol. 4: 89, 1931 (
*** Thysananthus amazonicus (Spruce) Schiffn., Hepat. (Engl.-Prantl): 130, 1893 (
*** Thysananthus appendiculatus Steph., Sp. Hepat. (Stephani) 4: 794, 1912 (
** Thysananthus ciliaris (Sande Lac.) Sukkharak, Nova Hedwigia 99 (3/4): 339, 2014 (
*** Thysananthus comosus Lindenb., Nov. Stirp. Pug. 8: 25, 1844 (
*** Thysananthus convolutus Lindenb., Syn. Hepat. 2: 288, 1845 (
** Thysananthus convolutus var. laceratus (Steph.) Sukkharak, Phytotaxa 193 (1): 30, 2015 (
*** Thysananthus discretus Sukkharak et Gradst., Cryptog. Bryol. 31 (2): 113, 2010 (
*** Thysananthus gottschei (J.B.Jack et Steph.) Steph., Sp. Hepat. (Stephani) 4: 787, 1912 (
** Thysananthus gottschei var. continuus Sukkharak, Phytotaxa 193 (1): 33, 2015 (
*** Thysananthus spathulistipus (Reinw., Blume et Nees) Lindenb., Syn. Hepat. 2: 287, 1845 (
** sect. Vittatae Verd., Ann. Bryol., Suppl. 4: 182, 1934 (
** subsect. Sandeanthus (B.M.Thiers et Gradst.) Sukkharak, Phytotaxa 193 (1): 43, 2015 (
*** Thysananthus mollis Steph., Sp. Hepat. (Stephani) 4: 798, 1912 (
*** Thysananthus montanus Gradst., Xiao L.He et Piippo, Acta Bot. Fenn. 174: 77, 2002 (
*** Thysananthus retusus (Reinw., Blume et Nees) B.M.Thiers et Gradst., Mem. New York Bot. Gard. 52: 67, 1989 (
** Thysananthus retusus subsp. sellingii (Herzog) Sukkharak, Phytotaxa 193 (1): 47, 2015 (
** subsect. Vittatae (Verd.) Sukkharak, Phytotaxa 193 (1): 40, 2015 (
*** Thysananthus fruticosus (Lindenb. et Gottsche) Schiffn., Hepat. (Engl.-Prantl): 130, 1893 (
** Tuzibeanthus S.Hatt., Biosphaera 1 (1): 7, 1947 (
*** Tuzibeanthus chinensis (Steph.) Mizut., J. Hattori Bot. Lab. 24: 151, 1961 (
** Verdoornianthus Gradst., Bryologist 80 (4): 607, 1977 [1978] (
*** Verdoornianthus griffinii Gradst., Bryologist 80 (4): 609, 1977 [1978] (
*** Verdoornianthus marsupiifolius (Spruce) Gradst., Bryologist 80 (4): 609, 1977 [1978] (
by M. von Konrat and M.A.M. Renner
*** Goebeliella Steph., Hedwigia 51 (1): 61, 1911 (
*** Goebeliella cornigera (Mitt.) Steph., Hedwigia 51 (1): 62, 1911 (
by M. von Konrat
The generic composition of Lepidolaenaceae follows
*** Gackstroemia Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (
** subg. Gackstroemia
** Gackstroemia ljungneri (Herzog) Grolle, J. Hattori Bot. Lab. 30: 17, 1967 (
*** Gackstroemia magellanica (Lam.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (
*** Gackstroemia novae-zelandiae R.M.Schust. et J.J.Engel, Phytotaxa 118 (1): 10, 2013 (
*** Gackstroemia weindorferi (Herzog) Grolle, J. Hattori Bot. Lab. 30: 20, 1967 (
** subg. Hariotiella (Besch. et C.Massal. ex Schiffn.) Grolle, J. Hattori Bot. Lab. 30: 12, 1967 (
*** Gackstroemia hariotiana (Besch. et C.Massal.) Grolle, J. Hattori Bot. Lab. 30: 12, 1967 (
*** Gackstroemia patagonica (Steph.) Grolle, J. Hattori Bot. Lab. 30: 14, 1967 (
** Gackstroemia schwabei (Herzog) Grolle, J. Hattori Bot. Lab. 30: 16, 1967 (
Incertae sedis
*** Gackstroemia alpina R.M.Schust., J. Hattori Bot. Lab. 36: 349, 1972 [1973] (
*** Lepidogyna R.M.Schust., Phytologia 45 (5): 419, 1980 (
*** Lepidogyna hodgsoniae (Grolle) R.M.Schust., Phytologia 45 (5): 419, 1980 (
*** Lepidogyna menziesii (Hook.) R.M.Schust., Phytologia 45 (5): 419, 1980 (
*** Lepidolaena Dumort., Recueil Observ. Jungerm.: 13, 1835 (
*** Lepidolaena berggrenii E.A.Hodgs., Trans. Roy. Soc. New Zealand 87 (3/4): 205, 1959 (
** Lepidolaena brachyclada (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (
*** Lepidolaena clavigera (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (
*** Lepidolaena novae-zelandiae (E.A.Hodgs. et S.W.Arnell) von Konrat, L.Söderstr. et A.Hagborg, Phytotaxa 65: 51, 2012 (
*** Lepidolaena palpebrifolia (Hook.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (
*** Lepidolaena reticulata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (
*** Lepidolaena taylorii (Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 393, 1877 (
*** Ascidiota C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 5 (2): 256, 1898 (
*** Ascidiota blepharophylla C.Massal., Nuovo Giorn. Bot. Ital. (n.ser.) 5 (2): 257, 1898 (
** Ascidiota blepharophylla subsp. alaskana Steere et R.M.Schust., Bull. Torrey Bot. Club 87 (3): 213, 1960 (
*** Porella L., Sp. Pl. 1: 1106, 1753 (
** Porella abyssinica (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
** Porella abyssinica var. hoehnelii (Steph.) Pócs, Fragm. Florist. Geobot. 39 (1): 229, 1994 (
** Porella acutifolia (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877 (
** Porella acutifolia var. hattoriana (Pócs) S.Hatt., Misc. Bryol. Lichenol. 8 (4): 79, 1979 (
* Porella acutifolia var. linguifolia (Steph.) M.L.So, Syst. Bot. 27 (1): 5, 2002 (
** Porella acutifolia subsp. tosana (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 100, 1978 (
* Porella andica (Gottsche) Hässel, Beih. Nova Hedwigia 134: 452, 2009 (
*** Porella arboris-vitae (With.) Grolle, Trans. Brit. Bryol. Soc. 5 (4): 770, 1969 (
** Porella arboris-vitae subsp. nitidula (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 40: 123, 1976 (
** Porella baueri (Schiffn.) C.E.O.Jensen, Danmarks mosser: 240, 1915 (
** Porella bolanderi (Austin) Pearson, List. Canad. Hepat.: 7, 1890 (
** Porella borellii (Gola) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 15, 1962 (
*** Porella brachiata (Taylor) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 334, 1885 (
*** Porella brasiliensis (Raddi) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893 (
* Porella brasiliensis var. ciliata (Gottsche, Lindenb. et Nees) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893 (
* Porella brasiliensis var. laevior (Gottsche, Lindenb. et Nees) Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 246, 1893 (
** Porella caespitans (Steph.) S.Hatt., J. Hattori Bot. Lab. 33: 50, 1970 (
** Porella caespitans var. cordifolia (Steph.) S.Hatt. ex T.Katag. et T.Yamag., Bryol. Res. 10 (5): 133, 2011 (
** Porella caespitans subsp. latior (S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 40: 127, 1976 (
** Porella caespitans var. nipponica S.Hatt., J. Hattori Bot. Lab. 33: 57, 1970 (
** Porella caespitans var. reflexigastria (Pócs) S.Hatt., J. Hattori Bot. Lab. 40: 127, 1976 (
** Porella campylophylla (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877 (
** Porella campylophylla subsp. lancistipula (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 102, 1978 (
** Porella campylophylla var. ligulifera (Taylor) S.Hatt., J. Hattori Bot. Lab. 32: 333, 1969 (
** Porella campylophylla var. tixieri (Pócs) S.Hatt., J. Hattori Bot. Lab. 40: 128, 1976 (
** Porella canariensis (F.Weber) Underw., Rep. (Annual) Missouri Bot. Gard. 8: 186, 1897 (
* Porella capehorniensis Swails, Nova Hedwigia 19: 244, 1970 (Swails 1970).
** Porella capensis (Gottsche) Mitt., J. Linn. Soc., Bot. 22 (146): 323, 1886 (
* Porella caucasica Steph., Bot. Centralbl. 50 (3): 71, 1892 (
** Porella chenii S.Hatt., J. Hattori Bot. Lab. 30: 129, 1967 (
*** Porella chilensis (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
* Porella chilensis var. antucensis (Gottsche) Hässel, Beih. Nova Hedwigia 134: 452, 2009 (
** Porella chilensis var. fernandeziensis (Herzog) Swails, Nova Hedwigia 19: 236, 1970 (Swails 1970). Bas.: Madotheca chilensis f. fernandeziensis Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 736, 1942 (
** Porella chilensis var. microloba (Herzog) Swails, Nova Hedwigia 19: 236, 1970 (Swails 1970). Bas.: Madotheca chilensis f. microloba Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 736, 1942 (
** Porella chinensis (Steph.) S.Hatt., J. Hattori Bot. Lab. 30: 131, 1967 (
** Porella chinensis var. crispata Udar et Shaheen, Misc. Bryol. Lichenol. 9 (4): 74, 1982 (
** Porella chinensis var. decurrens (Steph.) S.Hatt., J. Hattori Bot. Lab. 44: 102, 1978 (
** Porella chinensis var. hattorii Udar et Shaheen, Misc. Bryol. Lichenol. 9 (7): 146, 1983 (
** Porella chinensis var. irregularis (Steph.) S.Hatt., J. Hattori Bot. Lab. 39: 270, 1975 (
** Porella circinnata Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 355, 1874 (
*** Porella cordaeana (Huebener) Moore, Proc. Roy. Irish Acad. (ser. 2) 2: 618, 1877 (
** Porella cranfordii Steph., Hedwigia 28 (4): 270, 1889 (
*** Porella crispata (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
** Porella cucullistipula Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 38, 1893 [1894] (
** Porella densifolia (Steph.) S.Hatt., J. Jap. Bot. 20: 109, 1944 (
** Porella densifolia subsp. andamana S.Hatt., J. Hattori Bot. Lab. 32: 346, 1969 (
** Porella densifolia subsp. appendiculata (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 343, 1969 (
** Porella densifolia var. oviloba (Steph.) N.Kitag., Acta Phytotax. Geobot. 19 (2/3): 64, 1962 (
** Porella densifolia var. paraphyllina (P.C.Chen) Pócs, J. Hattori Bot. Lab. 31: 84, 1968 (
** Porella densifolia var. pilosa S.Hatt. et K.C.Chang, Bull. Bot. Res., Harbin 8 (2): 44, 1988 (
** Porella densifolia var. robusta (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 343, 1969 (
*** Porella elegantula (Mont.) E.A.Hodgs., Svensk Bot. Tidskr. 42 (3): 277, 1948 (
** Porella faurieri (Steph.) S.Hatt., J. Jap. Bot. 20: 109, 1944 (
** Porella fengii P.C.Chen et S.Hatt., J. Hattori Bot. Lab. 30: 133, 1967 (
** Porella geheebii (Steph.) S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 114, 1951 (
** Porella gracillima Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 202, 1891 (
** Porella grandifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 30: 136, 1967 (
** Porella grandiloba Lindb., Acta Soc. Sci. Fenn. 10: 234, 1872 [1873] (
** Porella grollei S.Hatt., J. Hattori Bot. Lab. 34: 411, 1971 (
** Porella handelii S.Hatt., J. Hattori Bot. Lab. 33: 65, 1970 (
** Porella hattorii Udar et Shaheen, Lindbergia 9 (1): 70, 1983 (
** Porella hoeana S.Hatt., Misc. Bryol. Lichenol. 7 (5): 86, 1976 (
* Porella imbricata Lour., Fl. Cochinch. 2: 683, 1790 (
** Porella inaequalis (Gottsche) Perss., Arch. Soc. Zool. Bot. Fenn. “Vanamo”, suppl. 9: 225, 1955 (
** Porella japonica (Sande Lac.) Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 202, 1891 (
** Porella japonica subsp. appalachiana R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 682, 1980 (
** Porella japonica var. densespinosa S.Hatt. et M.X.Zhang, J. Jap. Bot. 60 (11): 324, 1985 (
** Porella javanica (Gottsche) Inoue, J. Hattori Bot. Lab. 30: 60, 1967 (
** Porella latifolia J.S.Lou et Q.Li, Acta Phytotax. Sin. 25 (6): 482, 1987 (
*** Porella leiboldii (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
** Porella longifolia (Steph.) S.Hatt., J. Hattori Bot. Lab. 32: 351, 1969 (
** Porella madagascariensis (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
* Porella maxima (Steph.) M.L.So, Syst. Bot. 27 (1): 11, 2002 (
*** Porella mexicana (Hampe ex Gottsche, Lindenb. et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
** Porella montantii (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 460, 1963 (
*** Porella navicularis (Lehm. et Lindenb.) Pfeiff., Fl. Niederhessen 2: 234, 1855 (
** Porella nitens (Steph.) S.Hatt., Fl. E. Himalaya: 525, 1966 (
** Porella oblongifolia S.Hatt., J. Jap. Bot. 19 (7): 200, 1943 (
*** Porella obtusata (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
** Porella obtusata var. macroloba (Steph.) S.Hatt. et M.X.Zhang, J. Jap. Bot. 60 (11): 325, 1985 (
** Porella obtusiloba S.Hatt., J. Hattori Bot. Lab. 33: 69, 1970 (
** Porella perrottetiana (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877 (
** Porella perrottetiana var. angustifolia Pócs, J. Hattori Bot. Lab. 31: 75, 1968 (
** Porella perrottetiana var. ciliatodentata (P.C.Chen et P.C.Wu) S.Hatt., J. Hattori Bot. Lab. 30: 144, 1967 (
** Porella perrottetiana var. triciliata (Steph.) Pócs, J. Hattori Bot. Lab. 31: 75, 1968 (
*** Porella pinnata L., Sp. Pl. 1: 1106, 1753 (
** Porella planifolia J.S.Lou, Coll. Pap. Quing-Zang Huang-Den 1: 277, 1983 (
*** Porella platyphylla (L.) Pfeiff., Fl. Niederhessen 2: 234, 1855 (
*** Porella platyphylloidea (Schwein.) Lindb., Morgonbladet (Helsinki) 1876 (287, 10 Dec.): 1, 1876 (
** Porella plicata J.S.Lou, Acta Phytotax. Sin. 18 (1): 119, 1980 (
** Porella plumosa (Mitt.) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 17, 1962 (
** Porella prolixa (Gottsche) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 460, 1963 (
** Porella pulcherrima Herzog et S.Hatt., Bull. Natl. Sci. Mus. Tokyo, B 12 (1): 34, 1986 (
*** Porella reflexa (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 408, 1877 (
** Porella revoluta (Lehm. et Lindenb.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
** Porella revoluta var. propinqua (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 30: 148, 1967 (
** Porella roellii Steph., Bot. Centralbl. 45: 203, 1891 (
*** Porella saccata M.L.So, New Zealand J. Bot. 43 (1): 302, 2005 (
** Porella sichuanensis S.Hatt. et K.C.Chang, Bull. Bot. Res., Harbin 8 (2): 43, 1988 (
** Porella spinulosa (Steph.) S.Hatt., J. Hattori Bot. Lab. 33: 74, 1970 (
*** Porella squamulifera (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
** Porella stephaniana (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 5: 81, 1951 (
** Porella subdentata (Mitt.) Steph., Hedwigia 30 (5): 203, 1891 (
** Porella subdentata var. camerunensis E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 456, 1963 (
** Porella subobtusa (Steph.) S.Hatt., J. Jap. Bot. 20: 111, 1944 (
** Porella subparaphyllina J.S.Lou, Acta Phytotax. Sin. 25 (6): 483, 1987 (
*** Porella subsquarrosa (Nees et Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
*** Porella swailsii Grolle, J. Bryol. 10 (3): 270, 1979 (
*** Porella swartziana (F.Weber) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 407, 1877 (
** Porella triquetra (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 454, 1963 (
** Porella truncata J.S.Lou, Acta Phytotax. Sin. 18 (1): 119, 1980 (
** Porella ulophylla (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 92, 1944 (
** Porella undatorevoluta J.S.Lou, Acta Phytotax. Sin. 25 (6): 485, 1987 (
** Porella urceolata S.Hatt., J. Hattori Bot. Lab. 33: 66, 1970 (
** Porella urogea (C.Massal.) S.Hatt., J. Hattori Bot. Lab. 32: 349, 1969 (
** Porella vallis-gratiae (Gottsche) E.W.Jones, Trans. Brit. Bryol. Soc. 4 (3): 450, 1963 (
** Porella variabilis (Kashyap et R.S.Chopra) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 17, 1962 (
** Porella vernicosa Lindb., Acta Soc. Sci. Fenn. 10: 223, 1872 [1873] (
** Porella viridissima (Mitt.) Grolle, J. Hattori Bot. Lab. 36: 83, 1972 [1973] (
** Porella wataugensis (Sull.) Underw. ex M.Howe, Bull. Torrey Bot. Club 24 (11): 519, 1897 (
by M.A.M. Renner
The treatment of Radulaceae follows
*** Radula Dumort., Commentat. Bot. (Dumortier): 112, 1822 (
*** subg. Amentuloradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1630, 2011 (
*** Radula amentulosa Mitt., Bonplandia 9 (24): 367, 1861 (
*** Radula aneurismalis (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 262, 1845 (
** Radula ceylanica K.Yamada, J. Jap. Bot. 50 (12): 373, 1975 (
*** Radula fissifolia Steph., Sp. Hepat. (Stephani) 6: 507, 1924 (
*** Radula formosa (C.F.W.Meissn. ex Spreng.) Nees, Syn. Hepat. 2: 258, 1845 (
*** Radula helix (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 260, 1845 (
*** Radula hicksiae K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 191, 1984 (
*** Radula iwatsukii K.Yamada, J. Hattori Bot. Lab. 45: 275, 1979 (
** Radula morobeana K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 358, 1989 (
** Radula multiamentula E.A.Hodgs., Rec. Domin. Mus. 4 (11): 122, 1962 (
*** Radula ornata E.A.Br. et Pócs, Telopea 9 (3): 436, 2001 (
*** Radula physoloba Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 255, 1843 (
*** Radula pseudoscripta M.A.M.Renner, New Zealand J. Bot. 44 (3): 340, 2006 (
** Radula queenslandica K.Yamada, J. Hattori Bot. Lab. 62: 192, 1987 (
*** Radula scariosa Mitt., Bonplandia 9 (24): 367, 1861 (
*** Radula splendida M.A.M.Renner et Devos, Nova Hedwigia 90 (1/2): 113, 2010 (
* Radula squarrosa K.Yamada, J. Jap. Bot. 65 (1): 1, 1990 (
*** Radula thiersiae K.Yamada, J. Hattori Bot. Lab. 62: 198, 1987 (
*** Radula uvifera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 258, 1845 (
* Radula vagans Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 85, 1911 (
** Radula verrucosa K.Yamada, J. Hattori Bot. Lab. 45: 277, 1979 (
*** subg. Cladoradula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 315, 1885 (
** Radula auriculata Steph., Bull. Herb. Boissier 5 (2): 105, 1897 (
* Radula bipinnata Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 166, 1863 (
*** Radula boryana (F.Weber) Nees ex Mont., Ann. Sci. Nat. Bot. (sér. 2) 18: 13, 1842 (
*** Radula campanigera Mont., London J. Bot. 3: 634, 1844 (
*** Radula campanigera subsp. obiensis (S.Hatt.) K.Yamada, J. Hattori Bot. Lab. 45: 309, 1979 (
** Radula chinensis Steph., Nuovo Giorn. Bot. Ital. (n.ser.) 13 (4): 355, 1906 (
*** Radula gottscheana Taylor, London J. Bot. 5: 374, 1846 (
*** Radula perrottetii Gottsche, Hedwigia 23 (10): 154, 1884 (
** Radula tenax Lindb., Acta Soc. Sci. Fenn. 10: 492, 1875 (
*** subg. Dactyloradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1630, 2011 (
*** Radula brunnea Steph., Sp. Hepat. (Stephani) 4: 232, 1910 (
*** subg. Metaradula R.M.Schust., Phytologia 56 (2): 69, 1984 (
** Radula acuminata Steph., Sp. Hepat. (Stephani) 4: 230, 1910 (
** Radula aguirrei R.M.Schust., Phytotaxa 202 (1): 70, 2015 (
** Radula anisotoma M.A.M.Renner, PhytoKeys 27: 30, 2013 (
*** Radula assamica Steph., Hedwigia 23 (10): 151, 1884 (
*** Radula australiana K.Yamada, J. Hattori Bot. Lab. 51: 323, 1982 (
*** Radula buccinifera (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845 (
*** Radula demissa M.A.M.Renner, PhytoKeys 27: 53, 2013 (
** Radula evansii Castle, Ann. Bryol. 11: 37, 1938 (
** Radula flaccida Lindenb. et Gottsche, Syn. Hepat. 5: 726, 1847 (
** Radula flaccida var. brachycalyx Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 321, 1885 (
*** Radula forficata M.A.M.Renner, Austral. Syst. Bot. 26 (4): 307, 2013 (
** Radula grevilleana Taylor, Ann. Mag. Nat. Hist. 20 (135): 380, 1847 (
*** Radula imposita M.A.M.Renner, PhytoKeys 27: 65, 2013 (
** Radula jovetiana K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 193, 1984 (
** Radula kilgourii M.A.M.Renner, Austral. Syst. Bot. 26 (4): 313, 2013 (
** Radula loriana Castle, J. Hattori Bot. Lab. 21: 6, 1959 (
*** Radula mammosa Spruce, Mem. Torrey Bot. Club 1 (3): 127, 1890 (
*** Radula mittenii Steph., Hedwigia 23 (10): 148, 1884 (
*** Radula myriopoda M.A.M.Renner, Austral. Syst. Bot. 26 (4): 323, 2013 (
*** Radula notabilis M.A.M.Renner, PhytoKeys 27: 77, 2013 (
** Radula nymannii Steph., Sp. Hepat. (Stephani) 4: 229, 1910 (
** Radula protensa Lindenb., Bot. Zeitung (Berlin) 6 (25): 462, 1848 (
** Radula protensa var. erectilobula Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 247, 1893 (
** Radula pseudoflaccida E.W.Jones, J. Bryol. 9 (4): 501, 1977 [1978] (
** Radula psychosis M.A.M.Renner, Austral. Syst. Bot. 26 (4): 328, 2013 (
*** Radula ratkowskiana K.Yamada, J. Jap. Bot. 59 (3): 94, 1984 (
*** Radula robinsonii Steph., Sp. Hepat. (Stephani) 4: 214, 1910 (
** Radula stenocalyx Mont., Ann. Sci. Nat. Bot. (sér. 4) 3 (5): 315, 1855 (
*** Radula strangulata Hook.f. et Taylor, London J. Bot. 5: 377, 1846 (
* Radula tjibodensis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 7 (1): 53, 1888 (
*** Radula ventricosa Steph., Sp. Hepat. (Stephani) 4: 187, 1910 (
*** Radula yanoella R.M.Schust., Phytologia 56 (2): 72, 1984 (
*** subg. Odontoradula K.Yamada, J. Hattori Bot. Lab. 45: 209, 1979 (
** Radula acuta Mitt., Fl. vit.: 410, 1871 [1873] (
** Radula allisonii Castle, Rev. Bryol. Lichénol. 31 (3/4): 148, 1962 [1963] (
** Radula amoena Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 192, 1931 (
** Radula anceps Sande Lac., Ned. Kruidk. Arch. 3: 419, 1854 [1855] (
* Radula crenulata Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 21, 1890 (
*** Radula cuspidata Steph., Sp. Hepat. (Stephani) 4: 156, 1910 (
*** Radula decora Gottsche, Hedwigia 23 (10): 145, 1884 (
** Radula emarginata K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 352, 1989 (
** Radula kojana Steph., Bull. Herb. Boissier 5 (2): 105, 1897 (
*** Radula lacerata Steph., Rev. Bryol. 35 (2): 33, 1908 (
*** Radula novae-hollandiae Hampe, Nov. Stirp. Pug. 7: 24, 1838 (
*** Radula ocellata K.Yamada, J. Hattori Bot. Lab. 45: 209, 1979 (
*** Radula plicata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 154, 1854 (
** Radula pugioniformis M.A.M.Renner, PhytoKeys 27: 84, 2013 (
*** Radula pulchella Mitt., Hedwigia 23 (10): 149, 1884 (
*** Radula retroflexa Taylor, London J. Bot. 5: 378, 1846 (
*** Radula tasmanica Steph., Sp. Hepat. (Stephani) 4: 212, 1910 (
*** Radula weymouthiana Steph., Sp. Hepat. (Stephani) 4: 190, 1910 (
*** subg. Radula
*** Radula acutiloba Steph., Hedwigia 28 (4): 271, 1889 (
** Radula appressa Mitt., Philos. Trans. 168: 397, 1879 (
*** Radula aquilegia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 260, 1845 (
** Radula australis Austin, Bot. Bull. (Hanover) 1 (7): 32, 1876 (
** Radula borneensis Steph., Sp. Hepat. (Stephani) 4: 209, 1910 (
** Radula caduca K.Yamada, J. Hattori Bot. Lab. 45: 225, 1979 (
*** Radula carringtonii J.B.Jack, Flora 64 (25): 385, 1881 (
*** Radula complanata (L.) Dumort., Syll. Jungerm. Europ.: 38, 1831 (
** Radula constricta Steph., Sp. Hepat. (Stephani) 6: 506, 1924 (
** Radula evelynae K.Yamada, J. Jap. Bot. 50 (4): 115, 1975 (
** Radula fendleri Gottsche, Hedwigia 23 (10): 146, 1884 (
*** Radula grandis Steph., J. Linn. Soc., Bot. 29 (201): 271, 1892 (
** Radula japonica Gottsche, Hedwigia 23 (10): 152, 1884 (
** Radula javanica Gottsche, Syn. Hepat. 2: 257, 1845 (
*** Radula jonesii Bouman, Dirkse et K.Yamada, J. Bryol. 15 (1): 161, 1988 (
*** Radula lindenbergiana Gottsche ex C.Hartm., Handb. Skand. fl. (ed.9): 98, 1864 (
** Radula madagascariensis Gottsche, Abh. Naturwiss. Vereins Bremen 7: 349, 1882 (
** Radula marojezica E.W.Jones, J. Bryol. 17 (2): 307, 1992 (
* Radula multiflora Gottsche ex Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 20, 1890 (
** Radula novoguineensis K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 360, 1989 (
** Radula obconica Sull., Manual (Gray): 688, 1848 (
** Radula obtusiloba Steph., Bull. Herb. Boissier 5 (2): 105, 1897 (
** Radula obtusiloba subsp. polyclada (A.Evans) S.Hatt., J. Hattori Bot. Lab. 29: 275, 1966 (
* Radula oceania Castle, Rev. Bryol. Lichénol. 33 (3/4): 390, 1965 (
** Radula oreopsis M.A.M.Renner, Telopea 17: 123, 2014 (
** Radula portoricensis Steph., Hedwigia 27 (11/12): 298, 1888 (
*** Radula prolifera Arnell, Ark. Bot. 13 (2): 12, 1913 (
*** Radula quadrata Gottsche, Syn. Hepat. 2: 255, 1845 (
*** Radula reflexa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 255, 1843 (
** Radula sharpii K.Yamada, J. Jap. Bot. 60 (9): 260, 1985 (
** Radula sumatrana Steph., Sp. Hepat. (Stephani) 4: 204, 1910 (
** Radula tokiensis Steph., Hedwigia 23 (10): 150, 1884 (
*** Radula van-zantenii K.Yamada, J. Hattori Bot. Lab. 45: 260, 1979 (
** Radula varilobula Castle, J. Hattori Bot. Lab. 21: 19, 1959 (
*** Radula wichurae Steph., Sp. Hepat. (Stephani) 4: 168, 1910 (
*** subg. Volutoradula Devos, M.A.M.Renner, Gradst., A.J.Shaw et Vanderp., Taxon 60 (6): 1629, 2011 (
** Radula ankefinensis Gottsche, Hedwigia 23 (10): 152, 1884 (
** Radula antilleana Castle, J. Hattori Bot. Lab. 21: 48, 1959 (
** Radula comorensis Steph., Hedwigia 23 (9): 132, 1884 (
** Radula cubensis K.Yamada, J. Hattori Bot. Lab. 54: 241, 1983 (
** Radula diversifolia Steph., Sp. Hepat. (Stephani) 4: 212, 1910 (
*** Radula eggersii K.Yamada, J. Hattori Bot. Lab. 82: 339, 1997 (
** Radula episcia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 318, 1885 (
** Radula floridana Castle, Rev. Bryol. Lichénol. 36 (1/2): 1, 1968 [1969] (
** Radula fulvifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845 (
*** Radula hastata Steph., Sp. Hepat. (Stephani) 4: 163, 1910 (
* Radula holstiana Steph., Bot. Jahrb. Syst. 20 (3): 320, 1895 (
*** Radula holtii Spruce, J. Bot. 25: 209, 1887 (
** Radula husnotii Castle, J. Hattori Bot. Lab. 21: 45, 1959 (
** Radula inflexa Gottsche, Hedwigia 23 (10): 148, 1884 (
** Radula kegelii Gottsche ex Steph., Hedwigia 23 (10): 152, 1884 (
** Radula macroloba Steph., Bull. Soc. Roy. Bot. Belgique 31: 121, 1892 (
** Radula mazarunensis K.Yamada, Trop. Bryol. 1: 38, 1989 (
** Radula mexicana Lindenb. et Gottsche, Mexik. Leverm.: 150, 1863 (
** Radula microloba Gottsche, Syn. Hepat. 2: 259, 1845 (
** Radula neotropica Castle, J. Hattori Bot. Lab. 21: 31, 1959 (
*** Radula nudicaulis Steph., Sp. Hepat. (Stephani) 4: 174, 1910 (
* Radula nudicaulis var. delicatula P.Allorge et V.Allorge, Rev. Bryol. Lichénol. 19 (1/2): 106, 1950 (
** Radula pocsii K.Yamada, J. Hattori Bot. Lab. 54: 245, 1983 (
** Radula recubans Taylor, London J. Bot. 5: 376, 1846 (
** Radula saccatiloba Steph., Hedwigia 23 (8): 129, 1884 (
** Radula schaefer-verwimpii K.Yamada, J. Jap. Bot. 65 (1): 3, 1990 (
** Radula schofieldiana K.Yamada, J. Hattori Bot. Lab. 82: 337, 1997 (
** Radula stipatiflora Steph., Sp. Hepat. (Stephani) 4: 159, 1910 (
** Radula striata Mitt., Hedwigia 23 (10): 155, 1884 (
** Radula subinflata Lindenb. et Gottsche, Syn. Hepat. 5: 724, 1847 (
** Radula sullivantii Austin, Hepat. bor.-amer.: 22, 1873 (
** Radula tenera Mitt., Hedwigia 23 (10): 149, 1884 (
*** Radula voluta Taylor, Syn. Hepat. 2: 255, 1845 (
Incertae sedis
*** Radula acutangula Steph., Bull. Herb. Boissier 5 (10): 848, 1897 (
** Radula angulata Steph., Hedwigia 23 (8): 114, 1884 (
** Radula bogotensis Steph., Hedwigia 23 (8): 115, 1884 (
*** Radula bolanderi Gottsche, Hedwigia 23 (10): 145, 1884 (
** Radula boninensis Furuki et K.Yamada, J. Jap. Bot. 61 (10): 312, 1986 (
** Radula brasilica K.Yamada, J. Hattori Bot. Lab. 74: 35, 1993 (
** Radula caespitosa Steph., Hedwigia 27 (3/4): 107, 1888 (
** Radula campanulata Lindenb. et Gottsche, Syn. Hepat. 2: 256, 1845 (
** Radula castlei Grolle, Bryologist 73 (4): 662, 1970 (
*** Radula cavifolia Hampe ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 259, 1845 (
** Radula cochabambaensis K.Yamada, J. Hattori Bot. Lab. 74: 37, 1993 (
** Radula conferta Lindenb. et Gottsche, Syn. Hepat. 5: 729, 1847 (
*** Radula cordata Mitt., Fl. vit.: 410, 1871 [1873] (
** Radula costaricensis Gottsche, J. Bot. 15: 226, 1877 (
*** Radula curvilobula M.L.So, J. Hattori Bot. Lab. 98: 176, 2005 (
* Radula decurrens Mitt., Fl. vit.: 410, 1871 [1873] (
** Radula densifolia Castle, Rev. Bryol. Lichénol. 33 (3/4): 385, 1965 (
** Radula diaphana K.I.Goebel, Organogr. Pfl., ed. 2, 2 (1): 677, 1915 (
** Radula dolabrata K.Yamada, J. Jap. Bot. 60 (9): 257, 1985 (
** Radula elliottii Castle, J. Hattori Bot. Lab. 21: 12, 1959 (
** Radula falcata Steph., Hedwigia 23 (8): 115, 1884 (
** Radula fauriana Steph., Sp. Hepat. (Stephani) 4: 207, 1910 (
** Radula fernandezana Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 84, 1911 (
** Radula flavifolia (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 259, 1845 (
** Radula fujitae Furuki, Bryol. Res. 9 (5): 143, 2007 (
** Radula galapagona Steph., Sp. Hepat. (Stephani) 4: 176, 1910 (
** Radula gedena Gottsche, Hedwigia 23 (10): 146, 1884 (
** Radula gracilis Mitt., Hedwigia 23 (10): 147, 1884 (
** Radula gradsteinii K.Yamada, Trop. Bryol. 1: 37, 1989 (
** Radula grandifolia Steph., Sp. Hepat. (Stephani) 4: 184, 1910 (
** Radula grollei K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 379, 1989 (
** Radula guyanensis K.Yamada, Trop. Bryol. 1: 38, 1989 (
** Radula hattorii K.Yamada, J. Jap. Bot. 60 (9): 259, 1985 (
*** Radula hawaiica M.L.So, J. Hattori Bot. Lab. 98: 177, 2005 (
* Radula hedingeri K.I.Goebel, Ann. Jard. Bot. Buitenzorg 7 (1): 51, 1888 (
*** Radula inouei K.Yamada, J. Hattori Bot. Lab. 45: 262, 1979 (
*** Radula involvens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 325, 1885 (
*** Radula iwatsukiana K.Yamada, J. Hattori Bot. Lab. 58: 114, 1985 (
*** Radula jamaicensis Pearson, Ann. Bryol. 4: 103, 1931 (
** Radula jamesonii Taylor, London J. Bot. 5: 375, 1846 (
** Radula kinabaluensis K.Yamada, Misc. Bryol. Lichenol. 6 (6): 97, 1973 (
*** Radula kitagawae K.Yamada, J. Hattori Bot. Lab. 58: 116, 1985 (
** Radula koponenii K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 364, 1989 (
** Radula kurzii Steph., Hedwigia 23 (10): 153, 1884 (
** Radula laxiramea Steph., Sp. Hepat. (Stephani) 4: 178, 1910 (
** Radula leiboldii Steph., Hedwigia 23 (8): 116, 1884 (
** Radula lewisii K.Yamada, J. Hattori Bot. Lab. 74: 39, 1993 (
*** Radula ligula Steph., Sp. Hepat. (Stephani) 4: 228, 1910 (
*** Radula lingulata Gottsche, Syn. Hepat. 2: 260, 1845 (
** Radula longiloba K.Yamada, J. Hattori Bot. Lab. 54: 243, 1983 (
* Radula longispica Steph., Sp. Hepat. (Stephani) 4: 183, 1910 (
*** Radula marginata Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 261, 1845 (
*** Radula mauiensis M.L.So, J. Hattori Bot. Lab. 98: 178, 2005 (
** Radula microlobula Castle, J. Hattori Bot. Lab. 21: 35, 1959 (
** Radula minutilobula K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 377, 1989 (
** Radula mizutanii K.Yamada, J. Jap. Bot. 48 (5): 134, 1973 (
*** Radula nigra Pearson, J. Linn. Soc., Bot. 46 (305): 31, 1922 (
** Radula nilgiriensis Udar et D.Kumar, J. Indian Bot. Soc. 61: 177, 1982 (
** Radula norrisii K.Yamada et Piippo, Ann. Bot. Fenn. 26 (4): 374, 1989 (
** Radula novivrieseana K.Yamada, J. Hattori Bot. Lab. 51: 326, 1982 (
*** Radula novocaledonica Hürl. et K.Yamada, J. Jap. Bot. 54 (8): 238, 1979 (
*** Radula novocaledoniensis K.Yamada, J. Hattori Bot. Lab. 58: 120, 1985 (
** Radula obovata Castle, J. Hattori Bot. Lab. 21: 16, 1959 (
** Radula obscura Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (
** Radula okamurana Steph., Sp. Hepat. (Stephani) 4: 209, 1910 (
** Radula onraedtii K.Yamada, Misc. Bryol. Lichenol. 8 (6): 113, 1979 (
** Radula opaciuscula (Spruce) Castle, J. Hattori Bot. Lab. 21: 22, 1959 (
** Radula ovalilobula K.Yamada, J. Hattori Bot. Lab. 45: 257, 1979 (
** Radula oyamensis Steph., Hedwigia 23 (10): 149, 1884 (
* Radula paganii Castle, J. Hattori Bot. Lab. 21: 33, 1959 (
*** Radula pallens (Sw.) Nees ex Mont., Voy. Amér. Mérid. 7 (2): 71, 1839 (
*** Radula pandei Udar et Dh.Kumar, Lindbergia 9 (2): 133, 1983 (
** Radula patens K.Yamada, Cryptog. Bryol. Lichénol. 5 (1/2): 197, 1984 (
** Radula peruviana K.Yamada, Beih. Nova Hedwigia 88: 79, 1987 (
** Radula philippinensis K.Yamada, J. Hattori Bot. Lab. 45: 299, 1979 (
* Radula pinnulata Mitt., Fl. vit.: 410, 1871 [1873] (
** Radula pseudostachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 319, 1885 (
** Radula punctata Steph., Hedwigia 23 (8): 135, 1884 (
** Radula pusilla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 320, 1885 (
*** Radula rhombiloba Steph., Sp. Hepat. (Stephani) 4: 204, 1910 (
* Radula rupicola K.Yamada, J. Hattori Bot. Lab. 58: 124, 1985 (
** Radula santacruziana K.Yamada et Gradst., Trop. Bryol. 4: 67, 1991 (
** Radula silvestris Gottsche, Abh. Naturwiss. Vereins Bremen 7: 349, 1882 (
** Radula sinskeana K.Yamada, J. Hattori Bot. Lab. 74: 41, 1993 (
** Radula sinuata Gottsche ex Steph., Sp. Hepat. (Stephani) 4: 161, 1910 (
* Radula socorana Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 475, 1947 (
** Radula sonsonensis Steph., Sp. Hepat. (Stephani) 4: 201, 1910 (
** Radula stellatogemmipara C.Gao et Y.H.Wu, Nova Hedwigia 80 (1/2): 239, 2005 (
** Radula subsimplex Steph., Hedwigia 23 (8): 130, 1884 (
** Radula subsquarrosa S.W.Arnell, Ark. Bot. (n.ser.) 4 (1): 15, 1957 (
** Radula tabularis Steph., Hedwigia 23 (9): 131, 1884 (
** Radula taylorii Steph., Hedwigia 23 (9): 133, 1884 (
** Radula tectiloba Steph., Hedwigia 27 (11/12): 298, 1888 (
** Radula tenuis K.Yamada, J. Hattori Bot. Lab. 54: 247, 1983 (
** Radula underwoodii Castle, J. Hattori Bot. Lab. 21: 37, 1959 (
** Radula venezuelensis K.Yamada, Misc. Bryol. Lichenol. 9 (6): 122, 1982 (
*** Radula vieillardii Gottsche, Hedwigia 23 (10): 150, 1884 (
** Radula visianica C.Massal., Ann. Bot. (Rome) 1 (4): 298, 1904 (
** Radula vrieseana Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 305, 1864 (
** Radula wrightii Castle, J. Hattori Bot. Lab. 21: 15, 1959 (
** Radula xalapensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 56, 1836 (
** Herzogianthus R.M.Schust., J. Hattori Bot. Lab. 23: 71, 1960 [1961] (
* Herzogianthus sanguineus R.M.Schust., Phytologia 56 (7): 457, 1985 (
*** Herzogianthus vaginatus (Herzog) R.M.Schust., J. Hattori Bot. Lab. 23: 71, 1960 [1961] (
** Neotrichocolea S.Hatt., J. Hattori Bot. Lab. 2: 9, 1947 [1948] (
** Neotrichocolea bissetii (Mitt.) S.Hatt., J. Hattori Bot. Lab. 2: 10, 1947 [1948] (
** Trichocoleopsis S.Okamura, Bot. Mag. (Tokyo) 25 (293): 159, 1911 (
** Trichocoleopsis sacculata (Mitt.) S.Okamura, Bot. Mag. (Tokyo) 25 (293): 159, 1911 (
by L. Söderström
Ptilidiaceae was recently studied molecularly by
*** Ptilidium Nees, Naturgesch. Eur. Leberm. 1: 95, 1833 (
*** Ptilidium californicum (Austin) Pearson, List. Canad. Hepat.: 7, 1890 (
*** Ptilidium ciliare (L.) Hampe, Prod. fl. hercyn.: 76, 1836 (
*** Ptilidium pulcherrimum (Weber) Vain., Meddel. Soc. Fauna Fl. Fenn. 3: 88, 1878 (
by M. Nebel
A molecular phylogeny of Aneuraceae was recently published by
*** Aneura Dumort., Commentat. Bot. (Dumortier): 115, 1822 (
* Aneura amboinensis Steph., Bull. Herb. Boissier 7 (9): 678 (219), 1899 (
* Aneura augustae Steph., Sp. Hepat. (Stephani) 6: 430, 1923 (
* Aneura biflora Colenso, Trans. & Proc. New Zealand Inst. 17: 262, 1885 (
*** Aneura blasioides (Horik.) Furuki, J. Hattori Bot. Lab. 70: 311, 1991 (
** Aneura brasiliensis (Ångstr.) Steph., Hedwigia 32 (3): 137, 1893 (
* Aneura brevissima Steph., Sp. Hepat. (Stephani) 6: 21, 1917 (
** Aneura cerebrata Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 185, 1970 (
*** Aneura crateriformis Furuki et D.G.Long, J. Bryol. 18 (2): 281, 1994 (
* Aneura crinita C.Massal., Bull. Soc. Bot. Ital. 1917 (8/9): 81, 1917 (
** Aneura crumii L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 43, 2012 (
* Aneura densa Steph., Sp. Hepat. (Stephani) 6: 24, 1917 (
* Aneura denticulata Mitt. ex Thurn, Timehri 5: 222, 1886 (
** Aneura eachamensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 184, 1970 (
** Aneura erronea Steph., Sp. Hepat. (Stephani) 6: 20, 1917 (
** Aneura eskuchei Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 294, 1986 (
** Aneura gemmifera Furuki, J. Hattori Bot. Lab. 70: 309, 1991 (
* Aneura giangena Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 190, 1970 (
** Aneura gibbsiana Steph., Sp. Hepat. (Stephani) 6: 28, 1917 (
** Aneura glaucescens Steph., Sp. Hepat. (Stephani) 6: 28, 1917 (
* Aneura goebeliana Steph., Sp. Hepat. (Stephani) 6: 28, 1917 (
*** Aneura hirsuta Furuki, J. Hattori Bot. Lab. 70: 317, 1991 (
* Aneura hunsteinii Steph., Sp. Hepat. (Stephani) 6: 431, 1923 (
** Aneura imbricata Colenso, Trans. & Proc. New Zealand Inst. 16: 359, 1884 (
** Aneura kaguaensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 190, 1970 (
** Aneura keniae Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916 (
* Aneura latemultifida Steph., Sp. Hepat. (Stephani) 6: 431, 1923 (
** Aneura latissima Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 544, 1885 (
* Aneura ledermannii Steph., Sp. Hepat. (Stephani) 6: 431, 1923 (
** Aneura macrostachya Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 550, 1885 (
*** Aneura marianensis Furuki, Bryologist 97 (1): 87, 1994 (
*** Aneura maxima (Schiffn.) Steph., Bull. Herb. Boissier 7 (10): 760 (270), 1899 (
*** Aneura mirabilis (Malmb.) Wickett et Goffinet, Bot. J. Linn. Soc. 156 (1): 11, 2008 (
** Aneura novaecaledoniae R.M.Schust., Phytologia 56 (7): 451, 1985 (
*** Aneura novaguineensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 189, 1970 (
* Aneura nymannii Steph., Sp. Hepat. (Stephani) 6: 35, 1917 (
** Aneura pellucida Colenso, Trans. & Proc. New Zealand Inst. 18: 252, 1886 (
*** Aneura pinguis (L.) Dumort., Syll. Jungerm. Europ.: 86, 1831 (
** Aneura polyantha Colenso, Trans. & Proc. New Zealand Inst. 17: 262, 1885 (
** Aneura punctata Colenso, Trans. & Proc. New Zealand Inst. 18: 254, 1886 (
** Aneura rodwayi Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 94 (2): 188, 1970 (
* Aneura roraimensis Steph., Trans. Linn. Soc. London, Bot. 6 (1): 94, 1901 (
** Aneura rotangicola Steph., Sp. Hepat. (Stephani) 6: 432, 1923 (
* Aneura serrulata Gottsche ex Steph., Sp. Hepat. (Stephani) 6: 42, 1917 (
** Aneura sharpii Inoue et N.G.Mill., Bull. Natl. Sci. Mus. Tokyo, B 11 (3): 96, 1985 (
* Aneura singalangana (Schiffn.) Steph., Bull. Herb. Boissier 7 (10): 751 (261), 1899 (
** Aneura subcanaliculata R.M.Schust., J. Hattori Bot. Lab. 67: 60, 1989 (
* Aneura subledermannii Steph., Sp. Hepat. (Stephani) 6: 432, 1923 (
* Aneura subtenerrima Steph., Sp. Hepat. (Stephani) 6: 432, 1923 (
* Aneura vincentina Steph., Sp. Hepat. (Stephani) 6: 45, 1917 (
** Lobatiriccardia (Mizut. et S.Hatt.) Furuki, J. Hattori Bot. Lab. 70: 319, 1991 (
*** Lobatiriccardia alterniloba (Hook.f. et Taylor) Furuki, J. Hattori Bot. Lab. 70: 319, 1991 (
** Lobatiriccardia alterniloba var. gigantea (Steph.) Nebel, Phytotaxa 81 (1): 10, 2013 (
** Lobatiriccardia alterniloba var. robusta (Rodway) Nebel, Phytotaxa 81 (1): 10, 2013 (
*** Lobatiriccardia athertonensis (Hewson) Furuki, J. Hattori Bot. Lab. 70: 319, 1991 (
*** Lobatiriccardia coronopus (De Not.) Furuki, J. Hattori Bot. Lab. 100: 90, 2006 (
** Lobatiriccardia coronopus subsp. australis (R.M.Schust.) Nebel, Preussing, Schäf.-Verw. et D.Quandt, Taxon 59 (5): 1434, 2010 (
** Lobatiriccardia oberwinkleri Nebel, Preussing, Schäf.-Verw. et D.Quandt, Taxon 59 (5): 1435, 2010 (
** Lobatiriccardia subaquatica (R.M.Schust.) Nebel, Phytotaxa 81 (1): 10, 2013 (
** Lobatiriccardia verdoornioides Nebel, Preussing, Schäf.-Verw. et D.Quandt, Taxon 59 (5): 1437, 2010 (
*** Lobatiriccardia yakusimensis (S.Hatt.) Furuki, J. Hattori Bot. Lab. 70: 321, 1991 (
*** Lobatiriccardia yunnanensis Furuki et D.G.Long, J. Bryol. 29 (3): 161, 2007 (
*** Riccardia Gray, Nat. Arr. Brit. Pl. 1: 679, 1821 (
** Riccardia pectinata var. fasciculata (Steph.) Hürl., Bauhinia 5 (4): 208, 1976 (
** subg. Arceoneura Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 204, 1972 (
** Riccardia marionensis R.M.Schust., J. Hattori Bot. Lab. 67: 65, 1989 (
*** Riccardia prehensilis (Hook.f. et Taylor) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 255, 1885 (
** subg. Corioneura Furuki, J. Hattori Bot. Lab. 70: 394, 1991 (
** Riccardia argentolimbata Hewson et Grolle, J. Hattori Bot. Lab. 29: 70, 1966 (
*** Riccardia hattorii Furuki, J. Hattori Bot. Lab. 75: 257, 1994 (
** subg. Hyaloneura R.M.Schust., Phytologia 56 (7): 452, 1985 (
* Riccardia albomarginata (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 53, 1898 (
** Riccardia canaliculata (Nees) Kuntze, Revis. Gen. Pl. 2: 838, 1891 (
** Riccardia pindensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 87, 1970 (
** subg. Lophoneura Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 218, 1972 (
** Riccardia fuegiensis C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 255, 1885 (
** subg. Neoneura Furuki, J. Hattori Bot. Lab. 70: 385, 1991 (
*** Riccardia spongiosa Furuki, J. Hattori Bot. Lab. 70: 385, 1991 (
** subg. Phycaneura R.M.Schust., J. Hattori Bot. Lab. 26: 294, 1963 (
** Riccardia aequicellularis (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 79, 1970 (
** Riccardia asperulata R.M.Schust., J. Hattori Bot. Lab. 27: 209, 1964 (
** subg. Riccardia
*** Riccardia aeruginosa Furuki, J. Hattori Bot. Lab. 70: 345, 1991 (
*** Riccardia arcuata Furuki, J. Hattori Bot. Lab. 70: 361, 1991 (
*** Riccardia chamedryfolia (With.) Grolle, Trans. Brit. Bryol. Soc. 5 (4): 772, 1969 (
*** Riccardia cochleata (Hook.f. et Taylor) Kuntze, Revis. Gen. Pl. 2: 838, 1891 (
*** Riccardia eriocaula (Hook.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 256, 1885 (
*** Riccardia flavovirens Furuki, J. Hattori Bot. Lab. 70: 333, 1991 (
*** Riccardia fruticosa (Steph.) Furuki, Nat. Hist. Res. 5 (1): 1, 1998 (
*** Riccardia glauca Furuki, J. Hattori Bot. Lab. 70: 352, 1991 (
** Riccardia kodamae Mizut. et S.Hatt., J. Hattori Bot. Lab. 18: 57, 1957 (
*** Riccardia latifrons (Lindb.) Lindb., Acta Soc. Sci. Fenn. 10: 513, 1875 (
** Riccardia latifrons subsp. arctica R.M.Schust. et Damsh., J. Hattori Bot. Lab. 62: 303, 1987 (
** Riccardia latifrons var. miyakeana (Schiffn.) Furuki, J. Hattori Bot. Lab. 70: 375, 1991 (
** Riccardia nagasakiensis (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (
*** Riccardia palmata (Hedw.) Carruth., J. Bot. 3: 302, 1865 (
** Riccardia planiflora (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (
** Riccardia planiflora var. aequatorialis Furuki, Nat. Hist. Res. 4 (2): 77, 1997 (
** Riccardia pseudodendroceros R.M.Schust., Phytologia 56 (7): 452, 1985 (
*** Riccardia pumila Furuki, J. Hattori Bot. Lab. 70: 361, 1991 (
** Riccardia pusilla Grolle, J. Jap. Bot. 41 (8): 231, 1966 (
*** Riccardia subalpina Furuki, J. Hattori Bot. Lab. 70: 367, 1991 (
** Riccardia tamariscina (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 58, 1898 (
*** Riccardia vitrea Furuki, J. Hattori Bot. Lab. 70: 327, 1991 (
** sect. Alcicornia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 17, 1972 (
** Riccardia alcicornis (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (
** Riccardia conimitra (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 156, 1921 (
** Riccardia corralensis (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 143, 1921 (
*** Riccardia furtiva E.A.Br. et Braggins, J. Hattori Bot. Lab. 66: 35, 1989 (
** Riccardia fuscobrunnea (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 152, 1921 (
** Riccardia longioleata Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 30, 1972 (
*** Riccardia multicorpora E.A.Br., J. Hattori Bot. Lab. 66: 40, 1989 (
* Riccardia umida E.A.Br., J. Hattori Bot. Lab. 66: 38, 1989 (
** sect. Crassantia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 41, 1972 (
*** Riccardia aequitexta (Steph.) E.A.Br., J. Hattori Bot. Lab. 66: 78, 1989 (
*** Riccardia alba (Colenso) E.A.Br., J. Hattori Bot. Lab. 66: 66, 1989 (
** Riccardia amnicola Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 92, 1972 (
** Riccardia bipinnatifida (Colenso) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 93, 1970 (
** Riccardia calva (Schiffn.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 134, 1921 (
*** Riccardia colensoi (Steph.) W.Martin, Trans. & Proc. Roy. Soc. New Zealand 78 (4): 499, 1950 (
*** Riccardia crassa (Schwägr.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885 (
** Riccardia crassicrispa (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 194, 1921 (
** Riccardia diderma Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 56, 1972 (
** Riccardia diversiflora A.Evans, Trans. Connecticut Acad. Arts 25 (2): 167, 1921 (
** Riccardia diversiflora subsp. paucigyna R.M.Schust., J. Hattori Bot. Lab. 67: 97, 1989 (
** Riccardia duriuscula Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 82, 1972 (
** Riccardia falsifloribunda Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 119, 1972 (
** Riccardia floribunda (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 182, 1921 (
** Riccardia fluvigena Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 73, 1972 (
** Riccardia georgiensis (Steph.) Hässel, Lindbergia 1 (1/2): 80, 1971 [1972] (
** Riccardia georgiensis subsp. sympodea R.M.Schust., J. Hattori Bot. Lab. 67: 83, 1989 (
*** Riccardia graeffei (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 118, 1970 (
*** Riccardia marginata (Colenso) Pearson, Univ. Calif. Publ. Bot. 10 (4): 309, 1923 (
*** Riccardia marginata var. pacifica Furuki, J. Hattori Bot. Lab. 70: 380, 1991 (
** Riccardia mycophora A.Evans, Trans. Connecticut Acad. Arts 25 (2): 175, 1921 (
** Riccardia negeri (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 172, 1921 (
** Riccardia nitida (Colenso) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 92, 1965 (
** Riccardia papillosa (C.Massal. et Steph.) Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 60, 1972 (
** Riccardia regularis (Steph.) Kühnem., Revista Centro Estud. Doct. Ci. Nat. 1: 171, 1937 (
** Riccardia rivularis Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 52, 1972 (
** Riccardia saxicola Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 137, 1972 (
** Riccardia spectabilis (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 140, 1921 (
** Riccardia spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885 (
** Riccardia tenax (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 186, 1921 (
** Riccardia tenerrima (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 164, 1921 (
** Riccardia theliophora Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 96, 1972 (
** sect. Pallidevirida Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 142, 1972 (
** Riccardia granulata (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 192, 1921 (
** Riccardia intercellula E.A.Br., J. Hattori Bot. Lab. 66: 58, 1989 (
** Riccardia opuntiiformis S.W.Arnell, Arch. Soc. Zool. Bot. Fenn. “Vanamo” 9: 52, 1954 (
** Riccardia pallidevirens (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 189, 1921 (
*** Riccardia pennata E.A.Br., J. Hattori Bot. Lab. 66: 53, 1989 (
*** Riccardia perspicua E.A.Br., J. Hattori Bot. Lab. 66: 55, 1989 (
** Riccardia xylophila Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 156, 1972 (
** sect. Riccardia
*** Riccardia australis (Lehm.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 110, 1970 (
** Riccardia autoica (Steph.) A.Evans, Trans. Connecticut Acad. Arts 25 (2): 159, 1921 (
** Riccardia breviala E.A.Br., J. Hattori Bot. Lab. 66: 46, 1989 (
** Riccardia breviramosa (Steph.) A.Evans, Nat. Hist. Juan Fernandez (Botany) 2 (20): 559, 1930 (
** Riccardia filicina (Colenso) E.A.Hodgs., Rec. Domin. Mus. 4 (11): 129, 1962 (
** Riccardia leptostachya A.Evans, Nat. Hist. Juan Fernandez (Botany) 2 (20): 570, 1930 (
** Riccardia lobulata (Colenso) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 92, 1965 (
** Riccardia mejlandii S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 30, 1958 (
*** Riccardia multifida (L.) Gray, Nat. Arr. Brit. Pl. 1: 684, 1821 (
*** Riccardia multifida subsp. decrescens (Steph.) Furuki, J. Hattori Bot. Lab. 70: 341, 1991 (
** Riccardia multifida subsp. synoica R.M.Schust., J. Hattori Bot. Lab. 62: 319, 1987 (
** Riccardia multioleata Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 195, 1972 (
*** Riccardia papulosa (Steph.) E.A.Br., J. Hattori Bot. Lab. 66: 49, 1989 (
** Riccardia patens Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 165, 1972 (
** Riccardia polyclada (Mitt.) Hässel, Bryologist 109 (1): 34, 2006 (
** Riccardia statensis Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 177, 1972 (
** Riccardia thaxteri A.Evans, Trans. Connecticut Acad. Arts 25 (2): 126, 1921 (
** Riccardia tristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 31, 1958 (
** subg. Spinella (Schiffn.) Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 214, 1972 (
** Riccardia spinulifera C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 254, 1885 (
** subg. Thornoneura Furuki, J. Hattori Bot. Lab. 70: 382, 1991 (
* Riccardia baumannii Hürl., Bauhinia 5 (4): 203, 1976 (
** Riccardia deguchii Furuki et K.T.Yong, Hikobia 16 (3): 285, 2013 (
*** Riccardia grossitexta (Steph.) Furuki, J. Hattori Bot. Lab. 70: 382, 1991 (
** Riccardia inconspicua (Steph.) Reeb et Bardat, Cryptog. Bryol. 35 (1): 61, 2014 (
** subg. Trichothallia Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 224, 1972 (
** Riccardia hyalitricha Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 4 (1): 225, 1972 (
Incertae sedis
** Riccardia aberrans (Steph.) Gradst., J. Hattori Bot. Lab. 45: 129, 1979 (
** Riccardia agumana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 82, 1970 (
** Riccardia algoides (Taylor) Meenks, J. Hattori Bot. Lab. 62: 168, 1987 (
*** Riccardia amazonica (Spruce) Schiffn. ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 129, 1979 (
*** Riccardia andina (Spruce) Herzog, Svensk Bot. Tidskr. 46 (1): 65, 1952 (
** Riccardia angustata Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 126, 1934 (
** Riccardia angustealata (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 111, 1970 (
** Riccardia angustissima (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (
** Riccardia aspera (Steph.) Grolle, J. Hattori Bot. Lab. 30: 117, 1967 (
* Riccardia baldwinii (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (
** Riccardia barbiflora (Steph.) Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (
** Riccardia bogotensis (Gottsche) Pagán, Bryologist 42 (1): 6, 1939 (
** Riccardia boliviensis (Steph.) Meenks, J. Hattori Bot. Lab. 62: 170, 1987 (
** Riccardia bongeriana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 101, 1970 (
** Riccardia brunnea (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (
** Riccardia calcarea (Steph.) Meenks, J. Hattori Bot. Lab. 62: 170, 1987 (
** Riccardia capillacea (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 6, 1985 (
** Riccardia capillacea var. dentata Meenks, J. Hattori Bot. Lab. 62: 170, 1987 (
** Riccardia cardotii (Steph.) Pandé et S.C.Srivast., Biol. Mem. 1 (1/2): 131, 1976 (
** Riccardia cataractarum (Spruce) Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 18, 1964 (
*** Riccardia cervicornis (Spruce) Herzog ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 129, 1979 (
** Riccardia changbaishanensis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981 (
** Riccardia chinensis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981 (
** Riccardia ciliolata (Spruce) Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 198, 1933 (
** Riccardia columbica (Steph.) Hässel ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 130, 1979 (
** Riccardia comata (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (
** Riccardia compacta (Steph.) S.W.Arnell, Bot. Not. 105: 141, 1952 (
** Riccardia comptonii (Pearson) H.A.Mill., Phytologia 47 (4): 323, 1981 (
** Riccardia costata (Steph.) Hürl., Bauhinia 5 (4): 206, 1976 (
*** Riccardia crassicaulis (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 8, 1985 (
** Riccardia crassiretis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898 (
** Riccardia crenulata Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898 (
** Riccardia crenuliformis R.M.Schust., J. Hattori Bot. Lab. 67: 72, 1989 (
* Riccardia decolyana Schiffn., J. Indian Bot. Soc. 38 (4): 538, 1959 [1960] (
** Riccardia densiramea (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (
** Riccardia devexa Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 12, 1964 (
** Riccardia diablotina (Spruce) Pagán, Bryologist 45 (4): 80, 1942 (
*** Riccardia digitiloba (Spruce) Pagán, Bryologist 42 (1): 6, 1939 (
** Riccardia dilatata (Spruce) Schäf.-Verw. et Pócs, Cryptog. Bryol. 31 (4): 389, 2010 (
* Riccardia diminuta Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 170, 1898 (
* Riccardia diminuta var. thermarum Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898 (
** Riccardia distans (Spruce) Pagán, Bryologist 45 (4): 80, 1942 (
** Riccardia elata (Steph.) Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 169, 1898 (
* Riccardia elata var. flaccida Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 170, 1898 (
* Riccardia elata var. intercedens Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 170, 1898 (
** Riccardia elegans (Steph.) Hürl., Bauhinia 5 (4): 196, 1976 (
** Riccardia elisabethae Thouvenot et Reeb, Telopea 17: 229, 2014 (
*** Riccardia emarginata (Steph.) K.G.Hell, Bol. Fac. Filos. Univ. São Paulo, Bot. 25: 100, 1969 (
** Riccardia erosa (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (1): 83, 1956 (
** Riccardia fastigiata (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (
** Riccardia fendleri (Steph.) Pagán, Bryologist 45 (4): 80, 1942 (
** Riccardia flaccida (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (
** Riccardia flaccidissima Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 167, 1898 (
** Riccardia flagellaris (A.Gepp) H.A.Mill., Phytologia 47 (4): 323, 1981 (
** Riccardia flagellifrons C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 209, 1981 (
** Riccardia fleischeri (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (
** Riccardia foliacea Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 8, 1985 (
** Riccardia formosensis (Steph.) Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 125, 1934 (
*** Riccardia fucoidea (Sw.) C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 256, 1885 (
** Riccardia geniana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 76, 1970 (
*** Riccardia glaziovii (Spruce) Meenks, J. Hattori Bot. Lab. 62: 173, 1987 (
** Riccardia gogolensis (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 102, 1970 (
** Riccardia gracilis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 295, 1963 (
** Riccardia grandiflora (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963 (
*** Riccardia grollei Furuki, Haussknechtia, Beih. 9: 139, 1999 (
** Riccardia grossidens (Steph.) Pagán, Bryologist 45 (4): 80, 1942 (
** Riccardia gunniana (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (
** Riccardia hamatiflora (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963 (
*** Riccardia hans-meyeri (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985 (
** Riccardia hans-meyeri var. dentata Meenks, J. Hattori Bot. Lab. 62: 173, 1987 (
** Riccardia hawaica (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963 (
** Riccardia hebridensis (Steph.) H.A.Mill., Phytologia 47 (4): 323, 1981 (
** Riccardia herzogiana (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985 (
** Riccardia heteroclada Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 175, 1898 (
* Riccardia hirtiflora (Steph.) Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 19, 1964 (
** Riccardia humilis (Gottsche) O.Yano, J. Hattori Bot. Lab. 56: 530, 1984 (
** Riccardia hyalina (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (
** Riccardia hydra Hürl., Bauhinia 5 (4): 210, 1976 (
** Riccardia hymenophylloides Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 175, 1898 (
* Riccardia hymenophylloides var. flaccida Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 176, 1898 (
*** Riccardia hymenophytoides (Spruce) Meenks, Beih. Nova Hedwigia 88: 101, 1987 (
** Riccardia hypipamensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 97, 1970 (
** Riccardia ibana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 78, 1970 (
*** Riccardia incurvata Lindb., Helsingf. Dagbl. 1878 (315, 18 Nov.): 2, 1878 (
** Riccardia innovans (Steph.) Pagán, Bryologist 45 (4): 80, 1942 (
** Riccardia insularis Schiffn., Deutsche Südpolar-Exped. 1901-1903, 8 (bot.) 1: 66, 1906 (
** Riccardia intricata (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (
** Riccardia jackii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898 (
* Riccardia jackii var. densa Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898 (
** Riccardia judithae Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 12, 1985 (
*** Riccardia jugata R.M.Schust., J. Hattori Bot. Lab. 62: 305, 1987 (
* Riccardia karstenii (Steph.) Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 177, 1898 (
** Riccardia laticostata (Spruce) Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 12, 1964 (
** Riccardia latifrondoides Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 168, 1898 (
** Riccardia lepidomitra (Spruce) Gradst., J. Hattori Bot. Lab. 45: 130, 1979 (
** Riccardia leptophylla (Spruce) Herzog, Svensk Bot. Tidskr. 46 (1): 65, 1952 (
** Riccardia leptothallus R.M.Schust., J. Hattori Bot. Lab. 67: 87, 1989 (
** Riccardia levieri Schiffn., Österr. Bot. Z. 49 (4): 130, 1899 (
** Riccardia lichenoides (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (
** Riccardia ligulata (Steph.) Pócs et Schäf.-Verw., Cryptog. Bryol. 31 (4): 390, 2010 (
** Riccardia lilliena (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (
** Riccardia limbata (Steph.) E.W.Jones, Trans. Brit. Bryol. Soc. 3 (1): 79, 1956 (
** Riccardia loefgrenii Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 14, 1964 (
** Riccardia longiflora (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 88, 1970 (
** Riccardia longispica (Steph.) Pearson, Forh. Vidensk.-Selsk. Kristiania 1892 (14): 4, 1893 (
** Riccardia loriana (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963 (
** Riccardia macdonaldiana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 93, 1970 (
** Riccardia macrantha (Pearson) H.A.Mill., Phytologia 47 (4): 324, 1981 (
*** Riccardia magnicellularis Furuki, J. Hattori Bot. Lab. 100: 93, 2006 (
*** Riccardia metzgeriiformis (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 295, 1963 (
** Riccardia microscopica (Nees) Kuntze, Revis. Gen. Pl. 2: 838, 1891 (
** Riccardia minuta (Steph.) W.Martin, Trans. & Proc. Roy. Soc. New Zealand 78 (4): 499, 1950 (
* Riccardia multifidoides Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 166, 1898 (
** Riccardia multispica (Steph.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 164, 1944 (
** Riccardia nadeaudii (Steph.) Hürl., Bauhinia 5 (4): 201, 1976 (
** Riccardia newellana (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 528, 1963 (
** Riccardia nigra (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (
** Riccardia nobilis (Steph.) Schiffn., Consp. Hepat. Arch. Ind.: 56, 1898 (
** Riccardia novo-amstelodamensis Schiffn., Deutsche Südpolar-Exped. 1901-1903, 8 (bot.) 1: 65, 1906 (
** Riccardia nudiflora (Steph.) Grolle, Bryophyt. Biblioth. 48: 130, 1995 (
** Riccardia obtusa S.W.Arnell, Bot. Not. 105: 142, 1952 (
** Riccardia obtusifrons (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (
** Riccardia omkaliensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 97, 1970 (
*** Riccardia pallida (Spruce) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 15, 1985 (
** Riccardia palmatifida (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (
* Riccardia palmatiformis Schiffn., J. Indian Bot. Soc. 38 (4): 538, 1959 [1960] (
** Riccardia papillata (Gottsche) Hässel ex Gradst. et Hekking, J. Hattori Bot. Lab. 45: 130, 1979 (
** Riccardia paramorum Meenks, J. Hattori Bot. Lab. 62: 176, 1987 (
*** Riccardia parasitans (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985 (
** Riccardia parvula Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898 (
** Riccardia pauciramea (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (
** Riccardia paulensis Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 16, 1964 (
** Riccardia pectinata (Austin) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963 (
** Riccardia pellucida Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (
** Riccardia pembaiensis (Steph.) Hürl., Bauhinia 5 (4): 205, 1976 (
** Riccardia pengagensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 108, 1970 (
** Riccardia perssonii S.C.Srivast. et Udar, Lindbergia 4 (1/2): 127, 1977 (
*** Riccardia philippinensis Furuki, J. Hattori Bot. Lab. 100: 94, 2006 (
** Riccardia phleganiana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 80, 1970 (
** Riccardia plana (Steph.) Hürl., Bauhinia 5 (4): 205, 1976 (
* Riccardia plana var. minor (Pearson) Hürl. ex H.A.Mill., H.Whittier et B.Whittier, Bryophyt. Biblioth. 25: 303, 1983 (
** Riccardia planifrons (Spruce) Pagán, Bryologist 45 (4): 80, 1942 (
*** Riccardia plumiformis (Spruce) Hässel ex Meenks, Beih. Nova Hedwigia 88: 101, 1987 (
** Riccardia plumosa (Mitt.) E.O.Campb., J. Roy. Soc. New Zealand 1 (1): 24, 1971 (
*** Riccardia poeppigiana (Lehm. et Lindenb.) Hässel ex Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985 (
** Riccardia porcina (Hewson) L.Söderstr., Phytotaxa 202 (1): 70, 2015 (
** Riccardia portoricensis (Steph.) Pagán, Bryologist 42 (1): 7, 1939 (
** Riccardia punahuina (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 526, 1963 (
** Riccardia ramosissima (Steph.) Grolle, Bryophyt. Biblioth. 48: 130, 1995 (
** Riccardia regina Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 17, 1985 (
*** Riccardia regnellii (Ångstr.) K.G.Hell, Bol. Fac. Filos. Univ. São Paulo, Bot. 25: 110, 1969 (
** Riccardia reyesiana Meenks, Acta Bot. Hung. 32 (1/4): 207, 1986 (
** Riccardia riccioides Pearson, Univ. Calif. Publ. Bot. 10 (4): 310, 1923 (
* Riccardia rigida Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898 (
** Riccardia robbinsii Hewson et Grolle, J. Hattori Bot. Lab. 29: 72, 1966 (
** Riccardia robusta (Steph.) H.A.Mill., Phytologia 47 (4): 324, 1981 (
** Riccardia rockii (Steph.) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 527, 1963 (
** Riccardia rupicola (Steph.) Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 99, 1970 (
** Riccardia russellii R.M.Schust., J. Hattori Bot. Lab. 67: 93, 1989 (
** Riccardia saccatiflora (Steph.) S.W.Arnell, Bot. Not. 105: 144, 1952 (
** Riccardia santapaui Udar et S.C.Srivast., Rev. Bryol. Lichénol. 39 (1): 155, 1973 (
*** Riccardia schwaneckei (Steph.) Pagán, Bryologist 42 (1): 7, 1939 (
** Riccardia singapurensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898 (
** Riccardia smaragdina Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 20, 1985 (
*** Riccardia sprucei (Steph.) Meenks et C.De Jong, Cryptog. Bryol. Lichénol. 6 (1): 22, 1985 (
** Riccardia squamifera Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 19, 1964 (
* Riccardia stipatiflora (Steph.) Pagán, Bryologist 45 (4): 81, 1942 (
** Riccardia stricta R.M.Schust., J. Hattori Bot. Lab. 62: 326, 1987 (
** Riccardia subantarctica Grolle et L.Söderstr., Phytotaxa 202 (1): 70, 2015 (
** Riccardia subexalata Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 163, 1898 (
* Riccardia subexalata var. procera Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 164, 1898 (
* Riccardia submersa (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (
** Riccardia submultifida Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 128, 1934 (
** Riccardia subpalmata (Steph.) Hürl., Bauhinia 5 (4): 194, 1976 (
** Riccardia subsimplex (Steph.) Pagán, Bryologist 45 (4): 81, 1942 (
** Riccardia sumatrana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 173, 1898 (
** Riccardia tahitensis (Steph.) Hürl., Bauhinia 5 (4): 201, 1976 (
* Riccardia tenella Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 95, 1970 (
*** Riccardia tenuicula (Spruce) Meenks, Beih. Nova Hedwigia 88: 101, 1987 (
** Riccardia tenuis (Steph.) Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 172, 1898 (
* Riccardia tjibodensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 165, 1898 (
*** Riccardia trichomanoides (Spruce) Hässel ex Meenks, Beih. Nova Hedwigia 88: 101, 1987 (
** Riccardia trukensis H.A.Mill. et Bonner, Beih. Nova Hedwigia 11: 70, 1963 (
** Riccardia tumbareriensis Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 87, 1970 (
** Riccardia upoluna (Steph.) Grolle, J. Hattori Bot. Lab. 36: 77, 1972 [1973] (
** Riccardia valida (Steph.) J.J.Engel, Bryologist 78 (3): 362, 1975 (
** Riccardia venosa (Steph.) Hürl., Bauhinia 5 (4): 208, 1976 (
* Riccardia villosa (Steph.) Pandé et S.C.Srivast., Biol. Mem. 1 (1/2): 129, 1976 (
** Riccardia virens (Steph.) Hürl., Bauhinia 5 (4): 209, 1976 (
** Riccardia virgata (Gottsche) Pagán, Bryologist 42 (1): 7, 1939 (
** Riccardia wallisii (Steph.) Gradst., J. Hattori Bot. Lab. 45: 130, 1979 (
** Riccardia wettsteinii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 162, 1898 (
* Riccardia wettsteinii var. angustilimbia Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 163, 1898 (
* Riccardia wettsteinii var. crassa Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 163, 1898 (
* Riccardia wettsteinii var. procera Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 162, 1898 (
* Riccardia wettsteinii var. tenuiretis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 162, 1898 (
** Riccardia womersleyana Hewson, Proc. Linn. Soc. New South Wales (ser. 2) 95 (1): 98, 1970 (
** Verdoornia R.M.Schust., J. Hattori Bot. Lab. 26: 291, 1963 (
*** Verdoornia succulenta R.M.Schust., J. Hattori Bot. Lab. 26: 291, 1963 (
by D. P. Costa
The treatment of Metzgeriaceae follows what was outlined in
*** Metzgeria Raddi, Jungermanniogr. Etrusca: 34, 1818 (
*** Metzgeria acuminata Steph., Bull. Herb. Boissier 7 (12): 934 (282), 1899 (
*** Metzgeria adscendens Steph. ex K.I.Goebel, Flora 77: 427, 1893 (
*** Metzgeria agnewiae Kuwah., Bryologist 76 (4): 569, 1973 (
*** Metzgeria albinea Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cci, 1889 [1890] (
*** Metzgeria albinea var. aberrans Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 27, 1964 (
*** Metzgeria albinea var. angusta (Steph.) D.P.Costa et Gradst., Bryologist 103 (4): 757, 2000 [2001] (
*** Metzgeria allionii Steph., Sp. Hepat. (Stephani) 6: 47, 1917 (
*** Metzgeria alpina R.M.Schust. et J.J.Engel, Brittonia 40 (2): 203, 1988 (
*** Metzgeria americana Masuzaki, Hikobia 15 (4): 441, 2010 (
*** Metzgeria attenuata Steph., Biblioth. Bot. 87 (2): 177, 1916 (
*** Metzgeria aurantiaca Steph., Bull. Herb. Boissier 7 (12): 938 (286), 1899 (
*** Metzgeria auriculata Grolle et Kuwah., Bryophyt. Biblioth. 28: 194, 1986 (
*** Metzgeria bahiensis Schiffn., Österr. Bot. Z. 61 (7/8): 262, 1911 (
*** Metzgeria bartlettii Kuwah., Mem. New York Bot. Gard. 45: 561, 1987 (
*** Metzgeria bischlerae Kuwah., J. Hattori Bot. Lab. 40: 264, 1976 (
*** Metzgeria bracteata Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 553, 1885 (
*** Metzgeria brasiliensis Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 22, 1964 (
*** Metzgeria chilensis Steph., Bull. Herb. Boissier 7 (12): 937 (285), 1899 (
*** Metzgeria ciliata Raddi, Critt. Brasil.: 17, 1822 (
*** Metzgeria claviflora Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 556, 1885 (
*** Metzgeria cleefii Kuwah., Proc. Kon. Ned. Akad. Wetensch. C 85 (3): 360, 1982 (
** Metzgeria comata Steph., Bull. Herb. Boissier 7 (12): 939 (287), 1899 (
*** Metzgeria conjugata Lindb., Acta Soc. Sci. Fenn. 10: 495, 1875 (
*** Metzgeria consanguinea Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 271, 1893 (
*** Metzgeria convoluta Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899 (
** Metzgeria coorgensis S.C.Srivast. et S.Srivast., Phytotaxonomy 4: 81, 2004 (
** Metzgeria corralensis Steph., Bull. Herb. Boissier 7 (12): 933 (281), 1899 (
*** Metzgeria crassipilis (Lindb.) A.Evans, Rhodora 11 (130): 188, 1909 (
*** Metzgeria cratoneura Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 24, 1964 (
*** Metzgeria cylindra Kuwah., Bryologist 81 (3): 405, 1978 (
** Metzgeria decrescens Steph., Bull. Herb. Boissier 7 (12): 932 (280), 1899 (
*** Metzgeria dichotoma (Sw.) Nees, Syn. Hepat. 4: 504, 1846 (
** Metzgeria divaricata A.Evans, Proc. Amer. Acad. Arts 58 (7): 288, 1923 (
*** Metzgeria dorsipara (Herzog) Kuwah., J. Hattori Bot. Lab. 40: 269, 1976 (
*** Metzgeria duricosta Steph., Sp. Hepat. (Stephani) 6: 50, 1917 (
** Metzgeria engelii Kuwah., Hikobia 8 (3/4): 275, 1980 (
** Metzgeria epiphylla A.Evans, Proc. Amer. Acad. Arts 58 (7): 303, 1923 (
*** Metzgeria filicina Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 361, 1851 (
** Metzgeria flavovirens Colenso, Trans. & Proc. New Zealand Inst. 21: 79, 1889 (
* Metzgeria foliicola Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 181, 1898 (
*** Metzgeria francana Steph., Sp. Hepat. (Stephani) 6: 51, 1917 (
** Metzgeria frontipilis Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 14, 1877 (
*** Metzgeria fruticola Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 554, 1885 (
*** Metzgeria furcata (L.) Corda, Gen. hepat.: 654, 1829 (
* Metzgeria furcata var. expansa Douin, Rev. Bryol. 30 (3): 47, 1903 (
* Metzgeria furcata var. pacifica Brinkm., Bryologist 34 (2): 15, 1931 (
*** Metzgeria grandiflora A.Evans, Torreya 16 (3): 68, 1916 (
** Metzgeria hasselii Kuwah., J. Hattori Bot. Lab. 40: 509, 1976 (
** Metzgeria hebridensis Steph., Sp. Hepat. (Stephani) 6: 51, 1917 (
*** Metzgeria hegewaldii Kuwah., Nova Hedwigia 34: 784, 1981 (
*** Metzgeria herminieri Schiffn., Österr. Bot. Z. 61 (7/8): 261, 1911 (
* Metzgeria heteroramea Steph., Biblioth. Bot. 87 (2): 178, 1916 (
** Metzgeria imberbis J.B.Jack et Steph., Hedwigia 34 (6): 316, 1895 (
*** Metzgeria inflata Steph., Bull. Herb. Boissier 7 (12): 936 (284), 1899 (
*** Metzgeria jamesonii Kuwah., Bryophyt. Biblioth. 28: 157, 1986 (
** Metzgeria kanaii Kuwah., Fl. E. Himalaya 2: 240, 1971 (
*** Metzgeria kinabaluensis Masuzaki, Hikobia 16 (1): 59, 2011 (
** Metzgeria kuwaharae Piippo, Acta Bot. Fenn. 143: 8, 1991 (
** Metzgeria laciniata Kuwah., Bryologist 81 (3): 406, 1978 (
*** Metzgeria lechleri Steph., Bull. Herb. Boissier 7 (12): 942 (290), 1899 (
*** Metzgeria leptoneura Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 555, 1885 (
** Metzgeria leptoneura var. breviseta (Schiffn.) O.Yano, J. Hattori Bot. Lab. 56: 532, 1984 (
** Metzgeria leptoneura var. polychaeta R.M.Schust., Phytotaxa 202 (1): 70, 2015 (
*** Metzgeria liebmaniana Lindenb. et Gottsche, Syn. Hepat. 4: 505, 1846 (
*** Metzgeria lindbergii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 182, 1898 (
** Metzgeria litoralis J.J.Engel et Kuwah., Bryologist 76 (2): 293, 1973 (
*** Metzgeria longitexta Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899 (
** Metzgeria macrospora Kuwah., J. Hattori Bot. Lab. 32: 17, 1969 (
** Metzgeria macveanii Kuwah., Rev. Bryol. Lichénol. 36 (3/4): 539, 1969 [1970] (
*** Metzgeria maegdefraui Kuwah., Hikobia 8 (3/4): 269, 1980 (
** Metzgeria magellanica Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 43, 1890 (
*** Metzgeria metaensis Kuwah., Proc. Kon. Ned. Akad. Wetensch. C 85 (3): 375, 1982 (
*** Metzgeria mexicana Steph., Sp. Hepat. (Stephani) 6: 55, 1917 (
** Metzgeria monoica Kuwah. et J.J.Engel, Hikobia 8 (3/4): 284, 1980 (
*** Metzgeria myriopoda Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 22, 1877 (
*** Metzgeria neotropica Kuwah., Nova Hedwigia 34: 779, 1981 (
*** Metzgeria nudifrons Steph., Hedwigia 31 (3): 126, 1892 (
*** Metzgeria parviinvolucrata Kuwah., Nova Hedwigia 34: 774, 1981 (
** Metzgeria patagonica Steph., Bull. Herb. Boissier 7 (12): 940 (288), 1899 (
*** Metzgeria polytricha Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 553, 1885 (
*** Metzgeria procera Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 166, 1855 (
*** Metzgeria psilocraspeda Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 25, 1964 (
*** Metzgeria pubescens (Schrank) Raddi, Jungermanniogr. Etrusca: 35, 1818 (
*** Metzgeria pulvinata Steph., Biblioth. Bot. 87 (2): 179, 1916 (
** Metzgeria quadrifaria Steph., Bull. Herb. Boissier 7 (12): 953 (301), 1899 (
** Metzgeria raoi S.C.Srivast. et S.Srivast., Phytotaxonomy 4: 83, 2004 (
** Metzgeria rigida Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 43, 1877 (
** Metzgeria robinsonii Steph., Sp. Hepat. (Stephani) 6: 60, 1917 (
** Metzgeria roivainenii Kuwah., J. Hattori Bot. Lab. 40: 516, 1976 (
*** Metzgeria rufula Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 555, 1885 (
** Metzgeria saccata Mitt., J. Linn. Soc., Bot. 22 (145): 241, 1886 (
** Metzgeria saxbyi Pearson, Ann. Cryptog. Exot. 4 (2): 70, 1931 (
** Metzgeria scobina Mitt., J. Linn. Soc., Bot. 22 (145): 243, 1886 (
*** Metzgeria scyphigera A.Evans, Trans. Connecticut Acad. Arts 18 (5): 299, 1914 (
*** Metzgeria senjoana Masuzaki, Hikobia 15 (4): 445, 2010 (
** Metzgeria setigera R.M.Schust. ex Crand.-Stotl. et L.Söderstr., Phytotaxa 202 (1): 69, 2015 (
** Metzgeria sikkimensis S.C.Srivast. et K.K.Rawat, Geophytology 31 (1/2): 71, 2001 [2003] (
* Metzgeria simplex Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 115, 1941 (
*** Metzgeria sinuata Loitl., Diagn. pl. nov.: 25, 1894 (
** Metzgeria sparrei Kuwah., Hikobia 8 (3/4): 278, 1980 (
*** Metzgeria spindleri Steph., Biblioth. Bot. 87 (2): 179, 1916 (
*** Metzgeria subaneura Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 22, 1964 (
** Metzgeria submarginata M.L.So, New Zealand J. Bot. 40 (2): 201, 2002 (
** Metzgeria subundulata (Lindb.) Kuwah., Bryologist 86 (3): 276, 1983 [1984] (
** Metzgeria temperata Kuwah., J. Hattori Bot. Lab. 40: 219, 1976 (
*** Metzgeria uncigera A.Evans, Ann. Bot. (Oxford) 24 (2): 276, 1910 (
*** Metzgeria undulata Kuwah., Nova Hedwigia 34: 792, 1981 (
*** Metzgeria violacea (Ach.) Dumort., Recueil Observ. Jungerm.: 26, 1835 (
** Steereella Kuwah., Amer. J. Bot. 60 (6): 602, 1973 (
*** Steereella lilliana (Steph.) Kuwah., Bryophyt. Biblioth. 28: 179, 1986 (
*** Steereella linearis (Sw.) Kuwah., Amer. J. Bot. 60 (6): 604, 1973 (
** Vandiemenia Hewson, J. Hattori Bot. Lab. 52: 163, 1982 (
** Vandiemenia ratkowskiana Hewson, J. Hattori Bot. Lab. 52: 163, 1982 (
by B. Thiers
*** Pleurozia Dumort., Recueil Observ. Jungerm.: 15, 1835 (
** subg. Constantifolia B.M.Thiers, Bryologist 96 (4): 526, 1993 (
*** Pleurozia conchifolia (Hook. et Arn.) Austin, Bull. Torrey Bot. Club 5 (3): 16, 1874 (
** Pleurozia conchifolia var. papillosa B.M.Thiers, Bryologist 96 (4): 528, 1993 (
*** Pleurozia purpurea Lindb., Acta Soc. Fauna Fl. Fenn. 1 (2): 27, 1877 (
** subg. Diversifolia B.M.Thiers, Bryologist 96 (4): 531, 1993 (
*** Pleurozia acinosa (Mitt.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 412, 1877 (
*** Pleurozia articulata (Lindb.) Lindb. et Lackström, Hepat. Scand. Exsicc.: 5 (adnot.), 1874 (
*** Pleurozia caledonica (Gottsche) Steph., Rev. Bryol. 33 (2): 29 (
*** Pleurozia curiosa B.M.Thiers, Bryologist 96 (4): 537, 1993 (
*** Pleurozia heterophylla Steph. ex Fulford, Mem. New York Bot. Gard. 23: 842, 1972 (
*** Pleurozia johannis-winkleri Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 195, 1931 (
*** Pleurozia paradoxa (J.B.Jack) Schiffn., Hepat. (Engl.-Prantl): 115, 1893 (
*** Pleurozia subinflata (Austin) Austin, Bull. Torrey Bot. Club 5 (3): 16, 1874 (
** subg. Pleurozia
*** Pleurozia gigantea (F.Weber) Lindb., Hepat. Scand. Exsicc.: no. 5, 1874 (
Incertae sedis
** Pleurozia pocsii Frank Müll., Polish Bot. J. 58 (1): 50, 2013 (
*** Calycularia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861] (
*** Calycularia crispula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 122, 1860 [1861] (
*** Calycularia laxa Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 66, 1889 (
*** Allisonia Herzog, Hedwigia 80 (1/2): 77, 1941 (
*** Allisonia cockaynei (K.I.Goebel) R.M.Schust., J. Hattori Bot. Lab. 26: 294, 1963 (
by R. Stotler, B. J. Crandall-Stotler and D. C. Cargill
*** Fossombronia Raddi, Jungermanniogr. Etrusca: 29, 1818 (
*** Fossombronia alaskana Steere et Inoue, Bryologist 77 (1): 66, 1974 (
*** Fossombronia alata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 340, 1984 (
*** Fossombronia altilamellosa G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (
*** Fossombronia angulifolia Perold, Bothalia 28 (2): 159, 1998 (
*** Fossombronia angulosa (Dicks.) Raddi, Jungermanniogr. Etrusca: 29, 1818 (
** Fossombronia areolata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 368, 1987 (
** Fossombronia auricolor G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (
*** Fossombronia australis Mitt., J. Linn. Soc., Bot. 15 (82): 73, 1876 (
*** Fossombronia caespitiformis (Raddi) De Not. ex Rabenh., Hepat. Eur., Leberm. 13-14: no. 123, 1860 [1861] (
*** Fossombronia caespitiformis subsp. multispira (Schiffn.) J.R.Bray et Cargill, Bryologist 106 (1): 131, 2003 (
*** Fossombronia caledonica Steph., Sp. Hepat. (Stephani) 6: 71, 1917 (
*** Fossombronia cederbergensis Perold, Bothalia 28 (1): 1, 1998 (
*** Fossombronia cerebriformis G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 343, 1984 (
*** Fossombronia crassifolia Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 527, 1885 (
*** Fossombronia crispa Nees, Syn. Hepat. 4: 469, 1846 (
* Fossombronia crispula (Brot.) R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 383, 1992 (
*** Fossombronia cristula Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 228, 1869 (
*** Fossombronia cultriformis G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 371, 1987 (
*** Fossombronia densa G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 372, 1987 (
*** Fossombronia densilamellata S.W.Arnell, Bot. Not. 105: 317, 1952 (
*** Fossombronia echinata Macvicar, Rev. Bryol. 38 (4): 73, 1911 (
*** Fossombronia elsieae Perold, Bothalia 29 (1): 25, 1999 (
*** Fossombronia fernandeziensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 15, 1911 (
*** Fossombronia fimbriata Paton, J. Bryol. 8 (1): 1, 1974 (
*** Fossombronia fleischeri Osterwald ex Loeske, Verh. Bot. Vereins Prov. Brandenburg 70: 125, 1928 (
*** Fossombronia foveolata Lindb., Helsingf. Dagbl. 1873 (353, 28 Dec): 2, 1873 (
** Fossombronia fuhreri G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 374, 1987 (
*** Fossombronia gemmifera Perold, Bothalia 27 (1): 19, 1997 (
*** Fossombronia glenii Perold, Bothalia 27 (1): 20, 1997 (
** Fossombronia grandis Steph., Mém. Herb. Boissier 16: 29 (383), 1900 (
* Fossombronia gregaria Colenso, Trans. & Proc. New Zealand Inst. 20: 252, 1888 (
* Fossombronia grossepapillata Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 105, 1914 (
*** Fossombronia hamatohirta Steph., Hedwigia 33 (1): 8, 1894 (
** Fossombronia hewsoniae G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 375, 1987 (
*** Fossombronia himalayensis Kashyap, New Phytol. 14 (1): 4, 1915 (
*** Fossombronia hyalorhiza Perold, Bothalia 29 (1): 83, 1999 (
*** Fossombronia incurva Lindb., Helsingf. Dagbl. 1873 (273, 7 Oct): 2, 1873 (
*** Fossombronia indica Steph., Sp. Hepat. (Stephani) 6: 73, 1917 (
* Fossombronia integerrima Steph., Mém. Herb. Boissier 16: 40 (394), 1900 (
* Fossombronia integrifolia Steph., Sp. Hepat. (Stephani) 6: 73, 1917 (
*** Fossombronia intestinalis Taylor, London J. Bot. 5: 408, 1846 (
*** Fossombronia japonica Schiffn., Österr. Bot. Z. 49 (11): 389, 1899 (
** Fossombronia laciniata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 375, 1987 (
*** Fossombronia lamellata Steph., Hedwigia 33 (1): 9, 1894 (
*** Fossombronia leucoxantha (Lehm.) Lehm. et Lindenb., Syn. Hepat. 4: 469, 1846 (
*** Fossombronia longiseta (Austin) Austin, Hepat. bor.-amer.: 30, 1873 (
*** Fossombronia lophoclada Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 529, 1885 (
** Fossombronia lophoscypha Hässel, Beih. Nova Hedwigia 131: 16, 2007 (
*** Fossombronia luetzelburgiana K.I.Goebel, Flora 105: 55, 1912 (
*** Fossombronia macrocalyx Steph., Sp. Hepat. (Stephani) 6: 74, 1917 (
* Fossombronia macrophylla Colenso, Trans. & Proc. New Zealand Inst. 18: 285, 1886 (
** Fossombronia magnaspora G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (
*** Fossombronia marindae Perold, Bothalia 29 (1): 86, 1999 (
** Fossombronia maritima (Paton) Paton, J. Bryol. 18 (2): 367, 1994 (
*** Fossombronia marshii J.R.Bray et Stotler, Phytologia 92 (2): 230, 2010 (
** Fossombronia microlamellata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 347, 1984 (
*** Fossombronia mittenii Tind., J. Bot. 36: 44, 1898 (
*** Fossombronia montaguensis S.W.Arnell, Bot. Not. 105: 316, 1952 (
*** Fossombronia monticola Perold, Bothalia 29 (1): 87, 1999 (
*** Fossombronia mylioides Inoue, J. Hattori Bot. Lab. 37: 296, 1973 (
* Fossombronia nigricaulis Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886 (
** Fossombronia nyikaensis Perold, Bothalia 31 (1): 48, 2001 (
*** Fossombronia papillata Steph., Hedwigia 28 (3): 157, 1889 (
*** Fossombronia paranapanemae Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 34, 1964 (
*** Fossombronia peruviana Gottsche et Hampe, Linnaea 27 (5): 555, 1854 (
*** Fossombronia porphyrorhiza (Nees) Prosk., Bryologist 58 (3): 197, 1955 (
** Fossombronia pulvinata Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 113, 1911 (
*** Fossombronia punctata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 341, 1984 (
*** Fossombronia purpureospora G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (
*** Fossombronia pusilla (L.) Nees, Naturgesch. Eur. Leberm. 3: 319, 1838 (
*** Fossombronia renateae Perold, Bothalia 29 (1): 77, 1999 (
*** Fossombronia reticulata Steph., Hedwigia 33 (1): 9, 1894 (
* Fossombronia rosulata Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886 (
** Fossombronia rudis G.A.M.Scott et D.C.Pike, Beih. Nova Hedwigia 90: 110, 1988 (
*** Fossombronia ruminata Cargill, Phytotaxa 65: 45, 2012 (
** Fossombronia rupestris G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (
*** Fossombronia rwandaensis Perold, Bothalia 28 (1): 45, 1998 (
*** Fossombronia scrobiculata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 341, 1984 (
*** Fossombronia spinifolia Steph., Mém. Herb. Boissier 16: 35 (389), 1900 (
* Fossombronia spinosa Perold, Bothalia 29 (1): 29, 1999 (
*** Fossombronia stephanii Schiffn. ex Steph., Mém. Herb. Boissier 16: 27 (381), 1900 (
*** Fossombronia straussiana Perold, Bothalia 27 (1): 24, 1997 (
* Fossombronia subsaccata Steph., Sp. Hepat. (Stephani) 6: 75, 1917 (
*** Fossombronia swaziensis Perold, Bothalia 28 (2): 162, 1998 (
** Fossombronia tesselata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 62: 382, 1987 (
*** Fossombronia texana Lindb., Acta Soc. Sci. Fenn. 10: 533, 1875 (
*** Fossombronia truncata G.A.M.Scott et D.C.Pike, Fl. Austral. Suppl. 21: 114, 2003 (
*** Fossombronia tumida Mitt., J. Linn. Soc., Bot. 16 (91): 193, 1877 (
** Fossombronia valparaisiana Hässel, Beih. Nova Hedwigia 131: 14, 2007 (
** Fossombronia vermiculata G.A.M.Scott et D.C.Pike, J. Hattori Bot. Lab. 56: 345, 1984 (
*** Fossombronia wattsii Steph., Sp. Hepat. (Stephani) 6: 75, 1917 (
*** Fossombronia wondraczekii (Corda) Dumort. ex Lindb., Helsingf. Dagbl. 1873 (273, 7 Oct): 2, 1873 (
*** Fossombronia wrightii Austin, Bot. Bull. (Hanover) 1 (8): 36, 1876 (
** Fossombronia zuurbergensis Perold, Bothalia 31 (1): 25, 2001 (
by R. Stotler and B.J. Crandall-Stotler
*** Petalophyllum Nees et Gottsche, Nov. Stirp. Pug. 8: 29, 1844 (
*** Petalophyllum americanum C.H.Ford et Crand.-Stotl., Novon 12 (3): 335, 2002 (
*** Petalophyllum hodgsoniae C.H.Ford et Crand.-Stotl., Novon 12 (3): 336, 2002 (
*** Petalophyllum indicum Kashyap, J. Indian Bot. Soc. 7: 14, 1928 (
*** Petalophyllum preissii Gottsche, Nov. Stirp. Pug. 8: 30, 1844 (
*** Petalophyllum ralfsii (Wilson) Nees et Gottsche, Nov. Stirp. Pug. 8: 30, 1844 (
*** Sewardiella Kashyap, New Phytol. 14 (1): 5, 1915 (
*** Sewardiella tuberifera Kashyap, New Phytol. 14 (1): 5, 1915 (
Makinoiineae He-Nygrén, Juslén, Ahonen, Glenny et Piippo
by R. Stotler and B.J. Crandall-Stotler
*** Makinoa Miyake, Bot. Mag. (Tokyo) 13 (144): 23, 1899 (
*** Makinoa crispata (Steph.) Miyake, Bot. Mag. (Tokyo) 13 (144): 23, 1899 (
by R. Stotler and B.J. Crandall-Stotler
*** Hymenophyton Dumort., Recueil Observ. Jungerm.: 25, 1835 (
*** Hymenophyton flabellatum (Labill.) Dumort. ex Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 430, 1877 (
*** Hymenophyton leptopodum (Hook.f. et Taylor) A.Evans, Trans. Connecticut Acad. Arts 8 (16): 274, 1893 (
*** Hymenophyton pedicellatum Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 11, 1911 (
by B.J. Crandall-Stotler and R. Stotler
*** Hattorianthus R.M.Schust. et Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 103, 1975 (
*** Hattorianthus erimonus (Steph.) R.M.Schust. et Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (3): 106, 1975 (
*** Moerckia Gottsche, Hepat. Eur., Leberm. 13-14: no. 121, 1860 [1861] (
*** Moerckia blyttii (Mørch) Brockm., Arch. Vereins Freunde Naturgesch. Mecklenburg 17: 190, 1863 (
*** Moerckia flotoviana (Nees) Schiffn., Österr. Bot. Z. 51 (2): 43, 1901 (
*** Moerckia hibernica (Hook.) Gottsche, Hepat. Eur., Leberm. 13-14: no. 121, 1860 [1861] (
by R. Stotler and B.J. Crandall-Stotler
*** Jensenia Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 13, 1868 (
*** Jensenia connivens (Colenso) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965] (
* Jensenia crassifrons (Steph.) S.Schuette et Stotler, J. Hattori Bot. Lab. 97: 300, 2005 (
*** Jensenia decipiens (Mitt.) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965] (
*** Jensenia difformis (Nees) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965] (
*** Jensenia florschuetzii Gronde, Proc. Kon. Ned. Akad. Wetensch. C 83 (3): 273, 1980 (
*** Jensenia spinosa (Lindenb. et Gottsche) Grolle, Acta Bot. Fenn. 133: 65, 1986 (
*** Jensenia wallisii (J.B.Jack et Steph.) Grolle, Rev. Bryol. Lichénol. 33 (1/2): 228, 1964 [1965] (
*** Pallavicinia Gray, Nat. Arr. Brit. Pl. 1: 775, 1821 (
*** Pallavicinia ambigua (Mitt.) Steph., Mém. Herb. Boissier 11: 8 (312), 1900 (
** Pallavicinia baldwinii (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891 (
* Pallavicinia bipinnata Steph., Sp. Hepat. (Stephani) 6: 62, 1917 (
** Pallavicinia camisassai Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 4, 1916 (
** Pallavicinia cylindrica (Austin) A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891 (
** Pallavicinia himalayensis Schiffn., Mém. Herb. Boissier 11: 13 (317), 1900 (
*** Pallavicinia indica Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 183, 1898 (
*** Pallavicinia levieri Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 184, 1898 (
*** Pallavicinia lyellii (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 775, 1821 (
** Pallavicinia pilifera Steph., Hedwigia 30 (6): 271, 1891 (
** Pallavicinia purpurea Steph., Sp. Hepat. (Stephani) 6: 64, 1917 (
** Pallavicinia ridleyi Steph., Sp. Hepat. (Stephani) 6: 64, 1917 (
*** Pallavicinia rubristipa Schiffn., Österr. Bot. Z. 56 (1): 24, 1906 (
*** Pallavicinia subciliata (Austin) Steph., Mém. Herb. Boissier 11: 9 (313), 1900 (
*** Pallavicinia xiphoides (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 427, 1877 (
*** Podomitrium Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 164, 1855 (
*** Podomitrium malaccense (Steph.) Campb., Amer. J. Bot. 2 (5): 199, 1915 (
*** Podomitrium marginatum (Steph.) Hürl., Bauhinia 4 (1): 78, 1968 [1969] (
*** Podomitrium phyllanthus (Hook.) Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 164, 1855 (
** Greeneothallus Hässel, J. Bryol. 11 (1): 115, 1980 (
*** Greeneothallus gemmiparus Hässel, J. Bryol. 11 (1): 115, 1980 (
*** Seppeltia Grolle, J. Hattori Bot. Lab. 60: 276, 1986 (
*** Seppeltia succuba Grolle, J. Hattori Bot. Lab. 60: 276, 1986 (
*** Symphyogyna Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 66, 1836 (
*** Symphyogyna apiculispina Steph., Biblioth. Bot. 87 (2): 180, 1916 (
*** Symphyogyna aspera Steph. ex F.A.McCormick, Bot. Gaz. 58 (5): 403, 1914 (
** Symphyogyna atronervia Taylor, London J. Bot. 5: 409, 1846 (
** Symphyogyna bogotensis Steph., Mém. Herb. Boissier 11: 42 (346), 1900 (
* Symphyogyna boliviensis Steph., Biblioth. Bot. 87 (2): 180, 1916 (
*** Symphyogyna brasiliensis Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 67, 1836 (
** Symphyogyna brasiliensis var. angustior (Gottsche, Lindenb. et Nees) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 183, 1864 (
** Symphyogyna brasiliensis var. subsinuata Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 32, 1964 (
*** Symphyogyna brongniartii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 265, 1843 (
*** Symphyogyna circinata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 69, 1836 (
*** Symphyogyna digitisquama Steph., Mém. Herb. Boissier 11: 31 (335), 1900 (
** Symphyogyna fuscovirens A.Evans, Trans. Connecticut Acad. Arts 28 (6): 320, 1927 (
*** Symphyogyna hochstetteri Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 68, 1836 (
*** Symphyogyna hymenophyllum (Hook.) Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 68, 1836 (
** Symphyogyna hymenophyllum var. heterogenum Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 536, 1885 (
** Symphyogyna ignambiensis Hürl., Bauhinia 4 (1): 79, 1968 [1969] (
** Symphyogyna interrupta Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1053, 1888 (
** Symphyogyna irregularis Steph., Mém. Herb. Boissier 11: 29 (333), 1900 (
** Symphyogyna lacerosquama Steph., Sp. Hepat. (Stephani) 6: 67, 1917 (
*** Symphyogyna leptothelia Taylor, London J. Bot. 5: 408, 1846 (
** Symphyogyna lindmanii A.Evans, Trans. Connecticut Acad. Arts 28 (6): 316, 1927 (
*** Symphyogyna luetzelburgii Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 22, 1925 (
*** Symphyogyna marginata Steph., Mém. Herb. Boissier 11: 30 (334), 1900 (
*** Symphyogyna mexicana Steph., Rev. Bryol. 36 (6): 140, 1909 (
** Symphyogyna multiflora Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 133, 1914 (
* Symphyogyna paucidens Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 13, 1911 (
*** Symphyogyna podophylla (Thunb.) Nees et Mont., Flora 29 (9): 135, 1846 (
* Symphyogyna purpureolimbata E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 95, 1965 (
*** Symphyogyna rectidens Grolle, Acta Bot. Fenn. 133: 68, 1986 (
** Symphyogyna rhodina (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 487, 1846 (
*** Symphyogyna rubescens Steph., Mém. Herb. Boissier 11: 29 (333), 1900 (
*** Symphyogyna rubritincta A.Evans, Trans. Connecticut Acad. Arts 27 (1): 38, 1925 (
** Symphyogyna semi-involucrata Austin, Bull. Torrey Bot. Club 5 (3): 15, 1874 (
** Symphyogyna similis Grolle, Acta Bot. Fenn. 133: 70, 1986 (
*** Symphyogyna sinuata (Sw.) Nees et Mont., Voy. Amér. Mérid., Bot. 7 (1): 61, 1839 (
*** Symphyogyna subsimplex Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 166, 1855 (
*** Symphyogyna tenuinervis (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 61: 253, 1986 [1987] (
*** Symphyogyna trivittata Spruce, J. Linn. Soc., Bot. 30 (210): 365, 1895 (
* Symphyogyna ulvoides (Reinw., Blume et Nees) Nees, Syn. Hepat. 4: 487, 1846 (
*** Symphyogyna undulata Colenso, Trans. & Proc. New Zealand Inst. 16: 356, 1884 (
** Symphyogyna volkensii Steph., Mém. Herb. Boissier 11: 35 (339), 1900 (
** Symphyogynopsis Grolle, Acta Bot. Fenn. 133: 72, 1986 (
*** Symphyogynopsis gottscheana (Mont. et Nees) Grolle, J. Hattori Bot. Lab. 63: 441, 1987 (
** Xenothallus R.M.Schust., J. Hattori Bot. Lab. 26: 293, 1963 (
*** Xenothallus vulcanicola R.M.Schust., J. Hattori Bot. Lab. 26: 293, 1963 (
** Sandeothallus R.M.Schust., Nova Hedwigia 36: 10, 1982 (
** Sandeothallus japonicus (Inoue) Crand.-Stotl. et Stotler, Beih. Nova Hedwigia 131: 58, 2007 (
** Sandeothallus radiculosus (Schiffn.) R.M.Schust., Nova Hedwigia 36: 11, 1982 (
*** Phyllothallia E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (19): 247, 1964 (
*** Phyllothallia fuegiana R.M.Schust., Trans. Brit. Bryol. Soc. 5 (2): 284, 1967 (
*** Phyllothallia nivicola E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 2 (19): 247, 1964 (
*** Noteroclada Taylor ex Hook.f. et Wilson, London J. Bot. 3: 166, 1844 (
*** Noteroclada confluens Taylor, London J. Bot. 3: 166, 1844 (
Excluded from the genus
* Noteroclada longiuscula Colenso, Trans. & Proc. New Zealand Inst. 19: 299, 1887 (
** Pellia Raddi, Jungermanniogr. Etrusca: 38, 1818 (
*** subg. Apopellia Grolle, J. Bryol. 12 (3): 427, 1983 (
** Pellia alpicola R.M.Schust. ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 76 (3): 39, 2013 (
*** Pellia endiviifolia (Dicks.) Dumort., Recueil Observ. Jungerm.: 27, 1835 (
** Pellia megaspora R.M.Schust., J. Bryol. 11 (3): 419, 1981 (
*** subg. Pellia
** Pellia appalachiana R.M.Schust., Phytotaxa 76 (3): 39, 2013 (
*** Pellia epiphylla (L.) Corda, Gen. hepat.: 654, 1829 (
** Pellia epiphylla subsp. borealis (Lorb.) Messe, Bull. Soc. Roy. Bot. Belgique 114 (1): 13, 1981 (
*** Pellia neesiana (Gottsche) Limpr., Hedwigia 15 (2): 18, 1876 (
Incertae sedis
* Pellia cordaeana Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 433, 1877 (
* Pellia crispa P.Kumm., Führer Leberm.: 60, 1875 (
* Pellia gottscheana Kreh, Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 90 (4): 237, 1909 (
* Pellia longifolia P.Kumm., Führer Leberm.: 60, 1875 (
* Pellia undulata P.Kumm., Führer Leberm.: 60, 1875 (
by L. Söderström
*** Blasia L., Sp. Pl. 1: 1138, 1753 (
*** Blasia pusilla L., Sp. Pl. 1: 1138, 1753 (
*** Cavicularia Steph., Bull. Herb. Boissier 5 (2): 87, 1897 (
*** Cavicularia densa Steph., Bull. Herb. Boissier 5 (2): 87, 1897 (
by D.G. Long
*** Lunularia Adans., Fam. Pl. (Adanson) 2: 15, 1763 (
*** Lunularia cruciata (L.) Dumort. ex Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 298, 1868 (
* Lunularia cruciata subsp. thaxteri (A.Evans et Herzog) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 91, 1992 (
by D.G. Long
The genera of Aytoniaceae follow the treatment of
*** Asterella P.Beauv., Dict. Sci. Nat. [F. Cuvier] 3: 257, 1805 (
** subg. Asterella
** sect. Asterella
*** Asterella tenella (L.) P.Beauv., Dict. Sci. Nat. [F. Cuvier] 3: 258, 1805 (
** sect. Brachyblepharis (Nees) D.G.Long, J. Bryol. 22 (2): 113, 2000 (
*** Asterella abyssinica (Gottsche) Grolle, Explor. Hydrobiol. Lac Bangweolo Luapula: 170, 1972 (
*** Asterella africana (Mont.) Underw. ex A.Evans, Contr. U.S. Natl. Herb. 20: 250, 1920 (
*** Asterella blumeana (Nees) Kachroo, J. Gauhati India Univ. 3: 130, 1952 (
*** Asterella chilensis (Nees et Mont.) A.Evans, Bull. Torrey Bot. Club 46 (12): 469, 1919 (
*** Asterella cruciata (Steph.) Horik., Hikobia 1 (2): 79, 1951 (
*** Asterella dissoluta (Steph.) Grolle, Wiss. Z. Friedrich-Schiller-Univ. Jena, Math.-Naturwiss. Reihe 38 (2): 237, 1989 (
*** Asterella dominicensis S.W.Arnell, Bryologist 61 (2): 140, 1958 (
*** Asterella khasyana (Griff.) Grolle, Khumbu Himal 1 (4): 267, 1966 (
*** Asterella leptophylla (Mont.) Grolle, Feddes Repert. 87 (3/4): 246, 1976 (
*** Asterella limbata D.G.Long et Grolle, J. Bryol. 18 (2): 287, 1994 (
** Asterella shimizuana Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 10 (3): 361, 1967 (
*** Asterella tenera (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (
* Asterella tenerrima (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
*** Asterella venosa (Lehm. et Lindenb.) A.Evans, Contr. U.S. Natl. Herb. 20: 286, 1920 (
** subg. Phragmoblepharis Grolle, Feddes Repert. 87 (3/4): 246, 1976 (
*** Asterella australis (Hook.f. et Taylor) Verd., Ann. Bryol. 5: 126, 1932 (
*** Asterella bachmannii (Steph.) S.W.Arnell, Hepat. South Africa: 62, 1963 (
*** Asterella bolanderi (Austin) Underw., Bot. Gaz. 20 (2): 61, 1895 (
* Asterella caucasica (Steph.) H.Buch, A.Evans et Verd., Ann. Bryol. 10: 8, 1937 [1938] (
*** Asterella conocephala (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (
** Asterella coronata (Steph.) H.A.Mill., Phytologia 47 (4): 319, 1981 (
** Asterella dioica (Steph.) H.A.Mill., Phytologia 47 (4): 319, 1981 (
* Asterella dognyensis H.A.Mill., Phytologia 47 (4): 319, 1981 (
*** Asterella drummondii (Taylor) R.M.Schust. ex D.G.Long, J. Bryol. 21 (1): 76, 1999 (
*** Asterella echinella (Gottsche) Underw., Bot. Gaz. 20 (2): 62, 1895 (
*** Asterella elegans (Spreng.) Trevis., Rendiconti Reale Ist. Lombardo Sci. (ser. 2) 7: 785, 1874 (
*** Asterella heteroflora (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
*** Asterella innovans (Austin) H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 529, 1963 (
*** Asterella lateralis M.Howe, Bull. Torrey Bot. Club 25 (4): 189, 1898 (
*** Asterella lindenbergiana (Corda ex Nees) Lindb. ex Arnell, Lebermoosstud. nördl. Norwegen: 2, 1892 (
*** Asterella linearis (Steph.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898 (
*** Asterella longebarbata (Steph.) H.A.Mill., Phytologia 47 (4): 320, 1981 (
*** Asterella macropoda (Spruce) A.Evans, Bull. Torrey Bot. Club 46 (12): 472, 1919 (
*** Asterella marginata (Nees) S.W.Arnell, Hepat. South Africa: 63, 1963 (
** Asterella muelleri (Gottsche) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (
*** Asterella multiflora (Steph.) Kachroo, J. Hattori Bot. Lab. 19: 3, 1958 (
*** Asterella mussuriensis (Kashyap) Verd., Ann. Bryol. 8: 156, 1935 (
*** Asterella mussuriensis subsp. crassa (Shimizu et S.Hatt.) D.G.Long, Lindbergia 26 (1): 44, 2001 (
*** Asterella pappii (Gola) Grolle, Feddes Repert. 87 (3/4): 246, 1976 (
*** Asterella persica (Steph.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898 (
* Asterella preussii (Schiffn.) M.Howe, Bull. Torrey Bot. Club 25 (4): 191, 1898 (
** Asterella setisquama (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (
*** Asterella syngenesica (Bory) Grolle, Lindbergia 2 (3/4): 230, 1974 (
** Asterella tasmanica (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (
*** Asterella versicolor A.Evans, Contr. U.S. Natl. Herb. 20: 307, 1920 (
*** Asterella vulcanica (Schiffn.) Kachroo et Bapna, J. Indian Bot. Soc. 56 (1): 75, 1977 (
*** Asterella whiteleggeana (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 298, 1963 (
*** Asterella wilmsii (Steph.) S.W.Arnell, Hepat. South Africa: 62, 1963 (
** subg. Saccatae (Grolle) D.G.Long, J. Bryol. 22 (2): 113, 2000 (
*** Asterella alpina (Steph.) D.G.Long, J. Hattori Bot. Lab. 93: 9, 2003 (
*** Asterella grollei D.G.Long, Bryologist 102 (2): 169, 1999 (
*** Asterella muscicola (Steph.) S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 263, 1957 (
*** Asterella palmeri (Austin) Underw., Bot. Gaz. 20 (2): 63, 1895 (
*** Asterella pringlei Underw., Bot. Gaz. 20 (2): 64, 1895 (
*** Asterella rugosa A.Evans, Contr. U.S. Natl. Herb. 20: 289, 1920 (
*** Asterella saccata (Wahlenb.) A.Evans, Contr. U.S. Natl. Herb. 20: 276, 1920 (
** subg. Wallichianae D.G.Long, Lindbergia 26 (1): 43, 2001 (
** sect. Californicae D.G.Long, J. Hattori Bot. Lab. 97: 257, 2005 (
*** Asterella californica (Hampe ex Austin) Underw., Bot. Gaz. 20 (2): 60, 1895 (
** sect. Wallichianae D.G.Long, Phytotaxa 173 (1): 87, 2014 (
*** Asterella wallichiana (Lehm. et Lindenb.) Grolle, Khumbu Himal 1 (4): 262, 1966 (
** Cryptomitrium Austin ex Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 36, 1884 (
*** Cryptomitrium himalayense Kashyap, New Phytol. 14 (1): 2, 1915 (
*** Cryptomitrium oreades Perold, Bothalia 24 (2): 149, 1994 (
*** Cryptomitrium tenerum (Hook.) Austin ex Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 36, 1884 (
*** Mannia Corda, Gen. hepat.: 646, 1829 (
** subg. Mannia
*** Mannia androgyna (L.) A.Evans, Chron. Bot. 4: 224, 1938 (
*** Mannia californica (Gottsche) L.C.Wheeler, Bryologist 37 (5): 88, 1934 [1935] (
*** Mannia controversa (Meyl.) D.B.Schill, Edinburgh J. Bot. 65 (1): 36, 2008 (
*** Mannia controversa subsp. asiatica D.B.Schill et D.G.Long, Edinburgh J. Bot. 65 (1): 45, 2008 (
*** Mannia fragrans (Balb.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 62, 1937 (
** Mannia fragrans subsp. orientalis R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 201, 1992 (
* Mannia perssonii Udar et V.Chandra, Canad. J. Bot. 43 (1): 150, 1965 (
*** Mannia sibirica (Müll.Frib.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 66, 1937 (
** subg. Neesiella (Schiffn.) D.B.Schill et D.G.Long, Bryologist 113 (1): 175, 2010 (
*** Mannia gracilis (F.Weber) D.B.Schill et D.G.Long, Bryologist 113 (1): 173, 2010 (
* Mannia hegewaldii Bischl., Fl. Neotrop. Monogr. 97: 184, 2005 (
*** Mannia pilosa (Hornem.) Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (1): 64, 1937 (
*** Mannia triandra (Scop.) Grolle, J. Bryol. 8 (4): 487, 1975 (
Incertae sedis
* Mannia paradoxa R.M.Schust., Phytologia 57 (6): 410, 1985 (
*** Plagiochasma Lehm., Nov. Stirp. Pug. 4: 13, 1832 (
* Plagiochasma megacarpon (Griff.) Steph., Bull. Herb. Boissier 6 (10): 789 (86), 1898 (
** subg. Micropylum Bischl., Rev. Bryol. Lichénol. 43 (1): 103, 1977 (
*** Plagiochasma rupestre (J.R.Forst. et G.Forst.) Steph., Bull. Herb. Boissier 6 (10): 783 (80), 1898 (
*** Plagiochasma rupestre var. volkii Bischl., Rev. Bryol. Lichénol. 44 (3): 289, 1978 (
** subg. Plagiochasma
*** Plagiochasma appendiculatum Lehm. et Lindenb., Nov. Stirp. Pug. 4: 14, 1832 (
*** Plagiochasma argentinicum Bischl., Rev. Bryol. Lichénol. 45 (3): 301, 1979 (
*** Plagiochasma beccarianum Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898 (
*** Plagiochasma cordatum Lehm. et Lindenb., Nov. Stirp. Pug. 4: 13, 1832 (
*** Plagiochasma crenulatum Gottsche, Mexik. Leverm.: 266, 1863 (
*** Plagiochasma cuneatum A.Evans, Amer. J. Bot. 19 (7): 627, 1932 (
*** Plagiochasma eximium (Schiffn.) Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898 (
*** Plagiochasma intermedium Lindenb. et Gottsche, Syn. Hepat. 4: 513, 1846 (
*** Plagiochasma jamaicense (Haynes) A.Evans, Bull. Torrey Bot. Club 42 (5): 292, 1915 (
*** Plagiochasma japonicum (Steph.) C.Massal., Hepat. Shen-si: 47, 1897 (
*** Plagiochasma landii A.Evans, Bull. Torrey Bot. Club 42 (5): 298, 1915 (
*** Plagiochasma microcephalum (Steph.) Steph., Bull. Herb. Boissier 6 (10): 781 (78), 1898 (
*** Plagiochasma microcephalum var. tunesicum Bischl., Rev. Bryol. Lichénol. 44 (3): 247, 1978 (
*** Plagiochasma muenchianum Steph., Sp. Hepat. (Stephani) 6: 9, 1917 (
*** Plagiochasma pterospermum C.Massal., Hepat. Shen-si: 46, 1897 (
*** Plagiochasma wrightii Sull., Musc. Hepat. U.S.: 688, 1856 (
Incertae sedis
** Plagiochasma udarii A.Alam et S.C.Srivast., Indian J. Forest. 32 (4): 631, 2009 (
*** Reboulia Raddi, Opusc. Sci. 2 (6): 357, 1818 (
*** Reboulia hemisphaerica (L.) Raddi, Opusc. Sci. 2 (6): 357, 1818 (
** Reboulia hemisphaerica subsp. acrogyna (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 168, 1992 (
** Reboulia hemisphaerica subsp. australis R.M.Schust., Phytologia 56 (7): 460, 1985 (
** Reboulia hemisphaerica subsp. dioica R.M.Schust., Phytologia 56 (7): 462, 1985 (
** Reboulia hemisphaerica var. fissisquama Herzog, Symb. Sin. 5: 5, 1930 (
** Reboulia hemisphaerica subsp. orientalis R.M.Schust., Phytologia 56 (7): 461, 1985 (
** Reboulia hemisphaerica var. turkestanica C.E.O.Jensen ex Herzog, Symb. Sin. 5: 5, 1930 (
by D.G. Long
The genera of Cleveaceae were re-defined by
*** Athalamia Falc., Ann. Mag. Nat. Hist. (ser. 2) 1 (5): 375, 1848 (
* Athalamia dioica Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 348, 1916 (
*** Athalamia pinguis Falc., Ann. Mag. Nat. Hist. (ser. 2) 1 (5): 375, 1848 (
* Athalamia pulcherrima (Steph.) S.Hatt., J. Hattori Bot. Lab. 12: 54, 1954 (
*** Clevea Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 289, 1868 (
*** Clevea hyalina (Sommerf.) Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 291, 1868 (
* Clevea hyalina var. californica M.Howe, Mem. Torrey Bot. Club 7: 38, 1899 (
* Clevea pedicellata (Griff.) Lindb., Acta Soc. Fauna Fl. Fenn. 2 (3): 11, 1882 (
*** Clevea pusilla (Steph.) Rubas. et D.G.Long, J. Bryol. 33 (2): 167, 2011 (
*** Clevea spathysii (Lindenb.) Müll.Frib., Hedwigia 79 (1/2): 75, 1940 (
*** Peltolepis Lindb., Morgonbladet (Helsinki) 1876 (106, 9 May): 1, 1876 (
** Peltolepis japonica (Shimizu et S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 14: 103, 1955 (
*** Peltolepis quadrata (Saut.) Müll.Frib., Hedwigia 79 (1/2): 74, 1940 (
*** Sauteria Nees, Naturgesch. Eur. Leberm. 4: 139, 1838 (
** sect. Sauchia (Kashyap) R.M.Schust., Phytologia 57 (6): 411, 1985 (
* Sauteria japonica (Shimizu et S.Hatt.) S.Hatt., J. Hattori Bot. Lab. 12: 62, 1954 (
*** Sauteria spongiosa (Kashyap) S.Hatt., J. Hattori Bot. Lab. 12: 62, 1954 (
** sect. Sauteria
*** Sauteria alpina (Nees) Nees, Naturgesch. Eur. Leberm. 4: 143, 1838 (
* Sauteria inflata C.Gao et K.C.Chang, Acta Bot. Yunnan. 3 (4): 389, 1981 (
Incertae sedis
* Sauteria chilensis (Lindenb.) Grolle, J. Hattori Bot. Lab. 58: 200, 1985 (
* Sauteria crassipes Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 229, 1869 (
* Sauteria nyikaensis Perold, Bothalia 33 (2): 167, 2003 (
by D.G. Long
*** Conocephalum Hill, Gener. Nat. Hist. 2 Hist. pl. (ed. 2): 118, 1773 (
** subg. Conocephalum
*** Conocephalum conicum (L.) Dumort., Commentat. Bot. (Dumortier): 115, 1822 (
*** Conocephalum salebrosum Szweyk., Buczk. et Odrzyk., Pl. Syst. Evol. 253 (1/4): 146, 2005 (
** subg. Sandea (Lindb.) Inoue, Ill. Jap. Hep. 2: 192, 1976 (
*** Conocephalum japonicum (Thunb.) Grolle, J. Hattori Bot. Lab. 55: 501, 1984 (
by D.G. Long
*** Corsinia Raddi, Opusc. Sci. 2 (6): 354, 1818 (
*** Corsinia coriandrina (Spreng.) Lindb., Hepaticol. Utveckl.: 30, 1877 (
*** Cronisia Berk., Introd. crypt. bot.: 434, 1857 (
*** Cronisia fimbriata (Nees) Whittem. et Bischl., Cryptog. Bryol. 22 (3): 170, 2001 (
*** Cronisia weddellii (Mont.) Grolle, J. Bryol. 9 (4): 532, 1977 [1978] (
by D.G. Long
*** Cyathodium Kunze, Nov. Stirp. Pug. 6: 17, 1834 (
*** Cyathodium aureonitens (Griff.) Mitten, J. Linn. Soc., Bot. 22 (146): 327 (
*** Cyathodium bischlerianum N.Salazar, Bryologist 104 (1): 141, 2001 (Salazar 2001).
*** Cyathodium cavernarum Kunze, Nov. Stirp. Pug. 6: 18, 1834 (
** Cyathodium denticulatum Udar et S.C.Srivast., Geophytology 1 (2): 166, 1971 (
*** Cyathodium foetidissimum Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 154, 1898 (
*** Cyathodium indicum Udar et D.K.Singh, J. Bryol. 10 (2): 139, 1978 [1979] (
*** Cyathodium mehranum D.K.Singh, Misc. Bryol. Lichenol. 9 (8): 173, 1983 (
*** Cyathodium smaragdinum Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 3: 480, 1910 (
*** Cyathodium spruceanum Prosk., Bryologist 54 (4): 243, 1951 [1952] (
* Cyathodium spurium (Dicks.) Lindb. ex Braithw., J. Bot. 16: 55, 1878 (
*** Cyathodium steerei Hässel, Rev. Bryol. Lichénol. 30 (3/4): 223, 1961 (
*** Cyathodium tuberculatum Udar et D.K.Singh, Bryologist 79 (2): 235, 1976 (
*** Cyathodium tuberosum Kashyap, New Phytol. 13 (6/7): 210, 1914 (
by D.G. Long
*** Dumortiera Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 410, 1824 [1825] (
*** Dumortiera hirsuta (Sw.) Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 410, 1824 [1825] (
* Dumortiera hirsuta subsp. nepalensis (Taylor) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 386, 1992 (
* Dumortiera hirsuta subsp. tatunoi Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 6: 38, 1951 (
by D.G. Long
*** Aitchisoniella Kashyap, New Phytol. 13 (6/7): 219, 1914 (
*** Aitchisoniella himalayensis Kashyap, New Phytol. 13 (6/7): 219, 1914 (
*** Exormotheca Mitt., Nat. hist. Azores: 325, 1870 (
** subg. Corbierella (Douin et Trab.) Schiffn., Hedwigia 81 (1/2): 71, 1942 (
*** Exormotheca bischlerae Furuki et Higuchi, Cryptog. Bryol. 27 (1): 98, 2006 (
*** Exormotheca holstii Steph., Bull. Herb. Boissier 7 (3): 219 (145), 1899 (
*** Exormotheca welwitschii Steph., Bull. Herb. Boissier 7 (3): 220 (146), 1899 (
** subg. Exormotheca
*** Exormotheca pustulosa Mitt., Nat. hist. Azores: 326, 1870 (
Incertae sedis
*** Exormotheca bulbigena Bornefeld, O.H.Volk et R.Wolf, Bothalia 26 (2): 159, 1996 (
*** Exormotheca ceylonensis Meijer, J. Hattori Bot. Lab. 16: 72, 1956 (
* Exormotheca gollanii Steph., Sp. Hepat. (Stephani) 6: 18, 1917 (
*** Exormotheca tuberifera Kashyap, New Phytol. 13 (9): 309, 1914 (
*** Stephensoniella Kashyap, New Phytol. 13 (9): 312, 1914 (
*** Stephensoniella brevipedunculata Kashyap, New Phytol. 13 (9): 312, 1914 (
by D.G. Long
The treatment of Marchantiaceae is mainly following
*** Bucegia Radian, Bull. Herb. Inst. Bot. Bucarest 3-4: 3, 1903 (
*** Bucegia romanica Radian, Bull. Herb. Inst. Bot. Bucarest 3-4: 4, 1903 (
*** Marchantia L., Sp. Pl. 1: 1137, 1753 (
** subg. Chlamidium (Corda) Bischl., Cryptog. Bryol. Lichénol. 3 (4): 362, 1982 (
*** Marchantia breviloba A.Evans, Trans. Connecticut Acad. Arts 21 (3): 265, 1917 (
*** Marchantia inflexa Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 43, 1838 (
** sect. Chlamidium (Corda) Nees, Naturgesch. Eur. Leberm. 4: 101, 1838 (
*** Marchantia chenopoda L., Sp. Pl. 1: 1137, 1753 (
*** Marchantia crenata Austin, Bull. Torrey Bot. Club 5 (3): 14, 1874 (
*** Marchantia foliacea Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 168, 1855 (
*** Marchantia formosana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 121, 1934 (
*** Marchantia globosa Brid., Hist. Musc. Hepat. Prodr.: 102, 1815 (
*** Marchantia hexaptera Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 957, 1866 (
*** Marchantia linearis Lehm. et Lindenb., Nov. Stirp. Pug. 4: 8, 1832 (
*** Marchantia miqueliana Lehm., Nov. Stirp. Pug. 10: 20, 1857 (
*** Marchantia novoguineensis Bischl., Bryophyt. Biblioth. 38: 130, 1989 (
*** Marchantia pappeana Lehm., Nov. Stirp. Pug. 10: 21, 1857 (
** Marchantia pappeana subsp. robusta (Steph.) Bischl., Bryophyt. Biblioth. 45: 91, 1993 (
*** Marchantia pileata Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 169, 1855 (
*** Marchantia pinnata Steph., Candollea 14: 109, 1953 (
*** Marchantia rubribarba Steph., Bull. Herb. Boissier 7 (5): 400 (172), 1899 (
*** Marchantia vitiensis Steph., Bull. Herb. Boissier 7 (7): 520 (182), 1899 (
** sect. Paleaceae Bischl., Bryophyt. Biblioth. 38: 90, 1989 (
*** Marchantia paleacea Bertol., Opusc. Sci. 1: 242, 1817 (
*** Marchantia paleacea subsp. diptera (Nees et Mont.) Inoue, J. Jap. Bot. 64 (7): 194, 1989 (
** sect. Papillatae Bischl., Cryptog. Bryol. Lichénol. 10 (1): 69, 1989 (
*** Marchantia debilis K.I.Goebel, Organogr. Pfl., ed. 2, 2 (1): 901, 1915 (
*** Marchantia emarginata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 192, 1824 [1825] (
*** Marchantia emarginata subsp. lecordiana (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 78, 1989 (
*** Marchantia emarginata subsp. tosana (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 77, 1989 (
*** Marchantia papillata Raddi, Critt. Brasil.: 20, 1822 (
*** Marchantia papillata subsp. grossibarba (Steph.) Bischl., Cryptog. Bryol. Lichénol. 10 (1): 78, 1989 (
** subg. Marchantia
*** Marchantia plicata Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 43, 1838 (
** sect. Berteroanae R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 310, 1992 (
*** Marchantia berteroana Lehm. et Lindenb., Nov. Stirp. Pug. 6: 21, 1834 (
** sect. Marchantia
*** Marchantia polymorpha L., Sp. Pl. 1: 1137, 1753 (
*** Marchantia polymorpha subsp. montivagans Bischl. et Boissel.-Dub., J. Bryol. 16 (3): 364, 1991 (
*** Marchantia polymorpha subsp. ruderalis Bischl. et Boissel.-Dub., J. Bryol. 16 (3): 364, 1991 (
** subg. Protomarchantia R.M.Schust., Phytologia 57 (6): 410, 1985 (
** sect. Protomarchantia (R.M.Schust.) L.Söderstr., Phytotaxa 202 (1): 69, 2015 (
*** Marchantia acaulis Steph., Bull. Herb. Boissier 7 (7): 533 (195), 1899 (
*** Marchantia antiqua Steph., Candollea 14: 103, 1953 (
*** Marchantia carrii Bischl., Bryophyt. Biblioth. 38: 256, 1989 (
*** Marchantia geminata Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 194, 1824 [1825] (
*** Marchantia hartlessiana Steph., Candollea 14: 107, 1953 (
*** Marchantia macropora Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 169, 1855 (
*** Marchantia philippinensis Bischl., Bryophyt. Biblioth. 38: 245, 1989 (
*** Marchantia solomonensis Bischl., Bryophyt. Biblioth. 38: 281, 1989 (
*** Marchantia streimannii Bischl., Bryophyt. Biblioth. 38: 250, 1989 (
*** Marchantia subintegra Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 125, 1860 [1861] (
*** Marchantia treubii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 160, 1898 (
*** Marchantia wallisii J.B.Jack et Steph., Bull. Herb. Boissier 7 (7): 520 (182), 1899 (
** sect. Subgeminatae Bischl., Bryophyt. Biblioth. 38: 219, 1989 (
*** Marchantia subgeminata Steph., Bull. Herb. Boissier 7 (7): 530 (192), 1899 (
Incertae sedis
* Marchantia assamica Griff., Not. pl. asiat. 2: 327, 1849 (
* Marchantia balboi Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 2, 1916 (
* Marchantia balboi var. acutisquamata Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 472, 1947 (
* Marchantia cagnii Gola, Ann. Bot. (Rome) 6 (2): 271, 1907 (
* Marchantia cengiana Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 473, 1947 (
* Marchantia friedrichsthaliana Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 438, 1877 (
* Marchantia keniae Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916 (
* Marchantia papyracea Gola, Ann. Bot. (Rome) 6 (2): 271, 1907 (
* Marchantia quadriloba Steph., Candollea 14: 110, 1953 (
* Marchantia sellae Gola, Ann. Bot. (Rome) 6 (2): 271, 1907 (
** Marchantia stoloniscyphulus (C.Gao et K.C.Chang) Piippo, J. Hattori Bot. Lab. 68: 134, 1990 (
* Marchantia trilocularis Roth, Tent. Fl. Germ. 1: 487, 1788 (
* Marchantia tusui Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 3, 1916 (
** Preissia Corda, Gen. hepat.: 647, 1829 (
*** Preissia quadrata (Scop.) Nees, Naturgesch. Eur. Leberm. 4: 135, 1838 (
** Preissia quadrata subsp. hyperborea R.M.Schust., Phytologia 57 (6): 410, 1985 (
by D.G. Long
*** Monoclea Hook., Musci Exot. 2: tab. clxxiv, 1820 (
*** Monoclea forsteri Hook., Musci Exot. 2: tab. clxxiv, 1820 (
*** Monoclea gottschei Lindb., Rev. Bryol. 13 (6): 102, 1886 (
*** Monoclea gottschei subsp. elongata Gradst. et Mues, Pl. Syst. Evol. 180 (1/2): 133, 1992 (
by D.G.Long
*** Monosolenium Griff., Not. pl. asiat. 2: 341, 1849 (
*** Monosolenium tenerum Griff., Not. pl. asiat. 2: 341, 1849 (
by D.G. Long
*** Oxymitra Bisch. ex Lindenb., Syn. hepat. eur: 124, 1829 (
*** Oxymitra cristata Garside, Bothalia 23 (2): 211, 1993 (
*** Oxymitra incrassata (Brot.) Sérgio et Sim-Sim, J. Bryol. 15 (4): 662, 1989 (
by R. Stotler, B.J. Crandall-Stotler and D.C. Cargill
** Riccia L., Sp. Pl. 1: 1138, 1753 (
** subg. Chartaceae Perold, Bothalia 16 (1): 29, 1986 (
*** Riccia schelpei O.H.Volk et Perold, Bothalia 16 (1): 29, 1986 (
** subg. Leptoriccia R.M.Schust., Phytologia 56 (2): 72, 1984 (
*** Riccia membranacea Gottsche et Lindenb., Syn. Hepat. 4: 608, 1846 (
** subg. Riccia
*** Riccia albida Sull. ex Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 231, 1869 (
*** Riccia albopunctata Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 237, 1991 (
*** Riccia australis Steph., Bull. Herb. Boissier 6 (4): 337 (29), 1898 (
*** Riccia boliviensis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 242, 1991 (
*** Riccia brasiliensis Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 6, 1964 (
*** Riccia breutelii Hampe, Bull. Herb. Boissier 6 (4): 325 (17), 1898 (
*** Riccia brittonii M.Howe, Ann. Missouri Bot. Gard. 2 (1/2): 50, 1915 (
* Riccia chudoana Steph., Sp. Hepat. (Stephani) 6: 1, 1917 (
*** Riccia coracina Jovet-Ast, Cryptog. Bryol. 24 (3): 212, 2003 (
*** Riccia corrugata Jovet-Ast, Cryptog. Bryol. 21 (4): 308, 2000 (
*** Riccia crassivenia Jovet-Ast, Cryptog. Bryol. 21 (4): 312, 2000 (
*** Riccia cubensis S.W.Arnell, Bryologist 61 (2): 142, 1958 (
*** Riccia discolor Lehm. et Lindenb., Nov. Stirp. Pug. 4: 1, 1832 (
*** Riccia ekmanii S.W.Arnell, Bryologist 61 (2): 143, 1958 (
*** Riccia elliottii Steph., Bull. Herb. Boissier 6 (4): 324 (16), 1898 (
*** Riccia enyae Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 230, 1991 (
*** Riccia erythrocarpa Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 257, 1991 (
*** Riccia fruchartii Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898 (
*** Riccia gangetica Ahmad ex L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 57, 2012 (
*** Riccia grandis Nees, Fl. Bras. (Martius) 1 (1): 300, 1833 (
*** Riccia helenae Jovet-Ast, J. Hattori Bot. Lab. 74: 96, 1993 (
** Riccia hirta (Austin) Underw., Bot. Gaz. 19 (7): 274, 1894 (
*** Riccia horrida Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 226, 1991 (
*** Riccia hortorum Bory, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 435, 1836 [1837] (
*** Riccia howellii M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 202, 1934 (
* Riccia ianthina Jovet-Ast, Rev. Bryol. Lichénol. 44 (4): 418, 1978 (
*** Riccia inflexa Taylor, London J. Bot. 5: 417, 1846 (
*** Riccia iodocheila M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 200, 1934 (
*** Riccia lanceolata Steph., Hedwigia 27 (3/4): 110, 1888 (
*** Riccia lindmanii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 29, 1897 (
** Riccia macallisteri M.Howe, Bryologist 20 (3): 35, 1917 (
*** Riccia macrospora Steph., Bull. Herb. Boissier 6 (4): 328 (20), 1898 (
** Riccia mamillata Trab. ex Steph., Rev. Bryol. 16 (5): 65, 1889 (
*** Riccia mauryana Steph., Bull. Herb. Boissier 6 (4): 327 (19), 1898 (
*** Riccia olgensis Na-Thalang, Brunonia 3 (1): 100, 1980 (
*** Riccia planobiconvexa Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 23 (III, 2): 29, 1897 (
*** Riccia ridleyi A.Gepp, J. Linn. Soc., Bot. 27 (181): 74, 1890 (
*** Riccia sanguineisporis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 253, 1991 (
*** Riccia squamata Nees, Fl. Bras. (Martius) 1 (1): 302, 1833 (
*** Riccia subdepilata Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 228, 1991 (
*** Riccia subplana Steph., Symb. Antill. (Urban) 3 (2): 275, 1902 (
*** Riccia taeniiformis Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 270, 1991 (
*** Riccia viannae Jovet-Ast, Cryptog. Bryol. Lichénol. 12 (3): 261, 1991 (
*** Riccia vitalii Jovet-Ast, Mem. New York Bot. Gard. 45: 285, 1987 (
*** Riccia weinionis Steph., Bull. Herb. Boissier 6 (4): 326 (18), 1898 (
** sect. Pilifer O.H.Volk, Mitt. Bot. Staatssamml. München 19: 455, 1983 (
*** Riccia alatospora O.H.Volk et Perold, Bothalia 15 (3/4): 534, 1985 (
*** Riccia albomarginata Bisch. ex C.Krauss, Flora 29 (9): 135, 1846 (
*** Riccia albovestita O.H.Volk, Mitt. Bot. Staatssamml. München 17: 245, 1981 (
*** Riccia ampullacea Perold, Bothalia 20 (2): 168, 1990 (
*** Riccia concava Bisch. ex C.Krauss, Flora 29 (9): 135, 1846 (
*** Riccia elongata Perold, Bothalia 20 (2): 167, 1990 (
*** Riccia furfuracea Perold, Bothalia 20 (2): 176, 1990 (
*** Riccia hantamensis Perold, Bothalia 19 (2): 157, 1989 (
*** Riccia hirsuta O.H.Volk et Perold, Bothalia 16 (2): 187, 1986 (
*** Riccia namaquensis Perold, Bothalia 20 (2): 180, 1990 (
*** Riccia parvoareolata O.H.Volk et Perold, Bothalia 15 (1/2): 117, 1984 (
*** Riccia pulveracea Perold, Bothalia 20 (2): 185, 1990 (
*** Riccia radicosa Pearson, Natuurw. Tijdschr. 4 (5/6): 142, 1922 (
*** Riccia simii Perold, Bothalia 20 (1): 36, 1990 (
*** Riccia trachyglossa Perold, Bothalia 20 (2): 172, 1990 (
*** Riccia villosa Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 724, 1913 (
*** Riccia vitrea Perold, Bothalia 20 (2): 178, 1990 (
** sect. Riccia
*** Riccia albolimbata S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 264, 1957 (
*** Riccia alboporosa Perold, Bothalia 19 (1): 12, 1989 (
*** Riccia albornata O.H.Volk et Perold, Bothalia 18 (2): 160, 1988 (
*** Riccia angolensis Steph., Bull. Herb. Boissier 6 (4): 323 (15), 1898 (
*** Riccia argenteolimbata O.H.Volk et Perold, Bothalia 18 (2): 155, 1988 (
** Riccia atlantica Sérgio et Perold, J. Bryol. 17 (1): 127, 1992 (
*** Riccia atromarginata Levier, Nuovo Giorn. Bot. Ital. 21 (2): 291, 1889 (
** Riccia atromarginata var. jovet-astiae Rauh et Buchloh, Rev. Bryol. Lichénol. 30 (1/2): 77, 1961 (
*** Riccia atropurpurea Sim, Trans. Roy. Soc. South Africa 15 (1): 11, 1926 (
*** Riccia beyrichiana Hampe, Nov. Stirp. Pug. 7: 1, 1838 (
*** Riccia bicarinata Lindb., Rev. Bryol. 4 (3): 41, 1877 (
*** Riccia bicolorata Perold, Bothalia 20 (2): 188, 1990 (
*** Riccia bifurca Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 95, 1795 [1796] (
*** Riccia billardierei Mont. et Nees, Syn. Hepat. 4: 602, 1846 (
*** Riccia breidleri Jur. ex Steph., Hedwigia 24 (1): 6, 1885 (
*** Riccia californica Austin, Bull. Torrey Bot. Club 6 (7): 46, 1875 (
*** Riccia campbelliana M.Howe, Mem. Torrey Bot. Club 7: 26, 1899 (
*** Riccia ciliata Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 95, 1795 [1796] (
*** Riccia ciliifera Link, Syn. hepat. eur: 119, 1829 (
* Riccia congoana Steph., Bull. Herb. Boissier 6 (4): 328 (20), 1898 (
*** Riccia crinita Taylor, London J. Bot. 5: 415, 1846 (
*** Riccia crozalsii Levier, Rev. Bryol. 29 (4): 73, 1902 (
* Riccia crustata Trab., Bull. Soc. Hist. Nat. Afrique N. 7: 87, 1916 (
** Riccia dictyospora M.Howe, Bull. Torrey Bot. Club 28 (3): 163, 1901 (
*** Riccia glauca L., Sp. Pl. 1: 1139, 1753 (
** Riccia glauca var. ciliaris Warnst., Verh. Bot. Vereins Prov. Brandenburg 27 (1): 87, 1886 (
** Riccia gothica Damsh. et Hallingb., Lindbergia 12 (2/3): 100, 1986 [1987] (
*** Riccia gougetiana Durieu et Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 35, 1849 (
** Riccia gougetiana var. armatissima Levier ex Müll.Frib., Lebermoose 1 (3): 161, 1907 (
*** Riccia lamellosa Raddi, Opusc. Sci. 2 (6): 351, 1818 (
*** Riccia ligula Steph., Bull. Herb. Boissier 6 (4): 315 (7), 1898 (
*** Riccia limbata Bisch. ex C.Krauss, Flora 29 (9): 135, 1846 (
*** Riccia macrocarpa Levier, Bull. Soc. Bot. Ital. 1894: 114, 1894 (
*** Riccia mammifera O.H.Volk et Perold, Bothalia 16 (2): 176, 1986 (
*** Riccia melitensis C.Massal., Bull. Soc. Bot. Ital. 1913 (2/3): 52, 1913 (
*** Riccia michelii Raddi, Opusc. Sci. 2 (6): 352, 1818 (
*** Riccia microciliata O.H.Volk et Perold, Bothalia 16 (2): 173, 1986 (
*** Riccia montana Perold, Bothalia 19 (1): 9, 1989 (
*** Riccia natalensis Sim, Trans. Roy. Soc. South Africa 15 (1): 9, 1926 (
*** Riccia nigrella DC., Fl. Franç. (DC. & Lamarck), 5 (6): 193, 1815 (
*** Riccia okahandjana S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 268, 1957 (
** Riccia ozarkiana McGregor, Bryologist 63 (1): 30, 1960 (
*** Riccia papillosa Moris, Stirp. Sard. Elench.: 18, 1829 (
*** Riccia pottsiana Sim, Trans. Roy. Soc. South Africa 15 (1): 10, 1926 (
*** Riccia rosea O.H.Volk et Perold, Bothalia 16 (2): 181, 1986 (
* Riccia runssorensis Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898 (
*** Riccia sommieri Levier, Isola Giglio: 119, 1900 (
*** Riccia sorocarpa Bisch., Bem. Leberm.: 145, 1835 (
** Riccia sorocarpa var. heegii Schiffn., Hedwigia 53 (1/2): 36, 1912 (
*** Riccia subbifurca Warnst. ex Croz., Rev. Bryol. 30 (4): 62, 1903 (
** Riccia tenella D.L.Jacobs, Bryologist 52 (4): 168, 1949 [1950] (
*** Riccia trabutiana Steph., Rev. Bryol. 16 (5): 65, 1889 (
*** Riccia violacea M.Howe, Ann. Missouri Bot. Gard. 2 (1/2): 51, 1915 (
*** Riccia violacea var. laevis Jovet-Ast, Cryptog. Bryol. Lichénol. 10 (2): 100, 1989 (
*** Riccia warnstorfii Limpr. ex Warnst., Verh. Bot. Vereins Prov. Brandenburg 27 (1): 85, 1886 (
** subg. Ricciella (A.Braun) Boulay, Musc. France 2: 198, 1904 (
*** Riccia cancellata Taylor, London J. Bot. 5: 414, 1846 (
*** Riccia cincta Jovet-Ast, Cryptog. Bryol. 21 (4): 303, 2000 (
*** Riccia cruciata Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 349, 1916 (
*** Riccia eburnea Jovet-Ast, Cryptog. Bryol. 21 (4): 300, 2000 (
*** Riccia hasskarliana Steph., Bull. Herb. Boissier 6 (5): 374 (49), 1898 (
*** Riccia junghuhniana Nees et Lindenb., Syn. Hepat. 4: 609, 1846 (
** Riccia junghuhniana var. simplex Schiffn., Hep. Fl. Buitenzorg: 14, 1900 (
*** Riccia mangalorica Ahmad ex Jovet-Ast, Cryptog. Bryol. 24 (3): 223, 2003 (
*** Riccia multifida (Steph.) Steph., Bull. Herb. Boissier 6 (5): 365 (40), 1898 (
** Riccia polycarpa (Trab.) Jelenc, Bull. Trimestriel Geogr. Archeol. Oran 73 (228): 88, 1950 (
*** Riccia porosa Taylor, London J. Bot. 5: 416, 1846 (
*** Riccia pullulans Jovet-Ast, Cryptog. Bryol. Lichénol. 18 (3): 183, 1997 (
** sect. Ricciella (A.Braun) Bisch., Bem. Leberm.: 160, 1835 (
* Riccia bahiensis Steph., Bull. Herb. Boissier 6 (5): 375 (50), 1898 (
*** Riccia canaliculata Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 96, 1795 [1796] (
*** Riccia chiapasensis Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 235, 1993 (
*** Riccia crassifrons Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 570, 1885 (
*** Riccia duplex Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 100, 1941 (
** Riccia duplex var. megaspora Na-Thalang, Brunonia 3 (1): 128, 1980 (
*** Riccia dussiana Steph., Symb. Antill. (Urban) 3 (2): 275, 1902 (
*** Riccia fluitans L., Sp. Pl. 1: 1139, 1753 (
*** Riccia frostii Austin, Bull. Torrey Bot. Club 6 (3): 17, 1875 (
* Riccia frostii var. crystallinoides Schiffn., Ann. K. K. Naturhist. Hofmus. 27: 503, 1913 (
*** Riccia geissleriana Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 236, 1993 (
*** Riccia hegewaldiana Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 238, 1993 (
*** Riccia huebeneriana Lindenb., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 504d, 1836 [1837] (
* Riccia huebeneriana subsp. sullivantii (Austin) R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 457, 1992 (
*** Riccia jovet-astiae E.Vianna, Bol. Inst. Bioci. Univ. Fed. Rio Grande do Sul 38: 165, 1985 (
*** Riccia limicola Jovet-Ast, Rev. Bryol. Lichénol. 44 (4): 422, 1978 (
*** Riccia paraguayensis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcvi, 1889 [1890] (
*** Riccia paranaensis Hässel, Opera Lilloana 7: 228, 1962 [1963] (
*** Riccia perennis Steph., Bull. Herb. Boissier 6 (5): 372 (47), 1898 (
*** Riccia purpurascens Lehm., Linnaea 4: 371, 1829 (
** Riccia rhenana Lorb. ex Müll.Frib., Hedwigia 80 (1/2): 94, 1941 (
** Riccia rhenana var. violacea M.F.Boiko, Chornom. Bot. J. 7: 93, 2011 (
*** Riccia stricta (Lindenb.) Perold, Bothalia 20 (2): 197, 1990 (
** sect. Spongodes Nees, Naturgesch. Eur. Leberm. 4: 391, 1838 (
*** Riccia bullosa Link, Syn. hepat. eur: 119, 1829 (
*** Riccia cavernosa Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 95, 1795 [1796] (
*** Riccia crystallina L., Sp. Pl. 1: 1138, 1753 (
*** Riccia cupulifera A.V.Duthie, Trans. Roy. Soc. South Africa 24 (2): 116, 1936 (
*** Riccia garsidei Sim, Trans. Roy. Soc. South Africa 15 (1): 13, 1926 (
*** Riccia moenkemeyeri Steph., Bot. Jahrb. Syst. 8 (2): 95, 1886 (
*** Riccia rubricollis Garside et A.V.Duthie ex Perold, Bothalia 21 (1): 51, 1991 (
*** Riccia volkii S.W.Arnell, Mitt. Bot. Staatssamml. München 2 (16): 271, 1957 (
*** Riccia vulcanicola Eb.Fisch., Trop. Bryol. 8: 70, 1993 (
** subg. Thallocarpus (Lindb.) Jovet-Ast, Cryptog. Bryol. Lichénol. 14 (3): 220, 1993 (
*** Riccia curtisii (Austin) Austin, Bull. Torrey Bot. Club 6 (52): 305, 1879 (
** Riccia leptothallus R.M.Schust., J. Hattori Bot. Lab. 71: 271, 1992 (
*** Riccia perssonii Sultan Khan, Svensk Bot. Tidskr. 49 (3): 433, 1955 (
** subg. Triseriata Jovet-Ast, Cryptog. Bryol. Lichénol. 17 (2): 132, 1996 (
*** Riccia singularis Jovet-Ast, Cryptog. Bryol. Lichénol. 17 (2): 127, 1996 (
Incertae sedis
** Riccia abuensis Bapna, Trans. Brit. Bryol. Soc. 4 (2): 249, 1962 (
** Riccia acutisulca Steph., Sp. Hepat. (Stephani) 6: 1, 1917 (
* Riccia amboinensis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 44, 1890 (
** Riccia aravalliensis Pandé et Udar, J. Indian Bot. Soc. 36 (3): 249, 1957 (
* Riccia arnellii Sultan Khan, Bryologist 60 (1): 29, 1957 (
** Riccia asprella Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1059, 1888 (
* Riccia asservanda De Not. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 138, 1919 (
** Riccia attenuata Pandé, Proc. Natl. Inst. Sci. India B 25 (2): 92, 1959 (
* Riccia balansae Steph., Bull. Herb. Boissier 6 (5): 370 (45), 1898 (
* Riccia bialbistrata Hässel, Opera Lilloana 7: 243, 1962 [1963] (
** Riccia biokoensis Perold, Nova Hedwigia 64 (1/2): 244, 1997 (
** Riccia blackii Na-Thalang, Brunonia 3 (1): 81, 1980 (
*** Riccia caroliniana Na-Thalang, Brunonia 3 (1): 72, 1980 (
** Riccia cartilaginosa Steph., Hedwigia 28 (4): 272, 1889 (
* Riccia chartacea K.I.Goebel, Organogr. Pfl., ed. 2, 2 (1): 630, 1915 (
** Riccia chinensis Herzog, Symb. Sin. 5: 1, 1930 (
** Riccia collata Na-Thalang, Brunonia 3 (1): 122, 1980 (
** Riccia compacta Garside, Trans. Roy. Soc. South Africa 27 (1): 17, 1939 (
** Riccia convexa Steph., Sp. Hepat. (Stephani) 6: 2, 1917 (
* Riccia coronata Sim, Trans. Roy. Soc. South Africa 15 (1): 9, 1926 (
** Riccia crassa Steph., Bull. Herb. Boissier 6 (5): 376 (51), 1898 (
** Riccia crenatodentata O.H.Volk, Nova Hedwigia 46 (1/2): 27, 1988 (
** Riccia delavayi Steph., Bull. Herb. Boissier 6 (5): 367 (42), 1898 (
** Riccia deserticola Steph., Bull. Herb. Boissier 6 (5): 373 (48), 1898 (
*** Riccia erubescens Perold, J. Bryol. 16 (3): 371, 1991 (
** Riccia esulcata Steph., Sp. Hepat. (Stephani) 6: 2, 1917 (
** Riccia fertilissima Steph., Sp. Hepat. (Stephani) 6: 2, 1917 (
* Riccia gemmifera O.H.Volk, Nova Hedwigia 39: 131, 1984 (
** Riccia grollei Udar, Curr. Sci. 34 (4): 126, 1965 (
** Riccia handelii Schiffn., Symb. sin. 2: 81, 1937 (
** Riccia hawaiiensis Hürl., Phytologia 61 (5): 339, 1986 (
** Riccia indica Udar et A.Gupta, Proc. V Indian Geophytol. Conf., Special Publ.: 307, 1984 (
** Riccia indira-gandhiensis Dabhade et A.Hasan, J. Bombay Nat. Hist. Soc. 83 (2): 400, 1986 (
* Riccia intermedia Roum., Mém. Soc. Arts Sci. Carcassonne 5: 198, 1888 (
** Riccia jodhpurensis Bapna, Bot. Not. 114 (2): 181, 1961 (
** Riccia kirinensis C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 117, 1978 (
*** Riccia laxisquamata (Steph.) Steph., Bull. Herb. Boissier 6 (5): 371 (46), 1898 (
** Riccia liaoningensis C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 113, 1978 (
** Riccia linearis (Schiffn.) Steph., Bull. Herb. Boissier 6 (5): 371 (46), 1898 (
** Riccia luticola Na-Thalang, Brunonia 3 (1): 123, 1980 (
** Riccia mamrensis Perold, Cryptog. Bryol. 26 (1): 68, 2005 (
* Riccia marginata Lindb., Meddel. Soc. Fauna Fl. Fenn. 1: 106, 1876 [1877] (
** Riccia melanospora Kashyap, Liverworts W. Himal. 1: 94, 1929 (
** Riccia miyakeana Schiffn., Österr. Bot. Z. 49 (11): 386, 1899 (
** Riccia muscicola Steph., Hedwigia 24 (1): 4, 1885 (
*** Riccia nigerica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (2): 225, 1957 (
* Riccia nigrescens Mont., Voy. Amér. Mérid., Bot. 7 (1): 15, 1839 (
** Riccia nipponica S.Hatt., J. Hattori Bot. Lab. 9: 38, 1953 (
** Riccia novo-hannoverana Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 44, 1890 (
** Riccia numeensis Steph., Bull. Herb. Boissier 6 (4): 343 (35), 1898 (
* Riccia obtusa Meijer, J. Hattori Bot. Lab. 20: 113, 1958 (
*** Riccia oerstediana Lindenb. et Hampe, Linnaea 24 (3): 304, 1851 [1852] (
** Riccia pandei Udar, J. Indian Bot. Soc. 38 (1): 149, 1959 (
*** Riccia papillispora Steph., Bull. Herb. Boissier 6 (4): 334 (26), 1898 (
** Riccia papulosa (Steph.) Steph., Bull. Herb. Boissier 6 (5): 377 (52), 1898 (
** Riccia papulosa var. variabilis Na-Thalang, Brunonia 3 (1): 112, 1980 (
** Riccia pathankotensis Kashyap, J. Bombay Nat. Hist. Soc. 24 (2): 349, 1916 (
* Riccia perthiana Steph. ex K.I.Goebel, Organogr. Pfl., ed. 2, 2 (1): 630, 1915 (
* Riccia prominens Meijer, J. Hattori Bot. Lab. 20: 111, 1958 (
** Riccia pseudofluitans C.Gao et K.C.Chang, Acta Phytotax. Sin. 16 (4): 116, 1978 (
** Riccia pubescens S.Hatt., Nat. Sci. Mus. 14 (6): 141, 1943 (
*** Riccia radiata Perold, Bothalia 34 (1): 23, 2004 (
** Riccia rechingeri Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 288, 1907 (
* Riccia reticulatula Udar, Bull. Bot. Soc. Univ. Saugar 13: 49, 1961 (
** Riccia rorida Na-Thalang, Brunonia 3 (1): 101, 1980 (
* Riccia saharensis Steph. ex Jovet-Ast, Rev. Bryol. Lichénol. 26 (1/2): 62, 1957 (
** Riccia satoi S.Hatt., Bot. Mag. (Tokyo) 62 (733/734): 109, 1949 (
* Riccia schroederi Steph., 52 (5): 304, 1912 (
*** Riccia schweinfurthii Steph., Bull. Herb. Boissier 6 (4): 339 (31), 1898 (
** Riccia sibayenii Perold, Bothalia 31 (1): 151, 2001 (
*** Riccia somaliensis Perold, J. Bryol. 16 (3): 367, 1991 (
** Riccia spongiosula Na-Thalang, Brunonia 3 (1): 113, 1980 (
* Riccia subtilis (Steph.) Steph., Bull. Herb. Boissier 6 (5): 364 (39), 1898 (
** Riccia sumatrana Meijer, J. Hattori Bot. Lab. 20: 114, 1958 (
*** Riccia symoensii Vanden Berghen, Explor. Hydrobiol. Lac Bangweolo Luapula: 191, 1972 (
** Riccia tasmanica Steph. ex Rodway, Tasm. Bryoph.: 4, 1917 (
*** Riccia tomentosa O.H.Volk et Perold, Bothalia 20 (1): 25, 1990 (
** Riccia treubiana Steph., Bull. Herb. Boissier 6 (4): 323 (15), 1898 (
* Riccia treubiana var. subrubescens Schiffn., Hep. Fl. Buitenzorg: 16, 1900 (
* Riccia triangularis Steph., Bull. Mus. Natl. Hist. Nat. 18 (2): 116, 1912 (
* Riccia tuberculata Poir., Encycl. (Lamarck) 6: 199, 1804 (
** Riccia udarii Kanwal, J. Indian Bot. Soc. 58 (3): 282, 1979 (
* Riccia velenovskyi Kavina, Arch. Přír. Výzk. Čech 16 (2): 75, 1915 (
** Riccia velimalaiana A.E.D.Daniels et P.Daniel, Bull. Bot. Surv. India 44 (1/4): 139, 2002 [2003] (
** Riccia victoriensis Steph., Bull. Herb. Boissier 6 (5): 370, 1898 (
** Riccia weymouthiana Steph. ex Rodway, Tasm. Bryoph.: 5, 1917 (
** Riccia wichurae Steph., Bull. Herb. Boissier 6 (4): 330 (22), 1898 (
** Ricciocarpos Corda, Gen. hepat.: 651, 1829 (
*** Ricciocarpos natans (L.) Corda, Gen. hepat.: 651, 1829 (
by D.G. Long
*** Targionia L., Sp. Pl. 1: 1136, 1753 (
** subg. Prototargionia R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 69, 1992 (
** Targionia stellaris (Müll.Frib.) Hässel, Opera Lilloana 7: 74, 1962 [1963] (
** subg. Targionia
*** Targionia hypophylla L., Sp. Pl. 1: 1136, 1753 (
** Targionia hypophylla subsp. linealis W.Frey et Kürschner, Nova Hedwigia 57 (1/2): 127, 1993 (
** Targionia lorbeeriana Müll.Frib., Hedwigia 79 (1/2): 78, 1940 (
Incertae sedis
* Targionia dioica Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 154, 1898 (
* Targionia elongata Bisch., Syn. Hepat. 4: 576, 1846 (
* Targionia fiorii Gola, Ann. Bot. (Rome) 13 (1): 62, 1914 (
* Targionia formosica Horik., J. Jap. Bot. 11: 499, 1935 (
* Targionia indica Udar et A.Gupta, Geophytology 13 (1): 83, 1983 (
by D.G.Long
*** Wiesnerella Schiffn., Österr. Bot. Z. 46 (3): 86, 1896 (
*** Wiesnerella denudata (Mitt.) Steph., Bull. Herb. Boissier 7 (5): 382 (154), 1899 (
* Wiesnerella fasciaria C.Gao et K.C.Chang, Acta Bot. Yunnan. 3 (4): 391, 1981 (
by D.G. Long
*** Neohodgsonia Perss., Bot. Not. 107 (1): 40, 1954 (
*** Neohodgsonia mirabilis (Perss.) Perss., Bot. Not. 107 (1): 40, 1954 (
by D.G. Long
*** Monocarpus D.J.Carr, Austral. J. Bot. 4 (2): 176, 1956 (
*** Monocarpus sphaerocarpus D.J.Carr, Austral. J. Bot. 4 (2): 176, 1956 (
by J. G. Segarra-Moragues and F. Puche
*** Austroriella Cargill et J.Milne, Polish Bot. J. 58 (1): 72, 2013 (
*** Austroriella salta J.Milne et Cargill, Polish Bot. J. 58 (1): 72, 2013 (
*** Riella Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 11, 1852 (
*** subg. Riella
*** Riella alatospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 317, 1937 (
*** Riella americana M.Howe et Underw., Bull. Torrey Bot. Club 30 (4): 218, 1903 (
* Riella battandieri Trab., Rev. Bryol. 13 (3): 35, 1886 (
*** Riella bialata Trab., Rev. Bryol. 35 (4): 96, 1908 (
*** Riella capensis Cavers, Rev. Bryol. 30 (5): 81, 1903 (
*** Riella choconensis Hässel, Symp. Biol. Hung. 35: 341, 1987 (
* Riella cyrenaica Maire, Bull. Soc. Hist. Nat. Afrique N. 30 (5): 312, 1939 (
*** Riella echinospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 321, 1937 (
* Riella gallica Balansa ex Trab., Rev. Gén. Bot. 3 (35): 450, 1891 (
*** Riella halophila Banwell, Trans. Brit. Bryol. Soc. 1 (5): 475, 1951 (
*** Riella helicophylla (Bory et Mont.) Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852 (
* Riella helicophylla var. macrocarpa P.Allorge, Sched. Br. Iber. (ser. 2): 4, 1929 (
* Riella indica Steph. ex Kashyap, J. Bombay Nat. Hist. Soc. 25 (2): 279, 1917 (
*** Riella notarisii (Mont.) Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852 (
** Riella numidica Trab., Bull. Soc. Hist. Nat. Afrique N. 25 (9): 391, 1934 [1935] (
*** Riella pampae Hässel, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 5 (9): 207, 1979 (
*** Riella parisii Gottsche, Hepat. Eur., Leberm. 38-39: no. 375, 1867 (
*** Riella purpureospora Wigglesw., J. Linn. Soc., Bot. 51 (339): 312, 1937 (
* Riella reuteri Mont., Ann. Sci. Nat. Bot. (sér. 3) 18: 12, 1852 (
* Riella sersuensis Trab., Bull. Soc. Hist. Nat. Afrique N. 25 (9): 392, 1934 [1935] (
** Riella spiculata J.Taylor, Kew Bull. 9 (1): 45, 1954 (
*** Riella trigonospora Segarra et Puche, S. African J. Bot. 94: 175, 2014 (
*** Riella undulata Hässel, Symp. Biol. Hung. 35: 341, 1987 (
*** subg. Trabutiella Porsild, Bot. Tidsskr. 24 (3): 327, 1902 (
*** Riella affinis M.Howe et Underw., Bull. Torrey Bot. Club 30 (4): 221, 1903 (
*** Riella cossoniana Trab., Atlas fl. Alger 1: 6, 1886 (
*** Riella echinata (Müll.Frib.) Segarra, Puche et Sabovlj., Phytotaxa 159 (3): 165, 2014 (
*** Riella gamundiae Hässel, Rev. Bryol. Lichénol. 38 (3/4): 580, 1972 [1973] (
*** Riella heliospora Segarra, Puche et Sabovlj., Syst. Bot. 37 (2): 315, 2012 (
*** Riella mediterranea Segarra, Puche, Sabovlj., M.Infante et Heras, Phytotaxa 159 (3): 170, 2014 (
by D. Long
*** Geothallus Campb., Bot. Gaz. 21 (1): 13, 1896 (
*** Geothallus tuberosus Campb., Bot. Gaz. 21 (1): 13, 1896 (
*** Sphaerocarpos Boehm., Def. gen. pl., ed. 3: 501, 1760 (
** subg. Austrosphaerocarpos R.M.Schust., Hepat. Anthocerotae N. Amer. 5: 813, 1992 (
*** Sphaerocarpos stipitatus Bisch. ex Lindenb., Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 18 (1): 504i, 1836 [1837] (
** subg. Sphaerocarpos
*** Sphaerocarpos michelii Bellardi, App. fl. pedem.: 52, 1792 (
** Sphaerocarpos texanus Austin, Bull. Torrey Bot. Club 6 (30): 158, 1877 (
Incertae sedis
*** Sphaerocarpos cristatus M.Howe, Mem. Torrey Bot. Club 7: 66, 1899 (
*** Sphaerocarpos donnellii Austin, Bull. Torrey Bot. Club 6 (30): 157, 1877 (
*** Sphaerocarpos drewiae Wigglesw., Univ. Calif. Publ. Bot. 16 (3): 129, 1929 (
*** Sphaerocarpos europaeus Lorb., Jahrb. Wiss. Bot. 80: 665, 1934 (
*** Sphaerocarpos hians Haynes, Bull. Torrey Bot. Club 37 (5): 225, 1910 (
** Sphaerocarpos muccilloi E.Vianna, Lindbergia 7 (1): 58, 1981 (
The following taxa in unsupported genera are all poorly understood. We list them here rather than making new combinations for names we do not know the status of.
Acrostolia Dumort., Recueil Observ. Jungerm.: 26, 1835 (
* Acrostolia alata (Gottsche et Rabenh.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (
* Acrostolia brevifolia (Gottsche et Rabenh.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 431, 1877 (
Aphanolejeunea A.Evans, Bull. Torrey Bot. Club 38 (6): 272, 1911 (
* Aphanolejeunea lancifera R.M.Schust., Phytologia 45 (5): 434, 1980 (
* Aphanolejeunea minima Tixier, Ann. Fac. Sci. Yaoundé 20: 7, 1975 (
Aspiromitus Steph., Sp. Hepat. (Stephani) 5: 957, 1916 (
* Aspiromitus asper Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 402, 1955 (
* Aspiromitus bullosus Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 403, 1955 (
* Aspiromitus crenatifrons Steph., Sp. Hepat. (Stephani) 5: 968, 1916 (
* Aspiromitus lobatus Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 405, 1955 (
* Aspiromitus squamulosus Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 407, 1955 (
Crossotolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
* Crossotolejeunea curvifolia Steph., Hedwigia 35 (3): 75, 1896 (
Eulejeunea Steph., Hedwigia 27 (2): 60, 1888 (
* Eulejeunea setulosa Steph., Sp. Hepat. (Stephani) 6: 421, 1923 (
* Eulejeunea subpililoba Steph., Sp. Hepat. (Stephani) 6: 420, 1923 (
Euosmolejeunea (Spruce) Steph., Hedwigia 28 (3): 170, 1889 (
* Euosmolejeunea parvistipula (Lindenb. et Gottsche) Steph., Hedwigia 29 (1): 80, 1890 (
* Euosmolejeunea tenerrima (Nees) Steph., Sp. Hepat. (Stephani) 5: 589, 1914 (
Fimbraria Nees, Horae Phys. Berol.: 44, 1820 (
* Fimbraria gigantea Steph., Bull. Herb. Boissier 7 (2): 93 (106), 1899 (
* Fimbraria incrassata Steph., Bull. Herb. Boissier 7 (2): 87 (100), 1899 (
* Fimbraria kamerunensis Steph., Sp. Hepat. (Stephani) 6: 14, 1917 (
* Fimbraria pirottae Gola, Ann. Bot. (Rome) 13 (1): 64, 1914 (
Hygrolejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 124, 1893 (
* Hygrolejeunea cubensis Steph., Sp. Hepat. (Stephani) 5: 533, 1914 (
* Hygrolejeunea harpaphylla Steph., Sp. Hepat. (Stephani) 5: 552, 1914 (
* Hygrolejeunea pacifica Steph., Sp. Hepat. (Stephani) 6: 411, 1923 (
* Hygrolejeunea parvicalycina Steph., Hedwigia 35 (3): 103, 1896 (
* Hygrolejeunea parvistipula Steph., Sp. Hepat. (Stephani) 5: 568, 1914 (
* Hygrolejeunea patellirostris Steph., Hedwigia 35 (3): 103, 1896 (
* Hygrolejeunea staudtiana Steph., Sp. Hepat. (Stephani) 5: 528, 1914 (
Hypenantron Corda, Gen. hepat.: 648, 1829 (
* Hypenantron brachypus Steph. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 107, 1919 (
* Hypenantron brasiliense Steph. ex Lamothe, Rech. Anat. Taxinom. Gamét. Marchantiales: 104, 1919 (
Jamesoniella (Spruce) Carrington, Cat. Brit. Moss. Hepat.: 25, 1881 (
* Jamesoniella convoluta Steph., Sp. Hepat. (Stephani) 6: 433, 1924 (
Kingiolejeunea H.Rob., Bryologist 70 (1): 53, 1967 (
* Kingiolejeunea guayanensis H.Rob., Bol. Soc. Venez. Ci. Nat. 32 (132/133): 259, 1976 (
Leptocolea (Spruce) A.Evans, Bull. Torrey Bot. Club 38 (6): 261, 1911 (
* Leptocolea sumatrana Herzog, Ann. Bryol. 5: 96, 1932 (
Mastigobryum (Nees) Lindenb. et Gottsche, Syn. Hepat. 2: 214, 1845 (
* Mastigobryum aberrans Steph., Sp. Hepat. (Stephani) 6: 486, 1924 (
* Mastigobryum asperum Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 121, 1914 (
* Mastigobryum deningeri Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921 (
* Mastigobryum karstenii Steph., Bull. Herb. Boissier (sér. 2) 8 (12): 952 (502), 1908 (
* Mastigobryum ledermannii Steph., Sp. Hepat. (Stephani) 6: 486, 1924 (
* Mastigobryum londbergii Steph., Sp. Hepat. (Stephani) 6: 487, 1924 (
* Mastigobryum longifolium Steph., Sp. Hepat. (Stephani) 6: 487, 1924 (
* Mastigobryum minutitextum Steph., Sp. Hepat. (Stephani) 6: 487, 1924 (
* Mastigobryum multidens Steph., Sp. Hepat. (Stephani) 6: 487, 1924 (
* Mastigobryum muscicola Steph., Sp. Hepat. (Stephani) 6: 486, 1924 (
* Mastigobryum nigricans Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921 (
* Mastigobryum nipuranum Steph., Sp. Hepat. (Stephani) 6: 485, 1924 (
* Mastigobryum palmicola Steph., Sp. Hepat. (Stephani) 6: 488, 1924 (
* Mastigobryum rajanum Herzog, Mitt. Inst. Allg. Bot. Hamburg 7 (3): 190, 1931 (
* Mastigobryum ribehanum Steph., Sp. Hepat. (Stephani) 6: 488, 1924 (
* Mastigobryum ruficaule Beauverd, Sp. Hepat. (Stephani) 6: 485, 1924 (
* Mastigobryum schraderbergii Steph., Sp. Hepat. (Stephani) 6: 489, 1924 (
* Mastigobryum squamulistipum Steph., Sp. Hepat. (Stephani) 6: 480, 1924 (
* Mastigobryum subhyalinum Steph., Sp. Hepat. (Stephani) 6: 482, 1924 (
* Mastigobryum venezuelanum Molk., Syn. hepat. jav.: 104, 1856 [1857] (
* Mastigobryum vermiculare Herzog, Hedwigia 66 (6): 339, 1926 (
Nemoursia Mérat, Ann. Agric. Franç. (ser. 4) 2 (7): 10, 1840 (
* Nemoursia tuberculata Mérat, Ann. Agric. Franç. (ser. 4) 2 (7): 10, 1840 (
Physocolea (Spruce) Steph., Sp. Hepat. (Stephani) 5: 863, 1916 (
* Physocolea tambillensis (Loitl.) Steph., Sp. Hepat. (Stephani) 5: 885, 1916 (
Plectocolea (Mitt.) Mitt., Fl. vit.: 405, 1871 [1873] (
* Plectocolea subamoena S.Winkl., Rev. Bryol. Lichénol. 42 (3): 821, 1976 (
Polyotus Gottsche, Syn. Hepat. 2: 244, 1845 (
* Polyotus peckianus Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 224, 1869 (
Schisma Dumort., Commentat. Bot. (Dumortier): 114, 1822 (
* Schisma orizabense (Gottsche) Steph., Sp. Hepat. (Stephani) 4: 19, 1909 (
* Schisma uleanum Steph., Hedwigia 44 (4): 225, 1905 (
Strepsilejeunea (Spruce) Schiffn., Hepat. (Engl.-Prantl): 127, 1893 (
* Strepsilejeunea apollinea (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 291, 1913 (
* Strepsilejeunea durelii Schiffn., Österr. Bot. Z. 49 (6): 206, 1899 (
* Strepsilejeunea hamatifolia Steph., Sp. Hepat. (Stephani) 6: 396, 1923 (
* Strepsilejeunea lanceolata (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 283, 1913 (
* Strepsilejeunea muscicola Herzog, Hedwigia 74 (2): 96, 1934 (
* Strepsilejeunea novae-guineae Steph., Sp. Hepat. (Stephani) 6: 397, 1923 (
* Strepsilejeunea obtusistipula Steph., Biblioth. Bot. 87 (2): 258, 1916 (
* Strepsilejeunea papulifolia Steph., Biblioth. Bot. 87 (2): 259, 1916 (
* Strepsilejeunea pectiniformis (Gottsche) Steph., Sp. Hepat. (Stephani) 5: 285, 1913 (
* Strepsilejeunea renistipula Steph., Sp. Hepat. (Stephani) 5: 289, 1913 (
* Strepsilejeunea vatovae Gerola, Lav. Bot. Ist. Bot. Univ. Padova 12: 479, 1947 (
Thysanolejeunea (Spruce) Steph., Hedwigia 31 (1): 20, 1892 (
* Thysanolejeunea africana Sim, Trans. Roy. Soc. South Africa 15 (1): 50, 1926 (
Trachylejeunea (Spruce) Steph., Hedwigia 28 (4): 262, 1889 (
* Trachylejeunea conifera Steph., Sp. Hepat. (Stephani) 5: 302, 1913 (
* Trachylejeunea cristuliflora Steph., Hedwigia 35 (3): 137, 1896 (
* Trachylejeunea englishii Steph., Bull. Misc. Inform. Kew 1899 (151/152): 126, 1899 (
* Trachylejeunea jamaicensis Pearson, Ann. Bryol. 4: 98, 1931 (
* Trachylejeunea kusaiensis Inoue et H.A.Mill., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 8 (2): 147, 1965 (
The Early Land Plants Today project has been a community driven and collaborative effort that has involved numerous dedicated individuals, many institutions, and funding agencies that have helped provide partial funding. The high quality of the data and information could not have been achieved without rapid responses to our requests for information regarding publications, PDFs or other forms of bibliographic data from committed individuals. These include Afroz Alam, Dulip Daniels, Rajendra Lavate, Shuvadeep Majumdar, Alexey Potemkin, Ana Séneca, Mafalda Silva, Devendra K. Singh, Sushil K. Singh, Louis Thouvenot, Praveen Verma, and Gerhard Winter. We wish to especially thank the many individuals who have provided a wealth of logistical support that has been broad ranging from access to data, assistance with translation, access to unpublished molecular results and information, access to type material or herbaria support, library support, advice on Latin usage as well as many other forms of support. These have included Silvia Aranda, Riccardo Baldini, Fred Barrie, Laura Campos, Tong Cao, Xia-Fang Cheng, Patricia Eckel, Len Ellis, Lars Hedenäs, Christine Giannoni, Qiong He, Nijole Kalinauskaitė, Lucia Kawasaki, Peter de Lange, Itambi Malombe, John McNeill, David Meagher, Chiara Nepi, Michelle Price, Qiong Qui, Steven Rogstad, Cecilia Sérgio, Blanka Shaw, Jon Shaw, Akiko Shinya, Lei Shu, Sorin Ştefănuţ, Alain Vanderpoorten, Anna Vilnet, Yu-Mei Wei, Ying Yu, and Lan-Ping Zhou. We gratefully acknowledge the Biodiversity Heritage Library as a resource and the Conservatoire et Jardin botaniques de la Ville de Genève for providing access to the Franz Stephani database and specimens and Missouri Botanical Garden for access to the TROPICOS database. Throughout the years, the project has been generously supported in part by the Global Biological Information Facility (GBIF) Seed Money Award No.2007-41 and supported in part by funding from the Biodiversity Synthesis Center of the Encyclopedia of Life. Funding from the National Science Foundation (DBI-0749762, DBI 1057418, DBI 1458300, DEB-1145898, EF-0531730, EF-1115116) has also partially supported aspects of the ELPT project. We also wish to express our gratitude to the editors and publication team of Phytotaxa that published the important series of notes that helped lead to the current publication and to the tireless efforts of the editorial and production team of PhytoKeys that led to the current publication.
Anthoceros includes Aspiromitus, but some taxa have neither been transferred nor synonymized. They are listed in the “Names in genera not currently accepted” section below.
Anthoceros agrestis is conserved against the earlier Anthoceros nagasakiensis Steph. 1916. Anthoceros cristatus Steph. 1916 is also an earlier synonym and a proposal to conserve Anthoceros agrestis against it is in progress.
Anthoceros angustifolius (type from Europe) has neither been recognized in any recent treatment nor synonymized. The name may have priority once its identity is determined.
Anthoceros ferdinandi-muelleri is similar to Anthoceros punctatus, but Anthoceros still needs to be revised for Australia.
Anthoceros helmsii was treated as conspecific with Anthoceros muscoides by
Anthoceros javanicoides belongs to the Anthoceros punctatus species complex (
Anthoceros punctatus is a species complex in need of a revision.
Anthoceros aethyopicus is probably a Phaeoceros species (
Anthoceros brunnthaleri was annotated by Hasegawa in 1993 as probably conspecific with Phymatoceros bulbiculosus but the synonymy seems not to have been published.
Anthoceros floribundus type specimen in G was annotated by Hasegawa in 1993 as conspecific with Anthoceros parvifrons which is a Phaeoceros species.
Anthoceros mildbraedii is probably conspecific with Phaeoceros carolinianus (
Anthoceros parvifrons is annotated as a Phaeoceros species by Hasegawa in 1993 but the new combination has never been published.
Anthoceros pseudocostus is a Phaeoceros species of uncertain status (
Anthoceros rossoi is a Phaeoceros species of uncertain status (
Dendroceros adglutinatus is conspecific with Dendroceros crispus in
Dendroceros cucullatus was treated as conspecific with Dendroceros difficilis by
Dendroceros exalatus was reported from the Moluccas in
Dendroceros reticulus was mentioned by
Megaceros flavens and Megaceros jamesonii are probably conspecific with Nothoceros vincentianus (
Mesoceros was described from a collection that contains a mixture of both Anthoceros and Phaeoceros material.
Phaeoceros dendroceroides may belong to Phaeomegaceros
Phaeoceros microsporus may be conspecific with Phaeoceros mohrii in
Phaeoceros striatisporus was nested within Phaeoceros in
Syzygiella includes Jamesoniella (cf.
Syzygiella securifolia is a sister taxon to Syzygiella subintegerrima (
Anastrophyllum ellipticum is only tentatively placed in the genus following Mamontov & Vilnet (2013).
Anastrophyllum lignicola is possibly conspecific with Anastrophyllum ellipticum and its generic placement is uncertain.
Barbilophozia sudetica was transferred from Lophozia by
Gymnocolea inflata subsp. acutiloba is a problematic taxon that sometimes has been treated as conspecific with Gymnocolea inflata, sometimes as a separate species.
Plicanthus birmensis seems to be closely related to Tetralophozia (
Plicanthus giganteus is often treated as a subspecies of Plicanthus hirtellus (e.g.
Schizophyllopsis was originally published as Schizophyllum (R.M.Schust.) Váňa et L.Söderstr. (
Cephalozia neesiana may be conspecific with Cephalozia hamatiloba (
Cephalozia hians is probably an Isotachis or Lophocolea species.
Cephalozia indica is a Lophocolea species and probably conspecific with “Cephalozia” kodaikanalensis.
Cephalozia kodaikanalensis is a Lophocolea species, possibly conspecific with “Cephalozia” indica.
Cephalozia parvifolia is a Cephaloziella species although the type specimen has not been found.
Cephalozia tricuspidata is a Lepidozia species (
Fuscocephaloziopsis lunulifolia is a species complex also including Fuscocephaloziopsis affinis and Fuscocephaloziopsis schusteri.
Odontoschisma obcordatum is to be excluded from Odontoschisma but its identity remains unclear as the type is not found. Based on the original description this species might belong to Alobiellopsis (Gradstein & Ilkiu-Borges 2014: 81).
Cephaloziella is a large genus and the subgeneric division is unclear. The traditional subdivisions used here do not necessary agree with recent phylogenetic studies.
The correct name for the variety was discussed by
Cephaloziella villaumei is possibly conspecific with Cephaloziella anthelioides.
Cephaloziella meghalayensis is probably conspecific with Cephaloziella acanthophora or Cephaloziella microphylla.
Cephaloziella flexuosa may be a form of Plicanthus birmensis or a juvenile form of Tetralophozia filiformis.
Cephaloziella pellucida is probably not a Cephaloziella, but the type specimen has not been found.
Cephaloziella pygmaea is possibly conspecific with Cephaloziella granatensis (
Cephaloziella secundifolia could not be studied by
Lophozia has many nomenclatural and taxonomic problems. This treatment follows
Lophozia jamaicensis is probably not a Lophozia, but the type specimen is missing in Nees’s herbarium (
Lophozia pallida may be conspecific with Lophozia guttulata (
Lophozia subapiculata may be a form of Lophozia guttulata, but
Lophozia udarii has recently been described using the generic concept of Lophozia in the wide sense (as e.g.
Lophozia ventricosa is very variable and many segregates at various subspecific levels have been described. The species complex also includes at least Lophozia austrosibirica, Lophozia schusteriana and Lophozia wenzelii.
Lophozia wenzelii is closely related to Lophozia ventricosa according to
Lophozia serpens is probably conspecific with Schistochilopsis incisa.
Lophoziopsis longidens var. arctica was treated as a form of Lophoziopsis pellucida by
Lophoziopsis propagulifera belongs to the Lophoziopsis excisa species complex. It is probably only a modification (cf. discussion by
Diplophyllum obtusifolium is a species complex also including Diplophyllum androgynum, Diplophyllum angustifolium, Diplophyllum gemmiparum, Diplophyllum incrassatum and Diplophyllum novum.
Scapania degenii is conspecific with Scapania brevicaulis in
Scapania glaucoviridis was treated as conspecific with Scapania parvitexta by
Scapania parvidens was treated as conspecific with Scapania parvitexta by
Scapania scapanioides is possibly conspecific with Scapania glaucocephala (
Schistochilopsis nakanishii is possibly conspecific with Schistochilopsis incisa.
Schistochilopsis opacifolia is sometimes treated as a subspecies of Schistochilopsis incisa (e.g.
Acrobolbus bispinosus is a Plagiochila species (
Isotachis multiceps is sometimes treated as a Hypoisotachis, but recent molecular studies (
Calypogeia fissa contains two well separated taxa (
Calypogeia lechleri was treated as conspecific with Calypogeia leptoloma by
Calypogeia muelleriana contains two well separated taxa (
Calypogeia sphagnicola is a species complex including at least three genetically and morphologically well separated taxa (
Calypogeia amazonica was treated as conspecific with Calypogeia miquelii in
Nardia kamtschatica may be conspecific with Nardia assamica (
Jackiella unica was identified as a Colura species before the B herbarium burned (
Jungermannia was used for almost all leafy liverworts from
Jungermannia erectii was described on the basis of sterile material and it is not certain if it belongs in Jungermannia or Solenostoma (
Jungermannia was used for almost all leafy hepatics from
Jungermannia brasiliensis is a taxon of uncertain status.
Jungermannia chinensis may be conspecific with Cephalozia connivens or Cephalozia bicuspidata.
Jungermannia incerta is a doubtful taxon (
Jungermannia lescuriana is a Cephaloziella species.
Jungermannia longiretis is an Isotachis species.
Jungermannia mastigophora is probably a Mastigophora species.
Jungermannia michelii is a Fossombronia species.
Jungermannia quadridigitata is conspecific with Kurzia pauciflora in
Jungermannia stereocaulis is a doubtful taxon.
Jungermannia sullivantiana is a Cephaloziella species.
Jungermannia tenuis is a Lejeunea species.
Saccogyna tridens is a Bazzania species, but the affinity is unclear and the type specimen was burned in B (
Solenostoma subdivisions are following
Solenostoma rupicola is conspecific with Solenostoma rosulans in
Solenostoma appalachianum is placed in synonymy with Solenostoma pyriflorum by
Solenostoma appressifolium seems to be a species complex (
Solenostoma novaezelandiae is possibly conspecific with Solenostoma orbiculatum (
Solenostoma pyriflorum is a species complex (
Solenostoma pyriflorum var. minutissimum was treated as a separate species, Solenostoma rishiriense Amakawa by
Solenostoma totopapillosum is possibly conspecific with Solenostoma inundatum (
Solenostoma ventroversum is possibly conspecific with Solenostoma appressifolium.
Herbertus includes Schisma, but a few taxa have neither been transferred nor synonymized. They are listed in the “Names in genera not currently accepted” section below.
Herbertus delavayi is a species complex that requires further studies (
Herbertus juniperoideus is here treated with two subspecies, but they may deserve species rank instead.
Herbertus subrotundus does not belong in Herbertus (
Lepicolea rara is possibly a species complex and warrants further investigation (
Acromastigum is here divided into two subgenera which are supported by recent molecular studies (
Acromastigum echinatum may be conspecific with Acromastigum inaequilaterum since all specimens identified by
Bazzania can not be subdivided based on recent molecular studies, so infrageneric taxa are not used here. The genus includes Mastigobryum, but several taxa have neither been transferred nor synonymized. They are listed in the “Names in genera not currently accepted” section below.
Bazzania asymmetrica is conspecific with Bazzania macgregorii in
Bazzania corbieri may be conspecific with Bazzania crenata (
Bazzania crenata from Europe has neither been recognized in any recent treatment nor synonymized. The name may have priority once the identity is established.
Bazzania decrescens is a species complex also including also Bazzania comorensis, Bazzania curvidens and Bazzania mascarena (
Bazzania distans is probably conspecific with Bazzania loricata (
Bazzania lehmanniana is possibly conspecific with Bazzania arcuata or Bazzania longa (
Bazzania serpentina is possibly conspecific with Bazzania bidens (
Bazzania sikkimensis is possibly conspecific with Bazzania sumbavensis (
Bazzania sumbavensis is possibly conspecific with Bazzania japonica (
Bazzania wallichiana seems to be a species complex also including Bazzania francana, Bazzania intermedia and Bazzania marginata (
Bazzania wattsiana is possibly conspecific with Bazzania tridens (
Mastigopelma may be better placed in Bazzania, but it is retained here until further studied.
Kurzia cucullifolia is of doubtful status (
Lepidozia seems to be a monophyletic genus, but its infrataxonomy still has to be tested using molecular data.
Lepidozia brasiliensis is conspecific with Kurzia brasiliensis in
Lepidozia massartiana has not been studied recently.
Lepidozia minor may be conspecific with a known taxon. Lepidozia minor sensu
Lepidozia pallida is possibly Lepidozia chordulifera since all specimens cited by
Lepidozia paucifolia is conspecific with Lepidozia fauriana in
Lepidozia hexiloba is possibly an Arachniopsis or Telaranea species (
Lepidozia parvistipa is possibly conspecific with Ceramanus clatritexta.
Micropterygium trachyphyllum was split into several varieties by
Telaranea is a heterogeneous genus and several segregates are warranted. Additional studies are needed to clarify the relationships of the species retained here (
Zoopsis ciliata was not studied by
Chiloscyphus pallescens var. fragilis is a problematic taxon sometimes treated as a species, sometimes as conspecific with Chiloscyphus pallescens, sometimes as conspecific with Chiloscyphus polyanthos.
Chiloscyphus polyanthos var. rivularis is a problematic taxon sometimes treated as a species, sometimes as conspecific with Chiloscyphus polyanthos.
Chiloscyphus is here treated in a very narrow sense (cf.
Chiloscyphus bifidus is possibly conspecific with Heteroscyphus aselliformis (
Chiloscyphus durus is conspecific with Leptoscyphus expansus in
Clasmatocolea is here treated without subdivisions although
Cryptolophocolea levieri is possibly conspecific with Cryptolophocolea ciliolata (
Cryptolophocolea lilliena is possibly conspecific with Cryptolophocolea martiana (
Cryptolophocolea massalongoana is possibly conspecific with Cryptolophocolea costata (
Cryptolophocolea stephanii is doubtfully distinct from Cryptolophocolea ciliolata (
Cryptolophocolea thermarum is possibly conspecific with Cryptolophocolea ciliolata (
Cryptolophocolea whittieriana is possibly conspecific with Cryptolophocolea helmsiana (
Heteroscyphus caesius is possibly conspecific with Heteroscyphus splendens (
Heteroscyphus caledonicus is possibly conspecific with Heteroscyphus amboinensis
The subdivision of Leptoscyphus follows
Leptoscyphus difficilis is conspecific with Leptoscyphus expansus in
Leptoscyphus huidobroanus is conspecific with Leptoscyphus expansus in
Lophocolea bidentata is a species complex as discussed by
Lophocolea coadunata belongs to the Lophocolea bidentata species complex with complicated nomenclature (
Lophocolea microstipula is possibly conspecific with Lophocolea sabuletorum (
Lophocolea mollis is possibly conspecific with Lophocolea difformis (
Lophocolea morobeana is conspecific with Chiloscyphus kurzii in
Lophocolea rectangulata may be a Cryptolophocolea species.
Lophocolea semiteres is a species complex also including Lophocolea platensis and Lophocolea undulata.
Pachyglossa exilis may be a Chiloscyphus species.
Mastigophora includes (Mastigophora appendiculata, Mastigophora guineensis, Mastigophora pyramidana and Mastigophora valida) which
Plagiochila arnelliana may be conspecific with Plagiochila bifaria (
Plagiochila fragilis may be conspecific with Plagiochila bifaria (
Plagiochila integrilobula is possibly conspecific with Plagiochila blepharophora (
Plagiochila kuhliana is possibly conspecific with Plagiochila sciophila (
Plagiochila stephanii is possibly conspecific with Plagiochila blepharophora (
Plagiochila fragmentissima may be conspecific with Plagiochila gregaria (M.L. So and D. Glenny, pers. comm.).
Plagiochila cuneata is conspecific with Plagiochila bursata in
Plagiochila steyermarkii is conspecific with Plagiochila aerea in Inoue in
Plagiochila aspera is conspecific with Plagiochila trigona Steph. (=Plagiochila metcalfii) in
Plagiochila heterostipa is possibly conspecific with Plagiochila corymbulosa (
Plagiochila loloënsis is probably an entire-leaved form of Plagiochila moenkemeyeri (
Plagiochila moenkemeyeri is possibly conspecific with Plagiochila winteri (
Plagiochila owaihiensis was considered conspecific with Plagiochila remyana by
Plagiochila aliena is conspecific with Plagiochila ludoviciana in
Plagiochila borneensis is a doubtful taxon. No specimen could be found by
Plagiochila distans is a doubtful taxon. The type specimen could not be found (
Plagiochila exilis is a doubtful taxon. The type specimen could not be found (
Plagiochila glauca is a doubtful taxon.
Plagiochila karstenii is possibly conspecific with Plagiochila bantamensis (
Plagiochila lecontei is allied to and perhaps conspecific with Plagiochila ledieui and Plagiochila multiflora (
Plagiochila ledermanniana is a doubtful taxon. The type specimen was burned in B (
Plagiochila ledieui is allied to and perhaps conspecific with Plagiochila lecontei and Plagiochila multiflora (
Plagiochila multiflora is allied to and perhaps conspecific with Plagiochila lecontei and Plagiochila ledieui (
Plagiochila schraderbergii is a doubtful taxon. The type specimen was burned in B (
Plagiochila scotica was described from Europe, but it has neither been recognized in any recent European study nor synonymized.
Plagiochila vetustisilva is a doubtful taxon. The type specimen was burned in B (
Trichocolea floccosa is treated as a variety of Trichocolea flaccida (=Leiomitra flaccida by Schultze-Motel & Menzel (1987).
Trichocolea sprucei is suggested to be a variety of Trichocolea flaccida (=Leiomitra flaccida) in Schultze-Motel & Menzel (1987)
Schistochila baileyana is possibly conspecific with Schistochila beccariana (D. Meagher, pers. comm.).
Frullania trinervis is possibly a species complex (
Frullania claviloba is possibly conspecific with Frullania gracilis (
Frullania cordistipula is a species complex also including Frullania serrata.
Frullania crenatiloba is conspecific with Frullania apiculata in
Frullania multilacera is possibly conspecific with Frullania vaga.
Frullania van-zantenii is possibly conspecific with Frullania serrata.
Frullania brunea belongs to the Frullania lindenbergii complex (
Frullania tricarinata is possibly conspecific with Frullania hypoleuca (
Frullania contracta is conspecific with Frullania squarrosa in
Frullania cornuta is closely related to the very variable Frullania ornithocephala.
Frullania flexuosa is possibly conspecific with Frullania rubella.
Frullania giraldiana belongs to the Frullania nepalensis complex (
Frullania hebridensis belongs to the Frullania ericoides species complex (
Frullania longistyla is closely related to Frullania brevicalycina (
Frullania ludoviciae seems to be close to Frullania pusilla (
Frullania montana is possibly conspecific with Frullania reflexistipula.
Frullania nepalensis is a species complex (
Frullania subpedicellata is possibly conspecific with Frullania pedicellata.
Frullania baladina is a species complex also including Frullania fulfordiae.
Frullania fugax is a species complex possibly including Frullania baileyana, Frullania belmorensis, Frullania cataractarum, Frullania media and Frullania subtropica (
Frullania solanderiana is morphologically similar to Frullania pentapleura and possibly conspecific.
Frullania tjibodensis is possibly conspecific with Frullania campanulata (
Frullania eboracensis is a species complex also including Frullania appalachiana, Frullania parvistipula and Frullania virginica.
Frullania ericoides is enormously variable (
Frullania astrolabea is possibly conspecific with Frullania scandens (
Frullania intermedia is a species complex (
Frullania intermedia var. non-apiculata was reduced to a synonym of subsp. intermedia by
Frullania brotheri is probably just a high-elevation form of Frullania nodulosa (
Frullania brasiliensis is a species complex with controversial synonymy (
Frullania breuteliana is possibly conspecific with Frullania beyrichiana.
Frullania compacta is conspecific with Frullania brasiliensis in
Frullania granatensis is possibly conspecific with Frullania beyrichiana, but the type specimen was lost in B (
Frullania borbonica is possibly conspecific with Frullania apicalis (
Frullania schimperi belongs to a species complex also including Frullania onraedtii, Frullania capensis, Frullania apicalis, Frullania imerinensis and Frullania meyeniana (
Frullania chevalieri is a species complex (
Frullania junghuhniana is a species complex (
Frullania rostrata is a species complex with several lineages, some of which have been recently described (e.g.
Frullania acicularis was shown to be a species separate from Frullania tamarisci (
Frullania appendiculata was shown by
Frullania asagrayana consists of two geographically separated clades (
Frullania calcarifera was shown to be a species separate from Frullania tamarisci by
Frullania moniliata is a species complex (
Frullania subarctica was shown to be a species separate from the Frullania tamarisci complex (
Frullania affinis is possibly conspecific with Frullania ecklonii (
Jubula hutchinsiae has subspecies that may be better recognized at the species level.
Jubula hutchinsiae subsp. bogotensis is closely related to subsp. pennsylvanica (
Jubula hutchinsiae subsp. javanica includes a cryptic taxon from China (
The treatment of Ceratolejeunea follows
Ceratolejeunea kuerschneri is very similar to Ceratolejeunea papuliflora and may be conspecific with it.
Ceratolejeunea sinensis may be conspecific with Drepanolejeunea erecta (
Otigoniolejeunea crenulata is a doubtful taxon. The type specimen was burned in B and it is unclear where the taxon belongs (
Otigoniolejeunea ledermannii is a doubtful taxon. The type specimen was burned in B and it is unclear where the taxon belongs (
Aureolejeunea is nested in Cheilolejeunea (
Cheilolejeunea is here organized into subgenera that are maintained for practical purposes. They may not be natural lineages. An improved infrageneric classification based on molecular analysis is in preparation (
Cheilolejeunea fukiensis may be conspecific with Lejeunea flava (
Cheilolejeunea laurentii is not a Cheilolejeunea species (
Cheilolejeunea longiflora is possibly conspecific with Cheilolejeunea trifaria (
Cheilolejeunea micholitzii is conspecific with Cheilolejeunea longidens in
Cheilolejeunea ulugurica is possibly conspecific with Cheilolejeunea chenii (
Cheilolejeunea udarii is possibly conspecific with Cheilolejeunea krakakammae (
Cheilolejeunea heteroclada was considered conspecific with Cheilolejeunea aneogyna by X.-L. He (in
Omphalanthus is nested in Cheilolejeunea (
Cololejeunea is here organized into subgenera that are not supported by the molecular study of
Cololejeunea rotundilobula is possibly conspecific with Cololejeunea sigmoidea (
Cololejeunea subg. Chlorolejeunea forms a separate clade in
Cololejeunea subg. Chondriolejeunea is supported by
Subg. Cololejeunea includes subg. Aphanolejeunea in the phylogeny by
Cololejeunea frahmii is probably only a depauperate form of Cololejeunea elegans or Cololejeunea zenkeri.
Cololejeunea kahuziensis is probably only a depauperate form of Cololejeunea elegans or Cololejeunea zenkeri.
Cololejeunea drepanolejeuneoides may based on its description be close to Cololejeunea angustiflora and especially to Cololejeunea inflectens, forming a natural group.
Cololejeunea chamlongiana is possibly conspecific with Cololejeunea papuliflora.
Cololejeunea crenulata is possibly conspecific with Cololejeunea angustiflora judging from the protologue.
Cololejeunea dentilobula is possibly conspecific with Cololejeunea decliviloba judging from Stephani’s Icones (unpublished).
Cololejeunea effusa may be conspecific with Cololejeunea trichomanis.
Cololejeunea fleischeri is near to or conspecific with Cololejeunea bidentula.
Cololejeunea fredericii is possibly conspecific with Cololejeunea polyantha.
Cololejeunea kohkongensis may be conspecific with Cololejeunea plagiophylla.
Cololejeunea lobulilineata seems to be a depauperate form of some other species, maybe Cololejeunea obtusifolia.
Cololejeunea micronesica is possibly conspecific with Cololejeunea cookeifide
Cololejeunea ninguana is possibly conspecific with Cololejeunea decliviloba (
Cololejeunea oblongiperianthia is possibly conspecific with Cololejeunea trichomanis (
Cololejeunea pteroporum is possibly conspecific with Cololejeunea spathulifolia.
Cololejeunea spathulifolia is possibly conspecific with Cololejeunea obliqua (
Cololejeunea takamakae is possibly conspecific with Cololejeunea angustiflora.
Cololejeunea tuksapiana is possibly conspecific with Cololejeunea aequalis.
Cololejeunea brunelii is conspecific with Cololejeunea madothecoides in
Cololejeunea chittagongensis is probably conspecific with Cololejeunea schwabei.
Cololejeunea deslooveri is possibly conspecific with Cololejeunea cristata.
Cololejeunea dzumacensis is possibly conspecific with Cololejeunea lanciloba, but the type specimen has not been localized.
Cololejeunea epiphylla is closely related to Cololejeunea chittagongensis and Cololejeunea schwabei.
Cololejeunea hoabinhiana is possibly conspecific with Cololejeunea lanciloba.
Cololejeunea malaccensis may be conspecific with Cololejeunea stylosa.
Cololejeunea punctata is a doubtful taxon. The type specimen has been destroyed and the identity is unclear (
Cololejeunea tribracteata is possibly conspecific with Cololejeunea africana.
Cololejeunea uchimae is conspecific with Cololejeunea raduliloba in
Cololejeunea chuahiana is probably a Myriocoleopsis species (
Cololejeunea dauphinii is probably a Myriocoleopsis species (
Cololejeunea disciflora is probably a Myriocoleopsis species (
Cololejeunea amoena is possibly conspecific with Cololejeunea floccosa (
Cololejeunea bontocensis is possibly conspecific with Cololejeunea pseudostephanii.
Cololejeunea flavida may be conspecific with Cololejeunea peraffinis (
Cololejeunea polisiana may be conspecific with Cololejeunea peraffinis.
Cololejeunea conchifolia is probably a Myriocoleopsis species (
Diplasiolejeunea is here only provisionally accepted with subgenera following
Diplasiolejeunea guadalupensis is a poorly known taxon with some similarities to the Diplasiolejeunea replicata species complex (
Diplasiolejeunea heimii is possibly conspecific with Diplasiolejeunea replicata (
Diplasiolejeunea montecristensis is possibly conspecific with Diplasiolejeunea replicata.
Diplasiolejeunea rudolphiana and Diplasiolejeunea unidentata are genetically very close and may be conspecific (
Diplasiolejeunea glaziovii may be conspecific with Diplasiolejeunea cavifolia (
Diplasiolejeunea evansii is a doubtful taxon and the type specimen could not be found in PC (
Diplasiolejeunea integerrima is a juvenile Acrolejeunea species, but the type is too poor to allow identification (
Prionolejeunea corbisieri is a Cheilolejeunea species (
Prionolejeunea maculata is a Cyclolejeunea species (
Drepanolejeunea deslooveri was treated as conspecific with Drepanolejeunea hamatifolia by
Drepanolejeunea lancifolia resembles Drepanolejeunea bidens and Drepanolejeunea araucariae, but the type material has not been found (
Drepanolejeunea obliqua is possibly conspecific with Drepanolejeunea ternatensis (
Drepanolejeunea ruandensis is conspecific with Drepanolejeunea cultrella in
Drepanolejeunea tenera is possibly conspecific with Drepanolejeunea pentadactyla (
Drepanolejeunea thwaitesiana is a species complex also including Drepanolejeunea hampeana, Drepanolejeunea longicornua, Drepanolejeunea nymannii and Drepanolejeunea serricalyx.
Drepanolejeunea integerrima is a Leptolejeunea species (
Harpalejeunea acuta is possibly conspecific with Harpalejeunea cinchonae (
Harpalejeunea tenuicuspis is possibly conspecific with Harpalejeunea oxyphylla (
Harpalejeunea grossearmata is a Lejeunea species (
Harpalejeunea renneri is a Lejeunea species and most likely conspecific with something (
Harpalejeunea spruceana is a Lejeunea species (non Lejeunea spruceana C.Massal. 1885) (
Harpalejeunea vitrea is a Lejeunea species (non Lejeunea vitrea Nees, Lindenb. et Gottsche 1843) (
Lejeunea is a large genus and its infrageneric classification, which has not been studied in a rigorous manner, needs investigation although
Lejeunea adpressa from South America is very similar to the palaeotropical Lejeunea anisophylla. The two taxa probably represent a single pantropical species (possibly with two subspecies).
Lejeunea angulifolia may be conspecific with Lejeunea cocoes (
Lejeunea caroliniana was poorly described and the type specimen so poor that
Lejeunea flava is a species complex also including Lejeunea grossecristata.
Lejeunea lamacerina forms a North American and a Macaronesian/European clade and the separation of two subspecies are justified if not treating them at species level (
Lejeunea leratii is probably distinct from Lejeunea mimula (
Lejeunea obscura is possibly conspecific with Lejeunea aloba (
Lejeunea pallidissima is possibly conspecific with Lejeunea flavovirens (
Lejeunea paratropa is closely related to or perhaps conspecific with Lejeunea raddiana Lindenb. (
Lejeunea rara is the valid name for Lejeunea sinclairii
Lejeunea ravenelii was poorly described and the type specimen so poor that
Lejeunea stephaniana is doubtfully distinct from Lejeunea discreta (
Lejeunea emarginuliflora is a Lejeunea (subg. Heterolejeunea) species (
Lejeunea oerstediana is a poorly known taxon often placed in Euosmolejeunea (=Cheilolejeunea). We can not tell where it belongs until the type specimen is re-studied (
Lejeunea elegans is a Cheilolejeunea species (
Lejeunea hieronymii is a Cheilolejeunea species of uncertain status.
Lejeunea proboscidea is a Drepanolejeunea species, but
Lejeunea scabriflora was rejected from Dicranolejeunea by
Lejeunea trochantha is a Cheilolejeunea species (
Microlejeunea cochlearifolia is conspecific with Microlejeunea kamerunensis in
Microlejeunea lunulatiloba is conspecific with Microlejeunea ulicina in
Microlejeunea minutistipula is probably either Microlejeunea cochlearifolia or Microlejeunea africana (
Microlejeunea ovistipula is possibly conspecific with Lejeunea hepaticola (
Microlejeunea punctiformis was treated as conspecific with Microlejeunea ulicina by
Microlejeunea szechuanensis may be conspecific with Microlejeunea punctiformis.
Lepidolejeunea includes Kingiolejeunea, but one taxon has neither been transferred nor synonymized. It is listed in the “Names in genera not currently accepted” section below.
Rectolejeunea lindenbergii is a Lejeunea species (
Rectolejeunea lindigiana is a Lejeunea species of doubtful status (
Rectolejeunea pachyderma is a Lejeunea species of doubtful status (
Leptolejeunea borneensis may be conspecific with Leptolejeunea vitrea.
Leptolejeunea curvatifolia is possibly conspecific with Drepanolejeunea thwaitesiana, but the type specimen was burned in B and the identity can not be ascertained (
Leptolejeunea emarginata is possibly conspecific with Leptolejeunea apiculata (
Leptolejeunea massartiana is very possibly conspecific with Leptolejeunea elliptica judging from the original description and illustration (
Leptolejeunea revoluta may be conspecific with Leptolejeunea elliptica (
Leptolejeunea udarii is possibly conspecific with Leptolejeunea latifolia (
Pycnolejeunea anotomensis is a Cheilolejeunea species.
Pycnolejeunea cavistipula is possibly a Cheilolejeunea species.
Pycnolejeunea convexifolia is possibly conspecific with Cheilolejeunea imbricata.
Pycnolejeunea grossiloba is a Cheilolejeunea species, but its taxonomic status is unclear (
Pycnolejeunea palmicola is not a Pycnolejeunea species.
Pycnolejeunea schlimiana is not a Pycnolejeunea species.
Acrolejeunea abnormis was excluded from Acrolejeunea by
Acrolejeunea comptonii is a Mastigolejeunea species (
Acrolejeunea inflexa was excluded from Acrolejeunea by
Archilejeunea is polyphyletic and several nomenclatural and taxonomic will be proposed (
Archilejeunea bilabiata is a doubtful taxon, the type specimen could not be found (
Archilejeunea planifolia is conspecific with Archilejeunea kiushiana in
Archilejeunea negrensis is probably not an Archilejeunea species, but the type specimen is too poor to permit identification (
Archilejeunea ovata is conspecific with Lopholejeunea nigricans or Lopholejeunea subfusca (
Bryopteris fissiloba is probably conspecific with Bryopteris diffusa (
Lopholejeunea proxima is possibly conspecific with Lopholejeunea nigricans (
Lopholejeunea yapensis is possibly conspecific with Lopholejeunea subfusca (
Lopholejeunea aberrantia is probably an Archilejeunea species, but the type specimen in HIRO is destroyed (
Lopholejeunea vietnamica is probably an Archilejeunea species (
Mastigolejeunea humilis has been treated as conspecific with Mastigolejeunea auriculata by many recent authors, but it merits recognition based on molecular and morphological evidence (
Ptychanthus stephensonianus was tentatively treated as a separate species by
Lepidolaena includes Polyotus, but a few taxa have neither been transferred nor synonymized. They are listed in the “Names in genera not currently accepted” section below.
Porella acutifolia var. linguifolia was accepted by
Madotheca andica was transferred to Porella by Hässel (in
Porella caespitans was resolved as polyphyletic in the phylogeny of
Porella capehorniensis may be conspecific with Plagiochila setigera and the type specimen is probably not from South America (
Porella densifolia was resolved as paraphyletic in the phylogeny of
Porella maxima was accepted by
Porella oblongifolia is closely related to Porella densifolia (
Porella stephaniana is closely related to Porella densifolia (
Radula squarrosa is probably conspecific with Radula morobeana.
Radula bipinnata is distinct from Radula boryana according to molecular evidence (
Radula acuminata may be conspecific with Radula tjibodensis.
Radula tjibodensis is a doubtful taxon. The type specimen has not been found.
Radula crenulata is conspecific with Radula acuta in
Radula acutiloba is tentatively placed in subg. Radula. The specimen labelled Radula acutiloba in
Radula fendleri may be conspecific with Radula madagascariensis.
Radula multiflora is similar to, and may be conspecific with Radula reflexa, but it was accepted by
Radula oceania was considered conspecific with Radula javanica by
Radula decurrens may be conspecific with Radula reflexa.
Radula longispica may be conspecific with Radula javanica.
Radula paganii is doubtfully distinct from Radula neotropica.
Radula pinnulata was accepted by
Radula rupicola may be conspecific with Radula vieillardii (
Herzogianthus sanguineus is possibly conspecific with Herzogianthus vaginatus.
Aneura includes also Acrostolia and Pseudoneura, but a few taxa have never been transferred nor synonymized. They are listed in the section “Names in genera not currently accepted” below. The types of Aneura augustae, Aneura hunsteinii, Aneura latemultifida, Aneura ledermannii, Aneura subledermannii and Aneura subtenerrima were destroyed in B and their identity is doubtful (
Aneura crinita (type from Europe) has neither been recognized in any recent European treatment nor synonymized.
Aneura denticulata is possibly conspecific with Riccardia ciliolata (
Aneura sharpii is conspecific with Aneura maxima in
Aneura singalangana may be a form of Riccardia subexalata (
Riccardia includes Acrostolia, but a few taxa have neither been transferred nor synonymized. They are listed under “Names in genera not currently accepted” below.
Riccardia albomarginata is possibly conspecific with Riccardia canaliculata (
Riccardia baumannii is possibly conspecific with Riccardia grossitexta or Riccardia tenuicostata (
Riccardia baldwinii is possibly conspecific with Riccardia flaccida (
Riccardia decolyana is possibly conspecific with Riccardia multifida (
Riccardia diminuta is possibly conspecific with Riccardia elata (
Riccardia hirtiflora is possibly conspecific with Riccardia emarginata (
Riccardia multifidoides is conspecific with Riccardia multifida in
Riccardia palmatiformis was treated as a doubtful taxon with some affinity with Riccardia levieri by
Metzgeria was divided into many subdivisions proposed by Kuwahara in a series of publications, but they are artificial and not accepted here.
Metzgeria agnewiae is conspecific with Metzgeria consanguinea in
Metzgeria ciliata is conspecific with Metzgeria furcata in
Metzgeria conjugata is a complex taxon.
Metzgeria foliicola is conspecific with Metzgeria albinea in
Metzgeria furcata includes several cryptic species as demonstrated by
Metzgeria mexicana is conspecific with Metzgeria subundulata in
Fossombronia crispula is based on the description most likely Fossombronia pusilla, but no material has been found.
Fossombronia gregaria is possibly conspecific with Fossombronia australis (
Fossombronia grossepapillata may be conspecific with Fossombronia papillata, but the type specimen is too immature to allow determination (
Fossombronia integerrima is possibly conspecific with Fossombronia australis.
Fossombronia integrifolia is possibly conspecific with Fossombronia foveolata (
Fossombronia nigricaulis is possibly conspecific with Fossombronia australis (
Fossombronia subsaccata has a sterile and unidentifiable type specimen (
Jensenia crassifrons is possibly conspecific with Jensenia difformis.
Symphyogyna paucidens is a doubtful taxon.
Symphyogyna ulvoides is probably a filmy fern (
Noteroclada longiuscula is neither a Noteroclada nor a Fossombronia species. The type specimen could not be found (
Lunularia cruciata subsp. thaxteri was synonymized with subsp. cruciata by
Asterella includes Fimbraria and Hypenantron, but a few taxa have neither been transferred nor synonymized. They are listed in the “Names in genera not currently accepted” section below.
Asterella shimizuana is conspecific with Asterella khasyana in
Asterella tenerrima is probably conspecific with Asterella tenera.
Asterella caucasica is probably conspecific with Asterella whiteleggeana (
Asterella dognyensis is possibly conspecific with Asterella heteroflora.
Asterella preussii was provisionally placed in synonymy with Asterella bachmannii (
Mannia perssonii is possibly conspecific with Mannia sibirica.
Mannia hegewaldii is possibly conspecific with Mannia triandra.
Mannia paradoxa is possibly a Reboulia species.
Plagiochasma megacarpon may not be a Plagiochasma species.
Sauteria japonica is possibly conspecific with Sauteria spongiosa.
Sauteria inflata is possibly conspecific with Sauteria alpina (
Cyathodium spurium may be conspecific with Cyathodium cavernarum (
Dumortiera hirsuta is a species complex (
Marchantia assamica is a doubtful taxon.
Marchantia balboi is a doubtful taxon.
Marchantia balboi var. acutisquamata is a doubtful taxon,
Marchantia cagnii is a doubtful taxon.
Marchantia cengiana is a doubtful taxon.
Marchantia keniae is a doubtful taxon.
Marchantia papyracea is a doubtful taxon.
Marchantia sellae is a doubtful taxon.
Marchantia trilocularis is a doubtful taxon which
Marchantia tusui is a doubtful taxon.
Riccia ianthina seems identical to glabrous phases of Riccia atromarginata (
Riccia congoana is possibly conspecific with Riccia billardierei (
Riccia crustata may be conspecific with Riccia albida (
Riccia runssorensis is possibly conspecific with Riccia macrospora (
Riccia bahiensis is conspecific with Riccia cavernosa in
Riccia paranaensis may be conspecific with Riccia huebeneriana (
Riccia bialbistrata is a doubtful taxon closely related to Riccia lindmanii (
Riccia coronata is only known from the type specimen, which could not be traced. The description is very brief and it is suspected that it refers to smaller plants of Riccia natalensis (
Riccia gemmifera is possibly conspecific with Riccia atropurpurea (
Riccia intermedia (type from France) has neither been recognized in any recent treatment nor synonymized. The name may have priority once its identity is determined.
Riccia marginata is possibly conspecific with Riccia beyrichiana (
Riccia nigrescens is an Anthoceros species (
Riccia obtusa may be conspecific with Riccia gangetica (
Riccia prominens may be conspecific with Riccia junghuhniana (
Riccia saharensis is possibly conspecific with Riccia argenteolimbata (
Riccia tuberculata (type from France) has neither been recognized in any recent European treatment nor synonymized. The name may have priority once its identity is determined.
Riccia velenovskyi (type from Czech Republic) has neither been recognized in any recent European treatment nor synonymized. The name may have priority once its identity is determined.
Targionia dioica is probably conspecific with Targionia hypophylla (
Riella notarisii is a species complex with poorly defined taxonomic boundaries including Riella battandieri, Riella cyrenaica, Riella gallica, Riella indica, Riella reuteri and Riella sersuensis.
Sphaerocarpos europaeus is clearly distinct from Sphaerocarpos texanus (
Acrostolia is congeneric with Riccardia, but a few taxa have neither been transferred nor synonymized.
Aphanolejeunea lancifera is a Drepanolejeunea species (
Aphanolejeunea minima seems to be close to Cololejeunea gracilis based on the drawing in the original description, but the type specimen (not available for study) seems to be very fragmentary, having only reduced leaves, so its identity is uncertain (
Aspiromitus is congeneric with Anthoceros, but some taxa have neither been transferred nor synonymized.
Crossotolejeunea is congeneric with Lejeunea, but one taxon has neither been transferred nor synonymized.
Crossotolejeunea curvifolia has doubtful status, the type specimen could not be found in G (
Eulejeunea is congeneric with Lejeunea subg. Lejeunea, but a few taxa have neither been transferred nor synonymized.
Eulejeunea subpililoba is morphologically similar to Lejeunea spiniloba.
Euosmolejeunea is here treated as a subgenus of Cheilolejeunea, but a few taxa have neither been transferred nor synonymized.
Euosmolejeunea tenerrima is a doubtful taxon described with three syntypes, two from Brazil and one from Java. The Java material is probably not the same species as the material from Brazil.
Fimbraria is congeneric with Asterella, but the following taxa have neither been transferred nor synonymized, and their status is doubtful.
Fimbraria gigantea belongs to Asterella subg. Asterella sect. Brachyblepharis.
Fimbraria incrassata was provisionally placed in synonymy with Asterella abyssinica (
Fimbraria kamerunensis was provisionally placed in synonymy with Asterella abyssinica (
Hygrolejeunea is congeneric with Lejeunea, but some taxa have neither been transferred nor synonymized.
Hygrolejeunea patellirostris is conspecific with Lejeunea acuta in
Hygrolejeunea staudtiana is likely to be either Lejeunea brenanii or Lejeunea isophylla (
Hypenantron is congeneric with Asterella, but a few taxa have neither been transferred nor synonymized.
Jamesoniella is congeneric with Syzygiella (
Kingiolejeunea is congeneric with Lepidolejeunea, but one taxon has neither been transferred nor synonymized.
Leptocolea is congeneric with Cololejeunea, but one taxon has neither been transferred nor synonymized.
Leptocolea sumatrana is possibly conspecific with Cololejeunea equialbi.
Mastigobryum is congeneric with Bazzania, but many species have not been studied recently and thus not transferred. It is possible that some prove to be older names of existing taxa when their identities are known.
Nemoursia tuberculata is a Conocephalum species.
Physocolea is congeneric with Cololejeunea, but one taxon has neither been transferred nor synonymized.
Physocolea tambillensis is possibly conspecific with Myriocoleopsis minutissima.
Plectocolea is now regarded a subgenus of Solenostoma, but one taxon has neither been transferred nor synonymized.
Plectocolea subamoena is probably conspecific with Solenostoma amoena.
Polyotus is a superfluous name for Lepidolaena, but a few taxa have neither been transferred nor synonymized.
Schisma is congeneric with Herbertus, but a few taxa have neither been transferred nor synonymized.
Strepsilejeunea is congeneric with Cheilolejeunea, but some taxa have neither been transferred nor synonymized.
Thysanolejeunea africana may be conspecific with Caudalejeunea africana.
Trachylejeunea englishii is not a Trachylejeunea species (see